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<taxonomicName authority="Fischer, 1809" authorityName="Fischer" authorityYear="1809" class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus karasini" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="karasini">Kytorhinus karasini Fischer, 1809</taxonomicName>
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Figs 31-32
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.
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1♀, Tianshan Mountain, Fukang, Changji, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="43.95">43.95°N</geoCoordinate>
<geoCoordinate id="D780F3B73FFB9AA9B452BE48D8F47251" direction="north" orientation="latitude" precision="555" value="43.95">43.95°N</geoCoordinate>
,
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, alt. ca 2150 m, 1957.VII.14, G. Wang leg., IOZ(E)115178; 1♀, Tianshan Mountain, Fukang, Changji, Xinjiang,
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,
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, alt. ca 2250 m, 1957.VII.14, G. Wang leg., IOZ(E)107522; 1♀, Zhao Su Xian, Yili, Xinjiang,
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<geoCoordinate id="A932BAEC3294F00DCD8FA2DA67CCE57C" direction="north" orientation="latitude" precision="555" value="42.66">42.66°N</geoCoordinate>
,
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<geoCoordinate id="8E9BCD22493388465C8E3685CEE05C01" direction="east" orientation="longitude" precision="555" value="80.18">80.18°E</geoCoordinate>
, alt. ca 2200 m, 1978.VIII.7, Y.H. Han leg., IOZ(E)1045574; 1♀, Takeshikenzhen, Qinggil, Altay, Xinjiang,
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<geoCoordinate id="16F33BF9E57E8D106B71CE7823405560" direction="north" orientation="latitude" precision="555" value="46.18">46.18°N</geoCoordinate>
,
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, alt. ca 1110 m, 2013.VII.28, Y. Li leg.
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<paragraph pageId="6" pageNumber="19">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="19">China, Russia.</paragraph>
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<paragraph id="EC47F5E67345636BD7E545B1D32947CE" pageId="6" pageNumber="19">Distribution.</paragraph>
<paragraph id="AAB7767B67259BF41B717AA7F3309C00" pageId="6" pageNumber="19">China, Russia.</paragraph>
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Figs 33-34, 39, 40
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<paragraph pageId="6" pageNumber="19">Material.</paragraph>
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1♀, Xinjiang, IOZ(E)115177; 1♀, 6 km northwest of Fuyun, Altay, Xinjiang,
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, alt. ca 650 m, 2009.VII.13, X.L. Huang leg.
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<paragraph pageId="6" pageNumber="19">China, Kazakhstan, Mongolia, Russia.</paragraph>
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@ -1,105 +1,104 @@
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Figs 37-38, 47-48
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2♀1♂, Haiziwan Reservo, Shawan, Qoqek, Xinjiang,
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,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="85.78">85.78°E</geoCoordinate>
<geoCoordinate id="87D78036A3ABA8C6A46F7D00C4637E44" direction="east" orientation="longitude" precision="555" value="85.78">85.78°E</geoCoordinate>
, alt. ca 390 m, 1957.VI.9, C.P. Hong leg., IOZ(E)107501-107503; 1♀, Takeshikenzhen, Qinggil, Altay, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="46.18">46.18°N</geoCoordinate>
<geoCoordinate id="3B708218B234021317EB49A73A805AE3" direction="north" orientation="latitude" precision="555" value="46.18">46.18°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="90.81">90.81°E</geoCoordinate>
<geoCoordinate id="45AA5D0BA40BC36A7B81E1492F3C6F01" direction="east" orientation="longitude" precision="555" value="90.81">90.81°E</geoCoordinate>
, alt.1110 m, 2013.VII.28, Y. Li leg.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="distribution">
<paragraph pageId="7" pageNumber="20">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="20">China, Kazakhstan, Mongolia, Russia.</paragraph>
<subSubSection id="52B7D082CD56E6281BE99B4F25BBD655" pageId="7" pageNumber="20" type="distribution">
<paragraph id="54B1E715576A27868C3BAC16DE1F3DD8" pageId="7" pageNumber="20">Distribution.</paragraph>
<paragraph id="3BCDD42605566D90FE651A86DCCB33B2" pageId="7" pageNumber="20">China, Kazakhstan, Mongolia, Russia.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,192 +1,191 @@
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Subfamily
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Figure 2
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<paragraph pageId="2" pageNumber="53">
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Simon, 1890: 95. Type genus
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Simon, 1890, by monotypy.
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
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: 212.
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<paragraph pageId="2" pageNumber="53">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="53">
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<subSubSection id="BB841F3C6C424A34BF619CD3FB8D3301" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53" type="remarks">
<paragraph id="60750DD46417F1842216E37820A69351" pageId="2" pageNumber="53">Remarks.</paragraph>
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are small to tiny ground-dwelling spiders that are largely restricted to arid environments (
<bibRefCitation author="Huber, BA" journalOrPublisher="Insect Systematics and Evolution" pageId="41" pageNumber="92" pagination="15 - 20" title="Ibotyporanga Mello-Leitao: tropical spiders in Brazilian semi-arid habitats (Araneae: Pholcidae)." url="https://doi.org/10.1163/187631203788964926" volume="34" year="2003">Huber and Brescovit 2003</bibRefCitation>
<bibRefCitation id="8224CB190C0297C84FA2604B53905E53" author="Huber, BA" journalOrPublisher="Insect Systematics and Evolution" pageId="41" pageNumber="92" pagination="15 - 20" title="Ibotyporanga Mello-Leitao: tropical spiders in Brazilian semi-arid habitats (Araneae: Pholcidae)." url="https://doi.org/10.1163/187631203788964926" volume="34" year="2003">Huber and Brescovit 2003</bibRefCitation>
; BA Huber, unpublished data). With only 31 described extant species, the subfamily is by far the smallest of the five currently recognized subfamilies in
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.
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seem to be diverse in the New World (ten named genera + about four unnamed genera; BA Huber, unpublished data) where they represent the most southern (Argentina) and most northern (Canada) autochthonous pholcid records worldwide. Only two genera (
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Simon, 1890 and one unnamed; BA Huber, unpublished data) are known from the Old World.
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Their short legs make them superficially strikingly different from
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long-legged pholcids. This distinctness was recognized as early as 1893, when
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Simon classified the only ninetine species available to him in a separate subfamily &quot;
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&quot;, as opposed to all other pholcids classified in
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(
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). Subsequent morphological and molecular phylogenies have partly supported this view (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="AFEDB5B2967EB326D902A066C990BFB4" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
,
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="55D2A632EF01CB5F23519E5D40791BA1" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
) but never convincingly with strong support.
</paragraph>
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<paragraph id="E6FA9EF5F283C137A584C86F6EF6B0AD" pageId="2" pageNumber="53">
Our present analyses include 15 species representing eight of the eleven described genera, originating from both the New World and the Old World (Figure 2). A sister-group relationship between
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and all other pholcids is not supported by our analyses. Instead, all four analyses put
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<taxonomicName id="F0031CB13E101667C9D8C9131CF49499" lsidName="" pageId="2" pageNumber="53" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
as sister to
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<taxonomicName id="68AB4B0905055C8F318A57D7DA74B631" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
Walckenaer, 1837, and this clade is in turn sister to all other pholcids. For reasons discussed below (under
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), we consider this relationship between
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
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and
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dubious. The conclusion here is that
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are
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, either with
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<taxonomicName id="5DFA91D80D16191BDD6264A1540A1B8E" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
or without, but in any case the external relationships of
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<taxonomicName id="DFE9851741FC92DB3AA8EB6989169026" lsidName="" pageId="2" pageNumber="53" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
remain unsatisfactorily resolved and need further study.
</paragraph>
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<paragraph pageId="2" pageNumber="53">
<caption id="B7925C7518A0CA022F59CCC0AD024DEC" pageId="2" pageNumber="53">
<paragraph id="A6A0BAC56BBF7BD87D85D857F2643FD1" pageId="2" pageNumber="53">
Figure 2.
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<taxonomicName id="50E36ECCF0A93087EAD90238F595C828" lsidName="" pageId="2" pageNumber="53" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
and
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<taxonomicName id="BD8D2307152A8C35B518938348F8066C" lsidName="" pageId="2" pageNumber="53" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
a
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<taxonomicName id="A10D24411F50126291DE73D38B9DBD8B" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
sp. n.
<normalizedToken originalValue="“Om14”">&quot;Om14&quot;</normalizedToken>
<normalizedToken id="6587EE4526719D94F00670E1E1299D68" originalValue="“Om14”">&quot;Om14&quot;</normalizedToken>
(Oman) b Gen. n. (Ninetinae) sp. n.
<normalizedToken originalValue="“Om6”">&quot;Om6&quot;</normalizedToken>
<normalizedToken id="676D9103AD0988F1B5B3407EE9603C52" originalValue="“Om6”">&quot;Om6&quot;</normalizedToken>
(Oman) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa diluta" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="diluta">Chisosa diluta</taxonomicName>
<taxonomicName id="E9B6AAE1F608E8A9511314312B296D96" class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa diluta" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="diluta">Chisosa diluta</taxonomicName>
(USA) d Gen. n. (Arteminae) sp. n.
<normalizedToken originalValue="“Ind82”">&quot;Ind82&quot;</normalizedToken>
<normalizedToken id="DFE5C32572548B9996A26AB43A2293F6" originalValue="“Ind82”">&quot;Ind82&quot;</normalizedToken>
(Sulawesi).
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="54">
<pageBreakToken pageId="3" pageNumber="54" start="start">The</pageBreakToken>
<paragraph id="0865F1E3A21D23F3E683DCF826CD23F0" pageId="3" pageNumber="54">
<pageBreakToken id="CB3335E9729F94B60FC42F8DD20D87C4" pageId="3" pageNumber="54" start="start">The</pageBreakToken>
monophyly of the subfamily receives high to full support in all analyses but the composition is slightly different from previous concepts: the North American
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
<taxonomicName id="02864F3DE3D68B8D37F11A06211EAADA" class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
Huber, 2000, originally thought to be a representative of
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="DCC4169187B3899A43E181A5D1C24C02" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
(
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="FFA02EBE8EEED10173F26A5109663A11" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
), is moved to
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="8B11A96B98FDD6C9FD87D653A56A63AF" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
. This move is also supported by male genitalic characters (massive palpal femur; procursus with dorsal apophysis and ventral pocket) and by somatic characters (exposed tarsal organ; reduction of epiandrous spigots;
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="FD2A7F709077FA7410D7F837DF748D06" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
). Another genus that was previously (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="45 - 53" title="On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar." volume="1635" year="2007">Huber and El Hennawy 2007</bibRefCitation>
<bibRefCitation id="9F5E38521BF6E8EC6E298A6175D9FD6C" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="45 - 53" title="On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar." volume="1635" year="2007">Huber and El Hennawy 2007</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
<bibRefCitation id="695C1020D3E5D43ACBBF3483165D5A70" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
) thought to be a member of
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="F10CDC781549A9608E28A5E604B03E5C" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
is
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="1702B929C85784810E8B1F725470CF6F" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
Huber &amp; El Hennawy, 2007. As already suggested in a previous analysis (
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="23C520E0F5678BB58D49C2C77FEE6D7D" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
),
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="47A6E5A3C2EA339CA4386C8BBD0CA879" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
is not a member of
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="3C72B6D6297BBA38147C8197A2F9FA91" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
but of
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="BAAEA862874B1B28B2252BC39D0A7B63" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
.
</paragraph>
<paragraph pageId="3" pageNumber="54">
<paragraph id="9F166E9FA286F4BA300FF90BDCB387F5" pageId="3" pageNumber="54">
The internal relationships of
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="E55882BBD151153080F2F13D7187F7B2" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
suggested by the molecular data are difficult to evaluate: they are mostly neither supported nor contradicted by morphological data. Two details are remarkable because they suggest that South America may not only be the most diverse region as far as
<taxonomicName lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="038FD45170E78AE071D545EF7B7EE54C" lsidName="" pageId="3" pageNumber="54" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
are concerned but also the ancestral region of the subfamily. First, the analyses fully support a monophyletic North and Central American/Caribbean clade (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcophora" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Pholcophora</taxonomicName>
<taxonomicName id="C29EE924CF0EEF4A6B3E6757BBEA8ACF" class="Arachnida" family="Pholcidae" genus="Pholcophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcophora" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Pholcophora</taxonomicName>
Banks, 1896;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Papiamenta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papiamenta" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Papiamenta</taxonomicName>
<taxonomicName id="EFB6CE6AF7354609F7FB0DF3540EF425" class="Arachnida" family="Pholcidae" genus="Papiamenta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papiamenta" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Papiamenta</taxonomicName>
Huber, 2000; and unidentified taxa from Cuba and Puerto Rico; &quot;clade 2e&quot; in
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
<bibRefCitation id="E16D5D467B0DA51550643CA217BFA66E" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
) that is either nested among South American ancestors or is sister to the South American
<taxonomicName class="Arachnida" family="Pholcidae" genus="Ibotyporanga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ibotyporanga" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ibotyporanga</taxonomicName>
<normalizedToken originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
<taxonomicName id="5CB905873483D07A0BC08245D2CA1324" class="Arachnida" family="Pholcidae" genus="Ibotyporanga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ibotyporanga" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ibotyporanga</taxonomicName>
<normalizedToken id="57049E453850B8E59DBBACEC45185F23" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1944 (with reasonable support in the 4+ genes tree only). Based on its geographic distribution, we predict that the Mexican
<taxonomicName class="Arachnida" family="Pholcidae" genus="Tolteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tolteca" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Tolteca</taxonomicName>
<taxonomicName id="A9EF69EA4716A7FEF723D908D069EEF5" class="Arachnida" family="Pholcidae" genus="Tolteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tolteca" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Tolteca</taxonomicName>
is also a member of this clade. Second, the two Old World genera (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Ninetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ninetis" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ninetis</taxonomicName>
<taxonomicName id="696AC4E98EDFCB6DBDD327CCEAF2C3D4" class="Arachnida" family="Pholcidae" genus="Ninetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ninetis" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ninetis</taxonomicName>
and an undescribed genus from Oman) are also sister taxa (with low to modest support) and in all analyses (except for the 4+ genes analysis where
<taxonomicName class="Arachnida" family="Pholcidae" genus="Ninetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ninetis" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ninetis</taxonomicName>
<taxonomicName id="D411235F2508F4FEB0296880296D6167" class="Arachnida" family="Pholcidae" genus="Ninetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ninetis" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Ninetis</taxonomicName>
is missing) nested among South American taxa.
</paragraph>
</subSubSection>

View file

@ -1,275 +1,274 @@
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<treatment ID-GBIF-Taxon="148671575" LSID="urn:lsid:plazi:treatment:2C6A08DB4D12751A58C844D35E7C10F5" httpUri="http://treatment.plazi.org/id/2C6A08DB4D12751A58C844D35E7C10F5" lastPageId="12" lastPageNumber="63" pageId="9" pageNumber="60">
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<paragraph pageId="9" pageNumber="60">
<treatment id="2C6A08DB4D12751A58C844D35E7C10F5" ID-GBIF-Taxon="148671575" LSID="urn:lsid:plazi:treatment:2C6A08DB4D12751A58C844D35E7C10F5" httpUri="http://treatment.plazi.org/id/2C6A08DB4D12751A58C844D35E7C10F5" lastPageId="12" lastPageNumber="63" pageId="9" pageNumber="60" scope_family="Pholcidae" scope_order="Araneae">
<subSubSection id="9D55B24EE4831384C0A1B2D6A5B2CBB5" pageId="9" pageNumber="60" type="nomenclature">
<paragraph id="E3CB491F65D6CA2B6C944B03CF77636D" pageId="9" pageNumber="60">
Subfamily
<taxonomicName authority="Simon, 1893" authorityName="Simon" authorityYear="1893" family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="9" pageNumber="60" rank="subFamily" subFamily="Smeringopinae">Smeringopinae Simon, 1893</taxonomicName>
<taxonomicName id="C66548ABA39B23A654DDF5FC615DD4D1" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Smeringopinae">Smeringopinae Simon, 1893</taxonomicName>
Figure 6
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="61" type="reference_group">
<paragraph pageId="10" pageNumber="61">
<taxonomicName family="Smeringopodeae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Modisimeae" order="Araneae" pageId="10" pageNumber="61" rank="family">
<pageBreakToken pageId="10" pageNumber="61" start="start">Smeringopodeae</pageBreakToken>
<subSubSection id="0775F9891EDD695B5E78FA7DBDB60983" pageId="10" pageNumber="61" type="reference_group">
<paragraph id="16F37E53AA31D704A474D24B5F215ABC" pageId="10" pageNumber="61">
<taxonomicName id="EF82E63D93E9BC2FE05C82A64448E4BD" class="Arachnida" family="Smeringopodeae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimeae" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="family">
<pageBreakToken id="D83DFDEE41EB8FC5ECA83BF74046CD51" pageId="10" pageNumber="61" start="start">Smeringopodeae</pageBreakToken>
</taxonomicName>
Simon, 1893: 474. Type genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="974D2E71F6A576A26EC35C88CD76F9F9" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
Simon, 1890, by subsequent designation (
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
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).
</paragraph>
<paragraph pageId="10" pageNumber="61">
<taxonomicName family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<paragraph id="1F0FE0ACFDF14D6E2F4B8A78D1D4306A" pageId="10" pageNumber="61">
<taxonomicName id="7B372341292BFCC2D42548433281D91F" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
Simon;
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
<bibRefCitation id="69823C4780431BF5CC84E4620BA89A00" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
: 217.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="63" pageId="10" pageNumber="61" type="remarks">
<paragraph pageId="10" pageNumber="61">Remarks.</paragraph>
<paragraph pageId="10" pageNumber="61">
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<subSubSection id="44D4C455DA911464A732D622F0E7D9CD" lastPageId="12" lastPageNumber="63" pageId="10" pageNumber="61" type="remarks">
<paragraph id="7ACCECDF47E6B1948B0698A32E53B869" pageId="10" pageNumber="61">Remarks.</paragraph>
<paragraph id="BDE91078EDF9A5667D25885B9AEEE538" pageId="10" pageNumber="61">
<taxonomicName id="F70842AD1CB436611BE37DB1D44116C3" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
is a relatively homogeneous subfamily (with respect to body shapes, colour, webs, and microhabitats), and in this sense similar to
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="EDB725AD1A38C25F140B69FC4F685294" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
and
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="E372E9CE970DDE14A5F05C966165C2F7" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
but very unlike
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="0C72B58B3E0B78BCE34E5D0A8FE53C17" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
and
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="941C3D840470838BABF85039F60A17AA" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
. Most of the 125 known species of
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="7187551B7D09B1DDC9A0423112DD0267" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
are medium-size to large, have long legs, elongated to cylindrical abdomens, and all have eight eyes. Another similarity to
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="490DD40B0860DE78B8A1A50A14AF0E87" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
and
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="DACF510261E195C59DCA04BF218C8B37" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
is that
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="7E40313E0A064505E928B0061C9F2118" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
are often found in rather arid regions. The most obvious exception is the largely humid tropical genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
<taxonomicName id="2AB18D230D838E8F25CDD62D689CB3B6" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
Kraus, 1957, which is also the genus with the smallest and largest representatives in the subfamily (with body lengths ranging from 2.5-10 mm) and with the widest range of microhabitats used (leaf litter to large sheltered spaces) (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
<bibRefCitation id="8CDA69DAA96D430D76A099F0272F8803" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
). The original distribution of the subfamily is Africa, the Mediterranean, and the Middle East. Three species have attained much wider distributions, resulting from human-mediated dispersal (
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
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).
