Add updates up until 2024-12-11 13:59:56

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<document id="128557993B860C1A8BD95D5C59E9FC5C" ID-DOI="10.11646/zootaxa.5551.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14390102" ID-ZooBank="844F8BB4-B2FC-4BF6-BA33-3DD7A18CD988" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924457367" checkinUser="plazi" docAuthor="Keetapithchayakul, Tosaphol Saetung, Makbun, Noppadon, Ignatius, Kaewpawika Jitthamma &amp; Phan, Quoc Toan" docDate="2024" docId="03A44734835A4A60FF71FC69C0B84002" docLanguage="en" docName="zootaxa.5551.1.5.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prodasineura sangkhla Keetapithchayakul, Makbun, Ignatius &amp; Phan, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="134" masterDocId="FF9D3F4C835B4A63FFE6FFC7C14D4733" masterDocTitle="Prodasineura sangkhla sp. nov., a new damselfly from western Thailand (Odonata: Platycnemididae)" masterLastPageNumber="140" masterPageNumber="131" pageNumber="132" updateTime="1733925609185" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4913E119FA7C3783C98A543301A5EC43">Prodasineura sangkhla sp. nov., a new damselfly from western Thailand (Odonata: Platycnemididae)</mods:title>
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<mods:namePart id="34FFBE16282FFF3C5ECB8FC339697CAA">Keetapithchayakul, Tosaphol Saetung</mods:namePart>
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<mods:namePart id="5A8AEF4D509A49C290DE954FB84D14C6">Makbun, Noppadon</mods:namePart>
<mods:affiliation id="AF871608BF4B20BD0B7C5B9838D0CD3B">211 / 5 Moo 4, Takhli, Nakhon Sawan, 60140, Thailand</mods:affiliation>
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<mods:namePart id="AC59DF2199B14434F0B527BC8C360B34">Ignatius, Kaewpawika Jitthamma</mods:namePart>
<mods:affiliation id="90A376B7846EB03231631AFA83B4C9AB">Forest and Plant Conservation Research Office, Department of National Parks, Wildlife and Plant Conservation, Bangkok 10900, Thailand</mods:affiliation>
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<mods:namePart id="22E7B6D2FB8CD6E0FD3FEBAA237A5425">Phan, Quoc Toan</mods:namePart>
<mods:affiliation id="F2174BA7699538A7A330F81A56D5B8A1">The Center for Entomology &amp; Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<mods:date id="2DC380F18026A2A28007D6D620B22C2D">2024</mods:date>
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<heading id="D0FA414E835A4A62FF71FC69C35A44FA" bold="true" box="[151,535,942,969]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<emphasis id="B9792A30835A4A62FF71FC69C35A44FA" bold="true" box="[151,535,942,969]" pageId="1" pageNumber="132">
<taxonomicName id="4C0D8DA1835A4A62FF71FC69C0FF44FA" authorityName="Keetapithchayakul &amp; Makbun &amp; Ignatius &amp; Phan" authorityYear="2024" box="[151,434,942,969]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="sangkhla" status="sp. nov.">
<emphasis id="B9792A30835A4A62FF71FC69C0FF44FA" bold="true" box="[151,434,942,969]" italics="true" pageId="1" pageNumber="132">Prodasineura sangkhla</emphasis>
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<taxonomicNameLabel id="A24A974B835A4A62FE5FFC68C35A44FA" box="[441,535,943,969]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C317A5A9835A4A62FF71FC14C0034392" pageId="1" pageNumber="132" type="description">
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(
