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<document ID-DOI="http://dx.doi.org/10.3897/zse.96.55210" ID-GBIF-Dataset="112beb96-2da2-4cc9-8a36-d5ddbfe87288" ID-GBIF-Taxon="170420551" ID-Pensoft-Pub="1860-0743-2-781" ID-Pensoft-UUID="F0701CC577025AD690EE3FDD31F2AEFC" ID-ZooBank="CCDFA5BA6E714D1088087D39DBB92210" LSID="urn:lsid:zoobank.org:act:ABAC0D71-01A5-4E81-A1EB-C037E9888747" ModsDocID="1860-0743-2-781" checkinTime="1605866853493" checkinUser="pensoft" docAuthor="Perger, Robert &amp; Rubio, Gonzalo D." docDate="2020" docId="403C58D3001D5471A00B3B4756A5FED3" docLanguage="en" docName="ZoosystEvol 96(2): 781-795" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.55210" docTitle="Sympolymnia shinahota Perger &amp; Rubio 2020, sp. nov." docType="treatment" docUuid="ABAC0D71-01A5-4E81-A1EB-C037E9888747" docUuidSource="ZooBank" docVersion="3" id="F0701CC577025AD690EE3FDD31F2AEFC" lastPageNumber="781" masterDocId="F0701CC577025AD690EE3FDD31F2AEFC" masterDocTitle="Sympolymnia, a new genus of Neotropical ant-like spider, with description of two new species and indirect evidence for transformational mimicry (Araneae, Salticidae, Simonellini)" masterLastPageNumber="795" masterPageNumber="781" pageId="0" pageNumber="781" updateTime="1643439542219" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Sympolymnia, a new genus of Neotropical ant-like spider, with description of two new species and indirect evidence for transformational mimicry (Araneae, Salticidae, Simonellini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perger, Robert</mods:namePart>
<mods:affiliation>Coleccion Boliviana de Fauna, La Paz, Bolivia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-9930-9638</mods:nameIdentifier>
<mods:nameIdentifier type="email">robertperger@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rubio, Gonzalo D.</mods:namePart>
<mods:affiliation>National Research Council of Argentina (CONICET), Experimental Station of Agriculture (EEA-INTA), R 14, Km 836, Cerro Azul, Misiones, Argentina</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4223-2980</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>781</mods:start>
<mods:end>795</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zse.96.55210</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.55210</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-2-781</mods:identifier>
<mods:identifier type="ZooBank">CCDFA5BA6E714D1088087D39DBB92210</mods:identifier>
<mods:identifier type="Pensoft-UUID">F0701CC577025AD690EE3FDD31F2AEFC</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170420551" LSID="urn:lsid:zoobank.org:act:ABAC0D71-01A5-4E81-A1EB-C037E9888747" httpUri="http://treatment.plazi.org/id/403C58D3001D5471A00B3B4756A5FED3" lastPageNumber="781" pageId="0" pageNumber="781">
<subSubSection pageId="0" pageNumber="781" type="nomenclature">
<paragraph pageId="0" pageNumber="781">
<taxonomicName LSID="403C58D3-001D-5471-A00B-3B4756A5FED3" authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia shinahota" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="shinahota" status="sp. nov.">Sympolymnia shinahota</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="781">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus of A. Synemosyna myrmeciaeformis (Taczanowski, 1871), adult female; B. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; C. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm." figureDoi="10.3897/zse.96.55210.figure3" httpUri="https://binary.pensoft.net/fig/477340" pageId="0" pageNumber="781">Figs 3C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral habitus of A. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; B. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm. Arrows indicate the two light patches between the cephalic and thoracic areas (these patches are translucent white in live specimens)." figureDoi="10.3897/zse.96.55210.figure4" httpUri="https://binary.pensoft.net/fig/477341" pageId="0" pageNumber="781">, 4B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia and chelicerae of Sympolymnia spp.: S. shinahota sp. nov., female holotype (IBSI-Ara 0726): A. Epigyne in ventral view, cleared (blue line course of copulatory ducts); B. Spermathecae, dorsal view; C. Epigyne, ventral view; D. Chelicera in anterior view. Sympolymnia cutleri sp. nov., female holotype (IBSI-Ara 1072): E. Chelicera in anterior view; F. Epigyne in ventral view, cleared (blue line course of copulatory ducts); G. Spermathecae, dorsal