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<document id="11FFA0112C1D6C0326D09469E9E6A008" ID-DOI="10.11646/zootaxa.5507.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757868" ID-ZooBank="76B5DE79-1A25-47D7-B46E-11531AD37F92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726221892432" checkinUser="plazi" docAuthor="Lopes, Rogério Botion, Moleiro, Henrique Pereira &amp; Lucena, Daercio A. A." docDate="2024" docId="007A1F0BFFDB1F59FF45FA5A929DF867" docLanguage="en" docName="zootaxa.5507.3.3.pdf" docOrigin="Zootaxa 5507 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Huarpea rotundicornis Lopes, Moleiros &amp; Lucena 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="445" masterDocId="FC436773FFD91F5FFFD2FF91967DFF89" masterDocTitle="A new species of Huarpea Pate, 1947 (Hymenoptera, Sapygidae) from the semiarid region of Brazil" masterLastPageNumber="455" masterPageNumber="439" pageNumber="441" updateTime="1726222335780" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="767D07580B6A0750A9A1423ABF692589">A new species of Huarpea Pate, 1947 (Hymenoptera, Sapygidae) from the semiarid region of Brazil</mods:title>
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<mods:namePart id="936669F90B9945E7CDD7D64DAA51108F">Lopes, Rogério Botion</mods:namePart>
<mods:affiliation id="A12E9BB3C5395D1C89CC8A1A003A32EC">Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Ribeirão Preto, São Paulo, Brazil</mods:affiliation>
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<mods:namePart id="3752D85C92D969B3382C6C2DD79B47FE">Moleiro, Henrique Pereira</mods:namePart>
<mods:affiliation id="EE0A4C5C6E3771EE096A0E05E2AE0388">Universidade de São Paulo (USP), Museu de Zoologia (MZUSP), São Paulo, SP, Brazil</mods:affiliation>
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<mods:namePart id="6D0AD675192FB9EC6B188D677C29BF03">Lucena, Daercio A. A.</mods:namePart>
<mods:affiliation id="EF34E84D62358DE24B70F3971FED71DA">Departamento de Ciências Biológicas, Instituto de Biociências, Letras e Ciências Exatas de São José do Rio Preto, Universidade Estadual Paulista “ Júlio de Mesquita Filho ” (UNESP), São José do Rio Preto, São Paulo, Brazil</mods:affiliation>
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<mods:date id="D17D2F5F9D8C7CACC9DD576D0195E016">2024</mods:date>
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<treatment id="007A1F0BFFDB1F59FF45FA5A929DF867" ID-DOI="http://doi.org/10.5281/zenodo.13757866" ID-Zenodo-Dep="13757866" LSID="urn:lsid:plazi:treatment:007A1F0BFFDB1F59FF45FA5A929DF867" httpUri="http://treatment.plazi.org/id/007A1F0BFFDB1F59FF45FA5A929DF867" lastPageId="6" lastPageNumber="445" pageId="2" pageNumber="441">
<subSubSection id="C0C9FD96FFDB1F5DFF45FA5A951EFA6C" box="[151,867,1482,1509]" pageId="2" pageNumber="441" type="nomenclature">
<paragraph id="886CAE1DFFDB1F5DFF45FA5A951EFA6C" blockId="2.[151,867,1482,1545]" box="[151,867,1482,1509]" pageId="2" pageNumber="441">
<heading id="D3241971FFDB1F5DFF45FA5A951EFA6C" bold="true" box="[151,867,1482,1509]" fontSize="11" level="1" pageId="2" pageNumber="441" reason="1">
<emphasis id="BAA7720FFFDB1F5DFF45FA5A951EFA6C" bold="true" box="[151,867,1482,1509]" pageId="2" pageNumber="441">
<taxonomicName id="4FD3D59EFFDB1F5DFF45FA5A9484FA6C" authority="Lopes, Moleiro &amp; Lucena" authorityName="Lopes, Moleiros &amp; Lucena" authorityYear="2024" box="[151,761,1482,1509]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="441" phylum="Tracheophyta" rank="species" species="rotundicornis" status="sp. nov.">
<emphasis id="BAA7720FFFDB1F5DFF45FA5A97D2FA6C" bold="true" box="[151,431,1482,1509]" italics="true" pageId="2" pageNumber="441">Huarpea rotundicornis</emphasis>
Lopes, Moleiro &amp; Lucena
</taxonomicName>
,
<taxonomicNameLabel id="A194CF74FFDB1F5DFCD7FA5A951EFA6C" box="[773,867,1483,1509]" pageId="2" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C0C9FD96FFDB1F5DFF45FA7E97BCF980" box="[151,449,1518,1545]" pageId="2" pageNumber="441" type="description">
<paragraph id="886CAE1DFFDB1F5DFF45FA7E97BCF980" blockId="2.[151,867,1482,1545]" box="[151,449,1518,1545]" pageId="2" pageNumber="441">
(
<figureCitation id="10E8B298FFDB1F5DFF4DFA7E976CF980" box="[159,273,1519,1545]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–10" captionStart-2="FIGURES 11–14" captionStartId-0="3.[151,264,1609,1633]" captionStartId-1="4.[151,264,796,820]" captionStartId-2="5.[151,264,1019,1043]" captionTargetBox-0="[151,1436,179,1584]" captionTargetBox-1="[151,1435,417,772]" captionTargetBox-2="[151,1427,597,996]" captionTargetId-0="figure-19@3.[151,1436,179,1584]" captionTargetId-1="figure-132@4.[151,1435,417,772]" captionTargetId-2="figure-219@5.[151,1436,597,996]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1–6. Huarpea rotundicornis sp. nov. and diagnostic characters for Sapyginae (1, 2, 5. Male, holotype. 3, 4, 6. Female, paratype). 1, 2. Habitus, lateral view. 2, 4. Head, frontal view. 5. Habitus, dorsal view. 6. Legs, arrow indicates rounded projection of hind coxa with carina. Scales: 1.0 mm." captionText-1="FIGURES 7–10. Diagnostic characters of Huarpea rotundicornis sp. nov. (7. Male, holotype. 8–10. Female, paratype). 7. Lateral view of flagellum. 8. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt). 9. Head, oblique view; triangles point frontal carina. 10. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm." captionText-2="FIGURES 11–14. Male genitalia of Huarpea rotundicornis sp. nov., holotype. 11. Genital capsule. 12. Aedeagus, ventral view. 13. Aedeagus, lateral view. 14. Paramere and associated structures. Scales: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13757870" figureDoi-1="http://doi.org/10.5281/zenodo.13757874" figureDoi-2="http://doi.org/10.5281/zenodo.13757876" httpUri-0="https://zenodo.org/record/13757870/files/figure.png" httpUri-1="https://zenodo.org/record/13757874/files/figure.png" httpUri-2="https://zenodo.org/record/13757876/files/figure.png" pageId="2" pageNumber="441">Figs 1–14</figureCitation>
,
<figureCitation id="10E8B298FFDB1F5DFECCFA7F9741F980" box="[286,316,1518,1545]" captionStart="FIGURE 25" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[174,1414,181,1768]" captionTargetId="figure-19@7.[174,1414,181,1768]" captionTargetPageId="7" captionText="FIGURE 25. Map showing known records of South American Sapygidae species." figureDoi="http://doi.org/10.5281/zenodo.13757880" httpUri="https://zenodo.org/record/13757880/files/figure.png" pageId="2" pageNumber="441">25</figureCitation>
,
<figureCitation id="10E8B298FFDB1F5DFE9AFA7E971BF980" box="[328,358,1519,1545]" captionStart="FIGURES 26–29" captionStartId="8.[151,264,1899,1923]" captionTargetBox="[151,1436,1550,1876]" captionTargetId="figure-1049@8.[151,1436,1550,1876]" captionTargetPageId="8" captionText="FIGURES 26–29. Female head, oblique view. 26. Huarpea wagneriella. 27. H. fallax. 28. H. rotundicornis sp. nov., paratype. 29. Sapyga quinquepunctata. Arrows indicate occipital carina. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13757882" httpUri="https://zenodo.org/record/13757882/files/figure.png" pageId="2" pageNumber="441">28</figureCitation>
,
<figureCitation id="10E8B298FFDB1F5DFEA0FA7E97C4F980" box="[370,441,1519,1545]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1234,1258]" captionTargetBox="[151,1436,181,1211]" captionTargetId="figure-19@10.[151,1436,181,1211]" captionTargetPageId="10" captionText="FIGURES 30–33. Characters of Huarpea rotundicornis sp. nov. that contrast with other sapygid genera. 30. Male flagellum, ventral view, holotype. 31. Mid tarsus, dorsal view. 32. S6 of female, paratype. 33. Fore wing, holotype. Scales: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.13757884" httpUri="https://zenodo.org/record/13757884/files/figure.png" pageId="2" pageNumber="441">30–33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0C9FD96FFDB1F5DFF45F9A7941AF81C" pageId="2" pageNumber="441" type="diagnosis">
<paragraph id="886CAE1DFFDB1F5DFF45F9A7934CF910" blockId="2.[151,1437,1590,1941]" pageId="2" pageNumber="441">
<emphasis id="BAA7720FFFDB1F5DFF45F9A79777F9D9" bold="true" box="[151,266,1590,1616]" pageId="2" pageNumber="441">Diagnosis</emphasis>
: The round last male flagellomere, which is also wider than long, is unique in the genus (
<figureCitation id="10E8B298FFDB1F5DFB30F9A69358F9D8" box="[1250,1317,1591,1617]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,796,820]" captionTargetBox="[151,1435,417,772]" captionTargetId="figure-132@4.[151,1435,417,772]" captionTargetPageId="4" captionText="FIGURES 7–10. Diagnostic characters of Huarpea rotundicornis sp. nov. (7. Male, holotype. 8–10. Female, paratype). 7. Lateral view of flagellum. 8. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt). 9. Head, oblique view; triangles point frontal carina. 10. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13757874" httpUri="https://zenodo.org/record/13757874/files/figure.png" pageId="2" pageNumber="441">Fig. 7</figureCitation>
). Also, the metapleura tightly constricted by the propodeum (
<figureCitation id="10E8B298FFDB1F5DFD1FF9CA956CF9FC" box="[717,785,1627,1653]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,796,820]" captionTargetBox="[151,1435,417,772]" captionTargetId="figure-132@4.[151,1435,417,772]" captionTargetPageId="4" captionText="FIGURES 7–10. Diagnostic characters of Huarpea rotundicornis sp. nov. (7. Male, holotype. 8–10. Female, paratype). 7. Lateral view of flagellum. 8. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt). 9. Head, oblique view; triangles point frontal carina. 10. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13757874" httpUri="https://zenodo.org/record/13757874/files/figure.png" pageId="2" pageNumber="441">Fig. 8</figureCitation>
), strong frontal carina present (
<figureCitation id="10E8B298FFDB1F5DFBA4F9CA92C7F9FC" box="[1142,1210,1627,1653]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,796,820]" captionTargetBox="[151,1435,417,772]" captionTargetId="figure-132@4.[151,1435,417,772]" captionTargetPageId="4" captionText="FIGURES 7–10. Diagnostic characters of Huarpea rotundicornis sp. nov. (7. Male, holotype. 8–10. Female, paratype). 7. Lateral view of flagellum. 8. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt). 9. Head, oblique view; triangles point frontal carina. 10. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13757874" httpUri="https://zenodo.org/record/13757874/files/figure.png" pageId="2" pageNumber="441">Fig. 9</figureCitation>
) and abrupt smooth depression above antennal socket contrasting with surroundings (
<figureCitation id="10E8B298FFDB1F5DFCA0F9EE95BEF910" box="[882,963,1663,1689]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,796,820]" captionTargetBox="[151,1435,417,772]" captionTargetId="figure-132@4.[151,1435,417,772]" captionTargetPageId="4" captionText="FIGURES 7–10. Diagnostic characters of Huarpea rotundicornis sp. nov. (7. Male, holotype. 8–10. Female, paratype). 7. Lateral view of flagellum. 8. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt). 9. Head, oblique view; triangles point frontal carina. 10. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13757874" httpUri="https://zenodo.org/record/13757874/files/figure.png" pageId="2" pageNumber="441">Fig. 10</figureCitation>
) readily distinguish this species.
</paragraph>
<paragraph id="886CAE1DFFDB1F5DFF15F933941AF81C" blockId="2.[151,1437,1590,1941]" pageId="2" pageNumber="441">
<emphasis id="BAA7720FFFDB1F5DFF15F93397CCF935" bold="true" box="[199,433,1698,1724]" pageId="2" pageNumber="441">Examined material.</emphasis>
<materialsCitation id="38BBA440FFDB1F5DFE67F932941AF81C" collectingDate="2022-11-09" collectingDateMax="2023-05-05" collectingDateMin="2022-11-09" collectionCode="S, USP, RPSP, BA, I, H, P, MZSP" collectorName="Lucena &amp; Utinga &amp; Bairro Barbado &amp; Area &amp; Chapada Diamantina &amp; Malaise &amp; Moleiro &amp; Bairro Cajueiro &amp; Malaise II" country="Ecuador" elevation="324" latitude="-11.945917" longLatPrecision="1" longitude="-41.049" pageId="2" pageNumber="441" specimenCode="RPSP 00013831, MZSP 0124755, MZSP 0124754" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<typeStatus id="576810BFFFDB1F5DFE67F9329431F934" box="[437,588,1699,1725]" pageId="2" pageNumber="441" type="holotype">HOLOTYPE</typeStatus>
,
<specimenCount id="9ED56594FFDB1F5DFD86F9329416F934" box="[596,619,1699,1725]" count="1" pageId="2" pageNumber="441" type="male">♂</specimenCount>
, “Brasil.RN [
<collectingRegion id="4A1760FFFFDB1F5DFCD3F9329590F934" box="[769,1005,1699,1725]" country="Brazil" name="Rio Grande do Norte" pageId="2" pageNumber="441">Rio Grande do Norte</collectingRegion>
],
<collectingCountry id="F0C4EE8DFFDB1F5DFC2EF9329226F934" box="[1020,1115,1699,1725]" name="Ecuador" pageId="2" pageNumber="441">Equador</collectingCountry>
\
<collectionCode id="EEC236D8FFDB1F5DFBBEF9329201F934" box="[1132,1148,1699,1725]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="441" type="Herbarium">S</collectionCode>
Malhada Grande. Malaise 10 \
<geoCoordinate id="EDE7C8DAFFDB1F5DFF14F957972EF968" box="[198,339,1734,1761]" degrees="06" direction="south" minutes="51.359" orientation="latitude" pageId="2" pageNumber="441" precision="1" value="-6.8559833">06°51.359’S</geoCoordinate>
<geoCoordinate id="EDE7C8DAFFDB1F5DFE8AF9569783F968" box="[344,510,1735,1761]" degrees="036" direction="west" minutes="36.980" orientation="longitude" pageId="2" pageNumber="441" precision="1" value="-36.616333">036°36.980’W</geoCoordinate>
\
<date id="FC6D88DDFFDB1F5DFDC2F95794BBF968" box="[528,710,1734,1761]" pageId="2" pageNumber="441" value="2023-04-17" valueMax="2023-05-05" valueMin="2023-04-17">
<collectingDate id="EC297135FFDB1F5DFDC2F95794BBF968" box="[528,710,1734,1761]" pageId="2" pageNumber="441" value="2023-04-17" valueMax="2023-05-05" valueMin="2023-04-17">17.iv–05.v.2023</collectingDate>
</date>
<quantity id="4F2B03F8FFDB1F5DFD19F9569576F968" box="[715,779,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="3.2399999999999998" pageId="2" pageNumber="441" unit="m" value="324.0">
<elevation id="03FE492EFFDB1F5DFD19F9569576F968" box="[715,779,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="3.2399999999999998" pageId="2" pageNumber="441" unit="m" value="324.0">324m</elevation>
</quantity>
\ DAA
<collectorName id="2526CBCBFFDB1F5DFC8CF95695CDF968" box="[862,944,1735,1761]" pageId="2" pageNumber="441">Lucena</collectorName>
&amp; eq cols \\
<collectionCode id="EEC236D8FFDB1F5DFBEAF9569217F968" box="[1080,1130,1735,1761]" country="Fiji" httpUri="http://grbio.org/cool/hb24-jpsb" name="University of the South Pacific" pageId="2" pageNumber="441">USP</collectionCode>
_
<specimenCode id="D8750666FFDB1F5DFBABF956934FF968" box="[1145,1330,1735,1761]" collectionCode="RPSP" country="Brazil" httpUri="http://grbio.org/cool/745q-57sa" name="Universidade de Sao Paulo" pageId="2" pageNumber="441">RPSP 00013831</specimenCode>
” [
<collectionCode id="EEC236D8FFDB1F5DFA9AF95693EFF968" box="[1352,1426,1735,1761]" country="Brazil" httpUri="http://grbio.org/cool/745q-57sa" name="Universidade de Sao Paulo" pageId="2" pageNumber="441">RPSP</collectionCode>
].
<typeStatus id="576810BFFFDB1F5DFF45F97A9748F88C" box="[151,309,1771,1797]" pageId="2" pageNumber="441" type="paratype">PARATYPES</typeStatus>
.
<specimenCount id="9ED56594FFDB1F5DFE93F97A9719F88C" box="[321,356,1771,1797]" count="1" pageId="2" pageNumber="441" type="female">1♀</specimenCount>
, same data as holotype \\ “DZSJRP-Hym. 9044” [DZSJRP-Hymenoptera];
<specimenCount id="9ED56594FFDB1F5DFB16F97A929AF88C" box="[1220,1255,1771,1797]" count="1" pageId="2" pageNumber="441" type="female">1♀</specimenCount>
, “
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;
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<collectorName id="2526CBCBFFDB1F5DFF2EF89E9735F8A0" box="[252,328,1807,1833]" pageId="2" pageNumber="441">Utinga</collectorName>
(
<collectorName id="2526CBCBFFDB1F5DFE8BF89E9470F8A0" box="[345,525,1807,1833]" pageId="2" pageNumber="441">Bairro Barbado</collectorName>
) \
<collectorName id="2526CBCBFFDB1F5DFDFCF8819419F8A2" box="[558,612,1808,1835]" pageId="2" pageNumber="441">Área</collectorName>
de caatinga—
<collectorName id="2526CBCBFFDB1F5DFCD8F89E9584F8A0" box="[778,1017,1807,1833]" pageId="2" pageNumber="441">Chapada Diamantina</collectorName>
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</date>
<collectorName id="2526CBCBFFDB1F5DFEC5F8A2970DF8C4" box="[279,368,1843,1869]" pageId="2" pageNumber="441">Malaise</collectorName>
<collectionCode id="EEC236D8FFDB1F5DFEABF8A297FFF8C4" box="[377,386,1843,1869]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="2" pageNumber="441" type="Herbarium">I</collectionCode>
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<collectionCode id="EEC236D8FFDB1F5DFE67F8A297BBF8C4" box="[437,454,1843,1869]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="441" type="Herbarium">P</collectionCode>
.
<collectorName id="2526CBCBFFDB1F5DFE00F8A29451F8C4" box="[466,556,1843,1869]" pageId="2" pageNumber="441">Moleiro</collectorName>
col. \\
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, “
<collectingCountry id="F0C4EE8DFFDB1F5DFC3CF8A2922AF8C4" box="[1006,1111,1843,1869]" name="Brazil" pageId="2" pageNumber="441">BRASIL</collectingCountry>
;
<collectionCode id="EEC236D8FFDB1F5DFBB7F8A292F7F8C4" box="[1125,1162,1843,1869]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="2" pageNumber="441" type="Herbarium">BA</collectionCode>
[
<collectingRegion id="4A1760FFFFDB1F5DFB4BF8A2929EF8C4" box="[1177,1251,1843,1869]" country="Brazil" name="Bahia" pageId="2" pageNumber="441">Bahia</collectingRegion>
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<collectorName id="2526CBCBFFDB1F5DFB25F8A2933EF8C4" box="[1271,1347,1843,1869]" pageId="2" pageNumber="441">Utinga</collectorName>
(
<collectorName id="2526CBCBFFDB1F5DFA81F8A29684F8F8" pageId="2" pageNumber="441">Bairro Cajueiro</collectorName>
) \
<collectorName id="2526CBCBFFDB1F5DFEC7F8C99736F8FA" box="[277,331,1880,1907]" pageId="2" pageNumber="441">Área</collectorName>
de caatinga—
<collectorName id="2526CBCBFFDB1F5DFE39F8C694ABF8F8" box="[491,726,1879,1905]" pageId="2" pageNumber="441">Chapada Diamantina</collectorName>
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<geoCoordinate id="EDE7C8DAFFDB1F5DFD38F8C795FFF8F8" box="[746,898,1878,1905]" degrees="11" direction="south" minutes="56" orientation="latitude" pageId="2" pageNumber="441" precision="1" seconds="45.3" value="-11.945917">11°56’45.3”S</geoCoordinate>
<geoCoordinate id="EDE7C8DAFFDB1F5DFC5AF8C79251F8F8" box="[904,1068,1878,1905]" degrees="41" direction="west" minutes="02" orientation="longitude" pageId="2" pageNumber="441" precision="1" seconds="56.4" value="-41.049">41°02’56.4”W</geoCoordinate>
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<collectorName id="2526CBCBFFDB1F5DFB3AF8C69327F8F8" box="[1256,1370,1879,1905]" pageId="2" pageNumber="441">Malaise II</collectorName>
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.
<collectionCode id="EEC236D8FFDB1F5DFA55F8C693E5F8F8" box="[1415,1432,1879,1905]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="441" type="Herbarium">P</collectionCode>
.
<collectorName id="2526CBCBFFDB1F5DFF45F8EA968CF81C" box="[151,241,1915,1941]" pageId="2" pageNumber="441">Moleiro</collectorName>
col. \\
<specimenCode id="D8750666FFDB1F5DFEEDF8EA978EF81C" box="[319,499,1914,1941]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="441" type="Museum">MZSP 0124754</specimenCode>
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<collectionCode id="EEC236D8FFDB1F5DFDD8F8EA9421F81C" box="[522,604,1915,1941]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="441" type="Museum">MZSP</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DCACFE95FFDA1F5CFF45F9D894D7F920" ID-DOI="http://doi.org/10.5281/zenodo.13757870" ID-Zenodo-Dep="13757870" httpUri="https://zenodo.org/record/13757870/files/figure.png" pageId="3" pageNumber="442" startId="3.[151,264,1609,1633]" targetBox="[151,1436,179,1584]" targetPageId="3" targetType="figure">
<paragraph id="886CAE1DFFDA1F5CFF45F9D894D7F920" blockId="3.[151,1436,1609,1706]" pageId="3" pageNumber="442">
<emphasis id="BAA7720FFFDA1F5CFF45F9D89745F9EB" bold="true" box="[151,312,1609,1634]" pageId="3" pageNumber="442">FIGURES 1–6</emphasis>
.
<taxonomicName id="4FD3D59EFFDA1F5CFE95F9DB9449F9EB" authorityName="Lopes &amp; Moleiro &amp; Lucena" authorityYear="2024" box="[327,564,1610,1634]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="442" phylum="Tracheophyta" rank="species" species="rotundicornis" status="sp. nov.">
<emphasis id="BAA7720FFFDA1F5CFE95F9DB9449F9EB" box="[327,564,1610,1634]" italics="true" pageId="3" pageNumber="442">Huarpea rotundicornis</emphasis>
</taxonomicName>
<emphasis id="BAA7720FFFDA1F5CFDEFF9DB94ECF9EB" bold="true" box="[573,657,1610,1634]" pageId="3" pageNumber="442">
<taxonomicNameLabel id="A194CF74FFDA1F5CFDEFF9DB94ECF9EB" box="[573,657,1610,1634]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and diagnostic characters for
<taxonomicName id="4FD3D59EFFDA1F5CFC00F9DB9241F9EB" authorityName="Latreille" authorityYear="1810" box="[978,1084,1610,1634]" class="Insecta" family="Sapygidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="subFamily" subFamily="Sapyginae">Sapyginae</taxonomicName>
(1, 2, 5. Male, holotype. 3, 4, 6. Female, paratype). 1, 2. Habitus, lateral view.
<emphasis id="BAA7720FFFDA1F5CFDB5F9FF9409F90F" bold="true" box="[615,628,1646,1670]" pageId="3" pageNumber="442">2</emphasis>
,
<emphasis id="BAA7720FFFDA1F5CFD52F9FF94F0F90F" bold="true" box="[640,653,1646,1670]" pageId="3" pageNumber="442">4</emphasis>
. Head, frontal view. 5. Habitus, dorsal view.
<emphasis id="BAA7720FFFDA1F5CFB80F9FF9222F90F" bold="true" box="[1106,1119,1646,1670]" pageId="3" pageNumber="442">6</emphasis>
. Legs, arrow indicates rounded projection of hind coxa with carina. Scales: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C0C9FD96FFDA1F5AFF15F96D95D4F9E2" lastPageId="5" lastPageNumber="444" pageId="3" pageNumber="442" type="description">
<paragraph id="886CAE1DFFDA1F5BFF15F96D97AAFE00" blockId="3.[151,1436,1788,2030]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442">
<emphasis id="BAA7720FFFDA1F5CFF15F96D972CF89F" bold="true" box="[199,337,1788,1814]" pageId="3" pageNumber="442">Description</emphasis>
. MALE.
<emphasis id="BAA7720FFFDA1F5CFE6FF96D9443F89F" bold="true" box="[445,574,1788,1814]" pageId="3" pageNumber="442">Coloration</emphasis>
. Predominantly black, with the following creamy yellow markings along the body: clypeus entirely marked except medial diamond-shaped black spot; large spot on mandible; ventral surface of scape; ventral surface of F-I to III, markings evanescent on following flagellomeres; triangular mark with wide base on interantennal region narrowing until tip that ends well before anterior ocellus; wide band along inner margin of eye, including entire sinus; wide genal band along outer eye margin, extending a little past upper lobe; pair of spots on vertex, postero-lateral to posterior ocelli; anterior band along dorsal surface of pronotum extending down to lateral surface; band along posterior margin of pronotum, followed by a thin translucent band; lateral margins of mesoscutum; tegula; axillae; anterolateral spot on scutellum; metanotum; scrobal spot; longitudinal stripe on anterior surface of fore coxae; ventral surface of fore femur; fore tibia except posterior surface; fore tarsus; anterior face of mid and hind coxae; apical longitudinal stripe on mid femur; longitudinal stripe on dorsal surface of hind tibia; articulations on legs; propodeum with large spot on the lateral face and a pair or small spots on the posterior face; wide apical band on T1 submedially interrupted; wide band on T2; bands of T4–5 well defined, not evanescent, but medially interrupted; apical half of T6; S2–6 with irregular evanescent bands. T7 and S7 creamy yellow with wide apical testaceous band.
</paragraph>
<caption id="DCACFE95FFDD1F5BFF45FC8D936FFCF5" ID-DOI="http://doi.org/10.5281/zenodo.13757874" ID-Zenodo-Dep="13757874" httpUri="https://zenodo.org/record/13757874/files/figure.png" pageId="4" pageNumber="443" startId="4.[151,264,796,820]" targetBox="[151,1435,417,772]" targetPageId="4" targetType="figure">
<paragraph id="886CAE1DFFDD1F5BFF45FC8D936FFCF5" blockId="4.[151,1436,796,892]" pageId="4" pageNumber="443">
<emphasis id="BAA7720FFFDD1F5BFF45FC8D9739FCBD" bold="true" box="[151,324,796,820]" pageId="4" pageNumber="443">FIGURES 7–10</emphasis>
. Diagnostic characters of
<taxonomicName id="4FD3D59EFFDD1F5BFD8BFC8C953BFCBD" authorityName="Lopes &amp; Moleiro &amp; Lucena" authorityYear="2024" box="[601,838,796,820]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="443" phylum="Tracheophyta" rank="species" species="rotundicornis" status="sp. nov.">
<emphasis id="BAA7720FFFDD1F5BFD8BFC8C953BFCBD" box="[601,838,796,820]" italics="true" pageId="4" pageNumber="443">Huarpea rotundicornis</emphasis>
</taxonomicName>
<emphasis id="BAA7720FFFDD1F5BFC9DFC8D95DFFCBD" bold="true" box="[847,930,796,820]" pageId="4" pageNumber="443">
<taxonomicNameLabel id="A194CF74FFDD1F5BFC9DFC8D95DFFCBD" box="[847,930,796,820]" pageId="4" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(7. Male, holotype. 8–10. Female, paratype).
<emphasis id="BAA7720FFFDD1F5BFA5BFC8D93EBFCBD" bold="true" box="[1417,1430,796,820]" pageId="4" pageNumber="443">7</emphasis>
. Lateral view of flagellum.
<emphasis id="BAA7720FFFDD1F5BFE71FCD197CDFCD1" bold="true" box="[419,432,832,856]" pageId="4" pageNumber="443">8</emphasis>
. Lateral view of mesosoma; arrow indicates constriction of metapleura (mt).
<emphasis id="BAA7720FFFDD1F5BFB6AFCD192B8FCD1" bold="true" box="[1208,1221,832,856]" pageId="4" pageNumber="443">9</emphasis>
. Head, oblique view; triangles point frontal carina.
<emphasis id="BAA7720FFFDD1F5BFE16FCF597A0FCF5" bold="true" box="[452,477,868,892]" pageId="4" pageNumber="443">10</emphasis>
. Head, oblique view; arrow indicates depression above antenna. Scales: 1.0 mm.
</paragraph>
</caption>
<paragraph id="886CAE1DFFDD1F5BFF15FC7797F5FA38" blockId="4.[151,1437,998,1889]" pageId="4" pageNumber="443">
<emphasis id="BAA7720FFFDD1F5BFF15FC779747FB89" bold="true" box="[199,314,998,1024]" pageId="4" pageNumber="443">Structure</emphasis>
. Clypeus 2.38× wider than high; not medially depressed apically, with an evident concave apical margin between two blunt teeth. Flagellum fusiform, widest at F-VII to IX, and short, reaching at most posterior margin of scutum. F-XI 1.75× longer than wide, reniform, ventrally curved. Vertex without tubercles. Ocelli size not reduced. Eyes with inner orbit deeply margined, outer margin regularly convex and upper lobe smaller than lower lobe. Mandible 3-toothed, with basal triangle strongly separated from apex by a perpendicular ridge. Pronotum with anterior and posterior margins slightly concave in dorsal view, with dorsal surface slightly convex gradually transitioning to lateral surface. Scutum trapezoid, wider anteriorly. Axillae triangular. Scutellum nearly rectangular, with strong beveled antero-lateral edges. Mesopleura nearly horizontal, easily separated in three surfaces: anterior (epicnemium), lateral and posterior, the latter two divided by steep sloping of posterior surface. Metanotum as wide as scutellum, but about only half as long. Metapleura strongly constricted by propodeum about half-height. Metacoxa with rounded projection. Propodeum regularly convex on declivity. T1 subconical, with anterior face slightly concave and apex with a light constriction. Metasoma fusiform. S7 only convex, with bilobed apex. Fore wing length:
<quantity id="4F2B03F8FFDD1F5BFEF9FA0697F9FA38" box="[299,388,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="443" unit="mm" value="4.9">4.9 mm</quantity>
.
</paragraph>
<paragraph id="886CAE1DFFDD1F5BFF15FA2B9502F8E9" blockId="4.[151,1437,998,1889]" pageId="4" pageNumber="443">
<emphasis id="BAA7720FFFDD1F5BFF15FA2B9746FA5D" bold="true" box="[199,315,1466,1492]" pageId="4" pageNumber="443">Sculpture</emphasis>
. Cuticle shiny. Clypeus with moderate punctation, with interspaces around one puncture diameter, and irregular longitudinal striae. F-IX to V with longitudinal calli, with the apical most on ventral surface and each preceding one progressively more deviated to the lateral surface. Frons with very dense punctation presenting cariniform interspaces. Punctation dense, interspaces with less than one puncture diameter presenting smaller punctures, on vertex, pronotum, mesoscutum and scutellum. Mesopleura with moderate punctation, intercalated by smaller punctures, dorsally becoming gradually sparser antero-ventrally where only the smaller punctures are seen. Posterior margin of scutellum and metanotum with dense punctation intercalated by smooth areas. Metapleura with very dense small punctures. Propodeum with irregularly distributed large, shallow punctures posteriorly and laterally with large shiny smooth area. T1 with moderate to sparse punctation intercalated by smaller punctures, smooth on median concavity. S1 with a medial band of dense small punctures, laterally smooth. T3–4 and S3–4 with dense small punctures and sparse large punctures, sparser on T4 and S4. T5 and S5 and following segments with transverse micro shagreening. S6 without median carina or sulcus.
</paragraph>
<paragraph id="886CAE1DFFDC1F5AFF15FF079263FE94" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="444">
<emphasis id="BAA7720FFFDC1F5AFF15FF07975DFF39" bold="true" box="[199,288,150,176]" pageId="5" pageNumber="444">Pilosity</emphasis>
. Dense erect short thin silvery setae covering most of the body. Two very long dark setae on dorsal margin of left eye, three on the right. Very long and sparse setae on hypostoma and occiput ventrally. Longer on proepisternum. Very dense and thick on forecoxae. Long and thin on propodeum T1 and S1. Tomentum on metasoma. Golden pilosity on apex of T3 and S3 and entirety of T4–7 and S4–7.
</paragraph>
<paragraph id="886CAE1DFFDC1F5AFF15FEB7928AFDB4" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="444">
<emphasis id="BAA7720FFFDC1F5AFF15FEB79748FEC9" bold="true" box="[199,309,294,320]" pageId="5" pageNumber="444">Genitalia</emphasis>
(
<figureCitation id="10E8B298FFDC1F5AFE97FEB697BAFEC8" box="[325,455,295,321]" captionStart="FIGURES 11–14" captionStartId="5.[151,264,1019,1043]" captionTargetBox="[151,1427,597,996]" captionTargetId="figure-219@5.[151,1436,597,996]" captionTargetPageId="5" captionText="FIGURES 11–14. Male genitalia of Huarpea rotundicornis sp. nov., holotype. 11. Genital capsule. 12. Aedeagus, ventral view. 13. Aedeagus, lateral view. 14. Paramere and associated structures. Scales: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757876" httpUri="https://zenodo.org/record/13757876/files/figure.png" pageId="5" pageNumber="444">Figs 11–14</figureCitation>
). Genital capsule with a well-developed basal ring, basally narrower shaped like a shirt collar. Aedeagus completely bipartite, not fused medially in any section; ventral projection absent; apodemes with light dorsally oriented curvature and gradually narrowing towards apex with subapical ventrally oriented reflection; medial expansion strongly projected as a long triangle with anterior side curved and posterior side straight; anterior portion very large and wide with a convex dorsal margin and concave ventral margin; external face of anterior section covered with sparse short thick setae; penis valve thickened with spines on external surface. Paramere strongly convex with a falciform ventral projection. Parameral spine wide and flat, covered in thin setae. Digitus large and sac-like, covered by dense thin pilosity. Cuspis digitiform, long and slender, and glabrous.
</paragraph>
<caption id="DCACFE95FFDC1F5AFF45FC6A959BFBBE" ID-DOI="http://doi.org/10.5281/zenodo.13757876" ID-Zenodo-Dep="13757876" httpUri="https://zenodo.org/record/13757876/files/figure.png" pageId="5" pageNumber="444" startId="5.[151,264,1019,1043]" targetBox="[151,1427,597,996]" targetPageId="5" targetType="figure">
<paragraph id="886CAE1DFFDC1F5AFF45FC6A959BFBBE" blockId="5.[151,1437,1019,1079]" pageId="5" pageNumber="444">
<emphasis id="BAA7720FFFDC1F5AFF45FC6A9731FB9A" bold="true" box="[151,332,1019,1043]" pageId="5" pageNumber="444">FIGURES 11–14</emphasis>
. Male genitalia of
<taxonomicName id="4FD3D59EFFDC1F5AFDDBFC6D948FFB9A" authorityName="Lopes &amp; Moleiro &amp; Lucena" authorityYear="2024" box="[521,754,1019,1043]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="444" phylum="Tracheophyta" rank="species" species="rotundicornis" status="sp. nov.">
<emphasis id="BAA7720FFFDC1F5AFDDBFC6D948FFB9A" box="[521,754,1019,1043]" italics="true" pageId="5" pageNumber="444">Huarpea rotundicornis</emphasis>
</taxonomicName>
<emphasis id="BAA7720FFFDC1F5AFD2AFC6A9535FB9A" bold="true" box="[760,840,1019,1043]" pageId="5" pageNumber="444">
<taxonomicNameLabel id="A194CF74FFDC1F5AFD2AFC6A9535FB9A" box="[760,840,1019,1043]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="BAA7720FFFDC1F5AFC68FC6A95AFFB9A" bold="true" box="[954,978,1019,1043]" pageId="5" pageNumber="444">11</emphasis>
. Genital capsule.
<emphasis id="BAA7720FFFDC1F5AFB54FC6A92E2FB9A" bold="true" box="[1158,1183,1019,1043]" pageId="5" pageNumber="444">12</emphasis>
. Aedeagus, ventral view.
<emphasis id="BAA7720FFFDC1F5AFF45FB8E96CDFBBE" bold="true" box="[151,176,1055,1079]" pageId="5" pageNumber="444">13</emphasis>
. Aedeagus, lateral view.
<emphasis id="BAA7720FFFDC1F5AFE7CFB8E97BAFBBE" bold="true" box="[430,455,1055,1079]" pageId="5" pageNumber="444">14</emphasis>
. Paramere and associated structures. Scales: 0.5 mm.
</paragraph>
</caption>
<paragraph id="886CAE1DFFDC1F5AFF15FB3395D4F9E2" blockId="5.[151,1437,1185,1644]" pageId="5" pageNumber="444">
FEMALE. As described for male, except as listed below.
<emphasis id="BAA7720FFFDC1F5AFCB1FB309599FB32" bold="true" box="[867,996,1185,1211]" pageId="5" pageNumber="444">Coloration</emphasis>
. Creamy yellow markings as follows: lateral third of clypeus frontal marking narrower inferiorly, superiorly reaching anterior ocellus; vertex without spots; Fore coxae with only an anterior line; T3 with a pair of very small subapical sublateral spots; Mandible and F-II to III not marked. T4–5 with irregular partially evanescent apical bands. Parts with testaceous markings instead of yellow: basal half of F-I; tegula; ventral surface of fore femur; fore tibia except posterior surface; fore tarsus; apical longitudinal stripe on mid femur; articulations on legs.
<emphasis id="BAA7720FFFDC1F5AFC90FAC495C8FAE6" bold="true" box="[834,949,1365,1391]" pageId="5" pageNumber="444">Structure</emphasis>
. Clypeus 2.08× wider than high; flat, with a small medial apical depression between broadly blunt apical teeth. F-X conical with rounded tip and slightly curved inwards. S6 conical.
<emphasis id="BAA7720FFFDC1F5AFE55FA0C9786FA3E" bold="true" box="[391,507,1437,1463]" pageId="5" pageNumber="444">Sculpture</emphasis>
. Punctation overall not so dense and lighter. Fore wing length:
<quantity id="4F2B03F8FFDC1F5AFB07FA0C934DFA31" box="[1237,1328,1437,1464]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="444" unit="mm" value="5.4">5.4 mm</quantity>
.
<emphasis id="BAA7720FFFDC1F5AFAEFFA0C93EBFA3E" bold="true" box="[1341,1430,1437,1463]" pageId="5" pageNumber="444">Pilosity</emphasis>
. Metasoma without golden pilosity.
<emphasis id="BAA7720FFFDC1F5AFDE2FA5094D6FA52" bold="true" box="[560,683,1473,1499]" pageId="5" pageNumber="444">Variations</emphasis>
. More extensively colored females will present wider bands and larger spots and specimens may vary from testaceous to creamy yellow in the legs (perhaps due to preservation differences). One of the specimens from
<collectingRegion id="4A1760FFFFDC1F5AFDB1F99894D9F9AA" box="[611,676,1545,1571]" country="Brazil" name="Bahia" pageId="5" pageNumber="444">Bahia</collectingRegion>
presents a basal triangular spot on the mandible, a large transversal spot on the posterior surface of the propodeum instead of the pair of small spots and band of T1 submedially narrow instead of interrupted. In the same specimen, there are no spots on T3.
</paragraph>
</subSubSection>
<caption id="DCACFE95FFDF1F59FF45FAE19747F991" ID-DOI="http://doi.org/10.5281/zenodo.13757878" ID-Zenodo-Dep="13757878" httpUri="https://zenodo.org/record/13757878/files/figure.png" pageId="6" pageNumber="445" startId="6.[151,264,1392,1416]" targetBox="[151,1436,181,1368]" targetPageId="6" targetType="figure">
<paragraph id="886CAE1DFFDF1F59FF45FAE19747F991" blockId="6.[151,1437,1392,1561]" pageId="6" pageNumber="445">
<emphasis id="BAA7720FFFDF1F59FF45FAE19730FA00" bold="true" box="[151,333,1392,1417]" pageId="6" pageNumber="445">FIGURES 15–24</emphasis>
. Key characters for identification of
<taxonomicName id="4FD3D59EFFDF1F59FD13FAE3950CFA00" box="[705,881,1393,1417]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="445" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="BAA7720FFFDF1F59FD13FAE39566FA00" box="[705,795,1394,1417]" italics="true" pageId="6" pageNumber="445">Huarpea</emphasis>
species.
</taxonomicName>
<emphasis id="BAA7720FFFDF1F59FCA4FAE095CBFA00" bold="true" box="[886,950,1393,1417]" pageId="6" pageNumber="445">15–17</emphasis>
.
<emphasis id="BAA7720FFFDF1F59FC10FAE392DFFA00" box="[962,1186,1393,1417]" italics="true" pageId="6" pageNumber="445">
<taxonomicName id="4FD3D59EFFDF1F59FC10FAE392E3FA00" baseAuthorityName="du Buysson" baseAuthorityYear="1904" box="[962,1182,1393,1417]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="445" phylum="Tracheophyta" rank="species" species="wagneriella">Huarpea wagneriella</taxonomicName>
.
</emphasis>
<emphasis id="BAA7720FFFDF1F59FB75FAE0929AFA00" bold="true" box="[1191,1255,1393,1417]" pageId="6" pageNumber="445">18–24</emphasis>
.
<taxonomicName id="4FD3D59EFFDF1F59FB21FAE39336FA00" baseAuthorityName="Gerstacker" baseAuthorityYear="1861" box="[1267,1355,1393,1417]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="445" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="BAA7720FFFDF1F59FB21FAE39336FA00" box="[1267,1355,1393,1417]" italics="true" pageId="6" pageNumber="445">H. fallax</emphasis>
</taxonomicName>
.
<emphasis id="BAA7720FFFDF1F59FA85FAE0930DFA00" bold="true" box="[1367,1392,1393,1417]" pageId="6" pageNumber="445">15</emphasis>
,
<emphasis id="BAA7720FFFDF1F59FAAFFAE093EBFA00" bold="true" box="[1405,1430,1393,1417]" pageId="6" pageNumber="445">18</emphasis>
. Clypeus, oblique view.
<emphasis id="BAA7720FFFDF1F59FE5AFA0497DCFA24" bold="true" box="[392,417,1429,1453]" pageId="6" pageNumber="445">16</emphasis>
,
<emphasis id="BAA7720FFFDF1F59FE7CFA0497BAFA24" bold="true" box="[430,455,1429,1453]" pageId="6" pageNumber="445">19</emphasis>
. Close up of setae above eye; triangle points overhanging longer setae.
<emphasis id="BAA7720FFFDF1F59FB7AFA0492BCFA24" bold="true" box="[1192,1217,1429,1453]" pageId="6" pageNumber="445">17</emphasis>
,
<emphasis id="BAA7720FFFDF1F59FB1DFA049295FA24" bold="true" box="[1231,1256,1429,1453]" pageId="6" pageNumber="445">20</emphasis>
. T1, dorsal view; arrow indicates protruding spiracles.
<emphasis id="BAA7720FFFDF1F59FDC5FA28944DFA58" bold="true" box="[535,560,1465,1489]" pageId="6" pageNumber="445">21</emphasis>
. Mesosoma, lateral view showing metapleura (
<emphasis id="BAA7720FFFDF1F59FBCEFA2B9248FA58" box="[1052,1077,1466,1489]" italics="true" pageId="6" pageNumber="445">mt</emphasis>
) not constricted.
<emphasis id="BAA7720FFFDF1F59FB3FFA28937BFA58" bold="true" box="[1261,1286,1465,1489]" pageId="6" pageNumber="445">22</emphasis>
. Head, frontal view.
<emphasis id="BAA7720FFFDF1F59FF01FA4C9691FA7C" bold="true" box="[211,236,1501,1525]" pageId="6" pageNumber="445">23</emphasis>
. Head, oblique view; triangles point frontal carina.
<emphasis id="BAA7720FFFDF1F59FD2AFA4C956CFA7C" bold="true" box="[760,785,1501,1525]" pageId="6" pageNumber="445">24</emphasis>
. Head, oblique view; arrow indicates depression above antenna. Scales: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C0C9FD96FFDF1F59FF15F9FA95AAF82C" pageId="6" pageNumber="445" type="distribution">
<paragraph id="886CAE1DFFDF1F59FF15F9FA951BF90F" blockId="6.[151,1437,1643,2030]" box="[199,870,1643,1670]" pageId="6" pageNumber="445">
<emphasis id="BAA7720FFFDF1F59FF15F9FA9724F90C" bold="true" box="[199,345,1643,1669]" pageId="6" pageNumber="445">Distribution</emphasis>
.
<collectingCountry id="F0C4EE8DFFDF1F59FEB5F9FA97CDF90C" box="[359,432,1643,1669]" name="Brazil" pageId="6" pageNumber="445">Brazil</collectingCountry>
:
<collectingRegion id="4A1760FFFFDF1F59FE69F9FA94D7F90F" box="[443,682,1643,1670]" country="Brazil" name="Rio Grande do Norte" pageId="6" pageNumber="445">Rio Grande do Norte</collectingRegion>
,
<collectingRegion id="4A1760FFFFDF1F59FD66F9FA9488F90C" box="[692,757,1643,1669]" country="Brazil" name="Bahia" pageId="6" pageNumber="445">Bahia</collectingRegion>
(
<figureCitation id="10E8B298FFDF1F59FCD7F9FA9524F90F" box="[773,857,1643,1670]" captionStart="FIGURE 25" captionStartId="7.[151,250,1792,1816]" captionTargetBox="[174,1414,181,1768]" captionTargetId="figure-19@7.[174,1414,181,1768]" captionTargetPageId="7" captionText="FIGURE 25. Map showing known records of South American Sapygidae species." figureDoi="http://doi.org/10.5281/zenodo.13757880" httpUri="https://zenodo.org/record/13757880/files/figure.png" pageId="6" pageNumber="445">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="886CAE1DFFDF1F59FF15F91E95AAF82C" blockId="6.[151,1437,1643,2030]" pageId="6" pageNumber="445">
<emphasis id="BAA7720FFFDF1F59FF15F91E9706F920" bold="true" box="[199,379,1679,1705]" pageId="6" pageNumber="445">Sex association</emphasis>
. The male
<typeStatus id="576810BFFFDF1F59FE24F91E9425F920" box="[502,600,1679,1705]" pageId="6" pageNumber="445" type="holotype">holotype</typeStatus>
and a female
<typeStatus id="576810BFFFDF1F59FD21F901952EF923" box="[755,851,1680,1706]" pageId="6" pageNumber="445" type="paratype">paratype</typeStatus>
were collected in the same area and during the same period. Despite some sexually dimorphic differences—general body coloration, clypeus shape, antennae structure, and shape of the last sternite—we confidently assigned them to
<taxonomicName id="4FD3D59EFFDF1F59FC86F9499271F978" authorityName="Lopes &amp; Moleiro &amp; Lucena" authorityYear="2024" box="[852,1036,1751,1777]" class="Magnoliopsida" family="Asteraceae" genus="Huarpea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="445" phylum="Tracheophyta" rank="species" species="rotundicornis" status="sp. nov.">
<emphasis id="BAA7720FFFDF1F59FC86F9499271F978" box="[852,1036,1751,1777]" italics="true" pageId="6" pageNumber="445">H. rotundicornis</emphasis>
</taxonomicName>
<emphasis id="BAA7720FFFDF1F59FBC0F9469217F978" bold="true" box="[1042,1130,1751,1777]" pageId="6" pageNumber="445">
<taxonomicNameLabel id="A194CF74FFDF1F59FBC0F9469217F978" box="[1042,1130,1751,1777]" pageId="6" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
based on specific structural features. These include the metapleura tightly constricted by the propodeum, a strongly carinate frons, and a distinct smooth depression above the antennal socket contrasting with its surroundings. Given the rarity of sapygids and the aforementioned evidence, we concluded that the examined specimens belong to the same species. Additionally, a comparison of female specimens from
<collectingRegion id="4A1760FFFFDF1F59FD9AF8F6954CF80B" box="[584,817,1895,1922]" country="Brazil" name="Rio Grande do Norte" pageId="6" pageNumber="445">Rio Grande do Norte</collectingRegion>
and
<collectingRegion id="4A1760FFFFDF1F59FCB4F8F695DAF808" box="[870,935,1895,1921]" country="Brazil" name="Bahia" pageId="6" pageNumber="445">Bahia</collectingRegion>
further supports they correspond to the same taxon, apparently with broad geographic range within the semiarid region.
</paragraph>
</subSubSection>
<subSubSection id="C0C9FD96FFDF1F59FF15F83E929DF867" pageId="6" pageNumber="445" type="etymology">
<paragraph id="886CAE1DFFDF1F59FF15F83E929DF867" blockId="6.[151,1437,1643,2030]" pageId="6" pageNumber="445">
<emphasis id="BAA7720FFFDF1F59FF15F83E9735F840" bold="true" box="[199,328,1967,1993]" pageId="6" pageNumber="445">Etymology</emphasis>
. The specific epithet is composed of the Latin words “
<emphasis id="BAA7720FFFDF1F59FC17F83E9255F840" box="[965,1064,1967,1993]" italics="true" pageId="6" pageNumber="445">rotundus</emphasis>
” meaning rounded and “
<emphasis id="BAA7720FFFDF1F59FA98F82193EDF840" box="[1354,1424,1968,1993]" italics="true" pageId="6" pageNumber="445">cornis</emphasis>
” meaning horn, referring to the rounded last male flagellomere that contrasts with other congeners.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="16DCA625590FC852F9A007CBC165883C">Description of new species of deep water Sthenolepis Willey, 1905 and Neoleanira Pettibone, 1970 (Annelida, Sigalionidae) from off Northern California, with the redescription of Sthenolepis spargens Fauchald, 1972</mods:title>
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<paragraph id="8B9D36E3FFE55654FF2BFCF4C019DD2E" blockId="11.[151,590,834,861]" box="[151,590,834,861]" pageId="11" pageNumber="235">
<heading id="D0D5818FFFE55654FF2BFCF4C019DD2E" bold="true" box="[151,590,834,861]" fontSize="11" level="1" pageId="11" pageNumber="235" reason="1">
<emphasis id="B956EAF1FFE55654FF2BFCF4C019DD2E" bold="true" box="[151,590,834,861]" pageId="11" pageNumber="235">
Genus
<taxonomicName id="4C224D60FFE55654FF52FCF4C019DD2E" ID-CoL="84QTX" authority="Pettibone, 1970 b" authorityName="Pettibone" authorityYear="1970" box="[238,590,834,861]" class="Polychaeta" family="Sigalionidae" genus="Neoleanira" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="235" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFE55654FF52FCF4C322DD2F" bold="true" box="[238,373,834,861]" italics="true" pageId="11" pageNumber="235">Neoleanira</emphasis>
<bibRefCitation id="EFB34B12FFE55654FEC1FCF4C019DD2E" author="Pettibone, M. H." box="[381,590,834,860]" pageId="11" pageNumber="235" pagination="365 - 386" refId="ref13111" refString="Pettibone, M. H. (1970 b) Two new genera of Sigalionidae (Polychaeta). Proceedings of the Biological Society of Washington, 83 (34), 365 - 386. [https: // www. biodiversitylibrary. org / page / 34573005, https: // repository. si. edu / handle / 10088 / 3407]" type="journal article" year="1970">Pettibone, 1970b</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFE55654FF2BFC3DC0E1DDD7" box="[151,694,906,935]" pageId="11" pageNumber="235" type="type_taxon">
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<emphasis id="B956EAF1FFE55654FF2BFC3DC365DDD6" bold="true" box="[151,306,906,933]" pageId="11" pageNumber="235">
<typeStatus id="54998841FFE55654FF2BFC3DC285DDD7" box="[151,210,907,933]" pageId="11" pageNumber="235">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C224D60FFE55654FE85FC3DC0E1DDD7" ID-CoL="6YN59" authority="Orsted, 1845" authorityName="Orsted" authorityYear="1845" box="[313,694,906,935]" class="Polychaeta" family="Sigalionidae" genus="Sigalion" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="235" phylum="Annelida" rank="species" species="tetragonum">
<emphasis id="B956EAF1FFE55654FE85FC3DC048DDD6" box="[313,543,907,933]" italics="true" pageId="11" pageNumber="235">Sigalion tetragonum</emphasis>
<bibRefCitation id="EFB34B12FFE55654FD9AFC3AC0E1DDD7" author="Orsted, A. S." box="[550,694,906,935]" pageId="11" pageNumber="235" pagination="400 - 427" refId="ref13011" refString="Orsted, A. S. (1845) Fortegnelse over Dry, samlede I Christianiafjord ved Drobak fra 21 - 24 Juli 1844. Naturhistorisk Tidsskrift, KJObenhavn, 2 (1), 400 - 427." type="journal article" year="1845">Örsted, 1845</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFE55654FF7BFC18C049DAD3" pageId="11" pageNumber="235" type="diagnosis">
<paragraph id="8B9D36E3FFE55654FF7BFC18C049DAD3" blockId="11.[151,1437,906,1185]" pageId="11" pageNumber="235">
<emphasis id="B956EAF1FFE55654FF7BFC18C315DDBA" bold="true" box="[199,322,942,968]" pageId="11" pageNumber="235">Diagnosis.</emphasis>
(modified after
<bibRefCitation id="EFB34B12FFE55654FE44FC19C0EFDDBB" author="Aungtonya, C." box="[504,696,943,969]" pageId="11" pageNumber="235" pagination="205 - 235" refId="ref11789" refString="Aungtonya, C. (2002) A preliminary study of Sigalionidae (Annelida: Polychaeta) from the Andaman Sea off Southwestern Thailand, with an overview of presently recognized genera. Phuket Marine Biological Center Special Publication, 24, 205 - 235." type="journal article" year="2002">Aungtonya 2002</bibRefCitation>
;
<bibRefCitation id="EFB34B12FFE55654FD7FFC19C1A0DDBB" author="Eibye-Jacobsen, D. &amp; Aungtonya, A. &amp; Gonzalez, B. C." box="[707,1015,943,969]" pageId="11" pageNumber="235" refId="ref12258" refString="Eibye-Jacobsen, D., Aungtonya, A. &amp; Gonzalez, B. C. (2022) Sigalionidae Kinberg, 1856. (In): Purschke, G., Boggemann, M. &amp; Westheide, W. (Eds.), Pleistoannelida, Errantia II. Vol. 4. Walter de Gruyter, Berlin, pp. 114 - 138. https: // doi. org / 10.1515 / 9783110647167 - 006" type="book" year="2022">
Eibye-Jacobsen
<emphasis id="B956EAF1FFE55654FCC6FC19C1E4DDBB" box="[890,947,943,969]" italics="true" pageId="11" pageNumber="235">et al.</emphasis>
2022
</bibRefCitation>
).
<taxonomicName id="4C224D60FFE55654FBB6FC19C6C1DDBB" authorityName="Kinberg" authorityYear="1856" box="[1034,1174,943,969]" class="Polychaeta" family="Sigalionidae" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="235" phylum="Annelida" rank="subFamily" subFamily="Sigalioninae">Sigalioninae</taxonomicName>
with oval prostomium. Median antenna with long style; median antennal ceratophore with a pair of auricles; lateral antennae rather long, fused with dorsal side of segment 1. Without facial tubercle. Segment 1 with dorsal ctenidial pads and inner tentacular lobes. Segment 3 with a long dorsal cirrus. Lateral lips of mouth aperture without labial lobes. Elytra colorless, smooth, or with tubercles and papillae. Neuropodia in median and posterior segment with three neuropodial lobes, one prechaetal and two postchaetal. Neurochaetae compound, with canaliculate blades; spines and simple chaetae might be present in upper position.
</paragraph>
</subSubSection>
</treatment>
</document>

298
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<mods:title id="4127E01D3461FF7C50612A18F2AD65FB">Description of new species of deep water Sthenolepis Willey, 1905 and Neoleanira Pettibone, 1970 (Annelida, Sigalionidae) from off Northern California, with the redescription of Sthenolepis spargens Fauchald, 1972</mods:title>
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<paragraph id="8B9D36E3FFEC565DFF2BF9E3C00AD81D" blockId="2.[151,821,1621,1720]" box="[151,605,1621,1648]" pageId="2" pageNumber="226">
<heading id="D0D5818FFFEC565DFF2BF9E3C00AD81D" bold="true" box="[151,605,1621,1648]" fontSize="11" level="1" pageId="2" pageNumber="226" reason="1">
<taxonomicName id="4C224D60FFEC565DFF2BF9E3C00AD81D" ID-CoL="52HK3" authority="Fauchald, 1972" authorityName="Fauchald" authorityYear="1972" box="[151,605,1621,1648]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEC565DFF2BF9E3C00AD81D" bold="true" box="[151,605,1621,1648]" pageId="2" pageNumber="226">
<emphasis id="B956EAF1FFEC565DFF2BF9E3C3C0D802" bold="true" box="[151,407,1621,1648]" italics="true" pageId="2" pageNumber="226">Sthenolepis spargens</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFE22F9E3C00AD81D" author="Fauchald, K." box="[414,605,1621,1647]" pageId="2" pageNumber="226" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
</emphasis>
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<subSubSection id="C3386568FFEC565DFF2BF9CFC337D8E1" box="[151,352,1657,1683]" pageId="2" pageNumber="226" type="description">
<paragraph id="8B9D36E3FFEC565DFF2BF9CFC337D8E1" blockId="2.[151,821,1621,1720]" box="[151,352,1657,1683]" pageId="2" pageNumber="226">
<figureCitation id="13192A66FFEC565DFF2BF9CFC356D8E1" box="[151,257,1657,1683]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1763,1787]" captionStartId-1="4.[151,250,1267,1291]" captionTargetBox-0="[306,1276,179,1740]" captionTargetBox-1="[264,1323,194,1243]" captionTargetId-0="figure-26@3.[306,1281,179,1740]" captionTargetId-1="figure-17@4.[264,1323,181,1243]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." captionText-1="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13757502" figureDoi-1="http://doi.org/10.5281/zenodo.13757506" httpUri-0="https://zenodo.org/record/13757502/files/figure.png" httpUri-1="https://zenodo.org/record/13757506/files/figure.png" pageId="2" pageNumber="226">Figs. 1–2</figureCitation>
,
<tableCitation id="C6A00358FFEC565DFEB1F9CFC337D8E1" box="[269,352,1657,1683]" captionStart="TABLE 1" captionStartId="17.[151,235,463,488]" captionText="TABLE 1. Comparative table of the valid species of Sthenolepis Willey, 1905 (Read &amp; Fauchald 2024), including the geographical and bathymetrical distribution. The number of times in sizes relationships between wide and length is given by ‘×’." pageId="2" pageNumber="226">Table 1</tableCitation>
</paragraph>
</subSubSection>
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<taxonomicName id="4C224D60FFEC565DFF2BF92BC0D9D8C5" ID-CoL="52HK3" authority="Fauchald, 1972: 36 - 38" authorityName="Fauchald" authorityPageNumber="36 - 38" authorityYear="1972" box="[151,654,1693,1719]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEC565DFF2BF92BC3D4D8C5" box="[151,387,1693,1719]" italics="true" pageId="2" pageNumber="226">Sthenolepis spargens</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFE36F92BC0D9D8C5" author="Fauchald, K." box="[394,654,1693,1719]" pageId="2" pageNumber="226" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald, 1972: 36–38</bibRefCitation>
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, Pl. I, figs. f–i.
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<emphasis id="B956EAF1FFEC565DFF2BF953C328D88D" bold="true" box="[151,383,1765,1791]" pageId="2" pageNumber="226">Material examined.</emphasis>
<materialsCitation id="3B4A3CBEFFEC565DFE38F953C698D956" collectingDate="1967-11-13" collectorName="V Velero IV &amp; Campbell" country="Mexico" county="Mazatlan" elevation="2926" latitude="22.733" location="North Pacific Ocean" longLatPrecision="75" longitude="-108.079" municipality="Mazatlan" pageId="2" pageNumber="226" specimenCount="1" typeStatus="holotype">
<emphasis id="B956EAF1FFEC565DFE38F953C3AFD88D" bold="true" box="[388,504,1765,1791]" pageId="2" pageNumber="226">
<typeStatus id="54998841FFEC565DFE38F953C3A4D88D" box="[388,499,1765,1791]" pageId="2" pageNumber="226" type="holotype">Holotype</typeStatus>
.
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,
<collectingCountry id="F3357673FFEC565DFD57F953C114D88D" box="[747,835,1765,1791]" name="Mexico" pageId="2" pageNumber="226">Mexico</collectingCountry>
,
<collectingMunicipality id="6BF9AC99FFEC565DFCF1F953C1EFD88D" box="[845,952,1765,1791]" pageId="2" pageNumber="226">Mazatlán</collectingMunicipality>
,
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from
<collectingCounty id="62FC4E6FFFEC565DFBEAF953C6E9D88D" box="[1110,1214,1765,1791]" pageId="2" pageNumber="226">Mazatlán</collectingCounty>
lighthouse,
<geoCoordinate id="EE165024FFEC565DFAF8F953C2F9D956" degrees="22.733" direction="north" orientation="latitude" pageId="2" pageNumber="226" precision="55" value="22.733">22.733° N</geoCoordinate>
,
<geoCoordinate id="EE165024FFEC565DFF05F8BFC316D956" box="[185,321,1801,1828]" degrees="108.079" direction="west" orientation="longitude" pageId="2" pageNumber="226" precision="55" value="-108.079">108.079° W</geoCoordinate>
,
<quantity id="4CDA9B06FFEC565DFEF0F8BFC3F4D956" box="[332,419,1801,1828]" metricMagnitude="3" metricUnit="m" metricValue="2.926" pageId="2" pageNumber="226" unit="m" value="2926.0">
<elevation id="000FD1D0FFEC565DFEF0F8BFC3F4D956" box="[332,419,1801,1828]" metricMagnitude="3" metricUnit="m" metricValue="2.926" pageId="2" pageNumber="226" unit="m" value="2926.0">2926 m</elevation>
</quantity>
,
<date id="FF9C1023FFEC565DFE13F8BFC013D951" box="[431,580,1801,1828]" pageId="2" pageNumber="226" value="1967-11-13">
<collectingDate id="EFD8E9CBFFEC565DFE13F8BFC013D951" box="[431,580,1801,1828]" pageId="2" pageNumber="226" value="1967-11-13">13 Nov 1967</collectingDate>
</date>
, R/
<collectorName id="26D75335FFEC565DFDD6F8BFC0BBD951" box="[618,748,1801,1827]" pageId="2" pageNumber="226">
V
<emphasis id="B956EAF1FFEC565DFD38F8BFC0BBD951" box="[644,748,1801,1827]" italics="true" pageId="2" pageNumber="226">Velero IV</emphasis>
</collectorName>
,
<collectorName id="26D75335FFEC565DFD46F8BFC130D951" box="[762,871,1801,1827]" pageId="2" pageNumber="226">Campbell</collectorName>
grab (LACM-AHF-Poly 1053).
</materialsCitation>
</paragraph>
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<paragraph id="8B9D36E3FFEC565DFF7BF89BC619D9C6" blockId="2.[151,1437,1765,1972]" pageId="2" pageNumber="226">
<emphasis id="B956EAF1FFEC565DFF7BF89BC30ED935" bold="true" box="[199,345,1837,1863]" pageId="2" pageNumber="226">Description.</emphasis>
<typeStatus id="54998841FFEC565DFEDEF89BC39DD935" box="[354,458,1837,1863]" pageId="2" pageNumber="226" type="holotype">Holotype</typeStatus>
incomplete with 73 segments,
<quantity id="4CDA9B06FFEC565DFC8EF89BC1D4D93A" box="[818,899,1837,1864]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="226" unit="mm" value="17.0">17 mm</quantity>
long,
<quantity id="4CDA9B06FFEC565DFC72F89BC645D93A" box="[974,1042,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="226" unit="mm" value="5.0">5 mm</quantity>
to segment 30,
<quantity id="4CDA9B06FFEC565DFB71F89BC747D93A" box="[1229,1296,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="226" unit="mm" value="1.0">1 mm</quantity>
wide. Body slender, tapered, fragile posteriorly due to poor condition; pale orange, slightly translucent (
<figureCitation id="13192A66FFEC565DFB12F8E7C75ED919" box="[1198,1289,1873,1899]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="2" pageNumber="226">Fig. 1A</figureCitation>
). Mid-dorsal line smooth, no elytra remain, venter smooth. Elytrophores on segments 2, 4, 5, 7, then alternate segment to 25, then present in all segments. Elytrophores short, longer and thinner in posterior segments.
</paragraph>
<caption id="DF5D666BFFED565CFF2BF955C11FD9A6" ID-DOI="http://doi.org/10.5281/zenodo.13757502" ID-Zenodo-Dep="13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="3" pageNumber="227" startId="3.[151,250,1763,1787]" targetBox="[306,1276,179,1740]" targetPageId="3" targetType="figure">
<paragraph id="8B9D36E3FFED565CFF2BF955C11FD9A6" blockId="3.[151,1437,1763,2004]" pageId="3" pageNumber="227">
<emphasis id="B956EAF1FFED565CFF2BF955C0F7D889" bold="true" box="[151,672,1763,1788]" pageId="3" pageNumber="227">
FIGURE 1.
<taxonomicName id="4C224D60FFED565CFEA5F955C0F7D889" authority="Fauchald, 1972" authorityName="Fauchald" authorityYear="1972" box="[281,672,1763,1788]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="227" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFED565CFEA5F955C3A2D889" bold="true" box="[281,501,1763,1787]" italics="true" pageId="3" pageNumber="227">Sthenolepis spargens</emphasis>
<bibRefCitation id="EFB34B12FFED565CFE47F952C0F7D889" author="Fauchald, K." box="[507,672,1763,1788]" pageId="3" pageNumber="227" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
</taxonomicName>
</emphasis>
. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta. Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm.
</paragraph>
</caption>
<caption id="DF5D666BFFEA565BFF2BFB45C1B2DBB2" ID-DOI="http://doi.org/10.5281/zenodo.13757506" ID-Zenodo-Dep="13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="4" pageNumber="228" startId="4.[151,250,1267,1291]" targetBox="[264,1323,194,1243]" targetPageId="4" targetType="figure">
<paragraph id="8B9D36E3FFEA565BFF2BFB45C1B2DBB2" blockId="4.[151,1437,1267,1472]" pageId="4" pageNumber="228">
<emphasis id="B956EAF1FFEA565BFF2BFB45C0E1DB79" bold="true" box="[151,694,1267,1292]" pageId="4" pageNumber="228">
FIGURE 2.
<taxonomicName id="4C224D60FFEA565BFE9EFB45C0E6DB79" authority="Fauchald 1972" authorityName="Fauchald" authorityYear="1972" box="[290,689,1267,1292]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="228" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEA565BFE9EFB45C055DB79" bold="true" box="[290,514,1267,1291]" italics="true" pageId="4" pageNumber="228">Sthenolepis spargens</emphasis>
<bibRefCitation id="EFB34B12FFEA565BFDB1FB42C0E6DB79" author="Fauchald, K." box="[525,689,1267,1292]" pageId="4" pageNumber="228" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald 1972</bibRefCitation>
</taxonomicName>
.
</emphasis>
Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm.
</paragraph>
</caption>
<paragraph id="8B9D36E3FFEA565BFF7BF9A4C353D9AE" blockId="4.[151,1437,1554,2012]" pageId="4" pageNumber="228">
Prostomium pale orange, darker on cerebral lobes; oval, wider than long. Eyes lacking. Lateral antennae short, inconspicuous, inserted on inner dorsal side of tentacular segment. Median antenna with ceratophore slightly shorter than prostomial length; style lost; inserted centrally on prostomium. Auricles semispherical, half as long as ceratophore; inserted basally and laterally on ceratophore. Tentacular segment uniramous, chaetae verticillate. Dorsal tentacular cirri 12× longer than tentacular neuropodia, ventral tentacular cirri short, 2× longer than tentacular neuropodia (
<figureCitation id="13192A66FFEA565BFE9FF970C32ED892" box="[291,377,1734,1760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="4" pageNumber="228">Fig. 1B</figureCitation>
). Inner tentacular lobes ¼ larger than inner palpal sheaths (
<figureCitation id="13192A66FFEA565BFBB8F970C60DD892" box="[1028,1114,1734,1760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="4" pageNumber="228">Fig. 1C</figureCitation>
). Palps reaching segment 20. Inner and outer palpal sheaths subequal. Buccal cirri, slightly larger than remaining ventral cirri. Buccal ctenidial pads, enlarged, inserted anterolaterally on buccal aperture (
<figureCitation id="13192A66FFEA565BFC9AF8B8C12AD95A" box="[806,893,1806,1832]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="4" pageNumber="228">Fig. 1C</figureCitation>
). Ctenidial pads from segment 1; segment 1 with only one dorsal ctenidial pad, succeeding segments with 3–6 ctenidial pads: 2 large and bulbous pads placed on dorsolateral surface of segment; 1–2 smaller, half as large as dorsolateral ones, placed on dorsal side of notopodia; 0–1, small pads, placed on anterior inner side of parapodia; and 1 truncated pad, inserted ventrally. Branchiae from segment 2, small, becoming slightly larger from segment 7, filiform, cilia not seen. Nephridial papillae not observed.
</paragraph>
<paragraph id="8B9D36E3FFEB565AFF7BFF21C3D4DEC3" blockId="5.[151,1437,151,1473]" box="[199,387,151,177]" pageId="5" pageNumber="229">Elytra unknown.</paragraph>
<paragraph id="8B9D36E3FFEB565AFF7BFF0DC75CDF87" blockId="5.[151,1437,151,1473]" pageId="5" pageNumber="229">
Segment 3 (
<figureCitation id="13192A66FFEB565AFEF7FF0DC3F6DEA7" box="[331,417,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="5" pageNumber="229">Fig. 1E</figureCitation>
): Notopodia conical, ½ as long as neuropodia. Notacicula slender, protruding from the body wall. Notochaetae with up to 10 simple verticillate chaetae (
<figureCitation id="13192A66FFEB565AFC98FF69C120DE8B" box="[804,887,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="5" pageNumber="229">Fig. 1E</figureCitation>
, inset), smallest 4× longer than notopodia, longest 7× as long. Neuropodia lanceolate. Prechaetal lobe entire, without stylodes. Postchaetal lobes well differentiated, without stylodes. Neuracicula thick, inserted in prechaetal lobe, protruding from the body wall. Neurochaetae only spinigers (
<figureCitation id="13192A66FFEB565AFEB2FEFDC33DDF17" box="[270,362,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="5" pageNumber="229">Fig. 1D</figureCitation>
). Upper group (unit A) with 4 chaetae, handles thick with a subdistal row of small denticles, blades medium-sized, 18–20× as long as wide. Upper middle group (unit B) with 5 chaetae, blades medium-sized to long, 24–26× as long as wide. Middle group (unit C) with 4 chaetae, blades short, 12–13× as long as wide. Lower middle group (subunit 1) with 6 chaetae, blades medium-sized to long, 23–24× as long as wide. Lowest group (unit D) with 5 chaetae, blades short, 13× as long as wide. Ventral cirri 1/3 as long as neuropodia (
<figureCitation id="13192A66FFEB565AFB1BFE6DC6ABDF87" box="[1191,1276,475,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1763,1787]" captionTargetBox="[306,1276,179,1740]" captionTargetId="figure-26@3.[306,1281,179,1740]" captionTargetPageId="3" captionText="FIGURE 1. Sthenolepis spargens Fauchald, 1972. Holotype (LACM-AHF-Poly 1053). A. Incomplete specimen, inset: closeup of stylodes emerging from the posterior surface on posterior segments. B. Drawing of anterior end, dorsal view. C. Drawing of anterior end, ventral view. D. Neurochaetae from segment 3. E. Parapodium from segment 3, anterior view, inset: close-up of a notochaeta.Abbreviations: au= auricle, ct= ctenidia, dtu= dorsal tubercle, dtc= dorsal tentacular cirrus, ftu= facial tubercle, ipas= inner palpal sheath, itl= inner tentacular lobe, lan= lateral antenna, man= median antenna, opas= outer palpal sheath, pa= palp, S1= lower middle group, UA= upper group, UB= upper middle group, UC=middle group, UD= lowest group, vtc= ventral tentacular cirrus. Scales bars: A, B, C: 1 mm, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757502" httpUri="https://zenodo.org/record/13757502/files/figure.png" pageId="5" pageNumber="229">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E3FFEB565AFF7BFE49C10EDD2F" blockId="5.[151,1437,151,1473]" pageId="5" pageNumber="229">
Segment 20 (middle segment) (
<figureCitation id="13192A66FFEB565AFD82FE49C097DC6B" box="[574,704,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1267,1291]" captionTargetBox="[264,1323,194,1243]" captionTargetId="figure-17@4.[264,1323,181,1243]" captionTargetPageId="4" captionText="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="5" pageNumber="229">Fig. 2A–C</figureCitation>
): Notopodia conical, as long as neuropodia. Notacicula thick. Notochaetae with up to 20 simple verticillate chaetae (
<figureCitation id="13192A66FFEB565AFD55FD95C16ADC4F" box="[745,829,547,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1267,1291]" captionTargetBox="[264,1323,194,1243]" captionTargetId="figure-17@4.[264,1323,181,1243]" captionTargetPageId="4" captionText="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="5" pageNumber="229">Fig. 2E</figureCitation>
, inset), smallest as long as notopodia, longest 2× longer than notopodia. Neuropodia conical. Prechaetal lobe entire with a few small dendritic stylodes on it. Postchaetal lobes well differentiated, without stylodes. Neuracicula thick, inserted in prechaetal lobe, protruding from body wall. Neurochaetae only spinigers (
<figureCitation id="13192A66FFEB565AFD94FD39C0D4DCDB" box="[552,643,655,681]" captionStart="FIGURE 2" captionStartId="4.[151,250,1267,1291]" captionTargetBox="[264,1323,194,1243]" captionTargetId="figure-17@4.[264,1323,181,1243]" captionTargetPageId="4" captionText="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="5" pageNumber="229">Fig. 2D</figureCitation>
). Upper group (unit A) with 4 chaetae, handles thick with an enlarged subdistal tooth (
<figureCitation id="13192A66FFEB565AFEF7FD05C3F3DCBF" box="[331,420,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1267,1291]" captionTargetBox="[264,1323,194,1243]" captionTargetId="figure-17@4.[264,1323,181,1243]" captionTargetPageId="4" captionText="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="5" pageNumber="229">Fig. 2D</figureCitation>
, inset); blades short, 14–15× as long as wide. Upper middle group (unit B) with 7 chaetae, handle thick with an enlarged subdistal tooth, blades medium-sized, 23× as long as wide. Middle group (unit C) with 6 chaetae, blades medium-sized, 19–20× as long as wide. Lower middle group (subunit 1) with 3 chaetae, blades medium-sized, 20–21× as long as wide. Lowest group (unit D) with 4 chaetae, blades medium-sized, 16–17× as long as wide. Ventral cirri half as long as neuropodia (
<figureCitation id="13192A66FFEB565AFD49FCF5C11BDD2F" box="[757,844,835,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1267,1291]" captionTargetBox="[264,1323,194,1243]" captionTargetId="figure-17@4.[264,1323,181,1243]" captionTargetPageId="4" captionText="FIGURE 2. Sthenolepis spargens Fauchald 1972. Holotype (LACM-AHF-Poly 1053). A. Drawing of parapodium from segment 20, anterior view. B. Drawing of same, posterior view. C. Diagrammatic of same, lateral view, color lines indicate the insertion of neurochaetal groups, color scheme following that shown in D. D. Neurochaetae from segment 20, inset: close-up of the distal margin of upper group handle. E. Parapodium from segment 20, anterior view, inset: close-up of a notochaeta. Abbreviations: as= anterior side; ps= posterior side. S1= lower middle group, UA= upper group, UB= upper middle group, UC= middle group, UD= lowest group. Scale bars: A–C: no scale, D: 50 µm, E: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13757506" httpUri="https://zenodo.org/record/13757506/files/figure.png" pageId="5" pageNumber="229">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E3FFEB565AFF7BFCD1C0CBDDF3" blockId="5.[151,1437,151,1473]" box="[199,668,871,897]" pageId="5" pageNumber="229">Posterior region lost. Pygidium unknown.</paragraph>
</subSubSection>
<subSubSection id="C3386568FFEB565AFF7BFC3DC694DBEF" pageId="5" pageNumber="229" type="discussion">
<paragraph id="8B9D36E3FFEB565AFF7BFC3DC06DDAD3" blockId="5.[151,1437,151,1473]" pageId="5" pageNumber="229">
<emphasis id="B956EAF1FFEB565AFF7BFC3DC36CDDD7" bold="true" box="[199,315,907,933]" pageId="5" pageNumber="229">Remarks.</emphasis>
<bibRefCitation id="EFB34B12FFEB565AFEFAFC3DC055DDD7" author="Fauchald, K." box="[326,514,907,933]" pageId="5" pageNumber="229" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald (1972)</bibRefCitation>
described
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<emphasis id="B956EAF1FFEB565AFD3EFC3DC125DDD6" box="[642,882,907,933]" italics="true" pageId="5" pageNumber="229">Sthenolepis spargens</emphasis>
</taxonomicName>
based on specimens from the southern Gulf of
<collectingRegion id="49E6F801FFEB565AFF2BFC19C35EDDBB" box="[151,265,943,969]" country="United States of America" name="California" pageId="5" pageNumber="229">California</collectingRegion>
collected at
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depth. In the original description,
<bibRefCitation id="EFB34B12FFEB565AFCCFFC19C67CDDBB" author="Fauchald, K." box="[883,1067,943,969]" pageId="5" pageNumber="229" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald (1972)</bibRefCitation>
stated that
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<emphasis id="B956EAF1FFEB565AFB16FC19C77DDDBA" box="[1194,1322,943,969]" italics="true" pageId="5" pageNumber="229">S. spargens</emphasis>
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possesses long lateral antennae, as well as a small dorsal cirrus on segment 3. However, examination of the
<typeStatus id="54998841FFEB565AFB69FC65C760DD9F" box="[1237,1335,979,1005]" pageId="5" pageNumber="229" type="holotype">holotype</typeStatus>
revealed that what Fauchald interpreted as lateral antennae are the dorsal tentacular cirri. The lateral antennae in
<taxonomicName id="4C224D60FFEB565AFAA7FC41C7CBDA63" authorityName="Willey" authorityYear="1905" box="[1307,1436,1015,1041]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEB565AFAA7FC41C7CBDA63" box="[1307,1436,1015,1041]" italics="true" pageId="5" pageNumber="229">Sthenolepis</emphasis>
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are small and located on the dorsal side of the tentacular segment but displaced to the inner side of the notopodium, very close to the anterior prostomial border covered by the auricles on the median antennal ceratophore. Therefore, they were likely overlooked by Fauchald. It was also noticed that the
<typeStatus id="54998841FFEB565AFC0CFBD5C645DA0F" box="[944,1042,1123,1149]" pageId="5" pageNumber="229" type="holotype">holotype</typeStatus>
lacks a dorsal cirrus on segment 3 and instead, has an enlarged tubercle.
</paragraph>
<paragraph id="8B9D36E3FFEB565AFF7BFB1DC694DBEF" blockId="5.[151,1437,151,1473]" pageId="5" pageNumber="229">
<bibRefCitation id="EFB34B12FFEB565AFF7BFB1DC32ADAB7" author="Fauchald, K." box="[199,381,1195,1221]" pageId="5" pageNumber="229" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald (1972)</bibRefCitation>
discussed in the remarks section of
<taxonomicName id="4C224D60FFEB565AFD43FB1DC6C8DAB7" authority="Fauchald, 1972" authorityName="Fauchald" authorityYear="1972" box="[767,1183,1195,1221]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="species" species="racemosa">
<emphasis id="B956EAF1FFEB565AFD43FB1DC1A6DAB6" box="[767,1009,1195,1221]" italics="true" pageId="5" pageNumber="229">Sthenolepis racemosa</emphasis>
<bibRefCitation id="EFB34B12FFEB565AFC49FB1DC6C8DAB7" author="Fauchald, K." box="[1013,1183,1195,1221]" pageId="5" pageNumber="229" pagination="1 - 575" refId="ref12338" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575. [http: // hdl. handle. net / 10088 / 6207]" type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
</taxonomicName>
that
<taxonomicName id="4C224D60FFEB565AFB6EFB1DC718DAB6" authorityName="Fauchald" authorityYear="1972" box="[1234,1359,1195,1221]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEB565AFB6EFB1DC718DAB6" box="[1234,1359,1195,1221]" italics="true" pageId="5" pageNumber="229">S. spargens</emphasis>
</taxonomicName>
differs from all the eye-less
<taxonomicName id="4C224D60FFEB565AFE35FB79C033DA9B" box="[393,612,1231,1257]" pageId="5" pageNumber="229">
<emphasis id="B956EAF1FFEB565AFE35FB79C05DDA9B" box="[393,522,1231,1257]" italics="true" pageId="5" pageNumber="229">Sthenolepis</emphasis>
species
</taxonomicName>
in having four fimbriae (=stylodes) in the notopodium. The comparison between
<taxonomicName id="4C224D60FFEB565AFF40FB45C32ADB7E" authorityName="Fauchald" authorityYear="1972" box="[252,381,1267,1293]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEB565AFF40FB45C32ADB7E" box="[252,381,1267,1293]" italics="true" pageId="5" pageNumber="229">S. spargens</emphasis>
</taxonomicName>
and the eye-less
<taxonomicName id="4C224D60FFEB565AFDFDFB45C14DDB7F" box="[577,794,1267,1293]" pageId="5" pageNumber="229">
<emphasis id="B956EAF1FFEB565AFDFDFB45C095DB7F" box="[577,706,1267,1293]" italics="true" pageId="5" pageNumber="229">Sthenolepis</emphasis>
species
</taxonomicName>
was pertinent at the time. However, all those species are now contained either in
<taxonomicName id="4C224D60FFEB565AFE14FAA1C071DB43" authorityName="Pettibone" authorityYear="1970" box="[424,550,1303,1329]" class="Polychaeta" family="Sigalionidae" genus="Neoleanira" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEB565AFE14FAA1C071DB43" box="[424,550,1303,1329]" italics="true" pageId="5" pageNumber="229">Neoleanira</emphasis>
</taxonomicName>
or
<taxonomicName id="4C224D60FFEB565AFDF0FAA1C0F2DB42" authorityName="Kinberg" authorityYear="1856" box="[588,677,1303,1328]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEB565AFDF0FAA1C0F2DB42" box="[588,677,1303,1328]" italics="true" pageId="5" pageNumber="229">Leanira</emphasis>
</taxonomicName>
(
<emphasis id="B956EAF1FFEB565AFD09FAA1C18FDB42" box="[693,984,1303,1329]" italics="true" pageId="5" pageNumber="229">
e.g.