<figureCitation id="1336EAA7835A4A62FF79FC14C06B44DE" box="[159,294,979,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1277,1301]" captionStartId-1="3.[151,250,1492,1516]" captionStartId-2="4.[151,250,619,643]" captionStartId-3="4.[151,250,1695,1719]" captionStartId-4="5.[151,250,1878,1902]" captionStartId-5="6.[151,250,1254,1278]" captionTargetBox-0="[151,1436,179,1251]" captionTargetBox-1="[151,1436,179,1467]" captionTargetBox-2="[151,1436,181,583]" captionTargetBox-3="[151,1436,702,1650]" captionTargetBox-4="[154,1434,179,1852]" captionTargetBox-5="[151,1436,414,1228]" captionTargetId-0="figure-18@2.[151,1436,179,1251]" captionTargetId-1="figure-18@3.[151,1436,179,1467]" captionTargetId-2="figure-18@4.[151,1436,181,596]" captionTargetId-3="figure-38@4.[151,1436,702,1670]" captionTargetId-4="figure-18@5.[154,1434,179,1852]" captionTargetId-5="figure-147@6.[151,1436,414,1228]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionText-0="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." captionText-1="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." captionText-2="FIGURE 3. Genital ligula of holotype ♂ (A), in ventral view &amp; (B), in lateral view." captionText-3="FIGURE 4. Prothorax of paratype ♀ of Prodasineura sangkhla sp. nov. (A, B), in lateral and dorsal view; (C, D), illustrations in oblique-dorsal and dorsal view." captionText-4="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." captionText-5="FIGURE 6. Prodasineura sangkhla sp. nov. in typical location. (A), a male; (B), a tandem in flight; (C), an ovipositing female in association with a male (Takhian Thong waterfall, 4.vi.2024, photographed by QTP)." figureDoi-0="http://doi.org/10.5281/zenodo.14390104" figureDoi-1="http://doi.org/10.5281/zenodo.14390108" figureDoi-2="http://doi.org/10.5281/zenodo.14390110" figureDoi-3="http://doi.org/10.5281/zenodo.14390114" figureDoi-4="http://doi.org/10.5281/zenodo.14390116" figureDoi-5="http://doi.org/10.5281/zenodo.14390120" httpUri-0="https://zenodo.org/record/14390104/files/figure.png" httpUri-1="https://zenodo.org/record/14390108/files/figure.png" httpUri-2="https://zenodo.org/record/14390110/files/figure.png" httpUri-3="https://zenodo.org/record/14390114/files/figure.png" httpUri-4="https://zenodo.org/record/14390116/files/figure.png" httpUri-5="https://zenodo.org/record/14390120/files/figure.png" pageId="1" pageNumber="132">Figures 16</figureCitation>
)
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<emphasis id="B9792A30835A4A62FF71FBDCC0334306" box="[151,382,1051,1077]" italics="true" pageId="1" pageNumber="132">Specimens examined</emphasis>
.
<materialsCitation id="3B65FC7F835A4A62FE6AFBDCC4D9436A" collectingDate="2024-06-04" collectionCode="T, TNHM" collectorName="Phan" country="Thailand" county="Thai-Myanmar" elevation="205" latitude="15.30327" location="Takhian Thong" longLatPrecision="1" longitude="98.45249" municipality="Sangkhla Buri District" pageId="1" pageNumber="132" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="holotype">
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<typeStatus id="54B64880835A4A62FE6AFBDCC0B64306" box="[396,507,1051,1077]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D0B3DAB835A4A62FDE0FBDCC3504306" box="[518,541,1051,1077]" count="1" pageId="1" pageNumber="132" type="male"></specimenCount>
,
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waterfall (
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N
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,
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E
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,
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elevation),
<collectingMunicipality id="6BD66C58835A4A62FAD2FBDCC069436A" pageId="1" pageNumber="132">Sangkhla Buri District</collectingMunicipality>
at the
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border,
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,
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,
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<collectingDate id="EFF7290A835A4A62FC1AFBF8C53D436A" box="[1020,1136,1087,1113]" pageId="1" pageNumber="132" value="2024-06-04">4.VI.2024</collectingDate>
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, Q.
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leg.
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[
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.
