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<document ID-CLB-Dataset="287343" ID-DOI="http://dx.doi.org/10.3897/mycokeys.102.117067" ID-GBIF-Dataset="39a17317-78a1-4640-92d4-0bd5e973e7cf" ID-Pensoft-Pub="1314-4049-102-29" ID-Pensoft-UUID="BFFD3833DA5359D3A737FBCD03307916" ModsDocID="1314-4049-102-29" checkinTime="1707224635089" checkinUser="pensoft" docAuthor="Gunaseelan, Sugantha, Kezo, Kezhocuyi, Karunarathna, Samantha C., Yang, Erfu, Zhao, Changlin, Elgorban, Abdallah M., Tibpromma, Saowaluck &amp; Kaliyaperumal, Malarvizhi" docDate="2024" docId="62082AE9516E542FB4B61BFE5B345743" docLanguage="en" docName="MycoKeys 102: 29-54" docOrigin="MycoKeys 102" docPubDate="2024-02-05" docSource="http://dx.doi.org/10.3897/mycokeys.102.117067" docTitle="Tropicoporus indicus S. Gunaseelan &amp; M. Kaliyaperumal 2024, sp. nov." docType="treatment" docVersion="2" id="BFFD3833DA5359D3A737FBCD03307916" lastPageNumber="29" masterDocId="BFFD3833DA5359D3A737FBCD03307916" masterDocTitle="New species of Tropicoporus (Basidiomycota, Hymenochaetales, Hymenochaetaceae) from India, with a key to Afro-Asian Tropicoporus species" masterLastPageNumber="54" masterPageNumber="29" pageNumber="29" updateTime="1707225250396" updateUser="ExternalLinkService">
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<mods:title id="A7EEB3B2724C14CA259B199CBC075CF1">New species of Tropicoporus (Basidiomycota, Hymenochaetales, Hymenochaetaceae) from India, with a key to Afro-Asian Tropicoporus species</mods:title>
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<mods:namePart id="6DD4B26E710A902DEC8824EF4B03214A">Gunaseelan, Sugantha</mods:namePart>
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<mods:affiliation id="4836CC2FEB1E4D941D2DE76F3DCFD866">Centre for Advanced Studies in Botany, University of Madras, Guindy Campus, Chennai 600025, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="96BDF8AE01498BB9B04BA76FB1C9F571">Kezo, Kezhocuyi</mods:namePart>
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<mods:affiliation id="E60226D6C1718EA64426969429C5CECE">Centre for Advanced Studies in Botany, University of Madras, Guindy Campus, Chennai 600025, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="7311CFCBFB39C28F1AE0097A96984B35">Karunarathna, Samantha C.</mods:namePart>
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<mods:affiliation id="14CFFF7D322631295A46F26A0355653B">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China &amp; National Institute of Fundamental Studies (NIFS), Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart id="DF2D897D143110C0AD21F03DB37531E0">Yang, Erfu</mods:namePart>
<mods:nameIdentifier id="72F7AA4D845926050B2BC346027946DE" type="ORCID">https://orcid.org/0000-0003-2385-6402</mods:nameIdentifier>
<mods:affiliation id="78361E3BB18E461DEB42109C5B0ED877">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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<mods:namePart id="7B577756B411E3537FBAAB12CBEEFE34">Zhao, Changlin</mods:namePart>
<mods:nameIdentifier id="D8FA60275A664DE8B72D038D9F1F8ACC" type="ORCID">https://orcid.org/0000-0002-1395-591X</mods:nameIdentifier>
<mods:affiliation id="4EF618DFF76D6B19880EF37B32130302">College of Biodiversity Conservation, Southwest Forestry University, Kunming 650224, China</mods:affiliation>
</mods:name>
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<mods:namePart id="49EDF20822646A8E7008751D5421E1B0">Elgorban, Abdallah M.</mods:namePart>
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<mods:affiliation id="02EC1637314B7AB6F1FE2AFE21807B53">Department of Botany and Microbiology, College of Science, King Saud University, Riyadh 11451, Saudi Arabia</mods:affiliation>
</mods:name>
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<mods:namePart id="AB805127CE9014BC88E9BAB69F9B7188">Tibpromma, Saowaluck</mods:namePart>
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<mods:affiliation id="183609F872890ED61B600406C3F09E1A">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
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<mods:namePart id="850B2C4B48C5646E93F6696689404C62">Kaliyaperumal, Malarvizhi</mods:namePart>
<mods:nameIdentifier id="BDF15F2C17062042453D5DF9843AC468" type="ORCID">https://orcid.org/0000-0002-1218-3778</mods:nameIdentifier>
<mods:affiliation id="177836F78E42235C317E8F32E117CC2F">Centre for Advanced Studies in Botany, University of Madras, Guindy Campus, Chennai 600025, Tamil Nadu, India</mods:affiliation>
<mods:nameIdentifier id="08759552596E12E708EA90B3BA1E9706" type="email">malar.kaliyaperumal@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:date id="427B699225A3FC3CADB7DD772CD597DC">2024</mods:date>
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<mods:number id="4300CCB16967C21E219B3C25CF3E927A">102</mods:number>
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<mods:url id="C072AA8426C26E2CEF3D38D684536E7B">http://dx.doi.org/10.3897/mycokeys.102.117067</mods:url>
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<mods:classification id="B2F91F571A0FBD746902D78A5B5963E8">journal article</mods:classification>
<mods:identifier id="473AB05837EB3A583F1E5BED6152799B" type="DOI">http://dx.doi.org/10.3897/mycokeys.102.117067</mods:identifier>
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<treatment id="62082AE9516E542FB4B61BFE5B345743" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743" httpUri="http://treatment.plazi.org/id/62082AE9516E542FB4B61BFE5B345743" lastPageNumber="29" pageId="0" pageNumber="29">
<subSubSection id="5AF2E0B5F77AC7BD11A8CD98C4C50A05" pageId="0" pageNumber="29" type="nomenclature">
<paragraph id="386DD79B1E686B57019C41C410DED53D" pageId="0" pageNumber="29">
<taxonomicName id="2BF937C4BFA1BCA84418456136C6565A" LSID="62082AE9-516E-542F-B4B6-1BFE5B345743" authority="S. Gunaseelan &amp; M. Kaliyaperumal" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus" status="sp. nov.">Tropicoporus indicus S. Gunaseelan &amp; M. Kaliyaperumal</taxonomicName>
<taxonomicNameLabel id="0BF4AFA386FBB85951355A4EDEEA2573" pageId="0" pageNumber="29">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="46251EDD8D25EBE3E279290BBF0DC49A" pageId="0" pageNumber="29" type="description">
<paragraph id="5B97403B05901000C06D035BE6EFF4FB" pageId="0" pageNumber="29">
<figureCitation id="2E5081CFE91A99CFEC099B4C696B618F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropicoporus indicus (MUBL 1083 holotype) A basidiomata (Holotype) B basidiome (SMK-MK 2 a- Isotype) C pore surface D cross-section of basidiome (arrows indicating stratified tube layers) E hymenial setae F cystidioles G-J basidiospores: G basidiospores in water H basidiospores in KOH I basidiospores in cotton blue J basidiospores in Melzer's reagent. Scale bars: 1 cm (A-D); 5 µm (E-J)." figureDoi="10.3897/mycokeys.102.117067.figure5" httpUri="https://binary.pensoft.net/fig/979926" pageId="0" pageNumber="29">Figs 5</figureCitation>
<figureCitation id="F671952AD4917BD5CC18DA591B58C2D5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tropicoporus indicus (MUBL 1083 holotype) A contextual hyphae B tramal hyphae C hymenial setae D cystidioles E basidioles F basidia G basidiospores. Scale bars: 5 µm." figureDoi="10.3897/mycokeys.102.117067.figure6" httpUri="https://binary.pensoft.net/fig/979927" pageId="0" pageNumber="29">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAF800DC178929EBB9A16C954EE6E41C" pageId="0" pageNumber="29" type="etymology">
<paragraph id="0B02D8C13FD011D749F2A58C7573D613" pageId="0" pageNumber="29">Etymology.</paragraph>
<paragraph id="C800399C1B4C0FA04A3167D43CA96491" pageId="0" pageNumber="29">
The species epithet &quot;
<emphasis id="044F20B73D669DB94A5EA2B61957BEAC" italics="true" pageId="0" pageNumber="29">indicus</emphasis>
&quot; (Lat.): referring to the species being collected from India.
</paragraph>
</subSubSection>
<subSubSection id="368C4DC75023C71E89CC7ADD0FDF838F" pageId="0" pageNumber="29" type="diagnosis">
<paragraph id="4D9597E8B1BF60BB4F27FA8D361F6607" pageId="0" pageNumber="29">Diagnosis.</paragraph>
<paragraph id="EF899FD667D892D618687B0837171020" pageId="0" pageNumber="29">
<taxonomicName id="FA0501C519CADD63F1CB94AAC0505925" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="744132A9C18066647965B63534AE0F56" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
is characterised by applanate to meagrely triquetrous basidiome with concentrically zonate, sulcate, glabrous, deeply cracked to rimose pilear surface, homogenous context, acute margin, mono-dimitic hyphal system, presence of cystidioles, subglobose to broadly ellipsoid basidiospores measuring 5-6
<normalizedToken id="FEB817D4525411C29B3E2585ADE237C0" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="7E918504528DB828C6EE214A14020E0E" originalValue="μm">μm</normalizedToken>
.
</paragraph>
<caption id="F7F7BEC721272C72A5CFC1E0E1B1016E" doi="10.3897/mycokeys.102.117067.figure5" httpUri="https://binary.pensoft.net/fig/979926" pageId="0" pageNumber="29" start="Figure 5" startId="F5">
<paragraph id="BEE5D3E91315B53A61FC158B7BA185C6" pageId="0" pageNumber="29">
<emphasis id="8F4AE627BCC5A69425279DEED6D03C90" bold="true" pageId="0" pageNumber="29">Figure 5.</emphasis>
<taxonomicName id="04BE451B3C8F09BCB2D7F23246C83DFB" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="D455BB614D1F11AA5CC819489486E994" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
(MUBL1083 holotype)
<emphasis id="1BFC7E586A241C25522011B93504BA69" bold="true" pageId="0" pageNumber="29">A</emphasis>
basidiomata (Holotype)
<emphasis id="0AF7E936E01BC0AD876868A59FC844C6" bold="true" pageId="0" pageNumber="29">B</emphasis>
basidiome (SMK-MK2a- Isotype)
<emphasis id="66CBFA7998079088C115A3F048A39DC5" bold="true" pageId="0" pageNumber="29">C</emphasis>
pore surface
<emphasis id="9DA68684183AE8E546AC05B172031183" bold="true" pageId="0" pageNumber="29">D</emphasis>
cross-section of basidiome (arrows indicating stratified tube layers)
<emphasis id="4C4572BF89BDE299D826DA962D96C3BC" bold="true" pageId="0" pageNumber="29">E</emphasis>
hymenial setae
<emphasis id="C1258520A83F2E3E47C8DF9CE8FDEA97" bold="true" pageId="0" pageNumber="29">F</emphasis>
cystidioles
<emphasis id="98117C7B4DA3A802CB87F1D21E80F871" bold="true" pageId="0" pageNumber="29">G-J</emphasis>
basidiospores:
<emphasis id="C71223762EED1995AD2EBDA60AAAE7B7" bold="true" pageId="0" pageNumber="29">G</emphasis>
basidiospores in water
<emphasis id="D3F7B13D88B643D602939871B3FB730A" bold="true" pageId="0" pageNumber="29">H</emphasis>
basidiospores in KOH
<emphasis id="D00A54486967153821948C4346314343" bold="true" pageId="0" pageNumber="29">I</emphasis>
basidiospores in cotton blue
<emphasis id="25A034D9FFD0E4FF95A1E322BEC90426" bold="true" pageId="0" pageNumber="29">J</emphasis>
basidiospores in
<normalizedToken id="395FCE2E8AA71DD46801BD7C9977FD0B" originalValue="Melzers">Melzer's</normalizedToken>
reagent. Scale bars: 1 cm (
<emphasis id="399895FDD4F5A6B8771A2C3E2CBBDFB7" bold="true" pageId="0" pageNumber="29">A-D</emphasis>
); 5
<normalizedToken id="573B4995A57AD8D6034AF5B2F45DD600" originalValue="µm">µm</normalizedToken>
(
<emphasis id="60FDFD3C201F4CAE26B9CAC8518D1794" bold="true" pageId="0" pageNumber="29">E-J</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B153723D747BFB2999D29B76464C52D4" pageId="0" pageNumber="29" type="materials_examined">
<paragraph id="C054F912DFEA41B6962D22E0E300BE2D" pageId="0" pageNumber="29">
<typeStatus id="DE05FB7F564E07A637D0135F674E73A8">Type</typeStatus>
.
</paragraph>
<paragraph id="7E1762E532BF4ADB76AE04320BA14B7E" pageId="0" pageNumber="29">
<materialsCitation id="65D5A8F4A463E2F68D1FC8D2E53391B5" collectingDate="2022-09-29" country="India" location="Kalvarayan Hills" municipality="Kallakurichi District" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="Holotype">
<collectingCountry id="838D83A17E568A2EAFFABB656510106E" name="India">India</collectingCountry>
,
<collectingRegion id="E03789AA88E377A64B85919D5C46A908" country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="C64899D78194C957A1374D7F56367A04">Kallakurichi District</collectingMunicipality>
,
<location id="B92D70A2586D84DCE5CA4F1DDCAF65D6" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743:B92D70A2586D84DCE5CA4F1DDCAF65D6" country="India" municipality="Kallakurichi District" name="Kalvarayan Hills" stateProvince="Tamil Nadu">Kalvarayan Hills</location>
; 11°91'30&quot;N, 78°57'86&quot;E;
<collectingDate id="A3E21C5CB676E09428D79FFE3028B91E" value="2022-09-29">29 Sep 2022</collectingDate>
; Sugantha Gunaseelan; on living angiosperm tree of
<taxonomicName id="7356494E7AE3E2204FE06E49D24E9E46" class="Dicotyledoneae" family="Fabaceae" genus="Albizia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Albizia amara" order="Rosales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="amara">
<emphasis id="D061194C9A6FD987B7D27FFC4E0A674B" italics="true" pageId="0" pageNumber="29">Albizia amara</emphasis>
</taxonomicName>
; SMK- MK2 (MUBL1083,
<typeStatus id="EF522058354CCD8E6F1D6F37889759D1" type="holotype">Holotype</typeStatus>
); GenBank: OR272293 (ITS); OR272338 (nLSU)
</materialsCitation>
.
</paragraph>
<caption id="0417FD3D2E1C7D5D6F79F5571170ECB1" doi="10.3897/mycokeys.102.117067.figure6" httpUri="https://binary.pensoft.net/fig/979927" pageId="0" pageNumber="29" start="Figure 6" startId="F6">
<paragraph id="3404361760D4ABE24912E5BB43A51F73" pageId="0" pageNumber="29">
<emphasis id="81EE7AA8516163E33DB81F4D737DA260" bold="true" pageId="0" pageNumber="29">Figure 6.</emphasis>
<taxonomicName id="3140D02FE7705B07BE1AC9FAFB909EA3" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="8A1381499FD1A636CECD2AB74F42B836" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
(MUBL1083
<typeStatus id="43CB44B3B279BC9EC3F70F74F354C36E" type="holotype">holotype</typeStatus>
)
<emphasis id="8B4B091459877DA2B9FA07399DB6C7B8" bold="true" pageId="0" pageNumber="29">A</emphasis>
contextual hyphae
<emphasis id="A710C7B2FEC6853120673896E6216D8D" bold="true" pageId="0" pageNumber="29">B</emphasis>
tramal hyphae
<emphasis id="571A649858CB441BA96CD9BF8CCD2735" bold="true" pageId="0" pageNumber="29">C</emphasis>
hymenial setae
<emphasis id="C603982024A17BA0804568D4C6DEA17E" bold="true" pageId="0" pageNumber="29">D</emphasis>
cystidioles
<emphasis id="0D67633B6FC55CBA2A7B3278D4BC234B" bold="true" pageId="0" pageNumber="29">E</emphasis>
basidioles
<emphasis id="432667E0EFB1A32AA1442E3E7E4A8806" bold="true" pageId="0" pageNumber="29">F</emphasis>
basidia
<emphasis id="E5C6F6A2267748611CB9FEF583FDF2A8" bold="true" pageId="0" pageNumber="29">G</emphasis>
basidiospores. Scale bars: 5
<normalizedToken id="84CD5828A7E50191D668CCFE9945EBC1" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1FDDB638A1442F8A46F22F5B0CE00762" pageId="0" pageNumber="29" type="description">
<paragraph id="98DBCB21A5C82F7F6F7998E865A6712C" pageId="0" pageNumber="29">Description.</paragraph>
<paragraph id="4FAF4EB773F10A8041B31FABF94617D1" pageId="0" pageNumber="29">
<emphasis id="4A07B64BD97536DC7316937B47CEB234" bold="true" italics="true" pageId="0" pageNumber="29">Basidiome</emphasis>
perennial, pileate, woody and without distinctive odour or taste when fresh, hard when dry.
<emphasis id="93D13D632F46E25A5324C9BBEF7E0C27" bold="true" italics="true" pageId="0" pageNumber="29">Pilei</emphasis>
applanate to meagrely triquetrous, projecting up to 5.5 cm, 7 cm wide and 4 cm thick at the base; pileal surface concentrically zonate, sulcate, glabrous, deeply cracked to rimose near attachment, dark brown (7E4), greyish-brown (7F3).
<emphasis id="A7E0D09CCC88EAACBA811B122321522D" bold="true" italics="true" pageId="0" pageNumber="29">Margin</emphasis>
velutinate, 2 mm thick, acute, brown (6E7).
<emphasis id="B3A7983052C52B00E24F82DC7479311E" bold="true" italics="true" pageId="0" pageNumber="29">Pore surface</emphasis>
light brown (5D5) to yellowish-brown (5E7); pores circular, 4-6 per mm.
<emphasis id="FAABF0438CCB8B9A8CCF780DB31BE4A5" bold="true" italics="true" pageId="0" pageNumber="29">Context</emphasis>
homogenous, up to 0.8 cm thick, brown (6E8) to dark brown (6F8).
<emphasis id="5773F0A59CEB0E64AD1DC1EA3068B995" bold="true" italics="true" pageId="0" pageNumber="29">Tubes</emphasis>
woody hard, up to 2.5 cm long, brown (6E7); tube layers stratified, each stratum up to 0.5 cm long.
</paragraph>
</subSubSection>
<subSubSection id="141A9D6385836279BCD32ED43E141274" pageId="0" pageNumber="29" type="hyphal structures">
<paragraph id="1C2E9AD4E03BA6AD7ACE1130AD4A0549" pageId="0" pageNumber="29">Hyphal structures.</paragraph>
<paragraph id="4B849D2824CEA3990096DA2F9029E339" pageId="0" pageNumber="29">Hyphal system monomitic in the context and dimitic in the trama, tissue darkening with KOH without swelling</paragraph>
</subSubSection>
<subSubSection id="0CD9E240ACCB5F7C663FFBD92544170A" pageId="0" pageNumber="29" type="context">
<paragraph id="443A9278FCEE2A993C7AEE546B8FD941" pageId="0" pageNumber="29">Context.</paragraph>
<paragraph id="62D8E7A78673314CE750C523DF96C78E" pageId="0" pageNumber="29">
Generative hyphae, thin to thick-walled, hyaline to golden yellow, simple septate, rarely branched, 2-5
<normalizedToken id="2567CF407934D51785B603BADD37E539" originalValue="μm">μm</normalizedToken>
diam.
</paragraph>
</subSubSection>
<subSubSection id="75E574BF614993C299550F9CCA1DA46F" pageId="0" pageNumber="29" type="trama">
<paragraph id="B9ECE6697971C4404F00486FADC0E489" pageId="0" pageNumber="29">Trama.</paragraph>
<paragraph id="2A73862BE2196029BBF1DD80DD26168D" pageId="0" pageNumber="29">
Generative hyphae dominant, thin to thick-walled, hyaline to pale yellow, septate, occasionally branched, 2-4.5
<normalizedToken id="1CDEC11197425FE8CA29F48EEFAA561F" originalValue="μm">μm</normalizedToken>
diam. Skeletal hyphae thick-walled with narrow to wide lumen, yellowish-brown, aseptate, unbranched, 2-4
<normalizedToken id="C9FCFA4431D5BA6841EA7F02CF2BCF93" originalValue="μm">μm</normalizedToken>
diam.
</paragraph>
</subSubSection>
<subSubSection id="FBDDA3000150531EB66506A207CC41A8" pageId="0" pageNumber="29" type="hymenium">
<paragraph id="EBDDC36B0F4FD6F84F2236C697940DD7" pageId="0" pageNumber="29">Hymenium.</paragraph>
<paragraph id="DAEB2E79F4C8CE6E6E8EF0C20D86E894" pageId="0" pageNumber="29">
Hymenial setae dark brown, thick-walled, ventricose to subulate with sharp to blunt tips, 7-28
<normalizedToken id="607304B009249B4C40DFB2FC3F93C9B7" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="EFAEE034537567CF407C25F7F5131E8B" originalValue="μm">μm</normalizedToken>
. Cystidia absent. Cystidioles hyaline, thin-walled, ventricose to fusoid with elongated tapering apical portion, 5-21
<normalizedToken id="09EBEC289E4A747B587B77E89B7ACD98" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="2D6FACE1178FDCC2FC6793021D69672B" originalValue="μm">μm</normalizedToken>
. Basidia clavate to broadly clavate, 7-17
<normalizedToken id="C7CE3238EBB532C414159EE1936ECD38" originalValue="×">x</normalizedToken>
3-6
<normalizedToken id="E989E5236DB34BFE676E74CB8F2CEF26" originalValue="μm">μm</normalizedToken>
, with four sterigmata and a simple septum at the base. Basidioles clavate, 5-14
<normalizedToken id="868F6BC28E599AB4402CA0090B14C50B" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="6E522470C071D757166C9A8D234A162E" originalValue="μm">μm</normalizedToken>
. Basidiospores broadly ellipsoid to subglobose, pale yellow in water, turning golden yellow to brown in KOH, thick-walled, smooth, CB ̄, IKI ̄, (5-) 5.3-5.8 (- 6)
<normalizedToken id="FC7D71B7C78D3840173D035BE0BB85CD" originalValue="×">x</normalizedToken>
(4.2-) 4.7-4.9
<normalizedToken id="E18EB24817901976567F0180CB8DF878" originalValue="μm">μm</normalizedToken>
(n = 50/2), Q = 1.16 (Q range 1.05-1.3).
</paragraph>
</subSubSection>
<subSubSection id="2F8EEA2EEBAF0BA881C42DCA287E5FBE" pageId="0" pageNumber="29" type="habitat">
<paragraph id="1A6CC74B66624F147CD6512684B29BAF" pageId="0" pageNumber="29">Habitat and distribution.</paragraph>
<paragraph id="F0747385A9392E373997D9D88F4538DD" pageId="0" pageNumber="29">
Basidiomes were found on living trees of
<taxonomicName id="768FD47B32D89D5C0490A0351D54C084" authorityName="L.W.Zhou, Y.C.Dai &amp; Sheng H.Wu" authorityYear="2015" family="Fabaceae" kingdom="Fungi" lsidName="" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="family">Fabaceae</taxonomicName>
members (
<taxonomicName id="5BF3EC14EF19DA47CDC26296A0BC433F" class="Dicotyledoneae" family="Fabaceae" genus="Albizia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Albizia amara" order="Rosales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="amara">
<emphasis id="3C3B38163648D176BF1F51C684ED06EF" italics="true" pageId="0" pageNumber="29">Albizia amara</emphasis>
</taxonomicName>
and
<taxonomicName id="EC65242800100FA474C5C95030CDD797" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Prosopis cineraria" order="Fabales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="cineraria">
<emphasis id="33CA6C2825C2B88E1DD2D8F40F362739" italics="true" pageId="0" pageNumber="29">Prosopis cineraria</emphasis>
</taxonomicName>
), distributed in Kalvarayan Hills, Kallakurichi District, Tamil Nadu, India.
</paragraph>
</subSubSection>
<subSubSection id="7BA82C9FD7C07A575AF531310B8DE92C" pageId="0" pageNumber="29" type="materials_examined">
<paragraph id="FF1D6B35849BA76C1B7E48E05707C2E8" pageId="0" pageNumber="29">Additional material examined.</paragraph>
<paragraph id="483A64701DF586D4F190A14F5B631C80" pageId="0" pageNumber="29">
<materialsCitation id="88EC66C3741B1BA40BE3BB40DC7AB17A" collectingDate="2022-09-29" country="India" location="Kalvarayan Hills" municipality="Kallakurichi District" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="Paratype">
<collectingCountry id="84B6BA94B8495944C0BEB96621C5CF14" name="India">India</collectingCountry>
,
<collectingRegion id="68AC01C23429F3484422E1175201FEE0" country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="367EDD920F008D3F2E3DC595057A54AA">Kallakurichi District</collectingMunicipality>
,
<location id="113318AC5CEE30896BCBCF5E5FEC6653" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743:113318AC5CEE30896BCBCF5E5FEC6653" country="India" municipality="Kallakurichi District" name="Kalvarayan Hills" stateProvince="Tamil Nadu">Kalvarayan Hills</location>
; 11°90'39&quot;N, 78°55'69&quot;E; on a living angiosperm tree (
<taxonomicName id="3332666F69E88D433B1F97A2BC251725" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Prosopis cineraria" order="Fabales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="cineraria">
<emphasis id="52B2E9050875A720B86A5ACBCD69457A" italics="true" pageId="0" pageNumber="29">Prosopis cineraria</emphasis>
</taxonomicName>
);
<collectingDate id="7F1186D5D9DA111D96C70E3B5A30F47A" value="2022-09-29">29 Sep 2022</collectingDate>
; Kezhocuyi Kezo; SMK-MK2a (MUBL1084,
<typeStatus id="124DBAB3BE02C44BEFA1DFC7191BAC66" type="paratype">Paratype</typeStatus>
); GenBank: OR272294 (ITS); OR272339 (nLSU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="21542072A96A093A2F4DAC979EEB7137" pageId="0" pageNumber="29" type="notes">
<paragraph id="7508C31DAE256A46DAD9257199805ABA" pageId="0" pageNumber="29">Notes.</paragraph>
<paragraph id="CBF2696AB924C521661F3942FFB366C1" pageId="0" pageNumber="29">
Phylogenetically,
<taxonomicName id="A7D15F9C6E91E1A2005759ABCD11FB23" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="F6B16A66DC6178BFFC34B79ABB5AE0D4" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
was recovered in the
<taxonomicName id="757C53F94C54C1AA1D425C16B4E455F0" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="706063A4D60CC79DE7983763834A5959" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
clade.
<taxonomicName id="290473D566A981A2EB27449C51AF4F8F" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="AB61F023984706CE3D47B8B0572DEBAC" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9ADBB6FF122D484DDE26F70CC2DF57B" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="04270A33FE3F0025C83083D15598BEF6" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
by sharing, pileate, dimidiate basidiomes, concentrically sulcate pilear surface, zonate context, smaller pores (5-7/mm), a mono-dimitic hyphal system and presence of cystidioles. While varying in the nature of cracks,
<taxonomicName id="0997FF0E8004F89E6C761DEB8ABF6600" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="B3977EB245F3D6E499F377572B9FADAB" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
has more or less cracked basidiomes,
<taxonomicName id="010052204A51053ADE6CF54A429B0E86" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="8C0C32E4AA0732285609918F39C76C90" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
has irregular deep cracks in basidiomes, with larger basidiospores (
<taxonomicName id="72830352DCE2554A7235626789FCD6DD" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="5F555F4C4C97E454B5E3BEFC19F9FEC5" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
4.8-5.7
<normalizedToken id="48D049455B43C16187C8E379AE62A386" originalValue="×">x</normalizedToken>
3.8-4.8
<normalizedToken id="0F748C7A7088E2942C1E5059F00FFF76" originalValue="μm">μm</normalizedToken>
and
<taxonomicName id="885DADA9BE758F105C9062726B5CB986" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="8DBB3F8BF3078F77C60B6DCF0673F6D1" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
5-6
<normalizedToken id="B6DDC768A90196223CB40D1A2DD4653C" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="3A4BC333663BCD1B7D0C6DD94C2C55C8" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="3E01B73412D6C6BC5C4B6666A1BB04AB" DOI="https://doi.org/10.1007/s13225-012-0202-9" author="Tian, XM" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="159 - 169" refId="B25" refString="Tian, XM, Yu, HY, Zhou, LW, Decock, C, Vlasak, J, Dai, YC, 2013. Phylogeny and taxonomy of the Inonotus linteus complex. Fungal Diversity 58 (1): 159 - 169, DOI: https://doi.org/10.1007/s13225-012-0202-9" title="Phylogeny and taxonomy of the Inonotus linteus complex." url="https://doi.org/10.1007/s13225-012-0202-9" volume="58" year="2013">Tian et al. 2013</bibRefCitation>
;
<bibRefCitation id="7AAEDE9F84BFC8CB20E630A9799C48F9" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="B85B63A409C56D98CF489E0C0BE724E7" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="0A32D642975E3FD1C106B145EE1FA9F2" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="C87F1423FBB08087E5E903B345951218" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="CFD24BEFE5F8CE1742D9D7CF1D7332B1" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
share a mono-dimitic hyphal system, but
<taxonomicName id="D7701BAFAF7D08BC69DFF9216C40D5F2" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="A49328DA1EAE65E722B428386F04EA60" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="62228A9067A01ECC79E5741838BED530" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="61E5AED4F8C4E1141AF242602451453C" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
in having zonate, sulcate, deeply cracked to rimose basidiomes and larger pores (4-6/mm). In these regards,
<taxonomicName id="06F0CAD7D2B8FF2AA67F100D4B920138" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="97190D5527A9B3C78C1EEC564343B500" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
is characterised by fulvous, velvety, concentrically zonate, matted, rimose basidiome and smaller pores (6-7/mm) (
<bibRefCitation id="D2E879DC954C6D669C9F1650E39E0E6B" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="51CFD021F464E69FD8BD19D25CB2E48B" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="BD68A9F67E1CFE47A3D7D769F69EA4E2" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="DC4ECCEBBADE26624948B0A6D011ECEF" lsidName="T. angustisulcatus" pageId="0" pageNumber="29" rank="species" species="angustisulcatus">
<emphasis id="1704375BF970D9C294A131BAC7D25E3D" italics="true" pageId="0" pageNumber="29">T. angustisulcatus</emphasis>
</taxonomicName>
,
<taxonomicName id="22EB3450D6EF3EB914A2ED3173DC8544" lsidName="T. excentrodendri" pageId="0" pageNumber="29" rank="species" species="excentrodendri">
<emphasis id="08F1F80D2B38E6906DFF76F454E4C563" italics="true" pageId="0" pageNumber="29">T. excentrodendri</emphasis>
</taxonomicName>
,
<taxonomicName id="8C0E34D1E4F32D5024ADC9BBBFA57F31" lsidName="T. lineatus" pageId="0" pageNumber="29" rank="species" species="lineatus">
<emphasis id="BED1EF3E56689C26F45FA7A7C7FE4039" italics="true" pageId="0" pageNumber="29">T. lineatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C3B1B86418960A0C08E0E27739B7E135" lsidName="T. substratificans" pageId="0" pageNumber="29" rank="species" species="substratificans">
<emphasis id="F67AFBEE50F5FCA3D161CA3AAD708340" italics="true" pageId="0" pageNumber="29">T. substratificans</emphasis>
</taxonomicName>
in having mono-dimitic hyphal system and a cracked basidiome (
<bibRefCitation id="64BB0B816B6959FB42B66D2782633F5B" DOI="https://doi.org/10.1007/s13225-015-0335-8" author="Zhou, LW" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="335 - 347" refId="B31" refString="Zhou, LW, Vlasak, J, Decock, C, Assefa, A, Stenlid, J, Abate, D, Wu, SH, Dai, YC, 2015. Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations. Fungal Diversity 77 (1): 335 - 347, DOI: https://doi.org/10.1007/s13225-015-0335-8" title="Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations." url="https://doi.org/10.1007/s13225-015-0335-8" volume="77" year="2015">Zhou et al. 2015</bibRefCitation>
;
<bibRefCitation id="365C9053C4A4877ADF6045036A0BC3A8" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="8A7D735461597D07595ACECA099D033E" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="7AC01A0EB90535B9052160AF5D9D1917" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="2887B4C182C3FD5C33E5DD64BE4944DE" lsidName="T. flabellatus" pageId="0" pageNumber="29" rank="species" species="flabellatus">
<emphasis id="B8DB698885E5BD77BC72C66FFF58FEF6" italics="true" pageId="0" pageNumber="29">T. flabellatus</emphasis>
</taxonomicName>
are similar in having a mono-dimitic hyphal system, but differ significantly by having concentrical zones, sulcate, glabrous, deeply-cracked to rimose pilear surface with larger pores (4-6/mm) and broadly ellipsoid to subglobose spores (5-6
<normalizedToken id="F6716CC838A4A323C91607FE08BB4A59" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="5FBD43EA80F88F4BE6381F7433FF7359" originalValue="μm">μm</normalizedToken>
).
<taxonomicName id="FC1B44D11C81C48381332AAA0C965DDA" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus flabellatus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="flabellatus">
<emphasis id="C7FE21E2F2A58AC3C3E3DDFEC2A6DA61" italics="true" pageId="0" pageNumber="29">Tropicoporus flabellatus</emphasis>
</taxonomicName>
, in contrast, has a velutinate pilear surface, uncracked basidiomes with smaller pores (7-9/mm) and smaller basidiospores (4.5-5
<normalizedToken id="5A4936E733DB2B94986096871DBDFACE" originalValue="×">x</normalizedToken>
3.5-4
<normalizedToken id="A4C1701FC7C12EC8FE79DD0C6B5C36BB" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="D3C5238A3AE64D51DC8678CA28D1A8DF" DOI="https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" author="Lima, VX" journalOrPublisher="Mycologie" pageId="0" pageNumber="29" pagination="1 - 21" refId="B19" refString="Lima, VX, de Oliveira, VRT, de Lima-Junior, NC, Oliveira-Filho, JRC, Santos, C, Lima, N, Gibertoni, TB, 2022. Taxonomy and phylogenetic analysis reveal one new genus and three new species in Inonotus s.l. (Hymenochaetaceae) from Brazil. Cryptogamie. Mycologie 43 (1): 1 - 21, DOI: https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" title="Taxonomy and phylogenetic analysis reveal one new genus and three new species in Inonotus s. l. (Hymenochaetaceae) from Brazil. Cryptogamie." url="https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" volume="43" year="2022">Lima et al. 2022</bibRefCitation>
).
<taxonomicName id="2230DFB9DD5EA61B590758E07106062A" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="A75CFFF4B858D143E8B78E340EFE5873" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="F87A8A4D81B35A7BABA342DF3938A9D8" lsidName="T. guanacastensis" pageId="0" pageNumber="29" rank="species" species="guanacastensis">
<emphasis id="39E69F2204BF61C8931144860F496456" italics="true" pageId="0" pageNumber="29">T. guanacastensis</emphasis>
</taxonomicName>
are similar in having sulcate, cracked basidiome, stratified tube, mono-dimitic hyphal system and ventricose setae. However, the former differs in larger pores (4-6/mm) and larger basidiospores (
<taxonomicName id="FCFBDDFAFD4F6B02B9A6E02AD699CA4A" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="FD4F9733CA341389FCAE2A0F3EFECB24" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
5-6
<normalizedToken id="EDB663B085763F9983F378D7839AB6DE" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="D92D1CE66BD07B1E073246BBF75440B0" originalValue="μm">μm</normalizedToken>
vs.
<taxonomicName id="9E8199B862C36D0A21F8DA0C4BBAC532" lsidName="T. guanacastensis" pageId="0" pageNumber="29" rank="species" species="guanacastensis">
<emphasis id="AAACAD3CF1864039C16CFB1990D8F1FE" italics="true" pageId="0" pageNumber="29">T. guanacastensis</emphasis>
</taxonomicName>
4.1-5
<normalizedToken id="725D64489769E4312F118087D3519BA1" originalValue="×">x</normalizedToken>
3.1-4
<normalizedToken id="43EDD852A765743E4BD7E9BA933C86E4" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="8911BFE71B3217A7E13918CE3335E80E" DOI="https://doi.org/10.1007/s13225-015-0335-8" author="Zhou, LW" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="335 - 347" refId="B31" refString="Zhou, LW, Vlasak, J, Decock, C, Assefa, A, Stenlid, J, Abate, D, Wu, SH, Dai, YC, 2015. Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations. Fungal Diversity 77 (1): 335 - 347, DOI: https://doi.org/10.1007/s13225-015-0335-8" title="Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations." url="https://doi.org/10.1007/s13225-015-0335-8" volume="77" year="2015">Zhou et al. 2015</bibRefCitation>
).
<taxonomicName id="9593665F2A4A44F9101CB12344E55438" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="ACE313B25FC7830F12A4A7AE2591B6F9" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="C4989550866730E10919384A41F807D3" lsidName="T. drechsleri" pageId="0" pageNumber="29" rank="species" species="drechsleri">
<emphasis id="8EC068E2DDC6E6DDD8B9DD5A532F3D83" italics="true" pageId="0" pageNumber="29">T. drechsleri</emphasis>
</taxonomicName>
share concentrically sulcate deeply-cracked pilei with mono-dimitic hyphal system, larger pores (&lt;6/mm) and presence of cystidioles, but the South American species differs in basidiospore size (4-5.5
<normalizedToken id="97247C47E8A131858A3A41F682495144" originalValue="×">x</normalizedToken>
3-4.5
<normalizedToken id="28DD91A828D881E6B15E4154B23252E6" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="804381AA5378D0B8092AA0ABBE7567FD" DOI="https://doi.org/10.11646/phytotaxa.338.1.6" author="Salvador-Montoya, CA" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="29" pagination="075 - 089" refId="B22" refString="Salvador-Montoya, CA, Costa-Rezende, DH, Ferreira-Lopes, V, Borba-Silva, MA, Popoff, OF, 2018. Tropicoporus drechsleri (Hymenochaetales, Basidiomycota), a new species in the &quot; Inonotus linteus &quot; complex from Northern Argentina. Phytotaxa 338: 075 - 089, DOI: https://doi.org/10.11646/phytotaxa.338.1.6" title="Tropicoporus drechsleri (Hymenochaetales, Basidiomycota), a new species in the &quot; Inonotus linteus &quot; complex from Northern Argentina." url="https://doi.org/10.11646/phytotaxa.338.1.6" volume="338" year="2018">Salvador-Montoya et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,113 @@
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<mods:title id="C8F5B13F63A1D74419331569B5A8F369">Review of the genus Tersilochus Holmgren (Hymenoptera, Ichneumonidae, Tersilochinae) from South Korea</mods:title>
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<treatment id="6208759AD960FD47DF47B88D617FFB13" ID-GBIF-Taxon="152017753" LSID="urn:lsid:zoobank.org:act:83AB92C8-1729-4ADA-9F15-286F4861DAE5" httpUri="http://treatment.plazi.org/id/6208759AD960FD47DF47B88D617FFB13" lastPageId="9" lastPageNumber="36" pageId="7" pageNumber="34">
<subSubSection id="D16DD6C4EEFA19985B35B704CB2166FE" pageId="7" pageNumber="34" type="nomenclature">
<paragraph id="026A6773F5E6846DB249836E733909C0" pageId="7" pageNumber="34">
<taxonomicName id="BB67401F0190FD8E24AD4FB6E08354B5" LSID="http://zoobank.org/83AB92C8-1729-4ADA-9F15-286F4861DAE5" authority="Khalaim &amp; Lee" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus (Tersilochus) iracundus" order="Hymenoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="iracundus" subGenus="Tersilochus">Tersilochus (Tersilochus) iracundus Khalaim &amp; Lee</taxonomicName>
<taxonomicNameLabel id="8B73E562334A4BBF820EDAD6FAF83068" pageId="7" pageNumber="34">sp. n.</taxonomicNameLabel>
Figs 18-27
</paragraph>
</subSubSection>
<subSubSection id="CABB84140ABC5E072A45A858EF43464C" lastPageId="9" lastPageNumber="36" pageId="7" pageNumber="34" type="description">
<paragraph id="39FC2E7081BD520A779D93626E996E10" pageId="7" pageNumber="34">Description.</paragraph>
<paragraph id="D99655AA16AFD9EDC795500A7F5BA942" pageId="7" pageNumber="34">Female (holotype). Body length 3.7 mm. Fore wing length 2.8 mm.</paragraph>
<paragraph id="AA80A4344D532171C255E2A9B552BECD" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34">
Head strongly rounded behind eyes in dorsal view (Fig. 19); temple 0.74 times as long as eye width. Inner eye orbits parallel (Fig. 23). Mandible with upper tooth much longer than lower tooth. Clypeus lenticular, almost 3.0 times as broad as long, in profile convex, with lower 0.4 bent backwards (Fig. 23); sparsely punctate, finely granulate, and dull in upper 0.7. Malar space 0.4 times as long as basal width of mandible. Flagellum of antenna weakly tapered towards apex, with 19 segments (Fig. 20); subbasal flagellomeres 1.3-1.4 times, and subapical flagellomeres about 1.2 times
<pageBreakToken id="938592AEE4C935A308C51A30A43234B2" pageId="8" pageNumber="35" start="start">as</pageBreakToken>
long as broad; flagellomeres 4-6 with distinct and flagellomere 7 with rudimental subapical finger-shaped structures on outer surface (Fig. 21). Face, frons, vertex, and temple distinctly granulate, dull, and impunctate (Figs 19, 23).
</paragraph>
<caption id="59784AE2DBCC07ECE01C871032C77BA3" pageId="8" pageNumber="35">
<paragraph id="4F6385D43824896D1FE73BB763E56969" pageId="8" pageNumber="35">
Figures 23-30.
<taxonomicName id="A39E51E9C6429BB2E77DE39C69FF5F22" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus iracundus" order="Hymenoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="iracundus">Tersilochus iracundus</taxonomicName>
sp. n., female, holotype: 23 head, frontal view 24 mesosoma, ventrolateral view 25 fore wings 26 metasoma with ovipositor, lateral view 27 tergites 1-3 of metasoma, dorsal view.
<taxonomicName id="6EE7EE21461DD3BBDFEBFDE61A7CB851" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus nigellus" order="Hymenoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="nigellus">Tersilochus nigellus</taxonomicName>
sp. n., female, holotype: 28 head with antenna and anterior part of mesosoma, lateral view 29 mesosoma, lateral view 30 propodeum and base of metasoma, dorsolateral view.
</paragraph>
</caption>
<paragraph id="EA5E2765BA364C530F5448A415B65FC3" pageId="9" pageNumber="36">
<pageBreakToken id="718AE4E7CDDC04C382C5A99838B6A748" pageId="9" pageNumber="36" start="start">Mesosoma</pageBreakToken>
entirely granulate, dull, impunctate; mesopleuron centrally with fine oblique striae on granulate background (Fig. 24). Notaulus absent. Foveate groove weak and short, oblique, situated in anterior half of mesopleuron (Fig. 24). Propodeum with narrow basal area, which is 0.4 times as long as apical area (Fig. 22). Propodeal spiracle separated from pleural carina by 1.5 times diameter of spiracle. Apical area slightly impressed, anteriorly rounded (Fig. 22). Apical longitudinal carinae developed in posterior half, anteriorly absent. Fore wing (Fig. 25) with intercubitus thickened, as long as abscissa of cubitus between intercubitus and second recurrent vein. First abscissa of radius slightly longer than width of pterostigma. Metacarpus ending far from apex of fore wing. Postnervulus intercepted below middle. Hind wing with nervellus vertical. First tergite almost 3.0 times as long as broad posteriorly, mostly smooth, with petiole more or less round in cross-section, well separated from postpetiole in dorsal view, finely striate laterally before glymma (Fig. 27). Glymma deep, situated somewhat behind center of first tergite, joining by distinct furrow to ventral part of postpetiole (Fig. 26). Second tergite as long as anteriorly broad (Fig. 27). Thyridial depression short, transverse. Ovipositor short, slender, almost straight basally and upcurved in apical 0.3, with fine teeth dorsally and ventrally at apex (Fig. 26); sheath 0.7 times as long as first tergite.
</paragraph>
<paragraph id="353BD09C879988D1537A21942D9FBF3F" pageId="9" pageNumber="36">Head, mesosoma, and first tergite black; palpi, mandible (except reddish black teeth), and lower 0.3 of clypeus yellow-brown; tegula yellow. Antenna dark brown. Pterostigma brown. Legs brownish yellow; fore and mid coxae weakly brown, and hind coxa strongly brown. Metasoma behind first tergite yellow-brown.</paragraph>
<paragraph id="C47448BFBCDCDF1BCC9C97742FCF6AF9" pageId="9" pageNumber="36">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D03B0777FBD4FABDB66D879F58B2CEEB" pageId="9" pageNumber="36" type="comparison">
<paragraph id="CF642D83E07A2D530BEEE52A0A4E75B9" pageId="9" pageNumber="36">Comparison.</paragraph>
<paragraph id="67509D0009D7564D9FF8B97F4C78E912" pageId="9" pageNumber="36">
Differs from other Palaearctic species of
<taxonomicName id="BE9CD147247ACC9076C14FA931CD7B60" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus" order="Hymenoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="genus">Tersilochus</taxonomicName>
by the combination short malar space, reddish brown behind first tergite of metasoma (Figs 26, 27) and short ovipositor (Fig. 26).
</paragraph>
</subSubSection>
<subSubSection id="1EC5A0D6BF1F3F79D8E1F829994F1F4F" pageId="9" pageNumber="36" type="type material">
<paragraph id="89C69654DE920C2464C4B25EE1C36A23" pageId="9" pageNumber="36">Type material.</paragraph>
<paragraph id="28A3CDAA85BFC415A391488C63BA1DE6" pageId="9" pageNumber="36">
Holotype female, South Korea, Chungbuk-do (CB), Jecheon-si, Deoksan-myeon, Worak-ri, Deoksanmaepyoso,
<geoCoordinate id="F4CE89986FF4B4032F83582DC3E9FA65" direction="north" orientation="latitude" precision="925" value="36.866665">36°52'N</geoCoordinate>
,
<geoCoordinate id="76CC461690A7A6D940AE4E93AE843352" direction="east" orientation="longitude" precision="925" value="128.21666">128°13'E</geoCoordinate>
, Malaise trap, 6-20.V.2006, coll. J.W. Lee (YUG).
</paragraph>
</subSubSection>
<subSubSection id="571EBABA336EEAE971BF9A4B8AA6C41D" pageId="9" pageNumber="36" type="distribution">
<paragraph id="603FC1573DE4751E398DE4DD4E0567B2" pageId="9" pageNumber="36">Distribution.</paragraph>
<paragraph id="2C025249658ED7CDB718D374BABAF19C" pageId="9" pageNumber="36">South Korea.</paragraph>
</subSubSection>
<subSubSection id="E83AB0B2FE257A79999682F119AB9C60" pageId="9" pageNumber="36" type="etymology">
<paragraph id="E5150D1BAA3804C058E94139EE9908BD" pageId="9" pageNumber="36">Etymology.</paragraph>
<paragraph id="FE58EB145CA2EA49EEAF37422D17AA9F" pageId="9" pageNumber="36">Named after the Latin iracundus (angry, hot-tempered, furious).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,215 @@
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<paragraph id="EA1E3692C301C75FE12E75F2FC12FE35" blockId="47.[264,936,483,510]" box="[264,936,483,510]" pageId="47" pageNumber="48">
<heading id="B15681FEC301C75FE12E75F2FC12FE35" bold="true" box="[264,936,483,510]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="D8D5EA80C301C75FE12E75F2FC12FE35" bold="true" box="[264,936,483,510]" pageId="47" pageNumber="48">
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13.
<taxonomicName id="2DA14D11C301C75FE11275F5FD20FE35" ID-CoL="6YBD9" authority="Gusarov" authorityName="Gusarov" box="[308,666,483,510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C301C75FE11275F5FD92FE36" bold="true" box="[308,552,484,510]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC301C75FE28175F2FD53FE35" box="[679,745,483,509]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C301C75FE2DF75F2FC1AFE35" box="[761,928,483,509]" captionStart-0="FIGURES 175 ­ 178" captionStart-1="FIGURES 179 ­ 189" captionStartId-0="47.[264,383,1629,1653]" captionStartId-1="48.[264,383,1595,1619]" captionTargetBox-0="[300,1285,812,1597]" captionTargetBox-1="[298,1302,460,1563]" captionTargetId-0="figure@47.[294,1294,804,1603]" captionTargetId-1="figure@48.[278,1310,449,1569]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionText-0="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 male tergum 8; 176 male sternum 8; 177 female tergum 8; 178 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri-0="https://zenodo.org/record/156451/files/figure.png" httpUri-1="https://zenodo.org/record/156452/files/figure.png" pageId="47" pageNumber="48">Figs. 175­189</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C301C75FE12E763FFECFFD09" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="EA1E3692C301C75FE12E763FFB8CFDB9" blockId="47.[264,1323,558,705]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE12E763FFE7AFD80" bold="true" box="[264,448,558,584]" pageId="47" pageNumber="48">
<typeStatus id="351A8830C301C75FE12E763FFEFFFD80" box="[264,325,558,584]" pageId="47" pageNumber="48">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C301C75FE1F7763EFDFBFD81" box="[465,577,559,585]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C301C75FE254763FFC53FD80" bold="true" box="[626,1001,558,584]" pageId="47" pageNumber="48">
<collectingCountry id="92B67602C301C75FE254763FFCFFFD80" box="[626,837,558,584]" name="El Salvador" pageId="47" pageNumber="48">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
Montecristo,
<quantity id="2D599B77C301C75FE4BF763EFB41FD81" box="[1177,1275,559,585]" metricMagnitude="4" metricUnit="m" metricValue="2.17" pageId="47" pageNumber="48" unit="km" value="21.7">21.7 km</quantity>
NE Metapan,
<quantity id="2D599B77C301C75FE15B7646FE63FDB9" box="[381,473,599,625]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="47" pageNumber="48" unit="m" value="2100.0">2100 m</quantity>
, forest litter (R.S.Anderson),
<date id="9E1F1052C301C75FE3127646FC78FDB9" box="[820,962,599,625]" pageId="47" pageNumber="48" value="1994-08-29">29.viii.1994</date>
(
<collectionCode id="8CB0AE57C301C75FE3F77646FB93FDB9" box="[977,1065,599,625]" pageId="47" pageNumber="48">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C301C75FE11E766EFECFFD09" blockId="47.[264,1323,558,705]" pageId="47" pageNumber="48">
<typeStatus id="351A8830C301C75FE11E766EFE15FD51" box="[312,431,639,665]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C301C75FE1EB766FFCECFD50" bold="true" box="[461,854,638,664]" pageId="47" pageNumber="48">
<collectingCountry id="92B67602C301C75FE1EB766FFD1EFD50" box="[461,676,638,664]" name="El Salvador" pageId="47" pageNumber="48">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
, 4, same data as the
<typeStatus id="351A8830C301C75FE4E2766EFA91FD51" box="[1220,1323,639,665]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C301C75FE13676B6FED2FD09" box="[272,360,679,705]" pageId="47" pageNumber="48">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC301C75FE12E724CFBEAF972" httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="47" pageNumber="48" targetBox="[300,1285,812,1597]" targetPageId="47">
<paragraph id="EA1E3692C301C75FE12E724CFBEAF972" blockId="47.[264,1322,1629,1722]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE12E724CFE65F9BD" bold="true" box="[264,479,1629,1653]" pageId="47" pageNumber="48">FIGURES 175­178</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C301C75FE2DA724CFB8FF9BC" authority="Gusarov" authorityName="Gusarov" box="[764,1077,1629,1652]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C301C75FE2DA724CFC69F9BC" box="[764,979,1629,1652]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC301C75FE466724CFBC6F9BD" box="[1088,1148,1629,1653]" pageId="47" pageNumber="48" rank="species">
<emphasis id="D8D5EA80C301C75FE466724CFBC6F9BD" bold="true" box="[1088,1148,1629,1653]" pageId="47" pageNumber="48">sp. n.</emphasis>
</taxonomicNameLabel>
(175­176, holotype; 177­178, paratype, 21.7 km NE Metapan, El Salvador). 175 male tergum 8; 176 male sternum 8; 177 female tergum 8; 178 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C301C741E11E731BFDBAFDEE" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="EA1E3692C301C740E11E731BFC4BFE4E" blockId="47.[264,1323,1802,1869]" lastBlockId="48.[264,1324,284,390]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE11E731BFE02F8EC" bold="true" box="[312,440,1802,1828]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C301C75FE1EA731AFD07F8EC" box="[460,701,1803,1828]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C301C75FE1EA731AFD07F8EC" box="[460,701,1803,1828]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C301C75FE4F5731AFEFDF884" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C301C75FE4F5731AFEFDF884" italics="true" pageId="47" pageNumber="48">Seeversiella</emphasis>
</taxonomicName>
by having brownish yellow; small eyes (temple length to eye length ratio 4.0­7.0); glossy pronotum with fine and weak microsculpture; elytra much shorter than pronotum; reduced wings; tergum 8 with four pairs of macrosetae; the shape of aedeagus (
<figureCitation id="729A2A17C31EC740E4947555FE1CFE4E" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49">Figs. 179­ 183, 185­189</figureCitation>
) and the distinct shape of spermatheca (
<figureCitation id="729A2A17C31EC740E35A757DFC59FE4E" box="[892,995,364,390]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49">Fig. 184</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31EC740E12E722AFECBF892" httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49" targetBox="[298,1302,460,1563]" targetPageId="48">
<paragraph id="EA1E3692C31EC740E12E722AFECBF892" blockId="48.[264,1323,1595,1882]" pageId="48" pageNumber="49">
<emphasis id="D8D5EA80C31EC740E12E722AFE5BF99B" bold="true" box="[264,481,1595,1619]" pageId="48" pageNumber="49">FIGURES 179­189</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C31EC740E2A2722DFC78F99B" authority="Gusarov" authorityName="Gusarov" box="[644,962,1596,1619]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C31EC740E2A2722DFCE5F99B" box="[644,863,1596,1619]" italics="true" pageId="48" pageNumber="49">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31EC740E3F6722AFBB4F99B" box="[976,1038,1595,1619]" pageId="48" pageNumber="49" rank="species">
<emphasis id="D8D5EA80C31EC740E3F6722AFBB4F99B" bold="true" box="[976,1038,1595,1619]" pageId="48" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
(179­183, 185­186, holotype; 184, 187­189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185­186), 0.1 mm (180, 182­184, 187­189).
</paragraph>
</caption>
<paragraph id="EA1E3692C31FC741E11E750DFDBAFDEE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<taxonomicName id="2DA14D11C31FC741E11E750DFD9EFEFD" box="[312,548,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C31FC741E11E750DFD9EFEFD" box="[312,548,284,309]" italics="true" pageId="49" pageNumber="50">Seeversiella minima</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C31FC741E378750DFC76FEFD" box="[862,972,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C31FC741E378750DFC76FEFD" box="[862,972,284,309]" italics="true" pageId="49" pageNumber="50">S. flavida</emphasis>
</taxonomicName>
and other similar species (24­ 28) with small body and very short elytra by the combination of the following characters: pubescence in lateral portions of pronotum directed towards midline and obliquely posteriorly; tergum 8 with four pairs of macrosetae; proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C31FC741E2F875ADFCFCFE1E" box="[734,838,444,470]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 183</figureCitation>
); apex of median lobe in parameral view wide (
<figureCitation id="729A2A17C31FC741E17275F5FE45FE36" box="[340,511,484,510]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Figs. 179­180</figureCitation>
); spermatheca J­shaped with bulbous proximal portion and without umbilicus (
<figureCitation id="729A2A17C31FC741E1AA761DFE48FDEE" box="[396,498,524,550]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31FC741E11E7622FDC7FA5E" pageId="49" pageNumber="50" type="description">
<paragraph id="EA1E3692C31FC741E11E7622FE7CFDBE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E7622FE6AFD85" bold="true" box="[312,464,563,589]" pageId="49" pageNumber="50">Description.</emphasis>
Length 1.6­2.0 mm. Body brownish­yellow with yellow legs, antennae and mouthparts.
</paragraph>
<paragraph id="EA1E3692C31FC741E11E7695FD35FCDE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">Head surface glossy, with fine and weak isodiametric microsculpture, fine and sparse punctation, distance between punctures equals 3­4 times their diameter. Temple length to eye length ratio 4.0­7.0. Antennal article 2 as long as 3, article 4 transverse (ratio 1.5), 5­ 10 strongly transverse (ratio 2.0).</paragraph>
<paragraph id="EA1E3692C31FC741E11E7735FE7EFBE6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C31FC741E40D7735FB6CFCF6" box="[1067,1238,804,830]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.0" metricValueMin="3.4" pageId="49" pageNumber="50" unit="mm" value="0.37" valueMax="0.4" valueMin="0.34">0.34­0.40 mm</quantity>
, length
<quantity id="2D599B77C31FC741E12E775DFE08FCAE" box="[264,434,844,870]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5999999999999996" metricValueMin="3.1" pageId="49" pageNumber="50" unit="mm" value="0.33499999999999996" valueMax="0.36" valueMin="0.31">0.31­0.36 mm</quantity>
, width to length ratio 1.1, surface glossy, microsculpture and punctation as on head. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.9 times wider than long, glossy, with fine and poorly visible isodiametric microsculpture; punctation asperate and stronger than on pronotum, distance between punctures equals 2­3 times their diameter. Wings shorter than elytra or absent.
</paragraph>
<paragraph id="EA1E3692C31FC741E11E702DFC1BFB06" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C31FC741E2C170A5FC2EFB06" box="[743,916,1204,1230]" captionStart="FIGURES 175 ­ 178" captionStartId="47.[264,383,1629,1653]" captionTargetBox="[300,1285,812,1597]" captionTargetId="figure@47.[294,1294,804,1603]" captionTargetPageId="47" captionText="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 male tergum 8; 176 male sternum 8; 177 female tergum 8; 178 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="49" pageNumber="50">Figs. 175, 177</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E70CDFBD7FAD6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
In males posterior angles of tergum 3 not projecting, tergum 7 without medial carina or tubercle, posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C31FC741E3DF7115FBDAFAD6" box="[1017,1120,1284,1310]" captionStart="FIGURES 175 ­ 178" captionStartId="47.[264,383,1629,1653]" captionTargetBox="[300,1285,812,1597]" captionTargetId="figure@47.[294,1294,804,1603]" captionTargetPageId="47" captionText="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 male tergum 8; 176 male sternum 8; 177 female tergum 8; 178 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="49" pageNumber="50">Fig. 175</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E713DFCF6FAA6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Aedeagus as in
<figureCitation id="729A2A17C31FC741E1DB713DFCA6FA8E" box="[509,796,1324,1350]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Figs. 179­183, 185­189</figureCitation>
. Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C31FC741E2FE7145FC85FAA6" box="[728,831,1364,1390]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 183</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E716DFDC7FA5E" blockId="49.[264,1324,284,1550]" box="[312,637,1404,1430]" pageId="49" pageNumber="50">
Spermatheca as in
<figureCitation id="729A2A17C31FC741E232716DFDC3FA5E" box="[532,633,1404,1430]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 median lobe, parameral view; 180 apex of median lobe, parameral view; 181 median lobe, lateral view; 182 apex of median lobe, lateral view; 183 apex of left paramere; 184 spermatheca; 185 details of internal sac retracted into median lobe, lateral view; 186 details of internal sac retracted into median lobe, abparameral view; 187 medial lamellae, parameral view; 188 medial lamellae, lateral view; 189 copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 184</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31FC741E11E71B2FE9CF9C6" pageId="49" pageNumber="50" type="distribution">
<paragraph id="EA1E3692C31FC741E11E71B2FBC2FA76" blockId="49.[264,1324,284,1550]" box="[312,1144,1443,1470]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E71B2FE62FA75" bold="true" box="[312,472,1443,1469]" pageId="49" pageNumber="50">Distribution.</emphasis>
Known from Cerro Montecristo,
<collectingCountry id="92B67602C31FC741E34D71B5FC4EFA76" box="[875,1012,1444,1470]" name="El Salvador" pageId="49" pageNumber="50">El Salvador</collectingCountry>
(
<figureCitation id="729A2A17C31FC741E42271B5FBD1FA76" box="[1028,1131,1444,1470]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="49" pageNumber="50">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E71DAFE9CF9C6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E71DAFDBFFA2D" bold="true" box="[312,517,1483,1509]" pageId="49" pageNumber="50">Natural History.</emphasis>
<taxonomicName id="2DA14D11C31FC741E23571DDFCBAFA2D" box="[531,768,1484,1509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C31FC741E23571DDFCBAFA2D" box="[531,768,1484,1509]" italics="true" pageId="49" pageNumber="50">Seeversiella minima</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C31FC741E4C971DDFE98F9C6" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="49" pageNumber="50" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,247 @@
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<mods:title id="B5EDD83F84BC62BC69BA65F86F9CF023">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C302C75CE12E775AFC23FCAD" box="[264,921,843,870]" pageId="44" pageNumber="45" type="key">
<paragraph id="EA1E3692C302C75CE12E775AFC23FCAD" blockId="44.[264,921,843,870]" box="[264,921,843,870]" pageId="44" pageNumber="45">
<heading id="B15681FEC302C75CE12E775AFC23FCAD" bold="true" box="[264,921,843,870]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<keyLead id="515095A7C302C75CE12E775AFC23FCAD" box="[264,921,843,870]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE12E775AFC23FCAD" bold="true" box="[264,921,843,870]" pageId="44" pageNumber="45">
<keyStep id="51552E37C302C75CE12E775AFC23FCAD" box="[264,921,843,870]" pageId="44" pageNumber="45">
<key id="9E3061A3C302C75CE12E775AFC23FCAD" box="[264,921,843,870]" pageId="44" pageNumber="45">
12.
<taxonomicName id="2DA14D11C302C75CE112775DFD31FCAD" ID-CoL="6YBDB" authority="Gusarov" authorityName="Gusarov" box="[308,651,843,870]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C302C75CE112775DFDA3FCAE" bold="true" box="[308,537,844,870]" italics="true" pageId="44" pageNumber="45">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC302C75CE2BE775AFD60FCAD" box="[664,730,843,869]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C302C75CE2CC775AFC2BFCAD" box="[746,913,843,869]" captionStart-0="FIGURES 161 ­ 164" captionStart-1="FIGURES 165 ­ 168" captionStart-2="FIGURES 169 ­ 174" captionStartId-0="44.[264,383,1760,1784]" captionStartId-1="45.[264,383,1583,1607]" captionStartId-2="46.[264,383,1661,1685]" captionTargetBox-0="[317,1270,975,1727]" captionTargetBox-1="[302,1289,779,1549]" captionTargetBox-2="[288,1298,900,1628]" captionTargetId-0="figure@44.[309,1279,964,1734]" captionTargetId-1="figure@45.[292,1295,769,1556]" captionTargetId-2="figure@46.[282,1304,889,1634]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIGURES 161 ­ 164. Abdominal segment 8 of Seeversiella similis Gusarov, sp. n. (161 ­ 162, holotype; 163 ­ 164, paratype from La Tigra, Honduras). 161 male tergum 8; 162 male sternum 8; 163 female tergum 8; 164 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-2="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156448/files/figure.png" httpUri-1="https://zenodo.org/record/156449/files/figure.png" httpUri-2="https://zenodo.org/record/156450/files/figure.png" pageId="44" pageNumber="45">Figs. 161­174</figureCitation>
)
</key>
</keyStep>
</emphasis>
</keyLead>
</heading>
</paragraph>
</subSubSection>
<caption id="BEDE661AC302C75CE12E72F1FC63F8F5" httpUri="https://zenodo.org/record/156448/files/figure.png" pageId="44" pageNumber="45" targetBox="[317,1270,975,1727]" targetPageId="44">
<paragraph id="EA1E3692C302C75CE12E72F1FC63F8F5" blockId="44.[264,1323,1760,1853]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE12E72F1FE65F930" bold="true" box="[264,479,1760,1784]" pageId="44" pageNumber="45">FIGURES 161­164</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C302C75CE32772F0FB89F930" authority="Gusarov" authorityName="Gusarov" box="[769,1075,1761,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C302C75CE32772F0FC75F930" box="[769,975,1761,1784]" italics="true" pageId="44" pageNumber="45">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C302C75CE41872F1FBC0F930" bold="true" box="[1086,1146,1760,1784]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="C3E657FBC302C75CE41872F1FBC0F930" box="[1086,1146,1760,1784]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(161­162, holotype; 163­164, paratype from La Tigra, Honduras). 161 male tergum 8; 162 male sternum 8; 163 female tergum 8; 164 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C303C75DE12E750AFD85FE36" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="EA1E3692C303C75DE12E750AFECFFE4E" blockId="45.[264,1324,283,710]" pageId="45" pageNumber="46">
<emphasis id="D8D5EA80C303C75DE12E750AFE7AFEFD" bold="true" box="[264,448,283,309]" pageId="45" pageNumber="46">
<typeStatus id="351A8830C303C75DE12E750AFEF9FEFD" box="[264,323,283,309]" pageId="45" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C303C75DE1FC750DFDF0FEFE" box="[474,586,284,310]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C303C75DE2A7750AFBF6FEFD" bold="true" box="[641,1100,283,309]" pageId="45" pageNumber="46">
<collectingCountry id="92B67602C303C75DE2A7750AFC97FEFD" box="[641,813,283,309]" name="Honduras" pageId="45" pageNumber="46">HONDURAS</collectingCountry>
: Francisco Morazán:
</emphasis>
<quantity id="2D599B77C303C75DE442750DFB72FEFE" box="[1124,1224,284,310]" metricMagnitude="4" metricUnit="m" metricValue="2.13" pageId="45" pageNumber="46" unit="km" value="21.3">21.3 km</quantity>
N Teguicigalpa, La Tigra,
<quantity id="2D599B77C303C75DE2237555FDDCFE96" box="[517,614,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="m" value="2100.0">2100 m</quantity>
,
<geoCoordinate id="8F955055C303C75DE2517555FD62FE96" box="[631,728,324,350]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="14.2">14°12'N</geoCoordinate>
<geoCoordinate id="8F955055C303C75DE2C37555FCF6FE96" box="[741,844,324,350]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-86.1">86°06'W</geoCoordinate>
, forest litter (R.S.Anderson),
<date id="9E1F1052C303C75DE49B7555FA96FE96" box="[1213,1324,324,350]" pageId="45" pageNumber="46" value="1994-06-07">7.vi.1994</date>
(
<collectionCode id="8CB0AE57C303C75DE136757DFED2FE4E" box="[272,360,364,390]" pageId="45" pageNumber="46">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C303C75DE11E7585FD85FE36" blockId="45.[264,1324,283,710]" pageId="45" pageNumber="46">
<typeStatus id="351A8830C303C75DE11E7585FE15FE66" box="[312,431,404,430]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C303C75DE1E17582FC37FE65" bold="true" box="[455,909,403,429]" pageId="45" pageNumber="46">
<collectingCountry id="92B67602C303C75DE1E17582FDCEFE65" box="[455,628,403,429]" name="Honduras" pageId="45" pageNumber="46">HONDURAS</collectingCountry>
: Francisco Morazán:
</emphasis>
4, same data as the
<typeStatus id="351A8830C303C75DE49A7585FA9CFE66" box="[1212,1318,404,430]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
; 2, ditto but
<date id="9E1F1052C303C75DE1EA75ADFDE7FE1E" box="[460,605,444,470]" pageId="45" pageNumber="46" value="1994-08-15">15.viii.1994</date>
; 2, 4, ditto but
<quantity id="2D599B77C303C75DE35875ADFC5CFE1E" box="[894,998,444,470]" metricMagnitude="4" metricUnit="m" metricValue="2.2199999999999998" pageId="45" pageNumber="46" unit="km" value="22.2">22.2 km</quantity>
N Teguicigalpa,
<quantity id="2D599B77C303C75DE4E275ADFA9DFE1E" box="[1220,1319,444,470]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="45" pageNumber="46" unit="m" value="2030.0">2030 m</quantity>
,
<date id="9E1F1052C303C75DE12E75F5FE2CFE36" box="[264,406,484,510]" pageId="45" pageNumber="46" value="1994-08-15">15.viii.1994</date>
(all ­
<collectionCode id="8CB0AE57C303C75DE1FB75F5FD8EFE36" box="[477,564,484,510]" pageId="45" pageNumber="46">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C303C75DE11E761AFB7FF8E9" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="EA1E3692C303C75DE11E761AFB6CFD0E" blockId="45.[264,1324,283,710]" pageId="45" pageNumber="46">
<emphasis id="D8D5EA80C303C75DE11E761AFE02FDED" bold="true" box="[312,440,523,549]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C303C75DE1E3761DFD1CFDED" box="[453,678,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE1E3761DFD1CFDED" box="[453,678,524,549]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C303C75DE4BB761DFA91FDED" box="[1181,1323,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C303C75DE4BB761DFA91FDED" box="[1181,1323,524,549]" italics="true" pageId="45" pageNumber="46">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.4­1.8 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra as long or slightly shorter than pronotum (elytral length to pronotal length ratio 1.0­0.9); reduced wings; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C303C75DE2FE76BDFCC7FD0E" box="[728,893,684,710]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="45" pageNumber="46">Figs. 165­173</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C303C75DE44476BDFB72FD0E" box="[1122,1224,684,710]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="45" pageNumber="46">Fig. 174</figureCitation>
).
</paragraph>
<caption id="BEDE661AC303C75DE12E723EFB71F943" httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="45" pageNumber="46" targetBox="[302,1289,779,1549]" targetPageId="45">
<paragraph id="EA1E3692C303C75DE12E723EFB71F943" blockId="45.[264,1322,1583,1675]" pageId="45" pageNumber="46">
<emphasis id="D8D5EA80C303C75DE12E723EFE5AF98F" bold="true" box="[264,480,1583,1607]" pageId="45" pageNumber="46">FIGURES 165­168</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C303C75DE2AE723EFC01F98E" authority="Gusarov" authorityName="Gusarov" box="[648,955,1583,1606]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C303C75DE2AE723EFCE2F98E" box="[648,856,1583,1606]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C303C75DE3EE723EFBBCF98F" bold="true" box="[968,1030,1583,1607]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="C3E657FBC303C75DE3EE723EFBBCF98F" box="[968,1030,1583,1607]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168).
</paragraph>
</caption>
<paragraph id="EA1E3692C303C75DE11E72CEFB7FF8E9" blockId="45.[264,1323,1759,1905]" pageId="45" pageNumber="46">
<taxonomicName id="2DA14D11C303C75DE11E72CEFDA0F930" box="[312,538,1759,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE11E72CEFDA0F930" box="[312,538,1759,1784]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C303C75DE34372CEFC7CF930" box="[869,966,1759,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C303C75DE34372CEFC7CF930" box="[869,966,1759,1784]" italics="true" pageId="45" pageNumber="46">S. badia</emphasis>
</taxonomicName>
in that apex of median lobe in lateral view lacks clear emargination on parameral side (
<figureCitation id="729A2A17C303C75DE3B97316FBFCF8E9" box="[927,1094,1799,1825]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="45" pageNumber="46">Figs. 167­168</figureCitation>
; 156­157).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C303C75EE11E733EFC0CF889" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="EA1E3692C303C75EE11E733EFCDCFE66" blockId="45.[264,1323,1759,1905]" lastBlockId="46.[264,1324,284,790]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="2DA14D11C303C75DE11E733EFDA0F880" box="[312,538,1839,1864]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE11E733EFDA0F880" box="[312,538,1839,1864]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C303C75DE37A733EFC4CF880" box="[860,1014,1839,1864]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C303C75DE37A733EFC4CF880" box="[860,1014,1839,1864]" italics="true" pageId="45" pageNumber="46">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 10) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have small spicules (
<figureCitation id="729A2A17C300C75EE2DD7555FCDFFE96" box="[763,869,324,350]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 173</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C300C75EE3F1757DFB3BFE4E" box="[983,1153,364,390]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="46" pageNumber="47">Figs. 167­168</figureCitation>
); spermatheca S­shaped with umbilicus facing laterally (
<figureCitation id="729A2A17C300C75EE2D47585FCE3FE66" box="[754,857,404,430]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C300C75EE11E75AAFC17FE36" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">
<emphasis id="D8D5EA80C300C75EE11E75AAFE6AFE1D" bold="true" box="[312,464,443,469]" pageId="46" pageNumber="47">Description.</emphasis>
Length
<quantity id="2D599B77C300C75EE21F75ADFD7FFE1E" box="[569,709,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="46" pageNumber="47" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C300C75EE3A575ADFB96FE1E" box="[899,1068,444,470]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="46" pageNumber="47" unit="mm" value="0.51" valueMax="0.54" valueMin="0.48">0.48­0.54 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E761DFC52FD86" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">
In all character states
<taxonomicName id="2DA14D11C300C75EE211761DFD1BFDED" box="[567,673,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C300C75EE211761DFD1BFDED" box="[567,673,524,549]" italics="true" pageId="46" pageNumber="47">S. similis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C300C75EE350761DFBB5FDED" box="[886,1039,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C300C75EE350761DFBB5FDED" box="[886,1039,524,549]" italics="true" pageId="46" pageNumber="47">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E764DFDE1FD0E" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">In males posterior angles of tergum 3 projecting as spines, up to 1 time as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C300C75EE11E76C5FD21FD26" blockId="46.[264,1324,284,790]" box="[312,667,724,750]" pageId="46" pageNumber="47">
Aedeagus as in
<figureCitation id="729A2A17C300C75EE1D776C5FD2DFD26" box="[497,663,724,750]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Figs. 165­173</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E76EDFDC4FCDE" blockId="46.[264,1324,284,790]" box="[312,638,764,790]" pageId="46" pageNumber="47">
Spermatheca as in
<figureCitation id="729A2A17C300C75EE23376EDFDC0FCDE" box="[533,634,764,790]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
.
</paragraph>
<caption id="BEDE661AC300C75EE12E726CFCCCF8D7" httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47" targetBox="[288,1298,900,1628]" targetPageId="46">
<paragraph id="EA1E3692C300C75EE12E726CFCCCF8D7" blockId="46.[264,1323,1661,1857]" pageId="46" pageNumber="47">
<emphasis id="D8D5EA80C300C75EE12E726CFE64F95D" bold="true" box="[264,478,1661,1685]" pageId="46" pageNumber="47">FIGURES 169­174</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C300C75EE253726FFC19F95C" authority="Gusarov" authorityName="Gusarov" box="[629,931,1661,1685]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C300C75EE253726FFCFBF95D" box="[629,833,1662,1685]" italics="true" pageId="46" pageNumber="47">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C300C75EE38B726CFC52F95D" bold="true" box="[941,1000,1661,1685]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3E657FBC300C75EE38B726CFC52F95D" box="[941,1000,1661,1685]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(171, 173, holotype; 169­170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172­173), 0.1 mm (170­171, 174).
</paragraph>
</caption>
<paragraph id="EA1E3692C300C75EE12E733BFC0CF889" blockId="46.[264,1323,1661,1857]" box="[264,950,1834,1857]" pageId="46" pageNumber="47">CP copulatory piece; LD lateral diverticulum of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C301C75FE11E750AFED3FE4E" pageId="47" pageNumber="48" type="distribution">
<paragraph id="EA1E3692C301C75FE11E750AFED3FE4E" blockId="47.[264,1323,283,390]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="47" pageNumber="48">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C301C75FE2A3750DFD4DFEFE" box="[645,759,284,310]" name="Honduras" pageId="47" pageNumber="48">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C301C75FE321750DFCD5FEFE" box="[775,879,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="47" pageNumber="48">Fig. 387</figureCitation>
).
