Add updates up until 2025-07-02 14:09:39

This commit is contained in:
ggserver 2025-07-02 14:15:44 +00:00
parent ce636872e3
commit b289c91f4a
11 changed files with 3926 additions and 0 deletions

View file

@ -0,0 +1,254 @@
<document id="D42808E0A16534647C8B5359509D86A8" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="552883EA6D205E00B0B8F7F06BB2B636" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Josephiella microcarpae Beardsley &amp; Rasplus 2001" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="191AD46FC75A6A57A32816FDDFCC62A2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2C1D904BD0D6047FD14FF6C16D2E71E">
<mods:title id="F40A2E2B8F030F9FB8CD272C6FB8A742">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="3BE479E194E98630AE8FDC4520E3AE21" type="personal">
<mods:role id="50915D8415549027AD8CF7B970A0850C">
<mods:roleTerm id="FC1D62B0F0DBE10770F94A09CED6571E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37CF9F9F5D268DC974C8F3FC747CBCAD">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="2A97E3DA78B305943C2B7032CCE088CF" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="7CA4BF6D087C39ABFE536725B8D66A18">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="98C71FE200B6508F17F524A2D0534F38" type="personal">
<mods:role id="42488FA5925213671296AAF6231B4735">
<mods:roleTerm id="3C395DAE56FA402B8C9CED3B9D83044D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D8F148ADDB40E32CFA6AF8C22623FCB">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="D37BB1174A717E0F6CB9E9189FBAFC58" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="41C7FA94F31380EEC82F4940DFBE0B97">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="E73936DBE4CE9EC71980AE549C294B0A" type="personal">
<mods:role id="6A3F36E104E586330551F1906619C39B">
<mods:roleTerm id="786B60D6101FDBA5A267696348E23644">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7171EE7E1483B1516C59E830AA87A7A3">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="ECBC0F99212E4C1A6135CB07D0D02B93" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="C6D00182D0503D394960E11A9E2888F7">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="0A00764BFFAA6380F37291CE8DBB4D62" type="personal">
<mods:role id="3439D703841D1A6AFFCEACC2B2868F48">
<mods:roleTerm id="11E005EDE512B85CB23CBC2D8146ECDD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A40D46D834DE80C621C28757797E4CCF">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="4E9DDCE0EB625A0E0982B02C55FE8588" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="D8E5BE34408FEA636F5DFC1649A9569D">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="A1FF77CBE48539FA4F9DE5A5F15E0F90" type="personal">
<mods:role id="C7F7FC0D1B759C467EED89B61046D623">
<mods:roleTerm id="E762A0D5338E6D25FE55E018DFE4BDC5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="87100265038C22210F0974E6B9B0C8EA">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="39EDDA2AA0B14B3F6E44851324207C52" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="FF43195AC34A61F3437D75DED952E386">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6FEAC0DE6102B02E75A11C2C31D1C525">text</mods:typeOfResource>
<mods:relatedItem id="D52A010EDC9E2CAE4E302E037A204CB4" type="host">
<mods:titleInfo id="5C40251FCAF6E4FD8B6A85951DFC9F0C">
<mods:title id="37DDAF67B53270C144DA92A64093A5B0">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="A715D4BC70ACF54E4FF7743A4D0EDC84">
<mods:date id="877D369F9CF873A224ABB9CC473014C4">2025</mods:date>
<mods:detail id="4E8E54FC98F776359BF0B88027FCEF3C" type="pubDate">
<mods:number id="046DB1A5C7F3B74644D5B6145E874702">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="20BF08D4502A039076A3E889A172E550" type="volume">
<mods:number id="F82D535D9448708FB8EF6EAC99C19E36">98</mods:number>
</mods:detail>
<mods:extent id="BFAF806289AFF8C794C5F31993F66536" unit="page">
<mods:start id="08D4F5A5B596B3A9644DA05750AC0011">667</mods:start>
<mods:end id="008CEECC0B5A2A0E40906A0B421DBD59">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A838C53C1E8ABB67BF2F7490004CFB7E">journal article</mods:classification>
<mods:identifier id="B70003611B85A314CC48915281BD5C75" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="CFB0E774C1BDF0868ED45BBAB2F33933" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="552883EA6D205E00B0B8F7F06BB2B636" ID-DOI="http://doi.org/10.5281/zenodo.15791856" ID-Zenodo-Dep="15791856" ID-arpha="552883EA-6D20-5E00-B0B8-F7F06BB2B636" LSID="urn:lsid:plazi:treatment:552883EA6D205E00B0B8F7F06BB2B636" httpUri="http://treatment.plazi.org/id/552883EA6D205E00B0B8F7F06BB2B636">
<subSubSection id="9F5636BFB928A3E656B9F242D3533773" type="nomenclature">
<paragraph id="1DB78E162E257EFD4077CF5039DA32E6">
<taxonomicName id="0717697DABA01912735A500B7A0D1658" ID-CoL="3QR4J" authority="Beardsley &amp; Rasplus, 2001" authorityName="Beardsley &amp; Rasplus" authorityYear="2001" class="Insecta" family="Agaonidae" genus="Josephiella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="microcarpae">
<emphasis id="F6C9B4A00DDE63CDBD200CDE09F4A8CD" italics="true">Josephiella microcarpae</emphasis>
<bibRefCitation id="57E5EC079FCB6DD0414098B31D4A944B" DOI="10.1080/002229301447871" author="Beardsley JW &amp; Rasplus J-Y" journalOrPublisher="Journal of Natural History" pagination="33-40" refId="B3" refString="Beardsley JW, Rasplus J-Y (2001) A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae). Journal of Natural History 35: 33-40. https://doi.org/10.1080/002229301447871" title="A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae)." volume="35" year="2001">Beardsley &amp; Rasplus, 2001</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGFAG" type="material">
<paragraph id="68FF843950A3774D49155C602E5C413F">
<heading id="822BFA18AE8D1F307489AA0F3A4EF8EB" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="EB05AA7B0907D623D5E117515D414F00">
<materialsCitation id="B2442AB9D1084EBA8743A7C893CB0CA6" collectingDate="2021-02-02" collectionCode="AC" collectorName="J. M. Royo" country="Spain" latitude="38.389034" location="Vicente Savall Pascual Street" longLatPrecision="1" longitude="-0.51813" municipality="San Vicente del Raspeig" specimenCount="4" specimenCount-female="4" stateProvince="Alicante">
<emphasis id="3107D719B5301D95877D38A4DBAC4F48" bold="true">
<collectingCountry id="BDBD470ED1FB753B7A9C17E5BD17E480" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="FBDD94DFF7591F29F5BCA584125022A4" bold="true">
<collectingRegion id="5665A237B98E072074251300922BCCA3">Alicante</collectingRegion>
</emphasis>
:
<collectingMunicipality id="EB707E525495B18CEF62ABA9F28FD254">San Vicente del Raspeig</collectingMunicipality>
:
<location id="D872CFD6FD99C7DA84319F453F3E8742" LSID="urn:lsid:plazi:treatment:552883EA6D205E00B0B8F7F06BB2B636:D872CFD6FD99C7DA84319F453F3E8742" country="Spain" latitude="38.389034" longLatPrecision="1" longitude="-0.51813" municipality="San Vicente del Raspeig" name="Vicente Savall Pascual Street" stateProvince="Alicante">Vicente Savall Pascual Street</location>
:
<specimenCount id="95DC731E9CBB39CAA76D56E950585E16" count="4" type="female">4 ♀</specimenCount>
<abbrev id="ABBRID0EQFAG" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="D6515DB81DE555B9C8DD1446D4E1776B">J. M. Royo</collectorName>
<abbrev id="ABBRID0EUFAG" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="933D9ADED3324207D968AE4DE33DF0D0" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EYFAG" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="864D1C66837293756B21D50FADAFA837" value="2021-02-02">
<collectingDate id="79C4882ECA893A043488EB840C5B2776" value="2021-02-02">
02 - II-
<quantity id="83700A54B454E38645806142C5826F09" metricMagnitude="6" metricUnit="m" metricValue="2.021193" unit="m" value="2021193.0">2021, 193 m</quantity>
</collectingDate>
</date>
a. s. l, found in dissecting leaf galls of
<taxonomicName id="3BF83BF1401C83138C012F5C876AF4F5" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="D041C74209DF5CAF231C16C3ECDAD3C2" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKGAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-0.518130,38.389033]}">
<geoCoordinate id="80DF6E40B8CDDEB7B2FCF5B72901FF0E" degrees="38.389033" direction="north" orientation="latitude" precision="1" value="38.389034">38.389033</geoCoordinate>
,
<geoCoordinate id="AC8D788AF096C256B6C5251FCDA4CFD4" degrees="0.518130" direction="west" orientation="longitude" precision="1" value="-0.51813">- 0.518130</geoCoordinate>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="FE88D3B2CCE1F074FB1F068E4BBF087C" collectingDate="2019-08-15" collectionCode="AC" collectorName="Barriada El Cobre &amp; I. Sanchez &amp; A. Perez" country="Spain" elevation="84" latitude="36.118237" location="Barriada El Cobre" longLatPrecision="1" longitude="-5.479646" municipality="Algeciras" specimenCount="9" specimenCount-female="8" specimenCount-male="1" stateProvince="Cadiz">
<emphasis id="EF946F7E8B9CC55C8F0FFB3AAA0F01F1" bold="true">
<collectingRegion id="1C0512A5F106941A21DCCA1F631D58A8">Cádiz</collectingRegion>
</emphasis>
:
<collectingMunicipality id="1E907BA196B0902B1C16A66A5694D62B">Algeciras</collectingMunicipality>
:
<collectorName id="B5E7390E306A3E5575496E2B79B3DD74">
<location id="54E3880243B02200DC94F262663BEC34" LSID="urn:lsid:plazi:treatment:552883EA6D205E00B0B8F7F06BB2B636:54E3880243B02200DC94F262663BEC34" country="Spain" latitude="36.118237" longLatPrecision="1" longitude="-5.479646" municipality="Algeciras" name="Barriada El Cobre" stateProvince="Cadiz">Barriada El Cobre</location>
</collectorName>
:
<specimenCount id="8A14A33FEECD15CFE60B4AADCA492517" count="8" type="female">8 ♀</specimenCount>
+
<specimenCount id="97D349D4C05FD34ED86FF689B5D31207" count="1" type="male">1 ♂</specimenCount>
<abbrev id="ABBRID0ERGAG" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="7405385888B7E63025C81F6210C3D21F">I. Sánchez</collectorName>
and
<collectorName id="DD4701C23ADCC1DD3B435600DC94AC22">Á. Pérez</collectorName>
<abbrev id="ABBRID0EVGAG" xlink_title="collector">leg</abbrev>
and
<collectionCode id="7DF6A1B9676DB25D5C1F523E89C9323C" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EZGAG" xlink_title="collection of deposit">deposit</abbrev>
.
<date id="94EAA0E9EF72F1076FDA5EABBDF5B4CC" value="2019-08-15">
<collectingDate id="5A4B97DD6220BF67520BB3F4CEBA675B" value="2019-08-15">15 - VIII- 2019</collectingDate>
</date>
,
<quantity id="4DD4DBF7F2D3B905C3D63B441CFD65BC" metricMagnitude="1" metricUnit="m" metricValue="8.4" unit="m" value="84.0">
<elevation id="63EF245731FB947332F257679D669D54" metricMagnitude="1" metricUnit="m" metricValue="8.4" unit="m" value="84.0">84 m</elevation>
</quantity>
a. s. l, found in dissecting leaf galls of
<taxonomicName id="84D4467DD2A1C0582006710D06EE5FB7" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="252E3BB444A6B1ED98C344FAABC99566" italics="true">F. microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELHAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-5.479646,36.118236]}">
<geoCoordinate id="52E15F94D6CEEEB6DBC93CEBEBB1E943" degrees="36.118236" direction="north" orientation="latitude" precision="1" value="36.118237">36.118236</geoCoordinate>
,
<geoCoordinate id="B4D9321F8523095F07631E65F307DBD1" degrees="5.479646" direction="west" orientation="longitude" precision="1" value="-5.479646">- 5.479646</geoCoordinate>
</named-content>
). See Fig.
<figureCitation id="6BF8F37B6718FD9EAA1D575304F9DA07" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 H</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUHAG" type="Ecological remarks">
<paragraph id="80AC8A0B24E522867F9E66348DBD81FF">
<heading id="6BA6312BF8CF0BEFACD9FDC94B866F20" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="0532307DE22A24D84B74694467BD6568">
Leaf galler of
<taxonomicName id="8CD6B6E362BB8620B2C62DFA5A1BAB33" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="A598F89C9055B7544BD21B86BE410393" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFIAG" type="distribution">
<paragraph id="BFE046CBD60DD6A47372B6B703C59DE9">
<heading id="9F952A29D1AC0284D8B4DCA5E62F4B60" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1D4C6F18C10EBE33C3CF339CA9DE7033">
The exact origin of the wasp is unknown but could be Southeast Asia as its host plant originates from there. The species was first reported in
<collectingRegion id="0B8823E1DA206E2C7EABD35BEF80AC85" country="United States of America" name="Hawaii">Hawaii</collectingRegion>
,
<collectingRegion id="69EBDA4718D0C0AB161726D2E1629BEA" country="United States of America" name="California">California</collectingRegion>
(
<collectingCountry id="B576D56D499F01F0216F8412EF4FF545" name="United States of America">USA</collectingCountry>
) and the
<collectingRegion id="1DB143656A0B3DEAFEF67DA274AD672D" country="Spain" name="Canarias">Canary Islands</collectingRegion>
(
<collectingCountry id="03EE75E42C330E264EC150B0B4C981C8" name="Spain">Spain</collectingCountry>
) (
<bibRefCitation id="0277D7D88D22F663742E6C3FAB130F9D" DOI="10.1080/002229301447871" author="Beardsley and Rasplus" firstAuthor="Beardsley" journalOrPublisher="Journal of Natural History" pagination="33-40" refId="B3" refString="Beardsley JW, Rasplus J-Y (2001) A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae). Journal of Natural History 35: 3340. https://doi.org/10.1080/002229301447871" title="A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae)." volume="35" year="2001">Beardsley and Rasplus 2001</bibRefCitation>
;
<bibRefCitation id="997C19F9FEA49BFABDC71A9CFEAAE231" author="Rodriguez and Rodriguez" firstAuthor="Rodriguez" refId="B84" refString="Rodríguez JM, Rodríguez R (2006) Josephiella microcarpa, avispita de las agallas. Ficha 292. Grupo de Trabajo de Laboratorios de Diagnóstico (Cabildo de Gran Canaria). MAPA, 2 pp." year="2006">Rodríguez and Rodríguez 2006</bibRefCitation>
). Subsequently it was recorded from several European areas like
<collectingRegion id="D58319C5ACBBF37C4EDDFAECE8587D85" country="Italy" name="Sicilia">Sicily</collectingRegion>
(
<collectingCountry id="4BB99C9FD79AED3DFD1DA056C8E48BFE" name="Italy">Italy</collectingCountry>
),
<collectingRegion id="C4A6CC021CF2A893CE056C659A40C4F4" country="Spain" name="Illes Baleares">Balearic Islands</collectingRegion>
(
<collectingCountry id="83A95BC5F2D5523346D3E02EE0F9885A" name="Spain">Spain</collectingCountry>
),
<collectingCountry id="12407FE2CC7D1A2C541C28AA91DC74D3" name="Malta">Malta</collectingCountry>
,
<collectingCountry id="8FA0F16D00021F4F90DB7472C30DAF95" name="Portugal">Portugal</collectingCountry>
,
<collectingCountry id="4859AE4BBA5506A2E5CE333401C38ECC" name="Greece">Greece</collectingCountry>
, and
<collectingCountry id="61859C7B421544614B091C4D2FCF7340" name="Cyprus">Cyprus</collectingCountry>
(
<bibRefCitation id="D3DC6F353E195D32A4B9D28AD272EC60" author="Lo Verde" etAl="et al." firstAuthor="Lo Verde" refId="B64" refString="Lo Verde G, Porcelli F, Bella S, Rasplus J-Y (2007) Imenotteri Agonidi nuovi per lEuropa e loro ruolo nella naturalizzazione di Ficus spp. in Italia. Atti XXI CNIE, Campobasso, 60." year="2007">Lo Verde et al. 2007</bibRefCitation>
;
<bibRefCitation id="669B42661C630982F506FEC1534C27D7" author="Caldwell" firstAuthor="Caldwell" journalOrPublisher="IFAS Extension, Collier County, Florida" refId="B8" refString="Caldwell D (2008) Ficus Trees Under Severe Insect Attack! University of Florida. IFAS Extension, Collier County, Florida, 2 pp." title="Ficus Trees Under Severe Insect Attack! University of Florida." year="2008">Caldwell 2008</bibRefCitation>
;
<bibRefCitation id="18A6A02F8340737F663E9A7C1BE49B23" DOI="10.3897/biorisk.4.55" author="Rasplus" editor="Roques A" etAl="et al." firstAuthor="Rasplus" journalOrPublisher="BioRisk" pagination="669-776" refId="B79" refString="Rasplus J-Y, Villemant C, Paiva MR, Delvare G, Roques A (2010) Hymenoptera. Chapter 12. In: Roques A. (Eds) Alien terrestrial arthropods of Europe. BioRisk 4: 669776. https://doi.org/10.3897/biorisk.4.55" title="Hymenoptera. Chapter 12." volume="4" volumeTitle="(Eds) Alien terrestrial arthropods of Europe." year="2010">Rasplus et al. 2010</bibRefCitation>
;
<bibRefCitation id="45E611412BC6794CC14F8A3B2CA85D5B" author="Mifsud" etAl="et al." firstAuthor="Mifsud" journalOrPublisher="Bulletin of the Entomological Society of Malta" pagination="5-34" refId="B68" refString="Mifsud D, Falzon A, Malumphy C, De Lillio E, Vovlas N, Porcelli F (2012) On some arthropods associated with Ficus species (Moraceae) in the Maltese Islands. Bulletin of the Entomological Society of Malta 5: 534." title="On some arthropods associated with Ficus species (Moraceae) in the Maltese Islands." volume="5" year="2012">Mifsud et al. 2012</bibRefCitation>
;
<bibRefCitation id="949A97219502C76A7CD42F5A50C3C3CD" DOI="10.1111/icad.12111" author="Wang R &amp; Aylwin R &amp; Barwell L &amp; Chen XY &amp; Chen Y &amp; Chou LS &amp; Cobb J &amp; Collette D &amp; Craine L &amp; Giblin-Davis RM &amp; Ghana S &amp; Harper M &amp; Harrison RD &amp; McPherson JR &amp; Peng Y-Q &amp; Pereira RAS &amp; Reyes-Betancort A &amp; Rodríguez LJV &amp; Strange E &amp; van Noort S &amp; Yang H-W &amp; Yu H &amp; Compton SG" issue="4" journalOrPublisher="Insect Conservation and Diversity" pagination="322-336" refId="B95" refString="Wang R, Aylwin R, Barwell L, Chen XY, Chen Y, Chou LS, Cobb J, Collette D, Craine L, Giblin-Davis RM, Ghana S, Harper M, Harrison RD, McPherson JR, Peng Y-Q, Pereira RAS, Reyes-Betancort A, Rodríguez LJV, Strange E, van Noort S, Yang H-W, Yu H, Compton SG (2015 a) The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa. Insect Conservation and Diversity 8 (4): 322336. https://doi.org/10.1111/icad.12111" title="The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa." volume="8">Wang et al. 2015 a</bibRefCitation>
;
<bibRefCitation id="CA90FF83777471709E70DFEF50F5C6C3" DOI="10.3391/bir.2023.12.2.19" author="Kalaentzis" etAl="et al." firstAuthor="Kalaentzis" issue="2" journalOrPublisher="BioInvasions Records" pagination="563-571" refId="B52" refString="Kalaentzis K, Koutsoukos E, Demetriou J, Kazilas C, Avtzis DN, Georgiadis C (2023) First record and DNA barcoding of the fig gall wasp Josephiella microcarpae (Hymenoptera: Chalcidoidea: Pteromalidae) in Greece. BioInvasions Records 12 (2): 563571. https://doi.org/10.3391/bir.2023.12.2.19" title="First record and DNA barcoding of the fig gall wasp Josephiella microcarpae (Hymenoptera: Chalcidoidea: Pteromalidae) in Greece." volume="12" year="2023">Kalaentzis et al. 2023</bibRefCitation>
). The species was recently reported in the Spanish peninsular area, namely Valencia (
<bibRefCitation id="A31EF7F7A75928EBF1E956F2516E97D7" author="Rodrigo" etAl="et al." firstAuthor="Rodrigo" journalOrPublisher="Boletín de la Sociedad Entomológica Aragonesa" pagination="390-392" refId="B83" refString="Rodrigo E, Laborda R, Sánchez-Domingo A, González-Rey S, Bertomeu S, Xamaní P, Hernández-Suárez E (2017) Primer registro de Josephiella microcarpae (Hymenoptera, Agaonidae) en Ficus microcarpa en España peninsular. Boletín de la Sociedad Entomológica Aragonesa 60: 390392." title="Primer registro de Josephiella microcarpae (Hymenoptera, Agaonidae) en Ficus microcarpa en España peninsular." volume="60" year="2017">Rodrigo et al. 2017</bibRefCitation>
), Alicante (
<bibRefCitation id="C3E525F442CCAF8F6A7CCC8A665B41E7" DOI="10.14198/cdbio.2019.57.03" author="Hernández-Martínez K &amp; Marcos-García MA" journalOrPublisher="Cuadernos de Biodiversidad" pagination="23-41" refId="B43" refString="Hernández-Martínez K, Marcos-García MA (2019) Entomofauna associated the plants of the campus of the University of Alicante (Spain). Cuadernos de Biodiversidad 57: 2341. https://doi.org/10.14198/cdbio.2019.57.03" title="Entomofauna associated the plants of the campus of the University of Alicante (Spain)." volume="57" year="2019">Hernández Martínez and Marcos García 2019</bibRefCitation>
) and Catalonia (
<bibRefCitation id="C47B99911D4C9B3C674EC41B509501D3" author="Riba-Flinch and Pujade-Villar" firstAuthor="Riba-Flinch" issue="4" journalOrPublisher="Butlletí de la Institució Catalana dHistòria Natural" pagination="187-190" refId="B81" refString="Riba-Flinch JM, Pujade-Villar J (2022) First detection of Josephiella microcarpa Beardsley &amp; Rasplus, 2001 (Hymenoptera: Chalcidoidea: Agaonidae) on Ficus microcarpa L. in Catalonia and other pests that can be found on it. Butlletí de la Institució Catalana dHistòria Natural 86 (4): 187190." title="First detection of Josephiella microcarpa Beardsley &amp; Rasplus, 2001 (Hymenoptera: Chalcidoidea: Agaonidae) on Ficus microcarpa L. in Catalonia and other pests that can be found on it." volume="86" year="2022">Riba-Flinch and Pujade-Villar 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXJAG" type="description">
<paragraph id="F93D754A111F43738777AA576CA8AA7E">
<heading id="528892A7127AF89A4DD53077442208C4" reason="title">Short description.</heading>
</paragraph>
<paragraph id="FCFBF87EC210637BDC7EEE5922659E7A">
The female has a body uniformly dark brown except antennal and legs pale yellow, wings hyaline. Head reticulate in dorsal occipital area behind ocelli. Pronotum reticulate without carinae, mesoscutum smooth centrally, scutellum smooth with weak, longitudinally oriented reticulation. Hind coxae reticulate. Gaster smooth and shiny (
<bibRefCitation id="756A7FA12667BB621AEC1F04B58F1A51" DOI="10.1080/002229301447871" author="Beardsley and Rasplus" firstAuthor="Beardsley" journalOrPublisher="Journal of Natural History" pagination="33-40" refId="B3" refString="Beardsley JW, Rasplus J-Y (2001) A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae). Journal of Natural History 35: 3340. https://doi.org/10.1080/002229301447871" title="A new species of Josephiella (Hymenoptera: Agaonidae) forming leaf galls on Ficus microcarpa L. (Moraceae)." volume="35" year="2001">Beardsley and Rasplus 2001</bibRefCitation>
). This species is recognized also by the spherical swellings or warty blisters that develop on the lower and upper surfaces of the leaves of
<taxonomicName id="4FFE3F778D940BA93B5471DE934F39B3" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpae">
<emphasis id="C08462CC68134496FED9ABD6F27EE1D0" italics="true">Ficus microcarpae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,174 @@
<document id="58D2EE6A7DC9DCFA0669E092078617CE" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="5AA43C0C196955F68835025C896920AB" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Philotrypesis okinavensis Ishii 1934" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="CEAF2BE28AD0172BFCDA354E652EF313" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4FF510656DA80CD83C289728B7CA4C97">
<mods:title id="1B29C5AA4318BE951C56B0C11F161987">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="2EBBC83A5FA2324CD4FD31BFB3E542A2" type="personal">
<mods:role id="F34C4ABB7382921CF46AFA91AE88DA2E">
<mods:roleTerm id="C268BC96ECBB3324CCB7C7B445087700">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD56A987C5C48464A4BE11B186DA4D7A">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="5E1981006FA6012D950D8442C0092E6C" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="2AD5156DCE9C85B11B42846F7A02CB36">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="FA1ECCB65087CFB7985EBA7E047A7DB2" type="personal">
<mods:role id="04F48CA1FCD08887210C7A168D1259A4">
<mods:roleTerm id="197366DEFD75F08A40E32FE530C47CCE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F68E9EA897328399419B1AB1E2F69DB">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="4665C50F48BE1D51653F6A83D448B9BE" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="C3011F32426F5670CEECE2BAC5199649">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="3751992C25F5327E3B730FA728824000" type="personal">
<mods:role id="20DE75EB4850B4A47F152D36972E5879">
<mods:roleTerm id="EC8FCB92FEBEA0973339A0321A30C9D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F50F24A18547E896BA28572B7F3EAFD">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="587D9E981864190258BE1F762B7CF011" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="914B364D7BC02A71EC6749265F20CC9F">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="62B0F86DC3293B3F20C871A978306DEF" type="personal">
<mods:role id="099481A6357A24A113366A95BD3899AF">
<mods:roleTerm id="5747103B4C694794BF357EEE64E5CD34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA7FAB6786E54A99C7F1DEB46798309C">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="1363FBABEC711C3668E867EEC23F104F" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="135D328EE4FE0B52588C3B79351256B1">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="7FC4D9B5292601DE2218590B6D242F0E" type="personal">
<mods:role id="CC8AC26576C851723DE6B330B40C520A">
<mods:roleTerm id="2A8A97CA666C0CD2F9FAC925B4FE7B15">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="531DC7AB35EE9C6E168FD9EBA3222DC3">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="11148E4A9D4D1767D4DD30EC10FF40FA" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="C53FF3A2CAB0EDA3186DE20215499D52">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EB11715672EA9AF3259F87D1ED673B24">text</mods:typeOfResource>
<mods:relatedItem id="F026C9E27C186E667A4A3CAB2A7CB41D" type="host">
<mods:titleInfo id="87A252F5282CEC36378D138499D2AB54">
<mods:title id="00B29606CD6BF4049641CBA285E6E63C">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="0DB2B3841B3FA034E9A15709A1822FE7">
<mods:date id="D7B06630A1E3C6B6C61210E5EA950779">2025</mods:date>
<mods:detail id="3064DD88D3C61E83FC2A2E5C16BBA9C8" type="pubDate">
<mods:number id="32BFE759FFA9508F3A58098CA7B13D78">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="A8B109C9D2DDA61151D263B29A14C81B" type="volume">
<mods:number id="1AD94081CB1A99514DA4A0A9BD331A1E">98</mods:number>
</mods:detail>
<mods:extent id="852008C59FDE495AC50D4B4526D40E96" unit="page">
<mods:start id="498649A31348B448147FE743E2275325">667</mods:start>
<mods:end id="20A66E78CAE1A71372EC32BCED546B96">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="75A7BB8DC8C485062B477E67D565CBE8">journal article</mods:classification>
<mods:identifier id="D1AC7D76DE0E64499C4DB11CC45FB2DF" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="B8F33581F7A8802502FFE046A1E1E65E" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="5AA43C0C196955F68835025C896920AB" ID-DOI="http://doi.org/10.5281/zenodo.15791860" ID-Zenodo-Dep="15791860" ID-arpha="5AA43C0C-1969-55F6-8835-025C896920AB" LSID="urn:lsid:plazi:treatment:5AA43C0C196955F68835025C896920AB" httpUri="http://treatment.plazi.org/id/5AA43C0C196955F68835025C896920AB">
<subSubSection id="39BA139F1BE6B8DEFCB827AA37BB150B" type="nomenclature">
<paragraph id="EA078EAE7F7BE5CB7FDEF109C13D3355">
<taxonomicName id="115766690592F05A0804EFF56324E7DC" ID-CoL="4GDJW" authority="Ishii, 1934" authorityName="Ishii" authorityYear="1934" class="Insecta" family="Agaonidae" genus="Philotrypesis" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="okinavensis">
<emphasis id="839607BD612C660D89FADB39A2F5D4B8" italics="true">Philotrypesis okinavensis</emphasis>
<bibRefCitation id="1D1C7AA4A1E2E9FC57626E362178699B" author="Ishii T" journalOrPublisher="Kontyû" pagination="84-100" refId="B47" refString="Ishii T (1934) Fig. chalcidoids of Japan. Kontyu 8: 84-100." title="Fig. chalcidoids of Japan." volume="8" year="1934">Ishii, 1934</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBUAG" type="material">
<paragraph id="3826C06167ECA2EA72663811AAC1B119">
<heading id="352FF68A028B27DACCB3533AF5E1E73B" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="A96F704F83CF7A685E7FA01FC7EAAAD1">
<materialsCitation id="4C711015A9C679333FF8502B093B68EC" collectingDate="2020-11-10" collectionCode="AC" collectorName="Puerto Real &amp; A. Perez-Gomez" country="Spain" latitude="36.5306" location="See in Fig." longLatPrecision="1" longitude="-6.211997" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Cadiz">
<emphasis id="6C6A423BE97553EAA53BAEFA555D9092" bold="true">
<collectingCountry id="F72A25D61120595BA976C96B46C9730D" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="21F9B9737889BB9440EDE64EDF16EBC5" bold="true">
<collectingRegion id="DF106D9F3EE1C05A87BF81E745A76B71">Cádiz</collectingRegion>
</emphasis>
:
<collectorName id="FC2A604B663F837CEE5191226448A8DB">Puerto Real</collectorName>
:
<specimenCount id="A6D95C5F5D9616C9ED5763FF1F1BBC8F" count="2" type="male">2 ♂</specimenCount>
+
<specimenCount id="57D8AA46EE66F7EFFFA89CBBE6FBB5CF" count="1" type="female">1 ♀</specimenCount>
<abbrev id="ABBRID0ELUAG" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="BFD2DA5265C08F7F053CB0AB244BEA65">A. Pérez-Gómez</collectorName>
<abbrev id="ABBRID0EPUAG" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="62F0BC09E1EC2044D9CC21DAF7642F8B" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
.
<abbrev id="ABBRID0ETUAG" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="D8707D43D44198DB3138AD96C2BE0742" value="2020-11-10">
<collectingDate id="F809F42550DB79DC923BAFE393980ACC" value="2020-11-10">
10 - XI-
<quantity id="9974B3E456A73A5A06B2C5239775607F" metricMagnitude="6" metricUnit="m" metricValue="2.020316" unit="m" value="2020316.0">2020, 316 m</quantity>
</collectingDate>
</date>
a. s. l., found dissecting fallen syconia of
<taxonomicName id="C35D9C465B6B56095222A170015AADA4" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="A596ACAC41463C80BBA99FB1AC3B72D8" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFVAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-6.211997,36.530600]}">
<geoCoordinate id="D730A3150733CEB03B6F75E311E44432" degrees="36.530600" direction="north" orientation="latitude" precision="1" value="36.5306">36.530600</geoCoordinate>
,
<geoCoordinate id="2F563FACCC1022260BF566E2C9689EB8" degrees="6.211997" direction="west" orientation="longitude" precision="1" value="-6.211997">- 6.211997</geoCoordinate>
</named-content>
).
