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<mods:title>The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species</mods:title>
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<mods:title id="413E9B6C534661BCC0B84FF0DF54CD98">The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species</mods:title>
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<mods:namePart>Giron, Jennifer C.</mods:namePart>
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<mods:affiliation>Department of Entomology, Purdue University, West Lafayette, Indiana, USA & Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, Texas, USA</mods:affiliation>
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<mods:affiliation id="0329CD8F38ACB92828C967E96918BBED">Department of Entomology, Purdue University, West Lafayette, Indiana, USA & Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, Texas, USA</mods:affiliation>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
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Genus
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<taxonomicName authority="Short & Garcia, 2007" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<taxonomicName id="82C916C03FF227625B3817E96C5E6CAE" ID-CoL="8X7X8" authority="Short & Garcia, 2007" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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Tobochares Short &
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, 2007
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Variation across Acidocerinae, dorsal and lateral views A Colossochares ellipticus B Peltochares sp. C Peltochares conspicuus D Aulonochares tubulus E Helochares sp. F Helochares tristis G Novochares sp. H Helopeltarium ferrugineum I Batochares sp. J Helobata larvalis K Radicitus sp. L Nanosaphes tricolor M Agraphydrus cf. attenuatus N Tobochares luteomargo O Tobochares sulcatus P Quadriops similaris Q Crucisternum ouboteri R Primocerus neutrum S Agraphydrus coomani T Agraphydrus sp. U Globulosis flavus V Crephelochares nitescens." figureDoi="10.3897/zookeys.1045.63810.figure1" httpUri="https://binary.pensoft.net/fig/557400" pageId="0" pageNumber="1">Figs 1N, O</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of the Acidocerinae simplified from Short et al. (2021), indicating the distribution, preferred habitat, and currently described number of species for each genus. For habitat, filled black circles indicate that at least some species of the genus are commonly found in this habitat; light grey circles indicate the genus has been found in this habitat, but is rare or not typical for the group; white circles indicate no species have been recorded for the genus in this habitat." figureDoi="10.3897/zookeys.1045.63810.figure2" httpUri="https://binary.pensoft.net/fig/557401" pageId="0" pageNumber="1">, 2</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">, 6</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">, 11A, B</figureCitation>
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">, 52</figureCitation>
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">, 53</figureCitation>
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<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">, 54</figureCitation>
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<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">, 55</figureCitation>
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<figureCitation id="2C83054C0B30C87C054DEC046F7C3487" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Variation across Acidocerinae, dorsal and lateral views A Colossochares ellipticus B Peltochares sp. C Peltochares conspicuus D Aulonochares tubulus E Helochares sp. F Helochares tristis G Novochares sp. H Helopeltarium ferrugineum I Batochares sp. J Helobata larvalis K Radicitus sp. L Nanosaphes tricolor M Agraphydrus cf. attenuatus N Tobochares luteomargo O Tobochares sulcatus P Quadriops similaris Q Crucisternum ouboteri R Primocerus neutrum S Agraphydrus coomani T Agraphydrus sp. U Globulosis flavus V Crephelochares nitescens." figureDoi="10.3897/zookeys.1045.63810.figure1" httpUri="https://binary.pensoft.net/fig/557400" pageId="0" pageNumber="1">Figs 1N, O</figureCitation>
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<figureCitation id="F3BF05C0B4DF35369D66B8E6D0AEACA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of the Acidocerinae simplified from Short et al. (2021), indicating the distribution, preferred habitat, and currently described number of species for each genus. For habitat, filled black circles indicate that at least some species of the genus are commonly found in this habitat; light grey circles indicate the genus has been found in this habitat, but is rare or not typical for the group; white circles indicate no species have been recorded for the genus in this habitat." figureDoi="10.3897/zookeys.1045.63810.figure2" httpUri="https://binary.pensoft.net/fig/557401" pageId="0" pageNumber="1">, 2</figureCitation>
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<figureCitation id="B6FDAAF60284A2A5638ACE04586CF22B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">, 6</figureCitation>
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<figureCitation id="DA5BB6DCAD9334AB34FFB33ECDB00960" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">, 11A, B</figureCitation>
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<figureCitation id="8F9001778D40A2ACA3E5EC59801BF28F" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">, 52</figureCitation>
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<figureCitation id="6A1A1A317D7713D0603897A88E34CECB" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">, 53</figureCitation>
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<figureCitation id="1DFAA9DBF8B4B2A023352ACA6070FC7C" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">, 54</figureCitation>
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<figureCitation id="1D2DA172E7B4F2B59375C12BBB40E705" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">, 55</figureCitation>
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<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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|
||||
<taxonomicName id="00DA5BA680AEA15C2D409E7B7D954CBA" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
|
||||
Short &
|
||||
<normalizedToken originalValue="García">Garcia</normalizedToken>
|
||||
<normalizedToken id="D924822E2591157FF321025964CE4F61" originalValue="García">Garcia</normalizedToken>
|
||||
, 2007: 2.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="gender">
|
||||
<paragraph pageId="0" pageNumber="1">Gender.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Masculine.</paragraph>
|
||||
<subSubSection id="261876DC81DE29059DD67181A698E2E1" pageId="0" pageNumber="1" type="gender">
|
||||
<paragraph id="43590F872B3DBD92AEE0384311F9E9A0" pageId="0" pageNumber="1">Gender.</paragraph>
|
||||
<paragraph id="9EE157B80131406ABF64ECCF50A8A254" pageId="0" pageNumber="1">Masculine.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type species">
|
||||
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Short & M.Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares sulcatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sulcatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares sulcatus</emphasis>
|
||||
<subSubSection id="C1E4A94EF15CFDB3C0BC4637759E83EA" pageId="0" pageNumber="1" type="type species">
|
||||
<paragraph id="2AAB196444347A7B8FBE0A3410177E5D" pageId="0" pageNumber="1">Type species.</paragraph>
|
||||
<paragraph id="AF114B675F720C7AC67EA588DB7DB957" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="2D505690D6322B72B9D58262AA6E603E" authorityName="Short & M.Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares sulcatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sulcatus">
|
||||
<emphasis id="F92D988270AFEFAD6182F9A6D714008E" italics="true" pageId="0" pageNumber="1">Tobochares sulcatus</emphasis>
|
||||
</taxonomicName>
|
||||
Short &
|
||||
<normalizedToken originalValue="García">Garcia</normalizedToken>
|
||||
<normalizedToken id="93390477657EBB4559CFABDA79DF33C6" originalValue="García">Garcia</normalizedToken>
|
||||
, 2007: 4; by original designation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="EDF86296175A6B053301C7C5FC1F6CC4" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="59585579409508D69F4DD29BC82E89B6" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="B3B4D13DFA7BCCEE2C5315E6BAF2105B" pageId="0" pageNumber="1">
|
||||
Small beetles, total body length 1.5-2.6 mm. Body shape oval in dorsal view; moderately to strongly convex in lateral view (Fig.
