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<mods:title id="B05CCF882630D590E2DEB0942E5E3967">A spectacular new antlion species of the subfamily Dendroleontinae (Neuroptera: Myrmeleontidae) from Malaysia</mods:title>
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<emphasis id="0DDFEAB0D800F905FF54FF14BC00FD6E" bold="true" box="[136,535,236,260]" pageId="5" pageNumber="294">
<emphasis id="0DDFEAB0D800F905FF54FF14BF95FD6E" bold="true" box="[136,386,236,260]" italics="true" pageId="5" pageNumber="294">Layahima wuzhishana</emphasis>
(
<bibRefCitation id="5B3A4B53D800F905FE4CFF14BC07FD6E" author="Yang" box="[400,528,236,260]" pageId="5" pageNumber="294" pagination="296 - 299" refId="ref6663" refString="Yang, C. K., Wang, X. L. 2002. Neuroptera: Myrmeleontidae. In: Huang, F. S. (ed.), Forest Insects of Hainan. Science Press, Beijing. pp. 296 - 299." type="book chapter" year="2002">Yang, 2002</bibRefCitation>
)
</emphasis>
</taxonomicName>
(
<figureCitation id="A7902A27D800F905FDE5FF14BC86FD6F" box="[569,657,236,261]" captionStart="Figure 2" captionStartId="4.[155,218,1987,2009]" captionTargetBox="[189,1277,358,1942]" captionTargetId="figure-71@4.[189,1398,348,1942]" captionTargetPageId="4" captionText="Figure 2. The Layahima wuzhishana species group, wings. A. L. weiweii sp. nov., male holotype. B. L. wuzhishana (Yang, 2002), male. Abbreviations: C—costa; Sc—subcosta; RA—radius anterior; RP—radius posterior; MA—media anterior; MP—media posterior; CuA—cubitus anterior; CuP—cubitus posterior; A—anal veins. Scale bars=10.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176114" httpUri="https://zenodo.org/record/7176114/files/figure.png" pageId="5" pageNumber="294">Figs 2B</figureCitation>
,
<figureCitation id="A7902A27D800F905FD40FF15BCCAFD6F" box="[668,733,237,261]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="5" pageNumber="294">3CD</figureCitation>
,
<figureCitation id="A7902A27D800F905FD35FF15BD8DFD6E" box="[745,922,237,261]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="5" pageNumber="294">4B, D, F, H, J, L</figureCitation>
,
<figureCitation id="A7902A27D800F905FC79FF14BDF0FD6E" box="[933,999,236,260]" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="5" pageNumber="294">5AC</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="77B16529D800F905FF54FEF7BC5DFD2F" pageId="5" pageNumber="294" type="reference_group">
<paragraph id="3F1436A2D800F905FF54FEF7BA85FD4F" blockId="5.[136,1170,236,325]" box="[136,1170,271,293]" pageId="5" pageNumber="294">
<taxonomicName id="F8AB4D21D800F905FF54FEF7BCD9FD4E" ID-CoL="34QDQ" authority="Yang, In Yang &amp; Wang, 2002: 297" authorityName="Yang, In Yang &amp; Wang" authorityPageNumber="297" authorityYear="2002" box="[136,718,271,292]" class="Insecta" family="Myrmeleontidae" genus="Dendroleon" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="wuzhishanus">
<emphasis id="0DDFEAB0D800F905FF54FEF7BF6CFD4E" box="[136,379,271,292]" italics="true" pageId="5" pageNumber="294">Dendroleon wuzhishanus</emphasis>
<materialsCitation id="8FC33CFFD800F905FE5DFEF7BCD9FD4E" box="[385,718,271,292]" country="China" location="Mt. Wuzhishan" pageId="5" pageNumber="294" specimenCount="1" stateProvince="Hainan">
Yang,
<emphasis id="0DDFEAB0D800F905FE62FEF7BFC5FD4E" box="[446,466,271,292]" italics="true" pageId="5" pageNumber="294">In</emphasis>
Yang &amp; Wang, 2002: 297
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.
<typeStatus id="E0108800D800F905FD0BFEF7BD10FD4E" box="[727,775,271,292]" pageId="5" pageNumber="294">Type</typeStatus>
locality:
<collectingCountry id="47BC7632D800F905FCBDFEF7BD8AFD4F" box="[865,925,271,293]" name="China" pageId="5" pageNumber="294">China</collectingCountry>
,
<collectingRegion id="FD6FF840D800F905FC7AFEF7BDF9FD4F" box="[934,1006,271,293]" country="China" name="Hainan" pageId="5" pageNumber="294">Hainan</collectingRegion>
,
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.
</paragraph>
<paragraph id="3F1436A2D800F905FF54FED7BC5DFD2F" blockId="5.[136,1170,236,325]" box="[136,586,303,325]" pageId="5" pageNumber="294">
<treatmentCitationGroup id="1FBB118CD800F905FF54FED7BC5DFD2F" box="[136,586,303,325]" pageId="5" pageNumber="294">
<treatmentCitation id="BE0A10B3D800F905FF54FED7BC51FD2F" author="Wang" box="[136,582,303,325]" page="65" pageId="5" pageNumber="294" year="2018">
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<emphasis id="0DDFEAB0D800F905FF54FED7BF7EFD2E" box="[136,361,303,324]" italics="true" pageId="5" pageNumber="294">Layahima wuzhishanus</emphasis>
:
<bibRefCitation id="5B3A4B53D800F905FEA9FED7BC51FD2F" author="Wang" box="[373,582,303,325]" pageId="5" pageNumber="294" pagination="1 - 285" refId="ref6570" refString="Wang, X. L., Zhan, Q. B., Wang, A. Q. 2018. Fauna Sinica Insecta Vol. 68 Neuroptera Myrmeleontoidea. Science Press, Beijing. pp. 1 - 285." type="book chapter" year="2018">
Wang
<emphasis id="0DDFEAB0D800F905FE6FFED7BFF4FD2E" box="[435,483,303,324]" italics="true" pageId="5" pageNumber="294">et al.</emphasis>
, 2018: 65
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.