<location id="92C3D112C40BB9C07FE7D59E626956FF" LSID="urn:lsid:plazi:treatment:5AA43C0C196955F68835025C896920AB:92C3D112C40BB9C07FE7D59E626956FF" country="Spain" latitude="36.5306" longLatPrecision="1" longitude="-6.211997" name="See in Fig." stateProvince="Cadiz">See in Fig.</location>
<figureCitation id="7C62F0337B77648B3DBDF5EACE02DE0F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 E, F</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOVAG" type="Ecological remarks">
<paragraph id="A5C1418D99542432B471341786D5841C">
<heading id="79B9AC2BAAED8875E1E78E528970BA59" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="CE7B0ED3BA372DAE24E86B34ACB873B5">
Parasitoid of
<taxonomicName id="48EDF0A1837A784F1DE335AD69CA1251" authorityName="Waterston" authorityYear="1921" class="Insecta" family="Agaonidae" genus="Eupristina" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="verticillata">
<emphasis id="7C6F2AADEEF5EC3CB40B96AE2D4ADA0F" italics="true">Eupristina verticillata</emphasis>
</taxonomicName>
and
<taxonomicName id="C290378D4DD85C41F158178093BE6D9B" authorityName="Boucek" authorityYear="1993" class="Insecta" family="Agaonidae" genus="Walkerella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="microcarpae">
<emphasis id="3E80AE7F6E05A9E354FDADD886000A83" italics="true">W. microcarpae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="29700AE774D91F31FC197AEE2367D4DF" DOI="10.1111/icad.12282" author="Compton" etAl="et al." firstAuthor="Compton" journalOrPublisher="Insect Conservation and Diversity" pagination="341-351" refId="B16" refString="Compton SG, Chen X-Y, Chen Y, Hatcher MJ, Peng Y-Q, Quinnell RJ, Rodriguez LJ, Yu H, Ouyang A, Wei F-L, Cai Z-T, Wang R (2018) Host-parasitoid relationships within figs of an invasive fig tree: a fig wasp community structured by gall size. Insect Conservation and Diversity 11: 341351. https://doi.org/10.1111/icad.12282" title="Host-parasitoid relationships within figs of an invasive fig tree: a fig wasp community structured by gall size." volume="11" year="2018">Compton et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOWAG" type="distribution">
<paragraph id="2034E57DAE961A188A0B5A023D75E276">
<heading id="2961B7283C4ECE877D266CCEEA77995D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2C3A55D9E8C77B3D53A1D68B78FD9148">
This species originates from eastern Asia, namely
<collectingCountry id="B4916B44984A16442579F3E99A32C406" name="China">China</collectingCountry>
,
<collectingCountry id="19A1AE28557C620252771077562BEB14" name="Hong Kong">Hong Kong</collectingCountry>
,
<collectingCountry id="E44D24C4A5A51301199961CB97A5D17E" name="Japan">Japan</collectingCountry>
and
<collectingCountry id="53BA7908FF8BC928AFB6010708B8E666" name="Taiwan">Taiwan</collectingCountry>
(
<bibRefCitation id="5ECF4D7CBB4C79AD9BD99D8C025BDE49" author="van Noort and Rasplus" firstAuthor="van Noort" refId="B91" refString="van Noort S, Rasplus J-Y (2024) Figweb: figs and fig wasps of the world. www. figweb. org [Accessed on 1 May 2024]" url="www. figweb. org" year="2024">van Noort and Rasplus 2024</bibRefCitation>
). This is the first record from
<collectingCountry id="1404693D09EAF2EBBFB015E43DC17728" name="Spain">Spain</collectingCountry>
and the Iberian Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYWAG" type="description">
<paragraph id="0506D12BA8CE62C01493BDE07B3ED9B7">
<heading id="DB3F0F67624048CD7CF5445F7250724F" reason="title">Short description.</heading>
</paragraph>
<paragraph id="9BF044942AC8131EB49EE435D2270E05">Female yellowish red brown in general. Antennae brown except the scape which is yellowish red brown; abdominal segments 56 with transverse brown band at the base; segments 79 with a median longitudinal brown band; ovipositor black. Mandible tridentate the lower tooth largest. Mesoscutum and scutellum with a shallow median longitudinal furrow; propodeum with three longitudinal keels, the median one only indicated in the anterior half; Wings hyaline, the veins pale brown. Legs yellowish red-brown. Abdomen almost as long as the head and thorax combined, long ovate; stalk very short; ovipositor if the abdomen, dilated in the middle.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,113 @@
<document id="3E51F4E80183F877F59AA7B62516E6FF" ID-DOI="10.3897/zookeys.1244.148112" ID-ZooBank="CCCA8237-6587-44BD-A25E-41BD00928A11" ID-publisher-id="148112" URI-arpha="9A0DFA26-0AED-523E-937A-37D6A902E41E" URI-zoobank="http://zoobank.org/CCCA8237-6587-44BD-A25E-41BD00928A11" article-type="research-article" checkinTime="1751465287194" checkinUser="pensoft" docAuthor="Huang, Chao, Shih, Hsi-Te &amp; Ahyong, Shane T." docDate="2025" docId="5B3B1E37FE7C52C9956DD1C067AFCC3A" docLanguage="en" docName="ZooKeys 1244: 1-27" docOrigin="ZooKeys 1244" docSource="https://zookeys.pensoft.net/article/148112/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megapleonum Huang, Shih &amp; Ahyong 2018" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="27" masterDocId="9A0DFA260AED523E937A37D6A902E41E" masterDocTitle="Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="1" updateTime="1751465704327" updateUser="ExternalLinkService">
<mods:mods id="6EB79BCF4E36543CA3A42A3EE612A6EE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EBE7435F722BF4A24C93148E808A1EC8">
<mods:title id="376ED1265742D33680EA22BEAEF5B67D">Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="7633DA4F717D64D025FF618AE611ADEA" type="personal">
<mods:role id="FECC416A8B749CE65DCCCEF00B3B9555">
<mods:roleTerm id="F11FB9BD9F3D5E8070B62F7B27F51BC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="191E7BB7486F60F9151E43749E6D31ED">Huang, Chao</mods:namePart>
<mods:nameIdentifier id="149A8F5C9BF669F04ADFF47266D6CF9F" type="ORCID">0000-0002-3782-9096</mods:nameIdentifier>
<mods:affiliation id="F26912C63AEDC119E322E90129DFA15A">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="476ACC1A45A9EDD5B7262967A0B0821F" type="personal">
<mods:role id="042527806CAE21BB82D625B07F18EDA2">
<mods:roleTerm id="2C0E7B04BD3F2E7F525F7A8150EC85B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED9D0B6EEFAE88DD741C47711FBF5D45">Shih, Hsi-Te</mods:namePart>
<mods:nameIdentifier id="80D6D67BEDBEA575EA80C6E06A69FA87" type="ORCID">0000-0003-1317-8783</mods:nameIdentifier>
<mods:affiliation id="9FED97FAFF9BEFBE6BD61C7FF40EE8F1">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="6413D8C18E48B683180CFD0D34944336" type="personal">
<mods:role id="CEAC1780B293AC862ED6E6FAC0C6D477">
<mods:roleTerm id="4D6F44D6D7D607413AD920F2D47E4F38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1634DD3D9226D8E558BB988545A99FE8">Ahyong, Shane T.</mods:namePart>
<mods:nameIdentifier id="4D6CE4AD67B9FE077B3C08698AF68FFC" type="ORCID">0000-0002-2820-4158</mods:nameIdentifier>
<mods:affiliation id="8891E907B597DAA3CFB483072D79C5F1">Australian Museum, 1 William St, Sydney NSW 2010, Australia &amp; School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6401F9AF07E62D5AB09F9B62F0ABADB8">text</mods:typeOfResource>
<mods:relatedItem id="921333DF36697324E61CEB753879FC5E" type="host">
<mods:titleInfo id="FF5F1C9A55FCB0CA4E71A49BB95150ED">
<mods:title id="668382561827FAE66B082BB24A6FEBDF">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F3DA09AE8214C046B8807FEAC6C395A1">
<mods:date id="67FDD9C6FC79ADE72AAD508B453493F7">2025</mods:date>
<mods:detail id="7DF4162E9E53FEE49FB8331E9533FD4A" type="pubDate">
<mods:number id="3871F5613C752E0F9FB260BBD8BDD023">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="1F8C4C3C38444A1B4844BE714958F46D" type="volume">
<mods:number id="66C1580B4039A07DD660BA0AB1ACB333">1244</mods:number>
</mods:detail>
<mods:extent id="693569FC34EC0E73DA0FB193E9A540F7" unit="page">
<mods:start id="E889D8182DC80AD7C2BF3181AF69ADDC">1</mods:start>
<mods:end id="00AD411C28AF66A91CB452448B79EE3F">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D18BC1CD4829D5828A89E3AE0ABBB576">journal article</mods:classification>
<mods:identifier id="7578339EA3ED8A453FC5E1837D628BC9" type="DOI">10.3897/zookeys.1244.148112</mods:identifier>
<mods:identifier id="F9B9EDC197CF97D7840FE20095E1DFF4" type="ZooBank">CCCA8237-6587-44BD-A25E-41BD00928A11</mods:identifier>
</mods:mods>
<treatment id="5B3B1E37FE7C52C9956DD1C067AFCC3A" ID-DOI="http://doi.org/10.5281/zenodo.15791864" ID-Zenodo-Dep="15791864" ID-arpha="5B3B1E37-FE7C-52C9-956D-D1C067AFCC3A" LSID="urn:lsid:plazi:treatment:5B3B1E37FE7C52C9956DD1C067AFCC3A" httpUri="http://treatment.plazi.org/id/5B3B1E37FE7C52C9956DD1C067AFCC3A">
<subSubSection id="33CAEE116D497E956CBB0D07F7BFF2F3" type="nomenclature">
<paragraph id="B96B809B39BBED5ED0D90DE5FF31E181">
<taxonomicName id="628D23ED82BED2879AC40F5BA9C03408" ID-CoL="847WM" authority="Huang, Shih &amp; Ahyong, 2018" authorityName="Huang, Shih &amp; Ahyong" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="53EF1B603B65F00880F9A1B2D8A05400" italics="true">Megapleonum</emphasis>
<bibRefCitation id="6EB53769B4D0B11FACEC87AAFDA4C9CE" DOI="10.1093/jcbiol/ruy050" author="Huang C &amp; Shih HT &amp; Ahyong ST" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614-624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang, Shih &amp; Ahyong, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGJAC" type="diagnosis">
<paragraph id="C6BE4224F9E8D570C5C21191216626A2">
<heading id="AB990EB9B94AD77D2E8D1D58C01F98B5" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7B068BCCEE8CAC0A4BAAEA32F45C375D">Carapace broader than long, dorsal surface slightly convex; postorbital, epigastric cristae confluent or almost confluent; external orbital angle bluntly triangular, confluent or almost confluent with anterolateral margin. Epistome buccal margin with triangular median lobe. Maxilliped 3 ischium longitudinally subquadrate, length ~ 0.7 × width; exopod reaching slightly beyond distal edge of ischium, flagellum absent to well-developed. Male anterior thoracic sternum very broad, width 1.82.0 × length. Male pleon large, broadly triangular to sublinguiform, tip of telson rounded. Female pleon linguiform to subovate. G 1 large, sinuous, reaching beyond male pleonal locking tubercle, subterminal segment outer margin slightly to strongly concave, terminal segment highly variable. G 2 subterminal segment tapering, bent to slightly bent outwards distally, thin to thick flagelliform terminal segment. Vulvae ovate, large, reaching suture of sternites 5 / 6, relatively widely separated.</paragraph>
</subSubSection>
<subSubSection id="SECID0ELJAC" type="Composition">
<paragraph id="FEEDF3FFE0764321FEDF13091F8E1123">
<heading id="9B08B4342C53133F5B1F5EB471BBFC1C" reason="title">Composition.</heading>
</paragraph>
<paragraph id="DD9D388821531D35399DA0A012B43098">
<taxonomicName id="E945AACCCE0D5332BF4331A079F041FE" authority="Huang, Shih &amp; Ahyong, 2018" authorityName="Huang, Shih &amp; Ahyong" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ehuangzhang">
<emphasis id="4062220E7E9E75CA58C95BFA81C1DEE4" italics="true">Megapleonum ehuangzhang</emphasis>
<bibRefCitation id="FFBE14E5F93E80687B4500625D86E7DC" DOI="10.1093/jcbiol/ruy050" author="Huang C &amp; Shih HT &amp; Ahyong ST" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614-624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang, Shih &amp; Ahyong, 2018</bibRefCitation>
</taxonomicName>
(
<typeStatus id="BCA19B1FF3C4BF5DBCE7C4580131E089">type</typeStatus>
species),
<taxonomicName id="6230D570744E704F8B4BCC2E0F6C25D6" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="DB246BFF4195AE1ED124CACE21844860" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AE4FE3C3B960EA50CF32CAD457A3FC8" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="F05E2266AFDC263B5B1421596EF3B1BB" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="DAE152B0E783E5FDDC0DC77C045A03D4" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9CBE1AB7BEF0A951723B097B1A48256" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="C95032506AE45FAE225F2089A695359A" authority="Huang &amp; Mao, 2021" authorityName="Huang &amp; Mao" authorityYear="2021" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="shenzhen">
<emphasis id="F96121155E180AC779B1AA0849C26D86" italics="true">Megapleonum shenzhen</emphasis>
<bibRefCitation id="E8534245CCC840E3B66F4445A283AB44" DOI="10.6620/ZS.2021.60-66" author="Huang C &amp; Mao SY" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang &amp; Mao, 2021</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B3D59301EDDF2D2809CB3BC730E1F94D" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="4E1384AEE4D7274039FA29B96599F7FB" italics="true">M. yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF858715E6A0A79A4689B7A9C2BAB6D" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="A6DB458558835BFF26F4924102136BA3" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="F03E7E189047DC5C7255A2FF3B30068B" italics="true">M. wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB97ED0D39D5D901993592C5C05B1C42" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETLAC" type="remarks">
<paragraph id="E3CB993D9D92E1AC7F654DD732B81F5A">
<heading id="F7E2B0CFF60B4E7C98320A46BF12195D" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="E0F476EFD3E44A34CE82FF708FD14D13">With the description of four new species herein, the diagnosis of the genus has to be revised. The significant variation within the genus, particularly in the form of the G 1 poses challenges to establishing a uniform diagnosis. However, the key distinguishing features include the confluent or almost confluent postorbital and epigastric cristae, and external orbital angle and anterolateral margin, very broad male anterior thoracic sternum, large male pleon and the ovate, large vulvae, which extend to the suture of sternites 5 / 6 and are relatively widely spaced.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,257 @@
<document id="CD2BDB5DA191151B5E8DBA9954C11575" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="6E389EC2FB145A7CA3F0D69B1FB30D87" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Walkerella microcarpae Boucek 1993" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="BC3077756227178191E794BC73922947" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="92AD6A4A7B9FC7367B2BD0282FB6F60E">
<mods:title id="2B423C373D9BA5BF618A51926B91F450">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="C0E9BE0BDE89126F370B4D3A53640C81" type="personal">
<mods:role id="2255C48AA69D070C110B1C482A7ADBEE">
<mods:roleTerm id="B89BF891A6F1332B0D332F1DD4825C55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDA48D8F1C3638F7A79FAC740F174B78">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="73DE2EBF8F5F844A39E5797201023021" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="5EEF64B0FEC1A83AECDF2BBF40D84178">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="73748DF45A87CA8D0148943538E884A2" type="personal">
<mods:role id="D204471C678A67BFF5A625DADAAF2BFC">
<mods:roleTerm id="A7FEC74CABE1A576BA127A1AE844A280">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1EE8480D60AD7CA2D4C69884B667B512">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="5955083185103FA11495757B2E2EFACC" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="8AA3DAE4163A642030AA509F017FF486">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="DD0A7D44210803947F3238DC79C35B31" type="personal">
<mods:role id="07B81E1CD99F3880BED801630490BBC6">
<mods:roleTerm id="2D0F9C9625F759F5C92A6004A62DCED6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D734040C105219E5E658C29FC630F104">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="AF7AB43966F6B64764DBA6AA5EA4C0F7" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="DB6C768366C5C0230A00E30B725C2E86">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="C0727A1EAE787709EA5B61B2D3803632" type="personal">
<mods:role id="4D5BD232E0995A7B3BA23BB8AF661B2F">
<mods:roleTerm id="CF429A534A855E58BB79080009463189">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5A356EF9F1522535921718749653136">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="1F89808753B0A7FAEA23D7E54ABA9A12" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="CA3369C976E545DFD464D1C0DD2DE818">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="7F95D9EEE7D801E4C22FE61686F93662" type="personal">
<mods:role id="5193C5EFAE190FBEA735B6B56BC776B6">
<mods:roleTerm id="84DE8C70D1ADEC2E9787233DA49392AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECCFF9717B6B70893401A17FFF6FA2B0">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="1FA6203155F14583B9BB251F12ADE687" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="890CCACB1FC74F5E147475C6CB1651EC">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="96BDD86E1F8708FBADF7C04F21214F84">text</mods:typeOfResource>
<mods:relatedItem id="F9BA6450B3154654170884AA6D64D355" type="host">
<mods:titleInfo id="3C69F8A9833E82731057BF7FE8A6F554">
<mods:title id="BC75625FFD4EAFEAEC6D53B3404913CE">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="6D73F32D94697E51BDAE455ECC00ECBF">
<mods:date id="A77C2170F4CD2A6845D84CDFDE033457">2025</mods:date>
<mods:detail id="AB567DE4C33039C3269CA60BDB2BEE41" type="pubDate">
<mods:number id="73D6E8F7BAEAF6376534C4BA44B5366B">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="E293CA83FDAC1868ED19C51CA1903561" type="volume">
<mods:number id="5419D0152FD5D426603D03994177B388">98</mods:number>
</mods:detail>
<mods:extent id="2E2B5F098CC7647C2D60453C283B1DB5" unit="page">
<mods:start id="08D2792776B848AA97783ED60BF6D4F8">667</mods:start>
<mods:end id="A9374B9025E09E661336165B351B52A0">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2394011A031869102215AF313B8D00C1">journal article</mods:classification>
<mods:identifier id="1FAEBA74EA2C20CB9C9F2F6E103DA6C5" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="69ACBFA50883979E2FAF313A7EE9F0D6" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="6E389EC2FB145A7CA3F0D69B1FB30D87" ID-DOI="http://doi.org/10.5281/zenodo.15791858" ID-Zenodo-Dep="15791858" ID-arpha="6E389EC2-FB14-5A7C-A3F0-D69B1FB30D87" LSID="urn:lsid:plazi:treatment:6E389EC2FB145A7CA3F0D69B1FB30D87" httpUri="http://treatment.plazi.org/id/6E389EC2FB145A7CA3F0D69B1FB30D87">
<subSubSection id="9BDF69F74894F45FA5CD5836C06FB6B4" type="nomenclature">
<paragraph id="0513EA5BF967C15D75475C11D2D101D8">
<taxonomicName id="BC5DCF4BFFCDF43C1EDC17EEDBF52184" ID-CoL="5BTYN" authority="Boucek, 1993" authorityName="Boucek" authorityYear="1993" class="Insecta" family="Agaonidae" genus="Walkerella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="microcarpae">
<emphasis id="E55E59688BC6477096BF61E0AA5F4F2C" italics="true">Walkerella microcarpae</emphasis>
<bibRefCitation id="204DA436221076DCD8418AC62CF1FB49" DOI="10.1080/00222939300770071" author="Bouček Z" journalOrPublisher="Journal of Natural History" pagination="173-217" refId="B6" refString="Boucek Z (1993) The genera of chalcidoid wasps from Ficus fruit in the New World. Journal of Natural History 27: 173-217. https://doi.org/10.1080/00222939300770071" title="The genera of chalcidoid wasps from Ficus fruit in the New World." volume="27" year="1993">Bouček, 1993</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5MAG" type="material">
<paragraph id="4C4B1F28B0284A087E054B1F3E499FD4">
<heading id="27CEB283FB223F1A5B3092AF1DA94FF9" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="7EA5F24612574BFC68D4365D9F9FB464">
<materialsCitation id="503093A8D685E5D5605616BC3EE64F65" collectingDate="2018-12-07" collectionCode="MNCN" collectorName="Jerez de la Frontera &amp; de Jerez &amp; I. Sanchez &amp; Ent" country="Spain" elevation="82" latitude="36.689316" location="Zoobotanico de Jerez" longLatPrecision="1" longitude="-6.150638" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Cadiz">
<emphasis id="E48895D6846B30555F024D6D507477BD" bold="true">
<collectingCountry id="8FCBFC00F39C2B783D569801F018A69B" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="5A981184FA465C53D91940A9F7B097C0" bold="true">
<collectingRegion id="13E7537DC9A13F46795F829D8FC78FDF">Cádiz</collectingRegion>
</emphasis>
:
<collectorName id="C1F2DCE3BF16B366B078246C1EC0AF38">Jerez de la Frontera</collectorName>
(
<location id="074C7A75529F1CBE3C0E03B4EA168FA1" LSID="urn:lsid:plazi:treatment:6E389EC2FB145A7CA3F0D69B1FB30D87:074C7A75529F1CBE3C0E03B4EA168FA1" country="Spain" latitude="36.689316" longLatPrecision="1" longitude="-6.150638" name="Zoobotanico de Jerez" stateProvince="Cadiz">
Zoobotánico
<collectorName id="950A9244AC082BDBE5E9B31F9C678D85">de Jerez</collectorName>
</location>
):
<specimenCount id="7B9BDDA66F144726645E56469D4A2021" count="3" type="male">3 ♂</specimenCount>
+
<specimenCount id="8E4F406492BCC681B02B2183BE8A8698" count="3" type="female">3 ♀</specimenCount>
<abbrev id="ABBRID0EINAG" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="149A3DF08327AB3AB506E05D726203E8">Í. Sánchez</collectorName>
<abbrev id="ABBRID0EMNAG" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="F86AC665F276120005E00596F9F0D1A9" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
_
<collectorName id="E6819869CD522765B90CFE6017051790">Ent</collectorName>
235063
<abbrev id="ABBRID0EQNAG" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="F719BBC0363BF0DF4FCB89BF86D4A36E" value="2018-12-07">
<collectingDate id="E7CA9CB8E07EBD521A237087363B99F1" value="2018-12-07">7 - XII- 2018</collectingDate>
</date>
,
<elevation id="7EB82BE8AC06AB078C31F2D5E3F2A039" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">
<quantity id="65201D697A11723CA4C3D5B2D73F6455" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">82 m</quantity>
a. s. l.
</elevation>
, found in dissected syconia of
<taxonomicName id="55161CB15D2709C8B2C8F23059BCB83F" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="42D04392007A7A27083F027F9DA6ADE4" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECOAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-6.150638,36.689315]}">
<geoCoordinate id="566E1867575D94FE686DA294D1C47833" degrees="36.689315" direction="north" orientation="latitude" precision="1" value="36.689316">36.689315</geoCoordinate>
,
<geoCoordinate id="6812694E25CC9E3E7FE69F0F1814D8EB" degrees="6.150638" direction="west" orientation="longitude" precision="1" value="-6.150638">- 6.150638</geoCoordinate>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="43530A1391B7BF0A86EA55E11892AC50" collectingDate="2023-07-05" collectionCode="AC" collectorName="de Alicante &amp; J. M. Royo" country="Spain" county="San Vicente del Raspeig" elevation="92" latitude="38.384266" location="See in Fig." longLatPrecision="1" longitude="-0.512455" municipality="Universidad de Alicante" specimenCount="16" specimenCount-female="11" specimenCount-male="5" stateProvince="Alicante">
<emphasis id="E30CC5005CB12F731A61B0BCF2D4BB40" bold="true">
<collectingRegion id="175001E268C239D97D0E862250A236DE">Alicante</collectingRegion>
</emphasis>
:
<collectingCounty id="9A2A945C735586C9DAF61BD8A3B3EB73">San Vicente del Raspeig</collectingCounty>
(
<collectingMunicipality id="F8817F2DB403F2E11F3B53673F3A526B">
Universidad
<collectorName id="32E9D0F8B9DA4640CBF338CFC69A3A69">de Alicante</collectorName>
</collectingMunicipality>
):
<specimenCount id="089DDA0464024E611DEAE521ED00228A" count="11" type="female">11 ♀</specimenCount>
+
<specimenCount id="359728A70932BE2002D9FDEDF11901C7" count="5" type="male">5 ♂</specimenCount>
<abbrev id="ABBRID0EJOAG" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="3F1AA9EF8CFF51B9C19E880ED730B20A">J. M. Royo</collectorName>
<abbrev id="ABBRID0ENOAG" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="395FA02F277FA93EA3CE9F48CCCC0C8C" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EROAG" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="4913DC7CD67C8F4FA32B9F3A382D2F21" value="2023-07-05">
<collectingDate id="4EC05D34A18378DEFEE90A71781577A1" value="2023-07-05">5 - VII- 2023</collectingDate>
</date>
,
<elevation id="C28F4B9DF1B6A00E970A9221B3DF6E91" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">
<quantity id="F8445AAA204DA9FA8E6C2C376CBD8517" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">92 m</quantity>
a. s. l.
</elevation>
, found dissecting fallen syconia of
<taxonomicName id="2D32F4F81E0A7E799B76B2C7FBDAEAF7" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="3AEEBF529EDD763E2945533123A2D951" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
and some flying specimens close to the tree (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDPAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-0.512455,38.384264]}">
<geoCoordinate id="7464453355F47C40229A49FDF5CDE48F" degrees="38.384264" direction="north" orientation="latitude" precision="1" value="38.384266">38.384264</geoCoordinate>
,
<geoCoordinate id="4A17B9A93385DE0A70A30FE7D63EF59D" degrees="0.512455" direction="west" orientation="longitude" precision="1" value="-0.512455">- 0.512455</geoCoordinate>
</named-content>
).
<location id="B7A0AA7F764625DA2E76798740B2CC51" LSID="urn:lsid:plazi:treatment:6E389EC2FB145A7CA3F0D69B1FB30D87:B7A0AA7F764625DA2E76798740B2CC51" country="Spain" county="San Vicente del Raspeig" latitude="38.384266" longLatPrecision="1" longitude="-0.512455" municipality="Universidad de Alicante" name="See in Fig." stateProvince="Alicante">See in Fig.</location>
<figureCitation id="BB4A81B20C9A168E733BFF7EDBA9AD0E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 B</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMPAG" type="Ecological remarks">
<paragraph id="9CF93753786F3F3CFCD8D70A87E0977D">
<heading id="C9D359A0DDFDB776D518E5AAE76D9766" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="1FA66465E0E6FD5AEAE1CFA50F9DE8AF">
Gall-forming wasp in figs of
<taxonomicName id="5FF20085D8EF8F4F785DA38855549073" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="6865889DC07F2B8580E6EA1193537FB3" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4PAG" type="distribution">
<paragraph id="3D94B7553BBB5B528734D0636A89E14D">
<heading id="003872945A77FAA74EA3186ED9549392" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="CC69F898FBE8D5F24CCEEE0A96E6FCC0">
This species probably originates from the Oriental region and was introduced to the Palearctic and the Nearctic (
<bibRefCitation id="85D8315AD3AD746AAB1C86171EFBFBE4" DOI="10.11646/zootaxa.3702.5.6" author="Ma" etAl="et al." firstAuthor="Ma" journalOrPublisher="Zootaxa" pagination="473-482" refId="B65" refString="Ma YC, Peng YQ, Yang DR (2013) Description of two new species of Walkerella (Pteromalidae, Otitesellinae) from China with a key to species of the genus. Zootaxa 3702: 473482. https://doi.org/10.11646/zootaxa.3702.5.6" title="Description of two new species of Walkerella (Pteromalidae, Otitesellinae) from China with a key to species of the genus." volume="3702" year="2013">Ma et al. 2013</bibRefCitation>
). In Europe, it is only known from the Canary Islands (Tenerife) (
<bibRefCitation id="F589F7994C73DA3592B1EDE04BA98CC0" author="Carnero Hernandez" etAl="et al." firstAuthor="Carnero Hernandez" refId="B11" refString="Carnero Hernández A, Hernández García M, Pérez Padrón F, Hernández Suárez E (1998) Walkerella microcarpae and Eupristina (Parapristina) verticillata (pollinator agent) cause fruit ripening of Old World fig (Ficus species) in the Canary Islands. Boletin de la Asociacion Espanola de Entomologia 22 (34): 255." year="1998">Carnero Hernandez et al. 1998</bibRefCitation>
), Majorca (
<bibRefCitation id="7AD8A0F85730C502FF102AC33AD9237D" DOI="10.1111/icad.12111" author="Wang R &amp; Aylwin R &amp; Barwell L &amp; Chen XY &amp; Chen Y &amp; Chou LS &amp; Cobb J &amp; Collette D &amp; Craine L &amp; Giblin-Davis RM &amp; Ghana S &amp; Harper M &amp; Harrison RD &amp; McPherson JR &amp; Peng Y-Q &amp; Pereira RAS &amp; Reyes-Betancort A &amp; Rodríguez LJV &amp; Strange E &amp; van Noort S &amp; Yang H-W &amp; Yu H &amp; Compton SG" issue="4" journalOrPublisher="Insect Conservation and Diversity" pagination="322-336" refId="B95" refString="Wang R, Aylwin R, Barwell L, Chen XY, Chen Y, Chou LS, Cobb J, Collette D, Craine L, Giblin-Davis RM, Ghana S, Harper M, Harrison RD, McPherson JR, Peng Y-Q, Pereira RAS, Reyes-Betancort A, Rodríguez LJV, Strange E, van Noort S, Yang H-W, Yu H, Compton SG (2015 a) The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa. Insect Conservation and Diversity 8 (4): 322336. https://doi.org/10.1111/icad.12111" title="The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa." volume="8">Wang et al. 2015 a</bibRefCitation>
), as well as
<collectingCountry id="7A908A9781E8DAE5576D67E520F378B2" name="Cyprus">Cyprus</collectingCountry>
(
<bibRefCitation id="B4CC64941FC1232FA6BC995F35CA900E" DOI="10.3391/bir.2023.12.2.20" author="Demetriou" etAl="et al." firstAuthor="Demetriou" issue="2" journalOrPublisher="BioInvasions Records" pagination="573-580" refId="B30" refString="Demetriou J, Koutsoukos E, Mavrovounioti N, Radea K, Arianoutsou M, Roy EH, Compton SG, Martinou A (2023) A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus. BioInvasions Records 12 (2): 573580. https://doi.org/10.3391/bir.2023.12.2.20" title="A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus." volume="12" year="2023">Demetriou et al. 2023</bibRefCitation>
) and
<collectingCountry id="31C67550657B3AFF3C8C68BF9DA03812" name="Greece">Greece</collectingCountry>
(
<bibRefCitation id="2051C6FE422E55C518AAF7CFF83C9A26" DOI="10.3897/neobiota.98.139650" author="Koutsoukos" etAl="et al." firstAuthor="Koutsoukos" journalOrPublisher="NeoBiota" pagination="93-115" refId="B60" refString="Koutsoukos E, Konstantopoulou A, Parmakelis A, Demetriou J, Compton SG, Georgiadis C, Constantinidis T (2025) The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece. NeoBiota 98: 93115. https://doi.org/10.3897/neobiota.98.139650" title="The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece." volume="98" year="2025">Koutsoukos et al. 2025</bibRefCitation>
). As the reports by
<bibRefCitation id="0D38A5D53D9B790E923F406EC4D029FE" author="Falcó-Garí JV &amp; Oltra-Moscardo MT &amp; Peris-Felipo FJ &amp; Jiménez-Peydro R" etAl="et al." firstAuthor="Falco-Gari" refId="B32" refString="Falcó-Garí JV, Oltra-Moscardo MT, Peris-Felipo FJ, Jiménez-Peydro R (2010) Exotic fig pollinators and associates (Chalcidoidea: Agaonidae) of Ficus trees in Europe. Seventh international Congress of Hymenopterists. Hungary, Programme and Abstracts." year="2010">Falcó-Garí et al. (2010)</bibRefCitation>
from eastern Iberian Peninsula have no localities, we consider this as a first confirmed record for the Iberian Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2QAG" type="description">
<paragraph id="974AB5BEE59B373F0CB71B640AB641B9">
<heading id="C5253FD0BFE34A1840C7708DCAA12ED5" reason="title">Short description.</heading>
</paragraph>
<paragraph id="DC0CA8660A2ED0E5D67385CA7B794521">
Difficult to separate morphologically from other genera of European
<taxonomicName id="B83C33C5325CD2E6843C1FC932F7D29E" authorityName="Dalman" authorityYear="1820" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">Pteromalidae</taxonomicName>
.
<taxonomicName id="570ACA33BC70652D0A33C8E491F83EBD" authorityName="Westwood" authorityYear="1883" class="Insecta" family="Agaonidae" genus="Walkerella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="AF003718C8494DEA57D813234328B852" italics="true">Walkerella</emphasis>
</taxonomicName>
is recognized by the following characters: body blackish, without conspicuous external ovipositor and apterous males with long mandibles. The genus
<taxonomicName id="AC9CC54E38D9D8E1A3C46C2192A3CABD" class="Insecta" family="Agaonidae" genus="Otitesella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="3ABBAD1B5D8DE15F7E58698365C2B96A" italics="true">Otitesella</emphasis>
</taxonomicName>
is also present in the Palearctic region (associated with
<taxonomicName id="6DF52B26ABDB8CB58547D96500732962" authorityName="Motschulsky" authorityYear="1863" class="Insecta" family="Agaonidae" genus="Platyscapa" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0DDFF9685D47BECF56C44084B0762BF0" italics="true">Platyscapa</emphasis>
</taxonomicName>
). Also, the genus
<taxonomicName id="4A86F4AFE5D17F4B8DBC56E3B6003414" class="Insecta" family="Agaonidae" genus="Micranisa" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="08DACC25780B3CB961F5B2943DE781C8" italics="true">Micranisa</emphasis>
</taxonomicName>
from
<collectingCountry id="1D14AAE2EF65FD1B080AC9D40A5C9E30" name="Greece">Greece</collectingCountry>
and
<collectingCountry id="CFF917596F323FE5029314FF9B2198ED" name="Cyprus">Cyprus</collectingCountry>
(
<bibRefCitation id="CCF0C9E71D776FF1524C62A388132574" DOI="10.3391/bir.2023.12.2.20" author="Demetriou" etAl="et al." firstAuthor="Demetriou" issue="2" journalOrPublisher="BioInvasions Records" pagination="573-580" refId="B30" refString="Demetriou J, Koutsoukos E, Mavrovounioti N, Radea K, Arianoutsou M, Roy EH, Compton SG, Martinou A (2023) A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus. BioInvasions Records 12 (2): 573580. https://doi.org/10.3391/bir.2023.12.2.20" title="A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus." volume="12" year="2023">Demetriou et al. 2023</bibRefCitation>
;
<bibRefCitation id="2D7363C824D90E89EF64C19E14E6137D" DOI="10.3897/neobiota.98.139650" author="Koutsoukos" etAl="et al." firstAuthor="Koutsoukos" journalOrPublisher="NeoBiota" pagination="93-115" refId="B60" refString="Koutsoukos E, Konstantopoulou A, Parmakelis A, Demetriou J, Compton SG, Georgiadis C, Constantinidis T (2025) The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece. NeoBiota 98: 93115. https://doi.org/10.3897/neobiota.98.139650" title="The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece." volume="98" year="2025">Koutsoukos et al. 2025</bibRefCitation>
), which is quite close to
<taxonomicName id="D913DB7C52EC420DD526B75300999A2E" authorityName="Westwood" authorityYear="1883" class="Insecta" family="Agaonidae" genus="Walkerella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="503E3449C98FF538A15F2C664834C83F" italics="true">Walkerella</emphasis>
</taxonomicName>
, although is easily separated by its ovipositor which is extended and curved downwards.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,263 @@
<document id="808D97A8956DA5437B7182D00E4EC1EB" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="79617C4480935728A52E2471BDD883F7" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eufroggattisca okinavensis" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="15BFF15787182994A8F78FDEE1302880" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E303138232DE21F6BC9163EE2A65489">
<mods:title id="ECBE79D72E8AB13BCA5FB72FBEFB6850">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="A8F50AA2B950CD7D9C6C4519A119282F" type="personal">
<mods:role id="5663823956C513261C2437DB277296AE">
<mods:roleTerm id="470D52B7E0032EE5E6DE65DF43D933E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="021DA9C2EDC6CFF2F6A8E9BD7C6CBF1C">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="9AC0E5357A48970C0EE7C0D4F977DCAC" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="31832E80A423FAEC786E04A19897795D">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="A0A64017DBA70623539D5DCF5179E3EC" type="personal">
<mods:role id="222CA987DC18A5863987753094F95C62">
<mods:roleTerm id="14AF2017B2C0399C5F1EB41EF3864857">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E22B8AFCBDDA40AEEFE39B0344025270">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="7256838AFDA33A67DFB151F7B8977A8B" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="8034F2F74A35592D0580DCBBCB1F837F">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="A0F6508500EA30A1EB4E8880AF2B3A67" type="personal">
<mods:role id="C861C438846553EA6185BF3603C9CAC5">
<mods:roleTerm id="9D3F1B607A96C1B98E856969D37EB213">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE9C0DAD0C370F2BE5386CD28D27EC78">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="B0D4CA8689B37E0127C9A32284C0A3BD" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="47AD27D5C16B8DC35DE4D9C9B72500C5">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="090A149C03BF22C470260898005FBF0A" type="personal">
<mods:role id="B1EB22E849631139EC96F29DB019C8FC">
<mods:roleTerm id="BDCD15FF30D5B5BA5252D1906C3E7337">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E9662E4900947E424C7819E9DE6E3EB">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="99A87A8E2798BF04F126E0913907D2F4" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="0CCB86619D0473F262E6B87A8BA81674">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="33F2F8D141EB268D7E809B69E5725EF2" type="personal">
<mods:role id="4728F53F562BB0701BC88AEA54E31D50">
<mods:roleTerm id="2E1A1B6E636179EAC37D23396B92CC1D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="350AB429083BB93B25FA476FF72DBD0B">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="A52195F40A809FA77B23F922E12E1A98" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="307459E4CED9DAEA3DF91B26F36E0533">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="282CA1F865B63D0BE06A5C8014169261">text</mods:typeOfResource>
<mods:relatedItem id="019B438134CEEFAD464D6DABA791FBB8" type="host">
<mods:titleInfo id="C546218CE524534F10A5EFBA2B7E7AAC">
<mods:title id="A113644FA58E8E1D04874212C9C72584">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="FD3A8402021FD949235770A4975EA0F8">
<mods:date id="219D4EB688FDCF025C2609FA6E44A147">2025</mods:date>
<mods:detail id="AF9EAA09357C568B18689179D6D604CF" type="pubDate">
<mods:number id="F2A652D322824F0A485A97A84089C96F">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="4FF3461A55A0C4FAFDBB3F64DE953D40" type="volume">
<mods:number id="EC57CFF78C52517915B61C44D92C9264">98</mods:number>
</mods:detail>
<mods:extent id="F4F9763CD5533DCEED2DE3A428466D0C" unit="page">
<mods:start id="D65A15F3680728CC9CB75F1B592319E4">667</mods:start>
<mods:end id="1319941C9C533715C3F96C3DB3FDDA85">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5A3B73351D764A487E3F9B03B3991CA6">journal article</mods:classification>
<mods:identifier id="F2FD787472B0569EB476293E84AC8024" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="926B4CC1C0D0B18AA2E4AECC4F434723" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="79617C4480935728A52E2471BDD883F7" ID-DOI="http://doi.org/10.5281/zenodo.15791854" ID-Zenodo-Dep="15791854" ID-arpha="79617C44-8093-5728-A52E-2471BDD883F7" LSID="urn:lsid:plazi:treatment:79617C4480935728A52E2471BDD883F7" httpUri="http://treatment.plazi.org/id/79617C4480935728A52E2471BDD883F7">
<subSubSection id="59E37058350A7068623BFAEA0EA7812A" type="nomenclature">
<paragraph id="5C9D62B976526DF4DF3607C2C4F6263A">
<taxonomicName id="4DDA181D2A551872E6B43C8330C26D33" ID-CoL="3C3K4" authority="(Ishii, 1934)" baseAuthorityName="Ishii" baseAuthorityYear="1934" class="Insecta" family="Agaonidae" genus="Eufroggattisca" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="okinavensis">
<emphasis id="6D75325229CAF50683E71B6FA3AE9723" italics="true">Eufroggattisca okinavensis</emphasis>
(
<bibRefCitation id="5AEB286AD19B2FDC8717BD405C7FF28C" author="Ishii T" journalOrPublisher="Kontyû" pagination="84-100" refId="B47" refString="Ishii T (1934) Fig. chalcidoids of Japan. Kontyu 8: 84-100." title="Fig. chalcidoids of Japan." volume="8" year="1934">Ishii, 1934</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EV4AE" type="material">
<paragraph id="DED96B6EB5F5CDB4E1028B8B133FEE65">
<heading id="1E7F6E530BE15E97993AA9CB5F49BF11" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="B577428D77E9ADE5507CA446A569C34C">
<materialsCitation id="3A8B7E5A65D972000B252856F576BC64" collectingDate="2018-12-07" collectionCode="MNCN" collectorName="Jerez de la Frontera &amp; de Jerez &amp; I. Sanchez &amp; Ent" country="Spain" elevation="82" latitude="36.689316" location="Zoobotanico de Jerez" longLatPrecision="1" longitude="-6.150638" municipality="Jerez de la Frontera" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Cadiz">
<emphasis id="838FADC57DD1B1220AB82336BC3B32A9" bold="true">
<collectingCountry id="F847A97D02CD16E286EB27872B118BE1" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="47853BDC9B80356E553296A85062431E" bold="true">
<collectingRegion id="AD99FC422426908144E7E97F19796C8F">Cádiz</collectingRegion>
</emphasis>
:
<collectorName id="FA24B19154DAF3867CE48761D3F5DF71">
<collectingMunicipality id="E1AA728D3D671EBF2E4663AED4C9CFC3">Jerez de la Frontera</collectingMunicipality>
</collectorName>
(
<location id="810BAC0194AD638EEAB369690291EFA3" LSID="urn:lsid:plazi:treatment:79617C4480935728A52E2471BDD883F7:810BAC0194AD638EEAB369690291EFA3" country="Spain" latitude="36.689316" longLatPrecision="1" longitude="-6.150638" municipality="Jerez de la Frontera" name="Zoobotanico de Jerez" stateProvince="Cadiz">
Zoobotánico
<collectorName id="43645EE28F3546EAD13D927AB7FEA16A">de Jerez</collectorName>
</location>
):
<specimenCount id="3F4C3D7CBFC91237A7464B75243BB241" count="2" type="male">2 ♂</specimenCount>
+
<specimenCount id="7526F3BFDF65D95F4D3141E07D4C77A8" count="3" type="female">3 ♀</specimenCount>
<abbrev id="ABBRID0E64AE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="1E71AA78F93853583BDFA41A712F82F6">Í. Sánchez</collectorName>
<abbrev id="ABBRID0ED5AE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="9E7866FAEA5030E9234074F323D70B39" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
_
<collectorName id="72CB526EBD676CA309B9B0FED6C2289E">Ent</collectorName>
235061
<abbrev id="ABBRID0EH5AE" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="0BE0BE79E8A535925EC2CAAC3268FA67" value="2018-12-07">
<collectingDate id="D02BFC5FE392E9621F26E2D8EA7B5F2D" value="2018-12-07">7 - XII- 2018</collectingDate>
</date>
,
<quantity id="D50523F1ADAD5E94038FB724D224143E" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">
<elevation id="6F18579D605EC5D1A73306CD3D104D7F" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">82 m</elevation>
</quantity>
a. s. l, found dissecting syconia of
<taxonomicName id="0BE2026D004B4046777BAB2281245D59" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="0036F9BB6AFC3209141F41AE0C65D59F" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZ5AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-6.150638,36.689315]}">
<geoCoordinate id="56254299A0817964AD11428769068260" degrees="36.689315" direction="north" orientation="latitude" precision="1" value="36.689316">36.689315</geoCoordinate>
,
<geoCoordinate id="BF714649D54C9A0D7BB1F67858FF2667" degrees="6.150638" direction="west" orientation="longitude" precision="1" value="-6.150638">- 6.150638</geoCoordinate>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="A203F1A4930306D011D4495A5AF95548" collectingDate="2024-05-19" collectionCode="AC" collectorName="de Alicante &amp; J. M. Royo" country="Spain" elevation="92" latitude="38.384266" location="Universidad de Alicante" longLatPrecision="1" longitude="-0.512455" municipality="San Vicente del Raspeig" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Alicante">
<emphasis id="14B313A5E1E3B7572ABEE44EE26B9CEF" bold="true">
<collectingRegion id="3D483BC91D48C7A89E23BADAB046A43A">Alicante</collectingRegion>
</emphasis>
:
<collectingMunicipality id="5D2256F9C2D22D7A5B427AD200C9C4F5">San Vicente del Raspeig</collectingMunicipality>
(
<location id="83A4996A293D1BB5221FA5F5B4825B1A" LSID="urn:lsid:plazi:treatment:79617C4480935728A52E2471BDD883F7:83A4996A293D1BB5221FA5F5B4825B1A" country="Spain" latitude="38.384266" longLatPrecision="1" longitude="-0.512455" municipality="San Vicente del Raspeig" name="Universidad de Alicante" stateProvince="Alicante">
Universidad
<collectorName id="508D3B80EE93B8D74F225E861F74207F">de Alicante</collectorName>
</location>
):
<specimenCount id="14197529747929B12B2E3B63F4EFEB0F" count="10" type="female">10 ♀</specimenCount>
+
<specimenCount id="4AF9552CA23595CB5400C3360DF74087" count="10" type="male">10 ♂</specimenCount>
<abbrev id="ABBRID0EA6AE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="15685B7F0A2D9D074F6960C922AE1849">J. M. Royo</collectorName>
<abbrev id="ABBRID0EE6AE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="43C428B9B741FF98B66240F7091CF566" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EI6AE" xlink_title="collection of deposit">deposit</abbrev>
.
