Add updates up until 2025-04-17 17:57:56
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<mods:title id="4FDBFA18D566CCA96937245437F2D5B5">Designation of lectotypes for some Spanish and other western European Melitaea taxa, some with mixed syntypic series of M. phoebe ([Denis & amp; Schiffermüller], 1775) and M. ornata Christoph, 1893 (Lepidoptera: Nymphalidae)</mods:title>
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<mods:nameIdentifier id="F566694D269F0E2A6BDE00334B8D98BF" type="email">peterjcrussell@yahoo.co.uk</mods:nameIdentifier>
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<mods:namePart id="5169905CD75CE2A7951A6163A23518F7">Bartolozzi, L.</mods:namePart>
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<mods:affiliation id="E2E2A471A52E1F7A22BC1DB6E513E028">Natural History Museum of the University of Florence Zoology Department ‘ La Specola’ via Romana, 17 I- 50125 Florence ITALIA / ITALY</mods:affiliation>
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<mods:namePart id="A29B43205FD912171255B4A3E81659B4">Hawkins, R. L.</mods:namePart>
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<mods:affiliation id="063A9BB0F0BCBF3EC7EB5EB63A2400ED">Museum of Comparative Zoology Harvard University 26 Oxford Street Cambridge MA, 02138 EE. UU. / USA</mods:affiliation>
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<mods:namePart id="B60CD96A192194470B6E3F281AD8B912">Tennent, W. J.</mods:namePart>
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<mods:affiliation id="27ABE1750FF775864AA5C26BA01723B1">Department of Life Sciences The Natural History Museum GB-London SW 7 5 BD REINO UNIDO / UNITED KINGDOM</mods:affiliation>
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<mods:namePart id="5100E6D422849D5B736BFF854CBCD7C8">Léger, T.</mods:namePart>
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<mods:affiliation id="4DB949525E38E9E19793CA1EF241328F">Museum fuer Naturkunde Leibniz-Institut fuer Evolutions und Biodiversitaetsforschung Invalidenstrasser, 43 D- 10115 Berlin ALEMANIA / GERMANY</mods:affiliation>
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<mods:nameIdentifier id="B5A31409AA41A4F2A42968EB05C10274" type="email">theo.leger@mfn.berlin</mods:nameIdentifier>
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<mods:part id="3EF109975BE20228C6092851B373C1C5">
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<mods:date id="D7D9E36015EA681F522399881F24D757">2020</mods:date>
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<mods:number id="1BFEB647CF26E38C0D0499AC8A7777E6">2020-09-30</mods:number>
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<mods:number id="596684CABB75025F9276E92176E4440A">48</mods:number>
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<mods:url id="87D084216E38D4CA86DA31CCB9118239">https://doi.org/10.57065/shilap.369</mods:url>
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<treatment id="03ECBD0540510B28FF277BB3FC9268CD" ID-DOI="http://doi.org/10.5281/zenodo.15237490" ID-Zenodo-Dep="15237490" LSID="urn:lsid:plazi:treatment:03ECBD0540510B28FF277BB3FC9268CD" httpUri="http://treatment.plazi.org/id/03ECBD0540510B28FF277BB3FC9268CD" lastPageNumber="453" pageId="4" pageNumber="453">
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<subSubSection id="C35F5F9840510B28FF277BB3FDCE6999" box="[179,571,1180,1202]" pageId="4" pageNumber="453" type="nomenclature">
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<paragraph id="8BFA0C1340510B28FF277BB3FDCE6999" blockId="4.[130,1128,201,1621]" box="[179,571,1180,1202]" pageId="4" pageNumber="453">
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<taxonomicName id="4C45779040510B28FF277BB3FDCE6999" ID-CoL="3ZKFM" authority="Turati 1920" authorityName="Turati" authorityYear="1920" box="[179,571,1180,1202]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="rostagnoi">
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<emphasis id="B931D00140510B28FF277BB3FE4B699A" box="[179,446,1181,1202]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe rostagnoi</emphasis>
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Turati 1920
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F5F9840510B28FDD07BB2FD926815" pageId="4" pageNumber="453" type="description">
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<paragraph id="8BFA0C1340510B28FDD07BB2FD926815" blockId="4.[130,1128,201,1621]" pageId="4" pageNumber="453">
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(223 and Tav. II, figs 10-12) [TL: Italy, Roma, Monte Autore]; from the figure 10, the male forewing apices appear distinctly acute as in
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<taxonomicName id="4C45779040510B28FC607B96FB9469E5" box="[1012,1121,1208,1229]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140510B28FC607B96FB9469E5" box="[1012,1121,1208,1229]" italics="true" pageId="4" pageNumber="453">M. phoebe</emphasis>
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</taxonomicName>
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. NEKRUTENKO (1993: 129) listed the Type material of Turati in the Museo Regionale di Scienze Naturali, Torino, Italy, (MRSN) and referring to this subspecies stated that there were two male and three female syntypes from Monte Autore, Province of Rome, Italy; he identified them as
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<taxonomicName id="4C45779040510B28FB867A22FF326815" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140510B28FB867A22FF326815" italics="true" pageId="4" pageNumber="453">Melitaea phoebe</emphasis>
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</taxonomicName>
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, a conclusion with which we cannot argue.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F5F9840510B28FF277A6AFC9268CD" pageId="4" pageNumber="453" type="discussion">
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<paragraph id="8BFA0C1340510B28FF277A6AFC9268CD" blockId="4.[130,1128,201,1621]" pageId="4" pageNumber="453">
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In view of the current long-term closure of the museum we cannot be sure of their identity, but
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<taxonomicName id="4C45779040510B28FBD97A6BFF32685E" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140510B28FBD97A6BFF32685E" italics="true" pageId="4" pageNumber="453">M. phoebe</emphasis>
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</taxonomicName>
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seems most likely. No specimen of
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<taxonomicName id="4C45779040510B28FDA77A4FFD63685E" authorityName="Christoph" authorityYear="1893" box="[563,662,1377,1398]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140510B28FDA77A4FFD63685E" box="[563,662,1377,1398]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
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</taxonomicName>
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has been observed in peninsular Italy north of San Marco Catola, Foggia, Apulia, Italy (c. 41º 30’ N.) (CAGNETTA 2016: 246). Similarly the aberration
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<taxonomicName id="4C45779040510B28FF7E7AB6FE246885" authority="Turati, 1921" authorityName="Turati" authorityYear="1921" box="[234,465,1432,1453]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="sterlineata">
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<emphasis id="B931D00140510B28FF7E7AB6FEA46885" box="[234,337,1432,1453]" italics="true" pageId="4" pageNumber="453">sterlineata</emphasis>
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Turati, 1921
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</taxonomicName>
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: (
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<figureCitation id="137E109640510B28FE777AB6FDDF6885" box="[483,554,1432,1453]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="4" pageNumber="453">Fig. 12</figureCitation>
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), with an almost unmarked discal area of the forewing and from the same locality, is most likely
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<taxonomicName id="4C45779040510B28FD937A9AFD8568E1" box="[519,624,1460,1481]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140510B28FD937A9AFD8568E1" box="[519,624,1460,1481]" italics="true" pageId="4" pageNumber="453">M. phoebe</emphasis>
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</taxonomicName>
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; VAN OORSCHOT & COUTSIS (2014: 61) and RUSSELL & TENNENT (2016: 52, note 75) agreed with this determination.
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</paragraph>
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</treatment>
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<mods:title id="C828175E3C4FBD8C4E53551FFBEE0B62">SHILAP Revista de lepidopterología</mods:title>
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<taxonomicName id="4C45779040510B28FF277DA7FDC76FB6" ID-CoL="72Y6M" authority="Turati, 1920" authorityName="Turati" authorityYear="1920" box="[179,562,649,670]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="phoebina">
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<emphasis id="B931D00140510B28FF277DA7FE416FB6" box="[179,436,649,670]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe phoebina</emphasis>
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Turati, 1920
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</taxonomicName>
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[TL: Italy, Calabria, Aspromonte, 1400 m]; this form was described on page 222 and the uppersides figured on tav. II figs 4 11 and 5 00, from which it appears to be a fairly heavily marked form, hence Verity’s suggestion that the ab.
|
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<taxonomicName id="4C45779040510B28FCC67DEFFC706FFE" authorityName="Stauder" authorityYear="1914" box="[850,901,705,726]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="totila">
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<emphasis id="B931D00140510B28FCC67DEFFC706FFE" box="[850,901,705,726]" italics="true" pageId="4" pageNumber="453">totila</emphasis>
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</taxonomicName>
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of Stauder (see above) equated to this subspecies. The undersides were not figured by Turati but the forewing apices appear rounded as in
|
||||
<taxonomicName id="4C45779040510B28FE9B7DD7FE876E26" authorityName="Christoph" authorityYear="1893" box="[271,370,761,782]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140510B28FE9B7DD7FE876E26" box="[271,370,761,782]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
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</taxonomicName>
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. According to CONCI & POGGI (1996) the collection of E. Turati is in the Museo Regionale di Scienze Naturali, Torino, Italy, (MRSN). NEKRUTENKO (1993: 129) listed the type material of Turati in this museum and referring to this subspecies stated that there were two female syntypes from the Aspromonte Mountains in the Province of Reggio Calabria, Italy and identified them as
|
||||
<taxonomicName id="4C45779040510B28FF087C46FEC96E55" box="[156,316,872,893]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
|
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<emphasis id="B931D00140510B28FF087C46FEC96E55" box="[156,316,872,893]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe</emphasis>
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|
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, being unaware of the specific separation of
|
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<taxonomicName id="4C45779040510B28FD7F7C46FCBE6E55" authorityName="Christoph" authorityYear="1893" box="[747,843,872,893]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
|
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<emphasis id="B931D00140510B28FD7F7C46FCBE6E55" box="[747,843,872,893]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
|
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</taxonomicName>
|
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.
