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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8051" ID-PMC="PMC5018104" ID-Pensoft-Pub="1314-2828-4-8051" ID-PubMed="27660527" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8051" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="The cicadas (Hemiptera: Cicadidae) of India, Bangladesh, Bhutan, Myanmar, Nepal and Sri Lanka: an annotated provisional catalogue, regional checklist and bibliography" checkinTime="1469054439337" checkinUser="pensoft" docAuthor="Price, Benjamin Wills, Allan, Elizabeth Louise, Marathe, Kiran, Sarkar, Vivek, Simon, Chris &amp; Kunte, Krushnamegh" docDate="2016" docId="2B2119B905CFB7A83795478912F5528C" docLanguage="en" docName="BiodivDatJour 4: e8051" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8051" docTitle="Haphsa bindusara Distant 1881" docType="treatment" docVersion="3" lastPageNumber="8051" masterDocId="2F66FFD9FFEBFFD3FF85AF11137AFFE4" masterDocTitle="The cicadas (Hemiptera: Cicadidae) of India, Bangladesh, Bhutan, Myanmar, Nepal and Sri Lanka: an annotated provisional catalogue, regional checklist and bibliography" masterLastPageNumber="8051" masterPageNumber="8051" pageNumber="8051" updateTime="1668126878091" updateUser="ExternalLinkService">
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<mods:title>The cicadas (Hemiptera: Cicadidae) of India, Bangladesh, Bhutan, Myanmar, Nepal and Sri Lanka: an annotated provisional catalogue, regional checklist and bibliography</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Price, Benjamin Wills</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Allan, Elizabeth Louise</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Marathe, Kiran</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sarkar, Vivek</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Simon, Chris</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kunte, Krushnamegh</mods:namePart>
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<mods:title>Biodiversity Data Journal</mods:title>
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<mods:date>2016</mods:date>
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<mods:number>4</mods:number>
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<subSubSection pageId="0" pageNumber="8051" type="nomenclature">
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<taxonomicName authority="Distant, 1881" authorityName="Distant" authorityYear="1881" class="Insecta" family="Cicadidae" genus="Haphsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haphsa bindusara" order="Hemiptera" pageId="0" pageNumber="8051" phylum="Arthropoda" rank="species" species="bindusara">Haphsa bindusara (Distant, 1881)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="8051" type="reference_group">
<paragraph pageId="0" pageNumber="8051">
<taxonomicName class="Insecta" family="Cicadidae" genus="Pomponia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pomponia bindusara" order="Hemiptera" pageId="0" pageNumber="8051" phylum="Arthropoda" rank="species" species="bindusara">Pomponia bindusara</taxonomicName>
Distant, 1881
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8051" type="materials_examined">
<paragraph pageId="0" pageNumber="8051">Materials</paragraph>
<paragraph pageId="0" pageNumber="8051">
<materialsCitation collectionCode="NHMUK" country="Myanmar" location="Upper Tenasserim" pageId="0" pageNumber="8051" specimenCode="BMNH (E) 1009459" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="8051">Holotype</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="8051">BMNH(E) 1009459</specimenCode>
; individualCount:
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; sex:
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; Taxon: scientificName: Haphsabindusara (Distant, 1881); Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="8051">Myanmar</collectingCountry>
; locality:
<location LSID="urn:lsid:plazi:treatment:2B2119B905CFB7A83795478912F5528C:29CA7942724316109EA63D2EF10DA6E4" country="Myanmar" name="Upper Tenasserim" pageId="0" pageNumber="8051">Upper Tenasserim</location>
; Record Level: institutionCode:
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; basisOfRecord: PreservedSpecimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8051" type="distribution">
<paragraph pageId="0" pageNumber="8051">Distribution</paragraph>
<paragraph pageId="0" pageNumber="8051">[Distant, 1906] Burma: Teinzo; Karen Hills. Upper Tenasserim. [Metcalf, 1963] Tenasserim; India; Burma; Laos; Indochina. [Sanborn, 2014] Pakistan (?), Burma, India, Yunnan, China, Laos, Thailand, Tenasserim, Indochina, North Vietnam, Myanmar, Vietnam, Bangladesh (?).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8051" type="notes">
<paragraph pageId="0" pageNumber="8051">Notes</paragraph>
<paragraph pageId="0" pageNumber="8051">
Authority:
<bibRefCitation author="Distant, William Lucas" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="8051" pagination="627 - 648" title="Descriptions of new species belonging to the Homopterous Family Cicadidae." volume="1881" year="1881">Distant 1881</bibRefCitation>
;
<bibRefCitation author="Metcalf, Z. P." journalOrPublisher="North Carolina State College Contribution" pageId="0" pageNumber="8051" pagination="587 - 919" title="General catalogue of the Homoptera. Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeaninae and Cicadinae." volume="1502" year="1963">Metcalf (1963)</bibRefCitation>
stated &quot;India&quot; in reference to
<bibRefCitation author="Atkinson, E. T." journalOrPublisher="Journal of the Asiatic Society Bengal" pageId="0" pageNumber="8051" pagination="143 - 223" title="Notes on Indian Rhynchota, No. 6. Addenda and index." volume="55" year="1886">Atkinson (1886)</bibRefCitation>
who was referring to the type specimen from Upper Tenasserim (Myanmar).
<bibRefCitation author="Metcalf, Z. P." journalOrPublisher="North Carolina State College Contribution" pageId="0" pageNumber="8051" pagination="587 - 919" title="General catalogue of the Homoptera. Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeaninae and Cicadinae." volume="1502" year="1963">Metcalf (1963)</bibRefCitation>
also listed India in reference to
<bibRefCitation author="Mathur, R. N." journalOrPublisher="Indian Forest Leaflet (ent.)" pageId="0" pageNumber="8051" pagination="138 - 187" title="A systematic catalogue of the main identified entomological collection at the Forest Research Institute, Dehra Dun. Part 21. Order Hemiptera (Continued)." volume="121" year="1953">Mathur (1953)</bibRefCitation>
who stated &quot;India.-Burma: Katha, Mohnyin Res.&quot; as a part of then &quot;British India&quot;. Listings of India since then are likely to be in reference to published localities, most likely that from
<bibRefCitation author="Metcalf, Z. P." journalOrPublisher="North Carolina State College Contribution" pageId="0" pageNumber="8051" pagination="587 - 919" title="General catalogue of the Homoptera. Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeaninae and Cicadinae." volume="1502" year="1963">Metcalf (1963)</bibRefCitation>
. In addition,
<bibRefCitation author="Sanborn, A" journalOrPublisher="Elsevier" pageId="0" pageNumber="8051" title="Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha)" year="2014">Sanborn (2014)</bibRefCitation>
listed Pakistan and Bangladesh in reference to a specimen collected in Chittagong (former &quot;East Pakistan&quot; now Bangladesh), however the identification of the specimen from
<bibRefCitation author="Chaudhry, G. - U." journalOrPublisher="Biological Sciences Research Division (Forest Entomology Branch), Pakistan Forest Institute, Peshwar" pageId="0" pageNumber="8051" title="Survey of insect fauna of forests of Pakistan. Volume II." year="1970">Chaudhry et al. (1970)</bibRefCitation>
(
<taxonomicName class="Insecta" family="Cicadidae" genus="Aola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aola" order="Hemiptera" pageId="0" pageNumber="8051" phylum="Arthropoda" rank="genus">Aola</taxonomicName>
sp.?
<taxonomicName class="Insecta" family="Erebidae" genus="Homoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homoptera bindusara" order="Lepidoptera" pageId="0" pageNumber="8051" phylum="Arthropoda" rank="species" species="bindusara">bindusara</taxonomicName>
) has not been confirmed thus the Bangladesh distribution is not conclusive at this point.
</paragraph>
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Largest record of
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<emphasis id="91FCEAA2FFFAF25FFEE24475FDC7FD60" bold="true" box="[366,614,635,662]" italics="true" pageId="5" pageNumber="143">Lophiodes triradiatus</emphasis>
(
<bibRefCitation id="C7194B41FFFAF25FFDF94472FD5CFD60" author="Lloyd, R. E." box="[629,765,636,662]" pageId="5" pageNumber="143" pagination="139 - 180" refId="ref4818" refString="Lloyd, R. E. (1909) A description of the deep-sea fish caught by the R. I. M. S. ship`Investigator' since the year 1900, with supposed evidence of mutation in Malthopsis. Memoirs of the Indian Museum, 2 (3), 139 - 180., pls. 44 - 50." type="journal article" year="1909">Lloyd 1909</bibRefCitation>
)
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<paragraph id="A33736B0FFFAF25FFF4B4491FBF1FCB5" blockId="5.[151,1437,635,976]" pageId="5" pageNumber="143">
<bibRefCitation id="C7194B41FFFAF25FFF4B4491FE20FD4F" author="Ho, H. - C. &amp; Seret, B. &amp; Shao, K. - T." box="[199,385,671,697]" pageId="5" pageNumber="143" pagination="62 - 68" refId="ref4433" refString="Ho, H. - C., Seret, B. &amp; Shao, K. - T. (2009) Redescription of Lophiodes infrabrunneus Smith and Radcliffe, 1912, a senior synonym of L. abdituspinus Ni, Wu and Li, 1990 (Lophiiformes: Lophiidae). Zootaxa, 2326 (1), 62 - 68. https: // doi. org / 10.11646 / zootaxa. 2326.1.5" type="journal article" year="2009">
Ho
<emphasis id="91FCEAA2FFFAF25FFF794491FE8AFD4F" box="[245,299,671,697]" italics="true" pageId="5" pageNumber="143">et al</emphasis>
. (2009
</bibRefCitation>
,
<bibRefCitation id="C7194B41FFFAF25FFE1D4491FE6AFD4F" author="Ho, H. - C. &amp; Bineesh, K. K. &amp; Akhilesh, K. V." box="[401,459,671,697]" pageId="5" pageNumber="143" pagination="587 - 592" refId="ref4360" refString="Ho, H. - C., Bineesh, K. K. &amp; Akhilesh, K. V. (2014) Rediscovery of Lophiodes triradiatus (Lloyd, 1909), a senior synonym of L. infrabrunneus Smith and Radcliffe (Lophiiformes: Lophiidae). Zootaxa, 3786 (5), 587 - 592. https: // doi. org / 10.11646 / zootaxa. 3786.5.6" type="journal article" year="2014">2014</bibRefCitation>
) examined a number of specimens and synonymized two junior synonyms with
<taxonomicName id="64884D33FFFAF25FFA0A4491FEAEFD2D" baseAuthorityName="Lloyd" baseAuthorityYear="1909" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="triradiatus">
<emphasis id="91FCEAA2FFFAF25FFA0A4491FEAEFD2D" italics="true" pageId="5" pageNumber="143">L. triradiatus</emphasis>
</taxonomicName>
. The largest specimen examined by them was
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SL (ZSI 878/1). The specimen reported herein (NMMB-P36204;
<figureCitation id="3BB32A35FFFAF25FFEE744EAFE10FD08" box="[363,433,740,766]" captionStart="FIGURE 3" captionStartId="5.[151,250,1854,1878]" captionTargetBox="[154,1434,999,1829]" captionTargetId="figure-380@5.[154,1434,999,1829]" captionTargetPageId="5" captionText="FIGURE 3. Lophiodes triradiatus (Lloyd, 1909), NMMB-P36204, 404 mm SL, from South China Sea." figureDoi="http://doi.org/10.5281/zenodo.7119357" httpUri="https://zenodo.org/record/7119357/files/figure.png" pageId="5" pageNumber="143">Fig. 3</figureCitation>
) was also collected from near Dongsha Island (Pratas Island), South
<collectingCountry id="DB9F7620FFFAF25FFB6944EAFA89FD08" box="[1253,1320,740,766]" name="China" pageId="5" pageNumber="143">China</collectingCountry>
Sea. It is measured
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SL and is a mature female with large ovaries containing numerous small immature eggs. As a result, this specimen represents the largest known individual of
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to date.
