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<mods:title id="85F03415EC6A09E05122FED8494149BA">Cryptic speciation within Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareatidae), the description of a new species from Peninsular Malaysia, and the resurrection of A. tropidonotus (Lidth de Jude, 1923)</mods:title>
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<mods:namePart id="064FF20ADC039725F5CB19DCC96BD196">Loredo, Ariel I.</mods:namePart>
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<mods:affiliation id="FF214269F31F670CC7D383F06F231B9E">Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California, 92515 - 8247 USA. & School of Veterinary Medicine, University of California, Davis, One Shields Avenue, Davis, CA 95616 USA.</mods:affiliation>
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<mods:namePart id="6C33CCA20A340997604F544A41CC7949">Wood, Perry L.</mods:namePart>
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<mods:affiliation id="4411B7C98AA9240EF149BDB445CC3BA0">Department of Biology, Brigham Young University, 150 East Bulldog Boulevard, Provo, Utah 84602 USA. E-mail: pwood @ byu. edu. School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Penang, Malaysia.</mods:affiliation>
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<mods:namePart id="66E3CC58C2DB4651BA6D73D14EF6ACD1">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="1B7A5858B964CDE2DCD96B94B22324F9">Anuar, Shahrul</mods:namePart>
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<mods:namePart id="330FE40B7ECAB796EE70FDE1C0EFCD82">Greer, Lee F.</mods:namePart>
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<mods:affiliation id="C0128072ABC34D3E5EA5E234F4D4254E">Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California, 92515 - 8247 USA.</mods:affiliation>
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<mods:namePart id="3C5CEF9A6C13B442BFBAF5198C990855">Ahmad, Norhayati</mods:namePart>
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<mods:affiliation id="67E16B5086BA6033FB9E3632E4982DD9">Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor Darul Ehsan, Malaysia. E-mail: yati _ 68 @ yahoo. co. uk</mods:affiliation>
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<mods:namePart id="06CCFE183C38D13CC4C43293127FBAFD">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="01C7687C163C29F4533AF6ADF3FD88C5">Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California, 92515 - 8247 USA. & Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor Darul Ehsan, Malaysia. E-mail: yati _ 68 @ yahoo. co. uk</mods:affiliation>
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<mods:title id="1FEE805CC41711F875965C06B9A7AAA9">Zootaxa</mods:title>
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<mods:part id="D7948C5138E81E6BA4E402FCD29F84CE">
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<mods:date id="AE265DB65F501A7C00D669FCA89BE90C">2013</mods:date>
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<mods:number id="A1C15133D4B31EE506DD95A4C9833389">2013-05-28</mods:number>
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<mods:number id="60DB307E08A107C68D40B76D4A40161D">3664</mods:number>
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<mods:identifier id="373C7CCFB71028E6894C7D3F78900A47" type="DOI">10.11646/zootaxa.3664.4.5</mods:identifier>
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<mods:identifier id="BC74C819693D6FCD00E4FD21D773ABAF" type="ISSN">1175-5326</mods:identifier>
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<paragraph id="4D7A3680DA0CFFFAFF19FCB2FDA20C07" blockId="9.[151,590,773,868]" box="[151,590,773,799]" pageId="9" pageNumber="514">
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<taxonomicName id="8AC54D03DA0CFFFAFF19FCB2FE010C06" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[151,493,773,799]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="514" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
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<emphasis id="7FB1EA92DA0CFFFAFF19FCB2FE010C06" bold="true" box="[151,493,773,799]" italics="true" pageId="9" pageNumber="514">Asthenodipsas lasgalenensis</emphasis>
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</taxonomicName>
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<emphasis id="7FB1EA92DA0CFFFAFE7BFCB1FDA20C07" bold="true" box="[501,590,774,798]" pageId="9" pageNumber="514">
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<taxonomicNameLabel id="648257E9DA0CFFFAFE7BFCB1FDA20C07" box="[501,590,774,798]" pageId="9" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="05DF650BDA0CFFFAFF19FC90FE330C26" box="[151,479,807,831]" pageId="9" pageNumber="514" type="vernacular_names">
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<paragraph id="4D7A3680DA0CFFFAFF19FC90FE330C26" blockId="9.[151,590,773,868]" box="[151,479,807,831]" pageId="9" pageNumber="514">Mirkwood Forest Slug Snake</paragraph>
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||||
</subSubSection>
|
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<subSubSection id="05DF650BDA0CFFFAFF19FCFBFEF90C7D" box="[151,277,844,868]" pageId="9" pageNumber="514" type="description">
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<paragraph id="4D7A3680DA0CFFFAFF19FCFBFEF90C7D" blockId="9.[151,590,773,868]" box="[151,277,844,868]" pageId="9" pageNumber="514">
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<figureCitation id="D5FE2A05DA0CFFFAFF19FCFBFEEE0C7D" box="[151,258,844,868]" captionStart="FIGURE 2" captionStartId="10.[151,250,1964,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure-50@10.[151,1436,193,1943]" captionTargetPageId="10" captionText="FIGURE 2. Upper. Adult male Asthenodipsas lasgalenensis sp. nov. from Bukit Larut, Perak (holotype LSUHC 8869). Lower. Adult female A. lasgalenensis sp. nov. from Cameron Highlands, Pahang (paratype LSUHC 6954)." figureDoi="http://doi.org/10.5281/zenodo.5266003" httpUri="https://zenodo.org/record/5266003/files/figure.png" pageId="9" pageNumber="514">Figures 2</figureCitation>
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||||
,
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||||
<figureCitation id="D5FE2A05DA0CFFFAFE89FCFBFEF90C7D" box="[263,277,844,868]" captionStart="FIGURE 3" captionStartId="11.[151,250,1055,1078]" captionTargetBox="[151,1436,193,1034]" captionTargetId="figure-429@11.[151,1436,193,1034]" captionTargetPageId="11" captionText="FIGURE 3. Coloration of a hatchling Asthenodipsas vertebralis from Fraser’s Hill, Pahang (upper left LSUHC 9100) and a subadult female from Bukit Larut, Perak lower right LSUHC 9837. Coloration of hatchling A. lasgalenensis from Bukit Larut, Perak (LSUHC 10798, upper right and LSUHC 10797, lower left)." figureDoi="http://doi.org/10.5281/zenodo.5266007" httpUri="https://zenodo.org/record/5266007/files/figure.png" pageId="9" pageNumber="514">3</figureCitation>
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</paragraph>
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<subSubSection id="05DF650BDA0CFFFAFF19FC38FCDA0B06" pageId="9" pageNumber="514" type="reference_group">
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<paragraph id="4D7A3680DA0CFFFAFF19FC38FDC50CF8" blockId="9.[151,1436,910,1055]" pageId="9" pageNumber="514">
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<taxonomicName id="8AC54D03DA0CFFFAFF19FC38FDF50CBD" authority="Smedley 1931: 122" authorityName="Smedley" authorityPageNumber="122" authorityYear="1931" box="[151,537,910,932]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="514" phylum="Chordata" rank="species" species="vertebralis">
|
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<emphasis id="7FB1EA92DA0CFFFAFF19FC38FEBD0CBD" box="[151,337,910,932]" italics="true" pageId="9" pageNumber="514">Pareas vertebralis</emphasis>