|
||||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52</figureCitation>
|
||||
<figureCitation id="63A92EBBE9B0C885C66135DBC9DC1D4E" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52</figureCitation>
|
||||
-
|
||||
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54</figureCitation>
|
||||
<figureCitation id="1CDEC0C8C238B4FB91325C89AB1399BE" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54</figureCitation>
|
||||
); dorsal outline usually evenly curved. Color yellowish brown, orange brown to dark brown, sometimes with paler spots on head, or paler margins of pronotum and elytra; ground punctation moderate to shallow. Shape of head somewhat oval. Eyes not emarginate (e.g., Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">11A</figureCitation>
|
||||
<figureCitation id="74564CB56AA9E713188C07C469D220E9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">11A</figureCitation>
|
||||
) to strongly emarginate (e.g., Fig.
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">11B</figureCitation>
|
||||
<figureCitation id="60AFA67A5F7D6AE7EF790BD2C5B2887A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Head of miscellaneous Acidocerinae A-D anterolateral view: A Tobochares luteomargo with white arrow pointing to straight anterior margin of eye B Tobochares emarginatus with white arrow pointing to Tobochares canthus emarginating anterior margin of eye C Quadriops politus with white arrow pointing to Tobochares canthus fully dividing the eye in dorsal and ventral faces D Batochares sp. black arrow pointing to transverse carina on labrum E-L dorsal view of head: E Batochares sp. F Helochares tristis G Crephelochares nitescens, H Chasmogenus australis with black arrow pointing to preclypeal membrane I Colossochares ellipticus J Aulonochares tubulus K Peltochares conspicuus L Helobata larvalis." figureDoi="10.3897/zookeys.1045.63810.figure11" httpUri="https://binary.pensoft.net/fig/557410" pageId="0" pageNumber="1">11B</figureCitation>
|
||||
), moderate to small in size, somewhat oval, slightly to strongly projected from outline of head. Clypeus trapezoid, with anterior margin broadly emarginate; membranous preclypeal area often visible. Labrum fully exposed, convex, and anteriorly emarginate. Mentum rather smooth, often medially depressed, or anteriorly shallowly crenulated; median anterior depression marked by transverse carina (e.g., Fig.
|
||||
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53C</figureCitation>
|
||||
<figureCitation id="9C1E601FB71492C50DDFDA2AF47A8EA2" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53C</figureCitation>
|
||||
). Submentum anteriorly smooth and shiny. Antennae with eight antennomeres, cupule slightly asymmetric with rounded outline. Maxillary palps from short and slender (slightly shorter than the width of the head; e.g., Fig.
|
||||
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53C</figureCitation>
|
||||
<figureCitation id="14191543E7349E6696AF08A9E275FFF1" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53C</figureCitation>
|
||||
) to very short and stout (nearly half the width of the head; Fig.
|
||||
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54E</figureCitation>
|
||||
<figureCitation id="50FE5CD8F521A31E13056B0BD97A1E3C" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54E</figureCitation>
|
||||
); maxillary palpomere 4 similar in length to slightly longer than palpomere 3; inner margin of maxillary palpomere 2 straight, outer margin strongly curved along apical 2/3. Elytra without sutural striae (in some species, stria 1 more strongly impressed along posterior half of elytra; Fig.
|
||||
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54C</figureCitation>
|
||||
<figureCitation id="0634D46D099588F498DACD8C8B1E3114" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Habitus of Tobochares spp. A, B T. kappel: A dorsal habitus B lateral habitus C, D T. akoerio: C dorsal habitus D lateral habitus E, F T. kolokoe: E dorsal habitus F lateral habitus G, H T. goias: G dorsal habitus H lateral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1">54C</figureCitation>
|
||||
); elytral punctures seemingly arranged in rows, in some species more pronounced; interserial punctures occasionally longitudinally aligned; serial punctures sometimes impressed into distinct grooves (e.g., Fig.
|
||||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52A</figureCitation>
|
||||
<figureCitation id="C98709A47A2FC3CFC62D8506D6B8EF9C" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52A</figureCitation>
|
||||
). Prosternum flat. Posterior elevation of mesoventrite either flat, bulging or with transverse or longitudinal ridge (Fig.
|
||||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14F, G</figureCitation>
|
||||
<figureCitation id="46C73FA53A326B1CF3C195EA4836E2CD" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14F, G</figureCitation>
|
||||
); anapleural sutures concave, separated at anterior margin by distance nearly 0.3-0.5
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="2230ECFA6024729A64CB635C9E8892D9" originalValue="×">x</normalizedToken>
|
||||
width of anterior margin of mesepisternum. Metaventrite densely pubescent, except for median glabrous patch, either ovoid and broad (Fig.
|
||||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14G</figureCitation>
|
||||
<figureCitation id="22DF2FB74C9A759984D928E538E9F667" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14G</figureCitation>
|
||||
) or longitudinal and narrow (Fig.
|
||||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14F</figureCitation>
|
||||
<figureCitation id="27FAED4A41D82A36C38A6119AFBF2E21" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Scanning electron micrographs of thorax in ventral view A, B prosternum: A Tobochares striatus with white arrow pointing to anterior projection B Quadriops reticulatus with white arrow pointing to anterior projection C-G mesoventrite and metaventrite: C Crucisternum ouboteri with white arrows pointing to anteriorly pointed transverse ridge and longitudinal carina of mesoventrite, metaventrite with median glabrous patch D Nanosaphes tricolor with black arrow pointing to longitudinal carina along mesoventrite and white arrows pointing to median and postero-lateral glabrous patches of metaventrite E Quadriops reticulatus with black arrow pointing to transverse carina across mesoventrite and metaventrite uniformly pubescent F Tobochares communis with black arrow pointing to longitudinal carina along mesoventrite and white arrow pointing to narrow postero-medial glabrous patch on metaventrite G Tobochares kasikasima with black arrow pointing to transverse elevation across mesoventrite and white arrow pointing to broad postero-medial glabrous patch on metaventrite. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1045.63810.figure14" httpUri="https://binary.pensoft.net/fig/557413" pageId="0" pageNumber="1">14F</figureCitation>
|
||||
). Protibiae with spines of anterior row hair-like, semi erect, relatively long and thick; apical spurs of protibia from very short and stout, to enlarged to reach apex of protarsomere 3. Tarsomeres 2-4 densely covered by hair-like spines on ventral face; metatarsomeres 1-4 similar in length, 5 nearly as long as 3 and 4 combined, or metatarsomere 2 similar in length to 5. Metafemora mostly glabrous, with only few scattered setae, sometimes with hydrofuge pubescence along basal half of anterodorsal margin (e.g., Figs
|
||||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52C, F</figureCitation>
|
||||
<figureCitation id="4261071403DB4A234B906D379AA3DBE3" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Habitus of Tobochares spp. A-C T. sulcatus: A dorsal habitus B lateral habitus C ventral habitus D-F T. luteomargo: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1">52C, F</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53 C, F</figureCitation>
|
||||
<figureCitation id="4C2AD81B331E13949FC0D9DDBEE6C306" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Habitus of Tobochares spp. A-C T. communis: A dorsal habitus B lateral habitus C ventral habitus D-F T. fusus: D dorsal habitus E lateral habitus F ventral habitus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1">53 C, F</figureCitation>
|
||||
). Fifth abdominal ventrite apically evenly rounded, without fringe of stout setae. Aedeagus trilobed (Fig.
