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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.93.1167" ID-GBIF-Dataset="704a30f8-6065-4db2-828f-822d9cfa3082" ID-PMC="PMC3095181" ID-Pensoft-Pub="1313-2970-93-9" ID-PubMed="21594077" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-93-9" ModsDocOrigin="ZooKeys 93" ModsDocTitle="The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda)" checkinTime="1451250468082" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Mikhaljova, Elena V. &amp; Chang, Hsueh-Wen" docDate="2011" docId="95DD5FFE3343DB6126C944477AED0505" docLanguage="en" docName="ZooKeys 93: 9-42" docOrigin="ZooKeys 93" docSource="http://dx.doi.org/10.3897/zookeys.93.1167" docTitle="Epanerchodus curtigonopus Golovatch, Mikhaljova &amp; Chang, 2011, sp. n." docType="treatment" docVersion="5" lastPageNumber="25" masterDocId="FFB1F12DFFDDFFEAFFF43B31FFF6B946" masterDocTitle="The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda)" masterLastPageNumber="42" masterPageNumber="9" pageNumber="23" updateTime="1668168924641" updateUser="ExternalLinkService">
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<mods:title>The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mikhaljova, Elena V.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chang, Hsueh-Wen</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>93</mods:number>
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<mods:start>9</mods:start>
<mods:end>42</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.93.1167</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.93.1167</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-93-9</mods:identifier>
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<treatment ID-GBIF-Taxon="159364985" LSID="urn:lsid:zoobank.org:act:CBDF0100-0AEF-4D12-B88A-3A59119AD5C0" httpUri="http://treatment.plazi.org/id/95DD5FFE3343DB6126C944477AED0505" lastPageId="16" lastPageNumber="25" pageId="14" pageNumber="23">
<subSubSection pageId="14" pageNumber="23" type="nomenclature">
<paragraph pageId="14" pageNumber="23">
<taxonomicName LSID="urn:lsid:zoobank.org:act:CBDF0100-0AEF-4D12-B88A-3A59119AD5C0" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus curtigonopus" order="Polydesmida" pageId="14" pageNumber="23" phylum="Arthropoda" rank="species" species="curtigonopus">Epanerchodus curtigonopus</taxonomicName>
<taxonomicNameLabel pageId="14" pageNumber="23">sp. n.</taxonomicNameLabel>
Figs 5460
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="23" type="type material">
<paragraph pageId="14" pageNumber="23">Type material:</paragraph>
<paragraph pageId="14" pageNumber="23">Holotype ♂ (NMNS-6560-001), Taiwan, Nantou County, Ren-ai Township, Mei-Feng, 15.10.2001, leg. S.H. Wu.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="23" type="name">
<paragraph pageId="14" pageNumber="23">Name:</paragraph>
<paragraph pageId="14" pageNumber="23">To emphasize the remarkably short gonopod telopodite.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="23" type="diagnosis">
<paragraph pageId="14" pageNumber="23">Diagnosis:</paragraph>
<paragraph pageId="14" pageNumber="23">
Differs from other
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus" order="Polydesmida" pageId="14" pageNumber="23" phylum="Arthropoda" rank="genus">Epanerchodus</taxonomicName>
species in the distal half of the gonopod telopodite being particularly stout, with the endomere represented only by a short spine supplied with a similarly short spine at its base (see also Key below). This new species seems to be particularly similar to
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus pinguis" order="Polydesmida" pageId="14" pageNumber="23" phylum="Arthropoda" rank="species" species="pinguis">Epanerchodus pinguis</taxonomicName>
sp. n. in sharing the gonopod which shows a densely setose coxa and a very stout telopodite (apparently, both being synapomorphies), but differs in the entire leg telopodites (prefemur+femur+postfemur+tibia+tarsus) being supplied with sphaerotrichomes ventrally, and the endomere and its basal process rudimentary and spiniform.
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="25" pageId="14" pageNumber="23" type="description">
<paragraph pageId="14" pageNumber="23">Description:</paragraph>
<paragraph pageId="14" pageNumber="23">Length ca 15 mm; width of pro- and metazona 1.3 and 2.5 mm, respectively. Coloration in alcohol pallid.</paragraph>
<paragraph pageId="14" pageNumber="23">
Superficially, also very similar to
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus pinguis" order="Polydesmida" pageId="14" pageNumber="23" phylum="Arthropoda" rank="species" species="pinguis">Epanerchodus pinguis</taxonomicName>
sp. n., except as follows.
</paragraph>
<paragraph lastPageId="15" lastPageNumber="24" pageId="14" pageNumber="23">
In width, collum &lt;head &lt;segment 2 = 3 &lt;4 &lt;5 &lt;6-15 (Figs 54), thereafter body gradually tapering towards telson (Fig. 55). Paraterga strongly developed, starting
<pageBreakToken pageId="15" pageNumber="24" start="start">from</pageBreakToken>
collum, set high, subhorizontal to very faintly upturned, lying slightly below dorsum only on collum and segment 2; mid-dorsum invariably extremely faintly convex, nearly flat; front shoulders mostly straight, drawn forward only on paraterga 2 and 3, thereafter directed increasingly caudolaterad and increasingly well rounded anterolaterally; caudal corner on paraterga 3-12 subrectangular and narrowly rounded, starting from 13th increasingly acutangular and evidently extending increasingly beyond rear tergal contour, but nearly pointed and beak-shaped only on segment 19 (Figs 54, 55). Paraterga 2-5 and all following pore-bearing segments with four, following poreless ones with three, small but evident incisions, each usually bearing a small seta on top at lateral margin. Metatergal sculpture typical, moderately developed, with three indistinct transverse rows of setiferous, polygonal bosses (Figs 54, 55). Tergal setae very short, mostly retained, a little longer only on collum and in rear row on metatergum 19. Stricture between pro- and metazona wide and smooth. Limbus very thin, microdenticulate. Epiproct rather short, conical (Figs 55, 56), only slightly bent ventrad, preapical papillae evident. Hypoproct semi-circular; caudal, paramedian, setiferous papillae distinct and well-separated (Fig. 56).