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<bibRefCitation id="29544B71DA0CFFFAFED6FC39FDF50CBD" author="Smedley, N." box="[344,537,910,932]" pageId="9" pageNumber="514" pagination="105 - 123" refId="ref11103" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum of Singapore, 6, 105 - 123." type="journal article" year="1931">Smedley 1931:122</bibRefCitation>
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</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="29544B71DA0CFFFAFDAAFC39FD3F0CBD" author="Tweedie, M. W. F." box="[548,723,910,932]" pageId="9" pageNumber="514" refId="ref11394" refString="Tweedie, M. W. F. (1953) The Snakes of Malaya. Government Printing Office, Singapore, 139 pp." type="book" year="1953">Tweedie 1953:34</bibRefCitation>
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(in part), 1957:35 (in part);
|
||||
<bibRefCitation id="29544B71DA0CFFFAFC7FFC39FB6F0CBD" author="Sly, G. R." box="[1009,1155,910,932]" pageId="9" pageNumber="514" pagination="155 - 157" refId="ref11070" refString="Sly, G. R. (1976) New locality records for some peninsular Amphibia, Reptilia, and Mammalia. The Malayan Nature Journal, 29, 155 - 157." type="journal article" year="1976">Sly 1976: 156</bibRefCitation>
|
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; Tweedie 1987:36 (in part);
|
||||
<bibRefCitation id="29544B71DA0CFFFAFF49FC19FE2A0CDA" author="Rao, D. - Q. & Yang, D. - T." box="[199,454,941,963]" pageId="9" pageNumber="514" pagination="139 - 150" refId="ref10930" refString="Rao, D. - Q. & Yang, D. - T. (1972) Phylogenetic systematics of Parinae (Serpents) of Southeastern Asia and adjacent islands with relationship between it and geology changes. Acta Zoologica Sinica, 38 (2), 139 - 150." type="journal article" year="1972">Rao and Yang 1972:144</bibRefCitation>
|
||||
; Lim
|
||||
<emphasis id="7FB1EA92DA0CFFFAFD89FC19FDD10CDA" box="[519,573,941,963]" italics="true" pageId="9" pageNumber="514">et al.</emphasis>
|
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2002:55; Manthey and Grossmann 1967:378;
|
||||
<bibRefCitation id="29544B71DA0CFFFAFBABFC1AFB010CDA" author="Cox, M. J. & Van Dijk, P. P. & Nabhitabhata, J. & Thirakhupt, K." box="[1061,1261,941,963]" pageId="9" pageNumber="514" refId="ref9124" refString="Cox, M. J., Van Dijk, P. P., Nabhitabhata, J. & Thirakhupt, K. (1998) A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. New Holland Publishers, London, England, 144 pp." type="book" year="1998">
|
||||
Cox
|
||||
<emphasis id="7FB1EA92DA0CFFFAFBD7FC19FB630CDA" box="[1113,1167,941,963]" italics="true" pageId="9" pageNumber="514">et al.</emphasis>
|
||||
1998:79
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="29544B71DA0CFFFAFB74FC19FEC50CF8" author="Leong, T. & Lim, K. K. P." pageId="9" pageNumber="514" pagination="123 - 136" refId="ref10563" refString="Leong, T. M & Lim, K. K. P. (2003) Herpetofaunal records from Fraser's Hill, peninsular Malaysia, with larval descriptions of Limnonectes nitidus and Theloderma asperum (Amphibia: Ranidae and Rhacophoridae). The Raffles Bulletin of Zoology, 51, 123 - 136." type="journal article" year="2003">Leong and Lim 2003:133</bibRefCitation>
|
||||
; Das & Yakkob 2007:74.
|
||||
</paragraph>
|
||||
<paragraph id="4D7A3680DA0CFFFAFF19FC5DFCDA0B06" blockId="9.[151,1436,910,1055]" pageId="9" pageNumber="514">
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<taxonomicName id="8AC54D03DA0CFFFAFF19FC5DFCEC0B19" authority="Grossman and Tillack 2003: 177" authorityName="Grossman and Tillack" authorityPageNumber="177" authorityYear="2003" box="[151,768,1002,1024]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="514" phylum="Chordata" rank="species" species="vertebralis">
|
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<emphasis id="7FB1EA92DA0CFFFAFF19FC5DFE4E0B19" box="[151,418,1002,1024]" italics="true" pageId="9" pageNumber="514">Asthenodipsas vertebralis</emphasis>
|
||||
Grossman and Tillack 2003:177
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="29544B71DA0CFFFAFC81FC5DFBCD0B19" author="Iskandar, D. T. & Colijn, E." box="[783,1057,1002,1024]" pageId="9" pageNumber="514" refId="ref10395" refString="Iskandar, D. T. & Colijn, E. (2002) A Checklist of Southeast Asian and New Guinean reptiles. Part I. Sperpentes. Biodiversity Conservation Project, Jakarta, 225 pp. http: // dx. doi. org / 10.11646 / zootaxa. 3616.3.2" type="book" year="2002">Iskandar and Colijn 2002</bibRefCitation>
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||||
; (in part);
|
||||
<bibRefCitation id="29544B71DA0CFFFAFB2CFC5CFADF0B19" author="Das, I." box="[1186,1331,1002,1024]" pageId="9" pageNumber="514" refId="ref9178" refString="Das, I. (2010) A Field Guide to the Reptiles of South-East Asia. New Holland Publishers, United Kingdom, 376 pp." type="book" year="2010">Das 2010:160</bibRefCitation>
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(in part);
|
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<bibRefCitation id="29544B71DA0CFFFAFF49FBBEFE540B06" author="Grismer, L. L. & Chan, K. O. & Grismer, J. L. & Wood, Jr., P. L. & Norhayati, A." box="[199,440,1033,1055]" pageId="9" pageNumber="514" pagination="147 - 160" refId="ref9715" refString="Grismer, L. L., Chan, K. O., Grismer, J. L., Wood, Jr., P. L. & Norhayati, A. (2010) A checklist of the herpetofauna of the Banjaran Bintang, Peninsular Malaysia. Russian Journal of Herpetology, 17, 147 - 160." type="journal article" year="2010">
|
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Grismer
|
||||
<emphasis id="7FB1EA92DA0CFFFAFEAEFBBDFEBF0B06" box="[288,339,1033,1055]" italics="true" pageId="9" pageNumber="514">et al.</emphasis>
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2010:155
|
||||
</bibRefCitation>
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(in part);
|
||||
<bibRefCitation id="29544B71DA0CFFFAFD90FBBEFD3B0B06" author="Grismer, L. L. & Grismer, J. L. & Wood, P. L. Jr. & Ngo, V. T. & Neang, T. & Chan, K. O." box="[542,727,1033,1055]" pageId="9" pageNumber="514" pagination="57 - 97" refId="ref9857" refString="Grismer, L. L., Grismer, J. L., Wood, P. L. Jr., Ngo, V. T., Neang, T. & Chan, K. O. (2011) Herpetology on the fringes of the Sunda Shelf: A Discussion of discovery, taxonomy, and biogeography. Bonner Zoologische Monographien, 57, 57 - 97." type="journal article" year="2011">Grismer 2011:170</bibRefCitation>
|
||||
(in part).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0CFFFAFF19FBF0FAE40A43" pageId="9" pageNumber="514" type="materials_examined">
|
||||
<paragraph id="4D7A3680DA0CFFFAFF19FBF0FB690B9B" blockId="9.[151,1436,1094,2024]" pageId="9" pageNumber="514">
|
||||
<materialsCitation id="FDAD3CDDDA0CFFFAFF19FBF0FB690B9B" collectingDate="2008-03-02" collectorName="L. Lee Grismer & Lee Grismer, P. L & Wood Jr., J. L & C. K. Onn" country="Malaysia" elevation="1184" latitude="4.7432666" location="Bukit Larut" longLatPrecision="1" longitude="100.75895" pageId="9" pageNumber="514" specimenCount="1" stateProvince="Perak" typeStatus="holotype">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF19FBF0FEE70B46" bold="true" box="[151,267,1095,1119]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
|
||||
<typeStatus id="927E8822DA0CFFFAFF19FBF0FEEA0B46" box="[151,262,1095,1119]" pageId="9" pageNumber="514" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
<collectorName id="E0305356DA0CFFFAFE9CFBF1FEBE0B47" box="[274,338,1094,1118]" pageId="9" pageNumber="514">Adult</collectorName>
|
||||
male, LSUHC 8869 collected by
|
||||
<collectorName id="E0305356DA0CFFFAFD41FBF0FC910B47" box="[719,893,1094,1118]" pageId="9" pageNumber="514">L. Lee Grismer</collectorName>
|
||||
, P. L.
