diff --git a/data/C3/6D/9C/C36D9C73FFDDCD405DDFFE8D492FFBAC.xml b/data/C3/6D/9C/C36D9C73FFDDCD405DDFFE8D492FFBAC.xml new file mode 100644 index 00000000000..93050b83377 --- /dev/null +++ b/data/C3/6D/9C/C36D9C73FFDDCD405DDFFE8D492FFBAC.xml @@ -0,0 +1,266 @@ + + + +Redescription of Tetebius latibunus as a new senior synonym of Malgaceros boviceps (Opiliones: Laniatores: Biantidae) + + + +Author + +Mamani, Vanesa +División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina. & Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Av. Int. Güiraldes s / n, Ciudad Universitaria, C 1428 EHA, Buenos Aires, Argentina. + + + +Author + +Pérez-González, Abel +0000-0002-4245-3302 +División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina. +abelaracno@gmail.com + +text + + +Zootaxa + + +2025 + +2025-01-17 + + +5570 + + +2 + + +325 +343 + + + + +https://doi.org/10.11646/zootaxa.5570.2.5 + +journal article +10.11646/zootaxa.5570.2.5 +1175-5326 +14735094 +2C8267DE-C36B-4516-AD18-7A35810DF5B1 + + + + + + + +Tetebius +Roewer, 1949 + + + + + + +urn:lsid:zoobank.org:act: +DE3F95EB-0A29-45C8-BB16-6D4D5AE4A7F6 + + + + + + + + +Tetebius +Roewer 1949: 288 + + +; + +Staręga 1992: 330 + +; + +Kury & Pérez-González 2015: 7 + +. + + + + + + +Malgaceros +Lawrence 1959: 79 + + +; + +Staręga 1992: 330 + +; + +Kury & Pérez-González 2015:6 + +[ +type +species: + +Malgaceros boviceps +Lawrence, 1959 + +, by original designation]. +New synonymy. + + + + + +Diagnosis. + +Tetebius + +can be distinguished from most genera of +Biantinae +by the absence of the typical biantid pedipalp shape with a thin and generally unarmed femur ( +e.g. +, +Martens 1978 +, fig. 2). By contrast, + +Tetebius + +exhibits a robust, curved, and armed femur, a feature shared with + +Hirstienus +Roewer, 1949 + +, + +Hovanoceros +Lawrence, 1959 + +, and + +Anaceros +Lawrence, 1959 + +. However, + +Tetebius + +can be separated from those genera by a male pedipalp ventrally concave ( +i.e. +, strongly arched). The male genital morphology is still unknown to + +Hirstienus + +and + +Anaceros + +avoiding comparisons, but it is known to one undetermined species of + +Hovanoceros + +(see +Kury & Pérez-González 2015 +). In this sense, + +Tetebius + +can separate from + +Hovanoceros + +by the presence of a longitudinal row of macrosetae at the sides of the +pars distalis +( +Figs 4B–D +; +9B–H +), which is more inflated. Also, + +Tetebius + +can be differentiated from + +Hovanoceros + +by the presence of a pair of setae on the base of the +lamina apicalis +( +Figs 4B–D +; +9B–H +). Lastly, in + +Tetebius + +, conductors are distally thin, apically pointed, and not as wide as in + +Hovanoceros + +( +Figs 4B–C +; +9B–C, E–F +vs. +fig. +2 in +Kury & Pérez-González 2015 +). + + + + +Remark. +The limits among + +Anaceros + +, + +Hovanoceros + +, + +Hirstienus + +, and + +Tetebius + +are not well defined, and the male genitalia morphology of + +Anaceros + +and + +Hirstienus + +remains unknown. Further studies are necessary to clarify the limits and validity of those genera. + + + + + +Type +species. + + +Tetebius latibunus +Roewer, 1949 + +(by original designation). + + +Included species. +Monotypic. + + + + \ No newline at end of file diff --git a/data/C3/6D/9C/C36D9C73FFDDCD4F5DDFFBF9499DFEC8.xml b/data/C3/6D/9C/C36D9C73FFDDCD4F5DDFFBF9499DFEC8.xml new file mode 100644 index 00000000000..d35ff0f6598 --- /dev/null +++ b/data/C3/6D/9C/C36D9C73FFDDCD4F5DDFFBF9499DFEC8.xml @@ -0,0 +1,1038 @@ + + + +Redescription of Tetebius latibunus as a new senior synonym of Malgaceros boviceps (Opiliones: Laniatores: Biantidae) + + + +Author + +Mamani, Vanesa +División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina. & Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Av. Int. Güiraldes s / n, Ciudad Universitaria, C 1428 EHA, Buenos Aires, Argentina. + + + +Author + +Pérez-González, Abel +0000-0002-4245-3302 +División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina. +abelaracno@gmail.com + +text + + +Zootaxa + + +2025 + +2025-01-17 + + +5570 + + +2 + + +325 +343 + + + + +https://doi.org/10.11646/zootaxa.5570.2.5 + +journal article +10.11646/zootaxa.5570.2.5 +1175-5326 +14735094 +2C8267DE-C36B-4516-AD18-7A35810DF5B1 + + + + + + + +Tetebius latibunus +Roewer, 1949 + + + + + + +urn:lsid:zoobank.org:act: +B8D24F87-16C6-40C1-B7F6-3486C4EF5741 + + + + + +( +Figs 1 +; +3–10 +, Table 1) + + + + + + + +Tetebius latibunus +Roewer 1949: 288 + + +, Taf. 12, fig. 98 a–e; + +Staręga 1992: 330 + +; + +Kury & Pérez-González 2015: 7 + +, fig. 3. + + + + +Malgaceros boviceps +Lawrence 1959: 80–82 + + +, fig. 27; + +Staręga 1992: 330 + +; + +Kury & Pérez-González 2015: 6–7 + +. +New synonymy + + + + + +Type material. + +Tetebius latibunus + +, + +♂ +holotype +( +SMF 9909102 +, previously cataloged as RII/1902/230, examined), +MOZAMBIQUE +, +Teté +( +Fig. 2 +) (spurious locality see further considerations) + +. + + + +Malgaceros boviceps + +, + +3 ♂ +, +1 ♀ +syntypes +( +MNHN +, destroyed), +MADAGASCAR +, +Sambirano +, +Nosy Be +, +Lokobe +, + +Oct. 1947 + +, +J. Millot +, under stones + +. + + + + +Remark +: the +syntypes +of + +Malgaceros boviceps + +were destroyed in the tragic fire that occurred in the MNRJ in 2018 (A.B. Kury pers. comm.). + + +Other material examined. + +2 major +♂ +(1 photo voucher), 3 minor +♂ +(1 photo voucher) (CASENT 9069465), +MADAGASCAR +, +Antsiranana +, Nosy Be, Réserve Naturelle Intégrale de Lokobe, +6.3 km +112° ESE Hellville, - 13.41944°; 48.33111°, + +30 m +a.s.l. + +, rainforest, EC19 sifted +Litter +, + +19–24 Mar 2001 + +, coll. +B. L. Fisher +et al. +, collection code: BLF3422. + + +1 major +♂ +, 1 minor +♂ +(MACN-Ar 45417) with the same data as for preceding. + + +1 major +♂ +( +SEM +voucher) (MACN-Ar 45346) with the same data as for preceding. + + +2 minor +♂ +, +9 ♀ +(1 photo voucher) ( +CASENT 9069463 +), +MADAGASCAR +, +Antsiranana +, +Nosy Be +, +Réserve Naturelle Intégrale de Lokobe +, + +6.3 km +112° ESE Hellville + +, -13.41944°; 48.33111°, + +30 m +a.s.l. + +, rainforest, EC19 sifted +Litter +, + +19–24 Mar 2001 + +, coll. +B. L. Fisher +et al. +, collection code: BLF3422. + + +1 ♀ +(MACN-Ar 45361) with the same data as for preceding. + + +1 minor +♂ +( +CASENT 9069464 +), +MADAGASCAR +, +Antsiranana +, +Nosy Be +, +Réserve Naturelle Intégrale de Lokobe +, + +6.3 km +112° ESE Hellville + +, -13.41944°; 48.33111°, +30 m +a.s.l., rainforest, EC27 pitfall trap, +19–24 Mar 2001 +, coll. B. L. Fisher +et al. +, collection code: BLF3418 + +. + + + + + +Redescription. Minor male +holotype +(SMF 9909102, formerly RII/1902/230) + +. Body measurements: Total body length 1.46, carapace length 0.50, +scutum magnum +length 1.15, carapace maximum width 0.86, abdominal +scutum +maximum width 0.99. Appendage measurements in Table 1. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
TrFePaTiMtTa +T +
Pedipalp +0.17 + +0.81 + +0.43 + +0.61 + +- + +0.34 + +2.36 +