<date id="6C05B0753239BE97123EA0094186DB16" value="2024-05-19">
<collectingDate id="69A270AFE3E1E8EF3D1CD0202B289D2B" value="2024-05-19">19 - V- 2024</collectingDate>
</date>
,
<quantity id="E25D1993185B93615154BB47F152F634" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">
<elevation id="2ED194C99AC66A516B7AEEA84EA15E7C" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">92 m</elevation>
</quantity>
a. s. l, found dissecting syconia of
<taxonomicName id="C393B417413952B64214E1D937E68841" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="CFA46366EDE1CDE0701782F3200B8EC3" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E16AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-0.512455,38.384264]}">
<geoCoordinate id="0016518B5A08D3834D75EFAF588FD689" degrees="38.384264" direction="north" orientation="latitude" precision="1" value="38.384266">38.384264</geoCoordinate>
,
<geoCoordinate id="5E3ABE7DC5C5E7428E5005B9A9376FC0" degrees="0.512455" direction="west" orientation="longitude" precision="1" value="-0.512455">- 0.512455</geoCoordinate>
</named-content>
). See Fig.
<figureCitation id="C3084013C81CFC436A650AAD4FE9ECC2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 G</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEAAG" type="Ecological remarks">
<paragraph id="12338574677C83E3E912F3F9697B2D62">
<heading id="500FBED36CF169CEED3EC4B8594DC69E" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="07F552F70A203FDF98AFE10146CC287A">
Large galler in syconium of
<taxonomicName id="233B37455F60B0F3C2B9A9670FEB0A1F" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="10EFA6C0744F5539D446AAAB7BA1B0EA" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
. In
<collectingCountry id="D598317AF24267F4D7A9D67460DCECDE" name="Greece">Greece</collectingCountry>
,
<taxonomicName id="6312B51247726748FA3D6CF7D35A3614" baseAuthorityName="Ishii" baseAuthorityYear="1934" class="Insecta" family="Agaonidae" genus="Eufroggattisca" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="okinavensis">
<emphasis id="E390C27F835670E50C02A16FD37EC2B3" italics="true">Eufroggattisca okinavensis</emphasis>
</taxonomicName>
was repeatedly recorded from the same syconia as
<taxonomicName id="0D3957F598FB7EEDA27ED9B1547FC36C" authorityName="Chen" authorityYear="1999" class="Insecta" family="Agaonidae" genus="Meselatus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B362D809F6EB6893357AFA05D6419DB4" italics="true">Meselatus bicolor</emphasis>
</taxonomicName>
, whose large galls suggest a possible host-parasitoid or inquiline relationship (
<bibRefCitation id="BA3EEE3227C330305780D45C4DE7F028" DOI="10.1016/j.biocontrol.2014.11.004" author="Wang R &amp; Aylwin R &amp; Cobb J &amp; Craine L &amp; Ghana S &amp; Reyes-Betancort J &amp; Quinnell R &amp; Compton SG" journalOrPublisher="Biological Control" pagination="21-30" refId="B96" refString="Wang R, Aylwin R, Cobb J, Craine L, Ghana S, Reyes-Betancort J, Quinnell R, Compton SG (2015 b) The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control. Biological Control 81: 2130. https://doi.org/10.1016/j.biocontrol.2014.11.004" title="The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control." volume="81">Wang et al. 2015 b</bibRefCitation>
;
<bibRefCitation id="C26F35132CBCEF092BFE347D052BCBCF" DOI="10.5852/ejt.2024.917.2397" author="Koutsoukos E &amp; Compton SG &amp; van Noort S &amp; Avtzis DN &amp; Askew RR" issue="1" journalOrPublisher="European Journal of Taxonomy" pagination="170-193" refId="B59" refString="Koutsoukos E, Compton SG, van Noort S, Avtzis DN, Askew RR (2024 b) A new species of Ormyrus Westwood (Hymenoptera, Ormyridae) developing in figs of Ficus microcarpa in Europe. European Journal of Taxonomy 917 (1): 170193. https://doi.org/10.5852/ejt.2024.917.2397" title="A new species of Ormyrus Westwood (Hymenoptera, Ormyridae) developing in figs of Ficus microcarpa in Europe." volume="917">Koutsoukos et al. 2024 b</bibRefCitation>
). This association raises the possibility that
<taxonomicName id="39CECA3437A94A7D54B8A167FE2932FF" authorityName="Chen" authorityYear="1999" class="Insecta" family="Agaonidae" genus="Meselatus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A4967D07DEF3B36481531E0902C3016E" italics="true">M. bicolor</emphasis>
</taxonomicName>
is also present in
<collectingCountry id="A47685E9E368D641BCCAE99C35C090C3" name="Spain">Spain</collectingCountry>
, although its presence remains unconfirmed.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5BAG" type="distribution">
<paragraph id="36D7E4A595D5D03E0667AB2466A79DB0">
<heading id="13A55D359B385F4FA3AD00364A818F2D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1AF91C9A765482CEAE5E999E1E90A9D0">
Originally from eastern Asia (
<collectingCountry id="04F1F1B4782CF60A01983663D4036C40" name="China">China</collectingCountry>
,
<collectingCountry id="A378A0963B5840891AD54CE9889CF43D" name="Japan">Japan</collectingCountry>
and Taiwan) and
<collectingCountry id="4313E18136C4FDBE35174BD1912E4413" name="Australia">Australia</collectingCountry>
(
<bibRefCitation id="91D32634103E49E11D4562C1A518B136" author="van Noort and Rasplus" firstAuthor="van Noort" refId="B91" refString="van Noort S, Rasplus J-Y (2024) Figweb: figs and fig wasps of the world. www. figweb. org [Accessed on 1 May 2024]" url="www. figweb. org" year="2024">van Noort and Rasplus 2024</bibRefCitation>
). It was reared from figs of
<taxonomicName id="C7A732A6F6E0893271CC6F1C15048B17" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="DA120B422B924E57976B106FAF8C14EA" italics="true">F. microcarpa</emphasis>
</taxonomicName>
collected in Málaga (
<collectingCountry id="FB9824A692811419E0A59A8331C4B351" name="Spain">Spain</collectingCountry>
) in 2018 and cited from
<collectingCountry id="2746FF1504BA75D43F3231F622865F56" name="Spain">Spain</collectingCountry>
as
<taxonomicName id="97180BFCE31219A91F53B7F792B724F2" class="Insecta" family="Agaonidae" genus="Eufroggattisca" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8EEDED2C796B1635F5DD25A61F6C4050" italics="true">Eufroggattisca</emphasis>
sp.
</taxonomicName>
(
<bibRefCitation id="C6E44592B8253A69A26A77ABB7F2BC77" DOI="10.3391/bir.2023.12.2.20" author="Demetriou" etAl="et al." firstAuthor="Demetriou" issue="2" journalOrPublisher="BioInvasions Records" pagination="573-580" refId="B30" refString="Demetriou J, Koutsoukos E, Mavrovounioti N, Radea K, Arianoutsou M, Roy EH, Compton SG, Martinou A (2023) A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus. BioInvasions Records 12 (2): 573580. https://doi.org/10.3391/bir.2023.12.2.20" title="A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus." volume="12" year="2023">Demetriou et al. 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5CAG" type="description">
<paragraph id="93265B6BB4ED4631D63F9A0B9CC9DE25">
<heading id="FB67D1B3F3961D75B82A974C3FD10CB6" reason="title">Short description.</heading>
</paragraph>
<paragraph id="3EDBD93AF6496A25CA5DF3ACA287EB06">
The species exhibits the following combination of characters: Head with occipital carina. In male petiole distinct, only slightly transverse, and pronotum long, much narrower than mesoscutum. Scutellum with two pairs of bristles (
<bibRefCitation id="5D3483C7F23CC4C171A08B2B259D8202" author="Boucek" firstAuthor="Boucek" journalOrPublisher="International, Wallingford, Oxon" refId="B5" refString="Bouček Z (1988) Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species. C. A. B. International, Wallingford, Oxon, 832 pp." title="Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species. C. A. B." year="1988">Bouček 1988</bibRefCitation>
). Female with gaster petiolated, either broadly convex or even slightly compressed from side-to-side. Body yellowish or brownish. This species present head and thorax mainly smooth, with sparse long setae, only rarely with some punctures sublaterally on thoracic dorsum. Notauli, usually formed by a line of punctures (
<bibRefCitation id="2F834956FC5F7D177E3516435DFB41B3" DOI="10.11646/zootaxa.2335.1.4" author="Feng and Huang" firstAuthor="Feng" journalOrPublisher="Zootaxa" pagination="40-48" refId="B33" refString="Feng G, Huang DW (2010) Description of a new species of Odontofroggatia (Chalcidoidea, Epichrysomallinae) associated with Ficus microcarpa (Moraceae) with a key to species of the genus. Zootaxa 2335: 4048. https://doi.org/10.11646/zootaxa.2335.1.4" title="Description of a new species of Odontofroggatia (Chalcidoidea, Epichrysomallinae) associated with Ficus microcarpa (Moraceae) with a key to species of the genus." volume="2335" year="2010">Feng and Huang 2010</bibRefCitation>
). Both sexes winged, this is true for this species and also for many species in
<taxonomicName id="2C12A0220EDC2CC63AED76BC427E00A6" authorityName="Hill &amp; Riek" authorityYear="1967" class="Insecta" family="Epichrysomallidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">Epichrysomallidae</taxonomicName>
, but not in general
<taxonomicName id="E23F4968937090D6AAF2D5B022415CA6" authorityName="Hill &amp; Riek" authorityYear="1967" class="Insecta" family="Epichrysomallidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">Epichrysomallidae</taxonomicName>
because of the fact that several species have apterous males.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,326 @@
<document id="100998C9F8DA5EC2F677B148D9578749" ID-DOI="10.3897/zookeys.1244.148112" ID-ZooBank="CCCA8237-6587-44BD-A25E-41BD00928A11" ID-publisher-id="148112" URI-arpha="9A0DFA26-0AED-523E-937A-37D6A902E41E" URI-zoobank="http://zoobank.org/CCCA8237-6587-44BD-A25E-41BD00928A11" article-type="research-article" checkinTime="1751465287194" checkinUser="pensoft" docAuthor="Huang, Chao, Shih, Hsi-Te &amp; Ahyong, Shane T." docDate="2025" docId="7FB5A7F23BF559619415E29FCBFBB8C3" docLanguage="en" docName="ZooKeys 1244: 1-27" docOrigin="ZooKeys 1244" docSource="https://zookeys.pensoft.net/article/148112/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megapleonum ferrumequinum Huang, Shih &amp; Ahyong, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="27" masterDocId="9A0DFA260AED523E937A37D6A902E41E" masterDocTitle="Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="1" updateTime="1751465704327" updateUser="ExternalLinkService">
<mods:mods id="F2506680DB4F71E057489F29FD62F3F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="580845E0B8F0AF063F9CFD45A45CF6BC">
<mods:title id="B1CFE46ACB3299232E88E41E65398E4C">Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="746E2D036EAFD98B9165238CBDE1CFA0" type="personal">
<mods:role id="CB678FD8DBDA28CD01EAC2E67C1A2A87">
<mods:roleTerm id="2D745A4EF1CE656CD2D68C9D344D1073">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97523414B0FFB57945EAB8EA5A666433">Huang, Chao</mods:namePart>
<mods:nameIdentifier id="096F3096AB38E2F00DD382990533B720" type="ORCID">0000-0002-3782-9096</mods:nameIdentifier>
<mods:affiliation id="3332B5814672760FB0F3161D41019F93">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="0CE3302E104F8BFF5C16586EFA616395" type="personal">
<mods:role id="7F6E350288540B89B68B7869F646CA2A">
<mods:roleTerm id="1F952D27B72CA4BB53321AAD00F524BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7035283EA18EFBA26C6A352B7975896E">Shih, Hsi-Te</mods:namePart>
<mods:nameIdentifier id="15C2C0D225937990B2FB5CEFA36DC645" type="ORCID">0000-0003-1317-8783</mods:nameIdentifier>
<mods:affiliation id="E32243B7198C83861A6B5DC6B5AACCB9">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="AE7573A693B43FBBB74A70D46FBC33EA" type="personal">
<mods:role id="6A8A53B01A3FA768F66D4C451D57F76C">
<mods:roleTerm id="18025C6C1BAD5D3AAE2E9392526EC7AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF6560E97761A41ACD11E51C573F1844">Ahyong, Shane T.</mods:namePart>
<mods:nameIdentifier id="CBA731CD0ED3DD3DB37BDF152FBC979A" type="ORCID">0000-0002-2820-4158</mods:nameIdentifier>
<mods:affiliation id="CAE4BE344FFD27909857FAEE65FDE2E5">Australian Museum, 1 William St, Sydney NSW 2010, Australia &amp; School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D1500B2769B91605B2C20BBF43BB184B">text</mods:typeOfResource>
<mods:relatedItem id="78A5DBD3633EF55645308175F301609B" type="host">
<mods:titleInfo id="BC814DBF9FFEC53410A54FAA127CA920">
<mods:title id="C28FFDBA717CDC0328DE7C73A6248D46">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F35F834238E627979C98E890F139DF31">
<mods:date id="A74E319242932095E74BD7C1B2AC5585">2025</mods:date>
<mods:detail id="10E2169DF5A8D162AC021D397186A3E9" type="pubDate">
<mods:number id="FC7621A12A7CC4CCCB37956702E4AB11">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="732634277A904BDB7FC1C38837877C5B" type="volume">
<mods:number id="4D48A17E25274D6CD152FED1AB2EA446">1244</mods:number>
</mods:detail>
<mods:extent id="46EA4A21B3D8BD8455109811041F8E4D" unit="page">
<mods:start id="37201193E29EED83E737C6178D9EDBED">1</mods:start>
<mods:end id="A123E6896F4D21ABC2DCF44FFC3637F1">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F47CF6855C673B921F4844525718862C">journal article</mods:classification>
<mods:identifier id="77C4EF3CE3B7D47B446DB06275F14F57" type="DOI">10.3897/zookeys.1244.148112</mods:identifier>
<mods:identifier id="07B5615ACD31D5B7DA76420C3EF2EFB2" type="ZooBank">CCCA8237-6587-44BD-A25E-41BD00928A11</mods:identifier>
</mods:mods>
<treatment id="7FB5A7F23BF559619415E29FCBFBB8C3" ID-DOI="http://doi.org/10.5281/zenodo.15791870" ID-Zenodo-Dep="15791870" ID-arpha="7FB5A7F2-3BF5-5961-9415-E29FCBFBB8C3" ID-zoobank="https://zoobank.org/3E312295-C1E5-47FD-B90E-086F02A72785" LSID="urn:lsid:plazi:treatment:7FB5A7F23BF559619415E29FCBFBB8C3" LSID-ZBK="urn:lsid:zoobank.org:act:3E312295-C1E5-47FD-B90E-086F02A72785" httpUri="http://treatment.plazi.org/id/7FB5A7F23BF559619415E29FCBFBB8C3">
<subSubSection id="8810544635DFBB8EBDC1B1E287B0F064" type="nomenclature">
<paragraph id="37A8D99252A807B2D94C57CA3927731F">
<taxonomicName id="D21CFB74667E16864B46B86ACCF31049" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="EB5126FACB434F90651DFA0269B12A87" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39B95C50084AA6A162E3F09CD56A27C4" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F64874B073851F6AB281CBD2131EA34D" type="description">
<paragraph id="17B57867431BC922754CDEF7CEA25FB9">
<figureCitation id="FFA985FA1950C6305C950CEFCC2DB4F3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">Figs 3</figureCitation>
,
<figureCitation id="B541CD0D5EAECCB037F3228CC4355095" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4</figureCitation>
,
<figureCitation id="3F5E3EFF32F8D61CBBFB71790C27DBA9" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELKAG" type="type material">
<paragraph id="E7733A4D3046691E56CC570D82F2FB86">
<heading id="12ABB677F533386C378EF7AE74CA63E1" reason="title">Type material.</heading>
</paragraph>
<paragraph id="A7D3848F30469FB070E1BD20CEF99715">
<materialsCitation id="473D2F93C498E6A62B6EFBDF301E54AA" collectingDate="2018-06" collectionCode="SYSBM" collectorName="Jian Wang" country="China" county="Maoming City" latitude="22.29" location="Datianding" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" specimenCode="SYSBM 002143" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="93E9372EB1B673A6B5ABC0B125C47DE2" bold="true" italics="true">
<typeStatus id="6B041343F75C392FE2397711C83536D3" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCode id="E7B72D3D7A42D67AC262DF5E04ACDB6F" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EWKAG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002143
</specimenCode>
, male (16.6 ×
<quantity id="70D88C76540877943F410A4503CA6D06" metricMagnitude="-2" metricUnit="m" metricValue="1.31" unit="mm" value="13.1">13.1 mm</quantity>
),
<location id="B810B1EDE613D0FADE9A7328BEBE6361" LSID="urn:lsid:plazi:treatment:7FB5A7F23BF559619415E29FCBFBB8C3:B810B1EDE613D0FADE9A7328BEBE6361" country="China" county="Maoming City" latitude="22.29" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" name="Datianding" stateProvince="Guangdong">Datianding</location>
,
<collectingMunicipality id="447B3A3E253AEF70B8D45B1B367F04F4">Dawuling Nature Reserve</collectingMunicipality>
,
<collectingCounty id="B9ACA5B72D434B1293F2F72E2B4D8951">Maoming City</collectingCounty>
,
<collectingRegion id="3C2FB037B4E3449CB2FB96FA0A3A2184" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="79365AA89DE48FF79F72350C22BC9D47" name="China">China</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5KAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.220000,22.290000]}">
<geoCoordinate id="2EC0BCEA79A60852BC2293234D962B5D" degrees="22.29" direction="north" orientation="latitude" precision="555" value="22.29">22.29 ° N</geoCoordinate>
,
<geoCoordinate id="347207F46365C3925682431A69967755" degrees="111.22" direction="east" orientation="longitude" precision="555" value="111.22">111.22 ° E</geoCoordinate>
</named-content>
, dirt road near the summit at night, coll.
<collectorName id="EEAEE5849D1FBDF2DF3768AC212504D8">Jian Wang</collectorName>
,
<date id="E4E3F0C39CD0B116C29FD5D5C8D684BE" value="2018-06">
<collectingDate id="E3D230CCEEF623791F5FF5EC9940CF67" value="2018-06">June 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDLAG" type="diagnosis">
<paragraph id="E5F325C0A5663A9193673C43296FD393">
<heading id="E17173D8A4798BE69899B1FEF6A732FC" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="48D6D106F32258EA0857EAB02FBAAC55">
Carapace broader than long, dorsal surface convex, lateral margins covered in dense setae; postorbital, epigastric cristae weak, almost confluent (Fig.
<figureCitation id="5648647E25E842FA0B108DE576C30C6B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Maxilliped 3 merus width ~ 1.3 × length; ischium width ~ 0.7 × length; exopod reaching slightly beyond anterior edge of ischium, without flagellum (Fig.
<figureCitation id="BA7DE47F45399470946004C5B84B46A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 A</figureCitation>
). Ambulatory legs densely setose; pereiopod dactylus shorter than propodus (Fig.
<figureCitation id="9E100258389EFBD53A0BA064AC5E240B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Male anterior thoracic sternum very broad, width ~ 1.9 × length (Fig.
<figureCitation id="40F5EF1CC1DD2ED3F61AB0D596477856" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 C</figureCitation>
). Male pleon large, sublinguiform, pleonite 6 width ~ 2.5 × length; telson width ~ 1.7 × length (Fig.
<figureCitation id="15F26089A250637E7E7CD68C2425CB5C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 D</figureCitation>
). G 1 large, slightly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="E0DD90AE42840D889E0AA1DF0BF24E08" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="A093C75CC40AAB0074F4012AA7E9A2EF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 E</figureCitation>
); subterminal segment length ~ 2.5 × length of terminal segment (Fig.
<figureCitation id="99C5BFB160529894DC0FA22847B9D8B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 C</figureCitation>
). Subterminal segment outer margin slightly concave, outer distal margin bulging; terminal segment curved inwards, tip pointed upwards in dissected view, connected to a large mesoanterior-facing horseshoe shaped structure, large thick proximal pad on dorsal side (Fig.
<figureCitation id="CBAE3BA31F97DF7DF80FA067A197006F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 C, F, G</figureCitation>
). G 2 subterminal segment tapering, slightly bent outwards distally, flagelliform terminal segment thick, ~ 1.6 × length of subterminal segment, apex blunt (Fig.
<figureCitation id="B785FE051E74177C4E2CDD2B74E4ABC3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENMAG" type="description">
<paragraph id="CF48ABA3BCE1E2A9DF1EE2F24F1F4753">
<heading id="AB08A4D91A9149AD3B1E4D39183D7339" reason="title">
Description of male
<typeStatus id="9ABA0E33644B83825D60E132E2764CC4" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="73276AB2737C08DB0E3B023BF7519DB0">
Carapace broader than long, ~ 1.3 × as wide as long; regions not pronounced, dorsal surface convex; surface finely pitted, dense setae at lateral margins (Fig.
<figureCitation id="4F4EED11D06A52F706D33804D4BBF46E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Frontal margin almost straight, deflexed (Fig.
<figureCitation id="567A1C6CC5E72EB3B767112FFFA3A9D9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Epigastric cristae and postorbital cristae rugose, low, almost confluent; bifurcated shallow groove between epigastric cristae (Fig.
<figureCitation id="FE86DB0FDD59CE496F9FC0791CA49F10" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Branchial regions not swollen (Fig.
<figureCitation id="C2CB2FB12C84EE6EE42CF5F1B6DA4503" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Cervical groove shallow, barely visible (Fig.
<figureCitation id="781EDC2FBD0B7B827017466A9118B9DF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Mesogastric region flat (Fig.
<figureCitation id="D6DF1E0E3FBEC4AB9E3B273C50752D56" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). External orbital angle broadly triangular, outer margin slightly convex, confluent with anterolateral margin (Fig.
<figureCitation id="395A327BB76C599A9D06196D14D00327" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Epibranchial tooth granular, indistinct (Fig.
<figureCitation id="B060ACE09FE2579EC1464F6A644CB83E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Anterolateral margin lined with indistinct single or partially fused granules; posterolateral margin posteriorly convergent (Fig.
<figureCitation id="17577893D8C915E6985D7A3EF615A1B7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
); posterolateral surface smooth (Fig.
<figureCitation id="BF1E90A7B7526B922C752F83BE0CAD96" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Orbits regular; supraorbital margins weakly cristate, infraorbital margins lined with fused granules (Fig.
<figureCitation id="E059AEE27D30FC5B397E34912AFC9D55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 B</figureCitation>
). Eyes normal (Fig.
<figureCitation id="91FD199E02AAD93DB1173722B5D12623" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Sub-orbital, pterygostomial and sub-hepatic regions generally smooth, pitted (Fig.
<figureCitation id="948F26DC23406A6954CCD4327C36C89E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
). Antennules large, folded within broad fossae; antennae very short (Fig.
<figureCitation id="17FF81DB809FE506C6A2022CC73BA90C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 B</figureCitation>
). Median lobe of epistome buccal margin broadly triangular, lateral margins straight (Fig.
<figureCitation id="77F6DBD467E29F595D51049096587BB5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 B</figureCitation>
).
</paragraph>
<paragraph id="83B8DAAD1C2CFDF83026E1B7AB2BAEC7">
Maxilliped 3 merus subtrapezoidal, with slight median depression, width ~ 1.4 × length; ischium subtrapezoidal with shallow median sulcus, distomesial margin rounded, width ~ 0.7 × length. Exopod reaching proximal one-third of merus; flagellum absent (Fig.
<figureCitation id="B1C1CC25C3F446E66B83F772B48480B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 A</figureCitation>
).
</paragraph>
<paragraph id="7B790542BD079510050E49F1CA698DDA">
Chelipeds (pereiopod 1) subequal (Fig.
<figureCitation id="7DA4FB6DBAA5EE96317F6656CB5D0203" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 D, E</figureCitation>
). Merus trigonal in cross section, surfaces generally smooth, margins lined with long setae; outer dorsal margin slightly crenulated margins slightly crenulated, inner and ventral margin lined with large granules (Fig.
<figureCitation id="FDF4BE53E2E13E1FE73262A555C583C8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A, B</figureCitation>
). Carpus dorsal surface slightly rugose, with large spike at inner-distal angle, spinule at base (Fig.
<figureCitation id="8E1D7EA0307DFD8734A702A52571AE77" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Major cheliped palm length ~ 1.5 × height; dactylus 0.7 × palm length (Fig.
<figureCitation id="591E7B341F3B41A47E7957FC972ACB07" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 D, E</figureCitation>
). Palm surface pitted, occlusal margin of fingers with 79 irregular blunt teeth, with small gape when closed (Fig.
<figureCitation id="194E51DF57EE37E651DDF462D2D95493" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 D, E</figureCitation>
).
</paragraph>
<paragraph id="906ED1ABC52B65894BAC0CBC17DE5404">
Ambulatory legs (pereiopods 25) covered with setae, especially dense on pereiopods 23 (Fig.
<figureCitation id="A10D7928934BA6C7435B92F984876950" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Pereiopod 3 merus 0.6 ×
<abbrev id="ABBRID0ERPAG" xlink_title="Carapace length">CL</abbrev>
(Fig.
<figureCitation id="2D60F099746EF1F6F8AB9B091179525C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). Pereiopod 5 propodus length 1.7 × height, longer than dactylus (Fig.
<figureCitation id="331DD2BB0867C1AC9B99FC4A60F811E1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
).
</paragraph>
<paragraph id="A72CE3F7FF80E91198E32F3139B67972">
Male thoracic sternum generally smooth, pitted, setae at margins and sparsely on sternites; sternites 14 width ~ 1.9 × length; sternites 1, 2 fused to form broad triangle; fused sternites 1, 2 demarcated from sternite 3 by sinuous transverse sulcus, sulcus lined with setae; sternites 3, 4 fused without obvious demarcation (Fig.
<figureCitation id="B4C7E870F9189FD6A2552196664C9417" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 C</figureCitation>
). Male sterno-pleonal cavity reaching anteriorly slightly beyond mid-length of cheliped coxa (Fig.
<figureCitation id="8507342E429E653651056D46F731ACC4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 C</figureCitation>
). Male pleonal locking tubercle positioned slightly posterior to mid-length of sternite 5 (Fig.
<figureCitation id="2DB92614060A5F090E3CF2E5D6E36FE0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 E</figureCitation>
).
</paragraph>
<paragraph id="85C37E4142956033B31350B1CCC9818E">
Male pleon large, sublinguiform; somites 36 progressively narrower; somite 6 width approximately 2.5 × length; telson width 1.7 × length; lateral margins almost straight, apex rounded (Fig.
<figureCitation id="E5D12D77E30C9D62232964C64D506675" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 D</figureCitation>
).
</paragraph>
<paragraph id="90FE445E07B57D2D47C93A59D7F363D9">
G 1 large, slightly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="2A1E69D96384F52420343FB2D962EEFB" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="BF29A11641693469BEF8231E21214A08" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 E</figureCitation>
); subterminal segment length ~ 2.5 × length of terminal segment (Fig.
<figureCitation id="76D01CEFAB863428F5E1CD88E8BE4A88" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 C</figureCitation>
). Subterminal segment outer margin slightly concave, outer distal margin bulging; terminal segment curved inwards, tip pointed upwards in dissected view, connected to a large meso-anterior-facing horseshoe shaped structure, large thick proximal pad on dorsal side (Fig.
<figureCitation id="725FE8D2FE16A1D86CABA842EDAFEED4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 C, F, G</figureCitation>
). G 2 subterminal segment tapering, slightly bent outwards distally, flagelliform terminal segment thick, ~ 1.6 × length of subterminal segment, apex blunt (Fig.
<figureCitation id="EA06795011BBA94B90C8549A188AE818" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDRAG" type="Colour in life">
<paragraph id="A91D0C29C62FBD8E2A9FC84ECDCF5FE4">
<heading id="5BDDD84EF12F6D22E756BACC5C10FA97" reason="title">Colour in life.</heading>
</paragraph>
<paragraph id="64D53D9462FA2033A29CF9D288BD7C01">
Generally drab camouflage-brown all over (Fig.
<figureCitation id="EF6EB61C9EAE21ED8F5BDDF4DB5CF888" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENRAG" type="habitat">
<paragraph id="C758535DF71F3A9A48D866E1D0776412">
<heading id="643D87CB983B2C5039A0242EAE7A205B" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="664295A3561EAF049F0E5105C2B1A793">
Typical habitat unknown. The only specimen collected was found at night on the side of a dirt road near the summit at approximately
<quantity id="9BFB2208C05BF670A6CABF04EE414277" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</quantity>
above sea level. Multiple subsequent attempts to find this species in the same area failed to locate any more specimens with only
<taxonomicName id="3C4BAE8FA69839EF581F0D00801BC2C1" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="EFC4CBEA3A9534233BEDB3E35A6ACCF8" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DA3A0E9F746FC3D6CBB2E43E97A9530" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="898ED8A1F1310CA5265D4D814E29E63C" baseAuthorityName="Dai &amp; Turkay" baseAuthorityYear="1997" class="Malacostraca" family="Potamidae" genus="Eurusamon" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="51B18A4E468FB018A3EEA0FF00F7F948" italics="true">Eurusamon guangdongense</emphasis>
</taxonomicName>
being found. The true habitat of this species remains elusive.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJSAG" type="distribution">
<paragraph id="201DBD3708F8F745D403DD2BB7C9D157">
<heading id="0E1E0613345B2CE67B9279C23FC531E4" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0DA64B912661501BDACF627635A7DA4E">
Dawuling Nature Reserve, Maoming City,
<collectingRegion id="854C62F90E6C5F042E267AD9830182BB" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="3D73C2B3FE1024DA790A328C432A38B5" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOSAG" type="etymology">
<paragraph id="764DCC3D158ADD71014A8B3380098F07">
<heading id="4C577FDBEB16A6268B4552AC442BC759" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="59F422789AE2DCDC1B6CFF25C7579482">
The species name is the Latin word
<emphasis id="962540D30FB1A052D88EEBC82BBB6FF8" italics="true">ferrumequinum</emphasis>
, meaning horseshoe. It alludes to the horseshoe-shaped structure on the G 1 terminal segment of this species. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5SAG" type="remarks">
<paragraph id="FC2DD1AC2ADEAB53DF3A9E1904DFF389">
<heading id="F621030104E61D858E7A43B5A1FBC0D1" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="AAE684299076413A53E6A3C4F6D5AB7B">
<taxonomicName id="AD2C4A657E979332338E7E3E2E2ECA83" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="056DBFD590C2F194EED6EB5805D67698" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="082E437B02D29A8659E83E088BB87DAD" rank="species">sp. nov.</taxonomicNameLabel>
is unique in its genus by its sublinguiform male abdomen (Fig.
<figureCitation id="43162EB8970227542B4AE6E836F28C54" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 D</figureCitation>
, vs broadly triangular in all other congeners) and its peculiar G 1, which is only slightly sinuous (Fig.
<figureCitation id="0EA2B5FAC93B9A260B313D5A803268C5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 C</figureCitation>
, vs more strongly sinuous in all other congeners) with a horseshoe-shaped structure in the terminal segment (Fig.
<figureCitation id="85C1707F01A6982315D8F76D90F01527" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 F H</figureCitation>
, vs absent in all other congeners). In possessing dense setation on the carapace margins and ambulatory legs,
<taxonomicName id="1D5747B09E44826F00847287A0C9EA15" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="7D9452506F9790E5463AB82E9A56D13B" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="400BCF8C6DFDBABF3D354C0CEE2F623E" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="B1C14C0BB3151CBF91152DE55230ABF6" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="36A0EDED244A0CF8A34A5A635276DD19" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31A76E46B0EB942B2E18A3F22844FE9F" rank="species">sp. nov.</taxonomicNameLabel>
(see remarks for
<taxonomicName id="D552B69E7CDE2F14FB134F137CCF70B9" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="3A6A27588E48F91457458A689A9E71B9" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40412C664D184A6DAF72F1E034133E66" rank="species">sp. nov.</taxonomicNameLabel>
). More detailed comparisons can be found in Table
<tableCitation id="4BF8CABF187D8178C464BF073CBE7DA4" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparisons between the species within the genus Megapleonum." httpUri="http://table.plazi.org/id/69D766E64FF15F0F8BB8C5782AADA37B" tableUuid="69D766E64FF15F0F8BB8C5782AADA37B">1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,580 @@
<document id="2C638CEBDE6A013AE42520DA0256674A" ID-DOI="10.3897/zookeys.1244.148112" ID-ZooBank="CCCA8237-6587-44BD-A25E-41BD00928A11" ID-publisher-id="148112" URI-arpha="9A0DFA26-0AED-523E-937A-37D6A902E41E" URI-zoobank="http://zoobank.org/CCCA8237-6587-44BD-A25E-41BD00928A11" article-type="research-article" checkinTime="1751465287194" checkinUser="pensoft" docAuthor="Huang, Chao, Shih, Hsi-Te &amp; Ahyong, Shane T." docDate="2025" docId="9E1C850AB36F51BD922339DC594E89F8" docLanguage="en" docName="ZooKeys 1244: 1-27" docOrigin="ZooKeys 1244" docSource="https://zookeys.pensoft.net/article/148112/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megapleonum yangdongense Huang, Shih &amp; Ahyong, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="27" masterDocId="9A0DFA260AED523E937A37D6A902E41E" masterDocTitle="Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="1" updateTime="1751465704327" updateUser="ExternalLinkService">
<mods:mods id="9C7128B2FBDB6941C0E00670C0ECDCEE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F96C0E7BED9FBA58AB38DE2FB64B4D3E">
<mods:title id="86580ACBC40E88D3324FFBD98268FD97">Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="1814B21B408CA28C9302974F6963737B" type="personal">
<mods:role id="B9B73CDF0314D9645594FD41ED9BDF81">
<mods:roleTerm id="4A96AE85C85189A48B1F3988949BD7CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0AB6CA11AFDF7156E9AD0D587586DFC9">Huang, Chao</mods:namePart>
<mods:nameIdentifier id="20AE5030228B2914400C50D594056A4E" type="ORCID">0000-0002-3782-9096</mods:nameIdentifier>
<mods:affiliation id="C239E3CE589671E8FCE094ACB6DCDA58">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="CEFABB9FB05B9726CE6FF5C65DA485F8" type="personal">
<mods:role id="5B35F81EFDE0567EB3209A38B547E634">
<mods:roleTerm id="DE7D0EF3BC2F1B932CD13BF7EF50EDE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78A03DC59175DFDBAB507B082C25F8FA">Shih, Hsi-Te</mods:namePart>
<mods:nameIdentifier id="05465FE1E755AA9DACB66E1E3CA2E456" type="ORCID">0000-0003-1317-8783</mods:nameIdentifier>
<mods:affiliation id="8512D34470E8804C846F0DEEDBDAA693">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="B93F52F058498B93341BA4A5A5D6303D" type="personal">
<mods:role id="ACFECD9B75BE6DCC43AA6048FC997215">
<mods:roleTerm id="4E19491362CCEFCA105483E540C626C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2497D3F573C8BF169C4B78E43FB664E7">Ahyong, Shane T.</mods:namePart>
<mods:nameIdentifier id="AF666091A2C17F1383542EE67CD40D50" type="ORCID">0000-0002-2820-4158</mods:nameIdentifier>
<mods:affiliation id="EA5425133EC038B15A95CDED087FE105">Australian Museum, 1 William St, Sydney NSW 2010, Australia &amp; School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="07CAC647C2CB1E976C55BBC4EAE86943">text</mods:typeOfResource>
<mods:relatedItem id="771B239C9025AEED72795AF2C590E388" type="host">
<mods:titleInfo id="E65C143767AD7733ACE7B69632671285">
<mods:title id="0D10C636666521D2733A840293B32C5E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="B325BA73F97500422B5B7FF87C371DAE">
<mods:date id="96068876B641F80860ED32D79345002C">2025</mods:date>
<mods:detail id="83C4AF54436093DC93A2941FDCBEF2E9" type="pubDate">
<mods:number id="4295B9AA877FFDB134E4DB1999177978">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="A134E40B2391D188377A078885EE2691" type="volume">
<mods:number id="5AD4466CD5D40C3185F452B423D89DEF">1244</mods:number>
</mods:detail>
<mods:extent id="4FBB3A83C37CD468AF18C635CA4DE879" unit="page">
<mods:start id="F1B75A13CF1D7FF719A7AB42BF04ACF8">1</mods:start>
<mods:end id="A9F8F99A122001CE1B1D62813A061EF0">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6C1274EF47D8E12B5F4E79873C565C61">journal article</mods:classification>
<mods:identifier id="A01BFCA87904F5502903808C62FD8272" type="DOI">10.3897/zookeys.1244.148112</mods:identifier>
<mods:identifier id="0C3FD4657EDC645D15FA38066CBEF859" type="ZooBank">CCCA8237-6587-44BD-A25E-41BD00928A11</mods:identifier>
</mods:mods>
<treatment id="9E1C850AB36F51BD922339DC594E89F8" ID-DOI="http://doi.org/10.5281/zenodo.15791874" ID-Zenodo-Dep="15791874" ID-arpha="9E1C850A-B36F-51BD-9223-39DC594E89F8" ID-zoobank="https://zoobank.org/63BEBAEA-6C90-4FCE-809D-7E4C7FBD62FF" LSID="urn:lsid:plazi:treatment:9E1C850AB36F51BD922339DC594E89F8" LSID-ZBK="urn:lsid:zoobank.org:act:63BEBAEA-6C90-4FCE-809D-7E4C7FBD62FF" httpUri="http://treatment.plazi.org/id/9E1C850AB36F51BD922339DC594E89F8">
<subSubSection id="8E8A62135D8D485DBF7A5E4D9C592D34" type="nomenclature">
<paragraph id="771061FCD4FDD80FFFFC071F65B4C9A0">
<taxonomicName id="D76177E36A77F4E56C4BB55719F4ED38" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="0FE2DC7795CD794FB280401FE74EF349" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9ED87EC00DD61BD47FE51AA2F43E23B9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D2659B0130367CF1E661436BD7DD8309" type="description">
<paragraph id="23C640A8465743122A1AF7D692FA755B">
<figureCitation id="5FD34250B198F633482A6EF6DABA54C5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">Figs 8</figureCitation>
,
<figureCitation id="FDF266460E1FF47A15A0F327F8C85281" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9</figureCitation>
,
<figureCitation id="8EDE5904BF3079F2CF1AF61F2E27BB46" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10</figureCitation>
,
<figureCitation id="DEFAA742E5446C3358B2C0D5FD4CA0E1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 C, D</figureCitation>
,
<figureCitation id="F0BFB5C6F354A614090598386C99FD7D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWMBG" type="type material">
<paragraph id="10BDFD8BA4AC3F40B416414CF8F0D405">
<heading id="77C3A2228F69A96D11F307339D8A0943" reason="title">Type material.</heading>
</paragraph>
<paragraph id="BFD18CC0C5A7F96716A8C8C257C0294B">
<materialsCitation id="BF1AC2053315862617F76AC75854E74D" collectingDate="2019-06-01" collectionCode="SYSBM" collectorName="Chao Huang &amp; Shih &amp; Bernhard Bein" country="China" county="Yangdong County" latitude="21.97" location="Gaozhai" longLatPrecision="757" longitude="112.07" municipality="Yangjiang City" specimenCode="SYSBM 002152" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="7F6289421EC10A90F16482599016C240" bold="true" italics="true">
<typeStatus id="B912CA75CAE444B86FFB5084DFAAB80A" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCode id="A6B8C67EFF3B86388EA26D2CF58A55CE" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EBNBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002152
</specimenCode>
, male (17.8 ×
<quantity id="C1C38B40DA9716F96318E5527B7293E5" metricMagnitude="-2" metricUnit="m" metricValue="1.45" unit="mm" value="14.5">14.5 mm</quantity>
),
<location id="4F8FCB62D001B69490300493B8E6D577" LSID="urn:lsid:plazi:treatment:9E1C850AB36F51BD922339DC594E89F8:4F8FCB62D001B69490300493B8E6D577" country="China" county="Yangdong County" latitude="21.97" longLatPrecision="757" longitude="112.07" municipality="Yangjiang City" name="Gaozhai" stateProvince="Guangdong">Gaozhai</location>
,
<collectingCounty id="A93F85EE0E60BDC13AD9F940F02D629A">Yangdong County</collectingCounty>
,
<collectingMunicipality id="3C63BB2695A3D70DAD7F10F61F7FA61F">Yangjiang City</collectingMunicipality>
,
<collectingRegion id="6B6829B42733193713E96935DB1561D6" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="F05DCE4730ADACA293D47313D0FDD512" name="China">China</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJNBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[112.070000,21.970000]}">
<geoCoordinate id="8EA9EF1F3C30FC33C3B2CF6A2DBBE049" degrees="21.97" direction="north" orientation="latitude" precision="555" value="21.97">21.97 ° N</geoCoordinate>
;
<geoCoordinate id="47AFCD1E3827653BE32F8201E3F423A4" degrees="112.07" direction="east" orientation="longitude" precision="555" value="112.07">112.07 ° E</geoCoordinate>
</named-content>
, under rocks in hillstream, coll.
