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3205 College Ave., Davie, FL 33314 - 7719, USA + + + +Author + +Bartlett, Charles R. +University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA + + + +Author + +Martinez Aponte, Laura V. +0009-0003-8030-0651 +laura.martinez9@upr.edu + + + +Author + +Ruiz, Alfredo Rodriguez +0000-0002-1311-1637 +arodr412@asu.edu + + + +Author + +Bloch, Melody +University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA + + + +Author + +Bahder, Brian W. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +341 +352 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.3 + +journal article +10.11646/zootaxa.5481.3.3 +1175-5326 +12758762 +798238C5-D310-4FA1-BF2D-6D365250655C + + + + + + + +Colpoptera stonei +Bahder & Bartlett + +, +sp. nov. + + + + + + +( +Figures 2–6 +) + + + + +Type locality. + +Manzanillo +, +Limón Province +, +Costa Rica +( +9.631917 +, +-84.672517 +) ( +Fig. 1 +) + +. + + + + +Etymology. +The specific epithet + +‘ +stonei + +’ is a patronym in honor of Dan and Eugene Stone, very close friends (almost older brothers) to the senior author. Their friendship, comedic relief, being fellow bandmates in Shutter and overall love and support was instrumental in his development and success in early academic years. Rock on Dan and Eugene! + + + + +Diagnosis. +Pale species, body generally uniformly testaceous, slight fuscous mottling on pronotum and mesonotum, forewing transparent, slight fuscous patch at apex, face without markings.Aedeagus lightly sclerotized, nearly symmetrical, pair of strongly bifurcated dorsal processes of moderate length arising subapically on lateral margins with ventral bifurcations emerging mesad near midline (not on outer lateral margin) and pair of shorter, curved processes on ventral margin. + + + + +Description. +Pale species, body generally uniformly testaceous, slight fuscous mottling on pronotum and mesonotum, forewing transparent, slight fuscous patch at apex, face without markings ( +Fig. 2 +). + + +Structure. +Body length with with wings; males ( +n +=4) +4.2–4.3 mm +, females ( +n= +6) 4.4–4.6 ( +Table 3 +). + + + +TABLE 3. +Biometric data for + +Colpoptera stonei + +sp. nov. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
Character +Male ( +n= +4) + +Female ( +n +=6) +
RangeAverage ± SERangeAverage±SE
Body length, with wings4.2–4.34.3±0.14.4–4.64.5±0.1
Body length, no wings2.5–2.52.5±0.02.8–2.92.9±0.1
Forewing length3.8–3.83.8±0.04.0–4.24.1±0.1
Vertex length0.1–0.10.1±0.00.1–0.10.1±0.0
Vertex width, basal margin0.3–0.30.3±0.00.3–0.30.3±0.0
Vertex width, distal margin0.3–0.30.3±0.00.3–0.30.3±0.0
Pronotum length, midline0.2–0.20.2±0.00.2–0.20.2±0.0
Mesonotum length, midline0.7–0.80.8±0.00.8–0.90.9±0.1
Mesonotum width0.8–0.80.8±0.00.8–0.80.8±0.0
Frons width, dorsal margin0.4–0.50.5±0.00.4–0.50.5±0.0
Frons width, clypeal suture0.2–0.30.3±0.00.2–0.30.3±0.0
Frons width, widest0.5–0.50.5±0.00.5–0.50.5±0.0
Frons width, narrowest0.4–0.50.5±0.00.4–0.50.5±0.0
Frons length, midline0.6–0.60.6±0.00.6–0.60.6±0.0
Clypeus length0.2–0.20.2±0.00.3–0.30.3±0.0
+ + +Head +. In dorsal view, vertex nearly rectangular, anterior margin linear, approximately level with anterior margin of eyes, posterior margin concave, median carina present, transverse carina at fastigium evident ( +Fig. 3A +). In lateral view, generally rounded, slightly angled at fastigium, vertex slightly projected above eyes, frons curved dorsad, becoming linear near ventral margin of eyes ( +Fig. 3B +). In frontal view, transverse carina at fastigium evident, lateral margins expanding from dorsal margin, reaching widest point just below antennae then constricting strongly, frontoclypeal suture curved dorsad ( +Fig. 3C +). + + +Thorax +. Pronotum in dorsal view widest at midpoint with anterior margin moderately convex, posterior margin moderately concave, tricarinate, carina at midline continuous from anterior to posterior margin, lateral carinae extending from anterior margin of pronotum, laterad, becoming obsolete ventrally ( +Fig. 3A +). Mesonotum approximately as wide as long at midpoint, tricarinate at anterior margin, two additional carinae arising at mid carina near anterior margin, extending diagonally to just beyond lateral carinae, slightly sinuate ( +Fig. 3A +). In lateral view, greatly raised, angled where diagonal carinae arise at mid carina ( +Fig. 3B +). Hind tibiae with single lateral spine approximately 2/3 distance from femoral-tibial joint, spinulation 6-7-2. + + +Forewing reticulate, broadest basally, constricting at claval apex, expanding slightly, apex broadly rounded ( +Fig. 4 +). + + +Male terminalia +. Pyfoger in lateral view nearly uniform width along entire length, anterior margin sinuate, slightly concave, posterior margin nearly linear, dorsal and ventral margin equal length, slightly sinuate, medioventral process lacking ( +Fig. 5 +). Gonostyli in lateral view generally rounded with large dorsal process arising basally, with helical sclerotized ridges, apex blunt with posterior corner rounded, anterior corner hooked ( +Fig. 5 +). Aedeagus nearly symmetrical with three pairs of processes ( +Fig. 6 +); first pair (A1 & A2) arising on subapical lateral margin, angled dorsad and cephalad, both processes straight, slightly more sclerotized than rest of aedeagus ( +Fig. 6A, B +), second pair (A3 & A4) arising on inner ventral margin just basad of A1 and A2, angled dorsad and cephalad, slightly curved mesad, apices barely crossing at midpoint, approximately twice the length of A1 and A2 ( +Fig. 6C +). Third pair (A5 & A6) arising on ventral margin midline, short, slightly curved distad from base, equally sclerotized relative to A1–A4, approximately half length of A1 and A2 ( +Fig. 6C, D +). Anal segment elongate, irregularly sinuate on dorsal and ventral margins, dorsal margin angled ventrad approximately ¼ from base, ventral margin with strong invagination approximately 2/3 from base, apex distad of invagination comma-shaped. + + + + +Distribution. +Manzanillo, +Limón Province +, +Costa Rica +; Almonds and Corals Hotel. + + + + +Type material. + +Holotype +, male “ +Costa Rica +, +Limón +Pr. / nr. +Manzanillo +/ + +16.VI.2018 + +/ sweeping weeds / +Coll. +: +B.W.Bahder +// +Holotype +/ + +Colpoptera stonei + +♂ +” ( +FLREC +). + + + + +FIGURE 2. +Adult habitus + +Colpoptera stonei + + +sp. nov. + +(A) Male lateral view; (B) male dorsal view. + + + + +FIGURE 3. +Adult + +Colpoptera stonei + + +sp. nov. + +; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm. + + + + +FIGURE 4. +Forewing venation of + +Colpoptera stonei + + +sp. nov. + +Vein labeling nomenclature following + +Bourgoin +et al +. (2015) + +. + + + + +FIGURE 5. +Adult male + +Colpoptera stonei + + +sp. nov. +, + +terminalia in left lateral view. + + + +Paratypes +: +2 male +and +4 females +same as + +Holotype +, +1 male +and +2 females +from +La Sleva Biological Station +, +Heredia Province +, + +20.VI.2018 + + +. + + +Plant associations. +Unknown, collected sweeping herbaceous weeds. + + +Sequence Data. +For the COI gene, a 657 bp product was generated (GenBank Accession No. PP378885), for the 18S gene, a 1,265 bp product (GenBank Accession No. PP386391) was generated and for the 28S D9– D10 expansion region, a 829 bp product was generated (GenBank Accession No. PP386392). The phylogenies constructed demonstrate strong bootstrap support for the monophyly of + +Colpoptera + +based on 18S, 28S, COI data and concatenated sequences of the three loci (100, 99 and 80, respectively) with + +Colpoptera stonei + + +sp. nov. + +resolving within this clade ( +Fig. 7 +). + + + + +FIGURE 6. +Aedeagus of + +Colpoptera stonei + + +sp. nov. + +: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view. + + + + +FIGURE 7. +Maximum Likelihood trees (1,000 replicates) exhibiting relationship of + +Colpoptera stonei + + +sp. nov +. + +to available nogodinid taxa: (A) 18S, (B) 28S, (C) COI and (D) consensus tree based on concatenated data for the three loci; scale bar = percent nucleotide difference. + + + + +Remarks. +The general habitus of + +Colpoptera stonei + + +sp. nov. + +appears similar to many described species of + +Colpoptera + +and both the form of the terminalia and geography appear to exclude the other described genera in the +Colpopterini +(most genera are Antillean). The closest species in terms of coloration and general form of the aedeagus appears to be + +C. albavenosa +Caldwell + +from +San Luis Potosí +in northern +Mexico +( +Caldwell 1945 +, plate IV, fig. 2). However, + +Colpoptera stonei + + +sp. nov. + +differs from + +C. albavenosa + +in the anal segment where + +C. albavenosa + +is noted as having a nearly straight segment whereas + +Colpoptera stonei + + +sp. nov. + +has deep invaginations on the ventral margin.Also, the processes on the aedeagus of + +C. albavenosa + +are noted as subequal in length whereas the difference in size is substantial in + +Colpoptera stonei + + +sp. nov. + +(also +Caldwell, 1945 +, depicts three processes on each side of the aedeagus for + +albavenosa + +rather than two per side for + +C. stonei + + +sp. nov. + +). Finally, the gonostyli in + +C. albavenosa + +are shown as elongated, hooked, and narrow where they are stout with helical ridges + +C. stonei + + +sp. nov. + +Another species with similar form in the terminalia is + +C. acutata +Caldwell. + +The general armature of the aedeagus in general is similar, however, in + +C. stonei + + +sp. nov. + +the ventral bifurcation of the processes arises mesad and in + +C. acutata + +(based on the description) appear to arise on the lateral margins. Additionally, + +C. acutata + +is described as a red-brown species with white spots on the frons whereas + +C. stonei + + +sp. nov. + +is not red-brown and has an unmarked frons. Finally, the apex of the anal segment in + +C. acutata + +appears narrowed and slightly hooked based on the description and it is much broaded and not hooked in + +C. stonei + + +sp. nov. + + + +While molecular data is useful for elucidating + +Colpoptera + +, very little data is available for other nogodinids taxa that would allow for generating a consensus tree among COI, 18S, and 28S. As new taxa are discovered and previously described taxa become available for analysis, this shortcoming can be addressed. + + + + \ No newline at end of file diff --git a/data/03/9B/0E/039B0E04FF88FFEBFF538218FC0C0940.xml b/data/03/9B/0E/039B0E04FF88FFEBFF538218FC0C0940.xml new file mode 100644 index 00000000000..6d959cc6761 --- /dev/null +++ b/data/03/9B/0E/039B0E04FF88FFEBFF538218FC0C0940.xml @@ -0,0 +1,158 @@ + + + +A new species of planthopper in the genus Colpoptera (Hemiptera: Fulgoroidea: Nogodinidae) from the Caribbean coast of Costa Rica + + + +Author + +Barrantes Barrantes, Edwin A. +Universidad de Costa Rica - Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica + + + +Author + +Zumbado Echavarria, Marco A. +Universidad de Costa Rica - Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica + + + +Author + +Van Dam, Alex R. +0000-0002-1966-0338 +alex.vandam@upr.edu + + + +Author + +Helmick, Ericka E. +University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA + + + +Author + +Bartlett, Charles R. +University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA + + + +Author + +Martinez Aponte, Laura V. +0009-0003-8030-0651 +laura.martinez9@upr.edu + + + +Author + +Ruiz, Alfredo Rodriguez +0000-0002-1311-1637 +arodr412@asu.edu + + + +Author + +Bloch, Melody +University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA + + + +Author + +Bahder, Brian W. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +341 +352 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.3 + +journal article +10.11646/zootaxa.5481.3.3 +1175-5326 +12758762 +798238C5-D310-4FA1-BF2D-6D365250655C + + + + + + +Genus + +Colpoptera +Burmeister, 1835 + + + + + + + + +Type +species. + +Colpoptera sinuata +Burmeister, 1835 + +( +type +by subsequent designation of Distant 1910: 314) + + + + +Type +species. + + +Colpoptera sinuata +Burmeister, 1835 + + + + + +Diagnosis +. (Abridged from +Gnezdilov & O’Brien 2008 +). Body elongated, head about as broad as prothorax, eyes prominent. Frons elongate, with distinct median carina. Vertex transverse, anterior margin truncate, posterior margin concave. Pronotum narrow, anterior margin strongly convex, posterior margin concave. Mesonotum at midline longer than vertex + pronotum combined, bearing arcuate anterior transverse carinae and median carina. Forewings elongate and narrowed to rounded apex, with narrow hypocostal plate and many transverse veins apically, hindwings large and functional. Hind tibia with single lateral tooth distally. Male pygofer with nearly straight hind margin (elongate process absent). Aedeagus relatively simple, bilaterally symmetrical, with elongated, usually bifurcated, subapical lateral processes (“phallobase lobe” of +Gnezdilov & O’Brien 2008 +); and with pair of shorter, simple. Gonostyli bearing broad capitulum, narrowing apically with broad lateral tooth.Anal tube elongated. Female sternite VII with rounded median process ( +Gnezdilov & O’Brien 2008 +, fig. 59). Gonoplacs nearly triangular, fused medially; gonocoxa VIII with straight hind margin ( +Gnezdilov & O’Brien 2008 +, figs. 60–62). Anal tube long and narrow, distinctly longer than gonoplacs +Gnezdilov & O’Brien 2008 +, fig. 57). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF1CF711ED9FE640E3505EC.xml b/data/03/D7/87/03D78790FFF1CF711ED9FE640E3505EC.xml new file mode 100644 index 00000000000..de57f2831d0 --- /dev/null +++ b/data/03/D7/87/03D78790FFF1CF711ED9FE640E3505EC.xml @@ -0,0 +1,206 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus motuoensis +Hassan & Xing + +sp. nov. + + + + + + +( +Figs 1–13 +) + + +Description. +Body coloration light yellowish green ( +Figs 1–4 +). Head including eyes as broader as pronotum, crown light yellowish green ( +Fig. 1 +), face light green, without markings. Antennae long, 1.5 x width of the head or longer, scape and pedicel brownish yellow, flagellum dark brown ( +Fig. 3 +). Pronotum light yellowish green, posterior margin brownish yellow, with some dark brown irroration on anterior margin. Scutellum brownish yellow, with a pair median of longitudinal light green bands present or absent in some individuals ( +Fig. 4 +). Forewing light brownish, venation dark brown. Legs pale yellow, unmarked, except for darker bases of macrosetae ( +Fig. 2 +). + + + +FIGURES 1–4. + +Fistulatus motuoensis +Hassan & Xing + + +sp. nov. + +Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view. + + + + +FIGURES 5–13. + +Fistulatus motuoensis +Hassan & Xing + + +sp. nov. + +Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10. Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex. + + + +Male genitalia ( +Figs 5–13 +). Pygofer slightly wider than long, posteroventral margin of lobe with long, upward directed process, posterodorsally rounded with triangular apex, covered with numerous stout setae at posterolaterally ( +Fig. 6 +). Valve triangular. Subgenital plate triangular with narrow and elongated apex ( +Fig. 7 +). Connective very short, Y-shaped, stem subequal to arms in length ( +Fig. 9 +). Style moderately long, broad at base, curved laterally with apex rostriform ( +Fig. 8 +). Aedeagus long and straight, with a pair of slender, retrorse subapical lateral spines arising from convex lateral flanges; shaft broadened at base and subapically in dorsal view, with apex fish-shaped in anteroventral view ( +Figs 10–13 +). + + +Measurement. +Length (including tegmen): +♂ +, 6.9–7.0 mm, +♀ +7.5 mm +. + + + + +Type material. + +Holotype +♂ +, +CHINA +: +Xizang +Autonomous Region +, +Motuo County +, + +08 August 2020 + +, coll. +Yongjin Sui +( +GUGC +); +paratypes +: +1♂ +, +1♀ +, same data as holotype. + + + + + +Etymology. +The specific epithet + +‘ +motuoensis + +’ refers to the +type +locality of the new species. + + + + +Remarks. + +F. motuoensis +Hassan & Xing + + +sp. nov. + +is unique among its known congeners in having the ventral pygofer appendage relatively long and straight and by the aedeagal shaft with slender paired lateral prepical spines and the apex broadened and hammer-like in lateral view.. + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF4CF7D1ED9F9660FD506C4.xml b/data/03/D7/87/03D78790FFF4CF7D1ED9F9660FD506C4.xml new file mode 100644 index 00000000000..61bd57463da --- /dev/null +++ b/data/03/D7/87/03D78790FFF4CF7D1ED9F9660FD506C4.xml @@ -0,0 +1,299 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus sinensis +Zhang, Zhang & Chen, 1997 + + + + + + + +( +Figs 50–54 +) + + + + + + + +Fistulatus sinensis +Zhang, Zhang & Chen, 1997: 237 + + +. + + + + + +Material examined. + + +CHINA +: + +1♂ +, +Hebei Prov. +, +Zhangjiakou City +, +Xiaowutai +, + +12–13 August 2010 + +, coll. +Zhihua Fan +( +GUGC +) + +; + +3♂♂ +6♀♀ +, +Shanxi Prov. +, +Linfen City +, +Yicheng County +, +Lishan +, + +23–26 July 2012 + +, coll. +Qiongzhang Song +( +GUGC +) + +; + +10♀♀ +, +Shanxi Prov. +, +Linfen City +, +Yicheng County +, +Lishan +, + +24–25 July 2011 + +, coll. +Hu Li +( +GUGC +) + +; + + + +6♂♂ +4♀♀ +, +Guizhou Prov. +, +Dushan County +, +Duliu Jiangyuan Wetland +, + +12–17 July 2012 + +, coll. +Qiongzhang Song +( +GUGC +) + +; + +1♂ +, +Hebei Prov. +, +Chengde City +, +Xinglong County +, +Wulingshan +, + +8 July 2011 + +, coll. +Hu Li +( +GUGC +) + +; + +1♂ +, +Shaanxi Prov. +, +Liuba County +, +Yingpan Town +, + +21 July 2010 + +, coll. +Hu Li +and +Zhihua Fan +( +GUGC +) + +; + +3♂♂ +, +Shaanxi +Pro. +, +Ningshan Huoditang Forest +Farm, + +12 July 2012 + +, coll. +Weicheng Yang +and +Weibin Zheng +( +GUGC +) + +. + + + + +Remarks. +This species can be distinguished from its congeners by the aedeagal shaft with a pair of spinelike lateral preapical processes in posterior view and the style with and apical extension in lateral view. + + + + +Distribution: +China +: +Shaanxi +, +Henan +, +Gansu +( + +Zhang +et al. +1997 + +; +Cen & Cai, 2002 +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +), +Hebei +, +Shanxi +, +Guizhou +. + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF5CF731ED9F9AA087A0034.xml b/data/03/D7/87/03D78790FFF5CF731ED9F9AA087A0034.xml new file mode 100644 index 00000000000..936e66b63fd --- /dev/null +++ b/data/03/D7/87/03D78790FFF5CF731ED9F9AA087A0034.xml @@ -0,0 +1,166 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus rectilineus +Shang & Zhang, 2006 + + + + + + + +( +Figs 44–49 +) + + + + + + +Fistulatus rectilineus +Shang & Zhang + +in + + +Shang +et al. +2006: 152 + + +. + + + + + +Material examined. + + +CHINA +: + +2♂♂ +, +Sichuan Prov. +, +Kangding County +, + +31 July 2012 + +, coll. +Zhihua Fan +and +Qiuxiao Tang +( +GUGC +) + +. + + + + +Remarks. +This species can be distinguished by the medially directed pygofer processes and the cylindrical, straight, and elongated aedeagal shaft, without processes. + + + + +Distribution: +China +: +Sichuan +( + +Zhang +et al. +1997 + +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF5CF731ED9FBD508A302DB.xml b/data/03/D7/87/03D78790FFF5CF731ED9FBD508A302DB.xml new file mode 100644 index 00000000000..525b47114f1 --- /dev/null +++ b/data/03/D7/87/03D78790FFF5CF731ED9FBD508A302DB.xml @@ -0,0 +1,133 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus monkoboensis +Shang & Zhang, 2003 + + + + + + + +( +Figs 40–44 +) + + + + + + + +Fistulatus monkoboensis +Shang & Zhang, 2003: 21 + + +. + + + + + +Remarks. +This species is similar to + +F. athena +Shang & Zhang + +in the shape of the pygofer processes but can be distinguished by the absence of lateral processes on the connective and by the style with apical process strongly curved and not digitate in lateral view. + + + + +Distribution: +Malaysia +: +Sabah +( +Shang & Zhang 2003 +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF5CF731ED9FDB40E3D0343.xml b/data/03/D7/87/03D78790FFF5CF731ED9FDB40E3D0343.xml new file mode 100644 index 00000000000..998b60b4274 --- /dev/null +++ b/data/03/D7/87/03D78790FFF5CF731ED9FDB40E3D0343.xml @@ -0,0 +1,272 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus luteolus +Cen & Cai, 2002 + + + + + + + +( +Figs 35–39 +) + + + + + + + +Fistulatus luteolus +Cen & Cai, 2002: 119 + + +. + + + + + +Material examined. + + +CHINA +: + +3♂♂ +, +Hubei Prov. +, +Shenlongjia Guanmenshan +, + +16 July 2013 + +, coll. +Dongliang Xing +, +Zhimin Chang +( +GUGC +) + +; + +8♂♂ +, +Hunan Prov. +, +Badashan Tianpingshan Management Station +, + +5 August 2013 + +, coll. +Ling Qu +and +Yunfei Wu +( +GUGC +) + +; + +1♂ +, +Guizhou Prov. +, +Dushan County +, +Duliu Jiangyuan Wetland +, + +12–17 August 2012 + +, coll. +Qiongzhang Song +( +GUGC +) + +; + +1♂ +14♀♀ +, +Shaanxi Prov. +, +Heihe National Forest +Park, + +10–12 August 2010 + +, coll. +Zhimin Chang +and +Yanli Zheng +( +GUGC +) + +; + +1♂ +1♀ +, +Guizhou Prov. +, +Kuankuoshui +, + +14 August 2010 + +, coll. +Jichun Xing +( +GUGC +) + +; + +1♂ +, +Hubei Prov. +, +Shenlongjia Guanmenshan +, + +15 July 2013 + +, coll. +Ling Qu +and +Yunfei Wu +( +GUGC +) + +. + + + + +Remarks. +This species is similar to + +F. quadrispinosus +Lu & Zhang + +in having two pairs of pygofer processes but can be distinguished by the absence of processes on the aedeagal shaft. + + + + +Distribution: +China +: +Zhejiang +, +Henan +( + +Zhang +et al. +1997 + +; +Cen & Cai, 2002 +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +), +Hubei +, +Hunan +, +Guizhou +, +Shaanxi +. + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF5CF731ED9FF3E0FC506A5.xml b/data/03/D7/87/03D78790FFF5CF731ED9FF3E0FC506A5.xml new file mode 100644 index 00000000000..6c4b1502871 --- /dev/null +++ b/data/03/D7/87/03D78790FFF5CF731ED9FF3E0FC506A5.xml @@ -0,0 +1,143 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus brevimarginalis +Yang & Dai, 2024 + + + + + + + +( +Figs 30–34 +) + + + + + + + +Fistulatus brevimarginalis + +Yang & Dai, 2024: 117 + + + +. +Type +locality: +China +( +Yunnan +: +Yingjiang Country +). + + + + + +Remarks. +This species is similar to + +F. brevimarginalis +Yang & Dai + +, + +F. luteolus +Cen & Cai + +and + +F. quadrispinosus +Lu & Zhang + +in having the pygofer with two pairs of processes but can be distinguished by the aedeagal shaft with a single dorsal preapical process and no paired apical processes. + + + + +Distribution: +China +: +Yunnan +( + +Yang +et al. +2024 + +). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF7CF711ED9F93E0FD90098.xml b/data/03/D7/87/03D78790FFF7CF711ED9F93E0FD90098.xml new file mode 100644 index 00000000000..04217003b3c --- /dev/null +++ b/data/03/D7/87/03D78790FFF7CF711ED9F93E0FD90098.xml @@ -0,0 +1,155 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus biconjugara +Yang & Dai, 2024 + + + + + + + +( +Figs 25–29 +) + + + + + + + +Fistulatus biconjugara + +Yang & Dai, 2024: 117 + + + +. +Type +locality: +China +( +Guizhou +: +Fanjing Mountain +). + + + + + +Remarks. +This species is similar to + +F. brevimarginalis +Yang & Dai + +, + +F. luteolus +Cen & Cai + +and + +F. quadrispinosus +Lu & Zhang + +in having the pygofer with two pairs of processes but can be distinguished by the aedeagal shaft with only small distal processes (aedeagal shaft with two pairs of processes in + +F. quadrispinosus + +and without processes in + +F. brevimarginalis + +and + +F. luteolus + +). + + + + +Distribution: +China +: +Guizhou +( + +Yang +et al. +2024 + +). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF7CF711ED9FB280CD1012F.xml b/data/03/D7/87/03D78790FFF7CF711ED9FB280CD1012F.xml new file mode 100644 index 00000000000..17d68d7d43c --- /dev/null +++ b/data/03/D7/87/03D78790FFF7CF711ED9FB280CD1012F.xml @@ -0,0 +1,223 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus bidentatus +Cen & Cai, 2002 + + + + + + + +( +Figs 20–24 +) + + + + + + + +Fistulatus bidentatus + +Cen & Cai, 2002: 117 + + + +. +Type +locality: +China +( +Zhejiang +: +Tianmushan +). + + + + + +Material examined. + + +CHINA +: + +1♂ +, +Guangxi +Autonomous Region +., +Damingshan +, + +15 May 2012 + +, coll. +Hu Li +( +GUGC +) + +; + +3♂♂ +, +Guizhou Prov. +, +Leigongshan +, + +4–5 July 2011 + +, coll. +Jiankun Long +and +Weibin Zheng +( +GUGC +) + +; + +1♂ +, +Zhejiang Prov. +, +Tianmushan +, + +20 July 2009 + +, coll. +Zehong Meng +and +Yong Chen +( +GUGC +) + +. + + + + +Remarks. +This species resembles + +F. luteolus +Cen & Cai + +and + +F. quadrispinosus +Lu & Zhang + +in having a distinctive elongated aedeagal shaft with short processes near the midlength of the shaft but differs in having a pair of short distal processes on the shaft. + + + + +Distribution: +China +: +Zhejiang +( + +Zhang +et al. +1997 + +; +Cen & Cai, 2002 +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +); +Guangxi +, +Guizhou +. + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFF7CF711ED9FD7F08A30359.xml b/data/03/D7/87/03D78790FFF7CF711ED9FD7F08A30359.xml new file mode 100644 index 00000000000..97d84137104 --- /dev/null +++ b/data/03/D7/87/03D78790FFF7CF711ED9FD7F08A30359.xml @@ -0,0 +1,141 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus athena +Shang & Zhang, 2003 + + + + + + + +( +Figs 14–19 +) + + + + + + + +Fistulatus athena + +Shang & Zhang, 2003: 21 + + + +. +Type +locality: +Malaysia +( +Sabah +). + + + + + +Remarks. +This species is similar to + +F. monkoboensis +Shang & Zhang + +in having a distinctive shape of pygofer processes, which are directed ventrally and slightly backward in lateral view, but can be distinguished by the presence of lateral processes on connective and by the style with the apical process slightly curved and digitate in lateral view. + + + + +Distribution: +Malaysia +: +Sabah +( +Shang & Zhang 2003 +; + +Shang +et al. +2006 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +). + + + + \ No newline at end of file diff --git a/data/03/D7/87/03D78790FFFBCF7D1ED9FE5709A10479.xml b/data/03/D7/87/03D78790FFFBCF7D1ED9FE5709A10479.xml new file mode 100644 index 00000000000..e2b491a5ea6 --- /dev/null +++ b/data/03/D7/87/03D78790FFFBCF7D1ED9FE5709A10479.xml @@ -0,0 +1,131 @@ + + + +Key to species of the leafhopper genus Fistulatus Zhang, Zhang & Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species + + + +Author + +Hassan, Muhammad Asghar +0000-0003-2590-5781 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China +kakojan112@gmail.com + + + +Author + +Shah, Bismillah +0000-0002-8407-8627 +Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A & F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China +bismillahshah1990@yahoo.com + + + +Author + +Xing, Jichun +0000-0002-3738-7996 +Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +363 +372 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.5 + +journal article +10.11646/zootaxa.5481.3.5 +1175-5326 +12758855 +126C97BD-856B-44DA-8EED-91F26CAD1C32 + + + + + + + +Fistulatus quadrispinosus +Lu & Zhang, 2014 + + + + + + + +( +Figs 55–59 +) + + + + + + + +Fistulatus quadrispinosus +Lu & Zhang, 2014: 247 + + +. + + + + + +Remarks. +This species is similar to + +F. luteolus +Cen & Cai + +in having two pairs of processes on the pygofer but can be distinguished by a pair of narrow medial processes on the aedeagal shaft in ventral view. + + + + +Distribution: +China +: +Zhejiang +( + +Zhang +et al. +1997 + +; +Lu & Zhang 2014 +; + +Qu +et al. +2015 + +). + + + + \ No newline at end of file diff --git a/data/03/DD/87/03DD87BA932E904855FA5D5E4169FA29.xml b/data/03/DD/87/03DD87BA932E904855FA5D5E4169FA29.xml index df546ab14f5..0a6da484b91 100644 --- a/data/03/DD/87/03DD87BA932E904855FA5D5E4169FA29.xml +++ b/data/03/DD/87/03DD87BA932E904855FA5D5E4169FA29.xml @@ -1,92 +1,95 @@ - - - -Hippocrepis fuelleborni (Digenea: Notocotylidae) from Argentina, morphological molecular and phylogenetic studies + + + +Hippocrepis fuelleborni (Digenea: Notocotylidae) from Argentina, morphological molecular and phylogenetic studies - - -Author + + +Author -Martins, Natalia Beatriz Guerreiro -Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. +Martins, Natalia Beatriz Guerreiro +0000-0001-6677-420X +Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. +natalia_gmartins@cepave.edu.ar - - -Author + + +Author -Robles, María Del Rosario -0000-0003-4197-8040 -Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. -rosario@cepave.edu.ar +Robles, María Del Rosario +0000-0003-4197-8040 +Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. +rosario@cepave.edu.ar - - -Author + + +Author -Canova, Victoria -0000-0001-9242-7017 -Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. -victoriac@cepave.edu.ar +Canova, Victoria +0000-0001-9242-7017 +Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. +victoriac@cepave.edu.ar - - -Author + + +Author -Diaz, Julia Inés -0000-0002-8500-0880 -Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. -jidiaz@cepave.edu.ar +Diaz, Julia Inés +0000-0002-8500-0880 +Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. +jidiaz@cepave.edu.ar - - -Author + + +Author -Balcazar, Dario -/0000-0002-8490-7683 -Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina. -dbalcazar@cepave.edu.ar +Callejón, Rocío +0000-0002-6276-3818 +Departamento de Microbiología y Parasitología, Facultad de Farmacia, Universidad de Sevilla, Sevilla, España. +callejon@us.es - - -Author + + +Author -Callejón, Rocío -Departamento de Microbiología y Parasitología, Facultad de Farmacia, Universidad de Sevilla, Sevilla, España. +Balcazar, Dario +0000-0002-8490-7683 +Centro de Estudios Parasitológicos y de Vectores (CEPAVE), FCNyM, UNLP, CONICET, Boulevard 120 entre Av. 60 y Calle 64, 1900 La Plata, Argentina +dbalcazar@cepave.edu.ar -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-07-16 + +2024 + +2024-07-16 - -5481 + +5481 - -2 + +2 - -225 -240 + +225 +240 - -http://dx.doi.org/10.11646/zootaxa.5481.2.3 + +http://dx.doi.org/10.11646/zootaxa.5481.2.3 -journal article -299923 -10.11646/zootaxa.5481.2.3 -f09b8715-e7a9-4dbf-a8a8-27c8a753888f -1175-5326 -12749559 -F55D186D-8A64-47F7-8338-7C6ECF4A0B55 +journal article +10.11646/zootaxa.5481.2.3 +f09b8715-e7a9-4dbf-a8a8-27c8a753888f +1175-5326 +12749559 +F55D186D-8A64-47F7-8338-7C6ECF4A0B55 - + @@ -104,37 +107,37 @@ Travassos & Vogelsong, 1930 Description (based on 22 specimens ): body flat, elongated ( -Figs. 2A–B +Figs. 2A–B , -3A +3A ), 2.81 ± 0.44 (1.68–3.52) long by 0.77 ± 0.13 (0.45–1.04) wide. Lateral projections on each side of the oral sucker ( -Figs. 2B +Figs. 2B , -3C +3C ). Tegument covered with spines ( -Fig. 3E–F +Fig. 3E–F ), smooth oral sucker edges, and covered with small spines from the oral sucker to the genital pore ( -Fig. 3C–D +Fig. 3C–D ). Ventral surface concave and provided with numerous longitudinal rows of ventral papillae arranged irregularly, extending from the middle zone of the cirrus sac to posterior extremity ( -Figs. 2B +Figs. 2B , -3A–B +3A–B , -4A +4A ), with a diameter of 51.97 ± 6.34 (43.02–57.81) µm. Ventral sucker absent. Oral sucker subterminal, 0.28 ± 0.04 (0.21–0.34) long by 0.22 ± 0.003 (0.17–0.32) wide. Pharynx absent; oesophagus 0.05 ± 0.01 (0.04–0.07). Caeca with lateral diverticula in its entire extension, bordering vitelline follicles, fuse posteriorly to the ovary passing between testes, up to level of their posterior border ( -Figs. 2A +Figs. 2A , -4C +4C ). Testes irregularly lobed, oblique, posterior to caecal union, right testis 0.24 ± 0.05 (0.15–0.34) long by 0.19 ± 0.04 (0.13–0.27) wide, left testis 0.23 ± 0.05 (0.15–0.32) long by 0.19 ± 0.05 (0.13–0.28) wide. Cirrus sac median, elongate 0.68 ± 0.08 (0.55–0.88) long, containing cirrus and internal seminal vesicle ( -Fig. 2C +Fig. 2C ). Cirrus not everted, with small spines ( -Fig. 4B +Fig. 4B ). Genital pore median, posterior to caecal bifurcation. Ovary median, pretesticular, transversally elongated ( -Fig. 2D +Fig. 2D ), 0.13 ± 0.03 (0.10–0.19) long by 0.24 ± 0.06 (0.10–0.36) wide. Vitelline follicles arranged in longitudinal series, extracaecal and pre-ovarian, ending close to testes, 0.60 ± 0.07 (0.50–0.70) long. Uterus transverse coils located in intercaecal space. Eggs oval, without polar filaments ( -Figs. 2E +Figs. 2E , -4D +4D ), 26.38 ± 11.94 (20.70–69.15) µm long by 8.54 ± 1.09 (6.72–10.25) µm wide. diff --git a/data/1A/2C/55/1A2C557E854F3E5B9EACFB88FE56F97B.xml b/data/1A/2C/55/1A2C557E854F3E5B9EACFB88FE56F97B.xml new file mode 100644 index 00000000000..186f3b2f805 --- /dev/null +++ b/data/1A/2C/55/1A2C557E854F3E5B9EACFB88FE56F97B.xml @@ -0,0 +1,295 @@ + + + +A new species of Leptomastax Pirazzoli in Israel (Coleoptera: Staphylinidae: Scydmaeninae) + + + +Author + +Jałoszyński, Paweł + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +391 +396 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.8 + +journal article +10.11646/zootaxa.5481.3.8 +1175-5326 +12758924 +F885B41C-3D17-4CD2-8046-85C2D7EE4721 + + + + + + + +Leptomastax minutissima + +sp. n. + + + + + + +( +Figs 1‒13 +) + + + + +Type material. + + +Holotype +: + +( + +ISRAEL +): + +♂ +, two labels: “ISRAEL: Galilea / Eilon, +N. Betzel +/ valley, + +20.IV.1982 + +/ Löbl, Bes. nr. 13a” [white, printed], “ + +LEPTOMASTAX + +/ + +minutissima + +m. / +P. Jałoszyński +, '24 / +HOLOTYPUS +” [red, printed] ( +MHNG +). + + + + + +Diagnosis. +Body extremely small, below +1.2 mm +; eyes lacking; subocular and elytral macrosetae lacking; prosternum with two large lateral patches of modified setae; anterior region of mesoventral process much wider than its posterior intermesocoxal region; elytral rows of punctures irregular, subhumeral and marginal rows lacking; male with U-shaped impression with distinct lateral margins on posterior 1/3 of metaventrite; aedeagus with simple (i.e., unbranched) and nearly straight endophallus; each paramere with evenly rounded subapical and apical margin and with pointed apex directed mesally. + + + + +Description. +Body of male ( +Fig. 1 +) strongly elongate, strongly flattened, yellowish brown. BL +1.13 mm +. + + +Head ( +Fig. 2 +) broadest near middle, eyeless, HL +0.15 mm +, HW +0.28 mm +; vertex and frons together slightly more than twice as wide as long; dorsal postmandibular impressions distinct; subocular (sensu Castellini (1994)) macrosetae absent, vertexal macrosetae (= +temporo-occipitale +of Castellini) present. Dorsum of head and genae ( +Fig. 5 +) impunctate. Antenna ( +Fig. 3 +) slender, AnL +0.50 mm +, scape as long as antennomeres 2–5 together, pedicel 1.3 times as long as broad, antennomeres 3–10 each transverse (3 strongly, 10 weakly so), antennomere 11 nearly as long as 9–10 combined, 1.8 times as long as broad. + +Labial palps moderately long, palpomere 2 about twice as long as broad, palpomere 3 setiform, slightly longer than 2. + +Pronotum ( +Fig. 2 +) subtrapezoidal, broadest near anterior fourth; PL +0.30 mm +, PW +0.28 mm +. Anterior margin evenly and weakly rounded; lateral margins strongly rounded in anterior third and nearly straight in posterior half; posterior corners strongly obtuse-angled; posterior margin weakly and evenly rounded. Pronotal disc impunctate and asetose. Prosternum ( +Fig. 5 +) with pair of large anterolateral patches of modified, strongly flattened and broadened scale-like setae and narrow row of similar setae along median portion of anterior margin. + + + +FIGURES 1–4. + +Leptomastax minutissima + +sp. n. +, holotype male. Dorsal habitus ( +1 +); head, pronotum and elytral base in dorsal view ( +2 +); left antenna in dorsal view ( +3 +); elytra in dorsal view ( +4 +). + + + +Mesoventrite ( +Figs 8‒9 +) with scale-like setae on procoxal rests and elongate flattened setae on mesoventral process. Metaventrite ( +Figs 8‒9 +) with scale-like setae filling postmesocoxal foveae and with elongate flattened setae evenly distributed on remaining surface. Posterior third of metaventrite with median U-shaped impression. + + +Elytra ( +Fig. 4 +) together oval and strongly elongate, broadest near middle; EL +0.68 mm +, EW +0.35 mm +, EI 1.93; humeral region strongly narrowing towards prothorax; lateral margins weakly rounded; apices separately rounded. Each elytron with sutural and humeral row of punctures, lacking subhumeral and marginal rows; punctures distinct but unevenly distributed, so that rows are somewhat irregular. Macrosetae (and setiferous punctures) in humeral and apical regions lacking. Elytra asetose and outside rows of punctures impunctate. + + +Legs ( +Figs 1 +, +9 +) moderately long and slender, unmodified; each mesocoxa with two macrosetae. + + +Abdomen ( +Fig. 8 +) with regular setae almost evenly covering all exposed sternites. + + +Aedeagus ( +Figs 10‒13 +) slender, AeL +0.18 mm +; median lobe in ventral view gradually narrowing towards subtriangular apex, in lateral view distal region nearly straight; sclerotized endophallus unipartite, tubular and unbranched, nearly straight; parameres with evenly rounded outer margins in subapical and apical regions, apex of each paramere pointed and directed mesally. + +Female. Unknown. + + + +FIGURES 5–7. + +Leptomastax minutissima + +sp. n. +, holotype male. Head, prothorax and anterior region of mesothorax in ventral view ( +5 +); mouthparts in ventral view ( +6 +); left labial palp in ventral view ( +7 +). + + + + +Distribution. +Northern +Israel +. + + + + +Etymology. +The specific adjective + +minutissima + +refers to the smallest body among all known species of + +Leptomastax + +. + + + + +Remarks. +This is the smallest known species of + +Leptomastax + +; it can be easily identified by the slender, strongly elongate elytra, reduced and irregular elytral rows of punctures, lack of eyes, and aedeagal structures. The simple, unbranched endophallus in + +L. minutissima + +resembles that of + +L. cretica +Meybohm + +(in + +Assing +et al. +(2019) + +, fig. 149), but the latter species is much larger (BL +1.73‒1.83 mm +) and has large eyes, each composed of “about 14 ommatidia” ( + +Assing +et al. +2019 + +). The apical region of median lobe and the endophallus in + +L. cretica + +in lateral view are strongly curved (nearly straight in + +L. minutissima + +), and the parameral apices in ventral view in + +L. cretica + +are rapidly bent mesally (evenly and gradually curved in + +L. minutissima + +). + + +Notes containing detailed collecting data for the studied specimen have been lost; is is only known that most specimens taken during the Löbl-Besuchet 1982 fieldwork in Eilon come from sifted oak leaf litter (email from Ivan Löbl dated +18/06/2024 +). + + + + \ No newline at end of file diff --git a/data/83/08/87/830887F6FFFA0A14FF7BFD69FD24AB4F.xml b/data/83/08/87/830887F6FFFA0A14FF7BFD69FD24AB4F.xml new file mode 100644 index 00000000000..9ab81946eb6 --- /dev/null +++ b/data/83/08/87/830887F6FFFA0A14FF7BFD69FD24AB4F.xml @@ -0,0 +1,115 @@ + + + +Description of Neosilphitrombium rhipicephalum sp. nov. (Prostigmata: Neothrombiidae) ectoparasite on hard ticks from Iran + + + +Author + +Kohansal, Mahnaz +Department of Plant Protection, College of Agriculture, University of Zabol, Zabol, Iran + + + +Author + +Noei, Javad +Department of Plant Protection, Faculty of Agriculture, University of Birjand, Birjand, Iran + + + +Author + +Ramroodi, Sara +Department of Plant Protection, College of Agriculture, University of Zabol, Zabol, Iran + + + +Author + +Rakhshani, Ehsan +Department of Plant Protection, College of Agriculture, University of Zabol, Zabol, Iran + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +326 +340 + + + + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +journal article +10.11646/zootaxa.5481.3.2 +1175-5326 +12758788 +46EB61BE-A976-4367-A448-7FFE15162212 + + + + + + +Genus + +Neosilphitrombium +Fain, 1992 + + + + + + + + +Type +species: + + +Neosilphitrombium gratum +Fain, 1992 + + + + + + +Diagnosis of larva (According to + +Saboori +et al +. (2011) + +with some modifications): + + + +Neothrombiids with eyes 2 + 2; two dorsal scuta, AL setae bilobed and posterior or posteromedially to AM setae, sensilla posteromedially to PL setae, scutellum with two or more normal setae; coxal setal formula 3-4-2, all coxal setae bifid except lateral seta on coxa II ( + +2b +4 + +); subcapitular setae ( +bs +) bilobed; tarsal claws 3-3-3; palpal tibial claw