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<materialsCitation id="3B65FC7F835A4A62FF21FBA4C23D434F" box="[199,880,1122,1149]" collectingDate="2024-06-04" collectionCode="ZCDTU" collectorName="Phan" country="Thailand" county="Thai-Myanmar" elevation="205" latitude="15.30327" location="Takhian Thong" longLatPrecision="1" longitude="98.45249" municipality="Sangkhla Buri District" pageId="1" pageNumber="132" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Kanchanaburi" typeStatus="paratype">
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<typeStatus id="54B64880835A4A62FF21FBA4C072434E" box="[199,319,1123,1149]" pageId="1" pageNumber="132" type="paratype">Paratypes</typeStatus>
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.
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,
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, same data as holotype
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[
<collectionCode id="ED1C6EE7835A4A62FCE4FBA5C224434F" box="[770,873,1122,1148]" pageId="1" pageNumber="132">ZCDTU</collectionCode>
]
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;
<materialsCitation id="3B65FC7F835A4A62FC67FBA4C00A4393" collectingDate="2024-06-04" collectionCode="NMC, ECNP" collectorName="Phan" country="Thailand" county="Thai-Myanmar" elevation="205" latitude="15.30327" location="Takhian Thong" longLatPrecision="1" longitude="98.45249" municipality="Sangkhla Buri District" pageId="1" pageNumber="132" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Kanchanaburi" typeStatus="paratype">
<specimenCount id="9D0B3DAB835A4A62FC67FBA4C288434E" box="[897,965,1123,1149]" count="4" pageId="1" pageNumber="132" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D0B3DAB835A4A62FC35FBA4C54D434E" box="[979,1024,1123,1149]" count="1" pageId="1" pageNumber="132" type="female">1 ♀</specimenCount>
(in tandem), same data as holotype
<emphasis id="B9792A30835A4A62FF71FB41C00A4393" bold="true" box="[151,327,1158,1184]" pageId="1" pageNumber="132">
[
<collectionCode id="ED1C6EE7835A4A62FF78FB41C1A94393" box="[158,228,1158,1184]" country="USA" lsid="urn:lsid:biocol.org:col:15488" name="New Mexico State University" pageId="1" pageNumber="132" type="Herbarium">NMC</collectionCode>
,
<collectionCode id="ED1C6EE7835A4A62FF17FB41C00D4393" box="[241,320,1158,1184]" pageId="1" pageNumber="132">ECNP</collectionCode>
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.
</paragraph>
</subSubSection>
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<emphasis id="B9792A30835A4A62FF21FB6CC00543F6" bold="true" box="[199,328,1195,1221]" pageId="1" pageNumber="132">Etymology</emphasis>
. The new species is named after the Sangkhla Buri District, where the new species was discovered. The specific epithet is a noun in apposition. The word of &quot;Sangkhla&quot; is a Burmese/Thai language, meaning diversity or combination, reflecting the area's rich ethnic people combination, which similarly corresponds to the diversity of
<taxonomicName id="4C0D8DA1835A4A62FF71FAD0C0C54202" box="[151,392,1303,1329]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF71FAD0C07D4202" box="[151,304,1303,1329]" italics="true" pageId="1" pageNumber="132">Prodasineura</emphasis>
species
</taxonomicName>
found this area.
</paragraph>
<paragraph id="8BB2F622835A4A62FF21FAFDC0BD4266" blockId="1.[151,1437,1050,1869]" box="[199,496,1338,1365]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21FAFDC0A44266" bold="true" box="[199,489,1338,1365]" pageId="1" pageNumber="132">
Descriptions of
<typeStatus id="54B64880835A4A62FE64FAFCC0A44266" box="[386,489,1339,1365]" pageId="1" pageNumber="132" type="holotype">holotype</typeStatus>
</emphasis>
.
</paragraph>
<paragraph id="8BB2F622835A4A62FF21FA98C46942AE" blockId="1.[151,1437,1050,1869]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21FA98C049424A" box="[199,260,1375,1401]" italics="true" pageId="1" pageNumber="132">Head</emphasis>
(
<figureCitation id="1336EAA7835A4A62FEF0FA98C0EB424A" box="[278,422,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[151,1436,179,1467]" captionTargetId="figure-18@3.[151,1436,179,1467]" captionTargetPageId="3" captionText="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="1" pageNumber="132">Fig. 2A, 2C</figureCitation>
). Head and antennae shining black except the base of antennae, the space between eye margins and genae yellowish, and a transverse opaque white band at the level of vertex to eyes margins.
