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Genus
<taxonomicName authority="Short &amp; Garcia, 2014" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<taxonomicName id="5B9DD909263CB3CC5EDDFA6ADDA09D1A" ID-CoL="8XPKV" authority="Short &amp; Garcia, 2014" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
Radicitus Short &amp;
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, 2014
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Variation across Acidocerinae, dorsal and lateral views A Colossochares ellipticus B Peltochares sp. C Peltochares conspicuus D Aulonochares tubulus E Helochares sp. F Helochares tristis G Novochares sp. H Helopeltarium ferrugineum I Batochares sp. J Helobata larvalis K Radicitus sp. L Nanosaphes tricolor M Agraphydrus cf. attenuatus N Tobochares luteomargo O Tobochares sulcatus P Quadriops similaris Q Crucisternum ouboteri R Primocerus neutrum S Agraphydrus coomani T Agraphydrus sp. U Globulosis flavus V Crephelochares nitescens." figureDoi="10.3897/zookeys.1045.63810.figure1" httpUri="https://binary.pensoft.net/fig/557400" pageId="0" pageNumber="1">Figs 1K</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of the Acidocerinae simplified from Short et al. (2021), indicating the distribution, preferred habitat, and currently described number of species for each genus. For habitat, filled black circles indicate that at least some species of the genus are commonly found in this habitat; light grey circles indicate the genus has been found in this habitat, but is rare or not typical for the group; white circles indicate no species have been recorded for the genus in this habitat." figureDoi="10.3897/zookeys.1045.63810.figure2" httpUri="https://binary.pensoft.net/fig/557401" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">, 49G-L</figureCitation>
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">, 50</figureCitation>
<figureCitation id="51B78D90978E2A4BA092446FDEF98E4E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Variation across Acidocerinae, dorsal and lateral views A Colossochares ellipticus B Peltochares sp. C Peltochares conspicuus D Aulonochares tubulus E Helochares sp. F Helochares tristis G Novochares sp. H Helopeltarium ferrugineum I Batochares sp. J Helobata larvalis K Radicitus sp. L Nanosaphes tricolor M Agraphydrus cf. attenuatus N Tobochares luteomargo O Tobochares sulcatus P Quadriops similaris Q Crucisternum ouboteri R Primocerus neutrum S Agraphydrus coomani T Agraphydrus sp. U Globulosis flavus V Crephelochares nitescens." figureDoi="10.3897/zookeys.1045.63810.figure1" httpUri="https://binary.pensoft.net/fig/557400" pageId="0" pageNumber="1">Figs 1K</figureCitation>
<figureCitation id="FF74C4B8BD455E1D2C3A0E8BF9476697" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of the Acidocerinae simplified from Short et al. (2021), indicating the distribution, preferred habitat, and currently described number of species for each genus. For habitat, filled black circles indicate that at least some species of the genus are commonly found in this habitat; light grey circles indicate the genus has been found in this habitat, but is rare or not typical for the group; white circles indicate no species have been recorded for the genus in this habitat." figureDoi="10.3897/zookeys.1045.63810.figure2" httpUri="https://binary.pensoft.net/fig/557401" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation id="4403CDC21931E9B2821DDDBE74D318F4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation id="E6C4675574C61FD6E0B10DFB4B222468" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">, 49G-L</figureCitation>
<figureCitation id="4F2092989923126E8D3A4B7532FCBC57" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">, 50</figureCitation>
</paragraph>
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<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Radicitus</taxonomicName>
<subSubSection id="1113721FAFBD8186828F074B1B24D124" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="EF76733AE2DB57DEF425F3C0ABA5AC94" pageId="0" pageNumber="1">
<taxonomicName id="C1A7C943D689D71DED4F2F3C1D72FC96" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Radicitus</taxonomicName>
Short &amp;
<normalizedToken originalValue="García">Garcia</normalizedToken>
<normalizedToken id="19AA0101235DADA828F749F07D2006BA" originalValue="García">Garcia</normalizedToken>
, 2014: 252.