</paragraph>
<paragraph pageId="10" pageNumber="61">
<paragraph id="A964C7E64C657FAC4021995A75602FD1" pageId="10" pageNumber="61">
As in previous molecular analyses (
<bibRefCitation author="Bruvo-Mađaric, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="39" pageNumber="90" pagination="661 - 673" title="Phylogeny of pholcid spiders (Araneae: Pholcidae): combined analysis using morphology and molecules." url="https://doi.org/10.1016/j.ympev.2005.08.016" volume="37" year="2005">
<normalizedToken originalValue="Bruvo-Mađarić">Bruvo-Mađaric</normalizedToken>
<bibRefCitation id="54ADD98449B96B2F5BFB8D69D563ED93" author="Bruvo-Mađaric, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="39" pageNumber="90" pagination="661 - 673" title="Phylogeny of pholcid spiders (Araneae: Pholcidae): combined analysis using morphology and molecules." url="https://doi.org/10.1016/j.ympev.2005.08.016" volume="37" year="2005">
<normalizedToken id="7F347596D21C071E35977F97A8ED1600" originalValue="Bruvo-Mađarić">Bruvo-Mađaric</normalizedToken>
et al. 2005
</bibRefCitation>
,
<bibRefCitation author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
<bibRefCitation id="8C7306BF74C371637F366A962501BA4C" author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
,
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="BD7544C6B196B396F0FE9F3A9CC030DD" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
),
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="86301D70EBBE3D214CF48DD17316A973" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
is sister to
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="7451282CDD9A8D0F0F6B540E3B412C93" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
(Figure 1) with reasonable to high support. This relationship is also supported by morphology: the two taxa share tarsus IV comb-hairs spread over the entire length of the tarsus (
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="232 - 240" title="Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae)." url="https://doi.org/10.1636/CSh07-71.1" volume="36" year="2008">Huber and Fleckenstein 2008</bibRefCitation>
<bibRefCitation id="98A9CAEB4ECB42C7D76C3B04282A1FC2" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="232 - 240" title="Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae)." url="https://doi.org/10.1636/CSh07-71.1" volume="36" year="2008">Huber and Fleckenstein 2008</bibRefCitation>
).
</paragraph>
<paragraph pageId="10" pageNumber="61">
<paragraph id="DC03B30A36C7767C8AC0DED63F22F3C4" pageId="10" pageNumber="61">
The monophyly of
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="7CAAE9036E26CE8E6B593DF760E85057" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
receives reasonable to high support in all our analyses. Previous molecular analyses have partly supported
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="61F431C14409AB8E7EE89B9133708664" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
, but also suggested rather obscure relationships [e.g., the position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus pluchei" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="pluchei">Holocnemus pluchei</taxonomicName>
<taxonomicName id="BFE102063E81811197F190BC3BF8D332" class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus pluchei" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="pluchei">Holocnemus pluchei</taxonomicName>
(Scopoli, 1763) among
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="4C580DC4ECCC0C96818D4B9E4A54A1C5" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
in
<bibRefCitation author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
<bibRefCitation id="2B216E65EF2FE38A1A10A720004D0E6D" author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
].
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus pluchei" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="pluchei">Holocnemus pluchei</taxonomicName>
<taxonomicName id="34E339EBF6E3B9273EEF647731813524" class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus pluchei" order="Araneae" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="pluchei">Holocnemus pluchei</taxonomicName>
was included in preliminary analyses of the present data but its position was drastically unstable, so we decided to exclude it from the final analyses.
<taxonomicName lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="94BFCF4183A1C1C480AB46DE10D96CF2" lsidName="" pageId="10" pageNumber="61" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
monophyly is rather weakly supported by morphology, i.e., by the presence of a large thoracic pit on the carapace (rather than a narrow furrow or an evenly domed carapace; cf.
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
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).
</paragraph>
<paragraph pageId="11" pageNumber="62">
<pageBreakToken pageId="11" pageNumber="62" start="start">Within</pageBreakToken>
<taxonomicName lsidName="" pageId="11" pageNumber="62" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<paragraph id="6DB13A210DE6AF8DC1FD4A52F1A8145B" pageId="11" pageNumber="62">
<pageBreakToken id="D9ADE29C3865CE92FB52A429E77AF559" pageId="11" pageNumber="62" start="start">Within</pageBreakToken>
<taxonomicName id="36D551D341021E937BC7E82023D5D021" lsidName="" pageId="11" pageNumber="62" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
, our data strongly support a basal split between a northern clade (Mediterranean, northern Africa, Middle East, Central Asia) and a southern clade (Sub-Sahara) (Figure 6). This basal split was also recovered in a morphological cladistic analysis (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber 2012</bibRefCitation>
<bibRefCitation id="5BC192C5EBEA3A0237382E170B1D33AF" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber 2012</bibRefCitation>
). Within the northern clade,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
<taxonomicName id="160962F756293188DF39C3C5C7F49BDC" class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
Kulczynski, 1908 is sister to all other genera and not close to
<taxonomicName class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Stygopholcus</taxonomicName>
<taxonomicName id="9D2220A952B8D3590AB4B0A8EA2A9347" class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Stygopholcus</taxonomicName>
Kratochvil, 1932 as repeatedly claimed by
<bibRefCitation author="Brignoli, PM" journalOrPublisher="Mitteilungen des Zoologischen Museums Berlin" pageId="39" pageNumber="90" pagination="255 - 267" title="Beitrag zur Kenntnis der mediterranen Pholcidae (Arachnida, Araneae)." url="https://doi.org/10.1002/mmnz.19710470203" volume="47" year="1971">Brignoli (1971</bibRefCitation>
<bibRefCitation id="28B363C60D128104A4FF384DA0A85EE1" author="Brignoli, PM" journalOrPublisher="Mitteilungen des Zoologischen Museums Berlin" pageId="39" pageNumber="90" pagination="255 - 267" title="Beitrag zur Kenntnis der mediterranen Pholcidae (Arachnida, Araneae)." url="https://doi.org/10.1002/mmnz.19710470203" volume="47" year="1971">Brignoli (1971</bibRefCitation>
,
<bibRefCitation author="Brignoli, PM" journalOrPublisher="Revue suisse de Zoologie" pageId="39" pageNumber="90" pagination="539 - 578" title="Ragni di Grecia IX. Specie nuove o interessanti delle famiglie Leptonetidae, Dysderidae, Pholcidae ed Agelenidae (Araneae)." volume="83" year="1976">1976</bibRefCitation>
<bibRefCitation id="8896A977635055C9B79F28BB97AE899E" author="Brignoli, PM" journalOrPublisher="Revue suisse de Zoologie" pageId="39" pageNumber="90" pagination="539 - 578" title="Ragni di Grecia IX. Specie nuove o interessanti delle famiglie Leptonetidae, Dysderidae, Pholcidae ed Agelenidae (Araneae)." volume="83" year="1976">1976</bibRefCitation>
,
<bibRefCitation author="Brignoli, PM" journalOrPublisher="Bulletin of the British arachnological Society" pageId="39" pageNumber="90" pagination="350 - 352" title="Spiders from Lebanon, V. On Hoplopholcuscecconii Kulczynski, 1908 (Pholcidae)." volume="4" year="1979">1979</bibRefCitation>
<bibRefCitation id="EC995F8CBDC307937A5BC2A8E14EAE8D" author="Brignoli, PM" journalOrPublisher="Bulletin of the British arachnological Society" pageId="39" pageNumber="90" pagination="350 - 352" title="Spiders from Lebanon, V. On Hoplopholcuscecconii Kulczynski, 1908 (Pholcidae)." volume="4" year="1979">1979</bibRefCitation>
) but contested by
<bibRefCitation pageId="11" pageNumber="62">Senglet (1971</bibRefCitation>
<bibRefCitation id="7BB1261862A500B95B1DE85C033104A8" pageId="11" pageNumber="62">Senglet (1971</bibRefCitation>
,
<bibRefCitation author="Senglet, A" journalOrPublisher="Mitteilungen der schweizerischen entomologischen Gesellschaft" pageId="43" pageNumber="94" pagination="43 - 67" title="Copulatory mechanisms in Hoplopholcus, Stygopholcus (revalidated), Pholcus, Spermophora and Spermophorides (Araneae, Pholcidae), with additional faunistic and taxonomic data." volume="74" year="2001">2001</bibRefCitation>
<bibRefCitation id="A488C4AD39F62966C1E7BD7637D0563C" author="Senglet, A" journalOrPublisher="Mitteilungen der schweizerischen entomologischen Gesellschaft" pageId="43" pageNumber="94" pagination="43 - 67" title="Copulatory mechanisms in Hoplopholcus, Stygopholcus (revalidated), Pholcus, Spermophora and Spermophorides (Araneae, Pholcidae), with additional faunistic and taxonomic data." volume="74" year="2001">2001</bibRefCitation>
). The genera
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
<taxonomicName id="9B2FD9428E4FEC2311039DE01F5D841C" class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Stygopholcus</taxonomicName>
<taxonomicName id="3C0015FCD155B9D323D801256AA83A3F" class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Stygopholcus</taxonomicName>
, and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Crossopriza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crossopriza" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Crossopriza</taxonomicName>
<taxonomicName id="5BE6ECE44AE88217301A930F98FF93B0" class="Arachnida" family="Pholcidae" genus="Crossopriza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crossopriza" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Crossopriza</taxonomicName>
Simon, 1893 all receive full support, but the small Mediterranean genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
<taxonomicName id="EEF75DCCAA894500D909A8608B36D5E5" class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
Simon, 1873 (only three described species) continues to be problematic even after the exclusion of
<taxonomicName lsidName="H. pluchei" pageId="11" pageNumber="62" rank="species" species="pluchei">H. pluchei</taxonomicName>
<taxonomicName id="BA2B5E51E9388095AA72E997436E82A5" lsidName="H. pluchei" pageId="11" pageNumber="62" rank="species" species="pluchei">H. pluchei</taxonomicName>
. The two species of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
<taxonomicName id="A1E94F8071A3764958252A828E6E2246" class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
included in our analyses never group together, and no morphological synapomorphy is known to suggest their sister-group relationship (in fact,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
<taxonomicName id="3D4B803280A9E59C080C1F9AF637FED1" class="Arachnida" family="Pholcidae" genus="Holocnemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocnemus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Holocnemus</taxonomicName>
has never been revised).
</paragraph>
<caption pageId="11" pageNumber="62">
<paragraph pageId="11" pageNumber="62">
<caption id="8AC6A2CB8C949436D712B6F9EA9CEB8E" pageId="11" pageNumber="62">
<paragraph id="1CA11D671F79380F63F39D8213F82811" pageId="11" pageNumber="62">
Figure 6. Smeringopinae a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
<taxonomicName id="7F4076C1890911210728981ED2D151C0" class="Arachnida" family="Pholcidae" genus="Hoplopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopholcus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Hoplopholcus</taxonomicName>
sp. n.
<normalizedToken originalValue="“Mar66”">&quot;Mar66&quot;</normalizedToken>
<normalizedToken id="846B189F0410A83C0E8F47474CAE55DE" originalValue="“Mar66”">&quot;Mar66&quot;</normalizedToken>
(Turkey) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus absoloni" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="absoloni">Stygopholcus absoloni</taxonomicName>
<taxonomicName id="C2D9ED0603F050FDF90320D339FF7377" class="Arachnida" family="Pholcidae" genus="Stygopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygopholcus absoloni" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="absoloni">Stygopholcus absoloni</taxonomicName>
? (Bosnia and Herzegovina) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Crossopriza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crossopriza" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Crossopriza</taxonomicName>
<taxonomicName id="B73E060FD67EAE6C593C7F333E1F837F" class="Arachnida" family="Pholcidae" genus="Crossopriza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crossopriza" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">Crossopriza</taxonomicName>
sp. n.
<normalizedToken originalValue="“Om11”">&quot;Om11&quot;</normalizedToken>
<normalizedToken id="6F7CA0A04560A142B61DE6C85FB514A9" originalValue="“Om11”">&quot;Om11&quot;</normalizedToken>
(Oman) d
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<taxonomicName id="CDB3BFC6EBB194325ECBD7D9D8B0BD0C" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus pallidus" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="pallidus">Smeringopus pallidus</taxonomicName>
(Philippines) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina pulchra" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="pulchra">Smeringopina pulchra</taxonomicName>
<taxonomicName id="C1C2CD6B7687F2DEE11AAEC1F57F05EC" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina pulchra" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="pulchra">Smeringopina pulchra</taxonomicName>
(Ghana) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina ankasa" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="ankasa">Smeringopina ankasa</taxonomicName>
<taxonomicName id="F37D43AAD6672FF9D447543A18488596" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina ankasa" order="Araneae" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="ankasa">Smeringopina ankasa</taxonomicName>
(Ghana).
</paragraph>
</caption>
<paragraph pageId="12" pageNumber="63">
<pageBreakToken pageId="12" pageNumber="63" start="start">The</pageBreakToken>
<paragraph id="EBC31628DEDC8795EE2D7859350AFCC6" pageId="12" pageNumber="63">
<pageBreakToken id="4766D9B613C35BB7588B4CA9FCD3B253" pageId="12" pageNumber="63" start="start">The</pageBreakToken>
southern (Sub-Saharan) clade includes
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="C7E82FDE349F36B0243E0A5C64AC2C5F" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
Simon, 1890 and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
<taxonomicName id="6090F4BBAC5FD6082A54AEF492C0989A" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
, and is also supported by a unique number of epiandrous spigots (two) (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber 2012</bibRefCitation>
<bibRefCitation id="EA323C12432C6D263FA3F71F654E3F51" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber 2012</bibRefCitation>
). The paraphyly of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="80F943C7B1875391AAE071867B081B6D" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
has been suggested before (
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="C0C46BBE00097303A5CB8F9275A5119E" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
), and our larger data set supports this view, but with low support values. Two of the species groups of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="F27FEA9F68A26B88D4B84F4C1CF86461" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
proposed in
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber (2012)</bibRefCitation>
<bibRefCitation id="017AA2E502E57AB66FC0DD7967E0351A" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber (2012)</bibRefCitation>
appear closer to
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<taxonomicName id="5C784C0AEA3530BBCB15C362897D2462" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
than to other
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="4A36A050996784AFC3EB5E2EFD9A7149" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
: the
<taxonomicName genus="Modisimeae" lsidName="Modisimeae chogoria" pageId="12" pageNumber="63" rank="species" species="chogoria">chogoria</taxonomicName>
<taxonomicName id="86134A9D132261370D2E67B6C5D270F9" genus="Modisimeae" lsidName="Modisimeae chogoria" pageId="12" pageNumber="63" rank="species" species="chogoria">chogoria</taxonomicName>
group and the rubrotinctus group. Morphological data do not support this view but they neither strongly contradict it: the two species groups lack the distinctive arrangement of pores on the pore plates (in groups or
<normalizedToken originalValue="islands">'islands'</normalizedToken>
<normalizedToken id="C0B4DDFD3B6880ADD7ED4E4343F35952" originalValue="islands">'islands'</normalizedToken>
) and the retrolateral furrow on the male palpal femur present in all other species of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="8ABDEE4BE794171B1B491CF595CDABA6" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
(
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber 2012</bibRefCitation>
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). Remarkably,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="913F52556FDA480991EB902E7C063E69" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
<taxonomicName id="051B622A0F4AFB0004B5E7E35F699D33" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
are largely separated geographically, with
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
<taxonomicName id="DD9E257488189A4BAF18EFA467384575" class="Arachnida" family="Pholcidae" genus="Smeringopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopus</taxonomicName>
being most diverse in southern and eastern Africa, and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
<taxonomicName id="6C85C0B2A014A5E2E885F67A08ED1CCD" class="Arachnida" family="Pholcidae" genus="Smeringopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smeringopina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Smeringopina</taxonomicName>
in western and central Africa (
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,
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<bibRefCitation id="0E9F964BD43CD475EF6E69C2FA265A31" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">2013</bibRefCitation>
). The
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and rubrotinctus groups are geographically restricted to an area where Central Africa (the Guineo-Congolian center of endemism) meets East Africa (
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). Other than that, our sampling in
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is not dense enough to test the species groups proposed in
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 138" title="Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)." volume="3461" year="2012">Huber (2012)</bibRefCitation>
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. Remarkably, though, the isolated
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position of
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Huber, 2012 is supported by the present analyses.
</paragraph>
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<paragraph id="70C4F90C99BE7B1B036BD69E88797857" pageId="12" pageNumber="63">
Our analyses include 30 of the 44 described species of
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(68%), and all species groups proposed in
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except two monotypic
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(
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<taxonomicName id="2BDF1E5C4AEE3C091AF185D8EF1D9C97" lsidName="S. fon" pageId="12" pageNumber="63" rank="species" species="fon">S. fon</taxonomicName>
Huber, 2013;
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Huber, 2013). Even though for some species only one gene (CO1) was sequenced, our analyses support several species groups and deeper relationships proposed previously (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
<bibRefCitation id="5DB5F0D0442B792D41A5253301B028EC" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
), based on cladistic analysis of morphological characters. Morphology placed the West African guineensis group as sister to all other
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; all our analyses support both the monophyly of the guineensis group and its sister-group relationship with all other congeners. The next two branches are composed of representatives of the lekoni group, which is thus here considered paraphyletic rather than monophyletic. The
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and cornigera groups are both supported, as is their sister group relationship to each other. The attuleh group is supported, but not as sister to the
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<taxonomicName id="E4147584E4A34649A911577B754659D9" genus="Modisimeae" lsidName="Modisimeae ankasa" pageId="12" pageNumber="63" rank="species" species="ankasa">ankasa</taxonomicName>
+ cornigera groups but as sister to the following group. The last clade is composed of representatives of the simplex and beninensis groups, but the clear dichotomy in the molecular trees is not equivalent to these groups. Instead, the simplex group includes all
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representatives originally assigned to the beninensis group; the beninensis group includes only those species that have a light transversal element ventrally on the abdomen (character 9 in
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
<bibRefCitation id="C541239F0FE65C718CC57C0575C610F3" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 160" title="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae)." url="https://doi.org/10.11646/zootaxa.3713.1.1" volume="3713" year="2013">Huber 2013</bibRefCitation>
, which is thus less homoplastic than previously thought).
