From bc7b071f59077173ef5b827e80a539b4e0bbcbdb Mon Sep 17 00:00:00 2001 From: ggserver Date: Tue, 5 Nov 2024 19:11:44 +0000 Subject: [PATCH] Add updates up until 2024-11-05 19:05:37 --- ...l => 004A87E8FFCCFFB6FF2AFEF3F898F886.xml} | 389 ++++----- .../C6/627EC678F743FF95FF4EFB607FD1FA92.xml | 383 +++++++++ .../C6/627EC678F74BFF91FF4EF98E79FCF965.xml | 350 ++++++-- .../C6/627EC678F74DFF94FF4EFD927858F97B.xml | 297 +++++++ .../C6/627EC678F753FF83FF4EF89B7E45FD42.xml | 563 +++++++++++++ ...l => 627EC678F75AFF83FF4EFA1279FDF82D.xml} | 162 ++-- .../C6/627EC678F761FFB7FF4EF9427DE5FEA2.xml | 100 +-- .../C6/627EC678F761FFB8FF4EFBB27EC4F97A.xml | 100 +-- .../C6/627EC678F763FFB8FF4EF94F7F2BFC2A.xml | 106 +-- .../C6/627EC678F763FFBAFF4EFE8E78F2F97F.xml | 284 +++++++ .../C6/627EC678F76CFFB5FF4EFD187875F864.xml | 303 +++++++ .../C6/627EC678F76DFFACFF4EFC487E68FBBA.xml | 757 ++++++++++++++++++ .../C6/627EC678F76EFFB7FF4EFBCB7CB9F95F.xml | 96 +-- .../C6/627EC678F76EFFB7FF4EFE3B7D0EFBF3.xml | 96 +-- .../C6/627EC678F772FFABFF4EFC077D48F800.xml | 236 ++++++ .../C7/DE0CC76FE16AFF8DFF4EF9A9FC09F90C.xml | 140 ++-- .../C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml | 429 +++++++--- 17 files changed, 4007 insertions(+), 784 deletions(-) rename data/00/4A/87/{004A87E8FFCCFFBFFF2AFEF3F8AEFDB2.xml => 004A87E8FFCCFFB6FF2AFEF3F898F886.xml} (93%) create mode 100644 data/62/7E/C6/627EC678F743FF95FF4EFB607FD1FA92.xml create mode 100644 data/62/7E/C6/627EC678F74DFF94FF4EFD927858F97B.xml create mode 100644 data/62/7E/C6/627EC678F753FF83FF4EF89B7E45FD42.xml rename data/62/7E/C6/{627EC678F75AFF83FF4EFA1279F9F82D.xml => 627EC678F75AFF83FF4EFA1279FDF82D.xml} (63%) create mode 100644 data/62/7E/C6/627EC678F763FFBAFF4EFE8E78F2F97F.xml create mode 100644 data/62/7E/C6/627EC678F76CFFB5FF4EFD187875F864.xml create mode 100644 data/62/7E/C6/627EC678F76DFFACFF4EFC487E68FBBA.xml create mode 100644 data/62/7E/C6/627EC678F772FFABFF4EFC077D48F800.xml diff --git a/data/00/4A/87/004A87E8FFCCFFBFFF2AFEF3F8AEFDB2.xml b/data/00/4A/87/004A87E8FFCCFFB6FF2AFEF3F898F886.xml similarity index 93% rename from data/00/4A/87/004A87E8FFCCFFBFFF2AFEF3F8AEFDB2.xml rename to data/00/4A/87/004A87E8FFCCFFB6FF2AFEF3F898F886.xml index 4c989888f5b..fa156520f48 100644 --- a/data/00/4A/87/004A87E8FFCCFFBFFF2AFEF3F8AEFDB2.xml +++ b/data/00/4A/87/004A87E8FFCCFFB6FF2AFEF3F898F886.xml @@ -1,68 +1,70 @@ - - - -Lethrus (Heteroplistodus) kucerai (Coleoptera: Geotrupidae: Lethrinae), a new species from China + + + +Lethrus (Heteroplistodus) kucerai (Coleoptera: Geotrupidae: Lethrinae), a new species from China - - -Author + + +Author -Hillert, Oliver -Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. +Hillert, Oliver +Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. - - -Author + + +Author -Král, David -Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00 Praha 2, Czechia. +Král, David +Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00 Praha 2, Czechia. - - -Author + + +Author -Bagaturov, Mikhail -Zoological Institute of Russian Academy of Sciences, St. Petersburg, Universitetskaya naberezhnaja 1, Russia; Athens Institute for Education and Research, 7 Eptanisou Street, Vrilissia, Athens, Greece. +Bagaturov, Mikhail +Zoological Institute of Russian Academy of Sciences, St. Petersburg, Universitetskaya naberezhnaja 1, Russia; Athens Institute for Education and Research, 7 Eptanisou Street, Vrilissia, Athens, Greece. - - -Author + + +Author -Chuluunbaatar, Gantigmaa -Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. & Entomological Laboratory Institute of Biology, Mongolian Academy of Science, Peace Avenue- 54 B, Bayanzurkh, Ulaanbaatar- 13330, Mongolia. & Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. +Chuluunbaatar, Gantigmaa +Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. & Entomological Laboratory Institute of Biology, Mongolian Academy of Science, Peace Avenue- 54 B, Bayanzurkh, Ulaanbaatar- 13330, Mongolia. & Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-24 + +2024 + +2024-10-24 - -5529 + +5529 - -1 + +1 - -123 -143 + +123 +143 - -http://dx.doi.org/10.11646/zootaxa.5529.1.6 + +http://dx.doi.org/10.11646/zootaxa.5529.1.6 -journal article -10.11646/zootaxa.5529.1.6 -1175-5326 -14022196 -598225C3-9B8F-4C0D-8E75-098F9943091D +journal article +304988 +10.11646/zootaxa.5529.1.6 +2d502b69-b834-4a2e-a615-d3a396bf0b96 +1175-5326 +14022196 +598225C3-9B8F-4C0D-8E75-098F9943091D - + @@ -123,7 +125,9 @@ Jakovlev 1890: 565 -(key, classification). +(key, classification). + + @@ -389,7 +393,8 @@ Yang (distribution). - + + Type @@ -535,8 +540,6 @@ Nikolajev [p, hw] || HNHM ). - - Additional material examined ( @@ -555,9 +558,9 @@ Nikolajev [p, hw] || ( ZSMC ), “China, -Prov. Kansu. +Prov. Kansu. , -Tscholo Shan +Tscholo Shan ” [not mapped]; @@ -569,7 +572,7 @@ Nikolajev [p, hw] || , Zhangye , -Shandan +Shandan , 12.v.2011 @@ -581,7 +584,7 @@ Nikolajev [p, hw] || lgt . - + China , @@ -596,9 +599,7 @@ lgt , col. Bain-Khuduk , 2.– - -4.vi. - +4.vi. [19]09, ex. of Kozlov; @@ -606,7 +607,7 @@ lgt ( ZMAS ), -Alashan desert +Alashan desert , beginning of x.–xi. [19]01, Kozlov and @@ -618,7 +619,7 @@ and ( HBUM ), -Alashanyouqi +Alashanyouqi , 21.vii.1988 @@ -689,7 +690,7 @@ lgt.; E of Bayan Hot [= -Alxa +Alxa ], 38°51'N 105°47'E @@ -714,7 +715,7 @@ lgt.; , E -Bayan Hot +Bayan Hot , 1780 m @@ -739,7 +740,7 @@ lg.; , E -Bayan Hot +Bayan Hot , 38.805N 105.748E @@ -758,7 +759,7 @@ lgt.; , E -Bayan Hot +Bayan Hot , 1700-1780 m @@ -790,7 +791,7 @@ lgt.; , E -Bayan Hot +Bayan Hot , 1750–1800 m @@ -812,7 +813,7 @@ lgt.; , -Dengkou +Dengkou , 27.vii.1988 @@ -827,7 +828,7 @@ lgt.; , -Tangsag Bulag +Tangsag Bulag , 8.vii.1988 @@ -849,7 +850,7 @@ lgt.; SW Wuhai , -Wuda. +Wuda. 8.v.1995 @@ -863,7 +864,7 @@ lgt.; , -Wulatehouqi +Wulatehouqi , 20.vii.2020 @@ -889,7 +890,7 @@ lgt Zhongning , -Baima +Baima , 3.iv.2016 @@ -906,7 +907,7 @@ lgt.; Zhongwei , -Gantang +Gantang , 6.v.1992 @@ -921,7 +922,7 @@ lgt.; , -Haiyuan +Haiyuan , 30.v.1995 @@ -936,7 +937,7 @@ lgt.; , -Lingwu +Lingwu , 19.v.1992 @@ -951,7 +952,7 @@ lgt.; , -Pingluo +Pingluo , 5.viii.1989 @@ -966,7 +967,7 @@ lgt.; , -Pingluo +Pingluo , 5.viii.1989 @@ -981,7 +982,7 @@ lgt.; , -Yongning Xian +Yongning Xian , 12.vii.1997 @@ -996,7 +997,7 @@ lgt.; , -Zhongwei +Zhongwei , iv.1985 @@ -1015,7 +1016,7 @@ lgt.; Xixia , -Western Xia Tombs +Western Xia Tombs env. 38°26'20"N 105°58'32"E @@ -1041,7 +1042,7 @@ lgt.; Helan Shan , -road to Suyukou National Forest Park +road to Suyukou National Forest Park , 28 km @@ -1086,7 +1087,7 @@ Mongolia, , -Bulgan somon +Bulgan somon , 4.ix.1970 @@ -1103,7 +1104,7 @@ lgt.; Omnogov’ , -Bulgan +Bulgan env., 16.iv.1972 @@ -1118,7 +1119,7 @@ lgt.; , -S. Gobi aimak +S. Gobi aimak , 30 km @@ -1145,16 +1146,14 @@ N of Bulgan ) , - + Eastern Gobi aimak , Agarut env., -Emelyanov +Emelyanov lgt., - -27.vi. - +27.vi. [1]971; 2 ♂♂ ( @@ -1170,7 +1169,7 @@ aimak, N of Agarut , -Emelyanov +Emelyanov lgt., 28.vi.1971 @@ -1183,7 +1182,7 @@ lgt., , - + 5 km N Agaruta @@ -1209,7 +1208,7 @@ aimak, , S. Sain-Shand , -Eglon +Eglon lgt., 24-25.vii.1949 @@ -1222,7 +1221,7 @@ lgt., , -Dalanjargalan +Dalanjargalan , 28.vii.2016 @@ -1237,7 +1236,7 @@ lgt.; , -Hubsgula +Hubsgula , 25 km @@ -1259,7 +1258,7 @@ lgt.; , -Kharayra +Kharayra 50 km SO Kharayra @@ -1280,7 +1279,7 @@ lgt.; , -East-Gobi Aimak +East-Gobi Aimak , 1120 m @@ -1309,7 +1308,7 @@ lgt.; Donggebi , -Khatanbulag +Khatanbulag , 15.vii.2015 @@ -1324,7 +1323,7 @@ lgt.; , -East-Gobi Aimak +East-Gobi Aimak , 7 km @@ -1347,7 +1346,7 @@ lgt.; , -Eastern Gobi aimak +Eastern Gobi aimak , 25 km @@ -1369,7 +1368,7 @@ NNW Khuvsgel , -Mandakh +Mandakh 12 km NO Mandakh @@ -1392,7 +1391,7 @@ lgt.; Mongolia , -S. Gobi aimak +S. Gobi aimak , 60 km @@ -1413,7 +1412,7 @@ lgt.; , same data but -Emelyanov +Emelyanov lgt., 1 ♂ ( @@ -1423,9 +1422,7 @@ lgt., , same data but - -17.vi. - +17.vi. [1]971, Kozlov lgt.; @@ -1445,7 +1442,7 @@ pr., S of Saynshand , -Burdene Bulag mts. +Burdene Bulag mts. , 1000 m @@ -1486,7 +1483,7 @@ lgt.; Zuun bayan -Argalant +Argalant , 80 km @@ -1536,7 +1533,7 @@ lgt.; Bulgan , -Bayanzag +Bayanzag , 21.vi.1971 @@ -1551,13 +1548,13 @@ lgt.; , -Bulgan +Bulgan , 12.v.1962 , -Mongol-Germany joint expedition +Mongol-Germany joint expedition ; 1 ♂ ( @@ -1566,13 +1563,13 @@ lgt.; , -Bulgan +Bulgan , 26.v.1962 , -Mongol-Germany joint expedition +Mongol-Germany joint expedition ; 1 ♀ ( @@ -1581,7 +1578,7 @@ lgt.; , -Bulgan +Bulgan , 24.vi.1971 @@ -1596,7 +1593,7 @@ lgt.; , -Bulgan +Bulgan , 31.vii.1971 @@ -1611,7 +1608,7 @@ lgt.; , -Bulgan +Bulgan , 08.viii.1971 @@ -1626,7 +1623,7 @@ lgt.; , -Bulgan +Bulgan , 24.vii.1971 @@ -1641,7 +1638,7 @@ lgt.; , -Gobi +Gobi , 100 km @@ -1668,7 +1665,7 @@ lgt.; SW Dalanzadgad , -Bayan-Dalai +Bayan-Dalai , 1.–2.vii.2003 @@ -1682,19 +1679,17 @@ lgt.; ) , - -S. Gobi aimak + +S. Gobi aimak , -Khongorin-els +Khongorin-els , 60 km WNW Bann-Dalay , - -30.–31.vii. - +30.–31.vii. [1]976, Zaitsev lgt.; @@ -1707,7 +1702,7 @@ lgt.; Omnogov’ aimak, -env. of Khurmek +env. of Khurmek [not mapped], 15.v.1962 @@ -1724,7 +1719,7 @@ lgt.; , -Manlai +Manlai , 24.viii.2011 @@ -1739,7 +1734,7 @@ lgt.; , -Manlay +Manlay , 60 km @@ -1761,7 +1756,7 @@ lgt.; , -Manlay +Manlay , 45 km @@ -1783,7 +1778,7 @@ lgt.; , -Mušugaj +Mušugaj ; 1 ♀ ( @@ -1791,7 +1786,7 @@ lgt.; ) , - + Mongolia , @@ -1799,9 +1794,7 @@ lgt.; SSW of Nomgon , - -9.viii. - +9.viii. [19]67, Kerzhner lgt.; @@ -1811,17 +1804,15 @@ lgt.; ) , - -Birzon-Gobi + +Birzon-Gobi , 50 km S Nomgon , - -9.viii. - +9.viii. [1]967, Zaitsev lgt.; @@ -1834,9 +1825,7 @@ lgt.; E of brook Talin-Bilgekh-Bulak , Zaitsev lgt., - -19.viii. - +19.viii. [1]1969 . @@ -1856,7 +1845,7 @@ E of brook Talin-Bilgekh-Bulak Omno-gov’ aimak, -env. of Bajan-Dalai +env. of Bajan-Dalai , 15.vi.1967 @@ -1873,7 +1862,7 @@ aimak, Hanbogd Bayan brigade , -Sookheni sand +Sookheni sand , 20.vii.1974 @@ -1887,8 +1876,8 @@ aimak, ) , - -Eastern Gobi + +Eastern Gobi aimak, 20 km @@ -1897,9 +1886,7 @@ ENE kol. [?] Tal-Khongorin-Khuduk, Emelyanov lgt. - -2.vii. - +2.vii. [1]971; 1 ♂ ( @@ -1910,7 +1897,7 @@ lgt. Bulgan , -Ulaan Nuur +Ulaan Nuur , 07.ix.1971 @@ -1925,7 +1912,7 @@ lgt.; , -S. Gobi aimak +S. Gobi aimak , Mount. Tzagan-Ula , @@ -1946,7 +1933,7 @@ lgt., Uvurkhangai [= Öwörchangai], -Artz-Bogd +Artz-Bogd [not located], 18.vi.1962 @@ -1958,7 +1945,7 @@ lgt., ( ELIB ), -Mongol-Germany +Mongol-Germany joint expedition, 26.v.1962 @@ -1981,7 +1968,7 @@ joint expedition, SW of Ulanbatar , -river Tula +river Tula , vi.1983 @@ -2023,12 +2010,10 @@ lgt. ) , - -Budurganin-khuduk + +Budurganin-khuduk , - -24.–28.v. - +24.–28.v. [19]09, ex. of Kozlov ; @@ -2038,15 +2023,13 @@ lgt. ) , - + Centr. Mongolia , S. pr. -Dundu-sankhang +Dundu-sankhang [?], - -18.vi.–2.vii. - +18.vi.–2.vii. [19]09, ex. of Kozlov ; @@ -2056,13 +2039,11 @@ Centr. ) , - + Din-uan-in, -Kureto +Kureto [?], 4.– - -7.v. - +7.v. [19]09, ex. of Kozlov ; @@ -2072,13 +2053,11 @@ Din-uan-in, ) , - + col. Kuku-shande , Dizhintata, 4.– - -5.vi. - +5.vi. [19]09, ex. of Kozlov; @@ -2088,13 +2067,9 @@ of Kozlov; ) , - -Alashan -, Dyn-Yuan-in, - -20.–25.iv. - -[10]09, ex. of Kozlov; + +Alashan +, Dyn-Yuan-in, 20.–25.iv.