view." figureDoi="10.3897/zse.96.55210.figure5" httpUri="https://binary.pensoft.net/fig/477342" pageId="0" pageNumber="781">, 5A-D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live habitus of Sympolymnia spp. and potential ant models. Please note the ontogenetic shift of shine and abdomen shape in the spiders. Sympolymnia shinahota sp. nov.: A. Juvenile female, Villa Tunari, Cochabamba Dept. (please note the shiny, pointed abdomen); B, C. Adult females, same location; D. Holotype female, Buena Vista, Santa Cruz Dept.; E. Pseudomyrmex ethicus, Villa Tunari, Cochabamba Dept.; F. Crematogaster sp., Villa Tunari, Cochabamba Dept.; G. Camponotus sanctaefidei, La Guardia, Santa Cruz Dept.; H. C. latangulus, Buena Vista, Santa Cruz Dept.; Sympolymnia lauretta: I. Juvenile, Bermejo, Santa Cruz Dept. (please note the shiny and pointed abdomen); J. Sub-adult male, Santiago de Chiquitos, Santa Cruz Dept.; K. Adult female, Santa Rosa de la Mina, Santa Cruz Dept.; L. Sympolymnia cutleri sp. nov.: adult female, Chairo, La Paz Dept." figureDoi="10.3897/zse.96.55210.figure8" httpUri="https://binary.pensoft.net/fig/477345" pageId="0" pageNumber="781">, 8A-D</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="type deposit">
<paragraph pageId="0" pageNumber="781">Type deposit.</paragraph>
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" italics="true" pageId="0" pageNumber="781">Holotype</emphasis>
♀; Bolivia: Santa Cruz Department, Buena Vista, Cafetal;
<geoCoordinate degrees="17.4658" direction="south" orientation="latitude" precision="5" value="-17.4658">17.4658°S</geoCoordinate>
,
<geoCoordinate degrees="63.6969" direction="west" orientation="longitude" precision="5" value="-63.6969">63.6969°W</geoCoordinate>
; 342 m a.s.l.; beating tray sampling; 21 Jan 2016; R. Perger leg.; IBSI-Ara 0726. Paratypes 1 ♂; Brazil: Amazonas State, Manaus, Taruma
<normalizedToken originalValue="Mirím">Mirim</normalizedToken>
; 26 Feb 1988; SMNK-ARA 00364 • 2 ♀; Bolivia: Cochabamba Department, Villa Tunari;
<geoCoordinate degrees="16.9844" direction="south" orientation="latitude" precision="5" value="-16.9844">16.9844°S</geoCoordinate>
,
<geoCoordinate degrees="65.4094" direction="west" orientation="longitude" precision="5" value="-65.4094">65.4094°W</geoCoordinate>
; 335 m a.s.l.; beating tray sampling; 6 Dec 2017; R. Perger leg.; IBSI-Ara 1033 • 1 ♀; same data as for preceding; CBF.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="diagnosis">
<paragraph pageId="0" pageNumber="781">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="781">
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia shinahota" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia shinahota</emphasis>
</taxonomicName>
sp. nov. is distinguished from congeners by a distinct dorsal constriction in the basal half of the abdomen (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus of A. Synemosyna myrmeciaeformis (Taczanowski, 1871), adult female; B. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; C. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm." figureDoi="10.3897/zse.96.55210.figure3" httpUri="https://binary.pensoft.net/fig/477340" pageId="0" pageNumber="781">3C</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral habitus of A. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; B. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm. Arrows indicate the two light patches between the cephalic and thoracic areas (these patches are translucent white in live specimens)." figureDoi="10.3897/zse.96.55210.figure4" httpUri="https://binary.pensoft.net/fig/477341" pageId="0" pageNumber="781">4B</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live habitus of Sympolymnia spp. and potential ant models. Please note the ontogenetic shift of shine and abdomen shape in the spiders. Sympolymnia shinahota sp. nov.: A. Juvenile female, Villa Tunari, Cochabamba Dept. (please note the shiny, pointed abdomen); B, C. Adult females, same location; D. Holotype female, Buena Vista, Santa Cruz Dept.; E. Pseudomyrmex ethicus, Villa Tunari, Cochabamba Dept.; F. Crematogaster sp., Villa Tunari, Cochabamba Dept.; G. Camponotus sanctaefidei, La Guardia, Santa Cruz Dept.; H. C. latangulus, Buena Vista, Santa Cruz Dept.; Sympolymnia lauretta: I. Juvenile, Bermejo, Santa Cruz Dept. (please note the shiny and pointed abdomen); J. Sub-adult male, Santiago de Chiquitos, Santa Cruz Dept.; K. Adult female, Santa Rosa de la Mina, Santa Cruz Dept.; L. Sympolymnia cutleri sp. nov.: adult female, Chairo, La Paz Dept." figureDoi="10.3897/zse.96.55210.figure8" httpUri="https://binary.pensoft.net/fig/477345" pageId="0" pageNumber="781">8A-C</figureCitation>
