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<mods:title id="728579ED8BDDF785E5DDCA3131FB944F">Redescription of Tetebius latibunus as a new senior synonym of Malgaceros boviceps (Opiliones: Laniatores: Biantidae)</mods:title>
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<mods:affiliation id="37A9FCB64474B0CA4491984554ADCAA6">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina.</mods:affiliation>
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<taxonomicName id="8CC456E6FFDDCD405DDFFE8D4A9EFE95" ID-CoL="646ZM" authority="Roewer, 1949" authorityName="Roewer" authorityYear="1949" box="[151,426,258,285]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405DDFFE8D4A9EFE95" bold="true" box="[151,426,258,285]" pageId="2" pageNumber="327">
<emphasis id="79B0F177FFDDCD405DDFFE8D4BCFFE94" bold="true" box="[151,251,258,285]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
<bibRefCitation id="2F555094FFDDCD405C4AFE8C4A9EFE95" author="Roewer, C. F." box="[258,426,258,285]" pageId="2" pageNumber="327" pagination="247 - 289" refId="ref9357" refString="Roewer, C. F. (1949) Uber Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30 (4), 247 - 289." type="journal article" year="1949">Roewer, 1949</bibRefCitation>
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urn:lsid:zoobank.org:act:
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<treatmentCitation id="CA650B74FFDDCD405DDFFEE44A9BFE0A" author="Roewer, C. F." box="[151,431,363,387]" page="288" pageId="2" pageNumber="327" year="1949">
<taxonomicName id="8CC456E6FFDDCD405DDFFEE44A9BFE0A" ID-CoL="646ZM" authority="Roewer 1949: 288" authorityName="Roewer" authorityPageNumber="288" authorityYear="1949" box="[151,431,363,387]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405DDFFEE44BDFFE0A" box="[151,235,363,387]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
<bibRefCitation id="2F555094FFDDCD405DB9FEE34A9BFE0A" author="Roewer, C. F." box="[241,431,363,387]" pageId="2" pageNumber="327" pagination="247 - 289" refId="ref9357" refString="Roewer, C. F. (1949) Uber Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30 (4), 247 - 289." type="journal article" year="1949">Roewer 1949: 288</bibRefCitation>
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;
<treatmentCitation id="CA650B74FFDDCD405CF2FEE44941FE0A" author="Starega, W." box="[442,629,363,387]" page="330" pageId="2" pageNumber="327" year="1992">
<bibRefCitation id="2F555094FFDDCD405CF2FEE44941FE0A" author="Starega, W." box="[442,629,363,387]" pageId="2" pageNumber="327" pagination="271 - 336" refId="ref9480" refString="Starega, W. (1992) An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum, 33 (2), 271 - 336." type="journal article" year="1992">Staręga 1992: 330</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CA650B74FFDDCD405F37FEE348FDFE0A" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[639,969,363,388]" page="7" pageId="2" pageNumber="327" year="2015">
<bibRefCitation id="2F555094FFDDCD405F37FEE348FDFE0A" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[639,969,363,388]" pageId="2" pageNumber="327" refId="ref8940" refString="Kury, A. B. &amp; Perez-Gonzalez, A. (2015) A companion to Part 2 of the World Checklist of Opiliones species (Arachnida): Laniatores - Samooidea, Zalmoxoidea and Grassatores incertae sedis. Biodiversity Data Journal, 3, e 6663. https: // doi. org / 10.3897 / BDJ. 3. e 6663" type="journal volume" year="2015">Kury &amp; Pérez-González 2015: 7</bibRefCitation>
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.