<emphasis id="D8D5EA80C301C75FE11E7552FDBFFE95" bold="true" box="[312,517,323,349]" pageId="47" pageNumber="48">Natural History.</emphasis>
<taxonomicName id="2DA14D11C301C75FE2327555FD4CFE95" box="[532,758,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C301C75FE2327555FD4CFE95" box="[532,758,324,349]" italics="true" pageId="47" pageNumber="48">Seeversiella similis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C301C75FE4C07555FEDEFE4E" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="47" pageNumber="48" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000­ 2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,223 @@
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<subSubSection id="A2BB6519C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43" type="key">
<paragraph id="EA1E3692C304C75AE12E750AFC36FEFD" blockId="42.[264,908,283,310]" box="[264,908,283,310]" pageId="42" pageNumber="43">
<heading id="B15681FEC304C75AE12E750AFC36FEFD" bold="true" box="[264,908,283,310]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="D8D5EA80C304C75AE12E750AFC36FEFD" bold="true" box="[264,908,283,310]" pageId="42" pageNumber="43">
<key id="9E3061A3C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43">
<keyStep id="51552E37C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43">
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11.
<taxonomicName id="2DA14D11C304C75AE115750DFDC4FEFD" ID-CoL="6YBCN" authority="Gusarov" authorityName="Gusarov" box="[307,638,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C304C75AE115750DFDB6FEFE" bold="true" box="[307,524,284,310]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC304C75AE2AD750AFD74FEFD" box="[651,718,283,309]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C304C75AE2FB750AFC3EFEFD" box="[733,900,283,309]" captionStart-0="FIGURES 150 ­ 153" captionStart-1="FIGURES 154 ­ 160" captionStartId-0="42.[264,382,1453,1477]" captionStartId-1="43.[264,383,1715,1739]" captionTargetBox-0="[286,1300,613,1419]" captionTargetBox-1="[292,1295,611,1684]" captionTargetId-0="figure@42.[279,1308,604,1426]" captionTargetId-1="figure@43.[281,1307,601,1691]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 150 ­ 153. Abdominal segment 8 of Seeversiella badia Gusarov, sp. n. (150 ­ 151, holotype; 152 ­ 153, paratype, 21.7 km NE Metapan, El Salvador). 150 male tergum 8; 151 male sternum 8; 152 female tergum 8; 153 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri-0="https://zenodo.org/record/156446/files/figure.png" httpUri-1="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 150­160</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C304C75AE12E7577FED1FE31" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="EA1E3692C304C75AE12E7577FBC7FE61" blockId="42.[264,1323,358,505]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="42" pageNumber="43">
<typeStatus id="351A8830C304C75AE12E7577FEFFFE48" box="[264,325,358,384]" pageId="42" pageNumber="43">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C304C75AE1F77576FDFBFE49" box="[465,577,359,385]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C304C75AE2547577FC53FE48" bold="true" box="[626,1001,358,384]" pageId="42" pageNumber="43">
<collectingCountry id="92B67602C304C75AE2547577FCFFFE48" box="[626,837,358,384]" name="El Salvador" pageId="42" pageNumber="43">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
Montecristo,
<quantity id="2D599B77C304C75AE4BF7576FB41FE49" box="[1177,1275,359,385]" metricMagnitude="4" metricUnit="m" metricValue="2.17" pageId="42" pageNumber="43" unit="km" value="21.7">21.7 km</quantity>
NE Metapan,
<quantity id="2D599B77C304C75AE15B759EFE63FE61" box="[381,473,399,425]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="42" pageNumber="43" unit="m" value="2100.0">2100 m</quantity>
, cloud forest litter (R.S.Anderson),
<date id="9E1F1052C304C75AE35D759EFBB2FE61" box="[891,1032,399,425]" pageId="42" pageNumber="43" value="1994-08-29">29.viii.1994</date>
(
<collectionCode id="8CB0AE57C304C75AE43E759EFBCAFE61" box="[1048,1136,399,425]" pageId="42" pageNumber="43">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C304C75AE11E75A6FED1FE31" blockId="42.[264,1323,358,505]" pageId="42" pageNumber="43">
<typeStatus id="351A8830C304C75AE11E75A6FE15FE19" box="[312,431,439,465]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C304C75AE1EE75A7FCF2FE18" bold="true" box="[456,840,438,464]" pageId="42" pageNumber="43">
<collectingCountry id="92B67602C304C75AE1EE75A7FD27FE18" box="[456,669,438,464]" name="El Salvador" pageId="42" pageNumber="43">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
2, same data as the
<typeStatus id="351A8830C304C75AE45B75A6FB5EFE19" box="[1149,1252,439,465]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(all ­
<collectionCode id="8CB0AE57C304C75AE12E75CEFEE4FE31" box="[264,350,479,505]" pageId="42" pageNumber="43">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC304C75AE12E71BCFBEAF9C1" httpUri="https://zenodo.org/record/156446/files/figure.png" pageId="42" pageNumber="43" targetBox="[286,1300,613,1419]" targetPageId="42">
<paragraph id="EA1E3692C304C75AE12E71BCFBEAF9C1" blockId="42.[264,1322,1453,1545]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE12E71BCFE65FA0D" bold="true" box="[264,479,1453,1477]" pageId="42" pageNumber="43">FIGURES 150­153</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C304C75AE32F71BCFB88FA0C" authority="Gusarov" authorityName="Gusarov" box="[777,1074,1453,1476]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C304C75AE32F71BCFC75FA0C" box="[777,975,1453,1476]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C304C75AE41B71BCFBC0FA0D" bold="true" box="[1085,1146,1453,1477]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="C3E657FBC304C75AE41B71BCFBC0FA0D" box="[1085,1146,1453,1477]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(150­151, holotype; 152­153, paratype, 21.7 km NE Metapan, El Salvador). 150 male tergum 8; 151 male sternum 8; 152 female tergum 8; 153 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C304C75AE11E724EFB26F88A" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="EA1E3692C304C75AE11E724EFB6CF93A" blockId="42.[264,1324,1631,1858]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE11E724EFE02F9B1" bold="true" box="[312,440,1631,1657]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C304C75AE1E17271FD25F9B1" box="[455,671,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C304C75AE1E17271FD25F9B1" box="[455,671,1632,1657]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C304C75AE4BB7271FA91F9B1" box="[1181,1323,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C304C75AE4BB7271FA91F9B1" box="[1181,1323,1632,1657]" italics="true" pageId="42" pageNumber="43">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.7­2.0 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra as long as pronotum; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C304C75AE2FE72C9FCC7F93A" box="[728,893,1752,1778]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 154­159</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C304C75AE44472C9FB72F93A" box="[1122,1224,1752,1778]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Fig. 160</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C304C75AE11E7311FB26F88A" blockId="42.[264,1324,1631,1858]" pageId="42" pageNumber="43">
<taxonomicName id="2DA14D11C304C75AE11E7311FDA9F8D1" box="[312,531,1792,1817]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C304C75AE11E7311FDA9F8D1" box="[312,531,1792,1817]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C304C75AE3567311FC67F8D1" box="[880,989,1792,1817]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C304C75AE3567311FC67F8D1" box="[880,989,1792,1817]" italics="true" pageId="42" pageNumber="43">S. similis</emphasis>
</taxonomicName>
in that the apex of median lobe in lateral view is emarginate on parameral side (
<figureCitation id="729A2A17C304C75AE3537339FBA7F88A" box="[885,1053,1832,1858]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 156­157</figureCitation>
; 167­168).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C305C75CE11E750DFDC4FDBE" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="EA1E3692C305C75BE11E750DFCF6FE36" blockId="43.[264,1324,284,510]" pageId="43" pageNumber="44">
<taxonomicName id="2DA14D11C305C75BE11E750DFDABFEFD" box="[312,529,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C305C75BE11E750DFDABFEFD" box="[312,529,284,309]" italics="true" pageId="43" pageNumber="44">Seeversiella badia</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C305C75BE371750DFC48FEFD" box="[855,1010,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C305C75BE371750DFC48FEFD" box="[855,1010,284,309]" italics="true" pageId="43" pageNumber="44">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 10) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have small spicules (
<figureCitation id="729A2A17C305C75BE2DD7585FCDFFE66" box="[763,869,404,430]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Fig. 158</figureCitation>
); apex of median lobe in lateral view widened subapically and emarginate on parameral side (
<figureCitation id="729A2A17C305C75BE39775ADFBE1FE1E" box="[945,1115,444,470]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Figs. 156­157</figureCitation>
); spermatheca Sshaped with umbilicus facing laterally (
<figureCitation id="729A2A17C305C75BE2FE75F5FC84FE36" box="[728,830,484,510]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Fig. 160</figureCitation>
).
</paragraph>
<caption id="BEDE661AC305C75BE12E72A2FB46F89D" httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44" targetBox="[292,1295,611,1684]" targetPageId="43">
<paragraph id="EA1E3692C305C75BE12E72A2FB46F89D" blockId="43.[264,1324,1715,1877]" pageId="43" pageNumber="44">
<emphasis id="D8D5EA80C305C75BE12E72A2FE59F903" bold="true" box="[264,483,1715,1739]" pageId="43" pageNumber="44">FIGURES 154­160</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C305C75BE2A872A5FC07F902" authority="Gusarov" authorityName="Gusarov" box="[654,957,1715,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C305C75BE2A872A5FCE2F903" box="[654,856,1716,1739]" italics="true" pageId="43" pageNumber="44">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C305C75BE3EB72A2FBB7F903" bold="true" box="[973,1037,1715,1739]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="C3E657FBC305C75BE3EB72A2FBB7F903" box="[973,1037,1715,1739]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(154­159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159­160).
</paragraph>
</caption>
<paragraph id="EA1E3692C302C75CE11E750AFC17FE96" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="44" pageNumber="45">Description.</emphasis>
Length
<quantity id="2D599B77C302C75CE262750DFD1FFEFE" box="[580,677,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="44" pageNumber="45" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C302C75CE348750DFBA1FEFE" box="[878,1051,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="44" pageNumber="45" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E757DFC52FE66" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
In all character states
<taxonomicName id="2DA14D11C302C75CE21C757DFD21FE4D" box="[570,667,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C302C75CE21C757DFD21FE4D" box="[570,667,364,389]" italics="true" pageId="44" pageNumber="45">S. badia</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C302C75CE355757DFBB6FE4D" box="[883,1036,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C302C75CE355757DFBB6FE4D" box="[883,1036,364,389]" italics="true" pageId="44" pageNumber="45">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E75ADFCE8FDEE" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">In the only known male specimen the posterior angles of tergum 3 projecting as spines, as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C302C75CE11E7625FD26FD86" blockId="44.[264,1323,283,750]" box="[312,668,564,590]" pageId="44" pageNumber="45">
Aedeagus as in
<figureCitation id="729A2A17C302C75CE1D77625FD2DFD86" box="[497,663,564,590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="44" pageNumber="45">Figs. 154­159</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E764DFDC4FDBE" blockId="44.[264,1323,283,750]" box="[312,638,604,630]" pageId="44" pageNumber="45">
Spermatheca as in
<figureCitation id="729A2A17C302C75CE233764DFDC1FDBE" box="[533,635,604,630]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="44" pageNumber="45">Fig. 160</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C302C75CE11E7692FED2FD26" pageId="44" pageNumber="45" type="distribution">
<paragraph id="EA1E3692C302C75CE11E7692FC29FD56" blockId="44.[264,1323,283,750]" box="[312,915,643,670]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E7692FE62FD55" bold="true" box="[312,472,643,669]" pageId="44" pageNumber="45">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C302C75CE2A37695FCB4FD56" box="[645,782,644,670]" name="El Salvador" pageId="44" pageNumber="45">El Salvador</collectingCountry>
(
<figureCitation id="729A2A17C302C75CE3397695FC3DFD56" box="[799,903,644,670]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="44" pageNumber="45">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C302C75CE11E76BAFED2FD26" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E76BAFDBCFD0D" bold="true" box="[312,518,683,709]" pageId="44" pageNumber="45">Natural History.</emphasis>
<taxonomicName id="2DA14D11C302C75CE23176BDFD4AFD0D" box="[535,752,684,709]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C302C75CE23176BDFD4AFD0D" box="[535,752,684,709]" italics="true" pageId="44" pageNumber="45">Seeversiella badia</emphasis>
</taxonomicName>
was collected in cloud forest litter at altitude of
<quantity id="2D599B77C302C75CE12E76C5FEDEFD26" box="[264,356,724,750]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="44" pageNumber="45" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,246 @@
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<subSubSection id="A2BB6519C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40" type="key">
<paragraph id="EA1E3692C309C757E12E76C2FC7BFD25" blockId="39.[264,961,723,750]" box="[264,961,723,750]" pageId="39" pageNumber="40">
<heading id="B15681FEC309C757E12E76C2FC7BFD25" bold="true" box="[264,961,723,750]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="D8D5EA80C309C757E12E76C2FC7BFD25" bold="true" box="[264,961,723,750]" pageId="39" pageNumber="40">
<keyStep id="51552E37C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
<key id="9E3061A3C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
<keyLead id="515095A7C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
10.
<taxonomicName id="2DA14D11C309C757E11276C5FD08FD25" ID-CoL="6YBD6" authority="Gusarov" authorityName="Gusarov" box="[308,690,723,750]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C309C757E11276C5FDFBFD26" bold="true" box="[308,577,724,750]" italics="true" pageId="39" pageNumber="40">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC309C757E29976C2FCB8FD25" box="[703,770,723,749]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C309C757E33776C2FC03FD25" box="[785,953,723,749]" captionStart-0="FIGURES 135 ­ 138" captionStart-1="FIGURES 139 ­ 142" captionStart-2="FIGURES 143 ­ 149" captionStartId-0="39.[264,383,1739,1763]" captionStartId-1="40.[264,383,1421,1445]" captionStartId-2="41.[264,383,1483,1507]" captionTargetBox-0="[313,1272,932,1705]" captionTargetBox-1="[306,1279,781,1384]" captionTargetBox-2="[298,1288,744,1449]" captionTargetId-0="figure@39.[304,1283,924,1712]" captionTargetId-1="figure@40.[299,1288,769,1395]" captionTargetId-2="figure@41.[289,1297,729,1456]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="FIGURES 135 ­ 138. Abdominal segment 8 of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (135 ­ 136) and Suchixtepec (137 ­ 138), Mexico). 135 male tergum 8; 136 male sternum 8; 137 female tergum 8; 138 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-2="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156443/files/figure.png" httpUri-1="https://zenodo.org/record/156444/files/figure.png" httpUri-2="https://zenodo.org/record/156445/files/figure.png" pageId="39" pageNumber="40">Figs. 135­149</figureCitation>
)
</keyLead>
</key>
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C309C758E12E770FFD44FE1E" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="EA1E3692C309C757E12E770FFBA9FCA9" blockId="39.[264,1323,798,865]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E12E770FFE07FCF0" bold="true" box="[264,445,798,824]" pageId="39" pageNumber="40">
<typeStatus id="351A8830C309C757E12E770FFEF9FCF0" box="[264,323,798,824]" pageId="39" pageNumber="40">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C309C757E1F6770EFDFAFCF1" box="[464,576,799,825]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C309C757E257770FFCCAFCF0" bold="true" box="[625,880,798,824]" pageId="39" pageNumber="40">
<collectingCountry id="92B67602C309C757E257770FFD4BFCF0" box="[625,753,798,824]" name="Mexico" pageId="39" pageNumber="40">MEXICO</collectingCountry>
: Oaxaca:
</emphasis>
<quantity id="2D599B77C309C757E3A7770EFC6EFCF1" box="[897,980,799,825]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="39" pageNumber="40" unit="km" value="4.6">4.6 km</quantity>
S Suchixtepec,
<quantity id="2D599B77C309C757E4B3770EFB48FCF1" box="[1173,1266,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="39" pageNumber="40" unit="m" value="2150.0">2150 m</quantity>
, leaf litter in riparian alder forest (R.S.Anderson),
<date id="9E1F1052C309C757E33C7756FC1AFCA9" box="[794,928,839,865]" pageId="39" pageNumber="40" value="1992-07-23">23.vii.1992</date>
(
<collectionCode id="8CB0AE57C309C757E3897756FBBDFCA9" box="[943,1031,839,865]" pageId="39" pageNumber="40">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC309C757E12E72DAFB31F8EF" httpUri="https://zenodo.org/record/156443/files/figure.png" pageId="39" pageNumber="40" targetBox="[313,1272,932,1705]" targetPageId="39">
<paragraph id="EA1E3692C309C757E12E72DAFB31F8EF" blockId="39.[264,1323,1739,1831]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E12E72DAFE5BF92B" bold="true" box="[264,481,1739,1763]" pageId="39" pageNumber="40">FIGURES 135­138</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C309C757E33472DAFBDDF92A" authority="Gusarov" authorityName="Gusarov" box="[786,1127,1739,1762]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C309C757E33472DAFBB8F92A" box="[786,1026,1739,1762]" italics="true" pageId="39" pageNumber="40">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C309C757E45272DAFB08F92B" bold="true" box="[1140,1202,1739,1763]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="C3E657FBC309C757E45272DAFB08F92B" box="[1140,1202,1739,1763]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo (135­136) and Suchixtepec (137­138), Mexico). 135 male tergum 8; 136 male sternum 8; 137 female tergum 8; 138 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C306C758E11E750DFD44FE1E" blockId="40.[264,1324,283,710]" pageId="40" pageNumber="41">
<typeStatus id="351A8830C306C758E11E750DFE14FEFE" box="[312,430,284,310]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C306C758E1E6750AFD06FEFD" bold="true" box="[448,700,283,309]" pageId="40" pageNumber="41">
<collectingCountry id="92B67602C306C758E1E6750AFDFAFEFD" box="[448,576,283,309]" name="Mexico" pageId="40" pageNumber="41">MEXICO</collectingCountry>
: Oaxaca:
</emphasis>
2, 5, same data as the
<typeStatus id="351A8830C306C758E43F750DFB39FEFE" box="[1049,1155,284,310]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
;, 2, ditto but
<quantity id="2D599B77C306C758E1107555FE33FE96" box="[310,393,324,350]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="40" pageNumber="41" unit="km" value="5.1">5.1 km</quantity>
S Suchixtepec, oak, pine and alder forest litter,
<date id="9E1F1052C306C758E3EE7555FBEBFE96" box="[968,1105,324,350]" pageId="40" pageNumber="41" value="1992-07-25">25.vii.1992</date>
;, 4, ditto but
<date id="9E1F1052C306C758E12E757DFE2BFE4E" box="[264,401,364,390]" pageId="40" pageNumber="41" value="1992-07-24">24.vii.1992</date>
;
<emphasis id="D8D5EA80C306C758E186757AFDA5FE4D" bold="true" box="[416,543,363,389]" pageId="40" pageNumber="41">Guerrero:</emphasis>
3, 4,
<quantity id="2D599B77C306C758E2E7757DFCB4FE4E" box="[705,782,364,390]" metricMagnitude="4" metricUnit="m" metricValue="7.1" pageId="40" pageNumber="41" unit="km" value="71.0">71 km</quantity>
NE Atoyac de Alvarez,
<quantity id="2D599B77C306C758E41C757DFB23FE4E" box="[1082,1177,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="40" pageNumber="41" unit="m" value="1700.0">1700 m</quantity>
, forest litter (J.S.Ashe),
<date id="9E1F1052C306C758E1B57585FDA6FE66" box="[403,540,404,430]" pageId="40" pageNumber="41" value="1992-07-25">25.vii.1992</date>
; 3,,
<quantity id="2D599B77C306C758E2B17585FD51FE66" box="[663,747,404,430]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="40" pageNumber="41" unit="km" value="5.6">5.6 km</quantity>
SW Filo de Caballo,
<quantity id="2D599B77C306C758E3D07585FBE9FE66" box="[1014,1107,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.31" pageId="40" pageNumber="41" unit="m" value="2310.0">2310 m</quantity>
, alder forest litter (R.S.Anderson),
<date id="9E1F1052C306C758E1E975ADFDEEFE1E" box="[463,596,444,470]" pageId="40" pageNumber="41" value="1992-07-13">13.vii.1992</date>
(all ­
<collectionCode id="8CB0AE57C306C758E2BD75ADFD48FE1E" box="[667,754,444,470]" pageId="40" pageNumber="41">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C306C758E11E75F2FCD6F8A2" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="EA1E3692C306C758E11E75F2FE3FFD0E" blockId="40.[264,1324,283,710]" pageId="40" pageNumber="41">
<emphasis id="D8D5EA80C306C758E11E75F2FE0DFE35" bold="true" box="[312,439,483,509]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C306C758E1EE75F5FD75FE35" box="[456,719,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C306C758E1EE75F5FD75FE35" box="[456,719,484,509]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C306C758E4F575F5FEFDFDED" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C306C758E4F575F5FEFDFDED" italics="true" pageId="40" pageNumber="41">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.2­2.0 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer or a little shorter than pronotum (elytral length to pronotal length ratio 1.2­0.9); wings fully developed or reduced; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C306C758E3887695FBEFFD56" box="[942,1109,644,670]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Figs. 139­148</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C306C758E13776BDFEC2FD0E" box="[273,376,684,710]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 149</figureCitation>
).
</paragraph>
<caption id="BEDE661AC306C758E12E719CFDAEF9C5" httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41" targetBox="[306,1279,781,1384]" targetPageId="40">
<paragraph id="EA1E3692C306C758E12E719CFDAEF9C5" blockId="40.[264,1323,1421,1549]" pageId="40" pageNumber="41">
<emphasis id="D8D5EA80C306C758E12E719CFE58FA6D" bold="true" box="[264,482,1421,1445]" pageId="40" pageNumber="41">FIGURES 139­142</emphasis>
. Aedeagus
<taxonomicName id="2DA14D11C306C758E254719FFC70FA6D" authority="Gusarov" authorityName="Gusarov" box="[626,970,1422,1445]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C306C758E254719FFCDEFA6D" box="[626,868,1422,1445]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C306C758E3FF719CFBA2FA6D" bold="true" box="[985,1048,1421,1445]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="C3E657FBC306C758E3FF719CFBA2FA6D" box="[985,1048,1421,1445]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142).
</paragraph>
</caption>
<paragraph id="EA1E3692C306C758E11E7271FCD6F8A2" blockId="40.[264,1323,1632,1898]" pageId="40" pageNumber="41">
<taxonomicName id="2DA14D11C306C758E11E7271FDFAF9B1" box="[312,576,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C306C758E11E7271FDFAF9B1" box="[312,576,1632,1657]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C306C758E3AD7271FB9DF9B1" box="[907,1063,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C306C758E3AD7271FB9DF9B1" box="[907,1063,1632,1657]" italics="true" pageId="40" pageNumber="41">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 11­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have strong spicules (
<figureCitation id="729A2A17C306C758E36D72C9FC09F93A" box="[843,947,1752,1778]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 146</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C306C758E46F7311FB42F8D2" box="[1097,1272,1792,1818]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41">Figs. 141­142</figureCitation>
), in parameral view narrow and with weak basal constriction (
<figureCitation id="729A2A17C306C758E3917339FBDAF88A" box="[951,1120,1832,1858]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41">Figs. 139­140</figureCitation>
); spermatheca Sshaped with umbilicus facing proximally (
<figureCitation id="729A2A17C306C758E2DE7341FCE5F8A2" box="[760,863,1872,1898]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 149</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C307C759E11E750AFDCDF947" pageId="41" pageNumber="42" type="description">
<paragraph id="EA1E3692C307C759E11E750AFC17FE96" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="41" pageNumber="42">Description.</emphasis>
Length
<quantity id="2D599B77C307C759E21F750DFD7FFEFE" box="[569,709,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="41" pageNumber="42" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C307C759E3A5750DFB96FEFE" box="[899,1068,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="41" pageNumber="42" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C307C759E11E757DFB89FE66" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">
In all character states
<taxonomicName id="2DA14D11C307C759E264757DFD68FE4D" box="[578,722,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C307C759E264757DFD68FE4D" box="[578,722,364,389]" italics="true" pageId="41" pageNumber="42">S. mexicana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C307C759E392757DFBEAFE4D" box="[948,1104,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C307C759E392757DFBEAFE4D" box="[948,1104,364,389]" italics="true" pageId="41" pageNumber="42">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C307C759E11E75ADFD73FDEE" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">In large males posterior angles of male tergum 3 projecting as spines up to twice as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C307C759E11E7625FD26FD86" blockId="41.[264,1323,283,630]" box="[312,668,564,590]" pageId="41" pageNumber="42">
Aedeagus as in
<figureCitation id="729A2A17C307C759E1D77625FD2DFD86" box="[497,663,564,590]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42">Figs. 139­148</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C307C759E11E764DFDC4FDBE" blockId="41.[264,1323,283,630]" box="[312,638,604,630]" pageId="41" pageNumber="42">
Spermatheca as in
<figureCitation id="729A2A17C307C759E233764DFDC0FDBE" box="[533,634,604,630]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42">Fig. 149</figureCitation>
.
</paragraph>
<caption id="BEDE661AC307C759E12E71DAFBB5F9A5" httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42" targetBox="[298,1288,744,1449]" targetPageId="41">
<paragraph id="EA1E3692C307C759E12E71DAFBB5F9A5" blockId="41.[264,1325,1483,1679]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E12E71DAFE5AFA2B" bold="true" box="[264,480,1483,1507]" pageId="41" pageNumber="42">FIGURES 143­149</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C307C759E2A471DDFC6FFA2A" authority="Gusarov" authorityName="Gusarov" box="[642,981,1483,1507]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C307C759E2A471DDFCC8FA2B" box="[642,882,1484,1507]" italics="true" pageId="41" pageNumber="42">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C307C759E3C471DAFB9BFA2B" bold="true" box="[994,1057,1483,1507]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="C3E657FBC307C759E3C471DAFB9BFA2B" box="[994,1057,1483,1507]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo (143, 145­148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145­146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145­146), 0.1 mm (144, 147­149).
</paragraph>
</caption>
<paragraph id="EA1E3692C307C759E12E7269FDCDF947" blockId="41.[264,1325,1483,1679]" box="[264,631,1656,1679]" pageId="41" pageNumber="42">DS distal sclerite of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C307C759E11E72FAFE17F89D" pageId="41" pageNumber="42" type="distribution">
<paragraph id="EA1E3692C307C759E11E72FAFE17F89D" blockId="41.[264,1323,1771,1877]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E11E72FAFE62F8CD" bold="true" box="[312,472,1771,1797]" pageId="41" pageNumber="42">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C307C759E2A372FAFD65F8CD" box="[645,735,1771,1797]" name="Mexico" pageId="41" pageNumber="42">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C307C759E2C972FAFCEDF8CD" box="[751,855,1771,1797]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="41" pageNumber="42">Fig. 387</figureCitation>
).
<emphasis id="D8D5EA80C307C759E11E7302FDB3F8E5" bold="true" box="[312,521,1811,1837]" pageId="41" pageNumber="42">Natural History.</emphasis>
<taxonomicName id="2DA14D11C307C759E2387302FC9DF8E4" box="[542,807,1811,1836]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C307C759E2387302FC9DF8E4" box="[542,807,1811,1836]" italics="true" pageId="41" pageNumber="42">Seeversiella mexicana</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C307C759E12E732AFE12F89D" box="[264,424,1851,1877]" metricMagnitude="3" metricUnit="m" metricValue="2.005" metricValueMax="2.31" metricValueMin="1.7" pageId="41" pageNumber="42" unit="m" value="2005.0" valueMax="2310.0" valueMin="1700.0">1700­2310 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,168 @@
<document id="ACA95A174DE776F477FFA89B0B8121D9" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C30BC757E12E77D2FD1DFDBE" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella grandis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="40" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="38" updateTime="1698203169024" updateUser="ExternalLinkService">
<mods:mods id="E1C05000137E0608DB556E06D6ED651B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="198894F31CE0F8F0D2AD6C5D7A9A9FF9">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="73896104A7AC13E46A7DF51F20832D9E" type="personal">
<mods:role id="398F55EEF31BDB43F1721AD8FCB6500E">
<mods:roleTerm id="1FA026B7E4AF924D86A9D8CDAD61BE23">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7E93F4324CE5BDD5FF468D37F5DB32C">Gusarov, Vladimir I.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A415F67582E7DC856CAF1D03DE7AA52F">text</mods:typeOfResource>
<mods:relatedItem id="B74DD9B24F79FB495107EFA3AD703C08" type="host">
<mods:titleInfo id="5FC69895BF58411CD433AB492588A9B8">
<mods:title id="CFF15FE5AEE47978F1581955153A8A56">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="08B58322871260C53811795634500079">
<mods:date id="B7302C34997E5F590ED39FDFCB5AD902">2003</mods:date>
<mods:detail id="51FE691B03E38A49B6AE7FEC1B7C1FA3" type="volume">
<mods:number id="930357A50EAB655EA51A8DE2CAD7B391">142</mods:number>
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<mods:start id="86596337BC0EB7D8BC1629EAC76903A7">1</mods:start>
<mods:end id="1486910A4C387AB2A7493D336C3DB3CD">102</mods:end>
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<mods:identifier id="4998B9D9F9E5B2574AA19C8CC13D9E8A" type="CLB-Dataset">51371</mods:identifier>
<mods:identifier id="2FB06E28A5772A7AE70826F917510392" type="DOI">10.5281/zenodo.156420</mods:identifier>
<mods:identifier id="B7DDE3A9AE908BEF8A1546940E199305" type="GBIF-Dataset">76c096e3-d27c-4abc-a504-a62ae8c7ba92</mods:identifier>
<mods:identifier id="BAB7AEA7F8B744A9DC404D38F506BCDC" type="ISSN">1175­5326</mods:identifier>
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<subSubSection id="A2BB6519C30BC755E12E77D2FC22FC15" box="[264,920,963,990]" pageId="37" pageNumber="38" type="key">
<paragraph id="EA1E3692C30BC755E12E77D2FC22FC15" blockId="37.[264,920,963,990]" box="[264,920,963,990]" pageId="37" pageNumber="38">
<heading id="B15681FEC30BC755E12E77D2FC22FC15" bold="true" box="[264,920,963,990]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="D8D5EA80C30BC755E12E77D2FC22FC15" bold="true" box="[264,920,963,990]" pageId="37" pageNumber="38">
<key id="9E3061A3C30BC755E12E77D2FC22FC15" box="[264,920,963,990]" pageId="37" pageNumber="38">
<keyStep id="51552E37C30BC755E12E77D2FC22FC15" box="[264,920,963,990]" pageId="37" pageNumber="38">
<keyLead id="515095A7C30BC755E12E77D2FC22FC15" box="[264,920,963,990]" pageId="37" pageNumber="38">
9.
<taxonomicName id="2DA14D11C30BC755E10077D5FD30FC15" ID-CoL="6YBD7" authority="Gusarov" authorityName="Gusarov" box="[294,650,963,990]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="D8D5EA80C30BC755E10077D5FDADFC16" bold="true" box="[294,535,964,990]" italics="true" pageId="37" pageNumber="38">Seeversiella grandis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30BC755E2B177D2FD63FC15" box="[663,729,963,989]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30BC755E2CF77D2FC2BFC15" box="[745,913,963,989]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="37" pageNumber="38">Figs. 126­134</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E12E701FFBF7FB99" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="EA1E3692C30BC755E12E701FFBF7FB99" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E12E701FFE07FBE0" bold="true" box="[264,445,1038,1064]" pageId="37" pageNumber="38">
<typeStatus id="351A8830C30BC755E12E701FFEF9FBE0" box="[264,323,1038,1064]" pageId="37" pageNumber="38">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30BC755E1F4701EFDF8FBE1" box="[466,578,1039,1065]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30BC755E255701FFBA7FBE0" bold="true" box="[627,1053,1038,1064]" pageId="37" pageNumber="38">
<collectingCountry id="92B67602C30BC755E255701FFC82FBE0" box="[627,824,1038,1064]" name="Guatemala" pageId="37" pageNumber="38">GUATEMALA</collectingCountry>
: Quetzaltenango:
</emphasis>
<quantity id="2D599B77C30BC755E409701EFBD7FBE1" box="[1071,1133,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="km" value="8.0">8 km</quantity>
SE Zunil,
<quantity id="2D599B77C30BC755E4D6701EFE98FB99" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="37" pageNumber="38" unit="m" value="2620.0">2620 m</quantity>
, flight intercept trap (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C30BC755E3137026FC60FB99" box="[821,986,1079,1105]" pageId="37" pageNumber="38" value="1993-06-19" valueMax="1993-06-21" valueMin="1993-06-19">19­21.vi.1993</date>
(
<collectionCode id="8CB0AE57C30BC755E3CF7026FBFBFB99" box="[1001,1089,1079,1105]" pageId="37" pageNumber="38">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E11E704FFBEAFAD1" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="EA1E3692C30BC755E11E704FFBEAFAD1" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E704FFE0DFBB0" bold="true" box="[312,439,1118,1144]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30BC755E1EA704EFD04FBB0" box="[460,702,1119,1144]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="D8D5EA80C30BC755E1EA704EFD04FBB0" box="[460,702,1119,1144]" italics="true" pageId="37" pageNumber="38">Seeversiella grandis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30BC755E4F5704EFEFDFB68" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30BC755E4F5704EFEFDFB68" italics="true" pageId="37" pageNumber="38">Seeversiella</emphasis>
</taxonomicName>
by having large body (
<quantity id="2D599B77C30BC755E2457096FD7FFB69" box="[611,709,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="37" pageNumber="38" unit="mm" value="3.7">3.7 mm</quantity>
), brownish black body with brown elytra and legs; large eyes (temple length to eye length ratio 0.9); matte pronotum with strong isodiametric microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C30BC755E3BD70EEFBF9FAD1" box="[923,1091,1279,1305]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="37" pageNumber="38">Figs. 129­134</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC757E11E7137FDB1FE36" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="EA1E3692C30BC755E11E7137FB0CFA89" blockId="37.[264,1324,1038,1904]" box="[312,1206,1318,1345]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E7137FE6AFA88" bold="true" box="[312,464,1318,1344]" pageId="37" pageNumber="38">Description.</emphasis>
Length
<quantity id="2D599B77C30BC755E21F7136FD2CFA89" box="[569,662,1319,1345]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="37" pageNumber="38" unit="mm" value="3.7">3.7 mm</quantity>
. Body brownish black, elytra and legs brown.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E715EFBF7FA28" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">Head surface matte, with fine isodiametric microsculpture, fine punctation, distance between punctures equals 1­2 times their diameter, anteromedial portion of the disc without punctation. Temple length to eye length ratio 0.9. Antennal article 2 as long as 3, article 4 elongate (ratio 1.2), 5 quadrate, 6­10 transverse (ratio 1.2­1.3).</paragraph>
<paragraph id="EA1E3692C30BC755E11E71FFFC99F930" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C30BC755E40D71FFFB23F9C0" box="[1067,1177,1518,1544]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="37" pageNumber="38" unit="mm" value="0.74">0.74 mm</quantity>
, length
<quantity id="2D599B77C30BC755E4DF71FFFE83F9F8" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="37" pageNumber="38" unit="mm" value="0.66">0.66 mm</quantity>
, width to length ratio 1.1, surface matte, with fine isodiametric microsculpture and fine punctation, distance between punctures equals ½­1 times their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to ½­ 1 time their diameter. Wings fully developed.
</paragraph>
<paragraph id="EA1E3692C30BC756E11E7317FDCAFE4E" blockId="37.[264,1324,1038,1904]" lastBlockId="38.[264,1325,284,390]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Abdominal terga glossy, terga 3­6 with fine microsculpture consisting of transverse waves and strongly transverse meshes, with fine punctation, distance between punctures equals 3­6 times their diameter. Tergum 7 with microsculpture consisting of transverse meshes and with fine and sparse punctation, distance between punctures equal to 4­7 times their diameter. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C308C756E1DA757DFDD9FE4E" box="[508,611,364,390]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="38" pageNumber="39">Fig. 126</figureCitation>
).
</paragraph>
<caption id="BEDE661AC308C756E12E7264FB85F880" httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="38" pageNumber="39" targetBox="[283,1299,500,1615]" targetPageId="38">
<paragraph id="EA1E3692C308C756E12E7264FB85F880" blockId="38.[264,1323,1653,1864]" pageId="38" pageNumber="39">
<emphasis id="D8D5EA80C308C756E12E7264FE5AF945" bold="true" box="[264,480,1653,1677]" pageId="38" pageNumber="39">FIGURES 126­134</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C308C756E3857267FB58F944" authority="Gusarov" authorityName="Gusarov" box="[931,1250,1653,1677]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="D8D5EA80C308C756E3857267FBC5F945" box="[931,1151,1654,1677]" italics="true" pageId="38" pageNumber="39">Seeversiella grandis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C308C756E4CB7264FA90F945" bold="true" box="[1261,1322,1653,1677]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="C3E657FBC308C756E4CB7264FA90F945" box="[1261,1322,1653,1677]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126­127), 0.3 mm (128, 130, 132­133), 0.1 mm (129, 131, 134).
</paragraph>
</caption>
<paragraph id="EA1E3692C309C757E11E750DFC61FE1E" blockId="39.[264,1323,284,630]" pageId="39" pageNumber="40">
In the only known male specimen pronotum with wide medial impression, posterior margin of male tergum 3 with two short and obtuse projections (as in
<figureCitation id="729A2A17C309C757E4777555FB13FE96" box="[1105,1193,324,350]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
), tergum 7 with medial tubercle in front of posterior margin. Posterior margin of male tergum 8 without emargination (
<figureCitation id="729A2A17C309C757E1C77585FDF2FE66" box="[481,584,404,430]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Fig. 126</figureCitation>
). Aedeagus as in
<figureCitation id="729A2A17C309C757E3047585FC73FE66" box="[802,969,404,430]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Figs. 129­134</figureCitation>
. Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C309C757E34175ADFC74FE1E" box="[871,974,444,470]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 male tergum 8; 127 male sternum 8; 128 median lobe, parameral view; 129 apex of median lobe, parameral view; 130 median lobe, lateral view; 131 apex of median lobe, lateral view; 132 details of internal sac retracted into median lobe, abparameral view; 133 details of internal sac retracted into median lobe, lateral view; 134 apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Fig. 134</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C309C757E11E75F5FDB1FE36" blockId="39.[264,1323,284,630]" box="[312,523,484,510]" pageId="39" pageNumber="40">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C309C757E11E761AFD1DFDBE" pageId="39" pageNumber="40" type="distribution">
<paragraph id="EA1E3692C309C757E11E761AFC33FDEE" blockId="39.[264,1323,284,630]" box="[312,905,523,550]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E11E761AFE62FDED" bold="true" box="[312,472,523,549]" pageId="39" pageNumber="40">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C309C757E2A3761DFCBEFDEE" box="[645,772,524,550]" name="Guatemala" pageId="39" pageNumber="40">Guatemala</collectingCountry>
(
<figureCitation id="729A2A17C309C757E332761DFCC6FDEE" box="[788,892,524,550]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="39" pageNumber="40">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C309C757E11E7622FD1DFDBE" blockId="39.[264,1323,284,630]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E11E7622FDBFFD85" bold="true" box="[312,517,563,589]" pageId="39" pageNumber="40">Natural History.</emphasis>
The only known specimen of
<taxonomicName id="2DA14D11C309C757E3527625FC56FD85" box="[884,1004,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="D8D5EA80C309C757E3527625FC56FD85" box="[884,1004,564,589]" italics="true" pageId="39" pageNumber="40">S. grandis</emphasis>
</taxonomicName>
was collected with a flight intercept trap at altitude of
<quantity id="2D599B77C309C757E261764DFD19FDBE" box="[583,675,604,630]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="39" pageNumber="40" unit="m" value="2620.0">2620 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="EB82048C6B182BBF0062EB58A2E0ADBD">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:date id="DA96ABEF505564803FA020FDD25F972A">2003</mods:date>
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<heading id="B15681FEC30DC753E12E7252FC13F995" bold="true" box="[264,937,1603,1629]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="D8D5EA80C30DC753E12E7252FC13F995" bold="true" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<key id="9E3061A3C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<keyStep id="51552E37C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
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8.
<taxonomicName id="2DA14D11C30DC753E1007252FD27F995" ID-CoL="6YBCY" authority="Gusarov" authorityName="Gusarov" box="[294,669,1603,1629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C30DC753E1007252FD91F995" bold="true" box="[294,555,1603,1629]" italics="true" pageId="35" pageNumber="36">Seeversiella nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30DC753E28C7252FD56F995" box="[682,748,1603,1629]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30DC753E2DA7252FC18F995" box="[764,930,1603,1629]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="35" pageNumber="36">Figs. 117­125</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC753E12E729CFD4BF918" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="EA1E3692C30DC753E12E729CFD4BF918" blockId="35.[264,1324,1677,1904]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E12E729CFE04F96F" bold="true" box="[264,446,1677,1703]" pageId="35" pageNumber="36">
<typeStatus id="351A8830C30DC753E12E729CFEFFF96F" box="[264,325,1677,1703]" pageId="35" pageNumber="36">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30DC753E1EB729FFD87F960" box="[461,573,1678,1704]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30DC753E24D729CFCE4F96F" bold="true" box="[619,862,1677,1703]" pageId="35" pageNumber="36">
<collectingCountry id="92B67602C30DC753E24D729CFD51F96F" box="[619,747,1677,1703]" name="Mexico" pageId="35" pageNumber="36">MEXICO</collectingCountry>
: Puebla:
</emphasis>
<quantity id="2D599B77C30DC753E348729FFC05F960" box="[878,959,1678,1704]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="36" unit="km" value="1.6">1.6 km</quantity>
E Nicolas Bravo,
<quantity id="2D599B77C30DC753E4B1729FFB48F960" box="[1175,1266,1678,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="35" pageNumber="36" unit="m" value="2410.0">2410 m</quantity>
, forest litter (J.S.Ashe),
<date id="9E1F1052C30DC753E1D172A7FDC7F918" box="[503,637,1718,1744]" pageId="35" pageNumber="36" value="1992-07-17">17.vii.1992</date>
(
<collectionCode id="8CB0AE57C30DC753E2AA72A7FD5EF918" box="[652,740,1718,1744]" pageId="35" pageNumber="36">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC754E11E72CCFC6DFE96" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="EA1E3692C30DC753E11E72CCFA91F8B8" blockId="35.[264,1324,1677,1904]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E72CCFE0DF93F" bold="true" box="[312,439,1757,1783]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30DC753E1EE72CFFD76F93F" box="[456,716,1758,1783]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30DC753E1EE72CFFD76F93F" box="[456,716,1758,1783]" italics="true" pageId="35" pageNumber="36">Seeversiella nigriceps</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30DC753E4F572CFFEFDF8D7" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30DC753E4F572CFFEFDF8D7" italics="true" pageId="35" pageNumber="36">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples twice as long as eyes; glossy pronotum with weak and partially missing microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C30DC753E4517347FAA5F8B8" box="[1143,1311,1878,1904]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="35" pageNumber="36">Figs. 119­125</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30AC754E11E750DFC6DFE96" blockId="36.[264,1323,284,350]" pageId="36" pageNumber="37">
<taxonomicName id="2DA14D11C30AC754E11E750DFD80FEFD" box="[312,570,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30AC754E11E750DFD80FEFD" box="[312,570,284,309]" italics="true" pageId="36" pageNumber="37">Seeversiella nigriceps</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C30AC754E3A0750DFBFFFEFD" box="[902,1093,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C30AC754E3A0750DFBFFFEFD" box="[902,1093,284,309]" italics="true" pageId="36" pageNumber="37">S. sonomotoides</emphasis>
</taxonomicName>
in having narrower distal sclerites of internal sac of aedeagus (
<figureCitation id="729A2A17C30AC754E2D97555FCCAFE96" box="[767,880,324,350]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="36" pageNumber="37">Figs. 124</figureCitation>
; 63, 65).
</paragraph>
</subSubSection>
<caption id="BEDE661AC30AC754E12E727DFDCDF8F8" httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="36" pageNumber="37" targetBox="[297,1299,423,1607]" targetPageId="36">
<paragraph id="EA1E3692C30AC754E12E727DFDCDF8F8" blockId="36.[264,1322,1644,1840]" pageId="36" pageNumber="37">
<emphasis id="D8D5EA80C30AC754E12E727DFE67F94C" bold="true" box="[264,477,1644,1668]" pageId="36" pageNumber="37">FIGURES 117­125</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C30AC754E3B1727DFB59F94B" authority="Gusarov" authorityName="Gusarov" box="[919,1251,1644,1667]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C30AC754E3B1727DFB3BF94B" box="[919,1153,1644,1667]" italics="true" pageId="36" pageNumber="37">Seeversiella nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C30AC754E4C9727DFA90F94C" bold="true" box="[1263,1322,1644,1668]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="C3E657FBC30AC754E4C9727DFA90F94C" box="[1263,1322,1644,1668]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123­124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117­118), 0.2 mm (119, 121, 123­124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac.
</paragraph>
</caption>
<subSubSection id="A2BB6519C30BC755E11E750DFDB0FD26" pageId="37" pageNumber="38" type="description">
<paragraph id="EA1E3692C30BC755E11E750DFB6EFE66" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<taxonomicName id="2DA14D11C30BC755E11E750DFD84FEFD" box="[312,574,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E11E750DFD84FEFD" box="[312,574,284,309]" italics="true" pageId="37" pageNumber="38">Seeversiella nigriceps</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30BC755E3AC750DFB9CFEFD" box="[906,1062,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30BC755E3AC750DFB9CFEFD" box="[906,1062,284,309]" italics="true" pageId="37" pageNumber="38">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (5­7, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac sclerotized and dentiform (
<figureCitation id="729A2A17C30BC755E4DA757DFE82FE66" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Fig. 124</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C30BC755E4397585FB7DFE66" box="[1055,1223,404,430]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Figs. 121­122</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30BC755E11E75AAFE2BFE36" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E75AAFE6AFE1D" bold="true" box="[312,464,443,469]" pageId="37" pageNumber="38">Description.</emphasis>
Length
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, pronotal width
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. Body dark brown, elytra and legs brown.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E761DFBF5FD86" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
In all character states
<taxonomicName id="2DA14D11C30BC755E21F761DFD79FDED" box="[569,707,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E21F761DFD79FDED" box="[569,707,524,549]" italics="true" pageId="37" pageNumber="38">S. nigriceps</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C30BC755E3BC761DFBE3FDED" box="[922,1113,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C30BC755E3BC761DFBE3FDED" box="[922,1113,524,549]" italics="true" pageId="37" pageNumber="38">S. sonomotoides</emphasis>
</taxonomicName>
, but differs in the shape of the distal sclerites of internal sac as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E764DFBCDFD56" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
In the only known male specimen the posterior angles of male tergum 3 not projecting as spines, tergum
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front of posterior margin with small medial tubercle.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E76BDFD20FD0E" blockId="37.[264,1324,284,870]" box="[312,666,684,710]" pageId="37" pageNumber="38">
Aedeagus as in
<figureCitation id="729A2A17C30BC755E1D676BDFD2CFD0E" box="[496,662,684,710]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Figs. 119­125</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E76C5FDB0FD26" blockId="37.[264,1324,284,870]" box="[312,522,724,750]" pageId="37" pageNumber="38">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E11E76EAFD90FCAE" pageId="37" pageNumber="38" type="distribution">
<paragraph id="EA1E3692C30BC755E11E76EAFCD8FCDE" blockId="37.[264,1324,284,870]" box="[312,866,763,790]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E76EAFE62FCDD" bold="true" box="[312,472,763,789]" pageId="37" pageNumber="38">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30BC755E2A376EDFD65FCDE" box="[645,735,764,790]" name="Mexico" pageId="37" pageNumber="38">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C30BC755E2C876EDFCEFFCDE" box="[750,853,764,790]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="37" pageNumber="38">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30BC755E11E7732FD90FCAE" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E7732FDBEFCF5" bold="true" box="[312,516,803,829]" pageId="37" pageNumber="38">Natural History.</emphasis>
The only known specimen of
<taxonomicName id="2DA14D11C30BC755E34A7735FC4CFCF5" box="[876,1014,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E34A7735FC4CFCF5" box="[876,1014,804,829]" italics="true" pageId="37" pageNumber="38">S. nigriceps</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C30BC755E1EC775DFD9FFCAE" box="[458,549,844,870]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="37" pageNumber="38" unit="m" value="2410.0">2410 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,201 @@
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<subSubSection id="A2BB6519C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34" type="key">
<paragraph id="EA1E3692C30FC751E12E7162FC48FA45" blockId="33.[264,1010,1395,1422]" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<heading id="B15681FEC30FC751E12E7162FC48FA45" bold="true" box="[264,1010,1395,1422]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="D8D5EA80C30FC751E12E7162FC48FA45" bold="true" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<keyStep id="51552E37C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<key id="9E3061A3C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<keyLead id="515095A7C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
7.
<taxonomicName id="2DA14D11C30FC751E1007165FD5FFA45" ID-CoL="6YBBJ" authority="Gusarov" authorityName="Gusarov" box="[294,741,1395,1422]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C30FC751E1007165FDC8FA46" bold="true" box="[294,626,1396,1422]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30FC751E2D47162FC8EFA45" box="[754,820,1395,1421]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30FC751E3627162FC50FA45" box="[836,1002,1395,1421]" captionStart-0="FIGURES 105 ­ 108" captionStart-1="FIGURES 109 ­ 116" captionStartId-0="34.[264,383,1348,1372]" captionStartId-1="35.[264,383,1334,1358]" captionTargetBox-0="[296,1289,499,1312]" captionTargetBox-1="[306,1282,441,1310]" captionTargetId-0="figure@34.[285,1302,489,1322]" captionTargetId-1="figure@35.[293,1295,433,1320]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURES 105 ­ 108. Abdominal segment 8 of Seeversiella tuberculicauda Gusarov, sp. n. (105 ­ 106, holotype; 107 ­ 108, paratype from Jalapa, Mexico). 105 male tergum 8; 106 male sternum 8; 107 female tergum 8; 108 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri-0="https://zenodo.org/record/156439/files/figure.png" httpUri-1="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Figs. 105­116</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E12E71AFFB19F9E0" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="EA1E3692C30FC751E12E71AFFD45F9C8" blockId="33.[264,1323,1470,1896]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E12E71AFFE07FA10" bold="true" box="[264,445,1470,1496]" pageId="33" pageNumber="34">
<typeStatus id="351A8830C30FC751E12E71AFFEF9FA10" box="[264,323,1470,1496]" pageId="33" pageNumber="34">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30FC751E1F471AFFDF8FA10" box="[466,578,1470,1496]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30FC751E25571AFFC3DFA10" bold="true" box="[627,903,1470,1496]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E25571AFFD49FA10" box="[627,755,1470,1496]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
: Veracruz:
</emphasis>
<quantity id="2D599B77C30FC751E3BF71AFFC57FA10" box="[921,1005,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="34" unit="km" value="2.3">2.3 km</quantity>
S Jalapa,
<quantity id="2D599B77C30FC751E44E71AFFB7CFA10" box="[1128,1222,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="33" pageNumber="34" unit="m" value="1320.0">1320 m</quantity>
, streamside litter (J.S.Ashe),
<date id="9E1F1052C30FC751E22071F7FD36F9C8" box="[518,652,1510,1536]" pageId="33" pageNumber="34" value="1992-07-13">13.vii.1992</date>
(
<collectionCode id="8CB0AE57C30FC751E2BD71F7FD49F9C8" box="[667,755,1510,1536]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E721FFB19F9E0" blockId="33.[264,1323,1470,1896]" box="[312,1187,1550,1576]" pageId="33" pageNumber="34">
<typeStatus id="351A8830C30FC751E11E721FFE14F9E0" box="[312,430,1550,1576]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C30FC751E1E6721FFD6AF9E0" bold="true" box="[448,720,1550,1576]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E1E6721FFDFAF9E0" box="[448,576,1550,1576]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
: Veracruz:
</emphasis>
, same data as the
<typeStatus id="351A8830C30FC751E3EE721FFB94F9E0" box="[968,1070,1550,1576]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C30FC751E418721FFB2CF9E0" box="[1086,1174,1550,1576]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC752E11E7227FD0AFE66" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="EA1E3692C30FC751E11E7227FD31F938" blockId="33.[264,1323,1470,1896]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E7227FE02F998" bold="true" box="[312,440,1590,1616]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30FC751E1E87226FCA2F998" box="[462,792,1591,1616]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30FC751E1E87226FCA2F998" box="[462,792,1591,1616]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30FC751E12E724EFE2CF9B0" box="[264,406,1631,1656]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30FC751E12E724EFE2CF9B0" box="[264,406,1631,1656]" italics="true" pageId="33" pageNumber="34">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.2 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C30FC751E49472BFFE8EF938" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Figs. 110­ 115</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C30FC751E23E72C7FDC4F938" box="[536,638,1750,1776]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Fig. 116</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC752E11E72EEFD0AFE66" blockId="33.[264,1323,1470,1896]" lastBlockId="34.[264,1323,284,430]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<taxonomicName id="2DA14D11C30FC751E11E72EEFDC7F8D0" box="[312,637,1791,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30FC751E11E72EEFDC7F8D0" box="[312,637,1791,1816]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30FC751E39E72EEFBEBF8D0" box="[952,1105,1791,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E39E72EEFBEBF8D0" box="[952,1105,1791,1816]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­6, 8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have weak spicules (
<figureCitation id="729A2A17C30CC752E3BD750DFBB8FEFE" box="[923,1026,284,310]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 114</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C30CC752E47F7555FB45FE96" box="[1113,1279,324,350]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 112­113</figureCitation>
), in parameral view wide (
<figureCitation id="729A2A17C30CC752E23D757DFD7FFE4E" box="[539,709,364,390]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 110­111</figureCitation>
); spermatheca L­shaped with small umbilicus and straight proximal portion (
<figureCitation id="729A2A17C30CC752E21B7585FD19FE66" box="[573,675,404,430]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 116</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC30CC752E12E7155FC4EFA69" httpUri="https://zenodo.org/record/156439/files/figure.png" pageId="34" pageNumber="35" targetBox="[296,1289,499,1312]" targetPageId="34">
<paragraph id="EA1E3692C30CC752E12E7155FC4EFA69" blockId="34.[264,1323,1348,1441]" pageId="34" pageNumber="35">
<emphasis id="D8D5EA80C30CC752E12E7155FE5AFA94" bold="true" box="[264,480,1348,1372]" pageId="34" pageNumber="35">FIGURES 105­108</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C30CC752E3297154FB27FA94" authority="Gusarov" authorityName="Gusarov" box="[783,1181,1349,1372]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C30CC752E3297154FB83FA94" box="[783,1081,1349,1372]" italics="true" pageId="34" pageNumber="35">Seeversiella tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30CC752E48C7155FB52FA94" box="[1194,1256,1348,1372]" pageId="34" pageNumber="35" rank="species">
<emphasis id="D8D5EA80C30CC752E48C7155FB52FA94" bold="true" box="[1194,1256,1348,1372]" pageId="34" pageNumber="35">sp. n.</emphasis>
</taxonomicNameLabel>
(105­ 106, holotype; 107­108, paratype from Jalapa, Mexico). 105 male tergum 8; 106 male sternum 8; 107 female tergum 8; 108 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C30CC752E11E71E7FDC7F899" pageId="34" pageNumber="35" type="description">
<paragraph id="EA1E3692C30CC752E11E71E7FEE0F9F1" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
<emphasis id="D8D5EA80C30CC752E11E71E7FE6AF9D8" bold="true" box="[312,464,1526,1552]" pageId="34" pageNumber="35">Description.</emphasis>
Length
<quantity id="2D599B77C30CC752E26D71E6FD14F9D9" box="[587,686,1527,1553]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="34" pageNumber="35" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C30CC752E35871E6FC4AF9D9" box="[894,1008,1527,1553]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="34" pageNumber="35" unit="mm" value="0.49">0.49 mm</quantity>
. Body dark brown, legs brown.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7256FB75F941" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
In all external character states
<taxonomicName id="2DA14D11C30CC752E28E7256FCC2F9A8" box="[680,888,1607,1632]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30CC752E28E7256FCC2F9A8" box="[680,888,1607,1632]" italics="true" pageId="34" pageNumber="35">S. tuberculicauda</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C30CC752E47F7256FB4EF9A8" box="[1113,1268,1607,1632]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30CC752E47F7256FB4EF9A8" box="[1113,1268,1607,1632]" italics="true" pageId="34" pageNumber="35">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7286FED4F8C9" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
In the only known male specimen the posterior angles of male tergum 3 projecting as spines twice as long as tergum 3 (measured medially), tergum
<quantity id="2D599B77C30CC752E3CE72AEFBAFF911" box="[1000,1045,1727,1753]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="34" pageNumber="35" unit="in" value="7.0">7 in</quantity>
basal third with medial tubercle.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E731EFD23F8E1" blockId="34.[264,1324,1526,1873]" box="[312,665,1807,1833]" pageId="34" pageNumber="35">
Aedeagus as in
<figureCitation id="729A2A17C30CC752E1D6731EFD2FF8E1" box="[496,661,1807,1833]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 110­115</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7326FDC7F899" blockId="34.[264,1324,1526,1873]" box="[312,637,1847,1873]" pageId="34" pageNumber="35">
Spermatheca as in
<figureCitation id="729A2A17C30CC752E2327326FDC2F899" box="[532,632,1847,1873]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 116</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC753E11E750AFE5DFE4E" pageId="35" pageNumber="36" type="distribution">
<paragraph id="EA1E3692C30DC753E11E750AFCD9FEFE" blockId="35.[264,1323,283,390]" box="[312,867,283,310]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="35" pageNumber="36">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30DC753E2A3750DFD65FEFE" box="[645,735,284,310]" name="Mexico" pageId="35" pageNumber="36">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C30DC753E2C9750DFCEDFEFE" box="[751,855,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="35" pageNumber="36">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30DC753E11E7552FE5DFE4E" blockId="35.[264,1323,283,390]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E7552FDB0FE95" bold="true" box="[312,522,323,349]" pageId="35" pageNumber="36">Natural History.</emphasis>
<taxonomicName id="2DA14D11C30DC753E2077555FCD1FE95" box="[545,875,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30DC753E2077555FCD1FE95" box="[545,875,324,349]" italics="true" pageId="35" pageNumber="36">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
was collected in streamside litter at altitude of
<quantity id="2D599B77C30DC753E1A1757DFE5EFE4E" box="[391,484,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="35" pageNumber="36" unit="m" value="1320.0">1320 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,301 @@
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<mods:title id="99A1D3AFCC9A1785EEC9E3F58ABCEC30">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<treatment id="62088784C311C733E12E7780FE17FE4E" ID-DOI="http://doi.org/10.5281/zenodo.6274674" ID-Zenodo-Dep="6274674" LSID="urn:lsid:plazi:treatment:62088784C311C733E12E7780FE17FE4E" httpUri="http://treatment.plazi.org/id/62088784C311C733E12E7780FE17FE4E" lastPageId="67" lastPageNumber="68" pageId="63" pageNumber="64">
<subSubSection id="A2BB6519C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64" type="key">
<paragraph id="EA1E3692C311C74FE12E7780FC6FFC63" blockId="63.[264,981,913,939]" box="[264,981,913,939]" pageId="63" pageNumber="64">
<heading id="B15681FEC311C74FE12E7780FC6FFC63" bold="true" box="[264,981,913,939]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
<key id="9E3061A3C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<keyStep id="51552E37C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<keyLead id="515095A7C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE12E7780FC6FFC63" bold="true" box="[264,981,913,939]" pageId="63" pageNumber="64">
18.
<taxonomicName id="2DA14D11C311C74FE1127780FD7DFC63" ID-CoL="6YBCS" authority="Gusarov" authorityName="Gusarov" box="[308,711,913,939]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C311C74FE1127780FDEEFC63" bold="true" box="[308,596,913,939]" italics="true" pageId="63" pageNumber="64">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC311C74FE2F57780FCAFFC63" box="[723,789,913,939]" pageId="63" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C311C74FE3037780FC77FC63" box="[805,973,913,939]" captionStart-0="FIGURES 247 ­ 250" captionStart-1="FIGURES 251 ­ 255" captionStart-2="FIGURES 256 ­ 264" captionStartId-0="63.[264,383,1778,1802]" captionStartId-1="64.[264,383,1694,1718]" captionStartId-2="66.[264,383,1579,1603]" captionTargetBox-0="[283,1297,1044,1745]" captionTargetBox-1="[284,1301,785,1656]" captionTargetBox-2="[278,1308,342,1490]" captionTargetId-0="figure@63.[271,1315,1033,1752]" captionTargetId-1="figure@64.[270,1316,769,1667]" captionTargetId-2="figure@66.[270,1318,331,1497]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="66" captionText-0="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." captionText-2="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri-0="https://zenodo.org/record/156463/files/figure.png" httpUri-1="https://zenodo.org/record/156464/files/figure.png" httpUri-2="https://zenodo.org/record/156465/files/figure.png" pageId="63" pageNumber="64">Figs. 247­264</figureCitation>
)
</emphasis>
</keyLead>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C311C74FE12E72E3FD11F887" blockId="63.[264,1323,1778,1871]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE12E72E3FE65F8C2" bold="true" box="[264,479,1778,1802]" pageId="63" pageNumber="64">FIGURES 247­250</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C311C74FE32272E2FBDDF8C2" authority="Gusarov" authorityName="Gusarov" box="[772,1127,1779,1802]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C311C74FE32272E2FBBEF8C2" box="[772,1028,1779,1802]" italics="true" pageId="63" pageNumber="64">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC311C74FE45772E3FB17F8C2" box="[1137,1197,1778,1802]" pageId="63" pageNumber="64" rank="species">
<emphasis id="D8D5EA80C311C74FE45772E3FB17F8C2" bold="true" box="[1137,1197,1778,1802]" pageId="63" pageNumber="64">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C36EC730E12E750AFC57FE1E" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="EA1E3692C36EC730E12E750AFB35FE96" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E12E750AFE07FEFD" bold="true" box="[264,445,283,309]" pageId="64" pageNumber="65">
<typeStatus id="351A8830C36EC730E12E750AFEF9FEFD" box="[264,323,283,309]" pageId="64" pageNumber="65">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36EC730E1F7750DFDFBFEFE" box="[465,577,284,310]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36EC730E254750AFC0EFEFD" bold="true" box="[626,948,283,309]" pageId="64" pageNumber="65">
<collectingCountry id="92B67602C36EC730E254750AFC97FEFD" box="[626,813,283,309]" name="Costa Rica" pageId="64" pageNumber="65">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
P. N. Volcán Irazú,
<geoCoordinate id="8F955055C36EC730E489750DFA96FEFE" box="[1199,1324,284,310]" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="15" value="9.974999">9°58'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36EC730E12E7555FE23FE96" box="[264,409,324,350]" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="15" value="-83.85833">83°51'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36EC730E1827555FDBAFE96" box="[420,512,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="64" pageNumber="65" unit="m" value="3300.0">3300 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C36EC730E3B87555FBA1FE96" box="[926,1051,324,350]" pageId="64" pageNumber="65" value="1997-06-18">18.vi.1997</date>
(
<collectionCode id="8CB0AE57C36EC730E40D7555FB39FE96" box="[1067,1155,324,350]" pageId="64" pageNumber="65">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36EC730E11E757DFC57FE1E" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<typeStatus id="351A8830C36EC730E11E757DFE14FE4E" box="[312,430,364,390]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36EC730E1E6757AFD47FE4D" bold="true" box="[448,765,363,389]" pageId="64" pageNumber="65">
<collectingCountry id="92B67602C36EC730E1E6757AFDC3FE4D" box="[448,633,363,389]" name="Costa Rica" pageId="64" pageNumber="65">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC36EC730E32A757DFC17FE4E" box="[780,941,364,390]" pageId="64" pageNumber="65" type="generic">28 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C36EC730E459757DFB53FE4E" box="[1151,1257,364,390]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
;
<emphasis id="D8D5EA80C36EC730E4DD757AFEFDFE65" bold="true" pageId="64" pageNumber="65">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC36EC730E17E7585FE56FE66" box="[344,492,404,430]" pageId="64" pageNumber="65" type="generic">4 specimens</specimenCount>
, Estación Cuerici,
<quantity id="2D599B77C36EC730E2EA7585FCA4FE66" box="[716,798,404,430]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="64" pageNumber="65" unit="km" value="4.6">4.6 km</quantity>
E Villa Mills,
<geoCoordinate id="8F955055C36EC730E3F67585FBF6FE66" box="[976,1100,404,430]" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36EC730E4727585FB5EFE66" box="[1108,1252,404,430]" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="15" value="-83.666664">83°40'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36EC730E4D77585FE98FE1E" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="64" pageNumber="65" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C36EC730E2E775ADFC84FE1E" box="[705,830,444,470]" pageId="64" pageNumber="65" value="1997-06-26">26.vi.1997</date>
(all
<collectionCode id="8CB0AE57C36EC730E3AC75ADFC5BFE1E" box="[906,993,444,470]" pageId="64" pageNumber="65">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36EC731E11E75F2FC95FDEE" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="EA1E3692C36EC730E11E75F2FAB3FD56" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E11E75F2FE02FE35" bold="true" box="[312,440,483,509]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C36EC730E1ED75F5FD5DFE35" box="[459,743,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36EC730E1ED75F5FD5DFE35" box="[459,743,484,509]" italics="true" pageId="64" pageNumber="65">Seeversiella curtipennis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C36EC730E4DE75F5FED7FDED" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36EC730E4DE75F5FED7FDED" italics="true" pageId="64" pageNumber="65">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.9­2.0 times as long as eyes; elytra shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of the male tergum 7 (
<figureCitation id="729A2A17C36EC730E2F3764DFC86FDBE" box="[725,828,604,630]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="64" pageNumber="65">Fig. 247</figureCitation>
); tergum 8 with three pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C36EC730E2BD7695FCFBFD56" box="[667,833,644,670]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65">Figs. 251­254</figureCitation>
,
<figureCitation id="729A2A17C36EC730E36B7695FC0BFD56" box="[845,945,644,670]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="64" pageNumber="65">256­264</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C36EC730E4B27695FB46FD56" box="[1172,1276,644,670]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65">Fig. 255</figureCitation>
).
</paragraph>
<caption id="BEDE661AC36EC730E12E728FFD66F8D5" httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65" targetBox="[284,1301,785,1656]" targetPageId="64">
<paragraph id="EA1E3692C36EC730E12E728FFD66F8D5" blockId="64.[264,1323,1694,1821]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E12E728FFE67F97E" bold="true" box="[264,477,1694,1718]" pageId="64" pageNumber="65">FIGURES 251­255</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C36EC730E253728FFC6CF97D" authority="Gusarov" authorityName="Gusarov" box="[629,982,1694,1717]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C36EC730E253728FFCCEF97D" box="[629,884,1694,1717]" italics="true" pageId="64" pageNumber="65">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36EC730E3C6728FFBA1F97E" box="[992,1051,1694,1718]" pageId="64" pageNumber="65" rank="species">
<emphasis id="D8D5EA80C36EC730E3C6728FFBA1F97E" bold="true" box="[992,1051,1694,1718]" pageId="64" pageNumber="65">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254­255).
</paragraph>
</caption>
<paragraph id="EA1E3692C36FC731E11E750DFC95FDEE" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<taxonomicName id="2DA14D11C36FC731E11E750DFDEAFEFD" box="[312,592,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E11E750DFDEAFEFD" box="[312,592,284,309]" italics="true" pageId="65" pageNumber="66">Seeversiella curtipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36FC731E2CF750DFC35FEFD" box="[745,911,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C36FC731E2CF750DFC35FEFD" box="[745,911,284,309]" italics="true" pageId="65" pageNumber="66">S. scabricollis</emphasis>
</taxonomicName>
,
<taxonomicName id="2DA14D11C36FC731E3BB750DFBEDFEFD" box="[925,1111,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C36FC731E3BB750DFBEDFEFD" box="[925,1111,284,309]" italics="true" pageId="65" pageNumber="66">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C36FC731E4A9750DFAB7FEFD" box="[1167,1293,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C36FC731E4A9750DFAB7FEFD" box="[1167,1293,284,309]" italics="true" pageId="65" pageNumber="66">S. brunnea</emphasis>
</taxonomicName>
in having shorter elytra (shorter than pronotum); reduced wings (shorter than elytra); posterior margin of male tergum 8 without medial emargination (
<figureCitation id="729A2A17C36FC731E3FF757DFBFBFE4E" box="[985,1089,364,390]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Fig. 247</figureCitation>
); broad apex of the median lobe (in parameral view) (
<figureCitation id="729A2A17C36FC731E2B57585FC80FE66" box="[659,826,404,430]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­252</figureCitation>
; 194­196; 212­213; 239­240); long medial lamellae of the internal sac (
<figureCitation id="729A2A17C36FC731E27175ADFD45FE1E" box="[599,767,444,470]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 258­259</figureCitation>
; 201) protruding apically when the internal sac is retracted (
<figureCitation id="729A2A17C36FC731E18475F5FDF5FE36" box="[418,591,484,510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 256­257</figureCitation>
); lateral diverticula without sclerotized denticles (
<figureCitation id="729A2A17C36FC731E49775F5FA9DFE36" box="[1201,1319,484,510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 259</figureCitation>
; 200); and longer spermatheca (
<figureCitation id="729A2A17C36FC731E252761DFD5FFDEE" box="[628,741,524,550]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 255</figureCitation>
; 205).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36FC731E11E7625FDC4F9FE" pageId="65" pageNumber="66" type="description">
<paragraph id="EA1E3692C36FC731E11E7625FBDCFDBE" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<taxonomicName id="2DA14D11C36FC731E11E7625FDEBFD85" box="[312,593,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E11E7625FDEBFD85" box="[312,593,564,589]" italics="true" pageId="65" pageNumber="66">Seeversiella curtipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36FC731E2CB7625FC38FD85" box="[749,898,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36FC731E2CB7625FC38FD85" box="[749,898,564,589]" italics="true" pageId="65" pageNumber="66">S. lativentris</emphasis>
</taxonomicName>
in having weaker basal constriction of the apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C36FC731E366764DFC5DFDBE" box="[832,999,604,630]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­252</figureCitation>
; 269­270).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E7692FCCBFD56" blockId="65.[264,1324,284,1870]" box="[312,881,643,670]" pageId="65" pageNumber="66">
<emphasis id="D8D5EA80C36FC731E11E7692FE6AFD55" bold="true" box="[312,464,643,669]" pageId="65" pageNumber="66">Description.</emphasis>
Length
<quantity id="2D599B77C36FC731E21F7695FD7EFD56" box="[569,708,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="65" pageNumber="66" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6­3.1 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C36FC731E11E76BDFD8BFCF6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">Head surface matte, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 1.9­2.0 times as long as eyes. Antennal article 2 as long as 3, article 4 slightly transverse, 5­10 transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C36FC731E11E775DFEE8FBB6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C36FC731E40D775DFB6CFCAE" box="[1067,1238,844,870]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="65" pageNumber="66" unit="mm" value="0.53" valueMax="0.56" valueMin="0.5">0.50­0.56 mm</quantity>
, length
<quantity id="2D599B77C36FC731E12E7765FE0BFC46" box="[264,433,884,910]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="65" pageNumber="66" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
, width to length ratio 1.1, surface matte, with fine isodiametric microsculpture; punctation fine and weak, distance between punctures equals 1­2 times their diameter. Elytra wider and shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.2), 1.7 times wider than long, glossy, with fine and weak isodiametric microsculpture, and fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C36FC731E11E709DFC16FAD6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­4 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C36FC731E2D47115FC1AFAD6" box="[754,928,1284,1310]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Figs. 247, 249</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E713DFEFFFA76" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
In large males posterior angles of tergum 3 projecting as spines up to ½ as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C36FC731E4DD716DFE82FA76" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 male tergum 8; 248 male sternum 8; 249 female tergum 8; 250 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Fig. 247</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E71DDFC19F9C6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Aedeagus as in
<figureCitation id="729A2A17C36FC731E1D971DDFD13FA2E" box="[511,681,1484,1510]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­254</figureCitation>
,
<figureCitation id="729A2A17C36FC731E29F71DDFCA5FA2E" box="[697,799,1484,1510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">256­264</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C36FC731E30971E5FC2CF9C6" box="[815,918,1524,1550]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261 ­ 262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Fig. 260</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E720DFDC4F9FE" blockId="65.[264,1324,284,1870]" box="[312,638,1564,1590]" pageId="65" pageNumber="66">
Spermatheca as in
<figureCitation id="729A2A17C36FC731E233720DFDC1F9FE" box="[533,635,1564,1590]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Fig. 255</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36FC732E11E7252FC0DF8FF" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="discussion">
<paragraph id="EA1E3692C36FC731E11E7252FB57F886" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<emphasis id="D8D5EA80C36FC731E11E7252FE79F995" bold="true" box="[312,451,1603,1629]" pageId="65" pageNumber="66">Discussion.</emphasis>
Volcán Irazú and Cerro Cuerici, the localities where
<taxonomicName id="2DA14D11C36FC731E4687255FB49F995" box="[1102,1267,1604,1629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E4687255FB49F995" box="[1102,1267,1604,1629]" italics="true" pageId="65" pageNumber="66">S. curtipennis</emphasis>
</taxonomicName>
was found, are separated by the distance of
<quantity id="2D599B77C36FC731E2FF727DFC9FF94E" box="[729,805,1644,1670]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="65" pageNumber="66" unit="km" value="50.0">50 km</quantity>
and the gap as low as
<quantity id="2D599B77C36FC731E415727DFB2AF94E" box="[1075,1168,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="65" pageNumber="66" unit="m" value="1400.0">1400 m</quantity>
. Considering the distribution of the other species of
<taxonomicName id="2DA14D11C36FC731E2FB7285FCD1F965" box="[733,875,1684,1709]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36FC731E2FB7285FCD1F965" box="[733,875,1684,1709]" italics="true" pageId="65" pageNumber="66">Seeversiella</emphasis>
</taxonomicName>
in
<collectingCountry id="92B67602C36FC731E3B27285FBA3F966" box="[916,1049,1684,1710]" name="Costa Rica" pageId="65" pageNumber="66">Costa Rica</collectingCountry>
, and the fact that in
<taxonomicName id="2DA14D11C36FC731E5337285FE36F91D" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E5337285FE36F91D" italics="true" pageId="65" pageNumber="66">S. curtipennis</emphasis>
</taxonomicName>
the wings are reduced to vestiges that are shorter than elytra, it is clear that the two known populations of this species are isolated from each other. Despite this, both are identical in the structure of genitalia, including the internal sac. Apparently, the isolation of the two population is recent and has not allowed enough time for their divergence.