</paragraph>
</subSubSection>

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@ -1,125 +1,124 @@
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<paragraph pageId="6" pageNumber="19">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<subSubSection id="6872D8BD84C59726730AA6BAE10D96ED" pageId="6" pageNumber="19" type="nomenclature">
<paragraph id="F7FACCA2D29CD149E7B2DBF5352EB07F" pageId="6" pageNumber="19">
<taxonomicName id="76B658466515F4DC6CD466558181EC9B" authority="Delobel &amp; Legalov, 2009" authorityName="Delobel &amp; Legalov" authorityYear="2009" genus="Kytorhinus" lsidName="Kytorhinus kergoati" pageId="6" pageNumber="19" rank="species" species="kergoati">Kytorhinus kergoati Delobel &amp; Legalov, 2009</taxonomicName>
Figs 35-36, 41-42, 68
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="19" type="material">
<paragraph pageId="6" pageNumber="19">Material.</paragraph>
<paragraph pageId="6" pageNumber="19">
<subSubSection id="23FBB473DC08F100432C64E7E6EC8BB1" pageId="6" pageNumber="19" type="material">
<paragraph id="E6FF656518C353C107EBF970C9348A73" pageId="6" pageNumber="19">Material.</paragraph>
<paragraph id="823B7AA2B6A7B4788DEC28E8490B95B6" pageId="6" pageNumber="19">
1♀, Pochengzi, Wensu, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
<geoCoordinate id="00CF5A41E70E428A16CE243C57C3051F" direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.91">80.91°E</geoCoordinate>
<geoCoordinate id="E2B4BE95BAA770B89B628936BEAF8519" direction="east" orientation="longitude" precision="555" value="80.91">80.91°E</geoCoordinate>
, alt. ca 1930 m, 1978.VI.3, X.Z. Zhang leg., IOZ(E)1045598; 1♀1♂, Tomur peak, Wensu, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.74">41.74°N</geoCoordinate>
<geoCoordinate id="3129EA0DA69BF16D7BE03F9DD4B3516A" direction="north" orientation="latitude" precision="555" value="41.74">41.74°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.58">80.58°E</geoCoordinate>
<geoCoordinate id="F57C293D3FA62B9DA9690F59739DA6EB" direction="east" orientation="longitude" precision="555" value="80.58">80.58°E</geoCoordinate>
, alt. ca 2400 m, 1977.VI.19, C.J. Li leg., IOZ(E)632265-632266; 1♂, Tomur peak, Wensu, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.81">41.81°N</geoCoordinate>
<geoCoordinate id="002B07597583F276B61C4806BA15C85E" direction="north" orientation="latitude" precision="555" value="41.81">41.81°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.59">80.59°E</geoCoordinate>
<geoCoordinate id="9B47647965414923B1AF6D0831D3AA97" direction="east" orientation="longitude" precision="555" value="80.59">80.59°E</geoCoordinate>
, alt. ca 3200 m, 1977.VII.11, Y.H. Han leg., IOZ(E)632264; 1♂, Tomur peak, Wensu, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.81">41.81°N</geoCoordinate>
<geoCoordinate id="DE8BE6B1B495DAA0AA4242EB29444341" direction="north" orientation="latitude" precision="555" value="41.81">41.81°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.59">80.59°E</geoCoordinate>
<geoCoordinate id="7A2C22FC9B1652A6466A3B1905A5472D" direction="east" orientation="longitude" precision="555" value="80.59">80.59°E</geoCoordinate>
, alt. ca 3200 m, 1977.VII.14, C.J. Li leg., IOZ(E)632263; 1♂, Yangbajingzhen, Damxung, Lhasa, Tibet, China,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.02">30.02°N</geoCoordinate>
<geoCoordinate id="0D561B02FE7904A62D637AE69295653D" direction="north" orientation="latitude" precision="555" value="30.02">30.02°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="90.39">90.39°E</geoCoordinate>
<geoCoordinate id="5D61ECA946B38DDB3287FAF094164ADE" direction="east" orientation="longitude" precision="555" value="90.39">90.39°E</geoCoordinate>
, alt. ca 4310 m, 1960.VI.2, C.G. Wang leg., IOZ(E)1016165.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="19" type="distribution">
<paragraph pageId="6" pageNumber="19">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="19">New record for China, Tajikistan (Gorno-Badakhshan autonomous province).</paragraph>
<subSubSection id="9BCFB07E9DCAE263E5F0FA77A4C13B4B" pageId="6" pageNumber="19" type="distribution">
<paragraph id="A865AB6FF06F0B14D50B7F61D70C579E" pageId="6" pageNumber="19">Distribution.</paragraph>
<paragraph id="7F223476EA0B657DF52DAE09DDAF4DD9" pageId="6" pageNumber="19">New record for China, Tajikistan (Gorno-Badakhshan autonomous province).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="19" type="host">
<paragraph pageId="6" pageNumber="19">Host.</paragraph>
<paragraph pageId="6" pageNumber="19">Unknown.</paragraph>
<subSubSection id="6B6234CF03AB98BCC676B20EDB427D49" pageId="6" pageNumber="19" type="host">
<paragraph id="85DAE4E05C8A5E94E3A49A01EFC3791B" pageId="6" pageNumber="19">Host.</paragraph>
<paragraph id="FFB7DF9D74E27A0C52F108125AC7E6E7" pageId="6" pageNumber="19">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="19" type="remarks">
<paragraph pageId="6" pageNumber="19">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="19">
<bibRefCitation author="Delobel, A" journalOrPublisher="Genus" pageId="11" pageNumber="24" pagination="71 - 76" title="A new species of Kytorhinus Fisch. - Wald. from western Pamir (Coleoptera: Bruchinae)." volume="20" year="2009">Delobel and Legalov (2009)</bibRefCitation>
<subSubSection id="B66E0FB988B9D121021EA309C8204ECE" pageId="6" pageNumber="19" type="remarks">
<paragraph id="A54893B46F9C968CA2B6049C4BED6A22" pageId="6" pageNumber="19">Remarks.</paragraph>
<paragraph id="099EF5B3264EB0995B89221AD32DBDB3" pageId="6" pageNumber="19">
<bibRefCitation id="4ECA54336774CA1A044D6C5D0DBC8CA0" author="Delobel, A" journalOrPublisher="Genus" pageId="11" pageNumber="24" pagination="71 - 76" title="A new species of Kytorhinus Fisch. - Wald. from western Pamir (Coleoptera: Bruchinae)." volume="20" year="2009">Delobel and Legalov (2009)</bibRefCitation>
described this species based on a single male. We examined a female specimen and found that its antennae are serrated and are slightly longer than half of the body length (Fig. 35, 41, 42). Additionally, the elytra of the female are almost yellow, with the exception of a black, elongate triangular area, which extends from the base to one-third of the elytral suture (Fig. 35) corresponding to main distinguishing characters of a single known male of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus kergoati" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="kergoati">Kytorhinus kergoati</taxonomicName>
<taxonomicName id="16C7588FF828BA64E088242016C170C6" genus="Kytorhinus" lsidName="Kytorhinus kergoati" pageId="6" pageNumber="19" rank="species" species="kergoati">Kytorhinus kergoati</taxonomicName>
.
</paragraph>
</subSubSection>

View file

@ -1,126 +1,125 @@
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<paragraph pageId="5" pageNumber="18">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
<treatment id="847CDF36DB80C5418CFE8EE4582BE08F" ID-GBIF-Taxon="152057188" LSID="urn:lsid:plazi:treatment:847CDF36DB80C5418CFE8EE4582BE08F" httpUri="http://treatment.plazi.org/id/847CDF36DB80C5418CFE8EE4582BE08F" lastPageNumber="18" pageId="5" pageNumber="18" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Bruchinae" scope_tribe="Kytorhinini">
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<paragraph id="00A958FCFB8E8FEFCB2BA3855D0143A3" pageId="5" pageNumber="18">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<paragraph pageId="5" pageNumber="18">
<taxonomicName authority="Motschulsky, 1874" authorityName="Motschulsky" authorityYear="1874" class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus immixtus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="immixtus">Kytorhinus immixtus Motschulsky, 1874</taxonomicName>
<subSubSection id="A54CFAE8687A936C8DA0FE6005EAC03D" pageId="5" pageNumber="18" type="nomenclature">
<paragraph id="18E98CAF849F225CD0D6010A235E0118" pageId="5" pageNumber="18">
<taxonomicName id="6D80E93EB6B3B58E4EA31DE5AD871930" authority="Motschulsky, 1874" authorityName="Motschulsky" authorityYear="1874" genus="Kytorhinus" lsidName="Kytorhinus immixtus" pageId="5" pageNumber="18" rank="species" species="immixtus">Kytorhinus immixtus Motschulsky, 1874</taxonomicName>
Figs 29-30, 43
</paragraph>
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<paragraph pageId="5" pageNumber="18">Material.</paragraph>
<paragraph pageId="5" pageNumber="18">
<subSubSection id="6FADC515E4907B4DCAA2CDE9A21C5F9B" pageId="5" pageNumber="18" type="material">
<paragraph id="20206FC256C16A54CBA8DC9CED5FE75E" pageId="5" pageNumber="18">Material.</paragraph>
<paragraph id="4B9C941798F606E5735A7A7B17C6B4CB" pageId="5" pageNumber="18">
1♂, Pochengzi, Wensu, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
<geoCoordinate id="387D79CEC90654FE1861CD2D5A294B49" direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.99">80.99°E</geoCoordinate>
<geoCoordinate id="ADF5D78E4877A972FD9318F65A072C88" direction="east" orientation="longitude" precision="555" value="80.99">80.99°E</geoCoordinate>
, alt. ca 2000 m, 1978.VI.15.
</paragraph>
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<subSubSection pageId="5" pageNumber="18" type="distribution">
<paragraph pageId="5" pageNumber="18">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="18">China, Kyrgyzstan, Russia.</paragraph>
<caption pageId="5" pageNumber="18">
<paragraph pageId="5" pageNumber="18">
<subSubSection id="73816FE0B3255186B827C86413A7659D" pageId="5" pageNumber="18" type="distribution">
<paragraph id="C2839C707353A17985EE04DC71C78F7D" pageId="5" pageNumber="18">Distribution.</paragraph>
<paragraph id="362C3BB1EE86ACC9851DA839E3267357" pageId="5" pageNumber="18">China, Kyrgyzstan, Russia.</paragraph>
<caption id="74E9DBFDA7332D155FFC8C19C9C1B6F4" pageId="5" pageNumber="18">
<paragraph id="F755E3707FCD8025BEF3C3B0A1D89EFC" pageId="5" pageNumber="18">
Figures 39-42. Antennae. 39
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus thermopsis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="thermopsis">Kytorhinus thermopsis</taxonomicName>
<taxonomicName id="FF6BAC2B566BCA383AF206E7D26A62A8" genus="Kytorhinus" lsidName="Kytorhinus thermopsis" pageId="5" pageNumber="18" rank="species" species="thermopsis">Kytorhinus thermopsis</taxonomicName>
male 40
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus thermopsis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="thermopsis">Kytorhinus thermopsis</taxonomicName>
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female 41
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus kergoati" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="kergoati">Kytorhinus kergoati</taxonomicName>
<taxonomicName id="C1E47508CCACA005FBE345A7EC4E7F55" genus="Kytorhinus" lsidName="Kytorhinus kergoati" pageId="5" pageNumber="18" rank="species" species="kergoati">Kytorhinus kergoati</taxonomicName>
male 42
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus kergoati" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="kergoati">Kytorhinus kergoati</taxonomicName>
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female 43-44 Head 43
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus immixtus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="immixtus">Kytorhinus immixtus</taxonomicName>
<taxonomicName id="D9C501A299B937A782AA87BBDBE2786C" genus="Kytorhinus" lsidName="Kytorhinus immixtus" pageId="5" pageNumber="18" rank="species" species="immixtus">Kytorhinus immixtus</taxonomicName>
44
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Kytorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kytorhinus thermopsis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="thermopsis">Kytorhinus thermopsis</taxonomicName>
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45-46 Lateral pronotal margin. 45
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus affinis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="affinis">Bruchus affinis</taxonomicName>
<taxonomicName id="F875E4EC550022B1F015AA4734361013" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus affinis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="affinis">Bruchus affinis</taxonomicName>
46
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus rufimanus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="rufimanus">Bruchus rufimanus</taxonomicName>
<taxonomicName id="77D05BB9E6375F7418FAB9967A4AF453" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus rufimanus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="rufimanus">Bruchus rufimanus</taxonomicName>
47 Hind trochanters 47
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Rhaebus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhaebus solskyi" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="solskyi">Rhaebus solskyi</taxonomicName>
<taxonomicName id="091B541B2069D8A34C52BD57E81F52E4" genus="Rhaebus" lsidName="Rhaebus solskyi" pageId="5" pageNumber="18" rank="species" species="solskyi">Rhaebus solskyi</taxonomicName>
48-51 Hind femur. 48
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Rhaebus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhaebus solskyi" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="solskyi">Rhaebus solskyi</taxonomicName>
<taxonomicName id="538A71E423172481386A4F071D1745F0" genus="Rhaebus" lsidName="Rhaebus solskyi" pageId="5" pageNumber="18" rank="species" species="solskyi">Rhaebus solskyi</taxonomicName>
male 49
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthoscelides pallidipennis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="pallidipennis">Acanthoscelides pallidipennis</taxonomicName>
<taxonomicName id="E3D9170F916E802F23EE540BF90EFD01" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthoscelides pallidipennis" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="pallidipennis">Acanthoscelides pallidipennis</taxonomicName>
50
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculatus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="maculatus">Callosobruchus maculatus</taxonomicName>
<taxonomicName id="566FEA37787E0E4FE212EFEE1F81A53E" class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculatus" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="maculatus">Callosobruchus maculatus</taxonomicName>
outside view 51
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculates" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="maculates">Callosobruchus maculates</taxonomicName>
<taxonomicName id="E60D65C6CF397592A9D561C5FAF23C1F" class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculates" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="maculates">Callosobruchus maculates</taxonomicName>
inside view.
</paragraph>
</caption>

View file

@ -1,154 +1,153 @@
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<paragraph pageId="2" pageNumber="15">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<paragraph id="F54113B67353E8E334D60D7CF2849FA2" pageId="2" pageNumber="15">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<paragraph pageId="2" pageNumber="15">
<taxonomicName authority="Geoffroy, 1785" authorityName="Geoffroy" authorityYear="1785" class="Insecta" family="Chrysomelidae" genus="Spermophagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Spermophagus sericeus" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="sericeus">Spermophagus sericeus (Geoffroy, 1785)</taxonomicName>
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<paragraph id="F5115DB8AD7CA7AF6DE4E1C09B386122" pageId="2" pageNumber="15">
<taxonomicName id="A74F362A31F72F1B0B7FE33C7666BEFA" authority="Geoffroy, 1785" authorityName="Geoffroy" authorityYear="1785" genus="Spermophagus" lsidName="Spermophagus sericeus" pageId="2" pageNumber="15" rank="species" species="sericeus">Spermophagus sericeus (Geoffroy, 1785)</taxonomicName>
Figs 1-2
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<paragraph pageId="2" pageNumber="15">Material.</paragraph>
<paragraph pageId="2" pageNumber="15">
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<paragraph id="5979AECF32FB26D219099C88208E0DC9" pageId="2" pageNumber="15">Material.</paragraph>
<paragraph id="E9101B4B9D727339CFEBCCF25C3A2281" pageId="2" pageNumber="15">
2♂, Akqi, Kizilsu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="40.98">40.98°N</geoCoordinate>
<geoCoordinate id="B910CA41BD372DE0FD4DECCFB8E2FBAB" direction="north" orientation="latitude" precision="555" value="40.98">40.98°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="78.7">78.70°E</geoCoordinate>
<geoCoordinate id="7BEF3C454A59E38EF7D5E830A58D5DD1" direction="east" orientation="longitude" precision="555" value="78.7">78.70°E</geoCoordinate>
, alt. ca 1970 m, 2005.VI.14, H.Y. Hu leg.; 1♀6♂, Aksu, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="40.94">40.94°N</geoCoordinate>
<geoCoordinate id="A71561E9466986E3A96F924E56E357F3" direction="north" orientation="latitude" precision="555" value="40.94">40.94°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="80.11">80.11°E</geoCoordinate>
<geoCoordinate id="D2F77F958971EABCB23924A339DC056E" direction="east" orientation="longitude" precision="555" value="80.11">80.11°E</geoCoordinate>
, alt. ca 1180 m, 1978.VI.19, Y.H. Han leg.; IOZ(E)1016347-1016583; 1♀, Qinggil, Altay, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="46.69">46.69°N</geoCoordinate>
<geoCoordinate id="98634C7A399210F8F1373DA167E4E92A" direction="north" orientation="latitude" precision="555" value="46.69">46.69°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="90.39">90.39°E</geoCoordinate>
<geoCoordinate id="67E0E28C19D8AB6D2CE3E57C6082C5FE" direction="east" orientation="longitude" precision="555" value="90.39">90.39°E</geoCoordinate>
, alt. ca 1390 m, 1956.VIII.1, W.Y. Yang leg., IOZ(E)632314; 1♀1♂, Turpan, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="42.93">42.93°N</geoCoordinate>
<geoCoordinate id="30BFE7516FFC0EFB4E009244C47B97A7" direction="north" orientation="latitude" precision="555" value="42.93">42.93°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="89.27">89.27°E</geoCoordinate>
<geoCoordinate id="2CDCCA8C261E45AB2190A8575765F952" direction="east" orientation="longitude" precision="555" value="89.27">89.27°E</geoCoordinate>
, alt. ca 140 m, 1958.V.20, C.Q. Li and G. Wang leg., IOZ(E)632431, IOZ(E)632433; 3♀3♂, Baicheng, Aksu,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.78">41.78°N</geoCoordinate>
<geoCoordinate id="661837AE0D762CE5D709341CCCAB86BF" direction="north" orientation="latitude" precision="555" value="41.78">41.78°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="81.92">81.92°E</geoCoordinate>
<geoCoordinate id="CA87172A66F1CCB9AFBEDA21250C04F4" direction="east" orientation="longitude" precision="555" value="81.92">81.92°E</geoCoordinate>
, alt. ca 1310 m, 1959.VII.22, A.F. Tian leg., IOZ(E)115170-115176; 1♀3♂, Yuli, Mila, Bayingol, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
<geoCoordinate id="E44CBC9116C7553F94763A85A18A5F84" direction="north" orientation="latitude" precision="555" value="41.77">41.77°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="84.24">84.24°E</geoCoordinate>
<geoCoordinate id="8A311E8728056F08863D44B390CAC76A" direction="east" orientation="longitude" precision="555" value="84.24">84.24°E</geoCoordinate>
, alt. ca 1000 m, 1958.VII.13, C.Q. Li leg., IOZ(E)115143-115146; 1♀1♂, Urumqi, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="43.83">43.83°N</geoCoordinate>
<geoCoordinate id="1DEF3590850DCBA31F6338777BEF807B" direction="north" orientation="latitude" precision="555" value="43.83">43.83°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="87.55">87.55°E</geoCoordinate>
<geoCoordinate id="2D2ADEA80A01233799B6516677E42A69" direction="east" orientation="longitude" precision="555" value="87.55">87.55°E</geoCoordinate>
, alt. ca 820 m, 1955.VII.25, S.J. Ma, K.L. Xia and Y.L.Chen leg., IOZ(E)115133-115134; 2♀2♂, Usu, Qoqek, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="45.02">45.02°N</geoCoordinate>
<geoCoordinate id="8A1DDEB656300C310EE95997A01A9259" direction="north" orientation="latitude" precision="555" value="45.02">45.02°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="84.78">84.78°E</geoCoordinate>
<geoCoordinate id="3108E3D201685A08E833A0B4FFBD5A11" direction="east" orientation="longitude" precision="555" value="84.78">84.78°E</geoCoordinate>
, alt. ca 290 m, 1957.VI.16, G. Wang leg., IOZ(E)115106-115109; 10♀16♂, Shihezi, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="44.28">44.28°N</geoCoordinate>
<geoCoordinate id="8F3DA15A1A49F00DB22614596E5D01B8" direction="north" orientation="latitude" precision="555" value="44.28">44.28°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="86.27">86.27°E</geoCoordinate>
<geoCoordinate id="00F663E41282355BC9AD76CE25032128" direction="east" orientation="longitude" precision="555" value="86.27">86.27°E</geoCoordinate>
, alt. ca 500 m, 1957.VI.7, G. Wang and C.P. Hong leg., IOZ(E)115080-115106; 1♀6♂, Shawan, Qoqek, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="44.46">44.46°N</geoCoordinate>
<geoCoordinate id="9ACEB9FA7E0A96D3974EF11310C8DDD8" direction="north" orientation="latitude" precision="555" value="44.46">44.46°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="85.66">85.66°E</geoCoordinate>
<geoCoordinate id="93FA0EB14A46C42C4D937D3927D3EA54" direction="east" orientation="longitude" precision="555" value="85.66">85.66°E</geoCoordinate>
, alt. ca 420 m, 1957.VI.11, G. Wang and C.P. Hong leg., IOZ(E)115063-115169; 1♀, Jeminay, Altay, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="47.43">47.43°N</geoCoordinate>
<geoCoordinate id="E62F5313522A95195FAFE61B45D29C53" direction="north" orientation="latitude" precision="555" value="47.43">47.43°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="85.87">85.87°E</geoCoordinate>
<geoCoordinate id="F84113597447C3862554596254E7E653" direction="east" orientation="longitude" precision="555" value="85.87">85.87°E</geoCoordinate>
, alt. ca 970 m, 1956.IX.17, W.Y. Yang leg., IOZ(E)115054.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="15" type="distribution">
<paragraph pageId="2" pageNumber="15">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="15">Widely distributed around the Palaearctic region.</paragraph>
<caption pageId="2" pageNumber="15">
<paragraph pageId="2" pageNumber="15">
<subSubSection id="AD20CE72227B993C60F5FA0606718030" pageId="2" pageNumber="15" type="distribution">
<paragraph id="11151E45B9D5F66EBC032401BCBAC02F" pageId="2" pageNumber="15">Distribution.</paragraph>
<paragraph id="35F4E4BE9633D978B558E0B10D618A6C" pageId="2" pageNumber="15">Widely distributed around the Palaearctic region.</paragraph>
<caption id="8141F566C82FA6B3A8076D5410983C06" pageId="2" pageNumber="15">
<paragraph id="63A0B20BF4AAB9123886C3C290B0C321" pageId="2" pageNumber="15">
Figures 1-12.