|
||||
</paragraph>
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|
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<subSubSection id="C35F5F9840510B28FF277CABFDED69BD" pageId="4" pageNumber="453" type="discussion">
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Unfortunately, the entomology collections in the Turin Museum are not currently accessible for administrative reasons and it has not been possible to inspect type material. However, it is possible to make a speculative identification, based on current knowledge of the distributions of the two species. Based on the collecting locality, this subspecies is almost certainly
|
||||
<taxonomicName id="4C45779040510B28FCA67CF7FC626EC6" authorityName="Christoph" authorityYear="1893" box="[818,919,985,1006]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140510B28FCA67CF7FC626EC6" box="[818,919,985,1006]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
|
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</taxonomicName>
|
||||
as
|
||||
<taxonomicName id="4C45779040510B28FC2A7CF7FBDD6EC6" box="[958,1064,985,1006]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
|
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<emphasis id="B931D00140510B28FC2A7CF7FBDD6EC6" box="[958,1064,985,1006]" italics="true" pageId="4" pageNumber="453">M. phoebe</emphasis>
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</taxonomicName>
|
||||
sensu strictu has not been recorded this far south in peninsular Italy; its limit appears to be Calabria, Cosenza, Monte Martinelli (RUSSELL & PATEMAN, 2011), where
|
||||
<taxonomicName id="4C45779040510B28FD5E7B3EFCD8690D" authorityName="Christoph" authorityYear="1893" box="[714,813,1040,1061]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
|
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<emphasis id="B931D00140510B28FD5E7B3EFCD8690D" box="[714,813,1040,1061]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
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</taxonomicName>
|
||||
also occurs. Six males and one female of this subspecies are present in the Rothschild collection in the Natural History Museum London (Russell and Tennent, pers obs.). This taxon was not considered by VAN OORSCHOT & COUTSIS (2014) but RUSSELL & TENNENT (2016: 51, note 67) suggested it was
|
||||
<taxonomicName id="4C45779040510B28FC217B4AFBE06951" authorityName="Christoph" authorityYear="1893" box="[949,1045,1124,1145]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
|
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<emphasis id="B931D00140510B28FC217B4AFBE06951" box="[949,1045,1124,1145]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
because of its location in southern peninsular Italy.
|
||||
</paragraph>
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129
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<taxonomicName id="4C45779040510B28FF277E2FFDCF6C3E" ID-CoL="5JM9Q" authority="Verity 1919" authorityName="Verity" authorityYear="1919" box="[179,570,257,278]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="emipunica">
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<emphasis id="B931D00140510B28FF277E2FFE346C3E" box="[179,449,257,278]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe emipunica</emphasis>
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Verity 1919
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[TL: Italy, Sicily, Palermo]: there are four male and one female specimens present in the Museo Zoologico de “La Specola” dell’Università, Firenza, Italy (cf. RUSSELL & BARTOLOZZI, 2019:
|
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<figureCitation id="137E109640510B28FE677E17FDD86C66" box="[499,557,313,334]" captionStart="Figs 1-5.– 1" captionStartId="15.[157,195,1424,1443]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-146@15.[136,1134,203,1619]" captionTargetPageId="15" captionText="Figs 1-5.– 1. Melitaea phoebe male underside, Republic of North Macedonia, Skopje, Kriva Palanka, c. 650 m, 9 June 2013. © P. Russell. 2. Melitaea ornata male underside, Republic of North Macedonia, Skopje, Suva Planina, c. 1000 m, 8 June 2013. © P. Russell. 3. Melitaea phoebe female underside, Italy, Calabria, Cosenza, San Fili, Monte Martinelli, 880 m, 10 June 2007. © P. Russell. 4. Melitaea ornata female underside, Italy, Calabria, Cosenza, San Fili, Monte Martinelli, 600 m, 3 June 1913 (Stauder Leg.) © W. J. Tennent. 5A, B & C. Melitaea phoebe occitanica Staudinger, 1871, lectotype male Barcelona, Spain. © T. Léger. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="4" pageNumber="453">Fig. 1</figureCitation>
|
||||
). The male, which was figured by VERITY (1950: 152; Tav. 43: fig. 63) and given more accurate locality data: “San Martino della Scala m. 800 (Monreale Palermo); 5 V”, is here designated as the lectotype of
|
||||
<taxonomicName id="4C45779040510B28FD0A7E5FFC586CAE" authorityName="Verity" authorityYear="1919" box="[670,941,369,390]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="emipunica">
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<emphasis id="B931D00140510B28FD0A7E5FFC586CAE" box="[670,941,369,390]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe emipunica</emphasis>
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</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E109640510B28FC287E5FFBDA6CAE" box="[956,1071,369,390]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="4" pageNumber="453">Figs 12a, b</figureCitation>
|
||||
). The squat triangular submarginal black markings barely touching the black intervening veins (
|
||||
<figureCitation id="137E109640510B28FB9E7EA3FB956C8A" box="[1034,1120,397,418]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="4" pageNumber="453">Fig. 12b</figureCitation>
|
||||
) clearly place the taxon
|
||||
<taxonomicName id="4C45779040510B28FEFE7E87FE3B6C96" authorityName="Verity" authorityYear="1919" box="[362,462,425,446]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="emipunica">
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<emphasis id="B931D00140510B28FEFE7E87FE3B6C96" box="[362,462,425,446]" italics="true" pageId="4" pageNumber="453">emipunica</emphasis>
|
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</taxonomicName>
|
||||
with
|
||||
<taxonomicName id="4C45779040510B28FD9D7E87FD996C96" authorityName="Christoph" authorityYear="1893" box="[521,620,425,446]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140510B28FD9D7E87FD996C96" box="[521,620,425,446]" italics="true" pageId="4" pageNumber="453">M. ornata</emphasis>
|
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</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="4C45779040510B28FDEE7E87FD176C96" box="[634,738,425,446]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140510B28FDEE7E87FD176C96" box="[634,738,425,446]" italics="true" pageId="4" pageNumber="453">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
has not so far been proven to occur in Sicily (RUSSELL, 2018: 258). The labels on the pin (
|
||||
<figureCitation id="137E109640510B28FD0E7EEBFD196CF2" box="[666,748,453,474]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="4" pageNumber="453">Fig. 12c</figureCitation>
|
||||
) are as follows: black print on yellow paper “Monreale (San Martino)m. 800/ PALERMO ISOLA di SICILIA/ 6 Maggio 1919 Querci”; black print on white paper “Ex coll. R. Verity”; black print on pink paper “Syntypus”; printed on yellow paper “FIGURATO DA/ R. VERITY FARF. D. IT./ TAV. 43 (hand-written)/ FIG. 63 (hand-written)”; Black print on red paper
|
||||
<emphasis id="B931D00140510B28FEE67D1BFDEF6F62" box="[370,538,565,586]" italics="true" pageId="4" pageNumber="453">
|
||||
“
|
||||
<taxonomicName id="4C45779040510B28FEED7D1BFDEF6F62" authorityName="Christoph" authorityYear="1893" box="[377,538,565,586]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="ornata">Melitaea ornata</taxonomicName>
|
||||
</emphasis>
|
||||
/
|
||||
<taxonomicName id="4C45779040510B28FDB37D1BFCF46F61" authority="Verity 1919" authorityName="Verity" authorityYear="1919" box="[551,769,564,586]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="emipunica">
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||||
<emphasis id="B931D00140510B28FDB37D1BFD7E6F62" box="[551,651,565,586]" italics="true" pageId="4" pageNumber="453">emipunica</emphasis>
|
||||
Verity 1919
|
||||
</taxonomicName>
|
||||
/ LECTOTYPUS/ P. RUSSELL & L. BARTOLOZZI 2019”. Both VAN OORSCHOT & COUTSIS (2014) and RUSSELL & TENNENT (2016) agreed with this determination.
|
||||
</paragraph>
|
||||
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|
||||
163
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163
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</mods:mods>
|
||||
<treatment id="03ECBD0540510B29FF277AC3FB9F6EB1" ID-DOI="http://doi.org/10.5281/zenodo.15237492" ID-Zenodo-Dep="15237492" LSID="urn:lsid:plazi:treatment:03ECBD0540510B29FF277AC3FB9F6EB1" httpUri="http://treatment.plazi.org/id/03ECBD0540510B29FF277AC3FB9F6EB1" lastPageId="5" lastPageNumber="454" pageId="4" pageNumber="453">
|
||||
<subSubSection id="C35F5F9840510B28FF277AC3FDBB6B2A" box="[179,590,1517,1538]" pageId="4" pageNumber="453" type="nomenclature">
|
||||
<paragraph id="8BFA0C1340510B28FF277AC3FDBB6B2A" blockId="4.[130,1128,201,1621]" box="[179,590,1517,1538]" pageId="4" pageNumber="453">
|
||||
<taxonomicName id="4C45779040510B28FF277AC3FDBB6B2A" ID-CoL="6QXTZ" authority="Ragusa, 1921" authorityName="Ragusa" authorityYear="1921" box="[179,590,1517,1538]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="punicata">
|
||||
<emphasis id="B931D00140510B28FF277AC3FE496B2A" box="[179,444,1517,1538]" italics="true" pageId="4" pageNumber="453">Melitaea phoebe punicata</emphasis>
|
||||
Ragusa, 1921
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F9840510B29FDCC7AC3FDEA6F39" lastPageId="5" lastPageNumber="454" pageId="4" pageNumber="453" type="distribution">
|
||||
<paragraph id="8BFA0C1340510B29FDCC7AC3FDEA6F39" blockId="4.[130,1128,201,1621]" lastBlockId="5.[136,1135,201,1621]" lastPageId="5" lastPageNumber="454" pageId="4" pageNumber="453">
|
||||
[TL: Italy, Sicily, Palermo District]. Apparently the Sicilian Macrolepidoptera collection of É. E. Ragusa was sold to Walter, Lord Rothschild (HORN
|
||||
<emphasis id="B931D00140510B28FBC27927FF6E6B12" italics="true" pageId="4" pageNumber="453">et al.</emphasis>
|
||||
, 1990). There are nine males and five females in his collection at NHM, London, UK. (Russell and Tennent, pers. obs.); no indication that the material was syntypic was present on any of the data labels. According to GREGORI (1926) another part of the butterfly collection of Énrico Ragusa was in the Instituto di Zoologia, Universita di Napoli, Portici, Italy (ZIUP); at our request Roberta Improta made a thorough search of the Naples Museum collections but was unable to find any of Ragusa’s
|
||||
<taxonomicName id="4C45779040500B29FB937E2FFB9B6C3E" box="[1031,1134,257,278]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FB937E2FFB9B6C3E" box="[1031,1134,257,278]" italics="true" pageId="5" pageNumber="454">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
specimens (see Acknowledgements). From the NHM specimens, which are clearly
|
||||
<taxonomicName id="4C45779040500B29FC287E33FBEB6C1A" authorityName="Christoph" authorityYear="1893" box="[956,1054,285,306]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140500B29FC287E33FBEB6C1A" box="[956,1054,285,306]" italics="true" pageId="5" pageNumber="454">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
, a male captured by Ragusa in the District of Palermo is herewith designated as a lectotype. The specific characters of
|
||||
<taxonomicName id="4C45779040500B29FE9E7E7BFEBF6C42" authorityName="Christoph" authorityYear="1893" box="[266,330,341,362]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140500B29FE9E7E7BFEBF6C42" box="[266,330,341,362]" italics="true" pageId="5" pageNumber="454">ornata</emphasis>
|
||||
</taxonomicName>
|
||||
are clearly visible in the photographs of the lectotype (
|
||||
<figureCitation id="137E109640500B29FCF57E7AFC256C41" box="[865,976,340,361]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="5" pageNumber="454">Figs 13a, b</figureCitation>
|
||||
).