+Minor ♂ +Leg I +0.17 + +0.63 + +0.32 + +0.42 + +0.64 + +0.47 + +2.65 +
+SMF +Leg II +0.23 + +0.90 + +0.42 + +0.71 + +0.98 + +0.76 + +4.00 +
+9909102 +Leg III +0.19 + +0.68 + +0.29 + +0.53 + +0.73 + +0.51 + +2.93 +
Leg IV +0.21 + +0.83 + +0.36 + +0.69 + +0.96 + +0.64 + +3.69 +
Pedipalp0.251.050.620.74-0.40 +3.06 +
Major ♂Leg I0.200.870.340.530.800.50 +3.24 +
CASENTLeg II0.251.200.540.941.230.88 +5.04 +
9069465Leg III0.200.850.300.640.910.61 +3.51 +
Leg IV0.261.010.390.821.180.78 +4.44 +
Pedipalp0.190.720.440.51-0.31 +2.17 +
♀Leg I0.160.650.270.430.630.46 +2.60 +
CASENTLeg II0.220.870.410.710.870.80 +3.88 +
9069463Leg III0.190.660.240.530.720.53 +2.87 +
Leg IV0.240.850.350.730.970.653.79
+ + +Dorsum +: Outline hourglass-shaped with Eta (η) shape, with a slightly accentuated constriction posterior to eye level ( +Figs 1A +; +3A +). Carapace coarsely granulated, wider than long ( +Figs 1A +; +3A +), with a marked and rounded frontal hump ( +Fig. 1C +); anterior border of carapace with three granules on each side ( +Fig. 3A +). Cheliceral sockets not marked ( +Figs 1A +; +3A +). Eyes separated and located on the sides of a wide and low +ocularium +( +Figs 1A, C +; +3A +); +ocularium +shorter than the height of the abdominal +scutum +and occupies the posterior region of the carapace up to the frontal hump ( +Fig. 1C +). Sulcus I deep and well-marked, in dorsal view curved to anterior body region ( +Figs 1A +; +3A +). Mesotergal areas I–IV granulated and not completely defined; sulci II–IV shallow and limited to the medial region ( +Figs 1A +; +3A +). Mesotergal area I with larger medial granules ( +Figs 1A +; +3A +); mesotergal area II with a medial row of larger granules and one medial tubercle; mesotergal areas III–IV with two medial tubercles; tubercles of mesotergal area IV more separated from each other than tubercles of mesotergal area III ( +Figs 1A +; +3A +). Mesotergal area V granulated with one medial tubercle ( +Figs 1A +; +3A +). Lateral margins of +scutum magnum +with rows of granules ( +Fig. 3A +). Free tergites granulated; free tergites I–III with a posterior row of larger granules and three medial setiferous tubercles ( +Fig. 3A +). + + +Venter +: Without any conspicuous armature ( +Fig. 1B +); coxa I with setiferous granules; anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively ( +Fig. 1B +); coxa IV with lateral margin granulated ( +Fig. 1B +); anal operculum with several conspicuous granules ( +Fig. 1B +). Spiracles concealed by a bridge-like projection from distal coxa IV connecting to the posterior margin of the stigmatic area ( +Fig. 1B +). + + +Chelicerae +: Basichelicerite with a marked +bulla +and two dorsal setiferous granules on the proximal margin ( +Fig. 3A, E +). Cheliceral hand with an ectal concavity, sparse setae and three frontal rounded setiferous tubercles ( +Fig. 3E–F +). Fixed finger proximally with one large triangular tooth followed by smaller teeth ( +Fig. 3F +). + + +Pedipalps +: Coxa proximally with two dorsomesal granules and one dorsoectal granule ( +Fig. 3A +). Femur curved; ventral surface with two large proximal spines, medially with one small mesal spine and one small ectal setiferous tubercle ( +Fig. 3B–D +). Patella distally with a small mesal spine ( +Fig. 3B +). Tibia ventrally with two short mesal spines and three large ectal spines ( +Fig. 3B–C +). Tarsus shorter than tibia; ventral surface with two ectal spines and two mesal spines ( +Fig. 3B–C +). + + +Legs +: Tarsi III and IV with a dense scopula. Tarsal formula: 3(2):4(3):5:5. + + +Color (specimen preserved in 80% ethanol) +: Body completely light yellow ( +Fig. 1A–C +). + + + +FIGURE 1. + +Tetebius latibunus +Roewer, 1949 + +, holotype minor male (SMF 9909102, formerly RII/1902/230), photos of habitus: A. Dorsal view; B. Ventral view; C. Lateral view. Scale bars: 500 µm. + + + + +FIGURE 2. +Label for holotype minor male of + +Tetebius latibunus +Roewer, 1949 + +(SMF 9909102, formerly RII/1902/230). + + + + +FIGURE 3. + +Tetebius latibunus +Roewer, 1949 + +, holotype minor male (SMF 9909102, formerly RII/1902/230), drawings of habitus, pedipalp and chelicera: A. Habitus, dorsal view; B–D. Left pedipalp: B. Mesal view; C. Ectal view; D. Femur, ventral view; E–F. Left chelicera: E. Ectal view; F. Frontal view. Scale bars: A–D = 500 µm; E–F = 200 µm. + + + +Genitalia +: Penis with distinguishable limits between +pars basalis +and +pars distalis +( +Fig. 4A +). +Pars basalis +tubular, +pars distalis +slightly swollen at the glans penis level ( +Fig. 4A +). Apical edge laminar ( +i.e. +, dorsoventrally flat) with a well-defined basal constriction and a broadened, rounded, and ventrally concave edge forming a +lamina apicalis +( +Fig. 4A–D +). +Pars distalis +with 17 spatula-shaped macrosetae: basally with a lateral row of five (right) and six (left) macrosetae ( +Fig. 4A, C +), +lamina apicalis +with one basal pair of macrosetae and two distal pairs of macrosetae ( +Fig. 4B–D +). + +Capsula externa + +in the everted state inflated with two elongated and curl apically titillators covered by digitiform projections on the inner surface ( +Fig. 4B–C +). + +Capsula interna + +formed by two laminar conductors broadened ventrally and apically pointed, and one laminar style apically with a pointed ventral projection ( +Fig. 4C +). + + + +FIGURE 4. + +Tetebius latibunus +Roewer, 1949 + +, holotype minor male (SMF 9909102, formerly RII/1902/230), everted penis drawing: A–B. Dorsal view; C. Lateral view; D. Ventral view. Abbreviations: PB, +pars basalis +; PD, +pars distalis +. Stylus in green, conductors in red, titillators in magenta. Scale bars: 100 µm. + + + +Major male (CASENT 9069465). +Body measurements: Total body length 1.57, carapace length 0.62, +scutum magnum +length 1.31, carapace maximum width 1.00, abdominal +scutum +maximum width 1.11. Appendage measurements in Table 1. + + +Dorsum +: Outline hourglass-shaped with Eta (η) shape, with a slightly accentuated constriction posterior to eye level ( +Figs 5A +; +6A +). Carapace coarsely granulated, wider than long; frontal hump well-marked, rounded, and granulated; anterior border straight with two granules on each lateral margin ( +Fig. 6A +). Cheliceral sockets not marked ( +Figs 5A +; +6A +). Eyes separated, located at the base of a wide and high +ocularium +, which exceeds the height of the abdominal +scutum +( +Figs 5C +; +6D +); +ocularium +occupies the posterior region of the carapace up to the area behind the frontal hump ( +Figs 5A, C +; +6A, D +). Mesotergal areas coarsely granulated ( +Fig. 6A, D +). Mesotergal areas I–IV not well defined; sulci II–IV shallow and limited to the medial region ( +Figs 5A, C +; +6A, D +). Mesotergal area I with two medial setiferous granules; mesotergal area II with a medial row of setiferous granules and one medial setiferous tubercle; mesotergal areas III–IV with a row of setiferous granules and two medial