bifurcate; palp femur and genu without seta. + + + + \ No newline at end of file diff --git a/data/A6/4A/2B/A64A2B29256A163142FEFE5EFD669E2F.xml b/data/A6/4A/2B/A64A2B29256A163142FEFE5EFD669E2F.xml new file mode 100644 index 00000000000..9b0e60075bb --- /dev/null +++ b/data/A6/4A/2B/A64A2B29256A163142FEFE5EFD669E2F.xml @@ -0,0 +1,158 @@ + + + +On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +384 +390 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.7 + +journal article +10.11646/zootaxa.5481.3.7 +1175-5326 +12758944 +F86F4482-5F80-444F-A5DE-F1F1BAC04E41 + + + + + + + +Hygrodromicus coriaceus +( +Cameron, 1924 +) + + + + + + + + + + +Geodromicus coriaceus +Cameron, 1924: 172 + + +. + + + + + + +Geodromicus perplexus +Cameron, 1941: 144 + + +. + + + + + + +Geodromicus palpalis +Coiffait, 1977: 271 + + +. + + + + + + +Hygrogeus ledouxi +Coiffait, 1983: 282 + + + +syn. n. + + + + + + +Hygrodromicus coriaceus + +: + +Zerche 1992: 120 + +, + +Shavrin 2021: 338 + +, + +2024: 237 + +. + + + + + +Remarks. + +Hygrogeus ledouxi + +based on the single +holotype +was originally described from “Cachemire, Pahalgam, Sheshnag…”. Based on the original description and the shape of the aedeagus ( +Figs 2I and 2J +in +Coiffait 1983 +), + +H. ledouxi + +is conspecific with the Himalayan + +H. coriaceus + +, also known from Kashmir ( +Shavrin 2021 +, +2024 +). Thus, + +H. ledouxi + +is synonymized with the latter species. + + + + \ No newline at end of file diff --git a/data/A6/4A/2B/A64A2B29256A163342FEFC83FED99F95.xml b/data/A6/4A/2B/A64A2B29256A163342FEFC83FED99F95.xml new file mode 100644 index 00000000000..080c17dfeaa --- /dev/null +++ b/data/A6/4A/2B/A64A2B29256A163342FEFC83FED99F95.xml @@ -0,0 +1,300 @@ + + + +On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +384 +390 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.7 + +journal article +10.11646/zootaxa.5481.3.7 +1175-5326 +12758944 +F86F4482-5F80-444F-A5DE-F1F1BAC04E41 + + + + + + + +Hygrodromicus incrassatus + +sp. n. + + + + + + +( +Figs 1 +, +3–4 +, +7 +) + + + + +Type material examined. +Holotype +♂ +(a plastic plate with the aedeagus in Canada balsam was pinned under the card with the beetle; abdominal tergite VIII, sternite VIII, and the apical segment are glued on the same card under the specimen): ‘ +PAKISTAN +: NW Frontier | province, Kaghan valley, nr. | Sighban, +2200 m +, | +20.VIII.2004 +, leg. +V +. Gurko’ <printed>, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ +HOLOTYPE +| + +Hygrodromicus + +| + +incrassatus + +sp. n. +| Shavrin A. +V +. des. 2024’ <red, printed> (cSch). + + +Paratypes +: +2 ♀♀ +( +Fig. 1 +; +one specimen +without right fore leg): same data as the holotype, with additional red printed label: ‘ +PARATYPE +| + +Hygrodromicus + +| + +incrassatus + +sp. n. +| Shavrin A. +V +. des. 2024’ (cSch, cSh). + + + + +Description. +Measurements (n = 3): HW: 1.05–1.10; HL: 0.70–0.73; OL: 0.12–0.14; LT: 0.10–0.13; AL ( +holotype +): 4.15; PL: 0.92–1.05; PWmax: 1.26–1.45; PWmin: 1.03–1.20; ESL: 1.52–1.85; EW: 1.94–2.30; MTbL( +holotype +): 1.55; MTrL ( +holotype +): 0.57 (MTrL 1–4: 0.32; MTrL 5: 0.25); AW: 2.24–2.40; AedL: 1.25; BL: 5.90–6.75 ( +holotype +: 5.92). + + +Habitus as in +Fig. 1 +. Body reddish-brown (elytra of +holotype +distinctly paler); mouthparts, antennae and legs brownish; apical segment of maxillary palpi and tarsi yellowish. Head with regular, dense and fine isodiametric microreticulation, coarser in middle portion; neck with isodiametric sculpture, denser and coarser than that in middle portion of head; pronotum with dense isodiametric microsculpture similar to that in head, finer in middle; scutellum with dense transverse meshes in middle; abdomen with dense and moderately fine transverse microreticulation. + +Head 1.5 times as broad as long, with markedly elevated portion between anterior margins of eyes and infraorbital portions; supra-antennal protuberances moderately wide and distinctly elevated; medioapical depression deep and wide, narrowing posteriad and indistinctly connected with moderately deep subrectangular interocellar depression; anteocellar foveae narrow and long, deep, convergent diagonally toward level of anterior margins of eyes; temples convex and slightly shorter than longitudinal length of eyes. Eyes moderately small, strongly convex. Punctation dense and moderately fine, sparser and finer in middle. Ocelli small, located distinctly below level of posterior margins of eyes, distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi short, about three times shorter than preapical segment. Antenna long, almost reaching apical margins of elytra when reclined; basal antennomere distinctly more than three times as long as broad, antennomere 2 narrower and significantly shorter than basal antennomere, 3 distinctly longer than 2, 4–5 slightly longer than 3, 6–9 indistinctly longer and broader than 5, 10 slightly shorter than 9, apical antennomere about 1.3 times as long as 10. + +Pronotum moderately convex, 1.3 times as broad as long, 1.2–1.3 times as broad as head, widest in anterior portion, strongly narrowed posteriad toward obtuse posterior angles, with indistinctly elevated longitudinal band in middle ( +holotype +); posterior margin rounded, about as long as posterior margin slightly concave in middle.Punctation about as that in middle portion of head, but distinctly denser, sparser in median part, with narrow impunctate portion in middle. Scutellum with several fine and sparse punctation. + + + +FIGURES 1–2. +Habitus of + +Hygrodromicus + +: 1– + +H. incrassatus + +(paratype), 2– + +H. splendidus + +(Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm. + + + +Elytra flattened, 1.2 times as broad as long, 1.6–1.7 times as long as pronotum, strongly broadened posteriad, with straight or widely rounded apical margins. Punctation dense, distinctly larger and deeper than that on pronotum, denser and finer around scutellum, with interstices between punctures in middle about as long as diameter of one-two nearest puncture; +paratypes +with finer and sparser punctation along suture. Hind wings fully developed. + +Abdomen distinctly broader than elytra, with two small and transverse tomentose spots in the middle of abdominal tergite V; apical margin of abdominal tergite VII with narrow palisade fringe. Punctation dense and fine. + +Male. Protarsomeres 1–4 wide. Apical portion of pronotum wide. Apical margin of abdominal tergite VIII somewhat straight. Apical margin of abdominal sternite VIII sinuate. Aedeagus with small narrow basal portion and elongate moderately wide median lobe, strongly narrowed in apical third, slightly broadened in preapical portion and narrowed toward rounded apex; parameres narrow, almost reaching apex of median lobe, each with four short apical setae; internal sac moderately long with two pairs of narrow elongate sclerotized structures and long narrow flagellum between them, spirally folded in basal portion ( +Fig. 3 +). Lateral aspect of the aedeagus as in +Fig. 4 +. + +Female. Protarsomeres 1–4 moderately narrow. Apical portion of pronotum relatively narrow. Apical margins of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII somewhat rounded. + +Comparative notes. +Based on the general shape of the body and the aedeagus, + +H. incrassatus + + +sp. n. + +is similar to + +H. wrasei +Zerche, 1992 + +, known from +Tajikistan +( +Zerche 1992 +, +Shavrin 2024 +). It can be distinguished from it by the larger body, the presence of ocelli, longer antennomeres 6–10, more transverse pronotum (especially in male) with more narrowed laterobasal margins, longer median lobe of the aedeagus with different shape of the apical portion, and the presence of the flagellum (missing in + +H +. +wrasei + +). Based on the general shape of the aedeagus, + +H. incrassatus + + +sp. n. + +is also similar to + +H. cachemiricum +Coiffait, 1983 + +( +India +: Kashmir), + +H. klapperichi +Coiffait, 1982 + +( +Afghanistan +), and Middle Asian + +H. reitteri +(Scheerpeltz, 1933) + +and + +H. splendidus + +. It can be distinguished from these species by the following morphological features: from + +H. cachemiricus + +by the broader pronotum, narrower and longer median lobe, with shorter apical portion; from + +H. klapperichi + +and + +H. reitteri + +by the distinctly broader pronotum and elytra, shorter median lobe with different shape of the apical portion; from + +H. splendidus + +by narrower antennae and the less convex pronotum, the longer parameres and different shape of the apical portion of the median lobe. From all these species + +H. incrassatus + + +sp. n. + +can be distinguished by the details of the internal structure of the aedeagus. + + + + +Distribution. + +Hygrodromicus incrassatus + + +sp. n. + +is known only from the +type +locality in northern +Pakistan +( +Fig. 7 +). + + +Bionomics. +The +type +specimens were collected at an elevation of +2200 m +a.s.l. The detailed bionomical data remains unknown. + + + + +Etymology. +The specific epithet is the Latin + +incrassatus + +, - +a +, - +um +(thickened). It alludes to the wide pronotum of the male. + + + + \ No newline at end of file diff --git a/data/A6/4A/2B/A64A2B29256F163542FEFF4EFCB09B35.xml b/data/A6/4A/2B/A64A2B29256F163542FEFF4EFCB09B35.xml new file mode 100644 index 00000000000..2e024a173cf --- /dev/null +++ b/data/A6/4A/2B/A64A2B29256F163542FEFF4EFCB09B35.xml @@ -0,0 +1,351 @@ + + + +On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +384 +390 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.7 + +journal article +10.11646/zootaxa.5481.3.7 +1175-5326 +12758944 +F86F4482-5F80-444F-A5DE-F1F1BAC04E41 + + + + + + + +Hygrodromicus splendidus +Zerche, 1992 + + + + + + + +( +Figs 2 +, +5–7 +) + + + + + + + +Hygrodromicus splendidus +Zerche, 1992: 116 + + + + + + + +Material examined. + + +KAZAKHSTAN +: +ALMATY +: + +9 ♂♂ +, +21 ♀♀ +: +Ketmen Ridge +, +Dolaity +R + +. + + +15.07.1988 + +. +V + +. +Kastcheev +leg. (cSh, +NMW +, +ZIN +); + +1 ♂ +, +1 ♀ +: same data, + +09.07.1988 + +(cSh) + +; + +1 ♂ +, +1 ♀ +: same data, +Avat +R + +. +14.07.2009 +(cSh); + +13 ♂♂ +, +6 ♀♀ +: same data, +Kulbastau +[=Kol`bastau] +R + +. + + +27.07.1988 + +(cSh, +ZIN +) + +; + +3 ♀♀ +: same data, +Bolshoy Kyrgyzsay. + +18.05.1991 + +(cSh) + +; + +1 ♀ +: same data, + +10.06.1988 + +(cSh) + +; + + +JAMBYL +: + +1 ♂ +: +Aksu-Dzhabagly +, +Koksay +R + +. + + +02.07.2001 + +. +V + +. Kastcheev leg. (cSh); + + +KYRGYZSTAN +: +CHÜY +: + +1 ♀ +: +Kegety Gorge. + +09.06.1994 + +. +S. Ovchinnikov +leg. (cSh) + +. + + + + +Redescription. +Measurements (n = 50): HW: 0.94–1.01; HL: 0.74–0.75; OL: 0.25–0.26; LT: 0.17–0.18; AL (averaged): 3.46; PL: 0.88–0.94; PWmax: 1.33–1.41; PWmin: 1.23–1.25; ESL: 1.47–1.63; EW: 2.07–2.35; MTbL(averaged): 1.55; MTrL (averaged): 0.34 (MTrL 1–4: 0.19; MTrL 5: 0.15); AW: 2.13–2.36; AedL: 1.28–1.34; BL: 5.45–7.00 ( +holotype +: +Zerche (1992)) +. + + + +FIGURE 7. +Distribution of + +Hygrodromicus incrassatus + +(square) and + +H. splendidus + +(circles). + + + +Habitus as in +Fig. 2 +. Body reddish-brown to black, usually with distinctly paler elytra; mouthparts, antennae and legs brownish; apical segment of maxillary palpi and tarsi yellowish. Head with dense and fine isodiametric microculpture, coarser on infraorbital portions; neck with dense transverse or isodiametric sculpture; pronotum with dense and coarse isodiametric meshes, usually finer along middle and mediobasal third; scutellum with dense and fine transverse meshes; abdomen with dense and sometimes coarse isodametric or transverse microsculpture. + + +Head 1.2–1.3 times as broad as long, with slightly elevated middle portion; medioapical depression deep and wide, narrowing posteriad; interocellar depression wide, relatively shallow or deep; anteocellar foveae narrow and long, strongly convergent diagonally toward level of anterior margins of eyes. Punctation irregular, fine, sometimes invisible in some specimens, denser and slightly larger between eyes; neck without visible punctation or with several punctures in middle. Ocelli small and indistinct (invisible in +holotype +) or moderately large. Antenna reaching basal third or apical margins of elytra when reclined, with somewhat widened antennomeres; antennomere 3 slightly longer than 2, 4–5 slightly longer and usually indistinctly broader than 3, 6–10 slightly longer than 5 (sometimes antennomere 5 indistinctly shorter). + +Pronotum markedly convex and distinctly transverse, 1.5 times as broad as long, 1.3–1.4 times as broad as head, from widest anterior portion strongly narrowed posteriad toward obtuse or subacute posterior angles; anterior angles widely rounded, sometimes slightly protruded anteriad; posterior margin straight or rounded, slightly narrower than posterior margin usually widely concave in middle. Punctation fine and dense, more distinct than that on head, finer in middle and sometimes with narrow impunctate median band. Scutellum without or with several fine punctures in middle. +Elytra 1.1–1.3 times as broad as long. Punctation dense, distinctly larger and deeper than that on pronotum, with interstices between punctures in middle about one-two diameters of nearest punctures. + +Abdomen slightly narrower ( +holotype +) or broader than elytra, with two large and transverse tometose spots in the middle of abdominal tergite IV and two small spots in tergite