</paragraph>
<caption id="BEDE661AC36CC732E12E723AFE5EF8DB" httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="66" pageNumber="67" targetBox="[278,1308,342,1490]" targetPageId="66">
<paragraph id="EA1E3692C36CC732E12E723AFE5EF8DB" blockId="66.[264,1323,1579,1847]" pageId="66" pageNumber="67">
<emphasis id="D8D5EA80C36CC732E12E723AFE59F98B" bold="true" box="[264,483,1579,1603]" pageId="66" pageNumber="67">FIGURES 256­264</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C36CC732E2BE723DFBB9F98B" authority="Gusarov" authorityName="Gusarov" box="[664,1027,1580,1603]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C36CC732E2BE723DFC26F98B" box="[664,924,1580,1603]" italics="true" pageId="66" pageNumber="67">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C36CC732E435723AFBEEF98B" bold="true" box="[1043,1108,1579,1603]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="C3E657FBC36CC732E435723AFBEEF98B" box="[1043,1108,1579,1603]" pageId="66" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, Volcán Irazú, Costa Rica). 256 details of internal sac retracted into median lobe, abparameral view; 257 details of internal sac retracted into median lobe, parameral view; 258 medial lamellae, parameral view; 259 everted internal sac, parameral view; 260 apex of left paramere; 261­262 details of internal sac retracted into median lobe, lateral view; 263 right distal sclerite of internal sac; 264 everted internal sac, lateral view. Scale bar 0.2 mm (256­257, 259, 261­262, 264), 0.1 mm (258, 260, 263).
</paragraph>
</caption>
<paragraph id="EA1E3692C36CC732E12E7331FC0DF8FF" blockId="66.[264,1323,1579,1847]" box="[264,951,1824,1847]" pageId="66" pageNumber="67">LD lateral diverticulum of internal sac; ML medial lamellae.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36DC733E11E750AFE17FE4E" pageId="67" pageNumber="68" type="distribution">
<paragraph id="EA1E3692C36DC733E11E750AFE17FE4E" blockId="67.[264,1323,283,390]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="67" pageNumber="68">Distribution.</emphasis>
Known from Volcán Irazú and Cerro Cuerici,
<collectingCountry id="92B67602C36DC733E427750DFB3BFEFE" box="[1025,1153,284,310]" name="Costa Rica" pageId="67" pageNumber="68">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C36DC733E4B7750DFB43FEFE" box="[1169,1273,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="67" pageNumber="68">Fig. 388</figureCitation>
).
<emphasis id="D8D5EA80C36DC733E11E7552FDBDFE95" bold="true" box="[312,519,323,349]" pageId="67" pageNumber="68">Natural History.</emphasis>
<taxonomicName id="2DA14D11C36DC733E23F7555FC8EFE95" box="[537,820,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36DC733E23F7555FC8EFE95" box="[537,820,324,349]" italics="true" pageId="67" pageNumber="68">Seeversiella curtipennis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C36DC733E12E757DFE12FE4E" box="[264,424,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="67" pageNumber="68" unit="m" value="2950.0" valueMax="3300.0" valueMin="2600.0">2600­3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,269 @@
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<paragraph id="EA1E3692C313C74DE12E761AFC08FDED" blockId="61.[264,946,523,550]" box="[264,946,523,550]" pageId="61" pageNumber="62">
<heading id="B15681FEC313C74DE12E761AFC08FDED" bold="true" box="[264,946,523,550]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
<emphasis id="D8D5EA80C313C74DE12E761AFC08FDED" bold="true" box="[264,946,523,550]" pageId="61" pageNumber="62">
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<key id="9E3061A3C313C74DE12E761AFC08FDED" box="[264,946,523,550]" pageId="61" pageNumber="62">
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17.
<taxonomicName id="2DA14D11C313C74DE112761DFD1EFDED" ID-CoL="6YBCR" authority="Gusarov" authorityName="Gusarov" box="[308,676,523,550]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C313C74DE112761DFD88FDEE" bold="true" box="[308,562,524,550]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC313C74DE297761AFD49FDED" box="[689,755,523,549]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C313C74DE324761AFC10FDED" box="[770,938,523,549]" captionStart-0="FIGURES 235 ­ 238" captionStart-1="FIGURES 239 ­ 246" captionStartId-0="61.[264,382,1659,1683]" captionStartId-1="62.[264,383,1630,1654]" captionTargetBox-0="[313,1272,852,1625]" captionTargetBox-1="[313,1277,660,1592]" captionTargetId-0="figure@61.[304,1283,844,1632]" captionTargetId-1="figure@62.[302,1285,649,1603]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURES 235 ­ 238. Abdominal segment 8 of Seeversiella brunnea Gusarov, sp. n. (235 ­ 236, holotype; 237 ­ 238, paratype, P. N. Tapantí, Costa Rica). 235 male tergum 8; 236 male sternum 8; 237 female tergum 8; 238 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri-0="https://zenodo.org/record/156461/files/figure.png" httpUri-1="https://zenodo.org/record/156462/files/figure.png" pageId="61" pageNumber="62">Figs. 235­246</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C313C74DE12E7647FE29FD21" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="EA1E3692C313C74DE12E7647FB3BFD51" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E7647FE78FDB8" bold="true" box="[264,450,598,624]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE12E7647FEF9FDB8" box="[264,323,598,624]" pageId="61" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C313C74DE1C67646FDEAFDB9" box="[480,592,599,625]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C313C74DE2AC7647FC61FDB8" bold="true" box="[650,987,598,624]" pageId="61" pageNumber="62">
<collectingCountry id="92B67602C313C74DE2AC7647FCF0FDB8" box="[650,842,598,624]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
P. N. Tapantí,
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<geoCoordinate id="8F955055C313C74DE12E766EFE23FD51" box="[264,409,639,665]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.816666">83°49'00&quot;W</geoCoordinate>
,
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, berlese forest litter (R.Anderson),
<date id="9E1F1052C313C74DE3B8766EFBB7FD51" box="[926,1037,639,665]" pageId="61" pageNumber="62" value="1997-06-04">4.vi.1997</date>
(
<collectionCode id="8CB0AE57C313C74DE43A766EFBCEFD51" box="[1052,1140,639,665]" pageId="61" pageNumber="62">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C313C74DE11E76B6FE29FD21" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE11E76B6FE15FD09" box="[312,431,679,705]" pageId="61" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C313C74DE1E576B7FDC7FD08" box="[451,637,678,704]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
,, ditto but,
<geoCoordinate id="8F955055C313C74DE3E976B6FBF0FD09" box="[975,1098,679,705]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="15" value="9.741667">9°44'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C313C74DE47476B6FB59FD09" box="[1106,1251,679,705]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.808334">83°48'30&quot;W</geoCoordinate>
,
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(
<collectionCode id="8CB0AE57C313C74DE10876DEFE3CFD21" box="[302,390,719,745]" pageId="61" pageNumber="62">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C313C74DE12E726AFC4EF91F" blockId="61.[264,1323,1659,1751]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E726AFE53F95B" bold="true" box="[264,489,1659,1683]" pageId="61" pageNumber="62">FIGURES 235­238.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C313C74DE338726AFBD2F95A" authority="Gusarov" authorityName="Gusarov" box="[798,1128,1659,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C313C74DE338726AFBB8F95A" box="[798,1026,1659,1682]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C313C74DE451726AFB0DF95B" bold="true" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="C3E657FBC313C74DE451726AFB0DF95B" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(235­236, holotype; 237­238, paratype, P. N. Tapantí, Costa Rica). 235 male tergum 8; 236 male sternum 8; 237 female tergum 8; 238 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74EE11E733CFB24FE1E" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="EA1E3692C313C74EE11E733CFC51FE4E" blockId="61.[264,1323,1837,1904]" lastBlockId="62.[264,1323,284,590]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E733CFE0DF88F" bold="true" box="[312,439,1837,1863]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C313C74DE4F5733FFEFDF8A7" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C313C74DE4F5733FFEFDF8A7" italics="true" pageId="61" pageNumber="62">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 0.8­1.2 times as long as eyes; matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or partially reduced, 2­4 times as long as elytra; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C310C74EE1CB757DFD29FE4E" box="[493,659,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 239­245</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C310C74EE351757DFC64FE4E" box="[887,990,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Fig. 246</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C310C74EE11E7585FB24FE1E" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" italics="true" pageId="62" pageNumber="63">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" italics="true" pageId="62" pageNumber="63">S. furcativentris</emphasis>
</taxonomicName>
in having concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE32075ADFC14FE1E" box="[774,942,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 197­198; 214­215).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C310C74FE11E75F5FDC4FE1E" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
<paragraph id="EA1E3692C310C74EE11E75F5FD68FD86" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" italics="true" pageId="62" pageNumber="63">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; and concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE18D7625FDE9FD86" box="[427,595,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 228­229).
</paragraph>
<caption id="BEDE661AC310C74EE12E724FFD50F8EA" httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63" targetBox="[313,1277,660,1592]" targetPageId="62">
<paragraph id="EA1E3692C310C74EE12E724FFD50F8EA" blockId="62.[264,1324,1630,1863]" pageId="62" pageNumber="63">
<emphasis id="D8D5EA80C310C74EE12E724FFE5AF9BE" bold="true" box="[264,480,1630,1654]" pageId="62" pageNumber="63">FIGURES 239­246</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C310C74EE2A4724FFC72F9BD" authority="Gusarov" authorityName="Gusarov" box="[642,968,1630,1653]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C310C74EE2A4724FFCDEF9BD" box="[642,868,1630,1653]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C310C74EE3F3724FFBA8F9BE" bold="true" box="[981,1042,1630,1654]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="C3E657FBC310C74EE3F3724FFBA8F9BE" box="[981,1042,1630,1654]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(239­244, holotype; 245­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246).
</paragraph>
</caption>
<paragraph id="EA1E3692C310C74EE12E7321FE47F88F" blockId="62.[264,1324,1630,1863]" box="[264,509,1840,1863]" pageId="62" pageNumber="63">CB connecting band.</paragraph>
<paragraph id="EA1E3692C311C74FE11E750AFB5BFEFE" blockId="63.[264,1323,283,830]" box="[312,1249,283,310]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="63" pageNumber="64">Description.</emphasis>
Length
<quantity id="2D599B77C311C74FE21F750DFD7EFEFE" box="[569,708,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="63" pageNumber="64" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Pronotal width
<quantity id="2D599B77C311C74FE3AA750DFB8FFEFE" box="[908,1077,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.0" metricValueMin="4.4" pageId="63" pageNumber="64" unit="mm" value="0.47" valueMax="0.5" valueMin="0.44">0.44­0.50 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7555FBEFFE4E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
In all character states
<taxonomicName id="2DA14D11C311C74FE2117555FD0CFE95" box="[567,694,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2117555FD0CFE95" box="[567,694,324,349]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7585FD26FE66" blockId="63.[264,1323,283,830]" box="[312,668,404,430]" pageId="63" pageNumber="64">
Aedeagus as in
<figureCitation id="729A2A17C311C74FE1D77585FD2DFE66" box="[497,663,404,430]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­245</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E75ADFDC4FE1E" blockId="63.[264,1323,283,830]" box="[312,638,444,470]" pageId="63" pageNumber="64">
Spermatheca as in
<figureCitation id="729A2A17C311C74FE23375ADFDC1FE1E" box="[533,635,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Fig. 246</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E75F2FBBFFD0E" pageId="63" pageNumber="64" type="discussion">
<paragraph id="EA1E3692C311C74FE11E75F2FBBFFD0E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E75F2FE79FE35" bold="true" box="[312,451,483,509]" pageId="63" pageNumber="64">Discussion.</emphasis>
Despite the similarity of
<taxonomicName id="2DA14D11C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
to
<taxonomicName id="2DA14D11C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
in external characters and the shape of genitalia, the difference between
<taxonomicName id="2DA14D11C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
and the two species in the shape of the apex of median lobe (
<figureCitation id="729A2A17C311C74FE35B7625FB9FFD86" box="[893,1061,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 median lobe, parameral view; 240 apex of median lobe, parameral view; 241 median lobe, lateral view; 242 apex of median lobe, lateral view; 243 details of internal sac retracted into median lobe, lateral view; 244 apex of left paramere; 245 everted internal sac, parameral view; 246 spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­242</figureCitation>
; 194­198; 212­215) is of the same scale as the difference between
<taxonomicName id="2DA14D11C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
. Since no intermediate forms have been found,
<taxonomicName id="2DA14D11C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is considered to represent a separate species, possibly a local endemic of the area near Tapantí (
<figureCitation id="729A2A17C311C74FE3B776BDFC42FD0E" box="[913,1016,684,710]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E76C2FE17FCF6" pageId="63" pageNumber="64" type="distribution">
<paragraph id="EA1E3692C311C74FE11E76C2FB29FD26" blockId="63.[264,1323,283,830]" box="[312,1171,723,750]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76C2FE62FD25" bold="true" box="[312,472,723,749]" pageId="63" pageNumber="64">Distribution.</emphasis>
Known from the area near Tapantí,
<collectingCountry id="92B67602C311C74FE3A976C5FBB5FD26" box="[911,1039,724,750]" name="Costa Rica" pageId="63" pageNumber="64">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C311C74FE43976C5FB3DFD26" box="[1055,1159,724,750]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C311C74FE11E76EAFE17FCF6" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76EAFDB0FCDD" bold="true" box="[312,522,763,789]" pageId="63" pageNumber="64">Natural History.</emphasis>
<taxonomicName id="2DA14D11C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" italics="true" pageId="63" pageNumber="64">Seeversiella brunnea</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C311C74FE12E7735FE12FCF6" box="[264,424,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="1.7399999999999998" metricValueMin="1.5" pageId="63" pageNumber="64" unit="m" value="1620.0" valueMax="1740.0" valueMin="1500.0">1500­1740 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="6690E386CD041993E6B5C8C93D5AE505" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C317C74DE12E72B7FED2FE66" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella brevipennis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="62" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="58" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="2F04BCC81938C116CA5C4FDB180A847D">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="5EA46DE9C9654DCC953E3FCDC514A639">Gusarov, Vladimir I.</mods:namePart>
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<subSubSection id="A2BB6519C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58" type="key">
<paragraph id="EA1E3692C317C749E12E72B7FC62F908" blockId="57.[264,984,1702,1729]" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<heading id="B15681FEC317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<keyLead id="515095A7C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<keyStep id="51552E37C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<key id="9E3061A3C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
16.
<taxonomicName id="2DA14D11C317C749E11272B6FD70F908" ID-CoL="6YBCM" authority="Gusarov" authorityName="Gusarov" box="[308,714,1702,1729]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C317C749E11272B6FDEDF909" bold="true" box="[308,599,1703,1729]" italics="true" pageId="57" pageNumber="58">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC317C749E2F072B7FCA3F908" box="[726,793,1702,1728]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C317C749E30E72B7FC6AF908" box="[808,976,1702,1728]" captionStart-0="FIGURES 222 ­ 225" captionStart-1="FIGURES 226 ­ 229" captionStart-2="FIGURES 230 ­ 234" captionStartId-0="58.[264,382,1219,1243]" captionStartId-1="59.[264,383,1174,1198]" captionStartId-2="60.[264,382,1729,1753]" captionTargetBox-0="[284,1303,333,1185]" captionTargetBox-1="[307,1283,459,1141]" captionTargetBox-2="[294,1292,824,1693]" captionTargetId-0="figure@58.[273,1314,320,1192]" captionTargetId-1="figure@59.[296,1291,449,1147]" captionTargetId-2="figure@60.[281,1305,809,1702]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionTargetPageId-2="60" captionText-0="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 male tergum 8; 223 male sternum 8; 224 female tergum 8; 225 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-2="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156458/files/figure.png" httpUri-1="https://zenodo.org/record/156459/files/figure.png" httpUri-2="https://zenodo.org/record/156460/files/figure.png" pageId="57" pageNumber="58">Figs. 222­234</figureCitation>
)
</key>
</keyStep>
</emphasis>
</keyLead>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C317C74AE12E72E0FB6CFA58" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="EA1E3692C317C749E12E72E0FCC8F894" blockId="57.[264,1323,1777,1884]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72E0FE04F8C3" bold="true" box="[264,446,1777,1803]" pageId="57" pageNumber="58">
<typeStatus id="351A8830C317C749E12E72E0FEF9F8C3" box="[264,323,1777,1803]" pageId="57" pageNumber="58">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C317C749E1F372E3FDFFF8C4" box="[469,581,1778,1804]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C317C749E25E72E0FC31F8C3" bold="true" box="[632,907,1777,1803]" pageId="57" pageNumber="58">
<collectingCountry id="92B67602C317C749E25E72E0FD47F8C3" box="[632,765,1777,1803]" name="Panama" pageId="57" pageNumber="58">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
<quantity id="2D599B77C317C749E3B972E3FC4EF8C4" box="[927,1012,1778,1804]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="57" pageNumber="58" unit="km" value="5.9">5.9 km</quantity>
N Cerro Punta, Par. Nac. Volcán Barú,
<geoCoordinate id="8F955055C317C749E181730BFE43F8FC" box="[423,505,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="8.866667">8°52'N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C317C749E30D730BFC22F8FC" box="[811,920,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="15" value="8.366667">8°22'0&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C317C749E41A730BFB07F8FC" box="[1084,1213,1818,1844]" direction="west" orientation="longitude" pageId="57" pageNumber="58" precision="15" value="-82.566666">82°34'0&quot;W</geoCoordinate>
,
<quantity id="2D599B77C317C749E4EF730BFA9EF8FC" box="[1225,1316,1818,1844]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="57" pageNumber="58" unit="m" value="2150.0">2150 m</quantity>
, alder forest litter (R.Anderson),
<date id="9E1F1052C317C749E2A77353FD44F894" box="[641,766,1858,1884]" pageId="57" pageNumber="58" value="1995-06-14">14.vi.1995</date>
(
<collectionCode id="8CB0AE57C317C749E3287353FCDCF894" box="[782,870,1858,1884]" pageId="57" pageNumber="58">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC314C74AE12E70D2FBEAFAE8" httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="58" pageNumber="59" targetBox="[284,1303,333,1185]" targetPageId="58">
<paragraph id="EA1E3692C314C74AE12E70D2FBEAFAE8" blockId="58.[264,1323,1219,1312]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE12E70D2FE5CFB13" bold="true" box="[264,486,1219,1243]" pageId="58" pageNumber="59">FIGURES 222­225.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C314C74AE32C70D5FBC8FB12" authority="Gusarov" authorityName="Gusarov" box="[778,1138,1219,1243]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C314C74AE32C70D5FBB4FB13" box="[778,1038,1220,1243]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C314C74AE45B70D2FB00FB13" bold="true" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="C3E657FBC314C74AE45B70D2FB00FB13" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(222­223, holotype; 224­225, paratype, 5.9 km N Cerro Punta, Panama). 222 male tergum 8; 223 male sternum 8; 224 female tergum 8; 225 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C314C74AE11E7167FB6CFA58" blockId="58.[264,1324,1397,1904]" box="[312,1238,1397,1424]" pageId="58" pageNumber="59">
<typeStatus id="351A8830C314C74AE11E7167FE14FA58" box="[312,430,1398,1424]" pageId="58" pageNumber="59" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C314C74AE1E67164FD74FA47" bold="true" box="[448,718,1397,1423]" pageId="58" pageNumber="59">
<collectingCountry id="92B67602C314C74AE1E67164FDFFFA47" box="[448,581,1397,1423]" name="Panama" pageId="58" pageNumber="59">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
,, same data as the
<typeStatus id="351A8830C314C74AE3DD7167FBD8FA58" box="[1019,1122,1398,1424]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C314C74AE4547167FB70FA58" box="[1138,1226,1398,1424]" pageId="58" pageNumber="59">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74AE11E718CFCA0F930" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="EA1E3692C314C74AE11E718CFBF2F948" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE11E718CFE02FA7F" bold="true" box="[312,440,1437,1463]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C314C74AE4DE718FFED7FA17" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C314C74AE4DE718FFED7FA17" italics="true" pageId="58" pageNumber="59">Seeversiella</emphasis>
</taxonomicName>
by having brown or reddish brown body with yellowish brown elytra, abdominal segments 3­4, legs and mouthparts; large eyes (temple length to eye length ratio 1.6­2.0); glossy pronotum with weak isodiametric microsculpture and asperate punctation; elytra as long as pronotum; wings partially reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C314C74AE26D7277FD4AF948" box="[587,752,1638,1664]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Figs. 226­233</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C314C74AE3F27277FB86F948" box="[980,1084,1638,1664]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Fig. 234</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C314C74AE11E729FFCA0F930" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" italics="true" pageId="58" pageNumber="59">S. scabricollis</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; apex of median lobe in parameral view constricted basally (
<figureCitation id="729A2A17C314C74AE1D572CFFD20F930" box="[499,666,1758,1784]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­227</figureCitation>
; 194­196).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74CE11E7317FDC4FD0E" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59" type="description">
<paragraph id="EA1E3692C314C74AE11E7317FE56F8B8" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" italics="true" pageId="58" pageNumber="59">S. furcativentris</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; larger median lobe with longer apex (
<figureCitation id="729A2A17C314C74AE4D7733FFED7F8B8" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­229</figureCitation>
; 212­215).
</paragraph>
<paragraph id="EA1E3692C315C74BE11E750DFD3BFE4E" blockId="59.[264,1323,284,390]" pageId="59" pageNumber="60">
<taxonomicName id="2DA14D11C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" italics="true" pageId="59" pageNumber="60">S. brunnea</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C315C74BE17D757DFDB8FE4E" box="[347,514,364,390]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60">Figs. 228­229</figureCitation>
; 241­242).
</paragraph>
<caption id="BEDE661AC315C74BE12E7087FEFAFADD" httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60" targetBox="[307,1283,459,1141]" targetPageId="59">
<paragraph id="EA1E3692C315C74BE12E7087FEFAFADD" blockId="59.[264,1323,1174,1301]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE12E7087FE59FB66" bold="true" box="[264,483,1174,1198]" pageId="59" pageNumber="60">FIGURES 226­229</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C315C74BE2B37087FBB9FB65" authority="Gusarov" authorityName="Gusarov" box="[661,1027,1174,1197]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C315C74BE2B37087FC26FB65" box="[661,924,1174,1197]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC315C74BE4347087FBE9FB66" box="[1042,1107,1174,1198]" pageId="59" pageNumber="60" rank="species">
<emphasis id="D8D5EA80C315C74BE4347087FBE9FB66" bold="true" box="[1042,1107,1174,1198]" pageId="59" pageNumber="60">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229).
</paragraph>
</caption>
<paragraph id="EA1E3692C315C74BE11E717AFC1BFA66" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE11E717AFE75FA4D" bold="true" box="[312,463,1387,1413]" pageId="59" pageNumber="60">Description.</emphasis>
Length 2.6­3.0 mm. Body brown or reddish brown, elytra, abdominal segments 3 and 4, legs and mouthparts yellowish brown.
</paragraph>
<paragraph id="EA1E3692C315C74BE11E71ADFE87F986" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">Head surface glossy, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2­3 times their diameter. Temple length to eye length ratio 1.6­2.0. Antennal article 2 as long as 3, articles 4­10 transverse (ratio 1.3­ 1.6).</paragraph>
<paragraph id="EA1E3692C315C74BE11E724DFB27F8AE" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C315C74BE40D724DFB6CF9BE" box="[1067,1238,1628,1654]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="59" pageNumber="60" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53­0.56 mm</quantity>
, length
<quantity id="2D599B77C315C74BE12E7295FE1FF956" box="[264,421,1668,1694]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="59" pageNumber="60" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47­0.5 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine and weak isodiametric microsculpture; punctation fine and asperate, distance between punctures equal to 1­1.5 times their diameter. Elytra wider and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.4 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings partially reduced.
</paragraph>
<paragraph id="EA1E3692C312C74CE11E750DFC3CFE66" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 3­6 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C312C74CE2ED7585FCC3FE66" box="[715,889,404,430]" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 male tergum 8; 223 male sternum 8; 224 female tergum 8; 225 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Figs. 222, 224</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E75ADFEFFFD86" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
In males posterior angles of tergum 3 projecting as spines up to 2 times as long as tergum 3 (measured medially); tergum
<quantity id="2D599B77C312C74CE2E775F5FD4AFE36" box="[705,752,484,510]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="60" pageNumber="61" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial tubercle; posterior margin of male tergum 8 with emargination delimited by short projections (
<figureCitation id="729A2A17C312C74CE4DD761DFE83FD86" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 male tergum 8; 223 male sternum 8; 224 female tergum 8; 225 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Fig. 222</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E764DFD48FD56" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Aedeagus as in
<figureCitation id="729A2A17C312C74CE1D4764DFD23FDBE" box="[498,665,604,630]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Figs. 226­233</figureCitation>
. Apex of paramere broad, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C312C74CE2587695FD5FFD56" box="[638,741,644,670]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 233</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E76BDFDC4FD0E" blockId="60.[264,1324,284,710]" box="[312,638,684,710]" pageId="60" pageNumber="61">
Spermatheca as in
<figureCitation id="729A2A17C312C74CE23276BDFDC0FD0E" box="[532,634,684,710]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 234</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="BEDE661AC312C74CE12E72D0FB63F888" httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61" targetBox="[294,1292,824,1693]" targetPageId="60">
<paragraph id="EA1E3692C312C74CE12E72D0FB63F888" blockId="60.[264,1323,1729,1856]" pageId="60" pageNumber="61">
<emphasis id="D8D5EA80C312C74CE12E72D0FE67F911" bold="true" box="[264,477,1729,1753]" pageId="60" pageNumber="61">FIGURES 230­234</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C312C74CE25272D3FC6DF910" authority="Gusarov" authorityName="Gusarov" box="[628,983,1729,1753]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C312C74CE25272D3FCCCF911" box="[628,886,1730,1753]" italics="true" pageId="60" pageNumber="61">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C312C74CE3C772D0FBA6F911" bold="true" box="[993,1052,1729,1753]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3E657FBC312C74CE3C772D0FBA6F911" box="[993,1052,1729,1753]" pageId="60" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(230­233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230­231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230­232), 0.1 mm (233­234).
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74DE11E750AFED2FE66" pageId="61" pageNumber="62" type="distribution">
<paragraph id="EA1E3692C313C74DE11E750AFD25FE96" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="61" pageNumber="62">Distribution.</emphasis>
Known from Cerro Punta,
<collectingCountry id="92B67602C313C74DE302750DFC3BFEFE" box="[804,897,284,310]" name="Panama" pageId="61" pageNumber="62">Panama</collectingCountry>
(
<figureCitation id="729A2A17C313C74DE3B4750DFC40FEFE" box="[914,1018,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="61" pageNumber="62">Fig. 388</figureCitation>
). In this locality
<taxonomicName id="2DA14D11C313C74DE4EE750DFEEFFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE4EE750DFEEFFE95" italics="true" pageId="61" pageNumber="62">S. brevipennis</emphasis>
</taxonomicName>
is sympatric
<taxonomicName id="2DA14D11C313C74DE1D47555FD22FE95" box="[498,664,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C313C74DE1D47555FD22FE95" box="[498,664,324,349]" italics="true" pageId="61" pageNumber="62">S. scabricollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EA1E3692C313C74DE11E757AFED2FE66" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E757AFDBCFE4D" bold="true" box="[312,518,363,389]" pageId="61" pageNumber="62">Natural History.</emphasis>
<taxonomicName id="2DA14D11C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" italics="true" pageId="61" pageNumber="62">Seeversiella brevipennis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C313C74DE12E7585FEDEFE66" box="[264,356,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="61" pageNumber="62" unit="m" value="2150.0">2150 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,489 @@
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<heading id="B15681FEC318C746E12E71B2FC55FA75" bold="true" box="[264,1007,1443,1470]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="D8D5EA80C318C746E12E71B2FC55FA75" bold="true" box="[264,1007,1443,1470]" pageId="54" pageNumber="55">
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15.
<taxonomicName id="2DA14D11C318C746E11271B5FD5AFA75" ID-CoL="6YBCP" authority="Gusarov" authorityName="Gusarov" box="[308,736,1443,1470]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="furcativentris" status="sp. nov.">
<emphasis id="D8D5EA80C318C746E11271B5FDD4FA76" bold="true" box="[308,622,1444,1470]" italics="true" pageId="54" pageNumber="55">Seeversiella furcativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC318C746E2CB71B2FC8AFA75" box="[749,816,1443,1469]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C318C746E31971B2FC5CFA75" box="[831,998,1443,1469]" captionStart-0="FIGURES 208 ­ 211" captionStart-1="FIGURES 212 ­ 221" captionStartId-0="56.[264,383,1366,1390]" captionStartId-1="57.[264,383,1351,1375]" captionTargetBox-0="[299,1287,536,1332]" captionTargetBox-1="[279,1264,462,1310]" captionTargetId-0="figure@56.[292,1294,529,1339]" captionTargetId-1="figure@57.[252,1336,433,1340]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="FIGURES 208 ­ 211. Abdominal segment 8 of Seeversiella furcativentris Gusarov, sp. n. (208 ­ 209, holotype; 210 ­ 211, paratype, Zurquí de Moravia, Costa Rica). 208 male tergum 8; 209 male sternum 8; 210 female tergum 8; 211 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri-0="https://zenodo.org/record/156456/files/figure.png" httpUri-1="https://zenodo.org/record/156457/files/figure.png" pageId="54" pageNumber="55">Figs. 208­221</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C318C747E12E71FFFBD4FA8E" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="EA1E3692C318C746E12E71FFFCC8F9F8" blockId="54.[264,1325,1518,1904]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E12E71FFFE7BF9C0" bold="true" box="[264,449,1518,1544]" pageId="54" pageNumber="55">
<typeStatus id="351A8830C318C746E12E71FFFEF9F9C0" box="[264,323,1518,1544]" pageId="54" pageNumber="55">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C318C746E1F871FEFDF4F9C1" box="[478,590,1519,1545]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C318C746E2AE71FFFC45F9C0" bold="true" box="[648,1023,1518,1544]" pageId="54" pageNumber="55">
<collectingCountry id="92B67602C318C746E2AE71FFFCFDF9C0" box="[648,839,1518,1544]" name="Costa Rica" pageId="54" pageNumber="55">COSTA RICA</collectingCountry>
: Puntarenas:
</emphasis>
Monte Verde,
<quantity id="2D599B77C318C746E4EA71FEFA91F9C1" box="[1228,1323,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="54" pageNumber="55" unit="m" value="1800.0">1800 m</quantity>
(J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C318C746E2BF7207FD45F9F8" box="[665,767,1558,1584]" pageId="54" pageNumber="55" value="1989-05-05">5.v.1989</date>
(
<collectionCode id="8CB0AE57C318C746E3287207FCDCF9F8" box="[782,870,1558,1584]" pageId="54" pageNumber="55">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C318C747E11E722FFBD4FA8E" blockId="54.[264,1325,1518,1904]" lastBlockId="55.[264,1324,284,1830]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<typeStatus id="351A8830C318C746E11E722FFE14F990" box="[312,430,1598,1624]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C318C746E1E6722FFC9CF990" bold="true" box="[448,806,1598,1624]" pageId="54" pageNumber="55">
<collectingCountry id="92B67602C318C746E1E6722FFDC0F990" box="[448,634,1598,1624]" name="Costa Rica" pageId="54" pageNumber="55">COSTA RICA</collectingCountry>
: Puntarenas:
</emphasis>
<specimenCount id="FCA7FD1BC318C746E312722FFC7DF990" box="[820,967,1598,1624]" pageId="54" pageNumber="55" type="generic">5 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C318C746E4BF722FFAB9F990" box="[1177,1283,1598,1624]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
;, ditto but
<quantity id="2D599B77C318C746E15E7277FE6DF948" box="[376,471,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="54" pageNumber="55" unit="m" value="1580.0">1580 m</quantity>
, ex
<taxonomicName id="2DA14D11C318C746E22B7276FD2CF948" box="[525,662,1639,1664]" class="Agaricomycetes" family="Auriculariaceae" genus="Auricularia" kingdom="Animalia" order="Auriculariales" pageId="54" pageNumber="55" phylum="Basidiomycota" rank="genus">
<emphasis id="D8D5EA80C318C746E22B7276FD2CF948" box="[525,662,1639,1664]" italics="true" pageId="54" pageNumber="55">Auricularia</emphasis>
</taxonomicName>
sp.;, ditto but
<quantity id="2D599B77C318C746E34C7277FC73F948" box="[874,969,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="54" pageNumber="55" unit="m" value="1520.0">1520 m</quantity>
, flight intercept trap (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C318C746E21A729FFD08F960" box="[572,690,1678,1704]" pageId="54" pageNumber="55" value="1989-05-11">11.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E2E4729FFCE2F960" box="[706,856,1678,1704]" pageId="54" pageNumber="55" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C318C746E3F1729FFBF4F960" box="[983,1102,1678,1704]" pageId="54" pageNumber="55" value="1989-05-14">14.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E47B729FFB52F960" box="[1117,1256,1678,1704]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E11072A7FE24F918" box="[310,414,1718,1744]" pageId="54" pageNumber="55" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E18D72A7FD8EF918" box="[427,564,1718,1744]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but pitfall trap,
<date id="9E1F1052C318C746E31F72A7FC15F918" box="[825,943,1718,1744]" pageId="54" pageNumber="55" value="1989-05-24">24.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E39B72A7FBEBF918" box="[957,1105,1718,1744]" pageId="54" pageNumber="55" type="generic">2 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C318C746E4EC72A7FA9DF918" box="[1226,1319,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="54" pageNumber="55" unit="m" value="1550.0">1550 m</quantity>
, flight intercept trap,
<date id="9E1F1052C318C746E1DC72CFFDCBF930" box="[506,625,1758,1784]" pageId="54" pageNumber="55" value="1989-05-23">23.v.1989</date>
;, ditto but
<quantity id="2D599B77C318C746E32F72CFFCDCF930" box="[777,870,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="54" pageNumber="55" unit="m" value="1570.0">1570 m</quantity>
, flight intercept trap,
<date id="9E1F1052C318C746E44572CFFB71F930" box="[1123,1227,1758,1784]" pageId="54" pageNumber="55" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E4F172CFFEFCF8E8" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E1EF7317FD85F8E8" box="[457,575,1798,1824]" pageId="54" pageNumber="55" value="1989-05-11">11.v.1989</date>
;, ditto but
<quantity id="2D599B77C318C746E2CC7317FCF1F8E8" box="[746,843,1798,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="54" pageNumber="55" unit="m" value="1770.0">1770 m</quantity>
, ex
<taxonomicName id="2DA14D11C318C746E3A47316FC5DF8E8" box="[898,999,1799,1824]" class="Agaricomycetes" family="Tricholomataceae" genus="Collybia" kingdom="Fungi" order="Agaricales" pageId="54" pageNumber="55" phylum="Basidiomycota" rank="genus">
<emphasis id="D8D5EA80C318C746E3A47316FC5DF8E8" box="[898,999,1799,1824]" italics="true" pageId="54" pageNumber="55">Collybia</emphasis>
</taxonomicName>
sp.,
<date id="9E1F1052C318C746E40E7317FB25F8E8" box="[1064,1183,1798,1824]" pageId="54" pageNumber="55" value="1989-05-21">21.v.1989</date>
;, Monteverde reserve, trail near lab, flight intercept trap (C.Michalski),
<date id="9E1F1052C318C746E43A733FFB34F880" box="[1052,1166,1838,1864]" pageId="54" pageNumber="55" value="1993-06-03">3.vi.1993</date>
;
<specimenCount id="FCA7FD1BC318C746E4BB733FFA9DF880" box="[1181,1319,1838,1864]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E15F7347FE51F8B8" box="[377,491,1878,1904]" pageId="54" pageNumber="55" value="1993-06-01">1.vi.1993</date>
;
<specimenCount id="FCA7FD1BC318C746E1DD7347FD28F8B8" box="[507,658,1878,1904]" pageId="54" pageNumber="55" type="generic">9 specimens</specimenCount>
, Monte Verde, Cerro Amigos,
<quantity id="2D599B77C318C746E42A7347FBD6F8B8" box="[1036,1132,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="54" pageNumber="55" unit="m" value="1780.0">1780 m</quantity>
, flight intercept trap (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C319C747E2C3750DFCE6FEFE" box="[741,860,284,310]" pageId="55" pageNumber="56" value="1989-05-14">14.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E34A750DFC43FEFE" box="[876,1017,284,310]" pageId="55" pageNumber="56" type="generic">1 specimen</specimenCount>
, ditto but pitfall trap;
<specimenCount id="FCA7FD1BC319C747E529750DFE3FFE96" pageId="55" pageNumber="56" type="generic">11 specimens</specimenCount>
, ditto but flight intercept trap,
<date id="9E1F1052C319C747E2D27555FCD1FE96" box="[756,875,324,350]" pageId="55" pageNumber="56" value="1989-05-21">21.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E35E7555FBA6FE96" box="[888,1052,324,350]" pageId="55" pageNumber="56" type="generic">12 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E4B07555FAB6FE96" box="[1174,1292,324,350]" pageId="55" pageNumber="56" value="1989-05-24">24.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E53C7555FE3FFE4E" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DB757DFDC9FE4E" box="[509,627,364,390]" pageId="55" pageNumber="56" value="1989-05-17">17.v.1989</date>
;
<emphasis id="D8D5EA80C319C747E2A7757AFC7FFE4D" bold="true" box="[641,965,363,389]" pageId="55" pageNumber="56">Puntarenas / Guanacaste:</emphasis>
, Monte Verde, Cerro Amigos,
<quantity id="2D599B77C319C747E1647585FE1AFE66" box="[322,416,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="55" pageNumber="56" unit="m" value="1780.0">1780 m</quantity>
, sifted leaf litter (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C319C747E42D7585FBC9FE66" box="[1035,1139,404,430]" pageId="55" pageNumber="56" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E4A47585FA9FFE66" box="[1154,1317,404,430]" pageId="55" pageNumber="56" type="generic">11 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C319C747E15675ADFE71FE1E" box="[368,459,444,470]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="55" pageNumber="56" unit="m" value="1760.0">1760 m</quantity>
, but flight intercept trap,
<date id="9E1F1052C319C747E2D675ADFCDDFE1E" box="[752,871,444,470]" pageId="55" pageNumber="56" value="1989-05-12">12.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E35475ADFBBFFE1E" box="[882,1029,444,470]" pageId="55" pageNumber="56" type="generic">6 specimens</specimenCount>
, ditto but pitfall traps;
<specimenCount id="FCA7FD1BC319C747E52D75ADFE3FFE36" pageId="55" pageNumber="56" type="generic">29 specimens</specimenCount>
, ditto but flight intercept trap,
<date id="9E1F1052C319C747E2CB75F5FCDEFE36" box="[749,868,484,510]" pageId="55" pageNumber="56" value="1989-05-10">10.v.1989</date>
;
<emphasis id="D8D5EA80C319C747E35775F2FBB7FE35" bold="true" box="[881,1037,483,509]" pageId="55" pageNumber="56">Guanacaste:</emphasis>
, Guanacaste Conservacion Area, Cacao Field Station,
<quantity id="2D599B77C319C747E2EF761DFC95FDEE" box="[713,815,524,550]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="55" pageNumber="56" unit="m" value="1400.0">1400 m</quantity>
, wet cloud forest litter (R.S.Anderson),
<date id="9E1F1052C319C747E12E7625FE39FD86" box="[264,387,564,590]" pageId="55" pageNumber="56" value="1996-02-12">12.ii.1996</date>
;
<emphasis id="D8D5EA80C319C747E1B57622FDB8FD85" bold="true" box="[403,514,563,589]" pageId="55" pageNumber="56">Heredia:</emphasis>
,, Cerro Chompipe,
<quantity id="2D599B77C319C747E3637625FC3FFD86" box="[837,901,564,590]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="km" value="2.0">2 km</quantity>
N Monte de la Cruz,
<geoCoordinate id="8F955055C319C747E4847625FA96FD86" box="[1186,1324,564,590]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="10.088888">10°05'20&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E12E764DFE23FDBE" box="[264,409,604,630]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-84.075">84°04'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E181764DFDBCFDBE" box="[423,518,604,630]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="2000.0">2000 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C319C747E394764DFB89FDBE" box="[946,1075,604,630]" pageId="55" pageNumber="56" value="1997-06-12">12.vi.1997</date>
;
<specimenCount id="FCA7FD1BC319C747E464764DFB5DFDBE" box="[1090,1255,604,630]" pageId="55" pageNumber="56" type="generic">13 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C319C747E11E7695FE37FD56" box="[312,397,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="55" pageNumber="56" unit="m" value="2100.0">2100m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C319C747E36E7695FC55FD56" box="[840,1007,644,670]" pageId="55" pageNumber="56" value="1997-06-12" valueMax="1997-06-27" valueMin="1997-06-12">12­27.vi.1997</date>
;
<emphasis id="D8D5EA80C319C747E3D97692FBC3FD55" bold="true" box="[1023,1145,643,669]" pageId="55" pageNumber="56">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC319C747E4A97695FA9CFD56" box="[1167,1318,644,670]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, Zurquí de Moravia [
<geoCoordinate id="8F955055C319C747E1D576BDFDECFD0E" box="[499,598,684,710]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="925" value="10.05">10°03'N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E27B76BDFD73FD0E" box="[605,713,684,710]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="925" value="-84.01667">84°01'W</geoCoordinate>
],
<quantity id="2D599B77C319C747E2FA76BDFC8DFD0E" box="[732,823,684,710]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="55" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, malaise traps (P.Hanson),
<date id="9E1F1052C319C747E45476BDFB76FD0E" box="[1138,1228,684,710]" pageId="55" pageNumber="56" value="1992-04">iv.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E4FE76BDFEEBFD26" pageId="55" pageNumber="56" type="generic">7 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1E076C5FDA3FD26" box="[454,537,724,750]" pageId="55" pageNumber="56" value="1992-05">v.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E20276C5FD0DFD26" box="[548,695,724,750]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E30B76C5FC33FD26" box="[813,905,724,750]" pageId="55" pageNumber="56" value="1992-06">vi.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E3B376C5FB92FD26" box="[917,1064,724,750]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E4BB76C5FAB8FD26" box="[1181,1282,724,750]" pageId="55" pageNumber="56" value="1992-07">vii.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E52876C5FE3FFCDE" pageId="55" pageNumber="56" type="generic">10 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DA76EDFDE8FCDE" box="[508,594,764,790]" pageId="55" pageNumber="56" value="1993-02">ii.1993</date>
;
<specimenCount id="FCA7FD1BC319C747E27876EDFD48FCDE" box="[606,754,764,790]" pageId="55" pageNumber="56" type="generic">6 specimens</specimenCount>
, ditto but iii­iv.1993;
<specimenCount id="FCA7FD1BC319C747E3D776EDFB2EFCDE" box="[1009,1172,764,790]" pageId="55" pageNumber="56" type="generic">36 specimens</specimenCount>
, ditto but ivv.1993;
<specimenCount id="FCA7FD1BC319C747E1417735FE41FCF6" box="[359,507,804,830]" pageId="55" pageNumber="56" type="generic">7 specimens</specimenCount>
, ditto but ix­x.1993;
<specimenCount id="FCA7FD1BC319C747E2D37735FC33FCF6" box="[757,905,804,830]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but xi­xii.1993;
<specimenCount id="FCA7FD1BC319C747E4B57735FA9DFCF6" box="[1171,1319,804,830]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E150775DFE71FCAE" box="[374,459,844,870]" pageId="55" pageNumber="56" value="1994-02">ii.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E1FF775DFDD4FCAE" box="[473,622,844,870]" pageId="55" pageNumber="56" type="generic">8 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E2CF775DFCFDFCAE" box="[745,839,844,870]" pageId="55" pageNumber="56" value="1994-03">iii.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E373775DFC50FCAE" box="[853,1002,844,870]" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E443775DFB05FCAE" box="[1125,1215,844,870]" pageId="55" pageNumber="56" value="1994-04">iv.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E4EB775DFEF3FC46" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1F67765FD99FC46" box="[464,547,884,910]" pageId="55" pageNumber="56" value="1994-05">v.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E2137765FD74FC46" box="[565,718,884,910]" pageId="55" pageNumber="56" type="generic">9 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E3737765FC5AFC46" box="[853,992,884,910]" pageId="55" pageNumber="56" value="1995-01-01" valueMax="1995-01-31" valueMin="1995-01-01">1­31.i.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E3D47765FB31FC46" box="[1010,1163,884,910]" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E5357765FE3DFC7E" pageId="55" pageNumber="56" value="1995-04-01" valueMax="1995-04-30" valueMin="1995-04-01">1­ 30.iv.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E1BE778DFD8AFC7E" box="[408,560,924,950]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E292778DFCF7FC7E" box="[692,845,924,950]" pageId="55" pageNumber="56" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">1­30.ix.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E379778DFC4CFC7E" box="[863,1014,924,950]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E45D778DFAB6FC7E" box="[1147,1292,924,950]" pageId="55" pageNumber="56" value="1995-10-01" valueMax="1995-10-30" valueMin="1995-10-01">1­30.x.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E53B778DFEC0FC16" pageId="55" pageNumber="56" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DC77D5FD29FC16" box="[506,659,964,990]" pageId="55" pageNumber="56" value="1996-06-01" valueMax="1996-06-30" valueMin="1996-06-01">1­30.vi.1996</date>
;,
<quantity id="2D599B77C319C747E2EE77D5FCACFC16" box="[712,790,964,990]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="55" pageNumber="56" unit="km" value="26.0">26 km</quantity>
N San Isidro,
<quantity id="2D599B77C319C747E3EB77D5FB90FC16" box="[973,1066,964,990]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="55" pageNumber="56" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson),
<date id="9E1F1052C319C747E4EE77D5FA98FC16" box="[1224,1314,964,990]" pageId="55" pageNumber="56" value="1992-11">xi.1992</date>
­
<date id="9E1F1052C319C747E12E77FDFEEFFBCE" box="[264,341,1004,1030]" pageId="55" pageNumber="56" value="1993-01">i.1993</date>
;,, San Antonio de Escazú,
<quantity id="2D599B77C319C747E2E177FDFC99FBCE" box="[711,803,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="2000.0">2000 m</quantity>
, flight intercept trap (C.
<collectingCountry id="92B67602C319C747E41E77FDFB3CFBCE" box="[1080,1158,1004,1030]" name="Indonesia" pageId="55" pageNumber="56">Flores</collectingCountry>
&amp; P.Hanson),
<date id="9E1F1052C319C747E12E7005FE0DFBE6" box="[264,439,1044,1070]" pageId="55" pageNumber="56" value="1996-09-01" valueMax="1996-10-30" valueMin="1996-09-01">1.ix­30.x.1996</date>
;
<emphasis id="D8D5EA80C319C747E1E37002FD06FBE5" bold="true" box="[453,700,1043,1069]" pageId="55" pageNumber="56">San Jose / Cartago:</emphasis>
, km 45, Int. Amer. Hwy.,
<quantity id="2D599B77C319C747E43D7005FBE2FBE6" box="[1051,1112,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="56" unit="km" value="6.0">6 km</quantity>
NE El Empalme,
<geoCoordinate id="8F955055C319C747E12E702DFE3EFB9E" box="[264,388,1084,1110]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="9.75">9°45'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E1AB702DFDA4FB9E" box="[397,542,1084,1110]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-83.975">83°58'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E20B702DFD31FB9E" box="[557,651,1084,1110]" metricMagnitude="3" metricUnit="m" metricValue="1.975" pageId="55" pageNumber="56" unit="m" value="1975.0">1975 m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C319C747E419702DFB62FB9E" box="[1087,1240,1084,1110]" pageId="55" pageNumber="56" value="1997-06-08" valueMax="1997-06-26" valueMin="1997-06-08">8­26.vi.1997</date>
;
<emphasis id="D8D5EA80C319C747E4C0702AFE84FBB5" bold="true" pageId="55" pageNumber="56">Cartago:</emphasis>
10,
<specimenCount id="FCA7FD1BC319C747E1897075FDE8FBB6" box="[431,594,1124,1150]" pageId="55" pageNumber="56" type="generic">26 specimens</specimenCount>
, La Cangreja,
<quantity id="2D599B77C319C747E3247075FCDAFBB6" box="[770,864,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="55" pageNumber="56" unit="m" value="1950.0">1950 m</quantity>
, (P.Hanson), ix­xii.1992;
<specimenCount id="FCA7FD1BC319C747E4B57075FA9DFBB6" box="[1171,1319,1124,1150]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but iii­v.1992;
<specimenCount id="FCA7FD1BC319C747E1DC709DFD24FB6E" box="[506,670,1164,1190]" pageId="55" pageNumber="56" type="generic">17 specimens</specimenCount>
, ditto but vi­vii.1992; 2, ditto but
<quantity id="2D599B77C319C747E443709DFB79FB6E" box="[1125,1219,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="55" pageNumber="56" unit="m" value="1450.0">1450 m</quantity>
, malaise trap,
<date id="9E1F1052C319C747E16270A5FE4BFB06" box="[324,497,1204,1230]" pageId="55" pageNumber="56" value="1995-04-01" valueMax="1995-05-30" valueMin="1995-04-01">1.iv­30.v.1995</date>
;, Tapantí, (M.L.Jameson),
<date id="9E1F1052C319C747E36070A5FC7DFB06" box="[838,967,1204,1230]" pageId="55" pageNumber="56" value="1992-06-23">23.vi.1992</date>
;
<emphasis id="D8D5EA80C319C747E3F570A2FB5AFB05" bold="true" box="[979,1248,1203,1229]" pageId="55" pageNumber="56">
<collectingCountry id="92B67602C319C747E3F570A2FBE2FB05" box="[979,1112,1203,1229]" name="Panama" pageId="55" pageNumber="56">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
,,
<quantity id="2D599B77C319C747E12E70CDFEE1FB3E" box="[264,347,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="55" pageNumber="56" unit="km" value="5.6">5.6 km</quantity>
N Boquete, La Culebra Trail,
<geoCoordinate id="8F955055C319C747E2E870CDFCF3FB3E" box="[718,841,1244,1270]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="8.823055">8°49'23&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E37570CDFC59FB3E" box="[851,995,1244,1270]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-82.42166">82°25'18&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E3D770CDFBF5FB3E" box="[1009,1103,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="55" pageNumber="56" unit="m" value="1800.0">1800 m</quantity>
, cloud forest litter (R.S.Anderson),
<date id="9E1F1052C319C747E1E97115FDEAFAD6" box="[463,592,1284,1310]" pageId="55" pageNumber="56" value="1996-06-15">15.vi.1996</date>
; 3, ditto but 6.0 km NE Boquete,
<geoCoordinate id="8F955055C319C747E3D97115FBEBFAD6" box="[1023,1105,1284,1310]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="925" value="8.8">8°48'N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E47E7115FB05FAD6" box="[1112,1215,1284,1310]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="925" value="-82.433334">82°26'W</geoCoordinate>
,
<quantity id="2D599B77C319C747E4ED7115FA9DFAD6" box="[1227,1319,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="55" pageNumber="56" unit="m" value="1650.0">1650 m</quantity>
, flight intercept traps (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C319C747E33D713DFC05FA8E" box="[795,959,1324,1350]" pageId="55" pageNumber="56" value="1996-06-14" valueMax="1996-06-19" valueMin="1996-06-14">14­19.vi.1996</date>
(all
<collectionCode id="8CB0AE57C319C747E42D713DFBD8FA8E" box="[1035,1122,1324,1350]" pageId="55" pageNumber="56">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C319C747E11E7142FBF8F996" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="EA1E3692C319C747E11E7142FEFFF9C6" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<emphasis id="D8D5EA80C319C747E11E7142FE0DFAA5" bold="true" box="[312,439,1363,1389]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C319C747E1E07145FD43FAA5" box="[454,761,1364,1389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E1E07145FD43FAA5" box="[454,761,1364,1389]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C319C747E4DE7145FED7FA5D" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C319C747E4DE7145FED7FA5D" italics="true" pageId="55" pageNumber="56">Seeversiella</emphasis>
</taxonomicName>
by brown body; large eyes (temples as long as eyes); matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or partially reduced, longer than elytra; the distinct shape of aedeagus (
<figureCitation id="729A2A17C319C747E34F71DDFBABFA2E" box="[873,1041,1484,1510]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­220</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C319C747E4DA71DDFE82F9C6" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Fig. 221</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C319C747E11E720DFBF8F996" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E720DFDD7F9FD" box="[312,621,1564,1589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E720DFDD7F9FD" box="[312,621,1564,1589]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E337720DFC01F9FD" box="[785,955,1564,1589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C319C747E337720DFC01F9FD" box="[785,955,1564,1589]" italics="true" pageId="55" pageNumber="56">S. scabricollis</emphasis>
</taxonomicName>
in having the apex of median lobe constricted basally (in parameral view) (
<figureCitation id="729A2A17C319C747E33D7255FC79F996" box="[795,963,1604,1630]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­213</figureCitation>
; 194­196).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C319C748E11E727DFDC7F983" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="description">
<paragraph id="EA1E3692C319C747E11E727DFC89F966" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E727DFDD6F94D" box="[312,620,1644,1669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E727DFDD6F94D" box="[312,620,1644,1669]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E32B727DFC34F94D" box="[781,910,1644,1669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C319C747E32B727DFC34F94D" box="[781,910,1644,1669]" italics="true" pageId="55" pageNumber="56">S. brunnea</emphasis>
</taxonomicName>
in having straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C319C747E22B7285FD0EF966" box="[525,692,1684,1710]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 214­215</figureCitation>
; 241­242).
</paragraph>
<paragraph id="EA1E3692C319C747E11E72ADFCB0F8EE" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E72ADFDCBF91D" box="[312,625,1724,1749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E72ADFDCBF91D" box="[312,625,1724,1749]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E30472ADFC75F91D" box="[802,975,1724,1749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C319C747E30472ADFC75F91D" box="[802,975,1724,1749]" italics="true" pageId="55" pageNumber="56">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; and smaller median lobe with shorter apex (
<figureCitation id="729A2A17C319C747E1C5731DFD31F8EE" box="[483,651,1804,1830]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­215</figureCitation>
; 226­229).
</paragraph>
<paragraph id="EA1E3692C316C748E11E750AFCABFE96" blockId="56.[264,1323,283,430]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="56" pageNumber="57">Description.</emphasis>
Length
<quantity id="2D599B77C316C748E219750DFD76FEFE" box="[575,716,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="56" pageNumber="57" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C316C748E3B7750DFB86FEFE" box="[913,1084,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="56" pageNumber="57" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
. Body brown, with darker head and lighter legs and mouthparts.
</paragraph>
<paragraph id="EA1E3692C316C748E11E757DFC0DFE66" blockId="56.[264,1323,283,430]" pageId="56" pageNumber="57">
In all character states
<taxonomicName id="2DA14D11C316C748E219757DFD41FE4D" box="[575,763,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E219757DFD41FE4D" box="[575,763,364,389]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C316C748E3FE757DFB3AFE4D" box="[984,1152,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E3FE757DFB3AFE4D" box="[984,1152,364,389]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<caption id="BEDE661AC316C748E12E7147FBF5FA7A" httpUri="https://zenodo.org/record/156456/files/figure.png" pageId="56" pageNumber="57" targetBox="[299,1287,536,1332]" targetPageId="56">
<paragraph id="EA1E3692C316C748E12E7147FBF5FA7A" blockId="56.[264,1325,1366,1458]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E12E7147FE5EFAA6" bold="true" box="[264,484,1366,1390]" pageId="56" pageNumber="57">FIGURES 208­211.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C316C748E2D87147FBCCFAA5" authority="Gusarov" authorityName="Gusarov" box="[766,1142,1366,1389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris" status="sp. nov.">
<emphasis id="D8D5EA80C316C748E2D87147FBAEFAA5" box="[766,1044,1366,1389]" italics="true" pageId="56" pageNumber="57">Seeversiella furcativentris</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C316C748E4A67147FB01FAA6" bold="true" box="[1152,1211,1366,1390]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="C3E657FBC316C748E4A67147FB01FAA6" box="[1152,1211,1366,1390]" pageId="56" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(208­209, holotype; 210­211, paratype, Zurquí de Moravia, Costa Rica). 208 male tergum 8; 209 male sternum 8; 210 female tergum 8; 211 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C316C748E11E7218FD21F9EB" blockId="56.[264,1324,1545,1891]" box="[312,667,1545,1571]" pageId="56" pageNumber="57">
Aedeagus as in
<figureCitation id="729A2A17C316C748E1D67218FD2DF9EB" box="[496,663,1545,1571]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="56" pageNumber="57">Figs. 212­220</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C316C748E11E7220FDC7F983" blockId="56.[264,1324,1545,1891]" box="[312,637,1585,1611]" pageId="56" pageNumber="57">
Spermatheca as in
<figureCitation id="729A2A17C316C748E2327220FDC3F983" box="[532,633,1585,1611]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 median lobe, parameral view; 213 apex of median lobe, parameral view; 214 apex of median lobe, lateral view; 215 median lobe, lateral view; 216 apex of left paramere, 217 details of internal sac retracted into median lobe, abparameral view; 218 details of internal sac retracted into median lobe, lateral view; 219 right distal sclerite of internal sac; 220 left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="56" pageNumber="57">Fig. 221</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C316C748E11E7249FD88F8AB" pageId="56" pageNumber="57" type="discussion">
<paragraph id="EA1E3692C316C748E11E7249FD88F8AB" blockId="56.[264,1324,1545,1891]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E11E7249FE79F9BA" bold="true" box="[312,451,1624,1650]" pageId="56" pageNumber="57">Discussion.</emphasis>
Despite the fact that
<taxonomicName id="2DA14D11C316C748E2ED7248FC3CF9BA" box="[715,902,1625,1650]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E2ED7248FC3CF9BA" box="[715,902,1625,1650]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C316C748E3E77248FBD2F9BA" box="[961,1128,1625,1650]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E3E77248FBD2F9BA" box="[961,1128,1625,1650]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
are very similar in their genitalia, the difference in the shape of the apex of median lobe is stable and no intermediate forms have been observed. In one locality both species have been collected in the same sample, which also confirms their status of separate species. The two species appear to be parapatric:
<taxonomicName id="2DA14D11C316C748E20E72E8FD5BF8DA" box="[552,737,1785,1810]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E20E72E8FD5BF8DA" box="[552,737,1785,1810]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
occurs mostly below
<quantity id="2D599B77C316C748E3C672E8FB83F8DB" box="[992,1081,1785,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="m" value="2000.0">2000 m</quantity>
(has never been collected above
<quantity id="2D599B77C316C748E1877330FE47F8F3" box="[417,509,1825,1851]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="56" pageNumber="57" unit="m" value="2100.0">2100 m</quantity>
), while
<taxonomicName id="2DA14D11C316C748E27F7330FD45F8F2" box="[601,767,1825,1850]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E27F7330FD45F8F2" box="[601,767,1825,1850]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
occurs mostly above
<quantity id="2D599B77C316C748E3DA7330FBEEF8F3" box="[1020,1108,1825,1851]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="m" value="2000.0">2000 m</quantity>
(occasionally collected as low as
<quantity id="2D599B77C316C748E1EE7358FD9FF8AB" box="[456,549,1865,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="56" pageNumber="57" unit="m" value="1450.0">1450 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C317C749E11E750AFC59FE4E" pageId="57" pageNumber="58" type="distribution">
<paragraph id="EA1E3692C317C749E11E750AFBA5FEFE" blockId="57.[264,1323,283,390]" box="[312,1055,283,310]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="57" pageNumber="58">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C317C749E2A3750DFCBFFEFE" box="[645,773,284,310]" name="Costa Rica" pageId="57" pageNumber="58">Costa Rica</collectingCountry>
and
<collectingCountry id="92B67602C317C749E318750DFC21FEFE" box="[830,923,284,310]" name="Panama" pageId="57" pageNumber="58">Panama</collectingCountry>
(
<figureCitation id="729A2A17C317C749E38D750DFBA9FEFE" box="[939,1043,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="57" pageNumber="58">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C317C749E11E7552FC59FE4E" blockId="57.[264,1323,283,390]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E11E7552FDB3FE95" bold="true" box="[312,521,323,349]" pageId="57" pageNumber="58">Natural History.</emphasis>
<taxonomicName id="2DA14D11C317C749E2397555FCEFFE95" box="[543,853,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C317C749E2397555FCEFFE95" box="[543,853,324,349]" italics="true" pageId="57" pageNumber="58">Seeversiella furcativentris</emphasis>
</taxonomicName>
was collected in forest litter and with flight intercept and malaise traps, at altitudes of
<quantity id="2D599B77C317C749E318757DFC65FE4E" box="[830,991,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="57" pageNumber="58" unit="m" value="1750.0" valueMax="2100.0" valueMin="1400.0">1400­2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,647 @@
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<mods:title id="0752D9A2180494B5354A7E032ACFA5A4">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C31FC741E12E727AFB90F94D" box="[264,1066,1643,1670]" pageId="49" pageNumber="50" type="key">
<paragraph id="EA1E3692C31FC741E12E727AFB90F94D" blockId="49.[264,1066,1643,1670]" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<heading id="B15681FEC31FC741E12E727AFB90F94D" bold="true" box="[264,1066,1643,1670]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="D8D5EA80C31FC741E12E727AFB90F94D" bold="true" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<key id="9E3061A3C31FC741E12E727AFB90F94D" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<keyStep id="51552E37C31FC741E12E727AFB90F94D" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<keyLead id="515095A7C31FC741E12E727AFB90F94D" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
14.