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<treatmentCitationGroup id="6BD40A4BFFDDCD405DDFFE05482AFE48" pageId="2" pageNumber="327">
<treatmentCitation id="CA650B74FFDDCD405DDFFE054ADEFE2B" author="Lawrence, R. F." box="[151,490,394,418]" page="79" pageId="2" pageNumber="327" year="1959">
<taxonomicName id="8CC456E6FFDDCD405DDFFE054ADEFE2B" ID-CoL="5KJR" authority="Lawrence 1959: 79" authorityName="Lawrence" authorityPageNumber="79" authorityYear="1959" box="[151,490,394,418]" class="Arachnida" family="Biantidae" genus="Malgaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405DDFFE054A3BFE2B" box="[151,271,394,418]" italics="true" pageId="2" pageNumber="327">Malgaceros</emphasis>
<bibRefCitation id="2F555094FFDDCD405C52FE054ADEFE2B" author="Lawrence, R. F." box="[282,490,394,418]" pageId="2" pageNumber="327" pagination="1 - 121" refId="ref9004" refString="Lawrence, R. F. (1959) Arachnides-Opilions. Faune de Madagascar, Publications de L'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 9, 1 - 121." type="journal article" year="1959">Lawrence 1959: 79</bibRefCitation>
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;
<treatmentCitation id="CA650B74FFDDCD405CB1FE054989FE2B" author="Starega, W." box="[505,701,394,418]" page="330" pageId="2" pageNumber="327" year="1992">
<bibRefCitation id="2F555094FFDDCD405CB1FE054989FE2B" author="Starega, W." box="[505,701,394,418]" pageId="2" pageNumber="327" pagination="271 - 336" refId="ref9480" refString="Starega, W. (1992) An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum, 33 (2), 271 - 336." type="journal article" year="1992">Staręga 1992: 330</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CA650B74FFDDCD405F84FE054F2FFE2B" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[716,1051,394,418]" page="6" pageId="2" pageNumber="327" year="2015">
<bibRefCitation id="2F555094FFDDCD405F84FE054F2FFE2B" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[716,1051,394,418]" pageId="2" pageNumber="327" refId="ref8940" refString="Kury, A. B. &amp; Perez-Gonzalez, A. (2015) A companion to Part 2 of the World Checklist of Opiliones species (Arachnida): Laniatores - Samooidea, Zalmoxoidea and Grassatores incertae sedis. Biodiversity Data Journal, 3, e 6663. https: // doi. org / 10.3897 / BDJ. 3. e 6663" type="journal volume" year="2015">Kury &amp; Pérez-González 2015:6</bibRefCitation>
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[
<typeStatus id="947F93C7FFDDCD405964FE054F6EFE2B" box="[1068,1114,394,418]" pageId="2" pageNumber="327">type</typeStatus>
species:
<taxonomicName id="8CC456E6FFDDCD405989FE054A59FE49" ID-CoL="7VRJV" authority="Lawrence, 1959" authorityName="Lawrence" authorityYear="1959" class="Arachnida" family="Biantidae" genus="Malgaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="boviceps">
<emphasis id="79B0F177FFDDCD405989FE054EA9FE2B" box="[1217,1437,394,418]" italics="true" pageId="2" pageNumber="327">Malgaceros boviceps</emphasis>
<bibRefCitation id="2F555094FFDDCD405D8FFE264A59FE49" author="Lawrence, R. F." box="[199,365,424,448]" pageId="2" pageNumber="327" pagination="1 - 121" refId="ref9004" refString="Lawrence, R. F. (1959) Arachnides-Opilions. Faune de Madagascar, Publications de L'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 9, 1 - 121." type="journal article" year="1959">Lawrence, 1959</bibRefCitation>
</taxonomicName>
, by original designation].
