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<document id="8351ECCEDBBC9C691FFC2C13A1C6FBDE" ID-DOI="10.11646/zootaxa.5525.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14021479" ID-ZooBank="7F2F59B2-E0FB-4E17-BAF1-31228DB9428E" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1730450777893" checkinUser="plazi" docAuthor="Sak, Serdar, Karaytuğ, Süphan &amp; Huys, Rony" docDate="2024" docId="627EC678F743FF95FF4EFB607FD1FA92" docLanguage="en" docName="zootaxa.5525.1.1.pdf" docOrigin="Zootaxa 5525 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5525.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Neoleptastacus supersetosus Sak, Karaytuğ &amp; Huys, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="43" masterDocId="9E47BE00F766FFBFFFD9FFA97C60FFC7" masterDocTitle="Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov." masterLastPageNumber="66" masterPageNumber="1" pageNumber="38" updateTime="1730833424424" updateUser="guilherme">
<mods:mods id="4AAA788439FFB3B75BAD3AE441855F86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6F385673E3C00F80F602785833CB659E">
<mods:title id="B9B8AC284098412CD1CAB4CBEEF9A9B4">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
</mods:titleInfo>
<mods:name id="CE859EF1780A86543143CB5F741DFD31" type="personal">
<mods:role id="859FFD54306F2DBB3400C2D112D6A06A">
<mods:roleTerm id="4E8685C94247AE549794768DA0435BE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A51454A3D83A3363A9ECA91F68C0D79">Sak, Serdar</mods:namePart>
<mods:nameIdentifier id="D08A91A15AD53ECCF443FB15ADA0DD8B" type="ORCID">0000-0002-4955-3162</mods:nameIdentifier>
<mods:affiliation id="A887EFA9EA3E0DC1BB7BE966A926BB34">Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162</mods:affiliation>
<mods:nameIdentifier id="7E2D49FE6F82B0686AC95F6CEAB53010" type="email">serdarsak@gmail.com.</mods:nameIdentifier>
</mods:name>
<mods:name id="5E7138B66429879FA63323C012D016E8" type="personal">
<mods:role id="14F3BF34063730778A99605D1E434295">
<mods:roleTerm id="7984D12AB1D0101ED6C24E3838D2ACBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9983245203591FCB2FBD5061F162565C">Karaytuğ, Süphan</mods:namePart>
<mods:nameIdentifier id="9E54508366E19EB0FB6CA96BB7039E07" type="ORCID">0000-0001-8980-4133</mods:nameIdentifier>
<mods:affiliation id="7F690418C8DD4E3C0D51BE3E498A72A2">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
<mods:nameIdentifier id="238B1EDDD377631D0B00699BB3A572D6" type="email">suphankaraytug@gmail.com.</mods:nameIdentifier>
</mods:name>
<mods:name id="9275D9EE830F63D8FEC63BE6778CBC59" type="personal">
<mods:role id="3CE6BF45E39BC3E60865A30ED2C3E81D">
<mods:roleTerm id="2EAA17DD781028AD7D00DE9B608B72C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3359A39E23FFEBC010865439EDFA9B15">Huys, Rony</mods:namePart>
<mods:nameIdentifier id="93C2E2926F079C3CD44A515805D9920A" type="ORCID">0000-0003-2411-7003</mods:nameIdentifier>
<mods:affiliation id="1737F2FDA0E2F739EF8A86A695B7B315">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="61440C6184040D90F817927F10D85C7E" type="email">r.huys@nhm.ac.uk.</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="9974EB9194351382BA63BFA4A0EDEACD">text</mods:typeOfResource>
<mods:relatedItem id="8833184AFD96A4F7119DB5C747C04C07" type="host">
<mods:titleInfo id="F8FCC90B17E91F5A29B239BC503999DF">
<mods:title id="3F4C4E08F91D105F2642437110682C44">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="311D588E51769F1DB2FC9FE4C9819FE5">
<mods:date id="C6D9281E02C5435B72DB0DC565F45E9D">2024</mods:date>
<mods:detail id="D604A31ECB9514F99D39437DF10051D2" type="pubDate">
<mods:number id="091AA4ECEE3D7BC758C295C7E682893F">2024-10-21</mods:number>
</mods:detail>
<mods:detail id="4C4F579AF3330E468DAC060EE4CCCABD" type="volume">
<mods:number id="644143C58520FC13F9BF583EDDCA31F3">5525</mods:number>
</mods:detail>
<mods:detail id="48B5DD2EEC8977034A1B6FC89FCB410D" type="issue">
<mods:number id="2B01B70DE00D2C2FDFC49BF571A805DF">1</mods:number>
</mods:detail>
<mods:extent id="C4B93756357F0474AD56ABD08920410D" unit="page">
<mods:start id="A8B294567F7D12214D7E9A4655A8CF29">1</mods:start>
<mods:end id="D5832462CA2CC7E70B0D6343F509EDDC">66</mods:end>
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<mods:location id="B60B8B867944533614BC1AE3AA7D597F">
<mods:url id="11B61F72C09A3A9D8005B00ADC34A9CB">http://dx.doi.org/10.11646/zootaxa.5525.1.1</mods:url>
</mods:location>
<mods:classification id="2B69EDFD4FC92EE10FA3DBECEE6DC67F">journal article</mods:classification>
<mods:identifier id="736E2F0C90B553D6EB96A06A4E85C717" type="DOI">10.11646/zootaxa.5525.1.1</mods:identifier>
<mods:identifier id="BDB48EBF47F5243699C15F3D23B41308" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="E346A91D7D7579FE8F84045B6651220B" type="Zenodo-Dep">14021479</mods:identifier>
<mods:identifier id="12CE44A852D0330DA5E7F60D91D9CE6F" type="ZooBank">7F2F59B2-E0FB-4E17-BAF1-31228DB9428E</mods:identifier>
</mods:mods>
<treatment id="627EC678F743FF95FF4EFB607FD1FA92" LSID="urn:lsid:plazi:treatment:627EC678F743FF95FF4EFB607FD1FA92" httpUri="http://treatment.plazi.org/id/627EC678F743FF95FF4EFB607FD1FA92" lastPageId="42" lastPageNumber="43" pageId="37" pageNumber="38">
<subSubSection id="A2CD24E5F743FF9AFF4EFB607E33FB23" box="[151,595,1225,1252]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="EA68776EF743FF9AFF4EFB607E33FB23" blockId="37.[151,881,1225,1325]" box="[151,595,1225,1252]" pageId="37" pageNumber="38">
<heading id="B120C002F743FF9AFF4EFB607E33FB23" bold="true" box="[151,595,1225,1252]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="D8A3AB7CF743FF9AFF4EFB607E33FB23" bold="true" box="[151,595,1225,1252]" pageId="37" pageNumber="38">
<taxonomicName id="2DD70CEDF743FF9AFF4EFB607D8EFB23" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[151,494,1225,1252]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF743FF9AFF4EFB607D8EFB23" bold="true" box="[151,494,1225,1252]" italics="true" pageId="37" pageNumber="38">Neoleptastacus supersetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3901607F743FF9AFE2FFB637E33FB23" box="[502,595,1226,1252]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F743FF9AFF4EFB477F11FAEA" pageId="37" pageNumber="38" type="description">
<paragraph id="EA68776EF743FF9AFF4EFB477D78FACF" blockId="37.[151,881,1225,1325]" box="[151,280,1262,1288]" pageId="37" pageNumber="38">
(
<figureCitation id="72EC6BEBF743FF9AFF46FB477D70FACF" box="[159,272,1262,1288]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="38.[151,250,1943,1967]" captionStartId-1="39.[151,250,1928,1952]" captionStartId-2="40.[151,250,1943,1967]" captionTargetBox-0="[192,1396,191,1907]" captionTargetBox-1="[207,1380,189,1892]" captionTargetBox-2="[196,1382,193,1907]" captionTargetId-0="figure-17@38.[179,1408,181,1919]" captionTargetId-1="figure-17@39.[195,1392,181,1904]" captionTargetId-2="figure-17@40.[194,1394,181,1919]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionText-0="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." captionText-1="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." captionText-2="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.14021498" figureDoi-1="http://doi.org/10.5281/zenodo.14021500" figureDoi-2="http://doi.org/10.5281/zenodo.14021502" httpUri-0="https://zenodo.org/record/14021498/files/figure.png" httpUri-1="https://zenodo.org/record/14021500/files/figure.png" httpUri-2="https://zenodo.org/record/14021502/files/figure.png" pageId="37" pageNumber="38">Figs 9–11</figureCitation>
)
</paragraph>
<paragraph id="EA68776EF743FF9AFF4EFABB7F11FAEA" blockId="37.[151,881,1225,1325]" box="[151,881,1298,1325]" pageId="37" pageNumber="38">
https://zoobank.org/
<uuid id="9E714DBBF743FF9AFEA3FABB7F11FAEA" box="[378,881,1298,1325]" pageId="37" pageNumber="38">316E69F8-8817-4599-9126-F24A67BE3579</uuid>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F743FF9AFF4EFAFE7F60FAA9" box="[151,768,1366,1390]" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="EA68776EF743FF9AFF4EFAFE7F60FAA9" blockId="37.[151,768,1366,1390]" box="[151,768,1366,1390]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="CAC75040F743FF9AFF4EFAFE7F60FAA9" box="[151,768,1366,1390]" pageId="37" pageNumber="38">
<emphasis id="D8A3AB7CF743FF9AFF4EFAFE7E57FAA9" box="[151,567,1366,1390]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="2DD70CEDF743FF9AFF4EFAFE7DEEFAA9" authority="Sak, 2004" box="[151,398,1366,1390]" class="Copepoda" family="Arenopontiidae" genus="Pararenopontia" kingdom="Animalia" order="Harpacticoida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="poliseta">Pararenopontia poliseta</taxonomicName>
—nomen nudum
</emphasis>
by
<treatmentCitation id="6B76517FF743FF9AFD84FAFF7F60FAA9" author="Sak, S." box="[605,768,1366,1390]" page="273" pageId="37" pageNumber="38" year="2004">
<bibRefCitation id="8E460A9FF743FF9AFD84FAFF7F60FAA9" author="Sak, S." box="[605,768,1366,1390]" pageId="37" pageNumber="38" refId="ref49361" refString="Sak, S. (2004) Arenopontiidae (Copepoda: Harpacticoida) familyasinin taksonomisi ve filogenisi. [Taxonomy and phylogeny of the family Arenopontiidae (Copepoda: Harpacticoida)]. PhD dissertation, University of Balikesir, Balikesir, 312 pp." type="book" year="2004">Sak (2004: 273)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F743FF9AFF4EFA347F9CF9AC" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="EA68776EF743FF9AFF4EFA347865FA39" blockId="37.[151,1437,1436,2038]" pageId="37" pageNumber="38">
<emphasis id="D8A3AB7CF743FF9AFF4EFA347D55FA71" bold="true" box="[151,309,1436,1463]" pageId="37" pageNumber="38">Type locality.</emphasis>
<collectingCountry id="92C037FEF743FF9AFEE5FA357DF1FA71" box="[316,401,1436,1462]" name="Kuwait" pageId="37" pageNumber="38">Kuwait</collectingCountry>
,
<location id="EF0821B5F743FF9AFE45FA357E6CFA71" LSID="urn:lsid:plazi:treatment:627EC678F743FF95FF4EFB607FD1FA92:EF0821B5F743FF9AFE45FA357E6CFA71" box="[412,524,1436,1462]" country="Kuwait" county="Relatively" latitude="29.361612" longLatPrecision="1" longitude="47.955334" municipality="Kuwait" name="Shuwaikh" pageId="37" pageNumber="38">Shuwaikh</location>
(
<geoCoordinate id="8FE311A9F743FF9AFDC2FA347EA3FA70" box="[539,707,1437,1463]" degrees="29" direction="north" minutes="21" orientation="latitude" pageId="37" pageNumber="38" precision="1" seconds="41.8" value="29.361612">29°21’41.8” N</geoCoordinate>
,
<geoCoordinate id="8FE311A9F743FF9AFD17FA357F10FA70" box="[718,880,1436,1463]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="37" pageNumber="38" precision="1" seconds="19.2" value="47.955334">47°57’19.2” E</geoCoordinate>
) mid- and upper tide levels on public recreational beach within
<collectingMunicipality id="0A0CED14F743FF9AFEF7FA697DE0FA1D" box="[302,384,1472,1498]" pageId="37" pageNumber="38">Kuwait</collectingMunicipality>
Bay adjacent to the commercial port, surrounded by extensive urban development. Relatively steeply sloping sandy beach, bounded to seaward by extensive limestone flats.
</paragraph>
<paragraph id="EA68776EF743FF9AFF1EF9A07F9CF9AC" blockId="37.[151,1437,1436,2038]" pageId="37" pageNumber="38">
<emphasis id="D8A3AB7CF743FF9AFF1EF9A07D18F9E5" bold="true" box="[199,376,1544,1571]" pageId="37" pageNumber="38">Type material.</emphasis>
<materialsCitation id="5ABF7D33F743FF9AFE5BF9A178C5F9E5" box="[386,1189,1544,1571]" collectionCode="NHMUK" country="Kuwait" latitude="29.361612" location="Shuwaikh" longLatPrecision="1" longitude="47.955334" municipality="Kuwait" pageId="37" pageNumber="38" specimenCode="NHMUK 2024.1063" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="356CC9CCF743FF9AFE5BF9A17D8AF9E5" box="[386,490,1544,1570]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
<specimenCount id="FCD1BCE7F743FF9AFE2DF9A07E69F9E4" box="[500,521,1545,1571]" count="1" pageId="37" pageNumber="38" type="female">♀</specimenCount>
(dissected on eight slides) (reg. no
<specimenCode id="BA71DF15F743FF9AFC75F9A078FCF9E5" box="[940,1180,1544,1571]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="37" pageNumber="38" type="Museum">NHMUK 2024.1063</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="5ABF7D33F743FF9AFB6BF9A07F98F9AC" collectingDate="2004-06-03" collectionCode="NHMUK" collectorName="S. Livesey &amp; N. O'Brien" country="Kuwait" latitude="29.361612" location="Shuwaikh" longLatPrecision="1" longitude="47.955334" municipality="Kuwait" pageId="37" pageNumber="38" specimenCode="NHMUK 2024.1064-1066" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="356CC9CCF743FF9AFB6BF9A0797FF9E4" box="[1202,1311,1545,1571]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
are
<specimenCount id="FCD1BCE7F743FF9AFA8CF9A079FCF9E4" box="[1365,1436,1545,1571]" count="1" pageId="37" pageNumber="38" type="male">one ♂</specimenCount>
(dissected) and
<specimenCount id="FCD1BCE7F743FF9AFE93F9847DF1F980" box="[330,401,1581,1607]" count="1" pageId="37" pageNumber="38" type="female">one ♀</specimenCount>
and
<specimenCount id="FCD1BCE7F743FF9AFE13F9847E71F980" box="[458,529,1581,1607]" count="1" pageId="37" pageNumber="38" type="male">one ♂</specimenCount>
preserved in alcohol (reg. nos
<specimenCode id="BA71DF15F743FF9AFCACF98478CAF981" box="[885,1194,1580,1607]" pageId="37" pageNumber="38">NHMUK 2024.1064–1066</specimenCode>
). All specimens were collected from the type locality; leg.
<collectorName id="472212B8F743FF9AFDEDF9F87ECBF9AD" box="[564,683,1616,1643]" pageId="37" pageNumber="38">S. Livesey</collectorName>
and
<collectorName id="472212B8F743FF9AFD38F9F87F01F9AD" box="[737,865,1616,1643]" pageId="37" pageNumber="38">N. O’Brien</collectorName>
,
<date id="9E6951AEF743FF9AFCB4F9F87F98F9AC" box="[877,1016,1617,1643]" pageId="37" pageNumber="38" value="2004-06-03">
<collectingDate id="8E2DA846F743FF9AFCB4F9F87F98F9AC" box="[877,1016,1617,1643]" pageId="37" pageNumber="38" value="2004-06-03">3 June 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F743FF95FF1EF9DD7F29FC9A" lastPageId="42" lastPageNumber="43" pageId="37" pageNumber="38" type="description">
<paragraph id="EA68776EF743FF9AFF1EF9DD7840F8A0" blockId="37.[151,1437,1436,2038]" pageId="37" pageNumber="38">
<emphasis id="D8A3AB7CF743FF9AFF1EF9DD7DAFF949" bold="true" box="[199,463,1652,1678]" pageId="37" pageNumber="38">Description of female.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 225 μm (
<emphasis id="D8A3AB7CF743FF9AFAAEF9DC79E5F949" box="[1399,1413,1653,1678]" italics="true" pageId="37" pageNumber="38">n</emphasis>
= 1). Body (
<figureCitation id="72EC6BEBF743FF9AFEDEF9317D3EF974" box="[263,350,1688,1715]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="37" pageNumber="38">Fig. 9A</figureCitation>
) slender and cylindrical without clear distinction between prosome and urosome. Hyaline frills of thoracic somites weakly developed and smooth; those of genital double-somite and free abdominal somites strongly developed and consisting of rectangular digitate lappets (
<figureCitation id="72EC6BEBF743FF9AFCD4F9497F0AF93C" box="[781,874,1760,1787]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="37" pageNumber="38">Figs 9A</figureCitation>
;
<figureCitation id="72EC6BEBF743FF9AFCACF9487FC8F93C" box="[885,936,1761,1787]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="37" pageNumber="38">10A</figureCitation>
;
<figureCitation id="72EC6BEBF743FF9AFC6AF9487F83F93C" box="[947,995,1761,1787]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="37" pageNumber="38">11C</figureCitation>
). Somites connected by well-developed intersomitic membranes. Somites bearing P2–P5, genital double-somite and second abdominal somite with internal semicircular reinforcements forming pattern as figured. Genital double-somite (
<figureCitation id="72EC6BEBF743FF9AFBD0F8817806F884" box="[1033,1126,1832,1859]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="37" pageNumber="38">Figs 9A</figureCitation>
;
<figureCitation id="72EC6BEBF743FF9AFBA8F88078C2F884" box="[1137,1186,1833,1859]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="37" pageNumber="38">10A</figureCitation>
) about as long as wide (measured in ventral aspect); with two ventral pores in posterior half (
<figureCitation id="72EC6BEBF743FF9AFC7FF8E57873F8A0" box="[934,1043,1868,1895]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="37" pageNumber="38">Figs 10A</figureCitation>
).
</paragraph>
<paragraph id="EA68776EF743FF9AFF1EF8D97DB9F831" blockId="37.[151,1437,1436,2038]" pageId="37" pageNumber="38">
Anal somite (
<figureCitation id="72EC6BEBF743FF9AFE84F8D97DA4F84C" box="[349,452,1904,1931]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="37" pageNumber="38">Fig. 10A</figureCitation>
;
<figureCitation id="72EC6BEBF743FF9AFE17F8D87E7EF84C" box="[462,542,1905,1931]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="37" pageNumber="38">11C, D</figureCitation>
) without paired, dorsally recurved, spinous processes; with two sensilla dorsally and few spinules either side of ventral midline.Anal operculum well developed, semicircular, without ornamentation; covering terminally positioned anal opening. Anal frill triradiate, distinctly incised (giving a spinulose appearance underneath anal operculum).
</paragraph>
<caption id="BEA827E6F740FF99FF4EF83E7EE2F83F" ID-DOI="http://doi.org/10.5281/zenodo.14021498" ID-Zenodo-Dep="14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1943,1967]" targetBox="[192,1396,191,1907]" targetPageId="38" targetType="figure">
<paragraph id="EA68776EF740FF99FF4EF83E7EE2F83F" blockId="38.[151,1436,1943,2040]" pageId="38" pageNumber="39">
<emphasis id="D8A3AB7CF740FF99FF4EF83E7ECAF877" bold="true" box="[151,682,1943,1968]" pageId="38" pageNumber="39">
FIGURE 9.
<taxonomicName id="2DD70CEDF740FF99FEF9F83E7E2CF868" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[288,588,1943,1967]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF740FF99FEF9F83E7E2CF868" bold="true" box="[288,588,1943,1967]" italics="true" pageId="38" pageNumber="39">Neoleptastacus supersetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3901607F740FF99FD8FF8317ECAF877" box="[598,682,1944,1968]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior.
</paragraph>
</caption>
<caption id="BEA827E6F741FF98FF4EF8217D75F803" ID-DOI="http://doi.org/10.5281/zenodo.14021500" ID-Zenodo-Dep="14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1928,1952]" targetBox="[207,1380,189,1892]" targetPageId="39" targetType="figure">
<paragraph id="EA68776EF741FF98FF4EF8217D75F803" blockId="39.[151,1436,1928,1988]" pageId="39" pageNumber="40">
<emphasis id="D8A3AB7CF741FF98FF4EF8217EC3F867" bold="true" box="[151,675,1928,1952]" pageId="39" pageNumber="40">
FIGURE 10.
<taxonomicName id="2DD70CEDF741FF98FEFCF8217E2DF867" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[293,589,1928,1952]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF741FF98FEFCF8217E2DF867" bold="true" box="[293,589,1928,1952]" italics="true" pageId="39" pageNumber="40">Neoleptastacus supersetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3901607F741FF98FD8AF8217EC3F867" box="[595,675,1928,1952]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral.
</paragraph>
</caption>
<caption id="BEA827E6F74EFF97FF4EF83E7D16F83F" ID-DOI="http://doi.org/10.5281/zenodo.14021502" ID-Zenodo-Dep="14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1943,1967]" targetBox="[196,1382,193,1907]" targetPageId="40" targetType="figure">
<paragraph id="EA68776EF74EFF97FF4EF83E7D16F83F" blockId="40.[151,1437,1943,2040]" pageId="40" pageNumber="41">
<emphasis id="D8A3AB7CF74EFF97FF4EF83E7ECCF877" bold="true" box="[151,684,1943,1968]" pageId="40" pageNumber="41">
FIGURE 11.
<taxonomicName id="2DD70CEDF74EFF97FEF1F83E7E31F868" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[296,593,1943,1967]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF74EFF97FEF1F83E7E31F868" bold="true" box="[296,593,1943,1967]" italics="true" pageId="40" pageNumber="41">Neoleptastacus supersetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3901607F74EFF97FD80F8317ECCF877" box="[601,684,1944,1968]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal.
</paragraph>
</caption>
<paragraph id="EA68776EF74FFF96FF1EFF3E7FB0FE6A" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
Caudal rami (
<figureCitation id="72EC6BEBF74FFF96FEBFFF3E7DB6FF76" box="[358,470,151,177]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="41" pageNumber="42">Figs 10A</figureCitation>
;
<figureCitation id="72EC6BEBF74FFF96FE3DFF3E7E58FF76" box="[484,568,151,177]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="41" pageNumber="42">11C, D</figureCitation>
) about 2.2 times longer than wide (measured in lateral view from anterior margin to apex of spinous process), distinctly tapering posteriorly with outer margin almost straight and inner margin distinctly convex; outer distal corner produced into short posteriorly directed, dorsally recurved spinous process; with pore ventrally near proximal margin (
<figureCitation id="72EC6BEBF74FFF96FD3BFEAA7F2AFEDA" box="[738,842,259,285]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="41" pageNumber="42">Fig. 10A</figureCitation>
) and dorsally at base of spinous process (
<figureCitation id="72EC6BEBF74FFF96FAF1FEAA79EEFEDA" box="[1320,1422,259,285]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="41" pageNumber="42">Fig. 11C</figureCitation>
); inner margin with conspicuous spinules near base of seta VII (
<figureCitation id="72EC6BEBF74FFF96FC82FE8E7FA1FE86" box="[859,961,295,321]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="41" pageNumber="42">Fig. 11C</figureCitation>
). Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, naked, located between seta V and posterior spinous process; seta V long, naked, with proximal fracture plane; seta VI small, naked and located at inner distal corner; seta VII distinctly foliaceous and tri-articulate at base.
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFE1E7E39FE32" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
Rostrum (
<figureCitation id="72EC6BEBF74FFF96FEECFE1E7DF7FE16" box="[309,407,439,465]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="41" pageNumber="42">Fig. 11B</figureCitation>
) small, broadly subtriangular, tapering distally; apical part lobate and demarcated by bilateral constrictions; with two delicate sensilla.
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFE5678E2FD6E" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
Antennule (
<figureCitation id="72EC6BEBF74FFF96FE95FE567DCFFDDE" box="[332,431,511,537]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="41" pageNumber="42">Fig. 11E</figureCitation>
) moderately long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about three times longer than wide. Segment 4 with long aesthetasc (L: 21 μm) fused at base with seta. Distal segment with seven setae (none distinctly spatulate) and apical acrothek consisting of short aesthetasc (L: 12 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[7 + acrothek].
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFD1A783AFC0E" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
Antenna (
<figureCitation id="72EC6BEBF74FFF96FEEDFD1A7DCCFD0A" box="[308,428,691,717]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="41" pageNumber="42">Fig. 9B, C</figureCitation>
). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment forming incompletely fused allobasis, about 2.5 times as long as maximum width; original basis-endopod boundary marked by short transverse surface suture at level of exopodal articulation and by constriction at abexopodal side; proximal part representing original basis with single spinules at distal corner. Exopod one-segmented, unornamented and elongate, with a naked apical seta (about 1.2 times longer than exopod). Free endopodal segment with few lateral spinules proximally and transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in
<figureCitation id="72EC6BEBF74FFF96FE79FC227D97FC62" box="[416,503,907,933]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="41" pageNumber="42">Fig. 9B</figureCitation>
); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta.
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFC7A79FBFC2A" blockId="41.[151,1437,151,1761]" box="[199,1435,979,1005]" pageId="41" pageNumber="42">
Mandible, maxillule, maxilla and maxilliped as in
<taxonomicName id="2DD70CEDF74FFF96FD25FC7A7FD9FC2A" authorityName="Nicholls" authorityYear="1945" box="[764,953,979,1005]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF74FFF96FD25FC7A7FD9FC2A" box="[764,953,979,1005]" italics="true" pageId="41" pageNumber="42">N. spinicaudatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8E460A9FF74FFF96FC2DFC7A78C0FC2A" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[1012,1184,979,1005]" pageId="41" pageNumber="42" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF74FFF96FBFCFC7A783DFC2A" box="[1061,1117,979,1005]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2008
</bibRefCitation>
:
<figureCitation id="72EC6BEBF74FFF96FB72FC7A7954FC2A" box="[1195,1332,979,1005]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="41" pageNumber="42">Figs 16D, E</figureCitation>
;
<figureCitation id="72EC6BEBF74FFF96FA99FC7A79EEFC2A" box="[1344,1422,979,1005]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="41" pageNumber="42">17E, F</figureCitation>
).
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFC5E7D6BFB2E" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
P1 (
<figureCitation id="72EC6BEBF74FFF96FF2FFC5E7D32FBD6" box="[246,338,1015,1041]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="41" pageNumber="42">Fig. 9D</figureCitation>
). Intercoxal sclerite wide and subrectangular. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis without spinular ornamentation; anterior surface with small naked seta near medial margin. Exopod two-segmented; segments with several spinules around outer margin; exp-1 unarmed; exp-2 with unipinnate outer spine and two geniculate setae distally. Endopod two-segmented, not prehensile, slightly longer than exopod; enp-1 about 1.4 times longer than enp-2, with a serrate inner seta arising from distal third and four coarse spinules along outer margin; enp-2 without spinules, distal margin with naked outer spine and geniculate inner seta.
</paragraph>
<paragraph id="EA68776EF74FFF96FF1EFB5A7F7DF926" blockId="41.[151,1437,151,1761]" pageId="41" pageNumber="42">
P2–P4 (
<figureCitation id="72EC6BEBF74FFF96FEC6FB5A7DF8FACA" box="[287,408,1267,1293]" captionStart="FIGURE 9" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[192,1396,191,1907]" captionTargetId="figure-17@38.[179,1408,181,1919]" captionTargetPageId="38" captionText="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021498" httpUri="https://zenodo.org/record/14021498/files/figure.png" pageId="41" pageNumber="42">Fig. 9F–G</figureCitation>
). Intercoxal sclerites rectangular with deeply concave ventral margins. Praecoxae triangular, small and naked. Coxae wider than long and without ornamentation. Bases smaller than coxae; anterior surface with a pore; outer basal seta absent in P2, naked and long (P3) or short (P4) in remaining legs; without ornamentation except for spinular row near base of P4 endopod. Exopods three-segmented; segments with coarse spinular ornamentation along outer margin, as illustrated; outer spine of segments naked (except for P2 exp-2 sparsely unipinnate); exp-3 with an outer unipinnate spine, and two setae distally (both naked in P2, one unipinnate and one bipinnate in P3–P4); outer spine of P2 exp-2 very long, extending far beyond exp-3 and about as long as entire exopod; P4 exp-2 elongate, distinctly longer than exp-1; segments of P4 about equally long, exp-3 with inner serrate seta in distal quarter originating from posterior surface. Endopods one- (P2–P3) or two-segmented (P4); P2–P3 endopods with few coarse spinules along both inner and outer margin; P4 enp-1 elongate, 3.7 times as long as enp-2, with few spinules along outer margin; P4 enp-2 without ornamentation; P2 endopod with long, apically serrate, backwardly directed seta along inner margin and one long unipinnate seta apically; P3 endopod shorter than P2 endopod, with short, naked spine apically; apical margin of P4 enp-2 with long, distally serrate and basally fused, inner seta, and short naked outer seta. Spine and seta formula as follows:
</paragraph>
<paragraph id="EA68776EF74FFF96FE6EF8A67F76F87E" pageId="41" pageNumber="42">
<table id="98D785CEF74F0040FF2EF8A67F5CF87E" box="[247,828,1807,1977]" gridcols="3" gridrows="4" inLine="true" pageId="41" pageNumber="42">
<tr id="54E7752CF74F0040FF2EF8A67F5CF8EE" box="[247,828,1807,1833]" gridrow="0" pageId="41" pageNumber="42" rowspan-0="1">
<th id="17361C50F74F0040FE6EF8A67E6EF8EE" box="[439,526,1807,1833]" gridcol="1" gridrow="0" pageId="41" pageNumber="42">Exopod</th>
<th id="17361C50F74F0040FD0EF8A67F5CF8EE" box="[727,828,1807,1833]" gridcol="2" gridrow="0" pageId="41" pageNumber="42">Endopod</th>
</tr>
<tr id="54E7752CF74F0040FF2EF8FE7F5CF8B6" box="[247,828,1879,1905]" gridrow="1" pageId="41" pageNumber="42">
<th id="17361C50F74F0040FF2EF8FE7D75F8B6" box="[247,277,1879,1905]" gridcol="0" gridrow="1" pageId="41" pageNumber="42">P2</th>
<td id="17361C50F74F0040FE6EF8FE7E6EF8B6" box="[439,526,1879,1905]" gridcol="1" gridrow="1" pageId="41" pageNumber="42">0.0.021</td>
<td id="17361C50F74F0040FD0EF8FE7F5CF8B6" box="[727,828,1879,1905]" gridcol="2" gridrow="1" pageId="41" pageNumber="42">110</td>
</tr>
<tr id="54E7752CF74F0040FF2EF8D27F5CF852" box="[247,828,1915,1941]" gridrow="2" pageId="41" pageNumber="42">
<th id="17361C50F74F0040FF2EF8D27D75F852" box="[247,277,1915,1941]" gridcol="0" gridrow="2" pageId="41" pageNumber="42">P3</th>
<td id="17361C50F74F0040FE6EF8D27E6EF852" box="[439,526,1915,1941]" gridcol="1" gridrow="2" pageId="41" pageNumber="42">0.0.021</td>
<td id="17361C50F74F0040FD0EF8D27F5CF852" box="[727,828,1915,1941]" gridcol="2" gridrow="2" pageId="41" pageNumber="42">010</td>
</tr>
<tr id="54E7752CF74F0040FF2EF8367F5CF87E" box="[247,828,1951,1977]" gridrow="3" pageId="41" pageNumber="42">
<th id="17361C50F74F0040FF2EF8367D75F87E" box="[247,277,1951,1977]" gridcol="0" gridrow="3" pageId="41" pageNumber="42">P4</th>
<td id="17361C50F74F0040FE6EF8367E6EF87E" box="[439,526,1951,1977]" gridcol="1" gridrow="3" pageId="41" pageNumber="42">0.0.121</td>
<td id="17361C50F74F0040FD0EF8367F5CF87E" box="[727,828,1951,1977]" gridcol="2" gridrow="3" pageId="41" pageNumber="42">0.020</td>
</tr>
</table>
</paragraph>
<paragraph id="EA68776EF74CFF95FF4EFF3E7D71FE86" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
Fifth legs (
<figureCitation id="72EC6BEBF74CFF95FECFFF3E7D1EFF76" box="[278,382,151,177]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="42" pageNumber="43">Fig. 10A</figureCitation>
) closely set together, but not touching in ventral midline. Baseoendopod and exopod fused, forming a subrectangular plate; with one pore on anterior surface and two pores along inner margin; inner distal corner with strong, bipinnate, spinous process (homologous to inner spine); distal margin with a minute outer seta, one long naked seta, and two short, subequally long (outer one slightly longer), naked spines; outer basal seta long and naked.
</paragraph>
<paragraph id="EA68776EF74CFF95FF1EFEE27885FE4E" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
Sixth pair of legs (
<figureCitation id="72EC6BEBF74CFF95FE48FEE27D96FEA2" box="[401,502,331,357]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="42" pageNumber="43">Fig. 10A</figureCitation>
) reduced, fused medially forming common genital operculum; armature represented by vestigial spiniform element on either side. Copulatory pore conspicuous (arrowed in
<figureCitation id="72EC6BEBF74CFF95FBA8FEC678B8FE4E" box="[1137,1240,367,393]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="42" pageNumber="43">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="EA68776EF74CFF95FF1EFE3B7DF0FDDE" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
<emphasis id="D8A3AB7CF74CFF95FF1EFE3B7DD7FE6A" bold="true" box="[199,439,402,429]" pageId="42" pageNumber="43">Description of male.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 235 µm (
<emphasis id="D8A3AB7CF74CFF95FA8BFE3A7900FE6B" box="[1362,1376,403,428]" italics="true" pageId="42" pageNumber="43">n</emphasis>
= 1). Body ornamentation essentially as in female except for internal semicircular reinforcements present only on somites bearing P2–P4 (
<figureCitation id="72EC6BEBF74CFF95FE96FE727DD8FE32" box="[335,440,475,501]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="42" pageNumber="43">Fig. 11A</figureCitation>
). Sexual dimorphism in antennule, genital segmentation and P6. Spermatophore length approximately 33 µm.
</paragraph>
<paragraph id="EA68776EF74CFF95FF1EFD8A7936FCD2" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
Antennule (
<figureCitation id="72EC6BEBF74CFF95FE93FD8A7DCAFDFA" box="[330,426,547,573]" captionStart="FIGURE 11" captionStartId="40.[151,250,1943,1967]" captionTargetBox="[196,1382,193,1907]" captionTargetId="figure-17@40.[194,1394,181,1919]" captionTargetPageId="40" captionText="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021502" httpUri="https://zenodo.org/record/14021502/files/figure.png" pageId="42" pageNumber="43">Fig. 11F</figureCitation>
) 9-segmented, haplocer; geniculation between segments 7 and 8. Segment 1 with a slender naked seta; segment 2 longest and about 2.4 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae and a spine; segment 4 an incomplete sclerite with one seta and one spiniform element; segment 5 with a long aesthetasc (40 µm) fused basally to slender seta; segment 6 with a seta; segment 7 with a seta and a spine; segment apparently unarmed; distal segment with six setae (none of which spatulate) and apical acrothek consisting of short aesthetasc (10 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4 + 1 spine], 4-[(1 + 1 spine)], 5-[1 + (1 + ae)], 6-[1], 7-[1 + 1 spine], 8-[0], 9-[6 + acrothek].
</paragraph>
<paragraph id="EA68776EF74CFF95FF1EFCB67F29FC9A" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
Sixth legs (
<figureCitation id="72EC6BEBF74CFF95FE9CFCB67DC9FCFE" box="[325,425,799,825]" captionStart="FIGURE 10" captionStartId="39.[151,250,1928,1952]" captionTargetBox="[207,1380,189,1892]" captionTargetId="figure-17@39.[195,1392,181,1904]" captionTargetPageId="39" captionText="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021500" httpUri="https://zenodo.org/record/14021500/files/figure.png" pageId="42" pageNumber="43">Fig. 10B</figureCitation>
) slightly asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer seta and a short inner spine, both elements being naked.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F74CFF95FF1EFCCE7876FC62" pageId="42" pageNumber="43" type="etymology">
<paragraph id="EA68776EF74CFF95FF1EFCCE7876FC62" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
<emphasis id="D8A3AB7CF74CFF95FF1EFCCE7D2EFC46" bold="true" box="[199,334,871,897]" pageId="42" pageNumber="43">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="D8A3AB7CF74CFF95FC91FCCE7FE6FC47" box="[840,902,871,896]" italics="true" pageId="42" pageNumber="43">super</emphasis>
, meaning over, and
<emphasis id="D8A3AB7CF74CFF95FBBDFCCE794FFC46" box="[1124,1327,871,897]" italics="true" pageId="42" pageNumber="43">s(a)eta, s(a)etosus</emphasis>
, meaning bristle, bristly, and refers to the supernumerary element on the P5 of both sexes.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F74CFF95FF1EFC067FD1FA92" pageId="42" pageNumber="43" type="discussion">
<paragraph id="EA68776EF74CFF95FF1EFC067FD1FA92" blockId="42.[151,1437,151,1365]" pageId="42" pageNumber="43">
<emphasis id="D8A3AB7CF74CFF95FF1EFC067D5BFC0E" bold="true" box="[199,315,943,969]" pageId="42" pageNumber="43">Remarks.</emphasis>
<taxonomicName id="2DD70CEDF74CFF95FE9DFC067EE7FC0F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[324,647,943,969]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF74CFF95FE9DFC067EE7FC0F" box="[324,647,943,969]" italics="true" pageId="42" pageNumber="43">Neoleptastacus supersetosus</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF74CFF95FD49FC067E8BFC0E" bold="true" box="[656,747,943,969]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="C3901607F74CFF95FD49FC067E8BFC0E" box="[656,747,943,969]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is assigned to the
<taxonomicName id="2DD70CEDF74CFF95FC1AFC06784EFC0F" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[963,1070,943,968]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF74CFF95FC1AFC06784EFC0F" box="[963,1070,943,968]" italics="true" pageId="42" pageNumber="43">trisetosus</emphasis>
</taxonomicName>
-group based on (1) the absence of paired spinous processes on the anal somite, (2) a well-developed anal operculum, (3) P1 exp-1 without outer spine and exp-2 with three setae/spines, (4) P2 exp-2 with very long outer setiform element extending far beyond distal margin of exp-3, (5) endopod P2–P3 1-segmented with inner seta and one distal spine on P2 and one distal spine on P3, and (6) outer seta of P4 enp-2 reduced. Within this group it is morphologically closest to
<taxonomicName id="2DD70CEDF74CFF95FAC9FB9679FCFB9F" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[1296,1436,1087,1112]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF74CFF95FAC9FB9679FCFB9F" box="[1296,1436,1087,1112]" italics="true" pageId="42" pageNumber="43">N. trisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF74CFF95FF13FBCA7D12FBBB" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[202,370,1123,1148]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="D8A3AB7CF74CFF95FF13FBCA7D12FBBB" box="[202,370,1123,1148]" italics="true" pageId="42" pageNumber="43">N. panamensis</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF74CFF95FEA4FBCA7DBAFBBA" bold="true" box="[381,474,1123,1149]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="C3901607F74CFF95FEA4FBCA7DBAFBBA" box="[381,474,1123,1149]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with which it shares the 2-segmented condition of the P1 exopod. Characters that differentiate the new species from these two congeners include (1) somites bearing P2–P5, genital double-somite and second abdominal somite with internal semicircular reinforcements in
<specimenCount id="FCD1BCE7F74CFF95FC3BFB027F97FB02" box="[994,1015,1195,1221]" count="1" pageId="42" pageNumber="43" type="female">♀</specimenCount>
, and on somites bearing P2–P
<quantity id="2D2FDA8BF74CFF95FA97FB027919FB02" box="[1358,1401,1195,1221]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="42" pageNumber="43" unit="in" value="4.0">4 in</quantity>
<specimenCount id="FCD1BCE7F74CFF95FA59FB0279F7FB02" box="[1408,1431,1195,1221]" count="1" pageId="42" pageNumber="43" type="male">♂</specimenCount>
, (2) inner spine of P5 fused to segment, forming spinous process in both sexes (
<emphasis id="D8A3AB7CF74CFF95FBCEFB66784EFB2F" box="[1047,1070,1231,1256]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
delimited at base), and (3) male P6 with outer seta and inner spiniform element (
<emphasis id="D8A3AB7CF74CFF95FD1BFB5A7EB9FACB" box="[706,729,1267,1292]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
outer seta only). An additional character that differentiates
<taxonomicName id="2DD70CEDF74CFF95FA5AFB5A7D46FAF7" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
<emphasis id="D8A3AB7CF74CFF95FA5AFB5A7D46FAF7" italics="true" pageId="42" pageNumber="43">N. supersetosus</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF74CFF95FEF4FABE7DE6FAF6" bold="true" box="[301,390,1303,1329]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="C3901607F74CFF95FEF4FABE7DE6FAF6" box="[301,390,1303,1329]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
readily from all known species of
<taxonomicName id="2DD70CEDF74CFF95FCD3FABE7FD5FAF6" authorityName="Nicholls" authorityYear="1945" box="[778,949,1303,1329]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="D8A3AB7CF74CFF95FCD3FABE7FD5FAF6" box="[778,949,1303,1329]" italics="true" pageId="42" pageNumber="43">Neoleptastacus</emphasis>
</taxonomicName>
(and in fact all other
<taxonomicName id="2DD70CEDF74CFF95FB7BFABE7932FAF6" authorityName="Martinez Arbizu &amp; Moura" authorityYear="1994" box="[1186,1362,1303,1329]" class="Copepoda" family="Arenopontiidae" kingdom="Animalia" order="Harpacticoida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Arenopontiidae</taxonomicName>
) is the presence of a supernumerary setiform element on the P5 of both sexes.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C3B137723D20775C4CA28F999BE22D2E">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title> <mods:title id="E41DE21965D46E9C217F020276FFD57A">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
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<mods:namePart id="04AFF071257C940BF67CE3B06EFD94B0">Sak, Serdar</mods:namePart> <mods:namePart id="23BF70D6243169F6D0F0786061473869">Sak, Serdar</mods:namePart>
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<mods:affiliation id="B4BFC4F17B66159D385EF134AADD3B33">Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162</mods:affiliation> <mods:affiliation id="8A0F0F8C28230FDD5B04FAEEA4CCA840">Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162</mods:affiliation>
<mods:nameIdentifier id="BF10902350646751D162427F9C8253AF" type="email">serdarsak@gmail.com.</mods:nameIdentifier> <mods:nameIdentifier id="729A0002742462831FC55D100B5840D2" type="email">serdarsak@gmail.com.</mods:nameIdentifier>
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<mods:namePart id="34D505456A8315093F95E2AEFC05D058">Karaytuğ, Süphan</mods:namePart> <mods:namePart id="44A83D832E85551E3175F14146686A81">Karaytuğ, Süphan</mods:namePart>
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<mods:affiliation id="73AD3D025751A8DFB8B9A8E4D42E9E84">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation> <mods:affiliation id="8C5B13D7DC5933DCDB9CC9D7C9A15093">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
<mods:nameIdentifier id="7DD8112CCD7F9BF7A68A36D92DC4F58B" type="email">suphankaraytug@gmail.com.</mods:nameIdentifier> <mods:nameIdentifier id="679185DA7FED3A84B3D8AD2647786FF1" type="email">suphankaraytug@gmail.com.</mods:nameIdentifier>
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<mods:namePart id="16CBDB78CB3C1480FA7BADA26CBCD2A5">Huys, Rony</mods:namePart> <mods:namePart id="7F77C1FD8183794E869255479289CDC6">Huys, Rony</mods:namePart>
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<mods:affiliation id="7BE4DE34C8C154CF80B7A43512F6149E">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation> <mods:affiliation id="DA8167D40D881EE96E188AA49BCC10BF">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation>
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<emphasis id="D8A3AB7CF74BFF92FF4EF98E7D27F986" bold="true" box="[151,327,1575,1601]" pageId="45" pageNumber="46">Key to species</emphasis> <emphasis id="D8A3AB7CF74BFF92FF4EF98E7D27F986" bold="true" box="[151,327,1575,1601]" pageId="45" pageNumber="46">
Key to
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Redefinition of the generic boundaries in the Redefinition of the generic boundaries in the
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@ -96,48 +101,93 @@ and
(both (both
<emphasis id="D8A3AB7CF74BFF92FDC7F9117E9DF916" box="[542,765,1719,1745]" italics="true" pageId="45" pageNumber="46">species inquirendae</emphasis> <emphasis id="D8A3AB7CF74BFF92FDC7F9117E9DF916" box="[542,765,1719,1745]" italics="true" pageId="45" pageNumber="46">species inquirendae</emphasis>
) are included in ) are included in
<tableCitation id="A75542D5F74BFF92FC18F91E7876F916" box="[961,1046,1719,1745]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." pageId="45" pageNumber="46">Table 2</tableCitation> <tableCitation id="A75542D5F74BFF92FC18F91E7876F916" box="[961,1046,1719,1745]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="45" pageNumber="46" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
but not in the updated key below. Identifications made with this key must be confirmed by reference to the original descriptions in the literature. but not in the updated key below. Identifications made with this key must be confirmed by reference to the original descriptions in the literature.
</paragraph> </paragraph>
<paragraph id="EA68776EF74BFF92FF4EF88979FDF8F2" blockId="45.[151,1437,1824,2022]" box="[151,1437,1824,1846]" pageId="45" pageNumber="46">1. Anal somite with paired dorsolateral spinous spurs or processes................................................2.</paragraph> </subSubSection>
<paragraph id="EA68776EF74BFF92FF3BF89479FCF8B7" blockId="45.[151,1437,1824,2022]" pageId="45" pageNumber="46">Anal somite without paired dorsolateral spurs or processes...................................................13. 2. Abdominal somites with integumental pattern of rectangular plates dorsally and ventrally; P2 enp-2 without inner seta.....3.</paragraph> <subSubSection id="A2CD24E5F74BFF91FF4EF88979FCF965" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="key">
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Abdominal somites without conspicuous surface sculpturing; P2 enp-2 with inner seta..............................4. 3. P3 enp-2 with two distal elements; P5 of both sexes with four articulating setae/spines................... <keyStep id="51236FCBF74BFF92FF4EF88979FDF894" pageId="45" pageNumber="46">
<paragraph id="EA68776EF74BFF92FF4EF88979FDF8F2" blockId="45.[151,1437,1824,2022]" box="[151,1437,1824,1846]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF4EF88979FDF8F2" box="[151,1437,1824,1846]" pageId="45" pageNumber="46">1. Anal somite with paired dorsolateral spinous spurs or processes................................................2.</keyLead>
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<keyLead id="5126D45BF74BFF92FF3BF89479FDF894" box="[226,1437,1853,1875]" pageId="45" pageNumber="46">Anal somite without paired dorsolateral spurs or processes...................................................13.</keyLead>
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<paragraph id="EA68776EF74BFF92FF4EF8F279FCF8B7" blockId="45.[151,1437,1824,2022]" box="[151,1436,1883,1905]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF4EF8F279FCF8B7" box="[151,1436,1883,1905]" pageId="45" pageNumber="46">2. Abdominal somites with integumental pattern of rectangular plates dorsally and ventrally; P2 enp-2 without inner seta.....3.</keyLead>
</paragraph>
<paragraph id="EA68776EF74BFF92FF3BF8D179FDF84A" blockId="45.[151,1437,1824,2022]" box="[226,1437,1912,1934]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF3BF8D179FDF84A" box="[226,1437,1912,1934]" pageId="45" pageNumber="46">Abdominal somites without conspicuous surface sculpturing; P2 enp-2 with inner seta..............................4.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF74BFF92FF4EF83C79FDF80F" pageId="45" pageNumber="46">
<paragraph id="EA68776EF74BFF92FF4EF83C79FDF86C" blockId="45.[151,1437,1824,2022]" box="[151,1437,1941,1963]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF4EF83C79FDF86C" box="[151,1437,1941,1963]" pageId="45" pageNumber="46">
3. P3 enp-2 with two distal elements; P5 of both sexes with four articulating setae/spines...................
<emphasis id="D8A3AB7CF74BFF92FAD0F83F79FDF86C" box="[1289,1437,1942,1963]" italics="true" pageId="45" pageNumber="46"> <emphasis id="D8A3AB7CF74BFF92FAD0F83F79FDF86C" box="[1289,1437,1942,1963]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="2DD70CEDF74BFF92FAD0F83F79F9F86C" ID-CoL="46KMP" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[1289,1433,1942,1963]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ornamentus">N. ornamentus</taxonomicName> <taxonomicName id="2DD70CEDF74BFF92FAD0F83F79F9F86C" ID-CoL="46KMP" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[1289,1433,1942,1963]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ornamentus">N. ornamentus</taxonomicName>
. .
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF74BFF92FF3BF81A79FCF821" blockId="45.[151,1437,1824,2022]" pageId="45" pageNumber="46"> <paragraph id="EA68776EF74BFF92FF3BF81A79FDF80F" blockId="45.[151,1437,1824,2022]" box="[226,1437,1971,1993]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF3BF81A79FDF80F" box="[226,1437,1971,1993]" pageId="45" pageNumber="46">
P3 enp-2 with one distal element; P5 of both sexes with three articulating setae/spines................... P3 enp-2 with one distal element; P5 of both sexes with three articulating setae/spines...................
<emphasis id="D8A3AB7CF74BFF92FB27F81A79FDF80F" box="[1278,1437,1971,1992]" italics="true" pageId="45" pageNumber="46"> <emphasis id="D8A3AB7CF74BFF92FB27F81A79FDF80F" box="[1278,1437,1971,1992]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="2DD70CEDF74BFF92FB27F81A79FAF80F" ID-CoL="GFQQ" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[1278,1434,1971,1992]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="reductaspina">N. reductaspina</taxonomicName> <taxonomicName id="2DD70CEDF74BFF92FB27F81A79FAF80F" ID-CoL="GFQQ" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[1278,1434,1971,1992]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="reductaspina">N. reductaspina</taxonomicName>
. .
</emphasis> </emphasis>
4. P3 enp-2 with two distal elements........................................................................5. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF3BFF3179FDFF6A" blockId="46.[151,1438,151,1699]" box="[226,1437,151,173]" pageId="46" pageNumber="47">P3 enp-2 with one distal element.........................................................................7.</paragraph> </keyStep>
<paragraph id="EA68776EF748FF91FF4EFF1C79FDFF2F" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <keyStep id="51236FCBF74BFF91FF4EF87979FDFF6A" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<paragraph id="EA68776EF74BFF92FF4EF87979FCF821" blockId="45.[151,1437,1824,2022]" box="[151,1436,2000,2022]" pageId="45" pageNumber="46">
<keyLead id="5126D45BF74BFF92FF4EF87979FCF821" box="[151,1436,2000,2022]" pageId="45" pageNumber="46">4. P3 enp-2 with two distal elements........................................................................5.</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFF3179FDFF6A" blockId="46.[151,1438,151,1699]" box="[226,1437,151,173]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFF3179FDFF6A" box="[226,1437,151,173]" pageId="46" pageNumber="47">P3 enp-2 with one distal element.........................................................................7.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFF1C79FDFF2F" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFF1C79FDFF0D" blockId="46.[151,1438,151,1699]" box="[151,1437,181,203]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFF1C79FDFF0D" box="[151,1437,181,203]" pageId="46" pageNumber="47">
5. P1 exp-3 with three setae/spines.................................................................... 5. P1 exp-3 with three setae/spines....................................................................
<emphasis id="D8A3AB7CF748FF91FA91FF1C79FDFF0D" box="[1352,1437,181,202]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA91FF1C79FDFF0D" box="[1352,1437,181,202]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FA91FF1C79FAFF0D" ID-CoL="46KMK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Karanovic" baseAuthorityYear="2000" box="[1352,1434,181,202]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="huysi">N. huysi</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FA91FF1C79FAFF0D" ID-CoL="46KMK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Karanovic" baseAuthorityYear="2000" box="[1352,1434,181,202]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="huysi">N. huysi</taxonomicName>
. .