<taxonomicName lsidName="" pageId="2" pageNumber="15" rank="subfamily" subfamily="Bruchinae">Bruchinae</taxonomicName>
<taxonomicName id="EB2A5A77161D865B1F413DE60D404981" lsidName="" pageId="2" pageNumber="15" rank="subfamily" subfamily="Bruchinae">Bruchinae</taxonomicName>
in Xinjiang, dorsal and lateral view. 1-2
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Spermophagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Spermophagus sericeus" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="sericeus">Spermophagus sericeus</taxonomicName>
<taxonomicName id="E8571EC17D0985866A8D84B0887AC7F0" genus="Spermophagus" lsidName="Spermophagus sericeus" pageId="2" pageNumber="15" rank="species" species="sericeus">Spermophagus sericeus</taxonomicName>
3-4
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthoscelides pallidipennis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="pallidipennis">Acanthoscelides pallidipennis</taxonomicName>
<taxonomicName id="750BAD8BC6CC84274DD9A8AC99EE4805" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthoscelides pallidipennis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="pallidipennis">Acanthoscelides pallidipennis</taxonomicName>
5-6
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchidius apicipennis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="apicipennis">Bruchidius apicipennis</taxonomicName>
<taxonomicName id="C9B79323BF69A1CBA68222077A23805B" class="Insecta" family="Chrysomelidae" genus="Bruchidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchidius apicipennis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="apicipennis">Bruchidius apicipennis</taxonomicName>
7-8
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchidius tuberculicauda" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="tuberculicauda">Bruchidius tuberculicauda</taxonomicName>
<taxonomicName id="5BB025E15637E6F1A830974CF4A0F44D" class="Insecta" family="Chrysomelidae" genus="Bruchidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchidius tuberculicauda" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="tuberculicauda">Bruchidius tuberculicauda</taxonomicName>
9-10
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus chinensis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="chinensis">Callosobruchus chinensis</taxonomicName>
<taxonomicName id="B168287C8A79FAE62C72DE382EDD26DB" class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus chinensis" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="chinensis">Callosobruchus chinensis</taxonomicName>
11-12
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculatus" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="maculatus">Callosobruchus maculatus</taxonomicName>
<taxonomicName id="F5CAA87B164B41DDFF1F4CECCF17D95B" class="Insecta" family="Chrysomelidae" genus="Callosobruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callosobruchus maculatus" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="maculatus">Callosobruchus maculatus</taxonomicName>
; black bar = 1 mm.
</paragraph>
</caption>

View file

@ -1,116 +1,115 @@
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<taxonomicName authority="Fabraeus, 1839" authorityName="Fabraeus" authorityYear="1839" class="Insecta" family="Chrysomelidae" genus="Megabruchidius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megabruchidius dorsalis" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="dorsalis">Megabruchidius dorsalis (Fabraeus, 1839)</taxonomicName>
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<taxonomicName id="793ED17E9478EC17FA846E622EBEE977" ID-CoL="6MNCK" authority="Fabraeus, 1839" authorityName="Fabraeus" authorityYear="1839" genus="Megabruchidius" lsidName="Megabruchidius dorsalis" pageId="4" pageNumber="17" rank="species" species="dorsalis">Megabruchidius dorsalis (Fabraeus, 1839)</taxonomicName>
Figs 13-14, 66-67
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="17" type="material">
<paragraph pageId="4" pageNumber="17">Material.</paragraph>
<paragraph pageId="4" pageNumber="17">
<subSubSection id="F998696DF00E30A16DA1BA34D79C387A" pageId="4" pageNumber="17" type="material">
<paragraph id="9FF9D5A3AAF25090E42FA6BBBD9BDA16" pageId="4" pageNumber="17">Material.</paragraph>
<paragraph id="78580A171B10CD8C6FE588111A5337B6" pageId="4" pageNumber="17">
10♀5♂, Ili Forestry Science Research Institute, Gulja, Ili, Xinjiang,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="43.94">43.94°N</geoCoordinate>
<geoCoordinate id="E959A22B0272FC004E15FBC37132F7E4" direction="north" orientation="latitude" precision="555" value="43.94">43.94°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="81.33">81.33°E</geoCoordinate>
<geoCoordinate id="62AF3568B6E7ACEB76E4660931EC4E02" direction="east" orientation="longitude" precision="555" value="81.33">81.33°E</geoCoordinate>
, alt. ca 660 m, 1973.VII.5, IOZ(E)109814-109818, 632556-632565.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="17" type="distribution">
<paragraph pageId="4" pageNumber="17">Distribution.</paragraph>
<paragraph pageId="4" pageNumber="17">Bulgaria, China, France, Greece, Hong Kong, Hungary, India, Italy, Japan, Mongolia, Papua New Guinea, Switzerland, Turkmenistan.</paragraph>
<caption pageId="4" pageNumber="17">
<paragraph pageId="4" pageNumber="17">
<subSubSection id="56F74CB05FD1D61D153E845839DA9296" pageId="4" pageNumber="17" type="distribution">
<paragraph id="9B1DA69FBCE76DC699006C1790EF7B0A" pageId="4" pageNumber="17">Distribution.</paragraph>
<paragraph id="483390A764E905A5053D799B1B4DC540" pageId="4" pageNumber="17">Bulgaria, China, France, Greece, Hong Kong, Hungary, India, Italy, Japan, Mongolia, Papua New Guinea, Switzerland, Turkmenistan.</paragraph>
<caption id="56EDD54A004F51C44EFB371DB93BECFE" pageId="4" pageNumber="17">
<paragraph id="403074255FA34DD77D6E2D8826411605" pageId="4" pageNumber="17">
Figures 13-26.
<taxonomicName lsidName="" pageId="4" pageNumber="17" rank="subfamily" subfamily="Bruchinae">Bruchinae</taxonomicName>
<taxonomicName id="341A8772F8BC00847BB9844BC2BDA442" lsidName="" pageId="4" pageNumber="17" rank="subfamily" subfamily="Bruchinae">Bruchinae</taxonomicName>
in Xinjiang, dorsal and lateral view. 13-14
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Megabruchidius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megabruchidius dorsalis" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="dorsalis">Megabruchidius dorsalis</taxonomicName>
<taxonomicName id="7298E2718437272F1C17FD64D180101E" genus="Megabruchidius" lsidName="Megabruchidius dorsalis" pageId="4" pageNumber="17" rank="species" species="dorsalis">Megabruchidius dorsalis</taxonomicName>
15-16
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus affinis" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="affinis">Bruchus affinis</taxonomicName>
<taxonomicName id="CAFAEDC80EA1668514F8322CF09B8E17" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus affinis" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="affinis">Bruchus affinis</taxonomicName>
17-18
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus atomarius" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="atomarius">Bruchus atomarius</taxonomicName>
<taxonomicName id="3517CC7B24EDEB094BAB2C320733B5CF" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus atomarius" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="atomarius">Bruchus atomarius</taxonomicName>
19-20
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus dentipes" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="dentipes">Bruchus dentipes</taxonomicName>
<taxonomicName id="2BCF072DDA14A794BEFEC71F6FDE3E8F" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus dentipes" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="dentipes">Bruchus dentipes</taxonomicName>
21-22
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus loti" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="loti">Bruchus loti</taxonomicName>
<taxonomicName id="92C9FACB3BA814C89CA637A276C87CC7" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus loti" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="loti">Bruchus loti</taxonomicName>
23-24
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus pisorum" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="pisorum">Bruchus pisorum</taxonomicName>
<taxonomicName id="94A2033D995AA325D7FA746794A1F841" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus pisorum" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="pisorum">Bruchus pisorum</taxonomicName>
25-26
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus rufimanus" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="rufimanus">Bruchus rufimanus</taxonomicName>
<taxonomicName id="2CB6CDC11D4C9C0ECACF892BC0E6DBC9" class="Insecta" family="Chrysomelidae" genus="Bruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bruchus rufimanus" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="rufimanus">Bruchus rufimanus</taxonomicName>
; black bar = 1 mm.
</paragraph>
</caption>

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@ -1,223 +1,222 @@
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<treatment id="ED7A28A8921CB02C09F787A55533E70A" ID-GBIF-Taxon="148671565" LSID="urn:lsid:plazi:treatment:ED7A28A8921CB02C09F787A55533E70A" httpUri="http://treatment.plazi.org/id/ED7A28A8921CB02C09F787A55533E70A" lastPageId="5" lastPageNumber="56" pageId="3" pageNumber="54" scope_family="Pholcidae" scope_order="Araneae">
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Subfamily
<taxonomicName authority="Simon, 1893" authorityName="Simon" authorityYear="1893" family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="3" pageNumber="54" rank="subFamily" subFamily="Arteminae">Arteminae Simon, 1893</taxonomicName>
<taxonomicName id="473B64DD300E6851F9C7953F5751D5F3" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="subFamily" subFamily="Arteminae">Arteminae Simon, 1893</taxonomicName>
Figure 2
</paragraph>
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<subSubSection pageId="4" pageNumber="55" type="reference_group">
<paragraph pageId="4" pageNumber="55">
<taxonomicName family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Artemeae" order="Araneae" pageId="4" pageNumber="55" rank="subFamily" subFamily="Artemeae">
<pageBreakToken pageId="4" pageNumber="55" start="start">Artemeae</pageBreakToken>
<subSubSection id="666F0618B99762421ED282D44FA75553" pageId="4" pageNumber="55" type="reference_group">
<paragraph id="B68FBA40C629B82CBA20095E8E139324" pageId="4" pageNumber="55">
<taxonomicName id="73130B5491362B3D3E9CD0A58C50AA7C" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artemeae" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="subFamily" subFamily="Artemeae">
<pageBreakToken id="953A20960D2003851F693C63ADAC318A" pageId="4" pageNumber="55" start="start">Artemeae</pageBreakToken>
</taxonomicName>
Simon, 1893: 463. Type genus
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<taxonomicName id="2E3BE943C50626477E89A4A324D5C811" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
Walckenaer, 1837, by monotypy.
</paragraph>
<paragraph pageId="4" pageNumber="55">
<taxonomicName family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<paragraph id="CAADAC53BAF25094E4C1D38BA7E9477D" pageId="4" pageNumber="55">
<taxonomicName id="30601DE2AB87077A12CFFC15B4D649B5" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
Simon;
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
<bibRefCitation id="AF7BA0F6EC0CB77ED7DAB2336683BFE2" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
: 212.
</paragraph>
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<paragraph pageId="4" pageNumber="55">Remarks.</paragraph>
<paragraph lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
<subSubSection id="0526F3BC756AB4F84F24615E8BE85C51" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55" type="remarks">
<paragraph id="1B667D55F54211EF5C601045B3721F05" pageId="4" pageNumber="55">Remarks.</paragraph>
<paragraph id="F2A4459A4D444A4023CC6BE8EE1F88D6" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
All our analyses exclude the name-giving genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="41D23A2794DDA4D46CF7FBE5925F026B" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
from the clade containing all other
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="D291DD36683B5B7418DFDB22986422D5" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
and invariably place
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="D29C7E0A360FE912BC4EB0D41DE39E78" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
as sister to
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="5C246E02650C18A2F7B76DF9206EC5F1" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
(Figure 2), formally precluding the use of the name
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="2B95756B0F9081891416F80E61B6EE42" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
for this clade. We do not propose a new subfamily name for this clade but treat it as 'other
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="79A3A5A2344821545439646C436EC7CF" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' because we consider the position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="A256402583E15A70B30CE89EE03BBA50" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
dubious.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="191132B34BAFD7EB8EDEFFBBE19AC5D4" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
shares with 'other
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="299BA865FD57D624C1B344C492CE68B4" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' a unique pair of structures on the procursus: a ventral pocket and a dorsal apophysis. These structures are associated with asymmetric palp insertion in both species studied with respect to this detail [
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus globosus" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="globosus">Physocyclus globosus</taxonomicName>
<taxonomicName id="062E2FC5D379508EA68681FBB0C60A91" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus globosus" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="globosus">Physocyclus globosus</taxonomicName>
(Taczanowski, 1874),
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema nephilit" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="nephilit">Artema nephilit</taxonomicName>
<taxonomicName id="B80E4C91A165B28E99F9724A6CFFC5CE" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema nephilit" order="Araneae" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="nephilit">Artema nephilit</taxonomicName>
Aharon et al., 2017:
<bibRefCitation author="Huber, BA" journalOrPublisher="Canadian Journal of Zoology" pageId="41" pageNumber="92" pagination="905 - 918" title="Courtship, copulation, and genital mechanics in Physocyclusglobosus (Araneae, Pholcidae)." url="https://doi.org/10.1139/z97-109" volume="74" year="1997">Huber and Eberhard 1997</bibRefCitation>
<bibRefCitation id="6DB20144EFA6799AF88EC47B5C00F40E" author="Huber, BA" journalOrPublisher="Canadian Journal of Zoology" pageId="41" pageNumber="92" pagination="905 - 918" title="Courtship, copulation, and genital mechanics in Physocyclusglobosus (Araneae, Pholcidae)." url="https://doi.org/10.1139/z97-109" volume="74" year="1997">Huber and Eberhard 1997</bibRefCitation>
,
<bibRefCitation author="Aharon, S" journalOrPublisher="European Journal of Taxonomy" pageId="38" pageNumber="89" pagination="1 - 57" title="Daddy-long-leg giants: Revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)." url="https://doi.org/10.5852/ejt.2017.376" volume="376" year="2017">Aharon et al. 2017</bibRefCitation>
<bibRefCitation id="4415124BDC4784D3BA3DAE9ACCAD58AB" author="Aharon, S" journalOrPublisher="European Journal of Taxonomy" pageId="38" pageNumber="89" pagination="1 - 57" title="Daddy-long-leg giants: Revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)." url="https://doi.org/10.5852/ejt.2017.376" volume="376" year="2017">Aharon et al. 2017</bibRefCitation>
]. The structures are present in all
<taxonomicName lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="34BAF35430CEFCEFD7E32B6734F28FE2" lsidName="" pageId="4" pageNumber="55" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
, even in taxa that were previously thought to be representatives
<pageBreakToken pageId="5" pageNumber="56" start="start">of</pageBreakToken>
<pageBreakToken id="3CD48715F387F61045EED9EEEF81DA2C" pageId="5" pageNumber="56" start="start">of</pageBreakToken>
other subfamilies, such as
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
<taxonomicName id="0ECFEB00E7D8D817EB8D6CEB866C43BB" class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="CF05A7DA997F0E869208522074A5AF71" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
(previously in
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="48AC14F7DEBEA739E62EEDB46C98D74D" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
; see above), and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
<taxonomicName id="BB99271DF7EBDB7182EA3FF8AF119289" class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
Huber, 2001 (previously in
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="6BCDDFDCC9082B48FABD55B5D017BB4B" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
; see below) (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="4806002A1EE95FEA058DF407B75B7E22" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">2001</bibRefCitation>
<bibRefCitation id="990886D989A064940B40D8A7AE97F2DA" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">2001</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="45 - 53" title="On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar." volume="1635" year="2007">Huber and El Hennawy 2007</bibRefCitation>
<bibRefCitation id="9D410FF9D16232CD5B6D39381D9BEE7E" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="45 - 53" title="On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar." volume="1635" year="2007">Huber and El Hennawy 2007</bibRefCitation>
). By contrast, these structures are apparently absent in all other
<taxonomicName family="Pholcidae" lsidName="" pageId="5" pageNumber="56" rank="family">Pholcidae</taxonomicName>
<taxonomicName id="A5C430F385DF19F06883B4268460835D" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
. Curiously and unexplainable to us, previous molecular analyses have supported a position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="54FA87E6E82A4CA94221AE6FFCBCF08D" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
among 'other
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="A51E1B33E2991C5E9C216343C3133476" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' (
<bibRefCitation author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
<bibRefCitation id="65A5F152AE336DA762DC5BEBBB31CF0E" author="Astrin, JJ" journalOrPublisher="Zootaxa" pageId="39" pageNumber="90" pagination="17 - 30" title="The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae)." volume="1646" year="2007">Astrin et al. 2007</bibRefCitation>
: fig. 1,
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="1D8C0BB093B6BD1CA973F51923F1AB71" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
).
</paragraph>
<paragraph pageId="5" pageNumber="56">
<paragraph id="1519FDCFFF9B67CB40466E92A1338134" pageId="5" pageNumber="56">
Some of the 99 currently known species of
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="B3A8DF1D58CE4F5A3F3505953EA3C335" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
are relatively large spiders with long, strong legs and high globose abdomens. The genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="F86DA446AF7E597E766588FA48605E55" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
, in particular, includes probably the largest pholcids in terms of body mass (
<bibRefCitation author="Aharon, S" journalOrPublisher="European Journal of Taxonomy" pageId="38" pageNumber="89" pagination="1 - 57" title="Daddy-long-leg giants: Revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)." url="https://doi.org/10.5852/ejt.2017.376" volume="376" year="2017">Aharon et al. 2017</bibRefCitation>
<bibRefCitation id="B3C80CE8FD269564BA6B09A26CF3C1C7" author="Aharon, S" journalOrPublisher="European Journal of Taxonomy" pageId="38" pageNumber="89" pagination="1 - 57" title="Daddy-long-leg giants: Revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)." url="https://doi.org/10.5852/ejt.2017.376" volume="376" year="2017">Aharon et al. 2017</bibRefCitation>
). However, tiny species that were previously assigned to
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="930FCD38211A18B48B3EBFCDB6515DAE" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
partly because of their size (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
<taxonomicName id="D02079B48B7EBD3D38153B0DC7F8B0EF" class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="2012E5F8223685C8608707A9AB939BCA" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
) are now included in
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="2A9EDC23697D91FF77373378A540D307" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
, and their
<normalizedToken originalValue="basal">'basal'</normalizedToken>
<normalizedToken id="2B4341D011D616214ACAEA112174E90B" originalValue="basal">'basal'</normalizedToken>
position in the cladogram suggests that ancestral
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="C4D59A20BDCCC0546D6B80A7DCD2F52D" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
may in fact have been tiny. Just like
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
<taxonomicName id="3726EFBBB08753B7F0526D15E5BA6D78" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
,
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="506971BE01A9FE56134FEEAF83F05BD5" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
often occur in rather dry regions, sometimes even in deserts like the Australian
<taxonomicName class="Arachnida" family="Pholcidae" genus="Trichocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Trichocyclus</taxonomicName>
<taxonomicName id="C5ED420AB66CD150E74364C42DDC01DF" class="Arachnida" family="Pholcidae" genus="Trichocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Trichocyclus</taxonomicName>
Simon, 1908. They have a wide distribution, but are apparently absent from Sub-Saharan Africa and from South America (except for
<normalizedToken originalValue="“Geneve59”">&quot;Geneve59&quot;</normalizedToken>
<normalizedToken id="8B8674C6D91481F55D3B9A2900172B92" originalValue="“Geneve59”">&quot;Geneve59&quot;</normalizedToken>
, a tiny undescribed species representing a new undescribed genus on
<normalizedToken originalValue="Curaçao">Curacao</normalizedToken>
<normalizedToken id="DA093CF9425ED10D0E6CE196BE505673" originalValue="Curaçao">Curacao</normalizedToken>
and Aruba).