</paragraph>
<paragraph id="8BB2F622835A4A62FF21FA60C381413A" blockId="1.[151,1437,1050,1869]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21FA60C05542F2" box="[199,280,1447,1473]" italics="true" pageId="1" pageNumber="132">Thorax</emphasis>
(
<figureCitation id="1336EAA7835A4A62FEC0FA60C0C542F2" box="[294,392,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="1" pageNumber="132">Figs. 1A</figureCitation>
,
<figureCitation id="1336EAA7835A4A62FE74FA60C0AD42F2" box="[402,480,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[151,1436,179,1467]" captionTargetId="figure-18@3.[151,1436,179,1467]" captionTargetPageId="3" captionText="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="1" pageNumber="132">2C, 2E</figureCitation>
) black, mostly covered by white pruinescence with some black in the area surrounding thoracic spiracle and metepimeron.
<emphasis id="B9792A30835A4A62FDCDFA0CC32D42D7" box="[555,608,1483,1508]" italics="true" pageId="1" pageNumber="132">Legs</emphasis>
(
<figureCitation id="1336EAA7835A4A62FD96FA0CC38A42D6" box="[624,711,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[151,1436,179,1467]" captionTargetId="figure-18@3.[151,1436,179,1467]" captionTargetPageId="3" captionText="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="1" pageNumber="132">Fig. 2C</figureCitation>
) mostly black with yellowish marking at the base of femur, and white pruinescence on coxa, trochanter and femur.
</paragraph>
<paragraph id="8BB2F622835A4A62FF21F9D4C2364162" blockId="1.[151,1437,1050,1869]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21F9D4C046411F" box="[199,267,1555,1580]" italics="true" pageId="1" pageNumber="132">Wings</emphasis>
(
<figureCitation id="1336EAA7835A4A62FEFBF9D4C038411E" box="[285,373,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="1" pageNumber="132">Fig. 1A</figureCitation>
) completely hyaline with black veins. Px
<quantity id="4CF55BC7835A4A62FCBCF9D5C2D8411E" box="[858,917,1554,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="1" pageNumber="132" unit="in" value="15.0">15 in</quantity>
forewings and
<quantity id="4CF55BC7835A4A62FBAEF9D4C5CE411E" box="[1096,1155,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="1" pageNumber="132" unit="in" value="13.0">13 in</quantity>
hindwings, respectively. Anal bridge vein on both wings has short arch. Pterostigma black.
</paragraph>
<paragraph id="8BB2F622835A4A62FF21F99CC418401A" blockId="1.[151,1437,1050,1869]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21F99CC07C4146" box="[199,305,1627,1653]" italics="true" pageId="1" pageNumber="132">Abdomen</emphasis>
(
<figureCitation id="1336EAA7835A4A62FEA7F99CC0D24146" box="[321,415,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="1" pageNumber="132">Figs 1A</figureCitation>
,
<figureCitation id="1336EAA7835A4A62FE4AF99DC0B24146" box="[428,511,1626,1653]" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="1" pageNumber="132">5A, 5C</figureCitation>
) mostly black except the following: yellow spots on lateral and lower margins of S1; tiny yellowish markings on ventrolateral surfaces of S27; S89 entirely black; S10 entirely blue on dorsum and extending to upper half of lateral side.