<taxonomicName id="2DA14D11C31FC741E112727DFD7CF94D" ID-CoL="6YBDD" authority="Gusarov" authorityName="Gusarov" box="[308,710,1643,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31FC741E112727DFDEEF94E" bold="true" box="[308,596,1644,1670]" italics="true" pageId="49" pageNumber="50">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31FC741E2F5727AFCAFF94D" box="[723,789,1643,1669]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C31FC741E302727AFC0BF94D" box="[804,945,1643,1669]" captionStart="FIGURES 10 ­ 16" captionStartId="6.[264,383,1647,1671]" captionTargetBox="[319,1271,331,1612]" captionTargetId="figure@6.[304,1283,320,1620]" captionTargetPageId="6" captionText="FIGURES 10 ­ 16. Details of Seeversiella globicollis (Bernhauer) (10, Anticosti, Quebec; 13, Waterton Lakes National Park, Alberta), S. scabricollis Gusarov, sp. n. (11, 15, paratype from Volcán Barva, Costa Rica) and S. micralymma Gusarov, sp. n. (12, 14, 16, paratype from Cerro Buenavista, Costa Rica). 10 ­ 12 pronotum; 13 ­ 14 mesometathorax; 15 male tergum 3 and paratergites; 16 male tergum 3. Scale bar 0.4 mm (10 ­ 11, 15), 0.3 mm (12 ­ 13, 16), 0.2 mm (14)." httpUri="https://zenodo.org/record/156423/files/figure.png" pageId="49" pageNumber="50">Figs. 11, 15</figureCitation>
,
<figureCitation id="729A2A17C31FC741E399727AFB98F94D" box="[959,1058,1643,1669]" captionStart-0="FIGURES 190 ­ 193" captionStart-1="FIGURES 194 ­ 198" captionStart-2="FIGURES 199 ­ 207" captionStartId-0="51.[264,383,1774,1798]" captionStartId-1="52.[264,383,1328,1352]" captionStartId-2="53.[264,383,1638,1662]" captionTargetBox-0="[282,1304,939,1740]" captionTargetBox-1="[289,1303,584,1286]" captionTargetBox-2="[306,1280,457,1604]" captionTargetId-0="figure@51.[275,1312,929,1748]" captionTargetId-1="figure@52.[272,1315,569,1301]" captionTargetId-2="figure@53.[299,1288,449,1611]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 male tergum 8; 191 male sternum 8; 192 female tergum 8; 193 female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-2="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156453/files/figure.png" httpUri-1="https://zenodo.org/record/156454/files/figure.png" httpUri-2="https://zenodo.org/record/156455/files/figure.png" pageId="49" pageNumber="50">190­207</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31FC743E12E72A7FB0CFCAE" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="EA1E3692C31FC741E12E72A7FECFF8E8" blockId="49.[264,1323,1718,1904]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E12E72A7FE7BF918" bold="true" box="[264,449,1718,1744]" pageId="49" pageNumber="50">
<typeStatus id="351A8830C31FC741E12E72A7FEF9F918" box="[264,323,1718,1744]" pageId="49" pageNumber="50">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C31FC741E1FA72A7FDF6F918" box="[476,588,1718,1744]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C31FC741E2A372A7FC68F918" bold="true" box="[645,978,1718,1744]" pageId="49" pageNumber="50">
<collectingCountry id="92B67602C31FC741E2A372A7FCF9F918" box="[645,835,1718,1744]" name="Costa Rica" pageId="49" pageNumber="50">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Sendero a Cerro Chirripó,
<geoCoordinate id="8F955055C31FC741E12E72CFFE3EF930" box="[264,388,1758,1784]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="15" value="9.446389">9°26'47&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31FC741E1B072CFFD9DF930" box="[406,551,1758,1784]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="15" value="-83.53667">83°32'12&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31FC741E21B72CFFD1FF930" box="[573,677,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="49" pageNumber="50" unit="m" value="2800.0">2800 m</quantity>
, mixed oak forest litter (R.Anderson),
<date id="9E1F1052C31FC741E48B72CFFA91F930" box="[1197,1323,1758,1784]" pageId="49" pageNumber="50" value="1999-06-27">27.vi.1999</date>
(
<collectionCode id="8CB0AE57C31FC741E1367317FED2F8E8" box="[272,360,1798,1824]" pageId="49" pageNumber="50">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C31FC742E11E733FFA91FA8E" blockId="49.[264,1323,1718,1904]" lastBlockId="50.[264,1325,284,1870]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<typeStatus id="351A8830C31FC741E11E733FFE15F880" box="[312,431,1838,1864]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C31FC741E1E3733FFCBAF880" bold="true" box="[453,768,1838,1864]" pageId="49" pageNumber="50">
<collectingCountry id="92B67602C31FC741E1E3733FFD3AF880" box="[453,640,1838,1864]" name="Costa Rica" pageId="49" pageNumber="50">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC31FC741E335733FFC0DF880" box="[787,951,1838,1864]" pageId="49" pageNumber="50" type="generic">42 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C31FC741E4B2733FFB44F880" box="[1172,1278,1838,1864]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC31FC741E52A733FFE3FF8B8" pageId="49" pageNumber="50" type="generic">50 specimens</specimenCount>
, ditto but
<geoCoordinate id="8F955055C31FC741E2227347FD3AF8B8" box="[516,640,1878,1904]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="15" value="9.443611">9°26'37&quot;N</geoCoordinate>
83°32'14&quot;,
<quantity id="2D599B77C31FC741E3377347FCCAF8B8" box="[785,880,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="49" pageNumber="50" unit="m" value="2950.0">2950 m</quantity>
, oak forest litter;, Cerro Chirripó, Base Crestones,
<geoCoordinate id="8F955055C31CC742E1C5750DFDE4FEFE" box="[483,606,284,310]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.454722">9°27'17&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E257750DFCB8FEFE" box="[625,770,284,310]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.50445">83°30'16&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E33C750DFC38FEFE" box="[794,898,284,310]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="50" pageNumber="51" unit="m" value="3350.0">3350 m</quantity>
, elfin forest litter (R.Anderson),
<date id="9E1F1052C31CC742E12E7555FE32FE96" box="[264,392,324,350]" pageId="50" pageNumber="51" value="1999-06-25">25.vi.1999</date>
;
<specimenCount id="FCA7FD1BC31CC742E1B97555FDF0FE96" box="[415,586,324,350]" pageId="50" pageNumber="51" type="generic">17 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E2797555FD1FFE96" box="[607,677,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
S Villa Mills,
<quantity id="2D599B77C31CC742E3517555FC66FE96" box="[887,988,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="m" value="3000.0">3000 m</quantity>
, ridgetop oak forest litter (R.S.Anderson),
<date id="9E1F1052C31CC742E1E9757DFDF0FE4E" box="[463,586,364,390]" pageId="50" pageNumber="51" value="1996-02-10">10.ii.1996</date>
;, 2.0 km E Villa Mills,
<geoCoordinate id="8F955055C31CC742E35F757DFC4FFE4E" box="[889,1013,364,390]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3DA757DFB36FE4E" box="[1020,1164,364,390]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.69722">83°41'50&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4BF757DFB4FFE4E" box="[1177,1269,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="50" pageNumber="51" unit="m" value="2750.0">2750 m</quantity>
, oak forest litter (R.S.Anderson),
<date id="9E1F1052C31CC742E24A7585FD5DFE66" box="[620,743,404,430]" pageId="50" pageNumber="51" value="1998-02-15">15.ii.1998</date>
;, Cerro Buenavista,
<geoCoordinate id="8F955055C31CC742E42A7585FB32FE66" box="[1036,1160,404,430]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.717222">9°43'02&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4B07585FA9CFE66" box="[1174,1318,404,430]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.75389">83°45'14&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E12E75ADFED3FE1E" box="[264,361,444,470]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="50" pageNumber="51" unit="m" value="3300.0">3300 m</quantity>
, subparamo litter (R.Anderson);
<specimenCount id="FCA7FD1BC31CC742E2D875ADFC1FFE1E" box="[766,933,444,470]" pageId="50" pageNumber="51" type="generic">10 specimens</specimenCount>
, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E49375ADFB4CFE1E" box="[1205,1270,444,470]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="51" unit="km" value="4.0">4 km</quantity>
NE Cañon,
<quantity id="2D599B77C31CC742E14375F5FE7BFE36" box="[357,449,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="50" pageNumber="51" unit="m" value="2350.0">2350 m</quantity>
, malaise trap (P.Hanson),
<date id="9E1F1052C31CC742E2DF75F5FCFCFE36" box="[761,838,484,510]" pageId="50" pageNumber="51" value="1995-01">i.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E37275F5FC52FE36" box="[852,1000,484,510]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E44475F5FAB6FE36" box="[1122,1292,484,510]" pageId="50" pageNumber="51" value="1995-01-01" valueMax="1995-03-30" valueMin="1995-01-01">1.i­30.iii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C75F5FE3FFDEE" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E225761DFD20FDEE" box="[515,666,524,550]" pageId="50" pageNumber="51" value="1995-04-01" valueMax="1995-04-30" valueMin="1995-04-01">1­30.iv.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E28F761DFC89FDEE" box="[681,819,524,550]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E397761DFBFAFDEE" box="[945,1088,524,550]" pageId="50" pageNumber="51" value="1995-05-01" valueMax="1995-05-30" valueMin="1995-05-01">1­30.v.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E469761DFB5FFDEE" box="[1103,1253,524,550]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E1667625FE62FD86" box="[320,472,564,590]" pageId="50" pageNumber="51" value="1995-06-01" valueMax="1995-06-30" valueMin="1995-06-01">1­30.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E1D67625FD34FD86" box="[496,654,564,590]" pageId="50" pageNumber="51" type="generic">6 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E3027625FC7FFD86" box="[804,965,564,590]" pageId="50" pageNumber="51" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1­30.vii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E3FA7625FBC0FD86" box="[988,1146,564,590]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E5377625FE23FDBE" pageId="50" pageNumber="51" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">1­ 30.viii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E18F764DFD8FFDBE" box="[425,565,604,630]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E290764DFCFAFDBE" box="[694,832,604,630]" pageId="50" pageNumber="51" value="1996-01-01" valueMax="1996-01-30" valueMin="1996-01-01">1­30.i.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E376764DFC66FDBE" box="[848,988,604,630]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E47B764DFAB0FDBE" box="[1117,1290,604,630]" pageId="50" pageNumber="51" value="1996-04-01" valueMax="1996-05-30" valueMin="1996-04-01">1.iv­30.v.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C764DFE3FFD56" pageId="50" pageNumber="51" type="generic">7 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E1DF7695FD20FD56" box="[505,666,644,670]" pageId="50" pageNumber="51" value="1996-07-01" valueMax="1996-07-30" valueMin="1996-07-01">1­30.vii.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E2807695FC82FD56" box="[678,824,644,670]" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E38A7695FBFEFD56" box="[940,1092,644,670]" pageId="50" pageNumber="51" value="1996-09-01" valueMax="1996-09-30" valueMin="1996-09-01">1­30.ix.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E4767695FB59FD56" box="[1104,1251,644,670]" pageId="50" pageNumber="51" type="generic">4 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E4C87695FA93FD56" box="[1262,1321,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
N P. N. Volcán Irazú,
<geoCoordinate id="8F955055C31CC742E22E76BDFD28FD0E" box="[520,658,684,710]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.0">10°00'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E2BC76BDFC93FD0E" box="[666,809,684,710]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85">83°51'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E31076BDFC28FD0E" box="[822,914,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="50" pageNumber="51" unit="m" value="2900.0">2900 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E12E76C5FE32FD26" box="[264,392,724,750]" pageId="50" pageNumber="51" value="1997-06-18">18.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E1B376C5FD92FD26" box="[405,552,724,750]" pageId="50" pageNumber="51" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E21576C5FDD4FD26" box="[563,622,724,750]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
N P. N. Volcán Irazú,
<geoCoordinate id="8F955055C31CC742E35376C5FC4BFD26" box="[885,1009,724,750]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.995833">9°59'45&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3DE76C5FB32FD26" box="[1016,1160,724,750]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85">83°51'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4B276C5FB4AFD26" box="[1172,1264,724,750]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="50" pageNumber="51" unit="m" value="2750.0">2750 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E25F76EDFD43FCDE" box="[633,761,764,790]" pageId="50" pageNumber="51" value="1997-06-18">18.vi.1997</date>
;, Madreselva, Finca Los Lagos, A. C. Amistad,
<quantity id="2D599B77C31CC742E11C7735FE63FCF6" box="[314,473,804,830]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="50" pageNumber="51" unit="m" value="2300.0" valueMax="2600.0" valueMin="2000.0">2000­2600 m</quantity>
,
<date id="9E1F1052C31CC742E1C27735FD36FCF6" box="[484,652,804,830]" pageId="50" pageNumber="51" value="1993-06-12" valueMax="1993-06-19" valueMin="1993-06-12">12­19.vi.1993</date>
; 2, ditto but
<date id="9E1F1052C31CC742E3627735FBA9FCF6" box="[836,1043,804,830]" pageId="50" pageNumber="51" value="1993-06-28" valueMax="1993-07-10" valueMin="1993-06-28">28.vi­10.vii.1993</date>
;, ditto but
<date id="9E1F1052C31CC742E48E7735FABEFCF6" box="[1192,1284,804,830]" pageId="50" pageNumber="51" value="1993-09">ix.1993</date>
;, ditto but
<date id="9E1F1052C31CC742E155775DFE7DFCAE" box="[371,455,844,870]" pageId="50" pageNumber="51" value="1993-10">x.1993</date>
;
<emphasis id="D8D5EA80C31CC742E1F2775AFD73FCAD" bold="true" box="[468,713,843,869]" pageId="50" pageNumber="51">San Jose / Cartago:</emphasis>
<specimenCount id="FCA7FD1BC31CC742E2FF775DFCC1FCAE" box="[729,891,844,870]" pageId="50" pageNumber="51" type="generic">11 specimens</specimenCount>
, km 71, Int. Amer. Hwy., nr. Tres de Junio,
<geoCoordinate id="8F955055C31CC742E17F7765FE6EFC46" box="[345,468,884,910]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.628889">9°37'44&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1F87765FDD4FC46" box="[478,622,884,910]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.836945">83°50'13&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E25A7765FD61FC46" box="[636,731,884,910]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="50" pageNumber="51" unit="m" value="2900.0">2900 m</quantity>
, wet cloud forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4807765FA9DFC46" box="[1190,1319,884,910]" pageId="50" pageNumber="51" value="1999-06-23">23.vi.1999</date>
; 3, km 69, Int. Amer. Hwy., nr. Tres de Junio,
<geoCoordinate id="8F955055C31CC742E37D778DFC6DFC7E" box="[859,983,924,950]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.658333">9°39'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3C6778DFBCBFC7E" box="[992,1137,924,950]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85833">83°51'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E459778DFB64FC7E" box="[1151,1246,924,950]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31CC742E27C77D5FD49FC16" box="[602,755,964,990]" pageId="50" pageNumber="51" value="1997-06-07" valueMax="1997-06-19" valueMin="1997-06-07">7­19.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E32777D5FC18FC16" box="[769,930,964,990]" pageId="50" pageNumber="51" type="generic">11 specimens</specimenCount>
, km 78, Int. Amer. Hwy., nr. Ojo de Agua,
<geoCoordinate id="8F955055C31CC742E15B77FDFE43FBCE" box="[381,505,1004,1030]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.608334">9°36'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E22477FDFD28FBCE" box="[514,658,1004,1030]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.83889">83°50'20&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E28677FDFD47FBCE" box="[672,765,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
, cloud forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4B477FDFAB6FBCE" box="[1170,1292,1004,1030]" pageId="50" pageNumber="51" value="1999-07-08">8.vii.1999</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C77FDFE3FFBE6" pageId="50" pageNumber="51" type="generic">9 specimens</specimenCount>
, km 72, Int. Amer. Hwy.,
<quantity id="2D599B77C31CC742E29B7005FD40FBE6" box="[701,762,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
W Ojo de Agua,
<geoCoordinate id="8F955055C31CC742E3E97005FBF0FBE6" box="[975,1098,1044,1070]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.625">9°37'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4757005FB59FBE6" box="[1107,1251,1044,1070]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.84167">83°50'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4D77005FE98FB9E" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="50" pageNumber="51" unit="m" value="2950.0">2950 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E2F1702DFCF2FB9E" box="[727,840,1084,1110]" pageId="50" pageNumber="51" value="1997-06-07">7.vi.1997</date>
;, km 87, Int. Amer. Hwy., nr. Cerro Buenavista,
<geoCoordinate id="8F955055C31CC742E1BF7075FDAFFBB6" box="[409,533,1124,1150]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.608334">9°36'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E23A7075FD17FBB6" box="[540,685,1124,1150]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.76667">83°46'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E29F7075FCACFBB6" box="[697,790,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="50" pageNumber="51" unit="m" value="3150.0">3150 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4927075FA9CFBB6" box="[1204,1318,1124,1150]" pageId="50" pageNumber="51" value="1997-06-08">8.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E12E709DFE21FB6E" box="[264,411,1164,1190]" pageId="50" pageNumber="51" type="generic">8 specimens</specimenCount>
, Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C31CC742E38A709DFB92FB6E" box="[940,1064,1164,1190]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E409709DFB7AFB6E" box="[1071,1216,1164,1190]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4EA709DFA93FB6E" box="[1228,1321,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="50" pageNumber="51" unit="m" value="3200.0">3200 m</quantity>
, berlese leaf litter (R.Anderson),
<date id="9E1F1052C31CC742E2AD70A5FCB1FB06" box="[651,779,1204,1230]" pageId="50" pageNumber="51" value="1998-06-18">18.vi.1998</date>
;, 2, km 55, Int. Amer. Hwy.,
<quantity id="2D599B77C31CC742E49670A5FB57FB06" box="[1200,1261,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
S El Empalme,
<geoCoordinate id="8F955055C31CC742E1AF70CDFDBEFB3E" box="[393,516,1244,1270]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.708333">9°42'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E22870CDFD25FB3E" box="[526,671,1244,1270]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.95">83°57'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E28B70CDFCB1FB3E" box="[685,779,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="50" pageNumber="51" unit="m" value="2350.0">2350 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E49270CDFA9CFB3E" box="[1204,1318,1244,1270]" pageId="50" pageNumber="51" value="1997-06-08">8.vi.1997</date>
;
<emphasis id="D8D5EA80C31CC742E12E7112FEC4FAD5" bold="true" box="[264,382,1283,1309]" pageId="50" pageNumber="51">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC31CC742E1A97115FD8BFAD6" box="[399,561,1284,1310]" pageId="50" pageNumber="51" type="generic">15 specimens</specimenCount>
, Estación Cuerici,
<quantity id="2D599B77C31CC742E3367115FCD9FAD6" box="[784,867,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="50" pageNumber="51" unit="km" value="4.6">4.6 km</quantity>
E Villa Mills,
<geoCoordinate id="8F955055C31CC742E4357115FB35FAD6" box="[1043,1167,1284,1310]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4B07115FA9DFAD6" box="[1174,1319,1284,1310]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.666664">83°40'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E12E713DFEDFFA8E" box="[264,357,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E32C713DFC31FA8E" box="[778,907,1324,1350]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E3BE713DFB96FA8E" box="[920,1068,1324,1350]" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E480713DFA9CFA8E" box="[1190,1318,1324,1350]" pageId="50" pageNumber="51" value="1997-06-19">19.vi.1997</date>
;
</paragraph>
<paragraph id="EA1E3692C31CC743E13A7145FA97FD86" blockId="50.[264,1325,284,1870]" lastBlockId="51.[264,1325,284,870]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
, 2.0 km E Villa Mills,
<geoCoordinate id="8F955055C31CC742E21E7145FD0EFAA6" box="[568,692,1364,1390]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.558333">9°33'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E2987145FCF5FAA6" box="[702,847,1364,1390]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.7">83°42'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E37B7145FC01FAA6" box="[861,955,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E16A716DFE77FA5E" box="[332,461,1404,1430]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
;, 2, km 97, Int. Amer. Hwy., Villa Mills,
<quantity id="2D599B77C31CC742E428716DFBD6FA5E" box="[1038,1132,1404,1430]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="m" value="3000.0">3000 m</quantity>
, flight intercept traps (S. &amp; J.Peck),
<date id="9E1F1052C31CC742E23371B5FD2CFA76" box="[533,662,1444,1470]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
; 3,, Cerros de Escazú,
<quantity id="2D599B77C31CC742E43271B5FBEDFA76" box="[1044,1111,1444,1470]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
S San Antonio,
<geoCoordinate id="8F955055C31CC742E12E71DDFE3EFA2E" box="[264,388,1484,1510]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.891666">9°53'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1A871DDFDA5FA2E" box="[398,543,1484,1510]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.15">84°09'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E20871DDFD37FA2E" box="[558,653,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="50" pageNumber="51" unit="m" value="1650.0">1650 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E41A71DDFB07FA2E" box="[1084,1213,1484,1510]" pageId="50" pageNumber="51" value="1997-06-13">13.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E4EA71DDFEF3F9C6" pageId="50" pageNumber="51" type="generic">8 specimens</specimenCount>
, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E26571E5FDC5F9C6" box="[579,639,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="51" unit="km" value="6.0">6 km</quantity>
N San Gerardo,
<quantity id="2D599B77C31CC742E36071E5FC25F9C6" box="[838,927,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
(P.Hanson), viii­ix.1992;
<specimenCount id="FCA7FD1BC31CC742E4F771E5FEF3F9FE" pageId="50" pageNumber="51" type="generic">4 specimens</specimenCount>
, ditto but x­xii.1992;
<specimenCount id="FCA7FD1BC31CC742E270720DFD57F9FE" box="[598,749,1564,1590]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E348720DFC72F9FE" box="[878,968,1564,1590]" pageId="50" pageNumber="51" value="1992-04">iv.1992</date>
; 2, malaise trap, ditto but
<date id="9E1F1052C31CC742E12E7255FED9F996" box="[264,355,1604,1630]" pageId="50" pageNumber="51" value="1993-04">iv.1993</date>
;, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E2A07255FD68F996" box="[646,722,1604,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="51" unit="km" value="19.0">19 km</quantity>
S and
<quantity id="2D599B77C31CC742E3007255FCDEF996" box="[806,868,1604,1630]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
W of Empalme,
<quantity id="2D599B77C31CC742E4147255FB34F996" box="[1074,1166,1604,1630]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, malaise trap (P.Hanson), ix­x.1992; 7, 26 km N San Isidro,
<quantity id="2D599B77C31CC742E347727DFC00F94E" box="[865,954,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="50" pageNumber="51" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson),
<date id="9E1F1052C31CC742E472727DFB0AF94E" box="[1108,1200,1644,1670]" pageId="50" pageNumber="51" value="1992-11">xi.1992</date>
­
<date id="9E1F1052C31CC742E49F727DFABEF94E" box="[1209,1284,1644,1670]" pageId="50" pageNumber="51" value="1993-01">i.1993</date>
;, ditto but vi­vii.1990;, ditto but ix­x.1992;
<specimenCount id="FCA7FD1BC31CC742E33C7285FC14F966" box="[794,942,1684,1710]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E39D7285FC42F966" box="[955,1016,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
W Empalme, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E14872ADFE70F91E" box="[366,458,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="50" pageNumber="51" unit="m" value="2300.0">2300 m</quantity>
(P.Hanson),
<date id="9E1F1052C31CC742E24172ADFCB2F91E" box="[615,776,1724,1750]" pageId="50" pageNumber="51" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1­30.vii.1995</date>
, malaise trap;
<emphasis id="D8D5EA80C31CC742E39C72AAFB93F91D" bold="true" box="[954,1065,1723,1749]" pageId="50" pageNumber="51">Heredia:</emphasis>
2, 2,
<specimenCount id="FCA7FD1BC31CC742E4EA72ADFEF3F936" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, Porrosati,
<quantity id="2D599B77C31CC742E1F672F5FDB4F936" box="[464,526,1764,1790]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="51" unit="km" value="6.0">6 km</quantity>
N San José de la Montana,
<geoCoordinate id="8F955055C31CC742E37972F5FC53F936" box="[863,1001,1764,1790]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.091666">10°05'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3D472F5FB38F936" box="[1010,1154,1764,1790]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.11667">84°07'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4A972F5FB56F936" box="[1167,1260,1764,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="51" unit="m" value="1900.0">1900 m</quantity>
, berlese forest litter (R.Anderson);
<specimenCount id="FCA7FD1BC31CC742E2BF731DFCF9F8EE" box="[665,835,1804,1830]" pageId="50" pageNumber="51" type="generic">44 specimens</specimenCount>
, P. N. Braulio Carrillo Volcán Barva,
<geoCoordinate id="8F955055C31CC742E12E7325FE28F886" box="[264,402,1844,1870]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.125">10°07'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1BC7325FD90F886" box="[410,554,1844,1870]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.125">84°07'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E2117325FD29F886" box="[567,659,1844,1870]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4127325FB09F886" box="[1076,1203,1844,1870]" pageId="50" pageNumber="51" value="1997-06-11">11.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC743E4E67325FEF3FEFE" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="generic">21 specimens</specimenCount>
, ditto but, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31DC743E3B7750DFB82FEFE" box="[913,1080,284,310]" pageId="51" pageNumber="52" value="1997-06-11" valueMax="1997-06-27" valueMin="1997-06-11">11­27.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31DC743E46D750DFB5EFEFE" box="[1099,1252,284,310]" pageId="51" pageNumber="52" type="generic">4 specimens</specimenCount>
, Vara Blanca, Finca Georgina,
<quantity id="2D599B77C31DC743E2147555FD37FE96" box="[562,653,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="51" pageNumber="52" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson), vi­vii.1990;
<emphasis id="D8D5EA80C31DC743E39C7552FB97FE95" bold="true" box="[954,1069,323,349]" pageId="51" pageNumber="52">Alajuela:</emphasis>
<specimenCount id="FCA7FD1BC31DC743E4197555FB59FE96" box="[1087,1251,324,350]" pageId="51" pageNumber="52" type="generic">28 specimens</specimenCount>
, P. N. Volcán Poás,
<geoCoordinate id="8F955055C31DC743E19B757DFDFCFE4E" box="[445,582,364,390]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="10.191667">10°11'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E27E757DFD53FE4E" box="[600,745,364,390]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-84.23333">84°14'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E326757DFCD2FE4E" box="[768,872,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="51" pageNumber="52" unit="m" value="2500.0">2500 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31DC743E12E7585FEC0FE66" box="[264,378,404,430]" pageId="51" pageNumber="52" value="1997-06-06">6.vi.1997</date>
;,, ditto but flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31DC743E3DF7585FB28FE66" box="[1017,1170,404,430]" pageId="51" pageNumber="52" value="1997-06-06" valueMax="1997-06-28" valueMin="1997-06-06">6­28.vi.1997</date>
;
<emphasis id="D8D5EA80C31DC743E4877582FEC7FE1D" bold="true" pageId="51" pageNumber="52">
<collectingCountry id="92B67602C31DC743E4877582FA9CFE65" box="[1185,1318,403,429]" name="Panama" pageId="51" pageNumber="52">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
<specimenCount id="FCA7FD1BC31DC743E1AA75ADFD94FE1E" box="[396,558,444,470]" pageId="51" pageNumber="52" type="generic">31 specimens</specimenCount>
,
<quantity id="2D599B77C31DC743E21F75ADFD31FE1E" box="[569,651,444,470]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="51" pageNumber="52" unit="km" value="5.9">5.9 km</quantity>
NE Cerro Punta, Par. Nac. Volcán Barú,
<geoCoordinate id="8F955055C31DC743E44B75ADFB7AFE1E" box="[1133,1216,444,470]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="8.866667">8°52'N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C31DC743E1FF75F5FDF2FE36" box="[473,584,484,510]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.366667">8°22'0&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C31DC743E2CD75F5FCE8FE36" box="[747,850,484,510]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="-82.566666">82°34'W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E37875F5FC03FE36" box="[862,953,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="51" pageNumber="52" unit="m" value="2100.0">2100 m</quantity>
, flight intercept traps (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C31DC743E193761DFDE7FDEE" box="[437,605,524,550]" pageId="51" pageNumber="52" value="1995-06-14" valueMax="1995-06-16" valueMin="1995-06-14">14­16.vi.1995</date>
;, ditto but
<quantity id="2D599B77C31DC743E322761DFCD8FDEE" box="[772,866,524,550]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="52" unit="m" value="2400.0">2400 m</quantity>
, bamboo forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E12E7625FE33FD86" box="[264,393,564,590]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E1B37625FD82FD86" box="[405,568,564,590]" pageId="51" pageNumber="52" type="generic">12 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C31DC743E28B7625FCB3FD86" box="[685,777,564,590]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
, alder forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E4817625FA92FD86" box="[1191,1320,564,590]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
</paragraph>
<paragraph id="EA1E3692C31DC743E13A764DFB0CFCAE" blockId="51.[264,1325,284,870]" pageId="51" pageNumber="52">
,
<quantity id="2D599B77C31DC743E10D764DFE37FDBE" box="[299,397,604,630]" metricMagnitude="4" metricUnit="m" metricValue="2.77" pageId="51" pageNumber="52" unit="km" value="27.7">27.7 km</quantity>
W Volcán Hartmann's Finca,
<quantity id="2D599B77C31DC743E2D5764DFCEAFDBE" box="[755,848,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E37B764DFC6DFDBE" box="[861,983,604,630]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.863334">8°51'48&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E3C6764DFBCBFDBE" box="[992,1137,604,630]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.74333">82°44'36&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E458764DFB61FDBE" box="[1150,1243,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450 m</quantity>
, forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E22B7695FD34FD56" box="[525,654,644,670]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E2BF7695FC81FD56" box="[665,827,644,670]" pageId="51" pageNumber="52" type="generic">22 specimens</specimenCount>
, NNE of Las Nubes @ continental divide,
<quantity id="2D599B77C31DC743E12E76BDFEDBFD0E" box="[264,353,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.4899999999999998" pageId="51" pageNumber="52" unit="m" value="2490.0">2490 m</quantity>
(A.R.Gillogly),
<date id="9E1F1052C31DC743E20576BDFD70FD0E" box="[547,714,684,710]" pageId="51" pageNumber="52" value="1995-06-14" valueMax="1995-06-15" valueMin="1995-06-14">14­15.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E2F076BDFCD3FD0E" box="[726,873,684,710]" pageId="51" pageNumber="52" type="generic">5 specimens</specimenCount>
, N of Las Nubes,
<quantity id="2D599B77C31DC743E41D76BDFB61FD0E" box="[1083,1243,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.45" metricValueMin="2.35" pageId="51" pageNumber="52" unit="m" value="2400.0" valueMax="2450.0" valueMin="2350.0">2350­2450 m</quantity>
, continental divide (A.R.Gillogly),
<date id="9E1F1052C31DC743E2AE76C5FCB2FD26" box="[648,776,724,750]" pageId="51" pageNumber="52" value="1995-06-15">15.vi.1995</date>
; 2, Cerro Pando,
<quantity id="2D599B77C31DC743E40976C5FB2DFD26" box="[1071,1175,724,750]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="51" pageNumber="52" unit="m" value="1875.0">1875 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E48976C5FA91FD26" box="[1199,1323,724,750]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.911666">8°54'42&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E12E76EDFE23FCDE" box="[264,409,764,790]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.721664">82°43'18&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E18E76EDFDBDFCDE" box="[424,519,764,790]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="51" pageNumber="52" unit="m" value="1875.0">1875 m</quantity>
, flight intercept trap (J.S.Ashe, R.Brooks),
<date id="9E1F1052C31DC743E43176EDFB04FCDE" box="[1047,1214,764,790]" pageId="51" pageNumber="52" value="1996-06-17" valueMax="1996-06-18" valueMin="1996-06-17">17­18.vi.1996</date>
;, ditto but
<quantity id="2D599B77C31DC743E1187735FE18FCF6" box="[318,418,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="51" pageNumber="52" unit="m" value="1850.0">1850 m</quantity>
;
<specimenCount id="FCA7FD1BC31DC743E19E7735FDEEFCF6" box="[440,596,804,830]" pageId="51" pageNumber="52" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C31DC743E24E7735FD00FCF6" box="[616,698,804,830]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="51" pageNumber="52" unit="km" value="11.0">11 km</quantity>
NW Boquete, Volcán Barú,
<quantity id="2D599B77C31DC743E4107735FB21FCF6" box="[1078,1179,804,830]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E4967735FA91FCF6" box="[1200,1323,804,830]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.8">8°48'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E12E775DFE23FCAE" box="[264,409,844,870]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.48333">82°29'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E183775DFDBBFCAE" box="[421,513,844,870]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
, oak forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E3A8775DFBB6FCAE" box="[910,1036,844,870]" pageId="51" pageNumber="52" value="1995-06-18">18.vi.1995</date>
(all ­
<collectionCode id="8CB0AE57C31DC743E475775DFB10FCAE" box="[1107,1194,844,870]" pageId="51" pageNumber="52">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC31DC743E12E72FFFB34F883" httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="51" pageNumber="52" targetBox="[282,1304,939,1740]" targetPageId="51">
<paragraph id="EA1E3692C31DC743E12E72FFFB34F883" blockId="51.[264,1325,1774,1867]" pageId="51" pageNumber="52">
<emphasis id="D8D5EA80C31DC743E12E72FFFE5CF8CE" bold="true" box="[264,486,1774,1798]" pageId="51" pageNumber="52">FIGURES 190­193.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C31DC743E32F72FEFBC8F8CE" authority="Gusarov" authorityName="Gusarov" box="[777,1138,1775,1798]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31DC743E32F72FEFBB5F8CE" box="[777,1039,1775,1798]" italics="true" pageId="51" pageNumber="52">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31DC743E45B72FFFB00F8CE" box="[1149,1210,1774,1798]" pageId="51" pageNumber="52" rank="species">
<emphasis id="D8D5EA80C31DC743E45B72FFFB00F8CE" bold="true" box="[1149,1210,1774,1798]" pageId="51" pageNumber="52">sp. n.</emphasis>
</taxonomicNameLabel>
(190­191, holotype; 192­193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 male tergum 8; 191 male sternum 8; 192 female tergum 8; 193 female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C31AC744E11E750AFC04F98A" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="EA1E3692C31AC744E11E750AFDA5FE36" blockId="52.[264,1323,283,510]" pageId="52" pageNumber="53">
<emphasis id="D8D5EA80C31AC744E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C31AC744E1EC750DFD51FEFD" box="[458,747,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E1EC750DFD51FEFD" box="[458,747,284,309]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C31AC744E4DE750DFED7FE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C31AC744E4DE750DFED7FE95" italics="true" pageId="52" pageNumber="53">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body with brown elytra, legs and mouthparts; large eyes (temple length to eye length ratio 0.9­1.1); matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C31AC744E31F75ADFBE9FE1E" box="[825,1107,444,470]" captionStart-0="FIGURES 194 ­ 198" captionStart-1="FIGURES 199 ­ 207" captionStartId-0="52.[264,383,1328,1352]" captionStartId-1="53.[264,383,1638,1662]" captionTargetBox-0="[289,1303,584,1286]" captionTargetBox-1="[306,1280,457,1604]" captionTargetId-0="figure@52.[272,1315,569,1301]" captionTargetId-1="figure@53.[299,1288,449,1611]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-1="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156454/files/figure.png" httpUri-1="https://zenodo.org/record/156455/files/figure.png" pageId="52" pageNumber="53">Figs. 194­204, 206­207</figureCitation>
) and the shape of spermatheca (
<figureCitation id="729A2A17C31AC744E18D75F5FDA8FE36" box="[427,530,484,510]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="52" pageNumber="53">Fig. 205</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31AC744E12E7121FBB5FA67" httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53" targetBox="[289,1303,584,1286]" targetPageId="52">
<paragraph id="EA1E3692C31AC744E12E7121FBB5FA67" blockId="52.[264,1322,1328,1455]" pageId="52" pageNumber="53">
<emphasis id="D8D5EA80C31AC744E12E7121FE5BFA80" bold="true" box="[264,481,1328,1352]" pageId="52" pageNumber="53">FIGURES 194­198</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C31AC744E2AC7121FC4CFA8F" authority="Gusarov" authorityName="Gusarov" box="[650,1014,1328,1351]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31AC744E2AC7121FC2BFA8F" box="[650,913,1328,1351]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C31AC744E4247121FBFAFA80" bold="true" box="[1026,1088,1328,1352]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="C3E657FBC31AC744E4247121FBFAFA80" box="[1026,1088,1328,1352]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(194, 196­198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194­195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194­195, 197), 0.1 mm (196, 198).
</paragraph>
</caption>
<paragraph id="EA1E3692C31AC744E11E7211FC04F98A" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7211FDECF9D1" box="[312,598,1536,1561]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7211FDECF9D1" box="[312,598,1536,1561]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E2D57211FC17F9D1" box="[755,941,1536,1561]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C31AC744E2D57211FC17F9D1" box="[755,941,1536,1561]" italics="true" pageId="52" pageNumber="53">S. furcativentris</emphasis>
</taxonomicName>
in having parallel­sided apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C31AC744E2B17239FC84F98A" box="[663,830,1576,1602]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 194­196</figureCitation>
; 212­213).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31AC746E11E7241FDC4FB06" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="description">
<paragraph id="EA1E3692C31AC744E11E7241FC89F95A" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7241FDE0F9A1" box="[312,602,1616,1641]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7241FDE0F9A1" box="[312,602,1616,1641]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E3267241FC39F9A1" box="[768,899,1616,1641]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C31AC744E3267241FC39F9A1" box="[768,899,1616,1641]" italics="true" pageId="52" pageNumber="53">S. brunnea</emphasis>
</taxonomicName>
in having straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C31AC744E22B7269FD0EF95A" box="[525,692,1656,1682]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 197­198</figureCitation>
; 241­242).
</paragraph>
<paragraph id="EA1E3692C31AC744E11E72B1FCD1F8C2" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E72B1FDECF971" box="[312,598,1696,1721]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E72B1FDECF971" box="[312,598,1696,1721]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E2D672B1FC2CF971" box="[752,918,1696,1721]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C31AC744E2D672B1FC2CF971" box="[752,918,1696,1721]" italics="true" pageId="52" pageNumber="53">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; apex of median lobe parallelsided (in parameral view) (
<figureCitation id="729A2A17C31AC744E26372E1FD56F8C2" box="[581,748,1776,1802]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 194­196</figureCitation>
; 226­227).
</paragraph>
<paragraph id="EA1E3692C31AC745E11E7309FB51FE4E" blockId="52.[264,1324,1536,1882]" lastBlockId="53.[264,1324,284,390]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7309FDE6F8F9" box="[312,604,1816,1841]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7309FDE6F8F9" box="[312,604,1816,1841]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E32D7309FC09F8F9" box="[779,947,1816,1841]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C31AC744E32D7309FC09F8F9" box="[779,947,1816,1841]" italics="true" pageId="52" pageNumber="53">S. curtipennis</emphasis>
</taxonomicName>
in having larger eyes; longer elytra; wings longer than elytra; posterior margin of male tergum 8 with medial emargination (
<figureCitation id="729A2A17C31BC745E175750DFE00FEFE" box="[339,442,284,310]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 male tergum 8; 191 male sternum 8; 192 female tergum 8; 193 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="53" pageNumber="54">Fig. 190</figureCitation>
); narrow apex of the median lobe (in parameral view) (
<figureCitation id="729A2A17C31BC745E464750DFB53FEFE" box="[1090,1257,284,310]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="53" pageNumber="54">Figs. 194­196</figureCitation>
; 251­ 252); short medial lamellae of the internal sac (
<figureCitation id="729A2A17C31BC745E3687555FC79FE96" box="[846,963,324,350]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 201</figureCitation>
; 258); lateral diverticula with sclerotized denticles (
<figureCitation id="729A2A17C31BC745E220757DFD09FE4E" box="[518,691,364,390]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 200, 207</figureCitation>
; 259); and shorter spermatheca (
<figureCitation id="729A2A17C31BC745E417757DFB18FE4E" box="[1073,1186,364,390]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 205</figureCitation>
,
<figureCitation id="729A2A17C31BC745E488757DFB65FE4E" box="[1198,1247,364,390]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 median lobe, parameral view; 252 apex of median lobe, parameral view; 253 median lobe, lateral view; 254 apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="53" pageNumber="54">255</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31BC745E12E7277FB09F885" httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54" targetBox="[306,1280,457,1604]" targetPageId="53">
<paragraph id="EA1E3692C31BC745E12E7277FB09F885" blockId="53.[264,1323,1638,1904]" pageId="53" pageNumber="54">
<emphasis id="D8D5EA80C31BC745E12E7277FE58F9B6" bold="true" box="[264,482,1638,1662]" pageId="53" pageNumber="54">FIGURES 199­207</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C31BC745E2AE7277FC4EF9B5" authority="Gusarov" authorityName="Gusarov" box="[648,1012,1638,1661]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31BC745E2AE7277FC35F9B5" box="[648,911,1638,1661]" italics="true" pageId="53" pageNumber="54">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C31BC745E4247277FBF8F9B6" bold="true" box="[1026,1090,1638,1662]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="C3E657FBC31BC745E4247277FBF8F9B6" box="[1026,1090,1638,1662]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(199, 204, holotype; 200­203, 205­207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199­200, 203­204), 0.1 mm (201­202, 205­207).
</paragraph>
</caption>
<paragraph id="EA1E3692C31BC745E12E7348FC0CF8B8" blockId="53.[264,1323,1638,1904]" box="[264,950,1881,1904]" pageId="53" pageNumber="54">CP copulatory piece; LD lateral diverticulum of internal sac.</paragraph>
<paragraph id="EA1E3692C318C746E11E750AFE2EFE96" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="54" pageNumber="55">Description.</emphasis>
Length
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. Body dark brown, with brown elytra, legs and mouthparts.
</paragraph>
<paragraph id="EA1E3692C318C746E11E757DFCE9FE36" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">Head surface matte, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2 times their diameter. Temple length to eye length ratio 0.9­1.1. Antennal article 2 longer than article 3, article 4 subquadrate, 5 slightly transverse, 6­10 transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C318C746E11E761DFBE5FCDE" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C318C746E40D761DFB6CFDEE" box="[1067,1238,524,550]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.4" metricValueMin="4.9" pageId="54" pageNumber="55" unit="mm" value="0.515" valueMax="0.54" valueMin="0.49">0.49­0.54 mm</quantity>
, length
<quantity id="2D599B77C318C746E12E7625FE0BFD86" box="[264,433,564,590]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="54" pageNumber="55" unit="mm" value="0.435" valueMax="0.46" valueMin="0.41">0.41­0.46 mm</quantity>
, width to length ratio 1.2, surface matte, with fine isodiametric microsculpture; punctation fine and weak, distance between punctures equal to their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times as wide as long, glossy, with fine and weak isodiametric microsculpture; with fine punctation, distance between punctures equal to ½­1 time their diameter. Wings fully developed or partially reduced, longer than elytra.
</paragraph>
<paragraph id="EA1E3692C318C746E11E7735FBBDFC7E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Abdominal terga glossy, with fine microsculpture consisting of strongly transverse meshes, with fine punctation, distance between punctures equals 2­4 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C318C746E36B778DFC40FC7E" box="[845,1018,924,950]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 male tergum 8; 191 male sternum 8; 192 female tergum 8; 193 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="54" pageNumber="55">Figs. 190, 192</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E77D5FC99FB9E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
In males posterior angles of tergum 3 projecting as spines up to 3 times as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to 2/ 3 as long as tergum length; posterior margin of tergum 8 with medial emargination delimited by short dentiform projections (
<figureCitation id="729A2A17C318C746E289702DFCACFB9E" box="[687,790,1084,1110]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 male tergum 8; 191 male sternum 8; 192 female tergum 8; 193 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="54" pageNumber="55">Fig. 190</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E7075FC19FB6E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Aedeagus as in
<figureCitation id="729A2A17C318C746E2247075FC9FFBB6" box="[514,805,1124,1150]" captionStart-0="FIGURES 194 ­ 198" captionStart-1="FIGURES 199 ­ 207" captionStartId-0="52.[264,383,1328,1352]" captionStartId-1="53.[264,383,1638,1662]" captionTargetBox-0="[289,1303,584,1286]" captionTargetBox-1="[306,1280,457,1604]" captionTargetId-0="figure@52.[272,1315,569,1301]" captionTargetId-1="figure@53.[299,1288,449,1611]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 median lobe, parameral view; 196 apex of median lobe, parameral view; 197 median lobe, lateral view; 198 apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-1="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156454/files/figure.png" httpUri-1="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Figs. 194­204, 206­207</figureCitation>
. Apex of paramere broad, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C318C746E309709DFC2CFB6E" box="[815,918,1164,1190]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Fig. 202</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E70A5FDC4FB06" blockId="54.[264,1324,283,1350]" box="[312,638,1204,1230]" pageId="54" pageNumber="55">
Spermatheca as in
<figureCitation id="729A2A17C318C746E23370A5FDC1FB06" box="[533,635,1204,1230]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 details of internal sac retracted into median lobe, abparameral view; 200 everted internal sac, parameral view; 201 medial lamellae, parameral view; 202 apex of left paramere; 203 everted internal sac, lateral view; 204 details of internal sac retracted into median lobe, lateral view; 205 spermatheca; 206 right distal sclerite of internal sac; 207 left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Fig. 205</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C318C746E11E70CAFC21FA8E" pageId="54" pageNumber="55" type="distribution">
<paragraph id="EA1E3692C318C746E11E70CAFBA5FB3E" blockId="54.[264,1324,283,1350]" box="[312,1055,1243,1270]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E70CAFE62FB3D" bold="true" box="[312,472,1243,1269]" pageId="54" pageNumber="55">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C318C746E2A370CDFCBFFB3E" box="[645,773,1244,1270]" name="Costa Rica" pageId="54" pageNumber="55">Costa Rica</collectingCountry>
and
<collectingCountry id="92B67602C318C746E31870CDFC21FB3E" box="[830,923,1244,1270]" name="Panama" pageId="54" pageNumber="55">Panama</collectingCountry>
(
<figureCitation id="729A2A17C318C746E38D70CDFBA9FB3E" box="[939,1043,1244,1270]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="54" pageNumber="55">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E7112FC21FA8E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E7112FDBCFAD5" bold="true" box="[312,518,1283,1309]" pageId="54" pageNumber="55">Natural History.</emphasis>
<taxonomicName id="2DA14D11C318C746E2317115FC8DFAD5" box="[535,823,1284,1309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C318C746E2317115FC8DFAD5" box="[535,823,1284,1309]" italics="true" pageId="54" pageNumber="55">Seeversiella scabricollis</emphasis>
</taxonomicName>
was collected in leaf litter and with flight intercept and malaise traps, at altitudes of
<quantity id="2D599B77C318C746E2D1713DFC2DFA8E" box="[759,919,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="3.3" metricValueMin="1.45" pageId="54" pageNumber="55" unit="m" value="2375.0" valueMax="3300.0" valueMin="1450.0">1450­3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C195F2F9F8657ED8F0160998A3987E3D">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C327C779E12E750AFCF3FEFE" box="[264,841,283,310]" pageId="9" pageNumber="10" type="nomenclature">
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<emphasis id="D8D5EA80C327C779E12E750AFCF3FEFE" bold="true" box="[264,841,283,310]" pageId="9" pageNumber="10">
Key for identification of species of
<taxonomicName id="2DA14D11C327C779E29F750DFCF3FEFE" ID-CoL="8VS3L" box="[697,841,284,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C327C779E29F750DFCF3FEFE" bold="true" box="[697,841,284,310]" italics="true" pageId="9" pageNumber="10">Seeversiella</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<keyStep id="51552E37C327C779E12E7576FA91FE31" pageId="9" pageNumber="10">
<subSubSection id="A2BB6519C327C779E12E7576FA96FE61" pageId="9" pageNumber="10" type="key">
<paragraph id="EA1E3692C327C779E12E7576FA96FE61" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7576FA96FE61" pageId="9" pageNumber="10">1 Elytra longer or only a little shorter than pronotum. Pronotal length to elytral length (measured from humeral angle) ratio 1.1 or less ......................................................... 2</keyLead>
</paragraph>
</subSubSection>
<paragraph id="EA1E3692C327C779E12E75A6FA91FE31" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E75A6FA91FE31" pageId="9" pageNumber="10">
Elytra much shorter than pronotum (
<figureCitation id="729A2A17C327C779E2D075A6FC3EFE19" box="[758,900,439,465]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="9" pageNumber="10">Figs. 18­19</figureCitation>
). Pronotal length to elytral length (measured from humeral angle) ratio more than 1.1 ................................................. 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7616FA91FD81" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7616FA96FDE9" blockId="9.[264,1327,359,1904]" box="[264,1324,519,545]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7616FA96FDE9" box="[264,1324,519,545]" pageId="9" pageNumber="10">2 Pronotum glossy, with weak isodiametric microsculpture or without microsculpture 3</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E763EFA91FD81" blockId="9.[264,1327,359,1904]" box="[264,1323,559,585]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E763EFA91FD81" box="[264,1323,559,585]" pageId="9" pageNumber="10"> Pronotum matte, with strong isodiametric microsculpture ....................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7646FA91FC41" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7646FA91FCD8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7646FA91FCD8" pageId="9" pageNumber="10">
3 Punctation of pronotum asperate. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C327C779E4907646FED3FD51" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 male tergum 8; 223 male sternum 8; 224 female tergum 8; 225 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="9" pageNumber="10">Figs. 222, 224</figureCitation>
). Proximal seta of the apex of paramere is approximately as long as the other three setae (
<figureCitation id="729A2A17C327C779E1EA76B6FD8DFD09" box="[460,567,679,705]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Fig. 233</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C327C779E2CB76B6FC2DFD09" box="[749,919,679,705]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 median lobe, parameral view; 227 apex of median lobe, parameral view; 228 median lobe, lateral view; 229 apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Figs. 226­233</figureCitation>
. Spermatheca:
<figureCitation id="729A2A17C327C779E44E76B6FB68FD09" box="[1128,1234,679,705]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 details of internal sac retracted into median lobe, lateral view; 232 details of internal sac retracted into median lobe, abparameral view; 233 apex of left paramere; 234 spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Fig. 234</figureCitation>
. Body length 2.6­3.0 mm, pronotal width
<quantity id="2D599B77C327C779E2F576DEFCC7FD21" box="[723,893,719,745]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="9" pageNumber="10" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53­0.56 mm</quantity>
. Known from Cerro Punta,
<collectingCountry id="92B67602C327C779E4EF76DEFA93FD21" box="[1225,1321,719,745]" name="Panama" pageId="9" pageNumber="10">Panama</collectingCountry>
, at altitude of
<quantity id="2D599B77C327C779E1F576E6FD96FCD9" box="[467,556,759,785]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="10" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C327C779E21A76E6FD1AFCD9" box="[572,672,759,785]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="9" pageNumber="10">Fig. 388</figureCitation>
) .............................. 16.
<taxonomicName id="2DA14D11C327C779E3EA76E6FB67FCD9" authority="Gusarov" authorityName="Gusarov" box="[972,1245,759,785]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C327C779E3EA76E6FBC8FCD8" box="[972,1138,759,784]" italics="true" pageId="9" pageNumber="10">S. brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C327C779E4CF76E7FA91FCD8" bold="true" box="[1257,1323,758,784]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="C3E657FBC327C779E4CF76E7FA91FCD8" box="[1257,1323,758,784]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E770EFA91FC41" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E770EFA91FC41" pageId="9" pageNumber="10">
Punctation of pronotum not asperate. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C327C779E4D7770EFE32FCA9" captionStart="FIGURES 20 ­ 23" captionStartId="15.[264,383,1258,1282]" captionTargetBox="[344,1240,531,1222]" captionTargetId="figure@15.[337,1250,524,1230]" captionTargetPageId="15" captionText="FIGURES 20 ­ 23. Abdominal segment 8 of Seeversiella globicollis (Bernhauer) (20 ­ 21, Cloudcroft, New Mexico; 22 ­ 23, Salmo, British Columbia). 20 male tergum 8; 21 male sternum 8; 22 female tergum 8; 23 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156425/files/figure.png" pageId="9" pageNumber="10">Figs. 20, 22</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C327C779E1A4777EFE6DFC41" box="[386,471,879,905]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Fig. 33</figureCitation>
)............................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7786FA96FC11" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7786FA96FC79" blockId="9.[264,1327,359,1904]" box="[264,1324,919,945]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7786FA96FC79" box="[264,1324,919,945]" pageId="9" pageNumber="10">4 Pronotum with weak isodiametric microsculpture ...................................................... 5</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E77AEFA96FC11" blockId="9.[264,1327,359,1904]" box="[264,1324,959,985]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E77AEFA96FC11" box="[264,1324,959,985]" pageId="9" pageNumber="10"> Pronotum without microsculpture ............................................................................. 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E77F6FA96FBE1" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E77F6FA91FBC9" blockId="9.[264,1327,359,1904]" box="[264,1323,999,1025]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E77F6FA91FBC9" box="[264,1323,999,1025]" pageId="9" pageNumber="10">
<specimenCount id="FCA7FD1BC327C779E12E77F6FEC5FBC9" box="[264,383,999,1025]" pageId="9" pageNumber="10" type="male">5 Males</specimenCount>
............................................................................................................................ 6
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E701EFA96FBE1" blockId="9.[264,1327,359,1904]" box="[264,1324,1039,1065]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E701EFA96FBE1" box="[264,1324,1039,1065]" pageId="9" pageNumber="10">
Females (Females of
<taxonomicName id="2DA14D11C327C779E215701EFD07FBE0" box="[563,701,1039,1064]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C327C779E215701EFD07FBE0" box="[563,701,1039,1064]" italics="true" pageId="9" pageNumber="10">S. nigriceps</emphasis>
</taxonomicName>
are unknown) ........................................................ 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7027FA94FB00" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7027FA96FBB0" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7027FA96FBB0" pageId="9" pageNumber="10">
6 Distal sclerites of internal sac strongly sclerotized, hook­shaped (
<figureCitation id="729A2A17C327C779E41D7027FB71FB98" box="[1083,1227,1078,1104]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 31, 35</figureCitation>
; 48, 51; DS) or dentiform (
<figureCitation id="729A2A17C327C779E234704FFD1BFBB0" box="[530,673,1118,1144]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
; DS) 7
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E7097FA94FB00" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7097FA94FB00" pageId="9" pageNumber="10">
Distal sclerites of internal sac poorly sclerotized, not hook­shaped or dentiform and may have numerous spicules (
<figureCitation id="729A2A17C327C779E2BC70BFFD47FB00" box="[666,765,1198,1224]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="9" pageNumber="10">Figs. 87</figureCitation>
; 99, 101; 124; 146; DS) ............................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E70C7FA95FAD0" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E70C7FA95FB38" blockId="9.[264,1327,359,1904]" box="[264,1327,1238,1264]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E70C7FA95FB38" box="[264,1327,1238,1264]" pageId="9" pageNumber="10">
7 Distal sclerites of internal sac hook­shaped (
<figureCitation id="729A2A17C327C779E36670C7FC75FB38" box="[832,975,1238,1264]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 31, 35</figureCitation>
; 48, 51) ............................. 8
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E70EFFA95FAD0" blockId="9.[264,1327,359,1904]" box="[264,1327,1278,1304]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E70EFFA95FAD0" box="[264,1327,1278,1304]" pageId="9" pageNumber="10">
Distal sclerites of internal sac dentiform (
<figureCitation id="729A2A17C327C779E33870EFFC16FAD0" box="[798,940,1278,1304]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
; 124) ...................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7137FA90F8E8" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7137FA91F9F8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7137FA91F9F8" pageId="9" pageNumber="10">
8 Body dark brown. Microsculpture on head and pronotum less distinct. Apex of median lobe in lateral view straight (
<figureCitation id="729A2A17C327C779E2DF715FFC3DFAA0" box="[761,903,1358,1384]" captionStart="FIGURES 24 ­ 27" captionStartId="18.[264,383,1768,1792]" captionTargetBox="[301,1288,1258,1750]" captionTargetId="figure@18.[287,1300,1249,1757]" captionTargetPageId="18" captionText="FIGURES 24 ­ 27. Aedeagus of Seeversiella globicollis (Bernhauer) (Cypress Hills Provincial Park, Alberta). 24 median lobe, parameral view; 25 apex of median lobe, parameral view; 26 median lobe, lateral view; 27 apex of median lobe, lateral view. Scale bar 0.2 mm (24, 26), 0.1 mm (25, 27)." httpUri="https://zenodo.org/record/156426/files/figure.png" pageId="9" pageNumber="10">Figs. 26­27</figureCitation>
). Copulatory piece of internal sac long (
<figureCitation id="729A2A17C327C779E1A77167FDA3FA58" box="[385,537,1398,1424]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 29, 34</figureCitation>
). Elytra (measured from humeral angle) longer than pronotum. Wings fully developed, 4 times as long as elytra. Aedeagus:
<figureCitation id="729A2A17C327C779E464718FFA9DFA70" box="[1090,1319,1438,1464]" captionStart-0="FIGURES 24 ­ 27" captionStart-1="FIGURES 28 ­ 36" captionStartId-0="18.[264,383,1768,1792]" captionStartId-1="19.[264,383,1598,1622]" captionTargetBox-0="[301,1288,1258,1750]" captionTargetBox-1="[297,1289,603,1577]" captionTargetId-0="figure@18.[287,1300,1249,1757]" captionTargetId-1="figure@19.[285,1302,592,1587]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 24 ­ 27. Aedeagus of Seeversiella globicollis (Bernhauer) (Cypress Hills Provincial Park, Alberta). 24 median lobe, parameral view; 25 apex of median lobe, parameral view; 26 median lobe, lateral view; 27 apex of median lobe, lateral view. Scale bar 0.2 mm (24, 26), 0.1 mm (25, 27)." captionText-1="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri-0="https://zenodo.org/record/156426/files/figure.png" httpUri-1="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 24­31, 33­36</figureCitation>
. Body length
<quantity id="2D599B77C327C779E1F471D7FDE5FA28" box="[466,607,1478,1504]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.3" metricValueMin="2.4" pageId="9" pageNumber="10" unit="mm" value="2.8499999999999996" valueMax="3.3" valueMin="2.4">2.4­3.3 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E30771D7FC76FA28" box="[801,972,1478,1504]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.4" metricValueMin="4.4" pageId="9" pageNumber="10" unit="mm" value="0.49" valueMax="0.54" valueMin="0.44">0.44­0.54 mm</quantity>
. Widespread in North
<collectingCountry id="92B67602C327C779E4C671D7FEE7F9C0" name="United States of America" pageId="9" pageNumber="10">America</collectingCountry>
, known from
<collectingCountry id="92B67602C327C779E22671FFFDE2F9C0" box="[512,600,1518,1544]" name="Canada" pageId="9" pageNumber="10">Canada</collectingCountry>
to
<collectingCountry id="92B67602C327C779E25B71FFFD55F9C0" box="[637,751,1518,1544]" name="Honduras" pageId="9" pageNumber="10">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C327C779E2D971FFFC1EF9C0" box="[767,932,1518,1544]" captionStart-0="FIGURE 385" captionStart-1="FIGURE 386" captionStart-2="FIGURE 387" captionStartId-0="95.[264,368,1458,1482]" captionStartId-1="96.[264,368,1354,1378]" captionStartId-2="97.[264,368,1861,1885]" captionTargetBox-0="[309,1276,654,1420]" captionTargetBox-1="[315,1274,412,1319]" captionTargetBox-2="[313,1275,771,1826]" captionTargetId-0="figure@95.[301,1286,649,1431]" captionTargetId-1="figure@96.[309,1279,409,1327]" captionTargetId-2="figure@97.[304,1282,766,1834]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionTargetPageId-2="97" captionText-0="FIGURE 385. Geographical distribution of Seeversiella globicollis (Bernhauer)." captionText-1="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." captionText-2="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri-0="https://zenodo.org/record/156487/files/figure.png" httpUri-1="https://zenodo.org/record/156488/files/figure.png" httpUri-2="https://zenodo.org/record/156489/files/figure.png" pageId="9" pageNumber="10">Figs. 385­387</figureCitation>
) .................................................. ...............................................................................1.
<taxonomicName id="2DA14D11C327C779E3B87206FB76F9F8" authority="Bernhauer" authorityName="Bernhauer" box="[926,1228,1558,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C327C779E3B87206FB8DF9F8" box="[926,1079,1559,1584]" italics="true" pageId="9" pageNumber="10">S. globicollis</emphasis>
(Bernhauer)
</taxonomicName>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E722FFA90F8E8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E722FFA90F8E8" pageId="9" pageNumber="10">
Body brown. Microsculpture on head and pronotum more distinct. Apex of median lobe in lateral view bent paramerally (
<figureCitation id="729A2A17C327C779E3277277FC35F948" box="[769,911,1638,1664]" captionStart="FIGURES 41 ­ 44" captionStartId="22.[264,383,1298,1322]" captionTargetBox="[293,1296,576,1260]" captionTargetId="figure@22.[281,1306,569,1271]" captionTargetPageId="22" captionText="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="9" pageNumber="10">Figs. 43­44</figureCitation>
). Copulatory piece of internal sac short (
<figureCitation id="729A2A17C327C779E1A5729FFD80F960" box="[387,570,1678,1704]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="9" pageNumber="10">Figs. 46, 49­50</figureCitation>
). Elytra (measured from humeral angle) shorter than pronotum. Wings short, 2 times as long as elytra. Aedeagus:
<figureCitation id="729A2A17C327C779E3B872A7FB93F918" box="[926,1065,1718,1744]" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="9" pageNumber="10">Figs. 41­52</figureCitation>
. Body length
<quantity id="2D599B77C327C779E4FE72A7FED3F930" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="9" pageNumber="10" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E20F72CFFD69F930" box="[553,723,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="9" pageNumber="10" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44­0.49 mm</quantity>
. Known from the
<collectingCountry id="92B67602C327C779E39772CFFB8AF930" box="[945,1072,1758,1784]" name="Guadeloupe" pageId="9" pageNumber="10">Guadalupe</collectingCountry>
Mountains, Texas, at altitude of
<quantity id="2D599B77C327C779E1917317FDAAF8E8" box="[439,528,1798,1824]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="10" unit="m" value="2400.0">2400 m</quantity>
(
<figureCitation id="729A2A17C327C779E2397317FD39F8E8" box="[543,643,1798,1824]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="9" pageNumber="10">Fig. 386</figureCitation>
) ............................... 2.
<taxonomicName id="2DA14D11C327C779E3817316FBC7F8E8" authority="Gusarov" authorityName="Gusarov" box="[935,1149,1798,1824]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C327C779E3817316FBA8F8E8" box="[935,1042,1799,1824]" italics="true" pageId="9" pageNumber="10">S. texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C327C779E4AF7317FB76F8E8" bold="true" box="[1161,1228,1798,1824]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="C3E657FBC327C779E4AF7317FB76F8E8" box="[1161,1228,1798,1824]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C77AE12E733FFA91FE1E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E733FFA97F8B8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E733FFA97F8B8" pageId="9" pageNumber="10">
9 Distal sclerites of internal sac wider (
<figureCitation id="729A2A17C327C779E2D8733FFC29F880" box="[766,915,1838,1864]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C327C779E463733FFA92F880" box="[1093,1320,1838,1864]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 58­61, 63­67</figureCitation>
. Body length
<quantity id="2D599B77C327C779E1F27347FDDBF8B8" box="[468,609,1878,1904]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="9" pageNumber="10" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E3007347FC68F8B8" box="[806,978,1878,1904]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="9" pageNumber="10" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
. Known from the Huachuca
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE11E750DFA91FE1E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
Mountains, Arizona, at altitude of
<quantity id="2D599B77C324C77AE2EA750DFC9FFEFE" box="[716,805,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="11" unit="m" value="2150.0">2150 m</quantity>
(
<figureCitation id="729A2A17C324C77AE313750DFC22FEFE" box="[821,920,284,310]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="10" pageNumber="11">Fig. 386</figureCitation>
) ................................................... ........................................................ 3.
<taxonomicName id="2DA14D11C324C77AE2DF7555FB9EFE96" authority="Gusarov" authorityName="Gusarov" box="[761,1060,324,350]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE2DF7555FC02FE95" box="[761,952,324,349]" italics="true" pageId="10" pageNumber="11">S. sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4097552FBCBFE95" bold="true" box="[1071,1137,323,349]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4097552FBCBFE95" box="[1071,1137,323,349]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males, in part)
<keyLead id="515095A7C324C77AE12E757DFA91FE1E" pageId="10" pageNumber="11">
Distal sclerites of internal sac narrower (
<figureCitation id="729A2A17C324C77AE33A757DFC3EFE4E" box="[796,900,364,390]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="10" pageNumber="11">Fig. 124</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE416757DFB6DFE4E" box="[1072,1239,364,390]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 male tergum 8; 118 male sternum 8; 119 median lobe, parameral view; 120 apex of median lobe, parameral view; 121 median lobe, lateral view; 122 apex of median lobe, lateral view; 123 ­ 124 details of internal sac retracted into median lobe, lateral view; 125 apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="10" pageNumber="11">Figs. 119­125</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE1AF7585FE52FE66" box="[393,488,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE28F7585FCADFE66" box="[681,791,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="10" pageNumber="11" unit="mm" value="0.47">0.47 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE3EC7585FB9DFE66" box="[970,1063,404,430]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE4F47585FA96FE66" box="[1234,1324,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="m" value="2400.0">2400 m</quantity>
(
<figureCitation id="729A2A17C324C77AE16775ADFE1FFE1E" box="[321,421,444,470]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
)...............................................8.
<taxonomicName id="2DA14D11C324C77AE30875ADFB99FE1E" authority="Gusarov" authorityName="Gusarov" box="[814,1059,444,470]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE30875ADFC02FE1D" box="[814,952,444,469]" italics="true" pageId="10" pageNumber="11">S. nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE40975AAFBCBFE1D" bold="true" box="[1071,1137,443,469]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE40975AAFBCBFE1D" box="[1071,1137,443,469]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males, in part)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E75F5FA97FDEE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E75F5FA91FE36" blockId="10.[264,1326,284,1870]" box="[264,1323,484,510]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E75F5FA91FE36" box="[264,1323,484,510]" pageId="10" pageNumber="11">
10 Apex of median lobe in lateral view widened subapically (
<figureCitation id="729A2A17C324C77AE3C375F5FB31FE36" box="[997,1163,484,510]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 112­113</figureCitation>
; 156­157) 11
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E761DFA97FDEE" blockId="10.[264,1326,284,1870]" box="[264,1325,524,550]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E761DFA97FDEE" box="[264,1325,524,550]" pageId="10" pageNumber="11">
Apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C324C77AE435761DFB20FDEE" box="[1043,1178,524,550]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Figs. 85­86</figureCitation>
) ............. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7625FA96FCDE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7625FA90FD0E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7625FA90FD0E" pageId="10" pageNumber="11">
11 Apex of median lobe in lateral view emarginate on parameral side (
<figureCitation id="729A2A17C324C77AE4527625FA9AFD86" box="[1140,1312,564,590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 156­157</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE1F6764DFDC7FDBE" box="[464,637,604,630]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 154­159</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE364764DFC1FFDBE" box="[834,933,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE45E764DFA92FDBE" box="[1144,1320,604,630]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="10" pageNumber="11" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE1F07695FDDBFD56" box="[470,609,644,670]" name="El Salvador" pageId="10" pageNumber="11">El Salvador</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE32E7695FCDBFD56" box="[776,865,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="11" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C324C77AE3577695FC6EFD56" box="[881,980,644,670]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ........................................... ................................................................................11.
<taxonomicName id="2DA14D11C324C77AE39476BDFBC7FD0E" authority="Gusarov" authorityName="Gusarov" box="[946,1149,684,710]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE39476BDFBA8FD0D" box="[946,1042,684,709]" italics="true" pageId="10" pageNumber="11">S. badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF76BAFB76FD0D" bold="true" box="[1161,1228,683,709]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF76BAFB76FD0D" box="[1161,1228,683,709]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E76C5FA96FCDE" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E76C5FA96FCDE" pageId="10" pageNumber="11">
Apex of median lobe in lateral view may be slightly bent paramerally (
<figureCitation id="729A2A17C324C77AE45B76C5FA99FD26" box="[1149,1315,724,750]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 141­142</figureCitation>
) but never clearly emarginate (
<figureCitation id="729A2A17C324C77AE2B076EDFC81FCDE" box="[662,827,764,790]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 112­113</figureCitation>
; 141­142) ............................................ 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7735FA97FC7E" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7735FA90FC46" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7735FA90FC46" pageId="10" pageNumber="11">
12 Apex of median lobe in parameral view wide (
<figureCitation id="729A2A17C324C77AE3447735FBB3FCF6" box="[866,1033,804,830]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 110­111</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4947735FEDCFCAE" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 apex of left paramere; 110 median lobe, parameral view; 111 apex of median lobe, parameral view; 112 median lobe, lateral view; 113 apex of median lobe, lateral view; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 110­ 115</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE234775DFDD5FCAE" box="[530,623,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE308775DFC21FCAE" box="[814,923,844,870]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="11" unit="mm" value="0.49">0.49 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE46A775DFB13FCAE" box="[1100,1193,844,870]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE17E7765FE0BFC46" box="[344,433,884,910]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="m" value="1300.0">1300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE1E67765FD9EFC46" box="[448,548,884,910]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................... 7.
<taxonomicName id="2DA14D11C324C77AE3637765FBC4FC46" authority="Gusarov" authorityName="Gusarov" box="[837,1150,884,910]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3637765FBA8FC45" box="[837,1042,884,909]" italics="true" pageId="10" pageNumber="11">S. tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF7762FB76FC45" bold="true" box="[1161,1228,883,909]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF7762FB76FC45" box="[1161,1228,883,909]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E778DFA97FC7E" blockId="10.[264,1326,284,1870]" box="[264,1325,924,950]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E778DFA97FC7E" box="[264,1325,924,950]" pageId="10" pageNumber="11">
Apex of median lobe in parameral view narrow (
<figureCitation id="729A2A17C324C77AE353778DFBA6FC7E" box="[885,1052,924,950]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 139­140</figureCitation>
; 165­166) .............. 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E77D5FA90FB3E" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E77D5FA90FB9E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E77D5FA90FB9E" pageId="10" pageNumber="11">
13 Apex of median lobe in parameral view with weak basal constriction (
<figureCitation id="729A2A17C324C77AE45077D5FAA5FC16" box="[1142,1311,964,990]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 139­140</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE1EE77FDFDCBFBCE" box="[456,625,1004,1030]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 median lobe, parameral view; 140 apex of median lobe, parameral view; 141 median lobe, lateral view; 142 apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Figs. 139­148</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE30177FDFC0FFBCE" box="[807,949,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="10" pageNumber="11" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE45A77FDFA92FBCE" box="[1148,1320,1004,1030]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="10" pageNumber="11" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE1F07005FD88FBE6" box="[470,562,1044,1070]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitudes of
<quantity id="2D599B77C324C77AE2C27005FC3BFBE6" box="[740,897,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="2000.0" valueMax="2300.0" valueMin="1700.0">1700­2300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE3B77005FC4FFBE6" box="[913,1013,1044,1070]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ....................................... ........................................................................ 10.
<taxonomicName id="2DA14D11C324C77AE3A3702DFBC4FB9E" authority="Gusarov" authorityName="Gusarov" box="[901,1150,1084,1110]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3A3702DFBA8FB9D" box="[901,1042,1084,1109]" italics="true" pageId="10" pageNumber="11">S. mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF702AFB76FB9D" bold="true" box="[1161,1228,1083,1109]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF702AFB76FB9D" box="[1161,1228,1083,1109]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E7075FA90FB3E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7075FA90FB3E" pageId="10" pageNumber="11">
Apex of median lobe in parameral view gradually widened basally, not constricted (
<figureCitation id="729A2A17C324C77AE167709DFE50FB6E" box="[321,490,1164,1190]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="10" pageNumber="11">Figs. 165­166</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE2B5709DFC80FB6E" box="[659,826,1164,1190]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 median lobe, parameral view; 166 apex of median lobe, parameral view; 167 median lobe, lateral view; 168 apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="10" pageNumber="11">Figs. 165­173</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE3CE709DFBCEFB6E" box="[1000,1140,1164,1190]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE11E70A5FE58FB06" box="[312,482,1204,1230]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="10" pageNumber="11" unit="mm" value="0.51" valueMax="0.54" valueMin="0.48">0.48­0.54 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE2B470A5FCBDFB06" box="[658,775,1204,1230]" name="Honduras" pageId="10" pageNumber="11">Honduras</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE38970A5FBF6FB06" box="[943,1100,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="10" pageNumber="11" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000­2100 m</quantity>
(
<figureCitation id="729A2A17C324C77AE47A70A5FB7BFB06" box="[1116,1217,1204,1230]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............ ............................................................................. 12.
<taxonomicName id="2DA14D11C324C77AE38E70CDFBC4FB3E" authority="Gusarov" authorityName="Gusarov" box="[936,1150,1244,1270]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE38E70CDFBA8FB3D" box="[936,1042,1244,1269]" italics="true" pageId="10" pageNumber="11">S. similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF70CAFB76FB3D" bold="true" box="[1161,1228,1243,1269]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF70CAFB76FB3D" box="[1161,1228,1243,1269]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7115FA91F9C6" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7115FA90FAA6" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7115FA90FAA6" pageId="10" pageNumber="11">
14 Distal sclerites of internal sac have weak spicules (
<figureCitation id="729A2A17C324C77AE3B17115FC4BFAD6" box="[919,1009,1284,1310]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Fig. 87</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4BB7115FA9DFAD6" box="[1181,1319,1284,1310]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Figs. 83­88</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE1F7713DFDE6FA8E" box="[465,604,1324,1350]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3­2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE33A713DFC7CFA8E" box="[796,966,1324,1350]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="10" pageNumber="11" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE45E713DFB6FFA8E" box="[1144,1237,1324,1350]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE1B77145FD94FAA6" box="[401,558,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700­1900 m</quantity>
(
<figureCitation id="729A2A17C324C77AE2187145FD18FAA6" box="[574,674,1364,1390]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................. 5.
<taxonomicName id="2DA14D11C324C77AE3917145FBC4FAA6" authority="Gusarov" authorityName="Gusarov" box="[951,1150,1364,1390]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3917145FBA8FAA5" box="[951,1042,1364,1389]" italics="true" pageId="10" pageNumber="11">S. fusca</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC324C77AE4AF7142FB76FAA5" box="[1161,1228,1363,1389]" pageId="10" pageNumber="11" rank="species">
<emphasis id="D8D5EA80C324C77AE4AF7142FB76FAA5" bold="true" box="[1161,1228,1363,1389]" pageId="10" pageNumber="11">sp. n.</emphasis>
</taxonomicNameLabel>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E716DFA91F9C6" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E716DFA91F9C6" pageId="10" pageNumber="11">
Distal sclerites of internal sac have strong spicules (
<figureCitation id="729A2A17C324C77AE385716DFBFFFA5E" box="[931,1093,1404,1430]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">Figs. 99, 101</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4D7716DFE35FA76" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">Figs. 94­103</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE21871B5FD71FA76" box="[574,715,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE3AB71B5FB83FA76" box="[909,1081,1444,1470]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="10" pageNumber="11" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE4C871B5FEE6FA2E" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
and
<collectingCountry id="92B67602C324C77AE1B271DDFDB3FA2E" box="[404,521,1484,1510]" name="Honduras" pageId="10" pageNumber="11">Honduras</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE29671DDFCF7FA2E" box="[688,845,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="10" pageNumber="11" unit="m" value="1950.0" valueMax="2400.0" valueMin="1500.0">1500­2400 m</quantity>
(
<figureCitation id="729A2A17C324C77AE37B71DDFC7BFA2E" box="[861,961,1484,1510]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................................................................................................................6.
<taxonomicName id="2DA14D11C324C77AE3AB71E5FBC4F9C6" authority="Gusarov" authorityName="Gusarov" box="[909,1150,1524,1550]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3AB71E5FBA8F9C5" box="[909,1042,1524,1549]" italics="true" pageId="10" pageNumber="11">S. castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF71E2FB76F9C5" bold="true" box="[1161,1228,1523,1549]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF71E2FB76F9C5" box="[1161,1228,1523,1549]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E720DFA91F996" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E720DFA96F9FE" blockId="10.[264,1326,284,1870]" box="[264,1324,1564,1590]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E720DFA96F9FE" box="[264,1324,1564,1590]" pageId="10" pageNumber="11">
15 Spermatheca S­shaped (
<figureCitation id="729A2A17C324C77AE277720DFD7BF9FE" box="[593,705,1564,1590]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Figs. 149</figureCitation>
,
<figureCitation id="729A2A17C324C77AE2E8720DFD41F9FE" box="[718,763,1564,1590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">160</figureCitation>
) .................................................................... 16
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E7255FA91F996" blockId="10.[264,1326,284,1870]" box="[264,1323,1604,1630]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7255FA91F996" box="[264,1323,1604,1630]" pageId="10" pageNumber="11">
Spermatheca L­, J­ or C­shaped (
<figureCitation id="729A2A17C324C77AE29D7255FCA7F996" box="[699,797,1604,1630]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 details of internal sac retracted into median lobe, abparameral view; 29 everted internal sac, parameral view; 30 medial lamellae, parameral view; 31 right hook of internal sac; 32 spermatheca; 33 apex of left paramere; 34 ­ 35 details of internal sac retracted into median lobe, lateral view; 36 copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="10" pageNumber="11">Figs. 32</figureCitation>
,
<figureCitation id="729A2A17C324C77AE30F7255FCF3F996" box="[809,841,1604,1630]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="10" pageNumber="11">53</figureCitation>
,
<figureCitation id="729A2A17C324C77AE3737255FCCFF996" box="[853,885,1604,1630]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="10" pageNumber="11">62</figureCitation>
,
<figureCitation id="729A2A17C324C77AE3A77255FC1BF996" box="[897,929,1604,1630]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">89</figureCitation>
,
<figureCitation id="729A2A17C324C77AE38B7255FC60F996" box="[941,986,1604,1630]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">104</figureCitation>
) ....................................... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E727DFA97F8EE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E727DFA91F91E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E727DFA91F91E" pageId="10" pageNumber="11">
16 Umbilicus facing proximally (
<figureCitation id="729A2A17C324C77AE28F727DFCAEF94E" box="[681,788,1644,1670]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 everted internal sac, parameral view; 144 medial lamellae, parameral view; 145 ­ 146 details of internal sac retracted into median lobe, lateral view; 147 copulatory piece, lateral view; 148 apex of left paramere; 149 spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Fig. 149</figureCitation>
). Body length
<quantity id="2D599B77C324C77AE3FD727DFBD1F94E" box="[987,1131,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="10" pageNumber="11" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE11E7285FE58F966" box="[312,482,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="10" pageNumber="11" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE2B47285FD55F966" box="[658,751,1684,1710]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitudes of
<quantity id="2D599B77C324C77AE3847285FB85F966" box="[930,1087,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="2000.0" valueMax="2300.0" valueMin="1700.0">1700­2300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE4697285FB0EF966" box="[1103,1204,1684,1710]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............. ..................................................................... 10.
<taxonomicName id="2DA14D11C324C77AE34872ADFBD2F91E" authority="Gusarov" authorityName="Gusarov" box="[878,1128,1724,1750]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE34872ADFC41F91D" box="[878,1019,1724,1749]" italics="true" pageId="10" pageNumber="11">S. mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE45572AAFB0FF91D" bold="true" box="[1139,1205,1723,1749]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE45572AAFB0FF91D" box="[1139,1205,1723,1749]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(females)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E72F5FA97F8EE" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E72F5FA97F8EE" pageId="10" pageNumber="11">
Umbilicus facing laterally (
<figureCitation id="729A2A17C324C77AE25D72F5FD56F936" box="[635,748,1764,1790]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 160</figureCitation>
,
<figureCitation id="729A2A17C324C77AE2DE72F5FC92F936" box="[760,808,1764,1790]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 everted internal sac, parameral view; 170 medial lamellae, parameral view; 171 apex of left paramere; 172 everted internal sac, lateral view; 173 details of internal sac retracted into median lobe, lateral view; 174 spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="10" pageNumber="11">174</figureCitation>
). Two closely related species which cannot be distinguished by females ....................................................................................... 17
</keyLead>
</paragraph>
</keyStep>
<paragraph id="EA1E3692C324C77BE12E7325FA91FE96" blockId="10.[264,1326,284,1870]" lastBlockId="11.[264,1325,284,1870]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7325FA91F886" box="[264,1323,1844,1870]" pageId="10" pageNumber="11">
<keyStep id="51552E37C324C77AE12E7325FA91F886" box="[264,1323,1844,1870]" pageId="10" pageNumber="11">
17 Body length
<quantity id="2D599B77C324C77AE1F67325FD94F886" box="[464,558,1844,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE2C87325FC22F886" box="[750,920,1844,1870]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="10" pageNumber="11" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Spermatheca:
<figureCitation id="729A2A17C324C77AE47D7325FB7AF886" box="[1115,1216,1844,1870]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 median lobe, parameral view; 155 apex of median lobe, parameral view; 156 median lobe, lateral view; 157 apex of median lobe, lateral view; 158 details of internal sac retracted into median lobe, lateral view; 159 apex of left paramere; 160 spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Fig. 160</figureCitation>
. Known
</keyStep>
</keyLead>
<subSection id="832E2F79C325C77BE11E750DFA91FE96" pageId="11" pageNumber="12" type="multiple">
from
<collectingCountry id="92B67602C325C77BE15E750DFDBEFEFE" box="[376,516,284,310]" name="El Salvador" pageId="11" pageNumber="12">El Salvador</collectingCountry>
, at altitude of
<quantity id="2D599B77C325C77BE28C750DFCB9FEFE" box="[682,771,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="12" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C325C77BE335750DFCCDFEFE" box="[787,887,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="11" pageNumber="12">Fig. 387</figureCitation>
) .....................................................................................................................................11.
<taxonomicName id="2DA14D11C325C77BE3BD7555FBD2FE96" authority="Gusarov" authorityName="Gusarov" box="[923,1128,324,350]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C325C77BE3BD7555FC46FE95" box="[923,1020,324,349]" italics="true" pageId="11" pageNumber="12">S. badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C325C77BE4557552FB0FFE95" bold="true" box="[1139,1205,323,349]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="C3E657FBC325C77BE4557552FB0FFE95" box="[1139,1205,323,349]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(females)
</subSection>
</paragraph>
</treatment>
</document>

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<document id="F91FBA367D17F0CB3F3FC9F01E09BF67" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C331C751E12E76BAFE17FADE" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella castanea Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="32" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:titleInfo id="90F71EB48D0806AD0BBDFC6447A294A9">
<mods:title id="DB14E98F9B5B764BF4DF829AA1BD855E">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="FF9AC7EEF5CB0B87851A3965D30EC021">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="B44B1C8EB798FD5DF6EA939AA2366BCB">2003</mods:date>
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<subSubSection id="A2BB6519C331C76FE12E76BAFC22FD0D" box="[264,920,683,710]" pageId="31" pageNumber="32" type="key">
<paragraph id="EA1E3692C331C76FE12E76BAFC22FD0D" blockId="31.[264,920,683,710]" box="[264,920,683,710]" pageId="31" pageNumber="32">
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6.
<taxonomicName id="2DA14D11C331C76FE10076BDFD22FD0D" ID-CoL="6YBD3" authority="Gusarov" authorityName="Gusarov" box="[294,664,683,710]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C331C76FE10076BDFD9CFD0E" bold="true" box="[294,550,684,710]" italics="true" pageId="31" pageNumber="32">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC331C76FE28076BAFD52FD0D" box="[678,744,683,709]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C331C76FE2D176BAFC2AFD0D" box="[759,912,683,709]" captionStart-0="FIGURES 90 ­ 93" captionStart-1="FIGURES 94 ­ 104" captionStartId-0="31.[264,383,1803,1827]" captionStartId-1="32.[264,383,1439,1463]" captionTargetBox-0="[290,1295,933,1772]" captionTargetBox-1="[296,1295,339,1401]" captionTargetId-0="figure@31.[281,1306,921,1779]" captionTargetId-1="figure@32.[282,1305,329,1412]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURES 90 ­ 93. Abdominal segment 8 of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (90 ­ 91) and Las Piedrecitas (92 ­ 93), Mexico). 90 male tergum 8; 91 male sternum 8; 92 female tergum 8; 93 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri-0="https://zenodo.org/record/156437/files/figure.png" httpUri-1="https://zenodo.org/record/156438/files/figure.png" pageId="31" pageNumber="32">Figs. 90­104</figureCitation>
)
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C331C751E12E76E7FD55FE49" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="EA1E3692C331C76FE12E76E7FE40FCA9" blockId="31.[264,1324,758,865]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE12E76E7FE7AFCD8" bold="true" box="[264,448,758,784]" pageId="31" pageNumber="32">
<typeStatus id="351A8830C331C76FE12E76E7FEFFFCD8" box="[264,325,758,784]" pageId="31" pageNumber="32">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C331C76FE1F576E6FDF9FCD9" box="[467,579,759,785]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C331C76FE25376E7FCC4FCD8" bold="true" box="[629,894,758,784]" pageId="31" pageNumber="32">
<collectingCountry id="92B67602C331C76FE25376E7FD4FFCD8" box="[629,757,758,784]" name="Mexico" pageId="31" pageNumber="32">MEXICO</collectingCountry>
: Chiapas:
</emphasis>
<quantity id="2D599B77C331C76FE3B776E6FC5FFCD9" box="[913,997,759,785]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="km" value="2.8">2.8 km</quantity>
NW Las Piedrecitas,
<quantity id="2D599B77C331C76FE4C876E6FA96FCD9" box="[1262,1324,759,785]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="km" value="5.0">5 km</quantity>
NW
<collectingCountry id="92B67602C331C76FE16F770EFE4BFCF1" box="[329,497,799,825]" name="Saint Kitts and Nevis" pageId="31" pageNumber="32">San Cristobal</collectingCountry>
de las Casas,
<quantity id="2D599B77C331C76FE29F770EFCA7FCF1" box="[697,797,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="2400.0">2400 m</quantity>
, oak and pine forest litter (R.Anderson),
<date id="9E1F1052C331C76FE12E7756FE3CFCA9" box="[264,390,839,865]" pageId="31" pageNumber="32" value="1991-09-20">20.ix.1991</date>
(
<collectionCode id="8CB0AE57C331C76FE1B37756FE57FCA9" box="[405,493,839,865]" pageId="31" pageNumber="32">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC331C76FE12E731AFC26F8A0" httpUri="https://zenodo.org/record/156437/files/figure.png" pageId="31" pageNumber="32" targetBox="[290,1295,933,1772]" targetPageId="31">
<paragraph id="EA1E3692C331C76FE12E731AFC26F8A0" blockId="31.[264,1323,1803,1896]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE12E731AFE79F8EB" bold="true" box="[264,451,1803,1827]" pageId="31" pageNumber="32">FIGURES 90­93</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C331C76FE2CF731DFB88F8EA" authority="Gusarov" authorityName="Gusarov" box="[745,1074,1803,1827]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C331C76FE2CF731DFC6AF8EB" box="[745,976,1804,1827]" italics="true" pageId="31" pageNumber="32">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C331C76FE41B731AFBC0F8EB" bold="true" box="[1085,1146,1803,1827]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="C3E657FBC331C76FE41B731AFBC0F8EB" box="[1085,1146,1803,1827]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Rayon (90­91) and Las Piedrecitas (92­93), Mexico). 90 male tergum 8; 91 male sternum 8; 92 female tergum 8; 93 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<caption id="BEDE661AC30EC750E12E718EFE4BF94E" httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="32" pageNumber="33" targetBox="[296,1295,339,1401]" targetPageId="32">
<paragraph id="EA1E3692C30EC750E12E718EFE4BF94E" blockId="32.[264,1323,1439,1704]" pageId="32" pageNumber="33">
<emphasis id="D8D5EA80C30EC750E12E718EFE6BFA7F" bold="true" box="[264,465,1439,1463]" pageId="32" pageNumber="33">FIGURES 94­104</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C30EC750E24D718EFC09FA7E" authority="Gusarov" authorityName="Gusarov" box="[619,947,1439,1462]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C30EC750E24D718EFCE8FA7E" box="[619,850,1439,1462]" italics="true" pageId="32" pageNumber="33">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C30EC750E398718EFC40FA7F" bold="true" box="[958,1018,1439,1463]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="C3E657FBC30EC750E398718EFC40FA7F" box="[958,1018,1439,1463]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Rayon (94­ 97, 99­102) and Las Piedrecitas (98, 103­104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100­101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100­101), 0.1 mm (95, 97, 99, 102­104).