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</paragraph>
</subSubSection>
<subSubSection id="77B16529D800F907FF61FE9CBFA8FE35" lastPageId="7" lastPageNumber="296" pageId="5" pageNumber="294" type="diagnosis">
<paragraph id="3F1436A2D800F905FF61FE9CBB60FE26" blockId="5.[136,1452,356,1108]" pageId="5" pageNumber="294">Diagnosis. Body overall pale yellow. Antennae nearly four times as long as pronotum; vertex with a black transverse band anteriorly, a pair of large dark brown markings present medially and extending to lateral part, a pair of dark brown spots present medially; frons with a nearly pentagonal dark brown marking. Pronotum with a pair of curved dark brown stripes and some dark brown dots medially; mesonotum with a pair of tufts consisted of long black setae, and with a pair of long trapezoid dark brown stripes laterally. Tibial spurs long, barely curved. Abdominal terga 38 each with an arrow-shaped pale yellowish marking. Male sternum 9 pentagonal, basally narrow; median part of gonocoxites 11 prominent and wide in ventral view. Female anterior gonocoxites 8 absent; gonapophyses 8 sclerotized; digging setae on gonocoxites 9 blunt.</paragraph>
<paragraph id="3F1436A2D800F905FF61FDAFBBB2FE04" blockId="5.[136,1452,356,1108]" box="[189,1445,598,623]" pageId="5" pageNumber="294">
Redescription. Size. Head width
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, forewing length
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, hind wing length
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Head. Vertex raised, covered with some short setae, a black transverse band present anteriorly, a transverse dark brown lines present medially, a pair of large dark brown markings present medially and extending to lateral part, and a pair of dark brown spots present posteromedially (
<figureCitation id="A7902A27D800F905FDC7FD47BC79FEBD" box="[539,622,703,727]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="5" pageNumber="294">Fig. 3C</figureCitation>
). Antennae nearly four times as long as pronotum; scape and pedicel generally pale yellow, covered with some short setae; non-intumescent part of flagellum generally pale yellow, each segment with a brown marking; intumescent part of flagellum generally black dorsally and pink ventrally. Frons with a nearly pentagonal dark brown marking (
<figureCitation id="A7902A27D800F905FEAEFCDFBFD0FF55" box="[370,455,807,831]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="5" pageNumber="294">Fig. 3D</figureCitation>
). Clypeus pale yellow with a few setae. Labrum pale brown. Maxillary palpus generally pale yellow. Labial palpus pale brown, apical segment fusiform, with a brown marking. Mandibles pale yellowish brown with distal half dark brown.
</paragraph>
<paragraph id="3F1436A2D800F905FF61FC77BD46F85B" blockId="5.[136,1452,356,1108]" pageId="5" pageNumber="294">
Thorax (
<figureCitation id="A7902A27D800F905FEC5FC77BF7AFFCD" box="[281,365,911,935]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="5" pageNumber="294">Fig. 3C</figureCitation>
). Pale yellow, with dark markings. Pronotum longer than wide, covered with some long black setae; a pair of curved dark brown stripes and some dark brown dots present medially, and a pair of longitudinal black stripe present laterally. Mesonotum covered with a pair of tufts consisted of long black setae anterolaterally; anterior part generally dark brown, laterally with a pair of long trapezoid dark brown stripes; mesoscutellum with a pair of black markings posteriorly. Metanotum generally dark brown. Pleurae with three black stripes.
</paragraph>
<paragraph id="3F1436A2D800F906FF61FBC4BDAAFD9D" blockId="5.[136,1452,356,1108]" lastBlockId="6.[136,1452,236,1023]" lastPageId="6" lastPageNumber="295" pageId="5" pageNumber="294">Legs. Short, pale yellow, with dark markings. All femora covered with many short black setae; all tibiae covered with some long black setae; tibial spurs brown, elongate and barely curved; tarsomeres 14 generally brown; tarsomere 5 brown with distal half dark brown, and with a few short tapered setae ventrally. Foreleg: Coxa pale yellow, with a brown marking; femur generally pale yellow, dark brown dorsally; tibia generally pale yellow, an irregular brown marking present basally, distal part dark brown; tibial spurs reaching base of tarsomere 5. Midleg: Coxa generally pale yellow, brown basally; femur generally pale yellow, pale brown distally; tibia generally pale yellow, with some brown dots, distal part dark brown; tibial spurs reaching base of tarsomere 5. Hind leg: Coxa generally pale yellow, brown basally; femur generally yellow, pale brown distally; tibia generally pale yellow, basal 1/3 with a short transverse dark brown line, distal part dark brown; tibial spurs reaching tip of tarsomere 4; tarsomere 1 longer than that of fore- and midleg.</paragraph>
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Figure 3. The
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<emphasis id="0DDFEAB0D800F905FEFEF83DBFEEFBB0" box="[290,505,1989,2010]" italics="true" pageId="5" pageNumber="294">Layahima wuzhishana</emphasis>
</taxonomicName>
species group. A.
<taxonomicName id="F8AB4D21D800F905FD75F83DBD1CFBB0" authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" box="[681,779,1989,2010]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis id="0DDFEAB0D800F905FD75F83DBD1CFBB0" box="[681,779,1989,2010]" italics="true" pageId="5" pageNumber="294">L. weiweii</emphasis>
</taxonomicName>
<emphasis id="0DDFEAB0D800F905FCCDF83DBD4AFBB0" bold="true" box="[785,861,1989,2010]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="16EC57CBD800F905FCCDF83DBD4AFBB0" box="[785,861,1989,2010]" pageId="5" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, head and thorax, dorsal view. B.
<taxonomicName id="F8AB4D21D800F905FAEAF83DBB8FFBB0" authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" box="[1334,1432,1989,2010]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis id="0DDFEAB0D800F905FAEAF83DBB8FFBB0" box="[1334,1432,1989,2010]" italics="true" pageId="5" pageNumber="294">L. weiweii</emphasis>
</taxonomicName>
<emphasis id="0DDFEAB0D800F905FF47F81DBEFEFB90" bold="true" box="[155,233,2021,2042]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="16EC57CBD800F905FF47F81DBEFEFB90" box="[155,233,2021,2042]" pageId="5" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, frons. C.
<taxonomicName id="F8AB4D21D800F905FE37F81DBD17FB90" authority="(Yang, 2002)" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[491,768,2021,2042]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D800F905FE37F81DBC61FB90" box="[491,630,2021,2042]" italics="true" pageId="5" pageNumber="294">L. wuzhishana</emphasis>
(
<bibRefCitation id="5B3A4B53D800F905FD59F81DBCEFFB90" author="Yang" box="[645,760,2021,2042]" pageId="5" pageNumber="294" pagination="296 - 299" refId="ref6663" refString="Yang, C. K., Wang, X. L. 2002. Neuroptera: Myrmeleontidae. In: Huang, F. S. (ed.), Forest Insects of Hainan. Science Press, Beijing. pp. 296 - 299." type="book chapter" year="2002">Yang, 2002</bibRefCitation>
)
</taxonomicName>
, male, head and thorax, dorsal view. D.
<taxonomicName id="F8AB4D21D800F905FB50F81DBB00FB90" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[1164,1303,2021,2042]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D800F905FB50F81DBB00FB90" box="[1164,1303,2021,2042]" italics="true" pageId="5" pageNumber="294">L. wuzhishana</emphasis>
</taxonomicName>
, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="3F1436A2D803F906FF61FDFABEABFE80" blockId="6.[136,1452,236,1023]" pageId="6" pageNumber="295">
Wings (
<figureCitation id="A7902A27D803F906FECCFDFABF74FE70" box="[272,355,514,538]" captionStart="Figure 2" captionStartId="4.[155,218,1987,2009]" captionTargetBox="[189,1277,358,1942]" captionTargetId="figure-71@4.[189,1398,348,1942]" captionTargetPageId="4" captionText="Figure 2. The Layahima wuzhishana species group, wings. A. L. weiweii sp. nov., male holotype. B. L. wuzhishana (Yang, 2002), male. Abbreviations: C—costa; Sc—subcosta; RA—radius anterior; RP—radius posterior; MA—media anterior; MP—media posterior; CuA—cubitus anterior; CuP—cubitus posterior; A—anal veins. Scale bars=10.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176114" httpUri="https://zenodo.org/record/7176114/files/figure.png" pageId="6" pageNumber="295">Fig. 2B</figureCitation>
). Long, distally rounded. Banksian lines absent. Male pilula axillaris absent. Pterostigma pale. Forewing slightly longer than hindwing. Veins dark brown alternating with pale. Forewing subcostal area with a series of discontinuous dark brown markings and dots; mediocubital area with some indistinct brown markings; rhegma present as two indistinct brown markings; base of cubital area with a brown spot; crossvein on posterior costal space mostly branched; RP origin align with CuA fork; presectoral area with four to five crossveins; RP with eight to nine branches. Hindwing generally transparent; subcostal area with a few indistinct brownish dots; presectoral area with one crossvein; RP origin basal to MP fork.