<emphasis id="B9792A30835A4A62FD95F964C279418E" box="[627,820,1699,1725]" italics="true" pageId="1" pageNumber="132">Anal appendages</emphasis>
(
<figureCitation id="1336EAA7835A4A62FCA5F964C2F6418E" box="[835,955,1698,1725]" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="1" pageNumber="132">Fig. 5AF</figureCitation>
). Cerci hammer-like shaped and paraprocts broad at base, tapering abruptly to a rounded tip; cerci blue with the two robust ventral spines, with the inner spine curved inward, tiny-shaped in lateral view (
<figureCitation id="1336EAA7835A4A62FD66F92CC2784036" box="[640,821,1770,1797]" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="1" pageNumber="132">Fig. 5B, 5D, 5F</figureCitation>
); paraprocts black with a tiny yellow spot at base (
<figureCitation id="1336EAA7835A4A62FA89F92CC1F6401A" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="1" pageNumber="132">Fig. 5B</figureCitation>
).
<emphasis id="B9792A30835A4A62FF29F8C8C026401A" box="[207,363,1807,1833]" italics="true" pageId="1" pageNumber="132">Genital ligula</emphasis>
(
<figureCitation id="1336EAA7835A4A62FE9DF8C8C0F0401A" box="[379,445,1807,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,619,643]" captionTargetBox="[151,1436,181,583]" captionTargetId="figure-18@4.[151,1436,181,596]" captionTargetPageId="4" captionText="FIGURE 3. Genital ligula of holotype ♂ (A), in ventral view &amp; (B), in lateral view." figureDoi="http://doi.org/10.5281/zenodo.14390110" httpUri="https://zenodo.org/record/14390110/files/figure.png" pageId="1" pageNumber="132">Fig. 3</figureCitation>
) with a pair of lateral robust flagella with curved tips and two apical robust lobes.
</paragraph>
</subSubSection>
<subSubSection id="C317A5A9835A4A60FF21F8F4C0B84002" lastPageId="3" lastPageNumber="134" pageId="1" pageNumber="132" type="description">
<paragraph id="8BB2F622835A4A62FF21F8F4C5C6407E" blockId="1.[151,1437,1050,1869]" box="[199,1163,1842,1869]" pageId="1" pageNumber="132">
<emphasis id="B9792A30835A4A62FF21F8F4C0A9407F" bold="true" box="[199,484,1842,1869]" pageId="1" pageNumber="132">Measurements (in mm).</emphasis>
Hindwing 19.8; abdomen (including anal appendages) 28.9.
</paragraph>
<caption id="DF72A6AA83594A61FF71FB3AC50C4227" ID-DOI="http://doi.org/10.5281/zenodo.14390104" ID-Zenodo-Dep="14390104" box="[151,1089,1277,1301]" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="2" pageNumber="133" startId="2.[151,250,1277,1301]" targetBox="[151,1436,179,1251]" targetPageId="2" targetType="figure">
<paragraph id="8BB2F62283594A61FF71FB3AC50C4227" blockId="2.[151,1089,1277,1301]" box="[151,1089,1277,1301]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF71FB3AC05E4226" bold="true" box="[151,275,1277,1301]" pageId="2" pageNumber="133">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="4C0D8DA183594A61FE76FB3AC3314226" authorityName="Keetapithchayakul &amp; Makbun &amp; Ignatius &amp; Phan" authorityYear="2024" box="[400,636,1277,1301]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="sangkhla" status="sp. nov.">
<emphasis id="B9792A3083594A61FE76FB3AC3314226" box="[400,636,1277,1301]" italics="true" pageId="2" pageNumber="133">Prodasineura sangkhla</emphasis>
</taxonomicName>
<emphasis id="B9792A3083594A61FD65FB3AC39E4226" bold="true" box="[643,723,1277,1301]" pageId="2" pageNumber="133">
<taxonomicNameLabel id="A24A974B83594A61FD65FB3AC39E4226" box="[643,723,1277,1301]" pageId="2" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A), holotype ♂ &amp; (B), paratype ♀.
</paragraph>
</caption>
<paragraph id="8BB2F62283594A61FF21FAA0C08042B1" blockId="2.[151,1437,1383,1985]" box="[199,461,1383,1410]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21FAA0C08B42B2" bold="true" box="[199,454,1383,1409]" pageId="2" pageNumber="133">Description of female</emphasis>
.