[10]09, ex. of Kozlov; 1 ♂ ( ZMAS @@ -2102,7 +2077,7 @@ of Kozlov; , -Gobi +Gobi , col. Sabura ; @@ -2112,13 +2087,11 @@ of Kozlov; ) , - + Col. Tsatkholung, Kharmiktan-Kuduk , - -1.vi. - +1.vi. [19]09, ex. of Kozlov; 2 ♂♂ ( @@ -2126,13 +2099,11 @@ Col. Tsatkholung, ) , - + Lan-gzhou-fu, -Khuan-khe +Khuan-khe [river] coast, - -25.iii. - +25.iii. [19]09, ex. of Kozlov ; @@ -2142,8 +2113,8 @@ Lan-gzhou-fu, ) , - -S. Gobi aimak, + +S. Gobi aimak, 25 km ENE Monlay @@ -2151,9 +2122,7 @@ ENE Monlay , lgt. Kerzhner , - -16.vi. - +16.vi. [1]971; 2 ♂♂ ( @@ -2161,7 +2130,7 @@ ENE Monlay ) , - + Eastern Gobi aimak, @@ -2171,9 +2140,7 @@ W of lake Tenger-nur , Kozlov lgt., - -25.vi. - +25.vi. [1]971; 3 ♂♂ ( @@ -2181,7 +2148,7 @@ lgt., ) , - + Eastern Gobi aimak, @@ -2191,9 +2158,7 @@ W of lake Tenger-nur , Emelyanov lgt., - -25.vi. - +25.vi. [1]971; 2 ♂♂ ( @@ -2201,7 +2166,7 @@ lgt., ), S. Gobi aimak, - + 30 km NE Tsailan @@ -2696,20 +2661,22 @@ considerably short, almost as long as wide, glabrous. Legs . Profemora with indistinct transversal edge anteroventrally; protibia with row of nine gradually proximad diminishing exterior denticles, and with row of pointed or broad tubercles on ventromedial edge. Apical spur of protibia pointed. - + Aedeagus ( Figs. 20 , 22 -). Parameres broadly rounded apically. +). Parameres broadly rounded apically. + + Differential diagnosis. Refer to character matrix on -Table 1 +Table 1 . - - + + TABLE 1. Character matrix for the differential diagnosis of males of the @@ -2719,10 +2686,9 @@ Character matrix for the differential diagnosis of males of the ) potanini -species +species group. -group. @@ -2805,7 +2771,7 @@ Total body length . - + Distribution. China @@ -2833,6 +2799,8 @@ and Ulaanbaatar . + + Note. In the type series we had examined two different taxa we found. A @@ -2960,8 +2928,5 @@ is designated here to fix the uniqueness of the species ). - - - \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F743FF95FF4EFB607FD1FA92.xml b/data/62/7E/C6/627EC678F743FF95FF4EFB607FD1FA92.xml new file mode 100644 index 00000000000..3233b5c83a6 --- /dev/null +++ b/data/62/7E/C6/627EC678F743FF95FF4EFB607FD1FA92.xml @@ -0,0 +1,383 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Neoleptastacus supersetosus + +sp. nov. + + + + + + +( +Figs 9–11 +) + + +https://zoobank.org/ +316E69F8-8817-4599-9126-F24A67BE3579 + + + + + + +Pararenopontia poliseta +—nomen nudum + +by + +Sak (2004: 273) + + + + + + +Type locality. +Kuwait +, +Shuwaikh +( +29°21’41.8” N +, +47°57’19.2” E +) mid- and upper tide levels on public recreational beach within +Kuwait +Bay adjacent to the commercial port, surrounded by extensive urban development. Relatively steeply sloping sandy beach, bounded to seaward by extensive limestone flats. + + +Type material. + +Holotype +♀ +(dissected on eight slides) (reg. no +NHMUK 2024.1063 +) + +. + +Paratypes +are +one ♂ +(dissected) and +one ♀ +and +one ♂ +preserved in alcohol (reg. nos +NHMUK 2024.1064–1066 +). All specimens were collected from the type locality; leg. +S. Livesey +and +N. O’Brien +, + +3 June 2004 + + +. + + + + +Description of female. +Total body length from tip of rostrum to posterior margin of caudal rami 225 μm ( +n += 1). Body ( +Fig. 9A +) slender and cylindrical without clear distinction between prosome and urosome. Hyaline frills of thoracic somites weakly developed and smooth; those of genital double-somite and free abdominal somites strongly developed and consisting of rectangular digitate lappets ( +Figs 9A +; +10A +; +11C +). Somites connected by well-developed intersomitic membranes. Somites bearing P2–P5, genital double-somite and second abdominal somite with internal semicircular reinforcements forming pattern as figured. Genital double-somite ( +Figs 9A +; +10A +) about as long as wide (measured in ventral aspect); with two ventral pores in posterior half ( +Figs 10A +). + + +Anal somite ( +Fig. 10A +; +11C, D +) without paired, dorsally recurved, spinous processes; with two sensilla dorsally and few spinules either side of ventral midline.Anal operculum well developed, semicircular, without ornamentation; covering terminally positioned anal opening. Anal frill triradiate, distinctly incised (giving a spinulose appearance underneath anal operculum). + + + + + +Caudal rami ( +Figs 10A +; +11C, D +) about 2.2 times longer than wide (measured in lateral view from anterior margin to apex of spinous process), distinctly tapering posteriorly with outer margin almost straight and inner margin distinctly convex; outer distal corner produced into short posteriorly directed, dorsally recurved spinous process; with pore ventrally near proximal margin ( +Fig. 10A +) and dorsally at base of spinous process ( +Fig. 11C +); inner margin with conspicuous spinules near base of seta VII ( +Fig. 11C +). Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, naked, located between seta V and posterior spinous process; seta V long, naked, with proximal fracture plane; seta VI small, naked and located at inner distal corner; seta VII distinctly foliaceous and tri-articulate at base. + + +Rostrum ( +Fig. 11B +) small, broadly subtriangular, tapering distally; apical part lobate and demarcated by bilateral constrictions; with two delicate sensilla. + + +Antennule ( +Fig. 11E +) moderately long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about three times longer than wide. Segment 4 with long aesthetasc (L: 21 μm) fused at base with seta. Distal segment with seven setae (none distinctly spatulate) and apical acrothek consisting of short aesthetasc (L: 12 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[7 + acrothek]. + + +Antenna ( +Fig. 9B, C +). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment forming incompletely fused allobasis, about 2.5 times as long as maximum width; original basis-endopod boundary marked by short transverse surface suture at level of exopodal articulation and by constriction at abexopodal side; proximal part representing original basis with single spinules at distal corner. Exopod one-segmented, unornamented and elongate, with a naked apical seta (about 1.2 times longer than exopod). Free endopodal segment with few lateral spinules proximally and transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in +Fig. 9B +); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta. + + +Mandible, maxillule, maxilla and maxilliped as in + +N. spinicaudatus + +(see + +Sak +et al. +2008 + +: +Figs 16D, E +; +17E, F +). + + +P1 ( +Fig. 9D +). Intercoxal sclerite wide and subrectangular. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis without spinular ornamentation; anterior surface with small naked seta near medial margin. Exopod two-segmented; segments with several spinules around outer margin; exp-1 unarmed; exp-2 with unipinnate outer spine and two geniculate setae distally. Endopod two-segmented, not prehensile, slightly longer than exopod; enp-1 about 1.4 times longer than enp-2, with a serrate inner seta arising from distal third and four coarse spinules along outer margin; enp-2 without spinules, distal margin with naked outer spine and geniculate inner seta. + + +P2–P4 ( +Fig. 9F–G +). Intercoxal sclerites rectangular with deeply concave ventral margins. Praecoxae triangular, small and naked. Coxae wider than long and without ornamentation. Bases smaller than coxae; anterior surface with a pore; outer basal seta absent in P2, naked and long (P3) or short (P4) in remaining legs; without ornamentation except for spinular row near base of P4 endopod. Exopods three-segmented; segments with coarse spinular ornamentation along outer margin, as illustrated; outer spine of segments naked (except for P2 exp-2 sparsely unipinnate); exp-3 with an outer unipinnate spine, and two setae distally (both naked in P2, one unipinnate and one bipinnate in P3–P4); outer spine of P2 exp-2 very long, extending far beyond exp-3 and about as long as entire exopod; P4 exp-2 elongate, distinctly longer than exp-1; segments of P4 about equally long, exp-3 with inner serrate seta in distal quarter originating from posterior surface. Endopods one- (P2–P3) or two-segmented (P4); P2–P3 endopods with few coarse spinules along both inner and outer margin; P4 enp-1 elongate, 3.7 times as long as enp-2, with few spinules along outer margin; P4 enp-2 without ornamentation; P2 endopod with long, apically serrate, backwardly directed seta along inner margin and one long unipinnate seta apically; P3 endopod shorter than P2 endopod, with short, naked spine apically; apical margin of P4 enp-2 with long, distally serrate and basally fused, inner seta, and short naked outer seta. Spine and seta formula as follows: + + +
 + + +FIGURE 9. + +Neoleptastacus supersetosus + +sp. nov. + +(♀): (A) habitus, lateral; (B) antenna, medial [lateral endopodal spines indicated by arrows]; (C) antennary endopod, outer [lateral spines omitted]; (D) P1, anterior; (E) P2, anterior [outer spine of exp-1 broken]; (F) P3, anterior; (G) P4, anterior. + + + + +FIGURE 10. + +Neoleptastacus supersetosus + +sp. nov. + +: (A) urosome (♀), ventral [arrow indicating copulatory pore]; (B) urosome (♂), ventral. + + + + +FIGURE 11. + +Neoleptastacus supersetosus + +sp. nov. + +: (A) habitus (♂), lateral; (B) rostrum and cephalothorax (♀), dorsal; (C) anal somite and caudal rami (♀), dorsal; (D) anal somite and left caudal ramus (♀), lateral; (E) antennule (♀), dorsal; (F) antennule (♂), dorsal. + +
+ + + + + + + + + + + + + + + + + + + +
ExopodEndopod
P20.0.021110
P30.0.021010
P40.0.1210.020
+ + +Fifth legs ( +Fig. 10A +) closely set together, but not touching in ventral midline. Baseoendopod and exopod fused, forming a subrectangular plate; with one pore on anterior surface and two pores along inner margin; inner distal corner with strong, bipinnate, spinous process (homologous to inner spine); distal margin with a minute outer seta, one long naked seta, and two short, subequally long (outer one slightly longer), naked spines; outer basal seta long and naked. + + +Sixth pair of legs ( +Fig. 10A +) reduced, fused medially forming common genital operculum; armature represented by vestigial spiniform element on either side. Copulatory pore conspicuous (arrowed in +Fig. 10A +). + + +Description of male. +Total body length from tip of rostrum to posterior margin of caudal rami 235 µm ( +n += 1). Body ornamentation essentially as in female except for internal semicircular reinforcements present only on somites bearing P2–P4 ( +Fig. 11A +). Sexual dimorphism in antennule, genital segmentation and P6. Spermatophore length approximately 33 µm. + + +Antennule ( +Fig. 11F +) 9-segmented, haplocer; geniculation between segments 7 and 8. Segment 1 with a slender naked seta; segment 2 longest and about 2.4 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae and a spine; segment 4 an incomplete sclerite with one seta and one spiniform element; segment 5 with a long aesthetasc (40 µm) fused basally to slender seta; segment 6 with a seta; segment 7 with a seta and a spine; segment apparently unarmed; distal segment with six setae (none of which spatulate) and apical acrothek consisting of short aesthetasc (10 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4 + 1 spine], 4-[(1 + 1 spine)], 5-[1 + (1 + ae)], 6-[1], 7-[1 + 1 spine], 8-[0], 9-[6 + acrothek]. + + +Sixth legs ( +Fig. 10B +) slightly asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer seta and a short inner spine, both elements being naked. + + + + +Etymology. +The specific epithet is derived from the Latin +super +, meaning over, and +s(a)eta, s(a)etosus +, meaning bristle, bristly, and refers to the supernumerary element on the P5 of both sexes. + + + + +Remarks. + +Neoleptastacus supersetosus + + +sp. nov. + +is assigned to the + +trisetosus + +-group based on (1) the absence of paired spinous processes on the anal somite, (2) a well-developed anal operculum, (3) P1 exp-1 without outer spine and exp-2 with three setae/spines, (4) P2 exp-2 with very long outer setiform element extending far beyond distal margin of exp-3, (5) endopod P2–P3 1-segmented with inner seta and one distal spine on P2 and one distal spine on P3, and (6) outer seta of P4 enp-2 reduced. Within this group it is morphologically closest to + +N. trisetosus + +and + +N. panamensis + + +sp. nov. + +with which it shares the 2-segmented condition of the P1 exopod. Characters that differentiate the new species from these two congeners include (1) somites bearing P2–P5, genital double-somite and second abdominal somite with internal semicircular reinforcements in +♀ +, and on somites bearing P2–P +4 in +♂ +, (2) inner spine of P5 fused to segment, forming spinous process in both sexes ( +vs +delimited at base), and (3) male P6 with outer seta and inner spiniform element ( +vs +outer seta only). An additional character that differentiates + +N. supersetosus + + +sp. nov. + +readily from all known species of + +Neoleptastacus + +(and in fact all other +Arenopontiidae +) is the presence of a supernumerary setiform element on the P5 of both sexes. + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F74BFF91FF4EF98E79FCF965.xml b/data/62/7E/C6/627EC678F74BFF91FF4EF98E79FCF965.xml index ca8352d626d..aa339c18d06 100644 --- a/data/62/7E/C6/627EC678F74BFF91FF4EF98E79FCF965.xml +++ b/data/62/7E/C6/627EC678F74BFF91FF4EF98E79FCF965.xml @@ -1,71 +1,76 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E - + -Key to species + +Key to +species + + + Redefinition of the generic boundaries in the @@ -96,48 +101,93 @@ and (both species inquirendae ) are included in -Table 2 +Table 2 but not in the updated key below. Identifications made with this key must be confirmed by reference to the original descriptions in the literature. -1. Anal somite with paired dorsolateral spinous spurs or processes................................................2. -Anal somite without paired dorsolateral spurs or processes...................................................13. 