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<emphasis id="B961BB030E70FF99FF4EFDAE59DE3D23" bold="true" box="[221,406,591,615]" italics="true" pageId="8" pageNumber="105">Agaue kurilensis</emphasis>
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(
<figureCitation id="132E7B940E70FF99FECDFD9759833DC8" box="[350,459,630,652]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[163,196,1306,1328]" captionStartId-1="8.[163,196,1689,1711]" captionTargetBox-0="[148,661,1077,1272]" captionTargetBox-1="[147,662,1400,1655]" captionTargetId-0="figure-456@8.[146,663,1075,1274]" captionTargetId-1="figure-470@8.[146,663,1399,1656]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionText-0="Fig. 27.ɹAgaue kurilensis, glass slide prepared by Dr. Makarova." captionText-1="Fig. 28.ɹAgaue kurilensis, holotype ˂ NSMT-Ac 14712. Idiosoma (A), gnathosoma (B), first leg (C). Scale bars for A=200 µm, B=50 µm, C=100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760479" figureDoi-1="http://doi.org/10.5281/zenodo.12760481" httpUri-0="https://zenodo.org/record/12760479/files/figure.png" httpUri-1="https://zenodo.org/record/12760481/files/figure.png" pageId="8" pageNumber="105">Figs. 27–28</figureCitation>
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series
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.ɹHolotype: ˂ NSMT-Ac 14712 (1/8052).
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<emphasis id="B961BB030E70FF99FF34FCE3595A3C5E" box="[167,274,770,794]" italics="true" pageId="8" pageNumber="105">Condition</emphasis>
.ɹEach body part is visible, and chaetotaxies of idiosoma, gnathosoma, palps, and legs are approximately recognizable.
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˂: Idiosoma length 1000, width 650; gnathosoma length 350, width 180. Idiosoma is somewhat broken.
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<paragraph id="8BAA67110E70FF99FF34FC335C253EA6" blockId="8.[140,669,699,1037]" lastBlockId="8.[707,1237,215,482]" pageId="8" pageNumber="105">
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.ɹThe specimen number given on the glass slide corresponds exactly to that in the original description by
<bibRefCitation id="EF841AE00E70FF99FC03FF365C053FAB" author="Makarova, N. G." box="[912,1101,215,239]" pageId="8" pageNumber="105" pagination="131 - 149" refId="ref4506" refString="Makarova, N. G. 1978. Marine mites (Acarina, Halacaridae) from the intertidal zone of the Gulf of Anadyr (Bering Sea). In Kussakin, O. G. (ed.): The intertidal zone of the Bering Sea and South-Eastern Kamchatka, pp. 131 - 149 (in Russian). Publishing House Nauka, Moscow." type="book chapter" year="1978">Makarova (1978)</bibRefCitation>
. The species name on the slide is given as l
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<taxonomicName id="4C151C920E70FF99FB9CFF1B5AAA3E70" authorityName="Makarova" authorityYear="1977" class="Arachnida" family="Halacaridae" genus="Agaue" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="105" phylum="Arthropoda" rank="species" species="kurilensis">Agaue aff kurilensis</taxonomicName>
.
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z Considering that the word l
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z is clearly separated from l
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<emphasis id="B961BB030E70FF99FCE8FEDE5BAB3E13" box="[891,995,319,343]" italics="true" pageId="8" pageNumber="105">kurilensis</emphasis>
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z in the species name on the slide (
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), it is probable that Dr. Makarova initially intended to describe this species as a subspecies of
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<emphasis id="B961BB030E70FF99FC45FE495C213EFB" box="[982,1129,423,447]" italics="true" pageId="8" pageNumber="105">Agaue affinis</emphasis>
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(=
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<emphasis id="B961BB030E70FF99FCBAFE2B5BFA3EA6" box="[809,946,458,482]" italics="true" pageId="8" pageNumber="105">Agaue parva</emphasis>
(
<bibRefCitation id="EF841AE00E70FF99FC53FE2B5C1F3EA6" author="Chilton, C." box="[960,1111,458,482]" pageId="8" pageNumber="105" pagination="190 - 192" refId="ref4159" refString="Chilton, C. 1883. On two marine mites. Transactions and Proceedings of the New Zealand Institute, Zoology 15: 190 - 192." type="journal article" year="1883">Chilton, 1883</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
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</treatment>
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211
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<emphasis id="B961BB030E79FF90FF25FC4859F53C85" bold="true" box="[182,445,937,961]" italics="true" pageId="1" pageNumber="98">Acanthopalpus hirsutus</emphasis>
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(
<figureCitation id="132E7B940E79FF90FEFAFC3059F73CA2" box="[361,447,977,999]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[729,761,473,495]" captionStartId-1="1.[729,761,1002,1024]" captionTargetBox-0="[715,1228,223,438]" captionTargetBox-1="[714,1229,567,968]" captionTargetId-0="figure-371@1.[713,1230,213,440]" captionTargetId-1="figure-385@1.[713,1230,566,969]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="Fig. 1.ɹAcanthopalpus hirsutus, glass slide prepared by Dr. Makarova." captionText-1="Fig. 2.ɹAcanthopalpus hirsutus, holotype ˂ NSMT- Ac 14693 (A, B, D) and paratype ˂ NSMT-Ac 14694 (C, E). Idiosoma (A), genitoanal region (B), body (C), gnathosoma (D), first leg (E). Scale bars for AC=100 µm, BDE=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760413" figureDoi-1="http://doi.org/10.5281/zenodo.12760417" httpUri-0="https://zenodo.org/record/12760413/files/figure.png" httpUri-1="https://zenodo.org/record/12760417/files/figure.png" pageId="1" pageNumber="98">Figs. 1–2</figureCitation>
)
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<paragraph id="8BAA67110E79FF90FD4AFC0B5CC53B31" blockId="1.[729,1214,1001,1141]" pageId="1" pageNumber="98">
Fig. 2.ɹ
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<emphasis id="B961BB030E79FF90FCBCFC0B5C483CBB" box="[815,1024,1002,1023]" italics="true" pageId="1" pageNumber="98">Acanthopalpus hirsutus</emphasis>
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, holotype ˂ NSMTAc 14693 (A, B, D) and paratype ˂ NSMT-Ac 14694 (C, E). Idiosoma (A), genitoanal region (B), body (C), gnathosoma (D), first leg (E). Scale bars for AC=100
<emphasis id="B961BB030E79FF90FC72FB815BA63B31" box="[993,1006,1120,1141]" italics="true" pageId="1" pageNumber="98">µ</emphasis>
m, BDE=50
<emphasis id="B961BB030E79FF90FBFAFB815C3E3B31" box="[1129,1142,1120,1141]" italics="true" pageId="1" pageNumber="98">µ</emphasis>
m.
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<emphasis id="B961BB030E79FF90FF34FBF9596A3B6B" box="[167,290,1048,1071]" italics="true" pageId="1" pageNumber="98">Type series</emphasis>
.ɹHolotype: ˂ NSMT-Ac 14693 (1/8087) and
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: 1 ˂ NSMT-Ac 14694 (2/8087).
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<emphasis id="B961BB030E79FF90FF34FB9E59513BD3" box="[167,281,1151,1175]" italics="true" pageId="1" pageNumber="98">Condition.</emphasis>
ɹThe glass slide is partly broken and missing. Each body part is visible in both type specimens. However, the chaetotaxies of idiosoma, gnathosoma, palps, and legs are only partly recognizable.
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˂: Idiosoma length 540, width 350; gnathosoma length 160, width 145.
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˂: Idiosoma length 475, width 320; gnathosoma length 160, width 145.
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.ɹThe specimen number given on the glass slide corresponds completely to that in the original description of
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. The species was described by
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based on the specimen from the Gulf of Anadyr on the Bering Sea.
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,
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) mentioned that this species might be synonymous with
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<emphasis id="B961BB030E79FF90FDA8F984597E39DB" italics="true" pageId="1" pageNumber="98">Acanthopalpus spinosus</emphasis>
(
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)
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as per Article
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the International Code of Zoological Nomenclature.
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(
<figureCitation id="132E7B940E79FF90FC33FB0E5BBE3A40" box="[928,1014,1263,1285]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="2.[163,195,443,465]" captionStartId-1="2.[163,196,868,890]" captionStartId-2="2.[163,196,1348,1370]" captionStartId-3="2.[729,762,668,690]" captionStartId-4="2.[729,762,1275,1297]" captionTargetBox-0="[148,656,223,407]" captionTargetBox-1="[147,662,540,834]" captionTargetBox-2="[147,662,1024,1314]" captionTargetBox-3="[714,1229,214,634]" captionTargetBox-4="[714,1229,824,1242]" captionTargetId-0="figure-172@2.[146,663,213,409]" captionTargetId-1="figure-186@2.[146,663,539,835]" captionTargetId-2="figure-218@2.[146,663,1023,1315]" captionTargetId-3="figure-250@2.[713,1230,213,635]" captionTargetId-4="figure-282@2.[713,1230,823,1243]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="2" captionTargetPageId-4="2" captionText-0="Fig. 3.ɹThalassarachna aculeata, glass slide prepared by Dr. Makarova." captionText-1="Fig. 4.ɹThalassarachna aculeata, holotype ˁ NSMT-Ac 14695. Body (A), genitoanal region (B), gnathosoma (C). Scale bars for A=100 µm, BC=50 µm." captionText-2="Fig. 5.ɹThalassarachna aculeata, paratype ˂ NSMT-Ac 14696. Body (A), genitoanal region (B), gnathosoma (C). Scale bars for A=100 µm, BC=50 µm." captionText-3="Fig. 6.ɹThalassarachna aculeata, paratype deutonymph NSMT-Ac 14697 (No. 4). Body (A), genitoanal region (B). Scale bars for A=100 µm, B=50 µm." captionText-4="Fig. 7.ɹThalassarachna aculeata, paratype deutonymph NSMT-Ac 14698 (No. 5). Body (A), genitoanal region (B). Scale bars for A=100 µm, B=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760419" figureDoi-1="http://doi.org/10.5281/zenodo.12760424" figureDoi-2="http://doi.org/10.5281/zenodo.12760431" figureDoi-3="http://doi.org/10.5281/zenodo.12760421" figureDoi-4="http://doi.org/10.5281/zenodo.12760427" httpUri-0="https://zenodo.org/record/12760419/files/figure.png" httpUri-1="https://zenodo.org/record/12760424/files/figure.png" httpUri-2="https://zenodo.org/record/12760431/files/figure.png" httpUri-3="https://zenodo.org/record/12760421/files/figure.png" httpUri-4="https://zenodo.org/record/12760427/files/figure.png" pageId="1" pageNumber="98">Figs. 3–7</figureCitation>
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.ɹHolotype: ˁ NSMT-Ac 14695 (2/8110) and
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: 1 ˂ NSMT-Ac 14696 (3/8110), 2 deutonymphs NSMT-Ac 14697– 14698 (4/8110 and 5/8110).
</paragraph>
<paragraph id="8BAA67110E79FF93FD4EFA565A363938" blockId="1.[707,1236,1322,1799]" lastBlockId="2.[140,670,1498,1799]" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98">
<emphasis id="B961BB030E79FF90FD4EFA565B003A8B" box="[733,840,1463,1487]" italics="true" pageId="1" pageNumber="98">Condition</emphasis>
.ɹThe body of the female is barely held in the mounting medium. The
<typeStatus id="54AED9B30E79FF90FBC0FA3B5AB33950" isOtherTypeStatus="true" pageId="1" pageNumber="98">other specimens</typeStatus>
(male and two deutonymphs) are preserved in reasonably good condition. Each body part is visible, and the chaetotaxies of idiosoma, gnathosoma, palps, and legs are almost recognizable in all specimens.
<typeStatus id="54AED9B30E79FF90FCE6F9665B9039DB" box="[885,984,1671,1695]" pageId="1" pageNumber="98" type="holotype">Holotype</typeStatus>
ˁ: Idiosoma length 860, width 660; gnathosoma length 320, width 180; the right first leg is missing.
<typeStatus id="54AED9B30E79FF90FBDAF92C5CEE39A1" box="[1097,1190,1741,1765]" pageId="1" pageNumber="98" type="paratype">Paratype</typeStatus>
˂: Idiosoma length 750, width 500; gnathosoma length 280, width 160.
<typeStatus id="54AED9B30E7AFF93FE0FFA3B59B13AB6" box="[412,505,1498,1522]" pageId="2" pageNumber="99" type="paratype">Paratype</typeStatus>
deutonymphs: NSMT-Ac 14697 (4/8110): Idiosoma length 880, width 640; gnathosoma length 250, width 150; NSMT-Ac 14698 (5/8110): Idiosoma length 650, width 480; gnathosoma length 250, width 150.
</paragraph>
<caption id="DF6A37990E7AFF93FF30FE5A59393EAA" ID-DOI="http://doi.org/10.5281/zenodo.12760419" ID-Zenodo-Dep="12760419" httpUri="https://zenodo.org/record/12760419/files/figure.png" pageId="2" pageNumber="99" startId="2.[163,195,443,465]" targetBox="[148,656,223,407]" targetPageId="2" targetType="figure">
<paragraph id="8BAA67110E7AFF93FF30FE5A59393EAA" blockId="2.[163,646,442,494]" pageId="2" pageNumber="99">
Fig. 3.ɹ
<taxonomicName id="4C151C920E7AFF93FF66FE5A59983E94" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[245,464,443,464]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B961BB030E7AFF93FF66FE5A59983E94" box="[245,464,443,464]" italics="true" pageId="2" pageNumber="99">Thalassarachna aculeata</emphasis>
</taxonomicName>
, glass slide prepared by Dr. Makarova.
</paragraph>
</caption>
<caption id="DF6A37990E7AFF93FD4AFD7D5BAF3C4E" ID-DOI="http://doi.org/10.5281/zenodo.12760421" ID-Zenodo-Dep="12760421" httpUri="https://zenodo.org/record/12760421/files/figure.png" pageId="2" pageNumber="99" startId="2.[729,762,668,690]" targetBox="[714,1229,214,634]" targetPageId="2" targetType="figure">
<paragraph id="8BAA67110E7AFF93FD4AFD7D5BAF3C4E" blockId="2.[729,1213,667,778]" pageId="2" pageNumber="99">
Fig. 6.ɹ
<taxonomicName id="4C151C920E7AFF93FCA3FD7D5C753DF5" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[816,1085,668,689]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B961BB030E7AFF93FCA3FD7D5C753DF5" box="[816,1085,668,689]" italics="true" pageId="2" pageNumber="99">Thalassarachna aculeata</emphasis>
</taxonomicName>
, paratype deutonymph NSMT-Ac 14697 (No. 4). Body (A), genitoanal region (B). Scale bars for A=100
<emphasis id="B961BB030E7AFF93FCC6FD145B2A3C4E" box="[853,866,757,778]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7AFF93FC51FD145B873C4E" box="[962,975,757,778]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="DF6A37990E7AFF93FF30FC85590E3C95" ID-DOI="http://doi.org/10.5281/zenodo.12760424" ID-Zenodo-Dep="12760424" httpUri="https://zenodo.org/record/12760424/files/figure.png" pageId="2" pageNumber="99" startId="2.[163,196,868,890]" targetBox="[147,662,540,834]" targetPageId="2" targetType="figure">
<paragraph id="8BAA67110E7AFF93FF30FC85590E3C95" blockId="2.[163,647,866,977]" pageId="2" pageNumber="99">
Fig. 4.ɹ
<taxonomicName id="4C151C920E7AFF93FF6AFC8559A73C3D" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[249,495,868,889]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B961BB030E7AFF93FF6AFC8559A73C3D" box="[249,495,868,889]" italics="true" pageId="2" pageNumber="99">Thalassarachna aculeata</emphasis>
</taxonomicName>
, holotype ˁ NSMT-Ac 14695. Body (A), genitoanal region (B), gnathosoma (C). Scale bars for A=100
<emphasis id="B961BB030E7AFF93FDF1FC7E5A273CF0" box="[610,623,927,948]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m, BC=50
<emphasis id="B961BB030E7AFF93FEB2FC5D59663C95" box="[289,302,956,977]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="DF6A37990E7AFF93FD4AFB1A5BAF3A2D" ID-DOI="http://doi.org/10.5281/zenodo.12760427" ID-Zenodo-Dep="12760427" httpUri="https://zenodo.org/record/12760427/files/figure.png" pageId="2" pageNumber="99" startId="2.[729,762,1275,1297]" targetBox="[714,1229,824,1242]" targetPageId="2" targetType="figure">
<paragraph id="8BAA67110E7AFF93FD4AFB1A5BAF3A2D" blockId="2.[729,1213,1274,1385]" pageId="2" pageNumber="99">
Fig. 7.ɹ
<taxonomicName id="4C151C920E7AFF93FCA3FB1A5C753A54" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[816,1085,1275,1296]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B961BB030E7AFF93FCA3FB1A5C753A54" box="[816,1085,1275,1296]" italics="true" pageId="2" pageNumber="99">Thalassarachna aculeata</emphasis>
</taxonomicName>
, paratype deutonymph NSMT-Ac 14698 (No. 5). Body (A), genitoanal region (B). Scale bars for A=100
<emphasis id="B961BB030E7AFF93FCC6FAB55B2A3A2D" box="[853,866,1364,1385]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7AFF93FC51FAB55B873A2D" box="[962,975,1364,1385]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="DF6A37990E7AFF93FF30FAA5590E3AF6" ID-DOI="http://doi.org/10.5281/zenodo.12760431" ID-Zenodo-Dep="12760431" httpUri="https://zenodo.org/record/12760431/files/figure.png" pageId="2" pageNumber="99" startId="2.[163,196,1348,1370]" targetBox="[147,662,1024,1314]" targetPageId="2" targetType="figure">
<paragraph id="8BAA67110E7AFF93FF30FAA5590E3AF6" blockId="2.[163,647,1347,1458]" pageId="2" pageNumber="99">
Fig. 5.ɹ
<taxonomicName id="4C151C920E7AFF93FF6AFAA559B83A1D" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[249,496,1348,1369]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B961BB030E7AFF93FF6AFAA559B83A1D" box="[249,496,1348,1369]" italics="true" pageId="2" pageNumber="99">Thalassarachna aculeata</emphasis>
</taxonomicName>
, paratype ˂ NSMT-Ac 14696. Body (A), genitoanal region (B), gnathosoma (C). Scale bars for A=100
<emphasis id="B961BB030E7AFF93FDF1FA9E5A273AD0" box="[610,623,1407,1428]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m, BC=50
<emphasis id="B961BB030E7AFF93FEB2FA7C59663AF6" box="[289,302,1437,1458]" italics="true" pageId="2" pageNumber="99">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="8BAA67110E7AFF92FF34F96659B93FAB" blockId="2.[140,670,1498,1799]" lastBlockId="3.[140,497,215,239]" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99">
<emphasis id="B961BB030E7AFF93FF34F966594339DB" box="[167,267,1671,1695]" italics="true" pageId="2" pageNumber="99">Remarks.</emphasis>
ɹThe specimen number of the
<typeStatus id="54AED9B30E7AFF93FDF6F96658F23986" pageId="2" pageNumber="99" type="holotype">holotype</typeStatus>
given on the glass slide corresponds completely to that in the original description of
<bibRefCitation id="EF841AE00E7AFF93FF1FF90E59013843" author="Makarova, N. G." box="[140,329,1775,1799]" pageId="2" pageNumber="99" pagination="131 - 149" refId="ref4506" refString="Makarova, N. G. 1978. Marine mites (Acarina, Halacaridae) from the intertidal zone of the Gulf of Anadyr (Bering Sea). In Kussakin, O. G. (ed.): The intertidal zone of the Bering Sea and South-Eastern Kamchatka, pp. 131 - 149 (in Russian). Publishing House Nauka, Moscow." type="book chapter" year="1978">Makarova (1978)</bibRefCitation>
. On the other hand, the sex and specimen number of the
<typeStatus id="54AED9B30E7AFF93FC4AFA745C773AE9" box="[985,1087,1429,1453]" pageId="2" pageNumber="99" type="paratype">paratypes</typeStatus>
given on the slide do not match those in the original description (ˁ, ˂, deutonymph: 3–22/8111–8118). Judging from the general features of the specimens on the slide, it is probable that the female and deutonymph specimens on the slides are the
<typeStatus id="54AED9B30E7AFF93FD50F9845B573939" box="[707,799,1637,1661]" pageId="2" pageNumber="99" type="paratype">paratype</typeStatus>
of
<emphasis id="B961BB030E7AFF93FCDEF9845C2F3939" box="[845,1127,1637,1661]" italics="true" pageId="2" pageNumber="99">
<taxonomicName id="4C151C920E7AFF93FCDEF9845C2B3939" baseAuthorityName="Makarova" baseAuthorityYear="1978" box="[845,1123,1637,1661]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeata">Thalassarachna aculeata</taxonomicName>
.
</emphasis>
The species name on the glass slide is given as
<taxonomicName id="4C151C920E7AFF93FBEEF9665B1A3986" authorityName="Makarova" authorityYear="1978" class="Arachnida" family="Halacaridae" genus="Halacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="B961BB030E7AFF93FBEEF9665B1A3986" italics="true" pageId="2" pageNumber="99">Halacarus aculeatus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF841AE00E7AFF93FCF2F94B5C493986" author="Bartsch, I." box="[865,1025,1706,1730]" pageId="2" pageNumber="99" pagination="1223 - 1236" refId="ref4041" refString="Bartsch, I. 1997. Thalassarachna and Halacarellus (Halacaridae: Acari): two separate genera. Journal of Natural History 31: 1223 - 1236." type="journal article" year="1997">Bartsch (1997)</bibRefCitation>
moved this species to the genus
<taxonomicName id="4C151C920E7AFF93FCFBF92C5C5C39A1" authorityName="Packard" authorityYear="1871" box="[872,1044,1741,1765]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B961BB030E7AFF93FCFBF92C5C5C39A1" box="[872,1044,1741,1765]" italics="true" pageId="2" pageNumber="99">Thalassarachna</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF841AE00E7AFF93FBB8F92C5C9B39A1" author="Bartsch, I." box="[1067,1235,1741,1765]" pageId="2" pageNumber="99" pagination="1 - 170" refId="ref4101" refString="Bartsch, I. 2009. Checklist of marine and freshwater halacarid mite genera and species (Halacaridae; Acari) with notes on synonyms, habitats, distribution and descriptions of the taxa. Zootaxa 1998: 1 - 170." type="journal article" year="2009">Bartsch (2009)</bibRefCitation>
commented that this species might be a synonym of
<taxonomicName id="4C151C920E7BFF92FF3AFF3959A53FAB" authority="(Trouessart, 1902)" baseAuthorityName="Trouessart" baseAuthorityYear="1902" box="[169,493,215,239]" class="Arachnida" family="Halacaridae" genus="Thalassarachna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="B961BB030E7BFF92FF3AFF3959573FAB" box="[169,287,216,239]" italics="true" pageId="3" pageNumber="100">T. princeps</emphasis>
(
<bibRefCitation id="EF841AE00E7BFF92FEBEFF3659AD3FAB" author="Trouessart, E. L." box="[301,485,215,239]" pageId="3" pageNumber="100" pagination="66 - 70" refId="ref4732" refString="Trouessart, E. L. 1902. Note preliminaire sur les acariens marins (Halacaridae) recueillis par S. A. S. le Prince de Monaco, dans les mers arctiques. Bulletin de la Societe Zoologique de France 27: 66 - 70." type="journal article" year="1902">Trouessart, 1902</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="4C151C920E7CFF95FD4DFC805CF03C3D" authority="Makarova, 1974" authorityName="Makarova" authorityYear="1974" box="[734,1208,865,889]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B961BB030E7CFF95FD4DFC805C4B3C3D" bold="true" box="[734,1027,865,889]" italics="true" pageId="4" pageNumber="101">Copidognathus globulosus</emphasis>
<bibRefCitation id="EF841AE00E7CFF95FB99FC805CF03C3D" author="Makarova, N. G." box="[1034,1208,865,889]" pageId="4" pageNumber="101" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova, 1974</bibRefCitation>
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<paragraph id="8BAA67110E7CFF95FC1DFC695C413CDA" blockId="4.[734,1208,865,926]" box="[910,1033,904,926]" pageId="4" pageNumber="101">
(
<figureCitation id="132E7B940E7CFF95FC06FC695C4A3CDA" box="[917,1026,904,926]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[163,196,598,620]" captionStartId-1="5.[163,195,1153,1175]" captionTargetBox-0="[147,661,215,564]" captionTargetBox-1="[147,662,722,1118]" captionTargetId-0="figure-233@5.[146,663,213,565]" captionTargetId-1="figure-253@5.[146,663,721,1119]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Fig. 15.ɹCopidognathus globulosus, glass slide prepared by Dr. Makarova. A: front side, B: back side." captionText-1="Fig. 16.ɹCopidognathus globulosus, holotype ˁ NSMT-Ac 14701. Idiosoma (A), genitoanal region (B), gnathosoma (C), first and second legs (D). Scale bars for A=100 µm, BCD=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760452" figureDoi-1="http://doi.org/10.5281/zenodo.12760458" httpUri-0="https://zenodo.org/record/12760452/files/figure.png" httpUri-1="https://zenodo.org/record/12760458/files/figure.png" pageId="4" pageNumber="101">Figs. 15–16</figureCitation>
)
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<paragraph id="8BAA67110E7CFF95FD4EFC325B593B49" blockId="4.[707,1237,976,1730]" pageId="4" pageNumber="101">
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series
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.ɹHolotype: ˁ NSMT-Ac 14701 (8103).
</paragraph>
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<emphasis id="B961BB030E7CFF95FD4EFBF65B003B6B" box="[733,840,1047,1071]" italics="true" pageId="4" pageNumber="101">Condition</emphasis>
.ɹThe specimen is mounted on a square cover slip that is partly broken. Each body part is visible, and chaetotaxies of idiosoma, gnathosoma, palps, and legs are almost recognizable.
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ˁ: Idiosoma length 500, width 410; gnathosoma length 120, width 100.
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Fig. 13.ɹ
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<emphasis id="B961BB030E7CFF95FE96FB6959893BD9" box="[261,449,1160,1181]" italics="true" pageId="4" pageNumber="101">Halacarellus longus</emphasis>
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[
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], ˂ NSMT-Ac 14708 (No. 6) mounted on the glass slide of
<taxonomicName id="4C151C920E7CFF95FEBEFB235A433B93" authorityName="Makarova" authorityYear="1978" box="[301,523,1218,1239]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="rombus">
<emphasis id="B961BB030E7CFF95FEBEFB235A433B93" box="[301,523,1218,1239]" italics="true" pageId="4" pageNumber="101">Copidognathus rombus</emphasis>
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. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100
<emphasis id="B961BB030E7CFF95FE33FB1F59E53A57" box="[416,429,1278,1299]" italics="true" pageId="4" pageNumber="101">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7CFF95FD9FFB1F5A513A57" box="[524,537,1278,1299]" italics="true" pageId="4" pageNumber="101">µ</emphasis>
m.
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.ɹThe specimen number given on the glass slide (8103) does not match that in the original description (ˁ: 1/8023) by
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. However, the collecting locality shown on the slide is consistent with that given by
<bibRefCitation id="EF841AE00E7CFF95FD50FA745BC83AE9" author="Makarova, N. G." box="[707,896,1429,1453]" pageId="4" pageNumber="101" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova (1974)</bibRefCitation>
. Further, the general features of this
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specimen on the slide accord well with those in
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Fig.
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in the original description. Therefore, it is certain that this specimen is the
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<emphasis id="B961BB030E7CFF95FD50F9FE5BAA3973" box="[707,994,1567,1591]" italics="true" pageId="4" pageNumber="101">Copidognathus globulosus</emphasis>
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. The species name on the glass slide is given as
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. However,
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described this species under the name of
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<emphasis id="B961BB030E7CFF95FC5FF9665AAD3986" italics="true" pageId="4" pageNumber="101">Copidognathus globulosus</emphasis>
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based on a specimen from Iturup Island.
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<mods:title id="92CC9E5BE287B5BD56469CC1D3E49DDC">Bulletin of the National Museum of Nature and Science. Series A, Zoology</mods:title>
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<mods:date id="0AC122B34E49D1C381AA586C771AC757">2023</mods:date>
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<mods:number id="AFFD1BB6374091EBDD1009D558B7C931">49</mods:number>
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<mods:identifier id="8249B80C9BC29A847AD9A2029367BB37" type="DOI">10.50826/bnmnszool.49.3_97</mods:identifier>
<mods:identifier id="EA20653034734FAB9D170D6ECA6DF99A" type="ISSN">2434-091X</mods:identifier>
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<treatment id="03BCD6070E7DFF97FF21FAEF5AC13E3E" ID-DOI="http://doi.org/10.5281/zenodo.12760443" ID-Zenodo-Dep="12760443" LSID="urn:lsid:plazi:treatment:03BCD6070E7DFF97FF21FAEF5AC13E3E" httpUri="http://treatment.plazi.org/id/03BCD6070E7DFF97FF21FAEF5AC13E3E" lastPageId="6" lastPageNumber="102" pageId="5" pageNumber="102">
<subSubSection id="C30F349A0E7DFF94FF21FAEF5A3F3A62" box="[178,631,1294,1318]" pageId="5" pageNumber="102" type="nomenclature">
<paragraph id="8BAA67110E7DFF94FF21FAEF5A3F3A62" blockId="5.[178,631,1294,1318]" box="[178,631,1294,1318]" pageId="5" pageNumber="102">
<heading id="D0E2D07D0E7DFF94FF21FAEF5A3F3A62" box="[178,631,1294,1318]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="102" reason="4">
<taxonomicName id="4C151C920E7DFF94FF21FAEF5A3F3A62" authority="Makarova, 1974" authorityName="Makarova" authorityYear="1974" box="[178,631,1294,1318]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B961BB030E7DFF94FF21FAEF598A3A62" bold="true" box="[178,450,1294,1318]" italics="true" pageId="5" pageNumber="102">Copidognathus pacificus</emphasis>
<bibRefCitation id="EF841AE00E7DFF94FE5BFAEF5A3F3A62" author="Makarova, N. G." box="[456,631,1294,1318]" pageId="5" pageNumber="102" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova, 1974</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="C30F349A0E7DFF97FEC4FADC5AC13E3E" lastPageId="6" lastPageNumber="103" pageId="5" pageNumber="102" type="description">
<paragraph id="8BAA67110E7DFF94FEC4FADC599A3A17" blockId="5.[343,466,1341,1363]" box="[343,466,1341,1363]" pageId="5" pageNumber="102">
(
<figureCitation id="132E7B940E7DFF94FECDFADC59833A17" box="[350,459,1341,1363]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[729,762,441,463]" captionStartId-1="5.[729,762,1088,1110]" captionTargetBox-0="[715,1218,221,405]" captionTargetBox-1="[714,1229,536,1055]" captionTargetId-0="figure-289@5.[713,1230,213,407]" captionTargetId-1="figure-302@5.[713,1230,535,1056]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Fig. 17.ɹCopidognathus pacificus, glass slide prepared by Dr. Makarova." captionText-1="Fig. 18.ɹCopidognathus pacificus, holotype ˁ NSMT-Ac 14702. Idiosoma (A), genitoanal region (B), first leg (C) (Phase-contrast micrographs). Scale bars for ABC=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760450" figureDoi-1="http://doi.org/10.5281/zenodo.12760456" httpUri-0="https://zenodo.org/record/12760450/files/figure.png" httpUri-1="https://zenodo.org/record/12760456/files/figure.png" pageId="5" pageNumber="102">Figs. 17–18</figureCitation>
)
</paragraph>
<paragraph id="8BAA67110E7DFF94FF34FA9258A73AE9" blockId="5.[140,670,1392,1800]" pageId="5" pageNumber="102">
<emphasis id="B961BB030E7DFF94FF34FA92596A3ACE" box="[167,290,1395,1418]" italics="true" pageId="5" pageNumber="102">
<typeStatus id="54AED9B30E7DFF94FF34FA9258913ACE" box="[167,217,1395,1418]" pageId="5" pageNumber="102">Type</typeStatus>
series
</emphasis>
.ɹHolotype: ˁ NSMT-Ac 14702 (1/8031).
</paragraph>
<paragraph id="8BAA67110E7DFF94FF34FA565A583938" blockId="5.[140,670,1392,1800]" pageId="5" pageNumber="102">
<emphasis id="B961BB030E7DFF94FF34FA56595A3A8B" box="[167,274,1463,1487]" italics="true" pageId="5" pageNumber="102">Condition</emphasis>
.ɹThe idiosoma and first and second legs are visible, and the chaetotaxies of those body parts are approximately recognizable. On the contrary, the gnathosoma is not found, and the third and fourth legs are missing.
<typeStatus id="54AED9B30E7DFF94FDA9F9A35AD5391E" box="[570,669,1602,1626]" pageId="5" pageNumber="102" type="holotype">Holotype</typeStatus>
ˁ: Idiosoma length 350, width 215.
</paragraph>
<paragraph id="8BAA67110E7DFF97FF34F9665AC13E3E" blockId="5.[140,670,1392,1800]" lastBlockId="6.[140,669,215,378]" lastPageId="6" lastPageNumber="103" pageId="5" pageNumber="102">
<emphasis id="B961BB030E7DFF94FF34F966594C39DB" box="[167,260,1671,1695]" italics="true" pageId="5" pageNumber="102">Remarks</emphasis>
.ɹThe specimen number given on the glass slide partly corresponds to that in the original description (ˁ:1/31) of
<bibRefCitation id="EF841AE00E7DFF94FE46F92C5AD139A1" author="Makarova, N. G." box="[469,665,1741,1765]" pageId="5" pageNumber="102" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova (1974)</bibRefCitation>
. The general features of this type specimen and collecting locality shown on the slide are consistent with those given in the original description. Therefore, it is most likely that this specimen is the
<typeStatus id="54AED9B30E7DFF94FD61FAAE5B073A23" box="[754,847,1359,1383]" pageId="5" pageNumber="102" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C151C920E7DFF94FC17FAAE5CDD3A23" authorityName="Makarova" authorityYear="1974" box="[900,1173,1359,1383]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B961BB030E7DFF94FC17FAAE5CDD3A23" box="[900,1173,1359,1383]" italics="true" pageId="5" pageNumber="102">Copidognathus pacificus</emphasis>
</taxonomicName>
. The species name on the glass slide is given as
<taxonomicName id="4C151C920E7DFF94FB04FA935B803AE9" class="Arachnida" family="Halacaridae" genus="Copidognathopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B961BB030E7DFF94FB04FA935B803AE9" italics="true" pageId="5" pageNumber="102">Copidognathopsis orientalis</emphasis>
</taxonomicName>
. On the contrary, the specimen number given on the slide almost corresponds to that in the original description of
<taxonomicName id="4C151C920E7DFF94FB28FA3B5B6A3951" authorityName="Makarova" authorityYear="1974" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B961BB030E7DFF94FB28FA3B5B6A3951" italics="true" pageId="5" pageNumber="102">C. pacificus</emphasis>
</taxonomicName>
. At that time,
<taxonomicName id="4C151C920E7DFF94FC49FA1C5CD43951" box="[986,1180,1533,1557]" class="Arachnida" family="Halacaridae" genus="Copidognathopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B961BB030E7DFF94FC49FA1C5CD43951" box="[986,1180,1533,1557]" italics="true" pageId="5" pageNumber="102">Copidognathopsis</emphasis>
</taxonomicName>
was regarded as one subgenus in the genus
<taxonomicName id="4C151C920E7DFF94FBFCF9FE5B42391E" authorityName="Trouessart" authorityYear="1888" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B961BB030E7DFF94FBFCF9FE5B42391E" italics="true" pageId="5" pageNumber="102">Copidognathus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C151C920E7DFF94FC88F9A35C67391E" authorityName="Newell" authorityYear="1951" box="[795,1071,1602,1626]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B961BB030E7DFF94FC88F9A35C67391E" box="[795,1071,1602,1626]" italics="true" pageId="5" pageNumber="102">Copidognathus orientalis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EF841AE00E7DFF94FD76F9845BC83938" author="Newell, I. M." box="[741,896,1636,1661]" pageId="5" pageNumber="102" pagination="1 - 27" refId="ref4600" refString="Newell, I. M. 1951. Copidognathus curtus Hall, 1912 and other species of Copidognathus from western north America (Acari, Halacaridae). American Museum Novitates 1499: 1 - 27." type="journal article" year="1951">Newell (1951)</bibRefCitation>
based on a specimen from Attu Island in Alaska.
<bibRefCitation id="EF841AE00E7DFF94FC18F9665C0339DB" author="Makarova, N. G." box="[907,1099,1671,1695]" pageId="5" pageNumber="102" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova (1974)</bibRefCitation>
described a new species under the name of
<taxonomicName id="4C151C920E7DFF94FBA2F94B5BB739A1" authorityName="Makarova" authorityYear="1974" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="species" species="pacificus" subGenus="Copidognathopsis">
<emphasis id="B961BB030E7DFF94FBA2F94B5C9C3986" box="[1073,1236,1706,1730]" italics="true" pageId="5" pageNumber="102">Copidognathus</emphasis>
(
<emphasis id="B961BB030E7DFF94FD5FF92C5BC639A1" box="[716,910,1741,1765]" italics="true" pageId="5" pageNumber="102">Copidognathopsis</emphasis>
)
<emphasis id="B961BB030E7DFF94FC33F92C5BB739A1" box="[928,1023,1741,1765]" italics="true" pageId="5" pageNumber="102">pacificus</emphasis>
</taxonomicName>
, whereas the name of the new species was given as
<taxonomicName id="4C151C920E7DFF94FBA2F90E5C9C3843" authorityName="Trouessart" authorityYear="1888" box="[1073,1236,1775,1799]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B961BB030E7DFF94FBA2F90E5C9C3843" box="[1073,1236,1775,1799]" italics="true" pageId="5" pageNumber="102">Copidognathus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C151C920E7EFF97FF06FF3659813FAB" box="[149,457,215,239]" class="Arachnida" family="Halacaridae" genus="Copidognathopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B961BB030E7EFF97FF06FF3659813FAB" box="[149,457,215,239]" italics="true" pageId="6" pageNumber="103">Copidognathopsis orientalis</emphasis>
</taxonomicName>
) [
<emphasis id="B961BB030E7EFF97FE74FF395A4D3FAB" box="[487,517,216,239]" italics="true" pageId="6" pageNumber="103">sic</emphasis>
] in the summary. Makarova (1874) got
<taxonomicName id="4C151C920E7EFF97FE42FF1B5A783E56" authorityName="Makarova" authorityYear="1974" box="[465,560,250,274]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B961BB030E7EFF97FE42FF1B5A783E56" box="[465,560,250,274]" italics="true" pageId="6" pageNumber="103">pacificus</emphasis>
</taxonomicName>
confused with
<taxonomicName id="4C151C920E7EFF97FF51FEFC59603E71" box="[194,296,285,309]" class="Arachnida" family="Halacaridae" genus="Copidognathopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B961BB030E7EFF97FF51FEFC59603E71" box="[194,296,285,309]" italics="true" pageId="6" pageNumber="103">orientalis</emphasis>
</taxonomicName>
in writing the manuscript. The sex of the specimen indicated in
<figureCitation id="132E7B940E7EFF97FE59FEDE5A423E13" box="[458,522,319,343]" captionStart="Fig" captionStartId="2.[163,196,1348,1370]" captionTargetBox="[147,662,1024,1314]" captionTargetId="figure-218@2.[146,663,1023,1315]" captionTargetPageId="2" captionText="Fig. 5.ɹThalassarachna aculeata, paratype ˂ NSMT-Ac 14696. Body (A), genitoanal region (B), gnathosoma (C). Scale bars for A=100 µm, BC=50 µm." figureDoi="http://doi.org/10.5281/zenodo.12760431" httpUri="https://zenodo.org/record/12760431/files/figure.png" pageId="6" pageNumber="103">
Fig.
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in
<bibRefCitation id="EF841AE00E7EFF97FDA2FEDE589B3E3E" author="Makarova, N. G." pageId="6" pageNumber="103" pagination="276 - 288" refId="ref4343" refString="Makarova, N. G. 1974. Six new species of marine mites Copidognathus (Acarina, Halacaridae) from the coast of the Kuril Islands. Sbotnik Rabot, Instituta Biologii Morya 1: 276 - 288 (in Russian)." type="journal article" year="1974">Makarova (1974)</bibRefCitation>
is not female (самка), but male (самец).
</paragraph>
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<emphasis id="B961BB030E7EFF97FF2AFE5559F23E88" bold="true" box="[185,442,436,460]" italics="true" pageId="6" pageNumber="103">Copidognathus rombus</emphasis>
<bibRefCitation id="EF841AE00E7EFF97FE52FE545A273E89" author="Makarova, N. G." box="[449,623,437,461]" pageId="6" pageNumber="103" pagination="131 - 149" refId="ref4506" refString="Makarova, N. G. 1978. Marine mites (Acarina, Halacaridae) from the intertidal zone of the Gulf of Anadyr (Bering Sea). In Kussakin, O. G. (ed.): The intertidal zone of the Bering Sea and South-Eastern Kamchatka, pp. 131 - 149 (in Russian). Publishing House Nauka, Moscow." type="book chapter" year="1978">Makarova, 1978</bibRefCitation>
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(
<figureCitation id="132E7B940E7EFF97FECDFE0559833EBD" box="[350,459,484,506]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[162,195,1218,1240]" captionStartId-1="6.[163,196,1689,1711]" captionStartId-2="6.[729,762,1145,1167]" captionStartId-3="6.[729,762,1689,1711]" captionTargetBox-0="[147,660,1000,1183]" captionTargetBox-2="[714,1229,760,1112]" captionTargetBox-3="[147,1229,1298,1655]" captionTargetId-0="figure-262@6.[145,662,991,1185]" captionTargetId-2="figure-312@6.[713,1230,759,1113]" captionTargetPageId-0="6" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="Fig. 19.ɹCopidognathus rombus and Halacarellus longus [nomen nudum], glass slide prepared by Dr. Makarova." captionText-1="Fig. 20.ɹCopidognathus rombus, holotype ˂ NSMT-Ac 14703. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100 µm, B=50 µm." captionText-2="Fig. 21.ɹCopidognathus rombus, paratype ˂ NSMT-Ac 14704. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100 µm, B=50 µm." captionText-3="Fig. 22.ɹCopidognathus rombus, paratype ˁ NSMT-Ac 14705. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100 µm, B=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760464" figureDoi-2="http://doi.org/10.5281/zenodo.12760460" figureDoi-3="http://doi.org/10.5281/zenodo.12760466" httpUri-0="https://zenodo.org/record/12760464/files/figure.png" httpUri-2="https://zenodo.org/record/12760460/files/figure.png" httpUri-3="https://zenodo.org/record/12760466/files/figure.png" pageId="6" pageNumber="103">Figs. 19–22</figureCitation>
)
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.ɹHolotype: ˂ NSMT-Ac 14703 (1/8119),
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: 1 ˂ NSMT-Ac 14704 (2/8120) and 1ˁ NSMT-Ac 14705 (5/8120).
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<emphasis id="B961BB030E7EFF97FF34FD7B595A3DF6" box="[167,274,666,690]" italics="true" pageId="6" pageNumber="103">Condition</emphasis>
.ɹEach body part is visible in type specimens. However, chaetotaxies of idiosoma, gnathosoma, palps, and legs are partly recognizable.
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˂: Idiosoma length 450, width 325; gnathosoma length 105, width 95.
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˂: Idiosoma length 480, width 375; gnathosoma length 110, width 100; second legs are broken.
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ˁ: Idiosoma length 410, width 300; gnathosoma length 105, width 95; the distal part of the idiosoma is broken, and the right first and left third and fourth legs are missing.
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<emphasis id="B961BB030E7EFF97FD4EFEDE5B723E13" box="[733,826,319,343]" italics="true" pageId="6" pageNumber="103">Remarks</emphasis>
.ɹThe specimen number of the
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given on the glass slide corresponds to that in the original description in
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. However, the specimen number of the
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given on the slide does not match those in the original description (ˁ and ˂: 3, 4, 5/8120 and 2/8121). Although the general characteristics of the specimen on the slide are barely recognizable, the collecting date and locality given on the slide correspond to those in the original description. Consequently, the female specimen (2/8120) on the slide will probably be the
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(2/8121) in the original description. The species was described by
<bibRefCitation id="EF841AE00E7FFF96FE0CFEFC5A123E71" author="Makarova, N. G." box="[415,602,285,309]" pageId="7" pageNumber="104" pagination="131 - 149" refId="ref4506" refString="Makarova, N. G. 1978. Marine mites (Acarina, Halacaridae) from the intertidal zone of the Gulf of Anadyr (Bering Sea). In Kussakin, O. G. (ed.): The intertidal zone of the Bering Sea and South-Eastern Kamchatka, pp. 131 - 149 (in Russian). Publishing House Nauka, Moscow." type="book chapter" year="1978">Makarova (1978)</bibRefCitation>
based on a specimen from the Gulf of Anadyr in the Bering Sea.
</paragraph>
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Fig. 21.ɹ
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<emphasis id="B961BB030E7EFF97FCAFFB985C6A3BCA" box="[828,1058,1145,1166]" italics="true" pageId="6" pageNumber="103">Copidognathus rombus</emphasis>
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, paratype ˂ NSMT-Ac 14704. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100
<emphasis id="B961BB030E7EFF97FCC6FB335B2A3BA3" box="[853,866,1234,1255]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7EFF97FC51FB335B873BA3" box="[962,975,1234,1255]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="DF6A37990E7EFF97FF31FB23591C3A56" ID-DOI="http://doi.org/10.5281/zenodo.12760464" ID-Zenodo-Dep="12760464" httpUri="https://zenodo.org/record/12760464/files/figure.png" pageId="6" pageNumber="103" startId="6.[162,195,1218,1240]" targetBox="[147,660,1000,1183]" targetPageId="6" targetType="figure">
<paragraph id="8BAA67110E7EFF97FF31FB23591C3A56" blockId="6.[162,647,1217,1298]" pageId="6" pageNumber="103">
Fig. 19.ɹ
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<emphasis id="B961BB030E7EFF97FE97FB2359943B93" box="[260,476,1218,1239]" italics="true" pageId="6" pageNumber="103">Copidognathus rombus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C151C920E7EFF97FD82FB2359433BB0" class="Arachnida" family="Halacaridae" genus="Halacarellus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B961BB030E7EFF97FD82FB2359433BB0" italics="true" pageId="6" pageNumber="103">Halacarellus longus</emphasis>
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[
<emphasis id="B961BB030E7EFF97FE88FB0159D53BB0" box="[283,413,1247,1269]" italics="true" pageId="6" pageNumber="103">nomen nudum</emphasis>
], glass slide prepared by Dr. Makarova.
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</caption>
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<paragraph id="8BAA67110E7EFF97FF30F97859F83842" blockId="6.[163,647,1688,1799]" pageId="6" pageNumber="103">
Fig. 20.ɹ
<taxonomicName id="4C151C920E7EFF97FE96F97859A239EA" authorityName="Makarova" authorityYear="1978" box="[261,490,1689,1710]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="rombus">
<emphasis id="B961BB030E7EFF97FE96F97859A239EA" box="[261,490,1689,1710]" italics="true" pageId="6" pageNumber="103">Copidognathus rombus</emphasis>
</taxonomicName>
, holotype ˂ NSMT-Ac 14703. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100
<emphasis id="B961BB030E7EFF97FE8CF91359643843" box="[287,300,1778,1799]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7EFF97FE18F91359D03843" box="[395,408,1778,1799]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m.
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<paragraph id="8BAA67110E7EFF97FD4AF9785BAF3842" blockId="6.[729,1214,1688,1799]" pageId="6" pageNumber="103">
Fig. 22.ɹ
<taxonomicName id="4C151C920E7EFF97FCAFF9785C6A39EA" authorityName="Makarova" authorityYear="1978" box="[828,1058,1689,1710]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="rombus">
<emphasis id="B961BB030E7EFF97FCAFF9785C6A39EA" box="[828,1058,1689,1710]" italics="true" pageId="6" pageNumber="103">Copidognathus rombus</emphasis>
</taxonomicName>
, paratype ˁ NSMT-Ac 14705. Body (A), gnathosoma (B) (Phase-contrast micrographs). Scale bars for A=100
<emphasis id="B961BB030E7EFF97FCC6F9135B2A3843" box="[853,866,1778,1799]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m, B=50
<emphasis id="B961BB030E7EFF97FC51F9135B873843" box="[962,975,1778,1799]" italics="true" pageId="6" pageNumber="103">µ</emphasis>
m.
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</caption>
<paragraph id="8BAA67110E7FFF96FF34FE6459C03D40" blockId="7.[140,669,215,517]" pageId="7" pageNumber="104">
Apart from the
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of
<taxonomicName id="4C151C920E7FFF96FE2FFE6458F93EFB" authorityName="Makarova" authorityYear="1978" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="104" phylum="Arthropoda" rank="species" species="rombus">
<emphasis id="B961BB030E7FFF96FE2FFE6458F93EFB" italics="true" pageId="7" pageNumber="104">Copidognathus rombus</emphasis>
</taxonomicName>
, the present glass slide contains
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specimens with the name shown as
<taxonomicName id="4C151C920E7FFF96FD80FE2B589B3D41" class="Arachnida" family="Halacaridae" genus="Halacarellus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="104" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B961BB030E7FFF96FD80FE2B589B3D41" italics="true" pageId="7" pageNumber="104">Halacarellus longus</emphasis>
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[
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193
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<taxonomicName id="4C151C920E7FFF96FF33FDDE5AC13D13" authority="Makarova, 1978" authorityName="Makarova" authorityYear="1978" box="[160,649,575,599]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="104" phylum="Arthropoda" rank="species" species="beringiensis">
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[
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(
<figureCitation id="132E7B940E7FFF96FECDFD7059833DE2" box="[350,459,657,679]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[163,196,1346,1368]" captionStartId-1="7.[163,196,1689,1711]" captionTargetBox-0="[148,661,1121,1312]" captionTargetBox-1="[147,662,1440,1655]" captionTargetId-0="figure-238@7.[146,663,1119,1314]" captionTargetId-1="figure-253@7.[146,663,1439,1656]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Fig. 23.ɹCopidognathus beringiensis [nomen nudum], glass slide prepared by Dr. Makarova." captionText-1="Fig. 24.ɹCopidognathus beringiensis [nomen nudum], ˂ NSMT-Ac 14710. Body (A), genitoanal region (B), first leg (C). Scale bars for A=200 µm, BC=50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12760470" figureDoi-1="http://doi.org/10.5281/zenodo.12760474" httpUri-0="https://zenodo.org/record/12760470/files/figure.png" httpUri-1="https://zenodo.org/record/12760474/files/figure.png" pageId="7" pageNumber="104">Figs. 23–24</figureCitation>
)
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<emphasis id="B961BB030E7FFF96FF34FD0159553DB3" box="[167,285,736,759]" italics="true" pageId="7" pageNumber="104">
<typeStatus id="54AED9B30E7FFF96FF34FD0158913DB3" box="[167,217,736,759]" pageId="7" pageNumber="104">Type</typeStatus>
series
</emphasis>
.ɹ1˂ NSMT-Ac 14710 (1/8091).
</paragraph>
<paragraph id="8BAA67110E7FFF96FF34FCE358B53CE1" blockId="7.[140,669,733,1071]" pageId="7" pageNumber="104">
<emphasis id="B961BB030E7FFF96FF34FCE3595A3C5E" box="[167,274,770,794]" italics="true" pageId="7" pageNumber="104">Condition</emphasis>
. Each body part is visible and chaetotaxies of idiosoma, gnathosoma, palps, and legs are almost recognizable.
<typeStatus id="54AED9B30E7FFF96FE42FCA95A4E3C24" box="[465,518,840,864]" pageId="7" pageNumber="104">Type</typeStatus>
˂: Idiosoma length 570, width 390; gnathosoma length 160, width 110.
</paragraph>
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<emphasis id="B961BB030E7FFF96FF35FC4E594B3C83" box="[166,259,943,967]" italics="true" pageId="7" pageNumber="104">Remarks</emphasis>
.ɹThis species has not been properly described. Nevertheless, the species name,
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<emphasis id="B961BB030E7FFF96FDF3FC3359CF3B49" italics="true" pageId="7" pageNumber="104">Copidognathus beringiensis</emphasis>
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the list of animal species from the intertidal zone of eastern
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and the western coast of the Bering Sea (
<bibRefCitation id="EF841AE00E7FFF96FC05FF1B5C043E56" author="Makarova, N. G." box="[918,1100,250,274]" pageId="7" pageNumber="104" pagination="131 - 149" refId="ref4506" refString="Makarova, N. G. 1978. Marine mites (Acarina, Halacaridae) from the intertidal zone of the Gulf of Anadyr (Bering Sea). In Kussakin, O. G. (ed.): The intertidal zone of the Bering Sea and South-Eastern Kamchatka, pp. 131 - 149 (in Russian). Publishing House Nauka, Moscow." type="book chapter" year="1978">Makarova, 1978</bibRefCitation>
). The specimen number given on the glass slide cannot be found in any papers by Dr. Makarova.
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129
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(
<figureCitation id="13682A62FF89D90FFE9309DAFE79FE79" box="[357,452,450,472]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2N–P</figureCitation>
<figureCitation id="13682A62FF89D90FFE9309DAFE79FE79" box="[357,452,450,472]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2N–P</figureCitation>
)
</paragraph>
<paragraph id="8BEC36E7FF89D90FFF510A17FE97FD13" blockId="6.[140,670,527,1418]" pageId="6" pageNumber="169">
@ -112,7 +113,7 @@ HONSHU:
<geoCoordinate id="EE675020FF89D90FFE6F0A2AFE4CFDE8" box="[409,497,561,586]" degrees="33" direction="north" minutes="46" orientation="latitude" pageId="6" pageNumber="169" precision="925" value="33.766666">33°46′N</geoCoordinate>
<geoCoordinate id="EE675020FF89D90FFE0D0A2AFDDDFDE8" box="[507,608,561,586]" degrees="140" direction="east" minutes="10" orientation="longitude" pageId="6" pageNumber="169" precision="925" value="140.16667">140°10′E</geoCoordinate>
, coll. larva (
<figureCitation id="13682A62FF89D90FFF230A4DFE92FDCC" box="[213,303,597,621]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2N</figureCitation>
<figureCitation id="13682A62FF89D90FFF230A4DFE92FDCC" box="[213,303,597,621]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2N</figureCitation>
),
<collectedFrom id="384F751AFF89D90FFEB10A4DFDD4FDCC" box="[327,617,597,621]" pageId="6" pageNumber="169">
on
@ -127,7 +128,7 @@ on
, mat. 29.
<collectionCode id="ED42AE22FF89D90FFE860A60FE32FD2E" box="[368,399,632,655]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="169">VI</collectionCode>
. (
<figureCitation id="13682A62FF89D90FFE5E0A6FFDBDFD2E" box="[424,512,631,655]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2P</figureCitation>
<figureCitation id="13682A62FF89D90FFE5E0A6FFDBDFD2E" box="[424,512,631,655]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2P</figureCitation>
), em.
<date id="FFED1027FF89D90FFDB90A6FFF79FD13" pageId="6" pageNumber="169" value="2017-04-26">
<collectingDate id="EFA9E9CFFF89D90FFDB90A6FFF79FD13" pageId="6" pageNumber="169" value="2017-04-26">26. IV. 2017</collectingDate>
@ -140,11 +141,11 @@ on
<paragraph id="8BEC36E7FF89D90FFF510AA5FE45FC04" blockId="6.[140,670,527,1418]" pageId="6" pageNumber="169">
<emphasis id="B927EAF5FF89D90FFF510AA5FF5AFD75" box="[167,231,701,724]" italics="true" pageId="6" pageNumber="169">Larva</emphasis>
. Middle instar (
<figureCitation id="13682A62FF89D90FFE690AA5FE47FD74" box="[415,506,701,725]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2N</figureCitation>
<figureCitation id="13682A62FF89D90FFE690AA5FE47FD74" box="[415,506,701,725]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2N</figureCitation>
): head creamy white; trunk pale, covered with snow-white woolly wax. Late instar (
<figureCitation id="13682A62FF89D90FFE6F0B1AFE4DFCBB" box="[409,496,770,794]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2O</figureCitation>
<figureCitation id="13682A62FF89D90FFE6F0B1AFE4DFCBB" box="[409,496,770,794]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2O</figureCitation>
): head dark yellow; trunk pale greenish, covered with snow-white woolly wax; thoracic legs dark yellow. Mature larva (
<figureCitation id="13682A62FF89D90FFEDC0B72FEC2FC23" box="[298,383,874,898]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2P</figureCitation>
<figureCitation id="13682A62FF89D90FFEDC0B72FEC2FC23" box="[298,383,874,898]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2P</figureCitation>
): head dark yellow; trunk pale green, without wax covering.
</paragraph>
<paragraph id="8BEC36E7FF89D90FFF510BA8FD2CFBAC" blockId="6.[140,670,527,1418]" pageId="6" pageNumber="169">
@ -173,9 +174,9 @@ L. (new record).
<paragraph id="8BEC36E7FF89D90FFF510C0FFEC4FB5E" blockId="6.[140,670,527,1418]" pageId="6" pageNumber="169">
<emphasis id="B927EAF5FF89D90FFF510C0FFE99FB8E" box="[167,292,1047,1071]" italics="true" pageId="6" pageNumber="169">Life history</emphasis>
. This is a univoltine species with the adult emergence in April to May and the larval feeding stages in May to June in lowlands of central Honshu. The larva is a solitary external feeder of the host leaves (
<figureCitation id="13682A62FF89D90FFE5E0CBAFD9FFB1B" box="[424,546,1186,1210]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2N–O</figureCitation>
<figureCitation id="13682A62FF89D90FFE5E0CBAFD9FFB1B" box="[424,546,1186,1210]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2N–O</figureCitation>
). On maturity, the reared larva had an extra molt (
<figureCitation id="13682A62FF89D90FFDB20CDDFD28FB7C" box="[580,661,1221,1245]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="6" pageNumber="169">Fig. 2P</figureCitation>
<figureCitation id="13682A62FF89D90FFDB20CDDFD28FB7C" box="[580,661,1221,1245]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="6" pageNumber="169">Fig. 2P</figureCitation>
) and went into the soil.
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New Japanese name: Kunugi-habachi (
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<figureCitation id="13682A62FF8AD90CFEBF0EBAFE24F919" box="[329,409,1698,1720]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." figureDoi="http://doi.org/10.5281/zenodo.12760199" httpUri="https://zenodo.org/record/12760199/files/figure.png" pageId="5" pageNumber="168">Figs. 1D</figureCitation>
,
<figureCitation id="13682A62FF8AD90CFE520EBBFE5DF919" box="[420,480,1699,1720]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">2K–M</figureCitation>
<figureCitation id="13682A62FF8AD90CFE520EBBFE5DF919" box="[420,480,1699,1720]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">2K–M</figureCitation>
)
</paragraph>
</subSubSection>
@ -108,14 +109,14 @@ New Japanese name: Kunugi-habachi (
<paragraph id="8BEC36E7FF8AD90CFD3508CFFC17FEF6" blockId="5.[707,1237,212,1279]" pageId="5" pageNumber="168">
<materialsCitation id="3B3B3CBAFF8AD90CFD3508CFFC1BFEF6" collectingDate="2019-05-10" collectingDateMax="2019-10-18" collectingDateMin="2019-05-10" collectionCode="V" collectorName="Nakagawa &amp; S. Ibuki" latitude="36.766666" longLatPrecision="1185" longitude="140.13333" pageId="5" pageNumber="168" specimenCount="1">
1˂ (
<figureCitation id="13682A62FF8AD90CFCFC08CFFCD6FF4E" box="[778,875,215,239]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." pageId="5" pageNumber="168">Fig. 1D</figureCitation>
<figureCitation id="13682A62FF8AD90CFCFC08CFFCD6FF4E" box="[778,875,215,239]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." figureDoi="http://doi.org/10.5281/zenodo.12760199" httpUri="https://zenodo.org/record/12760199/files/figure.png" pageId="5" pageNumber="168">Fig. 1D</figureCitation>
), Koisago,
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<geoCoordinate id="EE675020FF8AD90CFB9D08CFFB6DFF4E" box="[1131,1232,214,239]" degrees="140" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="168" precision="925" value="140.13333">140°08′E</geoCoordinate>
,
<collectorName id="26A65331FF8AD90CFD3508E2FC8BFEB3" box="[707,822,250,274]" pageId="5" pageNumber="168">Nakagawa</collectorName>
, coll. larva (
<figureCitation id="13682A62FF8AD90CFC3008E2FB83FEB3" box="[966,1086,250,274]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2K–L</figureCitation>
<figureCitation id="13682A62FF8AD90CFC3008E2FB83FEB3" box="[966,1086,250,274]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2K–L</figureCitation>
),
<collectedFrom id="384F751AFF8AD90CFBA308E2FCADFE95" pageId="5" pageNumber="168">
on
@ -130,7 +131,7 @@ on
, mat. 17.
<collectionCode id="ED42AE22FF8AD90CFBE40905FB98FE95" box="[1042,1061,285,308]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="168" type="Museum">V</collectionCode>
. (
<figureCitation id="13682A62FF8AD90CFBCC0905FB2BFE94" box="[1082,1174,285,309]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2M</figureCitation>
<figureCitation id="13682A62FF8AD90CFBCC0905FB2BFE94" box="[1082,1174,285,309]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2M</figureCitation>
), em.
<date id="FFED1027FF8AD90CFD350927FCFEFEF6" box="[707,835,319,343]" pageId="5" pageNumber="168" value="2019-10-18">
<collectingDate id="EFA9E9CFFF8AD90CFD350927FCFEFEF6" box="[707,835,319,343]" pageId="5" pageNumber="168" value="2019-10-18">18. X. 2019</collectingDate>
@ -143,9 +144,9 @@ on
<paragraph id="8BEC36E7FF8AD90CFD2B097BFC5CFD2E" blockId="5.[707,1237,212,1279]" pageId="5" pageNumber="168">
<emphasis id="B927EAF5FF8AD90CFD2B097BFCA0FEDB" box="[733,797,355,378]" italics="true" pageId="5" pageNumber="168">Larva</emphasis>
. Late instar (
<figureCitation id="13682A62FF8AD90CFC31097AFBFAFEDB" box="[967,1095,354,378]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2K–L</figureCitation>
<figureCitation id="13682A62FF8AD90CFC31097AFBFAFEDB" box="[967,1095,354,378]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2K–L</figureCitation>
): head dark opaque gray; trunk and legs creamy white, dorsal region dark gray with creamy white line along middorsal line and each thoracic and abdominal segment with blackish spot in laterodorsal region. Mature larva (
<figureCitation id="13682A62FF8AD90CFC200A17FBFDFD86" box="[982,1088,527,551]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2M</figureCitation>
<figureCitation id="13682A62FF8AD90CFC200A17FBFDFD86" box="[982,1088,527,551]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2M</figureCitation>
): head pale orange; trunk and legs yellowish white, dorsal region dark gray, each segment with blackish spot in laterodorsal region.
</paragraph>
<paragraph id="8BEC36E7FF8AD90CFD2B0A83FCEEFD56" blockId="5.[707,1237,212,1279]" pageId="5" pageNumber="168">
@ -168,9 +169,9 @@ Murray
<paragraph id="8BEC36E7FF8AD90CFD2B0B1AFCA5FC4B" blockId="5.[707,1237,212,1279]" pageId="5" pageNumber="168">
<emphasis id="B927EAF5FF8AD90CFD2B0B1AFCE6FCBB" box="[733,859,770,794]" italics="true" pageId="5" pageNumber="168">Life history</emphasis>
. This is a univoltine species with the adult emergence in autumn, hibernation in the egg stage, and the larval feeding stages in spring. The larva is a solitary external feeder of the host leaves (
<figureCitation id="13682A62FF8AD90CFC860B95FC75FC04" box="[880,968,909,933]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2 L</figureCitation>
<figureCitation id="13682A62FF8AD90CFC860B95FC75FC04" box="[880,968,909,933]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2 L</figureCitation>
). On maturity, the reared larva had an extra molt (
<figureCitation id="13682A62FF8AD90CFC250BB7FB90FC66" box="[979,1069,943,967]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2M</figureCitation>
<figureCitation id="13682A62FF8AD90CFC250BB7FB90FC66" box="[979,1069,943,967]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2M</figureCitation>
) and went into the soil.
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(
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<figureCitation id="13682A62FF8CD90AFE840A2AFE0BFDE9" box="[370,438,562,584]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." figureDoi="http://doi.org/10.5281/zenodo.12760199" httpUri="https://zenodo.org/record/12760199/files/figure.png" pageId="3" pageNumber="166">Fig. 1C</figureCitation>
)
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@ -228,7 +229,7 @@ of dead trees of the above two newly recorded species standing on the broad rive
), Moriya City,
<collectingRegion id="4997F805FF8CD90AFEAE0CFFFDA4FB5E" box="[344,537,1255,1279]" country="Japan" name="Ibaraki" pageId="3" pageNumber="166">Ibaraki Prefecture</collectingRegion>
. Apparently, those females (one of them in
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<figureCitation id="13682A62FF8CD90AFE230D12FD9BFA83" box="[469,550,1290,1314]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." figureDoi="http://doi.org/10.5281/zenodo.12760199" httpUri="https://zenodo.org/record/12760199/files/figure.png" pageId="3" pageNumber="166">Fig. 1C</figureCitation>
) died after oviposition as often observed for siricid females.
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<figureCitation id="13682A62FF8CD90AFE930EB2FE79F961" box="[357,452,1706,1728]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2A–F</figureCitation>
<figureCitation id="13682A62FF8CD90AFE930EB2FE79F961" box="[357,452,1706,1728]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2A–F</figureCitation>
)
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, S. Ibuki; 1˂,
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same locality, coll. larva (
<figureCitation id="13682A62FF8CD90AFCC00927FC3BFEF6" box="[822,902,319,343]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2C</figureCitation>
<figureCitation id="13682A62FF8CD90AFCC00927FC3BFEF6" box="[822,902,319,343]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2C</figureCitation>
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on
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˂,
<materialsCitation id="3B3B3CBAFF8CD90AFD35099DFC2DFE40" collectingDate="2016-05-28" collectingDateMax="2017-04-24" collectingDateMin="2016-05-28" collectionCode="VI" collectorName="S. Ibuki" pageId="3" pageNumber="166" specimenCount="1">
same locality, coll. larva (
<figureCitation id="13682A62FF8CD90AFC26099DFBDDFE3C" box="[976,1120,389,413]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2B, D, F</figureCitation>
<figureCitation id="13682A62FF8CD90AFC26099DFBDDFE3C" box="[976,1120,389,413]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2B, D, F</figureCitation>
),
<collectedFrom id="384F751AFF8CD90AFB82099DFCFEFE1E" pageId="3" pageNumber="166">
on
@ -169,7 +170,7 @@ on
,
<materialsCitation id="3B3B3CBAFF8CD90AFC6809D2FBA3FD86" collectingDate="2018-05-31" collectingDateMax="2019-04-21" collectingDateMin="2018-05-31" collectionCode="VI" collectorName="S. Ibuki" pageId="3" pageNumber="166" specimenCount="1">
same locality, coll. larva (
<figureCitation id="13682A62FF8CD90AFB5D09D2FD58FDA4" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2A</figureCitation>
<figureCitation id="13682A62FF8CD90AFB5D09D2FD58FDA4" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2A</figureCitation>
),
<collectedFrom id="384F751AFF8CD90AFD0F09F5FBAEFDA4" box="[761,1043,493,517]" pageId="3" pageNumber="166">
on
@ -195,17 +196,17 @@ on
<paragraph id="8BEC36E7FF8CD90AFD2B0A2BFB03FC23" blockId="3.[707,1237,214,1557]" pageId="3" pageNumber="166">
<emphasis id="B927EAF5FF8CD90AFD2B0A2BFCA6FDEB" box="[733,795,563,586]" italics="true" pageId="3" pageNumber="166">Larva</emphasis>
. Early or middle instar (
<figureCitation id="13682A62FF8CD90AFBEA0A2AFB2CFDEB" box="[1052,1169,562,586]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2A–B</figureCitation>
<figureCitation id="13682A62FF8CD90AFBEA0A2AFB2CFDEB" box="[1052,1169,562,586]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2A–B</figureCitation>
): head pale grayish brown (
<figureCitation id="13682A62FF8CD90AFC430A4DFBACFDCC" box="[949,1041,597,621]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2A</figureCitation>
<figureCitation id="13682A62FF8CD90AFC430A4DFBACFDCC" box="[949,1041,597,621]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2A</figureCitation>
, probably earlier instar) or gray (
<figureCitation id="13682A62FF8CD90AFC8C0A6FFC6FFD2E" box="[890,978,631,655]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2B</figureCitation>
<figureCitation id="13682A62FF8CD90AFC8C0A6FFC6FFD2E" box="[890,978,631,655]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2B</figureCitation>
, probably later instar); trunk and legs creamy white. Late instar (
<figureCitation id="13682A62FF8CD90AFB5C0A82FCB9FD74" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2C–E</figureCitation>
<figureCitation id="13682A62FF8CD90AFB5C0A82FCB9FD74" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2C–E</figureCitation>
): head blackish; trunk and legs creamy white, most of dorsal region brown with paired whitish lines dorsally, and supraspiracular,
<typeStatus id="54E88845FF8CD90AFB5D0B1AFB69FCB8" box="[1195,1236,770,794]" pageId="3" pageNumber="166" type="epitype">epi-</typeStatus>
pleural and hypopleural regions of each segment and two or three terminal segments blackish. Mature larva (
<figureCitation id="13682A62FF8CD90AFCA10B72FC1AFC23" box="[855,935,874,898]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="3" pageNumber="166">Fig. 2F</figureCitation>
<figureCitation id="13682A62FF8CD90AFCA10B72FC1AFC23" box="[855,935,874,898]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2F</figureCitation>
): entirely vivid pale green.
</paragraph>
<paragraph id="8BEC36E7FF8CD90AFD2B0B95FC5CFBF3" blockId="3.[707,1237,214,1557]" pageId="3" pageNumber="166">