<taxonomicName id="4C224D60FFEB565AFD59FAA1C18FDB42" authorityName="Fauchald" authorityYear="1972" box="[741,984,1303,1329]" class="Polychaeta" family="Sigalionidae" genus="Neoleanira" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="species" species="racemosa">Neoleanira racemosa</taxonomicName>
</emphasis>
), and both genera are known for having members eyes-less (
<bibRefCitation id="EFB34B12FFEB565AFECAFA8DC065DB27" author="Pettibone, M. H." box="[374,562,1339,1365]" pageId="5" pageNumber="229" pagination="365 - 386" refId="ref13111" refString="Pettibone, M. H. (1970 b) Two new genera of Sigalionidae (Polychaeta). Proceedings of the Biological Society of Washington, 83 (34), 365 - 386. [https: // www. biodiversitylibrary. org / page / 34573005, https: // repository. si. edu / handle / 10088 / 3407]" type="journal article" year="1970">Pettibone 1970b</bibRefCitation>
). The observation on the stylodes was confirmed, but their number was noticed to vary according to the segment observed, increasing in posterior segments. Currently,
<taxonomicName id="4C224D60FFEB565AFBD1FAE9C6BADB0A" authorityName="Fauchald" authorityYear="1972" box="[1133,1261,1375,1401]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="229" phylum="Annelida" rank="species" species="spargens">
<emphasis id="B956EAF1FFEB565AFBD1FAE9C6BADB0A" box="[1133,1261,1375,1401]" italics="true" pageId="5" pageNumber="229">S. spargens</emphasis>
</taxonomicName>
differs from all
<taxonomicName id="4C224D60FFEB565AFF2BFA35C324DBEF" box="[151,371,1411,1437]" pageId="5" pageNumber="229">
<emphasis id="B956EAF1FFEB565AFF2BFA35C34FDBEF" box="[151,280,1411,1437]" italics="true" pageId="5" pageNumber="229">Sthenolepis</emphasis>
species
</taxonomicName>
, by lacking eyes and having long palps and dorsal tentacular cirri (
<tableCitation id="C6A00358FFEB565AFBE2FA35C6E5DBEF" box="[1118,1202,1411,1437]" captionStart="TABLE 1" captionStartId="17.[151,235,463,488]" captionText="TABLE 1. Comparative table of the valid species of Sthenolepis Willey, 1905 (Read &amp; Fauchald 2024), including the geographical and bathymetrical distribution. The number of times in sizes relationships between wide and length is given by ‘×’." pageId="5" pageNumber="229">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFEB565AFF7BFA10C773DBB3" box="[199,1316,1446,1473]" pageId="5" pageNumber="229" type="distribution">
<paragraph id="8B9D36E3FFEB565AFF7BFA10C773DBB3" blockId="5.[151,1437,151,1473]" box="[199,1316,1446,1473]" pageId="5" pageNumber="229">
<emphasis id="B956EAF1FFEB565AFF7BFA10C336DBB2" bold="true" box="[199,353,1446,1472]" pageId="5" pageNumber="229">Distribution.</emphasis>
Northeastern Temperate Pacific, Western
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, Gulf of
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,
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m
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.
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Genus
<taxonomicName id="4C224D60FFEC565DFF52FCD0C04CDDF2" ID-CoL="84TVS" authority="Willey, 1905" authorityName="Willey" authorityYear="1905" box="[238,539,870,897]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFF52FCD0C32DDDF3" bold="true" box="[238,378,870,897]" italics="true" pageId="2" pageNumber="226">Sthenolepis</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFE3EFCD0C04CDDF2" author="Willey, A." box="[386,539,870,896]" pageId="2" pageNumber="226" pagination="243 - 319" refId="ref13600" refString="Willey, A. (1905) Report on the Polychaeta collected by Professor Herdman, at Ceylon, in 1902. Ceylon Pearl Oyster Fisheries, Supplementary Reports, 4 (30), 243 - 319. [https: // biodiversitylibrary. org / page / 1936112]" type="book chapter" year="1905">Willey, 1905</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFEC565DFF2BFC1DC14EDDB6" box="[151,793,939,964]" pageId="2" pageNumber="226" type="type_taxon">
<paragraph id="8B9D36E3FFEC565DFF2BFC1DC14EDDB6" blockId="2.[151,793,939,964]" box="[151,793,939,964]" pageId="2" pageNumber="226">
<emphasis id="B956EAF1FFEC565DFF2BFC1DC374DDB1" bold="true" box="[151,291,939,963]" pageId="2" pageNumber="226">
<typeStatus id="54998841FFEC565DFF2BFC1DC29BDDB1" box="[151,204,939,963]" pageId="2" pageNumber="226">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C224D60FFEC565DFE96FC1AC0D5DDB1" ID-CoL="3SPFB" authority="McIntosh, 1885" authorityName="McIntosh" authorityYear="1885" box="[298,642,939,963]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="species" species="japonica">
<emphasis id="B956EAF1FFEC565DFE96FC1AC38EDDB1" box="[298,473,940,963]" italics="true" pageId="2" pageNumber="226">Leanira Japonica</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFE63FC1DC0D5DDB1" author="McIntosh, W. C." box="[479,642,939,963]" pageId="2" pageNumber="226" pagination="1 - 554" refId="ref12874" refString="McIntosh, W. C. (1885) On the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Challenger reports, 12, 34, 1 - 554. [https: // biodiversitylibary. org / page 50688432]" type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFEC565DFF2BFC47C2AEDB75" pageId="2" pageNumber="226" type="diagnosis">
<paragraph id="8B9D36E3FFEC565DFF2BFC47C2AEDB75" blockId="2.[151,1437,1009,1540]" pageId="2" pageNumber="226">
<emphasis id="B956EAF1FFEC565DFF2BFC47C345DA79" bold="true" box="[151,274,1009,1035]" pageId="2" pageNumber="226">Diagnosis.</emphasis>
(modified after
<bibRefCitation id="EFB34B12FFEC565DFE75FC44C0DEDA79" author="Aungtonya, C." box="[457,649,1009,1036]" pageId="2" pageNumber="226" pagination="205 - 235" refId="ref11789" refString="Aungtonya, C. (2002) A preliminary study of Sigalionidae (Annelida: Polychaeta) from the Andaman Sea off Southwestern Thailand, with an overview of presently recognized genera. Phuket Marine Biological Center Special Publication, 24, 205 - 235." type="journal article" year="2002">Aungtonya 2002</bibRefCitation>
;
<bibRefCitation id="EFB34B12FFEC565DFD29FC47C19DDA79" author="Eibye-Jacobsen, D. &amp; Aungtonya, A. &amp; Gonzalez, B. C." box="[661,970,1009,1035]" pageId="2" pageNumber="226" refId="ref12258" refString="Eibye-Jacobsen, D., Aungtonya, A. &amp; Gonzalez, B. C. (2022) Sigalionidae Kinberg, 1856. (In): Purschke, G., Boggemann, M. &amp; Westheide, W. (Eds.), Pleistoannelida, Errantia II. Vol. 4. Walter de Gruyter, Berlin, pp. 114 - 138. https: // doi. org / 10.1515 / 9783110647167 - 006" type="book" year="2022">
Eibye-Jacobsen
<emphasis id="B956EAF1FFEC565DFCF0FC44C1D2DA79" box="[844,901,1009,1035]" italics="true" pageId="2" pageNumber="226">et al.</emphasis>
2022
</bibRefCitation>
).
<taxonomicName id="4C224D60FFEC565DFC61FC47C63EDA79" authorityName="Kinberg" authorityYear="1856" box="[989,1129,1009,1035]" class="Polychaeta" family="Sigalionidae" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="subFamily" subFamily="Sigalioninae">Sigalioninae</taxonomicName>
with prostomium rounded. Median antenna with long style; median antennal ceratophore with a pair of auricles; lateral antennae short, fused to the inner side of segment 1. With facial tubercle. Segment 1 with dorsal ctenidial pads and inner tentacular lobes. Segment 3 without dorsal cirri, but dorsal tubercles. Lateral lips of mouth aperture without labial lobes. Elytra colorless, smooth, or with tubercles and papillae. Neuropodia in median and posterior segments with three neuropodial lobes, one prechaetal and two postchaetal. Neurochaetae only compound spinigers, with canaliculate blades, non-canaliculate blades may be present in lower position, simple spinose capillaries may be present in upper position.
</paragraph>
</subSubSection>
<subSubSection id="C3386568FFEC565DFF7BFAA7C7D0D871" pageId="2" pageNumber="226" type="discussion">
<paragraph id="8B9D36E3FFEC565DFF7BFAA7C7D0D871" blockId="2.[151,1437,1009,1540]" pageId="2" pageNumber="226">
<emphasis id="B956EAF1FFEC565DFF7BFAA7C36CDB59" bold="true" box="[199,315,1297,1323]" pageId="2" pageNumber="226">Remarks.</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFEFFFAA7C05ADB59" author="Pettibone, M. H." box="[323,525,1297,1323]" pageId="2" pageNumber="226" pagination="365 - 386" refId="ref13111" refString="Pettibone, M. H. (1970 b) Two new genera of Sigalionidae (Polychaeta). Proceedings of the Biological Society of Washington, 83 (34), 365 - 386. [https: // www. biodiversitylibrary. org / page / 34573005, https: // repository. si. edu / handle / 10088 / 3407]" type="journal article" year="1970">Pettibone (1970b)</bibRefCitation>
proposed two sigalionid genera using specimens of
<taxonomicName id="4C224D60FFEC565DFBE0FAA4C6E2DB59" authorityName="Kinberg" authorityYear="1856" box="[1116,1205,1298,1323]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFBE0FAA4C6E2DB59" box="[1116,1205,1298,1323]" italics="true" pageId="2" pageNumber="226">Leanira</emphasis>
</taxonomicName>
,
<taxonomicName id="4C224D60FFEC565DFB7FFAA7C716DB59" authorityName="Pettibone" authorityYear="1970" box="[1219,1345,1297,1323]" class="Polychaeta" family="Sigalionidae" genus="Neoleanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFB7FFAA7C716DB59" box="[1219,1345,1297,1323]" italics="true" pageId="2" pageNumber="226">Neoleanira</emphasis>
</taxonomicName>
, mostly from deep water, and
<taxonomicName id="4C224D60FFEC565DFE33FA83C070DB3D" authorityName="Pettibone" authorityYear="1970" box="[399,551,1333,1359]" class="Polychaeta" family="Sigalionidae" genus="Horstileanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFE33FA83C070DB3D" box="[399,551,1333,1359]" italics="true" pageId="2" pageNumber="226">Horstileanira</emphasis>
</taxonomicName>
, mostly from shallow water. The name
<taxonomicName id="4C224D60FFEC565DFC51FA83C6D2DB3D" authorityName="Pettibone" authorityYear="1970" box="[1005,1157,1333,1359]" class="Polychaeta" family="Sigalionidae" genus="Horstileanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFC51FA83C6D2DB3D" box="[1005,1157,1333,1359]" italics="true" pageId="2" pageNumber="226">Horstileanira</emphasis>
</taxonomicName>
indicates morphological closeness with
<taxonomicName id="4C224D60FFEC565DFE81FAECC3C1DB01" authorityName="Kinberg" authorityYear="1856" box="[317,406,1370,1395]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFE81FAECC3C1DB01" box="[317,406,1370,1395]" italics="true" pageId="2" pageNumber="226">Leanira</emphasis>
</taxonomicName>
: however,
<taxonomicName id="4C224D60FFEC565DFDB2FAEFC0D8DB01" authorityName="Willey" authorityYear="1905" box="[526,655,1369,1395]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFDB2FAEFC0D8DB01" box="[526,655,1369,1395]" italics="true" pageId="2" pageNumber="226">Sthenolepis</emphasis>
</taxonomicName>
seems to be the genus that
<emphasis id="B956EAF1FFEC565DFC0AFAEFC619DB01" box="[950,1102,1369,1395]" italics="true" pageId="2" pageNumber="226">Hortsileanira</emphasis>
resembles the most. The latter was noted by
<bibRefCitation id="EFB34B12FFEC565DFE8DFAC8C055DBE5" author="Aungtonya, C." box="[305,514,1405,1432]" pageId="2" pageNumber="226" pagination="205 - 235" refId="ref11789" refString="Aungtonya, C. (2002) A preliminary study of Sigalionidae (Annelida: Polychaeta) from the Andaman Sea off Southwestern Thailand, with an overview of presently recognized genera. Phuket Marine Biological Center Special Publication, 24, 205 - 235." type="journal article" year="2002">Aungtonya (2002)</bibRefCitation>
, who suggested that both genera need further study to understand the morphology of each properly. Furthermore, some former members of
<taxonomicName id="4C224D60FFEC565DFC86FA17C1ECDBC9" authorityName="Willey" authorityYear="1905" box="[826,955,1441,1467]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFC86FA17C1ECDBC9" box="[826,955,1441,1467]" italics="true" pageId="2" pageNumber="226">Sthenolepis</emphasis>
</taxonomicName>
, with unclear generic and specific status such as
<taxonomicName id="4C224D60FFEC565DFF4DFA70C03EDBAD" authority="Takahashi, 1938" authorityName="Takahashi" authorityYear="1938" box="[241,617,1477,1503]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="species" species="izuensis">
<emphasis id="B956EAF1FFEC565DFF4DFA70C3FDDBAD" box="[241,426,1478,1503]" italics="true" pageId="2" pageNumber="226">Leanira izuensis</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFE0DFA73C03EDBAD" author="Takahashi, K." box="[433,617,1477,1503]" pageId="2" pageNumber="226" refId="ref13450" refString="Takahashi, K. (1938) Polychaetous Annelids of Izu Peninsula 1. Polychaeta, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu Peninsula. Science Reports of the Tokyo Burnika Daigaku, Series B. 3,193 - 220." type="book" year="1938">Takahashi, 1938</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C224D60FFEC565DFD1DFA70C1F3DBAD" authority="Horst, 1917" authorityName="Horst" authorityYear="1917" box="[673,932,1477,1504]" class="Polychaeta" family="Sigalionidae" genus="Leanira" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="species" species="vulturis">
<emphasis id="B956EAF1FFEC565DFD1DFA70C142DBAD" box="[673,789,1477,1503]" italics="true" pageId="2" pageNumber="226">L. vulturis</emphasis>
<bibRefCitation id="EFB34B12FFEC565DFCA0FA70C1F3DBAD" author="Horst, R." box="[796,932,1477,1504]" pageId="2" pageNumber="226" pagination="1 - 192" refId="ref12589" refString="Horst, R. (1917) Polychaeta Errantia of the Siboga-Expedition. Part II. Aphroditidae and Chrysopetalidae. Siboga-Expeditie. Monography, 24 b, 1 - 192. [https: // www. biodiversitylibrary. org / page / 2219945]" type="journal article" year="1917">Horst, 1917</bibRefCitation>
</taxonomicName>
, possess a morphology that could agree with
<taxonomicName id="4C224D60FFEC565DFF2BFA5FC34FD871" authorityName="Willey" authorityYear="1905" box="[151,280,1513,1539]" class="Polychaeta" family="Sigalionidae" genus="Sthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="226" phylum="Annelida" rank="genus">
<emphasis id="B956EAF1FFEC565DFF2BFA5FC34FD871" box="[151,280,1513,1539]" italics="true" pageId="2" pageNumber="226">Sthenolepis</emphasis>
</taxonomicName>
, but also other known genera (see
<tableCitation id="C6A00358FFEC565DFD22FA5FC0A7D871" box="[670,752,1513,1539]" captionStart="TABLE 1" captionStartId="17.[151,235,463,488]" captionText="TABLE 1. Comparative table of the valid species of Sthenolepis Willey, 1905 (Read &amp; Fauchald 2024), including the geographical and bathymetrical distribution. The number of times in sizes relationships between wide and length is given by ‘×’." pageId="2" pageNumber="226">Table 1</tableCitation>
). Solving these issues is beyond the scope of this research.
</paragraph>
</subSubSection>
</treatment>
</document>

414
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<emphasis id="B9645F2B4178FF94A8BBBB8BFE5BF8D1" bold="true" box="[151,426,1789,1815]" pageId="1" pageNumber="457">
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<emphasis id="B9645F2B4178FF94A8BBBB8BFEB4F8D1" bold="true" box="[151,325,1789,1815]" italics="true" pageId="1" pageNumber="457">Ingenia major</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C30AD0B24178FF94A8BBBA57FE7EF8FD" box="[151,399,1825,1851]" pageId="1" pageNumber="457" type="description">
<paragraph id="8BAF83394178FF94A8BBBA57FE7EF8FD" blockId="1.[151,426,1789,1851]" box="[151,399,1825,1851]" pageId="1" pageNumber="457">
(
<figureCitation id="132B9FBC4178FF94A8B3BA57FED9F8FD" box="[159,296,1825,1851]" captionStart-0="FIGURE" captionStart-1="FIGURE 2" captionStartId-0="2.[152,169,1895,1921]" captionStartId-1="3.[152,255,1613,1637]" captionTargetBox-0="[177,1410,192,1857]" captionTargetBox-1="[164,1423,181,1589]" captionTargetId-0="figure-18@2.[164,1423,179,1870]" captionTargetId-1="figure-27@3.[164,1423,181,1589]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Ingenia major sp. nov. A. Lateral view of male anterior portion, showing cephalic setae, buccal cavity and amphidial fovea; B. Details of male head, showing cephalic setae and amphidial fovea; C. Lateral view of male posterior portion, showing spicules and gubernaculum; D. Spicules and gubernaculum; E. Lateral view of male entire body. Scales: A, B, C = 50 μm, E = 100 μm." captionText-1="FIGURE 2. Ingenia major sp. nov. A. Lateral view of male anterior end, showing amphidial fovea; B. Lateral view of male anterior end, showing buccal cavity; C. Lateral view of male gubernaculum region (arrow); D. Lateral view of male spicules region (arrow); E. Details of inner labial setae and outer labial setae; F. Details of spicules teeth-shaped structure. Scales: A, B, C, D = 20 μm, E = 10 μm, F = 5 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13757901" figureDoi-1="http://doi.org/10.5281/zenodo.13757905" httpUri-0="https://zenodo.org/record/13757901/files/figure.png" httpUri-1="https://zenodo.org/record/13757905/files/figure.png" pageId="1" pageNumber="457">Figures 1–2</figureCitation>
,
<tableCitation id="C692B6824178FF94A91FBA57FE75F8FD" box="[307,388,1825,1851]" captionStart="TABLE 1" captionStartId="4.[151,239,260,284]" captionText="TABLE 1. Individual measurements of Ingenia major sp. nov. and Linhystera filiformis sp. nov. (in µm except for de Man’s ratios)." pageId="1" pageNumber="457">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B24178FF94A8BBBA1FFDC7F808" pageId="1" pageNumber="457" type="materials_examined">
<paragraph id="8BAF83394178FF94A8BBBA1FFB79F845" blockId="1.[151,1436,1897,2032]" box="[151,1160,1897,1924]" pageId="1" pageNumber="457">
<emphasis id="B9645F2B4178FF94A8BBBA1FFEB4F845" bold="true" box="[151,325,1897,1923]" pageId="1" pageNumber="457">Type material.</emphasis>
<specimenCount id="9D1648B04178FF94A960BA1FFE46F845" box="[332,439,1897,1923]" count="1" pageId="1" pageNumber="457" type="male">One male</specimenCount>
was measured and studied.
<typeStatus id="54AB3D9B4178FF94AADEBA1FFCAEF845" box="[754,863,1897,1923]" pageId="1" pageNumber="457" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1648B04178FF94AB45BA1FFC70F845" box="[873,897,1897,1923]" count="1" pageId="1" pageNumber="457" type="male">♂</specimenCount>
1 on slide 19YD1-1-13.
</paragraph>
<paragraph id="8BAF83394178FF94A8EBBAFBFDC7F808" blockId="1.[151,1436,1897,2032]" pageId="1" pageNumber="457">
<materialsCitation id="3B7889644178FF94A8EBBAFBFE4BF80D" country="China" latitude="37.783333" location="Specimens" municipality="Yantai" pageId="1" pageNumber="457" specimenCount="1" stateProvince="Shandong" typeStatus="holotype">
<emphasis id="B9645F2B4178FF94A8EBBAFBFE07F861" bold="true" box="[199,502,1933,1959]" pageId="1" pageNumber="457">
<typeStatus id="54AB3D9B4178FF94A8EBBAFBFEF3F861" box="[199,258,1933,1959]" pageId="1" pageNumber="457">Type</typeStatus>
locality and habitat.
</emphasis>
<location id="8ECFD5E24178FF94A9D1BAFBFD87F861" LSID="urn:lsid:plazi:treatment:03B9322F4178FF90A8BBBB8BFE0DFE33:8ECFD5E24178FF94A9D1BAFBFD87F861" box="[509,630,1933,1959]" country="China" latitude="37.783333" municipality="Yantai" name="Specimens" pageId="1" pageNumber="457" stateProvince="Shandong">Specimens</location>
were obtained from intertidal sandy sediment at
<collectingMunicipality id="6BCB19434178FF94ACBBBAFBFB2EF861" box="[1175,1247,1933,1959]" pageId="1" pageNumber="457">Yantai</collectingMunicipality>
coast,
<collectingRegion id="49D44DDB4178FF94AD00BAFBFF0CF80D" country="China" name="Shandong" pageId="1" pageNumber="457">Shandong Province</collectingRegion>
,
<collectingCountry id="F307C3A94178FF94A925BAC7FEBEF80D" box="[265,335,1969,1995]" name="China" pageId="1" pageNumber="457">China</collectingCountry>
,
<geoCoordinate id="EE24E5FE4178FF94A976BAC7FE4BF80D" box="[346,442,1969,1995]" degrees="37" direction="north" minutes="47" orientation="latitude" pageId="1" pageNumber="457" precision="925" value="37.783333">37º47´N</geoCoordinate>
</materialsCitation>
, 121º48΄E.
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B24178FF94A8EBBAA3FC7AF829" box="[199,907,2005,2032]" pageId="1" pageNumber="457" type="etymology">
<paragraph id="8BAF83394178FF94A8EBBAA3FC7AF829" blockId="1.[151,1436,1897,2032]" box="[199,907,2005,2032]" pageId="1" pageNumber="457">
<emphasis id="B9645F2B4178FF94A8EBBAA3FEBFF829" bold="true" box="[199,334,2005,2031]" pageId="1" pageNumber="457">Etymology.</emphasis>
Species epithet
<taxonomicName id="4C10F8BA4178FF94AA28BAA0FDB6F829" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[516,583,2006,2031]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="species" species="major">
<emphasis id="B9645F2B4178FF94AA28BAA0FDB6F829" box="[516,583,2006,2031]" italics="true" pageId="1" pageNumber="457">major</emphasis>
</taxonomicName>
refers to the large body size.
</paragraph>
</subSubSection>
<caption id="DF6FD3B1417BFF97A8B4BA11FF1DF82B" ID-DOI="http://doi.org/10.5281/zenodo.13757901" ID-Zenodo-Dep="13757901" httpUri="https://zenodo.org/record/13757901/files/figure.png" pageId="2" pageNumber="458" startId="2.[152,169,1895,1921]" targetBox="[177,1410,192,1857]" targetPageId="2" targetType="figure">
<paragraph id="8BAF8339417BFF97A8B4BA11FF1DF82B" blockId="2.[151,1436,1895,2029]" pageId="2" pageNumber="458">
<emphasis id="B9645F2B417BFF97A8B4BA11FEE3F846" bold="true" box="[152,274,1895,1921]" pageId="2" pageNumber="458">FIGURE 1</emphasis>
.
<taxonomicName id="4C10F8BA417BFF97A930BA1FFE5AF846" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[284,427,1897,1920]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="2" pageNumber="458" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417BFF97A930BA1FFE5AF846" box="[284,427,1897,1920]" italics="true" pageId="2" pageNumber="458">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417BFF97A983BA1EFE0CF846" bold="true" box="[431,509,1896,1920]" pageId="2" pageNumber="458">
<taxonomicNameLabel id="A257E250417BFF97A983BA1EFE0CF846" box="[431,509,1896,1920]" pageId="2" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Lateral view of male anterior portion, showing cephalic setae, buccal cavity and amphidial fovea; B. Details of male head, showing cephalic setae and amphidial fovea; C. Lateral view of male posterior portion, showing spicules and gubernaculum; D. Spicules and gubernaculum; E. Lateral view of male entire body. Scales: A, B, C = 50 μm, E = 100 μm.
</paragraph>
</caption>
<caption id="DF6FD3B1417AFF96A8B4BB3BFDFDF914" ID-DOI="http://doi.org/10.5281/zenodo.13757905" ID-Zenodo-Dep="13757905" httpUri="https://zenodo.org/record/13757905/files/figure.png" pageId="3" pageNumber="459" startId="3.[152,255,1613,1637]" targetBox="[164,1423,181,1589]" targetPageId="3" targetType="figure">
<paragraph id="8BAF8339417AFF96A8B4BB3BFDFDF914" blockId="3.[151,1437,1613,1746]" pageId="3" pageNumber="459">
<emphasis id="B9645F2B417AFF96A8B4BB3BFEE2F9A3" bold="true" box="[152,275,1613,1637]" pageId="3" pageNumber="459">FIGURE 2</emphasis>
.
<taxonomicName id="4C10F8BA417AFF96A90DBB38FE42F9A3" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[289,435,1614,1637]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="3" pageNumber="459" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417AFF96A90DBB38FE42F9A3" box="[289,435,1614,1637]" italics="true" pageId="3" pageNumber="459">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417AFF96A996BB3BFDFAF9A3" bold="true" box="[442,523,1613,1637]" pageId="3" pageNumber="459">
<taxonomicNameLabel id="A257E250417AFF96A996BB3BFDFAF9A3" box="[442,523,1613,1637]" pageId="3" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Lateral view of male anterior end, showing amphidial fovea; B. Lateral view of male anterior end, showing buccal cavity; C. Lateral view of male gubernaculum region (arrow); D. Lateral view of male spicules region (arrow); E. Details of inner labial setae and outer labial setae; F. Details of spicules teeth-shaped structure. Scales: A, B, C, D = 20 μm, E = 10 μm, F = 5 μm.
</paragraph>
</caption>
<subSubSection id="C30AD0B2417AFF90A8EBBA55FE0DFE33" lastPageId="5" lastPageNumber="461" pageId="3" pageNumber="459" type="description">
<paragraph id="8BAF8339417AFF96A8EBBA55FEA8F8FB" blockId="3.[151,1437,1827,2034]" box="[199,345,1827,1853]" pageId="3" pageNumber="459">
<emphasis id="B9645F2B417AFF96A8EBBA55FEA8F8FB" bold="true" box="[199,345,1827,1853]" pageId="3" pageNumber="459">Description.</emphasis>
</paragraph>
<paragraph id="8BAF8339417AFF91A8EBBA31FCB6FF13" blockId="3.[151,1437,1827,2034]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="460" pageId="3" pageNumber="459">
<emphasis id="B9645F2B417AFF96A8EBBA31FEFAF8A7" bold="true" box="[199,267,1863,1889]" pageId="3" pageNumber="459">Male.</emphasis>
Body columnar, very long and thick. Cuticle striated. Anterior sensilla arranged in two separate circles: six inner labial setae in one circle, six outer labial setae and four cephalic setae arranged in another circle. Outer labial setae approximately equal to head diameter in length, and each setae divided into four segments, with apical end swollen. Buccal cavity large and barrel-shaped, with one large dorsal tooth, two small subventral teeth. Amphidial fovea unispiral-shaped, 20% of corresponding body diameter (c.b.d.) in diameter, and 32 µm from anterior end. Pharynx cylindrical and slight swollen proximally. Nerve ring located slightly anterior to 1/3 pharynx length (31% of pharynx length). Secretory-excretory system not observed.
</paragraph>
<caption id="DF6FD3B1417DFF91A8BBBC72FEDDFE86" pageId="4" pageNumber="460" startId="4.[151,239,260,284]" targetType="table">
<paragraph id="8BAF8339417DFF91A8BBBC72FEDDFE86" blockId="4.[151,1436,260,321]" pageId="4" pageNumber="460">
<emphasis id="B9645F2B417DFF91A8BBBC72FEFDFEDA" bold="true" box="[151,268,260,284]" pageId="4" pageNumber="460">TABLE 1.</emphasis>
Individual measurements of
<taxonomicName id="4C10F8BA417DFF91AA65BC73FD12FEDA" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[585,739,261,284]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="4" pageNumber="460" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417DFF91AA65BC73FD12FEDA" box="[585,739,261,284]" italics="true" pageId="4" pageNumber="460">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417DFF91AAC7BC73FCB3FEDB" bold="true" box="[747,834,261,285]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A257E250417DFF91AAC7BC73FCB3FEDB" box="[747,834,261,285]" pageId="4" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C10F8BA417DFF91AB54BC72FBA4FEDA" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[888,1109,260,284]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="4" pageNumber="460" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B417DFF91AB54BC72FBA4FEDA" box="[888,1109,260,284]" italics="true" pageId="4" pageNumber="460">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417DFF91AC71BC73FB45FEDB" bold="true" box="[1117,1204,261,285]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A257E250417DFF91AC71BC73FB45FEDB" box="[1117,1204,261,285]" pageId="4" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in µm except for de Man’s ratios).
</paragraph>
</caption>
<paragraph id="8BAF8339417DFF91A91FBC21FA77F9FA" pageId="4" pageNumber="460">
<table id="F9107199417D006AA891BC25FA77F9FA" box="[189,1414,339,1596]" gridcols="5" gridrows="26" pageId="4" pageNumber="460">
<tr id="3520817B417D006AA891BC25FA77FEA9" box="[189,1414,339,367]" gridrow="0" pageId="4" pageNumber="460" rowspan-2="1">
<th id="76F1E807417D006AA891BC25FE04FEA9" box="[189,501,339,367]" gridcol="0" gridrow="0" pageId="4" pageNumber="460">Species</th>
<th id="76F1E807417D006AAA04BC25FCFEFEA9" box="[552,783,339,367]" gridcol="1" gridrow="0" pageId="4" pageNumber="460">
<taxonomicName id="4C10F8BA417DFF91AA04BC25FD48FEAC" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[552,697,339,362]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="4" pageNumber="460" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417DFF91AA04BC25FD48FEAC" box="[552,697,339,362]" italics="true" pageId="4" pageNumber="460">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417DFF91AA92BC25FCFEFEAD" bold="true" box="[702,783,339,363]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A257E250417DFF91AA92BC25FCFEFEAD" box="[702,783,339,363]" pageId="4" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76F1E807417D006AAB9ABC25FA77FEA9" box="[950,1414,339,367]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="4" pageNumber="460">
<taxonomicName id="4C10F8BA417DFF91ABE7BC25FB6BFEAD" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[971,1178,339,363]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="4" pageNumber="460" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B417DFF91ABE7BC25FB6BFEAD" box="[971,1178,339,363]" italics="true" pageId="4" pageNumber="460">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417DFF91AC8DBC25FB00FEAD" bold="true" box="[1185,1265,339,363]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A257E250417DFF91AC8DBC25FB00FEAD" box="[1185,1265,339,363]" pageId="4" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="3520817B417D006AA891BC0BFA77FE53" box="[189,1414,381,405]" gridrow="1" pageId="4" pageNumber="460" rowspan-0="1" rowspan-2="1">
<td id="76F1E807417D006AAA04BC0BFCFEFE53" box="[552,783,381,405]" gridcol="1" gridrow="1" pageId="4" pageNumber="460">Male</td>
<td id="76F1E807417D006AAB9ABC0BFB77FE53" box="[950,1158,381,405]" gridcol="3" gridrow="1" pageId="4" pageNumber="460">Males</td>
<td id="76F1E807417D006AAC9ABC0BFA77FE53" box="[1206,1414,381,405]" gridcol="4" gridrow="1" pageId="4" pageNumber="460">Females</td>
</tr>
<tr id="3520817B417D006AA891BCD3FA77FE78" box="[189,1414,421,446]" gridrow="2" pageId="4" pageNumber="460" rowspan-0="1">
<td id="76F1E807417D006AAA04BCD3FCFEFE78" box="[552,783,421,446]" gridcol="1" gridrow="2" pageId="4" pageNumber="460">Holotype</td>
<td id="76F1E807417D006AAB03BCD3FC7CFE78" box="[815,909,421,446]" gridcol="2" gridrow="2" pageId="4" pageNumber="460">Holotype</td>
<td id="76F1E807417D006AAB9ABCD3FB77FE78" box="[950,1158,421,446]" gridcol="3" gridrow="2" pageId="4" pageNumber="460">3 Paratypes</td>
<td id="76F1E807417D006AAC9ABCD3FA77FE78" box="[1206,1414,421,446]" gridcol="4" gridrow="2" pageId="4" pageNumber="460">3 Paratypes</td>
</tr>
<tr id="3520817B417D006AA891BCA4FA77FE2C" box="[189,1414,466,490]" gridrow="3" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BCA4FE04FE2C" box="[189,501,466,490]" gridcol="0" gridrow="3" pageId="4" pageNumber="460">Total body length</th>
<td id="76F1E807417D006AAA04BCA4FCFEFE2C" box="[552,783,466,490]" gridcol="1" gridrow="3" pageId="4" pageNumber="460">3470</td>
<td id="76F1E807417D006AAB03BCA4FC7CFE2C" box="[815,909,466,490]" gridcol="2" gridrow="3" pageId="4" pageNumber="460">829</td>
<td id="76F1E807417D006AAB9ABCA4FB77FE2C" box="[950,1158,466,490]" gridcol="3" gridrow="3" pageId="4" pageNumber="460">1011±44(962–1048)</td>
<td id="76F1E807417D006AAC9ABCA4FA77FE2C" box="[1206,1414,466,490]" gridcol="4" gridrow="3" pageId="4" pageNumber="460">922±34(895–960)</td>
</tr>
<tr id="3520817B417D006AA891BF74FA77FDDC" box="[189,1414,514,538]" gridrow="4" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BF74FE04FDDC" box="[189,501,514,538]" gridcol="0" gridrow="4" pageId="4" pageNumber="460">Maximum body diameter</th>
<td id="76F1E807417D006AAA04BF74FCFEFDDC" box="[552,783,514,538]" gridcol="1" gridrow="4" pageId="4" pageNumber="460">36</td>
<td id="76F1E807417D006AAB03BF74FC7CFDDC" box="[815,909,514,538]" gridcol="2" gridrow="4" pageId="4" pageNumber="460">18</td>
<td id="76F1E807417D006AAB9ABF74FB77FDDC" box="[950,1158,514,538]" gridcol="3" gridrow="4" pageId="4" pageNumber="460">19±1(19–20)</td>
<td id="76F1E807417D006AAC9ABF74FA77FDDC" box="[1206,1414,514,538]" gridcol="4" gridrow="4" pageId="4" pageNumber="460">22±1(21–23)</td>
</tr>
<tr id="3520817B417D006AA891BF47FA77FD8C" box="[189,1414,561,586]" gridrow="5" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BF47FE04FD8C" box="[189,501,561,586]" gridcol="0" gridrow="5" pageId="4" pageNumber="460">Head diameter</th>
<td id="76F1E807417D006AAA04BF47FCFEFD8C" box="[552,783,561,586]" gridcol="1" gridrow="5" pageId="4" pageNumber="460">31</td>
<td id="76F1E807417D006AAB03BF47FC7CFD8C" box="[815,909,561,586]" gridcol="2" gridrow="5" pageId="4" pageNumber="460">6</td>
<td id="76F1E807417D006AAB9ABF47FB77FD8C" box="[950,1158,561,586]" gridcol="3" gridrow="5" pageId="4" pageNumber="460">7±0(7–7)</td>
<td id="76F1E807417D006AAC9ABF47FA77FD8C" box="[1206,1414,561,586]" gridcol="4" gridrow="5" pageId="4" pageNumber="460">6±0(6–6)</td>
</tr>
<tr id="3520817B417D006AA891BF17FA77FDBF" box="[189,1414,609,633]" gridrow="6" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BF17FE04FDBF" box="[189,501,609,633]" gridcol="0" gridrow="6" pageId="4" pageNumber="460">Length of inner labial setae</th>
<td id="76F1E807417D006AAA04BF17FCFEFDBF" box="[552,783,609,633]" gridcol="1" gridrow="6" pageId="4" pageNumber="460">11</td>
<td id="76F1E807417D006AAB03BF17FC7CFDBF" box="[815,909,609,633]" gridcol="2" gridrow="6" pageId="4" pageNumber="460">5</td>
<td id="76F1E807417D006AAB9ABF17FB77FDBF" box="[950,1158,609,633]" gridcol="3" gridrow="6" pageId="4" pageNumber="460">4±0(4–4)</td>
<td id="76F1E807417D006AAC9ABF17FA77FDBF" box="[1206,1414,609,633]" gridcol="4" gridrow="6" pageId="4" pageNumber="460">4±1(4–5)</td>
</tr>
<tr id="3520817B417D006AA891BFE7FA77FD6F" box="[189,1414,657,681]" gridrow="7" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BFE7FE04FD6F" box="[189,501,657,681]" gridcol="0" gridrow="7" pageId="4" pageNumber="460">Length of outer labial setae</th>
<td id="76F1E807417D006AAA04BFE7FCFEFD6F" box="[552,783,657,681]" gridcol="1" gridrow="7" pageId="4" pageNumber="460">29</td>
<td id="76F1E807417D006AAB03BFE7FC7CFD6F" box="[815,909,657,681]" gridcol="2" gridrow="7" pageId="4" pageNumber="460">5</td>
<td id="76F1E807417D006AAB9ABFE7FB77FD6F" box="[950,1158,657,681]" gridcol="3" gridrow="7" pageId="4" pageNumber="460">4±0(4–4)</td>
<td id="76F1E807417D006AAC9ABFE7FA77FD6F" box="[1206,1414,657,681]" gridcol="4" gridrow="7" pageId="4" pageNumber="460">4±1(4–5)</td>
</tr>
<tr id="3520817B417D006AA891BFB6FA77FD1F" box="[189,1414,704,729]" gridrow="8" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BFB6FE04FD1F" box="[189,501,704,729]" gridcol="0" gridrow="8" pageId="4" pageNumber="460">Length of cephalic setae</th>
<td id="76F1E807417D006AAA04BFB6FCFEFD1F" box="[552,783,704,729]" gridcol="1" gridrow="8" pageId="4" pageNumber="460">16</td>
<td id="76F1E807417D006AAB03BFB6FC7CFD1F" box="[815,909,704,729]" gridcol="2" gridrow="8" pageId="4" pageNumber="460">5</td>
<td id="76F1E807417D006AAB9ABFB6FB77FD1F" box="[950,1158,704,729]" gridcol="3" gridrow="8" pageId="4" pageNumber="460">4±0(4–4)</td>
<td id="76F1E807417D006AAC9ABFB6FA77FD1F" box="[1206,1414,704,729]" gridcol="4" gridrow="8" pageId="4" pageNumber="460">4±1(4–5)</td>
</tr>
<tr id="3520817B417D006AA891BF86FA77FCCF" box="[189,1414,752,777]" gridrow="9" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BF86FE04FCCF" box="[189,501,752,777]" gridcol="0" gridrow="9" pageId="4" pageNumber="460">Length of buccal cavity</th>
<td id="76F1E807417D006AAA04BF86FCFEFCCF" box="[552,783,752,777]" gridcol="1" gridrow="9" pageId="4" pageNumber="460">37</td>
<td id="76F1E807417D006AAB03BF86FC7CFCCF" box="[815,909,752,777]" gridcol="2" gridrow="9" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB9ABF86FB77FCCF" box="[950,1158,752,777]" gridcol="3" gridrow="9" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAC9ABF86FA77FCCF" box="[1206,1414,752,777]" gridcol="4" gridrow="9" pageId="4" pageNumber="460">-</td>
</tr>
<tr id="3520817B417D006AA891BE56FA77FCFE" box="[189,1414,800,824]" gridrow="10" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BE56FE04FCFE" box="[189,501,800,824]" gridcol="0" gridrow="10" pageId="4" pageNumber="460">Diameter of amphidial fovea</th>
<td id="76F1E807417D006AAA04BE56FCFEFCFE" box="[552,783,800,824]" gridcol="1" gridrow="10" pageId="4" pageNumber="460">6</td>
<td id="76F1E807417D006AAB03BE56FC7CFCFE" box="[815,909,800,824]" gridcol="2" gridrow="10" pageId="4" pageNumber="460">4</td>
<td id="76F1E807417D006AAB9ABE56FB77FCFE" box="[950,1158,800,824]" gridcol="3" gridrow="10" pageId="4" pageNumber="460">4±1(4–5)</td>
<td id="76F1E807417D006AAC9ABE56FA77FCFE" box="[1206,1414,800,824]" gridcol="4" gridrow="10" pageId="4" pageNumber="460">4±1(3–4)</td>
</tr>
<tr id="3520817B417D006AA891BE26FA77FCAE" box="[189,1414,848,872]" gridrow="11" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BE26FE04FCAE" box="[189,501,848,872]" gridcol="0" gridrow="11" pageId="4" pageNumber="460">Amphidial fovea from anterior</th>
<td id="76F1E807417D006AAA04BE26FCFEFCAE" box="[552,783,848,872]" gridcol="1" gridrow="11" pageId="4" pageNumber="460">32</td>
<td id="76F1E807417D006AAB03BE26FC7CFCAE" box="[815,909,848,872]" gridcol="2" gridrow="11" pageId="4" pageNumber="460">7</td>
<td id="76F1E807417D006AAB9ABE26FB77FCAE" box="[950,1158,848,872]" gridcol="3" gridrow="11" pageId="4" pageNumber="460">7±1(7–8)</td>
<td id="76F1E807417D006AAC9ABE26FA77FCAE" box="[1206,1414,848,872]" gridcol="4" gridrow="11" pageId="4" pageNumber="460">7±1(6–8)</td>
</tr>
<tr id="3520817B417D006AA891BE09FA77FC5E" box="[189,1414,895,920]" gridrow="12" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BE09FE04FC5E" box="[189,501,895,920]" gridcol="0" gridrow="12" pageId="4" pageNumber="460">Nerve ring from anterior</th>
<td id="76F1E807417D006AAA04BE09FCFEFC5E" box="[552,783,895,920]" gridcol="1" gridrow="12" pageId="4" pageNumber="460">144</td>
<td id="76F1E807417D006AAB03BE09FC7CFC5E" box="[815,909,895,920]" gridcol="2" gridrow="12" pageId="4" pageNumber="460">54</td>
<td id="76F1E807417D006AAB9ABE09FB77FC5E" box="[950,1158,895,920]" gridcol="3" gridrow="12" pageId="4" pageNumber="460">54±2(53–56)</td>
<td id="76F1E807417D006AAC9ABE09FA77FC5E" box="[1206,1414,895,920]" gridcol="4" gridrow="12" pageId="4" pageNumber="460">56±3(54–59)</td>
</tr>
<tr id="3520817B417D006AA891BED9FA77FC01" box="[189,1414,943,967]" gridrow="13" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BED9FE04FC01" box="[189,501,943,967]" gridcol="0" gridrow="13" pageId="4" pageNumber="460">Body diameter at nerve ring</th>
<td id="76F1E807417D006AAA04BED9FCFEFC01" box="[552,783,943,967]" gridcol="1" gridrow="13" pageId="4" pageNumber="460">34</td>
<td id="76F1E807417D006AAB03BED9FC7CFC01" box="[815,909,943,967]" gridcol="2" gridrow="13" pageId="4" pageNumber="460">14</td>
<td id="76F1E807417D006AAB9ABED9FB77FC01" box="[950,1158,943,967]" gridcol="3" gridrow="13" pageId="4" pageNumber="460">14±1(13–14)</td>
<td id="76F1E807417D006AAC9ABED9FA77FC01" box="[1206,1414,943,967]" gridcol="4" gridrow="13" pageId="4" pageNumber="460">14±1(13–14)</td>
</tr>
<tr id="3520817B417D006AA891BEA9FA77FC31" box="[189,1414,991,1015]" gridrow="14" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891BEA9FE04FC31" box="[189,501,991,1015]" gridcol="0" gridrow="14" pageId="4" pageNumber="460">Pharynx length</th>
<td id="76F1E807417D006AAA04BEA9FCFEFC31" box="[552,783,991,1015]" gridcol="1" gridrow="14" pageId="4" pageNumber="460">464</td>
<td id="76F1E807417D006AAB03BEA9FC7CFC31" box="[815,909,991,1015]" gridcol="2" gridrow="14" pageId="4" pageNumber="460">79</td>
<td id="76F1E807417D006AAB9ABEA9FB77FC31" box="[950,1158,991,1015]" gridcol="3" gridrow="14" pageId="4" pageNumber="460">101±2(99–103)</td>
<td id="76F1E807417D006AAC9ABEA9FA77FC31" box="[1206,1414,991,1015]" gridcol="4" gridrow="14" pageId="4" pageNumber="460">100±4(96–104)</td>
</tr>
<tr id="3520817B417D006AA891B97DFA77FB81" box="[189,1414,1035,1095]" gridrow="15" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B97DFE04FB81" box="[189,501,1035,1095]" gridcol="0" gridrow="15" pageId="4" pageNumber="460">Body diameter at the base of pharynx</th>
<td id="76F1E807417D006AAA04B97DFCFEFB81" box="[552,783,1035,1095]" gridcol="1" gridrow="15" pageId="4" pageNumber="460">35</td>
<td id="76F1E807417D006AAB03B97DFC7CFB81" box="[815,909,1035,1095]" gridcol="2" gridrow="15" pageId="4" pageNumber="460">15</td>
<td id="76F1E807417D006AAB9AB97DFB77FB81" box="[950,1158,1035,1095]" gridcol="3" gridrow="15" pageId="4" pageNumber="460">15±1(14–16)</td>
<td id="76F1E807417D006AAC9AB97DFA77FB81" box="[1206,1414,1035,1095]" gridcol="4" gridrow="15" pageId="4" pageNumber="460">15±1(15–16)</td>
</tr>
<tr id="3520817B417D006AA891B92DFA77FBB5" box="[189,1414,1115,1139]" gridrow="16" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B92DFE04FBB5" box="[189,501,1115,1139]" gridcol="0" gridrow="16" pageId="4" pageNumber="460">Spicules length along arc</th>
<td id="76F1E807417D006AAA04B92DFCFEFBB5" box="[552,783,1115,1139]" gridcol="1" gridrow="16" pageId="4" pageNumber="460">39</td>
<td id="76F1E807417D006AAB03B92DFC7CFBB5" box="[815,909,1115,1139]" gridcol="2" gridrow="16" pageId="4" pageNumber="460">19</td>
<td id="76F1E807417D006AAB9AB92DFB77FBB5" box="[950,1158,1115,1139]" gridcol="3" gridrow="16" pageId="4" pageNumber="460">21±0(21–21)</td>
<td id="76F1E807417D006AAC9AB92DFA77FBB5" box="[1206,1414,1115,1139]" gridcol="4" gridrow="16" pageId="4" pageNumber="460">-</td>
</tr>
<tr id="3520817B417D006AA891B9FDFA77FB65" box="[189,1414,1163,1187]" gridrow="17" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B9FDFE04FB65" box="[189,501,1163,1187]" gridcol="0" gridrow="17" pageId="4" pageNumber="460">Gubernaculum length</th>
<td id="76F1E807417D006AAA04B9FDFCFEFB65" box="[552,783,1163,1187]" gridcol="1" gridrow="17" pageId="4" pageNumber="460">30</td>
<td id="76F1E807417D006AAB03B9FDFC7CFB65" box="[815,909,1163,1187]" gridcol="2" gridrow="17" pageId="4" pageNumber="460">6</td>
<td id="76F1E807417D006AAB9AB9FDFB77FB65" box="[950,1158,1163,1187]" gridcol="3" gridrow="17" pageId="4" pageNumber="460">7±1(6–7)</td>
<td id="76F1E807417D006AAC9AB9FDFA77FB65" box="[1206,1414,1163,1187]" gridcol="4" gridrow="17" pageId="4" pageNumber="460">-</td>
</tr>
<tr id="3520817B417D006AA891B9CFFA77FB14" box="[189,1414,1209,1234]" gridrow="18" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B9CFFE04FB14" box="[189,501,1209,1234]" gridcol="0" gridrow="18" pageId="4" pageNumber="460">Cloacal/Anal body diameter</th>
<td id="76F1E807417D006AAA04B9CFFCFEFB14" box="[552,783,1209,1234]" gridcol="1" gridrow="18" pageId="4" pageNumber="460">32</td>
<td id="76F1E807417D006AAB03B9CFFC7CFB14" box="[815,909,1209,1234]" gridcol="2" gridrow="18" pageId="4" pageNumber="460">12</td>
<td id="76F1E807417D006AAB9AB9CFFB77FB14" box="[950,1158,1209,1234]" gridcol="3" gridrow="18" pageId="4" pageNumber="460">14±1(14–15)</td>
<td id="76F1E807417D006AAC9AB9CFFA77FB14" box="[1206,1414,1209,1234]" gridcol="4" gridrow="18" pageId="4" pageNumber="460">13±1(12–14)</td>
</tr>
<tr id="3520817B417D006AA891B991FA77FB39" box="[189,1414,1255,1279]" gridrow="19" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B991FE04FB39" box="[189,501,1255,1279]" gridcol="0" gridrow="19" pageId="4" pageNumber="460">Vulva from anterior</th>
<td id="76F1E807417D006AAA04B991FCFEFB39" box="[552,783,1255,1279]" gridcol="1" gridrow="19" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB03B991FC7CFB39" box="[815,909,1255,1279]" gridcol="2" gridrow="19" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB9AB991FB77FB39" box="[950,1158,1255,1279]" gridcol="3" gridrow="19" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAC9AB991FA77FB39" box="[1206,1414,1255,1279]" gridcol="4" gridrow="19" pageId="4" pageNumber="460">476±15(460–489)</td>
</tr>
<tr id="3520817B417D006AA891B863FA77FAEB" box="[189,1414,1301,1325]" gridrow="20" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B863FE04FAEB" box="[189,501,1301,1325]" gridcol="0" gridrow="20" pageId="4" pageNumber="460">Body diameter at vulva</th>
<td id="76F1E807417D006AAA04B863FCFEFAEB" box="[552,783,1301,1325]" gridcol="1" gridrow="20" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB03B863FC7CFAEB" box="[815,909,1301,1325]" gridcol="2" gridrow="20" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB9AB863FB77FAEB" box="[950,1158,1301,1325]" gridcol="3" gridrow="20" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAC9AB863FA77FAEB" box="[1206,1414,1301,1325]" gridcol="4" gridrow="20" pageId="4" pageNumber="460">20±3(18–23)</td>
</tr>
<tr id="3520817B417D006AA891B834FA77FA9C" box="[189,1414,1346,1370]" gridrow="21" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B834FE04FA9C" box="[189,501,1346,1370]" gridcol="0" gridrow="21" pageId="4" pageNumber="460">V%</th>
<td id="76F1E807417D006AAA04B834FCFEFA9C" box="[552,783,1346,1370]" gridcol="1" gridrow="21" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB03B834FC7CFA9C" box="[815,909,1346,1370]" gridcol="2" gridrow="21" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAB9AB834FB77FA9C" box="[950,1158,1346,1370]" gridcol="3" gridrow="21" pageId="4" pageNumber="460">-</td>
<td id="76F1E807417D006AAC9AB834FA77FA9C" box="[1206,1414,1346,1370]" gridcol="4" gridrow="21" pageId="4" pageNumber="460">51.7±2.6(49.8–54.6)</td>
</tr>
<tr id="3520817B417D006AA891B807FA77FA4F" box="[189,1414,1393,1417]" gridrow="22" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B807FE04FA4F" box="[189,501,1393,1417]" gridcol="0" gridrow="22" pageId="4" pageNumber="460">Tail length</th>
<td id="76F1E807417D006AAA04B807FCFEFA4F" box="[552,783,1393,1417]" gridcol="1" gridrow="22" pageId="4" pageNumber="460">82</td>
<td id="76F1E807417D006AAB03B807FC7CFA4F" box="[815,909,1393,1417]" gridcol="2" gridrow="22" pageId="4" pageNumber="460">187</td>
<td id="76F1E807417D006AAB9AB807FB77FA4F" box="[950,1158,1393,1417]" gridcol="3" gridrow="22" pageId="4" pageNumber="460">205±14(189–216)</td>
<td id="76F1E807417D006AAC9AB807FA77FA4F" box="[1206,1414,1393,1417]" gridcol="4" gridrow="22" pageId="4" pageNumber="460">205±44(154–233)</td>
</tr>
<tr id="3520817B417D006AA891B8E9FA77FA7E" box="[189,1414,1439,1464]" gridrow="23" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B8E9FE04FA7E" box="[189,501,1439,1464]" gridcol="0" gridrow="23" pageId="4" pageNumber="460">a</th>
<td id="76F1E807417D006AAA04B8E9FCFEFA7E" box="[552,783,1439,1464]" gridcol="1" gridrow="23" pageId="4" pageNumber="460">96.4</td>
<td id="76F1E807417D006AAB03B8E9FC7CFA7E" box="[815,909,1439,1464]" gridcol="2" gridrow="23" pageId="4" pageNumber="460">46.1</td>
<td id="76F1E807417D006AAB9AB8E9FB77FA7E" box="[950,1158,1439,1464]" gridcol="3" gridrow="23" pageId="4" pageNumber="460">52.3±1.7(50.6–53.9)</td>
<td id="76F1E807417D006AAC9AB8E9FA77FA7E" box="[1206,1414,1439,1464]" gridcol="4" gridrow="23" pageId="4" pageNumber="460">42.3±0.8(41.7–43.3)</td>
</tr>
<tr id="3520817B417D006AA891B8B8FA77FA20" box="[189,1414,1486,1510]" gridrow="24" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B8B8FE04FA20" box="[189,501,1486,1510]" gridcol="0" gridrow="24" pageId="4" pageNumber="460">b</th>
<td id="76F1E807417D006AAA04B8B8FCFEFA20" box="[552,783,1486,1510]" gridcol="1" gridrow="24" pageId="4" pageNumber="460">7.5</td>
<td id="76F1E807417D006AAB03B8B8FC7CFA20" box="[815,909,1486,1510]" gridcol="2" gridrow="24" pageId="4" pageNumber="460">10.4</td>
<td id="76F1E807417D006AAB9AB8B8FB77FA20" box="[950,1158,1486,1510]" gridcol="3" gridrow="24" pageId="4" pageNumber="460">10.0±0.7(9.3–10.6)</td>
<td id="76F1E807417D006AAC9AB8B8FA77FA20" box="[1206,1414,1486,1510]" gridcol="4" gridrow="24" pageId="4" pageNumber="460">9.2±0.1(9.1–9.3)</td>
</tr>
<tr id="3520817B417D006AA891B88CFA77F9FA" box="[189,1414,1530,1596]" gridrow="25" pageId="4" pageNumber="460">
<th id="76F1E807417D006AA891B88CFE04F9FA" box="[189,501,1530,1596]" gridcol="0" gridrow="25" pageId="4" pageNumber="460">c c’</th>
<td id="76F1E807417D006AAA04B88CFCFEF9FA" box="[552,783,1530,1596]" gridcol="1" gridrow="25" pageId="4" pageNumber="460">42.3 2.6</td>
<td id="76F1E807417D006AAB03B88CFC7CF9FA" box="[815,909,1530,1596]" gridcol="2" gridrow="25" pageId="4" pageNumber="460">4.4 15.6</td>
<td id="76F1E807417D006AAB9AB88CFB77F9FA" box="[950,1158,1530,1596]" gridcol="3" gridrow="25" pageId="4" pageNumber="460">4.9±0.2(4.7–5.1) 14.3±0.8(13.5–15.0)</td>
<td id="76F1E807417D006AAC9AB88CFA77F9FA" box="[1206,1414,1530,1596]" gridcol="4" gridrow="25" pageId="4" pageNumber="460">4.6±1.0(4.0–5.8) 16.1±3.1(12.8–18.9)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF8339417DFF91A8EBBBF9FD3DF8FF" blockId="4.[151,1437,1679,2029]" pageId="4" pageNumber="460">Reproductive system with single and outstretched testis, located on the right of intestine. Spicules thin and almost straight, 1.2 times of cloacal body diameter in length, with a pair of ventro-laterally directed teeth (8 µm in length) hooked distally. Gubernaculum kidney-shaped with triangular cuticularized projection, precloacal supplements absent. Tail conical, 2.6 times of cloacal body diameter, gradually tapering with swollen terminus. Three caudal glands in a line. Caudal setae absent.</paragraph>
<paragraph id="8BAF8339417DFF91A8EBBA32FE40F89B" blockId="4.[151,1437,1679,2029]" box="[199,433,1859,1886]" pageId="4" pageNumber="460">
<emphasis id="B9645F2B417DFF91A8EBBA32FEC1F898" bold="true" box="[199,304,1860,1886]" pageId="4" pageNumber="460">Females.</emphasis>
Not found.
</paragraph>
<paragraph id="8BAF8339417DFF91A8EBBA11FB78F82B" blockId="4.[151,1437,1679,2029]" pageId="4" pageNumber="460">
<emphasis id="B9645F2B417DFF91A8EBBA11FD59F847" bold="true" box="[199,680,1895,1922]" pageId="4" pageNumber="460">Differentiation diagnosis and discussion.</emphasis>
<taxonomicName id="4C10F8BA417DFF91AA82BA1EFCBCF847" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[686,845,1896,1921]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="4" pageNumber="460" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417DFF91AA82BA1EFCBCF847" box="[686,845,1896,1921]" italics="true" pageId="4" pageNumber="460">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417DFF91AB78BA1EFC5CF844" bold="true" box="[852,941,1896,1922]" pageId="4" pageNumber="460">
<taxonomicNameLabel id="A257E250417DFF91AB78BA1EFC5CF844" box="[852,941,1896,1922]" pageId="4" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by body large and thick, six outer labial setae articulated, amphidial fovea small and unispiral, buccal cavity large and barrel-shaped with three teeth, spicules thin and almost straight with a pair of ventro-laterally directed teeth hooked distally, gubernaculum kidney-shaped with triangular cuticularized projection, tail conical with swollen terminus.
</paragraph>
<paragraph id="8BAF8339417CFF90A8EBBDE1FE0DFE33" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="461">
Until the moment specimens of
<taxonomicName id="4C10F8BA417CFF90AA03BDE1FD74FF76" authorityName="Gerlach" authorityYear="1957" box="[559,645,151,176]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="5" pageNumber="461" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B417CFF90AA03BDE1FD74FF76" box="[559,645,151,176]" italics="true" pageId="5" pageNumber="461">Ingenia</emphasis>
</taxonomicName>
are rare and here we compare the only specimen, a female, of
<taxonomicName id="4C10F8BA417CFF90AD6BBDE1FF0DFF13" authorityName="Gerlach" authorityYear="1957" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="5" pageNumber="461" phylum="Nematoda" rank="species" species="mirabilis">
<emphasis id="B9645F2B417CFF90AD6BBDE1FF0DFF13" italics="true" pageId="5" pageNumber="461">Ingenia mirabilis</emphasis>
</taxonomicName>
with the only specimen, in this case a single male, of the new species.
<taxonomicName id="4C10F8BA417CFF90AC3FBDCDFB43FF12" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[1043,1202,187,212]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="5" pageNumber="461" phylum="Nematoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B9645F2B417CFF90AC3FBDCDFB43FF12" box="[1043,1202,187,212]" italics="true" pageId="5" pageNumber="461">Ingenia major</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417CFF90AC95BDCDFAE3FF13" bold="true" box="[1209,1298,187,213]" pageId="5" pageNumber="461">
<taxonomicNameLabel id="A257E250417CFF90AC95BDCDFAE3FF13" box="[1209,1298,187,213]" pageId="5" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C10F8BA417CFF90A8BBBDA9FEE5FF3F" authorityName="Gerlach" authorityYear="1957" box="[151,276,223,249]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="5" pageNumber="461" phylum="Nematoda" rank="species" species="mirabilis">
<emphasis id="B9645F2B417CFF90A8BBBDA9FEE5FF3F" box="[151,276,223,249]" italics="true" pageId="5" pageNumber="461">I. mirabilis</emphasis>
</taxonomicName>
in the articulated outer labial setae, large and barrel-shaped buccal cavity with three teeth, slender and proximal swollen pharynx. The new species significantly differs from
<taxonomicName id="4C10F8BA417CFF90AB85BC75FBD4FEDB" authorityName="Gerlach" authorityYear="1957" box="[937,1061,259,285]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="5" pageNumber="461" phylum="Nematoda" rank="species" species="mirabilis">
<emphasis id="B9645F2B417CFF90AB85BC75FBD4FEDB" box="[937,1061,259,285]" italics="true" pageId="5" pageNumber="461">I. mirabilis</emphasis>
</taxonomicName>
in longer body size (3470 µm vs. 1700 µm), slender body shape (36 µm vs. 52 µm in maximum body diameter), shorter inner labial setae length (11 µm vs. 18 µm), shorter outer labial setae length (29 µm vs. 45–50 µm), more sections of outer labial setae (four parts vs. two or three parts), smaller amphidial fovea size (6 µm vs. 10 µm in diameter), amphidial fovea located closer to the anterior end of the body (32 µm, at level of buccal cavity vs. 60 µm from anterior end, posterior to buccal cavity), higher de Man “a” ratio (96.4 vs. 33) and de Man “c” ratio (42.3 vs. 18.5), as well as different tail shape (conical vs. conico-cylindrical).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4A1C6306391B98E992E247F595F41403">Description of two new nematode species: Ingenia major sp. nov. (Tripyloididae) and Linhystera filiformis sp. nov. (Xyalidae) from an intertidal beach of the Yellow China Sea</mods:title>
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<paragraph id="8BAF83394178FF94A8BBB95FFDF2FB85" blockId="1.[151,515,1065,1091]" box="[151,515,1065,1091]" pageId="1" pageNumber="457">
<heading id="D0E734554178FF94A8BBB95FFDF2FB85" bold="true" box="[151,515,1065,1091]" fontSize="11" level="1" pageId="1" pageNumber="457" reason="1">
<emphasis id="B9645F2B4178FF94A8BBB95FFDF2FB85" bold="true" box="[151,515,1065,1091]" pageId="1" pageNumber="457">
Genus
<taxonomicName id="4C10F8BA4178FF94A8C2B95FFDF2FB85" ID-CoL="87GKF" authority="Gerlach, 1957" authorityName="Gerlach" authorityYear="1957" box="[238,515,1065,1091]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94A8C2B95FFEBCFB85" bold="true" box="[238,333,1065,1091]" italics="true" pageId="1" pageNumber="457">Ingenia</emphasis>
<bibRefCitation id="EF81FEC84178FF94A978B95FFDF2FB85" author="Gerlach, S. A." box="[340,515,1065,1091]" pageId="1" pageNumber="457" pagination="411 - 459" refId="ref4951" refString="Gerlach, S. A. (1957) Die Nematodenfauna des Sandstrandes an der Kuste von Mittelbrasilien (Brasilianische Meerse-Nematoden IV). Mitteilungen Aus Dem Zoologischen Museum in Berlin, 33 (2), 411 - 459. https: // doi. org / 10.1002 / mmnz. 19570330206" type="journal article" year="1957">Gerlach, 1957</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B24178FF94A8BBB907FC21FADD" pageId="1" pageNumber="457" type="diagnosis">
<paragraph id="8BAF83394178FF94A8BBB907FC21FADD" blockId="1.[151,1437,1137,1524]" pageId="1" pageNumber="457">
<emphasis id="B9645F2B4178FF94A8BBB907FEFBFB4D" bold="true" box="[151,266,1137,1163]" pageId="1" pageNumber="457">Diagnosis</emphasis>
(modified based on
<bibRefCitation id="EF81FEC84178FF94A9D5B907FD4AFB4D" author="Smol, N. &amp; Muthumbi, A. &amp; Sharma, J." box="[505,699,1137,1163]" pageId="1" pageNumber="457" refId="ref5350" refString="Smol, N., Muthumbi, A. &amp; Sharma, J. (2014) Order Enoplida. In: Schmidt-Rhaesa, A. (Ed.), Gastrotricha, Cycloneuralia, Gnathifera. Vol. 2. Nematoda. Handbook of Zoology. De Gruyter, Berlin, pp. 193 - 249. https: // doi. org / 10.1515 / 9783110274257.193" type="book" year="2014">
Smol
<emphasis id="B9645F2B4178FF94AA12B904FD89FB4D" box="[574,632,1137,1163]" italics="true" pageId="1" pageNumber="457">et al.</emphasis>
2014
</bibRefCitation>
) Cuticle with very fine transverse striations. Lips high. Anterior sensilla in two circles: six long inner labial setae; six very long and jointed outer labial setae and four short and thin cephalic setae in one circle. Amphids in subventral position. Buccal cavity large, consisting of only one part with a large, sharply pointed dorsal tooth and two smaller ventrosublateral teeth. Females didelphic-amphidelphic with antidromously reflexed ovaries. Males with single and outstretched testis.
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B24178FF94A8EBB853FBA5FA32" pageId="1" pageNumber="457" type="discussion">
<paragraph id="8BAF83394178FF94A8EBB853FBA5FA32" blockId="1.[151,1437,1137,1524]" pageId="1" pageNumber="457">
<emphasis id="B9645F2B4178FF94A8EBB853FECAFAF9" bold="true" box="[199,315,1317,1343]" pageId="1" pageNumber="457">Remarks.</emphasis>
<taxonomicName id="4C10F8BA4178FF94A96CB850FE67FAF9" authorityName="Gerlach" authorityYear="1957" box="[320,406,1318,1343]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94A96CB850FE67FAF9" box="[320,406,1318,1343]" italics="true" pageId="1" pageNumber="457">Ingenia</emphasis>
</taxonomicName>
is morphologically similar to
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<emphasis id="B9645F2B4178FF94AAF2B853FC9BFAF9" box="[734,874,1317,1343]" italics="true" pageId="1" pageNumber="457">Bathylaimus</emphasis>
<bibRefCitation id="EF81FEC84178FF94AB43B853FC1EFAF9" author="Cobb, N. A." box="[879,1007,1317,1343]" pageId="1" pageNumber="457" pagination="389 - 421" refId="ref4719" refString="Cobb, N. A. (1894) Tricoma and other nematode genera. The proceedings of the Linnean Society of New South Wales, 8, 389 - 421." type="journal article" year="1894">
Cobb,
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in outer labial setae jointed, amphideal fovea unispiral, buccal cavity large, generally with three teeth and males with large gubernaculum.
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<emphasis id="B9645F2B4178FF94ACDBB83CFABCFAA5" box="[1271,1357,1354,1379]" italics="true" pageId="1" pageNumber="457">Ingenia</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C10F8BA4178FF94A8FBB81BFE92FA41" authorityName="Cobb" authorityYear="1894" box="[215,355,1389,1415]" class="Enoplea" family="Tripyloididae" genus="Bathylaimus" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94A8FBB81BFE92FA41" box="[215,355,1389,1415]" italics="true" pageId="1" pageNumber="457">Bathylaimus</emphasis>
</taxonomicName>
mainly based on buccal cavity that presents only one part in
<taxonomicName id="4C10F8BA4178FF94AC0BB818FB8CFA41" authorityName="Gerlach" authorityYear="1957" box="[1063,1149,1390,1415]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94AC0BB818FB8CFA41" box="[1063,1149,1390,1415]" italics="true" pageId="1" pageNumber="457">Ingenia</emphasis>
</taxonomicName>
while it has two parts in
<taxonomicName id="4C10F8BA4178FF94A8BBB8E7FED2FA6D" authorityName="Cobb" authorityYear="1894" box="[151,291,1425,1451]" class="Enoplea" family="Tripyloididae" genus="Bathylaimus" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94A8BBB8E7FED2FA6D" box="[151,291,1425,1451]" italics="true" pageId="1" pageNumber="457">Bathylaimus</emphasis>
</taxonomicName>
. According to description of
<taxonomicName id="4C10F8BA4178FF94AA47B8E4FD04FA6D" authorityName="Gagarin &amp; Thanh" authorityYear="2007" box="[619,757,1426,1451]" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="species" species="communis">
<emphasis id="B9645F2B4178FF94AA47B8E4FD04FA6D" box="[619,757,1426,1451]" italics="true" pageId="1" pageNumber="457">I. communis</emphasis>
</taxonomicName>
by Gagarin and Thanh (2007), this species presents a buccal cavity with paired stomal pockets, which conforms to character of
<taxonomicName id="4C10F8BA4178FF94ABA6B8C3FBE7FA09" authorityName="Cobb" authorityYear="1894" box="[906,1046,1461,1487]" class="Enoplea" family="Tripyloididae" genus="Bathylaimus" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94ABA6B8C3FBE7FA09" box="[906,1046,1461,1487]" italics="true" pageId="1" pageNumber="457">Bathylaimus</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="4C10F8BA4178FF94AC47B8C0FEB0FA35" authority="Gagarin &amp; Thanh, 2007" authorityName="Gagarin &amp; Thanh" authorityYear="2007" class="Enoplea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="species" species="communis">
<emphasis id="B9645F2B4178FF94AC47B8C0FACAFA09" box="[1131,1339,1462,1487]" italics="true" pageId="1" pageNumber="457">Ingenia communis</emphasis>
Gagarin &amp; Thanh, 2007
</taxonomicName>
is transferred here to
<taxonomicName id="4C10F8BA4178FF94AA1AB8AFFD33FA35" authorityName="Cobb" authorityYear="1894" box="[566,706,1497,1523]" class="Enoplea" family="Tripyloididae" genus="Bathylaimus" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4178FF94AA1AB8AFFD33FA35" box="[566,706,1497,1523]" italics="true" pageId="1" pageNumber="457">Bathylaimus</emphasis>
</taxonomicName>
as
<taxonomicName id="4C10F8BA4178FF94AACBB8AFFC1DFA35" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" baseAuthorityName="Yanwei &amp; Wen &amp; Mengna" baseAuthorityYear="2007" box="[743,1004,1497,1523]" class="Enoplea" family="Tripyloididae" genus="Bathylaimus" kingdom="Animalia" order="Enoplida" pageId="1" pageNumber="457" phylum="Nematoda" rank="species" species="communis" status="comb. nov.">
<emphasis id="B9645F2B4178FF94AACBB8AFFC1DFA35" box="[743,1004,1497,1523]" italics="true" pageId="1" pageNumber="457">Bathylaimus communis</emphasis>
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Yu
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) Cuticle finely striated. Anterior sensilla in two circles (six + ten): inner labial papilla in one circle and difficult to recognize, outer labial setae and cephalic setae in the other circle with the same length. Four subcephalic setae might be present. Buccal cavity minute and slit-like. Amphidial fovea circular. Female reproductive system monorchic with an outstreched ovary to the left of intestine. Male reproductive system monorchic with the gonad to the left of intestine. Posterior testis reduced or absent. Spicules simple and slender. Gubernaculum apophysis absent or present. Tail filiform with three terminal setae.
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(
<figureCitation id="132B9FBC417CFF90A8B3B8EDFED9FA73" box="[159,296,1435,1461]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[152,255,1909,1933]" captionStartId-1="7.[152,255,1503,1527]" captionTargetBox-0="[221,1359,190,1873]" captionTargetBox-1="[151,1436,181,1479]" captionTargetId-0="figure-18@6.[216,1371,181,1884]" captionTargetId-1="figure-27@7.[151,1436,181,1479]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. Linhystera filiformis sp. nov. A. Lateral view of male anterior end, showing amphidial fovea and cephalic setae; B. Lateral view of female anterior end, showing amphidial fovea and cephalic setae; C. Lateral view of female entire body; D. Lateral view of male posterior portion; E. Spicules and gubernaculum; F. Lateral view of female posterior portion. Scales: A, B, D, F = 20 µm, C = 50 µm." captionText-1="FIGURE 4. Linhystera filiformis sp. nov. A. Lateral view of male anterior end, showing amphidial fovea (arrow); B. Lateral view of male anterior end, showing cephalic setae (arrow); C. Lateral view of male posterior portion, showing spicules (arrow); D. Lateral view of male posterior portion, showing gubernaculum (arrow); E. Lateral view of male posterior end, showing tail and terminal setae. Scales: A, B, C, D, E = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13757907" figureDoi-1="http://doi.org/10.5281/zenodo.13757909" httpUri-0="https://zenodo.org/record/13757907/files/figure.png" httpUri-1="https://zenodo.org/record/13757909/files/figure.png" pageId="5" pageNumber="461">Figures 3–4</figureCitation>
,
<tableCitation id="C692B682417CFF90A91FB8EDFE75FA73" box="[307,388,1435,1461]" captionStart="TABLE 1" captionStartId="4.[151,239,260,284]" captionText="TABLE 1. Individual measurements of Ingenia major sp. nov. and Linhystera filiformis sp. nov. (in µm except for de Man’s ratios)." pageId="5" pageNumber="461">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B2417CFF90A8BBB895FAA5F94B" pageId="5" pageNumber="461" type="materials_examined">
<paragraph id="8BAF8339417CFF90A8BBB895FD4FF983" blockId="5.[151,1437,1506,2037]" pageId="5" pageNumber="461">
<emphasis id="B9645F2B417CFF90A8BBB895FEB7FA3A" bold="true" box="[151,326,1506,1533]" pageId="5" pageNumber="461">Type material.</emphasis>
<specimenCount id="9D1648B0417CFF90A961B895FE3AFA3B" box="[333,459,1507,1533]" count="4" pageId="5" pageNumber="461" type="male">Four males</specimenCount>
and
<specimenCount id="9D1648B0417CFF90AA2FB895FD68FA3B" box="[515,665,1507,1533]" count="3" pageId="5" pageNumber="461" type="female">three females</specimenCount>
were measured and studied.
<typeStatus id="54AB3D9B417CFF90ABCDB895FBBFFA3B" box="[993,1102,1507,1533]" pageId="5" pageNumber="461" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1648B0417CFF90AC75B895FB80FA3B" box="[1113,1137,1507,1533]" count="1" pageId="5" pageNumber="461" type="male">♂</specimenCount>
1 on slide 19YMD2–1–47,
<typeStatus id="54AB3D9B417CFF90A8BBBB71FEF6F9E7" box="[151,263,1543,1569]" pageId="5" pageNumber="461" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D1648B0417CFF90A93CBB71FED5F9E7" box="[272,292,1543,1569]" count="1" pageId="5" pageNumber="461" type="male">♂</specimenCount>
2 on slide 22HI7–2–3,
<specimenCount id="9D1648B0417CFF90AA0EBB71FDC9F9E7" box="[546,568,1543,1569]" count="1" pageId="5" pageNumber="461" type="male">♂</specimenCount>
3 on slide 22HI7–3–5, and
<specimenCount id="9D1648B0417CFF90AB4EBB71FC87F9E7" box="[866,886,1543,1569]" count="1" pageId="5" pageNumber="461" type="male">♂</specimenCount>
4 on slide 22HI7–3–20,
<specimenCount id="9D1648B0417CFF90ACAEBB71FB66F9E7" box="[1154,1175,1543,1569]" count="1" pageId="5" pageNumber="461" type="female">♀</specimenCount>
1 on slide 22HI7–3–10,
<specimenCount id="9D1648B0417CFF90A8BBBB5DFF59F983" box="[151,168,1579,1605]" count="1" pageId="5" pageNumber="461" type="female">♀</specimenCount>
2 on slide 22HI7–3–9,
<specimenCount id="9D1648B0417CFF90A982BB5DFE31F983" box="[430,448,1579,1605]" count="1" pageId="5" pageNumber="461" type="female">♀</specimenCount>
3 on slide 22HI7–2–4.
</paragraph>
<paragraph id="8BAF8339417CFF90A8EBBB39FAA5F94B" blockId="5.[151,1437,1506,2037]" pageId="5" pageNumber="461">
<materialsCitation id="3B788964417CFF90A8EBBB39FD94F94B" country="China" county="Rizhao" location="Specimens" municipality="Yangma Island" pageId="5" pageNumber="461" specimenCount="1" stateProvince="Shandong" typeStatus="holotype">
<emphasis id="B9645F2B417CFF90A8EBBB39FDF0F9AE" bold="true" box="[199,513,1614,1641]" pageId="5" pageNumber="461">
<typeStatus id="54AB3D9B417CFF90A8EBBB39FEF3F9AF" box="[199,258,1615,1641]" pageId="5" pageNumber="461">Type</typeStatus>
locality and habitat.
</emphasis>
<location id="8ECFD5E2417CFF90AA26BB39FD72F9AF" LSID="urn:lsid:plazi:treatment:03B9322F417CFF9DA8BBB800FAE8FA6A:8ECFD5E2417CFF90AA26BB39FD72F9AF" box="[522,643,1615,1641]" country="China" county="Rizhao" municipality="Yangma Island" name="Specimens" pageId="5" pageNumber="461" stateProvince="Shandong">Specimens</location>
were obtained from intertidal sandy sediment at
<collectingMunicipality id="6BCB1943417CFF90AC93BB39FA98F9AF" box="[1215,1385,1615,1641]" pageId="5" pageNumber="461">Yangma Island</collectingMunicipality>
and
<collectingCounty id="62CEFBB5417CFF90A8BBBB05FF17F94B" box="[151,230,1651,1677]" pageId="5" pageNumber="461">Rizhao</collectingCounty>
coast,
<collectingRegion id="49D44DDB417CFF90A919BB05FDE3F94B" box="[309,530,1651,1677]" country="China" name="Shandong" pageId="5" pageNumber="461">Shandong Province</collectingRegion>
,
<collectingCountry id="F307C3A9417CFF90AA32BB05FD94F94B" box="[542,613,1651,1677]" name="China" pageId="5" pageNumber="461">China</collectingCountry>
</materialsCitation>
.
<geoCoordinate id="EE24E5FE417CFF90AA42BB04FD21F94B" box="[622,720,1650,1677]" degrees="35" direction="north" minutes="14" orientation="latitude" pageId="5" pageNumber="461" precision="925" value="35.233334">35º14´N</geoCoordinate>
, 121º38΄E (YMD);
<geoCoordinate id="EE24E5FE417CFF90AB82BB05FBFFF94B" box="[942,1038,1651,1677]" degrees="37" direction="north" minutes="28" orientation="latitude" pageId="5" pageNumber="461" precision="925" value="37.466667">37º28´N</geoCoordinate>
, 119º24΄E (HI), respectively.
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B2417CFF90A8EBBBE1FC12F977" box="[199,995,1687,1713]" pageId="5" pageNumber="461" type="etymology">
<paragraph id="8BAF8339417CFF90A8EBBBE1FC12F977" blockId="5.[151,1437,1506,2037]" box="[199,995,1687,1713]" pageId="5" pageNumber="461">
<emphasis id="B9645F2B417CFF90A8EBBBE1FEBFF977" bold="true" box="[199,334,1687,1713]" pageId="5" pageNumber="461">Etymology.</emphasis>
Species epithet
<taxonomicName id="4C10F8BA417CFF90AA28BBE1FD9AF977" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[516,619,1687,1713]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="5" pageNumber="461" phylum="Nematoda" rank="species" species="filiformis">
<emphasis id="B9645F2B417CFF90AA28BBE1FD9AF977" box="[516,619,1687,1713]" italics="true" pageId="5" pageNumber="461">filiformis</emphasis>
</taxonomicName>
refers to the filiform body shape.
</paragraph>
</subSubSection>
<subSubSection id="C30AD0B2417CFF9DA8EBBBCDFAE8FA6A" lastPageId="8" lastPageNumber="464" pageId="5" pageNumber="461" type="description">
<paragraph id="8BAF8339417CFF90A8EBBBCDFC97F913" blockId="5.[151,1437,1506,2037]" box="[199,870,1723,1749]" pageId="5" pageNumber="461">
<emphasis id="B9645F2B417CFF90A8EBBBCDFE8FF913" bold="true" box="[199,382,1723,1749]" pageId="5" pageNumber="461">Measurements.</emphasis>
All measurement data are given in
<tableCitation id="C692B682417CFF90AB20BBCDFC91F913" box="[780,864,1723,1749]" captionStart="TABLE 1" captionStartId="4.[151,239,260,284]" captionText="TABLE 1. Individual measurements of Ingenia major sp. nov. and Linhystera filiformis sp. nov. (in µm except for de Man’s ratios)." pageId="5" pageNumber="461">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BAF8339417CFF90A8EBBBA8FEA8F93E" blockId="5.[151,1437,1506,2037]" box="[199,345,1758,1784]" pageId="5" pageNumber="461">
<emphasis id="B9645F2B417CFF90A8EBBBA8FEA8F93E" bold="true" box="[199,345,1758,1784]" pageId="5" pageNumber="461">Description.</emphasis>
</paragraph>
<paragraph id="8BAF8339417CFF90A8EBBA75FDE7F833" blockId="5.[151,1437,1506,2037]" pageId="5" pageNumber="461">
<emphasis id="B9645F2B417CFF90A8EBBA75FEE7F8DB" bold="true" box="[199,278,1795,1821]" pageId="5" pageNumber="461">Males.</emphasis>
Body cylindrical, filiform, tapering towards both extremities. Cuticle striated. Six outer labial setae and four cephalic setae in the same circle and with the same length, 4–5 μm (0.6–0.8 head diameter). Amphidial fovea circular, 4–5 μm in diameter (0.4–0.6 c.b.d.), located at 7–8 μm (1.0–1.2 head diameter) from anterior body end. Buccal cavity minute, slit-like. Pharynx muscular and cylindrical, posterior end not widened and without posterior bulb. Cardia triangular. Nerve ring positioned slightly posterior to half of pharynx length (0.5–0.7 pharynx length). Excretory pore and ventral gland not observed. Tail filiform, 14–16 c.b.d. Three terminal setae, 5 μm in length. Three caudal glands in tail region.
</paragraph>
<caption id="DF6FD3B1417FFF93A8B4BA03FE31F83F" ID-DOI="http://doi.org/10.5281/zenodo.13757907" ID-Zenodo-Dep="13757907" httpUri="https://zenodo.org/record/13757907/files/figure.png" pageId="6" pageNumber="462" startId="6.[152,255,1909,1933]" targetBox="[221,1359,190,1873]" targetPageId="6" targetType="figure">
<paragraph id="8BAF8339417FFF93A8B4BA03FE31F83F" blockId="6.[151,1437,1909,2041]" pageId="6" pageNumber="462">
<emphasis id="B9645F2B417FFF93A8B4BA03FEE2F84B" bold="true" box="[152,275,1909,1933]" pageId="6" pageNumber="462">FIGURE 3</emphasis>
.
<taxonomicName id="4C10F8BA417FFF93A90CBA03FE00F84B" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[288,497,1909,1933]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="6" pageNumber="462" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B417FFF93A90CBA03FE00F84B" box="[288,497,1909,1933]" italics="true" pageId="6" pageNumber="462">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417FFF93A9DBBA03FDB8F84B" bold="true" box="[503,585,1909,1933]" pageId="6" pageNumber="462">
<taxonomicNameLabel id="A257E250417FFF93A9DBBA03FDB8F84B" box="[503,585,1909,1933]" pageId="6" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Lateral view of male anterior end, showing amphidial fovea and cephalic setae; B. Lateral view of female anterior end, showing amphidial fovea and cephalic setae; C. Lateral view of female entire body; D. Lateral view of male posterior portion; E. Spicules and gubernaculum; F. Lateral view of female posterior portion. Scales: A, B, D, F = 20 µm, C = 50 µm.