|
||||
<collectorName id="E0305356DA0CFFFAFC4CFBF1FB8D0B47" box="[962,1121,1094,1118]" pageId="9" pageNumber="514">Wood Jr., J. L</collectorName>
|
||||
. Grismer, and
|
||||
<collectorName id="E0305356DA0CFFFAFA8BFBF1FA940B47" box="[1285,1400,1094,1118]" pageId="9" pageNumber="514">C. K. Onn</collectorName>
|
||||
on
|
||||
<date id="397B1040DA0CFFFAFF19FBDDFED80B9B" box="[151,308,1130,1154]" pageId="9" pageNumber="514" value="2008-03-02">
|
||||
<collectingDate id="293FE9A8DA0CFFFAFF19FBDDFED80B9B" box="[151,308,1130,1154]" pageId="9" pageNumber="514" value="2008-03-02">2 March 2008</collectingDate>
|
||||
</date>
|
||||
from
|
||||
<location id="481A605BDA0CFFFAFEF6FBDDFE100B9B" LSID="urn:lsid:plazi:treatment:C56C8796DA0CFFFFFF19FCB2FB910DB7:481A605BDA0CFFFAFEF6FBDDFE100B9B" box="[376,508,1130,1154]" country="Malaysia" latitude="4.7432666" longLatPrecision="1" longitude="100.75895" name="Bukit Larut" pageId="9" pageNumber="514" stateProvince="Perak">Bukit Larut</location>
|
||||
,
|
||||
<collectingRegion id="8F01F862DA0CFFFAFD86FBDDFDA70B9B" box="[520,587,1130,1154]" country="Malaysia" name="Perak" pageId="9" pageNumber="514">Perak</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="35D27610DA0CFFFAFDD8FBDDFD510B9B" box="[598,701,1130,1154]" name="Malaysia" pageId="9" pageNumber="514">Malaysia</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="28F15047DA0CFFFAFD45FBDDFCB30B9B" box="[715,863,1130,1154]" degrees="4" direction="north" minutes="44.596" orientation="latitude" pageId="9" pageNumber="514" precision="1" value="4.7432666">4º 44.596’ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="28F15047DA0CFFFAFCE4FBDDFBF90B9B" box="[874,1045,1130,1154]" degrees="100" direction="east" minutes="45.537" orientation="longitude" pageId="9" pageNumber="514" precision="1" value="100.75895">100º 45.537’ E</geoCoordinate>
|
||||
;
|
||||
<quantity id="8A3D9B65DA0CFFFAFBAFFBDDFB950B9B" box="[1057,1145,1130,1154]" metricMagnitude="3" metricUnit="m" metricValue="1.184" pageId="9" pageNumber="514" unit="m" value="1184.0">
|
||||
<elevation id="C6E8D1B3DA0CFFFAFBAFFBDDFB950B9B" box="[1057,1145,1130,1154]" metricMagnitude="3" metricUnit="m" metricValue="1.184" pageId="9" pageNumber="514" unit="m" value="1184.0">1184 m</elevation>
|
||||
</quantity>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="4D7A3680DA0CFFFAFF47FB38FAE40A43" blockId="9.[151,1436,1094,2024]" pageId="9" pageNumber="514">
|
||||
<materialsCitation id="FDAD3CDDDA0CFFFAFF47FB38FBDB0BF7" collectingDate="2008-06-06" collectorName="L. Lee Grismer & Lee Grismer, P. L & Wood Jr., J. L & C. K. Onn" country="Malaysia" county="West" elevation="1203" latitude="3.7205667" location="Pine Tree Trail" longLatPrecision="1" longitude="101.757965" municipality="Fraser's Hill" pageId="9" pageNumber="514" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47FB38FEA40BBE" bold="true" box="[201,328,1167,1191]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
|
||||
<typeStatus id="927E8822DA0CFFFAFF47FB38FEAF0BBE" box="[201,323,1167,1191]" pageId="9" pageNumber="514" type="paratype">Paratypes</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
<collectorName id="E0305356DA0CFFFAFEDCFB39FE7E0BBF" box="[338,402,1166,1190]" pageId="9" pageNumber="514">Adult</collectorName>
|
||||
female (LSUHC 9050) bears the same data as the holotype.
|
||||
<collectorName id="E0305356DA0CFFFAFBEEFB39FB4C0BBF" box="[1120,1184,1166,1190]" pageId="9" pageNumber="514">Adult</collectorName>
|
||||
male (LSUHC 9098) collected by
|
||||
<collectorName id="E0305356DA0CFFFAFEA4FB04FE300BD3" box="[298,476,1202,1226]" pageId="9" pageNumber="514">L. Lee Grismer</collectorName>
|
||||
, P. L.
|
||||
<collectorName id="E0305356DA0CFFFAFDA9FB05FD270BD3" box="[551,715,1202,1226]" pageId="9" pageNumber="514">Wood Jr., J. L</collectorName>
|
||||
. Grismer, and
|
||||
<collectorName id="E0305356DA0CFFFAFCFAFB05FC070BD3" box="[884,1003,1202,1226]" pageId="9" pageNumber="514">C. K. Onn</collectorName>
|
||||
on
|
||||
<date id="397B1040DA0CFFFAFB94FB05FB490BD3" box="[1050,1189,1202,1226]" pageId="9" pageNumber="514" value="2008-06-06">
|
||||
<collectingDate id="293FE9A8DA0CFFFAFB94FB05FB490BD3" box="[1050,1189,1202,1226]" pageId="9" pageNumber="514" value="2008-06-06">6 June 2008</collectingDate>
|
||||
</date>
|
||||
on
|
||||
<location id="481A605BDA0CFFFAFB5DFB05FA920BD3" LSID="urn:lsid:plazi:treatment:C56C8796DA0CFFFFFF19FCB2FB910DB7:481A605BDA0CFFFAFB5DFB05FA920BD3" box="[1235,1406,1202,1226]" country="Malaysia" county="West" latitude="3.7205667" longLatPrecision="1" longitude="101.757965" municipality="Fraser's Hill" name="Pine Tree Trail" pageId="9" pageNumber="514" stateProvince="Pahang">Pine Tree Trail</location>
|
||||
at
|
||||
<collectingMunicipality id="AD1EACFADA0CFFFAFF19FB61FEC60BF7" box="[151,298,1238,1262]" pageId="9" pageNumber="514">Fraser’s Hill</collectingMunicipality>
|
||||
,
|
||||
<collectingRegion id="8F01F862DA0CFFFAFEB5FB61FE7C0BF7" box="[315,400,1238,1262]" country="Malaysia" name="Pahang" pageId="9" pageNumber="514">Pahang</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="A41B4E0CDA0CFFFAFE2EFB60FE3B0BF7" box="[416,471,1239,1262]" pageId="9" pageNumber="514">West</collectingCounty>
|
||||
<collectingCountry id="35D27610DA0CFFFAFE6DFB61FDA50BF7" box="[483,585,1238,1262]" name="Malaysia" pageId="9" pageNumber="514">Malaysia</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="28F15047DA0CFFFAFDD0FB61FD160BF7" box="[606,762,1238,1262]" degrees="3" direction="north" minutes="43.234" orientation="latitude" pageId="9" pageNumber="514" precision="1" value="3.7205667">3º 43.234’ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="28F15047DA0CFFFAFC85FB61FC530BF7" box="[779,959,1238,1262]" degrees="101" direction="east" minutes="45.478" orientation="longitude" pageId="9" pageNumber="514" precision="1" value="101.757965">101º 45.478’ E</geoCoordinate>
|
||||
;
|
||||
<quantity id="8A3D9B65DA0CFFFAFC5EFB61FBC20BF7" box="[976,1070,1238,1262]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" pageId="9" pageNumber="514" unit="m" value="1203.0">
|
||||
<elevation id="C6E8D1B3DA0CFFFAFC5EFB61FBC20BF7" box="[976,1070,1238,1262]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" pageId="9" pageNumber="514" unit="m" value="1203.0">1203 m</elevation>
|
||||
</quantity>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="FDAD3CDDDA0CFFFAFBC8FB61FDA40A2F" collectorName="L. Lee Grismer & C. K. Onn" country="Malaysia" elevation="2050" latitude="4.52125" location="Pahang" longLatPrecision="1" longitude="101.38655" pageId="9" pageNumber="514" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
|
||||
Adult female (LSUHC 6954) collected by
|
||||
<collectorName id="E0305356DA0CFFFAFEA3FB4CFE0D0A0B" box="[301,481,1274,1298]" pageId="9" pageNumber="514">L. Lee Grismer</collectorName>
|
||||
and
|
||||
<collectorName id="E0305356DA0CFFFAFD91FB4DFD750A0B" box="[543,665,1274,1298]" pageId="9" pageNumber="514">C. K. Onn</collectorName>
|
||||
on trail 11 at Tanah Rata, Cameron Highlands,
|
||||