</paragraph>
</caption>
<paragraph id="EA1E3692C30EC750E12E7280FDCDF960" blockId="32.[264,1323,1439,1704]" box="[264,631,1681,1704]" pageId="32" pageNumber="33">DS distal sclerite of internal sac.</paragraph>
<paragraph id="EA1E3692C30EC751E11E72E8FD55FE49" blockId="32.[264,1324,1785,1886]" lastBlockId="33.[264,1324,283,1302]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<typeStatus id="351A8830C30EC750E11E72E8FE15F8DB" box="[312,431,1785,1811]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C30EC750E1E572E8FD70F8DB" bold="true" box="[451,714,1785,1811]" pageId="32" pageNumber="33">
<collectingCountry id="92B67602C30EC750E1E572E8FDF9F8DB" box="[451,579,1785,1811]" name="Mexico" pageId="32" pageNumber="33">MEXICO</collectingCountry>
: Chiapas:
</emphasis>
<specimenCount id="FCA7FD1BC30EC750E2FD72E8FCC4F8DB" box="[731,894,1785,1811]" pageId="32" pageNumber="33" type="generic">10 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C30EC750E47372E8FB05F8DB" box="[1109,1215,1785,1811]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC30EC750E4EB72E8FEF3F8F1" pageId="32" pageNumber="33" type="generic">6 specimens</specimenCount>
, Yerbabuena Reserve,
<quantity id="2D599B77C30EC750E271730EFD13F8F1" box="[599,681,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="32" pageNumber="33" unit="km" value="2.1">2.1 km</quantity>
NW Pueblo Nuevo Solistahuacan,
<quantity id="2D599B77C30EC750E469730EFB16F8F1" box="[1103,1196,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="32" pageNumber="33" unit="m" value="2070.0">2070 m</quantity>
, liquidambar forest litter (R.Anderson),
<date id="9E1F1052C30EC750E2507355FD4CF896" box="[630,758,1860,1886]" pageId="32" pageNumber="33" value="1992-09-23">23.ix.1992</date>
;
<specimenCount id="FCA7FD1BC30EC750E3237355FC20F896" box="[773,922,1860,1886]" pageId="32" pageNumber="33" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C30EC750E38F7355FC47F896" box="[937,1021,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="8.9" pageId="32" pageNumber="33" unit="km" value="8.9">8.9 km</quantity>
E Rayon,
<quantity id="2D599B77C30EC750E4597355FB66F896" box="[1151,1244,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="1500.0">1500 m</quantity>
, cloud forest litter (R.Anderson),
<date id="9E1F1052C30FC751E261750DFD72FEFE" box="[583,712,284,310]" pageId="33" pageNumber="34" value="1991-09-19">19.ix.1991</date>
;
<emphasis id="D8D5EA80C30FC751E2F0750AFC47FEFD" bold="true" box="[726,1021,283,309]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E2F0750AFC38FEFD" box="[726,898,283,309]" name="Honduras" pageId="33" pageNumber="34">HONDURAS</collectingCountry>
: La Paz:
</emphasis>
2, 4, Cord. Montecillos,
<quantity id="2D599B77C30FC751E1487550FE06FE93" box="[366,444,321,347]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="34" unit="km" value="18.0">18 km</quantity>
NW La Paz,
<geoCoordinate id="8F955055C30FC751E2407550FD7DFE93" box="[614,711,321,347]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="14.35">14°21'N</geoCoordinate>
<geoCoordinate id="8F955055C30FC751E2F77550FC83FE93" box="[721,825,321,347]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-87.78333">87°47'W</geoCoordinate>
,
<quantity id="2D599B77C30FC751E3617550FC1CFE93" box="[839,934,321,347]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="m" value="2000.0">2000 m</quantity>
, pine and hardwood forest litter (R.Anderson),
<date id="9E1F1052C30FC751E1917576FDFFFE49" box="[439,581,359,385]" pageId="33" pageNumber="34" value="1994-08-21">21.viii.1994</date>
(all ­
<collectionCode id="8CB0AE57C30FC751E2AA7576FD59FE49" box="[652,739,359,385]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E759AFDBAFCFE" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="EA1E3692C30FC751E11E759AFDEAFD8E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E759AFE0DFE6D" bold="true" box="[312,439,395,421]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30FC751E1EF759DFD72FE6D" box="[457,712,396,421]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E1EF759DFD72FE6D" box="[457,712,396,421]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30FC751E4F5759DFEFDFE05" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30FC751E4F5759DFEFDFE05" italics="true" pageId="33" pageNumber="34">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.5­1.7 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer or as long as pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C30FC751E4AB7615FA99FDD6" box="[1165,1315,516,542]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 94­103</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C30FC751E1FA763DFDF9FD8E" box="[476,579,556,582]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7645FDBAFCFE" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<taxonomicName id="2DA14D11C30FC751E11E7645FD82FDA5" box="[312,568,596,621]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E11E7645FD82FDA5" box="[312,568,596,621]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30FC751E3A17645FB9EFDA5" box="[903,1060,596,621]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E3A17645FB9EFDA5" box="[903,1060,596,621]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5, 7­8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have strong spicules (
<figureCitation id="729A2A17C30FC751E34B76DDFBB3FD2E" box="[877,1033,716,742]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 99, 101</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C30FC751E2C076E5FCCDFCC6" box="[742,887,756,782]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 96­97</figureCitation>
); spermatheca C­shaped with large umbilicus (
<figureCitation id="729A2A17C30FC751E1AA770DFE48FCFE" box="[396,498,796,822]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E7752FDC4FB56" pageId="33" pageNumber="34" type="description">
<paragraph id="EA1E3692C30FC751E11E7752FCD4FC4E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E7752FE6AFC95" bold="true" box="[312,464,835,861]" pageId="33" pageNumber="34">Description.</emphasis>
Length
<quantity id="2D599B77C30FC751E21F7755FD7FFC96" box="[569,709,836,862]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="33" pageNumber="34" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C30FC751E3A57755FB96FC96" box="[899,1068,836,862]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="33" pageNumber="34" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Body brown to dark brown with lighter pronotum and elytra, legs brown.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7785FB89FC1E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
In all external character states
<taxonomicName id="2DA14D11C30FC751E2B97785FC9EFC65" box="[671,804,916,941]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E2B97785FC9EFC65" box="[671,804,916,941]" italics="true" pageId="33" pageNumber="34">S. castanea</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="2DA14D11C30FC751E3997785FBE2FC65" box="[959,1112,916,941]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E3997785FBE2FC65" box="[959,1112,916,941]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E77F5FD73FB86" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">In large males posterior angles of male tergum 3 projecting as spines up to 1 time as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C30FC751E11E704DFD37FBBE" blockId="33.[264,1324,283,1302]" box="[312,653,1116,1142]" pageId="33" pageNumber="34">
Aedeagus as in
<figureCitation id="729A2A17C30FC751E1D7704DFD33FBBE" box="[497,649,1116,1142]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 94­103</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7095FDC4FB56" blockId="33.[264,1324,283,1302]" box="[312,638,1156,1182]" pageId="33" pageNumber="34">
Spermatheca as in
<figureCitation id="729A2A17C30FC751E2337095FDC0FB56" box="[533,634,1156,1182]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 median lobe, parameral view; 95 apex of median lobe, parameral view; 96 median lobe, lateral view; 97 apex of median lobe, lateral view; 98 everted internal sac, parameral view; 99 right hook of internal sac; 100 ­ 101 details of internal sac retracted into median lobe, lateral view; 102 apex of left paramere; 103 copulatory piece, lateral view; 104 spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E70BAFE17FADE" pageId="33" pageNumber="34" type="distribution">
<paragraph id="EA1E3692C30FC751E11E70BAFBB4FB0E" blockId="33.[264,1324,283,1302]" box="[312,1038,1195,1222]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E70BAFE62FB0D" bold="true" box="[312,472,1195,1221]" pageId="33" pageNumber="34">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30FC751E2A370BDFD65FB0E" box="[645,735,1196,1222]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
and
<collectingCountry id="92B67602C30FC751E33E70BDFC30FB0E" box="[792,906,1196,1222]" name="Honduras" pageId="33" pageNumber="34">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C30FC751E3BC70BDFBB8FB0E" box="[922,1026,1196,1222]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="33" pageNumber="34">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E70C2FE17FADE" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E70C2FDB3FB25" bold="true" box="[312,521,1235,1261]" pageId="33" pageNumber="34">Natural History.</emphasis>
<taxonomicName id="2DA14D11C30FC751E20670C5FC9BFB25" box="[544,801,1236,1261]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E20670C5FC9BFB25" box="[544,801,1236,1261]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C30FC751E12E70EDFE12FADE" box="[264,424,1276,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="33" pageNumber="34" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">2000­2400 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,219 @@
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<subSubSection id="A2BB6519C333C76DE12E750AFCDBFEFD" box="[264,865,283,310]" pageId="29" pageNumber="30" type="key">
<paragraph id="EA1E3692C333C76DE12E750AFCDBFEFD" blockId="29.[264,865,283,310]" box="[264,865,283,310]" pageId="29" pageNumber="30">
<heading id="B15681FEC333C76DE12E750AFCDBFEFD" bold="true" box="[264,865,283,310]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<key id="9E3061A3C333C76DE12E750AFCDBFEFD" box="[264,865,283,310]" pageId="29" pageNumber="30">
<keyStep id="51552E37C333C76DE12E750AFCDBFEFD" box="[264,865,283,310]" pageId="29" pageNumber="30">
<keyLead id="515095A7C333C76DE12E750AFCDBFEFD" box="[264,865,283,310]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE12E750AFCDBFEFD" bold="true" box="[264,865,283,310]" pageId="29" pageNumber="30">
5.
<taxonomicName id="2DA14D11C333C76DE100750DFDCAFEFD" ID-CoL="6YBCQ" authority="Gusarov" authorityName="Gusarov" box="[294,624,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C333C76DE100750DFE47FEFE" bold="true" box="[294,509,284,310]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC333C76DE25B750AFD05FEFD" box="[637,703,283,309]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C333C76DE2E9750AFCE3FEFD" box="[719,857,283,309]" captionStart-0="FIGURES 79 ­ 82" captionStart-1="FIGURES 83 ­ 89" captionStartId-0="29.[264,382,1486,1510]" captionStartId-1="30.[264,383,1432,1456]" captionTargetBox-0="[291,1296,652,1452]" captionTargetBox-1="[285,1297,538,1400]" captionTargetId-0="figure@29.[277,1311,641,1462]" captionTargetId-1="figure@30.[274,1314,526,1408]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURES 79 ­ 82. Abdominal segment 8 of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 79 male tergum 8; 80 male sternum 8; 81 female tergum 8; 82 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri-0="https://zenodo.org/record/156435/files/figure.png" httpUri-1="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Figs. 79­89</figureCitation>
)
</emphasis>
</keyLead>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C333C76DE12E7577FCF3FDE9" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="EA1E3692C333C76DE12E7577FC23FE61" blockId="29.[264,1323,358,545]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE12E7577FE01FE48" bold="true" box="[264,443,358,384]" pageId="29" pageNumber="30">
<typeStatus id="351A8830C333C76DE12E7577FEF9FE48" box="[264,323,358,384]" pageId="29" pageNumber="30">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C333C76DE1ED7576FD81FE49" box="[459,571,359,385]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C333C76DE24E7577FCC0FE48" bold="true" box="[616,890,358,384]" pageId="29" pageNumber="30">
<collectingCountry id="92B67602C333C76DE24E7577FD52FE48" box="[616,744,358,384]" name="Mexico" pageId="29" pageNumber="30">MEXICO</collectingCountry>
: Guerrero:
</emphasis>
<quantity id="2D599B77C333C76DE3AF7576FC69FE49" box="[905,979,359,385]" metricMagnitude="4" metricUnit="m" metricValue="7.1" pageId="29" pageNumber="30" unit="km" value="71.0">71 km</quantity>
NE Atoyac de Alvarez,
<quantity id="2D599B77C333C76DE4D67576FE98FE61" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="30" unit="m" value="1700.0">1700 m</quantity>
, flight intercept trap (J.S.Ashe),
<date id="9E1F1052C333C76DE286759EFC9FFE61" box="[672,805,399,425]" pageId="29" pageNumber="30" value="1992-07-27">27.vii.1992</date>
(
<collectionCode id="8CB0AE57C333C76DE313759EFC37FE61" box="[821,909,399,425]" pageId="29" pageNumber="30">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C333C76DE11E75A6FCF3FDE9" blockId="29.[264,1323,358,545]" pageId="29" pageNumber="30">
<typeStatus id="351A8830C333C76DE11E75A6FE15FE19" box="[312,431,439,465]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C333C76DE1E375A7FD66FE18" bold="true" box="[453,732,438,464]" pageId="29" pageNumber="30">
<collectingCountry id="92B67602C333C76DE1E375A7FDFFFE18" box="[453,581,438,464]" name="Mexico" pageId="29" pageNumber="30">MEXICO</collectingCountry>
: Guerrero:
</emphasis>
, 2, same data as the
<typeStatus id="351A8830C333C76DE40F75A6FB2EFE19" box="[1065,1172,439,465]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
;, ditto but
<date id="9E1F1052C333C76DE12E75CEFE2BFE31" box="[264,401,479,505]" pageId="29" pageNumber="30" value="1992-07-25">25.vii.1992</date>
, forest litter;
<emphasis id="D8D5EA80C333C76DE21675CFFD23FE30" bold="true" box="[560,665,478,504]" pageId="29" pageNumber="30">Oaxaca:</emphasis>
2,,
<quantity id="2D599B77C333C76DE33475CEFCCEFE31" box="[786,884,479,505]" metricMagnitude="4" metricUnit="m" metricValue="3.84" pageId="29" pageNumber="30" unit="km" value="38.4">38.4 km</quantity>
N Telixtlahuaca, Hwy. 131, 1880 m, forest litter (J.S.Ashe),
<date id="9E1F1052C333C76DE23C7616FD25FDE9" box="[538,671,519,545]" pageId="29" pageNumber="30" value="1992-07-19">19.vii.1992</date>
(all ­
<collectionCode id="8CB0AE57C333C76DE2C07616FC87FDE9" box="[742,829,519,545]" pageId="29" pageNumber="30">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C333C76DE12E71DFFD76F9E3" blockId="29.[264,1322,1486,1579]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE12E71DFFE7FFA2E" bold="true" box="[264,453,1486,1510]" pageId="29" pageNumber="30">FIGURES 79­82</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C333C76DE2D771DEFBADFA2E" authority="Gusarov" authorityName="Gusarov" box="[753,1047,1487,1510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C333C76DE2D771DEFC08FA2E" box="[753,946,1487,1510]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C333C76DE40471DFFBDAFA2E" bold="true" box="[1058,1120,1486,1510]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C3E657FBC333C76DE40471DFFBDAFA2E" box="[1058,1120,1486,1510]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, 71 km NE Atoyac de Alvarez, Mexico). 79 male tergum 8; 80 male sternum 8; 81 female tergum 8; 82 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C333C76EE11E7297FDCDF997" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="EA1E3692C333C76DE11E7297FB60F8D1" blockId="29.[264,1323,1670,1897]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE11E7297FE02F968" bold="true" box="[312,440,1670,1696]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C333C76DE1E17296FD21F968" box="[455,667,1671,1696]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C333C76DE1E17296FD21F968" box="[455,667,1671,1696]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C333C76DE4BB7296FA91F968" box="[1181,1323,1671,1696]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C333C76DE4BB7296FA91F968" box="[1181,1323,1671,1696]" italics="true" pageId="29" pageNumber="30">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body; temples as long as eyes; glossy pronotum with weak microsculpture; elytra longer than pronotum; wings fully developed, tergum 8 with four pairs of macrosetae, the distinct shape of aedeagus (
<figureCitation id="729A2A17C333C76DE32E72EEFC2AF8D1" box="[776,912,1791,1817]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Figs. 83­88</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C333C76DE45272EEFB76F8D1" box="[1140,1228,1791,1817]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Fig. 89</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C333C76EE11E7336FCF9FE66" blockId="29.[264,1323,1670,1897]" lastBlockId="30.[264,1324,284,430]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<taxonomicName id="2DA14D11C333C76DE11E7336FDB7F888" box="[312,525,1831,1856]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C333C76DE11E7336FDB7F888" box="[312,525,1831,1856]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C333C76DE3737336FC4AF888" box="[853,1008,1831,1856]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C333C76DE3737336FC4AF888" box="[853,1008,1831,1856]" italics="true" pageId="29" pageNumber="30">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 6­8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have weak spicules (
<figureCitation id="729A2A17C330C76EE3037555FCC6FE96" box="[805,892,324,350]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Fig. 87</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C330C76EE219757DFD76FE4E" box="[575,716,364,390]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Figs. 85­86</figureCitation>
); spermatheca L­shaped with small umbilicus and slightly curved in the proximal portion (
<figureCitation id="729A2A17C330C76EE2F87585FC8FFE66" box="[734,821,404,430]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Fig. 89</figureCitation>
).
</paragraph>
<caption id="BEDE661AC330C76EE12E7189FD53F9F2" httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31" targetBox="[285,1297,538,1400]" targetPageId="30">
<paragraph id="EA1E3692C330C76EE12E7189FD53F9F2" blockId="30.[264,1323,1432,1631]" pageId="30" pageNumber="31">
<emphasis id="D8D5EA80C330C76EE12E7189FE7EFA78" bold="true" box="[264,452,1432,1456]" pageId="30" pageNumber="31">FIGURES 83­89</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C330C76EE27B7188FCC5FA78" authority="Gusarov" authorityName="Gusarov" box="[605,895,1433,1456]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C330C76EE27B7188FCA7FA78" box="[605,797,1433,1456]" italics="true" pageId="30" pageNumber="31">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C330C76EE3AC7189FC7FFA78" bold="true" box="[906,965,1432,1456]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="C3E657FBC330C76EE3AC7189FC7FFA78" box="[906,965,1432,1456]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88­89).
</paragraph>
</caption>
<paragraph id="EA1E3692C330C76EE12E7259FDCDF997" blockId="30.[264,1323,1432,1631]" box="[264,631,1608,1631]" pageId="30" pageNumber="31">DS distal sclerite of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C330C76FE11E72ACFDD5FE1E" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="EA1E3692C330C76EE11E72ACFDCBF8C8" blockId="30.[264,1323,1725,1872]" pageId="30" pageNumber="31">
<emphasis id="D8D5EA80C330C76EE11E72ACFE6AF91F" bold="true" box="[312,464,1725,1751]" pageId="30" pageNumber="31">Description.</emphasis>
Length
<quantity id="2D599B77C330C76EE26772AFFD75F910" box="[577,719,1726,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="30" pageNumber="31" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3­2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C330C76EE3B372AFFBFBF910" box="[917,1089,1726,1752]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="30" pageNumber="31" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Body dark brown with lighter elytra, legs brown.
</paragraph>
<paragraph id="EA1E3692C330C76EE11E731FFB89F898" blockId="30.[264,1323,1725,1872]" pageId="30" pageNumber="31">
In all external character states
<taxonomicName id="2DA14D11C330C76EE2BF731FFD4EF8EF" box="[665,756,1806,1831]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C330C76EE2BF731FFD4EF8EF" box="[665,756,1806,1831]" italics="true" pageId="30" pageNumber="31">S. fusca</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C330C76EE3E3731FFBE7F8EF" box="[965,1117,1806,1831]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C330C76EE3E3731FFBE7F8EF" box="[965,1117,1806,1831]" italics="true" pageId="30" pageNumber="31">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C331C76FE11E750DFE03FE4E" blockId="31.[264,1324,284,590]" pageId="31" pageNumber="32">In four examined males posterior angles of tergum 3 not projecting at all or projecting as spines up to 1/3 as long as tergum 3 (measured medially), tergum 7 without medial carina or tubercle.</paragraph>
<paragraph id="EA1E3692C331C76FE11E7585FDC7FE66" blockId="31.[264,1324,284,590]" box="[312,637,404,430]" pageId="31" pageNumber="32">
Aedeagus as in
<figureCitation id="729A2A17C331C76FE1D67585FDC3FE66" box="[496,633,404,430]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="31" pageNumber="32">Figs. 83­88</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C331C76FE11E75ADFDD5FE1E" blockId="31.[264,1324,284,590]" box="[312,623,444,470]" pageId="31" pageNumber="32">
Spermatheca as in
<figureCitation id="729A2A17C331C76FE23275ADFDD1FE1E" box="[532,619,444,470]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 median lobe, parameral view; 84 apex of median lobe, parameral view; 85 median lobe, lateral view; 86 apex of median lobe, lateral view; 87 details of internal sac retracted into median lobe, lateral view; 88 apex of left paramere; 89 spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="31" pageNumber="32">Fig. 89</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C331C76FE11E75F2FD7BFD86" pageId="31" pageNumber="32" type="distribution">
<paragraph id="EA1E3692C331C76FE11E75F2FCD8FE36" blockId="31.[264,1324,284,590]" box="[312,866,483,510]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE11E75F2FE62FE35" bold="true" box="[312,472,483,509]" pageId="31" pageNumber="32">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C331C76FE2A375F5FD65FE36" box="[645,735,484,510]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C331C76FE2C875F5FCEFFE36" box="[750,853,484,510]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="31" pageNumber="32">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C331C76FE11E761AFD7BFD86" blockId="31.[264,1324,284,590]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE11E761AFDBEFDED" bold="true" box="[312,516,523,549]" pageId="31" pageNumber="32">Natural History.</emphasis>
<taxonomicName id="2DA14D11C331C76FE235761DFD5CFDED" box="[531,742,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C331C76FE235761DFD5CFDED" box="[531,742,524,549]" italics="true" pageId="31" pageNumber="32">Seeversiella fusca</emphasis>
</taxonomicName>
was collected in forest litter and with flight intercept traps at altitude of
<quantity id="2D599B77C331C76FE23A7625FD06FD86" box="[540,700,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="31" pageNumber="32" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700­1900 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,220 @@
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<keyStep id="51552E37C335C76BE12E750AFC2FFEFD" box="[264,917,283,310]" pageId="27" pageNumber="28">
4.
<taxonomicName id="2DA14D11C335C76BE100750DFD1EFEFD" ID-CoL="6YBCT" authority="Gusarov" authorityName="Gusarov" box="[294,676,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C335C76BE100750DFD8BFEFE" bold="true" box="[294,561,284,310]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC335C76BE297750AFD49FEFD" box="[689,755,283,309]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C335C76BE325750AFC37FEFD" box="[771,909,283,309]" captionStart-0="FIGURES 68 ­ 71" captionStart-1="FIGURES 72 ­ 78" captionStartId-0="27.[264,383,1474,1498]" captionStartId-1="28.[264,383,1298,1322]" captionTargetBox-0="[282,1307,629,1441]" captionTargetBox-1="[300,1285,615,1266]" captionTargetId-0="figure@27.[268,1319,604,1448]" captionTargetId-1="figure@28.[296,1291,609,1272]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURES 68 ­ 71. Abdominal segment 8 of Seeversiella liliputana Gusarov, sp. n. (68 ­ 69, holotype; 70 ­ 71, possible female of S. liliputana from Bear Saddle, Arizona). 68 male tergum 8; 69 male sternum 8; 70 female tergum 8; 71 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri-0="https://zenodo.org/record/156433/files/figure.png" httpUri-1="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 68­78</figureCitation>
)
</keyStep>
</key>
</keyLead>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C335C76BE12E7577FDB8FE31" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="EA1E3692C335C76BE12E7577FC01FE61" blockId="27.[264,1323,358,505]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="27" pageNumber="28">
<typeStatus id="351A8830C335C76BE12E7577FEF9FE48" box="[264,323,358,384]" pageId="27" pageNumber="28">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C335C76BE1F07576FDFCFE49" box="[470,582,359,385]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C335C76BE25D7577FC4AFE48" bold="true" box="[635,1008,358,384]" pageId="27" pageNumber="28">
<collectingCountry id="92B67602C335C76BE25D7577FCD2FE48" box="[635,872,358,384]" name="United States of America" pageId="27" pageNumber="28">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.: Huachuca Mts., Bear Saddle, 8100' (A.Smetana),
<date id="9E1F1052C335C76BE2F5759EFCE9FE61" box="[723,851,399,425]" pageId="27" pageNumber="28" value="1979-08-05">5.viii.1979</date>
(
<collectionCode id="8CB0AE57C335C76BE344759EFC14FE61" box="[866,942,399,425]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="28">CNCI</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C335C76BE11E75A7FDB8FE31" blockId="27.[264,1323,358,505]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE11E75A7FC0DFE18" bold="true" box="[312,951,438,464]" pageId="27" pageNumber="28">
Additional material.
<collectingCountry id="92B67602C335C76BE26175A7FC89FE18" box="[583,819,438,464]" name="United States of America" pageId="27" pageNumber="28">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.:, same data as the
<typeStatus id="351A8830C335C76BE11575CEFE20FE31" box="[307,410,479,505]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C335C76BE18C75CEFE4DFE31" box="[426,503,479,505]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="28">CNCI</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC335C76BE12E71D3FBCAF9D7" httpUri="https://zenodo.org/record/156433/files/figure.png" pageId="27" pageNumber="28" targetBox="[282,1307,629,1441]" targetPageId="27">
<paragraph id="EA1E3692C335C76BE12E71D3FBCAF9D7" blockId="27.[264,1322,1474,1567]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE12E71D3FE7FFA12" bold="true" box="[264,453,1474,1498]" pageId="27" pageNumber="28">FIGURES 68­71</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C335C76BE2D271D2FBF3FA12" authority="Gusarov" authorityName="Gusarov" box="[756,1097,1475,1498]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C335C76BE2D271D2FC5FFA12" box="[756,997,1475,1498]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C335C76BE47071D3FB29FA12" bold="true" box="[1110,1171,1474,1498]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="C3E657FBC335C76BE47071D3FB29FA12" box="[1110,1171,1474,1498]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(68­69, holotype; 70­71, possible female of
<taxonomicName id="2DA14D11C335C76BE27C71F7FD66FA35" box="[602,732,1510,1533]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE27C71F7FD66FA35" box="[602,732,1510,1533]" italics="true" pageId="27" pageNumber="28">S. liliputana</emphasis>
</taxonomicName>
from Bear Saddle, Arizona). 68 male tergum 8; 69 male sternum 8; 70 female tergum 8; 71 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C335C76BE11E726BFCF8F895" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="EA1E3692C335C76BE11E726BFD3FF8C5" blockId="27.[264,1323,1658,1885]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE11E726BFE0DF95C" bold="true" box="[312,439,1658,1684]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C335C76BE1EE726AFD75F95C" box="[456,719,1659,1684]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE1EE726AFD75F95C" box="[456,719,1659,1684]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C335C76BE4F5726AFEFDF974" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C335C76BE4F5726AFEFDF974" italics="true" pageId="27" pageNumber="28">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 2 times as long as eyes; pronotum without microsculpture; elytra as long as pronotum; tergum 8 with four pairs of macrosetae, and the distinct shape of aedeagus (
<figureCitation id="729A2A17C335C76BE1CB72E2FDC3F8C5" box="[493,633,1779,1805]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 72­77</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C335C76BE11E730AFCF8F895" blockId="27.[264,1323,1658,1885]" pageId="27" pageNumber="28">
<taxonomicName id="2DA14D11C335C76BE11E730AFDFAF8FC" box="[312,576,1819,1844]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE11E730AFDFAF8FC" box="[312,576,1819,1844]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C335C76BE2FB730AFC26F8FC" box="[733,924,1819,1844]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C335C76BE2FB730AFC26F8FC" box="[733,924,1819,1844]" italics="true" pageId="27" pageNumber="28">S. sonomotoides</emphasis>
</taxonomicName>
in having smaller median lobe of aedeagus with shorter apex (
<figureCitation id="729A2A17C335C76BE2707352FD5AF895" box="[598,736,1859,1885]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 72­75</figureCitation>
; 58­61).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C332C76CE11E750AFBB8F9BD" pageId="28" pageNumber="29" type="description">
<paragraph id="EA1E3692C332C76CE11E750AFC36FE96" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="28" pageNumber="29">Description.</emphasis>
Length
<quantity id="2D599B77C332C76CE21E750DFD2EFEFE" box="[568,660,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="29" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C332C76CE377750DFC0CFEFE" box="[849,950,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="28" pageNumber="29" unit="mm" value="0.41">0.41mm</quantity>
. Body brown, elytra and antennae light brown, legs and mouthparts brownish yellow.
</paragraph>
<paragraph id="EA1E3692C332C76CE11E757DFC30FE66" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">
In all character states
<taxonomicName id="2DA14D11C332C76CE211757DFD7CFE4D" box="[567,710,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE211757DFD7CFE4D" box="[567,710,364,389]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C332C76CE3BD757DFBE0FE4D" box="[923,1114,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C332C76CE3BD757DFBE0FE4D" box="[923,1114,364,389]" italics="true" pageId="28" pageNumber="29">S. sonomotoides</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C332C76CE11E75ADFD34FE36" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">In the only known male specimen the posterior angles of tergum 3 not projecting, tergum 7 without carina or tubercle.</paragraph>
<paragraph id="EA1E3692C332C76CE11E761DFDC7FDEE" blockId="28.[264,1323,283,550]" box="[312,637,524,550]" pageId="28" pageNumber="29">
Aedeagus as in
<figureCitation id="729A2A17C332C76CE1D6761DFDC3FDEE" box="[496,633,524,550]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29">Figs. 72­77</figureCitation>
.
</paragraph>
<caption id="BEDE661AC332C76CE12E7103FB28FA7C" httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29" targetBox="[300,1285,615,1266]" targetPageId="28">
<paragraph id="EA1E3692C332C76CE12E7103FB28FA7C" blockId="28.[264,1323,1298,1460]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE12E7103FE7FFAE2" bold="true" box="[264,453,1298,1322]" pageId="28" pageNumber="29">FIGURES 72­78</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C332C76CE2417102FC01FAE2" authority="Gusarov" authorityName="Gusarov" box="[615,955,1299,1322]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C332C76CE2417102FCE2FAE2" box="[615,856,1299,1322]" italics="true" pageId="28" pageNumber="29">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C332C76CE3EE7103FBBCFAE2" bold="true" box="[968,1030,1298,1322]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="C3E657FBC332C76CE3EE7103FBBCFAE2" box="[968,1030,1298,1322]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(72­77, holotype; 78, possible female of
<taxonomicName id="2DA14D11C332C76CE1967127FD89FA85" box="[432,563,1334,1357]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE1967127FD89FA85" box="[432,563,1334,1357]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77­78).
</paragraph>
</caption>
<paragraph id="EA1E3692C332C76CE11E721AFBB8F9BD" blockId="28.[264,1323,1547,1733]" pageId="28" pageNumber="29">
One female specimen, externally similar to the
<typeStatus id="351A8830C332C76CE347721AFC72F9ED" box="[865,968,1547,1573]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2DA14D11C332C76CE3C9721AFBC4F9EC" box="[1007,1150,1547,1572]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE3C9721AFBC4F9EC" box="[1007,1150,1547,1572]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
, was collected together with this
<typeStatus id="351A8830C332C76CE1FB7222FDFDF985" box="[477,583,1587,1613]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
. However that female specimen has the spermatheca like in
<taxonomicName id="2DA14D11C332C76CE5337222FE13F9BC" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C332C76CE5337222FE13F9BC" italics="true" pageId="28" pageNumber="29">S. sonomotoides</emphasis>
</taxonomicName>
(
<figureCitation id="729A2A17C332C76CE19F724AFDA6F9BD" box="[441,540,1627,1653]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 median lobe, parameral view; 73 apex of median lobe, parameral view; 74 median lobe, lateral view; 75 apex of median lobe, lateral view; 76 details of internal sac retracted into median lobe, lateral view; 77 apex of left paramere; 78 spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29">Figs. 78</figureCitation>
,
<figureCitation id="729A2A17C332C76CE20E724AFDFDF9BD" box="[552,583,1627,1653]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="28" pageNumber="29">62</figureCitation>
) and may belong to the latter species.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C332C76CE11E7293FB79F90D" pageId="28" pageNumber="29" type="distribution">
<paragraph id="EA1E3692C332C76CE11E7293FB21F955" blockId="28.[264,1323,1547,1733]" box="[312,1179,1666,1693]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E7293FE62F954" bold="true" box="[312,472,1666,1692]" pageId="28" pageNumber="29">Distribution.</emphasis>
Known from the Huachuca Mountains, Arizona (
<figureCitation id="729A2A17C332C76CE4017292FB35F955" box="[1063,1167,1667,1693]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="28" pageNumber="29">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C332C76CE11E72BBFB79F90D" blockId="28.[264,1323,1547,1733]" box="[312,1219,1706,1733]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E72BBFDBFF90C" bold="true" box="[312,517,1706,1732]" pageId="28" pageNumber="29">Natural History.</emphasis>
<taxonomicName id="2DA14D11C332C76CE23572BAFCA1F90C" box="[531,795,1707,1732]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE23572BAFCA1F90C" box="[531,795,1707,1732]" italics="true" pageId="28" pageNumber="29">Seeversiella liliputana</emphasis>
</taxonomicName>
was collected at altitude of
<quantity id="2D599B77C332C76CE44572BAFB05F90D" box="[1123,1215,1707,1733]" metricMagnitude="3" metricUnit="m" metricValue="2.4699999999999998" pageId="28" pageNumber="29" unit="m" value="2470.0">2470 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="435DF87D3BC9C770377D10F09EF21FA1" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C336C76AE12E77F7FDCDF8A0" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella sonomotoides Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="27" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="25" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="393DCAC902FA4C05EBCF86FA451CDF50">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25" type="key">
<paragraph id="EA1E3692C336C768E12E77F7FC7EFBC8" blockId="24.[264,964,998,1024]" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<heading id="B15681FEC336C768E12E77F7FC7EFBC8" bold="true" box="[264,964,998,1024]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<key id="9E3061A3C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<keyStep id="51552E37C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E12E77F7FC7EFBC8" bold="true" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<keyLead id="515095A7C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25">
3.
<taxonomicName id="2DA14D11C336C768E10077F7FD69FBC8" ID-CoL="6YBBF" authority="Gusarov" authorityName="Gusarov" box="[294,723,998,1024]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C336C768E10077F7FDDAFBC8" bold="true" box="[294,608,998,1024]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC336C768E2C677F7FC98FBC8" box="[736,802,998,1024]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C336C768E31477F7FC06FBC8" box="[818,956,998,1024]" captionStart-0="FIGURES 54 ­ 57" captionStart-1="FIGURES 58 ­ 67" captionStartId-0="25.[264,383,1800,1824]" captionStartId-1="26.[264,382,1663,1687]" captionTargetBox-0="[322,1265,1015,1769]" captionTargetBox-1="[289,1302,621,1628]" captionTargetId-0="figure@25.[313,1274,1006,1776]" captionTargetId-1="figure@26.[279,1307,609,1637]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 54 ­ 57. Abdominal segment 8 of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 54 male tergum 8; 55 male sternum 8; 56 female tergum 8; 57 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri-0="https://zenodo.org/record/156431/files/figure.png" httpUri-1="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 54­67</figureCitation>
)
</keyLead>
</emphasis>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E12E7020FBFBFB0B" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="EA1E3692C336C768E12E7020FB16FBBB" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E12E7020FE05FB83" bold="true" box="[264,447,1073,1099]" pageId="24" pageNumber="25">
<typeStatus id="351A8830C336C768E12E7020FEF9FB83" box="[264,323,1073,1099]" pageId="24" pageNumber="25">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C336C768E1F07020FDFCFB83" box="[470,582,1073,1099]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C336C768E25D7020FC4AFB83" bold="true" box="[635,1008,1073,1099]" pageId="24" pageNumber="25">
<collectingCountry id="92B67602C336C768E25D7020FCD2FB83" box="[635,872,1073,1099]" name="United States of America" pageId="24" pageNumber="25">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.: Huachuca Mts., Carr Cny.,
<quantity id="2D599B77C336C768E1EA7048FD92FBBB" box="[460,552,1113,1139]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="24" pageNumber="25" unit="m" value="2150.0">2150 m</quantity>
, leaf oak litter (P.M.Hammond),
<date id="9E1F1052C336C768E38D7048FB8BFBBB" box="[939,1073,1113,1139]" pageId="24" pageNumber="25" value="1976-07-23">23.vii.1976</date>
(
<collectionCode id="8CB0AE57C336C768E4677048FB1AFBBB" box="[1089,1184,1113,1139]" pageId="24" pageNumber="25">BMNH</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7090FBFBFB0B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<typeStatus id="351A8830C336C768E11E7090FE15FB53" box="[312,431,1153,1179]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C336C768E1E77090FC8AFB53" bold="true" box="[449,816,1153,1179]" pageId="24" pageNumber="25">
<collectingCountry id="92B67602C336C768E1E77090FD16FB53" box="[449,684,1153,1179]" name="United States of America" pageId="24" pageNumber="25">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.:
<specimenCount id="FCA7FD1BC336C768E3C17090FB33FB53" box="[999,1161,1153,1179]" pageId="24" pageNumber="25" type="generic">44 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C336C768E11570B8FE20FB0B" box="[307,410,1193,1219]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C336C768E18F70B8FDBDFB0B" box="[425,519,1193,1219]" pageId="24" pageNumber="25">BMNH</collectionCode>
); 2, ditto but (J.M.Campbell) (
<collectionCode id="8CB0AE57C336C768E3B570B8FC64FB0B" box="[915,990,1193,1219]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="24" pageNumber="25">CNCI</collectionCode>
,
<collectionCode id="8CB0AE57C336C768E3CF70B8FB8EFB0B" box="[1001,1076,1193,1219]" pageId="24" pageNumber="25">SPSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E11E70C0FC6DF9CB" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="EA1E3692C336C768E11E70C0FE65FA7B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E70C0FE0DFB23" bold="true" box="[312,439,1233,1259]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C336C768E1E070C3FD47FB23" box="[454,765,1234,1259]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E1E070C3FD47FB23" box="[454,765,1234,1259]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C336C768E4DE70C3FED7FADB" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C336C768E4DE70C3FED7FADB" italics="true" pageId="24" pageNumber="25">Seeversiella</emphasis>
</taxonomicName>
by having brown body; large eyes (temples 1.7­2.1 times as long as eyes); glossy pronotum without microsculpture or with weak isodiametric microsculpture; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C336C768E3D37158FB6DFAAB" box="[1013,1239,1353,1379]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 58­61, 63­67</figureCitation>
), especially the dentiform distal sclerites of internal sac (
<figureCitation id="729A2A17C336C768E3547160FBBDFA43" box="[882,1031,1393,1419]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
); and the shape of spermatheca (
<figureCitation id="729A2A17C336C768E15C7188FE6BFA7B" box="[378,465,1433,1459]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E71D3FC6DF9CB" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E71D3FDC9FA13" box="[312,627,1474,1499]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E71D3FDC9FA13" box="[312,627,1474,1499]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C336C768E3F771D3FBDAFA13" box="[977,1120,1474,1499]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C336C768E3F771D3FBDAFA13" box="[977,1120,1474,1499]" italics="true" pageId="24" pageNumber="25">S. nigriceps</emphasis>
</taxonomicName>
in having wider distal sclerites of internal sac of aedeagus (
<figureCitation id="729A2A17C336C768E2D971F8FC37F9CB" box="[767,909,1513,1539]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
; 124).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C76AE11E7203FDD5FE66" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="EA1E3692C336C768E11E7203FC86F99B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E7203FDCAF9E3" box="[312,624,1554,1579]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E7203FDCAF9E3" box="[312,624,1554,1579]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C336C768E3347203FC1EF9E3" box="[786,932,1554,1579]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C336C768E3347203FC1EF9E3" box="[786,932,1554,1579]" italics="true" pageId="24" pageNumber="25">S. liliputana</emphasis>
</taxonomicName>
in having larger median lobe of aedeagus with longer apex (
<figureCitation id="729A2A17C336C768E2767228FD63F99B" box="[592,729,1593,1619]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 58­61</figureCitation>
; 72­75).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7273FB5EF903" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E7273FDCBF9B3" box="[312,625,1634,1659]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E7273FDCBF9B3" box="[312,625,1634,1659]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from the similar species of
<taxonomicName id="2DA14D11C336C768E4067273FB14F9B3" box="[1056,1198,1634,1659]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C336C768E4067273FB14F9B3" box="[1056,1198,1634,1659]" italics="true" pageId="24" pageNumber="25">Seeversiella</emphasis>
</taxonomicName>
with long elytra and weak pronotal microsculpture in having strongly dentiform distal sclerites of internal sac (
<figureCitation id="729A2A17C336C768E1B872A0FD90F903" box="[414,554,1713,1739]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
) and L­shaped spermatheca with small umbilicus (
<figureCitation id="729A2A17C336C768E45872A0FB6CF903" box="[1150,1238,1713,1739]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E72C8FDB1F8D3" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E72C8FE6AF93B" bold="true" box="[312,464,1753,1779]" pageId="24" pageNumber="25">Description.</emphasis>
Length
<quantity id="2D599B77C336C768E21D72C8FD72F93B" box="[571,712,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="24" pageNumber="25" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
. Body brown, in some elytra and pronotum lighter, legs yellowish brown.
</paragraph>
<paragraph id="EA1E3692C336C769E11E7338FB84FE96" blockId="24.[264,1324,1073,1899]" lastBlockId="25.[264,1324,284,910]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">Head surface glossy, without microsculpture or with weak isodiametric microsculpture, with fine, weak and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Temples 1.7­2.1 times as long as eyes. Antennal article 2 longer than article 3, articles 4­10 transverse or strongly transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C337C769E11E757DFDAEFDBE" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C337C769E40D757DFB6CFE4E" box="[1067,1238,364,390]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="25" pageNumber="26" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
, length
<quantity id="2D599B77C337C769E12E7585FE09FE66" box="[264,435,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.5999999999999996" pageId="25" pageNumber="26" unit="mm" value="0.395" valueMax="0.43" valueMin="0.36">0.36­0.43 mm</quantity>
, width to length ratio 1.1, surface glossy, without microsculpture; punctation as on head. Elytra wider than pronotum and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.3 times wider than long, glossy, with weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings fully developed, 3­4 times as long as elytra.
</paragraph>
<paragraph id="EA1E3692C337C769E11E7695FCA7FCDE" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (54, 56).</paragraph>
<paragraph id="EA1E3692C337C769E11E7735FECCFC46" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">
In males posterior angles of tergum 3 not projecting, tergum 7 without medial carina. Posterior margin of male tergum 8 slightly concave medially, but without emargination (
<figureCitation id="729A2A17C337C769E1377765FED3FC46" box="[273,361,884,910]" captionStart="FIGURES 54 ­ 57" captionStartId="25.[264,383,1800,1824]" captionTargetBox="[322,1265,1015,1769]" captionTargetId="figure@25.[313,1274,1006,1776]" captionTargetPageId="25" captionText="FIGURES 54 ­ 57. Abdominal segment 8 of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 54 male tergum 8; 55 male sternum 8; 56 female tergum 8; 57 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156431/files/figure.png" pageId="25" pageNumber="26">Fig. 54</figureCitation>
).
</paragraph>
<caption id="BEDE661AC337C769E12E7319FD2DF8AD" httpUri="https://zenodo.org/record/156431/files/figure.png" pageId="25" pageNumber="26" targetBox="[322,1265,1015,1769]" targetPageId="25">
<paragraph id="EA1E3692C337C769E12E7319FD2DF8AD" blockId="25.[264,1323,1800,1893]" pageId="25" pageNumber="26">
<emphasis id="D8D5EA80C337C769E12E7319FE7EF8E8" bold="true" box="[264,452,1800,1824]" pageId="25" pageNumber="26">FIGURES 54­57</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C337C769E2CB7318FBD1F8D7" authority="Gusarov" authorityName="Gusarov" box="[749,1131,1800,1824]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C337C769E2CB7318FBB3F8E8" box="[749,1033,1801,1824]" italics="true" pageId="25" pageNumber="26">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C337C769E4517319FB0EF8E8" bold="true" box="[1143,1204,1800,1824]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="C3E657FBC337C769E4517319FB0EF8E8" box="[1143,1204,1800,1824]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Carr Canyon, Arizona). 54 male tergum 8; 55 male sternum 8; 56 female tergum 8; 57 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C334C76AE11E750DFE8CFE4E" blockId="26.[264,1324,284,550]" pageId="26" pageNumber="27">
Aedeagus as in
<figureCitation id="729A2A17C334C76AE1D7750DFD77FEFE" box="[497,717,284,310]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Figs. 58­61, 63­67</figureCitation>
. Distal sclerites of internal sac dentiform (
<figureCitation id="729A2A17C334C76AE4E2750DFE93FE96" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Figs. 63, 65</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C334C76AE4DA7555FE93FE4E" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C334C76AE11E7585FDD5FE66" blockId="26.[264,1324,284,550]" box="[312,623,404,430]" pageId="26" pageNumber="27">
Spermatheca as in
<figureCitation id="729A2A17C334C76AE2327585FDD1FE66" box="[532,619,404,430]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Fig. 62</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C334C76AE11E75AAFDCDF8A0" pageId="26" pageNumber="27" type="distribution">
<paragraph id="EA1E3692C334C76AE11E75AAFB20FE1E" blockId="26.[264,1324,284,550]" box="[312,1178,443,470]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE11E75AAFE62FE1D" bold="true" box="[312,472,443,469]" pageId="26" pageNumber="27">Distribution.</emphasis>
Known from the Huachuca Mountains, Arizona (
<figureCitation id="729A2A17C334C76AE40075ADFB34FE1E" box="[1062,1166,444,470]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="26" pageNumber="27">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C334C76AE11E75F2FE32FDEE" blockId="26.[264,1324,284,550]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE11E75F2FDBEFE35" bold="true" box="[312,516,483,509]" pageId="26" pageNumber="27">Natural History.</emphasis>
<taxonomicName id="2DA14D11C334C76AE23575F5FCF0FE35" box="[531,842,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C334C76AE23575F5FCF0FE35" box="[531,842,484,509]" italics="true" pageId="26" pageNumber="27">Seeversiella sonomotoides</emphasis>
</taxonomicName>
was collected in oak leaf litter at altitude of
<quantity id="2D599B77C334C76AE10E761DFE3EFDEE" box="[296,388,524,550]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="27" unit="m" value="2150.0">2150 m</quantity>
.
</paragraph>
<caption id="BEDE661AC334C76AE12E726EFC27F88C" httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27" targetBox="[289,1302,621,1628]" targetPageId="26">
<paragraph id="EA1E3692C334C76AE12E726EFC27F88C" blockId="26.[264,1324,1663,1896]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE12E726EFE7DF95F" bold="true" box="[264,455,1663,1687]" pageId="26" pageNumber="27">FIGURES 58­67</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C334C76AE24A7291FC54F95F" authority="Gusarov" authorityName="Gusarov" box="[620,1006,1664,1687]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C334C76AE24A7291FC30F95F" box="[620,906,1664,1687]" italics="true" pageId="26" pageNumber="27">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C334C76AE3DA726EFB81F95F" bold="true" box="[1020,1083,1663,1687]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="C3E657FBC334C76AE3DA726EFB81F95F" box="[1020,1083,1663,1687]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Carr Canyon, Arizona). 58 median lobe, parameral view; 59 apex of median lobe, parameral view; 60 median lobe, lateral view; 61 apex of median lobe, lateral view; 62 spermatheca; 63 details of internal sac retracted into median lobe, lateral view; 64 apex of left paramere; 65 right hook of internal sac; 66 copulatory piece, lateral view; 67 everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61­62, 64­66).
</paragraph>
</caption>
<paragraph id="EA1E3692C334C76AE12E7340FDCDF8A0" blockId="26.[264,1324,1663,1896]" box="[264,631,1873,1896]" pageId="26" pageNumber="27">DS distal sclerite of internal sac.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,255 @@
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<subSubSection id="A2BB6519C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22" type="key">
<paragraph id="EA1E3692C33BC765E12E75F2FCCBFE35" blockId="21.[264,881,483,510]" box="[264,881,483,510]" pageId="21" pageNumber="22">
<heading id="B15681FEC33BC765E12E75F2FCCBFE35" bold="true" box="[264,881,483,510]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<key id="9E3061A3C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
<keyStep id="51552E37C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E75F2FCCBFE35" bold="true" box="[264,881,483,510]" pageId="21" pageNumber="22">
<keyLead id="515095A7C33BC765E12E75F2FCCBFE35" box="[264,881,483,510]" pageId="21" pageNumber="22">
2.
<taxonomicName id="2DA14D11C33BC765E10075F5FD3AFE35" ID-CoL="6YBBD" authority="Gusarov" authorityName="Gusarov" box="[294,640,483,510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C33BC765E10075F5FDB7FE36" bold="true" box="[294,525,484,510]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC33BC765E2AB75F2FD6AFE35" box="[653,720,483,509]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C33BC765E2F975F2FCD0FE35" box="[735,874,483,509]" captionStart-0="FIGURES 37 ­ 40" captionStart-1="FIGURES 41 ­ 44" captionStart-2="FIGURES 45 ­ 53" captionStartId-0="21.[264,383,1773,1797]" captionStartId-1="22.[264,383,1298,1322]" captionStartId-2="23.[264,383,1314,1338]" captionTargetBox-0="[274,1312,896,1738]" captionTargetBox-1="[293,1296,576,1260]" captionTargetBox-2="[313,1274,323,1277]" captionTargetId-0="figure@21.[264,1323,884,1747]" captionTargetId-1="figure@22.[281,1306,569,1271]" captionTargetId-2="figure@23.[306,1280,313,1287]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-2="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156428/files/figure.png" httpUri-1="https://zenodo.org/record/156429/files/figure.png" httpUri-2="https://zenodo.org/record/156430/files/figure.png" pageId="21" pageNumber="22">Figs. 37­53</figureCitation>
)
</keyLead>
</emphasis>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C33BC765E12E763FFD6EFCD9" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="EA1E3692C33BC765E12E763FFC83FD51" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E763FFE7BFD80" bold="true" box="[264,449,558,584]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E12E763FFEFFFD80" box="[264,325,558,584]" pageId="21" pageNumber="22">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C33BC765E1F3763EFDFFFD81" box="[469,581,559,585]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C33BC765E25E763FFC75FD80" bold="true" box="[632,975,558,584]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E25E763FFCDCFD80" box="[632,870,558,584]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<collectingCountry id="92B67602C33BC765E48A763EFA91FD81" box="[1196,1323,559,585]" name="Guadeloupe" pageId="21" pageNumber="22">Guadalupe</collectingCountry>
Mts. N. P., The Bowl, jct. Tejas Tr. and Juniper Tr.,
<quantity id="2D599B77C33BC765E3457646FC05FDB9" box="[867,959,599,625]" metricMagnitude="3" metricUnit="m" metricValue="2.407" pageId="21" pageNumber="22" unit="m" value="2407.0">2407 m</quantity>
, berlese maple, oak, pine, ironwood litter (R.Anderson),
<date id="9E1F1052C33BC765E219766EFD7FFD51" box="[575,709,639,665]" pageId="21" pageNumber="22" value="1989-07-31">31.vii.1989</date>
(
<collectionCode id="8CB0AE57C33BC765E2F2766EFC96FD51" box="[724,812,639,665]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C33BC765E11E76B6FD6EFCD9" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E11E76B6FE15FD09" box="[312,431,679,705]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C33BC765E1E476B7FCA9FD08" bold="true" box="[450,787,678,704]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E1E476B7FD17FD08" box="[450,685,678,704]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<specimenCount id="FCA7FD1BC33BC765E3C076B6FB33FD09" box="[998,1161,679,705]" pageId="21" pageNumber="22" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C33BC765E11176DEFE1BFD21" box="[311,417,719,745]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC33BC765E19676DEFDFCFD21" box="[432,582,719,745]" pageId="21" pageNumber="22" type="generic">4 specimens</specimenCount>
, ditto but mixed hardwood litter;
<specimenCount id="FCA7FD1BC33BC765E3C276DEFBC0FD21" box="[996,1146,719,745]" pageId="21" pageNumber="22" type="generic">8 specimens</specimenCount>
, ditto but oak, conifer litter,
<date id="9E1F1052C33BC765E18176E6FD9FFCD9" box="[423,549,759,785]" pageId="21" pageNumber="22" value="1988-09-13">13.ix.1988</date>
(all
<collectionCode id="8CB0AE57C33BC765E25676E6FD7DFCD9" box="[624,711,759,785]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC33BC765E12E72FCFC32F882" httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="21" pageNumber="22" targetBox="[274,1312,896,1738]" targetPageId="21">
<paragraph id="EA1E3692C33BC765E12E72FCFC32F882" blockId="21.[264,1323,1773,1866]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E72FCFE7FF8CD" bold="true" box="[264,453,1773,1797]" pageId="21" pageNumber="22">FIGURES 37­40</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C33BC765E2D072FFFB90F8CD" authority="Gusarov" authorityName="Gusarov" box="[758,1066,1774,1797]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C33BC765E2D072FFFC7DF8CD" box="[758,967,1774,1797]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C33BC765E41E72FCFBCCF8CD" bold="true" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="C3E657FBC33BC765E41E72FCFBCCF8CD" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C338C766E11E750AFC9CF969" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="EA1E3692C338C766E11E750AFD8AFE36" blockId="22.[264,1323,283,510]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C338C766E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E4F5750DFEFDFE95" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
by having brown or brownish yellow body; small eyes (temples 2.1­2.7 times as long as eyes); glossy pronotum with weak microsculpture; wings reduced, 2 times as long as elytra; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C338C766E4E77585FE92FE1E" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 41­ 52</figureCitation>
), especially the hook­shaped distal sclerites of internal sac (
<figureCitation id="729A2A17C338C766E3C175ADFBCCFE1E" box="[999,1142,444,470]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 48, 51</figureCitation>
); and the shape of spermatheca (
<figureCitation id="729A2A17C338C766E1EC75F5FD98FE36" box="[458,546,484,510]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
<caption id="BEDE661AC338C766E12E7103FDCFFA59" httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23" targetBox="[293,1296,576,1260]" targetPageId="22">
<paragraph id="EA1E3692C338C766E12E7103FDCFFA59" blockId="22.[264,1323,1298,1425]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E12E7103FE7DFAE2" bold="true" box="[264,455,1298,1322]" pageId="22" pageNumber="23">FIGURES 41­44</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C338C766E2527103FC10FAE1" authority="Gusarov" authorityName="Gusarov" box="[628,938,1298,1321]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C338C766E2527103FCFFFAE1" box="[628,837,1298,1321]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C338C766E39E7103FC4CFAE2" bold="true" box="[952,1014,1298,1322]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="C3E657FBC338C766E39E7103FC4CFAE2" box="[952,1014,1298,1322]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44).
</paragraph>
</caption>
<paragraph id="EA1E3692C338C766E11E71F9FC9CF969" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" italics="true" pageId="22" pageNumber="23">S. globicollis</emphasis>
</taxonomicName>
in having lighter body colour; more distinct microsculpture of head and pronotum; elytra shorter than pronotum; reduced wings, 2 times as long as elytra; apex of median lobe bent paramerally (in lateral view) (
<figureCitation id="729A2A17C338C766E17B724EFE53F9B1" box="[349,489,1631,1657]" captionStart="FIGURES 41 ­ 44" captionStartId="22.[264,383,1298,1322]" captionTargetBox="[293,1296,576,1260]" captionTargetId="figure@22.[281,1306,569,1271]" captionTargetPageId="22" captionText="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23">Figs. 43­44</figureCitation>
; 26­27), shorter copulatory piece of internal sac (
<figureCitation id="729A2A17C338C766E41B724EFB42F9B1" box="[1085,1272,1631,1657]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 46, 49­50</figureCitation>
; 29, 34) and spermatheca with umbilicus (
<figureCitation id="729A2A17C338C766E2E77296FCA3F969" box="[705,793,1671,1697]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C338C768E11E72A1FDD5FCF6" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="EA1E3692C338C766E11E72A1FBF5F8D1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from the other similar species of
<taxonomicName id="2DA14D11C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
with long elytra and weak pronotal microsculpture in having strongly sclerotized hook­shaped distal sclerites of internal sac and J­shaped spermatheca with large umbilicus.
</paragraph>
<paragraph id="EA1E3692C338C766E11E7336FB32F8A1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E7336FE6AF889" bold="true" box="[312,464,1831,1857]" pageId="22" pageNumber="23">Description.</emphasis>
Length
<quantity id="2D599B77C338C766E2677336FD75F889" box="[577,719,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="22" pageNumber="23" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Body from brownish yellow to brown, head and abdominal segments 6­7 often darker, legs and mouthparts brownish yellow.
</paragraph>
<caption id="BEDE661AC339C767E12E7133FDC5FA2E" httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="23" pageNumber="24" targetBox="[313,1274,323,1277]" targetPageId="23">
<paragraph id="EA1E3692C339C767E12E7133FDC5FA2E" blockId="23.[264,1323,1314,1545]" pageId="23" pageNumber="24">
<emphasis id="D8D5EA80C339C767E12E7133FE7DFAF2" bold="true" box="[264,455,1314,1338]" pageId="23" pageNumber="24">FIGURES 45­53</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C339C767E2497133FC1DFAF1" authority="Gusarov" authorityName="Gusarov" box="[623,935,1314,1337]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C339C767E2497133FCFBFAF1" box="[623,833,1314,1337]" italics="true" pageId="23" pageNumber="24">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C339C767E3907133FC4FFAF2" bold="true" box="[950,1013,1314,1338]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="C3E657FBC339C767E3907133FC4FFAF2" box="[950,1013,1314,1338]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48­49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48­49, 53), 0.1 mm (46­47, 50­52).
</paragraph>
</caption>
<paragraph id="EA1E3692C339C767E12E71E3FDCDF9C1" blockId="23.[264,1323,1314,1545]" box="[264,631,1522,1545]" pageId="23" pageNumber="24">DS distal sclerite of internal sac.</paragraph>
<paragraph id="EA1E3692C339C767E11E7274FBD5F93F" blockId="23.[264,1324,1637,1903]" pageId="23" pageNumber="24">
Head surface glossy, with weak isodiametric microsculpture, with fine, weak and poorly visible (at
<date id="9E1F1052C339C767E1F0729CFDBEF96F" box="[470,516,1677,1703]" pageId="23" pageNumber="24" value="1970-10">70x</date>
) punctation, distance between punctures equals 2­3 times their diameter. Temples 2.1­2.7 times as long as eyes. Antennal article 2 longer than article 3, article 4 slightly transverse, 5­10 transverse or strongly transverse (ratio 1.5­2.0).
</paragraph>
<paragraph id="EA1E3692C339C768E11E7314FE14FE66" blockId="23.[264,1324,1637,1903]" lastBlockId="24.[264,1324,284,910]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C339C767E40D7314FB6CF8D7" box="[1067,1238,1797,1823]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="23" pageNumber="24" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44­0.49 mm</quantity>
, length
<quantity id="2D599B77C339C767E12E733CFE0FF88F" box="[264,437,1837,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="23" pageNumber="24" unit="mm" value="0.435" valueMax="0.46" valueMin="0.41">0.41­0.46 mm</quantity>
, width to length ratio 1.1, surface glossy, with weak isodiametric microsculpture; punctation as on head. Elytra wider and shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.1), 1.4 times wider than long, glossy, with weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings reduced, 2 times as long as elytra.
</paragraph>
<paragraph id="EA1E3692C336C768E11E75ADFCA7FD86" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (37, 39).</paragraph>
<paragraph id="EA1E3692C336C768E11E764DFA92FD0E" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
In males posterior angles of tergum 3 projecting as spines up to half as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to half as long as tergum length. Posterior margin of male tergum 8 without emargination (
<figureCitation id="729A2A17C336C768E4E576BDFAA0FD0E" box="[1219,1306,684,710]" captionStart="FIGURES 37 ­ 40" captionStartId="21.[264,383,1773,1797]" captionTargetBox="[274,1312,896,1738]" captionTargetId="figure@21.[264,1323,884,1747]" captionTargetPageId="21" captionText="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 male tergum 8; 38 male sternum 8; 39 female tergum 8; 40 female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="24" pageNumber="25">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E76C5FAA7FCDE" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
Aedeagus as in
<figureCitation id="729A2A17C336C768E1C976C5FDC3FD26" box="[495,633,724,750]" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 median lobe, parameral view; 42 apex of median lobe, parameral view; 43 median lobe, lateral view; 44 apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 41­52</figureCitation>
. Distal sclerites of internal sac hook­shaped (
<figureCitation id="729A2A17C336C768E4B676C5FAA5FD26" box="[1168,1311,724,750]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 48, 51</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C336C768E49E76EDFAAAFCDE" box="[1208,1296,764,790]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7735FDD5FCF6" blockId="24.[264,1324,284,910]" box="[312,623,804,830]" pageId="24" pageNumber="25">
Spermatheca as in
<figureCitation id="729A2A17C336C768E2337735FDD6FCF6" box="[533,620,804,830]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 details of internal sac retracted into median lobe, abparameral view; 46 copulatory piece, parameral view; 47 medial lamellae, parameral view; 48 ­ 49 details of internal sac retracted into median lobe, lateral view; 50 copulatory piece, lateral view; 51 right hook of internal sac; 52 apex of left paramere; 53 spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E11E775AFA96FC46" pageId="24" pageNumber="25" type="distribution">
<paragraph id="EA1E3692C336C768E11E775AFB31FCAE" blockId="24.[264,1324,284,910]" box="[312,1163,843,870]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E775AFE62FCAD" bold="true" box="[312,472,843,869]" pageId="24" pageNumber="25">Distribution.</emphasis>
Known from the
<collectingCountry id="92B67602C336C768E296775DFC95FCAE" box="[688,815,844,870]" name="Guadeloupe" pageId="24" pageNumber="25">Guadalupe</collectingCountry>
Mountains, Texas (
<figureCitation id="729A2A17C336C768E430775DFBC4FCAE" box="[1046,1150,844,870]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="24" pageNumber="25">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7762FA96FC46" blockId="24.[264,1324,284,910]" box="[312,1324,883,910]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E7762FDBFFC45" bold="true" box="[312,517,883,909]" pageId="24" pageNumber="25">Natural History.</emphasis>
<taxonomicName id="2DA14D11C336C768E2357765FD4CFC45" box="[531,758,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C336C768E2357765FD4CFC45" box="[531,758,884,909]" italics="true" pageId="24" pageNumber="25">Seeversiella texana</emphasis>
</taxonomicName>
was collected at altitude of
<quantity id="2D599B77C336C768E41B7765FB2CFC46" box="[1085,1174,884,910]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="25" unit="m" value="2400.0">2400 m</quantity>
in leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,231 @@
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<mods:title id="21FE1559D26C65FED59C6C55E033A700">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79" type="key">
<paragraph id="EA1E3692C360C73EE12E727AFC7FF94D" blockId="78.[264,965,1643,1670]" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<heading id="B15681FEC360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" fontSize="11" level="1" pageId="78" pageNumber="79" reason="1">
<emphasis id="D8D5EA80C360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<key id="9E3061A3C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<keyStep id="51552E37C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<keyLead id="515095A7C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
23.
<taxonomicName id="2DA14D11C360C73EE112727DFD0DF94D" ID-CoL="6YBBC" authority="Gusarov" authorityName="Gusarov" box="[308,695,1643,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C360C73EE112727DFDFEF94E" bold="true" box="[308,580,1644,1670]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC360C73EE2E5727AFCBCF94D" box="[707,774,1643,1669]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C360C73EE333727AFC06F94D" box="[789,956,1643,1669]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="78" pageNumber="79">Figs. 321­330</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C720E12E72A7FC86F8D7" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="EA1E3692C360C73EE12E72A7FECFF8E8" blockId="78.[264,1323,1718,1904]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE12E72A7FE04F918" bold="true" box="[264,446,1718,1744]" pageId="78" pageNumber="79">
<typeStatus id="351A8830C360C73EE12E72A7FEFFF918" box="[264,325,1718,1744]" pageId="78" pageNumber="79">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C360C73EE1EA72A7FD86F918" box="[460,572,1718,1744]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C360C73EE24C72A7FBA6F918" bold="true" box="[618,1052,1718,1744]" pageId="78" pageNumber="79">
<collectingCountry id="92B67602C360C73EE24C72A7FC99F918" box="[618,803,1718,1744]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C360C73EE1D972CFFDC1F930" box="[511,635,1758,1784]" direction="north" orientation="latitude" pageId="78" pageNumber="79" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C360C73EE2A472CFFCA8F930" box="[642,786,1758,1784]" direction="west" orientation="longitude" pageId="78" pageNumber="79" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C360C73EE33872CFFCC0F930" box="[798,890,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="78" pageNumber="79" unit="m" value="3300.0">3300 m</quantity>
, forest litter (R.Anderson),
<date id="9E1F1052C360C73EE49A72CFFA90F930" box="[1212,1322,1758,1784]" pageId="78" pageNumber="79" value="1997-06-08">8.vi.1997</date>
(
<collectionCode id="8CB0AE57C360C73EE1367317FED2F8E8" box="[272,360,1798,1824]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C360C73FE11E733FFD83FE1E" blockId="78.[264,1323,1718,1904]" lastBlockId="79.[264,1324,284,470]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<typeStatus id="351A8830C360C73EE11E733FFE14F880" box="[312,430,1838,1864]" pageId="78" pageNumber="79" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C360C73EE1E6733FFD44F880" bold="true" box="[448,766,1838,1864]" pageId="78" pageNumber="79">
<collectingCountry id="92B67602C360C73EE1E6733FFDC0F880" box="[448,634,1838,1864]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, same data as the
<typeStatus id="351A8830C360C73EE3D0733FFBE7F880" box="[1014,1117,1838,1864]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C360C73EE44A733FFB7EF880" box="[1132,1220,1838,1864]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
<emphasis id="D8D5EA80C360C73EE11E7347FE02F8B8" bold="true" box="[312,440,1878,1904]" pageId="78" pageNumber="79">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C360C73FE4F57346FEFDFEFD" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" lastPageId="79" lastPageNumber="80" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C360C73FE4F57346FEFDFEFD" italics="true" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">Seeversiella</emphasis>
</taxonomicName>
by having reddish brown body; temples 1.8­2.3 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by having small medial emargination delimited by two short and blunt projections at the posterior margin of the male tergum 8 (
<figureCitation id="729A2A17C361C73FE1AE7585FE49FE66" box="[392,499,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Fig. 330</figureCitation>
); tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C361C73FE1A575ADFD96FE1E" box="[387,556,444,470]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Figs. 321­328</figureCitation>
).
</paragraph>
<caption id="BEDE661AC361C73FE12E729EFB76F923" httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="79" pageNumber="80" targetBox="[283,1308,537,1644]" targetPageId="79">
<paragraph id="EA1E3692C361C73FE12E729EFB76F923" blockId="79.[264,1322,1679,1771]" pageId="79" pageNumber="80">
<emphasis id="D8D5EA80C361C73FE12E729EFE67F96F" bold="true" box="[264,477,1679,1703]" pageId="79" pageNumber="80">FIGURES 321­324</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C361C73FE25B729EFC68F96E" authority="Gusarov" authorityName="Gusarov" box="[637,978,1679,1702]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C361C73FE25B729EFCCAF96E" box="[637,880,1679,1702]" italics="true" pageId="79" pageNumber="80">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C361C73FE3FA729EFBADF96F" bold="true" box="[988,1047,1679,1703]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="C3E657FBC361C73FE3FA729EFBADF96F" box="[988,1047,1679,1703]" pageId="79" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324).
</paragraph>
</caption>
<caption id="BEDE661AC37EC720E12E7229FE10F934" httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="80" pageNumber="81" targetBox="[280,1306,336,1561]" targetPageId="80">
<paragraph id="EA1E3692C37EC720E12E7229FE10F934" blockId="80.[264,1323,1592,1823]" pageId="80" pageNumber="81">
<emphasis id="D8D5EA80C37EC720E12E7229FE67F998" bold="true" box="[264,477,1592,1616]" pageId="80" pageNumber="81">FIGURES 325­330</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C37EC720E3B77229FB5FF987" authority="Gusarov" authorityName="Gusarov" box="[913,1253,1592,1615]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C37EC720E3B77229FB3EF987" box="[913,1156,1592,1615]" italics="true" pageId="80" pageNumber="81">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37EC720E4C97229FA91F998" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81" rank="species">
<emphasis id="D8D5EA80C37EC720E4C97229FA91F998" bold="true" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81">sp. n.</emphasis>
</taxonomicNameLabel>
(325, 327­330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326­328), 0.4 mm (329­330).
</paragraph>
</caption>
<paragraph id="EA1E3692C37EC720E12E7319FC86F8D7" blockId="80.[264,1323,1592,1823]" box="[264,828,1800,1823]" pageId="80" pageNumber="81">SLD sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC721E11E750DFDB1F8EE" pageId="81" pageNumber="82" type="description">
<paragraph id="EA1E3692C37FC721E11E750DFE76FDEE" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" italics="true" pageId="81" pageNumber="82">S. impressicollis</emphasis>
</taxonomicName>
in having larger body; in males pronotum with medial furrow, lateral portions of posterior margin of tergum 3 extending as short and obtuse projections (as in
<figureCitation id="729A2A17C37FC721E298757DFCA9FE4E" box="[702,787,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
), tergum 7 with short medial carina; posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E3647585FC11FE66" box="[834,939,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E2C375ADFC28FE1E" box="[741,914,444,470]" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­322</figureCitation>
); copulatory piece of internal sac longer (
<figureCitation id="729A2A17C37FC721E14175F5FDA6FE36" box="[359,540,484,510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326, 328</figureCitation>
; 317); sclerites of lateral diverticula of internal sac more robust (
<figureCitation id="729A2A17C37FC721E137761DFE38FDEE" box="[273,386,524,550]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326</figureCitation>
; 318).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7625FEFFFD26" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C77625FC08FD85" box="[737,946,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37FC721E2C77625FC08FD85" box="[737,946,564,589]" italics="true" pageId="81" pageNumber="82">S. microphthalma</emphasis>
</taxonomicName>
in having more asperate pronotal punctation; in males pronotum with medial furrow, posterior angles of tergum 3 not projecting as spines, posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E4947695FAA6FD56" box="[1202,1308,644,670]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); and the apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E48976BDFE82FD26" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­ 322</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E76EAFC1AFCDE" blockId="81.[264,1324,284,1870]" box="[312,928,763,790]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E76EAFE6AFCDD" bold="true" box="[312,464,763,789]" pageId="81" pageNumber="82">Description.</emphasis>
Length 3.0 mm. Body reddish brown.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7735FDB1FC7E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 1.8­2.3 times as long as eyes. Antennal article 2 as long as 3, articles 4­6 subquadrate, 7­ 10 slightly transverse.</paragraph>
<paragraph id="EA1E3692C37FC721E11E77D5FCD3FB3E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C37FC721E40D77D5FB6CFC16" box="[1067,1238,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="81" pageNumber="82" unit="mm" value="0.665" valueMax="0.67" valueMin="0.66">0.66­0.67 mm</quantity>
, length
<quantity id="2D599B77C37FC721E12E77FDFE0BFBCE" box="[264,433,1004,1030]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.0" metricValueMin="5.8999999999999995" pageId="81" pageNumber="82" unit="mm" value="0.595" valueMax="0.6" valueMin="0.59">0.59­0.60 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­1.5 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7115FDD5FA2E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 1­4 times their diameter; tergum 7 with microsculpture consisting of isodiametric or slightly transverse meshes and with fine and sparse punctation, distance between punctures equals 3­6 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C37FC721E1DD71DDFDD9FA2E" box="[507,611,1484,1510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E71E5FD43F966" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
In males pronotum with medial furrow; lateral portions of posterior margin of tergum 3 extending as short and blunt projections up to 1/8 as long as tergum (measured medially) (as in
<figureCitation id="729A2A17C37FC721E1777255FE1DF996" box="[337,423,1604,1630]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
); tergum
<quantity id="2D599B77C37FC721E23F7255FDFDF996" box="[537,583,1604,1630]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="81" pageNumber="82" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina, up to 1/4 as long as the tergum; posterior margin of tergum 8 with small medial emargination delimited by two short and blunt projections (
<figureCitation id="729A2A17C37FC721E2A37285FD57F966" box="[645,749,1684,1710]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E72ADFD48F936" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Aedeagus as in
<figureCitation id="729A2A17C37FC721E1C872ADFD2EF91E" box="[494,660,1724,1750]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 median lobe, parameral view; 322 apex of median lobe, parameral view; 323 median lobe, lateral view; 324 apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 321­328</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C37FC721E25872F5FD5FF936" box="[638,741,1764,1790]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 apex of left paramere; 326 everted internal sac, parameral view; 327 details of internal sac retracted into median lobe, abparameral view; 328 details of internal sac retracted into median lobe, lateral view; 329 male sternum 8; 330 male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 325</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E731DFDB1F8EE" blockId="81.[264,1324,284,1870]" box="[312,523,1804,1830]" pageId="81" pageNumber="82">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC722E11E7322FED2FE96" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="distribution">
<paragraph id="EA1E3692C37FC721E11E7322FBCDF886" blockId="81.[264,1324,284,1870]" box="[312,1143,1843,1870]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E7322FE62F885" bold="true" box="[312,472,1843,1869]" pageId="81" pageNumber="82">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C37FC721E3547325FC49F886" box="[882,1011,1844,1870]" name="Costa Rica" pageId="81" pageNumber="82">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C37FC721E4257325FBD0F886" box="[1027,1130,1844,1870]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="81" pageNumber="82">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37CC722E11E750AFED2FE96" blockId="82.[264,1323,283,350]" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E11E750AFDB2FEFD" bold="true" box="[312,520,283,309]" pageId="82" pageNumber="83">Natural History.</emphasis>
<taxonomicName id="2DA14D11C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" italics="true" pageId="82" pageNumber="83">Seeversiella sulcicollis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C37CC722E12E7555FEDEFE96" box="[264,356,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="82" pageNumber="83" unit="m" value="3300.0">3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,238 @@
<document id="24DD095DE3D4B50DC20A29F21A463C5F" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C362C73EE12E750AFE5DF9C6" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella impressicollis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="79" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="77" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="43CB2D7B546D52CF95DD091CA37CFE25">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77" type="key">
<paragraph id="EA1E3692C362C73CE12E750AFC4AFEFD" blockId="76.[264,1008,283,310]" box="[264,1008,283,310]" pageId="76" pageNumber="77">
<heading id="B15681FEC362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="D8D5EA80C362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" pageId="76" pageNumber="77">
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22.
<taxonomicName id="2DA14D11C362C73CE112750DFD58FEFD" ID-CoL="6YBCW" authority="Gusarov" authorityName="Gusarov" box="[308,738,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C362C73CE112750DFDCAFEFE" bold="true" box="[308,624,284,310]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC362C73CE2C9750AFC8BFEFD" box="[751,817,283,309]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C362C73CE366750AFC52FEFD" box="[832,1000,283,309]" captionStart-0="FIGURES 307 ­ 310" captionStart-1="FIGURES 311 ­ 320" captionStartId-0="76.[264,383,1792,1816]" captionStartId-1="77.[264,383,1623,1647]" captionTargetBox-0="[290,1292,893,1759]" captionTargetBox-1="[286,1302,571,1605]" captionTargetId-0="figure@76.[282,1305,884,1766]" captionTargetId-1="figure@77.[276,1310,569,1613]" captionTargetPageId-0="76" captionTargetPageId-1="77" captionText-0="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri-0="https://zenodo.org/record/156474/files/figure.png" httpUri-1="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 307­320</figureCitation>
)
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</key>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A2BB6519C362C73CE12E7577FECFFDE9" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="EA1E3692C362C73CE12E7577FECFFE19" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="76" pageNumber="77">
<typeStatus id="351A8830C362C73CE12E7577FEF9FE48" box="[264,323,358,384]" pageId="76" pageNumber="77">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C362C73CE1F17576FDFDFE49" box="[471,583,359,385]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C362C73CE25D7577FC79FE48" bold="true" box="[635,963,358,384]" pageId="76" pageNumber="77">
<collectingCountry id="92B67602C362C73CE25D7577FC82FE48" box="[635,824,358,384]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Cerro Chirripó, Valle de los Conelos,
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,
<quantity id="2D599B77C362C73CE2B9759EFD44FE61" box="[671,766,399,425]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="76" pageNumber="77" unit="m" value="3600.0">3600 m</quantity>
, paramo shrub litter (R.Anderson),
<date id="9E1F1052C362C73CE48B759EFA90FE61" box="[1197,1322,399,425]" pageId="76" pageNumber="77" value="1999-06-26">26.vi.1999</date>
(
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).
</paragraph>
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<typeStatus id="351A8830C362C73CE11E75CEFE15FE31" box="[312,431,479,505]" pageId="76" pageNumber="77" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C362C73CE1ED75CFFD33FE30" box="[459,649,478,504]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC362C73CE31775CEFC62FE31" box="[817,984,479,505]" pageId="76" pageNumber="77" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C362C73CE4E575CEFA90FE31" box="[1219,1322,479,505]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
(
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).