setiferous tubercles; tubercles in area IV more separated and slightly larger than those in area III ( +Figs 5A +; +6A, D +). Mesotergal area V with a posterior row of setiferous granules and a medial setiferous tubercle ( +Fig. 6A, D +). Lateral margins of abdominal +scutum +with irregular longitudinal rows of granules ( +Fig. 6A, D +). Ozopore with an oval and narrow orifice with a descending channel that extends toward the posterior region ( +Fig. 6D–E +). Free tergites granulated, with a posterior row of setiferous granules and three medial setiferous tubercles ( +Fig. 6A, D +). + + + +FIGURE 5. + +Tetebius latibunus +Roewer, 1949 + +, major male (CASENT 9069465), photos of habitus: A. Dorsal view; B. Ventral view; C. Lateral view. Scale bars: 500 µm. + + + + +FIGURE 6. + +Tetebius latibunus +Roewer, 1949 + +, major male (MACN-Ar 45346), scanning electron micrographs of habitus: A. Dorsal view; B. Ventral view; C. Detail of sternum and medial region of coxae I–IV; D. Lateral view; E. Detail of ozopore. Scale bars: 200 µm. + + + +Venter +: Coxae I–II with setiferous granules ( +Figs 5B +; +6B–C +); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively; posterior border of the spiracular area and free sternites I–V with a row of setiferous granules; anal operculum with conspicuous setiferous granules ( +Fig. 6B, D +). Spiracles concealed by a bridge-like projection from distal coxa IV connecting to the posterior margin of the stigmatic area ( +Fig. 6B–C +). + + +Chelicerae +: Basichelicerite with an elongated +bulla +and three dorsal setiferous granules on the proximal margin ( +Figs 6A +; +7E–F +). Cheliceral hand with an ectal concavity, sparse frontal setae, prominent proximal rounded setiferous granules, and three distal setiferous tubercles: two large frontal and one smaller ectal ( +Fig. 7E–G +). Fixed finger with a protuberance; movable finger with two prominent teeth continued by small teeth ( +Fig. 7F–G +). + + +Pedipalps +: Coxa dorsoproximally with one ectal and two mesal tubercles; ventral surface with three small tubercles ( +Fig. 6A–B +). Trochanter with one dorsal tubercle and one ventral setiferous granule ( +Fig. 7A +). Femur strongly curved; ventrally with two large proximal spines: proximal spine with a rectangular socket and long seta; distal spine with a curved and pointed socket and short seta ( +Fig. 7A–B +); ventromedially with one small and rounded mesal spine and one ectal rounded granule ( +Fig. 7A–B +). Patella distally with a small spine ( +Fig. 7A +). Tibia ventrally with two short mesal spines: medial with conical socket and long subapical seta, distal with short socket and apical seta ( +Fig. 7A +); three large ectal spines: proximal with thickened and short socket; medial and distal with a narrow and elongated socket and long subapical seta ( +Fig. 7B +). Tarsus shorter than tibia, ventrally with two ectal and mesal spines, proximal spines with longer sockets ( +Fig. 7A–B +). Spines distally with sparse microtrichia ( +Fig. 7C–D +). + + + +FIGURE 7. + +Tetebius latibunus +Roewer, 1949 + +, major male (MACN-Ar 45346), scanning electron micrographs of pedipalp and chelicera: A–D. Left pedipalp: A. Mesal view; B. Ectal view; C. Detail of ventroproximal femoral spine with microtrichia; D. Detail of ectomedial tibial spine with microtrichia; E–G. Left chelicera: E. Mesal view; F. Ectal view; G. Frontal view. Scale bars: A–B, E–G = 200 µm; C–D = 20 µm. + + + +Legs +: unarmed ( +Fig. 