</paragraph>
<paragraph lastPageId="16" lastPageNumber="25" pageId="15" pageNumber="24">
Sterna without modifications, densely setose. Legs long and rather slender, only slightly enlarged (Figs 56, 57), ca 1.7-1.8 times as long as midbody height, prefemora not swollen dorsally, entire telopodite (prefemur+femur+postfemur+tibia+tarsus) with sphaerotrichomes ventrally (Fig. 58)
<pageBreakToken pageId="16" pageNumber="25" start="start">.</pageBreakToken>
</paragraph>
<paragraph pageId="16" pageNumber="25">Gonopods (Figs 57, 59, 60) with large, subquadrate, medially fused coxae carrying numerous setae ventrally. Telopodite especially stout, prefemoral portion relatively prominent, taking up about one-third of telopodite length, endomere (en) like a short spine supplied with another, somewhat shorter spine (p) near base; both accessory seminal chamber and hairy pulvillus evident.</paragraph>
<caption pageId="16" pageNumber="25">
<paragraph pageId="16" pageNumber="25">
Figures 54-57.
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus curtigonopus" order="Polydesmida" pageId="16" pageNumber="25" phylum="Arthropoda" rank="species" species="curtigonopus">Epanerchodus curtigonopus</taxonomicName>
sp. n., holotype. 54 anterior portion of body, dorsal view 55, 56 posterior portion of body, dorsal and ventral views, respectively 57 middle portion of body (with gonopods), ventral view. Photographed not to scale.
</paragraph>
</caption>
<caption pageId="16" pageNumber="25">
<paragraph pageId="16" pageNumber="25">
Figures 58-60.
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus curtigonopus" order="Polydesmida" pageId="16" pageNumber="25" phylum="Arthropoda" rank="species" species="curtigonopus">Epanerchodus curtigonopus</taxonomicName>
sp. n., holotype. 58 leg 9 59, 60 left gonopod, dorsomesal and ventrolateral views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="25" type="remarks">
<paragraph pageId="16" pageNumber="25">Remarks.</paragraph>
<paragraph pageId="16" pageNumber="25">This species is remarkable in showing an extremely short, simple, spiniform endomere (en), coupled with a single, even more rudimentary process (p) at en base.</paragraph>
<paragraph pageId="16" pageNumber="25">
In Taiwan,
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus curtigonopus" order="Polydesmida" pageId="16" pageNumber="25" phylum="Arthropoda" rank="species" species="curtigonopus">Epanerchodus curtigonopus</taxonomicName>
sp. n. occurs very locally, having been found only at a single locality (Map 2).
</paragraph>
</subSubSection>
</treatment>
</document>

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<figureCitation id="AF1FC01282CA7E2C5CD97E29DE73031E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">Fig. 13N</figureCitation>
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<taxonomicName id="6B22AD15F5B46093F3E365B8A41D163D" authorityName="T. -C. Luo, D. -N. Chen &amp; G. -Q. Zhang" authorityYear="2000" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia (Bensonella) tianxingqiaoensis" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="tianxingqiaoensis" subGenus="Bensonella">Boysidia (Bensonella) tianxingqiaoensis</taxonomicName>
<bibRefCitation id="E5DBA13632E23F36CF148C7BA17FAE26" author="Luo, T-C" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="157" pagination="147 - 151" refId="B34" refString="Luo, T-C, Chen, D-N, Zhang, G-Q, 2000. Two new species of Pupillidae from China (Pulmonata: Stylommatophora: Pupillidae). Acta Zootaxonomica Sinica 25 (2): 147 - 151" title="Two new species of Pupillidae from China (Pulmonata: Stylommatophora: Pupillidae)." volume="25" year="2000">Luo et al., 2000</bibRefCitation>
: 147, figs 1-4. Type locality. Tianxingqiao Town, Zhenning Bouyeizu Miaozu Zizhixian, Guizhou Province, China.
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.
<bibRefCitation id="2A16CE227321072B3BF36BBC83F2FDF5" DOI="https://doi.org/10.1127/arch.moll/151/131-139" author="Gojsina, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="157" pagination="131 - 139" refId="B24" refString="Gojsina, V, Chen, Z-U, Pall-Gergely, B, 2022. Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar. Archiv fuer Molluskenkunde 151 (2): 131 - 139, DOI: https://doi.org/10.1127/arch.moll/151/131-139" title="Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar." url="https://doi.org/10.1127/arch.moll/151/131-139" volume="151" year="2022">
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: 132-138, figs 1, 4b, 5c, d.
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This species was originally described in Guizhou Province, China and subsequently reported from
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[Mandalay Region, Myanmar] based on the historical A.E. Salisbury collection (
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;
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