</paragraph>
<paragraph pageId="5" pageNumber="56">
<paragraph id="607DECB1CCF191757EDB3E65CB4634D1" pageId="5" pageNumber="56">
The monophyly of 'other
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="C22235ECB50951CC08CACB23B7C4C061" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' is supported in all our analysis, even though with low support (possibly because of the dubious position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="B40D83E02537018BC43A270367C87B62" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
, see above). Similar to our previous analysis (i.e. except for the position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="EF264AEB56B40F77B2F8FB6A1376CEED" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
;
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="09D19627F75DE1A93C61C0BCF6751DC1" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
), 'other
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="B581D6A9AC687359E979248B25F0ECB6" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' is sister to
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="AE2B49F6FE20CF1F7BDD2BAAD2E02EB1" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
, with variable support (reasonable support only in the RogueNaRok tree; in other trees, bootstrap support is low but SH values range from 82 to 99). This sister group relationship is weakly supported by morphology: 'other
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="DA2B49C0A4BBB0E3FD83F51EF7A37E3A" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' and
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="BBD55C4A97552F93C9A9F40D7B18D569" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
lack epiandrous spigots. However, epiandrous spigots have been lost several times convergently in
<taxonomicName family="Pholcidae" lsidName="" pageId="5" pageNumber="56" rank="family">Pholcidae</taxonomicName>
<taxonomicName id="254C4ED57741EF95B3A6522B40CB05C9" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
(
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="1A8B4FC048B8848D603173178CDCD756" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
, BA Huber, unpubl. data).
</paragraph>
<paragraph pageId="5" pageNumber="56">
<paragraph id="4DC4203C97AFD2B749E7A8E9F904139D" pageId="5" pageNumber="56">
Internal relationships in 'other
<taxonomicName lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="806F1D9CC87D380E0BB02CB3A4EB2C72" lsidName="" pageId="5" pageNumber="56" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' are partly resolved with reasonable support. The data suggest a large Indomalayan-Australasian clade, including the genera
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<taxonomicName id="18C1CB4CDE8CB5593D7D347E2E03AAE4" class="Arachnida" family="Pholcidae" genus="Trichocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Trichocyclus</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
<taxonomicName id="4B3F991BADC31A14E83641F0C78269CF" class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
(Australia),
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocneminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocneminus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Holocneminus</taxonomicName>
<taxonomicName id="3BA962D7C08AEC05770065D2630324AE" class="Arachnida" family="Pholcidae" genus="Holocneminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocneminus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Holocneminus</taxonomicName>
Berland, 1942 (SE Asia and Pacific; excluding the misplaced and highly isolated
<taxonomicName lsidName="H. huangdi" pageId="5" pageNumber="56" rank="species" species="huangdi">H. huangdi</taxonomicName>
<taxonomicName id="ACC722EBD27D73D047D51698D949734C" lsidName="H. huangdi" pageId="5" pageNumber="56" rank="species" species="huangdi">H. huangdi</taxonomicName>
Tong &amp; Li, 2009), and a new undescribed genus (without any described species; ranging from Eastern Indonesia to the Pacific). Sister to this clade is either the New World genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
<taxonomicName id="1A397AB9A2E597D6CDB9736D945EC4DA" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
Simon, 1893 alone or
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
<taxonomicName id="33C434D7C14AC688A6DFC7EB048314DE" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
together with the Middle-Eastern monotypic
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="340F9CCEAE6A0FABF491E7CE0203C8DC" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
. However, support values for any of these options are low and morphological data do not favour (nor contradict) any of them. Finally, the
<normalizedToken originalValue="basal">'basal'</normalizedToken>
<normalizedToken id="B75CF8A8072644C25CD9E0EF5D3E0663" originalValue="basal">'basal'</normalizedToken>
branches, i.e., those leading to the taxa outside the Indomalayan-Australasian clade and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
<taxonomicName id="4ABA873D737A36828C6B57F65B32A9CE" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
(and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
<taxonomicName id="8F88C7D1AE6BD98DEBD2E24272F58242" class="Arachnida" family="Pholcidae" genus="Nita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nita" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Nita</taxonomicName>
in the case of the IQ-TREE analysis) lead to a group of North American and Caribbean taxa (the North American genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
<taxonomicName id="89E52CDBC8134DBE66C3AA62522966CD" class="Arachnida" family="Pholcidae" genus="Chisosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chisosa" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Chisosa</taxonomicName>
being sister to a tiny undescribed species representing a new undescribed genus on
<normalizedToken originalValue="Curaçao">Curacao</normalizedToken>
<normalizedToken id="8D1CDB536EDEEF6EA2F4D9C35C9DFFCD" originalValue="Curaçao">Curacao</normalizedToken>
and Aruba:
<normalizedToken originalValue="“Geneve59”">&quot;Geneve59&quot;</normalizedToken>
<normalizedToken id="37DF8BD899A37DD09F8A1DCB559950F7" originalValue="“Geneve59”">&quot;Geneve59&quot;</normalizedToken>
), and to the SE-Asian
<taxonomicName class="Arachnida" family="Pholcidae" genus="Holocneminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocneminus huangdi" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="huangdi">Holocneminus huangdi</taxonomicName>
<taxonomicName id="45754FED875A9EB58397B7B7B60EC389" class="Arachnida" family="Pholcidae" genus="Holocneminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holocneminus huangdi" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="huangdi">Holocneminus huangdi</taxonomicName>
, an isolated species that appears misplaced also by morphological criteria (A
<normalizedToken originalValue="Valdez-Mondragón">Valdez-Mondragon</normalizedToken>
<normalizedToken id="7314E0CB816676FCB71C47DD607455F2" originalValue="Valdez-Mondragón">Valdez-Mondragon</normalizedToken>
, pers. comm., Nov. 2015).
</paragraph>
</subSubSection>

View file

@ -1,296 +1,295 @@
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<treatment ID-GBIF-Taxon="148671563" LSID="urn:lsid:plazi:treatment:EE63F84511929B3C0CEF1E7BA1408A0A" httpUri="http://treatment.plazi.org/id/EE63F84511929B3C0CEF1E7BA1408A0A" lastPageNumber="63" pageId="12" pageNumber="63">
<subSubSection pageId="12" pageNumber="63" type="nomenclature">
<paragraph pageId="12" pageNumber="63">
<treatment id="EE63F84511929B3C0CEF1E7BA1408A0A" ID-GBIF-Taxon="148671563" LSID="urn:lsid:plazi:treatment:EE63F84511929B3C0CEF1E7BA1408A0A" httpUri="http://treatment.plazi.org/id/EE63F84511929B3C0CEF1E7BA1408A0A" lastPageNumber="63" pageId="12" pageNumber="63" scope_family="Pholcidae" scope_order="Araneae" scope_subFamily="Smeringopinae">
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<paragraph id="C7C27836AA19495625A86262813C3357" pageId="12" pageNumber="63">
Subfamily
<taxonomicName authority="C. L. Koch, 1850" authorityName="C. L. Koch" authorityYear="1850" family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae C.L. Koch, 1850</taxonomicName>
<taxonomicName id="7D69FEDB1D7002D8C97AE4F7CA067B20" authority="C. L. Koch, 1850" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="subFamily" subFamily="Pholcinae">Pholcinae C.L. Koch, 1850</taxonomicName>
Figs 7, 8, 9, 10, 11, 12
</paragraph>
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<paragraph pageId="12" pageNumber="63">
<taxonomicName family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" rank="family">Pholcidae</taxonomicName>
<subSubSection id="4FC8C9E87066F577C738766F6BB2E85A" pageId="12" pageNumber="63" type="reference_group">
<paragraph id="FCE99A2FAE0F4A8288ADCD00A56CDD28" pageId="12" pageNumber="63">
<taxonomicName id="5FFEAA561978D198F5E0B3CCD8946763" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
CL Koch, 1850: 31. Type genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="18B324A8D63A86B6CB85978D8B282431" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
Walckenaer, 1805, by monotypy.
</paragraph>
<paragraph pageId="12" pageNumber="63">
<taxonomicName family="Pholcidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<paragraph id="A2FE682E4FE51C0BF6089E9CDB43B7F2" pageId="12" pageNumber="63">
<taxonomicName id="BE4999268C0E7DB761ABCB28814999C7" class="Arachnida" family="Pholcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
CL Koch;
<bibRefCitation pageId="12" pageNumber="63">Simon 1893</bibRefCitation>
<bibRefCitation id="510319E5B8C8BE1E0C84ED5187106D3A" pageId="12" pageNumber="63">Simon 1893</bibRefCitation>
: 461;
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
<bibRefCitation id="F8A9F816BF9E50D2F701B36EDAC368CF" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="211 - 222" title="Phylogeny and classification of Pholcidae (Araneae): an update." url="https://doi.org/10.1636/CA10-57.1" volume="39" year="2011 b">Huber 2011b</bibRefCitation>
: 218.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="63" type="remarks">
<paragraph pageId="12" pageNumber="63">Remarks.</paragraph>
<paragraph pageId="12" pageNumber="63">
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<subSubSection id="EF0B5CA42F3B30B83750EC7983016672" pageId="12" pageNumber="63" type="remarks">
<paragraph id="F8D449E1AF64E0FE4F6DE6B37973B57F" pageId="12" pageNumber="63">Remarks.</paragraph>
<paragraph id="F0C96E3CFB3DB5F0D32F8EAE5D868A53" pageId="12" pageNumber="63">
<taxonomicName id="A43C2FE42CD44245DF379B004812423D" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
resemble
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="56D0C0D5206B8C85162EB10BF996C34E" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
in several respects. Their highest diversity is in the humid tropics and subtropics, and a large variety of body forms reflect adaptations to different microhabitats. With currently 922 species in 26 genera,
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="3F7EFEAE43FAA6D0F64C063D899BE5FE" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
is also similar to
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="85BBA04DD6D9D0A7A07917958BD50724" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
in diversity. In contrast to
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="FC0AB00CAF0750350E5A19787874AE92" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
,
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="DF8E7F5B275EC587791A049175924F96" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
is largely restricted to the Old World, with the notable exception of the New World endemic genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
<taxonomicName id="24896F586AB5A2CAED3117AC1D9CCE73" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
Simon, 1893 and a few possibly relict species in
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="CFF00E6F9C0E49808B354E89C2BC4AD8" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
<taxonomicName id="DDF10AEC5BC14CBDF59B4DBEAF41AF5C" class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
(
<bibRefCitation author="Huber, BA" journalOrPublisher="Bonner zoologische Monographien" pageId="40" pageNumber="91" pagination="1 - 509" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." volume="58" year="2011 a">Huber 2011a</bibRefCitation>
<bibRefCitation id="357A7A9D29A158606B87438DE30D85F8" author="Huber, BA" journalOrPublisher="Bonner zoologische Monographien" pageId="40" pageNumber="91" pagination="1 - 509" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." volume="58" year="2011 a">Huber 2011a</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Invertebrate Systematics" pageId="42" pageNumber="93" pagination="432 - 450" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." url="https://doi.org/10.1071/IS13050" volume="28" year="2014">Huber et al. 2014</bibRefCitation>
<bibRefCitation id="EB9CC6E0C8C484463390F8B2A82888ED" author="Huber, BA" journalOrPublisher="Invertebrate Systematics" pageId="42" pageNumber="93" pagination="432 - 450" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." url="https://doi.org/10.1071/IS13050" volume="28" year="2014">Huber et al. 2014</bibRefCitation>
). While only a single species of
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="28C9F764242B3D4E10953DBDCDFA9183" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
has followed humans around the globe [
<taxonomicName class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus culicinus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="culicinus">Modisimus culicinus</taxonomicName>
<taxonomicName id="02F210D9A4AFA2ACE1A274788749AC26" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus culicinus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="culicinus">Modisimus culicinus</taxonomicName>
(Simon, 1893)] and one further species has spread widely in Europe and neighboring regions [
<taxonomicName class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus simoni" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="simoni">Psilochorus simoni</taxonomicName>
<taxonomicName id="A319F045E01121FA0EAD4F05394CB29D" class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus simoni" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="simoni">Psilochorus simoni</taxonomicName>
(Berland, 1911)], several synanthropic species in
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="8635A83634EE803DF8CE210B789D0659" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
have attained worldwide distributions or extended their ranges to another continent [most notably
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus phalangioides" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="phalangioides">Pholcus phalangioides</taxonomicName>
<taxonomicName id="8D32223594997E9FA3359EE15352386D" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus phalangioides" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="phalangioides">Pholcus phalangioides</taxonomicName>
(Fuesslin, 1775);
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora senoculata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="senoculata">Spermophora senoculata</taxonomicName>
<taxonomicName id="82FB38E40FA6DEC75657935A76E1C027" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora senoculata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="senoculata">Spermophora senoculata</taxonomicName>
(
<normalizedToken originalValue="Dugès">Duges</normalizedToken>
<normalizedToken id="2D945476F66B3A5BB23FA11B25C9E0B9" originalValue="Dugès">Duges</normalizedToken>
, 1836);
<taxonomicName class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus fauroti" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="fauroti">Micropholcus fauroti</taxonomicName>
<taxonomicName id="57B35F44DDC36ECCC9A924C14A2153F2" class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus fauroti" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="fauroti">Micropholcus fauroti</taxonomicName>
(Simon, 1887);
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus manueli" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="manueli">Pholcus manueli</taxonomicName>
<taxonomicName id="49175907F2B8E79066491AAEEF31B51A" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus manueli" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="manueli">Pholcus manueli</taxonomicName>
Gertsch, 1937].
</paragraph>
<paragraph pageId="12" pageNumber="63">
<paragraph id="B0CA8D6174F30E8E346B8621D0CC7575" pageId="12" pageNumber="63">
The sister-group relationship between
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="DFC45555BEC86983B2545F79ED283DA5" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
and
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
<taxonomicName id="BC0328C2BC29459940D238C19D9EF688" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Smeringopinae">Smeringopinae</taxonomicName>
is well established (see above). The same is true for the monophyly of
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="6DE06F259E7E5A53BDE52C0ABB23839C" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
. All our analyses support this subfamily (reasonable to high support), and morphological data have also supported this group (presence of male lateral proximal cheliceral apophyses,
<bibRefCitation author="Huber, BA" journalOrPublisher="Acta Zoologica, Stockholm" pageId="40" pageNumber="91" pagination="291 - 300" title="Copulatory mechanism in Holocnemuspluchei and Pholcusopilionoides, with notes on male cheliceral apophyses and stridulatory organs in Pholcidae (Araneae)." url="https://doi.org/10.1111/j.1463-6395.1995.tb01001.x" volume="76" year="1995">Huber 1995</bibRefCitation>
<bibRefCitation id="D1D6B22761AA0AA9B508601FC25E9468" author="Huber, BA" journalOrPublisher="Acta Zoologica, Stockholm" pageId="40" pageNumber="91" pagination="291 - 300" title="Copulatory mechanism in Holocnemuspluchei and Pholcusopilionoides, with notes on male cheliceral apophyses and stridulatory organs in Pholcidae (Araneae)." url="https://doi.org/10.1111/j.1463-6395.1995.tb01001.x" volume="76" year="1995">Huber 1995</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">2000</bibRefCitation>
<bibRefCitation id="38A5CA8D0024480260B476BB181C711D" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">2000</bibRefCitation>
; tarsus IV comb hairs in a single row,
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="232 - 240" title="Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae)." url="https://doi.org/10.1636/CSh07-71.1" volume="36" year="2008">Huber and Fleckenstein 2008</bibRefCitation>
<bibRefCitation id="FE6A1840BFA5EF79614A376221BA3B92" author="Huber, BA" journalOrPublisher="Journal of Arachnology" pageId="41" pageNumber="92" pagination="232 - 240" title="Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae)." url="https://doi.org/10.1636/CSh07-71.1" volume="36" year="2008">Huber and Fleckenstein 2008</bibRefCitation>
).
</paragraph>
<paragraph pageId="12" pageNumber="63">
<paragraph id="174EC25A57BAAA415DB42F008DC41490" pageId="12" pageNumber="63">
Even though
<taxonomicName lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
<taxonomicName id="CB322BC9DE21B6474B94202A32542AD7" lsidName="" pageId="12" pageNumber="63" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
are well represented in our analyses (317 of 597 species, i.e., 53%) internal relationships in this subfamily continue to be problematic. Several
<normalizedToken originalValue="basal">'basal'</normalizedToken>
<normalizedToken id="E4ECFC1BFC6837F6B689E764980AFDF6" originalValue="basal">'basal'</normalizedToken>
nodes are poorly supported (Figure 1); in part the topology is highly sensitive to different algorithms of analysis; and some details appear dubious from the perspective of morphology. However, many details are strongly supported by morphology, including some deep nodes (e.g., the
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="A4E0371E193CE4F2D5F79A1AEDE27408" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
group of genera); and some nodes, even though weakly supported or in conflict with morphology, provide reasonable and testable predictions for further research (e.g., the polyphyly of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
<taxonomicName id="2E4DB5B16B20026D33F5F07EE19B4F03" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
Hentz, 1841; the close relationship of certain Sri Lankan taxa with African rather than Southeast Asian taxa; the monophyly of African
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="2E805A29F0B5A47F4FC929A3DE2864AC" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
).
</paragraph>
<paragraph pageId="12" pageNumber="63">
<paragraph id="4BBC29790B49E2781CB3AB33E4871EE8" pageId="12" pageNumber="63">
The subfamily is here divided into three operational groups, more for the sake of convenience than as a reflection of the support values they receive. Actually, support is low for all of them, but much of this division is consistent among different analyses and may well reflect real major groups. 'Group 1' (Figs 7, 8) is entirely composed of small six-eyed taxa, and is roughly equivalent to what was originally subsumed under the name
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
<taxonomicName id="A6D20EF8AB91C7F4C4F3FB515B8DC2BA" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
. 'Group 2' (Figure 9 part) is also entirely composed of six-eyed taxa and is remarkable because it places the exclusively New World genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
<taxonomicName id="A2C27649D568EB783FC4FF03834D1BCE" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
close to African and Madagascan taxa. 'Group 3' (Figure 9 part, Figs 10-12) includes the fully supported
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="52BCFDBC81D22DAA565231C3C789DA0D" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
group of genera as proposed previously (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bonner zoologische Monographien" pageId="40" pageNumber="91" pagination="1 - 509" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." volume="58" year="2011 a">Huber 2011a</bibRefCitation>
<bibRefCitation id="C1EE6DF13C49F3E6C025796E13D5B7DF" author="Huber, BA" journalOrPublisher="Bonner zoologische Monographien" pageId="40" pageNumber="91" pagination="1 - 509" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." volume="58" year="2011 a">Huber 2011a</bibRefCitation>
) and its sister genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
<taxonomicName id="934A886C1C8AB62C3C0AD7FCE7484153" class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
Huber, 2003, a sister-group relationship that has also been proposed before (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="40" pageNumber="91" pagination="477 - 527" title="Southern African pholcid spiders: revision and cladistic analysis of Quamtana gen. nov. and Spermophora Hentz (Araneae: Pholcidae), with notes on male-female covariation." url="https://doi.org/10.1046/j.0024-4082.2003.00082.x" volume="139" year="2003 c">Huber 2003c</bibRefCitation>
<bibRefCitation id="8F3977E0A7E6281420FEC8A397A61E59" author="Huber, BA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="40" pageNumber="91" pagination="477 - 527" title="Southern African pholcid spiders: revision and cladistic analysis of Quamtana gen. nov. and Spermophora Hentz (Araneae: Pholcidae), with notes on male-female covariation." url="https://doi.org/10.1046/j.0024-4082.2003.00082.x" volume="139" year="2003 c">Huber 2003c</bibRefCitation>
). In the tree shown here (and in the RogueNaRok tree), the '
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
<taxonomicName id="BE1082580036FE946E730CEB0A01E13B" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
'
<taxonomicName genus="Modisimeae" lsidName="Modisimeae dieke" pageId="12" pageNumber="63" rank="species" species="dieke">dieke</taxonomicName>
<taxonomicName id="19631FD90813FA13DF1A6EF564595E5B" genus="Modisimeae" lsidName="Modisimeae dieke" pageId="12" pageNumber="63" rank="species" species="dieke">dieke</taxonomicName>
group has an isolated position outside of the three operational groups. In the other trees, it is part of 'group 1'.