|
||||
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">55</figureCitation>
|
||||
<figureCitation id="A3E5BCC41FE1FFC12879C107CAF24C00" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">55</figureCitation>
|
||||
), with basal piece usually very short (nearly 1/3 length of parameres); median lobe usually broader than each paramere; median lobe and parameres apically rounded to truncate; apex of median lobe seldom medially emarginated; gonopore well developed.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
|
||||
<caption id="85A82FDE30B16232C1CF7EA0C473E2A1" doi="10.3897/zookeys.1045.63810.figure52" httpUri="https://binary.pensoft.net/fig/557451" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
|
||||
<paragraph id="10DF893BFACE0A2983BD6F8447B74AE6" pageId="0" pageNumber="1">
|
||||
<emphasis id="E7EBFF790500A399E3752D957BE1BDE0" bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="6FA95101B004F9CDCDA91F18DD6EBFB7" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="22DB4265674E1552B147FB63DDEF2E09" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
<taxonomicName lsidName="T. sulcatus" pageId="0" pageNumber="1" rank="species" species="sulcatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. sulcatus</emphasis>
|
||||
<emphasis id="5B5F4451860C35EB3C7B4A661E895976" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
<taxonomicName id="7CC85D8A769C19CFF568929DB948A4C8" lsidName="T. sulcatus" pageId="0" pageNumber="1" rank="species" species="sulcatus">
|
||||
<emphasis id="FAEFF4527D14C375F939E21ED72C6374" italics="true" pageId="0" pageNumber="1">T. sulcatus</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="F430DEEAC68F14A7A77C408CDF6143B2" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="29166C8EBEE371B7C7DB8D92EA5591F5" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="316D5B8A0B9EED9D26B31DD304D126DE" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
ventral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
|
||||
<taxonomicName lsidName="T. luteomargo" pageId="0" pageNumber="1" rank="species" species="luteomargo">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. luteomargo</emphasis>
|
||||
<emphasis id="297B772425CD48597057953136955C3B" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
|
||||
<taxonomicName id="D59E6A2546DE640DBC8ABD619005EB89" lsidName="T. luteomargo" pageId="0" pageNumber="1" rank="species" species="luteomargo">
|
||||
<emphasis id="0D167F906E9D11A8D012272AB01E9C3A" italics="true" pageId="0" pageNumber="1">T. luteomargo</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="2CFB9207B0E77F5D0C79F49802412313" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="1459CF62BDE07F85C08827A1E4D85C0D" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="B05913F87CF1FE68A1BC1853C07D9927" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
ventral habitus. Scale bars: 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1" start="Figure 53" startId="F53">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 53.</emphasis>
|
||||
<caption id="3FA892C6363DD9C48D4863931D66F661" doi="10.3897/zookeys.1045.63810.figure53" httpUri="https://binary.pensoft.net/fig/557452" pageId="0" pageNumber="1" start="Figure 53" startId="F53">
|
||||
<paragraph id="9910A52845474A9E7040A84FFA985317" pageId="0" pageNumber="1">
|
||||
<emphasis id="F1F155F299192D2A1A543D4B8A10FB0E" bold="true" pageId="0" pageNumber="1">Figure 53.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="6AF2D49621739C5CC3ABC7D7C42BC5CB" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="41717551EE810BDAAD2B7D5140D1C2F8" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
<taxonomicName lsidName="T. communis" pageId="0" pageNumber="1" rank="species" species="communis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. communis</emphasis>
|
||||
<emphasis id="488780AE992CBB87E6BB12F59466A0E8" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
<taxonomicName id="9FBEED7F76B71E92FED1EC3572C1EDD2" lsidName="T. communis" pageId="0" pageNumber="1" rank="species" species="communis">
|
||||
<emphasis id="6E534614233A60BA6314380F76B3E121" italics="true" pageId="0" pageNumber="1">T. communis</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="9A34299CE3351BC3800A0E8695BED854" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="BF9D7195077F5F6B23CDB8C9693D4B4B" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="5DF3CF92C440484A190BDC63AE70644E" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
ventral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
|
||||
<taxonomicName lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
<emphasis id="FBF5C2A47C02FE139660D712FFBC5866" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
|
||||
<taxonomicName id="9E716D1E5DF4ED5A089A4B0052187D7D" lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis id="5FDC559BE4AD1957365193199239F221" italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="35A532ED24CF70ECD004B90DF50A8700" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="BE17990AAD1057C65B7C2410BF896421" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="F3A8E5C30C560811F9A4B878EA9897D6" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
ventral habitus. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<subSubSection id="8C7615B8B2E534C2A48454903322D89F" pageId="0" pageNumber="1" type="differential diagnosis">
|
||||
<paragraph id="3A7C7375118EEF700A322E0190A9755B" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||||
<paragraph id="F9A8920DA2AECCB3805C77B1CCC2ADAD" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="2DB3D1D4A8FBF4E11E6E1369AD71B9C2" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="EB542E92089BB1B8E9363597EB420693" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
are among the smallest acidocerines. Some members of the group are unique in the presence of impressed elytral striae (
|
||||
<taxonomicName authorityName="Short" authorityYear="2013" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares striatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="striatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares striatus</emphasis>
|
||||
<taxonomicName id="EC632092122B10DE038A329C6A647AF4" authorityName="Short" authorityYear="2013" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares striatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="striatus">
|
||||
<emphasis id="F9BBA4BBAA9AD36BC72C5FD7401A529F" italics="true" pageId="0" pageNumber="1">Tobochares striatus</emphasis>
|
||||
</taxonomicName>
|
||||
species group;
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken originalValue="Girón">Giron</normalizedToken>
|
||||
<bibRefCitation id="F1AAAEA122AAA42B7B49756DAE427DC3" DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken id="E99F44F39DEACA17C09CE59680133E87" originalValue="Girón">Giron</normalizedToken>
|
||||
and Short 2021a
|
||||
</bibRefCitation>
|
||||
).