); male palp (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">9C-F</figureCitation>
) with small cymbium, bulb about 80% of cymbium length, tibial apophysis in retro-lateral view narrow, apex evenly tapering (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">9C, D</figureCitation>
); copulatory opening in female very small and quadrangular (width: ~ 0.05 mm) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia and chelicerae of Sympolymnia spp.: S. shinahota sp. nov., female holotype (IBSI-Ara 0726): A. Epigyne in ventral view, cleared (blue line course of copulatory ducts); B. Spermathecae, dorsal view; C. Epigyne, ventral view; D. Chelicera in anterior view. Sympolymnia cutleri sp. nov., female holotype (IBSI-Ara 1072): E. Chelicera in anterior view; F. Epigyne in ventral view, cleared (blue line course of copulatory ducts); G. Spermathecae, dorsal view." figureDoi="10.3897/zse.96.55210.figure5" httpUri="https://binary.pensoft.net/fig/477342" pageId="0" pageNumber="781">5C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.55210.figure4" httpUri="https://binary.pensoft.net/fig/477341" pageId="0" pageNumber="781" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">Figure 4.</emphasis>
Lateral habitus of
<emphasis bold="true" pageId="0" pageNumber="781">A.</emphasis>
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" baseAuthorityName="Peckham &amp; Peckham" baseAuthorityYear="1892" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia lauretta" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="lauretta">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia lauretta</emphasis>
</taxonomicName>
(Peckham &amp; Peckham, 1892), adult female;
<emphasis bold="true" pageId="0" pageNumber="781">B.</emphasis>
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm. Arrows indicate the two light patches between the cephalic and thoracic areas (these patches are translucent white in live specimens).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="comparisons">
<paragraph pageId="0" pageNumber="781">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="781">
The male palp of
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">9C-F</figureCitation>
) resembles that of
<taxonomicName genus="S." lsidName="S. lucasi." pageId="0" pageNumber="781" rank="species" species="lucasi.">
<emphasis italics="true" pageId="0" pageNumber="781">S. lucasi.</emphasis>
</taxonomicName>
However, the latter is distinguished by a male palp tibial apophysis with a spine-like apex (evenly tapering in
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov.), the thoracic part narrower than the cephalic region in dorsal view and distinctly concave posteriorly in lateral view and the epigyne opening transversely elliptical. In the other two species with known males, the tibial apophysis of the male palp is either broader (
<taxonomicName genus="S." lsidName="S. lauretta" pageId="0" pageNumber="781" rank="species" species="lauretta">
<emphasis italics="true" pageId="0" pageNumber="781">S. lauretta</emphasis>
</taxonomicName>
) or narrower (
<taxonomicName genus="S." lsidName="S. edwardsi" pageId="0" pageNumber="781" rank="species" species="edwardsi">
<emphasis italics="true" pageId="0" pageNumber="781">S. edwardsi</emphasis>
</taxonomicName>
) than in
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov. In all four congeners, the abdomen is not or only indistinctly constricted anteriorly.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="description">
<paragraph pageId="0" pageNumber="781">Description.</paragraph>
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">
Female
<emphasis italics="true" pageId="0" pageNumber="781">holotype</emphasis>
</emphasis>
(Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus of A. Synemosyna myrmeciaeformis (Taczanowski, 1871), adult female; B. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; C. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm." figureDoi="10.3897/zse.96.55210.figure3" httpUri="https://binary.pensoft.net/fig/477340" pageId="0" pageNumber="781">3C</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral habitus of A. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; B. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm. Arrows indicate the two light patches between the cephalic and thoracic areas (these patches are translucent white in live specimens)." figureDoi="10.3897/zse.96.55210.figure4" httpUri="https://binary.pensoft.net/fig/477341" pageId="0" pageNumber="781">4B</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia and chelicerae of Sympolymnia spp.: S. shinahota sp. nov., female holotype (IBSI-Ara 0726): A. Epigyne in ventral view, cleared (blue line course of copulatory ducts); B. Spermathecae, dorsal view; C. Epigyne, ventral view; D. Chelicera in anterior view. Sympolymnia cutleri sp. nov., female holotype (IBSI-Ara 1072): E. Chelicera in anterior view; F. Epigyne in ventral view, cleared (blue line course of copulatory ducts); G. Spermathecae, dorsal view." figureDoi="10.3897/zse.96.55210.figure5" httpUri="https://binary.pensoft.net/fig/477342" pageId="0" pageNumber="781">5A-D</figureCitation>
). Total length: 4.25 mm. Carapace length: 2.27 mm; width: 0.90 mm. Integument slightly shiny, orange yellowish with dark bands around eyes of last three rows, dorsum with sparse, simple, moderately long, whitish-yellow setae, denser and longer on anterior half of cephalic area and posterior half of abdomen; without pubescence. Carapace slender and elongated, cephalic portion a little longer than wide (width: 0.85 mm), as wide as widest thoracic part, smooth, marked constriction (width 0.52 mm) between cephalic and thoracic part, two translucent white areas at each side of constriction, separated by narrow black area (imitating part of femora I), constriction followed by globular, somewhat wrinkled knob, which is terminated behind by short pedicle which is more slender than the anterior constriction, evenly tapering when seen in lateral view and concave in dorsal view. Eyes arranged in four rows, quadrangle formed by the second and fourth rows of eyes wider than long, anterior eyes large, in contact, occupying entire front of vertical inclination of face; second pair placed on back behind eyes of first pair, but a little further from each other than distance between external borders of first ones, directed sideways; posterior eyes larger, separated by the same distance as those of second row, eyes of the third pair halfway between second and fourth. Chelicerae light brown, with three teeth on promargin and two on retromargin. Anterior half of sternum yellow and posterior half light brown. Abdomen length: 1.80 mm; width: 1.10 mm, of same length as carapace, broader, commencing by short pedicle that appears to constitute prolongation of that of thoracic part, dorsally completely covered by scutum, integument smooth, constriction in anterior portion. Slender and comparably long legs, in order 4, 3, 1, 2, third and fourth pair stouter, light brown. Epigyne (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia and chelicerae of Sympolymnia spp.: S. shinahota sp. nov., female holotype (IBSI-Ara 0726): A. Epigyne in ventral view, cleared (blue line course of copulatory ducts); B. Spermathecae, dorsal view; C. Epigyne, ventral view; D. Chelicera in anterior view. Sympolymnia cutleri sp. nov., female holotype (IBSI-Ara 1072): E. Chelicera in anterior view; F. Epigyne in ventral view, cleared (blue line course of copulatory ducts); G. Spermathecae, dorsal view." figureDoi="10.3897/zse.96.55210.figure5" httpUri="https://binary.pensoft.net/fig/477342" pageId="0" pageNumber="781">5A-C</figureCitation>
): epigynal plate forming part of the epigastric sclerite, copulatory opening in very small and quadrangular (width: ~ 0.05 mm), copulatory ducts long, starting in a chamber, forming a spiral with one loop and entering the spermathecae posteriorly; spermatheca lung-shaped; copulatory ducts anterior to spermathecae.
</paragraph>
<caption doi="10.3897/zse.96.55210.figure5" httpUri="https://binary.pensoft.net/fig/477342" pageId="0" pageNumber="781" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">Figure 5.</emphasis>
Genitalia and chelicerae of
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia</emphasis>
</taxonomicName>
spp.:
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov., female holotype (IBSI-Ara 0726):
<emphasis bold="true" pageId="0" pageNumber="781">A.</emphasis>
Epigyne in ventral view, cleared (blue line course of copulatory ducts);
<emphasis bold="true" pageId="0" pageNumber="781">B.</emphasis>
Spermathecae, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="781">C</emphasis>
. Epigyne, ventral view;
<emphasis bold="true" pageId="0" pageNumber="781">D.</emphasis>
Chelicera in anterior view.