</paragraph>
<paragraph id="3F1436A2D803F907FF61FD0CBA33FD6E" blockId="6.[136,1452,236,1023]" lastBlockId="7.[136,1452,236,607]" lastPageId="7" lastPageNumber="296" pageId="6" pageNumber="295">
Abdomen. Terga 12 generally dark brown; terga 38 each with an arrow-shaped pale yellowish marking. Male genitalia: Tergum 8 subrectanglar in lateral view. Sternum 9 covered with some long setae, pentagonal, basally narrow (
<figureCitation id="A7902A27D803F906FA5CFCEFBEBDFF38" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="6" pageNumber="295">Fig. 4B</figureCitation>
). Gonocoxites 9 medially curved internally; posterior part present as a pair of elongate plates, apex strongly sclerotized and scaly (
<figureCitation id="A7902A27D803F906FF25FCA4BF5DFF1F" box="[249,330,860,885]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="6" pageNumber="295">Fig. 4F</figureCitation>
). Gonocoxites 11 arrow-shaped, median part prominent and wide in ventral view (
<figureCitation id="A7902A27D803F906FB70FCA4BAEAFF1F" box="[1196,1277,860,885]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="6" pageNumber="295">Fig. 4F</figureCitation>
). Gonapophyses 11 weakly developed. Ectoproct covered with some long setae, rounded in lateral view (
<figureCitation id="A7902A27D803F906FBF0FC87BA95FFFD" box="[1068,1154,895,919]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="6" pageNumber="295">Fig. 4D</figureCitation>
). Female genitalia: Sternum 7 covered with many long setae. Pregenital plate sclerotized and rounded, external part with a tiny triangular prominent (
<figureCitation id="A7902A27D803F906FA5CFC5ABEBDFFB6" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="6" pageNumber="295">Fig. 5B</figureCitation>
). Anterior gonocoxites 8 absent; posterior gonocoxites 8 covered with many soft setae, thumb-shaped, depressed laterally (
<figureCitation id="A7902A27D803F906FF4CFC1FBF14FF95" box="[144,259,999,1023]" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="6" pageNumber="295">Fig. 5BC</figureCitation>
). Gonapophyses 8 sclerotized, anterior part wider than posterior part (
<figureCitation id="A7902A27D803F906FC3FFC1FBA21FF95" box="[995,1078,999,1023]" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="6" pageNumber="295">Fig. 5B</figureCitation>
). Gonocoxites 9 oval, covered with many blunt digging setae (
<figureCitation id="A7902A27D802F907FE78FF14BFEFFD6E" box="[420,504,236,260]" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="7" pageNumber="296">Fig. 5B</figureCitation>
). Ectoproct covered with some long setae (
<figureCitation id="A7902A27D802F907FC1FFF14BA00FD6E" box="[963,1047,236,260]" captionStart="Figure 5" captionStartId="7.[155,218,1639,1661]" captionTargetBox="[189,1398,662,1586]" captionTargetId="figure-305@7.[189,1398,662,1599]" captionTargetPageId="7" captionText="Figure 5. Layahima wuzhishana (Yang, 2002), female holotype. A. Habitus, type label and collecting label. B. Female holotype, genitalia, ventral view. C. Female holotype, genitalia, ventral view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp— gonapophyses; pp—pregenital plate; S—sternum. Scale bars: A=5.0mm; BC=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176120" httpUri="https://zenodo.org/record/7176120/files/figure.png" pageId="7" pageNumber="296">Fig. 5C</figureCitation>
).
</paragraph>
<caption id="6BD4662AD803F906FF47F85FBCD7F477" ID-DOI="http://doi.org/10.5281/zenodo.7176118" ID-Zenodo-Dep="7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="6" pageNumber="295" startId="6.[155,218,1959,1981]" targetBox="[190,1398,1083,1897]" targetPageId="6">
<paragraph id="3F1436A2D803F906FF47F85FBCD7F477" blockId="6.[155,1433,1959,2077]" pageId="6" pageNumber="295">
Figure 4. The
<taxonomicName id="F8AB4D21D803F906FEE4F85FBC01FBD6" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[312,534,1959,1980]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D803F906FEE4F85FBC01FBD6" box="[312,534,1959,1980]" italics="true" pageId="6" pageNumber="295">Layahima wuzhishana</emphasis>
</taxonomicName>
species group, male terminalia and genitalia. A, C, E, G, I, K.
<taxonomicName id="F8AB4D21D803F906FB11F850BB20FBD7" authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" box="[1229,1335,1960,1981]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis id="0DDFEAB0D803F906FB11F850BB20FBD7" box="[1229,1335,1960,1981]" italics="true" pageId="6" pageNumber="295">L. weiweii</emphasis>
</taxonomicName>
<emphasis id="0DDFEAB0D803F906FA98F850BB8FFBD7" bold="true" box="[1348,1432,1960,1981]" pageId="6" pageNumber="295">
<taxonomicNameLabel id="16EC57CBD803F906FA98F850BB8FFBD7" box="[1348,1432,1960,1981]" pageId="6" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
B, D, F, H, J, L.
<taxonomicName id="F8AB4D21D803F906FEE2F830BC46FBB7" authority="(Yang, 2002)" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[318,593,1991,2013]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D803F906FEE2F830BFDEFBB6" box="[318,457,1991,2013]" italics="true" pageId="6" pageNumber="295">L. wuzhishana</emphasis>
(
<bibRefCitation id="5B3A4B53D803F906FE0AF830BC5DFBB7" author="Yang" box="[470,586,1992,2013]" pageId="6" pageNumber="295" pagination="296 - 299" refId="ref6663" refString="Yang, C. K., Wang, X. L. 2002. Neuroptera: Myrmeleontidae. In: Huang, F. S. (ed.), Forest Insects of Hainan. Science Press, Beijing. pp. 296 - 299." type="book chapter" year="2002">Yang, 2002</bibRefCitation>
)
</taxonomicName>
. AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars =1.0 mm.
</paragraph>
</caption>
<paragraph id="3F1436A2D802F907FF61FEF7BF10FDD8" blockId="7.[136,1452,236,607]" pageId="7" pageNumber="296">
Material examined.
