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<document id="6AB3828AF553740DD293FEE3385500CE" ENCODING="UTF-8" ID-CLB-Dataset="276867" ID-DOI="https://doi.org/10.5281/zenodo.291971" ID-GBIF-Dataset="0bb2f033-2b79-4d87-803f-91694067c3f3" ID-ISBN="978-0-9506207-7-0" ID-Zenodo-Dep="291971" checkinTime="1588973351969" checkinUser="admin" docAuthor="Jarvis, Charlie" docDate="2007" docId="2B7B5496895A9A33C0C7B3F29468A01C" docLanguage="en" docName="ch7.gg.C.xml" docOrigin="Order out of Chaos. Linnaean Plant Types and their Types, London: Linnaean Society of London in association with the Natural History Museum" docTitle="Calystegia spithamaea Pursh" docType="treatment" docVersion="11" lastPageNumber="442" masterDocId="E213C9FA410C8D52CE14FE18C476F6F3" masterDocTitle="Chapter 7: Linnaean Plant Names and their Types (part C)" masterLastPageNumber="473" masterPageNumber="370" pageNumber="442" updateTime="1699548976333" updateUser="ExternalLinkService">
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<mods:title id="1872DC87FE9C0A152112F78D37B5DE80">Chapter 7: Linnaean Plant Names and their Types (part C)</mods:title>
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<mods:title id="4DBA5CAFD6BDDDE4AC7B715634B318B3">Order out of Chaos. Linnaean Plant Types and their Types</mods:title>
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<treatment id="2B7B5496895A9A33C0C7B3F29468A01C" ID-DOI="http://doi.org/10.5281/zenodo.4380115" ID-GBIF-Taxon="164003205" ID-Zenodo-Dep="4380115" LSID="urn:lsid:plazi:treatment:2B7B5496895A9A33C0C7B3F29468A01C" httpUri="http://treatment.plazi.org/id/2B7B5496895A9A33C0C7B3F29468A01C" lastPageNumber="442" pageNumber="442">
<subSubSection id="DC3569DD5E8CBD938A36AE7411263C9C" pageNumber="442" type="reference_group">
<paragraph id="CEF0CCEE78FB2AE01C603FC288E34D99" pageNumber="442">
<treatmentCitation id="89EC5A7CB7382582D668838C9760A2CB" ID-DOI="http://doi.org/10.5281/zenodo.4356360" ID-Zenodo-Dep="4356360" author="Linnaeus" httpUri="http://treatment.plazi.org/id/E868C71F7BEE07ACEE52AA1FE2CB12D7" pageNumber="158" year="1753">
<bibCitation id="96A903EB466C1A61362CD306BA47DDF4" author="Linnaeus" pagination="158" title="Convolvulus spithamaeus" volumeTitle="Species Plantarum 1" year="1753">
<taxonomicName id="C546D9F33CC8F72EDFDDB95EAA3C6BD0" ID-CoL="7JMNP" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Convolvulaceae" genus="Convolvulus" kingdom="Plantae" order="Solanales" phylum="Tracheophyta" rank="species" species="spithamaeus">
<emphasis id="955AF6202C8A606A966321E0B0F407B0" bold="true" pageNumber="442">Convolvulus spithamaeus</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="655C794321AF0C7AC27CF83BECFBACD5">
<emphasis id="8BB26AB9662FEAD0F0B2B612C469BCBD" italics="true" pageNumber="442">Species Plantarum</emphasis>
1
</volumeTitle>
: 158. 1753
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DF8F114B3DA99DA36A4C04625D06F75" pageNumber="442" type="distribution">
<paragraph id="09CE6425E2E60BF85086F7B8E242C45C" pageNumber="442">&quot;Habitat in Virginia.&quot; RCN: 1260.</paragraph>
</subSubSection>
<subSubSection id="50DBD50C1D2925399B18FE533BDD18A7" pageNumber="442" type="materials_examined">
<paragraph id="9383F8C3AAD807DC9178536D44145CB0" pageNumber="442">
<typeStatus id="75E5866EE76F88AB95A974A56873970D">Lectotype</typeStatus>
(Tryon in
<emphasis id="5836D74445C40DCD9ED301CC78718D02" italics="true" pageNumber="442">Rhodora</emphasis>
41: 417, pl. 557. 1939):
<emphasis id="57CB509738BD13AAC56B1C1946AA4A32" italics="true" pageNumber="442">Clayton 553</emphasis>
(BM-000051655).