</paragraph>
<paragraph id="A33736B0FFFAF25FFF4B4542FD48FC39" blockId="5.[151,1437,635,976]" pageId="5" pageNumber="143">
The measurements and proportions are showed in
<tableCitation id="EE0A030BFFFAF25FFC8A4542FCFAFC90" box="[774,859,844,870]" captionStart="TABLE 1" captionStartId="4.[152,249,258,284]" captionTargetBox="[163,1325,302,1536]" captionTargetPageId="4" captionText="TABLE 1. Morphometrics data of Lophiodes lugubris and L. triradiatus examined in present study." httpUri="http://table.plazi.org/id/F7F76638FFFBF25EFF14470CFB5BFEEB" pageId="5" pageNumber="143" tableUuid="F7F76638FFFBF25EFF14470CFB5BFEEB">Table 1</tableCitation>
. The morphology of present specimen agrees with previous description (
<bibRefCitation id="C7194B41FFFAF25FFE044561FD91FC7F" author="Ho, H. - C. &amp; Seret, B. &amp; Shao, K. - T." box="[392,560,879,905]" pageId="5" pageNumber="143" pagination="62 - 68" refId="ref4433" refString="Ho, H. - C., Seret, B. &amp; Shao, K. - T. (2009) Redescription of Lophiodes infrabrunneus Smith and Radcliffe, 1912, a senior synonym of L. abdituspinus Ni, Wu and Li, 1990 (Lophiiformes: Lophiidae). Zootaxa, 2326 (1), 62 - 68. https: // doi. org / 10.11646 / zootaxa. 2326.1.5" type="journal article" year="2009">
Ho
<emphasis id="91FCEAA2FFFAF25FFE3D4561FE49FC7F" box="[433,488,879,905]" italics="true" pageId="5" pageNumber="143">et al.</emphasis>
, 2009
</bibRefCitation>
,
<bibRefCitation id="C7194B41FFFAF25FFDB64561FDD7FC7F" author="Ho, H. - C. &amp; Bineesh, K. K. &amp; Akhilesh, K. V." box="[570,630,879,905]" pageId="5" pageNumber="143" pagination="587 - 592" refId="ref4360" refString="Ho, H. - C., Bineesh, K. K. &amp; Akhilesh, K. V. (2014) Rediscovery of Lophiodes triradiatus (Lloyd, 1909), a senior synonym of L. infrabrunneus Smith and Radcliffe (Lophiiformes: Lophiidae). Zootaxa, 3786 (5), 587 - 592. https: // doi. org / 10.11646 / zootaxa. 3786.5.6" type="journal article" year="2014">2014</bibRefCitation>
), except for most spines on head are relatively blunt or reduced. The esca is damaged and the leaf-like structure is not detectable. These blunt or reduced spines on head may be attributed to the large body size, which is likely an old individual.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,707 @@
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<paragraph id="A33736B0FFFEF25BFF1B4729FD95FEB7" blockId="1.[151,564,295,357]" box="[151,564,295,322]" pageId="1" pageNumber="139">
<heading id="F87F81DCFFFEF25BFF1B4729FD95FEB7" bold="true" box="[151,564,295,322]" fontSize="11" level="1" pageId="1" pageNumber="139" reason="1">
<taxonomicName id="64884D33FFFEF25BFF1B4729FD95FEB7" ID-CoL="3W3K3" authority="(Alcock 1894)" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[151,564,295,322]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="139" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFEF25BFF1B4729FD95FEB7" bold="true" box="[151,564,295,322]" pageId="1" pageNumber="139">
<emphasis id="91FCEAA2FFFEF25BFF1B4729FE20FEB4" bold="true" box="[151,385,295,322]" italics="true" pageId="1" pageNumber="139">Lophiodes lugubris</emphasis>
(Alcock 1894)
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFEF25BFF1B4745FEC0FE93" box="[151,353,331,357]" pageId="1" pageNumber="139" type="description">
<paragraph id="A33736B0FFFEF25BFF1B4745FEC0FE93" blockId="1.[151,564,295,357]" box="[151,353,331,357]" pageId="1" pageNumber="139">
<figureCitation id="3BB32A35FFFEF25BFF1B4745FEA3FE93" box="[151,258,331,357]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1967,1991]" captionStartId-1="3.[151,250,927,951]" captionTargetBox-0="[236,1351,200,1903]" captionTargetBox-1="[300,1284,191,891]" captionTargetId-0="figure-17@2.[227,1359,179,1941]" captionTargetId-1="figure-24@3.[275,1312,181,903]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Lophiodes lugubris (Alcock, 1894). A. Original drawing, reproduced from Alcock (1895). BC. NMMB-P36203, 214 mm SL, fresh caught specimen, from off Dongsha Island. B. After frozen, dorsal view. C. After preservation, ventral view." captionText-1="FIGURE 2. Proportions of lengths of illicium (IL), second dorsal-fin spine (DS2), and third dorsal-fin spine (DS3), versus SL, in Lophiodes lugubris, showing the growth changes." figureDoi-0="http://doi.org/10.5281/zenodo.7119353" figureDoi-1="http://doi.org/10.5281/zenodo.7119355" httpUri-0="https://zenodo.org/record/7119353/files/figure.png" httpUri-1="https://zenodo.org/record/7119355/files/figure.png" pageId="1" pageNumber="139">Figs. 12</figureCitation>
;
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</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFEF25BFF1B479EFF76FC91" pageId="1" pageNumber="139" type="reference_group">
<paragraph id="A33736B0FFFEF25BFF1B479EFAAEFE13" blockId="1.[151,1437,399,546]" pageId="1" pageNumber="139">
<taxonomicName id="64884D33FFFEF25BFF1B479EFE43FE5E" ID-CoL="3W2MD" authority="Alcock, 1894 a" authorityName="Alcock" authorityYear="1894" box="[151,482,400,424]" class="Actinopterygii" family="Lophiidae" genus="Lophius" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="139" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFEF25BFF1B479EFEE4FE5E" box="[151,325,400,424]" italics="true" pageId="1" pageNumber="139">Lophius lugubris</emphasis>
<bibRefCitation id="C7194B41FFFEF25BFEC0479EFE43FE5E" author="Alcock, A. W." box="[332,482,400,424]" pageId="1" pageNumber="139" pagination="115 - 137" refId="ref3911" refString="Alcock, A. W. (1894 a) Natural history notes from H. M. Indian marine survey steamer`Investigator, ', Commander C. F. Oldham, R. N., commanding. Series II., No. 11. An account of a recent collection of bathybial fishes from the Bay of Bengal and from the Laccadive Sea. Journal of the Asiatic Society of Bengal, 63 (pt. 2), 115 - 137., pls. 6 - 7." type="journal article" year="1894">Alcock, 1894a</bibRefCitation>
</taxonomicName>
(Type locality:
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north
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, Investigator station 151, 142400 fathoms [
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<typeStatus id="7C338812FFFEF25BFD1D47A1FD4FFE31" box="[657,750,431,455]" pageId="1" pageNumber="139" type="syntype">syntypes</typeStatus>
: ZSI F13467, 670/1, 671/1).
<bibRefCitation id="C7194B41FFFEF25BFB9947A0FB03FE30" author="Alcock, A. W." box="[1045,1186,430,454]" pageId="1" pageNumber="139" refId="ref4101" refString="Alcock, A. W. (1895) Illustrations of the zoology of the Royal Indian marine surveying steamer Investigator, under the command of Commander A. Carpenter, R. N., D. S. O., and of Commander R. F. Hoskyns, R. N. Fishes- - Part III. Office of the superintendent of government Printing, Calcutta, India, pls. XIV - XVI." type="book" year="1895">Alcock, 1895</bibRefCitation>
: pl. 14, fig. 1 (drawing).
<bibRefCitation id="C7194B41FFFEF25BFF4B47C3FED1FE13" author="Alcock, A. W." box="[199,368,461,485]" pageId="1" pageNumber="139" refId="ref4180" refString="Alcock, A. W. (1899) A descriptive catalogue of the Indian deep-sea fishes in the Indian Museum: being a revised account of the deep-sea fishes collected by the Royal Indian marine survey ship Investigator. Trustees of the Indian Museum, Calcutta, 211 pp. https: // doi. org / 10.5962 / bhl. title. 4684" type="book" year="1899">Alcock, 1899:55</bibRefCitation>
(based on type series).
<bibRefCitation id="C7194B41FFFEF25BFDD347C3FD54FE13" author="Lloyd, R. E." box="[607,757,461,485]" pageId="1" pageNumber="139" pagination="1 - 12" refId="ref4780" refString="Lloyd, R. E. (1907) Contributions to the fauna of the Arabian Sea, with descriptions of new fishes and crustacea. Records of the Indian Museum, 1, 1 - 12." type="journal article" year="1907">Lloyd, 1907: 5</bibRefCitation>
(listed,
<specimenCount id="B58EFD39FFFEF25BFCCB47C3FC69FE13" box="[839,968,461,485]" pageId="1" pageNumber="139" type="generic">3 specimens</specimenCount>
).
<bibRefCitation id="C7194B41FFFEF25BFC5547C3FB27FE13" author="Weber, M." box="[985,1158,461,485]" pageId="1" pageNumber="139" refId="ref5014" refString="Weber, M. (1913) Die Fische der Siboga-Expedition. Brill, Leiden, 710 pp." type="book" year="1913">Weber, 1913:557</bibRefCitation>
(mentioned).
</paragraph>
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<emphasis id="91FCEAA2FFFEF25BFF1B47E2FED7FDF2" box="[151,374,492,516]" italics="true" pageId="1" pageNumber="139">Chirolophius lugubris</emphasis>
(Alcock)
</taxonomicName>
:
<treatmentCitation id="222910A1FFFEF25BFE6947E2FD33FDF2" author="Regan, C. T." box="[485,658,492,516]" page="281" pageId="1" pageNumber="139" year="1903">
<bibRefCitation id="C7194B41FFFEF25BFE6947E2FD33FDF2" author="Regan, C. T." box="[485,658,492,516]" pageId="1" pageNumber="139" pagination="277 - 285" refId="ref4912" refString="Regan, C. T. (1903) A revision of the fishes of the family Lophiidae. The Annals and magazine of natural history; zoology, botany, and geology, 11 (7), 277 - 285. https: // doi. org / 10.1080 / 00222930308678765" type="journal article" year="1903">Regan, 1903:281</bibRefCitation>
</treatmentCitation>
(review).
<treatmentCitation id="222910A1FFFEF25BFD7647E2FC62FDF5" author="Le Danois, Y." box="[762,963,491,516]" page="88" pageId="1" pageNumber="139" year="1975">
<bibRefCitation id="C7194B41FFFEF25BFD7647E2FC62FDF5" author="Le Danois, Y." box="[762,963,491,516]" pageId="1" pageNumber="139" pagination="1 - 127" refId="ref4703" refString="Le Danois, Y. (1975) Etude osteo-myologique et revision systematique de la famille des Lophiidae (pediculates haplopterygiens). Memoires du Museum National d'Histoire Naturelle, Serie A Zoologie, 91, 1 - 127." type="journal article" year="1975">Le Danois, 1975:88</bibRefCitation>
</treatmentCitation>
(review).
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="8398119EFFFEF25BFF1B4404FCC5FDD4" box="[151,868,522,546]" pageId="1" pageNumber="139">
<taxonomicName id="64884D33FFFEF25BFF1B4404FE61FDD4" ID-CoL="3W3K3" authority="(Alcock)" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[151,448,522,546]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="139" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFEF25BFF1B4404FEFAFDD4" box="[151,347,522,546]" italics="true" pageId="1" pageNumber="139">Lophiodes lugubris</emphasis>
(Alcock)
</taxonomicName>
:
<treatmentCitation id="222910A1FFFEF25BFE454405FD20FDD4" author="Ho, H. - C. &amp; Kawai, T. &amp; Wudianto &amp; Satria, F." box="[457,641,522,546]" page="83" pageId="1" pageNumber="139" year="2016">
<bibRefCitation id="C7194B41FFFEF25BFE454405FD20FDD4" author="Ho, H. - C. &amp; Kawai, T. &amp; Wudianto &amp; Satria, F." box="[457,641,522,546]" pageId="1" pageNumber="139" pagination="77 - 85" refId="ref4584" refString="Ho, H. - C., Kawai, T., Wudianto &amp; Satria, F. (2016) Records of anglerfishes (Actinopterygii: Lophiiformes: Lophiidae) from Indonesia. Acta Ichthyologica et Piscatoria, 46 (2), 77 - 85. https: // doi. org / 10.3750 / AIP 2016.46.2.03" type="journal article" year="2016">
Ho
<emphasis id="91FCEAA2FFFEF25BFE634405FDBAFDD4" box="[495,539,522,546]" italics="true" pageId="1" pageNumber="139">et al</emphasis>
., 2016:83
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(record of
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).
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</paragraph>
<paragraph id="A33736B0FFFEF25BFF1B445EFF76FC91" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">
<emphasis id="91FCEAA2FFFEF25BFF1B445EFE38FD9C" bold="true" box="[151,409,592,618]" pageId="1" pageNumber="139">Specimens examined.</emphasis>
<materialsCitation id="13E03CEDFFFEF25BFE2D445EFD1FFD79" ID-GBIF-Occurrence="3914000302" county="Columbo Light House" elevation="731" location="Investigator station" municipality="Sri Lanka" pageId="1" pageNumber="139" specimenCount="731" typeStatus="syntype">
<emphasis id="91FCEAA2FFFEF25BFE2D445EFDB5FD9C" bold="true" box="[417,532,592,618]" pageId="1" pageNumber="139">
<typeStatus id="7C338812FFFEF25BFE2D445EFDAFFD9C" box="[417,526,592,618]" pageId="1" pageNumber="139" type="syntype">Syntypes</typeStatus>
.
</emphasis>
ZSI F670/1 (
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SL), ZSI F671/1 (94.3), west of
<collectingCounty id="4A564E3CFFFEF25BFB1B445EFA39FD9D" box="[1175,1432,592,619]" pageId="1" pageNumber="139">Columbo Light House</collectingCounty>
,
<collectingMunicipality id="4353ACCAFFFEF25BFF1B447AFEABFD78" box="[151,266,628,654]" pageId="1" pageNumber="139">Sri Lanka</collectingMunicipality>
,
<location id="A657606BFFFEF25BFE94447AFE53FD78" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFE94447AFE53FD78" box="[280,498,628,654]" county="Columbo Light House" municipality="Sri Lanka" name="Investigator station" pageId="1" pageNumber="139">Investigator station</location>
151, 260
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</quantity>
.