127
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@ -1,84 +1,85 @@
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<figureCitation id="13682A62FF8CD90AFC680EBAFC44F919" box="[926,1017,1698,1720]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="3" pageNumber="166">Fig. 2G–J</figureCitation>
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Fig. 2. A–F,
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<figureCitation id="13682A62FF8AD90CFF2308E2FE93FEB3" box="[213,302,250,274]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2G</figureCitation>
<figureCitation id="13682A62FF8AD90CFF2308E2FE93FEB3" box="[213,302,250,274]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2G</figureCitation>
), on
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<emphasis id="B927EAF5FF8AD90CFF51099DFF59FE3D" box="[167,228,389,412]" italics="true" pageId="5" pageNumber="168">Larva</emphasis>
. Early or middle instar (
<figureCitation id="13682A62FF8AD90CFE0A099DFDE9FE3C" box="[508,596,389,413]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2G</figureCitation>
<figureCitation id="13682A62FF8AD90CFE0A099DFDE9FE3C" box="[508,596,389,413]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2G</figureCitation>
): head pale gray; trunk and legs creamy white. Late instar (
<figureCitation id="13682A62FF8AD90CFF6509D2FEB8FE43" box="[147,261,458,482]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2H, J</figureCitation>
<figureCitation id="13682A62FF8AD90CFF6509D2FEB8FE43" box="[147,261,458,482]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2H, J</figureCitation>
): head pale pink; trunk and legs pale creamy white, with dorsal region pale greenish gray; thoracic legs tinted with pink. Mature larva (
<figureCitation id="13682A62FF8AD90CFF650A2AFF66FDEB" box="[147,219,562,586]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2I</figureCitation>
<figureCitation id="13682A62FF8AD90CFF650A2AFF66FDEB" box="[147,219,562,586]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2I</figureCitation>
): similar to late instar; head more yellowish.
</paragraph>
<paragraph id="8BEC36E7FF8AD90CFF510A4DFEC9FD74" blockId="5.[140,670,212,1037]" pageId="5" pageNumber="168">
@ -209,7 +210,7 @@ L. (new record).
<paragraph id="8BEC36E7FF8AD90CFF510AC7FD34FC23" blockId="5.[140,670,212,1037]" pageId="5" pageNumber="168">
<emphasis id="B927EAF5FF8AD90CFF510AC7FE9CFD56" box="[167,289,735,759]" italics="true" pageId="5" pageNumber="168">Life history</emphasis>
. This is a multivoltine species. The larva is solitary, though a few larvae may occur on the same leaf (
<figureCitation id="13682A62FF8AD90CFEBF0B3DFE22FC9C" box="[329,415,805,829]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." pageId="5" pageNumber="168">Fig. 2G</figureCitation>
<figureCitation id="13682A62FF8AD90CFEBF0B3DFE22FC9C" box="[329,415,805,829]" captionStart="Fig" captionStartId="4.[163,196,1617,1639]" captionTargetBox="[159,1216,215,1582]" captionTargetId="figure-12@4.[157,1218,213,1584]" captionTargetPageId="4" captionText="Fig. 2. A–F, Aglaostigma albicinctum: A, early or middle instar larva on Symplocarpus renifolius, 31. V. 2018; B, early or middle instar larva on Impatiens textorii, 29. V. 2016; C, late instar larva on Actinidia polygama, 25. VI. 2016; D, late instar larva on I. textorii, 30. V. 2016; E, late instar larva on A. polygama, 1. VII. 2016; F, mature larva, 10. VI. 2016. G–J, Allantus meridionalis: G, early instar, 20. IX. 2015; H, J, middle or late instar larva, 12. X. 2015; I, mature larva, 13. X. 2015. K–M, Apethymus kunugi: K, probably late instar larva, 10. V. 2019; L, same larva, 12. V. 2019; M, same larva, matured, and cast skin above, 17. V. 2019. N–P, Corymbas nipponica: N, middle instar larva, 17. VI. 2016; O, same larva, late instar, 26. VI. 2016; P, same larva, matured, and cast skin above, 29. VI. 2016.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760203" httpUri="https://zenodo.org/record/12760203/files/figure.png" pageId="5" pageNumber="168">Fig. 2G</figureCitation>
). The larva has an extra molt on maturation and enters a rotten wood, not soil, to make a cell for diapause (
<bibRefCitation id="EFC24B16FF8AD90CFE1A0B72FDC0FC23" author="Kondo, T." box="[492,637,874,898]" pageId="5" pageNumber="168" pagination="14 - 15" refId="ref14627" refString="Kondo, T. 1969. [Notes on the life history of Allantus meridionalis.] Suzumushi, 18: 14 - 15 (in Japanese)." type="book chapter" year="1969">Kondo, 1969</bibRefCitation>
).