<materialsCitation id="8FC33CFFD802F907FE4DFEF7BC6DFD20" collectingDate="1997-06-01" collectionCode="CAU" collectorName="Chikun Yang &amp; Jiafang Cheng" country="China" elevation="800" location="Wuzhishan International Resort" municipality="Mt. Wuzhishan" pageId="7" pageNumber="296" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="E0108800D802F907FE4DFEF7BFEFFD4D" box="[401,504,271,295]" pageId="7" pageNumber="296" type="holotype">Holotype</typeStatus>
.
<specimenCount id="29ADFD2BD802F907FDDDFEE8BC02FD4D" box="[513,533,272,295]" pageId="7" pageNumber="296" type="female"></specimenCount>
,
<collectingCountry id="47BC7632D802F907FDFDFEF7BC72FD4D" box="[545,613,271,295]" name="China" pageId="7" pageNumber="296">China</collectingCountry>
:
<collectingRegion id="FD6FF840D802F907FDB1FEF7BCAAFD4D" box="[621,701,271,295]" country="China" name="Hainan" pageId="7" pageNumber="296">Hainan</collectingRegion>
,
<collectingMunicipality id="DF70ACD8D802F907FD1BFEE8BD7CFD4D" box="[711,875,271,295]" pageId="7" pageNumber="296">Mt. Wuzhishan</collectingMunicipality>
,
<location id="3A746079D802F907FCA8FEF7BAD3FD4D" LSID="urn:lsid:plazi:treatment:B70287B4D800F907FF54FF14BFA8FE35:3A746079D802F907FCA8FEF7BAD3FD4D" box="[884,1220,271,295]" country="China" municipality="Mt. Wuzhishan" name="Wuzhishan International Resort" pageId="7" pageNumber="296" stateProvince="Hainan">Wuzhishan International Resort</location>
(
<quantity id="F8539B47D802F907FB0DFEF7BB02FD4D" box="[1233,1301,271,295]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="296" unit="m" value="800.0">
<elevation id="B486D191D802F907FB0DFEF7BB02FD4D" box="[1233,1301,271,295]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="296" unit="m" value="800.0">800 m</elevation>
</quantity>
),
<date id="4B151062D802F907FAFAFEF7BBBFFD4D" box="[1318,1448,271,295]" pageId="7" pageNumber="296" value="1997-06-01">
<collectingDate id="5B51E98AD802F907FAFAFEF7BBBFFD4D" box="[1318,1448,271,295]" pageId="7" pageNumber="296" value="1997-06-01">1 June 1997</collectingDate>
</date>
,
<collectorName id="925E5374D802F907FF54FECABF0DFD20" box="[136,282,306,330]" pageId="7" pageNumber="296">Chikun Yang</collectorName>
&amp;
<collectorName id="925E5374D802F907FE96FECABFFDFD20" box="[330,490,306,330]" pageId="7" pageNumber="296">Jiafang Cheng</collectorName>
leg. (
<collectionCode id="59BAAE67D802F907FDE8FECABC65FD20" box="[564,626,306,330]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="296">CAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8FC33CFFD802F907FD54FECABD0FFD07" collectingDate="2020-05-24" collectingDateMax="2020-05-27" collectingDateMin="2020-05-24" collectionCode="CAU" collectorName="Chao Wu" country="China" county="Ledong County" elevation="950" location="Mingfenggu" municipality="Jianfengling" pageId="7" pageNumber="296" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
Additional material.
<specimenCount id="29ADFD2BD802F907FCAFFECABD82FD20" box="[883,917,306,330]" pageId="7" pageNumber="296" type="male">1♂</specimenCount>
,
<collectingCountry id="47BC7632D802F907FC75FECABDFAFD20" box="[937,1005,306,330]" name="China" pageId="7" pageNumber="296">China</collectingCountry>
:
<collectingRegion id="FD6FF840D802F907FC22FECABA59FD20" box="[1022,1102,306,330]" country="China" name="Hainan" pageId="7" pageNumber="296">Hainan</collectingRegion>
,
<collectingCounty id="D6754E2ED802F907FB83FECABB18FD20" box="[1119,1295,306,330]" pageId="7" pageNumber="296">Ledong County</collectingCounty>
,
<collectingMunicipality id="DF70ACD8D802F907FAFDFECABBB0FD20" box="[1313,1447,306,330]" pageId="7" pageNumber="296">Jianfengling</collectingMunicipality>
,
<location id="3A746079D802F907FF54FEACBF1BFD06" LSID="urn:lsid:plazi:treatment:B70287B4D800F907FF54FF14BFA8FE35:3A746079D802F907FF54FEACBF1BFD06" box="[136,268,340,364]" country="China" county="Ledong County" municipality="Jianfengling" name="Mingfenggu" pageId="7" pageNumber="296" stateProvince="Hainan">Mingfenggu</location>
(
<quantity id="F8539B47D802F907FEC6FEACBF49FD07" box="[282,350,340,365]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="296" unit="m" value="950.0">
<elevation id="B486D191D802F907FEC6FEACBF49FD07" box="[282,350,340,365]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="296" unit="m" value="950.0">950m</elevation>
</quantity>
),
<date id="4B151062D802F907FEACFEADBC3FFD07" box="[368,552,341,365]" pageId="7" pageNumber="296" value="2020-05-24" valueMax="2020-05-27" valueMin="2020-05-24">
<collectingDate id="5B51E98AD802F907FEACFEADBC3FFD07" box="[368,552,341,365]" pageId="7" pageNumber="296" value="2020-05-24" valueMax="2020-05-27" valueMin="2020-05-24">2427 May 2020</collectingDate>
</date>
,
<collectorName id="925E5374D802F907FDEEFEACBC80FD06" box="[562,663,340,364]" pageId="7" pageNumber="296">Chao Wu</collectorName>
leg. (
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)
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;
<materialsCitation id="8FC33CFFD802F907FCFEFEADBFD4FDE5" collectingDate="2009-05-21" collectionCode="CAU" collectorName="Hu Li" country="China" county="Hekou County" elevation="132" location="Nanxi Town" pageId="7" pageNumber="296" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
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,
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:
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,
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,
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<collectingDate id="5B51E98AD802F907FA4CFEADBEEFFDE5" pageId="7" pageNumber="296" value="2009-05-21">21 May 2009</collectingDate>
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,
<collectorName id="925E5374D802F907FEDFFE80BF55FDE5" box="[259,322,375,399]" pageId="7" pageNumber="296">Hu Li</collectorName>
leg. (
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)
</materialsCitation>
;
<materialsCitation id="8FC33CFFD802F907FE12FE8FBF13FDD8" collectingDate="2012-05-22" collectionCode="CAU" collectorName="Xingyue Liu" country="Vietnam" elevation="330" location="Deo Gao Lae" municipality="Hoa An District" pageId="7" pageNumber="296" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
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,
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:
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,
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,
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(
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),
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<collectingDate id="5B51E98AD802F907FB54FE8FBB0EFDE5" box="[1160,1305,375,400]" pageId="7" pageNumber="296" value="2012-05-22">22 May 2012</collectingDate>
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,
<collectorName id="925E5374D802F907FAF8FE8FBBBCFDE5" box="[1316,1451,375,399]" pageId="7" pageNumber="296">Xingyue Liu</collectorName>
leg. (
<collectionCode id="59BAAE67D802F907FF62FE62BEEBFDD8" box="[190,252,410,434]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="296">CAU</collectionCode>
)
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.