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</materialsCitation>
<materialsCitation id="13E03CEDFFFEF25BFD4B447BFC7DFD45" ID-GBIF-Occurrence="3914000304" collectingDate="1911-04-27" collectorName="Non-types." country="India" elevation="433" latitude="9.236112" location="southwestern Indian" longLatPrecision="913" longitude="75.75" municipality="Investigator station" pageId="1" pageNumber="139" specimenCount="1" typeStatus="syntype">
<emphasis id="91FCEAA2FFFEF25BFD4B447BFC30FD78" bold="true" box="[711,913,628,655]" pageId="1" pageNumber="139">
<collectorName id="0E7D5366FFFEF25BFD4B447BFCE6FD79" box="[711,839,629,655]" pageId="1" pageNumber="139">Non-types.</collectorName>
<collectingCountry id="DB9F7620FFFEF25BFCDD447AFC30FD78" box="[849,913,628,654]" name="India" pageId="1" pageNumber="139">India</collectingCountry>
</emphasis>
: ZSI F6846/1 (115),
<collectingMunicipality id="4353ACCAFFFEF25BFB04447AFAC3FD78" box="[1160,1378,628,654]" pageId="1" pageNumber="139">Investigator station</collectingMunicipality>
391,
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,
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,
<location id="A657606BFFFEF25BFE014496FDD3FD44" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFE014496FDD3FD44" box="[397,626,664,690]" country="India" latitude="9.236112" longLatPrecision="913" longitude="75.75" municipality="Investigator station" name="southwestern Indian" pageId="1" pageNumber="139">southwestern Indian</location>
, 237 fms [
<quantity id="64709B55FFFEF25BFD6B4497FC94FD45" box="[743,821,665,691]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="1" pageNumber="139" unit="m" value="433.0">
<elevation id="28A5D183FFFEF25BFD6B4497FC94FD45" box="[743,821,665,691]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="1" pageNumber="139" unit="m" value="433.0">433 m</elevation>
</quantity>
],
<date id="D7361070FFFEF25BFCC44497FC7DFD45" box="[840,988,665,691]" pageId="1" pageNumber="139" value="1911-04-27">
<collectingDate id="C772E998FFFEF25BFCC44497FC7DFD45" box="[840,988,665,691]" pageId="1" pageNumber="139" value="1911-04-27">27 Apr. 1911</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="13E03CEDFFFEF25BFC6B4496FBABFCE9" ID-GBIF-Occurrence="3914000303" collectingDate="2022-03" collectionCode="NMMB-P" country="India" county="Gulf of Aden" elevation="238" latitude="13.6" location="southern Indian" longLatPrecision="1290" longitude="47.533333" municipality="Investigator station" pageId="1" pageNumber="139" specimenCode="NMMB-P36203" specimenCount="1" typeStatus="syntype">
<emphasis id="91FCEAA2FFFEF25BFC6B4496FB23FD44" bold="true" box="[999,1154,664,690]" pageId="1" pageNumber="139">
<collectingCounty id="4A564E3CFFFEF25BFC6B4496FB23FD44" box="[999,1154,664,690]" pageId="1" pageNumber="139">Gulf of Aden</collectingCounty>
</emphasis>
: ZSI F1281/1 (86.8, 1 of
<quantity id="64709B55FFFEF25BFF1B44B3FF63FD20" box="[151,194,700,727]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="139" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="C7194B41FFFEF25BFF4644B2FEF1FD21" author="Lloyd, R. E." box="[202,336,700,727]" pageId="1" pageNumber="139" pagination="1 - 12" refId="ref4780" refString="Lloyd, R. E. (1907) Contributions to the fauna of the Arabian Sea, with descriptions of new fishes and crustacea. Records of the Indian Museum, 1, 1 - 12." type="journal article" year="1907">Lloyd 1907</bibRefCitation>
),
<collectingMunicipality id="4353ACCAFFFEF25BFEEF44B2FD9AFD20" box="[355,571,700,726]" pageId="1" pageNumber="139">Investigator station</collectingMunicipality>
360,
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,
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, 130 fms [
<quantity id="64709B55FFFEF25BFC3344B3FBACFD21" box="[959,1037,701,727]" metricMagnitude="2" metricUnit="m" metricValue="2.38" pageId="1" pageNumber="139" unit="m" value="238.0">
<elevation id="28A5D183FFFEF25BFC3344B3FBACFD21" box="[959,1037,701,727]" metricMagnitude="2" metricUnit="m" metricValue="2.38" pageId="1" pageNumber="139" unit="m" value="238.0">238 m</elevation>
</quantity>
], 18971898; plus 18 uncatalogued specimens (78.8198), collected from off
<location id="A657606BFFFEF25BFDF744EEFC8CFD0C" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFDF744EEFC8CFD0C" box="[635,813,736,762]" country="India" county="Gulf of Aden" latitude="13.6" longLatPrecision="1290" longitude="47.533333" municipality="Investigator station" name="southern Indian" pageId="1" pageNumber="139">southern Indian</location>
coast, precise locality not recorded.
<emphasis id="91FCEAA2FFFEF25BFB5444EFFA34FD0D" bold="true" box="[1240,1429,737,763]" pageId="1" pageNumber="139">
<location id="A657606BFFFEF25BFB5444EFFA34FD0D" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFB5444EFFA34FD0D" box="[1240,1429,737,763]" country="India" county="Gulf of Aden" latitude="13.6" longLatPrecision="1290" longitude="47.533333" municipality="Investigator station" name="Dongsha Island" pageId="1" pageNumber="139">Dongsha Island</location>
</emphasis>
:
<specimenCode id="F32E9ECBFFFEF25BFF1B450AFEEEFCE8" box="[151,335,772,798]" collectionCode="NMMB-P" pageId="1" pageNumber="139">NMMB-P36203</specimenCode>
(214),
<location id="A657606BFFFEF25BFE2F450AFDC2FCE9" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFE2F450AFDC2FCE9" box="[419,611,772,799]" country="India" county="Gulf of Aden" latitude="13.6" longLatPrecision="1290" longitude="47.533333" municipality="Investigator station" name="South China Sea" pageId="1" pageNumber="139">South China Sea</location>
, bottom trawl, ca.
<quantity id="64709B55FFFEF25BFCB6450AFC25FCE9" box="[826,900,772,799]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="139" unit="m" value="500.0">
<elevation id="28A5D183FFFEF25BFCB6450AFC25FCE9" box="[826,900,772,799]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="139" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="D7361070FFFEF25BFC1D450BFBABFCE9" box="[913,1034,773,799]" pageId="1" pageNumber="139" value="2022-03">
<collectingDate id="C772E998FFFEF25BFC1D450BFBABFCE9" box="[913,1034,773,799]" pageId="1" pageNumber="139" value="2022-03">Mar. 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="13E03CEDFFFEF25BFB99450AFF72FC91" ID-GBIF-Occurrence="3914000301" collectingDate="2004-09-28" collectionCode="HUMZ" country="Indonesia" elevation="425" latitude="-3.3906667" location="Eastern Indian Ocean" longLatPrecision="13" longitude="100.326" pageId="1" pageNumber="139" specimenCode="HUMZ 190869" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="syntype">
<emphasis id="91FCEAA2FFFEF25BFB99450AFB2BFCE8" bold="true" box="[1045,1162,772,798]" pageId="1" pageNumber="139">
<collectingCountry id="DB9F7620FFFEF25BFB99450AFB2BFCE8" box="[1045,1162,772,798]" name="Indonesia" pageId="1" pageNumber="139">Indonesia</collectingCountry>
</emphasis>
:
<specimenCode id="F32E9ECBFFFEF25BFB16450BFAE8FCE8" box="[1178,1353,772,799]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="1" pageNumber="139">HUMZ 190869</specimenCode>
(41.6),
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,
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3°23.44S, 100°19.56E, off
<collectingRegion id="614CF852FFFEF25BFD714527FCFCFCB5" box="[765,861,809,835]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="139">Sumatra</collectingRegion>
,
<location id="A657606BFFFEF25BFCE64527FBC3FCB5" LSID="urn:lsid:plazi:treatment:2B2187A6FFFEF25FFF1B4729FBEEFDD8:A657606BFFFEF25BFCE64527FBC3FCB5" box="[874,1122,808,835]" country="Indonesia" latitude="-3.3906667" longLatPrecision="13" longitude="100.326" name="Eastern Indian Ocean" pageId="1" pageNumber="139" stateProvince="Sumatera Utara">Eastern Indian Ocean</location>
,
<quantity id="64709B55FFFEF25BFBE34526FB53FCB5" box="[1135,1266,808,835]" metricMagnitude="2" metricUnit="m" metricValue="4.245" metricValueMax="4.529999999999999" metricValueMin="3.96" pageId="1" pageNumber="139" unit="m" value="424.5" valueMax="453.0" valueMin="396.0">
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,
<date id="D7361070FFFEF25BFA8C4527FF72FC91" pageId="1" pageNumber="139" value="2004-09-28">
<collectingDate id="C772E998FFFEF25BFA8C4527FF72FC91" pageId="1" pageNumber="139" value="2004-09-28">28 September 2004</collectingDate>
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.
</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFEF25BFF4B457EFF52FC01" pageId="1" pageNumber="139" type="diagnosis">
<paragraph id="A33736B0FFFEF25BFF4B457EFF52FC01" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">
<emphasis id="91FCEAA2FFFEF25BFF4B457EFEE3FC7C" bold="true" box="[199,322,880,906]" pageId="1" pageNumber="139">Diagnosis.</emphasis>
A species of
<taxonomicName id="64884D33FFFEF25BFE50457EFDF0FC7C" authorityName="Goode &amp; T.H.Bean" authorityYear="1896" box="[476,593,880,906]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="91FCEAA2FFFEF25BFE50457EFDF0FC7C" box="[476,593,880,906]" italics="true" pageId="1" pageNumber="139">Lophiodes</emphasis>
</taxonomicName>
that can be separated from congeners in having only four dorsal-fin spine (fourth and sixth spines absent) and combination of following characters: esca a tiny bulb; third dorsal-fin spine thin and long, 42.173.5% SL (mean 56.4% SL), extending to origin to middle of soft dorsal-fin base; 1516 pectoralfin rays.
</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFEF259FF4B420EFD56FB46" lastPageId="3" lastPageNumber="141" pageId="1" pageNumber="139" type="description">
<paragraph id="A33736B0FFFEF25BFF4B420EFE96FB70" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">
<emphasis id="91FCEAA2FFFEF25BFF4B420EFEF8FBEC" bold="true" box="[199,345,1024,1050]" pageId="1" pageNumber="139">Description.</emphasis>
Morphometric data are provided in
<tableCitation id="EE0A030BFFFEF25BFD7B420EFCECFBED" box="[759,845,1024,1051]" captionStart="TABLE 1" captionStartId="4.[152,249,258,284]" captionTargetBox="[163,1325,302,1536]" captionTargetPageId="4" captionText="TABLE 1. Morphometrics data of Lophiodes lugubris and L. triradiatus examined in present study." httpUri="http://table.plazi.org/id/F7F76638FFFBF25EFF14470CFB5BFEEB" pageId="1" pageNumber="139" tableUuid="F7F76638FFFBF25EFF14470CFB5BFEEB">Table 1</tableCitation>
. Dorsal-fin spines 4, 3 cephalic and 1 post-cephalic (fourth and sixth spines absent); dorsal-fin rays 8; anal-fin rays 6; pectoral-fin rays 1516. Head relatively short (HL 34.638.5% SL) and narrow (head width 52.961.1% HL); interorbital space flat; eyes large and rounded, directed dorsolaterally.
</paragraph>
<paragraph id="A33736B0FFFEF25BFF4B429EFD9DFAA9" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">Anterior half of premaxilla with two irregular rows of enlarged teeth, loosely arranged, those on the innermost row largest, followed by a single row of small teeth on posterior half; lower jaw with three rows of teeth anteriorly, those on the innermost row largest, gradually becoming 2 rows of smaller teeth; vomer with 13 fang teeth on each end, the outer teeth largest; palatine with a single row of few fang teeth; fifth ceratobranchial with two rows of small teeth, forming a V-shaped patch; those on second and third pharyngobranchials forming small and rounded patches. No gill rakers. Pseudobranch present.</paragraph>
<paragraph id="A33736B0FFFEF25BFF4B4366FD94F9AC" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">Illicium relatively short to moderately long, 19.231.6% SL, slightly extending beyond posteriormost frontal spine to slightly beyond epiotic spine (when retracted); esca a small simple bulb, without cirri at tip; second dorsalfin spine short to moderately long 18.234.4% SL, generally longer than illicium, extending to the base of thirddorsal fin spine to postemporal spine; third dorsal-fin spine moderately long, 34.673.5% SL, rather thin, bearing many tendrils along the entire length, extending to origin (larger specimens) to middle (smaller specimens) of soft dorsal-fin base; fourth dorsal-fin spine absent; fifth dorsal-fin spine short to moderately long (11.824.6% SL), extending to origin of soft dorsal fin.</paragraph>
<paragraph id="A33736B0FFFEF25BFF4B406AFBCBF88C" blockId="1.[151,1437,592,1915]" pageId="1" pageNumber="139">
All spines associated with skull short, sharp (smaller specimens), blunt or weakly developed (larger specimens), mostly beneath skin but feeble though skin. Palatine spines pointed but short; frontal ridge elevated posteriorly, serrated with 3 spines, the last spine largest, or a smooth ridge without spines in
<quantity id="64709B55FFFEF25BFBBC40A3FB2FF931" box="[1072,1166,1709,1735]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="1" pageNumber="139" unit="mm" value="214.0">214 mm</quantity>
specimen; inner frontal spine absent; inner sphenotic spines pointed and short or reduced with a feeble knob; outer sphenotic spines blunt, broad and compressed; epiotic spines blunt, beneath skin completely; quadrate and anterior articular spines blunt to strongly reduced, both knob-like; hyomandibular bears two low and blunt spines; opercular spines blunt, forming small knobs; subopercular and interopercular spines sharp or reduced and knob-like; cleithral spines strong and blunt; humeral spines slightly developed, simple with a small spine or knob on its base.