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</subSubSection>
<subSubSection id="A2BB6519C362C73EE11E763FFE3BFE1E" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="EA1E3692C362C73CE11E763FFD78FCD9" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE11E763FFE0DFD80" bold="true" box="[312,439,558,584]" pageId="76" pageNumber="77">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
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<emphasis id="D8D5EA80C362C73CE4DE763EFED7FDB8" italics="true" pageId="76" pageNumber="77">Seeversiella</emphasis>
</taxonomicName>
by having brown or dark brown body with lighter pronotum and elytra; temples 1.8­2.2 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of male tergum 7 (
<figureCitation id="729A2A17C362C73CE13776DEFECDFD21" box="[273,375,719,745]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77">Fig. 307</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C362C73CE4D676DEFED0FCD9" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 311­319</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C362C73CE26876E6FD0CFCD9" box="[590,694,759,785]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Fig. 320</figureCitation>
).
</paragraph>
<caption id="BEDE661AC362C73CE12E7311FC2CF895" httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77" targetBox="[290,1292,893,1759]" targetPageId="76">
<paragraph id="EA1E3692C362C73CE12E7311FC2CF895" blockId="76.[264,1323,1792,1885]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7311FE52F8D0" bold="true" box="[264,488,1792,1816]" pageId="76" pageNumber="77">FIGURES 307­310</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C362C73CE3647310FB6BF8D0" authority="Gusarov" authorityName="Gusarov" box="[834,1233,1793,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C362C73CE3647310FBDCF8D0" box="[834,1126,1793,1816]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C362C73CE4C37311FA91F8D0" bold="true" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="C3E657FBC362C73CE4C37311FA91F8D0" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE11E750DFD43FE36" blockId="77.[264,1323,284,510]" pageId="77" pageNumber="78">
<taxonomicName id="2DA14D11C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" italics="true" pageId="77" pageNumber="78">S. sulcicollis</emphasis>
</taxonomicName>
in having smaller body; in males pronotum with broad medial impression, posterior margin of tergum 3 not projecting, tergum 7 without medial carina and posterior margin of tergum 8 without medial denticles (
<figureCitation id="729A2A17C363C73DE1377585FEC2FE66" box="[273,376,404,430]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="77" pageNumber="78">Fig. 307</figureCitation>
); apex of median lobe in parameral view without carinae (
<figureCitation id="729A2A17C363C73DE40D7585FB69FE66" box="[1067,1235,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 311­312</figureCitation>
); copulatory piece of internal sac shorter (
<figureCitation id="729A2A17C363C73DE28975ADFC99FE1E" box="[687,803,444,470]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 317</figureCitation>
; 326, 328); sclerites of lateral diverticula of internal sac more slender (
<figureCitation id="729A2A17C363C73DE21875F5FD15FE36" box="[574,687,484,510]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 318</figureCitation>
; 326).
</paragraph>
<caption id="BEDE661AC363C73DE12E7246FEFAF8F7" httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78" targetBox="[286,1302,571,1605]" targetPageId="77">
<paragraph id="EA1E3692C363C73DE12E7246FEFAF8F7" blockId="77.[264,1323,1623,1891]" pageId="77" pageNumber="78">
<emphasis id="D8D5EA80C363C73DE12E7246FE67F9A7" bold="true" box="[264,477,1623,1647]" pageId="77" pageNumber="78">FIGURES 311­320</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C363C73DE2537249FC4BF9A7" authority="Gusarov" authorityName="Gusarov" box="[629,1009,1624,1647]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C363C73DE2537249FC2AF9A7" box="[629,912,1624,1647]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C363C73DE3DA7246FB82F9A7" bold="true" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="C3E657FBC363C73DE3DA7246FB82F9A7" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312­313, 315, 317, 319­ 320).
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE12E735DFA91F8AB" blockId="77.[264,1323,1623,1891]" box="[264,1323,1868,1891]" pageId="77" pageNumber="78">CP copulatory piece; ML medial lamellae; SLD sclerite of lateral diverticulum of internal sac.</paragraph>
<paragraph id="EA1E3692C360C73EE11E750DFE3BFE1E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<taxonomicName id="2DA14D11C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" italics="true" pageId="78" pageNumber="79">S. microphthalma</emphasis>
</taxonomicName>
in having pronotum with more asperate punctation; in males pronotum with broad medial impression, posterior angles of tergum 3 not projecting as spines; lateral diverticula of internal sac with long spiniform sclerites (
<figureCitation id="729A2A17C360C73EE1DE7585FD11FE66" box="[504,683,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 316, 318</figureCitation>
; 343); spermatheca with smaller distal portion (
<figureCitation id="729A2A17C360C73EE4D77585FE8DFE1E" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 320</figureCitation>
; 341).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E75F2FDC4FA5E" pageId="78" pageNumber="79" type="description">
<paragraph id="EA1E3692C360C73EE11E75F2FD37FDEE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E75F2FE6AFE35" bold="true" box="[312,464,483,509]" pageId="78" pageNumber="79">Description.</emphasis>
Length
<quantity id="2D599B77C360C73EE21A75F5FD73FE36" box="[572,713,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="78" pageNumber="79" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2­2.7 mm</quantity>
. Body brown or dark brown with lighter pronotum and elytra; legs brownish yellow.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7625FCB1FD0E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals 2 times their diameter. Temples 1.8­2.2 times as long as eyes. Antennal article 2 longer than 3, article 4 subquadrate, 5­10 transverse (ratio 1.5­1.8).</paragraph>
<paragraph id="EA1E3692C360C73EE11E76C5FE5FFBCE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C360C73EE40D76C5FB6CFD26" box="[1067,1238,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.699999999999999" pageId="78" pageNumber="79" unit="mm" value="0.5" valueMax="0.53" valueMin="0.47">0.47­0.53 mm</quantity>
, length
<quantity id="2D599B77C360C73EE12E76EDFE0BFCDE" box="[264,433,764,790]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.9" metricValueMin="4.3" pageId="78" pageNumber="79" unit="mm" value="0.45999999999999996" valueMax="0.49" valueMin="0.43">0.43­0.49 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.5), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7005FC18FB6E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C360C73EE2C1709DFC2FFB6E" box="[743,917,1164,1190]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Figs. 307, 309</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E70A5FD9CFAD6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
In males pronotum with wide medial impression; posterior angles of tergum 3 not projecting; tergum 7 without medial carina or tubercle; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C360C73EE1947115FDA0FAD6" box="[434,538,1284,1310]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 male tergum 8; 308 male sternum 8; 309 female tergum 8; 310 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Fig. 307</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E713DFD48FAA6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Aedeagus as in
<figureCitation id="729A2A17C360C73EE1C9713DFD2EFA8E" box="[495,660,1324,1350]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 311­319</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C360C73EE2587145FD5FFAA6" box="[638,741,1364,1390]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 313</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E716DFDC4FA5E" blockId="78.[264,1324,284,1550]" box="[312,638,1404,1430]" pageId="78" pageNumber="79">
Spermatheca as in
<figureCitation id="729A2A17C360C73EE232716DFDC0FA5E" box="[532,634,1404,1430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 median lobe, parameral view; 312 apex of median lobe, parameral view; 313 apex of left paramere; 314 median lobe, lateral view; 315 apex of median lobe, lateral view; 316 details of internal sac retracted into median lobe, lateral view; 317 copulatory piece, lateral view; 318 everted internal sac, parameral view; 319 medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 320</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E71B2FE5DF9C6" pageId="78" pageNumber="79" type="distribution">
<paragraph id="EA1E3692C360C73EE11E71B2FBFEFA76" blockId="78.[264,1324,284,1550]" box="[312,1092,1443,1470]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71B2FE62FA75" bold="true" box="[312,472,1443,1469]" pageId="78" pageNumber="79">Distribution.</emphasis>
Known from Cerro Chirripó,
<collectingCountry id="92B67602C360C73EE36671B5FC7AFA76" box="[832,960,1444,1470]" name="Costa Rica" pageId="78" pageNumber="79">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C360C73EE3F671B5FB82FA76" box="[976,1080,1444,1470]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="78" pageNumber="79">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E71DAFE5DF9C6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71DAFDBDFA2D" bold="true" box="[312,519,1483,1509]" pageId="78" pageNumber="79">Natural History.</emphasis>
<taxonomicName id="2DA14D11C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
was collected in paramo shrub litter at altitude of
<quantity id="2D599B77C360C73EE1A171E5FE5EF9C6" box="[391,484,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="78" pageNumber="79" unit="m" value="3600.0">3600 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,286 @@
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<subSubSection id="A2BB6519C366C738E12E721DFB2EF9EE" box="[264,1172,1548,1574]" pageId="72" pageNumber="73" type="key">
<paragraph id="EA1E3692C366C738E12E721DFB2EF9EE" blockId="72.[264,1172,1548,1574]" box="[264,1172,1548,1574]" pageId="72" pageNumber="73">
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21.
<taxonomicName id="2DA14D11C366C738E112721DFD6EF9EE" ID-CoL="6YBCZ" authority="Gusarov" authorityName="Gusarov" box="[308,724,1548,1574]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="micralymma" status="sp. nov.">
<emphasis id="D8D5EA80C366C738E112721DFDD9F9EE" bold="true" box="[308,611,1548,1574]" italics="true" pageId="72" pageNumber="73">Seeversiella micralymma</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC366C738E2C4721DFC9EF9EE" box="[738,804,1548,1574]" pageId="72" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C366C738E315721DFC54F9EE" box="[819,1006,1548,1574]" captionStart="FIGURES 10 ­ 16" captionStartId="6.[264,383,1647,1671]" captionTargetBox="[319,1271,331,1612]" captionTargetId="figure@6.[304,1283,320,1620]" captionTargetPageId="6" captionText="FIGURES 10 ­ 16. Details of Seeversiella globicollis (Bernhauer) (10, Anticosti, Quebec; 13, Waterton Lakes National Park, Alberta), S. scabricollis Gusarov, sp. n. (11, 15, paratype from Volcán Barva, Costa Rica) and S. micralymma Gusarov, sp. n. (12, 14, 16, paratype from Cerro Buenavista, Costa Rica). 10 ­ 12 pronotum; 13 ­ 14 mesometathorax; 15 male tergum 3 and paratergites; 16 male tergum 3. Scale bar 0.4 mm (10 ­ 11, 15), 0.3 mm (12 ­ 13, 16), 0.2 mm (14)." httpUri="https://zenodo.org/record/156423/files/figure.png" pageId="72" pageNumber="73">Figs. 12, 14, 16</figureCitation>
,
<figureCitation id="729A2A17C366C738E3DA721DFBA1F9EE" box="[1020,1051,1548,1574]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="72" pageNumber="73">18</figureCitation>
,
<figureCitation id="729A2A17C366C738E40E721DFBEFF9EE" box="[1064,1109,1548,1574]" captionStart-0="FIGURES 291 ­ 294" captionStart-1="FIGURES 295 ­ 298" captionStart-2="FIGURES 299 ­ 306" captionStartId-0="73.[264,383,1507,1531]" captionStartId-1="74.[264,382,1006,1030]" captionStartId-2="75.[264,383,1507,1531]" captionTargetBox-0="[308,1282,701,1474]" captionTargetBox-1="[306,1279,497,970]" captionTargetBox-2="[297,1292,821,1475]" captionTargetId-0="figure@73.[295,1291,689,1481]" captionTargetId-1="figure@74.[299,1287,489,979]" captionTargetId-2="figure@75.[288,1299,809,1481]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionTargetPageId-2="75" captionText-0="FIGURES 291 ­ 294. Abdominal segment 8 of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (291 ­ 292) and Cerro Buenavista (293 ­ 294), Costa Rica). 291 male tergum 8; 292 male sternum 8; 293 female tergum 8; 294 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." captionText-2="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri-0="https://zenodo.org/record/156471/files/figure.png" httpUri-1="https://zenodo.org/record/156472/files/figure.png" httpUri-2="https://zenodo.org/record/156473/files/figure.png" pageId="72" pageNumber="73">291</figureCitation>
­306)
</key>
</keyStep>
</emphasis>
</keyLead>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C366C739E12E7247FED1FE96" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="EA1E3692C366C738E12E7247FECFF909" blockId="72.[264,1324,1622,1889]" pageId="72" pageNumber="73">
<emphasis id="D8D5EA80C366C738E12E7247FE04F9B8" bold="true" box="[264,446,1622,1648]" pageId="72" pageNumber="73">
<typeStatus id="351A8830C366C738E12E7247FEFFF9B8" box="[264,325,1622,1648]" pageId="72" pageNumber="73">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C366C738E1EA7246FD86F9B9" box="[460,572,1623,1649]" pageId="72" pageNumber="73" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C366C738E24C7247FBA6F9B8" bold="true" box="[618,1052,1622,1648]" pageId="72" pageNumber="73">
<collectingCountry id="92B67602C366C738E24C7247FC99F9B8" box="[618,803,1622,1648]" name="Costa Rica" pageId="72" pageNumber="73">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89 Int. Amer. Hwy.,
<geoCoordinate id="8F955055C366C738E1D8726EFDC3F951" box="[510,633,1663,1689]" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C366C738E2A7726EFCA8F951" box="[641,786,1663,1689]" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C366C738E339726EFCC6F951" box="[799,892,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="72" pageNumber="73" unit="m" value="3200.0">3200 m</quantity>
, leaf litter (R.Anderson),
<date id="9E1F1052C366C738E48B726EFA91F951" box="[1197,1323,1663,1689]" pageId="72" pageNumber="73" value="1998-06-18">18.vi.1998</date>
(
<collectionCode id="8CB0AE57C366C738E13672B6FED2F909" box="[272,360,1703,1729]" pageId="72" pageNumber="73">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C366C739E11E72DEFED1FE96" blockId="72.[264,1324,1622,1889]" lastBlockId="73.[264,1324,284,590]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<typeStatus id="351A8830C366C738E11E72DEFE14F921" box="[312,430,1743,1769]" pageId="72" pageNumber="73" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C366C738E1E672DFFCC8F920" bold="true" box="[448,882,1742,1768]" pageId="72" pageNumber="73">
<collectingCountry id="92B67602C366C738E1E672DFFDC3F920" box="[448,633,1742,1768]" name="Costa Rica" pageId="72" pageNumber="73">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
22, 28, same data as the
<typeStatus id="351A8830C366C738E4CB72DEFE84F8D9" pageId="72" pageNumber="73" type="holotype">holotype</typeStatus>
;, 2, ditto but
<quantity id="2D599B77C366C738E21572E6FD2EF8D9" box="[563,660,1783,1809]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="72" pageNumber="73" unit="m" value="3300.0">3300 m</quantity>
,
<date id="9E1F1052C366C738E28372E6FCADF8D9" box="[677,791,1783,1809]" pageId="72" pageNumber="73" value="1997-06-08">8.vi.1997</date>
;
<emphasis id="D8D5EA80C366C738E30E72E7FC22F8D8" bold="true" box="[808,920,1782,1808]" pageId="72" pageNumber="73">Cartago:</emphasis>
6, 10, Cerro Buenavista,
<geoCoordinate id="8F955055C366C738E12E730EFE39F8F1" box="[264,387,1823,1849]" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="15" value="9.550556">9°33'02&quot;N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C366C738E2E2730EFCFBF8F1" box="[708,833,1823,1849]" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="15" value="9.717222">9°43'02&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C366C738E3CA730EFBC6F8F1" box="[1004,1148,1823,1849]" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="15" value="-83.75389">83°45'14&quot;W</geoCoordinate>
,
<quantity id="2D599B77C366C738E4AC730EFB52F8F1" box="[1162,1256,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="72" pageNumber="73" unit="m" value="3300.0">3300 m</quantity>
, subparano litter (R.Anderson),
<date id="9E1F1052C366C738E2737356FD75F8A9" box="[597,719,1863,1889]" pageId="72" pageNumber="73" value="1998-02-15">15.ii.1998</date>
; 16, 19, Cerro de la Muerte, Pan American Highway, km 89,
<quantity id="2D599B77C367C739E1CA750DFDF5FEFE" box="[492,591,284,310]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="73" pageNumber="74" unit="m" value="3300.0">3300 m</quantity>
, elfin bamboo forest litter (R.Anderson),
<date id="9E1F1052C367C739E47D750DFB68FEFE" box="[1115,1234,284,310]" pageId="73" pageNumber="74" value="1996-02-10">10.ii.1996</date>
(all ­
<collectionCode id="8CB0AE57C367C739E12E7555FEE4FE96" box="[264,350,324,350]" pageId="73" pageNumber="74">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C367C739E11E757AFC64F887" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="EA1E3692C367C739E11E757AFD79FD86" blockId="73.[264,1324,284,590]" pageId="73" pageNumber="74">
<emphasis id="D8D5EA80C367C739E11E757AFE02FE4D" bold="true" box="[312,440,363,389]" pageId="73" pageNumber="74">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C367C739E1EE757DFD4BFE4D" box="[456,753,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C367C739E1EE757DFD4BFE4D" box="[456,753,364,389]" italics="true" pageId="73" pageNumber="74">Seeversiella micralymma</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C367C739E4DE757DFED7FE65" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C367C739E4DE757DFED7FE65" italics="true" pageId="73" pageNumber="74">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body with lighter pronotum and elytra; temples 1.1­1.9 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of the male tergum 7 (
<figureCitation id="729A2A17C367C739E137761DFECDFDEE" box="[273,375,524,550]" captionStart="FIGURES 291 ­ 294" captionStartId="73.[264,383,1507,1531]" captionTargetBox="[308,1282,701,1474]" captionTargetId="figure@73.[295,1291,689,1481]" captionTargetPageId="73" captionText="FIGURES 291 ­ 294. Abdominal segment 8 of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (291 ­ 292) and Cerro Buenavista (293 ­ 294), Costa Rica). 291 male tergum 8; 292 male sternum 8; 293 female tergum 8; 294 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156471/files/figure.png" pageId="73" pageNumber="74">Fig. 291</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C367C739E4D6761DFED1FD86" captionStart-0="FIGURES 295 ­ 298" captionStart-1="FIGURES 299 ­ 306" captionStartId-0="74.[264,382,1006,1030]" captionStartId-1="75.[264,383,1507,1531]" captionTargetBox-0="[306,1279,497,970]" captionTargetBox-1="[297,1292,821,1475]" captionTargetId-0="figure@74.[299,1287,489,979]" captionTargetId-1="figure@75.[288,1299,809,1481]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionText-0="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." captionText-1="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri-0="https://zenodo.org/record/156472/files/figure.png" httpUri-1="https://zenodo.org/record/156473/files/figure.png" pageId="73" pageNumber="74">Figs. 295­305</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C367C739E2697625FD0DFD86" box="[591,695,564,590]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="73" pageNumber="74">Fig. 306</figureCitation>
).
</paragraph>
<caption id="BEDE661AC367C739E12E71F2FAA5F988" httpUri="https://zenodo.org/record/156471/files/figure.png" pageId="73" pageNumber="74" targetBox="[308,1282,701,1474]" targetPageId="73">
<paragraph id="EA1E3692C367C739E12E71F2FAA5F988" blockId="73.[264,1323,1507,1600]" pageId="73" pageNumber="74">
<emphasis id="D8D5EA80C367C739E12E71F2FE64FA33" bold="true" box="[264,478,1507,1531]" pageId="73" pageNumber="74">FIGURES 291­294</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C367C739E32671F5FBD4FA33" authority="Gusarov" authorityName="Gusarov" box="[768,1134,1508,1531]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="micralymma" status="sp. nov.">
<emphasis id="D8D5EA80C367C739E32671F5FBB7FA33" box="[768,1037,1508,1531]" italics="true" pageId="73" pageNumber="74">Seeversiella micralymma</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C367C739E45F71F2FB0FFA33" bold="true" box="[1145,1205,1507,1531]" pageId="73" pageNumber="74">
<taxonomicNameLabel id="C3E657FBC367C739E45F71F2FB0FFA33" box="[1145,1205,1507,1531]" pageId="73" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro de la Muerte (291­292) and Cerro Buenavista (293­294), Costa Rica). 291 male tergum 8; 292 male sternum 8; 293 female tergum 8; 294 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C367C739E11E7287FC64F887" blockId="73.[264,1324,1686,1872]" pageId="73" pageNumber="74">
<taxonomicName id="2DA14D11C367C739E11E7287FDE4F967" box="[312,606,1686,1711]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C367C739E11E7287FDE4F967" box="[312,606,1686,1711]" italics="true" pageId="73" pageNumber="74">Seeversiella micralymma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C367C739E2DE7287FC23F967" box="[760,921,1686,1711]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C367C739E2DE7287FC23F967" box="[760,921,1686,1711]" italics="true" pageId="73" pageNumber="74">S. curtipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C367C739E3F77287FBDFF967" box="[977,1125,1686,1711]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C367C739E3F77287FBDFF967" box="[977,1125,1686,1711]" italics="true" pageId="73" pageNumber="74">S. lativentris</emphasis>
</taxonomicName>
in having glossy pronotum with weak microsculpture; tergum 8 with four pairs of macrosetae; medial lamellae of internal sac short (
<figureCitation id="729A2A17C367C739E2AA72F7FD40F8C8" box="[652,762,1766,1792]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="73" pageNumber="74">Fig. 305</figureCitation>
), the apices of lamellae not protruding from retracted sac (
<figureCitation id="729A2A17C367C739E194731FFDA6F8E0" box="[434,540,1806,1832]" captionStart="FIGURES 295 ­ 298" captionStartId="74.[264,382,1006,1030]" captionTargetBox="[306,1279,497,970]" captionTargetId="figure@74.[299,1287,489,979]" captionTargetPageId="74" captionText="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." httpUri="https://zenodo.org/record/156472/files/figure.png" pageId="73" pageNumber="74">Fig. 295</figureCitation>
); the apex of median lobe in parameral view narrow (
<figureCitation id="729A2A17C367C739E496731FFE82F898" captionStart="FIGURES 295 ­ 298" captionStartId="74.[264,382,1006,1030]" captionTargetBox="[306,1279,497,970]" captionTargetId="figure@74.[299,1287,489,979]" captionTargetPageId="74" captionText="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." httpUri="https://zenodo.org/record/156472/files/figure.png" pageId="73" pageNumber="74">Figs. 295­ 296</figureCitation>
); and J­shaped spermatheca without umbilicus (
<figureCitation id="729A2A17C367C739E34D7327FC75F898" box="[875,975,1846,1872]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="73" pageNumber="74">Fig. 306</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C364C73BE11E750DFDC4FDEE" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="description">
<paragraph id="EA1E3692C364C73AE11E750DFDDAFE4D" blockId="74.[264,1323,284,390]" pageId="74" pageNumber="75">
<taxonomicName id="2DA14D11C364C73AE11E750DFDDDFEFD" box="[312,615,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C364C73AE11E750DFDDDFEFD" box="[312,615,284,309]" italics="true" pageId="74" pageNumber="75">Seeversiella micralymma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C364C73AE33F750DFC05FEFD" box="[793,959,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C364C73AE33F750DFC05FEFD" box="[793,959,284,309]" italics="true" pageId="74" pageNumber="75">S. luridicollis</emphasis>
</taxonomicName>
in having narrower apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C364C73AE2937555FCD8FE96" box="[693,866,324,350]" captionStart="FIGURES 295 ­ 298" captionStartId="74.[264,382,1006,1030]" captionTargetBox="[306,1279,497,970]" captionTargetId="figure@74.[299,1287,489,979]" captionTargetPageId="74" captionText="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." httpUri="https://zenodo.org/record/156472/files/figure.png" pageId="74" pageNumber="75">Figs. 295­296</figureCitation>
; 282­283) and J­shaped spermatheca without umbilicus (
<figureCitation id="729A2A17C364C73AE1CB757DFDEBFE4E" box="[493,593,364,390]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="74" pageNumber="75">Fig. 306</figureCitation>
).
</paragraph>
<caption id="BEDE661AC364C73AE12E77FFFD39FBA5" httpUri="https://zenodo.org/record/156472/files/figure.png" pageId="74" pageNumber="75" targetBox="[306,1279,497,970]" targetPageId="74">
<paragraph id="EA1E3692C364C73AE12E77FFFD39FBA5" blockId="74.[264,1323,1006,1133]" pageId="74" pageNumber="75">
<emphasis id="D8D5EA80C364C73AE12E77FFFE65FBCE" bold="true" box="[264,479,1006,1030]" pageId="74" pageNumber="75">FIGURES 295­298</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C364C73AE2A277FFFC4FFBCD" authority="Gusarov" authorityName="Gusarov" box="[644,1013,1006,1029]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="micralymma" status="sp. nov.">
<emphasis id="D8D5EA80C364C73AE2A277FFFC28FBCD" box="[644,914,1006,1029]" italics="true" pageId="74" pageNumber="75">Seeversiella micralymma</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C364C73AE42777FFFB84FBCE" bold="true" box="[1025,1086,1006,1030]" pageId="74" pageNumber="75">sp. n.</emphasis>
</taxonomicNameLabel>
(paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298).
</paragraph>
</caption>
<paragraph id="EA1E3692C364C73AE11E70D2FC68FACE" blockId="74.[264,1324,1219,1886]" pageId="74" pageNumber="75">
<emphasis id="D8D5EA80C364C73AE11E70D2FE6AFB15" bold="true" box="[312,464,1219,1245]" pageId="74" pageNumber="75">Description.</emphasis>
Length
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. Body dark brown, pronotum and elytra brown to yellowish brown, antennae, mouthparts and legs dark brown.
</paragraph>
<paragraph id="EA1E3692C364C73AE11E7105FE5CFA6E" blockId="74.[264,1324,1219,1886]" pageId="74" pageNumber="75">Head surface glossy, with fine and weak isodiametric microsculpture, with fine punctation, distance between punctures equals 2­3 times their diameter. Temples 1.1­1.9 times as long as eyes. Antennal article 2 as long as 3, article 4 elongate, 5 subquadrate, 6­10 slightly transverse.</paragraph>
<paragraph id="EA1E3692C364C73AE11E71A5FE5FF976" blockId="74.[264,1324,1219,1886]" pageId="74" pageNumber="75">
Pronotum slightly transverse, 1.1 times as wide as head, width
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, length
<quantity id="2D599B77C364C73AE12E71CDFE08FA3E" box="[264,434,1500,1526]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.0" metricValueMin="3.4" pageId="74" pageNumber="75" unit="mm" value="0.37" valueMax="0.4" valueMin="0.34">0.34­0.40 mm</quantity>
, width to length ratio 1.1, surface glossy, microsculpture and punctation as on head disc. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equals 3­4 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C364C73AE11E72DDFC18F896" blockId="74.[264,1324,1219,1886]" pageId="74" pageNumber="75">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C364C73AE2C17355FC2FF896" box="[743,917,1860,1886]" captionStart="FIGURES 291 ­ 294" captionStartId="73.[264,383,1507,1531]" captionTargetBox="[308,1282,701,1474]" captionTargetId="figure@73.[295,1291,689,1481]" captionTargetPageId="73" captionText="FIGURES 291 ­ 294. Abdominal segment 8 of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (291 ­ 292) and Cerro Buenavista (293 ­ 294), Costa Rica). 291 male tergum 8; 292 male sternum 8; 293 female tergum 8; 294 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156471/files/figure.png" pageId="74" pageNumber="75">Figs. 291, 293</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C365C73BE11E750DFB07FE66" blockId="75.[264,1324,284,670]" pageId="75" pageNumber="76">
In males posterior angles of tergum 3 not projecting as spines; lateral portions of posterior margin convex, extending posteriorly beyond the level of the medial portion of the margin (
<figureCitation id="729A2A17C365C73BE14A757DFE75FE4E" box="[364,463,364,390]" captionStart="FIGURES 10 ­ 16" captionStartId="6.[264,383,1647,1671]" captionTargetBox="[319,1271,331,1612]" captionTargetId="figure@6.[304,1283,320,1620]" captionTargetPageId="6" captionText="FIGURES 10 ­ 16. Details of Seeversiella globicollis (Bernhauer) (10, Anticosti, Quebec; 13, Waterton Lakes National Park, Alberta), S. scabricollis Gusarov, sp. n. (11, 15, paratype from Volcán Barva, Costa Rica) and S. micralymma Gusarov, sp. n. (12, 14, 16, paratype from Cerro Buenavista, Costa Rica). 10 ­ 12 pronotum; 13 ­ 14 mesometathorax; 15 male tergum 3 and paratergites; 16 male tergum 3. Scale bar 0.4 mm (10 ­ 11, 15), 0.3 mm (12 ­ 13, 16), 0.2 mm (14)." httpUri="https://zenodo.org/record/156423/files/figure.png" pageId="75" pageNumber="76">Figs. 16</figureCitation>
,
<figureCitation id="729A2A17C365C73BE1FA757DFE46FE4E" box="[476,508,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="75" pageNumber="76">18</figureCitation>
); tergum
<quantity id="2D599B77C365C73BE249757DFD27FE4E" box="[623,669,364,390]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="75" pageNumber="76" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina up to half as long as tergum; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C365C73BE46F7585FB0AFE66" box="[1097,1200,404,430]" captionStart="FIGURES 291 ­ 294" captionStartId="73.[264,383,1507,1531]" captionTargetBox="[308,1282,701,1474]" captionTargetId="figure@73.[295,1291,689,1481]" captionTargetPageId="73" captionText="FIGURES 291 ­ 294. Abdominal segment 8 of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (291 ­ 292) and Cerro Buenavista (293 ­ 294), Costa Rica). 291 male tergum 8; 292 male sternum 8; 293 female tergum 8; 294 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156471/files/figure.png" pageId="75" pageNumber="76">Fig. 291</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C365C73BE11E75ADFD48FE36" blockId="75.[264,1324,284,670]" pageId="75" pageNumber="76">
Aedeagus as in
<figureCitation id="729A2A17C365C73BE1C875ADFD2EFE1E" box="[494,660,444,470]" captionStart-0="FIGURES 295 ­ 298" captionStart-1="FIGURES 299 ­ 306" captionStartId-0="74.[264,382,1006,1030]" captionStartId-1="75.[264,383,1507,1531]" captionTargetBox-0="[306,1279,497,970]" captionTargetBox-1="[297,1292,821,1475]" captionTargetId-0="figure@74.[299,1287,489,979]" captionTargetId-1="figure@75.[288,1299,809,1481]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionText-0="FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 median lobe, parameral view; 296 apex of median lobe, parameral view; 297 median lobe, lateral view; 298 apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298)." captionText-1="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri-0="https://zenodo.org/record/156472/files/figure.png" httpUri-1="https://zenodo.org/record/156473/files/figure.png" pageId="75" pageNumber="76">Figs. 295­305</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C365C73BE25875F5FD5FFE36" box="[638,741,484,510]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="75" pageNumber="76">Fig. 302</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C365C73BE11E761DFDC4FDEE" blockId="75.[264,1324,284,670]" box="[312,638,524,550]" pageId="75" pageNumber="76">
Spermatheca as in
<figureCitation id="729A2A17C365C73BE232761DFDC0FDEE" box="[532,634,524,550]" captionStart="FIGURES 299 ­ 306" captionStartId="75.[264,383,1507,1531]" captionTargetBox="[297,1292,821,1475]" captionTargetId="figure@75.[288,1299,809,1481]" captionTargetPageId="75" captionText="FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306)." httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="75" pageNumber="76">Fig. 306</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C365C73BE11E7622FA91F927" pageId="75" pageNumber="76" type="distribution">
<paragraph id="EA1E3692C365C73BE11E7622FBCFFD86" blockId="75.[264,1324,284,670]" box="[312,1141,563,590]" pageId="75" pageNumber="76">
<emphasis id="D8D5EA80C365C73BE11E7622FE62FD85" bold="true" box="[312,472,563,589]" pageId="75" pageNumber="76">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C365C73BE3547625FC48FD86" box="[882,1010,564,590]" name="Costa Rica" pageId="75" pageNumber="76">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C365C73BE4247625FBD3FD86" box="[1026,1129,564,590]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="75" pageNumber="76">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C365C73BE11E764AFE17FD56" blockId="75.[264,1324,284,670]" pageId="75" pageNumber="76">
<emphasis id="D8D5EA80C365C73BE11E764AFDBDFDBD" bold="true" box="[312,519,603,629]" pageId="75" pageNumber="76">Natural History.</emphasis>
<taxonomicName id="2DA14D11C365C73BE23D764DFCFEFDBD" box="[539,836,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C365C73BE23D764DFCFEFDBD" box="[539,836,604,629]" italics="true" pageId="75" pageNumber="76">Seeversiella micralymma</emphasis>
</taxonomicName>
was collected in leaf litter at altitude of
<quantity id="2D599B77C365C73BE12E7695FE12FD56" box="[264,424,644,670]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="75" pageNumber="76" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">3200­3300 m</quantity>
.
</paragraph>
<caption id="BEDE661AC365C73BE12E71F2FEFAF903" httpUri="https://zenodo.org/record/156473/files/figure.png" pageId="75" pageNumber="76" targetBox="[297,1292,821,1475]" targetPageId="75">
<paragraph id="EA1E3692C365C73BE12E71F2FEFAF903" blockId="75.[264,1323,1507,1775]" pageId="75" pageNumber="76">
<emphasis id="D8D5EA80C365C73BE12E71F2FE65FA33" bold="true" box="[264,479,1507,1531]" pageId="75" pageNumber="76">FIGURES 299­306</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C365C73BE25A71F5FC56FA33" authority="Gusarov" authorityName="Gusarov" box="[636,1004,1508,1531]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="micralymma" status="sp. nov.">
<emphasis id="D8D5EA80C365C73BE25A71F5FC33FA33" box="[636,905,1508,1531]" italics="true" pageId="75" pageNumber="76">Seeversiella micralymma</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C365C73BE3D171F2FB8EFA33" bold="true" box="[1015,1076,1507,1531]" pageId="75" pageNumber="76">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Cerro de la Muerte (299, 301­305) and Cerro Buenavista (300, 306), Costa Rica). 299 details of internal sac retracted into median lobe, lateral view; 300 copulatory piece, lateral view; 301 everted internal sac, lateral view; 302 apex of left paramere; 303 details of internal sac retracted into median lobe, abparameral view; 304 everted internal sac, parameral view; 305 medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303­304), 0.1 mm (300, 302, 305­ 306).
</paragraph>
</caption>
<paragraph id="EA1E3692C365C73BE12E72C9FA91F927" blockId="75.[264,1323,1507,1775]" box="[264,1323,1752,1775]" pageId="75" pageNumber="76">CP copulatory piece; ML medial lamellae; SLD sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,260 @@
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<heading id="B15681FEC36BC735E12E7252FC77F995" bold="true" box="[264,973,1603,1630]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
<emphasis id="D8D5EA80C36BC735E12E7252FC77F995" bold="true" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<keyStep id="51552E37C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<key id="9E3061A3C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
<keyLead id="515095A7C36BC735E12E7252FC77F995" box="[264,973,1603,1630]" pageId="69" pageNumber="70">
20.
<taxonomicName id="2DA14D11C36BC735E1127255FD05F995" ID-CoL="6YBD5" authority="Gusarov" authorityName="Gusarov" box="[308,703,1603,1630]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C36BC735E1127255FDF4F996" bold="true" box="[308,590,1604,1630]" italics="true" pageId="69" pageNumber="70">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36BC735E2EA7252FCB5F995" box="[716,783,1603,1629]" pageId="69" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C36BC735E3387252FC7FF995" box="[798,965,1603,1629]" captionStart-0="FIGURES 278 ­ 281" captionStart-1="FIGURES 282 ­ 285" captionStart-2="FIGURES 286 ­ 290" captionStartId-0="70.[264,383,1351,1375]" captionStartId-1="71.[264,383,1111,1135]" captionStartId-2="72.[264,382,1260,1284]" captionTargetBox-0="[308,1276,580,1326]" captionTargetBox-1="[293,1297,523,1086]" captionTargetBox-2="[297,1290,539,1225]" captionTargetId-0="figure@70.[302,1286,569,1332]" captionTargetId-1="figure@71.[282,1306,513,1098]" captionTargetId-2="figure@72.[291,1295,529,1233]" captionTargetPageId-0="70" captionTargetPageId-1="71" captionTargetPageId-2="72" captionText-0="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-2="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156468/files/figure.png" httpUri-1="https://zenodo.org/record/156469/files/figure.png" httpUri-2="https://zenodo.org/record/156470/files/figure.png" pageId="69" pageNumber="70">Figs. 278­290</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E12E729FFBB1F8B8" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="EA1E3692C36BC735E12E729FFECFF930" blockId="69.[264,1324,1678,1904]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E12E729FFE04F960" bold="true" box="[264,446,1678,1704]" pageId="69" pageNumber="70">
<typeStatus id="351A8830C36BC735E12E729FFEFFF960" box="[264,325,1678,1704]" pageId="69" pageNumber="70">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36BC735E1EA729FFD86F960" box="[460,572,1678,1704]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36BC735E24C729FFBA6F960" bold="true" box="[618,1052,1678,1704]" pageId="69" pageNumber="70">
<collectingCountry id="92B67602C36BC735E24C729FFC99F960" box="[618,803,1678,1704]" name="Costa Rica" pageId="69" pageNumber="70">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89 Int. Amer. Hwy.,
<geoCoordinate id="8F955055C36BC735E1D872A7FDC3F918" box="[510,633,1718,1744]" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36BC735E2A772A7FCA8F918" box="[641,786,1718,1744]" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36BC735E33972A7FCC6F918" box="[799,892,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="69" pageNumber="70" unit="m" value="3200.0">3200 m</quantity>
, leaf litter (R.Anderson),
<date id="9E1F1052C36BC735E48B72A7FA91F918" box="[1197,1323,1718,1744]" pageId="69" pageNumber="70" value="1998-06-18">18.vi.1998</date>
(
<collectionCode id="8CB0AE57C36BC735E13672CFFED2F930" box="[272,360,1758,1784]" pageId="69" pageNumber="70">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7317FBB1F8B8" blockId="69.[264,1324,1678,1904]" pageId="69" pageNumber="70">
<typeStatus id="351A8830C36BC735E11E7317FE15F8E8" box="[312,431,1798,1824]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36BC735E1E27317FCC4F8E8" bold="true" box="[452,894,1798,1824]" pageId="69" pageNumber="70">
<collectingCountry id="92B67602C36BC735E1E27317FD3AF8E8" box="[452,640,1798,1824]" name="Costa Rica" pageId="69" pageNumber="70">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
7, 7, same data as the
<typeStatus id="351A8830C36BC735E4CB7317FE84F880" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
;
<emphasis id="D8D5EA80C36BC735E16C733FFE00F880" bold="true" box="[330,442,1838,1864]" pageId="69" pageNumber="70">Cartago:</emphasis>
13, 4, Cerro de la Muerte, Pan American Highway, km 89,
<quantity id="2D599B77C36BC735E4ED733FFA9DF880" box="[1227,1319,1838,1864]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="69" pageNumber="70" unit="m" value="3300.0">3300 m</quantity>
, elfin bamboo forest litter (R.Anderson),
<date id="9E1F1052C36BC735E2C57347FCE0F8B8" box="[739,858,1878,1904]" pageId="69" pageNumber="70" value="1996-02-10">10.ii.1996</date>
(all ­
<collectionCode id="8CB0AE57C36BC735E38E7347FC45F8B8" box="[936,1023,1878,1904]" pageId="69" pageNumber="70">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C368C736E11E750AFBA0F96B" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="EA1E3692C368C736E11E750AFCB4FE36" blockId="70.[264,1324,283,510]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C368C736E1E3750DFD61FEFD" box="[453,731,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E1E3750DFD61FEFD" box="[453,731,284,309]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C368C736E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C368C736E4F5750DFEFDFE95" italics="true" pageId="70" pageNumber="71">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body with contrasting lighter pronotum and elytra; temples 1.4­2.1 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of the male tergum 7 (
<figureCitation id="729A2A17C368C736E10C75ADFE2EFE1E" box="[298,404,444,470]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="70" pageNumber="71">Fig. 278</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C368C736E13775F5FE0CFE36" box="[273,438,484,510]" captionStart-0="FIGURES 282 ­ 285" captionStart-1="FIGURES 286 ­ 290" captionStartId-0="71.[264,383,1111,1135]" captionStartId-1="72.[264,382,1260,1284]" captionTargetBox-0="[293,1297,523,1086]" captionTargetBox-1="[297,1290,539,1225]" captionTargetId-0="figure@71.[282,1306,513,1098]" captionTargetId-1="figure@72.[291,1295,529,1233]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-1="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156469/files/figure.png" httpUri-1="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Figs. 282­289</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C368C736E2BC75F5FCB8FE36" box="[666,770,484,510]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
<caption id="BEDE661AC368C736E12E7156FC91FA6B" httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="70" pageNumber="71" targetBox="[308,1276,580,1326]" targetPageId="70">
<paragraph id="EA1E3692C368C736E12E7156FC91FA6B" blockId="70.[264,1323,1351,1443]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E12E7156FE65FA97" bold="true" box="[264,479,1351,1375]" pageId="70" pageNumber="71">FIGURES 278­281</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C368C736E32C7156FBD0FA96" authority="Gusarov" authorityName="Gusarov" box="[778,1130,1351,1374]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C368C736E32C7156FBBDFA96" box="[778,1031,1351,1374]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC368C736E4517156FB09FA97" box="[1143,1203,1351,1375]" pageId="70" pageNumber="71" rank="species">
<emphasis id="D8D5EA80C368C736E4517156FB09FA97" bold="true" box="[1143,1203,1351,1375]" pageId="70" pageNumber="71">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C368C736E11E71FBFBA0F96B" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<taxonomicName id="2DA14D11C368C736E11E71FBFDF5F9CB" box="[312,591,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E11E71FBFDF5F9CB" box="[312,591,1514,1539]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C368C736E2CB71FBFC35F9CB" box="[749,911,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C368C736E2CB71FBFC35F9CB" box="[749,911,1514,1539]" italics="true" pageId="70" pageNumber="71">S. curtipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C368C736E3EC71FBFBDAF9CB" box="[970,1120,1514,1539]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C368C736E3EC71FBFBDAF9CB" box="[970,1120,1514,1539]" italics="true" pageId="70" pageNumber="71">S. lativentris</emphasis>
</taxonomicName>
in having glossy pronotum with weak microsculpture; tergum 8 with four pairs of macrosetae; medial lamellae of internal sac short (
<figureCitation id="729A2A17C368C736E2AA722BFD40F99C" box="[652,762,1594,1620]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 288</figureCitation>
), the apices of lamellae not protruding from retracted sac (
<figureCitation id="729A2A17C368C736E1897273FDA2F9B4" box="[431,536,1634,1660]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Fig. 282</figureCitation>
); the apex of median lobe in parameral view broad, without basal constriction (
<figureCitation id="729A2A17C368C736E184729BFDF1F96C" box="[418,587,1674,1700]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Figs. 282­283</figureCitation>
); and S­shaped spermatheca (
<figureCitation id="729A2A17C368C736E381729BFBB1F96C" box="[935,1035,1674,1700]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C368C738E11E72A3FDC4FEFE" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71" type="description">
<paragraph id="EA1E3692C368C736E11E72A3FAA3F93B" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<taxonomicName id="2DA14D11C368C736E11E72A3FDEAF903" box="[312,592,1714,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C368C736E11E72A3FDEAF903" box="[312,592,1714,1739]" italics="true" pageId="70" pageNumber="71">Seeversiella luridicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C368C736E2D772A3FC19F903" box="[753,931,1714,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C368C736E2D772A3FC19F903" box="[753,931,1714,1739]" italics="true" pageId="70" pageNumber="71">S. micralymma</emphasis>
</taxonomicName>
in having wider apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C368C736E21E72CBFD65F93C" box="[568,735,1754,1780]" captionStart="FIGURES 282 ­ 285" captionStartId="71.[264,383,1111,1135]" captionTargetBox="[293,1297,523,1086]" captionTargetId="figure@71.[282,1306,513,1098]" captionTargetPageId="71" captionText="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="70" pageNumber="71">Figs. 282­283</figureCitation>
; 295­296) and S­shaped spermatheca (
<figureCitation id="729A2A17C368C736E48272CBFAB3F93C" box="[1188,1289,1754,1780]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="70" pageNumber="71">Fig. 290</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C368C736E11E7310FE28F8A4" blockId="70.[264,1324,1514,1900]" pageId="70" pageNumber="71">
<emphasis id="D8D5EA80C368C736E11E7310FE6AF8D3" bold="true" box="[312,464,1793,1819]" pageId="70" pageNumber="71">Description.</emphasis>
Length 2.0­
<quantity id="2D599B77C368C736E24E7313FD7DF8D4" box="[616,711,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="70" pageNumber="71" unit="mm" value="2.5">2.5 mm</quantity>
. Head and abdomen dark brown; pronotum reddish brown to brownish orange; elytra brown to brownish yellow; antennae, mouthparts and legs brown.
</paragraph>
<paragraph id="EA1E3692C369C737E11E750DFC62FE66" blockId="71.[264,1325,284,430]" pageId="71" pageNumber="72">Head surface glossy, with fine and weak isodiametric microsculpture, and fine and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Temples 1.4­2.1 times as long as eyes. Antennal article 2 as long as 3, articles 4­5 elongate, 6­7 subquadrate, 8­10 slightly transverse (ratio 1.5­2.0).</paragraph>
<caption id="BEDE661AC369C737E12E7046FEFAFB1F" httpUri="https://zenodo.org/record/156469/files/figure.png" pageId="71" pageNumber="72" targetBox="[293,1297,523,1086]" targetPageId="71">
<paragraph id="EA1E3692C369C737E12E7046FEFAFB1F" blockId="71.[264,1323,1111,1239]" pageId="71" pageNumber="72">
<emphasis id="D8D5EA80C369C737E12E7046FE59FBA7" bold="true" box="[264,483,1111,1135]" pageId="71" pageNumber="72">FIGURES 282­285</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C369C737E2BE7049FBBBFBA7" authority="Gusarov" authorityName="Gusarov" box="[664,1025,1112,1135]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C369C737E2BE7049FC23FBA7" box="[664,921,1112,1135]" italics="true" pageId="71" pageNumber="72">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC369C737E4367046FBEBFBA7" box="[1040,1105,1111,1135]" pageId="71" pageNumber="72" rank="species">
<emphasis id="D8D5EA80C369C737E4367046FBEBFBA7" bold="true" box="[1040,1105,1111,1135]" pageId="71" pageNumber="72">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285).
</paragraph>
</caption>
<paragraph id="EA1E3692C369C737E11E7133FB52F9E4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Pronotum slightly transverse, 1.1 times as wide as head, width
<quantity id="2D599B77C369C737E40D7133FB6CFAF4" box="[1067,1238,1314,1340]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="71" pageNumber="72" unit="mm" value="0.485" valueMax="0.51" valueMin="0.46">0.46­0.51 mm</quantity>
, length
<quantity id="2D599B77C369C737E12E715BFE0BFAAC" box="[264,433,1354,1380]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="71" pageNumber="72" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.40­0.47 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, and fine and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.6 times wider than long, glossy, with fine and weak isodiametric microsculpture, and fine punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C369C737E11E722BFC18F904" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­3 times their diameter on terga 3­5 and 2­4 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C369C737E2C172A3FC2FF904" box="[743,917,1714,1740]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="71" pageNumber="72">Figs. 278, 280</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C369C737E11E72CBFBD4F8D4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
In males posterior angles of tergum 3 not projecting; tergum 7 without medial carina or tubercle; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C369C737E3DC7313FBD8F8D4" box="[1018,1122,1794,1820]" captionStart="FIGURES 278 ­ 281" captionStartId="70.[264,383,1351,1375]" captionTargetBox="[308,1276,580,1326]" captionTargetId="figure@70.[302,1286,569,1332]" captionTargetPageId="70" captionText="FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 male tergum 8; 279 male sternum 8; 280 female tergum 8; 281 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156468/files/figure.png" pageId="71" pageNumber="72">Fig. 278</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C369C737E11E733BFD48F8A4" blockId="71.[264,1324,1314,1900]" pageId="71" pageNumber="72">
Aedeagus as in
<figureCitation id="729A2A17C369C737E1C8733BFD2EF88C" box="[494,660,1834,1860]" captionStart-0="FIGURES 282 ­ 285" captionStart-1="FIGURES 286 ­ 290" captionStartId-0="71.[264,383,1111,1135]" captionStartId-1="72.[264,382,1260,1284]" captionTargetBox-0="[293,1297,523,1086]" captionTargetBox-1="[297,1290,539,1225]" captionTargetId-0="figure@71.[282,1306,513,1098]" captionTargetId-1="figure@72.[291,1295,529,1233]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 median lobe, parameral view; 283 apex of median lobe, parameral view; 284 median lobe, lateral view; 285 apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285)." captionText-1="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri-0="https://zenodo.org/record/156469/files/figure.png" httpUri-1="https://zenodo.org/record/156470/files/figure.png" pageId="71" pageNumber="72">Figs. 282­289</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C369C737E2587343FD5FF8A4" box="[638,741,1874,1900]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="71" pageNumber="72">Fig. 289</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C366C738E11E750DFDC4FEFE" blockId="72.[264,1322,284,430]" box="[312,638,284,310]" pageId="72" pageNumber="73">
Spermatheca as in
<figureCitation id="729A2A17C366C738E232750DFDC0FEFE" box="[532,634,284,310]" captionStart="FIGURES 286 ­ 290" captionStartId="72.[264,382,1260,1284]" captionTargetBox="[297,1290,539,1225]" captionTargetId="figure@72.[291,1295,529,1233]" captionTargetPageId="72" captionText="FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290)." httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="72" pageNumber="73">Fig. 290</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C366C738E11E7552FB8CFA78" pageId="72" pageNumber="73" type="distribution">
<paragraph id="EA1E3692C366C738E11E7552FE17FE66" blockId="72.[264,1322,284,430]" pageId="72" pageNumber="73">
<emphasis id="D8D5EA80C366C738E11E7552FE62FE95" bold="true" box="[312,472,323,349]" pageId="72" pageNumber="73">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C366C738E3547555FC48FE96" box="[882,1010,324,350]" name="Costa Rica" pageId="72" pageNumber="73">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C366C738E4247555FBD3FE96" box="[1026,1129,324,350]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="72" pageNumber="73">Fig. 389</figureCitation>
).
<emphasis id="D8D5EA80C366C738E11E757AFDBDFE4D" bold="true" box="[312,519,363,389]" pageId="72" pageNumber="73">Natural History.</emphasis>
<taxonomicName id="2DA14D11C366C738E23C757DFC88FE4D" box="[538,818,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C366C738E23C757DFC88FE4D" box="[538,818,364,389]" italics="true" pageId="72" pageNumber="73">Seeversiella luridicollis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C366C738E12E7585FE12FE66" box="[264,424,404,430]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="72" pageNumber="73" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">3200­3300 m</quantity>
.
</paragraph>
<caption id="BEDE661AC366C738E12E70FDFEFAFA46" httpUri="https://zenodo.org/record/156470/files/figure.png" pageId="72" pageNumber="73" targetBox="[297,1290,539,1225]" targetPageId="72">
<paragraph id="EA1E3692C366C738E12E70FDFEFAFA46" blockId="72.[264,1322,1260,1456]" pageId="72" pageNumber="73">
<emphasis id="D8D5EA80C366C738E12E70FDFE67FACC" bold="true" box="[264,477,1260,1284]" pageId="72" pageNumber="73">FIGURES 286­290</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C366C738E25270FCFC6BFACB" authority="Gusarov" authorityName="Gusarov" box="[628,977,1260,1284]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luridicollis" status="sp. nov.">
<emphasis id="D8D5EA80C366C738E25270FCFCCAFACC" box="[628,880,1261,1284]" italics="true" pageId="72" pageNumber="73">Seeversiella luridicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C366C738E3FD70FDFBADFACC" bold="true" box="[987,1047,1260,1284]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="C3E657FBC366C738E3FD70FDFBADFACC" box="[987,1047,1260,1284]" pageId="72" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(286, 289, holotype; 287­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 details of internal sac retracted into median lobe, lateral view; 287 copulatory piece, lateral view; 288 everted internal sac, parameral view; 289 apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287­ 290).
</paragraph>
</caption>
<paragraph id="EA1E3692C366C738E12E7188FB8CFA78" blockId="72.[264,1322,1260,1456]" box="[264,1078,1433,1456]" pageId="72" pageNumber="73">ML medial lamellae; SLD sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,238 @@
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<heading id="B15681FEC36DC733E12E75C8FC7FFE3B" bold="true" box="[264,965,473,499]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
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19.
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<emphasis id="D8D5EA80C36DC733E11275C8FDFEFE3B" bold="true" box="[308,580,473,499]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36DC733E2E575C8FCBCFE3B" box="[707,774,473,499]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C36DC733E33375C8FC07FE3B" box="[789,957,473,499]" captionStart-0="FIGURES 265 ­ 268" captionStart-1="FIGURES 269 ­ 277" captionStartId-0="67.[264,382,1514,1538]" captionStartId-1="68.[264,383,1662,1686]" captionTargetBox-0="[285,1297,756,1480]" captionTargetBox-1="[286,1302,460,1628]" captionTargetId-0="figure@67.[276,1311,746,1488]" captionTargetId-1="figure@68.[277,1309,449,1635]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionText-0="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri-0="https://zenodo.org/record/156466/files/figure.png" httpUri-1="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Figs. 265­277</figureCitation>
)
</emphasis>
</keyLead>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36DC733E12E760FFECFFD79" pageId="67" pageNumber="68" type="materials_examined">
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<typeStatus id="351A8830C36DC733E12E760FFEF9FDF0" box="[264,323,542,568]" pageId="67" pageNumber="68">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36DC733E1F2760EFDFEFDF1" box="[468,580,543,569]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36DC733E250760FFC78FDF0" bold="true" box="[630,962,542,568]" pageId="67" pageNumber="68">
<collectingCountry id="92B67602C36DC733E250760FFC8BFDF0" box="[630,817,542,568]" name="Costa Rica" pageId="67" pageNumber="68">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
<quantity id="2D599B77C36DC733E3F0760EFBA9FDF1" box="[982,1043,543,569]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="km" value="2.0">2 km</quantity>
S Villa Mills,
<quantity id="2D599B77C36DC733E4EE760EFA9CFDF1" box="[1224,1318,543,569]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="67" pageNumber="68" unit="m" value="3000.0">3000 m</quantity>
, ridgetop oak forest litter (R.Anderson),
<date id="9E1F1052C36DC733E2FC7656FCEBFDA9" box="[730,849,583,609]" pageId="67" pageNumber="68" value="1996-02-10">10.ii.1996</date>
(
<collectionCode id="8CB0AE57C36DC733E3477656FC03FDA9" box="[865,953,583,609]" pageId="67" pageNumber="68">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36DC733E11E767EFECFFD79" blockId="67.[264,1323,542,689]" pageId="67" pageNumber="68">
<typeStatus id="351A8830C36DC733E11E767EFE15FD41" box="[312,431,623,649]" pageId="67" pageNumber="68" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36DC733E1EF767FFCA4FD40" bold="true" box="[457,798,622,648]" pageId="67" pageNumber="68">
<collectingCountry id="92B67602C36DC733E1EF767FFD3DFD40" box="[457,647,622,648]" name="Costa Rica" pageId="67" pageNumber="68">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
<specimenCount id="FCA7FD1BC36DC733E310767EFC66FD41" box="[822,988,623,649]" pageId="67" pageNumber="68" type="generic">78 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C36DC733E4E2767EFA91FD41" box="[1220,1323,623,649]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C36DC733E1367686FED2FD79" box="[272,360,663,689]" pageId="67" pageNumber="68">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC36DC733E12E71FBFD63F98F" httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="67" pageNumber="68" targetBox="[285,1297,756,1480]" targetPageId="67">
<paragraph id="EA1E3692C36DC733E12E71FBFD63F98F" blockId="67.[264,1323,1514,1607]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E12E71FBFE5AF9CA" bold="true" box="[264,480,1514,1538]" pageId="67" pageNumber="68">FIGURES 265­268</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C36DC733E32971FAFBD2F9C9" authority="Gusarov" authorityName="Gusarov" box="[783,1128,1514,1538]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="D8D5EA80C36DC733E32971FAFBBEF9CA" box="[783,1028,1515,1538]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C36DC733E45371FBFB09F9CA" bold="true" box="[1141,1203,1514,1538]" pageId="67" pageNumber="68">
<taxonomicNameLabel id="C3E657FBC36DC733E45371FBFB09F9CA" box="[1141,1203,1514,1538]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C36DC735E11E729CFC43FE1D" lastPageId="69" lastPageNumber="70" pageId="67" pageNumber="68" type="diagnosis">
<paragraph id="EA1E3692C36DC733E11E729CFDA5F8A7" blockId="67.[264,1323,1677,1903]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E11E729CFE02F96F" bold="true" box="[312,440,1677,1703]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C36DC733E1E1729CFD69F96E" box="[455,723,1677,1702]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36DC733E1E1729CFD69F96E" box="[455,723,1677,1702]" italics="true" pageId="67" pageNumber="68">Seeversiella lativentris</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C36DC733E4F5729CFEFDF906" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36DC733E4F5729CFEFDF906" italics="true" pageId="67" pageNumber="68">Seeversiella</emphasis>
</taxonomicName>
by having brown body with lighter pronotum and elytra; temples 1.5­1.8 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at posterior margin of male tergum 7 (
<figureCitation id="729A2A17C36DC733E47B7314FB79F8D7" box="[1117,1219,1797,1823]" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="67" pageNumber="68">Fig. 265</figureCitation>
); tergum 8 with three pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C36DC733E463733CFB54F88F" box="[1093,1262,1837,1863]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Figs. 269­276</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C36DC733E18D7344FDA8F8A7" box="[427,530,1877,1903]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="67" pageNumber="68">Fig. 277</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36AC734E11E750DFBDCFE96" blockId="68.[264,1323,284,350]" pageId="68" pageNumber="69">
<taxonomicName id="2DA14D11C36AC734E11E750DFDF9FEFD" box="[312,579,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36AC734E11E750DFDF9FEFD" box="[312,579,284,309]" italics="true" pageId="68" pageNumber="69">Seeversiella lativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36AC734E2FA750DFCC7FEFD" box="[732,893,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36AC734E2FA750DFCC7FEFD" box="[732,893,284,309]" italics="true" pageId="68" pageNumber="69">S. curtipennis</emphasis>
</taxonomicName>
in having stronger basal constriction of the apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C36AC734E3667555FC5DFE96" box="[832,999,324,350]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="68" pageNumber="69">Figs. 269­270</figureCitation>
; 251­252).
</paragraph>
<caption id="BEDE661AC36AC734E12E726FFCA5F88A" httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="68" pageNumber="69" targetBox="[286,1302,460,1628]" targetPageId="68">
<paragraph id="EA1E3692C36AC734E12E726FFCA5F88A" blockId="68.[264,1323,1662,1895]" pageId="68" pageNumber="69">
<emphasis id="D8D5EA80C36AC734E12E726FFE58F95E" bold="true" box="[264,482,1662,1686]" pageId="68" pageNumber="69">FIGURES 269­277</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C36AC734E2AC726FFC52F95D" authority="Gusarov" authorityName="Gusarov" box="[650,1000,1662,1685]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="D8D5EA80C36AC734E2AC726FFC3BF95D" box="[650,897,1662,1685]" italics="true" pageId="68" pageNumber="69">Seeversiella lativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36AC734E3D3726FFB8FF95E" box="[1013,1077,1662,1686]" pageId="68" pageNumber="69" rank="species">
<emphasis id="D8D5EA80C36AC734E3D3726FFB8FF95E" bold="true" box="[1013,1077,1662,1686]" pageId="68" pageNumber="69">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274­276), 0.1 mm (270, 272­273, 277).
</paragraph>
</caption>
<paragraph id="EA1E3692C36AC734E12E7341FE46F8AF" blockId="68.[264,1323,1662,1895]" box="[264,508,1872,1895]" pageId="68" pageNumber="69">ML medial lamellae.</paragraph>
<paragraph id="EA1E3692C36BC735E11E750DFC43FE1D" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<taxonomicName id="2DA14D11C36BC735E11E750DFDFEFEFD" box="[312,580,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36BC735E11E750DFDFEFEFD" box="[312,580,284,309]" italics="true" pageId="69" pageNumber="70">Seeversiella lativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36BC735E2C7750DFC3AFEFD" box="[737,896,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="luridicollis">
<emphasis id="D8D5EA80C36BC735E2C7750DFC3AFEFD" box="[737,896,284,309]" italics="true" pageId="69" pageNumber="70">S. luridicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C36BC735E39C750DFBD6FEFD" box="[954,1132,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="micralymma">
<emphasis id="D8D5EA80C36BC735E39C750DFBD6FEFD" box="[954,1132,284,309]" italics="true" pageId="69" pageNumber="70">S. micralymma</emphasis>
</taxonomicName>
in having matte pronotum with strong isodiametric microsculpture; tergum 8 with three pairs of macrosetae; medial lamellae of internal sac long (
<figureCitation id="729A2A17C36BC735E2CB757DFCEEFE4E" box="[749,852,364,390]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 274</figureCitation>
), the apices of lamellae protruding from retracted sac (
<figureCitation id="729A2A17C36BC735E18B7585FDAEFE66" box="[429,532,404,430]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 269</figureCitation>
); different shape of the median lobe (
<figureCitation id="729A2A17C36BC735E3E97585FBCDFE66" box="[975,1143,404,430]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Figs. 269­270</figureCitation>
; 282­283; 295­ 296) and L­shaped spermatheca with large umbilicus (
<figureCitation id="729A2A17C36BC735E3A375ADFC53FE1E" box="[901,1001,444,470]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 277</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E11E75F2FDC4FAA6" pageId="69" pageNumber="70" type="description">
<paragraph id="EA1E3692C36BC735E11E75F2FAB0FDEE" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E75F2FE6AFE35" bold="true" box="[312,464,483,509]" pageId="69" pageNumber="70">Description.</emphasis>
Length 2.0­
<quantity id="2D599B77C36BC735E24A75F5FD71FE36" box="[620,715,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="69" pageNumber="70" unit="mm" value="2.6">2.6 mm</quantity>
. Head and abdomen brown to dark brown; pronotum reddish brown; elytra reddish brown or yellowish brown; antennae and legs brown.
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7625FB0DFD56" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">Head surface matte, with fine and dense isodiametric microsculpture, and fine and poorly visible punctation. Temples 1.5­1.8 times as long as eyes. Antennal article 2 as long as 3, article 4 transverse (ratio 1.2), 5­10 strongly transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C36BC735E11E76BDFD2EFC7E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C36BC735E40D76BDFB6CFD0E" box="[1067,1238,684,710]" metricMagnitude="-4" metricUnit="m" metricValue="4.3999999999999995" metricValueMax="4.699999999999999" metricValueMin="4.1" pageId="69" pageNumber="70" unit="mm" value="0.43999999999999995" valueMax="0.47" valueMin="0.41">0.41­0.47 mm</quantity>
, length
<quantity id="2D599B77C36BC735E12E76C5FE09FD26" box="[264,435,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.3" metricValueMin="3.4" pageId="69" pageNumber="70" unit="mm" value="0.385" valueMax="0.43" valueMin="0.34">0.34­0.43 mm</quantity>
, width to length ratio 1.1, surface matte, with fine and dense isodiametric microsculpture, and fine and poorly visible punctation. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.3), 1.7 times wider than long, matte, with fine and weak isodiametric microsculpture, and fine punctation, distance between punctures equals ½­1 time their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C36BC735E11E77D5FC16FB9E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C36BC735E2D4702DFC1AFB9E" box="[754,928,1084,1110]" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="69" pageNumber="70">Figs. 265, 267</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7075FEFFFB3E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
In large males posterior angles of tergum 3 projecting as spines up to half as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to ½ as long as tergum length; posterior margin of tergum 8 without clear emargination (
<figureCitation id="729A2A17C36BC735E4DD70A5FE82FB3E" captionStart="FIGURES 265 ­ 268" captionStartId="67.[264,382,1514,1538]" captionTargetBox="[285,1297,756,1480]" captionTargetId="figure@67.[276,1311,746,1488]" captionTargetPageId="67" captionText="FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 male tergum 8; 266 male sternum 8; 267 female tergum 8; 268 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156466/files/figure.png" pageId="69" pageNumber="70">Fig. 265</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7115FD48FA8E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
Aedeagus as in
<figureCitation id="729A2A17C36BC735E1C87115FD2FFAD6" box="[494,661,1284,1310]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Figs. 269­276</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C36BC735E258713DFD5FFA8E" box="[638,741,1324,1350]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 273</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E7145FDC4FAA6" blockId="69.[264,1324,284,1510]" box="[312,638,1364,1390]" pageId="69" pageNumber="70">
Spermatheca as in
<figureCitation id="729A2A17C36BC735E2327145FDC0FAA6" box="[532,634,1364,1390]" captionStart="FIGURES 269 ­ 277" captionStartId="68.[264,383,1662,1686]" captionTargetBox="[286,1302,460,1628]" captionTargetId="figure@68.[277,1309,449,1635]" captionTargetPageId="68" captionText="FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 median lobe, parameral view; 270 apex of median lobe, parameral view; 271 median lobe, lateral view; 272 apex of median lobe, lateral view; 273 apex of left paramere; 274 everted internal sac, parameral view; 275 everted internal sac, lateral view; 276 details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277)." httpUri="https://zenodo.org/record/156467/files/figure.png" pageId="69" pageNumber="70">Fig. 277</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36BC735E11E716AFED2FA2E" pageId="69" pageNumber="70" type="distribution">
<paragraph id="EA1E3692C36BC735E11E716AFBCFFA5E" blockId="69.[264,1324,284,1510]" box="[312,1141,1403,1430]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E716AFE62FA5D" bold="true" box="[312,472,1403,1429]" pageId="69" pageNumber="70">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C36BC735E354716DFC48FA5E" box="[882,1010,1404,1430]" name="Costa Rica" pageId="69" pageNumber="70">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C36BC735E424716DFBD3FA5E" box="[1026,1129,1404,1430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="69" pageNumber="70">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36BC735E11E71B2FED2FA2E" blockId="69.[264,1324,284,1510]" pageId="69" pageNumber="70">
<emphasis id="D8D5EA80C36BC735E11E71B2FDBDFA75" bold="true" box="[312,519,1443,1469]" pageId="69" pageNumber="70">Natural History.</emphasis>
<taxonomicName id="2DA14D11C36BC735E23A71B5FC91FA75" box="[540,811,1444,1469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36BC735E23A71B5FC91FA75" box="[540,811,1444,1469]" italics="true" pageId="69" pageNumber="70">Seeversiella lativentris</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C36BC735E12E71DDFEDEFA2E" box="[264,356,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="69" pageNumber="70" unit="m" value="3000.0">3000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,204 @@
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<mods:title id="B5E91E4E63344056840466F5F6EC0DEA">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="EA1E3692C373C72DE12E750AFC6DFEFD" blockId="93.[264,983,283,310]" box="[264,983,283,310]" pageId="93" pageNumber="94">
<heading id="B15681FEC373C72DE12E750AFC6DFEFD" bold="true" box="[264,983,283,310]" fontSize="11" level="1" pageId="93" pageNumber="94" reason="1">
<emphasis id="D8D5EA80C373C72DE12E750AFC6DFEFD" bold="true" box="[264,983,283,310]" pageId="93" pageNumber="94">
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<key id="9E3061A3C373C72DE12E750AFC6DFEFD" box="[264,983,283,310]" pageId="93" pageNumber="94">
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28.
<taxonomicName id="2DA14D11C373C72DE112750DFD73FEFD" ID-CoL="6YBDC" authority="Gusarov" authorityName="Gusarov" box="[308,713,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="paramoana" status="sp. nov.">
<emphasis id="D8D5EA80C373C72DE112750DFDECFEFE" bold="true" box="[308,598,284,310]" italics="true" pageId="93" pageNumber="94">Seeversiella paramoana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC373C72DE2F3750AFCADFEFD" box="[725,791,283,309]" pageId="93" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C373C72DE301750AFC75FEFD" box="[807,975,283,309]" captionStart-0="FIGURES 372 ­ 375" captionStart-1="FIGURES 376 ­ 384" captionStartId-0="92.[264,382,1784,1808]" captionStartId-1="93.[264,383,1671,1695]" captionTargetBox-0="[296,1289,901,1750]" captionTargetBox-1="[282,1309,574,1637]" captionTargetId-0="figure@92.[287,1301,889,1757]" captionTargetId-1="figure@93.[268,1319,558,1647]" captionTargetPageId-0="92" captionTargetPageId-1="93" captionText-0="FIGURES 372 ­ 375. Abdominal segment 8 of Seeversiella paramoana Gusarov, sp. n. (372 ­ 373, holotype; 374 ­ 375, paratype from Cerro Chirripó, Costa Rica). 372 male tergum 8; 373 male sternum 8; 374 female tergum 8; 375 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri-0="https://zenodo.org/record/156485/files/figure.png" httpUri-1="https://zenodo.org/record/156486/files/figure.png" pageId="93" pageNumber="94">Figs. 372­384</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C373C72DE12E7572FAAFFE3E" pageId="93" pageNumber="94" type="materials_examined">
<paragraph id="EA1E3692C373C72DE12E7572FECFFE06" blockId="93.[264,1323,355,502]" pageId="93" pageNumber="94">
<emphasis id="D8D5EA80C373C72DE12E7572FE05FEB5" bold="true" box="[264,447,355,381]" pageId="93" pageNumber="94">
<typeStatus id="351A8830C373C72DE12E7572FEF9FEB5" box="[264,323,355,381]" pageId="93" pageNumber="94">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C373C72DE1F17575FDFDFEB6" box="[471,583,356,382]" pageId="93" pageNumber="94" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C373C72DE25D7572FC79FEB5" bold="true" box="[635,963,355,381]" pageId="93" pageNumber="94">
<collectingCountry id="92B67602C373C72DE25D7572FC82FEB5" box="[635,824,355,381]" name="Costa Rica" pageId="93" pageNumber="94">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Cerro Chirripó, Valle de los Conelos,
<geoCoordinate id="8F955055C373C72DE15C759DFE4FFE6E" box="[378,501,396,422]" direction="north" orientation="latitude" pageId="93" pageNumber="94" precision="15" value="9.476944">9°28'37&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C373C72DE1D9759DFD35FE6E" box="[511,655,396,422]" direction="west" orientation="longitude" pageId="93" pageNumber="94" precision="15" value="-83.489716">83°29'23&quot;W</geoCoordinate>
,
<quantity id="2D599B77C373C72DE2B9759DFD44FE6E" box="[671,766,396,422]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="93" pageNumber="94" unit="m" value="3600.0">3600 m</quantity>
, paramo shrub litter (R.Anderson),
<date id="9E1F1052C373C72DE48B759DFA90FE6E" box="[1197,1322,396,422]" pageId="93" pageNumber="94" value="1999-06-26">26.vi.1999</date>
(
<collectionCode id="8CB0AE57C373C72DE13675A5FED2FE06" box="[272,360,436,462]" pageId="93" pageNumber="94">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C373C72DE11E75CDFAAFFE3E" blockId="93.[264,1323,355,502]" box="[312,1301,475,502]" pageId="93" pageNumber="94">
<typeStatus id="351A8830C373C72DE11E75CDFE14FE3E" box="[312,430,476,502]" pageId="93" pageNumber="94" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C373C72DE1E675CAFD44FE3D" bold="true" box="[448,766,475,501]" pageId="93" pageNumber="94">
<collectingCountry id="92B67602C373C72DE1E675CAFDC0FE3D" box="[448,634,475,501]" name="Costa Rica" pageId="93" pageNumber="94">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, 6, same data as the
<typeStatus id="351A8830C373C72DE41C75CDFB1BFE3E" box="[1082,1185,476,502]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C373C72DE49675CDFAB2FE3E" box="[1200,1288,476,502]" pageId="93" pageNumber="94">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C373C72DE12E7296FD0AF8A7" blockId="93.[264,1323,1671,1903]" pageId="93" pageNumber="94">
<emphasis id="D8D5EA80C373C72DE12E7296FE59F957" bold="true" box="[264,483,1671,1695]" pageId="93" pageNumber="94">FIGURES 376­384</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C373C72DE2A97299FC46F957" authority="Gusarov" authorityName="Gusarov" box="[655,1020,1672,1695]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="paramoana" status="sp. nov.">
<emphasis id="D8D5EA80C373C72DE2A97299FC2CF957" box="[655,918,1672,1695]" italics="true" pageId="93" pageNumber="94">Seeversiella paramoana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C373C72DE42D7296FBF1F957" bold="true" box="[1035,1099,1671,1695]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="C3E657FBC373C72DE42D7296FBF1F957" box="[1035,1099,1671,1695]" pageId="93" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(376­380, 382­383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380­ 381, 384), 0.2 mm (377, 379, 382­383).
</paragraph>
</caption>
<subSubSection id="A2BB6519C370C72EE11E750AFB75FDEE" pageId="94" pageNumber="95" type="diagnosis">
<paragraph id="EA1E3692C370C72EE11E750AFC8EFE1E" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
<emphasis id="D8D5EA80C370C72EE11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="94" pageNumber="95">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C370C72EE1ED750DFD50FEFD" box="[459,746,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C370C72EE1ED750DFD50FEFD" box="[459,746,284,309]" italics="true" pageId="94" pageNumber="95">Seeversiella paramoana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C370C72EE4DE750DFED7FE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C370C72EE4DE750DFED7FE95" italics="true" pageId="94" pageNumber="95">Seeversiella</emphasis>
</taxonomicName>
by having brownish yellow body; small eyes (temple length to eye length ratio 4.0­5.0); glossy pronotum with weak isodiametric microsculpture; elytra much shorter than pronotum; wings reduced, shorter than elytra; the distinct shape of aedeagus (
<figureCitation id="729A2A17C370C72EE4D67585FE66FE1E" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Figs. 376­380, 382­384</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C370C72EE2E675ADFC9CFE1E" box="[704,806,444,470]" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Fig. 381</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C370C72EE11E75F5FB75FDEE" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
<taxonomicName id="2DA14D11C370C72EE11E75F5FDEFFE35" box="[312,597,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C370C72EE11E75F5FDEFFE35" box="[312,597,484,509]" italics="true" pageId="94" pageNumber="95">Seeversiella paramoana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C370C72EE2D375F5FCDFFE35" box="[757,869,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C370C72EE2D375F5FCDFFE35" box="[757,869,484,509]" italics="true" pageId="94" pageNumber="95">S. flavida</emphasis>
</taxonomicName>
in having tergum 8 with four pairs of macrosetae; and pointed apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C370C72EE43F761DFB78FDEE" box="[1049,1218,524,550]" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Figs. 377­378</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C370C72EE11E7625FDC7FCAE" pageId="94" pageNumber="95" type="description">
<paragraph id="EA1E3692C370C72EE11E7625FBD0FDBE" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
<taxonomicName id="2DA14D11C370C72EE11E7625FDEFFD85" box="[312,597,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C370C72EE11E7625FDEFFD85" box="[312,597,564,589]" italics="true" pageId="94" pageNumber="95">Seeversiella paramoana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C370C72EE2D07625FCD9FD85" box="[758,867,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C370C72EE2D07625FCD9FD85" box="[758,867,564,589]" italics="true" pageId="94" pageNumber="95">S. adusta</emphasis>
</taxonomicName>
in having tergum 8 with four pairs of macrosetae; and longer proximal portion of spermatheca (
<figureCitation id="729A2A17C370C72EE389764DFB9AFDBE" box="[943,1056,604,630]" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Figs. 381</figureCitation>
; 358).