<collectingRegion id="8F01F862DA0CFFFAFB5FFB4DFACA0A0B" box="[1233,1318,1274,1298]" country="Malaysia" name="Pahang" pageId="9" pageNumber="514">Pahang</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="35D27610DA0CFFFAFABBFB4DFA700A0B" box="[1333,1436,1274,1298]" name="Malaysia" pageId="9" pageNumber="514">Malaysia</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="28F15047DA0CFFFAFF11FAA9FECA0A2F" box="[159,294,1310,1334]" degrees="4" direction="north" minutes="31.275" orientation="latitude" pageId="9" pageNumber="514" precision="1" value="4.52125">4°31.275’N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="28F15047DA0CFFFAFEBBFAA9FE390A2F" box="[309,469,1310,1334]" degrees="101" direction="east" minutes="23,193" orientation="longitude" pageId="9" pageNumber="514" precision="1" value="101.38655">101°23,193’E</geoCoordinate>
|
||||
;
|
||||
<quantity id="8A3D9B65DA0CFFFAFE6AFAA9FDAC0A2F" box="[484,576,1310,1334]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="9" pageNumber="514" unit="m" value="2050.0">
|
||||
<elevation id="C6E8D1B3DA0CFFFAFE6AFAA9FDAC0A2F" box="[484,576,1310,1334]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="9" pageNumber="514" unit="m" value="2050.0">2050 m</elevation>
|
||||
</quantity>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="FDAD3CDDDA0CFFFAFDD8FAA9FAE80A43" collectorName="Norhayati Ahmad & L. Lee Grismer" country="Malaysia" elevation="1239" latitude="3.7098334" location="Lee Grismer on Bishop's Trail" longLatPrecision="1" longitude="101.73727" pageId="9" pageNumber="514" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
|
||||
Adult female (LSUHC 9150) collected by
|
||||
<collectorName id="E0305356DA0CFFFAFBC9FAA9FAFE0A2F" box="[1095,1298,1310,1334]" pageId="9" pageNumber="514">Norhayati Ahmad</collectorName>
|
||||
and
|
||||
<collectorName id="E0305356DA0CFFFAFADEFAA8FF1F0A43" pageId="9" pageNumber="514">
|
||||
L.
|
||||
<location id="481A605BDA0CFFFAFAFCFAA8FF1F0A43" LSID="urn:lsid:plazi:treatment:C56C8796DA0CFFFFFF19FCB2FB910DB7:481A605BDA0CFFFAFAFCFAA8FF1F0A43" country="Malaysia" latitude="3.7098334" longLatPrecision="1" longitude="101.73727" name="Lee Grismer on Bishop's Trail" pageId="9" pageNumber="514" stateProvince="Pahang">Lee Grismer</location>
|
||||
</collectorName>
|
||||
on Bishop’s Trail at Fraser’s Hill,
|
||||
<collectingRegion id="8F01F862DA0CFFFAFDF9FAF5FD200A43" box="[631,716,1346,1370]" country="Malaysia" name="Pahang" pageId="9" pageNumber="514">Pahang</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="35D27610DA0CFFFAFD56FAF5FCD30A43" box="[728,831,1346,1370]" name="Malaysia" pageId="9" pageNumber="514">Malaysia</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="28F15047DA0CFFFAFCC0FAF5FC0D0A43" box="[846,993,1346,1370]" degrees="3" direction="north" minutes="42.590" orientation="latitude" pageId="9" pageNumber="514" precision="1" value="3.7098334">3º 42.590’ N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="28F15047DA0CFFFAFC62FAF5FB7B0A43" box="[1004,1175,1346,1370]" degrees="101" direction="east" minutes="44.236" orientation="longitude" pageId="9" pageNumber="514" precision="1" value="101.73727">101º 44.236’ E</geoCoordinate>
|
||||
;
|
||||
<quantity id="8A3D9B65DA0CFFFAFB2DFAF5FB100A43" box="[1187,1276,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.239" pageId="9" pageNumber="514" unit="m" value="1239.0">
|
||||
<elevation id="C6E8D1B3DA0CFFFAFB2DFAF5FB100A43" box="[1187,1276,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.239" pageId="9" pageNumber="514" unit="m" value="1239.0">1239 m</elevation>
|
||||
</quantity>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0CFFFAFF47FAD1FC940A67" box="[201,888,1382,1407]" pageId="9" pageNumber="514" type="description">
|
||||
<paragraph id="4D7A3680DA0CFFFAFF47FAD1FC940A67" blockId="9.[151,1436,1094,2024]" box="[201,888,1382,1407]" pageId="9" pageNumber="514">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47FAD1FDA60A66" bold="true" box="[201,586,1382,1407]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">Additional specimens examined.</emphasis>
|
||||
See
|
||||
<tableCitation id="0047033BDA0CFFFAFD0EFAD1FD3F0A67" box="[640,723,1382,1406]" captionStart="TABLE 5" captionStartId="12.[151,241,733,757]" captionTargetBox="[159,1394,816,1872]" captionTargetPageId="12" captionText="TABLE 5. Correlation between the presence of banding and a vertebral stripe with total length (TL) in Asthenodipsas lasgalenensis sp. nov. Asterisked species could not be included in the genetic analysis. ** Tail broken." pageId="9" pageNumber="514">Table 5</tableCitation>
|
||||
and appendix.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0CFFFAFF47FA3DFE3909DB" pageId="9" pageNumber="514" type="diagnosis">
|
||||
<paragraph id="4D7A3680DA0CFFFAFF47FA3DFE3909DB" blockId="9.[151,1436,1094,2024]" pageId="9" pageNumber="514">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47FA3DFEAF0ABB" box="[201,323,1418,1443]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47FA3DFED00ABA" bold="true" box="[201,316,1418,1443]" pageId="9" pageNumber="514">Diagnosis</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName id="8AC54D03DA0CFFFAFEC2FA3DFD620ABB" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[332,654,1418,1442]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="514" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFEC2FA3DFD620ABB" box="[332,654,1418,1442]" italics="true" pageId="9" pageNumber="514">Asthenodipsas lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0CFFFAFD19FA3DFD1E0ABB" bold="true" box="[663,754,1418,1442]" pageId="9" pageNumber="514">
|
||||
<taxonomicNameLabel id="648257E9DA0CFFFAFD19FA3DFD1E0ABB" box="[663,754,1418,1442]" pageId="9" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is distinguished from all other species of the
|
||||
<taxonomicName id="8AC54D03DA0CFFFAFA8CFA3DFA9A0ABB" authorityName="Romer" authorityYear="1956" box="[1282,1398,1418,1442]" class="Reptilia" family="Pareatidae" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="514" phylum="Chordata" rank="family">Pareatidae</taxonomicName>
|
||||
by the unique combination of having of a maximum TL 771 (
|
||||
<bibRefCitation id="29544B71DA0CFFFAFCA6FA19FC230ADF" author="Smedley, N." box="[808,975,1454,1478]" pageId="9" pageNumber="514" pagination="105 - 123" refId="ref11103" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum of Singapore, 6, 105 - 123." type="journal article" year="1931">Smedley 1931</bibRefCitation>
|
||||
); 15/15/14–15 dorsal scale rows167–190 ventrals; 54–77 subcaudal scales; one or two postorbitals; 1–2+1–3+1–2 temporals; 6–8 supralabials and infralabials; whitish ventral scales bearing small dark lateral spots; dorsum of adults unicolor dark brown; dorsum of hatchlings and juveniles bearing an orangish to light gray ground color overlain with 42–56 irregularly shaped rhomboidal darker bands beginning posterior to dark nuchal bar and extending the length of body and tail and onto lateral spots; edges of ventral scales but not encircling body; a light vertebral stripe; whitish labials usually edged in darker color; dark brown to black head; dark red to red-orange eyes. These character states are summarized across all species in Table 3.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0CFFF8FF47F979FAF20995" lastPageId="11" lastPageNumber="516" pageId="9" pageNumber="514" type="description">