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="D3ACEBDCA23D0ED0E1AFE7B1F33BB4DA" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1C306AA22E6C3B0664E09A51C9965B5C" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
without elytral striae may resemble some
|
||||
<taxonomicName authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
<taxonomicName id="911062818BC907FDA442926A512CBB24" authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="25D29460FFC99CCA4BDF4E0E1E786180" italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
</taxonomicName>
|
||||
(with eight antennomeres and mostly glabrous femora), and other than their distributions (
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="BAD2CCC5640A52648D485EC0AFB67A22" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="E7BA63073E7E0C66583159922056AC2F" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
in the New World,
|
||||
<taxonomicName authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
<taxonomicName id="F0A08B6FB3E01E4D79C46DD3740724FD" authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="91B20D5666B32ED943ED2AAC9E958A4D" italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
</taxonomicName>
|
||||
in the Old World) and slight differences in overall body shape, they can only be differentiated by the shape of the aedeagus (slender in
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="14466441CF7096C3B2C66BD88F03120D" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="8251A429477B9F97BE28F5EB364C3D33" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
, Fig.
|
||||
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">55</figureCitation>
|
||||
<figureCitation id="D5C6C1314783125365306CDA0E173579" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Aedeagi of Tobochares spp. A T. benettii B T. fusus C T. luteomargo D T. emarginatus E T. kusad F T. kasikasima G T. anthonyae H T. autures I T. communis J T. romanoae K T. akoreio. Scale bars: 0.5 mm (A-C); 0.1 mm (D-K)." figureDoi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1">55</figureCitation>
|
||||
; overall broader in
|
||||
<taxonomicName authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
<taxonomicName id="8CCAD630E914085856A97BD6E469B2CA" authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4ED61C02DD9ADA95B279AC057045B5CD" italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
|
||||
</taxonomicName>
|
||||
, Fig.
|
||||
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagi of Agraphydrus spp. A A. attenuatus B A. gracilipalpis C A. masatakai D A. chinensis E A. puzhelongi. Scale bars: 0.1 mm. Line drawings taken from Komarek (2018)." figureDoi="10.3897/zookeys.1045.63810.figure20" httpUri="https://binary.pensoft.net/fig/557419" pageId="0" pageNumber="1">20</figureCitation>
|
||||
<figureCitation id="4DA64F9E6156903508C22FEB6CA68CD6" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagi of Agraphydrus spp. A A. attenuatus B A. gracilipalpis C A. masatakai D A. chinensis E A. puzhelongi. Scale bars: 0.1 mm. Line drawings taken from Komarek (2018)." figureDoi="10.3897/zookeys.1045.63810.figure20" httpUri="https://binary.pensoft.net/fig/557419" pageId="0" pageNumber="1">20</figureCitation>
|
||||
). Within the New World,
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="F9E74F2CDC05B6F170E95848E7335229" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="F8BDCF7D060D03EEACB4A0D663641E7E" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
is most likely to be confused with
|
||||
<taxonomicName authorityName="Giron & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Ephydrolithus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephydrolithus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Ephydrolithus</emphasis>
|
||||
<taxonomicName id="A1E12C066F1EF4AD0C67443B2B70067C" authorityName="Giron & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Ephydrolithus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephydrolithus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A7BC7EB0FAA62081A90C3ED1607C01FC" italics="true" pageId="0" pageNumber="1">Ephydrolithus</emphasis>
|
||||
</taxonomicName>
|
||||
, which also contains small, seepage-inhabiting species, although currently the ranges of the two genera do not quite overlap. However, the difference in the number of antennomeres (nine in
|
||||
<taxonomicName authorityName="Giron & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Ephydrolithus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephydrolithus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Ephydrolithus</emphasis>
|
||||
<taxonomicName id="89DEA59CD565ECA25414838C50344C03" authorityName="Giron & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Ephydrolithus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephydrolithus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A9EDD5D60FE1F1D63CF6A8B2741B9CD7" italics="true" pageId="0" pageNumber="1">Ephydrolithus</emphasis>
|
||||
</taxonomicName>
|
||||
) provides a clear point of separation.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
|
||||
<caption id="C08F71CF02B3E267F069A9B62A3A11AF" doi="10.3897/zookeys.1045.63810.figure54" httpUri="https://binary.pensoft.net/fig/557453" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
|
||||
<paragraph id="82DC650B1D3155EDBC97E71832D5F271" pageId="0" pageNumber="1">
|
||||
<emphasis id="52B5A9602A1BA048125860E1AE57EAA5" bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="ED96997135C02AD6491AEE6FB9C8181E" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="61F594472E4E6958D20C50ED70EFBEB9" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||||
<taxonomicName lsidName="T. kappel" pageId="0" pageNumber="1" rank="species" species="kappel">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. kappel</emphasis>
|
||||
<emphasis id="E7FD1ADBD4F7C60B17345815A5896384" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||||
<taxonomicName id="A72FE01A36685F2382EB57A2E2BBA088" lsidName="T. kappel" pageId="0" pageNumber="1" rank="species" species="kappel">
|
||||
<emphasis id="90CB253428FD95EFACA233A66CD35681" italics="true" pageId="0" pageNumber="1">T. kappel</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="80AB2D4FB90F2815043A82BBDC67C2BD" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="356A791D2F84AFFE06193C610C881A8A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||||
<taxonomicName lsidName="T. akoerio" pageId="0" pageNumber="1" rank="species" species="akoerio">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. akoerio</emphasis>
|
||||
<emphasis id="3BBED99339E3114DA1F014D18E090B6C" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||||
<taxonomicName id="DDAB0F7CF7B3D5E778B6A89908018DFD" lsidName="T. akoerio" pageId="0" pageNumber="1" rank="species" species="akoerio">
|
||||
<emphasis id="06B4AD8AF591E3DE29BF5711325902D6" italics="true" pageId="0" pageNumber="1">T. akoerio</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="3D4B80D7199523B305EB5370D18B6FBE" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="A071986093A4B0DFE3BC9D520D9D4BB3" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||||
<taxonomicName lsidName="T. kolokoe" pageId="0" pageNumber="1" rank="species" species="kolokoe">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. kolokoe</emphasis>