<collectorName id="916D3199EB0B59DD5BE5AE1463F222EA">Chao Huang</collectorName>
, Hsi-Te
<collectorName id="82F69810A0E56351F413B804AE161B4B">Shih</collectorName>
and
<collectorName id="7884A68ECDBBE355F0B76688345993ED">Bernhard Bein</collectorName>
,
<date id="FB4CD233AFADEA3AB277829F7B64D5EC" value="2019-06-01">
<collectingDate id="CC3709849CBFB6CAD51B8ADB58BF1BC6" value="2019-06-01">1 June 2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="58A970E5CB1203AC251F98CD07766C09" collectingDate="2019-06-01" collectionCode="IHB" collectorName="Chao Huang &amp; Shih &amp; Bernhard Bein" country="China" county="Yangdong County" latitude="21.97" location="Gaozhai" longLatPrecision="757" longitude="112.07" municipality="Yangjiang City" specimenCode="SYSBM 002153, SYSBM 002154-002155" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="1A0FE6CF67F786C461C403C9618B6505" bold="true" italics="true">
<typeStatus id="AA730953B1A320F0B49B714320ACED19" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCode id="60C94EEAC78D6FD623211A27BFFB3050" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0ERNBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002153
</specimenCode>
, male (17.3 ×
<quantity id="973C2C42632BF437190B497762143466" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" unit="mm" value="13.7">13.7 mm</quantity>
), same data as holotype.
<specimenCode id="FA6875459E79EEBA9804557003F99DD9" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EWNBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002154002155
</specimenCode>
,
<specimenCount id="259DB5834CC3F1D3C03FC9BB770718BF" count="2" type="female">2 females</specimenCount>
(16.0 ×
<quantity id="39EE12A94C6AB293D715EBD0F31D78B1" metricMagnitude="-2" metricUnit="m" metricValue="1.24" unit="mm" value="12.4">12.4 mm</quantity>
, 15.3 × 12.0 mm), same data as
<typeStatus id="D8FB022B186DDC61EADDAEC9F06175E1" type="holotype">holotype</typeStatus>
.
<named-content id="039F35F2839C3867F7AECE687C1F8A0A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-hydrobiology-chinese-academy-sciences-0" xlink_title="Museum of Aquatic Organism, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, China">
<collectionCode id="1B9EDF4BA84FE05FE42048916A540106" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" type="">IHB</collectionCode>
</named-content>
,
<specimenCount id="45DEA91494E72C37EC848545791C0AD4" count="2" type="male">2 males</specimenCount>
(16.0 ×
<quantity id="888C19DFD746C0AEAAAB39D93ECEAA5F" metricMagnitude="-2" metricUnit="m" metricValue="1.27" unit="mm" value="12.7">12.7 mm</quantity>
, 15.4 × 12.0 mm), same data as
<typeStatus id="C15DA05BB3F9D7CF224DC4CBFA41CD39" type="holotype">holotype</typeStatus>
.
<named-content id="DC145668B4841FEE943AE35308F66864" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-chung-hsing-university" xlink_title="Zoological Collections of the Department of Life Science, National Chung Hsing University, Taichung, Taiwan">NCHUZOOL</named-content>
15306,
<specimenCount id="2199DE6BEFA3129E92CB0E182A4084A6" count="2" type="male">2 males</specimenCount>
(16.0 ×
<quantity id="652A6AE4BAAF1A371FD635642EB9EAB5" metricMagnitude="-2" metricUnit="m" metricValue="1.31" unit="mm" value="13.1">13.1 mm</quantity>
, 14.9 ×
<quantity id="4461F93A5031936D3C794CECB4B49EBB" metricMagnitude="-2" metricUnit="m" metricValue="1.18" unit="mm" value="11.8">11.8 mm</quantity>
),
<specimenCount id="4619172BCC4FC21F7F7AF5ABF68A8763" count="1" type="female">1 female</specimenCount>
(16.0 ×
<quantity id="954D9A6821D39E551F274658C16EAEA8" metricMagnitude="-2" metricUnit="m" metricValue="1.26" unit="mm" value="12.6">12.6 mm</quantity>
),
<named-content id="3BFA64BA4C0F131601F4233C4180F6B6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-chung-hsing-university" xlink_title="Zoological Collections of the Department of Life Science, National Chung Hsing University, Taichung, Taiwan">NCHUZOOL</named-content>
15307,
<specimenCount id="DE2DB792E9DB6D62B0990577F8BE797D" count="1" type="male">1 male</specimenCount>
(16.7 ×
<quantity id="E65EABCF76B52F6CD6F05F7B3C5B6267" metricMagnitude="-2" metricUnit="m" metricValue="1.38" unit="mm" value="13.8">13.8 mm</quantity>
), same data as
<typeStatus id="9DFDE20F4E8623B9FD1F041B9F2ED6FA" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKOBG" type="diagnosis">
<paragraph id="8FC409A0264EBE98B0EC4058319FC948">
<heading id="1F585A7D241999DCBE307B6B32B7E10D" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4156EE98386626935CF047B96AB664C2">
Carapace broader than long, dorsal surface slightly convex, postorbital, epigastric cristae confluent, sharp (Fig.
<figureCitation id="5B853B490C96B5F4B21F018FCA26C815" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Maxilliped 3 merus width ~ 1.3 × length; ischium width ~ 0.7 × length; exopod reaching slightly beyond anterior edge of ischium; flagellum short, as long as dactylus (Fig.
<figureCitation id="E09366A97682E6BAF1C0AD896BD45A61" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 A</figureCitation>
). Ambulatory legs without long setae, pereiopod dactylus shorter than propodus (Fig.
<figureCitation id="BC07563DE85D4DA6EC1AE46EC6726170" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Male anterior thoracic sternum very broad, width ~ 1.9 × length (Fig.
<figureCitation id="5D9948B84AEB5C63ECCEAE39EAC2E5DA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 B</figureCitation>
). Male pleon large, broadly triangular, pleonite 6 width ~ 2.4 × length, telson width ~ 1.8 × length (Fig.
<figureCitation id="372E00CEDDA98363003F1804CCCDE23A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 C</figureCitation>
). Female pleon subovate (Fig.
<figureCitation id="64E7C22297A1C00438FB98D3C8B03E91" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 E</figureCitation>
). G 1 large, sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="474BE540E03DBF2B9E1476E7DCEBA880" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="39DBDAC6CE9C1C9B974414DF82346B56" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 D</figureCitation>
); subterminal segment length ~ 2.7 × length of terminal segment (Fig.
<figureCitation id="521B7D75F3DBDD072521F1FD41020406" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 C E</figureCitation>
); subterminal segment thick, outer margin strongly concave, distal end slanted with outer-distal section highest; terminal segment short, both lateral margins slightly convex, bifurcated with one point being opening tip and other a smaller projection on outer margin, both pointing upwards (Figs
<figureCitation id="751A4CCAF4051141ADA924BB2B5B6852" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 C E</figureCitation>
,
<figureCitation id="4EC6D00421D53A765B7A423014EAFCB6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 C, D</figureCitation>
). G 2 subterminal segment thick, slightly bent outwards distally, flagelliform terminal segment thin, ~ 2.3 × length of subterminal segment (Fig.
<figureCitation id="3C56D43B0428E23CFDCA9538136C6E1A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 B</figureCitation>
).
</paragraph>
<caption id="A010981D11D658BB26217169DF0D40E4" ID-DOI="10.3897/zookeys.1244.148112.figure8" ID-arpha="1E61C6F8-24E8-5CCA-9EAE-4D696FE6D7C4" httpUri="https://binary.pensoft.net/fig/1366345" startId="F8">
<paragraph id="F6D1ED754D531A2380C2C92C3C856691">
<label id="7E940B41537A02BA7D62B4B4BD8ECD54">Figure 8.</label>
</paragraph>
<paragraph id="793D0208E5DAE7BF24B3229593EBD579">
Dorsal habitus.
<taxonomicName id="79B8E5B6C529B73CCB60B892C7A11690" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="46E647BF5FAF27A332F3279491FA87C9" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB02A8A93F1C775060C4AA14A0F0CE7" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (17.8 × 14.5 mm),
<abbrev content-type="institution" id="ABBRID0EPQBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002152 (
<emphasis id="CA670EE320D609AE12C802C6D4924962" bold="true">A</emphasis>
); female paratype (16.0 × 12.4 mm),
<abbrev content-type="institution" id="ABBRID0EWQBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002154 (
<emphasis id="2F4C49D22C9083997B4B1615CBDC9D8F" bold="true">B</emphasis>
).
</paragraph>
</caption>
<caption id="E511A504F160D7815BC35F87399AB291" ID-DOI="10.3897/zookeys.1244.148112.figure9" ID-arpha="41E4FE3F-A548-536E-9B2C-910C19F924B1" httpUri="https://binary.pensoft.net/fig/1366346" startId="F9">
<paragraph id="5523EAA88BAF07B960137FCA954F55B9">
<label id="729CD32488A481928645A2127CD5DEED">Figure 9.</label>
</paragraph>
<paragraph id="E5F5E941C14AD3CBE287176F2281B372">
<taxonomicName id="A76D3BD5FE1472DA53BFCDBFD5A803EB" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="98F8777508001DEF30BA9EE6F9492424" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE6F714FD44B6F30A71BEE8897D52AD9" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (17.8 × 14.5 mm),
<abbrev content-type="institution" id="ABBRID0EVRBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002152 (
<emphasis id="79D1A650B9BB4A57CEC7CACD290EDE2E" bold="true">A</emphasis>
<emphasis id="C2B757520DBFC217DBAB8C4A809454F4" bold="true">D</emphasis>
); female paratype (16.0 × 12.4 mm),
<abbrev content-type="institution" id="ABBRID0E5RBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002154 (
<emphasis id="F622D019216E4B7DBD3B5CDEAAB8D520" bold="true">E</emphasis>
,
<emphasis id="98BA941DC431FE90D8EE5E0B9EDBACA1" bold="true">F</emphasis>
). Cephalothorax, anterior view (
<emphasis id="639C3FFF944FD0D1B2C48A67F5D1B0C7" bold="true">A</emphasis>
); anterior thoracic sternum (
<emphasis id="03517FDE1AFC9DBC52EBE2D910FDCE7B" bold="true">B</emphasis>
); anterior thoracic sternum and pleon, ventral view (
<emphasis id="F2A809134A1BD2BF0D0BB8970CC45CFA" bold="true">C</emphasis>
); sterno-pleonal cavity with G 1 in situ, ventral view (
<emphasis id="B14E5970B3D9ABDD57A660887209225A" bold="true">D</emphasis>
); pleon, ventral view (
<emphasis id="02D8249856E2674FB420C1E183AA77EB" bold="true">E</emphasis>
); vulvae, ventral view (
<emphasis id="456055D081BA587D05A96A5F7127D7A5" bold="true">F</emphasis>
).
</paragraph>
</caption>
<caption id="38095358AB5E97E85588C62A5EEF8BBF" ID-DOI="10.3897/zookeys.1244.148112.figure10" ID-arpha="D4E2BD3C-7320-5C42-BE83-DCDEF408D74B" httpUri="https://binary.pensoft.net/fig/1366347" startId="F10">
<paragraph id="9EED0F5FA1A1549D6DD1A132D776EB6B">
<label id="81A1BA9A2EC97DDEB7FAADA546FCE6C2">Figure 10.</label>
</paragraph>
<paragraph id="236074A0585BEB641E2C216FBEB5B476">
<taxonomicName id="CAF12AC3C34ECEEB0E6DCEC87E196B33" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="8C0420A19F72EF02CF50B3CB01A52092" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4330F318CC75DD4DE8C8DFD0D914AA3B" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (17.8 × 14.5 mm),
<abbrev content-type="institution" id="ABBRID0ELTBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002152 (
<emphasis id="19EBA2042208EEA3932D06DEBAC91AD8" bold="true">A</emphasis>
<emphasis id="0A9A47D4BD4721550AD83503DC6DC18D" bold="true">C</emphasis>
,
<emphasis id="2C8E537C7DCFE7FF7218BF8BCCC67466" bold="true">F</emphasis>
,
<emphasis id="0EBCDB61506BD4CA9F29786C8D9856C6" bold="true">G</emphasis>
); male paratype (11.6 × 9.5 mm),
<abbrev content-type="institution" id="ABBRID0EYTBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002153 (
<emphasis id="88283E776F94DBFB2BC7943C57FB6C43" bold="true">D</emphasis>
); male paratype (16.0 × 12.7 mm),
<named-content id="5E90CEE357E1422B585978B234752860" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-hydrobiology-chinese-academy-sciences-0" xlink_title="Museum of Aquatic Organism, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, China">IHB</named-content>
(
<emphasis id="F2BEDFE809AE5DECA796F70A2AF16EE9" bold="true">E</emphasis>
); female paratype (16.0 × 12.4 mm),
<abbrev content-type="institution" id="ABBRID0EGUBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002154 (
<emphasis id="D2784BF0BA3DDAE8EFD7A430571E0714" bold="true">H</emphasis>
,
<emphasis id="D2D5BE600B298FEAF344B965E0E1373E" bold="true">I</emphasis>
). Left maxilliped 3 (
<emphasis id="AFFB8EAEF3414F793092E7301026689F" bold="true">A</emphasis>
); left G 2, pleonal view (
<emphasis id="CE4B1D033C9D5A999AB134BA63B8EDB6" bold="true">B</emphasis>
); left G 1, ventral view (
<emphasis id="12C9054470A8F01FFEECA7ECBCFD3FC4" bold="true">C</emphasis>
<emphasis id="56DCFAB311F9744A2C45AB2F5AE8B4D1" bold="true">E</emphasis>
); major cheliped (
<emphasis id="9B879BAC6B251121FF1419BF6F2676A9" bold="true">F</emphasis>
,
<emphasis id="93A6DA9CDA4D47C13A30C26572CB63E5" bold="true">H</emphasis>
); minor cheliped (
<emphasis id="0496DCC32B2E6184B2F390D674D4E271" bold="true">G</emphasis>
,
<emphasis id="CC3BAE38B4BAF543AF9891581762F5AE" bold="true">I</emphasis>
). Scale bars: 1.0 mm
<emphasis id="3047E069A8219B2B21D5DD4BC66B8026" bold="true">(A</emphasis>
<emphasis id="4AEE1B288A68B11FBE58342C0D363C34" bold="true">E)</emphasis>
, 5.0 mm
<emphasis id="2BD5CD08B59CF6404920BE48327227AB" bold="true">(F</emphasis>
<emphasis id="144364C7A800108E7F70D1ACDFEBB4A5" bold="true">I)</emphasis>
.
</paragraph>
</caption>
<caption id="04DF9B8D705C44D2E0E32EF359AA4403" ID-DOI="10.3897/zookeys.1244.148112.figure11" ID-arpha="E898A815-D593-526C-855D-757A916C9F0D" httpUri="https://binary.pensoft.net/fig/1366348" startId="F11">
<paragraph id="A3FB380B607FE64E302A6BE82F26D319">
<label id="2C1E67E1A048232B6E01BBFDDC02854F">Figure 11.</label>
</paragraph>
<paragraph id="2AE79781526A54998329034218B8C4E6">
Left G 1 terminal segments.
<taxonomicName id="4CD61A6547DF69D44A8EA48BCC8C67D8" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="480F30E4B093534022E54537B1FB4A34" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEA5E424F9BFEC8EA0D4BD6055715668" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (11.7 × 9.7 mm),
<abbrev content-type="institution" id="ABBRID0E6VBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002144 (
<emphasis id="0599881D105A7EF7C762925D47D48708" bold="true">A</emphasis>
,
<emphasis id="DC6248F00016BA0E8721C8BD672F9387" bold="true">B</emphasis>
);
<taxonomicName id="B780D830CE19E516EF0E9C496FE8234A" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="3ADEDC482DDA4AFA6B8AA2BE31ACE260" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2F9CCBC33D57BC96B5837B6F84E2A93E" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (17.8 × 14.5 mm),
<abbrev content-type="institution" id="ABBRID0ETWBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002152 (
<emphasis id="779CA2A5CDBB5995BE15CA99FECF7CA4" bold="true">C</emphasis>
,
<emphasis id="BCCD66477E90EE862ECDF556241B45A3" bold="true">D</emphasis>
). Ventral view (
<emphasis id="85C81AE7F8BD2B7E04793DEF8B9A21C8" bold="true">A</emphasis>
,
<emphasis id="B87C561A293F1EA992DA5171AF6A649C" bold="true">C</emphasis>
); dorsal view (
<emphasis id="76717273AEE2A82047951709ADF10FC6" bold="true">B</emphasis>
,
<emphasis id="21143A09A5D7FECF992688E28B0C82DA" bold="true">D</emphasis>
). Scale bar: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EJXBG" type="description">
<paragraph id="139262F7B5C9F93D0B753B6510B14E50">
<heading id="8249133DFF8D5F871FCE6B30A5AD8E52" reason="title">Description.</heading>
</paragraph>
<paragraph id="F4DD042C9E3253207BC2949121D6909C">
Carapace broader than long, males ~ 1.21.3 × as wide as long (
<emphasis id="3695E9B6530BDCADCA5DE113822E248B" italics="true">n</emphasis>
= 6); regions not all visible, dorsal surface slightly convex; surface pitted, anterolateral regions slightly rugose (Fig.
<figureCitation id="786BA7DF165F38EA86B2D3FE298A4178" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Frontal margin sinuous, deflexed (Fig.
<figureCitation id="DD9955A149D8FF67DE2E1867BA522D86" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Epigastric cristae and postorbital cristae confluent, sharp; bifurcated shallow groove between epigastric cristae (Fig.
<figureCitation id="1369F30E823989ADCCB618B71E90A674" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Branchial regions not swollen (Fig.
<figureCitation id="6B19BEDF90AE1CF62195311DB2097158" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Cervical groove obvious (Fig.
<figureCitation id="25ED1539C71E9698BCFDEA5991135B8B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Mesogastric region flat (Fig.
<figureCitation id="D7AA62C0184F2BDB96A31430FA117560" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). External orbital angle broadly triangular, outer margin slightly convex, almost confluent with anterolateral margin (Figs
<figureCitation id="E3D24B8A38148E9C930F63F1183DB94A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
,
<figureCitation id="560C4FA7A815351E6C215DF054EAFED9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Epibranchial tooth granular, indistinct (Figs
<figureCitation id="AB47B955BEC8553D73944A5B25A71330" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
,
<figureCitation id="7432526A5EF78D12FB0D8234C779D9A7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Anterolateral margin lined with 1014 granules; posterolateral margin posteriorly convergent (Fig.
<figureCitation id="EAD342F1C69FAB660F9806CFC79FFF49" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
); posterolateral with weak striae (Fig.
<figureCitation id="CED604CCBB4297F6CB36348B5A779B00" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Orbits regular; supraorbital margins cristate, infraorbital margins lined with fused granules (Fig.
<figureCitation id="1FAAD2A4FE7E24A95E8016DCE9731B23" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Eyes normal (Figs
<figureCitation id="612B13BB650EF0D2C32E592866D9DFBB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
,
<figureCitation id="F781AB6CA8ED82A613BE781E9C6AA090" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Sub-orbital, pterygostomial and sub-hepatic regions generally smooth, pitted (Fig.
<figureCitation id="C4D0BD04C5DBC44DE41D20E4EED15BB4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Antennules large, folded within broad fossae; antennae very short (Fig.
<figureCitation id="243AAE00877BA21C0620A28ABC6DFA46" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Median lobe of epistome buccal margin broadly triangular, lateral margins straight (Fig.
<figureCitation id="431AEB06641A3BBA481919AE619F7548" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
).
</paragraph>
<paragraph id="320385DB1AD5C67E7B253F458C7CA5DC">
Maxilliped 3 merus subtrapezoidal, with slight median depression, width ~ 1.3 × length; ischium subtrapezoidal with very shallow median sulcus, distomesial margin rounded, width ~ 0.7 × length. Exopod reaching proximal one-third of merus; flagellum short (Fig.
<figureCitation id="7A960F12B01419999D78FB3823713515" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 A</figureCitation>
).
</paragraph>
<paragraph id="F481EBEF177D109B0C8EB08C97536B33">
Chelipeds (pereiopod 1) subequal (Fig.
<figureCitation id="336551A5CF5DA384872D782B26AEBF8C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 F I</figureCitation>
). Merus trigonal in cross section, surfaces generally smooth; outer dorsal margin slightly crenulated, inner and ventral margin lined with large granules (Figs
<figureCitation id="BBA70AEAEE88814AFDC057E9FFB98DC1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
,
<figureCitation id="BAEB92DE7F83E18FC52683BB9EC56417" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 A</figureCitation>
). Carpus dorsal surface slightly rugose, with small blunt spine at inner-distal angle, spinule at base (Fig.
<figureCitation id="15FF25B23AF1549E2D050548A1D7C74B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Major cheliped palm length ~ 1.41.5 × height in males (
<emphasis id="ABF9755AC1B8140EBCB20A89B5BA4AAB" italics="true">n</emphasis>
= 3), 1.4 × height in females (
<emphasis id="08912AD5BA4F2F7093EAE780B5F7FDE4" italics="true">n</emphasis>
= 2); dactylus 0.9 × palm length in males (
<emphasis id="4340B7816292F497909FC21A9C6E228A" italics="true">n</emphasis>
= 3), 0.80.9 × palm length in females (
<emphasis id="C322498C53CF9C50E6D2D1D143B0E68E" italics="true">n</emphasis>
= 2) (Fig.
<figureCitation id="F574E925122A42D936DA59071AD17DA3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 F I</figureCitation>
). Palm surface pitted, occlusal margin of fingers with 913 irregular blunt teeth, with very small gape when closed (Fig.
<figureCitation id="240FE93EBC3586E5B47D53B52A5214C9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 F I</figureCitation>
).
</paragraph>
<paragraph id="C341038CF743CB17C31CF2E14287AD56">
Ambulatory legs slender (pereiopods 25), with only very short setae on margins (Fig.
<figureCitation id="9B34C87AC05F1567FCCDB160F069A2D7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Pereiopod 3 merus 0.7 ×
<abbrev id="ABBRID0EH2BG" xlink_title="Carapace length">CL</abbrev>
in both sexes (
<emphasis id="5DAEF456C71A772F7CF13F944B3546DA" italics="true">n</emphasis>
= 5, Fig.
<figureCitation id="0291BD3B9351F8208ABF5EB5528574E1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
). Pereiopod 5 propodus length 1.72.0 × height in males (
<emphasis id="00EC0ED3C4C3A2D800DA4903E51896F8" italics="true">n</emphasis>
= 3), 1.81.9 × height in females (
<emphasis id="6B37EAED7220D4EBA6D0836F395FE463" italics="true">n</emphasis>
= 2), longer than dactylus (Fig.
<figureCitation id="4527BBB4E143C83F0A3A6DFB66FE47BE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
).
</paragraph>
<paragraph id="B7B4481A9E5F666CBDC346BE7F7550C8">
Male thoracic sternum generally smooth, sparsely pitted; sternites 14 width ~ 1.9 × length; sternites 1, 2 fused to form broad triangle; fused sternites 1, 2 demarcated from sternite 3 by slightly sinuous sulcus; sternites 3, 4 fused with distinct sulcus (Fig.
<figureCitation id="E12BF1BD1E9AAB36FCE2760B3BE8ED0E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 B</figureCitation>
). Male sterno-pleonal cavity reaching anteriorly slightly beyond mid-length of cheliped coxa (Fig.
<figureCitation id="6ED336127FE7D7E661A71FB764F3B0AB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 B</figureCitation>
). Male pleonal locking tubercle positioned at mid-length of sternite 5 (Fig.
<figureCitation id="20B5ACC6136AD1330E7ED80AE963692E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 D</figureCitation>
). Female vulvae ovate, large, reaching suture of sternites 5 / 6, relatively widely separated (Fig.
<figureCitation id="7B91E1333AAA20DFF7E9BE25B7357FA9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 F</figureCitation>
).
</paragraph>
<paragraph id="A4E3EBEC7C36E65FD7D1A2A2B29C837B">
Male pleon large, broadly triangular; somites 36 progressively narrower; somite 6 width approximately 2.4 × length; telson width 1.8 × length; lateral margins almost straight, apex rounded (Fig.
<figureCitation id="CFBC146448333B5F36A379F3EF119D05" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 C</figureCitation>
). Female pleon sub-ovate (Fig.
<figureCitation id="72BAC4FDD672F8CE58AD6FFE722CA971" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 E</figureCitation>
).
</paragraph>
<paragraph id="3E02E6550553F0DD5B333E5A1F9B331F">
G 1 large, sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="95923A4145CA98DEBF6E85D8C0322EA6" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="D419EA42B5C1202A55978F1C8B75F160" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 D</figureCitation>
); subterminal segment length ~ 2.7 × length of terminal segment (Fig.
<figureCitation id="EE2FFBC775AA75050DDAFFB2B27BDD09" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 C E</figureCitation>
). Subterminal segment thick, outer margin strongly concave, distal end slanted with outer-distal section highest; terminal segment short, both lateral margins slightly convex, bifurcated with one point being opening tip and other being smaller projection on outer margin, both pointing upwards (Figs
<figureCitation id="0ACA2A59FB21893352D93C7094D712A6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 C E</figureCitation>
,
<figureCitation id="6002878268D6A5215DEAEB16DC8C194D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 C, D</figureCitation>
). G 2 subterminal segment thick, slightly bent outwards distally, flagelliform terminal segment thin, ~ 2.3 × length of subterminal segment (Fig.
<figureCitation id="BA5F6D21985F761A2755C7AEF9B6AD75" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL4BG" type="Colour in life">
<paragraph id="1313FCF6C7AB7B16B6D72E4097E3E671">
<heading id="8EACC05107603168C47DCE924199260A" reason="title">Colour in life.</heading>
</paragraph>
<paragraph id="B9F3B9541D9A4226176267102632CCE1">
Generally camouflaged in light mottled brown overall (Fig.
<figureCitation id="CAA79ED2204A18B5CD8CF2B550B6248F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EV4BG" type="habitat">
<paragraph id="17E1551ECC5FBDF5EDC46401BC6E7783">
<heading id="E2F5A36209C9D405C64AAE4E4C5710DF" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="661ED6223EFA6F91E3E744A9D9E14D50">
This is a typical aquatic hill stream species that can be found residing under rocks in the shallows of the hillstream. The sympatric
<taxonomicName id="F5E5F2B4995C785EE715D0A6D36BAA99" baseAuthorityName="Dai &amp; Turkay" baseAuthorityYear="1997" class="Malacostraca" family="Potamidae" genus="Eurusamon" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="CDBC0B1C6C347BD7B3CDDBA0C199CBAC" italics="true">Eurusamon guangdongense</emphasis>
</taxonomicName>
is also aquatic but can grow to a much larger size and mature individuals occupy the deeper areas of the hillstream.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG5BG" type="distribution">
<paragraph id="986684CB144C18518F7AFDD7F85DD412">
<heading id="1AD4D2EEE80D9332A495FE8B2A5ABCD9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="21046C41768A307E83A58E1833E16BFA">
Yangdong County, Yangjiang City,
<collectingRegion id="043B8B9931209A126B14C22A2E7CC710" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="AA2314AD19BF87F2279C9E4A13EB2CD7" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL5BG" type="etymology">
<paragraph id="4230BAAAD8AD21172FBE4C24679C8920">
<heading id="F9F56A33F954D019E295F88E2FD764ED" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="F91020885A49FE2072E0F6FF6E8D2F67">
This species is named after the
<typeStatus id="1E197A8AB490BDC241D0249385178D96">type</typeStatus>
locality, Yangdong County.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQ5BG" type="remarks">
<paragraph id="C8F1E87D4EFCDB6FE07034D6350E29E7">
<heading id="A43DDEBCF3E5F2245F97A457A91E0BBF" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="E2D47C6FBD3AB00975D77F09361A3FEF">
This new species is closest to
<taxonomicName id="C27AB2D06978043F9D3C15FDEE215FD9" authorityName="Huang, Shih &amp; Ahyong" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ehuangzhang">
<emphasis id="FFEDEC2D43DED6ACA373E7061E5E2135" italics="true">Megapleonum ehuangzhang</emphasis>
</taxonomicName>
and especially
<taxonomicName id="63856C05D2E718AEE3EFCA08B0E16315" authorityName="Huang &amp; Mao" authorityYear="2021" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="shenzhen">
<emphasis id="1C4D9ADBC6E6F6A8EF64F6D085F4CB7A" italics="true">M. shenzhen</emphasis>
</taxonomicName>
in terms of carapace and gonopodal morphology, and the three are no doubt closely related.
<taxonomicName id="CC64588EBDB4E645A9A20C34FDAD6949" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="6B9BC6FFB8A73E28860D222ED9351F21" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38775E4C3559FB084A1ABE8CCAC3CF76" rank="species">sp. nov.</taxonomicNameLabel>
, however, can be distinguished from the other two by its prong-shaped G 1 terminal segment (vs folded terminal segment in
<taxonomicName id="A1A798BE70371E70B23BFFD6C3A6C2F5" authority=", Huang et al. 2018" authorityName="Huang" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ehuangzhang">
<emphasis id="630A24CD08ED0839A09ED14D9DD991C8" italics="true">M. ehuangzhang</emphasis>
,
<bibRefCitation id="9EE40A1E6569BE5B383E55166D2C4F68" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
</taxonomicName>
: fig. 3 C, and goose-head-shaped terminal segment in
<taxonomicName id="46A75F785A274FF9EC4A9A90D94640A0" authority=", Huang and Mao 2021" authorityName="Huang and Mao" authorityYear="2021" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="shenzhen">
<emphasis id="949CA94E2AC8F80283D35F3D4E2D160B" italics="true">M. shenzhen</emphasis>
,
<bibRefCitation id="9E71EC8BF0FC15DD65065677161AC03B" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
</taxonomicName>
: fig. 3 C E). Otherwise, it can further be separated from the two aforementioned congeners in having a slightly narrower male abdominal somite 6, with the width ~ 2.4 × length (vs 2.6 × in both
<taxonomicName id="1D3F94777A804DFC3A4302BD4DF44760" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ehuangzhang">
<emphasis id="594F4DCD32231A312BB215F8B055085D" italics="true">M. ehuangzhang</emphasis>
</taxonomicName>
and
<taxonomicName id="04406F3C2FFFEEBA637C6FA2438F5CE9" authority=", Huang et al. 2018" authorityName="Huang" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="shenzhen">
<emphasis id="0ACD4E6B4649D9894D3710111D75CB6C" italics="true">M. shenzhen</emphasis>
,
<bibRefCitation id="847836D0D48FB09908C57BB1C9339BAC" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="11C8A48B8269B8003D0AA8B757634A6A" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
). More detailed comparisons can be found in Table
<tableCitation id="7975EA510E46D19053466F625708EF5C" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparisons between the species within the genus Megapleonum." httpUri="http://table.plazi.org/id/69D766E64FF15F0F8BB8C5782AADA37B" tableUuid="69D766E64FF15F0F8BB8C5782AADA37B">1</tableCitation>
. Interspecific variation of the G 1 is minimal, with the terminal segment slightly varying in size and the outer projection of the terminal segment slightly varying in sharpness and angle (Fig.
<figureCitation id="16654E9AD3B63C1F23F39D140C08D6E1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 C E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,301 @@
<document id="E64B0A64B1184721A593D446D8943508" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="A4D61C5D66B556D98A865B9133C26EBE" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pleistodontes imperialis Saunders 1883" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="6DC280C469A5F18D46CF6F52AD82F1F4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2B5DCF960599725BA4E018BB71883BC">
<mods:title id="BD856DE4B1541024311BEE3146583B93">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="AFE098B9774A88A6C504FB068B3B1EB2" type="personal">
<mods:role id="DD34074FC9C1C5D432538ADE294295CA">
<mods:roleTerm id="8F21ACFDD6AB51D23046AA53421FBD63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC66B35E7E89B9D46FE15B508E06AB86">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="50AE485B7A60A84C75CC14926C07E91D" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="320D5801EB12385CB6124F3DF143E4EE">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="172C3617D1FC9AEF92E64A0D61CA5522" type="personal">
<mods:role id="1F641C7A5ACF093021D8DED62A7053BA">
<mods:roleTerm id="444D2CA30F100B3AE50D645966F8816E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA8153ED03C74D2FF86D1B913C5AA1BE">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="0D950B7BD22B982F8CDA9511D2FBF246" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="218D6DC9B46507C2EDDBD00C892DF6A4">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="0A4485576FB23FAAB85C92210A419167" type="personal">
<mods:role id="579C08FACD8639515B649C0552FF09E5">
<mods:roleTerm id="4B60F3AC456D3B2375DAECA650630DF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C2D86FE21D0AFA026D43961B1D7D5EB">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="CEE25081D9E0DCECEFEF1F60BC5A5029" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="AE4ABBBE9346F1275BB3574A6846A16C">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="5F12939CBE6272DF341061BED7AE5584" type="personal">
<mods:role id="EC4E9749882AE859B75DB8633F7853F7">
<mods:roleTerm id="2EF9F0026D027385E4798C74B1B20AFD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C151E658901A33E3A8FA810DD3CF4ABA">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="C4FCAEC59B247AA876759CB06FBAF9A5" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="F13786AED27640DE8660E2BEFAFF2276">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="04A2A430B9E9DA62DE12A2DAA361DB22" type="personal">
<mods:role id="E45CAC481F0893F0F6D891732CB32BAA">
<mods:roleTerm id="68747C3F9D5BC59BC83629EFEFC66ACE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13F21461B9F0E66BD4BDB03096B07DED">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="BD045B6F835A58FDC415646CC6DA0893" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="77F6434D3BD7FC5707132195C57FDF8A">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EE1C90E0F3500759FD13ABCA47A50275">text</mods:typeOfResource>
<mods:relatedItem id="A7CD2A211F60587313E969880A9BF63C" type="host">
<mods:titleInfo id="B70D56CC2FA42E8766031F98869B99D3">
<mods:title id="9E8DD13AD4E5332C6CB78CF5091F762E">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="4722DE5231FAD9B076C8711FC8EE195C">
<mods:date id="53DAE48E756D5987B58E9C9C12054428">2025</mods:date>
<mods:detail id="2B3C23946B2FF615DD64289D7BE390B6" type="pubDate">
<mods:number id="9A783ED66DF6D8FA8838D7FC38D3BFF2">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="0D86472DC0187F45C364F119F1320ADB" type="volume">
<mods:number id="F573D088BBEE9D427433954749E3ABEA">98</mods:number>
</mods:detail>
<mods:extent id="1C5F69F37C1067C788AEB7C32961542B" unit="page">
<mods:start id="54F57FAE940765273C975B740B1F66CD">667</mods:start>
<mods:end id="29564815323C433D14B7ABEB352BA52D">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EBBBDD858C3BB33494DADB5E70486CB5">journal article</mods:classification>
<mods:identifier id="D1561089C69CF6F31F1E6C93F6E6C708" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="8E2DCEF7A2209F65DD7EB4FEABBF5450" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="A4D61C5D66B556D98A865B9133C26EBE" ID-DOI="http://doi.org/10.5281/zenodo.15791850" ID-Zenodo-Dep="15791850" ID-arpha="A4D61C5D-66B5-56D9-8A86-5B9133C26EBE" LSID="urn:lsid:plazi:treatment:A4D61C5D66B556D98A865B9133C26EBE" httpUri="http://treatment.plazi.org/id/A4D61C5D66B556D98A865B9133C26EBE">
<subSubSection id="BF6A90363A237280E0C471F8DB6D01CE" type="nomenclature">
<paragraph id="A44712C64168A54847D63921BD48999E">
<taxonomicName id="587F2266991799B2686031EEE2E69BE6" ID-CoL="4K6PG" authority="Saunders, 1883" authorityName="Saunders" authorityYear="1883" class="Insecta" family="Agaonidae" genus="Pleistodontes" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="5F310B12569567F8E05E5BAF3EAA6CB3" italics="true">Pleistodontes imperialis</emphasis>
Saunders, 1883
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYTAE" type="material">
<paragraph id="10507DCFD1003890AE714FBD63FCA15F">
<heading id="336D28DFA278018BD597BF43137E6A3B" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="90B1790E880018AEA42F46A1DDE9EAE1">
<materialsCitation id="DA93A72A28A82AE810598D202863DC08" collectingDate="2018-12-07" collectionCode="AC" collectorName="Jerez de la Frontera &amp; de Jerez &amp; I. Sanchez" country="Spain" elevation="82" latitude="36.689316" location="Zoobotanico de Jerez" longLatPrecision="1" longitude="-6.150638" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Cadiz">
<emphasis id="C953D5512A98A5ADF677B57C7BB89417" bold="true">
<collectingCountry id="F6143A2347776FB6F5225F25FC42CF39" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="B1493EE09DE3952E34DB3F4486E0E6CC" bold="true">
<collectingRegion id="63A0493D3FCDFC8C8B4ACAB1EF0275D3">Cádiz</collectingRegion>
</emphasis>
:
<collectorName id="A2E74020FD85EED0D8CD9377C20103E1">Jerez de la Frontera</collectorName>
(
<location id="FFC51D73D42D47CCBE2ACBCA7F657D63" LSID="urn:lsid:plazi:treatment:A4D61C5D66B556D98A865B9133C26EBE:FFC51D73D42D47CCBE2ACBCA7F657D63" country="Spain" latitude="36.689316" longLatPrecision="1" longitude="-6.150638" name="Zoobotanico de Jerez" stateProvince="Cadiz">
Zoobotánico
<collectorName id="27822E416BC7E40784DBE73A44142606">de Jerez</collectorName>
</location>
):
<specimenCount id="F9CAC6706DE4ADCBB55E28C192588358" count="2" type="male">2 ♂</specimenCount>
+
<specimenCount id="93AD7D1574F35CEFCFE4D383FEBDEA82" count="2" type="female">2 ♀</specimenCount>
<abbrev id="ABBRID0ECUAE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="DDFA5A7D687AD080BBAD5BC134CC95EB">Í. Sánchez</collectorName>
<abbrev id="ABBRID0EGUAE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="D10DF9FCA978C3D336113500F0E17BD8" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EKUAE" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="882DA2BEB58FE53DB96B5545093B1371" value="2018-12-07">
<collectingDate id="E1BC849FCB64D280F8CB8E94BCF37598" value="2018-12-07">7 - XII- 2018</collectingDate>
</date>
,
<elevation id="27E800EC62E2B961EDBDF038798D1176" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">
<quantity id="D25D60F5209EE54435A97460467F8709" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">82 m</quantity>
a. s. l.