V (some specimens witth indistinct spots); palisade fringe narrow or invisible. + + +Male.Apical margin of abdominal tergite VIII somewhat straight or rounded.Apical margin of abdominal sternite VIII sinuate. Aedeagus with small basal portion and moderately wide elongate median lobe, distinctly narrowed in apical third, slightly broadened in preapical portion and narrowed toward small rounded apex; parameres narrow, indistinctly broadened apically, usually not reaching apex of median lobe, with four short apical setae; internal sac relatively long, with two narrow sclerotized lobes in preapical portion and moderately long flagellum, spirally folded in basal portion ( +Fig. 5 +). Lateral aspect of the aedeagus as in +Fig. 6 +. + +Female. Apical margins of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. + +Comparative notes. +Based on the general shape of the aedeagus, + +H +. +splendidus + +is similar to + +H. cachemiricum + +, + +H. incrassatus + + +sp. n. + +, + +H. klapperichi + +, + +H. reitteri + +and + +H. wrasei + +. From all these species it can be distinguished by the broader antennae, more convex pronotum, and details of the external and internal morphology of the aedeagus. Additionally, it differs from + +H. cachemiricum + +, + +H. klapperichi + +and + +H. reitteri + +by the broader pronotum and elytra. + + + + +Distribution. + +Hygrodromicus splendidus + +is known from several locality in the Middle Asia: south-western +Kazakhstan +, north +Kyrgyzstan +and north +Tajikistan +( +Fig. 7 +). + + +Bionomics. +The +holotype +was collected at elevation +3000–4000 m +a.s.l. The detailed bionomical data is unknown. + + + + +Remarks. + +Hygrodromicus splendidus + +was originally described based on the +holotype +from +Tajikistan +(“… Pamir, Muksu Gebiet… Hochflâche bei Kishlak Ljarsch…”). The specimens studied by me are conspecific to the +holotype +, but have some insignificant morphological differences. Thus, I decided to redescribed this species. It is here recorded from +Kazakhstan +and +Kyrgyzstan +for the first time. + + + + \ No newline at end of file diff --git a/data/B8/7B/D2/B87BD23EE803FFA0FF55EF7FFDCB881B.xml b/data/B8/7B/D2/B87BD23EE803FFA0FF55EF7FFDCB881B.xml new file mode 100644 index 00000000000..e580d7aee6c --- /dev/null +++ b/data/B8/7B/D2/B87BD23EE803FFA0FF55EF7FFDCB881B.xml @@ -0,0 +1,691 @@ + + + +Two new species of the genus Sycanus Amyot & Serville (Insecta: Hemiptera: Reduviidae: Harpactorinae) from Vietnam + + + +Author + +Truong, Xuan Lam +0000-0002-1758-903X +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +txlam.iebr@gmail.com + + + +Author + +Giang, Phan Thi +0009-0006-3597-4077 +Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Vinh University, 182 Le Duan, Vinh, Nghe An, Vietnam +tragiangdhv8668@gmail.com + + + +Author + +Nguyen, Dai Dac +0009-0006-8024-2553 +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +daindiebr@gmail.com + + + +Author + +Chau, Tran Pham Minh +0009-0009-6264-0592 +Faculty of Biology, Vietnam National University - VNU University of Science, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam +tranphamminhchau_t66@hus.edu.vn + + + +Author + +Ha, Ngoc Linh +0000-0002-1723-1230 +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +linh.hangoc02@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +301 +325 + + + + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +journal article +10.11646/zootaxa.5481.3.1 +1175-5326 +12758808 +0803FD80-03E1-4751-BC89-0BC1D44CA26C + + + + + + + +Sycanus thuathienhuensis +Truong & Ha + +, +sp. nov. + + + + + + +( +Figs. 4A +, +6 +, +7 +) + + + + +Type material. + + +Holotype +. + +♂ +; TXL2024-012; +Vietnam +, +Thua Thien Hue Province +, +Bach Ma National Park +; + +02.ii.2024 + +; +XL Truong +leg.; +IEBR + +. + + +Paratypes +. + +1♂ +; TXL2018-004; +Vietnam +, +Gia Lai Province +, +Kon Chu Rang Nature Reserve +; + +05.v.2018 + +; +XL Truong +leg.; +IEBR + +. + +1♂ +; NDD2022-009; +Vietnam +, +Quang Nam Province +, +Tay Giang Forest +; + +01.v.2022 + +; +DD Nguyen +leg.; +IEBR + +. + +1♀ +; NDD2022-063; +Vietnam +, +Quang Nam Province +, +Tay Giang Forest +; + +1.v.2022 + +; +DD Nguyen +leg.; +IEBR + +. + +1♀ +; TXL2024-059; +Vietnam +, +Thua Thien Hue Province +, +Bach Ma National Park +; + +02.ii.2024 + +; +XL Truong +leg.; +IEBR + +. + +2♀ +; PTG2024-060; PTG2024-061; +Vietnam +, +Thua Thien Hue Province +, +Bach Ma National Park +; + +04.ii.2024 + +; +PT Giang +leg.; +IEBR + +. + +1♀ +; PTG2024-062; +Vietnam +, +Thua Thien Hue Province +, +Bach Ma National Park +; + +05.ii.2024 + +; +PT Giang +leg.; +IEBR + +. + + +Non-type material. + +1♂ +; TXL2016-514; +Vietnam +, +Gia Lai Province +, +Kon Chu Rang Nature Reserve +; + +26.iv.2016 + +; +XL Truong +leg.; +IEBR + +. + +1♀ +; TXL2016-521; +Vietnam +, +Gia Lai Province +, +Kon Chu Rang Nature Reserve +; + +27.iv.2016 + +; +XL Truong +leg.; +IEBR + +. + +1♀ +; TXL2016-604; +Vietnam +, +Dak Lak Province +, +Chu Yang Sin National Park +; + +4.v.2016 + +; +XL Truong +leg.; +IEBR + +. + +4♀ +; TXL2018-001; TXL2018-002; TXL2018-003; TXL2018-005; +Vietnam +, +Gia Lai Province +, +Kon Chu Rang Nature Reserve +; + +05.v.2018 + +; +XL Truong +leg.; +IEBR + +. + +2♀ +; TXL2018-043; TXL2018-044; +Vietnam +, +Dak Lak Province +, +Chu Yang Sin National Park +; + +08.v.2018 + +; +XL Truong +leg.; +IEBR + +. + +1♀ +; NDD2022-024; +Vietnam +, +Quang Nam Province +, +Tay Giang Forest +; + +02.v.2022 + +; +DD Nguyen +leg.; +IEBR + +. + + + + +Diagnosis. +Body ochreous; anterior pronotal lobe black with some rows of short bent cream-yellow short and slender setae ( +Fig. 6F +); posterior pronotal lobe ochreous, rugulose, punctured, covered with short erect slender setae ( +Fig. 6F +); scutellum black with yellow latero-posterior margin ( +Fig. 6F +); scutellar spine sub-vertical, short or almost tubercle-liked ( +0.44–1.53 mm +), not bifid at apex, yellow or brownish-yellow with black base ( +Fig. 6B, F +); coria yellow or ochreous with transection with clavus dark brown, clavus dark brown ( +Fig. 6G +); male pygophore brownish yellow with ventrally centrally largely suffused with dark brown or blackish brown ( +Fig. 6C +). + + +This species is very similar to + +Sycanus croceus + +in general appearance, especially in body colour, pronotum, and thoracic sterna. But the new species can be distinguished from + +S. croceus + +by a combination of the following characteristics: a brownish yellow stripe running between postero-upper corner of compound eye and ocellus of same side ( +Fig. 6E +) (in + +S. croceus + +without any yellow stripe), abdominal sternites with small longitudinal black markings segmentally laterally ( +Fig. 6C +) (in + +S. croceus + +with small round spots segmentally laterally), large marking of laterotergite V reach antero-lateral margin of laterotergite V (in + +S. croceus + +not extend to the antero-lateral margin), scutellar spine short or almost tubercle-liked, not bifid at apex, yellow or brownish-yellow with black base ( +Figs. 6B, F +) (in + +S. croceus + +long, apex bi-forked, wholly yellow or orange), distal dorsal lobe of endosoma (ddl) with two very large and long horn-shaped prickles centrally, surrounded by large prickles ( +Fig. 7I–K +) (in + +S. croceus + +ddl with two large horned processes, apical part laterally with three pairs of larger spines and eight pairs of smaller spines). + + +Moreover, the new species is also similar to + +Sycanus insularis + +in general morphology. But the former can be distinguished from the latter by the following features: abdominal sternites with small longitudinal black markings segmentally laterally ( +Fig. 6C +) (in + +S. insularis + +with small round spots segmentally laterally), scutellar spine short or almost tubercle-liked, not bifid at apex, yellow or brownish-yellow with black base ( +Fig. 6B, F +) (in + +S. insularis + +long, apex bifid, black or yellow), coria yellow or ochreous with transection with clavus dark brown, clavus dark brown brown ( +Fig. 6G +) (in + +S. insularis + +anterior half black or dark brown, remaining pale yellow or luteous). + + + +FIGURE 6. + +Sycanus thuathienhuensis +Truong & Ha + +, + +sp. nov. + +, +holotype, ♂, TXL2024-012. +A, +body in dorsal view; +B, +body in lateral view; +C, body +in ventral view; D, head in lateral view; +E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing. + + + +Furthermore, the new species has a very close relationship to + +Sycanus rufus + +in molecular phylogenetic analyses with a minimum interspecific divergence of 2.0% in p-distance and +0.02 in +K +2P. +However, the two species can be delimited by the external morphology, i.e., body ochreous (in + +S. rufus + +body red), posterior pronotal lobe ochreous, rugulose, punctured, covered with short erect slender setae ( +Fig. 6F +) (in + +S. rufus + +red, rugulose, not punctured, clothes with procumbent short yellow setae), distal dorsal lobe of endosoma (ddl) with two very large and long horn-shaped prickles centrally, surrounded by large prickles ( +Figs. 7I–K +) (in + +S. rufus + +ddl with two long and slender prickles, and laterally with two pairs of larger spines and about 10 pairs of smaller spines). + + +Male description. + + +Coloration. +Body ochreous. Head dorsum, neck, clypeus, maxillary plate, antenniferous tubercles, and posterior half of head venter black; labrum and anterior margin of gena dark brown; gena and anterior half of head venter luteous or yellow; a brownish yellow stripe running between postero-upper corner of compound eye and ocellus of same side; first visible labial segment blackish brown at basal half, paler as dark brown toward tip; base of second visible labial segment dark brown; remaining of labium brown, paler toward tip ( +Fig. 6D, E +). Scape, pedicel, and first flagellomere blackish brown; second flagellomere dark brown, paler toward tip. Collar, anterolateral angle, anterior margin of anterior pronotal lobe, acetabulum, prosternum, and stridulatory sulcus yellow ( +Fig. 6F +); posterior pronotal lobe yellow or ochreous ( +Fig. 6F +); scutellum black with yellow latero-posterior margin ( +Fig. 6F +); scutellar spine yellow or brownish-yellow with black base or wholly black ( +Fig. 6F +); meso- and metapleura, meso- and metasterna, black or blackish brown; coxae yellow; trochanters blackish brown; femora, tibiae, and tarsi black or blackish brown. Coria yellow or ochreous with transection with clavus dark brown; clavus dark brown; membrane pale bronzy brown, semi-hyaline ( +Fig. 6G +). Hind wings faintly semi-hyaline ( +Fig. 6H +). Abdominal mediotergites blackish brown or black with irregular brown or yellow suffusions; laterotergites yellow or orangish yellow with lateral half of red, reddish yellow, or orange ( +Fig. 6B +); laterotegites III–VII segmentally large black marking; large marking of laterotergites VI and V reach antero-lateral margin of latetotergites; large marking of laterotergites III, VI, and VII far from reaching the lateral margin of laterotergites; abdominal sternites yellow or luteous with broad transversal black strip in anterior margin of each segment and discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally ( +Fig. 6C +). Pygophore brownish yellow with ventrally centrally largely suffused with dark brown or blackish brown ( +Fig. 6C +). + + +Structure. +Body large-sized ( +17.40–19.08 mm +), elongated and posteriorly widened ( +Fig. 6A–C +). Head tubular, slender, elongated, head and neck together much longer than pronotum ( +Fig. 6A, B +); anteocular area of head elongate-conical; anteclypeus cylindrical and prominent; postocular area of head weakly globose, distinctly wider than anteocular area, shorter than anteocular area, constricted behind compound eyes, with a wide and deep interocular sulcus; neck long ( +Fig. 6D, E +). Compound eyes protruding laterally, nearly globose, with posterior margin sub-straight, oblique with respect to ventral margin of head; lateral ocelli produced, slightly elevated behind interocular sulcus, separated from each other; interspace between lateral ocelli wider than distance between compound eye and lateral ocellus ( +Fig. 6D, E +). First visible labial segment slightly thicker and much shorter than second segment, longer than anteocular area of head, not extending beyond level of middle of compound eye when labium laid backward ( +Fig. 6D +); proportional average length of first to third visible labial segments 2.4:3.7:0.6. Scape about 1.6 times as long as head, about 2.5 times as long as pedicel, almost as long as second flagellomere; first flagellomere slightly shorter than pedicel and about 1/3 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 7.0:2.9:2.3:6.8. Collar thick in dorsal view, with anterolateral angle roundly produced anteriorly; anterior pronotal lobe small, hemisphered and bulged, smooth, deeply depressed at base; posterior pronotal lobe rugulose and punctured, shallowly depressed on disc, with slightly swollen anteromedial elevation (never sulcate or concave); humerus bluntly triangular, with round apex; posterior margin of pronotum weakly concave, but slightly convex centrally; posterior angles round, slightly exceeded to posterior margin of pronotum ( +Fig. 6F +). Scutellum triangular, somewhat triangularly depressed basally, apically produced with a scutellar spine, and sloping downward, posterior apex round ( +Fig. 6F +); scutellar spine sub-vertical, short or almost tubercle-liked ( +0.44–1.21 mm +), not bifid at apex ( +Fig. 6B +). Femora elongated, slender but stout, moderate subnodulose apically; tibiae slender and elongated. Hemelytra surpassing beyond apex of abdomen when fully closed, 0.7 times as long as body length ( +Fig. 6A +); discal cell nearly diamond-shaped, longer than width ( +Fig. 6G +); Sc 0.8 times as long as hemelytron length, 1.4 times as long as R + M ( +Fig. 6G +). Hind wing about 3.0 times as long as maximum width ( +Fig. 6H +). Laterotergites much dilated and ascending with segmental incisures, posto-lateral margin of each laterotegite slightly produced than antero-lateral margin of following laterotegite ( +Fig. 6A–C +). Pygophore ovoid ( +Fig. 7B–D +); median process of pygophore (mpp) posteriorly produced, 0.52 times as long as wide in dorsal view, with apical margin convex, distinctly produced a round apex, apicolateral corner sub-round, distinctly produced posterolaterad ( +Fig. 7B–E +); paramere long, slender, clavate, somewhat incurved and larger in apical part, with round apex ( +Fig. 7F, G +). Aedeagus in dorsal view ovoid, dorsally sclerotized, and in lateral view long and narrow ( +Fig. 7H +); articulatory apparatus (aa) in ventral view with basal plate arms relatively slender and jointly forming a V-shape, and in lateral view arched strongly ( +Fig. 7H, I +); dorsal phallothecal sclerite (dps) in lateral view with posteromedian weakly produced ( +Fig. 7I +); spoon-like sclerites (sps) largely anteriorly produced, hyaline and glabrous ( +Fig. 7H, I +); membranous sac-like lobes almost disappeared ( +Fig. 7H +); distal dorsal lobe of endosoma (ddl) round and weakly bulged, with two very large and long horn-shaped prickles centrally, surrounded by large prickles ( +Fig. 7H–K +). + + +Vestiture. +Body clothed with pale, shining griseous, slender setae. Head covered with very short bent slender setae, interleaved with short erect slender setae; labium almost glabrous except base of first visible segment with a few tiny slender setae ( +Fig. 6D, E +). Scape and pedicel covered with short slender sub-erect setae; remaining antennae covered with short vertical setae, denser toward tip. Collar, anterolateral angles with tiny bent setae abundantly; anterior pronotal lobe with some rows of tiny bent setae, somewhat interleave with slender erect setae; posterior pronotal lobe covered with short slender setae and posteriorly cover with slender setae; scutellum densely covered with long, slender, vertical, erect black setae ( +Fig. 6F +). Coxae and trochanters covered with short bent setae; femora, tibiae, and tarsi densely covered with long, slender, erect setae. Coria and clavus densely covered with short bent setae. Abdominal mediotergites sparsely covered with slender erect setae; lateral margins of laterotergites densely covered with short erect setae; abdominal sternites and laterotergites covered with short oblique cream-yellow setae, somewhat interleaved with long slender erect setae. Male genitalia pygophore ventrally covered with short slender bent setae and posteriorly densely covered with short and long slender setae ( +Fig. 7B–E +); paramere almost glabrous except posterior apex densely covered with short and long thick erect setae ( +Fig. 7F, G +). + + +Female description. + +General external morphology similar to that of the male. + +Coloration. +Almost similar to male but differ in the following characters. Abdominal sternite VII yellow with dark brown suffusion near posterior margin, and blackish brown or back posterior margin; Gonocoxa VIII yellow with inner margin black; abdominal tergite IX and gonapophysis VIII black; abdominal laterotergite VIII orange or sanguineous ( +Fig. 7A +). + + +Structure. +Almost the same as male but larger than male and differ in the following characters. Body large-sized ( +22.18–24.87 mm +), elongated, and somewhat robust. Head shorter than or almost as long as pronotum. Proportional average length of first to third visible labial segments 2.6:4.2:0.7. Scape about 1.9 times as long as head, about 2.4 times as long as pedicel, slightly longer than second flagellomere; first flagellomere slightly shorter than pedicel and about 1/3 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 7.3:3.0:2.5:6.7. Abdominal laterotergite VIII with thin posterior margin, not exceed the posterior margin of abdominal tergite IX; abdominal sternite VII forming a semi-circular or wide sub-pentagonal median concavity, with posteromedian margin gently U-shaped, with inner posterolateral margin slightly concave; gonocoxa VIII subtriangular, produced mesad and forming a pointed apex at apical inner corner, and with inner margin weakly incurved in posterior 2/3; gonapophysis VIII subtriangular, posteriorly produced; abdominal tergite IX small ( +Fig. 7A +). + + +Vestiture. +Almost the same as male except for the following characters.Anterior 1/3 of gonocoxa VIII almost no setae; posterior 2/3 of gonocoxa VIII covered with short and long bent black setae; posterior margin of gonapophysis VIII densely covered with long bent black setae; abdominal laterotergite VIII and abdominal tergite IX covered with short and long bent black setae ( +Fig. 7A +). + + + + +FIGURE 7. + +Sycanus thuathienhuensis +Truong & Ha + +, + +sp. nov. + +, A, female genitalia in ventral view, paratype, ♀, NDD2022-063; B–K, male genitalia, +holotype, ♂, TXL2024-012. +B–E, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; F–G, left paramere; F, dorsal view; G, lateral view; H–J, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl). + + + + +Measurements. +All dimensions are given in mm. + +Holotype +( +♂ +): + +BL 18.89; HL 3.75; AoL 1.73; AoW 0.87; PoL 1.12; PoW 1.27; NL 1.17; OE 1.78; IE 0.70; ED 1.09; OD 0.22; OCD 0.36; COD 0.20; R1L 2.50; R2L 3.88; R3L 0.61; A1L 7.79; A2L 2.88; A3L 2.30; A4L 7.06; PnL 3.62; PnW 4.26; APL 1.31; PPL 2.30; SPL 0.46; HeL 13.84; HeW 4.58; Sc 10.23; R+M 7.38; HWL 9.63; HWW 3.17; AFL 7.09; ATL 8.29; MFL 6.34; MTL 7.89; PFL 8.41; PTL 11.08. + +Paratype +( +♂ +): + +BL 19.08; HL 3.61; AoL 1.62; AoW 0.88; PoL 1.10; PoW 1.23; NL 1.14; OE 1.69; IE 0.69; ED 1.03; OD 0.19; OCD 0.46; COD 0.20; R1L 2.32; R2L 3.73; R3L 0.60; A1L 6.82; A2L 3.08; A3L 2.15; A4L 6.60; PnL 3.52; PnW 4.22; APL 1.20; PPL 2.31; SPL 1.21; HeL 13.49; HeW 4.04; Sc 9.95; R+M 6.00; AFL 7.72; ATL 8.19; MFL 6.25; MTL 7.63; PFL 8.10; PTL 10.44. + +Paratype +( +♀ +): + +BL 22.18–24.87; HL 3.81–4.23; AoL 1.85–2.04; AoW 0.94–1.04; PoL 0.95–1.23; PoW 1.25–1.46; NL 1.35–1.42; OE 1.73–1.87; IE 0.76–0.84; ED 1.04–1.10; OD 0.21–0.28; OCD 0.45–0.58; COD 0.18–0.21; R1L 2.50–2.72; R2L 4.03–4.36; R3L 0.63–0.81; A1L 6.83–7.82; A2L 3.04–3.08; A3L 2.26–2.66; A4L 6.47–7.03; PnL 3.71–4.74; PnW 5.14–5.63; APL 1.03–1.45; PPL 2.69–3.27; SPL 0.96–1.53; HeL 16.05–18.06; HeW 4.67–5.13; Sc 11.69–12.98; R+M 7.93–8.83; AFL 7.33–8.08; ATL 8.58–9.00; MFL 6.19–6.71; MTL 7.89–8.40; PFL 8.11–9.22; PTL 11.37–12.05. + + + + +Distribution. +Vietnam +, Middle and Southern +Vietnam +(Thua Thien Hue, +Quang Nam +, +Binh Dinh +, +Gia Lai +, +Dak Lak +Provinces). + + + + + + +Type +locality. + +Vietnam +, +Thua Thien Hue +, +Bach Ma National Park + +. + + + + +Etymology. +The specific epithet refers to its occurrence in Thua Thien Hue, the North Central Coast region, the Central of +Vietnam +; an adjective. + + + + \ No newline at end of file diff --git a/data/B8/7B/D2/B87BD23EE81EFFA4FF55EE6BFE528DB3.xml b/data/B8/7B/D2/B87BD23EE81EFFA4FF55EE6BFE528DB3.xml new file mode 100644 index 00000000000..0abb19bfe24 --- /dev/null +++ b/data/B8/7B/D2/B87BD23EE81EFFA4FF55EE6BFE528DB3.xml @@ -0,0 +1,492 @@ + + + +Two new species of the genus Sycanus Amyot & Serville (Insecta: Hemiptera: Reduviidae: Harpactorinae) from Vietnam + + + +Author + +Truong, Xuan Lam +0000-0002-1758-903X +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +txlam.iebr@gmail.com + + + +Author + +Giang, Phan Thi +0009-0006-3597-4077 +Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Vinh University, 182 Le Duan, Vinh, Nghe An, Vietnam +tragiangdhv8668@gmail.com + + + +Author + +Nguyen, Dai Dac +0009-0006-8024-2553 +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +daindiebr@gmail.com + + + +Author + +Chau, Tran Pham Minh +0009-0009-6264-0592 +Faculty of Biology, Vietnam National University - VNU University of Science, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam +tranphamminhchau_t66@hus.edu.vn + + + +Author + +Ha, Ngoc Linh +0000-0002-1723-1230 +Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam +linh.hangoc02@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +301 +325 + + + + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +journal article +10.11646/zootaxa.5481.3.1 +1175-5326 +12758808 +0803FD80-03E1-4751-BC89-0BC1D44CA26C + + + + + + + +Sycanus taynguyenensis +Truong & Ha + +, +sp. nov. + + + + + + +( +Figs. 4B +, +8 +, +9 +) + + + + +Type material. + + +Holotype +. + +♂ +; NDD2019-285; +Vietnam +, +Dak Lak Province +, +Ea So Natural Reserve +; + +19.ix.2019 + +; +DD Nguyen +leg.; +IEBR + +. + + +Paratypes +. + +1♀ +; NDD2018-395; +Vietnam +, +Gia Lai Province +, +Kon Chu Rang Natural Reserve +; + +26.ix.2018 + +; +DD Nguyen +leg.; +IEBR + +. + + +Non-type material. + +1♂ +; Qhoa-M10-01; +Vietnam +, +Dak Lak Province +, +Krong Ana +, +Dray Sap +; + +30.iii.2019 + +; +Nguyen Thi Quynh Hoa +(Tay Nguyen University) leg.; +IEBR + +. + +1♂ +; TXL-M10-02; +Vietnam +, +Dak Lak Province +, +Ea So Natural Reserve +; + +15.ix.2023 + +; +XL Truong +leg.; +IEBR + +. + + + + +Diagnosis. +Body large-sized and pale orange brown; head long, almost black with a brownish yellow suffusion next to outer side of ocellus ( +Fig. 8D, E +); base of scape blackish brown, scape interleaved brownish yellow and blackish brown, pedicel dark brown with a brown suffusion medially, first and second flagellomeres brown; anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe rugulose, pale orange brown or reddish brown ( +Fig. 8F +); scutellum black with orange brown centrally and scutellar spine sub-vertical, short, bifid at apex, and wholly orange brown or reddish brown ( +Fig 8B, F +); basal 2/3 of coria and whole clavus black or dark brown; remaining of coria white or orange; clavus dark brown ( +Fig. 8G +). + + +The general appearance and male genitalia of this species are highly similar to that of the + +Sycanus sichuanensis, +However + +, this new species can be distinguished from the latter by the following features: pronotum with anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe pale orange brown or reddish brown ( +Fig. 8F +) (in + +S. sichuanensis + +pronotum wholly black); laterotergites much dilated, posto-lateral margin of each laterotegite elevated, slightly produced than antero-lateral margin of following laterotegite ( +Fig. 8A–C +) (in + +S. sichuanensis + +laterotergites distinctly roundly laterally dilated, middle part of each laterotergite elevated); basal 2/3 of coria and whole clavus black or dark brown and remaining of coria white or orange ( +Fig. 8G +) (in + +S. sichuanensis + +coria yellowish to milk-white with basal part and apical angle black); coxae orange brown or reddish brown (in + +S. sichuanensis + +coxae black). + + +Moreover, the new species and + +Sycanus minor + +have high similarity in general appearance, i.e., male genitalia structure and general body coloration, and a very close relationship in molecular phylogenetic analyses (minimum interspecific p-distance of 1.8 %). However, the new species is distinct from + +S. minor + +by the following features: pronotum with anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe pale orange brown or reddish brown ( +Fig. 8F +) (in + +S. minor + +pronotum black with lateral and posterior margins orange or anterior, lateral and posterior margins orange); abdominal sternites dark brown with discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally ( +Fig. 8C +) (in + +S. minor + +yellowish to red with median longitudinal stripe, segmental transversal stripe black); base of scape blackish brown, scape interleaved brownish yellow and blackish brown, pedicel dark brown with a brown suffusion medially, first and second flagellomeres brown (in + +S. minor + +black), distal dorsal lobe of endosoma (ddl) centrally covered longitudinally with tiny prickles, surrounded with four large and long horn-shaped prickles centrally, and a few smaller prickles ( +Fig. 9H–K +) (in + +S. minor + +ddl with two feebly sclerotized stripe-shaped sclerites, apically with a feebly sclerotized horned process and laterally with five pairs of small spines). + + + +FIGURE 8. + +Sycanus taynguyenensis Truong & Ha +, + + +sp. nov. + +, +holotype, ♂, NDD2019-285. +A, +body in dorsal view; +B, +body in lateral view; +C, body +in ventral view; D, head in lateral view; +E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra. + + + +Male description. + + +Colouration. +Body pale orange brown. Head dorsum, neck, clypeus, antenniferous tubercles, labrum, and head venter black; maxillary plate, gena black or blackish brown; a brownish yellow suffusion next to outer side of ocellus; first visible labial segment blackish brown; base of second visible labial segment dark brown; remaining of labium brown, paler as pale brown toward tip ( +Fig. 8D, E +). Base of scape blackish brown; scape alternating