</paragraph>
<paragraph id="8BB2F62283594A61FF21FA4CC3454106" blockId="2.[151,1437,1383,1985]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21FA4CC0494296" box="[199,260,1419,1445]" italics="true" pageId="2" pageNumber="133">Head</emphasis>
(
<figureCitation id="1336EAA783594A61FEF7FA4CC0DB4296" box="[273,406,1419,1445]" captionStart="FIGURE 2" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[151,1436,179,1467]" captionTargetId="figure-18@3.[151,1436,179,1467]" captionTargetPageId="3" captionText="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="2" pageNumber="133">Fig. 2B, 2D</figureCitation>
). Labrum yellowish with a black dot beneath upper margin and lower margin black; mandibles and genae yellowish; the yellow color extends from the eyes margin to the level of postclypeus; anteclypeus black, pale yellowish centrally; postclypeus black with two tiny yellow spots at corners of upper margin. Antennae black except for the second segment yellowish anteriorly. Front and rear of head black with a large transverse yellowish stripe at level of anterior ocellus.
</paragraph>
<paragraph id="8BB2F62283594A61FF21F9F8C58C41F6" blockId="2.[151,1437,1383,1985]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21F9F8C06D416A" box="[199,288,1599,1625]" italics="true" pageId="2" pageNumber="133">Thorax.</emphasis>
Prothorax black but laterally yellowish, this color forming a large stripe on lateral pronotal lobe, extending to a half of the projections of posterior pronotal lobe. Posterior lobe with a pair of horns which blunt apically and curving upward, and two adjacent ear-like structures (
<figureCitation id="1336EAA783594A61FC9EF940C2F14192" box="[888,956,1671,1697]" captionStart="FIGURE 4" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,702,1650]" captionTargetId="figure-38@4.[151,1436,702,1670]" captionTargetPageId="4" captionText="FIGURE 4. Prothorax of paratype ♀ of Prodasineura sangkhla sp. nov. (A, B), in lateral and dorsal view; (C, D), illustrations in oblique-dorsal and dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390114" httpUri="https://zenodo.org/record/14390114/files/figure.png" pageId="2" pageNumber="133">Fig. 4</figureCitation>
). Synthorax black with three yellow stripes as in
<figureCitation id="1336EAA783594A61FF34F96CC01841F6" box="[210,341,1707,1733]" captionStart="FIGURE 2" captionStartId="3.[151,250,1492,1516]" captionTargetBox="[151,1436,179,1467]" captionTargetId="figure-18@3.[151,1436,179,1467]" captionTargetPageId="3" captionText="FIGURE 2. Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of Prodasineura sangkhla sp. nov. (A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="2" pageNumber="133">
Fig. 2D,
<specimenCount id="9D0B3DAB83594A61FED0F96CC01841F6" box="[310,341,1707,1733]" count="2" pageId="2" pageNumber="133" type="generic">2F</specimenCount>
</figureCitation>
.
<emphasis id="B9792A3083594A61FEB9F96BC0D941F6" box="[351,404,1708,1733]" italics="true" pageId="2" pageNumber="133">Legs</emphasis>
(
<figureCitation id="1336EAA783594A61FE45F96CC0B541F6" box="[419,504,1707,1733]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="2" pageNumber="133">Fig. 1B</figureCitation>
) black, except for coxae, trochanter and outer femur yellowish.
</paragraph>
<paragraph id="8BB2F62283594A61FF21F917C025403E" blockId="2.[151,1437,1383,1985]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21F917C04641DA" box="[199,267,1744,1769]" italics="true" pageId="2" pageNumber="133">Wings</emphasis>
(
<figureCitation id="1336EAA783594A61FEFBF908C03841DA" box="[285,373,1743,1769]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="2" pageNumber="133">Fig. 1B</figureCitation>
) hyaline, forewings slightly longer than hindwings, Px
<quantity id="4CF55BC783594A61FC10F908C57F41DA" box="[1014,1074,1743,1769]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="2" pageNumber="133" unit="in" value="14.0">14 in</quantity>
both forewings and hindwings. Pterostigma black.