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="gender">
<paragraph pageId="0" pageNumber="1">Gender.</paragraph>
<paragraph pageId="0" pageNumber="1">Masculine.</paragraph>
<subSubSection id="7E79C9BFFFF7CEEACEEDEC143324B487" pageId="0" pageNumber="1" type="gender">
<paragraph id="8E3C8CD3673731868B542F0ED73A0C77" pageId="0" pageNumber="1">Gender.</paragraph>
<paragraph id="B74F99E17F8E2B512C8FC2D58314E9F4" pageId="0" pageNumber="1">Masculine.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Short &amp; M.Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus ayacucho" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ayacucho">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus ayacucho</emphasis>
<subSubSection id="1CE41404F26AACD7D747EBF335CBA0D8" pageId="0" pageNumber="1" type="type species">
<paragraph id="A779C992E9258268A8173D86982BC76D" pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph id="E0F03B51A5EEAB9381BEDF03BCFEE595" pageId="0" pageNumber="1">
<taxonomicName id="4F61AB58F113F4FDD8489D709CD88373" authorityName="Short &amp; M.Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus ayacucho" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ayacucho">
<emphasis id="6334DF7662BD688B87913791EE32ADB5" italics="true" pageId="0" pageNumber="1">Radicitus ayacucho</emphasis>
</taxonomicName>
Short &amp;
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<normalizedToken id="8B9676CF8014DC0099D75AD96B24BBCD" originalValue="García">Garcia</normalizedToken>
, 2014: 252; by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="14BE9AA10FAF7EFE8164EB07DB0F85BF" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="79E9BF3541BF78678538AFF7FF9C826F" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="DF130C99462CCC070E3C7A9A106A2442" pageId="0" pageNumber="1">
Medium sized beetles, body length 4.5-6.2 mm. Body shape oval in dorsal view; moderate to strongly convex in lateral view; dorsal outline nearly straight and anteriorly inclined along anterior half (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50</figureCitation>
<figureCitation id="202A15FA6689A466B73ABE23A30CB98D" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50</figureCitation>
). Color dark brown, usually uniform along body regions, sometimes margins of pronotum and elytra slightly paler; ground punctation fine, moderately marked (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50A, D</figureCitation>
<figureCitation id="FA45E2336A7DF365D029E91AF2FD79B6" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50A, D</figureCitation>
). Shape of head trapezoid and rather wide. Eyes moderate in size, not emarginated anteriorly, slightly projected from outline of head. Clypeus trapezoid, with anterior margin broadly, roundly, and weakly emarginate. Labrum fully exposed. Mentum medially rather broadly depressed, laterally longitudinally elevated; median anterior depression marked by transverse nearly straight carina (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C, F</figureCitation>
<figureCitation id="1D48177A718EB9198EC9C46BCC32C1DE" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C, F</figureCitation>
). Antennae with nine antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps short and stout, nearly as long as half width of head (e.g., Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C</figureCitation>
<figureCitation id="DD5E87A6D26CBCF9FA03651A454279FB" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C</figureCitation>
); inner margin of maxillary palpomere 2 nearly straight, outer margin strongly curved along apical 2/3; maxillary palpomere 4 slightly shorter than 3. Prosternum flat, only slightly carinate along midline of anterior projection. Elytra without sutural striae; elytral punctures shallow to moderately marked; ground punctures rather uniformly distributed; some species with serial punctures clearly visible along posterior third of elytra; outer margins of elytra slightly flared (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50A, D</figureCitation>
<figureCitation id="D0D67C6974F4595CE2C76B83819616DF" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50A, D</figureCitation>