</emphasis> </emphasis>
P1 exp-3 with four setae/spines..........................................................................6. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFF4679FDFE9B" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <paragraph id="EA68776EF748FF91FF3BFF7B79FDFF2F" blockId="46.[151,1438,151,1699]" box="[226,1437,210,232]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFF7B79FDFF2F" box="[226,1437,210,232]" pageId="46" pageNumber="47">P1 exp-3 with four setae/spines..........................................................................6.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFF4679FDFE9B" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFF4679FDFEE5" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFF4679FDFEE5" pageId="46" pageNumber="47">
6. P1 endopod clearly longer than exopod; P3 enp-2 about half the length of enp-1; P5 of both sexes with one seta and two short spines between inner spinous process and outer basal seta............................................ 6. P1 endopod clearly longer than exopod; P3 enp-2 about half the length of enp-1; P5 of both sexes with one seta and two short spines between inner spinous process and outer basal seta............................................
<emphasis id="D8A3AB7CF748FF91FAFDFEA479FDFEE5" box="[1316,1437,269,290]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FAFDFEA479FDFEE5" box="[1316,1437,269,290]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FAFDFEA479FAFEE5" ID-CoL="73VZK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1316,1434,269,290]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="acanthus">N. acanthus</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FAFDFEA479FAFEE5" ID-CoL="73VZK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1316,1434,269,290]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="acanthus">N. acanthus</taxonomicName>
. .
</emphasis> </emphasis>
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFE8379FDFE9B" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFE8379FDFE9B" pageId="46" pageNumber="47">
P1 endopod as long as exopod; P3 enp-2 only slightly shorter than enp-1; P5 of both sexes with one seta and one spur between inner spinous process and outer basal seta...................................................... P1 endopod as long as exopod; P3 enp-2 only slightly shorter than enp-1; P5 of both sexes with one seta and one spur between inner spinous process and outer basal seta......................................................
<emphasis id="D8A3AB7CF748FF91FADCFEE179FDFE9B" box="[1285,1437,327,349]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FADCFEE179FDFE9B" box="[1285,1437,327,349]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FADCFEE179FAFE9B" ID-CoL="46KMG" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" box="[1285,1434,327,349]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="chaufriassei">N. chaufriassei</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FADCFEE179FAFE9B" ID-CoL="46KMG" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" box="[1285,1434,327,349]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="chaufriassei">N. chaufriassei</taxonomicName>
. .
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFECC79FCFE50" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> </keyStep>
<keyStep id="51236FCBF748FF91FF4EFECC79FCFE50" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFECC79FCFEBD" blockId="46.[151,1438,151,1699]" box="[151,1436,357,379]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFECC79FCFEBD" box="[151,1436,357,379]" pageId="46" pageNumber="47">
7. Sternal plates of somites bearing P1–P4 with midventral hook-like processes..................... 7. Sternal plates of somites bearing P1–P4 with midventral hook-like processes.....................
<taxonomicName id="2DD70CEDF748FF91FB1EFECC792AFEBD" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1223,1354,357,378]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="emendatus" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FB1EFECC792AFEBD" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1223,1354,357,378]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="emendatus" status="sp. nov.">
<emphasis id="D8A3AB7CF748FF91FB1EFECC792AFEBD" box="[1223,1354,357,378]" italics="true" pageId="46" pageNumber="47">N. emendatus</emphasis> <emphasis id="D8A3AB7CF748FF91FB1EFECC792AFEBD" box="[1223,1354,357,378]" italics="true" pageId="46" pageNumber="47">N. emendatus</emphasis>
@ -145,10 +195,23 @@ P1 endopod as long as exopod; P3 enp-2 only slightly shorter than enp-1; P5 of b
<emphasis id="D8A3AB7CF748FF91FA89FECC79FCFEBD" bold="true" box="[1360,1436,357,378]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FECC79FCFEBD" bold="true" box="[1360,1436,357,378]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FECC79FCFEBD" box="[1360,1436,357,378]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FECC79FCFEBD" box="[1360,1436,357,378]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
Sternal plates of somites bearing P1–P4 without such posteriorly directed processes................................8. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFE0979FDFE15" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">8. Paired spinous processes on anal somite straight and backwardly directed.........................................9. Paired spinous processes on anal somite dorsally recurved....................................................10.</paragraph> <paragraph id="EA68776EF748FF91FF3BFE2B79FCFE50" blockId="46.[151,1438,151,1699]" box="[226,1436,386,408]" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFE7379FCFD80" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <keyLead id="5126D45BF748FF91FF3BFE2B79FCFE50" box="[226,1436,386,408]" pageId="46" pageNumber="47">Sternal plates of somites bearing P1–P4 without such posteriorly directed processes................................8.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFE0979FDFE15" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFE0979FDFE72" blockId="46.[151,1438,151,1699]" box="[151,1437,415,437]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFE0979FDFE72" box="[151,1437,415,437]" pageId="46" pageNumber="47">8. Paired spinous processes on anal somite straight and backwardly directed.........................................9.</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFE1479FDFE15" blockId="46.[151,1438,151,1699]" box="[226,1437,445,467]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFE1479FDFE15" box="[226,1437,445,467]" pageId="46" pageNumber="47">Paired spinous processes on anal somite dorsally recurved....................................................10.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFE7379FCFD80" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFE7379FCFDCA" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFE7379FCFDCA" pageId="46" pageNumber="47">
9. P1 endopod 1.15 times as long as exopod; P5 elongate in both sexes, length/maximum width ratio 3.0 ( 9. P1 endopod 1.15 times as long as exopod; P5 elongate in both sexes, length/maximum width ratio 3.0 (
<specimenCount id="FCD1BCE7F748FF91FB27FE73796EFE28" box="[1278,1294,474,495]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FB27FE73796EFE28" box="[1278,1294,474,495]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount>
) and 3.5 ( ) and 3.5 (
@ -157,7 +220,12 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc
<taxonomicName id="2DD70CEDF748FF91FACDFE5179F6FDCB" ID-CoL="GFQG" authorityName="Chappuis" authorityYear="1955" box="[1300,1430,503,525]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="longiremis"> <taxonomicName id="2DD70CEDF748FF91FACDFE5179F6FDCB" ID-CoL="GFQG" authorityName="Chappuis" authorityYear="1955" box="[1300,1430,503,525]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF748FF91FACDFE5179F6FDCB" box="[1300,1430,503,525]" italics="true" pageId="46" pageNumber="47">N. longiremis</emphasis> <emphasis id="D8A3AB7CF748FF91FACDFE5179F6FDCB" box="[1300,1430,503,525]" italics="true" pageId="46" pageNumber="47">N. longiremis</emphasis>
</taxonomicName> </taxonomicName>
. P1 endopod 1.30 times as long as exopod; P5 much shorter in both sexes, length/maximum width ratio 1.7 ( .
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFDBC79FCFD80" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFDBC79FCFD80" pageId="46" pageNumber="47">
P1 endopod 1.30 times as long as exopod; P5 much shorter in both sexes, length/maximum width ratio 1.7 (
<specimenCount id="FCD1BCE7F748FF91FB22FDBC796BFDED" box="[1275,1291,533,554]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FB22FDBC796BFDED" box="[1275,1291,533,554]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount>
) and 2.25 ( ) and 2.25 (
<specimenCount id="FCD1BCE7F748FF91FAA5FDBC79F0FDED" box="[1404,1424,533,554]" count="1" pageId="46" pageNumber="47" type="male">♂</specimenCount> <specimenCount id="FCD1BCE7F748FF91FAA5FDBC79F0FDED" box="[1404,1424,533,554]" count="1" pageId="46" pageNumber="47" type="male">♂</specimenCount>
@ -168,8 +236,12 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc
<emphasis id="D8A3AB7CF748FF91FA89FD9B79FCFD80" bold="true" box="[1360,1436,562,583]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FD9B79FCFD80" bold="true" box="[1360,1436,562,583]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FD9B79FCFD80" box="[1360,1436,562,583]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FD9B79FCFD80" box="[1360,1436,562,583]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFDF979FDFD45" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> </keyStep>
<keyStep id="51236FCBF748FF91FF4EFDF979FDFD45" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFDF979FCFDA2" blockId="46.[151,1438,151,1699]" box="[151,1436,591,613]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFDF979FCFDA2" box="[151,1436,591,613]" pageId="46" pageNumber="47">
10. P1 endopod shorter than exopod........................................................ 10. P1 endopod shorter than exopod........................................................
<taxonomicName id="2DD70CEDF748FF91FB64FDF97929FDA3" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1213,1353,591,613]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="abbreviatus" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FB64FDF97929FDA3" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1213,1353,591,613]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="abbreviatus" status="sp. nov.">
<emphasis id="D8A3AB7CF748FF91FB64FDF97929FDA3" box="[1213,1353,591,613]" italics="true" pageId="46" pageNumber="47">N. abbreviatus</emphasis> <emphasis id="D8A3AB7CF748FF91FB64FDF97929FDA3" box="[1213,1353,591,613]" italics="true" pageId="46" pageNumber="47">N. abbreviatus</emphasis>
@ -177,9 +249,15 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc
<emphasis id="D8A3AB7CF748FF91FA89FDF979FCFDA2" bold="true" box="[1360,1436,592,613]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FDF979FCFDA2" bold="true" box="[1360,1436,592,613]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FDF979FCFDA2" box="[1360,1436,592,613]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FDF979FCFDA2" box="[1360,1436,592,613]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
P1 endopod at least as long as exopod....................................................................11. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFD2379FDFD1D" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <paragraph id="EA68776EF748FF91FF3BFDC479FDFD45" blockId="46.[151,1438,151,1699]" box="[226,1437,621,643]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFDC479FDFD45" box="[226,1437,621,643]" pageId="46" pageNumber="47">P1 endopod at least as long as exopod....................................................................11.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFD2379FDFD1D" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFD2379FCFD7A" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFD2379FCFD7A" pageId="46" pageNumber="47">
11. P1 endopod as long as exopod; spinous process on caudal ramus relatively short, about 35% of ramus length.................................................................................................. 11. P1 endopod as long as exopod; spinous process on caudal ramus relatively short, about 35% of ramus length..................................................................................................
<taxonomicName id="2DD70CEDF748FF91FB03FD01792AFD7B" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1242,1354,679,701]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="chilensis" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FB03FD01792AFD7B" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1242,1354,679,701]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="chilensis" status="sp. nov.">
<emphasis id="D8A3AB7CF748FF91FB03FD01792AFD7B" box="[1242,1354,679,701]" italics="true" pageId="46" pageNumber="47">N. chilensis</emphasis> <emphasis id="D8A3AB7CF748FF91FB03FD01792AFD7B" box="[1242,1354,679,701]" italics="true" pageId="46" pageNumber="47">N. chilensis</emphasis>
@ -187,28 +265,55 @@ P1 endopod at least as long as exopod...........................................
<emphasis id="D8A3AB7CF748FF91FA89FD0179FCFD7A" bold="true" box="[1360,1436,680,701]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FD0179FCFD7A" bold="true" box="[1360,1436,680,701]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FD0179FCFD7A" box="[1360,1436,680,701]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FD0179FCFD7A" box="[1360,1436,680,701]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
P1 endopod 1.25–1.30 times as long as exopod; spinous process on caudal ramus longer, about 45–55% of ramus length..12. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFD4B79FCFC97" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <paragraph id="EA68776EF748FF91FF3BFD6C79FDFD1D" blockId="46.[151,1438,151,1699]" box="[226,1437,709,731]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFD6C79FDFD1D" box="[226,1437,709,731]" pageId="46" pageNumber="47">P1 endopod 1.25–1.30 times as long as exopod; spinous process on caudal ramus longer, about 45–55% of ramus length..12.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFD4B79FCFC97" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFD4B79FCFCD2" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFD4B79FCFCD2" pageId="46" pageNumber="47">
12. P1 enp-1 1.25 times as long as enp-2; P 12. P1 enp-1 1.25 times as long as enp-2; P
<specimenCount id="FCD1BCE7F748FF91FDACFD4B7EFBFD3F" box="[629,667,738,760]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FDACFD4B7EFBFD3F" box="[629,667,738,760]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount>
twice as long as maximum width; terminal process of caudal ramus dorsally recurved.................................................................................... twice as long as maximum width; terminal process of caudal ramus dorsally recurved....................................................................................
<taxonomicName id="2DD70CEDF748FF91FAF4FCA979F6FCD2" ID-CoL="5W3DL" authorityName="Cottarelli, 1973 a sensu Mielke" authorityYear="1987" box="[1325,1430,768,789]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gussoae"> <taxonomicName id="2DD70CEDF748FF91FAF4FCA979F6FCD2" ID-CoL="5W3DL" authorityName="Cottarelli, 1973 a sensu Mielke" authorityYear="1987" box="[1325,1430,768,789]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gussoae">
<emphasis id="D8A3AB7CF748FF91FAF4FCA979F6FCD2" box="[1325,1430,768,789]" italics="true" pageId="46" pageNumber="47">N. gussoae</emphasis> <emphasis id="D8A3AB7CF748FF91FAF4FCA979F6FCD2" box="[1325,1430,768,789]" italics="true" pageId="46" pageNumber="47">N. gussoae</emphasis>
</taxonomicName> </taxonomicName>
. P1 enp-1 1.65 times as long as enp-2; P .
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFCB479FCFC97" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFCB479FCFC97" pageId="46" pageNumber="47">
P1 enp-1 1.65 times as long as enp-2; P
<specimenCount id="FCD1BCE7F748FF91FD80FCB47E1CFCF5" box="[601,636,797,819]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FD80FCB47E1CFCF5" box="[601,636,797,819]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount>
2.7 times as long as maximum width; terminal process of caudal ramus not dorsally recurved.................................................................................... 2.7 times as long as maximum width; terminal process of caudal ramus not dorsally recurved....................................................................................
<taxonomicName id="2DD70CEDF748FF91FAECFC9279F6FC88" ID-CoL="46KML" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Rao" baseAuthorityYear="1967" box="[1333,1430,826,848]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="indicus"> <taxonomicName id="2DD70CEDF748FF91FAECFC9279F6FC88" ID-CoL="46KML" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Rao" baseAuthorityYear="1967" box="[1333,1430,826,848]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="D8A3AB7CF748FF91FAECFC9279F6FC88" box="[1333,1430,826,848]" italics="true" pageId="46" pageNumber="47">N. indicus</emphasis> <emphasis id="D8A3AB7CF748FF91FAECFC9279F6FC88" box="[1333,1430,826,848]" italics="true" pageId="46" pageNumber="47">N. indicus</emphasis>
</taxonomicName> </taxonomicName>
. .
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFCF179FDFC4C" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">13. P1 exopod 2-segmented...............................................................................14. P1 exopod 3-segmented...............................................................................16.</paragraph> </keyStep>
<paragraph id="EA68776EF748FF91FF4EFC3B79FCFC02" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <keyStep id="51236FCBF748FF91FF4EFCF179FDFC4C" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFCF179FDFCAA" blockId="46.[151,1438,151,1699]" box="[151,1437,855,877]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFCF179FDFCAA" box="[151,1437,855,877]" pageId="46" pageNumber="47">13. P1 exopod 2-segmented...............................................................................14.</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFCDC79FDFC4C" blockId="46.[151,1438,151,1699]" box="[226,1437,885,907]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFCDC79FDFC4C" box="[226,1437,885,907]" pageId="46" pageNumber="47">P1 exopod 3-segmented...............................................................................16.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFC3B79FCFC02" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFC3B79FCFC6F" blockId="46.[151,1438,151,1699]" box="[151,1436,914,936]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFC3B79FCFC6F" box="[151,1436,914,936]" pageId="46" pageNumber="47">
14. P5 inner spine at least partly delimited at base; P 14. P5 inner spine at least partly delimited at base; P
<specimenCount id="FCD1BCE7F748FF91FD6FFC3B7EB9FC6F" box="[694,729,914,936]" count="6" pageId="46" pageNumber="47" type="male">6 ♂</specimenCount> <specimenCount id="FCD1BCE7F748FF91FD6FFC3B7EB9FC6F" box="[694,729,914,936]" count="6" pageId="46" pageNumber="47" type="male">6 ♂</specimenCount>
with one seta.............................................15. P5 inner spine fused at base, forming spinous process; P with one seta.............................................15.
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFC0679FCFC02" blockId="46.[151,1438,151,1699]" box="[226,1436,943,965]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFC0679FCFC02" box="[226,1436,943,965]" pageId="46" pageNumber="47">
P5 inner spine fused at base, forming spinous process; P
<specimenCount id="FCD1BCE7F748FF91FD27FC067F42FC02" box="[766,802,943,965]" count="6" pageId="46" pageNumber="47" type="male">6 ♂</specimenCount> <specimenCount id="FCD1BCE7F748FF91FD27FC067F42FC02" box="[766,802,943,965]" count="6" pageId="46" pageNumber="47" type="male">6 ♂</specimenCount>
with two elements.................. with two elements..................
<taxonomicName id="2DD70CEDF748FF91FB6AFC19792AFC02" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1203,1354,944,965]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FB6AFC19792AFC02" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1203,1354,944,965]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="supersetosus" status="sp. nov.">
@ -217,30 +322,56 @@ with two elements..................
<emphasis id="D8A3AB7CF748FF91FA89FC1979FCFC02" bold="true" box="[1360,1436,944,965]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FC1979FCFC02" bold="true" box="[1360,1436,944,965]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FC1979FCFC02" box="[1360,1436,944,965]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FC1979FCFC02" box="[1360,1436,944,965]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFC6479FCFB90" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> </keyStep>
<keyStep id="51236FCBF748FF91FF4EFC6479FCFB90" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFC6479FCFBC7" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFC6479FCFBC7" pageId="46" pageNumber="47">
15. P2 exp-2 outer spine shorter than exopod; P2 endopod shorter than exp-1, flask-shaped; P4 exp-3 four times as long as maximum width, with inner seta arising from distal fifth of inner margin......................................... 15. P2 exp-2 outer spine shorter than exopod; P2 endopod shorter than exp-1, flask-shaped; P4 exp-3 four times as long as maximum width, with inner seta arising from distal fifth of inner margin.........................................
<taxonomicName id="2DD70CEDF748FF91FAC7FC4279F6FBC7" ID-CoL="46KMX" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[1310,1430,1003,1024]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="trisetosus"> <taxonomicName id="2DD70CEDF748FF91FAC7FC4279F6FBC7" ID-CoL="46KMX" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[1310,1430,1003,1024]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF748FF91FAC7FC4279F6FBC7" box="[1310,1430,1003,1024]" italics="true" pageId="46" pageNumber="47">N. trisetosus</emphasis> <emphasis id="D8A3AB7CF748FF91FAC7FC4279F6FBC7" box="[1310,1430,1003,1024]" italics="true" pageId="46" pageNumber="47">N. trisetosus</emphasis>
</taxonomicName> </taxonomicName>
. P2 exp-2 outer spine distinctly longer than exopod; P2 endopod as long as exp-1, subrectangular and slightly tapering in distal third; P4 exp-3 2.75 times as long as maximum width, with inner seta arising from middle third of inner margin............................................................................................ .
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFBA179FCFB90" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFBA179FCFB90" pageId="46" pageNumber="47">
P2 exp-2 outer spine distinctly longer than exopod; P2 endopod as long as exp-1, subrectangular and slightly tapering in distal third; P4 exp-3 2.75 times as long as maximum width, with inner seta arising from middle third of inner margin............................................................................................
<taxonomicName id="2DD70CEDF748FF91FB65FBEA7929FB9F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1212,1353,1091,1112]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FB65FBEA7929FB9F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1212,1353,1091,1112]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="D8A3AB7CF748FF91FB65FBEA7929FB9F" box="[1212,1353,1091,1112]" italics="true" pageId="46" pageNumber="47">N. panamensis</emphasis> <emphasis id="D8A3AB7CF748FF91FB65FBEA7929FB9F" box="[1212,1353,1091,1112]" italics="true" pageId="46" pageNumber="47">N. panamensis</emphasis>
</taxonomicName> </taxonomicName>
<emphasis id="D8A3AB7CF748FF91FA89FBEB79FCFB90" bold="true" box="[1360,1436,1090,1111]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89FBEB79FCFB90" bold="true" box="[1360,1436,1090,1111]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89FBEB79FCFB90" box="[1360,1436,1090,1111]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89FBEB79FCFB90" box="[1360,1436,1090,1111]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFBF679FDFB55" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">16. P3 endopod 1-segmented, with one distal element..........................................................17. P3 endopod 2-segmented, enp-2 with two elements (outer one fused at base).....................................19.</paragraph> </keyStep>
<paragraph id="EA68776EF748FF91FF4EFB3379FDFB0A" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <keyStep id="51236FCBF748FF91FF4EFBF679FDFB55" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFBF679FDFBB2" blockId="46.[151,1438,151,1699]" box="[151,1437,1119,1141]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFBF679FDFBB2" box="[151,1437,1119,1141]" pageId="46" pageNumber="47">16. P3 endopod 1-segmented, with one distal element..........................................................17.</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFBD479FDFB55" blockId="46.[151,1438,151,1699]" box="[226,1437,1149,1171]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFBD479FDFB55" box="[226,1437,1149,1171]" pageId="46" pageNumber="47">P3 endopod 2-segmented, enp-2 with two elements (outer one fused at base).....................................19.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFB3379FDFB0A" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFB3379FDFB68" blockId="46.[151,1438,151,1699]" box="[151,1437,1178,1200]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFB3379FDFB68" box="[151,1437,1178,1200]" pageId="46" pageNumber="47">
17. P1 exp-3 with four elements; P2 endopod 2-segmented; P4 exp-3 without inner seta....................... 17. P1 exp-3 with four elements; P2 endopod 2-segmented; P4 exp-3 without inner seta.......................
<emphasis id="D8A3AB7CF748FF91FAFFFB3279FDFB68" box="[1318,1437,1178,1200]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FAFFFB3279FDFB68" box="[1318,1437,1178,1200]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FAFFFB3279F9FB68" ID-CoL="747ZK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1953" box="[1318,1433,1178,1200]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="australis">N. australis</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FAFFFB3279F9FB68" ID-CoL="747ZK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1953" box="[1318,1433,1178,1200]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="australis">N. australis</taxonomicName>
. .
</emphasis> </emphasis>
P1 exp-3 with three elements; P2 endopod 1-segmented; P4 exp-3 with inner seta.................................18. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFB7C79FDFAE2" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <paragraph id="EA68776EF748FF91FF3BFB1179FDFB0A" blockId="46.[151,1438,151,1699]" box="[226,1437,1207,1229]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFB1179FDFB0A" box="[226,1437,1207,1229]" pageId="46" pageNumber="47">P1 exp-3 with three elements; P2 endopod 1-segmented; P4 exp-3 with inner seta.................................18.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFB7C79FDFAE2" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFB7C79FCFAC0" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFB7C79FCFAC0" pageId="46" pageNumber="47">
18. Inner spinous process of P 18. Inner spinous process of P
<specimenCount id="FCD1BCE7F748FF91FE2BFB7C7E7AFB2D" box="[498,538,1237,1259]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FE2BFB7C7E7AFB2D" box="[498,538,1237,1259]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount>
distinctly curved outwardly (unknown in distinctly curved outwardly (unknown in
@ -252,6 +383,10 @@ distinctly curved outwardly (unknown in
<emphasis id="D8A3AB7CF748FF91FAF4FB5B79FCFAC0" bold="true" box="[1325,1436,1266,1287]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FAF4FB5B79FCFAC0" bold="true" box="[1325,1436,1266,1287]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FAF4FB5B79FCFAC0" box="[1325,1436,1266,1287]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FAF4FB5B79FCFAC0" box="[1325,1436,1266,1287]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFAB979FDFAE2" blockId="46.[151,1438,151,1699]" box="[226,1437,1295,1317]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFAB979FDFAE2" box="[226,1437,1295,1317]" pageId="46" pageNumber="47">
Inner spinous process of P Inner spinous process of P
<specimenCount id="FCD1BCE7F748FF91FE39FAA67D9FFAE2" box="[480,511,1295,1317]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount> <specimenCount id="FCD1BCE7F748FF91FE39FAA67D9FFAE2" box="[480,511,1295,1317]" count="5" pageId="46" pageNumber="47" type="female">5 ♀</specimenCount>
/ /
@ -261,41 +396,77 @@ and terminal process of caudal ramus straight...........................
<taxonomicName id="2DD70CEDF748FF91FAF9FAB979FAFAE2" ID-CoL="46KMC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[1312,1434,1296,1317]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="africanus">N. africanus</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FAF9FAB979FAFAE2" ID-CoL="46KMC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[1312,1434,1296,1317]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="africanus">N. africanus</taxonomicName>
. .
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFA8479FDFA98" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> </keyStep>
<keyStep id="51236FCBF748FF91FF4EFA8479FDFA98" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFA8479FEFA85" blockId="46.[151,1438,151,1699]" box="[151,1438,1325,1347]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFA8479FEFA85" box="[151,1438,1325,1347]" pageId="46" pageNumber="47">
19. P4 exp-3 without inner seta.................................................................... 19. P4 exp-3 without inner seta....................................................................
<emphasis id="D8A3AB7CF748FF91FAFDFA8479FEFA85" box="[1316,1438,1325,1346]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FAFDFA8479FEFA85" box="[1316,1438,1325,1346]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FAFDFA8479FAFA85" ID-CoL="46KMQ" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1316,1434,1325,1346]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pacificus">N. pacificus</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FAFDFA8479FAFA85" ID-CoL="46KMQ" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1316,1434,1325,1346]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pacificus">N. pacificus</taxonomicName>
. .
</emphasis> </emphasis>
P4 exp-3 with inner seta...............................................................................20. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFAC179FDFA70" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <paragraph id="EA68776EF748FF91FF3BFAE379FDFA98" blockId="46.[151,1438,151,1699]" box="[226,1437,1354,1376]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFAE379FDFA98" box="[226,1437,1354,1376]" pageId="46" pageNumber="47">P4 exp-3 with inner seta...............................................................................20.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFAC179FDFA70" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFAC179FDFA5D" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFAC179FDFA5D" pageId="46" pageNumber="47">
21. Urosome (except anal somite) with distinct surface ornamentation consisting of elongate rectangular plates............................................................................. 21. Urosome (except anal somite) with distinct surface ornamentation consisting of elongate rectangular plates.............................................................................
<emphasis id="D8A3AB7CF748FF91FAF7FA2C79FDFA5D" box="[1326,1437,1413,1434]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FAF7FA2C79FDFA5D" box="[1326,1437,1413,1434]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FAF7FA2C79F9FA5D" ID-CoL="46KMH" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1326,1433,1413,1434]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="clasingi">N. clasingi</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FAF7FA2C79F9FA5D" ID-CoL="46KMH" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1326,1433,1413,1434]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="clasingi">N. clasingi</taxonomicName>
. .
</emphasis> </emphasis>
Urosome without conspicuous surface ornamentation........................................................22. </keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EFA6979FDFA35" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">22. Caudal ramus with dorsolateral spur near base of seta VII....................................................23. Caudal ramus without dorsolateral spur near base of seta VII..................................................24.</paragraph> <paragraph id="EA68776EF748FF91FF3BFA0A79FDFA70" blockId="46.[151,1438,151,1699]" box="[226,1437,1442,1464]" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFA5379FCF9AF" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> <keyLead id="5126D45BF748FF91FF3BFA0A79FDFA70" box="[226,1437,1442,1464]" pageId="46" pageNumber="47">Urosome without conspicuous surface ornamentation........................................................22.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFA6979FDFA35" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFA6979FDFA12" blockId="46.[151,1438,151,1699]" box="[151,1437,1471,1493]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFA6979FDFA12" box="[151,1437,1471,1493]" pageId="46" pageNumber="47">22. Caudal ramus with dorsolateral spur near base of seta VII....................................................23.</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BFA7479FDFA35" blockId="46.[151,1438,151,1699]" box="[226,1437,1501,1523]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BFA7479FDFA35" box="[226,1437,1501,1523]" pageId="46" pageNumber="47">Caudal ramus without dorsolateral spur near base of seta VII..................................................24.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF748FF91FF4EFA5379FCF9AF" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EFA5379FCF9EA" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EFA5379FCF9EA" pageId="46" pageNumber="47">
23. Lappets of abdominal hyaline frills semi-incised obtusidigitate; inner seta of P2–P3 enp-2 longer than endopod; P5 three times as long as wide, with naked spinous process........................................................ 23. Lappets of abdominal hyaline frills semi-incised obtusidigitate; inner seta of P2–P3 enp-2 longer than endopod; P5 three times as long as wide, with naked spinous process........................................................
<taxonomicName id="2DD70CEDF748FF91FAF5F9B179F7F9EA" ID-CoL="46KMV" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1324,1431,1560,1581]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spicatus"> <taxonomicName id="2DD70CEDF748FF91FAF5F9B179F7F9EA" ID-CoL="46KMV" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1324,1431,1560,1581]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spicatus">
<emphasis id="D8A3AB7CF748FF91FAF5F9B179F7F9EA" box="[1324,1431,1560,1581]" italics="true" pageId="46" pageNumber="47">N. spicatus</emphasis> <emphasis id="D8A3AB7CF748FF91FAF5F9B179F7F9EA" box="[1324,1431,1560,1581]" italics="true" pageId="46" pageNumber="47">N. spicatus</emphasis>
</taxonomicName> </taxonomicName>
. Lappets of abdominal hyaline frills denticulate; inner seta of P2–P3 enp-2 shorter than endopod; P5 about 2.5 times as long as wide, with pinnate spinous process.......................................................... .
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BF99C79FCF9AF" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BF99C79FCF9AF" pageId="46" pageNumber="47">
Lappets of abdominal hyaline frills denticulate; inner seta of P2–P3 enp-2 shorter than endopod; P5 about 2.5 times as long as wide, with pinnate spinous process..........................................................
<taxonomicName id="2DD70CEDF748FF91FB2CF9FA79F6F9A0" ID-CoL="7WX6G" authorityName="Nicholls" authorityYear="1945" box="[1269,1430,1618,1640]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spinicaudatus"> <taxonomicName id="2DD70CEDF748FF91FB2CF9FA79F6F9A0" ID-CoL="7WX6G" authorityName="Nicholls" authorityYear="1945" box="[1269,1430,1618,1640]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF748FF91FB2CF9FA79F6F9A0" box="[1269,1430,1618,1640]" italics="true" pageId="46" pageNumber="47">N. spinicaudatus</emphasis> <emphasis id="D8A3AB7CF748FF91FB2CF9FA79F6F9A0" box="[1269,1430,1618,1640]" italics="true" pageId="46" pageNumber="47">N. spinicaudatus</emphasis>
</taxonomicName> </taxonomicName>
. .
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF748FF91FF4EF9D979FCF965" blockId="46.[151,1438,151,1699]" pageId="46" pageNumber="47"> </keyStep>
<keyStep id="51236FCBF748FF91FF4EF9D979FCF965" pageId="46" pageNumber="47">
<paragraph id="EA68776EF748FF91FF4EF9D979FDF943" blockId="46.[151,1438,151,1699]" box="[151,1437,1647,1669]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF4EF9D979FDF943" box="[151,1437,1647,1669]" pageId="46" pageNumber="47">
24. P1 endopod distinctly longer than exopod...................................................... 24. P1 endopod distinctly longer than exopod......................................................
<emphasis id="D8A3AB7CF748FF91FADDF9D979FDF943" box="[1284,1437,1647,1669]" italics="true" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FADDF9D979FDF943" box="[1284,1437,1647,1669]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="2DD70CEDF748FF91FADDF9D979FAF943" ID-CoL="46KMM" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Ito" baseAuthorityYear="1968" box="[1284,1434,1647,1669]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ishikarianus">N. ishikarianus</taxonomicName> <taxonomicName id="2DD70CEDF748FF91FADDF9D979FAF943" ID-CoL="46KMM" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Ito" baseAuthorityYear="1968" box="[1284,1434,1647,1669]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ishikarianus">N. ishikarianus</taxonomicName>
. .
</emphasis> </emphasis>
</keyLead>
</paragraph>
<paragraph id="EA68776EF748FF91FF3BF92479FCF965" blockId="46.[151,1438,151,1699]" box="[226,1436,1677,1699]" pageId="46" pageNumber="47">
<keyLead id="5126D45BF748FF91FF3BF92479FCF965" box="[226,1436,1677,1699]" pageId="46" pageNumber="47">
P1 rami equally long............................................................ P1 rami equally long............................................................
<taxonomicName id="2DD70CEDF748FF91FBA6F924792AF965" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1151,1354,1677,1698]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudishikarianus" status="sp. nov."> <taxonomicName id="2DD70CEDF748FF91FBA6F924792AF965" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1151,1354,1677,1698]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pseudishikarianus" status="sp. nov.">
<emphasis id="D8A3AB7CF748FF91FBA6F924792AF965" box="[1151,1354,1677,1698]" italics="true" pageId="46" pageNumber="47">N. pseudishikarianus</emphasis> <emphasis id="D8A3AB7CF748FF91FBA6F924792AF965" box="[1151,1354,1677,1698]" italics="true" pageId="46" pageNumber="47">N. pseudishikarianus</emphasis>
@ -303,7 +474,10 @@ P1 rami equally long............................................................
<emphasis id="D8A3AB7CF748FF91FA89F92479FCF965" bold="true" box="[1360,1436,1677,1698]" pageId="46" pageNumber="47"> <emphasis id="D8A3AB7CF748FF91FA89F92479FCF965" bold="true" box="[1360,1436,1677,1698]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3901607F748FF91FA89F92479FCF965" box="[1360,1436,1677,1698]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F748FF91FA89F92479FCF965" box="[1360,1436,1677,1698]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
</keyStep>
</key>
</subSubSection> </subSubSection>
</treatment> </treatment>
</document> </document>

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<mods:title id="B6939D857C86D3D7B03422F71DB89638">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
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<treatment id="627EC678F74DFF94FF4EFD927858F97B" LSID="urn:lsid:plazi:treatment:627EC678F74DFF94FF4EFD927858F97B" httpUri="http://treatment.plazi.org/id/627EC678F74DFF94FF4EFD927858F97B" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="A2CD24E5F74DFF94FF4EFD927E19FD91" box="[151,633,571,598]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="EA68776EF74DFF94FF4EFD927E19FD91" blockId="43.[151,633,571,598]" box="[151,633,571,598]" pageId="43" pageNumber="44">
<heading id="B120C002F74DFF94FF4EFD927E19FD91" bold="true" box="[151,633,571,598]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="4">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD927E19FD91" ID-CoL="46KMB" authority="(Lang, 1965)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[151,633,571,598]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF4EFD927E19FD91" bold="true" box="[151,633,571,598]" pageId="43" pageNumber="44">
<emphasis id="D8A3AB7CF74DFF94FF4EFD927DB4FD91" bold="true" box="[151,468,571,598]" italics="true" pageId="43" pageNumber="44">Neoleptastacus accraensis</emphasis>
(
<bibRefCitation id="8E460A9FF74DFF94FE3CFD957E11FD91" author="Lang, K." box="[485,625,572,598]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F74DFF94FF4EFD28788DFC9C" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="EA68776EF74DFF94FF4EFD287FD0FD5F" blockId="43.[151,1091,640,787]" box="[151,944,640,665]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="CAC75040F74DFF94FF4EFD287FD0FD5F" box="[151,944,640,665]" pageId="43" pageNumber="44">
<treatmentCitation id="6B76517FF74DFF94FF4EFD287FD0FD5F" author="Chappuis, P. - A. &amp; Rouch, R." box="[151,944,640,665]" page=")" pageId="43" pageNumber="44" year="1955">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD287FD0FD5F" ID-CoL="GFQG" authority="Chappuis, 1955 sensu Chappuis &amp; Rouch (1961: 608)" authorityName="Chappuis, 1955 sensu Chappuis &amp; Rouch" authorityPageNumber="608" authorityYear="1961" box="[151,944,640,665]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF74DFF94FF4EFD287DE6FD5F" box="[151,390,640,664]" italics="true" pageId="43" pageNumber="44">Arenopontia longiremis</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FE54FD297E4CFD5F" author="Chappuis P. - A." box="[397,556,640,664]" pageId="43" pageNumber="44" pagination="45 - 73" refId="ref45639" refString="Chappuis P. - A. (1955) Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar. IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut scientifique de Madagascar, Series A, 9, 45 - 73." type="journal article" year="1955">Chappuis, 1955</bibRefCitation>
<emphasis id="D8A3AB7CF74DFF94FDEBFD287E0AFD5F" box="[562,618,641,664]" italics="true" pageId="43" pageNumber="44">sensu</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FDA8FD297FD0FD5F" author="Chappuis, P. - A. &amp; Rouch, R." box="[625,944,640,665]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961: 608)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF74DFF94FF4EFD367E62FD70" blockId="43.[151,1091,640,787]" box="[151,514,671,696]" pageId="43" pageNumber="44">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD367E62FD70" ID-CoL="GFPZ" authority="Lang, 1965" authorityName="Lang" authorityYear="1965" box="[151,514,671,696]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF4EFD367DE7FD70" box="[151,391,671,695]" italics="true" pageId="43" pageNumber="44">Arenopontia accraensis</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FE57FD097E62FD70" author="Lang, K." box="[398,514,671,696]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="EA68776EF74DFF94FF4EFD177823FD11" blockId="43.[151,1091,640,787]" box="[151,1091,702,726]" pageId="43" pageNumber="44">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD177ED7FD11" ID-CoL="GFPZ" authority="Lang, 1965" authorityName="Lang" authorityYear="1965" box="[151,695,702,726]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF74DFF94FF4EFD177D75FD12" box="[151,277,702,725]" italics="true" pageId="43" pageNumber="44">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF74DFF94FEFDFD177DDFFD11" box="[292,447,702,726]" italics="true" pageId="43" pageNumber="44">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF74DFF94FE14FD177E59FD12" box="[461,569,702,725]" italics="true" pageId="43" pageNumber="44">accraensis</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FD99FD177ED7FD11" author="Lang, K." box="[576,695,702,726]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF74DFF94FD19FD177F0DFD11" author="Wells, J. B. J." box="[704,877,702,726]" page="324" pageId="43" pageNumber="44" year="1967">
<bibRefCitation id="8E460A9FF74DFF94FD19FD177F0DFD11" author="Wells, J. B. J." box="[704,877,702,726]" pageId="43" pageNumber="44" pagination="189 - 358" refId="ref49859" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / S 0080456800024017" type="journal article" year="1967">Wells (1967: 324</bibRefCitation>
</treatmentCitation>
—spelled
<taxonomicName id="2DD70CEDF74DFF94FC0FFD17785BFD12" ID-CoL="GFPZ" authorityName="Lang" authorityYear="1965" box="[982,1083,702,725]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FC0FFD17785BFD12" box="[982,1083,702,725]" italics="true" pageId="43" pageNumber="44">accraensis</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="EA68776EF74DFF94FF4EFD747FADFD33" blockId="43.[151,1091,640,787]" box="[151,973,732,757]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="CAC75040F74DFF94FF4EFD747FADFD33" box="[151,973,732,757]" pageId="43" pageNumber="44">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD747F6FFD33" ID-CoL="GFPZ" authority="Lang, 1965" authorityName="Lang" authorityYear="1965" box="[151,783,732,757]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="acantha" subGenus="Neoleptastacus" subSpecies="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF4EFD747D75FD33" box="[151,277,733,756]" italics="true" pageId="43" pageNumber="44">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF74DFF94FEFDFD757DDFFD33" box="[292,447,732,756]" italics="true" pageId="43" pageNumber="44">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF74DFF94FE14FD757EF1FD33" box="[461,657,732,756]" italics="true" pageId="43" pageNumber="44">acantha accraensis</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FD41FD747F6FFD33" author="Lang, K." box="[664,783,732,757]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF74DFF94FCC0FD747FADFD33" author="Kunz, H." box="[793,973,732,757]" page="356" pageId="43" pageNumber="44" year="1971">
<bibRefCitation id="8E460A9FF74DFF94FCC0FD747FADFD33" author="Kunz, H." box="[793,973,732,757]" pageId="43" pageNumber="44" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971: 356)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF74DFF94FF4EFD527F72FCD4" blockId="43.[151,1091,640,787]" box="[151,786,763,787]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="CAC75040F74DFF94FF4EFD527F72FCD4" box="[151,786,763,787]" pageId="43" pageNumber="44">
<treatmentCitation id="6B76517FF74DFF94FF4EFD527F72FCD4" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[151,786,763,787]" page=")" pageId="43" pageNumber="44" year="1965">
<taxonomicName id="2DD70CEDF74DFF94FF4EFD527F72FCD4" ID-CoL="46KMB" authority="(Lang, 1965) Sak et al. (2008: 412)" authorityName="Sak et al." authorityPageNumber="412" authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[151,786,763,787]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF4EFD527DC4FCD4" box="[151,420,763,787]" italics="true" pageId="43" pageNumber="44">Neoleptastacus accraensis</emphasis>
(
<bibRefCitation id="8E460A9FF74DFF94FE6BFD527E48FCD4" author="Lang, K." box="[434,552,763,787]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
)
<bibRefCitation id="8E460A9FF74DFF94FDEFFD527F72FCD4" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[566,786,763,787]" pageId="43" pageNumber="44" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF74DFF94FDBBFD557EF5FCD4" box="[610,661,763,787]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
(2008: 412)
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF74DFF94FF4EFCE8788DFC9C" blockId="43.[151,1437,833,1724]" box="[151,1261,833,860]" pageId="43" pageNumber="44">
<emphasis id="D8A3AB7CF74DFF94FF4EFCE87DF0FC9C" bold="true" box="[151,400,833,859]" pageId="43" pageNumber="44">Original description.</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FE4EFCE87ED4FC9C" author="Chappuis, P. - A. &amp; Rouch, R." box="[407,692,833,859]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961</bibRefCitation>
—as
<taxonomicName id="2DD70CEDF74DFF94FD29FCEB7FE6FC9C" ID-CoL="GFQG" authorityName="Chappuis, 1955 sensu Chappuis &amp; Rouch" authorityYear="1961" box="[752,902,833,860]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF74DFF94FD29FCEB7FE6FC9C" box="[752,902,833,860]" italics="true" pageId="43" pageNumber="44">A. longiremis</emphasis>
</taxonomicName>
): 607–608;
<figureCitation id="72EC6BEBF74DFF94FBD6FCE8781EFC9C" box="[1039,1150,833,859]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStartId-0="38.[151,250,1943,1967]" captionStartId-1="39.[151,250,1928,1952]" captionStartId-2="40.[151,250,1943,1967]" captionStartId-3="48.[151,250,1943,1967]" captionStartId-4="49.[151,250,1943,1967]" captionTargetBox-0="[192,1396,191,1907]" captionTargetBox-1="[207,1380,189,1892]" captionTargetBox-2="[196,1382,193,1907]" captionTargetBox-3="[187,1397,193,1907]" captionTargetBox-4="[171,1412,190,1907]" captionTargetId-0="figure-17@38.[179,1408,181,1919]" captionTargetId-1="figure-17@39.[195,1392,181,1904]" captionTargetId-2="figure-17@40.[194,1394,181,1919]" captionTargetId-3="figure-17@48.[179,1409,181,1919]" captionTargetId-4="figure-17@49.[163,1424,181,1919]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionTargetPageId-3="48" captionTargetPageId-4="49" captionText-0="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." captionText-1="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." captionText-2="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." captionText-3="FIGURE 12. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, lateral; (B) habitus, dorsal; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]: (D) anal somite and right caudal ramus, lateral." captionText-4="FIGURE 13. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) antennule (♀), dorsal; (B) antenna (♀), medial [lateral endopodal spines indicated by arrows]; (C) antenna (♀), outer; (D) antennule (♂), anterior; (E) antennule (♂), dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.14021498" figureDoi-1="http://doi.org/10.5281/zenodo.14021500" figureDoi-2="http://doi.org/10.5281/zenodo.14021502" figureDoi-3="http://doi.org/10.5281/zenodo.14021504" figureDoi-4="http://doi.org/10.5281/zenodo.14021506" httpUri-0="https://zenodo.org/record/14021498/files/figure.png" httpUri-1="https://zenodo.org/record/14021500/files/figure.png" httpUri-2="https://zenodo.org/record/14021502/files/figure.png" httpUri-3="https://zenodo.org/record/14021504/files/figure.png" httpUri-4="https://zenodo.org/record/14021506/files/figure.png" pageId="43" pageNumber="44">Figs 9–13</figureCitation>
(
<specimenCount id="FCD1BCE7F74DFF94FB48FCE878C4FC9C" box="[1169,1188,833,859]" count="1" pageId="43" pageNumber="44" type="female">♀</specimenCount>
only).
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F74DFF94FF1EFCCC7DBBFC64" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="EA68776EF74DFF94FF1EFCCC78A4FCB8" blockId="43.[151,1437,833,1724]" box="[199,1220,869,896]" pageId="43" pageNumber="44">
<emphasis id="D8A3AB7CF74DFF94FF1EFCCC7D06FCB8" bold="true" box="[199,358,869,895]" pageId="43" pageNumber="44">
<typeStatus id="356CC9CCF74DFF94FF1EFCCC7D62FCB8" box="[199,258,869,895]" pageId="43" pageNumber="44">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="92C037FEF74DFF94FEB5FCCC7DD8FCB8" box="[364,440,869,895]" name="Ghana" pageId="43" pageNumber="44">Ghana</collectingCountry>
,
<collectingRegion id="2813B98CF74DFF94FE1AFCCC7EDBFCB8" box="[451,699,869,896]" country="Ghana" name="Greater Accra" pageId="43" pageNumber="44">Greater Accra Region</collectingRegion>
,
<location id="EF0821B5F74DFF94FD1DFCCF7F6AFC47" LSID="urn:lsid:plazi:treatment:627EC678F74DFF94FF4EFD927858F97B:EF0821B5F74DFF94FD1DFCCF7F6AFC47" box="[708,778,870,896]" country="Ghana" name="Accra" pageId="43" pageNumber="44" stateProvince="Greater Accra">Accra</location>
; sandy beach in front of the lighthouse.
</paragraph>
<paragraph id="EA68776EF74DFF94FF1EFC207DBBFC64" blockId="43.[151,1437,833,1724]" box="[199,475,905,931]" pageId="43" pageNumber="44">
<emphasis id="D8A3AB7CF74DFF94FF1EFC207D3EFC64" bold="true" box="[199,350,905,931]" pageId="43" pageNumber="44">Body length.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F74DFF94FF1EFC047858F97B" pageId="43" pageNumber="44" type="discussion">
<paragraph id="EA68776EF74DFF94FF1EFC0478D9FB04" blockId="43.[151,1437,833,1724]" pageId="43" pageNumber="44">
<emphasis id="D8A3AB7CF74DFF94FF1EFC047D5BFC00" bold="true" box="[199,315,941,967]" pageId="43" pageNumber="44">Remarks.</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FE98FC047EF9FC00" author="Chappuis, P. - A. &amp; Rouch, R." box="[321,665,941,967]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961: 608</bibRefCitation>
,
<figureCitation id="72EC6BEBF74DFF94FD7AFC047F72FC00" box="[675,786,941,967]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStartId-0="38.[151,250,1943,1967]" captionStartId-1="39.[151,250,1928,1952]" captionStartId-2="40.[151,250,1943,1967]" captionStartId-3="48.[151,250,1943,1967]" captionStartId-4="49.[151,250,1943,1967]" captionTargetBox-0="[192,1396,191,1907]" captionTargetBox-1="[207,1380,189,1892]" captionTargetBox-2="[196,1382,193,1907]" captionTargetBox-3="[187,1397,193,1907]" captionTargetBox-4="[171,1412,190,1907]" captionTargetId-0="figure-17@38.[179,1408,181,1919]" captionTargetId-1="figure-17@39.[195,1392,181,1904]" captionTargetId-2="figure-17@40.[194,1394,181,1919]" captionTargetId-3="figure-17@48.[179,1409,181,1919]" captionTargetId-4="figure-17@49.[163,1424,181,1919]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionTargetPageId-3="48" captionTargetPageId-4="49" captionText-0="FIGURE 9. Neoleptastacus supersetosus sp. nov. (♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior." captionText-1="FIGURE 10. Neoleptastacus supersetosus sp. nov.: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral." captionText-2="FIGURE 11. Neoleptastacus supersetosus sp. nov.: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal." captionText-3="FIGURE 12. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, lateral; (B) habitus, dorsal; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]: (D) anal somite and right caudal ramus, lateral." captionText-4="FIGURE 13. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) antennule (♀), dorsal; (B) antenna (♀), medial [lateral endopodal spines indicated by arrows]; (C) antenna (♀), outer; (D) antennule (♂), anterior; (E) antennule (♂), dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.14021498" figureDoi-1="http://doi.org/10.5281/zenodo.14021500" figureDoi-2="http://doi.org/10.5281/zenodo.14021502" figureDoi-3="http://doi.org/10.5281/zenodo.14021504" figureDoi-4="http://doi.org/10.5281/zenodo.14021506" httpUri-0="https://zenodo.org/record/14021498/files/figure.png" httpUri-1="https://zenodo.org/record/14021500/files/figure.png" httpUri-2="https://zenodo.org/record/14021502/files/figure.png" httpUri-3="https://zenodo.org/record/14021504/files/figure.png" httpUri-4="https://zenodo.org/record/14021506/files/figure.png" pageId="43" pageNumber="44">Figs 9–13</figureCitation>
) recorded
<specimenCount id="FCD1BCE7F74DFF94FC50FC077870FC00" box="[905,1040,941,968]" count="2" pageId="43" pageNumber="44" type="female">two females</specimenCount>
from a sandy beach in Accra which they assigned to
<taxonomicName id="2DD70CEDF74DFF94FE8BFC7B7F75FC2B" authority="Chappuis, 1955" authorityName="Chappuis" authorityYear="1955" box="[338,789,977,1004]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF74DFF94FE8BFC7B7E3BFC2C" box="[338,603,977,1004]" italics="true" pageId="43" pageNumber="44">Arenopontia longiremis</emphasis>
<bibRefCitation id="8E460A9FF74DFF94FDBBFC787F75FC2B" author="Chappuis P. - A." box="[610,789,977,1004]" pageId="43" pageNumber="44" pagination="45 - 73" refId="ref45639" refString="Chappuis P. - A. (1955) Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar. IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut scientifique de Madagascar, Series A, 9, 45 - 73." type="journal article" year="1955">Chappuis, 1955</bibRefCitation>
</taxonomicName>
. They considered the differences with the
<typeStatus id="356CC9CCF74DFF94FB34FC7B797DFC2B" box="[1261,1309,978,1004]" pageId="43" pageNumber="44">type</typeStatus>
population as insignificant, being a likely reflection of the disjunct distribution of the species.
<bibRefCitation id="8E460A9FF74DFF94FBFDFC5F78CEFBD7" author="Lang, K." box="[1060,1198,1013,1040]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang (1965)</bibRefCitation>
considered
<bibRefCitation id="8E460A9FF74DFF94FAEBFC5C7D01FBF4" author="Chappuis, P. - A. &amp; Rouch, R." pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch’s (1961)</bibRefCitation>
Ghanaian specimens to fall outside the limits of variability of
<taxonomicName id="2DD70CEDF74DFF94FBF9FBB378D6FBF4" authorityName="Chappuis" authorityYear="1955" box="[1056,1206,1049,1076]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF74DFF94FBF9FBB378D6FBF4" box="[1056,1206,1049,1076]" italics="true" pageId="43" pageNumber="44">A. longiremis</emphasis>
</taxonomicName>
and proposed a new name,
<taxonomicName id="2DD70CEDF74DFF94FF38FB977D18FB90" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[225,376,1086,1112]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF38FB977D18FB90" box="[225,376,1086,1112]" italics="true" pageId="43" pageNumber="44">A. accraensis</emphasis>
</taxonomicName>
, for them.