|
||||
<paragraph id="4D7A3680DA0CFFFAFF47F979FAA408F1" blockId="9.[151,1436,1094,2024]" pageId="9" pageNumber="514">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47F979FE0409FF" box="[201,488,1742,1767]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
|
||||
<emphasis id="7FB1EA92DA0CFFFAFF47F979FE0D09FE" bold="true" box="[201,481,1742,1767]" pageId="9" pageNumber="514">
|
||||
Description of
|
||||
<typeStatus id="927E8822DA0CFFFAFEF4F978FE0D09FE" box="[378,481,1743,1767]" pageId="9" pageNumber="514" type="holotype">holotype</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Adult male SVL
|
||||
<quantity id="8A3D9B65DA0CFFFAFD3DF979FCFC09FF" box="[691,784,1742,1766]" metricMagnitude="-1" metricUnit="m" metricValue="4.12" pageId="9" pageNumber="514" unit="mm" value="412.0">412 mm</quantity>
|
||||
and Tal
|
||||
<quantity id="8A3D9B65DA0CFFFAFCFAF979FC3F09FF" box="[884,979,1742,1766]" metricMagnitude="-1" metricUnit="m" metricValue="1.17" pageId="9" pageNumber="514" unit="mm" value="117.0">117 mm</quantity>
|
||||
; rostral as wide as high; head somewhat bulbous in shape, longer than wide; nasals undivided; internasals shorter than prefrontals; prefrontals contact eye; frontal hexagonal, slightly longer than wide; loreals absent; supraoculars subpentagonal, half the length and same width as frontal; preoculars longer than wide; upper and lower postocular, lower postocular extending to below orbit; no suboculars; supralabials 7/7 with 3
|
||||
<superScript id="BAB09BC8DA0CFFFAFD0FF8E9FD620875" attach="left" box="[641,654,1886,1900]" fontSize="6" pageId="9" pageNumber="514">rd</superScript>
|
||||
and 4
|
||||
<superScript id="BAB09BC8DA0CFFFAFD5CF8E9FD320875" attach="left" box="[722,734,1886,1900]" fontSize="6" pageId="9" pageNumber="514">th</superScript>
|
||||
contacting orbit and 7
|
||||
<superScript id="BAB09BC8DA0CFFFAFC57F8E9FC090875" attach="left" box="[985,997,1886,1900]" fontSize="6" pageId="9" pageNumber="514">th</superScript>
|
||||
elongate; temporals 2+2+2/2+2+2, left side has one small scale beteween postoculars and first pair of temporals; mental triangular wider than long; anterior inframaxillary rectangular, in contact with infralabials 1–5; inframaxillaries slightly rounded, elongated ovals in three pairs following the anterior inframaxillary; infralabials 8/8 with the 1
|
||||
<superScript id="BAB09BC8DA0CFFFAFBB2F87AFBAA08C2" attach="left" box="[1084,1094,1997,2011]" fontSize="6" pageId="9" pageNumber="514">st</superScript>
|
||||
pair in medial contact.
|
||||
</paragraph>
|
||||
<caption id="19BA6608DA0FFFF9FF19F81BFB6A08F8" ID-DOI="http://doi.org/10.5281/zenodo.5266003" ID-Zenodo-Dep="5266003" httpUri="https://zenodo.org/record/5266003/files/figure.png" pageId="10" pageNumber="515" startId="10.[151,250,1964,1987]" targetBox="[151,1436,193,1943]" targetPageId="10">
|
||||
<paragraph id="4D7A3680DA0FFFF9FF19F81BFB6A08F8" blockId="10.[151,1437,1964,2018]" pageId="10" pageNumber="515">
|
||||
<emphasis id="7FB1EA92DA0FFFF9FF19F81BFEFE08DA" bold="true" box="[151,274,1964,1987]" pageId="10" pageNumber="515">FIGURE 2.</emphasis>
|
||||
Upper. Adult male
|
||||
<taxonomicName id="8AC54D03DA0FFFF9FE57F81AFD1608DA" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[473,762,1965,1987]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="515" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0FFFF9FE57F81AFD1608DA" box="[473,762,1965,1987]" italics="true" pageId="10" pageNumber="515">Asthenodipsas lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0FFFF9FD71F81AFCA308DB" bold="true" box="[767,847,1965,1986]" pageId="10" pageNumber="515">
|
||||
<taxonomicNameLabel id="648257E9DA0FFFF9FD71F81AFCA308DB" box="[767,847,1965,1986]" pageId="10" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
from Bukit Larut, Perak (holotype LSUHC 8869). Lower. Adult female
|
||||
<taxonomicName id="8AC54D03DA0FFFF9FEACF87BFE2A08FB" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[290,454,1996,2018]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="515" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0FFFF9FEACF87BFE2A08FB" box="[290,454,1996,2018]" italics="true" pageId="10" pageNumber="515">A. lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0FFFF9FE43F87BFDF108F8" bold="true" box="[461,541,1996,2017]" pageId="10" pageNumber="515">
|
||||
<taxonomicNameLabel id="648257E9DA0FFFF9FE43F87BFDF108F8" box="[461,541,1996,2017]" pageId="10" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
from Cameron Highlands, Pahang (paratype LSUHC 6954).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="19BA6608DA0EFFF8FF19FBA8FCD00B6B" ID-DOI="http://doi.org/10.5281/zenodo.5266007" ID-Zenodo-Dep="5266007" httpUri="https://zenodo.org/record/5266007/files/figure.png" pageId="11" pageNumber="516" startId="11.[151,250,1055,1078]" targetBox="[151,1436,193,1034]" targetPageId="11">
|
||||
<paragraph id="4D7A3680DA0EFFF8FF19FBA8FCD00B6B" blockId="11.[151,1436,1055,1138]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF19FBA8FEF80B2F" bold="true" box="[151,276,1055,1078]" pageId="11" pageNumber="516">FIGURE 3.</emphasis>
|
||||
Coloration of a hatchling
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FDA9FBA8FCDC0B2C" authorityName="Grossman and Tillack" authorityYear="2003" box="[551,816,1055,1077]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="vertebralis">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FDA9FBA8FCDC0B2C" box="[551,816,1055,1077]" italics="true" pageId="11" pageNumber="516">Asthenodipsas vertebralis</emphasis>
|
||||
</taxonomicName>
|
||||
from Fraser’s Hill, Pahang (upper left LSUHC 9100) and a subadult female from Bukit Larut, Perak lower right LSUHC 9837. Coloration of hatchling
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FBB2FB89FB0D0B4D" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[1084,1249,1086,1108]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="lasgalenensis">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FBB2FB89FB0D0B4D" box="[1084,1249,1086,1108]" italics="true" pageId="11" pageNumber="516">A. lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
from Bukit Larut, Perak (LSUHC 10798, upper right and LSUHC 10797, lower left).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47FB28FF070BE6" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
Body long, thin, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsals smooth, 15/ 15/
|
||||
<quantity id="8A3D9B65DA0EFFF8FF35FB74FF1F0BC2" box="[187,243,1219,1243]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="11" pageNumber="516" unit="in" value="15.0">15 in</quantity>
|
||||
number, vertebrals greatly enlarged; 189 ventrals; 64 divided subcaudals; anal scute entire; tail tapering to a point.