</paragraph>
</subSubSection>
<subSubSection id="C38C53AB5C041F3028CD963757341D01" pageNumber="442" type="nomenclature">
<paragraph id="9DEB91EC5E2D0C23B139AD167FA76982" pageNumber="442">
Current name:
<taxonomicName id="6D0767313CA536B7C82322F18A9D687F" ID-CoL="Q82Z" authority="(L.) Pursh" authorityName="Pursh" baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Convolvulaceae" genus="Calystegia" kingdom="Plantae" order="Solanales" phylum="Tracheophyta" rank="species" species="spithamaea">
<emphasis id="6B1E11F1AEDC7409EE9DD9FC48A4F135" italics="true" pageNumber="442">
<emphasis id="32FC42CECF23766019AD416405160965" bold="true" italics="true" pageNumber="442">Calystegia spithamaea</emphasis>
</emphasis>
(L.) Pursh
</taxonomicName>
(
<taxonomicName id="D79370FF285BD4A4B2B94544C7DCB184" authorityName="de Jussieu" authorityYear="1789" class="Magnoliopsida" family="Convolvulaceae" kingdom="Plantae" order="Solanales" phylum="Tracheophyta" rank="family">Convolvulaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C9E018BCC191601391EFA8E992F55829" pageNumber="442" type="discussion">
<paragraph id="9A814B44D22072637FC242D2FFE5FFD8" pageNumber="442">
<emphasis id="B6E90418F491327F51C50941765E9DDF" italics="true" pageNumber="442">Note:</emphasis>
<normalizedToken id="751F0B6040179E520A7F7DF84AEE4F76" originalValue="Tryon’s">Tryon's</normalizedToken>
pl. 557 clarifies that the &quot;type specimen&quot; is clearly
<emphasis id="CB022D3685F6BC83A51EF61C0A3A6734" italics="true" pageNumber="442">Clayton 553</emphasis>
(BM).
</paragraph>
</subSubSection>
</treatment>
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Key to the species of
<taxonomicName id="64D24D24FFC33C441AEE49FFFE22FDEB" ID-CoL="8W4T9" authorityName="Lindley" authorityYear="1826" box="[406,504,499,525]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="91A6EAB5FFC33C441AEE49FFFE22FDEB" bold="true" box="[406,504,499,525]" italics="true" pageId="2" pageNumber="67">Zeuxine</emphasis>
</taxonomicName>
from
<collectingCountry id="DBC57637FFC33C44193B49F8FD55FDE8" box="[579,655,500,526]" name="China" pageId="2" pageNumber="67">China</collectingCountry>
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<keyLead id="18239592FFC33C441BEF4A4DFA46FDBE" box="[151,1436,577,600]" pageId="2" pageNumber="67">1. Leaves linear to linear-lanceolate, not stalked, base directly sheathing stem.............................................................. 2.</keyLead>
</paragraph>
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<keyLead id="18239592FFC33C441BEF4A6DFA41FD9E" box="[151,1435,609,632]" pageId="2" pageNumber="67">- Leaves ovate-lanceolate to elliptic, base contracted into a petiole-like stalk sheathing stem ..................................... 3.</keyLead>
</paragraph>
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<keyLead id="18239592FFC33C441BEF4A8DFA46FD7E" box="[151,1436,641,664]" pageId="2" pageNumber="67">
2. Plants
<quantity id="642A9B42FFC33C441A684A8DFEB6FD7E" box="[272,364,641,664]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="2.4" metricValueMin="0.4" pageId="2" pageNumber="67" unit="cm" value="14.0" valueMax="24.0" valueMin="4.0">4–24 cm</quantity>
tall; ovary and sepals glabrous ....................................................................................
<taxonomicName id="64D24D24FFC33C441F954A8DFA46FD7E" ID-CoL="7GFNK" box="[1261,1436,641,664]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="strateumatica">
<emphasis id="91A6EAB5FFC33C441F954A8DFA46FD7E" box="[1261,1436,641,664]" italics="true" pageId="2" pageNumber="67">Z. strateumatica</emphasis>
</taxonomicName>
</keyLead>
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<keyLead id="18239592FFC33C441BEF4AADFA46FD5E" box="[151,1436,673,696]" pageId="2" pageNumber="67">
- Plants
<quantity id="642A9B42FFC33C441A684AADFEA0FD5E" box="[272,378,673,696]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="2" pageNumber="67" unit="cm" value="27.5" valueMax="35.0" valueMin="20.0">20–35 cm</quantity>
tall; ovary and sepals pubescent...............................................................................
<taxonomicName id="64D24D24FFC33C441F914AADFA46FD5E" ID-CoL="5D49J" box="[1257,1436,673,696]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="membranacea">
<emphasis id="91A6EAB5FFC33C441F914AADFA46FD5E" box="[1257,1436,673,696]" italics="true" pageId="2" pageNumber="67">Z. membranacea</emphasis>
</taxonomicName>
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<keyLead id="18239592FFC33C441BEF4ACDFA46FD3E" box="[151,1436,705,728]" pageId="2" pageNumber="67">
3. Midvein of leaves marked with silvery blotches or a white line (also on lateral veins in
<taxonomicName id="64D24D24FFC33C441F124ACDFAC7FD3E" ID-CoL="5D48Y" authorityName="Ormerod" authorityYear="2002" baseAuthorityName="K.Y.Lang" box="[1130,1309,705,728]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="gengmanensis">
<emphasis id="91A6EAB5FFC33C441F124ACDFAC7FD3E" box="[1130,1309,705,728]" italics="true" pageId="2" pageNumber="67">Z. gengmanensis</emphasis>
</taxonomicName>
).............. 4.