</paragraph>
</caption>
<caption id="DF6FD3B1417EFF92A8B4B8A9FD64F9A5" ID-DOI="http://doi.org/10.5281/zenodo.13757909" ID-Zenodo-Dep="13757909" httpUri="https://zenodo.org/record/13757909/files/figure.png" pageId="7" pageNumber="463" startId="7.[152,255,1503,1527]" targetBox="[151,1436,181,1479]" targetPageId="7" targetType="figure">
<paragraph id="8BAF8339417EFF92A8B4B8A9FD64F9A5" blockId="7.[151,1436,1503,1635]" pageId="7" pageNumber="463">
<emphasis id="B9645F2B417EFF92A8B4B8A9FEE2FA31" bold="true" box="[152,275,1503,1527]" pageId="7" pageNumber="463">FIGURE 4</emphasis>
.
<taxonomicName id="4C10F8BA417EFF92A90CB8A9FE00FA31" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[288,497,1503,1527]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="7" pageNumber="463" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B417EFF92A90CB8A9FE00FA31" box="[288,497,1503,1527]" italics="true" pageId="7" pageNumber="463">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417EFF92A9D4B8A9FDB8FA31" bold="true" box="[504,585,1503,1527]" pageId="7" pageNumber="463">
<taxonomicNameLabel id="A257E250417EFF92A9D4B8A9FDB8FA31" box="[504,585,1503,1527]" pageId="7" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Lateral view of male anterior end, showing amphidial fovea (arrow); B. Lateral view of male anterior end, showing cephalic setae (arrow); C. Lateral view of male posterior portion, showing spicules (arrow); D. Lateral view of male posterior portion, showing gubernaculum (arrow); E. Lateral view of male posterior end, showing tail and terminal setae. Scales: A, B, C, D, E = 20 µm.
</paragraph>
</caption>
<paragraph id="8BAF8339417EFF92A8EBBBC3FF17F8FD" blockId="7.[151,1437,1717,2031]" pageId="7" pageNumber="463">The reproductive system with one testis outstretched to the left of the intestine. Spicules paired and equal in length (1.4–1.6 cloacal body diameter), sickle-shaped, with a small capitulum in the proximal end, middle part enlarged and distal end tapered. Gubernaculum boat-shaped, 0.4–0.5 cloacal body diameter. Precloacal supplements absent.</paragraph>
<paragraph id="8BAF8339417EFF92A8EBBA33FA71F845" blockId="7.[151,1437,1717,2031]" pageId="7" pageNumber="463">
<emphasis id="B9645F2B417EFF92A8EBBA33FEC1F899" bold="true" box="[199,304,1861,1887]" pageId="7" pageNumber="463">Females.</emphasis>
Similar to males in most characters. Single anterior outstretched ovary to the left of the intestine.Vulva located at mid-body, 460–489 μm (49.8–54.6 % body length) to anterior body end. Vagina short and sclerotized.
</paragraph>
<paragraph id="8BAF8339417EFF92A8EBBAFBFBBCF829" blockId="7.[151,1437,1717,2031]" pageId="7" pageNumber="463">
<emphasis id="B9645F2B417EFF92A8EBBAFBFD5BF861" bold="true" box="[199,682,1933,1959]" pageId="7" pageNumber="463">Differentiation diagnosis and discussion.</emphasis>
<taxonomicName id="4C10F8BA417EFF92AA9DBAFBFC66F861" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[689,919,1933,1959]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="7" pageNumber="463" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B417EFF92AA9DBAFBFC66F861" box="[689,919,1933,1959]" italics="true" pageId="7" pageNumber="463">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B417EFF92ABB2BAFBFC09F861" bold="true" box="[926,1016,1933,1959]" pageId="7" pageNumber="463">
<taxonomicNameLabel id="A257E250417EFF92ABB2BAFBFC09F861" box="[926,1016,1933,1959]" pageId="7" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by filiform body, six outer labial setae and four cephalic setae in one circle with the same length, amphidial fovea circular, buccal cavity minute, slit-like, spicules sickle-shaped, gubernaculum boat-shaped and tail filiform.
</paragraph>
<caption id="DF6FD3B14171FF9DA8BBBDE1FED6FE86" pageId="8" pageNumber="464" startId="8.[151,239,151,175]" targetType="table">
<paragraph id="8BAF83394171FF9DA8BBBDE1FED6FE86" blockId="8.[151,1437,151,320]" pageId="8" pageNumber="464">
<emphasis id="B9645F2B4171FF9DA8BBBDE1FEFBFF69" bold="true" box="[151,266,151,175]" pageId="8" pageNumber="464">TABLE 2.</emphasis>
Morphometric characteristics of valid species of the genus
<taxonomicName id="4C10F8BA4171FF9DABA4BDE1FB79FF76" authority="Juario, 1974" authorityName="Juario" authorityYear="1974" box="[904,1160,151,176]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4171FF9DABA4BDE1FC0BFF69" box="[904,1018,151,175]" italics="true" pageId="8" pageNumber="464">Linhystera</emphasis>
<bibRefCitation id="EF81FEC84171FF9DAC2DBDE1FB79FF76" author="Juario, J. V." box="[1025,1160,151,176]" pageId="8" pageNumber="464" pagination="275 - 303" refId="ref5004" refString="Juario, J. V. (1974) Neue freilebende Nematoden aus dem Sublitoral der Deutschen Bucht. Veroffentlichungen Des Instituts fur Meeresforschung in Bremerhaven, 14, 275 - 303." type="journal article" year="1974">Juario, 1974</bibRefCitation>
</taxonomicName>
. *The value was obtained by measurements in the figure and description; the dash means the data are absent; L, body length in µm; CS, cephalic setae length in µm; c.b.d., corresponding body diameter; Amp, amphidial fovea diameter/ corresponding body diameter (%); Pharynx, pharynx length in µm; Spicules, spicules length in µm, Gubernacular apophysis, gubernacular apophysis length in µm.
</paragraph>
</caption>
<paragraph id="8BAF83394171FF9DAB4BBC26FA9FFBCA" pageId="8" pageNumber="464">
<table id="F91071994171006AA8E9BC26FA7CFBD8" box="[197,1421,336,1054]" gridcols="6" gridrows="13" pageId="8" pageNumber="464">
<tr id="3520817B4171006AA8E9BC26FA7CFEAE" box="[197,1421,336,360]" gridrow="0" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BC26FA7CFEAE" box="[197,1421,336,360]" colspan="6" colspanRight="5" gridcol="0" gridrow="0" pageId="8" pageNumber="464">Species</th>
</tr>
<tr id="3520817B4171006AA8E9BCFEFA7CFE72" box="[197,1421,392,436]" gridrow="1" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BCFEFEBEFE72" box="[197,335,392,436]" gridcol="0" gridrow="1" pageId="8" pageNumber="464">Characters</th>
<td id="76F1E8074171006AA9A2BCFEFDB1FE72" box="[398,576,392,436]" gridcol="1" gridrow="1" pageId="8" pageNumber="464">
<taxonomicName id="4C10F8BA4171FF9DA9A2BCEBFDB1FE72" authorityName="Yu, Huang &amp; Xu" authorityYear="2014" box="[398,576,412,436]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="breviapophysis">
<emphasis id="B9645F2B4171FF9DA9A2BCEBFDB1FE72" box="[398,576,412,436]" italics="true" pageId="8" pageNumber="464">L. breviapophysis</emphasis>
</taxonomicName>
</td>
<td id="76F1E8074171006AAA72BCFEFCDDFE72" box="[606,812,392,436]" gridcol="2" gridrow="1" pageId="8" pageNumber="464">
<taxonomicName id="4C10F8BA4171FF9DAA72BCEBFD24FE72" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[606,725,412,436]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B4171FF9DAA72BCEBFD24FE72" box="[606,725,412,436]" italics="true" pageId="8" pageNumber="464">L. filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B4171FF9DAAF0BCEAFCDDFE72" bold="true" box="[732,812,412,436]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A257E2504171FF9DAAF0BCEAFCDDFE72" box="[732,812,412,436]" pageId="8" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
<td id="76F1E8074171006AAB79BCFEFC3AFE72" box="[853,971,392,436]" gridcol="3" gridrow="1" pageId="8" pageNumber="464">
<taxonomicName id="4C10F8BA4171FF9DAB4ABCEBFC4AFE72" authorityName="Pastor de Ward" authorityYear="1985" box="[870,955,412,436]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="longa">
<emphasis id="B9645F2B4171FF9DAB4ABCEBFC4AFE72" box="[870,955,412,436]" italics="true" pageId="8" pageNumber="464">L. longa</emphasis>
</taxonomicName>
</td>
<td id="76F1E8074171006AABD5BCFEFB5CFE72" box="[1017,1197,392,436]" gridcol="4" gridrow="1" pageId="8" pageNumber="464">
<taxonomicName id="4C10F8BA4171FF9DABD5BCEBFB5CFE72" authorityName="Yu, Huang &amp; Xu" authorityYear="2014" box="[1017,1197,412,436]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="longiapophysis">
<emphasis id="B9645F2B4171FF9DABD5BCEBFB5CFE72" box="[1017,1197,412,436]" italics="true" pageId="8" pageNumber="464">L. longiapophysis</emphasis>
</taxonomicName>
</td>
<td id="76F1E8074171006AACFCBCFEFA7CFE72" box="[1232,1421,392,436]" gridcol="5" gridrow="1" pageId="8" pageNumber="464">
<taxonomicName id="4C10F8BA4171FF9DACF2BCEBFA8EFE72" authorityName="Juario" authorityYear="1974" box="[1246,1407,412,436]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="problematica">
<emphasis id="B9645F2B4171FF9DACF2BCEBFA8EFE72" box="[1246,1407,412,436]" italics="true" pageId="8" pageNumber="464">L. problematica</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3520817B4171006AA8E9BC9AFA7CFDC2" box="[197,1421,492,516]" gridrow="2" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BC9AFEBEFDC2" box="[197,335,492,516]" gridcol="0" gridrow="2" pageId="8" pageNumber="464">L</th>
<td id="76F1E8074171006AA9A2BC9AFDB1FDC2" box="[398,576,492,516]" gridcol="1" gridrow="2" pageId="8" pageNumber="464">633–787</td>
<td id="76F1E8074171006AAA72BC9AFCDDFDC2" box="[606,812,492,516]" gridcol="2" gridrow="2" pageId="8" pageNumber="464">829–1048</td>
<td id="76F1E8074171006AAB79BC9AFC3AFDC2" box="[853,971,492,516]" gridcol="3" gridrow="2" pageId="8" pageNumber="464">1250–1320</td>
<td id="76F1E8074171006AABD5BC9AFB5CFDC2" box="[1017,1197,492,516]" gridcol="4" gridrow="2" pageId="8" pageNumber="464">1232</td>
<td id="76F1E8074171006AACFCBC9AFA7CFDC2" box="[1232,1421,492,516]" gridcol="5" gridrow="2" pageId="8" pageNumber="464">720–833</td>
</tr>
<tr id="3520817B4171006AA8E9BF6AFA7CFDF2" box="[197,1421,540,564]" gridrow="3" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BF6AFEBEFDF2" box="[197,335,540,564]" gridcol="0" gridrow="3" pageId="8" pageNumber="464">
<emphasis id="B9645F2B4171FF9DA928BF6BFEE0FDF2" box="[260,273,541,564]" italics="true" pageId="8" pageNumber="464">a</emphasis>
</th>
<td id="76F1E8074171006AA9A2BF6AFDB1FDF2" box="[398,576,540,564]" gridcol="1" gridrow="3" pageId="8" pageNumber="464">43.4–59.2</td>
<td id="76F1E8074171006AAA72BF6AFCDDFDF2" box="[606,812,540,564]" gridcol="2" gridrow="3" pageId="8" pageNumber="464">41.7–53.9</td>
<td id="76F1E8074171006AAB79BF6AFC3AFDF2" box="[853,971,540,564]" gridcol="3" gridrow="3" pageId="8" pageNumber="464">50.0–72.2</td>
<td id="76F1E8074171006AABD5BF6AFB5CFDF2" box="[1017,1197,540,564]" gridcol="4" gridrow="3" pageId="8" pageNumber="464">54.0</td>
<td id="76F1E8074171006AACFCBF6AFA7CFDF2" box="[1232,1421,540,564]" gridcol="5" gridrow="3" pageId="8" pageNumber="464">45.0–52.1</td>
</tr>
<tr id="3520817B4171006AA8E9BF3DFA7CFDA2" box="[197,1421,587,612]" gridrow="4" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BF3DFEBEFDA2" box="[197,335,587,612]" gridcol="0" gridrow="4" pageId="8" pageNumber="464">
<emphasis id="B9645F2B4171FF9DA928BF3DFEE0FDA5" box="[260,273,587,611]" italics="true" pageId="8" pageNumber="464">b</emphasis>
</th>
<td id="76F1E8074171006AA9A2BF3DFDB1FDA2" box="[398,576,587,612]" gridcol="1" gridrow="4" pageId="8" pageNumber="464">8.9–9.9</td>
<td id="76F1E8074171006AAA72BF3DFCDDFDA2" box="[606,812,587,612]" gridcol="2" gridrow="4" pageId="8" pageNumber="464">9.1–10.6</td>
<td id="76F1E8074171006AAB79BF3DFC3AFDA2" box="[853,971,587,612]" gridcol="3" gridrow="4" pageId="8" pageNumber="464">9.5–10.4</td>
<td id="76F1E8074171006AABD5BF3DFB5CFDA2" box="[1017,1197,587,612]" gridcol="4" gridrow="4" pageId="8" pageNumber="464">10.0</td>
<td id="76F1E8074171006AACFCBF3DFA7CFDA2" box="[1232,1421,587,612]" gridcol="5" gridrow="4" pageId="8" pageNumber="464">6.3</td>
</tr>
<tr id="3520817B4171006AA8E9BF0DFA7CFD55" box="[197,1421,635,659]" gridrow="5" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BF0DFEBEFD55" box="[197,335,635,659]" gridcol="0" gridrow="5" pageId="8" pageNumber="464">
<emphasis id="B9645F2B4171FF9DA928BF0AFEFEFD55" box="[260,271,636,659]" italics="true" pageId="8" pageNumber="464">c</emphasis>
</th>
<td id="76F1E8074171006AA9A2BF0DFDB1FD55" box="[398,576,635,659]" gridcol="1" gridrow="5" pageId="8" pageNumber="464">4.2–5.3</td>
<td id="76F1E8074171006AAA72BF0DFCDDFD55" box="[606,812,635,659]" gridcol="2" gridrow="5" pageId="8" pageNumber="464">4.0–5.8</td>
<td id="76F1E8074171006AAB79BF0DFC3AFD55" box="[853,971,635,659]" gridcol="3" gridrow="5" pageId="8" pageNumber="464">4.0–4.8</td>
<td id="76F1E8074171006AABD5BF0DFB5CFD55" box="[1017,1197,635,659]" gridcol="4" gridrow="5" pageId="8" pageNumber="464">4.5</td>
<td id="76F1E8074171006AACFCBF0DFA7CFD55" box="[1232,1421,635,659]" gridcol="5" gridrow="5" pageId="8" pageNumber="464">7.6–7.8</td>
</tr>
<tr id="3520817B4171006AA8E9BFDDFA7CFD05" box="[197,1421,683,707]" gridrow="6" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BFDDFEBEFD05" box="[197,335,683,707]" gridcol="0" gridrow="6" pageId="8" pageNumber="464">CS</th>
<td id="76F1E8074171006AA9A2BFDDFDB1FD05" box="[398,576,683,707]" gridcol="1" gridrow="6" pageId="8" pageNumber="464">3–4</td>
<td id="76F1E8074171006AAA72BFDDFCDDFD05" box="[606,812,683,707]" gridcol="2" gridrow="6" pageId="8" pageNumber="464">4–5</td>
<td id="76F1E8074171006AAB79BFDDFC3AFD05" box="[853,971,683,707]" gridcol="3" gridrow="6" pageId="8" pageNumber="464">3–6</td>
<td id="76F1E8074171006AABD5BFDDFB5CFD05" box="[1017,1197,683,707]" gridcol="4" gridrow="6" pageId="8" pageNumber="464">6</td>
<td id="76F1E8074171006AACFCBFDDFA7CFD05" box="[1232,1421,683,707]" gridcol="5" gridrow="6" pageId="8" pageNumber="464">4</td>
</tr>
<tr id="3520817B4171006AA8E9BFADFA7CFD35" box="[197,1421,731,755]" gridrow="7" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BFADFEBEFD35" box="[197,335,731,755]" gridcol="0" gridrow="7" pageId="8" pageNumber="464">Amp</th>
<td id="76F1E8074171006AA9A2BFADFDB1FD35" box="[398,576,731,755]" gridcol="1" gridrow="7" pageId="8" pageNumber="464">40.9–59.1 c.b.d.</td>
<td id="76F1E8074171006AAA72BFADFCDDFD35" box="[606,812,731,755]" gridcol="2" gridrow="7" pageId="8" pageNumber="464">42.9–66.7 c.b.d.</td>
<td id="76F1E8074171006AAB79BFADFC3AFD35" box="[853,971,731,755]" gridcol="3" gridrow="7" pageId="8" pageNumber="464">62.5 c.b.d.*</td>
<td id="76F1E8074171006AABD5BFADFB5CFD35" box="[1017,1197,731,755]" gridcol="4" gridrow="7" pageId="8" pageNumber="464">44.1 c.b.d.</td>
<td id="76F1E8074171006AACFCBFADFA7CFD35" box="[1232,1421,731,755]" gridcol="5" gridrow="7" pageId="8" pageNumber="464">96.6–100.0 c.b.d.*</td>
</tr>
<tr id="3520817B4171006AA8E9BE7CFA7CFCE5" box="[197,1421,778,803]" gridrow="8" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BE7CFEBEFCE5" box="[197,335,778,803]" gridcol="0" gridrow="8" pageId="8" pageNumber="464">Pharynx</th>
<td id="76F1E8074171006AA9A2BE7CFDB1FCE5" box="[398,576,778,803]" gridcol="1" gridrow="8" pageId="8" pageNumber="464">71–81</td>
<td id="76F1E8074171006AAA72BE7CFCDDFCE5" box="[606,812,778,803]" gridcol="2" gridrow="8" pageId="8" pageNumber="464">79–104</td>
<td id="76F1E8074171006AAB79BE7CFC3AFCE5" box="[853,971,778,803]" gridcol="3" gridrow="8" pageId="8" pageNumber="464">125–132*</td>
<td id="76F1E8074171006AABD5BE7CFB5CFCE5" box="[1017,1197,778,803]" gridcol="4" gridrow="8" pageId="8" pageNumber="464">123</td>
<td id="76F1E8074171006AACFCBE7CFA7CFCE5" box="[1232,1421,778,803]" gridcol="5" gridrow="8" pageId="8" pageNumber="464">114–132*</td>
</tr>
<tr id="3520817B4171006AA8E9BE4CFA7CFC94" box="[197,1421,826,850]" gridrow="9" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BE4CFEBEFC94" box="[197,335,826,850]" gridcol="0" gridrow="9" pageId="8" pageNumber="464">Spicules</th>
<td id="76F1E8074171006AA9A2BE4CFDB1FC94" box="[398,576,826,850]" gridcol="1" gridrow="9" pageId="8" pageNumber="464">17–18</td>
<td id="76F1E8074171006AAA72BE4CFCDDFC94" box="[606,812,826,850]" gridcol="2" gridrow="9" pageId="8" pageNumber="464">19–21</td>
<td id="76F1E8074171006AAB79BE4CFC3AFC94" box="[853,971,826,850]" gridcol="3" gridrow="9" pageId="8" pageNumber="464">42–53</td>
<td id="76F1E8074171006AABD5BE4CFB5CFC94" box="[1017,1197,826,850]" gridcol="4" gridrow="9" pageId="8" pageNumber="464">24</td>
<td id="76F1E8074171006AACFCBE4CFA7CFC94" box="[1232,1421,826,850]" gridcol="5" gridrow="9" pageId="8" pageNumber="464">22</td>
</tr>
<tr id="3520817B4171006AA8E9BE10FA7CFC64" box="[197,1421,870,930]" gridrow="10" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BE10FEBEFC64" box="[197,335,870,930]" gridcol="0" gridrow="10" pageId="8" pageNumber="464">Gubernacular apophysis</th>
<td id="76F1E8074171006AA9A2BE10FDB1FC64" box="[398,576,870,930]" gridcol="1" gridrow="10" pageId="8" pageNumber="464">3</td>
<td id="76F1E8074171006AAA72BE10FCDDFC64" box="[606,812,870,930]" gridcol="2" gridrow="10" pageId="8" pageNumber="464">-</td>
<td id="76F1E8074171006AAB79BE10FC3AFC64" box="[853,971,870,930]" gridcol="3" gridrow="10" pageId="8" pageNumber="464">-</td>
<td id="76F1E8074171006AABD5BE10FB5CFC64" box="[1017,1197,870,930]" gridcol="4" gridrow="10" pageId="8" pageNumber="464">10</td>
<td id="76F1E8074171006AACFCBE10FA7CFC64" box="[1232,1421,870,930]" gridcol="5" gridrow="10" pageId="8" pageNumber="464">-</td>
</tr>
<tr id="3520817B4171006AA8E9BEC0FA7CFC08" box="[197,1421,950,974]" gridrow="11" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BEC0FEBEFC08" box="[197,335,950,974]" gridcol="0" gridrow="11" pageId="8" pageNumber="464">Tail</th>
<td id="76F1E8074171006AA9A2BEC0FDB1FC08" box="[398,576,950,974]" gridcol="1" gridrow="11" pageId="8" pageNumber="464">132–167</td>
<td id="76F1E8074171006AAA72BEC0FCDDFC08" box="[606,812,950,974]" gridcol="2" gridrow="11" pageId="8" pageNumber="464">154–233</td>
<td id="76F1E8074171006AAB79BEC0FC3AFC08" box="[853,971,950,974]" gridcol="3" gridrow="11" pageId="8" pageNumber="464">270–322*</td>
<td id="76F1E8074171006AABD5BEC0FB5CFC08" box="[1017,1197,950,974]" gridcol="4" gridrow="11" pageId="8" pageNumber="464">276</td>
<td id="76F1E8074171006AACFCBEC0FA7CFC08" box="[1232,1421,950,974]" gridcol="5" gridrow="11" pageId="8" pageNumber="464">95–106</td>
</tr>
<tr id="3520817B4171006AA8E9BE94FA7CFBD8" box="[197,1421,994,1054]" gridrow="12" pageId="8" pageNumber="464">
<th id="76F1E8074171006AA8E9BE94FEBEFBD8" box="[197,335,994,1054]" gridcol="0" gridrow="12" pageId="8" pageNumber="464">Reference</th>
<td id="76F1E8074171006AA9A2BE94FDB1FBD8" box="[398,576,994,1054]" gridcol="1" gridrow="12" pageId="8" pageNumber="464">
<bibRefCitation id="EF81FEC84171FF9DA98CBE83FDDEFBCA" author="Yu, T. T. &amp; Huang, Y. &amp; Xu, K. D." box="[416,559,1012,1036]" pageId="8" pageNumber="464" pagination="515 - 520" refId="ref5418" refString="Yu, T. T., Huang, Y. &amp; Xu, K. D. (2014) Two new nematode species, Linhystera breviapophysis and L. longiapophysis (Xyalidae, Nematoda), from the East China Sea. Journal of the Marine Biological Association of the United Kingdom, 94 (3), 515 - 520." type="journal article" year="2014">
Yu
<emphasis id="B9645F2B4171FF9DA9EFBE83FE07FBCA" box="[451,502,1012,1036]" italics="true" pageId="8" pageNumber="464">et al.</emphasis>
2014
</bibRefCitation>
</td>
<td id="76F1E8074171006AAA72BE94FCDDFBD8" box="[606,812,994,1054]" gridcol="2" gridrow="12" pageId="8" pageNumber="464">Data by the present author</td>
<td id="76F1E8074171006AAB79BE94FC3AFBD8" box="[853,971,994,1054]" gridcol="3" gridrow="12" pageId="8" pageNumber="464">
<bibRefCitation id="EF81FEC84171FF9DAB4DBE95FC3AFBD8" author="Pastor de Ward, C. T." pageId="8" pageNumber="464" pagination="113 - 130" refId="ref5187" refString="Pastor de Ward, C. T. (1985) Nematodes marinos de la ria deseado (Monhysteroidea, Xyalidae) Santa Cruz Argentina II. Physis, Buenos Aires, 43, 113 - 130." type="journal article" year="1985">Pastor de Ward, 1985</bibRefCitation>
</td>
<td id="76F1E8074171006AABD5BE94FB5CFBD8" box="[1017,1197,994,1054]" gridcol="4" gridrow="12" pageId="8" pageNumber="464">
<bibRefCitation id="EF81FEC84171FF9DAC20BE83FB6AFBCA" author="Yu, T. T. &amp; Huang, Y. &amp; Xu, K. D." box="[1036,1179,1012,1036]" pageId="8" pageNumber="464" pagination="515 - 520" refId="ref5418" refString="Yu, T. T., Huang, Y. &amp; Xu, K. D. (2014) Two new nematode species, Linhystera breviapophysis and L. longiapophysis (Xyalidae, Nematoda), from the East China Sea. Journal of the Marine Biological Association of the United Kingdom, 94 (3), 515 - 520." type="journal article" year="2014">
Yu
<emphasis id="B9645F2B4171FF9DAC03BE83FB93FBCA" box="[1071,1122,1012,1036]" italics="true" pageId="8" pageNumber="464">et al.</emphasis>
2014
</bibRefCitation>
</td>
<td id="76F1E8074171006AACFCBE94FA7CFBD8" box="[1232,1421,994,1054]" gridcol="5" gridrow="12" pageId="8" pageNumber="464">
<bibRefCitation id="EF81FEC84171FF9DACDCBE82FA9FFBCA" author="Juario, J. V." box="[1264,1390,1012,1036]" pageId="8" pageNumber="464" pagination="275 - 303" refId="ref5004" refString="Juario, J. V. (1974) Neue freilebende Nematoden aus dem Sublitoral der Deutschen Bucht. Veroffentlichungen Des Instituts fur Meeresforschung in Bremerhaven, 14, 275 - 303." type="journal article" year="1974">Juario, 1974</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF83394171FF9DA8EBB904FAE8FA6A" blockId="8.[151,1437,1137,1452]" pageId="8" pageNumber="464">
According to the diagnostic character of “presence and absence of gubernacular apophysis” (
<figureCitation id="132B9FBC4171FF9DACCEB904FAB5FB4A" box="[1250,1348,1137,1164]" captionStart="FIGURE 5" captionStartId="9.[152,255,1385,1409]" captionTargetBox="[160,1430,187,1355]" captionTargetId="figure-27@9.[151,1436,181,1361]" captionTargetPageId="9" captionText="FIGURE 5. Pictorial key for identification of the species in genus Linhystera Juario, 1974." figureDoi="http://doi.org/10.5281/zenodo.13757911" httpUri="https://zenodo.org/record/13757911/files/figure.png" pageId="8" pageNumber="464">Figure 5</figureCitation>
), genus
<taxonomicName id="4C10F8BA4171FF9DA8BBB9E0FEFEFB76" authorityName="Juario" authorityYear="1974" box="[151,271,1174,1200]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4171FF9DA8BBB9E0FEFEFB76" box="[151,271,1174,1200]" italics="true" pageId="8" pageNumber="464">Linhystera</emphasis>
</taxonomicName>
can be divided in two groups, and morphometric characters of valid species of
<taxonomicName id="4C10F8BA4171FF9DAC9CB9E0FAD9FB76" authorityName="Juario" authorityYear="1974" box="[1200,1320,1174,1200]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="genus">
<emphasis id="B9645F2B4171FF9DAC9CB9E0FAD9FB76" box="[1200,1320,1174,1200]" italics="true" pageId="8" pageNumber="464">Linhystera</emphasis>
</taxonomicName>
are given in
<tableCitation id="C692B6824171FF9DA89BB9CCFEC3FB15" box="[183,306,1210,1236]" captionStart="TABLE 2" captionStartId="8.[151,239,151,175]" captionText="TABLE 2. Morphometric characteristics of valid species of the genus Linhystera Juario, 1974. *The value was obtained by measurements in the figure and description; the dash means the data are absent; L, body length in µm; CS, cephalic setae length in µm; c.b.d., corresponding body diameter; Amp, amphidial fovea diameter/ corresponding body diameter (%); Pharynx, pharynx length in µm; Spicules, spicules length in µm, Gubernacular apophysis, gubernacular apophysis length in µm." pageId="8" pageNumber="464">
Table 2.
<taxonomicName id="4C10F8BA4171FF9DA933B9CCFEC3FB15" authorityName="Yu, Huang &amp; Xu" authorityYear="2014" box="[287,306,1210,1235]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="breviapophysis">
<emphasis id="B9645F2B4171FF9DA933B9CCFEC3FB15" box="[287,306,1210,1235]" italics="true" pageId="8" pageNumber="464">L</emphasis>
</taxonomicName>
</tableCitation>
. breviapophysis and
<taxonomicName id="4C10F8BA4171FF9DAA0AB9CCFD00FB12" authorityName="Yu, Huang &amp; Xu" authorityYear="2014" box="[550,753,1210,1236]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="longiapophysis">
<emphasis id="B9645F2B4171FF9DAA0AB9CCFD00FB12" box="[550,753,1210,1236]" italics="true" pageId="8" pageNumber="464">L. longiapophysis</emphasis>
</taxonomicName>
are characterized with the presence of gubernaculum with apophysis, while
<taxonomicName id="4C10F8BA4171FF9DA94AB9A8FE01FB3E" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[358,496,1246,1272]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B4171FF9DA94AB9A8FE01FB3E" box="[358,496,1246,1272]" italics="true" pageId="8" pageNumber="464">L. filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B4171FF9DA9D0B9A8FDAAFB3E" bold="true" box="[508,603,1246,1272]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A257E2504171FF9DA9D0B9A8FDAAFB3E" box="[508,603,1246,1272]" pageId="8" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C10F8BA4171FF9DAA43B9A8FD23FB3E" authorityName="Pastor de Ward" authorityYear="1985" box="[623,722,1246,1272]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="longa">
<emphasis id="B9645F2B4171FF9DAA43B9A8FD23FB3E" box="[623,722,1246,1272]" italics="true" pageId="8" pageNumber="464">L. longa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C10F8BA4171FF9DAB38B9A8FC3AFB3E" authorityName="Juario" authorityYear="1974" box="[788,971,1246,1272]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="problematica">
<emphasis id="B9645F2B4171FF9DAB38B9A8FC3AFB3E" box="[788,971,1246,1272]" italics="true" pageId="8" pageNumber="464">L. problematica</emphasis>
</taxonomicName>
with gubernaculum without apophysis.
<taxonomicName id="4C10F8BA4171FF9DA8BBB874FE8CFADA" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[151,381,1282,1308]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B4171FF9DA8BBB874FE8CFADA" box="[151,381,1282,1308]" italics="true" pageId="8" pageNumber="464">Linhystera filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B4171FF9DA9A8B874FE2FFADA" bold="true" box="[388,478,1282,1308]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A257E2504171FF9DA9A8B874FE2FFADA" box="[388,478,1282,1308]" pageId="8" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is differentiated from
<taxonomicName id="4C10F8BA4171FF9DAAF6B874FC7DFADA" authorityName="Juario" authorityYear="1974" box="[730,908,1282,1308]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="problematica">
<emphasis id="B9645F2B4171FF9DAAF6B874FC7DFADA" box="[730,908,1282,1308]" italics="true" pageId="8" pageNumber="464">L. problematica</emphasis>
</taxonomicName>
in the body length (829–1048 μm vs. 720–833 μm), amphidial fovea size (3–5 μm vs. 5–8 μm in diameter), amphidial fovea location (6–8 μm vs. 8–11 μm from anterior end) and the tail length (154–233 μm vs. 95–106 μm); differs from
<taxonomicName id="4C10F8BA4171FF9DAC28B83CFB92FAA2" authorityName="Pastor de Ward" authorityYear="1985" box="[1028,1123,1354,1380]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="longa">
<emphasis id="B9645F2B4171FF9DAC28B83CFB92FAA2" box="[1028,1123,1354,1380]" italics="true" pageId="8" pageNumber="464">L. longa</emphasis>
</taxonomicName>
in shorter body (829–1048 μm vs. 1250–1320 μm), shorter tail (154–233 μm vs. 270–322 μm) and shorter spicules (19–21 μm vs. 51–53 μm). Furthermore, differences between
<taxonomicName id="4C10F8BA4171FF9DAA34B8E4FD6CFA6A" authorityName="Yanwei &amp; Wen &amp; Mengna" authorityYear="2024" box="[536,669,1426,1452]" class="Chromadorea" family="Xyalidae" genus="Linhystera" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="464" phylum="Nematoda" rank="species" species="filiformis" status="sp. nov.">
<emphasis id="B9645F2B4171FF9DAA34B8E4FD6CFA6A" box="[536,669,1426,1452]" italics="true" pageId="8" pageNumber="464">L. filiformis</emphasis>
</taxonomicName>
<emphasis id="B9645F2B4171FF9DAA88B8E4FD0CFA6A" bold="true" box="[676,765,1426,1452]" pageId="8" pageNumber="464">
<taxonomicNameLabel id="A257E2504171FF9DAA88B8E4FD0CFA6A" box="[676,765,1426,1452]" pageId="8" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and other congeners are listed in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>

331
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<taxonomicName id="4C104D575122275350B9E683EC79FB1E" authority="Davidian" authorityName="Davidian &amp; Belokobylskij" authorityYear="2024" box="[151,551,1159,1186]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="yunnanicus" status="sp. nov.">
<emphasis id="B964EAC65122275350B9E683EFF2FB1E" bold="true" box="[151,428,1159,1186]" italics="true" pageId="1" pageNumber="396">Binodoxys yunnanicus</emphasis>
Davidian
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,
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http://zoobank.org/
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<figureCitation id="132B2A515122275350B9E6CBEE87FB56" box="[151,217,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="1" pageNumber="396">Fig. 1</figureCitation>
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.
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<typeStatus id="54AB8876512227535164E713EFE9FA8E" box="[330,439,1303,1329]" pageId="1" pageNumber="396" type="holotype">Holotype</typeStatus>
: female,
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, h =
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(
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) (
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).
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<emphasis id="B964EAC65122275350E9E75BEF54FAC6" bold="true" box="[199,266,1375,1401]" pageId="1" pageNumber="396">Notes</emphasis>
. The
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was originally preserved in the alcohol (70%), and consequently the integument of the body is partly deformed. During the preparation of the slide and its following investigation and photographing, part of the setae on the dorsal margin of prongs were lost; all these setae were drawn on the
<figureCitation id="132B2A5151222753545BE7A3EA95FA7E" box="[1141,1227,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="1" pageNumber="396">Fig. 1G</figureCitation>
at the locations of the setae points similar to those preserved ones.
</paragraph>
</subSubSection>
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<emphasis id="B964EAC65122275350E9E7EBEF0FF9B6" bold="true" box="[199,337,1519,1545]" pageId="1" pageNumber="396">Description</emphasis>
. Female. Body length 2.0 mm; length of antenna
<quantity id="4CE89B31512227535350E7EBED89F9B5" box="[894,983,1519,1546]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="396" unit="mm" value="1.2">1.2 mm</quantity>
; length of fore wing
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Head (
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) smooth and shiny, with sparse setae, wider than mesoscutum at level of tegulae. Eye large, oval, weakly convergent below (front view), with sparse setae in its lower half. Temple about as long as transverse diameter of eye (dorsal view). Ocelli in acute triangle. POL 2.0 × Od. Clypeus 2.0 × wider than its high. Tentorial index 0.4. Face 1.5 × wider than its height. Antenna with 11 antennomeres, filiform, not widened towards apex, with semi-erect setae, length of setae almost equal to medial width of antennomeres. F1 and F2 same length, their length 4.6 × width in middle. F1 without rhinaria (multiparous plate sensillae), F2 with 2 rhinaria.
</paragraph>
<paragraph id="8BAF36D45122275350E9E4E8EE89F8F1" blockId="1.[151,1437,1303,2014]" pageId="1" pageNumber="396">
Mesosoma. Mesoscutum smooth, shiny, with sparse setae. Notauli present only in anterior one third of mesoscutum. Propodeum (
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) with distinct, wide and delineated by carinae areola. Legs with semi-erect setae.
</paragraph>
<paragraph id="8BAF36D45122275350E9E553EB7EF82A" blockId="1.[151,1437,1303,2014]" pageId="1" pageNumber="396">
Wings (
<figureCitation id="132B2A51512227535132E553EF2DF8CE" box="[284,371,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="1" pageNumber="396">Fig. 1A</figureCitation>
). Fore wing hyaline, its length 2.4 × maximum width. Pterostigma 3.3 × longer than its maximum width and equal to metacarp (1-R1). Radial vein (r) 1.4 × longer than metacarp (1-R1) and pterostigma.
</paragraph>
<paragraph id="8BAF36D45122275050E9E5A4EEA8FF46" blockId="1.[151,1437,1303,2014]" lastBlockId="2.[151,1437,150,429]" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396">
Metasoma. Petiole (
<figureCitation id="132B2A51512227535189E59BEFA2F806" box="[423,508,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="1" pageNumber="396">Fig. 1C</figureCitation>
) narrow, with two pair lateral tubercles, length of petiole 3.6 × its width at level of spiracles. Spiracular tubercles situated on middle of petiole. Basal part of ovipositor sheath dorsally evenly curved, 0.35 × as long as its apical part. Ovipositor sheath (
<figureCitation id="132B2A515121275052F1E293ED6DFF0E" box="[735,819,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="2" pageNumber="397">Fig. 1E</figureCitation>
) 4.0 × longer its maximum width. Prongs (
<figureCitation id="132B2A51512127505531E293EBCAFF0E" box="[1311,1428,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="2" pageNumber="397">Fig. 1F, G</figureCitation>
) 1.5 × longer than ovipositor sheath, with four long setae on its dorsal margin, with one long seta-shaped bristle apically.
</paragraph>
<paragraph id="8BAF36D45121275050E9E307EDBAFEDA" blockId="2.[151,1437,150,429]" pageId="2" pageNumber="397">Colour. Head, antennomeres from third antennomere, mesosoma and most of metasoma dark brown. Scape, pedicel, F1 and F2, clypeus, mouthparts, legs (except dark last segments), petiole, posterior tergites of metasoma with ovipositor sheaths and posterior sternite with prongs yellowish brown.</paragraph>
<paragraph id="8BAF36D45121275050E9E36BEFDBFE36" blockId="2.[151,1437,150,429]" box="[199,389,367,393]" pageId="2" pageNumber="397">
<emphasis id="B964EAC65121275050E9E36BEF5FFE36" box="[199,257,367,393]" italics="true" pageId="2" pageNumber="397">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A655F5121275150E9E397ED85FA35" lastPageId="3" lastPageNumber="398" pageId="2" pageNumber="397" type="biology_ecology">
<paragraph id="8BAF36D45121275050E9E397EFD0FE12" blockId="2.[151,1437,150,429]" box="[199,398,403,429]" pageId="2" pageNumber="397">
<emphasis id="B964EAC65121275050E9E397EF54FE12" bold="true" box="[199,266,403,429]" pageId="2" pageNumber="397">Hosts</emphasis>
. Unknown.
</paragraph>
<caption id="DF6F665C5121275050B9E59CEA70F86A" ID-DOI="http://doi.org/10.5281/zenodo.13757473" ID-Zenodo-Dep="13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="2" pageNumber="397" startId="2.[151,250,1944,1968]" targetBox="[178,1410,463,1900]" targetPageId="2" targetType="figure">
<paragraph id="8BAF36D45121275050B9E59CEA70F86A" blockId="2.[151,1436,1944,2005]" pageId="2" pageNumber="397">
<emphasis id="B964EAC65121275050B9E59CEF4CF80F" bold="true" box="[151,274,1944,1968]" pageId="2" pageNumber="397">FIGURE 1.</emphasis>
Morphological details of
<taxonomicName id="4C104D5751212750523BE59DECA8F80F" authorityName="Davidian &amp; Belokobylskij" authorityYear="2024" box="[533,758,1945,1969]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="yunnanicus" status="sp. nov.">
<emphasis id="B964EAC651212750523BE59DECA8F80F" box="[533,758,1945,1969]" italics="true" pageId="2" pageNumber="397">Binodoxys yunnanicus</emphasis>
</taxonomicName>
<emphasis id="B964EAC65121275052D2E59DED15F80E" bold="true" box="[764,843,1945,1969]" pageId="2" pageNumber="397">
<taxonomicNameLabel id="A25757BD5121275052D2E59DED15F80E" box="[764,843,1945,1969]" pageId="2" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(female, holotype) (
<emphasis id="B964EAC6512127505437E59CEA75F80F" bold="true" box="[1049,1067,1944,1968]" pageId="2" pageNumber="397">A</emphasis>
) Habitus. (
<emphasis id="B964EAC65121275054B5E59DEAF2F80E" bold="true" box="[1179,1196,1945,1969]" pageId="2" pageNumber="397">B</emphasis>
) Head, dorsal view. (
<emphasis id="B964EAC65121275055AFE59CEBCDF80F" bold="true" box="[1409,1427,1944,1968]" pageId="2" pageNumber="397">C</emphasis>
) Propodeum. (
<emphasis id="B964EAC651212750510FE5B9EF6DF86A" bold="true" box="[289,307,1981,2005]" pageId="2" pageNumber="397">D</emphasis>
) Petiole. (
<emphasis id="B964EAC65121275051B0E5B9EFF1F86A" bold="true" box="[414,431,1981,2005]" pageId="2" pageNumber="397">E</emphasis>
) Ovipositor sheaths. (
<emphasis id="B964EAC65121275052BEE5B9ECC1F86A" bold="true" box="[656,671,1981,2005]" pageId="2" pageNumber="397">F</emphasis>
) Prongs (photo). (
<emphasis id="B964EAC6512127505374E5B9ED39F86A" bold="true" box="[858,871,1981,2005]" pageId="2" pageNumber="397">J</emphasis>
) Prongs (drawing).
</paragraph>
</caption>
<caption id="DF6F665C5120275150B9E701EA63FAA2" ID-DOI="http://doi.org/10.5281/zenodo.13757479" ID-Zenodo-Dep="13757479" box="[151,1085,1285,1309]" httpUri="https://zenodo.org/record/13757479/files/figure.png" pageId="3" pageNumber="398" startId="3.[151,250,1285,1309]" targetBox="[189,1398,181,1261]" targetPageId="3" targetType="figure">
<paragraph id="8BAF36D45120275150B9E701EA63FAA2" blockId="3.[151,1085,1285,1309]" box="[151,1085,1285,1309]" pageId="3" pageNumber="398">
<emphasis id="B964EAC65120275150B9E701EF4DFAA2" bold="true" box="[151,275,1285,1309]" pageId="3" pageNumber="398">FIGURE 2.</emphasis>
Photo of the
<taxonomicName id="4C104D575120275151B2E702EFB1FAA2" box="[412,495,1286,1309]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="398" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B964EAC65120275151B2E702EF92FAA2" box="[412,460,1286,1309]" italics="true" pageId="3" pageNumber="398">Acer</emphasis>
sp.
</taxonomicName>
tree, where
<taxonomicName id="4C104D57512027515242E701ED11FAA2" authorityName="Davidian &amp; Belokobylskij" authorityYear="2024" box="[620,847,1285,1309]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="yunnanicus" status="sp. nov.">
<emphasis id="B964EAC6512027515242E701ED11FAA2" box="[620,847,1285,1309]" italics="true" pageId="3" pageNumber="398">Binodoxys yunnanicus</emphasis>
</taxonomicName>
<emphasis id="B964EAC651202751537BE701EDF8FAA2" bold="true" box="[853,934,1285,1309]" pageId="3" pageNumber="398">
<taxonomicNameLabel id="A25757BD51202751537BE701EDF8FAA2" box="[853,934,1285,1309]" pageId="3" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected.
</paragraph>
</caption>
<paragraph id="8BAF36D45120275150E9E76BED85FA35" blockId="3.[151,1437,1391,1993]" box="[199,987,1391,1418]" pageId="3" pageNumber="398">
<emphasis id="B964EAC65120275150E9E76BEF15FA36" bold="true" box="[199,331,1391,1417]" pageId="3" pageNumber="398">Bionomics.</emphasis>
Specimen was collected on the leaves of
<taxonomicName id="4C104D57512027515333E774ED27FA35" box="[797,889,1392,1418]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="398" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B964EAC6512027515333E774ED0CFA35" box="[797,850,1392,1418]" italics="true" pageId="3" pageNumber="398">Acer</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="132B2A515120275153A6E76BED90FA35" box="[904,974,1391,1418]" captionStart="FIGURE 2" captionStartId="3.[151,250,1285,1309]" captionTargetBox="[189,1398,181,1261]" captionTargetId="figure-18@3.[189,1398,181,1261]" captionTargetPageId="3" captionText="FIGURE 2. Photo of the Acer sp. tree, where Binodoxys yunnanicus sp. nov. was collected." figureDoi="http://doi.org/10.5281/zenodo.13757479" httpUri="https://zenodo.org/record/13757479/files/figure.png" pageId="3" pageNumber="398">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A655F5120275150E9E790EEBBF876" pageId="3" pageNumber="398" type="etymology">
<paragraph id="8BAF36D45120275150E9E790EA9BFA12" blockId="3.[151,1437,1391,1993]" box="[199,1221,1427,1454]" pageId="3" pageNumber="398">
<emphasis id="B964EAC65120275150E9E790EF16FA11" bold="true" box="[199,328,1428,1454]" pageId="3" pageNumber="398">Etymology</emphasis>
. Named after
<collectingRegion id="49D4F8365120275151C9E790ECF6FA12" box="[487,680,1427,1454]" country="China" name="Yunnan" pageId="3" pageNumber="398">Yunnan Province</collectingRegion>
of
<collectingCountry id="F30776445120275152E3E797ED4DFA12" box="[717,787,1427,1453]" name="China" pageId="3" pageNumber="398">China</collectingCountry>
, where this new species was collected.
</paragraph>
<paragraph id="8BAF36D45120275150E9E7B3EC70F9DD" blockId="3.[151,1437,1391,1993]" pageId="3" pageNumber="398">
<emphasis id="B964EAC65120275150E9E7B3EF85FA6E" bold="true" box="[199,475,1463,1489]" pageId="3" pageNumber="398">Comparative diagnosis</emphasis>
. This new species is similar to species described from North-East
<collectingCountry id="F30776445120275154FAE7B3EB50FA6E" box="[1236,1294,1463,1489]" name="India" pageId="3" pageNumber="398">India</collectingCountry>
(
<collectingRegion id="49D4F836512027515530E7B3EE8DFA49" country="India" name="Meghalaya" pageId="3" pageNumber="398">Meghalaya State</collectingRegion>
),
<taxonomicName id="4C104D575120275150C2E7DFED24FA49" authority="Samanta &amp; Raychaudhuri, 1990" authorityName="Samanta &amp; Raychaudhuri" authorityYear="1990" box="[236,890,1499,1526]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="trichosiphae">
<emphasis id="B964EAC65120275150C2E7DFEFA6FA4A" box="[236,504,1499,1525]" italics="true" pageId="3" pageNumber="398">Binodoxys trichosiphae</emphasis>
Samanta &amp;
<bibRefCitation id="EF814B255120275152BCE7DFED24FA49" author="Raychaudhuri, D. &amp; Samanta, A. K. &amp; Pramanik, D. R. &amp; Tamili, D. K. &amp; Sarkar, S." box="[658,890,1499,1526]" pageId="3" pageNumber="398" refId="ref3013" refString="Raychaudhuri, D., Samanta, A. K., Pramanik, D. R., Tamili, D. K. &amp; Sarkar, S. (1990) Aphidiids (Hymenoptera) of Northeast India. Indira Publishing House, West Bloomfield, Michigan, 155 pp." type="book" year="1990">Raychaudhuri, 1990</bibRefCitation>
</taxonomicName>
[parasitoid of
<taxonomicName id="4C104D57512027515400E7DFEE8EF9A6" authority="van der Goot, 1916" authorityName="van der Goot" authorityYear="1916" class="Insecta" family="Aphididae" genus="Greenidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="psidii">
<emphasis id="B964EAC6512027515400E7DFEAB3FA4A" box="[1070,1261,1499,1525]" italics="true" pageId="3" pageNumber="398">Greenidea psidii</emphasis>
van der Goot, 1916
</taxonomicName>
) (
<bibRefCitation id="EF814B255120275150C5E7FBEC4BF9A5" author="Raychaudhuri, D. &amp; Samanta, A. K. &amp; Pramanik, D. R. &amp; Tamili, D. K. &amp; Sarkar, S." box="[235,533,1535,1562]" pageId="3" pageNumber="398" refId="ref3013" refString="Raychaudhuri, D., Samanta, A. K., Pramanik, D. R., Tamili, D. K. &amp; Sarkar, S. (1990) Aphidiids (Hymenoptera) of Northeast India. Indira Publishing House, West Bloomfield, Michigan, 155 pp." type="book" year="1990">
Raychaudhuri
<emphasis id="B964EAC65120275151BBE404EF8FF9A6" box="[405,465,1535,1561]" italics="true" pageId="3" pageNumber="398">et al.</emphasis>
1990
</bibRefCitation>
) and
<taxonomicName id="4C104D57512027515275E404EDFFF9A5" authority="Stary, 1978" authorityName="Stary" authorityYear="1978" box="[603,929,1535,1562]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="shillongensis">
<emphasis id="B964EAC6512027515275E404ED4EF9A6" box="[603,784,1535,1561]" italics="true" pageId="3" pageNumber="398">B. shillongensis</emphasis>
<bibRefCitation id="EF814B25512027515332E404EDFFF9A5" author="Stary, P. &amp; Ghosh, A. K." box="[796,929,1536,1562]" pageId="3" pageNumber="398" pagination="77 - 88" refId="ref3066" refString="Stary, P. &amp; Ghosh, A. K. (1978) Further records of aphid parasitoids (Hymenoptera: Aphidiidae) from Meghalaya, India. Oriental Insects, 12, 77 - 88. https: // doi. org / 10.1080 / 00305316.1978.10434553" type="journal article" year="1978">Starý, 1978</bibRefCitation>
</taxonomicName>
(parasitoid of
<taxonomicName id="4C104D5751202751547FE404EFDAF982" authority="(Ghosh &amp; Raychaudhuri, 1968)" baseAuthorityName="Ghosh &amp; Raychaudhuri" baseAuthorityYear="1968" class="Insecta" family="Aphididae" genus="Sinomegoura" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="pyri">
<emphasis id="B964EAC651202751547FE404EB40F9A6" box="[1105,1310,1536,1562]" italics="true" pageId="3" pageNumber="398">Sinomegoura pyri</emphasis>
(Ghosh &amp; Raychaudhuri, 1968)
</taxonomicName>
] (
<bibRefCitation id="EF814B255120275151B7E420ECCAF981" author="Stary, P. &amp; Ghosh, A. K." box="[409,660,1571,1598]" pageId="3" pageNumber="398" pagination="77 - 88" refId="ref3066" refString="Stary, P. &amp; Ghosh, A. K. (1978) Further records of aphid parasitoids (Hymenoptera: Aphidiidae) from Meghalaya, India. Oriental Insects, 12, 77 - 88. https: // doi. org / 10.1080 / 00305316.1978.10434553" type="journal article" year="1978">Starý and Ghosh 1978</bibRefCitation>
). These Indian species are also characterised the presence of the single apical seta-shaped bristle on prongs.
</paragraph>
<paragraph id="8BAF36D45120275150E9E46FED37F94D" blockId="3.[151,1437,1391,1993]" pageId="3" pageNumber="398">
<taxonomicName id="4C104D575120275150E9E46FEF9AF93A" authorityName="Davidian &amp; Belokobylskij" authorityYear="2024" box="[199,452,1643,1669]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="yunnanicus" status="sp. nov.">
<emphasis id="B964EAC65120275150E9E46FEF9AF93A" box="[199,452,1643,1669]" italics="true" pageId="3" pageNumber="398">Binodoxys yunnanicus</emphasis>
</taxonomicName>
<emphasis id="B964EAC65120275151E2E468EC78F939" bold="true" box="[460,550,1644,1670]" pageId="3" pageNumber="398">
<taxonomicNameLabel id="A25757BD5120275151E2E468EC78F939" box="[460,550,1644,1670]" pageId="3" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C104D57512027515290E468ED2EF93A" authorityName="Stary" authorityYear="1978" box="[702,880,1643,1670]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="shillongensis">
<emphasis id="B964EAC6512027515290E468EC91F93A" box="[702,719,1644,1669]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
<emphasis id="B964EAC65120275152F0E46FED2EF93A" box="[734,880,1643,1669]" italics="true" pageId="3" pageNumber="398">shillongensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D57512027515387E468EA08F93A" authorityName="Samanta &amp; Raychaudhuri" authorityYear="1990" box="[937,1110,1643,1670]" class="Insecta" family="Braconidae" genus="Binodoxys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="trichosiphae">
<emphasis id="B964EAC6512027515387E468EDE4F93A" box="[937,954,1644,1669]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
<emphasis id="B964EAC65120275153E4E46FEA08F93A" box="[970,1110,1643,1669]" italics="true" pageId="3" pageNumber="398">trichosiphae</emphasis>
</taxonomicName>
the following characters: F1 and F2 the same length; pterostigma as long as metacarp (1-R1) (
<figureCitation id="132B2A5151202751534BE48BEDE3F915" box="[869,957,1679,1706]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="3" pageNumber="398">Fig. 1A</figureCitation>
); petiole 3.6 × longer than its width at level of spiracular tubercles (
<figureCitation id="132B2A5151202751518CE4B7EFA7F971" box="[418,505,1715,1742]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="3" pageNumber="398">Fig. 1C</figureCitation>
); ovipositor sheath 4.0 × longer than its maximum width (
<figureCitation id="132B2A515120275154BEE4B7EAB8F971" box="[1168,1254,1715,1742]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[178,1410,463,1900]" captionTargetId="figure-108@2.[165,1423,450,1918]" captionTargetPageId="2" captionText="FIGURE 1. Morphological details of Binodoxys yunnanicus sp. nov. (female, holotype) (A) Habitus. (B) Head, dorsal view. (C) Propodeum. (D) Petiole. (E) Ovipositor sheaths. (F) Prongs (photo). (J) Prongs (drawing)." figureDoi="http://doi.org/10.5281/zenodo.13757473" httpUri="https://zenodo.org/record/13757473/files/figure.png" pageId="3" pageNumber="398">Fig. 1E</figureCitation>
); prongs almost straight, with four long setae on their dorsal margin (
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).
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In turn,
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<emphasis id="B964EAC651202751510DE4F8EF6AF8AA" box="[291,308,1788,1813]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
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</taxonomicName>
and
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<emphasis id="B964EAC6512027515221E4F8EC7EF8AA" box="[527,544,1788,1813]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
<emphasis id="B964EAC651202751521EE4FFECE2F8AA" box="[560,700,1787,1813]" italics="true" pageId="3" pageNumber="398">trichosiphae</emphasis>
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have F1 longer than F2, pterostigma 2.0 × longer than metacarp (1-R1), petiole only 3.0 × longer than its width at level of spiracular tubercles. Besides, in
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<emphasis id="B964EAC6512027515492E524EA93F886" box="[1212,1229,1824,1849]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
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the ovipositor sheath is 3.0 × longer than its maximum width, and the prongs sinuate and with three long setae on their dorsal margin and single apical seta-shaped bristle. In
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<emphasis id="B964EAC65120275152D2E56CED53F83E" box="[764,781,1896,1921]" italics="true" pageId="3" pageNumber="398">B</emphasis>
.
<emphasis id="B964EAC6512027515332E563EDF6F83E" box="[796,936,1895,1921]" italics="true" pageId="3" pageNumber="398">trichosiphae</emphasis>
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of the ovipositor sheath is 4.5 × longer than its maximum width, the prongs straight and with two long setae on their dorsal margin and single apical seta-shaped bristle.
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Genus
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Mackauer, 1960
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species.
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<emphasis id="B964EAC65122275350B9E11BEE91FC86" bold="true" box="[151,207,799,825]" pageId="1" pageNumber="396">Note</emphasis>
. Genus
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belongs to the specialised parasitoid of the aphids (
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,
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) and characterise the number of following characters. The head transverse or subcubical; maxillary palps 4-segmented, labial palps 2- segmented; antenna with 10–13 antennomeres in female and 12–15 antennomeres in male. Propodeum usually with closed areola. The venation of fore wing strongly reduced: absent medial (1-SR+M; 2M; 3M), recurrent (m-cu) and both radiomedial (2-SR and r-m) veins. Petiole of metasoma with two pairs of lateral tubercles, of which anterior pair is spiracular tubercles and posterior one is additional tubercles. Ovipositor sheath usually distinctly cut out on lower margin, evenly narrowed to apex. Prongs not fused, straight, weakly curved or sinuate, with one or two apical hair-shaped bristles (
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<treatment id="03B987C25127275650B9E0D8EA60FB5B" ID-DOI="http://doi.org/10.5281/zenodo.13757486" ID-Zenodo-Dep="13757486" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B" httpUri="http://treatment.plazi.org/id/03B987C25127275650B9E0D8EA60FB5B" lastPageNumber="399" pageId="4" pageNumber="399">
<subSubSection id="C30A655F5127275650B9E0D8EFFAFC86" pageId="4" pageNumber="399" type="nomenclature">
<paragraph id="8BAF36D45127275650B9E0D8ECA1FD49" blockId="4.[151,767,732,759]" box="[151,767,732,759]" pageId="4" pageNumber="399">
<heading id="D0E781B85127275650B9E0D8ECA1FD49" bold="true" box="[151,767,732,759]" fontSize="11" level="1" pageId="4" pageNumber="399" reason="1">
<emphasis id="B964EAC65127275650B9E0D8ECA1FD49" bold="true" box="[151,767,732,759]" pageId="4" pageNumber="399">
<taxonomicName id="4C104D575127275650B9E0D8EC44FD49" authority="Tian &amp; Chen" authorityName="Tian &amp; Chen" authorityYear="2017" box="[151,538,732,759]" class="Insecta" family="Braconidae" genus="Astigmapraon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="hei">
<emphasis id="B964EAC65127275650B9E0D8EF31FD48" bold="true" box="[151,367,732,759]" italics="true" pageId="4" pageNumber="399">Astigmapraon hei</emphasis>
Tian &amp; Chen
</taxonomicName>
in
<bibRefCitation id="EF814B2551272756526FE0D8ECA1FD49" author="Tian, H. W. &amp; van Achterberg, C. &amp; Chen, X. X." box="[577,767,732,759]" pageId="4" pageNumber="399" pagination="131 - 134" refId="ref3114" refString="Tian, H. W., van Achterberg, C. &amp; Chen, X. X. (2017) A new genus of the tribe Praini Mackauer (Hymenoptera: Braconidae: Aphidiinae) from China. Zootaxa, 4362 (1), 131 - 134. https: // doi. org / 10.11646 / zootaxa. 4362.1.7" type="journal article" year="2017">
Tian
<emphasis id="B964EAC65127275652ACE0D9ECE3FD48" bold="true" box="[642,701,732,759]" italics="true" pageId="4" pageNumber="399">et al.</emphasis>
2017
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF36D45127275650B9E125EFFAFC86" blockId="4.[151,670,801,856]" box="[151,420,801,825]" pageId="4" pageNumber="399">
<bibRefCitation id="EF814B255127275650B9E125EFFEFC86" author="Tian, H. W. &amp; van Achterberg, C. &amp; Chen, X. X." box="[151,416,801,825]" pageId="4" pageNumber="399" pagination="131 - 134" refId="ref3114" refString="Tian, H. W., van Achterberg, C. &amp; Chen, X. X. (2017) A new genus of the tribe Praini Mackauer (Hymenoptera: Braconidae: Aphidiinae) from China. Zootaxa, 4362 (1), 131 - 134. https: // doi. org / 10.11646 / zootaxa. 4362.1.7" type="journal article" year="2017">
Tian
<emphasis id="B964EAC65127275650E5E126EEA0FC86" box="[203,254,801,825]" italics="true" pageId="4" pageNumber="399">et al.</emphasis>
2017: 132–134
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A655F5127275650B9E144ECC0FCE7" box="[151,670,832,856]" pageId="4" pageNumber="399" type="reference_group">
<paragraph id="8BAF36D45127275650B9E144ECC0FCE7" blockId="4.[151,670,801,856]" box="[151,670,832,856]" pageId="4" pageNumber="399">
<treatmentCitationGroup id="AB0011FA5127275650B9E144ECC0FCE7" box="[151,670,832,856]" pageId="4" pageNumber="399">
<treatmentCitation id="0AB110C55127275650B9E144EC68FCE7" ID-DOI="http://doi.org/10.5281/zenodo.13274362" ID-Zenodo-Dep="13274362" LSID="urn:lsid:zoobank.org:act:129DE716-C0CE-4011-BA16-4574D85939B9" author="Davidian, E. M." box="[151,566,832,856]" httpUri="http://treatment.plazi.org/id/130DA42AFFEEBF443F1EC08BFE8AF81D" page="208" pageId="4" pageNumber="399" year="2020">
<taxonomicName id="4C104D575127275650B9E144EC68FCE7" authority="Davidian, 2020: 208" authorityName="Davidian" authorityPageNumber="208" authorityYear="2020" box="[151,566,832,856]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="longistigmus" status="syn. nov.">
<emphasis id="B964EAC65127275650B9E144EF01FCE7" box="[151,351,832,856]" italics="true" pageId="4" pageNumber="399">Praon longistigmus</emphasis>
<bibRefCitation id="EF814B2551272756514BE144EC68FCE7" author="Davidian, E. M." box="[357,566,832,856]" pageId="4" pageNumber="399" pagination="207 - 211" refId="ref2958" refString="Davidian, E. M. (2020) New species of the genus Praon Haliday, 1833 (Hymenoptera: Aphidiidae) from China. Caucasian Entomological Bulletin, 16 (2), 207 - 211. [in Russian] https: // doi. org / 10.23885 / 181433262020162 - 207211" type="journal article" year="2020">Davidian, 2020: 208</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B964EAC651272756526FE144ECC0FCE7" bold="true" box="[577,670,832,856]" pageId="4" pageNumber="399">
<taxonomicNameLabel id="A25757BD51272756526FE144ECC0FCE7" box="[577,670,832,856]" pageId="4" pageNumber="399">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30A655F5127275650B9E182EA60FB5B" pageId="4" pageNumber="399" type="discussion">
<paragraph id="8BAF36D45127275650B9E182EAF7FBB3" blockId="4.[151,1436,902,1254]" pageId="4" pageNumber="399">
<emphasis id="B964EAC65127275650B9E182EF5AFC1F" bold="true" box="[151,260,902,928]" pageId="4" pageNumber="399">Remarks</emphasis>
. A study of the original description of
<taxonomicName id="4C104D57512727565281E182ECB1FC1F" authorityName="Tian &amp; Chen" authorityYear="2017" box="[687,751,902,928]" class="Insecta" family="Braconidae" genus="Astigmapraon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="hei">
<emphasis id="B964EAC6512727565281E182ECB1FC1F" box="[687,751,902,928]" italics="true" pageId="4" pageNumber="399">A. hei</emphasis>
</taxonomicName>
shows that there are no significant morphological differences between it and
<taxonomicName id="4C104D5751272756517EE1AFECF5FC7B" authority="Davidian." authorityName="Davidian." box="[336,683,938,964]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="longistigmus">
<emphasis id="B964EAC651272756517EE1AFEC6EFC7B" box="[336,560,938,964]" italics="true" pageId="4" pageNumber="399">Praon longistigmus</emphasis>
Davidian.
</taxonomicName>
The only difference is that the male antenna of
<taxonomicName id="4C104D575127275654DFE1AEEB69FC7B" authorityName="Tian &amp; Chen" authorityYear="2017" box="[1265,1335,938,964]" class="Insecta" family="Braconidae" genus="Astigmapraon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="hei">
<emphasis id="B964EAC65127275654DFE1AEEB69FC7B" box="[1265,1335,938,964]" italics="true" pageId="4" pageNumber="399">A. hei</emphasis>
</taxonomicName>
with 20 antennomeres, but in
<taxonomicName id="4C104D575127275651ACE1CBEC75FC57" box="[386,555,974,1000]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="longistigmus">
<emphasis id="B964EAC65127275651ACE1CBEC75FC57" box="[386,555,974,1000]" italics="true" pageId="4" pageNumber="399">P. longistigmus</emphasis>
</taxonomicName>
there are 21 antennomeres. On this basis, the following synonymy is established here:
<taxonomicName id="4C104D575127275650F8E1F6EC25FBB3" authority="Tian et Chen, 2017" authorityName="Tian et Chen" authorityYear="2017" box="[214,635,1010,1036]" class="Insecta" family="Braconidae" genus="Astigmapraon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="hei">
<emphasis id="B964EAC65127275650F8E1F6EFC0FBB3" box="[214,414,1010,1036]" italics="true" pageId="4" pageNumber="399">Astigmapraon hei</emphasis>
<bibRefCitation id="EF814B2551272756518BE1F6EC25FBB3" author="Tian, H. W. &amp; van Achterberg, C. &amp; Chen, X. X." box="[421,635,1010,1036]" pageId="4" pageNumber="399" pagination="131 - 134" refId="ref3114" refString="Tian, H. W., van Achterberg, C. &amp; Chen, X. X. (2017) A new genus of the tribe Praini Mackauer (Hymenoptera: Braconidae: Aphidiinae) from China. Zootaxa, 4362 (1), 131 - 134. https: // doi. org / 10.11646 / zootaxa. 4362.1.7" type="journal article" year="2017">Tian et Chen, 2017</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C104D575127275652B6E1F7EA7FFBB3" authority="Davidian 2020" authorityName="Davidian" authorityYear="2020" box="[664,1057,1010,1036]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="longistigmus">
<emphasis id="B964EAC65127275652B6E1F7ED2AFBB3" box="[664,884,1010,1036]" italics="true" pageId="4" pageNumber="399">Praon longistigmus</emphasis>
<bibRefCitation id="EF814B25512727565352E1F6EA7FFBB3" author="Davidian, E. M." box="[892,1057,1010,1036]" pageId="4" pageNumber="399" pagination="207 - 211" refId="ref2958" refString="Davidian, E. M. (2020) New species of the genus Praon Haliday, 1833 (Hymenoptera: Aphidiidae) from China. Caucasian Entomological Bulletin, 16 (2), 207 - 211. [in Russian] https: // doi. org / 10.23885 / 181433262020162 - 207211" type="journal article" year="2020">Davidian 2020</bibRefCitation>
</taxonomicName>
(
<emphasis id="B964EAC651272756541CE1F6EAC7FBB3" bold="true" box="[1074,1177,1010,1036]" pageId="4" pageNumber="399">syn. nov.</emphasis>
).
</paragraph>
<paragraph id="8BAF36D45127275650E9E612EA60FB5B" blockId="4.[151,1436,902,1254]" pageId="4" pageNumber="399">
<materialsCitation id="3B783C895127275650E9E612EA64FB5B" collectingDate="2006-07-25" collectingDateMax="2013-08-21" collectingDateMin="2006-07-25" collectionCode="ZISP" collectorName="Tu Bin-Bin, No. &amp; Zhang Hong-Ying, No. &amp; I. Belousov &amp; G. Davidian &amp; I. Kabak" country="China" county="Pingwu" elevation="2770" latitude="33.346943" location="Additionally" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" pageId="4" pageNumber="399" specimenCount="3" specimenCount-male="3" stateProvince="Shaanxi" typeStatus="holotype">
<location id="8ECF600F5127275650E9E612EF08FB8F" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B:8ECF600F5127275650E9E612EF08FB8F" box="[199,342,1046,1072]" country="China" county="Pingwu" latitude="33.346943" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" name="Additionally" pageId="4" pageNumber="399" stateProvince="Shaanxi">Additionally</location>
, both taxa were described from same region.
<location id="8ECF600F51272756536CE612ED30FB8F" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B:8ECF600F51272756536CE612ED30FB8F" box="[834,878,1046,1072]" country="China" county="Pingwu" latitude="33.346943" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" name="The" pageId="4" pageNumber="399" stateProvince="Shaanxi">The</location>
type specimens of
<taxonomicName id="4C104D57512727565410E612EA23FB8F" authorityName="Tian et Chen" authorityYear="2017" box="[1086,1149,1046,1072]" class="Insecta" family="Braconidae" genus="Astigmapraon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="hei">
<emphasis id="B964EAC6512727565410E612EA23FB8F" box="[1086,1149,1046,1072]" italics="true" pageId="4" pageNumber="399">A. hei</emphasis>
</taxonomicName>
were collected in
<collectingRegion id="49D4F83651272756556CE612EBC2FB8F" box="[1346,1436,1046,1072]" country="China" name="Shaanxi" pageId="4" pageNumber="399">Shaanxi</collectingRegion>
and
<collectingRegion id="49D4F8365127275650E4E63EEF7AFBEB" box="[202,292,1082,1108]" country="China" name="Sichuan" pageId="4" pageNumber="399">Sichuan</collectingRegion>
<location id="8ECF600F512727565101E63EEFFCFBEB" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B:8ECF600F512727565101E63EEFFCFBEB" box="[303,418,1082,1108]" country="China" county="Pingwu" latitude="33.346943" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" name="Provinces" pageId="4" pageNumber="399" stateProvince="Shaanxi">Provinces</location>
:
<typeStatus id="54AB887651272756519EE63EEC4BFBEB" box="[432,533,1082,1108]" pageId="4" pageNumber="399" type="holotype">holotype</typeStatus>
, female, N.
<collectingCountry id="F3077644512727565289E63EECB3FBEB" box="[679,749,1082,1108]" name="China" pageId="4" pageNumber="399">China</collectingCountry>
,
<collectingRegion id="49D4F8365127275652D5E63EED06FBEB" box="[763,856,1082,1108]" country="China" name="Shaanxi" pageId="4" pageNumber="399">Shaanxi</collectingRegion>
,
<location id="8ECF600F512727565349E63EEDF9FBEB" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B:8ECF600F512727565349E63EEDF9FBEB" box="[871,935,1082,1108]" country="China" county="Pingwu" latitude="33.346943" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" name="Baoji" pageId="4" pageNumber="399" stateProvince="Shaanxi">Baoji</location>
,
<date id="FFAE101451272756539BE63EEA14FBEB" box="[949,1098,1082,1108]" pageId="4" pageNumber="399" value="2013-08-21">
<collectingDate id="EFEAE9FC51272756539BE63EEA14FBEB" box="[949,1098,1082,1108]" pageId="4" pageNumber="399" value="2013-08-21">21.VIII.2013</collectingDate>
</date>
,
<collectorName id="26E55302512727565477E63EEB4BFBEA" box="[1113,1301,1082,1109]" pageId="4" pageNumber="399">Tu Bin-Bin, No.</collectorName>
201306561;
<typeStatus id="54AB88765127275650B9E65AEF59FBC7" box="[151,263,1118,1144]" pageId="4" pageNumber="399" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D16FD5D51272756513CE65AEF3EFBC7" box="[274,352,1118,1144]" count="1" pageId="4" pageNumber="399" type="male">1 male</specimenCount>
, S.
<collectingCountry id="F30776445127275651A4E65AEF8EFBC7" box="[394,464,1118,1144]" name="China" pageId="4" pageNumber="399">China</collectingCountry>
,
<collectingRegion id="49D4F8365127275651F5E65AEC66FBC7" box="[475,568,1118,1144]" country="China" name="Sichuan" pageId="4" pageNumber="399">Sichuan</collectingRegion>
,
<location id="8ECF600F51272756526AE65AECC2FBC7" LSID="urn:lsid:plazi:treatment:03B987C25127275650B9E0D8EA60FB5B:8ECF600F51272756526AE65AECC2FBC7" box="[580,668,1118,1144]" country="China" county="Pingwu" latitude="33.346943" longLatPrecision="20" longitude="103.80862" municipality="Gao Zhi-Lei" name="Pingwu" pageId="4" pageNumber="399" stateProvince="Shaanxi">Pingwu</location>
,
<date id="FFAE1014512727565286E65AED6DFBC6" box="[680,819,1118,1145]" pageId="4" pageNumber="399" value="2006-07-25">
<collectingDate id="EFEAE9FC512727565286E65AED6DFBC6" box="[680,819,1118,1145]" pageId="4" pageNumber="399" value="2006-07-25">25.VII.2006</collectingDate>
</date>
,
<collectingMunicipality id="6BCBACAE512727565311E65AED93FBC7" box="[831,973,1118,1144]" pageId="4" pageNumber="399">Gao Zhi-Lei</collectingMunicipality>
, 200615767;
<specimenCount id="9D16FD5D512727565449E65AEA9EFBC7" box="[1127,1216,1118,1144]" count="2" pageId="4" pageNumber="399" type="male">2 males</specimenCount>
, S.
<collectingCountry id="F30776445127275654C4E65AEB6EFBC7" box="[1258,1328,1118,1144]" name="China" pageId="4" pageNumber="399">China</collectingCountry>
,
<collectingRegion id="49D4F836512727565515E65AEBC6FBC7" box="[1339,1432,1118,1144]" country="China" name="Sichuan" pageId="4" pageNumber="399">Sichuan</collectingRegion>
,
<collectingCounty id="62CE4E585127275650B9E686EEB1FB23" box="[151,239,1154,1180]" pageId="4" pageNumber="399">Pingwu</collectingCounty>
,
<date id="FFAE10145127275650D1E686EFD4FB22" box="[255,394,1154,1181]" pageId="4" pageNumber="399" value="2006-07-25">
<collectingDate id="EFEAE9FC5127275650D1E686EFD4FB22" box="[255,394,1154,1181]" pageId="4" pageNumber="399" value="2006-07-25">25.VII.2006</collectingDate>
</date>
,
<collectorName id="26E553025127275651B7E686ECFBFB23" box="[409,677,1154,1180]" pageId="4" pageNumber="399">Zhang Hong-Ying, No.</collectorName>
200614125, 200614768.
<taxonomicName id="4C104D575127275653E1E687EAF1FB23" box="[975,1199,1154,1180]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="longistigmus">
<emphasis id="B964EAC65127275653E1E687EAF1FB23" box="[975,1199,1154,1180]" italics="true" pageId="4" pageNumber="399">Praon longistigmus</emphasis>
</taxonomicName>
was described from a single male with following label:
<typeStatus id="54AB8876512727565212E6A2ECFFFB7F" box="[572,673,1190,1216]" pageId="4" pageNumber="399" type="holotype">holotype</typeStatus>
, male,
<collectingCountry id="F30776445127275652D7E6A2ED61FB7F" box="[761,831,1190,1216]" name="China" pageId="4" pageNumber="399">China</collectingCountry>
, N of
<collectingRegion id="49D4F8365127275653A1E6A2EA05FB7F" box="[911,1115,1190,1216]" country="China" name="Sichuan" pageId="4" pageNumber="399">Sichuan Province</collectingRegion>
,
<locationDeviation id="9A1D2FF6512727565444E6A2EB73FB7F" box="[1130,1325,1190,1216]" location="Jiuzhaigou" pageId="4" pageNumber="399">W of Jiuzhaigou</locationDeviation>
,
<quantity id="4CE89B31512727565512E6A2EBC9FB7F" box="[1340,1431,1190,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.77" pageId="4" pageNumber="399" unit="m" value="2770.0">
<elevation id="003DD1E7512727565512E6A2EBC9FB7F" box="[1340,1431,1190,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.77" pageId="4" pageNumber="399" unit="m" value="2770.0">2770 m</elevation>
</quantity>
,
<geoCoordinate id="EE2450135127275650B9E6CFEF40FB59" box="[151,286,1227,1254]" degrees="33" direction="north" minutes="20" orientation="latitude" pageId="4" pageNumber="399" precision="15" seconds="49" value="33.346943">33°20ʹ49ʺN</geoCoordinate>
,
<geoCoordinate id="EE245013512727565107E6CFEFE4FB59" box="[297,442,1227,1254]" degrees="103" direction="east" minutes="48" orientation="longitude" pageId="4" pageNumber="399" precision="15" seconds="31" value="103.80862">103°48ʹ31ʺE</geoCoordinate>
,
<date id="FFAE10145127275651E8E6CEEC1BFB5B" box="[454,581,1226,1252]" pageId="4" pageNumber="399" value="2012-06-21">
<collectingDate id="EFEAE9FC5127275651E8E6CEEC1BFB5B" box="[454,581,1226,1252]" pageId="4" pageNumber="399" value="2012-06-21">21.VI.2012</collectingDate>
</date>
(
<collectorName id="26E55302512727565278E6CEEC86FB5B" box="[598,728,1226,1252]" pageId="4" pageNumber="399">I. Belousov</collectorName>
,
<collectorName id="26E553025127275652CDE6CEED31FB5B" box="[739,879,1226,1252]" pageId="4" pageNumber="399">G. Davidian</collectorName>
,
<collectorName id="26E55302512727565354E6CEED82FB5B" box="[890,988,1226,1252]" pageId="4" pageNumber="399">I. Kabak</collectorName>
) (
<collectionCode id="ED01AE115127275653DCE6CEEA6FFB5B" box="[1010,1073,1226,1252]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="399">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="9DA8546FF8568C5A3D56B66914263705" ID-DOI="10.11646/zootaxa.5507.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757579" ID-ZooBank="C5A55C22-3D3B-4B49-9591-148B0FB7E8C3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726221267114" checkinUser="plazi" docAuthor="Telnov, Dmitry" docDate="2024" docId="03C351407B2BFFE4FF1C8C3E8337F90C" docLanguage="en" docName="zootaxa.5507.2.4.pdf" docOrigin="Zootaxa 5507 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nitorus sathulaz Telnov, 2024, sp. nov." docType="treatment" docUuid="DF37A008-F628-46AC-B811-B25072B2ADF7" docUuidSource="ZooBank" docVersion="2" lastPageNumber="308" masterDocId="FFFA29387B3EFFF3FF8B8A788038FFB1" masterDocTitle="New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum" masterLastPageNumber="312" masterPageNumber="285" pageNumber="306" updateTime="1726221726128" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1495E24F6546AD048F8F2EAC60A0FB7D">New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum</mods:title>
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<mods:namePart id="9ADBF47F2FE9E3EE1D044B8119AE9953">Telnov, Dmitry</mods:namePart>
<mods:affiliation id="A0E61B850212FA1CD8C64F8FCE90E5B8">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD London, United Kingdom &amp; Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401 Daugavpils, Latvia &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004 Rīga, Latvia</mods:affiliation>
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<mods:title id="5128397347BF660BD26BD78FCF01D51E">Zootaxa</mods:title>
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<mods:date id="E8680B36BF581008873D379FF434EC9B">2024</mods:date>
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<mods:number id="18E58FFD260E303900C1EDEA4486F65C">2024-09-12</mods:number>
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<subSubSection id="C370B3DD7B2BFFE6FF1C8C3E83F2F934" pageId="21" pageNumber="306" type="nomenclature">
<paragraph id="8BD5E0567B2BFFE6FF1C8C3E82B5F9D1" blockId="21.[151,970,1606,1669]" box="[151,653,1606,1633]" pageId="21" pageNumber="306">
<heading id="D09D573A7B2BFFE6FF1C8C3E82B5F9D1" bold="true" box="[151,653,1606,1633]" fontSize="11" level="1" pageId="21" pageNumber="306" reason="1">
<emphasis id="B91E3C447B2BFFE6FF1C8C3E82B5F9D1" bold="true" box="[151,653,1606,1633]" pageId="21" pageNumber="306">
<taxonomicName id="4C6A9BD57B2BFFE6FF1C8C3E8167F9D0" authorityName="Telnov" authorityYear="2024" box="[151,351,1606,1633]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="306" phylum="Arthropoda" rank="species" species="sathulaz" status="sp. nov.">
<emphasis id="B91E3C447B2BFFE6FF1C8C3E8167F9D0" bold="true" box="[151,351,1606,1633]" italics="true" pageId="21" pageNumber="306">Nitorus sathulaz</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2BFFE6FEED8C3F81FCF9D0" box="[358,452,1607,1633]" pageId="21" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1351FCD37B2BFFE6FE588C3E82BDF9D1" box="[467,645,1606,1632]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="22.[151,250,1476,1500]" captionStartId-1="24.[151,250,1560,1584]" captionTargetBox-0="[163,1423,193,1440]" captionTargetBox-1="[424,1171,197,1508]" captionTargetId-0="figure-28@22.[151,1435,181,1452]" captionTargetId-1="figure-28@24.[384,1202,181,1537]" captionTargetPageId-0="22" captionTargetPageId-1="24" captionText-0="FIGURE 17. Nitorus sathulaz sp. nov., holotype ♂. A—Habitus, dorsal view; B—ditto, lateral view." captionText-1="FIGURE 18. Nitorus sathulaz sp. nov., paratype ♂. A—Aedeagus; B—ditto, apex, magnified; C—ditto, median portion, magnified [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.13757631" figureDoi-1="http://doi.org/10.5281/zenodo.13757637" httpUri-0="https://zenodo.org/record/13757631/files/figure.png" httpUri-1="https://zenodo.org/record/13757637/files/figure.png" pageId="21" pageNumber="306">Figures 17–18</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2BFFE6FF1C8CCB828AF8A4" pageId="21" pageNumber="306" type="materials_examined">
<paragraph id="8BD5E0567B2BFFE6FF1C8CCB857BF940" blockId="21.[151,1436,1714,1813]" pageId="21" pageNumber="306">
<emphasis id="B91E3C447B2BFFE6FF1C8CCB81F6F97D" bold="true" box="[151,462,1714,1741]" pageId="21" pageNumber="306">Type material designated.</emphasis>
<materialsCitation id="3B02EA0B7B2BFFE6FE5C8CCA857BF940" collectingDate="1996-05-30" collectingDateMax="1996-06-14" collectingDateMin="1996-05-30" collectionCode="NME" collectorName="Na Hang &amp; Pac Ban &amp; A. Napolov" elevation="900" location="Hanoi, Tuyen Quang prov." pageId="21" pageNumber="306" specimenCount="1" specimenCount-male="1" stateProvince="Hanoi, Tuyen Quang prov." typeStatus="holotype">
<typeStatus id="54D15EF47B2BFFE6FE5C8CCA8207F97D" box="[471,575,1714,1740]" pageId="21" pageNumber="306" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6C2BDF7B2BFFE6FDCC8CCB8266F97C" box="[583,606,1715,1741]" count="1" pageId="21" pageNumber="306" type="male">♂</specimenCount>
,
<collectionCode id="ED7B78937B2BFFE6FDE78CCB8297F97C" box="[620,687,1715,1741]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="21" pageNumber="306">NME</collectionCode>
: Vietnam- N,
<quantity id="4C924DB37B2BFFE6FCC58CCA839CF97D" box="[846,932,1714,1740]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="306" unit="km" value="160.0">160 km</quantity>
NNW
<collectingRegion id="49AE2EB47B2BFFE6FC7C8CCA8510F97C" box="[1015,1320,1714,1741]" isEnumeration="true" pageId="21" pageNumber="306">Hanoi, Tuyen Quang prov.</collectingRegion>
,
<quantity id="4C924DB37B2BFFE6FABF8CCB8557F97D" box="[1332,1391,1714,1741]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="306" unit="km" value="3.0">3 km</quantity>
NE from
<collectorName id="269F85807B2BFFE6FF5E8CAF8103F940" box="[213,315,1751,1777]" pageId="21" pageNumber="306">Na Hang</collectorName>
,
<collectorName id="269F85807B2BFFE6FECC8CAF819BF940" box="[327,419,1751,1777]" pageId="21" pageNumber="306">Pac Ban</collectorName>
vill. env.,
<quantity id="4C924DB37B2BFFE6FD928CAF825AF940" box="[537,610,1751,1777]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="306" unit="m" value="900.0">
<elevation id="004707657B2BFFE6FD928CAF825AF940" box="[537,610,1751,1777]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="306" unit="m" value="900.0">900 m</elevation>
</quantity>
, primary rain forest,
<date id="FFD4C6967B2BFFE6FCC58CAF842DF941" box="[846,1045,1750,1777]" pageId="21" pageNumber="306" value="1996-05-30" valueMax="1996-06-14" valueMin="1996-05-30">
<collectingDate id="EF903F7E7B2BFFE6FCC58CAF842DF941" box="[846,1045,1750,1777]" pageId="21" pageNumber="306" value="1996-05-30" valueMax="1996-06-14" valueMin="1996-05-30">30.V–14.VI.1996</collectingDate>
</date>
, leg.