117
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described this species from Etorofu Island, southern Kuriles, and it was later recorded from Moskva, Jakutsk and southern Primorye,
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and Kunashiri Island, southern Kuriles (see references above). The specimen listed above represents the first distribution record from the main island of Hokkaido. This is a mainly black species, with characteristic yellow marks on the lower inner orbit, mandible, and each side of the abdominal terga 3 and 4 (
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<figureCitation id="13682A62FF8ED908FC5E0C2AFC53FBE9" box="[936,1006,1074,1096]" captionStart="Fig" captionStartId="2.[163,196,1201,1223]" captionTargetBox="[158,1212,215,1166]" captionTargetId="figure-170@2.[157,1218,213,1168]" captionTargetPageId="2" captionText="Fig. 1. A, Late instar larva of Sterictiphora nipponica, 25. V. 2015, photographed by Ibuki. B, Phylloecus etorofensis, female, photographed by Hara. C, Dead female of Tremex apicalis on trunk of Alnus japonica, 23. V. 2021, photographed by Shinohara. D, Apethymus kunugi, female, photographed by Hara." figureDoi="http://doi.org/10.5281/zenodo.12760199" httpUri="https://zenodo.org/record/12760199/files/figure.png" pageId="1" pageNumber="164">Fig. 1A</figureCitation>
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. Late instar larva pale green (
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); setae inconspicuous; spiracles dark green; thoracic legs apically brownish.
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<figureCitation id="13682A62FF9AD91CFC5F0A10FC50FDBC" box="[937,1005,520,542]" captionStart="Fig" captionStartId="18.[163,196,998,1020]" captionTargetBox="[157,1218,213,964]" captionTargetId="figure-270@18.[157,1218,213,964]" captionTargetPageId="18" captionText="Fig. 8. A–C, Late instar larvae: A, Masaakia longivaginata, 21. V. 2019; B, Monophadnoides montanus, 18. VI. 2019; C, Strongylogaster moiwana, 30. V. 2017.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760224" httpUri="https://zenodo.org/record/12760224/files/figure.png" pageId="21" pageNumber="184">Fig. 8C</figureCitation>
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<emphasis id="B927EAF5FF9AD91CFD2B0AF8FCA0FD56" box="[733,797,736,759]" italics="true" pageId="21" pageNumber="184">Larva</emphasis>
. Late instar larva (
<figureCitation id="13682A62FF9AD91CFC0F0AC7FBEFFD56" box="[1017,1106,735,759]" captionStart="Fig" captionStartId="18.[163,196,998,1020]" captionTargetBox="[157,1218,213,964]" captionTargetId="figure-270@18.[157,1218,213,964]" captionTargetPageId="18" captionText="Fig. 8. A–C, Late instar larvae: A, Masaakia longivaginata, 21. V. 2019; B, Monophadnoides montanus, 18. VI. 2019; C, Strongylogaster moiwana, 30. V. 2017.All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 8C</figureCitation>
<figureCitation id="13682A62FF9AD91CFC0F0AC7FBEFFD56" box="[1017,1106,735,759]" captionStart="Fig" captionStartId="18.[163,196,998,1020]" captionTargetBox="[157,1218,213,964]" captionTargetId="figure-270@18.[157,1218,213,964]" captionTargetPageId="18" captionText="Fig. 8. A–C, Late instar larvae: A, Masaakia longivaginata, 21. V. 2019; B, Monophadnoides montanus, 18. VI. 2019; C, Strongylogaster moiwana, 30. V. 2017.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760224" httpUri="https://zenodo.org/record/12760224/files/figure.png" pageId="21" pageNumber="184">Fig. 8C</figureCitation>
): head pale brown, black on vertex and lateral part; trunk and legs pale yellowish green.
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<figureCitation id="13682A62FF9AD91CFE9E093DFE7CFE9B" box="[360,449,293,315]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9H–I</figureCitation>
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, mat. 9. VI. (
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<figureCitation id="13682A62FF9AD91CFEF00A2AFEF1FDEB" box="[262,332,562,586]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9I</figureCitation>
), em.
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, S. Ibuki.
@ -137,9 +138,9 @@ or
<paragraph id="8BEC36E7FF9AD91CFF510A4DFDDDFC9C" blockId="21.[140,670,354,1453]" pageId="21" pageNumber="184">
<emphasis id="B927EAF5FF9AD91CFF510A4DFF5AFDCD" box="[167,231,597,620]" italics="true" pageId="21" pageNumber="184">Larva</emphasis>
. Late instar (
<figureCitation id="13682A62FF9AD91CFE7D0A4DFE57FDCC" box="[395,490,597,621]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9H</figureCitation>
<figureCitation id="13682A62FF9AD91CFE7D0A4DFE57FDCC" box="[395,490,597,621]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9H</figureCitation>
): head purplish black; trunk and legs creamy white, dorsal region dark gray, sparsely covered with irregular black spots, each segment with paired yellowish dorsolateral thorns, and black line along ventral margin of pleural region. Mature larva (
<figureCitation id="13682A62FF9AD91CFDCA0B1AFD2DFCBB" box="[572,656,770,794]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9I</figureCitation>
<figureCitation id="13682A62FF9AD91CFDCA0B1AFD2DFCBB" box="[572,656,770,794]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9I</figureCitation>
): blackish, shiny; thoracic legs creamy white.
</paragraph>
<paragraph id="8BEC36E7FF9AD91CFF510B50FE6CFC04" blockId="21.[140,670,354,1453]" pageId="21" pageNumber="184">
@ -184,9 +185,9 @@ seems to be a polyphagous species with long larval feeding period. The larva of
<emphasis id="B927EAF5FF9AD91CFDBE0CF0FD20FB5E" box="[584,669,1255,1279]" italics="true" pageId="21" pageNumber="184">S. ferox</emphasis>
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by the dark color pattern of the head and dorsal region of the trunk in the feeding stages (compare A–C, E–F with H in
<figureCitation id="13682A62FF9AD91CFE500D57FE5AFAC6" box="[422,487,1359,1383]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9</figureCitation>
<figureCitation id="13682A62FF9AD91CFE500D57FE5AFAC6" box="[422,487,1359,1383]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9</figureCitation>
) and the creamy white thoracic legs of the mature larva (compare D, G with I in
<figureCitation id="13682A62FF9AD91CFED10D8DFED5FA0C" box="[295,360,1429,1453]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9</figureCitation>
<figureCitation id="13682A62FF9AD91CFED10D8DFED5FA0C" box="[295,360,1429,1453]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>