<emphasis id="79B0F177FFDDCD405F3EFE26482AFE48" bold="true" box="[630,798,425,449]" pageId="2" pageNumber="327">New synonymy.</emphasis>
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</paragraph>
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<subSubSection id="03DE7EEEFFDDCD405DDFFE61494EFCF8" pageId="2" pageNumber="327" type="diagnosis">
<paragraph id="4B7B2D65FFDDCD405DDFFE61494EFCF8" blockId="2.[151,1437,494,1061]" pageId="2" pageNumber="327">
<emphasis id="79B0F177FFDDCD405DDFFE614A26FD81" bold="true" box="[151,274,494,520]" pageId="2" pageNumber="327">Diagnosis.</emphasis>
<taxonomicName id="8CC456E6FFDDCD405C55FE604A4DFD80" authorityName="Roewer" authorityYear="1949" box="[285,377,495,521]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405C55FE604A4DFD80" box="[285,377,495,521]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
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can be distinguished from most genera of
<taxonomicName id="8CC456E6FFDDCD405E30FE6048D7FD80" authorityName="Thorell" authorityYear="1889" box="[888,995,495,521]" class="Arachnida" family="Biantidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="subFamily" subFamily="Biantinae">Biantinae</taxonomicName>
by the absence of the typical biantid pedipalp shape with a thin and generally unarmed femur (
<emphasis id="79B0F177FFDDCD405E68FD9C487CFDA5" box="[800,840,531,556]" italics="true" pageId="2" pageNumber="327">e.g.</emphasis>
,
<bibRefCitation id="2F555094FFDDCD405E1EFD9C48C0FDA4" author="Martens, J." box="[854,1012,531,557]" pageId="2" pageNumber="327" pagination="347 - 414" refId="ref9035" refString="Martens, J. (1978) Opiliones aus dem Nepal-Himalaya. IV. Biantidae (Arachnida). Senckenbergiana biologica, 58 (5 / 6), 347 - 414." type="journal article" year="1978">Martens 1978</bibRefCitation>
, fig. 2). By contrast,
<taxonomicName id="8CC456E6FFDDCD4059A8FD9C4E08FDA4" authorityName="Roewer" authorityYear="1949" box="[1248,1340,531,557]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD4059A8FD9C4E08FDA4" box="[1248,1340,531,557]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
exhibits a robust, curved, and armed femur, a feature shared with
<taxonomicName id="8CC456E6FFDDCD405E55FDB84F0DFDD8" authority="Roewer, 1949" authorityName="Roewer" authorityYear="1949" box="[797,1081,567,593]" class="Arachnida" family="Biantidae" genus="Hirstienus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405E55FDB848A6FDD9" box="[797,914,567,592]" italics="true" pageId="2" pageNumber="327">Hirstienus</emphasis>
<bibRefCitation id="2F555094FFDDCD405ED1FDB84F0DFDD8" author="Roewer, C. F." box="[921,1081,567,593]" pageId="2" pageNumber="327" pagination="247 - 289" refId="ref9357" refString="Roewer, C. F. (1949) Uber Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30 (4), 247 - 289." type="journal article" year="1949">Roewer, 1949</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC456E6FFDDCD40590CFDB84EACFDD8" authority="Lawrence, 1959" authorityName="Lawrence" authorityYear="1959" box="[1092,1432,566,593]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD40590CFDB84FECFDD9" box="[1092,1240,567,592]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
<bibRefCitation id="2F555094FFDDCD4059A8FDB84EACFDD8" author="Lawrence, R. F." box="[1248,1432,566,593]" pageId="2" pageNumber="327" pagination="1 - 121" refId="ref9004" refString="Lawrence, R. F. (1959) Arachnides-Opilions. Faune de Madagascar, Publications de L'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 9, 1 - 121." type="journal article" year="1959">Lawrence, 1959</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="8CC456E6FFDDCD405D8EFDD44AD9FDFC" authority="Lawrence, 1959" authorityName="Lawrence" authorityYear="1959" box="[198,493,602,629]" class="Arachnida" family="Biantidae" genus="Anaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405D8EFDD44A1BFDFC" box="[198,303,603,629]" italics="true" pageId="2" pageNumber="327">Anaceros</emphasis>