). Posterior elevation of mesoventrite with median longitudinal carina elevated and forming posteriorly pointing process; anapleural sutures strongly concave, separated at anterior margin by distance nearly half width of anterior margin of mesepisternum. Metaventrite sometimes with posteromesal glabrous patch. Protibiae with anterior row of spines completely reduced; apical spurs of protibiae stout, reaching apex of protarsomere 3. Metafemora with tibial grooves very sharply marked and covered by hydrofuge pubescence; hydrofuge pubescence restricted to dorsal half on basal three-quarters of anterior surface of metafemora (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C, F</figureCitation>
<figureCitation id="F514F40CB4934C6C2F37F069904FF505" captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Habitus of Radicitus spp. A-C R. ayacucho: A dorsal habitus B lateral habitus C ventral habitus D-F R. granitum: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1">50C, F</figureCitation>
). Tarsomeres 1-4 with long spiniform setae on ventral face; metatarsomere 2 nearly as long as 5 and as 3 and 4 combined. Fifth abdominal ventrite evenly rounded, without apical emargination or fringe of stout setae. Aedeagus either trilobed (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">49I-L</figureCitation>
<figureCitation id="CBA2B88F38D1FCE8B7538D57A0D5B727" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">49I-L</figureCitation>
) or divided (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">49G, H</figureCitation>
<figureCitation id="3EA7DC4F5A114479F47362F63246EADF" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Aedeagi of Quadriops, Radicitus and Sindolus spp. A Q. clusia B Q. depressus C Q. reticulatus D Q. similaris E S. sp. (Venezuela) F. S. sp. (Venezuela) G, H R. ayacucho: G dorsal view H lateral view I, J R. cf. granitum (Suriname): I dorsal view J lateral view K, L R. surinamensis: K dorsal view L lateral view. Scale bars: 0.1 mm (A-D); 0.5 mm (E-L)." figureDoi="10.3897/zookeys.1045.63810.figure49" httpUri="https://binary.pensoft.net/fig/557448" pageId="0" pageNumber="1">49G, H</figureCitation>
), with basal piece short and rather simple parameres separated from each other for most of their lengths; gonopore well developed.
</paragraph>
<caption doi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
<caption id="9668BAEF66837D19EA3B3BD423462DEA" doi="10.3897/zookeys.1045.63810.figure50" httpUri="https://binary.pensoft.net/fig/557449" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
<paragraph id="023DB07898E79C6ECF4BDD68972F74F0" pageId="0" pageNumber="1">
<emphasis id="03B98A84FD18DB28E0328DAA9D19A749" bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
Habitus of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<taxonomicName id="8578184225ACF2DCC97B3D073DD7D0FD" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EAD8898F4F3FF1B31261ED4719DD4848" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName lsidName="R. ayacucho" pageId="0" pageNumber="1" rank="species" species="ayacucho">
<emphasis italics="true" pageId="0" pageNumber="1">R. ayacucho</emphasis>
<emphasis id="D52E9527C44AE7547030BC6C3417122F" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName id="0200B78E9AC7C6452008776EE49079AC" lsidName="R. ayacucho" pageId="0" pageNumber="1" rank="species" species="ayacucho">
<emphasis id="DA31B76BC1D6B22D1C8B30F8A9A9E152" italics="true" pageId="0" pageNumber="1">R. ayacucho</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="A8CDD155F3CDA4AE993CB4C441FA5A63" bold="true" pageId="0" pageNumber="1">A</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="B770FB69B08EEACDFF7FE8BF130D8DDB" bold="true" pageId="0" pageNumber="1">B</emphasis>
lateral habitus
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="9A7761F0967F5B4352AD5DE42DA02AAA" bold="true" pageId="0" pageNumber="1">C</emphasis>
ventral habitus
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName lsidName="R. granitum" pageId="0" pageNumber="1" rank="species" species="granitum">
<emphasis italics="true" pageId="0" pageNumber="1">R. granitum</emphasis>
<emphasis id="AF9F8E015E0A8F18D0F9DF5414807D20" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName id="48475AC015B48249E72B2C682E66A865" lsidName="R. granitum" pageId="0" pageNumber="1" rank="species" species="granitum">