129
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@ -95,7 +96,7 @@ Togashi, 2009
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<paragraph id="8BEC36E7FF9AD91CFC630D72FBBCFA21" blockId="21.[777,1166,1350,1408]" box="[917,1025,1386,1408]" pageId="21" pageNumber="184">
(
<figureCitation id="13682A62FF9AD91CFC6A0D72FC46FA21" box="[924,1019,1386,1408]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9J–M</figureCitation>
<figureCitation id="13682A62FF9AD91CFC6A0D72FC46FA21" box="[924,1019,1386,1408]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9J–M</figureCitation>
)
</paragraph>
<paragraph id="8BEC36E7FF9AD91CFD2B0DAFFB69F945" blockId="21.[707,1237,1463,1799]" pageId="21" pageNumber="184">
@ -127,13 +128,13 @@ var.
,
<date id="FFED1027FF9AD91CFCD90E9FFC04F93E" box="[815,953,1671,1695]" pageId="21" pageNumber="184" value="2019-09-26">26. IX. 2019</date>
(
<figureCitation id="13682A62FF9AD91CFC3C0E9FFBA9F93E" box="[970,1044,1671,1695]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9J</figureCitation>
<figureCitation id="13682A62FF9AD91CFC3C0E9FFBA9F93E" box="[970,1044,1671,1695]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9J</figureCitation>
[larger larva], K), mat. 30. IX. (
<figureCitation id="13682A62FF9AD91CFC950EB2FC03F963" box="[867,958,1706,1730]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9 L</figureCitation>
<figureCitation id="13682A62FF9AD91CFC950EB2FC03F963" box="[867,958,1706,1730]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9 L</figureCitation>
), em.
<date id="FFED1027FF9AD91CFBFD0EB2FB29F963" box="[1035,1172,1706,1730]" pageId="21" pageNumber="184" value="2019-10-26">26. X. 2019</date>
(
<figureCitation id="13682A62FF9AD91CFB5F0EB2FD56F944" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="21" pageNumber="184">Fig. 9M</figureCitation>
<figureCitation id="13682A62FF9AD91CFB5F0EB2FD56F944" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="21" pageNumber="184">Fig. 9M</figureCitation>
), S. Ibuki; 1 ˂, same locality, coll. larva, on
</paragraph>
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@ -145,7 +146,7 @@ var.
,
<date id="FFED1027FF9AD91CFBCB0EF7FB69F8A6" box="[1085,1236,1775,1799]" pageId="21" pageNumber="184" value="2019-09-26">26. IX. 2019</date>
(
<figureCitation id="13682A62FF99D91FFF6208CFFF5CFF4E" box="[148,225,215,239]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="22" pageNumber="185">Fig. 9J</figureCitation>
<figureCitation id="13682A62FF99D91FFF6208CFFF5CFF4E" box="[148,225,215,239]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="22" pageNumber="185">Fig. 9J</figureCitation>
[smaller larva]), mat. 11. X., em.
<date id="FFED1027FF99D91FFD9708CFFF79FEB3" pageId="22" pageNumber="185" value="2020-01-23">23. I. 2020</date>
, S. Ibuki.
@ -153,11 +154,11 @@ var.
<paragraph id="8BEC36E7FF99D91FFF510905FE3DFDEB" blockId="22.[140,670,215,1799]" pageId="22" pageNumber="185">
<emphasis id="B927EAF5FF99D91FFF510905FF5AFE95" box="[167,231,285,308]" italics="true" pageId="22" pageNumber="185">Larva</emphasis>
. Middle instar (
<figureCitation id="13682A62FF99D91FFE680905FE56FE94" box="[414,491,285,309]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="22" pageNumber="185">Fig. 9J</figureCitation>
<figureCitation id="13682A62FF99D91FFE680905FE56FE94" box="[414,491,285,309]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="22" pageNumber="185">Fig. 9J</figureCitation>
[smaller larva]): head brownish white, with large median spot and smaller lateral spot on vertex black; trunk and legs creamy white, dorsal region pale greenish, each segment with small black spot laterally. Late instar (
<figureCitation id="13682A62FF99D91FFEE309D2FEDCFE43" box="[277,353,458,482]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="22" pageNumber="185">Fig. 9J</figureCitation>
<figureCitation id="13682A62FF99D91FFEE309D2FEDCFE43" box="[277,353,458,482]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="22" pageNumber="185">Fig. 9J</figureCitation>
[larger larva], K): similar to middle instar. Mature larva (
<figureCitation id="13682A62FF99D91FFE3009F5FDA2FDA4" box="[454,543,493,517]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="22" pageNumber="185">Fig. 9 L</figureCitation>
<figureCitation id="13682A62FF99D91FFE3009F5FDA2FDA4" box="[454,543,493,517]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="22" pageNumber="185">Fig. 9 L</figureCitation>
): yellowish white, shiny, with small blackish lateral spots on each segment of trunk.
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<figureCitation id="13682A62FF9CD91AFE950DCAFE7BFA49" box="[355,454,1490,1512]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9A–G</figureCitation>
<figureCitation id="13682A62FF9CD91AFE950DCAFE7BFA49" box="[355,454,1490,1512]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9A–G</figureCitation>
)
</paragraph>
<paragraph id="8BEC36E7FF9CD91AFF510E07FBA8FE94" blockId="19.[140,670,1567,1799]" lastBlockId="19.[707,1237,212,1661]" pageId="19" pageNumber="182">
@ -111,7 +112,7 @@
, mat. 18. VII., em.
<date id="FFED1027FF9CD91AFD7F0E7DFF59F93E" pageId="19" pageNumber="182" value="2017-05-02">2. V. 2017</date>
, S. Ibuki; 1 ˁ, Nakagawa, Bicchuzawa, coll. larva (
<figureCitation id="13682A62FF9CD91AFEFE0EB2FEE3F963" box="[264,350,1706,1730]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9A</figureCitation>
<figureCitation id="13682A62FF9CD91AFEFE0EB2FEE3F963" box="[264,350,1706,1730]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9A</figureCitation>
), on
<taxonomicName id="4C534D64FF9CD91AFE600EB2FDD0F963" box="[406,621,1706,1730]" class="Magnoliopsida" family="Betulaceae" genus="Actinidea" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="182" phylum="Tracheophyta" rank="species" species="polygama">
<emphasis id="B927EAF5FF9CD91AFE600EB2FDD0F963" box="[406,621,1706,1730]" italics="true" pageId="19" pageNumber="182">Actinidea polygama</emphasis>
@ -125,7 +126,7 @@ subsp.
<emphasis id="B927EAF5FF9CD91AFF2C0EF7FE96F8A6" box="[218,299,1775,1799]" italics="true" pageId="19" pageNumber="182">grandis</emphasis>
</taxonomicName>
27. VI., mat. 9. VIII. (
<figureCitation id="13682A62FF9CD91AFDC10EF7FD2CF8A6" box="[567,657,1775,1799]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9D</figureCitation>
<figureCitation id="13682A62FF9CD91AFDC10EF7FD2CF8A6" box="[567,657,1775,1799]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9D</figureCitation>
), em.
<date id="FFED1027FF9CD91AFD0508CFFCDAFF4E" box="[755,871,215,239]" pageId="19" pageNumber="182" value="2017-05-06">6. V. 2017</date>
, S. Ibuki; 1 ˂, Nasukarasuyama, coll. larva, on
@ -141,9 +142,9 @@ subsp.
<paragraph id="8BEC36E7FF9CD91AFD2B0958FCD5FDEB" blockId="19.[707,1237,212,1661]" pageId="19" pageNumber="182">
<emphasis id="B927EAF5FF9CD91AFD2B0958FCA0FEF6" box="[733,797,320,343]" italics="true" pageId="19" pageNumber="182">Larva</emphasis>
. Middle or late instar (
<figureCitation id="13682A62FF9CD91AFBE00927FB75FEF6" box="[1046,1224,319,343]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9A–C, E–F</figureCitation>
<figureCitation id="13682A62FF9CD91AFBE00927FB75FEF6" box="[1046,1224,319,343]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9A–C, E–F</figureCitation>
): head dark purplish gray to pale yellowish gray; trunk and legs creamy white, dorsal region pale gray or pale yellowish gray, sparsely covered with minute black spots, each segment with paired yellowish dorsolateral thorns. Mature larva (
<figureCitation id="13682A62FF9CD91AFCFF0A17FC3EFD86" box="[777,899,527,551]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9D, G</figureCitation>
<figureCitation id="13682A62FF9CD91AFCFF0A17FC3EFD86" box="[777,899,527,551]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9D, G</figureCitation>
): blackish, shiny; thoracic legs dark yellowish.
</paragraph>
<paragraph id="8BEC36E7FF9CD91AFD2B0A4DFC1AFC23" blockId="19.[707,1237,212,1661]" pageId="19" pageNumber="182">
@ -235,7 +236,7 @@ subsp.
noted that the larval period of this species was about 40 days. However, one of our cases of rearing indicated that the period may be much longer. The larva collected on
<date id="FFED1027FF9CD91AFCA00DA0FC4BFA6E" box="[854,1014,1463,1487]" pageId="19" pageNumber="182" value="2016-06-22">June 22, 2016</date>
(
<figureCitation id="13682A62FF9CD91AFBFC0DAFFBD8FA6E" box="[1034,1125,1463,1487]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." pageId="19" pageNumber="182">Fig. 9A</figureCitation>
<figureCitation id="13682A62FF9CD91AFBFC0DAFFBD8FA6E" box="[1034,1125,1463,1487]" captionStart="Fig" captionStartId="20.[163,196,1523,1545]" captionTargetBox="[159,1216,215,1487]" captionTargetId="figure-12@20.[157,1218,213,1489]" captionTargetPageId="20" captionText="Fig. 9. A–G, Siobla ferox: A, probably middle instar larva on Glechoma hederacea subsp. grandis, 27. VI. 2016; B, same larva, probably late instar, 12. VII. 2016; C, same larva, 16. VII. 2016; D, same larva, matured, 9. VIII. 2016; E. probably middle instar larva on Actinidia polygama, 14. VI. 2016; F. probably middle instar lar- vae on A. polygama, 19. VI. 2016; G. same larva, 15. VII. 2016. H–I, Siobla japonica: H, last instar larva, 5. VI. 2019; I, mature larva and cast skin above, 9. VI. 2019; J–M, Taxonus shinoharai: J, middle and late instar larvae on Agrimonia pilosa var. japonica, 29. IX. 2019; K, late instar, 29. IX. 2019; L, same larva, matured, 30. IX. 2019; M, male adult, just emerged, 26. X. 2019. All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760226" httpUri="https://zenodo.org/record/12760226/files/figure.png" pageId="19" pageNumber="182">Fig. 9A</figureCitation>
, probably middle instar) and reared indoors matured on August 9 (Fig. D). The rearing period from middle instar to maturity in this case lasted 48 days. The full larval period of this individual is esti- mated at 60 days at least.
</paragraph>
</subSubSection>