<collectorName id="269F85807B2BFFE6FBDB8CAF84EAF940" box="[1104,1234,1751,1777]" pageId="21" pageNumber="306">A. Napolov</collectorName>
[printed].
</materialsCitation>
</paragraph>
<paragraph id="8BD5E0567B2BFFE6FF4C8C83828AF8A4" blockId="21.[151,1436,1714,1813]" box="[199,690,1787,1813]" pageId="21" pageNumber="306">
<typeStatus id="54D15EF47B2BFFE6FF4C8C83810CF8A4" box="[199,308,1787,1813]" pageId="21" pageNumber="306" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D6C2BDF7B2BFFE6FEB08C838166F8A4" box="[315,350,1787,1813]" count="2" pageId="21" pageNumber="306" type="male">2♂</specimenCount>
DTC: same label as
<typeStatus id="54D15EF47B2BFFE6FDC38C838296F8A4" box="[584,686,1787,1813]" pageId="21" pageNumber="306" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF15B0DE7B28FFE5FF1C8FBC84AFFA6D" ID-DOI="http://doi.org/10.5281/zenodo.13757631" ID-Zenodo-Dep="13757631" box="[151,1175,1476,1500]" httpUri="https://zenodo.org/record/13757631/files/figure.png" pageId="22" pageNumber="307" startId="22.[151,250,1476,1500]" targetBox="[163,1423,193,1440]" targetPageId="22" targetType="figure">
<paragraph id="8BD5E0567B28FFE5FF1C8FBC84AFFA6D" blockId="22.[151,1175,1476,1500]" box="[151,1175,1476,1500]" pageId="22" pageNumber="307">
<emphasis id="B91E3C447B28FFE5FF1C8FBC8118FA6D" bold="true" box="[151,288,1476,1500]" pageId="22" pageNumber="307">FIGURE 17.</emphasis>
<taxonomicName id="4C6A9BD57B28FFE5FEAD8FBD81F4FA6D" authorityName="Telnov" authorityYear="2024" box="[294,460,1476,1500]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="307" phylum="Arthropoda" rank="species" species="sathulaz" status="sp. nov.">
<emphasis id="B91E3C447B28FFE5FEAD8FBD81F4FA6D" box="[294,460,1476,1500]" italics="true" pageId="22" pageNumber="307">Nitorus sathulaz</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B28FFE5FE588FBD821CFA6D" box="[467,548,1477,1500]" pageId="22" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂. A—Habitus, dorsal view; B—ditto, lateral view.
</paragraph>
</caption>
<subSubSection id="C370B3DD7B28FFE5FF4C8C56825CF9DD" pageId="22" pageNumber="307" type="etymology">
<paragraph id="8BD5E0567B28FFE5FF4C8C56825CF9DD" blockId="22.[151,1436,1582,1789]" pageId="22" pageNumber="307">
<emphasis id="B91E3C447B28FFE5FF4C8C568190F9F9" bold="true" box="[199,424,1582,1608]" pageId="22" pageNumber="307">Derivatio nominis.</emphasis>
Named from Proto-Germanic ‘sathulaz’ (saddle) to point on the saddle-like impressed, bigibbose pronotum. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B28FFE4FF4C8C0E82BFFA54" lastPageId="23" lastPageNumber="308" pageId="22" pageNumber="307" type="description">
<paragraph id="8BD5E0567B28FFE5FF4C8C0E8284F94D" blockId="22.[151,1436,1582,1789]" pageId="22" pageNumber="307">
<emphasis id="B91E3C447B28FFE5FF4C8C0E814EF921" bold="true" box="[199,374,1654,1680]" pageId="22" pageNumber="307">Measurements</emphasis>
,
<typeStatus id="54D15EF47B28FFE5FE0E8C0E81DFF921" box="[389,487,1654,1680]" pageId="22" pageNumber="307" type="holotype">holotype</typeStatus>
male, total body length excluding partially exposed terminalia
<quantity id="4C924DB37B28FFE5FB3F8C0E8535F920" box="[1204,1293,1654,1681]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="307" unit="mm" value="2.2">2.2 mm</quantity>
; head length
<quantity id="4C924DB37B28FFE5FF1C8CE280C9F904" box="[151,241,1690,1717]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="307" unit="mm" value="0.6">0.6 mm</quantity>
, head width across compound eyes
<quantity id="4C924DB37B28FFE5FD048CE282D1F904" box="[655,745,1690,1717]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="307" unit="mm" value="0.5">0.5 mm</quantity>
, pronotal length
<quantity id="4C924DB37B28FFE5FC278CE2843EF904" box="[940,1030,1690,1717]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="307" unit="mm" value="0.5">0.5 mm</quantity>
, maximum pronotal width
<quantity id="4C924DB37B28FFE5FAB58CE285A0F904" box="[1342,1432,1690,1717]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="307" unit="mm" value="0.4">0.4 mm</quantity>
, minimum pronotal width in basal half
<quantity id="4C924DB37B28FFE5FDDD8CC68288F968" box="[598,688,1726,1753]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="307" unit="mm" value="0.2">0.2 mm</quantity>
, elytral length
<quantity id="4C924DB37B28FFE5FCEB8CC68382F968" box="[864,954,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="307" unit="mm" value="1.1">1.1 mm</quantity>
, combined maximum elytral width across midlength
<quantity id="4C924DB37B28FFE5FE9B8C9A8151F94C" box="[272,361,1762,1789]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="307" unit="mm" value="0.8">0.8 mm</quantity>
. Male
<typeStatus id="54D15EF47B28FFE5FE3F8C9B8227F94C" box="[436,543,1763,1789]" pageId="22" pageNumber="307" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B28FFE5FDAD8C9A8244F94C" box="[550,636,1762,1789]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="307" unit="mm" value="2.4">2.4 mm</quantity>
long.
</paragraph>
<paragraph id="8BD5E0567B29FFE4FF4C8AEE8269FA70" blockId="23.[151,1437,150,1725]" pageId="23" pageNumber="308">
<emphasis id="B91E3C447B29FFE4FF4C8AEE8161FF01" bold="true" box="[199,345,150,176]" pageId="23" pageNumber="308">Description.</emphasis>
<typeStatus id="54D15EF47B29FFE4FED48AEF81FFFF00" box="[351,455,151,177]" pageId="23" pageNumber="308" type="holotype">Holotype</typeStatus>
male. Dorsal forebody dark rufous, ventral—pale rufous. Elytra narrowly dark rufous at base including scutellar shield, rest of surface black-brown. Antennomeres 1–6 pale rufous to yellowish, remaining antennomeres dark rufous to brown. Maxillary palpus dark rufous, mesal margin of terminal palpomere pale. Legs dark rufous, tarsi pale rufous to yellowish. Abdomen brown. Head elliptical, slightly longer than wide, slightly convex in dorsal aspect, moderately glossy. Labrum subtruncate at anterior margin. Frontoclypeal suture present, slightly arched (shallowly concave anteriorly). Compound eye small, moderately protruding from lateral outline of head, shorter than rather strongly converging tempus. Head base narrowly rounded. Head dorsal punctures large, irregularly shaped, dense and deep. Intervening spaces on anterior head microreticulate and much narrower than punctures, on posterior head smooth and glossy, generally slightly narrower than punctures. Dorsal cranial setae whitish, sparse, inconspicuous. Antenna extending towards base of elytra when directed posteriad, thickened in apical half. Antennomere three about 1.4× as long as antennomere two. Antennomeres 4 and 5 about same long, each slightly shorter than antennomere three. Antennomeres 8–11 thickened, of them 9–10 distinctly widened distally. Penultimate antennomere, somewhat shortened, about as long as wide. Terminal antennomere slightly asymmetrical, shortly conical, bluntly pointed, about 1.6–1.7× as long as penultimate antennomere, distinctly shorter the combined length of antennomeres 9–10. Penultimate maxillary palpomere slightly lobate at mesal margin distally. Terminal maxillary palpomere strongly securiform. Cranial ‘neck’ less than a third head width across compound eyes, smooth and glossy. Pronotum longer than wide, distinctly narrower than head across eyes, truncate medially at anterior margin. Pronotal disc glossy, bigibbose in dorsal aspect with a deep, saddle-like discal impression separating anterior and basal lobes of pronotum. Lateral margins of anterior lobe strongly protruding laterad (subangular), of basal lobe—widened towards base.Anterior pronotal rim distinct, wide dorsally, laterally and ventrally. Antebasal sulcus moderately wide dorsally, wide laterally. Pronotal punctures present mainly on anterior and basal pronotal humps only, smaller than those on head. Intervening spaces glossy and glabrous, generally as wide as punctures. Basal lobe with large and dense but shallow punctures, intervening spaces glossy and glabrous, narrower than to as wide as punctures. Lateral sides of anterior lobe glossy and glabrous, impunctured, of basal lobe and constriction area—glossy and glabrous, with dense subparallel longitudinal wrinkles. Pronotal setae as those on head, inconspicuous. Scutellar shield lanceolate, apically rounded, glossy and glabrous. Elytra elliptical, about 1.3–1.4× as long as wide, widest at midlength, dorsally strongly convex. Postbasal transverse impression not indicated. Humerus broadly rounded, humeral callosity slightly projecting in dorso-lateral aspect. Apical sutural angle rounded. Elytral surface glossy and smooth, punctures minute, shallow, sparse. Intervening spaces smooth and glossy, much wider than punctures. Elytral setae whitish, rather short, sparse, not fully appressed, inconspicuous. Scattered longer erect tactile setae on elytral disc. Metathoracic wing fully developed (functional). Legs moderately long, sparsely minutely setose. All tibiae somewhat thickened. Protibia somewhat sinuous on both anterior and posterior margin. Metatibia slightly sinuous in distal half, dorsal surface in posterior two-thirds slightly flattened, here densely punctured and with numerous whitish suberect setae. Tibial terminal spurs paired, very short. Basal metatarsomere longer than combined length of remaining metatarsomeres. Penultimate tarsomeres slightly bilobate. Tergite VII truncate at posterior margin. Morphological sternite VII broad, rounded at posterior margin. Aedeagus (
<figureCitation id="1351FCD37B29FFE4FF148FDF80C9FA70" box="[159,241,1447,1473]" captionStart="FIGURE 18" captionStartId="24.[151,250,1560,1584]" captionTargetBox="[424,1171,197,1508]" captionTargetId="figure-28@24.[384,1202,181,1537]" captionTargetPageId="24" captionText="FIGURE 18. Nitorus sathulaz sp. nov., paratype ♂. A—Aedeagus; B—ditto, apex, magnified; C—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757637" httpUri="https://zenodo.org/record/13757637/files/figure.png" pageId="23" pageNumber="308">Fig. 18</figureCitation>
) elongate, apex trident-shaped.
</paragraph>
<paragraph id="8BD5E0567B29FFE4FF4C8FB282BFFA54" blockId="23.[151,1437,150,1725]" box="[199,647,1482,1509]" pageId="23" pageNumber="308">
<emphasis id="B91E3C447B29FFE4FF4C8FB2818FFA55" bold="true" box="[199,439,1482,1508]" pageId="23" pageNumber="308">Sexual dimorphism.</emphasis>
Female unknown.
</paragraph>
</subSubSection>
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<paragraph id="8BD5E0567B29FFE4FF4C8F9681C9F9C4" blockId="23.[151,1437,150,1725]" pageId="23" pageNumber="308">
<emphasis id="B91E3C447B29FFE4FF4C8F9681F7F9B9" bold="true" box="[199,463,1518,1544]" pageId="23" pageNumber="308">Differential diagnosis.</emphasis>
<taxonomicName id="4C6A9BD57B29FFE4FE5C8F9782A9F9B8" authorityName="Telnov" authorityYear="2024" box="[471,657,1519,1545]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="308" phylum="Arthropoda" rank="species" species="sathulaz" status="sp. nov.">
<emphasis id="B91E3C447B29FFE4FE5C8F9782A9F9B8" box="[471,657,1519,1545]" italics="true" pageId="23" pageNumber="308">Nitorus sathulaz</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B29FFE4FD118F9782CAF9B8" box="[666,754,1519,1545]" pageId="23" pageNumber="308" rank="species">sp. nov.</taxonomicNameLabel>
appears most similar to
<taxonomicName id="4C6A9BD57B29FFE4FB808F978565F9B8" authority="(Pic, 1901)" baseAuthorityName="Pic" baseAuthorityYear="1901" box="[1035,1373,1519,1545]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="308" phylum="Arthropoda" rank="species" species="monstrosicollis">
<emphasis id="B91E3C447B29FFE4FB808F9784EFF9B8" box="[1035,1239,1519,1545]" italics="true" pageId="23" pageNumber="308">N. monstrosicollis</emphasis>
(Pic, 1901)
</taxonomicName>
from the Greater Sunda islands of Borneo and Sumatra but is specifically different in the shape of the apical portion of aedeagus, the comparatively coarser punctured head dorsum and the anterior pronotal lobe comparatively wider than that in the insular species.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B29FFE4FF4C8C078414F928" box="[199,1068,1663,1689]" pageId="23" pageNumber="308" type="biology_ecology">
<paragraph id="8BD5E0567B29FFE4FF4C8C078414F928" blockId="23.[151,1437,150,1725]" box="[199,1068,1663,1689]" pageId="23" pageNumber="308">
<emphasis id="B91E3C447B29FFE4FF4C8C078114F928" bold="true" box="[199,300,1663,1689]" pageId="23" pageNumber="308">Ecology.</emphasis>
Sampled at light at the edge of primary lowland rainforest at
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<quantity id="4C924DB37B29FFE4FC558C078410F928" box="[990,1064,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="308" unit="m" value="900.0">900 m</quantity>
.
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</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B29FFE4FF4C8CDA8337F90C" box="[199,783,1698,1725]" pageId="23" pageNumber="308" type="distribution">
<paragraph id="8BD5E0567B29FFE4FF4C8CDA8337F90C" blockId="23.[151,1437,150,1725]" box="[199,783,1698,1725]" pageId="23" pageNumber="308">
<emphasis id="B91E3C447B29FFE4FF4C8CDA8159F90D" bold="true" box="[199,353,1698,1724]" pageId="23" pageNumber="308">Distribution.</emphasis>
So far known from northern
<collectingCountry id="F37DA0C67B29FFE4FD238CDB8333F90C" box="[680,779,1699,1725]" name="Vietnam" pageId="23" pageNumber="308">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

291
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<mods:title id="161F0F3B3F6BF4789BE62A6C12C248DB">New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum</mods:title>
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<taxonomicName id="4C6A9BD57B2CFFE1FF1C88128146FD34" authorityName="Telnov" authorityYear="2024" box="[151,382,618,645]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2CFFE1FF1C88128146FD34" bold="true" box="[151,382,618,645]" italics="true" pageId="18" pageNumber="303">Microhoria taguan</emphasis>
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<taxonomicNameLabel id="A22D813F7B2CFFE1FE0E881381DBFD34" box="[389,483,619,645]" pageId="18" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1351FCD37B2CFFE1FE798812829CFD35" box="[498,676,618,644]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="19.[151,250,1943,1967]" captionStartId-1="20.[151,250,1616,1640]" captionStartId-2="21.[151,250,924,948]" captionTargetBox-0="[170,1418,194,1907]" captionTargetBox-1="[159,1425,464,1582]" captionTargetBox-2="[185,1412,194,886]" captionTargetId-0="figure-18@19.[157,1430,181,1919]" captionTargetId-1="figure-149@20.[151,1436,453,1593]" captionTargetId-2="figure-18@21.[164,1423,181,900]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 14. Microhoria taguan sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." captionText-1="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." captionText-2="FIGURE 16. Microhoria taguan sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.13757621" figureDoi-1="http://doi.org/10.5281/zenodo.13757625" figureDoi-2="http://doi.org/10.5281/zenodo.13757629" httpUri-0="https://zenodo.org/record/13757621/files/figure.png" httpUri-1="https://zenodo.org/record/13757625/files/figure.png" httpUri-2="https://zenodo.org/record/13757629/files/figure.png" pageId="18" pageNumber="303">Figures 14–16</figureCitation>
)
</emphasis>
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This species belongs to the
<emphasis id="B91E3C447B2CFFE1FE4788AF820FFD40" box="[460,567,727,753]" italics="true" pageId="18" pageNumber="303">longiceps</emphasis>
species-group as of
<bibRefCitation id="EFFB9DA77B2CFFE1FC9688AF847FFD40" author="Kejval, Z. &amp; Chandler, D. S." box="[797,1095,727,753]" pageId="18" pageNumber="303" pagination="95 - 154" refId="ref13339" refString="Kejval, Z. &amp; Chandler, D. S. (2020) Generic revision of the Microhoriini with new species and synonymies from the Palaearctic Region (Coleoptera: Anthicidae). Acta entomologica musei nationalis pragae, 60 (1), 95 - 154. https: // doi. org / 10.37520 / aemnp. 2020.007" type="journal article" year="2020">Kejval &amp; Chandler (2020)</bibRefCitation>
.
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<emphasis id="B91E3C447B2CFFE1FF4C88838230FCA5" bold="true" box="[199,520,762,789]" pageId="18" pageNumber="303">Type material designated.</emphasis>
<materialsCitation id="3B02EA0B7B2CFFE1FD9E888384C5FC88" collectingDate="2008-08-06" collectionCode="BMNH" collectorName="M. V. L. Barclay &amp; H. Mendel &amp; R. Ewers" country="Taiwan" elevation="1920" latitude="24.042166" location="Nantou County" longLatPrecision="1" longitude="121.20925" pageId="18" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54D15EF47B2CFFE1FD9E88838245FCA4" box="[533,637,763,789]" pageId="18" pageNumber="303" type="holotype">Holotype</typeStatus>
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,
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:
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,
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;
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beating etc.
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<collectingDate id="EF903F7E7B2CFFE1FEFF896781D5FC88" box="[372,493,799,825]" pageId="18" pageNumber="303" value="2008-08-06">6.viii.2008</collectingDate>
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<collectorName id="269F85807B2CFFE1FE7F8967829BFC88" box="[500,675,799,825]" pageId="18" pageNumber="303">M.V.L. Barclay</collectorName>
,
<collectorName id="269F85807B2CFFE1FD2589678327FC88" box="[686,799,799,825]" pageId="18" pageNumber="303">H.Mendel</collectorName>
&amp;
<collectorName id="269F85807B2CFFE1FCC889678391FC88" box="[835,937,799,825]" pageId="18" pageNumber="303">R. Ewers</collectorName>
<collectionCode id="ED7B78937B2CFFE1FC3B8967843FFC88" box="[944,1031,799,825]" pageId="18" pageNumber="303">BMNH</collectionCode>
(E) 2008-85 [printed].
</materialsCitation>
</paragraph>
<paragraph id="8BD5E0567B2CFFE1FF4C893B84D9FC30" blockId="18.[151,1437,727,2013]" pageId="18" pageNumber="303">
<materialsCitation id="3B02EA0B7B2CFFE1FF4C893B84E6FC30" collectingDate="2008-08-12" collectionCode="BMNH" collectorName="H. Mendel &amp; Barclay" country="Taiwan" elevation="2574" latitude="23.266167" location="County Nanheng Sanshan" longLatPrecision="1" longitude="120.93955" pageId="18" pageNumber="303" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Kaohsiung" typeStatus="paratype">
<typeStatus id="54D15EF47B2CFFE1FF4C893B810CFCEC" box="[199,308,835,861]" pageId="18" pageNumber="303" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D6C2BDF7B2CFFE1FEC9893B815CFCEC" box="[322,356,835,861]" count="2" pageId="18" pageNumber="303" type="male">2♂</specimenCount>
&amp;
<specimenCount id="9D6C2BDF7B2CFFE1FE1D893B8181FCEC" box="[406,441,835,861]" count="4" pageId="18" pageNumber="303" type="female">4♀</specimenCount>
<collectionCode id="ED7B78937B2CFFE1FE4C893B8227FCEC" box="[455,543,835,861]" pageId="18" pageNumber="303">BMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B2CFFE1FDBB893B82A7FCEC" box="[560,671,835,861]" name="Taiwan" pageId="18" pageNumber="303">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B2CFFE1FD3A893B8312FCEC" box="[689,810,835,861]" country="Taiwan" name="Kaohsiung" pageId="18" pageNumber="303">Kaohsiung</collectingRegion>
<location id="8EB5B68D7B2CFFE1FCB3893B8453FCEC" LSID="urn:lsid:plazi:treatment:03C351407B2CFFE6FF1C8812848FFA1C:8EB5B68D7B2CFFE1FCB3893B8453FCEC" box="[824,1131,835,861]" country="Taiwan" latitude="23.266167" longLatPrecision="1" longitude="120.93955" name="County Nanheng Sanshan" pageId="18" pageNumber="303" stateProvince="Kaohsiung">County Nanheng Sanshan</location>
,
<quantity id="4C924DB37B2CFFE1FBF6893A84F3FCEC" box="[1149,1227,834,861]" metricMagnitude="3" metricUnit="m" metricValue="2.574" pageId="18" pageNumber="303" unit="m" value="2574.0">
<elevation id="004707657B2CFFE1FBF6893A84F3FCEC" box="[1149,1227,834,861]" metricMagnitude="3" metricUnit="m" metricValue="2.574" pageId="18" pageNumber="303" unit="m" value="2574.0">2574m</elevation>
</quantity>
trail above road,
<date id="FFD4C6967B2CFFE1FF1C891F8127FC30" box="[151,287,871,897]" pageId="18" pageNumber="303" value="2008-08-12">
<collectingDate id="EF903F7E7B2CFFE1FF1C891F8127FC30" box="[151,287,871,897]" pageId="18" pageNumber="303" value="2008-08-12">12.viii.2008</collectingDate>
</date>
<geoCoordinate id="EE5E86917B2CFFE1FEAD891E818DFC30" box="[294,437,870,897]" degrees="23" direction="north" minutes="15.970" orientation="latitude" pageId="18" pageNumber="303" precision="1" value="23.266167">N23°15.970’</geoCoordinate>
<geoCoordinate id="EE5E86917B2CFFE1FE37891E826EFC30" box="[444,598,870,897]" degrees="120" direction="east" minutes="56.373" orientation="longitude" pageId="18" pageNumber="303" precision="1" value="120.93955">E120°56.373’</geoCoordinate>
<collectorName id="269F85807B2CFFE1FDD5891F82F6FC30" box="[606,718,871,897]" pageId="18" pageNumber="303">H.Mendel</collectorName>
&amp; MVL
<collectorName id="269F85807B2CFFE1FCBD891F83B5FC30" box="[822,909,871,897]" pageId="18" pageNumber="303">Barclay</collectorName>
<collectionCode id="ED7B78937B2CFFE1FC1E891F83D4FC30" box="[917,1004,871,897]" pageId="18" pageNumber="303">BMNH</collectionCode>
(E) 2008-85 [printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2CFFE1FF4C89F28262FC78" pageId="18" pageNumber="303" type="etymology">
<paragraph id="8BD5E0567B2CFFE1FF4C89F28262FC78" blockId="18.[151,1437,727,2013]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C89F2819EFC15" bold="true" box="[199,422,906,932]" pageId="18" pageNumber="303">Derivatio nominis.</emphasis>
Toponymic. Named after Taguan Mountain in central
<collectingCountry id="F37DA0C67B2CFFE1FB9C89F38451FC14" box="[1047,1129,907,933]" name="Taiwan" pageId="18" pageNumber="303">Taiwan</collectingCountry>
in the area where the
<typeStatus id="54D15EF47B2CFFE1FAE789F385A4FC14" box="[1388,1436,907,933]" pageId="18" pageNumber="303">type</typeStatus>
series was sampled. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2CFFE6FF4C89AB82B7FB64" lastPageId="21" lastPageNumber="306" pageId="18" pageNumber="303" type="description">
<paragraph id="8BD5E0567B2CFFE1FF4C89AB8476FBE8" blockId="18.[151,1437,727,2013]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C89AB814EFC5C" bold="true" box="[199,374,979,1005]" pageId="18" pageNumber="303">Measurements</emphasis>
,
<typeStatus id="54D15EF47B2CFFE1FE0C89AB81D1FC5C" box="[391,489,979,1005]" pageId="18" pageNumber="303" type="holotype">holotype</typeStatus>
male, total body length excluding partially exposed terminalia
<quantity id="4C924DB37B2CFFE1FB4F89AB8532FC5C" box="[1220,1290,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="303" unit="mm" value="3.0">3 mm</quantity>
; head length
<quantity id="4C924DB37B2CFFE1FF1C898F80C8FBA0" box="[151,240,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="303" unit="mm" value="0.6">0.6 mm</quantity>
, head width across compound eyes
<quantity id="4C924DB37B2CFFE1FD08898E82D2FBA0" box="[643,746,1014,1041]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="303" unit="mm" value="0.55">0.55 mm</quantity>
, pronotal length
<quantity id="4C924DB37B2CFFE1FC23898F8438FBA0" box="[936,1024,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="303" unit="mm" value="0.6">0.6 mm</quantity>
, maximum pronotal width
<quantity id="4C924DB37B2CFFE1FABA898E85A0FBA0" box="[1329,1432,1014,1041]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="303" unit="mm" value="0.45">0.45 mm</quantity>
, minimum pronotal width in constriction area
<quantity id="4C924DB37B2CFFE1FD148E62833FFB84" box="[671,775,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="303" unit="mm" value="0.35">0.35 mm</quantity>
, elytral length
<quantity id="4C924DB37B2CFFE1FC388E638435FB84" box="[947,1037,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="18" pageNumber="303" unit="mm" value="1.8">1.8 mm</quantity>
, combined maximum elytral width across midlength
<quantity id="4C924DB37B2CFFE1FED68E478199FBE8" box="[349,417,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="303" unit="mm" value="1.0">1 mm</quantity>
. Male
<typeStatus id="54D15EF47B2CFFE1FE678E47826FFBE8" box="[492,599,1087,1113]" pageId="18" pageNumber="303" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B2CFFE1FDD58E47828EFBE8" box="[606,694,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="303" unit="mm" value="2.9">2.9 mm</quantity>
, female
<typeStatus id="54D15EF47B2CFFE1FC9E8E4783B8FBE8" box="[789,896,1087,1113]" pageId="18" pageNumber="303" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B2CFFE1FC0C8E478436FBE8" box="[903,1038,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="18" pageNumber="303" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
long.
</paragraph>
<paragraph id="8BD5E0567B2CFFE7FF4C8E1A83F6FE1C" blockId="18.[151,1437,727,2013]" lastBlockId="20.[151,1436,151,429]" lastPageId="20" lastPageNumber="305" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C8E1A8161FBCD" bold="true" box="[199,345,1122,1148]" pageId="18" pageNumber="303">Description.</emphasis>
<typeStatus id="54D15EF47B2CFFE1FED48E1B81FFFBCC" box="[351,455,1123,1149]" pageId="18" pageNumber="303" type="holotype">Holotype</typeStatus>
male. Dorsal and ventral head brown, paler yellowish around antennal insertions and on clypeus. Pronotum yellowish brown. Elytra brown, each elytron with poorly defined yellowish humeral (smaller) and postmedian (larger) spot; spots distinctly separated, not confluent. Maxillary palpus, antenna and legs pale yellow-rufous. Ventral pterothorax and abdominal ventrites brown. Head elongate elliptical, hardly 1.1× as long as wide, slightly convex in dorsal aspect, glossy. Labrum subtruncate at anterior margin. Frontoclypeal suture present, slightly arched (shallowly concave anteriorly). Compound eye moderate, about 0.7× as long as converging tempus, strongly protruding from lateral outline of head. Head base rounded, posterior temporal angles obsolete. Head dorsal punctures moderately large, shallow, sparse. Intervening spaces smooth and glossy, as wide as to twice as wide as punctures. Dorsal cranial setae yellowish, sparse, and inconspicuous. Antenna extending slightly beyond elytral base when directed posteriad, hardly thickened in apical half. Antennomeres 7–10 somewhat widened and shortened, more regularly trapezoid in dorsal view, slightly widened distally. Penultimate antennomere about 1.1–1.2× as long as wide. Terminal antennomere elongate, bluntly pointed, about 2.2–2.3× as long as penultimate antennomere, about the combined length of antennomeres 9–10. Penultimate maxillary palpomere lobate at mesal margin. Terminal maxillary palpomere securiform. Cranial ‘neck’ less than one third head width across compound eyes, densely finely punctured, moderately glossy. Pronotum longer than wide, distinctly narrower than head across eyes, subtruncate at anterior margin. Pronotal disc moderately glossy, anterior lobe slightly convex in dorsal aspect. Lateral margins broadly rounded in anterior half, moderately constricted postmedium. Anterior pronotal rim distinct dorsally, laterally and ventrally. Antebasal sulcus wide, well-developed dorsally and laterally. Pronotal punctures on anterior portion of pronotal disc larger than those on head, minute and sparse on lateral sides of anterior lobe, large and dense on basal half of disc, in particular, in constriction area. Large punctures are circular or elliptical, puncture background microreticulate or corrugate. Intervening spaces glossy, about as wide as punctures on anterior half of pronotal disc, narrower than those on rest of pronotal dorsum. Pronotal setae as those on head, somewhat longer, not fully appressed. Scutellar shield nearly triangular, apically rounded, glossy. Elytra elongate, about 1.8× as long as wide, widest at midlength, dorsally flattened. Postbasal transverse impression very feeble, nearly obsolete. Humerus broadly rounded. Apical sutural angle rounded.Apex of elytron modified, with a short, obtuse denticle at opening of a gland channel. Elytral surface glossy and smooth, punctures smaller than those on pronotal constriction, but distinct and dense. Intervening spaces smooth and glossy, generally as wide as punctures. Elytral setae yellowish, long and dense, suberect, directed posteriad. Metathoracic wing fully developed (functional). Legs without modifications, moderately densely setose. Tibial terminal spurs paired, inconspicuous. Basal metatarsomere longer than combined length of remaining metatarsomeres. Tergite VII broadly rounded at posterior margin (
<figureCitation id="1351FCD37B2AFFE7FBFD8B5F84D8FEF0" box="[1142,1248,294,321]" captionStart="FIGURE 15" captionStartId="20.[151,250,1616,1640]" captionTargetBox="[159,1425,464,1582]" captionTargetId="figure-149@20.[151,1436,453,1593]" captionTargetPageId="20" captionText="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305">Fig. 15A</figureCitation>
). Morphological sternite VII broadly rounded at posterior margin (
<figureCitation id="1351FCD37B2AFFE7FD4A8B33831FFED4" box="[705,807,330,357]" captionStart="FIGURE 15" captionStartId="20.[151,250,1616,1640]" captionTargetBox="[159,1425,464,1582]" captionTargetId="figure-149@20.[151,1436,453,1593]" captionTargetPageId="20" captionText="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305">Fig. 15B</figureCitation>
). Morphological sternite VIII somewhat bow tie-shaped (
<figureCitation id="1351FCD37B2AFFE7FF148B17813EFE38" box="[159,262,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1616,1640]" captionTargetBox="[159,1425,464,1582]" captionTargetId="figure-149@20.[151,1436,453,1593]" captionTargetPageId="20" captionText="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305">Fig. 15C</figureCitation>
), sternite IX Y-shaped, arms moderately long (
<figureCitation id="1351FCD37B2AFFE7FC988B178344FE38" box="[787,892,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1616,1640]" captionTargetBox="[159,1425,464,1582]" captionTargetId="figure-149@20.[151,1436,453,1593]" captionTargetPageId="20" captionText="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305">Fig. 15D</figureCitation>
). Aedeagus (
<figureCitation id="1351FCD37B2AFFE7FB878B1784AAFE38" box="[1036,1170,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1616,1640]" captionTargetBox="[159,1425,464,1582]" captionTargetId="figure-149@20.[151,1436,453,1593]" captionTargetPageId="20" captionText="FIGURE 15. Microhoria taguan sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305">Fig. 15E–G</figureCitation>
) elongate; tegmen each side with acute, sabre-like spine; endophallic armature of irregular shape.
</paragraph>
<caption id="DF15B0DE7B2DFFE0FF1C8DEF835CF862" ID-DOI="http://doi.org/10.5281/zenodo.13757621" ID-Zenodo-Dep="13757621" httpUri="https://zenodo.org/record/13757621/files/figure.png" pageId="19" pageNumber="304" startId="19.[151,250,1943,1967]" targetBox="[170,1418,194,1907]" targetPageId="19" targetType="figure">
<paragraph id="8BD5E0567B2DFFE0FF1C8DEF835CF862" blockId="19.[151,1436,1943,2003]" pageId="19" pageNumber="304">
<emphasis id="B91E3C447B2DFFE0FF1C8DEF8118F81E" bold="true" box="[151,288,1943,1967]" pageId="19" pageNumber="304">FIGURE 14.</emphasis>
<taxonomicName id="4C6A9BD57B2DFFE0FEAD8DEF81DFF81E" authorityName="Telnov" authorityYear="2024" box="[294,487,1943,1967]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="304" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2DFFE0FEAD8DEF81DFF81E" box="[294,487,1943,1967]" italics="true" pageId="19" pageNumber="304">Microhoria taguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2DFFE0FE658DE08205F81E" box="[494,573,1944,1967]" pageId="19" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale].
</paragraph>
</caption>
<caption id="DF15B0DE7B2AFFE7FF1C8C2882A5F901" ID-DOI="http://doi.org/10.5281/zenodo.13757625" ID-Zenodo-Dep="13757625" httpUri="https://zenodo.org/record/13757625/files/figure.png" pageId="20" pageNumber="305" startId="20.[151,250,1616,1640]" targetBox="[159,1425,464,1582]" targetPageId="20" targetType="figure">
<paragraph id="8BD5E0567B2AFFE7FF1C8C2882A5F901" blockId="20.[151,1437,1616,1712]" pageId="20" pageNumber="305">
<emphasis id="B91E3C447B2AFFE7FF1C8C28811AF9D9" bold="true" box="[151,290,1616,1640]" pageId="20" pageNumber="305">FIGURE 15.</emphasis>
<taxonomicName id="4C6A9BD57B2AFFE7FEA08C2881D6F9D9" authorityName="Telnov" authorityYear="2024" box="[299,494,1616,1640]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="305" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2AFFE7FEA08C2881D6F9D9" box="[299,494,1616,1640]" italics="true" pageId="20" pageNumber="305">Microhoria taguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2AFFE7FE7C8C288270F9D6" box="[503,584,1616,1639]" pageId="20" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified; G—ditto, median portion, magnified [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B2AFFE7FF4C8D7A8298F845" blockId="20.[151,1437,1794,2037]" pageId="20" pageNumber="305">
<emphasis id="B91E3C447B2AFFE7FF4C8D7A8183F8AD" bold="true" box="[199,443,1794,1820]" pageId="20" pageNumber="305">Sexual dimorphism.</emphasis>
Female (
<figureCitation id="1351FCD37B2AFFE7FDA18D7A8281F8AD" box="[554,697,1794,1820]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[170,1418,194,1907]" captionTargetId="figure-18@19.[157,1430,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Microhoria taguan sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757621" httpUri="https://zenodo.org/record/13757621/files/figure.png" pageId="20" pageNumber="305">Figs 14C–D</figureCitation>
,
<figureCitation id="1351FCD37B2AFFE7FD428D7A82DEF8AD" box="[713,742,1794,1820]" captionStart="FIGURE 16" captionStartId="21.[151,250,924,948]" captionTargetBox="[185,1412,194,886]" captionTargetId="figure-18@21.[164,1423,181,900]" captionTargetPageId="21" captionText="FIGURE 16. Microhoria taguan sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757629" httpUri="https://zenodo.org/record/13757629/files/figure.png" pageId="20" pageNumber="305">16</figureCitation>
) slightly stouter, elytra stronger widened in median portion and comparatively stronger convex in dorsal aspect. Elytral sutural angle produced into a moderately long, acutely angulate process which is slightly curved ventrally at apex (
<figureCitation id="1351FCD37B2AFFE7FCD88D328385F8D5" box="[851,957,1866,1892]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[170,1418,194,1907]" captionTargetId="figure-18@19.[157,1430,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Microhoria taguan sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757621" httpUri="https://zenodo.org/record/13757621/files/figure.png" pageId="20" pageNumber="305">Fig. 14C</figureCitation>
). Tergite VII strongly modified, posterior margin with subcircular, short, irregularly serrate median prong; posterior margin strongly deflected dorsoanteriad forming irregularly shaped, medially broadly emarginate and laterally irregularly serrate ‘pocket’ (
<figureCitation id="1351FCD37B2AFFE7FAE48DEA80D5F861" captionStart="FIGURE 16" captionStartId="21.[151,250,924,948]" captionTargetBox="[185,1412,194,886]" captionTargetId="figure-18@21.[164,1423,181,900]" captionTargetPageId="21" captionText="FIGURE 16. Microhoria taguan sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757629" httpUri="https://zenodo.org/record/13757629/files/figure.png" pageId="20" pageNumber="305">Fig. 16A–B</figureCitation>
). Morphological sternite VII broad, deeply U-shapely emarginate medially at posterior margin (
<figureCitation id="1351FCD37B2AFFE7FAA38DCE85A8F861" box="[1320,1424,1974,2000]" captionStart="FIGURE 16" captionStartId="21.[151,250,924,948]" captionTargetBox="[185,1412,194,886]" captionTargetId="figure-18@21.[164,1423,181,900]" captionTargetPageId="21" captionText="FIGURE 16. Microhoria taguan sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757629" httpUri="https://zenodo.org/record/13757629/files/figure.png" pageId="20" pageNumber="305">Fig. 16C</figureCitation>
). Morphological sternite IX rod-like (
<figureCitation id="1351FCD37B2AFFE7FDA08DA282ABF845" box="[555,659,2010,2036]" captionStart="FIGURE 16" captionStartId="21.[151,250,924,948]" captionTargetBox="[185,1412,194,886]" captionTargetId="figure-18@21.[164,1423,181,900]" captionTargetPageId="21" captionText="FIGURE 16. Microhoria taguan sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757629" httpUri="https://zenodo.org/record/13757629/files/figure.png" pageId="20" pageNumber="305">Fig. 16D</figureCitation>
).
</paragraph>
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<emphasis id="B91E3C447B2BFFE6FF1C89E4811BFC05" bold="true" box="[151,291,924,948]" pageId="21" pageNumber="306">FIGURE 16.</emphasis>
<taxonomicName id="4C6A9BD57B2BFFE6FEA789E481C8FC05" authorityName="Telnov" authorityYear="2024" box="[300,496,924,948]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="306" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2BFFE6FEA789E481C8FC05" box="[300,496,924,948]" italics="true" pageId="21" pageNumber="306">Microhoria taguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2BFFE6FE7189E58276FC05" box="[506,590,925,948]" pageId="21" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Morphological sternite IX [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B2BFFE6FF4C8E5282B7FB64" blockId="21.[151,1437,1066,1453]" pageId="21" pageNumber="306">
<emphasis id="B91E3C447B2BFFE6FF4C8E5281D0FBF5" bold="true" box="[199,488,1066,1092]" pageId="21" pageNumber="306">Intraspecific variability.</emphasis>
Head or whole dorsal forebody darker brown to black-brown in some
<typeStatus id="54D15EF47B2BFFE6FA808E538542FBF4" box="[1291,1402,1067,1093]" pageId="21" pageNumber="306" type="paratype">paratypes</typeStatus>
. In
<specimenCount id="9D6C2BDF7B2BFFE6FF1C8E37811EFBD8" box="[151,294,1103,1129]" count="1" pageId="21" pageNumber="306" type="generic" typeStatus="paratype">one paratype</specimenCount>
anterior pale elytral spot not interrupted on suture, occupying whole basal fifth of elytra. Posterior pale paired spot extends to cover most of apical half of elytra except dark elytral apex but also in that case posterior spots are separated by dark sutural area. Dorsal pronotal punctures on constriction area vary strongly in shape and size. Total body length varies from
<quantity id="4C924DB37B2BFFE6FE608EC382B2FB64" box="[491,650,1211,1237]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="21" pageNumber="306" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8 to 3.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2BFFE6FF4C8EA68558FAF0" pageId="21" pageNumber="306" type="diagnosis">
<paragraph id="8BD5E0567B2BFFE6FF4C8EA68558FAF0" blockId="21.[151,1437,1066,1453]" pageId="21" pageNumber="306">
<emphasis id="B91E3C447B2BFFE6FF4C8EA681EBFB49" bold="true" box="[199,467,1246,1272]" pageId="21" pageNumber="306">Differential diagnosis.</emphasis>
<taxonomicName id="4C6A9BD57B2BFFE6FE548EA68281FB49" authorityName="Telnov" authorityYear="2024" box="[479,697,1246,1272]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="306" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2BFFE6FE548EA68281FB49" box="[479,697,1246,1272]" italics="true" pageId="21" pageNumber="306">Microhoria taguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2BFFE6FD4E8EA78318FB48" box="[709,800,1247,1273]" pageId="21" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
is externally similar to Taiwanese
<taxonomicName id="4C6A9BD57B2BFFE6FB488EA7850EFB49" authorityName="Telnov" authorityYear="2024" box="[1219,1334,1246,1272]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="306" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B2BFFE6FB488EA7850EFB49" box="[1219,1334,1246,1272]" italics="true" pageId="21" pageNumber="306">M. lingua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2BFFE6FACA8EA785A5FB48" box="[1345,1437,1247,1273]" pageId="21" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
(described herein) and several other similarly-coloured Himalayan and eastern-Palaearctic congeners but is specifically different in the shape of the male aedeagus and the female tergite and morphological sternite VII.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2BFFE6FF4C8F338474FA38" pageId="21" pageNumber="306" type="biology_ecology">
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<emphasis id="B91E3C447B2BFFE6FF4C8F338114FAD4" bold="true" box="[199,300,1355,1381]" pageId="21" pageNumber="306">Ecology.</emphasis>
Occurs at about
<quantity id="4C924DB37B2BFFE6FE628F3282BBFAD4" box="[489,643,1354,1381]" metricMagnitude="3" metricUnit="m" metricValue="2.247" metricValueMax="2.574" metricValueMin="1.92" pageId="21" pageNumber="306" unit="m" value="2247.0" valueMax="2574.0" valueMin="1920.0">1920–2574 m</quantity>
in forested area. The
<typeStatus id="54D15EF47B2BFFE6FCFD8F3283E0FAD5" box="[886,984,1354,1380]" pageId="21" pageNumber="306" type="holotype">holotype</typeStatus>
collected by beating forest vegetation, a typical microstation of many
<taxonomicName id="4C6A9BD57B2BFFE6FE6A8F16834CFA39" box="[481,884,1390,1417]" pageId="21" pageNumber="306">
<emphasis id="B91E3C447B2BFFE6FE6A8F168259FA39" box="[481,609,1390,1416]" italics="true" pageId="21" pageNumber="306">Microhoria</emphasis>
Chevrolat, 1877 species
</taxonomicName>
(see
<bibRefCitation id="EFFB9DA77B2BFFE6FC248F168478FA38" author="Telnov, D." box="[943,1088,1390,1417]" pageId="21" pageNumber="306" pagination="195 - 312" refId="ref13736" refString="Telnov, D. (2022) Revisional notes on the genus Microhoria Chevrolat, 1877 (Insecta: Coleoptera: Anthicidae) from the Eastern Mediterranean and Turkey, with new descriptions, an annotated catalogue, and a key. Baltic Journal of Coleopterology, 22 (2), 195 - 312." type="journal article" year="2022">Telnov 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2BFFE6FF4C8FEA848FFA1C" box="[199,1207,1426,1453]" pageId="21" pageNumber="306" type="distribution">
<paragraph id="8BD5E0567B2BFFE6FF4C8FEA848FFA1C" blockId="21.[151,1437,1066,1453]" box="[199,1207,1426,1453]" pageId="21" pageNumber="306">
<emphasis id="B91E3C447B2BFFE6FF4C8FEA8159FA1D" bold="true" box="[199,353,1426,1452]" pageId="21" pageNumber="306">Distribution.</emphasis>
So far known from mountainous central part of
<collectingCountry id="F37DA0C67B2BFFE6FCF58FEA83EBFA1D" box="[894,979,1426,1452]" name="Taiwan" pageId="21" pageNumber="306">Taiwan</collectingCountry>
,
<collectingRegion id="49AE2EB47B2BFFE6FC558FEA846FFA1D" box="[990,1111,1426,1452]" country="Taiwan" name="Kaohsiung" pageId="21" pageNumber="306">Kaohsiung</collectingRegion>
County.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="240B27843DBB94B6531C4A03ED49678F">New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum</mods:title>
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<taxonomicName id="4C6A9BD57B30FFFDFF1C8F83814FF9A7" authorityName="Telnov" authorityYear="2024" box="[151,375,1531,1558]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="299" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B30FFFDFF1C8F83814FF9A7" bold="true" box="[151,375,1531,1558]" italics="true" pageId="14" pageNumber="299">Microhoria lingua</emphasis>
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(
<figureCitation id="1351FCD37B30FFFDFE678F8382A4F9A4" box="[492,668,1531,1557]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="15.[151,250,1943,1967]" captionStartId-1="16.[151,250,1296,1320]" captionStartId-2="17.[151,250,1911,1935]" captionTargetBox-0="[331,1249,194,1907]" captionTargetBox-1="[163,1424,194,1259]" captionTargetBox-2="[197,1387,1041,1874]" captionTargetId-0="figure-18@15.[325,1261,181,1919]" captionTargetId-1="figure-28@16.[151,1436,181,1271]" captionTargetId-2="figure-399@17.[188,1399,1029,1887]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURE 11. Microhoria lingua sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." captionText-1="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." captionText-2="FIGURE 13. Microhoria lingua sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.13757611" figureDoi-1="http://doi.org/10.5281/zenodo.13757615" figureDoi-2="http://doi.org/10.5281/zenodo.13757617" httpUri-0="https://zenodo.org/record/13757611/files/figure.png" httpUri-1="https://zenodo.org/record/13757615/files/figure.png" httpUri-2="https://zenodo.org/record/13757617/files/figure.png" pageId="14" pageNumber="299">Figures 11–13</figureCitation>
)
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This species belongs to the
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species-group as of
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.
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<emphasis id="B91E3C447B30FFFDFF4C8CF4823DF914" bold="true" box="[199,517,1675,1702]" pageId="14" pageNumber="299">Type material designated.</emphasis>
<materialsCitation id="3B02EA0B7B30FFFDFD9A8CF38235F95C" collectingDate="2002-04-18" collectingDateMax="2002-04-19" collectingDateMin="2002-04-18" collectionCode="HMNH" collectorName="D. A. Anstine &amp; Gy. Fabian &amp; O. Merkl" country="Taiwan" elevation="1945" latitude="24.076017" location="Kao-Leng Dyi" longLatPrecision="1" longitude="121.1341" pageId="14" pageNumber="299" specimenCount="1" specimenCount-male="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54D15EF47B30FFFDFD9A8CF38241F914" box="[529,633,1675,1701]" pageId="14" pageNumber="299" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6C2BDF7B30FFFDFD0F8CF482A3F917" box="[644,667,1676,1702]" count="1" pageId="14" pageNumber="299" type="male">♂</specimenCount>
,
<collectionCode id="ED7B78937B30FFFDFD278CF4833EF917" box="[684,774,1676,1702]" country="Japan" httpUri="http://grbio.org/cool/jm3w-rdxm" name="Hayashibara Museum of Natural History" pageId="14" pageNumber="299">HMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B30FFFDFC9E8CF483BCF917" box="[789,900,1676,1702]" name="Taiwan" pageId="14" pageNumber="299">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B30FFFDFC188CF48407F917" box="[915,1087,1676,1702]" country="Taiwan" name="Nantou" pageId="14" pageNumber="299">Nantou county</collectingRegion>
,
<location id="8EB5B68D7B30FFFDFBC48CF484C1F914" LSID="urn:lsid:plazi:treatment:03C351407B30FFE1FF1C8F8384A8FDA8:8EB5B68D7B30FFFDFBC48CF484C1F914" box="[1103,1273,1675,1702]" country="Taiwan" latitude="24.076017" longLatPrecision="1" longitude="121.1341" name="Kao-Leng Dyi" pageId="14" pageNumber="299" stateProvince="Nantou">Kao-Leng Dyi</location>
,
<locationDeviation id="9A67F9747B30FFFDFA828CF480DFF978" location="Wushe" pageId="14" pageNumber="299">
<quantity id="4C924DB37B30FFFDFA828CF4856CF914" box="[1289,1364,1675,1702]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="299" unit="km" value="18.0">18 km</quantity>
W of Wushe
</locationDeviation>
,
<geoCoordinate id="EE5E86917B30FFFDFF798CD78146F97B" box="[242,382,1711,1738]" degrees="24" direction="north" minutes="4.561" orientation="latitude" pageId="14" pageNumber="299" precision="1" value="24.076017">24°4.561’ N</geoCoordinate>
,
<geoCoordinate id="EE5E86917B30FFFDFE028CD78227F97B" box="[393,543,1711,1738]" degrees="121" direction="east" minutes="8.046" orientation="longitude" pageId="14" pageNumber="299" precision="1" value="121.1341">121°8.046’ E</geoCoordinate>
, [printed] //
<quantity id="4C924DB37B30FFFDFD208CD7833BF97B" box="[683,771,1711,1738]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="14" pageNumber="299" unit="m" value="1945.0">
<elevation id="004707657B30FFFDFD208CD7833BF97B" box="[683,771,1711,1738]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="14" pageNumber="299" unit="m" value="1945.0">1945 m</elevation>
</quantity>
, swept from vegetation,
<date id="FFD4C6967B30FFFDFB938CC884F8F97B" box="[1048,1216,1712,1738]" pageId="14" pageNumber="299" value="2002-04-18" valueMax="2002-04-19" valueMin="2002-04-18">
<collectingDate id="EF903F7E7B30FFFDFB938CC884F8F97B" box="[1048,1216,1712,1738]" pageId="14" pageNumber="299" value="2002-04-18" valueMax="2002-04-19" valueMin="2002-04-18">18‒19.IV.2002</collectingDate>
</date>
, leg.
<collectorName id="269F85807B30FFFDFB778CC885A0F978" box="[1276,1432,1711,1738]" pageId="14" pageNumber="299">D. A. Anstine</collectorName>
,
<collectorName id="269F85807B30FFFDFF1C8CAC812AF95C" box="[151,274,1747,1774]" pageId="14" pageNumber="299">Gy. Fábián</collectorName>
&amp;
<collectorName id="269F85807B30FFFDFEBD8CAC81A4F95C" box="[310,412,1747,1774]" pageId="14" pageNumber="299">O. Merkl</collectorName>
[printed].
</materialsCitation>
</paragraph>
<paragraph id="8BD5E0567B30FFFDFF4C8C808531F8A0" blockId="14.[151,1437,1639,2026]" box="[199,1289,1783,1810]" pageId="14" pageNumber="299">
<typeStatus id="54D15EF47B30FFFDFF4C8C80810CF8A3" box="[199,308,1784,1810]" pageId="14" pageNumber="299" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D6C2BDF7B30FFFDFEB08C8081EEF8A0" box="[315,470,1783,1810]" count="13" pageId="14" pageNumber="299" type="generic">13 specimens</specimenCount>
.
<materialsCitation id="3B02EA0B7B30FFFDFE6A8C8F8291F8A3" box="[481,681,1783,1810]" collectionCode="HMNH" pageId="14" pageNumber="299" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D6C2BDF7B30FFFDFE6A8C8F823CF8A0" box="[481,516,1783,1809]" count="6" pageId="14" pageNumber="299" type="male">6♂</specimenCount>
&amp;
<specimenCount id="9D6C2BDF7B30FFFDFDAC8C808273F8A3" box="[551,587,1784,1810]" count="4" pageId="14" pageNumber="299" type="female">4♀</specimenCount>
<collectionCode id="ED7B78937B30FFFDFDDA8C808291F8A3" box="[593,681,1784,1810]" country="Japan" httpUri="http://grbio.org/cool/jm3w-rdxm" name="Hayashibara Museum of Natural History" pageId="14" pageNumber="299">HMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B02EA0B7B30FFFDFD3E8C80830EF8A3" box="[693,822,1784,1810]" collectionCode="BMNH" pageId="14" pageNumber="299" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D6C2BDF7B30FFFDFD3E8C8082E0F8A3" box="[693,728,1784,1810]" count="1" pageId="14" pageNumber="299" type="male">1♂</specimenCount>
<collectionCode id="ED7B78937B30FFFDFD548C80830EF8A3" box="[735,822,1784,1810]" pageId="14" pageNumber="299">BMNH</collectionCode>
</materialsCitation>
,
<specimenCount id="9D6C2BDF7B30FFFDFCCA8C80835CF8A3" box="[833,868,1784,1810]" count="1" pageId="14" pageNumber="299" type="male">1♂</specimenCount>
&amp;
<specimenCount id="9D6C2BDF7B30FFFDFC038C808393F8A3" box="[904,939,1784,1810]" count="1" pageId="14" pageNumber="299" type="female">1♀</specimenCount>
DTC: same labels as
<typeStatus id="54D15EF47B30FFFDFB2B8C8F853EF8A0" box="[1184,1286,1783,1809]" pageId="14" pageNumber="299" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B30FFFDFF4C8D63814EF8E8" pageId="14" pageNumber="299" type="etymology">
<paragraph id="8BD5E0567B30FFFDFF4C8D63814EF8E8" blockId="14.[151,1437,1639,2026]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8D63819BF884" bold="true" box="[199,419,1819,1845]" pageId="14" pageNumber="299">Derivatio nominis.</emphasis>
Named from Latin ‘lingua’ (tongue) to point on the tongue-shaped aedeagus of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B30FFE1FF4C8D1C80F2FEF0" lastPageId="18" lastPageNumber="303" pageId="14" pageNumber="299" type="description">
<paragraph id="8BD5E0567B30FFFDFF4C8D1C84CFF858" blockId="14.[151,1437,1639,2026]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8D1C814EF8CF" bold="true" box="[199,374,1892,1918]" pageId="14" pageNumber="299">Measurements</emphasis>
,
<typeStatus id="54D15EF47B30FFFDFE0E8D1B81DFF8CC" box="[389,487,1891,1917]" pageId="14" pageNumber="299" type="holotype">holotype</typeStatus>
male, total body length excluding partially exposed terminalia
<quantity id="4C924DB37B30FFFDFB3F8D1C8535F8CF" box="[1204,1293,1892,1918]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="299" unit="mm" value="2.4">2.4 mm</quantity>
; head length
<quantity id="4C924DB37B30FFFDFF1C8DFF80C8F813" box="[151,240,1927,1954]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="299" unit="mm" value="0.5">0.5 mm</quantity>
, head width across compound eyes
<quantity id="4C924DB37B30FFFDFD028DFF82C8F813" box="[649,752,1927,1954]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="299" unit="mm" value="0.45">0.45 mm</quantity>
, pronotal length
<quantity id="4C924DB37B30FFFDFC3A8DFF8432F813" box="[945,1034,1927,1954]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="299" unit="mm" value="0.5">0.5 mm</quantity>
, maximum pronotal width
<quantity id="4C924DB37B30FFFDFAB58DF085A0F813" box="[1342,1432,1928,1954]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="299" unit="mm" value="0.4">0.4 mm</quantity>
, minimum pronotal width in constriction area
<quantity id="4C924DB37B30FFFDFD2F8DD482C7F877" box="[676,767,1964,1990]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="299" unit="mm" value="0.3">0.3 mm</quantity>
, elytral length
<quantity id="4C924DB37B30FFFDFC248DD48431F877" box="[943,1033,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="299" unit="mm" value="1.4">1.4 mm</quantity>
, combined maximum elytral width across midlength
<quantity id="4C924DB37B30FFFDFED68DA8818EF85B" box="[349,438,2000,2026]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="299" unit="mm" value="0.8">0.8 mm</quantity>
. Selected male
<typeStatus id="54D15EF47B30FFFDFDEF8DA882F7F85B" box="[612,719,2000,2026]" pageId="14" pageNumber="299" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B30FFFDFD5D8DB78367F85B" box="[726,863,1999,2026]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="14" pageNumber="299" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
, female
<typeStatus id="54D15EF47B30FFFDFC358DA88411F85B" box="[958,1065,2000,2026]" pageId="14" pageNumber="299" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B30FFFDFBBB8DA8848FF85B" box="[1072,1207,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="14" pageNumber="299" unit="mm" value="3.15" valueMax="3.4" valueMin="2.9">2.9–3.4 mm</quantity>
long.
</paragraph>
<caption id="DF15B0DE7B31FFFCFF1C8DEF835CF862" ID-DOI="http://doi.org/10.5281/zenodo.13757611" ID-Zenodo-Dep="13757611" httpUri="https://zenodo.org/record/13757611/files/figure.png" pageId="15" pageNumber="300" startId="15.[151,250,1943,1967]" targetBox="[331,1249,194,1907]" targetPageId="15" targetType="figure">
<paragraph id="8BD5E0567B31FFFCFF1C8DEF835CF862" blockId="15.[151,1436,1943,2003]" pageId="15" pageNumber="300">
<emphasis id="B91E3C447B31FFFCFF1C8DEF8127F81E" bold="true" box="[151,287,1943,1967]" pageId="15" pageNumber="300">FIGURE 11.</emphasis>
<taxonomicName id="4C6A9BD57B31FFFCFEAD8DEF81DAF81E" authorityName="Telnov" authorityYear="2024" box="[294,482,1943,1967]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="300" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B31FFFCFEAD8DEF81DAF81E" box="[294,482,1943,1967]" italics="true" pageId="15" pageNumber="300">Microhoria lingua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B31FFFCFE638DE08200F81E" box="[488,568,1944,1967]" pageId="15" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale].
</paragraph>
</caption>
<caption id="DF15B0DE7B2EFFE3FF1C8F688543FAFD" ID-DOI="http://doi.org/10.5281/zenodo.13757615" ID-Zenodo-Dep="13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="16" pageNumber="301" startId="16.[151,250,1296,1320]" targetBox="[163,1424,194,1259]" targetPageId="16" targetType="figure">
<paragraph id="8BD5E0567B2EFFE3FF1C8F688543FAFD" blockId="16.[151,1436,1296,1356]" pageId="16" pageNumber="301">
<emphasis id="B91E3C447B2EFFE3FF1C8F68811BFA99" bold="true" box="[151,291,1296,1320]" pageId="16" pageNumber="301">FIGURE 12.</emphasis>
<taxonomicName id="4C6A9BD57B2EFFE3FEA78F6881D2FA99" authorityName="Telnov" authorityYear="2024" box="[300,490,1296,1320]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="301" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B2EFFE3FEA78F6881D2FA99" box="[300,490,1296,1320]" italics="true" pageId="16" pageNumber="301">Microhoria lingua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2EFFE3FE788F68827CFA96" box="[499,580,1296,1319]" pageId="16" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B2EFFE2FF4C8FE6842FFD40" blockId="16.[151,1437,1438,2041]" lastBlockId="17.[151,1437,151,1005]" lastPageId="17" lastPageNumber="302" pageId="16" pageNumber="301">
<emphasis id="B91E3C447B2EFFE3FF4C8FE68161FA09" bold="true" box="[199,345,1438,1464]" pageId="16" pageNumber="301">Description.</emphasis>
<typeStatus id="54D15EF47B2EFFE3FEE88FE681F3FA09" box="[355,459,1438,1464]" pageId="16" pageNumber="301" type="holotype">Holotype</typeStatus>
male. Dorsal and ventral head and pronotum brown, the latter slightly paler. Elytra brown, each elytron with poorly defined yellowish humeral (smaller) and postmedian (larger) spot; spots distinctly separated, not confluent. Maxillary palpus and antenna yellowish brown, three basal antennomeres yellow. Legs yellowish brown with paler tarsi. Ventral pterothorax and abdominal ventrites brown. Head elongate elliptical, about 1.1× as long as wide, slightly convex in dorsal aspect, moderately glossy. Labrum subtruncate at anterior margin. Frontoclypeal suture present, slightly arched (shallowly concave anteriorly). Compound eye moderate, about as long as slightly converging tempus, strongly protruding from lateral outline of head. Head base rounded, posterior temporal angles rounded. Head dorsal punctures minute, sparse. Intervening spaces smooth and glossy, much wider than punctures. Dorsal cranial setae whitish to yellowish, sparse, inconspicuous. Antenna extending slightly beyond elytral base when directed posteriad, hardly thickened in apical half. Antennomeres 6–10 somewhat widened and shortened, more regularly cylindrical to trapezoid in dorsal view, of them 9–10 hardly widened to not widened distally. Penultimate antennomere about 1.5× as long as wide. Terminal antennomere elongate, bluntly pointed, twice as long as penultimate antennomere, about the combined length of antennomeres 9–10. Penultimate maxillary palpomere lobate at mesal margin. Terminal maxillary palpomere securiform. Cranial ‘neck’ about one third head width across compound eyes, densely punctured, subopaque. Pronotum slightly longer than wide, distinctly narrower than head across eyes, subtruncate medially at anterior margin. Pronotal disc moderately glossy, slightly convex in dorsal aspect. Lateral margins broadly rounded in anterior half, strongly constricted postmedium. Anterior pronotal rim distinct dorsally, laterally and ventrally. Antebasal sulcus wide, dorsally poorly defined, well-developed laterally. Pronotal punctures minute and sparse on anterior portion of pronotal disc and on lateral sides of anterior lobe, large and dense on most of pronotal disc in particular in constriction area. Large punctures are of irregular shape, puncture background microreticulate or corrugate. Intervening spaces glossy, much wider than punctures near anterior margin, narrower than to about as wide as those on rest of pronotal dorsum. Pronotal setae as those on head, somewhat longer, suberect. Scutellar shield nearly triangular, apically rounded, glossy. Elytra elongate, about 1.7–1.8× as long as wide, widest at midlength, dorsally flattened to slightly convex. Postbasal transverse impression very feeble, nearly obsolete. Humerus broadly rounded. Apical sutural angle rounded. Apex of elytron modified, with a short, obtuse denticle at opening of a gland channel. Elytral surface glossy and smooth, punctures smaller than those on pronotal constriction, but distinct and dense. Intervening spaces smooth and glossy, generally as wide as punctures. Elytral setae yellowish, long and dense, subdecumbent, directed posteriad. Metathoracic wing fully developed (functional). Legs without modifications, moderately densely setose. Tibial terminal spurs paired. Basal metatarsomere slightly longer than combined length of remaining metatarsomeres. Tergite VII broadly rounded to subtruncate at posterior margin (
<figureCitation id="1351FCD37B2FFFE2FD8488138242FD34" box="[527,634,619,645]" captionStart="FIGURE 12" captionStartId="16.[151,250,1296,1320]" captionTargetBox="[163,1424,194,1259]" captionTargetId="figure-28@16.[151,1436,181,1271]" captionTargetPageId="16" captionText="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="17" pageNumber="302">Fig. 12A</figureCitation>
). Morphological sternite VII broadly triangular, shallowly emarginate medially at posterior margin (
<figureCitation id="1351FCD37B2FFFE2FE5488F7827DFD18" box="[479,581,655,681]" captionStart="FIGURE 12" captionStartId="16.[151,250,1296,1320]" captionTargetBox="[163,1424,194,1259]" captionTargetId="figure-28@16.[151,1436,181,1271]" captionTargetPageId="16" captionText="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="17" pageNumber="302">Fig. 12B</figureCitation>
). Tergite VIII broadly rounded at posterior margin (
<figureCitation id="1351FCD37B2FFFE2FBF588F784DCFD18" box="[1150,1252,655,681]" captionStart="FIGURE 12" captionStartId="16.[151,250,1296,1320]" captionTargetBox="[163,1424,194,1259]" captionTargetId="figure-28@16.[151,1436,181,1271]" captionTargetPageId="16" captionText="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="17" pageNumber="302">Fig. 12C</figureCitation>
). Morphological sternite IX Y-shaped, arms short (
<figureCitation id="1351FCD37B2FFFE2FD9C88CB82BAFD7C" box="[535,642,691,717]" captionStart="FIGURE 12" captionStartId="16.[151,250,1296,1320]" captionTargetBox="[163,1424,194,1259]" captionTargetId="figure-28@16.[151,1436,181,1271]" captionTargetPageId="16" captionText="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="17" pageNumber="302">Fig. 12D</figureCitation>
). Aedeagus (
<figureCitation id="1351FCD37B2FFFE2FC9F88CB83AEFD7C" box="[788,918,691,717]" captionStart="FIGURE 12" captionStartId="16.[151,250,1296,1320]" captionTargetBox="[163,1424,194,1259]" captionTargetId="figure-28@16.[151,1436,181,1271]" captionTargetPageId="16" captionText="FIGURE 12. Microhoria lingua sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite IX; E—Aedeagus; F—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757615" httpUri="https://zenodo.org/record/13757615/files/figure.png" pageId="17" pageNumber="302">Fig. 12E–F</figureCitation>
) elongate; endophallic armature of numerous hair-like spines gathered to form a ‘spikelet’ resembling those of some
<taxonomicName id="4C6A9BD57B2FFFE2FC3888AF842AFD40" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[947,1042,727,753]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="302" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
.
</paragraph>
<paragraph id="8BD5E0567B2FFFE2FF4C8882832FFC5C" blockId="17.[151,1437,151,1005]" pageId="17" pageNumber="302">
<emphasis id="B91E3C447B2FFFE2FF4C8882818DFCA5" bold="true" box="[199,437,762,788]" pageId="17" pageNumber="302">Sexual dimorphism.</emphasis>
Female (
<figureCitation id="1351FCD37B2FFFE2FD928883829BFCA4" box="[537,675,763,789]" captionStart="FIGURE 11" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[331,1249,194,1907]" captionTargetId="figure-18@15.[325,1261,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Microhoria lingua sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757611" httpUri="https://zenodo.org/record/13757611/files/figure.png" pageId="17" pageNumber="302">Figs 11C–D</figureCitation>
,
<figureCitation id="1351FCD37B2FFFE2FD27888382F0FCA4" box="[684,712,763,789]" captionStart="FIGURE 13" captionStartId="17.[151,250,1911,1935]" captionTargetBox="[197,1387,1041,1874]" captionTargetId="figure-399@17.[188,1399,1029,1887]" captionTargetPageId="17" captionText="FIGURE 13. Microhoria lingua sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757617" httpUri="https://zenodo.org/record/13757617/files/figure.png" pageId="17" pageNumber="302">13</figureCitation>
) larger and stouter, elytra stronger widened in median portion and comparatively stronger convex in dorsal aspect. Elytral sutural angle produced into a long, acutely angulate process which is slightly curved ventrally at apex (
<figureCitation id="1351FCD37B2FFFE2FD06893B82CDFCEC" box="[653,757,835,861]" captionStart="FIGURE 11" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[331,1249,194,1907]" captionTargetId="figure-18@15.[325,1261,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Microhoria lingua sp. nov. A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757611" httpUri="https://zenodo.org/record/13757611/files/figure.png" pageId="17" pageNumber="302">Fig. 11C</figureCitation>
). Tergite VII strongly modified, with dorsal median prong serrate at all margins, deep V-shaped incision at posterior margin and, each side of it, with posterolateral margin of tergite deflected dorsally, with irregularly serrate margins of each deflection (
<figureCitation id="1351FCD37B2FFFE2FC7589F384B0FC14" box="[1022,1160,907,933]" captionStart="FIGURE 13" captionStartId="17.[151,250,1911,1935]" captionTargetBox="[197,1387,1041,1874]" captionTargetId="figure-399@17.[188,1399,1029,1887]" captionTargetPageId="17" captionText="FIGURE 13. Microhoria lingua sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757617" httpUri="https://zenodo.org/record/13757617/files/figure.png" pageId="17" pageNumber="302">Fig. 13A–B</figureCitation>
). Morphological sternite VII broad, obtusely pointed and shallowly emarginate medially at posterior margin (
<figureCitation id="1351FCD37B2FFFE2FBF889D784E6FC78" box="[1139,1246,943,969]" captionStart="FIGURE 13" captionStartId="17.[151,250,1911,1935]" captionTargetBox="[197,1387,1041,1874]" captionTargetId="figure-399@17.[188,1399,1029,1887]" captionTargetPageId="17" captionText="FIGURE 13. Microhoria lingua sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757617" httpUri="https://zenodo.org/record/13757617/files/figure.png" pageId="17" pageNumber="302">Fig. 13C</figureCitation>
). Morphological sternite IX rod-like (
<figureCitation id="1351FCD37B2FFFE2FEF689AB81DDFC5C" box="[381,485,979,1005]" captionStart="FIGURE 13" captionStartId="17.[151,250,1911,1935]" captionTargetBox="[197,1387,1041,1874]" captionTargetId="figure-399@17.[188,1399,1029,1887]" captionTargetPageId="17" captionText="FIGURE 13. Microhoria lingua sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757617" httpUri="https://zenodo.org/record/13757617/files/figure.png" pageId="17" pageNumber="302">Fig. 13D</figureCitation>
). Ovipositor as in fig. 13D.
</paragraph>
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<paragraph id="8BD5E0567B2FFFE2FF1C8D0F817CF84A" blockId="17.[151,1437,1911,2043]" pageId="17" pageNumber="302">
<emphasis id="B91E3C447B2FFFE2FF1C8D0F811BF83E" bold="true" box="[151,291,1911,1935]" pageId="17" pageNumber="302">FIGURE 13.</emphasis>
<taxonomicName id="4C6A9BD57B2FFFE2FEA68D0F81D4F83E" authorityName="Telnov" authorityYear="2024" box="[301,492,1911,1935]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="302" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B2FFFE2FEA68D0F81D4F83E" box="[301,492,1911,1935]" italics="true" pageId="17" pageNumber="302">Microhoria lingua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2FFFE2FE7E8D0F8272F83F" box="[501,586,1911,1934]" pageId="17" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view [lateral margins of the tergite are deflected dorsally in the image, the real shape of this tergite is much wider triangular, than depicted]; C—Morphological sternite VII, ventral view; D—Ovipositor, morphological sternite IX [not to scale].
</paragraph>
</caption>
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<emphasis id="B91E3C447B2CFFE1FF4C8AEE81D0FF01" bold="true" box="[199,488,150,176]" pageId="18" pageNumber="303">Intraspecific variability.</emphasis>
Dorsal and ventral forebody dark reddish in some
<typeStatus id="54D15EF47B2CFFE1FBA78AEF84A3FF00" box="[1068,1179,151,177]" pageId="18" pageNumber="303" type="paratype">paratypes</typeStatus>
. The shape and size of pale elytral spots strongly vary from small to (jointly) occupying about a half of elytra. Posterior pale paired spot extends to cover whole apical half of elytra but also in that case posterior spots are separated by a dark sutural area. Dorsal pronotal punctures on constriction vary strongly in shape and size. Total body length varies from
<quantity id="4C924DB37B2CFFE1FAB98B7B80FEFEF0" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.4" metricValueMin="2.4" pageId="18" pageNumber="303" unit="mm" value="2.9" valueMax="3.4" valueMin="2.4">2.4 to 3.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2CFFE1FF4C8B328558FE1C" pageId="18" pageNumber="303" type="diagnosis">
<paragraph id="8BD5E0567B2CFFE1FF4C8B328558FE1C" blockId="18.[151,1437,150,537]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C8B3281EBFED5" bold="true" box="[199,467,330,356]" pageId="18" pageNumber="303">Differential diagnosis.</emphasis>
<taxonomicName id="4C6A9BD57B2CFFE1FE548B33828BFED4" authorityName="Telnov" authorityYear="2024" box="[479,691,331,357]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="species" species="lingua" status="sp. nov.">
<emphasis id="B91E3C447B2CFFE1FE548B33828BFED4" box="[479,691,331,357]" italics="true" pageId="18" pageNumber="303">Microhoria lingua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2CFFE1FD358B338322FED4" box="[702,794,331,357]" pageId="18" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
is externally similar to Taiwanese
<taxonomicName id="4C6A9BD57B2CFFE1FB368B33850DFED5" authorityName="Telnov" authorityYear="2024" box="[1213,1333,331,356]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="species" species="taguan" status="sp. nov.">
<emphasis id="B91E3C447B2CFFE1FB368B33850DFED5" box="[1213,1333,331,356]" italics="true" pageId="18" pageNumber="303">M. taguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B2CFFE1FACA8B3385A5FED4" box="[1345,1437,331,357]" pageId="18" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
(described herein) and several other similarly-coloured Himalayan and eastern-Palaearctic congeners but is specifically different in the shape of the male aedeagus and the female tergite and morphological sternite VII.
</paragraph>
</subSubSection>
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<paragraph id="8BD5E0567B2CFFE1FF4C8BCF8474FE44" blockId="18.[151,1437,150,537]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C8BCF8114FE60" bold="true" box="[199,300,439,465]" pageId="18" pageNumber="303">Ecology.</emphasis>
Occurs at about
<quantity id="4C924DB37B2CFFE1FE7D8BCE8276FE60" box="[502,590,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="18" pageNumber="303" unit="m" value="1945.0">1945 m</quantity>
in forested area. The studied specimens were ‘swept from vegetation’, a typical microstation of many
<taxonomicName id="4C6A9BD57B2CFFE1FE6A8BA3834CFE44" box="[481,884,475,501]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FE6A8BA38259FE44" box="[481,609,475,501]" italics="true" pageId="18" pageNumber="303">Microhoria</emphasis>
Chevrolat, 1877 species
</taxonomicName>
(see
<bibRefCitation id="EFFB9DA77B2CFFE1FC248BA38478FE44" author="Telnov, D." box="[943,1088,475,501]" pageId="18" pageNumber="303" pagination="195 - 312" refId="ref13736" refString="Telnov, D. (2022) Revisional notes on the genus Microhoria Chevrolat, 1877 (Insecta: Coleoptera: Anthicidae) from the Eastern Mediterranean and Turkey, with new descriptions, an annotated catalogue, and a key. Baltic Journal of Coleopterology, 22 (2), 195 - 312." type="journal article" year="2022">Telnov 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B2CFFE1FF4C8B8684A8FDA8" box="[199,1168,510,537]" pageId="18" pageNumber="303" type="distribution">
<paragraph id="8BD5E0567B2CFFE1FF4C8B8684A8FDA8" blockId="18.[151,1437,150,537]" box="[199,1168,510,537]" pageId="18" pageNumber="303">
<emphasis id="B91E3C447B2CFFE1FF4C8B868159FDA9" bold="true" box="[199,353,510,536]" pageId="18" pageNumber="303">Distribution.</emphasis>
So far known from mountainous central part of
<collectingCountry id="F37DA0C67B2CFFE1FCF58B8783EBFDA8" box="[894,979,511,537]" name="Taiwan" pageId="18" pageNumber="303">Taiwan</collectingCountry>
,
<collectingRegion id="49AE2EB47B2CFFE1FC558B8784B3FDA8" box="[990,1163,511,537]" country="Taiwan" name="Nantou" pageId="18" pageNumber="303">Nantou County</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
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315
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<emphasis id="B91E3C447B34FFF9FF1C8D2C82A1F8DF" bold="true" box="[151,665,1876,1903]" pageId="10" pageNumber="295">
<taxonomicName id="4C6A9BD57B34FFF9FF1C8D2C8141F8DE" authorityName="Telnov" authorityYear="2024" box="[151,377,1876,1903]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="295" phylum="Arthropoda" rank="species" species="bunun" status="sp. nov.">
<emphasis id="B91E3C447B34FFF9FF1C8D2C8141F8DE" bold="true" box="[151,377,1876,1903]" italics="true" pageId="10" pageNumber="295">Microhoria bunun</emphasis>
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(
<figureCitation id="1351FCD37B34FFF9FE668D2C82A8F8DF" box="[493,656,1876,1902]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="11.[151,250,1943,1967]" captionStartId-1="13.[151,250,1674,1698]" captionStartId-2="14.[151,250,1065,1089]" captionTargetBox-0="[318,1269,205,1911]" captionTargetBox-1="[169,1419,194,1637]" captionTargetBox-2="[217,1371,207,1028]" captionTargetId-0="figure-18@11.[307,1279,181,1919]" captionTargetId-1="figure-18@13.[156,1432,181,1650]" captionTargetId-2="figure-28@14.[204,1384,181,1041]" captionTargetPageId-0="11" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 8. Microhoria bunun sp. nov.A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." captionText-1="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." captionText-2="FIGURE 10. Microhoria bunun sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Ovipositor [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.13757601" figureDoi-1="http://doi.org/10.5281/zenodo.13757603" figureDoi-2="http://doi.org/10.5281/zenodo.13757607" httpUri-0="https://zenodo.org/record/13757601/files/figure.png" httpUri-1="https://zenodo.org/record/13757603/files/figure.png" httpUri-2="https://zenodo.org/record/13757607/files/figure.png" pageId="10" pageNumber="295">Figures 8–10</figureCitation>
)
</emphasis>
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</subSubSection>
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<paragraph id="8BD5E0567B34FFF9FF1C8DB88474F86A" blockId="10.[151,1100,1984,2011]" box="[151,1100,1984,2011]" pageId="10" pageNumber="295">
This species belongs to the
<emphasis id="B91E3C447B34FFF9FE478DB8820FF86B" box="[460,567,1984,2010]" italics="true" pageId="10" pageNumber="295">longiceps</emphasis>
species-group as of
<bibRefCitation id="EFFB9DA77B34FFF9FC968DB8847FF86A" author="Kejval, Z. &amp; Chandler, D. S." box="[797,1095,1984,2011]" pageId="10" pageNumber="295" pagination="95 - 154" refId="ref13339" refString="Kejval, Z. &amp; Chandler, D. S. (2020) Generic revision of the Microhoriini with new species and synonymies from the Palaearctic Region (Coleoptera: Anthicidae). Acta entomologica musei nationalis pragae, 60 (1), 95 - 154. https: // doi. org / 10.37520 / aemnp. 2020.007" type="journal article" year="2020">Kejval &amp; Chandler (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF15B0DE7B35FFF8FF1C8DEF837CF862" ID-DOI="http://doi.org/10.5281/zenodo.13757601" ID-Zenodo-Dep="13757601" httpUri="https://zenodo.org/record/13757601/files/figure.png" pageId="11" pageNumber="296" startId="11.[151,250,1943,1967]" targetBox="[318,1269,205,1911]" targetPageId="11" targetType="figure">
<paragraph id="8BD5E0567B35FFF8FF1C8DEF837CF862" blockId="11.[151,1436,1943,2003]" pageId="11" pageNumber="296">
<emphasis id="B91E3C447B35FFF8FF1C8DEF812AF81E" bold="true" box="[151,274,1943,1967]" pageId="11" pageNumber="296">FIGURE 8.</emphasis>
<taxonomicName id="4C6A9BD57B35FFF8FE938DEF81E9F81E" authorityName="Telnov" authorityYear="2024" box="[280,465,1943,1967]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="296" phylum="Arthropoda" rank="species" species="bunun" status="sp. nov.">
<emphasis id="B91E3C447B35FFF8FE938DEF81E9F81E" box="[280,465,1943,1967]" italics="true" pageId="11" pageNumber="296">Microhoria bunun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B35FFF8FE5C8DE0821DF81E" box="[471,549,1944,1967]" pageId="11" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale].
</paragraph>
</caption>
<subSubSection id="C370B3DD7B32FFFFFF4C8AEF843BFED4" pageId="12" pageNumber="297" type="materials_examined">
<paragraph id="8BD5E0567B32FFFFFF4C8AEF8524FF64" blockId="12.[151,1437,150,1761]" pageId="12" pageNumber="297">
<emphasis id="B91E3C447B32FFFFFF4C8AEF8231FF01" bold="true" box="[199,521,150,177]" pageId="12" pageNumber="297">Type material designated.</emphasis>
<materialsCitation id="3B02EA0B7B32FFFFFD9F8AEF8524FF64" collectingDate="2008-08-06" collectionCode="BMNH" collectorName="M. V. L. Barclay &amp; H. Mendel &amp; R. Ewers" country="Taiwan" elevation="1920" latitude="24.042166" location="Nantou County" longLatPrecision="1" longitude="121.20925" pageId="12" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54D15EF47B32FFFFFD9F8AEF8246FF00" box="[532,638,151,177]" pageId="12" pageNumber="297" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6C2BDF7B32FFFFFD038AEF82A7FF00" box="[648,671,151,177]" count="1" pageId="12" pageNumber="297" type="male">♂</specimenCount>
,
<collectionCode id="ED7B78937B32FFFFFD258AEF8330FF00" box="[686,776,151,177]" pageId="12" pageNumber="297">BMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B32FFFFFC9E8AEF83B3FF00" box="[789,907,151,177]" name="Taiwan" pageId="12" pageNumber="297">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B32FFFFFC118AEF8474FF00" box="[922,1100,151,177]" country="Taiwan" name="Nantou" pageId="12" pageNumber="297">Nantou County</collectingRegion>
<geoCoordinate id="EE5E86917B32FFFFFBDD8AEE84D6FF00" box="[1110,1262,150,177]" degrees="24" direction="north" minutes="02.530" orientation="latitude" pageId="12" pageNumber="297" precision="1" value="24.042166">N24°02.530’</geoCoordinate>
;
<geoCoordinate id="EE5E86917B32FFFFFB768AEE85A5FF00" box="[1277,1437,150,177]" degrees="121" direction="east" minutes="12.555" orientation="longitude" pageId="12" pageNumber="297" precision="1" value="121.20925">E121°12.555’</geoCoordinate>
beating etc.
<quantity id="4C924DB37B32FFFFFEA88AC3814AFF64" box="[291,370,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="12" pageNumber="297" unit="m" value="1920.0">
<elevation id="004707657B32FFFFFEA88AC3814AFF64" box="[291,370,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="12" pageNumber="297" unit="m" value="1920.0">1920m</elevation>
</quantity>
<date id="FFD4C6967B32FFFFFEF28AC381CEFF64" box="[377,502,187,213]" pageId="12" pageNumber="297" value="2008-08-06">
<collectingDate id="EF903F7E7B32FFFFFEF28AC381CEFF64" box="[377,502,187,213]" pageId="12" pageNumber="297" value="2008-08-06">6.viii.2008</collectingDate>
</date>
<collectorName id="269F85807B32FFFFFE768AC3828DFF64" box="[509,693,187,213]" pageId="12" pageNumber="297">M.V.L. Barclay</collectorName>
,
<collectorName id="269F85807B32FFFFFD4A8AC3830CFF64" box="[705,820,187,213]" pageId="12" pageNumber="297">H.Mendel</collectorName>
&amp;
<collectorName id="269F85807B32FFFFFCD38AC383F9FF64" box="[856,961,187,213]" pageId="12" pageNumber="297">R. Ewers</collectorName>
<collectionCode id="ED7B78937B32FFFFFC428AC3841AFF64" box="[969,1058,187,213]" pageId="12" pageNumber="297">BMNH</collectionCode>
(E) 2008-85 [printed].