</paragraph>
<paragraph id="3F1436A2D802F907FF61FE44BD51FDBF" blockId="7.[136,1452,236,607]" box="[189,838,444,469]" pageId="7" pageNumber="296">
Distribution.
<collectingCountry id="47BC7632D802F907FE90FE44BF9BFDBE" box="[332,396,444,468]" name="China" pageId="7" pageNumber="296">China</collectingCountry>
(
<collectingRegion id="FD6FF840D802F907FE45FE44BFFDFDBE" box="[409,490,444,468]" country="China" name="Hainan" pageId="7" pageNumber="296">Hainan</collectingRegion>
,
<collectingRegion id="FD6FF840D802F907FE29FE45BC59FDBF" box="[501,590,445,469]" country="China" name="Yunnan" pageId="7" pageNumber="296">Yunnan</collectingRegion>
);
<collectingCountry id="47BC7632D802F907FDBDFE44BCAAFDBE" box="[609,701,444,468]" name="Vietnam" pageId="7" pageNumber="296">Vietnam</collectingCountry>
(
<collectingRegion id="FD6FF840D802F907FD16FE45BD2DFDBF" box="[714,826,445,469]" country="Vietnam" name="Cao Bang" pageId="7" pageNumber="296">Cao Bang</collectingRegion>
).
</paragraph>
<paragraph id="3F1436A2D802F907FF61FE27BFA8FE35" blockId="7.[136,1452,236,607]" pageId="7" pageNumber="296">
Remarks. It is worth noting that the female gonapophyses 8 of
<taxonomicName id="F8AB4D21D802F907FC56FE18BA3FFD9D" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[906,1064,479,503]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D802F907FC56FE18BA3FFD9D" box="[906,1064,479,503]" italics="true" pageId="7" pageNumber="296">L. wuzhishana</emphasis>
</taxonomicName>
are sclerotized. However, in other members of
<taxonomicName id="F8AB4D21D802F907FED3FDFABF6DFE70" authorityName="Navas" authorityYear="1912" box="[271,378,514,538]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0DDFEAB0D802F907FED3FDFABF6DFE70" box="[271,378,514,538]" italics="true" pageId="7" pageNumber="296">Layahima</emphasis>
</taxonomicName>
with female records, the female gonapophyses 8 are absent. This unique female genital character of
<taxonomicName id="F8AB4D21D802F907FF54FDDDBF35FE56" baseAuthorityName="Yang" baseAuthorityYear="2002" box="[136,290,548,572]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis id="0DDFEAB0D802F907FF54FDDDBF35FE56" box="[136,290,548,572]" italics="true" pageId="7" pageNumber="296">L. wuzhishana</emphasis>
</taxonomicName>
indicates that the interspecific phylogeny of
<taxonomicName id="F8AB4D21D802F907FCDDFDDCBD7BFE56" authorityName="Navas" authorityYear="1912" box="[769,876,548,572]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0DDFEAB0D802F907FCDDFDDCBD7BFE56" box="[769,876,548,572]" italics="true" pageId="7" pageNumber="296">Layahima</emphasis>
</taxonomicName>
requires further study. Unfortunately, the female of
<taxonomicName id="F8AB4D21D802F907FA4AFDDDBECEFE35" authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis id="0DDFEAB0D802F907FA4AFDDDBECEFE35" italics="true" pageId="7" pageNumber="296">L. weiweii</emphasis>
</taxonomicName>
<emphasis id="0DDFEAB0D802F907FF3CFDBFBF21FE35" bold="true" box="[224,310,583,607]" pageId="7" pageNumber="296">
<taxonomicNameLabel id="16EC57CBD802F907FF3CFDBFBF21FE35" box="[224,310,583,607]" pageId="7" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,237 @@
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<mods:title>A spectacular new antlion species of the subfamily Dendroleontinae (Neuroptera: Myrmeleontidae) from Malaysia</mods:title>
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<mods:namePart>Zheng, Yuchen</mods:namePart>
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<mods:namePart>Liu, Xingyue</mods:namePart>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172476" ID-Zenodo-Dep="7172476" LSID="urn:lsid:plazi:treatment:B70287B4D807F904FF54FB3BBD4BFD6E" httpUri="http://treatment.plazi.org/id/B70287B4D807F904FF54FB3BBD4BFD6E" lastPageId="4" lastPageNumber="292" pageId="2" pageNumber="291">
<subSubSection box="[136,855,1219,1244]" pageId="2" pageNumber="291" type="nomenclature">
<paragraph blockId="2.[136,855,1219,1244]" box="[136,855,1219,1244]" pageId="2" pageNumber="291">
<heading box="[136,855,1219,1244]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="291" reason="2">
<emphasis bold="true" box="[136,428,1219,1244]" pageId="2" pageNumber="291">
<taxonomicName authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" box="[136,335,1219,1244]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis bold="true" box="[136,335,1219,1244]" italics="true" pageId="2" pageNumber="291">Layahima weiweii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[342,428,1220,1244]" pageId="2" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[461,529,1220,1244]" captionStart="Figure 1" captionStartId="3.[155,218,2054,2076]" captionTargetBox="[299,1285,1422,2003]" captionTargetId="figure-611@3.[189,1398,1417,2007]" captionTargetPageId="3" captionText="Figure 1. Layahima weiweii sp. nov., male holotype, habitus. Scale bar=10.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176112" httpUri="https://zenodo.org/record/7176112/files/figure.png" pageId="2" pageNumber="291">Figs 1</figureCitation>
,
<figureCitation box="[542,577,1220,1244]" captionStart="Figure 2" captionStartId="4.[155,218,1987,2009]" captionTargetBox="[189,1277,358,1942]" captionTargetId="figure-71@4.[189,1398,348,1942]" captionTargetPageId="4" captionText="Figure 2. The Layahima wuzhishana species group, wings. A. L. weiweii sp. nov., male holotype. B. L. wuzhishana (Yang, 2002), male. Abbreviations: C—costa; Sc—subcosta; RA—radius anterior; RP—radius posterior; MA—media anterior; MP—media posterior; CuA—cubitus anterior; CuP—cubitus posterior; A—anal veins. Scale bars=10.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176114" httpUri="https://zenodo.org/record/7176114/files/figure.png" pageId="2" pageNumber="291">2A</figureCitation>
,
<figureCitation box="[588,655,1220,1244]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="2" pageNumber="291">3AB</figureCitation>
,
<figureCitation box="[665,847,1220,1244]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="2" pageNumber="291">4A, C, E, G, I, K</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="293" pageId="2" pageNumber="291" type="diagnosis">
<paragraph blockId="2.[136,1452,1275,2062]" pageId="2" pageNumber="291">Diagnosis. Body largely yellow. Antennae nearly four times as long as pronotum; vertex with a black transverse band anteriorly, and with a pair of dark brown spots and indistinct transverse dark brown lines medially; frons with a nearly rhombic black marking. Pronotum with some indistinct brown markings; anterolateral mesonotum with a pair of tufts consisted of long black setae, and with a pair of long triangular dark brown stripes laterally. Tibial spurs long, barely curved. Abdominal terga 38 yellow alternating with dark brown. Male sternum 9 pentagonal; median part of gonocoxites 11 prominent in ventral view, protruding backward in lateral view.</paragraph>
<paragraph blockId="2.[136,1452,1275,2062]" box="[189,1421,1483,1508]" pageId="2" pageNumber="291">
Description. Size. Head width
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, forewing length
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, hind wing length
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.