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia cutleri" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="cutleri">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia cutleri</emphasis>
</taxonomicName>
sp. nov., female holotype (IBSI-Ara 1072):
<emphasis bold="true" pageId="0" pageNumber="781">E.</emphasis>
Chelicera in anterior view;
<emphasis bold="true" pageId="0" pageNumber="781">F.</emphasis>
Epigyne in ventral view, cleared (blue line course of copulatory ducts);
<emphasis bold="true" pageId="0" pageNumber="781">G.</emphasis>
Spermathecae, dorsal view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">
Male
<emphasis italics="true" pageId="0" pageNumber="781">paratype</emphasis>
</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">9</figureCitation>
). Total length: 3.81 mm. Carapace length: 1.94 mm; width: 0.74 mm. Cephalic portion a little longer than wide (width: 0.71 mm), thoracic constriction 0.50 mm wide. General form of carapace, chelicerae and sternum as female, colouration dark brown. Abdomen length: 1.70 mm; width: 0.83 mm, covered dorsally and completely by a scutum, with a conspicuous constriction in the anterior portion; colouration dark brown. Palp (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sympolymnia shinahota sp. nov., paratype male (SMNK-ARA: 00364): A. Dorsal view; B. Lateral view; palp; C. Retrolateral; D. Retroventral; E. Ventral; F. Prolateral (photographs by Hubert Hoefer, SMNK, Germany)." figureDoi="10.3897/zse.96.55210.figure9" httpUri="https://binary.pensoft.net/fig/477346" pageId="0" pageNumber="781">9C-F</figureCitation>
): small cymbium, bulb about 80% of cymbium length, irregularly shaped spherical, embolus long, arising at the basal side of the bulb, without complete circular revolution and is accommodated on a slight retro-lateral concavity of cymbium, lacking pars pendula, tibial apophysis tooth-like, narrow, moderately tapering, tip directed forward.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="variation">
<paragraph pageId="0" pageNumber="781">Variation.</paragraph>
<paragraph pageId="0" pageNumber="781">
Juvenile females had a shiny, dark brown-blackish body surface with dark orange to light brown cephalic part and a pointed abdomen (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live habitus of Sympolymnia spp. and potential ant models. Please note the ontogenetic shift of shine and abdomen shape in the spiders. Sympolymnia shinahota sp. nov.: A. Juvenile female, Villa Tunari, Cochabamba Dept. (please note the shiny, pointed abdomen); B, C. Adult females, same location; D. Holotype female, Buena Vista, Santa Cruz Dept.; E. Pseudomyrmex ethicus, Villa Tunari, Cochabamba Dept.; F. Crematogaster sp., Villa Tunari, Cochabamba Dept.; G. Camponotus sanctaefidei, La Guardia, Santa Cruz Dept.; H. C. latangulus, Buena Vista, Santa Cruz Dept.; Sympolymnia lauretta: I. Juvenile, Bermejo, Santa Cruz Dept. (please note the shiny and pointed abdomen); J. Sub-adult male, Santiago de Chiquitos, Santa Cruz Dept.; K. Adult female, Santa Rosa de la Mina, Santa Cruz Dept.; L. Sympolymnia cutleri sp. nov.: adult female, Chairo, La Paz Dept." figureDoi="10.3897/zse.96.55210.figure8" httpUri="https://binary.pensoft.net/fig/477345" pageId="0" pageNumber="781">8A</figureCitation>
). The constriction in the proximal half of the abdomen was marked by a relatively broad, light transverse band. Adult females had a matt, blackish body surface with a dark orange or completely black cephalic part, the bright band in the abdominal constriction indistinct or absent (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live habitus of Sympolymnia spp. and potential ant models. Please note the ontogenetic shift of shine and abdomen shape in the spiders. Sympolymnia shinahota sp. nov.: A. Juvenile female, Villa Tunari, Cochabamba Dept. (please note the shiny, pointed abdomen); B, C. Adult females, same location; D. Holotype female, Buena Vista, Santa Cruz Dept.; E. Pseudomyrmex ethicus, Villa Tunari, Cochabamba Dept.; F. Crematogaster sp., Villa Tunari, Cochabamba Dept.; G. Camponotus sanctaefidei, La Guardia, Santa Cruz Dept.; H. C. latangulus, Buena Vista, Santa Cruz Dept.; Sympolymnia lauretta: I. Juvenile, Bermejo, Santa Cruz Dept. (please note the shiny and pointed abdomen); J. Sub-adult male, Santiago de Chiquitos, Santa Cruz Dept.; K. Adult female, Santa Rosa de la Mina, Santa Cruz Dept.; L. Sympolymnia cutleri sp. nov.: adult female, Chairo, La Paz Dept." figureDoi="10.3897/zse.96.55210.figure8" httpUri="https://binary.pensoft.net/fig/477345" pageId="0" pageNumber="781">8B, C</figureCitation>