2. Abdominal somites with integumental pattern of rectangular plates dorsally and ventrally; P2 enp-2 without inner seta.....3. - -Abdominal somites without conspicuous surface sculpturing; P2 enp-2 with inner seta..............................4. 3. P3 enp-2 with two distal elements; P5 of both sexes with four articulating setae/spines................... + + + + + +1. Anal somite with paired dorsolateral spinous spurs or processes................................................2. + + +Anal somite without paired dorsolateral spurs or processes...................................................13. + + + + +2. Abdominal somites with integumental pattern of rectangular plates dorsally and ventrally; P2 enp-2 without inner seta.....3. + + +Abdominal somites without conspicuous surface sculpturing; P2 enp-2 with inner seta..............................4. + + + + + +3. P3 enp-2 with two distal elements; P5 of both sexes with four articulating setae/spines................... N. ornamentus . + - + + P3 enp-2 with one distal element; P5 of both sexes with three articulating setae/spines................... N. reductaspina . -4. P3 enp-2 with two distal elements........................................................................5. + -P3 enp-2 with one distal element.........................................................................7. - + + + +4. P3 enp-2 with two distal elements........................................................................5. + + +P3 enp-2 with one distal element.........................................................................7. + + + + + 5. P1 exp-3 with three setae/spines.................................................................... N. huysi . -P1 exp-3 with four setae/spines..........................................................................6. + - + +P1 exp-3 with four setae/spines..........................................................................6. + + + + + 6. P1 endopod clearly longer than exopod; P3 enp-2 about half the length of enp-1; P5 of both sexes with one seta and two short spines between inner spinous process and outer basal seta............................................ N. acanthus . + + + + P1 endopod as long as exopod; P3 enp-2 only slightly shorter than enp-1; P5 of both sexes with one seta and one spur between inner spinous process and outer basal seta...................................................... N. chaufriassei . + - + + + + 7. Sternal plates of somites bearing P1–P4 with midventral hook-like processes..................... N. emendatus @@ -145,10 +195,23 @@ P1 endopod as long as exopod; P3 enp-2 only slightly shorter than enp-1; P5 of b sp. nov. -Sternal plates of somites bearing P1–P4 without such posteriorly directed processes................................8. + -8. Paired spinous processes on anal somite straight and backwardly directed.........................................9. Paired spinous processes on anal somite dorsally recurved....................................................10. - + +Sternal plates of somites bearing P1–P4 without such posteriorly directed processes................................8. + + + + +8. Paired spinous processes on anal somite straight and backwardly directed.........................................9. + + +Paired spinous processes on anal somite dorsally recurved....................................................10. + + + + + 9. P1 endopod 1.15 times as long as exopod; P5 elongate in both sexes, length/maximum width ratio 3.0 ( ♀ ) and 3.5 ( @@ -157,7 +220,12 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc N. longiremis -. P1 endopod 1.30 times as long as exopod; P5 much shorter in both sexes, length/maximum width ratio 1.7 ( +. + + + + +P1 endopod 1.30 times as long as exopod; P5 much shorter in both sexes, length/maximum width ratio 1.7 ( ♀ ) and 2.25 ( ♂ @@ -168,8 +236,12 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc sp. nov. + - + + + + 10. P1 endopod shorter than exopod........................................................ N. abbreviatus @@ -177,9 +249,15 @@ Sternal plates of somites bearing P1–P4 without such posteriorly directed proc sp. nov. -P1 endopod at least as long as exopod....................................................................11. + - + +P1 endopod at least as long as exopod....................................................................11. + + + + + 11. P1 endopod as long as exopod; spinous process on caudal ramus relatively short, about 35% of ramus length.................................................................................................. N. chilensis @@ -187,28 +265,55 @@ P1 endopod at least as long as exopod........................................... sp. nov. -P1 endopod 1.25–1.30 times as long as exopod; spinous process on caudal ramus longer, about 45–55% of ramus length..12. + - + +P1 endopod 1.25–1.30 times as long as exopod; spinous process on caudal ramus longer, about 45–55% of ramus length..12. + + + + + 12. P1 enp-1 1.25 times as long as enp-2; P 5 ♀ twice as long as maximum width; terminal process of caudal ramus dorsally recurved.................................................................................... N. gussoae -. P1 enp-1 1.65 times as long as enp-2; P +. + + + + +P1 enp-1 1.65 times as long as enp-2; P 5 ♀ 2.7 times as long as maximum width; terminal process of caudal ramus not dorsally recurved.................................................................................... N. indicus . + -13. P1 exopod 2-segmented...............................................................................14. P1 exopod 3-segmented...............................................................................16. - + + + +13. P1 exopod 2-segmented...............................................................................14. + + +P1 exopod 3-segmented...............................................................................16. + + + + + 14. P5 inner spine at least partly delimited at base; P 6 ♂ -with one seta.............................................15. P5 inner spine fused at base, forming spinous process; P +with one seta.............................................15. + + + + +P5 inner spine fused at base, forming spinous process; P 6 ♂ with two elements.................. @@ -217,30 +322,56 @@ with two elements.................. sp. nov. + - + + + + 15. P2 exp-2 outer spine shorter than exopod; P2 endopod shorter than exp-1, flask-shaped; P4 exp-3 four times as long as maximum width, with inner seta arising from distal fifth of inner margin......................................... N. trisetosus -. P2 exp-2 outer spine distinctly longer than exopod; P2 endopod as long as exp-1, subrectangular and slightly tapering in distal third; P4 exp-3 2.75 times as long as maximum width, with inner seta arising from middle third of inner margin............................................................................................ +. + + + + +P2 exp-2 outer spine distinctly longer than exopod; P2 endopod as long as exp-1, subrectangular and slightly tapering in distal third; P4 exp-3 2.75 times as long as maximum width, with inner seta arising from middle third of inner margin............................................................................................ N. panamensis sp. nov. + -16. P3 endopod 1-segmented, with one distal element..........................................................17. P3 endopod 2-segmented, enp-2 with two elements (outer one fused at base).....................................19. - + + + +16. P3 endopod 1-segmented, with one distal element..........................................................17. + + +P3 endopod 2-segmented, enp-2 with two elements (outer one fused at base).....................................19. + + + + + 17. P1 exp-3 with four elements; P2 endopod 2-segmented; P4 exp-3 without inner seta....................... N. australis . -P1 exp-3 with three elements; P2 endopod 1-segmented; P4 exp-3 with inner seta.................................18. + - + +P1 exp-3 with three elements; P2 endopod 1-segmented; P4 exp-3 with inner seta.................................18. + + + + + 18. Inner spinous process of P 5 ♀ distinctly curved outwardly (unknown in @@ -252,6 +383,10 @@ distinctly curved outwardly (unknown in comb. nov. + + + + Inner spinous process of P 5 ♀ / @@ -261,41 +396,77 @@ and terminal process of caudal ramus straight........................... N. africanus . + - + + + + 19. P4 exp-3 without inner seta.................................................................... N. pacificus . -P4 exp-3 with inner seta...............................................................................20. + - + +P4 exp-3 with inner seta...............................................................................20. + + + + + 21. Urosome (except anal somite) with distinct surface ornamentation consisting of elongate rectangular plates............................................................................. N. clasingi . -Urosome without conspicuous surface ornamentation........................................................22. + -22. Caudal ramus with dorsolateral spur near base of seta VII....................................................23. Caudal ramus without dorsolateral spur near base of seta VII..................................................24. - + +Urosome without conspicuous surface ornamentation........................................................22. + + + + +22. Caudal ramus with dorsolateral spur near base of seta VII....................................................23. + + +Caudal ramus without dorsolateral spur near base of seta VII..................................................24. + + + + + 23. Lappets of abdominal hyaline frills semi-incised obtusidigitate; inner seta of P2–P3 enp-2 longer than endopod; P5 three times as long as wide, with naked spinous process........................................................ N. spicatus -. Lappets of abdominal hyaline frills denticulate; inner seta of P2–P3 enp-2 shorter than endopod; P5 about 2.5 times as long as wide, with pinnate spinous process.......................................................... +. + + + + +Lappets of abdominal hyaline frills denticulate; inner seta of P2–P3 enp-2 shorter than endopod; P5 about 2.5 times as long as wide, with pinnate spinous process.......................................................... N. spinicaudatus . + - + + + + 24. P1 endopod distinctly longer than exopod...................................................... N. ishikarianus . + + + + P1 rami equally long............................................................ N. pseudishikarianus @@ -303,7 +474,10 @@ P1 rami equally long............................................................ sp. nov. + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F74DFF94FF4EFD927858F97B.xml b/data/62/7E/C6/627EC678F74DFF94FF4EFD927858F97B.xml new file mode 100644 index 00000000000..3abf936571c --- /dev/null +++ b/data/62/7E/C6/627EC678F74DFF94FF4EFD927858F97B.xml @@ -0,0 +1,297 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Neoleptastacus accraensis +( +Lang, 1965 +) + + + + + + + + + + +Arenopontia longiremis +Chappuis, 1955 +sensu +Chappuis & Rouch (1961: 608) + + + + + + +Arenopontia accraensis +Lang, 1965 + + + + +Arenopontia +( +Neoleptastacus +) +accraensis +Lang, 1965 + +: + +Wells (1967: 324 + +—spelled + +accraensis + +) + + + + +Arenopontia +( +Neoleptastacus +) +acantha accraensis +Lang, 1965 + +: + +Kunz (1971: 356) + + + + + + + +Neoleptastacus accraensis +( +Lang, 1965 +) + +Sak +et al. +(2008: 412) + + + + + + +Original description. +Chappuis & Rouch (1961 +—as + +A. longiremis + +): 607–608; +Figs 9–13 +( +♀ +only). + + + + + +Type +locality. + +Ghana +, +Greater Accra Region +, +Accra +; sandy beach in front of the lighthouse. + + +Body length. +Unknown. + + + + +Remarks. +Chappuis & Rouch (1961: 608 +, +Figs 9–13 +) recorded +two females +from a sandy beach in Accra which they assigned to + +Arenopontia longiremis +Chappuis, 1955 + +. They considered the differences with the +type +population as insignificant, being a likely reflection of the disjunct distribution of the species. +Lang (1965) +considered +Chappuis & Rouch’s (1961) +Ghanaian specimens to fall outside the limits of variability of + +A. longiremis + +and proposed a new name, + +A. accraensis + +, for them. +Kunz (1971) +treated + +A. longiremis + +and + +A. accraensis + +as subspecies of + +A. acantha + +but this was not adopted by subsequent revisers [ +Itô 1978 +; +Bodiou & Colomines 1986 +; +Wells 2007 +; + +Sak +et al. +2008 + +—but see +Bodin (1997) +for a dissenting opinion]. The Angolan material identified by +Kunz (1971) +as + +A. +( +N. +) +acantha accraensis + +belongs to a different species, described here as + +N. emendatus + + +sp. nov. + +(see above). + + +Although +Lang (1965) +, +Bodiou & Colomines (1986) +and +Wells (2007) +claim that P4 exp-3 possesses an inner seta, the setation of this leg has yet to be revealed. Similarly, in the identification keys provided by these authorities and +Karanovic (2000) +it is assumed that the distal endopod segment of P3 has only one distal element. However, +Chappuis & Rouch (1961) +show a second short element on enp- +2 in +their illustration of the P3 endopod. This element is here interpreted as the outer distal spine using the spinule row on enp-1 as a reference point for correct orientation. Note also that +Chappuis & Rouch’s (1961) +illustration of the P1 shows it in an unnatural twisted position and that the terminal spinous process on the caudal ramus was figured as an articulating spine. The latter condition is unlikely since the caudal process in all members of the +Arenopontiidae +represents a posterior outgrowth of the ramus and not the homologue of an armature element that became secondarily incorporated. + + + +Neoleptastacus accraensis + +is in need of detailed redescription (including the as yet unknown male) and is treated here as a +species inquirenda +in the genus since it is practically unidentifiable based on the few drawings provided by +Chappuis & Rouch (1961) +. It is tentatively placed in the + +spinicaudatus + +-group based on the combined absence of paired anal processes and the retention of the plesiomorphic armature formula of P1–P4. The species is only known from the +type +locality where it occurs sympatrically with + +N. africanus + +. + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F753FF83FF4EF89B7E45FD42.xml b/data/62/7E/C6/627EC678F753FF83FF4EF89B7E45FD42.xml new file mode 100644 index 00000000000..3c723f283cf --- /dev/null +++ b/data/62/7E/C6/627EC678F753FF83FF4EF89B7E45FD42.xml @@ -0,0 +1,563 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Phreatipontia speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + +comb. nov. + + + + + + +( +Figs 16–19 +) + + + + + +Arenopontia +( +Neoleptastacus +) +speluncae +Cottarelli, Bruno & Venanzetti, 1994 + + + + + +Neoleptastacus turcicus +—nomen nudum + +by + +Sak (2004: 221) + + + + + + + +Neoleptastacus speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + +Sak +et al. +(2008: 412) + + + +Original +description. + + +Cottarelli +et al. +(1994) + +: 475 + +–480; +Figs 2–3 +. + + + + + + + +Type locality. +Italy +, +Lazio +, +Latina Province +, +Sperlonga +, +Tiberio +beach; on the banks of a little stream that originates from a spring on the beach; interstitial water. +Note +that + +Cottarelli +et al. +(1994) + +also collected +two females +from a second locality in the +Latina Province +( +S. Agostino +beach in +Gaeta +) but did not explicitly state where the female +holotype +originated from. Since they allocated numbers 2–13 to the +paratypes +( +six ♀♀ +and +five ♂♂ +in total were collected from Sperlonga, +two ♀♀ +from +Gaeta +) it appears rational to assume that number 1 was given to the +holotype +and that it came from +Sperlonga. The +etymology of the species name and the fact that additional drawings ( +Figs 2–e +, +3–g +) were presented of an atypical female of +S. Agostino +beach also seem to suggest that +Sperlonga +was the intended type locality. + + +Material examined. + +One ♀ +(dissected on eight slides) and +one ♂ +(dissected one seven slides); 10 individuals in ethanol. +All +specimens collected from +Ismailbeyli Village +beach ( +41°02.198’ N +; +38°56.901’ E +) close to a small stream, Giresun ( +Giresun Province +), northeastern +Türkiye +( +Black Sea region +); leg. +S. Karaytuğ +& +S. Sak +, + +10 September 2002 + + +. + + +Body length. +394 μm ( +♀ +), 346 μm ( +♂ +) [Cottarelli +ei al. +1994]; 356 μm ( +♀ +), 338 μm ( +♂ +) [present account]. + + + + +Redescription of female. +Total body length from tip of rostrum to posterior margin of caudal rami 356 μm ( +n += 1). Cephalothorax maximum width 42 μm measured at posterior margin. Maximum body width 47 μm, measured at P5-bearing somite. Body ( +Fig. 16A, B +) slender and cylindrical without clear distinction between prosome and urosome. Sensillar pattern on body as figured. Hyaline frills of thoracic somites weakly developed and plain; those of genital double-somite and free abdominal somites well developed and consisting of rectangular digitate lappets that taper towards their distal margins ( +Figs 16A–B +; +19A +). Somites connected by well-developed intersomitic membranes. Genital double-somite slightly longer than wide (measured in dorsal aspect); with two ventral pores ( +Fig. 19A +). Anal somite ( +Figs 16C, D +; +19A +) with two dorsal and four lateral pores. Anal frill triradiate, minutely incised (giving a spinulose appearance); anal operculum virtually straight, without ornamentation. + + +Caudal rami ( +Figs 16C, D +; +19A +) about 3.4 times longer than wide (measured in dorsal view from anterior margin to apex of spinous process), distinctly tapering posteriorly; outer and inner margins slightly convex; with two ventral pores in proximal third and two lateral pores near insertion of seta III; terminal spinous process slightly recurved dorsally; no spinular ornamentation discernible. Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, located between seta V and posterior spinous process, with long outer spinule; seta V with proximal fracture plane; seta VI small, naked and located at inner distal corner; seta VII spatulate and tri-articulate at base. + + +Rostrum ( +Fig. 16A +) small, subtriangular, tapering distally; with two delicate sensilla. + + +Antennule ( +Fig. 17C +) long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about 2.5 times longer than wide. Segment 4 with long aesthetasc (L: 29 μm) fused at base with seta. Distal segment with eight setae (none of them spatulate) and apical acrothek consisting of short aesthetasc (L: 13 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[8 + acrothek]. + + +Antenna ( +Fig. 17D +). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment completely separated. Basis with few spinules along exopodal margin and with oblique spinular row near base of exopod. Exopod one-segmented and elongate, with a naked apical seta (about 1.4 times longer than exopod). Free endopodal segment with transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in +Fig. 17D +); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta. + + +Mandible, maxillule, maxilla and maxilliped as in + +P. phreatica + +. + + +P1 ( +Fig. 18A +). Intercoxal sclerite wide, naked and subrectangular, with concave ventral margin. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis with spinular row near base of endopod; anterior surface with a small setiform spine near medial margin and one pore near articulation with coxa. Exopod three-segmented; all segments with several spinules around outer margin; exp-1 about as long as exp-3, with naked outer spine; exp-2 without outer element; exp-3 with short naked outer spine, and a longer naked spine and two geniculate setae distally. Endopod two-segmented, not prehensile, slightly longer than exopod; enp-1 with virtually straight inner margin, about 1.5 times longer than enp-2, with a serrate seta arising from about halfway down inner margin and two sets of two spinules along outer margin; enp-2 without spinules, distal margin with two geniculate setae, outermost of which shortest. + + + +FIGURE 16. + +Phreatipontia speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + + +comb. nov. + +(♀): (A) habitus, dorsal; (B) habitus, lateral; (C) anal somite and caudal rami, dorsal [arrow indicating modified seta IV]; (D) anal somite and right caudal ramus, lateral. + + + + +FIGURE 17. + +Phreatipontia speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + + +comb. nov. + +: (A) habitus (♂), dorsal; (B) antennule (♂), ventral; (C) antennule (♀), ventral; (D) antenna (♀), medial [lateral endopodal spines indicated by arrows]. + + + + +FIGURE 18. + +Phreatipontia speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + + +comb. nov. + +(♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. + + + + +FIGURE 19. + +Phreatipontia speluncae +( +Cottarelli, Bruno & Venanzetti, 1994 +) + + +comb. nov. + +: (A) urosome (♀), ventral; (B) urosome (♂), ventral. + + + +P2–P4 ( +Fig. 18B–D +). Intercoxal sclerite naked (P2–P4), rectangular (P2) or squarish (P3–P4) with deeply concave ventral margins. Praecoxae triangular, small and naked. Coxae squarish (P3) or slightly wider than long (P2, P4) and without ornamentation. Bases smaller than coxae, with a spinular row near base of endopod (P2–P3) and a few spinules around outer corner in P2 and P4; anterior surface with a pore in P3-P4; outer basal seta absent in P2, naked in P3–P4. Exopods three-segmented; segments with coarse spinular ornamentation along outer margin; outer spine of exp-1 and exp-2 naked; exp-3 with an outer unipinnate spine and two bipinnate setae distally, P4 exp-3 with additional serrate seta on inner margin; P4 exp-2 only marginally longer than exp-1. Endopods two-segmented; P2–P4 enp-1 unarmed, about 1.2, 1.5, and 4.2 times longer than their respective distal segments, with few coarse spinules along outer margin as figured, but without ornamentation along inner margin; P2–P3 enp-2 with few spinules around distal margin; P2 enp-2 with long, apically serrate, backwardly directed seta near proximal margin and one unipinnate setae apically; P3 enp-2 with long, bipinnate apically; distal margin of P4 enp-2 with long, distally serrate and basally fused, inner seta, and shorter unipinnate, outer seta. Spine and seta formula as follows: + + + + + + + + + + + + + + + + + + + + + + +
ExopodEndopod
P20.0.0210.110
P30.0.0210.010
P40.0.1210.020
+ + +Fifth legs ( +Fig. 19A +) closely set together, almost touching medially. Baseoendopod and exopod fused, forming a subrectangular plate with straight distal margin; with one pore on anterior surface; inner distal corner with long, bipinnate spinous process (homologous to inner spine); distal margin with long naked outer seta and two short equally long, bipinnate spines; outer basal seta long and plumose. + +Sixth legs vestigial, forming opercula closing off genital apertures. + +Redescription of male. +Total body length from tip of rostrum to posterior margin of caudal rami 338 µm ( +n += 1). Body ornamentation essentially as in female ( +Fig. 17A +). Sexual dimorphism in antennule, genital segmentation, P5, and P6. Spermatophore length approximately 70 μm. + + +Antennule ( +Fig. 17B +) 8-segmented, haplocer; geniculation between segments 6 and 7. Segment 1 with a naked seta; segment 2 longest and about 2.7 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae; segment 4 an incomplete sclerite with two setae; segment 5 with four setae, one pinnate spine and a long aesthetasc (43 µm) fused basally to a slender seta; segments 6–7 with one seta each; distal segment with seven setae (none of which spatulate) and apical acrothek consisting of short aesthetasc (16 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[2], 5-[4 + (1 + ae)], 6-[1], 7-[1], 8-[7 + acrothek]. + + +P5 ( +Fig. 19B +) essentially as in female but inner spinous process slightly more slender. + + +Sixth legs ( +Fig. 19B +) asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer plumose seta and a shorter inner bipinnate seta. + + + + +Remarks. +The report of an inner seta on P3 exp-2 by + +Cottarelli +et al. +(1994) + +is extremely doubtful and requires confirmation. No other member in the family +Arenopontiidae +displays an inner seta on this segment. Our specimens from the Black