<bibRefCitation id="8E460A9FF74DFF94FE28FB977E1FFB90" author="Kunz, H." box="[497,639,1085,1112]" pageId="43" pageNumber="44" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
treated
<taxonomicName id="2DD70CEDF74DFF94FD01FB977F0EFB90" authorityName="Chappuis" authorityYear="1955" box="[728,878,1085,1112]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF74DFF94FD01FB977F0EFB90" box="[728,878,1085,1112]" italics="true" pageId="43" pageNumber="44">A. longiremis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF74DFF94FC7AFB977859FB90" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[931,1081,1086,1112]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FC7AFB977859FB90" box="[931,1081,1086,1112]" italics="true" pageId="43" pageNumber="44">A. accraensis</emphasis>
</taxonomicName>
as subspecies of
<taxonomicName id="2DD70CEDF74DFF94FB20FB977912FB90" authorityName="Chappuis" authorityYear="1954" box="[1273,1394,1085,1112]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF74DFF94FB20FB977912FB90" box="[1273,1394,1085,1112]" italics="true" pageId="43" pageNumber="44">A. acantha</emphasis>
</taxonomicName>
but this was not adopted by subsequent revisers [
<bibRefCitation id="8E460A9FF74DFF94FD5EFBCB7E8AFBBC" author="Ito, T." box="[647,746,1121,1148]" pageId="43" pageNumber="44" pagination="47 - 55" refId="ref47150" refString="Ito, T. (1978) A new species of marine interstitial harpacticoid copepod of the genus Arenopontia from the Bonin Islands, southern Japan. Annotationes zoologicae japonenses, 51, 47 - 55." type="journal article" year="1978">Itô 1978</bibRefCitation>
;
<bibRefCitation id="8E460A9FF74DFF94FD2CFBC87843FBBC" author="Bodiou, J. - Y. &amp; Colomines, J. - C." box="[757,1059,1121,1147]" pageId="43" pageNumber="44" pagination="55 - 64" refId="ref45366" refString="Bodiou, J. - Y. &amp; Colomines, J. - C. (1986) Harpacticoides (Copepodes) des Iles Crozet I. - Description d'une espece nouvelle du genre Arenopontia Kunz. Harpacticoids (Crustacea, Copepoda) from the Crozet Islands. Description of a new species of the genus Arenopontia Kunz. Vie et Milieu, 36, 55 - 64." type="journal article" year="1986">Bodiou &amp; Colomines 1986</bibRefCitation>
;
<bibRefCitation id="8E460A9FF74DFF94FBF4FBC878CDFBBC" author="Wells, J. B. J." box="[1069,1197,1121,1147]" pageId="43" pageNumber="44" pagination="1 - 872" refId="ref50074" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1 - 872. https: // doi. org / 10.11646 / zootaxa. 1568.1.1" type="journal article" year="2007">Wells 2007</bibRefCitation>
;
<bibRefCitation id="8E460A9FF74DFF94FB61FBC8793BFBBC" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[1208,1371,1121,1147]" pageId="43" pageNumber="44" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF74DFF94FB31FBCB797FFBBC" box="[1256,1311,1121,1147]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
2008
</bibRefCitation>
—but see
<bibRefCitation id="8E460A9FF74DFF94FF1CFB2C7D3DFB58" author="Bodin, P." box="[197,349,1157,1183]" pageId="43" pageNumber="44" pagination="1 - 304" refId="ref45314" refString="Bodin, P. (1997) Catalogue of the new marine harpacticoid copepods (1997 Edition). Studiedocumenten van het K. B. I. N. / Documents de Travail de l'I. R. Sc. N. B., 89, 1 - 304." type="journal article" year="1997">Bodin (1997)</bibRefCitation>
for a dissenting opinion]. The Angolan material identified by
<bibRefCitation id="8E460A9FF74DFF94FBF6FB2F78DFFB58" author="Kunz, H." box="[1071,1215,1157,1184]" pageId="43" pageNumber="44" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
as
<taxonomicName id="2DD70CEDF74DFF94FB33FB2F7D6FFB04" authorityName="Lang" authorityYear="1965" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="acantha" subGenus="Neoleptastacus" subSpecies="accraensis">
<emphasis id="D8A3AB7CF74DFF94FB33FB2F7963FB67" box="[1258,1283,1158,1184]" italics="true" pageId="43" pageNumber="44">A.</emphasis>
(
<emphasis id="D8A3AB7CF74DFF94FACCFB2F7950FB58" box="[1301,1328,1158,1183]" italics="true" pageId="43" pageNumber="44">N.</emphasis>
)
<emphasis id="D8A3AB7CF74DFF94FA9BFB2C7D6FFB04" italics="true" pageId="43" pageNumber="44">acantha accraensis</emphasis>
</taxonomicName>
belongs to a different species, described here as
<taxonomicName id="2DD70CEDF74DFF94FCEBFB037FAAFB04" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[818,970,1193,1219]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="emendatus" status="sp. nov.">
<emphasis id="D8A3AB7CF74DFF94FCEBFB037FAAFB04" box="[818,970,1193,1219]" italics="true" pageId="43" pageNumber="44">N. emendatus</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF74DFF94FC08FB00784BFB04" bold="true" box="[977,1067,1193,1219]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="C3901607F74DFF94FC08FB00784BFB04" box="[977,1067,1193,1219]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see above).
</paragraph>
<paragraph id="EA68776EF74DFF94FF1EFB6478C0F9C0" blockId="43.[151,1437,833,1724]" pageId="43" pageNumber="44">
Although
<bibRefCitation id="8E460A9FF74DFF94FEE0FB677DA7FB2F" author="Lang, K." box="[313,455,1229,1256]" pageId="43" pageNumber="44" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang (1965)</bibRefCitation>
,
<bibRefCitation id="8E460A9FF74DFF94FE0DFB647F77FB20" author="Bodiou, J. - Y. &amp; Colomines, J. - C." box="[468,791,1229,1255]" pageId="43" pageNumber="44" pagination="55 - 64" refId="ref45366" refString="Bodiou, J. - Y. &amp; Colomines, J. - C. (1986) Harpacticoides (Copepodes) des Iles Crozet I. - Description d'une espece nouvelle du genre Arenopontia Kunz. Harpacticoids (Crustacea, Copepoda) from the Crozet Islands. Description of a new species of the genus Arenopontia Kunz. Vie et Milieu, 36, 55 - 64." type="journal article" year="1986">Bodiou &amp; Colomines (1986)</bibRefCitation>
and
<bibRefCitation id="8E460A9FF74DFF94FC96FB647F81FB20" author="Wells, J. B. J." box="[847,993,1229,1255]" pageId="43" pageNumber="44" pagination="1 - 872" refId="ref50074" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1 - 872. https: // doi. org / 10.11646 / zootaxa. 1568.1.1" type="journal article" year="2007">Wells (2007)</bibRefCitation>
claim that P4 exp-3 possesses an inner seta, the setation of this leg has yet to be revealed. Similarly, in the identification keys provided by these authorities and
<bibRefCitation id="8E460A9FF74DFF94FF11FABC7DF0FAE8" author="Karanovic, T." box="[200,400,1301,1327]" pageId="43" pageNumber="44" pagination="33 - 38" refId="ref47348" refString="Karanovic, T. (2000) Arenopontia (Neoleptastacus) huysi, sp. nov. (Crustacea, Copepoda, Harpacticoida) from marine interstitial of Montenegro (S. E. Europe). Helgoland marine Research, 54, 33 - 38. https: // doi. org / 10.1007 / s 101520050033" type="journal article" year="2000">Karanovic (2000)</bibRefCitation>
it is assumed that the distal endopod segment of P3 has only one distal element. However,
<bibRefCitation id="8E460A9FF74DFF94FF4EFA907DAAFA94" author="Chappuis, P. - A. &amp; Rouch, R." box="[151,458,1337,1363]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
show a second short element on enp-
<quantity id="2D2FDA8BF74DFF94FC51FA907FD6FA94" box="[904,950,1337,1363]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="43" pageNumber="44" unit="in" value="2.0">2 in</quantity>
their illustration of the P3 endopod. This element is here interpreted as the outer distal spine using the spinule row on enp-1 as a reference point for correct orientation. Note also that
<bibRefCitation id="8E460A9FF74DFF94FE65FA287E92FA5C" author="Chappuis, P. - A. &amp; Rouch, R." box="[444,754,1409,1435]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch’s (1961)</bibRefCitation>
illustration of the P1 shows it in an unnatural twisted position and that the terminal spinous process on the caudal ramus was figured as an articulating spine. The latter condition is unlikely since the caudal process in all members of the
<taxonomicName id="2DD70CEDF74DFF94FCF4FA607FBBFA24" authorityName="Martinez Arbizu &amp; Moura" authorityYear="1994" box="[813,987,1481,1507]" class="Copepoda" family="Arenopontiidae" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="family">Arenopontiidae</taxonomicName>
represents a posterior outgrowth of the ramus and not the homologue of an armature element that became secondarily incorporated.
</paragraph>
<paragraph id="EA68776EF74DFF94FF1EF9B87858F97B" blockId="43.[151,1437,833,1724]" pageId="43" pageNumber="44">
<taxonomicName id="2DD70CEDF74DFF94FF1EF9B87D95F9EC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[199,501,1553,1579]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF74DFF94FF1EF9B87D95F9EC" box="[199,501,1553,1579]" italics="true" pageId="43" pageNumber="44">Neoleptastacus accraensis</emphasis>
</taxonomicName>
is in need of detailed redescription (including the as yet unknown male) and is treated here as a
<emphasis id="D8A3AB7CF74DFF94FE85F99F7E50F988" box="[348,560,1589,1615]" italics="true" pageId="43" pageNumber="44">species inquirenda</emphasis>
in the genus since it is practically unidentifiable based on the few drawings provided by
<bibRefCitation id="8E460A9FF74DFF94FEFFF9F07E35F9B4" author="Chappuis, P. - A. &amp; Rouch, R." box="[294,597,1625,1651]" pageId="43" pageNumber="44" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
. It is tentatively placed in the
<taxonomicName id="2DD70CEDF74DFF94FC73F9F07826F9B4" authorityName="Nicholls" authorityYear="1945" box="[938,1094,1625,1651]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF74DFF94FC73F9F07826F9B4" box="[938,1094,1625,1651]" italics="true" pageId="43" pageNumber="44">spinicaudatus</emphasis>
</taxonomicName>
-group based on the combined absence of paired anal processes and the retention of the plesiomorphic armature formula of P1–P4. The species is only known from the
<typeStatus id="356CC9CCF74DFF94FE53F90B7DDAF97B" box="[394,442,1698,1724]" pageId="43" pageNumber="44">type</typeStatus>
locality where it occurs sympatrically with
<taxonomicName id="2DD70CEDF74DFF94FC7FF90B7851F97C" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[934,1073,1697,1723]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF74DFF94FC7FF90B7851F97C" box="[934,1073,1697,1723]" italics="true" pageId="43" pageNumber="44">N. africanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="195F6CF3571C9296DF90DC42B2336D1C">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
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<mods:namePart id="E17FB6D41A14BC56A82FAB1E12D67981">Sak, Serdar</mods:namePart>
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<mods:affiliation id="0AD5FA20964070E89C0713C3AFDF6A1F">Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162</mods:affiliation>
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<mods:namePart id="D11DE6BD1063054A4A7AF7B3E9A54E6C">Karaytuğ, Süphan</mods:namePart>
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<mods:affiliation id="3A855CEEA0D95E4523BCC10E98CCFD62">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
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<mods:namePart id="F8C4DC0BCC5F7A45DA42911508A60511">Huys, Rony</mods:namePart>
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<mods:affiliation id="2B4EA97595A278C86A4C0ED3E91AEB66">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation>
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<paragraph id="EA68776EF753FF8AFF4EF89B7851F88A" blockId="53.[151,1073,1842,1905]" box="[151,1073,1842,1869]" pageId="53" pageNumber="54">
<heading id="B120C002F753FF8AFF4EF89B7851F88A" bold="true" box="[151,1073,1842,1869]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
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<taxonomicName id="2DD70CEDF753FF8AFF4EF89B7FC3F88B" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994)" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[151,931,1842,1869]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF753FF8AFF4EF89B7DDEF88A" bold="true" box="[151,446,1842,1869]" italics="true" pageId="53" pageNumber="54">Phreatipontia speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF753FF8AFE17F89B7FFBF88B" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[462,923,1842,1869]" pageId="53" pageNumber="54" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
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<taxonomicNameLabel id="C3901607F753FF8AFC72F89A7851F88A" box="[939,1073,1843,1869]" pageId="53" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="EA68776EF753FF8AFF4EF8FE7D47F8B6" blockId="53.[151,1073,1842,1905]" box="[151,295,1879,1905]" pageId="53" pageNumber="54">
(
<figureCitation id="72EC6BEBF753FF8AFF46F8FE7D7EF8B6" box="[159,286,1879,1905]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStartId-0="55.[151,250,1943,1967]" captionStartId-1="56.[151,250,1943,1967]" captionStartId-2="57.[151,250,1943,1967]" captionStartId-3="58.[151,250,1943,1967]" captionTargetBox-0="[182,1406,193,1907]" captionTargetBox-1="[205,1381,188,1907]" captionTargetBox-2="[190,1399,183,1906]" captionTargetBox-3="[184,1402,193,1906]" captionTargetId-0="figure-17@55.[170,1418,181,1919]" captionTargetId-1="figure-17@56.[194,1393,181,1919]" captionTargetId-2="figure-17@57.[175,1412,181,1919]" captionTargetId-3="figure-17@58.[172,1415,181,1919]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." captionText-1="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." captionText-2="FIGURE 18. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." captionText-3="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi-0="http://doi.org/10.5281/zenodo.14021512" figureDoi-1="http://doi.org/10.5281/zenodo.14021514" figureDoi-2="http://doi.org/10.5281/zenodo.14021516" figureDoi-3="http://doi.org/10.5281/zenodo.14021518" httpUri-0="https://zenodo.org/record/14021512/files/figure.png" httpUri-1="https://zenodo.org/record/14021514/files/figure.png" httpUri-2="https://zenodo.org/record/14021516/files/figure.png" httpUri-3="https://zenodo.org/record/14021518/files/figure.png" pageId="53" pageNumber="54">Figs 16–19</figureCitation>
)
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<paragraph id="EA68776EF753FF8AFF4EF8327FCFF874" blockId="53.[151,1037,1947,2033]" box="[151,943,1947,1971]" pageId="53" pageNumber="54">
<taxonomicName id="2DD70CEDF753FF8AFF4EF8327FCFF874" ID-CoL="GFPX" authority="Cottarelli, Bruno &amp; Venanzetti, 1994" authorityName="Cottarelli, Bruno &amp; Venanzetti" authorityYear="1994" box="[151,943,1947,1971]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="speluncae" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF753FF8AFF4EF8327D75F875" box="[151,277,1947,1970]" italics="true" pageId="53" pageNumber="54">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF753FF8AFEFDF8327DDFF874" box="[292,447,1947,1971]" italics="true" pageId="53" pageNumber="54">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF753FF8AFE14F8327E52F874" box="[461,562,1947,1971]" italics="true" pageId="53" pageNumber="54">speluncae</emphasis>
<bibRefCitation id="8E460A9FF753FF8AFDE0F8327FCFF874" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[569,943,1947,1971]" pageId="53" pageNumber="54" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
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<paragraph id="EA68776EF753FF8AFF4EF8137E9AF815" blockId="53.[151,1037,1947,2033]" box="[151,762,1978,2002]" pageId="53" pageNumber="54">
<treatmentCitationGroup id="CAC75040F753FF8AFF4EF8137E9AF815" box="[151,762,1978,2002]" pageId="53" pageNumber="54">
<emphasis id="D8A3AB7CF753FF8AFF4EF8137E51F815" box="[151,561,1978,2002]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="2DD70CEDF753FF8AFF4EF8137DE9F816" authority="Sak, 2004" box="[151,393,1978,2002]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="turcicus">Neoleptastacus turcicus</taxonomicName>
—nomen nudum
</emphasis>
by
<treatmentCitation id="6B76517FF753FF8AFD8EF8137E9AF815" author="Sak, S." box="[599,762,1978,2002]" page="221" pageId="53" pageNumber="54" year="2004">
<bibRefCitation id="8E460A9FF753FF8AFD8EF8137E9AF815" author="Sak, S." box="[599,762,1978,2002]" pageId="53" pageNumber="54" refId="ref49361" refString="Sak, S. (2004) Arenopontiidae (Copepoda: Harpacticoida) familyasinin taksonomisi ve filogenisi. [Taxonomy and phylogeny of the family Arenopontiidae (Copepoda: Harpacticoida)]. PhD dissertation, University of Balikesir, Balikesir, 312 pp." type="book" year="2004">Sak (2004: 221)</bibRefCitation>
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<treatmentCitation id="6B76517FF753FF89FF4EF8717C9CFF77" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." lastPageId="54" lastPageNumber="55" page=")" pageId="53" pageNumber="54" year="1994">
<taxonomicName id="2DD70CEDF753FF89FF4EF8717C9CFF77" ID-CoL="6RVZJ" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994) Sak et al. (2008: 412) Original" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" lastPageId="54" lastPageNumber="55" order="Harpacticoida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="speluncae">
<paragraph id="EA68776EF753FF8AFF4EF871786DF836" blockId="53.[151,1037,1947,2033]" box="[151,1037,2008,2033]" pageId="53" pageNumber="54">
<emphasis id="D8A3AB7CF753FF8AFF4EF8717DFDF837" box="[151,413,2008,2032]" italics="true" pageId="53" pageNumber="54">Neoleptastacus speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF753FF8AFE72F8717F44F836" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[427,804,2008,2033]" pageId="53" pageNumber="54" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
<bibRefCitation id="8E460A9FF753FF8AFCE8F871786DF836" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[817,1037,2008,2033]" pageId="53" pageNumber="54" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF753FF8AFC84F8707FF1F837" box="[861,913,2008,2032]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2008: 412)
</bibRefCitation>
</paragraph>
<paragraph id="EA68776EF750FF89FF4EFF3F7F1AFF76" blockId="54.[151,1437,150,2013]" box="[151,890,150,177]" pageId="54" pageNumber="55">
<emphasis id="D8A3AB7CF750FF89FF4EFF3F7C9CFF77" bold="true" box="[151,252,150,176]" pageId="54" pageNumber="55">Original</emphasis>
<emphasis id="D8A3AB7CF750FF89FEDAFF3F7DF0FF77" bold="true" box="[259,400,150,176]" pageId="54" pageNumber="55">description.</emphasis>
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<bibRefCitation id="8E460A9FF750FF89FE4EFF3E7EF5FF76" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[407,661,151,177]" pageId="54" pageNumber="55" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF750FF89FDD1FF3E7E20FF76" box="[520,576,151,177]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(1994)
</bibRefCitation>
: 475
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–480;
<figureCitation id="72EC6BEBF750FF89FCC8FF3E7F15FF76" box="[785,885,151,177]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="13.[151,250,1943,1967]" captionStartId-1="14.[151,250,1844,1868]" captionTargetBox-0="[187,1401,193,1915]" captionTargetBox-1="[160,1424,322,1809]" captionTargetId-0="figure-17@13.[175,1413,181,1919]" captionTargetId-1="figure-17@14.[151,1436,307,1821]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" captionText-1="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi-0="http://doi.org/10.5281/zenodo.14021484" figureDoi-1="http://doi.org/10.5281/zenodo.14021486" httpUri-0="https://zenodo.org/record/14021484/files/figure.png" httpUri-1="https://zenodo.org/record/14021486/files/figure.png" pageId="54" pageNumber="55">Figs 2–3</figureCitation>
.
</paragraph>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="A2CD24E5F750FF89FF1EFF127913FD42" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="EA68776EF750FF89FF1EFF127DD9FE16" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="D8A3AB7CF750FF89FF1EFF127D09FF13" bold="true" box="[199,361,186,213]" pageId="54" pageNumber="55">Type locality.</emphasis>
<collectingCountry id="92C037FEF750FF89FEADFF127DC8FF12" box="[372,424,187,213]" name="Italy" pageId="54" pageNumber="55">Italy</collectingCountry>
,
<collectingRegion id="2813B98CF750FF89FE61FF127D9BFF12" box="[440,507,187,213]" country="Italy" name="Lazio" pageId="54" pageNumber="55">Lazio</collectingRegion>
,
<collectingCounty id="03090FE2F750FF89FDD0FF127EA2FF12" box="[521,706,187,213]" pageId="54" pageNumber="55">Latina Province</collectingCounty>
,
<collectingMunicipality id="0A0CED14F750FF89FD08FF127F25FF12" box="[721,837,187,213]" pageId="54" pageNumber="55">Sperlonga</collectingMunicipality>
,
<location id="EF0821B5F750FF89FC8DFF127FC5FF12" LSID="urn:lsid:plazi:treatment:627EC678F753FF83FF4EF89B7E45FD42:EF0821B5F750FF89FC8DFF127FC5FF12" box="[852,933,187,213]" country="Italy" county="Latina Province" municipality="Sperlonga" name="Tiberio" pageId="54" pageNumber="55" stateProvince="Lazio">Tiberio</location>
beach; on the banks of a little stream that originates from a spring on the beach; interstitial water.
<location id="EF0821B5F750FF89FCDEFF767F5DFF3E" LSID="urn:lsid:plazi:treatment:627EC678F753FF83FF4EF89B7E45FD42:EF0821B5F750FF89FCDEFF767F5DFF3E" box="[775,829,223,249]" country="Italy" county="Latina Province" municipality="Sperlonga" name="Note" pageId="54" pageNumber="55" stateProvince="Lazio">Note</location>
that
<bibRefCitation id="8E460A9FF750FF89FCACFF76780EFF3E" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[885,1134,223,249]" pageId="54" pageNumber="55" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF750FF89FC3CFF76787DFF3E" box="[997,1053,223,249]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(1994)
</bibRefCitation>
also collected
<specimenCount id="FCD1BCE7F750FF89FACDFF7679FCFF3E" box="[1300,1436,223,249]" count="2" pageId="54" pageNumber="55" type="female">two females</specimenCount>
from a second locality in the
<location id="EF0821B5F750FF89FE30FEAA7EFDFEDA" LSID="urn:lsid:plazi:treatment:627EC678F753FF83FF4EF89B7E45FD42:EF0821B5F750FF89FE30FEAA7EFDFEDA" box="[489,669,259,285]" country="Italy" county="Latina Province" municipality="Sperlonga" name="Latina Province" pageId="54" pageNumber="55" stateProvince="Lazio">Latina Province</location>
(
<collectorName id="472212B8F750FF89FD76FEAA7F54FEDA" box="[687,820,259,285]" pageId="54" pageNumber="55">S. Agostino</collectorName>
beach in
<location id="EF0821B5F750FF89FC7CFEAA7F88FEDA" LSID="urn:lsid:plazi:treatment:627EC678F753FF83FF4EF89B7E45FD42:EF0821B5F750FF89FC7CFEAA7F88FEDA" box="[933,1000,259,285]" country="Italy" county="Latina Province" municipality="Sperlonga" name="Gaeta" pageId="54" pageNumber="55" stateProvince="Lazio">Gaeta</location>
) but did not explicitly state where the female
<typeStatus id="356CC9CCF750FF89FF33FE8E7D2CFE86" box="[234,332,295,321]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
originated from. Since they allocated numbers 2–13 to the
<typeStatus id="356CC9CCF750FF89FC3FFE8E7831FE86" box="[998,1105,295,321]" pageId="54" pageNumber="55" type="paratype">paratypes</typeStatus>
(
<specimenCount id="FCD1BCE7F750FF89FBB9FE8E78D3FE86" box="[1120,1203,295,321]" count="6" pageId="54" pageNumber="55" type="female">six ♀♀</specimenCount>
and
<specimenCount id="FCD1BCE7F750FF89FB33FE8E7927FE86" box="[1258,1351,295,321]" count="5" pageId="54" pageNumber="55" type="male">five ♂♂</specimenCount>
in total were collected from Sperlonga,
<specimenCount id="FCD1BCE7F750FF89FDD8FEE27E3CFEA2" box="[513,604,331,357]" count="2" pageId="54" pageNumber="55" type="female">two ♀♀</specimenCount>
from
<collectorName id="472212B8F750FF89FD7BFEE27E85FEA2" box="[674,741,331,357]" pageId="54" pageNumber="55">Gaeta</collectorName>
) it appears rational to assume that number 1 was given to the
<typeStatus id="356CC9CCF750FF89FF4EFEC67C99FE4E" box="[151,249,367,393]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
and that it came from
<collectorName id="472212B8F750FF89FE2CFEC67EFFFE4E" box="[501,671,367,393]" pageId="54" pageNumber="55">Sperlonga. The</collectorName>
etymology of the species name and the fact that additional drawings (
<figureCitation id="72EC6BEBF750FF89FF46FE3A7D63FE6A" box="[159,259,403,429]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="54" pageNumber="55">Figs 2–e</figureCitation>
,
<figureCitation id="72EC6BEBF750FF89FED7FE3A7D59FE6A" box="[270,313,403,429]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="54" pageNumber="55">3–g</figureCitation>
) were presented of an atypical female of
<collectorName id="472212B8F750FF89FCD0FE3A7FECFE6A" box="[777,908,403,429]" pageId="54" pageNumber="55">S. Agostino</collectorName>
beach also seem to suggest that
<collectorName id="472212B8F750FF89FB21FE3A790AFE6A" box="[1272,1386,403,429]" pageId="54" pageNumber="55">Sperlonga</collectorName>
was the intended type locality.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFE737D36FDA6" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="D8A3AB7CF750FF89FF1EFE737DD1FE33" bold="true" box="[199,433,474,500]" pageId="54" pageNumber="55">Material examined.</emphasis>
<materialsCitation id="5ABF7D33F750FF89FE60FE727D32FDA6" collectingDate="2002-09-10" collectorName="S. Karaytug &amp; S. Sak" country="Turkey" county="Black Sea region" latitude="41.036633" location="All" longLatPrecision="1" longitude="38.94835" municipality="Ismailbeyli Village" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Giresun">
<specimenCount id="FCD1BCE7F750FF89FE60FE727E64FE32" box="[441,516,475,501]" count="1" pageId="54" pageNumber="55" type="female">One ♀</specimenCount>
(dissected on eight slides) and
<specimenCount id="FCD1BCE7F750FF89FCB1FE727FCDFE32" box="[872,941,475,501]" count="1" pageId="54" pageNumber="55" type="male">one ♂</specimenCount>
(dissected one seven slides); 10 individuals in ethanol.
<location id="EF0821B5F750FF89FECCFE567D59FDDE" LSID="urn:lsid:plazi:treatment:627EC678F753FF83FF4EF89B7E45FD42:EF0821B5F750FF89FECCFE567D59FDDE" box="[277,313,511,537]" country="Turkey" county="Black Sea region" latitude="41.036633" longLatPrecision="1" longitude="38.94835" municipality="Ismailbeyli Village" name="All" pageId="54" pageNumber="55" stateProvince="Giresun">All</location>
specimens collected from
<collectingMunicipality id="0A0CED14F750FF89FDB2FE567F21FDDE" box="[619,833,511,537]" pageId="54" pageNumber="55">Ismailbeyli Village</collectingMunicipality>
beach (
<geoCoordinate id="8FE311A9F750FF89FC42FE567857FDDE" box="[923,1079,511,537]" degrees="41" direction="north" minutes="02.198" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="41.036633">41°02.198’ N</geoCoordinate>
;
<geoCoordinate id="8FE311A9F750FF89FB9DFE5778BAFDDE" box="[1092,1242,510,537]" degrees="38" direction="east" minutes="56.901" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="38.94835">38°56.901’ E</geoCoordinate>
) close to a small stream, Giresun (
<collectingRegion id="2813B98CF750FF89FEB8FD8A7E50FDFA" box="[353,560,547,573]" country="Turkey" name="Giresun" pageId="54" pageNumber="55">Giresun Province</collectingRegion>
), northeastern
<collectingCountry id="92C037FEF750FF89FD07FD8A7F57FDFA" box="[734,823,547,573]" name="Turkey" pageId="54" pageNumber="55">Türkiye</collectingCountry>
(
<collectingCounty id="03090FE2F750FF89FC93FD8A787BFDFA" box="[842,1051,547,573]" pageId="54" pageNumber="55">Black Sea region</collectingCounty>
); leg.
<collectorName id="472212B8F750FF89FBB1FD8A7892FDFA" box="[1128,1266,547,573]" pageId="54" pageNumber="55">S. Karaytuğ</collectorName>
&amp;
<collectorName id="472212B8F750FF89FAF9FD8A7910FDFA" box="[1312,1392,547,573]" pageId="54" pageNumber="55">S. Sak</collectorName>
,
<date id="9E6951AEF750FF89FA59FD8A7D32FDA6" pageId="54" pageNumber="55" value="2002-09-10">
<collectingDate id="8E2DA846F750FF89FA59FD8A7D32FDA6" pageId="54" pageNumber="55" value="2002-09-10">10 September 2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFDC27913FD42" blockId="54.[151,1437,150,2013]" box="[199,1395,618,645]" pageId="54" pageNumber="55">
<emphasis id="D8A3AB7CF750FF89FF1EFDC27D3EFD42" bold="true" box="[199,350,619,645]" pageId="54" pageNumber="55">Body length.</emphasis>
394 μm (
<specimenCount id="FCD1BCE7F750FF89FE14FDC27D80FD42" box="[461,480,619,645]" count="1" pageId="54" pageNumber="55" type="female">♀</specimenCount>
), 346 μm (
<specimenCount id="FCD1BCE7F750FF89FD84FDC27E13FD42" box="[605,627,619,645]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
) [Cottarelli
<emphasis id="D8A3AB7CF750FF89FD24FDC27F55FD42" box="[765,821,619,645]" italics="true" pageId="54" pageNumber="55">ei al.</emphasis>
1994]; 356 μm (
<specimenCount id="FCD1BCE7F750FF89FC2DFDC27867FD42" box="[1012,1031,619,645]" count="1" pageId="54" pageNumber="55" type="female">♀</specimenCount>
), 338 μm (
<specimenCount id="FCD1BCE7F750FF89FB5DFDC278FAFD42" box="[1156,1178,619,645]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
) [present account].
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F750FF84FF1EFD277ECAFA27" lastPageId="59" lastPageNumber="60" pageId="54" pageNumber="55" type="description">
<paragraph id="EA68776EF750FF89FF1EFD277882FC0E" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="D8A3AB7CF750FF89FF1EFD277D8AFD6F" bold="true" box="[199,490,654,680]" pageId="54" pageNumber="55">Redescription of female.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 356 μm (
<emphasis id="D8A3AB7CF750FF89FA57FD2679FCFD6F" box="[1422,1436,655,680]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1). Cephalothorax maximum width 42 μm measured at posterior margin. Maximum body width 47 μm, measured at P5-bearing somite. Body (
<figureCitation id="72EC6BEBF750FF89FE39FD7E7E0AFD36" box="[480,618,727,753]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="54" pageNumber="55">Fig. 16A, B</figureCitation>
) slender and cylindrical without clear distinction between prosome and urosome. Sensillar pattern on body as figured. Hyaline frills of thoracic somites weakly developed and plain; those of genital double-somite and free abdominal somites well developed and consisting of rectangular digitate lappets that taper towards their distal margins (
<figureCitation id="72EC6BEBF750FF89FDB1FCEA7E98FC9A" box="[616,760,835,861]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="54" pageNumber="55">Figs 16A–B</figureCitation>
;
<figureCitation id="72EC6BEBF750FF89FCD0FCEA7F5CFC9A" box="[777,828,835,861]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="54" pageNumber="55">19A</figureCitation>
). Somites connected by well-developed intersomitic membranes. Genital double-somite slightly longer than wide (measured in dorsal aspect); with two ventral pores (
<figureCitation id="72EC6BEBF750FF89FF46FC227D69FC62" box="[159,265,907,933]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="54" pageNumber="55">Fig. 19A</figureCitation>
). Anal somite (
<figureCitation id="72EC6BEBF750FF89FE6EFC227E27FC62" box="[439,583,907,933]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="54" pageNumber="55">Figs 16C, D</figureCitation>
;
<figureCitation id="72EC6BEBF750FF89FD8CFC227EE6FC62" box="[597,646,907,933]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="54" pageNumber="55">19A</figureCitation>
) with two dorsal and four lateral pores. Anal frill triradiate, minutely incised (giving a spinulose appearance); anal operculum virtually straight, without ornamentation.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFC7A7EADFB02" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
Caudal rami (
<figureCitation id="72EC6BEBF750FF89FEBFFC7A7D99FC2A" box="[358,505,979,1005]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="54" pageNumber="55">Figs 16C, D</figureCitation>
;
<figureCitation id="72EC6BEBF750FF89FDD1FC7A7E5AFC2A" box="[520,570,979,1005]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="54" pageNumber="55">19A</figureCitation>
) about 3.4 times longer than wide (measured in dorsal view from anterior margin to apex of spinous process), distinctly tapering posteriorly; outer and inner margins slightly convex; with two ventral pores in proximal third and two lateral pores near insertion of seta III; terminal spinous process slightly recurved dorsally; no spinular ornamentation discernible. Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, located between seta V and posterior spinous process, with long outer spinule; seta V with proximal fracture plane; seta VI small, naked and located at inner distal corner; seta VII spatulate and tri-articulate at base.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFB66781FFB2E" blockId="54.[151,1437,150,2013]" box="[199,1151,1231,1257]" pageId="54" pageNumber="55">
Rostrum (
<figureCitation id="72EC6BEBF750FF89FEEEFB667DFDFB2E" box="[311,413,1231,1257]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="54" pageNumber="55">Fig. 16A</figureCitation>
) small, subtriangular, tapering distally; with two delicate sensilla.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFB5A7F95FA5A" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
Antennule (
<figureCitation id="72EC6BEBF750FF89FE96FB5A7DD7FACA" box="[335,439,1267,1293]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="54" pageNumber="55">Fig. 17C</figureCitation>
) long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about 2.5 times longer than wide. Segment 4 with long aesthetasc (L: 29 μm) fused at base with seta. Distal segment with eight setae (none of them spatulate) and apical acrothek consisting of short aesthetasc (L: 13 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[8 + acrothek].
</paragraph>
<paragraph id="EA68776EF750FF89FF1EFA0E7F90F9B2" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
Antenna (
<figureCitation id="72EC6BEBF750FF89FEEFFA0E7DFEFA06" box="[310,414,1447,1473]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="54" pageNumber="55">Fig. 17D</figureCitation>
). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment completely separated. Basis with few spinules along exopodal margin and with oblique spinular row near base of exopod. Exopod one-segmented and elongate, with a naked apical seta (about 1.4 times longer than exopod). Free endopodal segment with transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in
<figureCitation id="72EC6BEBF750FF89FEF0F99E7DF0F996" box="[297,400,1591,1617]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="54" pageNumber="55">Fig. 17D</figureCitation>
); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EF9D67FE9F95E" blockId="54.[151,1437,150,2013]" box="[199,905,1663,1689]" pageId="54" pageNumber="55">
Mandible, maxillule, maxilla and maxilliped as in
<taxonomicName id="2DD70CEDF750FF89FD25F9D67FE2F95E" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="2024" box="[764,898,1663,1689]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="phreatica">
<emphasis id="D8A3AB7CF750FF89FD25F9D67FE2F95E" box="[764,898,1663,1689]" italics="true" pageId="54" pageNumber="55">P. phreatica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EA68776EF750FF89FF1EF90A7E7DF81A" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
P1 (
<figureCitation id="72EC6BEBF750FF89FF2AF90A7D3AF97A" box="[243,346,1699,1725]" captionStart="FIGURE 18" captionStartId="57.[151,250,1943,1967]" captionTargetBox="[190,1399,183,1906]" captionTargetId="figure-17@57.[175,1412,181,1919]" captionTargetPageId="57" captionText="FIGURE 18. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021516" httpUri="https://zenodo.org/record/14021516/files/figure.png" pageId="54" pageNumber="55">Fig. 18A</figureCitation>
). Intercoxal sclerite wide, naked and subrectangular, with concave ventral margin. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis with spinular row near base of endopod; anterior surface with a small setiform spine near medial margin and one pore near articulation with coxa. Exopod three-segmented; all segments with several spinules around outer margin; exp-1 about as long as exp-3, with naked outer spine; exp-2 without outer element; exp-3 with short naked outer spine, and a longer naked spine and two geniculate setae distally. Endopod two-segmented, not prehensile, slightly longer than exopod; enp-1 with virtually straight inner margin, about 1.5 times longer than enp-2, with a serrate seta arising from about halfway down inner margin and two sets of two spinules along outer margin; enp-2 without spinules, distal margin with two geniculate setae, outermost of which shortest.
</paragraph>
<caption id="BEA827E6F751FF88FF4EF83E7CBEF83F" ID-DOI="http://doi.org/10.5281/zenodo.14021512" ID-Zenodo-Dep="14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1943,1967]" targetBox="[182,1406,193,1907]" targetPageId="55" targetType="figure">
<paragraph id="EA68776EF751FF88FF4EF83E7CBEF83F" blockId="55.[151,1436,1943,2040]" pageId="55" pageNumber="56">
<emphasis id="D8A3AB7CF751FF88FF4EF83E7D40F868" bold="true" box="[151,288,1943,1967]" pageId="55" pageNumber="56">FIGURE 16.</emphasis>
<taxonomicName id="2DD70CEDF751FF88FEFFF8317FC9F877" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994)" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[294,937,1944,1968]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF751FF88FEFFF8317E7DF877" box="[294,541,1944,1968]" italics="true" pageId="55" pageNumber="56">Phreatipontia speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF751FF88FDF3F8317FC2F877" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[554,930,1944,1968]" pageId="55" pageNumber="56" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="D8A3AB7CF751FF88FC76F8317843F877" bold="true" box="[943,1059,1944,1968]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="C3901607F751FF88FC76F8317843F877" box="[943,1059,1944,1968]" pageId="55" pageNumber="56" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral.
</paragraph>
</caption>
<caption id="BEA827E6F75EFF87FF4EF83E7966F814" ID-DOI="http://doi.org/10.5281/zenodo.14021514" ID-Zenodo-Dep="14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1943,1967]" targetBox="[205,1381,188,1907]" targetPageId="56" targetType="figure">
<paragraph id="EA68776EF75EFF87FF4EF83E7966F814" blockId="56.[151,1436,1943,2004]" pageId="56" pageNumber="57">
<emphasis id="D8A3AB7CF75EFF87FF4EF83E7D7EF868" bold="true" box="[151,286,1943,1967]" pageId="56" pageNumber="57">FIGURE 17.</emphasis>
<taxonomicName id="2DD70CEDF75EFF87FEFAF8317FC0F877" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994)" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[291,928,1944,1968]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75EFF87FEFAF8317E79F877" box="[291,537,1944,1968]" italics="true" pageId="56" pageNumber="57">Phreatipontia speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF75EFF87FDFCF8317FF9F877" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[549,921,1944,1968]" pageId="56" pageNumber="57" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="D8A3AB7CF75EFF87FC7CF8317877F877" bold="true" box="[933,1047,1944,1968]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="C3901607F75EFF87FC7CF8317877F877" box="[933,1047,1944,1968]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows].
</paragraph>
</caption>
<caption id="BEA827E6F75FFF86FF4EF83E7D89F813" ID-DOI="http://doi.org/10.5281/zenodo.14021516" ID-Zenodo-Dep="14021516" httpUri="https://zenodo.org/record/14021516/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1943,1967]" targetBox="[190,1399,183,1906]" targetPageId="57" targetType="figure">
<paragraph id="EA68776EF75FFF86FF4EF83E7D89F813" blockId="57.[151,1437,1943,2004]" pageId="57" pageNumber="58">
<emphasis id="D8A3AB7CF75FFF86FF4EF83E7D7FF868" bold="true" box="[151,287,1943,1967]" pageId="57" pageNumber="58">FIGURE 18.</emphasis>
<taxonomicName id="2DD70CEDF75FFF86FEFDF8317FC1F877" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994)" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[292,929,1944,1968]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75FFF86FEFDF8317E7AF877" box="[292,538,1944,1968]" italics="true" pageId="57" pageNumber="58">Phreatipontia speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF75FFF86FDFFF8317FFBF877" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[550,923,1944,1968]" pageId="57" pageNumber="58" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="D8A3AB7CF75FFF86FC7EF8317879F877" bold="true" box="[935,1049,1944,1968]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="C3901607F75FFF86FC7EF8317879F877" box="[935,1049,1944,1968]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior.
</paragraph>
</caption>
<caption id="BEA827E6F75CFF85FF4EF83E7D13F813" ID-DOI="http://doi.org/10.5281/zenodo.14021518" ID-Zenodo-Dep="14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[184,1402,193,1906]" targetPageId="58" targetType="figure">
<paragraph id="EA68776EF75CFF85FF4EF83E7D13F813" blockId="58.[151,1436,1943,2004]" pageId="58" pageNumber="59">
<emphasis id="D8A3AB7CF75CFF85FF4EF83E7D44F868" bold="true" box="[151,292,1943,1967]" pageId="58" pageNumber="59">FIGURE 19.</emphasis>
<taxonomicName id="2DD70CEDF75CFF85FEF7F8317FABF877" authority="(Cottarelli, Bruno &amp; Venanzetti, 1994)" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[302,971,1944,1968]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75CFF85FEF7F8317E49F877" box="[302,553,1944,1968]" italics="true" pageId="58" pageNumber="59">Phreatipontia speluncae</emphasis>
(
<bibRefCitation id="8E460A9FF75CFF85FDE2F8317FA4F877" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[571,964,1944,1968]" pageId="58" pageNumber="59" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">Cottarelli, Bruno &amp; Venanzetti, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="D8A3AB7CF75CFF85FC0CF831782DF877" bold="true" box="[981,1101,1944,1968]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="C3901607F75CFF85FC0CF831782DF877" box="[981,1101,1944,1968]" pageId="58" pageNumber="59" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: (A) urosome (♀), ventral; (B) urosome (♂), ventral.
</paragraph>
</caption>
<paragraph id="EA68776EF75DFF84FF1EFF3E7C94FDA6" blockId="59.[151,1437,151,609]" pageId="59" pageNumber="60">
P2–P4 (
<figureCitation id="72EC6BEBF75DFF84FEFDFF3E7DD1FF76" box="[292,433,151,177]" captionStart="FIGURE 18" captionStartId="57.[151,250,1943,1967]" captionTargetBox="[190,1399,183,1906]" captionTargetId="figure-17@57.[175,1412,181,1919]" captionTargetPageId="57" captionText="FIGURE 18. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021516" httpUri="https://zenodo.org/record/14021516/files/figure.png" pageId="59" pageNumber="60">Fig. 18B–D</figureCitation>
). Intercoxal sclerite naked (P2–P4), rectangular (P2) or squarish (P3–P4) with deeply concave ventral margins. Praecoxae triangular, small and naked. Coxae squarish (P3) or slightly wider than long (P2, P4) and without ornamentation. Bases smaller than coxae, with a spinular row near base of endopod (P2–P3) and a few spinules around outer corner in P2 and P4; anterior surface with a pore in P3-P4; outer basal seta absent in P2, naked in P3–P4. Exopods three-segmented; segments with coarse spinular ornamentation along outer margin; outer spine of exp-1 and exp-2 naked; exp-3 with an outer unipinnate spine and two bipinnate setae distally, P4 exp-3 with additional serrate seta on inner margin; P4 exp-2 only marginally longer than exp-1. Endopods two-segmented; P2–P4 enp-1 unarmed, about 1.2, 1.5, and 4.2 times longer than their respective distal segments, with few coarse spinules along outer margin as figured, but without ornamentation along inner margin; P2–P3 enp-2 with few spinules around distal margin; P2 enp-2 with long, apically serrate, backwardly directed seta near proximal margin and one unipinnate setae apically; P3 enp-2 with long, bipinnate apically; distal margin of P4 enp-2 with long, distally serrate and basally fused, inner seta, and shorter unipinnate, outer seta. Spine and seta formula as follows:
</paragraph>
<paragraph id="EA68776EF75DFF84FE6EFD267F76FCFE" pageId="59" pageNumber="60">
<table id="98D785CEF75D0040FF2EFD267F5CFCFE" box="[247,828,655,825]" gridcols="3" gridrows="4" inLine="true" pageId="59" pageNumber="60">
<tr id="54E7752CF75D0040FF2EFD267F5CFD6E" box="[247,828,655,681]" gridrow="0" pageId="59" pageNumber="60" rowspan-0="1">
<th id="17361C50F75D0040FE6EFD267E6EFD6E" box="[439,526,655,681]" gridcol="1" gridrow="0" pageId="59" pageNumber="60">Exopod</th>
<th id="17361C50F75D0040FD0EFD267F5CFD6E" box="[727,828,655,681]" gridcol="2" gridrow="0" pageId="59" pageNumber="60">Endopod</th>
</tr>
<tr id="54E7752CF75D0040FF2EFD7E7F5CFD36" box="[247,828,727,753]" gridrow="1" pageId="59" pageNumber="60">
<th id="17361C50F75D0040FF2EFD7E7D75FD36" box="[247,277,727,753]" gridcol="0" gridrow="1" pageId="59" pageNumber="60">P2</th>
<td id="17361C50F75D0040FE6EFD7E7E6EFD36" box="[439,526,727,753]" gridcol="1" gridrow="1" pageId="59" pageNumber="60">0.0.021</td>
<td id="17361C50F75D0040FD0EFD7E7F5CFD36" box="[727,828,727,753]" gridcol="2" gridrow="1" pageId="59" pageNumber="60">0.110</td>
</tr>
<tr id="54E7752CF75D0040FF2EFD527F5CFCD2" box="[247,828,763,789]" gridrow="2" pageId="59" pageNumber="60">
<th id="17361C50F75D0040FF2EFD527D75FCD2" box="[247,277,763,789]" gridcol="0" gridrow="2" pageId="59" pageNumber="60">P3</th>
<td id="17361C50F75D0040FE6EFD527E6EFCD2" box="[439,526,763,789]" gridcol="1" gridrow="2" pageId="59" pageNumber="60">0.0.021</td>
<td id="17361C50F75D0040FD0EFD527F5CFCD2" box="[727,828,763,789]" gridcol="2" gridrow="2" pageId="59" pageNumber="60">0.010</td>
</tr>
<tr id="54E7752CF75D0040FF2EFCB67F5CFCFE" box="[247,828,799,825]" gridrow="3" pageId="59" pageNumber="60">
<th id="17361C50F75D0040FF2EFCB67D75FCFE" box="[247,277,799,825]" gridcol="0" gridrow="3" pageId="59" pageNumber="60">P4</th>
<td id="17361C50F75D0040FE6EFCB67E6EFCFE" box="[439,526,799,825]" gridcol="1" gridrow="3" pageId="59" pageNumber="60">0.0.121</td>
<td id="17361C50F75D0040FD0EFCB67F5CFCFE" box="[727,828,799,825]" gridcol="2" gridrow="3" pageId="59" pageNumber="60">0.020</td>
</tr>
</table>
</paragraph>
<paragraph id="EA68776EF75DFF84FF4EFCCF7F17FC2C" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
Fifth legs (
<figureCitation id="72EC6BEBF75DFF84FECCFCCF7D1CFC47" box="[277,380,870,896]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="59" pageNumber="60">Fig. 19A</figureCitation>
) closely set together, almost touching medially. Baseoendopod and exopod fused, forming a subrectangular plate with straight distal margin; with one pore on anterior surface; inner distal corner with long, bipinnate spinous process (homologous to inner spine); distal margin with long naked outer seta and two short equally long, bipinnate spines; outer basal seta long and plumose.
</paragraph>
<paragraph id="EA68776EF75DFF84FF4EFC5C7FE3FBD7" blockId="59.[151,1437,870,2041]" box="[151,899,1013,1040]" pageId="59" pageNumber="60">Sixth legs vestigial, forming opercula closing off genital apertures.</paragraph>
<paragraph id="EA68776EF75DFF84FF1EFBB07F75FBBC" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
<emphasis id="D8A3AB7CF75DFF84FF1EFBB07DB4FBF4" bold="true" box="[199,468,1049,1075]" pageId="59" pageNumber="60">Redescription of male.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 338 µm (
<emphasis id="D8A3AB7CF75DFF84FAAEFBB379E5FBF4" box="[1399,1413,1050,1075]" italics="true" pageId="59" pageNumber="60">n</emphasis>
= 1). Body ornamentation essentially as in female (
<figureCitation id="72EC6BEBF75DFF84FD63FB947F43FB90" box="[698,803,1085,1111]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="59" pageNumber="60">Fig. 17A</figureCitation>
). Sexual dimorphism in antennule, genital segmentation, P5, and P6. Spermatophore length approximately 70 μm.
</paragraph>
<paragraph id="EA68776EF75DFF84FF1EFB2D7D69FAB2" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
Antennule (
<figureCitation id="72EC6BEBF75DFF84FE92FB2D7DCEFB59" box="[331,430,1156,1182]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="59" pageNumber="60">Fig. 17B</figureCitation>
) 8-segmented, haplocer; geniculation between segments 6 and 7. Segment 1 with a naked seta; segment 2 longest and about 2.7 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae; segment 4 an incomplete sclerite with two setae; segment 5 with four setae, one pinnate spine and a long aesthetasc (43 µm) fused basally to a slender seta; segments 6–7 with one seta each; distal segment with seven setae (none of which spatulate) and apical acrothek consisting of short aesthetasc (16 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[2], 5-[4 + (1 + ae)], 6-[1], 7-[1], 8-[7 + acrothek].
</paragraph>
<paragraph id="EA68776EF75DFF84FF1EFAD778E8FA5F" blockId="59.[151,1437,870,2041]" box="[199,1160,1406,1433]" pageId="59" pageNumber="60">
P5 (
<figureCitation id="72EC6BEBF75DFF84FF2DFAD77D38FA5F" box="[244,344,1406,1432]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="59" pageNumber="60">Fig. 19B</figureCitation>
) essentially as in female but inner spinous process slightly more slender.
</paragraph>
<paragraph id="EA68776EF75DFF84FF1EFA0B7ECAFA27" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
Sixth legs (
<figureCitation id="72EC6BEBF75DFF84FE91FA0B7DCDFA7B" box="[328,429,1442,1468]" captionStart="FIGURE 19" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[184,1402,193,1906]" captionTargetId="figure-17@58.[172,1415,181,1919]" captionTargetPageId="58" captionText="FIGURE 19. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) urosome (♀), ventral; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021518" httpUri="https://zenodo.org/record/14021518/files/figure.png" pageId="59" pageNumber="60">Fig. 19B</figureCitation>
) asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer plumose seta and a shorter inner bipinnate seta.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F75DFF83FF1EFA437E45FD42" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="discussion">
<paragraph id="EA68776EF75DFF84FF1EFA437D49F91D" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
<emphasis id="D8A3AB7CF75DFF84FF1EFA437D5BF9C3" bold="true" box="[199,315,1514,1540]" pageId="59" pageNumber="60">Remarks.</emphasis>
The report of an inner seta on P3 exp-2 by
<bibRefCitation id="8E460A9FF75DFF84FCC7FA407877F9C3" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[798,1047,1513,1540]" pageId="59" pageNumber="60" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF75DFF84FC57FA437FA6F9C4" box="[910,966,1513,1539]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
(1994)
</bibRefCitation>
is extremely doubtful and requires confirmation. No other member in the family
<taxonomicName id="2DD70CEDF75DFF84FD4BF9A47F20F9E0" box="[658,832,1549,1575]" class="Copepoda" family="Arenopontiidae" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="family">Arenopontiidae</taxonomicName>
displays an inner seta on this segment. Our specimens from the Black Sea agree in virtually all other morphological aspects with Cottarelli
<emphasis id="D8A3AB7CF75DFF84FB4BF99B78AFF98C" box="[1170,1231,1585,1611]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
’s (1994) original description, adding further credence to the notion that their illustration of P3 is based on an observational error. The presence of an outer basal seta on P2 (their
<figureCitation id="72EC6BEBF75DFF84FD67F9D17F7EF954" box="[702,798,1656,1683]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="59" pageNumber="60">Fig. 3–e</figureCitation>
), their statement that the somitic hyaline frills are plain, and the absence of spinular ornamentation on the spinous process of P5 are also questionable and require further examination.
</paragraph>
<paragraph id="EA68776EF75DFF84FF1EF94D7D15F877" blockId="59.[151,1437,870,2041]" pageId="59" pageNumber="60">
<bibRefCitation id="8E460A9FF75DFF84FF1EF94D7DA1F939" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[199,449,1764,1790]" pageId="59" pageNumber="60" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF75DFF84FEE1F94D7D10F939" box="[312,368,1764,1790]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
(1994)
</bibRefCitation>
did not observe any variability among the individuals from Sperlonga but illustrated the P5 and caudal ramus of a female from S. Agostino beach without further comment. The fifth leg of the latter deviates from that of the
<typeStatus id="356CC9CCF75DFF84FE96F8827DD1F882" box="[335,433,1835,1861]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
in the shorter spinous process, the longer outer marginal seta (extending well beyond the process) and the general appearance of the two medial elements which are unequal in length and setiform in shape. The dorsal view of the caudal ramus is difficult to compare with the
<typeStatus id="356CC9CCF75DFF84FC4EF8DA7F99F84A" box="[919,1017,1907,1933]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
since it was only illustrated in lateral aspect for the latter.