</paragraph>
<caption id="F7F76638FFFDF258FF1B41A1FA3DF81D" ID-DOI="http://doi.org/10.5281/zenodo.7119353" ID-Zenodo-Dep="7119353" httpUri="https://zenodo.org/record/7119353/files/figure.png" pageId="2" pageNumber="140" startId="2.[151,250,1967,1991]" targetBox="[236,1351,200,1903]" targetPageId="2">
<paragraph id="A33736B0FFFDF258FF1B41A1FA3DF81D" blockId="2.[151,1436,1967,2027]" pageId="2" pageNumber="140">
<emphasis id="91FCEAA2FFFDF258FF1B41A1FEB2F831" bold="true" box="[151,275,1967,1991]" pageId="2" pageNumber="140">FIGURE 1.</emphasis>
<taxonomicName id="64884D33FFFDF258FE9641A1FD22F831" authority="(Alcock, 1894)" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[282,643,1967,1991]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="140" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFDF258FE9641A1FE7FF831" box="[282,478,1967,1991]" italics="true" pageId="2" pageNumber="140">Lophiodes lugubris</emphasis>
(Alcock, 1894)
</taxonomicName>
. A. Original drawing, reproduced from
<bibRefCitation id="C7194B41FFFDF258FB9841A1FB0AF831" author="Alcock, A. W." box="[1044,1195,1967,1991]" pageId="2" pageNumber="140" refId="ref4101" refString="Alcock, A. W. (1895) Illustrations of the zoology of the Royal Indian marine surveying steamer Investigator, under the command of Commander A. Carpenter, R. N., D. S. O., and of Commander R. F. Hoskyns, R. N. Fishes- - Part III. Office of the superintendent of government Printing, Calcutta, India, pls. XIV - XVI." type="book" year="1895">Alcock (1895)</bibRefCitation>
. BC. NMMB-P36203, 214 mm SL, fresh caught specimen, from off Dongsha Island. B. After frozen, dorsal view. C. After preservation, ventral view.
</paragraph>
</caption>
<caption id="F7F76638FFFCF259FF1B4591FD0AFC2D" ID-DOI="http://doi.org/10.5281/zenodo.7119355" ID-Zenodo-Dep="7119355" httpUri="https://zenodo.org/record/7119355/files/figure.png" pageId="3" pageNumber="141" startId="3.[151,250,927,951]" targetBox="[300,1284,191,891]" targetPageId="3">
<paragraph id="A33736B0FFFCF259FF1B4591FD0AFC2D" blockId="3.[151,1436,927,987]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF1B4591FEB2FC41" bold="true" box="[151,275,927,951]" pageId="3" pageNumber="141">FIGURE 2.</emphasis>
Proportions of lengths of illicium (IL), second dorsal-fin spine (DS2), and third dorsal-fin spine (DS3), versus SL, in
<taxonomicName id="64884D33FFFCF259FF3D45CDFED4FC2D" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[177,373,963,987]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFCF259FF3D45CDFED4FC2D" box="[177,373,963,987]" italics="true" pageId="3" pageNumber="141">Lophiodes lugubris</emphasis>
</taxonomicName>
, showing the growth changes.
</paragraph>
</caption>
<paragraph id="A33736B0FFFCF259FF4B4223FD56FB46" blockId="3.[151,1437,1069,2038]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF4B4223FEEEFBB1" bold="true" box="[199,335,1069,1095]" pageId="3" pageNumber="141">Coloration.</emphasis>
When fresh, dorsal surface of body pinkish gray to brown; ventral surface pale with irregular light gray marbles and patches; pectoral fin with white margin. When preserved, dorsal surface grey to brown with many large light brown patches on dorsal surface, pale to light grey on ventral surface; color of illicium and esca as on dorsal surface; all fins pale distally. Peritoneum black.
</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFCF259FF6D42B6FB97F87F" pageId="3" pageNumber="141" type="distribution">
<paragraph id="A33736B0FFFCF259FF6D42B6FD06FAEE" blockId="3.[151,1437,1069,2038]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF6D42B6FEDBFB24" bold="true" box="[225,378,1208,1234]" pageId="3" pageNumber="141">Distribution.</emphasis>
Specimens examined were collected from off
<collectingCountry id="DB9F7620FFFCF259FC0242B6FC6BFB24" box="[910,970,1208,1234]" name="India" pageId="3" pageNumber="141">India</collectingCountry>
,
<collectingCountry id="DB9F7620FFFCF259FC5B42B6FBE7FB24" box="[983,1094,1208,1234]" name="Sri Lanka" pageId="3" pageNumber="141">Sri Lanka</collectingCountry>
(
<typeStatus id="7C338812FFFCF259FBDA42B6FB1AFB24" box="[1110,1211,1208,1234]" pageId="3" pageNumber="141" type="holotype">holotype</typeStatus>
) and eastern Indian Ocean side off
<collectingCountry id="DB9F7620FFFCF259FEC542D5FE17FB03" box="[329,438,1243,1269]" name="Indonesia" pageId="3" pageNumber="141">Indonesia</collectingCountry>
(
<bibRefCitation id="C7194B41FFFCF259FE4442D5FDDAFB03" author="Ho, H. - C. &amp; Kawai, T. &amp; Wudianto &amp; Satria, F." box="[456,635,1243,1270]" pageId="3" pageNumber="141" pagination="77 - 85" refId="ref4584" refString="Ho, H. - C., Kawai, T., Wudianto &amp; Satria, F. (2016) Records of anglerfishes (Actinopterygii: Lophiiformes: Lophiidae) from Indonesia. Acta Ichthyologica et Piscatoria, 46 (2), 77 - 85. https: // doi. org / 10.3750 / AIP 2016.46.2.03" type="journal article" year="2016">
Ho
<emphasis id="91FCEAA2FFFCF259FE7942D2FD88FB03" box="[501,553,1243,1269]" italics="true" pageId="3" pageNumber="141">et al</emphasis>
., 2016
</bibRefCitation>
) and now off Dong-sha Island (South
<collectingCountry id="DB9F7620FFFCF259FBB742D5FBDFFB03" box="[1083,1150,1243,1269]" name="China" pageId="3" pageNumber="141">China</collectingCountry>
Sea). Bathymetric range
<quantity id="64709B55FFFCF259FF1B42F0FEB7FAEE" box="[151,278,1278,1304]" metricMagnitude="2" metricUnit="m" metricValue="4.845" metricValueMax="7.31" metricValueMin="2.38" pageId="3" pageNumber="141" unit="m" value="484.5" valueMax="731.0" valueMin="238.0">238731 m</quantity>
based on the Indian Ocean records.
</paragraph>
<paragraph id="A33736B0FFFCF259FF4B432EFDF5FACD" blockId="3.[151,1437,1069,2038]" box="[199,596,1312,1339]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF4B432EFF5FFACC" bold="true" box="[199,254,1312,1338]" pageId="3" pageNumber="141">Size.</emphasis>
Adults attain at least
<quantity id="64709B55FFFCF259FE7C432FFDF1FACD" box="[496,592,1313,1339]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="3" pageNumber="141" unit="mm" value="214.0">214 mm</quantity>
.
</paragraph>
<paragraph id="A33736B0FFFCF259FF4B434DFB5AFA1E" blockId="3.[151,1437,1069,2038]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF4B434DFE85FAAB" bold="true" box="[199,292,1347,1373]" pageId="3" pageNumber="141">Otolith.</emphasis>
The otolith (CHLOL 15743) described here is collected from NMMB-P36203 (
<figureCitation id="3BB32A35FFFCF259FB16434DFB7FFAAB" box="[1178,1246,1347,1373]" captionStart="FIGURE 4" captionStartId="6.[151,250,1831,1855]" captionTargetBox="[169,1419,597,1807]" captionTargetId="figure-198@6.[169,1419,597,1807]" captionTargetPageId="6" captionText="FIGURE 4. Otoliths of species of Lophiidae from Taiwan. Images are view of left otoliths unless otherwise indicated. a, ventral views; b, inner views; scale bar = 1 mm. 1. Lophiodes bruchius, 279 mm SL, northern Taiwan. 25. Lophiomus setigerus: 2, 187 mm SL, northern Taiwan; 3, 70.6 mm SL, Daxi; 4, 89.6 mm SL, Dong-gang; 5, 272 mm SL, northern Taiwan. 6. Lophiodes lugubris, 214 mm SL, Dongsha Island. 79. Lophiodes naresi: 7, 96.0 mm SL, Dong-gang; 8, 157 mm SL, Dongsha Island; 9, 214 mm SL, Dongsha Island. 1012. Lophius litulon: 10, no data; 11, 66.2 mm TL, Dong-gang; 12, 94.6 mm TL, Dong-gang. 13. Lophiodes miacanthus, 90.0 mm SL, Daxi. 14. Lophiodes triradiatus, 404 mm SL, Dongsha Island." figureDoi="http://doi.org/10.5281/zenodo.7119359" httpUri="https://zenodo.org/record/7119359/files/figure.png" pageId="3" pageNumber="141">Fig. 4</figureCitation>
-6a, b). The inner face of the otolith is nearly flat, and the outer face is concave with its posterior part thicker than the anterior one. The otolith is characterized by two horn-shaped protrusions on the dorsal rim with a deeply incised notch in the middle of the otolith. The ventral rim is straight. The sulcus is only observable in the crista inferior, where it swells and becomes crest-like in the middle of the otolith. A dorsal depression occupies most of the dorsal area.
</paragraph>
<paragraph id="A33736B0FFFCF259FF4B43FEFB97F87F" blockId="3.[151,1437,1069,2038]" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF4B43FEFEC3F9FC" bold="true" box="[199,354,1520,1546]" pageId="3" pageNumber="141">Comparison.</emphasis>
<taxonomicName id="64884D33FFFCF259FEE643FEFDE2F9FC" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[362,579,1520,1546]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFCF259FEE643FEFDE2F9FC" box="[362,579,1520,1546]" italics="true" pageId="3" pageNumber="141">Lophiodes lugubris</emphasis>
</taxonomicName>
is closely similar to
<taxonomicName id="64884D33FFFCF259FCBD43FFFC00F9FC" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[817,929,1520,1546]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFCF259FCBD43FFFC00F9FC" box="[817,929,1520,1546]" italics="true" pageId="3" pageNumber="141">L. mutilus</emphasis>
</taxonomicName>
,
<taxonomicName id="64884D33FFFCF259FC3D43FFFAECF9FD" authority="Ho, Seret &amp; Shao, 2011" authorityName="Ho, Seret &amp; Shao" authorityYear="2011" box="[945,1357,1520,1547]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="iwamotoi">
<emphasis id="91FCEAA2FFFCF259FC3D43FFFB96F9FC" box="[945,1079,1521,1546]" italics="true" pageId="3" pageNumber="141">L. iwamotoi</emphasis>
<bibRefCitation id="C7194B41FFFCF259FBB343FFFAECF9FD" author="Ho, H. - C. &amp; Seret, B. &amp; Shao, K. - T." box="[1087,1357,1520,1547]" pageId="3" pageNumber="141" pagination="1722 - 1745" refId="ref4509" refString="Ho, H. - C., Seret, B. &amp; Shao, K. - T. (2011) Records of anglerfishes (Lophiiformes: Lophiidae) from the western South Pacific Ocean, with descriptions of two new species. Journal of Fish Biology, 79 (7), 1722 - 1745. https: // doi. org / 10.1111 / j. 1095 - 8649.2011.03106. x" type="journal article" year="2011">Ho, Séret &amp; Shao, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="64884D33FFFCF259FA0A43FFFD84F9DB" authority="Ho, Seret &amp; Shao, 2011" authorityName="Ho, Seret &amp; Shao" authorityYear="2011" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="91FCEAA2FFFCF259FA0A43FFFEABF9DB" italics="true" pageId="3" pageNumber="141">L. maculatus</emphasis>
<bibRefCitation id="C7194B41FFFCF259FE9E401AFD84F9DB" author="Ho, H. - C. &amp; Seret, B. &amp; Shao, K. - T." box="[274,549,1555,1582]" pageId="3" pageNumber="141" pagination="1722 - 1745" refId="ref4509" refString="Ho, H. - C., Seret, B. &amp; Shao, K. - T. (2011) Records of anglerfishes (Lophiiformes: Lophiidae) from the western South Pacific Ocean, with descriptions of two new species. Journal of Fish Biology, 79 (7), 1722 - 1745. https: // doi. org / 10.1111 / j. 1095 - 8649.2011.03106. x" type="journal article" year="2011">Ho, Séret &amp; Shao, 2011</bibRefCitation>
</taxonomicName>
. Except for the only four dorsal-fin spine (vs. five dorsal-fin spines), it can be separated from
<taxonomicName id="64884D33FFFCF259FEC44039FE18F9A6" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[328,441,1590,1616]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFCF259FEC44039FE18F9A6" box="[328,441,1590,1616]" italics="true" pageId="3" pageNumber="141">L. mutilus</emphasis>
</taxonomicName>
by having relatively broad head deep (66.271.4% HL, vs. 61.168.6% HL), relatively broad distance between inner sphenotic spines (43.050.7% HL, vs. 38.845.1% HL), relatively long snout length (56.963.2% HL, vs. 51.559.1% HL), relatively broad distance between pterotic and sphenotic spines (19.323.7% HL, vs. 16.920.1% HL); from
<taxonomicName id="64884D33FFFCF259FD874091FD32F94E" authorityName="Ho, Seret &amp; Shao" authorityYear="2011" box="[523,659,1695,1720]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="iwamotoi">
<emphasis id="91FCEAA2FFFCF259FD874091FD32F94E" box="[523,659,1695,1720]" italics="true" pageId="3" pageNumber="141">L. iwamotoi</emphasis>
</taxonomicName>
by having relatively long snout length (56.963.2% HL, vs. 55.4 57.5%) and relatively broad distance between inner sphenotic spines (43.050.7% HL, vs. 40.442.5% HL); and from
<taxonomicName id="64884D33FFFCF259FF5940EAFEC4F90B" authorityName="Ho, Seret &amp; Shao" authorityYear="2011" box="[213,357,1763,1789]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="91FCEAA2FFFCF259FF5940EAFEC4F90B" box="[213,357,1763,1789]" italics="true" pageId="3" pageNumber="141">L. maculatus</emphasis>
</taxonomicName>
in having relatively short third dorsal-fin spine (34.673.5% SL, vs. 71.987.6% SL), relatively narrow head width (52.961.1% HL, vs. 61.667.5% HL) and head depth (66.271.4% HL, vs. 73.378.9% HL), relatively short snout length (56.963.2% HL, vs. 63.268.5% HL), relatively narrow snout width (17.322.9% HL, 21.728.6% HL), distance between posterior frontal spines (38.045.6% HL, vs. 47.456.0% HL), and distance between opercular and subopercular spines (42.747.0% HL, vs. 50.060.3% HL).