<emphasis id="1322557B7E7BA92E4EED8429DE4111B5" italics="true" pageId="0" pageNumber="1">R. granitum</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="DB84592C88EF2DBABFE45F6A1E8090E5" bold="true" pageId="0" pageNumber="1">D</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="0DAA4BFB5DE0BC6636F7925F7ECC38F0" bold="true" pageId="0" pageNumber="1">E</emphasis>
lateral habitus
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<emphasis id="A04874482D4CD80401D47B3C00F226FB" bold="true" pageId="0" pageNumber="1">F</emphasis>
ventral habitus. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<subSubSection id="0232F3569F6BBB5795AC693453EAD139" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="98A342278F40B8621EDC7080F1BF8B74" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="74260F94A13EF3D85650B95CF63509B5" pageId="0" pageNumber="1">
<taxonomicName id="6373AAA5CDECBB243916547C12552A47" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="73569B5033B6261A4380769E83202551" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
may resemble some punctate
<taxonomicName authorityName="Giron &amp; Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Novochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Novochares</emphasis>
<taxonomicName id="CB04B186E0E73FB525809EC0248F852C" authorityName="Giron &amp; Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="E172CFB54D10A881F406CE123815724B" italics="true" pageId="0" pageNumber="1">Novochares</emphasis>
</taxonomicName>
but can be recognized by the short and stout maxillary palps, along with metafemora only partly covered by pubescence (long and slender maxillary palps with metafemora mostly covered by pubescence in
<taxonomicName authorityName="Giron &amp; Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Novochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Novochares</emphasis>
<taxonomicName id="A32F94D0A540D698DF32EF68620CB76F" authorityName="Giron &amp; Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="A7D1B870CBE8D6CCEEB6CDA2FE1A6487" italics="true" pageId="0" pageNumber="1">Novochares</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Neotropical</emphasis>
<subSubSection id="387FEFB58247297DD1D26C457DB7C095" pageId="0" pageNumber="1" type="distribution">
<paragraph id="4DB40DB793761DF06EF621156C5C2479" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="66AE8F3876F32ADAF91337B6159717D2" pageId="0" pageNumber="1">
<emphasis id="C6EC1091792C8B9FA0AE4F579BF8E574" bold="true" pageId="0" pageNumber="1">Neotropical</emphasis>
: Guyana, Suriname, Venezuela; Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">6</figureCitation>
<figureCitation id="851A2BC4494F4BCAEA5DBF034E581FA5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="natural history">
<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="8C02BDF1FEF310C83433C62093CEFAB9" pageId="0" pageNumber="1" type="natural history">
<paragraph id="B9B135BC7C1B93AEBFF3EAFC42C605DD" pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph id="1A904236D42ABD1E19EC7EA2C2E7ECDA" pageId="0" pageNumber="1">
Species of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<taxonomicName id="13E8DC71B46D9924C3DFC10DEE389D6E" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="7475D3152095B0C80EAC2A6ADDB7BCD8" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
have been found on a variety of habitats associated with streams and seeps on rock outcrops. Some have been collected by submerging root mats found along streams, and in the roots of vegetation growing on seepage areas on granite outcrops (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3835.2.5" author="Short, AEZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="251 - 262" refId="B268" refString="Short, AEZ, Garcia, M, 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 3835 (2): 251 - 262, DOI: https://doi.org/10.11646/zootaxa.3835.2.5" title="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.11646/zootaxa.3835.2.5" volume="3835" year="2014">
<bibRefCitation id="28A959590E66CE56B6E3644869E682CA" DOI="https://doi.org/10.11646/zootaxa.3835.2.5" author="Short, AEZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="251 - 262" refId="B268" refString="Short, AEZ, Garcia, M, 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 3835 (2): 251 - 262, DOI: https://doi.org/10.11646/zootaxa.3835.2.5" title="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.11646/zootaxa.3835.2.5" volume="3835" year="2014">