|
||||
<taxonomicName id="4C45779040500B29FC767E7BFBB26C42" box="[994,1095,341,362]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FC767E7BFBB26C42" box="[994,1095,341,362]" italics="true" pageId="5" pageNumber="454">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
has not been reported from Sicily (RUSSELL, 2018). The labels on the pin of the lectotype (
|
||||
<figureCitation id="137E109640500B29FC647E5EFBB46CAD" box="[1008,1089,368,389]" captionStart="Figs 12-13.– 12" captionStartId="19.[157,195,1475,1494]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-112@19.[136,1134,203,1619]" captionTargetPageId="19" captionText="Figs 12-13.– 12A, B & C. Melitaea phoebe emipunica Verity, 1919, male lectotype, designated Russell & Bartolozzi, 2019, Italy, Sicily, Palermo, Monreale, 800 m, 6 May 1918, Querci leg., misidentified M. ornata. © S. Bambi. 13A, B & C. Melitaea phoebe punicata Ragusa, 1921, male lectotype, designated Russell & Tennent, 2019, Italy, Palermo, Valle Corta, 10 May, [1]917, Ragusa leg., misidentified M. ornata. © W. J. Tennent. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="5" pageNumber="454">Fig. 13c</figureCitation>
|
||||
) are as follows: on beige paper, part printed part handwritten “Prov. Palermo/ Local. V[all] Corta/ Data 10.5.[1]917/ Coll. E. Ragusa”; on beige paper, printed “Sicily,/ coll. E. Ragusa”; on purple-bordered circular white paper printed “LECTO-/TYPE”; on beige paper printed “Rothschild/ Bequest/ B.M. 1939-1.”; on white paper printed “LECTOTYPE/
|
||||
<taxonomicName id="4C45779040500B29FD117ECEFCD96CDD" box="[645,812,480,501]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FD117ECEFCD96CDD" box="[645,812,480,501]" italics="true" pageId="5" pageNumber="454">Melitaea phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
forma/
|
||||
<taxonomicName id="4C45779040500B29FCEB7ECEFB936CDD" authority="Ragusa, 1919" authorityName="Ragusa" authorityYear="1919" box="[895,1126,480,501]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="punicata">
|
||||
<emphasis id="B931D00140500B29FCEB7ECEFC206CDD" box="[895,981,480,501]" italics="true" pageId="5" pageNumber="454">punicata</emphasis>
|
||||
Ragusa, 1919
|
||||
</taxonomicName>
|
||||
/ Designated by Russell/ & Tennent, 2019”.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F9840500B29FF2D7D37FB9F6EB1" pageId="5" pageNumber="454" type="discussion">
|
||||
<paragraph id="8BFA0C1340500B29FF2D7D37FB9F6EB1" blockId="5.[136,1135,201,1621]" pageId="5" pageNumber="454">
|
||||
We here designate the remaining 13 syntypes as paralectotypes, of which six have the following labels: on beige paper, printed “Sicilien”/ hand written “Ficuzza/ 5”/ printed “Geo.C.Kr.”; on beige paper printed “Sicily/ coll. E. Ragusa”; on beige paper printed “Rothschild/ Bequest/ B.M.1939-1.”; white circle with blue surround printed “PARA-/LECTO-/ TYPE”; on white paper printed “PARALECTOTYPE/
|
||||
<taxonomicName id="4C45779040500B29FEFB7DA6FDED6FB5" box="[367,536,648,669]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FEFB7DA6FDED6FB5" box="[367,536,648,669]" italics="true" pageId="5" pageNumber="454">Melitaea phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
forma/
|
||||
<taxonomicName id="4C45779040500B29FDF97DA6FCA06FB5" authority="Ragusa, 1919" authorityName="Ragusa" authorityYear="1919" box="[621,853,648,669]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="punicata">
|
||||
<emphasis id="B931D00140500B29FDF97DA6FD316FB5" box="[621,708,648,669]" italics="true" pageId="5" pageNumber="454">punicata</emphasis>
|
||||
Ragusa, 1919
|
||||
</taxonomicName>
|
||||
/ Designated by Russell/ & Tennent, 2019”, 4 have the following labels: on beige paper with black surround handwritten: “M. Cuccio/ 30.4.[1]916”; on beige paper printed: “Sicily/ coll. E. Ragusa”; on beige paper printed “Rothschild/ Bequest/ B.M.1939-1.”; white circle with blue surround printed “PARA-/LECTO-/ TYPE”; on white paper printed “PARALECTOTYPE/
|
||||
<taxonomicName id="4C45779040500B29FD0F7DD6FCC96E25" box="[667,828,760,781]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FD0F7DD6FCC96E25" box="[667,828,760,781]" italics="true" pageId="5" pageNumber="454">Melitaea phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
forma/
|
||||
<taxonomicName id="4C45779040500B29FC1D7DD6FB936E25" authority="Ragusa, 1919" authorityName="Ragusa" authorityYear="1919" box="[905,1126,760,781]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="punicata">
|
||||
<emphasis id="B931D00140500B29FC1D7DD6FC286E25" box="[905,989,760,781]" italics="true" pageId="5" pageNumber="454">punicata</emphasis>
|
||||
Ragusa, 1919
|
||||
</taxonomicName>
|
||||
/ Designated by Russell/ & Tennent, 2019” and finally three have the following labels: on beige paper printed: “Sicily/ coll. E. Ragusa”; on beige paper printed “Rothschild/ Bequest/ B.M.1939-1.”; white circle with blue surround printed “PARA-/LECTO-/ TYPE”; on white paper printed “PARALECTOTYPE/
|
||||
<taxonomicName id="4C45779040500B29FEFB7C46FDED6E55" box="[367,536,872,893]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140500B29FEFB7C46FDED6E55" box="[367,536,872,893]" italics="true" pageId="5" pageNumber="454">Melitaea phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
forma/
|
||||
<taxonomicName id="4C45779040500B29FDF97C46FCA06E55" authority="Ragusa, 1919" authorityName="Ragusa" authorityYear="1919" box="[621,853,872,893]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="punicata">
|
||||
<emphasis id="B931D00140500B29FDF97C46FD316E55" box="[621,708,872,893]" italics="true" pageId="5" pageNumber="454">punicata</emphasis>
|
||||
Ragusa, 1919
|
||||
</taxonomicName>
|
||||
/ Designated by Russell/ & Tennent, 2019”. We here formally identify all 14 specimens as
|
||||
<taxonomicName id="4C45779040500B29FD727CAAFB936EB1" authority="Ragusa, 1921" authorityName="Ragusa" authorityYear="1921" box="[742,1126,900,921]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="punicata">
|
||||
<emphasis id="B931D00140500B29FD727CAAFC2E6EB1" box="[742,987,900,921]" italics="true" pageId="5" pageNumber="454">Melitaea ornata punicata</emphasis>
|
||||
Ragusa, 1921
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
144
data/03/EC/BD/03ECBD0540520B2BFF2D7B03FBD768CD.xml
Normal file
144
data/03/EC/BD/03ECBD0540520B2BFF2D7B03FBD768CD.xml
Normal file
|
|
@ -0,0 +1,144 @@
|
|||
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<mods:mods id="C6C59AF3D552EA3F55A3B3E6058CF244" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="084F31CD33B0047306DBCBA357849DAC">
|
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<mods:title id="3DD9A295BAFB1BF2361351CF023C6DD5">Designation of lectotypes for some Spanish and other western European Melitaea taxa, some with mixed syntypic series of M. phoebe ([Denis & amp; Schiffermüller], 1775) and M. ornata Christoph, 1893 (Lepidoptera: Nymphalidae)</mods:title>
|
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</mods:titleInfo>
|
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<mods:name id="EA5C5A4E2F9221A1042F512DF389C3DD" type="personal">
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<mods:namePart id="976432304527DEF9A154E972B1A681AF">Russell, P. J. C.</mods:namePart>
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<mods:affiliation id="70CB5CFDAB9CAC4D015D9C8BC5314A8F">Oakmeadow Wessex Avenue, East Wittering GB-West Sussex PO 20 8 NP REINO UNIDO / UNITED KINGDOM</mods:affiliation>
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<mods:nameIdentifier id="38A4B8927438FF97CD16330153471A30" type="email">peterjcrussell@yahoo.co.uk</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="9D55B1BA8AEED6B4AE01360325945E2A">Bartolozzi, L.</mods:namePart>
|
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<mods:nameIdentifier id="9EC01E8DB58AF52C09333E4A0F4B8FF4" type="ORCID">0000-0002-9119-2540</mods:nameIdentifier>
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<mods:affiliation id="329B941E4BCA2E641B86CFA8995C702D">Natural History Museum of the University of Florence Zoology Department ‘ La Specola’ via Romana, 17 I- 50125 Florence ITALIA / ITALY</mods:affiliation>
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<mods:nameIdentifier id="DF5D44B4F7DC188F29B0C16E631BFFC3" type="email">luca.bartolozzi@unifi.it</mods:nameIdentifier>
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</mods:name>
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</mods:role>
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<mods:namePart id="14F7D9FDCEE6B3F8BD4AF99C5F460F59">Hawkins, R. L.</mods:namePart>
|
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<mods:nameIdentifier id="6CD08449D39C78D1DEFBBEBE6E6723B2" type="ORCID">0000-0002-2225-1084</mods:nameIdentifier>
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<mods:affiliation id="9867A50DABBF5992B9C3CA13F8F1AE27">Museum of Comparative Zoology Harvard University 26 Oxford Street Cambridge MA, 02138 EE. UU. / USA</mods:affiliation>
|
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<mods:nameIdentifier id="1CC74B6E131F0CA8CB83F8FB1F79F621" type="email">rhawkins@fas.harvard.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="0E78F89AC199991AB76F217B09E44A37" type="personal">
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<mods:role id="CA8EE59A9C57645D0C6476C91A789AA6">
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<mods:roleTerm id="678E6E92BF4E48262E7E5855747596AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5B25425A12354F657E0FD0F53BB08396">Tennent, W. J.</mods:namePart>
|
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<mods:nameIdentifier id="BDCE0CD7828AC2AD464357207CC96146" type="ORCID">000-0001-7096-4946</mods:nameIdentifier>
|
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<mods:affiliation id="C9F31186EA511DB435FDD2663FDEB5BE">Department of Life Sciences The Natural History Museum GB-London SW 7 5 BD REINO UNIDO / UNITED KINGDOM</mods:affiliation>
|
||||
<mods:nameIdentifier id="B0299318219E60B925D18587E1DC85D8" type="email">johntennent@hotmail.co.uk</mods:nameIdentifier>
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</mods:name>
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<mods:name id="9E7AF416B72CF1F4B6103FDE72D19716" type="personal">
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</mods:role>
|
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<mods:namePart id="9D1DEDE1BFB7D524765194E72B2C3020">Léger, T.</mods:namePart>
|
||||
<mods:nameIdentifier id="AA94F6BA07ED1E83B32A89A8C33B1ABA" type="ORCID">0000-0001-7330-3940</mods:nameIdentifier>
|
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<mods:affiliation id="8086FA531D001D81A4505A0D7A2F87FD">Museum fuer Naturkunde Leibniz-Institut fuer Evolutions und Biodiversitaetsforschung Invalidenstrasser, 43 D- 10115 Berlin ALEMANIA / GERMANY</mods:affiliation>
|
||||
<mods:nameIdentifier id="5F39A3A562E32B2359ECBA424356621C" type="email">theo.leger@mfn.berlin</mods:nameIdentifier>
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<mods:title id="62C2F20E9634130B6D966E39A0F987B7">SHILAP Revista de lepidopterología</mods:title>