|
||||
<emphasis id="0F1151388AA3916C46A460068CA94DAC" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||||
<taxonomicName id="B6237546616FE981956AE150FC563960" lsidName="T. kolokoe" pageId="0" pageNumber="1" rank="species" species="kolokoe">
|
||||
<emphasis id="467334CA50213FC9A0E00CBD01F27AD9" italics="true" pageId="0" pageNumber="1">T. kolokoe</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="05DF9702A30F3FEBA4703E9CA919BFEC" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="8B3499AEF2A37F6EE2A71F5F2FD60148" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
lateral habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
|
||||
<taxonomicName lsidName="T. goias" pageId="0" pageNumber="1" rank="species" species="goias">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. goias</emphasis>
|
||||
<emphasis id="74CCBAB922F3216E159766AA06BE88F2" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
|
||||
<taxonomicName id="3468589F9E568D6244AB37C92FD40EFA" lsidName="T. goias" pageId="0" pageNumber="1" rank="species" species="goias">
|
||||
<emphasis id="E924FA97515FD76B200631E4EDF5F5EA" italics="true" pageId="0" pageNumber="1">T. goias</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
<emphasis id="8CC9454441CF200961AA1A113D2EADFC" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
dorsal habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||||
<emphasis id="CFC16A98C10AA08987B148D5E678991F" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||||
lateral habitus. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
|
||||
<caption id="629578CEE6BF8271074DD64DCA998142" doi="10.3897/zookeys.1045.63810.figure55" httpUri="https://binary.pensoft.net/fig/557454" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
|
||||
<paragraph id="3F3EDE539E958772823A00C787FC4BB0" pageId="0" pageNumber="1">
|
||||
<emphasis id="E01F03338CA2216CAD0AE8773D1F75BF" bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
|
||||
Aedeagi of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="9048752059036502C2BE5AB1A40A60B1" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="23587C1933998318F81C56BFD2E01126" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
spp.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<taxonomicName lsidName="T. benettii" pageId="0" pageNumber="1" rank="species" species="benettii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. benettii</emphasis>
|
||||
<emphasis id="637BF03A341E4B5F771EB464DE0B71BB" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<taxonomicName id="9733CA38F74D0FFA671DD25E20FF5094" lsidName="T. benettii" pageId="0" pageNumber="1" rank="species" species="benettii">
|
||||
<emphasis id="C956345D459D0E8774249866F9632693" italics="true" pageId="0" pageNumber="1">T. benettii</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<taxonomicName lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
<emphasis id="54A361873CB0A77B35C3990A06BF4DB0" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<taxonomicName id="F3FAFA1E9BDE012A7B0D424C3A4DFB2F" lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis id="0CA428F1050C1CA9B066010EB5513309" italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<taxonomicName lsidName="T. luteomargo" pageId="0" pageNumber="1" rank="species" species="luteomargo">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. luteomargo</emphasis>
|
||||
<emphasis id="DF2B3DA2C345739610B9C77E77B9E1DB" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<taxonomicName id="2D880727CB28326032347EF03A019416" lsidName="T. luteomargo" pageId="0" pageNumber="1" rank="species" species="luteomargo">
|
||||
<emphasis id="160F79A6AEC45F05E42B59CE3672FAE3" italics="true" pageId="0" pageNumber="1">T. luteomargo</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<taxonomicName lsidName="T. emarginatus" pageId="0" pageNumber="1" rank="species" species="emarginatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. emarginatus</emphasis>
|
||||
<emphasis id="971D1B5B909E163B2A707FE1D03116E6" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<taxonomicName id="14C99F507572E539A70636D0C297144F" lsidName="T. emarginatus" pageId="0" pageNumber="1" rank="species" species="emarginatus">
|
||||
<emphasis id="6D9E3C668584EFDEF18DBF02409F9DD3" italics="true" pageId="0" pageNumber="1">T. emarginatus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<taxonomicName lsidName="T. kusad" pageId="0" pageNumber="1" rank="species" species="kusad">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. kusad</emphasis>
|
||||
<emphasis id="4C822553ADC60B2209BEB2D995FF2144" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<taxonomicName id="45E7611551F02B75145BE85CFDA467E4" lsidName="T. kusad" pageId="0" pageNumber="1" rank="species" species="kusad">
|
||||
<emphasis id="978AB76EEF6779B18279A9D8C7559DB3" italics="true" pageId="0" pageNumber="1">T. kusad</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<taxonomicName lsidName="T. kasikasima" pageId="0" pageNumber="1" rank="species" species="kasikasima">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. kasikasima</emphasis>
|
||||
<emphasis id="E592F89281140805D855BA42B30E6D0D" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<taxonomicName id="45F13A9168DD1F4831628698B1FAF304" lsidName="T. kasikasima" pageId="0" pageNumber="1" rank="species" species="kasikasima">
|
||||
<emphasis id="889167AA5B53F56509FF907DA4959A14" italics="true" pageId="0" pageNumber="1">T. kasikasima</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
<taxonomicName lsidName="T. anthonyae" pageId="0" pageNumber="1" rank="species" species="anthonyae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. anthonyae</emphasis>
|
||||
<emphasis id="6BC82185FC2FB9D5FDD40FE660E9D521" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||||
<taxonomicName id="A44F301A442FF5436545E224B19F02D8" lsidName="T. anthonyae" pageId="0" pageNumber="1" rank="species" species="anthonyae">
|
||||
<emphasis id="F988B07F191A9EEA01A113DBD7959AB8" italics="true" pageId="0" pageNumber="1">T. anthonyae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||||
<taxonomicName lsidName="T. autures" pageId="0" pageNumber="1" rank="species" species="autures">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. autures</emphasis>
|
||||
<emphasis id="2C00FE95C9CB74D49F13E86771998416" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||||
<taxonomicName id="3AA03A09B3F89ED2BCB1AD4AF5632D19" lsidName="T. autures" pageId="0" pageNumber="1" rank="species" species="autures">
|
||||
<emphasis id="A37E0C427E2DA4E467A1755319D2D5A3" italics="true" pageId="0" pageNumber="1">T. autures</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
|
||||
<taxonomicName lsidName="T. communis" pageId="0" pageNumber="1" rank="species" species="communis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. communis</emphasis>
|
||||
<emphasis id="A7EDF08C120355150BB68DCCD171F4AE" bold="true" pageId="0" pageNumber="1">I</emphasis>
|
||||
<taxonomicName id="93D5DA614BEF27591DC7B703382708A3" lsidName="T. communis" pageId="0" pageNumber="1" rank="species" species="communis">
|
||||
<emphasis id="D4199DEE210C63440D776C27CD479CDD" italics="true" pageId="0" pageNumber="1">T. communis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