|
||||
</paragraph>
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47FABCFD0E0A75" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47FABCFE1B0A3A" box="[201,503,1291,1316]" inLineHeading="true" pageId="11" pageNumber="516" reason="1">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47FABCFE1C0A3D" bold="true" box="[201,496,1291,1316]" pageId="11" pageNumber="516">
|
||||
Coloration in life (
|
||||
<figureCitation id="D5FE2A05DA0EFFF8FE2DFABCFE040A3D" box="[419,488,1291,1316]" captionStart="FIGURE 2" captionStartId="10.[151,250,1964,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure-50@10.[151,1436,193,1943]" captionTargetPageId="10" captionText="FIGURE 2. Upper. Adult male Asthenodipsas lasgalenensis sp. nov. from Bukit Larut, Perak (holotype LSUHC 8869). Lower. Adult female A. lasgalenensis sp. nov. from Cameron Highlands, Pahang (paratype LSUHC 6954)." figureDoi="http://doi.org/10.5281/zenodo.5266003" httpUri="https://zenodo.org/record/5266003/files/figure.png" pageId="11" pageNumber="516">Fig. 2</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head, body and tail unicolor dark brown; labials white, edged in dark brown; belly white, edged in dark brown and bearing scattered, dark lateral blotches extending into midventral region; subcaudal region gray laterally white midventrally; iris orange.
|
||||
</paragraph>
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47FAC0FD740AAA" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47FAC0FEAC0A96" box="[201,320,1399,1424]" inLineHeading="true" pageId="11" pageNumber="516" reason="1">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47FAC0FED50A89" bold="true" box="[201,313,1399,1424]" pageId="11" pageNumber="516">Variation</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
The
|
||||
<typeStatus id="927E8822DA0EFFF8FEF2FACFFE0B0A89" box="[380,487,1400,1424]" pageId="11" pageNumber="516" type="paratype">paratypes</typeStatus>
|
||||
approximate the
|
||||
<typeStatus id="927E8822DA0EFFF8FD20FAC0FCFC0A96" box="[686,784,1399,1423]" pageId="11" pageNumber="516" type="holotype">holotype</typeStatus>
|
||||
in all aspects of coloration except that LSUHC 6593 and 9150 have fewer dark lateral ventral blotches.
|
||||
</paragraph>
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47FA08FAF20995" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47FA08FDF50AC1" bold="true" box="[201,537,1471,1496]" inLineHeading="true" pageId="11" pageNumber="516" reason="1">
|
||||
Juvenile coloration (
|
||||
<figureCitation id="D5FE2A05DA0EFFF8FE4CFA08FDE60AC1" box="[450,522,1471,1496]" captionStart="FIGURE 3" captionStartId="11.[151,250,1055,1078]" captionTargetBox="[151,1436,193,1034]" captionTargetId="figure-429@11.[151,1436,193,1034]" captionTargetPageId="11" captionText="FIGURE 3. Coloration of a hatchling Asthenodipsas vertebralis from Fraser’s Hill, Pahang (upper left LSUHC 9100) and a subadult female from Bukit Larut, Perak lower right LSUHC 9837. Coloration of hatchling A. lasgalenensis from Bukit Larut, Perak (LSUHC 10798, upper right and LSUHC 10797, lower left)." figureDoi="http://doi.org/10.5281/zenodo.5266007" httpUri="https://zenodo.org/record/5266007/files/figure.png" pageId="11" pageNumber="516">Fig. 3</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
Juveniles LSUHC 7228, 7240, 9099, 9151, 10259–60, 10789, 10799, 10797, 10997 (see
|
||||
<tableCitation id="0047033BDA0EFFF8FE96FA54FE870AE2" box="[280,363,1507,1531]" captionStart="TABLE 2" captionStartId="4.[151,241,1644,1668]" captionTargetPageId="4" captionText="TABLE 2. Uncorrected p-distances for the major clades of A. vertebralis calculated with MEGA v5.02 (Tamura et al. 2012). Genetic distances in bold are within clades and distances in regular text are between major clades of the A. vertebralis groups." pageId="11" pageNumber="516">Table 2</tableCitation>
|
||||
for scalation and appendix for locality data) that are not part of the
|
||||
<typeStatus id="927E8822DA0EFFF8FBEFFA53FB7D0AE5" box="[1121,1169,1508,1532]" pageId="11" pageNumber="516">type</typeStatus>
|
||||
series bear a coloration and pattern of having dark brown on the top of the head; an orangish to light gray ground color overlain with 42–56 irregular hourglass-shaped darker bands beginning posterior to the dark nuchal bar and extending the length of the body and tail and onto lateral edges of tventral scales but not encircling body; a light vertebral stripe; whitish labials and ventral scales usually edged in darker color; dark brown to black head; and red-orange eyes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0EFFF8FF47F920FC9F0802" pageId="11" pageNumber="516" type="distribution">
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47F920FC9F0802" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47F920FE8E09B6" box="[201,354,1687,1712]" inLineHeading="true" pageId="11" pageNumber="516" reason="1">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47F920FEB709A9" bold="true" box="[201,347,1687,1712]" pageId="11" pageNumber="516">Distribution</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FEE3F920FD5D09B6" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[365,689,1687,1711]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FEE3F920FD5D09B6" box="[365,689,1687,1711]" italics="true" pageId="11" pageNumber="516">Asthenodipsas lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0EFFF8FD32F92FFCF609A9" bold="true" box="[700,794,1688,1712]" pageId="11" pageNumber="516">
|
||||
<taxonomicNameLabel id="648257E9DA0EFFF8FD32F92FFCF609A9" box="[700,794,1688,1712]" pageId="11" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is known only from Genting Highlands, Fraser’s Hill, Cameron Highlands,
|
||||
<collectingRegion id="8F01F862DA0EFFF8FE1AF90CFE0A09CA" box="[404,486,1723,1747]" country="Malaysia" name="Pahang" pageId="11" pageNumber="516">Pahang</collectingRegion>
|
||||
and Bukit Larut,
|
||||
<collectingRegion id="8F01F862DA0EFFF8FD46F90CFCE409CA" box="[712,776,1723,1747]" country="Malaysia" name="Perak" pageId="11" pageNumber="516">Perak</collectingRegion>
|
||||
(
|
||||
<figureCitation id="D5FE2A05DA0EFFF8FC90F90CFC8B09CA" box="[798,871,1723,1747]" captionStart="FIGURE 1" captionStartId="3.[151,250,1982,2005]" captionTargetBox="[151,1424,195,1952]" captionTargetId="figure-45@3.[151,1436,195,1958]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of Asthenodipsas vertebralis (yellow circles), A. lasgalenensis sp. nov. (blue circles), and A. tropidonotus (orange circles) in Peninsular Malaysia and Northern Sumatra." figureDoi="http://doi.org/10.5281/zenodo.5266001" httpUri="https://zenodo.org/record/5266001/files/figure.png" pageId="11" pageNumber="516">Fig. 1</figureCitation>
|
||||
). It is likely
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FB9AF90BFB3D09CD" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[1044,1233,1724,1748]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FB9AF90BFB3D09CD" box="[1044,1233,1724,1748]" italics="true" pageId="11" pageNumber="516">A. lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0EFFF8FB51F90BFAD309CD" bold="true" box="[1247,1343,1724,1748]" pageId="11" pageNumber="516">