</paragraph>
</subSubSection>
<subSubSection id="EB92653BFFFCF25FFF4B419DFBEEFDD8" lastPageId="5" lastPageNumber="143" pageId="3" pageNumber="141" type="discussion">
<paragraph id="A33736B0FFFCF25EFF4B419DFB75FF23" blockId="3.[151,1437,1069,2038]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141">
<emphasis id="91FCEAA2FFFCF259FF4B419DFE9AF85B" bold="true" box="[199,315,1939,1965]" pageId="3" pageNumber="141">Remarks.</emphasis>
<bibRefCitation id="C7194B41FFFCF259FEC9419AFDAEF858" author="Le Danois, Y." box="[325,527,1939,1966]" pageId="3" pageNumber="141" pagination="1 - 127" refId="ref4703" refString="Le Danois, Y. (1975) Etude osteo-myologique et revision systematique de la famille des Lophiidae (pediculates haplopterygiens). Memoires du Museum National d'Histoire Naturelle, Serie A Zoologie, 91, 1 - 127." type="journal article" year="1975">Le Danois (1975)</bibRefCitation>
reviewed the family
<taxonomicName id="64884D33FFFCF259FC84419DFCDBF85B" authorityName="Rafinesque" authorityYear="1810" box="[776,890,1939,1965]" class="Actinopterygii" family="Lophiidae" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="family">Lophiidae</taxonomicName>
and recognized
<taxonomicName id="64884D33FFFCF259FBB5419DFA93F85B" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[1081,1330,1939,1965]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFCF259FBB5419DFA93F85B" box="[1081,1330,1939,1965]" italics="true" pageId="3" pageNumber="141">Chirolophius lugubris</emphasis>
</taxonomicName>
as valid. She also put two junior synonyms,
<taxonomicName id="64884D33FFFCF259FD9041B9FC3DF827" authority="Weber, 1913" authorityName="Weber" authorityYear="1913" box="[540,924,1975,2001]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="papillosus">
<emphasis id="91FCEAA2FFFCF259FD9041B9FCA8F827" box="[540,777,1975,2001]" italics="true" pageId="3" pageNumber="141">Lophiodes papillosus</emphasis>
<bibRefCitation id="C7194B41FFFCF259FC8341B9FC3DF827" author="Weber, M." box="[783,924,1975,2001]" pageId="3" pageNumber="141" refId="ref5014" refString="Weber, M. (1913) Die Fische der Siboga-Expedition. Brill, Leiden, 710 pp." type="book" year="1913">Weber, 1913</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="64884D33FFFCF259FC5C41B9FA39F827" authority="Kamohara, 1938" authorityName="Kamohara" authorityYear="1938" box="[976,1432,1975,2001]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="91FCEAA2FFFCF259FC5C41B9FB77F827" box="[976,1238,1975,2001]" italics="true" pageId="3" pageNumber="141">Chirolophius japonicus</emphasis>
<bibRefCitation id="C7194B41FFFCF259FB5041B9FA39F827" author="Kamohara, T." box="[1244,1432,1975,2001]" pageId="3" pageNumber="141" pagination="1 - 86" refId="ref4673" refString="Kamohara, T. (1938) On the offshore bottom-fishes of Prov. Tosa, Shikoku, Japan. Maruzen Kobushiki Kaisha, Tokyo, 1 - 86." type="book chapter" year="1938">Kamohara, 1938</bibRefCitation>
</taxonomicName>
, under this name. These three names were further included in the synonymy of
<taxonomicName id="64884D33FFFCF259FBBA41D5FAD7F803" authority="(Alcock)" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[1078,1398,2011,2037]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="141" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFCF259FBBA41D5FAA6F803" box="[1078,1287,2011,2037]" italics="true" pageId="3" pageNumber="141">Lophiodes mutilus</emphasis>
(Alcock)
</taxonomicName>
by
<bibRefCitation id="C7194B41FFFBF25EFF1B4699FE9DFF47" author="Caruso, J. H." box="[151,316,151,177]" pageId="4" pageNumber="142" pagination="522 - 549" refId="ref4245" refString="Caruso, J. H. (1981) The systematics and distribution of the lophiid anglerfishes: I. A revision of the genus Lophiodes with the description of two new species. Copeia, 1981 (3), 522 - 549. https: // doi. org / 10.2307 / 1444556" type="journal article" year="1981">Caruso (1981)</bibRefCitation>
. However, the original descriptions of
<taxonomicName id="64884D33FFFBF25EFD614699FC21FF47" authorityName="Weber" authorityYear="1913" box="[749,896,151,177]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="papillosus">
<emphasis id="91FCEAA2FFFBF25EFD614699FC21FF47" box="[749,896,151,177]" italics="true" pageId="4" pageNumber="142">C. papillosus</emphasis>
</taxonomicName>
and
<taxonomicName id="64884D33FFFBF25EFC3B4699FBE5FF46" authorityName="Kamohara" authorityYear="1938" box="[951,1092,151,177]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="91FCEAA2FFFBF25EFC3B4699FBE5FF46" box="[951,1092,151,177]" italics="true" pageId="4" pageNumber="142">C. japonicus</emphasis>
</taxonomicName>
showed both have five dorsalfin spines, thus we retained in the junior synonyms of
<taxonomicName id="64884D33FFFBF25EFD7B46B5FCC7FF23" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[759,870,187,213]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFBF25EFD7B46B5FCC7FF23" box="[759,870,187,213]" italics="true" pageId="4" pageNumber="142">L. mutilus</emphasis>
</taxonomicName>
, as suggested by
<bibRefCitation id="C7194B41FFFBF25EFBA746B5FB71FF23" author="Caruso, J. H." box="[1067,1232,187,213]" pageId="4" pageNumber="142" pagination="522 - 549" refId="ref4245" refString="Caruso, J. H. (1981) The systematics and distribution of the lophiid anglerfishes: I. A revision of the genus Lophiodes with the description of two new species. Copeia, 1981 (3), 522 - 549. https: // doi. org / 10.2307 / 1444556" type="journal article" year="1981">Caruso (1981)</bibRefCitation>
.
</paragraph>
<caption id="F7F76638FFFBF25EFF14470CFB5BFEEB" ID-Table-UUID="F7F76638FFFBF25EFF14470CFB5BFEEB" box="[152,1274,258,285]" httpUri="http://table.plazi.org/id/F7F76638FFFBF25EFF14470CFB5BFEEB" pageId="4" pageNumber="142" startId="4.[152,249,258,284]" targetBox="[163,1325,302,1536]" targetIsTable="true" targetPageId="4">
<paragraph id="A33736B0FFFBF25EFF14470CFB5BFEEB" blockId="4.[152,1274,258,285]" box="[152,1274,258,285]" pageId="4" pageNumber="142">
<emphasis id="91FCEAA2FFFBF25EFF14470CFEB4FEEB" bold="true" box="[152,277,258,285]" pageId="4" pageNumber="142">TABLE 1.</emphasis>
Morphometrics data of
<taxonomicName id="64884D33FFFBF25EFDA8470DFD5DFEEB" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[548,764,259,285]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFDA8470DFD5DFEEB" box="[548,764,259,285]" italics="true" pageId="4" pageNumber="142">Lophiodes lugubris</emphasis>
</taxonomicName>
and
<taxonomicName id="64884D33FFFBF25EFCBF470DFC68FEEB" baseAuthorityName="Lloyd" baseAuthorityYear="1907" box="[819,969,259,285]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="triradiatus">
<emphasis id="91FCEAA2FFFBF25EFCBF470DFC68FEEB" box="[819,969,259,285]" italics="true" pageId="4" pageNumber="142">L. triradiatus</emphasis>
</taxonomicName>
examined in present study.