|
||||
<taxonomicName lsidName="T. romanoae" pageId="0" pageNumber="1" rank="species" species="romanoae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. romanoae</emphasis>
|
||||
<emphasis id="9D936A6BFC753863CE325270718CE6DD" bold="true" pageId="0" pageNumber="1">J</emphasis>
|
||||
<taxonomicName id="19B58483DFC43166F12CC62CC810588D" lsidName="T. romanoae" pageId="0" pageNumber="1" rank="species" species="romanoae">
|
||||
<emphasis id="D7F895F64DE70C00F7FCD78CE2DF2715" italics="true" pageId="0" pageNumber="1">T. romanoae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
|
||||
<taxonomicName lsidName="T. akoreio" pageId="0" pageNumber="1" rank="species" species="akoreio">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. akoreio</emphasis>
|
||||
<emphasis id="9787A3788F95AD9AE4D3FA3686FEC5A1" bold="true" pageId="0" pageNumber="1">K</emphasis>
|
||||
<taxonomicName id="E60FF2264C24FC90592B535E9044BA7E" lsidName="T. akoreio" pageId="0" pageNumber="1" rank="species" species="akoreio">
|
||||
<emphasis id="6F39F53E170A7C2BECF3378A5C5835AE" italics="true" pageId="0" pageNumber="1">T. akoreio</emphasis>
|
||||
</taxonomicName>
|
||||
. Scale bars: 0.5 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
<emphasis id="4F2767A5ECDE70DDA46285362F11E921" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||||
); 0.1 mm (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D-K</emphasis>
|
||||
<emphasis id="DE83DA898C0A9F364BFB4C2A793731F9" bold="true" pageId="0" pageNumber="1">D-K</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Neotropical</emphasis>
|
||||
<subSubSection id="FB3B0A42CAA827DC1A303D0F11E0284F" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="4E3D30D5D3D0D91EDAF3F0A049C81644" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph id="0A26B52926065412774CCB3426B95348" pageId="0" pageNumber="1">
|
||||
<emphasis id="A814C39231BF4F4BE77D5C63150E57EE" bold="true" pageId="0" pageNumber="1">Neotropical</emphasis>
|
||||
: Brazil (
|
||||
<normalizedToken originalValue="Amapá">Amapa</normalizedToken>
|
||||
<normalizedToken id="E1BA739A357DFAD63E5249C17BB795C0" originalValue="Amapá">Amapa</normalizedToken>
|
||||
, Amazonas,
|
||||
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
|
||||
<normalizedToken id="06E8DC0C51366EE52E71B17FF1C11371" originalValue="Goiás">Goias</normalizedToken>
|
||||
, Roraima), French Guiana, Guyana, Suriname, Venezuela; Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">6</figureCitation>
|
||||
<figureCitation id="3528AFFCB2DE2D1F7B11B79472833A9C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">6</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="natural history">
|
||||
<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="35D20FA2E6FEF7E88A0C76FA1F52242C" pageId="0" pageNumber="1" type="natural history">
|
||||
<paragraph id="0218995B6A8EF9F124900FB22ECE62B9" pageId="0" pageNumber="1">Natural history.</paragraph>
|
||||
<paragraph id="E464E4E7F72271E9E09B7AFC04E8802B" pageId="0" pageNumber="1">
|
||||
Most
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="0F333F74D645C3BA812F051FE566C165" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="328D12C47582581977C9FDCAAA5C294B" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
specimens have been collected at hygropetric habitats, including isolated hygropetric seeps as well as wet rock surfaces along rivers and waterfalls. They can sometimes be found in large numbers. One species,
|
||||
<taxonomicName lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
<taxonomicName id="13484316C7AEC08C086E219C65706A34" lsidName="T. fusus" pageId="0" pageNumber="1" rank="species" species="fusus">
|
||||
<emphasis id="089B010812E56465D7C134D002DA03A9" italics="true" pageId="0" pageNumber="1">T. fusus</emphasis>
|
||||
</taxonomicName>
|
||||
, has been collected in both seepage habitats as well as terrestrially in the rotten fruits of
|
||||
<taxonomicName class="Insecta" family="Clusiidae" genus="Clusia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Clusia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Clusia</emphasis>
|
||||
<taxonomicName id="BB1981D0F96CB06F9C15706EBA40059E" class="Insecta" family="Clusiidae" genus="Clusia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Clusia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="AA2A5FE9481C49786FBC76DB18A07654" italics="true" pageId="0" pageNumber="1">Clusia</emphasis>
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
|
||||
<bibRefCitation id="8F92200D797E1877F44ECCB104E9B30F" DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken originalValue="Girón">Giron</normalizedToken>
|
||||
<bibRefCitation id="548B1E8C26D2B36C7D9BFD74175589D5" DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken id="E42C6E3B8039D6C49A26DC3D4B204518" originalValue="Girón">Giron</normalizedToken>
|
||||
and Short 2021a
|
||||
</bibRefCitation>
|
||||
for more details).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="larvae">
|
||||
<paragraph pageId="0" pageNumber="1">Larvae.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="DDA12910F68BA959A2FACF9625BD4846" pageId="0" pageNumber="1" type="larvae">
|
||||
<paragraph id="47FAB02DCB4E7BFC91115AEC00126EF3" pageId="0" pageNumber="1">Larvae.</paragraph>
|
||||
<paragraph id="7488DE935F6F5E7F10F01C23353D6C85" pageId="0" pageNumber="1">
|
||||
The immature stages of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="33650A06186D66EED19FDDD3683B426C" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="6806539CFD9DAB3FD197E888D01D1D5D" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
remain unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="taxonomic history">
|
||||
<paragraph pageId="0" pageNumber="1">Taxonomic history.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/01650420701247869" author="Short, AEZ" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="1" pagination="1 - 7" refId="B267" refString="Short, AEZ, Garcia, M, 2007. Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae). Aquatic Insects 29 (1): 1 - 7, DOI: https://doi.org/10.1080/01650420701247869" title="Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae)." url="https://doi.org/10.1080/01650420701247869" volume="29" year="2007">
|
||||
<subSubSection id="0E1984E4FA266035163F0F687811A62B" pageId="0" pageNumber="1" type="taxonomic history">
|
||||
<paragraph id="816C8904FC2A94D418EFF4D48DDCFA6E" pageId="0" pageNumber="1">Taxonomic history.</paragraph>
|
||||
<paragraph id="57D1BFD748DE5E449CEF59CFC35D93DE" pageId="0" pageNumber="1">
|
||||
<bibRefCitation id="0DAA22A11FAE9BE94AFC28A853B74361" DOI="https://doi.org/10.1080/01650420701247869" author="Short, AEZ" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="1" pagination="1 - 7" refId="B267" refString="Short, AEZ, Garcia, M, 2007. Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae). Aquatic Insects 29 (1): 1 - 7, DOI: https://doi.org/10.1080/01650420701247869" title="Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae)." url="https://doi.org/10.1080/01650420701247869" volume="29" year="2007">
|
||||
Short and
|
||||
<normalizedToken originalValue="García">Garcia</normalizedToken>
|
||||
<normalizedToken id="DDFA6F2DC9A15ACC96848E217AB95A75" originalValue="García">Garcia</normalizedToken>
|
||||