</paragraph>
<paragraph id="EA68776EF75DFF83FF1EF813785CFEDA" blockId="59.[151,1437,870,2041]" lastBlockId="60.[151,1437,151,645]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
The two described species of
<taxonomicName id="2DD70CEDF75DFF84FDC4F8137ED8F813" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[541,696,1978,2004]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D8A3AB7CF75DFF84FDC4F8137ED8F813" box="[541,696,1978,2004]" italics="true" pageId="59" pageNumber="60">Phreatipontia</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75DFF84FD18F8137F4CF813" bold="true" box="[705,812,1978,2004]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C3901607F75DFF84FD18F8137F4CF813" box="[705,812,1978,2004]" pageId="59" pageNumber="60" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are morphologically very similar. The
<typeStatus id="356CC9CCF75DFF84FB37F812797EF812" box="[1262,1310,1979,2005]" pageId="59" pageNumber="60">type</typeStatus>
species,
<taxonomicName id="2DD70CEDF75DFF84FA51F8127D67F83F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75DFF84FA51F8127D67F83F" italics="true" pageId="59" pageNumber="60">P. speluncae</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75DFF84FED6F8777DF1F83F" bold="true" box="[271,401,2014,2040]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C3901607F75DFF84FED6F8777DF1F83F" box="[271,401,2014,2040]" pageId="59" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, can primarily be differentiated from
<taxonomicName id="2DD70CEDF75DFF84FCE7F8767FA4F83F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="2024" box="[830,964,2014,2040]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="phreatica" status="comb. nov.">
<emphasis id="D8A3AB7CF75DFF84FCE7F8767FA4F83F" box="[830,964,2014,2040]" italics="true" pageId="59" pageNumber="60">P. phreatica</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75DFF84FC15F877782EF83F" bold="true" box="[972,1102,2014,2040]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C3901607F75DFF84FC15F877782EF83F" box="[972,1102,2014,2040]" pageId="59" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
by morphometric differences in the caudal rami and swimming legs. The exopodal segments of P2–P3 are distinctly more slender and longer in the latter, P2 enp-2 is markedly shorter (enp-1:enp-2 length ratio 1.6
<emphasis id="D8A3AB7CF75AFF83FC47FF127FD5FF13" box="[926,949,187,212]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
1.2) while P4 enp-1 is longer (enp-1:enp-2 length ratio 5.8
<emphasis id="D8A3AB7CF75AFF83FE9EFF767D3EFF3F" box="[327,350,223,248]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
4.2). The caudal rami are generally more slender and longer (L:W ratio 3.9
<emphasis id="D8A3AB7CF75AFF83FB73FF7678A1FF3F" box="[1194,1217,223,248]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
3.4) in
<taxonomicName id="2DD70CEDF75AFF83FACEFF7679FCFF3E" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="2024" box="[1303,1436,223,249]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="phreatica" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FACEFF7679FCFF3E" box="[1303,1436,223,249]" italics="true" pageId="60" pageNumber="61">P. phreatica</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75AFF83FF4EFEAA7D78FEDA" bold="true" box="[151,280,259,285]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FF4EFEAA7D78FEDA" box="[151,280,259,285]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
while the dorsal seta VII is more foliaceous in
<taxonomicName id="2DD70CEDF75AFF83FCF0FEAA7FD5FEDA" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[809,949,259,285]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FCF0FEAA7FD5FEDA" box="[809,949,259,285]" italics="true" pageId="60" pageNumber="61">P. speluncae</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75AFF83FC65FEAA785CFEDA" bold="true" box="[956,1084,259,285]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FC65FEAA785CFEDA" box="[956,1084,259,285]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="EA68776EF75AFF83FF1EFE8E7E45FD42" blockId="60.[151,1437,151,645]" pageId="60" pageNumber="61">
In addition to the
<typeStatus id="356CC9CCF75AFF83FE46FE8E7DAFFE86" box="[415,463,295,321]" pageId="60" pageNumber="61">type</typeStatus>
locality,
<taxonomicName id="2DD70CEDF75AFF83FDE6FE8E7EAFFE86" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[575,719,295,321]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FDE6FE8E7EAFFE86" box="[575,719,295,321]" italics="true" pageId="60" pageNumber="61">P. speluncae</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75AFF83FD03FE8E7F3EFE86" bold="true" box="[730,862,295,321]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FD03FE8E7F3EFE86" box="[730,862,295,321]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
was also obtained in interstitial freshwater of S. Agostino beach (Gaeta, Latina Province, Lazio) on the banks of a little stream fed by a spring in Grotta del Serpente (
<bibRefCitation id="8E460A9FF75AFF83FF46FEC67DF3FE4E" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." box="[159,403,367,393]" pageId="60" pageNumber="61" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF75AFF83FECDFEC67D30FE4E" box="[276,336,367,393]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
1994
</bibRefCitation>
) and from the mouths of the Fiora River and Valfragida Stream, in the Viterbo Province (Lazio) (
<bibRefCitation id="8E460A9FF75AFF83FF2EFE3A7D82FE6A" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Berera, R." box="[247,482,403,429]" pageId="60" pageNumber="61" pagination="129 - 143" refId="ref46147" refString="Cottarelli, V., Bruno, M. C. &amp; Berera, R. (1998) Remarks on the genus Ichnusella (Crustacea, Copepoda, Harpacticoida) and description of two new species from subterranean freshwater habitats in Latium and Sardinia, Italy. Vie et Milieu, 49, 129 - 143." type="journal article" year="1998">
Cottarelli
<emphasis id="D8A3AB7CF75AFF83FEB0FE3A7DC1FE6A" box="[361,417,403,429]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="8E460A9FF75AFF83FE34FE3A7E1CFE6A" author="Berera, R." box="[493,636,403,429]" pageId="60" pageNumber="61" refId="ref45107" refString="Berera, R. (2000) Gli Psammopsyllinae Krishnaswamy 1956 (Copepoda, Harpacticoida) come esempio di colonizzazione delle acque continentali: osservazioni sugli aspetti biologici, ecologici ed evolutivi. PhD dissertation, Universita' degli Studi della Tuscia, Viterbo, 121 pp." type="book" year="2000">Berera 2000</bibRefCitation>
).
<bibRefCitation id="8E460A9FF75AFF83FD54FE3A7FD6FE6A" author="Cottarelli, V. &amp; Berera, R." box="[653,950,403,429]" pageId="60" pageNumber="61" pagination="27 - 30" refId="ref46077" refString="Cottarelli, V. &amp; Berera, R. (2004) Il fiume sotto il fiume: ricerche sul popolamento iporreico a crostacei del Fiume Fiora (Italia Centrale). [The river under the river: researches on the hyporheic crustacean community of the Fiora River (Central Italia).] Studi trentini di Scienze naturali, Acta biologica, 80, 27 - 30." type="journal article" year="2004">Cottarelli &amp; Berera (2004)</bibRefCitation>
also recorded it from the banks of the Fiora River near Montalto di Castro. Our material extends its distribution further into the Black Sea basin.
<bibRefCitation id="8E460A9FF75AFF83FB2CFE1E7C82FE32" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." pageId="60" pageNumber="61" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF75AFF83FABCFE1E79FDFE16" box="[1381,1437,439,465]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
(1994)
</bibRefCitation>
suggested that the species may be restricted to fresh or slightly brackish water but admitted that more data are needed before this can be corroborated. It is conceivable that
<taxonomicName id="2DD70CEDF75AFF83FCAAFE567F9FFDDE" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[883,1023,511,537]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FCAAFE567F9FFDDE" box="[883,1023,511,537]" italics="true" pageId="60" pageNumber="61">P. speluncae</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75AFF83FBDFFE5678E7FDDE" bold="true" box="[1030,1159,511,537]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FBDFFE5678E7FDDE" box="[1030,1159,511,537]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(and
<taxonomicName id="2DD70CEDF75AFF83FB1EFE56792CFDDE" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="2024" box="[1223,1356,511,537]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="phreatica" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FB1EFE56792CFDDE" box="[1223,1356,511,537]" italics="true" pageId="60" pageNumber="61">P. phreatica</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF75AFF83FA8DFE567CA8FDFA" bold="true" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FA8DFE567CA8FDFA" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
) is in the process of colonizing continental subterranean waters (
<bibRefCitation id="8E460A9FF75AFF83FC43FD8A7802FDFA" author="Bruno, M. C. &amp; Cottarelli, V. &amp; Berrera, R." box="[922,1122,547,573]" pageId="60" pageNumber="61" pagination="69 - 79" refId="ref45518" refString="Bruno, M. C., Cottarelli, V. &amp; Berrera, R. (1998) Preliminary remarks on the cladistic systematics in some taxa of Leptopontiidae and Parastenocarididae (Copepoda, Harpacticoida). Bolletino. Museo Civico di Storia naturale di Verona, 2 ° Serie, 13, 69 - 79." type="journal article" year="1998">
Bruno
<emphasis id="D8A3AB7CF75AFF83FC31FD8A7840FDFA" box="[1000,1056,547,573]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
1998
</bibRefCitation>
). A third species, apparently close to
<taxonomicName id="2DD70CEDF75AFF83FF2EFDEE7DE4FDA6" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[247,388,583,609]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="speluncae" status="comb. nov.">
<emphasis id="D8A3AB7CF75AFF83FF2EFDEE7DE4FDA6" box="[247,388,583,609]" italics="true" pageId="60" pageNumber="61">P. speluncae</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="C3901607F75AFF83FE55FDEE7E6DFDA6" box="[396,525,583,609]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
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and still under study, was discovered in freshwater habitats in
<collectingCountry id="92C037FEF75AFF83FB00FDEE7948FDA6" box="[1241,1320,583,609]" name="Greece" pageId="60" pageNumber="61">Greece</collectingCountry>
(
<bibRefCitation id="8E460A9FF75AFF83FAE1FDEE7C96FD42" author="Bruno, M. C. &amp; Cottarelli, V. &amp; Berrera, R." pageId="60" pageNumber="61" pagination="69 - 79" refId="ref45518" refString="Bruno, M. C., Cottarelli, V. &amp; Berrera, R. (1998) Preliminary remarks on the cladistic systematics in some taxa of Leptopontiidae and Parastenocarididae (Copepoda, Harpacticoida). Bolletino. Museo Civico di Storia naturale di Verona, 2 ° Serie, 13, 69 - 79." type="journal article" year="1998">
Bruno
<emphasis id="D8A3AB7CF75AFF83FA51FDEE7CD4FD42" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="8E460A9FF75AFF83FEDBFDC27E78FD42" author="Bruno, M. C. &amp; Cottarelli, V." box="[258,536,619,645]" pageId="60" pageNumber="61" pagination="305 - 315" refId="ref45466" refString="Bruno, M. C. &amp; V. Cottarelli, V. (1999) New records of harpacticoids (Crustacea, Copepoda) from subterranean fresh and brackish waters of Greece. Contributions to the Zoogeography and Ecology of the Eastern Mediterranean Region, 1, 305 - 315." type="journal article" year="1999">Bruno &amp; Cottarelli 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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162
data/62/7E/C6/627EC678F75AFF83FF4EFA1279F9F82D.xml → data/62/7E/C6/627EC678F75AFF83FF4EFA1279FDF82D.xml
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@ -1,90 +1,105 @@
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Key to genera of Key to genera of
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1. P1 endopod prehensile, enp-1 distinctly longer than exopod; innermost element of P1 exp-3 penicillate.................................................................................................. 1. P1 endopod prehensile, enp-1 distinctly longer than exopod; innermost element of P1 exp-3 penicillate..................................................................................................
<taxonomicName id="2DD70CEDF75AFF83FB70F9B479F9F9F5" authority="Kunz, 1937" authorityName="Kunz" authorityYear="1937" box="[1193,1433,1565,1586]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus"> <taxonomicName id="2DD70CEDF75AFF83FB70F9B479F9F9F5" authority="Kunz, 1937" authorityName="Kunz" authorityYear="1937" box="[1193,1433,1565,1586]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D8A3AB7CF75AFF83FB70F9B47940F9F5" box="[1193,1312,1565,1586]" italics="true" pageId="60" pageNumber="61">Arenopontia</emphasis> <emphasis id="D8A3AB7CF75AFF83FB70F9B47940F9F5" box="[1193,1312,1565,1586]" italics="true" pageId="60" pageNumber="61">Arenopontia</emphasis>
<bibRefCitation id="8E460A9FF75AFF83FAFFF9B479F9F9F5" author="Kunz, H." box="[1318,1433,1565,1586]" pageId="60" pageNumber="61" pagination="95 - 115" refId="ref47619" refString="Kunz, H. (1937) Zur Kenntnis der Harpacticoiden des Kustengrundwassers der Kieler Forde. (Studien an marinen Copepoden I.). Kieler Meeresforschungen, 2, 95 - 115." type="journal article" year="1937">Kunz, 1937</bibRefCitation> <bibRefCitation id="8E460A9FF75AFF83FAFFF9B479F9F9F5" author="Kunz, H." box="[1318,1433,1565,1586]" pageId="60" pageNumber="61" pagination="95 - 115" refId="ref47619" refString="Kunz, H. (1937) Zur Kenntnis der Harpacticoiden des Kustengrundwassers der Kieler Forde. (Studien an marinen Copepoden I.). Kieler Meeresforschungen, 2, 95 - 115." type="journal article" year="1937">Kunz, 1937</bibRefCitation>
</taxonomicName> </taxonomicName>
. .
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF99379FDF988" blockId="60.[151,1437,1536,2027]" box="[226,1437,1594,1616]" pageId="60" pageNumber="61">P1 endopod not prehensile, enp-1 not longer than exopod; innermost element of P1 exp-3 geniculate...................2.</paragraph> <paragraph id="EA68776EF75AFF83FF3BF99379FDF988" blockId="60.[151,1437,1536,2027]" box="[226,1437,1594,1616]" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF9F179FCF9AA" blockId="60.[151,1437,1536,2027]" box="[151,1436,1623,1646]" pageId="60" pageNumber="61">2. P5 of both sexes with innermost element forming a distinct spinous process which is incorporated in the segment.........3.</paragraph> <keyLead id="5126D45BF75AFF83FF3BF99379FDF988" box="[226,1437,1594,1616]" pageId="60" pageNumber="61">P1 endopod not prehensile, enp-1 not longer than exopod; innermost element of P1 exp-3 geniculate...................2.</keyLead>
<paragraph id="EA68776EF75AFF83FF3BF9DC79FDF94D" blockId="60.[151,1437,1536,2027]" box="[226,1437,1653,1675]" pageId="60" pageNumber="61">P5 of both sexes with innermost element not modified (occasionally fused at base).................................7.</paragraph> </paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF9F179FDF94D" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF9F179FCF9AA" blockId="60.[151,1437,1536,2027]" box="[151,1436,1623,1646]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF9F179FCF9AA" box="[151,1436,1623,1646]" pageId="60" pageNumber="61">2. P5 of both sexes with innermost element forming a distinct spinous process which is incorporated in the segment.........3.</keyLead>
</paragraph>
<paragraph id="EA68776EF75AFF83FF3BF9DC79FDF94D" blockId="60.[151,1437,1536,2027]" box="[226,1437,1653,1675]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF9DC79FDF94D" box="[226,1437,1653,1675]" pageId="60" pageNumber="61">P5 of both sexes with innermost element not modified (occasionally fused at base).................................7.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF93B79FCF902" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF93B79FCF960" blockId="60.[151,1437,1536,2027]" box="[151,1436,1682,1704]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF93B79FCF960" blockId="60.[151,1437,1536,2027]" box="[151,1436,1682,1704]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF93B79FCF960" box="[151,1436,1682,1704]" pageId="60" pageNumber="61">
3. Distal margin of P1 enp-2 with two geniculate setae........................................ 3. Distal margin of P1 enp-2 with two geniculate setae........................................
<taxonomicName id="2DD70CEDF75AFF83FB61F93B795CF960" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1208,1340,1682,1703]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov."> <taxonomicName id="2DD70CEDF75AFF83FB61F93B795CF960" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1208,1340,1682,1703]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D8A3AB7CF75AFF83FB61F93B795CF960" box="[1208,1340,1682,1703]" italics="true" pageId="60" pageNumber="61">Phreatipontia</emphasis> <emphasis id="D8A3AB7CF75AFF83FB61F93B795CF960" box="[1208,1340,1682,1703]" italics="true" pageId="60" pageNumber="61">Phreatipontia</emphasis>
@ -92,46 +107,70 @@ Key to genera of
<emphasis id="D8A3AB7CF75AFF83FA9BF93B79FCF960" bold="true" box="[1346,1436,1682,1703]" pageId="60" pageNumber="61"> <emphasis id="D8A3AB7CF75AFF83FA9BF93B79FCF960" bold="true" box="[1346,1436,1682,1703]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3901607F75AFF83FA9BF93B79FCF960" box="[1346,1436,1682,1703]" pageId="60" pageNumber="61" rank="genus">gen. nov.</taxonomicNameLabel> <taxonomicNameLabel id="C3901607F75AFF83FA9BF93B79FCF960" box="[1346,1436,1682,1703]" pageId="60" pageNumber="61" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis> </emphasis>
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF91979FCF902" blockId="60.[151,1437,1536,2027]" box="[226,1436,1711,1734]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF3BF91979FCF902" blockId="60.[151,1437,1536,2027]" box="[226,1436,1711,1734]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF91979FCF902" box="[226,1436,1711,1734]" pageId="60" pageNumber="61">
Distal margin of P1 enp-2 with outer naked spine and inner geniculate seta............ Distal margin of P1 enp-2 with outer naked spine and inner geniculate seta............
<taxonomicName id="2DD70CEDF75AFF83FBE3F919790EF902" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[1082,1390,1711,1734]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus"> <taxonomicName id="2DD70CEDF75AFF83FBE3F919790EF902" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[1082,1390,1711,1734]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D8A3AB7CF75AFF83FBE3F91978ADF902" box="[1082,1229,1712,1733]" italics="true" pageId="60" pageNumber="61">Neoleptastacus</emphasis> <emphasis id="D8A3AB7CF75AFF83FBE3F91978ADF902" box="[1082,1229,1712,1733]" italics="true" pageId="60" pageNumber="61">Neoleptastacus</emphasis>
<bibRefCitation id="8E460A9FF75AFF83FB0AF919790EF902" author="Nicholls, A. G." box="[1235,1390,1711,1734]" pageId="60" pageNumber="61" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation> <bibRefCitation id="8E460A9FF75AFF83FB0AF919790EF902" author="Nicholls, A. G." box="[1235,1390,1711,1734]" pageId="60" pageNumber="61" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</taxonomicName> </taxonomicName>
.......4. .......4.
</keyLead>
</paragraph> </paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF96479FCF8C7" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF96479FDF925" blockId="60.[151,1437,1536,2027]" box="[151,1437,1741,1763]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF96479FDF925" blockId="60.[151,1437,1536,2027]" box="[151,1437,1741,1763]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF96479FDF925" box="[151,1437,1741,1763]" pageId="60" pageNumber="61">
4. Anal somite with paired spinous processes either side of anal operculum............................. 4. Anal somite with paired spinous processes either side of anal operculum.............................
<taxonomicName id="2DD70CEDF75AFF83FAD9F9647937F925" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1280,1367,1741,1762]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="acanthus"> <taxonomicName id="2DD70CEDF75AFF83FAD9F9647937F925" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1280,1367,1741,1762]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF75AFF83FAD9F9647937F925" box="[1280,1367,1741,1762]" italics="true" pageId="60" pageNumber="61">acanthus</emphasis> <emphasis id="D8A3AB7CF75AFF83FAD9F9647937F925" box="[1280,1367,1741,1762]" italics="true" pageId="60" pageNumber="61">acanthus</emphasis>
</taxonomicName> </taxonomicName>
-group. -group.
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF94379FCF8C7" blockId="60.[151,1437,1536,2027]" box="[226,1436,1770,1792]" pageId="60" pageNumber="61">Anal somite without paired spinous processes either side of anal operculum.......................................5.</paragraph> <paragraph id="EA68776EF75AFF83FF3BF94379FCF8C7" blockId="60.[151,1437,1536,2027]" box="[226,1436,1770,1792]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF94379FCF8C7" box="[226,1436,1770,1792]" pageId="60" pageNumber="61">Anal somite without paired spinous processes either side of anal operculum.......................................5.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF8AE79FDF8FC" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF8AE79FDF8DA" blockId="60.[151,1437,1536,2027]" box="[151,1437,1799,1822]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF8AE79FDF8DA" blockId="60.[151,1437,1536,2027]" box="[151,1437,1799,1822]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF8AE79FDF8DA" box="[151,1437,1799,1822]" pageId="60" pageNumber="61">
5. P1 exp-1 without outer spine; distal segment of P1 exopod with three elements; P2 endopod 1-segmented... 5. P1 exp-1 without outer spine; distal segment of P1 exopod with three elements; P2 endopod 1-segmented...
<taxonomicName id="2DD70CEDF75AFF83FB23F8A17936F8DA" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[1274,1366,1800,1821]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="trisetosus"> <taxonomicName id="2DD70CEDF75AFF83FB23F8A17936F8DA" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[1274,1366,1800,1821]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF75AFF83FB23F8A17936F8DA" box="[1274,1366,1800,1821]" italics="true" pageId="60" pageNumber="61">trisetosus</emphasis> <emphasis id="D8A3AB7CF75AFF83FB23F8A17936F8DA" box="[1274,1366,1800,1821]" italics="true" pageId="60" pageNumber="61">trisetosus</emphasis>
</taxonomicName> </taxonomicName>
-group. -group.
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF88C79FDF8FC" blockId="60.[151,1437,1536,2027]" box="[226,1437,1829,1851]" pageId="60" pageNumber="61">P1 exp-1 with outer spine; distal segment of P1 exopod with four elements; P2 endopod 2-segmented..................6.</paragraph> <paragraph id="EA68776EF75AFF83FF3BF88C79FDF8FC" blockId="60.[151,1437,1536,2027]" box="[226,1437,1829,1851]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF88C79FDF8FC" box="[226,1437,1829,1851]" pageId="60" pageNumber="61">P1 exp-1 with outer spine; distal segment of P1 exopod with four elements; P2 endopod 2-segmented..................6.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF8EB79FDF8B2" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF8EB79FDF89F" blockId="60.[151,1437,1536,2027]" box="[151,1437,1858,1880]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF8EB79FDF89F" blockId="60.[151,1437,1536,2027]" box="[151,1437,1858,1880]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF8EB79FDF89F" box="[151,1437,1858,1880]" pageId="60" pageNumber="61">
6. Distal segment of P3 endopod with one apical element............................................ 6. Distal segment of P3 endopod with one apical element............................................
<taxonomicName id="2DD70CEDF75AFF83FADBF8EB7936F890" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1953" box="[1282,1366,1858,1879]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="australis"> <taxonomicName id="2DD70CEDF75AFF83FADBF8EB7936F890" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1953" box="[1282,1366,1858,1879]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="D8A3AB7CF75AFF83FADBF8EB7936F890" box="[1282,1366,1858,1879]" italics="true" pageId="60" pageNumber="61">australis</emphasis> <emphasis id="D8A3AB7CF75AFF83FADBF8EB7936F890" box="[1282,1366,1858,1879]" italics="true" pageId="60" pageNumber="61">australis</emphasis>
</taxonomicName> </taxonomicName>
-group. -group.
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF8C979FDF8B2" blockId="60.[151,1437,1536,2027]" box="[226,1437,1888,1910]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF3BF8C979FDF8B2" blockId="60.[151,1437,1536,2027]" box="[226,1437,1888,1910]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF8C979FDF8B2" box="[226,1437,1888,1910]" pageId="60" pageNumber="61">
Distal segment of P3 endopod with two apical elements....................................... Distal segment of P3 endopod with two apical elements.......................................
<taxonomicName id="2DD70CEDF75AFF83FB08F8C97936F8B2" authorityName="Nicholls" authorityYear="1945" box="[1233,1366,1888,1909]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="spinicaudatus"> <taxonomicName id="2DD70CEDF75AFF83FB08F8C97936F8B2" authorityName="Nicholls" authorityYear="1945" box="[1233,1366,1888,1909]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF75AFF83FB08F8C97936F8B2" box="[1233,1366,1888,1909]" italics="true" pageId="60" pageNumber="61">spinicaudatus</emphasis> <emphasis id="D8A3AB7CF75AFF83FB08F8C97936F8B2" box="[1233,1366,1888,1909]" italics="true" pageId="60" pageNumber="61">spinicaudatus</emphasis>
</taxonomicName> </taxonomicName>
-group. -group.
</keyLead>
</paragraph> </paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF8D479FCF868" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF8D479FDF855" blockId="60.[151,1437,1536,2027]" box="[151,1437,1917,1939]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF8D479FDF855" blockId="60.[151,1437,1536,2027]" box="[151,1437,1917,1939]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF8D479FDF855" box="[151,1437,1917,1939]" pageId="60" pageNumber="61">
7. P1 enp-2 with one geniculate seta and one spine; P3 endopod not modified in 7. P1 enp-2 with one geniculate seta and one spine; P3 endopod not modified in
<specimenCount id="FCD1BCE7F75AFF83FC17F8D47F80F855" box="[974,992,1917,1938]" count="1" pageId="60" pageNumber="61" type="male">♂</specimenCount> <specimenCount id="FCD1BCE7F75AFF83FC17F8D47F80F855" box="[974,992,1917,1938]" count="1" pageId="60" pageNumber="61" type="male">♂</specimenCount>
.. ..
@ -140,9 +179,15 @@ Distal segment of P3 endopod with two apical elements...........................
<bibRefCitation id="8E460A9FF75AFF83FB59F8D479F9F855" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[1152,1433,1917,1939]" pageId="60" pageNumber="61" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">Sak, Huys &amp; Karaytuğ, 2008</bibRefCitation> <bibRefCitation id="8E460A9FF75AFF83FB59F8D479F9F855" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[1152,1433,1917,1939]" pageId="60" pageNumber="61" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">Sak, Huys &amp; Karaytuğ, 2008</bibRefCitation>
</taxonomicName> </taxonomicName>
. .
</keyLead>
</paragraph> </paragraph>
<paragraph id="EA68776EF75AFF83FF3BF83379FCF868" blockId="60.[151,1437,1536,2027]" box="[226,1436,1946,1968]" pageId="60" pageNumber="61">P1 enp-2 with two geniculate setae; P3 endopod sexually dimorphic.............................................8.</paragraph> <paragraph id="EA68776EF75AFF83FF3BF83379FCF868" blockId="60.[151,1437,1536,2027]" box="[226,1436,1946,1968]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF3BF83379FCF868" box="[226,1436,1946,1968]" pageId="60" pageNumber="61">P1 enp-2 with two geniculate setae; P3 endopod sexually dimorphic.............................................8.</keyLead>
</paragraph>
</keyStep>
<keyStep id="51236FCBF75AFF83FF4EF81179FDF82D" pageId="60" pageNumber="61">
<paragraph id="EA68776EF75AFF83FF4EF81179FDF80A" blockId="60.[151,1437,1536,2027]" box="[151,1437,1976,1998]" pageId="60" pageNumber="61"> <paragraph id="EA68776EF75AFF83FF4EF81179FDF80A" blockId="60.[151,1437,1536,2027]" box="[151,1437,1976,1998]" pageId="60" pageNumber="61">
<keyLead id="5126D45BF75AFF83FF4EF81179FDF80A" box="[151,1437,1976,1998]" pageId="60" pageNumber="61">
8. P2 enp-2 with inner serrate seta; P3 endopod 8. P2 enp-2 with inner serrate seta; P3 endopod
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two-segmented...................... two-segmented......................
@ -151,8 +196,10 @@ two-segmented......................
<bibRefCitation id="8E460A9FF75AFF83FACDF81179F9F80A" author="Pennak, RW" box="[1300,1433,1976,1998]" pageId="60" pageNumber="61" pagination="274 - 285" refId="ref48658" refString="Pennak, RW. (1942) Harpacticoid copepods from some intertidal beaches near Woods Hole, Massachusetts. Transactions of the American microscopical Society, 61, 274 - 285. https: // doi. org / 10.2307 / 3222599" type="journal article" year="1942">Pennak, 1942</bibRefCitation> <bibRefCitation id="8E460A9FF75AFF83FACDF81179F9F80A" author="Pennak, RW" box="[1300,1433,1976,1998]" pageId="60" pageNumber="61" pagination="274 - 285" refId="ref48658" refString="Pennak, RW. (1942) Harpacticoid copepods from some intertidal beaches near Woods Hole, Massachusetts. Transactions of the American microscopical Society, 61, 274 - 285. https: // doi. org / 10.2307 / 3222599" type="journal article" year="1942">Pennak, 1942</bibRefCitation>
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P2 enp-2 without inner serrate seta; P3 endopod P2 enp-2 without inner serrate seta; P3 endopod
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one-segmented............ one-segmented............
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100
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<emphasis id="D8A3AB7CF761FFB8FF4EF89B7D72F88B" bold="true" box="[151,274,1842,1868]" pageId="7" pageNumber="8">Diagnosis.</emphasis> <emphasis id="D8A3AB7CF761FFB8FF4EF89B7D72F88B" bold="true" box="[151,274,1842,1868]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Anal somite without paired dorsolateral processes. Anal operculum weakly developed, without rounded medial extension. P1 exp-1 with outer spine; exp-3 with four setae/spines. P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp- 3). P2 endopod 2-segmented; enp-2 with inner seta and two distal spines. P3 endopod 1-segmented; with one distal spine. P4 enp-2 with normally developed outer seta. Anal somite without paired dorsolateral processes. Anal operculum weakly developed, without rounded medial extension. P1 exp-1 with outer spine; exp-3 with four setae/spines. P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp-3). P2 endopod 2-segmented; enp-2 with inner seta and two distal spines. P3 endopod 1-segmented; with one distal spine. P4 enp-2 with normally developed outer seta.
</paragraph> </paragraph>
</subSubSection> </subSubSection>
<subSubSection id="A2CD24E5F76EFFB7FF1EFF3F7E8DFF76" box="[199,749,150,177]" pageId="8" pageNumber="9" type="taxon_list"> <subSubSection id="A2CD24E5F76EFFB7FF1EFF3F7E8DFF76" box="[199,749,150,177]" pageId="8" pageNumber="9" type="taxon_list">
@ -104,7 +104,7 @@ This species group includes one incompletely described African species which doe
<emphasis id="D8A3AB7CF76EFFB7FD08FEAA7F35FEDB" box="[721,853,258,284]" italics="true" pageId="8" pageNumber="9">N. australis</emphasis> <emphasis id="D8A3AB7CF76EFFB7FD08FEAA7F35FEDB" box="[721,853,258,284]" italics="true" pageId="8" pageNumber="9">N. australis</emphasis>
</taxonomicName> </taxonomicName>
( (
<tableCitation id="A75542D5F76EFFB7FCBDFEAB7FDDFEDB" box="[868,957,258,284]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." pageId="8" pageNumber="9">Table 2</tableCitation> <tableCitation id="A75542D5F76EFFB7FCBDFEAB7FDDFEDB" box="[868,957,258,284]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="8" pageNumber="9" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
). More information is required before any statement can be made about its relationships, particularly whether it may be nested as an advanced member in the ). More information is required before any statement can be made about its relationships, particularly whether it may be nested as an advanced member in the
<taxonomicName id="2DD70CEDF76EFFB7FF4EFEE37D53FEA3" authorityName="Nicholls" authorityYear="1945" box="[151,307,330,356]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinicaudatus"> <taxonomicName id="2DD70CEDF76EFFB7FF4EFEE37D53FEA3" authorityName="Nicholls" authorityYear="1945" box="[151,307,330,356]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76EFFB7FF4EFEE37D53FEA3" box="[151,307,330,356]" italics="true" pageId="8" pageNumber="9">spinicaudatus</emphasis> <emphasis id="D8A3AB7CF76EFFB7FF4EFEE37D53FEA3" box="[151,307,330,356]" italics="true" pageId="8" pageNumber="9">spinicaudatus</emphasis>

100
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<mods:affiliation id="A9D4B9C25BD955B546941DBA37604D92">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation> <mods:affiliation id="5CF3DB451207F25F9EB6123262227D13">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
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<mods:affiliation id="DCD43AFA90F691DD8C4E0C424683CCCB">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation> <mods:affiliation id="9DACC5E5F0E168FBCB50EE75ACF80068">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation>
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<emphasis id="D8A3AB7CF761FFB8FF66FBB27D3BFBF2" box="[191,347,1051,1077]" italics="true" pageId="7" pageNumber="8">spinicaudatus</emphasis> <emphasis id="D8A3AB7CF761FFB8FF66FBB27D3BFBF2" box="[191,347,1051,1077]" italics="true" pageId="7" pageNumber="8">spinicaudatus</emphasis>
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<emphasis id="D8A3AB7CF761FFB8FACCFA0E79FCFA06" box="[1301,1436,1447,1473]" italics="true" pageId="7" pageNumber="8">N. pacificus</emphasis> <emphasis id="D8A3AB7CF761FFB8FACCFA0E79FCFA06" box="[1301,1436,1447,1473]" italics="true" pageId="7" pageNumber="8">N. pacificus</emphasis>
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which lacks the inner seta on P4 exp-3 ( which lacks the inner seta on P4 exp-3 (
<tableCitation id="A75542D5F761FFB8FD81FA627ED0FA22" box="[600,688,1483,1509]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." pageId="7" pageNumber="8">Table 2</tableCitation> <tableCitation id="A75542D5F761FFB8FD81FA627ED0FA22" box="[600,688,1483,1509]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="7" pageNumber="8" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
). In the absence of any morphology-based apomorphies that could support the common origin of these species, assessment of the potential monophyly of this species group will have to await the arrival of molecular sequence data. ). In the absence of any morphology-based apomorphies that could support the common origin of these species, assessment of the potential monophyly of this species group will have to await the arrival of molecular sequence data.
<bibRefCitation id="8E460A9FF761FFB8FD48F9BA7F2BF9EA" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[657,843,1555,1581]" pageId="7" pageNumber="8" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008"> <bibRefCitation id="8E460A9FF761FFB8FD48F9BA7F2BF9EA" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[657,843,1555,1581]" pageId="7" pageNumber="8" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak Sak

106
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@ -1,71 +1,71 @@
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<mods:affiliation id="BF3FB452223672DA356D4FBA520170DF">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation> <mods:affiliation id="1D6B9C259C76877AF08BD15E7DE7E63E">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
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). P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp-3). Endopod P2–P3 2-segmented. P2 enp-2 with or without inner seta; with two distal spines. P3 enp-2 with 1–2 distal spines. P4 enp-2 with well developed outer seta. ). P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp-3). Endopod P2–P3 2-segmented. P2 enp-2 with or without inner seta; with two distal spines. P3 enp-2 with 1–2 distal spines. P4 enp-2 with well developed outer seta.
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<emphasis id="D8A3AB7CF760FFB9FF41FF3E7D70FF68" bold="true" box="[152,272,151,175]" pageId="6" pageNumber="7">TABLE 1.</emphasis> <emphasis id="D8A3AB7CF760FFB9FF41FF3E7D70FF68" bold="true" box="[152,272,151,175]" pageId="6" pageNumber="7">TABLE 1.</emphasis>
Valid species and Valid species and
@ -511,7 +511,7 @@ Venanzetti, 1994)
<subSubSection id="A2CD24E5F760FFB9FF1EF9CD781FF9B9" box="[199,1151,1636,1662]" pageId="6" pageNumber="7" type="multiple"> <subSubSection id="A2CD24E5F760FFB9FF1EF9CD781FF9B9" box="[199,1151,1636,1662]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="EA68776EF760FFB9FF1EF9CD781FF9B9" blockId="6.[151,1437,1636,2023]" box="[199,1151,1636,1662]" pageId="6" pageNumber="7"> <paragraph id="EA68776EF760FFB9FF1EF9CD781FF9B9" blockId="6.[151,1437,1636,2023]" box="[199,1151,1636,1662]" pageId="6" pageNumber="7">
<emphasis id="D8A3AB7CF760FFB9FF1EF9CD7DDDF9B9" bold="true" box="[199,445,1636,1662]" pageId="6" pageNumber="7">Subgroups included.</emphasis> <emphasis id="D8A3AB7CF760FFB9FF1EF9CD7DDDF9B9" bold="true" box="[199,445,1636,1662]" pageId="6" pageNumber="7">Subgroups included.</emphasis>
<taxonomicName id="2DD70CEDF760FFB9FE1DF9CD7E49F9B9" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[452,553,1636,1662]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acanthus"> <taxonomicName id="2DD70CEDF760FFB9FE1DF9CD7E49F9B9" ID-CoL="73VZK" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[452,553,1636,1662]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF760FFB9FE1DF9CD7E49F9B9" box="[452,553,1636,1662]" italics="true" pageId="6" pageNumber="7">acanthus</emphasis> <emphasis id="D8A3AB7CF760FFB9FE1DF9CD7E49F9B9" box="[452,553,1636,1662]" italics="true" pageId="6" pageNumber="7">acanthus</emphasis>
</taxonomicName> </taxonomicName>
-subgroup, -subgroup,
@ -609,7 +609,7 @@ and
) )
</taxonomicName> </taxonomicName>
( (
<tableCitation id="A75542D5F761FFB8FC4BFF3E7F8BFF76" box="[914,1003,151,177]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." pageId="7" pageNumber="8">Table 2</tableCitation> <tableCitation id="A75542D5F761FFB8FC4BFF3E7F8BFF76" box="[914,1003,151,177]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="7" pageNumber="8" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
). ).
<bibRefCitation id="8E460A9FF761FFB8FBD9FF3E7887FF76" author="Wells, J. B. J. &amp; Rao, G. C." box="[1024,1255,151,177]" pageId="7" pageNumber="8" pagination="1 - 385" refId="ref50123" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao (1987)</bibRefCitation> <bibRefCitation id="8E460A9FF761FFB8FBD9FF3E7887FF76" author="Wells, J. B. J. &amp; Rao, G. C." box="[1024,1255,151,177]" pageId="7" pageNumber="8" pagination="1 - 385" refId="ref50123" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao (1987)</bibRefCitation>
, inspired by the extensive variability observed in their material of , inspired by the extensive variability observed in their material of
@ -743,7 +743,7 @@ and
<emphasis id="D8A3AB7CF761FFB8FBE8FCB67882FCFE" box="[1073,1250,799,825]" italics="true" pageId="7" pageNumber="8">N. reductaspina</emphasis> <emphasis id="D8A3AB7CF761FFB8FBE8FCB67882FCFE" box="[1073,1250,799,825]" italics="true" pageId="7" pageNumber="8">N. reductaspina</emphasis>
</taxonomicName> </taxonomicName>
share the unique absence of the inner seta on P2 enp-2 ( share the unique absence of the inner seta on P2 enp-2 (
<tableCitation id="A75542D5F761FFB8FD9EFCEA7EFEFC9A" box="[583,670,835,861]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." pageId="7" pageNumber="8">Table 2</tableCitation> <tableCitation id="A75542D5F761FFB8FD9EFCEA7EFEFC9A" box="[583,670,835,861]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="7" pageNumber="8" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
), giving further credence to the monophyly of this subgroup. ), giving further credence to the monophyly of this subgroup.
</paragraph> </paragraph>
<paragraph id="EA68776EF761FFB8FF1EFCCE7F2BFC2A" blockId="7.[151,1437,150,1005]" pageId="7" pageNumber="8"> <paragraph id="EA68776EF761FFB8FF1EFCCE7F2BFC2A" blockId="7.[151,1437,150,1005]" pageId="7" pageNumber="8">

284
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<mods:title id="EA481019B666E476930355D1C9366B60">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
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Genus
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<emphasis id="D8A3AB7CF763FFBAFF37FE8E7DC5FE85" bold="true" box="[238,421,295,322]" italics="true" pageId="5" pageNumber="6">Neoleptastacus</emphasis>
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<emphasis id="D8A3AB7CF763FFBAFF4EFEC57D58FE44" box="[151,312,364,387]" italics="true" pageId="5" pageNumber="6">Pararenopontia</emphasis>
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(
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species by original designation:
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<bibRefCitation id="8E460A9FF763FFBAFB26FEC579E8FE44" author="Mielke, W." box="[1279,1416,363,388]" pageId="5" pageNumber="6" pagination="1 - 134" refId="ref48153" refString="Mielke, W. (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens, 52, 1 - 134." type="journal article" year="1975">Mielke, 1975</bibRefCitation>
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)
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<emphasis id="D8A3AB7CF763FFBAFF4EFE187D6AFE0C" bold="true" box="[151,266,433,459]" pageId="5" pageNumber="6">Diagnosis</emphasis>
(adapted from
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Sak
<emphasis id="D8A3AB7CF763FFBAFE20FE1A7E56FE0B" box="[505,566,434,460]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2008
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).
<taxonomicName id="2DD70CEDF763FFBAFD4AFE1B7F25FE0B" authorityName="Martinez Arbizu &amp; Moura" authorityYear="1994" box="[659,837,434,460]" class="Copepoda" family="Arenopontiidae" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Arenopontiidae</taxonomicName>
. Urosomites occasionally with conspicuous surface ornamentation (
<taxonomicName id="2DD70CEDF763FFBAFE95FE7E7DA8FE37" authorityName="Mielke" authorityYear="1985" box="[332,456,470,496]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="clasingi">
<emphasis id="D8A3AB7CF763FFBAFE95FE7E7DA8FE37" box="[332,456,470,496]" italics="true" pageId="5" pageNumber="6">N. clasingi</emphasis>
</taxonomicName>
,
<taxonomicName id="2DD70CEDF763FFBAFE00FE7E7E3DFE37" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[473,605,471,496]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ornamentus">
<emphasis id="D8A3AB7CF763FFBAFE00FE7E7E3DFE37" box="[473,605,471,496]" italics="true" pageId="5" pageNumber="6">ornamentus</emphasis>
</taxonomicName>
,
<taxonomicName id="2DD70CEDF763FFBAFDB7FE7F7E9FFE37" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1987" box="[622,767,470,496]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reductaspina">
<emphasis id="D8A3AB7CF763FFBAFDB7FE7F7E9FFE37" box="[622,767,470,496]" italics="true" pageId="5" pageNumber="6">reductaspina</emphasis>
</taxonomicName>
). Anal somite with (
<taxonomicName id="2DD70CEDF763FFBAFC28FE7F7836FE37" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1009,1110,470,496]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF763FFBAFC28FE7F7836FE37" box="[1009,1110,470,496]" italics="true" pageId="5" pageNumber="6">acanthus</emphasis>
</taxonomicName>
-group) or without (all other species groups) paired dorsolateral spinous processes. Anal operculum sometimes with median extension. Hyaline frills of abdominal somites with rectangular digitate or non-digitate lappets. Caudal ramus usually with dorsolateral spur near medial margin. P1 exopod 2- or 3-segmented; exp-1 with/without outer spine; exp-3 (or exp-2 when exopod 2-segmented) with one–two spine(s) and two geniculate setae. P1 endopod not prehensile, at least as long as exopod; enp-2 with outer spine and inner geniculate seta at distal margin. P2–P3 endopods 1- or 2-segmented. P3 endopod with outer distal element (when present) usually fused at base. P4 enp-2 with well developed outer distal element (except in
<taxonomicName id="2DD70CEDF763FFBAFEB4FD7A7DB8FD2B" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[365,472,723,748]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF763FFBAFEB4FD7A7DB8FD2B" box="[365,472,723,748]" italics="true" pageId="5" pageNumber="6">trisetosus</emphasis>
</taxonomicName>
-group). Armature formula as follows:
</paragraph>
<paragraph id="EA68776EF763FFBAFE6EFCB37F16FC2F" pageId="5" pageNumber="6">
<table id="98D785CEF7630040FF1EFCB37842FC2F" box="[199,1058,794,1000]" gridcols="3" gridrows="5" inLine="true" pageId="5" pageNumber="6">
<tr id="54E7752CF7630040FF1EFCB37842FCF3" box="[199,1058,794,820]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1">
<th id="17361C50F7630040FE6EFCB37EFEFCF3" box="[439,670,794,820]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Exopod</th>
<th id="17361C50F7630040FCEEFCB37842FCF3" box="[823,1058,794,820]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">Endopod</th>
</tr>
<tr id="54E7752CF7630040FF1EFCCB7842FCBB" box="[199,1058,866,892]" gridrow="1" pageId="5" pageNumber="6">
<th id="17361C50F7630040FF1EFCCB7C85FCBB" box="[199,229,866,892]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">P1</th>
<td id="17361C50F7630040FE6EFCCB7EFEFCBB" box="[439,670,866,892]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">
0.0.02(1–2) or
<geoCoordinate id="8FE311A9F763FFBAFD86FCCB7EFEFCBB" box="[607,670,866,892]" degrees="0.021" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.021">0.021</geoCoordinate>
</td>
<td id="17361C50F7630040FCEEFCCB7842FCBB" box="[823,1058,866,892]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">
<geoCoordinate id="8FE311A9F763FFBAFCEEFCCB7F15FCBB" box="[823,885,866,892]" degrees="1.011" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="1.011">1.011</geoCoordinate>
</td>
</tr>
<tr id="54E7752CF7630040FF1EFC2F7842FC67" box="[199,1058,902,928]" gridrow="2" pageId="5" pageNumber="6">
<th id="17361C50F7630040FF1EFC2F7C85FC67" box="[199,229,902,928]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">P2</th>
<td id="17361C50F7630040FE6EFC2F7EFEFC67" box="[439,670,902,928]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">0.0.021</td>
<td id="17361C50F7630040FCEEFC2F7842FC67" box="[823,1058,902,928]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">0.(0–1)(1–2)0 or 110</td>
</tr>
<tr id="54E7752CF7630040FF1EFC037842FC03" box="[199,1058,938,964]" gridrow="3" pageId="5" pageNumber="6">
<th id="17361C50F7630040FF1EFC037C85FC03" box="[199,229,938,964]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">P3</th>
<td id="17361C50F7630040FE6EFC037EFEFC03" box="[439,670,938,964]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">0.0.021</td>
<td id="17361C50F7630040FCEEFC037842FC03" box="[823,1058,938,964]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">0.0(1–2)0 or 010</td>
</tr>
<tr id="54E7752CF7630040FF1EFC677842FC2F" box="[199,1058,974,1000]" gridrow="4" pageId="5" pageNumber="6">
<th id="17361C50F7630040FF1EFC677C85FC2F" box="[199,229,974,1000]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">P4</th>
<td id="17361C50F7630040FE6EFC677EFEFC2F" box="[439,670,974,1000]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">0.0.(0–1)21</td>
<td id="17361C50F7630040FCEEFC677842FC2F" box="[823,1058,974,1000]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">0.020</td>
</tr>
</table>
</paragraph>
<paragraph id="EA68776EF763FFBAFF4EFBBF7FC2FBBF" blockId="5.[151,1437,1046,1720]" pageId="5" pageNumber="6">
P3 endopod
<specimenCount id="FCD1BCE7F763FFBAFEFAFBBF7D58FBF7" box="[291,312,1046,1072]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
not sexually dimorphic. P5 with outer basal seta and one–four discrete elements; innermost element fused to segment forming spinous process (weakly delimited in
<taxonomicName id="2DD70CEDF763FFBAFC84FB927F88FB93" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[861,1000,1083,1108]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF763FFBAFC84FB927F88FB93" box="[861,1000,1083,1108]" italics="true" pageId="5" pageNumber="6">N. trisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF763FFBAFBC7FB9278A2FB93" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1054,1218,1083,1108]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="D8A3AB7CF763FFBAFBC7FB9278A2FB93" box="[1054,1218,1083,1108]" italics="true" pageId="5" pageNumber="6">N. panamensis</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF763FFBAFB11FB937941FB93" bold="true" box="[1224,1313,1082,1108]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="C3901607F763FFBAFB11FB937941FB93" box="[1224,1313,1082,1108]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); length of process sometimes sexually dimorphic. P
<specimenCount id="FCD1BCE7F763FFBAFDBEFBF77EF0FBBF" box="[615,656,1118,1144]" count="6" pageId="5" pageNumber="6" type="male">6 ♂</specimenCount>
with one–two elements.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F763FFBAFF1EFB2B7879FB5B" box="[199,1049,1153,1181]" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="EA68776EF763FFBAFF1EFB2B7879FB5B" blockId="5.[151,1437,1046,1720]" box="[199,1049,1153,1181]" pageId="5" pageNumber="6">
<emphasis id="D8A3AB7CF763FFBAFF1EFB2B7D02FB5C" bold="true" box="[199,354,1153,1180]" pageId="5" pageNumber="6">
<typeStatus id="356CC9CCF763FFBAFF1EFB2B7D62FB5B" box="[199,258,1154,1180]" pageId="5" pageNumber="6">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="2DD70CEDF763FFBAFEB0FB2B7F03FB5A" ID-CoL="46KMW" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[361,867,1154,1181]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF763FFBAFEB0FB2B7ED7FB5B" box="[361,695,1154,1180]" italics="true" pageId="5" pageNumber="6">Neoleptastacus spinicaudatus</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFD67FB2B7F03FB5A" author="Nicholls, A. G." box="[702,867,1154,1181]" pageId="5" pageNumber="6" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</taxonomicName>
[by monotypy].
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F763FFBAFF1EFB0C7CBFFA97" pageId="5" pageNumber="6" type="taxon_list">
<paragraph id="EA68776EF763FFBAFF1EFB0C7EC1FB07" blockId="5.[151,1437,1046,1720]" box="[199,673,1189,1216]" pageId="5" pageNumber="6">
<emphasis id="D8A3AB7CF763FFBAFF1EFB0C7D0FFB78" bold="true" box="[199,367,1189,1215]" pageId="5" pageNumber="6">Other species.</emphasis>
Twenty-three. See
<tableCitation id="A75542D5F763FFBAFD9EFB0F7EFBFB07" box="[583,667,1190,1216]" captionStart="TABLE 1" captionStartId="6.[152,245,151,175]" captionTargetBox="[151,1423,258,1518]" captionText="TABLE 1. Valid species and species inquirendae (indicated by *) of Neoleptastacus Nicholls, 1945 and Phreatipontia gen. nov." httpUri="http://table.plazi.org/id/BEA827E6F760FFB9FF41FF3E7C9CFF13" pageId="5" pageNumber="6" tableUuid="BEA827E6F760FFB9FF41FF3E7C9CFF13">Table 1</tableCitation>
.
</paragraph>
<paragraph id="EA68776EF763FFBAFF1EFB637965FB22" blockId="5.[151,1437,1046,1720]" box="[199,1285,1225,1253]" pageId="5" pageNumber="6">
Species inquirenda.
<taxonomicName id="2DD70CEDF763FFBAFE68FB637FA0FB22" ID-CoL="46KMT" authority="Krishnaswamy, 1957" authorityName="Krishnaswamy" authorityYear="1957" box="[433,960,1226,1253]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="D8A3AB7CF763FFBAFE68FB637EAAFB23" box="[433,714,1226,1252]" italics="true" pageId="5" pageNumber="6">Neoleptastacus secundus</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFD08FB637FA0FB22" author="Krishnaswamy, S." box="[721,960,1226,1253]" pageId="5" pageNumber="6" refId="ref47590" refString="Krishnaswamy, S. (1957) Studies on the Copepoda of Madras. PhD dissertation, University of Madras, Chennai, Tamil Nadu, 168 pp." type="book" year="1957">Krishnaswamy, 1957</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2DD70CEDF763FFBAFC15FB627961FB22" ID-CoL="46KMB" authority="(Lang, 1965)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[972,1281,1226,1253]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF763FFBAFC15FB627805FB23" box="[972,1125,1227,1252]" italics="true" pageId="5" pageNumber="6">N. accraensis</emphasis>
(
<bibRefCitation id="8E460A9FF763FFBAFBAAFB637898FB22" author="Lang, K." box="[1139,1272,1226,1253]" pageId="5" pageNumber="6" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="EA68776EF763FFBAFF1EFB477CBFFA97" blockId="5.[151,1437,1046,1720]" pageId="5" pageNumber="6">
<emphasis id="D8A3AB7CF763FFBAFF1EFB477D99FAC0" bold="true" box="[199,505,1261,1288]" pageId="5" pageNumber="6">Taxa of doubtful identity.</emphasis>
<taxonomicName id="2DD70CEDF763FFBAFDD8FB4778EDFACF" ID-CoL="GFPY" authority="Chappuis, 1954 sensu Bozic (1967)" authorityName="Chappuis, 1954 sensu Bozic" authorityYear="1967" box="[513,1165,1262,1289]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF763FFBAFDD8FB477E8EFACF" box="[513,750,1262,1288]" italics="true" pageId="5" pageNumber="6">Arenopontia acantha</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFD2EFB477FC8FACE" author="Chappuis, P. - A." box="[759,936,1262,1289]" pageId="5" pageNumber="6" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
<emphasis id="D8A3AB7CF763FFBAFC69FB467F8EFACF" box="[944,1006,1263,1288]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFC2FFB4778EDFACF" author="Bozic, B." box="[1014,1165,1262,1288]" pageId="5" pageNumber="6" pagination="869 - 873" refId="ref45435" refString="Bozic, B. (1967) Deux Copepodes Harpacticoides de l'Inde. Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 38, 869 - 873." type="journal article" year="1967">Božić (1967)</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2DD70CEDF763FFBAFB43FB477E2BFAEB" ID-CoL="5W3DL" authority="Cottarelli, 1973 a sensu Mielke (1982 b)" authorityName="Cottarelli, 1973 a sensu Mielke" authorityYear="1982" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gussoae">
<emphasis id="D8A3AB7CF763FFBAFB43FB477945FACF" box="[1178,1317,1262,1288]" italics="true" pageId="5" pageNumber="6">Arenopontia</emphasis>
?