119
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<figureCitation id="13682A62FF9DD91BFE840DCAFE0AFA49" box="[370,439,1490,1512]" captionStart="Fig" captionStartId="18.[163,196,998,1020]" captionTargetBox="[157,1218,213,964]" captionTargetId="figure-270@18.[157,1218,213,964]" captionTargetPageId="18" captionText="Fig. 8. A–C, Late instar larvae: A, Masaakia longivaginata, 21. V. 2019; B, Monophadnoides montanus, 18. VI. 2019; C, Strongylogaster moiwana, 30. V. 2017.All photographed by Ibuki." figureDoi="http://doi.org/10.5281/zenodo.12760224" httpUri="https://zenodo.org/record/12760224/files/figure.png" pageId="18" pageNumber="181">Fig. 8B</figureCitation>
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@ -126,7 +127,7 @@ VI. 2019, em.
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, coll. larva (
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@ -151,7 +152,7 @@ mat. 24.
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): head pale brownish green; trunk entirely pale green except for green middorsal stripe, with many branched spines; legs, spines and spiracles whitish. Mature larva:
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long, pale green, without spines.

119
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, coll. larva (
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. Late instar larva (
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) pale green; trunk with semitransparent branched spines; thoracic spines apically brownish. Mature larva
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long; head pale greenish yellow; trunk pale green, without spines; legs pale yellow green, with claws brownish.

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<figureCitation id="4FA254CCFFE1902DFEF8FD6BFE535B7D" box="[365,444,577,599]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="31" pageNumber="178">Fig. 11E</figureCitation>
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<figureCitation id="4FA254CCFFE1902DFEFEFB55FE245DBC" box="[363,459,1151,1175]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." pageId="31" pageNumber="178">Fig. 11E</figureCitation>
<figureCitation id="4FA254CCFFE1902DFEFEFB55FE245DBC" box="[363,459,1151,1175]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="31" pageNumber="178">Fig. 11E</figureCitation>
), the rostrum is triangular, with a wide basal part; both of two transverse carinae of the carapace dorsal surface are wholly distinct between the carapace lateral margins of both sides.
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(
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<figureCitation id="4FA254CCFFE3902FFC03FE2BFBEE583D" box="[918,1025,257,279]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14A–F</figureCitation>
)
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@ -96,17 +97,17 @@ KH-72-1 cruise, sta. 52, 1Ə (CB 10.1×CL
<paragraph id="D7264849FFE3902FFD48FE8DFB945AEC" blockId="29.[707,1237,319,1799]" pageId="29" pageNumber="176">
<emphasis id="E5ED945BFFE3902FFD48FE8DFCD55894" box="[733,826,423,447]" italics="true" pageId="29" pageNumber="176">Remarks</emphasis>
. The carapace (
<figureCitation id="4FA254CCFFE3902FFC6BFE8DFB6E5894" box="[1022,1153,423,447]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." pageId="29" pageNumber="176">Fig. 14A–B</figureCitation>
<figureCitation id="4FA254CCFFE3902FFC6BFE8DFB6E5894" box="[1022,1153,423,447]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14A–B</figureCitation>
) of the present specimen of
<taxonomicName id="109933CAFFE3902FFC38FEE0FB9258C9" authorityName="Alcock and Anderson" authorityYear="1899" box="[941,1149,458,482]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="176" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="E5ED945BFFE3902FFC38FEE0FB9258C9" box="[941,1149,458,482]" italics="true" pageId="29" pageNumber="176">Camatopsis rubida</emphasis>
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is subtrapezoidal and high, differing slightly in the pro- portional shape according to the photographing angles. The carapace dorsal surface, chelipeds and ambulatory legs are wholly covered with longish fine setae (
<figureCitation id="4FA254CCFFE3902FFC33FD5DFBC05BA4" box="[934,1071,631,655]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." pageId="29" pageNumber="176">Fig. 14A–F</figureCitation>
<figureCitation id="4FA254CCFFE3902FFC33FD5DFBC05BA4" box="[934,1071,631,655]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14A–F</figureCitation>
). In the male examined, the right cheliped (
<figureCitation id="4FA254CCFFE3902FFB92FDB0FB885B99" box="[1031,1127,666,690]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." pageId="29" pageNumber="176">Fig. 14E</figureCitation>
<figureCitation id="4FA254CCFFE3902FFB92FDB0FB885B99" box="[1031,1127,666,690]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14E</figureCitation>
) is larger, but slender, leaving a narrow gape between both fingers along their basal halves. In the small left cheliped (
<figureCitation id="4FA254CCFFE3902FFCA6FC28FC755A31" box="[819,922,770,794]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." pageId="29" pageNumber="176">Fig. 14F</figureCitation>
<figureCitation id="4FA254CCFFE3902FFCA6FC28FC755A31" box="[819,922,770,794]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14F</figureCitation>
), the palm and fingers are slenderer, being armed with small irregular teeth on both cutting edges and with a sharp spine at distal one third of immovable cutting edge. Both cheliped fingers are apparently different from the chelipeds of developing or mature males.
</paragraph>
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<figureCitation id="4FA254CCFFE3902FFEC8FD83FE235B95" box="[349,460,681,703]" captionStart="Fig" captionStartId="30.[163,196,1651,1673]" captionTargetBox="[158,1217,215,1617]" captionTargetId="figure-10@30.[157,1218,213,1618]" captionTargetPageId="30" captionText="Fig. 14. A–F: Camatopsis rubida Alcock and Anderson, Ə (NSMT-Cr 30942; CB 10.1×CL 8.9mm) from sta. 52. A–B: Dorsal views in different photographic angle (A–B), frontal view (C), pleon (B), right (E) and left (F) chelae. G–H: Camatopsis leptomerus Ng and Castro, 8 (NSMT-Cr 30941; CB 5.5×CL 4.8mm) from sta. 42. Dorsal views in different photographic angle (G–H)." figureDoi="http://doi.org/10.5281/zenodo.12760124" httpUri="https://zenodo.org/record/12760124/files/figure.png" pageId="29" pageNumber="176">Fig. 14G–H</figureCitation>
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is most characteristic in having the shortest and stoutest ambulatory meri among its congeners. The specimen at hand (
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@ -83,7 +84,7 @@ H.
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(
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<figureCitation id="4FA254CCFFE49028FC36FAC4FC1C5F2F" box="[931,1011,1518,1540]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="26" pageNumber="173">Fig. 11C</figureCitation>
)
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@ -97,7 +98,7 @@ KH-72-1 cruise, sta. 50, 1Ə (CB 7.5×CL
<paragraph id="D7264849FFE49029FD48F980FE435B2E" blockId="26.[707,1237,1602,1799]" lastBlockId="27.[140,669,215,621]" lastPageId="27" lastPageNumber="174" pageId="26" pageNumber="173">
<emphasis id="E5ED945BFFE49028FD48F980FCAD5FE9" box="[733,834,1706,1730]" italics="true" pageId="26" pageNumber="173">Remarks.</emphasis>
The male examined after ecdysis is wholly semitransparent due to decalcification (
<figureCitation id="4FA254CCFFE49028FD5EF9C5FCD85E2C" box="[715,823,1775,1799]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." pageId="26" pageNumber="173">Fig. 11C</figureCitation>
<figureCitation id="4FA254CCFFE49028FD5EF9C5FCD85E2C" box="[715,823,1775,1799]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="26" pageNumber="173">Fig. 11C</figureCitation>
), agreeing well with the detailed description, photographs and figures by
<bibRefCitation id="B30835B8FFE59029FDA8FFFDFF155839" author="Chia, D. G. B. &amp; P. K. L. Ng" pageId="27" pageNumber="174" pagination="15 - 56" refId="ref22895" refString="Chia, D. G. B. and P. K. L. Ng 2000. A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History 34: 15 - 56." type="journal article" year="2000">Chia and Ng (2000)</bibRefCitation>
who examined the type specimens and synonymized