</paragraph>
</caption>
<paragraph id="A33736B0FFFBF25EFE764720FB74F9F6" pageId="4" pageNumber="142">
<table id="D188C410FFFB0DA5FF2F4720FA8CF9F6" box="[163,1325,302,1536]" gridcols="4" gridrows="29" pageId="4" pageNumber="142">
<tr id="1DB834F2FFFB0DA5FF2F4720FA8CFEB0" box="[163,1325,302,326]" gridrow="0" pageId="4" pageNumber="142" rowspan-0="1" rowspan-2="1">
<th id="5E695D8EFFFB0DA5FE764720FD1FFEB0" box="[506,702,302,326]" gridcol="1" gridrow="0" pageId="4" pageNumber="142">
<taxonomicName id="64884D33FFFBF25EFE764720FD1FFEB0" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[506,702,302,326]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFE764720FD1FFEB0" box="[506,702,302,326]" italics="true" pageId="4" pageNumber="142">Lophiodes lugubris</emphasis>
</taxonomicName>
</th>
<th id="5E695D8EFFFB0DA5FBDD4720FA8CFEB0" box="[1105,1325,302,326]" gridcol="3" gridrow="0" pageId="4" pageNumber="142">
<emphasis id="91FCEAA2FFFBF25EFBDD4720FA8CFEB0" box="[1105,1325,302,326]" italics="true" pageId="4" pageNumber="142">Lophiodes triradiatus</emphasis>
</th>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4757FA8CFE87" box="[163,1325,345,369]" gridrow="1" pageId="4" pageNumber="142" rowspan-2="1">
<th id="5E695D8EFFFB0DA5FF2F4757FF5FFE87" box="[163,254,345,369]" gridcol="0" gridrow="1" pageId="4" pageNumber="142">SL (mm)</th>
<td id="5E695D8EFFFB0DA5FE764757FD1FFE87" box="[506,702,345,369]" gridcol="1" gridrow="1" pageId="4" pageNumber="142">78.8214 (n=21)</td>
<td id="5E695D8EFFFB0DA5FBDD4757FA8CFE87" box="[1105,1325,345,369]" gridcol="3" gridrow="1" pageId="4" pageNumber="142">404 (n=1)</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F478BFA8CFE6B" box="[163,1325,389,413]" gridrow="2" pageId="4" pageNumber="142" rowspan-0="1" rowspan-3="1">
<td id="5E695D8EFFFB0DA5FE76478BFD1FFE6B" box="[506,702,389,413]" gridcol="1" gridrow="2" pageId="4" pageNumber="142">Mean (Range)</td>
<td id="5E695D8EFFFB0DA5FCD1478BFC28FE6B" box="[861,905,389,413]" gridcol="2" gridrow="2" pageId="4" pageNumber="142">SD</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F47BEFA8CFE3E" box="[163,1325,432,456]" gridrow="3" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F47BEFA8CFE3E" box="[163,1325,432,456]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="4" pageNumber="142">% SL</th>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F47D5FA8CFE05" box="[163,1325,475,499]" gridrow="4" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F47D5FF5FFE05" box="[163,254,475,499]" gridcol="0" gridrow="4" pageId="4" pageNumber="142">HL</th>
<td id="5E695D8EFFFB0DA5FE7647D5FD1FFE05" box="[506,702,475,499]" gridcol="1" gridrow="4" pageId="4" pageNumber="142">36.4 (34.638.5)</td>
<td id="5E695D8EFFFB0DA5FCD147D5FC28FE05" box="[861,905,475,499]" gridcol="2" gridrow="4" pageId="4" pageNumber="142">1.2</td>
<td id="5E695D8EFFFB0DA5FBDD47D5FA8CFE05" box="[1105,1325,475,499]" gridcol="3" gridrow="4" pageId="4" pageNumber="142">33.4</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4408FA8CFDE8" box="[163,1325,518,542]" gridrow="5" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4408FF5FFDE8" box="[163,254,518,542]" gridcol="0" gridrow="5" pageId="4" pageNumber="142">HW</th>
<td id="5E695D8EFFFB0DA5FE764408FD1FFDE8" box="[506,702,518,542]" gridcol="1" gridrow="5" pageId="4" pageNumber="142">20.7 (19.223.5)</td>
<td id="5E695D8EFFFB0DA5FCD14408FC28FDE8" box="[861,905,518,542]" gridcol="2" gridrow="5" pageId="4" pageNumber="142">1.4</td>
<td id="5E695D8EFFFB0DA5FBDD4408FA8CFDE8" box="[1105,1325,518,542]" gridcol="3" gridrow="5" pageId="4" pageNumber="142">18.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F443FFA8CFDBF" box="[163,1325,561,585]" gridrow="6" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F443FFF5FFDBF" box="[163,254,561,585]" gridcol="0" gridrow="6" pageId="4" pageNumber="142">HD</th>
<td id="5E695D8EFFFB0DA5FE76443FFD1FFDBF" box="[506,702,561,585]" gridcol="1" gridrow="6" pageId="4" pageNumber="142">25.4 (23.927.3)</td>
<td id="5E695D8EFFFB0DA5FCD1443FFC28FDBF" box="[861,905,561,585]" gridcol="2" gridrow="6" pageId="4" pageNumber="142">1.2</td>
<td id="5E695D8EFFFB0DA5FBDD443FFA8CFDBF" box="[1105,1325,561,585]" gridcol="3" gridrow="6" pageId="4" pageNumber="142">24.8</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4452FA8CFD83" box="[163,1325,604,629]" gridrow="7" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4452FF5FFD83" box="[163,254,604,629]" gridcol="0" gridrow="7" pageId="4" pageNumber="142">IL</th>
<td id="5E695D8EFFFB0DA5FE764452FD1FFD83" box="[506,702,604,629]" gridcol="1" gridrow="7" pageId="4" pageNumber="142">28.1 (19.231.6)</td>
<td id="5E695D8EFFFB0DA5FCD14452FC28FD83" box="[861,905,604,629]" gridcol="2" gridrow="7" pageId="4" pageNumber="142">3.4</td>
<td id="5E695D8EFFFB0DA5FBDD4452FA8CFD83" box="[1105,1325,604,629]" gridcol="3" gridrow="7" pageId="4" pageNumber="142">-</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4486FA8CFD56" box="[163,1325,648,672]" gridrow="8" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4486FF5FFD56" box="[163,254,648,672]" gridcol="0" gridrow="8" pageId="4" pageNumber="142">DS2</th>
<td id="5E695D8EFFFB0DA5FE764486FD1FFD56" box="[506,702,648,672]" gridcol="1" gridrow="8" pageId="4" pageNumber="142">27.4 (18.234.4)</td>
<td id="5E695D8EFFFB0DA5FCD14486FC28FD56" box="[861,905,648,672]" gridcol="2" gridrow="8" pageId="4" pageNumber="142">4.1</td>
<td id="5E695D8EFFFB0DA5FBDD4486FA8CFD56" box="[1105,1325,648,672]" gridcol="3" gridrow="8" pageId="4" pageNumber="142">18.6</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F44BDFA8CFD3D" box="[163,1325,691,715]" gridrow="9" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F44BDFF5FFD3D" box="[163,254,691,715]" gridcol="0" gridrow="9" pageId="4" pageNumber="142">DS3</th>
<td id="5E695D8EFFFB0DA5FE7644BDFD1FFD3D" box="[506,702,691,715]" gridcol="1" gridrow="9" pageId="4" pageNumber="142">55.3 (34.673.5)</td>
<td id="5E695D8EFFFB0DA5FCD144BDFC28FD3D" box="[861,905,691,715]" gridcol="2" gridrow="9" pageId="4" pageNumber="142">10.2</td>
<td id="5E695D8EFFFB0DA5FBDD44BDFA8CFD3D" box="[1105,1325,691,715]" gridcol="3" gridrow="9" pageId="4" pageNumber="142">16.6</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F44D0FA8CFD00" box="[163,1325,734,758]" gridrow="10" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F44D0FF5FFD00" box="[163,254,734,758]" gridcol="0" gridrow="10" pageId="4" pageNumber="142">DS5</th>
<td id="5E695D8EFFFB0DA5FE7644D0FD1FFD00" box="[506,702,734,758]" gridcol="1" gridrow="10" pageId="4" pageNumber="142">16.4 (11.824.6)</td>
<td id="5E695D8EFFFB0DA5FCD144D0FC28FD00" box="[861,905,734,758]" gridcol="2" gridrow="10" pageId="4" pageNumber="142">3.3</td>
<td id="5E695D8EFFFB0DA5FBDD44D0FA8CFD00" box="[1105,1325,734,758]" gridcol="3" gridrow="10" pageId="4" pageNumber="142">N/A</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4507FA8CFCD7" box="[163,1325,777,801]" gridrow="11" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4507FF5FFCD7" box="[163,254,777,801]" gridcol="0" gridrow="11" pageId="4" pageNumber="142">IF</th>
<td id="5E695D8EFFFB0DA5FE764507FD1FFCD7" box="[506,702,777,801]" gridcol="1" gridrow="11" pageId="4" pageNumber="142">15.4 (13.817.6)</td>
<td id="5E695D8EFFFB0DA5FCD14507FC28FCD7" box="[861,905,777,801]" gridcol="2" gridrow="11" pageId="4" pageNumber="142">1.3</td>
<td id="5E695D8EFFFB0DA5FBDD4507FA8CFCD7" box="[1105,1325,777,801]" gridcol="3" gridrow="11" pageId="4" pageNumber="142">9.7</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F453AFA8CFCBB" box="[163,1325,820,845]" gridrow="12" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F453AFF5FFCBB" box="[163,254,820,845]" gridcol="0" gridrow="12" pageId="4" pageNumber="142">ISP</th>
<td id="5E695D8EFFFB0DA5FE76453AFD1FFCBB" box="[506,702,820,845]" gridcol="1" gridrow="12" pageId="4" pageNumber="142">16.9 (15.219.3)</td>
<td id="5E695D8EFFFB0DA5FCD1453AFC28FCBB" box="[861,905,820,845]" gridcol="2" gridrow="12" pageId="4" pageNumber="142">1.5</td>
<td id="5E695D8EFFFB0DA5FBDD453AFA8CFCBB" box="[1105,1325,820,845]" gridcol="3" gridrow="12" pageId="4" pageNumber="142">14.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F456EFA8CFC8E" box="[163,1325,864,888]" gridrow="13" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F456EFF5FFC8E" box="[163,254,864,888]" gridcol="0" gridrow="13" pageId="4" pageNumber="142">SNL</th>
<td id="5E695D8EFFFB0DA5FE76456EFD1FFC8E" box="[506,702,864,888]" gridcol="1" gridrow="13" pageId="4" pageNumber="142">21.7 (19.723.8)</td>
<td id="5E695D8EFFFB0DA5FCD1456EFC28FC8E" box="[861,905,864,888]" gridcol="2" gridrow="13" pageId="4" pageNumber="142">1.4</td>
<td id="5E695D8EFFFB0DA5FBDD456EFA8CFC8E" box="[1105,1325,864,888]" gridcol="3" gridrow="13" pageId="4" pageNumber="142">18.6</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4585FA8CFC55" box="[163,1325,907,931]" gridrow="14" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4585FF5FFC55" box="[163,254,907,931]" gridcol="0" gridrow="14" pageId="4" pageNumber="142">SNW1</th>
<td id="5E695D8EFFFB0DA5FE764585FD1FFC55" box="[506,702,907,931]" gridcol="1" gridrow="14" pageId="4" pageNumber="142">7.3 (6.18.1)</td>
<td id="5E695D8EFFFB0DA5FCD14585FC28FC55" box="[861,905,907,931]" gridcol="2" gridrow="14" pageId="4" pageNumber="142">0.6</td>
<td id="5E695D8EFFFB0DA5FBDD4585FA8CFC55" box="[1105,1325,907,931]" gridcol="3" gridrow="14" pageId="4" pageNumber="142">8.7</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F45B8FA8CFC38" box="[163,1325,950,974]" gridrow="15" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F45B8FF5FFC38" box="[163,254,950,974]" gridcol="0" gridrow="15" pageId="4" pageNumber="142">PTSP</th>
<td id="5E695D8EFFFB0DA5FE7645B8FD1FFC38" box="[506,702,950,974]" gridcol="1" gridrow="15" pageId="4" pageNumber="142">7.9 (6.89.1)</td>
<td id="5E695D8EFFFB0DA5FCD145B8FC28FC38" box="[861,905,950,974]" gridcol="2" gridrow="15" pageId="4" pageNumber="142">0.7</td>
<td id="5E695D8EFFFB0DA5FBDD45B8FA8CFC38" box="[1105,1325,950,974]" gridcol="3" gridrow="15" pageId="4" pageNumber="142">4.7</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F45EFFA8CFC0F" box="[163,1325,993,1017]" gridrow="16" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F45EFFF5FFC0F" box="[163,254,993,1017]" gridcol="0" gridrow="16" pageId="4" pageNumber="142">QPAL</th>
<td id="5E695D8EFFFB0DA5FE7645EFFD1FFC0F" box="[506,702,993,1017]" gridcol="1" gridrow="16" pageId="4" pageNumber="142">25.4 (22.929.1)</td>
<td id="5E695D8EFFFB0DA5FCD145EFFC28FC0F" box="[861,905,993,1017]" gridcol="2" gridrow="16" pageId="4" pageNumber="142">2.2</td>
<td id="5E695D8EFFFB0DA5FBDD45EFFA8CFC0F" box="[1105,1325,993,1017]" gridcol="3" gridrow="16" pageId="4" pageNumber="142">22.8</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4202FA8CFBD3" box="[163,1325,1036,1061]" gridrow="17" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4202FF5FFBD3" box="[163,254,1036,1061]" gridcol="0" gridrow="17" pageId="4" pageNumber="142">OPSOP</th>
<td id="5E695D8EFFFB0DA5FE764202FD1FFBD3" box="[506,702,1036,1061]" gridcol="1" gridrow="17" pageId="4" pageNumber="142">16.5 (15.018.3)</td>
<td id="5E695D8EFFFB0DA5FCD14202FC28FBD3" box="[861,905,1036,1061]" gridcol="2" gridrow="17" pageId="4" pageNumber="142">1.1</td>
<td id="5E695D8EFFFB0DA5FBDD4202FA8CFBD3" box="[1105,1325,1036,1061]" gridcol="3" gridrow="17" pageId="4" pageNumber="142">18.6</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4236FA8CFBA6" box="[163,1325,1080,1104]" gridrow="18" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4236FF5FFBA6" box="[163,254,1080,1104]" gridcol="0" gridrow="18" pageId="4" pageNumber="142">TL</th>
<td id="5E695D8EFFFB0DA5FE764236FD1FFBA6" box="[506,702,1080,1104]" gridcol="1" gridrow="18" pageId="4" pageNumber="142">26.8 (23.531.2)</td>
<td id="5E695D8EFFFB0DA5FCD14236FC28FBA6" box="[861,905,1080,1104]" gridcol="2" gridrow="18" pageId="4" pageNumber="142">2.6</td>
<td id="5E695D8EFFFB0DA5FBDD4236FA8CFBA6" box="[1105,1325,1080,1104]" gridcol="3" gridrow="18" pageId="4" pageNumber="142">34.7</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F426DFA8CFB8D" box="[163,1325,1123,1147]" gridrow="19" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F426DFA8CFB8D" box="[163,1325,1123,1147]" colspan="4" colspanRight="3" gridcol="0" gridrow="19" pageId="4" pageNumber="142">%HL</th>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4280FA8CFB50" box="[163,1325,1166,1190]" gridrow="20" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4280FF5FFB50" box="[163,254,1166,1190]" gridcol="0" gridrow="20" pageId="4" pageNumber="142">HW</th>