</elevation>
, found in dissected syconia of
<taxonomicName id="3A52F846D7E85C2742F5EB9F31EA651E" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="rubiginosa">
<emphasis id="F7246D6F72461735A15E3D89E483DE39" italics="true">Ficus rubiginosa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3UAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-6.150638,36.689315]}">
<geoCoordinate id="8AF02D45B736F7F220F9B27632580E5E" degrees="36.689315" direction="north" orientation="latitude" precision="1" value="36.689316">36.689315</geoCoordinate>
,
<geoCoordinate id="CAEA615423A0F9FA3253C63F25590185" degrees="6.150638" direction="west" orientation="longitude" precision="1" value="-6.150638">- 6.150638</geoCoordinate>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="D5A8C162606C4BECEA89917587BBED82" collectingDate="2023-07-05" collectionCode="AC" collectorName="Vicente Savall Pascual &amp; J. M. Royo" country="Spain" county="San Vicente del Raspeig" latitude="38.389034" location="See in Fig." longLatPrecision="1" longitude="-0.51813" municipality="University area" specimenCount="17" specimenCount-female="9" specimenCount-male="8" stateProvince="Alicante">
<emphasis id="C27F5E3432D376E66B67ED9695C14880" bold="true">
<collectingRegion id="8A086184FE9E66FDC57716DC08A0DDFC">Alicante</collectingRegion>
</emphasis>
:
<collectingCounty id="57D7B7308C5A897CADA984C488F33AA5">San Vicente del Raspeig</collectingCounty>
:
<collectorName id="40587DD45D95FD91FCEF284257843445">Vicente Savall Pascual</collectorName>
street and
<collectingMunicipality id="08DD9D77637C9A98BBEB102EB5038A22">University area</collectingMunicipality>
:
<specimenCount id="A01CC3105552A00A408B9B9C546061CA" count="7" type="female">7 ♀</specimenCount>
+
<specimenCount id="51A0F82B9E1030FBEE76FC16B9ADB253" count="8" type="male">8 ♂</specimenCount>
<abbrev id="ABBRID0EDVAE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="FAB86B1F0806278ED16A268618B19AD9">J. M. Royo</collectorName>
<abbrev id="ABBRID0EHVAE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="D3B47BD688418185C6FCA1F974CFF8D1" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0ELVAE" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="9F68513EADAC86D24895B53AA999E504" value="2023-07-05">
<collectingDate id="26164F2F538D93E32151994003576B65" value="2023-07-05">
5 - VII-
<quantity id="7D42353289DE5FD5CFA7D7A41C7582C7" metricMagnitude="6" metricUnit="m" metricValue="2.023193" unit="m" value="2023193.0">2023, 193 m</quantity>
</collectingDate>
</date>
a. s. l., found in dissecting fallen syconia of
<taxonomicName id="6FAB0CCA9C49F4B451FDA775CB5CB679" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="rubiginosa">
<emphasis id="DFBA8B7A025E5943F3EA5489A781A491" italics="true">Ficus rubiginosa</emphasis>
</taxonomicName>
tree, excepted
<specimenCount id="034A6950601022A328A1B67540C74923" count="2" type="female">two females</specimenCount>
found in syconia of
<taxonomicName id="CEB956B00C408F81DE3E47E32CD0BBE8" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="macrophylla">
<emphasis id="7B1603218946A60F48FDFB04B02A7CE2" italics="true">Ficus macrophylla</emphasis>
</taxonomicName>
in an urban area (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIWAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-0.518130,38.389033]}">
<geoCoordinate id="25B883599A53BFE771FD1ABA1907F862" degrees="38.389033" direction="north" orientation="latitude" precision="1" value="38.389034">38.389033</geoCoordinate>
,
<geoCoordinate id="D48C8F32C5D0114DD23915FC67C9FB7F" degrees="0.518130" direction="west" orientation="longitude" precision="1" value="-0.51813">- 0.518130</geoCoordinate>
</named-content>
).
<location id="17F1D7401D4C34FB115EB0A33ED50046" LSID="urn:lsid:plazi:treatment:A4D61C5D66B556D98A865B9133C26EBE:17F1D7401D4C34FB115EB0A33ED50046" country="Spain" county="San Vicente del Raspeig" latitude="38.389034" longLatPrecision="1" longitude="-0.51813" municipality="University area" name="See in Fig." stateProvince="Alicante">See in Fig.</location>
<figureCitation id="6C5DCD5E23C6B6C35DF77277C7697674" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 C, D</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERWAE" type="Ecological remarks">
<paragraph id="3A0338C0C068D01A5B46476EF87DA30A">
<heading id="43DD771941AE28F13EBC1C85F06BD241" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="C5016296CB974B5D9597A14FD29EF009">
Pollinator of
<taxonomicName id="875B791496AA6D1A92AF75397E1803A8" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="rubiginosa">
<emphasis id="DCEEC7F6C090FC45DB78385EEC359BE0" italics="true">Ficus rubiginosa</emphasis>
</taxonomicName>
.
<specimenCount id="C566A0B727339BD0D2BACCB0EB74114A" count="1" type="generic">One specimen</specimenCount>
of
<taxonomicName id="9AF154EF4AC13CF472CB70E9C259FF3B" authorityName="Saunders" authorityYear="1883" class="Insecta" family="Agaonidae" genus="Pleistodontes" isUncertain="true" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="D23045DFAF284EC1421B840529BB6A1D" italics="true">Pleistodontes</emphasis>
cf.
<emphasis id="B7DB0C6C2CA6B8410A1F34CFEF424D1C" italics="true">imperialis</emphasis>
</taxonomicName>
was found inside a syconia of
<emphasis id="62FE3B76DC672446D27BB00FBF4984F4" italics="true">
<taxonomicName id="D811F7706E82874CFA7DB2329F9CADFB" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="macrophylla">Ficus macrophylla</taxonomicName>
f. columnaris
</emphasis>
(C. Moore) D. J. Dixon (
<bibRefCitation id="865A4089AA1A42E2F1A1872548FB67EA" author="Speciale" etAl="et al." firstAuthor="Speciale" issue="2" journalOrPublisher="Il Naturalista siciliano" pagination="399-406" refId="B86" refString="Speciale M, Cerasa G, Lo Verde G (2015) First record in Europe of seedling of Ficus macrophylla f. columnaris (Moraceae) and of its pollinating waps Pleistodontes cf. imperialis (Chalcidoidea Agaonidae). Il Naturalista siciliano 39 (2): 399406." title="First record in Europe of seedling of Ficus macrophylla f. columnaris (Moraceae) and of its pollinating waps Pleistodontes cf. imperialis (Chalcidoidea Agaonidae)." volume="39" year="2015">Speciale et al. 2015</bibRefCitation>
). We have found
<specimenCount id="34132F20AA81C85DFE52BFFC03E9A5E0" count="2" type="female">two female</specimenCount>
specimens as well in syconia of
<taxonomicName id="063571AAC4FEF6A10AA41FCB4EC3704E" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="macrophylla">
<emphasis id="4A356E256A21169816FF1ABCDC929D72" italics="true">F. macrophylla</emphasis>
</taxonomicName>
. It has also been found in
<taxonomicName id="8AB54BC31E30C5E19D089C31CB024ECB" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="watkinsiana">
<emphasis id="10F791ED4DD8DF7F0B1681F7730851DA" italics="true">Ficus watkinsiana</emphasis>
</taxonomicName>
in
<collectingCountry id="8D248E7849F92D8AAC8D0355308F3402" name="Greece">Greece</collectingCountry>
(
<bibRefCitation id="2647701DBC6C34C0A4B0728CE50FF65D" DOI="10.1007/s12600-024-01158-2" author="Koutsoukos E &amp; Demetriou J &amp; Martinou AF &amp; Compton SG &amp; Avtzis DN &amp; Rasplus J-Y" refId="B58" refString="Koutsoukos E, Demetriou J, Martinou AF, Compton SG, Avtzis DN, Rasplus J-Y (2024 a) Playing both fig sides; the presence and host-switch of Pleistodontes imperialis (Hymenoptera: Agaonidae) is confirmed in Greece and Cyprus. Phytoparasitica 52: 39. https://doi.org/10.1007/s12600-024-01158-2">Koutsoukos et al. 2024 a</bibRefCitation>
) and mentioned in
<taxonomicName id="567A331922807A46A419CF29B23F2E4F" authority="Roxb. Ex Hornem." authorityName="Roxb. Ex Hornem." class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="elastica">
<emphasis id="4BEA162ED7206DD46CE61A7FA2350990" italics="true">Ficus elastica</emphasis>
Roxb. Ex Hornem.
</taxonomicName>
without additional data (
<bibRefCitation id="B34A7E967CD56CACE0FA18A6890F86FA" author="Falco-Gari" etAl="et al." firstAuthor="Falco-Gari" refId="B32" refString="Falcó-Garí JV, Oltra-Moscardo MT, Peris-Felipo FJ, Jiménez-Peydro R (2010) Exotic fig pollinators and associates (Chalcidoidea: Agaonidae) of Ficus trees in Europe. Seventh international Congress of Hymenopterists. Hungary, Programme and Abstracts." year="2010">Falcó-Garí et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENZAE" type="distribution">
<paragraph id="F34C1ACB2FF57E29A959657C15FAC764">
<heading id="86D22D314A2733403F366CD6D2CAB7A1" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A2F5539E767B665691634370907D82C2">
Native from eastern
<collectingCountry id="149ED1C0E83059165CA3DE5ED39D5789" name="Australia">Australia</collectingCountry>
(
<bibRefCitation id="5A2EDF0C8F60C62C5C835E5A3F56F58B" author="van Noort and Rasplus" firstAuthor="van Noort" refId="B91" refString="van Noort S, Rasplus J-Y (2024) Figweb: figs and fig wasps of the world. www. figweb. org [Accessed on 1 May 2024]" url="www. figweb. org" year="2024">van Noort and Rasplus 2024</bibRefCitation>
) and already reported from
<collectingCountry id="709276D23FD2669A6950E9AF8F1771EB" name="Israel">Israel</collectingCountry>
,
<collectingCountry id="EF48F9FA4EF5CBCA420CE690E8C7EA36" name="New Zealand">New Zealand</collectingCountry>
and
<collectingCountry id="31CDCE40F2CAB6D98488EA5A2D3915BC" name="United States of America">United States</collectingCountry>
(
<bibRefCitation id="7EE251145C6BFAE75BC828C9ED019AE5" DOI="10.1046/j.1096-3642.2002.00040.x" author="López-Vaamonde C &amp; Dixon DJ &amp; Cook JM &amp; Rasplus J-Y" issue="4" journalOrPublisher="Zoological journal of the Linnean Society" pagination="637-683" refId="B62" refString="López-Vaamonde C, Dixon DJ, Cook JM, Rasplus J-Y (2002) Revision of the Australian species of Pleistodontes (Hymenoptera: Agaonidae) fig-pollinating wasps and their host-plant associations. Zoological journal of the Linnean Society 136 (4): 637683. https://doi.org/10.1046/j.1096-3642.2002.00040.x" title="Revision of the Australian species of Pleistodontes (Hymenoptera: Agaonidae) fig-pollinating wasps and their host-plant associations." volume="136" year="2002">López- Vaamonde et al. 2002</bibRefCitation>
). The species has been found from several Mediterranean islands in which it was introduced recently:
<collectingRegion id="9B4393F8F2A67A514DDF6AFEFE536636" country="Italy" name="Sicilia">Sicily</collectingRegion>
in
<collectingCountry id="0A199392E512111CF6D6E4C84C9BE146" name="Italy">Italy</collectingCountry>
(
<bibRefCitation id="118B963377AF350CF87ECE1F5223F6E1" author="Lo Verde" etAl="et al." firstAuthor="Lo Verde" refId="B64" refString="Lo Verde G, Porcelli F, Bella S, Rasplus J-Y (2007) Imenotteri Agonidi nuovi per lEuropa e loro ruolo nella naturalizzazione di Ficus spp. in Italia. Atti XXI CNIE, Campobasso, 60." year="2007">Lo Verde et al. 2007</bibRefCitation>
;
<bibRefCitation id="369126B2E932A0B7CFF9622486045014" author="Speciale" etAl="et al." firstAuthor="Speciale" issue="2" journalOrPublisher="Il Naturalista siciliano" pagination="399-406" refId="B86" refString="Speciale M, Cerasa G, Lo Verde G (2015) First record in Europe of seedling of Ficus macrophylla f. columnaris (Moraceae) and of its pollinating waps Pleistodontes cf. imperialis (Chalcidoidea Agaonidae). Il Naturalista siciliano 39 (2): 399406." title="First record in Europe of seedling of Ficus macrophylla f. columnaris (Moraceae) and of its pollinating waps Pleistodontes cf. imperialis (Chalcidoidea Agaonidae)." volume="39" year="2015">Speciale et al. 2015</bibRefCitation>
),
<collectingRegion id="60AC0F2BBA63074E3DCDB9DD4591AA3B" country="Malta" name="Zebbug">Gozo</collectingRegion>
in
<collectingCountry id="21228DC69CC435728A996BE9690EB00C" name="Malta">Malta</collectingCountry>
(
<bibRefCitation id="6B72AB18B8B1756EAE532BD2A9E256B6" author="Mifsud D &amp; Falzon A &amp; Malumphy C &amp; De Lillio E &amp; Vovlas N &amp; Porcelli F" journalOrPublisher="Bulletin of the Entomological Society of Malta" pagination="5-34" refId="B68" refString="Mifsud D, Falzon A, Malumphy C, De Lillio E, Vovlas N, Porcelli F (2012) On some arthropods associated with Ficus species (Moraceae) in the Maltese Islands. Bulletin of the Entomological Society of Malta 5: 534." title="On some arthropods associated with Ficus species (Moraceae) in the Maltese Islands." volume="5" year="2012">Mitsud et al. 2012</bibRefCitation>
), Canary Islands (
<bibRefCitation id="00EE52DE33AC2B96AE7F50CE64B96872" author="Reyes-Betancort" etAl="et al." firstAuthor="Reyes-Betancort" issue="1-2" journalOrPublisher="Boletín de la Asociación Española de Entomología" pagination="85-87" refId="B80" refString="Reyes-Betancort JA, Hernández-Suárez E, Polaszek A (2013) First record of Pleistodontes imperialis Saunders, 1882 (Hymenoptera: Chalcidoidea, Agaonidae) in the Canary Islands. Boletín de la Asociación Española de Entomología 37 (12): 8587." title="First record of Pleistodontes imperialis Saunders, 1882 (Hymenoptera: Chalcidoidea, Agaonidae) in the Canary Islands." volume="37" year="2013">Reyes-Betancort et al. 2013</bibRefCitation>
) and
<collectingCountry id="2204E4DC0A613D2C61EB407A3BE859F7" name="Greece">Greece</collectingCountry>
and
<collectingCountry id="7608E45A7765F4DD8A520AE7D222FECF" name="Cyprus">Cyprus</collectingCountry>
(
<bibRefCitation id="B3BC47FDD3B8CFF962273297B45B8D04" DOI="10.31184/G00138894" author="Compton SG &amp; Newton H &amp; Stavrinides M &amp; Kaponas C" journalOrPublisher="Entomologists Gazette" pagination="121-124" refId="B17" refString="Compton SG, Newton H, Stavrinides M, Kaponas C (2020 a) First confirmed records of fig wasps (Hymenoptera: Chalcidoidea) associated with the Sycamore Fig. Ficus sycomorus on a Mediterranean Island. Entomologists Gazette 71: 121124. https://doi.org/10.31184/G00138894.712.1735" title="First confirmed records of fig wasps (Hymenoptera: Chalcidoidea) associated with the Sycamore Fig. Ficus sycomorus on a Mediterranean Island." volume="71">Compton et al. 2020 a</bibRefCitation>
;
<bibRefCitation id="483A9CED1328FA7ABF76E66A83C452D4" DOI="10.1007/s12600-024-01158-2" author="Koutsoukos E &amp; Demetriou J &amp; Martinou AF &amp; Compton SG &amp; Avtzis DN &amp; Rasplus J-Y" refId="B58" refString="Koutsoukos E, Demetriou J, Martinou AF, Compton SG, Avtzis DN, Rasplus J-Y (2024 a) Playing both fig sides; the presence and host-switch of Pleistodontes imperialis (Hymenoptera: Agaonidae) is confirmed in Greece and Cyprus. Phytoparasitica 52: 39. https://doi.org/10.1007/s12600-024-01158-2">Koutsoukos et al. 2024 a</bibRefCitation>
). As the reports by
<bibRefCitation id="9C5BDBC4B5EDE88E0CB85234051E8E58" author="Falcó-Garí JV &amp; Oltra-Moscardo MT &amp; Peris-Felipo FJ &amp; Jiménez-Peydro R" etAl="et al." firstAuthor="Falco-Gari" refId="B32" refString="Falcó-Garí JV, Oltra-Moscardo MT, Peris-Felipo FJ, Jiménez-Peydro R (2010) Exotic fig pollinators and associates (Chalcidoidea: Agaonidae) of Ficus trees in Europe. Seventh international Congress of Hymenopterists. Hungary, Programme and Abstracts." year="2010">Falcó-Garí et al. (2010)</bibRefCitation>
on eastern Spanish have no localities, we consider this as a first confirmed record for the Iberian Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX1AE" type="description">
<paragraph id="9275814A0361EFC353809A9CA338CC8D">
<heading id="8827E37E5C43E10BEEBDD26BEBA7898F" reason="title">Short description.</heading>
</paragraph>
<paragraph id="ECA40D0B4B866C95B46C9E00D15F65A0">
Small species
<quantity id="863ECD0AA47C09A8BBB8567A4AB14924" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
. Female of
<taxonomicName id="FDBF06F603CAACA2802694AECA0C9F85" authorityName="Saunders" authorityYear="1882" class="Insecta" family="Agaonidae" genus="Pleistodontes" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="63B17C34769B68D1E07FCA0EA13FB634" italics="true">Pleistodontes</emphasis>
</taxonomicName>
can easily be recognized by their elongated head, with long subparallel genae, mandibular appendage elongate, usually bearing more than 20 transverse laminae, rarely less, sometimes bearing transverse rows of small teeth (
<bibRefCitation id="6919BE44D6B001676CCEDB591FF82D7B" author="Boucek" firstAuthor="Boucek" journalOrPublisher="International, Wallingford, Oxon" refId="B5" refString="Bouček Z (1988) Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species. C. A. B. International, Wallingford, Oxon, 832 pp." title="Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species. C. A. B." year="1988">Bouček 1988</bibRefCitation>
). Antennal scape usually elongate, sometimes shorter, mostly bearing a dorsal lamina which can be curved outwards and downwards. Pedicel rather short without dorsal spines. Third antennal segment elongates into a triangular process, always undivided and frequently elongate. Some species have no triangular expansion on the third antennal segment. Mesosoma bears pollen pockets, sometimes reduced, or absent. The fore tibia usually bears 23 spines in the dorso-apical comb, rarely only one. Ovipositor sheaths about as long as or shorter than metasoma. Venation is always complete (
<bibRefCitation id="A4FE9D62E010CE891E003074A0DAD938" DOI="10.1046/j.1096-3642.2002.00040.x" author="López-Vaamonde C &amp; Dixon DJ &amp; Cook JM &amp; Rasplus J-Y" issue="4" journalOrPublisher="Zoological journal of the Linnean Society" pagination="637-683" refId="B62" refString="López-Vaamonde C, Dixon DJ, Cook JM, Rasplus J-Y (2002) Revision of the Australian species of Pleistodontes (Hymenoptera: Agaonidae) fig-pollinating wasps and their host-plant associations. Zoological journal of the Linnean Society 136 (4): 637683. https://doi.org/10.1046/j.1096-3642.2002.00040.x" title="Revision of the Australian species of Pleistodontes (Hymenoptera: Agaonidae) fig-pollinating wasps and their host-plant associations." volume="136" year="2002">López- Vaamonde et al. 2002</bibRefCitation>
).
<taxonomicName id="81AA1A0793E0D1E308BA3CBE00E6F8AC" authorityName="Saunders" authorityYear="1883" class="Insecta" family="Agaonidae" genus="Pleistodontes" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="7E9095AD9F5EABDE8AE58B2E64601D6C" italics="true">P. imperialis</emphasis>
</taxonomicName>
has been demonstrated to be a complex of morphologically related species (
<bibRefCitation id="433CB7DEF8E1F61A829C2B3B300D1A73" DOI="10.1071/ZO15011" author="Sutton" etAl="et al." firstAuthor="Sutton" issue="2" journalOrPublisher="Australian Journal of Zoology" pagination="122-126" refId="B89" refString="Sutton TL, Reuter C, Riegler M, Cook JM (2015) Characterisation of microsatellite markers for fig-pollinating wasps in the Pleistodontes imperialis species complex. Australian Journal of Zoology 63 (2): 122126. https://doi.org/10.1071/ZO15011" title="Characterisation of microsatellite markers for fig-pollinating wasps in the Pleistodontes imperialis species complex." volume="63" year="2015">Sutton et al. 2015</bibRefCitation>
). The wasps discussed here share all characters of the nominal species and morphologically the species occurring in Europe belong to the nominal entity.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,557 @@
<document id="986EB9FEEA81B9B07417F7BDD685BFDD" ID-DOI="10.3897/zookeys.1244.148112" ID-ZooBank="CCCA8237-6587-44BD-A25E-41BD00928A11" ID-publisher-id="148112" URI-arpha="9A0DFA26-0AED-523E-937A-37D6A902E41E" URI-zoobank="http://zoobank.org/CCCA8237-6587-44BD-A25E-41BD00928A11" article-type="research-article" checkinTime="1751465287194" checkinUser="pensoft" docAuthor="Huang, Chao, Shih, Hsi-Te &amp; Ahyong, Shane T." docDate="2025" docId="C80D0340D9645FCD8913422A01FB4647" docLanguage="en" docName="ZooKeys 1244: 1-27" docOrigin="ZooKeys 1244" docSource="https://zookeys.pensoft.net/article/148112/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megapleonum wangjiani Huang, Shih &amp; Ahyong, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="27" masterDocId="9A0DFA260AED523E937A37D6A902E41E" masterDocTitle="Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="1" updateTime="1751465704327" updateUser="ExternalLinkService">
<mods:mods id="F70941B4BE52050633EE746088215E6E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="70D54E46E009FB762D0B3457114C0723">
<mods:title id="5BC93766D8925399D29D313318AA6916">Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="6C35F9FC608C7A08BCD2885F804366C7" type="personal">
<mods:role id="7BC0D5AE5B443799A71B060455F54B37">
<mods:roleTerm id="7A900ED849C3257951DFB530BB9AFCC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99C89FFB0C2C40CCC9109803ABFDB038">Huang, Chao</mods:namePart>
<mods:nameIdentifier id="A4F52F01B797BABD9B3287D9362CDD2E" type="ORCID">0000-0002-3782-9096</mods:nameIdentifier>
<mods:affiliation id="EE100BF9FF1415249EC582A9C4BE1D39">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="263758182E9CB738FE32580F4B112D39" type="personal">
<mods:role id="FB42B522FF0768E66492BDCBBEAE8085">
<mods:roleTerm id="58C706959A528EC44FFF509D173AB452">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B94EDF28FFD393C8C043EBBF2BB9B932">Shih, Hsi-Te</mods:namePart>
<mods:nameIdentifier id="0AAD2E7F9823A924B31A0E7C2806B7A1" type="ORCID">0000-0003-1317-8783</mods:nameIdentifier>
<mods:affiliation id="46D8A43130CFE8022151BC9401B98BFB">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="08A7066DC82DEBF27A7A637447E73560" type="personal">
<mods:role id="12496BE889DD3053EF07D366E57BBA47">
<mods:roleTerm id="F173E1E31CCBD5FE32FC4954F204D4DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB868864EBC1F9FCC58ADAB679453068">Ahyong, Shane T.</mods:namePart>
<mods:nameIdentifier id="A231980261E4711F344152BD35E02039" type="ORCID">0000-0002-2820-4158</mods:nameIdentifier>
<mods:affiliation id="AF4A15DB2CA78FAC42C0B4E997DF800D">Australian Museum, 1 William St, Sydney NSW 2010, Australia &amp; School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="CCF0338974FF9F8059E926851406DCC8">text</mods:typeOfResource>
<mods:relatedItem id="7D237A37A9FFCA612424F5CCA51E1BEA" type="host">
<mods:titleInfo id="8A63E92EE7158955C631D2A59CB7973E">
<mods:title id="A916217138921D3DB67812355CFA0991">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F7AF91460133300DCEFF03C759A0B87C">
<mods:date id="59E353C68BE08153D85AF2B4CA1FFE81">2025</mods:date>
<mods:detail id="316521C664D06BF776456E5ECB92B81A" type="pubDate">
<mods:number id="04F042C26DC0265E10BFD6584D0B1D68">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="3CA0E595C0F8D9560ED0F657053B41A5" type="volume">
<mods:number id="38D9AA1E5F440D787A55B288DCA8C896">1244</mods:number>
</mods:detail>
<mods:extent id="AF446E32552A79EEEE7A625CABE5618B" unit="page">
<mods:start id="679EA16B7BE4F2BC84C37CF4E9703EB0">1</mods:start>
<mods:end id="285F7CB13B0A10034EC0A7AFE73D0EC6">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E17B7B9EB7E62AE0CCD30C948C10F684">journal article</mods:classification>
<mods:identifier id="E9B75F09AFCC77CEBCB51B90091AA62D" type="DOI">10.3897/zookeys.1244.148112</mods:identifier>
<mods:identifier id="7164D163D31259C48C7A7EC0E894EF23" type="ZooBank">CCCA8237-6587-44BD-A25E-41BD00928A11</mods:identifier>
</mods:mods>
<treatment id="C80D0340D9645FCD8913422A01FB4647" ID-DOI="http://doi.org/10.5281/zenodo.15791872" ID-Zenodo-Dep="15791872" ID-arpha="C80D0340-D964-5FCD-8913-422A01FB4647" ID-zoobank="https://zoobank.org/8E66307C-9F4C-4466-85D1-C7CBE7D6DB52" LSID="urn:lsid:plazi:treatment:C80D0340D9645FCD8913422A01FB4647" LSID-ZBK="urn:lsid:zoobank.org:act:8E66307C-9F4C-4466-85D1-C7CBE7D6DB52" httpUri="http://treatment.plazi.org/id/C80D0340D9645FCD8913422A01FB4647">
<subSubSection id="1D20A05709A2E9C3AC469812977A1612" type="nomenclature">
<paragraph id="8FF97CEAAB2B871D1E34AC83CC417081">
<taxonomicName id="38C8B2A7B4EADC5C4021ECF906559778" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="867CFE47521E8BE2B92FAF2617EA7DE0" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E19B98A9C3D1B5477D736AB1FEC543" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FA1BD04CFD09E371FDFA316B0146BB15" type="description">
<paragraph id="B6B290EA07F70D5D66D0FF4F142C3E57">
<figureCitation id="F8B64A0BCBDEBA05BA0B6F637E706F37" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">Figs 5</figureCitation>
,
<figureCitation id="A5ACB92C799DDB746A1A68658D630A23" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6</figureCitation>
,
<figureCitation id="2B8042AEF1C39B4ADF2DEF97B60D6DE8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7</figureCitation>
,
<figureCitation id="1489E96A4F03E81FC280EC015A1181C8" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 A, B</figureCitation>
,
<figureCitation id="6BDBC224568B3F6DF595F44A73D2BA6C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 B, C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4WAG" type="type material">
<paragraph id="D11D782FD07D350F6EEFC069CEC80366">
<heading id="1F938AB7E9CAF6259F3373619B2069EC" reason="title">Type material.</heading>
</paragraph>
<paragraph id="414BD280C14043D727509917950525BA">
<materialsCitation id="0BE7D778AB808958671A9D7FADCAD40C" collectingDate="2018-11" collectionCode="SYSBM" collectorName="Chao Huang" country="China" county="Maoming City" latitude="22.29" location="Datianding" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" specimenCode="SYSBM 002144" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="0E980FB34728277A55C37D71DB3CD4CF" bold="true" italics="true">
<typeStatus id="19907C5AF75CAB498472BBE09F861DCF" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCode id="F0A2A1CF6E11786989DDDD49AFE90202" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EIXAG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002144
</specimenCode>
, male (11.7 ×
<quantity id="C882A7665B31FB59F06117771FB62E3F" metricMagnitude="-3" metricUnit="m" metricValue="9.7" unit="mm" value="9.7">9.7 mm</quantity>
),
<location id="A599628CF9196E6A2DBB4587E38CE52E" LSID="urn:lsid:plazi:treatment:C80D0340D9645FCD8913422A01FB4647:A599628CF9196E6A2DBB4587E38CE52E" country="China" county="Maoming City" latitude="22.29" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" name="Datianding" stateProvince="Guangdong">Datianding</location>
,
<collectingMunicipality id="BF9B5CF4476BDC42646C5BCC34ABEC02">Dawuling Nature Reserve</collectingMunicipality>
,
<collectingCounty id="6393B6ADEEBA892F03635C02B8F77954">Maoming City</collectingCounty>
,
<collectingRegion id="B1F2CB702EF15FB5EAA5BAA02A72C7FF" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="6FD8A9A6A1B873A32EA7EF8E7D887EA2" name="China">China</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQXAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[111.220000,22.290000]}">
<geoCoordinate id="1CBEF4774D59B11EA74116D10C15F858" degrees="22.29" direction="north" orientation="latitude" precision="555" value="22.29">22.29 ° N</geoCoordinate>
,
<geoCoordinate id="5A48AA0662366B1B5AA3EC059E97813E" degrees="111.22" direction="east" orientation="longitude" precision="555" value="111.22">111.22 ° E</geoCoordinate>
</named-content>
, under rocks in small seepage, coll.
<collectorName id="42EEA7DAC3A174B6798195270B0AC58A">Chao Huang</collectorName>
,
<date id="12E345979A65BBDBEA88E2F178C03634" value="2018-11">
<collectingDate id="2280386249E16C781688919CA531A3F4" value="2018-11">November 2018</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="0128D157C9CA50DC2F8DA75F6EECA6D7" collectingDate="2018-11" collectionCode="SYSBM" collectorName="Chao Huang" country="China" county="Maoming City" latitude="22.29" location="Datianding" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" specimenCode="SYSBM 002145-002147, SYSBM 002148-002151" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="1E3D25F93F7523883D183CFA6724E70D" bold="true" italics="true">
<typeStatus id="582F90A96DB6949F4F7D989380B35104" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCode id="195637AED2DE8198DBBFA678D3999088" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EYXAG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002145002147
</specimenCode>
,
<specimenCount id="F3D74C6B89ED59018937A2F513A94CCA" count="3" type="male">3 males</specimenCount>
(11.6 ×
<quantity id="809234274B67EA8367AF86DC5E38D8F0" metricMagnitude="-3" metricUnit="m" metricValue="9.5" unit="mm" value="9.5">9.5 mm</quantity>
, 10.9 × 9.0 mm, 8.3 ×
<quantity id="4E53AF3113864C9129E3A26723F2E851" metricMagnitude="-3" metricUnit="m" metricValue="6.9" unit="mm" value="6.9">6.9 mm</quantity>
), same data as holotype.
<specimenCode id="71A2D288830EDE5364EF92335B09E9CC" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0E4XAG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002148002151
</specimenCode>
,
<specimenCount id="2B85061E00FAE48583FE8BB94428AFF4" count="4" type="female">4 females</specimenCount>
(16.7 ×
<quantity id="F4FA80D5CFFE1DEE86B0313B47576F08" metricMagnitude="-2" metricUnit="m" metricValue="1.28" unit="mm" value="12.8">12.8 mm</quantity>
, 15.0 ×
<quantity id="1674D0267BFCCBE6FF6B389E6F57C120" metricMagnitude="-2" metricUnit="m" metricValue="1.13" unit="mm" value="11.3">11.3 mm</quantity>
, 14.6 ×
<quantity id="EFEE4C9939DDD700E60133AB2FA124D2" metricMagnitude="-2" metricUnit="m" metricValue="1.14" unit="mm" value="11.4">11.4 mm</quantity>
, 9.3 ×
<quantity id="3D9DB2DF1332B346DC8C5EDAD806A23B" metricMagnitude="-3" metricUnit="m" metricValue="7.5" unit="mm" value="7.5">7.5 mm</quantity>
), same data as
<typeStatus id="D7F6169D81FE001F3CC8E4715EA70F51" type="holotype">holotype</typeStatus>
</materialsCitation>
. •
<materialsCitation id="A3EC77653CB1CF10749E7FDE2DA121DC" collectingDate="2018-11" collectionCode="MW" collectorName="Chao Huang" country="China" county="Maoming City" latitude="22.29" location="Datianding" longLatPrecision="756" longitude="111.22" municipality="Dawuling Nature Reserve" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="paratype">
<named-content id="A04638A3EC2D56FA580D09A3C60D60FD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/nanjing-normal-university" xlink_title="Jiangsu Key Laboratory for Biodiversity and Biotechnology, College of Life Sciences, Nanjing Normal University, Nanjing, China">NNU</named-content>
16 C- 201811
<collectionCode id="11F432FFE42401305194B505D064AE13" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
,
<specimenCount id="5501BB49A3448596738022F2D3251A4F" count="1" type="male">1 male</specimenCount>
(10.3 ×
<quantity id="2610D7F884D0FED8A60C9C31E243B40B" metricMagnitude="-3" metricUnit="m" metricValue="8.3" unit="mm" value="8.3">8.3 mm</quantity>
),
<specimenCount id="4E8FC67873A8C337E5417090ABA87E67" count="1" type="female">1 female</specimenCount>
(14.6 ×
<quantity id="C503F6ACCF7E65C62A4048B775B06086" metricMagnitude="-2" metricUnit="m" metricValue="1.14" unit="mm" value="11.4">11.4 mm</quantity>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHYAG" type="diagnosis">
<paragraph id="E3713EBFFCA63EEEFE54351DF69E64D4">
<heading id="E1B498914116C2616DF6A8A28FCDCC27" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="100B60DE31CE4A7EFD0277DC9CCF3F8F">
Carapace broader than long; dorsal surface slightly convex, postorbital, epigastric cristae weak, rugose, almost confluent (Fig.
<figureCitation id="09FEF603492A3EF25AF0D0083B856DF0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Maxilliped 3 merus width ~ 1.2 × length; ischium width ~ 0.7 × length; exopod reaching slightly beyond anterior edge of ischium, without flagellum (Fig.
<figureCitation id="7C3FE18BEFE73BFAE3B0A4B707C21548" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 A</figureCitation>
). Ambulatory legs with dense setae; pereiopod dactylus shorter than propodus (Fig.
<figureCitation id="D0A6EA11B11DD5D9BDFBEE8764A22BD6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7</figureCitation>
). Male anterior thoracic sternum very broad, width ~ 2.0 × length (Fig.
<figureCitation id="FBCA1520D2FE467C472F016572FBDC63" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 B</figureCitation>
). Male pleon large, broadly triangular, pleonite 6 width ~ 2.6 × length; telson width ~ 1.8 × length (Fig.
<figureCitation id="8126FD7975E47044998015BDCF47A05A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 C</figureCitation>
). Female pleon subovate (Fig.
<figureCitation id="057733469B4D53C66EE7EB0D359023F9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 E</figureCitation>
). G 1 large, strongly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="84D16B885E02AB06F00567C4DBFEB751" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="E1391D699B47C2E190082A59C517972C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 D</figureCitation>
); subterminal segment length ~ 2.3 × length of terminal segment (Fig.
<figureCitation id="46791A10BA2C20302322A64555722023" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
). Subterminal segment outer margin strongly concave; terminal segment short, rounded, distoventrally with long setae, directed inwards, inner-proximal margin concave; tip presenting as protrusion on higher two-thirds of outer margin (Figs
<figureCitation id="AE7CEA256D9AACBD043639679FFDED8C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
,
<figureCitation id="860D03A588C245AB5E8103E8DE71F79C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 A, B</figureCitation>
). G 2 subterminal segment slightly bent outwards distally, flagelliform terminal segment thick, ~ 1.7 × length of subterminal segment, apex blunt, slightly swollen (Fig.
<figureCitation id="6805FC8263216A7D55C327A38EA912E3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 B</figureCitation>
).
</paragraph>
<caption id="E1B0245E580557AEE5BC3FD4770185AD" ID-DOI="10.3897/zookeys.1244.148112.figure5" ID-arpha="0C2D3A26-331B-5A76-9804-0C11FE932E26" httpUri="https://binary.pensoft.net/fig/1366342" startId="F5">
<paragraph id="CC18E3351C7093DF5490D113C1E1FBCD">
<label id="B75D5E870B2EFF654CF0C88092BA2603">Figure 5.</label>
</paragraph>
<paragraph id="9FFFF476D28A7FE233E117E44A141F71">
Dorsal habitus.
<taxonomicName id="BE044C964429B2102E053928BC4A9058" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="78DE8EE02D6F6956D09BFB031FAA204F" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="452735B9FB10BA4D1C12C1D675D9053B" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (11.7 × 9.7 mm),
<abbrev content-type="institution" id="ABBRID0EM1AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002144 (
<emphasis id="514AA7E5425B3242A95F5988C3E59DC4" bold="true">A</emphasis>
); female paratype (16.7 × 12.8 mm),
<abbrev content-type="institution" id="ABBRID0ET1AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002148 (
<emphasis id="4066F3BABE55C3881364136EA9520822" bold="true">B</emphasis>
).