<emphasis id="D8A3AB7CF763FFBAFA9BFB4679FCFACF" box="[1346,1436,1263,1288]" italics="true" pageId="5" pageNumber="6">gussoae</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFF4EFABB7D32FAEB" author="Cottarelli V." box="[151,338,1298,1324]" pageId="5" pageNumber="6" pagination="49 - 59" refId="ref45971" refString="Cottarelli V. (1973 a) Arenopontia gussoae n. sp., nuovo arpacticoide di acque interstiziali litorali dell'isola di Cuba (Crustacea, Copepoda). Fragmenta entomologica, 9, 49 - 59." type="journal article" year="1973">Cottarelli, 1973a</bibRefCitation>
<emphasis id="D8A3AB7CF763FFBAFE81FABA7DF6FAEB" box="[344,406,1299,1324]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFE44FABB7E2BFAEB" author="Mielke, W." box="[413,587,1298,1324]" pageId="5" pageNumber="6" pagination="199 - 207" refId="ref48211" refString="Mielke, W. (1982 b) Three variable Arenopontia species (Crustacea, Copepoda) from Panama. Zoologica Scripta, 11, 199 - 207. https: // doi. org / 10.1111 / j. 1463 - 6409.1982. tb 00533. x" type="journal article" year="1982">Mielke (1982b)</bibRefCitation>
</taxonomicName>
[partim];
<taxonomicName id="2DD70CEDF763FFBAFD63FABB7903FAEB" ID-CoL="5W3DL" authority="Cottarelli, 1973 a sensu Mielke (1987)" authorityName="Cottarelli, 1973 a sensu Mielke" authorityYear="1987" box="[698,1379,1298,1324]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gussoae">
<emphasis id="D8A3AB7CF763FFBAFD63FABB7F25FAEB" box="[698,837,1298,1324]" italics="true" pageId="5" pageNumber="6">Arenopontia</emphasis>
?
<emphasis id="D8A3AB7CF763FFBAFC84FABA7FD7FAEB" box="[861,951,1299,1324]" italics="true" pageId="5" pageNumber="6">gussoae</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFC67FABB7818FAEB" author="Cottarelli V." box="[958,1144,1298,1324]" pageId="5" pageNumber="6" pagination="49 - 59" refId="ref45971" refString="Cottarelli V. (1973 a) Arenopontia gussoae n. sp., nuovo arpacticoide di acque interstiziali litorali dell'isola di Cuba (Crustacea, Copepoda). Fragmenta entomologica, 9, 49 - 59." type="journal article" year="1973">Cottarelli, 1973a</bibRefCitation>
<emphasis id="D8A3AB7CF763FFBAFBA6FABA78DDFAEB" box="[1151,1213,1299,1324]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
<bibRefCitation id="8E460A9FF763FFBAFB1AFABB7903FAEB" author="Mielke, W." box="[1219,1379,1298,1324]" pageId="5" pageNumber="6" pagination="309 - 361" refId="ref48334" refString="Mielke, W. (1987) Interstitielle Copepoda von Nord- und Sud-Chile. Microfauna marina, 3, 309 - 361." type="journal article" year="1987">Mielke (1987)</bibRefCitation>
</taxonomicName>
[2nd form].
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F763FFBAFF1EFAF078F2F97F" pageId="5" pageNumber="6" type="description">
<paragraph id="EA68776EF763FFBAFF1EFAF07DC9FAB3" blockId="5.[151,1437,1046,1720]" box="[199,425,1369,1396]" pageId="5" pageNumber="6">
<emphasis id="D8A3AB7CF763FFBAFF1EFAF07D46FAB4" bold="true" box="[199,294,1369,1395]" pageId="5" pageNumber="6">Gender.</emphasis>
Masculine.
</paragraph>
<paragraph id="EA68776EF763FFBAFF1EFAD778F2F97F" blockId="5.[151,1437,1046,1720]" pageId="5" pageNumber="6">
The genus shows considerable variation in swimming leg armature and segmentation, P5 morphology, abdominal ornamentation and caudal ramus structure. However, all species display a spinous apical process on the P5 which is derived from the fused innermost element (weakly delimited in
<taxonomicName id="2DD70CEDF763FFBAFC5CFA6E7872FA27" ID-CoL="46KMX" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[901,1042,1479,1504]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF763FFBAFC5CFA6E7872FA27" box="[901,1042,1479,1504]" italics="true" pageId="5" pageNumber="6">N. trisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF763FFBAFB93FA6E7890FA27" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" box="[1098,1264,1479,1504]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="D8A3AB7CF763FFBAFB93FA6E7890FA27" box="[1098,1264,1479,1504]" italics="true" pageId="5" pageNumber="6">N. panamensis</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF763FFBAFB21FA6F7933FA27" bold="true" box="[1272,1363,1478,1504]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="C3901607F763FFBAFB21FA6F7933FA27" box="[1272,1363,1478,1504]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). This apomorphic character state is unique in the
<taxonomicName id="2DD70CEDF763FFBAFD48FA437F5FF9C3" ID-CoL="7NHD6" authorityName="Martinez Arbizu &amp; Moura" authorityYear="1994" box="[657,831,1514,1540]" class="Copepoda" family="Arenopontiidae" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Arenopontiidae</taxonomicName>
and potentially serves as evidence in support of the monophyly of the genus. For practical reasons we have recognized several species groups within
<taxonomicName id="2DD70CEDF763FFBAFB33F9A779F5F9EF" ID-CoL="7PQ3W" authorityName="Nicholls" authorityYear="1945" box="[1258,1429,1550,1576]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="D8A3AB7CF763FFBAFB33F9A779F5F9EF" box="[1258,1429,1550,1576]" italics="true" pageId="5" pageNumber="6">Neoleptastacus</emphasis>
</taxonomicName>
, some of which are monophyletic and may eventually be attributed separate generic status (
<tableCitation id="A75542D5F763FFBAFB45F99B7895F98B" box="[1180,1269,1586,1612]" captionStart="TABLE 2" captionStartId="9.[152,245,151,175]" captionTargetPageId="9" captionText="TABLE 2. Species groups in Neoleptastacus Nicholls, 1945 sensu lato. [AP = paired spinous processes on anal somite; G+G = two geniculate setae; G+S = outer spine + inner geniculate seta; P5 = number of marginal elements (excluding outer basal seta) in both sexes (when known); p = spinous process; + = present; – = absent;? = missing data]." httpUri="http://table.plazi.org/id/BEA827E6F76FFFB6FF41FF3E7947FF30" pageId="5" pageNumber="6" tableUuid="BEA827E6F76FFFB6FF41FF3E7947FF30">Table 2</tableCitation>
). The majority of these groups are characterized by one or several autapomorphic character states with the notable exception of the
<taxonomicName id="2DD70CEDF763FFBAFF4EF9D37D53F953" ID-CoL="46KMW" authorityName="Nicholls" authorityYear="1945" box="[151,307,1658,1684]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF763FFBAFF4EF9D37D53F953" box="[151,307,1658,1684]" italics="true" pageId="5" pageNumber="6">spinicaudatus</emphasis>
</taxonomicName>
-group which contains the
<typeStatus id="356CC9CCF763FFBAFD80F9D37EE9F953" box="[601,649,1658,1684]" pageId="5" pageNumber="6">type</typeStatus>
species. For this reason, we have refrained from proposing new genera (except for the
<taxonomicName id="2DD70CEDF763FFBAFE98F9377DD1F97F" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Cottarelli, Bruno &amp; Venanzetti" baseAuthorityYear="1994" box="[321,433,1694,1720]" class="Copepoda" family="Arenopontiidae" genus="Phreatipontia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="speluncae">
<emphasis id="D8A3AB7CF763FFBAFE98F9377DD1F97F" box="[321,433,1694,1720]" italics="true" pageId="5" pageNumber="6">speluncae</emphasis>
</taxonomicName>
-group) as this would render
<taxonomicName id="2DD70CEDF763FFBAFD2FF9377FC1F97F" ID-CoL="7PQ3W" authorityName="Nicholls" authorityYear="1945" box="[758,929,1694,1720]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="D8A3AB7CF763FFBAFD2FF9377FC1F97F" box="[758,929,1694,1720]" italics="true" pageId="5" pageNumber="6">Neoleptastacus</emphasis>
</taxonomicName>
a paraphyletic taxon.
</paragraph>
</subSubSection>
</treatment>
</document>

303
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<mods:title id="0BADFEE59D4588C38D620C6528C65A5D">Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov.</mods:title>
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<mods:affiliation id="C72C315DE99A60CCF4E7F2205AC05A7B">Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162</mods:affiliation>
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<mods:affiliation id="46B9D832929BB4D5C09AABD0013FC5E9">Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133</mods:affiliation>
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<mods:affiliation id="4E1EEB45AE202BD163F2F002B00FF69B">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk</mods:affiliation>
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<paragraph id="EA68776EF76CFFB5FF4EFD187ED3FD0B" blockId="10.[151,691,689,716]" box="[151,691,689,716]" pageId="10" pageNumber="11">
<heading id="B120C002F76CFFB5FF4EFD187ED3FD0B" bold="true" box="[151,691,689,716]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="3">
<taxonomicName id="2DD70CEDF76CFFB5FF4EFD187ED3FD0B" ID-CoL="46KMW" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[151,691,689,716]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FF4EFD187ED3FD0B" bold="true" box="[151,691,689,716]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF4EFD187D9DFD0B" bold="true" box="[151,509,689,716]" italics="true" pageId="10" pageNumber="11">Neoleptastacus spinicaudatus</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FDDDFD187ED3FD0B" author="Nicholls, A. G." box="[516,691,689,716]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</emphasis>
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<paragraph id="EA68776EF76CFFB5FF4EFD5F7867FCC9" blockId="10.[151,1031,758,874]" box="[151,1031,758,782]" pageId="10" pageNumber="11">
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<taxonomicName id="2DD70CEDF76CFFB5FF4EFD5F7E0EFCC9" ID-CoL="7WX6G" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[151,622,758,782]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FF4EFD5F7DAFFCC9" box="[151,463,758,782]" italics="true" pageId="10" pageNumber="11">Paraleptastacus spinicaudatus</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FE0CFD5F7E0EFCC9" author="Nicholls, A. G." box="[469,622,758,782]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</taxonomicName>
:
<emphasis id="D8A3AB7CF76CFFB5FDA1FD5F7F64FCC9" box="[632,772,758,782]" italics="true" pageId="10" pageNumber="11">lapsus calami</emphasis>
by
<treatmentCitation id="6B76517FF76CFFB5FCF3FD5F7867FCC9" author="Chappuis, P. - A." box="[810,1031,758,782]" page="269" pageId="10" pageNumber="11" year="1954">
<bibRefCitation id="8E460A9FF76CFFB5FCF3FD5F7867FCC9" author="Chappuis, P. - A." box="[810,1031,758,782]" pageId="10" pageNumber="11" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis (1954: 269)</bibRefCitation>
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<paragraph id="EA68776EF76CFFB5FF4EFCBC7F41FCEA" blockId="10.[151,1031,758,874]" box="[151,801,789,813]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="CAC75040F76CFFB5FF4EFCBC7F41FCEA" box="[151,801,789,813]" pageId="10" pageNumber="11">
<treatmentCitation id="6B76517FF76CFFB5FF4EFCBC7F41FCEA" author="Chappuis P. - A." box="[151,801,789,813]" page=")" pageId="10" pageNumber="11" year="1955">
<taxonomicName id="2DD70CEDF76CFFB5FF4EFCBC7F41FCEA" ID-CoL="GFQW" authority="(Nicholls, 1945) Chappuis (1955: 56)" authorityName="Chappuis" authorityPageNumber="56" authorityYear="1955" baseAuthorityName="Nicholls" baseAuthorityYear="1945" box="[151,801,789,813]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudata">
<emphasis id="D8A3AB7CF76CFFB5FF4EFCBC7DFEFCEA" box="[151,414,789,813]" italics="true" pageId="10" pageNumber="11">Arenopontia spinicaudata</emphasis>
(
<bibRefCitation id="8E460A9FF76CFFB5FE72FCBC7E23FCEA" author="Nicholls, A. G." box="[427,579,789,813]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
)
<bibRefCitation id="8E460A9FF76CFFB5FD88FCBC7F41FCEA" author="Chappuis P. - A." box="[593,801,789,813]" pageId="10" pageNumber="11" pagination="45 - 73" refId="ref45639" refString="Chappuis P. - A. (1955) Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar. IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut scientifique de Madagascar, Series A, 9, 45 - 73." type="journal article" year="1955">Chappuis (1955: 56)</bibRefCitation>
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<treatmentCitationGroup id="CAC75040F76CFFB5FF4EFC9D7FA0FC8B" box="[151,960,819,844]" pageId="10" pageNumber="11">
<taxonomicName id="2DD70CEDF76CFFB5FF4EFC9D7F60FC8C" ID-CoL="GFQW" authority="(Nicholls, 1945)" authorityName="Chappuis" authorityYear="1955" baseAuthorityName="Nicholls" baseAuthorityYear="1945" box="[151,768,819,844]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudata" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF76CFFB5FF4EFC9D7D75FC8C" box="[151,277,820,843]" italics="true" pageId="10" pageNumber="11">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF76CFFB5FEFDFC9A7DDFFC8C" box="[292,447,819,843]" italics="true" pageId="10" pageNumber="11">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF76CFFB5FE14FC9A7E30FC8C" box="[461,592,819,843]" italics="true" pageId="10" pageNumber="11">spinicaudata</emphasis>
(
<bibRefCitation id="8E460A9FF76CFFB5FD84FC9A7E98FC8C" author="Nicholls, A. G." box="[605,760,819,843]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="6B76517FF76CFFB5FCD0FC9A7FA0FC8B" author="Wells, J. B. J." box="[777,960,819,844]" page="324" pageId="10" pageNumber="11" year="1967">
<bibRefCitation id="8E460A9FF76CFFB5FCD0FC9A7FA0FC8B" author="Wells, J. B. J." box="[777,960,819,844]" pageId="10" pageNumber="11" pagination="189 - 358" refId="ref49859" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / S 0080456800024017" type="journal article" year="1967">Wells (1967: 324)</bibRefCitation>
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<paragraph id="EA68776EF76CFFB5FF4EFCFB7F89FCAD" blockId="10.[151,1031,758,874]" box="[151,1001,850,874]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="CAC75040F76CFFB5FF4EFCFB7F89FCAD" box="[151,1001,850,874]" pageId="10" pageNumber="11">
<taxonomicName id="2DD70CEDF76CFFB5FF4EFCFB7E4DFCAD" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[151,557,850,874]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="D8A3AB7CF76CFFB5FF4EFCFB7DEEFCAD" box="[151,398,850,874]" italics="true" pageId="10" pageNumber="11">Neoleptastacus longipes</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FE4DFCFB7E4DFCAD" author="Nicholls, A. G." box="[404,557,850,874]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</taxonomicName>
:
<emphasis id="D8A3AB7CF76CFFB5FDEEFCFB7EA3FCAD" box="[567,707,850,874]" italics="true" pageId="10" pageNumber="11">lapsus calami</emphasis>
by
<treatmentCitation id="6B76517FF76CFFB5FD30FCFB7FCFFCAD" author="Karanovic, T." box="[745,943,850,874]" page="5" pageId="10" pageNumber="11" year="2008">
<bibRefCitation id="8E460A9FF76CFFB5FD30FCFB7FCFFCAD" author="Karanovic, T." box="[745,943,850,874]" pageId="10" pageNumber="11" pagination="1 - 336" refId="ref47406" refString="Karanovic, T. (2008) Marine interstitial Poecilostomatoida and Cyclopoida (Copepoda) of Australia. Crustaceana Monographs, 9, 1 - 336. https: // doi. org / 10.1163 / ej. 9789004164598. i- 332" type="journal article" year="2008">Karanovic (2008: 5</bibRefCitation>
</treatmentCitation>
, 331)
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76CFFB5FF4EFC317F45FC74" blockId="10.[151,1437,920,1955]" box="[151,805,920,947]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF4EFC317DF0FC75" bold="true" box="[151,400,920,946]" pageId="10" pageNumber="11">Original description.</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FE4EFC317E2CFC74" author="Nicholls, A. G." box="[407,588,920,947]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls (1945)</bibRefCitation>
: 22–23; Text fig. 3.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76CFFB5FF1EFC157FC0FC10" box="[199,928,956,983]" pageId="10" pageNumber="11" type="description">
<paragraph id="EA68776EF76CFFB5FF1EFC157FC0FC10" blockId="10.[151,1437,920,1955]" box="[199,928,956,983]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF1EFC157DBBFC11" bold="true" box="[199,475,956,982]" pageId="10" pageNumber="11">Additional description.</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FE3BFC157EC0FC10" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[482,672,956,983]" pageId="10" pageNumber="11" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF76CFFB5FDCAFC147E2BFC11" box="[531,587,956,982]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2008)
</bibRefCitation>
: 435–441;
<figureCitation id="72EC6BEBF76CFFB5FCC5FC157FFCFC10" box="[796,924,956,983]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStart-4="FIGURE 17" captionStartId-0="49.[151,250,1943,1967]" captionStartId-1="50.[151,250,1943,1967]" captionStartId-2="51.[151,250,1942,1966]" captionStartId-3="55.[151,250,1943,1967]" captionStartId-4="56.[151,250,1943,1967]" captionTargetBox-0="[171,1412,190,1907]" captionTargetBox-1="[204,1383,186,1908]" captionTargetBox-2="[500,1087,189,1913]" captionTargetBox-3="[182,1406,193,1907]" captionTargetBox-4="[205,1381,188,1907]" captionTargetId-0="figure-17@49.[163,1424,181,1919]" captionTargetId-1="figure-17@50.[191,1397,181,1919]" captionTargetId-2="figure-17@51.[500,1087,181,1918]" captionTargetId-3="figure-17@55.[170,1418,181,1919]" captionTargetId-4="figure-17@56.[194,1393,181,1919]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionTargetPageId-2="51" captionTargetPageId-3="55" captionTargetPageId-4="56" captionText-0="FIGURE 13. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) antennule (♀), dorsal; (B) antenna (♀), medial [lateral endopodal spines indicated by arrows]; (C) antenna (♀), outer; (D) antennule (♂), anterior; (E) antennule (♂), dorsal." captionText-1="FIGURE 14. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." captionText-2="FIGURE 15. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: urosome (♀), ventral." captionText-3="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." captionText-4="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi-0="http://doi.org/10.5281/zenodo.14021506" figureDoi-1="http://doi.org/10.5281/zenodo.14021508" figureDoi-2="http://doi.org/10.5281/zenodo.14021510" figureDoi-3="http://doi.org/10.5281/zenodo.14021512" figureDoi-4="http://doi.org/10.5281/zenodo.14021514" httpUri-0="https://zenodo.org/record/14021506/files/figure.png" httpUri-1="https://zenodo.org/record/14021508/files/figure.png" httpUri-2="https://zenodo.org/record/14021510/files/figure.png" httpUri-3="https://zenodo.org/record/14021512/files/figure.png" httpUri-4="https://zenodo.org/record/14021514/files/figure.png" pageId="10" pageNumber="11">Figs 13–17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76CFFB5FF1EFC487FC0FB4C" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="EA68776EF76CFFB5FF1EFC487FC0FB4C" blockId="10.[151,1437,920,1955]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF1EFC487D08FC3D" bold="true" box="[199,360,992,1019]" pageId="10" pageNumber="11">Type locality.</emphasis>
<collectingCountry id="92C037FEF76CFFB5FEA8FC497DBBFC3D" box="[369,475,992,1018]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
,
<collectingRegion id="2813B98CF76CFFB5FE3EFC487ED4FC3D" box="[487,692,992,1019]" country="Australia" name="Western Australia" pageId="10" pageNumber="11">Western Australia</collectingRegion>
.
<bibRefCitation id="8E460A9FF76CFFB5FD18FC497F14FC3C" author="Nicholls, A. G." box="[705,884,992,1019]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls (1945)</bibRefCitation>
did not specify a type locality but collected the species from two sandy beaches some
<quantity id="2D2FDA8BF76CFFB5FD87FBAD7ED7FBD9" box="[606,695,1028,1055]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="km" value="500.0">500 km</quantity>
apart. The first, Leighton Beach (in the vicinity of Perth), was sampled in
<date id="9E6951AEF76CFFB5FEC5FB817DD7FB84" box="[284,439,1064,1091]" pageId="10" pageNumber="11" value="1939-10">October 1939</date>
while samples from the second site, locally known as ‘Back Beach’ at Dongarra, were collected in
<date id="9E6951AEF76CFFB5FEFBFBE57DCFFBA0" box="[290,431,1100,1127]" pageId="10" pageNumber="11" value="1940-03">March 1940</date>
. All
<typeStatus id="356CC9CCF76CFFB5FE3FFBE47E28FBA0" box="[486,584,1101,1127]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
deposited at the Natural History Museum in London originated from Back Beach which was designated as the type locality by
<bibRefCitation id="8E460A9FF76CFFB5FD07FBD97FFBFB4C" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[734,923,1136,1163]" pageId="10" pageNumber="11" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF76CFFB5FCD6FBD87F27FB4D" box="[783,839,1136,1162]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2008)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76CFFB5FF1EFB3C7F21FB68" box="[199,833,1172,1199]" pageId="10" pageNumber="11" type="description">
<paragraph id="EA68776EF76CFFB5FF1EFB3C7F21FB68" blockId="10.[151,1437,920,1955]" box="[199,833,1172,1199]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF1EFB3C7D3EFB68" bold="true" box="[199,350,1173,1199]" pageId="10" pageNumber="11">Body length.</emphasis>
280–300 μm (both sexes) [
<bibRefCitation id="8E460A9FF76CFFB5FD56FB3D7F56FB68" author="Nicholls, A. G." box="[655,822,1172,1199]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls 1945</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76CFFB5FF1EFB107875F864" pageId="10" pageNumber="11" type="discussion">
<paragraph id="EA68776EF76CFFB5FF1EFB107F34F998" blockId="10.[151,1437,920,1955]" pageId="10" pageNumber="11">
<emphasis id="D8A3AB7CF76CFFB5FF1EFB107D5BFB14" bold="true" box="[199,315,1209,1235]" pageId="10" pageNumber="11">Remarks.</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FE9FFB117E6BFB14" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[326,523,1208,1235]" pageId="10" pageNumber="11" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF76CFFB5FEA3FB107DD6FB15" box="[378,438,1208,1234]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2008)
</bibRefCitation>
re-examined the
<typeStatus id="356CC9CCF76CFFB5FD01FB107F68FB14" box="[728,776,1209,1235]" pageId="10" pageNumber="11">type</typeStatus>
material and redescribed both sexes in detail. Nicholls’s (1945) original description is adequate by contemporary standards except for the reported absence of the inner seta on P4 exp-3, an error that was perpetuated in some morphological comparisons (
<bibRefCitation id="8E460A9FF76CFFB5FBA6FAA9797AFADC" author="Noodt, W." box="[1151,1306,1280,1307]" pageId="10" pageNumber="11" pagination="86 - 109" refId="ref48577" refString="Noodt, W. (1955 b) Harpacticiden (Crust. Cop.) aus dem Sandstrand der franzosischen Biscaya-Kuste. Kieler Meeresforschungen, 11, 86 - 109." type="journal article" year="1955">Noodt 1955b</bibRefCitation>
; Bodiou &amp; Colomines 1996) and identification keys (
<bibRefCitation id="8E460A9FF76CFFB5FDA6FA8C7F60FAF8" author="Lang, K." box="[639,768,1316,1343]" pageId="10" pageNumber="11" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang 1965</bibRefCitation>
;
<bibRefCitation id="8E460A9FF76CFFB5FCD6FA8D7FAAFAF8" author="Karanovic, T." box="[783,970,1316,1343]" pageId="10" pageNumber="11" pagination="33 - 38" refId="ref47348" refString="Karanovic, T. (2000) Arenopontia (Neoleptastacus) huysi, sp. nov. (Crustacea, Copepoda, Harpacticoida) from marine interstitial of Montenegro (S. E. Europe). Helgoland marine Research, 54, 33 - 38. https: // doi. org / 10.1007 / s 101520050033" type="journal article" year="2000">Karanovic 2000</bibRefCitation>
;
<bibRefCitation id="8E460A9FF76CFFB5FC01FA8D783EFAF8" author="Wells, J. B. J." box="[984,1118,1316,1343]" pageId="10" pageNumber="11" pagination="1 - 872" refId="ref50074" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1 - 872. https: // doi. org / 10.11646 / zootaxa. 1568.1.1" type="journal article" year="2007">Wells 2007</bibRefCitation>
). Within the
<taxonomicName id="2DD70CEDF76CFFB5FB2EFA8D79F3FAF9" authorityName="Nicholls" authorityYear="1945" box="[1271,1427,1316,1342]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FB2EFA8D79F3FAF9" box="[1271,1427,1316,1342]" italics="true" pageId="10" pageNumber="11">spinicaudatus</emphasis>
</taxonomicName>
- group,
<taxonomicName id="2DD70CEDF76CFFB5FF3FFAE07DC2FAA5" authorityName="Nicholls" authorityYear="1945" box="[230,418,1352,1378]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FF3FFAE07DC2FAA5" box="[230,418,1352,1378]" italics="true" pageId="10" pageNumber="11">N. spinicaudatus</emphasis>
</taxonomicName>
is morphologically closest to
<taxonomicName id="2DD70CEDF76CFFB5FD29FAE07F0CFAA5" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[752,876,1353,1378]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spicatus">
<emphasis id="D8A3AB7CF76CFFB5FD29FAE07F0CFAA5" box="[752,876,1353,1378]" italics="true" pageId="10" pageNumber="11">N. spicatus</emphasis>
</taxonomicName>
, both of which exhibit a dorsolateral spur near the base of caudal ramus seta VII. The
<typeStatus id="356CC9CCF76CFFB5FDC5FAC47E2CFA40" box="[540,588,1389,1415]" pageId="10" pageNumber="11">type</typeStatus>
species can be differentiated from
<taxonomicName id="2DD70CEDF76CFFB5FC16FAC4782BFA41" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[975,1099,1389,1414]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spicatus">
<emphasis id="D8A3AB7CF76CFFB5FC16FAC4782BFA41" box="[975,1099,1389,1414]" italics="true" pageId="10" pageNumber="11">N. spicatus</emphasis>
</taxonomicName>
by the shape of the abdominal hyaline frills (lappets denticulate
<emphasis id="D8A3AB7CF76CFFB5FDC1FA387E4FFA6D" box="[536,559,1425,1450]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
semi-incised obtusidigitate), the proportional size of the inner distal seta on P2–P3 enp-2 (shorter
<emphasis id="D8A3AB7CF76CFFB5FE4BFA1C7DC9FA09" box="[402,425,1461,1486]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
longer than endopod) and the morphometric dimensions of the P5 (2.5 times as long as wide
<emphasis id="D8A3AB7CF76CFFB5FF0FFA707C8DFA35" box="[214,237,1497,1522]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
3.0 times). The apical spinous process on the P5 of both sexes is minutely bipinnate in
<taxonomicName id="2DD70CEDF76CFFB5FB07FA7079FDFA35" authorityName="Nicholls" authorityYear="1945" box="[1246,1437,1496,1522]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FB07FA7079FDFA35" box="[1246,1437,1496,1522]" italics="true" pageId="10" pageNumber="11">N. spinicaudatus</emphasis>
</taxonomicName>
(and still delimited at its base on the posterior surface in the
<specimenCount id="FCD1BCE7F76CFFB5FCEAFA547F26F9D0" box="[819,838,1533,1559]" count="1" pageId="10" pageNumber="11" type="female">♀</specimenCount>
) while it is lacking such ornamentation in
<taxonomicName id="2DD70CEDF76CFFB5FAF8FA5479FCF9D1" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1985" box="[1313,1436,1533,1558]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spicatus">
<emphasis id="D8A3AB7CF76CFFB5FAF8FA5479FCF9D1" box="[1313,1436,1533,1558]" italics="true" pageId="10" pageNumber="11">N. spicatus</emphasis>
</taxonomicName>
(and completely fused to the segment in the
<specimenCount id="FCD1BCE7F76CFFB5FD52F9887EFFF9FC" box="[651,671,1569,1595]" count="1" pageId="10" pageNumber="11" type="female">♀</specimenCount>
). The linear egg-sac typically contains 2–4 large eggs; occasionally
<specimenCount id="FCD1BCE7F76CFFB5FF4EF9ED7C92F998" box="[151,242,1604,1631]" count="6" pageId="10" pageNumber="11" type="egg">six eggs</specimenCount>
are found in which case they overlap (
<bibRefCitation id="8E460A9FF76CFFB5FD7DF9ED7F28F998" author="Nicholls, A. G." box="[676,840,1604,1631]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls 1945</bibRefCitation>
).
</paragraph>
<paragraph id="EA68776EF76CFFB5FF1EF9C17875F864" blockId="10.[151,1437,920,1955]" pageId="10" pageNumber="11">
The species is so far known only from two sandy beaches in Western Australia (
<bibRefCitation id="8E460A9FF76CFFB5FBE9F9C178B2F944" author="Nicholls, A. G." box="[1072,1234,1640,1667]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls 1945</bibRefCitation>
).
<bibRefCitation id="8E460A9FF76CFFB5FB3DF9C17CABF961" author="Chappuis, P. - A." pageId="10" pageNumber="11" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis (1954: 269)</bibRefCitation>
claimed to have found
<specimenCount id="FCD1BCE7F76CFFB5FE05F9247E3AF961" box="[476,602,1676,1703]" count="1" pageId="10" pageNumber="11" type="female">one female</specimenCount>
and
<specimenCount id="FCD1BCE7F76CFFB5FD4DF9247E9CF961" box="[660,764,1676,1703]" count="1" pageId="10" pageNumber="11" type="male">one male</specimenCount>
in
<collectingRegion id="2813B98CF76CFFB5FCFBF9257F19F961" box="[802,889,1676,1702]" country="Algeria" name="Annaba" pageId="10" pageNumber="11">Annaba</collectingRegion>
(= Bône) in northeastern
<collectingCountry id="92C037FEF76CFFB5FB7BF9257896F961" box="[1186,1270,1676,1702]" name="Algeria" pageId="10" pageNumber="11">Algeria</collectingCountry>
which closely resemble
<taxonomicName id="2DD70CEDF76CFFB5FEDAF9187DDFF90D" authorityName="Nicholls" authorityYear="1945" box="[259,447,1712,1738]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FEDAF9187DDFF90D" box="[259,447,1712,1738]" italics="true" pageId="10" pageNumber="11">N. spinicaudatus</emphasis>
</taxonomicName>
but did not present any illustrations to substantiate his record. In a later report,
<bibRefCitation id="8E460A9FF76CFFB5FAEBF9197D73F928" author="Chappuis P. - A." pageId="10" pageNumber="11" pagination="409 - 422" refId="ref45686" refString="Chappuis P. - A. (1958) Harpacticoides psammiques marins des environs de Seattle (Washington, U. S. A.). Vie et Milieu, 8, 409 - 422." type="journal article" year="1958">Chappuis (1958: 414</bibRefCitation>
,
<figureCitation id="72EC6BEBF76CFFB5FEC4F97D7D1EF928" box="[285,382,1748,1775]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="13.[151,250,1943,1967]" captionStartId-1="14.[151,250,1844,1868]" captionStartId-2="15.[151,250,1943,1967]" captionStartId-3="16.[151,250,1943,1967]" captionTargetBox-0="[187,1401,193,1915]" captionTargetBox-1="[160,1424,322,1809]" captionTargetBox-2="[185,1411,193,1907]" captionTargetBox-3="[203,1384,193,1907]" captionTargetId-0="figure-17@13.[175,1413,181,1919]" captionTargetId-1="figure-17@14.[151,1436,307,1821]" captionTargetId-2="figure-17@15.[173,1414,181,1919]" captionTargetId-3="figure-17@16.[191,1396,181,1919]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" captionText-1="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" captionText-2="FIGURE 4. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." captionText-3="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi-0="http://doi.org/10.5281/zenodo.14021484" figureDoi-1="http://doi.org/10.5281/zenodo.14021486" figureDoi-2="http://doi.org/10.5281/zenodo.14021488" figureDoi-3="http://doi.org/10.5281/zenodo.14021490" httpUri-0="https://zenodo.org/record/14021484/files/figure.png" httpUri-1="https://zenodo.org/record/14021486/files/figure.png" httpUri-2="https://zenodo.org/record/14021488/files/figure.png" httpUri-3="https://zenodo.org/record/14021490/files/figure.png" pageId="10" pageNumber="11">Figs 2–5</figureCitation>
) provided illustrations of the female antennule, P2 endopod, P5 and caudal ramus of specimens he collected in Puget Sound, Washington State. Despite differences in the P5, length of P3–P4 endopods and caudal rami,
<bibRefCitation id="8E460A9FF76CFFB5FF03F8B57DF6F8F0" author="Chappuis P. - A." box="[218,406,1820,1847]" pageId="10" pageNumber="11" pagination="409 - 422" refId="ref45686" refString="Chappuis P. - A. (1958) Harpacticoides psammiques marins des environs de Seattle (Washington, U. S. A.). Vie et Milieu, 8, 409 - 422." type="journal article" year="1958">Chappuis (1958)</bibRefCitation>
assigned his material to
<taxonomicName id="2DD70CEDF76CFFB5FD6BF8B47F0FF8F1" authorityName="Nicholls" authorityYear="1945" box="[690,879,1820,1846]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF76CFFB5FD6BF8B47F0FF8F1" box="[690,879,1820,1846]" italics="true" pageId="10" pageNumber="11">N. spinicaudatus</emphasis>
</taxonomicName>
, stating that such morphological minutiae do not warrant the proposal of a distinct subspecies. As pointed out by
<bibRefCitation id="8E460A9FF76CFFB5FCB0F8E97848F89C" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[873,1064,1856,1883]" pageId="10" pageNumber="11" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF76CFFB5FC42F8E87FB4F89D" box="[923,980,1856,1882]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2008)
</bibRefCitation>
, the morphology of the fifth legs suggests that
<bibRefCitation id="8E460A9FF76CFFB5FEF2F8CD7D85F8B8" author="Chappuis P. - A." box="[299,485,1892,1919]" pageId="10" pageNumber="11" pagination="409 - 422" refId="ref45686" refString="Chappuis P. - A. (1958) Harpacticoides psammiques marins des environs de Seattle (Washington, U. S. A.). Vie et Milieu, 8, 409 - 422." type="journal article" year="1958">Chappuis (1958)</bibRefCitation>
was almost certainly dealing with
<taxonomicName id="2DD70CEDF76CFFB5FCBBF8CC797EF8B8" authority="(Lang, 1965)" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[866,1310,1892,1919]" class="Copepoda" family="Arenopontiidae" genus="Mesopontia" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dillonbeachia">
<emphasis id="D8A3AB7CF76CFFB5FCBBF8CC78E5F8B9" box="[866,1157,1892,1918]" italics="true" pageId="10" pageNumber="11">Mesopontia dillonbeachia</emphasis>
(
<bibRefCitation id="8E460A9FF76CFFB5FB4AF8CC7976F8B8" author="Lang, K." box="[1171,1302,1892,1919]" pageId="10" pageNumber="11" pagination="1 - 560" refId="ref47749" refString="Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska vetenskapsakademiens Handlingar, Series 4, 10 (2), 1 - 560." type="journal article" year="1965">Lang, 1965</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="8E460A9FF76CFFB5FAF1F8CD7C82F864" author="Karanovic, T." pageId="10" pageNumber="11" pagination="1 - 336" refId="ref47406" refString="Karanovic, T. (2008) Marine interstitial Poecilostomatoida and Cyclopoida (Copepoda) of Australia. Crustaceana Monographs, 9, 1 - 336. https: // doi. org / 10.1163 / ej. 9789004164598. i- 332" type="journal article" year="2008">Karanovic (2008)</bibRefCitation>
erroneously cited the species as
<taxonomicName id="2DD70CEDF76CFFB5FD88F8217871F864" authority="Nicholls, 1945" authorityName="Nicholls" authorityYear="1945" box="[593,1041,1928,1955]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="D8A3AB7CF76CFFB5FD88F8217F02F865" box="[593,866,1928,1954]" italics="true" pageId="10" pageNumber="11">Neoleptastacus longipes</emphasis>
<bibRefCitation id="8E460A9FF76CFFB5FCB0F8217871F864" author="Nicholls, A. G." box="[873,1041,1928,1955]" pageId="10" pageNumber="11" pagination="17 - 24" refId="ref48495" refString="Nicholls, A. G. (1945) Marine Copepoda from Western Australia. IV. - Psammophilous harpacticoids. Journal of the Royal Society of Western Australia, 29, 17 - 24." type="journal article" year="1945">Nicholls, 1945</bibRefCitation>
</taxonomicName>
.
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757
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<emphasis id="D8A3AB7CF76DFFB4FF4EFC487DA4FC3B" bold="true" box="[151,452,993,1020]" italics="true" pageId="11" pageNumber="12">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF76DFFB4FE0CFC487EF8FC3C" author="Chappuis, P. - A." box="[469,664,993,1019]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
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(
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)
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<taxonomicName id="2DD70CEDF76DFFB4FF4EFBE27E73FBA5" ID-CoL="GFPY" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[151,531,1098,1122]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
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<bibRefCitation id="8E460A9FF76DFFB4FEAAFBE37E73FBA5" author="Chappuis, P. - A." box="[371,531,1098,1122]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
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<paragraph id="EA68776EF76DFFB4FF4EFBC07FE7FB46" blockId="11.[151,990,1098,1276]" box="[151,903,1129,1153]" pageId="11" pageNumber="12">
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<taxonomicName id="2DD70CEDF76DFFB4FF4EFBC07EA8FB46" ID-CoL="GFPY" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[151,712,1129,1153]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF76DFFB4FF4EFBC07D75FB47" box="[151,277,1129,1152]" italics="true" pageId="11" pageNumber="12">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF76DFFB4FEFDFBC07DDFFB46" box="[292,447,1129,1153]" italics="true" pageId="11" pageNumber="12">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF76DFFB4FE14FBC07E7FFB46" box="[461,543,1129,1153]" italics="true" pageId="11" pageNumber="12">acantha</emphasis>
<bibRefCitation id="8E460A9FF76DFFB4FDFCFBC07EA8FB46" author="Chappuis, P. - A." box="[549,712,1129,1153]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF76DFFB4FD08FBC07FE7FB46" author="Wells, J. B. J." box="[721,903,1129,1153]" page="324" pageId="11" pageNumber="12" year="1967">
<bibRefCitation id="8E460A9FF76DFFB4FD08FBC07FE7FB46" author="Wells, J. B. J." box="[721,903,1129,1153]" pageId="11" pageNumber="12" pagination="189 - 358" refId="ref49859" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / S 0080456800024017" type="journal article" year="1967">Wells (1967: 324)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76DFFB4FF4EFB217FCEFB67" blockId="11.[151,990,1098,1276]" box="[151,942,1159,1184]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="CAC75040F76DFFB4FF4EFB217FCEFB67" box="[151,942,1159,1184]" pageId="11" pageNumber="12">
<taxonomicName id="2DD70CEDF76DFFB4FF4EFB217E42FB58" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[151,546,1159,1184]" class="Copepoda" family="Arenopontiidae" genus="Arenoponthia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF76DFFB4FF4EFB217D1AFB67" box="[151,378,1160,1184]" italics="true" pageId="11" pageNumber="12">Arenoponthia acantha</emphasis>
<bibRefCitation id="8E460A9FF76DFFB4FE59FB217E42FB58" author="Chappuis, P. - A." box="[384,546,1159,1184]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
</taxonomicName>
:
<emphasis id="D8A3AB7CF76DFFB4FDF5FB217ED8FB67" box="[556,696,1160,1184]" italics="true" pageId="11" pageNumber="12">lapsus calami</emphasis>
by
<treatmentCitation id="6B76517FF76DFFB4FD07FB217FCEFB67" author="Cottarelli, V." box="[734,942,1159,1184]" page="20" pageId="11" pageNumber="12" year="1969">
<bibRefCitation id="8E460A9FF76DFFB4FD07FB217FCEFB67" author="Cottarelli, V." box="[734,942,1159,1184]" pageId="11" pageNumber="12" pagination="8 - 21" refId="ref45879" refString="Cottarelli, V. (1969) Un nuovo Crostaceo di acque interstiziali Italiane: Psammopsyllus pasquinii n. sp. - (Harpacticoida, Cylindropsyllidae). Rendiconti dell'Istituto Lombardo di Scienze e Lettere, (B) 103, 8 - 21." type="journal article" year="1969">Cottarelli (1969: 20)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76DFFB4FF4EFB0E7FBEFB79" blockId="11.[151,990,1098,1276]" box="[151,990,1190,1214]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="CAC75040F76DFFB4FF4EFB0E7FBEFB79" box="[151,990,1190,1214]" pageId="11" pageNumber="12">
<taxonomicName id="2DD70CEDF76DFFB4FF4EFB0E7F40FB79" ID-CoL="GFPY" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[151,800,1190,1214]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="acantha" subGenus="Neoleptastacus" subSpecies="acantha">
<emphasis id="D8A3AB7CF76DFFB4FF4EFB0E7D75FB79" box="[151,277,1191,1214]" italics="true" pageId="11" pageNumber="12">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF76DFFB4FEFDFB0F7DDFFB79" box="[292,447,1190,1214]" italics="true" pageId="11" pageNumber="12">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF76DFFB4FE14FB0F7E17FB79" box="[461,631,1190,1214]" italics="true" pageId="11" pageNumber="12">acantha acantha</emphasis>
<bibRefCitation id="8E460A9FF76DFFB4FDA4FB0F7F40FB79" author="Chappuis, P. - A." box="[637,800,1190,1214]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF76DFFB4FCF3FB0E7FBEFB79" author="Kunz, H." box="[810,990,1190,1214]" page="356" pageId="11" pageNumber="12" year="1971">
<bibRefCitation id="8E460A9FF76DFFB4FCF3FB0E7FBEFB79" author="Kunz, H." box="[810,990,1190,1214]" pageId="11" pageNumber="12" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971: 356)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76DFFB4FF4EFB6C7F4DFB1A" blockId="11.[151,990,1098,1276]" box="[151,813,1221,1245]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="CAC75040F76DFFB4FF4EFB6C7F4DFB1A" box="[151,813,1221,1245]" pageId="11" pageNumber="12">
<treatmentCitation id="6B76517FF76DFFB4FF4EFB6C7F4DFB1A" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[151,813,1221,1245]" page=")" pageId="11" pageNumber="12" year="1954">
<taxonomicName id="2DD70CEDF76DFFB4FF4EFB6C7F4DFB1A" ID-CoL="73VZK" authority="(Chappuis, 1954) Sak et al. (2008: 412)" authorityName="Sak et al." authorityPageNumber="412" authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[151,813,1221,1245]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF76DFFB4FF4EFB6C7DF3FB1A" box="[151,403,1221,1245]" italics="true" pageId="11" pageNumber="12">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF76DFFB4FE78FB6C7E23FB1A" author="Chappuis, P. - A." box="[417,579,1221,1245]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
<bibRefCitation id="8E460A9FF76DFFB4FD89FB6C7F4DFB1A" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[592,813,1221,1245]" pageId="11" pageNumber="12" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF76DFFB4FDA4FB6F7ED0FB1A" box="[637,688,1221,1245]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2008: 412)
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76DFFB4FF4EFB4D7F7FFB3B" blockId="11.[151,990,1098,1276]" box="[151,799,1252,1276]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="CAC75040F76DFFB4FF4EFB4D7F7FFB3B" box="[151,799,1252,1276]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF4EFB4D7E35FB3B" box="[151,597,1252,1276]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="2DD70CEDF76DFFB4FF4EFB4D7DCDFB3B" authority="Sak, 2004" box="[151,429,1252,1276]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="knidosensis">Neoleptastacus knidosensis</taxonomicName>
—nomen nudum
</emphasis>
by
<treatmentCitation id="6B76517FF76DFFB4FDA2FB4D7F7FFB3B" author="Sak, S." box="[635,799,1252,1276]" page="230" pageId="11" pageNumber="12" year="2004">
<bibRefCitation id="8E460A9FF76DFFB4FDA2FB4D7F7FFB3B" author="Sak, S." box="[635,799,1252,1276]" pageId="11" pageNumber="12" refId="ref49361" refString="Sak, S. (2004) Arenopontiidae (Copepoda: Harpacticoida) familyasinin taksonomisi ve filogenisi. [Taxonomy and phylogeny of the family Arenopontiidae (Copepoda: Harpacticoida)]. PhD dissertation, University of Balikesir, Balikesir, 312 pp." type="book" year="2004">Sak (2004: 230)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA68776EF76DFFB4FF4EFA807F37FA82" blockId="11.[151,1437,1321,2032]" box="[151,855,1321,1349]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF4EFA807DF0FA84" bold="true" box="[151,400,1321,1347]" pageId="11" pageNumber="12">Original description.</emphasis>
<bibRefCitation id="8E460A9FF76DFFB4FE4EFA837E36FA82" author="Chappuis, P. - A." box="[407,598,1322,1349]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
: 268–270; Figs 46–53.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76DFFB4FF1EFAE47874FA77" pageId="11" pageNumber="12" type="description">
<paragraph id="EA68776EF76DFFB4FF1EFAE47874FA77" blockId="11.[151,1437,1321,2032]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF1EFAE47D86FAA0" bold="true" box="[199,486,1357,1383]" pageId="11" pageNumber="12">Additional descriptions.</emphasis>
<bibRefCitation id="8E460A9FF76DFFB4FE34FAE77E1BFAAF" author="Masry, D." box="[493,635,1358,1384]" pageId="11" pageNumber="12" pagination="229 - 258" refId="ref48094" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970</bibRefCitation>
—as
<taxonomicName id="2DD70CEDF76DFFB4FD61FAE77F51FAAF" authorityName="Chappuis" authorityYear="1954" box="[696,817,1358,1384]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF76DFFB4FD61FAE77F51FAAF" box="[696,817,1358,1384]" italics="true" pageId="11" pageNumber="12">A. acantha</emphasis>
</taxonomicName>
): 251–253;
<figureCitation id="72EC6BEBF76DFFB4FC63FAE7786DFAAF" box="[954,1037,1358,1384]" captionStart="FIGURE 13" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[171,1412,190,1907]" captionTargetId="figure-17@49.[163,1424,181,1919]" captionTargetPageId="49" captionText="FIGURE 13. Phreatipontia phreatica (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) antennule (♀), dorsal; (B) antenna (♀), medial [lateral endopodal spines indicated by arrows]; (C) antenna (♀), outer; (D) antennule (♂), anterior; (E) antennule (♂), dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021506" httpUri="https://zenodo.org/record/14021506/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
.
<bibRefCitation id="8E460A9FF76DFFB4FBC1FAE778B5FAAF" author="Cottarelli V." box="[1048,1237,1358,1384]" pageId="11" pageNumber="12" pagination="49 - 59" refId="ref45971" refString="Cottarelli V. (1973 a) Arenopontia gussoae n. sp., nuovo arpacticoide di acque interstiziali litorali dell'isola di Cuba (Crustacea, Copepoda). Fragmenta entomologica, 9, 49 - 59." type="journal article" year="1973">Cottarelli (1973a</bibRefCitation>
—as
<taxonomicName id="2DD70CEDF76DFFB4FACBFAE779EBFAAF" authorityName="Chappuis" authorityYear="1954" box="[1298,1419,1358,1384]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF76DFFB4FACBFAE779EBFAAF" box="[1298,1419,1358,1384]" italics="true" pageId="11" pageNumber="12">A. acantha</emphasis>
</taxonomicName>
): Fig. 22 [mandibular palp].
<bibRefCitation id="8E460A9FF76DFFB4FE10FADB7F5AFA4B" author="Martinez Arbizu, P. &amp; Moura, G." box="[457,826,1394,1420]" pageId="11" pageNumber="12" pagination="55 - 77" refId="ref48048" refString="Martinez Arbizu, P. &amp; Moura, G. (1994) The phylogenetic position of the Cylindropsyllinae Sars (Copepoda, Harpacticoida) and the systematic status of the Leptopontiinae Lang. Zoologische Beitrage, Neue Folge, 35, 55 - 77." type="journal article" year="1994">Martínez Arbizu &amp; Moura (1994</bibRefCitation>
—as
<taxonomicName id="2DD70CEDF76DFFB4FCAEFADB7F91FA4B" authorityName="Chappuis" authorityYear="1954" box="[887,1009,1394,1420]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF76DFFB4FCAEFADB7F91FA4B" box="[887,1009,1394,1420]" italics="true" pageId="11" pageNumber="12">A. acantha</emphasis>
</taxonomicName>
): 63;
<figureCitation id="72EC6BEBF76DFFB4FBEFFADB78C4FA4B" box="[1078,1188,1394,1420]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="11" pageNumber="12">Fig. 2d, g</figureCitation>
[rostrum, anal somite and caudal rami].
<bibRefCitation id="8E460A9FF76DFFB4FEB9FA3F7D95FA77" author="Alper, A." box="[352,501,1430,1456]" pageId="11" pageNumber="12" refId="ref44916" refString="Alper, A. (2009) Datca-Bozburun ozel cevre koruma bolgesi (Mugla, Turkiye) sahillerinin kumici ve fital Harpacticoida (Crustacea, Copepoda) faunasinin belirlenmesi. Determination of the interstitial and phytal Harpacticoida (Crustacea, Copepoda) fauna of the coast of Datca-Bozburun (Mugla, Turkey) specially protected area. PhD dissertation, University of Balikesir, Turkiye, 150 pp. [in Turkish with English summary]" type="book" year="2009">Alper (2009)</bibRefCitation>
: 45–46;
<figureCitation id="72EC6BEBF76DFFB4FD8CFA3F7ECBFA77" box="[597,683,1430,1456]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="11" pageNumber="12">Fig. 3.6</figureCitation>
[
<specimenCount id="FCD1BCE7F76DFFB4FD65FA3F7EB0FA77" box="[700,720,1430,1456]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
lateral habitus, photograph].
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76DFFB4FF1EFA1378EBF933" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="EA68776EF76DFFB4FF1EFA137E7FF9A3" blockId="11.[151,1437,1321,2032]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF1EFA137D0AFA14" bold="true" box="[199,362,1465,1492]" pageId="11" pageNumber="12">Type locality.</emphasis>
<location id="EF0821B5F76DFFB4FEADFA137EE5FA13" LSID="urn:lsid:plazi:treatment:627EC678F76DFFACFF4EFC487E68FBBA:EF0821B5F76DFFB4FEADFA137EE5FA13" box="[372,645,1466,1492]" country="Italy, Algeria" name="Western Mediterranean." pageId="11" pageNumber="12">Western Mediterranean.</location>
<bibRefCitation id="8E460A9FF76DFFB4FD49FA137F2FFA12" author="Chappuis, P. - A." box="[656,847,1466,1493]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
collected the species in various localities in
<collectingCountry id="92C037FEF76DFFB4FABDFA137C8BFA3F" isEnumeration="true" pageId="11" pageNumber="12">Italy, Algeria</collectingCountry>
and
<collectingCountry id="92C037FEF76DFFB4FEFAFA777D16FA3F" box="[291,374,1502,1528]" name="Tunisia" pageId="11" pageNumber="12">Tunisia</collectingCountry>
(see below) but did not specify a
<emphasis id="D8A3AB7CF76DFFB4FD25FA777FEEFA3F" box="[764,910,1502,1528]" italics="true" pageId="11" pageNumber="12">locus typicus</emphasis>
.