Sea agree in virtually all other morphological aspects with Cottarelli +et al. +’s (1994) original description, adding further credence to the notion that their illustration of P3 is based on an observational error. The presence of an outer basal seta on P2 (their +Fig. 3–e +), their statement that the somitic hyaline frills are plain, and the absence of spinular ornamentation on the spinous process of P5 are also questionable and require further examination. + + + +Cottarelli +et al. +(1994) + +did not observe any variability among the individuals from Sperlonga but illustrated the P5 and caudal ramus of a female from S. Agostino beach without further comment. The fifth leg of the latter deviates from that of the +holotype +in the shorter spinous process, the longer outer marginal seta (extending well beyond the process) and the general appearance of the two medial elements which are unequal in length and setiform in shape. The dorsal view of the caudal ramus is difficult to compare with the +holotype +since it was only illustrated in lateral aspect for the latter. + + +The two described species of + +Phreatipontia + + +gen. nov. + +are morphologically very similar. The +type +species, + +P. speluncae + + +comb. nov. + +, can primarily be differentiated from + +P. phreatica + + +comb. nov. + +by morphometric differences in the caudal rami and swimming legs. The exopodal segments of P2–P3 are distinctly more slender and longer in the latter, P2 enp-2 is markedly shorter (enp-1:enp-2 length ratio 1.6 +vs +1.2) while P4 enp-1 is longer (enp-1:enp-2 length ratio 5.8 +vs +4.2). The caudal rami are generally more slender and longer (L:W ratio 3.9 +vs +3.4) in + +P. phreatica + + +comb. nov. + +while the dorsal seta VII is more foliaceous in + +P. speluncae + + +comb. nov. + + + +In addition to the +type +locality, + +P. speluncae + + +comb. nov. + +was also obtained in interstitial freshwater of S. Agostino beach (Gaeta, Latina Province, Lazio) on the banks of a little stream fed by a spring in Grotta del Serpente ( + +Cottarelli +et al. +1994 + +) and from the mouths of the Fiora River and Valfragida Stream, in the Viterbo Province (Lazio) ( + +Cottarelli +et al. +1998 + +; +Berera 2000 +). +Cottarelli & Berera (2004) +also recorded it from the banks of the Fiora River near Montalto di Castro. Our material extends its distribution further into the Black Sea basin. + +Cottarelli +et al. +(1994) + +suggested that the species may be restricted to fresh or slightly brackish water but admitted that more data are needed before this can be corroborated. It is conceivable that + +P. speluncae + + +comb. nov. + +(and + +P. phreatica + + +comb. nov. + +) is in the process of colonizing continental subterranean waters ( + +Bruno +et al. +1998 + +). A third species, apparently close to + +P. speluncae + + +comb. nov. + +and still under study, was discovered in freshwater habitats in +Greece +( + +Bruno +et al. +1998 + +; +Bruno & Cottarelli 1999 +). + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F75AFF83FF4EFA1279F9F82D.xml b/data/62/7E/C6/627EC678F75AFF83FF4EFA1279FDF82D.xml similarity index 63% rename from data/62/7E/C6/627EC678F75AFF83FF4EFA1279F9F82D.xml rename to data/62/7E/C6/627EC678F75AFF83FF4EFA1279FDF82D.xml index b4a1ffdcff2..93e31f3fb88 100644 --- a/data/62/7E/C6/627EC678F75AFF83FF4EFA1279F9F82D.xml +++ b/data/62/7E/C6/627EC678F75AFF83FF4EFA1279FDF82D.xml @@ -1,90 +1,105 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E - + Key to genera of -Arenopontiidae +Arenopontiidae - + + + + 1. P1 endopod prehensile, enp-1 distinctly longer than exopod; innermost element of P1 exp-3 penicillate.................................................................................................. Arenopontia Kunz, 1937 . + -P1 endopod not prehensile, enp-1 not longer than exopod; innermost element of P1 exp-3 geniculate...................2. -2. P5 of both sexes with innermost element forming a distinct spinous process which is incorporated in the segment.........3. -P5 of both sexes with innermost element not modified (occasionally fused at base).................................7. + +P1 endopod not prehensile, enp-1 not longer than exopod; innermost element of P1 exp-3 geniculate...................2. + + + + +2. P5 of both sexes with innermost element forming a distinct spinous process which is incorporated in the segment.........3. + + +P5 of both sexes with innermost element not modified (occasionally fused at base).................................7. + + + + 3. Distal margin of P1 enp-2 with two geniculate setae........................................ Phreatipontia @@ -92,46 +107,70 @@ Key to genera of gen. nov. + + Distal margin of P1 enp-2 with outer naked spine and inner geniculate seta............ Neoleptastacus Nicholls, 1945 .......4. + + + + 4. Anal somite with paired spinous processes either side of anal operculum............................. acanthus -group. + -Anal somite without paired spinous processes either side of anal operculum.......................................5. + +Anal somite without paired spinous processes either side of anal operculum.......................................5. + + + + 5. P1 exp-1 without outer spine; distal segment of P1 exopod with three elements; P2 endopod 1-segmented... trisetosus -group. + -P1 exp-1 with outer spine; distal segment of P1 exopod with four elements; P2 endopod 2-segmented..................6. + +P1 exp-1 with outer spine; distal segment of P1 exopod with four elements; P2 endopod 2-segmented..................6. + + + + 6. Distal segment of P3 endopod with one apical element............................................ australis -group. + + Distal segment of P3 endopod with two apical elements....................................... spinicaudatus -group. + + + + 7. P1 enp-2 with one geniculate seta and one spine; P3 endopod not modified in ♂ .. @@ -140,9 +179,15 @@ Distal segment of P3 endopod with two apical elements........................... Sak, Huys & Karaytuğ, 2008 . + -P1 enp-2 with two geniculate setae; P3 endopod sexually dimorphic.............................................8. + +P1 enp-2 with two geniculate setae; P3 endopod sexually dimorphic.............................................8. + + + + 8. P2 enp-2 with inner serrate seta; P3 endopod ♂ two-segmented...................... @@ -151,8 +196,10 @@ two-segmented...................... Pennak, 1942 . + + P2 enp-2 without inner serrate seta; P3 endopod ♂ one-segmented............ @@ -161,7 +208,10 @@ one-segmented............ Sak, Huys & Karaytuğ, 2008 . + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F761FFB7FF4EF9427DE5FEA2.xml b/data/62/7E/C6/627EC678F761FFB7FF4EF9427DE5FEA2.xml index 7a02323c446..408cf66b6f6 100644 --- a/data/62/7E/C6/627EC678F761FFB7FF4EF9427DE5FEA2.xml +++ b/data/62/7E/C6/627EC678F761FFB7FF4EF9427DE5FEA2.xml @@ -1,71 +1,71 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E (3) - + australis -group @@ -74,7 +74,7 @@ Diagnosis. -Anal somite without paired dorsolateral processes. Anal operculum weakly developed, without rounded medial extension. P1 exp-1 with outer spine; exp-3 with four setae/spines. P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp- 3). P2 endopod 2-segmented; enp-2 with inner seta and two distal spines. P3 endopod 1-segmented; with one distal spine. P4 enp-2 with normally developed outer seta. +Anal somite without paired dorsolateral processes. Anal operculum weakly developed, without rounded medial extension. P1 exp-1 with outer spine; exp-3 with four setae/spines. P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp-3). P2 endopod 2-segmented; enp-2 with inner seta and two distal spines. P3 endopod 1-segmented; with one distal spine. P4 enp-2 with normally developed outer seta. @@ -104,7 +104,7 @@ This species group includes one incompletely described African species which doe N. australis ( -Table 2 +Table 2 ). More information is required before any statement can be made about its relationships, particularly whether it may be nested as an advanced member in the spinicaudatus diff --git a/data/62/7E/C6/627EC678F761FFB8FF4EFBB27EC4F97A.xml b/data/62/7E/C6/627EC678F761FFB8FF4EFBB27EC4F97A.xml index a589e8f9da2..a723dfddce9 100644 --- a/data/62/7E/C6/627EC678F761FFB8FF4EFBB27EC4F97A.xml +++ b/data/62/7E/C6/627EC678F761FFB8FF4EFBB27EC4F97A.xml @@ -1,71 +1,71 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E - + (2) - + spinicaudatus -group @@ -139,7 +139,7 @@ The N. pacificus which lacks the inner seta on P4 exp-3 ( -Table 2 +Table 2 ). In the absence of any morphology-based apomorphies that could support the common origin of these species, assessment of the potential monophyly of this species group will have to await the arrival of molecular sequence data. Sak diff --git a/data/62/7E/C6/627EC678F763FFB8FF4EF94F7F2BFC2A.xml b/data/62/7E/C6/627EC678F763FFB8FF4EF94F7F2BFC2A.xml index 5fc0e48741f..dc0dda2a7be 100644 --- a/data/62/7E/C6/627EC678F763FFB8FF4EF94F7F2BFC2A.xml +++ b/data/62/7E/C6/627EC678F763FFB8FF4EF94F7F2BFC2A.xml @@ -1,71 +1,71 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E - + (1) - + acanthus -group @@ -81,7 +81,7 @@ Anal somite with paired dorsolateral processes. Anal operculum weakly developed, ). P1 enp-2 with outer spine and inner geniculate seta distally. P2 exp-2 with outer spine of normal length (not extending far beyond distal margin of exp-3). Endopod P2–P3 2-segmented. P2 enp-2 with or without inner seta; with two distal spines. P3 enp-2 with 1–2 distal spines. P4 enp-2 with well developed outer seta. - + TABLE 1. Valid species and @@ -511,7 +511,7 @@ Venanzetti, 1994) Subgroups included. - + acanthus -subgroup, @@ -609,7 +609,7 @@ and ) ( -Table 2 +Table 2 ). Wells & Rao (1987) , inspired by the extensive variability observed in their material of @@ -743,7 +743,7 @@ and N. reductaspina share the unique absence of the inner seta on P2 enp-2 ( -Table 2 +Table 2 ), giving further credence to the monophyly of this subgroup. diff --git a/data/62/7E/C6/627EC678F763FFBAFF4EFE8E78F2F97F.xml b/data/62/7E/C6/627EC678F763FFBAFF4EFE8E78F2F97F.xml new file mode 100644 index 00000000000..ee1682bee3d --- /dev/null +++ b/data/62/7E/C6/627EC678F763FFBAFF4EFE8E78F2F97F.xml @@ -0,0 +1,284 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + +Genus + +Neoleptastacus +Nicholls, 1945 + + + + + + + +Pararenopontia +Bodiou & Colomines, 1986 + +( +type +species by original designation: + +Arenopontia breviarticulata +Mielke, 1975 + +) + + + + +Diagnosis +(adapted from + +Sak +et al. +2008 + +). +Arenopontiidae +. Urosomites occasionally with conspicuous surface ornamentation ( + +N. clasingi + +, + +ornamentus + +, + +reductaspina + +). Anal somite with ( + +acanthus + +-group) or without (all other species groups) paired dorsolateral spinous processes. Anal operculum sometimes with median extension. Hyaline frills of abdominal somites with rectangular digitate or non-digitate lappets. Caudal ramus usually with dorsolateral spur near medial margin. P1 exopod 2- or 3-segmented; exp-1 with/without outer spine; exp-3 (or exp-2 when exopod 2-segmented) with one–two spine(s) and two geniculate setae. P1 endopod not prehensile, at least as long as exopod; enp-2 with outer spine and inner geniculate seta at distal margin. P2–P3 endopods 1- or 2-segmented. P3 endopod with outer distal element (when present) usually fused at base. P4 enp-2 with well developed outer distal element (except in + +trisetosus + +-group). Armature formula as follows: + + + + + + + + + + + + + + + + + + + + + + + + + + + +
ExopodEndopod
P1 +0.0.02(1–2) or +0.021 + +1.011 +
P20.0.0210.(0–1)(1–2)0 or 110
P30.0.0210.0(1–2)0 or 010
P40.0.(0–1)210.020