</paragraph>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="1CC594EC941EF32C35F1E10BF785E431" pageId="12" pageNumber="63">
<paragraph id="C575B05124059BB649E73438195F7585" pageId="12" pageNumber="63">
Figure 7. Pholcinae 'group 1' (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
<taxonomicName id="6FA806DC833E5DEC975BFFA4F7E77BC7" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
and relatives) a Gen. n., sp. n.
<normalizedToken originalValue="“Ind206”">&quot;Ind206&quot;</normalizedToken>
<normalizedToken id="48A3F5C02C8BBE9F023255E82594DA25" originalValue="“Ind206”">&quot;Ind206&quot;</normalizedToken>
(Halmahera);
<normalizedToken originalValue="b">b'</normalizedToken>
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
<normalizedToken id="0733E653E155E904B17E3E903D34241E" originalValue="b">b'</normalizedToken>
<taxonomicName id="51F85E1BA6F2773032DDDB476F433850" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Spermophora</taxonomicName>
sp. n.
<normalizedToken originalValue="“Ind27”">&quot;Ind27&quot;</normalizedToken>
<normalizedToken id="1670F3E90690E0619DA26441325F7F3B" originalValue="“Ind27”">&quot;Ind27&quot;</normalizedToken>
(Sumatra) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Aetana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aetana baganihan" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="baganihan">Aetana baganihan</taxonomicName>
<taxonomicName id="2FA42DD6C574CBE3FDE979689A32AB9E" class="Arachnida" family="Pholcidae" genus="Aetana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aetana baganihan" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="baganihan">Aetana baganihan</taxonomicName>
(Philippines) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora senoculata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="senoculata">Spermophora senoculata</taxonomicName>
<taxonomicName id="FBCC71E8150E63BD270B452EAD47FBBF" class="Arachnida" family="Pholcidae" genus="Spermophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spermophora senoculata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="senoculata">Spermophora senoculata</taxonomicName>
(Turkey) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Savarna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Savarna tessellata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="tessellata">Savarna tessellata</taxonomicName>
<taxonomicName id="FE967C8B70B48425EA40D106FE4285EE" class="Arachnida" family="Pholcidae" genus="Savarna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Savarna tessellata" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="tessellata">Savarna tessellata</taxonomicName>
(Thailand) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wanniyala agrabopath" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="agrabopath">Wanniyala agrabopath</taxonomicName>
<taxonomicName id="70CFD5874C6D0D7D0DB3BF4E37F22C06" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wanniyala agrabopath" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="agrabopath">Wanniyala agrabopath</taxonomicName>
(Sri Lanka).
</paragraph>
</caption>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="F7918BC8D94DC403D751DCF124F6C5BC" pageId="12" pageNumber="63">
<paragraph id="3DC13E09BCF623E84AC6502A244966BE" pageId="12" pageNumber="63">
Figure 8.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
<taxonomicName id="70F728B56CC7199E294D9BABBCE1CAC9" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Hantu</taxonomicName>
<taxonomicName id="6046FC7E7D5868BEBC7CF1F9FE226F2C" class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Hantu</taxonomicName>
. For
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
<taxonomicName id="8DC99A12BE4DACE6522C90F16B482ADA" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
, the background colours signify microhabitat: red = ground; green = leaf. D, domed web; R, highly regular
<normalizedToken originalValue="curtain">'curtain'</normalizedToken>
<normalizedToken id="ACED52657E34D26816C3B0E5AE65C7D1" originalValue="curtain">'curtain'</normalizedToken>
web. Photos a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu niah" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="niah">Hantu niah</taxonomicName>
<taxonomicName id="7C308E226855ECE6E87ECA1C3AD85431" class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu niah" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="niah">Hantu niah</taxonomicName>
(Sarawak) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu kapit" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="kapit">Hantu kapit</taxonomicName>
<taxonomicName id="0835326E03143ED6758B17E04142C1F7" class="Arachnida" family="Pholcidae" genus="Hantu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hantu kapit" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="kapit">Hantu kapit</taxonomicName>
(Sarawak) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana sandakan" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sandakan">Belisana sandakan</taxonomicName>
<taxonomicName id="8410182907E87F8CD54480980437FDB4" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana sandakan" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sandakan">Belisana sandakan</taxonomicName>
(Sumatra) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana sabah" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sabah">Belisana sabah</taxonomicName>
<taxonomicName id="B6B2824DB1A652B70C7E08AB0BE167E5" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana sabah" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sabah">Belisana sabah</taxonomicName>
(Sabah) e domed web of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
<taxonomicName id="E4D02C987F113F8B0E514903ACA20B52" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Belisana</taxonomicName>
sp. n.
<normalizedToken originalValue="“Mal77”">&quot;Mal77&quot;</normalizedToken>
<normalizedToken id="CF2FC1D7A8274F7DEB02631ED9DBF2C0" originalValue="“Mal77”">&quot;Mal77&quot;</normalizedToken>
(Malaysia) f regular
<normalizedToken originalValue="curtain">'curtain'</normalizedToken>
<normalizedToken id="13F2F80CBEE781316BC3E271CFD48D17" originalValue="curtain">'curtain'</normalizedToken>
web of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana bohorok" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="bohorok">Belisana bohorok</taxonomicName>
<taxonomicName id="AFEB7B7480740EF2FCDDE632CACFB5CD" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana bohorok" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="bohorok">Belisana bohorok</taxonomicName>
(Sarawak).
</paragraph>
</caption>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="2F36F5373AD96DD09DC2A1A457B99953" pageId="12" pageNumber="63">
<paragraph id="378D9F460AE22EB86B2F6AC409DB9543" pageId="12" pageNumber="63">
Figure 9. Pholcinae 'group 2' (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Zatavua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zatavua" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Zatavua</taxonomicName>
<taxonomicName id="EAC81167DDEC23814D80177961ECD755" class="Arachnida" family="Pholcidae" genus="Zatavua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zatavua" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Zatavua</taxonomicName>
and relatives,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
<taxonomicName id="B2CC4960617E27E16CDAECB701D4C669" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
), and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
<taxonomicName id="B963B15A02234329A83D4C1AC606C430" class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
(marked: non-South African species). Photos a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia taruma" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="taruma">Metagonia taruma</taxonomicName>
<taxonomicName id="E930B33024BC5E18ADA814512498B384" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia taruma" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="taruma">Metagonia taruma</taxonomicName>
(Brazil) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
<taxonomicName id="2CACA248C36E80DB86EE3553AA3EC3EA" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Metagonia</taxonomicName>
sp. n.
<normalizedToken originalValue="“Br07-1”">&quot;Br07-1&quot;</normalizedToken>
<normalizedToken id="71DBAEA1DD198747FE363D60AB525870" originalValue="“Br07-1”">&quot;Br07-1&quot;</normalizedToken>
(Brazil) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia bifida" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="bifida">Metagonia bifida</taxonomicName>
<taxonomicName id="BEDF6707F69364916C298BFA890D99FB" class="Arachnida" family="Pholcidae" genus="Metagonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metagonia bifida" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="bifida">Metagonia bifida</taxonomicName>
? (Brazil) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
<taxonomicName id="D9A421F56069BC0512073D96B65C0B25" class="Arachnida" family="Pholcidae" genus="Quamtana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quamtana" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Quamtana</taxonomicName>
sp. n. (cf. mabusai) (Germany).
</paragraph>
</caption>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="A9F14C02A1605D55261FFD84E6E0A670" pageId="12" pageNumber="63">
<paragraph id="DA16F29A14FE26B12F3C6C1E41B4D094" pageId="12" pageNumber="63">
Figure 10.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Calapnita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calapnita" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Calapnita</taxonomicName>
<taxonomicName id="EE526D7F0B4B044DB70CC340A5227EEC" class="Arachnida" family="Pholcidae" genus="Calapnita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calapnita" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Calapnita</taxonomicName>
-
<taxonomicName class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panjange" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Panjange</taxonomicName>
<taxonomicName id="4221EEB2FBCDB9A3ADC8030CD55B8B66" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panjange" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Panjange</taxonomicName>
clade a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Kintaqa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kintaqa satun" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="satun">Kintaqa satun</taxonomicName>
<taxonomicName id="0D8A0259022C9860432C23ED269AA3B8" class="Arachnida" family="Pholcidae" genus="Kintaqa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kintaqa satun" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="satun">Kintaqa satun</taxonomicName>
(Malaysia) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Meraha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meraha narathiwat" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="narathiwat">Meraha narathiwat</taxonomicName>
<taxonomicName id="168562F1667F527FD11776F946BF3A0F" class="Arachnida" family="Pholcidae" genus="Meraha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meraha narathiwat" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="narathiwat">Meraha narathiwat</taxonomicName>
(Thailand) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Calapnita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calapnita vermiformis" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="vermiformis">Calapnita vermiformis</taxonomicName>
<taxonomicName id="0C572A8A915FF48C66DA95375D6B3E5F" class="Arachnida" family="Pholcidae" genus="Calapnita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calapnita vermiformis" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="vermiformis">Calapnita vermiformis</taxonomicName>
(Philippines) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Apokayana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apokayana kapit" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="kapit">Apokayana kapit</taxonomicName>
<taxonomicName id="CBE7EF188173CB5A30BDF39C6E897694" class="Arachnida" family="Pholcidae" genus="Apokayana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apokayana kapit" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="kapit">Apokayana kapit</taxonomicName>
(Sarawak) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Uthina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uthina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Uthina</taxonomicName>
<taxonomicName id="FB725B749C0BA6D16909E0EE19C0B54D" class="Arachnida" family="Pholcidae" genus="Uthina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uthina" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Uthina</taxonomicName>
sp. n.
<normalizedToken originalValue="“Ind121”">&quot;Ind121&quot;</normalizedToken>
<normalizedToken id="C61C9BB5A278F95143DB7907CE1F1A3F" originalValue="“Ind121”">&quot;Ind121&quot;</normalizedToken>
(Indonesia) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panjange casaroro" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="casaroro">Panjange casaroro</taxonomicName>
<taxonomicName id="C4A4D76A7C92EC9CF26B18A20C405EEC" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panjange casaroro" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="casaroro">Panjange casaroro</taxonomicName>
(Philippines).
</paragraph>
</caption>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="64CCF05C9EFC8A5F2478F1319254AA17" pageId="12" pageNumber="63">
<paragraph id="CDD643C6CC91D9167E0E91804C91AA5C" pageId="12" pageNumber="63">
Figure 11.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
<taxonomicName id="F0479537A795CF88AED3D7411D2F6C1C" class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
-
<taxonomicName class="Arachnida" family="Pholcidae" genus="Leptopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Leptopholcus</taxonomicName>
<taxonomicName id="4CCA6C2DABDD39AF6E841F3759B7C658" class="Arachnida" family="Pholcidae" genus="Leptopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Leptopholcus</taxonomicName>
clade a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
<taxonomicName id="9685E7D3076A21303C7CEA4DA7A61783" class="Arachnida" family="Pholcidae" genus="Micropholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micropholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
sp. n.
<normalizedToken originalValue="“Br15-152”">&quot;Br15-152&quot;</normalizedToken>
<normalizedToken id="66594A1E8D256A3CD5706B6893BCD025" originalValue="“Br15-152”">&quot;Br15-152&quot;</normalizedToken>
(Brazil) b
<taxonomicName genus="Canticus" lsidName="Canticus sepaku" pageId="12" pageNumber="63" rank="species" species="sepaku">Canticus sepaku</taxonomicName>
<taxonomicName id="E7DA2657119D9F3061814D483456C772" genus="Canticus" lsidName="Canticus sepaku" pageId="12" pageNumber="63" rank="species" species="sepaku">Canticus sepaku</taxonomicName>
(East Kalimantan) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Micromerys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromerys baiteta" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="baiteta">Micromerys baiteta</taxonomicName>
<taxonomicName id="F92AC316A16392F3A46B0AC33F8DEADD" class="Arachnida" family="Pholcidae" genus="Micromerys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromerys baiteta" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="baiteta">Micromerys baiteta</taxonomicName>
(West Papua) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Leptopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopholcus borneensis" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="borneensis">Leptopholcus borneensis</taxonomicName>
<taxonomicName id="576FCDC257580E29BA33AB4A65662417" class="Arachnida" family="Pholcidae" genus="Leptopholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopholcus borneensis" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="borneensis">Leptopholcus borneensis</taxonomicName>
(Singapore).
</paragraph>
</caption>
<caption pageId="12" pageNumber="63">
<paragraph pageId="12" pageNumber="63">
<caption id="7CDC7ADB21DA699B38C03944D6E5898C" pageId="12" pageNumber="63">
<paragraph id="771D80FEE9400846F96528BA13DAB5AC" pageId="12" pageNumber="63">
Figure 12.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
<taxonomicName id="A63A67748F8EB1A7485D4534EB4F901D" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pholcus" order="Araneae" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
a
<taxonomicName lsidName="P. creticus" pageId="12" pageNumber="63" rank="species" species="creticus">P. creticus</taxonomicName>
<taxonomicName id="12BCF509CF8A92B26B453ADAFF021783" lsidName="P. creticus" pageId="12" pageNumber="63" rank="species" species="creticus">P. creticus</taxonomicName>
(Crete) b
<taxonomicName lsidName="P. camba" pageId="12" pageNumber="63" rank="species" species="camba">P. camba</taxonomicName>
<taxonomicName id="8BC0CFDF7BA98D5303D5FCBD895F0F77" lsidName="P. camba" pageId="12" pageNumber="63" rank="species" species="camba">P. camba</taxonomicName>
(Sulawesi) c
<taxonomicName lsidName="P. mulu" pageId="12" pageNumber="63" rank="species" species="mulu">P. mulu</taxonomicName>
<taxonomicName id="45BAC57F9C369175720B5F0111A84E28" lsidName="P. mulu" pageId="12" pageNumber="63" rank="species" species="mulu">P. mulu</taxonomicName>
(Sarawak) d
<taxonomicName lsidName="P. baka" pageId="12" pageNumber="63" rank="species" species="baka">P. baka</taxonomicName>
<taxonomicName id="730562B8E9371ACDD8D9164450817C5F" lsidName="P. baka" pageId="12" pageNumber="63" rank="species" species="baka">P. baka</taxonomicName>
(Gabon) e
<taxonomicName genus="P." lsidName="P." pageId="12" pageNumber="63" rank="genus">P.</taxonomicName>
<taxonomicName id="6FA1FD91C8EA20E50DE0070B8FF9C9DA" genus="P." lsidName="P." pageId="12" pageNumber="63" rank="genus">P.</taxonomicName>
sp. n.
<normalizedToken originalValue="“SL43”">&quot;SL43&quot;</normalizedToken>
<normalizedToken id="8224BC4B577E98D463E364A58D4002F5" originalValue="“SL43”">&quot;SL43&quot;</normalizedToken>
(Sri Lanka).
</paragraph>
</caption>

View file

@ -1,518 +1,517 @@
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Figs 3, 4, 5
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<paragraph pageId="6" pageNumber="57">
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Simon, 1893: 484. Type genus
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Simon, 1893, by subsequent designation (
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).
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<paragraph id="966EE8F8F083B7B74D194B92EDC95590" pageId="6" pageNumber="57">
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Simon;
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: 216.
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<paragraph id="AC8304D8EBA75C89F0D49CFF372A31CD" pageId="6" pageNumber="57">Remarks.</paragraph>
<paragraph id="41423CAC8C0826F306A36AB8665A5D90" pageId="6" pageNumber="57">
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are the typical pholcids of the humid Neotropics, where they occupy a wide variety of microhabitats from leaf litter to high among the vegetation. This ecological variability is paralleled by a wide range of body forms, from tiny ground-dwelling forms (e.g.,
<bibRefCitation pageId="6" pageNumber="57">Gertsch 1982</bibRefCitation>
<bibRefCitation id="39DDB1128CB5242A5B4C3CE49BE8D6C3" pageId="6" pageNumber="57">Gertsch 1982</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Natural History" pageId="42" pageNumber="93" pagination="275 - 301" title="Diversity and endemism of pholcid spiders in Brazil's Atlantic Forest, with descriptions of four new species of the Atlantic Forest endemic genus Tupigea (Araneae: Pholcidae)." url="https://doi.org/10.1080/00222933.2010.524319" volume="45" year="2011">Huber and Rheims 2011</bibRefCitation>
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) to some of the largest pholcids with leg spans of over 15 cm (e.g.,
<bibRefCitation author="Huber, BA" journalOrPublisher="Invertebrate Systematics" pageId="41" pageNumber="92" pagination="281 - 300" title="Increased sampling blurs morphological and molecular species limits: revision of the Hispaniolan endemic spider genus Tainonia (Araneae: Pholcidae)." url="https://doi.org/10.1071/IS09017" volume="23" year="2009">Huber and Astrin 2009</bibRefCitation>
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,
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 60" title="Small scale endemism in Brazil's Atlantic Forest: 14 new species of Mesabolivar (Araneae, Pholcidae), each known from a single locality." url="https://doi.org/10.11646/zootaxa.3942.1.1" volume="3942" year="2015">Huber 2015</bibRefCitation>
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,
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">2018</bibRefCitation>
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). With currently 480 species in 24 genera,
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="AEAE279276DACEA75BB7879345C72815" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
is one of the two large subfamilies of
<taxonomicName family="Pholcidae" lsidName="" pageId="6" pageNumber="57" rank="family">Pholcidae</taxonomicName>
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, with several species-rich genera (e.g.,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Anopsicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anopsicus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Anopsicus</taxonomicName>
<taxonomicName id="789139EAB1DFFCE7FEA4FE1FEAF6736B" class="Arachnida" family="Pholcidae" genus="Anopsicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anopsicus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Anopsicus</taxonomicName>
Chamberlin &amp; Ivie, 1938;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Psilochorus</taxonomicName>
<taxonomicName id="F74FCD083D33044D5FC21B48B9006A4B" class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Psilochorus</taxonomicName>
Simon, 1893;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
<taxonomicName id="E05CC9D52358BCCAFF4E608FAC040205" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
Simon, 1893;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
<taxonomicName id="1971FB25C15AEDDDE7DB05EBFC387DB8" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<normalizedToken id="BEDD2D81636F8B4E2DE9C7DCFC8D1D30" originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 1998;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
<taxonomicName id="B676535CC4BB87F0A8331E2DCE7E5324" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<normalizedToken id="614D0A1D38EAA059182173512D88FA8D" originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 1998) and many undescribed species.
</paragraph>
<paragraph pageId="6" pageNumber="57">
<paragraph id="6289658ADB1F551413A1595F6F4978A8" pageId="6" pageNumber="57">
All previous analyses have supported this group (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="F4B2DA596FF38B4D167BB70BD6EAB17C" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">2001</bibRefCitation>
<bibRefCitation id="1607737C7EF48EBCEF39FC1C4A8C3838" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">2001</bibRefCitation>
,
<bibRefCitation author="Bruvo-Mađaric, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="39" pageNumber="90" pagination="661 - 673" title="Phylogeny of pholcid spiders (Araneae: Pholcidae): combined analysis using morphology and molecules." url="https://doi.org/10.1016/j.ympev.2005.08.016" volume="37" year="2005">
<bibRefCitation id="FD46AD6032E48E3AC6B4B1C3EB673DB8" author="Bruvo-Mađaric, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="39" pageNumber="90" pagination="661 - 673" title="Phylogeny of pholcid spiders (Araneae: Pholcidae): combined analysis using morphology and molecules." url="https://doi.org/10.1016/j.ympev.2005.08.016" volume="37" year="2005">
Bruvo
<normalizedToken originalValue="Mađarić">Mađaric</normalizedToken>
<normalizedToken id="6AD673F1B1EBE65CFFCBA4CD7572427E" originalValue="Mađarić">Mađaric</normalizedToken>
et al. 2005
</bibRefCitation>
,
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="999790E3611ED8720C248BEB20A00759" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
), even though with minor differences in composition. The equivalent 'New World
<normalizedToken originalValue="clade">clade'</normalizedToken>
<normalizedToken id="01CF32E1BC95A7B2002E69A019938A41" originalValue="clade">clade'</normalizedToken>
in
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">Huber (2001)</bibRefCitation>
<bibRefCitation id="2CDD0C9A987AD83E8CE2FD8341418CFA" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 144" title="The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships." url="https://doi.org/10.1206/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2" volume="260" year="2001">Huber (2001)</bibRefCitation>
still included the Australian
<taxonomicName class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
<taxonomicName id="56345B61EFA11F964EEAB099F831E41B" class="Arachnida" family="Pholcidae" genus="Wugigarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wugigarra" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Wugigarra</taxonomicName>
, a genus that has since been moved to
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="F238C7AD7949E01823BCCA914D25B715" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
(
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="3B6383A76E544E2EDFBCBE3AFC8B3A59" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
). As a result,
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="7841F7CF8B1E3683CC72FD37D14E9AEE" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
is now considered to be restricted to the New World.