). The holotype (BL 4.25 mm), collected in Cafetal, Buena Vista (Santa Cruz Department), had a matt, orange yellowish body except for some darker patches around the second and third eyes and the abdominal apex (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus of A. Synemosyna myrmeciaeformis (Taczanowski, 1871), adult female; B. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; C. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm." figureDoi="10.3897/zse.96.55210.figure3" httpUri="https://binary.pensoft.net/fig/477340" pageId="0" pageNumber="781">3C</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral habitus of A. Sympolymnia lauretta (Peckham &amp; Peckham, 1892), adult female; B. S. shinahota sp. nov. (IBSI-Ara 0726), female holotype. Scale bars: 1 cm. Arrows indicate the two light patches between the cephalic and thoracic areas (these patches are translucent white in live specimens)." figureDoi="10.3897/zse.96.55210.figure4" httpUri="https://binary.pensoft.net/fig/477341" pageId="0" pageNumber="781">4B</figureCitation>
). The abdominal constriction becomes less pronounced with increasing body length.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="etymology">
<paragraph pageId="0" pageNumber="781">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="781">
The specific epithet,
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="shinahota" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="shinahota">shinahota</taxonomicName>
, refers to a place with many ants or an ant nest in the
<normalizedToken originalValue="Yuracaré">Yuracare</normalizedToken>
language (
<bibRefCitation author="Querejazu Lewis, R" journalOrPublisher="Biota Neotropica" pageId="0" pageNumber="781" refId="B45" refString="Querejazu Lewis, R, 2005. La Cultura de los Yuracares su Habitat y su Proceso de Cambio. Cochabamba UMSS / ASDI. SAREC." title="La Cultura de los Yuracares su Habitat y su Proceso de Cambio. Cochabamba UMSS / ASDI. SAREC." year="2005">Querejazu 2005</bibRefCitation>
), spoken by the
<normalizedToken originalValue="Yuracaré">Yuracare</normalizedToken>
people living along the Chapare River in the Amazon Basin of Bolivia.
</paragraph>
<caption doi="10.3897/zse.96.55210.figure6" httpUri="https://binary.pensoft.net/fig/477343" pageId="0" pageNumber="781" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">Figure 6.</emphasis>
Holotype of
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" baseAuthorityName="Taczanowski" baseAuthorityYear="1871" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia lucasi" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="lucasi">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia lucasi</emphasis>
</taxonomicName>
(Taczanowski, 1871):
<emphasis bold="true" pageId="0" pageNumber="781">A.</emphasis>
Habitus dorsal and lateral views;
<emphasis bold="true" pageId="0" pageNumber="781">B.</emphasis>
Spermatheca and copulatory duct;
<emphasis bold="true" pageId="0" pageNumber="781">C.</emphasis>
Data label.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="distribution">
<paragraph pageId="0" pageNumber="781">
Geographical and ecological distribution (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ecoregion distribution: &quot; Sympolymnia lucasi (Taczanowski, 1871); 3 / 4 S. cutleri sp. nov.; · S. shinahota sp. nov.; l S. lauretta (Peckham &amp; Peckham, 1892)." figureDoi="10.3897/zse.96.55210.figure7" httpUri="https://binary.pensoft.net/fig/477344" pageId="0" pageNumber="781">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="781">
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia shinahota" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia shinahota</emphasis>
</taxonomicName>
sp. nov. is known from Brazil, Amazonas State, Manaus and from the Bolivian Departments of Cochabamba (Villa Tunari) and Santa Cruz (Buena Vista). The collection locations of this species were situated in moist Amazonas forest regions, including Uatuma-Trombetas moist forest (Amazonas State, Manaus) and pre-Andean Southwest Amazon rainforest (Villa Tunari, Cochabamba Dept. and Buena Vista, Santa Cruz Dept.). However, in all areas that were surveyed in the present study,
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov. was exclusively collected in early successional forests in large tree-fall gaps or secondary forest from isolated, small trees that were densely overgrown with climbing plants, several metres away from the edge of primary forest.