103
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(
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<figureCitation id="4FA254CCFFE59029FECBFA34FE245C1F" box="[350,459,1310,1332]" captionStart="Fig" captionStartId="24.[163,196,1358,1380]" captionTargetBox="[158,1217,215,1324]" captionTargetId="figure-154@24.[157,1218,213,1325]" captionTargetPageId="24" captionText="Fig. 10. A–B: Pilumnus minutus (De Haan), Ə (NSMT-Cr 30937; CB 8.0×CL 6.1mm) from sta. 45. C–D: Actumnus forficigerus (Stimpson), Ə (NSMT-Cr 30929; CB 10.0mm×CL 7.0 mm) from sta. 45. E–F: Actum- nus squamosus (De Haan), 8 (NSMT-Cr 30932; CB 7.7×CL 5.5 mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760112" httpUri="https://zenodo.org/record/12760112/files/figure.png" pageId="27" pageNumber="174">Fig. 10A–B</figureCitation>
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@ -102,7 +103,7 @@ KH-72-1 cruise, sta. 45, 1 Ə (CB 8.0×CL
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<emphasis id="E5ED945BFFE59029FF33FAD7FEEC5F3E" box="[166,259,1533,1557]" italics="true" pageId="27" pageNumber="174">Remarks</emphasis>
. The male at hand (
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<figureCitation id="4FA254CCFFE59029FD98FAD7FD7A5F3E" box="[525,661,1533,1557]" captionStart="Fig" captionStartId="24.[163,196,1358,1380]" captionTargetBox="[158,1217,215,1324]" captionTargetId="figure-154@24.[157,1218,213,1325]" captionTargetPageId="24" captionText="Fig. 10. A–B: Pilumnus minutus (De Haan), Ə (NSMT-Cr 30937; CB 8.0×CL 6.1mm) from sta. 45. C–D: Actumnus forficigerus (Stimpson), Ə (NSMT-Cr 30929; CB 10.0mm×CL 7.0 mm) from sta. 45. E–F: Actum- nus squamosus (De Haan), 8 (NSMT-Cr 30932; CB 7.7×CL 5.5 mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760112" httpUri="https://zenodo.org/record/12760112/files/figure.png" pageId="27" pageNumber="174">Fig. 10A–B</figureCitation>
) agrees well with the specimens from the Sahul Shelf recorded by Takeda
<emphasis id="E5ED945BFFE59029FE23F969FE1E5F71" box="[438,497,1602,1626]" italics="true" pageId="27" pageNumber="174">et al.</emphasis>
(2002b) in the setation of the carapace, chelipeds and ambulatory legs with stiff setae of various sizes and the spination of the carapace anterolateral margins and chelipeds.

111
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(
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<figureCitation id="4FA254CCFFE59029FC3FFB63FC035D75" box="[938,1004,1097,1119]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." figureDoi="http://doi.org/10.5281/zenodo.12760120" httpUri="https://zenodo.org/record/12760120/files/figure.png" pageId="27" pageNumber="174">Fig. 13</figureCitation>
)
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@ -138,16 +139,16 @@ by
</paragraph>
<paragraph id="D7264849FFE5902EFD48F9E7FCF25E2C" blockId="27.[707,1237,1151,1799]" lastBlockId="28.[707,1237,1463,1799]" lastPageId="28" lastPageNumber="175" pageId="27" pageNumber="174">
The carapace dorsal surface (
<figureCitation id="4FA254CCFFE59029FBBCF9E7FB415FCE" box="[1065,1198,1741,1765]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." pageId="27" pageNumber="174">Fig. 13A–B</figureCitation>
<figureCitation id="4FA254CCFFE59029FBBCF9E7FB415FCE" box="[1065,1198,1741,1765]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." figureDoi="http://doi.org/10.5281/zenodo.12760120" httpUri="https://zenodo.org/record/12760120/files/figure.png" pageId="27" pageNumber="174">Fig. 13A–B</figureCitation>
) is smooth and glabrous without areolar indication, and evenly convex in both directions; the front prominently extends forward, with transverse margin and without median incision, slightly more than one-third as wide as carapace; the carapace anterolateral margin (
<figureCitation id="4FA254CCFFE2902EFE55F968FDAB5F71" box="[448,580,1602,1626]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." pageId="28" pageNumber="175">Fig. 13A–B</figureCitation>
<figureCitation id="4FA254CCFFE2902EFE55F968FDAB5F71" box="[448,580,1602,1626]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." figureDoi="http://doi.org/10.5281/zenodo.12760120" httpUri="https://zenodo.org/record/12760120/files/figure.png" pageId="28" pageNumber="175">Fig. 13A–B</figureCitation>
) is narrowly edged and cut into three lobes; first lobe is isolated from the less prominent external orbital angle with a short distance, and fringed with minute granules; the second lobe is small, but tooth-like and subacute at the tip, separated from the first lobe by a small but distinct notch; the third lobe obscurely isolated from the second lobe is obscurely convex and weakly convergent toward the carapace posterolateral corner. Both chelipeds (
<figureCitation id="4FA254CCFFE2902EFCACF968FC545F71" box="[825,955,1602,1626]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." pageId="28" pageNumber="175">Fig. 13D–E</figureCitation>
<figureCitation id="4FA254CCFFE2902EFCACF968FC545F71" box="[825,955,1602,1626]" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." figureDoi="http://doi.org/10.5281/zenodo.12760120" httpUri="https://zenodo.org/record/12760120/files/figure.png" pageId="28" pageNumber="175">Fig. 13D–E</figureCitation>
) are different in size and shape; in the right, or larger cheliped in this male, the palm is stout, fairly bulged, but the left cheliped palm is compressed, with sharp lower margin. The ambulatory legs are all slender (
<figureCitation id="4FA254CCFFE2902EFB3CF9E7FCFF5E2C" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." pageId="28" pageNumber="175">Fig. 13A, F</figureCitation>
<figureCitation id="4FA254CCFFE2902EFB3CF9E7FCFF5E2C" captionStart="Fig" captionStartId="28.[163,196,1321,1343]" captionTargetBox="[158,1217,215,1286]" captionTargetId="figure-162@28.[157,1218,213,1287]" captionTargetPageId="28" captionText="Fig. 13. Xestopilumnus cultripollex Ng and Dai, Ə (NSMT-Cr 30940; CB 5.6×CL 4.7 mm) from sta. 42. Dorsal (A), frontal (B) and ventral (C) views, right (larger) chela in dorsal (D) and outer (E) views, left (smaller) chela in outer view (F)." figureDoi="http://doi.org/10.5281/zenodo.12760120" httpUri="https://zenodo.org/record/12760120/files/figure.png" pageId="28" pageNumber="175">Fig. 13A, F</figureCitation>
).
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Fig. 13.
<taxonomicName id="109933CAFFE2902EFE90FA03FD855C14" authority="Ng and Dai" authorityName="Ng and Dai" authorityYear="1997" box="[261,618,1321,1343]" class="Malacostraca" family="Pilumnidae" genus="Xestopilumnus" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="175" phylum="Arthropoda" rank="species" species="cultripollex">

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(
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<figureCitation id="4FA254CCFFE7902BFC3FF9FBFC035FCC" box="[938,1004,1745,1767]" captionStart="Fig" captionStartId="26.[163,196,765,787]" captionTargetBox="[158,1217,214,731]" captionTargetId="figure-406@26.[157,1218,213,732]" captionTargetPageId="26" captionText="Fig. 12. Ceratoplax truncatifrons Rathbun, 8 (NSMT-Cr 30934; CB 5.3×CL 3.3mm) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760118" httpUri="https://zenodo.org/record/12760118/files/figure.png" pageId="25" pageNumber="172">Fig. 12</figureCitation>
)
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Fig. 12.
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@ -150,7 +151,7 @@ specimens of Rathbun`s two species. According to
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. It is impossible at present to apply the sex-associated characters with the female at hand, and the differences of the carapace shape may be subtle for the species identification as shown at the photographic angles (
<figureCitation id="4FA254CCFFE49028FECCF9ADFE0B5FB4" box="[345,484,1671,1695]" captionStart="Fig" captionStartId="26.[163,196,765,787]" captionTargetBox="[158,1217,214,731]" captionTargetId="figure-406@26.[157,1218,213,732]" captionTargetPageId="26" captionText="Fig. 12. Ceratoplax truncatifrons Rathbun, 8 (NSMT-Cr 30934; CB 5.3×CL 3.3mm) from sta. 42." pageId="26" pageNumber="173">Fig. 12A–B</figureCitation>
<figureCitation id="4FA254CCFFE49028FECCF9ADFE0B5FB4" box="[345,484,1671,1695]" captionStart="Fig" captionStartId="26.[163,196,765,787]" captionTargetBox="[158,1217,214,731]" captionTargetId="figure-406@26.[157,1218,213,732]" captionTargetPageId="26" captionText="Fig. 12. Ceratoplax truncatifrons Rathbun, 8 (NSMT-Cr 30934; CB 5.3×CL 3.3mm) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760118" httpUri="https://zenodo.org/record/12760118/files/figure.png" pageId="26" pageNumber="173">Fig. 12A–B</figureCitation>
). In the present female, however, the antero-external corner of the third maxilliped merus is sharply developed as in the figure given by
<bibRefCitation id="B30835B8FFE49028FE03F9DAFD805E2C" author="Ng, P. K. L. &amp; P. F. Clark" box="[406,623,1775,1800]" pageId="26" pageNumber="173" pagination="323 - 331" refId="ref25872" refString="Ng, P. K. L. and P. F. Clark 2015. Ceratoplax margarita n. sp., a new rhizopine crab (Crustacea: Brachyura: Pilumnidae) from Papua New Guinea, with rediagnoses of C. truncatifrons Rathbun, 1914, and C. fulgida Rathbun, 1914. Zoosystema 37: 323 - 331." type="journal article" year="2015">Ng and Clark (2015</bibRefCitation>
, fig. 4C).

101
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@ -83,7 +84,7 @@ Semper
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<figureCitation id="4FA254CCFFE89024FEFBFEABFE5458BD" box="[366,443,385,407]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="22" pageNumber="169">Fig. 11F</figureCitation>
)
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@ -101,7 +102,7 @@ including lateral teeth, CL 17.0 mm including frontal teeth), NSMT-Cr 30924.
<paragraph id="D7264849FFE89024FF32FD5DFD965AA9" blockId="22.[140,669,458,1037]" pageId="22" pageNumber="169">
<emphasis id="E5ED945BFFE89024FF32FD5DFEE45BA4" box="[167,267,631,655]" italics="true" pageId="22" pageNumber="169">Remarks.</emphasis>
The female at hand (
<figureCitation id="4FA254CCFFE89024FDBCFD5DFD7F5BA4" box="[553,656,631,655]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." pageId="22" pageNumber="169">Fig. 11F</figureCitation>
<figureCitation id="4FA254CCFFE89024FDBCFD5DFD7F5BA4" box="[553,656,631,655]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="22" pageNumber="169">Fig. 11F</figureCitation>
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<taxonomicName id="109933CAFFE89024FEA4FDB0FE755B99" box="[305,410,666,690]" class="Maxillopoda" family="Sacculinidae" genus="Sacculina" kingdom="Animalia" order="Kentrogonida" pageId="22" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="E5ED945BFFE89024FEA4FDB0FE755B99" box="[305,410,666,690]" italics="true" pageId="22" pageNumber="169">Sacculina</emphasis>

103
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@ -1,74 +1,75 @@
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@ -82,7 +83,7 @@ Alcock, 1899
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<figureCitation id="4FA254CCFFE89024FEE7FBFEFE595DC2" box="[370,438,1236,1258]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760087" httpUri="https://zenodo.org/record/12760087/files/figure.png" pageId="22" pageNumber="169">Fig. 1C</figureCitation>
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@ -96,7 +97,7 @@ KH-72-1 cruise, sta. 45, 1 8 (CB 11.0×CL
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. The female at hand (
<figureCitation id="4FA254CCFFE89024FD94FA58FDBD5CA1" box="[513,594,1394,1418]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." pageId="22" pageNumber="169">Fig. 1C</figureCitation>
<figureCitation id="4FA254CCFFE89024FD94FA58FDBD5CA1" box="[513,594,1394,1418]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760087" httpUri="https://zenodo.org/record/12760087/files/figure.png" pageId="22" pageNumber="169">Fig. 1C</figureCitation>
) is not fully mature, but is clearly identifiable with
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(
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, fig. 2a) and
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, fig. 196). In the present specimens (
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<figureCitation id="4FA254CCFFE89024FC5FFBCDFBD15DD4" box="[970,1086,1255,1279]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="22" pageNumber="169">Fig. 9E–F</figureCitation>
), the anterior margin of the outer lobe is shallowly concave at the subterminal part and shallowly separated from the supraorbital angle. In this case, the frontal margin is indistinctly six-lobed rather than four-lobed, being somewhat different from the figures given by
<bibRefCitation id="B30835B8FFE89024FCECFA9DFBC95CE4" author="Crosnier, A." box="[889,1062,1463,1487]" pageId="22" pageNumber="169" pagination="1 - 154" refId="ref23044" refString="Crosnier, A. 1962. Crustaces Decapodes Portunidae. Fauna Madagascar 16: 1 - 154, pls. 1 - 13." type="journal article" year="1962">Crosnier (1962)</bibRefCitation>
,
@ -114,7 +115,7 @@ characterized by small body size, the pilose carapace surface, the large orbital
. This condition of the frontal lobes approaching to the six-lobed appearance was already mentioned by
<bibRefCitation id="B30835B8FFE89024FBE0F935FC275F71" author="Stephenson, W. &amp; J. J. Hudson" pageId="22" pageNumber="169" pagination="312 - 368" refId="ref28568" refString="Stephenson, W. and J. J. Hudson 1957. The Australian portunids (Crustacea: Portunidae). I. The genus Thalamita. Australian Journal of Marine and Freshwater Research 8: 312 - 368, pls. 1 - 10." type="journal article" year="1957">Stephenson and Hudson (1957)</bibRefCitation>
. The carapace anterolateral margin is armed with five anteriorly-directed teeth including the external orbital tooth which is the largest of all; the second is slightly smaller than the first, but larger than the subequal third and fifth teeth; the fourth is apparently smaller than the others, but distinct in the specimens examined (
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<figureCitation id="4FA254CCFFE99025FE97FFD0FEBA5839" box="[258,341,250,274]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="23" pageNumber="170">Fig. 9E</figureCitation>
).
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101
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(
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<figureCitation id="4FA254CCFFE99025FC0DFEB4FC10589F" box="[920,1023,414,436]" captionStart="Fig" captionStartId="24.[163,196,1358,1380]" captionTargetBox="[158,1217,215,1324]" captionTargetId="figure-154@24.[157,1218,213,1325]" captionTargetPageId="24" captionText="Fig. 10. A–B: Pilumnus minutus (De Haan), Ə (NSMT-Cr 30937; CB 8.0×CL 6.1mm) from sta. 45. C–D: Actumnus forficigerus (Stimpson), Ə (NSMT-Cr 30929; CB 10.0mm×CL 7.0 mm) from sta. 45. E–F: Actum- nus squamosus (De Haan), 8 (NSMT-Cr 30932; CB 7.7×CL 5.5 mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760112" httpUri="https://zenodo.org/record/12760112/files/figure.png" pageId="23" pageNumber="170">Fig. 10E–F</figureCitation>
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clearly figured this feature with a photograph of both chelae in comparing the new species,
<emphasis id="E5ED945BFFE99025FB1BFC62FCEC5AA9" italics="true" pageId="23" pageNumber="170">A. tsu- rukaii</emphasis>
, with the related species. The present female (
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<figureCitation id="4FA254CCFFE99025FC8AFCA7FC705A8E" box="[799,927,909,933]" captionStart="Fig" captionStartId="24.[163,196,1358,1380]" captionTargetBox="[158,1217,215,1324]" captionTargetId="figure-154@24.[157,1218,213,1325]" captionTargetPageId="24" captionText="Fig. 10. A–B: Pilumnus minutus (De Haan), Ə (NSMT-Cr 30937; CB 8.0×CL 6.1mm) from sta. 45. C–D: Actumnus forficigerus (Stimpson), Ə (NSMT-Cr 30929; CB 10.0mm×CL 7.0 mm) from sta. 45. E–F: Actum- nus squamosus (De Haan), 8 (NSMT-Cr 30932; CB 7.7×CL 5.5 mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760112" httpUri="https://zenodo.org/record/12760112/files/figure.png" pageId="23" pageNumber="170">Fig. 10E–F</figureCitation>
) agrees with the ovigerous female from the Sahul Shelf in having the carapace covered with a thick felt of short setae and in not being strongly convex dorsally, and also in having the characteristic chelipeds reported by
<bibRefCitation id="B30835B8FFE99025FD56FB10FC445D79" author="Takeda, M. &amp; S. T. Ahyong &amp; N. Ohtsuchi &amp; H. Komatsu" box="[707,939,1082,1106]" pageId="23" pageNumber="170" pagination="35 - 83" refId="ref29099" refString="Takeda, M., S. T. Ahyong, N. Ohtsuchi and H. Komatsu 2022 b. Crabs (Crustacea, Decapoda) from the seas of East and Southeast Asia collected by the RV Hakuho Maru (KH- 72 - 1 cruise). 3. Sahul Shelf. Bulletin of the National Museum of Nature and Science, Tokyo (A) 49: 35 - 83." type="journal article" year="2022">
Takeda