(2007)
|
||||
</bibRefCitation>
|
||||
described the genus and one species from Venezuela. Additional species were described from Suriname, one by
|
||||
<bibRefCitation author="Short, AEZ" editor="O'Shea, BJ" journalOrPublisher="RAP Bulletin of Biological Assessment 63, Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 90" publicationUrl="http://www.bioone.org/doi/abs/10.1896/054.063.0107" refId="B271" refString="Short, AEZ, Kadosoe, V, 2011. Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera). In: O'Shea, BJ, Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname. RAP Bulletin of Biological Assessment 63, Conservation International, Arlington: 79 - 90, http://www.bioone.org/doi/abs/10.1896/054.063.0107" title="Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera)." url="http://www.bioone.org/doi/abs/10.1896/054.063.0107" volumeTitle="A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname." year="2011">Short and Kadosoe (2011)</bibRefCitation>
|
||||
<bibRefCitation id="0435A5BC3CFFE543DCB5B819503D2B28" author="Short, AEZ" editor="O'Shea, BJ" journalOrPublisher="RAP Bulletin of Biological Assessment 63, Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 90" publicationUrl="http://www.bioone.org/doi/abs/10.1896/054.063.0107" refId="B271" refString="Short, AEZ, Kadosoe, V, 2011. Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera). In: O'Shea, BJ, Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname. RAP Bulletin of Biological Assessment 63, Conservation International, Arlington: 79 - 90, http://www.bioone.org/doi/abs/10.1896/054.063.0107" title="Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera)." url="http://www.bioone.org/doi/abs/10.1896/054.063.0107" volumeTitle="A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname." year="2011">Short and Kadosoe (2011)</bibRefCitation>
|
||||
and two more by
|
||||
<bibRefCitation author="Short, AEZ" editor="Alonso, LE" journalOrPublisher="RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 89" publicationUrl="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" refId="B263" refString="Short, AEZ, 2013. Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera). In: Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname. RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington: 79 - 89, https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" title="Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera)." url="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" volumeTitle="A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname." year="2013">Short (2013)</bibRefCitation>
|
||||
<bibRefCitation id="EAE40A8D06D6701A58CCDA1C4FE8BEA5" author="Short, AEZ" editor="Alonso, LE" journalOrPublisher="RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 89" publicationUrl="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" refId="B263" refString="Short, AEZ, 2013. Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera). In: Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname. RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington: 79 - 89, https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" title="Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera)." url="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" volumeTitle="A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname." year="2013">Short (2013)</bibRefCitation>
|
||||
. The genus was revised by
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short (2017)</bibRefCitation>
|
||||
<bibRefCitation id="D039776FF5E13A526F5F4D8E191004FC" DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short (2017)</bibRefCitation>
|
||||
, including the description of five new species and the characterization of one specimen from
|
||||
<normalizedToken originalValue="Tobogán">Tobogan</normalizedToken>
|
||||
<normalizedToken id="512C2E33305068BBFF98FF9BC040671C" originalValue="Tobogán">Tobogan</normalizedToken>
|
||||
de la Selva (Venezuela) left undescribed until additional material can be studied. The genus was reviewed again just a few years later by
|
||||
<normalizedToken originalValue="Girón">Giron</normalizedToken>
|
||||
<normalizedToken id="9021B5E8D629B8B6037D3837B67EFDFF" originalValue="Girón">Giron</normalizedToken>
|
||||
and Short (2021), in the light of new molecular evidence, describing 15 additional new species and establishing four diagnosable species groups.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="FE0DF68413C974D818A5A70B997747EF" pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph id="83957D71EFD8B047FA4A1AC94CBBD09C" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph id="95A38976BB39F73482244E6A45B77286" pageId="0" pageNumber="1">
|
||||
There are 24 described species of
|
||||
<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
<taxonomicName id="17FBA050FA3545DE3B3CD4711EB2986C" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="BBDDAC615683D55617117EF26B756CC9" italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
|
||||
</taxonomicName>
|
||||
. The genus is rather highly variable in its external morphology: there is variation in coloration, the degree of emargination of the eyes and the degree of development and extension of the elytral striae. The form of the aedeagus is also somewhat variable, although not as extreme as in some genera such as
|
||||
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Chasmogenus</emphasis>
|
||||
<taxonomicName id="ACE65ABD80B02F42ECF5A5303DD863F1" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1CB2950D25BF19A89E9A2E440B964D43" italics="true" pageId="0" pageNumber="1">Chasmogenus</emphasis>
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Helochares</emphasis>
|
||||
<taxonomicName id="50B1684AA8345485166249E728592588" authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="0D0333D4AC7935E09EE7F6488ECFFB8F" italics="true" pageId="0" pageNumber="1">Helochares</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">The genus is much richer in species and more broadly distributed in the Amazon region than as currently published. We have examined numerous additional specimens from around the Amazonian region, particularly the southern Amazon (e.g., Brazil: Rondonia) from where the genus is currently unknown. We would not be surprised if the genus exceeded 50 species when more attention is paid to seepage habitats in this region.</paragraph>
|
||||
<paragraph id="7A80C47BC9ED76809897E225947B7207" pageId="0" pageNumber="1">The genus is much richer in species and more broadly distributed in the Amazon region than as currently published. We have examined numerous additional specimens from around the Amazonian region, particularly the southern Amazon (e.g., Brazil: Rondonia) from where the genus is currently unknown. We would not be surprised if the genus exceeded 50 species when more attention is paid to seepage habitats in this region.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="species examined">
|
||||
<paragraph pageId="0" pageNumber="1">Species examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Holotypes, paratypes, and additional specimens of all described species, as well as several undescribed species were examined for this study.</paragraph>
|
||||
<subSubSection id="EE04C64A496A74AD71F86AF9B7EF7F10" pageId="0" pageNumber="1" type="species examined">