</paragraph>
<paragraph id="8BB2F62283594A61FF21F8D0C5CB40AE" blockId="2.[151,1437,1383,1985]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21F8D0C07C4002" box="[199,305,1815,1841]" italics="true" pageId="2" pageNumber="133">Abdomen</emphasis>
(
<figureCitation id="1336EAA783594A61FED9F8D0C0D74002" box="[319,410,1815,1841]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1301]" captionTargetBox="[151,1436,179,1251]" captionTargetId="figure-18@2.[151,1436,179,1251]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Prodasineura sangkhla sp. nov. (A), holotype ♂ &amp; (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.14390104" httpUri="https://zenodo.org/record/14390104/files/figure.png" pageId="2" pageNumber="133">Figs 1B</figureCitation>
,
<figureCitation id="1336EAA783594A61FE40F8D0C0A14001" box="[422,492,1815,1842]" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="2" pageNumber="133">5GH</figureCitation>
) black with the following yellowish pattern: S1 with a large spot on the lateral sides; S29 with narrow yellowish stripes along lower margins; S36 brownyellowish with an oval yellowish spot on the anterior margin of each segment; dorsal S810 with longitudinal yellow stripe centro-dorsally. Cerci (
<figureCitation id="1336EAA783594A61FAFFF898C4D94049" box="[1305,1428,1887,1914]" captionStart="FIGURE 5" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[154,1434,179,1852]" captionTargetId="figure-18@5.[154,1434,179,1852]" captionTargetPageId="5" captionText="FIGURE 5. Abdominal tip including appendages of Prodasineura sangkhla sp. nov. [AF] holotype ♂ &amp; [GH] paratype ♀. (A, B, G, H), abdominal tip in dorsal and lateral view; (C, D), appendages, in posterodorsal view; (E, F), appendages in obliqueventral view." figureDoi="http://doi.org/10.5281/zenodo.14390116" httpUri="https://zenodo.org/record/14390116/files/figure.png" pageId="2" pageNumber="133">Fig. 5GH</figureCitation>
) yellowish except for lower margin black. Ovipositor black with distal end dark yellowish.
</paragraph>
<paragraph id="8BB2F62283594A61FF21F860C5CB40F2" blockId="2.[151,1437,1383,1985]" box="[199,1158,1959,1985]" pageId="2" pageNumber="133">
<emphasis id="B9792A3083594A61FF21F860C03B40F2" bold="true" box="[199,374,1959,1985]" pageId="2" pageNumber="133">Measurements</emphasis>
(in mm). Hindwing 18.3; abdomen (including anal appendages) 28.3.
</paragraph>
<caption id="DF72A6AA83584A60FF71FA13C2B64122" ID-DOI="http://doi.org/10.5281/zenodo.14390108" ID-Zenodo-Dep="14390108" httpUri="https://zenodo.org/record/14390108/files/figure.png" pageId="3" pageNumber="134" startId="3.[151,250,1492,1516]" targetBox="[151,1436,179,1467]" targetPageId="3" targetType="figure">
<paragraph id="8BB2F62283584A60FF71FA13C2B64122" blockId="3.[151,1437,1492,1553]" pageId="3" pageNumber="134">
<emphasis id="B9792A3083584A60FF71FA13C05942DF" bold="true" box="[151,276,1492,1516]" pageId="3" pageNumber="134">FIGURE 2.</emphasis>
Structures of holotype ♂ [A, C, E] and paratype ♀ [B, D, F] of
<taxonomicName id="4C0D8DA183584A60FC54FA12C5D242DE" authorityName="Keetapithchayakul &amp; Makbun &amp; Ignatius &amp; Phan" authorityYear="2024" box="[946,1183,1493,1517]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="sangkhla" status="sp. nov.">
<emphasis id="B9792A3083584A60FC54FA12C5D242DE" box="[946,1183,1493,1517]" italics="true" pageId="3" pageNumber="134">Prodasineura sangkhla</emphasis>
</taxonomicName>
<emphasis id="B9792A3083584A60FB41FA12C5B442DE" bold="true" box="[1191,1273,1493,1517]" pageId="3" pageNumber="134">
<taxonomicNameLabel id="A24A974B83584A60FB41FA12C5B442DE" box="[1191,1273,1493,1517]" pageId="3" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A, B), head in frontal view; (C, D), head and thorax in lateral view; (E, F), synthorax in dorsal view.