</materialsCitation>
</paragraph>
<paragraph id="8BD5E0567B32FFFFFF4C8AA7843BFED4" blockId="12.[151,1437,150,1761]" pageId="12" pageNumber="297">
<typeStatus id="54D15EF47B32FFFFFF4C8AA7810EFF48" box="[199,310,223,249]" pageId="12" pageNumber="297" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D6C2BDF7B32FFFFFEB68AA781F6FF48" box="[317,462,223,249]" count="7" pageId="12" pageNumber="297" type="generic">7 specimens</specimenCount>
.
<specimenCount id="9D6C2BDF7B32FFFFFE538AA78238FF48" box="[472,512,223,249]" count="4" pageId="12" pageNumber="297" type="male">4♂</specimenCount>
<specimenCount id="9D6C2BDF7B32FFFFFD8A8AA78229FF48" box="[513,529,223,249]" count="1" pageId="12" pageNumber="297" type="female">♀</specimenCount>
<materialsCitation id="3B02EA0B7B32FFFFFD938AA783BBFF48" box="[536,899,223,249]" collectionCode="BMNH" pageId="12" pageNumber="297" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED7B78937B32FFFFFD938AA7824AFF48" box="[536,626,223,249]" pageId="12" pageNumber="297">BMNH</collectionCode>
: same label as
<typeStatus id="54D15EF47B32FFFFFC908AA783BBFF48" box="[795,899,223,249]" pageId="12" pageNumber="297" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<specimenCount id="9D6C2BDF7B32FFFFFC058AA78389FF48" box="[910,945,223,249]" count="1" pageId="12" pageNumber="297" type="female">1♀</specimenCount>
<materialsCitation id="3B02EA0B7B32FFFFFC328AA78119FEF0" collectingDate="2008-08-06" collectionCode="BMNH" collectorName="H. Mendel &amp; Barclay" country="Taiwan" elevation="1165" latitude="24.01725" location="0.6 km SE of Lushan" longLatPrecision="1" longitude="121.18847" pageId="12" pageNumber="297" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
<collectionCode id="ED7B78937B32FFFFFC328AA7842BFF48" box="[953,1043,223,249]" pageId="12" pageNumber="297">BMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B32FFFFFB968AA784ABFF48" box="[1053,1171,223,249]" name="Taiwan" pageId="12" pageNumber="297">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B32FFFFFB158AA78575FF48" box="[1182,1357,223,249]" country="Taiwan" name="Nantou" pageId="12" pageNumber="297">Nantou County</collectingRegion>
<location id="8EB5B68D7B32FFFFFADF8AA7810BFEAC" LSID="urn:lsid:plazi:treatment:03C351407B34FFFDFF1C8D2C84A8FA1B:8EB5B68D7B32FFFFFADF8AA7810BFEAC" country="Taiwan" latitude="24.01725" longLatPrecision="1" longitude="121.18847" name="0.6 km SE of Lushan" pageId="12" pageNumber="297" stateProvince="Nantou">
<quantity id="4C924DB37B32FFFFFADF8AA785A4FF48" box="[1364,1436,223,249]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="297" unit="km" value="0.6">0.6km</quantity>
SE of Lushan
</location>
,
<quantity id="4C924DB37B32FFFFFEB58B7A81B5FEAC" box="[318,397,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.165" pageId="12" pageNumber="297" unit="m" value="1165.0">
<elevation id="004707657B32FFFFFEB58B7A81B5FEAC" box="[318,397,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.165" pageId="12" pageNumber="297" unit="m" value="1165.0">1165m</elevation>
</quantity>
<geoCoordinate id="EE5E86917B32FFFFFE188B7A8227FEAC" box="[403,543,258,285]" degrees="24" direction="north" minutes="01.035" orientation="latitude" pageId="12" pageNumber="297" precision="1" value="24.01725">N24°01.035</geoCoordinate>
<geoCoordinate id="EE5E86917B32FFFFFDAE8B7B8283FEAC" box="[549,699,259,285]" degrees="121" direction="east" minutes="11.308" orientation="longitude" pageId="12" pageNumber="297" precision="1" value="121.18847">E121°11.308</geoCoordinate>
at light,
<date id="FFD4C6967B32FFFFFC958B7B83A2FEAC" box="[798,922,259,285]" pageId="12" pageNumber="297" value="2008-08-06">
<collectingDate id="EF903F7E7B32FFFFFC958B7B83A2FEAC" box="[798,922,259,285]" pageId="12" pageNumber="297" value="2008-08-06">6.viii.2008</collectingDate>
</date>
<collectorName id="269F85807B32FFFFFC2B8B7B842CFEAC" box="[928,1044,259,285]" pageId="12" pageNumber="297">H.Mendel</collectorName>
&amp; MVL
<collectorName id="269F85807B32FFFFFBF08B7B84ECFEAC" box="[1147,1236,259,285]" pageId="12" pageNumber="297">Barclay</collectorName>
<collectionCode id="ED7B78937B32FFFFFB518B7B850BFEAC" box="[1242,1331,259,285]" pageId="12" pageNumber="297">BMNH</collectionCode>
(E) 2008- 85 [printed]
</materialsCitation>
;
<specimenCount id="9D6C2BDF7B32FFFFFEA18B5F8176FEF0" box="[298,334,295,321]" count="2" pageId="12" pageNumber="297" type="female">2♀</specimenCount>
<materialsCitation id="3B02EA0B7B32FFFFFED88B5F83C7FED4" collectingDate="2008-08-08" collectionCode="BMNH" collectorName="M. V. L. Barclay &amp; H. Mendel" country="Taiwan" elevation="1268" latitude="24.024683" location="0.5 km NW of Lushan" longLatPrecision="1" longitude="121.18127" pageId="12" pageNumber="297" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
<collectionCode id="ED7B78937B32FFFFFED88B5F8195FEF0" box="[339,429,295,321]" pageId="12" pageNumber="297">BMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B32FFFFFE3E8B5F8213FEF0" box="[437,555,295,321]" name="Taiwan" pageId="12" pageNumber="297">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B32FFFFFDBE8B5F82DAFEF0" box="[565,738,295,321]" country="Taiwan" name="Nantou" pageId="12" pageNumber="297">Nantou County</collectingRegion>
<location id="8EB5B68D7B32FFFFFD6C8B5E83E2FEF0" LSID="urn:lsid:plazi:treatment:03C351407B34FFFDFF1C8D2C84A8FA1B:8EB5B68D7B32FFFFFD6C8B5E83E2FEF0" box="[743,986,294,321]" country="Taiwan" latitude="24.024683" longLatPrecision="1" longitude="121.18127" name="0.5 km NW of Lushan" pageId="12" pageNumber="297" stateProvince="Nantou">
<quantity id="4C924DB37B32FFFFFD6C8B5E8308FEF0" box="[743,816,294,321]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="297" unit="km" value="0.5">0.5km</quantity>
NW of Lushan
</location>
<geoCoordinate id="EE5E86917B32FFFFFC548B5F8452FEF0" box="[991,1130,295,321]" degrees="24" direction="north" minutes="01.481" orientation="latitude" pageId="12" pageNumber="297" precision="1" value="24.024683">N24°01.481</geoCoordinate>
<geoCoordinate id="EE5E86917B32FFFFFBE48B5F853DFEF0" box="[1135,1285,295,321]" degrees="121" direction="east" minutes="10.876" orientation="longitude" pageId="12" pageNumber="297" precision="1" value="121.18127">E121°10.876</geoCoordinate>
<date id="FFD4C6967B32FFFFFA808B5F85AFFEF0" box="[1291,1431,295,321]" pageId="12" pageNumber="297" value="2008-08-08">
<collectingDate id="EF903F7E7B32FFFFFA808B5F85AFFEF0" box="[1291,1431,295,321]" pageId="12" pageNumber="297" value="2008-08-08">08.viii.2008</collectingDate>
</date>
,
<quantity id="4C924DB37B32FFFFFF1C8B3380D1FED4" box="[151,233,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="12" pageNumber="297" unit="m" value="1268.0">
<elevation id="004707657B32FFFFFF1C8B3380D1FED4" box="[151,233,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="12" pageNumber="297" unit="m" value="1268.0">1268m</elevation>
</quantity>
, at light,
<collectorName id="269F85807B32FFFFFEDF8B338231FED4" box="[340,521,331,357]" pageId="12" pageNumber="297">M.V.L. Barclay</collectorName>
&amp;
<collectorName id="269F85807B32FFFFFDA58B338290FED4" box="[558,680,331,357]" pageId="12" pageNumber="297">H. Mendel</collectorName>
<collectionCode id="ED7B78937B32FFFFFD3B8B338331FED4" box="[688,777,331,357]" pageId="12" pageNumber="297">BMNH</collectionCode>
(E) 2008-85 [printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B32FFFFFF4C8B16852BFE38" box="[199,1299,366,393]" pageId="12" pageNumber="297" type="etymology">
<paragraph id="8BD5E0567B32FFFFFF4C8B16852BFE38" blockId="12.[151,1437,150,1761]" box="[199,1299,366,393]" pageId="12" pageNumber="297">
<emphasis id="B91E3C447B32FFFFFF4C8B168191FE39" bold="true" box="[199,425,366,392]" pageId="12" pageNumber="297">Derivatio nominis.</emphasis>
Named after the Taiwan’s Bunun people and language. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B32FFFDFF4C8BEB83DDFB63" lastPageId="14" lastPageNumber="299" pageId="12" pageNumber="297" type="description">
<paragraph id="8BD5E0567B32FFFFFF4C8BEB84E6FDA8" blockId="12.[151,1437,150,1761]" pageId="12" pageNumber="297">
<emphasis id="B91E3C447B32FFFFFF4C8BEB8142FE1C" bold="true" box="[199,378,403,429]" pageId="12" pageNumber="297">Measurements</emphasis>
,
<typeStatus id="54D15EF47B32FFFFFE068BEB81C9FE1C" box="[397,497,403,429]" pageId="12" pageNumber="297" type="holotype">holotype</typeStatus>
male, total body length excluding partially exposed terminalia
<quantity id="4C924DB37B32FFFFFB618BEA8560FE1C" box="[1258,1368,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="12" pageNumber="297" unit="mm" value="2.65">2.65 mm</quantity>
; head length
<quantity id="4C924DB37B32FFFFFF6C8BCE8169FE60" box="[231,337,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="297" unit="mm" value="0.55">0.55 mm</quantity>
, head width across compound eyes
<quantity id="4C924DB37B32FFFFFD7A8BCE8374FE60" box="[753,844,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="297" unit="mm" value="0.5">0.5 mm</quantity>
, pronotal length
<quantity id="4C924DB37B32FFFFFB9B8BCE8453FE60" box="[1040,1131,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="297" unit="mm" value="0.5">0.5 mm</quantity>
, maximum pronotal width
<quantity id="4C924DB37B32FFFFFF1C8BA380C8FE44" box="[151,240,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="297" unit="mm" value="0.4">0.4 mm</quantity>
, minimum pronotal width in constriction area
<quantity id="4C924DB37B32FFFFFD768BA3836FFE44" box="[765,855,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="297" unit="mm" value="0.3">0.3 mm</quantity>
, elytral length
<quantity id="4C924DB37B32FFFFFC748BA38460FE44" box="[1023,1112,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="297" unit="mm" value="1.6">1.6 mm</quantity>
, combined maximum elytral width across midlength
<quantity id="4C924DB37B32FFFFFE218B8781C8FDA8" box="[426,496,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="297" unit="mm" value="1.0">1 mm</quantity>
. Male
<typeStatus id="54D15EF47B32FFFFFDB78B878292FDA8" box="[572,682,511,537]" pageId="12" pageNumber="297" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B32FFFFFD3A8B868305FDA8" box="[689,829,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="12" pageNumber="297" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
, female
<typeStatus id="54D15EF47B32FFFFFC158B878434FDA8" box="[926,1036,511,537]" pageId="12" pageNumber="297" type="paratype">paratypes</typeStatus>
<quantity id="4C924DB37B32FFFFFB988B8684A5FDA8" box="[1043,1181,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="12" pageNumber="297" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
long.
</paragraph>
<paragraph id="8BD5E0567B32FFFFFF4C885A818FF950" blockId="12.[151,1437,150,1761]" pageId="12" pageNumber="297">
<emphasis id="B91E3C447B32FFFFFF4C885A8159FD8D" bold="true" box="[199,353,546,572]" pageId="12" pageNumber="297">Description.</emphasis>
<typeStatus id="54D15EF47B32FFFFFEFA885B81E7FD8C" box="[369,479,547,573]" pageId="12" pageNumber="297" type="holotype">Holotype</typeStatus>
male. Dorsum and venter uniformly black to black-brown. Antennomeres 2–4, tibiae and tarsi yellowish brown. Head elliptical, hardly longer than wide, slightly convex in dorsal aspect, moderately glossy. Labrum subtruncate at anterior margin. Frontoclypeal suture present, slightly arched (shallowly concave anteriorly). Compound eye moderate, strongly protruding from lateral outline of head. Head base rounded in one broad arc with tempora. Head dorsal punctures moderately large, dense. Intervening spaces smooth and glossy, as wide as to twice as wide as punctures. Dorsal cranial setae whitish to yellowish, sparse, inconspicuous. Antenna extending slightly beyond elytral base when directed posteriad, hardly thickened in apical half. Antennomeres 7–10 widened distally. Penultimate antennomere hardly longer than wide. Terminal antennomere elongate, bluntly pointed, about 1.8× as long as penultimate antennomere, about the combined length of antennomeres 9–10. Penultimate maxillary palpomere lobate at mesal margin. Terminal maxillary palpomere securiform. Cranial ‘neck’ less than one third head width across compound eyes, densely punctured, moderately glossy. Pronotum longer than wide, narrower than head across eyes, subtruncate medially at anterior margin. Pronotal disc moderately glossy, flattened in dorsal aspect. Lateral margins broadly rounded in anterior half, strongly constricted postmedium. Anterior pronotal rim distinct dorsally, laterally and ventrally. Antebasal sulcus wide, less prominent dorsally, well-developed laterally. Pronotal punctures on anterior portion of pronotal disc larger than those on head, minute and sparse on lateral sides of anterior lobe, large, dense and coarse on basal half of disc in particular in constriction area. Large punctures are irregularly shaped, crateriform. Intervening spaces glossy, about as wide as punctures on anterior half of pronotal disc, much narrower than those on rest of pronotal dorsum. Pronotal setae as those on head, not fully appressed. Scutellar shield apically rounded, moderately glossy. Elytra elongate, about 1.6× as long as wide, widest at midlength, dorsally flattened. Postbasal transverse impression obsolete. Humerus broadly rounded. Apical sutural angle rounded. Apex of elytron modified, with a short, semicircular protuberance at opening of a gland channel, adjacent to apical sutural angle of elytron. Elytral surface moderately glossy and smooth, punctures smaller than those on pronotal constriction, but distinct and dense. Intervening spaces glossy, generally as wide as to twice as wide as punctures. Elytral setae whitish to yellowish, moderately long and dense, not fully appressed, directed posteriad. Metathoracic wing fully developed (functional). Legs without modifications, moderately densely setose. Tibial terminal spurs paired, inconspicuous. Basal metatarsomere longer than combined length of remaining metatarsomeres. Tergite VII broadly rounded to subtruncate at posterior margin (
<figureCitation id="1351FCD37B32FFFFFDCD8C6B8291F99C" box="[582,681,1555,1581]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9A</figureCitation>
). Morphological sternite VII subtruncate medially at posterior margin (
<figureCitation id="1351FCD37B32FFFFFF778C4F8163F9E0" box="[252,347,1591,1617]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9B</figureCitation>
). Morphological tergite VIII rounded at posterior margin (
<figureCitation id="1351FCD37B32FFFFFB938C4F844FF9E0" box="[1048,1143,1591,1617]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9C</figureCitation>
). Morphological sternite VIII glasses-shaped, sclerites narrowly interconnected (
<figureCitation id="1351FCD37B32FFFFFCB38C2383A1F9C4" box="[824,921,1627,1653]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9D</figureCitation>
), sternite IX Y-shaped, arms short, arched (
<figureCitation id="1351FCD37B32FFFFFF2B8C078138F928" box="[160,256,1663,1689]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9E</figureCitation>
). Aedeagus (
<figureCitation id="1351FCD37B32FFFFFE298C07821AF928" box="[418,546,1663,1689]" captionStart="FIGURE 9" captionStartId="13.[151,250,1674,1698]" captionTargetBox="[169,1419,194,1637]" captionTargetId="figure-18@13.[156,1432,181,1650]" captionTargetPageId="13" captionText="FIGURE 9. Microhoria bunun sp. nov. holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757603" httpUri="https://zenodo.org/record/13757603/files/figure.png" pageId="12" pageNumber="297">Fig. 9F–H</figureCitation>
) with three central spines nearly as long as tegmen; tegmen preapically trispinose (spines distinctly curved), tegmen apex tongue-shaped, with serrate periphery; endophallic armature somewhat braid-shaped.
</paragraph>
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<paragraph id="8BD5E0567B33FFFEFF1C8CF28344F95B" blockId="13.[151,1437,1674,1770]" pageId="13" pageNumber="298">
<emphasis id="B91E3C447B33FFFEFF1C8CF2812AF913" bold="true" box="[151,274,1674,1698]" pageId="13" pageNumber="298">FIGURE 9.</emphasis>
<taxonomicName id="4C6A9BD57B33FFFEFE928CF281E9F913" authorityName="Telnov" authorityYear="2024" box="[281,465,1674,1698]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="298" phylum="Arthropoda" rank="species" species="bunun" status="sp. nov.">
<emphasis id="B91E3C447B33FFFEFE928CF281E9F913" box="[281,465,1674,1698]" italics="true" pageId="13" pageNumber="298">Microhoria bunun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B33FFFEFE538CF3821EF913" box="[472,550,1675,1698]" pageId="13" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus; G—Median portion, magnified; H—ditto, apex, magnified [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B33FFFEFF4C8D2C84FBF86A" blockId="13.[151,1437,1876,2011]" pageId="13" pageNumber="298">
<emphasis id="B91E3C447B33FFFEFF4C8D2C81F9F8DF" bold="true" box="[199,449,1876,1902]" pageId="13" pageNumber="298">Sexual dimorphism.</emphasis>
Female (
<figureCitation id="1351FCD37B33FFFEFDBC8D2D8285F8DE" box="[567,701,1877,1903]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[318,1269,205,1911]" captionTargetId="figure-18@11.[307,1279,181,1919]" captionTargetPageId="11" captionText="FIGURE 8. Microhoria bunun sp. nov.A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757601" httpUri="https://zenodo.org/record/13757601/files/figure.png" pageId="13" pageNumber="298">Figs 8C–D</figureCitation>
,
<figureCitation id="1351FCD37B33FFFEFD448D2D82D4F8DE" box="[719,748,1877,1903]" captionStart="FIGURE 10" captionStartId="14.[151,250,1065,1089]" captionTargetBox="[217,1371,207,1028]" captionTargetId="figure-28@14.[204,1384,181,1041]" captionTargetPageId="14" captionText="FIGURE 10. Microhoria bunun sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Ovipositor [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757607" httpUri="https://zenodo.org/record/13757607/files/figure.png" pageId="13" pageNumber="298">10</figureCitation>
) elytral apex suntruncate (
<figureCitation id="1351FCD37B33FFFEFBBF8D2D84ACF8DE" box="[1076,1172,1877,1903]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[318,1269,205,1911]" captionTargetId="figure-18@11.[307,1279,181,1919]" captionTargetPageId="11" captionText="FIGURE 8. Microhoria bunun sp. nov.A—Holotype ♂, habitus, dorsal view; B—ditto, forebody, dorsal view; C—Paratype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757601" httpUri="https://zenodo.org/record/13757601/files/figure.png" pageId="13" pageNumber="298">Fig. 8C</figureCitation>
). Tergite VII strongly modified, with deep median apical incision and here medially with a sword-like projection; projection distinctly shorter than corresponding incision (
<figureCitation id="1351FCD37B33FFFEFDDE8DE582D0F806" box="[597,744,1949,1975]" captionStart="FIGURE 10" captionStartId="14.[151,250,1065,1089]" captionTargetBox="[217,1371,207,1028]" captionTargetId="figure-28@14.[204,1384,181,1041]" captionTargetPageId="14" captionText="FIGURE 10. Microhoria bunun sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Ovipositor [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757607" httpUri="https://zenodo.org/record/13757607/files/figure.png" pageId="13" pageNumber="298">Fig. 10A–B</figureCitation>
). Morphological sternite VII broadly triangular, medially angulate projecting (
<figureCitation id="1351FCD37B33FFFEFE0F8DB981D5F86A" box="[388,493,1985,2011]" captionStart="FIGURE 10" captionStartId="14.[151,250,1065,1089]" captionTargetBox="[217,1371,207,1028]" captionTargetId="figure-28@14.[204,1384,181,1041]" captionTargetPageId="14" captionText="FIGURE 10. Microhoria bunun sp. nov., paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Ovipositor [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757607" httpUri="https://zenodo.org/record/13757607/files/figure.png" pageId="13" pageNumber="298">Fig. 10C</figureCitation>
). Morphological sternite IX rod-like. Ovipositor as in fig. 10D.
</paragraph>
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<paragraph id="8BD5E0567B30FFFDFF1C8E5184E6FBD4" blockId="14.[151,1436,1065,1125]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF1C8E51811BFBF0" bold="true" box="[151,291,1065,1089]" pageId="14" pageNumber="299">FIGURE 10.</emphasis>
<taxonomicName id="4C6A9BD57B30FFFDFEA68E5181D2FBF0" authorityName="Telnov" authorityYear="2024" box="[301,490,1065,1089]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="299" phylum="Arthropoda" rank="species" species="bunun" status="sp. nov.">
<emphasis id="B91E3C447B30FFFDFEA68E5181D2FBF0" box="[301,490,1065,1089]" italics="true" pageId="14" pageNumber="299">Microhoria bunun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B30FFFDFE7F8E528271FBF0" box="[500,585,1066,1089]" pageId="14" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀. A—Tergite VII, dorsal view; B—ditto, median portion of posterior margin, magnified, dorsal view; C—Morphological sternite VII, ventral view; D—Ovipositor [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B30FFFDFF4C8ECF83DDFB63" blockId="14.[151,1436,1207,1450]" box="[199,997,1207,1234]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8ECF81DFFB60" bold="true" box="[199,487,1207,1233]" pageId="14" pageNumber="299">Intraspecific variability.</emphasis>
Total body length varies from
<quantity id="4C924DB37B30FFFDFCCA8EC083D9FB63" box="[833,993,1207,1234]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="14" pageNumber="299" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3 to 2.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B30FFFDFF4C8EA381EAFA8F" pageId="14" pageNumber="299" type="diagnosis">
<paragraph id="8BD5E0567B30FFFDFF4C8EA381EAFA8F" blockId="14.[151,1436,1207,1450]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8EA381EBFB44" bold="true" box="[199,467,1243,1269]" pageId="14" pageNumber="299">Differential diagnosis.</emphasis>
<taxonomicName id="4C6A9BD57B30FFFDFE6B8EA3828BFB44" authorityName="Telnov" authorityYear="2024" box="[480,691,1243,1269]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="299" phylum="Arthropoda" rank="species" species="bunun" status="sp. nov.">
<emphasis id="B91E3C447B30FFFDFE6B8EA3828BFB44" box="[480,691,1243,1269]" italics="true" pageId="14" pageNumber="299">Microhoria bunun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B30FFFDFD348EA48324FB47" box="[703,796,1244,1270]" pageId="14" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
is externally similar to several entirely dark-coloured Himalayan and eastern-Palaearctic congeners but is readily different in the shape of the aedeagus and the peculiarly modified female tergite VII.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B30FFFDFF4C8F3084D6FA34" pageId="14" pageNumber="299" type="biology_ecology">
<paragraph id="8BD5E0567B30FFFDFF4C8F3084D6FA34" blockId="14.[151,1436,1207,1450]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8F308114FAD3" bold="true" box="[199,300,1352,1378]" pageId="14" pageNumber="299">Ecology.</emphasis>
Occurs at about
<quantity id="4C924DB37B30FFFDFE718F3F82A1FAD3" box="[506,665,1351,1378]" metricMagnitude="3" metricUnit="m" metricValue="1.5425" metricValueMax="1.92" metricValueMin="1.165" pageId="14" pageNumber="299" unit="m" value="1542.5" valueMax="1920.0" valueMin="1165.0">1165–1920 m</quantity>
in forested area. Collected by beating forest vegetation, a typical microstation of many
<taxonomicName id="4C6A9BD57B30FFFDFE058F138319FA34" box="[398,801,1387,1414]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFE058F138236FA34" box="[398,526,1387,1413]" italics="true" pageId="14" pageNumber="299">Microhoria</emphasis>
Chevrolat, 1877 species
</taxonomicName>
(see
<bibRefCitation id="EFFB9DA77B30FFFDFCD78F1383D4FA37" author="Telnov, D." box="[860,1004,1387,1414]" pageId="14" pageNumber="299" pagination="195 - 312" refId="ref13736" refString="Telnov, D. (2022) Revisional notes on the genus Microhoria Chevrolat, 1877 (Insecta: Coleoptera: Anthicidae) from the Eastern Mediterranean and Turkey, with new descriptions, an annotated catalogue, and a key. Baltic Journal of Coleopterology, 22 (2), 195 - 312." type="journal article" year="2022">Telnov 2022</bibRefCitation>
), and attracted to light.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B30FFFDFF4C8FF784A8FA1B" box="[199,1168,1423,1450]" pageId="14" pageNumber="299" type="distribution">
<paragraph id="8BD5E0567B30FFFDFF4C8FF784A8FA1B" blockId="14.[151,1436,1207,1450]" box="[199,1168,1423,1450]" pageId="14" pageNumber="299">
<emphasis id="B91E3C447B30FFFDFF4C8FF78159FA18" bold="true" box="[199,353,1423,1449]" pageId="14" pageNumber="299">Distribution.</emphasis>
So far known from mountainous central part of
<collectingCountry id="F37DA0C67B30FFFDFCF58FF783EBFA18" box="[894,979,1423,1449]" name="Taiwan" pageId="14" pageNumber="299">Taiwan</collectingCountry>
,
<collectingRegion id="49AE2EB47B30FFFDFC558FE884B3FA1B" box="[990,1163,1424,1450]" country="Taiwan" name="Nantou" pageId="14" pageNumber="299">Nantou County</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="4C6A9BD57B36FFFBFF1C8C8881C6F8B8" authority="(Pic, 1893)" baseAuthorityName="Pic" baseAuthorityYear="1893" box="[151,510,1775,1802]" class="Insecta" family="Anthicidae" genus="Sapintus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="293" phylum="Arthropoda" rank="species" species="marseuli">
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<emphasis id="B91E3C447B36FFFBFF1C8C88814AF8BB" bold="true" box="[151,370,1775,1802]" italics="true" pageId="8" pageNumber="293">Sapintus marseuli</emphasis>
(Pic, 1893)
</emphasis>
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<emphasis id="B91E3C447B36FFFBFF1C8D408182F8E0" bold="true" box="[151,442,1847,1874]" pageId="8" pageNumber="293">New material examined.</emphasis>
<materialsCitation id="3B02EA0B7B36FFFBFE4B8D408243F8C7" collectingDate="1996-07-02" collectionCode="NHMB" collectorName="G. Csorba &amp; L. Nemeth" country="Taiwan" determinerName="Kejval" elevation="150" location="Chintiun" pageId="8" pageNumber="293" specimenCount="1" stateProvince="Taitung">
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:
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, Prov.
<collectingRegion id="49AE2EB47B36FFFBFCEF8D408384F8E3" box="[868,956,1848,1874]" country="Taiwan" name="Taitung" pageId="8" pageNumber="293">Taitung</collectingRegion>
,
<location id="8EB5B68D7B36FFFBFC4C8D408414F8E3" LSID="urn:lsid:plazi:treatment:03C351407B36FFFBFF1C8C8882F9F8C7:8EB5B68D7B36FFFBFC4C8D408414F8E3" box="[967,1068,1848,1874]" country="Taiwan" name="Chintiun" pageId="8" pageNumber="293" stateProvince="Taitung">Chintiun</location>
,
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,
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<collectingDate id="EF903F7E7B36FFFBFB078D408532F8E3" box="[1164,1290,1848,1874]" pageId="8" pageNumber="293" value="1996-07-02">2.VII.1996</collectingDate>
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,
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&amp;
<collectorName id="269F85807B36FFFBFF1C8D248130F8C7" box="[151,264,1884,1910]" pageId="8" pageNumber="293">L.Nemeth</collectorName>
//
<taxonomicName id="4C6A9BD57B36FFFBFEAD8D2481C0F8C7" box="[294,504,1884,1910]" class="Insecta" family="Anthicidae" genus="Sapintus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="293" phylum="Arthropoda" rank="species" species="undetermined">Sapintus Casey sp.</taxonomicName>
<determinerName id="7E090C927B36FFFBFE748D248272F8C7" box="[511,586,1884,1910]" pageId="8" pageNumber="293">Kejval</determinerName>
det.
</materialsCitation>
1997.
</paragraph>
</subSubSection>
</treatment>
</document>

119
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<mods:title id="92E331520825BC410F469CD55AA6331D">New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum</mods:title>
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<taxonomicName id="4C6A9BD57B36FFFBFF1C8F8C8216F9BC" authority="(Marseul, 1876)" baseAuthorityName="Marseul" baseAuthorityYear="1876" box="[151,558,1523,1550]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="293" phylum="Arthropoda" rank="species" species="trigibber">
<emphasis id="B91E3C447B36FFFBFF1C8F8C815AF9BF" bold="true" box="[151,354,1523,1550]" italics="true" pageId="8" pageNumber="293">Nitorus trigibber</emphasis>
(Marseul, 1876)
</taxonomicName>
(
<figureCitation id="1351FCD37B36FFFBFDB58F8B8291F9BC" box="[574,681,1523,1549]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="9.[151,250,1407,1431]" captionStartId-1="10.[151,250,1662,1686]" captionTargetBox-0="[271,1317,192,1369]" captionTargetBox-1="[406,1155,207,1605]" captionTargetId-0="figure-18@9.[258,1330,181,1382]" captionTargetId-1="figure-28@10.[386,1202,181,1638]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 6. Nitorus trigibber (Marseul, 1876). A—♀ from ‘14km W of Chihshang’, habitus, dorsal view; B—ditto, lateral view; C—♀ from ‘Japan’ collected by Lewis, forebody, lateral view [not to scale]." captionText-1="FIGURE 7. Nitorus trigibber (Marseul, 1876), ♂ from ‘Tien Tsin’, China. A—Aedeagus; B—ditto, apex, magnified [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.13757595" figureDoi-1="http://doi.org/10.5281/zenodo.13757597" httpUri-0="https://zenodo.org/record/13757595/files/figure.png" httpUri-1="https://zenodo.org/record/13757597/files/figure.png" pageId="8" pageNumber="293">Figs 6–7</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C370B3DD7B36FFFBFF1C8C4482D7F92F" pageId="8" pageNumber="293" type="materials_examined">
<paragraph id="8BD5E0567B36FFFBFF1C8C4482D7F92F" blockId="8.[151,1437,1595,1694]" pageId="8" pageNumber="293">
<emphasis id="B91E3C447B36FFFBFF1C8C4481F8F9E4" bold="true" box="[151,448,1595,1622]" pageId="8" pageNumber="293">New material examined.</emphasis>
<materialsCitation id="3B02EA0B7B36FFFBFE408C448288F9E7" box="[459,688,1596,1622]" collectionCode="NHMB" pageId="8" pageNumber="293" specimenCount="1">
<specimenCount id="9D6C2BDF7B36FFFBFE408C448275F9E7" box="[459,589,1596,1622]" count="1" pageId="8" pageNumber="293" type="generic">1 specimen</specimenCount>
<collectionCode id="ED7B78937B36FFFBFDD38C448288F9E7" box="[600,688,1596,1622]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="293">NHMB</collectionCode>
</materialsCitation>
:
<materialsCitation id="3B02EA0B7B36FFFBFD368C4481A5F9CB" collectingDate="1997-06-10" collectorName="Chihpen &amp; B. Herczig &amp; L. Ronkay" country="Taiwan" elevation="390" location="Taitung Prov." pageId="8" pageNumber="293" specimenCount="1" stateProvince="Taitung">
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,
<collectingRegion id="49AE2EB47B36FFFBFCB08C4483E0F9E7" box="[827,984,1596,1622]" country="Taiwan" name="Taitung" pageId="8" pageNumber="293">Taitung Prov.</collectingRegion>
,
<collectorName id="269F85807B36FFFBFC6D8C448470F9E7" box="[998,1096,1596,1622]" pageId="8" pageNumber="293">Chihpen</collectorName>
,
<quantity id="4C924DB37B36FFFBFBDD8C44849BF9E7" box="[1110,1187,1596,1622]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="8" pageNumber="293" unit="m" value="390.0">
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, at light,
<date id="FFD4C6967B36FFFBFA9D8C4485A0F9E7" box="[1302,1432,1596,1622]" pageId="8" pageNumber="293" value="1997-06-10">
<collectingDate id="EF903F7E7B36FFFBFA9D8C4485A0F9E7" box="[1302,1432,1596,1622]" pageId="8" pageNumber="293" value="1997-06-10">10.VI.1997</collectingDate>
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&amp;
<collectorName id="269F85807B36FFFBFEA18C1881A5F9CB" box="[298,413,1632,1658]" pageId="8" pageNumber="293">L.Ronkay</collectorName>
</materialsCitation>
;
<materialsCitation id="3B02EA0B7B36FFFBFE2C8C1882B9F9CB" box="[423,641,1632,1658]" collectionCode="BMNH" country="Taiwan" pageId="8" pageNumber="293" specimenCount="1" stateProvince="Taitung">
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<collectionCode id="ED7B78937B36FFFBFDA28C1882B9F9CB" box="[553,641,1632,1658]" pageId="8" pageNumber="293">BMNH</collectionCode>
</materialsCitation>
:
<materialsCitation id="3B02EA0B7B36FFFBFD018C1882D2F92F" collectingDate="2006-11-16" collectionCode="BMNH" collectorName="L. Dembicky" country="Taiwan" elevation="900" latitude="23.15" location="Co." longLatPrecision="1257" longitude="121.066666" pageId="8" pageNumber="293" specimenCode="BMNH 2008-85" specimenCount="1" stateProvince="Taitung">
<collectingCountry id="F37DA0C67B36FFFBFD018C1882C1F9CB" box="[650,761,1632,1658]" name="Taiwan" pageId="8" pageNumber="293">TAIWAN</collectingCountry>
,
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<location id="8EB5B68D7B36FFFBFCD68C1883BFF9CB" LSID="urn:lsid:plazi:treatment:03C351407B36FFFBFF1C8F8C82D7F92F:8EB5B68D7B36FFFBFCD68C1883BFF9CB" box="[861,903,1632,1658]" country="Taiwan" latitude="23.15" longLatPrecision="1257" longitude="121.066666" name="Co." pageId="8" pageNumber="293" stateProvince="Taitung">Co.</location>
,
<quantity id="4C924DB37B36FFFBFC1A8C1883E9F9CB" box="[913,977,1632,1658]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="293" unit="km" value="14.0">14km</quantity>
W of CHIHSHANG,
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,
<quantity id="4C924DB37B36FFFBFF1C8CFC80E2F92F" box="[151,218,1668,1694]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="293" unit="m" value="900.0">
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,
<collectorName id="269F85807B36FFFBFF6E8CFC8149F92F" box="[229,369,1668,1694]" pageId="8" pageNumber="293">L.Dembický</collectorName>
leg.,
<date id="FFD4C6967B36FFFBFE3B8CFC8210F92F" box="[432,552,1668,1694]" pageId="8" pageNumber="293" value="2006-11-16">
<collectingDate id="EF903F7E7B36FFFBFE3B8CFC8210F92F" box="[432,552,1668,1694]" pageId="8" pageNumber="293" value="2006-11-16">16.xi.2006</collectingDate>
</date>
<specimenCode id="DBCC482D7B36FFFBFDA48CFC82D2F92F" box="[559,746,1667,1694]" collectionCode="BMNH" pageId="8" pageNumber="293">BMNH 2008-85</specimenCode>
</materialsCitation>
.
</paragraph>
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143
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, swept, leg.
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:
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isl.
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road
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// J.
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collection.
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(E) 2021-16
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.
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Lewis, 1895
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:
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,
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,
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garden, swept from vegetation,
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, leg.
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;
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:
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Co.,
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, before ESRI
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, //
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, secondary broadleaved forest,
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, L.Dányi&amp;E.Lazányi,TO09-57.
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251
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:
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light
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&amp;
<collectorName id="269F85807B3FFFF2FC2F8D208473F8C0" box="[932,1099,1879,1906]" pageId="1" pageNumber="286">M.V.L.Barclay</collectorName>
<collectionCode id="ED7B78937B3FFFF2FBDA8D2F8490F8C0" box="[1105,1192,1879,1905]" pageId="1" pageNumber="286">BMNH</collectionCode>
(E) 2008-85 [printed].
<collectorName id="269F85807B3FFFF2FF1C8D0380FBF824" box="[151,195,1915,1941]" pageId="1" pageNumber="286">The</collectorName>
left protibia- and tarsus and the right metatarsomeres 3–4 are missing.
</materialsCitation>
</paragraph>
<paragraph id="8BD5E0567B3FFFF2FF4C8DD883D4F86C" blockId="1.[151,1437,1843,2014]" pageId="1" pageNumber="286">
<materialsCitation id="3B02EA0B7B3FFFF2FF4C8DD883D1F86C" collectingDate="2008-08-08" collectionCode="BMNH" collectorName="M. V. L. Barclay &amp; H. Mendel" country="Taiwan" elevation="1268" latitude="24.024683" location="Lushan" longLatPrecision="1" longitude="121.18127" pageId="1" pageNumber="286" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="paratype">
<typeStatus id="54D15EF47B3FFFF2FF4C8DD88111F80B" box="[199,297,1952,1978]" pageId="1" pageNumber="286" type="paratype">Paratype</typeStatus>
<specimenCount id="9D6C2BDF7B3FFFF2FEBB8DE7816BF808" box="[304,339,1951,1977]" count="1" pageId="1" pageNumber="286" type="female">1♀</specimenCount>
<collectionCode id="ED7B78937B3FFFF2FED18DD8818AF80B" box="[346,434,1952,1978]" pageId="1" pageNumber="286">BMNH</collectionCode>
:
<collectingCountry id="F37DA0C67B3FFFF2FE378DE78213F808" box="[444,555,1951,1977]" name="Taiwan" pageId="1" pageNumber="286">TAIWAN</collectingCountry>
,
<collectingRegion id="49AE2EB47B3FFFF2FDBC8DD882DAF808" box="[567,738,1951,1978]" country="Taiwan" name="Nantou" pageId="1" pageNumber="286">Nantou County</collectingRegion>
<quantity id="4C924DB37B3FFFF2FD628DE78308F80B" box="[745,816,1951,1978]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="286" unit="km" value="0.5">0.5km</quantity>
NW of
<location id="8EB5B68D7B3FFFF2FC008DE783E5F808" LSID="urn:lsid:plazi:treatment:03C351407B3FFFF7FF1C8CBF8197FAE4:8EB5B68D7B3FFFF2FC008DE783E5F808" box="[907,989,1951,1977]" country="Taiwan" latitude="24.024683" longLatPrecision="1" longitude="121.18127" name="Lushan" pageId="1" pageNumber="286" stateProvince="Nantou">Lushan</location>
<geoCoordinate id="EE5E86917B3FFFF2FC6F8DE78455F808" box="[996,1133,1951,1977]" degrees="24" direction="north" minutes="01.481" orientation="latitude" pageId="1" pageNumber="286" precision="1" value="24.024683">N24°01.481</geoCoordinate>
<geoCoordinate id="EE5E86917B3FFFF2FBFF8DE7853FF808" box="[1140,1287,1951,1977]" degrees="121" direction="east" minutes="10.876" orientation="longitude" pageId="1" pageNumber="286" precision="1" value="121.18127">E121°10.876</geoCoordinate>
<date id="FFD4C6967B3FFFF2FA858DE785A0F808" box="[1294,1432,1951,1977]" pageId="1" pageNumber="286" value="2008-08-08">
<collectingDate id="EF903F7E7B3FFFF2FA858DE785A0F808" box="[1294,1432,1951,1977]" pageId="1" pageNumber="286" value="2008-08-08">08.viii.2008</collectingDate>
</date>
,
<quantity id="4C924DB37B3FFFF2FF1C8DBB80D0F86C" box="[151,232,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="1" pageNumber="286" unit="m" value="1268.0">
<elevation id="004707657B3FFFF2FF1C8DBB80D0F86C" box="[151,232,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="1" pageNumber="286" unit="m" value="1268.0">1268m</elevation>
</quantity>
, at light,
<collectorName id="269F85807B3FFFF2FEDB8DBC81C5F86C" box="[336,509,1987,2014]" pageId="1" pageNumber="286">M.V.L. Barclay</collectorName>
&amp;
<collectorName id="269F85807B3FFFF2FDAA8DBC82A1F86C" box="[545,665,1987,2014]" pageId="1" pageNumber="286">H. Mendel</collectorName>
<collectionCode id="ED7B78937B3FFFF2FD2B8DBB82CFF86C" box="[672,759,1987,2013]" pageId="1" pageNumber="286">BMNH</collectionCode>
(E) 2008-85 [printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF15B0DE7B3CFFF1FF1C8F5080E1FAD5" ID-DOI="http://doi.org/10.5281/zenodo.13757581" ID-Zenodo-Dep="13757581" httpUri="https://zenodo.org/record/13757581/files/figure.png" pageId="2" pageNumber="287" startId="2.[151,250,1320,1344]" targetBox="[181,1395,190,1232]" targetPageId="2" targetType="figure">
<paragraph id="8BD5E0567B3CFFF1FF1C8F5080E1FAD5" blockId="2.[151,1437,1320,1380]" pageId="2" pageNumber="287">
<emphasis id="B91E3C447B3CFFF1FF1C8F50812EFAF1" bold="true" box="[151,278,1320,1344]" pageId="2" pageNumber="287">FIGURE 1.</emphasis>
<taxonomicName id="4C6A9BD57B3CFFF1FEAB8F5181E8FAF1" authorityName="Telnov" authorityYear="2024" box="[288,464,1320,1344]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="mediator" status="sp. nov.">
<emphasis id="B91E3C447B3CFFF1FEAB8F5181E8FAF1" box="[288,464,1320,1344]" italics="true" pageId="2" pageNumber="287">Nitorus mediator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B3CFFF1FE518F508214FA8E" box="[474,556,1320,1343]" pageId="2" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
A—Paratype ♀, habitus, dorsal view; B—Holotype ♂, habitus, lateral view [not to scale].
</paragraph>
</caption>
<subSubSection id="C370B3DD7B3CFFF1FF4C8FCE8420FA61" box="[199,1048,1462,1488]" pageId="2" pageNumber="287" type="etymology">
<paragraph id="8BD5E0567B3CFFF1FF4C8FCE8420FA61" blockId="2.[151,1437,1462,2029]" box="[199,1048,1462,1488]" pageId="2" pageNumber="287">
<emphasis id="B91E3C447B3CFFF1FF4C8FCE819CFA61" bold="true" box="[199,420,1462,1488]" pageId="2" pageNumber="287">Derivatio nominis.</emphasis>
Named from Latin ‘mediātor’ (intermediary, mediator).
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B3CFFF7FF4C8FA283FCFB84" lastPageId="4" lastPageNumber="289" pageId="2" pageNumber="287" type="description">
<paragraph id="8BD5E0567B3CFFF1FF4C8FA2850EF98D" blockId="2.[151,1437,1462,2029]" pageId="2" pageNumber="287">
<emphasis id="B91E3C447B3CFFF1FF4C8FA2814EFA45" bold="true" box="[199,374,1498,1524]" pageId="2" pageNumber="287">Measurements</emphasis>
,
<typeStatus id="54D15EF47B3CFFF1FE0E8FA281DFFA45" box="[389,487,1498,1524]" pageId="2" pageNumber="287" type="holotype">holotype</typeStatus>
male, total body length
<quantity id="4C924DB37B3CFFF1FD768FA2836FFA44" box="[765,855,1498,1525]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="287" unit="mm" value="2.8">2.8 mm</quantity>
; head length
<quantity id="4C924DB37B3CFFF1FC658FA2847FFA44" box="[1006,1095,1498,1525]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="287" unit="mm" value="0.6">0.6 mm</quantity>
, head width across compound eyes
<quantity id="4C924DB37B3CFFF1FF5B8F868110F9A8" box="[208,296,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="287" unit="mm" value="0.6">0.6 mm</quantity>
, pronotal length
<quantity id="4C924DB37B3CFFF1FE6D8F868206F9A8" box="[486,574,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="287" unit="mm" value="0.6">0.6 mm</quantity>
, maximum pronotal width
<quantity id="4C924DB37B3CFFF1FCE68F8683FEF9A8" box="[877,966,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="287" unit="mm" value="0.4">0.4 mm</quantity>
, minimum pronotal width
<quantity id="4C924DB37B3CFFF1FB7A8F868571F9A8" box="[1265,1353,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="287" unit="mm" value="0.3">0.3 mm</quantity>
, elytral length
<quantity id="4C924DB37B3CFFF1FF6F8C5A8104F98C" box="[228,316,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="287" unit="mm" value="1.6">1.6 mm</quantity>
, combined maximum elytral width across midlength
<quantity id="4C924DB37B3CFFF1FC1A8C5A83D2F98C" box="[913,1002,1570,1597]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="287" unit="mm" value="0.9">0.9 mm</quantity>
. Female
<typeStatus id="54D15EF47B3CFFF1FBC58C5B8496F98C" box="[1102,1198,1571,1597]" pageId="2" pageNumber="287" type="paratype">paratype</typeStatus>
<quantity id="4C924DB37B3CFFF1FB3E8C5A84CEF98C" box="[1205,1270,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="287" unit="mm" value="3.0">3 mm</quantity>
long.
</paragraph>
<paragraph id="8BD5E0567B3CFFF7FF4C8C3E81DEFC78" blockId="2.[151,1437,1462,2029]" lastBlockId="4.[151,1437,151,1365]" lastPageId="4" lastPageNumber="289" pageId="2" pageNumber="287">
<emphasis id="B91E3C447B3CFFF1FF4C8C3E8161F9D1" bold="true" box="[199,345,1606,1632]" pageId="2" pageNumber="287">Description.</emphasis>
Head black-brown, pronotum reddish brown. Elytra black-brown, base narrowly reddish brown, postbasal transverse impression with a yellowish rufous transverse band. Antenna yellowish rufous, two terminal antennomeres brown. Maxillary palpus yellowish rufous. Legs brown, basal area of each tibia and trochanters yellowish. Head elliptical, about as long as wide, slightly convex in dorsal aspect, glossy. Labrum subtruncate at anterior margin. Frontoclypeal suture barely indicated, slightly arched. Compound eye moderate, strongly protruding from lateral outline of head, about as long as converging tempus. Head base narrowly rounded. Head dorsal punctures moderate, deep, rather dense between compound eyes, sparse beyond. Intervening spaces smooth and glossy, as wide as (on frons) to 3× as wide as punctures (on vertex). Dorsal cranial setae yellowish, moderately dense, appressed. Antenna extending towards postbasal transverse impression of elytra when directed posteriad, not or hardly thickened in apical half. Antennomere three about 1.4× as long as antennomere two. Antennomeres 4 and 5 about same length, each about 1.1–1.2× as long as antennomere three. Antennomeres 8–9 slightly, penultimate antennomere distinctly widened distally. Terminal antennomere elongate, pointed, about 1.7× as long as penultimate antennomere, distinctly shorter the combined length of antennomeres 9–10. Penultimate maxillary palpomere short, slightly transverse, lobate and long setose at mesal margin. Terminal maxillary palpomere strongly securiform. Cranial ‘neck’ less than one third head width across compound eyes, densely punctured, moderately glossy. Pronotum longer than wide, distinctly narrower than head across eyes, subtruncate medially at anterior margin. Pronotal disc glossy, bigibbose in dorsal aspect with deep, saddle-like discal impression separating anterior and basal lobe of pronotum. Anterior gibbosity smaller and lower, located medially on anterior lobe. Lateral margins of anterior lobe broadly rounded, of basal lobe—subparallel. Anterior pronotal rim distinct, wide dorsally, laterally and ventrally. Antebasal sulcus wide dorsally and laterally. Pronotal punctures smaller and sparser than those on head, intervening spaces glossy and glabrous, generally larger than punctures. Basal lobe with larger and denser (compared to anterior lobe) but shallow punctures, intervening spaces here glossy and glabrous, about as wide as punctures. Lateral sides of anterior lobe glossy and glabrous, nearly impunctured, of basal lobe—glossy and glabrous, with several irregular subparallel longitudinal wrinkles. Pronotal setae as those on head, subdecumbent. Scutellar shield lanceolate, apically rounded, glossy and glabrous, dorsally slightly convex. Elytra elongate, about 1.7–1.8× as long as wide, widest at midlength, dorsally slightly convex in apical two thirds. Postbasal transverse impression shallow but distinct. Humerus broadly rounded. Apical sutural angle rounded. Elytral surface glossy and smooth, dorsal punctures moderate, shallow, moderately dense. Intervening spaces smooth and glossy, about as wide as to wider than punctures. A yellowish, short, slightly curved, nearly appressed seta raises from each puncture. Scattered, longer erect tactile setae on elytral disc. Metathoracic wing fully developed (functional). Legs long and slender, moderately densely, appressedly setose. Tibial terminal spurs appear single, short. Penultimate tarsomeres slightly bilobate. Tergite VII truncate at posterior margin (
<figureCitation id="1351FCD37B3AFFF7FC26893B8431FCEC" box="[941,1033,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1836,1860]" captionTargetBox="[163,1424,210,1800]" captionTargetId="figure-18@3.[151,1436,181,1812]" captionTargetPageId="3" captionText="FIGURE 2. Nitorus mediator sp. nov., holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="4" pageNumber="289">Fig. 2A</figureCitation>
). Morphological sternite VII broad, rounded at posterior margin (
<figureCitation id="1351FCD37B3AFFF7FE6C891F8279FC30" box="[487,577,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1836,1860]" captionTargetBox="[163,1424,210,1800]" captionTargetId="figure-18@3.[151,1436,181,1812]" captionTargetPageId="3" captionText="FIGURE 2. Nitorus mediator sp. nov., holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="4" pageNumber="289">Fig. 2B</figureCitation>
). Morphological tergite VIII rounded, long setose at posterior margin (
<figureCitation id="1351FCD37B3AFFF7FAE4891F8084FC14" captionStart="FIGURE 2" captionStartId="3.[151,250,1836,1860]" captionTargetBox="[163,1424,210,1800]" captionTargetId="figure-18@3.[151,1436,181,1812]" captionTargetPageId="3" captionText="FIGURE 2. Nitorus mediator sp. nov., holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="4" pageNumber="289">Fig. 2C</figureCitation>
). Morphological sternite VIII horseshoe-shaped (
<figureCitation id="1351FCD37B3AFFF7FD6E89F38307FC14" box="[741,831,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1836,1860]" captionTargetBox="[163,1424,210,1800]" captionTargetId="figure-18@3.[151,1436,181,1812]" captionTargetPageId="3" captionText="FIGURE 2. Nitorus mediator sp. nov., holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="4" pageNumber="289">Fig. 2D</figureCitation>
), sternite IX Y-shaped, arms short, stem sinuous (
<figureCitation id="1351FCD37B3AFFF7FAE489F38082FC78" captionStart="FIGURE 2" captionStartId="3.[151,250,1836,1860]" captionTargetBox="[163,1424,210,1800]" captionTargetId="figure-18@3.[151,1436,181,1812]" captionTargetPageId="3" captionText="FIGURE 2. Nitorus mediator sp. nov., holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="4" pageNumber="289">Fig. 2E</figureCitation>
). Aedeagus as in fig. 2F‒I.
</paragraph>
<caption id="DF15B0DE7B3DFFF0FF1C8D5485B2F83D" ID-DOI="http://doi.org/10.5281/zenodo.13757585" ID-Zenodo-Dep="13757585" httpUri="https://zenodo.org/record/13757585/files/figure.png" pageId="3" pageNumber="288" startId="3.[151,250,1836,1860]" targetBox="[163,1424,210,1800]" targetPageId="3" targetType="figure">
<paragraph id="8BD5E0567B3DFFF0FF1C8D5485B2F83D" blockId="3.[151,1436,1836,1933]" pageId="3" pageNumber="288">
<emphasis id="B91E3C447B3DFFF0FF1C8D54812BF8F5" bold="true" box="[151,275,1836,1860]" pageId="3" pageNumber="288">FIGURE 2.</emphasis>
<taxonomicName id="4C6A9BD57B3DFFF0FE928D5581FFF8F5" authorityName="Telnov" authorityYear="2024" box="[281,455,1836,1860]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="mediator" status="sp. nov.">
<emphasis id="B91E3C447B3DFFF0FE928D5581FFF8F5" box="[281,455,1836,1860]" italics="true" pageId="3" pageNumber="288">Nitorus mediator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D813F7B3DFFF0FE468D558226F8F5" box="[461,542,1837,1860]" pageId="3" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂. A—Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Tergite VIII, dorsal view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX, ventral view; F— Aedeagus; G—ditto, apex, magnified; H—ditto, median portion; I—ditto, internal sac with gonopore armature [not to scale].
</paragraph>
</caption>
<paragraph id="8BD5E0567B3AFFF7FF4C89AA83FCFB84" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="289">
<emphasis id="B91E3C447B3AFFF7FF4C89AA8180FC5D" bold="true" box="[199,440,978,1004]" pageId="4" pageNumber="289">Sexual dimorphism.</emphasis>
Female externally similar to male, but antenna comparatively shorter, anterior gibbosity of pronotum in lateral view comparatively stronger developed than that of male. Tergite VII truncate at posterior margin. Morphological sternite VII broadly rounded at posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B3AFFF7FF4C8E468323FB58" pageId="4" pageNumber="289" type="diagnosis">
<paragraph id="8BD5E0567B3AFFF7FF4C8E468323FB58" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="289">
<emphasis id="B91E3C447B3AFFF7FF4C8E4681E9FBE9" bold="true" box="[199,465,1086,1112]" pageId="4" pageNumber="289">Differential diagnosis.</emphasis>
This species differs from
<taxonomicName id="4C6A9BD57B3AFFF7FC8E8E478452FBE8" authority="(Pic, 1913)" baseAuthorityName="Pic" baseAuthorityYear="1913" box="[773,1130,1087,1113]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="mediomaculatus">
<emphasis id="B91E3C447B3AFFF7FC8E8E4783D9FBE8" box="[773,993,1087,1113]" italics="true" pageId="4" pageNumber="289">N. mediomaculatus</emphasis>
(
<bibRefCitation id="EFFB9DA77B3AFFF7FC798E47845AFBE8" author="Pic, M." box="[1010,1122,1087,1113]" pageId="4" pageNumber="289" pagination="129 - 135" refId="ref13523" refString="Pic, M. (1913) H. Sauter's Formosa-Ausbeute. Anthicidae, Pedilidae und Hylophilidae. Archiv fur Naturgeschichte, Series A, 79, 129 - 135." type="journal article" year="1913">Pic, 1913</bibRefCitation>
)
</taxonomicName>
(see the new combination and rescued name below),
<taxonomicName id="4C6A9BD57B3AFFF7FE5B8E1B8321FBCC" authority="(Marseul, 1876)" baseAuthorityName="Marseul" baseAuthorityYear="1876" box="[464,793,1123,1149]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="trigibber">
<emphasis id="B91E3C447B3AFFF7FE5B8E1B8260FBCC" box="[464,600,1123,1149]" italics="true" pageId="4" pageNumber="289">N. trigibber</emphasis>
(Marseul, 1876)
</taxonomicName>
and other congeners with strongly bigibbose pronotum in the shape of the aedeagus (also see the Key below). It appears different from
<taxonomicName id="4C6A9BD57B3AFFF7FBA38EFF8496FB10" baseAuthorityName="Marseul" baseAuthorityYear="1876" box="[1064,1198,1159,1185]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="trigibber">
<emphasis id="B91E3C447B3AFFF7FBA38EFF8496FB10" box="[1064,1198,1159,1185]" italics="true" pageId="4" pageNumber="289">N. trigibber</emphasis>
</taxonomicName>
also in the distinctly longer antenna (extending towards postbasal transverse impression area of elytra
<emphasis id="B91E3C447B3AFFF7FBA68ED3844DFB75" box="[1069,1141,1195,1220]" italics="true" pageId="4" pageNumber="289">versus</emphasis>
towards elytral base in
<taxonomicName id="4C6A9BD57B3AFFF7FA088ED380C3FB58" baseAuthorityName="Marseul" baseAuthorityYear="1876" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="trigibber">
<emphasis id="B91E3C447B3AFFF7FA088ED380C3FB58" italics="true" pageId="4" pageNumber="289">N. trigibber</emphasis>
</taxonomicName>
) and the comparatively stronger elongate elytra.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B3AFFF7FF4C8E8B81C1FABC" box="[199,505,1267,1293]" pageId="4" pageNumber="289" type="biology_ecology">
<paragraph id="8BD5E0567B3AFFF7FF4C8E8B81C1FABC" blockId="4.[151,1437,151,1365]" box="[199,505,1267,1293]" pageId="4" pageNumber="289">
<emphasis id="B91E3C447B3AFFF7FF4C8E8B8114FABC" bold="true" box="[199,300,1267,1293]" pageId="4" pageNumber="289">Ecology.</emphasis>
Attracted to light.
</paragraph>
</subSubSection>
<subSubSection id="C370B3DD7B3AFFF7FF4C8F6E8197FAE4" pageId="4" pageNumber="289" type="distribution">
<paragraph id="8BD5E0567B3AFFF7FF4C8F6E8197FAE4" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="289">
<emphasis id="B91E3C447B3AFFF7FF4C8F6E8159FA81" bold="true" box="[199,353,1302,1328]" pageId="4" pageNumber="289">Distribution.</emphasis>
So far known from mountainous central (
<collectingRegion id="49AE2EB47B3AFFF7FCB48F6F83C8FA80" box="[831,1008,1303,1329]" country="Taiwan" name="Nantou" pageId="4" pageNumber="289">Nantou County</collectingRegion>
) and southern (the former
<collectingRegion id="49AE2EB47B3AFFF7FAA88F6F85A4FA80" box="[1315,1436,1303,1329]" country="Taiwan" name="Kaohsiung" pageId="4" pageNumber="289">Kaohsiung</collectingRegion>
County) parts of
<collectingCountry id="F37DA0C67B3AFFF7FEDE8F438193FAE4" box="[341,427,1339,1365]" name="Taiwan" pageId="4" pageNumber="289">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

293
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<mods:title id="2AE891347053CDFA0EB249BC71BBCDBC">First record of subgenus Corynopus Lepeletier and Brullé, 1835 (Genus Rhopalum Stephens, Hymenoptera: Crabronidae: Crabronini) from India: a new species and a key to Asian species of the subgenus</mods:title>
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<paragraph id="8BDF36E9A512FFD5FF738939FC815517" blockId="2.[151,852,1358,1421]" box="[151,852,1358,1385]" pageId="2" pageNumber="469">
<heading id="D0978185A512FFD5FF738939FC815517" bold="true" box="[151,852,1358,1385]" fontSize="11" level="1" pageId="2" pageNumber="469" reason="1">
<emphasis id="B914EAFBA512FFD5FF738939FC815517" bold="true" box="[151,852,1358,1385]" pageId="2" pageNumber="469">
<taxonomicName id="4C604D6AA512FFD5FF738939FD3C5517" authority="Saini &amp; Dey" authorityName="Saini &amp; Dey" authorityYear="2024" box="[151,745,1358,1385]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="chettalicum" status="sp. nov." subGenus="Corynopus">
<emphasis id="B914EAFBA512FFD5FF738939FD9E5517" bold="true" box="[151,587,1358,1385]" italics="true" pageId="2" pageNumber="469">Rhopalum (Corynopus) chettalicum</emphasis>
Saini &amp; Dey
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,
<taxonomicNameLabel id="A2275780A512FFD5FD128938FC815517" box="[758,852,1359,1385]" pageId="2" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C37A6562A512FFD5FF738905FECC55F3" box="[151,281,1394,1421]" pageId="2" pageNumber="469" type="description">
<paragraph id="8BDF36E9A512FFD5FF738905FECC55F3" blockId="2.[151,852,1358,1421]" box="[151,281,1394,1421]" pageId="2" pageNumber="469">
(
<figureCitation id="135B2A6CA512FFD5FF7B8905FEC555F3" box="[159,272,1394,1421]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–8" captionStart-2="FIGURES 9–11" captionStart-3="FIGURES 12–15" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,1943,1967]" captionStartId-3="6.[151,264,1534,1558]" captionTargetBox-0="[210,1377,191,1905]" captionTargetBox-1="[220,1368,192,1909]" captionTargetBox-2="[182,1406,193,1907]" captionTargetBox-3="[163,1410,193,1499]" captionTargetId-0="figure-17@3.[198,1389,179,1917]" captionTargetId-1="figure-21@4.[209,1379,181,1920]" captionTargetId-2="figure-17@5.[170,1418,181,1919]" captionTargetId-3="figure-21@6.[151,1436,181,1511]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1–2. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 1. Habitus, lateral view; 2. Habitus, ventral view." captionText-1="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." captionText-2="FIGURES 9–11. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 9. Mesosoma, dorsal view; 10. Mesosoma, lateral view; 11. Propodeum." captionText-3="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13757937" figureDoi-1="http://doi.org/10.5281/zenodo.13757939" figureDoi-2="http://doi.org/10.5281/zenodo.13757945" figureDoi-3="http://doi.org/10.5281/zenodo.13757947" httpUri-0="https://zenodo.org/record/13757937/files/figure.png" httpUri-1="https://zenodo.org/record/13757939/files/figure.png" httpUri-2="https://zenodo.org/record/13757945/files/figure.png" httpUri-3="https://zenodo.org/record/13757947/files/figure.png" pageId="2" pageNumber="469">Figs 1–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A6562A512FFD5FF7389CDFE7356D2" pageId="2" pageNumber="469" type="diagnosis">
<paragraph id="8BDF36E9A512FFD5FF7389CDFE7356D2" blockId="2.[151,1437,1466,1925]" pageId="2" pageNumber="469">
<emphasis id="B914EAFBA512FFD5FF7389CDFEC755AA" bold="true" box="[151,274,1466,1492]" pageId="2" pageNumber="469">Diagnosis.</emphasis>
The new species is close to
<taxonomicName id="4C604D6AA512FFD5FDAD89CCFCBF55AB" authority="Wesmael, 1852" authorityName="Wesmael" authorityYear="1852" box="[585,874,1466,1493]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B914EAFBA512FFD5FDAD89CCFD6355AA" box="[585,694,1466,1492]" italics="true" pageId="2" pageNumber="469">R. gracile</emphasis>
<bibRefCitation id="EFF14B18A512FFD5FD5889CDFCBF55AB" author="Wesmael, C." box="[700,874,1466,1493]" pageId="2" pageNumber="469" pagination="589 - 635" refId="ref6461" refString="Wesmael, C. (1852) Revue critique des Hymenopteres Fouisseurs de Belgique. Suite. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 19, 589 - 635. [in French] https: // doi. org / 10.5962 / bhl. title. 67728" type="journal article" year="1852">Wesmael, 1852</bibRefCitation>
</taxonomicName>
, sharing with it the following characters: mandible ferruginous; posterior lateral corner of propleuron without small tooth; hind tibiae black with yellow ring basally and markedly clavate &amp; spinose; posterior surface of propodeum carinate; tergum I longer than hind femur; pygidium triangular and surrounded by carina. The new species differs from
<taxonomicName id="4C604D6AA512FFD5FC788A50FBD8563E" authorityName="Wesmael" authorityYear="1852" box="[924,1037,1574,1600]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B914EAFBA512FFD5FC788A50FBD8563E" box="[924,1037,1574,1600]" italics="true" pageId="2" pageNumber="469">R. gracile</emphasis>
</taxonomicName>
and other species of the subgenus
<taxonomicName id="4C604D6AA512FFD5FF738A3CFEC7561B" authorityName="Lepeletier de Saint Fargeau and Brulle" authorityYear="1835" box="[151,274,1611,1637]" pageId="2" pageNumber="469" rank="subGenus" subGenus="Corynopus">
<emphasis id="B914EAFBA512FFD5FF738A3CFEC7561B" box="[151,274,1611,1637]" italics="true" pageId="2" pageNumber="469">Corynopus</emphasis>
</taxonomicName>
by a slight clypeal emargination of medial lobe, orbital foveae distinct, fine longitudinal furrow between posterior ocelli present, pygidial plate with prominent medio-basal hump, coloration and punctuation on body and legs as described below.
</paragraph>
</subSubSection>
<subSubSection id="C37A6562A512FFD0FF238AC1FE4A544B" lastPageId="7" lastPageNumber="474" pageId="2" pageNumber="469" type="description">
<paragraph id="8BDF36E9A512FFD5FF238AC1FCB9571F" blockId="2.[151,1437,1466,1925]" pageId="2" pageNumber="469">
<emphasis id="B914EAFBA512FFD5FF238AC1FE7056AF" bold="true" box="[199,421,1718,1745]" pageId="2" pageNumber="469">
Description.
<emphasis id="B914EAFBA512FFD5FE848AC1FE7056AF" bold="true" box="[352,421,1718,1745]" italics="true" pageId="2" pageNumber="469">Head.</emphasis>
</emphasis>
Densely micro-sculptured dorsally but impunctate; clypeus medial emargination shallow; orbital fovea distinct, elliptical, basally joining compound eye; fine longitudinal line between posterior ocelli present (
<figureCitation id="135B2A6CA512FFD5FF138A89FEE95767" box="[247,316,1790,1817]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="2" pageNumber="469">Fig. 4</figureCitation>
).
<emphasis id="B914EAFBA512FFD5FEB48A88FE075767" bold="true" box="[336,466,1791,1817]" pageId="2" pageNumber="469">
<taxonomicName id="4C604D6AA512FFD5FEB48A88FE1E5767" box="[336,459,1791,1817]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAFBA512FFD5FEB48A88FE1E5767" bold="true" box="[336,459,1791,1817]" italics="true" pageId="2" pageNumber="469">Mesosoma</emphasis>
</taxonomicName>
.
</emphasis>
Scutum and scutellum punctate; mesopleuron and lateral surface of propodeum micro-sculptured dorsally (
<figureCitation id="135B2A6CA512FFD5FE9B8B55FE065743" box="[383,467,1826,1853]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@5.[170,1418,181,1919]" captionTargetPageId="5" captionText="FIGURES 9–11. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 9. Mesosoma, dorsal view; 10. Mesosoma, lateral view; 11. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.13757945" httpUri="https://zenodo.org/record/13757945/files/figure.png" pageId="2" pageNumber="469">Fig. 10</figureCitation>
).
<emphasis id="B914EAFBA512FFD5FE0D8B54FDB25743" bold="true" box="[489,615,1827,1853]" italics="true" pageId="2" pageNumber="469">Metasoma.</emphasis>
Gt
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longer than hind femur, straight, bend basally, nodose apically (
<figureCitation id="135B2A6CA512FFD5FA8B8B55FF63571F" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1534,1558]" captionTargetBox="[163,1410,193,1499]" captionTargetId="figure-21@6.[151,1436,181,1511]" captionTargetPageId="6" captionText="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi="http://doi.org/10.5281/zenodo.13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="2" pageNumber="469">Fig. 12</figureCitation>
); pygidial plate with distinct medio-basal hump (
<figureCitation id="135B2A6CA512FFD5FD388B31FC8A571F" box="[732,863,1862,1889]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1534,1558]" captionTargetBox="[163,1410,193,1499]" captionTargetId="figure-21@6.[151,1436,181,1511]" captionTargetPageId="6" captionText="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi="http://doi.org/10.5281/zenodo.13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="2" pageNumber="469">Figs 14–15</figureCitation>
).
</paragraph>
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<emphasis id="B914EAFBA512FFD5FF238B1DFE9357FA" bold="true" box="[199,326,1898,1924]" pageId="2" pageNumber="469">FEMALE.</emphasis>
<typeStatus id="54DB884BA512FFD5FEA98B1DFE6057FA" box="[333,437,1898,1924]" pageId="2" pageNumber="469" type="holotype">Holotype</typeStatus>
<specimenCount id="9D66FD60A512FFD5FE588B1CFE0757FB" box="[444,466,1899,1925]" count="1" pageId="2" pageNumber="469" type="female">♀</specimenCount>
. BL=
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; FWL=
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<emphasis id="B914EAFBA513FFD4FF738BE0FEEF57D1" bold="true" box="[151,314,1943,1967]" pageId="3" pageNumber="470">FIGURES 1–2.</emphasis>
<taxonomicName id="4C604D6AA513FFD4FEA58BE0FD7957D1" authorityName="Saini &amp; Dey" authorityYear="2024" box="[321,684,1943,1967]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="470" phylum="Arthropoda" rank="species" species="chettalicum" status="sp. nov." subGenus="Corynopus">
<emphasis id="B914EAFBA513FFD4FEA58BE0FD7957D1" box="[321,684,1943,1967]" italics="true" pageId="3" pageNumber="470">Rhopalum (Corynopus) chettalicum</emphasis>
</taxonomicName>
<emphasis id="B914EAFBA513FFD4FD568BE0FCD657D1" bold="true" box="[690,771,1943,1967]" pageId="3" pageNumber="470">
<taxonomicNameLabel id="A2275780A513FFD4FD568BE0FCD657D1" box="[690,771,1943,1967]" pageId="3" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype ♀. 1. Habitus, lateral view; 2. Habitus, ventral view.
</paragraph>
</caption>
<caption id="DF1F6661A514FFD3FF738BE0FD3A57AA" ID-DOI="http://doi.org/10.5281/zenodo.13757939" ID-Zenodo-Dep="13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="4" pageNumber="471" startId="4.[151,264,1943,1967]" targetBox="[220,1368,192,1909]" targetPageId="4" targetType="figure">
<paragraph id="8BDF36E9A514FFD3FF738BE0FD3A57AA" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="471">
<emphasis id="B914EAFBA514FFD3FF738BE0FEEF57D1" bold="true" box="[151,314,1943,1967]" pageId="4" pageNumber="471">FIGURES 3–8.</emphasis>
<taxonomicName id="4C604D6AA514FFD3FEA48BEFFD7F57CE" authorityName="Saini &amp; Dey" authorityYear="2024" box="[320,682,1944,1968]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="471" phylum="Arthropoda" rank="species" species="chettalicum" status="sp. nov." subGenus="Corynopus">
<emphasis id="B914EAFBA514FFD3FEA48BEFFD7F57CE" box="[320,682,1944,1968]" italics="true" pageId="4" pageNumber="471">Rhopalum (Corynopus) chettalicum</emphasis>
</taxonomicName>
<emphasis id="B914EAFBA514FFD3FD548BEFFCD557CE" bold="true" box="[688,768,1944,1968]" pageId="4" pageNumber="471">
<taxonomicNameLabel id="A2275780A514FFD3FD548BEFFCD557CE" box="[688,768,1944,1968]" pageId="4" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view.
</paragraph>
</caption>
<caption id="DF1F6661A515FFD2FF738BE0FEA257AD" ID-DOI="http://doi.org/10.5281/zenodo.13757945" ID-Zenodo-Dep="13757945" httpUri="https://zenodo.org/record/13757945/files/figure.png" pageId="5" pageNumber="472" startId="5.[151,264,1943,1967]" targetBox="[182,1406,193,1907]" targetPageId="5" targetType="figure">
<paragraph id="8BDF36E9A515FFD2FF738BE0FEA257AD" blockId="5.[151,1436,1943,2003]" pageId="5" pageNumber="472">
<emphasis id="B914EAFBA515FFD2FF738BE0FE9257D1" bold="true" box="[151,327,1943,1967]" pageId="5" pageNumber="472">FIGURES 9–11.</emphasis>
<taxonomicName id="4C604D6AA515FFD2FEAA8BE0FD6F57D1" authorityName="Saini &amp; Dey" authorityYear="2024" box="[334,698,1943,1967]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="472" phylum="Arthropoda" rank="species" species="chettalicum" status="sp. nov." subGenus="Corynopus">
<emphasis id="B914EAFBA515FFD2FEAA8BE0FD6F57D1" box="[334,698,1943,1967]" italics="true" pageId="5" pageNumber="472">Rhopalum (Corynopus) chettalicum</emphasis>
</taxonomicName>
<emphasis id="B914EAFBA515FFD2FD268BE0FCC657D1" bold="true" box="[706,787,1943,1967]" pageId="5" pageNumber="472">
<taxonomicNameLabel id="A2275780A515FFD2FD268BE0FCC657D1" box="[706,787,1943,1967]" pageId="5" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype ♀. 9.
<taxonomicName id="4C604D6AA515FFD2FC5F8BEFFBFE57D1" box="[955,1067,1944,1967]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="474" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, dorsal view; 10.
<taxonomicName id="4C604D6AA515FFD2FB068BEFFA8757D1" box="[1250,1362,1944,1967]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="474" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, lateral view; 11. Propodeum.
</paragraph>
</caption>
<caption id="DF1F6661A516FFD1FF738989FDDE5645" ID-DOI="http://doi.org/10.5281/zenodo.13757947" ID-Zenodo-Dep="13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="6" pageNumber="473" startId="6.[151,264,1534,1558]" targetBox="[163,1410,193,1499]" targetPageId="6" targetType="figure">
<paragraph id="8BDF36E9A516FFD1FF738989FDDE5645" blockId="6.[151,1437,1534,1595]" pageId="6" pageNumber="473">
<emphasis id="B914EAFBA516FFD1FF738989FE815668" bold="true" box="[151,340,1534,1558]" pageId="6" pageNumber="473">FIGURES 12–15.</emphasis>
<taxonomicName id="4C604D6AA516FFD1FEBE8988FD105669" authorityName="Saini &amp; Dey" authorityYear="2024" box="[346,709,1535,1559]" class="Insecta" family="Crabronidae" genus="Rhopalum" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="473" phylum="Arthropoda" rank="species" species="chettalicum" status="sp. nov." subGenus="Corynopus">
<emphasis id="B914EAFBA516FFD1FEBE8988FD105669" box="[346,709,1535,1559]" italics="true" pageId="6" pageNumber="473">Rhopalum (Corynopus) chettalicum</emphasis>
</taxonomicName>
<emphasis id="B914EAFBA516FFD1FD2F8988FCC95669" bold="true" box="[715,796,1535,1559]" pageId="6" pageNumber="473">
<taxonomicNameLabel id="A2275780A516FFD1FD2F8988FCC95669" box="[715,796,1535,1559]" pageId="6" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view.
</paragraph>
</caption>
<paragraph id="8BDF36E9A516FFD1FF238AFBFBA75795" blockId="6.[151,1437,1676,2027]" pageId="6" pageNumber="473">
<emphasis id="B914EAFBA516FFD1FF238AFBFDE856D9" bold="true" box="[199,573,1676,1703]" pageId="6" pageNumber="473">
<emphasis id="B914EAFBA516FFD1FF238AFBFDE356D9" bold="true" box="[199,566,1676,1703]" italics="true" pageId="6" pageNumber="473">Colour pattern and pubescence</emphasis>
.
</emphasis>
Body black, except the following yellow: scape and pedicel (except dorsal spot), flagellum ventrally, pronotal lobe, maxillary &amp; labial palpi, fore femur with large apical spot, fore tibia except small dorsal spot, fore and mid basitarsi completely, fore and mid tarsomeres except apically, mid tibia except large dorsal spot, hind femur with small apical spot and hind tibia basally (
<figureCitation id="135B2A6CA516FFD1FC588A8EFBF6576D" box="[956,1059,1785,1811]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[210,1377,191,1905]" captionTargetId="figure-17@3.[198,1389,179,1917]" captionTargetPageId="3" captionText="FIGURES 1–2. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 1. Habitus, lateral view; 2. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757937" httpUri="https://zenodo.org/record/13757937/files/figure.png" pageId="6" pageNumber="473">Figs 1–2</figureCitation>
); apical portion of all coxae and pygidial plate apically yellowish-brown; mandible except apically, petiole, basal portion of Gt
<subScript id="17E434ACA516FFD1FB5E8B58FB175743" attach="left" box="[1210,1218,1839,1853]" fontSize="6" pageId="6" pageNumber="473">2</subScript>
to Gt
<subScript id="17E434ACA516FFD1FAE68B58FADF5743" attach="left" box="[1282,1290,1839,1853]" fontSize="6" pageId="6" pageNumber="473">5</subScript>
dorsally and sterna reddish brown; wing veins and flagellum light to dark brown. Body mostly with creamish-white setae; clypeus and scapal basin covered with dense silvery setae; upper frons and vertex with small, sparse, silvery setae; lateral surface of head covered with dense creamish-white setae; scutum covered with fine silvery setae; mesopleuron and lateral side of propodeum covered with dense creamish-white setae; Gt
<subScript id="17E434ACA516FFD1FC5D8BC8FC1457B3" attach="left" box="[953,961,1983,1997]" fontSize="6" pageId="6" pageNumber="473">2</subScript>
to Gt
<subScript id="17E434ACA516FFD1FBE68BC8FBDF57B3" attach="left" box="[1026,1034,1983,1997]" fontSize="6" pageId="6" pageNumber="473">5</subScript>
covered with small creamish-white setae (
<figureCitation id="135B2A6CA516FFD1FF3A8BA6FEB45795" box="[222,353,2001,2027]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1534,1558]" captionTargetBox="[163,1410,193,1499]" captionTargetId="figure-21@6.[151,1436,181,1511]" captionTargetPageId="6" captionText="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi="http://doi.org/10.5281/zenodo.13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="6" pageNumber="473">Figs 12–13</figureCitation>
); pygidial plate laterally covered with elongate sparse brownish setae.
</paragraph>
<paragraph id="8BDF36E9A517FFD0FF238CE1FBF6518B" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF238CE1FED050CF" bold="true" box="[199,261,150,177]" italics="true" pageId="7" pageNumber="474">Head</emphasis>
. Cubical, shiny, distinctly micro-sculptured dorsally, microsculpture dense near ocelli and vertex, length 0.7× of width; ocelli slightly broader than high (
<figureCitation id="135B2A6CA517FFD0FD378CCCFCC950AB" box="[723,796,187,213]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Fig. 4</figureCitation>
); distance between compound eyes and antennal toruli very small, almost nil; inner margins of compound eyes convergent below, with carina; supra-antennal projection inconspicuous, covered with silvery setae; scapal basin shiny and slightly concave, inner orbit of compound eyes near scapal basin with silvery pubescence (
<figureCitation id="135B2A6CA517FFD0FD938D50FD6F513F" box="[631,698,295,321]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Fig. 3</figureCitation>
); indistinct longitudinal furrow extending from anterior ocellus to frontoclypeal groove; face narrow; clypeus broad, slightly swollen, anterior margin emarginate, with blunt projection on each side (
<figureCitation id="135B2A6CA517FFD0FEC88D18FE5A51F7" box="[300,399,367,393]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Figs 3, 6</figureCitation>
); vertex with fine longitudinal furrow starting from anterior ocellus and not reaching occipital carina; hypostomal carina wide U-shaped, separated from occipital carina (
<figureCitation id="135B2A6CA517FFD0FC318DE4FBC251D3" box="[981,1047,403,429]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Fig. 8</figureCitation>
); mandibles bifid at apex; antennal sockets contiguous; POD equal to 0.77
<emphasis id="B914EAFBA517FFD0FDA18DC0FD8051AF" bold="true" box="[581,597,439,465]" pageId="7" pageNumber="474">×</emphasis>
of OOD; relative lengths of antennomeres I: II: III: IV: V: VI: VII: VIII: IX: X: XI and XII = 4.3: 1.1: 0.9: 0.9: 0.9: 0.8: 0.8: 0.9: 0.7: 0.7: 0.8 and 1.4 (
<figureCitation id="135B2A6CA517FFD0FC348DACFBC0518B" box="[976,1045,475,501]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BDF36E9A517FFD0FF238D88FEA65323" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="474">
<taxonomicName id="4C604D6AA517FFD0FF238D88FE975267" box="[199,322,511,537]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAFBA517FFD0FF238D88FE975267" bold="true" box="[199,322,511,537]" italics="true" pageId="7" pageNumber="474">Mesosoma</emphasis>
</taxonomicName>
. Dull compared to head; dorsally covered with fine creamish-white setae; pronotal collar laterally dentate, slightly notched medially; prosternum with slight indentation (
<figureCitation id="135B2A6CA517FFD0FC4A8E54FBC45243" box="[942,1041,546,573]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[220,1368,192,1909]" captionTargetId="figure-21@4.[209,1379,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–8. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 3. Head, frontal view; 4. Head, dorsal view; 5. Head, lateral view; 6. Clypeus; 7. Antenna; 8. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13757939" httpUri="https://zenodo.org/record/13757939/files/figure.png" pageId="7" pageNumber="474">Figs 5, 8</figureCitation>
); scutum densely micro-punctured, with well-defined admedian and parapsidal lines; scutellum broad and metanotum micro-punctate dorsally (
<figureCitation id="135B2A6CA517FFD0FA8B8E30FF7252FB" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@5.[170,1418,181,1919]" captionTargetPageId="5" captionText="FIGURES 9–11. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 9. Mesosoma, dorsal view; 10. Mesosoma, lateral view; 11. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.13757945" httpUri="https://zenodo.org/record/13757945/files/figure.png" pageId="7" pageNumber="474">Fig. 9</figureCitation>
); mesopleuron shiny, micro-sculptured dorsally; epicnemial sulcus with distinct large foveolae, reaching tegula; precoxal area with small foveolae compared to those on epicnemial sulcus; mesopleural suture without fovea; metapleuron shiny, micro-sculptured dorsally (
<figureCitation id="135B2A6CA517FFD0FD488EC4FCD752B3" box="[684,770,691,717]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@5.[170,1418,181,1919]" captionTargetPageId="5" captionText="FIGURES 9–11. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 9. Mesosoma, dorsal view; 10. Mesosoma, lateral view; 11. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.13757945" httpUri="https://zenodo.org/record/13757945/files/figure.png" pageId="7" pageNumber="474">Fig. 10</figureCitation>
); propodeum smooth, shiny, micro-sculptured dorsally, its anterior margin with fine foveolae, longitudinal furrow conspicuous and narrow, posterior surface dorsally micro-sculptured, lateral surface with fine transverse striae (
<figureCitation id="135B2A6CA517FFD0FD198E8CFC85536B" box="[765,848,763,789]" captionStart="FIGURES 9–11" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[182,1406,193,1907]" captionTargetId="figure-17@5.[170,1418,181,1919]" captionTargetPageId="5" captionText="FIGURES 9–11. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 9. Mesosoma, dorsal view; 10. Mesosoma, lateral view; 11. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.13757945" httpUri="https://zenodo.org/record/13757945/files/figure.png" pageId="7" pageNumber="474">Fig. 11</figureCitation>
); fore tibia without dorsal spines, mid tibia simple, with 2-4 spines dorsally, hind tibia largely inflated apically with 8-10 short spines dorsally, ventrally with dense silvery pubescence.
</paragraph>
<paragraph id="8BDF36E9A517FFD0FF238F10FBC653FF" blockId="7.[151,1437,150,1185]" box="[199,1043,871,897]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF238F10FEDA53FF" bold="true" box="[199,271,871,897]" italics="true" pageId="7" pageNumber="474">Wings</emphasis>
. Recurrent vein joining sub marginal cell slightly beyond its middle.