|
||||
<paragraph id="CC998C6B9159678E8F2357824C0B0F45" pageId="0" pageNumber="1">Species examined.</paragraph>
|
||||
<paragraph id="5B84F48A1D948D9136EC078A6240F88E" pageId="0" pageNumber="1">Holotypes, paratypes, and additional specimens of all described species, as well as several undescribed species were examined for this study.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="selected references">
|
||||
<paragraph pageId="0" pageNumber="1">Selected references.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/01650420701247869" author="Short, AEZ" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="1" pagination="1 - 7" refId="B267" refString="Short, AEZ, Garcia, M, 2007. Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae). Aquatic Insects 29 (1): 1 - 7, DOI: https://doi.org/10.1080/01650420701247869" title="Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae)." url="https://doi.org/10.1080/01650420701247869" volume="29" year="2007">
|
||||
<subSubSection id="9E78D53B90E0702DDA9F7AA0B30F4BB7" pageId="0" pageNumber="1" type="selected references">
|
||||
<paragraph id="7C354EE20C5282B18683B06532585922" pageId="0" pageNumber="1">Selected references.</paragraph>
|
||||
<paragraph id="137AD08AE577E764E3D68A6FCFA3A676" pageId="0" pageNumber="1">
|
||||
<bibRefCitation id="5B1D813F48D6CE0D911B0FFCF72B1107" DOI="https://doi.org/10.1080/01650420701247869" author="Short, AEZ" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="1" pagination="1 - 7" refId="B267" refString="Short, AEZ, Garcia, M, 2007. Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae). Aquatic Insects 29 (1): 1 - 7, DOI: https://doi.org/10.1080/01650420701247869" title="Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae)." url="https://doi.org/10.1080/01650420701247869" volume="29" year="2007">
|
||||
Short and
|
||||
<normalizedToken originalValue="García">Garcia</normalizedToken>
|
||||
<normalizedToken id="4641C91435D2688507D64C4E4343062E" originalValue="García">Garcia</normalizedToken>
|
||||
2007
|
||||
</bibRefCitation>
|
||||
: original description of the genus and its type species;
|
||||
<bibRefCitation author="Short, AEZ" editor="O'Shea, BJ" journalOrPublisher="RAP Bulletin of Biological Assessment 63, Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 90" publicationUrl="http://www.bioone.org/doi/abs/10.1896/054.063.0107" refId="B271" refString="Short, AEZ, Kadosoe, V, 2011. Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera). In: O'Shea, BJ, Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname. RAP Bulletin of Biological Assessment 63, Conservation International, Arlington: 79 - 90, http://www.bioone.org/doi/abs/10.1896/054.063.0107" title="Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera)." url="http://www.bioone.org/doi/abs/10.1896/054.063.0107" volumeTitle="A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname." year="2011">Short and Kadosoe 2011</bibRefCitation>
|
||||
<bibRefCitation id="FE3F77D7B6965A668FE200EA8C89480B" author="Short, AEZ" editor="O'Shea, BJ" journalOrPublisher="RAP Bulletin of Biological Assessment 63, Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 90" publicationUrl="http://www.bioone.org/doi/abs/10.1896/054.063.0107" refId="B271" refString="Short, AEZ, Kadosoe, V, 2011. Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera). In: O'Shea, BJ, Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname. RAP Bulletin of Biological Assessment 63, Conservation International, Arlington: 79 - 90, http://www.bioone.org/doi/abs/10.1896/054.063.0107" title="Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera)." url="http://www.bioone.org/doi/abs/10.1896/054.063.0107" volumeTitle="A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname." year="2011">Short and Kadosoe 2011</bibRefCitation>
|
||||
: description of one additional species;
|
||||
<bibRefCitation author="Short, AEZ" editor="Alonso, LE" journalOrPublisher="RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 89" publicationUrl="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" refId="B263" refString="Short, AEZ, 2013. Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera). In: Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname. RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington: 79 - 89, https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" title="Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera)." url="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" volumeTitle="A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname." year="2013">Short 2013</bibRefCitation>
|
||||
<bibRefCitation id="C3DBCC2D6F66A40189661AB5844760E1" author="Short, AEZ" editor="Alonso, LE" journalOrPublisher="RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington" pageId="0" pageNumber="1" pagination="79 - 89" publicationUrl="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" refId="B263" refString="Short, AEZ, 2013. Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera). In: Alonso, LE, Larsen, TH, Eds., A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname. RAP Bulletin of Biological Assessment (Vol. 67). Conservation International, Arlington: 79 - 89, https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" title="Chapter 4. Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera)." url="https://bioone.org/ebooks/RAP-Bulletin-of-Biological-Assessment/A-Rapid-Biological-Assessment-of-the-Upper-Palumeu-River-Watershed/Chapter/Aquatic-Beetles-of-the-Grensgebergte-and-Kasikasima-Regions-Suriname-Insecta/10.1896/054.067.0112" volumeTitle="A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname." year="2013">Short 2013</bibRefCitation>
|
||||
: description of two additional species;
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
|
||||
<bibRefCitation id="B742163AD7ED2CB1B5AC4F6DF5B486F0" DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="113 - 146" refId="B191" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
|
||||
: revision of the genus and description of five new species;
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken originalValue="Girón">Giron</normalizedToken>
|
||||
<bibRefCitation id="B52D609B91591D7B42FA73E87AC6AFF6" DOI="https://doi.org/10.3897/zookeys.1019.59881" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="93 - 140" refId="B133" refString="Giron, JC, Short, AEZ, 2021a. New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys 1019: 93 - 140, DOI: https://doi.org/10.3897/zookeys.1019.59881" title="New subgenera, new species and new records of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.3897/zookeys.1019.59881" volume="1019" year="2021 a">
|
||||
<normalizedToken id="D06DA4D4FA4B76DF9E43F2A4D361875E" originalValue="Girón">Giron</normalizedToken>
|
||||
and Short 2021a
|
||||
</bibRefCitation>
|
||||
: review of the genus with description of 15 new species and establishment of four species groups;
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. 2021</bibRefCitation>
|
||||
<bibRefCitation id="F788ED6A85E1B59935D1979CF7DFCCC1" DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. 2021</bibRefCitation>
|
||||
: phylogenetic placement.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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