|
||||
<taxonomicNameLabel id="648257E9DA0EFFF8FB51F90BFAD309CD" box="[1247,1343,1724,1748]" pageId="11" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FA0AF90BFEF209EE" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="vertbebralis">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FA0AF90BFEF209EE" italics="true" pageId="11" pageNumber="516">A. vertbebralis</emphasis>
|
||||
</taxonomicName>
|
||||
range throughout the montane cloud forests of northeastern Peninsular
|
||||
<collectingCountry id="35D27610DA0EFFF8FBCEF968FB4B09EE" box="[1088,1191,1759,1783]" name="Malaysia" pageId="11" pageNumber="516">Malaysia</collectingCountry>
|
||||
as well as throughout similar habitats in southern
|
||||
<collectingCountry id="35D27610DA0EFFF8FE40F8B4FDDD0802" box="[462,561,1795,1819]" name="Thailand" pageId="11" pageNumber="516">Thailand</collectingCountry>
|
||||
south of the Isthmus of Kra.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0EFFF8FF47F89FFE4D089E" pageId="11" pageNumber="516" type="biology_ecology">
|
||||
<paragraph id="4D7A3680DA0EFFF8FF47F89FFE4D089E" blockId="11.[151,1436,1183,2035]" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47F89FFE650859" bold="true" box="[201,393,1831,1856]" pageId="11" pageNumber="516">Natural History</emphasis>
|
||||
. The localities from which
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FD5AF890FBFB0826" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[724,1047,1831,1855]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FD5AF890FBFB0826" box="[724,1047,1831,1855]" italics="true" pageId="11" pageNumber="516">Asthenodipsas lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA0EFFF8FBACF89FFB6C0859" bold="true" box="[1058,1152,1832,1856]" pageId="11" pageNumber="516">
|
||||
<taxonomicNameLabel id="648257E9DA0EFFF8FBACF89FFB6C0859" box="[1058,1152,1832,1856]" pageId="11" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
was found are montane cloud forests above
|
||||
<quantity id="8A3D9B65DA0EFFF8FEF4F8FCFE2D087A" box="[378,449,1867,1891]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="516" unit="m" value="800.0">800 m</quantity>
|
||||
in elevation. All specimens were found at least
|
||||
<quantity id="8A3D9B65DA0EFFF8FC6FF8FCFBE1087A" box="[993,1037,1867,1891]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="516" unit="m" value="1.0">1 m</quantity>
|
||||
above the ground at night foraging or resting in vegetation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA0EFFFFFF47F823FD530FCD" lastPageId="12" lastPageNumber="517" pageId="11" pageNumber="516" type="etymology">
|
||||
<paragraph id="4D7A3680DA0EFFFFFF47F823FD530FCD" blockId="11.[151,1436,1183,2035]" lastBlockId="12.[151,1436,151,686]" lastPageId="12" lastPageNumber="517" pageId="11" pageNumber="516">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47F823FEA308B2" box="[201,335,1940,1964]" inLineHeading="true" pageId="11" pageNumber="516" reason="1">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FF47F823FEA408B5" bold="true" box="[201,328,1940,1964]" pageId="11" pageNumber="516">Etymology</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
The specific epithet
|
||||
<taxonomicName id="8AC54D03DA0EFFF8FDCDF823FD3608B5" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[579,730,1940,1964]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="516" phylum="Chordata" rank="species" species="lasgalenensis">
|
||||
<emphasis id="7FB1EA92DA0EFFF8FDCDF823FD3608B5" box="[579,730,1940,1964]" italics="true" pageId="11" pageNumber="516">lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
is derived from the name
|
||||
<emphasis id="7FB1EA92DA0EFFF8FB83F823FB5F08B5" box="[1037,1203,1940,1964]" italics="true" pageId="11" pageNumber="516">Eryn Lasgalen</emphasis>
|
||||
which means in the “Wood of Greenleaves” in the fictional Sindarian language from J.R.R Tolkien’s
|
||||
<emphasis id="7FB1EA92DA0EFFF8FBBEF800FAC508D6" box="[1072,1321,1975,1999]" italics="true" pageId="11" pageNumber="516">The Lord of the Rings</emphasis>
|
||||
(1955). It was the name used by the Wood Elves for the Mirkwood Forest after its cleansing following the War of the Ring. This name was chosen because
|
||||
<bibRefCitation id="29544B71DA09FFFFFD88FF20FD2D0FB6" author="Tolkien, J. R. R." box="[518,705,151,175]" pageId="12" pageNumber="517" refId="ref11362" refString="Tolkien, J. R. R. (1955) The Lord of the Rings: Return of the King. George Allen & Unwin, Australia, 560 pp." type="book" year="1955">Tolkien’s (1955)</bibRefCitation>
|
||||
description of this forest showed great similarity to the cloudy, upland forests within which this species is found.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="05DF650BDA09FFFFFF47FF68FB910DB7" pageId="12" pageNumber="517" type="discussion">
|
||||
<paragraph id="4D7A3680DA09FFFFFF47FF68FB910DB7" blockId="12.[151,1436,151,686]" pageId="12" pageNumber="517">
|
||||
<emphasis id="7FB1EA92DA09FFFFFF47FF68FE820FEE" box="[201,366,223,248]" inLineHeading="true" pageId="12" pageNumber="517" reason="1">
|
||||
<emphasis id="7FB1EA92DA09FFFFFF47FF68FE840FE1" bold="true" box="[201,360,223,248]" pageId="12" pageNumber="517">Comparisons</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName id="8AC54D03DA09FFFFFEF9FF57FD540FE1" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[375,696,224,248]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA09FFFFFEF9FF57FD540FE1" box="[375,696,224,248]" italics="true" pageId="12" pageNumber="517">Asthenodipsas lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA09FFFFFD4EFF57FCF70FE1" bold="true" box="[704,795,224,248]" pageId="12" pageNumber="517">
|
||||
<taxonomicNameLabel id="648257E9DA09FFFFFD4EFF57FCF70FE1" box="[704,795,224,248]" pageId="12" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is differentiated from all other species of
|
||||
<taxonomicName id="8AC54D03DA09FFFFFB74FF57FA700FE1" baseAuthorityName="Grossmann & Tillack" baseAuthorityYear="2003" box="[1274,1436,224,248]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="genus">
|
||||
<emphasis id="7FB1EA92DA09FFFFFB74FF57FA700FE1" box="[1274,1436,224,248]" italics="true" pageId="12" pageNumber="517">Asthenodipsas</emphasis>
|
||||
</taxonomicName>
|
||||
on the basis of phylogenetics (
|
||||
<figureCitation id="D5FE2A05DA09FFFFFE78FEB3FDD20E05" box="[502,574,260,284]" captionStart="FIGURE 4" captionStartId="13.[151,250,1010,1033]" captionTargetBox="[156,1436,198,989]" captionTargetId="figure-404@13.[151,1436,193,989]" captionTargetPageId="13" captionText="FIGURE 4. Bayesian Inference tree based on 1,073 bp of cyt b showing the relationships between Pareatidae and the placement of Asthenodipsas lasgalenensis sp. nov. Bayesian posterior probabilities (BPP) and ML and MP bootstrap support values, respectively (BPP/ML/MP), at the nodes." figureDoi="http://doi.org/10.5281/zenodo.5266009" httpUri="https://zenodo.org/record/5266009/files/figure.png" pageId="12" pageNumber="517">Fig. 4</figureCitation>
|
||||
;
|
||||