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|
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<mods:part id="E02ACF0894F29EC6E12709B7112701E5">
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<mods:date id="617F3BA365DEB33AC8D6BFB143F49BC6">2020</mods:date>
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<treatment id="03ECBD0540520B2BFF2D7B03FBD768CD" ID-DOI="http://doi.org/10.5281/zenodo.15237504" ID-Zenodo-Dep="15237504" LSID="urn:lsid:plazi:treatment:03ECBD0540520B2BFF2D7B03FBD768CD" httpUri="http://treatment.plazi.org/id/03ECBD0540520B2BFF2D7B03FBD768CD" lastPageNumber="456" pageId="7" pageNumber="456">
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<subSubSection id="C35F5F9840520B2BFF2D7B03FD406969" box="[185,693,1068,1090]" pageId="7" pageNumber="456" type="nomenclature">
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<paragraph id="8BFA0C1340520B2BFF2D7B03FD406969" blockId="7.[136,1135,201,1621]" box="[185,693,1068,1090]" pageId="7" pageNumber="456">
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<taxonomicName id="4C45779040520B2BFF2D7B03FD406969" ID-CoL="7KBT5" authority="Verity, 1938" authorityName="Fruhstorfer" authorityYear="1908" box="[185,693,1068,1090]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ogygia">
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<emphasis id="B931D00140520B2BFF2D7B03FDD9696A" box="[185,556,1069,1090]" italics="true" pageId="7" pageNumber="456">Melitaea phoebe ogygia postogygia</emphasis>
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Verity, 1938
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F5F9840520B2BFD2A7B02FBAA6809" pageId="7" pageNumber="456" type="distribution">
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<paragraph id="8BFA0C1340520B2BFD2A7B02FBAA6809" blockId="7.[136,1135,201,1621]" pageId="7" pageNumber="456">
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[TL: uncertain - three syntypes from two different localities in Greece: Salonica (= Thessalonica), Macedonia @ 1000 ft. (=
|
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<emphasis id="B931D00140520B2BFC577B67FC066976" box="[963,1011,1097,1118]" italics="true" pageId="7" pageNumber="456">circa</emphasis>
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300 m) and Olympus, bordering Thessaly/Macedonia @ 2500 ft. (=
|
||||
<emphasis id="B931D00140520B2BFD3B7B4AFD2A6951" box="[687,735,1124,1145]" italics="true" pageId="7" pageNumber="456">circa</emphasis>
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750 m)]. Although the name
|
||||
<taxonomicName id="4C45779040520B2BFB917B4AFB9B6951" authorityName="Verity" authorityYear="1938" box="[1029,1134,1124,1145]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="postogygia">
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<emphasis id="B931D00140520B2BFB917B4AFB9B6951" box="[1029,1134,1124,1145]" italics="true" pageId="7" pageNumber="456">postogygia</emphasis>
|
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</taxonomicName>
|
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has no formal nomenclatural standing as part of a quadrinomial, it is considered here because of its association with the names
|
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<taxonomicName id="4C45779040520B2BFE007BB2FE206999" authorityName="Fruhstorfer" authorityYear="1908" box="[404,469,1180,1201]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="ogygia">
|
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<emphasis id="B931D00140520B2BFE007BB2FE206999" box="[404,469,1180,1201]" italics="true" pageId="7" pageNumber="456">ogygia</emphasis>
|
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</taxonomicName>
|
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and
|
||||
<taxonomicName id="4C45779040520B2BFD917BB2FD986999" authorityName="Verity" authorityYear="1939" box="[517,621,1180,1201]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="nigrogygia">
|
||||
<emphasis id="B931D00140520B2BFD917BB2FD986999" box="[517,621,1180,1201]" italics="true" pageId="7" pageNumber="456">nigrogygia</emphasis>
|
||||
</taxonomicName>
|
||||
, which are associated with
|
||||
<taxonomicName id="4C45779040520B2BFCE87BB2FC286999" authorityName="Christoph" authorityYear="1893" box="[892,989,1180,1201]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140520B2BFCE87BB2FC286999" box="[892,989,1180,1201]" italics="true" pageId="7" pageNumber="456">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(cf above, and RUSSELL & BARTOLOZZI, 2019). Verity’s description (1938: (16)) indicated that the name was proposed for a second generation of “
|
||||
<taxonomicName id="4C45779040520B2BFD997BFAFD3169C1" authorityName="Fruhstorfer" authorityYear="1908" box="[525,708,1236,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ogygia">
|
||||
<emphasis id="B931D00140520B2BFD997BFAFD3169C1" box="[525,708,1236,1257]" italics="true" pageId="7" pageNumber="456">M. phoebe ogygia</emphasis>
|
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</taxonomicName>
|
||||
”. This is unusual:
|
||||
<taxonomicName id="4C45779040520B2BFC1E7BFAFC1A69C1" authorityName="Christoph" authorityYear="1893" box="[906,1007,1236,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="ornata">
|
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<emphasis id="B931D00140520B2BFC1E7BFAFC1A69C1" box="[906,1007,1236,1257]" italics="true" pageId="7" pageNumber="456">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
is generally univoltine (RUSSELL & PATEMAN, 2011), although second generations occur when rearing the species in the U.K., if the larvae are exposed to very wet conditions (RUSSELL & PATEMAN, 2013).
|
||||
</paragraph>
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</subSubSection>
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<subSubSection id="C35F5F9840520B2BFF2D7A06FBD768CD" pageId="7" pageNumber="456" type="discussion">
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<paragraph id="8BFA0C1340520B2BFF2D7A06FBD768CD" blockId="7.[136,1135,201,1621]" pageId="7" pageNumber="456">
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Examination of the photographs of the three syntypes and their associated labels revealed that the two syntypic males from Salonica, taken 12 and 13 August 1936 are almost certainly
|
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<taxonomicName id="4C45779040520B2BFC477A6BFBC06872" authorityName="Christoph" authorityYear="1893" box="[979,1077,1349,1370]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="ornata">
|
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<emphasis id="B931D00140520B2BFC477A6BFBC06872" box="[979,1077,1349,1370]" italics="true" pageId="7" pageNumber="456">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E109640520B2BFBD77A6AFF29685D" captionStart="Figs 16-17.– 16" captionStartId="21.[157,196,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@21.[136,1134,144,1716]" captionTargetPageId="21" captionText="Figs 16-17.– 16A, B & C. Melitaea phoebe ogygia postogygia Verity, 1938, male syntype, Thessalonika, Greece, misidentified M. ornata. © S. Bambi. 17A, B & C. Melitaea phoebe ogygia postogygia Verity, 1938, female syntype, Olymbos, Greece. © S. Bambi. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="7" pageNumber="456">Figs 16a, b, c</figureCitation>
|
||||
); whereas a female from Olympus taken on “Aug[ust]. 17, 1935”, is
|
||||
<taxonomicName id="4C45779040520B2BFCE27A4EFC29685D" box="[886,988,1376,1397]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140520B2BFCE27A4EFC29685D" box="[886,988,1376,1397]" italics="true" pageId="7" pageNumber="456">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E109640520B2BFC7D7A4EFF6068B9" captionStart="Figs 16-17.– 16" captionStartId="21.[157,196,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@21.[136,1134,144,1716]" captionTargetPageId="21" captionText="Figs 16-17.– 16A, B & C. Melitaea phoebe ogygia postogygia Verity, 1938, male syntype, Thessalonika, Greece, misidentified M. ornata. © S. Bambi. 17A, B & C. Melitaea phoebe ogygia postogygia Verity, 1938, female syntype, Olymbos, Greece. © S. Bambi. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="7" pageNumber="456">Figs 17a, b & c</figureCitation>
|
||||
). All three specimens were captured by Romei. These identifications, admittedly based only on antenna and wing morphology, were agreed by John Coutsis and Jim Pateman. (See Acknowledgements). The designation of a lectotype in this case is not relevant because the name
|
||||
<taxonomicName id="4C45779040520B2BFF1C7AFEFF0468CD" authorityName="Verity" authorityYear="1938" box="[136,241,1488,1509]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="postogygia">
|
||||
<emphasis id="B931D00140520B2BFF1C7AFEFF0468CD" box="[136,241,1488,1509]" italics="true" pageId="7" pageNumber="456">postogygia</emphasis>
|
||||
</taxonomicName>
|
||||
is part of a quadrinomial (infrasubspecific) and thus not nomenclaturally significant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
108
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|
||||
<taxonomicName id="4C45779040530B2AFF277BB3FDC4699A" ID-CoL="5JM9Y" authority="Gaede, 1930" authorityName="Gaede" authorityYear="1930" box="[179,561,1181,1202]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="malvida">
|
||||
<emphasis id="B931D00140530B2AFF277BB3FE59699A" box="[179,428,1181,1202]" italics="true" pageId="6" pageNumber="455">Melitaea phoebe malvida</emphasis>
|
||||
Gaede, 1930
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F9840530B2AFDAE7BB3FEF068CD" pageId="6" pageNumber="455" type="description">
|
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<paragraph id="8BFA0C1340530B2AFDAE7BB3FEF068CD" blockId="6.[130,1129,201,1621]" pageId="6" pageNumber="455">
|
||||
[TL: Bosnia & Herzegovina, Bosnia, Maklen (also spelt Makljen) Pass]; Gaede described (page 207) this subspecies and figured the upperside (Plate 13, d5). Gaede attributed this name to Fruhstorfer, but without a date; like HIGGINS (1941: 340) the present authors were unable to find any original description by Fruhstorfer. This subspecies was not mentioned by VAN OORSCHOT & COUTSIS (2014). The specimen figured and the description by Gaede indicated that the apices of the forewings were quite acute, indicating that this subspecies belongs to
|
||||
<taxonomicName id="4C45779040530B2AFBD97A07FF3D6872" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140530B2AFBD97A07FF3D6872" italics="true" pageId="6" pageNumber="455">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
, as described. Gaede also indicated an association with the subspecies
|
||||
<taxonomicName id="4C45779040530B2AFC347A6BFF41685D" authority="Fruhstorfer, 1917" authorityName="Fruhstorfer" authorityYear="1917" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="narenta">
|
||||
<emphasis id="B931D00140530B2AFC347A6BFC1E6872" box="[928,1003,1349,1370]" italics="true" pageId="6" pageNumber="455">narenta</emphasis>
|
||||
Fruhstorfer, 1917
|
||||
</taxonomicName>
|
||||
, from Herzegovina (see above), also identified here as
|
||||