brownish yellow and blackish brown; pedicel dark brown with a brown suffusion medially; first and second flagellomeres brown. Collar pale orange brown or pale reddish brown ( +Fig. 8F +); anterolateral angle, anterior pronotal lobe and prosternum orange brown, somewhat with irregular dark brown suffusion ( +Fig. 8F +); anterior acetabulum orange brown; stridulatory sulcus luteous; posterior pronotal lobe pale orange brown or reddish brown ( +Fig. 8F +); scutellum black with orange brown centrally ( +Fig. 8F +); scutellar spine wholly orange brown or reddish brown ( +Fig. 8F +); meso- and metapleura, meso- and metasterna, dark brown with irregular blackish brown sufusions; coxae orange brown or reddish brown; trochanters and basal half of femora dark brown; apical half of femora, tibiae, and tarsi blackish brown. Basal 2/3 of coria and whole clavus black or dark brown; remaining of coria white or orange; clavus dark brown; membrane pale bronzy brown, semi-hyaline ( +Fig. 8G +). Hind wings faintly semi-hyaline. Abdominal mediotergites dark brown with irregular brown suffusions; laterotergites sanguineous or reddish orange ( +Fig. 8A, B +); each laterotegites III–VII with large black markings in anterior and posterior margins (( +Fig. 8B +); abdominal sternites dark brown with discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally ( +Fig. 8C +). Pygophore orange or sanguineous ( +Fig. 8C +). + + +Var. +abdomen beneath black. + + +Structure. +Body large-sized ( +15.50 mm +), elongated and posteriorly widened. Head tubular, slender, elongated ( +Fig. 8D, E +); head and neck together much longer than pronotum ( +Fig. 8A, B +); anteocular area of head elongate-conical; anteclypeus cylindrical and prominent; postocular area of head weakly globose, distinctly wider than anteocular area, shorter than anteocular area, constricted behind compound eyes, with a wide and deep interocular sulcus; neck long ( +Fig. 8D, E +). Compound eyes protruding laterally, nearly globose, oblique with respect to ventral margin of head; lateral ocelli produced, slightly elevated behind interocular sulcus, separated from each other; interspace between lateral ocelli wider than distance between compound eye and lateral ocellus ( +Fig. 8D, E +). First visible labial segment slightly thicker and much shorter than second segment, longer than anteocular area of head, not extending beyond level of middle of compound eye when labium laid backward ( +Fig. 8D +); proportional length of first to third visible labial segments 2.2:3.7:0.7. Scape about 1.7 times as long as head, about 2.4 times as long as pedicel, longer than second flagellomere; first flagellomere about half as long as pedicel and about 1/4 times as long as second flagellomere; proportional length of scape, pedicel, first and second flagellomeres 5.5:2.3:1.3:4.8. Collar thin, almost invisible in dorsal view, with anterolateral angle roundly produced anteriorly; anterior pronotal lobe small, hemisphered and bulged, smooth, deeply depressed at base; posterior pronotal lobe rugulose, shallowly depressed on disc, with slightly swollen anteromedial elevation (never sulcate or concave); humerus bluntly triangular, with round apex; posterior margin of pronotum substraight; posterior angles round, weakly exceeded to posterior margin of pronotum ( +Fig. 8F +). Scutellum triangular, somewhat triangularly depressed basally, apically produced with a scutellar spine, and sloping downward, posterior apex round ( +Fig. 8F +); scutellar spine sub-vertical, short ( +1.45 mm +), bifid at apex ( +Fig. 8B, F +). Femora elongated, slender but stout, moderate subnodulose apically; tibiae slender and elongated. Hemelytra surpassing beyond apex of abdomen when fully closed, 0.7 times as long as body length ( +Fig. 8A +); discal cell nearly diamond-shaped, longer than width ( +Fig. 8G +); Sc 0.7 times as long as hemelytron length, 1.4 times as long as R + M ( +Fig. 8G +). Hind wing damaged. Laterotergites much dilated and ascending with segmental incisures, posto-lateral margin of each laterotegite slightly produced than antero-lateral margin of following laterotegite ( +Fig. 8B +). Pygophore ovoid ( +Fig. 9B–D +); median process of pygophore (mpp) posteriorly produced, 0.39 times as long as wide in dorsal view, with apical margin slightly convex, distinctly produced a round apex, apicolateral corner sub-round, distinctly produced posterolaterad ( +Fig. 9B–E +); paramere long, slender, clavate, somewhat incurved and larger in apical part, with round apex ( +Fig. 9F–G +). Aedeagus in dorsal view ovoid, dorsally sclerotized and in lateral view long and narrow ( +Figs. 9H, I +); articulatory apparatus (aa) in ventral view with basal plate arms relatively slender and jointly forming a V-shape, and in lateral view arched strongly ( +Fig. 9H, I +); dorsal phallothecal sclerite (dps) in lateral view with posteromedian weakly produced ( +Fig. 9I +); spoon-like sclerites (sps) weakly anteriorly produced, hyaline and glabrous ( +Fig. 9H, I +); membranous sac-like lobes almost disappeared ( +Fig. 9H +); distal dorsal lobe of endosoma (ddl) round and weakly bulged, centrally covered longitudinally with tiny prickles, surrounded with four large and long horn-shaped prickles centrally, and a few smaller prickles ( +Fig. 9H–K +). + + + +FIGURE 9. + +Sycanus taynguyenensis Truong & Ha +, + + +sp. nov. + +, A, female genitalia in ventral view, paratype, ♀, EG2018-395; B–K, male genitalia, +holotype, ♂, NDD2019-285. +B–E, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; F–G, left paramere; F, dorsal view; G, lateral view; H–J, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl). + + + +Vestiture. +Body clothed with yellow, shining, griseous setae interleaved with black setae. Head with moderately densely bent setae ( +Fig. 8D, E +); first visible labial segment somewhat with a few bent setae; remaining labium without setae ( +Fig. 8D +). Scape with some short, slender, oblique setae, denser apically; pedicel with short, oblique, setae, denser than first joint, and denser apically; first and second flagellomeres with abundantly short, slender, more oblique than second joint setae. Collar, anterolateral angles, posterior pronotal lobe, scutellum, pleuron, sternum, coxae, coria with dense of bent, yellowish griseous setae, interleaved with long, erect, vertical, black setae ( +Fig. 8H +); abdomen beneath with straight, slender, oblique griseous setae; trochanter, femora, tibia with long, slender, erect, vertical, black setae; Male pygophore ventrally moderately densely with bent, slender, yellowish griseous setae, somewhat with black setae ( +Fig. 9B–E +); paramere apically with some long, yellowish griseous setae ( +Fig. 9F, G +). + + +Female description. + +General external morphology similar to that of the male. + +Coloration. +Almost similar to male but differ in the following characters. Abdominal sternites wholly black; Gonocoxa VIII black with a reddish-brown spot near posterior apex; abdominal laterotergite VIII, abdominal tergite IX, and gonapophysis VIII black ( +Fig. 9A +). + + +Structure. +Almost the same as male but larger than male and differ in the following characters. Body large-sized ( +21.34 mm +), elongated, and somewhat robust. Head together with neck shorter than or almost as long as pronotum. Proportional length of first to third visible labial segments 2.2:4.1:0.9. Scape about 1.8 times as long as head, about 2.5 times as long as pedicel, longer than second flagellomere; first flagellomere about half as long as pedicel and about 1/5 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 6.4:2.5:1.3:5.9. Abdominal laterotergite VIII with thin posterior margin, not exceed the posterior margin of abdominal tergite IX; abdominal sternite VII forming a wide sub-pentagonal median concavity, with posteromedian margin gently U-shaped, with inner posterolateral margin slightly concave but weakly convex medially; gonocoxa VIII subtriangular, produced mesad and forming a pointed apex at apical inner corner, and with inner margin weakly incurved in posterior 3/4; gonapophysis VIII subtriangular, posteriorly produced; abdominal tergite IX small ( +Fig. 9A +). + + +Vestiture. +Almost the same as male except for the following characters. Anterior 1/3 of gonocoxa VIII almost no setae; posterior 2/3 of gonocoxa VIII covered with short and long slender black setae; posterior margin of gonapophysis VIII densely covered with long erect black setae; abdominal laterotergite VIII and abdominal tergite IX covered with short and long black setae ( +Fig. 9A +). + + + + +Measurements. +All dimensions are given in mm. + +Holotype +( + +♂ +): +BL 15.50; HL 3.29; AoL 1.47; AoW 0.77; PoL 1.02; PoW 1.08; NL 1.16; OE 1.42; IE 0.63; ED 0.89; OD 0.16; OCD 0.43; COD 0.17; R1L 2.22; R2L 3.68; R3L 0.71; A1L 5.47; A2L 2.25; A3L 1.33; A4L 4.80; PnL 2.82; PnW 3.36; APL 0.87; PPL 1.96; SPL 1.45; HeL 10.39; HeW 3.52; Sc 7.66; R+M 5.32; AFL 5.60; ATL 6.31; MFL 5.37; MTL 6.21; PFL 6.79; PTL 8.30. + +Paratype +( + +♀ +): +BL 21.34; HL 3.50; AoL 1.74; AoW 0.89; PoL 0.95; PoW 1.16; NL 1.37; OE 1.56; IE 0.70; ED 0.91; OD 0.15; OCD 0.42; COD 0.15; R1L 2.21; R2L 4.07; R3L 0.90; A1L 6.39; A2L 2.53; A3L 1.32; A4L 5.86; PnL 3.89; PnW 4.54; APL 1.17; PPL 2.72; SPL 2.31; HeL 12.24; HeW 4.58; Sc 9.43; R+M 6.53; AFL 6.20; ATL 7.24; MFL 5.63; MTL 6.87; PFL 7.36; PTL 9.67. + + + + +Distribution. +Vietnam +, Central Highlands ( +Dak Lak +and +Gia Lai +Provinces). + + + + + + +Type +locality. + +Vietnam +, +Central Highlands +( +Dak Lak +and +Gia Lai +Provinces +) + +. + + + + +Etymology. +This new species is named after the Tay Nguyen region, the local name of Central Highlands, +Vietnam +; an adjective. + + + + \ No newline at end of file diff --git a/data/CE/6E/87/CE6E879DE12CFF94FF6CFA81FB8CA045.xml b/data/CE/6E/87/CE6E879DE12CFF94FF6CFA81FB8CA045.xml new file mode 100644 index 00000000000..842ea48f6bd --- /dev/null +++ b/data/CE/6E/87/CE6E879DE12CFF94FF6CFA81FB8CA045.xml @@ -0,0 +1,1308 @@ + + + +Sycetta papillata sp. nov. (Porifera, Calcarea): the first record of Sycetta to the Southwestern Atlantic Ocean + + + +Author + +Mattedi, Alice + + + +Author + +Cavalcanti, Fernanda F. + +text + + +Zootaxa + + +2024 + +2024-07-17 + + +5481 + + +3 + + +353 +362 + + + + +http://dx.doi.org/10.11646/zootaxa.5481.3.4 + +journal article +10.11646/zootaxa.5481.3.4 +1175-5326 +12758841 +2400DCEF-B04F-480A-B248-038A27C8601E + + + + + + + +Sycetta papillata + +sp. nov. + + + + + + +( +Figs 1–3 +; +Tables 1 +, +2 +) + + + + +Diagnosis: + +Sycetta + +with only one category of diactines and atrial region formed exclusively by tetractines. + + + + +Etymology: +The name is related to the body formed by papillae (i.e., the radial tubes separated from each other). + + + + +Type material: + +UFBAPOR 1917 [ +Holotype +. +Camaçari +, +Bahia +, +Brazil +( +12°46'40" S +, +38°07'50" W +); collected by +W.Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1922 [ +Paratype +. Camaçari, +Bahia +, +Brazil +( +12°45'19" S +, +38°06'07" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +] + +. + + +Additional material: + +UFBAPOR 1645 [ +Stella Maris +, Salvador, +Bahia +, +Brazil +( +12°59'00" S +, +38°12'00" W +); collected by +E. Hajdu +; + +21/I/1997 + +; 0.5 to 1.0 m depth]. UFBAPOR 1918 [ +Camaçari +, +Bahia +, +Brazil +( +12°46'40" S +, +38°07'50" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1948 [Camaçari, +Bahia +, +Brazil +( +12°46'40" S +, +38°07'50" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1919 [Camaçari, +Bahia +, +Brazil +( +12°46'16" S +, +38°07'01" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1921 [Camaçari, +Bahia +, +Brazil +( +12°46'16" S +, +38°07'01" W +); collected by +W.Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1924 [Camaçari, +Bahia +, +Brazil +( +12°46'16" S +, +38°07'01" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 1923 [Camaçari, +Bahia +, +Brazil +( +12°46'06" S +, +38°07'15" W +), collected by +W. Andrade +; + +28/XI/1995 + +, + +25 m +depth + +]. UFBAPOR 1947 [Camaçari, +Bahia +, +Brazil +( +12°43'17" S +, +38°02'19.92" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +]. UFBAPOR 5022 [Camaçari, +Bahia +, +Brazil +( +12°45'19" S +, +38°06'07" W +); collected by +W. Andrade +; + +28/XI/1995 + +; + +25 m +depth + +] + +. + + + + +Type +locality: + +Camaçari +, +Bahia +, +Brazil +[Marine ecoregion: Northeastern +Brazil +( + +Spalding +et al +. 2007 + +)] + +. + + + +FIGURE 1. +External morphology of preserved specimens of + +Sycetta papillata + + +sp. nov. + +A—UFBAPOR 1645. B—UFBAPOR 5022. C—UFBAPOR 1922 (paratype). D—Detail of the oscular membrane (UFBAPOR 1922). E—Detail of the surface formed by papillae (UFBAPOR 1645). F—Detail of the base of UFBAPOR 1917 (holotype). + + + + +TABLE 1 +. Measurements of spicules of + +Sycetta papillata + + +sp. nov. + +(H—holotype; Pa—paratype; P—paired; U—unpaired; + +A—apical). + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
Spicules/ specimensActinesLength (µm)Width (µm)N
MinMeanSDMaxMeanSD
Diactines
UFBAPOR1917 (H)276.9410.280.9525.87.21.420
UFBAPOR1922 (Pa)130.6198.049.5337.95.40.720
Triactines of the distal cones
UFBAPOR1917 (H)P41.167.716.498.94.70.820
UFBAPOR1922 (Pa)61.982.811.8105.66.70.820
UFBAPOR1917 (H)U47.883.530.6142.15.60.920
UFBAPOR1922 (Pa)78.5123.121.2168.46.20.620
Tetractines of the distal cones
UFBAPOR1917 (H)P47.372.314.297.45.40.520