</paragraph>
<paragraph id="8BDF36E9A517FFD0FF238FFCFDC1546F" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF238FFCFE9353DB" bold="true" box="[199,326,907,933]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF238FFCFEEA53DB" bold="true" box="[199,319,907,933]" italics="true" pageId="7" pageNumber="474">Metasoma</emphasis>
.
</emphasis>
Petiolate, petiole longer than combined length of hind trochanter and hind femur, distinctly nodose at apex; Gt
<subScript id="17E434ACA517FFD0FF118FB6FF2853B1" attach="left" box="[245,253,961,975]" fontSize="6" pageId="7" pageNumber="474">2</subScript>
longer than wide, Gt
<subScript id="17E434ACA517FFD0FE018FB6FE3853B1" attach="left" box="[485,493,961,975]" fontSize="6" pageId="7" pageNumber="474">1</subScript>
to Gt
<subScript id="17E434ACA517FFD0FDCE8FB6FDE753B1" attach="left" box="[554,562,961,975]" fontSize="6" pageId="7" pageNumber="474">5</subScript>
covered with short pubescence dorsally (
<figureCitation id="135B2A6CA517FFD0FC1C8FD8FBAF53B7" box="[1016,1146,943,969]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1534,1558]" captionTargetBox="[163,1410,193,1499]" captionTargetId="figure-21@6.[151,1436,181,1511]" captionTargetPageId="6" captionText="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi="http://doi.org/10.5281/zenodo.13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="7" pageNumber="474">Figs 12–13</figureCitation>
); pygidial plate triangular, broader at base, with medio-basal gibbosity, depressed and rounded apically, carinate laterally, ferruginous with thick brown bristles (
<figureCitation id="135B2A6CA517FFD0FE608F80FDD2546F" box="[388,519,1014,1041]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1534,1558]" captionTargetBox="[163,1410,193,1499]" captionTargetId="figure-21@6.[151,1436,181,1511]" captionTargetPageId="6" captionText="FIGURES 12–15. Rhopalum (Corynopus) chettalicum sp. nov. Holotype ♀. 12. Metasoma, lateral view; 13. Tergum I; 14–15. Pygidial plate, dorsal &amp; lateral view." figureDoi="http://doi.org/10.5281/zenodo.13757947" httpUri="https://zenodo.org/record/13757947/files/figure.png" pageId="7" pageNumber="474">Figs 14–15</figureCitation>
).
</paragraph>
<paragraph id="8BDF36E9A517FFD0FF23886DFE4A544B" blockId="7.[151,1437,150,1185]" box="[199,415,1050,1077]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF23886DFEF7544A" bold="true" box="[199,290,1050,1076]" pageId="7" pageNumber="474">MALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37A6562A517FFD0FF238848FC1E5427" box="[199,971,1087,1113]" pageId="7" pageNumber="474" type="etymology">
<paragraph id="8BDF36E9A517FFD0FF238848FC1E5427" blockId="7.[151,1437,150,1185]" box="[199,971,1087,1113]" pageId="7" pageNumber="474">
<emphasis id="B914EAFBA517FFD0FF238848FE9B5427" bold="true" box="[199,334,1087,1113]" pageId="7" pageNumber="474">Etymology.</emphasis>
The species name is based on the
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).
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<emphasis id="B914EAFBA517FFD0FF238815FE645402" bold="true" box="[199,433,1122,1148]" pageId="7" pageNumber="474">Material examined.</emphasis>
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,
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,
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(= Kodagu),
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<collectingDate id="EF9AE9C1A517FFD0FBB38814FB045403" box="[1111,1233,1123,1149]" pageId="7" pageNumber="474" value="2012-06-10">10.vi.2012</collectingDate>
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, coll.
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(NPC-IARI).
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<emphasis id="B914EAFBA518FFDFFF7388DBFDDD54B9" bold="true" box="[151,520,1196,1223]" italics="true" pageId="8" pageNumber="475">Rhopalum coarctatum koreense</emphasis>
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Only known from
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Distribution:
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Q.
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Only known from
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Distribution:
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A. Morawitz, 1866: 267
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<emphasis id="B914EAFBA518FFDFFF738A06FEAA56F7" box="[151,383,1649,1673]" italics="true" pageId="8" pageNumber="475">Corynopus simplicipes</emphasis>
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Distribution: Eastern
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Q.
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Distribution:
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<emphasis id="B914EAFBA518FFDFFF738E50FE46523C" bold="true" box="[151,403,551,578]" italics="true" pageId="8" pageNumber="475">Rhopalum beaumonti</emphasis>
<bibRefCitation id="EFF14B18A518FFDFFE7D8E51FDEC523F" author="Moczar, L." box="[409,569,550,577]" pageId="8" pageNumber="475" pagination="423 - 426" refId="ref5968" refString="Moczar, L. (1957) Zwei neue Formen der Unterfamilie Crabroninae aus Ungarn (Hymenoptera). Rovartani Kozlemenyek, Series Nova, 10, 423 - 426. [in German]" type="journal article" year="1957">Móczár, 1957</bibRefCitation>
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<collectingCountry id="F3777679A518FFDFFF738E3FFF21521E" box="[151,244,584,608]" name="Hungary" pageId="8" pageNumber="475">Hungary</collectingCountry>
:
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(
<collectingMunicipality id="6BBBAC93A518FFDFFE958E3FFE2A521E" box="[369,511,584,608]" pageId="8" pageNumber="475">Lake Balaton</collectingMunicipality>
),
<collectingCounty id="62BE4E65A518FFDFFDF48E3FFD50521E" box="[528,645,584,608]" pageId="8" pageNumber="475">Gyenesdiás</collectingCounty>
(
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<specimenCount id="9D66FD60A518FFDFFD168E3FFCD2521E" box="[754,775,584,608]" count="1" pageId="8" pageNumber="475" type="male">♂</specimenCount>
)
</materialsCitation>
.
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<paragraph id="8BDF36E9A518FFDFFF738E11FC3B5200" blockId="8.[151,1006,550,639]" box="[151,1006,614,639]" pageId="8" pageNumber="475">
Distribution:
<collectingCountry id="F3777679A518FFDFFEFA8E11FEBE5200" box="[286,363,614,638]" name="Austria" pageId="8" pageNumber="475">Austria</collectingCountry>
, Crimea,
<collectingCountry id="F3777679A518FFDFFE288E11FDFE5200" box="[460,555,614,638]" name="Germany" pageId="8" pageNumber="475">Germany</collectingCountry>
,
<collectingCountry id="F3777679A518FFDFFDD18E10FD5A5201" box="[565,655,615,639]" name="Hungary" pageId="8" pageNumber="475">Hungary</collectingCountry>
,
<collectingCountry id="F3777679A518FFDFFD7D8E11FCC75200" box="[665,786,614,638]" name="Kazakhstan" pageId="8" pageNumber="475">Kazakhstan</collectingCountry>
,
<collectingCountry id="F3777679A518FFDFFCFF8E11FCB65200" box="[795,867,614,638]" name="Turkey" pageId="8" pageNumber="475">Turkey</collectingCountry>
and
<collectingCountry id="F3777679A518FFDFFC718E11FC3F5200" box="[917,1002,614,638]" name="Ukraine" pageId="8" pageNumber="475">Ukraine</collectingCountry>
.
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Genus
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<emphasis id="84C7405CFFC22D6AFF4D1174FE39F99F" bold="true" box="[238,434,1549,1576]" italics="true" pageId="6" pageNumber="377">Huananphasma</emphasis>
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— Ho G.W.C., 2013. Stick Insects of
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. 92.
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<emphasis id="84C7405CFFC22D6AFF3411E1FEB8F905" bold="true" box="[151,307,1688,1714]" pageId="6" pageNumber="377">Type species.</emphasis>
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<emphasis id="84C7405CFFC22D6AFE9911E1FDC7F905" box="[314,588,1688,1714]" italics="true" pageId="6" pageNumber="377">Huananphasma amicum</emphasis>
(Bey-Sienko, 1959)
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:
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1818, 1820, fig. 4 (type locality:
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(
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);
<typeStatus id="690822ECFFC22D6AFF3411C5FF72F961" box="[151,249,1724,1750]" pageId="6" pageNumber="377" type="holotype">holotype</typeStatus>
in
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)
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.
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<emphasis id="84C7405CFFC22D6AFF641199FEEAF94D" bold="true" box="[199,353,1760,1786]" pageId="6" pageNumber="377">Distribution.</emphasis>
<collectingCountry id="CEA4DCDEFFC22D6AFEC41199FE21F94D" box="[359,426,1760,1786]" name="China" pageId="6" pageNumber="377">China</collectingCountry>
(
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Genus
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— Chen S.C. &amp; He Y.H., 2008.
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(
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12, figs. 7–8 (type locality:
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.
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(
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,
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,
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,
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,
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,
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,
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, Hainan);
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<mods:namePart id="8D1A88E401485B6676454A6029EE7DB9">Xie, Chong-Xin</mods:namePart>
<mods:nameIdentifier id="00E1ADBB97EF34743738BFCBC8A5E406" type="ORCID">0000-0003-2104-4806</mods:nameIdentifier>
<mods:affiliation id="B4C356F352FF9B75004540CBE37D248E">Key Laboratory of Forest Disaster Warning and Control in Yunnan Province, Faculty of Forestry, Southwest Forestry University, Kunming, Yunnan 650224, China</mods:affiliation>
</mods:name>
<mods:name id="750B88C3F27BD66F4DC3335A1873D528" type="personal">
<mods:role id="1AEFA5522E550D6FFEEFDDE234DCA32B">
<mods:roleTerm id="2316A63B804A06FF81901CC00574B04C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0584E6E3C9BE8683F2C6A1FADD1E7CF">Chen, Rui</mods:namePart>
<mods:nameIdentifier id="B6BD076F645A2047AF8CCA4FB260367D" type="ORCID">0009-0002-8746-9962</mods:nameIdentifier>
<mods:affiliation id="B418AD4B87294BF8717D831C3F939E26">Key Laboratory of Forest Disaster Warning and Control in Yunnan Province, Faculty of Forestry, Southwest Forestry University, Kunming, Yunnan 650224, China</mods:affiliation>
<mods:nameIdentifier id="ECE8E4D31D9E9AD3301620C53FF6B3B5" type="email">3051769337@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="1D3BB7EF00B3EC46E7F02F8F82443513" type="personal">
<mods:role id="EECE1C2105EDA83E55173D2A87B50A5B">
<mods:roleTerm id="CEA8B780531B1B7BEFA1294892233426">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EDE1B0C6AFEBF3B1E22F893230C2DEE8">Wang, Chen</mods:namePart>
<mods:nameIdentifier id="125B9737F6A95198D2D1E633D97311AE" type="ORCID">0000-0002-2711-2358</mods:nameIdentifier>
<mods:affiliation id="DD95778BC8E0C94C2BFAEB22140B34AE">College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083 China</mods:affiliation>
<mods:nameIdentifier id="18F10309D7C293A9D09629665C0BFA78" type="email">2284482002@qq.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="67F9468963842570CB62FA3D24960CC6">text</mods:typeOfResource>
<mods:relatedItem id="7A91CF1A10105B6C9716252559AE3C29" type="host">
<mods:titleInfo id="7A21425CD0C56CEF120B60AA5109D81D">
<mods:title id="F61C60F5781CE0480D5F76F75C004340">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="056D21996B76CF1DACD1D3EE1363CDAC">
<mods:date id="ABBD1389165DE2B979CB86B505425DC0">2024</mods:date>
<mods:detail id="D2957F90FBFF92CA21594CDDA3EC75CD" type="pubDate">
<mods:number id="4FC5CEE0B0BB5B1651FF11A734B8B376">2024-09-12</mods:number>
</mods:detail>
<mods:detail id="839B974798F3CF1C13B568120456FBB8" type="volume">
<mods:number id="12DE4CAB0588ACBE4177EA9B55C224E9">5507</mods:number>
</mods:detail>
<mods:detail id="2181FFC329A9514D812317F258131623" type="issue">
<mods:number id="3C93C28070EA3EAA33AF3807EEB401EB">2</mods:number>
</mods:detail>
<mods:extent id="D2DD428FEFBBB04F5CD611293EE09807" unit="page">
<mods:start id="B52AEAEC03BB9A646ED6B929C452C8D1">371</mods:start>
<mods:end id="983D6585BC83CF523BD99AB02686FA86">383</mods:end>
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<mods:location id="FBD4A63B50DC725B0C1DDB36270D5F4B">
<mods:url id="2B5FC0E5B7875031FD06AA5AC8C570F9">http://dx.doi.org/10.11646/zootaxa.5507.2.8</mods:url>
</mods:location>
<mods:classification id="462EAC1075FD30A45A96A77E3EEF75E2">journal article</mods:classification>
<mods:identifier id="5E42BAA6DADAB6C19A0E76DE4A835820" type="DOI">10.11646/zootaxa.5507.2.8</mods:identifier>
<mods:identifier id="C61706B824D5EF1AF9A6D39D6B3A071E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="96FF400E8A723E008E9BE868B1201174" type="Zenodo-Dep">13757787</mods:identifier>
<mods:identifier id="52E05145B1D8F9E3B80F7CB30ABC2344" type="ZooBank">673E5A94-62C3-49FC-8719-65E4380586EE</mods:identifier>
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<treatment id="3E1A2D58FFC72D6AFF3417EEFD19FA0B" ID-DOI="http://doi.org/10.5281/zenodo.13757785" ID-Zenodo-Dep="13757785" LSID="urn:lsid:plazi:treatment:3E1A2D58FFC72D6AFF3417EEFD19FA0B" httpUri="http://treatment.plazi.org/id/3E1A2D58FFC72D6AFF3417EEFD19FA0B" lastPageId="6" lastPageNumber="377" pageId="3" pageNumber="374">
<subSubSection id="FEA9CFC5FFC72D6FFF3417EEFD6AFF06" box="[151,737,151,178]" pageId="3" pageNumber="374" type="nomenclature">
<paragraph id="B60C9C4EFFC72D6FFF3417EEFD6AFF06" blockId="3.[151,737,151,178]" box="[151,737,151,178]" pageId="3" pageNumber="374">
<heading id="ED442B22FFC72D6FFF3417EEFD6AFF06" bold="true" box="[151,737,151,178]" fontSize="11" level="1" pageId="3" pageNumber="374" reason="1">
<emphasis id="84C7405CFFC72D6FFF3417EEFD6AFF06" bold="true" box="[151,737,151,178]" pageId="3" pageNumber="374">
<taxonomicName id="71B3E7CDFFC72D6FFF3417EEFE7AFF05" authorityName="Wen &amp; Qian &amp; Xie &amp; Chen &amp; Wang" authorityYear="2024" box="[151,497,151,178]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="ledongense" status="sp. nov.">
<emphasis id="84C7405CFFC72D6FFF3417EEFE7AFF05" bold="true" box="[151,497,151,178]" italics="true" pageId="3" pageNumber="374">Parasinophasma ledongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF4FD27FFC72D6FFE5A17EEFDDDFF06" box="[505,598,151,177]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="2E8880CBFFC72D6FFDC517EEFD53FF06" box="[614,728,151,177]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1800,1824]" captionStartId-1="5.[151,250,1778,1802]" captionTargetBox-0="[151,1435,181,1769]" captionTargetBox-1="[151,1435,181,1754]" captionTargetId-0="figure-22@4.[151,1435,181,1776]" captionTargetId-1="figure-17@5.[151,1435,181,1754]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Parasinophasma ledongense sp. nov., holotype, female A–G, egg H–K. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view. H. egg, dorsal view; I. egg, lateral view; J. egg, top view; K. egg, polar view." captionText-1="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi-0="http://doi.org/10.5281/zenodo.13757789" figureDoi-1="http://doi.org/10.5281/zenodo.13757793" httpUri-0="https://zenodo.org/record/13757789/files/figure.png" httpUri-1="https://zenodo.org/record/13757793/files/figure.png" pageId="3" pageNumber="374">Figs. 1–2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEA9CFC5FFC72D6FFF3417A6FD67FEF6" pageId="3" pageNumber="374" type="materials_examined">
<paragraph id="B60C9C4EFFC72D6FFF3417A6FD67FEF6" blockId="3.[151,1437,223,1869]" pageId="3" pageNumber="374">
<materialsCitation id="06DB9613FFC72D6FFF3417A6FCFAFEA9" collectingDate="2023-07-30" country="China" county="Ledong Li Autonomous County" elevation="800" location="Jianfengling District of National Park" municipality="Female" pageId="3" pageNumber="374" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<emphasis id="84C7405CFFC72D6FFF3417A6FEEFFF4E" bold="true" box="[151,356,223,250]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF3417A6FE8DFF4D" bold="true" box="[151,262,223,250]" italics="true" pageId="3" pageNumber="374">
<typeStatus id="690822ECFFC72D6FFF3417A6FE89FF4D" box="[151,258,223,250]" pageId="3" pageNumber="374" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingMunicipality id="56680634FFC72D6FFEAE17A6FEEFFF4E" box="[269,356,223,249]" pageId="3" pageNumber="374">Female</collectingMunicipality>
</emphasis>
,
<collectingCountry id="CEA4DCDEFFC72D6FFED217A6FE3CFF4E" box="[369,439,223,249]" name="China" pageId="3" pageNumber="374">China</collectingCountry>
,
<collectingRegion id="747752ACFFC72D6FFE6117A6FDF4FF4E" box="[450,639,223,249]" country="China" name="Hainan" pageId="3" pageNumber="374">Hainan Province</collectingRegion>
,
<collectingCounty id="5F6DE4C2FFC72D6FFD2917A6FC64FF4E" box="[650,1007,223,249]" pageId="3" pageNumber="374">Ledong Li Autonomous County</collectingCounty>
,
<location id="B36CCA95FFC72D6FFC5A17A6FA17FF4E" LSID="urn:lsid:plazi:treatment:3E1A2D58FFC72D6AFF3417EEFD19FA0B:B36CCA95FFC72D6FFC5A17A6FA17FF4E" box="[1017,1436,223,249]" country="China" county="Ledong Li Autonomous County" municipality="Female" name="Jianfengling District of National Park" pageId="3" pageNumber="374" stateProvince="Hainan">Jianfengling District of National Park</location>
of
<collectingRegion id="747752ACFFC72D6FFF17167AFE8EFEAA" box="[180,261,259,285]" country="China" name="Hainan" pageId="3" pageNumber="374">Hainan</collectingRegion>
<location id="B36CCA95FFC72D6FFEA8167AFE6DFEAA" LSID="urn:lsid:plazi:treatment:3E1A2D58FFC72D6AFF3417EEFD19FA0B:B36CCA95FFC72D6FFEA8167AFE6DFEAA" box="[267,486,259,285]" country="China" county="Ledong Li Autonomous County" municipality="Female" name="Tropical Rainforest" pageId="3" pageNumber="374" stateProvince="Hainan">Tropical Rainforest</location>
,
<quantity id="714B31ABFFC72D6FFE53167AFDB3FEA9" box="[496,568,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="374" unit="m" value="800.0">
<elevation id="3D9E7B7DFFC72D6FFE53167AFDB3FEA9" box="[496,568,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="374" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="C20DBA8EFFC72D6FFDE1167AFD46FEAA" box="[578,717,259,285]" pageId="3" pageNumber="374" value="2023-07-30">
<collectingDate id="D2494366FFC72D6FFDE1167AFD46FEAA" box="[578,717,259,285]" pageId="3" pageNumber="374" value="2023-07-30">30 Jul. 2023</collectingDate>
</date>
, Leg. Jun Wen
</materialsCitation>
.
<materialsCitation id="06DB9613FFC72D6FFCD8167AFD63FEF6" collectingDate="2023-07-30" collectionCode="SWFU" country="China" county="Ledong Li Autonomous County" elevation="800" location="Jianfengling District of National Park" municipality="Female" pageId="3" pageNumber="374" specimenCount="10" specimenCount-egg="8" specimenCount-female="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="84C7405CFFC72D6FFCD8167AFC63FEAA" bold="true" box="[891,1000,259,285]" italics="true" pageId="3" pageNumber="374">
<typeStatus id="690822ECFFC72D6FFCD8167AFC6FFEAA" box="[891,996,259,285]" pageId="3" pageNumber="374" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCount id="A0B557C7FFC72D6FFC4C167AFBB0FEAA" box="[1007,1083,259,285]" count="1" pageId="3" pageNumber="374" type="male">1 male</specimenCount>
,
<specimenCount id="A0B557C7FFC72D6FFBE5167AFB2DFEAA" box="[1094,1190,259,285]" count="1" pageId="3" pageNumber="374" type="female">1 female</specimenCount>
and
<specimenCount id="A0B557C7FFC72D6FFB79167AFAAFFEA9" box="[1242,1316,259,286]" count="8" pageId="3" pageNumber="374" type="egg">8 eggs</specimenCount>
, same data as holotype (all type specimens deposited in
<collectionCode id="D0A2048BFFC72D6FFD2E165EFD6BFEF6" box="[653,736,295,321]" pageId="3" pageNumber="374">SWFU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEA9CFC5FFC72D6FFF641632FE4EF8FA" pageId="3" pageNumber="374" type="description">
<paragraph id="B60C9C4EFFC72D6FFF641632FA16FE65" blockId="3.[151,1437,223,1869]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF641632FE31FED2" bold="true" box="[199,442,331,357]" pageId="3" pageNumber="374">Description. Female</emphasis>
(
<figureCitation id="2E8880CBFFC72D6FFE6E1632FDC6FED2" box="[461,589,331,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1824]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@4.[151,1435,181,1776]" captionTargetPageId="4" captionText="FIGURE 1. Parasinophasma ledongense sp. nov., holotype, female A–G, egg H–K. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view. H. egg, dorsal view; I. egg, lateral view; J. egg, top view; K. egg, polar view." figureDoi="http://doi.org/10.5281/zenodo.13757789" httpUri="https://zenodo.org/record/13757789/files/figure.png" pageId="3" pageNumber="374">Fig. 1A–G</figureCitation>
). General coloration of body is brown.
<emphasis id="84C7405CFFC72D6FFBBF1632FBE8FED2" bold="true" box="[1052,1123,331,357]" pageId="3" pageNumber="374">Head.</emphasis>
Oblong, longer than wide, smooth, with irregular dark maculation. Occiput distinctly convex, median and lateral longitudinal furrows distinct. Compound eyes rounded and prominent, lacking ocelli. Antennae filiform, densely covered with short bristles, longer than forelegs. Scapus flattened basally, longer than pedicellus, pedicellus about as long as third segment.
</paragraph>
<paragraph id="B60C9C4EFFC72D6FFF3416A2FE5BFC5A" blockId="3.[151,1437,223,1869]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF3416A2FF7CFE42" bold="true" box="[151,247,475,501]" pageId="3" pageNumber="374">Thorax.</emphasis>
Pronotum smooth, rectangular, longer than wide, shorter than head; anterior margin incurved, posterior margin rounded, transverse and longitudinal sulci crossing before middle point. Mesonotum parallel-sided, about 3.5X as long as pronotum, covered with some black granules, median longitudinal line distinct. Mesosternum densely covered with granules, with a transverse carina near anterior margin, with 2–3 short transverse rugose on each side in middle area. Metanotum is about 3/4 the length of the median segment.
<emphasis id="84C7405CFFC72D6FFC431512FBB8FD32" bold="true" box="[992,1075,619,645]" pageId="3" pageNumber="374">Wings.</emphasis>
Tegmina squamiform, posterior margin rounded, slightly longer than metanotum. Anal region of wing gray, with transparent spots. Posterior apices of alae surpassing anterior margin of abdominal tergum VI.
<emphasis id="84C7405CFFC72D6FFCEA15CAFC4EFD7A" bold="true" box="[841,965,691,717]" pageId="3" pageNumber="374">Abdomen.</emphasis>
Cylindrical, smooth. Sternum VII with a distinct crest-like praeopercular organ posteromedially, posterior margin of praeopercular organ extending beyond the tergum VII posterior margin. Tergum VIII longer than tergum IX and anal segment, tergum IX shorter than anal segment. Anal segment median longitudinal ridge distinctly, with a U-shaped emargination on posterior margin. Supra-anal plate small, posterior margin almost truncate, not surpassing posterolateral angles of anal segment. Subgenital plate scoop-shaped, apex pointed and surpassing anterior margin of anal segment. Gonapophyses exposed. Cerci cylindrical, distinctly surpassing posterior margin of anal segment.
<emphasis id="84C7405CFFC72D6FFBF714F2FB18FC12" bold="true" box="[1108,1171,907,933]" pageId="3" pageNumber="374">Legs.</emphasis>
Legs sparsely covered with short bristles, unarmed. Forelegs brown with black maculation, midleg and hindleg green. Hindleg shorter than foreleg, longer than midleg.
</paragraph>
<paragraph id="B60C9C4EFFC72D6FFF64148EFC82F92E" blockId="3.[151,1437,223,1869]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF64148EFE8FFBA6" bold="true" box="[199,260,1015,1041]" pageId="3" pageNumber="374">Male</emphasis>
(
<figureCitation id="2E8880CBFFC72D6FFEB0148EFE1AFBA6" box="[275,401,1015,1041]" captionStart="FIGURE 2" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1435,181,1754]" captionTargetId="figure-17@5.[151,1435,181,1754]" captionTargetPageId="5" captionText="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi="http://doi.org/10.5281/zenodo.13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="3" pageNumber="374">Fig. 2A–H</figureCitation>
). Smaller than female. General coloration of body is light brown.
<emphasis id="84C7405CFFC72D6FFBCD148EFB3EFBA6" bold="true" box="[1134,1205,1015,1041]" pageId="3" pageNumber="374">Head.</emphasis>
Oblong, longer than wide, longer and broader than pronotum. Vertex flat, there are two small depressions in the central position. Occiput weakly convex, median and lateral longitudinal furrows distinct. Compound eyes rounded and larger than female, lacking ocelli. Genae with two black postocular stripes. Antennae filiform, densely covered with short bristles, longer than forelegs. Scapus flattened basally, longer than pedicellus and third segment, pedicellus shorter than third segment.
<emphasis id="84C7405CFFC72D6FFEA713D2FEEFFB72" bold="true" box="[260,356,1195,1221]" pageId="3" pageNumber="374">Thorax.</emphasis>
Pronotum trapezium, anterior margin wide, posterior margin narrow, with a black herringbone stripe in the center, transverse and longitudinal sulci crossing before middle point. Mesonotum gradually broader on posterior, about 4.5X as long as pronotum, sparsely covered with a few granules, median longitudinal line distinct, a black longitudinal stripe at each side of mesonotum. Mesosternum with 2–3 short transverse rugose on each side in first half part. Metanotum about 2/3 the length of the median segment.
<emphasis id="84C7405CFFC72D6FFC7D1242FBBBFAE2" bold="true" box="[990,1072,1339,1365]" pageId="3" pageNumber="374">Wings.</emphasis>
Tegmina squamiform, posterior margin rounded, shorter than metanotum. Anal region of wing gray, with transparent spots. Posterior apices of alae reaching anterior margin of abdominal tergum VI.
<emphasis id="84C7405CFFC72D6FFD7A12FAFCDEFA2A" bold="true" box="[729,853,1411,1437]" pageId="3" pageNumber="374">Abdomen.</emphasis>
Cylindrical, smooth, unarmed, with some irregular maculation. Tergum VII to anal segment shorter than other abdomen segments. Tergum VIII about as long as anal segment, shorter than tergum IX. Anal segment median longitudinal carina distinct, with a U-shaped emargination on posterior margin, the two posterolateral angles spine shaped. Vomer small and symmetric, apex rounded, with two longitudinal ridges. Poculum cup-shaped, posterior margin truncate and surpassing anterior margin of anal segment. Cerci cylindrical, tapering posteriorly, apices pointed and curved down, not reaching posterolateral angles of anal segment.
<emphasis id="84C7405CFFC72D6FFEFD1122FE16F9C2" bold="true" box="[350,413,1627,1653]" pageId="3" pageNumber="374">Legs.</emphasis>
Legs sparsely covered with short bristles, unarmed. Forelegs brown, midleg and hindleg green. Foreleg shorter than hindleg, longer than midleg.
</paragraph>
<paragraph id="B60C9C4EFFC72D6FFF6411DAFE84F89E" blockId="3.[151,1437,223,1869]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF6411DAFE8AF90A" bold="true" box="[199,257,1699,1725]" pageId="3" pageNumber="374">Eggs</emphasis>
(
<figureCitation id="2E8880CBFFC72D6FFEB711DAFE1FF90A" box="[276,404,1699,1725]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1824]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@4.[151,1435,181,1776]" captionTargetPageId="4" captionText="FIGURE 1. Parasinophasma ledongense sp. nov., holotype, female A–G, egg H–K. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view. H. egg, dorsal view; I. egg, lateral view; J. egg, top view; K. egg, polar view." figureDoi="http://doi.org/10.5281/zenodo.13757789" httpUri="https://zenodo.org/record/13757789/files/figure.png" pageId="3" pageNumber="374">Fig. 1H–K</figureCitation>
). Capsule oval, general brown and with irregular dark maculation, covered with densely granulation and long bristles. Micropylar drop-shaped, with dense granulation, micropylar cup position is near the bottom of micropylar plate. Median line indistinct. Operculum almost rounded, convex medially, lacking capitulum.
</paragraph>
<paragraph id="B60C9C4EFFC72D6FFF64104AFE4EF8FA" blockId="3.[151,1437,223,1869]" box="[199,453,1843,1869]" pageId="3" pageNumber="374">
<emphasis id="84C7405CFFC72D6FFF64104AFE4EF8FA" bold="true" box="[199,453,1843,1869]" pageId="3" pageNumber="374">Measurements (mm).</emphasis>
</paragraph>
</subSubSection>
<caption id="E2CCCCC6FFC02D68FF341071FBB8F8DF" ID-DOI="http://doi.org/10.5281/zenodo.13757789" ID-Zenodo-Dep="13757789" httpUri="https://zenodo.org/record/13757789/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,250,1800,1824]" targetBox="[151,1435,181,1769]" targetPageId="4" targetType="figure">
<paragraph id="B60C9C4EFFC02D68FF341071FBB8F8DF" blockId="4.[151,1437,1800,1897]" pageId="4" pageNumber="375">
<emphasis id="84C7405CFFC02D68FF341071FBCAF897" bold="true" box="[151,1089,1800,1824]" pageId="4" pageNumber="375">
FIGURE 1.
<taxonomicName id="71B3E7CDFFC02D68FEBC1071FDC5F897" authorityName="Wen &amp; Qian &amp; Xie &amp; Chen &amp; Wang" authorityYear="2024" box="[287,590,1800,1824]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="ledongense" status="sp. nov.">
<emphasis id="84C7405CFFC02D68FEBC1071FDC5F897" bold="true" box="[287,590,1800,1824]" italics="true" pageId="4" pageNumber="375">Parasinophasma ledongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF4FD27FFC02D68FDF41071FD27F897" box="[599,684,1800,1824]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female A–G, egg H–K. A.
</emphasis>
habitus, dorsal view;
<emphasis id="84C7405CFFC02D68FA8F1071FAC8F897" bold="true" box="[1324,1347,1800,1824]" pageId="4" pageNumber="375">B.</emphasis>
habitus, lateral view;
<emphasis id="84C7405CFFC02D68FEBE1055FEBDF8F3" bold="true" box="[285,310,1836,1860]" pageId="4" pageNumber="375">C.</emphasis>
head, dorsal view;
<emphasis id="84C7405CFFC02D68FE5C1055FD93F8F3" bold="true" box="[511,536,1836,1860]" pageId="4" pageNumber="375">D.</emphasis>
head, lateral view;
<emphasis id="84C7405CFFC02D68FD411055FD72F8F3" bold="true" box="[738,761,1836,1860]" pageId="4" pageNumber="375">E.</emphasis>
terminalia, dorsal view;
<emphasis id="84C7405CFFC02D68FC5A1055FB87F8F3" bold="true" box="[1017,1036,1836,1860]" pageId="4" pageNumber="375">F.</emphasis>
terminalia, lateral view;
<emphasis id="84C7405CFFC02D68FAAD1055FAA3F8F3" bold="true" box="[1294,1320,1836,1860]" pageId="4" pageNumber="375">G.</emphasis>
terminalia, ventral view.
<emphasis id="84C7405CFFC02D68FE831029FEB1F8DF" bold="true" box="[288,314,1872,1896]" pageId="4" pageNumber="375">H.</emphasis>
egg, dorsal view;
<emphasis id="84C7405CFFC02D68FE561029FD8EF8DF" bold="true" box="[501,517,1872,1896]" pageId="4" pageNumber="375">I.</emphasis>
egg, lateral view;
<emphasis id="84C7405CFFC02D68FD621029FD5FF8DF" bold="true" box="[705,724,1872,1896]" pageId="4" pageNumber="375">J.</emphasis>
egg, top view;
<emphasis id="84C7405CFFC02D68FCD31029FC00F8DF" bold="true" box="[880,907,1872,1896]" pageId="4" pageNumber="375">K.</emphasis>
egg, polar view.
</paragraph>
</caption>
<caption id="E2CCCCC6FFC12D69FF34118BFDBAF8E5" ID-DOI="http://doi.org/10.5281/zenodo.13757793" ID-Zenodo-Dep="13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="5" pageNumber="376" startId="5.[151,250,1778,1802]" targetBox="[151,1435,181,1754]" targetPageId="5" targetType="figure">
<paragraph id="B60C9C4EFFC12D69FF34118BFDBAF8E5" blockId="5.[151,1436,1778,1874]" pageId="5" pageNumber="376">
<emphasis id="84C7405CFFC12D69FF34118BFB86F8BD" bold="true" box="[151,1037,1778,1802]" pageId="5" pageNumber="376">
FIGURE 2.
<taxonomicName id="71B3E7CDFFC12D69FEBA118BFDCFF8BD" authorityName="Wen &amp; Qian &amp; Xie &amp; Chen &amp; Wang" authorityYear="2024" box="[281,580,1778,1802]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="ledongense" status="sp. nov.">
<emphasis id="84C7405CFFC12D69FEBA118BFDCFF8BD" bold="true" box="[281,580,1778,1802]" italics="true" pageId="5" pageNumber="376">Parasinophasma ledongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF4FD27FFC12D69FDE9118BFD17F8BD" box="[586,668,1778,1802]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, male A–H, habitat I. A.
</emphasis>
habitus, dorsal view;
<emphasis id="84C7405CFFC12D69FB49118BFA8AF8BD" bold="true" box="[1258,1281,1778,1802]" pageId="5" pageNumber="376">B.</emphasis>
habitus, lateral view;
<emphasis id="84C7405CFFC12D69FF75106FFF64F899" bold="true" box="[214,239,1814,1838]" pageId="5" pageNumber="376">C.</emphasis>
head, dorsal view;
<emphasis id="84C7405CFFC12D69FE14106FFE44F899" bold="true" box="[439,463,1814,1838]" pageId="5" pageNumber="376">D.</emphasis>
head, lateral view;
<emphasis id="84C7405CFFC12D69FD3B106FFD24F899" bold="true" box="[664,687,1814,1838]" pageId="5" pageNumber="376">E.</emphasis>
terminalia, dorsal view;
<emphasis id="84C7405CFFC12D69FC0D106FFC4AF899" bold="true" box="[942,961,1814,1838]" pageId="5" pageNumber="376">F.</emphasis>
terminalia, lateral view;
<emphasis id="84C7405CFFC12D69FB62106FFB50F899" bold="true" box="[1217,1243,1814,1838]" pageId="5" pageNumber="376">G.</emphasis>
terminalia, ventral view;
<emphasis id="84C7405CFFC12D69FF751043FF7BF8E5" bold="true" box="[214,240,1850,1874]" pageId="5" pageNumber="376">H.</emphasis>
vomer, ventral view;
<emphasis id="84C7405CFFC12D69FE6D1043FE55F8E5" bold="true" box="[462,478,1850,1874]" pageId="5" pageNumber="376">I.</emphasis>
habitat.
</paragraph>
</caption>
<paragraph id="B60C9C4EFFC22D6AFF0017CEFC52FD71" pageId="6" pageNumber="377">
<table id="C4B36EEEFFC2D293FF0017DAFAE5FD71" box="[163,1390,163,710]" gridcols="6" gridrows="13" pageId="6" pageNumber="377">
<tr id="08839E0CFFC2D293FF0017DAFAE5FF54" box="[163,1390,163,227]" gridrow="0" pageId="6" pageNumber="377">
<th id="4B52F770FFC2D293FF0017DAFEC1FF54" box="[163,330,163,227]" gridcol="0" gridrow="0" pageId="6" pageNumber="377">Length(mm)</th>
<th id="4B52F770FFC2D293FE5E17DAFDD0FF54" box="[509,603,163,227]" gridcol="1" gridrow="0" pageId="6" pageNumber="377">Holotype Female</th>
<th id="4B52F770FFC2D293FD7417DAFC01FF54" box="[727,906,163,227]" gridcol="2" gridrow="0" pageId="6" pageNumber="377">Paratype Female</th>
<th id="4B52F770FFC2D293FC0A17DAFC56FF54" box="[937,989,163,227]" gridcol="3" gridrow="0" pageId="6" pageNumber="377">Male</th>
<th id="4B52F770FFC2D293FBCB17DAFAE5FF54" box="[1128,1390,163,227]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="6" pageNumber="377">Egg (n=8)</th>
</tr>
<tr id="08839E0CFFC2D293FF00178AFAE5FEBC" box="[163,1390,243,267]" gridrow="1" pageId="6" pageNumber="377">
<th id="4B52F770FFC2D293FF00178AFEC1FEBC" box="[163,330,243,267]" gridcol="0" gridrow="1" pageId="6" pageNumber="377">Body</th>
<td id="4B52F770FFC2D293FE5E178AFDD0FEBC" box="[509,603,243,267]" gridcol="1" gridrow="1" pageId="6" pageNumber="377">92.6</td>
<td id="4B52F770FFC2D293FD74178AFC01FEBC" box="[727,906,243,267]" gridcol="2" gridrow="1" pageId="6" pageNumber="377">92.2</td>
<td id="4B52F770FFC2D293FC0A178AFC56FEBC" box="[937,989,243,267]" gridcol="3" gridrow="1" pageId="6" pageNumber="377">66.1</td>
<td id="4B52F770FFC2D293FBCB178AFB3BFEBC" box="[1128,1200,243,267]" gridcol="4" gridrow="1" pageId="6" pageNumber="377">Length</td>
<td id="4B52F770FFC2D293FA81178AFAE5FEBC" box="[1314,1390,243,267]" gridcol="5" gridrow="1" pageId="6" pageNumber="377">2.8–3.0</td>
</tr>
<tr id="08839E0CFFC2D293FF001662FAE5FE84" box="[163,1390,283,307]" gridrow="2" pageId="6" pageNumber="377">
<th id="4B52F770FFC2D293FF001662FEC1FE84" box="[163,330,283,307]" gridcol="0" gridrow="2" pageId="6" pageNumber="377">Head</th>
<td id="4B52F770FFC2D293FE5E1662FDD0FE84" box="[509,603,283,307]" gridcol="1" gridrow="2" pageId="6" pageNumber="377">5.5</td>
<td id="4B52F770FFC2D293FD741662FC01FE84" box="[727,906,283,307]" gridcol="2" gridrow="2" pageId="6" pageNumber="377">5.4</td>
<td id="4B52F770FFC2D293FC0A1662FC56FE84" box="[937,989,283,307]" gridcol="3" gridrow="2" pageId="6" pageNumber="377">3.5</td>
<td id="4B52F770FFC2D293FBCB1662FB3BFE84" box="[1128,1200,283,307]" gridcol="4" gridrow="2" pageId="6" pageNumber="377">Width</td>
<td id="4B52F770FFC2D293FA811662FAE5FE84" box="[1314,1390,283,307]" gridcol="5" gridrow="2" pageId="6" pageNumber="377">1.7–1.8</td>
</tr>
<tr id="08839E0CFFC2D293FF00163DFAE5FEEB" box="[163,1390,324,348]" gridrow="3" pageId="6" pageNumber="377">
<th id="4B52F770FFC2D293FF00163DFEC1FEEB" box="[163,330,324,348]" gridcol="0" gridrow="3" pageId="6" pageNumber="377">Pronotum</th>
<td id="4B52F770FFC2D293FE5E163DFDD0FEEB" box="[509,603,324,348]" gridcol="1" gridrow="3" pageId="6" pageNumber="377">4.0</td>
<td id="4B52F770FFC2D293FD74163DFC01FEEB" box="[727,906,324,348]" gridcol="2" gridrow="3" pageId="6" pageNumber="377">3.9</td>
<td id="4B52F770FFC2D293FC0A163DFC56FEEB" box="[937,989,324,348]" gridcol="3" gridrow="3" pageId="6" pageNumber="377">2.8</td>
<td id="4B52F770FFC2D293FBCB163DFB3BFEEB" box="[1128,1200,324,348]" gridcol="4" gridrow="3" pageId="6" pageNumber="377">Height</td>
<td id="4B52F770FFC2D293FA81163DFAE5FEEB" box="[1314,1390,324,348]" gridcol="5" gridrow="3" pageId="6" pageNumber="377">1.9–2.0</td>
</tr>
<tr id="08839E0CFFC2D293FF001615FAE5FE33" box="[163,1390,364,388]" gridrow="4" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF001615FEC1FE33" box="[163,330,364,388]" gridcol="0" gridrow="4" pageId="6" pageNumber="377">Mesonotum</th>
<td id="4B52F770FFC2D293FE5E1615FDD0FE33" box="[509,603,364,388]" gridcol="1" gridrow="4" pageId="6" pageNumber="377">13.9</td>
<td id="4B52F770FFC2D293FD741615FC01FE33" box="[727,906,364,388]" gridcol="2" gridrow="4" pageId="6" pageNumber="377">13.7</td>
<td id="4B52F770FFC2D293FC0A1615FC56FE33" box="[937,989,364,388]" gridcol="3" gridrow="4" pageId="6" pageNumber="377">10.6</td>
</tr>
<tr id="08839E0CFFC2D293FF0016EDFAE5FE1B" box="[163,1390,404,428]" gridrow="5" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF0016EDFEC1FE1B" box="[163,330,404,428]" gridcol="0" gridrow="5" pageId="6" pageNumber="377">Metanotum</th>
<td id="4B52F770FFC2D293FE5E16EDFDD0FE1B" box="[509,603,404,428]" gridcol="1" gridrow="5" pageId="6" pageNumber="377">5.2</td>
<td id="4B52F770FFC2D293FD7416EDFC01FE1B" box="[727,906,404,428]" gridcol="2" gridrow="5" pageId="6" pageNumber="377">5.2</td>
<td id="4B52F770FFC2D293FC0A16EDFC56FE1B" box="[937,989,404,428]" gridcol="3" gridrow="5" pageId="6" pageNumber="377">4.1</td>
</tr>
<tr id="08839E0CFFC2D293FF0016C5FAE5FE62" box="[163,1390,444,469]" gridrow="6" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF0016C5FEC1FE62" box="[163,330,444,469]" gridcol="0" gridrow="6" pageId="6" pageNumber="377">Median segment</th>
<td id="4B52F770FFC2D293FE5E16C5FDD0FE62" box="[509,603,444,469]" gridcol="1" gridrow="6" pageId="6" pageNumber="377">6.1</td>
<td id="4B52F770FFC2D293FD7416C5FC01FE62" box="[727,906,444,469]" gridcol="2" gridrow="6" pageId="6" pageNumber="377">5.9</td>
<td id="4B52F770FFC2D293FC0A16C5FC56FE62" box="[937,989,444,469]" gridcol="3" gridrow="6" pageId="6" pageNumber="377">4.6</td>
</tr>
<tr id="08839E0CFFC2D293FF00169DFAE5FE4A" box="[163,1390,484,509]" gridrow="7" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF00169DFEC1FE4A" box="[163,330,484,509]" gridcol="0" gridrow="7" pageId="6" pageNumber="377">Profemora</th>
<td id="4B52F770FFC2D293FE5E169DFDD0FE4A" box="[509,603,484,509]" gridcol="1" gridrow="7" pageId="6" pageNumber="377">19.1</td>
<td id="4B52F770FFC2D293FD74169DFC01FE4A" box="[727,906,484,509]" gridcol="2" gridrow="7" pageId="6" pageNumber="377">18.8</td>
<td id="4B52F770FFC2D293FC0A169DFC56FE4A" box="[937,989,484,509]" gridcol="3" gridrow="7" pageId="6" pageNumber="377">17.6</td>
</tr>
<tr id="08839E0CFFC2D293FF001574FAE5FD92" box="[163,1390,525,549]" gridrow="8" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF001574FEC1FD92" box="[163,330,525,549]" gridcol="0" gridrow="8" pageId="6" pageNumber="377">Mesofemora</th>
<td id="4B52F770FFC2D293FE5E1574FDD0FD92" box="[509,603,525,549]" gridcol="1" gridrow="8" pageId="6" pageNumber="377">11.1</td>
<td id="4B52F770FFC2D293FD741574FC01FD92" box="[727,906,525,549]" gridcol="2" gridrow="8" pageId="6" pageNumber="377">10.9</td>
<td id="4B52F770FFC2D293FC0A1574FC56FD92" box="[937,989,525,549]" gridcol="3" gridrow="8" pageId="6" pageNumber="377">12.4</td>
</tr>
<tr id="08839E0CFFC2D293FF00154CFAE5FDFA" box="[163,1390,565,589]" gridrow="9" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF00154CFEC1FDFA" box="[163,330,565,589]" gridcol="0" gridrow="9" pageId="6" pageNumber="377">Metafemora</th>
<td id="4B52F770FFC2D293FE5E154CFDD0FDFA" box="[509,603,565,589]" gridcol="1" gridrow="9" pageId="6" pageNumber="377">16.8</td>
<td id="4B52F770FFC2D293FD74154CFC01FDFA" box="[727,906,565,589]" gridcol="2" gridrow="9" pageId="6" pageNumber="377">16.5</td>
<td id="4B52F770FFC2D293FC0A154CFC56FDFA" box="[937,989,565,589]" gridcol="3" gridrow="9" pageId="6" pageNumber="377">17.3</td>
</tr>
<tr id="08839E0CFFC2D293FF001524FAE5FDC2" box="[163,1390,605,629]" gridrow="10" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF001524FEC1FDC2" box="[163,330,605,629]" gridcol="0" gridrow="10" pageId="6" pageNumber="377">Protibiae</th>
<td id="4B52F770FFC2D293FE5E1524FDD0FDC2" box="[509,603,605,629]" gridcol="1" gridrow="10" pageId="6" pageNumber="377">16.9</td>
<td id="4B52F770FFC2D293FD741524FC01FDC2" box="[727,906,605,629]" gridcol="2" gridrow="10" pageId="6" pageNumber="377">16.7</td>
<td id="4B52F770FFC2D293FC0A1524FC56FDC2" box="[937,989,605,629]" gridcol="3" gridrow="10" pageId="6" pageNumber="377">16.2</td>
</tr>
<tr id="08839E0CFFC2D293FF0015FCFAE5FD29" box="[163,1390,645,670]" gridrow="11" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF0015FCFEC1FD29" box="[163,330,645,670]" gridcol="0" gridrow="11" pageId="6" pageNumber="377">Mesotibiae</th>
<td id="4B52F770FFC2D293FE5E15FCFDD0FD29" box="[509,603,645,670]" gridcol="1" gridrow="11" pageId="6" pageNumber="377">9.8</td>
<td id="4B52F770FFC2D293FD7415FCFC01FD29" box="[727,906,645,670]" gridcol="2" gridrow="11" pageId="6" pageNumber="377">9.5</td>
<td id="4B52F770FFC2D293FC0A15FCFC56FD29" box="[937,989,645,670]" gridcol="3" gridrow="11" pageId="6" pageNumber="377">10.4</td>
</tr>
<tr id="08839E0CFFC2D293FF0015D7FAE5FD71" box="[163,1390,686,710]" gridrow="12" pageId="6" pageNumber="377" rowspan-4="1" rowspan-5="1">
<th id="4B52F770FFC2D293FF0015D7FEC1FD71" box="[163,330,686,710]" gridcol="0" gridrow="12" pageId="6" pageNumber="377">Metatibiae</th>
<td id="4B52F770FFC2D293FE5E15D7FDD0FD71" box="[509,603,686,710]" gridcol="1" gridrow="12" pageId="6" pageNumber="377">13.5</td>
<td id="4B52F770FFC2D293FD7415D7FC01FD71" box="[727,906,686,710]" gridcol="2" gridrow="12" pageId="6" pageNumber="377">13.3</td>
<td id="4B52F770FFC2D293FC0A15D7FC56FD71" box="[937,989,686,710]" gridcol="3" gridrow="12" pageId="6" pageNumber="377">16.3</td>
</tr>
</table>
</paragraph>
<subSubSection id="FEA9CFC5FFC22D6AFF641460FD97FC84" box="[199,540,793,819]" pageId="6" pageNumber="377" type="distribution">
<paragraph id="B60C9C4EFFC22D6AFF641460FD97FC84" blockId="6.[151,1437,793,1468]" box="[199,540,793,819]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF641460FEEAFC84" bold="true" box="[199,353,793,819]" pageId="6" pageNumber="377">Distribution.</emphasis>
<collectingCountry id="CEA4DCDEFFC22D6AFEC41460FE21FC84" box="[359,426,793,819]" name="China" pageId="6" pageNumber="377">China</collectingCountry>
(
<collectingRegion id="747752ACFFC22D6AFE1A1460FD9BFC84" box="[441,528,793,819]" country="China" name="Hainan" pageId="6" pageNumber="377">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="FEA9CFC5FFC22D6AFF641444FEF3FC17" pageId="6" pageNumber="377" type="diagnosis">
<paragraph id="B60C9C4EFFC22D6AFF641444FEF3FC17" blockId="6.[151,1437,793,1468]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF641444FEC9FCE0" bold="true" box="[199,322,829,855]" pageId="6" pageNumber="377">Diagnosis.</emphasis>
The females of new species are distinguished by a U-shaped emargination on the posterior margin of the anal segment, and gonapophyses are exposed. Males are distinguished by two spiny situated posterolaterally to the anal segment.
</paragraph>
</subSubSection>
<subSubSection id="FEA9CFC5FFC22D6AFF6414D0FC9AFAF8" pageId="6" pageNumber="377" type="discussion">
<paragraph id="B60C9C4EFFC22D6AFF6414D0FC9AFAF8" blockId="6.[151,1437,793,1468]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF6414D0FEB0FC74" bold="true" box="[199,315,937,963]" pageId="6" pageNumber="377">Remarks.</emphasis>
The female of the new species is similar to
<taxonomicName id="71B3E7CDFFC22D6AFC8014D3FB38FC74" authority="Ho, 2017" authorityName="Ho" authorityYear="2017" box="[803,1203,937,964]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="luchunense">
<emphasis id="84C7405CFFC22D6AFC8014D3FBCAFC74" box="[803,1089,937,963]" italics="true" pageId="6" pageNumber="377">P. luchunense luchunense</emphasis>
<bibRefCitation id="D222E1BFFFC22D6AFBE414D3FB38FC74" author="Ho, G. W. C." box="[1095,1203,937,964]" pageId="6" pageNumber="377" pagination="181 - 194" refId="ref8165" refString="Ho, G. W. C. (2017) Contribution to the knowledge of Oriental Phasmatodea I: A taxonomic study of the genus Parasinophasma (Phasmatodea: Necrosciinae). Journal of Orthoptera Research, 26 (2), 181 - 194. https: // doi. org / 10.3897 / jor. 26.15289" type="journal article" year="2017">Ho, 2017</bibRefCitation>
</taxonomicName>
. In the female of this new species, the sternum VII has a distinct crest-like praeopercular organ posteromedially and the gonapophyses are exposed (
<figureCitation id="2E8880CBFFC22D6AFEA21488FE2EFBBC" box="[257,421,1009,1036]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1824]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@4.[151,1435,181,1776]" captionTargetPageId="4" captionText="FIGURE 1. Parasinophasma ledongense sp. nov., holotype, female A–G, egg H–K. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view. H. egg, dorsal view; I. egg, lateral view; J. egg, top view; K. egg, polar view." figureDoi="http://doi.org/10.5281/zenodo.13757789" httpUri="https://zenodo.org/record/13757789/files/figure.png" pageId="6" pageNumber="377">Fig. 1 F and G</figureCitation>
). In
<taxonomicName id="71B3E7CDFFC22D6AFE76148BFD79FBBC" authorityName="Ho" authorityYear="2017" box="[469,754,1009,1035]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="luchunense">
<emphasis id="84C7405CFFC22D6AFE76148BFD79FBBC" box="[469,754,1009,1035]" italics="true" pageId="6" pageNumber="377">P. luchunense luchunense</emphasis>
</taxonomicName>
, the sternum VII has a dorsoventrally flattened praeopercular organ posteromedially and gonapophyses are not exposed (
<bibRefCitation id="D222E1BFFFC22D6AFC90136CFC29FB98" author="Ho, G. W. C." box="[819,930,1045,1071]" pageId="6" pageNumber="377" pagination="181 - 194" refId="ref8165" refString="Ho, G. W. C. (2017) Contribution to the knowledge of Oriental Phasmatodea I: A taxonomic study of the genus Parasinophasma (Phasmatodea: Necrosciinae). Journal of Orthoptera Research, 26 (2), 181 - 194. https: // doi. org / 10.3897 / jor. 26.15289" type="journal article" year="2017">Ho, 2017</bibRefCitation>
). The male of the new species is also similar to
<taxonomicName id="71B3E7CDFFC22D6AFF161343FDCDFBE4" authority="Ho, 2017" authorityName="Ho" authorityYear="2017" box="[181,582,1081,1108]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="luchunense">
<emphasis id="84C7405CFFC22D6AFF161343FE5EFBE4" box="[181,469,1081,1107]" italics="true" pageId="6" pageNumber="377">P. luchunense luchunense</emphasis>
<bibRefCitation id="D222E1BFFFC22D6AFE7E1343FDCDFBE4" author="Ho, G. W. C." box="[477,582,1081,1108]" pageId="6" pageNumber="377" pagination="181 - 194" refId="ref8165" refString="Ho, G. W. C. (2017) Contribution to the knowledge of Oriental Phasmatodea I: A taxonomic study of the genus Parasinophasma (Phasmatodea: Necrosciinae). Journal of Orthoptera Research, 26 (2), 181 - 194. https: // doi. org / 10.3897 / jor. 26.15289" type="journal article" year="2017">Ho, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="71B3E7CDFFC22D6AFDDD1343FC78FBE4" authority="Ho, 2017" authorityName="Ho" authorityYear="2017" box="[638,1011,1081,1108]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="xingyuei">
<emphasis id="84C7405CFFC22D6AFDDD1343FCF4FBE4" box="[638,895,1081,1107]" italics="true" pageId="6" pageNumber="377">P. luchunense xingyuei</emphasis>
<bibRefCitation id="D222E1BFFFC22D6AFC251343FC78FBE4" author="Ho, G. W. C." box="[902,1011,1081,1108]" pageId="6" pageNumber="377" pagination="181 - 194" refId="ref8165" refString="Ho, G. W. C. (2017) Contribution to the knowledge of Oriental Phasmatodea I: A taxonomic study of the genus Parasinophasma (Phasmatodea: Necrosciinae). Journal of Orthoptera Research, 26 (2), 181 - 194. https: // doi. org / 10.3897 / jor. 26.15289" type="journal article" year="2017">Ho, 2017</bibRefCitation>
</taxonomicName>
. But male of the new species has two longitudinal ridges and a rounded posterior margin of vomer (
<figureCitation id="2E8880CBFFC22D6AFCFB1324FC31FBC0" box="[856,954,1117,1143]" captionStart="FIGURE 2" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1435,181,1754]" captionTargetId="figure-17@5.[151,1435,181,1754]" captionTargetPageId="5" captionText="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi="http://doi.org/10.5281/zenodo.13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="6" pageNumber="377">Fig. 2 H</figureCitation>
) and the posterior apices of the alae reach the anterior margin of tergum VI (
<figureCitation id="2E8880CBFFC22D6AFDB413F8FD36FB2C" box="[535,701,1153,1179]" captionStart="FIGURE 2" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1435,181,1754]" captionTargetId="figure-17@5.[151,1435,181,1754]" captionTargetPageId="5" captionText="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi="http://doi.org/10.5281/zenodo.13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="6" pageNumber="377">Fig. 2 A and E</figureCitation>
), additionally, posterior margin of poculum slightly surpasses the anterior margin of anal segment, with a black herringbone stripe on its pronotum (
<figureCitation id="2E8880CBFFC22D6AFB8F13DCFB5AFB08" box="[1068,1233,1189,1215]" captionStart="FIGURE 2" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1435,181,1754]" captionTargetId="figure-17@5.[151,1435,181,1754]" captionTargetPageId="5" captionText="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi="http://doi.org/10.5281/zenodo.13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="6" pageNumber="377">Fig. 2 C and F</figureCitation>
). In
<taxonomicName id="71B3E7CDFFC22D6AFAA013DFFE9EFB54" authorityName="Ho" authorityYear="2017" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="luchunense">
<emphasis id="84C7405CFFC22D6AFAA013DFFE9EFB54" italics="true" pageId="6" pageNumber="377">P. luchunense luchunense</emphasis>
</taxonomicName>
, there is a weak emargination forming two indistinct lateral elevations at the apex of the vomer and the posterior apices of the alae reach only to the middle area of tergum IV; in
<emphasis id="84C7405CFFC22D6AFB871397FAA4FAB0" box="[1060,1327,1261,1287]" italics="true" pageId="6" pageNumber="377">
<taxonomicName id="71B3E7CDFFC22D6AFB871397FAA0FAB0" authorityName="Ho" authorityYear="2017" box="[1060,1323,1261,1287]" class="Insecta" family="Lonchodidae" genus="Parasinophasma" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="377" phylum="Arthropoda" rank="subSpecies" species="luchunense" subSpecies="xingyuei">P. luchunense xingyuei</taxonomicName>
,
</emphasis>
posterior margin of poculum does not reach the anterior margin of anal segment and pronotum has two short blackish brown longitudinal stripes behind transverse sulcus (
<bibRefCitation id="D222E1BFFFC22D6AFD34124CFC8EFAF8" author="Ho, G. W. C." box="[663,773,1333,1359]" pageId="6" pageNumber="377" pagination="181 - 194" refId="ref8165" refString="Ho, G. W. C. (2017) Contribution to the knowledge of Oriental Phasmatodea I: A taxonomic study of the genus Parasinophasma (Phasmatodea: Necrosciinae). Journal of Orthoptera Research, 26 (2), 181 - 194. https: // doi. org / 10.3897 / jor. 26.15289" type="journal article" year="2017">Ho, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FEA9CFC5FFC22D6AFF641220FD19FA0B" pageId="6" pageNumber="377" type="etymology">
<paragraph id="B60C9C4EFFC22D6AFF641220FB65FAC4" blockId="6.[151,1437,793,1468]" box="[199,1262,1369,1396]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF641220FEC5FAC4" bold="true" box="[199,334,1369,1395]" pageId="6" pageNumber="377">Etymology.</emphasis>
This new species is named after the
<typeStatus id="690822ECFFC22D6AFD4A1223FC92FAC3" box="[745,793,1370,1396]" pageId="6" pageNumber="377">type</typeStatus>
locality; Ledong Li Autonomous County.
</paragraph>
<paragraph id="B60C9C4EFFC22D6AFF641204FA0CFA20" blockId="6.[151,1437,793,1468]" box="[199,1415,1405,1432]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF641204FEA1FA20" bold="true" box="[199,298,1405,1431]" pageId="6" pageNumber="377">Habitat.</emphasis>
The new species inhabits forest at an elevation of
<quantity id="714B31ABFFC22D6AFCC31204FC2CFA2F" box="[864,935,1405,1432]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="377" unit="m" value="800.0">800 m</quantity>
and feeds on leaves of
<taxonomicName id="71B3E7CDFFC22D6AFB0E1207FA9EFA2F" authorityName="Dumortier" authorityYear="1829" box="[1197,1301,1406,1432]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="377" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
(
<figureCitation id="2E8880CBFFC22D6AFA861204FAF2FA20" box="[1317,1401,1405,1431]" captionStart="FIGURE 2" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1435,181,1754]" captionTargetId="figure-17@5.[151,1435,181,1754]" captionTargetPageId="5" captionText="FIGURE 2. Parasinophasma ledongense sp. nov., paratype, male A–H, habitat I. A. habitus, dorsal view; B. habitus, lateral view; C. head, dorsal view; D. head, lateral view; E. terminalia, dorsal view; F. terminalia, lateral view; G. terminalia, ventral view; H. vomer, ventral view; I. habitat." figureDoi="http://doi.org/10.5281/zenodo.13757793" httpUri="https://zenodo.org/record/13757793/files/figure.png" pageId="6" pageNumber="377">Fig. 2 I</figureCitation>
).
</paragraph>
<paragraph id="B60C9C4EFFC22D6AFF6412D8FD19FA0B" blockId="6.[151,1437,793,1468]" box="[199,658,1441,1468]" pageId="6" pageNumber="377">
<emphasis id="84C7405CFFC22D6AFF6412D8FE68FA0C" bold="true" box="[199,483,1441,1467]" pageId="6" pageNumber="377">Chinese common name.</emphasis>
乐东副华ü䗛
</paragraph>
</subSubSection>
</treatment>
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<heading id="A41C81E1FFA3FFC4C2BFFBDDF44CFB88" bold="true" box="[151,585,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="490" reason="1">
<emphasis id="CD9FEA9FFFA3FFC4C2BFFBDDF44CFB88" bold="true" box="[151,585,1086,1113]" pageId="1" pageNumber="490">
<taxonomicName id="38EB4D0EFFA3FFC4C2BFFBDDF7E1FB88" authority="Nakamura" authorityName="Nakamura" authorityYear="2024" box="[151,484,1086,1113]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="490" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA3FFC4C2BFFBDDF750FB88" bold="true" box="[151,341,1086,1113]" italics="true" pageId="1" pageNumber="490">Lycocerus todai</emphasis>
Nakamura
</taxonomicName>
<taxonomicNameLabel id="D6AC57E4FFA3FFC4C3C3FBDDF44CFB88" box="[491,585,1087,1113]" pageId="1" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
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</subSubSection>
<subSubSection id="B7F16506FFA3FFC4C2BFFB65F784FB70" box="[151,385,1159,1185]" pageId="1" pageNumber="490" type="description">
<paragraph id="FF54368DFFA3FFC4C2BFFB65F784FB70" blockId="1.[151,585,1086,1185]" box="[151,385,1159,1185]" pageId="1" pageNumber="490">
(
<figureCitation id="67D02A08FFA3FFC4C2B7FB65F72DFB70" box="[159,296,1159,1185]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–9" captionStartId-0="2.[151,264,1763,1787]" captionStartId-1="3.[151,264,1924,1948]" captionTargetBox-0="[175,1412,194,1726]" captionTargetBox-1="[172,1415,195,1888]" captionTargetId-0="figure-28@2.[162,1425,181,1739]" captionTargetId-1="figure-15@3.[159,1427,181,1901]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Habitus of Lycocerus spp., male. 1 &amp; 4–6, L. todai sp. nov., 1: holotype, dorsal; 4: paratype from Jana, Oppadale, ventral; 5: ditto, alive, dorsal; 6: ditto, alive, lateral; 2, L. okinawanus, dorsal; 3, L. yonaensis, dorsal. Scale bars: 3.0 mm." captionText-1="FIGURES 7–9.Aedeagus, male tergite IX and male sternite IX of Lycocerus spp. 7, L. todai sp. nov., paratype; 8, L. okinawanus; 9, L. yonaensis. A: aedeagus, ventral; B: ditto, lateral; C: ditto, dorsal; D: tergite IX and sternite IX, ventral. Scale bars: 0.5 mm." pageId="1" pageNumber="490">Figs. 1, 4–7</figureCitation>
,
<figureCitation id="67D02A08FFA3FFC4C31BFB65F77DFB70" box="[307,376,1159,1185]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="5.[151,250,1800,1824]" captionStartId-1="6.[151,250,1652,1676]" captionTargetBox-0="[151,1436,813,1777]" captionTargetBox-1="[169,1422,980,1627]" captionTargetId-0="figure-263@5.[151,1436,813,1777]" captionTargetId-1="figure-289@6.[166,1422,977,1627]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 10. Collecting site of Lycocerus todai sp. nov. (Nakijin-son, Jana, Oppadake), taken by the author when a paratype was collected." captionText-1="FIGURE 11. Distribution map of three black-colored Lycocerus species in the Ryukyu Islands, created based on the distributional information presented in this paper." pageId="1" pageNumber="490">10–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA3FFC4C2BFFB2AF49FFAA3" pageId="1" pageNumber="490" type="materials_examined">
<paragraph id="FF54368DFFA3FFC4C2BFFB2AF49FFAA3" blockId="1.[151,1437,1224,2043]" pageId="1" pageNumber="490">
<emphasis id="CD9FEA9FFFA3FFC4C2BFFB2AF755FB33" bold="true" box="[151,336,1224,1250]" pageId="1" pageNumber="490">Type materials.</emphasis>
<materialsCitation id="4F833CD0FFA3FFC4C37FFB2AF7CFFAFB" collectingDate="2019-02-15" collectionCode="KURA" collectorName="Naoki Toda" country="Japan" latitude="26.67047" location="Nakijin-son" longLatPrecision="1" longitude="127.96964" municipality="Tamashiro" pageId="1" pageNumber="490" specimenCount="1" typeStatus="holotype">
<typeStatus id="2050882FFFA3FFC4C37FFB2AF7C7FB33" box="[343,450,1224,1250]" pageId="1" pageNumber="490" type="holotype">Holotype</typeStatus>
, male, “[
<emphasis id="CD9FEA9FFFA3FFC4C000FB2BF4DDFB32" bold="true" box="[552,728,1225,1251]" pageId="1" pageNumber="490">ṗäŝ 乙ṟē</emphasis>
]
<emphasis id="CD9FEA9FFFA3FFC4C0C0FB2BF5E0FB32" bold="true" box="[744,997,1225,1251]" pageId="1" pageNumber="490">国DZō÷ṣ仁ťīĸ</emphasis>
26. 670470, 127. 969643,
<date id="8B55104DFFA3FFC4C73AFB2AF39DFB33" box="[1298,1432,1224,1251]" pageId="1" pageNumber="490" value="2019-02-15">
<collectingDate id="9B11E9A5FFA3FFC4C73AFB2AF39DFB33" box="[1298,1432,1224,1251]" pageId="1" pageNumber="490" value="2019-02-15">15. II. 2019</collectingDate>
</date>
, leg.
<emphasis id="CD9FEA9FFFA3FFC4C2E2FB0FF740FAD6" bold="true" box="[202,325,1261,1287]" pageId="1" pageNumber="490">DƂ 尚*</emphasis>
/
<collectingCountry id="87FC761DFFA3FFC4C348FB0EF7BCFAD7" box="[352,441,1260,1286]" name="Japan" pageId="1" pageNumber="490">JAPAN</collectingCountry>
: Okinawa-jima Is.,
<location id="FA346056FFA3FFC4C0B4FB0EF526FAD7" LSID="urn:lsid:plazi:treatment:7742879BFFA3FFC1C2BFFBDDF561FC74:FA346056FFA3FFC4C0B4FB0EF526FAD7" box="[668,803,1260,1286]" country="Japan" latitude="26.67047" longLatPrecision="1" longitude="127.96964" municipality="Tamashiro" name="Nakijin-son" pageId="1" pageNumber="490">Nakijin-son</location>
,
<collectingMunicipality id="1F30ACF7FFA3FFC4C118FB0EF5ACFAD7" box="[816,937,1260,1286]" pageId="1" pageNumber="490">Tamashiro</collectingMunicipality>
,
<geoCoordinate id="9ADF504AFFA3FFC4C19FFB0EF24BFAD7" box="[951,1102,1260,1286]" degrees="26. 670470" direction="north" orientation="latitude" pageId="1" pageNumber="490" precision="1" value="26.67047">26. 670470N</geoCoordinate>
,
<geoCoordinate id="9ADF504AFFA3FFC4C675FB0EF2FAFAD7" box="[1117,1279,1260,1286]" degrees="127. 969643" direction="east" orientation="longitude" pageId="1" pageNumber="490" precision="1" value="127.96964">127. 969643E</geoCoordinate>
,
<date id="8B55104DFFA3FFC4C725FB0EF39DFAD7" box="[1293,1432,1260,1287]" pageId="1" pageNumber="490" value="2019-02-15">
<collectingDate id="9B11E9A5FFA3FFC4C725FB0EF39DFAD7" box="[1293,1432,1260,1287]" pageId="1" pageNumber="490" value="2019-02-15">15. II. 2019</collectingDate>
</date>
,
<collectorName id="521E535BFFA3FFC4C2BFFAF2F719FAFB" box="[151,284,1296,1322]" pageId="1" pageNumber="490">Naoki Toda</collectorName>
leg.” (
<collectionCode id="99FAAE48FFA3FFC4C344FAF2F7C7FAFB" box="[364,450,1296,1322]" pageId="1" pageNumber="490">KURA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2050882FFFA3FFC4C3FEFAF3F446FAFA" box="[470,579,1297,1323]" pageId="1" pageNumber="490" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="E9EDFD04FFA3FFC4C07DFAF2F4B1FAFB" box="[597,692,1296,1322]" count="3" pageId="1" pageNumber="490" type="male">3 males</specimenCount>
):
<materialsCitation id="4F833CD0FFA3FFC4C0E0FAF2F2CDFAFB" box="[712,1224,1296,1323]" collectingDate="2019-02-15" collectionCode="KURA" collectorName="Naoki Toda" country="Japan" latitude="26.67047" location="Nakijin-son" longLatPrecision="1" longitude="127.96964" municipality="Tamashiro" pageId="1" pageNumber="490" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA3FFC4C0E0FAF2F526FAFB" box="[712,803,1296,1322]" count="2" pageId="1" pageNumber="490" type="male">2 males</specimenCount>
, same data as the holotype (
<collectionCode id="99FAAE48FFA3FFC4C643FAF2F2C4FAFB" box="[1131,1217,1296,1322]" pageId="1" pageNumber="490">KURA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA3FFC4C6FEFAF2F493FAA3" collectingDate="2024-02-15" collectionCode="FZUT" collectorName="Ryo Nakamura" country="Japan" county="Mt. Oppadake" elevation="260" latitude="26.6701" location="Nakijin-son" longLatPrecision="6" longitude="127.9622" municipality="Jana" pageId="1" pageNumber="490" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA3FFC4C6FEFAF2F323FAFB" box="[1238,1318,1296,1322]" count="1" pageId="1" pageNumber="490" type="male">1 male</specimenCount>
, “
<collectingCountry id="87FC761DFFA3FFC4C715FAF2F39CFAFB" box="[1341,1433,1296,1322]" name="Japan" pageId="1" pageNumber="490">JAPAN</collectingCountry>
: Okinawa-jima Is.,
<location id="FA346056FFA3FFC4C340FAD6F7EAFA9F" LSID="urn:lsid:plazi:treatment:7742879BFFA3FFC1C2BFFBDDF561FC74:FA346056FFA3FFC4C340FAD6F7EAFA9F" box="[360,495,1332,1358]" country="Japan" county="Mt. Oppadake" latitude="26.6701" longLatPrecision="6" longitude="127.9622" municipality="Jana" name="Nakijin-son" pageId="1" pageNumber="490">Nakijin-son</location>
,
<collectingMunicipality id="1F30ACF7FFA3FFC4C3D3FAD7F435FA9E" box="[507,560,1333,1359]" pageId="1" pageNumber="490">Jana</collectingMunicipality>
,
<collectingCounty id="16354E01FFA3FFC4C013FAD7F4DBFA9F" box="[571,734,1332,1359]" pageId="1" pageNumber="490">Mt. Oppadake</collectingCounty>
, alt.
<quantity id="38139B68FFA3FFC4C13CFAD6F55BFA9E" box="[788,862,1332,1359]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="490" unit="m" value="260.0">
<elevation id="74C6D1BEFFA3FFC4C13CFAD6F55BFA9E" box="[788,862,1332,1359]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="490" unit="m" value="260.0">260 m</elevation>
</quantity>
,
<geoCoordinate id="9ADF504AFFA3FFC4C142FAD6F5D9FA9F" box="[874,988,1332,1358]" degrees="26.6701" direction="north" orientation="latitude" pageId="1" pageNumber="490" precision="5" value="26.6701">26.6701N</geoCoordinate>
,
<geoCoordinate id="9ADF504AFFA3FFC4C1CFFAD6F261FA9F" box="[999,1124,1332,1358]" degrees="127.9622" direction="east" orientation="longitude" pageId="1" pageNumber="490" precision="5" value="127.9622">127.9622E</geoCoordinate>
,
<date id="8B55104DFFA3FFC4C658FAD6F2E2FA9E" box="[1136,1255,1332,1359]" pageId="1" pageNumber="490" value="2024-02-15">
<collectingDate id="9B11E9A5FFA3FFC4C658FAD6F2E2FA9E" box="[1136,1255,1332,1359]" pageId="1" pageNumber="490" value="2024-02-15">15.II.2024</collectingDate>
</date>
,
<collectorName id="521E535BFFA3FFC4C6DBFAD7F398FA9F" box="[1267,1437,1332,1359]" pageId="1" pageNumber="490">Ryo Nakamura</collectorName>
leg.,
<emphasis id="CD9FEA9FFFA3FFC4C2E6FABBF423FAA2" bold="true" box="[206,550,1369,1395]" pageId="1" pageNumber="490">ṗäŝ÷ṣ仁ťā名 乙ṟē</emphasis>
” (
<collectionCode id="99FAAE48FFA3FFC4C069FABAF488FAA3" box="[577,653,1368,1394]" pageId="1" pageNumber="490">FZUT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA3FFC1C2EFFA9EF79BFE58" lastPageId="4" lastPageNumber="493" pageId="1" pageNumber="490" type="description">
<paragraph id="FF54368DFFA3FFC4C2EFFA9EF509FA0F" blockId="1.[151,1437,1224,2043]" pageId="1" pageNumber="490">
<emphasis id="CD9FEA9FFFA3FFC4C2EFFA9EF75CFA47" bold="true" box="[199,345,1404,1430]" pageId="1" pageNumber="490">Description.</emphasis>
Male. BL
<quantity id="38139B68FFA3FFC4C3FBFA9EF472FA46" box="[467,631,1404,1431]" metricMagnitude="-3" metricUnit="m" metricValue="9.62" metricValueMax="10.1" metricValueMin="9.14" pageId="1" pageNumber="490" unit="mm" value="9.62" valueMax="10.1" valueMin="9.14">9.14–10.1 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C0AEFA9EF4E9FA46" box="[646,748,1404,1431]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="490" unit="mm" value="10.1">10.1 mm</quantity>
in the
<typeStatus id="2050882FFFA3FFC4C112FA9EF599FA47" box="[826,924,1404,1430]" pageId="1" pageNumber="490" type="holotype">holotype</typeStatus>
= HT). Body almost entirely black but claws reddish brown and abdomen yellowish orange with a pair of small dark brown maculae on basal five segments. Body closely covered with yellowish white pubescence.
</paragraph>
<paragraph id="FF54368DFFA3FFC4C2EFFA0AF52FF942" blockId="1.[151,1437,1224,2043]" pageId="1" pageNumber="490">
Head shorter than width, dorsum slightly depressed between antennal sockets, HW
<quantity id="38139B68FFA3FFC4C642FA0AF308F9D2" box="[1130,1293,1512,1539]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="1" pageNumber="490" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.60–1.80 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C733FA0AF385F9D2" box="[1307,1408,1512,1539]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="490" unit="mm" value="1.8">1.80 mm</quantity>
in HT); eyes large, globular and prominent, interocular distance 2.69 times as wide as the diameter of an eye; mandible arcuate and pointed at apex; apical segments of labial and maxillary palpomere knife-shaped; antennae filiform, without any grooves, reaching 3/5 of elytra, comparative lengths of antennomeres of the
<typeStatus id="2050882FFFA3FFC4C65CF9B6F2D3F9BF" box="[1140,1238,1620,1646]" pageId="1" pageNumber="490" type="holotype">holotype</typeStatus>
as follows: 2.08: 1: 2.06: 2.54: 2.67: 2.69: 2.73: 2.67: 2.63: 2.31: 2.75.
</paragraph>
<paragraph id="FF54368DFFA3FFC4C2EFF97FF35DF863" blockId="1.[151,1437,1224,2043]" pageId="1" pageNumber="490">
Pronotum rectangular, widest at posterior angles, anterior margin arcuate, posterior margin feebly arcuate, lateral margins almost parallel but slightly emarginated in the middle, anterior angles obtuse and rounded, posterior angles nearly right-angled and slightly protruding laterally; PL
<quantity id="38139B68FFA3FFC4C17FF906F5FCF92E" box="[855,1017,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="1.425" metricValueMax="1.54" metricValueMin="1.31" pageId="1" pageNumber="490" unit="mm" value="1.425" valueMax="1.54" valueMin="1.31">1.31–1.54 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C62FF906F269F92E" box="[1031,1132,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="490" unit="mm" value="1.54">1.54 mm</quantity>
in HT), PW
<quantity id="38139B68FFA3FFC4C6D2F906F398F92E" box="[1274,1437,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.41" metricValueMin="1.2" pageId="1" pageNumber="490" unit="mm" value="1.305" valueMax="1.41" valueMin="1.2">1.20–1.41 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C288F8EAF701F8F2" box="[160,260,1800,1827]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="490" unit="mm" value="1.41">1.41 mm</quantity>
in HT), HW/PW 1.28–1.34 (
<quantity id="38139B68FFA3FFC4C061F8EAF49DF8F3" box="[585,664,1800,1826]" metricMagnitude="-2" metricUnit="m" metricValue="3.2512" pageId="1" pageNumber="490" unit="in" value="1.28">1.28 in</quantity>
HT), PL/PW 1.07–1.09 (
<quantity id="38139B68FFA3FFC4C19BF8EAF207F8F3" box="[947,1026,1800,1826]" metricMagnitude="-2" metricUnit="m" metricValue="2.7686" pageId="1" pageNumber="490" unit="in" value="1.09">1.09 in</quantity>
HT); disc with a medio-longitudinal furrow not reaching the anterior and posterior margins, moderately convex and gradually flattened in lateral halves. Scutellum triangular with acute and round apex. Elytra long, parallel in lateral sides, apex normally rounded, EL
<quantity id="38139B68FFA3FFC4C2BFF896F739F85E" box="[151,316,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="7.025" metricValueMax="7.27" metricValueMin="6.78" pageId="1" pageNumber="490" unit="mm" value="7.025" valueMax="7.27" valueMin="6.78">6.78–7.27 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C363F896F7B6F85E" box="[331,435,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="7.27" pageId="1" pageNumber="490" unit="mm" value="7.27">7.27 mm</quantity>
in HT), EW
<quantity id="38139B68FFA3FFC4C062F896F4EAF85E" box="[586,751,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="2.025" metricValueMax="2.17" metricValueMin="1.88" pageId="1" pageNumber="490" unit="mm" value="2.025" valueMax="2.17" valueMin="1.88">1.88–2.17 mm</quantity>
(
<quantity id="38139B68FFA3FFC4C0D7F896F563F85E" box="[767,870,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="1" pageNumber="490" unit="mm" value="2.17">2.17 mm</quantity>
in HT), EW/PW 1.52–1.57 (
<quantity id="38139B68FFA3FFC4C698F896F304F85F" box="[1200,1281,1908,1935]" metricMagnitude="-2" metricUnit="m" metricValue="3.9116" pageId="1" pageNumber="490" unit="in" value="1.54">1.54 in</quantity>
HT), EL/EW 3.35–3.61 (
<quantity id="38139B68FFA3FFC4C33EF87AF763F863" box="[278,358,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="8.509" pageId="1" pageNumber="490" unit="in" value="3.35">3.35 in</quantity>
HT). Legs slender, each femur mostly straight, each tibia feebly arcuate, all claws simple.
</paragraph>
<paragraph id="FF54368DFFA3FFC4C2EFF85EF368F82B" blockId="1.[151,1437,1224,2043]" pageId="1" pageNumber="490">Male sternite IX (fig. 7D) cymbiform, as long as tergite IX, 3.43 times longer than wide, pubescence in apical 1/3, lateral margins parallel in the middle, feebly narrowing in apical 1/4 to form a round and slightly bifid tip.</paragraph>
<caption id="AB946605FFA0FFC7C2BFF901F6C3F892" pageId="2" pageNumber="491" startId="2.[151,264,1763,1787]" targetBox="[175,1412,194,1726]" targetPageId="2" targetType="figure">
<paragraph id="FF54368DFFA0FFC7C2BFF901F6C3F892" blockId="2.[151,1437,1763,1859]" pageId="2" pageNumber="491">
<emphasis id="CD9FEA9FFFA0FFC7C2BFF901F73BF92A" bold="true" box="[151,318,1763,1787]" pageId="2" pageNumber="491">FIGURES 1–6.</emphasis>
Habitus of
<taxonomicName id="38EB4D0EFFA0FFC7C3E8F906F45BF92A" box="[448,606,1764,1787]" class="Insecta" family="Cantharidae" genus="Lycocerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="491" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="CD9FEA9FFFA0FFC7C3E8F906F422F92A" box="[448,551,1764,1787]" italics="true" pageId="2" pageNumber="491">Lycocerus</emphasis>
spp.
</taxonomicName>
, male. 1 &amp; 4–6,
<taxonomicName id="38EB4D0EFFA0FFC7C130F906F56FF92A" authorityName="Nakamura" authorityYear="2024" box="[792,874,1763,1787]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="491" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA0FFC7C130F906F56FF92A" box="[792,874,1763,1787]" italics="true" pageId="2" pageNumber="491">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA0FFC7C15CF906F5CDF92D" bold="true" box="[884,968,1764,1788]" pageId="2" pageNumber="491">
<taxonomicNameLabel id="D6AC57E4FFA0FFC7C15CF906F5CDF92D" box="[884,968,1764,1788]" pageId="2" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 1: holotype, dorsal; 4: paratype from Jana, Oppadale, ventral; 5: ditto, alive, dorsal; 6: ditto, alive, lateral; 2,
<taxonomicName id="38EB4D0EFFA0FFC7C106F8EAF5C7F8CE" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[814,962,1799,1823]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="491" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA0FFC7C106F8EAF5C7F8CE" box="[814,962,1799,1823]" italics="true" pageId="2" pageNumber="491">L. okinawanus</emphasis>
</taxonomicName>
, dorsal; 3,
<taxonomicName id="38EB4D0EFFA0FFC7C61BF8EAF2B4F8CE" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[1075,1201,1800,1823]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="491" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA0FFC7C61BF8EAF2B4F8CE" box="[1075,1201,1800,1823]" italics="true" pageId="2" pageNumber="491">L. yonaensis</emphasis>
</taxonomicName>
, dorsal. Scale bars: 3.0 mm.