</paragraph>
<caption doi="10.3897/zse.96.55210.figure7" httpUri="https://binary.pensoft.net/fig/477344" pageId="0" pageNumber="781" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="781">
<emphasis bold="true" pageId="0" pageNumber="781">Figure 7.</emphasis>
Ecoregion distribution:&quot;
<taxonomicName authority="Perger &amp; Rubio, 2020" authorityName="Perger &amp; Rubio" authorityYear="2020" baseAuthorityName="Taczanowski" baseAuthorityYear="1871" class="Arachnida" family="Salticidae" genus="Sympolymnia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sympolymnia lucasi" order="Araneae" pageId="0" pageNumber="781" phylum="Arthropoda" rank="species" species="lucasi">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia lucasi</emphasis>
</taxonomicName>
(Taczanowski, 1871);
<normalizedToken originalValue="¾">3/4</normalizedToken>
<taxonomicName genus="S." lsidName="S. cutleri" pageId="0" pageNumber="781" rank="species" species="cutleri">
<emphasis italics="true" pageId="0" pageNumber="781">S. cutleri</emphasis>
</taxonomicName>
sp. nov.; ·
<taxonomicName genus="S." lsidName="S. shinahota" pageId="0" pageNumber="781" rank="species" species="shinahota">
<emphasis italics="true" pageId="0" pageNumber="781">S. shinahota</emphasis>
</taxonomicName>
sp. nov.; l
<taxonomicName genus="S." lsidName="S. lauretta" pageId="0" pageNumber="781" rank="species" species="lauretta">
<emphasis italics="true" pageId="0" pageNumber="781">S. lauretta</emphasis>
</taxonomicName>
(Peckham &amp; Peckham, 1892).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="781" type="remarks">
<paragraph pageId="0" pageNumber="781">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="781">
Photographs of two individuals (
<bibRefCitation author="Maddison, WP" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="781" publicationUrl="http://salticidae.org/salticidImages/pages/ecuador/images174.html" refId="B23" refString="Maddison, WP, 2018. Salticid Images (Ecuador). http://salticidae.org/salticidImages/pages/ecuador/images174.html" title="Salticid Images (Ecuador)." url="http://salticidae.org/salticidImages/pages/ecuador/images174.html" year="2018">Maddison 2018</bibRefCitation>
) suggest that this species also occurs in the Amazon rainforest in Ecuador (Yasuni National Park). Light variants were also reported in
<taxonomicName genus="S." lsidName="S. lauretta" pageId="0" pageNumber="781" rank="species" species="lauretta">
<emphasis italics="true" pageId="0" pageNumber="781">S. lauretta</emphasis>
</taxonomicName>
and
<taxonomicName genus="S." lsidName="S. lucasi" pageId="0" pageNumber="781" rank="species" species="lucasi">
<emphasis italics="true" pageId="0" pageNumber="781">S. lucasi</emphasis>
</taxonomicName>
(
<bibRefCitation author="Galiano, ME" journalOrPublisher="Revista de la Sociedad Argentina de Ciencias Naturales" pageId="0" pageNumber="781" pagination="27 - 39" refId="B14" refString="Galiano, ME, 1967. Salticidae (Araneae) formiciformes. VIII. Nuevas descripciones. Physis. Revista de la Sociedad Argentina de Ciencias Naturales 27: 27 - 39" title="Salticidae (Araneae) formiciformes. VIII. Nuevas descripciones. Physis." volume="27" year="1967">Galiano 1967</bibRefCitation>
). However, these reports refer to specimens that were stored in alcohol or formaldehyde for a longer time and may have faded due to the preservative (
<bibRefCitation author="Galiano, ME" journalOrPublisher="Revista de la Sociedad Argentina de Ciencias Naturales" pageId="0" pageNumber="781" pagination="27 - 39" refId="B14" refString="Galiano, ME, 1967. Salticidae (Araneae) formiciformes. VIII. Nuevas descripciones. Physis. Revista de la Sociedad Argentina de Ciencias Naturales 27: 27 - 39" title="Salticidae (Araneae) formiciformes. VIII. Nuevas descripciones. Physis." volume="27" year="1967">Galiano 1967</bibRefCitation>
). This study reports for the first time an in situ observation of an orange variant of
<taxonomicName genus="Sympolymnia." lsidName="Sympolymnia." pageId="0" pageNumber="781" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="781">Sympolymnia.</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="8D73CD4FC5EB2149F4A1389D12CA9CBB" pageId="null" pageNumber="131">
<taxonomicName id="80D85C01175D3F649A15CC593DE30709" authority="Poir." authorityName="Poir." class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="null" pageNumber="131" phylum="Tracheophyta" rank="species" species="olympica">
Stachys
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Poir.