<tableCitation id="0047033BDA09FFFFFDC5FEB3FD4E0E05" box="[587,674,260,284]" captionStart="TABLE 2" captionStartId="4.[151,241,1644,1668]" captionTargetPageId="4" captionText="TABLE 2. Uncorrected p-distances for the major clades of A. vertebralis calculated with MEGA v5.02 (Tamura et al. 2012). Genetic distances in bold are within clades and distances in regular text are between major clades of the A. vertebralis groups." pageId="12" pageNumber="517">Table 2</tableCitation>
|
||||
), morphology, color pattern (Tables 3,4;
|
||||
<figureCitation id="D5FE2A05DA09FFFFFBF4FEB3FB240E05" box="[1146,1224,260,284]" captionStart="FIGURE 2" captionStartId="10.[151,250,1964,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure-50@10.[151,1436,193,1943]" captionTargetPageId="10" captionText="FIGURE 2. Upper. Adult male Asthenodipsas lasgalenensis sp. nov. from Bukit Larut, Perak (holotype LSUHC 8869). Lower. Adult female A. lasgalenensis sp. nov. from Cameron Highlands, Pahang (paratype LSUHC 6954)." figureDoi="http://doi.org/10.5281/zenodo.5266003" httpUri="https://zenodo.org/record/5266003/files/figure.png" pageId="12" pageNumber="517">Figs. 2</figureCitation>
|
||||
–5,7), and perhaps microhabitat preference. It is discretely diagnosable from
|
||||
<taxonomicName id="8AC54D03DA09FFFFFCB3FE9FFC360E59" authorityName="Grossman and Tillack" authorityYear="2003" box="[829,986,296,320]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="vertebralis">
|
||||
<emphasis id="7FB1EA92DA09FFFFFCB3FE9FFC360E59" box="[829,986,296,320]" italics="true" pageId="12" pageNumber="517">A. vertebralis</emphasis>
|
||||
</taxonomicName>
|
||||
(a species to which it was previously ascribed) and
|
||||
<taxonomicName id="8AC54D03DA09FFFFFEBAFEFBFE0D0E7D" baseAuthorityName="Lidth de Jeude" baseAuthorityYear="1923" box="[308,481,332,356]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="tropidonotus">
|
||||
<emphasis id="7FB1EA92DA09FFFFFEBAFEFBFE0D0E7D" box="[308,481,332,356]" italics="true" pageId="12" pageNumber="517">A. tropidonotus</emphasis>
|
||||
</taxonomicName>
|
||||
based on having 167–190 ventral scales as opposed to having 195–215 and 199–204 ventral scales, respectively. It is further separated from
|
||||
<taxonomicName id="8AC54D03DA09FFFFFC8AFEC7FC710E91" authorityName="Grossman and Tillack" authorityYear="2003" box="[772,925,368,392]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="vertebralis">
|
||||
<emphasis id="7FB1EA92DA09FFFFFC8AFEC7FC710E91" box="[772,925,368,392]" italics="true" pageId="12" pageNumber="517">A. vertebralis</emphasis>
|
||||
</taxonomicName>
|
||||
in having a significantly lower mean number (
|
||||
<emphasis id="7FB1EA92DA09FFFFFF2AFE2CFF5E0EAB" box="[164,178,411,434]" italics="true" pageId="12" pageNumber="517">x</emphasis>
|
||||
=46.8; 40–56) of trapezoidal-shaped dorsal bands as opposed to a higher mean number (
|
||||
<emphasis id="7FB1EA92DA09FFFFFB2AFE2CFB5E0EAB" box="[1188,1202,411,434]" italics="true" pageId="12" pageNumber="517">x</emphasis>
|
||||
=62.0; 57–64) and a significant difference in the mean number of subcaudal scales (
|
||||
<emphasis id="7FB1EA92DA09FFFFFCEFFE75FC830EC3" box="[865,879,450,474]" italics="true" pageId="12" pageNumber="517">x</emphasis>
|
||||
=64.5; 54–77 vs.
|
||||
<emphasis id="7FB1EA92DA09FFFFFBB2FE75FBA60EC3" box="[1084,1098,450,474]" italics="true" pageId="12" pageNumber="517">x</emphasis>
|
||||
=72.3; 65–77). It is discretely diagnosable from
|
||||
<taxonomicName id="8AC54D03DA09FFFFFEEFFE55FE160EE3" authorityName="Grossman and Tillack" authorityYear="2003" box="[353,506,482,506]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="vertebralis">
|
||||
<emphasis id="7FB1EA92DA09FFFFFEEFFE55FE160EE3" box="[353,506,482,506]" italics="true" pageId="12" pageNumber="517">A. vertebralis</emphasis>
|
||||
</taxonomicName>
|
||||
in having yellow to orangish labials and ventrals as opposed to whitish labials and ventrals; a yellowish vertebral stripe throughout life as opposed to losing the light vertebral stripe between
|
||||
<quantity id="8A3D9B65DA09FFFFFACEFDB1FA770D04" box="[1344,1435,518,542]" metricMagnitude="-1" metricUnit="m" metricValue="3.16" pageId="12" pageNumber="517" unit="mm" value="316.0">316 mm</quantity>
|
||||
and
|
||||
<quantity id="8A3D9B65DA09FFFFFF47FD9EFECB0D58" box="[201,295,553,577]" metricMagnitude="-1" metricUnit="m" metricValue="3.4899999999999998" pageId="12" pageNumber="517" unit="mm" value="349.0">349 mm</quantity>
|
||||
TL; and a dorsal banding pattern throughout life as opposed to transforming from a dorsal banded pattern to a unicolor pattern between
|
||||
<quantity id="8A3D9B65DA09FFFFFDCEFDF9FD710D7C" box="[576,669,590,614]" metricMagnitude="-1" metricUnit="m" metricValue="3.16" pageId="12" pageNumber="517" unit="mm" value="316.0">316 mm</quantity>
|
||||
and
|
||||
<quantity id="8A3D9B65DA09FFFFFD58FDF9FCDF0D7C" box="[726,819,590,614]" metricMagnitude="-1" metricUnit="m" metricValue="3.4899999999999998" pageId="12" pageNumber="517" unit="mm" value="349.0">349 mm</quantity>
|
||||
TL. Furthermore, the top of the head in hatchling and juvenile
|
||||
<taxonomicName id="8AC54D03DA09FFFFFF72FDC5FE590D93" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[252,437,626,650]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA09FFFFFF72FDC5FE590D93" box="[252,437,626,650]" italics="true" pageId="12" pageNumber="517">A. lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA09FFFFFE4EFDC5FDF10D93" bold="true" box="[448,541,626,650]" pageId="12" pageNumber="517">
|
||||
<taxonomicNameLabel id="648257E9DA09FFFFFE4EFDC5FDF10D93" box="[448,541,626,650]" pageId="12" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is dark brown as opposed to orange. Additionally,
|
||||
<taxonomicName id="8AC54D03DA09FFFFFBF4FDC5FAD80D93" authority="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer, 2013" authorityName="Loredo & Wood & Quah & Anuar & Greer & Ahmad & Grismer" authorityYear="2013" box="[1146,1332,626,650]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="lasgalenensis" status="sp. nov.">
|
||||
<emphasis id="7FB1EA92DA09FFFFFBF4FDC5FAD80D93" box="[1146,1332,626,650]" italics="true" pageId="12" pageNumber="517">A. lasgalenensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7FB1EA92DA09FFFFFAB1FDC5FA700D93" bold="true" box="[1343,1436,626,650]" pageId="12" pageNumber="517">
|
||||
<taxonomicNameLabel id="648257E9DA09FFFFFAB1FDC5FA700D93" box="[1343,1436,626,650]" pageId="12" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
appears to be terrestrial when sympatric with
|
||||
<taxonomicName id="8AC54D03DA09FFFFFD1BFD21FCC20DB7" authorityName="Grossman and Tillack" authorityYear="2003" box="[661,814,662,686]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="517" phylum="Chordata" rank="species" species="vertebralis">
|
||||
<emphasis id="7FB1EA92DA09FFFFFD1BFD21FCC20DB7" box="[661,814,662,686]" italics="true" pageId="12" pageNumber="517">A. vertebralis</emphasis>
|
||||
</taxonomicName>
|
||||
as opposed to being arboreal.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
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