+ + +P3 endopod +♂ +not sexually dimorphic. P5 with outer basal seta and one–four discrete elements; innermost element fused to segment forming spinous process (weakly delimited in + +N. trisetosus + +and + +N. panamensis + + +sp. nov. + +); length of process sometimes sexually dimorphic. P +6 ♂ +with one–two elements. + + + + + +Type +species. + + +Neoleptastacus spinicaudatus +Nicholls, 1945 + +[by monotypy]. + + + + +Other species. +Twenty-three. See +Table 1 +. + + +Species inquirenda. + +Neoleptastacus secundus +Krishnaswamy, 1957 + +; + +N. accraensis +( +Lang, 1965 +) + +. + + +Taxa of doubtful identity. + +Arenopontia acantha +Chappuis, 1954 +sensu +Božić (1967) + +; + +Arenopontia +? +gussoae +Cottarelli, 1973a +sensu +Mielke (1982b) + +[partim]; + +Arenopontia +? +gussoae +Cottarelli, 1973a +sensu +Mielke (1987) + +[2nd form]. + + + + +Gender. +Masculine. + + +The genus shows considerable variation in swimming leg armature and segmentation, P5 morphology, abdominal ornamentation and caudal ramus structure. However, all species display a spinous apical process on the P5 which is derived from the fused innermost element (weakly delimited in + +N. trisetosus + +and + +N. panamensis + + +sp. nov. + +). This apomorphic character state is unique in the +Arenopontiidae +and potentially serves as evidence in support of the monophyly of the genus. For practical reasons we have recognized several species groups within + +Neoleptastacus + +, some of which are monophyletic and may eventually be attributed separate generic status ( +Table 2 +). The majority of these groups are characterized by one or several autapomorphic character states with the notable exception of the + +spinicaudatus + +-group which contains the +type +species. For this reason, we have refrained from proposing new genera (except for the + +speluncae + +-group) as this would render + +Neoleptastacus + +a paraphyletic taxon. + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F76CFFB5FF4EFD187875F864.xml b/data/62/7E/C6/627EC678F76CFFB5FF4EFD187875F864.xml new file mode 100644 index 00000000000..d6d2c373719 --- /dev/null +++ b/data/62/7E/C6/627EC678F76CFFB5FF4EFD187875F864.xml @@ -0,0 +1,303 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Neoleptastacus spinicaudatus +Nicholls, 1945 + + + + + + + + + +Paraleptastacus spinicaudatus +Nicholls, 1945 + +: +lapsus calami +by + +Chappuis (1954: 269) + + + + + + + +Arenopontia spinicaudata +( +Nicholls, 1945 +) +Chappuis (1955: 56) + + + + + + + +Arenopontia +( +Neoleptastacus +) +spinicaudata +( +Nicholls, 1945 +) + +: + +Wells (1967: 324) + + + + + + +Neoleptastacus longipes +Nicholls, 1945 + +: +lapsus calami +by + +Karanovic (2008: 5 + +, 331) + + + +Original description. +Nicholls (1945) +: 22–23; Text fig. 3. + + + + +Additional description. + +Sak +et al. +(2008) + +: 435–441; +Figs 13–17 +. + + + + +Type locality. +Australia +, +Western Australia +. +Nicholls (1945) +did not specify a type locality but collected the species from two sandy beaches some +500 km +apart. The first, Leighton Beach (in the vicinity of Perth), was sampled in +October 1939 +while samples from the second site, locally known as ‘Back Beach’ at Dongarra, were collected in +March 1940 +. All +syntypes +deposited at the Natural History Museum in London originated from Back Beach which was designated as the type locality by + +Sak +et al. +(2008) + +. + + + + +Body length. +280–300 μm (both sexes) [ +Nicholls 1945 +]. + + + + +Remarks. + +Sak +et al. +(2008) + +re-examined the +type +material and redescribed both sexes in detail. Nicholls’s (1945) original description is adequate by contemporary standards except for the reported absence of the inner seta on P4 exp-3, an error that was perpetuated in some morphological comparisons ( +Noodt 1955b +; Bodiou & Colomines 1996) and identification keys ( +Lang 1965 +; +Karanovic 2000 +; +Wells 2007 +). Within the + +spinicaudatus + +- group, + +N. spinicaudatus + +is morphologically closest to + +N. spicatus + +, both of which exhibit a dorsolateral spur near the base of caudal ramus seta VII. The +type +species can be differentiated from + +N. spicatus + +by the shape of the abdominal hyaline frills (lappets denticulate +vs +semi-incised obtusidigitate), the proportional size of the inner distal seta on P2–P3 enp-2 (shorter +vs +longer than endopod) and the morphometric dimensions of the P5 (2.5 times as long as wide +vs +3.0 times). The apical spinous process on the P5 of both sexes is minutely bipinnate in + +N. spinicaudatus + +(and still delimited at its base on the posterior surface in the +♀ +) while it is lacking such ornamentation in + +N. spicatus + +(and completely fused to the segment in the +♀ +). The linear egg-sac typically contains 2–4 large eggs; occasionally +six eggs +are found in which case they overlap ( +Nicholls 1945 +). + + +The species is so far known only from two sandy beaches in Western Australia ( +Nicholls 1945 +). +Chappuis (1954: 269) +claimed to have found +one female +and +one male +in +Annaba +(= Bône) in northeastern +Algeria +which closely resemble + +N. spinicaudatus + +but did not present any illustrations to substantiate his record. In a later report, +Chappuis (1958: 414 +, +Figs 2–5 +) provided illustrations of the female antennule, P2 endopod, P5 and caudal ramus of specimens he collected in Puget Sound, Washington State. Despite differences in the P5, length of P3–P4 endopods and caudal rami, +Chappuis (1958) +assigned his material to + +N. spinicaudatus + +, stating that such morphological minutiae do not warrant the proposal of a distinct subspecies. As pointed out by + +Sak +et al. +(2008) + +, the morphology of the fifth legs suggests that +Chappuis (1958) +was almost certainly dealing with + +Mesopontia dillonbeachia +( +Lang, 1965 +) + +. +Karanovic (2008) +erroneously cited the species as + +Neoleptastacus longipes +Nicholls, 1945 + +. + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F76DFFACFF4EFC487E68FBBA.xml b/data/62/7E/C6/627EC678F76DFFACFF4EFC487E68FBBA.xml new file mode 100644 index 00000000000..c78d64aa9f3 --- /dev/null +++ b/data/62/7E/C6/627EC678F76DFFACFF4EFC487E68FBBA.xml @@ -0,0 +1,757 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + + + + + + + +( +Figs 1–5 +) + + + + + +Arenopontia acantha +Chappuis, 1954 + + + + + +Arenopontia +( +Neoleptastacus +) +acantha +Chappuis, 1954 + +: + +Wells (1967: 324) + + + + + + +Arenoponthia acantha +Chappuis, 1954 + +: +lapsus calami +by + +Cottarelli (1969: 20) + + + + + + +Arenopontia +( +Neoleptastacus +) +acantha acantha +Chappuis, 1954 + +: + +Kunz (1971: 356) + + + + + + + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +Sak +et al. +(2008: 412) + + + + + + + + +Neoleptastacus knidosensis +—nomen nudum + +by + +Sak (2004: 230) + + + + +Original description. +Chappuis (1954) +: 268–270; Figs 46–53. + + + + +Additional descriptions. +Masry (1970 +—as + +A. acantha + +): 251–253; +Fig. 13 +. +Cottarelli (1973a +—as + +A. acantha + +): Fig. 22 [mandibular palp]. +Martínez Arbizu & Moura (1994 +—as + +A. acantha + +): 63; +Fig. 2d, g +[rostrum, anal somite and caudal rami]. +Alper (2009) +: 45–46; +Fig. 3.6 +[ +♀ +lateral habitus, photograph]. + + + + +Type locality. +Western Mediterranean. +Chappuis (1954) +collected the species in various localities in +Italy, Algeria +and +Tunisia +(see below) but did not specify a +locus typicus +. +Since P.-A. +Chappuis habitually ignored type fixation, all the specimens on which he established + +A. acantha + +are +syntypes +and collectively constitute the name-bearing type ( +ICZN +Art. 72.1.1). According to +ICZN +Art. 73.2.3 the type locality encompasses all of the places where the +syntypes +were collected. + + +Material examined. + +One ♀ +(dissected on eight slides) (reg. no +NHMUK 2024.1050 +) + +and + +one ♂ +(dissected on seven slides) (reg. no +NHMUK 2024.1051 +) + +; + +seven ♀♀ +and +three ♂♂ +(in ethanol) (reg. nos +NHMUK 2024.1052 +– +1061 +) + +; + +five ♀♀ +and +two ♂♂ +(in ethanol) deposited in the collections of +BUZM +. +All +specimens collected from +Hastanealtı Beach +, +Datça +( +Muğla Province +), +Türkiye +on + +24 November 2002 + +; leg. +A. Alper. + + + + + +Body length. +450 μm ( +♀ +) [ +Chappuis 1954 +]; 380–420 μm ( +♀ +), 365–390 μm ( +♂ +) [ +Masry 1970 +]; 331 μm ( +♀ +), 325 μm ( +♂ +) [present account]. + + +Redescription of female. +Total body length from tip of rostrum to posterior margin of caudal rami 350 μm (mean = 331 μm; +n += 10). Maximum body width 36 μm (mean = 34 μm; +n += 10), measured near posterior margin of cephalothorax. Body slender and cylindrical without clear distinction between prosome and urosome. Sensillar pattern on body as figured. Hyaline frills of thoracic somites weakly developed and crenulated; those of genital double-somite and free abdominal somites strongly developed and consisting of rectangular digitate lappets ( +Figs 1A, B +; +2B +; +5A +). Somites connected by well-developed intersomitic membranes. Genital double-somite about 1.3 times longer than wide (measured in dorsal aspect); with two middorsal, four lateral and four ventral pores ( +Figs 1A, B +; +5A +). Anal somite ( +Figs 2B, C +; +5A +) with paired, dorsally recurved, spinous processes near posterior border either side of anal opening, and two lateral pores near anterior margin; with two conspicuous pores near ventral posterior margin. Anal frill triradiate, minutely incised (giving a spinulose appearance); anal operculum slightly convex, without ornamentation. + + + +FIGURE 1. + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +(♀): (A) habitus, dorsal; (B) habitus, lateral; (C) rostrum, dorsal. + + + + +FIGURE 2. + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +: (A) habitus (♂), dorsal; (B) anal somite and caudal rami (♀), dorsal; (C) anal somite and left caudal ramus (♀), lateral; (D) antenna (♀), medial; (E) antennary endopod (♀), outer. [Lateral spines of antennary endopod indicated by arrows in D–E] + + + + +FIGURE 3. + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +: (A) antennule (♀), dorsal; (B) antennule (♂), ventral. [spatulate setae indicated by arrows] + + + + +FIGURE 4. + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +(♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. + + + + +FIGURE 5. + +Neoleptastacus acanthus +( +Chappuis, 1954 +) + +: (A) urosome (♀), ventral [caudal ramus seta V figured at full length in inset]; (B) urosome (♂), ventral. + + + +Caudal rami ( +Figs 2B, C +; +5A +) about three times longer than wide (measured in dorsal view from anterior margin to apex of spinous process), distinctly tapering posteriorly; with a pore near ventral proximal margin ( +Fig. 5A +); outer distal corner produced into posteriorly directed, dorsally recurved spinous process; no spinular ornamentation discernible. Armature consisting of seven setae; seta I small; setae II and III (displaced to dorsal surface) long and naked; seta IV short, naked, located between seta V and posterior spinous process; seta V long, naked, with proximal fracture plane ( +Fig. 5A +); seta VI small, naked and located at inner distal corner; seta VII distinctly foliaceous and tri-articulate at base. + + +Rostrum ( +Fig. 1C +) small, broadly subtriangular, tapering distally; apical part lobate and demarcated by bilateral constrictions, with two delicate sensilla. + + +Antennule ( +Fig. 3A +) long and slender, 6-segmented. Segment 1 with small seta near anterodistal margin. Segment 2 longest, about 3.4 times longer than wide. Segment 4 with long aesthetasc (L: 45 μm) fused at base with seta. Distal segment with eight setae (two distinctly spatulate; indicated by arrows in +Fig. 3A +) and apical acrothek consisting of short aesthetasc (L: 20 μm) and two setae. All setal elements naked except for plumose seta on dorsal surface of segment 2. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[8 + acrothek]. + + +Antenna ( +Fig. 2D, E +). Coxa small, without ornamentation (not figured). Basis and proximal endopodal segment forming incompletely fused allobasis, about 2.9 times as long as maximum width; original basis-endopod boundary marked by transverse surface suture at level of exopodal articulation; proximal part representing original basis with longitudinal row of small spinules near base of exopod. Exopod one-segmented, unornamented, and elongate, with a naked apical seta (about 1.3 times longer than exopod). Free endopodal segment with few lateral spinules proximally and transverse spinular row distally; medial armature consisting of two short spines (indicated by arrows in +Fig. 2D, E +); apical armature consisting of two naked spines and three geniculate setae, longest of which with spinules around geniculation and fused basally to naked accessory seta. + + +Mandible, maxillule, maxilla and maxilliped as in + +N. spinicaudatus + +(see + +Sak +et al. +2008 + +: +Figs 16D, E +; +17E, F +). Mandibular gnathobase elongate, about as long as palp; with several curved, minute teeth and one tiny recurved seta at dorsal corner. Mandibular palp uniramous, consisting of elongate, unisetose basis and one-segmented endopod with one inner, two outer, and two apical setae. Maxillule comprising praecoxa, coxa, basis and vestigial rami; praecoxal arthrite with one anterior surface seta and five spines and two setae around distal margin; coxal endite cylindrical, with two recurved spines; basis elongate, with rami completely incorporated; basal armature consisting of three apical setae; exopod and endopod represented by one and three setae, respectively. Maxilla comprising syncoxa, allobasis and endopod; syncoxa with two cylindrical endites, proximal endite with three setae (one fused at base) and distal endite with two setae (one fused at base); allobasis drawn out into long claw with one accessory seta; endopod one-segmented, with three setae; all elements naked. Maxilliped comprising syncoxa, basis and endopod; syncoxa longer than wide, unarmed, with few spinules; basis elongate and unarmed; endopod with small accessory seta and slightly curved claw, bearing strong, subterminal spinule. + + +P1 ( +Fig. 4A +). Intercoxal sclerite wide and subrectangular. Praecoxa small, triangular and naked. Coxa wider than long, without ornamentation. Basis with spinular row near base of endopod; anterior surface with a small inner spine near medial margin. Exopod three-segmented; exp-1 and -2 with several spinules around outer margin, exp-3 with single spinule; exp-1 longest, with unipinnate outer spine; exp-2 without outer element; exp-3 with short unipinnate outer spine, a long curved unipinnate spine and two geniculate setae distally. Endopod two-segmented, not prehensile, longer than exopod; enp-1 about 1.25 times longer than enp-2, with a serrate seta arising from halfway down inner margin and three coarse spinules along outer margin; enp-2 without spinules, distal margin with naked outer spine and geniculate inner seta. + + +P2–P4 ( +Fig. 4B–D +). Intercoxal sclerites rectangular (P2) or squarish (P3–P4) with concave ventral margins. Praecoxae triangular, small and naked. Coxae squarish and without ornamentation. Bases smaller than coxae, with a spinular row near base of endopod (P3–P4) and a few spinules around outer corner in P2 and P4; anterior surface with a pore; outer basal seta absent in P2, long and plumose in P3–P4. Exopods three-segmented; segments with coarse spinular ornamentation, as illustrated; outer spine of exp-1 and -2 naked (except for P3 exp-2 sparsely unipinnate); exp-3 with an outer unipinnate spine, and two setae distally (one unipinnate and one bipinnate in P2–P3 or two bipinnate setae in P4); P4 exp-2 elongate, distinctly longer than exp-1; inner seta of P4 exp-3 serrate in distal half and originating near distal margin from posterior surface. Endopods two-segmented; P2–P4 enp-1 unarmed, about 1.1, 1.6, and 4.1 times longer than their respective distal segments, with few coarse spinules along outer margin as figured, but without ornamentation along inner margin (except for single minute spinule near inner distal corner of P4 enp-1); P2 enp-2 with long, apically serrate, backwardly directed seta near proximal margin and two unequal unipinnate setae around distal margin, with two spinules along outer margin subdistally and with one spinule at inner distal corner; P3 enp-2 with three coarse spinules on anterior surface, apical margin with long, unipinnate, inner seta and short, naked, outer spine; P4 enp-2 with a spinule halfway along outer margin, apical margin with long, distally serrate and basally fused, inner seta, and long, unipinnate, outer seta. Spine and seta formula as follows: + + + + + + + + + + + + + + + + + + + + + + +
ExopodEndopod
P20.0.0210.120
P30.0.0210.020
P40.0.1210.020
+ + +Fifth legs ( +Fig. 5A +) closely set together, almost touching medially. Baseoendopod and exopod fused, forming a subrectangular plate; anterior surface with two pores; inner distal corner with strong, minutely bipinnate, spinous process (homologous to inner spine); distal margin with plumose outer basal seta, one long, naked seta, and two short, equally long, bipinnate spines. + + +Redescription of male. +Total body length from tip of rostrum to posterior margin of caudal rami 336 µm (mean = 325 μm; +n += 6). Maximum body width 31 μm (mean = 32 μm; +n += 6), measured at posterior margin of cephalothorax. Body ornamentation ( +Fig. 2A +) essentially as in female. Sexual dimorphism in antennule, genital segmentation, P5, and P6. Spermatophore length approximately 53 µm. + + +Antennule ( +Fig. 3B +) 8-segmented, haplocer; geniculation between segments 6 and 7. Segment 1 with a slender naked seta; segment 2 longest and about 3.6 times longer than wide, with one plumose and seven naked setae; segment 3 with four setae and a spine; segment 4 an incomplete sclerite with two spiniform elements; segment 5 with six setae and a long aesthetasc (50 µm) fused basally to a slender seta; segments 6 and 7 with a seta; distal segment with seven setae (three of which spatulate and indicated by arrows in +Fig. 3B +) and apical acrothek consisting of short aesthetasc (12 µm) fused basally to two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4 + 2 spines], 4-[2 spines], 5-[6 + (1 + ae)], 6-[1], 7-[1], 8-[7 + acrothek]. + + +P5 ( +Fig. 5B +) with armature as in female but inner spinous process slightly more slender. + + +Sixth legs ( +Fig. 5B +) asymmetrical, with smallest P6 closing off functional gonopore; each with a long outer seta and a short inner spine, both elements naked. + + + + +Remarks. +Kunz (1971) +treated + +A. longiremis + +and + +A. accraensis + +as subspecies of + +A. acantha + +. +Bodin (1976) +adopted the division of + +A. acantha + +in three subspecies but reinstated + +N. longiremis + +in later editions of his catalogue ( +Bodin 1979 +, +1988 +, +1997 +). +Kunz’s (1971) +subspecific classification was rejected by later revisers ( +Itô 1978 +; +Bodiou & Colomines 1986 +; +Wells 2007 +; + +Sak +et al. +2008 + +) who treated all three subspecific taxa as valid species. Within the + +acanthus + +-subgroup, + +N. acanthus + +differs from + +N. chaufriassei + +primarily in the proportional lengths of the endopodal segments of P1–P3 (enp-1:enp-2 length ratio 1.25, 1.20, 1.65 +vs +1.65, 0.90, 1.20, respectively), the length and appearance of the inner subdistal seta on P4 exp-3, and the morphology of the P +5 in +both sexes. + +Neoleptastacus huysi + +is morphologically very close to its Mediterranean congener, + +N. acanthus + +, sharing the shape of the paired anal processes, the elongate P4 exp-2 (about 1.3 times the length of exp-1) and the morphology of P5. The only difference that distinguishes both species is the presence of only one outer spine on P1 exp- +3 in + +N. huysi + +, raising the suspicion that the +holotype +female (and only known specimen) of the latter species was based on an aberrant or damaged individual. + + + +Neoleptastacus acanthus + +has been recorded on multiple occasions in the Indian Ocean and the North and Southern Atlantic. Unfortunately, most of these records are not accompanied by illustrations which could have confirmed their authenticity. The species is widely distributed in the Western Mediterranean basin. +Chappuis (1954) +and +Delamare Deboutteville (1953a +– +c, 1960 +) recorded it from +Italy +( +Lazio +, Fregene), +Algeria +(El Kala = La Calle; +Skikda += Philippeville; +Jijel += Djidjelli; Beni Saf; Terga = Plage Turgot; Mers el Hadjad = Port aux Poules) and +Tunisia +(Gamarth-Plage, Rass Salakta). Reliable Italian records are those from near Cagliari ( +Cottarelli 1975 +) and Isola Tavolara ( +Cottarelli & Venanzetti 1989 +) in Sardinia, from Sperlonga in Lazio ( +Cottarelli 1969 +; +1971 +; + +Cottarelli +et al. +1994 + +), and from two localities (Tombolo della Giannella, +Marina di Alberese +) near Porto S. Stefano in Tuscany ( +Cottarelli 1973b +). +Martínez Arbizu & Moura (1994) +published the only record from +Spain +( +Valencia +, El Saler). + + +Records from the Eastern Mediterranean include those from Elafonisi Beach and Pahia Ammos on the island of +Crete +, +Greece +( +Sevastou 2005 +; + +Sevastou +et al. +2011 + +) and from the Datça-Bozburun peninsula ( +Muğla Province +) ( +Sak 2004 +; +Alper 2009 +; + +Alper +et al. +2010 + +; present account) and Dilek peninsula ( +Aydın Province +) ( + +Alper +et al. +2015 + +) in +Türkiye +. +Masry’s (1970) +records from Nahariyya and Akhziv in +Israel +are the only published outliers in the Levantine Sea but note that his setal formula of P1 endopod and P4 exopod contradicts his illustrations, and that his report of an outer basal seta on P1–P2 is most probably false. The great variability alluded to by some authors ( +e.g. +Karanovic 2000 +) has not yet been established as genuine. + + +Božić’s (1967: 873 +, +Fig. 3-3 +, +4 +) record from the Indian west coast (locality unknown but possibly +Kerala +) conceivably refers to + +N. indicus + +or a related species (see below). His illustrations of the female P5 and the anal somite including the caudal rami in dorsal aspect are of no use in deciding the identity of his material. +Božić (1967) +stated that the body length is about 400 μm and that the number of eggs in the egg-sac ranges between one and four. + +Munro +et al. +(1978) + +also reported “ + +Arenopontia acantha + +” as one of the most abundant harpacticoids in the sandy beach of Cherthala (formerly Shertallai) in +Kerala +, +India +. + + +Lindgren’s (1972 +, +1976 +) records from the beaches at Bogue Sound and Iron Steamer Pier, +North Carolina +( +U.S.A. +) are not accompanied by illustrations and are more likely based on the geographically close + +N. gussoae + +. Various South African workers ( +e.g. +McLachlan 1980 +; + +Fricke +et al. +1981 + +; + +Hennig +et al. +1983 + +) have reported “ + +Arenopontia acantha + +” in significant numbers in several beaches from around Cape Town in the west to Port Elizabeth and Algoa Bay in the east (Western Cape Province). It is highly likely that these authors were dealing with a different member of the + +acanthus + +-group which is probably conspecific with the species recorded as + +Arenopontia +sp. + +from Algoa Bay ( +McLachlan & Furstenberg 1977 +). +Wandeness (1998) +recorded + +N. acanthus + +from a sandy beach in the Macaé region in +Rio de Janeiro State +, +Brazil +, but it is conceivable that this geographically disjunct record is also based on a different species. + + + + \ No newline at end of file diff --git a/data/62/7E/C6/627EC678F76EFFB7FF4EFBCB7CB9F95F.xml b/data/62/7E/C6/627EC678F76EFFB7FF4EFBCB7CB9F95F.xml index c187c5a42da..999a07ba47a 100644 --- a/data/62/7E/C6/627EC678F76EFFB7FF4EFBCB7CB9F95F.xml +++ b/data/62/7E/C6/627EC678F76EFFB7FF4EFBCB7CB9F95F.xml @@ -1,71 +1,71 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E (5) - + speluncae -group diff --git a/data/62/7E/C6/627EC678F76EFFB7FF4EFE3B7D0EFBF3.xml b/data/62/7E/C6/627EC678F76EFFB7FF4EFE3B7D0EFBF3.xml index 504c5e15358..ce6f26c36ac 100644 --- a/data/62/7E/C6/627EC678F76EFFB7FF4EFE3B7D0EFBF3.xml +++ b/data/62/7E/C6/627EC678F76EFFB7FF4EFE3B7D0EFBF3.xml @@ -1,71 +1,71 @@ - - - -Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. - - -Author + + +Author -Sak, Serdar -0000-0002-4955-3162 -Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 -serdarsak@gmail.com. +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. - - -Author + + +Author -Karaytuğ, Süphan -0000-0001-8980-4133 -Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 -suphankaraytug@gmail.com. +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. - - -Author + + +Author -Huys, Rony -0000-0003-2411-7003 -Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk -r.huys@nhm.ac.uk. +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-10-21 + +2024 + +2024-10-21 - -5525 + +5525 - -1 + +1 - -1 -66 + +1 +66 - -http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 -journal article -10.11646/zootaxa.5525.1.1 -1175-5334 -14021479 -7F2F59B2-E0FB-4E17-BAF1-31228DB9428E +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E (4) - + trisetosus -group diff --git a/data/62/7E/C6/627EC678F772FFABFF4EFC077D48F800.xml b/data/62/7E/C6/627EC678F772FFABFF4EFC077D48F800.xml new file mode 100644 index 00000000000..b3910c929ca --- /dev/null +++ b/data/62/7E/C6/627EC678F772FFABFF4EFC077D48F800.xml @@ -0,0 +1,236 @@ + + + +Review of Neoleptastacus Nicholls, 1945 (Harpacticoida, Arenopontiidae), including an updated key to species and proposal of Phreatipontia