</paragraph>
</caption>
<caption id="AB946605FFA1FFC6C2BFF866F6C3F835" pageId="3" pageNumber="492" startId="3.[151,264,1924,1948]" targetBox="[172,1415,195,1888]" targetPageId="3" targetType="figure">
<paragraph id="FF54368DFFA1FFC6C2BFF866F6C3F835" blockId="3.[151,1437,1924,2020]" pageId="3" pageNumber="492">
<emphasis id="CD9FEA9FFFA1FFC6C2BFF866F73CF84C" bold="true" box="[151,313,1924,1949]" pageId="3" pageNumber="492">FIGURES 7–9.</emphasis>
Aedeagus, male tergite IX and male sternite IX of
<taxonomicName id="38EB4D0EFFA1FFC6C104F867F5D1F84D" box="[812,980,1925,1948]" class="Insecta" family="Cantharidae" genus="Lycocerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="undefined-7">
<emphasis id="CD9FEA9FFFA1FFC6C104F867F596F84D" box="[812,915,1925,1948]" italics="true" pageId="3" pageNumber="492">Lycocerus</emphasis>
spp. 7
</taxonomicName>
,
<taxonomicName id="38EB4D0EFFA1FFC6C1F5F867F22FF84C" authorityName="Nakamura" authorityYear="2024" box="[989,1066,1925,1949]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA1FFC6C1F5F867F22FF84C" box="[989,1066,1925,1949]" italics="true" pageId="3" pageNumber="492">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA1FFC6C607F867F278F84C" bold="true" box="[1071,1149,1925,1949]" pageId="3" pageNumber="492">
<taxonomicNameLabel id="D6AC57E4FFA1FFC6C607F867F278F84C" box="[1071,1149,1925,1949]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype; 8,
<taxonomicName id="38EB4D0EFFA1FFC6C72BF867F390F84C" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[1283,1429,1925,1949]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA1FFC6C72BF867F390F84C" box="[1283,1429,1925,1949]" italics="true" pageId="3" pageNumber="492">L. okinawanus</emphasis>
</taxonomicName>
; 9,
<taxonomicName id="38EB4D0EFFA1FFC6C29AF84BF736F811" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[178,307,1961,1984]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA1FFC6C29AF84BF736F811" box="[178,307,1961,1984]" italics="true" pageId="3" pageNumber="492">L. yonaensis</emphasis>
</taxonomicName>
. A: aedeagus, ventral; B: ditto, lateral; C: ditto, dorsal; D: tergite IX and sternite IX, ventral. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="FF54368DFFA6FFC1C2EFFF75F7D4FEB4" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="493">Aedeagus (figs. 7A–C) without hairs; ventral process of each paramere slender, feebly bent inwards and slightly expanded at apex in ventral view, almost straight and gradually expanded toward apex in lateral view; dorsal plates of parameres broad with apical margins arcuate, lateral margins approaching and inner margins almost parallel each other in dorsal view, as long as ventral process in lateral view; laterophyse almost straight and thin in ventral view, slightly not reaching the same level as the tip of ventral process and dorsal plates, bent dorsally toward apex of dorsal plates in lateral view.</paragraph>
<paragraph id="FF54368DFFA6FFC1C2EFFE8DF79BFE58" blockId="4.[151,1437,151,933]" box="[199,414,367,393]" pageId="4" pageNumber="493">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2EFFE70F545FE7C" box="[199,832,402,429]" pageId="4" pageNumber="493" type="distribution">
<paragraph id="FF54368DFFA6FFC1C2EFFE70F545FE7C" blockId="4.[151,1437,151,933]" box="[199,832,402,429]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFE70F764FE7D" bold="true" box="[199,353,402,428]" pageId="4" pageNumber="493">Distribution.</emphasis>
<collectingCountry id="87FC761DFFA6FFC1C34FFE71F7A2FE7C" box="[359,423,403,429]" name="Japan" pageId="4" pageNumber="493">Japan</collectingCountry>
(Ryukyu:
<collectingRegion id="3D2FF86FFFA6FFC1C036FE71F486FE7C" box="[542,643,403,429]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa</collectingRegion>
Island) (fig. 11).
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2EFFE54F213FE24" pageId="4" pageNumber="493" type="biology_ecology">
<paragraph id="FF54368DFFA6FFC1C2EFFE54F213FE24" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFE54F722FE01" bold="true" box="[199,295,438,464]" pageId="4" pageNumber="493">Biology.</emphasis>
<specimenCount id="E9EDFD04FFA6FFC1C318FE55F79AFE00" box="[304,415,439,465]" count="1" pageId="4" pageNumber="493" type="male">One male</specimenCount>
<typeStatus id="2050882FFFA6FFC1C380FE55F40DFE00" box="[424,520,439,465]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
from Jana, Oppadake was collected by sweeping green leaves in the evergreen broad-leaved forest at noon (fig. 10). Adults are collected only in mid-February.
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2EFFE1DF711FDEC" pageId="4" pageNumber="493" type="etymology">
<paragraph id="FF54368DFFA6FFC1C2EFFE1DF711FDEC" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFE1DF74BFDC8" bold="true" box="[199,334,511,537]" pageId="4" pageNumber="493">Etymology.</emphasis>
The specific name is dedicated to Mr. Naoki Toda, a Japanese colleopterist who collected the
<typeStatus id="2050882FFFA6FFC1C744FE1DF399FDC8" box="[1388,1436,511,537]" pageId="4" pageNumber="493">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2EFFDA4F561FC74" pageId="4" pageNumber="493" type="diagnosis">
<paragraph id="FF54368DFFA6FFC1C2EFFDA4F561FC74" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFDA4F7D4FDB1" bold="true" box="[199,465,582,608]" pageId="4" pageNumber="493">Differential diagnosis.</emphasis>
This new species closely resembles to
<taxonomicName id="38EB4D0EFFA6FFC1C1B5FDA5F34CFDB0" authority="(Ishida, 1985)" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[925,1353,582,609]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C1B5FDA5F29AFDB0" box="[925,1183,583,609]" italics="true" pageId="4" pageNumber="493">Lycocerus okinawanus</emphasis>
(Ishida, 1985)
</taxonomicName>
and
<taxonomicName id="38EB4D0EFFA6FFC1C7AEFDA5F42CFD54" authority="(Sato et Okushima, 1992)" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C7AEFDA5F700FD55" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
(
<bibRefCitation id="9B7A4B7CFFA6FFC1C33BFD89F425FD54" author="Sato, M. &amp; Okushima, Y." box="[275,544,619,645]" pageId="4" pageNumber="493" pagination="857 - 865" refId="ref5090" refString="Sato, M. &amp; Okushima, Y. (1992) The genus Athemus (Coleoptera, Cantharidae) of the Ryukyu Islands, southwest Japan. Japanese Journal of Entomology, 60, 857 - 865." type="journal article" year="1992">Satô et Okushima, 1992</bibRefCitation>
)
</taxonomicName>
from the same island but differs in the following characters (see also
<bibRefCitation id="9B7A4B7CFFA6FFC1C701FD89F6EAFD78" author="Okushima, Y." pageId="4" pageNumber="493" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima (2005))</bibRefCitation>
.
<emphasis id="CD9FEA9FFFA6FFC1C2D2FD6DF768FD79" bold="true" box="[250,365,654,681]" pageId="4" pageNumber="493">Body size</emphasis>
: larger in
<taxonomicName id="38EB4D0EFFA6FFC1C3CBFD6DF439FD78" authorityName="Nakamura" authorityYear="2024" box="[483,572,655,681]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C3CBFD6DF439FD78" box="[483,572,655,681]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C06BFD6DF49BFD78" bold="true" box="[579,670,655,681]" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C06BFD6DF49BFD78" box="[579,670,655,681]" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="38EB4D0EFFA6FFC1C0FFFD6DF566FD79" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[727,867,655,680]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C0FFFD6DF566FD79" box="[727,867,655,680]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
(more than
<quantity id="38139B68FFA6FFC1C1C7FD6DF242FD78" box="[1007,1095,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="493" unit="mm" value="9.1">9.1 mm</quantity>
) vs. smaller in
<taxonomicName id="38EB4D0EFFA6FFC1C6D0FD6DF399FD78" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[1272,1436,655,681]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C6D0FD6DF399FD78" box="[1272,1436,655,681]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
(less than
<quantity id="38139B68FFA6FFC1C323FD51F763FD1C" box="[267,358,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="493" unit="mm" value="8.6">8.6 mm</quantity>
);
<emphasis id="CD9FEA9FFFA6FFC1C353FD51F420FD1C" bold="true" box="[379,549,690,717]" pageId="4" pageNumber="493">color of elytra</emphasis>
: dull black in
<taxonomicName id="38EB4D0EFFA6FFC1C0E3FD51F521FD1C" authorityName="Nakamura" authorityYear="2024" box="[715,804,691,717]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C0E3FD51F521FD1C" box="[715,804,691,717]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C104FD51F582FD1C" bold="true" box="[812,903,691,717]" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C104FD51F582FD1C" box="[812,903,691,717]" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="38EB4D0EFFA6FFC1C197FD51F249FD1D" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[959,1100,691,716]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C197FD51F249FD1D" box="[959,1100,691,716]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
vs. glossy black with blueish lustre in
<taxonomicName id="38EB4D0EFFA6FFC1C2D7FD35F7A0FD20" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[255,421,727,753]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2D7FD35F7A0FD20" box="[255,421,727,753]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
;
<emphasis id="CD9FEA9FFFA6FFC1C39FFD35F489FD20" bold="true" box="[439,652,726,753]" pageId="4" pageNumber="493">color of abdomen</emphasis>
: almost entirely yellowish orange in
<taxonomicName id="38EB4D0EFFA6FFC1C616FD35F29CFD20" authorityName="Nakamura" authorityYear="2024" box="[1086,1177,727,753]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C616FD35F29CFD20" box="[1086,1177,727,753]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C68AFD35F2FBFD20" bold="true" box="[1186,1278,727,753]" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C68AFD35F2FBFD20" box="[1186,1278,727,753]" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. black but marginally yellowish-brown in
<taxonomicName id="38EB4D0EFFA6FFC1C3DFFD19F49FFCC4" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[503,666,763,789]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C3DFFD19F49FFCC4" box="[503,666,763,789]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
vs. entirely black in
<taxonomicName id="38EB4D0EFFA6FFC1C1ABFD19F20BFCC5" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[899,1038,763,788]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C1ABFD19F20BFCC5" box="[899,1038,763,788]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
;
<emphasis id="CD9FEA9FFFA6FFC1C635FD18F298FCC4" bold="true" box="[1053,1181,762,789]" pageId="4" pageNumber="493">sternite IX</emphasis>
: narrow in
<taxonomicName id="38EB4D0EFFA6FFC1C735FD19F370FCC4" authorityName="Nakamura" authorityYear="2024" box="[1309,1397,763,789]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C735FD19F370FCC4" box="[1309,1397,763,789]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C753FD19F6CDFCE8" bold="true" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C753FD19F6CDFCE8" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(more than 3.4 times longer than wide) vs. wide in
<taxonomicName id="38EB4D0EFFA6FFC1C122FCFDF5A8FCE8" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[778,941,799,825]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C122FCFDF5A8FCE8" box="[778,941,799,825]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
and
<taxonomicName id="38EB4D0EFFA6FFC1C1CBFCFDF26AFCE9" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[995,1135,799,824]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C1CBFCFDF26AFCE9" box="[995,1135,799,824]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
(less than 2.8 times longer than wide);
<emphasis id="CD9FEA9FFFA6FFC1C333FCA1F465FC8C" bold="true" box="[283,608,834,861]" pageId="4" pageNumber="493">ventral process of aedeagus</emphasis>
: stick-like in
<taxonomicName id="38EB4D0EFFA6FFC1C0D0FCA1F54AFC8C" authorityName="Nakamura" authorityYear="2024" box="[760,847,835,861]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C0D0FCA1F54AFC8C" box="[760,847,835,861]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C17DFCA1F5ABFC8C" bold="true" box="[853,942,835,861]" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C17DFCA1F5ABFC8C" box="[853,942,835,861]" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="38EB4D0EFFA6FFC1C1CBFCA1F280FC8C" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[995,1157,835,861]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C1CBFCA1F280FC8C" box="[995,1157,835,861]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
vs. broad and angular on dorsal side in
<taxonomicName id="38EB4D0EFFA6FFC1C31CFC85F7C5FC51" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[308,448,871,896]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C31CFC85F7C5FC51" box="[308,448,871,896]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
;
<emphasis id="CD9FEA9FFFA6FFC1C3E7FC85F4E9FC50" bold="true" box="[463,748,870,897]" pageId="4" pageNumber="493">laterophyse of aedeagus</emphasis>
: thin in ventral view in
<taxonomicName id="38EB4D0EFFA6FFC1C1D2FC85F257FC50" authorityName="Nakamura" authorityYear="2024" box="[1018,1106,871,897]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="todai" status="sp. nov.">
<emphasis id="CD9FEA9FFFA6FFC1C1D2FC85F257FC50" box="[1018,1106,871,897]" italics="true" pageId="4" pageNumber="493">L. todai</emphasis>
</taxonomicName>
<emphasis id="CD9FEA9FFFA6FFC1C670FC85F2B7FC50" bold="true" box="[1112,1202,871,897]" pageId="4" pageNumber="493">
<taxonomicNameLabel id="D6AC57E4FFA6FFC1C670FC85F2B7FC50" box="[1112,1202,871,897]" pageId="4" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="38EB4D0EFFA6FFC1C6C1FC85F370FC51" baseAuthorityName="M. Sato et Okushima" baseAuthorityYear="1992" box="[1257,1397,871,896]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="yonaensis">
<emphasis id="CD9FEA9FFFA6FFC1C6C1FC85F370FC51" box="[1257,1397,871,896]" italics="true" pageId="4" pageNumber="493">L. yonaensis</emphasis>
</taxonomicName>
vs. stout and widend toward apex in ventral view in
<taxonomicName id="38EB4D0EFFA6FFC1C092FC69F558FC74" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[698,861,907,933]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C092FC69F558FC74" box="[698,861,907,933]" italics="true" pageId="4" pageNumber="493">L. okinawanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="7742879BFFA6FFC0C2BFFC15F6ECFCC4" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4" httpUri="http://treatment.plazi.org/id/7742879BFFA6FFC0C2BFFC15F6ECFCC4" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
<subSubSection id="B7F16506FFA6FFC1C2BFFC15F45BFBC1" box="[151,606,1014,1041]" pageId="4" pageNumber="493" type="nomenclature">
<paragraph id="FF54368DFFA6FFC1C2BFFC15F45BFBC1" blockId="4.[151,606,1014,1077]" box="[151,606,1014,1041]" pageId="4" pageNumber="493">
<heading id="A41C81E1FFA6FFC1C2BFFC15F45BFBC1" bold="true" box="[151,606,1014,1041]" fontSize="11" level="1" pageId="4" pageNumber="493" reason="1">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFC15F45BFBC1" authority="(Ishida, 1985)" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[151,606,1014,1041]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFC15F45BFBC1" bold="true" box="[151,606,1014,1041]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFC15F7A9FBC0" bold="true" box="[151,428,1014,1041]" italics="true" pageId="4" pageNumber="493">Lycocerus okinawanus</emphasis>
(Ishida, 1985)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2BFFBF9F73EFBE4" box="[151,315,1051,1077]" pageId="4" pageNumber="493" type="description">
<paragraph id="FF54368DFFA6FFC1C2BFFBF9F73EFBE4" blockId="4.[151,606,1014,1077]" box="[151,315,1051,1077]" pageId="4" pageNumber="493">
(
<figureCitation id="67D02A08FFA6FFC1C2B7FBF9F6EAFBE4" box="[159,239,1051,1077]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1763,1787]" captionTargetBox="[175,1412,194,1726]" captionTargetId="figure-28@2.[162,1425,181,1739]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Lycocerus spp., male. 1 &amp; 4–6, L. todai sp. nov., 1: holotype, dorsal; 4: paratype from Jana, Oppadale, ventral; 5: ditto, alive, dorsal; 6: ditto, alive, lateral; 2, L. okinawanus, dorsal; 3, L. yonaensis, dorsal. Scale bars: 3.0 mm." pageId="4" pageNumber="493">Figs. 2</figureCitation>
,
<figureCitation id="67D02A08FFA6FFC1C2D3FBF9F70FFBE4" box="[251,266,1051,1077]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1924,1948]" captionTargetBox="[172,1415,195,1888]" captionTargetId="figure-15@3.[159,1427,181,1901]" captionTargetPageId="3" captionText="FIGURES 7–9.Aedeagus, male tergite IX and male sternite IX of Lycocerus spp. 7, L. todai sp. nov., paratype; 8, L. okinawanus; 9, L. yonaensis. A: aedeagus, ventral; B: ditto, lateral; C: ditto, dorsal; D: tergite IX and sternite IX, ventral. Scale bars: 0.5 mm." pageId="4" pageNumber="493">8</figureCitation>
,
<figureCitation id="67D02A08FFA6FFC1C33FFBF9F737FBE4" box="[279,306,1051,1077]" captionStart="FIGURE 11" captionStartId="6.[151,250,1652,1676]" captionTargetBox="[169,1422,980,1627]" captionTargetId="figure-289@6.[166,1422,977,1627]" captionTargetPageId="6" captionText="FIGURE 11. Distribution map of three black-colored Lycocerus species in the Ryukyu Islands, created based on the distributional information presented in this paper." pageId="4" pageNumber="493">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC1C2BFFBBDF335FB02" pageId="4" pageNumber="493" type="reference_group">
<paragraph id="FF54368DFFA6FFC1C2BFFBBDF238FBA6" blockId="4.[151,1436,1119,1235]" box="[151,1085,1119,1143]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFBBDF238FBA6" box="[151,1085,1119,1143]" pageId="4" pageNumber="493">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFBBDF7B7FBA6" authority="Ishida" authorityName="Ishida" box="[151,434,1119,1143]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFBBDF76BFBA6" box="[151,366,1119,1143]" italics="true" pageId="4" pageNumber="493">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
in
<treatmentCitation id="7E4A109CFFA6FFC1C3FAFBBDF476FBA6" author="Sato, M." box="[466,627,1119,1143]" page="113" pageId="4" pageNumber="493" year="1985">
<bibRefCitation id="9B7A4B7CFFA6FFC1C3FAFBBDF476FBA6" author="Sato, M." box="[466,627,1119,1143]" pageId="4" pageNumber="493" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô, 1985: 113</bibRefCitation>
</treatmentCitation>
, pl. 18, fig. 23 (Okinawa-hontô Is., Ryukyu).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FF54368DFFA6FFC1C2BFFB9CF79EFB64" blockId="4.[151,1436,1119,1235]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFB9CF79EFB64" pageId="4" pageNumber="493">
<treatmentCitation id="7E4A109CFFA6FFC1C2BFFB9CF433FB47" author="Ishida, K." box="[151,566,1150,1174]" page="196" pageId="4" pageNumber="493" year="1986">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFB9CF433FB47" authority="Ishida, 1986: 196" authorityName="Ishida" authorityPageNumber="196" authorityYear="1986" box="[151,566,1150,1174]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFB9CF777FB47" box="[151,370,1150,1174]" italics="true" pageId="4" pageNumber="493">Athemus okinawanus</emphasis>
<bibRefCitation id="9B7A4B7CFFA6FFC1C354FB9CF433FB47" author="Ishida, K." box="[380,566,1150,1174]" pageId="4" pageNumber="493" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida, 1986: 196</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 4–7 (Ginama-mura, Kunigami,
<collectingRegion id="3D2FF86FFFA6FFC1C1EEFB9CF224FB47" box="[966,1057,1150,1174]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa</collectingRegion>
Is.);
<treatmentCitation id="7E4A109CFFA6FFC1C676FB9CF39DFB47" author="Sato, M. &amp; Okushima, Y." box="[1118,1432,1150,1174]" page="858" pageId="4" pageNumber="493" year="1992">
<bibRefCitation id="9B7A4B7CFFA6FFC1C676FB9CF39DFB47" author="Sato, M. &amp; Okushima, Y." box="[1118,1432,1150,1174]" pageId="4" pageNumber="493" pagination="857 - 865" refId="ref5090" refString="Sato, M. &amp; Okushima, Y. (1992) The genus Athemus (Coleoptera, Cantharidae) of the Ryukyu Islands, southwest Japan. Japanese Journal of Entomology, 60, 857 - 865." type="journal article" year="1992">Satô &amp; Okushima, 1992: 858</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="7E4A109CFFA6FFC1C2EFFB7EF793FB64" author="Takahashi, K." box="[199,406,1180,1205]" page="40" pageId="4" pageNumber="493" year="1998">
<bibRefCitation id="9B7A4B7CFFA6FFC1C2EFFB7EF793FB64" author="Takahashi, K." box="[199,406,1180,1205]" pageId="4" pageNumber="493" pagination="29 - 48" refId="ref5129" refString="Takahashi, K. (1998) Check-list of Cantharidae of Japan. Kanagawa-Chuho, 122, 29 - 48. [in Japanese, with English title]" type="journal article" year="1998">Takahashi, 1998: 40</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FF54368DFFA6FFC1C2BFFB5EF335FB02" blockId="4.[151,1436,1119,1235]" box="[151,1328,1211,1235]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFB5EF335FB02" box="[151,1328,1211,1235]" pageId="4" pageNumber="493">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFB5EF413FB02" authority="(Ishida, 1986)" baseAuthorityName="Ishida" baseAuthorityYear="1986" box="[151,534,1211,1235]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFB5EF77BFB02" box="[151,382,1211,1235]" italics="true" pageId="4" pageNumber="493">Lycocerus okinawanus</emphasis>
(
<bibRefCitation id="9B7A4B7CFFA6FFC1C3A3FB59F40BFB02" author="Ishida, K." box="[395,526,1211,1235]" pageId="4" pageNumber="493" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida, 1986</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="7E4A109CFFA6FFC1C008FB59F4F8FB02" author="Okushima, Y." box="[544,765,1211,1235]" page="262" pageId="4" pageNumber="493" year="2005">
<bibRefCitation id="9B7A4B7CFFA6FFC1C008FB59F4F8FB02" author="Okushima, Y." box="[544,765,1211,1235]" pageId="4" pageNumber="493" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005: 262</bibRefCitation>
</treatmentCitation>
, figs. 65, 136, 179;
<treatmentCitation id="7E4A109CFFA6FFC1C1EEFB5EF329FB02" author="Kazantsev, S. &amp; Brancucci, M." box="[966,1324,1211,1235]" page="252" pageId="4" pageNumber="493" year="2007">
<bibRefCitation id="9B7A4B7CFFA6FFC1C1EEFB5EF329FB02" author="Kazantsev, S. &amp; Brancucci, M." box="[966,1324,1211,1235]" pageId="4" pageNumber="493" refId="ref4627" refString="Kazantsev, S. &amp; Brancucci, M. (2007) Cantharidae. In Lo ¨ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 234 - 298." type="book" year="2007">Kazantsev &amp; Brancucci, 2007: 252</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA6FFC0C2BFFAE3F7ECFE58" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493" type="materials_examined">
<paragraph id="FF54368DFFA6FFC1C2BFFAE3F6FEFA56" blockId="4.[151,1437,1281,2028]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFAE3F7CCFACA" bold="true" box="[151,457,1281,1307]" pageId="4" pageNumber="493">Type materials examined.</emphasis>
<materialsCitation id="4F833CD0FFA6FFC1C3F8FAE3F6F3FAEE" collectingDate="1978-04-01" collectionCode="EUMJ" collectorName="N. Kanie" location="Kunigami" municipality="Ginama Vil." pageId="4" pageNumber="493" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount id="E9EDFD04FFA6FFC1C3F8FAE3F41FFACA" box="[464,538,1281,1307]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C001FAE3F494FACA" box="[553,657,1281,1307]" pageId="4" pageNumber="493" type="holotype">holotype</typeStatus>
), “
<collectingMunicipality id="1F30ACF7FFA6FFC1C086FAE3F53EFACA" box="[686,827,1281,1307]" pageId="4" pageNumber="493">Ginama Vil.</collectingMunicipality>
,
<location id="FA346056FFA6FFC1C16EFAE3F5BCFACA" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4:FA346056FFA6FFC1C16EFAE3F5BCFACA" box="[838,953,1281,1307]" municipality="Ginama Vil." name="Kunigami" pageId="4" pageNumber="493" stateProvince="Okinawa">Kunigami</location>
,
<collectingRegion id="3D2FF86FFFA6FFC1C1EBFAE3F26DFACA" box="[963,1128,1281,1307]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa Pref.</collectingRegion>
,
<date id="8B55104DFFA6FFC1C65AFAE3F2F4FACA" box="[1138,1265,1281,1308]" pageId="4" pageNumber="493" value="1978-04-01">
<collectingDate id="9B11E9A5FFA6FFC1C65AFAE3F2F4FACA" box="[1138,1265,1281,1308]" pageId="4" pageNumber="493" value="1978-04-01">1. IV. 1978</collectingDate>
</date>
,
<collectorName id="521E535BFFA6FFC1C6D3FAE0F365FACA" box="[1275,1376,1281,1308]" pageId="4" pageNumber="493">N. Kanie</collectorName>
leg.” (
<collectionCode id="99FAAE48FFA6FFC1C2B7FAC7F6EBFAEE" box="[159,238,1317,1343]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA6FFC1C32AFAC7F4BAFAEE" box="[258,703,1317,1344]" collectingDate="1978-04-01" collectionCode="EUMJ" collectorName="N. Kanie" location="Kunigami" municipality="Ginama Vil." pageId="4" pageNumber="493" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA6FFC1C32AFAC7F767FAEE" box="[258,354,1317,1343]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C359FAC7F7D2FAEE" box="[369,471,1317,1343]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
), same data (
<collectionCode id="99FAAE48FFA6FFC1C040FAC7F4B2FAEE" box="[616,695,1317,1343]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA6FFC1C0E2FAC7F79AFAB2" collectingDate="1978-04-02" collectionCode="EUMJ" collectorName="N. Kanie" county="Yona" location="Is." municipality="Kunigami" pageId="4" pageNumber="493" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA6FFC1C0E2FAC7F52EFAEE" box="[714,811,1317,1343]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C112FAC7F5A5FAEE" box="[826,928,1317,1343]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
), “
<collectingCounty id="16354E01FFA6FFC1C195FAC7F5F9FAEE" box="[957,1020,1317,1343]" pageId="4" pageNumber="493">Yona</collectingCounty>
,
<collectingMunicipality id="1F30ACF7FFA6FFC1C62FFAC7F27FFAEE" box="[1031,1146,1317,1343]" pageId="4" pageNumber="493">Kunigami</collectingMunicipality>
,
<collectingRegion id="3D2FF86FFFA6FFC1C6ADFAC7F2EFFAEE" box="[1157,1258,1317,1343]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa</collectingRegion>
<location id="FA346056FFA6FFC1C6D9FAC4F308FA91" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4:FA346056FFA6FFC1C6D9FAC4F308FA91" box="[1265,1293,1318,1344]" county="Yona" municipality="Kunigami" name="Is." pageId="4" pageNumber="493" stateProvince="Okinawa">Is.</location>
,
<date id="8B55104DFFA6FFC1C732FAC7F39DFAEE" box="[1306,1432,1317,1344]" pageId="4" pageNumber="493" value="1978-04-02">
<collectingDate id="9B11E9A5FFA6FFC1C732FAC7F39DFAEE" box="[1306,1432,1317,1344]" pageId="4" pageNumber="493" value="1978-04-02">2. IV. 1978</collectingDate>
</date>
,
<collectorName id="521E535BFFA6FFC1C2BFFAA8F6F9FAB2" box="[151,252,1353,1380]" pageId="4" pageNumber="493">N. Kanie</collectorName>
leg.” (
<collectionCode id="99FAAE48FFA6FFC1C36FFAABF793FAB2" box="[327,406,1353,1379]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA6FFC1C382FAABF6F2FA56" collectingDate="1981-06-07" collectionCode="EUMJ" location="OKINAWA" pageId="4" pageNumber="493" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA6FFC1C382FAABF40FFAB2" box="[426,522,1353,1379]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C031FAABF47AFAB2" box="[537,639,1353,1379]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
), “IERINDOH, KUNIGAMI V,
<collectingRegion id="3D2FF86FFFA6FFC1C1C1FAABF275FAB2" box="[1001,1136,1353,1379]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">OKINAWA</collectingRegion>
,
<date id="8B55104DFFA6FFC1C653FAA8F2E8FAB2" box="[1147,1261,1353,1380]" pageId="4" pageNumber="493" value="1981-06-07">
<collectingDate id="9B11E9A5FFA6FFC1C653FAA8F2E8FAB2" box="[1147,1261,1353,1380]" pageId="4" pageNumber="493" value="1981-06-07">7 VI 1981</collectingDate>
</date>
, T. OHMOTO” (
<collectionCode id="99FAAE48FFA6FFC1C2B7FA8FF6EBFA56" box="[159,238,1389,1415]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FF54368DFFA6FFC0C2EFFA73F27FFECC" blockId="4.[151,1437,1281,2028]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFA73F444FA7A" bold="true" box="[199,577,1425,1451]" pageId="4" pageNumber="493">Additional materials examined.</emphasis>
<specimenCount id="E9EDFD04FFA6FFC1C061FA73F4A6FA7A" box="[585,675,1425,1451]" count="3" pageId="4" pageNumber="493" type="male">3 males</specimenCount>
, “Yona, Kunigami-son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C64DFA73F2FBFA7D" box="[1125,1278,1425,1452]" pageId="4" pageNumber="493" value="1995-02-28">28 Feb. 1995</date>
, H. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C2B7FA57F6F0FA1E" box="[159,245,1461,1487]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C321FA57F75DFA1E" box="[265,344,1461,1487]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C34DFA57F7CFFA1E" box="[357,458,1461,1487]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C0AEFA57F51AFA1E" box="[646,799,1461,1487]" pageId="4" pageNumber="493" value="1994-02-13">13 Feb. 1994</date>
, T. Shimizu leg.” (
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, same data but “
<date id="8B55104DFFA6FFC1C757FA57F70EFA22" pageId="4" pageNumber="493" value="1994-02-25">25 Feb. 1994</date>
, T. Shimizu leg.” (
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);
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,
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, same data but “
<date id="8B55104DFFA6FFC1C1E6FA3BF25EFA22" box="[974,1115,1497,1524]" pageId="4" pageNumber="493" value="1991-03-06">6 Mar. 1991</date>
, Y. Okushima leg. (
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);
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, same data but “
<date id="8B55104DFFA6FFC1C3EEFA1FF451F9C6" box="[454,596,1533,1560]" pageId="4" pageNumber="493" value="1991-03-07">7 Mar. 1991</date>
, Y. Okushima leg.” (
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);
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, same data but “
<date id="8B55104DFFA6FFC1C6F8FA1FF39DF9C6" box="[1232,1432,1533,1560]" pageId="4" pageNumber="493" value="1988-03-12" valueMax="1988-03-14" valueMin="1988-03-12">12–14 Mar. 1988</date>
, Coll. Teruhisa Ueno” (
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,
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, “Yona, Kunigami-son, Okinawa, Ryukyus,
<collectingCountry id="87FC761DFFA6FFC1C69DF9C0F2FDF9ED" box="[1205,1272,1570,1596]" name="Japan" pageId="4" pageNumber="493">Japan</collectingCountry>
,
<date id="8B55104DFFA6FFC1C72BF9C3F37EF9ED" box="[1283,1403,1569,1596]" pageId="4" pageNumber="493" value="1991-03-05">5-III-1991</date>
, T. Ueno leg.” (
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);
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, “
<collectingRegion id="3D2FF86FFFA6FFC1C3D6F9A7F486F98E" box="[510,643,1605,1631]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">OKINAWA</collectingRegion>
IS., Yona,
<date id="8B55104DFFA6FFC1C0D5F9A7F571F98E" box="[765,884,1605,1631]" pageId="4" pageNumber="493" value="1988-03-13">1988-3-13</date>
, T. Ueno leg.” (
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);
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, “Yona, Okinawa, Ryukyus,
<date id="8B55104DFFA6FFC1C322F98BF78AF952" box="[266,399,1641,1667]" pageId="4" pageNumber="493" value="1994-04-20">20-IV-1994</date>
, M. Kawanabe” (
<collectionCode id="99FAAE48FFA6FFC1C070F98BF4A2F952" box="[600,679,1641,1667]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
);
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,
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, “Mt. Yonaha-dake, Kunigami Son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C2F5F96FF768F976" box="[221,365,1677,1704]" pageId="4" pageNumber="493" value="1991-03-08">8 Mar. 1991</date>
, Y. Okushima leg. (
<collectionCode id="99FAAE48FFA6FFC1C07BF96FF4ACF976" box="[595,681,1677,1703]" pageId="4" pageNumber="493">KURA</collectionCode>
);
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, same data but “
<date id="8B55104DFFA6FFC1C1F5F96FF26BF979" box="[989,1134,1677,1704]" pageId="4" pageNumber="493" value="1995-03-03">3 Mar. 1995</date>
, H. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C70BF96FF37CF976" box="[1315,1401,1677,1703]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C7A6F96FF6D5F91A" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
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, same data but “
<date id="8B55104DFFA6FFC1C02EF953F492F91A" box="[518,663,1713,1740]" pageId="4" pageNumber="493" value="1999-03-02">2 Mar. 1999</date>
, Coll. T. Nonaka” (
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);
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, same data but “
<date id="8B55104DFFA6FFC1C724F953F39DF91A" box="[1292,1432,1713,1740]" pageId="4" pageNumber="493" value="1990-04-05">5 Apr. 1990</date>
, Coll. Y. Okushima” (
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);
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, “Ohkuni-ridou, Okinawa, Ryukyus,
<date id="8B55104DFFA6FFC1C625F937F28CF93E" box="[1037,1161,1749,1775]" pageId="4" pageNumber="493" value="2003-02-10">10-II-2003</date>
, M. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C769F937F395F93E" box="[1345,1424,1749,1775]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C2BFF91BF6F6F8C2" box="[151,243,1785,1811]" count="2" pageId="4" pageNumber="493" type="male">2 males</specimenCount>
,
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, “
<collectingCountry id="87FC761DFFA6FFC1C3A5F91BF7ECF8C2" box="[397,489,1785,1811]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
: Okinawa-jima Is., Kunigami-son, Ura, Ôkuni-rindô,
<date id="8B55104DFFA6FFC1C67DF91BF2E7F8C2" box="[1109,1250,1785,1812]" pageId="4" pageNumber="493" value="2024-02-14">14. II. 2024</date>
, Ryo Nakamura leg.
<emphasis id="CD9FEA9FFFA6FFC1C2E0F8FCF7EDF8E9" bold="true" box="[200,488,1822,1848]" pageId="4" pageNumber="493">ṗäŝ 国DZť大国ṳã</emphasis>
” (
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);
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, “[
<emphasis id="CD9FEA9FFFA6FFC1C0DBF8FCF5A6F8E9" bold="true" box="[755,931,1822,1848]" pageId="4" pageNumber="493">ṗäŝ ạüƜ</emphasis>
]
<emphasis id="CD9FEA9FFFA6FFC1C19CF8FCF301F8E9" bold="true" box="[948,1284,1822,1848]" pageId="4" pageNumber="493">ṗäḁ国DZō国DZťṉǭẹ</emphasis>
,
<date id="8B55104DFFA6FFC1C73AF8FFF39DF8E9" box="[1298,1432,1821,1848]" pageId="4" pageNumber="493" value="2015-02-28">28. II. 2015</date>
, leg.
<emphasis id="CD9FEA9FFFA6FFC1C2E2F8A0F73FF88D" bold="true" box="[202,314,1858,1884]" pageId="4" pageNumber="493">DƂ尚*</emphasis>
” (
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,
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, “Benoki-dam, Kunigami Son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C680F8A3F33DF88D" box="[1192,1336,1857,1884]" pageId="4" pageNumber="493" value="1995-03-02">2 Mar. 1995</date>
, H. Sato leg.” (
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);
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, “
<emphasis id="CD9FEA9FFFA6FFC1C380F884F549F851" bold="true" box="[424,844,1894,1920]" pageId="4" pageNumber="493">mṵṗäḛŝṗä×ŝṉǭẹダム</emphasis>
, [
<collectingCountry id="87FC761DFFA6FFC1C148F884F5BAF851" box="[864,959,1894,1920]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
]: the Ryukyus, Okinawa Isls., Okinawa Is., Benoki,
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128°16’106.7”E,
<date id="8B55104DFFA6FFC1C074F86BF4DEF872" box="[604,731,1929,1956]" pageId="4" pageNumber="493" value="2023-03-05">5. III. 2023</date>
. J. Souma leg.” (
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, “
<emphasis id="CD9FEA9FFFA6FFC1C640F868F6CAF819" bold="true" pageId="4" pageNumber="493">mṵṗäḛŝṗä×ŝṉ ǭẹ</emphasis>
, [
<collectingCountry id="87FC761DFFA6FFC1C2CBF84CF747F819" box="[227,322,1966,1992]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
]: the Ryukyus, Okinawa Isls., Okinawa Is., Benoki Dam,
<geoCoordinate id="9ADF504AFFA6FFC1C196F84FF258F816" box="[958,1117,1965,1991]" degrees="26" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="493" precision="1" seconds="09.9" value="26.786083">26°47’09.9”N</geoCoordinate>
<geoCoordinate id="9ADF504AFFA6FFC1C64BF84FF30BF819" box="[1123,1294,1965,1992]" degrees="128" direction="east" minutes="15" orientation="longitude" pageId="4" pageNumber="493" precision="1" seconds="15.9" value="128.25441">128°15’15.9”E</geoCoordinate>
,
<date id="8B55104DFFA6FFC1C731F84FF39CF816" box="[1305,1433,1965,1992]" pageId="4" pageNumber="493" value="2023-03-05">5. III. 2023</date>
. J. Souma leg.” (
<collectionCode id="99FAAE48FFA6FFC1C364F833F79DF83A" box="[332,408,2001,2027]" pageId="4" pageNumber="493">FZUT</collectionCode>
);
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, “Haneji, Nago City, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C19EF833F24AF83A" box="[950,1103,2001,2028]" pageId="4" pageNumber="493" value="1991-03-11">11 Mar. 1991</date>
, Y. Okushima leg.” (
<collectionCode id="99FAAE48FFA6FFC1C712F833F395F83A" box="[1338,1424,2001,2027]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C2BFFF75F6E3FF60" box="[151,230,151,177]" count="1" pageId="5" pageNumber="494" type="male">1 male</specimenCount>
,
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, “Haneji, Okinawa,
<date id="8B55104DFFA7FFC0C063FF75F4D9FF60" box="[587,732,151,177]" pageId="5" pageNumber="494" value="1986-03-23">23. III. 1986</date>
, M. Sawai leg.” (
<collectionCode id="99FAAE48FFA7FFC0C18CFF75F5F6FF60" box="[932,1011,151,177]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="5" pageNumber="494">EUMJ</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C62FFF75F252FF60" box="[1031,1111,151,177]" count="1" pageId="5" pageNumber="494" type="male">1 male</specimenCount>
, “HANEJI, Is. Okinawa,
<date id="8B55104DFFA7FFC0C751FF75F6F9FF04" pageId="5" pageNumber="494" value="1987-03-16">16. III. 1987</date>
, M. SAWAI” (
<collectionCode id="99FAAE48FFA7FFC0C3B7FF59F7EEFF04" box="[415,491,187,213]" pageId="5" pageNumber="494">FZUT</collectionCode>
);
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, “Afuso, Onna Son, Okinawa Pref.,
<date id="8B55104DFFA7FFC0C1D0FF59F27BFF04" box="[1016,1150,187,213]" pageId="5" pageNumber="494" value="1989-04-03">3 Apr. 1989</date>
, Coll. T. Ueno” (
<collectionCode id="99FAAE48FFA7FFC0C712FF59F395FF04" box="[1338,1424,187,213]" pageId="5" pageNumber="494">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C2BFFF3DF6FBFF28" box="[151,254,223,249]" count="1" pageId="5" pageNumber="494" type="female">1 female</specimenCount>
, “Afuso, Okinawa-hontô Is., Okinawa Pref.,
<date id="8B55104DFFA7FFC0C122FF3DF5A0FF28" box="[778,933,223,249]" pageId="5" pageNumber="494" value="1989-04-21">21 Apr. 1989</date>
, Coll. Teruhisa Ueno” (
<collectionCode id="99FAAE48FFA7FFC0C692FF3DF315FF28" box="[1210,1296,223,249]" pageId="5" pageNumber="494">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C70EFF3DF39DFF28" box="[1318,1432,223,249]" count="2" pageId="5" pageNumber="494" type="female">2 females</specimenCount>
, “Kenminnomori, Onna Son, Okinawa Pref.,
<date id="8B55104DFFA7FFC0C0A2FEE1F526FECC" box="[650,803,259,285]" pageId="5" pageNumber="494" value="1990-03-31">31 Mar. 1990</date>
, Coll. Y. Okushima” (
<collectionCode id="99FAAE48FFA7FFC0C630FEE1F26BFECC" box="[1048,1134,259,285]" pageId="5" pageNumber="494">KURA</collectionCode>
).
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFEC4F7ECFE58" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFEC4F417FE91" bold="true" box="[199,530,294,320]" pageId="5" pageNumber="494">Supplementary description.</emphasis>
Male sternite IX (fig. 8D) cymbiform, slightly shorter than tergite IX, 2.80 times longer than wide, lateral margins parallel in the middle, abruptly narrowing in apical 1/4 to form a triangle with a rounded acute angle at the tip.
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA7FFC0C2EFFE70F545FE7C" box="[199,832,402,429]" pageId="5" pageNumber="494" type="distribution">
<paragraph id="FF54368DFFA7FFC0C2EFFE70F545FE7C" blockId="5.[151,1437,151,789]" box="[199,832,402,429]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFE70F764FE7D" bold="true" box="[199,353,402,428]" pageId="5" pageNumber="494">Distribution.</emphasis>
<collectingCountry id="87FC761DFFA7FFC0C34FFE71F7A2FE7C" box="[359,423,403,429]" name="Japan" pageId="5" pageNumber="494">Japan</collectingCountry>
(Ryukyu:
<collectingRegion id="3D2FF86FFFA7FFC0C036FE71F486FE7C" box="[542,643,403,429]" country="Japan" name="Okinawa" pageId="5" pageNumber="494">Okinawa</collectingRegion>
Island) (fig. 11).
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA7FFC0C2EFFE54F6ECFCC4" pageId="5" pageNumber="494" type="biology_ecology">
<paragraph id="FF54368DFFA7FFC0C2EFFE54F729FDC8" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFE54F722FE01" bold="true" box="[199,295,438,464]" pageId="5" pageNumber="494">Biology.</emphasis>
This species can be caught by sweeping blossoms and green leaves in the primitive or secondary broad-leaved forest (
<bibRefCitation id="9B7A4B7CFFA7FFC0C3A9FE39F43AFE24" author="Okushima, Y." box="[385,575,474,501]" pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005</bibRefCitation>
; the author’s personal observation). Adults are collected from mid-February to mid-April.
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFDC1F38EFD1C" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFDC1F404FDEC" bold="true" box="[199,513,547,573]" pageId="5" pageNumber="494">Nomenclatural comments.</emphasis>
<bibRefCitation id="9B7A4B7CFFA7FFC0C020FDC1F49BFDEC" author="Ishida, K." box="[520,670,547,573]" pageId="5" pageNumber="494" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida (1986)</bibRefCitation>
has hitherto been regarded as the original description of this species (
<bibRefCitation id="9B7A4B7CFFA7FFC0C2B7FDA5F7A8FDB0" author="Sato, M. &amp; Okushima, Y." box="[159,429,583,609]" pageId="5" pageNumber="494" pagination="857 - 865" refId="ref5090" refString="Sato, M. &amp; Okushima, Y. (1992) The genus Athemus (Coleoptera, Cantharidae) of the Ryukyu Islands, southwest Japan. Japanese Journal of Entomology, 60, 857 - 865." type="journal article" year="1992">Satô &amp; Okushima, 1992</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C39EFDA5F46AFDB0" author="Takahashi, K." box="[438,623,583,609]" pageId="5" pageNumber="494" pagination="29 - 48" refId="ref5129" refString="Takahashi, K. (1998) Check-list of Cantharidae of Japan. Kanagawa-Chuho, 122, 29 - 48. [in Japanese, with English title]" type="journal article" year="1998">Takahashi, 1998</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C051FDA5F536FDB0" author="Okushima, Y." box="[633,819,582,609]" pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C115FDA5F28EFDB0" author="Kazantsev, S. &amp; Brancucci, M." box="[829,1163,583,609]" pageId="5" pageNumber="494" refId="ref4627" refString="Kazantsev, S. &amp; Brancucci, M. (2007) Cantharidae. In Lo ¨ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 234 - 298." type="book" year="2007">Kazantsev &amp; Brancucci, 2007</bibRefCitation>
), but the name “
<taxonomicName id="38EB4D0EFFA7FFC0C714FDA5F769FD54" authority="Ishida" authorityName="Ishida" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA7FFC0C714FDA5F718FD54" italics="true" pageId="5" pageNumber="494">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
” did appear precedently in
<bibRefCitation id="9B7A4B7CFFA7FFC0C08EFD89F52FFD54" author="Sato, M." box="[678,810,618,645]" pageId="5" pageNumber="494" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô (1985)</bibRefCitation>
along with color photograph and a short description in the key. Based on ICZN Article 10 and 13 (
<bibRefCitation id="9B7A4B7CFFA7FFC0C0BCFD6DF35FFD78" author="International Commission on Zoological Nomenclature &amp; Tipografia La" box="[660,1370,655,681]" pageId="5" pageNumber="494" refId="ref4522" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. Tipografia La Garangola, Padova, xxix + 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature, 1999</bibRefCitation>
), it is considered that the name
<taxonomicName id="38EB4D0EFFA7FFC0C39DFD51F4E8FD1C" authority="Ishida" authorityName="Ishida" box="[437,749,691,717]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA7FFC0C39DFD51F4A6FD1C" box="[437,675,691,717]" italics="true" pageId="5" pageNumber="494">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
was available at the time of the publication by
<bibRefCitation id="9B7A4B7CFFA7FFC0C728FD51F38DFD1C" author="Sato, M." box="[1280,1416,690,717]" pageId="5" pageNumber="494" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô (1985)</bibRefCitation>
.
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFD35F6ECFCC4" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFD35F703FD20" bold="true" box="[199,262,727,753]" pageId="5" pageNumber="494">Note.</emphasis>
This species is placed in the
<taxonomicName id="38EB4D0EFFA7FFC0C05AFD35F554FD21" box="[626,849,727,752]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="aegrotus">
<emphasis id="CD9FEA9FFFA7FFC0C05AFD35F554FD21" box="[626,849,727,752]" italics="true" pageId="5" pageNumber="494">Lycocerus aegrotus</emphasis>
</taxonomicName>
subgroup of the
<taxonomicName id="38EB4D0EFFA7FFC0C60CFD35F2A2FD21" box="[1060,1191,727,752]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="aegrotus">
<emphasis id="CD9FEA9FFFA7FFC0C60CFD35F2A2FD21" box="[1060,1191,727,752]" italics="true" pageId="5" pageNumber="494">L. aegrotus</emphasis>
</taxonomicName>
group by
<bibRefCitation id="9B7A4B7CFFA7FFC0C701FD35F6E0FCC4" author="Okushima, Y." pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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320
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<paragraph id="F07136CD5247FFE15F80EACA4F9EFF4C" blockId="9.[151,925,151,178]" box="[151,925,151,178]" pageId="9" pageNumber="436">
<heading id="AB3981A15247FFE15F80EACA4F9EFF4C" bold="true" box="[151,925,151,178]" fontSize="11" level="1" pageId="9" pageNumber="436" reason="1">
<emphasis id="C2BAEADF5247FFE15F80EACA4F9EFF4C" bold="true" box="[151,925,151,178]" pageId="9" pageNumber="436">
<taxonomicName id="37CE4D4E5247FFE15F80EACA4EF8FF4C" authority="Timokhov &amp; Belokobylskij, 2020" authorityName="Timokhov &amp; Belokobylskij" authorityYear="2020" box="[151,763,151,178]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="436" phylum="Arthropoda" rank="species" species="yurii">
<emphasis id="C2BAEADF5247FFE15F80EACA4D5BFF4C" bold="true" box="[151,344,151,178]" italics="true" pageId="9" pageNumber="436">Vanhornia yurii</emphasis>
<bibRefCitation id="945F4B3C5247FFE15E48EACA4EF8FF4C" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[351,763,151,177]" pageId="9" pageNumber="436" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij, 2020</bibRefCitation>
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(
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<emphasis id="C2BAEADF5247FFE15F80EA814D9DFF09" box="[151,414,220,244]" italics="true" pageId="9" pageNumber="436">Vanhornia eucnemidarum</emphasis>
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:
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<bibRefCitation id="945F4B3C5247FFE15EBCEA814E88FF0E" author="Choi, M. B. &amp; Lee, J. W." box="[427,651,219,244]" pageId="9" pageNumber="436" pagination="59 - 61" refId="ref6362" refString="Choi, M. B. &amp; Lee, J. W. (2012) First record of Vanhorniidae (Hymenoptera: Proctotrupoidea) from Korea. Journal of Asia- Pacific Entomology, 15, 59 - 61. https: // doi. org / 10.1016 / j. aspen. 2011.08.003" type="journal article" year="2012">Choi &amp; Lee, 2012: 60</bibRefCitation>
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. Misidentification.
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<treatmentCitationGroup id="D0DE11E35247FFE15F80EAA74D73FECC" pageId="9" pageNumber="436">
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<taxonomicName id="37CE4D4E5247FFE15F80EAA74EEBFEEF" authority="Timokhov &amp; Belokobylskij, 2020: 60 - 67" authorityName="Timokhov &amp; Belokobylskij" authorityPageNumber="60 - 67" authorityYear="2020" box="[151,744,250,274]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="436" phylum="Arthropoda" rank="species" species="yurii">
<emphasis id="C2BAEADF5247FFE15F80EAA74D3BFEEF" box="[151,312,250,274]" italics="true" pageId="9" pageNumber="436">Vanhornia yurii</emphasis>
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<bibRefCitation id="945F4B3C5247FFE15E29EAA74EEBFEEF" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[318,744,250,274]" pageId="9" pageNumber="436" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij, 2020: 60–67</bibRefCitation>
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.
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(not examined): female; type locality:
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,
<collectingCountry id="88D9765D5247FFE15E32EB444D6FFECC" box="[293,364,281,305]" name="Russia" pageId="9" pageNumber="436">Russia</collectingCountry>
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<emphasis id="C2BAEADF5247FFE15F80EB024D11FE84" bold="true" box="[151,274,351,377]" pageId="9" pageNumber="436">Diagnosis.</emphasis>
Frontal transverse carina absent; apical margin of labrum straight. Scutoscutellar sulcus with seven foveae; mesodiscrimen with about 10 strong foveae; median longitudinal carina on dorsal part of propodeum present; smooth area on postero-lateral part of propodeum present. Vein Rs+M not connected to Cu. Syntergite with longitudinal ridges, reaching to a maximum of 0.4 length of syntergite, touching first thyridium; antero-dorsal margin of syntergite almost straight.
</paragraph>
<caption id="A4B166455247FFE15F8FEFA14E93F9A1" ID-DOI="http://doi.org/10.5281/zenodo.13757864" ID-Zenodo-Dep="13757864" httpUri="https://zenodo.org/record/13757864/files/figure.png" pageId="9" pageNumber="436" startId="9.[152,255,1532,1556]" targetBox="[151,1435,545,1509]" targetPageId="9" targetType="figure">
<paragraph id="F07136CD5247FFE15F8FEFA14E93F9A1" blockId="9.[151,1437,1532,1629]" pageId="9" pageNumber="436">
<emphasis id="C2BAEADF5247FFE15F8FEFA14D4BF9E9" bold="true" box="[152,328,1532,1556]" pageId="9" pageNumber="436">FIGURE 7A–F.</emphasis>
<taxonomicName id="37CE4D4E5247FFE15E58EFA14DF3F9E9" authorityName="Timokhov &amp; Belokobylskij" authorityYear="2020" box="[335,496,1532,1556]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="436" phylum="Arthropoda" rank="species" species="yurii">
<emphasis id="C2BAEADF5247FFE15E58EFA14DF3F9E9" box="[335,496,1532,1556]" italics="true" pageId="9" pageNumber="436">Vanhornia yurii</emphasis>
</taxonomicName>
(Japan, female). A, lateral habitus; B, frontal view of head; C, dorsal view of mesoscutum; D, ventral view of mesodiscrimen; E, dorsal view of scutellum, metanotum, and propodeum; F, dorsal view of metasoma Scale bars: A—2.0 mm, B, D–F—0.5 mm, C—0.2 mm.
</paragraph>
</caption>
<paragraph id="F07136CD5247FFE15FD0ECF24E14F85C" blockId="9.[151,1436,1710,2025]" pageId="9" pageNumber="436">
<emphasis id="C2BAEADF5247FFE15FD0ECF24D4BF934" bold="true" box="[199,328,1711,1737]" pageId="9" pageNumber="436">Non-types.</emphasis>
<materialsCitation id="40A63C905247FFE15E58ECF34FF4F910" collectionCode="MT" collectorName="Chongdo-gun &amp; Unmun-myeon" country="South Korea" latitude="35.647224" location="Mt. Unmun" longLatPrecision="19" longitude="128.97472" pageId="9" pageNumber="436" specimenCount="1" stateProvince="Gyeongbuk">
<emphasis id="C2BAEADF5247FFE15E58ECF34DE5F934" bold="true" box="[335,486,1710,1737]" pageId="9" pageNumber="436">
<collectingCountry id="88D9765D5247FFE15E58ECF34DE5F934" box="[335,486,1710,1737]" name="South Korea" pageId="9" pageNumber="436">South Korea</collectingCountry>
</emphasis>
: 1f (recorded as
<taxonomicName id="37CE4D4E5247FFE15DB2ECF24F65F934" authorityName="Crawford" authorityYear="1909" box="[677,870,1711,1737]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="436" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5247FFE15DB2ECF24F65F934" box="[677,870,1711,1737]" italics="true" pageId="9" pageNumber="436">V. eucnemidarum</emphasis>
</taxonomicName>
in
<bibRefCitation id="945F4B3C5247FFE15C9CECF2486EF934" author="Choi, M. B. &amp; Lee, J. W." box="[907,1133,1711,1737]" pageId="9" pageNumber="436" pagination="59 - 61" refId="ref6362" refString="Choi, M. B. &amp; Lee, J. W. (2012) First record of Vanhorniidae (Hymenoptera: Proctotrupoidea) from Korea. Journal of Asia- Pacific Entomology, 15, 59 - 61. https: // doi. org / 10.1016 / j. aspen. 2011.08.003" type="journal article" year="2012">Choi &amp; Lee (2012))</bibRefCitation>
, GB (
<collectingRegion id="320AF82F5247FFE15BA6ECF24992F934" box="[1201,1425,1711,1737]" country="South Korea" name="Gyeongbuk" pageId="9" pageNumber="436">Gyeongsangbuk-do</collectingRegion>
),
<collectorName id="5D3B531B5247FFE15F80EC8E4D37F910" box="[151,308,1747,1773]" pageId="9" pageNumber="436">Chongdo-gun</collectorName>
,
<collectorName id="5D3B531B5247FFE15E57EC8E4DEEF910" box="[320,493,1747,1773]" pageId="9" pageNumber="436">Unmun-myeon</collectorName>
,
<location id="F51160165247FFE15EEEEC8E4E7CF910" LSID="urn:lsid:plazi:treatment:786787DB5247FFE25F80EACA4951FE2C:F51160165247FFE15EEEEC8E4E7CF910" box="[505,639,1747,1773]" country="South Korea" latitude="35.647224" longLatPrecision="19" longitude="128.97472" name="Mt. Unmun" pageId="9" pageNumber="436" stateProvince="Gyeongbuk">Mt. Unmun</location>
,
<geoCoordinate id="95FA500A5247FFE15D9CEC8F4F18F910" box="[651,795,1746,1773]" degrees="35" direction="north" minutes="38" orientation="latitude" pageId="9" pageNumber="436" precision="15" seconds="50" value="35.647224">35°38’50’’N</geoCoordinate>
,
<geoCoordinate id="95FA500A5247FFE15C3FEC8F4FC1F910" box="[808,962,1746,1773]" degrees="128" direction="east" minutes="58" orientation="longitude" pageId="9" pageNumber="436" precision="15" seconds="29" value="128.97472">128°58’29’’E</geoCoordinate>
,
<collectionCode id="96DFAE085247FFE15CD8EC8E4FF4F910" box="[975,1015,1747,1773]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="9" pageNumber="436">MT</collectionCode>
</materialsCitation>
,
<date id="8470100D5247FFE15B11EC8E48CCF910" box="[1030,1231,1746,1773]" pageId="9" pageNumber="436" value="2009-05-30" valueMax="2009-06-16" valueMin="2009-05-30">
30 V–16 VI
<materialsCitation id="40A63C905247FFE15B83EC8E48CCF910" box="[1172,1231,1747,1773]" collectionCode="YNU" collectorName="Lee" country="South Korea" pageId="9" pageNumber="436" specimenCount="1">2009</materialsCitation>
</date>
, J.W.
<collectorName id="5D3B531B5247FFE15A03EC8E493DF910" box="[1300,1342,1747,1773]" pageId="9" pageNumber="436">Lee</collectorName>
(YNU).
<materialsCitation id="40A63C905247FFE15F80ECAA4F02F8C8" collectingDate="2010-06-22" collectingDateMax="2011-07-04" collectingDateMin="2010-06-22" collectionCode="VI, VII, MT" collectorName="Sapporo &amp; Hitsujigaoka" country="Japan" latitude="43.0025" location="Hokkaido" longLatPrecision="19" longitude="141.41554" pageId="9" pageNumber="436" specimenCount="1" stateProvince="Hokkaido">
<emphasis id="C2BAEADF5247FFE15F80ECAA4D78F8ED" bold="true" box="[151,379,1782,1809]" pageId="9" pageNumber="436">
<collectingCountry id="88D9765D5247FFE15F80ECAA4CE7F8EC" box="[151,228,1783,1809]" name="Japan" pageId="9" pageNumber="436">Japan</collectingCountry>
: [
<collectingRegion id="320AF82F5247FFE15FEEECAB4D77F8ED" box="[249,372,1782,1808]" country="Japan" name="Hokkaido" pageId="9" pageNumber="436">Hokkaido</collectingRegion>
]
</emphasis>
3f (
<date id="8470100D5247FFE15EBCECAA4E5FF8EC" box="[427,604,1783,1809]" pageId="9" pageNumber="436" value="2010-06-22" valueMax="2010-06-29" valueMin="2010-06-22">
<collectingDate id="9434E9E55247FFE15EBCECAA4E5FF8EC" box="[427,604,1783,1809]" pageId="9" pageNumber="436" value="2010-06-22" valueMax="2010-06-29" valueMin="2010-06-22">
22–29
<collectionCode id="96DFAE085247FFE15EEDECAA4E1BF8EC" box="[506,536,1783,1809]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="9" pageNumber="436">VI</collectionCode>
2010
</collectingDate>
</date>
), 1f (
<date id="8470100D5247FFE15D8FECAA4F69F8EC" box="[664,874,1782,1809]" pageId="9" pageNumber="436" value="2010-06-29" valueMax="2010-07-06" valueMin="2010-06-29">
<collectingDate id="9434E9E55247FFE15D8FECAA4F69F8EC" box="[664,874,1782,1809]" pageId="9" pageNumber="436" value="2010-06-29" valueMax="2010-07-06" valueMin="2010-06-29">
29
<collectionCode id="96DFAE085247FFE15DAAECAB4ED8F8ED" box="[701,731,1782,1808]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="9" pageNumber="436">VI</collectionCode>
–6
<collectionCode id="96DFAE085247FFE15DE8ECAA4F25F8EC" box="[767,806,1783,1809]" pageId="9" pageNumber="436">VII</collectionCode>
2010
</collectingDate>
</date>
), 1f (
<date id="8470100D5247FFE15CB1ECAB4851F8EC" box="[934,1106,1782,1809]" pageId="9" pageNumber="436" value="2010-07-06" valueMax="2010-07-13" valueMin="2010-07-06">
<collectingDate id="9434E9E55247FFE15CB1ECAB4851F8EC" box="[934,1106,1782,1809]" pageId="9" pageNumber="436" value="2010-07-06" valueMax="2010-07-13" valueMin="2010-07-06">
6–13
<collectionCode id="96DFAE085247FFE15CF0ECAA480DF8EC" box="[999,1038,1783,1809]" pageId="9" pageNumber="436">VII</collectionCode>
2010
</collectingDate>
</date>
), and 2f (
<date id="8470100D5247FFE15BD7ECAA4993F8EC" box="[1216,1424,1783,1809]" pageId="9" pageNumber="436" value="2011-06-27" valueMax="2011-07-04" valueMin="2011-06-27">
<collectingDate id="9434E9E55247FFE15BD7ECAA4993F8EC" box="[1216,1424,1783,1809]" pageId="9" pageNumber="436" value="2011-06-27" valueMax="2011-07-04" valueMin="2011-06-27">
27
<collectionCode id="96DFAE085247FFE15BF2ECAA4901F8EC" box="[1253,1282,1783,1809]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="9" pageNumber="436">VI</collectionCode>
–4
<collectionCode id="96DFAE085247FFE15A31ECAA494EF8EC" box="[1318,1357,1783,1809]" pageId="9" pageNumber="436">VII</collectionCode>
2011
</collectingDate>
</date>
),
<collectorName id="5D3B531B5247FFE15F80ED464CF4F8C8" box="[151,247,1819,1845]" pageId="9" pageNumber="436">Sapporo</collectorName>
,
<collectorName id="5D3B531B5247FFE15E16ED464D90F8C8" box="[257,403,1819,1845]" pageId="9" pageNumber="436">Hitsujigaoka</collectorName>
,
<geoCoordinate id="95FA500A5247FFE15E8AED464E2FF8C8" box="[413,556,1819,1845]" degrees="43" direction="north" minutes="00" orientation="latitude" pageId="9" pageNumber="436" precision="15" seconds="09" value="43.0025">43°00’09” N</geoCoordinate>
<geoCoordinate id="95FA500A5247FFE15D24ED474ECDF8C8" box="[563,718,1818,1845]" degrees="141" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="436" precision="15" seconds="56" value="141.41554">141°24’56” E</geoCoordinate>
,
<collectionCode id="96DFAE085247FFE15DCEED464F02F8C8" box="[729,769,1819,1845]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="9" pageNumber="436">MT</collectionCode>
</materialsCitation>
,
<materialsCitation id="40A63C905247FFE15C19ED464D06F8A4" collectionCode="ELKU" collectorName="K. Konishi" country="Japan" latitude="43.033333" location="Mt. Maruyama" longLatPrecision="1146" longitude="141.3" municipality="Sapporo" pageId="9" pageNumber="436" specimenCount="1">
<collectorName id="5D3B531B5247FFE15C19ED464F8BF8C8" box="[782,904,1819,1845]" pageId="9" pageNumber="436">K. Konishi</collectorName>
(
<collectionCode id="96DFAE085247FFE15C81ED464FE0F8C8" box="[918,995,1819,1845]" pageId="9" pageNumber="436">ELKU</collectionCode>
); 1f,
<collectingMunicipality id="1015ACB75247FFE15B0AED46487EF8C8" box="[1053,1149,1819,1845]" pageId="9" pageNumber="436">Sapporo</collectingMunicipality>
,
<location id="F51160165247FFE15B9FED464932F8C8" LSID="urn:lsid:plazi:treatment:786787DB5247FFE25F80EACA4951FE2C:F51160165247FFE15B9FED464932F8C8" box="[1160,1329,1819,1845]" country="Japan" latitude="43.033333" longLatPrecision="1146" longitude="141.3" municipality="Sapporo" name="Mt. Maruyama" pageId="9" pageNumber="436">Mt. Maruyama</location>
,
<geoCoordinate id="95FA500A5247FFE15A2CED46499FF8C8" box="[1339,1436,1819,1845]" degrees="43" direction="north" minutes="02" orientation="latitude" pageId="9" pageNumber="436" precision="925" value="43.033333">43°02’N</geoCoordinate>
<geoCoordinate id="95FA500A5247FFE15F80ED624D06F8A4" box="[151,261,1855,1881]" degrees="141" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="436" precision="925" value="141.3">141°18’E</geoCoordinate>
</materialsCitation>
,
<date id="8470100D5247FFE15E05ED624D91F8A4" box="[274,402,1855,1881]" pageId="9" pageNumber="436" value="2017-07-06">
6 VII
<materialsCitation id="40A63C905247FFE15E40ED624D91F8A4" box="[343,402,1855,1881]" collectionCode="SEHU" collectorName="Juriya Okayasu" country="Japan" latitude="43.605" location="Tsunemoto" longLatPrecision="6" longitude="143.344" municipality="Oketo Town" pageId="9" pageNumber="436" specimenCount="1">2017</materialsCitation>
</date>
,
<collectorName id="5D3B531B5247FFE15E89ED624E4EF8A4" box="[414,589,1855,1881]" pageId="9" pageNumber="436">Juriya Okayasu</collectorName>
(SEHU); 1f,
<collectingMunicipality id="1015ACB75247FFE15DFFED624F75F8A4" box="[744,886,1855,1881]" pageId="9" pageNumber="436">Oketo Town</collectingMunicipality>
,
<location id="F51160165247FFE15C94ED624801F8A4" LSID="urn:lsid:plazi:treatment:786787DB5247FFE25F80EACA4951FE2C:F51160165247FFE15C94ED624801F8A4" box="[899,1026,1855,1881]" country="Japan" latitude="43.605" longLatPrecision="6" longitude="143.344" municipality="Oketo Town" name="Tsunemoto" pageId="9" pageNumber="436">Tsunemoto</location>
,
<geoCoordinate id="95FA500A5247FFE15B19ED63487EF8A4" box="[1038,1149,1854,1881]" degrees="43.6050" direction="north" orientation="latitude" pageId="9" pageNumber="436" precision="5" value="43.605">43.6050N</geoCoordinate>
<geoCoordinate id="95FA500A5247FFE15B92ED624901F8A4" box="[1157,1282,1855,1881]" degrees="143.3440" direction="east" orientation="longitude" pageId="9" pageNumber="436" precision="5" value="143.344">143.3440E</geoCoordinate>
,
<date id="8470100D5247FFE15A18ED62499BF8A4" box="[1295,1432,1855,1881]" pageId="9" pageNumber="436" value="2013-08-03">3 VIII 2013</date>
, R.
<materialsCitation id="40A63C905247FFE15FA0ED3E48E2F880" box="[183,1249,1890,1917]" collectionCode="OMNH" collectorName="Matsumoto" country="Japan" latitude="35.333332" location="Mt. Hyono-sen" longLatPrecision="1194" longitude="134.51666" municipality="Odanganaru" pageId="9" pageNumber="436" specimenCount="1" stateProvince="Hyogo">
<collectorName id="5D3B531B5247FFE15FA0ED3E4D34F880" box="[183,311,1891,1917]" pageId="9" pageNumber="436">Matsumoto</collectorName>
(
<collectionCode id="96DFAE085247FFE15E50ED3E4D9EF880" box="[327,413,1891,1917]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="9" pageNumber="436" type="Museum">OMNH</collectionCode>
).
<emphasis id="C2BAEADF5247FFE15EA4ED3E4E20F880" bold="true" box="[435,547,1891,1917]" pageId="9" pageNumber="436">[Honshu]</emphasis>
<collectingRegion id="320AF82F5247FFE15D3EED3E4EB6F880" box="[553,693,1891,1917]" country="Japan" name="Hyogo" pageId="9" pageNumber="436">Hyogo Pref.</collectingRegion>
,
<collectingMunicipality id="1015ACB75247FFE15DA8ED3E4F48F880" box="[703,843,1891,1917]" pageId="9" pageNumber="436">Odanganaru</collectingMunicipality>
,
<location id="F51160165247FFE15C41ED3E4801F880" LSID="urn:lsid:plazi:treatment:786787DB5247FFE25F80EACA4951FE2C:F51160165247FFE15C41ED3E4801F880" box="[854,1026,1891,1917]" country="Japan" latitude="35.333332" longLatPrecision="1194" longitude="134.51666" municipality="Odanganaru" name="Mt. Hyono-sen" pageId="9" pageNumber="436" stateProvince="Hyogo">Mt. Hyono-sen</location>
,
<geoCoordinate id="95FA500A5247FFE15B1BED3F486EF880" box="[1036,1133,1890,1917]" degrees="35" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="436" precision="925" value="35.333332">35°20’N</geoCoordinate>
<geoCoordinate id="95FA500A5247FFE15B64ED3E48E2F880" box="[1139,1249,1891,1917]" degrees="134" direction="east" minutes="31" orientation="longitude" pageId="9" pageNumber="436" precision="925" value="134.51666">134°31’E</geoCoordinate>
</materialsCitation>
,
<date id="8470100D5247FFE15BFBED3E4974F880" box="[1260,1399,1891,1917]" pageId="9" pageNumber="436" value="2012-07-21">
21 VII
<materialsCitation id="40A63C905247FFE15A2BED3E4974F880" box="[1340,1399,1891,1917]" collectionCode="KPMNH" collectorName="K. Watanabe &amp; Kanno" country="Japan" pageId="9" pageNumber="436" specimenCount="1">2012</materialsCitation>
</date>
,
<collectorName id="5D3B531B5247FFE15A96ED3E4D06F85C" pageId="9" pageNumber="436">K. Watanabe</collectorName>
&amp; Y.
<collectorName id="5D3B531B5247FFE15E51EDDA4D92F85C" box="[326,401,1927,1953]" pageId="9" pageNumber="436">Kanno</collectorName>
(KPMNH).
</paragraph>
</subSubSection>
<subSubSection id="B8D465465247FFE15FD0EDF64D80F838" box="[199,387,1963,1989]" pageId="9" pageNumber="436" type="biology_ecology">
<paragraph id="F07136CD5247FFE15FD0EDF64D80F838" blockId="9.[151,1436,1710,2025]" box="[199,387,1963,1989]" pageId="9" pageNumber="436">
<emphasis id="C2BAEADF5247FFE15FD0EDF64D05F838" bold="true" box="[199,262,1963,1989]" pageId="9" pageNumber="436">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465247FFE15FD0ED93495EF814" box="[199,1373,1998,2025]" pageId="9" pageNumber="436" type="distribution">
<paragraph id="F07136CD5247FFE15FD0ED93495EF814" blockId="9.[151,1436,1710,2025]" box="[199,1373,1998,2025]" pageId="9" pageNumber="436">
<emphasis id="C2BAEADF5247FFE15FD0ED934D62F815" bold="true" box="[199,353,1998,2024]" pageId="9" pageNumber="436">Distribution.</emphasis>
<collectingCountry id="88D9765D5247FFE15E70ED924DA4F814" box="[359,423,1999,2025]" name="Japan" pageId="9" pageNumber="436">Japan</collectingCountry>
(new record),
<collectingCountry id="88D9765D5247FFE15D5BED924E94F814" box="[588,663,1999,2025]" name="Russia" pageId="9" pageNumber="436">Russia</collectingCountry>
(
<bibRefCitation id="945F4B3C5247FFE15DB0ED92481CF814" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[679,1055,1999,2025]" pageId="9" pageNumber="436" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij 2020</bibRefCitation>
),
<collectingCountry id="88D9765D5247FFE15B25ED9248BDF814" box="[1074,1214,1999,2025]" name="South Korea" pageId="9" pageNumber="436">South Korea</collectingCountry>
(new record).
</paragraph>
</subSubSection>
<subSubSection id="B8D465465244FFE25FD0EACA4951FE2C" pageId="10" pageNumber="437" type="discussion">
<paragraph id="F07136CD5244FFE25FD0EACA4F02FF04" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="437">
<emphasis id="C2BAEADF5244FFE25FD0EACA4D38FF4C" bold="true" box="[199,315,151,177]" pageId="10" pageNumber="437">Remarks.</emphasis>
<bibRefCitation id="945F4B3C5244FFE25E52EACA4ED5FF4C" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[325,726,151,177]" pageId="10" pageNumber="437" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij (2020)</bibRefCitation>
suggested that the South Korean record of
<taxonomicName id="37CE4D4E5244FFE25BC0EACA499FFF4C" authorityName="Crawford" authorityYear="1909" box="[1239,1436,151,177]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="437" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5244FFE25BC0EACA499FFF4C" box="[1239,1436,151,177]" italics="true" pageId="10" pageNumber="437">V. eucnemidarum</emphasis>
</taxonomicName>
needed to be reconfirmed. We examined the voucher specimen of this record and photographs in
<bibRefCitation id="945F4B3C5244FFE25BDEEAE6499FFF28" author="Choi, M. B. &amp; Lee, J. W." box="[1225,1436,187,213]" pageId="10" pageNumber="437" pagination="59 - 61" refId="ref6362" refString="Choi, M. B. &amp; Lee, J. W. (2012) First record of Vanhorniidae (Hymenoptera: Proctotrupoidea) from Korea. Journal of Asia- Pacific Entomology, 15, 59 - 61. https: // doi. org / 10.1016 / j. aspen. 2011.08.003" type="journal article" year="2012">Choi &amp; Lee (2012)</bibRefCitation>
and identified the species from
<collectingCountry id="88D9765D5244FFE25EE0EA824E87FF04" box="[503,644,223,249]" name="South Korea" pageId="10" pageNumber="437">South Korea</collectingCountry>
as
<taxonomicName id="37CE4D4E5244FFE25DBEEA824EF9FF05" authorityName="Timokhov &amp; Belokobylskij" authorityYear="2020" box="[681,762,223,248]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="437" phylum="Arthropoda" rank="species" species="yurii">
<emphasis id="C2BAEADF5244FFE25DBEEA824EF9FF05" box="[681,762,223,248]" italics="true" pageId="10" pageNumber="437">V. yurii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F07136CD5244FFE25FD0EB5E4F59FE74" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="437">The morphological variations were recognized as follows: median longitudinal sulcus on vertex sometimes present, but weak; foveae on notauli sometimes fused posteriorly; syntergite 1.96–2.4 times as long as its maximum width in dorsal view; length of ovipositor 1.1–1.9 times as long as metasoma in lateral view (normally reaching to around pronotum, but sometimes not reaching to middle coxa).</paragraph>
<paragraph id="F07136CD5244FFE25FD0EBCE4951FE2C" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="437">
The specimen from Oketo Town,
<collectingRegion id="320AF82F5244FFE25D58EBCE4EC3FE50" box="[591,704,403,429]" country="Japan" name="Hokkaido" pageId="10" pageNumber="437">Hokkaido</collectingRegion>
,
<collectingCountry id="88D9765D5244FFE25DD9EBCE4F0DFE50" box="[718,782,403,429]" name="Japan" pageId="10" pageNumber="437">Japan</collectingCountry>
was collected near some logs in the forest (
<figureCitation id="68F52A485244FFE25A05EBCE495AFE50" box="[1298,1369,402,429]" captionStart="FIGURE 5" captionStartId="7.[152,255,1978,2002]" captionTargetBox="[365,1222,1311,1954]" captionTargetId="figure-75@7.[365,1222,1311,1954]" captionTargetPageId="7" captionText="FIGURE 5. Logs in the forest in Hokkaido, Japan, where Vanhornia leileri and V. yurii were collected. Photo taken by R. Matsumoto." figureDoi="http://doi.org/10.5281/zenodo.13757858" httpUri="https://zenodo.org/record/13757858/files/figure.png" pageId="10" pageNumber="437">Fig. 5</figureCitation>
). The specimen from
<collectingRegion id="320AF82F5244FFE25E52EBEA4E0CFE2C" box="[325,527,439,465]" country="Japan" name="Hyogo" pageId="10" pageNumber="437">Hyogo Prefecture</collectingRegion>
,
<collectingCountry id="88D9765D5244FFE25D0DEBEA4E59FE2C" box="[538,602,439,465]" name="Japan" pageId="10" pageNumber="437">Japan</collectingCountry>
was found at night on a dead walnut tree (K. Watanabe pers. com.).
</paragraph>
</subSubSection>
</treatment>
</document>

401
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<paragraph id="F07136CD5248FFEE5F80E98F4ECCFC11" blockId="6.[151,719,978,1005]" box="[151,719,978,1005]" pageId="6" pageNumber="433">
<heading id="AB3981A15248FFEE5F80E98F4ECCFC11" bold="true" box="[151,719,978,1005]" fontSize="11" level="1" pageId="6" pageNumber="433" reason="1">
<emphasis id="C2BAEADF5248FFEE5F80E98F4ECCFC11" bold="true" box="[151,719,978,1005]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5F80E98F4D1DFC10" bold="true" box="[151,286,978,1005]" italics="true" pageId="6" pageNumber="433">Vamhornia</emphasis>
<taxonomicName id="37CE4D4E5248FFEE5E31E98F4E2EFC11" authority="Hedqvist, 1976" authorityName="Hedqvist" authorityYear="1976" box="[294,557,978,1005]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="leileri">
<emphasis id="C2BAEADF5248FFEE5E31E98F4D6FFC10" bold="true" box="[294,364,978,1005]" italics="true" pageId="6" pageNumber="433">leileri</emphasis>
<bibRefCitation id="945F4B3C5248FFEE5E64E98F4E2EFC11" author="Hedqvist, K. J." box="[371,557,978,1004]" pageId="6" pageNumber="433" pagination="315 - 316" refId="ref6598" refString="Hedqvist, K. J. (1976) Vanhornia leileri n. sp. from Central Sweden (Hymenoptera: Proctotrupidae, Vanhorniinae). Entomologica Scandinavia, 7, 315 - 316. https: // doi. org / 10.1163 / 187631276 X 00540" type="journal article" year="1976">Hedqvist, 1976</bibRefCitation>
</taxonomicName>
(
<figureCitation id="68F52A485248FFEE5D2AE98F4EC4FC11" box="[573,711,978,1004]" captionStart="FIGURE 4" captionStartId="7.[152,255,1159,1183]" captionTargetBox="[158,1430,181,1135]" captionTargetId="figure-18@7.[158,1430,181,1135]" captionTargetPageId="7" captionText="FIGURE 4A–F. Vanhornia leileri (Japan, female). A, lateral habitus; B, frontal view of head; C, dorsal view of mesosoma; D, ventral view of head, prothorax, and mesothorax; E, dorsal view of metasoma; F, ventral view of metasoma. Scale bars: A—2.0 mm, C and E—0.5 mm, B, D, and F—0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13757856" httpUri="https://zenodo.org/record/13757856/files/figure.png" pageId="6" pageNumber="433">Figs. 4A–F</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B8D465465248FFEE5F80EE4A4912FBD2" box="[151,1297,1047,1072]" pageId="6" pageNumber="433" type="reference_group">
<paragraph id="F07136CD5248FFEE5F80EE4A4912FBD2" blockId="6.[151,1297,1047,1072]" box="[151,1297,1047,1072]" pageId="6" pageNumber="433">
<treatmentCitationGroup id="D0DE11E35248FFEE5F80EE4A4912FBD2" box="[151,1297,1047,1072]" pageId="6" pageNumber="433">
<treatmentCitation id="716F10DC5248FFEE5F80EE4A4E4DFBD2" author="Hedqvist, K. J." box="[151,590,1047,1071]" page="316" pageId="6" pageNumber="433" year="1976">
<taxonomicName id="37CE4D4E5248FFEE5F80EE4A4E4DFBD2" authority="Hedqvist, 1976: 315 - 316" authorityName="Hedqvist" authorityPageNumber="315 - 316" authorityYear="1976" box="[151,590,1047,1071]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="leileri">
<emphasis id="C2BAEADF5248FFEE5F80EE4A4D47FBD2" box="[151,324,1047,1071]" italics="true" pageId="6" pageNumber="433">Vanhornia leileri</emphasis>
<materialsCitation id="40A63C905248FFEE5E5DEE4A4E4DFBD2" box="[330,590,1047,1071]" country="Sweden" location="Sodermanland" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Sodermanland" typeStatus="holotype">
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.
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(examined): female; type locality:
<collectingRegion id="320AF82F5248FFEE5B0DEE4A48ACFBD2" box="[1050,1199,1047,1071]" country="Sweden" name="Sodermanland" pageId="6" pageNumber="433">Sӧdermanland</collectingRegion>
,
<collectingCountry id="88D9765D5248FFEE5BAEEE4A490EFBD2" box="[1209,1293,1047,1071]" name="Sweden" pageId="6" pageNumber="433">Sweden</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5F80EE004E2DFB1E" pageId="6" pageNumber="433" type="diagnosis">
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<emphasis id="C2BAEADF5248FFEE5F80EE004D11FB8A" bold="true" box="[151,274,1117,1143]" pageId="6" pageNumber="433">Diagnosis.</emphasis>
Frontal transverse carina present; apical margin of labrum straight. Scutoscutellar sulcus with 4–6 foveae; mesodiscrimen with about 10 strong foveae; transverse carina on dorsal part of propodeum complete. Vein Rs+M connected to Cu. Syntergite with longitudinal ridges, reaching to 0.7–0.8 length of syntergite; antero-dorsal margin of syntergite almost straight.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0EEB04932F9B6" pageId="6" pageNumber="433" type="materials_examined">
<paragraph id="F07136CD5248FFEE5FD0EEB04DC5FAD6" blockId="6.[151,1437,1117,2008]" pageId="6" pageNumber="433">
<materialsCitation id="40A63C905248FFEE5FD0EEB04DC5FAD6" collectingDate="1971-07-27" collectionCode="T, NHRS-, NRM" collectorName="Female. Sdm &amp; Tullgarn &amp; E. Leiler" pageId="6" pageNumber="433" specimenCode="HEVA 000002533" specimenCount="1" typeStatus="holotype">
<emphasis id="C2BAEADF5248FFEE5FD0EEB04D38FAFA" bold="true" box="[199,315,1261,1287]" pageId="6" pageNumber="433">
<typeStatus id="2F75886F5248FFEE5FD0EEB04D35FAFA" box="[199,310,1261,1287]" pageId="6" pageNumber="433" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="5D3B531B5248FFEE5E54EEB04DD8FAFA" box="[323,475,1261,1287]" pageId="6" pageNumber="433">Female. Sdm</collectorName>
,
<collectorName id="5D3B531B5248FFEE5EFFEEB04E49FAFA" box="[488,586,1261,1287]" pageId="6" pageNumber="433">Tullgarn</collectorName>
,
<date id="8470100D5248FFEE5D41EEB04EC8FAFA" box="[598,715,1261,1287]" pageId="6" pageNumber="433" value="1971-07-27">
<collectingDate id="9434E9E55248FFEE5D41EEB04EC8FAFA" box="[598,715,1261,1287]" pageId="6" pageNumber="433" value="1971-07-27">27/7 1971</collectingDate>
</date>
,
<collectionCode id="96DFAE085248FFEE5DC0EEB34EE5FAF5" box="[727,742,1262,1288]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="433">T</collectionCode>
.
<collectorName id="5D3B531B5248FFEE5DE1EEB34F5EFAFA" box="[758,861,1261,1288]" pageId="6" pageNumber="433">E. Leiler</collectorName>
:
<taxonomicName id="37CE4D4E5248FFEE5C7EEEB048C4FAFA" authority="K. J. Hedqist" authorityName="K. J. Hedqist" box="[873,1223,1261,1288]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="leileri">
<emphasis id="C2BAEADF5248FFEE5C7EEEB0482AFAFA" box="[873,1065,1261,1287]" italics="true" pageId="6" pageNumber="433">Vanhornia leileri</emphasis>
K. J. Hedqist
</taxonomicName>
det. 1976.