</paragraph>
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Head. Vertex raised, covered with some short setae, a black transverse band present anteriorly, a pair of dark brown spots and indistinct transverse dark brown lines present medially, and a pair of dark brown markings present posterolaterally (
<figureCitation box="[144,231,1587,1612]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="2" pageNumber="291">Fig. 3A</figureCitation>
). Antennae nearly four times as long as pronotum; scape and pedicel generally yellow, covered with some short setae; flagellomeres generally yellow, each segment with a circular brown marking. Frons with a nearly rhombic black marking (
<figureCitation box="[242,327,1657,1681]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="2" pageNumber="291">Fig. 3B</figureCitation>
). Clypeus yellow with a few setae. Labrum brown. Maxillary palpus generally pale yellow, but brown distally. Labial palpus pale brown, apical segment fusiform, with a brown marking. Mandibles yellowish brown with distal half dark brown.
</paragraph>
<paragraph blockId="2.[136,1452,1275,2062]" pageId="2" pageNumber="291">
Thorax (
<figureCitation box="[281,367,1761,1785]" captionStart="Figure 3" captionStartId="5.[155,218,1989,2011]" captionTargetBox="[189,1398,1186,1942]" captionTargetId="figure-513@5.[189,1398,1186,1942]" captionTargetPageId="5" captionText="Figure 3. The Layahima wuzhishana species group. A. L. weiweii sp. nov., male holotype, head and thorax, dorsal view. B. L. weiweii sp. nov., male holotype, frons. C. L. wuzhishana (Yang, 2002), male, head and thorax, dorsal view. D. L. wuzhishana, male, frons. Red arrow indicates the tuft of long black setae on mesonotum. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176116" httpUri="https://zenodo.org/record/7176116/files/figure.png" pageId="2" pageNumber="291">Fig. 3A</figureCitation>
). Yellow, with dark markings. Pronotum longer than wide, covered with some long black setae; three indistinct brown markings present anteriorly, a pair of longitudinal black stripes present laterally, and an indistinct brown marking present medially. Mesonotum covered with a pair of tufts consisted of long black setae anterolaterally; anterior part generally dark brown, laterally with a pair of long triangular dark brown stripes and a pair of small spots; mesoscutellum with a pair of black marking posteriorly. Metanotum generally dark brown. Pleurae with three black stripes.
</paragraph>
<paragraph blockId="2.[136,1452,1275,2062]" lastBlockId="3.[136,1452,236,1370]" lastPageId="3" lastPageNumber="292" pageId="2" pageNumber="291">Legs. Short, pale yellow, with dark markings. All femora covered with many short black setae; all tibiae covered with some long black setae; tibial spurs brown, elongate and barely curved; tarsomeres 14 generally brown; tarsomeres 5 brown, with distal half dark brown, and with a few short tapered setae ventrally. Foreleg: Coxa brown, with a dark brown spot; femur generally yellow, dark brown dorsally; tibia generally pale yellow, an irregular brown marking present basally, distal part dark brown; tibial spurs reaching base of tarsomere 5. Midleg: Coxa with basal half black and distal half yellow; femur generally yellow, dark brown distally; tibia generally pale yellow, with some brown dots, basal 1/3 with a dark brown spot, distal part dark brown; tibial spurs reaching base of tarsomere 5. Hind leg: Coxa with basal half black and distal half yellow; femur generally yellow, dark brown distally; tibia generally pale yellow, basal 1/3 with a dark brown spot, distal part dark brown; tibial spurs reaching tip of tarsomere 4; tarsomere 1 longer than that of fore- and midleg.</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" pageId="3" pageNumber="292">
Wings (
<figureCitation box="[272,357,410,434]" captionStart="Figure 2" captionStartId="4.[155,218,1987,2009]" captionTargetBox="[189,1277,358,1942]" captionTargetId="figure-71@4.[189,1398,348,1942]" captionTargetPageId="4" captionText="Figure 2. The Layahima wuzhishana species group, wings. A. L. weiweii sp. nov., male holotype. B. L. wuzhishana (Yang, 2002), male. Abbreviations: C—costa; Sc—subcosta; RA—radius anterior; RP—radius posterior; MA—media anterior; MP—media posterior; CuA—cubitus anterior; CuP—cubitus posterior; A—anal veins. Scale bars=10.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176114" httpUri="https://zenodo.org/record/7176114/files/figure.png" pageId="3" pageNumber="292">Fig. 2A</figureCitation>
). Long, distally rounded. Banksian lines absent. Male pilula axillaris absent. Pterostigma pale. Forewing slightly longer than hindwing. Veins dark brown alternating with pale stripes. Forewing subcostal area with a series of discontinuous dark brown markings and dots; mediocubital area with some indistinct brown marking; rhegma present as two indistinct brown markings; base of cubital area with a brown spot; crossvein on posterior costal space mostly branched; RP origin with CuA fork alignment; presectoral area with five crossveins; RP with eight branches. Hindwing generally transparent; subcostal area with some brown dots; presectoral area with one crossvein; RP origin basal to MP fork.
</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" pageId="3" pageNumber="292">
Abdomen. Tergum 1 generally pale yellow, dark brown basally; tergum 2 generally dark brown; Yellow alternating with dark brown on terga 38. Male genitalia: Tergum 8 subrectanglar in lateral view. Sternum 9 covered with some long setae, pentagonal (
<figureCitation box="[270,359,687,711]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="3" pageNumber="292">Fig. 4A</figureCitation>
). Gonocoxites 9 medially curved internally; posterior part present as a pair of elongate plates, apex strongly sclerotized and scaly (
<figureCitation box="[471,556,722,746]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="3" pageNumber="292">Fig. 4E</figureCitation>
). Gonocoxites 11 arrow-shaped, median part prominent in ventral view; protruding backward in lateral view (
<figureCitation box="[420,539,756,781]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="3" pageNumber="292">Fig. 4E, G</figureCitation>
). Gonapophyses 11 weakly developed. Ectoproct covered with long setae (
<figureCitation box="[1353,1440,756,781]" captionStart="Figure 4" captionStartId="6.[155,218,1959,1981]" captionTargetBox="[190,1398,1083,1897]" captionTargetId="figure-505@6.[189,1399,1077,1908]" captionTargetPageId="6" captionText="Figure 4. The Layahima wuzhishana species group, male terminalia and genitalia. A, C, E, G, I, K. L. weiweii sp. nov. B, D, F, H, J, L. L. wuzhishana (Yang, 2002). AB. Terminalia, ventral view. CD. Terminalia, lateral view. EF. Genitalia, ventral view. GH. Genitalia, lateral view. IJ. Dorsal view. KL. Genitalia, caudal view. Abbreviations: ect—ectoproct; gx—gonocoxites; gp—gonapophyses; S9—sternum 9. Scale bars=1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7176118" httpUri="https://zenodo.org/record/7176118/files/figure.png" pageId="3" pageNumber="292">Fig. 4C</figureCitation>
). Female genitalia unknown.