</paragraph>
<paragraph id="EA1E3692C370C72EE11E7692FE53FD0E" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
<emphasis id="D8D5EA80C370C72EE11E7692FE6AFD55" bold="true" box="[312,464,643,669]" pageId="94" pageNumber="95">Description.</emphasis>
Length
<quantity id="2D599B77C370C72EE21C7695FD7CFD56" box="[570,710,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="94" pageNumber="95" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2­2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C370C72EE3A37695FB95FD56" box="[901,1071,644,670]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="94" pageNumber="95" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.40­0.47 mm</quantity>
. Body brownish yellow to light brown.
</paragraph>
<paragraph id="EA1E3692C370C72EE11E76C5FD02FCDE" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
In all external character states
<taxonomicName id="2DA14D11C370C72EE2BB76C5FCFBFD25" box="[669,833,724,749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C370C72EE2BB76C5FCFBFD25" box="[669,833,724,749]" italics="true" pageId="94" pageNumber="95">S. paramoana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C370C72EE43076C5FB3FFD25" box="[1046,1157,724,749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C370C72EE43076C5FB3FFD25" box="[1046,1157,724,749]" italics="true" pageId="94" pageNumber="95">S. flavida</emphasis>
</taxonomicName>
, but differs in the characters listed in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C370C72EE11E7735FCB1FCF6" blockId="94.[264,1324,283,990]" box="[312,779,804,830]" pageId="94" pageNumber="95">
Aedeagus as in
<figureCitation id="729A2A17C370C72EE1D67735FCB2FCF6" box="[496,776,804,830]" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Figs. 376­380, 382­384</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C370C72EE11E775DFDC7FCAE" blockId="94.[264,1324,283,990]" box="[312,637,844,870]" pageId="94" pageNumber="95">
Spermatheca as in
<figureCitation id="729A2A17C370C72EE232775DFDC3FCAE" box="[532,633,844,870]" captionStart="FIGURES 376 ­ 384" captionStartId="93.[264,383,1671,1695]" captionTargetBox="[282,1309,574,1637]" captionTargetId="figure@93.[268,1319,558,1647]" captionTargetPageId="93" captionText="FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 apex of left paramere; 377 median lobe, parameral view; 378 apex of median lobe, parameral view; 379 median lobe, lateral view; 380 apex of median lobe, lateral view; 381 ­ spermatheca; 382 details of internal sac retracted into median lobe, abparameral view; 383 details of internal sac retracted into median lobe, lateral view; 384 everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383)." httpUri="https://zenodo.org/record/156486/files/figure.png" pageId="94" pageNumber="95">Fig. 381</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C370C72EE11E7762FE78FC16" pageId="94" pageNumber="95" type="distribution">
<paragraph id="EA1E3692C370C72EE11E7762FBFEFC46" blockId="94.[264,1324,283,990]" box="[312,1092,883,910]" pageId="94" pageNumber="95">
<emphasis id="D8D5EA80C370C72EE11E7762FE62FC45" bold="true" box="[312,472,883,909]" pageId="94" pageNumber="95">Distribution.</emphasis>
Known from Cerro Chirripó,
<collectingCountry id="92B67602C370C72EE3667765FC7AFC46" box="[832,960,884,910]" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C370C72EE3F67765FB82FC46" box="[976,1080,884,910]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="94" pageNumber="95">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C370C72EE11E778AFE78FC16" blockId="94.[264,1324,283,990]" pageId="94" pageNumber="95">
<emphasis id="D8D5EA80C370C72EE11E778AFDBEFC7D" bold="true" box="[312,516,923,949]" pageId="94" pageNumber="95">Natural History.</emphasis>
<taxonomicName id="2DA14D11C370C72EE235778DFC8AFC7D" box="[531,816,924,949]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C370C72EE235778DFC8AFC7D" box="[531,816,924,949]" italics="true" pageId="94" pageNumber="95">Seeversiella paramoana</emphasis>
</taxonomicName>
was collected in paramo shrub litter at altitude of
<quantity id="2D599B77C370C72EE14777D5FE07FC16" box="[353,445,964,990]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="94" pageNumber="95" unit="m" value="3600.0">3600 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,168 @@
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26.
<taxonomicName id="2DA14D11C376C728E112764DFD37FDBD" ID-CoL="6YBCH" authority="Gusarov" authorityName="Gusarov" box="[308,653,603,630]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="adusta" status="sp. nov.">
<emphasis id="D8D5EA80C376C728E112764DFDA1FDBE" bold="true" box="[308,539,604,630]" italics="true" pageId="88" pageNumber="89">Seeversiella adusta</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC376C728E2BC764AFD66FDBD" box="[666,732,603,629]" pageId="88" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C376C728E2CA764AFC2EFDBD" box="[748,916,603,629]" captionStart="FIGURES 356 ­ 358" captionStartId="88.[264,382,1791,1815]" captionTargetBox="[333,1254,860,1773]" captionTargetId="figure@88.[322,1264,844,1781]" captionTargetPageId="88" captionText="FIGURES 356 ­ 358. Abdominal segment 8 and spermatheca of Seeversiella adusta Gusarov, sp. n. (holotype). 356 female tergum 8; 357 female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356 ­ 357), 0.1 mm (358)." httpUri="https://zenodo.org/record/156482/files/figure.png" pageId="88" pageNumber="89">Figs. 356­358</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C376C728E12E76B7FECFFCD9" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="EA1E3692C376C728E12E76B7FECFFCD9" blockId="88.[264,1323,678,785]" pageId="88" pageNumber="89">
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<typeStatus id="351A8830C376C728E12E76B7FEFFFD08" box="[264,325,678,704]" pageId="88" pageNumber="89">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C376C728E1EA76B6FD86FD09" box="[460,572,679,705]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C376C728E24C76B7FBA6FD08" bold="true" box="[618,1052,678,704]" pageId="88" pageNumber="89">
<collectingCountry id="92B67602C376C728E24C76B7FC99FD08" box="[618,803,678,704]" name="Costa Rica" pageId="88" pageNumber="89">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C376C728E1D976DEFDC1FD21" box="[511,635,719,745]" direction="north" orientation="latitude" pageId="88" pageNumber="89" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C376C728E2A476DEFCA8FD21" box="[642,786,719,745]" direction="west" orientation="longitude" pageId="88" pageNumber="89" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
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, forest litter (R.Anderson),
<date id="9E1F1052C376C728E49A76DEFA90FD21" box="[1212,1322,719,745]" pageId="88" pageNumber="89" value="1997-06-08">8.vi.1997</date>
(
<collectionCode id="8CB0AE57C376C728E13676E6FED2FCD9" box="[272,360,759,785]" pageId="88" pageNumber="89">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
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<emphasis id="D8D5EA80C376C728E12E72EEFE64F8DF" bold="true" box="[264,478,1791,1815]" pageId="88" pageNumber="89">FIGURES 356­358</emphasis>
. Abdominal segment 8 and spermatheca of
<taxonomicName id="2DA14D11C376C728E3907311FB5CF8DF" authority="Gusarov" authorityName="Gusarov" box="[950,1254,1792,1815]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="adusta" status="sp. nov.">
<emphasis id="D8D5EA80C376C728E3907311FB3EF8DF" box="[950,1156,1792,1815]" italics="true" pageId="88" pageNumber="89">Seeversiella adusta</emphasis>
Gusarov
</taxonomicName>
,
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<taxonomicNameLabel id="C3E657FBC376C728E4C972EEFA90F8DF" box="[1263,1322,1791,1815]" pageId="88" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 356 female tergum 8; 357 female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356­357), 0.1 mm (358).
</paragraph>
</caption>
<subSubSection id="A2BB6519C377C729E11E750AFD8EFE36" pageId="89" pageNumber="90" type="diagnosis">
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<emphasis id="D8D5EA80C377C729E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="89" pageNumber="90">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C377C729E1E8750DFD0FFEFD" box="[462,693,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C377C729E1E8750DFD0FFEFD" box="[462,693,284,309]" italics="true" pageId="89" pageNumber="90">Seeversiella adusta</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C377C729E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C377C729E4F5750DFEFDFE95" italics="true" pageId="89" pageNumber="90">Seeversiella</emphasis>
</taxonomicName>
by having uniformly brown body; small eyes (temples 3.3 times as long as eyes); elytra much shorter than pronotum; reduced wings (shorter than elytra); tergum 8 with three pairs of macrosetae; and by the distinct shape of spermatheca (
<figureCitation id="729A2A17C377C729E40E7585FB2AFE66" box="[1064,1168,404,430]" captionStart="FIGURES 356 ­ 358" captionStartId="88.[264,382,1791,1815]" captionTargetBox="[333,1254,860,1773]" captionTargetId="figure@88.[322,1264,844,1781]" captionTargetPageId="88" captionText="FIGURES 356 ­ 358. Abdominal segment 8 and spermatheca of Seeversiella adusta Gusarov, sp. n. (holotype). 356 female tergum 8; 357 female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356 ­ 357), 0.1 mm (358)." httpUri="https://zenodo.org/record/156482/files/figure.png" pageId="89" pageNumber="90">Fig. 358</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C377C729E11E75ADFD8EFE36" blockId="89.[264,1324,283,830]" pageId="89" pageNumber="90">
<taxonomicName id="2DA14D11C377C729E11E75ADFDA6FE1D" box="[312,540,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C377C729E11E75ADFDA6FE1D" box="[312,540,444,469]" italics="true" pageId="89" pageNumber="90">Seeversiella adusta</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C377C729E29F75ADFC93FE1D" box="[697,809,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C377C729E29F75ADFC93FE1D" box="[697,809,444,469]" italics="true" pageId="89" pageNumber="90">S. flavida</emphasis>
</taxonomicName>
in having shorter proximal portion of spermatheca (
<figureCitation id="729A2A17C377C729E15C75F5FE51FE36" box="[378,491,484,510]" captionStart="FIGURES 356 ­ 358" captionStartId="88.[264,382,1791,1815]" captionTargetBox="[333,1254,860,1773]" captionTargetId="figure@88.[322,1264,844,1781]" captionTargetPageId="88" captionText="FIGURES 356 ­ 358. Abdominal segment 8 and spermatheca of Seeversiella adusta Gusarov, sp. n. (holotype). 356 female tergum 8; 357 female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356 ­ 357), 0.1 mm (358)." httpUri="https://zenodo.org/record/156482/files/figure.png" pageId="89" pageNumber="90">Figs. 358</figureCitation>
; 371).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C377C729E11E761AFE4BFD0E" pageId="89" pageNumber="90" type="description">
<paragraph id="EA1E3692C377C729E11E761AFE68FD86" blockId="89.[264,1324,283,830]" pageId="89" pageNumber="90">
<emphasis id="D8D5EA80C377C729E11E761AFE6AFDED" bold="true" box="[312,464,523,549]" pageId="89" pageNumber="90">Description.</emphasis>
Length 2.0 mm, pronotal width
<quantity id="2D599B77C377C729E37D761DFC73FDEE" box="[859,969,524,550]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="89" pageNumber="90" unit="mm" value="0.43">0.43 mm</quantity>
. Body uniformly brown, legs brownish yellow.
</paragraph>
<paragraph id="EA1E3692C377C729E11E764DFC2BFD56" blockId="89.[264,1324,283,830]" pageId="89" pageNumber="90">
In all external character states
<taxonomicName id="2DA14D11C377C729E2BA764DFCB2FDBD" box="[668,776,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C377C729E2BA764DFCB2FDBD" box="[668,776,604,629]" italics="true" pageId="89" pageNumber="90">S. adusta</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C377C729E3FA764DFBF0FDBD" box="[988,1098,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C377C729E3FA764DFBF0FDBD" box="[988,1098,604,629]" italics="true" pageId="89" pageNumber="90">S. flavida</emphasis>
</taxonomicName>
, but differs in having shorter proximal portion of spermatheca (
<figureCitation id="729A2A17C377C729E33B7695FC3EFD56" box="[797,900,644,670]" captionStart="FIGURES 356 ­ 358" captionStartId="88.[264,382,1791,1815]" captionTargetBox="[333,1254,860,1773]" captionTargetId="figure@88.[322,1264,844,1781]" captionTargetPageId="88" captionText="FIGURES 356 ­ 358. Abdominal segment 8 and spermatheca of Seeversiella adusta Gusarov, sp. n. (holotype). 356 female tergum 8; 357 female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356 ­ 357), 0.1 mm (358)." httpUri="https://zenodo.org/record/156482/files/figure.png" pageId="89" pageNumber="90">Fig. 358</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C377C729E11E76BDFE4BFD0E" blockId="89.[264,1324,283,830]" box="[312,497,684,710]" pageId="89" pageNumber="90">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C377C729E11E76C2FE9CFCF6" pageId="89" pageNumber="90" type="distribution">
<paragraph id="EA1E3692C377C729E11E76C2FBCDFD26" blockId="89.[264,1324,283,830]" box="[312,1143,723,750]" pageId="89" pageNumber="90">
<emphasis id="D8D5EA80C377C729E11E76C2FE62FD25" bold="true" box="[312,472,723,749]" pageId="89" pageNumber="90">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C377C729E35476C5FC49FD26" box="[882,1011,724,750]" name="Costa Rica" pageId="89" pageNumber="90">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C377C729E42576C5FBD0FD26" box="[1027,1130,724,750]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="89" pageNumber="90">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C377C729E11E76EAFE9CFCF6" blockId="89.[264,1324,283,830]" pageId="89" pageNumber="90">
<emphasis id="D8D5EA80C377C729E11E76EAFDBCFCDD" bold="true" box="[312,518,763,789]" pageId="89" pageNumber="90">Natural History.</emphasis>
<taxonomicName id="2DA14D11C377C729E22976EDFD4EFCDD" box="[527,756,764,789]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C377C729E22976EDFD4EFCDD" box="[527,756,764,789]" italics="true" pageId="89" pageNumber="90">Seeversiella adusta</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C377C729E4D676EDFE98FCF6" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="89" pageNumber="90" unit="m" value="3300.0">3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,298 @@
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<mods:title id="9125364C35ED1749BA2088D4FB77066F">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="9DEF2D6BA2017D7AEC1A571A185CCFBD">Gusarov, Vladimir I.</mods:namePart>
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<subSubSection id="A2BB6519C377C729E12E778AFC70FC7D" box="[264,970,923,950]" pageId="89" pageNumber="90" type="key">
<paragraph id="EA1E3692C377C729E12E778AFC70FC7D" blockId="89.[264,970,923,950]" box="[264,970,923,950]" pageId="89" pageNumber="90">
<heading id="B15681FEC377C729E12E778AFC70FC7D" bold="true" box="[264,970,923,950]" fontSize="11" level="1" pageId="89" pageNumber="90" reason="1">
<emphasis id="D8D5EA80C377C729E12E778AFC70FC7D" bold="true" box="[264,970,923,950]" pageId="89" pageNumber="90">
<key id="9E3061A3C377C729E12E778AFC70FC7D" box="[264,970,923,950]" pageId="89" pageNumber="90">
<keyStep id="51552E37C377C729E12E778AFC70FC7D" box="[264,970,923,950]" pageId="89" pageNumber="90">
<keyLead id="515095A7C377C729E12E778AFC70FC7D" box="[264,970,923,950]" pageId="89" pageNumber="90">
27.
<taxonomicName id="2DA14D11C377C729E112778DFD2AFC7D" ID-CoL="6YBD4" authority="Gusarov" authorityName="Gusarov" box="[308,656,923,950]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida" status="sp. nov.">
<emphasis id="D8D5EA80C377C729E112778DFDA4FC7E" bold="true" box="[308,542,924,950]" italics="true" pageId="89" pageNumber="90">Seeversiella flavida</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC377C729E2BB778AFD5AFC7D" box="[669,736,923,949]" pageId="89" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C377C729E2C9778AFCE8FC7D" box="[751,850,923,949]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="89" pageNumber="90">Figs. 19</figureCitation>
,
<figureCitation id="729A2A17C377C729E379778AFC78FC7D" box="[863,962,923,949]" captionStart-0="FIGURES 359 ­ 362" captionStart-1="FIGURES 363 ­ 371" captionStartId-0="90.[264,382,1323,1347]" captionStartId-1="91.[264,383,1333,1357]" captionTargetBox-0="[295,1288,324,1290]" captionTargetBox-1="[306,1292,428,1296]" captionTargetId-0="figure@90.[288,1299,313,1297]" captionTargetId-1="figure@91.[284,1303,409,1306]" captionTargetPageId-0="90" captionTargetPageId-1="91" captionText-0="FIGURES 359 ­ 362. Abdominal segment 8 of Seeversiella flavida Gusarov, sp. n. (359 ­ 360, holotype; 361 ­ 362, paratype from Cerro Buenavista, Costa Rica). 359 male tergum 8; 360 male sternum 8; 361 female tergum 8; 362 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri-0="https://zenodo.org/record/156483/files/figure.png" httpUri-1="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">359­371</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C377C729E12E77F7FDCAFAD1" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="EA1E3692C377C729E12E77F7FECFFB99" blockId="89.[264,1324,998,1864]" pageId="89" pageNumber="90">
<emphasis id="D8D5EA80C377C729E12E77F7FE04FBC8" bold="true" box="[264,446,998,1024]" pageId="89" pageNumber="90">
<typeStatus id="351A8830C377C729E12E77F7FEFFFBC8" box="[264,325,998,1024]" pageId="89" pageNumber="90">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C377C729E1EA77F6FD86FBC9" box="[460,572,999,1025]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C377C729E24C77F7FBA6FBC8" bold="true" box="[618,1052,998,1024]" pageId="89" pageNumber="90">
<collectingCountry id="92B67602C377C729E24C77F7FC99FBC8" box="[618,803,998,1024]" name="Costa Rica" pageId="89" pageNumber="90">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C377C729E224701EFDC4FBE1" box="[514,638,1039,1065]" direction="north" orientation="latitude" pageId="89" pageNumber="90" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C377C729E2A0701EFCACFBE1" box="[646,790,1039,1065]" direction="west" orientation="longitude" pageId="89" pageNumber="90" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C377C729E304701EFCC5FBE1" box="[802,895,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="89" pageNumber="90" unit="m" value="3200.0">3200 m</quantity>
, leaf litter (R.Anderson),
<date id="9E1F1052C377C729E48B701EFA91FBE1" box="[1197,1323,1039,1065]" pageId="89" pageNumber="90" value="1997-06-18">18.vi.1997</date>
(
<collectionCode id="8CB0AE57C377C729E1367026FED2FB99" box="[272,360,1079,1105]" pageId="89" pageNumber="90">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C377C729E11E704EFDCAFAD1" blockId="89.[264,1324,998,1864]" pageId="89" pageNumber="90">
<typeStatus id="351A8830C377C729E11E704EFE15FBB1" box="[312,431,1119,1145]" pageId="89" pageNumber="90" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C377C729E1EE704FFCABFBB0" bold="true" box="[456,785,1118,1144]" pageId="89" pageNumber="90">
<collectingCountry id="92B67602C377C729E1EE704FFD3FFBB0" box="[456,645,1118,1144]" name="Costa Rica" pageId="89" pageNumber="90">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC377C729E301704EFC07FBB1" box="[807,957,1119,1145]" pageId="89" pageNumber="90" type="generic">8 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C377C729E484704EFAB6FBB1" box="[1186,1292,1119,1145]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC377C729E53D704EFE3FFB69" pageId="89" pageNumber="90" type="generic">6 specimens</specimenCount>
, Cerro Buenavista,
<geoCoordinate id="8F955055C377C729E2527096FD4AFB69" box="[628,752,1159,1185]" direction="north" orientation="latitude" pageId="89" pageNumber="90" precision="15" value="9.550556">9°33'02&quot;N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C377C729E4137096FB08FB69" box="[1077,1202,1159,1185]" direction="north" orientation="latitude" pageId="89" pageNumber="90" precision="15" value="9.717222">9°43'02&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C377C729E16270BEFE6FFB01" box="[324,469,1199,1225]" direction="west" orientation="longitude" pageId="89" pageNumber="90" precision="15" value="-83.75389">83°45'14&quot;W</geoCoordinate>
,
<quantity id="2D599B77C377C729E1C770BEFD86FB01" box="[481,572,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="89" pageNumber="90" unit="m" value="3300.0">3300 m</quantity>
, subparano litter (R.Anderson),
<date id="9E1F1052C377C729E39270BEFB94FB01" box="[948,1070,1199,1225]" pageId="89" pageNumber="90" value="1998-02-15">15.ii.1998</date>
;
<specimenCount id="FCA7FD1BC377C729E41F70BEFB61FB01" box="[1081,1243,1199,1225]" pageId="89" pageNumber="90" type="generic">23 specimens</specimenCount>
, Cerro de la Muerte, Pan American Highway, km 89,
<quantity id="2D599B77C377C729E30570C6FCC5FB39" box="[803,895,1239,1265]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="89" pageNumber="90" unit="m" value="3300.0">3300 m</quantity>
, elfin bamboo forest litter (R.Anderson),
<date id="9E1F1052C377C729E16F70EEFE7AFAD1" box="[329,448,1279,1305]" pageId="89" pageNumber="90" value="1996-02-10">10.ii.1996</date>
(all ­
<collectionCode id="8CB0AE57C377C729E22B70EEFDDEFAD1" box="[525,612,1279,1305]" pageId="89" pageNumber="90">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C377C729E11E7137FEFFF960" pageId="89" pageNumber="90" type="diagnosis">
<paragraph id="EA1E3692C377C729E11E7137FDEAF9C1" blockId="89.[264,1324,998,1864]" pageId="89" pageNumber="90">
<emphasis id="D8D5EA80C377C729E11E7137FE0DFA88" bold="true" box="[312,439,1318,1344]" pageId="89" pageNumber="90">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C377C729E1E87136FD02FA88" box="[462,696,1319,1344]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C377C729E1E87136FD02FA88" box="[462,696,1319,1344]" italics="true" pageId="89" pageNumber="90">Seeversiella flavida</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C377C729E4F57136FEFDFAA0" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C377C729E4F57136FEFDFAA0" italics="true" pageId="89" pageNumber="90">Seeversiella</emphasis>
</taxonomicName>
by having uniformly brownish yellow body; small eyes (temples 4.0­5.0 times as long as eyes); elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking medial emargination at the posterior margin of the male tergum 8 (
<figureCitation id="729A2A17C377C729E4A2718EFB57FA71" box="[1156,1261,1439,1465]" captionStart="FIGURES 359 ­ 362" captionStartId="90.[264,382,1323,1347]" captionTargetBox="[295,1288,324,1290]" captionTargetId="figure@90.[288,1299,313,1297]" captionTargetPageId="90" captionText="FIGURES 359 ­ 362. Abdominal segment 8 of Seeversiella flavida Gusarov, sp. n. (359 ­ 360, holotype; 361 ­ 362, paratype from Cerro Buenavista, Costa Rica). 359 male tergum 8; 360 male sternum 8; 361 female tergum 8; 362 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156483/files/figure.png" pageId="89" pageNumber="90">Fig. 359</figureCitation>
); tergum 8 with three pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C377C729E45D71D6FA98FA29" box="[1147,1314,1479,1505]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Figs. 363­370</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C377C729E1FA71FEFDF9F9C1" box="[476,579,1519,1545]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Fig. 371</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C377C729E11E7206FEFFF960" blockId="89.[264,1324,998,1864]" pageId="89" pageNumber="90">
<taxonomicName id="2DA14D11C377C729E11E7206FDA5F9F8" box="[312,543,1559,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C377C729E11E7206FDA5F9F8" box="[312,543,1559,1584]" italics="true" pageId="89" pageNumber="90">Seeversiella flavida</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C377C729E29A7206FCC6F9F8" box="[700,892,1559,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C377C729E29A7206FCC6F9F8" box="[700,892,1559,1584]" italics="true" pageId="89" pageNumber="90">S. impressicollis</emphasis>
</taxonomicName>
,
<taxonomicName id="2DA14D11C377C729E3AD7206FBA5F9F8" box="[907,1055,1559,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C377C729E3AD7206FBA5F9F8" box="[907,1055,1559,1584]" italics="true" pageId="89" pageNumber="90">S. sulcicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C377C729E47D7206FA91F9F8" box="[1115,1323,1559,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C377C729E47D7206FA91F9F8" box="[1115,1323,1559,1584]" italics="true" pageId="89" pageNumber="90">S. microphthalma</emphasis>
</taxonomicName>
in having smaller body size; pronotal punctation not asperate; tergum 8 with three pairs of macrosetae; and by different shape of aedeagus (
<figureCitation id="729A2A17C377C729E3477277FBB0F948" box="[865,1034,1638,1664]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Figs. 363­370</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C377C729E4DD7277FE82F960" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Fig. 371</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C377C72CE11E72A6FDC4FD0E" lastPageId="92" lastPageNumber="93" pageId="89" pageNumber="90" type="description">
<paragraph id="EA1E3692C377C729E11E72A6FAAAF880" blockId="89.[264,1324,998,1864]" pageId="89" pageNumber="90">
<taxonomicName id="2DA14D11C377C729E11E72A6FD98F918" box="[312,546,1719,1744]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C377C729E11E72A6FD98F918" box="[312,546,1719,1744]" italics="true" pageId="89" pageNumber="90">Seeversiella flavida</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C377C729E2E072A6FCC0F918" box="[710,890,1719,1744]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C377C729E2E072A6FCC0F918" box="[710,890,1719,1744]" italics="true" pageId="89" pageNumber="90">S. geostiboides</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed towards the midline and obliquely posteriorly; tergum 8 with three pairs of macrosetae; wider apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C377C729E4957317FE8DF880" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Figs. 363­ 364</figureCitation>
; 348­349); and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C377C729E473733FFB7DF880" box="[1109,1223,1838,1864]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="89" pageNumber="90">Figs. 371</figureCitation>
; 355).
</paragraph>
<caption id="BEDE661AC374C72AE12E713AFBEAFA40" httpUri="https://zenodo.org/record/156483/files/figure.png" pageId="90" pageNumber="91" targetBox="[295,1288,324,1290]" targetPageId="90">
<paragraph id="EA1E3692C374C72AE12E713AFBEAFA40" blockId="90.[264,1322,1323,1416]" pageId="90" pageNumber="91">
<emphasis id="D8D5EA80C374C72AE12E713AFE64FA8B" bold="true" box="[264,478,1323,1347]" pageId="90" pageNumber="91">FIGURES 359­362</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C374C72AE324713DFB8FFA8B" authority="Gusarov" authorityName="Gusarov" box="[770,1077,1324,1347]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavida" status="sp. nov.">
<emphasis id="D8D5EA80C374C72AE324713DFC6EFA8B" box="[770,980,1324,1347]" italics="true" pageId="90" pageNumber="91">Seeversiella flavida</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C374C72AE466713AFBC6FA8B" bold="true" box="[1088,1148,1323,1347]" pageId="90" pageNumber="91">
<taxonomicNameLabel id="C3E657FBC374C72AE466713AFBC6FA8B" box="[1088,1148,1323,1347]" pageId="90" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(359­360, holotype; 361­362, paratype from Cerro Buenavista, Costa Rica). 359 male tergum 8; 360 male sternum 8; 361 female tergum 8; 362 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C374C72AE11E71C8FD8EF9A3" blockId="90.[264,1324,1497,1883]" pageId="90" pageNumber="91">
<taxonomicName id="2DA14D11C374C72AE11E71C8FDA5FA3A" box="[312,543,1497,1522]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C374C72AE11E71C8FDA5FA3A" box="[312,543,1497,1522]" italics="true" pageId="90" pageNumber="91">Seeversiella flavida</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C374C72AE29A71C8FCDBFA3A" box="[700,865,1497,1522]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C374C72AE29A71C8FCDBFA3A" box="[700,865,1497,1522]" italics="true" pageId="90" pageNumber="91">S. paramoana</emphasis>
</taxonomicName>
in having tergum 8 with three pairs of macrosetae; and obtuse apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C374C72AE42A7210FB0FF9D3" box="[1036,1205,1537,1563]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="90" pageNumber="91">Figs. 363­364</figureCitation>
).
<taxonomicName id="2DA14D11C374C72AE11E7238FD9AF98A" box="[312,544,1577,1602]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C374C72AE11E7238FD9AF98A" box="[312,544,1577,1602]" italics="true" pageId="90" pageNumber="91">Seeversiella flavida</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C374C72AE2987238FC91F98A" box="[702,811,1577,1602]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C374C72AE2987238FC91F98A" box="[702,811,1577,1602]" italics="true" pageId="90" pageNumber="91">S. adusta</emphasis>
</taxonomicName>
in having longer proximal portion of spermatheca (
<figureCitation id="729A2A17C374C72AE15C7240FE51F9A3" box="[378,491,1617,1643]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="90" pageNumber="91">Figs. 371</figureCitation>
; 358).
</paragraph>
<paragraph id="EA1E3692C374C72BE11E7268FB70FE96" blockId="90.[264,1324,1497,1883]" lastBlockId="91.[264,1323,284,350]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
<taxonomicName id="2DA14D11C374C72AE11E7268FDA4F95A" box="[312,542,1657,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C374C72AE11E7268FDA4F95A" box="[312,542,1657,1682]" italics="true" pageId="90" pageNumber="91">Seeversiella flavida</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C374C72AE2917268FC91F95A" box="[695,811,1657,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C374C72AE2917268FC91F95A" box="[695,811,1657,1682]" italics="true" pageId="90" pageNumber="91">S. minima</emphasis>
</taxonomicName>
in having tergum 8 with three pairs of macrosetae; proximal seta of the apex of paramere is approximately as long as the other three setae (
<figureCitation id="729A2A17C374C72AE17072D8FE7BF92B" box="[342,449,1737,1763]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="90" pageNumber="91">Fig. 370</figureCitation>
); apex of median lobe in parameral view narrower (
<figureCitation id="729A2A17C374C72AE41D72D8FB5CF92B" box="[1083,1254,1737,1763]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="90" pageNumber="91">Figs. 363­364</figureCitation>
; 179­ 180); spermatheca with thin proximal portion and with umbilicus (
<figureCitation id="729A2A17C374C72AE43372E0FB3DF8C3" box="[1045,1159,1777,1803]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="90" pageNumber="91">Figs. 371</figureCitation>
; 184).
<emphasis id="D8D5EA80C374C72AE11E7309FE6AF8FA" bold="true" box="[312,464,1816,1842]" pageId="90" pageNumber="91">Description.</emphasis>
Length
<quantity id="2D599B77C374C72AE21F7308FD7EF8FB" box="[569,708,1817,1843]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="90" pageNumber="91" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2­2.5 mm</quantity>
. Body uniformly brownish­yellow to light brown. Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation. Temples 4.0­5.0 times as long as eyes. Antennal article 2 longer than 3, article 4 slightly transverse, 5­10 strongly transverse (ratio 1.6­2.0).
</paragraph>
<caption id="BEDE661AC375C72BE12E7124FB3DFA31" httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="91" pageNumber="92" targetBox="[306,1292,428,1296]" targetPageId="91">
<paragraph id="EA1E3692C375C72BE12E7124FB3DFA31" blockId="91.[264,1325,1333,1529]" pageId="91" pageNumber="92">
<emphasis id="D8D5EA80C375C72BE12E7124FE5BFA85" bold="true" box="[264,481,1333,1357]" pageId="91" pageNumber="92">FIGURES 363­371</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C375C72BE2AF7124FC79FA84" authority="Gusarov" authorityName="Gusarov" box="[649,963,1333,1356]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="flavida" status="sp. nov.">
<emphasis id="D8D5EA80C375C72BE2AF7124FCE4FA84" box="[649,862,1333,1356]" italics="true" pageId="91" pageNumber="92">Seeversiella flavida</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C375C72BE3F77124FBABFA85" bold="true" box="[977,1041,1333,1357]" pageId="91" pageNumber="92">
<taxonomicNameLabel id="C3E657FBC375C72BE3F77124FBABFA85" box="[977,1041,1333,1357]" pageId="91" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(363­367, 370, holotype; 368­369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368­371).
</paragraph>
</caption>
<paragraph id="EA1E3692C375C72BE11E723CFDE8F897" blockId="91.[264,1324,1581,1887]" pageId="91" pageNumber="92">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C375C72BE40D723CFB6CF98F" box="[1067,1238,1581,1607]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="91" pageNumber="92" unit="mm" value="0.41" valueMax="0.43" valueMin="0.39">0.39­0.43 mm</quantity>
, length
<quantity id="2D599B77C375C72BE12E7244FE0BF9A7" box="[264,433,1621,1647]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.5999999999999996" pageId="91" pageNumber="92" unit="mm" value="0.375" valueMax="0.39" valueMin="0.36">0.36­0.39 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine punctation, distance between punctures equals 2­3 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.7 times wider than long, glossy, with poorly visible microsculpture, with fine and asperate punctation, distance between punctures equals 1­ 1.5 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C372C72CE11E750DFD42FE1E" blockId="92.[264,1325,284,830]" pageId="92" pageNumber="93">
Abdominal terga glossy, with microsculpture consisting of strongly transverse meshes or transverse waves and with fine punctation, on terga 3­5 distance between punctures equals 1­3 times their diameter; on tergum 7 distance between punctures equals 3­7 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C372C72CE21875ADFD50FE1E" box="[574,746,444,470]" captionStart="FIGURES 359 ­ 362" captionStartId="90.[264,382,1323,1347]" captionTargetBox="[295,1288,324,1290]" captionTargetId="figure@90.[288,1299,313,1297]" captionTargetPageId="90" captionText="FIGURES 359 ­ 362. Abdominal segment 8 of Seeversiella flavida Gusarov, sp. n. (359 ­ 360, holotype; 361 ­ 362, paratype from Cerro Buenavista, Costa Rica). 359 male tergum 8; 360 male sternum 8; 361 female tergum 8; 362 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156483/files/figure.png" pageId="92" pageNumber="93">Figs. 359, 361</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C372C72CE11E75F5FCCAFDBE" blockId="92.[264,1325,284,830]" pageId="92" pageNumber="93">
In males lateral portions of posterior margin of tergum 3 extending as short and obtuse projections (
<figureCitation id="729A2A17C372C72CE1BC761DFE4AFDEE" box="[410,496,524,550]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="92" pageNumber="93">Fig. 19</figureCitation>
), up to 1/3 as long as tergum (measured medially); tergum
<quantity id="2D599B77C372C72CE486761DFB76FDEE" box="[1184,1228,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="92" pageNumber="93" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina up to half as long as tergum; posterior margin of tergum 8 without clear medial emargination (
<figureCitation id="729A2A17C372C72CE2DA764DFCDEFDBE" box="[764,868,604,630]" captionStart="FIGURES 359 ­ 362" captionStartId="90.[264,382,1323,1347]" captionTargetBox="[295,1288,324,1290]" captionTargetId="figure@90.[288,1299,313,1297]" captionTargetPageId="90" captionText="FIGURES 359 ­ 362. Abdominal segment 8 of Seeversiella flavida Gusarov, sp. n. (359 ­ 360, holotype; 361 ­ 362, paratype from Cerro Buenavista, Costa Rica). 359 male tergum 8; 360 male sternum 8; 361 female tergum 8; 362 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156483/files/figure.png" pageId="92" pageNumber="93">Fig. 359</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C372C72CE11E7695FD26FD56" blockId="92.[264,1325,284,830]" box="[312,668,644,670]" pageId="92" pageNumber="93">
Aedeagus as in
<figureCitation id="729A2A17C372C72CE1D77695FD22FD56" box="[497,664,644,670]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="92" pageNumber="93">Figs. 363­370</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C372C72CE11E76BDFDC4FD0E" blockId="92.[264,1325,284,830]" box="[312,638,684,710]" pageId="92" pageNumber="93">
Spermatheca as in
<figureCitation id="729A2A17C372C72CE23376BDFDC0FD0E" box="[533,634,684,710]" captionStart="FIGURES 363 ­ 371" captionStartId="91.[264,383,1333,1357]" captionTargetBox="[306,1292,428,1296]" captionTargetId="figure@91.[284,1303,409,1306]" captionTargetPageId="91" captionText="FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 median lobe, parameral view; 364 apex of median lobe, parameral view; 365 median lobe, lateral view; 366 apex of median lobe, lateral view; 367 details of internal sac retracted into median lobe, lateral view; 368 copulatory piece, lateral view; 369 everted internal sac, parameral view; 370 apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371)." httpUri="https://zenodo.org/record/156484/files/figure.png" pageId="92" pageNumber="93">Fig. 371</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C372C72CE11E76C2FED2FCF6" pageId="92" pageNumber="93" type="distribution">
<paragraph id="EA1E3692C372C72CE11E76C2FBCDFD26" blockId="92.[264,1325,284,830]" box="[312,1143,723,750]" pageId="92" pageNumber="93">
<emphasis id="D8D5EA80C372C72CE11E76C2FE62FD25" bold="true" box="[312,472,723,749]" pageId="92" pageNumber="93">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C372C72CE35476C5FC49FD26" box="[882,1011,724,750]" name="Costa Rica" pageId="92" pageNumber="93">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C372C72CE42576C5FBD0FD26" box="[1027,1130,724,750]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="92" pageNumber="93">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C372C72CE11E76EAFED2FCF6" blockId="92.[264,1325,284,830]" pageId="92" pageNumber="93">
<emphasis id="D8D5EA80C372C72CE11E76EAFDBCFCDD" bold="true" box="[312,518,763,789]" pageId="92" pageNumber="93">Natural History.</emphasis>
<taxonomicName id="2DA14D11C372C72CE23E76EDFCBAFCDD" box="[536,768,764,789]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C372C72CE23E76EDFCBAFCDD" box="[536,768,764,789]" italics="true" pageId="92" pageNumber="93">Seeversiella flavida</emphasis>
</taxonomicName>
was collected in leaf litter at altitude of
<quantity id="2D599B77C372C72CE4C176EDFEDEFCF6" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="92" pageNumber="93" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">3200­ 3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,270 @@
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<mods:title id="75B875CDE60CC34593E2DA6A244321CF">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C37BC725E12E764AFC64FDBD" box="[264,990,603,630]" pageId="85" pageNumber="86" type="key">
<paragraph id="EA1E3692C37BC725E12E764AFC64FDBD" blockId="85.[264,990,603,630]" box="[264,990,603,630]" pageId="85" pageNumber="86">
<heading id="B15681FEC37BC725E12E764AFC64FDBD" bold="true" box="[264,990,603,630]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
<emphasis id="D8D5EA80C37BC725E12E764AFC64FDBD" bold="true" box="[264,990,603,630]" pageId="85" pageNumber="86">
<key id="9E3061A3C37BC725E12E764AFC64FDBD" box="[264,990,603,630]" pageId="85" pageNumber="86">
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25.
<taxonomicName id="2DA14D11C37BC725E112764DFD6AFDBD" ID-CoL="6YBD2" authority="Gusarov" authorityName="Gusarov" box="[308,720,603,630]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C37BC725E112764DFDE4FDBE" bold="true" box="[308,606,604,630]" italics="true" pageId="85" pageNumber="86">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37BC725E2FB764AFCA5FDBD" box="[733,799,603,629]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C37BC725E309764AFC6DFDBD" box="[815,983,603,629]" captionStart-0="FIGURES 344 ­ 347" captionStart-1="FIGURES 348 ­ 355" captionStartId-0="85.[264,382,1733,1757]" captionStartId-1="86.[264,383,1647,1671]" captionTargetBox-0="[305,1286,853,1699]" captionTargetBox-1="[290,1290,620,1609]" captionTargetId-0="figure@85.[292,1295,844,1706]" captionTargetId-1="figure@86.[284,1303,609,1620]" captionTargetPageId-0="85" captionTargetPageId-1="86" captionText-0="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri-0="https://zenodo.org/record/156480/files/figure.png" httpUri-1="https://zenodo.org/record/156481/files/figure.png" pageId="85" pageNumber="86">Figs. 344­355</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37BC725E12E76B7FB6AFCD9" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="EA1E3692C37BC725E12E76B7FBDAFD21" blockId="85.[264,1323,678,785]" pageId="85" pageNumber="86">
<emphasis id="D8D5EA80C37BC725E12E76B7FE04FD08" bold="true" box="[264,446,678,704]" pageId="85" pageNumber="86">
<typeStatus id="351A8830C37BC725E12E76B7FEFFFD08" box="[264,325,678,704]" pageId="85" pageNumber="86">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C37BC725E1ED76B6FD81FD09" box="[459,571,679,705]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
,,
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<collectingCountry id="92B67602C37BC725E24E76B7FC9BFD08" box="[616,801,678,704]" name="Costa Rica" pageId="85" pageNumber="86">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
2.0 km E Villa Mills,
<geoCoordinate id="8F955055C37BC725E48976B6FA91FD09" box="[1199,1323,679,705]" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C37BC725E12E76DEFE22FD21" box="[264,408,719,745]" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="15" value="-83.69722">83°41'50&quot;W</geoCoordinate>
,
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, oak forest litter (R.Anderson),
<date id="9E1F1052C37BC725E35376DEFC56FD21" box="[885,1004,719,745]" pageId="85" pageNumber="86" value="1998-02-15">15.ii.1998</date>
(
<collectionCode id="8CB0AE57C37BC725E3DA76DEFBEEFD21" box="[1020,1108,719,745]" pageId="85" pageNumber="86">KSEM</collectionCode>
).
</paragraph>
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<typeStatus id="351A8830C37BC725E11E76E6FE14FCD9" box="[312,430,759,785]" pageId="85" pageNumber="86" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C37BC725E1E676E7FDC0FCD8" box="[448,634,758,784]" name="Costa Rica" pageId="85" pageNumber="86">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, same data as the
<typeStatus id="351A8830C37BC725E3D076E6FBE7FCD9" box="[1014,1117,759,785]" pageId="85" pageNumber="86" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C37BC725E44A76E6FB7EFCD9" box="[1132,1220,759,785]" pageId="85" pageNumber="86">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
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<emphasis id="D8D5EA80C37BC725E12E72D4FE64F915" bold="true" box="[264,478,1733,1757]" pageId="85" pageNumber="86">FIGURES 344­347</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C37BC725E32272D4FBC9F914" authority="Gusarov" authorityName="Gusarov" box="[772,1139,1733,1756]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C37BC725E32272D4FBABF914" box="[772,1041,1733,1756]" italics="true" pageId="85" pageNumber="86">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C37BC725E45972D4FB00F915" bold="true" box="[1151,1210,1733,1757]" pageId="85" pageNumber="86">sp. n.</emphasis>
</taxonomicNameLabel>
(344­345, holotype; 346­347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C378C727E11E750AFB4DFE1E" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="diagnosis">
<paragraph id="EA1E3692C378C726E11E750AFA91FE36" blockId="86.[264,1324,283,510]" pageId="86" pageNumber="87">
<emphasis id="D8D5EA80C378C726E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="86" pageNumber="87">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C378C726E1EE750DFD48FEFD" box="[456,754,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C378C726E1EE750DFD48FEFD" box="[456,754,284,309]" italics="true" pageId="86" pageNumber="87">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C378C726E4DE750DFED7FE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C378C726E4DE750DFED7FE95" italics="true" pageId="86" pageNumber="87">Seeversiella</emphasis>
</taxonomicName>
by having uniformly brown body; small eyes (temples 3.5­4.0 times as long as eyes); pronotal pubescence in lateral portions of the disc directed posteriorly; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at the posterior margin of male tergum 8 (
<figureCitation id="729A2A17C378C726E34E75ADFC6AFE1E" box="[872,976,444,470]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="86" pageNumber="87">Fig. 344</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C378C726E30875F5FC69FE36" box="[814,979,484,510]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87">Figs. 348­354</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C378C726E49175F5FAA4FE36" box="[1207,1310,484,510]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87">Fig. 355</figureCitation>
).
</paragraph>
<caption id="BEDE661AC378C726E12E727EFC5AF8FB" httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="86" pageNumber="87" targetBox="[290,1290,620,1609]" targetPageId="86">
<paragraph id="EA1E3692C378C726E12E727EFC5AF8FB" blockId="86.[264,1324,1647,1843]" pageId="86" pageNumber="87">
<emphasis id="D8D5EA80C378C726E12E727EFE5BF94F" bold="true" box="[264,481,1647,1671]" pageId="86" pageNumber="87">FIGURES 348­355</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C378C726E2A1727EFC46F94E" authority="Gusarov" authorityName="Gusarov" box="[647,1020,1647,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="geostiboides" status="sp. nov.">
<emphasis id="D8D5EA80C378C726E2A1727EFC2DF94E" box="[647,919,1647,1670]" italics="true" pageId="86" pageNumber="87">Seeversiella geostiboides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC378C726E42C727EFBF3F94F" box="[1034,1097,1647,1671]" pageId="86" pageNumber="87" rank="species">
<emphasis id="D8D5EA80C378C726E42C727EFBF3F94F" bold="true" box="[1034,1097,1647,1671]" pageId="86" pageNumber="87">sp. n.</emphasis>
</taxonomicNameLabel>
(348­354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353­354), 0.1 mm (349, 351­352, 355).
</paragraph>
</caption>
<paragraph id="EA1E3692C379C727E11E750DFB4DFE1E" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E750DFDD8FEFD" box="[312,610,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E750DFDD8FEFD" box="[312,610,284,309]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C379C727E38C750DFBD6FEFD" box="[938,1132,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C379C727E38C750DFBD6FEFD" box="[938,1132,284,309]" italics="true" pageId="87" pageNumber="88">S. impressicollis</emphasis>
</taxonomicName>
by not asperate pronotal punctation; in males pronotum without broad medial impression, lateral portions of posterior margin of tergum 3 extending as short and blunt projections (as in
<figureCitation id="729A2A17C379C727E4E2757DFAA6FE4E" box="[1220,1308,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="87" pageNumber="88">Fig. 19</figureCitation>
), and tergum 7 with medial carina; the apex of median lobe in parameral view more rounded (
<figureCitation id="729A2A17C379C727E13775ADFE02FE1E" box="[273,440,444,470]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
; 311­312); and spermatheca with larger distal portion (
<figureCitation id="729A2A17C379C727E41A75ADFB17FE1E" box="[1084,1197,444,470]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 320).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C379C728E11E75F5FDC7FE4E" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
<paragraph id="EA1E3692C379C727E11E75F5FDAAFDBE" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E75F5FDDCFE35" box="[312,614,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E75F5FDDCFE35" box="[312,614,484,509]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C379C727E3E275F5FBE5FE35" box="[964,1119,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C379C727E3E275F5FBE5FE35" box="[964,1119,484,509]" italics="true" pageId="87" pageNumber="88">S. sulcicollis</emphasis>
</taxonomicName>
by not asperate pronotal punctation; in males pronotum without medial furrow and posterior margin of tergum 8 without denticles; the apex of median lobe without divergent carinae (in parameral view) (
<figureCitation id="729A2A17C379C727E17D764DFDB9FDBE" box="[347,515,604,630]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C379C727E11E7695FC14FCDE" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E7695FDDBFD55" box="[312,609,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E7695FDDBFD55" box="[312,609,644,669]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C379C727E3267695FC6BFD55" box="[768,977,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C379C727E3267695FC6BFD55" box="[768,977,644,669]" italics="true" pageId="87" pageNumber="88">S. microphthalma</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C379C727E42B7695FB08FD55" box="[1037,1202,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C379C727E42B7695FB08FD55" box="[1037,1202,644,669]" italics="true" pageId="87" pageNumber="88">S. paramoana</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed posteriorly; in narrower apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C379C727E28676C5FCF0FD26" box="[672,842,724,750]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 348­349</figureCitation>
; 335­336; 377­378); and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C379C727E29E76EDFC93FCDE" box="[696,809,764,790]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 341; 381).
</paragraph>
<paragraph id="EA1E3692C379C727E11E7735FB7DFC45" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
<taxonomicName id="2DA14D11C379C727E11E7735FDE5FCF5" box="[312,607,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C379C727E11E7735FDE5FCF5" box="[312,607,804,829]" italics="true" pageId="87" pageNumber="88">Seeversiella geostiboides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C379C727E2DE7735FCDCFCF5" box="[760,870,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C379C727E2DE7735FCDCFCF5" box="[760,870,804,829]" italics="true" pageId="87" pageNumber="88">S. flavida</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C379C727E3B97735FBB0FCF5" box="[927,1034,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C379C727E3B97735FBB0FCF5" box="[927,1034,804,829]" italics="true" pageId="87" pageNumber="88">S. adusta</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed posteriorly; tergum 8 with four pairs of macrosetae; and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C379C727E3F77765FBF8FC46" box="[977,1090,884,910]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="87" pageNumber="88">Figs. 355</figureCitation>
; 358; 371).
</paragraph>
<paragraph id="EA1E3692C379C727E11E778AFC05FC7E" blockId="87.[264,1324,284,1870]" box="[312,959,923,950]" pageId="87" pageNumber="88">
<emphasis id="D8D5EA80C379C727E11E778AFE6AFC7D" bold="true" box="[312,464,923,949]" pageId="87" pageNumber="88">Description.</emphasis>
Length
<quantity id="2D599B77C379C727E21F778DFD2CFC7E" box="[569,662,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="87" pageNumber="88" unit="mm" value="2.6">2.6 mm</quantity>
. Body uniformly brown.
</paragraph>
<paragraph id="EA1E3692C379C727E11E77D5FD49FB9E" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals 2 times their diameter. Temples 3.5­4.0 times as long as eyes. Antennal article 2 as long as 3, article 4 slightly transverse, 5­10 transverse (ratio 1.6­2.0).</paragraph>
<paragraph id="EA1E3692C379C727E11E7075FEE8FA76" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C379C727E40F7075FB69FBB6" box="[1065,1235,1124,1150]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="87" pageNumber="88" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47­0.50 mm</quantity>
, length
<quantity id="2D599B77C379C727E12E709DFE0AFB6E" box="[264,432,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.6" metricValueMin="4.3" pageId="87" pageNumber="88" unit="mm" value="0.445" valueMax="0.46" valueMin="0.43">0.43­0.46 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine punctation, distance between punctures equals 1­2 times their diameter. Pronotal pubescence in lateral portions of the disc directed posteriorly. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with poorly visible microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C379C727E11E71DDFE71F966" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 2­3 times their diameter; tergum 7 with microsculpture consisting of isodiametric or transverse meshes and with fine and sparse punctation, distance between punctures equals 3­7 times. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C379C727E1377285FE05F966" box="[273,447,1684,1710]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="87" pageNumber="88">Figs. 344, 346</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C379C727E11E72ADFB01F886" blockId="87.[264,1324,284,1870]" pageId="87" pageNumber="88">
In the only known male the posterior angles of tergum 3 not projecting as spines, lateral portions of posterior margin slightly extending as short and blunt projections (as in
<figureCitation id="729A2A17C379C727E12E731DFEDAF8EE" box="[264,352,1804,1830]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="87" pageNumber="88">Fig. 19</figureCitation>
); tergum
<quantity id="2D599B77C379C727E1F2731DFDB9F8EE" box="[468,515,1804,1830]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="87" pageNumber="88" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina 2/3 as long as the tergum; posterior margin of tergum 8 without clear medial emargination (
<figureCitation id="729A2A17C379C727E4617325FB15F886" box="[1095,1199,1844,1870]" captionStart="FIGURES 344 ­ 347" captionStartId="85.[264,382,1733,1757]" captionTargetBox="[305,1286,853,1699]" captionTargetId="figure@85.[292,1295,844,1706]" captionTargetPageId="85" captionText="FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 male tergum 8; 345 male sternum 8; 346 female tergum 8; 347 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156480/files/figure.png" pageId="87" pageNumber="88">Fig. 344</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E750DFD48FE96" blockId="88.[264,1323,284,510]" pageId="88" pageNumber="89">
Aedeagus as in
<figureCitation id="729A2A17C376C728E1C8750DFD2EFEFE" box="[494,660,284,310]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Figs. 348­354</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C376C728E2587555FD5FFE96" box="[638,741,324,350]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Fig. 352</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E757DFDC7FE4E" blockId="88.[264,1323,284,510]" box="[312,637,364,390]" pageId="88" pageNumber="89">
Spermatheca as in
<figureCitation id="729A2A17C376C728E232757DFDC3FE4E" box="[532,633,364,390]" captionStart="FIGURES 348 ­ 355" captionStartId="86.[264,383,1647,1671]" captionTargetBox="[290,1290,620,1609]" captionTargetId="figure@86.[284,1303,609,1620]" captionTargetPageId="86" captionText="FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 median lobe, parameral view; 349 apex of median lobe, parameral view; 350 median lobe, lateral view; 351 apex of median lobe, lateral view; 352 apex of left paramere; 353 details of internal sac retracted into median lobe, abparameral view; 354 details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355)." httpUri="https://zenodo.org/record/156481/files/figure.png" pageId="88" pageNumber="89">Fig. 355</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C376C728E11E7582FED2FE36" pageId="88" pageNumber="89" type="distribution">
<paragraph id="EA1E3692C376C728E11E7582FBCFFE66" blockId="88.[264,1323,284,510]" box="[312,1141,403,430]" pageId="88" pageNumber="89">
<emphasis id="D8D5EA80C376C728E11E7582FE62FE65" bold="true" box="[312,472,403,429]" pageId="88" pageNumber="89">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C376C728E3547585FC48FE66" box="[882,1010,404,430]" name="Costa Rica" pageId="88" pageNumber="89">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C376C728E4247585FBD3FE66" box="[1026,1129,404,430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="88" pageNumber="89">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C376C728E11E75AAFED2FE36" blockId="88.[264,1323,284,510]" pageId="88" pageNumber="89">
<emphasis id="D8D5EA80C376C728E11E75AAFDBFFE1D" bold="true" box="[312,517,443,469]" pageId="88" pageNumber="89">Natural History.</emphasis>
<taxonomicName id="2DA14D11C376C728E23375ADFC84FE1D" box="[533,830,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C376C728E23375ADFC84FE1D" box="[533,830,444,469]" italics="true" pageId="88" pageNumber="89">Seeversiella geostiboides</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C376C728E12E75F5FEDEFE36" box="[264,356,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="88" pageNumber="89" unit="m" value="2750.0">2750 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,270 @@
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<subSubSection id="A2BB6519C37CC722E12E75AAFBBFFE1D" box="[264,1029,443,470]" pageId="82" pageNumber="83" type="key">
<paragraph id="EA1E3692C37CC722E12E75AAFBBFFE1D" blockId="82.[264,1029,443,470]" box="[264,1029,443,470]" pageId="82" pageNumber="83">
<heading id="B15681FEC37CC722E12E75AAFBBFFE1D" bold="true" box="[264,1029,443,470]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
<emphasis id="D8D5EA80C37CC722E12E75AAFBBFFE1D" bold="true" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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24.
<taxonomicName id="2DA14D11C37CC722E11275ADFD4DFE1D" ID-CoL="6YBCX" authority="Gusarov" authorityName="Gusarov" box="[308,759,443,470]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
<emphasis id="D8D5EA80C37CC722E11275ADFD3FFE1E" bold="true" box="[308,645,444,470]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37CC722E32275AAFCFCFE1D" box="[772,838,443,469]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C37CC722E37375AAFC47FE1D" box="[853,1021,443,469]" captionStart-0="FIGURES 331 ­ 334" captionStart-1="FIGURES 335 ­ 343" captionStartId-0="82.[264,383,1659,1683]" captionStartId-1="83.[264,383,1660,1684]" captionTargetBox-0="[274,1308,735,1617]" captionTargetBox-1="[283,1301,504,1615]" captionTargetId-0="figure@82.[264,1323,724,1632]" captionTargetId-1="figure@83.[270,1318,489,1633]" captionTargetPageId-0="82" captionTargetPageId-1="83" captionText-0="FIGURES 331 ­ 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 ­ 332, holotype; 333 ­ 334, paratype from Villa Mills, Costa Rica). 331 male tergum 8; 332 male sternum 8; 333 female tergum 8; 334 female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri-0="https://zenodo.org/record/156478/files/figure.png" httpUri-1="https://zenodo.org/record/156479/files/figure.png" pageId="82" pageNumber="83">Figs. 331­343</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37CC722E12E7617FB42FDB9" pageId="82" pageNumber="83" type="materials_examined">
<paragraph id="EA1E3692C37CC722E12E7617FC7FFD81" blockId="82.[264,1323,518,625]" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E12E7617FE04FDE8" bold="true" box="[264,446,518,544]" pageId="82" pageNumber="83">
<typeStatus id="351A8830C37CC722E12E7617FEF9FDE8" box="[264,323,518,544]" pageId="82" pageNumber="83">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C37CC722E1F27616FDFEFDE9" box="[468,580,519,545]" pageId="82" pageNumber="83" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C37CC722E2507617FC78FDE8" bold="true" box="[630,962,518,544]" pageId="82" pageNumber="83">
<collectingCountry id="92B67602C37CC722E2507617FC8BFDE8" box="[630,817,518,544]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
<quantity id="2D599B77C37CC722E3F07616FBA9FDE9" box="[982,1043,519,545]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="82" pageNumber="83" unit="km" value="2.0">2 km</quantity>
S Villa Mills,
<quantity id="2D599B77C37CC722E4EE7616FA9CFDE9" box="[1224,1318,519,545]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="82" pageNumber="83" unit="m" value="3000.0">3000 m</quantity>
, ridgetop oak forest litter (R.Anderson),
<date id="9E1F1052C37CC722E2FC763EFCEBFD81" box="[730,849,559,585]" pageId="82" pageNumber="83" value="1996-02-10">10.ii.1996</date>
(
<collectionCode id="8CB0AE57C37CC722E347763EFC03FD81" box="[865,953,559,585]" pageId="82" pageNumber="83">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C37CC722E11E7646FB42FDB9" blockId="82.[264,1323,518,625]" box="[312,1272,598,625]" pageId="82" pageNumber="83">
<typeStatus id="351A8830C37CC722E11E7646FE14FDB9" box="[312,430,599,625]" pageId="82" pageNumber="83" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C37CC722E1E67647FCB9FDB8" bold="true" box="[448,771,598,624]" pageId="82" pageNumber="83">
<collectingCountry id="92B67602C37CC722E1E67647FDC0FDB8" box="[448,634,598,624]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
: San Jose:
</emphasis>
8, same data as the
<typeStatus id="351A8830C37CC722E43B7646FB3EFDB9" box="[1053,1156,599,625]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C37CC722E4B27646FB56FDB9" box="[1172,1260,599,625]" pageId="82" pageNumber="83">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC37CC722E12E726AFBEAF91F" httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="82" pageNumber="83" targetBox="[274,1308,735,1617]" targetPageId="82">
<paragraph id="EA1E3692C37CC722E12E726AFBEAF91F" blockId="82.[264,1323,1659,1751]" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E12E726AFE5AF95B" bold="true" box="[264,480,1659,1683]" pageId="82" pageNumber="83">FIGURES 331­334</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C37CC722E329726AFB27F95A" authority="Gusarov" authorityName="Gusarov" box="[783,1181,1659,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
<emphasis id="D8D5EA80C37CC722E329726AFB80F95A" box="[783,1082,1659,1682]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C37CC722E48C726AFB52F95B" bold="true" box="[1194,1256,1659,1683]" pageId="82" pageNumber="83">sp. n.</emphasis>
</taxonomicNameLabel>
(331­ 332, holotype; 333­334, paratype from Villa Mills, Costa Rica). 331 male tergum 8; 332 male sternum 8; 333 female tergum 8; 334 female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C37CC724E11E733BFEFFFE1D" lastPageId="84" lastPageNumber="85" pageId="82" pageNumber="83" type="diagnosis">
<paragraph id="EA1E3692C37CC723E11E733BFD37FE66" blockId="82.[264,1323,1834,1901]" lastBlockId="83.[264,1324,284,430]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E11E733BFE02F88C" bold="true" box="[312,440,1834,1860]" pageId="82" pageNumber="83">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C37CC722E1E8733AFCA0F88C" box="[462,794,1835,1860]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37CC722E1E8733AFCA0F88C" box="[462,794,1835,1860]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C37CC722E12E7342FE2CF8A4" box="[264,406,1875,1900]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C37CC722E12E7342FE2CF8A4" box="[264,406,1875,1900]" italics="true" pageId="82" pageNumber="83">Seeversiella</emphasis>
</taxonomicName>
by having uniformly brown body; small eyes (temples 4.0­4.3 times as long as eyes); elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at the posterior margin of male tergum 8 (
<figureCitation id="729A2A17C37DC723E47A7555FB79FE96" box="[1116,1219,324,350]" captionStart="FIGURES 331 ­ 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 ­ 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 ­ 332, holotype; 333 ­ 334, paratype from Villa Mills, Costa Rica). 331 male tergum 8; 332 male sternum 8; 333 female tergum 8; 334 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="83" pageNumber="84">Fig. 331</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C37DC723E41A757DFE8FFE66" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84">Figs. 335­340, 342­ 343</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C37DC723E23F7585FD3AFE66" box="[537,640,404,430]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84">Fig. 341</figureCitation>
).
</paragraph>
<caption id="BEDE661AC37DC723E12E726DFB1FF888" httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84" targetBox="[283,1301,504,1615]" targetPageId="83">
<paragraph id="EA1E3692C37DC723E12E726DFB1FF888" blockId="83.[264,1323,1660,1856]" pageId="83" pageNumber="84">
<emphasis id="D8D5EA80C37DC723E12E726DFE65F95C" bold="true" box="[264,479,1660,1684]" pageId="83" pageNumber="84">FIGURES 335­343</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C37DC723E25B726CFBB0F95B" authority="Gusarov" authorityName="Gusarov" box="[637,1034,1660,1684]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
<emphasis id="D8D5EA80C37DC723E25B726CFC12F95C" box="[637,936,1661,1684]" italics="true" pageId="83" pageNumber="84">Seeversiella microphthalma</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37DC723E430726DFBE9F95C" box="[1046,1107,1660,1684]" pageId="83" pageNumber="84" rank="species">
<emphasis id="D8D5EA80C37DC723E430726DFBE9F95C" bold="true" box="[1046,1107,1660,1684]" pageId="83" pageNumber="84">sp. n.</emphasis>
</taxonomicNameLabel>
(335­340, 342­343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342­343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342­343), 0.1 mm (336­337, 339, 341).
</paragraph>
</caption>
<paragraph id="EA1E3692C37AC724E11E750DFEFFFE1D" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
<taxonomicName id="2DA14D11C37AC724E11E750DFD3CFEFD" box="[312,646,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37AC724E11E750DFD3CFEFD" box="[312,646,284,309]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37AC724E311750DFC44FEFD" box="[823,1022,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C37AC724E311750DFC44FEFD" box="[823,1022,284,309]" italics="true" pageId="84" pageNumber="85">S. impressicollis</emphasis>
</taxonomicName>
in having less asperate pronotal punctation; in males pronotum without medial impression, posterior angles of male tergum 3 projecting as spines; lateral diverticula of internal sac without long spiniform sclerites (
<figureCitation id="729A2A17C37AC724E19F7585FD91FE66" box="[441,555,404,430]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 343</figureCitation>
; 316, 318); L­shaped spermatheca with larger distal portion (
<figureCitation id="729A2A17C37AC724E4DD7585FE8FFE1E" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Fig. 341</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37AC725E11E75F5FDC4FE4E" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="description">
<paragraph id="EA1E3692C37AC724E11E75F5FD4AFDBD" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
<taxonomicName id="2DA14D11C37AC724E11E75F5FD3FFE35" box="[312,645,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37AC724E11E75F5FD3FFE35" box="[312,645,484,509]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37AC724E31275F5FC75FE35" box="[820,975,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37AC724E31275F5FC75FE35" box="[820,975,484,509]" italics="true" pageId="84" pageNumber="85">S. sulcicollis</emphasis>
</taxonomicName>
in having larger body, less asperate pronotal punctation; in males pronotum without medial furrow, posterior angles of male tergum 3 projecting as spines; lateral diverticula of internal sac without strong denticulate sclerites (
<figureCitation id="729A2A17C37AC724E226764DFDCBFDBE" box="[512,625,604,630]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 343</figureCitation>
; 326­328).
</paragraph>
<paragraph id="EA1E3692C37AC724E11E7695FC07FCDE" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
<taxonomicName id="2DA14D11C37AC724E11E7695FD3BFD55" box="[312,641,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37AC724E11E7695FD3BFD55" box="[312,641,644,669]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37AC724E3047695FC6EFD55" box="[802,980,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="geostiboides">
<emphasis id="D8D5EA80C37AC724E3047695FC6EFD55" box="[802,980,644,669]" italics="true" pageId="84" pageNumber="85">S. geostiboides</emphasis>
</taxonomicName>
in having pubescence in the lateral portions of pronotum directed towards the midline and obliquely posteriorly; in wider apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C37AC724E37F76C5FBBEFD26" box="[857,1028,724,750]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 335­336</figureCitation>
; 348­349); and different shape of the distal portion of spermatheca (
<figureCitation id="729A2A17C37AC724E32476EDFCC9FCDE" box="[770,883,764,790]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 341</figureCitation>
; 355).
</paragraph>
<paragraph id="EA1E3692C37AC724E11E7735FD62FCAE" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
<taxonomicName id="2DA14D11C37AC724E11E7735FDC5FCF5" box="[312,639,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37AC724E11E7735FDC5FCF5" box="[312,639,804,829]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37AC724E33A7735FC31FCF5" box="[796,907,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C37AC724E33A7735FC31FCF5" box="[796,907,804,829]" italics="true" pageId="84" pageNumber="85">S. flavida</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C37AC724E3E37735FB8BFCF5" box="[965,1073,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="D8D5EA80C37AC724E3E37735FB8BFCF5" box="[965,1073,804,829]" italics="true" pageId="84" pageNumber="85">S. adusta</emphasis>
</taxonomicName>
in having abdominal tergum 8 with four pairs of macrosetae.
</paragraph>
<paragraph id="EA1E3692C37AC724E11E7765FBC2FC7D" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
<taxonomicName id="2DA14D11C37AC724E11E7765FD3EFC45" box="[312,644,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37AC724E11E7765FD3EFC45" box="[312,644,884,909]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37AC724E3097765FC60FC45" box="[815,986,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="paramoana">
<emphasis id="D8D5EA80C37AC724E3097765FC60FC45" box="[815,986,884,909]" italics="true" pageId="84" pageNumber="85">S. paramoana</emphasis>
</taxonomicName>
in having concave but not emarginate apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C37AC724E374778DFC40FC7E" box="[850,1018,924,950]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 338­339</figureCitation>
; 379­380).
</paragraph>
<paragraph id="EA1E3692C37AC724E11E77D2FCCBFC16" blockId="84.[264,1324,284,1870]" box="[312,881,963,990]" pageId="84" pageNumber="85">
<emphasis id="D8D5EA80C37AC724E11E77D2FE6AFC15" bold="true" box="[312,464,963,989]" pageId="84" pageNumber="85">Description.</emphasis>
Length
<quantity id="2D599B77C37AC724E21F77D5FD7EFC16" box="[569,708,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="84" pageNumber="85" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C37AC724E11E77FDFE3DFBB6" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 4.0­4.3 times as long as eyes. Antennal article 2 as long as 3, article 4 subquadrate, 5­10 transverse.</paragraph>
<paragraph id="EA1E3692C37AC724E11E709DFD2EFA76" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C37AC724E40D709DFB6CFB6E" box="[1067,1238,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="84" pageNumber="85" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
, length
<quantity id="2D599B77C37AC724E12E70A5FE0BFB06" box="[264,433,1204,1230]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="84" pageNumber="85" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43­0.47 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with poorly visible microsculpture, with fine and asperate punctation, distance between punctures equals 1­1.5 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C37AC724E11E71DDFE71F966" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 2­4 times their diameter; tergum 7 with microsculpture consisting of transverse meshes and with fine and sparse punctation, distance between punctures equals 3­5 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C37AC724E1377285FE04F966" box="[273,446,1684,1710]" captionStart="FIGURES 331 ­ 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 ­ 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 ­ 332, holotype; 333 ­ 334, paratype from Villa Mills, Costa Rica). 331 male tergum 8; 332 male sternum 8; 333 female tergum 8; 334 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="84" pageNumber="85">Figs. 331, 333</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37AC724E11E72ADFE56F886" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
In the only known male the posterior angles of tergum 3 projecting as sharp spines half as long as tergum (measured medially); tergum
<quantity id="2D599B77C37AC724E31B72F5FCD1F936" box="[829,875,1764,1790]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="84" pageNumber="85" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina half as long as the tergum; posterior margin of tergum 8 without clear medial emargination (
<figureCitation id="729A2A17C37AC724E15E7325FE65F886" box="[376,479,1844,1870]" captionStart="FIGURES 331 ­ 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 ­ 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 ­ 332, holotype; 333 ­ 334, paratype from Villa Mills, Costa Rica). 331 male tergum 8; 332 male sternum 8; 333 female tergum 8; 334 female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="84" pageNumber="85">Fig. 331</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37BC725E11E750DFC19FE96" blockId="85.[264,1324,284,510]" pageId="85" pageNumber="86">
Aedeagus as in
<figureCitation id="729A2A17C37BC725E1D9750DFCA5FEFE" box="[511,799,284,310]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Figs. 335­340, 342­343</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C37BC725E3097555FC2CFE96" box="[815,918,324,350]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Fig. 337</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37BC725E11E757DFDC4FE4E" blockId="85.[264,1324,284,510]" box="[312,638,364,390]" pageId="85" pageNumber="86">
Spermatheca as in
<figureCitation id="729A2A17C37BC725E233757DFDC0FE4E" box="[533,634,364,390]" captionStart="FIGURES 335 ­ 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 median lobe, parameral view; 336 apex of median lobe, parameral view; 337 apex of left paramere; 338 median lobe, lateral view; 339 apex of median lobe, lateral view; 340 details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Fig. 341</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37BC725E11E7582FE32FE36" pageId="85" pageNumber="86" type="distribution">
<paragraph id="EA1E3692C37BC725E11E7582FBCDFE66" blockId="85.[264,1324,284,510]" box="[312,1143,403,430]" pageId="85" pageNumber="86">
<emphasis id="D8D5EA80C37BC725E11E7582FE62FE65" bold="true" box="[312,472,403,429]" pageId="85" pageNumber="86">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C37BC725E3547585FC49FE66" box="[882,1011,404,430]" name="Costa Rica" pageId="85" pageNumber="86">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C37BC725E4257585FBD0FE66" box="[1027,1130,404,430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="85" pageNumber="86">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37BC725E11E75AAFE32FE36" blockId="85.[264,1324,284,510]" pageId="85" pageNumber="86">
<emphasis id="D8D5EA80C37BC725E11E75AAFDBCFE1D" bold="true" box="[312,518,443,469]" pageId="85" pageNumber="86">Natural History.</emphasis>
<taxonomicName id="2DA14D11C37BC725E23075ADFCE7FE1D" box="[534,861,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37BC725E23075ADFCE7FE1D" box="[534,861,444,469]" italics="true" pageId="85" pageNumber="86">Seeversiella microphthalma</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C37BC725E10E75F5FE3EFE36" box="[296,388,484,510]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="85" pageNumber="86" unit="m" value="3000.0">3000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,352 @@
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<mods:title id="BDB92D0CFDB76DB58076A49B12762B13">A new species of Siamoglaris from Thailand with complementary description of the type species (Psocodea: Psocoptera: Prionoglarididae)</mods:title>
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<paragraph id="EA1E369DAD361F23FF400D7B2A1CBE02" blockId="2.[136,1058,1306,1331]" box="[136,1058,1306,1331]" pageId="2" pageNumber="295">
<heading id="B15681F1AD361F23FF400D7B2A1CBE02" box="[136,1058,1306,1331]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="295" reason="2">
<taxonomicName id="2DA14D1EAD361F23FF400D7B2C31BE02" ID-CoL="4X5LY" authority="Lienhard, 2004" authorityName="Lienhard" authorityYear="2004" box="[136,527,1306,1331]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD361F23FF400D7B2F5ABE03" bold="true" box="[136,356,1306,1330]" italics="true" pageId="2" pageNumber="295">Siamoglaris zebrina</emphasis>
<bibRefCitation id="8E304B6CAD361F23FEA40D7A2C31BE02" author="LIENHARD, C." box="[364,527,1307,1331]" pageId="2" pageNumber="295" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
</taxonomicName>
Figs 1-4
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</paragraph>
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<treatmentCitationGroup id="CAB111B3AD361F23FF400D5F2CDABE62" box="[136,740,1342,1364]" pageId="2" pageNumber="295">
<treatmentCitation id="6B00108CAD361F23FF400D5F2C28BE65" author="LIENHARD, C." box="[136,534,1342,1364]" page="866" pageId="2" pageNumber="295" year="2004">
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<emphasis id="D8D5EA8FAD361F23FF400D5F2F75BE62" box="[136,331,1342,1363]" italics="true" pageId="2" pageNumber="295">Siamoglaris zebrina</emphasis>
<bibRefCitation id="8E304B6CAD361F23FE990D5F2C28BE65" author="LIENHARD, C." box="[337,534,1342,1364]" pageId="2" pageNumber="295" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004: 866</bibRefCitation>
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; description of male.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD361F23FF1C0D012D6ABD46" pageId="2" pageNumber="295" type="materials_examined">
<paragraph id="EA1E369DAD361F23FF1C0D012CB1BE96" blockId="2.[136,1059,1375,1656]" pageId="2" pageNumber="295">
<materialsCitation id="5AC93CC0AD361F23FF1C0D012CB1BE96" collectingDate="2003-12-09" collectionCode="TYPE, MATERIAL, MHNG" collectorName="P. Schwendinger." country="Thailand" location="Wang Badan Cave" municipality="Sai Yok District" pageId="2" pageNumber="295" specimenCount="3" stateProvince="Kanchanaburi" typeStatus="holotype">
<collectionCode id="8CB0AE58AD361F23FF1C0D012F59BE45" box="[212,359,1376,1397]" pageId="2" pageNumber="295">TYPE MATERIAL</collectionCode>
:
<collectionCode id="8CB0AE58AD361F23FEBD0D3E2F80BE45" box="[373,446,1375,1396]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="295" type="Museum">MHNG</collectionCode>
,
<specimenCount id="FCA7FD14AD361F23FE030D3E2C04BE44" box="[459,570,1375,1397]" pageId="2" pageNumber="295" type="generic" typeStatus="holotype">3 holotype</specimenCount>
(re-examined),
<collectingCountry id="92B6760DAD361F23FD1E0D3E2D15BE44" box="[726,811,1375,1397]" name="Thailand" pageId="2" pageNumber="295">Thailand</collectingCountry>
,
<collectingRegion id="2865F87FAD361F23FCF10D3E2A22BE44" box="[825,1052,1375,1397]" country="Thailand" name="Kanchanaburi" pageId="2" pageNumber="295">Kanchanaburi Province</collectingRegion>
,
<collectingMunicipality id="0A7AACE7AD361F23FF400D182F1DBEBE" box="[136,291,1401,1423]" pageId="2" pageNumber="295">Sai Yok District</collectingMunicipality>
, near
<location id="EF7E6046AD361F23FEA80D182C2DBEBF" LSID="urn:lsid:plazi:treatment:6208878BAD361F29FF400D7B2C36B8CB:EF7E6046AD361F23FEA80D182C2DBEBF" box="[352,531,1401,1423]" country="Thailand" municipality="Sai Yok District" name="Wang Badan Cave" pageId="2" pageNumber="295" stateProvince="Kanchanaburi">Wang Badan Cave</location>
, ca.