<td id="5E695D8EFFFB0DA5FE764280FD1FFB50" box="[506,702,1166,1190]" gridcol="1" gridrow="20" pageId="4" pageNumber="142">56.3 (52.961.1)</td>
<td id="5E695D8EFFFB0DA5FCD14280FC28FB50" box="[861,905,1166,1190]" gridcol="2" gridrow="20" pageId="4" pageNumber="142">2.5</td>
<td id="5E695D8EFFFB0DA5FBDD4280FA8CFB50" box="[1105,1325,1166,1190]" gridcol="3" gridrow="20" pageId="4" pageNumber="142">54.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F42B7FA8CFB27" box="[163,1325,1209,1233]" gridrow="21" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F42B7FF5FFB27" box="[163,254,1209,1233]" gridcol="0" gridrow="21" pageId="4" pageNumber="142">HD</th>
<td id="5E695D8EFFFB0DA5FE7642B7FD1FFB27" box="[506,702,1209,1233]" gridcol="1" gridrow="21" pageId="4" pageNumber="142">69.3 (66.271.4)</td>
<td id="5E695D8EFFFB0DA5FCD142B7FC28FB27" box="[861,905,1209,1233]" gridcol="2" gridrow="21" pageId="4" pageNumber="142">1.6</td>
<td id="5E695D8EFFFB0DA5FBDD42B7FA8CFB27" box="[1105,1325,1209,1233]" gridcol="3" gridrow="21" pageId="4" pageNumber="142">74.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F42EAFA8CFB0B" box="[163,1325,1252,1277]" gridrow="22" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F42EAFF5FFB0B" box="[163,254,1252,1277]" gridcol="0" gridrow="22" pageId="4" pageNumber="142">IF</th>
<td id="5E695D8EFFFB0DA5FE7642EAFD1FFB0B" box="[506,702,1252,1277]" gridcol="1" gridrow="22" pageId="4" pageNumber="142">42.1 (38.045.6)</td>
<td id="5E695D8EFFFB0DA5FCD142EAFC28FB0B" box="[861,905,1252,1277]" gridcol="2" gridrow="22" pageId="4" pageNumber="142">2.5</td>
<td id="5E695D8EFFFB0DA5FBDD42EAFA8CFB0B" box="[1105,1325,1252,1277]" gridcol="3" gridrow="22" pageId="4" pageNumber="142">28.9</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F431EFA8CFADE" box="[163,1325,1296,1320]" gridrow="23" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F431EFF5FFADE" box="[163,254,1296,1320]" gridcol="0" gridrow="23" pageId="4" pageNumber="142">ISP</th>
<td id="5E695D8EFFFB0DA5FE76431EFD1FFADE" box="[506,702,1296,1320]" gridcol="1" gridrow="23" pageId="4" pageNumber="142">46.3 (43.050.7)</td>
<td id="5E695D8EFFFB0DA5FCD1431EFC28FADE" box="[861,905,1296,1320]" gridcol="2" gridrow="23" pageId="4" pageNumber="142">3.0</td>
<td id="5E695D8EFFFB0DA5FBDD431EFA8CFADE" box="[1105,1325,1296,1320]" gridcol="3" gridrow="23" pageId="4" pageNumber="142">42.2</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4335FA8CFAA5" box="[163,1325,1339,1363]" gridrow="24" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4335FF5FFAA5" box="[163,254,1339,1363]" gridcol="0" gridrow="24" pageId="4" pageNumber="142">SNL</th>
<td id="5E695D8EFFFB0DA5FE764335FD1FFAA5" box="[506,702,1339,1363]" gridcol="1" gridrow="24" pageId="4" pageNumber="142">59.3 (56.963.2)</td>
<td id="5E695D8EFFFB0DA5FCD14335FC28FAA5" box="[861,905,1339,1363]" gridcol="2" gridrow="24" pageId="4" pageNumber="142">2.1</td>
<td id="5E695D8EFFFB0DA5FBDD4335FA8CFAA5" box="[1105,1325,1339,1363]" gridcol="3" gridrow="24" pageId="4" pageNumber="142">55.6</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F4368FA8CFA88" box="[163,1325,1382,1406]" gridrow="25" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F4368FF5FFA88" box="[163,254,1382,1406]" gridcol="0" gridrow="25" pageId="4" pageNumber="142">SNW</th>
<td id="5E695D8EFFFB0DA5FE764368FD1FFA88" box="[506,702,1382,1406]" gridcol="1" gridrow="25" pageId="4" pageNumber="142">20.1 (17.322.9)</td>
<td id="5E695D8EFFFB0DA5FCD14368FC28FA88" box="[861,905,1382,1406]" gridcol="2" gridrow="25" pageId="4" pageNumber="142">1.9</td>
<td id="5E695D8EFFFB0DA5FBDD4368FA8CFA88" box="[1105,1325,1382,1406]" gridcol="3" gridrow="25" pageId="4" pageNumber="142">25.9</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F439FFA8CFA5F" box="[163,1325,1425,1449]" gridrow="26" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F439FFF5FFA5F" box="[163,254,1425,1449]" gridcol="0" gridrow="26" pageId="4" pageNumber="142">PTSP</th>
<td id="5E695D8EFFFB0DA5FE76439FFD1FFA5F" box="[506,702,1425,1449]" gridcol="1" gridrow="26" pageId="4" pageNumber="142">21.5 (19.323.7)</td>
<td id="5E695D8EFFFB0DA5FCD1439FFC28FA5F" box="[861,905,1425,1449]" gridcol="2" gridrow="26" pageId="4" pageNumber="142">1.5</td>
<td id="5E695D8EFFFB0DA5FBDD439FFA8CFA5F" box="[1105,1325,1425,1449]" gridcol="3" gridrow="26" pageId="4" pageNumber="142">14.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F43B2FA8CFA23" box="[163,1325,1468,1493]" gridrow="27" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F43B2FF5FFA23" box="[163,254,1468,1493]" gridcol="0" gridrow="27" pageId="4" pageNumber="142">QPAL</th>
<td id="5E695D8EFFFB0DA5FE7643B2FD1FFA23" box="[506,702,1468,1493]" gridcol="1" gridrow="27" pageId="4" pageNumber="142">69.6 (62.277.6)</td>
<td id="5E695D8EFFFB0DA5FCD143B2FC28FA23" box="[861,905,1468,1493]" gridcol="2" gridrow="27" pageId="4" pageNumber="142">5.2</td>
<td id="5E695D8EFFFB0DA5FBDD43B2FA8CFA23" box="[1105,1325,1468,1493]" gridcol="3" gridrow="27" pageId="4" pageNumber="142">68.1</td>
</tr>
<tr id="1DB834F2FFFB0DA5FF2F43E6FA8CF9F6" box="[163,1325,1512,1536]" gridrow="28" pageId="4" pageNumber="142">
<th id="5E695D8EFFFB0DA5FF2F43E6FF5FF9F6" box="[163,254,1512,1536]" gridcol="0" gridrow="28" pageId="4" pageNumber="142">OPSOP</th>
<td id="5E695D8EFFFB0DA5FE7643E6FD1FF9F6" box="[506,702,1512,1536]" gridcol="1" gridrow="28" pageId="4" pageNumber="142">44.7 (42.747.0)</td>
<td id="5E695D8EFFFB0DA5FCD143E6FC28F9F6" box="[861,905,1512,1536]" gridcol="2" gridrow="28" pageId="4" pageNumber="142">1.4</td>
<td id="5E695D8EFFFB0DA5FBDD43E6FA8CF9F6" box="[1105,1325,1512,1536]" gridcol="3" gridrow="28" pageId="4" pageNumber="142">55.6</td>
</tr>
</table>
</paragraph>
<paragraph id="A33736B0FFFBF25EFF4B403FFAC3F945" blockId="4.[151,1437,1585,2027]" pageId="4" pageNumber="142">
<bibRefCitation id="C7194B41FFFBF25EFF4B403FFED0F9BD" author="Ho, H. - C. &amp; Kawai, T. &amp; Wudianto &amp; Satria, F." box="[199,369,1585,1611]" pageId="4" pageNumber="142" pagination="77 - 85" refId="ref4584" refString="Ho, H. - C., Kawai, T., Wudianto &amp; Satria, F. (2016) Records of anglerfishes (Actinopterygii: Lophiiformes: Lophiidae) from Indonesia. Acta Ichthyologica et Piscatoria, 46 (2), 77 - 85. https: // doi. org / 10.3750 / AIP 2016.46.2.03" type="journal article" year="2016">
Ho
<emphasis id="91FCEAA2FFFBF25EFF62403CFEBDF9BD" box="[238,284,1585,1611]" italics="true" pageId="4" pageNumber="142">et al</emphasis>
. (2016)
</bibRefCitation>
examined
<specimenCount id="B58EFD39FFFBF25EFE6A403FFDD5F9BD" box="[486,628,1585,1611]" pageId="4" pageNumber="142" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
and many additional individuals of
<taxonomicName id="64884D33FFFBF25EFC79403CFBCAF9BD" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[1013,1131,1585,1611]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFC79403CFBCAF9BD" box="[1013,1131,1585,1611]" italics="true" pageId="4" pageNumber="142">L. lugubris</emphasis>
</taxonomicName>
collected from off
<collectingCountry id="DB9F7620FFFBF25EFABB403FFACEF9BD" box="[1335,1391,1585,1611]" name="India" pageId="4" pageNumber="142">India</collectingCountry>
and concluded that
<taxonomicName id="64884D33FFFBF25EFEB0405AFE12F99B" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[316,435,1619,1645]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFEB0405AFE12F99B" box="[316,435,1619,1645]" italics="true" pageId="4" pageNumber="142">L. lugubris</emphasis>
</taxonomicName>
is a valid species, with comments on the type series. In the large among specimens collected from the Indo-west Pacific Ocean examined by the first author, we found that only
<taxonomicName id="64884D33FFFBF25EFBBF4079FB00F966" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[1075,1185,1654,1680]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFBF25EFBBF4079FB00F966" box="[1075,1185,1654,1680]" italics="true" pageId="4" pageNumber="142">L. mutilus</emphasis>
</taxonomicName>
present in the western Pacific Ocean and none of them can be referred to
<taxonomicName id="64884D33FFFBF25EFD4E4094FC9BF945" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[706,826,1689,1715]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFD4E4094FC9BF945" box="[706,826,1689,1715]" italics="true" pageId="4" pageNumber="142">L. lugubris</emphasis>
</taxonomicName>
, except for the
<quantity id="64709B55FFFBF25EFC684097FB9EF945" box="[996,1087,1689,1715]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="4" pageNumber="142" unit="mm" value="214.0">214 mm</quantity>
specimen reported herein.
</paragraph>
<paragraph id="A33736B0FFFBF25EFF4B40B5FA2BF90E" blockId="4.[151,1437,1585,2027]" pageId="4" pageNumber="142">
The
<quantity id="64709B55FFFBF25EFF7040B2FEFDF920" box="[252,348,1724,1750]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="4" pageNumber="142" unit="mm" value="214.0">214 mm</quantity>
specimen is by far the largest individual of
<taxonomicName id="64884D33FFFBF25EFCD240B2FC7AF923" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[862,987,1723,1749]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="142" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFBF25EFCD240B2FC7AF923" box="[862,987,1723,1749]" italics="true" pageId="4" pageNumber="142">L. lugubris</emphasis>
</taxonomicName>
. It has a pair of moderately developed gonads with sex cannot be determinate. It is presumed that this species is matured at slightly larger than this size.
</paragraph>
<paragraph id="A33736B0FFFBF25EFF4B410FFDF6F81D" blockId="4.[151,1437,1585,2027]" pageId="4" pageNumber="142">
The
<quantity id="64709B55FFFBF25EFF70410FFEFDF8ED" box="[252,348,1793,1819]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="4" pageNumber="142" unit="mm" value="214.0">214 mm</quantity>
specimen has first threes spines relatively short. An analysis on the proportions of these three spines versus SL shows that the lengths of illicium and second dorsal-fin spine have very minor negative trend, whereas that of third dorsal-fin spine has a strong negative trend (
<figureCitation id="3BB32A35FFFBF25EFCFA4148FC1DF896" box="[886,956,1862,1888]" captionStart="FIGURE 2" captionStartId="3.[151,250,927,951]" captionTargetBox="[300,1284,191,891]" captionTargetId="figure-24@3.[275,1312,181,903]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of lengths of illicium (IL), second dorsal-fin spine (DS2), and third dorsal-fin spine (DS3), versus SL, in Lophiodes lugubris, showing the growth changes." figureDoi="http://doi.org/10.5281/zenodo.7119355" httpUri="https://zenodo.org/record/7119355/files/figure.png" pageId="4" pageNumber="142">Fig. 2</figureCitation>
). The relatively short third dorsal-fin spine is attributed to the growth change, whereas the lengths of illicium and second dorsal-fin do not change significantly. In addition, we also found that all head spines associated with skeleton are either strongly reduced, knob-like or blunt. Judging from the relatively large size, these reduced spines may also attribute to the growth change, which is common in congeners (Ho, pers. obse.).
</paragraph>
<paragraph id="A33736B0FFFAF25FFF4B4699FE11FECD" blockId="5.[151,1437,151,558]" pageId="5" pageNumber="143">
It is also notable that the
<quantity id="64709B55FFFAF25FFE694699FDE3FF47" box="[485,578,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="5" pageNumber="143" unit="mm" value="214.0">214 mm</quantity>
specimen has its ventral surface pale with irregular gray marbles or patches (
<figureCitation id="3BB32A35FFFAF25FFF1346B7FF57FF25" box="[159,246,185,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1991]" captionTargetBox="[236,1351,200,1903]" captionTargetId="figure-17@2.[227,1359,179,1941]" captionTargetPageId="2" captionText="FIGURE 1. Lophiodes lugubris (Alcock, 1894). A. Original drawing, reproduced from Alcock (1895). BC. NMMB-P36203, 214 mm SL, fresh caught specimen, from off Dongsha Island. B. After frozen, dorsal view. C. After preservation, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7119353" httpUri="https://zenodo.org/record/7119353/files/figure.png" pageId="5" pageNumber="143">Fig. 1C</figureCitation>
). Because all other specimens examined are either small individuals or had been preserved for a long time, the coloration of ventral surface is rather pale or faded. When more specimens are available, it may be proved that the coloration on ventral surface is a diagnostic character since most members we examined have a uniformly gray to brown ventral surface.
</paragraph>
<paragraph id="A33736B0FFFAF25FFF4B474AFBEEFDD8" blockId="5.[151,1437,151,558]" pageId="5" pageNumber="143">
Although
<bibRefCitation id="C7194B41FFFAF25FFEB6474AFDB8FEA8" author="Fricke, R. &amp; Mahafina, J. &amp; Behivoke, F. &amp; Jaonalison, H. &amp; Leopold, M. &amp; Ponton, D." box="[314,537,324,351]" pageId="5" pageNumber="143" pagination="1 - 432" refId="ref4300" refString="Fricke, R., Mahafina, J., Behivoke, F., Jaonalison, H., Leopold, M. &amp; Ponton, D. (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa, 3 (1), 1 - 432." type="journal article" year="2018">
Fricke
<emphasis id="91FCEAA2FFFAF25FFE07474BFE1FFEA8" box="[395,446,324,350]" italics="true" pageId="5" pageNumber="143">et al</emphasis>
. (2018)
</bibRefCitation>
recorded
<taxonomicName id="64884D33FFFAF25FFD01474AFCC9FEA8" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[653,872,324,350]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFAF25FFD01474AFCC9FEA8" box="[653,872,324,350]" italics="true" pageId="5" pageNumber="143">Lophiodes lugubris</emphasis>
</taxonomicName>
from
<collectingCountry id="DB9F7620FFFAF25FFC3E474AFB9AFEA8" box="[946,1083,324,350]" name="Madagascar" pageId="5" pageNumber="143">Madagascar</collectingCountry>
, most of specimens they listed were identified as
<taxonomicName id="64884D33FFFAF25FFED34769FD8AFE77" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[351,555,359,385]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="mutilus">
<emphasis id="91FCEAA2FFFAF25FFED34769FD8AFE77" box="[351,555,359,385]" italics="true" pageId="5" pageNumber="143">Lophiodes mutilus</emphasis>
</taxonomicName>
by the first author.
<bibRefCitation id="C7194B41FFFAF25FFD714769FC6EFE77" author="Fricke, R. &amp; Mahafina, J. &amp; Behivoke, F. &amp; Jaonalison, H. &amp; Leopold, M. &amp; Ponton, D." box="[765,975,359,385]" pageId="5" pageNumber="143" pagination="1 - 432" refId="ref4300" refString="Fricke, R., Mahafina, J., Behivoke, F., Jaonalison, H., Leopold, M. &amp; Ponton, D. (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa, 3 (1), 1 - 432." type="journal article" year="2018">
Fricke
<emphasis id="91FCEAA2FFFAF25FFCC64769FCD8FE77" box="[842,889,359,385]" italics="true" pageId="5" pageNumber="143">et al</emphasis>
. (2018)
</bibRefCitation>
probably wrongly inferred “
<taxonomicName id="64884D33FFFAF25FFA854769FD24FE55" authorityName="Le Danois" authorityYear="1975" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="variety" species="lugubris" subGenus="Lophiodes" variety="madagascariensis">
<emphasis id="91FCEAA2FFFAF25FFA854769FA3DFE77" box="[1289,1436,359,385]" italics="true" pageId="5" pageNumber="143">Chirolophius</emphasis>
(
<emphasis id="91FCEAA2FFFAF25FFF2D4787FEB7FE55" box="[161,278,393,419]" italics="true" pageId="5" pageNumber="143">Lophiodes</emphasis>
)
<emphasis id="91FCEAA2FFFAF25FFEAA4787FE22FE55" box="[294,387,393,419]" italics="true" pageId="5" pageNumber="143">lugubris</emphasis>
var.
<emphasis id="91FCEAA2FFFAF25FFE374787FD24FE55" box="[443,645,393,419]" italics="true" pageId="5" pageNumber="143">madagascariensis</emphasis>
</taxonomicName>
” described by
<bibRefCitation id="C7194B41FFFAF25FFCA34784FC57FE52" author="Le Danois, Y." box="[815,1014,393,420]" pageId="5" pageNumber="143" pagination="1 - 127" refId="ref4703" refString="Le Danois, Y. (1975) Etude osteo-myologique et revision systematique de la famille des Lophiidae (pediculates haplopterygiens). Memoires du Museum National d'Histoire Naturelle, Serie A Zoologie, 91, 1 - 127." type="journal article" year="1975">Le Danois (1975)</bibRefCitation>
to
<taxonomicName id="64884D33FFFAF25FFB964784FB34FE55" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[1050,1173,393,419]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFAF25FFB964784FB34FE55" box="[1050,1173,393,419]" italics="true" pageId="5" pageNumber="143">L. lugubris</emphasis>
</taxonomicName>
. Because this variation was described after 1960 and thus not available according to Article 15.2 of the International Code of Zoological Nomenclature (ICZN, 1999). Moreover, the original drawing of “
<taxonomicName id="64884D33FFFAF25FFCF647C1FBD6FE1F" authorityName="Le Danois" authorityYear="1975" box="[890,1143,463,489]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="variety" species="lugubris" variety="madagascariensis">
var.
<emphasis id="91FCEAA2FFFAF25FFC2147C1FBD6FE1F" box="[941,1143,463,489]" italics="true" pageId="5" pageNumber="143">madagascariensis</emphasis>
</taxonomicName>
” has six dorsal-fin spines and is clearly different from the definition of
<taxonomicName id="64884D33FFFAF25FFD3847FCFC92FDFD" baseAuthorityName="Alcock" baseAuthorityYear="1894" box="[692,819,497,523]" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="91FCEAA2FFFAF25FFD3847FCFC92FDFD" box="[692,819,497,523]" italics="true" pageId="5" pageNumber="143">L. lugubris</emphasis>
</taxonomicName>
. Base on the original drawing and description, “
<taxonomicName id="64884D33FFFAF25FFAFD47FFFEC0FDD8" authorityName="Le Danois" authorityYear="1975" class="Actinopterygii" family="Lophiidae" genus="Chirolophius" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="variety" species="lugubris" variety="madagascariensis">
var.
<emphasis id="91FCEAA2FFFAF25FFF1B441AFEC0FDD8" box="[151,353,532,558]" italics="true" pageId="5" pageNumber="143">madagascariensis</emphasis>
</taxonomicName>
” is likely misidentification of
<taxonomicName id="64884D33FFFAF25FFD34441AFBEDFDD8" authority="(Ragan, 1921)" baseAuthorityName="Ragan" baseAuthorityYear="1921" box="[696,1100,532,558]" class="Actinopterygii" family="Lophiidae" genus="Lophiodes" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="143" phylum="Chordata" rank="species" species="insidiator">
<emphasis id="91FCEAA2FFFAF25FFD34441AFC00FDD8" box="[696,929,532,558]" italics="true" pageId="5" pageNumber="143">Lophiodes insidiator</emphasis>
(Ragan, 1921)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,207 @@
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<mods:title>Integrative taxonomy resolves the cryptic and pseudo-cryptic Radula buccinifera complex (Porellales, Jungermanniopsida), including two reinstated and five new species</mods:title>
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<mods:namePart>Renner, Matt A. M.</mods:namePart>
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<treatment ID-GBIF-Taxon="152024753" LSID="urn:lsid:plazi:treatment:2B21D3AE52DBF4723972687E2FB1F3B3" httpUri="http://treatment.plazi.org/id/2B21D3AE52DBF4723972687E2FB1F3B3" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
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<taxonomicName LSID="2B21D3AE-52DB-F472-3972-687E2FB1F3B3" authority="Pearson. Journal of the Linnean Society, Botany 46: 29. 1922" authorityName="Pearson. Journal of the Linnean Society, Botany 46: 29." authorityYear="1922" class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Radula farmeri" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="farmeri">Radula farmeri Pearson. Journal of the Linnean Society, Botany 46: 29. 1922</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="type">
<paragraph pageId="62" pageNumber="63">Type.</paragraph>
<paragraph pageId="62" pageNumber="63">
Ignambi on rocks, 3000 ft, New Caledonia, 30 Jul 1914,
<emphasis italics="true" pageId="62" pageNumber="63">R.H. Compton</emphasis>
. 1530 N.C. (holotype: BM000825023!).
</paragraph>
<paragraph pageId="62" pageNumber="63">
The type specimens of
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="62" pageNumber="63">Radula farmeri</emphasis>
</taxonomicName>
and
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="novae-hollandiae">
<emphasis italics="true" pageId="62" pageNumber="63">Radula novae-hollandiae</emphasis>
</taxonomicName>
(S-B43474!) share the same suite of characteristics;the leaf-lobe apex is obtuse to subacute, the lobules are longitudinally rectangular with a prominent notch in their apex within which sits the lobule papilla, the lobule marginal cells are irregularly crenulate, and the two specimens are also of equivalent size. The similarity between the types suggest the only substantive difference between
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="62" pageNumber="63">Radula farmeri</emphasis>
</taxonomicName>
and
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="novae-hollandiae">
<emphasis italics="true" pageId="62" pageNumber="63">Radula novae-hollandiae</emphasis>
</taxonomicName>
might be the perianth mouth, which is fimbriate in the former and entire in the latter. We have only Pearsons (1922) observations on this, as the type specimen of
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="62" pageNumber="63">Radula farmeri</emphasis>
</taxonomicName>
in BM today bears no perianths.
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="australian specimens of r. farmeri">
<paragraph pageId="62" pageNumber="63">
Australian specimens of
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="62" pageNumber="63">Radula farmeri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="62" pageNumber="63">
Australia: Norfolk Island: Mt Pitt Road, Mount Pitt Reserve, 230 m,
<geoCoordinate degrees="29" direction="south" minutes="1.5" orientation="latitude" precision="92" value="-29.025">29°1.5'S</geoCoordinate>
,
<geoCoordinate degrees="167" direction="east" minutes="56.25" orientation="longitude" precision="9" value="167.9375">167°56.25'E</geoCoordinate>
, 2 Dec 1984,
<emphasis italics="true" pageId="62" pageNumber="63">H. Streimann 31867</emphasis>
, CANB650456, is
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Radula</emphasis>
</taxonomicName>
subg.
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Odontoradula" higherTaxonomySource="GBIF" kingdom="Viridiplantae" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Streptophyta" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Odontoradula</emphasis>
</taxonomicName>
sp. indet.; Mount Pitt Reserve, Filmy Fern Trail, off Selwyn Pine Road,
<geoCoordinate degrees="29" direction="south" minutes="01" orientation="latitude" precision="925" value="-29.016666">29°01'S</geoCoordinate>
,
<geoCoordinate degrees="167" direction="east" minutes="58" orientation="longitude" precision="925" value="167.96666">167°58'E</geoCoordinate>
, 130 m, 3 Dec 1984,
<emphasis italics="true" pageId="62" pageNumber="63">H. Streimann 32084A</emphasis>
, CANB650459 is
<taxonomicName authorityName="M. A. M. Renner" authorityYear="2013" class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="anisotoma">
<emphasis italics="true" pageId="62" pageNumber="63">Radula anisotoma</emphasis>
</taxonomicName>
mixed with
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Radula</emphasis>
</taxonomicName>
subg.
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Odontoradula" higherTaxonomySource="GBIF" kingdom="Viridiplantae" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Streptophyta" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Odontoradula</emphasis>
</taxonomicName>
sp. indet.; Mount Pitt Reserve, Filmy Fern Trail, off Selwyn Pine Road,
<geoCoordinate degrees="29" direction="south" minutes="1.3" orientation="latitude" precision="92" value="-29.021667">29°1.3'S</geoCoordinate>
,
<geoCoordinate degrees="167" direction="east" minutes="57.6" orientation="longitude" precision="92" value="167.96">167°57.6'E</geoCoordinate>
, 130 m, 3 Dec 1984,
<emphasis italics="true" pageId="62" pageNumber="63">H. Streimann 32078</emphasis>
(CANB650457, NICH, NY, EGR, H) is
<taxonomicName authorityName="M. A. M. Renner" authorityYear="2013" class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="62" pageNumber="63" phylum="Bryophyta" rank="species" species="anisotoma">
<emphasis italics="true" pageId="62" pageNumber="63">Radula anisotoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="remarks">
<paragraph pageId="63" pageNumber="64">
<pageBreakToken pageId="63" pageNumber="64" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="63" pageNumber="64">
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="63" pageNumber="64" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="63" pageNumber="64">Radula farmeri</emphasis>
</taxonomicName>
was recorded for Australia by
<bibRefCitation author="So, ML" journalOrPublisher="Journal of the Hattori Botanical Laboratory" pageId="69" pageNumber="70" pagination="207 - 232" publicationUrl="10.1093/bioinformatics/btl446" refId="B118" refString="So, ML, 2006. Radula (Radulaceae, Marchantiophyta) in the south pacific. Journal of the Hattori Botanical Laboratory 99: 207 - 232, 10.1093/bioinformatics/btl446" title="Radula (Radulaceae, Marchantiophyta) in the south pacific." url="10.1093/bioinformatics/btl446" volume="99" year="2006">So (2006)</bibRefCitation>
. The specimens CANB650456 and CANB650457 are sterile, and while specimen CANB650459 is fertile the perianth mouth is entire. As identification of
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Porellales" pageId="63" pageNumber="64" phylum="Bryophyta" rank="species" species="farmeri">
<emphasis italics="true" pageId="63" pageNumber="64">Radula farmeri</emphasis>
</taxonomicName>
is dependent on observation of perianths with fimbriate mouths; her determinations are not substantiated by the specimens, and this species should be excluded from the Australian flora.
</paragraph>
</subSubSection>
</treatment>
</document>