<bibRefCitation id="2F555094FFDDCD405C7EFDD44AD9FDFC" author="Lawrence, R. F." box="[310,493,602,629]" pageId="2" pageNumber="327" pagination="1 - 121" refId="ref9004" refString="Lawrence, R. F. (1959) Arachnides-Opilions. Faune de Madagascar, Publications de L'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 9, 1 - 121." type="journal article" year="1959">Lawrence, 1959</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="8CC456E6FFDDCD405F23FDD449F3FDFC" authorityName="Roewer" authorityYear="1949" box="[619,711,603,629]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405F23FDD449F3FDFC" box="[619,711,603,629]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
can be separated from those genera by a male pedipalp ventrally concave (
<emphasis id="79B0F177FFDDCD405C4BFDF04A12FD11" box="[259,294,639,664]" italics="true" pageId="2" pageNumber="327">i.e.</emphasis>
, strongly arched). The male genital morphology is still unknown to
<taxonomicName id="8CC456E6FFDDCD405968FDF04FA1FD11" authorityName="Roewer" authorityYear="1949" box="[1056,1173,639,664]" class="Arachnida" family="Biantidae" genus="Hirstienus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405968FDF04FA1FD11" box="[1056,1173,639,664]" italics="true" pageId="2" pageNumber="327">Hirstienus</emphasis>
</taxonomicName>
and
<taxonomicName id="8CC456E6FFDDCD405983FDF04E00FD10" authorityName="Lawrence" authorityYear="1959" box="[1227,1332,639,665]" class="Arachnida" family="Biantidae" genus="Anaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405983FDF04E00FD10" box="[1227,1332,639,665]" italics="true" pageId="2" pageNumber="327">Anaceros</emphasis>
</taxonomicName>
avoiding comparisons, but it is known to one undetermined species of
<taxonomicName id="8CC456E6FFDDCD405E29FD2C48C1FD35" authorityName="Lawrence" authorityYear="1959" box="[865,1013,675,700]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405E29FD2C48C1FD35" box="[865,1013,675,700]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
</taxonomicName>
(see
<bibRefCitation id="2F555094FFDDCD40597EFD2C4EA5FD34" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[1078,1425,674,701]" pageId="2" pageNumber="327" refId="ref8940" refString="Kury, A. B. &amp; Perez-Gonzalez, A. (2015) A companion to Part 2 of the World Checklist of Opiliones species (Arachnida): Laniatores - Samooidea, Zalmoxoidea and Grassatores incertae sedis. Biodiversity Data Journal, 3, e 6663. https: // doi. org / 10.3897 / BDJ. 3. e 6663" type="journal volume" year="2015">Kury &amp; Pérez-González 2015</bibRefCitation>
). In this sense,
<taxonomicName id="8CC456E6FFDDCD405C7CFD484AA4FD68" authorityName="Roewer" authorityYear="1949" box="[308,400,711,737]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405C7CFD484AA4FD68" box="[308,400,711,737]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
can separate from
<taxonomicName id="8CC456E6FFDDCD405F22FD4849CAFD69" authorityName="Lawrence" authorityYear="1959" box="[618,766,711,736]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405F22FD4849CAFD69" box="[618,766,711,736]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
</taxonomicName>
by the presence of a longitudinal row of macrosetae at the sides of the
<emphasis id="79B0F177FFDDCD405C6EFD644A87FC8C" box="[294,435,747,773]" italics="true" pageId="2" pageNumber="327">pars distalis</emphasis>