<locationDeviation id="FBAC2FBFAD361F23FD8B0D182DBBBEBE" box="[579,901,1401,1423]" location="Sai Yok Noi Waterfall" pageId="2" pageNumber="295">
<quantity id="2D599B78AD361F23FD8B0D182C4BBEBE" box="[579,629,1401,1423]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="295" unit="km" value="2.0">2 km</quantity>
N of Sai Yok Noi Waterfall
</locationDeviation>
, dry stream bed (on low vegetation),
<date id="9E1F105DAD361F23FE9A0DF32F8DBE96" box="[338,435,1426,1448]" pageId="2" pageNumber="295" value="2003-12-09">
<collectingDate id="8E5BE9B5AD361F23FE9A0DF32F8DBE96" box="[338,435,1426,1448]" pageId="2" pageNumber="295" value="2003-12-09">9.xii.2003</collectingDate>
</date>
, leg.
<collectorName id="4754534BAD361F23FE220DF32CB1BE96" box="[490,655,1426,1448]" pageId="2" pageNumber="295">P. Schwendinger.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="EA1E369DAD361F23FF1C0DD02D6ABD46" blockId="2.[136,1059,1375,1656]" pageId="2" pageNumber="295">
<collectionCode id="8CB0AE58AD361F23FF1C0DD02F3FBEF7" box="[212,257,1457,1478]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="2" pageNumber="295">NEW</collectionCode>
<collectionCode id="8CB0AE58AD361F23FEC70DD42FE9BEF7" box="[271,471,1461,1478]" pageId="2" pageNumber="295">MATERIAL EXAMINED</collectionCode>
:
<collectionCode id="8CB0AE58AD361F23FE240DD02C0BBEF7" box="[492,565,1457,1478]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="295" type="Museum">MHNG</collectionCode>
and
<collectionCode id="8CB0AE58AD361F23FDBD0DD02C8BBEF7" box="[629,693,1457,1478]" pageId="2" pageNumber="295">QSBG</collectionCode>
,
<materialsCitation id="5AC93CC0AD361F23FD010DD02D70BD49" collectingDate="2008-09-11" collectingDateMax="2008-11-13" collectingDateMin="2008-09-11" collectionCode="IO" country="Thailand" county="Khuean Srinagarindra National Park" location="Tourist" municipality="Huai Mae Kamint" pageId="2" pageNumber="295" specimenCode="T3442, T3443, T3444, T3445, T3463, T3464, T3472, T4431, D 1" specimenCount="11" specimenCount-female="10" stateProvince="Kanchanaburi">
<collectingCountry id="92B6760DAD361F23FD010DD02D20BEF6" box="[713,798,1457,1479]" name="Thailand" pageId="2" pageNumber="295">Thailand</collectingCountry>
,
<collectingRegion id="2865F87FAD361F23FCFA0DD02A22BEF6" box="[818,1052,1457,1479]" country="Thailand" name="Kanchanaburi" pageId="2" pageNumber="295">Kanchanaburi Province</collectingRegion>
,
<collectingCounty id="037F4E11AD361F23FF400DAB2FD9BED1" box="[136,487,1482,1504]" pageId="2" pageNumber="295">Khuean Srinagarindra National Park</collectingCounty>
, 93 and
<specimenCount id="FCA7FD14AD361F23FD8B0DAB2C5DBEEE" box="[579,611,1482,1504]" pageId="2" pageNumber="295" type="female">5♀</specimenCount>
<collectingMethod id="33E04E8AAD361F23FDA10DAB2D3DBEEE" box="[617,771,1482,1504]" pageId="2" pageNumber="295">in Malaise traps</collectingMethod>
from the following localities, leg. TIGER-project:
<collectingMunicipality id="0A7AACE7AD361F23FE9D0D822C37BEC8" box="[341,521,1507,1529]" pageId="2" pageNumber="295">Huai Mae Kamint</collectingMunicipality>
/
<location id="EF7E6046AD361F23FDEA0D822C58BEC8" LSID="urn:lsid:plazi:treatment:6208878BAD361F29FF400D7B2C36B8CB:EF7E6046AD361F23FDEA0D822C58BEC8" box="[546,614,1507,1529]" country="Thailand" county="Khuean Srinagarindra National Park" municipality="Huai Mae Kamint" name="Tourist" pageId="2" pageNumber="295" stateProvince="Kanchanaburi">Tourist</location>
center (13,
<date id="9E1F105DAD361F23FD220D822D4EBEC9" box="[746,880,1507,1529]" pageId="2" pageNumber="295" value="2008-09-11" valueMax="2008-09-18" valueMin="2008-09-11">
<collectingDate id="8E5BE9B5AD361F23FD220D822D4EBEC9" box="[746,880,1507,1529]" pageId="2" pageNumber="295" value="2008-09-11" valueMax="2008-09-18" valueMin="2008-09-11">11-18.ix.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FCB60D852D83BEC9" box="[894,957,1507,1529]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3442</specimenCode>
; 23,
<specimenCount id="FCA7FD14AD361F23FC340D822A22BEC8" box="[1020,1052,1507,1529]" pageId="2" pageNumber="295" type="female">2♀</specimenCount>
,
<date id="9E1F105DAD361F23FF400D9C2F31BD23" box="[136,271,1533,1555]" pageId="2" pageNumber="295" value="2008-09-18" valueMax="2008-09-25" valueMin="2008-09-18">
<collectingDate id="8E5BE9B5AD361F23FF400D9C2F31BD23" box="[136,271,1533,1555]" pageId="2" pageNumber="295" value="2008-09-18" valueMax="2008-09-25" valueMin="2008-09-18">18-25.ix.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FEEA0D9C2F5EBD23" box="[290,352,1533,1554]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3443</specimenCode>
;
<specimenCount id="FCA7FD14AD361F23FEBC0D9C2FAABD23" box="[372,404,1533,1554]" pageId="2" pageNumber="295" type="female">1♀</specimenCount>
,
<date id="9E1F105DAD361F23FE600D9C2C0ABD23" box="[424,564,1533,1555]" pageId="2" pageNumber="295" value="2008-09-25" valueMax="2008-10-02" valueMin="2008-09-25">
<collectingDate id="8E5BE9B5AD361F23FE600D9C2C0ABD23" box="[424,564,1533,1555]" pageId="2" pageNumber="295" value="2008-09-25" valueMax="2008-10-02" valueMin="2008-09-25">25.ix-2.x.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FD8F0D9C2CB8BD23" box="[583,646,1533,1554]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3444</specimenCode>
; 13,
<specimenCount id="FCA7FD14AD361F23FD060D9C2CD0BD23" box="[718,750,1533,1554]" pageId="2" pageNumber="295" type="female">1♀</specimenCount>
,
<date id="9E1F105DAD361F23FCC90D9C2D57BD23" box="[769,873,1533,1554]" pageId="2" pageNumber="295" value="2008-10-02" valueMax="2008-10-09" valueMin="2008-10-02">
<collectingDate id="8E5BE9B5AD361F23FCC90D9C2D57BD23" box="[769,873,1533,1554]" pageId="2" pageNumber="295" value="2008-10-02" valueMax="2008-10-09" valueMin="2008-10-02">2-9.x.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FCB40D9C2D85BD22" box="[892,955,1533,1555]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3445</specimenCode>
; 13,
<date id="9E1F105DAD361F23FBCA0D9C2ED6BD1A" pageId="2" pageNumber="295" value="2008-10-23" valueMax="2008-10-30" valueMin="2008-10-23">
<collectingDate id="8E5BE9B5AD361F23FBCA0D9C2ED6BD1A" pageId="2" pageNumber="295" value="2008-10-23" valueMax="2008-10-30" valueMin="2008-10-23">23- 30.x.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FF3C0E772F0DBD1D" box="[244,307,1558,1580]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3463</specimenCode>
; 13,
<date id="9E1F105DAD361F23FEA40E772FC7BD1A" box="[364,505,1558,1580]" pageId="2" pageNumber="295" value="2008-10-30" valueMax="2008-11-06" valueMin="2008-10-30">
<collectingDate id="8E5BE9B5AD361F23FEA40E772FC7BD1A" box="[364,505,1558,1580]" pageId="2" pageNumber="295" value="2008-10-30" valueMax="2008-11-06" valueMin="2008-10-30">30.x-6.xi.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FDCC0E772C7DBD1D" box="[516,579,1558,1580]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3464</specimenCode>
);
<location id="EF7E6046AD361F23FD900E772CDEBD1D" LSID="urn:lsid:plazi:treatment:6208878BAD361F29FF400D7B2C36B8CB:EF7E6046AD361F23FD900E772CDEBD1D" box="[600,736,1558,1580]" country="Thailand" county="Khuean Srinagarindra National Park" municipality="Huai Mae Kamint" name="Tha Thung-na" pageId="2" pageNumber="295" stateProvince="Kanchanaburi">Tha Thung-na</location>
/
<location id="EF7E6046AD361F23FD3B0E772DB4BD1D" LSID="urn:lsid:plazi:treatment:6208878BAD361F29FF400D7B2C36B8CB:EF7E6046AD361F23FD3B0E772DB4BD1D" box="[755,906,1558,1580]" country="Thailand" county="Khuean Srinagarindra National Park" municipality="Huai Mae Kamint" name="Chong Kraborg" pageId="2" pageNumber="295" stateProvince="Kanchanaburi">Chong Kraborg</location>
(13,
<specimenCount id="FCA7FD14AD361F23FC0C0E772DDABD1A" box="[964,996,1558,1579]" pageId="2" pageNumber="295" type="female">1♀</specimenCount>
,
<date id="9E1F105DAD361F23FC380E772EDDBD75" pageId="2" pageNumber="295" value="2008-10-30" valueMax="2008-11-06" valueMin="2008-10-30">
<collectingDate id="8E5BE9B5AD361F23FC380E772EDDBD75" pageId="2" pageNumber="295" value="2008-10-30" valueMax="2008-11-06" valueMin="2008-10-30">30.x- 6.xi.2008</collectingDate>
</date>
,
<specimenCode id="BA079EE6AD361F23FF390E512F0EBD75" box="[241,304,1583,1605]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="295">T3472</specimenCode>
; 23, lacking abdomen,
<date id="9E1F105DAD361F23FDEC0E4E2CA0BD75" box="[548,670,1583,1605]" pageId="2" pageNumber="295" value="2008-11-06" valueMax="2008-11-13" valueMin="2008-11-06">
<collectingDate id="8E5BE9B5AD361F23FDEC0E4E2CA0BD75" box="[548,670,1583,1605]" pageId="2" pageNumber="295" value="2008-11-06" valueMax="2008-11-13" valueMin="2008-11-06">6-13.xi.2008</collectingDate>
</date>
,
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, these
<specimenCount id="FCA7FD14AD361F23FCFC0E512DFBBD74" box="[820,965,1583,1605]" pageId="2" pageNumber="295" type="generic">two specimens</specimenCount>
could be identified as males due to their small body size and their relatively large compound eyes, i.e.
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/
<specimenCode id="BA079EE6AD361F23FF600E032EF2BD46" box="[168,204,1634,1655]" collectionCode="IO" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="2" pageNumber="295">D 1</specimenCode>
.16 and 1.15, see also Complementary description of male, below)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD371F24FD800AA52A34B8BD" lastPageId="5" lastPageNumber="298" pageId="3" pageNumber="296" type="description">
<paragraph id="EA1E369DAD371F22FD800AA52CBFB9E8" blockId="3.[151,1074,708,785]" box="[584,641,708,729]" pageId="3" pageNumber="296">FIG. 1</paragraph>
<paragraph id="EA1E369DAD371F22FF5F0A832D3FB820" blockId="3.[151,1074,708,785]" pageId="3" pageNumber="296">
<taxonomicName id="2DA14D1EAD371F22FF5F0A832FFCB9C9" authority="Lienhard" authorityName="Lienhard" authorityYear="2004" box="[151,450,738,760]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD371F22FF5F0A832F61B9C6" box="[151,351,738,759]" italics="true" pageId="3" pageNumber="296">Siamoglaris zebrina</emphasis>
Lienhard
</taxonomicName>
, male: Habitus, lateral view (body length
<quantity id="2D599B78AD371F22FCB50A832DF2B9C9" box="[893,972,738,760]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="296" unit="mm" value="2.5">2.5 mm</quantity>
); antenna incomplete; pilosity, hindwing and right appendages not shown.
</paragraph>
<paragraph id="EA1E369DAD371F25FF2B0B5F2C4CBD21" blockId="3.[151,1074,829,1654]" lastBlockId="4.[136,1058,1531,1660]" lastPageId="4" lastPageNumber="297" pageId="3" pageNumber="296">
DESCRIPTION OF FEMALE: Colouration and general morphology as described for male by
<bibRefCitation id="8E304B6CAD371F22FF3C0B3C2F9CB844" author="LIENHARD, C." box="[244,418,861,885]" pageId="3" pageNumber="296" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004)</bibRefCitation>
. Body size distinctly larger than in male, but compound eyes relatively smaller (see Complementary description of male, below). Femora lacking dark brown transversal band of hypodermal pigment in 2/3 of their length (Fig. 4d, cf. Fig. 5i showing this band in
<taxonomicName id="2DA14D1EAD371F22FE000BDC2C7AB8E4" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[456,580,957,981]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD371F22FE000BDC2C7AB8E4" box="[456,580,957,981]" italics="true" pageId="3" pageNumber="296">S. theresiae</emphasis>
</taxonomicName>
). Colour pattern of eyes not well-preserved in the specimens examined (observed after two years in alcohol), but still partially visible (cf. Fig. 5a). Pterostigma colourless, transparent or very slightly opaque. Antennae damaged in all specimens examined. Anterodorsal region of abdomen without small humps (see also Fig. 6f, showing pair of humps present in male of
<taxonomicName id="2DA14D1EAD371F22FC690C5C2A1CBF64" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[929,1058,1085,1109]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD371F22FC690C5C2A1CBF64" box="[929,1058,1085,1109]" italics="true" pageId="3" pageNumber="296">S. theresiae</emphasis>
</taxonomicName>
). Terminalia shown in Figs 2, 3; see also revised generic diagnosis, above. Epiproct relatively small, pilosity only developed in apical half, consisting of a pair of long setae and some shorter hairs. Paraproct in middle with a transversal row of some long setae, in its apical half with a dense group of hairs, some of them differentiated as papilliferous setae, similar to those on the external gonapophysis (in some cases weakly differentiated, see Fig. 2b: 4); sense cushion well-differentiated, bearing one normal seta and numerous fine trichobothria, the latter without basal rosettes. Subgenital plate short, with a anteriorly narrowing pigmented area. External gonapophyses welldeveloped, bearing some long stout setae and many shorter acuminate hairs; towards posteroventral margin with numerous apically thin-walled and usually slightly curved papilliferous spatulate or club-shaped setae of unknown function (see Discussion, below). An approximately oval structure present on each side at base of external gonapophysis (shown by interrupted lines in Fig. 2a), originating from anteroventral margin of clunium, covered by rounded basal part of gonapophysis and lateral part of subgenital plate (rudiments of ventral and dorsal gonapophyses?). Spermapore region with a weakly sclerotized suboval posterior area and a membranous anterior area bearing the spermapore. Spermathecal duct very long and more or less spirally coiled FIG. 2
</paragraph>
<paragraph id="EA1E369DAD301F25FF400E7B2DA4BD4A" blockId="4.[136,1058,1531,1660]" pageId="4" pageNumber="297">
<taxonomicName id="2DA14D1EAD301F25FF400E7B2F98BD01" authority="Lienhard" authorityName="Lienhard" authorityYear="2004" box="[136,422,1562,1584]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="297" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD301F25FF400E7B2F74BD1E" box="[136,330,1562,1583]" italics="true" pageId="4" pageNumber="297">Siamoglaris zebrina</emphasis>
Lienhard
</taxonomicName>
, female: (a) Subgenital plate, ovipositor valvulae (pilosity on right side not figured), ventrolateral parts of clunium, spermapore region and distal part of spermathecal duct. (b) Papilliferous setae on external gonapophysis and paraproct; 1 erect seta, 2 curved seta, 3 slender seta, 4 weakly differentiated seta. (c) Epiproct and right paraproct.
</paragraph>
<paragraph id="EA1E369DAD311F24FF5F08822D4ABA6A" blockId="5.[151,1074,227,348]" pageId="5" pageNumber="298">
(at least more than twice as long as the distal part of the spermathecal duct shown in Fig. 2a), spermathecal sac not examined (lost during dissection). Measurements (female of sample T3472): BL =
<quantity id="2D599B78AD311F24FE3309452C73BA0A" box="[507,589,291,316]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="298" unit="mm" value="3.5">3.5 mm</quantity>
; FW =
<quantity id="2D599B78AD311F24FD6909452CCDBA0A" box="[673,755,292,316]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="298" unit="mm" value="4.2">4.2 mm</quantity>
; HW =
<quantity id="2D599B78AD311F24FC8409452DA3BA0A" box="[844,925,292,316]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="298" unit="mm" value="2.8">2.8 mm</quantity>
; F = 917
<emphasis id="D8D5EA8FAD311F24FBC009452A2BBA0A" box="[1032,1045,292,315]" italics="true" pageId="5" pageNumber="298">µ</emphasis>
m; T = 1610
<emphasis id="D8D5EA8FAD311F24FEC909242F30BA6D" box="[257,270,325,348]" italics="true" pageId="5" pageNumber="298">µ</emphasis>
m; t1 = 818
<emphasis id="D8D5EA8FAD311F24FE5C09242F9FBA6D" box="[404,417,325,348]" italics="true" pageId="5" pageNumber="298">µ</emphasis>
m; t2 = 182
<emphasis id="D8D5EA8FAD311F24FDEF09242C0ABA6D" box="[551,564,325,348]" italics="true" pageId="5" pageNumber="298">µ</emphasis>
m; t3 = 185
<emphasis id="D8D5EA8FAD311F24FD7109242CF8BA6D" box="[697,710,325,348]" italics="true" pageId="5" pageNumber="298">µ</emphasis>
m; IO/D = 1.35.
</paragraph>
<paragraph id="EA1E369DAD311F24FF2B090E2A34B8BD" blockId="5.[151,1074,367,909]" pageId="5" pageNumber="298">
COMPLEMENTARY DESCRIPTION OF MALE: Body size smaller than in female (BL3 &lt;
<quantity id="2D599B78AD311F24FF6509F12ED5BA96" box="[173,235,400,424]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="298" unit="mm" value="3.0">3 mm</quantity>
; BL
<specimenCount id="FCA7FD14AD311F24FED409F12F0CBA96" box="[284,306,400,423]" pageId="5" pageNumber="298" type="female"></specimenCount>
&gt;
<quantity id="2D599B78AD311F24FE8709F12FB3BA96" box="[335,397,400,424]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="298" unit="mm" value="3.0">3 mm</quantity>
), but compound eyes relatively larger (IO/D3 &lt;1.2; IO/D
<specimenCount id="FCA7FD14AD311F24FBCD09F12A25BA96" box="[1029,1051,400,423]" pageId="5" pageNumber="298" type="female"></specimenCount>
&gt; 1.3). Colouration as in female. Antennae damaged in all specimens examined. Anterodorsal region of abdomen without small humps (see also Fig. 6f, showing pair of humps present in male of
<taxonomicName id="2DA14D1EAD311F24FE0E09902C7FB939" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[454,577,496,520]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD311F24FE0E09902C7FB939" box="[454,577,496,520]" italics="true" pageId="5" pageNumber="298">S. theresiae</emphasis>
</taxonomicName>
). Terminalia (Fig. 4a-c): Paraproct simple (see
<bibRefCitation id="8E304B6CAD311F24FF5F0A702F7CB918" author="LIENHARD, C." box="[151,322,529,553]" pageId="5" pageNumber="298" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
: fig. 14), its pilosity normal (i.e. lacking the papilliferous setae observed in female). Shape of apical part of phallosome variable due to different positions of the eversible distal structures and variable tumescence of the membranous blisters. For a view of all structures completely everted and blisters swollen see Fig. 4c and the figures of the
<typeStatus id="351A883FAD311F24FE4F0AF32FDAB99B" box="[391,484,658,682]" pageId="5" pageNumber="298" type="holotype">holotype</typeStatus>
in
<bibRefCitation id="8E304B6CAD311F24FDC10AF32C86B99B" author="LIENHARD, C." box="[521,696,658,682]" pageId="5" pageNumber="298" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004)</bibRefCitation>
; for retracted apical structures and collapsed blisters see Fig. 4a; for everted apical structures but collapsed blisters see Fig. 4b. Characteristical distal pair of sclerotized hooks visible in all positions. Rounded medioventral process (sensu
<bibRefCitation id="8E304B6CAD311F24FD840A922CCBB83A" author="LIENHARD, C." box="[588,757,755,779]" pageId="5" pageNumber="298" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
) posteriorly delimited by a transversal reticulate membrane; slightly concave mediodistal margin of this membrane well visible in ventral view (Fig. 4b), but also visible in dorsal view on cleared phallosome (Fig. 4c, interrupted line), always lacking sclerotized posterolateral lobes (see also Figs 5f, g and 6d, e showing posterolateral lobes present in
<taxonomicName id="2DA14D1EAD311F24FCB60B142DC4B8BD" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[894,1018,884,908]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD311F24FCB60B142DC4B8BD" box="[894,1018,884,908]" italics="true" pageId="5" pageNumber="298">S. theresiae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD311F26FF2B0BC12CBFBECB" lastPageId="7" lastPageNumber="300" pageId="5" pageNumber="298" type="discussion">
<paragraph id="EA1E369DAD311F24FF2B0BC12EE2BD2F" blockId="5.[151,1074,928,1663]" pageId="5" pageNumber="298">
DISCUSSION: The female terminalia of
<taxonomicName id="2DA14D1EAD311F24FD430BC12CC7B889" authorityName="Lienhard" authorityYear="2004" box="[651,761,928,952]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD311F24FD430BC12CC7B889" box="[651,761,928,952]" italics="true" pageId="5" pageNumber="298">S. zebrina</emphasis>
</taxonomicName>
resemble those of the other known prionoglaridine genera due to symplesiomorphy. However, the presence of special papilliferous setae on the gonapophyses (some of them also on paraprocts) can be interpreted as an autapomorphy of
<taxonomicName id="2DA14D1EAD311F24FDE70C602C8EBF28" authorityName="Lienhard" authorityYear="2004" box="[559,688,1025,1049]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA8FAD311F24FDE70C602C8EBF28" box="[559,688,1025,1049]" italics="true" pageId="5" pageNumber="298">Siamoglaris</emphasis>
</taxonomicName>
. This interesting character deserves additional comments. The socket and the basal part of these modified setae are identical to those of the adjacent acuminate hairs. The basal part is brown, thick-walled and longitudinally grooved (Fig. 2b) as usual in chaetal sensilla (grooves not visible in the scanning electron micrographs of Fig. 3, but clearly visible under a light microscope; see also
<bibRefCitation id="8E304B6CAD311F24FF3B0CC32FDBBF8A" author="SLIFER, E. H. &amp; SEKHON, S. S." box="[243,485,1186,1211]" pageId="5" pageNumber="298" pagination="315 - 323" refId="ref5899" refString="SLIFER, E. H. &amp; SEKHON, S. S. 1977. Sense organs on the antennal flagellum of psocids (Insecta, Psocoptera). Journal of Morphology 151 (3): 315 - 323." type="journal article" year="1977">Slifer &amp; Sekhon, 1977</bibRefCitation>
and
<bibRefCitation id="8E304B6CAD311F24FDD30CC22C82BF8A" author="HU, F. &amp; ZHANG, G. N. &amp; WANG, J. J." box="[539,700,1186,1211]" pageId="5" pageNumber="298" pagination="628 - 634" refId="ref5537" refString="HU, F., ZHANG, G. N. &amp; WANG, J. J. 2009. Antennal sensillae of five stored-product psocids pests (Psocoptera: Liposcelididae). Micron 40: 628 - 634." type="journal article" year="2009">
Hu
<emphasis id="D8D5EA8FAD311F24FD8B0CC22C4DBF8B" box="[579,627,1186,1210]" italics="true" pageId="5" pageNumber="298">et al</emphasis>
., 2009
</bibRefCitation>
). However, the distal half of these modified setae is thin-walled and completely transparent, and the surface area is augmented by its broadening (spatulate to club-shaped) and by the presence of many small papilliform spicules (Figs 2b; 3d, f). Several more acuminate spicules are also present in the basal half of many of these setae (Figs 2b, 3c). As far as I know, it is the first time that such modified setae have been observed in Psocoptera. In general, the pilosity of the female gonapophyses of psocids consists of normal acuminate setae (probably mechanoreceptors). The particular morphology of these modified setae suggests a possible function as scent organs involved in sexual behaviour (emitting an aphrodisiac pheromone?). Brush-like pheromone-diffusing scent scales of somewhat similar morphology have been observed in several Lepidoptera (
<bibRefCitation id="8E304B6CAD311F24FC850D872DE6BECF" author="GRASSE, P. P." box="[845,984,1510,1534]" pageId="5" pageNumber="298" pagination="48 - 67" refId="ref5477" refString="GRASSE, P. P. 1975. Phaneres epidermiques (pp. 48 - 67). In: GRASSE. P. P. (ed.). Insectes: teguments, systeme nerveux, organes sensoriels. Traite de Zoologie, vol. VIII, fasc. III. Masson, Paris, 910 pp." type="book chapter" year="1975">Grassé, 1975</bibRefCitation>
;
<bibRefCitation id="8E304B6CAD311F24FC2F0D872EF2BD2F" author="WUEST, J." pageId="5" pageNumber="298" pagination="47 - 56" refId="ref5970" refString="WUEST, J. 1996. L'appareil a pheromone d' Argynnis paphia et de Mesoacidalia aglaja males (Lepidopteres, Nymphalides) en microscopie electronique a balayage. Bulletin romand d'entomologie 14: 47 - 56." type="journal article" year="1996">Wüest, 1996</bibRefCitation>
).
</paragraph>
<paragraph id="EA1E369DAD311F27FF2B0E472D09BD4B" blockId="5.[151,1074,928,1663]" lastBlockId="6.[136,1058,1509,1658]" lastPageId="6" lastPageNumber="299" pageId="5" pageNumber="298">
The presence of eversible distal structures on the phallosome, often retracted in alcohol preserved specimens, may pose a problem for the interpretation of the micromorphology of the posterior part of the phallosome. Therefore they have been illus- FIG. 3.
<taxonomicName id="2DA14D1EAD321F27FF050D842FD3BECA" authority="Lienhard" authorityName="Lienhard" authorityYear="2004" box="[205,493,1509,1531]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="299" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD321F27FF050D842FAEBECB" box="[205,400,1509,1530]" italics="true" pageId="6" pageNumber="299">Siamoglaris zebrina</emphasis>
Lienhard
</taxonomicName>
, female, scanning electron micrographs (made by A. Piuz, MHNG): (a) Abdominal apex (ventral side above, paraprocts left). (b) Posteroventral margin of external gonapophyses (same position as in Fig. 3a). (c) Papilliferous setae on posteroventral margin of external gonapophysis. (d) Papilliferous seta on posteroventral margin of external gonapophysis (same seta also recognizable in Fig. 3b, slightly left above middle). (e) Ditto (other seta, not recognizable in Fig. 3b). (f) Detail of apical part of same seta.
</paragraph>
<paragraph id="EA1E369DAD331F26FD800D842CBFBECB" blockId="7.[151,1074,1509,1663]" box="[584,641,1509,1530]" pageId="7" pageNumber="300">FIG. 4</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD331F29FF5F0E622C36B8CB" lastPageId="8" lastPageNumber="301" pageId="7" pageNumber="300" type="description">
<paragraph id="EA1E369DAD331F26FF5F0E622F37BD4F" blockId="7.[151,1074,1509,1663]" pageId="7" pageNumber="300">
<taxonomicName id="2DA14D1EAD331F26FF5F0E622FFABD28" authority="Lienhard" authorityName="Lienhard" authorityYear="2004" box="[151,452,1539,1561]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="7" pageNumber="300" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD331F26FF5F0E622F5EBD29" box="[151,352,1539,1560]" italics="true" pageId="7" pageNumber="300">Siamoglaris zebrina</emphasis>
Lienhard
</taxonomicName>
, male: (a) Posterior part of phallosome (dorsal view) with retracted distal structures. (b) Ditto (same specimen, ventral view) after artificially provoked eversion of distal structures (see Material and methods). (c) Ditto (other specimen, dorsal view) with naturally everted distal structures and swollen membranous blisters. (d) Hindleg (pilosity not shown).
</paragraph>
<paragraph id="EA1E369DAD3C1F29FF4008822F24B82B" blockId="8.[136,1059,227,1019]" pageId="8" pageNumber="301">
trated here in the retracted (Fig. 4a) and in the everted position (Fig. 4b, c). In Fig. 4c the membranous blisters of this region of the phallosome are presented in the swollen position, though not exhibiting the maximal swelling as shown for the
<typeStatus id="351A883FAD3C1F29FC6809422DC3BA0A" box="[928,1021,291,315]" pageId="8" pageNumber="301" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="8E304B6CAD3C1F29FF4009222F12BA6A" author="LIENHARD, C." box="[136,300,323,347]" pageId="8" pageNumber="301" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004</bibRefCitation>
: fig. 11). In this figure of the
<typeStatus id="351A883FAD3C1F29FDAF09222CFABA6A" box="[615,708,323,347]" pageId="8" pageNumber="301" type="holotype">holotype</typeStatus>
the slightly concave mediodistal margin of the reticulate membrane is not well recognizable (probably due to the excessive swelling of lateral blisters), but in the re-examined phallosome of the
<typeStatus id="351A883FAD3C1F29FC2209E22E88BA8A" pageId="8" pageNumber="301" type="holotype">holotype</typeStatus>
it is well visible (blisters collapsed after permanent slide-mounting). The eversion of the distal structures of the phallosome could be directly observed under the stereomicroscope for the specimen shown in Fig. 4a, b (see Material and methods). In the retracted position the distal part of the phallosome is anterodorsally folded and the tips of the pair of sclerotized claspers are anteriorly directed, well visible in dorsal view (Fig. 4a). During eversion this part rises dorsally and finally unfolds backwards, so that the free tips of the hooked claspers are situated at the posterior end of the phallosome (Fig. 4b). Several intermediate states can be observed in alcohol preserved material. When identifying males of
<taxonomicName id="2DA14D1EAD3C1F29FE7C0AC22C0BB98A" authorityName="Lienhard" authorityYear="2004" box="[436,565,675,699]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA8FAD3C1F29FE7C0AC22C0BB98A" box="[436,565,675,699]" italics="true" pageId="8" pageNumber="301">Siamoglaris</emphasis>
</taxonomicName>
one has to pay attention to this phenomenon and not mistake it for variation in the morphology of the terminal structures of the phallosome. For a distinction between
<taxonomicName id="2DA14D1EAD3C1F29FDEB0A822CB3B9CA" authorityName="Lienhard" authorityYear="2004" box="[547,653,739,763]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FDEB0A822CB3B9CA" box="[547,653,739,763]" italics="true" pageId="8" pageNumber="301">S. zebrina</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D1EAD3C1F29FD080A822D02B9CA" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[704,828,739,763]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3C1F29FD080A822D02B9CA" box="[704,828,739,763]" italics="true" pageId="8" pageNumber="301">S. theresiae</emphasis>
</taxonomicName>
see Discussion of the latter species.
</paragraph>
<paragraph id="EA1E369DAD3C1F29FF1C0B422C36B8CB" blockId="8.[136,1059,227,1019]" pageId="8" pageNumber="301">
Almost nothing is known about the biology of this species. The
<typeStatus id="351A883FAD3C1F29FC4C0B422D8CB80A" box="[900,946,803,827]" pageId="8" pageNumber="301">type</typeStatus>
locality is situated about
<quantity id="2D599B78AD3C1F29FEE20B222F9FB86B" box="[298,417,835,859]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="301" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200-300 m</quantity>
from the main entrance of Wang Badan Cave, in a rocky limestone region full of subterranean crevices (
<bibRefCitation id="8E304B6CAD3C1F29FD4A0B022D1BB84A" author="LIENHARD, C." box="[642,805,867,891]" pageId="8" pageNumber="301" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
). The new material has been collected in a very similar limestone region of
<collectingRegion id="2865F87FAD3C1F29FD7A0BE22D93B8AA" box="[690,941,899,923]" country="Thailand" name="Kanchanaburi" pageId="8" pageNumber="301">Kanchanaburi Province</collectingRegion>
. Therefore it seems possible that
<taxonomicName id="2DA14D1EAD3C1F29FEBD0BC22C70B88A" authorityName="Lienhard" authorityYear="2004" box="[373,590,931,955]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FEBD0BC22C70B88A" box="[373,590,931,955]" italics="true" pageId="8" pageNumber="301">Siamoglaris zebrina</emphasis>
</taxonomicName>
has some affinities to caves or similar subterranean habitats, at least during its nymphal life, as known for most
<taxonomicName id="2DA14D1EAD3C1F29FCB90BA22A1DB8EA" authorityName="Azar, Huang &amp; Nel" authorityYear="2017" box="[881,1059,963,987]" class="Insecta" family="Prionoglarididae" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="family">Prionoglarididae</taxonomicName>
(see Lienhard
<emphasis id="D8D5EA8FAD3C1F29FEEA0B852F6FB8CA" box="[290,337,995,1019]" italics="true" pageId="8" pageNumber="301">et al</emphasis>
., 2010a, 2010b).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,311 @@
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<mods:title id="FF8A12A7F6444BD909545CE3A409F877">A new species of Siamoglaris from Thailand with complementary description of the type species (Psocodea: Psocoptera: Prionoglarididae)</mods:title>
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<paragraph id="EA1E369DAD3C1F29FF400C792A1CBF01" blockId="8.[136,1058,1048,1072]" box="[136,1058,1048,1072]" pageId="8" pageNumber="301">
<heading id="B15681F1AD3C1F29FF400C792A1CBF01" box="[136,1058,1048,1072]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="301" reason="2">
<taxonomicName id="2DA14D1EAD3C1F29FF400C792F48BF01" ID-CoL="4X5LX" authority="Lienhard &amp; Ch-, 2011" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[136,374,1048,1072]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="theresiae" status="sp. nov.">
<emphasis id="D8D5EA8FAD3C1F29FF400C792F48BF01" bold="true" box="[136,374,1048,1072]" italics="true" pageId="8" pageNumber="301">Siamoglaris theresiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E657F4AD3C1F29FEB60C782F86BF01" box="[382,440,1049,1072]" pageId="8" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
Figs 5-6
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD3C1F29FF1C0C5D2FC2BFE4" pageId="8" pageNumber="301" type="materials_examined">
<paragraph id="EA1E369DAD3C1F29FF1C0C5D2F7BBFB2" blockId="8.[136,1059,1083,1237]" pageId="8" pageNumber="301">
<materialsCitation id="5AC93CC0AD3C1F29FF1C0C5D2F7BBFB2" collectingDate="2007-09-30" collectingDateMax="2007-10-07" collectingDateMin="2007-09-30" collectionCode="QSBG" collectorName="Songkran &amp; Apichart" country="Thailand" elevation="491" location="Doi Chiang Dao Wildlife Sanctuary" pageId="8" pageNumber="301" specimenCode="T3174" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<typeStatus id="351A883FAD3C1F29FF1C0C5D2F09BF61" box="[212,311,1084,1105]" pageId="8" pageNumber="301" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="8CB0AE58AD3C1F29FE8D0C5A2FBBBF61" box="[325,389,1083,1104]" pageId="8" pageNumber="301">QSBG</collectionCode>
, 3 (on 3 microscopical slides),
<collectingCountry id="92B6760DAD3C1F29FD080C5A2D2BBF60" box="[704,789,1083,1105]" name="Thailand" pageId="8" pageNumber="301">Thailand</collectingCountry>
,
<collectingRegion id="2865F87FAD3C1F29FCEA0C5A2DCFBF60" box="[802,1009,1083,1105]" country="Thailand" name="Chiang Mai" pageId="8" pageNumber="301">Chiang Mai Province</collectingRegion>
,
<location id="EF7E6046AD3C1F29FC360C5A2F89BF5B" LSID="urn:lsid:plazi:treatment:6208878BAD3C1F2CFF400C792D33BA0B:EF7E6046AD3C1F29FC360C5A2F89BF5B" country="Thailand" name="Doi Chiang Dao Wildlife Sanctuary" pageId="8" pageNumber="301" stateProvince="Chiang Mai">Doi Chiang Dao Wildlife Sanctuary</location>
, nature trail,
<quantity id="2D599B78AD3C1F29FDF20C342C4FBF5B" box="[570,625,1109,1130]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="8" pageNumber="301" unit="m" value="491.0">
<elevation id="618CD1AEAD3C1F29FDF20C342C4FBF5B" box="[570,625,1109,1130]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="8" pageNumber="301" unit="m" value="491.0">491m</elevation>
</quantity>
,
<collectingMethod id="33E04E8AAD3C1F29FDB60C342CCBBF5B" box="[638,757,1109,1131]" pageId="8" pageNumber="301">Malaise trap</collectingMethod>
,
<date id="9E1F105DAD3C1F29FCCB0C342DB1BF5B" box="[771,911,1109,1131]" pageId="8" pageNumber="301" value="2007-09-30" valueMax="2007-10-07" valueMin="2007-09-30">
<collectingDate id="8E5BE9B5AD3C1F29FCCB0C342DB1BF5B" box="[771,911,1109,1131]" pageId="8" pageNumber="301" value="2007-09-30" valueMax="2007-10-07" valueMin="2007-09-30">30.ix-7.x.2007</collectingDate>
</date>
, leg.
<collectorName id="4754534BAD3C1F29FC0E0C342A1CBF5A" box="[966,1058,1109,1131]" pageId="8" pageNumber="301">Songkran</collectorName>
&amp;
<collectorName id="4754534BAD3C1F29FF680C0F2ECABFB5" box="[160,244,1134,1156]" pageId="8" pageNumber="301">Apichart</collectorName>
,
<specimenCode id="BA079EE6AD3C1F29FEC80C0F2F01BFB2" box="[256,319,1134,1155]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="301">T3174</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="EA1E369DAD3C1F29FF1C0CEC2FC2BFE4" blockId="8.[136,1059,1083,1237]" pageId="8" pageNumber="301">
<materialsCitation id="5AC93CC0AD3C1F29FF1C0CEC2FABBF93" box="[212,405,1165,1186]" collectionCode="MHNG" pageId="8" pageNumber="301" specimenCount="1" typeStatus="paratype">
<typeStatus id="351A883FAD3C1F29FF1C0CEC2F04BF93" box="[212,314,1165,1186]" pageId="8" pageNumber="301" type="paratype">PARATYPES</typeStatus>
:
<collectionCode id="8CB0AE58AD3C1F29FE840CEC2FABBF93" box="[332,405,1165,1186]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="301" type="Museum">MHNG</collectionCode>
</materialsCitation>
,
<materialsCitation id="5AC93CC0AD3C1F29FE6E0CEC2FC8BFE4" collectingDate="2007-10-08" collectingDateMax="2008-03-24" collectingDateMin="2007-10-08" collectionCode="T" collectorName="C. Piluek &amp; A. Inpuang" country="Thailand" elevation="1306" location="Chong Yen" municipality="Mae Wong National Park" pageId="8" pageNumber="301" specimenCode="T3686, T3641" specimenCount="1" stateProvince="Kamphaeng Phet" typeStatus="paratype">
23,
<collectingCountry id="92B6760DAD3C1F29FE1E0CEC2C15BF92" box="[470,555,1165,1187]" name="Thailand" pageId="8" pageNumber="301">Thailand</collectingCountry>
,
<collectingRegion id="2865F87FAD3C1F29FDF50CEC2D78BF92" box="[573,838,1165,1187]" country="Thailand" name="Kamphaeng Phet" pageId="8" pageNumber="301">Kamphaeng Phet Province</collectingRegion>
,
<collectingMunicipality id="0A7AACE7AD3C1F29FC9F0CEC2E8ABF8D" pageId="8" pageNumber="301">Mae Wong National Park</collectingMunicipality>
,
<location id="EF7E6046AD3C1F29FF090CC72F13BF8A" LSID="urn:lsid:plazi:treatment:6208878BAD3C1F2CFF400C792D33BA0B:EF7E6046AD3C1F29FF090CC72F13BF8A" box="[193,301,1190,1212]" country="Thailand" municipality="Mae Wong National Park" name="Chong Yen" pageId="8" pageNumber="301" stateProvince="Kamphaeng Phet">Chong Yen</location>
,
<quantity id="2D599B78AD3C1F29FEF20CC72F43BF8A" box="[314,381,1190,1212]" metricMagnitude="3" metricUnit="m" metricValue="1.3059999999999998" pageId="8" pageNumber="301" unit="m" value="1306.0">
<elevation id="618CD1AEAD3C1F29FEF20CC72F43BF8A" box="[314,381,1190,1212]" metricMagnitude="3" metricUnit="m" metricValue="1.3059999999999998" pageId="8" pageNumber="301" unit="m" value="1306.0">1306m</elevation>
</quantity>
,
<collectingMethod id="33E04E8AAD3C1F29FE420CC72C3CBF8A" box="[394,514,1190,1212]" pageId="8" pageNumber="301">Malaise trap</collectingMethod>
,
<date id="9E1F105DAD3C1F29FDD80CC72CBABF8A" box="[528,644,1190,1212]" pageId="8" pageNumber="301" value="2007-10-08" valueMax="2007-10-15" valueMin="2007-10-08">
<collectingDate id="8E5BE9B5AD3C1F29FDD80CC72CBABF8A" box="[528,644,1190,1212]" pageId="8" pageNumber="301" value="2007-10-08" valueMax="2007-10-15" valueMin="2007-10-08">8-15.x.2007</collectingDate>
</date>
, leg.
<collectorName id="4754534BAD3C1F29FD730CC72D2BBF8D" box="[699,789,1190,1212]" pageId="8" pageNumber="301">C. Piluek</collectorName>
&amp;
<collectorName id="4754534BAD3C1F29FCFD0CC72D9DBF8A" box="[821,931,1190,1211]" pageId="8" pageNumber="301">A. Inpuang</collectorName>
(
<specimenCode id="BA079EE6AD3C1F29FC7A0CC72DCEBF8D" box="[946,1008,1190,1212]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="301">T3686</specimenCode>
) and
<date id="9E1F105DAD3C1F29FF400CDE2F2EBFE5" box="[136,272,1215,1237]" pageId="8" pageNumber="301" value="2008-03-17" valueMax="2008-03-24" valueMin="2008-03-17">
<collectingDate id="8E5BE9B5AD3C1F29FF400CDE2F2EBFE5" box="[136,272,1215,1237]" pageId="8" pageNumber="301" value="2008-03-17" valueMax="2008-03-24" valueMin="2008-03-17">17-24.iii.2008</collectingDate>
</date>
, leg.
<collectorName id="4754534BAD3C1F29FE8F0CDE2F9EBFE4" box="[327,416,1215,1237]" pageId="8" pageNumber="301">C. Piluek</collectorName>
(
<specimenCode id="BA079EE6AD3C1F29FE670CA12FD0BFE4" box="[431,494,1215,1237]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="301">T3641</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD3C1F2BFF1C0C872C85BEF5" lastPageId="10" lastPageNumber="303" pageId="8" pageNumber="301" type="description">
<paragraph id="EA1E369DAD3C1F28FF1C0C872CBFBEF8" blockId="8.[136,1059,1253,1662]" lastBlockId="9.[151,1074,1459,1664]" lastPageId="9" lastPageNumber="302" pageId="8" pageNumber="301">
DESCRIPTION OF MALE (female unknown): Habitus as in
<taxonomicName id="2DA14D1EAD3C1F29FCFA0C872DA0BFCC" authorityName="Lienhard" authorityYear="2004" box="[818,926,1253,1277]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FCFA0C872DA0BFCC" box="[818,926,1253,1277]" italics="true" pageId="8" pageNumber="301">S. zebrina</emphasis>
</taxonomicName>
(cf. Fig. 1). Colouration in general as describd for
<taxonomicName id="2DA14D1EAD3C1F29FD980D672CFCBE2C" authorityName="Lienhard" authorityYear="2004" box="[592,706,1285,1309]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FD980D672CFCBE2C" box="[592,706,1285,1309]" italics="true" pageId="8" pageNumber="301">S. zebrina</emphasis>
</taxonomicName>
by
<bibRefCitation id="8E304B6CAD3C1F29FD310D642D91BE2F" author="LIENHARD, C." box="[761,943,1285,1310]" pageId="8" pageNumber="301" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004)</bibRefCitation>
, with the following slight differences: head anteroventrally with more dark pigment (Fig. 5a); colour pattern of the legs highly contrasted, femora with a dark brown transversal band of hypodermal pigment in 2/3 of their length (Fig. 5i); some brown pigmentation present along the pterostigmal veins in forewing (Fig. 6a of PT, forewings of HT damaged). Colour pattern of eyes not well-preserved in the specimens examined (after 2-3 years in alcohol) but still partially visible (Fig. 5a). General morphology as described for
<taxonomicName id="2DA14D1EAD3C1F29FEE90D872FAEBECC" authorityName="Lienhard" authorityYear="2004" box="[289,400,1509,1533]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FEE90D872FAEBECC" box="[289,400,1509,1533]" italics="true" pageId="8" pageNumber="301">S. zebrina</emphasis>
</taxonomicName>
by
<bibRefCitation id="8E304B6CAD3C1F29FE760D842C4EBECF" author="LIENHARD, C." box="[446,624,1509,1534]" pageId="8" pageNumber="301" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004)</bibRefCitation>
, with the following differences. Vertex with a pair of small lateral protuberances near compound eyes (Fig. 5a). Antennae damaged in all specimens examined. Both maxillary palps broken in HT. Maxillary palp of PT (sample T3641) as figured for
<taxonomicName id="2DA14D1EAD3C1F29FD8C0E272C90BD6C" authorityName="Lienhard" authorityYear="2004" box="[580,686,1605,1629]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3C1F29FD8C0E272C90BD6C" box="[580,686,1605,1629]" italics="true" pageId="8" pageNumber="301">S. zebrina</emphasis>
</taxonomicName>
by
<bibRefCitation id="8E304B6CAD3C1F29FD1E0E242D44BD6F" author="LIENHARD, C." box="[726,890,1605,1630]" pageId="8" pageNumber="301" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard (2004</bibRefCitation>
: figs 1 and 10), its terminal article with 5 thin-walled conical sensilla in apical half (as figured for FIG. 5
</paragraph>
<paragraph id="EA1E369DAD3D1F28FF5F0DB32F90BD4E" blockId="9.[151,1074,1459,1664]" pageId="9" pageNumber="302">
<taxonomicName id="2DA14D1EAD3D1F28FF5F0DB32F52BED6" authority="Lienhard &amp; Ch-, 2011" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[151,364,1490,1511]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="theresiae" status="sp. nov.">
<emphasis id="D8D5EA8FAD3D1F28FF5F0DB32F52BED6" box="[151,364,1490,1511]" italics="true" pageId="9" pageNumber="302">Siamoglaris theresiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E657F4AD3D1F28FEBD0DB32F92BED6" box="[373,428,1490,1511]" pageId="9" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
, male
<typeStatus id="351A883FAD3D1F28FE3B0DB32C79BED9" box="[499,583,1490,1512]" pageId="9" pageNumber="302" type="holotype">holotype</typeStatus>
: (a) Head (frontal view, pilosity not shown). (b) Remnant of left lacinia (length about 70
<emphasis id="D8D5EA8FAD3D1F28FD830D8D2C69BD30" box="[587,599,1516,1537]" italics="true" pageId="9" pageNumber="302">µ</emphasis>
m). (c) Remnant of right lacinia. (d) Posterior pretarsal claw of midleg (internal view). (e) Anterior pretarsal claw of midleg (external view). (f) Phallosome (ventral view) with naturally everted distal structures and slightly swollen blisters. (g) Posterior part of phallosome (ventral view) showing details of everted distal structures. (h) Labial palpus (pilosity not shown, except for thin-walled internal sensilla). (i) hindleg (pilosity not shown).
</paragraph>
<paragraph id="EA1E369DAD3E1F2BFDF10D4A2C4CBE70" blockId="10.[136,1059,1323,1477]" box="[569,626,1323,1345]" pageId="10" pageNumber="303">FIG. 6</paragraph>
<paragraph id="EA1E369DAD3E1F2BFF400D2B2C85BEF5" blockId="10.[136,1059,1323,1477]" pageId="10" pageNumber="303">
<taxonomicName id="2DA14D1EAD3E1F2BFF400D2B2F62BE6E" authority="Lienhard &amp; Ch-, 2011" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[136,348,1354,1375]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="theresiae" status="sp. nov.">
<emphasis id="D8D5EA8FAD3E1F2BFF400D2B2F62BE6E" box="[136,348,1354,1375]" italics="true" pageId="10" pageNumber="303">Siamoglaris theresiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E657F4AD3E1F2BFEAD0D2B2FA5BE6E" box="[357,411,1354,1375]" pageId="10" pageNumber="303" rank="species">sp. n.</taxonomicNameLabel>
, male
<typeStatus id="351A883FAD3E1F2BFE290D2B2C0ABE6E" box="[481,564,1354,1375]" pageId="10" pageNumber="303" type="paratype">paratype</typeStatus>
(sample T3686): (a) Forewing. (b) Hindwing. (c) Phallosome (lateral view) with retracted distal structures. (d) Ditto (dorsal view). (e) Posterior part of phallosome (ventral view) with artificially provoked partial eversion of distal structures (see Material and methods). (f) Abdominal base (dorsal view) showing pair of small hemispherical humps anteromedially of spiracles of segment 3.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD3E1F2AFF400D842D34B94F" lastPageId="11" lastPageNumber="304" pageId="10" pageNumber="303" type="reference_group">
<paragraph id="EA1E369DAD3E1F2AFF400D842D34B94F" blockId="10.[136,1059,1509,1661]" lastBlockId="11.[151,1074,227,639]" lastPageId="11" lastPageNumber="304" pageId="10" pageNumber="303">
<taxonomicName id="2DA14D1EAD3E1F2BFF400D842ECCBECC" ID-CoL="4X5LY" authorityName="Lienhard" authorityYear="2004" box="[136,242,1509,1533]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3E1F2BFF400D842ECCBECC" box="[136,242,1509,1533]" italics="true" pageId="10" pageNumber="303">S. zebrina</emphasis>
</taxonomicName>
by
<bibRefCitation id="8E304B6CAD3E1F2BFED00D842F84BECC" author="LIENHARD, C." box="[280,442,1509,1533]" pageId="10" pageNumber="303" pagination="441 - 469" refId="ref5679" refString="LIENHARD, C. 2007. Description of a new African genus and a new tribe of Speleketorinae (Psocodea: ' Psocoptera': Prionoglarididae). Revue suisse de Zoologie 114 (3): 441 - 469." type="journal article" year="2007">Lienhard, 2007</bibRefCitation>
: fig. 3b). Remnant of lacinia relatively short (length about 70
<emphasis id="D8D5EA8FAD3E1F2BFF7D0E642EFDBD2D" box="[181,195,1541,1564]" italics="true" pageId="10" pageNumber="303">µ</emphasis>
m), apically weakly sclerotized and slightly bidenticulate (Fig. 5b, c). Anterodorsal region of abdomen with a pair of small hemispherical humps (Fig. 6f). Terminalia (Figs 5f-g, 6c-e): Epiproct, paraproct and hypandrium simple, similar to those of
<taxonomicName id="2DA14D1EAD3E1F2BFF2F0E042F6DBD4C" ID-CoL="4X5LY" authorityName="Lienhard" authorityYear="2004" box="[231,339,1637,1661]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3E1F2BFF2F0E042F6DBD4C" box="[231,339,1637,1661]" italics="true" pageId="10" pageNumber="303">S. zebrina</emphasis>
</taxonomicName>
(
<bibRefCitation id="8E304B6CAD3E1F2BFEAC0E042C36BD4C" author="LIENHARD, C." box="[356,520,1637,1661]" pageId="10" pageNumber="303" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
: figs 14, 15). Apex of dorsolateral appendages of phallosome rather abruptly narrowed (Figs 5f, 6d). Shape of apical part of phallosome variable due to movable distal structures and inflatable membranous blisters. For a view of all structures completely everted see Fig. 5f, g; for apical structures retracted see Fig. 6c, d; for partially everted apical structures see Fig. 6e. Eversible structures lacking conspicuous hooked claspers but bearing a small finely spiculate conical lobe near opening of ejaculatory duct; this median lobe directed anteriad after eversion of the apical structures of the phallosome (Figs 5f, g and 6e); distal part of ejaculatory duct curved in retracted state (Fig. 6c, interrupted lines). Medioventral process (sensu
<bibRefCitation id="8E304B6CAD3F1F2AFF5F09842F05BACF" author="LIENHARD, C." box="[151,315,485,510]" pageId="11" pageNumber="304" pagination="865 - 875" refId="ref5633" refString="LIENHARD, C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
) posteriorly delimited by a transversal reticulate membrane bearing a pair of slightly sclerotized posterolateral lobes (Figs 5f, g and 6e); these conspicuous lobes also visible in retracted state (Fig. 6d). Measurements (HT except for wing lengths): BL =
<quantity id="2D599B78AD3F1F2AFEFF0A262FB9B96F" box="[311,391,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="304" unit="mm" value="2.3">2.3 mm</quantity>
; FW = 4.0 mm (PT); HW =
<quantity id="2D599B78AD3F1F2AFD7C0A262D3BB96F" box="[692,773,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="304" unit="mm" value="2.7">2.7 mm</quantity>
(PT); F = 846
<emphasis id="D8D5EA8FAD3F1F2AFC6A0A262D91B96F" box="[930,943,583,606]" italics="true" pageId="11" pageNumber="304">µ</emphasis>
m; T = 1450
<emphasis id="D8D5EA8FAD3F1F2AFF5F0A092E9AB94E" box="[151,164,616,639]" italics="true" pageId="11" pageNumber="304">µ</emphasis>
m; t1 = 804
<emphasis id="D8D5EA8FAD3F1F2AFEE20A092F09B94E" box="[298,311,616,639]" italics="true" pageId="11" pageNumber="304">µ</emphasis>
m; t2 = 210
<emphasis id="D8D5EA8FAD3F1F2AFE740A092FF7B94E" box="[444,457,616,639]" italics="true" pageId="11" pageNumber="304">µ</emphasis>
m; t3 = 193
<emphasis id="D8D5EA8FAD3F1F2AFD870A092C62B94E" box="[591,604,616,639]" italics="true" pageId="11" pageNumber="304">µ</emphasis>
m; IO/D = 1.06.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD3F1F2AFF2B0AF22CDCB87A" pageId="11" pageNumber="304" type="etymology">
<paragraph id="EA1E369DAD3F1F2AFF2B0AF22CDCB87A" blockId="11.[151,1074,658,844]" pageId="11" pageNumber="304">
ETYMOLOGY: The species is dedicated to my friend and colleague Thérèse Cuche (MHNG) in recognition of her invaluable assistance in my research on Psocoptera during more than 20 years. After retiring in 2007 she continues to work voluntarily for the
<collectingRegion id="2865F87FAD3F1F2AFEAE0A922F89B83A" box="[358,439,755,779]" country="Switzerland" name="Geneve" pageId="11" pageNumber="304">Geneva</collectingRegion>
Museum. Due to her competent sorting to morphospecies and labeling of the thousands of psocids collected by the TIGER-project this fascinating material became accessible to scientific studies.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD3F1F2DFF2B0B012C4CBED1" lastPageId="12" lastPageNumber="305" pageId="11" pageNumber="304" type="discussion">
<paragraph id="EA1E369DAD3F1F2AFF2B0B012CD2BECC" blockId="11.[151,1074,863,1663]" pageId="11" pageNumber="304">
DISCUSSION: The new species is easy to distinguish from
<taxonomicName id="2DA14D1EAD3F1F2AFCA70B3E2DE1B846" authorityName="Lienhard" authorityYear="2004" box="[879,991,863,887]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFCA70B3E2DE1B846" box="[879,991,863,887]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
by the presence of a pair of small protuberances of the vertex, near the compound eyes (Fig. 5a), and of a pair of small hemispherical humps anterodorsally on the abdomen (Fig. 6f). These characters are probably present in both sexes, but as the female is unknown, the possibility of sexual dimorphism cannot be ruled out. Two colour characters should also be mentioned here, although probably of limited diagnostic value, i.e. the presence, in
<taxonomicName id="2DA14D1EAD3F1F2AFE7C0C402C0EBF08" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[436,560,1057,1081]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFE7C0C402C0EBF08" box="[436,560,1057,1081]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
, of a brown transversal band in about 2/3 of the femora (Fig. 5i) and of some brown pigmentation along the pterostigmal veins in the forewing (Fig. 6a). The phallosomes of the two species of
<taxonomicName id="2DA14D1EAD3F1F2AFCD70C032D9EBF4B" authorityName="Lienhard" authorityYear="2004" box="[799,928,1122,1146]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA8FAD3F1F2AFCD70C032D9EBF4B" box="[799,928,1122,1146]" italics="true" pageId="11" pageNumber="304">Siamoglaris</emphasis>
</taxonomicName>
differ by the shape of the dorsolateral appendages, distally more abruptly narrowed in
<taxonomicName id="2DA14D1EAD3F1F2AFC7D0CE32A0CBFAB" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[949,1074,1154,1178]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFC7D0CE32A0CBFAB" box="[949,1074,1154,1178]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
(Figs 5f, 6d) than in
<taxonomicName id="2DA14D1EAD3F1F2AFEA70CC22FE7BF8B" authorityName="Lienhard" authorityYear="2004" box="[367,473,1186,1210]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFEA70CC22FE7BF8B" box="[367,473,1186,1210]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
(Fig. 4), by the presence of a pair of sclerotized terminal claspers in
<taxonomicName id="2DA14D1EAD3F1F2AFEDE0CA22FBBBFEA" authorityName="Lienhard" authorityYear="2004" box="[278,389,1219,1243]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFEDE0CA22FBBBFEA" box="[278,389,1219,1243]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
(Fig. 4), absent in
<taxonomicName id="2DA14D1EAD3F1F2AFDA90CA22CDEBFEA" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[609,736,1219,1243]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFDA90CA22CDEBFEA" box="[609,736,1219,1243]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
(Figs 5f, g and 6d, e), by the presence of a pair of slightly sclerotized lateral lobes at the posterior margin of the medioventral part in
<taxonomicName id="2DA14D1EAD3F1F2AFEB40D652FC7BE2A" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[380,505,1283,1307]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFEB40D652FC7BE2A" box="[380,505,1283,1307]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
(Figs 5f, g and 6d, e), absent in
<taxonomicName id="2DA14D1EAD3F1F2AFCAC0D652DEEBE2A" authorityName="Lienhard" authorityYear="2004" box="[868,976,1283,1307]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFCAC0D652DEEBE2A" box="[868,976,1283,1307]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
(Fig. 4), and by some details of the eversible posterior structures, especially the presence, medially near the opening of the ejaculatory duct, of a spiculate conical lobe in
<taxonomicName id="2DA14D1EAD3F1F2AFF5F0D042F2ABE4D" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[151,276,1380,1404]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFF5F0D042F2ABE4D" box="[151,276,1380,1404]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
(Figs
<quantity id="2D599B78AD3F1F2AFE930D052F4BBE4C" box="[347,373,1380,1405]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="11" pageNumber="304" unit="g" value="5.0">5g</quantity>
, 6e) which could not be observed in
<taxonomicName id="2DA14D1EAD3F1F2AFCC40D042D46BE4D" authorityName="Lienhard" authorityYear="2004" box="[780,888,1380,1404]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFCC40D042D46BE4D" box="[780,888,1380,1404]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
(Fig. 4b, c). The mechanism of everting the distal parts of the phallosome (observed in
<specimenCount id="FCA7FD14AD3F1F2AFC550DE42A17BEAC" box="[925,1065,1413,1437]" pageId="11" pageNumber="304" type="generic" typeStatus="paratype">one paratype</specimenCount>
) corresponds to that described for
<taxonomicName id="2DA14D1EAD3F1F2AFE350DC42C56BE8C" authorityName="Lienhard" authorityYear="2004" box="[509,616,1445,1469]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD3F1F2AFE350DC42C56BE8C" box="[509,616,1445,1469]" italics="true" pageId="11" pageNumber="304">S. zebrina</emphasis>
</taxonomicName>
. For
<taxonomicName id="2DA14D1EAD3F1F2AFD690DC42D23BE8C" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[673,797,1445,1469]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFD690DC42D23BE8C" box="[673,797,1445,1469]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
it has to be noted that the characteristic spiculate conical lobe near the opening of the ejaculatory duct is difficult or impossible to observe in the retracted state (Fig. 6d).
</paragraph>
<paragraph id="EA1E369DAD3F1F2AFF2B0E672A0FBD4E" blockId="11.[151,1074,863,1663]" pageId="11" pageNumber="304">
Almost nothing is known about the biology of
<taxonomicName id="2DA14D1EAD3F1F2AFD120E672D68BD2F" authorityName="Lienhard &amp; Ch-" authorityYear="2011" box="[730,854,1542,1566]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="theresiae">
<emphasis id="D8D5EA8FAD3F1F2AFD120E672D68BD2F" box="[730,854,1542,1566]" italics="true" pageId="11" pageNumber="304">S. theresiae</emphasis>
</taxonomicName>
. The type locality is situated at the foot of a large limestone mountain with caves (e. g. Chiang Dao Cave) and subterranean crevices (Peter Schwendinger, pers. comm.). The
<typeStatus id="351A883FAD3F1F2AFC440E262DCCBD6E" box="[908,1010,1607,1631]" pageId="11" pageNumber="304" type="paratype">paratypes</typeStatus>
were collected in a area with isolated limestone outcrops. Therefore it seems possible that
</paragraph>
<paragraph id="EA1E369DAD381F2DFDF10DAA2C4CBED1" blockId="12.[136,1059,1483,1662]" box="[569,626,1483,1504]" pageId="12" pageNumber="305">FIG. 7</paragraph>
</subSubSection>
<subSubSection id="A2BB6516AD381F2CFF400D882D33BA0B" lastPageId="13" lastPageNumber="306" pageId="12" pageNumber="305" type="description">
<paragraph id="EA1E369DAD381F2DFF400D882ECFBD4C" blockId="12.[136,1059,1483,1662]" pageId="12" pageNumber="305">
<emphasis id="D8D5EA8FAD381F2DFF400D882F76BECE" box="[136,328,1513,1535]" italics="true" pageId="12" pageNumber="305">
<taxonomicName id="2DA14D1EAD381F2DFF400D882F3DBECF" authorityName="Enderlein" authorityYear="1909" box="[136,259,1513,1534]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="305" phylum="Arthropoda" rank="genus">Prionoglaris</taxonomicName>
stygia
</emphasis>
Enderlein: (a) Posterior pretarsal claw of hindleg (internal view). (b) Anterior pretarsal claw of hindleg (external view). (c) Left paraproct of female (specimen from the
<typeStatus id="351A883FAD381F2DFF650E7D2EE8BD00" box="[173,214,1564,1585]" pageId="12" pageNumber="305">type</typeStatus>
locality). (d) Epiproct of female (same specimen; posterior margin pointing to bottom of figure plate). (e) Subgenital plate, ovipositor valvulae (pilosity on right side not shown), ventrolateral parts of clunium, spermapore region and distal part of spermathecal duct (same specimen).
</paragraph>
<paragraph id="EA1E369DAD391F2CFF5F08822D33BA0B" blockId="13.[151,1074,227,315]" pageId="13" pageNumber="306">
also this species has some affinities to caves or similar subterranean habitats, at least during its nymphal life, as has been observed in most
<taxonomicName id="2DA14D1EAD391F2CFD1509622DB1BA2A" authorityName="Azar, Huang &amp; Nel" authorityYear="2017" box="[733,911,259,283]" class="Insecta" family="Prionoglarididae" kingdom="Animalia" order="Psocodea" pageId="13" pageNumber="306" phylum="Arthropoda" rank="family">Prionoglarididae</taxonomicName>
and postulated for
<taxonomicName id="2DA14D1EAD391F2CFF7609422F17BA0A" authorityName="Lienhard" authorityYear="2004" box="[190,297,291,315]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="13" pageNumber="306" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="D8D5EA8FAD391F2CFF7609422F17BA0A" box="[190,297,291,315]" italics="true" pageId="13" pageNumber="306">S. zebrina</emphasis>
</taxonomicName>
(see Discussion of the latter species, above).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,351 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1123.90308" ID-Pensoft-Pub="1313-2970-1123-63" ID-Pensoft-UUID="1BCB0D62026158E7B962D3F2EE741FB4" ID-ZooBank="13556B2ABE8744F69A43FCDAFE1C68CB" ModsDocID="1313-2970-1123-63" checkinTime="1664944213626" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana &amp; Gattolliat, Jean-Luc" docDate="2022" docId="62089C43D7B3558384831625E704FD8A" docLanguage="en" docName="ZooKeys 1123: 63-81" docOrigin="ZooKeys 1123" docPubDate="2022-10-04" docSource="http://dx.doi.org/10.3897/zookeys.1123.90308" docTitle="Labiobaetis kui Kaltenbach &amp; Garces &amp; Gattolliat 2022, sp. nov." docType="treatment" docUuid="03B09E8B-57E2-40AA-8BAD-911A6D969606" docUuidSource="ZooBank" docVersion="1" id="1BCB0D62026158E7B962D3F2EE741FB4" lastPageNumber="63" masterDocId="1BCB0D62026158E7B962D3F2EE741FB4" masterDocTitle="First contribution to Labiobaetis Novikova &amp; Kluge in Cambodia (Ephemeroptera, Baetidae), with description of two new species" masterLastPageNumber="81" masterPageNumber="63" pageNumber="63" updateTime="1664944213626" updateUser="pensoft">
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<mods:title>First contribution to Labiobaetis Novikova &amp; Kluge in Cambodia (Ephemeroptera, Baetidae), with description of two new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kaltenbach, Thomas</mods:namePart>
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<mods:namePart>Garces, Jhoana</mods:namePart>
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<mods:affiliation>Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-10-04</mods:number>
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<treatment LSID="urn:lsid:zoobank.org:act:03B09E8B-57E2-40AA-8BAD-911A6D969606" httpUri="http://treatment.plazi.org/id/62089C43D7B3558384831625E704FD8A" lastPageNumber="63" pageId="0" pageNumber="63">
<subSubSection pageId="0" pageNumber="63" type="nomenclature">
<paragraph pageId="0" pageNumber="63">
<taxonomicName LSID="https://zoobank.org/03B09E8B-57E2-40AA-8BAD-911A6D969606" authority="Kaltenbach &amp; Garces &amp; Gattolliat, 2022" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">Labiobaetis kui</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">, 5c, d</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Labiobaetis in Cambodia a overview map b Labiobaetis species." figureDoi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63">, 6b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="diagnosis">
<paragraph pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Larva.</emphasis>
Following combination of characters: A) antennal scape without process (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4g</figureCitation>
); B) dorsal surface of labrum with submarginal arc of 16-18 long, clavate setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3a</figureCitation>
); C) labial palp segment II with an extended, slightly hooked, thumb-like distomedial protuberance (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3i</figureCitation>
); D) left mandible without setae at apex of mola (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3e</figureCitation>
); E) fore femur rather slender, length ca 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with 10-15 curved, spine-like setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4a</figureCitation>
); F) hind protoptera absent; G) six pairs of terga-lii; H) paraproct distally slightly expanded, with 33-38 stout, marginal spines (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4f</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
labrum (left: ventral view, right: dorsal view)
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
seta of submarginal arc
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
right mandible
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
right prostheca
<emphasis bold="true" pageId="0" pageNumber="63">e</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="63">f</emphasis>
left prostheca
<emphasis bold="true" pageId="0" pageNumber="63">g</emphasis>
hypopharynx and superlinguae
<emphasis bold="true" pageId="0" pageNumber="63">h</emphasis>
maxilla
<emphasis bold="true" pageId="0" pageNumber="63">i</emphasis>
labium (left: ventral view, right: dorsal view).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
foreleg
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
seta of tibia dorsal margin
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
fore claw
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
tergum IV
<emphasis bold="true" pageId="0" pageNumber="63">e</emphasis>
tergalius IV
<emphasis bold="true" pageId="0" pageNumber="63">f</emphasis>
paraproct
<emphasis bold="true" pageId="0" pageNumber="63">g</emphasis>
antennal base.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">Description.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Larva</emphasis>
(Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">5c, d</figureCitation>
). Body length ca 4.9 mm. Caudalii broken. Antenna broken.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Colouration</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, larvae a Labiobaetis brao sp. nov., dorsal view b Labiobaetis brao sp. nov., ventral view c Labiobaetis kui sp. nov., dorsal view d Labiobaetis kui sp. nov., ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63">5c, d</figureCitation>
). Head, thorax, and abdomen dorsally uniform brown. Head, thorax, and abdomen ventrally light brown. Legs light brown; femur with a brown medial spot, darker on ventral margin, dorsal margin and apex brown. Caudalii light brown.
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure5" httpUri="https://binary.pensoft.net/fig/752847" pageId="0" pageNumber="63" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 5.</emphasis>
Habitus, larvae
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis brao" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="brao">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis brao</emphasis>
</taxonomicName>
sp. nov., dorsal view
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis brao" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="brao">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis brao</emphasis>
</taxonomicName>
sp. nov., ventral view
<emphasis bold="true" pageId="0" pageNumber="63">c</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., dorsal view
<emphasis bold="true" pageId="0" pageNumber="63">d</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kui" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="kui">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis kui</emphasis>
</taxonomicName>
sp. nov., ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4g</figureCitation>
) with scape and pedicel subcylindrical, without distolateral process at scape.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Labrum</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3a, b</figureCitation>
). Rectangular, length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 16-18 long, clavate setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae; ventral surface with ca three short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Right mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3c, d</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles, inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Left mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3e, f</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Tuft of setae at apex of mola absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3g</figureCitation>
). Lingua approx. as long as superlinguae. Lingua longer than broad; medial tuft of stout setae well developed; distal half laterally expanded. Superlinguae distally almost straight; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3h</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid. Medially with one pectinate, spine-like seta and three or four medium, simple setae. Maxillary palp 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; 2-segmented; palp segment II 1.2
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Labium</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis kui sp. nov., larva morphology a labrum (left: ventral view, right: dorsal view) b seta of submarginal arc c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium (left: ventral view, right: dorsal view)." figureDoi="10.3897/zookeys.1123.90308.figure3" httpUri="https://binary.pensoft.net/fig/752845" pageId="0" pageNumber="63">3i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca six spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with ca four spine-like setae; ventral surface with fine, simple, scattered setae. Paraglossa subrectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and a row of 2-4 medium, simple setae in anteromedial area; dorsally with a row of four or five long, spine-like, simple setae near inner margin. Labial palp with segment I 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with extended, slightly hooked, thumb-like distomedial protuberance; distomedial protuberance 0.7
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with two long, spine-like, simple setae near outer margin. Segment III slightly pentagonal; apex rounded, inner apical margin slightly concave; length subequal to width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Foreleg</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4a-c</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.4:0.2.
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Femur</emphasis>
.
</emphasis>
Length ca 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with 10-15 long, curved, spine-like setae; length of setae 0.23
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded, with a pair of long, curved, spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch absent.
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Tibia</emphasis>
.
</emphasis>
Dorsal margin with row of short, stout, apically rounded setae, on apex one longer, spine-like seta. Ventral margin with row of short, curved, spine-like setae, on apex some longer setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae near ventral margin. Patellatibial suture present on basal 1/3 area.
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Tarsus</emphasis>
.
</emphasis>
Dorsal margin with some fine, simple setae. Ventral margin with row of curved, spine-like setae. Claw with one row of nine or ten denticles; distally pointed; with ca. five stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Terga</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4d</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Tergalii</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4e</figureCitation>
). Present on segments II-VII. Margins with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Tergalius IV as long as length of segments V and 1/3 VI combined. Tergalius VII as long as length of segment VIII.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Paraproct</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis kui sp. nov., larva morphology a foreleg b seta of tibia dorsal margin c fore claw d tergum IV e tergalius IV f paraproct g antennal base." figureDoi="10.3897/zookeys.1123.90308.figure4" httpUri="https://binary.pensoft.net/fig/752846" pageId="0" pageNumber="63">4f</figureCitation>
). Distally slightly expanded, with 33-38 stout, marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with numerous small, marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="etymology">
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="63">The new species is dedicated to the indigenous Kui people from northeastern Cambodia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="distribution">
<paragraph pageId="0" pageNumber="63">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="63">
Cambodia (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Labiobaetis in Cambodia a overview map b Labiobaetis species." figureDoi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63">6b</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1123.90308.figure6" httpUri="https://binary.pensoft.net/fig/752848" pageId="0" pageNumber="63" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 6.</emphasis>
Distribution of
<taxonomicName authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis</emphasis>
</taxonomicName>
in Cambodia
<emphasis bold="true" pageId="0" pageNumber="63">a</emphasis>
overview map
<emphasis bold="true" pageId="0" pageNumber="63">b</emphasis>
<taxonomicName authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Labiobaetis</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="biological aspects">
<paragraph pageId="0" pageNumber="63">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="63">The specimens were collected from 100 m to 640 m, mostly on littoral gravel.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="type material">
<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
.
</emphasis>
Cambodia • larva; Kampong Speu Province, Chambok River, 1.83 Km from Chambok Community; 240 m;
<geoCoordinate degrees="11" direction="north" minutes="21" orientation="latitude" precision="15" seconds="58" value="11.366112">11°21'58&quot;N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="06" orientation="longitude" precision="15" seconds="17" value="104.10472">104°06'17&quot;E</geoCoordinate>
; 11.07.2018; leg. H. Freitag and J. Garces; on slide; GBIFCH00592702; MZL.
<emphasis bold="true" pageId="0" pageNumber="63">
<emphasis italics="true" pageId="0" pageNumber="63">Paratypes</emphasis>
.
</emphasis>
Cambodia • 14 larvae; same data as holotype; 1 on slide; GBIFCH00592701; MZL; 13 in alcohol; GenBank ON982737, ON982738; GBIFCH00515681, GBIFCH00829876, GBIFCH00829877, GBIFCH00975577, GBIFCH00975578; MZL • 7 larvae; Kampong Speu Province, waterfall at Kirirom National Park; 640 m;
<geoCoordinate degrees="11" direction="north" minutes="20" orientation="latitude" precision="15" seconds="26" value="11.340555">11°20'26&quot;N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="02" orientation="longitude" precision="15" seconds="14" value="104.037224">104°02'14&quot;E</geoCoordinate>
; 13.07.2018; leg. H. Freitag and J. Garces; 1 on slide; GBIFCH00592698; RUPP; 6 in alcohol; GBIFCH00975579; MZL • 1 larva; Kampong Speu Province, Kokie waterfall, secondary forest remnants; 110 m;
<geoCoordinate degrees="11" direction="north" minutes="12" orientation="latitude" precision="15" seconds="11" value="11.203055">11°12'11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="03" orientation="longitude" precision="15" seconds="49" value="104.063614">104°03'49&quot;E</geoCoordinate>
; 12.07.2018; leg. H. Freitag and J. Garces; on slide; GBIFCH00592699; RUPP.
</paragraph>
</subSubSection>
</treatment>
</document>