</paragraph>
</caption>
<caption id="50CB8700FF4E4B6D42D83C3C2487FF47" ID-DOI="10.3897/zookeys.1244.148112.figure6" ID-arpha="87E8755B-0889-52C2-83B3-6AD5054E1549" httpUri="https://binary.pensoft.net/fig/1366343" startId="F6">
<paragraph id="F772AF0608FB924DEAEBFF24FB9CF96D">
<label id="87A227665748134C1414DC3300DFEE90">Figure 6.</label>
</paragraph>
<paragraph id="D81245ECE43A7AE3A8855B5793C4D6AE">
<taxonomicName id="A4CDE3A141D33B17206F4C5E622E3C0C" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="28B1E3285C319B3FC2DD9BFC6BF0DF12" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9CCF5F5CC6FF016270250C22AFF00FE" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (11.7 × 9.7 mm),
<abbrev content-type="institution" id="ABBRID0ES2AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002144 (
<emphasis id="6A5EC0DFC7FBF5B540FD34B1B3B74248" bold="true">A</emphasis>
<emphasis id="42797F2E160FD60C0AC087B3FE282731" bold="true">D</emphasis>
); female paratype (16.7 × 12.8 mm),
<abbrev content-type="institution" id="ABBRID0E22AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002148 (
<emphasis id="6C606A32FF10566C8787A5C67FBF5796" bold="true">E</emphasis>
,
<emphasis id="8D37C2544B91D10F65B666BF3E890686" bold="true">F</emphasis>
). Cephalothorax, anterior view (
<emphasis id="1BD0A2C44692B4829D2B9647467814A1" bold="true">A</emphasis>
); anterior thoracic sternum (
<emphasis id="2CCFAA68C3841F28C2A9B44ACCE3AAE7" bold="true">B</emphasis>
); anterior thoracic sternum and pleon, ventral view (
<emphasis id="EACE9D34CE3EBFC1584E6DBA8D826528" bold="true">C</emphasis>
); sterno-pleonal cavity with G 1 in situ, ventral view (
<emphasis id="20E7534890586E7EFFD78FE2E4B8DC85" bold="true">D</emphasis>
); pleon, ventral view (
<emphasis id="4E886DFA50EAA2A332B769A637F01040" bold="true">E</emphasis>
); vulvae, ventral view (
<emphasis id="5378E5A888B3A411C2A661DFDFADF85C" bold="true">F</emphasis>
).
</paragraph>
</caption>
<caption id="DB0E51FD0A94A6FC1BCB99351F6544D7" ID-DOI="10.3897/zookeys.1244.148112.figure7" ID-arpha="85600192-8AD6-59DE-8876-F189707AB03A" httpUri="https://binary.pensoft.net/fig/1366344" startId="F7">
<paragraph id="6004DED723863C89B3931B54F010DAE2">
<label id="83DCDB1AFE0D5541FD90E80990317189">Figure 7.</label>
</paragraph>
<paragraph id="4BABD5DF0EB54DC5BD426644AD0D39FF">
<taxonomicName id="701775AA864855E13CF9C5A0218097E6" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="4045AC0C033E539B7FB2AF21F82C751A" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="058845C8ECB763CA8D71AF8F68E81820" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (11.7 × 9.7 mm),
<abbrev content-type="institution" id="ABBRID0EI4AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002144 (
<emphasis id="31DA0E617A3823A121E391A28C3F04AC" bold="true">A</emphasis>
<emphasis id="3AB44811FDDD7C10DF1AD0616ADF75A4" bold="true">C</emphasis>
,
<emphasis id="2CF519A66D2FE534A2F5C05EFED98381" bold="true">F</emphasis>
,
<emphasis id="3FA53B65EB10A137F1AADB4F7D843B47" bold="true">G</emphasis>
); male paratype (11.6 × 9.5 mm),
<abbrev content-type="institution" id="ABBRID0EV4AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002145 (
<emphasis id="057111B42B9E0A30CCEAF85FD3F55776" bold="true">D</emphasis>
); male paratype (10.9 × 9.0 mm),
<abbrev content-type="institution" id="ABBRID0E34AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002146 (
<emphasis id="47777A0214A9AF652A072FE4A219F251" bold="true">E</emphasis>
); female paratype (16.7 × 12.8 mm),
<abbrev content-type="institution" id="ABBRID0ED5AG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
00 2148 (
<emphasis id="BCA6EE105B9539138DE4652F11B14F66" bold="true">H</emphasis>
,
<emphasis id="0A3AAB15EFDEC1EA9DCA42BE9C0ACA5D" bold="true">I</emphasis>
). Left maxilliped 3 (
<emphasis id="A7A458DE20390E739EE583C713D49A4D" bold="true">A</emphasis>
); left G 2, pleonal view (
<emphasis id="3FFD5D8B186B441D2398452C9A0DBF81" bold="true">B</emphasis>
); left G 1, ventral view (
<emphasis id="30D5BC8FCCE887AE44FD3BA7DB18D9BA" bold="true">C</emphasis>
<emphasis id="D127F0A8C02BF0A687DE9F8FEF02379A" bold="true">E</emphasis>
); major cheliped (
<emphasis id="48B69B6DF3376605DBCFB9660A8E960D" bold="true">F</emphasis>
,
<emphasis id="C5F2BC5D73AC3CC1795D634CBD1613FE" bold="true">H</emphasis>
); minor cheliped (
<emphasis id="A7301D3F02B8C5CAA53F8DE7972C2420" bold="true">G</emphasis>
,
<emphasis id="2DF9CB02DD38A9449F54EBD40F9F4FD4" bold="true">I</emphasis>
). Scale bars: 1.0 mm
<emphasis id="08C4F8DB4C19E7681DC8F512155D73AB" bold="true">(A</emphasis>
<emphasis id="D32522D68D05CB793F6E0BC6FE46087E" bold="true">E)</emphasis>
, 5.0 mm
<emphasis id="71A667BD55D1A6608B4E81019083DEE9" bold="true">(F</emphasis>
<emphasis id="565C625D3469183153C22A4602859E4E" bold="true">I)</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EJ6AG" type="description">
<paragraph id="89A6B2552C293999B3457C5EA74CEA7B">
<heading id="1430FCE9CC038123A1AEACF276D7AA14" reason="title">Description.</heading>
</paragraph>
<paragraph id="4C769362D8B11A199B6310CF91DDB316">
Carapace broader than long, ~ 1.2 × as wide as long in males (
<emphasis id="2C97FAED602DCB91800D2856302C4548" italics="true">n</emphasis>
= 4), ~ 1.3 × as wide in mature females (
<emphasis id="C1D9AE0A0F0542C3932BBF13757BA3CF" italics="true">n</emphasis>
= 4); regions not pronounced, dorsal surface convex; surface finely pitted, anterolateral regions slightly rugose (Fig.
<figureCitation id="DD403C507644345C8E8AB7B29355CAF6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Frontal margin slightly sinuous, deflexed (Fig.
<figureCitation id="E40568BEEE7D7944A17AA66BBF8ADFC2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Epigastric cristae and postorbital cristae rugose, low, almost confluent; bifurcated shallow groove between epigastric cristae (Fig.
<figureCitation id="0FA2C5A08399C666E5578DF6B207FD9D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Branchial regions not swollen (Fig.
<figureCitation id="0958A26BB7B9DF369595EAA870774C77" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Cervical groove shallow (Fig.
<figureCitation id="0DCADAB0C29DB2B1EEE03CD73E9A8469" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Mesogastric region flat (Fig.
<figureCitation id="D4B85EFF706D49192C2593CF68C90502" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). External orbital angle broadly triangular, outer margin slightly convex, confluent with anterolateral margin (Figs
<figureCitation id="985D1E8E157AE162785B11CECACC4F91" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
,
<figureCitation id="D24D3B50A6CE7871D3833F6DEEACC6ED" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Epibranchial tooth granular, indistinct (Figs
<figureCitation id="F3BBA7A089E4A41411778FCA7E353A46" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
,
<figureCitation id="659BF075C19F7A2929D5147B83313E70" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Anterolateral margin lined with 1014 granules; posterolateral margin posteriorly convergent (Fig.
<figureCitation id="50E52F31D279DA571BA7FC2CF4211948" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
); posterolateral surface smooth (Fig.
<figureCitation id="1CCC79CF99687A0F72230A89AD16DFE3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Orbits regular; supraorbital margins weakly cristate, infraorbital margins lined with fused granules (Fig.
<figureCitation id="AF09A8BB08C3AA0A0D369129BD365828" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Eyes normal (Figs
<figureCitation id="74794F3CCCDE54A08CCEBC40031DE1B7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
,
<figureCitation id="BE17C8EC93B4B1C3161F3305B2EDF0EB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Sub-orbital, pterygostomial and sub-hepatic regions generally smooth, pitted (Fig.
<figureCitation id="3047699E62723527555012FDD98EF7BF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Antennules large, folded within broad fossae; antennae very short (Fig.
<figureCitation id="C3BED4108228C8DB62952B9384DD86E4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Median lobe of epistome buccal margin triangular, lateral margins straight (Fig.
<figureCitation id="7D2829ED87098CF2804EF3903690D757" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
).
</paragraph>
<paragraph id="A46CB42AB96EFA2494D9CD05E72EA0DC">
Maxilliped 3 merus subtrapezoidal, with slight median depression, width ~ 1.2 × length; ischium subtrapezoidal with shallow median sulcus, distomesial margin rounded, width ~ 0.7 × length. Exopod reaching proximal one-third of merus; flagellum absent (Fig.
<figureCitation id="6D5ECD3CC26650F4FA4DBFACC4C1E109" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 A</figureCitation>
).
</paragraph>
<paragraph id="9B5A83D4172DF7136C56A15190B2E6EA">
Chelipeds (pereiopod 1) subequal (Fig.
<figureCitation id="715436C6EB28A5C44C3913E9386288E4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 F I</figureCitation>
). Merus trigonal in cross section, surfaces generally smooth; outer dorsal margin slightly crenulated, inner and ventral margin lined with large granules (Figs
<figureCitation id="D818E3F402834CA42F02CBD43126CB1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
,
<figureCitation id="BE4C6CF428642E3A6010019E214E8718" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 A</figureCitation>
). Carpus dorsal surface slightly rugose, with small blunt spine at inner-distal angle, spinule at base (Fig.
<figureCitation id="4EF542C0496DB213E77CC1CAC98797F4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Major cheliped palm length ~ 1.31.4 × height in males (
<emphasis id="1BAEA7AB65FB5ECC942FD3C3B1B49DCF" italics="true">n</emphasis>
= 2), 1.41.5 × height in females (
<emphasis id="875229C2C2BD125E03E6B74A72974237" italics="true">n</emphasis>
= 4); dactylus 0.9 × palm length in males (
<emphasis id="0FF0C7E8471C57B1E9C0338094A21742" italics="true">n</emphasis>
= 2), 0.91.0 × palm length in females (
<emphasis id="FF189C8E8EBAC802ADEA8E8AC5648484" italics="true">n</emphasis>
= 4) (Fig.
<figureCitation id="FF3F645DA606CA9D87C4063A87B77AD8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 F I</figureCitation>
). Palm surface pitted, occlusal margin of fingers with 811 irregular blunt teeth, with very small gape when closed (Fig.
<figureCitation id="519F17CF3C3B113B6F4E17D02542B393" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 F I</figureCitation>
).
</paragraph>
<paragraph id="CE8109367AC2D6012FD8ED35D2CBAACE">
Ambulatory legs slender (pereiopods 25) covered with setae, especially dense on pereiopods 2, 3 (Fig.
<figureCitation id="A6DB142B77CCB42F82636F6E2B328EF7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Pereiopod 3 merus 0.60.7 ×
<abbrev id="ABBRID0EJDBG" xlink_title="Carapace length">CL</abbrev>
in both sexes (
<emphasis id="C520ACC94E2486E3F146CAA357404A9A" italics="true">n</emphasis>
= 8, Fig.
<figureCitation id="8F7EA0D44B0939C14E22C559B8FBF1E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
). Pereiopod 5 propodus length 1.92.2 × height in males (
<emphasis id="8F475EEAE6ECDB2DA6316942D54E129B" italics="true">n</emphasis>
= 4), 2.12.4 × height in females (
<emphasis id="F585BB540A7964B89D060E13D6BAA638" italics="true">n</emphasis>
= 4), shorter than dactylus (Fig.
<figureCitation id="D53A367CD2CE80BAC27B14F2B5FF389D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
).
</paragraph>
<paragraph id="C58AD4C5377E255BA9EC02F1520D78D4">
Male thoracic sternum generally smooth, sparsely pitted; sternites 14 width ~ 2.0 × length; sternites 1, 2 fused to form broad triangle; fused sternites 1, 2 demarcated from sternite 3 by almost straight transverse sulcus; sternites 3, 4 fused without obvious demarcation (Fig.
<figureCitation id="E3322EC65A39826B826A01BC652D8674" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 B</figureCitation>
). Male sterno-pleonal cavity reaching anteriorly slightly beyond mid-length of cheliped coxa (Fig.
<figureCitation id="892821742D42709D9301B132004EA97B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 B</figureCitation>
). Male pleonal locking tubercle positioned at mid-length of sternite 5 (Fig.
<figureCitation id="D53E3D2F2A1D620C0E8927AD838AE5E9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 D</figureCitation>
). Female vulvae ovate, large, reaching suture of sternites 5 / 6, relatively widely separated (Fig.
<figureCitation id="86DF0C8E968AC1E1ABAFC29AEC36097A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 F</figureCitation>
).
</paragraph>
<paragraph id="8DEC0DF68079301BFB16DCEE53DCB293">
Male pleon large, broadly triangular; somites 36 progressively narrower; somite 6 width ~ 2.6 × length; telson width 1.8 × length; lateral margins almost straight, apex rounded (Fig.
<figureCitation id="B98BEAB4B2947E82B261A3877C864F8B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 C</figureCitation>
). Female pleon sub-ovate (Fig.
<figureCitation id="7E0C5B4544B202183181E1806AD05C09" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 E</figureCitation>
).
</paragraph>
<paragraph id="63CB9CEE411F48495B48635BF6B06B00">
G 1 large, strongly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="D2862FB382B5594C1C177D87C000C8F5" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="8B7F00D32ECCFB2DFF5D2335F9889DC9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 D</figureCitation>
); subterminal segment length ~ 2.3 × length of terminal segment (Fig.
<figureCitation id="BD64F7994DE56C3C7BA443DEB40B33C8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
). Subterminal segment outer margin strongly concave; terminal segment short, rounded, distoventral region with long setae, directed inwards, inner-proximal margin concave; tip presenting as protrusion on higher two-thirds of outer margin (Figs
<figureCitation id="88486972817803F3D6E5B0EF0BDF2CA5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
,
<figureCitation id="7E33B77557D586AED143953D4A382766" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 A, B</figureCitation>
). G 2 subterminal segment slightly bent outwards distally, flagelliform terminal segment thick, ~ 1.7 × length of subterminal segment, apex blunt, slightly swollen (Fig.
<figureCitation id="A96E540CC6AA9F228B019A16E5542202" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENFBG" type="Colour in life">
<paragraph id="603535C274C335E3D4D1A527FA1DEA54">
<heading id="74514D5B94B78B1DE69CF0736F37F279" reason="title">Colour in life.</heading>
</paragraph>
<paragraph id="218046A7DDFFD63911259F748413A616">
Generally drab camouflage-brown all over, some individuals exhibit a reddish hue (Fig.
<figureCitation id="1F17948B30BE00854FE51F49C414E6E5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Colour in life. Megapleonum ferrumequinum sp. nov., male (A); Megapleonum wangjiani sp. nov., male (B); Megapleonum wangjiani sp. nov., female (C); Megapleonum yangdongense sp. nov., male." figureDoi="10.3897/zookeys.1244.148112.figure13" httpUri="https://binary.pensoft.net/fig/1366350">13 B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXFBG" type="habitat">
<paragraph id="7EE8E3EFC90400397145D8455448E01A">
<heading id="734FE48CD3B4575788F457DD0E847D8D" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="101914545F204B3FEF1F5F7FE6D068C3">
Little is known about the ecology of this new species, aside from its occurrence at high elevations, where it inhabits seepages and is occasionally seen roaming the forest floor.
<taxonomicName id="519FBE5BBD8FA2115C83B40931D9F241" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="28B8484065C309BB2FFA4A2CC7CD0376" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6D3702881376D4028726823411773BAC" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4E1A73620A6AE9A36082FD16F5716FDF" baseAuthorityName="Dai &amp; Turkay" baseAuthorityYear="1997" class="Malacostraca" family="Potamidae" genus="Eurusamon" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="C16988F5764EB61BBD84B06193D3E111" italics="true">Eurusamon guangdongense</emphasis>
</taxonomicName>
are also found on the same mountain, but the three species apparently occupy different niches.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETGBG" type="distribution">
<paragraph id="7B58DBC53BCEF1A34924E7C8BFD6BF1E">
<heading id="E70DCB551A7B8856B9B4D1B8D9E69B10" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="99688FBB1F24B646CA80E95D14C81CD3">
Dawuling Nature Reserve, Maoming City,
<collectingRegion id="690E00B30E334B7C8A42E0996878C12D" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="ABCB242BF5DD2798AFB07F2E9E1DDF75" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYGBG" type="etymology">
<paragraph id="3387510FB5C61F35BF3260CF036EE943">
<heading id="0B3FB03C907C4CAF4801C75269EC601F" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9CFAD2893FCA4681BF65EAF7B5BFF98A">This species is named in honour of its discoverer, Jian Wang, in recognition of his contribution to this study.</paragraph>
</subSubSection>
<subSubSection id="SECID0E4GBG" type="remarks">
<paragraph id="A918C74BFD0336BEF154247ED53DFDBC">
<heading id="CDCB2EE0779DDF9C362294D4C709250D" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="568D2724E4401B181C425858E90C8080">
Like many species in this genus,
<taxonomicName id="9EBEF9012376D71320B11C0D8EB540D0" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="3E132B01F298BE0C5A533BB4F49A9FC6" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE89016C2F9B19213C59B357B870EDBB" rank="species">sp. nov.</taxonomicNameLabel>
can immediately be distinguished by its distinctive G 1, especially in the rounded terminal segment with the tip presenting as a protrusion on the higher two-thirds of the outer margin (Figs
<figureCitation id="6C8134A3F73DB0F5E825159BBC200947" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
,
<figureCitation id="DA3A142BE48842E5800034AD8AADCD7A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 A, B</figureCitation>
). Apart from the G 1,
<taxonomicName id="C5E2958A898B91310E4B49C21504F872" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani">
<emphasis id="90AC8A5D40835DD99B223D295678819A" italics="true">M. wangjiani</emphasis>
</taxonomicName>
can be separated from the sympatric
<taxonomicName id="6634AB0C92567AA4E520FE003758A1B2" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="A6227DED5889E249306171F19A616205" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D65F09EADE36391E9BB14B0ECD032B9C" rank="species">sp. nov.</taxonomicNameLabel>
by its slenderer legs, with the pereiopod 5 propodus length 1.92.4 × height, shorter than dactylus, whereas in the latter the pereiopod 5 propodus length is 1.7 × height, longer than dactylus (Fig.
<figureCitation id="873F314C82949F838DC7107E67A96CED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5 A, B</figureCitation>
vs Fig.
<figureCitation id="3C3415FD5F3EF44F44090BFF14B84619" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
).
<taxonomicName id="C9A82AB042BA45B39EC9FA9FECF46D6D" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="812C79230A6D06C4B3F36A2816E85FE8" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7681FF052FF63631E33C3483EE9A7C76" rank="species">sp. nov.</taxonomicNameLabel>
also has a broader male abdomen than
<taxonomicName id="50516D209FBEF79F34E580A03DCCEF53" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="n. sp.">
<emphasis id="B4CE186F075C93142DE788AF90A8E5AD" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="687BE8669A5DE00315E8CFE050369F5E" rank="species">n. sp.</taxonomicNameLabel>
(Fig.
<figureCitation id="6778687BD21237F9E3495D143CE136F4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 C</figureCitation>
vs Fig.
<figureCitation id="789B1A644C23A523656343DC2D8039AE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 D</figureCitation>
). More detailed comparisons can be found in Table
<tableCitation id="EEAFD46E066C62A511AAA542E87E1A65" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparisons between the species within the genus Megapleonum." httpUri="http://table.plazi.org/id/69D766E64FF15F0F8BB8C5782AADA37B" tableUuid="69D766E64FF15F0F8BB8C5782AADA37B">1</tableCitation>
.
</paragraph>
<paragraph id="A1D5A3E0B3D0F971AAC7A8608065E06E">
There is noticeable sexual dimorphism in
<taxonomicName id="2719B09C632A388A3803185E779CE277" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="C9FDCCF201328C33537C8A7D5CB15B95" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="519966836A8F75AD79E4AC71DF47CE97" rank="species">sp. nov.</taxonomicNameLabel>
, with males being smaller and maturing at a smaller size than females (the male
<abbrev content-type="institution" id="ABBRID0EDKBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">
<collectionCode id="76A475EF492519A141B97410D2F89DD2">SYSBM</collectionCode>
</abbrev>
002147 at
<abbrev id="ABBRID0EIKBG" xlink_title="Carapace width">CW</abbrev>
<quantity id="5326C08993AAD4CCB5449FBAB8B0F6E9" metricMagnitude="-3" metricUnit="m" metricValue="8.3" unit="mm" value="8.3">8.3 mm</quantity>
has a full length G 1 whereas the female
<abbrev content-type="institution" id="ABBRID0EMKBG" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">
<collectionCode id="E9A8FC55C1852092A0938EAB45DD3F8F">SYSBM</collectionCode>
</abbrev>
002151 at
<abbrev id="ABBRID0ERKBG" xlink_title="Carapace width">CW</abbrev>
<quantity id="3B08D74B57178E55A1A15BC919D0352D" metricMagnitude="-3" metricUnit="m" metricValue="9.3" unit="mm" value="9.3">9.3 mm</quantity>
has an immature narrow abdomen). Females also have a proportionally wider carapace (1.2 × as wide as long in mature males vs ~ 1.3 × in mature females) and more slender legs than females (pereiopod 5 propodus length 1.92.2 × height in males vs 2.12.4 × height in females), but these differences might also be related to size and are only obvious due to the apparent inability for the males to reach the size of females. Interspecific variation of the G 1 is small, with the tip opening varying slightly in size and shape (Fig.
<figureCitation id="8A84A53C8BD108B7AD14A22729DB4550" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 C E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,311 @@
<document id="A23E6DA92B6891746C50FBF8C65E0F9E" ID-DOI="10.3897/jhr.98.156087" ID-ZooBank="A8629F7B-FB5C-459A-BFB2-93EEC00493E6" ID-publisher-id="156087" URI-arpha="0C3B68B5-FD1F-5B6D-B6AC-610BFE7DCD22" URI-zoobank="http://zoobank.org/A8629F7B-FB5C-459A-BFB2-93EEC00493E6" article-type="research-article" checkinTime="1751465250129" checkinUser="pensoft" docAuthor="Pérez-Gómez, Álvaro, Sánchez-García, Íñigo, Royo, José Manuel, Rasplus, Jean-Yves &amp; Robla, Jairo" docDate="2025" docId="FB48926A020C5819B0E41C6C4F81D913" docLanguage="en" docName="Journal of Hymenoptera Research 98: 667-687" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/156087/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eupristina verticillata Waterston 1921" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="687" masterDocId="0C3B68B5FD1F5B6DB6AC610BFE7DCD22" masterDocTitle="Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe" masterLastPageNumber="687" masterPageNumber="667" pageNumber="667" updateTime="1751465673923" updateUser="ExternalLinkService">
<mods:mods id="2A25A2CE96DFA7334CED3EE0BD42AD08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="74A25883BE4E3F8CC29C8CBE79A3771E">
<mods:title id="E5E6AE4FD9B454FF66A9529D42D5E87B">Unveiling an intricate relationship: Ficus trees, their associated wasps (Hymenoptera, Chalcidoidea) and another story of invasion in the Iberian Peninsula and Europe</mods:title>
</mods:titleInfo>
<mods:name id="780751D2F34A4DF2DAE1E90A12933611" type="personal">
<mods:role id="8E681ADED2717FBC3F9FE277BCBF046A">
<mods:roleTerm id="94D429AE96252010B067E02763AF4AB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1497515268028D539377FF55E11DECE5">Pérez-Gómez, Álvaro</mods:namePart>
<mods:nameIdentifier id="300B01A6012C81F2F2ECAD88E04EB53B" type="ORCID">0000-0002-6752-0348</mods:nameIdentifier>
<mods:affiliation id="436A357BD2B499B28855D85BC2041D70">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:name id="BFD622BC83408B8D9DE9FA2C0745C613" type="personal">
<mods:role id="33E68BB31C109DAD6AA87A6DEC1292AE">
<mods:roleTerm id="DB3799C3AFB12479192B35910F99202B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FCBE1B397CF050D83ADDC5CF837CB235">Sánchez-García, Íñigo</mods:namePart>
<mods:nameIdentifier id="D5DFD70490D82D2103E053396032C3B5" type="ORCID">0000-0003-4280-9206</mods:nameIdentifier>
<mods:affiliation id="CE976A6EB2DD6B4B4DF546D708A61DA1">Zoobotánico de Jerez, Jerez de la Frontera, Spain</mods:affiliation>
</mods:name>
<mods:name id="3922CE40F67292650AFA39CA9B92B6B5" type="personal">
<mods:role id="EDF731D4A5AF2C628202DEB1AD3DA5D6">
<mods:roleTerm id="AB12540258C6D0B66BFE70BC48932036">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3BCDE2F4F6704B6DC785693D62D1D372">Royo, José Manuel</mods:namePart>
<mods:nameIdentifier id="14C7B547098AAB9A0536D99C6123EB42" type="ORCID">https://orcid.org/0009-0005-9759-7767</mods:nameIdentifier>
<mods:affiliation id="3DA8E00A5089B75700B2CCF80BFBB97E">Department of Environmental Sciences and Natural Resources, University of Alicante, Alicante, Spain</mods:affiliation>
</mods:name>
<mods:name id="18B8C677AB8E1EB2144481E450094683" type="personal">
<mods:role id="D68B4188962ACA3E7797220079000746">
<mods:roleTerm id="2A4DE6B32A2ACD28041FDA6826AD9599">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2167467540FCD6006C23DD2C18FA8E70">Rasplus, Jean-Yves</mods:namePart>
<mods:nameIdentifier id="D784B47CA973749BEB6CE15613E5A92D" type="ORCID">0000-0001-8614-6665</mods:nameIdentifier>
<mods:affiliation id="1E39CC9490162EBF7B1C2C318889B240">CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, Université de Montpellier, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="15C770A5282E28789C3E0BC864C052A9" type="personal">
<mods:role id="22CD6D685C371852DEEEEFE266FF7513">
<mods:roleTerm id="A2D94CA2CC8B5AC96643FBEB9860A8B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="40231D07AB403BB2CB6001851B1B190C">Robla, Jairo</mods:namePart>
<mods:nameIdentifier id="CAF5212114A9F777EC4250C14FCB914F" type="ORCID">0000-0001-8866-1740</mods:nameIdentifier>
<mods:affiliation id="0D7B67C55BF589556535B035704188F7">Estación Biológica de Doñana (EBD-CSIC), Americo Vespucio 26, 41092. Sevilla, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource id="25064A0563D01705AE121C6922CC66C6">text</mods:typeOfResource>
<mods:relatedItem id="89FEABAD8303AD8116A6A35A3534FE63" type="host">
<mods:titleInfo id="1608A4A6A57E0DBB27E4CE06A2CDCC28">
<mods:title id="0618EAD852477B9041C78F3D455D99E4">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="2C8CED6EFF2FDC3FA7FD9C3914C1069D">
<mods:date id="4568138AA35BF52A91CE68A6A58DE29C">2025</mods:date>
<mods:detail id="A561F9C9C9ED46B23BCBBD29AB077054" type="pubDate">
<mods:number id="4A59DB327E68D2736C3E04909D4761E2">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="AA5F8B674C9F172F5565F9274857DA36" type="volume">
<mods:number id="7C6889333639B02A42F550F8F591D031">98</mods:number>
</mods:detail>
<mods:extent id="1C57B9D1F02550CB0A6A64DD14B1F142" unit="page">
<mods:start id="19FD4E84F29348C1C82293B5C63E8840">667</mods:start>
<mods:end id="F9282F58E62F58030D49501A6D50FCA4">687</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="26F60729C1D35D6445185C6F7ACD354E">journal article</mods:classification>
<mods:identifier id="AEF6B64E7BDC559EC1E6B62BF8A6897B" type="DOI">10.3897/jhr.98.156087</mods:identifier>
<mods:identifier id="48D182768ACF33F9950D16A2F7BBF1A5" type="ZooBank">A8629F7B-FB5C-459A-BFB2-93EEC00493E6</mods:identifier>
</mods:mods>
<treatment id="FB48926A020C5819B0E41C6C4F81D913" ID-DOI="http://doi.org/10.5281/zenodo.15791845" ID-Zenodo-Dep="15791845" ID-arpha="FB48926A-020C-5819-B0E4-1C6C4F81D913" LSID="urn:lsid:plazi:treatment:FB48926A020C5819B0E41C6C4F81D913" httpUri="http://treatment.plazi.org/id/FB48926A020C5819B0E41C6C4F81D913">
<subSubSection id="2914F24138F83B749A08F4ACCC30EBD8" type="nomenclature">
<paragraph id="ABACCBAC317F6757BA8F6630148F66E1">
<taxonomicName id="DB2640EEA123B34A81466AC35A74DCFC" ID-CoL="3CXGD" authority="Waterston, 1921" authorityName="Waterston" authorityYear="1921" class="Insecta" family="Agaonidae" genus="Eupristina" isUncertain="true" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="verticillata">
<emphasis id="DDB702527DD59F49DC732E8A2EBCDAE5" italics="true">Eupristina</emphasis>
aff.
<emphasis id="48ABE18ED8A51C0E99F79402FC4BAB3C" italics="true">verticillata</emphasis>
<bibRefCitation id="54AA052B5DB2BCB7211BA08D02055323" DOI="10.1017/S0007485300044837" author="Waterston J" issue="1" journalOrPublisher="Bulletin of Entomological Research" pagination="35-40" refId="B97" refString="Waterston J (1921) On some Borean fig-insects (Agaonidae - Hymenoptera Chalcidoidea). Bulletin of Entomological Research 12 (1): 35-40. https://doi.org/10.1017/S0007485300044837" title="On some Borean fig-insects (Agaonidae - Hymenoptera Chalcidoidea)." volume="12" year="1921">Waterston, 1921</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGKAE" type="material">
<paragraph id="942AED5F5F368EA1044ED43C9D11B328">
<heading id="EEF9271B9CA36155454342A55E61254B" reason="title">Material studied.</heading>
</paragraph>
<paragraph id="1506EFD323352A530AE25CB2B65AA370">
<materialsCitation id="A24A8A291008594BA7E6C0A5E857ABAC" collectingDate="2018-12-07" collectionCode="MNCN" collectorName="Jerez de la Frontera &amp; de Jerez &amp; I. Sanchez &amp; Ent" country="Spain" elevation="82" latitude="36.689316" location="Zoobotanico de Jerez" longLatPrecision="1" longitude="-6.150638" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Cadiz">
<emphasis id="C2F49CCDDBF613B5DA39423B53F65F11" bold="true">
<collectingCountry id="F7C2016FCB5364ACD26972A454404E47" name="Spain">Spain</collectingCountry>
</emphasis>
: •
<emphasis id="183D36CB20793CB1557CC1551A6F965F" bold="true">
<collectingRegion id="0CCF2B125D0E09633E53321B2D319275">Cádiz</collectingRegion>
</emphasis>
:
<collectorName id="0F5FD30EE5BC55AD42ABE5EA35D4ADC8">Jerez de la Frontera</collectorName>
(
<location id="79F9BE76E47A31F0B30FE79081A0017F" LSID="urn:lsid:plazi:treatment:FB48926A020C5819B0E41C6C4F81D913:79F9BE76E47A31F0B30FE79081A0017F" country="Spain" latitude="36.689316" longLatPrecision="1" longitude="-6.150638" name="Zoobotanico de Jerez" stateProvince="Cadiz">
Zoobotánico
<collectorName id="7709FBDA9FDB822BCE874AE7AB9495C6">de Jerez</collectorName>
</location>
):
<specimenCount id="9FEA544625204451EAE5A9C24AE81C3E" count="2" type="male">2 ♂</specimenCount>
+
<specimenCount id="B4F01BA6FDB513AA886D1891C9EEF81A" count="2" type="female">2 ♀</specimenCount>
<abbrev id="ABBRID0EQKAE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="0458ED63401D8F038A0DF3BB3E045C97">Í. Sánchez</collectorName>
<abbrev id="ABBRID0EUKAE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="48E3243B69D9D39EDD504C7B29EF6766" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
_
<collectorName id="C0FFDA5088A03C8E362BC3350C2E8117">Ent</collectorName>
235062
<abbrev id="ABBRID0EYKAE" xlink_title="collection of deposit">deposit</abbrev>
.,
<date id="3DFC2B3DD1A6E4BDD65865B090ECCAA2" value="2018-12-07">
<collectingDate id="F5222DD4D0F80FDE65A5583586616983" value="2018-12-07">7 - XII- 2018</collectingDate>
</date>
,
<quantity id="3F0AED692D4CCA4181AF98E6CDDA270D" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">
<elevation id="6EA86D0C1B9807E2AE3DFB317BF52914" metricMagnitude="1" metricUnit="m" metricValue="8.2" unit="m" value="82.0">82 m</elevation>
</quantity>
a. s. l, found in dissected syconia of
<taxonomicName id="8646761F2783971651F2D7C174FACE51" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="E0A201217E32B851DFE584CB3BE21A54" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKLAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-6.150638,36.689315]}">
<geoCoordinate id="53B6BE24B81E04A51D57C334DA2702F4" degrees="36.689315" direction="north" orientation="latitude" precision="1" value="36.689316">36.689315</geoCoordinate>
,
<geoCoordinate id="8BE54A7D462CF6F3016F543A41CBA219" degrees="6.150638" direction="west" orientation="longitude" precision="1" value="-6.150638">- 6.150638</geoCoordinate>
</named-content>
)
</materialsCitation>
. •
<materialsCitation id="1CB67FAAB1B4126B1CDA9D700CE2C64C" collectingDate="2023-07-05" collectionCode="AC" collectorName="de Alicante &amp; J. M. Royo" country="Spain" county="San Vicente del Raspeig" elevation="92" latitude="38.384266" location="See in Fig." longLatPrecision="1" longitude="-0.512455" municipality="Universidad de Alicante" specimenCount="17" specimenCount-female="10" specimenCount-male="7" stateProvince="Alicante">
<emphasis id="DB5B94455820887943CB6763EA3CCF12" bold="true">
<collectingRegion id="8F468D3579B817682ADBF143DE405200">Alicante</collectingRegion>
</emphasis>
:
<collectingCounty id="15C077CAF8C4A8D450BA8C5F54355D18">San Vicente del Raspeig</collectingCounty>
(
<collectingMunicipality id="CB870D62342DFB4AB82874C9ABDA041E">
Universidad
<collectorName id="CB534B3466DC7589177226ECEC03D951">de Alicante</collectorName>
</collectingMunicipality>
):
<specimenCount id="E80F0BBD596F22450FCE0C03D61C8A4A" count="10" type="female">10 ♀</specimenCount>
+
<specimenCount id="2D82D6C5A8C7A060702D6C55C9BFB511" count="7" type="male">7 ♂</specimenCount>
<abbrev id="ABBRID0ERLAE" xlink_title="specimen">ex</abbrev>
.,
<collectorName id="FE2D026BE418D377B8A84C8DBFE189E5">J. M. Royo</collectorName>
<abbrev id="ABBRID0EVLAE" xlink_title="collector">leg</abbrev>
. and
<collectionCode id="2F4E8FBBF7C15EFD7167A05BE6F73B75" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" type="University or college">AC</collectionCode>
<abbrev id="ABBRID0EZLAE" xlink_title="collection of deposit">deposit</abbrev>
.
<date id="2EC2818648A62449CE116623A2D4D88A" value="2023-07-05">
<collectingDate id="08036FCD362908E923BC8AECEC1F7CF2" value="2023-07-05">5 - VII- 2023</collectingDate>
</date>
,
<quantity id="BCA527C3427C9237FF746E68515A2796" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">
<elevation id="D1512D1706F71F8F561319A2A1BB3985" metricMagnitude="1" metricUnit="m" metricValue="9.2" unit="m" value="92.0">92 m</elevation>
</quantity>
a. s. l, found dissecting fallen syconia of
<taxonomicName id="58DC6B45F89667870B3B95339E2DCF05" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="985F1B7EABFFBD6835A31BF64337E788" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
. This day, several specimens were seen flying close to the tree (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELMAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-0.512455,38.384264]}">
<geoCoordinate id="B9499FACFE3EC6AE1E1D06DD12EB2780" degrees="38.384264" direction="north" orientation="latitude" precision="1" value="38.384266">38.384264</geoCoordinate>
,
<geoCoordinate id="CBB4B839E9AA0985319B207581DA0A5F" degrees="0.512455" direction="west" orientation="longitude" precision="1" value="-0.512455">- 0.512455</geoCoordinate>
</named-content>
).