</paragraph>
</caption>
<paragraph id="8BB2F62283584A60FF21F9A4C0B84002" blockId="3.[151,1437,1635,1841]" pageId="3" pageNumber="134">
<emphasis id="B9792A3083584A60FF21F9A4C3E7414E" bold="true" box="[199,682,1635,1661]" pageId="3" pageNumber="134">
Variation in males and females
<typeStatus id="54B6488083584A60FDD2F9A4C3E7414E" box="[564,682,1635,1661]" pageId="3" pageNumber="134" type="paratype">paratypes</typeStatus>
</emphasis>
. Other male and female
<typeStatus id="54B6488083584A60FC50F9A4C56C414E" box="[950,1057,1635,1661]" pageId="3" pageNumber="134" type="paratype">paratypes</typeStatus>
from the type locality provided no significant differential features from the
<typeStatus id="54B6488083584A60FDBFF940C3F64192" box="[601,699,1671,1697]" pageId="3" pageNumber="134" type="holotype">holotype</typeStatus>
male and the above-described female, except their measurements: In male
<typeStatus id="54B6488083584A60FF14F96CC02F41F6" box="[242,354,1707,1733]" pageId="3" pageNumber="134" type="paratype">paratypes</typeStatus>
: hindwing ranges from
<quantity id="4CF55BC783584A60FD8EF96CC38641F6" box="[616,715,1707,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="3" pageNumber="134" unit="mm" value="18.4">18.4 mm</quantity>
to
<quantity id="4CF55BC783584A60FD0BF96CC21D41F6" box="[749,848,1707,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="3" pageNumber="134" unit="mm" value="19.4">19.4 mm</quantity>
and abdomen including anal appendages is
<quantity id="4CF55BC783584A60FADFF96CC4D041F6" box="[1337,1437,1707,1733]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="3" pageNumber="134" unit="mm" value="28.9">28.9 mm</quantity>
to
<quantity id="4CF55BC783584A60FF52F908C05641DA" box="[180,283,1743,1769]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="3" pageNumber="134" unit="mm" value="31.2">31.2 mm</quantity>
; with Px is
<quantity id="4CF55BC783584A60FE7BF908C34D41DA" box="[413,512,1743,1770]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="3" pageNumber="134" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">1415 in</quantity>
forewing and
<quantity id="4CF55BC783584A60FD45F908C24B41DA" box="[675,774,1743,1769]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="3" pageNumber="134" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">1314 in</quantity>
hindwing. In female
<typeStatus id="54B6488083584A60FC10F908C52941DA" box="[1014,1124,1743,1769]" pageId="3" pageNumber="134" type="paratype">paratypes</typeStatus>
, hindwing is
<quantity id="4CF55BC783584A60FB1CF908C4D041DA" box="[1274,1437,1743,1769]" metricMagnitude="-2" metricUnit="m" metricValue="1.9700000000000002" metricValueMax="2.0100000000000002" metricValueMin="1.9300000000000002" pageId="3" pageNumber="134" unit="mm" value="19.7" valueMax="20.1" valueMin="19.3">19.320.1 mm</quantity>
and abdomen including anal appendages is 28.0
<quantity id="4CF55BC783584A60FD2DF934C27E403E" box="[715,819,1779,1805]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="3" pageNumber="134" unit="mm" value="28.3">28.3 mm</quantity>
. Forewing and hindwing of
<typeStatus id="54B6488083584A60FB9CF934C597403E" box="[1146,1242,1779,1805]" pageId="3" pageNumber="134" type="paratype">paratype</typeStatus>
females with Px 1314 and 1213, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>