</paragraph>
<paragraph pageId="6" pageNumber="57">
<paragraph id="896A570752672274BD47FE21936B6A21" pageId="6" pageNumber="57">
Our analyses all recover
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="68779F9D51D3DAB7B1488F2F77165E67" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
, but with very low support values. This is possibly due to the mysterious Andean genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="1D44921DA3FEF88A5F70D551834AEF9D" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
Simon, 1893 (Figure 3) that is either included in
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="1015D085755459286391D0816EE582E9" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
(IQ-TREE) or not (RAxML). The position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="1147C5139E4A8B17B9DF224BA664A180" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
has always been considered problematic.
<bibRefCitation pageId="6" pageNumber="57">Simon (1893)</bibRefCitation>
<bibRefCitation id="E5DE18CEFDC59881003F0D51FFE9F8F0" pageId="6" pageNumber="57">Simon (1893)</bibRefCitation>
created a separate taxon
<normalizedToken originalValue="“Prisculeae”">&quot;Prisculeae&quot;</normalizedToken>
<normalizedToken id="DCB0F86C5A620468A335399D76393F4F" originalValue="“Prisculeae”">&quot;Prisculeae&quot;</normalizedToken>
for this genus;
<bibRefCitation author="Brignoli, PM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="90" pagination="90 - 100" title="Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae)." volume="170" year="1981">Brignoli (1981)</bibRefCitation>
<bibRefCitation id="399FA96B93B469E03C6C093AF78E45B2" author="Brignoli, PM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="90" pagination="90 - 100" title="Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae)." volume="170" year="1981">Brignoli (1981)</bibRefCitation>
synonymized it with
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
<taxonomicName id="4C8B289A84C273F0528CB3A3D221E51E" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
; the first morphological cladistic analysis (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="4FB70F5B5F024A0413E58A1BC82E7F3B" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
) supported the position of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="64D02D5954F62637BFFAFFF4F9D12D67" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
near
<taxonomicName class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
<taxonomicName id="54150E98DF2507F1946B9C109043308B" class="Arachnida" family="Pholcidae" genus="Physocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Physocyclus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Physocyclus</taxonomicName>
but this result was explicitly doubted (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="B7D24F71F51AB042A47861626D8192BD" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
: 129). In the molecular analysis of
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. (2013)</bibRefCitation>
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<bibRefCitation id="02DE1BF98A4792226D56DB2031C53B4B" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. (2013)</bibRefCitation>
<taxonomicName id="A8338972AA2C63E8A281FF1CC8CC58E4" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
was excluded because the positions of the included species varied dramatically among different types of analyses. Morphologically,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="9B28FD775252AB60EED1562B3259FE7A" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
differs from (other)
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="8DF001477A4FC01FA86F4018A6B79009" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
by the presence of ALS piriform gland spigots and by the absence of a retrolateral apophysis on the male palpal coxa (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="BBA4422F3CD71ECAEAE6CADD9F3F8AA2" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
), i.e., it has retained plesiomorphic characters. A sister-group relationship between
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="05019221107AE5502C8564F67A095EAB" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
and other
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="D9F603685AA5E4716F02A3D21A6A7699" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
appears thus plausible from a morphological point of view.
</paragraph>
<caption pageId="6" pageNumber="57">
<paragraph pageId="6" pageNumber="57">
<caption id="3BE0C0759EC19E0DA2FC0FF3445BB83B" pageId="6" pageNumber="57">
<paragraph id="FA3C1BBCA892CF69C405839CB38D4C40" pageId="6" pageNumber="57">
Figure 3.
<normalizedToken originalValue="Basal">'Basal'</normalizedToken>
<normalizedToken id="A8D527130B3112DF6C067F8DB992A841" originalValue="Basal">'Basal'</normalizedToken>
Modisiminae a Gen. n., sp. n.
<normalizedToken originalValue="“Br16-50”">&quot;Br16-50&quot;</normalizedToken>
<normalizedToken id="9B185D10FAA89844001FFB5CD8B5DC66" originalValue="“Br16-50”">&quot;Br16-50&quot;</normalizedToken>
(Brazil) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula andinensis" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="andinensis">Priscula andinensis</taxonomicName>
<taxonomicName id="767B078C7CB35EB05F0D2E6662DBA58D" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula andinensis" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="andinensis">Priscula andinensis</taxonomicName>
? (Venezuela) c Gen. n., sp. n.
<normalizedToken originalValue="“Br16-196”">&quot;Br16-196&quot;</normalizedToken>
<normalizedToken id="288B76E738F2467C5E083DDCDBD2F82A" originalValue="“Br16-196”">&quot;Br16-196&quot;</normalizedToken>
(Brazil) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Tupigea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tupigea" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Tupigea</taxonomicName>
<taxonomicName id="937A3C38C3FD098D5F1D9D378BC10D9E" class="Arachnida" family="Pholcidae" genus="Tupigea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tupigea" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Tupigea</taxonomicName>
sp. n.
<normalizedToken originalValue="“Br14-47”">&quot;Br14-47&quot;</normalizedToken>
<normalizedToken id="ED53C610D2E380B1C730DBF7EEFDDCC6" originalValue="“Br14-47”">&quot;Br14-47&quot;</normalizedToken>
(Brazil) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa silvae" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="silvae">Pisaboa silvae</taxonomicName>
<taxonomicName id="270DBB83DACB74A30B232CF1FD0C488E" class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa silvae" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="silvae">Pisaboa silvae</taxonomicName>
(Brazil) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus imitatus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="imitatus">Psilochorus imitatus</taxonomicName>
<taxonomicName id="EDC4A16800433D9AB2FC24060CCD86C1" class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus imitatus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="imitatus">Psilochorus imitatus</taxonomicName>
(USA) g
<taxonomicName class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus incertus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="incertus">Modisimus incertus</taxonomicName>
<taxonomicName id="8E231C5B85AB5CAF52B13996E4A5E856" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus incertus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="incertus">Modisimus incertus</taxonomicName>
(Cuba).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="57">
<paragraph id="51AEAFA2713132AF0AC4800B56FD9BB3" pageId="6" pageNumber="57">
Despite the low support values, we thus consider
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="AA1AC5E7834EF1BEA86AE8E487A3036B" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
(including
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="AA7D8FB567ED4B340A0644576A6EEC2B" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
or not) a likely monophyletic group. Several morphological characters support
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="BD692B1541C71699DB3CC7882DDA55EB" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
(incl.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="CE6F4FB44D266710CEA0DB2049957E33" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
): an exposed tarsal organ; the reduction of epiandrous spigots (shared with 'other
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="D10DCBCBE9BF7C504A2075FF404B833A" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
'; see above); and a large distance between ALE and PME (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="473916B6BFDAA4E574CCA919800F935E" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
). As indicated above (section
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="section" section="Arteminae" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="27729E798C59B0DDC7747888E3379FB2" lsidName="" pageId="6" pageNumber="57" rank="section" section="Arteminae" subFamily="Arteminae">Arteminae</taxonomicName>
) our data weakly support a sister-group relationship between 'other
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="FEF734AF481C9FE2EB417D8F88EAF6C7" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' and
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="F2FD73AFC432776564EEA84859E0CC41" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
.
</paragraph>
<paragraph pageId="6" pageNumber="57">
<paragraph id="1654D780814654842CCDBA6C3F067818" pageId="6" pageNumber="57">
Within
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="F790BF860E8BD2A834A0470F3D5B1E83" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
, many support values are extremely low, and the suggested relationships are thus unreliable (Figure 3). In addition, taxon sampling is very uneven, with some genera well represented (e.g.,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<taxonomicName id="F74D1F0CBC3B0E7C6843D5BB6F824484" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="FA80E136707B1EC03794B38D6BC437CA" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
<taxonomicName id="638A0A36643A9C1057B257D39F996D22" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
), and others poorly represented or entirely missing (see below). However, several results are consistent among analyses and noteworthy for various reasons: they suggest groups that appear feasible in terms of biogeography; they suggest interesting evolutionary scenarios; and they suggest formal taxonomic changes, some of which have been suggested before based on morphology.
</paragraph>
<paragraph lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
<paragraph id="2B90D031564B73A8013326BDFB33994F" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
Apart from
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="2B1C1C2AFE781043ED4C15FBF0EBFDF9" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
, the
<normalizedToken originalValue="basal">'basal'</normalizedToken>
<normalizedToken id="333614FA5F231C85C5D0519001269579" originalValue="basal">'basal'</normalizedToken>
branches within
<taxonomicName lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="CF36551844B80370B7153B18C46AADB4" lsidName="" pageId="6" pageNumber="57" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
lead to small South American unnamed taxa (Figure 3). In particular, the two species
<normalizedToken originalValue="“Br16-44”">&quot;Br16-44&quot;</normalizedToken>
<normalizedToken id="3F922D2C3E66BA0A7FBFA8CF68DBCD0A" originalValue="“Br16-44”">&quot;Br16-44&quot;</normalizedToken>
and
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<pageBreakToken id="CC157EF93F3152BAC15AD936B7AA873C" pageId="7" pageNumber="58" start="start">&quot;</pageBreakToken>
MACN270&quot; are both tiny, with body lengths of 0.9 and 1.3 mm, respectively. Other
<normalizedToken originalValue="basal">'basal'</normalizedToken>
<normalizedToken id="E8548B54125F3268F4CC4D0D7E18C15A" originalValue="basal">'basal'</normalizedToken>
branches lead to an unnamed Amazonian genus (
<normalizedToken originalValue="“Br16-178”">&quot;Br16-178&quot;</normalizedToken>
<normalizedToken id="4D45C19EDBB5A4F5CC903CE52570037B" originalValue="“Br16-178”">&quot;Br16-178&quot;</normalizedToken>
and
<normalizedToken originalValue="“Br16-50”">&quot;Br16-50&quot;</normalizedToken>
<normalizedToken id="420739B4117BAEC4D7CDDEEE3AE6A9C9" originalValue="“Br16-50”">&quot;Br16-50&quot;</normalizedToken>
; body lengths: 1.5-1.8 mm) and the Atlantic Forest genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Tupigea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tupigea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Tupigea</taxonomicName>
<taxonomicName id="4C2CF563AF7434BDB52D6B5C23DF6F0E" class="Arachnida" family="Pholcidae" genus="Tupigea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tupigea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Tupigea</taxonomicName>
Huber, 2000 (body lengths: 1.3-1.9 mm;
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="883BCEDAA998D464F1ACFF38752E934A" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Natural History" pageId="42" pageNumber="93" pagination="275 - 301" title="Diversity and endemism of pholcid spiders in Brazil's Atlantic Forest, with descriptions of four new species of the Atlantic Forest endemic genus Tupigea (Araneae: Pholcidae)." url="https://doi.org/10.1080/00222933.2010.524319" volume="45" year="2011">Huber and Rheims 2011</bibRefCitation>
<bibRefCitation id="398C85C1BBFC26D9B5740DC0676DD20F" author="Huber, BA" journalOrPublisher="Journal of Natural History" pageId="42" pageNumber="93" pagination="275 - 301" title="Diversity and endemism of pholcid spiders in Brazil's Atlantic Forest, with descriptions of four new species of the Atlantic Forest endemic genus Tupigea (Araneae: Pholcidae)." url="https://doi.org/10.1080/00222933.2010.524319" volume="45" year="2011">Huber and Rheims 2011</bibRefCitation>
). This suggests a similar evolutionary scenario as proposed for 'other
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="01AF82616F48E9E7EED8A8BEDC64C966" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
' above, i.e., that ancestral
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="A3B5D8764F214D0E7B8C324F22AD8AF8" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
may have been small ground-dwelling species.
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="E20BB1C3A5CD7DD60E9E8A25BACE81E2" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
is once again the disturbing factor in this scenario: all known representatives of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="0F89E9B6D326D6B0A4A49E2C719607D1" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
are medium-size to large spiders (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="C1A78E1B0C6763B64AC72CDF9B44EAB0" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
), possibly surpassed (as far as body mass is concerned) by
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="805EA43961DC8AA8FF434180C4FD7359" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
only. In both
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="E87CD5229CD9107D3729DC9C1642D73B" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
and
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="DC7BBEB7B12F90DBD3D01BCFFF0B28FE" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
, the emerging picture is one of medium-sized forms missing or disappearing early, large forms experiencing little subsequent changes in body shape and poor subsequent speciation (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
<taxonomicName id="DD9E3F18E39BE1CD59F4E46226F54BE5" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artema" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Artema</taxonomicName>
: currently eight species;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="9897E4DCD6D8B56EB44937E1DC48C97B" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
: currently 17 species), and small forms diversifying dramatically in size, shape, and numbers ('other
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
<taxonomicName id="4093AA244D46ACA725CD7874A0E6C18B" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Arteminae">Arteminae</taxonomicName>
': currently 91 species;
<taxonomicName lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
<taxonomicName id="374344B807924EDF2906E23674315F1E" lsidName="" pageId="7" pageNumber="58" rank="subFamily" subFamily="Modisiminae">Modisiminae</taxonomicName>
without
<taxonomicName class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
<taxonomicName id="81E58B23406566C2BC57B99659F9526E" class="Arachnida" family="Pholcidae" genus="Priscula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Priscula" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Priscula</taxonomicName>
: currently 463 species).
</paragraph>
<paragraph pageId="7" pageNumber="58">
<paragraph id="8B4F6ECB450943EF9B45AEA89E61CFE5" pageId="7" pageNumber="58">
The next branch (Figure 3;
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
<taxonomicName id="EF29CC168777E8250E46344392CD6845" class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
Huber, 2000 to
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pisaboa</taxonomicName>
<taxonomicName id="5D2471AAC99AC0F86E5D7D6698C8EDC9" class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pisaboa</taxonomicName>
Huber, 2000) includes several South American genera, some of them diverse but poorly represented in our analyses (e.g.,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
<taxonomicName id="732812BED35D4119B65505359CD26FB2" class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
). The close relationship between
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pisaboa</taxonomicName>
<taxonomicName id="365E66739F3178587F7FCF2C5EBFA82E" class="Arachnida" family="Pholcidae" genus="Pisaboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisaboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pisaboa</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Waunana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Waunana" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Waunana</taxonomicName>
<taxonomicName id="1B95E01AB8073D217F00E1676334C6E0" class="Arachnida" family="Pholcidae" genus="Waunana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Waunana" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Waunana</taxonomicName>
Huber, 2000 was already suggested in the original descriptions of these genera (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="A0C80CCEA17B89B70694FB7877137B59" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
), even though based on highly homoplastic characters (vertical hairs on male leg tibiae in high density; shape of apophysis on male palpal femur). A close relationship of these two genera with
<taxonomicName class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
<taxonomicName id="C1CAC837CB1C78AC3F4AD19B2E6CA04E" class="Arachnida" family="Pholcidae" genus="Chibchea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chibchea" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Chibchea</taxonomicName>
either receives very low support (IQ-TREE, RAxML) or is not recovered (RogueNaRok); it is neither supported nor contradicted by morphology. Clearly, this clade needs a much denser sampling and the addition of missing taxa that are possibly related (e.g.,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Pomboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pomboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pomboa</taxonomicName>
<taxonomicName id="2A343ECC85AB429375EC5271E5402EEA" class="Arachnida" family="Pholcidae" genus="Pomboa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pomboa" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Pomboa</taxonomicName>
).
</paragraph>
<paragraph lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<paragraph id="2828F2C5070602E9C9D379B2FF9578C5" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
The next clade (Figure 3) includes all North and Central American and Caribbean taxa, suggesting that the ancestor of this clade arrived in the region from South America. This scenario was explicitly rejected by
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. (2013)</bibRefCitation>
<bibRefCitation id="6A42C4B9AF013052A8A99E6BBF49DE06" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. (2013)</bibRefCitation>
based on the supposed age of the group (~120-170 Ma). However, our upcoming analysis has not been able to confirm this age (
<bibRefCitation author="Eberle, J" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" url="https://doi.org/10.1186/s12862-018-1244-8" year="2018">Eberle et al. 2018</bibRefCitation>
<bibRefCitation id="F59746CE4B175CBDEFB9E03D95B262E2" author="Eberle, J" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" url="https://doi.org/10.1186/s12862-018-1244-8" year="2018">Eberle et al. 2018</bibRefCitation>
; we were not able to calculate convincing absolute ages from the data). The clade is recovered in most analyses (it is paraphyletic in the 4+ genes tree) but always with low support (only SH values are reasonable to high). The only geographic outlier in this clade is South American '
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'. North American (
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)
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and South American '
<taxonomicName class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Psilochorus</taxonomicName>
<taxonomicName id="8E02E1F9C42D67004F1C92B383CB3449" class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Psilochorus</taxonomicName>
' each receive high to full support but are never resolved as sister taxa. Whether South American '
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<taxonomicName id="A7660797A737CBD9BDD7D82983364D20" class="Arachnida" family="Pholcidae" genus="Psilochorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psilochorus" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Psilochorus</taxonomicName>
' are ancestral within this large clade or represent a case of back-colonization is currently impossible to say; the internal nodes in this clade have partly too low support to favour a particular scenario. The inclusion of the Central American
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<taxonomicName id="55F95BB42557FA15A8CABF36DD5850E5" class="Arachnida" family="Pholcidae" genus="Ixchela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ixchela" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Ixchela</taxonomicName>
Huber, 2000 in this clade fits the geographic pattern and contradicts a previous speculation (in
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
<bibRefCitation id="48A01096C0876ABC879919FD13D83EF7" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
) that
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<taxonomicName id="92A84E3D3C36155667E2DBE26D11B1DB" class="Arachnida" family="Pholcidae" genus="Ixchela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ixchela" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Ixchela</taxonomicName>
might be close to the South American genus
<taxonomicName class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
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Huber, 2000. In much the same way, the only Central American representative of
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<taxonomicName id="D9983FD071467D1484FCD24A2A97B5F7" class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
Simon, 1893 included in our analyses is placed in this group, far away from
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South American
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(the polyphyly of
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has long been suspected:
<bibRefCitation author="Gertsch, WJ" journalOrPublisher="Association for Mexican Cave Studies, Bulletin" pageId="39" pageNumber="90" pagination="47 - 111" title="A report on some Mexican cave spiders." volume="4" year="1971">Gertsch 1971</bibRefCitation>
<bibRefCitation id="7D6CB0A29293F775D6BC2E8723E48075" author="Gertsch, WJ" journalOrPublisher="Association for Mexican Cave Studies, Bulletin" pageId="39" pageNumber="90" pagination="47 - 111" title="A report on some Mexican cave spiders." volume="4" year="1971">Gertsch 1971</bibRefCitation>
,
<bibRefCitation author="Brignoli, PM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="90" pagination="90 - 100" title="Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae)." volume="170" year="1981">Brignoli 1981</bibRefCitation>
<bibRefCitation id="2AA8C3001B996B10D666021764BCA802" author="Brignoli, PM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="90" pagination="90 - 100" title="Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae)." volume="170" year="1981">Brignoli 1981</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Journal of Zoology, London" pageId="40" pageNumber="91" pagination="587 - 599" title="Genital mechanics in some neotropical pholcid spiders (Araneae: Pholcidae), with implications for systematics." url="https://doi.org/10.1111/j.1469-7998.1998.tb00063.x" volume="244" year="1998">Huber 1998</bibRefCitation>
<bibRefCitation id="0C11934E5F545C8A3E7AB4F421E245F8" author="Huber, BA" journalOrPublisher="Journal of Zoology, London" pageId="40" pageNumber="91" pagination="587 - 599" title="Genital mechanics in some neotropical pholcid spiders (Araneae: Pholcidae), with implications for systematics." url="https://doi.org/10.1111/j.1469-7998.1998.tb00063.x" volume="244" year="1998">Huber 1998</bibRefCitation>
,
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">2000</bibRefCitation>
<bibRefCitation id="E11B4A8A03CCB8FBB6ADDCE989ED32F6" author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">2000</bibRefCitation>
). The Cuban endemic
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<normalizedToken originalValue="Özdikmen">Oezdikmen</normalizedToken>
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&amp; Demir, 2009 is deeply nested within the large genus
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. It is resolved as sister to a distinctive group of Hispaniolan leaf-dwelling representatives of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
<taxonomicName id="8DF373379A6D81A7183C90DCE42981C2" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
(the &quot;leaf-dwelling species group&quot; in
<bibRefCitation author="Huber, BA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="42" pageNumber="93" pagination="244 - 299" title="High level of endemism in Haiti's last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules." url="https://doi.org/10.1111/j.1096-3642.2009.00559.x" volume="158" year="2010">Huber et al. 2010</bibRefCitation>
<bibRefCitation id="773FD265922F2A628BF09E994ABE0B06" author="Huber, BA" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="42" pageNumber="93" pagination="244 - 299" title="High level of endemism in Haiti's last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules." url="https://doi.org/10.1111/j.1096-3642.2009.00559.x" volume="158" year="2010">Huber et al. 2010</bibRefCitation>
) and synonymized below. Finally, the large genus
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<taxonomicName id="F1A0F3E9DD88D3A6D6CEE4E67C6B95B4" class="Arachnida" family="Pholcidae" genus="Anopsicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anopsicus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Anopsicus</taxonomicName>
(63 described species) is poorly represented in our analyses. The three species included are all undescribed, do not group together, and are nested among
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<taxonomicName id="053943A210EFCB926B983F58FC6E425A" class="Arachnida" family="Pholcidae" genus="Modisimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Modisimus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Modisimus</taxonomicName>
. Since neither the type species of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Anopsicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anopsicus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Anopsicus</taxonomicName>
<taxonomicName id="11A54072A045EF1609A99D2FFF0BA8E4" class="Arachnida" family="Pholcidae" genus="Anopsicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anopsicus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Anopsicus</taxonomicName>
is included nor is a potential close relative (or at least another species from
<normalizedToken originalValue="Yucatán">Yucatan</normalizedToken>
<normalizedToken id="6CAD42155AC2036CF47D5AD317BA6067" originalValue="Yucatán">Yucatan</normalizedToken>
), the monophyly and position of
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both remain dubious.