</keyLead>
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<keyLead id="18239592FFC33C441BEF4AEDFA41FD1E" box="[151,1435,737,760]" pageId="2" pageNumber="67">- Leaves uniformly green or occasionally tinged with red............................................................................................. 8.</keyLead>
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<keyLead id="18239592FFC33C441BEF4B0DFA46FCFE" box="[151,1436,769,792]" pageId="2" pageNumber="67">
4. Lip entire, rhombic, not adnate to column, base contracted and lacking calli.....................................
<taxonomicName id="64D24D24FFC33C441F964B0DFA46FCFE" ID-CoL="5D495" authorityName="C.S.Leou" authorityYear="1994" box="[1262,1436,769,792]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="integrilabella">
<emphasis id="91A6EAB5FFC33C441F964B0DFA46FCFE" box="[1262,1436,769,792]" italics="true" pageId="2" pageNumber="67">Z. integrilabella</emphasis>
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<keyLead id="18239592FFC33C441BEF4B2DFA41FCBE" pageId="2" pageNumber="67">- Lip bi-tripartite, base adnate to column, hypochile concave and containing 2 subulate, lamellate, or 2-3-digitate calli near base....................................................................................................................................................................... 5.</keyLead>
</paragraph>
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<keyLead id="18239592FFC33C441BEF4B6DFA46FC9E" box="[151,1436,865,888]" pageId="2" pageNumber="67">5. Epichile of lip transversely dilated but as broad as or narrower than hypochile, entire to shallowly bilobed ............ 6.</keyLead>
</paragraph>
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<keyLead id="18239592FFC33C441BEF4B8DFA41FC7E" box="[151,1435,897,920]" pageId="2" pageNumber="67">- Epichile of lip dilated, broader than hypochile, distinctly bilobed.............................................................................. 7.</keyLead>
</paragraph>
</keyStep>
<keyStep id="18262E02FFC33C441BEF4BADFA46FC3E" pageId="2" pageNumber="67">
<paragraph id="A36D36A7FFC33C441BEF4BADFA41FC5E" blockId="2.[151,1436,577,1656]" box="[151,1435,929,952]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4BADFA41FC5E" box="[151,1435,929,952]" pageId="2" pageNumber="67">
6. Ovary sparsely hairy; sepals hairy abaxially ......................................................................................
<taxonomicName id="64D24D24FFC33C441F924BADFA41FC5E" ID-CoL="5D49Z" authorityName="Ames" authorityYear="1938" baseAuthorityName="Ames" box="[1258,1435,929,952]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="philippinensis">
<emphasis id="91A6EAB5FFC33C441F924BADFA41FC5E" box="[1258,1435,929,952]" italics="true" pageId="2" pageNumber="67">Z. philippinensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4BCDFA46FC3E" blockId="2.[151,1436,577,1656]" box="[151,1436,961,984]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4BCDFA46FC3E" box="[151,1436,961,984]" pageId="2" pageNumber="67">
- Ovary glabrous; sepals glabrous ...........................................................................................................
<taxonomicName id="64D24D24FFC33C441F894BCDFA46FC3E" ID-CoL="5D493" box="[1265,1436,961,984]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="goodyeroides">
<emphasis id="91A6EAB5FFC33C441F894BCDFA46FC3E" box="[1265,1436,961,984]" italics="true" pageId="2" pageNumber="67">Z. goodyeroides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="18262E02FFC33C441BEF4BEDFA46FBBE" pageId="2" pageNumber="67">
<paragraph id="A36D36A7FFC33C441BEF4BEDFA46FBFE" blockId="2.[151,1436,577,1656]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4BEDFA46FBFE" pageId="2" pageNumber="67">
7. Stem stout, ca.
<quantity id="642A9B42FFC33C441A144BEDFE73FC1E" box="[364,425,993,1016]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="67" unit="mm" value="5.0">5 mm</quantity>
in diam.; lateral veins of leaves marked with white lines; flowers pink; hypochile containing 2 short, rounded calli...............................................................................................................................