<location id="C34070C5368B8C1F8545D2795BDB7BE9" LSID="urn:lsid:plazi:treatment:FB48926A020C5819B0E41C6C4F81D913:C34070C5368B8C1F8545D2795BDB7BE9" country="Spain" county="San Vicente del Raspeig" latitude="38.384266" longLatPrecision="1" longitude="-0.512455" municipality="Universidad de Alicante" name="See in Fig." stateProvince="Alicante">See in Fig.</location>
<figureCitation id="298FFF5D65DF0F49A64102EE5A900E48" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wasps cited in this work A female of Eupristina aff. verticillata Waterston, 1921 B Walkerella microcarpae Bouček, 1993 C female and D male of Pleistodontes imperialis Saunders, 1883 E female and F males of Philotrypesis okinavensis Ishii, 1934 G Eufroggattisca okinavensis (Ishii, 1934) H Josephiella microcarpae Beardsley &amp; Rasplus, 2001." figureDoi="10.3897/jhr.98.156087.figure1" httpUri="https://binary.pensoft.net/fig/1366113">1 A</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMAE" type="Ecological remarks">
<paragraph id="A0A4D6F12BDDC11501F60084F20A1551">
<heading id="9228481F26DC445F42628D64CDCD02EA" reason="title">Ecological remarks.</heading>
</paragraph>
<paragraph id="08148287924EB35B304C6A5BFF0633CE">
Pollinator of
<taxonomicName id="1A836D4BB853926999E6AF05AF8E0D94" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="E680888F30A21659FA35496D90F500D7" italics="true">Ficus microcarpa</emphasis>
</taxonomicName>
. Also mentioned to be associated with
<taxonomicName id="D47E73534AFF47CF249BAEA0083D7B01" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="benjamina">
<emphasis id="707EEB7C8ED4F403FCC51C59FF47748B" italics="true">Ficus benjamina</emphasis>
</taxonomicName>
L. but without additional data (
<bibRefCitation id="57BE87E9390B6026940C50EFFD0D10D1" author="Falco-Gari" etAl="et al." firstAuthor="Falco-Gari" refId="B32" refString="Falcó-Garí JV, Oltra-Moscardo MT, Peris-Felipo FJ, Jiménez-Peydro R (2010) Exotic fig pollinators and associates (Chalcidoidea: Agaonidae) of Ficus trees in Europe. Seventh international Congress of Hymenopterists. Hungary, Programme and Abstracts." year="2010">Falcó-Garí et al. 2010</bibRefCitation>
), probably misidentified.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUNAE" type="distribution">
<paragraph id="B4C1B4A86002FCFD7E28C8C0B29BB5D2">
<heading id="FBCE4B0899E94E13CB82DF73EFB06FBA" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="D8DF7BEB40BAA819D40EB1CAE4C59017">
This pollinator was described from
<collectingCountry id="242D7AAA75C3920C5A958640A22B424E" name="Malaysia">Malaysia</collectingCountry>
(
<bibRefCitation id="E16136B8BAF2E3DFE4917614461BF467" DOI="10.1017/S0007485300044837" author="Waterston" firstAuthor="Waterston" issue="1" journalOrPublisher="Bulletin of Entomological Research" pagination="35-40" refId="B97" refString="Waterston J (1921) On some Borean fig-insects (Agaonidae - Hymenoptera Chalcidoidea). Bulletin of Entomological Research 12 (1): 3540. https://doi.org/10.1017/S0007485300044837" title="On some Borean fig-insects (Agaonidae - Hymenoptera Chalcidoidea)." volume="12" year="1921">Waterston 1921</bibRefCitation>
) and have been subsequently cited from
<collectingCountry id="A3A3E52303BE9BA7E139ED547E6ABD29" name="Indonesia">Indonesia</collectingCountry>
and
<collectingCountry id="9F8C7ACB95FFCAEF98E69DDBED029E19" name="Philippines">Philippines</collectingCountry>
(
<bibRefCitation id="900F7555769ADC341A80DF59699E5E0A" DOI="10.14203/treubia.v8i3-4.1752" author="Grandi" firstAuthor="Grandi" journalOrPublisher="Jacobson Descriptions préliminaires Treubia" pagination="352-364" refId="B37" refString="Grandi G (1926) Hyménoptères sycophiles récoltés à Sumatra et à Java par E. Jacobson Descriptions préliminaires Treubia 8: 352364. https://doi.org/10.14203/treubia.v8i3-4.1752" title="Hyménoptères sycophiles récoltés à Sumatra et à Java par E." volume="8" year="1926">Grandi 1926</bibRefCitation>
), although these last records may be based on misidentification. The species is supposed to be native in
<collectingCountry id="31B96C6826FC882DD932C3EE67801107" name="China">China</collectingCountry>
or
<collectingCountry id="F439537D02975191242E47B70EDE650A" name="Japan">Japan</collectingCountry>
(
<bibRefCitation id="3FA95DFC0E452F1778875D3508FCC2FF" author="Grandi G" journalOrPublisher="Bollettino della Società entomologica italiana" pagination="18-24" refId="B38" refString="Grandi G (1927 b) Una nuova specie di Blastophaga del Giappone. Bollettino della Società entomologica italiana 59: 1824." title="Una nuova specie di Blastophaga del Giappone." volume="59">Grandi 1927 b</bibRefCitation>
;
<bibRefCitation id="51832ED29F48D1D0CD550849B6569ABD" author="Ishii" firstAuthor="Ishii" journalOrPublisher="Kontyû" pagination="84-100" refId="B47" refString="Ishii T (1934) Fig. chalcidoids of Japan. Kontyû 8: 84100." title="Fig. chalcidoids of Japan." volume="8" year="1934">Ishii 1934</bibRefCitation>
;
<bibRefCitation id="D9B3CEC9BDD2BFDBF0DC5B0B0098E087" author="Corner" firstAuthor="Corner" journalOrPublisher="The Gardens bulletin, Singapore" pagination="1-186" refId="B22" refString="Corner EJH (1965) Check-list of Ficus in Asia and Australasia with keys to identification. The Gardens bulletin, Singapore 21: 1186." title="Check-list of Ficus in Asia and Australasia with keys to identification." volume="21" year="1965">Corner 1965</bibRefCitation>
) and has been introduced in numerous countries with its host plants (
<bibRefCitation id="D19978F7FDA3EA8411E4C8299F26BFCC" author="van Noort and Rasplus" firstAuthor="van Noort" refId="B91" refString="van Noort S, Rasplus J-Y (2024) Figweb: figs and fig wasps of the world. www. figweb. org [Accessed on 1 May 2024]" url="www. figweb. org" year="2024">van Noort and Rasplus 2024</bibRefCitation>
).
<bibRefCitation id="AEE17921A92228A8A0495411E1D7878C" author="Falcó-Garí JV &amp; Oltra-Moscardo MT &amp; Peris-Felipo FJ &amp; Jiménez-Peydro R" etAl="et al." firstAuthor="Falco-Gari" refId="B32" refString="Falcó-Garí JV, Oltra-Moscardo MT, Peris-Felipo FJ, Jiménez-Peydro R (2010) Exotic fig pollinators and associates (Chalcidoidea: Agaonidae) of Ficus trees in Europe. Seventh international Congress of Hymenopterists. Hungary, Programme and Abstracts." year="2010">Falcó-Garí et al. (2010)</bibRefCitation>
has not provided specific data for the eastern localities of
<collectingCountry id="4EDF986B09966811759A2A8B586F2033" name="Spain">Spain</collectingCountry>
. Thus, we can consider this the first confirmed record for the Iberian Peninsula. Furthermore, this species was cited from continental Europe in
<collectingCountry id="9FD804254A82B66C3B71B5A5B887D46C" name="Italy">Italy</collectingCountry>
(
<bibRefCitation id="2E0E89E02B62BE2C561EE8CE9843823C" author="Lo Verde" etAl="et al." firstAuthor="Lo Verde" refId="B64" refString="Lo Verde G, Porcelli F, Bella S, Rasplus J-Y (2007) Imenotteri Agonidi nuovi per lEuropa e loro ruolo nella naturalizzazione di Ficus spp. in Italia. Atti XXI CNIE, Campobasso, 60." year="2007">Lo Verde et al. 2007</bibRefCitation>
) and
<collectingCountry id="C0A72B4976464F77C958A64AC125D921" name="Greece">Greece</collectingCountry>
(
<bibRefCitation id="855CE5E3593574AEDB3D92F93256ACFF" DOI="10.3897/neobiota.98.139650" author="Koutsoukos" etAl="et al." firstAuthor="Koutsoukos" journalOrPublisher="NeoBiota" pagination="93-115" refId="B60" refString="Koutsoukos E, Konstantopoulou A, Parmakelis A, Demetriou J, Compton SG, Georgiadis C, Constantinidis T (2025) The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece. NeoBiota 98: 93115. https://doi.org/10.3897/neobiota.98.139650" title="The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece." volume="98" year="2025">Koutsoukos et al. 2025</bibRefCitation>
). Moreover, it was cited from several islands such as
<collectingRegion id="5424217072E1B0A2F9A92DE1A82CF0EE" country="Italy" name="Sicilia">Sicily</collectingRegion>
(
<collectingCountry id="E5680160CC3A181AD42928B9B8A4FFF3" name="Italy">Italy</collectingCountry>
) (
<bibRefCitation id="2259D789BAA0645C98C012774DB9D418" author="Lo Verde" etAl="et al." firstAuthor="Lo Verde" refId="B63" refString="Lo Verde G, Porcelli F, Sinacori A (1991) Presenza di Parapristina verticillata (Waterst.) e Odontofroggatia galili Wiebes (Hymenoptera: Chalcidoidea Agaonidae) in Sicilia. Atti XVI CNIE, Bari-Martina Franca (TA), 139143." year="1991">Lo Verde et al. 1991</bibRefCitation>
),
<collectingCountry id="EC39117CA7ACA0313C96A7B0D9A3F6F1" name="Cyprus">Cyprus</collectingCountry>
(
<bibRefCitation id="B59AB99CC9EC17A497EB0B91EE127263" DOI="10.3391/bir.2023.12.2.20" author="Demetriou" etAl="et al." firstAuthor="Demetriou" issue="2" journalOrPublisher="BioInvasions Records" pagination="573-580" refId="B30" refString="Demetriou J, Koutsoukos E, Mavrovounioti N, Radea K, Arianoutsou M, Roy EH, Compton SG, Martinou A (2023) A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus. BioInvasions Records 12 (2): 573580. https://doi.org/10.3391/bir.2023.12.2.20" title="A rather unfruitful relationship? Fig. wasp diversity (Hymenoptera: Chalcidoidea) of the alien invasive Ficus microcarpa in Cyprus." volume="12" year="2023">Demetriou et al. 2023</bibRefCitation>
), Madeira (
<bibRefCitation id="A4277C827995602D51B412AEEDC2E168" author="Koponen and Askew" firstAuthor="Koponen" journalOrPublisher="Vieraea" pagination="112-117" refId="B57" refString="Koponen M, Askew RR (2002) Chalcids from Madeira, Canary Islands and Azores (Hymenoptera, Chalcidoidea). Vieraea 30: 112117." title="Chalcids from Madeira, Canary Islands and Azores (Hymenoptera, Chalcidoidea)." volume="30" year="2002">Koponen and Askew 2002</bibRefCitation>
), Canary and Balearic islands (
<bibRefCitation id="C2BD67B6C687FA200F7CBC30AF790AF2" author="Baez" firstAuthor="Baez" refId="B2" refString="Baez M (1998) Sobre la presencia del himenóptero polinizador de los “ Laureles de Indias ” en Canarias. Boletín de la Asociación Española de Entomología 22 (12): 225." year="1998">Báez 1998</bibRefCitation>
;
<bibRefCitation id="9E0BA10DD6DB506CF4E752E812DD5280" author="Carnero" etAl="et al." firstAuthor="Carnero" refId="B11" refString="Carnero Hernández A, Hernández García M, Pérez Padrón F, Hernández Suárez E (1998) Walkerella microcarpae and Eupristina (Parapristina) verticillata (pollinator agent) cause fruit ripening of Old World fig (Ficus species) in the Canary Islands. Boletin de la Asociacion Espanola de Entomologia 22 (34): 255." year="1998">Carnero et al. 1998</bibRefCitation>
;
<bibRefCitation id="376B6618E20D9A297AE7E201E5ABE037" DOI="10.1111/icad.12111" author="Wang R &amp; Aylwin R &amp; Barwell L &amp; Chen XY &amp; Chen Y &amp; Chou LS &amp; Cobb J &amp; Collette D &amp; Craine L &amp; Giblin-Davis RM &amp; Ghana S &amp; Harper M &amp; Harrison RD &amp; McPherson JR &amp; Peng Y-Q &amp; Pereira RAS &amp; Reyes-Betancort A &amp; Rodríguez LJV &amp; Strange E &amp; van Noort S &amp; Yang H-W &amp; Yu H &amp; Compton SG" issue="4" journalOrPublisher="Insect Conservation and Diversity" pagination="322-336" refId="B95" refString="Wang R, Aylwin R, Barwell L, Chen XY, Chen Y, Chou LS, Cobb J, Collette D, Craine L, Giblin-Davis RM, Ghana S, Harper M, Harrison RD, McPherson JR, Peng Y-Q, Pereira RAS, Reyes-Betancort A, Rodríguez LJV, Strange E, van Noort S, Yang H-W, Yu H, Compton SG (2015 a) The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa. Insect Conservation and Diversity 8 (4): 322336. https://doi.org/10.1111/icad.12111" title="The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa." volume="8">Wang et al. 2015 a</bibRefCitation>
,
<bibRefCitation id="B838EA7B984C1F34174F54634C3EDB0E" DOI="10.1016/j.biocontrol.2014.11.004" author="Wang R &amp; Aylwin R &amp; Cobb J &amp; Craine L &amp; Ghana S &amp; Reyes-Betancort J &amp; Quinnell R &amp; Compton SG" journalOrPublisher="Biological Control" pagination="21-30" refId="B96" refString="Wang R, Aylwin R, Cobb J, Craine L, Ghana S, Reyes-Betancort J, Quinnell R, Compton SG (2015 b) The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control. Biological Control 81: 2130. https://doi.org/10.1016/j.biocontrol.2014.11.004" title="The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control." volume="81">2015 b</bibRefCitation>
), and various Greek islands such as
<collectingRegion id="E9248856C3969D92CF9AECB028ECE21B" country="Greece" name="Kriti">Crete</collectingRegion>
, Cyclades,
<collectingCountry id="F3DBA1F3B35CC9C9C00F19D597E5F41C" name="Greece">Dodecanese</collectingCountry>
islands or
<collectingCountry id="2D9A7DD273842EB3812BB25E277038DD" name="Greece">Ionian islands</collectingCountry>
(
<bibRefCitation id="AC698909B3CC9D5B7F69ED0AA20569B7" DOI="10.1111/icad.12111" author="Wang R &amp; Aylwin R &amp; Barwell L &amp; Chen XY &amp; Chen Y &amp; Chou LS &amp; Cobb J &amp; Collette D &amp; Craine L &amp; Giblin-Davis RM &amp; Ghana S &amp; Harper M &amp; Harrison RD &amp; McPherson JR &amp; Peng Y-Q &amp; Pereira RAS &amp; Reyes-Betancort A &amp; Rodríguez LJV &amp; Strange E &amp; van Noort S &amp; Yang H-W &amp; Yu H &amp; Compton SG" issue="4" journalOrPublisher="Insect Conservation and Diversity" pagination="322-336" refId="B95" refString="Wang R, Aylwin R, Barwell L, Chen XY, Chen Y, Chou LS, Cobb J, Collette D, Craine L, Giblin-Davis RM, Ghana S, Harper M, Harrison RD, McPherson JR, Peng Y-Q, Pereira RAS, Reyes-Betancort A, Rodríguez LJV, Strange E, van Noort S, Yang H-W, Yu H, Compton SG (2015 a) The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa. Insect Conservation and Diversity 8 (4): 322336. https://doi.org/10.1111/icad.12111" title="The fig wasp followers and colonists of a widely introduced fig tree, Ficus microcarpa." volume="8">Wang et al. 2015 a</bibRefCitation>
,
<bibRefCitation id="8E723D42E7E79EE2E0C2C65FF2009CB6" DOI="10.1016/j.biocontrol.2014.11.004" author="Wang R &amp; Aylwin R &amp; Cobb J &amp; Craine L &amp; Ghana S &amp; Reyes-Betancort J &amp; Quinnell R &amp; Compton SG" journalOrPublisher="Biological Control" pagination="21-30" refId="B96" refString="Wang R, Aylwin R, Cobb J, Craine L, Ghana S, Reyes-Betancort J, Quinnell R, Compton SG (2015 b) The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control. Biological Control 81: 2130. https://doi.org/10.1016/j.biocontrol.2014.11.004" title="The impact of fig wasps (Chalcidoidea), new to the Mediterranean, on reproduction of an invasive fig tree Ficus microcarpa (Moraceae) and their potential for its biological control." volume="81">2015 b</bibRefCitation>
;
<bibRefCitation id="6EA7390A64D41842C8ABC7870A4FD647" DOI="10.3897/neobiota.98.139650" author="Koutsoukos" etAl="et al." firstAuthor="Koutsoukos" journalOrPublisher="NeoBiota" pagination="93-115" refId="B60" refString="Koutsoukos E, Konstantopoulou A, Parmakelis A, Demetriou J, Compton SG, Georgiadis C, Constantinidis T (2025) The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece. NeoBiota 98: 93115. https://doi.org/10.3897/neobiota.98.139650" title="The fellowship of the fig continues its journey: Fig. wasps (Hymenoptera, Chalcidoidea) associated with Ficus microcarpa in Greece." volume="98" year="2025">Koutsoukos et al. 2025</bibRefCitation>
) where
<taxonomicName id="908FBDF9AB5CD6E191EB5BAE3055E013" authorityName="Boucek" authorityYear="1993" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="D467E17DDB76AB5C5A2ADFAD9851A51F" italics="true">F. microcarpa</emphasis>
</taxonomicName>
is a frequent ornamental tree.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERQAE" type="description">
<paragraph id="E7A866A13123DD2BD0BF67F629BCCA29">
<heading id="6E0639806E06ED332AE0EAD638107556" reason="title">Short description.</heading>
</paragraph>
<paragraph id="1A5F3EA92961AE65DB0359124666EF49">
Female with head subquadrate, shorter than wide across compound eyes which are as long as the gena; with antennal scrobes separated. Antennae filiform; flagellar segments (except for the proximal two) with a whorl of long black
<emphasis id="BEEC0D7A4B5DA9A79B3D8516743C7F44" italics="true">sensilla chaetica</emphasis>
(
<bibRefCitation id="DD8BB7300D980A8D16E7A3E7C4ADB91C" author="Wiebes" firstAuthor="Wiebes" issue="1" journalOrPublisher="Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C" pagination="191-125" refId="B100" refString="Wiebes JT (1992) Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l. Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C 95 (1): 191125." title="Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l." volume="95" year="1992">Wiebes 1992</bibRefCitation>
;
<bibRefCitation id="00CECC58ABCB20642E80ECC196F345ED" DOI="10.1080/00222939300770071" author="Boucek" firstAuthor="Boucek" journalOrPublisher="Journal of Natural History" pagination="173-217" refId="B6" refString="Bouček Z (1993) The genera of chalcidoid wasps from Ficus fruit in the New World. Journal of Natural History 27: 173217. https://doi.org/10.1080/00222939300770071" title="The genera of chalcidoid wasps from Ficus fruit in the New World." volume="27" year="1993">Bouček 1993</bibRefCitation>
). Dorso-apical comb of fore tibia bearing two teeth; the hypopygium has a short acute spine. Body flattened. Pronotum extended laterally. Mesonotum and propodeum fused, the limit between them only marked by short lateral septa (
<bibRefCitation id="3BA8F4BA9F542548101F93799B7DDCAE" author="Wiebes" firstAuthor="Wiebes" issue="1" journalOrPublisher="Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C" pagination="191-125" refId="B100" refString="Wiebes JT (1992) Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l. Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C 95 (1): 191125." title="Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l." volume="95" year="1992">Wiebes 1992</bibRefCitation>
). Fore
<abbrev id="ABBRID0EFRAE" xlink_title="collector">leg</abbrev>
with three tarsomeres; mid and hind legs with four tarsomeres. Molecular analyses have demonstrated that
<taxonomicName id="21D98EF58C4512434C3BCF71C6297812" authorityName="Waterston" authorityYear="1921" class="Insecta" family="Agaonidae" genus="Eupristina" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="verticillata">
<emphasis id="50A388463669345A9CE07EA9BD19A68A" italics="true">E. verticillata</emphasis>
</taxonomicName>
, as presently understood, is in fact a species complex of at least three species (
<bibRefCitation id="DCE007990529FE1377437EC1593AAF2E" DOI="10.1186/1471-2148-11-86" author="Sun" etAl="et al." firstAuthor="Sun" refId="B88" refString="Sun XJ, Xiao JH, Cook JM, Feng G, Huang DW (2011) Comparisons of host mitochondrial, nuclear and endosymbiont bacterial genes reveal cryptic fig wasp species and the effects of Wolbachia on host mtDNA evolution and diversity. BMC Evolutionary Biology 11: 86. https://doi.org/10.1186/1471-2148-11-86" year="2011">Sun et al. 2011</bibRefCitation>
;
<bibRefCitation id="95D70538D9E3C39B0AFDD39D83B213D9" author="Wang" firstAuthor="Wang" refId="B94" refString="Wang R (2014) The Fig. Wasps associated with Ficus microcarpa, an Invasive Fig. Tree. PhD thesis. University of Leeds." year="2014">Wang 2014</bibRefCitation>
;
<bibRefCitation id="B317710BB8131C8D4747C3AC38024E25" DOI="10.1007/s10530-019-02132-4" author="Compton SG &amp; Stavrinides M &amp; Kaponas C &amp; Thomas PJ" journalOrPublisher="Biological Invasions" pagination="211-216" refId="B18" refString="Compton SG, Stavrinides M, Kaponas C, Thomas PJ (2020 b) No escape: most insect colonisers of an introduced fig tree in Cyprus come from the plants native range. Biological Invasions 22: 211216. https://doi.org/10.1007/s10530-019-02132-4" title="No escape: most insect colonisers of an introduced fig tree in Cyprus come from the plants native range." volume="22">Compton et al. 2020 b</bibRefCitation>
). This may explain the discrepancies observed in diagnostic characters used by several authors, such as the ovipositor length, which has been classified as shorter (
<bibRefCitation id="7514FA1DB7AAFF1AB80B7A3BC64D46FF" author="Wiebes" firstAuthor="Wiebes" issue="1" journalOrPublisher="Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C" pagination="191-125" refId="B100" refString="Wiebes JT (1992) Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l. Proceedings of the Koninklijke Nederlandse Akademie Wetenschappen Amsterdam C 95 (1): 191125." title="Agaonidae (Hymenoptera: Chalcidoidea) and Ficus (Moraceae): fig wasps and their figs. VIII. Eupristina s. l." volume="95" year="1992">Wiebes 1992</bibRefCitation>
) or longer (
<bibRefCitation id="31346B10476027A9FE17D09A045E2198" DOI="10.1080/00222939300770071" author="Boucek" firstAuthor="Boucek" journalOrPublisher="Journal of Natural History" pagination="173-217" refId="B6" refString="Bouček Z (1993) The genera of chalcidoid wasps from Ficus fruit in the New World. Journal of Natural History 27: 173217. https://doi.org/10.1080/00222939300770071" title="The genera of chalcidoid wasps from Ficus fruit in the New World." volume="27" year="1993">Bouček 1993</bibRefCitation>
) than the gaster.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,790 @@
<document id="26567DD7C4CC995CBA8B24262E5AC42D" ID-DOI="10.3897/zookeys.1244.148112" ID-ZooBank="CCCA8237-6587-44BD-A25E-41BD00928A11" ID-publisher-id="148112" URI-arpha="9A0DFA26-0AED-523E-937A-37D6A902E41E" URI-zoobank="http://zoobank.org/CCCA8237-6587-44BD-A25E-41BD00928A11" article-type="research-article" checkinTime="1751465287194" checkinUser="pensoft" docAuthor="Huang, Chao, Shih, Hsi-Te &amp; Ahyong, Shane T." docDate="2025" docId="FEBCE5AD8B235E49A60056E5B8027C78" docLanguage="en" docName="ZooKeys 1244: 1-27" docOrigin="ZooKeys 1244" docSource="https://zookeys.pensoft.net/article/148112/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megapleonum falx Huang, Shih &amp; Ahyong, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="27" masterDocId="9A0DFA260AED523E937A37D6A902E41E" masterDocTitle="Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="1" updateTime="1751465704327" updateUser="ExternalLinkService">
<mods:mods id="EBD81771E6CACAC536396C1BF15FBA80" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="68FEBCED1F919D4FCB92C31AECE83ED5">
<mods:title id="CCBBCBD2119E2320F5BDAE001787BFF6">Four new freshwater crab species of the genus Megapleonum Huang, Shih &amp; Ahyong, 2018 (Crustacea, Decapoda, Potamidae) from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="9300BC864CBCA9E4A750364A987DA5E1" type="personal">
<mods:role id="EDC9B410124E49559C0B3476798931E1">
<mods:roleTerm id="A5A84A53A4DC42AE46F8E956F9B55EB3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="598CBDD5EC3D59649FF6390428A9F905">Huang, Chao</mods:namePart>
<mods:nameIdentifier id="C04FD463741A182D04B467CCB42702CA" type="ORCID">0000-0002-3782-9096</mods:nameIdentifier>
<mods:affiliation id="E0A21328D23954768561BEA00DC9500A">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="AE989737ACF3765B200C364CA1360D40" type="personal">
<mods:role id="177BC4F6D171675DB1B97F788DE4A033">
<mods:roleTerm id="6EBEC33087E66952BF8E5F41090DAD53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0EC5F4225399DB7611474731B4D9BA0">Shih, Hsi-Te</mods:namePart>
<mods:nameIdentifier id="AD51FD8CAC08357D3F7E2668612C56DC" type="ORCID">0000-0003-1317-8783</mods:nameIdentifier>
<mods:affiliation id="E4022413F6D0AC46151A6C936DE9FAD8">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="934EDFE01B19D1F0FCA5EF9DFC26C2E4" type="personal">
<mods:role id="7EBF3D1EEBF7157298E0E6EE833DCA55">
<mods:roleTerm id="6B778EC325360312616180461DA7399F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA13B8216F49A71D7D4C5E10AE71E416">Ahyong, Shane T.</mods:namePart>
<mods:nameIdentifier id="F2D87FB3840337CD0C586C27451F1A9F" type="ORCID">0000-0002-2820-4158</mods:nameIdentifier>
<mods:affiliation id="82DD2CA7D0803319BE14ADAFB0E67C18">Australian Museum, 1 William St, Sydney NSW 2010, Australia &amp; School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E2B462D9A0FF1C31065DECD2F2ECA88A">text</mods:typeOfResource>
<mods:relatedItem id="971AC38F62B5B173F841F00D56207C73" type="host">
<mods:titleInfo id="1BD8C5BA2BB778BEA7BBAEFD15AE605D">
<mods:title id="18A93159D958A23755BE33D9BFB51F29">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="01504EFC484C3CE63D44B3E86990E7C7">
<mods:date id="4BBB569D1E7495DB0955E4D6230A2452">2025</mods:date>
<mods:detail id="E2FC3A5FA573FD4EC5FB607252134247" type="pubDate">
<mods:number id="6DFE81D91FA130CE83DE3B78E8D7CBD4">2025-07-02</mods:number>
</mods:detail>
<mods:detail id="C354244D7BE8F27ECE68556398A189B0" type="volume">
<mods:number id="AF4BD0DE41AF74B92E947B70DA27D07B">1244</mods:number>
</mods:detail>
<mods:extent id="C9C49023405B1E9E28C920F29F887536" unit="page">
<mods:start id="B1CAF76C3C42D32033CCF3629D949096">1</mods:start>
<mods:end id="325419E6310DA42EB3118DAF9BC3CB13">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="42F5BDE5A8856397682CAEB8C09A43A3">journal article</mods:classification>
<mods:identifier id="43D280DFCB576DFAF358BEAEA8550F98" type="DOI">10.3897/zookeys.1244.148112</mods:identifier>
<mods:identifier id="37A10EF88F720A008A35854F86CE2D81" type="ZooBank">CCCA8237-6587-44BD-A25E-41BD00928A11</mods:identifier>
</mods:mods>
<treatment id="FEBCE5AD8B235E49A60056E5B8027C78" ID-DOI="http://doi.org/10.5281/zenodo.15791866" ID-Zenodo-Dep="15791866" ID-arpha="FEBCE5AD-8B23-5E49-A600-56E5B8027C78" ID-zoobank="https://zoobank.org/8F1A2D8E-5CC3-4DA2-AF9D-50DC0C1885D7" LSID="urn:lsid:plazi:treatment:FEBCE5AD8B235E49A60056E5B8027C78" LSID-ZBK="urn:lsid:zoobank.org:act:8F1A2D8E-5CC3-4DA2-AF9D-50DC0C1885D7" httpUri="http://treatment.plazi.org/id/FEBCE5AD8B235E49A60056E5B8027C78">
<subSubSection id="3FB1C83D29F7112F2D9E27EE55822601" type="nomenclature">
<paragraph id="2A76B0F0B6D93290CC5D2690492A6704">
<taxonomicName id="279384596EE92A9C28C3F90F7F8E968A" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="7A9C3DAE86BACA502B7DFFFB227FDF3B" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0738B3F0A1541BD510AD9AC2820DEB7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="44FB9B83927076ACA9F6FFEBDCF775C1" type="description">
<paragraph id="9C9B5D6F67898BAD06926898E863819C">
<figureCitation id="901DC6E281028795966F796BA5C36936" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">Figs 1</figureCitation>
,
<figureCitation id="FD67828F3CD94E1E4324AE1448192F0A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKFAE" type="type material">
<paragraph id="DA7AA751E89FAC21D08562E998663488">
<heading id="A2C16613A0B4286680F86F0CED8722D9" reason="title">Type material.</heading>
</paragraph>
<paragraph id="9775C1C19B394188C7548021D3ED82D4">
<materialsCitation id="6BC85D90FAD3BD24F790C84A0C7A52F3" collectingDate="2019-08" collectionCode="SYSBM" country="China" county="Huidong County" latitude="23.03" location="Baima Village" longLatPrecision="754" longitude="115.07" municipality="Wang" specimenCode="SYSBM 002142" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="9C197A837FFA1A0E7EFC8FCEA8C0C3FF" bold="true" italics="true">
<typeStatus id="C92ABDAB9D1736122009618EDA522DAE" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCode id="B38E0E5ADFA43C1F73B26D87EC52C551" collectionCode="SYSBM">
<abbrev content-type="institution" id="ABBRID0EVFAE" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002142
</specimenCode>
, male (19.6 ×
<quantity id="C79954869B58E0EC69A11A39FDAC047E" metricMagnitude="-2" metricUnit="m" metricValue="1.58" unit="mm" value="15.8">15.8 mm</quantity>
),
<location id="FAB0043785DDCE44C51E38BF2FAD1EF1" LSID="urn:lsid:plazi:treatment:FEBCE5AD8B235E49A60056E5B8027C78:FAB0043785DDCE44C51E38BF2FAD1EF1" country="China" county="Huidong County" latitude="23.03" longLatPrecision="754" longitude="115.07" municipality="Wang" name="Baima Village" stateProvince="Guangdong">Baima Village</location>
,
<collectingCounty id="5CD66496DBCA6E6445ECFAD85D5A6F25">Huidong County</collectingCounty>
,
<location id="B0F8338C84BC3D725B352E1E2D45B100" LSID="urn:lsid:plazi:treatment:FEBCE5AD8B235E49A60056E5B8027C78:B0F8338C84BC3D725B352E1E2D45B100" country="China" county="Huidong County" latitude="23.03" longLatPrecision="754" longitude="115.07" municipality="Wang" name="Huizhou City" stateProvince="Guangdong">Huizhou City</location>
,
<collectingRegion id="CC0EC205AE2C401D323D649226A4F783" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="EF8E8DB0F5F84D371F61F493D3CF018C" name="China">China</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4FAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.070000,23.030000]}">
<geoCoordinate id="E60E6F5D00C706C50C8E59FC23DA01CF" degrees="23.03" direction="north" orientation="latitude" precision="555" value="23.03">23.03 ° N</geoCoordinate>
,
<geoCoordinate id="18F6416BF2BEB994218130FE34F7BA1C" degrees="115.07" direction="east" orientation="longitude" precision="555" value="115.07">115.07 ° E</geoCoordinate>
</named-content>
, hill stream under rock, coll. Song-Bo
<collectingMunicipality id="476842BE050E3635E26691BFE6392249">Wang</collectingMunicipality>
,
<date id="C70F608793BE5B4D6AB1F6D041C50CA3" value="2019-08">
<collectingDate id="0A23017AD3083D456A4678EBC3A2A08C" value="2019-08">August 2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECGAE" type="diagnosis">
<paragraph id="EC1E5C1599B59358FAA092EBA7077A48">
<heading id="3B9908F5BEB6B18FDDC032CA03373090" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="91A0F986AF703B474C95537AE17BDA72">
Carapace broader than long, dorsal surface slightly convex, lateral margins covered in dense setae; postorbital, epigastric cristae confluent (Fig.
<figureCitation id="91D1C4BD240F21347B0F3E6E790119E1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Maxilliped 3 sparsely covered in long setae; merus width ~ 1.4 × length; ischium width ~ 0.7 × length; exopod reaching beyond anterior edge of ischium, flagellum as long as merus width (Fig.
<figureCitation id="4D71AA576FBCD9800B2261D5B7E43763" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 A</figureCitation>
). Ambulatory legs with dense setae; pereiopod 5 dactylus ~ as long as propodus (Fig.
<figureCitation id="69FF300A54C455BA029CF3F099E780E7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Male anterior thoracic sternum very broad, width ~ 1.8 × length (Fig.
<figureCitation id="6E5F05BF70E38C59BACF7B0A20B28CA5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 C</figureCitation>
). Male pleon large, broadly triangular, pleonite 6 width ~ 2.6 × length, telson width ~ 1.8 × length (Fig.
<figureCitation id="979FFC19EEECBDB597984F86B7D0FA05" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 D</figureCitation>
). G 1 large, strongly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="1BB6453E0CB142A5E226E23770630844" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="B8E6810E4277EF12DC34793490016FAA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 E</figureCitation>
); subterminal segment length ~ 3.2 × length of terminal segment (Fig.
<figureCitation id="1E50FDA5B5588560C2A0C359A741D90F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 C</figureCitation>
). Subterminal segment outer margin strongly concave; terminal segment strongly curved inwards, strongly tapering, sickle shaped, tip pointed downwards in dissected view (Fig.
<figureCitation id="8C4A51D0AAEFF849C0C8DDE11A8D0018" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 C, F, G</figureCitation>
). G 2 subterminal segment tapering, bent outwards distally, flagelliform terminal segment ~ 1.8 × length of subterminal segment, apex blunt (Fig.
<figureCitation id="D6AE67B1F6EDD8DE1D9A4F24068594A0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 B</figureCitation>
).
</paragraph>
<caption id="E2157A7E6EE402D9677CB7E9085977A4" ID-DOI="10.3897/zookeys.1244.148112.figure1" ID-arpha="042CE975-295E-5FF3-884F-FD4A41212858" httpUri="https://binary.pensoft.net/fig/1366338" startId="F1">
<paragraph id="9533BF636A24385E112375C629FA1E9D">
<label id="DD5E2FE5919931CF9B8205A4680941C1">Figure 1.</label>
</paragraph>
<paragraph id="8A95F619E0231EE8F079471B41989759">
<taxonomicName id="7DACC68F274849CBFB4993BB5E058C9C" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="7BBE6D4189BA83674AF9E3C027288596" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEC9219B28DF114B99FC970B87A39B0B" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (19.6 × 15.8 mm),
<abbrev content-type="institution" id="ABBRID0E6HAE" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002142. Dorsal habitus (
<emphasis id="EEA60BBC6A7B3E821B93849A8F470EE5" bold="true">A</emphasis>
); cephalothorax, anterior view (
<emphasis id="AFA6926CC092D02FDBAC299F7087C03E" bold="true">B</emphasis>
); anterior thoracic sternum (
<emphasis id="EAD962958C6120EB94A37BBB3A793455" bold="true">C</emphasis>
); anterior thoracic sternum and pleon, ventral view (
<emphasis id="BF063CBAEDCB67E279ED3C78AACBB584" bold="true">D</emphasis>
); sterno-pleonal cavity with G 1 in situ, ventral view (
<emphasis id="165C491B7883F799A223F1C30733940A" bold="true">E</emphasis>
).
</paragraph>
</caption>
<caption id="6FA9CC3DA3620CB14197FF37FFA8B852" ID-DOI="10.3897/zookeys.1244.148112.figure2" ID-arpha="7487EAB4-317A-56F1-9668-58252134843F" httpUri="https://binary.pensoft.net/fig/1366339" startId="F2">
<paragraph id="F121C2650A8962C16FDAF17BF3D0D813">
<label id="143A82DDE1801EF0DA761054E676A22A">Figure 2.</label>
</paragraph>
<paragraph id="ED15CFBEAB8FC4B1B0BB282A7694D59D">
<taxonomicName id="35F4579B91C4F2A7D8D7EA953B97A2EF" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="F57A8DDA10A46A3CDBC302DF19459E3D" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DB1EBC2DB51D03E7B9E371CE83158CCB" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (19.6 × 15.8 mm),
<abbrev content-type="institution" id="ABBRID0EGJAE" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002142. Left maxilliped 3 (
<emphasis id="28B54BAE46BA43FFFE4DCF4E9752F7D3" bold="true">A</emphasis>
); left G 2, pleonal view (
<emphasis id="5361701FF6B15632772E18638E9CAC5B" bold="true">B</emphasis>
); left G 1, ventral view (
<emphasis id="676EBD7143D5F0F9CE5705A2B29BA485" bold="true">C</emphasis>
); major cheliped, partially damaged (
<emphasis id="3661A961ADAAB80150E08D8DEDD857F1" bold="true">D</emphasis>
); minor cheliped (
<emphasis id="432F09A67620CBEB8E9B12B50794CE59" bold="true">E</emphasis>
); left G 1 terminal segment, ventral view (
<emphasis id="2E6C02020A91C35526CA7B4B5A985AC0" bold="true">F</emphasis>
); left G 1 terminal segment, dorsal view (
<emphasis id="9FFFCA334088BB722D9DFA8A0AC56DEE" bold="true">G</emphasis>
). Scale bars: 1.0 mm (
<emphasis id="BC850B79057CA6F059219D0F0B5A0488" bold="true">A</emphasis>
<emphasis id="700D92A5E7E22011C6F9B1AB4BDD035F" bold="true">C</emphasis>
,
<emphasis id="E5A2B2A16D7C097F6553037C34436306" bold="true">F</emphasis>
,
<emphasis id="CD993681CF9D650F53C201E6AF1381F3" bold="true">G)</emphasis>
, 5.0 mm
<emphasis id="BE799B6D96120B9BF872924E18E74492" bold="true">(D</emphasis>
,
<emphasis id="0058ED9CA2A2E929E94E5260B1FE53A1" bold="true">E)</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKKAE" type="description">
<paragraph id="80F16326209F67C0ACA9A95D846C8F60">
<heading id="88D332A9D52D81185AA7AB70E0695D91" reason="title">
Description of male
<typeStatus id="A84467A294D2074E59BE7277E9E4E3FB" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="EB5BF41814277E29CCD24D11FDDEBFA7">
Carapace broader than long, ~ 1.2 × as wide as long; regions not pronounced, dorsal surface slightly convex, finely pitted; dense setae along lateral margins (Fig.
<figureCitation id="259E8D80C6FD8AA7712877025DDBE7A5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Frontal margin almost straight, deflexed (Fig.
<figureCitation id="F96FF7B50DD115C8DE346DF212D9B7DA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Epigastric cristae and postorbital cristae relatively smooth, confluent; bifurcated groove between epigastric cristae (Figs
<figureCitation id="26908519595915D1B5421FFD068C0FB0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
,
<figureCitation id="77085E4C3C301BB25A50A9F59214E41D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 A</figureCitation>
). Branchial regions not swollen (Fig.
<figureCitation id="813D0883661C4B957F83C0C0DA4C8140" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Cervical groove shallow but visible (Fig.
<figureCitation id="B7C13D7A3711621CFAB87052926F42F2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Mesogastric region flat (Fig.