</paragraph>
<paragraph pageId="8" pageNumber="59">
<paragraph id="B0E115CF8CE3CDECE315F523F4B78C37" pageId="8" pageNumber="59">
Sister to the previous North and Central American and Caribbean clade is another large, entirely South American clade (Figure 3, bottom). The sister-group relationship is very poorly supported, but the monophyly of the South American clade has modest (4+ genes) to reasonable (RogueNaRok) support. It is divided into three subclades with reasonable to full support plus the genus
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<taxonomicName id="9E957B10F5A91672D8B195B96FB86587" class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
that is represented by a single species and whose position within this clade is not convincingly resolved. The first subclade included is here informally called the '
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<normalizedToken originalValue="clade">clade'</normalizedToken>
<taxonomicName id="BA66618DF5AF5A5A55FD474029006DBE" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<normalizedToken id="7243F9454608479FDCAEFC2395FC5D34" originalValue="clade">clade'</normalizedToken>
(Figure 4); the second subclade is largely Venezuelan and thus called 'Venezuelan
<normalizedToken originalValue="clade">clade'</normalizedToken>
<normalizedToken id="A0EA136E1BA157CC694CDFC26946BB94" originalValue="clade">clade'</normalizedToken>
(Figure 5); the third subclade is the genus
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<taxonomicName id="FFF1C668537F0201C108EDD0CB31FF82" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
(Figure 5).
</paragraph>
<caption pageId="8" pageNumber="59">
<paragraph pageId="8" pageNumber="59">
<caption id="1B33071463EE1934F8B19BEB5FEC31E4" pageId="8" pageNumber="59">
<paragraph id="4592AB0AB579D5A6DE95A85C07C60D73" pageId="8" pageNumber="59">
Figure 4.
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<taxonomicName id="BCF7243E483E3E2A8656A33903128AB4" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
clade a
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<taxonomicName id="3A659568BC45D4BDE498816B8C2C9EFA" class="Arachnida" family="Pholcidae" genus="Otavaloa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otavaloa lisei" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="lisei">Otavaloa lisei</taxonomicName>
(Brazil) b
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<taxonomicName id="8919B750908038A6D60E75F839ED97D3" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar maraba" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="maraba">Mesabolivar maraba</taxonomicName>
(Brazil) c
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<taxonomicName id="4FDF2A38F150FCEF8B06DC4B872CAD58" class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
sp. n.
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<normalizedToken id="D6248795357631466CB238447DED0A8B" originalValue="“Br16-153”">&quot;Br16-153&quot;</normalizedToken>
(Brazil) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar cyaneotaeniatus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="cyaneotaeniatus">Mesabolivar cyaneotaeniatus</taxonomicName>
<taxonomicName id="0E7B654D9619CA22551B69053E24BB0A" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar cyaneotaeniatus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="cyaneotaeniatus">Mesabolivar cyaneotaeniatus</taxonomicName>
(Brazil) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar kathrinae" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="kathrinae">Mesabolivar kathrinae</taxonomicName>
<taxonomicName id="7F8846C8B5E3804EAFFF069ED288470E" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar kathrinae" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="kathrinae">Mesabolivar kathrinae</taxonomicName>
(Brazil) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar saci" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="saci">Mesabolivar saci</taxonomicName>
<taxonomicName id="49A93C39CE3641D89E3E2C1C0575BA95" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar saci" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="saci">Mesabolivar saci</taxonomicName>
(Brazil).
</paragraph>
</caption>
<caption pageId="8" pageNumber="59">
<paragraph pageId="8" pageNumber="59">
<caption id="881E26134E4F2D010063FC52C79E8DE6" pageId="8" pageNumber="59">
<paragraph id="FC5185BB722A29ED30B7688013D3758A" pageId="8" pageNumber="59">
Figure 5. Venezuelan clade +
<taxonomicName class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
<taxonomicName id="C43B79C9F5129C62A766EBDEE37D9F9D" class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
+
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<taxonomicName id="07EE22A1EFE2F73D5DD9A0C40405DEF3" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
a
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus yawaperi" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="yawaperi">Mecolaesthus yawaperi</taxonomicName>
<taxonomicName id="F9045AC8BD106A5B9EA3ABCC14271A79" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus yawaperi" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="yawaperi">Mecolaesthus yawaperi</taxonomicName>
(Brazil) b
<taxonomicName class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
<taxonomicName id="8E1C176CE9B4BB6E6406F83DFB6C0597" class="Arachnida" family="Pholcidae" genus="Aymaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aymaria" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Aymaria</taxonomicName>
sp. n.
<normalizedToken originalValue="“Br16-188”">&quot;Br16-188&quot;</normalizedToken>
<normalizedToken id="FC96E454B465BDB2D2230CCD61CF0FD1" originalValue="“Br16-188”">&quot;Br16-188&quot;</normalizedToken>
(Brazil) c
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia rubra" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="rubra">Carapoia rubra</taxonomicName>
<taxonomicName id="D1FE2D74B4226944752940439DCC0FF1" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia rubra" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="rubra">Carapoia rubra</taxonomicName>
(Brazil) d
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia kaxinawa" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="kaxinawa">Carapoia kaxinawa</taxonomicName>
<taxonomicName id="6B603ECEF9B733EA6BA6460BB4FB6F85" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia kaxinawa" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="kaxinawa">Carapoia kaxinawa</taxonomicName>
(Brazil) e
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia pulchra" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="pulchra">Carapoia pulchra</taxonomicName>
<taxonomicName id="A4D6F00B25CEF39CBAB3E2F7951B564C" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia pulchra" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="pulchra">Carapoia pulchra</taxonomicName>
(Brazil) f
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia agilis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="agilis">Carapoia agilis</taxonomicName>
<taxonomicName id="B6A236AF0B56A1CA0C6351C66519EF10" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia agilis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="agilis">Carapoia agilis</taxonomicName>
(Brazil).
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="59">
<paragraph id="54D4024997B1692DB013C3FD37A34DA9" pageId="8" pageNumber="59">
Within the '
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<normalizedToken originalValue="clade">clade'</normalizedToken>
<taxonomicName id="35998BE8FE63561170BAEC641BF6A6CB" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<normalizedToken id="13AFBA768EAE85E7ED04F1ECA7034C42" originalValue="clade">clade'</normalizedToken>
(Figure 4), our analyses suggest two specific relationships that are likely to have drastic taxonomic consequences. First,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
<taxonomicName id="72D4FD12BB37E26CA7CA73FB4C53D700" class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
Simon, 1893 is nested among
<normalizedToken originalValue="true">'true'</normalizedToken>
<normalizedToken id="1862E8DE4C9F668B642E026F34C2EFE2" originalValue="true">'true'</normalizedToken>
northern South American
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="69BDB1189ED45C45425B5E1F868D1363" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
. This has been suggested before (
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
<bibRefCitation id="2654CF9F4786D67D0A85A53C906E61AF" author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
), but that previous analysis included a single species of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
<taxonomicName id="BC0ADC55F868A044EF3C087B4EFE5B6D" class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
whose generic identity was uncertain (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zoologischer Anzeiger" pageId="42" pageNumber="93" pagination="511 - 521" title="Litoporusiguassuensis Mello-Leitao, 1918 (Araneae, Pholcidae): camouflaged retreat, sexual dimorphism, female color polymorphism, intra-specific genital variation, and description of the male." url="https://doi.org/10.1016/j.jcz.2012.12.001" volume="252" year="2013">Huber et al. 2013</bibRefCitation>
<bibRefCitation id="B4CBE422C41F2C47B7ECA0C63D00979E" author="Huber, BA" journalOrPublisher="Zoologischer Anzeiger" pageId="42" pageNumber="93" pagination="511 - 521" title="Litoporusiguassuensis Mello-Leitao, 1918 (Araneae, Pholcidae): camouflaged retreat, sexual dimorphism, female color polymorphism, intra-specific genital variation, and description of the male." url="https://doi.org/10.1016/j.jcz.2012.12.001" volume="252" year="2013">Huber et al. 2013</bibRefCitation>
). The present analyses include several unambiguous (Amazonian) representatives of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
<taxonomicName id="414318A92D6CFF1738B896463D1192BF" class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
. Our data support the monophyly of
<taxonomicName class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
<taxonomicName id="F67346A0954897F4A37E5940A1F3DDC0" class="Arachnida" family="Pholcidae" genus="Litoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoporus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Litoporus</taxonomicName>
(full support) but also its position within
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="BC121B37F1DDDA99F0EDD305AB8A7A16" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
(reasonable to high support). Second,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="B43511D85E831BFC5E66995048A81721" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
is composed of two sub-clades:
<normalizedToken originalValue="true">'true'</normalizedToken>
<normalizedToken id="7E225E28F4045695C40BEDAA286F3548" originalValue="true">'true'</normalizedToken>
northern South American
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="58113663495F2B45BF367C600BDF8AE7" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
, and southern South American (largely Atlantic Forest) '
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="94060945060D1D8A4FE21450A9689B15" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
'. The southern sub-clade includes the monotypic genus
<taxonomicName family="Pholcidae" genus="Teuia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Teuia</taxonomicName>
<taxonomicName id="B0AE7897FD4A7E86F66E0374B212CAF2" class="Arachnida" family="Pholcidae" genus="Teuia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Teuia</taxonomicName>
Huber, 2000 (synonymized with
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="88DD6E32F9A166546FA86DB20B2AAE5D" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
in
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber 2018</bibRefCitation>
<bibRefCitation id="8CB772CF110DF42AE97424DBA36241C4" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber 2018</bibRefCitation>
; the type species of
<taxonomicName family="Pholcidae" genus="Teuia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Teuia</taxonomicName>
<taxonomicName id="3E2FC81824A9015612728B36262EDC09" class="Arachnida" family="Pholcidae" genus="Teuia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Teuia</taxonomicName>
is not included but a putatively closely related species:
<taxonomicName lsidName="M. sepitus" pageId="8" pageNumber="59" rank="species" species="sepitus">M. sepitus</taxonomicName>
<taxonomicName id="E4A32A2DFEED8294CE2348987674C264" lsidName="M. sepitus" pageId="8" pageNumber="59" rank="species" species="sepitus">M. sepitus</taxonomicName>
). Potential formal taxonomic changes are discussed in the Taxonomy section below. The close relationship between
<taxonomicName class="Arachnida" family="Pholcidae" genus="Otavaloa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otavaloa" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Otavaloa</taxonomicName>
<taxonomicName id="6AB32DEE989441C05968EA2E4A1DDC42" class="Arachnida" family="Pholcidae" genus="Otavaloa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otavaloa" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Otavaloa</taxonomicName>
Huber, 2000 and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="6AA6138F27AB0D4C661345E001F75677" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
is neither supported nor contradicted by morphological data.
</paragraph>
<paragraph lastPageId="9" lastPageNumber="60" pageId="8" pageNumber="59">
<paragraph id="FE9C0D4A8BEBEAE5DF11C6AC9986F481" lastPageId="9" lastPageNumber="60" pageId="8" pageNumber="59">
The 'Venezuelan
<normalizedToken originalValue="clade">clade'</normalizedToken>
<normalizedToken id="718E1E52AAD5F537553477BED73313A1" originalValue="clade">clade'</normalizedToken>
(Figure 5) receives high to full support and is composed of several genera that are either known from Venezuela only (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Systenita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenita" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Systenita</taxonomicName>
<taxonomicName id="0C8ED9D443116739F1AEC65D6E18D84E" class="Arachnida" family="Pholcidae" genus="Systenita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenita" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Systenita</taxonomicName>
Simon, 1893,
<taxonomicName class="Arachnida" family="Pholcidae" genus="Stenosfemuraia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenosfemuraia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Stenosfemuraia</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
<taxonomicName id="D196B84104344D535FA26C7E747C7728" class="Arachnida" family="Pholcidae" genus="Stenosfemuraia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenosfemuraia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Stenosfemuraia</taxonomicName>
<normalizedToken id="FDEFBCAE59964724E810AF5CC9890AB4" originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 1998), from Venezuela and Trinidad and Tobago (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
<taxonomicName id="DE167E96AF6A1733BB8872072A2FA1BB" class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
), or from Venezuela plus neighboring countries (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
<taxonomicName id="49E773AA5F56F31553ED6E5E114FA6E4" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
Simon, 1893). A close relationship among these genera had been suspected before based on morphology (
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
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), and molecular data have always supported this (
<bibRefCitation author="Bruvo-Mađaric, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="39" pageNumber="90" pagination="661 - 673" title="Phylogeny of pholcid spiders (Araneae: Pholcidae): combined analysis using morphology and molecules." url="https://doi.org/10.1016/j.ympev.2005.08.016" volume="37" year="2005">
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et al. 2005
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: 28S data and combined analysis;
<bibRefCitation author="Dimitrov, D" journalOrPublisher="Cladistics" pageId="39" pageNumber="90" pagination="132 - 146" title="Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group." url="https://doi.org/10.1111/j.1096-0031.2012.00419.x" volume="29" year="2013">Dimitrov et al. 2013</bibRefCitation>
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). Our data suggest that
<taxonomicName class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
<taxonomicName id="4A739964D29B8A4441BC795E449142DF" class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
may be nested within
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
<taxonomicName id="CDDBD5F8E4892E0C0343D00560411606" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
, but our taxon sampling is weak, the topology is unstable (
<taxonomicName class="Arachnida" family="Pholcidae" genus="Systenita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenita" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Systenita</taxonomicName>
<taxonomicName id="12ACBAAD7B41C08A71B18DE669F67FA5" class="Arachnida" family="Pholcidae" genus="Systenita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Systenita" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Systenita</taxonomicName>
is either nested within
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
<taxonomicName id="622D55FA13E50AACA262289D780A399F" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecolaesthus" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Mecolaesthus</taxonomicName>
or not), and several internal nodes in the clade have low support. Formally,
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<taxonomicName id="E085BBBBC00065643E1D34FBDBF3E611" class="Arachnida" family="Pholcidae" genus="Coryssocnemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryssocnemis" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">Coryssocnemis</taxonomicName>
still includes several obviously misplaced species: several Central American species (see
<pageBreakToken pageId="9" pageNumber="60" start="start">above</pageBreakToken>
<pageBreakToken id="5FC3F849DF34845EEB5BE358A731B748" pageId="9" pageNumber="60" start="start">above</pageBreakToken>
), and several Atlantic Forest (Brazilian) species whose identity is probably impossible to resolve (poor descriptions, lost types; see
<bibRefCitation author="Huber, BA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="91" pagination="1 - 348" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." url="https://doi.org/10.1206/0003-0090(2000)254&lt;0001:NWPSAP&gt;2.0.CO;2" volume="254" year="2000">Huber 2000</bibRefCitation>
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,
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">2018</bibRefCitation>
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).
</paragraph>
<paragraph pageId="9" pageNumber="60">
<paragraph id="427D943FB35C4E4653E7FE8FD5C8D17C" pageId="9" pageNumber="60">
The third subclade in the South American clade is
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<taxonomicName id="9C04473C20CF26C0818ECE4F424B7506" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
(Figure 5). Unlike
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="0414FEAFA5DD7DAB5E98BA82F785C4E5" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
it is monophyletic and apparently less problematic, but just as
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="0437503D21A4985404E5939993A5732B" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
, the genus has become very difficult to diagnose, mainly because of
<normalizedToken originalValue="untypical">'untypical'</normalizedToken>
<normalizedToken id="C258A43370939192D2F148AAB009A260" originalValue="untypical">'untypical'</normalizedToken>
species added to the genus based in large part on the present molecular data (
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber 2018</bibRefCitation>
<bibRefCitation id="38A74036D2259FED30CB9BEE090709C0" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber 2018</bibRefCitation>
). Both for
<taxonomicName class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
<taxonomicName id="B9537E7D80A971543EC20A0F4DD0D725" class="Arachnida" family="Pholcidae" genus="Mesabolivar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesabolivar" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Mesabolivar</taxonomicName>
and
<taxonomicName class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
<taxonomicName id="C87E659CC8B684E8427DDF98F80F6E4A" class="Arachnida" family="Pholcidae" genus="Carapoia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carapoia" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Carapoia</taxonomicName>
our analyses suggest several species groups that are also supported by morphological data. For a detailed discussion of these groups, see
<bibRefCitation author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber (2018)</bibRefCitation>
<bibRefCitation id="B6CC0B74B1CC1878C2F68FF05EF243F6" author="Huber, BA" journalOrPublisher="Zootaxa" pageId="41" pageNumber="92" pagination="1 - 178" title="The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits." url="https://doi.org/10.11646/zootaxa.4395.1.1" volume="4395" year="2018">Huber (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>