<taxonomicName id="64D24D24FFC33C441F914C0DFA46FBFE" ID-CoL="5D48Y" authorityName="Ormerod" authorityYear="2002" baseAuthorityName="K.Y.Lang" box="[1257,1436,1025,1048]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="gengmanensis">
<emphasis id="91A6EAB5FFC33C441F914C0DFA46FBFE" box="[1257,1436,1025,1048]" italics="true" pageId="2" pageNumber="67">Z. gengmanensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4C2DFA46FBBE" blockId="2.[151,1436,577,1656]" pageId="2" pageNumber="67">
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- Stem slender,
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in diam.; lateral veins of leaves without white lines; flowers white or pale yellow; hypochile containing 2 2-3-digitate to narrowly lamellate calli .....................................................................................
<taxonomicName id="64D24D24FFC33C441E534C4DFA46FBBE" ID-CoL="5D49N" box="[1323,1436,1089,1112]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="nervosa">
<emphasis id="91A6EAB5FFC33C441E534C4DFA46FBBE" box="[1323,1436,1089,1112]" italics="true" pageId="2" pageNumber="67">Z. nervosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<keyLead id="18239592FFC33C441BEF4C6DFA4FFB9E" box="[151,1429,1121,1144]" pageId="2" pageNumber="67">
8. Plants robust,
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tall; leaves 5–12 ×
<quantity id="642A9B42FFC33C4419E44C6DFD30FB9E" box="[668,746,1121,1144]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="67" unit="cm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 cm</quantity>
....................................................................................
<taxonomicName id="64D24D24FFC33C441E5C4C6DFA4FFB9E" ID-CoL="5D49R" box="[1316,1429,1121,1144]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="odorata">
<emphasis id="91A6EAB5FFC33C441E5C4C6DFA4FFB9E" box="[1316,1429,1121,1144]" italics="true" pageId="2" pageNumber="67">Z. odorata</emphasis>
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</keyLead>
</paragraph>
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- Plants slender to robust, to
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tall; leaves 2–6 ×
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....................................................................................... 9.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="18262E02FFC33C441BEF4CADFA46FB3E" pageId="2" pageNumber="67">
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<keyLead id="18239592FFC33C441BEF4CADFA46FB5E" box="[151,1436,1185,1208]" pageId="2" pageNumber="67">9. Leaves not usually withering at anthesis; epichile of lip as broad as or narrower than hypochile............................ 10.</keyLead>
</paragraph>
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<keyLead id="18239592FFC33C441BEF4CCDFA46FB3E" box="[151,1436,1217,1240]" pageId="2" pageNumber="67">- Leaves withering or pendulous at anthesis; epichile of lip dilated, broader than hypochile .................................... 11.</keyLead>
</paragraph>
</keyStep>
<keyStep id="18262E02FFC33C441BEF4CEDFA46FAFE" pageId="2" pageNumber="67">
<paragraph id="A36D36A7FFC33C441BEF4CEDFA46FB1E" blockId="2.[151,1436,577,1656]" box="[151,1436,1249,1272]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4CEDFA46FB1E" box="[151,1436,1249,1272]" pageId="2" pageNumber="67">
10. Leaves ovate-oblong, 3.0–
<quantity id="642A9B42FFC33C441A9B4CEDFDF5FB1E" box="[483,559,1249,1272]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="67" unit="cm" value="4.5">4.5 cm</quantity>
; bract entire, white hypochile containing 2 subulate, calli .................
<taxonomicName id="64D24D24FFC33C441E734CEDFA46FB1E" ID-CoL="5D48B" authorityName="Fukuyama" authorityYear="1934" box="[1291,1436,1249,1272]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="agyokuana">
<emphasis id="91A6EAB5FFC33C441E734CEDFA46FB1E" box="[1291,1436,1249,1272]" italics="true" pageId="2" pageNumber="67">Z. agyokuana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4D0DFA46FAFE" blockId="2.[151,1436,577,1656]" box="[151,1436,1281,1304]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4D0DFA46FAFE" box="[151,1436,1281,1304]" pageId="2" pageNumber="67">
- Leaves ovate,
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; bract ciliate, yellow hypochile containing 2 2-3-digitate calli.........................
<taxonomicName id="64D24D24FFC33C441E624D0DFA46FAFE" ID-CoL="5D49S" authorityName="L. Li &amp; S. J. Li" authorityYear="2013" box="[1306,1436,1281,1304]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="ovalifolia">
<emphasis id="91A6EAB5FFC33C441E624D0DFA46FAFE" box="[1306,1436,1281,1304]" italics="true" pageId="2" pageNumber="67">Z. ovalifolia</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="18262E02FFC33C441BEF4D2DFA46FABE" pageId="2" pageNumber="67">
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<keyLead id="18239592FFC33C441BEF4D2DFA46FADE" box="[151,1436,1313,1336]" pageId="2" pageNumber="67">11. Lip T-shaped; epichile lobes diverging at right angles to axis of lip ......................................................................... 12.</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4D4DFA46FABE" blockId="2.[151,1436,577,1656]" box="[151,1436,1345,1368]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4D4DFA46FABE" box="[151,1436,1345,1368]" pageId="2" pageNumber="67">- Lip Y-shaped; epichile lobes diverging at acute angles to axis of lip; epichile not mucronate at apex..................... 13.</keyLead>
</paragraph>
</keyStep>
<keyStep id="18262E02FFC33C441BEF4D6DFA41FA7E" pageId="2" pageNumber="67">
<paragraph id="A36D36A7FFC33C441BEF4D6DFA46FA9E" blockId="2.[151,1436,577,1656]" box="[151,1436,1377,1400]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4D6DFA46FA9E" box="[151,1436,1377,1400]" pageId="2" pageNumber="67">
12. Lip white .....................................................................................................................................................