(
<figureCitation id="D3FF31E0FFDDCD405C8DFD644972FC8C" box="[453,582,747,773]" captionStart="FIGURE 4" captionStartId="6.[151,250,1252,1276]" captionTargetBox="[151,1435,417,1229]" captionTargetId="figure-128@6.[151,1435,417,1229]" captionTargetPageId="6" captionText="FIGURE 4. Tetebius latibunus Roewer, 1949, holotype minor male (SMF 9909102, formerly RII/1902/230), everted penis drawing: A–B. Dorsal view; C. Lateral view; D. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735102" httpUri="https://zenodo.org/record/14735102/files/figure.png" pageId="2" pageNumber="327">Figs 4B–D</figureCitation>
;
<figureCitation id="D3FF31E0FFDDCD405F1DFD6449AFFC8C" box="[597,667,747,773]" captionStart="FIGURE 9" captionStartId="11.[151,250,1763,1787]" captionTargetBox="[151,1435,454,1740]" captionTargetId="figure-152@11.[151,1435,453,1740]" captionTargetPageId="11" captionText="FIGURE 9. Tetebius latibunus Roewer, 1949, major males. A–E. Penis drawings (CASENT 9069465): A, B. Dorsal view; C. Lateral view; D. Ventral view; E. Expanded penis, lateral view; F–H. Scanning electron micrographs of penis (MACN-Ar 45346): F. Dorsal view; G. Lateral view; H. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: A = 500 µm; B–H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735116" httpUri="https://zenodo.org/record/14735116/files/figure.png" pageId="2" pageNumber="327">9B–H</figureCitation>
), which is more inflated. Also,
<taxonomicName id="8CC456E6FFDDCD405942FD644F52FC8C" authorityName="Roewer" authorityYear="1949" box="[1034,1126,747,773]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405942FD644F52FC8C" box="[1034,1126,747,773]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="8CC456E6FFDDCD405DDFFC804A1FFCA1" authorityName="Lawrence" authorityYear="1959" box="[151,299,783,808]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405DDFFC804A1FFCA1" box="[151,299,783,808]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
</taxonomicName>
by the presence of a pair of setae on the base of the
<emphasis id="79B0F177FFDDCD405EC2FC804F0EFCA0" box="[906,1082,783,809]" italics="true" pageId="2" pageNumber="327">lamina apicalis</emphasis>
(
<figureCitation id="D3FF31E0FFDDCD405902FC804FFEFCA0" box="[1098,1226,783,809]" captionStart="FIGURE 4" captionStartId="6.[151,250,1252,1276]" captionTargetBox="[151,1435,417,1229]" captionTargetId="figure-128@6.[151,1435,417,1229]" captionTargetPageId="6" captionText="FIGURE 4. Tetebius latibunus Roewer, 1949, holotype minor male (SMF 9909102, formerly RII/1902/230), everted penis drawing: A–B. Dorsal view; C. Lateral view; D. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735102" httpUri="https://zenodo.org/record/14735102/files/figure.png" pageId="2" pageNumber="327">Figs 4B–D</figureCitation>
;
<figureCitation id="D3FF31E0FFDDCD40599FFC804E2AFCA0" box="[1239,1310,783,809]" captionStart="FIGURE 9" captionStartId="11.[151,250,1763,1787]" captionTargetBox="[151,1435,454,1740]" captionTargetId="figure-152@11.[151,1435,453,1740]" captionTargetPageId="11" captionText="FIGURE 9. Tetebius latibunus Roewer, 1949, major males. A–E. Penis drawings (CASENT 9069465): A, B. Dorsal view; C. Lateral view; D. Ventral view; E. Expanded penis, lateral view; F–H. Scanning electron micrographs of penis (MACN-Ar 45346): F. Dorsal view; G. Lateral view; H. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: A = 500 µm; B–H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735116" httpUri="https://zenodo.org/record/14735116/files/figure.png" pageId="2" pageNumber="327">9B–H</figureCitation>
). Lastly, in