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<paragraph id="063C5E362A0ACE18B2807BA12C7124C6" pageId="null" pageNumber="131">
(
<taxonomicName id="FB40D47340843B1B2A00B795EF238F9E" authority="K. Koch" authorityName="K. Koch" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="null" pageNumber="131" phylum="Tracheophyta" rank="species" species="byzantina">
<emphasis id="722A939FC7B814079FFA15039C5A2E14" italics="true" pageId="null" pageNumber="131">S. byzantina</emphasis>
K. Koch
</taxonomicName>
,
<taxonomicName id="F35EE47BD6C175D2CBF4061F2685B5D4" authority="Jacquin" authorityName="Jacquin" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="null" pageNumber="131" phylum="Tracheophyta" rank="species" species="lanata">
<emphasis id="DDE6DB7F74F6A79E6A25EFCA6A2613C7" italics="true" pageId="null" pageNumber="131">S. lanata</emphasis>
Jacquin
</taxonomicName>
)
</paragraph>
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<subSubSection id="F245C6C87DFDD65218A914E3C15544CB" pageId="null" pageNumber="131" type="vernacular_names">
<paragraph id="8398E12742D8510554D6D1014B53C11C" pageId="null" pageNumber="131">Wolliger Ziest</paragraph>
</subSubSection>
</title>
<subSubSection id="F2861A761263F808C8C292CB0040197B" pageId="null" pageNumber="131" type="description">
<paragraph id="5894E783E8FB1CE150500C8BD2537570" pageId="null" pageNumber="131">
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wie
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<emphasis id="5E5ED70558FDE641AD7B9A379129740D" italics="true" pageId="null" pageNumber="131">S. germanica</emphasis>
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(Nr. 7), aber
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(
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die Epidermis der Stengel und
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unter der
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Behaarung nicht mehr sichtbar
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).
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am Grunde in den Stiel
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</emphasis>
, nicht
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, sehr fein und stumpf
<normalizedToken id="8CB976AE3637C9F755EBAED61C01628E" originalValue="gezähnt" pageId="null" pageNumber="131">gezaehnt</normalizedToken>
oder fast ganzrandig, auffallend runzelig,
<emphasis id="33E4DA390EC5DFE1D1EDDA53DA0625BF" italics="true" pageId="null" pageNumber="131">
<normalizedToken id="178D44F9C4E8341CAB6631C640AB27BE" originalValue="weiß" pageId="null" pageNumber="131">weiss</normalizedToken>
</emphasis>
. -
<normalizedToken id="22BFE27F10A5AC67D03DAC7F63D26EA1" originalValue="Blüte" pageId="null" pageNumber="131">Bluete</normalizedToken>
: Sommer.
</paragraph>
<paragraph id="5C2A198F7E55D0A96A77340FEC5B6CCB" pageId="null" pageNumber="131">
<emphasis id="399CC6A08580DC3378DAD19218D5E3DB" italics="true" pageId="null" pageNumber="131">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="67A8099C93BFEE246A04331211CA7726" italics="true" pageId="null" pageNumber="131">30:</emphasis>
Material aus botanischen
<normalizedToken id="99CD140D0F126FE7EAF5663212AF41E8" originalValue="Gärten" pageId="null" pageNumber="131">Gaerten</normalizedToken>
(Lang 1940), aus der Sowjetunion (Chuksanova und Kaplanbekova 1971).
</paragraph>
<paragraph id="5AD7270E886BF50271614BB331E7756A" pageId="null" pageNumber="131">
<emphasis id="6060BAAEF4644BA1094A7588C35A7BBB" italics="true" pageId="null" pageNumber="131">Standort</emphasis>
. Kollin. Lockere, stickstoffreiche, steinige
<normalizedToken id="979D9A99C558318D94B6D7806A955BC9" originalValue="Böden" pageId="null" pageNumber="131">Boeden</normalizedToken>
in warmen Lagen.
<normalizedToken id="FCD89E372F642588FB82F0D2F27223EB" originalValue="Schuttplätze" pageId="null" pageNumber="131">Schuttplaetze</normalizedToken>
.
</paragraph>
<paragraph id="5252849CB3FB1942C0A2DAF4E9BAF036" pageId="null" pageNumber="131">
<emphasis id="F66366E350105C4C1151F8A0D549BA21" italics="true" pageId="null" pageNumber="131">Verbreitung. Ostmediterrane Pflanze:</emphasis>
Thrazien, Kleinasien, Krim, Kaukasus, Nordpersien. - Im Gebiet gelegentlich in
<normalizedToken id="6C1025EE4D15F4C4AA09D5E7819D7F5B" originalValue="Gärten" pageId="null" pageNumber="131">Gaerten</normalizedToken>
als Zierpflanze kultiviert und selten verwildert (so z. B. bei La Sarraz und Vaumarcus im
<normalizedToken id="A480328DC236BEB5DC3C28B29ED975D2" originalValue="Waadtländer" pageId="null" pageNumber="131">Waadtlaender</normalizedToken>
und Neuenburger Jura, bei Mandello am Comersee).
</paragraph>
</subSubSection>
</treatment>
</document>