<taxonomicName id="64D24D24FFC33C441E6E4D6DFA46FA9E" ID-CoL="8X9S7" box="[1302,1436,1377,1400]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="91A6EAB5FFC33C441E6E4D6DFA46FA9E" box="[1302,1436,1377,1400]" italics="true" pageId="2" pageNumber="67">Z. parvifolia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4D8DFA41FA7E" blockId="2.[151,1436,577,1656]" box="[151,1435,1409,1432]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4D8DFA41FA7E" box="[151,1435,1409,1432]" pageId="2" pageNumber="67">
- Lip yellow ...........................................................................................................................................................
<taxonomicName id="64D24D24FFC33C441E324D8DFA41FA7E" ID-CoL="5D48W" box="[1354,1435,1409,1432]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="flava">
<emphasis id="91A6EAB5FFC33C441E324D8DFA41FA7E" box="[1354,1435,1409,1432]" italics="true" pageId="2" pageNumber="67">Z. flava</emphasis>
</taxonomicName>
</keyLead>
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</keyStep>
<keyStep id="18262E02FFC33C441BEF4DADFA46F9FE" pageId="2" pageNumber="67">
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<keyLead id="18239592FFC33C441BEF4DADFA41FA3E" pageId="2" pageNumber="67">
13. Flower with lip
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; hypochile containing 2 lamellate calli, each often with a linear extension; column wings lacking.............................................................................................................................................................
<taxonomicName id="64D24D24FFC33C441E564DCDFA41FA3E" ID-CoL="7YFLW" box="[1326,1435,1473,1496]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="91A6EAB5FFC33C441E564DCDFA41FA3E" box="[1326,1435,1473,1496]" italics="true" pageId="2" pageNumber="67">Z. grandis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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- Flower with lip to
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; hypochile containing 2 subulate, hooked calli; column with triangular or transversely oblong wings .............................................................................................................................................................. 14.
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<keyLead id="18239592FFC33C441BEF4E2DFA46F9DE" box="[151,1436,1569,1592]" pageId="2" pageNumber="67">
14. Epichile rotated gently downward; epichile lobes obovate-flabellate, 2.0–2.5 × 2.0–
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........................
<taxonomicName id="64D24D24FFC33C441E454E2DFA46F9DE" ID-CoL="5D489" box="[1341,1436,1569,1592]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="91A6EAB5FFC33C441E454E2DFA46F9DE" box="[1341,1436,1569,1592]" italics="true" pageId="2" pageNumber="67">Z. affinis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="A36D36A7FFC33C441BEF4E4DFA46F99E" blockId="2.[151,1436,577,1656]" pageId="2" pageNumber="67">
<keyLead id="18239592FFC33C441BEF4E4DFA46F99E" pageId="2" pageNumber="67">
- Epichile reflexed sharply downward from apex of mesochile; epichile lobes oblong to irregularly rhombic, 2.5–3.0 ×
<quantity id="642A9B42FFC33C441BBF4E6DFE9CF99E" box="[199,326,1633,1656]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="2" pageNumber="67" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.2–1.8 mm</quantity>
......................................................................................................................................................