<taxonomicName id="8CC456E6FFDDCD405DDFFCBC4BC7FCC4" authorityName="Roewer" authorityYear="1949" box="[151,243,819,845]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405DDFFCBC4BC7FCC4" box="[151,243,819,845]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
, conductors are distally thin, apically pointed, and not as wide as in
<taxonomicName id="8CC456E6FFDDCD405EBAFCBC4FB2FCC5" authorityName="Lawrence" authorityYear="1959" box="[1010,1158,819,844]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405EBAFCBC4FB2FCC5" box="[1010,1158,819,844]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
</taxonomicName>
(
<figureCitation id="D3FF31E0FFDDCD4059DDFCBC4E26FCC4" box="[1173,1298,819,845]" captionStart="FIGURE 4" captionStartId="6.[151,250,1252,1276]" captionTargetBox="[151,1435,417,1229]" captionTargetId="figure-128@6.[151,1435,417,1229]" captionTargetPageId="6" captionText="FIGURE 4. Tetebius latibunus Roewer, 1949, holotype minor male (SMF 9909102, formerly RII/1902/230), everted penis drawing: A–B. Dorsal view; C. Lateral view; D. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735102" httpUri="https://zenodo.org/record/14735102/files/figure.png" pageId="2" pageNumber="327">Figs 4B–C</figureCitation>
;
<figureCitation id="D3FF31E0FFDDCD405856FCBC4EA8FCC4" box="[1310,1436,819,845]" captionStart="FIGURE 9" captionStartId="11.[151,250,1763,1787]" captionTargetBox="[151,1435,454,1740]" captionTargetId="figure-152@11.[151,1435,453,1740]" captionTargetPageId="11" captionText="FIGURE 9. Tetebius latibunus Roewer, 1949, major males. A–E. Penis drawings (CASENT 9069465): A, B. Dorsal view; C. Lateral view; D. Ventral view; E. Expanded penis, lateral view; F–H. Scanning electron micrographs of penis (MACN-Ar 45346): F. Dorsal view; G. Lateral view; H. Ventral view. Abbreviations: PB, pars basalis; PD, pars distalis. Stylus in green, conductors in red, titillators in magenta. Scale bars: A = 500 µm; B–H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14735116" httpUri="https://zenodo.org/record/14735116/files/figure.png" pageId="2" pageNumber="327">9B–C, E–F</figureCitation>
<emphasis id="79B0F177FFDDCD405DDFFCD84B82FCF9" box="[151,182,855,880]" italics="true" pageId="2" pageNumber="327">vs.</emphasis>
fig.
<quantity id="8C3C8080FFDDCD405DA2FCD84A21FCF8" box="[234,277,855,881]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="327" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="2F555094FFDDCD405C53FCD8495AFCF8" author="Kury, A. B. &amp; Perez-Gonzalez, A." box="[283,622,854,881]" pageId="2" pageNumber="327" refId="ref8940" refString="Kury, A. B. &amp; Perez-Gonzalez, A. (2015) A companion to Part 2 of the World Checklist of Opiliones species (Arachnida): Laniatores - Samooidea, Zalmoxoidea and Grassatores incertae sedis. Biodiversity Data Journal, 3, e 6663. https: // doi. org / 10.3897 / BDJ. 3. e 6663" type="journal volume" year="2015">Kury &amp; Pérez-González 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03DE7EEEFFDDCD405D8FFCF44929FC54" pageId="2" pageNumber="327" type="discussion">
<paragraph id="4B7B2D65FFDDCD405D8FFCF44929FC54" blockId="2.[151,1437,494,1061]" pageId="2" pageNumber="327">
<emphasis id="79B0F177FFDDCD405D8FFCF44A04FC1C" bold="true" box="[199,304,891,917]" pageId="2" pageNumber="327">Remark.</emphasis>
The limits among
<taxonomicName id="8CC456E6FFDDCD405CB7FCF4495CFC1C" authorityName="Lawrence" authorityYear="1959" box="[511,616,891,917]" class="Arachnida" family="Biantidae" genus="Anaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405CB7FCF4495CFC1C" box="[511,616,891,917]" italics="true" pageId="2" pageNumber="327">Anaceros</emphasis>