<figureCitation id="2D7E7C6C992625BC750D7CF82EB60287" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). External orbital angle broadly triangular, outer margin slightly convex, confluent with anterolateral margin (Fig.
<figureCitation id="B8AA76CDCC68B5AFB6D75F3FC4678712" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Epibranchial tooth granular, indistinct (Fig.
<figureCitation id="B6C14A84B14B67E61DBA1D71908A89FF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Anterolateral margin lined with indistinct single or partially fused granules; posterolateral margin posteriorly convergent (Fig.
<figureCitation id="FAF1B11CA7B639C143903297A3BAB13C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
); posterolateral surface slightly rugose (Fig.
<figureCitation id="DA65941BB160C87B7C516BF8B6C1C0FF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Orbits regular; supraorbital margins weakly cristate, infraorbital margins lined with fused granules (Fig.
<figureCitation id="CDC766ED877EEA2D1836506CEF5A1144" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
). Eyes normal (Fig.
<figureCitation id="CC948B608A9C4E68C06C8B15F99DA30D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Sub-orbital, pterygostomial and sub-hepatic regions generally smooth, pitted (Fig.
<figureCitation id="7F92547D111C37F5C9FDA30F6216566C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
). Antennules large, folded within broad fossae; antennae very short (Fig.
<figureCitation id="81C53B6A61B515BEE07915C072BBFB54" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
). Median lobe of epistome buccal margin broadly triangular, lateral margins slightly sinuous (Fig.
<figureCitation id="3A4B0EA12C21DDFFD369F6A37EADD682" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
).
</paragraph>
<paragraph id="28DB8B4044CE4A1AAFE0D68F27B686C5">
Maxilliped 3 sparsely covered in long setae, merus subtrapezoidal, with slight median depression, width ~ 1.4 × length; ischium subtrapezoidal with shallow median sulcus, distomesial margin rounded, width ~ 0.7 × length. Exopod reaching proximal one-third of merus; flagellum as long as merus width (Fig.
<figureCitation id="77DF92721FB35FBBD1242369BC421684" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 A</figureCitation>
).
</paragraph>
<paragraph id="F2C73B5C68B16A9072013BF95B59E06C">
Chelipeds (pereiopod 1) subequal (Fig.
<figureCitation id="C169DFB4D6FA0B6E6A8669BB02D90ADE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 D, E</figureCitation>
). Merus trigonal in cross-section, surfaces generally smooth; outer dorsal and ventral margins slightly crenulated, inner margin lined with large granules (Fig.
<figureCitation id="2270EF138ECD4E78856FCB5D550E67D9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A, B</figureCitation>
). Carpus dorsal surface slightly rugose, with large spike at inner-distal angle, spinule at base (Fig.
<figureCitation id="B5B3D29EB050F1F5B63F0370828BC37C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Major cheliped palm length ~ 1.5 × height; dactylus 0.7 × palm length (Fig.
<figureCitation id="0DC06AD4FBEA5F2C95CF1993D64C3622" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 D, E</figureCitation>
). Palm surface pitted, occlusal margin of fingers with 79 irregular blunt teeth, with small gape when closed (Fig.
<figureCitation id="E2722BC6C0616F9A55358080E41E97A7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 D, E</figureCitation>
).
</paragraph>
<paragraph id="6EE7F2EBDE77ECA7749D8E1279D27E30">
Ambulatory legs (pereiopods 25) slender, covered with setae (Fig.
<figureCitation id="202C9091EEB2683BCE88D1378ADD231E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Pereiopod 3 merus 0.6 ×
<abbrev id="ABBRID0ESNAE" xlink_title="Carapace length">CL</abbrev>
(Fig.
<figureCitation id="63C68B1277B7D3EF158BCE19C7AEC6BE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
). Pereiopod 5 propodus length 1.7 × height, approximately as long as dactylus (Fig.
<figureCitation id="79621C9DBAF0A870CFB17E2622BD1538" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
).
</paragraph>
<paragraph id="5D09F0464F9242EDD27E7AB12A358E18">
Male thoracic sternum generally smooth, pitted, setae at margins and sparsely on sternites; sternites 14 width ~ 1.8 × length; sternites 1, 2 fused to form broad triangle; fused sternites 1, 2 demarcated from sternite 3 by sinuous transverse sulcus, sulcus lined with setae; sternites 3, 4 fused without obvious demarcation other than a line of setae (Fig.
<figureCitation id="3FB64604522C2FF50446287F53C84292" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 C</figureCitation>
). Male sterno-pleonal cavity reaching anteriorly slightly beyond mid-length of cheliped coxa (Fig.
<figureCitation id="E3874200DC2A6B00B8A1437E59A478DF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 C</figureCitation>
). Male pleonal locking tubercle positioned at mid-length of sternite 5 (Fig.
<figureCitation id="7BDF1936655D63A1C243EAF46BCF0B4D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 E</figureCitation>
).
</paragraph>
<paragraph id="AC102494B42FA272B689B6B0104671DB">
Male pleon large, broadly triangular; somites 36 progressively narrower, not entirely confluent with each other; somite 6 width approximately 2.6 × length; telson width 1.8 × length; lateral margins almost straight, apex rounded (Fig.
<figureCitation id="520503E2953068BA1713777DB48F590C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 D</figureCitation>
).
</paragraph>
<paragraph id="2F8D1D3119D55CB697D5386FCC18C421">
G 1 large, strongly sinuous, tip exceeding suture between thoracic sternites 4 /
<quantity id="CAD870B9CAB32AE67EA27E16A380DD9D" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
situ (Fig.
<figureCitation id="C8443B669AF7AAA2E0F7F4717B3F3121" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 E</figureCitation>
); subterminal segment length ~ 3.2 × length of terminal segment (Fig.
<figureCitation id="392F3269D31744EB394137F706F269DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 C</figureCitation>
). Subterminal segment outer margin strongly concave; terminal segment strongly curved inwards, strongly tapering, sickle shaped, tip pointed downwards in dissected view (Fig.
<figureCitation id="DD9978316BC3247CC29B0B1BD90242DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 C, F, G</figureCitation>
). G 2 subterminal segment tapering, bent outwards distally, flagelliform terminal segment ~ 1.8 × length of subterminal segment, apex blunt (Fig.
<figureCitation id="6261D4545BD4E272D597D6DEEF1332F6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPAE" type="Colour in life">
<paragraph id="1AB2BB15A8AEDF68DB017F5A54F4D3EA">
<heading id="E63A3DCD18D2E9201F698EBFA85B714C" reason="title">Colour in life.</heading>
</paragraph>
<paragraph id="8DDF9D98A936FB3A11AAF6927570DCE3">Generally camouflage-brown all over.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJPAE" type="habitat">
<paragraph id="CED9DDB271388CCC3EE9CD55B2E0C314">
<heading id="94C57F6096E203BD6D266D4B8AA4E37F" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="4EB21A322B468D4BA31814C887D0CF8A">
Unknown. The only specimen collected was found in the lower reaches of the hill stream, but further collection efforts in the same area and further upstream yielded no crabs at all. We consider it likely that the primary habitat of this species is higher up the mountain, which reaches above
<quantity id="8F160FB56421C77FEDDE9AF84182FCE0" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
above sea level, and that this lone specimen that was collected was washed downstream in a flooding event.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOPAE" type="distribution">
<paragraph id="0EAB146D740D127E28AC7617C24D2943">
<heading id="F21053279F7D90AF1AE54496D557F6F9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="6F3DDA89DBD0E689EA624FAD53781901">
Baima Village, Huidong County, Huizhou City,
<collectingRegion id="A93311309E2B4AF06D568CC0ED80E2B5" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCountry id="DD072AD3021C5A98260B0CADFDFA9605" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETPAE" type="etymology">
<paragraph id="214038A964BD3DBD5641D9D6AA5082D8">
<heading id="1C9A6410124A38AE9420F92B521FBE1D" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="445B588847A953234C9BDBD0690BDEA4">
The species name is the Latin word
<emphasis id="1F9309DB62B70F20762B31FBA3E35E5F" italics="true">falx</emphasis>
which means sickle-shaped. It alludes to the sickle-shaped G 1 terminal segment of this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDQAE" type="remarks">
<paragraph id="4899C078C58F43DAD31498738E553EE4">
<heading id="79F51705F6CD27F7A3F9E92FA8E1F544" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="3B0AE5488E67B895FE5D99BA68C31CEE">
The wide male anterior thoracic sternum, large and broadly triangular male abdomen and large and sinuous G 1 of
<taxonomicName id="E2684CF4C748694A16C0481A789FBEE3" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="19F087276F505DA037071D4943E3B100" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81C1BF0D8D8729B3590E506E79382A35" rank="species">sp. nov.</taxonomicNameLabel>
fit the diagnosis of the genus. In possessing thick setae on the carapace margins and ambulatory legs,
<taxonomicName id="4E202E1E3A16B940E563594BFF82CE63" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="A0171F6110F6E8978D3D2B2C5C63EBE9" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F47E4797FF590BBA8D32D78E0B88CC2" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="FBDC6319B530FDACE4E0FB66C78E075C" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="F6021D11E0783600C092B04CC7CD4B87" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BDC25CB8B7E0145F7D1B33BF08354C7" rank="species">sp. nov.</taxonomicNameLabel>
The sickle-shaped G 1 terminal segment of
<taxonomicName id="5254E9AD6615C3FE20212DD24F6A5B03" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="30588BC1DE4DC75D4CB8F1FF3E9356E0" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="710B521584B662166AFE33F229D834E2" rank="species">sp. nov.</taxonomicNameLabel>
, however, is unlike any other congener and immediately distinguishes it (Fig.
<figureCitation id="B8FD5552196496A3305AD9DB9EA89B76" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 C, F, G</figureCitation>
). Apart from the G 1,
<taxonomicName id="C5BF09A3D3B55B2B0511B4114C354238" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="959EB93347FE099BCB0221D16D6783FA" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2CDE32D8B88284CF92AF07216D98094" rank="species">sp. nov.</taxonomicNameLabel>
also has a flatter carapace dorsal surface when compared to
<taxonomicName id="16FBB9B382C6F9063CF46DAE3574B39B" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="6426D0EA57CAD99BA202FDF03B731EFA" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7A5614D623D63DED8BC1724FB4791E37" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="11B7BFDCC56DDD71CBD01E389CE180DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 B</figureCitation>
,
<figureCitation id="27C26B3E714B8C3535143ECE4CBFED7D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 B</figureCitation>
). The flagellum of the maxilliped 3 exopod is present in
<taxonomicName id="C66281B86B3A2DA2E5F3039C9145FC49" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="C53B3C6ADAC4CA4006F7E0DE9CF5CFD6" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FC6940770FD10D1C83EBEEF6715D0A39" rank="species">sp. nov.</taxonomicNameLabel>
whereas it is absent in
<taxonomicName id="A465604A680B3EA4F290153962C63AB4" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="02E2CC849534904559ACE5E61A283ACD" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24E3DF8081EC249FDBA31333DF24C7D0" rank="species">sp. nov.</taxonomicNameLabel>
(Fig.
<figureCitation id="70357C09A41F436F78D9939B4FAD8CDA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 A</figureCitation>
vs Fig.
<figureCitation id="978B15B8077E7F1B2815C77FD1269C6F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 A</figureCitation>
). The male abdomen is also obviously wider in
<taxonomicName id="8662817E6630AF20304B4F93DEB830A6" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="ED8AC6329802228B4609BD47E47EE2DA" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29D572B2AA4C6FEE16A80F309881C2ED" rank="species">sp. nov.</taxonomicNameLabel>
, with the somite 6 width approximately 2.6 × length (Fig.
<figureCitation id="CDB62AD5B2E9082800699574F094930F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 D</figureCitation>
) (vs narrower in
<taxonomicName id="4DCD2CDEE88FF0F38DBA200436FF8B2F" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="9E376813FBD548BF43A443FBD351EF92" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E53B24B61DEE9D3F458120A383ECB1AE" rank="species">sp. nov.</taxonomicNameLabel>
, sixth somite width approximately 2.5 × length, Fig.
<figureCitation id="4EF2FC77BDEBCBC719D51925EB2DC3B8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 D</figureCitation>
). The relative length of the pereiopod dactylus to the propodus is also different, being approximately the same length in
<taxonomicName id="2EF266E9FD59908F8E76FC1B2B768DC2" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="011CC3F02DFBBDCB7AF53C51C3344363" italics="true">M. falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0D513C2D782B9C9B9BA4033FEB37A5B1" rank="species">sp. nov.</taxonomicNameLabel>
and in
<taxonomicName id="FE3F03FFFE673C4B9D60A2EC8DD3794E" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="0AA7C335CD51C36F7BC54D5C54A82943" italics="true">M. ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBF4570543345A41A552D49C7037B124" rank="species">sp. nov.</taxonomicNameLabel>
, having a shorter dactylus (Figs
<figureCitation id="442BCEED6A14B9151EE1FF6A5B43A395" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
,
<figureCitation id="89D46B8809CE5C3B46A2E5C1112A14EC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
). More detailed comparisons can be found in Table
<tableCitation id="5DA116EC7AD7B3E20FFC815E9EBB6A9F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparisons between the species within the genus Megapleonum." httpUri="http://table.plazi.org/id/69D766E64FF15F0F8BB8C5782AADA37B" tableUuid="69D766E64FF15F0F8BB8C5782AADA37B">1</tableCitation>
.
</paragraph>
<caption id="B320431E95AEEA2B51FCD7F80A175F23" ID-DOI="10.3897/zookeys.1244.148112.figure3" ID-arpha="D32D36F2-2B51-5BEC-A838-B7D42BA6E98B" httpUri="https://binary.pensoft.net/fig/1366340" startId="F3">
<paragraph id="5CAEF7A82C2814038C6D1EB8DFDEF8D0">
<label id="0EE1FDD8481D860AD8119F1B535952A2">Figure 3.</label>
</paragraph>
<paragraph id="422AB329DD329E2B42C6AC529B953DA5">
<taxonomicName id="67E346F99AD8493E3D148FA3AB42CC4F" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="EB033E19767782BD464DD644855EA1E3" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="774ABEC31998012C6B7117C4D2D112C4" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (16.6 × 13.1 mm),
<abbrev content-type="institution" id="ABBRID0EIWAE" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002143. Dorsal habitus (
<emphasis id="D0EC3C33C1883E30F9C65A5536EA451D" bold="true">A</emphasis>
); cephalothorax, anterior view (
<emphasis id="224A7B20144A1A22201A3F83F4C49D14" bold="true">B</emphasis>
); anterior thoracic sternum (
<emphasis id="06D7A19758D94DFFC26C8FD7CB3FD4EB" bold="true">C</emphasis>
); anterior thoracic sternum and pleon, ventral view (
<emphasis id="90C22F5ACA574F2FE5A318FB61FCC47A" bold="true">D</emphasis>
); sterno-pleonal cavity with G 1 in situ, ventral view (
<emphasis id="F91B1E6B21DB3D8170A364815FE02CDC" bold="true">E</emphasis>
).
</paragraph>
</caption>
<caption id="34E0FFDE5736ACD04F50FB988F10A50E" ID-DOI="10.3897/zookeys.1244.148112.figure4" ID-arpha="B567F8F1-75BB-5099-B9CD-10191EBDFD5F" httpUri="https://binary.pensoft.net/fig/1366341" startId="F4">
<paragraph id="24349B19D4B9A9E48A4CACCAC435A00B">
<label id="34FAE3DC622429FC5FB24D06FF52E08E">Figure 4.</label>
</paragraph>
<paragraph id="655400CEEC683ED1E5A590334094129E">
<taxonomicName id="478C8A5539914102FB46DFF394C66F73" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="97289E14810053ADC972D625A9F8A99B" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0A4B2C6C04E61ACC2AD92C1C96E38D9C" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (16.6 × 13.1 mm),
<abbrev content-type="institution" id="ABBRID0EPXAE" xlink_title="Sun Yat-sen Museum of Biology, Sun Yat-sen University, Guangzhou, China">SYSBM</abbrev>
002143. Left maxilliped 3 (
<emphasis id="A550EE6A857E6F114F529A37CCF72DBA" bold="true">A</emphasis>
); left G 2, pleonal view (
<emphasis id="C47F59C7EBACDC33CB90160099EC93E3" bold="true">B</emphasis>
); left G 1, ventral view (
<emphasis id="C86E797C44F49D18D9117D4A4B60E04F" bold="true">C</emphasis>
); major cheliped (
<emphasis id="98232940F68A7BBF242760F27AAAC905" bold="true">D</emphasis>
); minor cheliped (
<emphasis id="2F17CA54B267EDC05D792C3C86016AB9" bold="true">E</emphasis>
); left G 1 terminal segment, ventral view (
<emphasis id="D4962BCF0E462FCEF3A8412CABAC47C8" bold="true">F</emphasis>
); left G 1 horse-shoe structure viewed from the top (
<emphasis id="E6F1BB56260EDA9F10038C9C8C4543C2" bold="true">G</emphasis>
); left G 1 terminal segment, dorsal view (
<emphasis id="4FCDC874BC5BA3E30F0EB08044B0FC44" bold="true">H</emphasis>
). Scale bars: 1.0 mm
<emphasis id="A2BB3EA90E3F1C408B3740B91D085A01" bold="true">(A</emphasis>
<emphasis id="0C1F1790E822174906C036BF38E0D76D" bold="true">C</emphasis>
,
<emphasis id="7D7DDCB3364F305DA0EB37C3D97A0EF0" bold="true">F</emphasis>
<emphasis id="61F1C107FB073E15BC03BA65FF500968" bold="true">H)</emphasis>
, 5.0 mm
<emphasis id="19D09EACE636C8D91E3C3907BCACB050" bold="true">(D</emphasis>
,
<emphasis id="91952955BB359458C24CC52FA60A67DF" bold="true">E)</emphasis>
.
</paragraph>
</caption>
<caption id="69D766E64FF15F0F8BB8C5782AADA37B" ID-Table-UUID="69D766E64FF15F0F8BB8C5782AADA37B" httpUri="http://table.plazi.org/id/69D766E64FF15F0F8BB8C5782AADA37B" startId="T1" targetIsTable="true">
<paragraph id="46E21B7CFB198F093481EFE82EC5EFBA">
<label id="FD33A34AF564C92AB6850A9B2EBC488D">Table 1.</label>
</paragraph>
<paragraph id="BFFEB59EAD9823F5E6E302A13755AC39">
Morphological comparisons between the species within the genus
<taxonomicName id="33B6F24F11467C066FCE940FA307FBCA" authorityName="Huang, Shih &amp; Ahyong" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
<emphasis id="92C2B0158866E9CC13283D5F555259D2" italics="true">Megapleonum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A973CFD70AB3195EC1D104342D68C9B6">
<table id="TID0E2SBG" rules="all">
<tbody id="399EA8F53140D9B65CDC2E43793A6FC1">
<tr id="6AEBECF6285D964B132E5A2904D20AAE">
<th id="0CB4A3BE255C43268C0662498118A228" colspan="1" rowspan="1">Character</th>
<th id="F02669309B7FAC0467969CAD834C4E92" colspan="1" rowspan="1">
<taxonomicName id="AA90A8FD27787ED68DDA3A35B8BF4DB2" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="falx" status="sp. nov.">
<emphasis id="B6B6666E8FC8DF5A34C3035C606BB78D" italics="true">Megapleonum falx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD30954D6DBCFC4A70BE2E3D3549CB53" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="236D5F121CB7742EEB288E259A1F34C5" colspan="1" rowspan="1">
<taxonomicName id="0C2381E439C2274DAFFB4EBDF1A913AB" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ferrumequinum" status="sp. nov.">
<emphasis id="93289083B16FD3F028933BE1CA635033" italics="true">Megapleonum ferrumequinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E5E70CA81EC6A5A3031FE53F8F6E5B6" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="12332F79855A494B30DED0640B495F07" colspan="1" rowspan="1">
<taxonomicName id="039DB568E0D61F10717ABC2C9F0F9625" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">
<emphasis id="E2CAC108E18283CF6B3F375FB19BD006" italics="true">Megapleonum wangjiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1513F5F0D8037AD7A1410EBC8F1F26D" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="43D7FA7D8D3247A8EB5A275EA3D4612C" colspan="1" rowspan="1">
<taxonomicName id="AF07F07C0F66C4BA09AE8CDB3E7FBF65" authorityName="Huang &amp; Shih &amp; Ahyong" authorityYear="2025" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="yangdongense" status="sp. nov.">
<emphasis id="F770306E635AA8D2D952490FDABC8D34" italics="true">Megapleonum yangdongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AD032A29FA98DDD9F0110B343E5D46B3" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<emphasis id="E222E56CFAA87A8661037AA5C09D16FA" italics="true">
<th id="D0ECFC4AB06DB55A0206FB57DF5E6720" colspan="1" rowspan="1">
<taxonomicName id="AB82F38028E8FF83D531E51600F6B7FB" authorityName="Huang, Shih &amp; Ahyong" authorityYear="2018" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="ehuangzhang">Megapleonum ehuangzhang</taxonomicName>
</th>
<th id="4A78888A181C4B965F7ACBF03B581C5B" colspan="1" rowspan="1">
<taxonomicName id="82FEF7712C3BE1772F7EAE7892ABC52E" authorityName="Huang &amp; Mao" authorityYear="2021" class="Malacostraca" family="Potamidae" genus="Megapleonum" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="shenzhen">Megapleonum shenzhen</taxonomicName>
</th>
</emphasis>
</tr>
<tr id="DB196DC35F5A7091F15B5BD87570ABA0">
<td id="53D0672507780EFC7E9AC74211ECBB81" colspan="1" rowspan="1">Carapace</td>
<td id="85A36AA3F7BCF3CA10303CC96DEC4204" colspan="1" rowspan="1">
Dense long setae at lateral margins, epigastric cristae and postorbital cristae confluent (Fig.
<figureCitation id="FF2544F35573CE521BCE13392D85421C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
)
</td>
<td id="D2118BE409F9604CB1787E1B13D0C806" colspan="1" rowspan="1">
Dense long setae at lateral margins, epigastric cristae and postorbital cristae almost confluent (Fig.
<figureCitation id="5648C48D2EBBF663FCFF621191FC5D86" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
)
</td>
<td id="1A1AAF75280B5D3352DD5A191F7E9122" colspan="1" rowspan="1">
Sparse short setae at lateral margins, epigastric cristae and postorbital cristae almost confluent (Fig.
<figureCitation id="3A271B0E57E555A3198FAE15748E8D5A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
)
</td>
<td id="CC289AD5D8A90E40B94BE9AA7A1BADBD" colspan="1" rowspan="1">
No setae at lateral margins, epigastric cristae and postorbital cristae confluent (Fig.
<figureCitation id="65EEE2EA788D49204EB9DC92474EBA8F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
)
</td>
<td id="C6C28B354238C80C84E91C35DBA6298E" colspan="1" rowspan="1">
No setae at lateral margins, epigastric cristae and postorbital cristae confluent (
<bibRefCitation id="6282BC8B9EF2DC98153D5423BF104184" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 2)
</td>
<td id="1887834E4301FF494A82BF9E9A0BBCD5" colspan="1" rowspan="1">
No setae at lateral margins, epigastric cristae and postorbital cristae confluent (
<bibRefCitation id="44EC1F49C65418EEE8585EC0BFA5F6F1" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 1)
</td>
</tr>
<tr id="E936880B630BCFF746FF3BCF106C698C">
<td id="4BAEA19673B3AC88CE4CAAB83681A579" colspan="1" rowspan="1">Maxilliped 3 exopod</td>
<td id="8346F9AE087D7583DF133E073B8358A4" colspan="1" rowspan="1">
With long flagellum (Fig.
<figureCitation id="B1131A42590F03B64118690B9F34E686" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 A</figureCitation>
)
</td>
<td id="8044CAE846150076E57E29D43B3CAC34" colspan="1" rowspan="1">
With no flagellum (Fig.
<figureCitation id="0F2D3CB710EED20F4B1DC49B9FEE5611" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 A</figureCitation>
)
</td>
<td id="61694CA1D31F69E8C746E8468939E1CD" colspan="1" rowspan="1">
With no flagellum (Fig.
<figureCitation id="C60EE643F5ECFEB7DA0005117B40547A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 A</figureCitation>
)
</td>
<td id="1A1445324BEC2954AAB944C63F9D60B9" colspan="1" rowspan="1">
With short flagellum (Fig.
<figureCitation id="0F1AEC06BAA5A9B4F23B165795839DA7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 A</figureCitation>
)
</td>
<td id="E6AED81C0A644A3A20F5BBCBAAE57439" colspan="1" rowspan="1">
With no flagellum (
<bibRefCitation id="07572D819F39B24EE33A7C33E1B3053B" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 3 D)
</td>
<td id="0D52F107A39485A868F09A002DAA3320" colspan="1" rowspan="1">
With vestigial flagellum (
<bibRefCitation id="FFE897010F0E9367D037562D0C06D6D6" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 3 A)
</td>
</tr>
<tr id="A2F3C05A6F39892A99D9419D0CBE7F40">
<td id="B17A6B740D302E04F42A0D14FB88B4D5" colspan="1" rowspan="1">Ambulatory legs</td>
<td id="03179D9F9FDEA814027DDBD6087C76B1" colspan="1" rowspan="1">
With dense long setae (Fig.
<figureCitation id="F1F5F79CBE350068CA3B15912B8B9010" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure1" httpUri="https://binary.pensoft.net/fig/1366338">1 A</figureCitation>
)
</td>
<td id="CCB43F2E84DDDE12AAE8BEF687A7404D" colspan="1" rowspan="1">
With dense long setae, particularly pereiopods 2 and 3 (Fig.
<figureCitation id="17ABAADA25D60624B70A0A036FB1260E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Dorsal habitus (A); cephalothorax, anterior view (B); anterior thoracic sternum (C); anterior thoracic sternum and pleon, ventral view (D); sterno-pleonal cavity with G 1 in situ, ventral view (E)." figureDoi="10.3897/zookeys.1244.148112.figure3" httpUri="https://binary.pensoft.net/fig/1366340">3 A</figureCitation>
)
</td>
<td id="BF258201BE7F3AAF4610DB3BB9E1D140" colspan="1" rowspan="1">
With dense long setae, particularly pereiopods 2 and 3 (Fig.
<figureCitation id="0862DE4054BEA160DCBDE2A7C1C18299" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal habitus. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A); female paratype (16.7 × 12.8 mm), SYSBM 002148 (B)." figureDoi="10.3897/zookeys.1244.148112.figure5" httpUri="https://binary.pensoft.net/fig/1366342">5</figureCitation>
)
</td>
<td id="04B2B145CD077680AC14635AB567FB21" colspan="1" rowspan="1">
With short setae (Fig.
<figureCitation id="9EC415930AE83402D0C2FC95B3DADB4D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal habitus. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A); female paratype (16.0 × 12.4 mm), SYSBM 002154 (B)." figureDoi="10.3897/zookeys.1244.148112.figure8" httpUri="https://binary.pensoft.net/fig/1366345">8</figureCitation>
)
</td>
<td id="1ECDC73FEECC31322B6A132ACBEF044A" colspan="1" rowspan="1">
With short setae (
<bibRefCitation id="6D4CA1F092DE82A721ECDD2FA9692114" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 2)
</td>
<td id="CE53EA19C2950F80D90BEBCAD1F3C167" colspan="1" rowspan="1">
With short setae (
<bibRefCitation id="A94FBBBAB271C5F672BE610CDD1E5BCA" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 1)
</td>
</tr>
<tr id="0E613D5CD0F875F7FE53DC31F2DF9AA7">
<td id="BBBA27DB0112EF478774F081F6D85567" colspan="1" rowspan="1">Male anterior thoracic sternum width length ratio</td>
<td id="A5647ECA9104F936CC76CFB39CB2C59D" colspan="1" rowspan="1">1.8</td>
<td id="8F48E7EC0B88FB4AC3E351A27A7131F5" colspan="1" rowspan="1">1.9</td>
<td id="9032E82C1B3B71A924440F5B187ACFDB" colspan="1" rowspan="1">2.0</td>
<td id="A997D4A42108F7E637CF16C4528047E8" colspan="1" rowspan="1">1.9</td>
<td id="285123F5841FC51B82536B5896BEE495" colspan="1" rowspan="1">
1.9 (
<bibRefCitation id="23493C538A7DADE4FC6FA8B81B83D661" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
)
</td>
<td id="3E9240F3C2ED29A084372E2841801F1D" colspan="1" rowspan="1">
1.9 (
<bibRefCitation id="A3FDD0D64804FADDC21755692537CDA3" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
)
</td>
</tr>
<tr id="406D4C9DDA76B908695748342590653F">
<td id="0BF29963D2A0EA01804562ABA9551BC9" colspan="1" rowspan="1">Male pleonite 6 width to length ratio</td>
<td id="F85C0149EF2957C58A1C3ED5575DACEF" colspan="1" rowspan="1">2.6</td>
<td id="410FC1056BA866B828A258117548CFFE" colspan="1" rowspan="1">2.5</td>
<td id="B202FA23433564B686CFA1EAFFAD42DC" colspan="1" rowspan="1">2.6</td>
<td id="676AC54658553391C9096251F7044366" colspan="1" rowspan="1">2.4</td>
<td id="62531B97C44F372D496D9A0A5547C701" colspan="1" rowspan="1">
2.6 (
<bibRefCitation id="D78776D0F6FD6932C7CAE3B335C2B88C" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
)
</td>
<td id="ABD844D78C1B1D55A398E33D35252476" colspan="1" rowspan="1">
2.6 (
<bibRefCitation id="C7C54E9FD0E5F2C7CFF3F564C88C15BE" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
)
</td>
</tr>
<tr id="C664D25B6A2043D4CF07F7E239443E9C">
<td id="1ADCE2C9C9D1300572D317230F82D4AD" colspan="1" rowspan="1">Male telson width to length ratio</td>
<td id="C300596837657A60F8B736D8A7150401" colspan="1" rowspan="1">1.8</td>
<td id="8F5AA1A76DC18FB4488C4A47B75E8E83" colspan="1" rowspan="1">1.7</td>
<td id="8227BF24D1409551577E330858BF9733" colspan="1" rowspan="1">1.8</td>
<td id="F8BFCB8A457D2CF9A9E9A281F96AA133" colspan="1" rowspan="1">1.8</td>
<td id="513743F9B6EFD3F9B41EBB77685C6BC0" colspan="1" rowspan="1">
1.7 (
<bibRefCitation id="1BA57C83BED42E2816B45F8835C365EF" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
)
</td>
<td id="D0696C898504114FD5BA7770C50B8808" colspan="1" rowspan="1">
2.0 (
<bibRefCitation id="7DF2F4BA5C80CF128920ED9D19143409" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
)
</td>
</tr>
<tr id="067BC50427582B7194888BA5471750BB">
<td id="F3DC817335CD53002B492A5EC67C9CEE" colspan="1" rowspan="1">G 1 terminal segment</td>
<td id="75EECDC911EBC906830AC4987F111E5B" colspan="1" rowspan="1">
Strongly tapering, sickle shaped (Fig.
<figureCitation id="AC99305F6329A293BC9D7EF64345C44F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 F, G</figureCitation>
)
</td>
<td id="6E9E8970C3B8DFD6D62EF4A4E393B59D" colspan="1" rowspan="1">
Long and curved with horseshoe shaped structure (Fig.
<figureCitation id="BFC4F3FD0E58DEB7254F78F6134DB9AB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 F H</figureCitation>
)
</td>
<td id="E474E6C49AF0DCF2C2417BF1A07A7BD0" colspan="1" rowspan="1">
Short, rounded, distal-ventral region with long setae, tip presenting as protrusion on outer margin (Fig.
<figureCitation id="9D3FBFB6D690652552261A4CB5717522" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 A, B</figureCitation>
)
</td>
<td id="F17BB77EB7354BFBCE10F5CC17365D8B" colspan="1" rowspan="1">
Short, bifurcated, prong shaped (Fig.
<figureCitation id="758E97E93F9DFB03858CDED245955F66" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Left G 1 terminal segments. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A, B); Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (C, D). Ventral view (A, C); dorsal view (B, D). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1244.148112.figure11" httpUri="https://binary.pensoft.net/fig/1366348">11 C, D</figureCitation>
)
</td>
<td id="94DDB5EF0A701F81426DC5EEEB759405" colspan="1" rowspan="1">
Bent outwards and folded inwards with large flap on ventral side (
<bibRefCitation id="A0ECE7F427005B91E29553C9889C1921" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 3 C)
</td>
<td id="2E3A1A9F90C7BC809AA0815AB023C578" colspan="1" rowspan="1">
Stout, goose-head-shaped (
<bibRefCitation id="D3CCA1658A06B1B693C4BD22E40543D3" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 4 G)
</td>
</tr>
<tr id="BEC60FBD06DC86C6BB9DD0C057FCA529">
<td id="59A12987259939D39A26FE4497B9A6FE" colspan="1" rowspan="1">G 2 subterminal segment proximal region</td>
<td id="F6DBEC09F08E89CB1CAEA0333A798C72" colspan="1" rowspan="1">
Small (Fig.
<figureCitation id="F74E00F5AE82D3967AAAAA4D11F248F7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megapleonum falx sp. nov., male holotype (19.6 × 15.8 mm), SYSBM 002142. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped, partially damaged (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 terminal segment, dorsal view (G). Scale bars: 1.0 mm (A C, F, G), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure2" httpUri="https://binary.pensoft.net/fig/1366339">2 B</figureCitation>
)
</td>
<td id="65E0B1DA74FB1CE27297DD8ED6939CCF" colspan="1" rowspan="1">
Small (Fig.
<figureCitation id="07428240A632EB898DAA43E54C002C87" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megapleonum ferrumequinum sp. nov., male holotype (16.6 × 13.1 mm), SYSBM 002143. Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C); major cheliped (D); minor cheliped (E); left G 1 terminal segment, ventral view (F); left G 1 horse-shoe structure viewed from the top (G); left G 1 terminal segment, dorsal view (H). Scale bars: 1.0 mm (A C, F H), 5.0 mm (D, E)." figureDoi="10.3897/zookeys.1244.148112.figure4" httpUri="https://binary.pensoft.net/fig/1366341">4 B</figureCitation>
)
</td>
<td id="ECAFDAE4E38C4FFFBDC2A09C8B60DB7C" colspan="1" rowspan="1">
Very small (Fig.
<figureCitation id="2E64F5CF20F6EB93D04A322130B30B42" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002145 (D); male paratype (10.9 × 9.0 mm), SYSBM 002146 (E); female paratype (16.7 × 12.8 mm), SYSBM 00 2148 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure7" httpUri="https://binary.pensoft.net/fig/1366344">7 B</figureCitation>
)
</td>
<td id="BD0704EBDD451B66A28AAE4C4BF2F18E" colspan="1" rowspan="1">
Large (Fig.
<figureCitation id="AAAB831A1F83BC3B93DE01E32FD6F7A4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A C, F, G); male paratype (11.6 × 9.5 mm), SYSBM 002153 (D); male paratype (16.0 × 12.7 mm), IHB (E); female paratype (16.0 × 12.4 mm), SYSBM 002154 (H, I). Left maxilliped 3 (A); left G 2, pleonal view (B); left G 1, ventral view (C E); major cheliped (F, H); minor cheliped (G, I). Scale bars: 1.0 mm (A E), 5.0 mm (F I)." figureDoi="10.3897/zookeys.1244.148112.figure10" httpUri="https://binary.pensoft.net/fig/1366347">10 B</figureCitation>
)
</td>
<td id="E979EC1C17CB5303FD06094BDD1F7CAD" colspan="1" rowspan="1">
Large (
<bibRefCitation id="65F90CDC73CDE6FECF226257526E4B5B" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 3 A)
</td>
<td id="3F9A4D6661C35738D9B10AA8FD6C6F62" colspan="1" rowspan="1">
Large (
<bibRefCitation id="CAC76BB56995CE7C4B1C1755BE9953E3" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 3 B)
</td>
</tr>
<tr id="5E87E87FC4BD45FD4BD06EC43D3646FD">
<td id="5F96FF6EB202DDFAF82F13E9C1465668" colspan="1" rowspan="1">Female pleon</td>
<td id="4724284EB072A185F13C9FF61153B1D7" colspan="1" rowspan="1">Ovate</td>
<td id="D08001CC7B5439B3EE6CAE42FF6980E6" colspan="1" rowspan="1">Ovate</td>
<td id="91EEC3C7E69F30BC4E645CAE24DB339D" colspan="1" rowspan="1">
Ovate (Fig.
<figureCitation id="77950DF9CDDCFA44B4E820786DDFB30C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megapleonum wangjiani sp. nov., male holotype (11.7 × 9.7 mm), SYSBM 002144 (A D); female paratype (16.7 × 12.8 mm), SYSBM 002148 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure6" httpUri="https://binary.pensoft.net/fig/1366343">6 E</figureCitation>
)
</td>
<td id="654791985B95E0E0C107009C3D0B1331" colspan="1" rowspan="1">
Ovate (Fig.
<figureCitation id="92CD034F85601652F6361CF13BAAD320" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Megapleonum yangdongense sp. nov., male holotype (17.8 × 14.5 mm), SYSBM 002152 (A D); female paratype (16.0 × 12.4 mm), SYSBM 002154 (E, F). Cephalothorax, anterior view (A); anterior thoracic sternum (B); anterior thoracic sternum and pleon, ventral view (C); sterno-pleonal cavity with G 1 in situ, ventral view (D); pleon, ventral view (E); vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1244.148112.figure9" httpUri="https://binary.pensoft.net/fig/1366346">9 E</figureCitation>
)
</td>
<td id="D43E8B804D5801612D93AD1C60A67DC0" colspan="1" rowspan="1">
Linguiform (
<bibRefCitation id="1ED393432A0F77B31E43605E7C0C5913" DOI="10.1093/jcbiol/ruy050" author="Huang" etAl="et al." firstAuthor="Huang" issue="5" journalOrPublisher="Journal of Crustacean Biology" pagination="614-624" refId="B10" refString="Huang C, Shih HT, Ahyong ST (2018) Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae). Journal of Crustacean Biology 38 (5): 614624. https://doi.org/10.1093/jcbiol/ruy050" title="Two new genera and two new species of narrow-range freshwater crabs from Guangdong, China (Decapoda: Brachyura: Potamidae)." volume="38" year="2018">Huang et al. 2018</bibRefCitation>
: fig. 4 C)
</td>
<td id="A42FAE36F383494FA7CB4A85689055C5" colspan="1" rowspan="1">
Linguiform (
<bibRefCitation id="9CCFECF8BB9FCAD9058ECF30ABCD4DD2" DOI="10.6620/ZS.2021.60-66" author="Huang and Mao" firstAuthor="Huang" refId="B8" refString="Huang C, Mao SY (2021) The hillstream decapod crustaceans of Shenzhen, China, with description of a new species of freshwater crab (Crustacea: Brachyura: Potamidae) in the genus Megapleonum Huang, Shih &amp; Ahyong, 2018. Zoological Studies (Taipei, Taiwan) 60: 66. https://doi.org/10.6620/ZS.2021.60-66" year="2021">Huang and Mao 2021</bibRefCitation>
: fig. 2 E)
</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>