8A–D +). Tarsi III–IV with a dense scopula ( +Fig. 8E–F +). Tarsal formula: 3(2):4(3):5:5. +Color (specimen preserved in 80% ethanol) +: Body yellowish-brown, with dark brown reticulations on pedipalps, chelicerae, legs, and carapace ( +Fig. 5A–B +). Mesotergal areas, free tergites, and free sternites dark brown ( +Fig. 5 A–C +). + + + +FIGURE 8. + +Tetebius latibunus +Roewer, 1949 + +, major male (MACN-Ar 45346), scanning electron micrographs of left legs: A. Leg I, prolateral view: B. Leg II, prolateral view; C. Leg III, prolateral view; D. Leg IV, prolateral view; E. Scopula of leg III, prolateral view; F. Scopula of leg IV, prolateral view. Scale bars: A = 200 µm; B–D = 500 µm; E–F = 50 µm. + + + +Genitalia +: Penis with distinguishable limits between +pars basalis +and +pars distalis +( +Fig. 9A +). +Pars basalis +tubular; +pars distalis +slightly swollen ( +Fig. 9A–B, D, F, H +). Apical edge laminar ( +i.e. +, flattened dorsoventrally) with a well-defined basal constriction and a broadened, rounded, and ventrally concave edge forming a +lamina apicalis +( +Fig. 9A–D, F–H +). +Pars distalis +with 18 spatula-shaped macrosetae: basally with a lateral row of six macrosetae ( +Fig. 9B, D, F, H +), +lamina apicalis +with one basal pair of macrosetae and two subapical pairs of macrosetae ( +Fig. 9B–H +). + +Capsula externa + +with two titillators covered by digitiform projections on the inner surface ( +Fig. 9A–B, E–H +). + +Capsula interna + +formed by two laminar conductors broadened ventrally and apically pointed and one laminar style apically with a pointed ventral projection ( +Fig. 9B, E, F +). + + + +FIGURE 9. + +Tetebius latibunus +Roewer, 1949 + +, major males. A–E. Penis drawings (CASENT 9069465): A, B. Dorsal view; C. Lateral view; D. Ventral view; E. Expanded penis, lateral view; F–H. Scanning electron micrographs of penis (MACN-Ar 45346): F. Dorsal view; G. Lateral view; H. Ventral view. Abbreviations: PB, +pars basalis +; PD, +pars distalis +. Stylus in green, conductors in red, titillators in magenta. Scale bars: A = 500 µm; B–H = 100 µm. + + + + +FIGURE 10. + +Tetebius latibunus +Roewer, 1949 + +, sexual dimorphism, photos of habitus. A, E. Female (CASENT 9069463), habitus: A. Dorsal view, E. Lateral view; B, F. Holotype minor male (SMF 9909102, formerly RII/1902/230), habitus: B. Dorsal view, F. Lateral view; C, G. Minor male (CASENT 9069465), habitus: C. Dorsal view; G. Lateral view; D, H. Major male (CASENT 9069465), habitus: D. Dorsal view, H. Lateral view. Scale bars: A–D = 500 µm; E–H = 1 mm. + + + +Female (CASENT 9069463). +Body measurements: Total body length 1.55, carapace length 0.51, +scutum magnum +length 1.15, maximum carapace width 0.83, maximum abdominal +scutum +width 1.03. Appendage measurements in Table 1. Female resembles both minor and major males in the armature of the +scutum magnum +but with lower tubercles on the abdominal +scutum +and free tergites. Female differs from males by having less development chelicerae ( +Fig. 10A +vs. +Fig. 10B–D +), and smaller +ocularium +( +Fig. 10E +vs. +Fig. 10F–H +). Additionally, female differs by having thinner pedipalps with a less pronounced curvature in the femur ( +Fig. 10E +vs. +Fig. 10F–H +). Tarsal formula 3(2):4(3):5:5. + + + + +Distribution +: Lokobe Strict Nature Reserve, Nosy Be, Diana, +Madagascar +( +Fig. 11 +). Spurious record in +Mozambique +, +Tete +(see further considerations). + + + + \ No newline at end of file