<taxonomicName id="4C45779040530B2AFD487A4EFCB6685D" box="[732,835,1376,1397]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140530B2AFD487A4EFCB6685D" box="[732,835,1376,1397]" italics="true" pageId="6" pageNumber="455">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
. No indication of the precise location or date of capture was provided, making for difficulties in finding this subspecies at the location given, which has a maximum elevation of 1123 m (Yugoslav Coast, Lascelles, scale 1:300,000, dated 1988/9). Until such time as further specimens become available, this identification requires confirmation.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
108
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Normal file
108
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|
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@ -0,0 +1,108 @@
|
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<mods:affiliation id="5ECB2EBF62B83F078638A45F8782FB0A">Department of Life Sciences The Natural History Museum GB-London SW 7 5 BD REINO UNIDO / UNITED KINGDOM</mods:affiliation>
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<mods:nameIdentifier id="3FECF13CBCA1378E7367097ADDF5D3F7" type="email">johntennent@hotmail.co.uk</mods:nameIdentifier>
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<taxonomicName id="4C45779040530B2AFF277C63FD876E4A" ID-CoL="5JM9S" authority="Verity, 1928" authorityName="Verity" authorityYear="1928" box="[179,626,845,866]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="galliaemontium">
|
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<emphasis id="B931D00140530B2AFF277C63FE016E4A" box="[179,500,845,866]" italics="true" pageId="6" pageNumber="455">Melitaea phoebe galliaemontium</emphasis>
|
||||
Verity, 1928
|
||||
</taxonomicName>
|
||||
</paragraph>
|
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</subSubSection>
|
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<subSubSection id="C35F5F9840530B2AFDEE7C63FEB869BD" pageId="6" pageNumber="455" type="distribution">
|
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<paragraph id="8BFA0C1340530B2AFDEE7C63FEB869BD" blockId="6.[130,1129,201,1621]" pageId="6" pageNumber="455">
|
||||
[TL: Mont-Dore, Puy-de-Dôme, France]. This is a name given by Verity to an unnamed
|
||||
<taxonomicName id="4C45779040530B2AFE6F7C47FD7D6E56" box="[507,648,873,894]" pageId="6" pageNumber="455" rank="subSpecies" subSpecies="described">race described</taxonomicName>
|
||||
but not named by FRUHSTORFER (1918: 42). This was a small
|
||||
<taxonomicName id="4C45779040530B2AFEB97CABFE726EB2" box="[301,391,901,922]" pageId="6" pageNumber="455" rank="subSpecies" subSpecies="with">race with</taxonomicName>
|
||||
part of the forewings and all of hindwings covered in a greenish suffusion. HIGGINS (1941: 340) mistakenly gave the description of these specimens as being covered in black suffusion; this actually applied to the previous description of
|
||||
<taxonomicName id="4C45779040530B2AFC927C93FBF36EFA" authority="Verity, 1928" authorityName="Verity" authorityYear="1928" box="[774,1030,957,978]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="crassenigra">
|
||||
<emphasis id="B931D00140530B2AFC927C93FC886EFA" box="[774,893,957,978]" italics="true" pageId="6" pageNumber="455">crassenigra</emphasis>
|
||||
Verity, 1928
|
||||
</taxonomicName>
|
||||
, given to specimens from Gironde, Lozère and Pyrénées Orientales (VERITY, 1928: 162). There were no specimens of this subspecies extant in Verity’s collection in Florence in the early 1980’s (KUDRNA, 1983) and thus no further comment can be made; the name was included here simply to correct the description given by Higgins and to confirm that the problem of identity is insoluble until further samples are collected from the Mont-Dore area. This taxon was not considered by VAN OORSCHOT & COUTSIS (2014) but RUSSELL & TENNENT (2016: 47, note 38) considered it to be
|
||||
<taxonomicName id="4C45779040530B2AFC6E7B4AFB946951" box="[1018,1121,1124,1145]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140530B2AFC6E7B4AFB946951" box="[1018,1121,1124,1145]" italics="true" pageId="6" pageNumber="455">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
, based on its location.
|
||||
</paragraph>
|
||||
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|
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||||
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|
||||
175
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Normal file
175
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<mods:namePart id="A4B8A0AB4498069F84AB9843885E2641">Bartolozzi, L.</mods:namePart>
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<mods:affiliation id="DED0DB7E88CC903B683A2F0F86A7E903">Natural History Museum of the University of Florence Zoology Department ‘ La Specola’ via Romana, 17 I- 50125 Florence ITALIA / ITALY</mods:affiliation>
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<mods:nameIdentifier id="7463AC672E1D37579D40E4E8CCF83237" type="email">luca.bartolozzi@unifi.it</mods:nameIdentifier>
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<mods:namePart id="5F57DDF4560776CA28F33A4BA9F22B58">Hawkins, R. L.</mods:namePart>
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<mods:affiliation id="5C73C8B2045E1F2568716F9C7BD49A19">Museum of Comparative Zoology Harvard University 26 Oxford Street Cambridge MA, 02138 EE. UU. / USA</mods:affiliation>
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<mods:nameIdentifier id="E3250D657F1A6ECF0FD6A60002E141B0" type="email">rhawkins@fas.harvard.edu</mods:nameIdentifier>
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<mods:namePart id="8DC2D16810D5B77D53497CB0C42C68D1">Tennent, W. J.</mods:namePart>
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<mods:affiliation id="440D32F678ED9BEC0F7185EA236BE689">Department of Life Sciences The Natural History Museum GB-London SW 7 5 BD REINO UNIDO / UNITED KINGDOM</mods:affiliation>
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<mods:nameIdentifier id="5EDAD3CE00F54F4214F53B3B1CD0900C" type="email">johntennent@hotmail.co.uk</mods:nameIdentifier>
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<mods:namePart id="ED745352ED351F78A9D4E97E6C9CBC88">Léger, T.</mods:namePart>
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<mods:affiliation id="4418F322B612C6D06BBF372052F9ACB5">Museum fuer Naturkunde Leibniz-Institut fuer Evolutions und Biodiversitaetsforschung Invalidenstrasser, 43 D- 10115 Berlin ALEMANIA / GERMANY</mods:affiliation>
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<mods:nameIdentifier id="F6820B8FAF53893C6DA1A5BCC75C95C9" type="email">theo.leger@mfn.berlin</mods:nameIdentifier>
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<mods:typeOfResource id="8AB20151321362656D412FE3AB860814">text</mods:typeOfResource>
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<mods:title id="38D12ACA3A6D27B797D70FAD37BF9054">SHILAP Revista de lepidopterología</mods:title>
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</mods:titleInfo>
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<mods:part id="F48AD5AB3044B426ED2F178434446A76">
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<mods:date id="6D4DB02CA7F6F9C503AB06687D0EFF26">2020</mods:date>
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<mods:number id="A031BDE3C79C36FCE7B8F547C657655E">2020-09-30</mods:number>
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<mods:detail id="8D3A71AEF25F0F4CC28E65790C384588" type="volume">
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<mods:number id="E3342A9DBE93CA8B311E3CD5A2DE5F65">48</mods:number>
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<mods:number id="FB5498B39C978A8690E7D10F3840CC7E">191</mods:number>
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<mods:start id="CE8C3F005E2EE2132763193229047578">449</mods:start>
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<mods:url id="6023445482AE9BE79D8D9B9795FA54C3">https://doi.org/10.57065/shilap.369</mods:url>
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<mods:classification id="0A5D4FFB6C4BC7F5F51B68BDCB3B73A2">journal article</mods:classification>
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<mods:identifier id="4DC08401D20EB27083303F06302DBA65" type="DOI">10.57065/shilap.369</mods:identifier>
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<mods:identifier id="4D59FC23EB4A7F77D95040412A287575" type="Zenodo-Dep">15237535</mods:identifier>
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<treatment id="03ECBD0540560B28FF2D7C69FCA66DD1" ID-DOI="http://doi.org/10.5281/zenodo.15237484" ID-Zenodo-Dep="15237484" LSID="urn:lsid:plazi:treatment:03ECBD0540560B28FF2D7C69FCA66DD1" httpUri="http://treatment.plazi.org/id/03ECBD0540560B28FF2D7C69FCA66DD1" lastPageId="4" lastPageNumber="452" pageId="3" pageNumber="452">
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<subSubSection id="C35F5F9840560B2FFF2D7C69FD956E74" box="[185,608,839,860]" pageId="3" pageNumber="452" type="nomenclature">
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<paragraph id="8BFA0C1340560B2FFF2D7C69FD956E74" blockId="3.[136,1135,201,1621]" box="[185,608,839,860]" pageId="3" pageNumber="452">
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<taxonomicName id="4C45779040560B2FFF2D7C69FD956E74" ID-CoL="7KC4M" authority="Fruhstorfer, 1917" authorityName="Fruhstorfer" authorityYear="1917" box="[185,608,839,860]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="guevara">
|
||||
<emphasis id="B931D00140560B2FFF2D7C69FE446E74" box="[185,433,839,860]" italics="true" pageId="3" pageNumber="452">Melitaea phoebe guevara</emphasis>
|
||||
Fruhstorfer, 1917
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="C35F5F9840560B2FFDF37C69FBAD687A" pageId="3" pageNumber="452" type="description">
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<paragraph id="8BFA0C1340560B2FFDF37C69FBAD687A" blockId="3.[136,1135,201,1621]" pageId="3" pageNumber="452">
|
||||
[TL: Spain, Castilien, (Cuenca mont.)] was described from three males in the “Leonhard collection”. There is a significant statement in the description given by FRUHSTORFER (1917: 19), who said that this subspecies bore a close relationship to both subspecies
|
||||
<taxonomicName id="4C45779040560B2FFF6C7CB5FECC6E98" authorityName="Fruhstorfer" authorityYear="1908" box="[248,313,923,944]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="ogygia">
|
||||
<emphasis id="B931D00140560B2FFF6C7CB5FECC6E98" box="[248,313,923,944]" italics="true" pageId="3" pageNumber="452">ogygia</emphasis>
|
||||
</taxonomicName>
|
||||
from Greece and
|
||||
<taxonomicName id="4C45779040560B2FFE667CB5FDDB6E98" authorityName="Fruhstorfer" authorityYear="1908" box="[498,558,923,944]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="telona">
|
||||
<emphasis id="B931D00140560B2FFE667CB5FDDB6E98" box="[498,558,923,944]" italics="true" pageId="3" pageNumber="452">telona</emphasis>
|
||||
</taxonomicName>
|
||||
from “Palästina” (considered to be near Jerusalem, Israel [HIGGINS, 1941: 335]). Both are morphologically very similar and were considered to be