<taxonomicName id="64D24D24FFC33C441E4F4E6DFA46F99E" ID-CoL="7GFNQ" box="[1335,1436,1633,1656]" class="Liliopsida" family="Orchidaceae" genus="Zeuxine" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="reflexa">
<emphasis id="91A6EAB5FFC33C441E4F4E6DFA46F99E" box="[1335,1436,1633,1656]" italics="true" pageId="2" pageNumber="67">Z. reflexa</emphasis>
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<mods:namePart id="5DF2E3E053006E5ED3F84F50DBF27212">Vijayakumar, S. P.</mods:namePart>
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</mods:role>
<mods:namePart id="9622539E24EF099A7F162AF5B8227C5C">Channakeshavamurthy, B. H.</mods:namePart>
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<mods:name id="1BCACDFE2FB64E2AA63829543D996662" type="personal">
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<mods:namePart id="9E9FBB69073D5B4930981751A480EBF2">Torsekar, Varun R.</mods:namePart>
</mods:name>
<mods:name id="22AE01922995F9B491B2F126B85258ED" type="personal">
<mods:role id="22A291264649E5BAA9472A360F3D8F7A">
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</mods:role>
<mods:namePart id="42D2CD57636815B21C5CFA9F838674B8">Kulkarni, Nirmal U.</mods:namePart>
</mods:name>
<mods:name id="D3062B557F1637766B86D04B6EE509B0" type="personal">
<mods:role id="CDFFB097336F4F38C5A8A5F364907330">
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</mods:role>
<mods:namePart id="9AEE55299CB6974E471DC2C2F43C028E">Shanker, Kartik</mods:namePart>
</mods:name>
<mods:typeOfResource id="A33FE7C2446FB96D196492A6B0DAD6CE">text</mods:typeOfResource>
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<mods:title id="8A1EAC2D737346787D3EF54A6F0EDE1E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A1CB50C3C075618FC45AD92FAF9FC012">
<mods:date id="D111A7363CD4ED1149D92B49EC420645">2015</mods:date>
<mods:detail id="704C8506A09EA6F9686B38C07F1BF88C" type="volume">
<mods:number id="99ABC61D8D24AE1F768C6871D7A5A2AE">3999</mods:number>
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<mods:start id="B12B5705A1CADB7BA713A52003C584F4">79</mods:start>
<mods:end id="DF5A6714BAA882ABC5ECF493DA5633EE">94</mods:end>
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<mods:classification id="844A8AB6FCAFE815E4FE05BD20EC18EC">journal article</mods:classification>
<mods:identifier id="602625420E648C6C8E3053E6BD70A42B" type="DOI">10.11646/zootaxa.3999.1.5</mods:identifier>
<mods:identifier id="2BA69222CE157CBA3CDE71F0B7226AE7" type="GBIF-Dataset">3ef5a58e-b058-4c33-a79d-f768fdc73427</mods:identifier>
<mods:identifier id="2E299887DDC65CE86BE0730C077BAA04" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="78864C9DC2FDB0A90DAABE573AB01128" type="Zenodo-Dep">289212</mods:identifier>
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<treatment id="2B7BF863FFAF880AFF23FDAE026CFE53" ID-DOI="http://doi.org/10.5281/zenodo.5672931" ID-GBIF-Taxon="127692704" ID-Zenodo-Dep="5672931" LSID="urn:lsid:plazi:treatment:2B7BF863FFAF880AFF23FDAE026CFE53" httpUri="http://treatment.plazi.org/id/2B7BF863FFAF880AFF23FDAE026CFE53" lastPageId="4" lastPageNumber="82" pageId="3" pageNumber="82">
<subSubSection id="EBC81AFEFFAF880DFF23FDAE0191FDA6" box="[151,452,544,571]" pageId="3" pageNumber="82" type="nomenclature">
<paragraph id="A36D4975FFAF880DFF23FDAE0191FDA6" blockId="3.[151,452,544,571]" box="[151,452,544,571]" pageId="3" pageNumber="82">
<heading id="F825FE19FFAF880DFF23FDAE0191FDA6" bold="true" box="[151,452,544,571]" fontSize="11" level="1" pageId="3" pageNumber="82" reason="1">
<emphasis id="91A69567FFAF880DFF23FDAE0191FDA6" bold="true" box="[151,452,544,571]" pageId="3" pageNumber="82">
Phylogeny of
<taxonomicName id="64D232F6FFAF880DFEF4FDAF0191FDA6" ID-CoL="4JPG" box="[320,452,544,570]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFEF4FDAF0191FDA6" bold="true" box="[320,452,544,570]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EBC81AFEFFAF880AFF23FDE8026CFE53" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82" type="description">
<paragraph id="A36D4975FFAF880DFF23FDE80558FCE1" blockId="3.[151,1436,615,893]" pageId="3" pageNumber="82">
The phylogeny recovered a well supported clade consisting of all
<taxonomicName id="64D232F6FFAF880DFCCEFDE703A3FDE3" box="[890,1014,616,639]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFCCEFDE703A3FDE3" box="[890,1014,616,639]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
</taxonomicName>
from South and South
<collectingCountry id="DBC509E5FFAF880DFB4FFDE70533FD1C" box="[1275,1382,615,640]" name="China" pageId="3" pageNumber="82">East Asia</collectingCountry>
(
<figureCitation id="3BE955F0FFAF880DFAC1FDE800FDFD38" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
). We also recovered two sister clades within
<taxonomicName id="64D232F6FFAF880DFD0AFD02036FFD38" box="[702,826,653,676]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFD0AFD02036FFD38" box="[702,826,653,676]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
</taxonomicName>
. The geographical ranges of the two clades broadly overlap around the Northeast Indian regions and
<collectingCountry id="DBC509E5FFAF880DFD09FD200374FD54" box="[701,801,687,712]" name="Thailand" pageId="3" pageNumber="82">Thailand</collectingCountry>
(
<figureCitation id="3BE955F0FFAF880DFC84FD20033BFD54" box="[816,878,687,712]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
). The
<taxonomicName id="64D232F6FFAF880DFC02FD3F0466FD5B" box="[950,1075,688,711]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFC02FD3F0466FD5B" box="[950,1075,688,711]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
</taxonomicName>
clade was recovered as sister to
<taxonomicName id="64D232F6FFAF880DFF23FD5B0165FD70" box="[151,304,724,748]" class="Amphibia" family="Dicroglossidae" genus="Sphaerotheca" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFF23FD5B0165FD70" box="[151,304,724,748]" italics="true" pageId="3" pageNumber="82">Sphaerotheca</emphasis>
</taxonomicName>
. The unidentified individuals in our collection formed a monophyletic group and showed a sister relationship with
<taxonomicName id="64D232F6FFAF880DFED9FD77023AFC8C" box="[365,623,760,784]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="sahyadris">
<emphasis id="91A69567FFAF880DFED9FD77023AFC8C" box="[365,623,760,784]" italics="true" pageId="3" pageNumber="82">Minervarya sahyadris</emphasis>
</taxonomicName>
(
<figureCitation id="3BE955F0FFAF880DFD33FD78029AFC8C" box="[647,719,759,784]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
). The clade consisting of the unidentified individuals and
<taxonomicName id="64D232F6FFAF880DFF23FC9201C0FCA8" box="[151,405,796,820]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="sahyadris">
<emphasis id="91A69567FFAF880DFF23FC9201C0FCA8" box="[151,405,796,820]" italics="true" pageId="3" pageNumber="82">Minervarya sahyadris</emphasis>
</taxonomicName>
was deeply nested within the larger
<taxonomicName id="64D232F6FFAF880DFCECFC920380FCA8" box="[856,981,797,820]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFCECFC920380FCA8" box="[856,981,797,820]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
</taxonomicName>
clade. Based on this, we synonymize
<taxonomicName id="64D232F6FFAF880DFF23FCCF022FFCC4" authority="Dubois, Ohler and Biju, 2001" authorityName="Dubois, Ohler and Biju" authorityYear="2001" box="[151,634,831,856]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFF23FCCF014EFCCB" box="[151,283,832,855]" italics="true" pageId="3" pageNumber="82">Minervarya</emphasis>
<bibRefCitation id="C7433484FFAF880DFE90FCB0022FFCC4" author="Dubois" box="[292,634,831,856]" pageId="3" pageNumber="82" refString="Dubois, A., Ohler, A. &amp; Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article" year="2001">Dubois, Ohler and Biju, 2001</bibRefCitation>
</taxonomicName>
under
<taxonomicName id="64D232F6FFAF880DFD78FCCF03BCFCC4" authority="Bolkay, 1915" authorityName="Bolkay" authorityYear="1915" box="[716,1001,831,856]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFAF880DFD78FCCF031DFCCB" box="[716,840,832,855]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
Bolkay, 1915
</taxonomicName>
(see discussion below) and provide a diagnosis and describe the unidentified individuals in our collection as
<taxonomicName id="64D232F6FFAF880DFC07FCEA04F9FCE0" box="[947,1196,868,892]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="gomantaki" status="sp. nov.">
<emphasis id="91A69567FFAF880DFC07FCEA04F9FCE0" box="[947,1196,868,892]" italics="true" pageId="3" pageNumber="82">Fejervarya gomantaki</emphasis>
</taxonomicName>
<emphasis id="91A69567FFAF880DFB00FCEB0558FCE1" bold="true" box="[1204,1293,868,893]" pageId="3" pageNumber="82">
<taxonomicNameLabel id="8A95281CFFAF880DFB00FCEB0558FCE1" box="[1204,1293,868,893]" pageId="3" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="F7AD19FDFFAF880DFF23F8EE0365F809" httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82" targetBox="[175,1424,957,1852]" targetPageId="3">
<paragraph id="A36D4975FFAF880DFF23F8EE0365F809" blockId="3.[151,1436,1889,1941]" pageId="3" pageNumber="82">
<emphasis id="91A69567FFAF880DFF23F8EE0146F8EA" bold="true" box="[151,275,1889,1911]" pageId="3" pageNumber="82">FIGURE 2.</emphasis>
Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16S and 12S) and nuclear genes (BDNF, Rhod, Tyr, RAG-1, RAG-2, NCX1, and CXCR4).