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(
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in having the usual tubercles on the outer surface of the cheliped palm instead of the thick pavement of the tubercles. In the smaller male and all of the females, both chelipeds are stout and covered with some rows of conical granules on the palm outer surfaces, with short and stout fingers, but in the larger male (
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105
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(
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<figureCitation id="4FA254CCFFE99025FC00FAE6FBED5CCA" box="[917,1026,1484,1506]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="23" pageNumber="170">Fig. 11A–B</figureCitation>
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@ -122,10 +123,10 @@ The female specimen examined agrees well with the descriptions and figures given
<emphasis id="E5ED945BFFE6902AFF19FAD7FF1E5F3E" box="[140,241,1533,1557]" italics="true" pageId="24" pageNumber="171">Pilumnus</emphasis>
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). The carapace is robust in appearance, vaulted in both directions, with the ill-defined surface covered with stiff longish setae, the chelipeds are stout, and different in size but similar in the armature, with some rows of equidistant long, curved spiniform tubercles on both carpi and palms (
<figureCitation id="4FA254CCFFE6902AFE92F9E7FE665FCE" box="[263,393,1741,1765]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." pageId="24" pageNumber="171">Fig. 11A–B</figureCitation>
<figureCitation id="4FA254CCFFE6902AFE92F9E7FE665FCE" box="[263,393,1741,1765]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="24" pageNumber="171">Fig. 11A–B</figureCitation>
).
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Fig. 10. A–B:
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103
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(
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<figureCitation id="4FA254CCFFEA9026FC3DFB0CFC005D17" box="[936,1007,1062,1084]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760087" httpUri="https://zenodo.org/record/12760087/files/figure.png" pageId="20" pageNumber="167">Fig. 1D</figureCitation>
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is characterised by having many transverse striae on the carapace dorsal surface (
<figureCitation id="4FA254CCFFEA9026FC26FA20FBE15C09" box="[947,1038,1290,1314]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." pageId="20" pageNumber="167">Fig. 1D</figureCitation>
<figureCitation id="4FA254CCFFEA9026FC26FA20FBE15C09" box="[947,1038,1290,1314]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760087" httpUri="https://zenodo.org/record/12760087/files/figure.png" pageId="20" pageNumber="167">Fig. 1D</figureCitation>
). The records of occurrence are few, but the characteristics are sufficiently illustrated by
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,

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(
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<figureCitation id="4FA254CCFFEA9026FEF9FC54FE525ABF" box="[364,445,894,916]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="20" pageNumber="167">Fig. 11D</figureCitation>
)
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In this specimen (
<figureCitation id="4FA254CCFFEA9026FEF8FAF0FE3E5CD9" box="[365,465,1498,1522]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." pageId="20" pageNumber="167">Fig. 11D</figureCitation>
<figureCitation id="4FA254CCFFEA9026FEF8FAF0FE3E5CD9" box="[365,465,1498,1522]" captionStart="Fig" captionStartId="25.[163,196,1415,1437]" captionTargetBox="[158,1217,215,1381]" captionTargetId="figure-62@25.[157,1218,213,1382]" captionTargetPageId="25" captionText="Fig. 11. A–B: Bathypilumnus sinensis (Gordon), 8 (NSMT-Cr 30933; CB 8.7×CL 7.0mm) from sta. 45. C: Eumedonus niger H. Milne Edwards, Ə (NSMT-Cr 30935; CB 7.5×CL 7.7mm) from sta. 50. D: Incultus tuberculosus (A. Milne-Edwards), 8 (NSMT-Cr 30928; CB 25.3mm with epibranchial tubercles, CL 14.5mm) from sta. 45. E: Retropluma denticulata Rathbun, Ə (NSMT-Cr 30943; CB 7.8×CL 6.6 mm) from sta. 42. F: Lupocyclus philippinensis Semper, Ə infested by a Sacculina (NSMT-Cr 30924; CB 20.5mm including lateral teeth, CL 17.0mm including frontal teeth) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760116" httpUri="https://zenodo.org/record/12760116/files/figure.png" pageId="20" pageNumber="167">Fig. 11D</figureCitation>
), the lateral tubercles of the carapace posterior margin are stout and directed obliquely outward. This species is characteristic in the carapace ornamentation and armature: the dorsal surface is flattened overall, areolated with a distinct gastric region, a strong ridge running from the epibranchial tubercle toward the gastric region, and several obtuse tubercles arranged symmetrically around the cardiac region. The carapace anterolateral margin is armed with nine teeth including the obtuse external orbital angle and the strong epibranchial tubercle as usual, but the first, second, third and fifth teeth are distinctly smaller than the fourth, sixth and seventh teeth.
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(
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@ -163,9 +164,9 @@ and the G1 morphology. The shape of the front lobes seems to be similar in both
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The following account of the carapace is based on the present male examined (
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<figureCitation id="4FA254CCFFEB9027FB87FDB0FB875B99" box="[1042,1128,666,690]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="21" pageNumber="168">Fig. 9A</figureCitation>
). The carapace dorsal surface is wholly covered with a thick coat of short setae; each protogastric region is provided with two transverse rows of minute granules along the anterior and median parts, which are united at the lateral end; the protogastric regions of both sides are isolated by a longitudinal median row of minute granules that reaches to the mesogastric region; a patch of minute granules on each side of the metagastric region and the cardiac region is indicated by a pair of low, granulated mounds arranged side by side. The frontal margin is cut into four shallow lobes, the inner pair being about half as wide as the outer pair. The carapace anterolateral margin is gently curved, with nine serrated teeth that are subequal or becoming slightly larger posteriorly; the last (epibranchial tooth) is about two times longer than the eighth tooth, directed laterally and weakly curved anteriorly, with the sharp tip. The basal half of the G1 is stout, and distal half tapers and strongly curved outwards (
<figureCitation id="4FA254CCFFEB9027FBC1FA58FB475CA1" box="[1108,1192,1394,1418]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." pageId="21" pageNumber="168">Fig. 9B</figureCitation>
<figureCitation id="4FA254CCFFEB9027FBC1FA58FB475CA1" box="[1108,1192,1394,1418]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="21" pageNumber="168">Fig. 9B</figureCitation>
).
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(
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<figureCitation id="4FA254CCFFEF9023FC25FD43FC0A5B55" box="[944,997,617,639]" captionStart="Fig" captionStartId="18.[163,196,1551,1573]" captionTargetBox="[158,1217,214,1516]" captionTargetId="figure-85@18.[157,1218,213,1517]" captionTargetPageId="18" captionText="Fig. 8. Charybdis (Archias) hongkongensis Shen, Ə (NSMT-Cr 30919; CB 36.7×CL 25.7mm) from sta. 42. Dorsal view (A), right chela in lower view (B), left chela in outer view (C), pleon (D), and G1s in situ (E)." figureDoi="http://doi.org/10.5281/zenodo.12760103" httpUri="https://zenodo.org/record/12760103/files/figure.png" pageId="17" pageNumber="164">Fig. 8</figureCitation>
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@ -115,7 +116,7 @@ of this species as
and
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. The shape of the six frontal lobes is somewhat variable and so more emphasised in the present male (
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), as useful to discriminate
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@ -82,7 +83,7 @@ Alcock, 1899
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<paragraph id="D7264849FFEF9023FC01F93CFBED5F07" blockId="17.[916,1026,1558,1580]" box="[916,1026,1558,1580]" pageId="17" pageNumber="164">
(
<figureCitation id="4FA254CCFFEF9023FC0EF93CFC125F07" box="[923,1021,1558,1580]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." pageId="17" pageNumber="164">Fig. 9C–D</figureCitation>
<figureCitation id="4FA254CCFFEF9023FC0EF93CFC125F07" box="[923,1021,1558,1580]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="17" pageNumber="164">Fig. 9C–D</figureCitation>
)
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@ -110,7 +111,7 @@ Alcock, 1899
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as its synonym.
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<paragraph id="D7264849FFEC9020FF36F925FB7A5F6A" blockId="18.[163,1214,1549,1602]" pageId="18" pageNumber="165">
Fig. 8.
<taxonomicName id="109933CAFFEC9020FF6CF925FD915F0E" authority="Shen" authorityName="Shen" authorityYear="1934" box="[249,638,1551,1573]" class="Malacostraca" family="Portunidae" genus="Charybdis" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="165" phylum="Arthropoda" rank="species" species="hongkongensis" subGenus="Archias">
@ -127,14 +128,14 @@ Shen
is a small species, reaching at most about
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in the carapace breadth including epibranchial teeth. The carapace surface (
<figureCitation id="4FA254CCFFEC9020FC90F980FCB15FE9" box="[773,862,1706,1730]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." pageId="18" pageNumber="165">Fig. 9C</figureCitation>
<figureCitation id="4FA254CCFFEC9020FC90F980FCB15FE9" box="[773,862,1706,1730]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="18" pageNumber="165">Fig. 9C</figureCitation>
) is wholly pilose, with granular patches on the regions; most characteristic is the somewhat Y-shaped, thick cardiac region; otherwise, it is noted on close examination that the metagastric transverse ridge and metagastric region are connected by a median longitudinal line of granules. The front (
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<figureCitation id="4FA254CCFFED9021FE59F935FDAB5F1C" box="[460,580,1567,1591]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="19" pageNumber="166">Fig. 9C–D</figureCitation>
) is low and six-lobed; the median lobe is inclined laterally, the submedian lobe is subtruncated along the inner margin, with the lateral angle hardly attaining the level of the median lobe; the inner part of the submedian lobe slightly overlaps the median lobe; the lateral lobe is narrow and separated from the submedian lobe by a deep notch. The orbit is large, with the diameter wider than half the frontal length. As seen in
<figureCitation id="4FA254CCFFED9021FBAEFAD7FB5E5F3E" box="[1083,1201,1533,1557]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." pageId="19" pageNumber="166">Fig. 9C–D</figureCitation>
<figureCitation id="4FA254CCFFED9021FBAEFAD7FB5E5F3E" box="[1083,1201,1533,1557]" captionStart="Fig" captionStartId="19.[163,195,1340,1362]" captionTargetBox="[158,1217,215,1306]" captionTargetId="figure-168@19.[157,1218,213,1307]" captionTargetPageId="19" captionText="Fig. 9. A–B: Lupocycloporus innominatus (Rathbun), Ə (NSMT-Cr 30923; CB 15.1mm including lateral teeth, CL 10.5mm) from sta. 45. C–D: Charybdis (Archias) vadorum Alcock, Ə (NSMT-Cr 30920; CB 10.7×CL 7.6mm) from sta. 50. E–F: Thalamita sexlobata Miers, ovig.8 (NSMT-Cr 30926; CB 7.0×CL 6.0mm) from sta. 45." figureDoi="http://doi.org/10.5281/zenodo.12760109" httpUri="https://zenodo.org/record/12760109/files/figure.png" pageId="19" pageNumber="166">Fig. 9C–D</figureCitation>
, of the six anterolateral teeth of the carapace, the first (the external orbital tooth) is directed forward and slightly larger than the second and fourth teeth, but slightly smaller than the third and fourth teeth; the last (the epibranchial tooth) is narrow, much longer than the precedings and sharply directed laterally.
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Fig. 9. A–B:
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(
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<figureCitation id="4FA254CCFFF2903EFC09FC76FC145A5A" box="[924,1019,860,882]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." figureDoi="http://doi.org/10.5281/zenodo.12760091" httpUri="https://zenodo.org/record/12760091/files/figure.png" pageId="12" pageNumber="159">Fig. 3C–E</figureCitation>
)
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@ -106,7 +107,7 @@ excluding lateral plates, PCL
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<emphasis id="E5ED945BFFF2903EFD48FB55FCD55DBC" box="[733,826,1151,1175]" italics="true" pageId="12" pageNumber="159">Remarks</emphasis>
. The specimens (
<figureCitation id="4FA254CCFFF2903EFB9BFB55FB6F5DBC" box="[1038,1152,1151,1175]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." pageId="12" pageNumber="159">Fig. 3C–E</figureCitation>
<figureCitation id="4FA254CCFFF2903EFB9BFB55FB6F5DBC" box="[1038,1152,1151,1175]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." figureDoi="http://doi.org/10.5281/zenodo.12760091" httpUri="https://zenodo.org/record/12760091/files/figure.png" pageId="12" pageNumber="159">Fig. 3C–E</figureCitation>
) examined agree well with the original description (
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), and the accounts (

101
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(
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<figureCitation id="4FA254CCFFF6903AFEE4FA34FE575C1F" box="[369,440,1310,1332]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." figureDoi="http://doi.org/10.5281/zenodo.12760091" httpUri="https://zenodo.org/record/12760091/files/figure.png" pageId="8" pageNumber="155">Fig. 3A</figureCitation>
)
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@ -104,7 +105,7 @@ A fully-grown female at hand generally agrees with the original description (
<emphasis id="E5ED945BFFF6903AFF7EF935FE8C5F1C" box="[235,355,1567,1591]" italics="true" pageId="8" pageNumber="155">Achaeopsis</emphasis>
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), though the protogastric and epibranchial spines (
<figureCitation id="4FA254CCFFF6903AFEFBF968FE2A5F71" box="[366,453,1602,1626]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." pageId="8" pageNumber="155">Fig. 3A</figureCitation>
<figureCitation id="4FA254CCFFF6903AFEFBF968FE2A5F71" box="[366,453,1602,1626]" captionStart="Fig" captionStartId="9.[163,196,1216,1238]" captionTargetBox="[158,1217,215,1182]" captionTargetId="figure-192@9.[157,1218,213,1183]" captionTargetPageId="9" captionText="Fig. 3. A: Dorhynchus rostratus (Sakai), 8 (NSMT-Cr 30908; CB 6.6×PCL 8.5mm) from sta. 54. B: Oncino- pus angustifrons Takeda and Miyake, Ə (NSMT-Cr 30909; CB 3.4×PCL 4.3mm) from sta. 48. C–E: Oxypleurodon sphaenocarcinoides (Rathbun), ovig.8 (NSMT-Cr 30913; CB 7.8×PCL 12.7mm) (C–D) from sta. 54; 8(NSMT-Cr 30914; 5.0×8.3mm) (E) from sta. 54." figureDoi="http://doi.org/10.5281/zenodo.12760091" httpUri="https://zenodo.org/record/12760091/files/figure.png" pageId="8" pageNumber="155">Fig. 3A</figureCitation>
) are not prominent as in the figures represented by
<bibRefCitation id="B30835B8FFF6903AFE7CF94FFD825F56" author="Sakai, T." box="[489,621,1637,1661]" pageId="8" pageNumber="155" pagination="41 - 59" refId="ref27728" refString="Sakai, T. 1932. Notes on some rare materials of Japanese Oxyrhyncha. Science Reports of the Tokyo Bunrika Daigaku, (B) 1: 41 - 59, pls. 2 - 3." type="journal article" year="1932">Sakai (1932</bibRefCitation>
, fig. 3; 1938, fig. 17) and also in the female from the Timor Sea (

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(
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<figureCitation id="4FA254CCFFFA9036FC0EFD84FC135BEF" box="[923,1020,686,708]" captionStart="Fig" captionStartId="5.[163,196,1344,1366]" captionTargetBox="[158,1217,215,1309]" captionTargetId="figure-174@5.[157,1218,213,1310]" captionTargetPageId="5" captionText="Fig. 2. A–B: Cosmonotus grayii White, 8 (NSMT-Cr 30903; CB 5.1×CL 6.1mm) from sta. 45. C–D: Cosmonotus genkaiae Takeda and Miyake, Ə (NSMT-Cr 13917; CB 14.2×CL 19.4mm) from the East China Sea, 182–167m depth." figureDoi="http://doi.org/10.5281/zenodo.12760089" httpUri="https://zenodo.org/record/12760089/files/figure.png" pageId="4" pageNumber="151">Fig. 2A–B</figureCitation>
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@ -151,7 +152,7 @@ Seo
; the main part of the carapace dorsal surface is smooth, but the anterolateral portion is distinctly roughened with minute granules of irregular size, and the carapace midline is rather thickened and ridged for much of its length, somewhat like a lbackbone.z
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Fig. 2. A–B:
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@ -213,7 +214,7 @@ since the original description. The photograph is not clear, but the diagnostic
Sea, 182–
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depth (NSMT-Cr 13917,
<figureCitation id="4FA254CCFFF89034FDB4FEAFFD7A58B6" box="[545,661,389,413]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." pageId="6" pageNumber="153">Fig. 1C–D</figureCitation>
<figureCitation id="4FA254CCFFF89034FDB4FEAFFD7A58B6" box="[545,661,389,413]" captionStart="Fig" captionStartId="3.[163,196,1420,1442]" captionTargetBox="[158,1217,214,1386]" captionTargetId="figure-90@3.[157,1218,213,1387]" captionTargetPageId="3" captionText="Fig. 1. A: Mclaydromia colini Guinot and Tavares, 8 (NSMT-Cr 30898; CB 13.1×CL 15.0mm including pseudorostral teeth) from sta. 45. B: Mursia australiensis Campbell, juv. (NSMT-Cr 30905; CB 8.2mm excluding lateral tubercles, CL 7.4 mm) from sta. 52. C: Podophthalmus nacreus Alcock, 8 (NSMT-Cr 30925; CB 11.0×CL 7.1mm) from sta. 45. D: Lissocarcinus arkati Kemp, 8 (NSMT-Cr 30921; CB 9.8×CL 8.7mm) from sta. 45. E–F: Parilia pattersoni Ng, Devi and Kumar, juv. (NSMT-Cr 30906; CB 5.6×CL 6.6mm including posterior median tubercle) from sta. 42." figureDoi="http://doi.org/10.5281/zenodo.12760087" httpUri="https://zenodo.org/record/12760087/files/figure.png" pageId="6" pageNumber="153">Fig. 1C–D</figureCitation>
) preserved in the Tsukuba Research Departments, National Museum of Nature and Science, Tokyo. In this specimen, the carapace dorsal surface is much more strongly striated than in the
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, with the longer and deeper striae closely set together, and the carapace dorsal midline is weakly ridged throughout the whole length; the frontal median sinus is provided with a small tubercle ventrally, as mentioned in the original description, and the supraorbital margin is interrupted only by one notch.