|
||||
<taxonomicName id="4C45779040560B2FFB987C99FB9B6EE4" authorityName="Christoph" authorityYear="1893" box="[1036,1134,951,972]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140560B2FFB987C99FB9B6EE4" box="[1036,1134,951,972]" italics="true" pageId="3" pageNumber="452">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(RUSSELL & TENNENT, 2016: notes 59 and 91); however, TÓTH
|
||||
<emphasis id="B931D00140560B2FFCAE7CFCFC926ECF" box="[826,871,978,999]" italics="true" pageId="3" pageNumber="452">et al</emphasis>
|
||||
. (2014) suggested that
|
||||
<taxonomicName id="4C45779040560B2FFBC07CFCFF31692B" authorityName="Fruhstorfer" authorityYear="1908" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="telona">
|
||||
<emphasis id="B931D00140560B2FFBC07CFCFF31692B" italics="true" pageId="3" pageNumber="452">M. telona</emphasis>
|
||||
</taxonomicName>
|
||||
may be a fourth species in this group. HIGGINS (1941: 349) suggested a similarity between
|
||||
<taxonomicName id="4C45779040560B2FFF1C7B24FF226937" box="[136,215,1034,1055]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="guevara">
|
||||
<emphasis id="B931D00140560B2FFF1C7B24FF226937" box="[136,215,1034,1055]" italics="true" pageId="3" pageNumber="452">guevara</emphasis>
|
||||
</taxonomicName>
|
||||
and subspecies
|
||||
<taxonomicName id="4C45779040560B2FFEE27B24FE0A6937" authorityName="de Sagarra" authorityYear="1926" box="[374,511,1034,1055]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="bethunebakeri">
|
||||
<emphasis id="B931D00140560B2FFEE27B24FE0A6937" box="[374,511,1034,1055]" italics="true" pageId="3" pageNumber="452">bethunebakeri</emphasis>
|
||||
</taxonomicName>
|
||||
(see below). A “holotype” male and two male “paratypes” (i.e. three syntypes) were recognised by BERNARDI & DE LESSE (1951: 141). HESSELBARTH
|
||||
<emphasis id="B931D00140560B2FFBAF7B08FB9D6913" box="[1083,1128,1062,1083]" italics="true" pageId="3" pageNumber="452">et al</emphasis>
|
||||
. (1995: 1030/1031 stated that they should be considered more correctly as lectotype and paralectotypes, with which the present authors agree. An inspection of the Fruhstorfer collection in MNHNP by RR revealed that there are three male specimens present, two of which have “PARATYPE” labels attached (presumably, since they were the first to mention “paratypes”, by BERNARDI & DE LESSE, 1951: 141) but the third specimen has no “type” label. The question arose: was this third specimen one of the syntypes with the “HOLOTYPE” label missing? A close inspection of the wing and antenna morphology from high quality photographs (
|
||||
<figureCitation id="137E109640560B2FFDAE7BC7FD6B69D6" box="[570,670,1257,1278]" captionStart="Figs 8-9.– 8" captionStartId="17.[157,195,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@17.[136,1134,144,1716]" captionTargetPageId="17" captionText="Figs 8-9.– 8A, B. & C. Melitaea phoebe ogygia Fruhstorfer, 1908, female lectotype, Greece, Poros, 14 June 1909, misidentified M. ornata. © R. Rougeri. 9A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="3" pageNumber="452">Figs 9a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="137E109640560B2FFD3D7BC7FCDB69D6" box="[681,814,1257,1278]" captionStart="Figs 10-11.– 10" captionStartId="18.[150,188,1475,1494]" captionTargetBox="[130,1127,203,1619]" captionTargetId="graphics-91@18.[130,1128,203,1619]" captionTargetPageId="18" captionText="Figs 10-11.– 10A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. 11A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male, ‘non-type specimen’ Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="3" pageNumber="452">10a, b, 11a, b</figureCitation>
|
||||
) indicated that the specimens all belong the same species,
|
||||
<taxonomicName id="4C45779040560B2FFE177A28FE116833" authorityName="Christoph" authorityYear="1893" box="[387,484,1286,1307]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140560B2FFE177A28FE116833" box="[387,484,1286,1307]" italics="true" pageId="3" pageNumber="452">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
, in contrast to VAN OORSCHOT & COUTSIS (2014: 61), based on studies of genitalia, and RUSSELL & TENNENT (2016: 48, note 41), who both considered, prior to the confirmation of the presence of
|
||||
<taxonomicName id="4C45779040560B2FFE4B7A13FDCA687A" authorityName="Christoph" authorityYear="1893" box="[479,575,1341,1362]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D00140560B2FFE4B7A13FDCA687A" box="[479,575,1341,1362]" italics="true" pageId="3" pageNumber="452">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
in Spain, that all Iberian subspecies were of
|
||||
<taxonomicName id="4C45779040560B2FFC797A13FBA7687A" box="[1005,1106,1341,1362]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140560B2FFC797A13FBA7687A" box="[1005,1106,1341,1362]" italics="true" pageId="3" pageNumber="452">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F9840560B28FF2D7A4FFCA66DD1" lastPageId="4" lastPageNumber="453" pageId="3" pageNumber="452" type="discussion">
|
||||
<paragraph id="8BFA0C1340560B28FF2D7A4FFCA66DD1" blockId="3.[136,1135,201,1621]" lastBlockId="4.[130,1128,201,1621]" lastPageId="4" lastPageNumber="453" pageId="3" pageNumber="452">
|
||||
Inspection of the labels (
|
||||
<figureCitation id="137E109640560B2FFE397A4FFE0D685E" box="[429,504,1377,1398]" captionStart="Figs 8-9.– 8" captionStartId="17.[157,195,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@17.[136,1134,144,1716]" captionTargetPageId="17" captionText="Figs 8-9.– 8A, B. & C. Melitaea phoebe ogygia Fruhstorfer, 1908, female lectotype, Greece, Poros, 14 June 1909, misidentified M. ornata. © R. Rougeri. 9A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="3" pageNumber="452">Figs 9c</figureCitation>
|
||||
,
|
||||
<figureCitation id="137E109640560B2FFD907A4FFDA2685E" box="[516,599,1377,1398]" captionStart="Figs 10-11.– 10" captionStartId="18.[150,188,1475,1494]" captionTargetBox="[130,1127,203,1619]" captionTargetId="graphics-91@18.[130,1128,203,1619]" captionTargetPageId="18" captionText="Figs 10-11.– 10A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. 11A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male, ‘non-type specimen’ Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="3" pageNumber="452">10c, 11c</figureCitation>
|
||||
) revealed no indication that any of the specimens had originated from the Leonhard collection. The location labels were similar in all respects, being handwritten as follows: “Castilien [underlined with printed dots]/ Cuenca/ mont./ 1900 Korb”. Conversely, the identification labels, also handwritten, were not all the same: the two specimens with “PARATYPE” labels were handwritten as follows: “
|
||||
<emphasis id="B931D00140560B2FFD117AFFFCBD68CE" box="[645,840,1489,1510]" italics="true" pageId="3" pageNumber="452">
|
||||
<taxonomicName id="4C45779040560B2FFD117AFFFD1268CE" box="[645,743,1489,1510]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">M. phoebe</taxonomicName>
|
||||
/
|
||||
<taxonomicName id="4C45779040560B2FFD6D7AFFFC6268CE" authority="Fruhst." authorityName="Fruhst." box="[761,919,1489,1510]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="guevara">guevara Fruhst.</taxonomicName>
|
||||
</emphasis>
|
||||
”, whereas the label of the “non-type” specimen was written in a different hand as follows:
|
||||
<emphasis id="B931D00140560B2FFCA67AC3FC166B2A" box="[818,995,1517,1538]" italics="true" pageId="3" pageNumber="452">
|
||||
“
|
||||
<taxonomicName id="4C45779040560B2FFCAD7AC3FC166B2A" box="[825,995,1517,1538]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">Melitaea phoebe</taxonomicName>
|
||||
</emphasis>
|
||||
/
|
||||
<taxonomicName id="4C45779040560B2FFC667AC3FF306B35" authority="Frhst." authorityName="Fruhstorfer" authorityYear="1917" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="guevara">
|
||||
ssp.
|
||||
<emphasis id="B931D00140560B2FFB8B7AC3FB9B6B2A" box="[1055,1134,1517,1538]" italics="true" pageId="3" pageNumber="452">guevara</emphasis>
|
||||
Frhst.
|
||||
</taxonomicName>
|
||||
/ 1917 (
|
||||
<emphasis id="B931D00140560B2FFE8F7927FE896B36" box="[283,380,1545,1566]" italics="true" pageId="3" pageNumber="452">Soc. Ent.</emphasis>
|
||||
p. 19)”. Although it is possible that this is the specimen observed by BERNARDI & DE LESSE (1958), it cannot be assumed that this is their presumed ‘holotype’. Since it is possible that the specimen with the “HOLOTYPE” label may turn up in the future, it was considered unwise to designate a lectotype under these circumstances. We here formally identify the three available syntypes as
|
||||
<taxonomicName id="4C45779040510B28FEC67FCBFD046DD2" authority="Fruhstorfer, 1917" authorityName="Fruhstorfer" authorityYear="1917" box="[338,753,229,250]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="ornata" status="comb. n." subSpecies="guevara">
|
||||
<emphasis id="B931D00140510B28FEC67FCBFDB76DD2" box="[338,578,229,250]" italics="true" pageId="4" pageNumber="453">Melitaea ornata guevara</emphasis>
|
||||
Fruhstorfer, 1917
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B931D00140510B28FD6F7FCAFCA66DD1" bold="true" box="[763,851,228,249]" pageId="4" pageNumber="453">
|
||||
<taxonomicNameLabel id="A2026D7A40510B28FD6F7FCAFCA66DD1" box="[763,851,228,249]" pageId="4" pageNumber="453" rank="species">comb. n.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
99
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99
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<taxonomicName id="4C45779040560B2FFF2D7D4EFDA66F5C" ID-CoL="5JM9T" authority="Fruhstorfer, 1917" authorityName="Fruhstorfer" authorityYear="1917" box="[185,595,607,629]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="gerinia">
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<emphasis id="B931D00140560B2FFF2D7D4EFE506F5D" box="[185,421,608,629]" italics="true" pageId="3" pageNumber="452">Melitaea phoebe gerinia</emphasis>
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Fruhstorfer, 1917
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</taxonomicName>
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[TL: Portugal, Lissabon {Lisbon}]; BERNARDI & DE LESSE (1951: 141) reported that there were no specimens of this subspecies in Fruhstorfer’s collection in MNHNP. However, Rodolphe Rougeri found a male specimen there. From photographs of the underside it can be concluded that, despite the somewhat triangular shape of the submarginal black markings, they touched the intervening veins, the antennal tips were club shaped and not spatulate and the forewing apices appeared more acute than rounded. This specimen is correctly identified as
|
||||
<taxonomicName id="4C45779040560B2FFBC07DC5FF386E34" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140560B2FFBC07DC5FF386E34" italics="true" pageId="3" pageNumber="452">M. phoebe</emphasis>
|
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</taxonomicName>
|
||||
, as described by Fruhstorfer and in agreement with VAN OORSCHOT & COUTSIS (2014: 61) and RUSSELL & TENNENT (2016: note 39).