<collectorName id="472212B8F76DFFB4FC44FA77787CFA3F" box="[925,1052,1502,1528]" pageId="11" pageNumber="12">Since P.-A.</collectorName>
Chappuis habitually ignored type fixation, all the specimens on which he established
<taxonomicName id="2DD70CEDF76DFFB4FD3AF9AB7F3DF9DB" authorityName="Chappuis" authorityYear="1954" box="[739,861,1538,1564]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF76DFFB4FD3AF9AB7F3DF9DB" box="[739,861,1538,1564]" italics="true" pageId="11" pageNumber="12">A. acantha</emphasis>
</taxonomicName>
are
<typeStatus id="356CC9CCF76DFFB4FC49F9AB7F92F9DB" box="[912,1010,1538,1564]" pageId="11" pageNumber="12" type="syntype">syntypes</typeStatus>
and collectively constitute the name-bearing type (
<collectionCode id="8CC6EFABF76DFFB4FEEFF98F7D19F987" box="[310,377,1574,1600]" pageId="11" pageNumber="12">ICZN</collectionCode>
Art. 72.1.1). According to
<collectionCode id="8CC6EFABF76DFFB4FD6CF98F7E96F987" box="[693,758,1574,1600]" pageId="11" pageNumber="12">ICZN</collectionCode>
Art. 73.2.3 the type locality encompasses all of the places where the
<typeStatus id="356CC9CCF76DFFB4FED5F9E37D0EF9A3" box="[268,366,1610,1636]" pageId="11" pageNumber="12" type="syntype">syntypes</typeStatus>
were collected.
</paragraph>
<paragraph id="EA68776EF76DFFB4FF1EF9C478EBF933" blockId="11.[151,1437,1321,2032]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF1EF9C47DD1F940" bold="true" box="[199,433,1645,1671]" pageId="11" pageNumber="12">Material examined.</emphasis>
<materialsCitation id="5ABF7D33F76DFFB4FE61F9C778E5F94E" box="[440,1157,1646,1673]" collectionCode="NHMUK" pageId="11" pageNumber="12" specimenCode="NHMUK 2024.1050" specimenCount="1" specimenCount-female="1">
<specimenCount id="FCD1BCE7F76DFFB4FE61F9C77E63F94F" box="[440,515,1646,1672]" count="1" pageId="11" pageNumber="12" type="female">One ♀</specimenCount>
(dissected on eight slides) (reg. no
<specimenCode id="BA71DF15F76DFFB4FC4DF9C7781EF94E" box="[916,1150,1646,1673]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">NHMUK 2024.1050</specimenCode>
)
</materialsCitation>
and
<materialsCitation id="5ABF7D33F76DFFB4FB64F9C77EE7F96A" collectionCode="NHMUK" pageId="11" pageNumber="12" specimenCode="NHMUK 2024.1051" specimenCount="1" specimenCount-male="1">
<specimenCount id="FCD1BCE7F76DFFB4FB64F9C77961F94F" box="[1213,1281,1646,1672]" count="1" pageId="11" pageNumber="12" type="male">one ♂</specimenCount>
(dissected on seven slides) (reg. no
<specimenCode id="BA71DF15F76DFFB4FE49F93B7E1EF96A" box="[400,638,1682,1709]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">NHMUK 2024.1051</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="5ABF7D33F76DFFB4FD4AF93B7CBCF917" collectionCode="NHMUK" pageId="11" pageNumber="12" specimenCode="NHMUK 2024.1052, 1061" specimenCount="10" specimenCount-female="7" specimenCount-male="3">
<specimenCount id="FCD1BCE7F76DFFB4FD4AF93B7F64F96B" box="[659,772,1682,1708]" count="7" pageId="11" pageNumber="12" type="female">seven ♀♀</specimenCount>
and
<specimenCount id="FCD1BCE7F76DFFB4FCE4F93B7FC7F96B" box="[829,935,1682,1708]" count="3" pageId="11" pageNumber="12" type="male">three ♂♂</specimenCount>
(in ethanol) (reg. nos
<specimenCode id="BA71DF15F76DFFB4FB7CF93B79EEF96A" box="[1189,1422,1682,1709]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">NHMUK 2024.1052</specimenCode>
–
<specimenCode id="BA71DF15F76DFFB4FF4EF91F7CB3F917" box="[151,211,1718,1744]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">1061</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="5ABF7D33F76DFFB4FF32F91F78EBF933" collectingDate="2002-11-24" collectionCode="BUZM" collectorName="A. Alper." country="Turkey" county="Datca" location="All" municipality="Hastanealti Beach" pageId="11" pageNumber="12" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Mugla">
<specimenCount id="FCD1BCE7F76DFFB4FF32F91F7D2BF917" box="[235,331,1718,1744]" count="5" pageId="11" pageNumber="12" type="female">five ♀♀</specimenCount>
and
<specimenCount id="FCD1BCE7F76DFFB4FE51F91F7D86F917" box="[392,486,1718,1744]" count="2" pageId="11" pageNumber="12" type="male">two ♂♂</specimenCount>
(in ethanol) deposited in the collections of
<collectionCode id="8CC6EFABF76DFFB4FC31F91F785DF917" box="[1000,1085,1718,1744]" pageId="11" pageNumber="12">BUZM</collectionCode>
.
<location id="EF0821B5F76DFFB4FB90F91F780DF917" LSID="urn:lsid:plazi:treatment:627EC678F76DFFACFF4EFC487E68FBBA:EF0821B5F76DFFB4FB90F91F780DF917" box="[1097,1133,1718,1744]" country="Turkey" county="Datca" municipality="Hastanealti Beach" name="All" pageId="11" pageNumber="12" stateProvince="Mugla">All</location>
specimens collected from
<collectingMunicipality id="0A0CED14F76DFFB4FF4EF9737D05F933" box="[151,357,1754,1780]" pageId="11" pageNumber="12">Hastanealtı Beach</collectingMunicipality>
,
<collectingCounty id="03090FE2F76DFFB4FEA9F9737DD1F933" box="[368,433,1754,1780]" pageId="11" pageNumber="12">Datça</collectingCounty>
(
<collectingRegion id="2813B98CF76DFFB4FE19F9737E1AF933" box="[448,634,1754,1780]" country="Turkey" name="Mugla" pageId="11" pageNumber="12">Muğla Province</collectingRegion>
),
<collectingCountry id="92C037FEF76DFFB4FD55F9737E85F933" box="[652,741,1754,1780]" name="Turkey" pageId="11" pageNumber="12">Türkiye</collectingCountry>
on
<date id="9E6951AEF76DFFB4FCD6F9737F8BF933" box="[783,1003,1754,1780]" pageId="11" pageNumber="12" value="2002-11-24">
<collectingDate id="8E2DA846F76DFFB4FCD6F9737F8BF933" box="[783,1003,1754,1780]" pageId="11" pageNumber="12" value="2002-11-24">24 November 2002</collectingDate>
</date>
; leg.
<collectorName id="472212B8F76DFFB4FBFDF97378EBF933" box="[1060,1163,1754,1780]" pageId="11" pageNumber="12">A. Alper.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F76DFFADFF1EF9577F0CFA06" lastPageId="18" lastPageNumber="19" pageId="11" pageNumber="12" type="description">
<paragraph id="EA68776EF76DFFB4FF1EF9577DDBF8FB" blockId="11.[151,1437,1321,2032]" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF1EF9577D3DF8DF" bold="true" box="[199,349,1790,1816]" pageId="11" pageNumber="12">Body length.</emphasis>
450 μm (
<specimenCount id="FCD1BCE7F76DFFB4FE10F9577DBBF8DF" box="[457,475,1790,1816]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
) [
<bibRefCitation id="8E460A9FF76DFFB4FE2BF9577EC3F8DE" author="Chappuis, P. - A." box="[498,675,1790,1817]" pageId="11" pageNumber="12" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis 1954</bibRefCitation>
]; 380–420 μm (
<specimenCount id="FCD1BCE7F76DFFB4FC8BF9577F05F8DF" box="[850,869,1790,1816]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), 365–390 μm (
<specimenCount id="FCD1BCE7F76DFFB4FBC1F957784EF8DF" box="[1048,1070,1790,1816]" count="1" pageId="11" pageNumber="12" type="male">♂</specimenCount>
) [
<bibRefCitation id="8E460A9FF76DFFB4FB9AF95778B1F8DF" author="Masry, D." box="[1091,1233,1790,1816]" pageId="11" pageNumber="12" pagination="229 - 258" refId="ref48094" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry 1970</bibRefCitation>
]; 331 μm (
<specimenCount id="FCD1BCE7F76DFFB4FA90F957793CF8DF" box="[1353,1372,1790,1816]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), 325 μm (
<specimenCount id="FCD1BCE7F76DFFB4FF15F88B7C82F8FB" box="[204,226,1826,1852]" count="1" pageId="11" pageNumber="12" type="male">♂</specimenCount>
) [present account].
</paragraph>
<paragraph id="EA68776EF76DFFAEFF1EF8EC7D9CFEA2" blockId="11.[151,1437,1321,2032]" lastBlockId="17.[151,1437,150,2013]" lastPageId="17" lastPageNumber="18" pageId="11" pageNumber="12">
<emphasis id="D8A3AB7CF76DFFB4FF1EF8EC7D8EF898" bold="true" box="[199,494,1861,1887]" pageId="11" pageNumber="12">Redescription of female.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 350 μm (mean = 331 μm;
<emphasis id="D8A3AB7CF76DFFB4FEBAF8C27D11F843" box="[355,369,1899,1924]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 10). Maximum body width 36 μm (mean = 34 μm;
<emphasis id="D8A3AB7CF76DFFB4FC0FF8C27F84F843" box="[982,996,1899,1924]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 10), measured near posterior margin of cephalothorax. Body slender and cylindrical without clear distinction between prosome and urosome. Sensillar pattern on body as figured. Hyaline frills of thoracic somites weakly developed and crenulated; those of genital double-somite and free abdominal somites strongly developed and consisting of rectangular digitate lappets (
<figureCitation id="72EC6BEBF76DFFAEFAB3F87F7CBEFF76" captionStart="FIGURE 1" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[191,1395,191,1904]" captionTargetId="figure-17@12.[179,1407,179,1917]" captionTargetPageId="12" captionText="FIGURE 1. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021482" httpUri="https://zenodo.org/record/14021482/files/figure.png" lastPageId="17" lastPageNumber="18" pageId="11" pageNumber="12">Figs 1A, B</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFF35FF3E7D70FF76" box="[236,272,151,177]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="17" pageNumber="18">2B</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFEC4FF3F7D23FF76" box="[285,323,150,177]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
). Somites connected by well-developed intersomitic membranes. Genital double-somite about 1.3 times longer than wide (measured in dorsal aspect); with two middorsal, four lateral and four ventral pores (
<figureCitation id="72EC6BEBF777FFAEFAB3FF127CBDFF3E" captionStart="FIGURE 1" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[191,1395,191,1904]" captionTargetId="figure-17@12.[179,1407,179,1917]" captionTargetPageId="12" captionText="FIGURE 1. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021482" httpUri="https://zenodo.org/record/14021482/files/figure.png" pageId="17" pageNumber="18">Figs 1A, B</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFF32FF777D71FF3E" box="[235,273,222,249]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
). Anal somite (
<figureCitation id="72EC6BEBF777FFAEFE64FF767E5CFF3E" box="[445,572,223,249]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="17" pageNumber="18">Figs 2B, C</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFD90FF777E0CFF3E" box="[585,620,222,249]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
) with paired, dorsally recurved, spinous processes near posterior border either side of anal opening, and two lateral pores near anterior margin; with two conspicuous pores near ventral posterior margin. Anal frill triradiate, minutely incised (giving a spinulose appearance); anal operculum slightly convex, without ornamentation.
</paragraph>
<caption id="BEA827E6F76AFFB3FF4EF83E7902F877" ID-DOI="http://doi.org/10.5281/zenodo.14021482" ID-Zenodo-Dep="14021482" box="[151,1378,1943,1968]" httpUri="https://zenodo.org/record/14021482/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1943,1967]" targetBox="[191,1395,191,1904]" targetPageId="12" targetType="figure">
<paragraph id="EA68776EF76AFFB3FF4EF83E7902F877" blockId="12.[151,1378,1943,1968]" box="[151,1378,1943,1968]" pageId="12" pageNumber="13">
<emphasis id="D8A3AB7CF76AFFB3FF4EF83E7D73F868" bold="true" box="[151,275,1943,1967]" pageId="12" pageNumber="13">FIGURE 1.</emphasis>
<taxonomicName id="2DD70CEDF76AFFB3FEC0F8317EACF868" authority="(Chappuis, 1954)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[281,716,1943,1968]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF76AFFB3FEC0F8317E76F877" box="[281,534,1944,1968]" italics="true" pageId="12" pageNumber="13">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF76AFFB3FDFAF8317EA5F868" author="Chappuis, P. - A." box="[547,709,1943,1968]" pageId="12" pageNumber="13" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
(♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal.
</paragraph>
</caption>
<caption id="BEA827E6F76BFFB2FF4EF83E7E1EF830" ID-DOI="http://doi.org/10.5281/zenodo.14021484" ID-Zenodo-Dep="14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1943,1967]" targetBox="[187,1401,193,1915]" targetPageId="13" targetType="figure">
<paragraph id="EA68776EF76BFFB2FF4EF83E7E1EF830" blockId="13.[151,1436,1943,2040]" pageId="13" pageNumber="14">
<emphasis id="D8A3AB7CF76BFFB2FF4EF83E7D74F868" bold="true" box="[151,276,1943,1967]" pageId="13" pageNumber="14">FIGURE 2.</emphasis>
<taxonomicName id="2DD70CEDF76BFFB2FEC2F8317EB4F868" authority="(Chappuis, 1954)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[283,724,1943,1968]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF76BFFB2FEC2F8317E78F877" box="[283,536,1944,1968]" italics="true" pageId="13" pageNumber="14">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF76BFFB2FDFEF8317EACF868" author="Chappuis, P. - A." box="[551,716,1943,1968]" pageId="13" pageNumber="14" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
: (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]
</paragraph>
</caption>
<caption id="BEA827E6F768FFB1FF4EF89D7D08F8B7" ID-DOI="http://doi.org/10.5281/zenodo.14021486" ID-Zenodo-Dep="14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1844,1868]" targetBox="[160,1424,322,1809]" targetPageId="14" targetType="figure">
<paragraph id="EA68776EF768FFB1FF4EF89D7D08F8B7" blockId="14.[151,1437,1844,1904]" pageId="14" pageNumber="15">
<emphasis id="D8A3AB7CF768FFB1FF4EF89D7D73F88B" bold="true" box="[151,275,1844,1868]" pageId="14" pageNumber="15">FIGURE 3.</emphasis>
<taxonomicName id="2DD70CEDF768FFB1FEC3F89D7EB2F88B" authority="(Chappuis, 1954)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[282,722,1844,1868]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF768FFB1FEC3F89D7E76F88B" box="[282,534,1844,1868]" italics="true" pageId="14" pageNumber="15">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF768FFB1FDFCF89D7EAAF88B" author="Chappuis, P. - A." box="[549,714,1844,1868]" pageId="14" pageNumber="15" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
: (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]
</paragraph>
</caption>
<caption id="BEA827E6F769FFB0FF4EF83E7C8CF813" ID-DOI="http://doi.org/10.5281/zenodo.14021488" ID-Zenodo-Dep="14021488" httpUri="https://zenodo.org/record/14021488/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1943,1967]" targetBox="[185,1411,193,1907]" targetPageId="15" targetType="figure">
<paragraph id="EA68776EF769FFB0FF4EF83E7C8CF813" blockId="15.[151,1436,1943,2004]" pageId="15" pageNumber="16">
<emphasis id="D8A3AB7CF769FFB0FF4EF83E7D77F868" bold="true" box="[151,279,1943,1967]" pageId="15" pageNumber="16">FIGURE 4.</emphasis>
<taxonomicName id="2DD70CEDF769FFB0FEFBF8317E82F868" authority="(Chappuis, 1954)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[290,738,1943,1968]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF769FFB0FEFBF8317E42F877" box="[290,546,1944,1968]" italics="true" pageId="15" pageNumber="16">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF769FFB0FDECF8317EBBF868" author="Chappuis, P. - A." box="[565,731,1943,1968]" pageId="15" pageNumber="16" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
(♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior.
</paragraph>
</caption>
<caption id="BEA827E6F776FFAFFF4EF83E7D81F813" ID-DOI="http://doi.org/10.5281/zenodo.14021490" ID-Zenodo-Dep="14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1943,1967]" targetBox="[203,1384,193,1907]" targetPageId="16" targetType="figure">
<paragraph id="EA68776EF776FFAFFF4EF83E7D81F813" blockId="16.[151,1437,1943,2004]" pageId="16" pageNumber="17">
<emphasis id="D8A3AB7CF776FFAFFF4EF83E7D72F877" bold="true" box="[151,274,1943,1968]" pageId="16" pageNumber="17">FIGURE 5.</emphasis>
<taxonomicName id="2DD70CEDF776FFAFFEC0F8317EAEF868" authority="(Chappuis, 1954)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[281,718,1943,1968]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF776FFAFFEC0F8317E74F877" box="[281,532,1944,1968]" italics="true" pageId="16" pageNumber="17">Neoleptastacus acanthus</emphasis>
(
<bibRefCitation id="8E460A9FF776FFAFFDFBF8317EA6F868" author="Chappuis, P. - A." box="[546,710,1943,1968]" pageId="16" pageNumber="17" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
: (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral.
</paragraph>
</caption>
<paragraph id="EA68776EF777FFAEFF1EFEC67D1DFDA6" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
Caudal rami (
<figureCitation id="72EC6BEBF777FFAEFE84FEC67DB8FE4E" box="[349,472,367,393]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="17" pageNumber="18">Figs 2B, C</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFE3AFEC77E66FE4E" box="[483,518,366,393]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
) about three times longer than wide (measured in dorsal view from anterior margin to apex of spinous process), distinctly tapering posteriorly; with a pore near ventral proximal margin (
<figureCitation id="72EC6BEBF777FFAEFAEBFE3A79EFFE6A" box="[1330,1423,402,429]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">Fig. 5A</figureCitation>
); outer distal corner produced into posteriorly directed, dorsally recurved spinous process; no spinular ornamentation discernible. Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, naked, located between seta V and posterior spinous process; seta V long, naked, with proximal fracture plane (
<figureCitation id="72EC6BEBF777FFAEFE9AFD8A7DFFFDFA" box="[323,415,546,573]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="17" pageNumber="18">Fig. 5A</figureCitation>
); seta VI small, naked and located at inner distal corner; seta VII distinctly foliaceous and tri-articulate at base.
</paragraph>
<paragraph id="EA68776EF777FFAEFF1EFDC27E38FD6E" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
Rostrum (
<figureCitation id="72EC6BEBF777FFAEFEEFFDC27DEAFD42" box="[310,394,619,645]" captionStart="FIGURE 1" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[191,1395,191,1904]" captionTargetId="figure-17@12.[179,1407,179,1917]" captionTargetPageId="12" captionText="FIGURE 1. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.14021482" httpUri="https://zenodo.org/record/14021482/files/figure.png" pageId="17" pageNumber="18">Fig. 1C</figureCitation>
) small, broadly subtriangular, tapering distally; apical part lobate and demarcated by bilateral constrictions, with two delicate sensilla.
</paragraph>
<paragraph id="EA68776EF777FFAEFF1EFD1A793DFC9A" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
Antennule (
<figureCitation id="72EC6BEBF777FFAEFE89FD1A7DCDFD0A" box="[336,429,691,717]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="17" pageNumber="18">Fig. 3A</figureCitation>
) long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about 3.4 times longer than wide. Segment 4 with long aesthetasc (L: 45 μm) fused at base with seta. Distal segment with eight setae (two distinctly spatulate; indicated by arrows in
<figureCitation id="72EC6BEBF777FFAEFB83FD5278D2FCD2" box="[1114,1202,763,789]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="17" pageNumber="18">Fig. 3A</figureCitation>
) and apical acrothek consisting of short aesthetasc (L: 20 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[8 + acrothek].
</paragraph>
<paragraph id="EA68776EF777FFAEFF1EFCCE7FD7FBBA" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
Antenna (
<figureCitation id="72EC6BEBF777FFAEFEEDFCCE7DCCFC46" box="[308,428,871,897]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="17" pageNumber="18">Fig. 2D, E</figureCitation>
). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment forming incompletely fused allobasis, about 2.9 times as long as maximum width; original basis-endopod boundary marked by transverse surface suture at level of exopodal articulation; proximal part representing original basis with longitudinal row of small spinules near base of exopod. Exopod one-segmented, unornamented, and elongate, with a naked apical seta (about 1.3 times longer than exopod). Free endopodal segment with few lateral spinules proximally and transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in
<figureCitation id="72EC6BEBF777FFAEFF6FFB967D52FB9E" box="[182,306,1087,1113]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="17" pageNumber="18">Fig. 2D, E</figureCitation>
); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta.
</paragraph>
<paragraph id="EA68776EF777FFAEFF1EFB2E7F1BF9EA" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
Mandible, maxillule, maxilla and maxilliped as in
<taxonomicName id="2DD70CEDF777FFAEFD24FB2E7FDAFB66" authorityName="Nicholls" authorityYear="1945" box="[765,954,1159,1185]" class="Copepoda" family="Leptastacidae" genus="Paraleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spinicaudatus">
<emphasis id="D8A3AB7CF777FFAEFD24FB2E7FDAFB66" box="[765,954,1159,1185]" italics="true" pageId="17" pageNumber="18">N. spinicaudatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8E460A9FF777FFAEFC2CFB2E78C1FB66" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[1013,1185,1159,1185]" pageId="17" pageNumber="18" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF777FFAEFBFFFB2E783EFB66" box="[1062,1118,1159,1185]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2008
</bibRefCitation>
:
<figureCitation id="72EC6BEBF777FFAEFB75FB2E7955FB66" box="[1196,1333,1159,1185]" captionStart="FIGURE 16" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@55.[170,1418,181,1919]" captionTargetPageId="55" captionText="FIGURE 16. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14021512" httpUri="https://zenodo.org/record/14021512/files/figure.png" pageId="17" pageNumber="18">Figs 16D, E</figureCitation>
;
<figureCitation id="72EC6BEBF777FFAEFA98FB2E79EFFB66" box="[1345,1423,1159,1185]" captionStart="FIGURE 17" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[205,1381,188,1907]" captionTargetId="figure-17@56.[194,1393,181,1919]" captionTargetPageId="56" captionText="FIGURE 17. Phreatipontia speluncae (Cottarelli, Bruno &amp; Venanzetti, 1994) comb. nov.: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]." figureDoi="http://doi.org/10.5281/zenodo.14021514" httpUri="https://zenodo.org/record/14021514/files/figure.png" pageId="17" pageNumber="18">17E, F</figureCitation>
). Mandibular gnathobase elongate, about as long as palp; with several curved, minute teeth and one tiny recurved seta at dorsal corner. Mandibular palp uniramous, consisting of elongate, unisetose basis and one-segmented endopod with one inner, two outer, and two apical setae. Maxillule comprising praecoxa, coxa, basis and vestigial rami; praecoxal arthrite with one anterior surface seta and five spines and two setae around distal margin; coxal endite cylindrical, with two recurved spines; basis elongate, with rami completely incorporated; basal armature consisting of three apical setae; exopod and endopod represented by one and three setae, respectively. Maxilla comprising syncoxa, allobasis and endopod; syncoxa with two cylindrical endites, proximal endite with three setae (one fused at base) and distal endite with two setae (one fused at base); allobasis drawn out into long claw with one accessory seta; endopod one-segmented, with three setae; all elements naked. Maxilliped comprising syncoxa, basis and endopod; syncoxa longer than wide, unarmed, with few spinules; basis elongate and unarmed; endopod with small accessory seta and slightly curved claw, bearing strong, subterminal spinule.
</paragraph>
<paragraph id="EA68776EF777FFAEFF1EF99E7EE1F88A" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="18">
P1 (
<figureCitation id="72EC6BEBF777FFAEFF2FF99E7D33F996" box="[246,339,1591,1617]" captionStart="FIGURE 4" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[185,1411,193,1907]" captionTargetId="figure-17@15.[173,1414,181,1919]" captionTargetPageId="15" captionText="FIGURE 4. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021488" httpUri="https://zenodo.org/record/14021488/files/figure.png" pageId="17" pageNumber="18">Fig. 4A</figureCitation>
). Intercoxal sclerite wide and subrectangular. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis with spinular row near base of endopod; anterior surface with a small inner spine near medial margin. Exopod three-segmented; exp-1 and -2 with several spinules around outer margin, exp-3 with single spinule; exp-1 longest, with unipinnate outer spine; exp-2 without outer element; exp-3 with short unipinnate outer spine, a long curved unipinnate spine and two geniculate setae distally. Endopod two-segmented, not prehensile, longer than exopod; enp-1 about 1.25 times longer than enp-2, with a serrate seta arising from halfway down inner margin and three coarse spinules along outer margin; enp-2 without spinules, distal margin with naked outer spine and geniculate inner seta.
</paragraph>
<paragraph id="EA68776EF777FFADFF1EF8FE7D12FDFA" blockId="17.[151,1437,150,2013]" lastBlockId="18.[151,1437,151,573]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
P2–P4 (
<figureCitation id="72EC6BEBF777FFAEFEFBF8FE7DC1F8B6" box="[290,417,1879,1905]" captionStart="FIGURE 4" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[185,1411,193,1907]" captionTargetId="figure-17@15.[173,1414,181,1919]" captionTargetPageId="15" captionText="FIGURE 4. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021488" httpUri="https://zenodo.org/record/14021488/files/figure.png" pageId="17" pageNumber="18">Fig. 4B–D</figureCitation>
). Intercoxal sclerites rectangular (P2) or squarish (P3–P4) with concave ventral margins. Praecoxae triangular, small and naked. Coxae squarish and without ornamentation. Bases smaller than coxae, with a spinular row near base of endopod (P3–P4) and a few spinules around outer corner in P2 and P4; anterior surface with a pore; outer basal seta absent in P2, long and plumose in P3–P4. Exopods three-segmented; segments with coarse spinular ornamentation, as illustrated; outer spine of exp-1 and -2 naked (except for P3 exp-2 sparsely unipinnate); exp-3 with an outer unipinnate spine, and two setae distally (one unipinnate and one bipinnate in P2–P3 or two bipinnate setae in P4); P4 exp-2 elongate, distinctly longer than exp-1; inner seta of P4 exp-3 serrate in distal half and originating near distal margin from posterior surface. Endopods two-segmented; P2–P4 enp-1 unarmed, about 1.1, 1.6, and 4.1 times longer than their respective distal segments, with few coarse spinules along outer margin as figured, but without ornamentation along inner margin (except for single minute spinule near inner distal corner of P4 enp-1); P2 enp-2 with long, apically serrate, backwardly directed seta near proximal margin and two unequal unipinnate setae around distal margin, with two spinules along outer margin subdistally and with one spinule at inner distal corner; P3 enp-2 with three coarse spinules on anterior surface, apical margin with long, unipinnate, inner seta and short, naked, outer spine; P4 enp-2 with a spinule halfway along outer margin, apical margin with long, distally serrate and basally fused, inner seta, and long, unipinnate, outer seta. Spine and seta formula as follows:
</paragraph>
<paragraph id="EA68776EF774FFADFE6EFDC27F76FCD2" pageId="18" pageNumber="19">
<table id="98D785CEF7740040FF2EFDC27F5CFCD2" box="[247,828,619,789]" gridcols="3" gridrows="4" inLine="true" pageId="18" pageNumber="19">
<tr id="54E7752CF7740040FF2EFDC27F5CFD42" box="[247,828,619,645]" gridrow="0" pageId="18" pageNumber="19" rowspan-0="1">
<th id="17361C50F7740040FE6EFDC27E6EFD42" box="[439,526,619,645]" gridcol="1" gridrow="0" pageId="18" pageNumber="19">Exopod</th>
<th id="17361C50F7740040FD0EFDC27F5CFD42" box="[727,828,619,645]" gridcol="2" gridrow="0" pageId="18" pageNumber="19">Endopod</th>
</tr>
<tr id="54E7752CF7740040FF2EFD1A7F5CFD0A" box="[247,828,691,717]" gridrow="1" pageId="18" pageNumber="19">
<th id="17361C50F7740040FF2EFD1A7D75FD0A" box="[247,277,691,717]" gridcol="0" gridrow="1" pageId="18" pageNumber="19">P2</th>
<td id="17361C50F7740040FE6EFD1A7E6EFD0A" box="[439,526,691,717]" gridcol="1" gridrow="1" pageId="18" pageNumber="19">0.0.021</td>
<td id="17361C50F7740040FD0EFD1A7F5CFD0A" box="[727,828,691,717]" gridcol="2" gridrow="1" pageId="18" pageNumber="19">0.120</td>
</tr>
<tr id="54E7752CF7740040FF2EFD7E7F5CFD36" box="[247,828,727,753]" gridrow="2" pageId="18" pageNumber="19">
<th id="17361C50F7740040FF2EFD7E7D75FD36" box="[247,277,727,753]" gridcol="0" gridrow="2" pageId="18" pageNumber="19">P3</th>
<td id="17361C50F7740040FE6EFD7E7E6EFD36" box="[439,526,727,753]" gridcol="1" gridrow="2" pageId="18" pageNumber="19">0.0.021</td>
<td id="17361C50F7740040FD0EFD7E7F5CFD36" box="[727,828,727,753]" gridcol="2" gridrow="2" pageId="18" pageNumber="19">0.020</td>
</tr>
<tr id="54E7752CF7740040FF2EFD527F5CFCD2" box="[247,828,763,789]" gridrow="3" pageId="18" pageNumber="19">
<th id="17361C50F7740040FF2EFD527D75FCD2" box="[247,277,763,789]" gridcol="0" gridrow="3" pageId="18" pageNumber="19">P4</th>
<td id="17361C50F7740040FE6EFD527E6EFCD2" box="[439,526,763,789]" gridcol="1" gridrow="3" pageId="18" pageNumber="19">0.0.121</td>
<td id="17361C50F7740040FD0EFD527F5CFCD2" box="[727,828,763,789]" gridcol="2" gridrow="3" pageId="18" pageNumber="19">0.020</td>
</tr>
</table>
</paragraph>
<paragraph id="EA68776EF774FFADFF4EFCEA7E53FC0E" blockId="18.[151,1437,834,2013]" pageId="18" pageNumber="19">
Fifth legs (
<figureCitation id="72EC6BEBF774FFADFECEFCEA7D11FC9A" box="[279,369,834,861]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="18" pageNumber="19">Fig. 5A</figureCitation>
) closely set together, almost touching medially. Baseoendopod and exopod fused, forming a subrectangular plate; anterior surface with two pores; inner distal corner with strong, minutely bipinnate, spinous process (homologous to inner spine); distal margin with plumose outer basal seta, one long, naked seta, and two short, equally long, bipinnate spines.
</paragraph>
<paragraph id="EA68776EF774FFADFF1EFC7B7FD9FB9E" blockId="18.[151,1437,834,2013]" pageId="18" pageNumber="19">
<emphasis id="D8A3AB7CF774FFADFF1EFC7B7DBAFC2A" bold="true" box="[199,474,978,1005]" pageId="18" pageNumber="19">Redescription of male.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 336 µm (mean = 325 μm;
<emphasis id="D8A3AB7CF774FFADFEBEFC5E7D15FBD7" box="[359,373,1015,1040]" italics="true" pageId="18" pageNumber="19">n</emphasis>
= 6). Maximum body width 31 μm (mean = 32 μm;
<emphasis id="D8A3AB7CF774FFADFC00FC5E7F87FBD7" box="[985,999,1015,1040]" italics="true" pageId="18" pageNumber="19">n</emphasis>
= 6), measured at posterior margin of cephalothorax. Body ornamentation (
<figureCitation id="72EC6BEBF774FFADFD98FBB27EFCFBF2" box="[577,668,1051,1077]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[187,1401,193,1915]" captionTargetId="figure-17@13.[175,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" figureDoi="http://doi.org/10.5281/zenodo.14021484" httpUri="https://zenodo.org/record/14021484/files/figure.png" pageId="18" pageNumber="19">Fig. 2A</figureCitation>
) essentially as in female. Sexual dimorphism in antennule, genital segmentation, P5, and P6. Spermatophore length approximately 53 µm.
</paragraph>
<paragraph id="EA68776EF774FFADFF1EFBCA7F4FFA92" blockId="18.[151,1437,834,2013]" pageId="18" pageNumber="19">
Antennule (
<figureCitation id="72EC6BEBF774FFADFE92FBCA7DC1FBBA" box="[331,417,1123,1149]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="18" pageNumber="19">Fig. 3B</figureCitation>
) 8-segmented, haplocer; geniculation between segments 6 and 7. Segment 1 with a slender naked seta; segment 2 longest and about 3.6 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae and a spine; segment 4 an incomplete sclerite with two spiniform elements; segment 5 with six setae and a long aesthetasc (50 µm) fused basally to a slender seta; segments 6 and 7 with a seta; distal segment with seven setae (three of which spatulate and indicated by arrows in
<figureCitation id="72EC6BEBF774FFADFC4BFB5A7F85FACA" box="[914,997,1267,1293]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="18" pageNumber="19">Fig. 3B</figureCitation>
) and apical acrothek consisting of short aesthetasc (12 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4 + 2 spines], 4-[2 spines], 5-[6 + (1 + ae)], 6-[1], 7-[1], 8-[7 + acrothek].
</paragraph>
<paragraph id="EA68776EF774FFADFF1EFAF778C0FABE" blockId="18.[151,1437,834,2013]" box="[199,1184,1374,1401]" pageId="18" pageNumber="19">
P5 (
<figureCitation id="72EC6BEBF774FFADFF2DFAF67D2AFABE" box="[244,330,1374,1401]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="18" pageNumber="19">Fig. 5B</figureCitation>
) with armature as in female but inner spinous process slightly more slender.
</paragraph>
<paragraph id="EA68776EF774FFADFF1EFA2A7F0CFA06" blockId="18.[151,1437,834,2013]" pageId="18" pageNumber="19">
Sixth legs (
<figureCitation id="72EC6BEBF774FFADFE9FFA2A7DFCFA5A" box="[326,412,1410,1437]" captionStart="FIGURE 5" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1384,193,1907]" captionTargetId="figure-17@16.[191,1396,181,1919]" captionTargetPageId="16" captionText="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." figureDoi="http://doi.org/10.5281/zenodo.14021490" httpUri="https://zenodo.org/record/14021490/files/figure.png" pageId="18" pageNumber="19">Fig. 5B</figureCitation>
) asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer seta and a short inner spine, both elements naked.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F774FFACFF1EFA627E68FBBA" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="discussion">
<paragraph id="EA68776EF774FFADFF1EFA627D1DF8B6" blockId="18.[151,1437,834,2013]" pageId="18" pageNumber="19">
<emphasis id="D8A3AB7CF774FFADFF1EFA627D5BFA22" bold="true" box="[199,315,1483,1509]" pageId="18" pageNumber="19">Remarks.</emphasis>
<bibRefCitation id="8E460A9FF774FFADFE9CFA627DB6FA22" author="Kunz, H." box="[325,470,1483,1509]" pageId="18" pageNumber="19" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
treated
<taxonomicName id="2DD70CEDF774FFADFDEEFA627EB0FA22" baseAuthorityName="Chappuis" baseAuthorityYear="1955" box="[567,720,1483,1509]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF774FFADFDEEFA627EB0FA22" box="[567,720,1483,1509]" italics="true" pageId="18" pageNumber="19">A. longiremis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF774FFADFCD4FA627FC7FA23" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Lang" baseAuthorityYear="1965" box="[781,935,1483,1509]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="accraensis">
<emphasis id="D8A3AB7CF774FFADFCD4FA627FC7FA23" box="[781,935,1483,1509]" italics="true" pageId="18" pageNumber="19">A. accraensis</emphasis>
</taxonomicName>
as subspecies of
<taxonomicName id="2DD70CEDF774FFADFBACFA627892FA22" authorityName="Chappuis" authorityYear="1954" box="[1141,1266,1483,1509]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF774FFADFBACFA627892FA22" box="[1141,1266,1483,1509]" italics="true" pageId="18" pageNumber="19">A. acantha</emphasis>
</taxonomicName>
.
<bibRefCitation id="8E460A9FF774FFADFADAFA6279FCFA22" author="Bodin, P." box="[1283,1436,1483,1509]" pageId="18" pageNumber="19" pagination="265 - 278" refId="ref45222" refString="Bodin, P. (1976) Catalogue des nouveaux Copepodes Harpacticoides marins. Additif no. 2. Tethys, 7 (2 - 3), 265 - 278." type="journal article" year="1976">Bodin (1976)</bibRefCitation>
adopted the division of
<taxonomicName id="2DD70CEDF774FFADFE46FA467E78F9CE" authorityName="Chappuis" authorityYear="1954" box="[415,536,1519,1545]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF774FFADFE46FA467E78F9CE" box="[415,536,1519,1545]" italics="true" pageId="18" pageNumber="19">A. acantha</emphasis>
</taxonomicName>
in three subspecies but reinstated
<taxonomicName id="2DD70CEDF774FFADFC41FA467850F9CE" baseAuthorityName="Chappuis" baseAuthorityYear="1955" box="[920,1072,1519,1545]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="D8A3AB7CF774FFADFC41FA467850F9CE" box="[920,1072,1519,1545]" italics="true" pageId="18" pageNumber="19">N. longiremis</emphasis>
</taxonomicName>
in later editions of his catalogue (
<bibRefCitation id="8E460A9FF774FFADFF46F9BA7D46F9EA" author="Bodin, P." box="[159,294,1555,1581]" pageId="18" pageNumber="19" refId="ref45254" refString="Bodin, P. (1979) Catalogue des nouveaux Copepodes Harpacticoides marins (Nouvelle edition). Universite de Bretagne Occidentale, Brest, 228 pp." type="book" year="1979">Bodin 1979</bibRefCitation>
,
<bibRefCitation id="8E460A9FF774FFADFEE8F9BA7D0CF9EA" author="Bodin, P." box="[305,364,1555,1581]" pageId="18" pageNumber="19" refId="ref45282" refString="Bodin, P. (1988) Catalogue des nouveaux Copepodes Harpacticoides marins (Edition 1988). Universite de Bretagne Occidentale, Laboratoire d'Oceanographie biologique, Brest, 288 pp." type="book" year="1988">1988</bibRefCitation>
,
<bibRefCitation id="8E460A9FF774FFADFEAFF9BA7DD2F9EA" author="Bodin, P." box="[374,434,1555,1581]" pageId="18" pageNumber="19" pagination="1 - 304" refId="ref45314" refString="Bodin, P. (1997) Catalogue of the new marine harpacticoid copepods (1997 Edition). Studiedocumenten van het K. B. I. N. / Documents de Travail de l'I. R. Sc. N. B., 89, 1 - 304." type="journal article" year="1997">1997</bibRefCitation>
).
<bibRefCitation id="8E460A9FF774FFADFE1CF9BA7E05F9EA" author="Kunz, H." box="[453,613,1555,1581]" pageId="18" pageNumber="19" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz’s (1971)</bibRefCitation>
subspecific classification was rejected by later revisers (
<bibRefCitation id="8E460A9FF774FFADFB05F9BA795EF9EA" author="Ito, T." box="[1244,1342,1555,1581]" pageId="18" pageNumber="19" pagination="47 - 55" refId="ref47150" refString="Ito, T. (1978) A new species of marine interstitial harpacticoid copepod of the genus Arenopontia from the Bonin Islands, southern Japan. Annotationes zoologicae japonenses, 51, 47 - 55." type="journal article" year="1978">Itô 1978</bibRefCitation>
;
<bibRefCitation id="8E460A9FF774FFADFA93F9BA7D10F996" author="Bodiou, J. - Y. &amp; Colomines, J. - C." pageId="18" pageNumber="19" pagination="55 - 64" refId="ref45366" refString="Bodiou, J. - Y. &amp; Colomines, J. - C. (1986) Harpacticoides (Copepodes) des Iles Crozet I. - Description d'une espece nouvelle du genre Arenopontia Kunz. Harpacticoids (Crustacea, Copepoda) from the Crozet Islands. Description of a new species of the genus Arenopontia Kunz. Vie et Milieu, 36, 55 - 64." type="journal article" year="1986">Bodiou &amp; Colomines 1986</bibRefCitation>
;
<bibRefCitation id="8E460A9FF774FFADFEA2F99E7D9CF996" author="Wells, J. B. J." box="[379,508,1591,1617]" pageId="18" pageNumber="19" pagination="1 - 872" refId="ref50074" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1 - 872. https: // doi. org / 10.11646 / zootaxa. 1568.1.1" type="journal article" year="2007">Wells 2007</bibRefCitation>
;
<bibRefCitation id="8E460A9FF774FFADFDD1F99E7ED3F996" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[520,691,1591,1617]" pageId="18" pageNumber="19" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF774FFADFDE3F99E7E12F996" box="[570,626,1591,1617]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2008
</bibRefCitation>
) who treated all three subspecific taxa as valid species. Within the
<taxonomicName id="2DD70CEDF774FFADFF4EF9F27C9CF9B2" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[151,252,1627,1653]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF774FFADFF4EF9F27C9CF9B2" box="[151,252,1627,1653]" italics="true" pageId="18" pageNumber="19">acanthus</emphasis>
</taxonomicName>
-subgroup,
<taxonomicName id="2DD70CEDF774FFADFEA5F9F27E61F9B2" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[380,513,1627,1653]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF774FFADFEA5F9F27E61F9B2" box="[380,513,1627,1653]" italics="true" pageId="18" pageNumber="19">N. acanthus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DD70CEDF774FFADFD4DF9F27F5DF9B2" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" box="[660,829,1627,1653]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chaufriassei">
<emphasis id="D8A3AB7CF774FFADFD4DF9F27F5DF9B2" box="[660,829,1627,1653]" italics="true" pageId="18" pageNumber="19">N. chaufriassei</emphasis>
</taxonomicName>
primarily in the proportional lengths of the endopodal segments of P1–P3 (enp-1:enp-2 length ratio 1.25, 1.20, 1.65
<emphasis id="D8A3AB7CF774FFADFCB3F9D67FE1F95F" box="[874,897,1663,1688]" italics="true" pageId="18" pageNumber="19">vs</emphasis>
1.65, 0.90, 1.20, respectively), the length and appearance of the inner subdistal seta on P4 exp-3, and the morphology of the P
<quantity id="2D2FDA8BF774FFADFBECF90B7801F97A" box="[1077,1121,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="19" unit="in" value="5.0">5 in</quantity>
both sexes.
<taxonomicName id="2DD70CEDF774FFADFB28F90A7CB2F926" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Karanovic" baseAuthorityYear="2000" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="huysi">
<emphasis id="D8A3AB7CF774FFADFB28F90A7CB2F926" italics="true" pageId="18" pageNumber="19">Neoleptastacus huysi</emphasis>
</taxonomicName>
is morphologically very close to its Mediterranean congener,
<taxonomicName id="2DD70CEDF774FFADFC79F96E7848F926" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[928,1064,1735,1761]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF774FFADFC79F96E7848F926" box="[928,1064,1735,1761]" italics="true" pageId="18" pageNumber="19">N. acanthus</emphasis>
</taxonomicName>
, sharing the shape of the paired anal processes, the elongate P4 exp-2 (about 1.3 times the length of exp-1) and the morphology of P5. The only difference that distinguishes both species is the presence of only one outer spine on P1 exp-
<quantity id="2D2FDA8BF774FFADFB68F8A678BBF8EE" box="[1201,1243,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="19" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="2DD70CEDF774FFADFB3AF8A67920F8EE" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Karanovic" baseAuthorityYear="2000" box="[1251,1344,1807,1833]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="huysi">
<emphasis id="D8A3AB7CF774FFADFB3AF8A67920F8EE" box="[1251,1344,1807,1833]" italics="true" pageId="18" pageNumber="19">N. huysi</emphasis>
</taxonomicName>
, raising the suspicion that the
<typeStatus id="356CC9CCF774FFADFE52F89A7D8DF88A" box="[395,493,1843,1869]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
female (and only known specimen) of the latter species was based on an aberrant or damaged individual.
</paragraph>
<paragraph id="EA68776EF774FFACFF1EF8D27919FEA2" blockId="18.[151,1437,834,2013]" lastBlockId="19.[151,1437,150,1149]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<taxonomicName id="2DD70CEDF774FFADFF1EF8D27D82F852" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[199,482,1915,1941]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF774FFADFF1EF8D27D82F852" box="[199,482,1915,1941]" italics="true" pageId="18" pageNumber="19">Neoleptastacus acanthus</emphasis>
</taxonomicName>
has been recorded on multiple occasions in the Indian Ocean and the North and Southern Atlantic. Unfortunately, most of these records are not accompanied by illustrations which could have confirmed their authenticity. The species is widely distributed in the Western Mediterranean basin.
<bibRefCitation id="8E460A9FF774FFADFB3BF86A79FCF81A" author="Chappuis, P. - A." box="[1250,1436,1986,2013]" pageId="18" pageNumber="19" pagination="254 - 276" refId="ref45608" refString="Chappuis, P. - A. (1954) Harpacticides psammiques recoltes par Cl. Delamare Deboutteville en Mediterranee. Vie et Milieu, 4, 254 - 276." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
and
<bibRefCitation id="8E460A9FF775FFACFF1EFF3E7E4AFF76" author="Delamare Deboutteville, C." box="[199,554,150,177]" pageId="19" pageNumber="20" pagination="141 - 170" refId="ref46409" refString="Delamare Deboutteville, C. (1953 a) La faune des eaux souterraines littorales des plages de Tunisie. Vie et Milieu, 4, 141 - 170." type="journal article" year="1953">Delamare Deboutteville (1953a</bibRefCitation>
–
<bibRefCitation id="8E460A9FF775FFACFDE3FF3F7EEBFF76" author="Delamare Deboutteville, C." box="[570,651,150,177]" pageId="19" pageNumber="20" pagination="1 - 740" refId="ref46503" refString="Delamare Deboutteville, C. (1960) Biologie des eaux souterraines littorales et continentales. Vie et Milieu, Supplement 9, 1 - 740." type="journal article" year="1960">c, 1960</bibRefCitation>
) recorded it from
<collectingCountry id="92C037FEF775FFACFC80FF3E7FECFF76" box="[857,908,151,177]" name="Italy" pageId="19" pageNumber="20">Italy</collectingCountry>
(
<collectingRegion id="2813B98CF775FFACFC45FF3E7F80FF76" box="[924,992,151,177]" country="Italy" name="Lazio" pageId="19" pageNumber="20">Lazio</collectingRegion>
, Fregene),
<collectingCountry id="92C037FEF775FFACFB85FF3E78D0FF76" box="[1116,1200,151,177]" name="Algeria" pageId="19" pageNumber="20">Algeria</collectingCountry>
(El Kala = La Calle;
<collectingRegion id="2813B98CF775FFACFF4EFF127C85FF12" box="[151,229,187,213]" country="Algeria" name="Skikda" pageId="19" pageNumber="20">Skikda</collectingRegion>
= Philippeville;
<collectingRegion id="2813B98CF775FFACFE7EFF127DB6FF12" box="[423,470,187,213]" country="Algeria" name="Jijel" pageId="19" pageNumber="20">Jijel</collectingRegion>
= Djidjelli; Beni Saf; Terga = Plage Turgot; Mers el Hadjad = Port aux Poules) and
<collectingCountry id="92C037FEF775FFACFF4EFF767C8AFF3E" box="[151,234,223,249]" name="Tunisia" pageId="19" pageNumber="20">Tunisia</collectingCountry>
(Gamarth-Plage, Rass Salakta). Reliable Italian records are those from near Cagliari (
<bibRefCitation id="8E460A9FF775FFACFB6EFF767904FF3E" author="Cottarelli, V." box="[1207,1380,222,249]" pageId="19" pageNumber="20" pagination="65 - 72" refId="ref46044" refString="Cottarelli, V. (1975) Una nuova Arenopontia di acque interstiziali litorali della Sardegna (Crustacea, Copepoda, Harpacticoida). Fragmenta entomologica, 11, 65 - 72." type="journal article" year="1975">Cottarelli 1975</bibRefCitation>
) and Isola Tavolara (
<bibRefCitation id="8E460A9FF775FFACFE9BFEAA7EE4FEDA" author="Cottarelli, V. &amp; Venanzetti, F." box="[322,644,259,285]" pageId="19" pageNumber="20" pagination="183 - 235" refId="ref46340" refString="Cottarelli, V. &amp; Venanzetti, F. (1989) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. II. Cylindropsyllidae del meiobenthos di Montecristo e delle isole circumsarde (Crustacea, Copepoda, Harpacticoida). Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 87, 183 - 235." type="journal article" year="1989">Cottarelli &amp; Venanzetti 1989</bibRefCitation>
) in Sardinia, from Sperlonga in Lazio (
<bibRefCitation id="8E460A9FF775FFACFBECFEAA7882FEDA" author="Cottarelli, V." box="[1077,1250,259,285]" pageId="19" pageNumber="20" pagination="8 - 21" refId="ref45879" refString="Cottarelli, V. (1969) Un nuovo Crostaceo di acque interstiziali Italiane: Psammopsyllus pasquinii n. sp. - (Harpacticoida, Cylindropsyllidae). Rendiconti dell'Istituto Lombardo di Scienze e Lettere, (B) 103, 8 - 21." type="journal article" year="1969">Cottarelli 1969</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFB34FEAA7948FEDA" author="Cottarelli, V." box="[1261,1320,259,285]" pageId="19" pageNumber="20" pagination="105 - 132" refId="ref45924" refString="Cottarelli, V. (1971) Nuove Parastenocaris (Copepoda, Harpacticoida) dell'Italia centro-meridionale. Some new species of Parastenocaris (Copepoda, Harpacticoida) of the central and southern Italy. Rivista di Idrobiologia, 8 (1), 105 - 132." type="journal article" year="1971">1971</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFAEAFEAA7D70FE86" author="Cottarelli, V. &amp; Bruno, M. C. &amp; Venanzetti, F." pageId="19" pageNumber="20" pagination="471 - 485" refId="ref46257" refString="Cottarelli, V., Bruno, M. C. &amp; Venanzetti, F. (1994) Ricerche zoologiche della nave oceanografica &quot; Minerva &quot; (C. N. R.) sulle isole circumsarde. XVIII. First record in phreatic freshwater of harpacticoids belonging to the genus Arenopontia (Crustacea, Copepoda) and description of two new species. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 90, 471 - 485." type="journal article" year="1994">
Cottarelli
<emphasis id="D8A3AB7CF775FFACFF4EFE8E7CAEFE86" box="[151,206,295,321]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
1994
</bibRefCitation>
), and from two localities (Tombolo della Giannella,
<taxonomicName id="2DD70CEDF775FFACFC89FE8E7846FE86" authority="di Alberese" authorityName="di Alberese" box="[848,1062,295,321]" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Marina di Alberese</taxonomicName>
) near Porto S. Stefano in Tuscany (
<bibRefCitation id="8E460A9FF775FFACFF46FEE27D3CFEA2" author="Cottarelli, V." box="[159,348,331,357]" pageId="19" pageNumber="20" pagination="19 - 30" refId="ref46009" refString="Cottarelli, V. (1973 b) Paramesochridae (Copepoda, Harpacticoida) di acque interstiziali litorali Italiana. Rivista di Idrobiologia, 10 (1 - 2), 19 - 30." type="journal article" year="1973">Cottarelli 1973b</bibRefCitation>
).