UFBAPOR1922 (Pa)72.688.48.9103.66.50.820
+ + +.....continued on the next page + + +TABLE 1 +. (Continued) + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
Spicules/ specimensActinesLength (µm)Width (µm)N
MinMeanSDMaxMeanSD
UFBAPOR1917 (H)U60.184.717.1126.95.80.820
UFBAPOR1922 (Pa)74.6111.527.2188.96.40.820
UFBAPOR1917 (H)A11.018.64.527.93.70.817
UFBAPOR1922 (Pa)11.515.93.623.55.11.018
Triactines of the tubes
UFBAPOR1917 (H)P55.376.510.695.15.10.820
UFBAPOR1922 (Pa)70.291.49.0110.07.10.720
UFBAPOR1917 (H)U73.6125.321.6154.05.70.620
UFBAPOR1922 (Pa)92.7131.821.6170.76.70.720
Tetractines of the tubes
UFBAPOR1917 (H)P58.684.613.6112.85.40.720
UFBAPOR1922 (Pa)71.9109.521.0150.55.41.220
UFBAPOR1917 (H)U77.7121.123.9164.75.90.920
UFBAPOR1922 (Pa)106.0142.729.4208.66.10.820
UFBAPOR1917 (H)A12.230.912.849.14.80.820
UFBAPOR1922 (Pa)14.328.710.046.45.61.020
Subatrial triactines
UFBAPOR1917 (H)P29.343.09.163.14.30.820
UFBAPOR1922 (Pa)29.750.48.266.95.70.920
UFBAPOR1917 (H)U62.1103.130.0162.35.50.820
UFBAPOR1922 (Pa)48.996.726.6144.55.50.820
Subatrial tetractines
UFBAPOR1917 (H)P33.558.616.293.05.11.320
UFBAPOR1922 (Pa)37.543.38.052.45.90.23
UFBAPOR1917 (H)U51.8110.025.4168.15.81.020
UFBAPOR1922 (Pa)63.578.819.4100.65.81.13
UFBAPOR1917 (H)A11.014.44.722.54.91.55
UFBAPOR1922 (Pa)------0
Atrial tetractines
UFBAPOR1917 (H)P53.981.018.5115.14.71.010
UFBAPOR1922 (Pa)90.4123.023.3174.56.50.920
UFBAPOR1917 (H)U87.2113.121.0152.55.80.810
UFBAPOR1922 (Pa)110.0146.821.3194.36.60.820
UFBAPOR1917 (H)A16.637.514.358.24.90.89
UFBAPOR1922 (Pa)12.724.98.650.65.00.920
+ + + +FIGURE 2. +Skeleton of + +Sycetta papillata + + +sp. nov. + +(UFBAPOR 1917, holotype). A—Preserved specimen with an incipient peduncle (arrowhead) and apical osculum (arrow). Body covered by papillae (in detail). B—Tangential section showing diactines (arrowhead) and the membrane around the osculum (arrow). C—Transversal section of the skeleton showing separated distal cones and the syconoid aquiferous system. D—Distal cones with triactines and tetractines and tubar tetractines some of them with folded apical actines (arrowhead and in detail). E—Tubar skeleton with triactines and tetractines (arrowhead). F—Tubar tetractines with the apical displaced from the center of the basal actines (arrowhead). G—Subatrial skeleton with triactines (arrowhead). H—Subatrial and atrial regions. Subatrial tetractine (arrowhead) and atrial tetractines with the apical facing the lumen of the atrium (arrow). dc: distal cone; at: atrium. + + + + +FIGURE 3. +Spicules of + +Sycetta papillata + + +sp. nov. + +(UFBAPOR 1922—paratype—except for G, which belongs to the holotype). A—Diactines. B—Triactine of the distal cones. C—Tetractines of the distal cones. D—Tubar triactines. E—Tubar tetractine. F—Subatrial triactines. G—Subatrial tetractine. H—Atrial tetractine. + + + + +Description: +Colour in alcohol is beige. The +holotype +(UFBAPOR 1917) measures 0.8 x +0.2 cm +(height x width), while the other specimens vary from +0.5 to 1.3 cm +in height x +0.2 to 0.3 cm +in width ( +Figs 1A, B, C +). It has a cylindrical shape with terminal osculum surrounded by an incipient membrane ( +Figs 2A, B +). There is no fringe of trichoxeas, and this membrane is the only ornamentation of the osculum ( +Figs 1 +, +2A, B +). The external surface has numerous radial tubes separated from each other, which has been called conical diverticula ( +Dendy 1893 +) or papillae—nomenclature adopted here ( +Haeckel 1872 +; +Van Soest & De Voogd 2015 +) ( +Figs 1 +, +2A +). In some cases, small diactines can be observed protruding from the surface ( +Figs 1E +, +2B +), and these spicules are more easily visualized in tangential sections of the body surface. Some specimens (UFBAPOR 1917, UFBAPOR 1922, and UFBAPOR 1923) have small extensions at the base, resembling a very incipient stalk or stolon ( +Figs 1F +, +2A +). The aquiferous system is syconoid ( +Fig 2C +). + + +The skeleton is articulated, and in most sections, the radial tubes are separated from the proximal region ( +Fig 2C +). As typical of +Syconidae +, there is no cortex. The diactines are not organized in tufts, they occur individually. In UFBAPOR 1947, these diactines extend from the distal cone to the tubar region and, in the +holotype +, they can be visualized tangentially ( +Fig 2B +). In other specimens, they were observed only in dissociated spicules’ slides. In addition to diactines, the distal cone contains triactines and tetractines, which in some cases have the unpaired actine perforating the surface ( +Fig 2D +). The tubar skeleton is formed by triactines and tetractines ( +Figs 2D, E, F +), and these spicules are larger and have different shapes from those present in the distal cones (see below for spicules’ description). They point the unpaired actine towards the distal region ( +Figs 2D, E, F +). At the wall of the radial tubes, tetractines point their apical actines into the lumen ( +Figs 2D, E +). Some of these apical actines have an abrupt fold at the tip ( +Fig 2D +). Both in the distal cones and the tubar skeleton, tetractines were observed with the apical actine displaced from the center of the basal actines ( +Figs 2E, F +). More internally, in the subatrium, there are triactines and tetractines arranged with the unpaired actine facing the distal region ( +Figs 2G, H +). These spicules can be recognized by their obtuse unpaired angle and by their curved paired actines (see below). Tetractines are frequent in the +holotype +but were rarely observed in the +paratype +. The atrial region is the only one that does not have triactines, being formed exclusively by tetractines. Their apical actines face the interior of the atrium, giving the atrial surface a hispid appearance ( +Figs 2G, H +). + + +Spicules +( +Figure 3 +, +Table 1 +)—The measurements presented here are an average of the measurements of the +holotype +and +paratype +. + +Diactines: Straight or slightly curved, with sharp tips [130—304.1 (±126.2)—525.8/ 4.3—6.3 (±1.5)—9.8 μm]. +Triactines of the distal cones: Sagittal.Actines are slightly conical with sharp tips. Paired actines curve downward in a "U" shape. Unpaired actine is straight and larger than the paired ones [paired: 41.1—75.2 (±16.1)—105.6/ 2.9—5.7 (±1.3)—8.3 μm; unpaired: 47.8—103.3 (±32.8)—168.4/ 3.5—5.9 (±0.8)—7.7 μm]. +Tetractines of the distal cones: Sagittal. Actines are slightly conical, with sharp tips. Paired actines curve downward. Unpaired actine is straight and slightly larger or the same size as the paired ones. The apical is short, and it may be displaced from the center of the basal actines [paired: 47.3—80.3 (±14.3)—103.6/ 4.4—6.0 (±0.9)—8.2 μm; unpaired: 60.1—98.1 (±26.2)—188.9/ 4.3—6.1 (±0.8)—7.6 μm; apical: 11.0—17.2 (±4.2)—27.9/ 2.4—4.4 (±1.1)—7.0 μm]. +Triactines of the tubes: Sagittal. Actines are slightly conical, with sharp tips. The paired actines are long and straight, sometimes slightly curved downward in direction to the atrial region. In some cases, one of the paired actines may be shorter than the other. The unpaired angle increases as it approaches the atrial region. The unpaired actine is long and straight [paired: 55.3—83.9 (±12.3)—110.0/ 3.2—6.1 (±1.3)—9.1 μm; unpaired: 73.6—128.6 (±21.6)—170.7/ 4.3—6.2 (±0.8)—7.9 μm]. +Tetractines of the tubes: Sagittal. Actines are slightly conical with sharp tips. The paired actines are long and straight, and the unpaired is larger. Apical actines are long and curved, and in some spicules, an abrupt fold at the tip of the apical can be observed [paired: 58.6—97.1 (±21.5)—150.5/ 3.6—5.4 (±1.0)—7.4 μm; unpaired: 77.7—131.9 (±28.6)—208.6/ 3.9—6.0 (±0.8)—7.6 μm; apical 12.2—29.8 (±11.4)—49.1/3.4—5.2 (±1.0)—7.3 μm]. +Subatrial triactines: Sagittal. Actines are cylindrical with sharp tips. The paired ones are curved toward the distal region, but in some cases, they are straight and short. The unpaired angle is wide. The unpaired actine is long and straight, directed toward the distal cone [paired: 29.3—46.7 (±9.4)—66.9/ 2.8—5.0 (±1.1)—7.1 μm; paired: 48.9—99.9 (±28.2)—162.3/ 3.7—5.5 (±0.8)—7.6 μm]. +Subatrial tetractines: Sagittal. Actines are cylindrical with sharp tips. The paired actines are slightly curved. The unpaired actine is straight and long. The apical actines are short and straight [paired: 33.5—57.1 (±16.8)— 93.0/2.9—5.1 (±1.3)—7.5 μm; paired: 51.8—104.5 (±27.4)—168.1/ 3.4—5.8 (±1.0)—7.4 μm; apical: 11.0—14.4 (±4.7)—22.5/4.0—4.9 (±1.5)—7.5 μm]. +Atrial tetractines: Sagittal. Actines are slightly conical with sharp tips. The paired actines are long and slightly curved. Unpaired actine is long and straight, while apical actine is long and curved [paired: 53.9—112.0 (±28.5)— 174.5/ 3.1—6.0 (±1.3)—7.9 μm; unpaired: 87.2—136.9 (±27.1)—194.3/ 4.6—6.3 (±0.9)—8.1 μm; apical: 12.7— 29.3 (±12.2)—58.2/ 3.6—5.0 (± 0.9)—6.6 μm]. + + + +Remarks: +To date, + +Sycetta + +consists of nine species. Comparing + +Sycetta papillata + + +sp. nov. + +with these species reveals that the composition of its skeleton is unique for the genus: unlike any other + +Sycetta +species + +, + +S. papillata + + +sp. nov. + +has diactines, triactines and tetractines in the distal cones, triactines and tetractines in the tubar and subatrial regions, and an atrial skeleton composed only of tetractines ( +Table 2 +). This composition also differs from + +Sycon tuba +Lendenfeld 1891 + +and + +Sycon conulosum +Cóndor-Luján, Louzada, Hajdu & Klautau, 2018 + +, which have apparent separate distal cones, as tetractines are absent from their tubar and subatrial skeletons. Moreover, in these species the atrium is surrounded by both triactines and tetractines. + + +The main differences between the new species and others species of + +Sycetta + +lie in the presence of a single category of diactines and the composition of the atrial skeleton ( +Table 2 +). + +Sycetta antarctica +Brøndsted, 1931 + +may be considered the most similar to the new species, but it has two diactines’ categories and the atrial skeleton is made from triactines and tetractines ( +Table 2 +). + + + +TABLE 2. +Surface, skeleton composition, type locality and depth of + +Sycetta +species. + +Di—diactines, Tri—triactines, Tetra—tetractines. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
SpeciesSurfaceDistal conesTubesSubatriumAtriumType locality/ depth
+ +S. antarctica +Brøndsted, 1931 + +With papillae-Tri, di (two types)-Tri, tetraAntarctica (350-385 m)
+ +S. asconoides +Breitfuss, 1896 + +SmoothTri (?)TriTriTetraSpitsbergen Islands, Artic Ocean (100-256 m)
+ +S. conifera +( +Haeckel, 1870 +) + +With papillae-TriTriTetraAdriatic Sea (60-100 m)
+ +S. perforata +( +Haeckel, 1872 +) + +SmoothTriTri, tetra-Tri, tetra (a few)Sand-Key, Florida, United States of America (230 m)
+ +S. primitiva +Haeckel, 1872 + +* +With papillaeTriTriTriTriBass Strait (South Australia)
+ +S. quadriradiata +Hôzawa, 1929 + +Even granular-TriTetraTetraKagoshina Bay, Yamakawa, Japan (100 m)
+ +S. sagitta +de Laubenfels, 1942 + +Nearly smooth-Tri-TetraFoxe Channel (Canada)
+ +S. sagittifera +Haeckel, 1872 + +With papillae-Tri-TriSri Lanka
+ +S. vinitincta +Van Soest & De Voogd, 2015 + +With papillaeTri, tetra-Tri, tetraTri, tetra (two types)Samama Island, Indonesia (2 m)
+ +S. papillata + + +sp. nov. + +With papillaeDi, tri, tetraTri, tetraTri, tetraTetraCamaçari, Bahia, Brazil (0.5-25 m)
+ + + +Compiled based on original descriptions and +Burton (1963) +. + + +* + +Sycetta primitiva + +presents all triactines of the skeleton regular and of the same size. + +(-) Not mentioned along the descriptions + +? Mentioned by +Burton (1963) +as doubtful + + + +Among the species previously described within the genus, only two are found in the Atlantic Ocean— + +Sycetta perforata +( +Haeckel, 1872 +) + +in the Tropical Atlantic and + +S. conifera +( +Haeckel, 1870 +) + +in the Temperate Northern Atlantic. Two species are reported for the Arctic— + +S. asconoides +Breitfuss, 1896 + +, and + +S. sagitta +de Laubenfels, 1942 + +, and one for the Southern Ocean— + +S. antarctica + +. The remaining species are distributed across various realms as follows ( + +Spalding +et al. +2007 + +): Central Indo-Pacific ( + +S. vinitincta +Van Soest & De Voogd 2015 + +), Temperate Northern Pacific ( + +S. quadriradiata +Hôzawa, 1929 + +), Western Indo-Pacific ( + +S. sagittifera +Haeckel, 1872 + +), and Temperate Australasia ( + +S. primitiva +Haeckel, 1872 + +). More details on the distribution of these species are provided in +Table 2 +. + + +Another noteworthy factor is the depth in which these species are found. While complete sampling data are lacking for some of them, there are records of + +S. conifera + +occurring between 60 and 100 meters ( +Haeckel 1870 +; +Burton 1963 +), and + +S. sagitta + +at a depth of 100 meters ( +Burton 1963 +). + +Sycetta asconoides + +has been observed at greater depths ranging from 100 to 256 meters ( +de Laubenfels 1942 +; +Burton 1963 +), whereas + +S. antarctica + +is, so far, the species recorded at the greatest depths within the genus, found between 350 and 385 meters ( +Brøndsted 1931 +; +Burton 1963 +). Apart from + +Sycetta papillata + + +sp. nov. + +, which inhabits depths ranging from 0.5 to 25 meters, only + +S. vinitincta + +has been documented in shallow waters, specifically at a depth of 2 meters ( +Van Soest & De Voogd 2015 +). + + + + \ No newline at end of file