</taxonomicName>
,
<taxonomicName id="8CC456E6FFDDCD405F3CFCF4483CFC1D" authorityName="Lawrence" authorityYear="1959" box="[628,776,891,916]" class="Arachnida" family="Biantidae" genus="Hovanoceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405F3CFCF4483CFC1D" box="[628,776,891,916]" italics="true" pageId="2" pageNumber="327">Hovanoceros</emphasis>
</taxonomicName>
,
<taxonomicName id="8CC456E6FFDDCD405E5DFCF448BEFC1D" authorityName="Roewer" authorityYear="1949" box="[789,906,891,916]" class="Arachnida" family="Biantidae" genus="Hirstienus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405E5DFCF448BEFC1D" box="[789,906,891,916]" italics="true" pageId="2" pageNumber="327">Hirstienus</emphasis>
</taxonomicName>
, and
<taxonomicName id="8CC456E6FFDDCD405E8DFCF44F15FC1C" authorityName="Roewer" authorityYear="1949" box="[965,1057,891,917]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405E8DFCF44F15FC1C" box="[965,1057,891,917]" italics="true" pageId="2" pageNumber="327">Tetebius</emphasis>
</taxonomicName>
are not well defined, and the male genitalia morphology of
<taxonomicName id="8CC456E6FFDDCD405CFDFC10492AFC30" authorityName="Lawrence" authorityYear="1959" box="[437,542,927,953]" class="Arachnida" family="Biantidae" genus="Anaceros" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405CFDFC10492AFC30" box="[437,542,927,953]" italics="true" pageId="2" pageNumber="327">Anaceros</emphasis>
</taxonomicName>
and
<taxonomicName id="8CC456E6FFDDCD405F11FC1049FAFC31" authorityName="Roewer" authorityYear="1949" box="[601,718,927,952]" class="Arachnida" family="Biantidae" genus="Hirstienus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="79B0F177FFDDCD405F11FC1049FAFC31" box="[601,718,927,952]" italics="true" pageId="2" pageNumber="327">Hirstienus</emphasis>
</taxonomicName>
remains unknown. Further studies are necessary to clarify the limits and validity of those genera.
</paragraph>
</subSubSection>
<subSubSection id="03DE7EEEFFDDCD405D8FFC68492FFBAC" pageId="2" pageNumber="327" type="type_taxon">
<paragraph id="4B7B2D65FFDDCD405D8FFC6848CFFB88" blockId="2.[151,1437,494,1061]" box="[199,1019,998,1025]" pageId="2" pageNumber="327">
<emphasis id="79B0F177FFDDCD405D8FFC684A56FB89" bold="true" box="[199,354,998,1025]" pageId="2" pageNumber="327">
<typeStatus id="947F93C7FFDDCD405D8FFC684A36FB88" box="[199,258,999,1025]" pageId="2" pageNumber="327">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8CC456E6FFDDCD405C21FC6849ECFB88" ID-CoL="7X259" authority="Roewer, 1949" authorityName="Roewer" authorityYear="1949" box="[361,728,999,1025]" class="Arachnida" family="Biantidae" genus="Tetebius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="latibunus">
<emphasis id="79B0F177FFDDCD405C21FC684901FB88" box="[361,565,999,1025]" italics="true" pageId="2" pageNumber="327">Tetebius latibunus</emphasis>
<bibRefCitation id="2F555094FFDDCD405F74FC6849ECFB88" author="Roewer, C. F." box="[572,728,999,1025]" pageId="2" pageNumber="327" pagination="247 - 289" refId="ref9357" refString="Roewer, C. F. (1949) Uber Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30 (4), 247 - 289." type="journal article" year="1949">Roewer, 1949</bibRefCitation>
</taxonomicName>
(by original designation).
</paragraph>
<paragraph id="4B7B2D65FFDDCD405D8FFB84492FFBAC" blockId="2.[151,1437,494,1061]" box="[199,539,1034,1061]" pageId="2" pageNumber="327">
<emphasis id="79B0F177FFDDCD405D8FFB844AA6FBAD" bold="true" box="[199,402,1034,1061]" pageId="2" pageNumber="327">Included species.</emphasis>
Monotypic.
</paragraph>
</subSubSection>
</treatment>
</document>

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