<bibRefCitation id="8E460A9FF775FFACFEB6FEE27E87FEA2" author="Martinez Arbizu, P. &amp; Moura, G." box="[367,743,331,357]" pageId="19" pageNumber="20" pagination="55 - 77" refId="ref48048" refString="Martinez Arbizu, P. &amp; Moura, G. (1994) The phylogenetic position of the Cylindropsyllinae Sars (Copepoda, Harpacticoida) and the systematic status of the Leptopontiinae Lang. Zoologische Beitrage, Neue Folge, 35, 55 - 77." type="journal article" year="1994">Martínez Arbizu &amp; Moura (1994)</bibRefCitation>
published the only record from
<collectingCountry id="92C037FEF775FFACFB89FEE278F0FEA2" box="[1104,1168,331,357]" name="Spain" pageId="19" pageNumber="20">Spain</collectingCountry>
(
<collectingRegion id="2813B98CF775FFACFB47FEE27963FEA2" box="[1182,1283,331,357]" country="Spain" name="Comunitat Valenciana" pageId="19" pageNumber="20">Valencia</collectingRegion>
, El Saler).
</paragraph>
<paragraph id="EA68776EF775FFACFF1EFEC67F79FDA6" blockId="19.[151,1437,150,1149]" pageId="19" pageNumber="20">
Records from the Eastern Mediterranean include those from Elafonisi Beach and Pahia Ammos on the island of
<collectingRegion id="2813B98CF775FFACFF6FFE3A7C95FE6A" box="[182,245,403,429]" country="Greece" name="Kriti" pageId="19" pageNumber="20">Crete</collectingRegion>
,
<collectingCountry id="92C037FEF775FFACFED8FE3A7D30FE6A" box="[257,336,403,429]" name="Greece" pageId="19" pageNumber="20">Greece</collectingCountry>
(
<bibRefCitation id="8E460A9FF775FFACFEB9FE3A7E69FE6A" author="Sevastou, K." box="[352,521,402,429]" pageId="19" pageNumber="20" refId="ref49561" refString="Sevastou, K. (2005) On the ecology of meiofauna of the sandy shores of Crete: distribution patterns of meiobenthic copepods over space and time. PhD dissertation, University of Crete, Heraklion, 233 pp. [in Greek with English summary]" type="book" year="2005">Sevastou 2005</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFDCCFE3A7E9BFE6A" author="Sevastou, K. &amp; Lampadariou, N. &amp; Eleftheriou, A." box="[533,763,403,429]" pageId="19" pageNumber="20" pagination="205 - 214" refId="ref49609" refString="Sevastou, K., Lampadariou, N. &amp; Eleftheriou, A. (2011) Meiobenthic diversity in space and time: The case of harpacticoid copepods in two Mediterranean microtidal sandy beaches. Journal of Sea Research, 66, 205 - 214. https: // doi. org / 10.1016 / j. seares. 2011.06.004" type="journal article" year="2011">
Sevastou
<emphasis id="D8A3AB7CF775FFACFD5BFE3A7EDBFE6A" box="[642,699,403,429]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2011
</bibRefCitation>
) and from the Datça-Bozburun peninsula (
<collectingRegion id="2813B98CF775FFACFB04FE3A79F4FE6A" box="[1245,1428,403,429]" country="Turkey" name="Mugla" pageId="19" pageNumber="20">Muğla Province</collectingRegion>
) (
<bibRefCitation id="8E460A9FF775FFACFF46FE1E7D71FE16" author="Sak, S." box="[159,273,439,465]" pageId="19" pageNumber="20" refId="ref49361" refString="Sak, S. (2004) Arenopontiidae (Copepoda: Harpacticoida) familyasinin taksonomisi ve filogenisi. [Taxonomy and phylogeny of the family Arenopontiidae (Copepoda: Harpacticoida)]. PhD dissertation, University of Balikesir, Balikesir, 312 pp." type="book" year="2004">Sak 2004</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFEC4FE1E7DC2FE16" author="Alper, A." box="[285,418,439,465]" pageId="19" pageNumber="20" refId="ref44916" refString="Alper, A. (2009) Datca-Bozburun ozel cevre koruma bolgesi (Mugla, Turkiye) sahillerinin kumici ve fital Harpacticoida (Crustacea, Copepoda) faunasinin belirlenmesi. Determination of the interstitial and phytal Harpacticoida (Crustacea, Copepoda) fauna of the coast of Datca-Bozburun (Mugla, Turkey) specially protected area. PhD dissertation, University of Balikesir, Turkiye, 150 pp. [in Turkish with English summary]" type="book" year="2009">Alper 2009</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFE69FE1E7E1BFE16" author="Alper, A. &amp; Karaytug, S. &amp; Sak, S." box="[432,635,439,465]" pageId="19" pageNumber="20" pagination="16 - 28" refId="ref44992" refString="Alper, A., Karaytug, S. &amp; Sak, S. (2010) Interstitial and phytal Harpacticoida (Crustacea: Copepoda) inhabiting the mediolittoral zone of the Datca-Bozburun Peninsulas (Mugla, Turkey). SDU Journal of Science, 5, 16 - 28." type="journal article" year="2010">
Alper
<emphasis id="D8A3AB7CF775FFACFE23FE1E7E55FE16" box="[506,565,439,465]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2010
</bibRefCitation>
; present account) and Dilek peninsula (
<collectingRegion id="2813B98CF775FFACFB9EFE1E789CFE16" box="[1095,1276,439,465]" country="Turkey" name="Aydin" pageId="19" pageNumber="20">Aydın Province</collectingRegion>
) (
<bibRefCitation id="8E460A9FF775FFACFACCFE1E7CB1FE32" author="Alper, A. &amp; Sonmez, S. &amp; Sak, S. &amp; Karaytug, S." pageId="19" pageNumber="20" pagination="580 - 586" refId="ref45043" refString="Alper, A., Sonmez, S., Sak, S. &amp; Karaytug, S. (2015) Marine harpacticoid (Copepoda, Harpacticoida) fauna of the Dilek Peninsula (Aydin, Turkey). Turkish Journal of Zoology, 39, 580 - 586. https: // doi. org / 10.3906 / zoo- 1405 - 67" type="journal article" year="2015">
Alper
<emphasis id="D8A3AB7CF775FFACFAB8FE1E79FDFE16" box="[1377,1437,439,465]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2015
</bibRefCitation>
) in
<collectingCountry id="92C037FEF775FFACFF23FE727D36FE32" box="[250,342,475,501]" name="Turkey" pageId="19" pageNumber="20">Türkiye</collectingCountry>
.
<bibRefCitation id="8E460A9FF775FFACFEB9FE727E6BFE32" author="Masry, D." box="[352,523,475,501]" pageId="19" pageNumber="20" pagination="229 - 258" refId="ref48094" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry’s (1970)</bibRefCitation>
records from Nahariyya and Akhziv in
<collectingCountry id="92C037FEF775FFACFC1AFE727861FE32" box="[963,1025,475,501]" name="Israel" pageId="19" pageNumber="20">Israel</collectingCountry>
are the only published outliers in the Levantine Sea but note that his setal formula of P1 endopod and P4 exopod contradicts his illustrations, and that his report of an outer basal seta on P1–P2 is most probably false. The great variability alluded to by some authors (
<emphasis id="D8A3AB7CF775FFACFAADFD8A79FCFDFB" box="[1396,1436,547,572]" italics="true" pageId="19" pageNumber="20">e.g.</emphasis>
<bibRefCitation id="8E460A9FF775FFACFF4EFDEE7D2DFDA6" author="Karanovic, T." box="[151,333,583,609]" pageId="19" pageNumber="20" pagination="33 - 38" refId="ref47348" refString="Karanovic, T. (2000) Arenopontia (Neoleptastacus) huysi, sp. nov. (Crustacea, Copepoda, Harpacticoida) from marine interstitial of Montenegro (S. E. Europe). Helgoland marine Research, 54, 33 - 38. https: // doi. org / 10.1007 / s 101520050033" type="journal article" year="2000">Karanovic 2000</bibRefCitation>
) has not yet been established as genuine.
</paragraph>
<paragraph id="EA68776EF775FFACFF1EFDC27F76FCFE" blockId="19.[151,1437,150,1149]" pageId="19" pageNumber="20">
<bibRefCitation id="8E460A9FF775FFACFF1EFDC27DC5FD42" author="Bozic, B." box="[199,421,619,645]" pageId="19" pageNumber="20" pagination="869 - 873" refId="ref45435" refString="Bozic, B. (1967) Deux Copepodes Harpacticoides de l'Inde. Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 38, 869 - 873." type="journal article" year="1967">Božić’s (1967: 873</bibRefCitation>
,
<figureCitation id="72EC6BEBF775FFACFE6CFDC27E73FD42" box="[437,531,619,645]" captionStart="FIGURE 3" captionStartId="14.[151,250,1844,1868]" captionTargetBox="[160,1424,322,1809]" captionTargetId="figure-17@14.[151,1436,307,1821]" captionTargetPageId="14" captionText="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" figureDoi="http://doi.org/10.5281/zenodo.14021486" httpUri="https://zenodo.org/record/14021486/files/figure.png" pageId="19" pageNumber="20">Fig. 3-3</figureCitation>
,
<figureCitation id="72EC6BEBF775FFACFDFBFDC27E51FD42" box="[546,561,619,645]" captionStart="FIGURE 4" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[185,1411,193,1907]" captionTargetId="figure-17@15.[173,1414,181,1919]" captionTargetPageId="15" captionText="FIGURE 4. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.14021488" httpUri="https://zenodo.org/record/14021488/files/figure.png" pageId="19" pageNumber="20">4</figureCitation>
) record from the Indian west coast (locality unknown but possibly
<collectingRegion id="2813B98CF775FFACFA91FDC279F4FD42" box="[1352,1428,619,645]" country="India" name="Kerala" pageId="19" pageNumber="20">Kerala</collectingRegion>
) conceivably refers to
<taxonomicName id="2DD70CEDF775FFACFE49FD267E64FD6E" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Rao" baseAuthorityYear="1967" box="[400,516,655,681]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="D8A3AB7CF775FFACFE49FD267E64FD6E" box="[400,516,655,681]" italics="true" pageId="19" pageNumber="20">N. indicus</emphasis>
</taxonomicName>
or a related species (see below). His illustrations of the female P5 and the anal somite including the caudal rami in dorsal aspect are of no use in deciding the identity of his material.
<bibRefCitation id="8E460A9FF775FFACFAD3FD1A79FCFD0A" author="Bozic, B." box="[1290,1436,691,717]" pageId="19" pageNumber="20" pagination="869 - 873" refId="ref45435" refString="Bozic, B. (1967) Deux Copepodes Harpacticoides de l'Inde. Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 38, 869 - 873." type="journal article" year="1967">Božić (1967)</bibRefCitation>
stated that the body length is about 400 μm and that the number of eggs in the egg-sac ranges between one and four.
<bibRefCitation id="8E460A9FF775FFACFF4EFD527D18FCD2" author="Munro, A. L. S. &amp; Wells, J. B. J. &amp; McIntyre, A. D." box="[151,376,763,789]" pageId="19" pageNumber="20" pagination="297 - 315" refId="ref48389" refString="Munro, A. L. S., Wells, J. B. J. &amp; McIntyre, A. D. (1978) Energy flow in the flora and meiofauna of sandy beaches. Proceedings of the Royal Society of Edinburgh, 76 B, 297 - 315. https: // doi. org / 10.1017 / S 0269727000002943" type="journal article" year="1978">
Munro
<emphasis id="D8A3AB7CF775FFACFF35FD527D46FCD2" box="[236,294,763,789]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(1978)
</bibRefCitation>
also reported “
<taxonomicName id="2DD70CEDF775FFACFDF0FD527F76FCD2" authorityName="Chappuis" authorityYear="1954" box="[553,790,763,789]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF775FFACFDF0FD527F76FCD2" box="[553,790,763,789]" italics="true" pageId="19" pageNumber="20">Arenopontia acantha</emphasis>
</taxonomicName>
” as one of the most abundant harpacticoids in the sandy beach of Cherthala (formerly Shertallai) in
<collectingRegion id="2813B98CF775FFACFDA5FCB67EA9FCFE" box="[636,713,799,825]" country="India" name="Kerala" pageId="19" pageNumber="20">Kerala</collectingRegion>
,
<collectingCountry id="92C037FEF775FFACFD0CFCB67F72FCFE" box="[725,786,799,825]" name="India" pageId="19" pageNumber="20">India</collectingCountry>
.
</paragraph>
<paragraph id="EA68776EF775FFACFF1EFCEA7E68FBBA" blockId="19.[151,1437,150,1149]" pageId="19" pageNumber="20">
<bibRefCitation id="8E460A9FF775FFACFF1EFCEA7DEFFC9A" author="Lindgren, E. W." box="[199,399,835,861]" pageId="19" pageNumber="20" refId="ref47781" refString="Lindgren, E. W. (1972) Systematics and ecology of North Carolina marine Sandy-beach Harpacticoida (Copepoda: Crustacea). PhD dissertation, University of North Carolina, Chapel Hill, North Carolina, 212 pp." type="book" year="1972">Lindgren’s (1972</bibRefCitation>
,
<bibRefCitation id="8E460A9FF775FFACFE47FCEA7DB7FC9A" author="Lindgren, E. W." box="[414,471,835,861]" pageId="19" pageNumber="20" pagination="229 - 240" refId="ref47822" refString="Lindgren, E. W. (1976) Five species of Arenopontia (Copepoda, Harpacticoida) from a North Carolina beach, U. S. A. Crustaceana, 30, 229 - 240. https: // doi. org / 10.1163 / 156854076 X 00611" type="journal article" year="1976">1976</bibRefCitation>
) records from the beaches at Bogue Sound and Iron Steamer Pier,
<collectingRegion id="2813B98CF775FFACFB29FCEA79FCFC9A" box="[1264,1436,835,861]" country="United States of America" name="North Carolina" pageId="19" pageNumber="20">North Carolina</collectingRegion>
(
<collectingCountry id="92C037FEF775FFACFF79FCCE7C8EFC46" box="[160,238,871,897]" name="United States of America" pageId="19" pageNumber="20">U.S.A.</collectingCountry>
) are not accompanied by illustrations and are more likely based on the geographically close
<taxonomicName id="2DD70CEDF775FFACFAC1FCCE79F5FC47" baseAuthorityName="Cottarelli" baseAuthorityYear="1973" box="[1304,1429,871,896]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="gussoae">
<emphasis id="D8A3AB7CF775FFACFAC1FCCE79F5FC47" box="[1304,1429,871,896]" italics="true" pageId="19" pageNumber="20">N. gussoae</emphasis>
</taxonomicName>
. Various South African workers (
<emphasis id="D8A3AB7CF775FFACFDCFFC227E5EFC63" box="[534,574,907,932]" italics="true" pageId="19" pageNumber="20">e.g.</emphasis>
<bibRefCitation id="8E460A9FF775FFACFD93FC227F70FC62" author="McLachlan, A." box="[586,784,907,933]" pageId="19" pageNumber="20" pagination="213 - 223" refId="ref47872" refString="McLachlan, A. (1980) Intertidal zonation of macrofauna and stratification of meiofauna on high energy sandy beaches in the Eastern Cape, South Africa. Transactions of the Royal Society of South Africa, 44, 213 - 223. https: // doi. org / 10.1080 / 00359198009520563" type="journal article" year="1980">McLachlan 1980</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFCF8FC227F99FC62" author="Fricke, A. H. &amp; Hennig, H. F. - K. O. &amp; Orren, M. J." box="[801,1017,907,933]" pageId="19" pageNumber="20" pagination="59 - 77" refId="ref46530" refString="Fricke, A. H., Hennig, H. F. - K. O. &amp; Orren, M. J. (1981) Relationship between oil pollution and psammolittoral meiofauna density of two South African beaches. Marine environmental Research, 5, 59 - 77. https: // doi. org / 10.1016 / 0141 - 1136 (81) 90023 - 4" type="journal article" year="1981">
Fricke
<emphasis id="D8A3AB7CF775FFACFCACFC227FD2FC62" box="[885,946,907,933]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
1981
</bibRefCitation>
;
<bibRefCitation id="8E460A9FF775FFACFBD0FC22788AFC62" author="Hennig, H. F. - K. O. &amp; Eagle, G. A. &amp; Fielder, L. &amp; Fricke, A. H. &amp; Gledhill, W. J. &amp; Greenwood, P. J. &amp; Orren, M. J." box="[1033,1258,907,933]" pageId="19" pageNumber="20" pagination="45 - 60" refId="ref46600" refString="Hennig, H. F. - K. O., Eagle, G. A., Fielder, L., Fricke, A. H., Gledhill, W. J., Greenwood, P. J. &amp; Orren, M. J. (1983) Ratio and population density of psammolittoral meiofauna as a perturbation indicator of sandy beaches in South Africa. Environmental Monitoring and Assessment, 3, 45 - 60. https: // doi. org / 10.1007 / BF 00394031" type="journal article" year="1983">
Hennig
<emphasis id="D8A3AB7CF775FFACFBB1FC2278C5FC62" box="[1128,1189,907,933]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
1983
</bibRefCitation>
) have reported “
<taxonomicName id="2DD70CEDF775FFACFF7DFC067DF4FC0E" authorityName="Chappuis" authorityYear="1954" box="[164,404,943,969]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acantha">
<emphasis id="D8A3AB7CF775FFACFF7DFC067DF4FC0E" box="[164,404,943,969]" italics="true" pageId="19" pageNumber="20">Arenopontia acantha</emphasis>
</taxonomicName>
” in significant numbers in several beaches from around Cape Town in the west to Port Elizabeth and Algoa Bay in the east (Western Cape Province). It is highly likely that these authors were dealing with a different member of the
<taxonomicName id="2DD70CEDF775FFACFE67FC5E7E43FBD6" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[446,547,1015,1041]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF775FFACFE67FC5E7E43FBD6" box="[446,547,1015,1041]" italics="true" pageId="19" pageNumber="20">acanthus</emphasis>
</taxonomicName>
-group which is probably conspecific with the species recorded as
<taxonomicName id="2DD70CEDF775FFACFAC8FC5E7CD7FBF2" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D8A3AB7CF775FFACFAC8FC5E79FCFBD6" box="[1297,1436,1015,1041]" italics="true" pageId="19" pageNumber="20">Arenopontia</emphasis>
sp.
</taxonomicName>
from Algoa Bay (
<bibRefCitation id="8E460A9FF775FFACFEA7FBB27E8AFBF2" author="McLachlan, A. &amp; Furstenberg, J. P." box="[382,746,1051,1077]" pageId="19" pageNumber="20" pagination="61 - 71" refId="ref47925" refString="McLachlan, A. &amp; Furstenberg, J. P. (1977) Studies on the psammolittoral meiofauna of Algoa Bay. III. A quantitative analysis of the nematode and crustacean communities. Zoologica africana, 12, 61 - 71. https: // doi. org / 10.1080 / 00445096.1977.11447549" type="journal article" year="1977">McLachlan &amp; Furstenberg 1977</bibRefCitation>
).
<bibRefCitation id="8E460A9FF775FFACFD25FBB27FAAFBF2" author="Wandeness, A. P." box="[764,970,1051,1077]" pageId="19" pageNumber="20" refId="ref49795" refString="Wandeness, A. P. (1998) Estrutura espaco-temporal dos Copepoda Harpacticoida em um perfil na regiao de Macae, RJ. (Spatio-temporal structure of the Copepoda Harpacticoida on a transect in the region of Macae, RJ.). MSc dissertation, Universidade Federal Fluminense, Niteroi, Rio de Janeiro, 69 pp. [in Portuguese]" type="book" year="1998">Wandeness (1998)</bibRefCitation>
recorded
<taxonomicName id="2DD70CEDF775FFACFBE1FBB278DDFBF2" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[1080,1213,1051,1077]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="D8A3AB7CF775FFACFBE1FBB278DDFBF2" box="[1080,1213,1051,1077]" italics="true" pageId="19" pageNumber="20">N. acanthus</emphasis>
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from a sandy beach in the Macaé region in
<collectingRegion id="2813B98CF775FFACFE44FB967EE2FB9E" box="[413,642,1087,1113]" country="Brazil" name="Rio de Janeiro" pageId="19" pageNumber="20">Rio de Janeiro State</collectingRegion>
,
<collectingCountry id="92C037FEF775FFACFD57FB967EB5FB9E" box="[654,725,1087,1113]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
, but it is conceivable that this geographically disjunct record is also based on a different species.
</paragraph>
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</treatment>
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<mods:date id="D2EF5DBEA96050BA93F950FFCE555D11">2024</mods:date>
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<mods:number id="F574409E981CB8251938BB39DAF2C87B">5525</mods:number>
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<subSubSection id="A2CD24E5F772FFABFF4EFC077F7EFC0F" box="[151,798,942,969]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="EA68776EF772FFABFF4EFC077F7EFC0F" blockId="20.[151,798,942,969]" box="[151,798,942,969]" pageId="20" pageNumber="21">
<heading id="B120C002F772FFABFF4EFC077F7EFC0F" bold="true" box="[151,798,942,969]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
<taxonomicName id="2DD70CEDF772FFABFF4EFC077F7EFC0F" ID-CoL="46KMC" authority="(Chappuis &amp; Rouch, 1961)" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[151,798,942,969]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFF4EFC077F7EFC0F" bold="true" box="[151,798,942,969]" pageId="20" pageNumber="21">
<emphasis id="D8A3AB7CF772FFABFF4EFC077DA9FC0F" bold="true" box="[151,457,942,969]" italics="true" pageId="20" pageNumber="21">Neoleptastacus africanus</emphasis>
(
<bibRefCitation id="8E460A9FF772FFABFE03FC077F76FC0F" author="Chappuis, P. - A. &amp; Rouch, R." box="[474,790,942,969]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch, 1961</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F772FFABFF4EFC5A7FC7FB68" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="EA68776EF772FFABFF4EFC5A7E19FBCC" blockId="20.[151,1118,1011,1127]" box="[151,633,1011,1035]" pageId="20" pageNumber="21">
<taxonomicName id="2DD70CEDF772FFABFF4EFC5A7E19FBCC" ID-CoL="GFQ2" authority="Chappuis &amp; Rouch, 1961" authorityName="Chappuis &amp; Rouch" authorityYear="1961" box="[151,633,1011,1035]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="D8A3AB7CF772FFABFF4EFC5A7D11FBCC" box="[151,369,1011,1035]" italics="true" pageId="20" pageNumber="21">Arenopontia africana</emphasis>
<bibRefCitation id="8E460A9FF772FFABFEAEFC5A7E19FBCC" author="Chappuis, P. - A. &amp; Rouch, R." box="[375,633,1011,1035]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch, 1961</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="EA68776EF772FFABFF4EFBBB7F8EFBED" blockId="20.[151,1118,1011,1127]" box="[151,1006,1042,1066]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="CAC75040F772FFABFF4EFBBB7F8EFBED" box="[151,1006,1042,1066]" pageId="20" pageNumber="21">
<taxonomicName id="2DD70CEDF772FFABFF4EFBBB7F4EFBED" ID-CoL="GFQ2" authority="Chappuis &amp; Rouch, 1961" authorityName="Chappuis &amp; Rouch" authorityYear="1961" box="[151,814,1042,1066]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africana" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF772FFABFF4EFBBB7D75FBEE" box="[151,277,1042,1065]" italics="true" pageId="20" pageNumber="21">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF772FFABFEFDFBBB7DDFFBED" box="[292,447,1042,1066]" italics="true" pageId="20" pageNumber="21">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF772FFABFE14FBBB7E43FBED" box="[461,547,1042,1066]" italics="true" pageId="20" pageNumber="21">africana</emphasis>
<bibRefCitation id="8E460A9FF772FFABFDF0FBBB7F4EFBED" author="Chappuis, P. - A. &amp; Rouch, R." box="[553,814,1042,1066]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch, 1961</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF772FFABFCE1FBBB7F8EFBED" author="Wells, J. B. J." box="[824,1006,1042,1066]" page="324" pageId="20" pageNumber="21" year="1967">
<bibRefCitation id="8E460A9FF772FFABFCE1FBBB7F8EFBED" author="Wells, J. B. J." box="[824,1006,1042,1066]" pageId="20" pageNumber="21" pagination="189 - 358" refId="ref49859" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / S 0080456800024017" type="journal article" year="1967">Wells (1967: 324)</bibRefCitation>
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</paragraph>
<paragraph id="EA68776EF772FFABFF4EFB98783EFB8F" blockId="20.[151,1118,1011,1127]" box="[151,1118,1072,1097]" pageId="20" pageNumber="21">
<taxonomicName id="2DD70CEDF772FFABFF4EFB987E43FB8E" ID-CoL="GFQ2" authorityName="Chappuis &amp; Rouch" authorityYear="1961" box="[151,547,1072,1097]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africana" subGenus="Neoleptastacus">
<emphasis id="D8A3AB7CF772FFABFF4EFB987D75FB8F" box="[151,277,1073,1096]" italics="true" pageId="20" pageNumber="21">Arenopontia</emphasis>
(
<emphasis id="D8A3AB7CF772FFABFEFDFB997DDFFB8F" box="[292,447,1072,1096]" italics="true" pageId="20" pageNumber="21">Neoleptastacus</emphasis>
)
<emphasis id="D8A3AB7CF772FFABFE14FB987E43FB8E" box="[461,547,1073,1097]" italics="true" pageId="20" pageNumber="21">africana</emphasis>
</taxonomicName>
f.
<taxonomicName id="2DD70CEDF772FFABFDE6FB987FC0FB8F" ID-CoL="GFQ2" authority="Chappuis &amp; Rouch, 1961" authorityName="Chappuis &amp; Rouch" authorityYear="1961" box="[575,928,1072,1097]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="D8A3AB7CF772FFABFDE6FB987EF5FB8E" box="[575,661,1073,1097]" italics="true" pageId="20" pageNumber="21">africana</emphasis>
<bibRefCitation id="8E460A9FF772FFABFD42FB997FC0FB8F" author="Chappuis, P. - A. &amp; Rouch, R." box="[667,928,1072,1097]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch, 1961</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="6B76517FF772FFABFC73FB98783EFB8F" author="Kunz, H." box="[938,1118,1072,1097]" page="358" pageId="20" pageNumber="21" year="1971">
<bibRefCitation id="8E460A9FF772FFABFC73FB98783EFB8F" author="Kunz, H." box="[938,1118,1072,1097]" pageId="20" pageNumber="21" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971: 358)</bibRefCitation>
</treatmentCitation>
</paragraph>
<paragraph id="EA68776EF772FFABFF4EFBE67FF3FBA0" blockId="20.[151,1118,1011,1127]" box="[151,915,1103,1127]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="CAC75040F772FFABFF4EFBE67FF3FBA0" box="[151,915,1103,1127]" pageId="20" pageNumber="21">
<treatmentCitation id="6B76517FF772FFABFF4EFBE67FF3FBA0" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[151,915,1103,1127]" page=")" pageId="20" pageNumber="21" year="1961">
<taxonomicName id="2DD70CEDF772FFABFF4EFBE67FF3FBA0" ID-CoL="46KMC" authority="(Chappuis &amp; Rouch, 1961) Sak et al. (2008: 412)" authorityName="Sak et al." authorityPageNumber="412" authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[151,915,1103,1127]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFF4EFBE67DF8FBA0" box="[151,408,1103,1127]" italics="true" pageId="20" pageNumber="21">Neoleptastacus africanus</emphasis>
(
<bibRefCitation id="8E460A9FF772FFABFE7CFBE67ECAFBA0" author="Chappuis, P. - A. &amp; Rouch, R." box="[421,682,1103,1127]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch, 1961</bibRefCitation>
)
<bibRefCitation id="8E460A9FF772FFABFD6EFBE67FF3FBA0" author="Sak, S. &amp; Huys, R. &amp; Karaytug, S." box="[695,915,1103,1127]" pageId="20" pageNumber="21" pagination="409 - 458" refId="ref49406" refString="Sak, S., Huys, R. &amp; Karaytug, S. (2008) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00373. x" type="journal article" year="2008">
Sak
<emphasis id="D8A3AB7CF772FFABFD3AFBF97F77FBA0" box="[739,791,1103,1127]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
(2008: 412)
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</paragraph>
<paragraph id="EA68776EF772FFABFF4EFB3C7FC7FB68" blockId="20.[151,1437,1173,1992]" box="[151,935,1173,1199]" pageId="20" pageNumber="21">
<emphasis id="D8A3AB7CF772FFABFF4EFB3C7DF0FB68" bold="true" box="[151,400,1173,1199]" pageId="20" pageNumber="21">Original description.</emphasis>
<bibRefCitation id="8E460A9FF772FFABFE4EFB3C7EA4FB68" author="Chappuis, P. - A. &amp; Rouch, R." box="[407,708,1173,1199]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
: 606–607;
<figureCitation id="72EC6BEBF772FFABFCE6FB3C7FC3FB68" box="[831,931,1173,1199]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="12.[151,250,1943,1967]" captionStartId-1="13.[151,250,1943,1967]" captionStartId-2="14.[151,250,1844,1868]" captionStartId-3="15.[151,250,1943,1967]" captionStartId-4="16.[151,250,1943,1967]" captionStartId-5="31.[151,250,1943,1967]" captionStartId-6="32.[151,250,1943,1967]" captionStartId-7="33.[151,250,1936,1960]" captionTargetBox-0="[191,1395,191,1904]" captionTargetBox-1="[187,1401,193,1915]" captionTargetBox-2="[160,1424,322,1809]" captionTargetBox-3="[185,1411,193,1907]" captionTargetBox-4="[203,1384,193,1907]" captionTargetBox-5="[177,1407,192,1907]" captionTargetBox-6="[196,1400,187,1907]" captionTargetBox-7="[188,1400,186,1898]" captionTargetId-0="figure-17@12.[179,1407,179,1917]" captionTargetId-1="figure-17@13.[175,1413,181,1919]" captionTargetId-2="figure-17@14.[151,1436,307,1821]" captionTargetId-3="figure-17@15.[173,1414,181,1919]" captionTargetId-4="figure-17@16.[191,1396,181,1919]" captionTargetId-5="figure-17@31.[177,1410,181,1919]" captionTargetId-6="figure-17@32.[187,1400,181,1919]" captionTargetId-7="figure-17@33.[175,1412,181,1912]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionTargetPageId-5="31" captionTargetPageId-6="32" captionTargetPageId-7="33" captionText-0="FIGURE 1. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal." captionText-1="FIGURE 2. Neoleptastacus acanthus (Chappuis, 1954): (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E]" captionText-2="FIGURE 3. Neoleptastacus acanthus (Chappuis, 1954): (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows]" captionText-3="FIGURE 4. Neoleptastacus acanthus (Chappuis, 1954) (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." captionText-4="FIGURE 5. Neoleptastacus acanthus (Chappuis, 1954): (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral." captionText-5="FIGURE 6. Neoleptastacus emendatus sp. nov. (♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal; (D) anal somite and right caudal ramus, lateral." captionText-6="FIGURE 7. Neoleptastacus emendatus sp. nov. (♀): (A) urosome, ventral; (B) antennule, dorsal; (C) antenna, outer; (D) antennary endopod, medial; (E) rostrum, dorsal; (F) midventral hook-like processes [indicated by arrows] on pedigerous somites; (G) hook-like process, lateral. [lateral endopodal spines indicated by arrows in C–D]" captionText-7="FIGURE 8. Neoleptastacus emendatus sp. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi-0="http://doi.org/10.5281/zenodo.14021482" figureDoi-1="http://doi.org/10.5281/zenodo.14021484" figureDoi-2="http://doi.org/10.5281/zenodo.14021486" figureDoi-3="http://doi.org/10.5281/zenodo.14021488" figureDoi-4="http://doi.org/10.5281/zenodo.14021490" figureDoi-5="http://doi.org/10.5281/zenodo.14021492" figureDoi-6="http://doi.org/10.5281/zenodo.14021494" figureDoi-7="http://doi.org/10.5281/zenodo.14021496" httpUri-0="https://zenodo.org/record/14021482/files/figure.png" httpUri-1="https://zenodo.org/record/14021484/files/figure.png" httpUri-2="https://zenodo.org/record/14021486/files/figure.png" httpUri-3="https://zenodo.org/record/14021488/files/figure.png" httpUri-4="https://zenodo.org/record/14021490/files/figure.png" httpUri-5="https://zenodo.org/record/14021492/files/figure.png" httpUri-6="https://zenodo.org/record/14021494/files/figure.png" httpUri-7="https://zenodo.org/record/14021496/files/figure.png" pageId="20" pageNumber="21">Figs 1–8</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F772FFABFF1EFB107ECBFB3F" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="EA68776EF772FFABFF1EFB10786AFB14" blockId="20.[151,1437,1173,1992]" box="[199,1034,1209,1236]" pageId="20" pageNumber="21">
<emphasis id="D8A3AB7CF772FFABFF1EFB107D06FB14" bold="true" box="[199,358,1209,1235]" pageId="20" pageNumber="21">
<typeStatus id="356CC9CCF772FFABFF1EFB107D62FB14" box="[199,258,1209,1235]" pageId="20" pageNumber="21">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="92C037FEF772FFABFEB5FB107DD8FB14" box="[364,440,1209,1235]" name="Ghana" pageId="20" pageNumber="21">Ghana</collectingCountry>
,
<collectingRegion id="2813B98CF772FFABFE1AFB107EDBFB14" box="[451,699,1209,1236]" country="Ghana" name="Greater Accra" pageId="20" pageNumber="21">Greater Accra Region</collectingRegion>
,
<location id="EF0821B5F772FFABFD1DFB137F6AFB13" LSID="urn:lsid:plazi:treatment:627EC678F772FFABFF4EFC077D48F800:EF0821B5F772FFABFD1DFB137F6AFB13" box="[708,778,1210,1236]" country="Ghana" name="Accra" pageId="20" pageNumber="21" stateProvince="Greater Accra">Accra</location>
; in front of lighthouse.
</paragraph>
<paragraph id="EA68776EF772FFABFF1EFB747ECBFB3F" blockId="20.[151,1437,1173,1992]" box="[199,683,1245,1272]" pageId="20" pageNumber="21">
<emphasis id="D8A3AB7CF772FFABFF1EFB747D3EFB30" bold="true" box="[199,350,1245,1271]" pageId="20" pageNumber="21">Body length.</emphasis>
380 μm (
<specimenCount id="FCD1BCE7F772FFABFE14FB747D80FB30" box="[461,480,1245,1271]" count="1" pageId="20" pageNumber="21" type="female">♀</specimenCount>
); unknown for
<specimenCount id="FCD1BCE7F772FFABFD56FB777EC7FB3F" box="[655,679,1246,1272]" count="1" pageId="20" pageNumber="21" type="male">♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CD24E5F772FFABFF1EFAA87D48F800" pageId="20" pageNumber="21" type="discussion">
<paragraph id="EA68776EF772FFABFF1EFAA87D48F800" blockId="20.[151,1437,1173,1992]" pageId="20" pageNumber="21">
<emphasis id="D8A3AB7CF772FFABFF1EFAA87D5BFADC" bold="true" box="[199,315,1281,1307]" pageId="20" pageNumber="21">Remarks.</emphasis>
<taxonomicName id="2DD70CEDF772FFABFE9BFAA87E3DFADC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[322,605,1281,1307]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFE9BFAA87E3DFADC" box="[322,605,1281,1307]" italics="true" pageId="20" pageNumber="21">Neoleptastacus africanus</emphasis>
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belongs to the
<taxonomicName id="2DD70CEDF772FFABFCDEFAAB7F12FADC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Mielke" baseAuthorityYear="1982" box="[775,882,1282,1307]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="trisetosus">
<emphasis id="D8A3AB7CF772FFABFCDEFAAB7F12FADC" box="[775,882,1282,1307]" italics="true" pageId="20" pageNumber="21">trisetosus</emphasis>
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-group based on (1) P1 exp-1 without outer spine, (2) distal segment (exp-3, or exp-2 when exopod 2-segmented) with only one outer spine, (3) P2 exp-2 with very long outer setiform element (extending far beyond distal margin of exp-3), (4) P2–P3 endopods 1-segmented with reduced armature (010), and (5) outer seta of P4 enp-2 reduced. Within this group it is morphologically most similar to its geographically closest member,
<taxonomicName id="2DD70CEDF772FFABFD90FA3B7E84FA6C" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" box="[585,740,1425,1451]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angolensis" status="comb. nov.">
<emphasis id="D8A3AB7CF772FFABFD90FA3B7E84FA6C" box="[585,740,1425,1451]" italics="true" pageId="20" pageNumber="21">N. angolensis</emphasis>
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<emphasis id="D8A3AB7CF772FFABFD34FA387F0FFA6C" bold="true" box="[749,879,1425,1451]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="C3901607F772FFABFD34FA387F0FFA6C" box="[749,879,1425,1451]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
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, with which it shares the 3-segmented condition of the P1 exopod, a distinctly elongate P4 exopod, the reduced armature on the fifth legs, and the presence of a middorsal process on the anal operculum.
<bibRefCitation id="8E460A9FF772FFABFD45FA737F4DFA34" author="Kunz, H." box="[668,813,1497,1524]" pageId="20" pageNumber="21" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
listed seven differences between
<taxonomicName id="2DD70CEDF772FFABFB6EFA737926FA34" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[1207,1350,1497,1523]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFB6EFA737926FA34" box="[1207,1350,1497,1523]" italics="true" pageId="20" pageNumber="21">N. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD70CEDF772FFABFA5AFA737D6FF9D0" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angolensis" status="comb. nov.">
<emphasis id="D8A3AB7CF772FFABFA5AFA737D6FF9D0" italics="true" pageId="20" pageNumber="21">N. angolensis</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF772FFABFEC1FA547DFAF9D0" bold="true" box="[280,410,1533,1559]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="C3901607F772FFABFEC1FA547DFAF9D0" box="[280,410,1533,1559]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
but at least five of them appear unreliable or incorrect,
<emphasis id="D8A3AB7CF772FFABFBC4FA577825F9D0" box="[1053,1093,1534,1559]" italics="true" pageId="20" pageNumber="21">e.g.</emphasis>
(1)
<bibRefCitation id="8E460A9FF772FFABFBA1FA577968F9D0" author="Kunz, H." box="[1144,1288,1533,1560]" pageId="20" pageNumber="21" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
claimed that the outer spine on P1 exp-1 is present in
<taxonomicName id="2DD70CEDF772FFABFDA8F98B7E9EF9FC" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[625,766,1569,1595]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFDA8F98B7E9EF9FC" box="[625,766,1569,1595]" italics="true" pageId="20" pageNumber="21">N. africanus</emphasis>
</taxonomicName>
but the short outer element figured by
<bibRefCitation id="8E460A9FF772FFABFB1BF9887C82F998" author="Chappuis, P. - A. &amp; Rouch, R." pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
appears too small to be a genuine spine (when present, its size invariably approaches that of the outer spines on exp-3) and is interpreted here as a spinule (as found in the same position in other species of the genus); (2) according to
<bibRefCitation id="8E460A9FF772FFABFEF0F9277DDAF960" author="Kunz, H." box="[297,442,1677,1704]" pageId="20" pageNumber="21" pagination="348 - 357" refId="ref47685" refString="Kunz, H. (1971) Harpacticoiden (Crustacea Copepoda) von einem Sandstrand Angolas. Zoologischer Anzeiger, 186, 348 - 357." type="journal article" year="1971">Kunz (1971)</bibRefCitation>
, P4 enp-2 has only one seta but
<bibRefCitation id="8E460A9FF772FFABFCF8F9247828F960" author="Chappuis, P. - A. &amp; Rouch, R." box="[801,1096,1677,1703]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
clearly stated (and illustrated) that the long distal seta is accompanied by “... une fine soie au bord externe”; (3) the difference related to the antennary exopod (represented by seta
<emphasis id="D8A3AB7CF772FFABFD87F97F7E15F928" box="[606,629,1750,1775]" italics="true" pageId="20" pageNumber="21">vs</emphasis>
unisetose segment) is ambiguous because
<bibRefCitation id="8E460A9FF772FFABFBB0F97C79FCF928" author="Chappuis, P. - A. &amp; Rouch, R." box="[1129,1436,1749,1775]" pageId="20" pageNumber="21" pagination="605 - 614" refId="ref45768" refString="Chappuis, P. - A. &amp; Rouch, R. (1961) Harpacticoides psammiques d'une plage pres d'Accra (Ghana). Vie et Milieu, 11, 605 - 614." type="journal article" year="1961">Chappuis &amp; Rouch (1961)</bibRefCitation>
were not explicit about its condition (“Exopodite de l’antenne II, une soie courte et mince”; and (4) the pinnules bordering the anal operculum in
<taxonomicName id="2DD70CEDF772FFABFDDFF8B77EF1F8F0" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[518,657,1821,1847]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFDDFF8B77EF1F8F0" box="[518,657,1821,1847]" italics="true" pageId="20" pageNumber="21">N. africanus</emphasis>
</taxonomicName>
are not part of the opercular ornamentation but a misinterpretation of the underlying incised anal frill. Both species require thorough redescription but can be distinguished at present by the shape/curvature of the P5 and caudal rami. The number of spiniform armature elements on the P5 is reduced in both species, being one in
<taxonomicName id="2DD70CEDF772FFABFE18F8237E3AF864" authorityName="Sak &amp; Karaytuğ &amp; Huys" authorityYear="2024" baseAuthorityName="Bodiou &amp; Colomines" baseAuthorityYear="1986" box="[449,602,1929,1955]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angolensis" status="comb. nov.">
<emphasis id="D8A3AB7CF772FFABFE18F8237E3AF864" box="[449,602,1929,1955]" italics="true" pageId="20" pageNumber="21">N. angolensis</emphasis>
</taxonomicName>
<emphasis id="D8A3AB7CF772FFABFDB8F8207E82F864" bold="true" box="[609,738,1929,1955]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="C3901607F772FFABFDB8F8207E82F864" box="[609,738,1929,1955]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and none in
<taxonomicName id="2DD70CEDF772FFABFCAAF8237F9EF864" authorityName="Sak et al." authorityYear="2008" baseAuthorityName="Chappuis &amp; Rouch" baseAuthorityYear="1961" box="[883,1022,1929,1955]" class="Copepoda" family="Arenopontiidae" genus="Neoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="D8A3AB7CF772FFABFCAAF8237F9EF864" box="[883,1022,1929,1955]" italics="true" pageId="20" pageNumber="21">N. africanus</emphasis>
</taxonomicName>
. The species is known only from the
<typeStatus id="356CC9CCF772FFABFF4EF8077CA7F80F" box="[151,199,1966,1992]" pageId="20" pageNumber="21">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

140
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@ -1,73 +1,73 @@
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<subSubSection id="1EBF25F2E16AFF85FF4EF9A9FC51F9E4" box="[151,1001,1548,1575]" pageId="2" pageNumber="61" type="nomenclature"> <subSubSection id="1EBF25F2E16AFF85FF4EF9A9FC51F9E4" box="[151,1001,1548,1575]" pageId="2" pageNumber="61" type="nomenclature">
@ -231,12 +231,16 @@ det.
. .
<emphasis id="64D1AA6BE16AFF85FAECF8E0FDD1F8BF" bold="true" pageId="2" pageNumber="61"> <emphasis id="64D1AA6BE16AFF85FAECF8E0FDD1F8BF" bold="true" pageId="2" pageNumber="61">
syn. nov. syn. nov.
<materialsCitation id="E6CD7C24E16AFF85FF1EF8C1FB99F87A" collectingDate="1997-11" collectionCode="NMW, NHMW" collectorName="Br. v. W. &amp; Brunner v. W." country="Brazil" determinerName="det. Br. v. W. &amp; O. Zompro" location="Espirito-Santo" pageId="2" pageNumber="61" specimenCode="20.264" specimenCount="1" stateProvince="Espirito-Santo" typeStatus="holotype"> <materialsCitation id="E6CD7C24E16AFF85FF1EF8C1FB99F87A" collectingDate="1997-11-31" collectionCode="NMW, NHMW" collectorName="Br. v. W. &amp; Brunner v. W." country="Brazil" determinerName="det. Br. v. W. &amp; O. Zompro" location="Espirito-Santo" pageId="2" pageNumber="61" specimenCode="20.264" specimenCount="1" stateProvince="Espirito-Santo" typeStatus="holotype">
&gt;&gt; &gt;&gt;
<typeStatus id="891EC8DBE16AFF85FF32F8C1FEF7F8BF" box="[235,335,1892,1916]" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus> <typeStatus id="891EC8DBE16AFF85FF32F8C1FEF7F8BF" box="[235,335,1892,1916]" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus>
, ♀ ( ,
<specimenCount id="40A3BDF0E16AFF85FE83F8C0FED5F8BD" box="[346,365,1893,1918]" count="1" pageId="2" pageNumber="61" type="female">♀</specimenCount>
(
<collectionCode id="30B4EEBCE16AFF85FEA3F8C6FE03F8B8" box="[378,443,1891,1915]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NMW</collectionCode> <collectionCode id="30B4EEBCE16AFF85FEA3F8C6FE03F8B8" box="[378,443,1891,1915]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NMW</collectionCode>
), with two eggs: ), with
<specimenCount id="40A3BDF0E16AFF85FDDDF8C1FDDCF8BF" box="[516,612,1892,1916]" count="2" pageId="2" pageNumber="61" type="egg">two eggs</specimenCount>
:
<determinerName id="A3C69ABDE16AFF85FDA1F8C1FD48F8B8" box="[632,752,1892,1916]" pageId="2" pageNumber="61">det. Br.v.W.</determinerName> <determinerName id="A3C69ABDE16AFF85FDA1F8C1FD48F8B8" box="[632,752,1892,1916]" pageId="2" pageNumber="61">det. Br.v.W.</determinerName>
<taxonomicName id="91A50DFAE16AFF85FD2FF8C1FC6BF8B8" ID-CoL="48H3C" authority="Br" authorityName="Br" box="[758,979,1892,1916]" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra"> <taxonomicName id="91A50DFAE16AFF85FD2FF8C1FC6BF8B8" ID-CoL="48H3C" authority="Br" authorityName="Br" box="[758,979,1892,1916]" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="64D1AA6BE16AFF85FD2FF8C1FC09F8B8" box="[758,945,1892,1916]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis> <emphasis id="64D1AA6BE16AFF85FD2FF8C1FC09F8B8" box="[758,945,1892,1916]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis>
@ -257,18 +261,19 @@ Zompro
</taxonomicName> </taxonomicName>
, ,
<taxonomicNameLabel id="7FE21710E16AFF85FCC6F826FCEEF859" box="[799,854,1923,1946]" pageId="2" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel> <taxonomicNameLabel id="7FE21710E16AFF85FCC6F826FCEEF859" box="[799,854,1923,1946]" pageId="2" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="64D1AA6BE16AFF85FC85F821FCD7F85E" bold="true" box="[860,879,1924,1949]" pageId="2" pageNumber="61">♀</emphasis> <emphasis id="64D1AA6BE16AFF85FC85F821FCD7F85E" bold="true" box="[860,879,1924,1949]" pageId="2" pageNumber="61">
and 2 eggs, HOLOTYPUS, PLT of <specimenCount id="40A3BDF0E16AFF85FC85F821FCD7F85E" box="[860,879,1924,1949]" count="1" pageId="2" pageNumber="61" type="female">♀</specimenCount>
<taxonomicName id="91A50DFAE16AFF85FB38F827FEA2F87B" ID-CoL="48H3C" authority="Brunner" authorityName="Brunner" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra"> </emphasis>
and
<specimenCount id="40A3BDF0E16AFF85FC78F827FC5EF858" box="[929,998,1922,1947]" count="2" pageId="2" pageNumber="61" type="egg">2 eggs</specimenCount>
, HOLOTYPUS, PLT of
<taxonomicName id="91A50DFAE16AFF85FB38F827FA24F859" ID-CoL="48H3C" authority="Brunner" authorityName="Brunner" box="[1249,1436,1922,1946]" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="64D1AA6BE16AFF85FB38F827FA24F859" box="[1249,1436,1922,1946]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis> <emphasis id="64D1AA6BE16AFF85FB38F827FA24F859" box="[1249,1436,1922,1946]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis>
<collectorName id="FB5013AFE16AFF85FF1EF804FEE6F87B" box="[199,350,1953,1976]" pageId="2" pageNumber="61">Brunner v. W.</collectorName>
</taxonomicName> </taxonomicName>
<collectorName id="FB5013AFE16AFF85FF1EF804FEE6F87B" box="[199,350,1953,1976]" pageId="2" pageNumber="61">Brunner v. W.</collectorName>
, 1907, det. , 1907, det.
<determinerName id="A3C69ABDE16AFF85FE07F804FDF0F87B" box="[478,584,1953,1977]" pageId="2" pageNumber="61">O.Zompro</determinerName> <determinerName id="A3C69ABDE16AFF85FE07F804FDF0F87B" box="[478,584,1953,1977]" pageId="2" pageNumber="61">O.Zompro</determinerName>
<date id="221B50B9E16AFF85FD8BF804FD1EF87A" box="[594,678,1953,1977]" pageId="2" pageNumber="61" value="1997-11"> XI.1997 (
<collectingDate id="325FA951E16AFF85FD8BF804FD1EF87A" box="[594,678,1953,1977]" pageId="2" pageNumber="61" value="1997-11">XI.1997</collectingDate>
</date>
(
<collectionCode id="30B4EEBCE16AFF85FD6EF804FCB5F87A" box="[695,781,1953,1977]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NHMW</collectionCode> <collectionCode id="30B4EEBCE16AFF85FD6EF804FCB5F87A" box="[695,781,1953,1977]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NHMW</collectionCode>
) ( ) (
<figureCitation id="CE9E6AFCE16AFF85FCFCF804FC3CF87A" box="[805,900,1953,1977]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,899,923]" captionStartId-1="4.[151,250,1667,1691]" captionStartId-2="5.[151,250,1772,1796]" captionTargetBox-0="[159,1436,188,866]" captionTargetBox-1="[151,1436,202,1643]" captionTargetBox-2="[151,1436,181,1740]" captionTargetId-0="figure-19@3.[151,1436,181,875]" captionTargetId-1="figure-32@4.[151,1436,181,1643]" captionTargetId-2="figure-19@5.[151,1436,181,1748]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Exocnophila brevitarsata comb. nov., female holotype of Exocnophila exintegra syn. nov. A, body in dorsal view; B, habitus in dorsal view; C, habitus in lateral view (mirrored). Scale bars: 2 mm." captionText-1="FIGURE 2. Exocnophila brevitarsata comb. nov., head and thorax, female holotype of Exocnophila exintegra syn. nov. A, dorsal view; B, right lateral view; C, left lateral view; D, ventral view. Scale bars: 2 mm." captionText-2="FIGURE 3. Exocnophila brevitarsata comb. nov., thorax and abdomen, female holotype of Exocnophila exintegra syn. nov. A, metathorax in dorsal view; B, terminala (end of the abdomen) in dorsal view; C, terminalia, lateral view; D, terminalia, ventral view. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14024023" figureDoi-1="http://doi.org/10.5281/zenodo.14024027" figureDoi-2="http://doi.org/10.5281/zenodo.14024031" httpUri-0="https://zenodo.org/record/14024023/files/figure.png" httpUri-1="https://zenodo.org/record/14024027/files/figure.png" httpUri-2="https://zenodo.org/record/14024031/files/figure.png" pageId="2" pageNumber="61">Figs 1–3</figureCitation> <figureCitation id="CE9E6AFCE16AFF85FCFCF804FC3CF87A" box="[805,900,1953,1977]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,899,923]" captionStartId-1="4.[151,250,1667,1691]" captionStartId-2="5.[151,250,1772,1796]" captionTargetBox-0="[159,1436,188,866]" captionTargetBox-1="[151,1436,202,1643]" captionTargetBox-2="[151,1436,181,1740]" captionTargetId-0="figure-19@3.[151,1436,181,875]" captionTargetId-1="figure-32@4.[151,1436,181,1643]" captionTargetId-2="figure-19@5.[151,1436,181,1748]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Exocnophila brevitarsata comb. nov., female holotype of Exocnophila exintegra syn. nov. A, body in dorsal view; B, habitus in dorsal view; C, habitus in lateral view (mirrored). Scale bars: 2 mm." captionText-1="FIGURE 2. Exocnophila brevitarsata comb. nov., head and thorax, female holotype of Exocnophila exintegra syn. nov. A, dorsal view; B, right lateral view; C, left lateral view; D, ventral view. Scale bars: 2 mm." captionText-2="FIGURE 3. Exocnophila brevitarsata comb. nov., thorax and abdomen, female holotype of Exocnophila exintegra syn. nov. A, metathorax in dorsal view; B, terminala (end of the abdomen) in dorsal view; C, terminalia, lateral view; D, terminalia, ventral view. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14024023" figureDoi-1="http://doi.org/10.5281/zenodo.14024027" figureDoi-2="http://doi.org/10.5281/zenodo.14024031" httpUri-0="https://zenodo.org/record/14024023/files/figure.png" httpUri-1="https://zenodo.org/record/14024027/files/figure.png" httpUri-2="https://zenodo.org/record/14024031/files/figure.png" pageId="2" pageNumber="61">Figs 1–3</figureCitation>
@ -277,7 +282,7 @@ and 2 eggs, HOLOTYPUS, PLT of
). ).