</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" pageId="3" pageNumber="292">
Type material.
<materialsCitation collectingDate="2018-04-30" collectionCode="CAU" collectorName="Zhuo Chen" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus box="[351,454,826,850]" pageId="3" pageNumber="292">Holotype</typeStatus>
.
<specimenCount box="[463,485,826,849]" pageId="3" pageNumber="292" type="male"></specimenCount>
,
<collectingCountry box="[496,598,826,850]" name="Malaysia" pageId="3" pageNumber="292">Malaysia</collectingCountry>
:
<collectingRegion box="[608,676,826,850]" country="Malaysia" name="Sabah" pageId="3" pageNumber="292">Sabah</collectingRegion>
,
<collectingMunicipality box="[686,835,826,850]" pageId="3" pageNumber="292">Mt. Trusmadi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:B70287B4D807F904FF54FB3BBD4BFD6E:3A746079D806F903FC91FCC2BA49FF38" box="[845,1118,826,850]" country="Malaysia" municipality="Mt. Trusmadi" name="Borneo Jungle Girl Camp" pageId="3" pageNumber="292" stateProvince="Sabah">Borneo Jungle Girl Camp</location>
(
<quantity box="[1133,1215,826,850]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="292" unit="m" value="1200.0">
<elevation box="[1133,1215,826,850]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="292" unit="m" value="1200.0">1200 m</elevation>
</quantity>
),
<date box="[1233,1385,826,850]" pageId="3" pageNumber="292" value="2018-04-30">
<collectingDate box="[1233,1385,826,850]" pageId="3" pageNumber="292" value="2018-04-30">30 April 2018</collectingDate>
</date>
,
<collectorName pageId="3" pageNumber="292">Zhuo Chen</collectorName>
leg. (
<collectionCode box="[252,314,861,885]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation box="[331,1226,860,885]" collectingDate="2015-09-27" collectionCode="CAU" collectorName="Chao Wu" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus box="[331,438,861,885]" pageId="3" pageNumber="292">Paratypes</typeStatus>
.
<specimenCount box="[448,482,861,885]" pageId="3" pageNumber="292" type="male">1♂</specimenCount>
, same locality as holotype,
<date box="[776,985,860,885]" pageId="3" pageNumber="292" value="2015-09-27">
<collectingDate box="[776,985,860,885]" pageId="3" pageNumber="292" value="2015-09-27">27 September 2015</collectingDate>
</date>
,
<collectorName box="[995,1097,860,884]" pageId="3" pageNumber="292">Chao Wu</collectorName>
leg. (
<collectionCode box="[1156,1218,861,885]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2017-05-01" collectingDateMax="2017-05-07" collectingDateMin="2017-05-01" collectionCode="CAU" collectorName="Chao Wu" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount box="[1236,1271,861,885]" pageId="3" pageNumber="292" type="male">1♂</specimenCount>
, same locality as holotype,
<date box="[243,401,895,920]" pageId="3" pageNumber="292" value="2017-05-01" valueMax="2017-05-07" valueMin="2017-05-01">
<collectingDate box="[243,401,895,920]" pageId="3" pageNumber="292" value="2017-05-01" valueMax="2017-05-07" valueMin="2017-05-01">17 May 2017</collectingDate>
</date>
,
<collectorName box="[412,514,895,919]" pageId="3" pageNumber="292">Chao Wu</collectorName>
leg. (
<collectionCode box="[573,635,895,919]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[654,1340,895,919]" collectingDate="2016-04-27" collectionCode="CAU" collectorName="Hu Li" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<specimenCount box="[654,689,895,919]" pageId="3" pageNumber="292" type="male">3♂</specimenCount>
, same locality as holotype,
<date box="[984,1136,895,919]" pageId="3" pageNumber="292" value="2016-04-27">
<collectingDate box="[984,1136,895,919]" pageId="3" pageNumber="292" value="2016-04-27">27 April 2016</collectingDate>
</date>
,
<collectorName box="[1147,1210,895,919]" pageId="3" pageNumber="292">Hu Li</collectorName>
leg. (
<collectionCode box="[1270,1332,895,919]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2016-04-29" collectionCode="CAU" collectorName="Xiumei Lu" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<specimenCount box="[1350,1386,895,919]" pageId="3" pageNumber="292" type="male">2♂</specimenCount>
, same locality as holotype,
<date box="[358,509,930,954]" pageId="3" pageNumber="292" value="2016-04-29">
<collectingDate box="[358,509,930,954]" pageId="3" pageNumber="292" value="2016-04-29">29 April 2016</collectingDate>
</date>
,
<collectorName box="[520,636,930,954]" pageId="3" pageNumber="292">Xiumei Lu</collectorName>
leg. (
<collectionCode box="[696,758,930,954]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2019-05-05" collectionCode="CAU" collectorName="Guanyuan Cao" country="Malaysia" elevation="1200" location="Borneo Jungle Girl Camp" municipality="Mt. Trusmadi" pageId="3" pageNumber="292" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount box="[776,811,930,954]" pageId="3" pageNumber="292" type="male">1♂</specimenCount>
, same locality as holotype,
<date box="[1105,1236,930,954]" pageId="3" pageNumber="292" value="2019-05-05">
<collectingDate box="[1105,1236,930,954]" pageId="3" pageNumber="292" value="2019-05-05">5 May 2019</collectingDate>
</date>
,
<collectorName box="[1247,1406,930,954]" pageId="3" pageNumber="292">Guanyuan Cao</collectorName>
leg. (
<collectionCode box="[144,206,965,989]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="292">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" box="[189,526,999,1023]" pageId="3" pageNumber="292">
Distribution.
<collectingCountry box="[332,430,999,1023]" name="Malaysia" pageId="3" pageNumber="292">Malaysia</collectingCountry>
(
<collectingRegion box="[444,515,999,1023]" country="Malaysia" name="Sabah" pageId="3" pageNumber="292">Sabah</collectingRegion>
).
</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" pageId="3" pageNumber="292">
Etymology. The new species is dedicated to the entomologist, science writer and amber collector Weiwei Zhang (
<collectingRegion box="[144,268,1068,1092]" country="China" name="Chongqing" pageId="3" pageNumber="292">Chongqing</collectingRegion>
), who kindly helped us to obtain these important specimens.
</paragraph>
<paragraph blockId="3.[136,1452,236,1370]" lastBlockId="4.[136,860,236,261]" lastPageId="4" lastPageNumber="293" pageId="3" pageNumber="292">
Remarks. This species is very similar to
<taxonomicName baseAuthorityName="Yang" baseAuthorityYear="2002" box="[620,774,1103,1127]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis box="[620,774,1103,1127]" italics="true" pageId="3" pageNumber="292">L. wuzhishana</emphasis>
</taxonomicName>
, but can be distinguished from the latter by its yellow body, the vertex with a pair of dark brown spots and indistinct transverse dark brown lines, the pronotum with indistinct brown markings anteriorly and medially, the mesonotum with a pair of long triangular dark brown stripes laterally, the median part of male gonocoxites 11 narrower in ventral view, and the anterior gonocoxites 11 protruding backward in lateral view.