|
||||
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117
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<taxonomicName id="4C45779040570B2EFF277B00FDAC696B" ID-CoL="7KBT5" authority="Fruhstorfer, 1908" authorityName="Fruhstorfer" authorityYear="1908" box="[179,601,1070,1091]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ogygia">
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<emphasis id="B931D00140570B2EFF277B00FE56696B" box="[179,419,1070,1091]" italics="true" pageId="2" pageNumber="451">Melitaea phoebe ogygia</emphasis>
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Fruhstorfer, 1908
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[TL: Greece, Poros Island]; name used by VARGA (1967) for Hungarian populations of what proved later to be
|
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<taxonomicName id="4C45779040570B2EFD4F7B64FCC86977" authorityName="Christoph" authorityYear="1893" box="[731,829,1098,1119]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140570B2EFD4F7B64FCC86977" box="[731,829,1098,1119]" italics="true" pageId="2" pageNumber="451">M. ornata</emphasis>
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</taxonomicName>
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. According to BERNARDI & DE LESSE (1951: 140), a single female “holotype” is present in the Museum National d’Histoire Naturelle de Paris (MNHNP) (
|
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<figureCitation id="137E109640570B2EFE3B7BAFFDE069BE" box="[431,533,1153,1174]" captionStart="Figs 8-9.– 8" captionStartId="17.[157,195,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@17.[136,1134,144,1716]" captionTargetPageId="17" captionText="Figs 8-9.– 8A, B. & C. Melitaea phoebe ogygia Fruhstorfer, 1908, female lectotype, Greece, Poros, 14 June 1909, misidentified M. ornata. © R. Rougeri. 9A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="2" pageNumber="451">Figs 8a, b</figureCitation>
|
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). Fruhstorfer did not routinely designate holotypes but since there is only one specimen it can be regarded as the holotype. From the photograph (
|
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<figureCitation id="137E109640570B2EFC417BB3FBE8699A" box="[981,1053,1181,1202]" captionStart="Figs 8-9.– 8" captionStartId="17.[157,195,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@17.[136,1134,144,1716]" captionTargetPageId="17" captionText="Figs 8-9.– 8A, B. & C. Melitaea phoebe ogygia Fruhstorfer, 1908, female lectotype, Greece, Poros, 14 June 1909, misidentified M. ornata. © R. Rougeri. 9A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="2" pageNumber="451">Fig. 8b</figureCitation>
|
||||
) of the underside of this specimen, it can be seen that it has centrally thickened triangular black submarginal markings not touching the intervening veins on both fore- and hindwings and also foreshortened tips to the antenna (spatulate). Thus
|
||||
<taxonomicName id="4C45779040570B2EFE237BDFFE0E682E" authorityName="Fruhstorfer" authorityYear="1908" box="[439,507,1265,1286]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="ogygia">
|
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<emphasis id="B931D00140570B2EFE237BDFFE0E682E" box="[439,507,1265,1286]" italics="true" pageId="2" pageNumber="451">ogygia</emphasis>
|
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</taxonomicName>
|
||||
is
|
||||
<taxonomicName id="4C45779040570B2EFD8B7BDFFD73682E" authorityName="Christoph" authorityYear="1893" box="[543,646,1265,1286]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="ornata">
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<emphasis id="B931D00140570B2EFD8B7BDFFD73682E" box="[543,646,1265,1286]" italics="true" pageId="2" pageNumber="451">M. ornata</emphasis>
|
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</taxonomicName>
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||||
and not
|
||||
<taxonomicName id="4C45779040570B2EFD717BDFFCA7682E" box="[741,850,1265,1286]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="B931D00140570B2EFD717BDFFCA7682E" box="[741,850,1265,1286]" italics="true" pageId="2" pageNumber="451">M. phoebe</emphasis>
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</taxonomicName>
|
||||
as described originally by Fruhstorfer, a suitable label has been attached (
|
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<figureCitation id="137E109640570B2EFDD17A23FD7F680A" box="[581,650,1293,1314]" captionStart="Figs 8-9.– 8" captionStartId="17.[157,195,1501,1520]" captionTargetBox="[136,1134,203,1619]" captionTargetId="graphics-18@17.[136,1134,144,1716]" captionTargetPageId="17" captionText="Figs 8-9.– 8A, B. & C. Melitaea phoebe ogygia Fruhstorfer, 1908, female lectotype, Greece, Poros, 14 June 1909, misidentified M. ornata. © R. Rougeri. 9A, B & C. Melitaea phoebe guevara Fruhstorfer, 1917, male paralectotype, Spain, Castilien, Cuenca Mont., 1900 Korb, misidentified M. ornata. © R. Rougeri. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="2" pageNumber="451">Fig. 8c</figureCitation>
|
||||
).
|
||||
</paragraph>
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</subSubSection>
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</treatment>
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</document>
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107
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Fruhstorfer, 1917
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(a)
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<paragraph id="8BFA0C1340570B2FFD0E790AFD786C41" blockId="2.[130,1128,203,1621]" lastBlockId="3.[136,1135,201,1621]" lastPageId="3" lastPageNumber="452" pageId="2" pageNumber="451">[TL: Bosnia and Herzegovina, Herzegovina, Jablanica] was described from five females in the “Leonhard collection” (Leonhard does not appear to be in any list of entomologists and thus was presumably an amateur butterfly collector) and a pair of “Types” in Fruhstorfer’s collection (FRUHSTORFER, 1916: 1) in the MNHN, Paris (MNHNP). BERNARDI & DE LESSE (1951: 141) reported only a single male “holotype” in Fruhstorfer’s collection but both a male and a female are present in the collection, thus there appear to be two syntypes. A further “paratype” (i.e. a syntype) was reported as being in the Natural History Museum, London (VAN OORSCHOT & COUTSIS (2014: 61).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F9840560B2FFF2D7E56FE626F79" pageId="3" pageNumber="452" type="discussion">
|
||||
<paragraph id="8BFA0C1340560B2FFF2D7E56FE626F79" blockId="3.[136,1135,201,1621]" pageId="3" pageNumber="452">
|
||||
Jablanica lies on the Neretva River at an elevation of c. 200 m; VAN OORSCHOT & COUTSIS (2014: 61) unfortunately misconstrued this as Mount Jablanica, which is on the North Macedonia/Albania border and mostly above 2000 m. The true locality, adjacent to both the Neretva River and Jablanica Lake, suggests a humid biotope typical of
|
||||
<taxonomicName id="4C45779040560B2FFD767EE2FCB26CC9" box="[738,839,460,481]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140560B2FFD767EE2FCB26CC9" box="[738,839,460,481]" italics="true" pageId="3" pageNumber="452">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
; however prior to the building of the dam in 1954, the habitat was more xerophilous. On close inspection of the photographs of the undersides of the two specimens it was concluded that they belonged to
|
||||
<taxonomicName id="4C45779040560B2FFCF67D2AFC3F6F31" box="[866,970,516,537]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D00140560B2FFCF67D2AFC3F6F31" box="[866,970,516,537]" italics="true" pageId="3" pageNumber="452">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
as described by Fruhstorfer and in agreement with VAN OORSCHOT & COUTSIS (2014: 61) and RUSSELL & TENNENT (2016: note 55).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
197
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<mods:title id="C962029F5B78020016B20823643EA470">Designation of lectotypes for some Spanish and other western European Melitaea taxa, some with mixed syntypic series of M. phoebe ([Denis & amp; Schiffermüller], 1775) and M. ornata Christoph, 1893 (Lepidoptera: Nymphalidae)</mods:title>
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<mods:namePart id="1DC9AA761970D555CE7C938F746775A3">Bartolozzi, L.</mods:namePart>
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<mods:affiliation id="B8079DA7AED2EDD33AE69843B6AB2093">Natural History Museum of the University of Florence Zoology Department ‘ La Specola’ via Romana, 17 I- 50125 Florence ITALIA / ITALY</mods:affiliation>
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<mods:affiliation id="40962344CE4F789A9E840B2E01CBCA0D">Department of Life Sciences The Natural History Museum GB-London SW 7 5 BD REINO UNIDO / UNITED KINGDOM</mods:affiliation>
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<treatment id="03ECBD05405D0B24FF277E87FCC56950" ID-DOI="http://doi.org/10.5281/zenodo.15237508" ID-Zenodo-Dep="15237508" LSID="urn:lsid:plazi:treatment:03ECBD05405D0B24FF277E87FCC56950" httpUri="http://treatment.plazi.org/id/03ECBD05405D0B24FF277E87FCC56950" lastPageNumber="457" pageId="8" pageNumber="457">
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<subSubSection id="C35F5F98405D0B24FF277E87FD906C96" box="[179,613,425,446]" pageId="8" pageNumber="457" type="nomenclature">
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<paragraph id="8BFA0C13405D0B24FF277E87FD906C96" blockId="8.[130,1128,201,1145]" box="[179,613,425,446]" pageId="8" pageNumber="457">
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<taxonomicName id="4C457790405D0B24FF277E87FD906C96" ID-CoL="7KC4H" authority="Verity, 1952" authorityName="Verity" authorityYear="1952" box="[179,613,425,446]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="subtusca">
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<emphasis id="B931D001405D0B24FF277E87FEA36C96" box="[179,342,425,446]" italics="true" pageId="8" pageNumber="457">Melitaea phoebe</emphasis>
|
||||
race
|
||||
<emphasis id="B931D001405D0B24FE047E87FE116C96" box="[400,484,425,446]" italics="true" pageId="8" pageNumber="457">subtusca</emphasis>
|
||||
Verity, 1952
|
||||
</taxonomicName>
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</paragraph>
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<subSubSection id="C35F5F98405D0B24FDFA7E87FD9C6EF8" pageId="8" pageNumber="457" type="description">
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[TL: France, Var, La Sainte Baume, Nans-les-Pins, 300 m], the syntypic series consists of seven specimens (4 11 and 3 00) from the Type Locality taken between 24 May 1926 and 24 May 1936 and held in the Museo Zoologico de “La Specola” dell’Università, Firenza, Italy. From an examination of the photographs of the undersides of all seven specimens, it would appear that they include both