gen. nov. + + + +Author + +Sak, Serdar +0000-0002-4955-3162 +Department of Biology, Faculty of Science and Literature, Balıkesir University, Balıkesir, Türkiye. serdarsak @ gmail. com. https: // orcid. org / 0000 - 0002 - 4955 - 3162 +serdarsak@gmail.com. + + + +Author + +Karaytuğ, Süphan +0000-0001-8980-4133 +Department of Biology, Faculty of Science, Mersin University, Mersin, Türkiye. suphankaraytug @ gmail. com. https: // orcid. org / 0000 - 0001 - 8980 - 4133 +suphankaraytug@gmail.com. + + + +Author + +Huys, Rony +0000-0003-2411-7003 +Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Corresponding author. r. huys @ nhm. ac. uk +r.huys@nhm.ac.uk. + +text + + +Zootaxa + + +2024 + +2024-10-21 + + +5525 + + +1 + + +1 +66 + + + + +http://dx.doi.org/10.11646/zootaxa.5525.1.1 + +journal article +10.11646/zootaxa.5525.1.1 +1175-5334 +14021479 +7F2F59B2-E0FB-4E17-BAF1-31228DB9428E + + + + + + + +Neoleptastacus africanus +( +Chappuis & Rouch, 1961 +) + + + + + + + + +Arenopontia africana +Chappuis & Rouch, 1961 + + + + + +Arenopontia +( +Neoleptastacus +) +africana +Chappuis & Rouch, 1961 + +: + +Wells (1967: 324) + + + + + +Arenopontia +( +Neoleptastacus +) +africana + +f. + +africana +Chappuis & Rouch, 1961 + +: + +Kunz (1971: 358) + + + + + + +Neoleptastacus africanus +( +Chappuis & Rouch, 1961 +) + +Sak +et al. +(2008: 412) + + + + + + +Original description. +Chappuis & Rouch (1961) +: 606–607; +Figs 1–8 +. + + + + + +Type +locality. + +Ghana +, +Greater Accra Region +, +Accra +; in front of lighthouse. + + +Body length. +380 μm ( +♀ +); unknown for +♂ +. + + + + +Remarks. + +Neoleptastacus africanus + +belongs to the + +trisetosus + +-group based on (1) P1 exp-1 without outer spine, (2) distal segment (exp-3, or exp-2 when exopod 2-segmented) with only one outer spine, (3) P2 exp-2 with very long outer setiform element (extending far beyond distal margin of exp-3), (4) P2–P3 endopods 1-segmented with reduced armature (010), and (5) outer seta of P4 enp-2 reduced. Within this group it is morphologically most similar to its geographically closest member, + +N. angolensis + + +comb. nov. + +, with which it shares the 3-segmented condition of the P1 exopod, a distinctly elongate P4 exopod, the reduced armature on the fifth legs, and the presence of a middorsal process on the anal operculum. +Kunz (1971) +listed seven differences between + +N. africanus + +and + +N. angolensis + + +comb. nov. + +but at least five of them appear unreliable or incorrect, +e.g. +(1) +Kunz (1971) +claimed that the outer spine on P1 exp-1 is present in + +N. africanus + +but the short outer element figured by +Chappuis & Rouch (1961) +appears too small to be a genuine spine (when present, its size invariably approaches that of the outer spines on exp-3) and is interpreted here as a spinule (as found in the same position in other species of the genus); (2) according to +Kunz (1971) +, P4 enp-2 has only one seta but +Chappuis & Rouch (1961) +clearly stated (and illustrated) that the long distal seta is accompanied by “... une fine soie au bord externe”; (3) the difference related to the antennary exopod (represented by seta +vs +unisetose segment) is ambiguous because +Chappuis & Rouch (1961) +were not explicit about its condition (“Exopodite de l’antenne II, une soie courte et mince”; and (4) the pinnules bordering the anal operculum in + +N. africanus + +are not part of the opercular ornamentation but a misinterpretation of the underlying incised anal frill. Both species require thorough redescription but can be distinguished at present by the shape/curvature of the P5 and caudal rami. The number of spiniform armature elements on the P5 is reduced in both species, being one in + +N. angolensis + + +comb. nov. + +and none in + +N. africanus + +. The species is known only from the +type +locality. + + + + \ No newline at end of file diff --git a/data/DE/0C/C7/DE0CC76FE16AFF8DFF4EF9A9FC09F90C.xml b/data/DE/0C/C7/DE0CC76FE16AFF8DFF4EF9A9FC09F90C.xml index b1522887518..091724f07be 100644 --- a/data/DE/0C/C7/DE0CC76FE16AFF8DFF4EF9A9FC09F90C.xml +++ b/data/DE/0C/C7/DE0CC76FE16AFF8DFF4EF9A9FC09F90C.xml @@ -1,73 +1,73 @@ - - - -On the Brazilian Exocnophila stick insects (Insecta: Phasmatodea), with the description of a new species + + + +On the Brazilian Exocnophila stick insects (Insecta: Phasmatodea), with the description of a new species - - -Author + + +Author -Ghirotto, Victor Morais -Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil & Projeto Phasma, São Paulo, Brazil +Ghirotto, Victor Morais +Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil & Projeto Phasma, São Paulo, Brazil - - -Author + + +Author -Conle, Oskar -Am Freischütz 16, 47058 Duisburg, Germany +Conle, Oskar +Am Freischütz 16, 47058 Duisburg, Germany - - -Author + + +Author -Hennemann, Frank -Tannenwaldallee 53, 61348 Bad Homburg, Germany +Hennemann, Frank +Tannenwaldallee 53, 61348 Bad Homburg, Germany - - -Author + + +Author -Valero, Pablo -Aachrain 1, 87534 Oberstaufen, Germany +Valero, Pablo +Aachrain 1, 87534 Oberstaufen, Germany - - -Author + + +Author -Cancello, Eliana M. -Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil +Cancello, Eliana M. +Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-01 + +2024 + +2024-11-01 - -5536 + +5536 - -1 + +1 - -59 -98 + +59 +98 - -http://dx.doi.org/10.11646/zootaxa.5536.1.2 + +http://dx.doi.org/10.11646/zootaxa.5536.1.2 -journal article -10.11646/zootaxa.5536.1.2 -1175-5326 -14024019 -47C8D2C3-7FA9-4E5E-8167-97016ABBECD5 +journal article +10.11646/zootaxa.5536.1.2 +1175-5326 +14024019 +47C8D2C3-7FA9-4E5E-8167-97016ABBECD5 @@ -231,12 +231,16 @@ det. . syn. nov. - + >> Holotype -, ♀ ( +, +♀ +( NMW -), with two eggs: +), with +two eggs +: det. Br.v.W. Ocnophila integra @@ -257,18 +261,19 @@ Zompro , n. sp. -♀ -and 2 eggs, HOLOTYPUS, PLT of - + +♀ + +and +2 eggs +, HOLOTYPUS, PLT of + Ocnophila integra -Brunner v. W. +Brunner v. W. , 1907, det. O.Zompro - -XI.1997 - -( +XI.1997 ( NHMW ) ( Figs 1–3 @@ -277,7 +282,7 @@ and 2 eggs, HOLOTYPUS, PLT of ). - + Paratype , @@ -306,16 +311,13 @@ Zompro n. sp. ♀ , PARATYPUS, PLT of - + Ocnophila integra -Brunner v. W. +Brunner v. W. , 1907, det. O.Zompro - -XI.1997 - -( +XI.1997 ( NHMW ). diff --git a/data/DE/0C/C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml b/data/DE/0C/C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml index 51dff4b6a47..8ff64f1c238 100644 --- a/data/DE/0C/C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml +++ b/data/DE/0C/C7/DE0CC76FE17FFFA2FF4EF934FAB0FE6E.xml @@ -1,75 +1,75 @@ - - - -On the Brazilian Exocnophila stick insects (Insecta: Phasmatodea), with the description of a new species + + + +On the Brazilian Exocnophila stick insects (Insecta: Phasmatodea), with the description of a new species - - -Author + + +Author -Ghirotto, Victor Morais -Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil & Projeto Phasma, São Paulo, Brazil +Ghirotto, Victor Morais +Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil & Projeto Phasma, São Paulo, Brazil - - -Author + + +Author -Conle, Oskar -Am Freischütz 16, 47058 Duisburg, Germany +Conle, Oskar +Am Freischütz 16, 47058 Duisburg, Germany - - -Author + + +Author -Hennemann, Frank -Tannenwaldallee 53, 61348 Bad Homburg, Germany +Hennemann, Frank +Tannenwaldallee 53, 61348 Bad Homburg, Germany - - -Author + + +Author -Valero, Pablo -Aachrain 1, 87534 Oberstaufen, Germany +Valero, Pablo +Aachrain 1, 87534 Oberstaufen, Germany - - -Author + + +Author -Cancello, Eliana M. -Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil +Cancello, Eliana M. +Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-01 + +2024 + +2024-11-01 - -5536 + +5536 - -1 + +1 - -59 -98 + +59 +98 - -http://dx.doi.org/10.11646/zootaxa.5536.1.2 + +http://dx.doi.org/10.11646/zootaxa.5536.1.2 -journal article -10.11646/zootaxa.5536.1.2 -1175-5326 -14024019 -47C8D2C3-7FA9-4E5E-8167-97016ABBECD5 +journal article +10.11646/zootaxa.5536.1.2 +1175-5326 +14024019 +47C8D2C3-7FA9-4E5E-8167-97016ABBECD5 - + @@ -90,7 +90,7 @@ Ghirotto, Conle, Hennemann & Valero ) - + >> Holotype @@ -106,10 +106,13 @@ Ghirotto, Conle, Hennemann & Valero Brasil . Rio de Janeiro -. Itatiaia. -PARNA -Itatiaia -, próx ao alojamento dos pesquisadores, +. +Itatiaia +. +PARNA Itatiaia +, +próx ao alojamento dos pesquisadores +, 750m @@ -126,16 +129,25 @@ col. + Paratypes , ♀ -(MZUSP): +( +MZUSP +): 1♂ -(MZUSP V0044), +( +MZUSP +V0044 +), 1♀ -(MZUSP V0043), eggs: +( +MZUSP +V0043 +), eggs: Brasil . Rio de Janeiro @@ -143,116 +155,313 @@ col. 22°26’07.7”S 44°36’46.3”W . -i.2017 -. VM Ghirotto, PABA Neves, PW Engelking, PIC Machado col.; + +i.2017 + +. +VM Ghirotto +, +PABA Neves +, +PW Engelking +, +PIC Machado +col.; + + 2♂ -(MZUSP V0292, V0293), +( +MZUSP +V0292 +, +V0293 +), 1♀ -(MZUSP V0309), eggs: +( +MZUSP +V0309 +), eggs: Brasil . Rio de Janeiro -. Itatiaia. PARNA Itatiaia, próx ao alojamento dos pesquisadores, -750m -, +. +Itatiaia +. PARNA Itatiaia, +próx ao alojamento dos pesquisadores +,750m, 22°27’10.5”S 44°36’33.7”W ,on cf. -Synedrella nodiflora + +Synedrella nodiflora + , Asteraceae . -05.v.2019 -. VM Ghirotto, RP Indicatti, EF Trova col.; + +05.v.2019 + +. +VM Ghirotto +, +RP Indicatti +, +EF Trova +col.; + + 1♂ -juvenile (MZUSP V0400): +juvenile +( +MZUSP +V0400 +): Brasil . Rio de Janeiro -. Itatiaia. PARNA Itatiaia, próx ao alojamento dos pesquisadores, -750m +. +Itatiaia +. +PARNA Itatiaia , +próx ao alojamento dos pesquisadores +, 750m, 22°27’10.5”S 44°36’33.7”W . -07.v.2019 -. VM Ghirotto, RP Indicatti col.; + +07.v.2019 + +. +VM Ghirotto +, +RP Indicatti +col. + +; + 1♂ -(MZUSP0262): +( +MZUSP0262 +): Brasil . Est do Rio [ Rio de Janeiro -]. Itatiaia (1.100 mtr.). -i.1972 -. Dirigns [col.].; +]. +Itatiaia +(1.100 mtr.). + +i.1972 + +. +Dirigns +[col.].; + + 1♂ -(MZUSP0282): +( +MZUSP0282 +): Brasil . Est do Rio [ Rio de Janeiro -]. Itatiaia (1.100 mtr.). -iv.1960 -. Dirigns [col.].; +]. +Itatiaia +(1.100 mtr.). + +iv.1960 + +. +Dirigns +[col.]. + +; + 1♂ -(MZUSP0685), +( +MZUSP0685 +), 1 subadult ♂ -(MZUSP0686), +( +MZUSP0686 +), 2♀ -(MZUSP0738, 0783): +( +MZUSP0738 +, +0783 +): Brasil . Rio de Janeiro -. Itatiaia. Parque Nacional de Itatiaia, parte baixa. +. +Itatiaia +. +Parque Nacional de Itatiaia +, parte baixa. 22°25’35”S 44°37’05”W . -31.x–2.xi.2016 -. P.I.Chiquetto-Machado col.; + +31.x–2.xi.2016 + +. +P.I.Chiquetto-Machado +col. + +; + 2♂ -(MNRJ-ENT15-17, MNRJ-ENT15-18): +( +MNRJ-ENT15-17 +, +MNRJ-ENT15-18 +): Brasil . Rio de Janeiro -. Itatiaia. PN Itatiaia, parte baixa, ao redor da Casa do Pesquisador. -06–08.ii.2020 -. P. Souza-Dias, Siqueira, Lima, Borille col.; +. +Itatiaia +. +PN Itatiaia +, +parte baixa, ao redor da Casa do Pesquisador +. + +06–08.ii.2020 + +. +P. Souza-Dias +, +Siqueira +, +Lima +, +Borille +col. + +; + 1♂ -(FIOC): +( +FIOC +): Brasil . E. do Rio [ Rio de Janeiro -]. Itatiaia. 10.viii.[19]44. J.F. Zikan col; +]. +Itatiaia +. 10.viii.[19]44. +J.F. Zikan +col + +; + 13♂ -(PNI022, PNI037–40, PNI094), +( +PNI022 +, +PNI037–40 +, +PNI094 +), 7♀ -(PNI119–120, PNI206, PNI218–221), eggs (FIOC): +( +PNI119–120 +, +PNI206 +, +PNI218–221 +), +eggs +( +FIOC +): Brasil . Rio de Janeiro -. Parque Nacional do Itatiaia, along BR 485 betw. park entrance & Cachoeira de Maromba, 22°45’97 S 44°59’91 W – -22°25’47 S -44°37’05 W +. +Parque Nacional do Itatiaia , -720–1120 m -a.s.l. -30.x.–06.xi.2016 -. OV Conle, FH Hennemann, Uhlig, Y Gutiérrez, P Valero, J Costa, Rodrigues, PABA Neves, PI Chiquetto-Machado, VM Ghirotto, PW Engelking col.; +along BR 485 betw. park entrance & Cachoeira de Maromba +, +22°45’97 S 44°59’91 W +– 22°25’47 S 44°37’05 W, 720–1120 m a.s.l. + +30.x.–06.xi.2016 + +. +OV Conle +, +FH Hennemann +, +Uhlig +, +Y Gutiérrez +, +P Valero +, +J Costa +, +Rodrigues +, +PABA Neves +, +PI Chiquetto-Machado +, +VM Ghirotto +, +PW Engelking +col. + +; + 1♀ -(FIOC PNI036): +( +FIOC +PNI036 +): Brasil . Rio de Janeiro -. Parque Nacional do Itatiaia, near Lago Azul, +. +Parque Nacional do Itatiaia +, +near Lago Azul +, 22°27’10 S 44°36’36 W +, 750–780 m a.s.l. + +04.11.2016 + +. +OV Conle , -750–780 m -a.s.l. -04.11.2016 -. OV Conle, FH Hennemann, Uhlig, Y Gutiérrez, P Valero, J Costa, Rodrigues, PABA Neves, PI Chiquetto-Machado, VM Ghirotto, PW Engelking col. +FH Hennemann +, +Uhlig +, +Y Gutiérrez +, +P Valero +, +J Costa +, +Rodrigues +, +PABA Neves +, +PI Chiquetto-Machado +, +VM Ghirotto +, +PW Engelking +col. +