Short and
<normalizedToken originalValue="García">Garcia</normalizedToken>
<normalizedToken id="0208536CAEEBF3F396BD9474A54C3CA3" originalValue="García">Garcia</normalizedToken>
2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="larvae">
<paragraph pageId="0" pageNumber="1">Larvae.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="4513A081261EDC01D6AFA79566FC9DFB" pageId="0" pageNumber="1" type="larvae">
<paragraph id="5EFF7E1C3FE3754EFCA6A6B9781774E2" pageId="0" pageNumber="1">Larvae.</paragraph>
<paragraph id="EAF94D63D4FA9704CF548B215A4CECFF" pageId="0" pageNumber="1">
The immature stages of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<taxonomicName id="A72C6DC1AC0B1C72B7F7B6BAC2FAB9F2" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="A486FFFC402B6ED977AB0D223643C066" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
remain unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="taxonomic history">
<paragraph pageId="0" pageNumber="1">Taxonomic history.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<subSubSection id="4BC2C5FA6AB4872D658E76595BD74BAE" pageId="0" pageNumber="1" type="taxonomic history">
<paragraph id="DD867016939290F7406EC4767E862423" pageId="0" pageNumber="1">Taxonomic history.</paragraph>
<paragraph id="CCF26B8377A793399814BEE346647FFE" pageId="0" pageNumber="1">
<taxonomicName id="819609CD560FE48988FA28B2E1D1B1CD" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="094AE5808D7B4332B13BED99A33B8429" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
was only recently described.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="C3CC6E86F179D73B37FAD2A9A100CC0B" pageId="0" pageNumber="1" type="remarks">
<paragraph id="EC5AF504EB436531E64A7C470CE8EE25" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="42C2DA1FF9871E52D7547AE1A88D1E45" pageId="0" pageNumber="1">
There are three known species of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
<taxonomicName id="969DD1B5C65BB5D3F86459EB3DD8C7F3" authorityName="Short &amp; Garcia" authorityYear="2014" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="DBB7E08AA09A706C1EAB65A0E5B64DB5" italics="true" pageId="0" pageNumber="1">Radicitus</emphasis>
</taxonomicName>
, all currently endemic to the Guiana Shield.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="selected references">
<paragraph pageId="0" pageNumber="1">Selected references.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3835.2.5" author="Short, AEZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="251 - 262" refId="B268" refString="Short, AEZ, Garcia, M, 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 3835 (2): 251 - 262, DOI: https://doi.org/10.11646/zootaxa.3835.2.5" title="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.11646/zootaxa.3835.2.5" volume="3835" year="2014">
<subSubSection id="E7906F487ED4ED6B59AB8DA15017CF77" pageId="0" pageNumber="1" type="selected references">
<paragraph id="ABFF385922DB14CC7FFC827157601172" pageId="0" pageNumber="1">Selected references.</paragraph>
<paragraph id="B9076D41798690777730208EA46939D1" pageId="0" pageNumber="1">
<bibRefCitation id="8D98D77BC837D939D768B90B9C5B0FA5" DOI="https://doi.org/10.11646/zootaxa.3835.2.5" author="Short, AEZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="251 - 262" refId="B268" refString="Short, AEZ, Garcia, M, 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 3835 (2): 251 - 262, DOI: https://doi.org/10.11646/zootaxa.3835.2.5" title="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.11646/zootaxa.3835.2.5" volume="3835" year="2014">
Short and
<normalizedToken originalValue="García">Garcia</normalizedToken>
<normalizedToken id="3CED0A80AC7151A58D20D59FA801A072" originalValue="García">Garcia</normalizedToken>
2014
</bibRefCitation>
: original description of the genus and all known species;
<bibRefCitation DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. 2021</bibRefCitation>
<bibRefCitation id="2A086A561177708A3E48FAEB6DECB807" DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. 2021</bibRefCitation>
: phylogenetic placement.
</paragraph>
</subSubSection>