<taxonomicName baseAuthorityName="Yang" baseAuthorityYear="2002" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="wuzhishana">
<emphasis italics="true" pageId="3" pageNumber="292">L. wuzhishana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName authority="Zheng &amp; Liu, 2021" authorityName="Zheng &amp; Liu" authorityYear="2021" box="[549,657,1243,1266]" class="Insecta" family="Myrmeleontidae" genus="Layahima" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis box="[549,657,1243,1266]" italics="true" pageId="3" pageNumber="292">L. weiweii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[664,750,1242,1266]" pageId="3" pageNumber="292">
<taxonomicNameLabel box="[664,750,1242,1266]" pageId="3" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the following characters: the body pale yellow, the vertex with a pair of large dark brown markings medially and then extending to lateral part, the pronotum with a pair of curved dark brown stripes medially, the mesonotum with a pair of long trapezoid dark brown stripes laterally, the abdominal terga 38 each with an arrow-shaped pale yellowish marking, the median part of male gonocoxites 11 prominent and wide in ventral view, and the anterior gonocoxites 11 not protruding in lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/abs.6.e54151" ID-GBIF-Dataset="ffbe838f-508c-4c3d-ad77-20349236c7ea" ID-Pensoft-Pub="2412-1908-6-249" ID-Pensoft-UUID="871DBC1F1DFB5B7A8150200B335888A9" ID-ZooBank="BD65A575E6AB4E97B3EB199B17BA64A9" ModsDocID="2412-1908-6-249" checkinTime="1633362629850" checkinUser="pensoft" docAuthor="Vinokurov, Nikolay N. &amp; Rudoi, Valentin V." docDate="2020" docId="B7029A56647E53328EF52640D2B1859C" docLanguage="en" docName="FF9BFFAEFF8EFFFFFFF2AD03FFCF4D73" docOrigin="Acta Biologica Sibirica 6" docPubDate="2020-09-18" docSource="http://dx.doi.org/10.3897/abs.6.e54151" docTitle="Coranus aethiops Jakovlev 1893" docType="treatment" docVersion="4" id="871DBC1F1DFB5B7A8150200B335888A9" lastPageNumber="249" masterDocId="871DBC1F1DFB5B7A8150200B335888A9" masterDocTitle="Materials on the fauna of true bugs (Heteroptera) of East Kazakhstan Region of the Republic of Kazakhstan" masterLastPageNumber="277" masterPageNumber="249" pageNumber="249" updateTime="1635288858150" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Materials on the fauna of true bugs (Heteroptera) of East Kazakhstan Region of the Republic of Kazakhstan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vinokurov, Nikolay N.</mods:namePart>
<mods:affiliation>Institute for Biological Problems of Cryolithozone, Siberian Branch RAS, 41 Lenin Av., Yakutsk, 677980, Russia</mods:affiliation>
<mods:nameIdentifier type="email">vinok@ibpc.ysn.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rudoi, Valentin V.</mods:namePart>
<mods:affiliation>Altai State University, 61 Lenin Av., Barnaul, 656049, Russia</mods:affiliation>
</mods:name>
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<mods:date>2020</mods:date>
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<mods:number>2020-09-18</mods:number>
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<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/abs.6.e54151</mods:identifier>
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<mods:identifier type="Pensoft-UUID">871DBC1F1DFB5B7A8150200B335888A9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="189470697" LSID="urn:lsid:plazi:treatment:B7029A56647E53328EF52640D2B1859C" httpUri="http://treatment.plazi.org/id/B7029A56647E53328EF52640D2B1859C" lastPageNumber="249" pageId="0" pageNumber="249">
<subSubSection pageId="0" pageNumber="249" type="nomenclature">
<paragraph pageId="0" pageNumber="249">
<taxonomicName LSID="B7029A56-647E-5332-8EF5-2640D2B1859C" authority="Jakovlev, 1893" authorityName="Jakovlev" authorityYear="1893" class="Insecta" family="Reduviidae" genus="Coranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coranus aethiops" order="Reduviidae" pageId="0" pageNumber="249" phylum="Arthropoda" rank="species" species="Aethiops">Coranus aethiops Jakovlev, 1893</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="249" type="materials_examined">
<paragraph pageId="0" pageNumber="249">Material.</paragraph>
<paragraph pageId="0" pageNumber="249">
Berelsk Vill.,
<date value="2017-09-27">27.09.2017</date>
(G. Amanbaeva)
<specimenCount type="female">1 female</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="249" type="distribution">
<paragraph pageId="0" pageNumber="249">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="249">Euro-Siberian. First record from East Kazakhstan Region.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F646D0563607E86CAEE5D8FA2AD4E21F">Chapter 7: Linnaean Plant Names and their Types (part C)</mods:title>
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<treatmentCitation id="E43588EA820762B4AF41837CF4D7D968" author="Linnaeus" pageNumber="909" year="1753">
<bibCitation id="276779231EF7FEE2B2F6AD82939F2694" author="Linnaeus" pagination="909" title="Coreopsis alternifolia" volumeTitle="Species Plantarum 2" year="1753">
<taxonomicName id="AD02538F86FEC4993C5DF4089FCAB1FB" ID-CoL="YDLZ" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Coreopsis" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="alternifolia">
<emphasis id="87ED0FC3AF9C16732FA5AA7A89787719" bold="true" pageNumber="446">Coreopsis alternifolia</emphasis>
Linnaeus
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,
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<emphasis id="34A4B7B5E53187EAECC1CD09AEA3B9F5" italics="true" pageNumber="446">Species Plantarum</emphasis>
2
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: 909. 1753
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.
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<paragraph id="C1E24685D8B2D7DEB5B723ADE454C78C" pageNumber="446">&quot;Habitat in Virginia, Canada.&quot; RCN: 6566.</paragraph>
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<paragraph id="563D6B4D1EDA3C7B8DB567BE96D8285C" pageNumber="446">
<materialsCitation id="1197B6150B630EC439B8D605E2D484BE" ID-GBIF-Occurrence="2612072316" collectionCode="LINN" specimenCode="1026.11" typeStatus="lectotype">
<typeStatus id="25F940C61EB5A0A818A66F3F8767754F">Lectotype</typeStatus>
(Reveal in Jarvis &amp; Turland in
<emphasis id="9251935A99FD4A1BBA90E7D928037671" italics="true" pageNumber="446">Taxon</emphasis>
47: 359. 1998): Herb. Linn. No. 1026.11 (
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)
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.
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Current name:
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<emphasis id="D950BFC1749B22B473B4A8EFBE2A9CFC" bold="true" italics="true" pageNumber="446">Verbesina alternifolia</emphasis>
</emphasis>
(L.) Britton ex Kearney
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(
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).
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