101
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<heading id="425681E1FF8F391B04DDADCBFDE2FDA8" box="[174,635,505,530]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="118" reason="4">
<taxonomicName id="DEA14D0EFF8F391B04DDADCBFDE2FDA8" ID-CoL="7FQNS" authority="Ng, Dai and Yang, 1997" authorityName="Ng, Dai and Yang" authorityYear="1997" box="[174,635,505,530]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="118" phylum="Arthropoda" rank="species" species="sentus">
<emphasis id="2BD5EA9FFF8F391B04DDADCBFEEDFDAB" bold="true" box="[174,372,505,529]" italics="true" pageId="11" pageNumber="118">Viaderiana sentus</emphasis>
<bibRefCitation id="7D304B7CFF8F391B0509ADC8FDE2FDA8" author="Ng, P. K. L. &amp; A. - Y. Dai &amp; S. - L. Yang" box="[378,635,506,530]" pageId="11" pageNumber="118" pagination="145 - 159" refId="ref6816" refString="Ng, P. K. L., A. - Y. Dai and S. - L. Yang 1997. The Indo- West Pacific Pilumnidae X. New species and records from the South China Sea (Crustacea: Decapoda: Brachyura). The Raffles Bulletin of Zoology 45: 145 - 159." type="journal article" year="1997">Ng, Dai and Yang, 1997</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="51BB6506FF8F391B0510AE2BFE5EFD94" box="[355,455,537,559]" pageId="11" pageNumber="118" type="description">
<paragraph id="191E368DFF8F391B0510AE2BFE5EFD94" blockId="11.[174,635,505,559]" box="[355,455,537,559]" pageId="11" pageNumber="118">
(
<figureCitation id="819A2A08FF8F391B0519AE2BFE27FD94" box="[362,446,537,559]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[163,196,1539,1561]" captionStartId-1="13.[163,196,1416,1438]" captionTargetBox-0="[159,1216,215,1504]" captionTargetBox-1="[159,1217,216,1380]" captionTargetId-0="figure-67@12.[157,1218,213,1506]" captionTargetId-1="figure-132@13.[156,1220,213,1383]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 8. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Habitus in dorsal view (A), and carapace, enlarged (B). In Fig. A, the right half of the carapace is denuded, with the horny tip of each anterolateral tooth broken off." captionText-1="Fig. 9. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Right chela in dorsal view (A), right first to fourth ambulatory legs in dorsal view (B–E); left G1 in sternal view (F) and its distal part in abdominal view (G). Scales:A–E=5mm; G=4mm; F=2mm." figureDoi-0="http://doi.org/10.5281/zenodo.12760379" figureDoi-1="http://doi.org/10.5281/zenodo.12760381" httpUri-0="https://zenodo.org/record/12760379/files/figure.png" httpUri-1="https://zenodo.org/record/12760381/files/figure.png" pageId="11" pageNumber="118">Figs. 8–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="51BB6506FF8F391B04FFAE7BFD80FDC6" pageId="11" pageNumber="118" type="reference_group">
<paragraph id="191E368DFF8F391B04FFAE7BFD80FDC6" blockId="11.[140,670,585,636]" pageId="11" pageNumber="118">
<treatmentCitationGroup id="39B111A3FF8F391B04FFAE7BFD80FDC6" pageId="11" pageNumber="118">
<treatmentCitation id="9800109CFF8F391B04FFAE7BFDC3FDE5" author="Ng, P. K. L. &amp; A. - Y. Dai &amp; S. - L. Yang" box="[140,602,585,607]" page="156" pageId="11" pageNumber="118" year="1997">
<taxonomicName id="DEA14D0EFF8F391B04FFAE7BFDC3FDE5" ID-CoL="7FQNS" authority="Ng, Dai &amp; Yang, 1997, p. 156" authorityName="Ng, Dai &amp; Yang" authorityPageNumber="156" authorityYear="1997" box="[140,602,585,607]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="118" phylum="Arthropoda" rank="species" species="sentus">
<emphasis id="2BD5EA9FFF8F391B04FFAE7BFEA8FDE5" box="[140,305,585,607]" italics="true" pageId="11" pageNumber="118">Viaderiana sentus</emphasis>
<bibRefCitation id="7D304B7CFF8F391B054AAE78FD88FDE4" author="Ng, P. K. L. &amp; A. - Y. Dai &amp; S. - L. Yang" box="[313,529,585,607]" pageId="11" pageNumber="118" pagination="145 - 159" refId="ref6816" refString="Ng, P. K. L., A. - Y. Dai and S. - L. Yang 1997. The Indo- West Pacific Pilumnidae X. New species and records from the South China Sea (Crustacea: Decapoda: Brachyura). The Raffles Bulletin of Zoology 45: 145 - 159." type="journal article" year="1997">Ng, Dai &amp; Yang, 1997</bibRefCitation>
, p. 156
</taxonomicName>
</treatmentCitation>
, fig. 6. —
<treatmentCitation id="9800109CFF8F391B04CEAE55FE3DFDC6" author="Ng, P. K. L. &amp; C. - W. Lin &amp; P. - H. Ho" box="[189,420,614,636]" page="7" pageId="11" pageNumber="118" year="2018">
<bibRefCitation id="7D304B7CFF8F391B04CEAE55FEE1FDC6" author="Ng, P. K. L. &amp; C. - W. Lin &amp; P. - H. Ho" box="[189,376,614,636]" pageId="11" pageNumber="118" refId="ref7036" refString="Ng, P. K. L., C. - W. Lin and P. - H. Ho 2018. On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on Heteropilumnus De Man, 2895 (Crustacea: Brachyura). Zoological Studies 57: 12 (17 pp.). doi: 10.6620 / ZS. 2018.57 - 12." type="book" year="2018">Ng, Lin &amp; Ho, 2018</bibRefCitation>
, p. 7
</treatmentCitation>
, figs. 10–13.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="51BB6506FF8F391B04D4AEA8FBA4FB2D" pageId="11" pageNumber="118" type="materials_examined">
<paragraph id="191E368DFF8F391B04D4AEA8FE84FC87" blockId="11.[140,669,665,1799]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B04D4AEA8FE18FD08" box="[167,385,666,690]" italics="true" pageId="11" pageNumber="118">Material examined.</emphasis>
<materialsCitation id="A9C93CD0FF8F391B05E9AEA8FE84FC87" collectingDate="2018-09-30" collectorName="Luzon I. &amp; H. Takakura" country="Philippines" elevation="4" location="Dive" municipality="Anilao" pageId="11" pageNumber="118" specimenCount="1" stateProvince="Batangas">
<location id="1C7E6056FF8F391B05E9AEA8FE57FD08" LSID="urn:lsid:plazi:treatment:9108879BFF8F391B04DDADCBFBA4FB2D:1C7E6056FF8F391B05E9AEA8FE57FD08" box="[410,462,666,690]" country="Philippines" municipality="Anilao" name="Dive" pageId="11" pageNumber="118" stateProvince="Batangas">Dive</location>
site lSecret Gar- denz,
<collectingMunicipality id="F97AACF7FF8F391B04A3AE8FFE82FD6F" box="[208,283,701,725]" pageId="11" pageNumber="118">Anilao</collectingMunicipality>
,
<collectingRegion id="DB65F86FFF8F391B055CAE8FFE0AFD6F" box="[303,403,701,725]" country="Philippines" name="Batangas" pageId="11" pageNumber="118">Batangas</collectingRegion>
,
<collectorName id="B454535BFF8F391B05D5AE8FFD93FD6E" box="[422,522,701,724]" pageId="11" pageNumber="118">Luzon I.</collectorName>
,
<collectingCountry id="61B6761DFF8F391B066DAE8FFD00FD6F" box="[542,665,701,725]" name="Philippines" pageId="11" pageNumber="118">Philippines</collectingCountry>
,
<elevation id="928CD1BEFF8F391B04FFAEEDFF62FD4D" box="[140,251,735,759]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="118" unit="m" value="4.0">
<quantity id="DE599B68FF8F391B04FFAEEDFF2BFD4D" box="[140,178,735,759]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="118" unit="m" value="4.0">4 m</quantity>
depth
</elevation>
, sandy/muddy bottom; 1 Ə (NMCR-92509; cb
<quantity id="DE599B68FF8F391B0577AF30FECCFCA3" box="[260,341,770,794]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="118" unit="mm" value="8.4">8.4 mm</quantity>
, cl
<quantity id="DE599B68FF8F391B05F2AF30FE4DFCA0" box="[385,468,770,794]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="118" unit="mm" value="6.4">6.4 mm</quantity>
);
<date id="6D1F104DFF8F391B0599AF30FDEDFCA0" box="[490,628,770,794]" pageId="11" pageNumber="118" value="2018-09-30">
<collectingDate id="7D5BE9A5FF8F391B0599AF30FDEDFCA0" box="[490,628,770,794]" pageId="11" pageNumber="118" value="2018-09-30">30–IX–2018</collectingDate>
</date>
;
<collectorName id="B454535BFF8F391B06F0AF30FF76FC87" pageId="11" pageNumber="118">H. Takakura</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="191E368DFF8F391B04D4AF7AFD7FFE27" blockId="11.[140,669,665,1799]" lastBlockId="11.[707,1240,215,1175]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B04D4AF7AFE83FCE5" box="[167,282,840,863]" italics="true" pageId="11" pageNumber="118">Diagnosis.</emphasis>
Carapace (
<figureCitation id="819A2A08FF8F391B05C3AF75FDB5FCE5" box="[432,556,839,863]" captionStart="Fig" captionStartId="12.[163,196,1539,1561]" captionTargetBox="[159,1216,215,1504]" captionTargetId="figure-67@12.[157,1218,213,1506]" captionTargetPageId="12" captionText="Fig. 8. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Habitus in dorsal view (A), and carapace, enlarged (B). In Fig. A, the right half of the carapace is denuded, with the horny tip of each anterolateral tooth broken off." figureDoi="http://doi.org/10.5281/zenodo.12760379" httpUri="https://zenodo.org/record/12760379/files/figure.png" pageId="11" pageNumber="118">Fig. 8A–B</figureCitation>
) rounded quadrate; dorsal surface moderately convex in both directions, smooth, without granules, shallowly divided into regions by indistinct linear furrows; hairs short, soft, regularly cover whole surface, but not disguise areolation; some tufts of longer, club-shaped hairs on frontal, protogastric, branchial regions, a similar tuft of some club-shaped hairs at each lateral end of transverse furrow in front of cardiac region; external orbital angle armed with a sharp, forward-directed spine; infraorbital margin armed with four sharp, equidistant spines along inner part, inner infraorbital angle terminating in a sharp spine; anterolateral margin armed with three sharp spines on conical or weakly bulged bases behind external orbital spine, last spine smaller than others, subequal to external orbital spine. Both chelipeds (
<figureCitation id="819A2A08FF8F391B04E7A985FF77FA75" box="[148,238,1463,1487]" captionStart="Fig" captionStartId="13.[163,196,1416,1438]" captionTargetBox="[159,1217,216,1380]" captionTargetId="figure-132@13.[156,1220,213,1383]" captionTargetPageId="13" captionText="Fig. 9. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Right chela in dorsal view (A), right first to fourth ambulatory legs in dorsal view (B–E); left G1 in sternal view (F) and its distal part in abdominal view (G). Scales:A–E=5mm; G=4mm; F=2mm." figureDoi="http://doi.org/10.5281/zenodo.12760381" httpUri="https://zenodo.org/record/12760381/files/figure.png" pageId="11" pageNumber="118">Fig. 9A</figureCitation>
) covered with short hairs interspaced with long hairs, some long hairs club-shaped; carpus armed with several distant, dark-colored, curved spines on outer surface, inner angle strongly produced, elongated, tipped with curved, dark-colored spine; palm also armed with sharp, curved, distant spines on outer surface, in smaller chela, spines more prominent than in larger chela. Ambulatory legs (
<figureCitation id="819A2A08FF8F391B0655AAFFFD0CF95F" box="[550,661,1741,1765]" captionStart="Fig" captionStartId="13.[163,196,1416,1438]" captionTargetBox="[159,1217,216,1380]" captionTargetId="figure-132@13.[156,1220,213,1383]" captionTargetPageId="13" captionText="Fig. 9. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Right chela in dorsal view (A), right first to fourth ambulatory legs in dorsal view (B–E); left G1 in sternal view (F) and its distal part in abdominal view (G). Scales:A–E=5mm; G=4mm; F=2mm." figureDoi="http://doi.org/10.5281/zenodo.12760381" httpUri="https://zenodo.org/record/12760381/files/figure.png" pageId="11" pageNumber="118">Fig. 9B–E</figureCitation>
) moderately long; each merus armed with one to three sharp spines on anterior margin and a terminal spine; each carpus of all pairs armed anteriorly with a terminal, strong erect spine. G1 (
<figureCitation id="819A2A08FF8F391B06B8AD0DFCA0FEED" box="[715,825,319,343]" captionStart="Fig" captionStartId="13.[163,196,1416,1438]" captionTargetBox="[159,1217,216,1380]" captionTargetId="figure-132@13.[156,1220,213,1383]" captionTargetPageId="13" captionText="Fig. 9. Viaderiana sentus Ng, Dai and Yang, 1Ə (NMCR-92509; cb 8.4×cl 6.4mm). Right chela in dorsal view (A), right first to fourth ambulatory legs in dorsal view (B–E); left G1 in sternal view (F) and its distal part in abdominal view (G). Scales:A–E=5mm; G=4mm; F=2mm." figureDoi="http://doi.org/10.5281/zenodo.12760381" httpUri="https://zenodo.org/record/12760381/files/figure.png" pageId="11" pageNumber="118">Fig. 9F, G</figureCitation>
) long, not strongly curved, with distal beak comparatively long and sharply pointed at tip.
</paragraph>
<paragraph id="191E368DFF8F391B06AEAD95FD67FDD6" blockId="11.[707,1240,215,1175]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B06AEAD95FCFFFE05" box="[733,870,423,447]" italics="true" pageId="11" pageNumber="118">Color in life</emphasis>
. The carapace dorsal surface is dark reddish brown, with some bunds of long feathered, grayish or whitish hairs symmetrically arranged. The chelipeds and ambulatory legs are marked finely with light purplish reticulate patterns.
</paragraph>
<paragraph id="191E368DFF8F391B06AEAE45FB18FC38" blockId="11.[707,1240,215,1175]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B06AEAE45FCA3FD35" box="[733,826,631,655]" italics="true" pageId="11" pageNumber="118">Remarks</emphasis>
. This species was fully described and figured in the original description by
<bibRefCitation id="7D304B7CFF8F391B00E5AEA8FCA9FD6F" author="Ng, P. K. L. &amp; A. - Y. Dai &amp; S. - L. Yang" pageId="11" pageNumber="118" pagination="145 - 159" refId="ref6816" refString="Ng, P. K. L., A. - Y. Dai and S. - L. Yang 1997. The Indo- West Pacific Pilumnidae X. New species and records from the South China Sea (Crustacea: Decapoda: Brachyura). The Raffles Bulletin of Zoology 45: 145 - 159." type="journal article" year="1997">
Ng
<emphasis id="2BD5EA9FFF8F391B00B2AEA9FD41FD6F" italics="true" pageId="11" pageNumber="118">et al</emphasis>
. (1997)
</bibRefCitation>
, to which the present specimen agrees well in all the respects. Ng
<emphasis id="2BD5EA9FFF8F391B078BAED2FBB3FD4D" box="[1016,1066,735,759]" italics="true" pageId="11" pageNumber="118">et al</emphasis>
. (2008) treated the genus
<taxonomicName id="DEA14D0EFF8F391B0742AF30FC3EFCA0" authorityName="Ward" authorityYear="1942" box="[817,935,770,794]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="2BD5EA9FFF8F391B0742AF30FC3EFCA0" box="[817,935,770,794]" italics="true" pageId="11" pageNumber="118">Viaderiana</emphasis>
</taxonomicName>
to include 17 valid species; among them,
<taxonomicName id="DEA14D0EFF8F391B0725AF17FC2DFC86" authorityName="Ng, Dai &amp; Yang" authorityYear="1997" box="[854,948,805,828]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="118" phylum="Arthropoda" rank="species" species="sentus">
<emphasis id="2BD5EA9FFF8F391B0725AF17FC2DFC86" box="[854,948,805,828]" italics="true" pageId="11" pageNumber="118">V. sentus</emphasis>
</taxonomicName>
is characteristic especially in the arrangement of anterior marginal spines of the meri and carpi of the ambulatory legs.
</paragraph>
<paragraph id="191E368DFF8F391B06AEAFBFFBA4FB2D" blockId="11.[707,1240,215,1175]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B06AEAFBFFCC6FC1F" box="[733,863,909,933]" italics="true" pageId="11" pageNumber="118">Distribution</emphasis>
. Previously recorded from the Nansha Islands in the South
<collectingCountry id="61B6761DFF8F391B0055AF9DFBFFFC7D" box="[1062,1126,943,967]" name="China" pageId="11" pageNumber="118">China</collectingCountry>
Sea and southern
<collectingCountry id="61B6761DFF8F391B0756AFE0FCEAFC50" box="[805,883,978,1002]" name="Taiwan" pageId="11" pageNumber="118">Taiwan</collectingCountry>
(
<bibRefCitation id="7D304B7CFF8F391B07F1AFE0FBBEFC50" author="Ng, P. K. L. &amp; A. - Y. Dai &amp; S. - L. Yang" box="[898,1063,978,1002]" pageId="11" pageNumber="118" pagination="145 - 159" refId="ref6816" refString="Ng, P. K. L., A. - Y. Dai and S. - L. Yang 1997. The Indo- West Pacific Pilumnidae X. New species and records from the South China Sea (Crustacea: Decapoda: Brachyura). The Raffles Bulletin of Zoology 45: 145 - 159." type="journal article" year="1997">
Ng
<emphasis id="2BD5EA9FFF8F391B07D8AFE1FC42FC50" box="[939,987,978,1002]" italics="true" pageId="11" pageNumber="118">et al</emphasis>
., 1997
</bibRefCitation>
,
<bibRefCitation id="7D304B7CFF8F391B0041AFE0FBF5FC50" author="Ng, P. K. L. &amp; C. - W. Lin &amp; P. - H. Ho" box="[1074,1132,978,1002]" pageId="11" pageNumber="118" refId="ref7036" refString="Ng, P. K. L., C. - W. Lin and P. - H. Ho 2018. On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on Heteropilumnus De Man, 2895 (Crustacea: Brachyura). Zoological Studies 57: 12 (17 pp.). doi: 10.6620 / ZS. 2018.57 - 12." type="book" year="2018">2018</bibRefCitation>
). Another
<taxonomicName id="DEA14D0EFF8F391B06B0AFC7FC0CFBB7" box="[707,917,1013,1037]" pageId="11" pageNumber="118">
<emphasis id="2BD5EA9FFF8F391B06B0AFC7FCA0FBB7" box="[707,825,1013,1037]" italics="true" pageId="11" pageNumber="118">Viaderiana</emphasis>
species
</taxonomicName>
from the
<collectingCountry id="61B6761DFF8F391B006BAFC7FB09FBB7" box="[1048,1168,1013,1037]" name="Philippines" pageId="11" pageNumber="118">Philippines</collectingCountry>
is
<taxonomicName id="DEA14D0EFF8F391B00B3AFC7FBAEFB95" authority="Takeda and Manuel, 2003" authorityName="Takeda and Manuel" authorityYear="2003" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="118" phylum="Arthropoda" rank="species" species="kasei">
<emphasis id="2BD5EA9FFF8F391B00B3AFC7FD63FB95" italics="true" pageId="11" pageNumber="118">V. kasei</emphasis>
<bibRefCitation id="7D304B7CFF8F391B077BA825FBAEFB95" author="Takeda, M. &amp; M. Manuel" box="[776,1079,1047,1071]" pageId="11" pageNumber="118" pagination="205 - 211" refId="ref7403" refString="Takeda, M. and M. Manuel 2003. A new crab of the family Pilumnidae (Crustacea, Decapoda), Viaderiana kasei from the Philippines. Bulletin of the National Science Museum, Tokyo, Series A 29: 205 - 211." type="journal article" year="2003">Takeda and Manuel, 2003</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="DB65F86FFF8F391B00FFA825FB49FB95" box="[1164,1232,1047,1071]" country="Philippines" name="Bohol" pageId="11" pageNumber="118">Bohol</collectingRegion>
, about
<quantity id="DE599B68FF8F391B0775A808FCDFFBEB" box="[774,838,1082,1106]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="118" unit="m" value="150.0">150 m</quantity>
depth, in which the ambulatory legs are remarkably long, with the third pair exceeding three times the carapace length.
</paragraph>
</subSubSection>
</treatment>
</document>