</materialsCitation> </materialsCitation>
</emphasis> </emphasis>
<materialsCitation id="E6CD7C24E16AFF85FBF3F804FBB6F835" collectingDate="1997-11" collectionCode="NMW, NHMW" collectorName="Br. v. W. &amp; Brunner v. W." determinerName="Br. v. W &amp; O. Zompro" pageId="2" pageNumber="61" specimenCode="14.978" specimenCount="1" specimenCount-female="1" typeStatus="paratype"> <materialsCitation id="E6CD7C24E16AFF85FBF3F804FBB6F835" collectingDate="1997-11-31" collectionCode="NMW, NHMW" collectorName="Br. v. W. &amp; Brunner v. W." determinerName="Br. v. W &amp; O. Zompro" pageId="2" pageNumber="61" specimenCode="14.978" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="64D1AA6BE16AFF85FBF3F804FAB6F87A" bold="true" box="[1066,1294,1953,1979]" pageId="2" pageNumber="61"> <emphasis id="64D1AA6BE16AFF85FBF3F804FAB6F87A" bold="true" box="[1066,1294,1953,1979]" pageId="2" pageNumber="61">
<typeStatus id="891EC8DBE16AFF85FBF3F804FB36F87A" box="[1066,1166,1953,1977]" pageId="2" pageNumber="61" type="paratype">Paratype</typeStatus> <typeStatus id="891EC8DBE16AFF85FBF3F804FB36F87A" box="[1066,1166,1953,1977]" pageId="2" pageNumber="61" type="paratype">Paratype</typeStatus>
, ,
@ -306,16 +311,13 @@ Zompro
<taxonomicNameLabel id="7FE21710E16AFF85FB76F865FB5CF814" box="[1199,1252,1984,2007]" pageId="2" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel> <taxonomicNameLabel id="7FE21710E16AFF85FB76F865FB5CF814" box="[1199,1252,1984,2007]" pageId="2" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="64D1AA6BE16AFF85FB30F864FB44F819" bold="true" box="[1257,1276,1985,2010]" pageId="2" pageNumber="61">♀</emphasis> <emphasis id="64D1AA6BE16AFF85FB30F864FB44F819" bold="true" box="[1257,1276,1985,2010]" pageId="2" pageNumber="61">♀</emphasis>
, PARATYPUS, PLT of , PARATYPUS, PLT of
<taxonomicName id="91A50DFAE16AFF85FECAF87BFD9FF835" ID-CoL="48H3C" authority="Brunner" authorityName="Brunner" box="[275,551,2014,2038]" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra"> <taxonomicName id="91A50DFAE16AFF85FECAF87BFE75F835" ID-CoL="48H3C" authority="Brunner" authorityName="Brunner" box="[275,461,2014,2038]" class="Insecta" family="Diapheromeridae" genus="Ocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="64D1AA6BE16AFF85FECAF87BFE75F835" box="[275,461,2014,2038]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis> <emphasis id="64D1AA6BE16AFF85FECAF87BFE75F835" box="[275,461,2014,2038]" italics="true" pageId="2" pageNumber="61">Ocnophila integra</emphasis>
<collectorName id="FB5013AFE16AFF85FE0DF87AFDDCF835" box="[468,612,2015,2038]" pageId="2" pageNumber="61">Brunner v. W.</collectorName>
</taxonomicName> </taxonomicName>
<collectorName id="FB5013AFE16AFF85FE0DF87AFDDCF835" box="[468,612,2015,2038]" pageId="2" pageNumber="61">Brunner v. W.</collectorName>
, 1907, det. , 1907, det.
<determinerName id="A3C69ABDE16AFF85FD01F87BFCFAF835" box="[728,834,2014,2038]" pageId="2" pageNumber="61">O.Zompro</determinerName> <determinerName id="A3C69ABDE16AFF85FD01F87BFCFAF835" box="[728,834,2014,2038]" pageId="2" pageNumber="61">O.Zompro</determinerName>
<date id="221B50B9E16AFF85FC90F87BFC25F835" box="[841,925,2014,2038]" pageId="2" pageNumber="61" value="1997-11"> XI.1997 (
<collectingDate id="325FA951E16AFF85FC90F87BFC25F835" box="[841,925,2014,2038]" pageId="2" pageNumber="61" value="1997-11">XI.1997</collectingDate>
</date>
(
<collectionCode id="30B4EEBCE16AFF85FC73F87BFBBBF835" box="[938,1027,2014,2038]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NHMW</collectionCode> <collectionCode id="30B4EEBCE16AFF85FC73F87BFBBBF835" box="[938,1027,2014,2038]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="61">NHMW</collectionCode>
). ).
</materialsCitation> </materialsCitation>

429
data/DE/0C/C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml
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@ -1,75 +1,75 @@
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<materialsCitation id="E6CD7C24E17FFF90FF4EF8FAFA24F85E" collectingDate="2017-01" collectionCode="MZUSP" collectorName="VM Ghirotto &amp; PABA Neves &amp; PW Engelking &amp; PIC Machado" country="Brazil" latitude="-22.43547" location="Rio de Janeiro" longLatPrecision="1" longitude="-44.612858" pageId="23" pageNumber="82" specimenCode="V0044, V0043" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<emphasis id="64D1AA6BE17FFF90FF4EF8FAFE03F8BB" bold="true" box="[151,443,1886,1916]" pageId="23" pageNumber="82"> <emphasis id="64D1AA6BE17FFF90FF4EF8FAFE03F8BB" bold="true" box="[151,443,1886,1916]" pageId="23" pageNumber="82">
<typeStatus id="891EC8DBE17FFF90FF4EF8FAFEA8F8BA" box="[151,272,1887,1913]" pageId="23" pageNumber="82" type="paratype">Paratypes</typeStatus> <typeStatus id="891EC8DBE17FFF90FF4EF8FAFEA8F8BA" box="[151,272,1887,1913]" pageId="23" pageNumber="82" type="paratype">Paratypes</typeStatus>
, ,
<specimenCount id="40A3BDF0E17FFF90FEC6F8C5FE8CF8BF" box="[287,308,1888,1916]" count="1" pageId="23" pageNumber="82" type="female">♀</specimenCount> <specimenCount id="40A3BDF0E17FFF90FEC6F8C5FE8CF8BF" box="[287,308,1888,1916]" count="1" pageId="23" pageNumber="82" type="female">♀</specimenCount>
(MZUSP): (
<collectionCode id="30B4EEBCE17FFF90FE9CF8FBFE16F8BB" box="[325,430,1886,1912]" pageId="23" pageNumber="82">MZUSP</collectionCode>
):
</emphasis> </emphasis>
<specimenCount id="40A3BDF0E17FFF90FE1DF8FAFE5FF8BA" box="[452,487,1887,1913]" count="1" pageId="23" pageNumber="82" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E17FFF90FE1DF8FAFE5FF8BA" box="[452,487,1887,1913]" count="1" pageId="23" pageNumber="82" type="male">1♂</specimenCount>
(MZUSP V0044), (
<collectionCode id="30B4EEBCE17FFF90FE2EF8FAFDEDF8BA" box="[503,597,1887,1913]" pageId="23" pageNumber="82">MZUSP</collectionCode>
<specimenCode id="0603DE02E17FFF90FD87F8FAFD17F8BA" box="[606,687,1887,1913]" pageId="23" pageNumber="82">V0044</specimenCode>
),
<specimenCount id="40A3BDF0E17FFF90FD1DF8FAFD5EF8BA" box="[708,742,1887,1913]" count="1" pageId="23" pageNumber="82" type="female">1♀</specimenCount> <specimenCount id="40A3BDF0E17FFF90FD1DF8FAFD5EF8BA" box="[708,742,1887,1913]" count="1" pageId="23" pageNumber="82" type="female">1♀</specimenCount>
(MZUSP V0043), eggs: (
<collectionCode id="30B4EEBCE17FFF90FD2EF8FAFCEDF8BA" box="[759,853,1887,1913]" pageId="23" pageNumber="82">MZUSP</collectionCode>
<specimenCode id="0603DE02E17FFF90FC87F8FAFC16F8BA" box="[862,942,1887,1913]" pageId="23" pageNumber="82">V0043</specimenCode>
), eggs:
<collectingCountry id="2EB236E9E17FFF90FBDEF8FAFBF5F8BA" box="[1031,1101,1887,1913]" name="Brazil" pageId="23" pageNumber="82">Brasil</collectingCountry> <collectingCountry id="2EB236E9E17FFF90FBDEF8FAFBF5F8BA" box="[1031,1101,1887,1913]" name="Brazil" pageId="23" pageNumber="82">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE17FFF90FB83F8FAFABCF8BA" box="[1114,1284,1887,1913]" country="Brazil" name="Rio de Janeiro" pageId="23" pageNumber="82">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE17FFF90FB83F8FAFABCF8BA" box="[1114,1284,1887,1913]" country="Brazil" name="Rio de Janeiro" pageId="23" pageNumber="82">Rio de Janeiro</collectingRegion>
@ -143,116 +155,313 @@ col.
<geoCoordinate id="339110BEE17FFF90FEC3F827FE0BF85F" box="[282,435,1922,1948]" degrees="22" direction="south" minutes="26" orientation="latitude" pageId="23" pageNumber="82" precision="1" seconds="07.7" value="-22.43547">22°26’07.7”S</geoCoordinate> <geoCoordinate id="339110BEE17FFF90FEC3F827FE0BF85F" box="[282,435,1922,1948]" degrees="22" direction="south" minutes="26" orientation="latitude" pageId="23" pageNumber="82" precision="1" seconds="07.7" value="-22.43547">22°26’07.7”S</geoCoordinate>
<geoCoordinate id="339110BEE17FFF90FE64F827FDDAF85F" box="[445,610,1922,1948]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="23" pageNumber="82" precision="1" seconds="46.3" value="-44.612858">44°36’46.3”W</geoCoordinate> <geoCoordinate id="339110BEE17FFF90FE64F827FDDAF85F" box="[445,610,1922,1948]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="23" pageNumber="82" precision="1" seconds="46.3" value="-44.612858">44°36’46.3”W</geoCoordinate>
. .
<date id="221B50B9E17FFF90FDB6F826FD01F85E" box="[623,697,1923,1949]" pageId="23" pageNumber="82" value="2017-01">i.2017</date> <date id="221B50B9E17FFF90FDB6F826FD01F85E" box="[623,697,1923,1949]" pageId="23" pageNumber="82" value="2017-01">
. VM Ghirotto, PABA Neves, PW Engelking, PIC Machado col.; <collectingDate id="325FA951E17FFF90FDB6F826FD01F85E" box="[623,697,1923,1949]" pageId="23" pageNumber="82" value="2017-01">i.2017</collectingDate>
</date>
.
<collectorName id="FB5013AFE17FFF90FD1CF826FCE5F85E" box="[709,861,1923,1949]" pageId="23" pageNumber="82">VM Ghirotto</collectorName>
,
<collectorName id="FB5013AFE17FFF90FCB3F826FBBBF85E" box="[874,1027,1923,1949]" pageId="23" pageNumber="82">PABA Neves</collectorName>
,
<collectorName id="FB5013AFE17FFF90FBC9F826FB00F85E" box="[1040,1208,1923,1949]" pageId="23" pageNumber="82">PW Engelking</collectorName>
,
<collectorName id="FB5013AFE17FFF90FB1CF826FADAF85E" box="[1221,1378,1923,1949]" pageId="23" pageNumber="82">PIC Machado</collectorName>
col.;
</materialsCitation>
<materialsCitation id="E6CD7C24E17FFF9FFF4EF802FDCDFF72" collectedFrom="Synedrella nodiflora" collectingDate="2019-05-05" collectionCode="MZUSP" collectorName="VM Ghirotto &amp; RP Indicatti &amp; EF Trova" country="Brazil" elevation="750" lastPageId="24" lastPageNumber="83" latitude="-22.452917" location="Itatiaia" longLatPrecision="1" longitude="-44.60936" municipality="Itatiaia" pageId="23" pageNumber="82" specimenCode="V0292, V0293, V0309" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E17FFF90FF4EF802FF02F802" box="[151,186,1959,1985]" count="2" pageId="23" pageNumber="82" type="male">2♂</specimenCount> <specimenCount id="40A3BDF0E17FFF90FF4EF802FF02F802" box="[151,186,1959,1985]" count="2" pageId="23" pageNumber="82" type="male">2♂</specimenCount>
(MZUSP V0292, V0293), (
<collectionCode id="30B4EEBCE17FFF90FF10F802FE9FF802" box="[201,295,1959,1985]" pageId="23" pageNumber="82">MZUSP</collectionCode>
<specimenCode id="0603DE02E17FFF90FEF7F802FEC6F802" box="[302,382,1959,1985]" pageId="23" pageNumber="82">V0292</specimenCode>
,
<specimenCode id="0603DE02E17FFF90FE51F802FE60F802" box="[392,472,1959,1985]" pageId="23" pageNumber="82">V0293</specimenCode>
),
<specimenCount id="40A3BDF0E17FFF90FE35F802FDB7F802" box="[492,527,1959,1985]" count="1" pageId="23" pageNumber="82" type="female">1♀</specimenCount> <specimenCount id="40A3BDF0E17FFF90FE35F802FDB7F802" box="[492,527,1959,1985]" count="1" pageId="23" pageNumber="82" type="female">1♀</specimenCount>
(MZUSP V0309), eggs: (
<collectionCode id="30B4EEBCE17FFF90FDC7F802FDC4F802" box="[542,636,1959,1985]" pageId="23" pageNumber="82">MZUSP</collectionCode>
<specimenCode id="0603DE02E17FFF90FD5AF802FD6BF802" box="[643,723,1959,1985]" pageId="23" pageNumber="82">V0309</specimenCode>
), eggs:
<collectingCountry id="2EB236E9E17FFF90FCF3F802FCC8F802" box="[810,880,1959,1985]" name="Brazil" pageId="23" pageNumber="82">Brasil</collectingCountry> <collectingCountry id="2EB236E9E17FFF90FCF3F802FCC8F802" box="[810,880,1959,1985]" name="Brazil" pageId="23" pageNumber="82">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE17FFF90FCA2F802FB9AF802" box="[891,1058,1959,1985]" country="Brazil" name="Rio de Janeiro" pageId="23" pageNumber="82">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE17FFF90FCA2F802FB9AF802" box="[891,1058,1959,1985]" country="Brazil" name="Rio de Janeiro" pageId="23" pageNumber="82">Rio de Janeiro</collectingRegion>
. Itatiaia. PARNA Itatiaia, próx ao alojamento dos pesquisadores, .
<quantity id="915DDB9CE17FFF90FE3EF86FFD92F826" box="[487,554,1994,2021]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="82" unit="m" value="750.0">750m</quantity> <collectingMunicipality id="B67EEC03E17FFF90FBF5F802FBC5F802" box="[1068,1149,1959,1985]" pageId="23" pageNumber="82">Itatiaia</collectingMunicipality>
, . PARNA Itatiaia,
<locationDeviation id="47A86F5BE17FFF90FA9EF802FE58F826" pageId="23" pageNumber="82">próx ao alojamento dos pesquisadores</locationDeviation>
,750m,
<geoCoordinate id="339110BEE17FFF90FDE8F86FFD73F826" box="[561,715,1994,2021]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="23" pageNumber="82" precision="1" seconds="10.5" value="-22.452917">22°27’10.5”S</geoCoordinate> <geoCoordinate id="339110BEE17FFF90FDE8F86FFD73F826" box="[561,715,1994,2021]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="23" pageNumber="82" precision="1" seconds="10.5" value="-22.452917">22°27’10.5”S</geoCoordinate>
<geoCoordinate id="339110BEE17FFF90FD17F86FFCCAF827" box="[718,882,1994,2020]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="23" pageNumber="82" precision="1" seconds="33.7" value="-44.60936">44°36’33.7”W</geoCoordinate> <geoCoordinate id="339110BEE17FFF90FD17F86FFCCAF827" box="[718,882,1994,2020]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="23" pageNumber="82" precision="1" seconds="33.7" value="-44.60936">44°36’33.7”W</geoCoordinate>
,on cf. ,on cf.
<taxonomicName id="91A50DFAE17FFF90FC61F86EFB22F826" box="[952,1178,1995,2021]" class="Magnoliopsida" family="Asteraceae" genus="Synedrella" kingdom="Plantae" order="Asterales" pageId="23" pageNumber="82" phylum="Tracheophyta" rank="species" species="nodiflora"> <taxonomicName id="91A50DFAE17FFF90FC61F86EFB22F826" box="[952,1178,1995,2021]" class="Magnoliopsida" family="Asteraceae" genus="Synedrella" kingdom="Plantae" order="Asterales" pageId="23" pageNumber="82" phylum="Tracheophyta" rank="species" species="nodiflora">
<emphasis id="64D1AA6BE17FFF90FC61F86EFB22F826" box="[952,1178,1995,2021]" italics="true" pageId="23" pageNumber="82">Synedrella nodiflora</emphasis> <emphasis id="64D1AA6BE17FFF90FC61F86EFB22F826" box="[952,1178,1995,2021]" italics="true" pageId="23" pageNumber="82">
<collectedFrom id="E5B93584E17FFF90FC61F86EFB22F826" box="[952,1178,1995,2021]" pageId="23" pageNumber="82">Synedrella nodiflora</collectedFrom>
</emphasis>
</taxonomicName> </taxonomicName>
, ,
<taxonomicName id="91A50DFAE17FFF90FB7BF86EFA99F826" box="[1186,1313,1995,2021]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="23" pageNumber="82" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName> <taxonomicName id="91A50DFAE17FFF90FB7BF86EFA99F826" box="[1186,1313,1995,2021]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="23" pageNumber="82" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
. .
<date id="221B50B9E17FFF90FAFEF86FFA20F826" box="[1319,1432,1994,2021]" pageId="23" pageNumber="82" value="2019-05-05">05.v.2019</date> <date id="221B50B9E17FFF90FAFEF86FFA20F826" box="[1319,1432,1994,2021]" pageId="23" pageNumber="82" value="2019-05-05">
. VM Ghirotto, RP Indicatti, EF Trova col.; <collectingDate id="325FA951E17FFF90FAFEF86FFA20F826" box="[1319,1432,1994,2021]" pageId="23" pageNumber="82" value="2019-05-05">05.v.2019</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFF4EFF32FE97FF72" box="[151,303,151,177]" pageId="24" pageNumber="83">VM Ghirotto</collectorName>
,
<collectorName id="FB5013AFE170FF9FFEE2FF32FE7CFF72" box="[315,452,151,177]" pageId="24" pageNumber="83">RP Indicatti</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE08FF32FD84FF72" box="[465,572,151,177]" pageId="24" pageNumber="83">EF Trova</collectorName>
col.;
</materialsCitation>
<materialsCitation id="E6CD7C24E170FF9FFDA4FF32FE9FFF3A" collectingDate="2019-05-07" collectionCode="MZUSP" collectorName="VM Ghirotto &amp; RP Indicatti" country="Brazil" elevation="750" latitude="-22.452917" location="PARNA Itatiaia" longLatPrecision="1" longitude="-44.60936" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCode="V0400" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFDA4FF32FD18FF72" box="[637,672,151,177]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFDA4FF32FD18FF72" box="[637,672,151,177]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount>
juvenile (MZUSP V0400): <specimenCount id="40A3BDF0E170FF9FFD70FF32FCBBFF72" box="[681,771,151,177]" pageId="24" pageNumber="83" type="juvenile">juvenile</specimenCount>
(
<collectionCode id="30B4EEBCE170FF9FFCCBFF32FCC8FF72" box="[786,880,151,177]" pageId="24" pageNumber="83">MZUSP</collectionCode>
<specimenCode id="0603DE02E170FF9FFCA0FF32FC73FF72" box="[889,971,151,177]" pageId="24" pageNumber="83">V0400</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFC07FF32FB9CFF72" box="[990,1060,151,177]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFC07FF32FB9CFF72" box="[990,1060,151,177]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE170FF9FFBE9FF32FB61FF72" box="[1072,1241,151,177]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFBE9FF32FB61FF72" box="[1072,1241,151,177]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
. Itatiaia. PARNA Itatiaia, próx ao alojamento dos pesquisadores, .
<quantity id="915DDB9CE170FF9FFD6FFF1FFD40FF16" box="[694,760,186,213]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="24" pageNumber="83" unit="m" value="750.0">750m</quantity> <collectingMunicipality id="B67EEC03E170FF9FFB3DFF32FA8DFF72" box="[1252,1333,151,177]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
.
<location id="537A20A2E170FF9FFA98FF32FF50FF16" LSID="urn:lsid:plazi:treatment:DE0CC76FE17FFFA2FF4EF934FAB0FE6E:537A20A2E170FF9FFA98FF32FF50FF16" country="Brazil" latitude="-22.452917" longLatPrecision="1" longitude="-44.60936" municipality="Itatiaia" name="PARNA Itatiaia" pageId="24" pageNumber="83" stateProvince="Rio de Janeiro">PARNA Itatiaia</location>
, ,
<locationDeviation id="47A86F5BE170FF9FFF2CFF1EFD11FF16" box="[245,681,187,213]" pageId="24" pageNumber="83">próx ao alojamento dos pesquisadores</locationDeviation>
, 750m,
<geoCoordinate id="339110BEE170FF9FFCDCFF1FFC27FF16" box="[773,927,186,213]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="24" pageNumber="83" precision="1" seconds="10.5" value="-22.452917">22°27’10.5”S</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFCDCFF1FFC27FF16" box="[773,927,186,213]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="24" pageNumber="83" precision="1" seconds="10.5" value="-22.452917">22°27’10.5”S</geoCoordinate>
<geoCoordinate id="339110BEE170FF9FFC7EFF1EFBF4FF16" box="[935,1100,187,213]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="24" pageNumber="83" precision="1" seconds="33.7" value="-44.60936">44°36’33.7”W</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFC7EFF1EFBF4FF16" box="[935,1100,187,213]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="24" pageNumber="83" precision="1" seconds="33.7" value="-44.60936">44°36’33.7”W</geoCoordinate>
. .
<date id="221B50B9E170FF9FFB80FF1EFB73FF16" box="[1113,1227,187,213]" pageId="24" pageNumber="83" value="2019-05-07">07.v.2019</date> <date id="221B50B9E170FF9FFB80FF1EFB73FF16" box="[1113,1227,187,213]" pageId="24" pageNumber="83" value="2019-05-07">
. VM Ghirotto, RP Indicatti col.; <collectingDate id="325FA951E170FF9FFB80FF1EFB73FF16" box="[1113,1227,187,213]" pageId="24" pageNumber="83" value="2019-05-07">07.v.2019</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFB0FFF1EFAD6FF16" box="[1238,1390,187,213]" pageId="24" pageNumber="83">VM Ghirotto</collectorName>
,
<collectorName id="FB5013AFE170FF9FFAA2FF1EFF4CFF3A" pageId="24" pageNumber="83">RP Indicatti</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFEE8FF7AFACBFF3A" box="[305,1395,223,249]" collectingDate="1972-01" collectionCode="MZUSP" collectorName="Dirigns" country="Brazil" location="Itatiaia" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCode="MZUSP0262" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFEE8FF7AFEECFF3A" box="[305,340,223,249]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFEE8FF7AFEECFF3A" box="[305,340,223,249]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount>
(MZUSP0262): (
<specimenCode id="0603DE02E170FF9FFEBBFF7AFE47FF3A" box="[354,511,223,249]" pageId="24" pageNumber="83">MZUSP0262</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFDD6FF7AFDEDFF3A" box="[527,597,223,249]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFDD6FF7AFDEDFF3A" box="[527,597,223,249]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. Est do Rio [ . Est do Rio [
<collectingRegion id="9461B89BE170FF9FFD06FF7AFC3FFF3A" box="[735,903,223,249]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFD06FF7AFC3FFF3A" box="[735,903,223,249]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
]. Itatiaia (1.100 mtr.). ].
<date id="221B50B9E170FF9FFBA2FF7AFB7DFF3A" box="[1147,1221,223,249]" pageId="24" pageNumber="83" value="1972-01">i.1972</date> <collectingMunicipality id="B67EEC03E170FF9FFC4EFF7AFC5DFF3A" box="[919,997,223,249]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
. Dirigns [col.].; (1.100 mtr.).
<date id="221B50B9E170FF9FFBA2FF7AFB7DFF3A" box="[1147,1221,223,249]" pageId="24" pageNumber="83" value="1972-01">
<collectingDate id="325FA951E170FF9FFBA2FF7AFB7DFF3A" box="[1147,1221,223,249]" pageId="24" pageNumber="83" value="1972-01">i.1972</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFB16FF7AFA9BFF3A" box="[1231,1315,223,249]" pageId="24" pageNumber="83">Dirigns</collectorName>
[col.].;
</materialsCitation>
<materialsCitation id="E6CD7C24E170FF9FFAA0FF7AFB01FEDE" collectingDate="1960-04" collectionCode="MZUSP" collectorName="Dirigns" country="Brazil" location="Itatiaia" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCode="MZUSP0282" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFAA0FF7AFA24FF3A" box="[1401,1436,223,249]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFAA0FF7AFA24FF3A" box="[1401,1436,223,249]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount>
(MZUSP0282): (
<specimenCode id="0603DE02E170FF9FFF46FEA6FE84FEDE" box="[159,316,259,285]" pageId="24" pageNumber="83">MZUSP0282</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFE94FEA6FE2BFEDE" box="[333,403,259,285]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFE94FEA6FE2BFEDE" box="[333,403,259,285]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. Est do Rio [ . Est do Rio [
<collectingRegion id="9461B89BE170FF9FFDC4FEA6FD7EFEDE" box="[541,710,259,285]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFDC4FEA6FD7EFEDE" box="[541,710,259,285]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
]. Itatiaia (1.100 mtr.). ].
<date id="221B50B9E170FF9FFC63FEA6FBA9FEDE" box="[954,1041,259,285]" pageId="24" pageNumber="83" value="1960-04">iv.1960</date> <collectingMunicipality id="B67EEC03E170FF9FFD0FFEA6FC9CFEDE" box="[726,804,259,285]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
. Dirigns [col.].; (1.100 mtr.).
<date id="221B50B9E170FF9FFC63FEA6FBA9FEDE" box="[954,1041,259,285]" pageId="24" pageNumber="83" value="1960-04">
<collectingDate id="325FA951E170FF9FFC63FEA6FBA9FEDE" box="[954,1041,259,285]" pageId="24" pageNumber="83" value="1960-04">iv.1960</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFBC3FEA6FBD6FEDE" box="[1050,1134,259,285]" pageId="24" pageNumber="83">Dirigns</collectorName>
[col.].
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFB1DFEA6FB89FEA6" collectingDate="2016-10-31" collectingDateMax="2016-11-02" collectingDateMin="2016-10-31" collectionCode="MZUSP" collectorName="P. I. Chiquetto-Machado" country="Brazil" latitude="-22.426388" location="Parque Nacional de Itatiaia" longLatPrecision="21" longitude="-44.618053" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCode="MZUSP0685, MZUSP0686, MZUSP0738, 0783" specimenCount="5" specimenCount-female="2" specimenCount-male="2" specimenCount-subadult="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFB1DFEA6FB5FFEDE" box="[1220,1255,259,285]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFB1DFEA6FB5FFEDE" box="[1220,1255,259,285]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount>
(MZUSP0685), (
<specimenCode id="0603DE02E170FF9FFB2CFEA7FA28FEDE" box="[1269,1424,258,285]" pageId="24" pageNumber="83">MZUSP0685</specimenCode>
),
<specimenCount id="40A3BDF0E170FF9FFF4EFE82FEB3FE82" box="[151,267,295,321]" count="1" pageId="24" pageNumber="83" type="subadult">1 subadult</specimenCount> <specimenCount id="40A3BDF0E170FF9FFF4EFE82FEB3FE82" box="[151,267,295,321]" count="1" pageId="24" pageNumber="83" type="subadult">1 subadult</specimenCount>
<specimenCount id="40A3BDF0E170FF9FFECBFE82FE9FFE82" box="[274,295,295,321]" count="1" pageId="24" pageNumber="83" type="male">♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFECBFE82FE9FFE82" box="[274,295,295,321]" count="1" pageId="24" pageNumber="83" type="male">♂</specimenCount>
(MZUSP0686), (
<specimenCode id="0603DE02E170FF9FFEECFE82FE69FE82" box="[309,465,295,321]" pageId="24" pageNumber="83">MZUSP0686</specimenCode>
),
<specimenCount id="40A3BDF0E170FF9FFE3DFE82FDBEFE82" box="[484,518,295,321]" count="2" pageId="24" pageNumber="83" type="female">2♀</specimenCount> <specimenCount id="40A3BDF0E170FF9FFE3DFE82FDBEFE82" box="[484,518,295,321]" count="2" pageId="24" pageNumber="83" type="female">2♀</specimenCount>
(MZUSP0738, 0783): (
<specimenCode id="0603DE02E170FF9FFDCCFE82FD17FE82" box="[533,687,295,321]" pageId="24" pageNumber="83">MZUSP0738</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFD63FE82FD4FFE82" box="[698,759,295,321]" pageId="24" pageNumber="83">0783</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFCD3FE82FCE8FE82" box="[778,848,295,321]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFCD3FE82FCE8FE82" box="[778,848,295,321]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE170FF9FFC82FE82FBB8FE82" box="[859,1024,295,321]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFC82FE82FBB8FE82" box="[859,1024,295,321]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
. Itatiaia. Parque Nacional de Itatiaia, parte baixa. .
<collectingMunicipality id="B67EEC03E170FF9FFBD3FE82FBE3FE82" box="[1034,1115,295,321]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
.
<location id="537A20A2E170FF9FFBBFFE82FA20FE82" LSID="urn:lsid:plazi:treatment:DE0CC76FE17FFFA2FF4EF934FAB0FE6E:537A20A2E170FF9FFBBFFE82FA20FE82" box="[1126,1432,295,321]" country="Brazil" latitude="-22.426388" longLatPrecision="21" longitude="-44.618053" municipality="Itatiaia" name="Parque Nacional de Itatiaia" pageId="24" pageNumber="83" stateProvince="Rio de Janeiro">Parque Nacional de Itatiaia</location>
, parte baixa.
<geoCoordinate id="339110BEE170FF9FFEF9FEEFFE1CFEA6" box="[288,420,330,357]" degrees="22" direction="south" minutes="25" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="35" value="-22.426388">22°25’35”S</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFEF9FEEFFE1CFEA6" box="[288,420,330,357]" degrees="22" direction="south" minutes="25" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="35" value="-22.426388">22°25’35”S</geoCoordinate>
<geoCoordinate id="339110BEE170FF9FFE73FEEFFD83FEA6" box="[426,571,330,357]" degrees="44" direction="west" minutes="37" orientation="longitude" pageId="24" pageNumber="83" precision="15" seconds="05" value="-44.618053">44°37’05”W</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFE73FEEFFD83FEA6" box="[426,571,330,357]" degrees="44" direction="west" minutes="37" orientation="longitude" pageId="24" pageNumber="83" precision="15" seconds="05" value="-44.618053">44°37’05”W</geoCoordinate>
. .
<date id="221B50B9E170FF9FFD9CFEEEFD49FEA6" box="[581,753,331,357]" pageId="24" pageNumber="83" value="2016-10-31" valueMax="2016-11-02" valueMin="2016-10-31">31.x–2.xi.2016</date> <date id="221B50B9E170FF9FFD9CFEEEFD49FEA6" box="[581,753,331,357]" pageId="24" pageNumber="83" value="2016-10-31" valueMax="2016-11-02" valueMin="2016-10-31">
. P.I.Chiquetto-Machado col.; <collectingDate id="325FA951E170FF9FFD9CFEEEFD49FEA6" box="[581,753,331,357]" pageId="24" pageNumber="83" value="2016-10-31" valueMax="2016-11-02" valueMin="2016-10-31">31.x–2.xi.2016</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFD22FEEEFC46FEA6" box="[763,1022,331,357]" pageId="24" pageNumber="83">P.I.Chiquetto-Machado</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFBE5FEEEFDC2FE6E" collectingDate="2020-02-06" collectingDateMax="2020-02-08" collectingDateMin="2020-02-06" collectorName="P. Souza-Dias &amp; Siqueira &amp; Lima &amp; Borille" country="Brazil" location="PN Itatiaia" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCode="MNRJ-ENT15-17, MNRJ-ENT15-18" specimenCount="2" specimenCount-male="2" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFBE5FEEEFBE7FEA6" box="[1084,1119,331,357]" count="2" pageId="24" pageNumber="83" type="male">2♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFBE5FEEEFBE7FEA6" box="[1084,1119,331,357]" count="2" pageId="24" pageNumber="83" type="male">2♂</specimenCount>
(MNRJ-ENT15-17, MNRJ-ENT15-18): (
<specimenCode id="0603DE02E170FF9FFBB7FEEFFA86FEA6" box="[1134,1342,330,357]" pageId="24" pageNumber="83">MNRJ-ENT15-17</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFA91FEEEFEABFE4A" pageId="24" pageNumber="83">MNRJ-ENT15-18</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFEFFFECAFED4FE4A" box="[294,364,367,393]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFEFFFECAFED4FE4A" box="[294,364,367,393]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE170FF9FFEAFFECAFDA1FE4A" box="[374,537,367,393]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFEAFFECAFDA1FE4A" box="[374,537,367,393]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
. Itatiaia. PN Itatiaia, parte baixa, ao redor da Casa do Pesquisador. .
<date id="221B50B9E170FF9FFB23FECAFA20FE4A" box="[1274,1432,367,393]" pageId="24" pageNumber="83" value="2020-02-06" valueMax="2020-02-08" valueMin="2020-02-06">06–08.ii.2020</date> <collectingMunicipality id="B67EEC03E170FF9FFDFBFECAFDCBFE4A" box="[546,627,367,393]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
. P. Souza-Dias, Siqueira, Lima, Borille col.; .
<location id="537A20A2E170FF9FFDA5FECAFD4FFE4A" LSID="urn:lsid:plazi:treatment:DE0CC76FE17FFFA2FF4EF934FAB0FE6E:537A20A2E170FF9FFDA5FECAFD4FFE4A" box="[636,759,367,393]" country="Brazil" municipality="Itatiaia" name="PN Itatiaia" pageId="24" pageNumber="83" stateProvince="Rio de Janeiro">PN Itatiaia</location>
,
<locationDeviation id="47A86F5BE170FF9FFCD9FECAFB49FE4A" box="[768,1265,367,393]" pageId="24" pageNumber="83">parte baixa, ao redor da Casa do Pesquisador</locationDeviation>
.
<date id="221B50B9E170FF9FFB23FECAFA20FE4A" box="[1274,1432,367,393]" pageId="24" pageNumber="83" value="2020-02-06" valueMax="2020-02-08" valueMin="2020-02-06">
<collectingDate id="325FA951E170FF9FFB23FECAFA20FE4A" box="[1274,1432,367,393]" pageId="24" pageNumber="83" value="2020-02-06" valueMax="2020-02-08" valueMin="2020-02-06">06–08.ii.2020</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFF4EFE36FE8EFE6E" box="[151,310,403,429]" pageId="24" pageNumber="83">P. Souza-Dias</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE9BFE36FE1AFE6E" box="[322,418,403,429]" pageId="24" pageNumber="83">Siqueira</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE76FE36FE55FE6E" box="[431,493,403,429]" pageId="24" pageNumber="83">Lima</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE20FE36FDFFFE6E" box="[505,583,403,429]" pageId="24" pageNumber="83">Borille</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFD5EFE36FE8AFE12" collectingDate="1944-08-10" collectionCode="FIOC" collectorName="J. F. Zikan" country="Brazil" location="Itatiaia" municipality="Itatiaia" pageId="24" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFD5EFE36FD12FE6E" box="[647,682,403,429]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFD5EFE36FD12FE6E" box="[647,682,403,429]" count="1" pageId="24" pageNumber="83" type="male">1♂</specimenCount>
(FIOC): (
<collectionCode id="30B4EEBCE170FF9FFD63FE36FCB9FE6E" box="[698,769,403,429]" collectionName="Brazil, Rio de Janeiro, Rio de Janeiro, Fundacao Instituto Oswaldo Cruz" pageId="24" pageNumber="83">FIOC</collectionCode>
):
<collectingCountry id="2EB236E9E170FF9FFCCDFE36FCE2FE6E" box="[788,858,403,429]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFCCDFE36FCE2FE6E" box="[788,858,403,429]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. E. do Rio [ . E. do Rio [
<collectingRegion id="9461B89BE170FF9FFC39FE36FB35FE6E" box="[992,1165,403,429]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFC39FE36FB35FE6E" box="[992,1165,403,429]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
]. Itatiaia. 10.viii.[19]44. J.F. Zikan col; ].
<collectingMunicipality id="B67EEC03E170FF9FFB46FE36FB48FE6E" box="[1183,1264,403,429]" pageId="24" pageNumber="83">Itatiaia</collectingMunicipality>
. 10.viii.[19]44.
<collectorName id="FB5013AFE170FF9FFF4EFE12FEBDFE12" box="[151,261,439,465]" pageId="24" pageNumber="83">J.F. Zikan</collectorName>
col
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFEE7FE12FA20FDFE" collectingDate="2016-10-30" collectingDateMax="2016-11-06" collectingDateMin="2016-10-30" collectionCode="FIOC" collectorName="OV Conle &amp; FH Hennemann &amp; Uhlig &amp; Y Gutierrez &amp; P Valero &amp; J Costa &amp; Rodrigues &amp; PABA Neves &amp; PI Chiquetto-Machado &amp; VM Ghirotto &amp; PW Engelking" country="Brazil" elevation="920" latitude="-22.429722" location="Parque Nacional do Itatiaia" longLatPrecision="21" longitude="-44.618053" pageId="24" pageNumber="83" specimenCode="PNI022, PNI037-40, PNI094, PNI119-120, PNI206, PNI218-221" specimenCount="21" specimenCount-egg="1" specimenCount-female="7" specimenCount-male="13" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFEE7FE12FED3FE12" box="[318,363,439,465]" count="13" pageId="24" pageNumber="83" type="male">13♂</specimenCount> <specimenCount id="40A3BDF0E170FF9FFEE7FE12FED3FE12" box="[318,363,439,465]" count="13" pageId="24" pageNumber="83" type="male">13♂</specimenCount>
(PNI022, PNI037–40, PNI094), (
<specimenCode id="0603DE02E170FF9FFEAAFE12FE6AFE12" box="[371,466,439,465]" pageId="24" pageNumber="83">PNI022</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFE07FE12FDD9FE12" box="[478,609,439,465]" pageId="24" pageNumber="83">PNI037–40</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFDB4FE12FD71FE12" box="[621,713,439,465]" pageId="24" pageNumber="83">PNI094</specimenCode>
),
<specimenCount id="40A3BDF0E170FF9FFD02FE12FD47FE12" box="[731,767,439,465]" count="7" pageId="24" pageNumber="83" type="female">7♀</specimenCount> <specimenCount id="40A3BDF0E170FF9FFD02FE12FD47FE12" box="[731,767,439,465]" count="7" pageId="24" pageNumber="83" type="female">7♀</specimenCount>
(PNI119–120, PNI206, PNI218–221), eggs (FIOC): (
<specimenCode id="0603DE02E170FF9FFCD4FE12FC18FE12" box="[781,928,439,465]" pageId="24" pageNumber="83">PNI119–120</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFC72FE12FBBDFE12" box="[939,1029,439,465]" pageId="24" pageNumber="83">PNI206</specimenCode>
,
<specimenCode id="0603DE02E170FF9FFBC9FE12FB1BFE12" box="[1040,1187,439,465]" pageId="24" pageNumber="83">PNI218–221</specimenCode>
),
<specimenCount id="40A3BDF0E170FF9FFB6EFE12FB52FE12" box="[1207,1258,439,465]" pageId="24" pageNumber="83" type="egg">eggs</specimenCount>
(
<collectionCode id="30B4EEBCE170FF9FFB20FE12FAF8FE12" box="[1273,1344,439,465]" pageId="24" pageNumber="83">FIOC</collectionCode>
):
<collectingCountry id="2EB236E9E170FF9FFA8BFE12FA20FE12" box="[1362,1432,439,465]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFA8BFE12FA20FE12" box="[1362,1432,439,465]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE170FF9FFF4EFE7EFE84FE36" box="[151,316,475,501]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFF4EFE7EFE84FE36" box="[151,316,475,501]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
. Parque Nacional do Itatiaia, along BR 485 betw. park entrance &amp; Cachoeira de Maromba, 22°45’97 S 44°59’91 W – .
<geoCoordinate id="339110BEE170FF9FFE8FFE5BFE6FFDDA" box="[342,471,510,537]" degrees="22" direction="south" minutes="25" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="47" value="-22.429722">22°25’47 S</geoCoordinate> <location id="537A20A2E170FF9FFE9FFE7EFDC1FE36" LSID="urn:lsid:plazi:treatment:DE0CC76FE17FFFA2FF4EF934FAB0FE6E:537A20A2E170FF9FFE9FFE7EFDC1FE36" box="[326,633,475,501]" country="Brazil" latitude="-22.429722" longLatPrecision="21" longitude="-44.618053" name="Parque Nacional do Itatiaia" pageId="24" pageNumber="83" stateProvince="Rio de Janeiro">Parque Nacional do Itatiaia</location>
<geoCoordinate id="339110BEE170FF9FFE07FE5BFDD1FDDA" box="[478,617,510,537]" degrees="44" direction="west" minutes="37" orientation="longitude" pageId="24" pageNumber="83" precision="15" seconds="05" value="-44.618053">44°37’05 W</geoCoordinate>
, ,
<quantity id="915DDB9CE170FF9FFDACFE5AFCBAFDDA" box="[629,770,511,537]" metricMagnitude="2" metricUnit="m" metricValue="9.2" metricValueMax="11.2" metricValueMin="7.2" pageId="24" pageNumber="83" unit="m" value="920.0" valueMax="1120.0" valueMin="720.0">720–1120 m</quantity> <locationDeviation id="47A86F5BE170FF9FFD5AFE7EFA91FE36" box="[643,1321,474,501]" pageId="24" pageNumber="83">along BR 485 betw. park entrance &amp; Cachoeira de Maromba</locationDeviation>
a.s.l. ,
<date id="221B50B9E170FF9FFC9CFE5AFBBFFDDA" box="[837,1031,511,537]" pageId="24" pageNumber="83" value="2016-10-30" valueMax="2016-11-06" valueMin="2016-10-30">30.x.–06.xi.2016</date> <geoCoordinate id="339110BEE170FF9FFAEDFE7FFE81FDDA" degrees="22" direction="south" minutes="45" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="97" value="-22.426388">22°45’97 S 44°59’91 W</geoCoordinate>
. OV Conle, FH Hennemann, Uhlig, Y Gutiérrez, P Valero, J Costa, Rodrigues, PABA Neves, PI Chiquetto-Machado, VM Ghirotto, PW Engelking col.; – 22°25’47 S 44°37’05 W, 720–1120 m a.s.l.
<date id="221B50B9E170FF9FFC9CFE5AFBBFFDDA" box="[837,1031,511,537]" pageId="24" pageNumber="83" value="2016-10-30" valueMax="2016-11-06" valueMin="2016-10-30">
<collectingDate id="325FA951E170FF9FFC9CFE5AFBBFFDDA" box="[837,1031,511,537]" pageId="24" pageNumber="83" value="2016-10-30" valueMax="2016-11-06" valueMin="2016-10-30">30.x.–06.xi.2016</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFBCBFE5AFB30FDDA" box="[1042,1160,511,537]" pageId="24" pageNumber="83">OV Conle</collectorName>
,
<collectorName id="FB5013AFE170FF9FFB4DFE5AFAF1FDDA" box="[1172,1353,511,537]" pageId="24" pageNumber="83">FH Hennemann</collectorName>
,
<collectorName id="FB5013AFE170FF9FFA8CFE5AFA20FDDA" box="[1365,1432,511,537]" pageId="24" pageNumber="83">Uhlig</collectorName>
,
<collectorName id="FB5013AFE170FF9FFF4EFD86FEA6FDFE" box="[151,286,547,573]" pageId="24" pageNumber="83">Y Gutiérrez</collectorName>
,
<collectorName id="FB5013AFE170FF9FFEF1FD86FE31FDFE" box="[296,393,547,573]" pageId="24" pageNumber="83">P Valero</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE4DFD86FE50FDFE" box="[404,488,547,573]" pageId="24" pageNumber="83">J Costa</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE2AFD86FDD1FDFE" box="[499,617,547,573]" pageId="24" pageNumber="83">Rodrigues</collectorName>
,
<collectorName id="FB5013AFE170FF9FFDADFD86FCB2FDFE" box="[628,778,547,573]" pageId="24" pageNumber="83">PABA Neves</collectorName>
,
<collectorName id="FB5013AFE170FF9FFCCDFD86FBAFFDFE" box="[788,1047,547,573]" pageId="24" pageNumber="83">PI Chiquetto-Machado</collectorName>
,
<collectorName id="FB5013AFE170FF9FFBF8FD86FB0FFDFE" box="[1057,1207,547,573]" pageId="24" pageNumber="83">VM Ghirotto</collectorName>
,
<collectorName id="FB5013AFE170FF9FFB18FD86FADCFDFE" box="[1217,1380,547,573]" pageId="24" pageNumber="83">PW Engelking</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="E6CD7C24E170FF9FFF4EFDE2FCD6FD6A" collectingDate="2016-04-11" collectionCode="FIOC" collectorName="OV Conle &amp; FH Hennemann &amp; Uhlig &amp; Y Gutierrez &amp; P Valero &amp; J Costa &amp; Rodrigues &amp; PABA Neves &amp; PI Chiquetto-Machado &amp; VM Ghirotto &amp; PW Engelking" country="Brazil" elevation="765" latitude="-22.452778" location="Parque Nacional do Itatiaia" longLatPrecision="21" longitude="-44.609997" pageId="24" pageNumber="83" specimenCode="PNI036" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro">
<specimenCount id="40A3BDF0E170FF9FFF4EFDE2FF08FDA2" box="[151,176,583,609]" count="1" pageId="24" pageNumber="83" type="female">1♀</specimenCount> <specimenCount id="40A3BDF0E170FF9FFF4EFDE2FF08FDA2" box="[151,176,583,609]" count="1" pageId="24" pageNumber="83" type="female">1♀</specimenCount>
(FIOC PNI036): (
<collectionCode id="30B4EEBCE170FF9FFF61FDE2FEBBFDA2" box="[184,259,583,609]" pageId="24" pageNumber="83">FIOC</collectionCode>
<specimenCode id="0603DE02E170FF9FFED5FDE2FED0FDA2" box="[268,360,583,609]" pageId="24" pageNumber="83">PNI036</specimenCode>
):
<collectingCountry id="2EB236E9E170FF9FFEA5FDE2FE7AFDA2" box="[380,450,583,609]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry> <collectingCountry id="2EB236E9E170FF9FFEA5FDE2FE7AFDA2" box="[380,450,583,609]" name="Brazil" pageId="24" pageNumber="83">Brasil</collectingCountry>
. .
<collectingRegion id="9461B89BE170FF9FFE17FDE2FDCFFDA2" box="[462,631,583,609]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion> <collectingRegion id="9461B89BE170FF9FFE17FDE2FDCFFDA2" box="[462,631,583,609]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="83">Rio de Janeiro</collectingRegion>
. Parque Nacional do Itatiaia, near Lago Azul, .
<location id="537A20A2E170FF9FFD5BFDE2FC02FDA2" LSID="urn:lsid:plazi:treatment:DE0CC76FE17FFFA2FF4EF934FAB0FE6E:537A20A2E170FF9FFD5BFDE2FC02FDA2" box="[642,954,583,609]" country="Brazil" latitude="-22.452778" longLatPrecision="21" longitude="-44.609997" name="Parque Nacional do Itatiaia" pageId="24" pageNumber="83" stateProvince="Rio de Janeiro">Parque Nacional do Itatiaia</location>
,
<locationDeviation id="47A86F5BE170FF9FFC1FFDE2FBC0FDA2" box="[966,1144,583,609]" pageId="24" pageNumber="83">near Lago Azul</locationDeviation>
,
<geoCoordinate id="339110BEE170FF9FFB5DFDE2FABDFDA2" box="[1156,1285,583,609]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="10" value="-22.452778">22°27’10 S</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFB5DFDE2FABDFDA2" box="[1156,1285,583,609]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="24" pageNumber="83" precision="15" seconds="10" value="-22.452778">22°27’10 S</geoCoordinate>
<geoCoordinate id="339110BEE170FF9FFAD4FDE2FA20FDA2" box="[1293,1432,583,609]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="24" pageNumber="83" precision="15" seconds="36" value="-44.609997">44°36’36 W</geoCoordinate> <geoCoordinate id="339110BEE170FF9FFAD4FDE2FA20FDA2" box="[1293,1432,583,609]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="24" pageNumber="83" precision="15" seconds="36" value="-44.609997">44°36’36 W</geoCoordinate>
, 750–780 m a.s.l.
<date id="221B50B9E170FF9FFE80FDCEFE62FD46" box="[345,474,619,645]" pageId="24" pageNumber="83" value="2016-04-11">
<collectingDate id="325FA951E170FF9FFE80FDCEFE62FD46" box="[345,474,619,645]" pageId="24" pageNumber="83" value="2016-04-11">04.11.2016</collectingDate>
</date>
.
<collectorName id="FB5013AFE170FF9FFE3CFDCEFDE2FD46" box="[485,602,619,645]" pageId="24" pageNumber="83">OV Conle</collectorName>
, ,
<quantity id="915DDB9CE170FF9FFF4EFDCFFEAFFD46" box="[151,279,618,645]" metricMagnitude="2" metricUnit="m" metricValue="7.65" metricValueMax="7.8" metricValueMin="7.5" pageId="24" pageNumber="83" unit="m" value="765.0" valueMax="780.0" valueMin="750.0">750–780 m</quantity> <collectorName id="FB5013AFE170FF9FFDBFFDCEFCA3FD46" box="[614,795,619,645]" pageId="24" pageNumber="83">FH Hennemann</collectorName>
a.s.l. ,
<date id="221B50B9E170FF9FFE80FDCEFE62FD46" box="[345,474,619,645]" pageId="24" pageNumber="83" value="2016-04-11">04.11.2016</date> <collectorName id="FB5013AFE170FF9FFCFEFDCEFCD2FD46" box="[807,874,619,645]" pageId="24" pageNumber="83">Uhlig</collectorName>
. OV Conle, FH Hennemann, Uhlig, Y Gutiérrez, P Valero, J Costa, Rodrigues, PABA Neves, PI Chiquetto-Machado, VM Ghirotto, PW Engelking col. ,
<collectorName id="FB5013AFE170FF9FFCADFDCEFC43FD46" box="[884,1019,619,645]" pageId="24" pageNumber="83">Y Gutiérrez</collectorName>
,
<collectorName id="FB5013AFE170FF9FFBDFFDCEFBD0FD46" box="[1030,1128,619,645]" pageId="24" pageNumber="83">P Valero</collectorName>
,
<collectorName id="FB5013AFE170FF9FFBAAFDCEFB71FD46" box="[1139,1225,619,645]" pageId="24" pageNumber="83">J Costa</collectorName>
,
<collectorName id="FB5013AFE170FF9FFB0DFDCEFAF2FD46" box="[1236,1354,619,645]" pageId="24" pageNumber="83">Rodrigues</collectorName>
,
<collectorName id="FB5013AFE170FF9FFA8FFDCEFF58FD6A" pageId="24" pageNumber="83">PABA Neves</collectorName>
,
<collectorName id="FB5013AFE170FF9FFF32FD2AFE56FD6A" box="[235,494,655,681]" pageId="24" pageNumber="83">PI Chiquetto-Machado</collectorName>
,
<collectorName id="FB5013AFE170FF9FFE21FD2AFD37FD6A" box="[504,655,655,681]" pageId="24" pageNumber="83">VM Ghirotto</collectorName>
,
<collectorName id="FB5013AFE170FF9FFD40FD2AFC85FD6A" box="[665,829,655,681]" pageId="24" pageNumber="83">PW Engelking</collectorName>
col.
</materialsCitation>
</paragraph> </paragraph>
</subSubSection> </subSubSection>
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