|
||||
<taxonomicName id="4C457790405D0B24FDFC7D36FD3B6F05" box="[616,718,536,557]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="phoebe">
|
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<emphasis id="B931D001405D0B24FDFC7D36FD3B6F05" box="[616,718,536,557]" italics="true" pageId="8" pageNumber="457">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C457790405D0B24FD6A7D36FCAA6F05" authorityName="Christoph" authorityYear="1893" box="[766,863,536,557]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D001405D0B24FD6A7D36FCAA6F05" box="[766,863,536,557]" italics="true" pageId="8" pageNumber="457">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
. A syntype of each species is illustrated for comparison: one male has morphological characters tending towards those of
|
||||
<taxonomicName id="4C457790405D0B24FBD97D1AFF376F4D" authorityName="Christoph" authorityYear="1893" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B931D001405D0B24FBD97D1AFF376F4D" italics="true" pageId="8" pageNumber="457">M. ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(Figs 19a, b); another male has characters closely resembling
|
||||
<taxonomicName id="4C457790405D0B24FCBE7D7EFC646F4D" box="[810,913,592,613]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D001405D0B24FCBE7D7EFC646F4D" box="[810,913,592,613]" italics="true" pageId="8" pageNumber="457">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E1096405D0B24FC347D7EFBD36F4D" box="[928,1062,592,613]" captionStart="Fig" captionStartId="23.[157,188,1073,1092]" captionTargetId="figure-92@23.[399,870,635,1036]" captionTargetPageId="23" captionText="Fig. 20A, B & C.– Melitaea phoebe subtusca Verity, 1952, male syntype Nans les Pins, Var, France, 24 May 1926, misidentified M. ornata. © S. Bambi. [A = upperside (dorsal surface), B = underside (ventral surface), C = labels on specimen’s pin]." pageId="8" pageNumber="457">Figs 20a & b</figureCitation>
|
||||
). Both are labelled as having been taken on the same day, 24 May 1926 (Figs 19c, 20c), but handwriting on the data labels suggests by different collectors; this raises the possibility that they were captured some distance apart. The only indication of who captured another of the specimens is a label “23-V-[19]33 Nans (Var) Foulquier leg.’’; he must have been accompanied by another collector because there is another label dated ‘Nans 23 Mai [19]33’ again in a different hand! These two simultaneous captures by different collectors could suggest that the two specimens captured on each occasion (24 May 1926 and 23 May 1933) were taken some distance apart but with the nearest location reference for the data labels both being ‘Nans’. It is unsurprising that Gédéon Foulquier (1855-1941) collected on the Massif de la St. Baume since he lived in Marseille, just to the south of the mountain ridge. Currently, it is not possible to reliably place
|
||||
<taxonomicName id="4C457790405D0B24FEE37C46FE3E6E55" authorityName="Verity" authorityYear="1952" box="[375,459,872,893]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="subtusca">
|
||||
<emphasis id="B931D001405D0B24FEE37C46FE3E6E55" box="[375,459,872,893]" italics="true" pageId="8" pageNumber="457">subtusca</emphasis>
|
||||
</taxonomicName>
|
||||
with either species.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA0C13405D0B24FF277CAAFD9C6EF8" blockId="8.[130,1128,201,1145]" pageId="8" pageNumber="457">
|
||||
The following
|
||||
<taxonomicName id="4C457790405D0B24FED07CAAFE6C6EB1" authorityName="Fabricius" authorityYear="1807" box="[324,409,900,921]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B931D001405D0B24FED07CAAFE6C6EB1" box="[324,409,900,921]" italics="true" pageId="8" pageNumber="457">Melitaea</emphasis>
|
||||
</taxonomicName>
|
||||
subspecies described by Fruhstorfer from damp Alpine regions, based on photographs of the syntypes are all
|
||||
<taxonomicName id="4C457790405D0B24FE487C8EFDB76E9D" box="[476,578,928,949]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D001405D0B24FE487C8EFDB76E9D" box="[476,578,928,949]" italics="true" pageId="8" pageNumber="457">M. phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
, as originally described:
|
||||
<taxonomicName id="4C457790405D0B24FCAC7C8EFC9E6E9D" authorityName="Fruhstorfer" authorityYear="1908" box="[824,875,928,949]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="koios">
|
||||
<emphasis id="B931D001405D0B24FCAC7C8EFC9E6E9D" box="[824,875,928,949]" italics="true" pageId="8" pageNumber="457">koios</emphasis>
|
||||
</taxonomicName>
|
||||
(1908b),
|
||||
<taxonomicName id="4C457790405D0B24FC5E7C8EFBE56E9D" authorityName="Fruhstorfer" authorityYear="1917" box="[970,1040,928,949]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="virgilia">
|
||||
<emphasis id="B931D001405D0B24FC5E7C8EFBE56E9D" box="[970,1040,928,949]" italics="true" pageId="8" pageNumber="457">virgilia</emphasis>
|
||||
</taxonomicName>
|
||||
(1917a),
|
||||
<taxonomicName id="4C457790405D0B24FF167C92FF386EF9" authorityName="Fruhstorfer" authorityYear="1917" box="[130,205,956,977]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="sylleion">
|
||||
<emphasis id="B931D001405D0B24FF167C92FF386EF9" box="[130,205,956,977]" italics="true" pageId="8" pageNumber="457">sylleion</emphasis>
|
||||
</taxonomicName>
|
||||
(1917a),
|
||||
<taxonomicName id="4C457790405D0B24FEBD7C92FE906EF9" authorityName="Fruhstorfer" authorityYear="1917" box="[297,357,956,977]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="minoa">
|
||||
<emphasis id="B931D001405D0B24FEBD7C92FE906EF9" box="[297,357,956,977]" italics="true" pageId="8" pageNumber="457">minoa</emphasis>
|
||||
</taxonomicName>
|
||||
(1917a), and
|
||||
<taxonomicName id="4C457790405D0B24FE7E7C92FDE96EF9" authorityName="Fruhstorfer" authorityYear="1923" box="[490,540,956,977]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="rovia">
|
||||
<emphasis id="B931D001405D0B24FE7E7C92FDE96EF9" box="[490,540,956,977]" italics="true" pageId="8" pageNumber="457">rovia</emphasis>
|
||||
</taxonomicName>
|
||||
(1919).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F5F98405D0B24FF277CF6FCC56950" pageId="8" pageNumber="457" type="distribution">
|
||||
<paragraph id="8BFA0C13405D0B24FF277CF6FCC56950" blockId="8.[130,1128,201,1145]" pageId="8" pageNumber="457">
|
||||
Additional subspecies described by Verity and present in the La Specola Museum in Florence, from France, Switzerland and Italy were all examined and confirmed to be
|
||||
<taxonomicName id="4C457790405D0B24FCF67CDAFBF16921" box="[866,1028,1012,1033]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="phoebe">
|
||||
<emphasis id="B931D001405D0B24FCF67CDAFBF16921" box="[866,1028,1012,1033]" italics="true" pageId="8" pageNumber="457">Melitaea phoebe</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<taxonomicName id="4C457790405D0B24FB867CDAFB926921" authorityName="Verity" authorityYear="1928" box="[1042,1127,1012,1033]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="monilata">
|
||||
<emphasis id="B931D001405D0B24FB867CDAFB926921" box="[1042,1127,1012,1033]" italics="true" pageId="8" pageNumber="457">monilata</emphasis>
|
||||
</taxonomicName>
|
||||
(1919),
|
||||
<taxonomicName id="4C457790405D0B24FF4D7B3EFEE5690D" authorityName="Verity" authorityYear="1919" box="[217,272,1040,1061]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="tusca">
|
||||
<emphasis id="B931D001405D0B24FF4D7B3EFEE5690D" box="[217,272,1040,1061]" italics="true" pageId="8" pageNumber="457">tusca</emphasis>
|
||||
</taxonomicName>
|
||||
(1919),
|
||||
<taxonomicName id="4C457790405D0B24FEE67B3EFE1A690D" authorityName="Verity" authorityYear="1928" box="[370,495,1040,1061]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="crassenigra">
|
||||
<emphasis id="B931D001405D0B24FEE67B3EFE1A690D" box="[370,495,1040,1061]" italics="true" pageId="8" pageNumber="457">crassenigra</emphasis>
|
||||
</taxonomicName>
|
||||
(1928),
|
||||
<taxonomicName id="4C457790405D0B24FDC57B3EFD1E690D" authorityName="Verity" authorityYear="1928" box="[593,747,1040,1061]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="subcorythallia">
|
||||
<emphasis id="B931D001405D0B24FDC57B3EFD1E690D" box="[593,747,1040,1061]" italics="true" pageId="8" pageNumber="457">subcorythallia</emphasis>
|
||||
</taxonomicName>
|
||||
(1928),
|
||||
<taxonomicName id="4C457790405D0B24FCD97B3EFC14690D" authorityName="Verity" authorityYear="1928" box="[845,993,1040,1061]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="suboccitanica">
|
||||
<emphasis id="B931D001405D0B24FCD97B3EFC14690D" box="[845,993,1040,1061]" italics="true" pageId="8" pageNumber="457">suboccitanica</emphasis>
|
||||
</taxonomicName>
|
||||
(1928), and
|
||||
<taxonomicName id="4C457790405D0B24FF167B02FEFB6969" authorityName="Verity" authorityYear="1950" box="[130,270,1068,1089]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="medioastricta">
|
||||
<emphasis id="B931D001405D0B24FF167B02FEFB6969" box="[130,270,1068,1089]" italics="true" pageId="8" pageNumber="457">medioastricta</emphasis>
|
||||
</taxonomicName>
|
||||
(1950). The following available names were not considered because of a lack of surviving specimens in what remains of Verity’s collection in Florence:
|
||||
<taxonomicName id="4C457790405D0B24FCE57B66FBE06975" authorityName="Verity" authorityYear="1919" box="[881,1045,1096,1117]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="monilataeformis">
|
||||
<emphasis id="B931D001405D0B24FCE57B66FBE06975" box="[881,1045,1096,1117]" italics="true" pageId="8" pageNumber="457">monilataeformis</emphasis>
|
||||
</taxonomicName>
|
||||
(1919),
|
||||
<taxonomicName id="4C457790405D0B24FF167B4AFEE86951" authorityName="Verity" authorityYear="1919" box="[130,285,1124,1145]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="aethereaeformis">
|
||||
<emphasis id="B931D001405D0B24FF167B4AFEE86951" box="[130,285,1124,1145]" italics="true" pageId="8" pageNumber="457">aethereaeformis</emphasis>
|
||||
</taxonomicName>
|
||||
(1919),
|
||||
<emphasis id="B931D001405D0B24FEFB7B4AFE086951" box="[367,509,1124,1145]" italics="true" pageId="8" pageNumber="457">nigroalternans</emphasis>
|
||||
(1919) and
|
||||
<taxonomicName id="4C457790405D0B24FDE57B4AFD166951" authorityName="Verity" authorityYear="1939" box="[625,739,1124,1145]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="postnarenta">
|
||||
<emphasis id="B931D001405D0B24FDE57B4AFD166951" box="[625,739,1124,1145]" italics="true" pageId="8" pageNumber="457">postnarenta</emphasis>
|
||||
</taxonomicName>
|
||||
(1939).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue