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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="2B6B6F92021F6D12F2F9EBCF4B07EB46" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Allocota cyanipennis Heller 1923" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="53" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="52" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
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<mods:detail type="volume">
<mods:number>284</mods:number>
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<mods:start>1</mods:start>
<mods:end>129</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
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<treatment ID-GBIF-Taxon="152042275" LSID="urn:lsid:plazi:treatment:2B6B6F92021F6D12F2F9EBCF4B07EB46" httpUri="http://treatment.plazi.org/id/2B6B6F92021F6D12F2F9EBCF4B07EB46" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<subSubSection pageId="51" pageNumber="52" type="nomenclature">
<paragraph pageId="51" pageNumber="52">
<taxonomicName LSID="2B6B6F92-021F-6D12-F2F9-EBCF4B07EB46" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Allocota cyanipennis" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="cyanipennis">Allocota cyanipennis Heller, 1923</taxonomicName>
<figureCitation captionStart="Figures 13-18" captionStartId="F3" captionText="Figures 13 - 18. Habitus, scale bars = 2.0 mm: 13 Diamella cupreomicans (Oberthuer), a specimen from Hainan 14 Allocota viridipennis Motschulsky, a specimen from Java 15 Allocota viridipennis Motschulsky, a specimen from Singapore 16 Allocota cyanipennis Heller, a specimen from Philippines 17 Allocota bicolor sp. n., a paratype from Hainan 18 Allocota aurata (Bates), a specimen from Japan." httpUri="https://binary.pensoft.net/fig/29675" pageId="51" pageNumber="52">Habitus: Figs 16</figureCitation>
<figureCitation captionStart="Figures 4954" captionStartId="F9" captionText="Figures 49 - 54. Type materials, scale bars = 2.0 mm: 49 Lectotype of Allocota caerulea Andrewes [= Allocota viridipennis Motschulsky] 50 Holotype of Allocota cyanipennis Heller 51 Lectotype of Taicona aurata Bates [= Allocota aurata (Bates)] 52 Holotype of Allocota bicolor sp. n. 53 Syntype of Dasiosoma testaceum Britton 54 Holotype of Dasiosoma hirsutum Basilewsky [= Dasiosoma basilewskyi nom. n.]." httpUri="https://binary.pensoft.net/fig/29682" pageId="51" pageNumber="52">50</figureCitation>
<figureCitation captionStart="Figure 81" captionStartId="F26" captionText="Figure 81. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Allocota cyanipennis Heller, a specimen from Philippines." httpUri="https://binary.pensoft.net/fig/29699" pageId="51" pageNumber="52">male genitalia: Fig. 81</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="reference_group">
<paragraph pageId="51" pageNumber="52">
<taxonomicName authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="cyanipennis">Allocota cyanipennis</taxonomicName>
<bibRefCitation author="Heller, KM" journalOrPublisher="The Philippine Journal of Science" pageId="87" pageNumber="88" pagination="295 - 307" refId="B52" refString="Heller, KM, 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 307" title="Some new Malayan Carabidae, especially Philippine." volume="23" year="1923">Heller 1923</bibRefCitation>
: 305 (original:
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota</emphasis>
</taxonomicName>
; type locality: Mindanao; holotype deposited in SNSD);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Zoologische Mededeelingen, Leiden" pageId="85" pageNumber="86" refId="B14" refString="Andrewes, HE, 1930d. Catalogue of Indian Insects, Part 18-Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." title="Catalogue of Indian Insects, Part 18 - Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." year="1930 d">Andrewes 1930d</bibRefCitation>
: 13 (
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota</emphasis>
</taxonomicName>
; catalogue);
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 306 (
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota</emphasis>
</taxonomicName>
);
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="757 - 799" refId="B62" refString="Kirschenhofer, E, 1996. Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil. Linzer Biologische Beitraege 28 (2): 757 - 799" title="Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil." volume="28" year="1996">Kirschenhofer 1996</bibRefCitation>
: 763 (
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="type examined">
<paragraph pageId="51" pageNumber="52">Type examined.</paragraph>
<paragraph pageId="51" pageNumber="52">
<emphasis bold="true" pageId="51" pageNumber="52">Holotype</emphasis>
of
<taxonomicName authorityName="Heller" authorityYear="1923" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="cyanipennis">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota cyanipennis</emphasis>
</taxonomicName>
Heller, monotypy (SNSD): male, body length = 6.3 mm, board mounted, &quot;Tangcolan / Bukidnon /Baker&quot;; &quot;
<emphasis italics="true" pageId="51" pageNumber="52">cyanipennis /</emphasis>
Typus&quot; [red label]; &quot;
<emphasis italics="true" pageId="51" pageNumber="52">14261</emphasis>
&quot;; &quot;1921 /
<emphasis italics="true" pageId="51" pageNumber="52">3</emphasis>
&quot; [yellow label]; &quot;Staatl. Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
/ Tierkunde, Dresden&quot;[Fig. 50].
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="notes on types">
<paragraph pageId="51" pageNumber="52">Notes on types.</paragraph>
<paragraph pageId="51" pageNumber="52">
<bibRefCitation author="Heller, KM" journalOrPublisher="The Philippine Journal of Science" pageId="87" pageNumber="88" pagination="295 - 307" refId="B52" refString="Heller, KM, 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 307" title="Some new Malayan Carabidae, especially Philippine." volume="23" year="1923">Heller (1923)</bibRefCitation>
<normalizedToken originalValue="didnt">didn't</normalizedToken>
clearly state this species was based on a single specimen, but he only cited one specimen with a serial number
<normalizedToken originalValue="“14261”">&quot;14261&quot;</normalizedToken>
. So, the holotype (
<figureCitation captionStart="Figures 4954" captionStartId="F9" captionText="Figures 49 - 54. Type materials, scale bars = 2.0 mm: 49 Lectotype of Allocota caerulea Andrewes [= Allocota viridipennis Motschulsky] 50 Holotype of Allocota cyanipennis Heller 51 Lectotype of Taicona aurata Bates [= Allocota aurata (Bates)] 52 Holotype of Allocota bicolor sp. n. 53 Syntype of Dasiosoma testaceum Britton 54 Holotype of Dasiosoma hirsutum Basilewsky [= Dasiosoma basilewskyi nom. n.]." httpUri="https://binary.pensoft.net/fig/29682" pageId="51" pageNumber="52">Fig. 50</figureCitation>
) was originally fixed by monotypy, as the description implies a single specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="non-type material examined">
<paragraph pageId="51" pageNumber="52">Non-type material examined</paragraph>
<paragraph pageId="51" pageNumber="52">
(Total 5 specimens).
<emphasis bold="true" pageId="51" pageNumber="52">The</emphasis>
<emphasis bold="true" pageId="51" pageNumber="52">Philippines</emphasis>
: 1 specimen (NHML), &quot;Philippine Is., Coll. Bottcher., B. M. 1929-201&quot;. 1 male (NHML), &quot;Island of Basilan, Baker&quot;; &quot;Ex Mus. Coll. Agric. Phil. Is.&quot;; &quot;Allocota cyanipennis Heller, Compared with type H. E. A.&quot;[Figs 16, 81]. 1 specimen (NHML), &quot;Tangcolan, Bukienon, Baker&quot;.
<emphasis bold="true" pageId="51" pageNumber="52">Sulawesi</emphasis>
: 1 male and 1 female (NHML),
<normalizedToken originalValue="“Celebes”">&quot;Celebes&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="diagnosis">
<paragraph pageId="51" pageNumber="52">Diagnosis.</paragraph>
<paragraph pageId="51" pageNumber="52">
Elytra metallic blue; metasternum and abdomen much darker than prosternum; front angles of pronotum with setae relative long (
<figureCitation captionStart="Figures 138157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="51" pageNumber="52">Fig. 152</figureCitation>
); elytral 7th interval without setigerous pores; internal sac of aedeagus with setose area divided into three parts, apex of secondary flagellum forming a large triangular sclerite.
</paragraph>
<paragraph pageId="51" pageNumber="52">
This species is closest to
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis italics="true" pageId="51" pageNumber="52">Allocota viridipennis</emphasis>
</taxonomicName>
Motschulsky; differences are presented in the diagnosis part under the latter species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
<paragraph pageId="51" pageNumber="52">Description.</paragraph>
<paragraph lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
Body length 6.3-7.1 mm; head and pronotum orange red or brown; mouthparts earth yellow, palpomeres brownish, apex of terminal palpomeres yellow; 1st antennomere reddish, the remaining antennomeres usually slightly darker; elytra metallic blue, elytral suture and lateral margins metallic; legs usually much darker than pronotum, darkest on apical half of femora, basal half usually the same color as pronotum; ventral side of head and prosternum the same color as dorsal side; metasternum and abdomen brownish to piceous, much darker than prosternum.
<emphasis bold="true" pageId="51" pageNumber="52">Head</emphasis>
glabrous, without punctures, microsculpture indistinct; males with terminal labial palpomeres fusiform, slightly expanded in middle.
<emphasis bold="true" pageId="51" pageNumber="52">Pronotum</emphasis>
glabrous,cordiform, widest at anterior third; ratio PW/PL 1.28 to 1.32; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles with setae relative long (
<figureCitation captionStart="Figures 138157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="51" pageNumber="52">Fig. 152</figureCitation>
); lateral margins rounded, slightly expanded in middle,
<pageBreakToken pageId="52" pageNumber="53" start="start">strongly</pageBreakToken>
sinuate before hind angles; lateral explanate areas moderately wide, without punctures; hind angles slightly acute, less distinct; basal foveae very shallow, without punctures; median line very fine, usually indistinct.
<emphasis bold="true" pageId="52" pageNumber="53">Elytra</emphasis>
with striae very shallow, finely punctate; intervals nearly flat, without accessory setae, with a row of very fine punctures; microsculpture absent; 3rd and 5th intervals with four to ten setigerous pores, their positions variable; 7th interval without setigerous pores; setae on lateral margins fine and short, but more distinct than in
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis italics="true" pageId="52" pageNumber="53">Allocota viridipennis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="52" pageNumber="53">Legs.</emphasis>
Protibiae with cleaning spur absent (
<figureCitation captionStart="Figures 138157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="52" pageNumber="53">Fig. 140</figureCitation>
); males with adhesive hairs well developed (two whole rows) on first two mesotarsomeres, rudimentary (two rows, weakly present near apex) on 3rd mesotarsomere.
<emphasis bold="true" pageId="52" pageNumber="53">Male genitalia</emphasis>
with median lobe of aedeagus stout, more distinctly bent to right side near apex than in
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis italics="true" pageId="52" pageNumber="53">Allocota viridipennis</emphasis>
</taxonomicName>
in dorsal view, right margin slightly curved before apical lamella; apical lamella placed on right side, narrow, slightly elongate; internal sac with setose area divided into three parts; apex of secondary flagellum expanded, formed a large triangular sclerite; trumpet-form expansion with ventral margin more or less expanded. Female genitalia not studied (
<figureCitation captionStart="Figure 81" captionStartId="F26" captionText="Figure 81. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Allocota cyanipennis Heller, a specimen from Philippines." httpUri="https://binary.pensoft.net/fig/29699" pageId="52" pageNumber="53">Fig. 81</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="distribution">
<paragraph pageId="52" pageNumber="53">Distribution</paragraph>
<paragraph pageId="52" pageNumber="53">
(
<figureCitation captionStart="Map 6" captionStartId="F50" captionText="Map 6. Known distribution of genus Allocota: ● Allocota aurata (Bates) ★ Allocota cyanipennis Heller ▲ Allocota viridipennis Motschulsky ◆ Allocota bicolor sp. n." httpUri="https://binary.pensoft.net/fig/29728" pageId="52" pageNumber="53">Map 6</figureCitation>
). The Philippines, Sulawesi.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="geographical variation">
<paragraph pageId="52" pageNumber="53">Geographical variation.</paragraph>
<paragraph pageId="52" pageNumber="53">Pronotum and head of two specimens from Sulawesi we examined are brown, much darker than those from the Philippines which are vivid orange.</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="remarks">
<paragraph pageId="52" pageNumber="53">Remarks.</paragraph>
<paragraph pageId="52" pageNumber="53">
We regard specimens from the Philippines and Sulawesi as a distinct species from
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis italics="true" pageId="52" pageNumber="53">Allocota viridipennis</emphasis>
</taxonomicName>
Motschulsky for the constant difference in internal sac of aedeagus as mentioned above in the diagnosis section under
<taxonomicName authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis italics="true" pageId="52" pageNumber="53">Allocota viridipennis</emphasis>
</taxonomicName>
. In external characters these two species are very similar, with only slight differences in setae on pronotal front angles (
<figureCitation captionStart="Figures 138157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="52" pageNumber="53">Figs 151, 152</figureCitation>
) and elytral lateral margins.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.423.7536" ID-GBIF-Dataset="1f17876d-2290-4d36-aa1f-10b951701679" ID-PMC="PMC4106094" ID-Pensoft-Pub="1313-2970-423-1" ID-PubMed="25061393" ID-ZBK="55B4F9C858934F88841660FF730E8872" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-423-1" ModsDocOrigin="ZooKeys 423" ModsDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" checkinTime="1451245649604" checkinUser="pensoft" docAuthor="Brunke, Adam J. &amp; Solodovnikov, Alexey" docDate="2014" docId="2B6BA06A9B404CCDE6DF5AA721670BA2" docLanguage="en" docName="ZooKeys 423: 1-113" docOrigin="ZooKeys 423" docSource="http://dx.doi.org/10.3897/zookeys.423.7536" docTitle="Bolitogyrus apicofasciatus Brunke, sp. n." docType="treatment" docUuid="CD87F64C-3FF2-4E43-AA32-9F1197BC15C6" docUuidSource="ZooBank" docVersion="4" lastPageNumber="36" masterDocId="FFAB12065F4CFFED925DFFB46A5FFFFB" masterDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" masterLastPageNumber="113" masterPageNumber="1" pageNumber="35" updateTime="1668158857111" updateUser="ExternalLinkService">
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<mods:title>A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Brunke, Adam J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Alexey</mods:namePart>
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<mods:number>423</mods:number>
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<treatment ID-GBIF-Taxon="152054119" LSID="urn:lsid:zoobank.org:act:CD87F64C-3FF2-4E43-AA32-9F1197BC15C6" httpUri="http://treatment.plazi.org/id/2B6BA06A9B404CCDE6DF5AA721670BA2" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection pageId="34" pageNumber="35" type="multiple">
<paragraph pageId="34" pageNumber="35">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="nomenclature">
<paragraph pageId="34" pageNumber="35">
<taxonomicName LSID="http://zoobank.org/CD87F64C-3FF2-4E43-AA32-9F1197BC15C6" authority="Brunke" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus apicofasciatus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="apicofasciatus">Bolitogyrus apicofasciatus Brunke</taxonomicName>
<taxonomicNameLabel pageId="34" pageNumber="35">sp. n.</taxonomicNameLabel>
Figs 6D, 11H, 14B, 18
<normalizedToken originalValue="FH">F-H</normalizedToken>
, 23O, 26L, 30B (map)
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="type locality">
<paragraph pageId="34" pageNumber="35">Type locality.</paragraph>
<paragraph pageId="34" pageNumber="35">MEXICO, Chiapas, 2 km SE Custepec.</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="type material">
<paragraph pageId="34" pageNumber="35">Type material.</paragraph>
<paragraph pageId="34" pageNumber="35">
Holotype ♂ (SEMC): MEXICO, Chiapas, 2 km SE Custepec,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.72219">15.72219°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-92.9464">92.94640°W</geoCoordinate>
, 1610 m, 19-V-2008, beating vegetation, cloud forest, LLAMA08 Go-A-02-3-03 [white printed label] / SM0822854 [white barcode label] / SUBJECT TO RETENTION AGREEMENT w/ ECOSUR, see KUNHM acc. # 2010-EN-089, to be removed with fulfilled [green printed label] / Holotype,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus apicofasciatus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="apicofasciatus">Bolitogyrus apicofasciatus</taxonomicName>
Brunke, sp. n. [red printed label].
</paragraph>
<paragraph pageId="34" pageNumber="35">
Paratypes (3♀ SEMC): same as holotype except:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.72236">15.72236°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-92.94585">92.94585°W</geoCoordinate>
, 1620 m, mesophil forest, LLAMA08 Go-A-02-3-02, SM0839743 (SEMC). GUATEMALA:Quetzaltenango, 12.5 km SW Zunil, 1520 m, 21 June 1993, R. Anderson, #93-11, second-growth, SM0037995, 1 ♀ (SEMC) [missing head and pronotum], same except: 20 June 1993, J. Ashe, R. Brooks, #067, ex.
<taxonomicName class="Holothuroidea" family="Synaptidae" genus="Auricularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Auricularia" order="Apodida" pageId="34" pageNumber="35" phylum="Echinodermata" rank="genus">Auricularia</taxonomicName>
sp., SM0037984, 1 ♀ (SEMC).
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="diagnosis">
<paragraph pageId="34" pageNumber="35">Diagnosis.</paragraph>
<paragraph pageId="34" pageNumber="35">Within Bullatus Lineage: one setose puncture in dorsal row of pronotum; elytra dark, without bright metallic reflection; scutellum dark, not reddish; elytra without strigose patch; femur with dark band separated from the apex by a distance distinctly shorter than half the length of the band (Fig. 11H).</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="description">
<paragraph pageId="35" pageNumber="36">
<pageBreakToken pageId="35" pageNumber="36" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="35" pageNumber="36">Measurements ♂ (n=1): HW/HL 1.45; PW/PL 1.50; EW/EL 1.14; ESut/PL 1.00; PW/HW 1.03; forebody length 3.5 mm.</paragraph>
<paragraph pageId="35" pageNumber="36">Measurements ♀ (n=2): HW/HL 1.41-1.43; PW/PL 1.36-1.45; EW/EL 1.18-1.19; ESut/PL 0.92-1.00; PW/HW 1.07-1.10; forebody length 4.2-4.5 mm.</paragraph>
<paragraph pageId="35" pageNumber="36">
Similar to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus bullatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bullatus">Bolitogyrus bullatus</taxonomicName>
and differing only in the following: pronotum dark brown throughout, abdominal segments (except VIII) broadly paler at apex, without paler lateral stripes, antennomeres
<normalizedToken originalValue="IX">I-X</normalizedToken>
grading from reddish-yellow (I) to dark brown, apical segment not or barely paler than previous, brown to dark brown; procoxa light yellow, meso- and metacoxa yellowish-brown (male) or darker, brown (female); profemur without (male) or with short, nearly apical dark band, meso and metafemur with dark band close to apex, band on metafemur separated from the apex by a distance distinctly shorter than half the length of the band; median frontal impression present as in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus bullatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bullatus">Bolitogyrus bullatus</taxonomicName>
but sometimes obscured by irregular sculpture; central protuberance without fine median sulcus, base of head without distinct posterior protuberances, punctures of head impressed but these impressions not forming sulci, disc of pronotum markedly glossy, with very few punctures; protuberance of pronotum only weakly visible in lateral view; macrosetae of elytra relatively short, not distinct from overall surface sculpture in lateral view; abdominal sternites with relatively coarse microsculpture, interspaces wider than lines; median lobe in lateral view gently curved ventrad from base, apical portion more slender and more strongly curved ventrad near apex (Fig. 18G), similar to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus bullatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bullatus">Bolitogyrus bullatus</taxonomicName>
in parameral view (Fig. 18F), paramere in parameral view not divided apically but with median suture, distinctly extended to midlength (Fig. 18F), paramere slightly shorter than median lobe; paramere with peg setae arranged in a pair of lateral rows, which are wider (2-3 peg setae wide) near apex and base of row (Fig. 18H); male sternite VIII with transverse basal line complete; male sternite IX more densely setose, asymmetrical at base and with much shallower emargination at apex than
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus bullatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bullatus">Bolitogyrus bullatus</taxonomicName>
(Fig. 23O); female tergite X strongly constricted at base, with broadly rounded apex (Fig. 26L); basal margin of laterotergal sclerites fused and thickened across base of tergite X (Fig. 26L); female laterotergal sclerites strongly expanded at base and overlapping with tergite X (Fig. 26L).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="distribution">
<paragraph pageId="35" pageNumber="36">Distribution.</paragraph>
<paragraph pageId="35" pageNumber="36">Figure 30B. Known from Chiapas, Mexico and Quetzaltenango, Guatemala.</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="bionomics">
<paragraph pageId="35" pageNumber="36">Bionomics.</paragraph>
<paragraph pageId="35" pageNumber="36">
Specimens were collected in cloud and mesophil forests at 1520-1620 m, by beating vegetation and from an
<taxonomicName class="Holothuroidea" family="Synaptidae" genus="Auricularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Auricularia" order="Apodida" pageId="35" pageNumber="36" phylum="Echinodermata" rank="genus">Auricularia</taxonomicName>
(jelly) fungus, during May and June.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="etymology">
<paragraph pageId="35" pageNumber="36">Etymology.</paragraph>
<paragraph pageId="35" pageNumber="36">
The specific epithet means 'apical
<normalizedToken originalValue="band">band'</normalizedToken>
and refers to the dark color band of the hind femur, which is situated distinctly closer to the apex than in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus bullatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bullatus">Bolitogyrus bullatus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="comments">
<paragraph pageId="35" pageNumber="36">Comments.</paragraph>
<paragraph pageId="35" pageNumber="36">
Within its distribution,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus apicofasciatus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="apicofasciatus">Bolitogyrus apicofasciatus</taxonomicName>
cannot be confused with any described species of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Bolitogyrus</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Distribution of wild bee (Hymenoptera: Anthophila) and hoverfly (Diptera: Syrphidae) communities within farms undergoing ecological transition</mods:title>
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<mods:namePart>Noel, Gregoire</mods:namePart>
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<mods:namePart>Bonnet, Julie</mods:namePart>
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<treatment LSID="urn:lsid:plazi:treatment:2B6BC5BE26CA50EDA3CB0A6837186BBF" httpUri="http://treatment.plazi.org/id/2B6BC5BE26CA50EDA3CB0A6837186BBF" lastPageNumber="60665" pageId="0" pageNumber="60665">
<subSubSection pageId="0" pageNumber="60665" type="nomenclature">
<paragraph pageId="0" pageNumber="60665">
<taxonomicName LSID="2B6BC5BE-26CA-50ED-A3CB-0A6837186BBF" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Hexapoda" family="Syrphidae" genus="Sphaerophoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerophoria scripta" order="Hymenoptera" pageId="0" pageNumber="60665" phylum="Arthropoda" rank="species" species="scripta">Sphaerophoria scripta (Linnaeus 1758)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="60665" type="ecological interactions">
<paragraph pageId="0" pageNumber="60665">Ecological interactions</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="60665" type="feeds on">
<paragraph pageId="0" pageNumber="60665">Feeds on</paragraph>
<paragraph pageId="0" pageNumber="60665">Polylectic</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="60665" type="conservation status">
<paragraph pageId="0" pageNumber="60665">Conservation status</paragraph>
<paragraph pageId="0" pageNumber="60665">Not Applicable</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="60665" type="notes">
<paragraph pageId="0" pageNumber="60665">Notes</paragraph>
<paragraph pageId="0" pageNumber="60665">
Table
<tableCitation captionStart="Table 2" captionStartId="T5947970" captionText="Table 2. Abundance of each pollinator species according to the habitat of its collection. The habitat details are given in Table 1." httpUri="http://table.plazi.org/id/38825501005D93B791340A54111FC5E7" pageId="0" pageNumber="60665" tableUuid="38825501005D93B791340A54111FC5E7">2</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>