</paragraph>
</caption>
<paragraph id="A36D4975FFAF880AFF73F84D026CFE53" blockId="3.[199,1436,1986,2011]" lastBlockId="4.[151,1436,151,464]" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82">
<emphasis id="91A69567FFAF880DFF73F84D01E5F847" bold="true" box="[199,432,1986,2011]" pageId="3" pageNumber="82">Generic allocation.</emphasis>
There is considerable confusion in generic level taxonomy and systematics within dicroglossid frogs, specifically the distinction between
<taxonomicName id="64D232F6FFA8880AFC95FF1703C8FF33" box="[801,925,152,175]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFC95FF1703C8FF33" box="[801,925,152,175]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
</taxonomicName>
,
<taxonomicName id="64D232F6FFA8880AFC05FF170460FF33" box="[945,1077,152,175]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFC05FF170460FF33" box="[945,1077,152,175]" italics="true" pageId="4" pageNumber="83">Minervarya</emphasis>
</taxonomicName>
and
<taxonomicName id="64D232F6FFA8880AFBC2FF1704B4FF2C" box="[1142,1249,152,176]" class="Amphibia" family="Dicroglossidae" genus="Zakerana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFBC2FF1704B4FF2C" box="[1142,1249,152,176]" italics="true" pageId="4" pageNumber="83">Zakerana</emphasis>
</taxonomicName>
(see discussion below). Apart from the strong phylogenetic support (
<figureCitation id="3BE955F0FFA8880AFD5EFF33037DFF48" box="[746,808,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="4" pageNumber="83">Fig 2</figureCitation>
), the individuals are assignable to the genus
<taxonomicName id="64D232F6FFA8880AFAABFF3205C9FF48" box="[1311,1436,189,212]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFAABFF3205C9FF48" box="[1311,1436,189,212]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
</taxonomicName>
based on the diagnosable generic morphological characters assigned by
<bibRefCitation id="C7433484FFA8880AFC63FF500495FF64" box="[983,1216,223,248]" pageId="4" pageNumber="83" refString="Dubois, A., Ohler, A. &amp; Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article">
Dubois
<emphasis id="91A69567FFA8880AFB85FF6E0439FF64" box="[1073,1132,224,248]" italics="true" pageId="4" pageNumber="83">et al.</emphasis>
(2001)
</bibRefCitation>
for the then genus
<taxonomicName id="64D232F6FFA8880AFF23FE8A0224FE80" authority="Dubois, Ohler and Biju, 2001" authorityName="Dubois, Ohler and Biju" authorityYear="2001" box="[151,625,260,285]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFF23FE8A014EFE80" box="[151,283,261,284]" italics="true" pageId="4" pageNumber="83">Minervarya</emphasis>
<bibRefCitation id="C7433484FFA8880AFE97FE8B0224FE80" author="Dubois" box="[291,625,260,285]" pageId="4" pageNumber="83" refString="Dubois, A., Ohler, A. &amp; Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article" year="2001">Dubois, Ohler and Biju, 2001</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="64D232F6FFA8880AFD1DFE8A0393FE81" authority="Bolkay, 1915" authorityName="Bolkay" authorityYear="1915" box="[681,966,260,285]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFD1DFE8A0373FE80" box="[681,806,261,284]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
Bolkay, 1915
</taxonomicName>
; namely to the group of small sized frogs having snout not greatly pointed or slightly rounded; relatively small tympanum; small rounded or laterally compressed internal metatarsal tubercles; rudimentary webbing on feet and small tibia; presence of rictal glands, fejervaryan lines, and horizontal band on the upper lip. Within
<taxonomicName id="64D232F6FFA8880AFCD1FEFF03B4FE1B" box="[869,993,368,391]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="91A69567FFA8880AFCD1FEFF03B4FE1B" box="[869,993,368,391]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
</taxonomicName>
, presence of a rictal gland is a derived character unique to the clade composed of
<taxonomicName id="64D232F6FFA8880AFD2CFE1A03D8FE30" box="[664,909,404,428]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="sahyadris">
<emphasis id="91A69567FFA8880AFD2CFE1A03D8FE30" box="[664,909,404,428]" italics="true" pageId="4" pageNumber="83">Fejervarya sahyadris</emphasis>
</taxonomicName>
,
<taxonomicName id="64D232F6FFA8880AFC14FE1A04CBFE30" box="[928,1182,404,428]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="gomantaki" status="sp. nov.">
<emphasis id="91A69567FFA8880AFC14FE1A04CBFE30" box="[928,1182,404,428]" italics="true" pageId="4" pageNumber="83">Fejervarya gomantaki</emphasis>
</taxonomicName>
<emphasis id="91A69567FFA8880AFB1EFE1B055CFE31" bold="true" box="[1194,1289,404,429]" pageId="4" pageNumber="83">
<taxonomicNameLabel id="8A95281CFFA8880AFB1EFE1B055CFE31" box="[1194,1289,404,429]" pageId="4" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and a north eastern species
<taxonomicName id="64D232F6FFA8880AFEF1FE370267FE4C" box="[325,562,440,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="chilapata">
<emphasis id="91A69567FFA8880AFEF1FE370267FE4C" box="[325,562,440,464]" italics="true" pageId="4" pageNumber="83">Fejervarya chilapata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>