<collectionCode id="96DFAE085248FFEE5A5FEEB0499FFAFA" box="[1352,1436,1261,1287]" pageId="6" pageNumber="433">NHRS-</collectionCode>
<specimenCode id="A0689EB65248FFEE5F80EF4F4D66FAD1" box="[151,357,1297,1324]" collectionCode="HEVA" pageId="6" pageNumber="433">HEVA 000002533</specimenCode>
(
<collectionCode id="96DFAE085248FFEE5E63EF4C4DB9FAD6" box="[372,442,1297,1323]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="6" pageNumber="433" type="Museum">NRM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F07136CD5248FFEE5FD0EF684D34FA8E" blockId="6.[151,1437,1117,2008]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0EF684D4BFAB2" bold="true" box="[199,328,1333,1359]" pageId="6" pageNumber="433">Non-types.</emphasis>
<materialsCitation id="40A63C905248FFEE5E5BEF684D30FA8E" collectingDate="2011-06-12" collectingDateMax="2011-06-19" collectingDateMin="2011-06-12" collectionCode="MT, MNHN" collectorName="Claire Villemant" country="France" elevation="1" location="Piege" municipality="Chafosse" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Picardie">
<emphasis id="C2BAEADF5248FFEE5E5BEF684DA3FAB2" bold="true" box="[332,416,1333,1359]" pageId="6" pageNumber="433">
<collectingCountry id="88D9765D5248FFEE5E5BEF684DA3FAB2" box="[332,416,1333,1359]" name="France" pageId="6" pageNumber="433">France</collectingCountry>
</emphasis>
:
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,
<collectingRegion id="320AF82F5248FFEE5ECBEF684E38FAB2" box="[476,571,1333,1359]" country="France" name="Picardie" pageId="6" pageNumber="433">Picardie</collectingRegion>
, 02600 St-Pierre-Aigle,
<collectingMunicipality id="1015ACB75248FFEE5C51EF684FB3FAB2" box="[838,944,1333,1359]" pageId="6" pageNumber="433">Chafosse</collectingMunicipality>
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,
<collectionCode id="96DFAE085248FFEE5B15EF6B4829FAAD" box="[1026,1066,1334,1360]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6" pageNumber="433">MT</collectionCode>
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<date id="8470100D5248FFEE5B21EF6848DFFAB2" box="[1078,1244,1333,1360]" pageId="6" pageNumber="433" value="2011-06-12" valueMax="2011-06-19" valueMin="2011-06-12">
<collectingDate id="9434E9E55248FFEE5B21EF6848DFFAB2" box="[1078,1244,1333,1360]" pageId="6" pageNumber="433" value="2011-06-12" valueMax="2011-06-19" valueMin="2011-06-12">12–19 VI 2011</collectingDate>
</date>
,
<collectorName id="5D3B531B5248FFEE5BF2EF68499FFAB2" box="[1253,1436,1333,1359]" pageId="6" pageNumber="433">Claire Villemant</collectorName>
rec. (
<collectionCode id="96DFAE085248FFEE5FD8EF044D28FA8E" box="[207,299,1369,1395]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="433">MNHN</collectionCode>
)
</materialsCitation>
.
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12
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(
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); 1f (
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24–31
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2009
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13–20
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2010
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),
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,
<collectorName id="5D3B531B5248FFEE5C86EF984820FA22" box="[913,1059,1477,1503]" pageId="6" pageNumber="433">Hitsujigaoka</collectorName>
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<collectorName id="5D3B531B5248FFEE5F80EFB74D12F9FE" box="[151,273,1513,1540]" pageId="6" pageNumber="433">K. Konishi</collectorName>
(
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); 1f (
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13
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2015
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10
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2015
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,
<collectorName id="5D3B531B5248FFEE5B2BEFB44929F9FE" box="[1084,1322,1513,1539]" pageId="6" pageNumber="433">Maruyama-genshirin</collectorName>
,
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,
<collectorName id="5D3B531B5248FFEE5E05EC504DCBF9DA" box="[274,456,1549,1575]" pageId="6" pageNumber="433">Namiki Kikuchi</collectorName>
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,
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,
<date id="8470100D5248FFEE5BC1EC50497CF9DA" box="[1238,1407,1549,1576]" pageId="6" pageNumber="433" value="2007-07-04" valueMax="2007-07-27" valueMin="2007-07-04">
4–27
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<materialsCitation id="40A63C905248FFEE5A53EC50497CF9DA" box="[1348,1407,1549,1575]" collectionCode="SEHU" collectorName="A Ueda" country="Japan" latitude="43.605" location="Tsunemoto" longLatPrecision="6" longitude="143.344" municipality="Oketo Town" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Hokkaido">2007</materialsCitation>
</date>
,
<collectorName id="5D3B531B5248FFEE5A9DEC534CD1F9B6" pageId="6" pageNumber="433">A Ueda</collectorName>
(
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); 1f,
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,
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,
<date id="8470100D5248FFEE5C9EEC6C4813F9B6" box="[905,1040,1585,1612]" pageId="6" pageNumber="433" value="2013-08-03">
3
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2013
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,
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.
<materialsCitation id="40A63C905248FFEE5B2BEC6F492FF9B6" box="[1084,1324,1585,1612]" collectionCode="OMNH" collectorName="Matsumoto" country="Japan" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Hokkaido">
<collectorName id="5D3B531B5248FFEE5B2BEC6F48BFF9B1" box="[1084,1212,1586,1612]" pageId="6" pageNumber="433">Matsumoto</collectorName>
(
<collectionCode id="96DFAE085248FFEE5BDBEC6C4921F9B6" box="[1228,1314,1585,1611]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="6" pageNumber="433" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0EC084F75F992" box="[199,886,1621,1647]" pageId="6" pageNumber="433" type="biology_ecology">
<paragraph id="F07136CD5248FFEE5FD0EC084F75F992" blockId="6.[151,1437,1117,2008]" box="[199,886,1621,1647]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0EC084D05F992" bold="true" box="[199,262,1621,1647]" pageId="6" pageNumber="433">Host.</emphasis>
<taxonomicName id="37CE4D4E5248FFEE5E1AEC084E0EF992" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[269,525,1621,1647]" class="Insecta" family="Eucnemidae" genus="Hypocoelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="cariniceps">
<emphasis id="C2BAEADF5248FFEE5E1AEC084E0EF992" box="[269,525,1621,1647]" italics="true" pageId="6" pageNumber="433">Hypocoelus cariniceps</emphasis>
</taxonomicName>
(
<taxonomicName id="37CE4D4E5248FFEE5D0CEC084EAAF992" baseAuthorityName="Crawford" baseAuthorityYear="1909" box="[539,681,1621,1647]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
) (
<bibRefCitation id="945F4B3C5248FFEE5DD6EC084F6AF992" author="Hedqvist, K. J." box="[705,873,1621,1647]" pageId="6" pageNumber="433" pagination="315 - 316" refId="ref6598" refString="Hedqvist, K. J. (1976) Vanhornia leileri n. sp. from Central Sweden (Hymenoptera: Proctotrupidae, Vanhorniinae). Entomologica Scandinavia, 7, 315 - 316. https: // doi. org / 10.1163 / 187631276 X 00540" type="journal article" year="1976">Hedqvist 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0EC244D92F926" pageId="6" pageNumber="433" type="distribution">
<paragraph id="F07136CD5248FFEE5FD0EC244D92F926" blockId="6.[151,1437,1117,2008]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0EC244D62F96E" bold="true" box="[199,353,1657,1683]" pageId="6" pageNumber="433">Distribution.</emphasis>
<collectingCountry id="88D9765D5248FFEE5E7BEC274DBBF969" box="[364,440,1658,1684]" name="France" pageId="6" pageNumber="433">France</collectingCountry>
(new record with specimen data),
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? (Baur et al 2010),
<collectingCountry id="88D9765D5248FFEE5BA4EC2748F0F969" box="[1203,1267,1658,1684]" name="Japan" pageId="6" pageNumber="433">Japan</collectingCountry>
(
<bibRefCitation id="945F4B3C5248FFEE5A10EC244D76F94A" author="Timokhov, A. V. &amp; Belokobylskij, S. A." pageId="6" pageNumber="433" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij 2020</bibRefCitation>
),
<collectingCountry id="88D9765D5248FFEE5E91ECC04E0EF94A" box="[390,525,1693,1719]" name="Netherlands" pageId="6" pageNumber="433">Netherlands</collectingCountry>
(Belgers
<emphasis id="C2BAEADF5248FFEE5D62ECC34EA8F94A" box="[629,683,1693,1719]" italics="true" pageId="6" pageNumber="433">et al.</emphasis>
2020),
<collectingCountry id="88D9765D5248FFEE5DEAECC04F4BF94A" box="[765,840,1693,1719]" name="Russia" pageId="6" pageNumber="433">Russia</collectingCountry>
(
<bibRefCitation id="945F4B3C5248FFEE5C44ECC04FEBF94A" author="Kozlov, M. A." box="[851,1000,1693,1719]" pageId="6" pageNumber="433" pagination="675 - 677" refId="ref6713" refString="Kozlov, M. A. (1998) Fam. Vanhorniidae. In: Lehr, P. A. (Ed.), Key to insects of Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt. 3. Dal'nauka, Vladivostok, pp. 675 - 677." type="book chapter" year="1998">Kozlov 1998</bibRefCitation>
),
<collectingCountry id="88D9765D5248FFEE5CEEECC04851F94A" box="[1017,1106,1693,1719]" name="Sweden" pageId="6" pageNumber="433">Sweden</collectingCountry>
(
<bibRefCitation id="945F4B3C5248FFEE5B77ECC04906F94A" author="Hedqvist, K. J." box="[1120,1285,1693,1719]" pageId="6" pageNumber="433" pagination="315 - 316" refId="ref6598" refString="Hedqvist, K. J. (1976) Vanhornia leileri n. sp. from Central Sweden (Hymenoptera: Proctotrupidae, Vanhorniinae). Entomologica Scandinavia, 7, 315 - 316. https: // doi. org / 10.1163 / 187631276 X 00540" type="journal article" year="1976">Hedqvist 1976</bibRefCitation>
),
<collectingCountry id="88D9765D5248FFEE5A01ECC0499FF94A" box="[1302,1436,1693,1719]" name="Switzerland" pageId="6" pageNumber="433">Switzerland</collectingCountry>
(
<bibRefCitation id="945F4B3C5248FFEE5F89EC9C4D86F926" author="Artmann-Graf, G." box="[158,389,1729,1755]" pageId="6" pageNumber="433" pagination="64 - 67" refId="ref6271" refString="Artmann-Graf, G. (2016) Vanhornia leileri Hedqvist, 1976: eine fur Mitteleuropa neue Art und Familie parasitischer Wespen in der Region Olten. Oltner Neujahrsblatter, 2016, 64 - 67. https: // doi. org / 10.5169 / SEALS- 659392" type="journal article" year="2016">Artmann-Graf 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0ECB848F5F825" pageId="6" pageNumber="433" type="discussion">
<paragraph id="F07136CD5248FFEE5FD0ECB84994F8DE" blockId="6.[151,1437,1117,2008]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0ECB84D38F902" bold="true" box="[199,315,1765,1791]" pageId="6" pageNumber="433">Remarks.</emphasis>
The records from
<collectingCountry id="88D9765D5248FFEE5D1FECBB4E57F8FD" box="[520,596,1766,1792]" name="France" pageId="6" pageNumber="433">France</collectingCountry>
and
<collectingCountry id="88D9765D5248FFEE5D9EECB84EF2F902" box="[649,753,1765,1791]" name="Germany" pageId="6" pageNumber="433">Germany</collectingCountry>
reported by Baur
<emphasis id="C2BAEADF5248FFEE5CADECBB4FF2F902" box="[954,1009,1765,1791]" italics="true" pageId="6" pageNumber="433">et al.</emphasis>
(2010) are without specimen data. We newly record this species from
<collectingCountry id="88D9765D5248FFEE5EE1ED574E41F8D9" box="[502,578,1802,1828]" name="France" pageId="6" pageNumber="433">France</collectingCountry>
with specimen data. The specimen from
<collectingCountry id="88D9765D5248FFEE5B18ED544874F8DE" box="[1039,1143,1801,1827]" name="Germany" pageId="6" pageNumber="433">Germany</collectingCountry>
needs to be re-examined.
</paragraph>
<paragraph id="F07136CD5248FFEE5FD0ED7048E5F84E" blockId="6.[151,1437,1117,2008]" pageId="6" pageNumber="433">
The following morphological variations were observed: head round, upper part of gena 1.1–1.5 times as long as width of eye in lateral view; longitudinal ridges on dorsal part of propodeum weak, sometimes widened posteriorly. The male of this species was described by
<bibRefCitation id="945F4B3C5248FFEE5D61ED284803F872" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[630,1024,1909,1935]" pageId="6" pageNumber="433" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij (2020)</bibRefCitation>
. We report that the male antenna has tyloids on F1–6 that form ridges. This character may be useful to distinguish males of this species.
</paragraph>
<paragraph id="F07136CD5248FFEE5FD0EDE048F5F825" blockId="6.[151,1437,1117,2008]" box="[199,1270,1981,2008]" pageId="6" pageNumber="433">
The specimen from Oketo Town,
<collectingRegion id="320AF82F5248FFEE5D57EDE04EB2F82A" box="[576,689,1981,2007]" country="Japan" name="Hokkaido" pageId="6" pageNumber="433">Hokkaido</collectingRegion>
,
<collectingCountry id="88D9765D5248FFEE5DAAEDE34EFEF825" box="[701,765,1982,2008]" name="Japan" pageId="6" pageNumber="433">Japan</collectingCountry>
was collected near logs in the forest (
<figureCitation id="68F52A485248FFEE5BB3EDE048EBF825" box="[1188,1256,1981,2008]" captionStart="FIGURE 5" captionStartId="7.[152,255,1978,2002]" captionTargetBox="[365,1222,1311,1954]" captionTargetId="figure-75@7.[365,1222,1311,1954]" captionTargetPageId="7" captionText="FIGURE 5. Logs in the forest in Hokkaido, Japan, where Vanhornia leileri and V. yurii were collected. Photo taken by R. Matsumoto." figureDoi="http://doi.org/10.5281/zenodo.13757858" httpUri="https://zenodo.org/record/13757858/files/figure.png" pageId="6" pageNumber="433">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

198
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@ -0,0 +1,198 @@
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<subSubSection id="B8D46546524AFFEC5F80ED774819F8B9" box="[151,1050,1834,1861]" pageId="4" pageNumber="431" type="nomenclature">
<paragraph id="F07136CD524AFFEC5F80ED774819F8B9" blockId="4.[151,1050,1834,1861]" box="[151,1050,1834,1861]" pageId="4" pageNumber="431">
<heading id="AB3981A1524AFFEC5F80ED774819F8B9" bold="true" box="[151,1050,1834,1861]" fontSize="11" level="1" pageId="4" pageNumber="431" reason="1">
<emphasis id="C2BAEADF524AFFEC5F80ED774819F8B9" bold="true" box="[151,1050,1834,1861]" pageId="4" pageNumber="431">
<taxonomicName id="37CE4D4E524AFFEC5F80ED774F0EF8B9" authority="Abe, Yamagishi et Konishi" authorityName="Abe, Yamagishi et Konishi" authorityYear="2024" box="[151,781,1834,1861]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="hikosanensis" status="sp. nov.">
<emphasis id="C2BAEADF524AFFEC5F80ED774DB8F8B8" bold="true" box="[151,443,1834,1861]" italics="true" pageId="4" pageNumber="431">Vanhornia hikosanensis</emphasis>
Abe, Yamagishi et Konishi
</taxonomicName>
,
<taxonomicNameLabel id="D98957A4524AFFEC5C0DED764F7BF8B8" box="[794,888,1835,1861]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="68F52A48524AFFEC5C90ED774811F8B9" box="[903,1042,1834,1860]" captionStart="FIGURE 3" captionStartId="5.[152,255,1169,1193]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-18@5.[151,1436,181,1145]" captionTargetPageId="5" captionText="FIGURE 3A–F. Vanhornia hikosanensis (Japan, female, holotype).A, lateral habitus; B, dorso-frontal view of head; C, frontal view of head; D, dorsal view of mesosoma; E, dorsal view of propodeum; F, antero-dorsal view of metasoma. Scale bars:A—1.0 mm, B–F—0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13757852" httpUri="https://zenodo.org/record/13757852/files/figure.png" pageId="4" pageNumber="431">Figs. 3A–F</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5F80ED2F48F2F804" pageId="4" pageNumber="431" type="diagnosis">
<paragraph id="F07136CD524AFFEC5F80ED2F48F2F804" blockId="4.[151,1437,1906,2041]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5F80ED2F4D11F871" bold="true" box="[151,274,1906,1932]" pageId="4" pageNumber="431">Diagnosis.</emphasis>
Frontal transverse carina absent; apical margin of labrum weakly convex. Scutoscutellar sulcus with five foveae; mesodiscrimen with 10 strong foveae; median longitudinal carina on dorsal part of propodeum absent, smooth area on postero-lateral part of propodeum absent. Syntergite with longitudinal ridges, reaching to a maximum of 0.4 the length of syntergite, touching first thyridium; antero-dorsal margin of syntergite concave.
</paragraph>
</subSubSection>
<caption id="A4B16645524BFFED5F8FEECC4D66FB0D" ID-DOI="http://doi.org/10.5281/zenodo.13757852" ID-Zenodo-Dep="13757852" httpUri="https://zenodo.org/record/13757852/files/figure.png" pageId="5" pageNumber="432" startId="5.[152,255,1169,1193]" targetBox="[151,1436,181,1145]" targetPageId="5" targetType="figure">
<paragraph id="F07136CD524BFFED5F8FEECC4D66FB0D" blockId="5.[151,1437,1169,1265]" pageId="5" pageNumber="432">
<emphasis id="C2BAEADF524BFFED5F8FEECC4D45FB54" bold="true" box="[152,326,1169,1193]" pageId="5" pageNumber="432">FIGURE 3A–F.</emphasis>
<taxonomicName id="37CE4D4E524BFFED5E5CEECC4E3FFB54" authorityName="Abe, Yamagishi et Konishi" authorityYear="2024" box="[331,572,1169,1193]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="hikosanensis">
<emphasis id="C2BAEADF524BFFED5E5CEECC4E3FFB54" box="[331,572,1169,1193]" italics="true" pageId="5" pageNumber="432">Vanhornia hikosanensis</emphasis>
</taxonomicName>
(Japan, female, holotype). A, lateral habitus; B, dorso-frontal view of head; C, frontal view of head; D, dorsal view of mesosoma; E, dorsal view of propodeum; F, antero-dorsal view of metasoma. Scale bars: A—1.0 mm, B–F—0.2 mm.
</paragraph>
</caption>
<subSubSection id="B8D46546524BFFEE5FD0EF1E4D86FE08" lastPageId="6" lastPageNumber="433" pageId="5" pageNumber="432" type="description">
<paragraph id="F07136CD524BFFED5FD0EF1E4E76FA7C" blockId="5.[151,1437,1347,2022]" pageId="5" pageNumber="432">
<emphasis id="C2BAEADF524BFFED5FD0EF1E4D5AFAA0" bold="true" box="[199,345,1347,1373]" pageId="5" pageNumber="432">Description.</emphasis>
Female. Body length
<quantity id="37369B28524BFFED5D45EF1E4EA8FAA3" box="[594,683,1347,1374]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="432" unit="mm" value="3.8">3.8 mm</quantity>
. Color. Body black; antenna, coxae, trochanters, femora, and tibiae brown; tarsi and ovipositor reddish brown.
</paragraph>
<paragraph id="F07136CD524BFFED5FD0EFD64ED9FA34" blockId="5.[151,1437,1347,2022]" pageId="5" pageNumber="432">Antenna. Scape widened apically, 1.5 times as long as width its anterior margin, 3.5 times as long as pedicel; length to width ratio of F9–11: 1.5: 1; 1.6: 1; 2.6: 1.</paragraph>
<paragraph id="F07136CD524BFFED5FD0EF8E496AF93B" blockId="5.[151,1437,1347,2022]" pageId="5" pageNumber="432">Head. Head punctate, setose, 1.2 times as wide as its medium length and 1.2 times as wide as mesoscutum in dorsal; face 1.1 times as long as inner distance of eyes; upper part of gena 1.2 times as long as width of eye laterally; eyes oval, narrowed ventrally; POL: OOL=2.1: 1; frontal transverse carina absent; frons bulged dorsally; diameter of antennal socket 0.12 times as long as inner distance of eyes; area between antennal sockets weakly rugose; antennal scrobe present; epistomal sulcus distinct; apical margin of clypeus weakly concave, 0.3 times as wide as inner distance of eyes; malar space very short; median longitudinal sulcus on postero-dorsal part of vertex and antero-dorsal part of occiput weak. Mandible with four distinct teeth; apical margin of labrum weakly convex.</paragraph>
<paragraph id="F07136CD524BFFEE5FD0EC8D496AFF28" blockId="5.[151,1437,1347,2022]" lastBlockId="6.[151,1437,151,897]" lastPageId="6" lastPageNumber="433" pageId="5" pageNumber="432">Mesosoma. Pronotum weakly punctate; pronotal scrobe and pronotal collar rugose; dorsal margin of pronotal shoulder developed; epomia present. Anterior margin of mesopleuron rugose, ventral 1/3 of mesopleuron with setae and weakly punctate; mesopleural furrow smooth, its anterior part punctate; mesopleural carina (anterior ridge of mesopleuron) extending to postpectal carina; mesepimeral sulcus foveolate incompletely, its postero-ventral 1/5 smooth; mesodiscrimen with 10 strong foveae; mesoscutum 1.4 times as wide as long, anterior margin foveolate; notaulus foveolate, with seven foveae; parapsidial sulcus weak; scutoscutellar sulcus with five foveae; scutellum smooth, its posterior margin with some foveae; postscutellum strongly foveolate, with median longitudinal carina. Metapleuron areolate-rugose, its anterior margin strongly foveolate; metapleural smooth area present, anterior margin of the area punctate; metapleural carina present; propodeum areolate; median longitudinal carina on dorsal part of propodeum absent, transverse carina present; smooth area on postero-lateral part of propodeum absent.</paragraph>
<paragraph id="F07136CD5248FFEE5FD0EA824D3BFEBC" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="433">
Wings. Fore wing
<quantity id="37369B285248FFEE5E83EA824DEAFF04" box="[404,489,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="433" unit="mm" value="2.7">2.7 mm</quantity>
long. Vein Rs+M not connected to Cu; radial cell 4.0 times as long as wide; R (costal vein of radial cell) 1.5 times as long as width of stigma; 1cu-a interstitial to M; hind wing with two hamuli on anteroapical margin.
</paragraph>
<paragraph id="F07136CD5248FFEE5FD0EB164D4AFE2C" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="433">Metasoma. Syntergite 2.0 times as long as maximum width, some longitudinal ridges reaching 0.4 the length of syntergite, touching first thyridium; antero-dorsal margin of syntergite concave; tergite 6 punctate, its antero-dorsal margin smooth, dorso-lateral margin finely punctate; ovipositor reaching around anterior margin of mesodiscrimen.</paragraph>
<paragraph id="F07136CD5248FFEE5FD0EB864D86FE08" blockId="6.[151,1437,151,897]" box="[199,389,475,501]" pageId="6" pageNumber="433">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0EBA24F2BFDC0" pageId="6" pageNumber="433" type="materials_examined">
<paragraph id="F07136CD5248FFEE5FD0EBA24F2BFDC0" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="433">
<materialsCitation id="40A63C905248FFEE5FD0EBA24F2BFDC0" collectingDate="2022-05-28" collectingDateMax="2022-06-20" collectingDateMin="2022-05-28" collectionCode="MT, ELKU" collectorName="Junta Abe" country="Japan" county="Kyushu" elevation="650" latitude="33.482" location="Soeda-cho" longLatPrecision="72" longitude="130.908" municipality="Tagawa-gun" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Fukuoka" typeStatus="holotype">
<emphasis id="C2BAEADF5248FFEE5FD0EBA24D38FDE4" bold="true" box="[199,315,511,537]" pageId="6" pageNumber="433">
<typeStatus id="2F75886F5248FFEE5FD0EBA24D35FDE4" box="[199,310,511,537]" pageId="6" pageNumber="433" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female.
<collectingCountry id="88D9765D5248FFEE5EB0EBA24E03FDE4" box="[423,512,511,537]" name="Japan" pageId="6" pageNumber="433">JAPAN</collectingCountry>
:
<collectingCounty id="19104E415248FFEE5D1AEBA24E64FDE4" box="[525,615,511,537]" pageId="6" pageNumber="433">Kyushu</collectingCounty>
,
<collectingRegion id="320AF82F5248FFEE5D63EBA24F1BFDE4" box="[628,792,511,537]" country="Japan" name="Fukuoka" pageId="6" pageNumber="433">Fukuoka Pref.</collectingRegion>
,
<collectingMunicipality id="1015ACB75248FFEE5C32EBA24FB1FDE4" box="[805,946,511,537]" pageId="6" pageNumber="433">Tagawa-gun</collectingMunicipality>
,
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,
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,
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, alt.
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,
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,
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,
<collectorName id="5D3B531B5248FFEE5D09E87E4E8EFDC0" box="[542,653,547,573]" pageId="6" pageNumber="433">Junta Abe</collectorName>
leg. (
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).
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</paragraph>
</subSubSection>
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<paragraph id="F07136CD5248FFEE5FD0E81A4D80FD9C" blockId="6.[151,1437,151,897]" box="[199,387,583,609]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0E81A4D05FD9C" bold="true" box="[199,262,583,609]" pageId="6" pageNumber="433">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0E8374DACFD78" box="[199,431,618,645]" pageId="6" pageNumber="433" type="distribution">
<paragraph id="F07136CD5248FFEE5FD0E8374DACFD78" blockId="6.[151,1437,151,897]" box="[199,431,618,645]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0E8374D62FD79" bold="true" box="[199,353,618,644]" pageId="6" pageNumber="433">Distribution.</emphasis>
<collectingCountry id="88D9765D5248FFEE5E70E8364DA9FD78" box="[359,426,619,645]" name="Japan" pageId="6" pageNumber="433">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0E8D24900FD54" box="[199,1283,655,681]" pageId="6" pageNumber="433" type="etymology">
<paragraph id="F07136CD5248FFEE5FD0E8D24900FD54" blockId="6.[151,1437,151,897]" box="[199,1283,655,681]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0E8D24D4DFD54" bold="true" box="[199,334,655,681]" pageId="6" pageNumber="433">Etymology.</emphasis>
The specific epithet is derived from the
<typeStatus id="2F75886F5248FFEE5C04E8D24F40FD54" box="[787,835,655,681]" pageId="6" pageNumber="433">type</typeStatus>
locality, “Hiko-san” called in Japanese.
</paragraph>
</subSubSection>
<subSubSection id="B8D465465248FFEE5FD0E8EE4947FC7C" pageId="6" pageNumber="433" type="discussion">
<paragraph id="F07136CD5248FFEE5FD0E8EE4947FC7C" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="433">
<emphasis id="C2BAEADF5248FFEE5FD0E8EE4D38FD30" bold="true" box="[199,315,691,717]" pageId="6" pageNumber="433">Remarks.</emphasis>
This new species is closely related to
<taxonomicName id="37CE4D4E5248FFEE5DE7E8EE4FB1FD30" authorityName="Crawford" authorityYear="1909" box="[752,946,691,717]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5248FFEE5DE7E8EE4FB1FD30" box="[752,946,691,717]" italics="true" pageId="6" pageNumber="433">V. eucnemidarum</emphasis>
</taxonomicName>
in having following characteristics: frontal transverse carina absent; scutoscutellar sulcus with five foveae; mesodiscrimen with about 10 foveae; ridges on syntergite touching first thyridium. However, it can be distinguished by following characteristics: apical margin of clypeus weakly convex (straight in
<taxonomicName id="37CE4D4E5248FFEE5D5FE9424F08FCC4" authorityName="Crawford" authorityYear="1909" box="[584,779,799,825]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5248FFEE5D5FE9424F08FCC4" box="[584,779,799,825]" italics="true" pageId="6" pageNumber="433">V. eucnemidarum</emphasis>
</taxonomicName>
); median longitudinal carina on dorsal part of propodeum absent (present in
<taxonomicName id="37CE4D4E5248FFEE5E64E91E4E3AFCA0" authorityName="Crawford" authorityYear="1909" box="[371,569,835,861]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5248FFEE5E64E91E4E3AFCA0" box="[371,569,835,861]" italics="true" pageId="6" pageNumber="433">V. eucnemidarum</emphasis>
</taxonomicName>
); smooth area on postero-lateral part of propodeum absent (present in
<taxonomicName id="37CE4D4E5248FFEE5A9FE91E4D3EFC7C" authorityName="Crawford" authorityYear="1909" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5248FFEE5A9FE91E4D3EFC7C" italics="true" pageId="6" pageNumber="433">V. eucnemidarum</emphasis>
</taxonomicName>
); antero-dorsal margin of syntergite strongly concave (almost straight in
<taxonomicName id="37CE4D4E5248FFEE5B65E93A4930FC7C" authorityName="Crawford" authorityYear="1909" box="[1138,1331,871,897]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF5248FFEE5B65E93A4930FC7C" box="[1138,1331,871,897]" italics="true" pageId="6" pageNumber="433">V. eucnemidarum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

280
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<mods:title id="437306709CC2010D4B866AC8D46926A4">Review of the family Vanhorniidae (Hymenoptera: Proctotrupoidea), with the description of two new species</mods:title>
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<subSubSection id="B8D46546524CFFEA5F80EB0A4F46FE8C" box="[151,837,343,370]" pageId="2" pageNumber="429" type="nomenclature">
<paragraph id="F07136CD524CFFEA5F80EB0A4F46FE8C" blockId="2.[151,837,343,370]" box="[151,837,343,370]" pageId="2" pageNumber="429">
<heading id="AB3981A1524CFFEA5F80EB0A4F46FE8C" bold="true" box="[151,837,343,370]" fontSize="11" level="1" pageId="2" pageNumber="429" reason="1">
<emphasis id="C2BAEADF524CFFEA5F80EB0A4F46FE8C" bold="true" box="[151,837,343,370]" pageId="2" pageNumber="429">
<taxonomicName id="37CE4D4E524CFFEA5F80EB0A4EA0FE8F" authority="Crawford, 1909" authorityName="Crawford" authorityYear="1909" box="[151,675,343,370]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF524CFFEA5F80EB0A4DD5FE8F" bold="true" box="[151,470,343,370]" italics="true" pageId="2" pageNumber="429">Vanhornia eucnemidarum</emphasis>
<bibRefCitation id="945F4B3C524CFFEA5ECAEB054EA0FE8F" author="Crawford, J." box="[477,675,344,370]" pageId="2" pageNumber="429" pagination="63 - 64" refId="ref6416" refString="Crawford, J. (1909) A new family of parasitica Hymenoptara. Proceedings of the Entomological Society of Washington, 11, 63 - 64." type="journal article" year="1909">Crawford, 1909</bibRefCitation>
</taxonomicName>
(
<figureCitation id="68F52A48524CFFEA5DA5EB0A4F3FFE8C" box="[690,828,343,369]" captionStart="FIGURE 1" captionStartId="2.[152,255,1628,1652]" captionTargetBox="[151,1435,638,1602]" captionTargetId="figure-197@2.[151,1435,638,1602]" captionTargetPageId="2" captionText="FIGURE 1A–F. Vanhornia eucnemidarum (USA, female). A, lateral habitus; B, ventral view of head, arrow indicates apical margin of labrum; C, dorsal view of pronotum and mesothorax; D, ventral view of mesodiscrimen; E, dorsal view of propodeum; F, antero-dorsal view of metasoma. Scale bars: A—2.0 mm, B–F—0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13757844" httpUri="https://zenodo.org/record/13757844/files/figure.png" pageId="2" pageNumber="429">Figs. 1A–F</figureCitation>
)
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<paragraph id="F07136CD524CFFEA5F80EBC14932FE49" blockId="2.[151,1329,412,437]" box="[151,1329,412,437]" pageId="2" pageNumber="429">
<treatmentCitationGroup id="D0DE11E3524CFFEA5F80EBC14932FE49" box="[151,1329,412,437]" pageId="2" pageNumber="429">
<treatmentCitation id="716F10DC524CFFEA5F80EBC14E96FE49" author="Crawford, J." box="[151,661,412,436]" page="64" pageId="2" pageNumber="429" year="1909">
<taxonomicName id="37CE4D4E524CFFEA5F80EBC14E96FE49" authority="Crawford, 1909: 63 - 64" authorityName="Crawford" authorityPageNumber="63 - 64" authorityYear="1909" box="[151,661,412,436]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF524CFFEA5F80EBC14D9DFE49" box="[151,414,412,436]" italics="true" pageId="2" pageNumber="429">Vanhornia eucnemidarum</emphasis>
<materialsCitation id="40A63C90524CFFEA5EB3EBC14E96FE49" box="[420,661,412,436]" country="United States of America" location="Maryland" pageId="2" pageNumber="429" specimenCount="1" stateProvince="Maryland" typeStatus="holotype">
<bibRefCitation id="945F4B3C524CFFEA5EB3EBC14E96FE49" author="Crawford, J." box="[420,661,412,436]" pageId="2" pageNumber="429" pagination="63 - 64" refId="ref6416" refString="Crawford, J. (1909) A new family of parasitica Hymenoptara. Proceedings of the Entomological Society of Washington, 11, 63 - 64." type="journal article" year="1909">Crawford, 1909: 63–64</bibRefCitation>
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.
<typeStatus id="2F75886F524CFFEA5D89EBC14EFFFE49" box="[670,764,412,436]" pageId="2" pageNumber="429" type="holotype">Holotype</typeStatus>
(not examined): female; type locality:
<collectingRegion id="320AF82F524CFFEA5B90EBC148EFFE49" box="[1159,1260,412,436]" country="United States of America" name="Maryland" pageId="2" pageNumber="429">Maryland</collectingRegion>
,
<collectingCountry id="88D9765D524CFFEA5BE0EBC1492EFE49" box="[1271,1325,412,436]" name="United States of America" pageId="2" pageNumber="429">USA</collectingCountry>
.
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<emphasis id="C2BAEADF524CFFEA5F80EBBF4D11FE01" bold="true" box="[151,274,482,508]" pageId="2" pageNumber="429">Diagnosis.</emphasis>
Frontal transverse carina absent; apical margin of labrum straight. Scutoscutellar sulcus with five foveae; mesodiscrimen with about 10 strong foveae; median longitudinal carina on dorsal part of propodeum present, smooth area on postero-lateral part of propodeum present. Syntergite with longitudinal ridges, reaching to a maximum of 0.4 length of syntergite, touching first thyridium; antero-dorsal margin of syntergite almost straight.
</paragraph>
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<paragraph id="F07136CD524CFFEA5F8FEC014F96F946" blockId="2.[151,1437,1628,1724]" pageId="2" pageNumber="429">
<emphasis id="C2BAEADF524CFFEA5F8FEC014D44F989" bold="true" box="[152,327,1628,1652]" pageId="2" pageNumber="429">FIGURE 1A–F.</emphasis>
<taxonomicName id="37CE4D4E524CFFEA5E58EC014E55F989" authorityName="Crawford" authorityYear="1909" box="[335,598,1628,1652]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF524CFFEA5E58EC014E55F989" box="[335,598,1628,1652]" italics="true" pageId="2" pageNumber="429">Vanhornia eucnemidarum</emphasis>
</taxonomicName>
(USA, female). A, lateral habitus; B, ventral view of head, arrow indicates apical margin of labrum; C, dorsal view of pronotum and mesothorax; D, ventral view of mesodiscrimen; E, dorsal view of propodeum; F, antero-dorsal view of metasoma. Scale bars: A—2.0 mm, B–F—0.2 mm.
</paragraph>
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<emphasis id="C2BAEADF524CFFEA5FD0ED534E40F8D5" bold="true" box="[199,579,1806,1832]" pageId="2" pageNumber="429">Specimen examined. Non-types.</emphasis>
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: 1f,
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,
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,
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16 VII
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.
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(ELMU); 1f,
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,
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,
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19 VII
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, B. E.
<collectorName id="5D3B531B524CFFEA5DB9ED6F4F03F8B1" box="[686,768,1842,1868]" pageId="2" pageNumber="429">Cooper</collectorName>
(ELMU).
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<emphasis id="C2BAEADF524CFFEA5C61ED6F4FADF8B1" bold="true" box="[886,942,1842,1868]" pageId="2" pageNumber="429">
<collectingCountry id="88D9765D524CFFEA5C61ED6F4FADF8B1" box="[886,942,1842,1868]" name="United States of America" pageId="2" pageNumber="429">USA</collectingCountry>
</emphasis>
: 1f (
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15–21
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2011
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</date>
), 4f (
<date id="8470100D524CFFEA5BA9ED6F4962F8B1" box="[1214,1377,1842,1869]" pageId="2" pageNumber="429" value="2011-05-22" valueMax="2011-05-28" valueMin="2011-05-22">
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22–28
<collectionCode id="96DFAE08524CFFEA5A1BED6E4923F8B0" box="[1292,1312,1843,1869]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="429" type="Museum">V</collectionCode>
2011
</collectingDate>
</date>
), and 2f (
<date id="8470100D524CFFEA5FA9ED0B4D79F88D" box="[190,378,1878,1905]" pageId="2" pageNumber="429" value="2011-05-29" valueMax="2011-06-04" valueMin="2011-05-29">
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29
<collectionCode id="96DFAE08524CFFEA5FF4ED0B4CF4F88D" box="[227,247,1878,1904]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="429" type="Museum">V</collectionCode>
–4
<collectionCode id="96DFAE08524CFFEA5E0DED0A4D3BF88C" box="[282,312,1879,1905]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="429">VI</collectionCode>
2011
</collectingDate>
</date>
), VA,
<collectingCounty id="19104E41524CFFEA5ED6ED0B4E47F88D" box="[449,580,1878,1904]" pageId="2" pageNumber="429">Fairfax Co.</collectingCounty>
,
<collectorName id="5D3B531B524CFFEA5D47ED0B4EE2F88D" box="[592,737,1878,1904]" pageId="2" pageNumber="429">Holmes Run</collectorName>
~1/
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NE, jct,
<collectorName id="5D3B531B524CFFEA5C82ED0B4818F88D" box="[917,1051,1878,1904]" pageId="2" pageNumber="429">Gallows Rd</collectorName>
&amp; I-495,
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,
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,
<collectionCode id="96DFAE08524CFFEA5A7AED0A4996F88C" box="[1389,1429,1879,1905]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="429">MT</collectionCode>
</materialsCitation>
, D. R.
<materialsCitation id="40A63C90524CFFEA5FC9ED274CBCF844" collectionCode="SEHU, MT" collectorName="Smith" country="United States of America" county="Prince William Co." latitude="38.826668" location="Chestnut Ridge Trail" longLatPrecision="117" longitude="-77.706665" municipality="Bull Run Mtns" pageId="2" pageNumber="429" specimenCount="1">
<collectorName id="5D3B531B524CFFEA5FC9ED274D22F869" box="[222,289,1914,1940]" pageId="2" pageNumber="429">Smith</collectorName>
(
<collectionCode id="96DFAE08524CFFEA5E25ED274D7EF869" box="[306,381,1914,1940]" pageId="2" pageNumber="429">SEHU</collectionCode>
); 1f, VA,
<collectingCounty id="19104E41524CFFEA5EE5ED274ED3F869" box="[498,720,1914,1940]" pageId="2" pageNumber="429">Prince William Co.</collectingCounty>
,
<collectingMunicipality id="1015ACB7524CFFEA5DC9ED274F88F868" box="[734,907,1914,1941]" pageId="2" pageNumber="429">Bull Run Mtns</collectingMunicipality>
,
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,
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,
<geoCoordinate id="95FA500A524CFFEA5A0BED27499BF869" box="[1308,1432,1914,1940]" degrees="77" direction="west" minutes="42.4" orientation="longitude" pageId="2" pageNumber="429" precision="92" value="-77.706665">77°42.4’W</geoCoordinate>
,
<collectionCode id="96DFAE08524CFFEA5F80EDC24CBCF844" box="[151,191,1951,1977]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="429">MT</collectionCode>
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,
<date id="8470100D524CFFEA5FDAEDC34D95F845" box="[205,406,1950,1977]" pageId="2" pageNumber="429" value="2011-06-25" valueMax="2011-07-07" valueMin="2011-06-25">25 VI–7 VII 2011</date>
, D. R.
<materialsCitation id="40A63C90524CFFEA5EF5EDC34855F844" box="[482,1110,1950,1977]" collectionCode="SEHU, MT" collectorName="Smith &amp; Annandale" country="United States of America" county="Fairfax Co." location="Fairfax Co." pageId="2" pageNumber="429" specimenCount="1">
<collectorName id="5D3B531B524CFFEA5EF5EDC34E26F845" box="[482,549,1950,1976]" pageId="2" pageNumber="429">Smith</collectorName>
(
<collectionCode id="96DFAE08524CFFEA5D24EDC34E7DF845" box="[563,638,1950,1976]" pageId="2" pageNumber="429">SEHU</collectionCode>
); 2f, VA,
<collectingCounty id="19104E41524CFFEA5DFBEDC34F6FF845" box="[748,876,1950,1976]" pageId="2" pageNumber="429">Fairfax Co.</collectingCounty>
near
<collectorName id="5D3B531B524CFFEA5CB0EDC34820F845" box="[935,1059,1950,1976]" pageId="2" pageNumber="429">Annandale</collectorName>
,
<collectionCode id="96DFAE08524CFFEA5B39EDC24855F844" box="[1070,1110,1951,1977]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="429">MT</collectionCode>
</materialsCitation>
,
<date id="8470100D524CFFEA5B73EDC348E7F845" box="[1124,1252,1950,1977]" pageId="2" pageNumber="429" value="1988-06-11">
11 VI
<materialsCitation id="40A63C90524CFFEA5BBEEDC348E7F845" box="[1193,1252,1950,1976]" collectionCode="R" collectorName="David" country="United States of America" pageId="2" pageNumber="429" specimenCount="1">1988</materialsCitation>
</date>
,
<collectorName id="5D3B531B524CFFEA5BF9EDC34931F845" box="[1262,1330,1950,1976]" pageId="2" pageNumber="429">David</collectorName>
R.
<materialsCitation id="40A63C90524CFFEA5A4EEDC34CFAF821" collectionCode="NSMT" collectorName="Smith" country="United States of America" pageId="2" pageNumber="429" specimenCount="1">
<collectorName id="5D3B531B524CFFEA5A4EEDC3499FF845" box="[1369,1436,1950,1976]" pageId="2" pageNumber="429">Smith</collectorName>
(
<collectionCode id="96DFAE08524CFFEA5F88ED9F4CECF821" box="[159,239,1986,2012]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="2" pageNumber="429">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524DFFEB5FD0EACA48D4FEBC" pageId="3" pageNumber="430" type="biology_ecology">
<paragraph id="F07136CD524DFFEB5FD0EACA48D4FEBC" blockId="3.[151,1436,151,321]" pageId="3" pageNumber="430">
<emphasis id="C2BAEADF524DFFEB5FD0EACA4D05FF4C" bold="true" box="[199,262,151,177]" pageId="3" pageNumber="430">Host.</emphasis>
<taxonomicName id="37CE4D4E524DFFEB5E1AEACA4F1AFF4C" authority="(Say, 1823) (Muona 2021)" authorityName="Muona" authorityYear="2021" baseAuthorityName="Say" baseAuthorityYear="1823" box="[269,793,151,177]" class="Insecta" family="Eucnemidae" genus="Isorhipis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="C2BAEADF524DFFEB5E1AEACA4DE6FF4C" box="[269,485,151,177]" italics="true" pageId="3" pageNumber="430">Isorhipis ruficornis</emphasis>
(Say, 1823) (
<bibRefCitation id="945F4B3C524DFFEB5D6BEACA4F13FF4C" author="Muona, J." box="[636,784,151,177]" pageId="3" pageNumber="430" pagination="1 - 10" refId="ref6811" refString="Muona, J. (2021) How do Vanhorniidae (Hymenoptera) parasitize Eucnemidae (Coleoptera)? Insecta Mundi, 0867, 1 - 10." type="book chapter" year="2021">Muona 2021</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="C2BAEADF524DFFEB5FD0EAE74D62FF29" bold="true" box="[199,353,186,212]" pageId="3" pageNumber="430">Distribution.</emphasis>
<collectingCountry id="88D9765D524DFFEB5E70EAE64DB8FF28" box="[359,443,187,213]" name="Canada" pageId="3" pageNumber="430">Canada</collectingCountry>
(
<bibRefCitation id="945F4B3C524DFFEB5EDBEAE64E9BFF28" author="Hogan, J. &amp; Bass, A. &amp; Zhang, Y. M. &amp; Sharanowski, B." box="[460,664,187,213]" pageId="3" pageNumber="430" refId="ref6645" refString="Hogan, J., Bass, A., Zhang, Y. M. &amp; Sharanowski, B. (2019) Integrating multiple sources of biodiversity information greatly expands the range of a rare species of Hymenoptera (Vanhorniidae). Biodiversity Data Journal, 7, e 37569. https: // doi. org / 10.3897 / BDJ. 7. e 37569" type="journal volume" year="2019">
Hogan
<emphasis id="C2BAEADF524DFFEB5D0AEAE64E55FF28" box="[541,598,187,213]" italics="true" pageId="3" pageNumber="430">et al.</emphasis>
2019
</bibRefCitation>
),
<collectingCountry id="88D9765D524DFFEB5DBBEAE64EE1FF28" box="[684,738,187,213]" name="United States of America" pageId="3" pageNumber="430">USA</collectingCountry>
(
<bibRefCitation id="945F4B3C524DFFEB5DE5EAE64868FF28" author="Timokhov, A. V. &amp; Belokobylskij, S. A." box="[754,1131,187,213]" pageId="3" pageNumber="430" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij 2020</bibRefCitation>
).
<emphasis id="C2BAEADF524DFFEB5FD0EA824D38FF04" bold="true" box="[199,315,223,249]" pageId="3" pageNumber="430">Remarks.</emphasis>
This species was erroneously recorded from
<collectingCountry id="88D9765D524DFFEB5C76EA824FF7FF04" box="[865,1012,223,249]" name="South Korea" pageId="3" pageNumber="430">South Korea</collectingCountry>
(
<bibRefCitation id="945F4B3C524DFFEB5B1EEA8248E5FF04" author="Choi, M. B. &amp; Lee, J. W." box="[1033,1254,223,249]" pageId="3" pageNumber="430" pagination="59 - 61" refId="ref6362" refString="Choi, M. B. &amp; Lee, J. W. (2012) First record of Vanhorniidae (Hymenoptera: Proctotrupoidea) from Korea. Journal of Asia- Pacific Entomology, 15, 59 - 61. https: // doi. org / 10.1016 / j. aspen. 2011.08.003" type="journal article" year="2012">Choi &amp; Lee 2012</bibRefCitation>
). We examined the voucher specimen deposited in YNU and identified it as
<taxonomicName id="37CE4D4E524DFFEB5C40EB5E4FA8FEE1" authorityName="Timokhov &amp; Belokobylskij" authorityYear="2020" box="[855,939,259,284]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="yurii">
<emphasis id="C2BAEADF524DFFEB5C40EB5E4FA8FEE1" box="[855,939,259,284]" italics="true" pageId="3" pageNumber="430">V. yurii</emphasis>
</taxonomicName>
(as preliminary suggested by
<bibRefCitation id="945F4B3C524DFFEB5A1EEB5E4D7CFEBC" author="Timokhov, A. V. &amp; Belokobylskij, S. A." pageId="3" pageNumber="430" pagination="57 - 76" refId="ref6839" refString="Timokhov, A. V. &amp; Belokobylskij, S. A. (2020) Review of the rare genus Vanhornia Crawford, 1909 (Hymenoptera, Proctotrupoidea, Vanhorniidae) with description of a new species from the Russian Far East. Journal of Hymenoptera Research, 79, 57 - 76. https: // doi. org / 10.3897 / jhr. 79.56481" type="journal article" year="2020">Timokhov &amp; Belokobylskij, 2020</bibRefCitation>
). Thus,
<taxonomicName id="37CE4D4E524DFFEB5EC0EB7A4E9AFEBC" authorityName="Crawford" authorityYear="1909" box="[471,665,295,321]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="eucnemidarum">
<emphasis id="C2BAEADF524DFFEB5EC0EB7A4E9AFEBC" box="[471,665,295,321]" italics="true" pageId="3" pageNumber="430">V. eucnemidarum</emphasis>
</taxonomicName>
is not part of the vanhorniid fauna of
<collectingCountry id="88D9765D524DFFEB5B55EB7A48D0FEBC" box="[1090,1235,295,321]" name="South Korea" pageId="3" pageNumber="430">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

191
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<mods:role id="5DAB2E9AC82B46BA19ABFA581BF6BF52">
<mods:roleTerm id="1D3FE5F354F4C1EC6D59766536E0E511">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62DBC23183791FF0A6ACC6FA0566F1EE">Yamagishi, Kenzo</mods:namePart>
<mods:nameIdentifier id="B3E0975FD2308C932348FF927A463613" type="ORCID">0009-0001-0322-5745</mods:nameIdentifier>
<mods:nameIdentifier id="B7704F6BB5BCA32B543218CEF80B4BA7" type="email">kentatsu@tg.commufa.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="7660CA4E0337A8075082C83886B1F7FC" type="personal">
<mods:role id="1BC690479D986989B95646A836C70F62">
<mods:roleTerm id="C526BB1765F26D95EA7779E591F506D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8718ABEF096C4804871D88DE07DF65B2">Konishi, Kazuhiko</mods:namePart>
<mods:affiliation id="B089FC171A1FCAF5A42827C3EDF18201">Hokkaido University Museum, Kita 10 - jo, Nishi 8 - chome, Kita-ku, Sapporo, Hokkaido 060 - 0810 Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3D9D5D047EE93E8F7A8E4CA723CD2B9E">text</mods:typeOfResource>
<mods:relatedItem id="69876909FFE1F1CA6970BC7C81851FE9" type="host">
<mods:titleInfo id="93406AA576E0C4342E4EEF40EAE4D55E">
<mods:title id="4903309B84C886BE5E35F4EC37C183DA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9D9F4925709483BFEE4450EB3F6B9484">
<mods:date id="9EC5B3BE2F4A1CEDFBF5609FA74E86FC">2024</mods:date>
<mods:detail id="F1ABCAC9ADC6B95BFDA586884F56CAD8" type="pubDate">
<mods:number id="DF4B6A38B488C227A431B446F78B0EF5">2024-09-13</mods:number>
</mods:detail>
<mods:detail id="03CCB708B0AB7C0A76607869B1C6AF45" type="volume">
<mods:number id="DD685C65EAF28A328DAF8D4DC261AD2C">5507</mods:number>
</mods:detail>
<mods:detail id="7892AD7AF9C47528FD64220F4F200929" type="issue">
<mods:number id="1E19E5EF2A497A8D4917035719A508BB">3</mods:number>
</mods:detail>
<mods:extent id="165849C7944058137D1091725353D2CA" unit="page">
<mods:start id="82365D91B5DA93C1DCD837AC94B6A4C1">427</mods:start>
<mods:end id="CC1EF7DDF85E25E571D2E51F79B828E5">438</mods:end>
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<mods:location id="9C4B3D5F296CD35E80BB1B6B832344A6">
<mods:url id="145A6395004603F44902A0F75AD164DA">http://dx.doi.org/10.11646/zootaxa.5507.3.2</mods:url>
</mods:location>
<mods:classification id="3B69EFB051D0523031061AA629FC4CB6">journal article</mods:classification>
<mods:identifier id="C1F26D312CA7C678191E5F6449DF9F33" type="DOI">10.11646/zootaxa.5507.3.2</mods:identifier>
<mods:identifier id="B9992349C4B9F605EAF430ED413D2BFA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B2988F9A0BDA2F283AC81C553C97BB9E" type="Zenodo-Dep">13757842</mods:identifier>
<mods:identifier id="14A18A4E45C1434622FAE5C8D0F15C17" type="ZooBank">873ABA0C-039A-4B68-84AC-98E505D95A92</mods:identifier>
</mods:mods>
<treatment id="786787DB524DFFEC5F80EBD74EF2F91C" ID-DOI="http://doi.org/10.5281/zenodo.13757850" ID-Zenodo-Dep="13757850" LSID="urn:lsid:plazi:treatment:786787DB524DFFEC5F80EBD74EF2F91C" httpUri="http://treatment.plazi.org/id/786787DB524DFFEC5F80EBD74EF2F91C" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430">
<subSubSection id="B8D46546524DFFEB5F80EBD74807FE59" box="[151,1028,394,421]" pageId="3" pageNumber="430" type="nomenclature">
<paragraph id="F07136CD524DFFEB5F80EBD74807FE59" blockId="3.[151,1028,394,421]" box="[151,1028,394,421]" pageId="3" pageNumber="430">
<heading id="AB3981A1524DFFEB5F80EBD74807FE59" bold="true" box="[151,1028,394,421]" fontSize="11" level="1" pageId="3" pageNumber="430" reason="1">
<emphasis id="C2BAEADF524DFFEB5F80EBD74807FE59" bold="true" box="[151,1028,394,421]" pageId="3" pageNumber="430">
<taxonomicName id="37CE4D4E524DFFEB5F80EBD74EF0FE59" authority="Abe, Yamagishi et Konishi" authorityName="Abe, Yamagishi et Konishi" authorityYear="2024" box="[151,755,394,421]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="formosana" status="sp. nov.">
<emphasis id="C2BAEADF524DFFEB5F80EBD74DA2FE59" bold="true" box="[151,417,394,421]" italics="true" pageId="3" pageNumber="430">Vanhornia formosana</emphasis>
Abe, Yamagishi et Konishi
</taxonomicName>
,
<taxonomicNameLabel id="D98957A4524DFFEB5C17EBD64F5EFE58" box="[768,861,395,421]" pageId="3" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="68F52A48524DFFEB5C7AEBD74FFEFE59" box="[877,1021,394,420]" captionStart="FIGURE 2" captionStartId="3.[152,255,1901,1925]" captionTargetBox="[183,1403,657,1868]" captionTargetId="figure-180@3.[183,1403,657,1877]" captionTargetPageId="3" captionText="FIGURE 2A–F. Vanhornia formosana (Taiwan, female, holotype). A, lateral habitus; B, anterior view of head, arrow indicates frontal transverse carina; C, dorsal view of head; D, dorsal view of mesosoma; E, dorsal view of propodeum, arrow indicates transverse carina (incomplete); F, antero-dorsal view of metasoma, arrow indicates first thyridium; G, wings. Scale bars:A—1.0 mm, B–F—0.2 mm, G—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757848" httpUri="https://zenodo.org/record/13757848/files/figure.png" pageId="3" pageNumber="430">Figs. 2A–G</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524DFFEB5F80EB9348F6FD84" pageId="3" pageNumber="430" type="diagnosis">
<paragraph id="F07136CD524DFFEB5F80EB9348F6FD84" blockId="3.[151,1437,462,633]" pageId="3" pageNumber="430">
<emphasis id="C2BAEADF524DFFEB5F80EB934D09FE15" bold="true" box="[151,266,462,488]" pageId="3" pageNumber="430">Diagnosis</emphasis>
. Punctures between median ocellus and frontal transverse carina large, larger than punctures on anterior half of head; frontal transverse carina present; apical margin of labrum convex. Scutoscutellar sulcus with six foveae; mesodiscrimen with six strong foveae; transverse carina on dorsal part of propodeum incomplete; postero-lateral part of propodeum with smooth area. Syntergite with longitudinal ridges, reaching to a maximum of 0.4 length of syntergite, not touching first thyridium; antero-dorsal margin of syntergite almost straight.
</paragraph>
</subSubSection>
<caption id="A4B16645524DFFEB5F8FED304DE8F80C" ID-DOI="http://doi.org/10.5281/zenodo.13757848" ID-Zenodo-Dep="13757848" httpUri="https://zenodo.org/record/13757848/files/figure.png" pageId="3" pageNumber="430" startId="3.[152,255,1901,1925]" targetBox="[183,1403,657,1868]" targetPageId="3" targetType="figure">
<paragraph id="F07136CD524DFFEB5F8FED304DE8F80C" blockId="3.[151,1436,1901,2033]" pageId="3" pageNumber="430">
<emphasis id="C2BAEADF524DFFEB5F8FED304D45F878" bold="true" box="[152,326,1901,1925]" pageId="3" pageNumber="430">FIGURE 2A–F.</emphasis>
<taxonomicName id="37CE4D4E524DFFEB5E5CED304E24F878" authorityName="Abe, Yamagishi et Konishi" authorityYear="2024" box="[331,551,1901,1925]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C2BAEADF524DFFEB5E5CED304E24F878" box="[331,551,1901,1925]" italics="true" pageId="3" pageNumber="430">Vanhornia formosana</emphasis>
</taxonomicName>
(Taiwan, female, holotype). A, lateral habitus; B, anterior view of head, arrow indicates frontal transverse carina; C, dorsal view of head; D, dorsal view of mesosoma; E, dorsal view of propodeum, arrow indicates transverse carina (incomplete); F, antero-dorsal view of metasoma, arrow indicates first thyridium; G, wings. Scale bars: A—1.0 mm, B–F—0.2 mm, G—0.5 mm.
</paragraph>
</caption>
<subSubSection id="B8D46546524AFFEC5FD0EACB4D86FACC" pageId="4" pageNumber="431" type="description">
<paragraph id="F07136CD524AFFEC5FD0EACB485AFF28" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5FD0EACB4D5AFF4D" bold="true" box="[199,345,150,176]" pageId="4" pageNumber="431">Description.</emphasis>
Female. Body length
<quantity id="37369B28524AFFEC5D51EACB4E9EFF4C" box="[582,669,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="431" unit="mm" value="5.1">5.1 mm</quantity>
. Color. Body black; scape, apical part of flagellum, coxae, trochanters, and femora brown; basal part of flagellum, tibiae, tarsi, and ovipositor reddish brown.
</paragraph>
<paragraph id="F07136CD524AFFEC5FD0EA824ED9FEE0" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">Antenna. Scape widened apically, 1.3 times as long as width of its anterior margin, 2.8 times as long as pedicel; length to width ratio of F9–11: 1.8: 1; 1.9: 1; 2.8: 1.</paragraph>
<paragraph id="F07136CD524AFFEC5FD0EB7A4D67FD9C" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">Head. Head punctate, with sparse setae, 1.3 times as wide as its medium length and 1.1 times as wide as mesoscutum in dorsal view; face 1.1 times as long as inner distance of eyes; upper part of gena 1.7 times as long as width of eye laterally; eyes suboval, slightly narrowed ventrally; POL: OOL=1.5: 1; frontal transverse carina present; frons bulged in dorsal view, punctures between median ocellus and frontal transverse carina large, larger than punctures on anterior half of head; diameter of antennal socket 0.13 times as long as inner distance of eyes, area between antennal sockets rugose; antennal scrobe present; epistomal sulcus distinct; apical margin of clypeus weakly concave, 0.4 times as wide as inner distance of eyes; malar space very short; median longitudinal sulcus on postero-dorsal part of vertex and antero-dorsal part of occiput weak. Mandible with five distinct teeth; apical margin of labrum convex.</paragraph>
<paragraph id="F07136CD524AFFEC5FD0E8364CCEFBEC" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">Mesosoma. Dorsal and postero-lateral part of pronotum punctate, antero-lateral part of pronotum rugose; epomia present; pronotal collar punctate, sparsely setose; dorso-lateral margin of pronotal shoulder developed. Anterior margin of mesopleuron rugose, ventral part of mesopleuron weakly punctate, with setae; mesopleural carina (anterior ridge of mesopleuron) extending to postpectal carina; mesopleural furrow smooth, its anterior part weakly punctate; mesepimeral sulcus strongly foveolate; mesodiscrimen with six strong foveae; mesoscutum punctate, 1.6 times as wide as long; anterior margin foveolate; notaulus with seven foveae; parapsidial sulcus weakly indicated; axilla foveolate; scutoscutellar sulcus with six foveae; lateral margin of scutellum weakly foveolate, posterior margin with some foveae; postscutellum foveolate, with median longitudinal carina; dorsal part of metanotum scrobiculate. Metapleuron areolate-rugose, with small smooth area; metapleural carina (carina connecting metapleural smooth area and dorso-anterior margin of propodeum) present. Dorsal part of propodeum with three longitudinal carinae, transverse carina incomplete; posterior part of propodeum areolate; postero-lateral part of propodeum with smooth area.</paragraph>
<paragraph id="F07136CD524AFFEC5FD0EE464D6BFB80" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">
Wings. Fore wing
<quantity id="37369B28524AFFEC5E83EE464DEAFBC8" box="[404,489,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="431" unit="mm" value="3.2">3.2 mm</quantity>
long. Vein Rs+M not connected to Cu; radial cell 3.9 times as long as wide; R (costal vein of radial cell) 1.5 times as long as wide of stigma; 1cu-a interstitial to M; hind wing with two hamuli on anterosubmedial margin.
</paragraph>
<paragraph id="F07136CD524AFFEC5FD0EEDA4EE0FAF0" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">Metasoma.Syntergite1.9 times as long as maximum width, some longitudinal ridges reaching basal0.4 or shorter, not touching first thyridium; antero-dorsal margin of syntergite almost straight; posterior margin of synsternite with some wrinkles; tergite 6 punctate, antero-dorsal margin smooth, dorso-lateral margin finely punctate; ovipositor reaching around posterior margin of mesodiscrimen.</paragraph>
<paragraph id="F07136CD524AFFEC5FD0EF4A4D86FACC" blockId="4.[151,1437,150,1761]" box="[199,389,1303,1329]" pageId="4" pageNumber="431">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5FD0EF664F1AFA84" pageId="4" pageNumber="431" type="materials_examined">
<paragraph id="F07136CD524AFFEC5FD0EF664F1AFA84" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">
<materialsCitation id="40A63C90524AFFEC5FD0EF664F1AFA84" collectingDate="2019-03-31" collectionCode="ELKU" collectorName="K. Uemori" country="Taiwan" county="Country" elevation="463" latitude="22.130945" location="Mt. Gaoshifo-shan" longLatPrecision="1" longitude="120.81023" municipality="Mudan Township" pageId="4" pageNumber="431" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="C2BAEADF524AFFEC5FD0EF664D38FAA8" bold="true" box="[199,315,1339,1365]" pageId="4" pageNumber="431">
<typeStatus id="2F75886F524AFFEC5FD0EF664D35FAA8" box="[199,310,1339,1365]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female.
<collectingCountry id="88D9765D524AFFEC5EA0EF664E0EFAA8" box="[439,525,1339,1365]" name="Taiwan" pageId="4" pageNumber="431">Taiwan</collectingCountry>
:
<collectingRegion id="320AF82F524AFFEC5D35EF664E8EFAA8" box="[546,653,1339,1365]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="431">Pingdong</collectingRegion>
<collectingCounty id="19104E41524AFFEC5D88EF664EFFFAA8" box="[671,764,1339,1365]" pageId="4" pageNumber="431">Country</collectingCounty>
,
<collectingMunicipality id="1015ACB7524AFFEC5C06EF664FE1FAA8" box="[785,994,1339,1365]" pageId="4" pageNumber="431">Mudan Township</collectingMunicipality>
,
<location id="F5116016524AFFEC5CEFEF6648D4FAA8" LSID="urn:lsid:plazi:treatment:786787DB524DFFEC5F80EBD74EF2F91C:F5116016524AFFEC5CEFEF6648D4FAA8" box="[1016,1239,1339,1365]" country="Taiwan" county="Country" latitude="22.130945" longLatPrecision="1" longitude="120.81023" municipality="Mudan Township" name="Mt. Gaoshifo-shan" pageId="4" pageNumber="431" stateProvince="Pingtung">Mt. Gaoshifo-shan</location>
,
<geoCoordinate id="95FA500A524AFFEC5BFAEF67499FFAA8" box="[1261,1436,1338,1365]" degrees="22" direction="north" minutes="07" orientation="latitude" pageId="4" pageNumber="431" precision="1" seconds="51.4" value="22.130945">22°07’51.4” N</geoCoordinate>
<geoCoordinate id="95FA500A524AFFEC5F80EF024D4AFA84" box="[151,329,1375,1401]" degrees="120" direction="east" minutes="48" orientation="longitude" pageId="4" pageNumber="431" precision="1" seconds="36.8" value="120.81023">120°48’36.8” E</geoCoordinate>
,
<quantity id="37369B28524AFFEC5E42EF024D9DFA84" box="[341,414,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.63" pageId="4" pageNumber="431" unit="m" value="463.0">
<elevation id="7BE3D1FE524AFFEC5E42EF024D9DFA84" box="[341,414,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.63" pageId="4" pageNumber="431" unit="m" value="463.0">463 m</elevation>
</quantity>
,
<date id="8470100D524AFFEC5EBDEF024E28FA84" box="[426,555,1375,1401]" pageId="4" pageNumber="431" value="2019-03-31">
<collectingDate id="9434E9E5524AFFEC5EBDEF024E28FA84" box="[426,555,1375,1401]" pageId="4" pageNumber="431" value="2019-03-31">31 III 2019</collectingDate>
</date>
,
<collectorName id="5D3B531B524AFFEC5D21EF024EADFA84" box="[566,686,1375,1401]" pageId="4" pageNumber="431">K. Uemori</collectorName>
(
<collectionCode id="96DFAE08524AFFEC5DABEF024F0AFA84" box="[700,777,1375,1401]" pageId="4" pageNumber="431">ELKU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5FD0EFDE4D80FA60" box="[199,387,1411,1437]" pageId="4" pageNumber="431" type="biology_ecology">
<paragraph id="F07136CD524AFFEC5FD0EFDE4D80FA60" blockId="4.[151,1437,150,1761]" box="[199,387,1411,1437]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5FD0EFDE4D05FA60" bold="true" box="[199,262,1411,1437]" pageId="4" pageNumber="431">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5FD0EFFB4DC2FA3C" box="[199,449,1446,1473]" pageId="4" pageNumber="431" type="distribution">
<paragraph id="F07136CD524AFFEC5FD0EFFB4DC2FA3C" blockId="4.[151,1437,150,1761]" box="[199,449,1446,1473]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5FD0EFFB4D62FA3D" bold="true" box="[199,353,1446,1472]" pageId="4" pageNumber="431">Distribution.</emphasis>
<collectingCountry id="88D9765D524AFFEC5E70EFFA4DBEFA3C" box="[359,445,1447,1473]" name="Taiwan" pageId="4" pageNumber="431">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5FD0EF964D51F9F4" pageId="4" pageNumber="431" type="etymology">
<paragraph id="F07136CD524AFFEC5FD0EF964D51F9F4" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5FD0EF964D4DFA18" bold="true" box="[199,334,1483,1509]" pageId="4" pageNumber="431">Etymology.</emphasis>
The specific epithet is derived from the name of
<typeStatus id="2F75886F524AFFEC5C9FEF964FBBFA18" box="[904,952,1483,1509]" pageId="4" pageNumber="431">type</typeStatus>
locality, “formosa” which is the historical name of
<collectingCountry id="88D9765D524AFFEC5FEEEFB24D4CF9F4" box="[249,335,1519,1545]" name="Taiwan" pageId="4" pageNumber="431">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8D46546524AFFEC5FD0EC4E4EF2F91C" pageId="4" pageNumber="431" type="discussion">
<paragraph id="F07136CD524AFFEC5FD0EC4E4EF2F91C" blockId="4.[151,1437,150,1761]" pageId="4" pageNumber="431">
<emphasis id="C2BAEADF524AFFEC5FD0EC4E4D38F9D0" bold="true" box="[199,315,1555,1581]" pageId="4" pageNumber="431">Remarks.</emphasis>
This new species is closely related to
<taxonomicName id="37CE4D4E524AFFEC5C1DEC4E4FB8F9D0" baseAuthorityName="He &amp; Chu" baseAuthorityYear="1990" box="[778,955,1555,1581]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="quizhouensis">
<emphasis id="C2BAEADF524AFFEC5C1DEC4E4FB8F9D0" box="[778,955,1555,1581]" italics="true" pageId="4" pageNumber="431">V. quizhouensis</emphasis>
</taxonomicName>
in having the following character states: frontal transverse carina present; apical margin of clypeus convex; mesodiscrimen with about six foveae; ridges on syntergite not touching first thyridium. However, it can be distinguished by following characters: punctures between median ocellus and frontal transverse carina larger than ones on anterior half of head (same size in
<taxonomicName id="37CE4D4E524AFFEC5A9FEC224D2BF940" baseAuthorityName="He &amp; Chu" baseAuthorityYear="1990" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="quizhouensis">
<emphasis id="C2BAEADF524AFFEC5A9FEC224D2BF940" italics="true" pageId="4" pageNumber="431">V. quizhouensis</emphasis>
</taxonomicName>
); scutoscutellar sulcus with six foveae (five in
<taxonomicName id="37CE4D4E524AFFEC5C5CECFE4FF9F940" baseAuthorityName="He &amp; Chu" baseAuthorityYear="1990" box="[843,1018,1699,1725]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="quizhouensis">
<emphasis id="C2BAEADF524AFFEC5C5CECFE4FF9F940" box="[843,1018,1699,1725]" italics="true" pageId="4" pageNumber="431">V. quizhouensis</emphasis>
</taxonomicName>
); transverse carina on dorsal part of propodeum incomplete (complete in
<taxonomicName id="37CE4D4E524AFFEC5D22EC9A4EE2F91C" baseAuthorityName="He &amp; Chu" baseAuthorityYear="1990" box="[565,737,1735,1761]" class="Insecta" family="Vanhorniidae" genus="Vanhornia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="quizhouensis">
<emphasis id="C2BAEADF524AFFEC5D22EC9A4EE2F91C" box="[565,737,1735,1761]" italics="true" pageId="4" pageNumber="431">V. quizhouensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7A82F987C0F475DC9041737F0BFD8210">How high can trumpet moths occur: documentation of mountainous leaf-mining Tischeriidae, featuring a species from record-high elevations</mods:title>
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<taxonomicName id="328F3A81AA18057037DAFE19FD88FDB0" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[151,622,574,601]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="andina" status="sp. nov.">
<emphasis id="C7FB9D10AA18057037DAFE19FE7BFDB1" bold="true" box="[151,413,574,601]" italics="true" pageId="6" pageNumber="207">Astrotischeria andina</emphasis>
Diškus &amp; Stonis
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http://zoobank.org/NomenclaturalActs/
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(
<figureCitation id="6DB45D87AA18057037D2FEA0FEF4FD49" box="[159,274,647,673]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1768,1792]" captionTargetBox="[212,1375,181,1744]" captionTargetId="figure-18@7.[212,1375,181,1744]" captionTargetPageId="7" captionText="FIGURES 4–10. Adults of mountainous Astrotischeria species. 4, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, male holotype; 5, 6, same, female paratype; 7, A. andina Diškus &amp; Stonis, sp. nov., male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="6" pageNumber="207">Figs 7–10</figureCitation>
,
<figureCitation id="6DB45D87AA1805703653FEA0FE80FD49" box="[286,358,647,673]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">34–42</figureCitation>
,
<figureCitation id="6DB45D87AA180570363FFEA1FE5DFD49" box="[370,443,646,673]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="6" pageNumber="207">45, 46</figureCitation>
,
<figureCitation id="6DB45D87AA180570368BFEA1FDEBFD49" box="[454,525,646,673]" captionStart="FIGURES 53–59" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[160,1421,181,1885]" captionTargetId="figure-18@15.[154,1434,181,1885]" captionTargetPageId="15" captionText="FIGURES 53–59. Bionomics of Astrotischeria andina Diškus &amp; Stonis, sp. nov. 53, 54, habitat, Saqsaywaman, Cusco, Peru, 3,595–3,600 m; 55–57, host plant Baccharis buxifolia (Lam.) Pers., Asteraceae; 58, 59, leaf mines." figureDoi="http://doi.org/10.5281/zenodo.13757559" httpUri="https://zenodo.org/record/13757559/files/figure.png" pageId="6" pageNumber="207">53–59</figureCitation>
)
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<emphasis id="C7FB9D10AA18057037DAFEECFEA2FD0C" bold="true" box="[151,324,714,741]" pageId="6" pageNumber="207">Type material.</emphasis>
<materialsCitation id="45E74B5FAA1805703607FEECFD9AFCC5" collectingDate="2008-10-24" collectingDateMax="2008-11" collectingDateMin="2008-10-24" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="3595" latitude="-13.508611" location="Cusco Province" longLatPrecision="21" longitude="-71.979164" pageId="6" pageNumber="207" specimenCount="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="holotype">
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:
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,
<collectingCountry id="8D980192AA18057036BAFEECFDA7FD0D" box="[503,577,715,741]" name="Peru" pageId="6" pageNumber="207">PERU</collectingCountry>
,
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,
<collectingRegion id="374B8FE0AA1805703445FEECFCA8FD0D" box="[776,846,715,741]" country="Peru" name="Cusco" pageId="6" pageNumber="207">Cusco</collectingRegion>
(Saqsaywaman),
<geoCoordinate id="90BB27C5AA1805703359FEECFB70FD0D" box="[1044,1174,715,741]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="6" pageNumber="207" precision="15" seconds="31" value="-13.508611">13°30'31&quot;S</geoCoordinate>
,
<geoCoordinate id="90BB27C5AA18057033ECFEEDFACDFD0D" box="[1185,1323,714,741]" degrees="71" direction="west" minutes="58" orientation="longitude" pageId="6" pageNumber="207" precision="15" seconds="45" value="-71.979164">71°58'45&quot;W</geoCoordinate>
, elevation ca.
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<elevation id="7EA2A631AA180570378DFEC9FEC6FCE1" box="[192,288,750,777]" metricMagnitude="3" metricUnit="m" metricValue="3.595" pageId="6" pageNumber="207" unit="m" value="3595.0">3,595 m</elevation>
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, from feeding larva
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on
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<emphasis id="C7FB9D10AA180570357CFEC8FCE9FCE1" box="[561,783,751,777]" italics="true" pageId="6" pageNumber="207">Baccharis buxifolia</emphasis>
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<collectingDate id="91759E2AAA1805703452FEC8FC77FCE1" box="[799,913,751,777]" pageId="6" pageNumber="207" value="2008-10-24">24.x.2008</collectingDate>
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, ex pupa
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<collectingDate id="91759E2AAA180570334FFEC8FBBFFCE1" box="[1026,1113,751,777]" pageId="6" pageNumber="207" value="2008-11">xi.2008</collectingDate>
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, field card no. 4952, leg.
<collectorName id="587A24D4AA18057032CCFEC8FF0EFCC5" pageId="6" pageNumber="207">A. Diškus</collectorName>
, genitalia slide no. AD1160 (
<collectionCode id="939ED9C7AA180570357AFF34FD92FCC5" box="[567,628,787,813]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="6" pageNumber="207" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2A34FFA0AA18057035C4FF34FD10FCC5" box="[649,758,787,813]" pageId="6" pageNumber="207" type="paratype">Paratypes</typeStatus>
<specimenCount id="E3898A8BAA18057035B2FF34FCCBFCC5" box="[767,813,787,813]" count="2" pageId="6" pageNumber="207" type="male">2 ♂</specimenCount>
,
<specimenCount id="E3898A8BAA1805703476FF34FC81FCC5" box="[827,871,787,813]" count="1" pageId="6" pageNumber="207" type="female">1 ♀</specimenCount>
,
<materialsCitation id="45E74B5FAA180570343AFF34FF24FCB9" collectingDate="2008-10-24" collectingDateMax="2008-11" collectingDateMin="2008-10-24" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="3595" latitude="-13.508611" location="Cusco Province" longLatPrecision="21" longitude="-71.979164" pageId="6" pageNumber="207" specimenCount="1" stateProvince="Cusco" typeStatus="paratype">same label data as holotype, genitalia slides nos AD</materialsCitation>
<specimenCount id="E3898A8BAA180570378FFF10FEE7FCB9" box="[194,257,823,849]" count="748" pageId="6" pageNumber="207" type="male">748♂</specimenCount>
, AD1179
<materialsCitation id="45E74B5FAA180570363FFF10FE35FCB9" box="[370,467,823,849]" collectionCode="MfN" country="Peru" pageId="6" pageNumber="207" specimenCount="1" specimenCount-female="1" stateProvince="Cusco" typeStatus="paratype">
<specimenCount id="E3898A8BAA180570363FFF10FE66FCB9" box="[370,384,823,849]" count="1" pageId="6" pageNumber="207" type="female">♀</specimenCount>
(
<collectionCode id="939ED9C7AA18057036C3FF10FE2DFCB9" box="[398,459,823,849]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="6" pageNumber="207" type="Museum">MfN</collectionCode>
)
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.
</paragraph>
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<emphasis id="C7FB9D10AA180570378AFF7DFEA4FC9C" bold="true" box="[199,322,858,884]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
Externally, this species is characterized by striking sexual dimorphism; the males are distinguished by a densely irrorated forewing, while the females exhibit an entirely yellowish cream forewing. In the male genitalia, the combination of an uncus with slender lateral lobes and pronounced rounded median lobes, sinuous, dentate dorsal valval lobes, and a phallus that is widely bilobed distinguishes the new species from all other species of the genus.
</paragraph>
<paragraph id="F5304102AA180570378AF828FC4FFACD" blockId="6.[151,1437,714,2037]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AF828FEE2FBC1" bold="true" box="[199,260,1039,1065]" pageId="6" pageNumber="207">Male</emphasis>
(
<figureCitation id="6DB45D87AA1805703658F828FE98FBC1" box="[277,382,1039,1065]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1768,1792]" captionTargetBox="[212,1375,181,1744]" captionTargetId="figure-18@7.[212,1375,181,1744]" captionTargetPageId="7" captionText="FIGURES 4–10. Adults of mountainous Astrotischeria species. 4, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, male holotype; 5, 6, same, female paratype; 7, A. andina Diškus &amp; Stonis, sp. nov., male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="6" pageNumber="207">Figs 7, 8</figureCitation>
). Forewing length 3.5–4.0 mm; wingspan
<quantity id="3277ECE7AA1805703425F828FC14FBC1" box="[872,1010,1039,1065]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="8.7" metricValueMin="7.7" pageId="6" pageNumber="207" unit="mm" value="8.2" valueMax="8.7" valueMin="7.7">7.7–8.7 mm</quantity>
(n = 2). Head: frons and palpi ochre cream; frontal tuft ochreous brown to yellowish ochre laterally, dark ochreous brown distally; collar pale ochreous brown to ochre; antenna slightly longer than half the length of forewing; flagellum glossy dark brown on the upper side, blackish brown on the underside. Thorax pale ochreous brown to ochre. Tegula grey-brown. Forewing pale brownish grey densely irrorated with dark brown ochre-tipped scales; fringe pale brownish grey to grey; fringe line absent; forewing underside grey-brown, without spots or androconia. Hindwing and its fringe pale brownish to grey, without androconia. Legs glossy grey; forelegs and midlegs densely covered with dark brown or dark grey scales on the upper side. Abdomen glossy grey on the upper side and underside.
</paragraph>
<paragraph id="F5304102AA180570378AF908FAA1FA15" blockId="6.[151,1437,714,2037]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AF908FE94FAA0" bold="true" box="[199,370,1326,1353]" pageId="6" pageNumber="207">Male genitalia</emphasis>
(
<figureCitation id="6DB45D87AA18057036CCF908FDE3FAA1" box="[385,517,1327,1353]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Figs 34–42</figureCitation>
). Capsule 540–595 µm long, 280 µm wide. Uncus with long, slender lateral lobes (
<figureCitation id="6DB45D87AA18057037D2F974FEAFFA85" box="[159,329,1363,1389]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Figs 36, 37, 41</figureCitation>
) and short but rounded median lobes; the gap between the median rounded lobes is distinctly wide and thickened (
<figureCitation id="6DB45D87AA1805703609F950FE7FFA79" box="[324,409,1399,1425]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Fig. 38</figureCitation>
). Socii relatively small, membranous, covered with tiny spines. Valva divided: ventral lobe (main body) very slender and slightly sinuous (
<figureCitation id="6DB45D87AA18057035F6F9BCFCF4FA5D" box="[699,786,1435,1461]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Fig. 41</figureCitation>
); dorsal lobe slightly sinuous, inwardly curved, strongly chitinized, distinctly pointed distally, dentate on the inner side (
<figureCitation id="6DB45D87AA1805703403F998FC47FA31" box="[846,929,1471,1497]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Fig. 40</figureCitation>
). Transtilla absent.Anellus weakly chitinized, indistinctive. Phallus (
<figureCitation id="6DB45D87AA18057036DCF9C4FDF7FA15" box="[401,529,1506,1533]" captionStart="FIGURES 34–42" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-21@12.[159,1428,181,1881]" captionTargetPageId="12" captionText="FIGURES 34–42. Male genitalia of Astrotischeria andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m. 34, phallus, paratype, genitalia slide no. AD748 (MfN). 35, same, holotype, genitalia slide no. AD1160 (MfN); 36–39, details of genitalia capsule, holotype, genitalia slide no.AD1160 (MfN); 40, dentate dorsal lobe of valva, paratype, genitalia slide no.AD748 (MfN); 41, 42, general view of genitalia capsule with phallus removed, holotype, genitalia slide no. AD1160 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757551" httpUri="https://zenodo.org/record/13757551/files/figure.png" pageId="6" pageNumber="207">Figs 34, 35</figureCitation>
) 410–500 µm long, slender, divided apically, with two wide apical lobes.
</paragraph>
<paragraph id="F5304102AA180570378AFA20FAB9F965" blockId="6.[151,1437,714,2037]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFA20FEF8F9C9" bold="true" box="[199,286,1543,1569]" pageId="6" pageNumber="207">Female</emphasis>
(
<figureCitation id="6DB45D87AA1805703666FA20FE7AF9C9" box="[299,412,1543,1569]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1768,1792]" captionTargetBox="[212,1375,181,1744]" captionTargetId="figure-18@7.[212,1375,181,1744]" captionTargetPageId="7" captionText="FIGURES 4–10. Adults of mountainous Astrotischeria species. 4, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, male holotype; 5, 6, same, female paratype; 7, A. andina Diškus &amp; Stonis, sp. nov., male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="6" pageNumber="207">Figs 9, 10</figureCitation>
). Forewing length
<quantity id="3277ECE7AA1805703527FA20FD24F9C9" box="[618,706,1543,1569]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="207" unit="mm" value="3.2">3.2 mm</quantity>
; wingspan
<quantity id="3277ECE7AA1805703471FA20FC76F9C9" box="[828,912,1543,1569]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="6" pageNumber="207" unit="mm" value="7.1">7.1 mm</quantity>
(n = 1). Head: frons, palpi, and collar yellowish cream; frontal tuft yellowish cream, but grey-brown laterally; antenna slightly longer than half the length of forewing; flagellum glossy grey. Thorax, tegula, and forewing yellowish cream; forewing underside pale brownishgrey, without spots or androconia. Hindwing and its fringe cream, without androconia. Legs brownish cream.
</paragraph>
<paragraph id="F5304102AA180570378AFAB0FB95F861" blockId="6.[151,1437,714,2037]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFAB0FE6BF958" bold="true" box="[199,397,1686,1713]" pageId="6" pageNumber="207">Female genitalia</emphasis>
(
<figureCitation id="6DB45D87AA18057036D0FAB0FDF9F959" box="[413,543,1686,1713]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="6" pageNumber="207">Figs 45, 46</figureCitation>
) 1650 µm long. Ovipositor lobes large, distinctly flattened, clothed with short, modified setae (‘peg setae’); area between ovipositor lobes medium wide, with tiny indistinctive papillae and some short setae. Second pair of lobes, lateral and anterior to the ovipositor lobes, three to four times smaller than ovipositor lobes, but bearing setae. Anterior and posterior apophyses almost equal in length; prela comprised of three pairs of unique projections (
<figureCitation id="6DB45D87AA180570355BFB00FD8DF8A9" box="[534,619,1831,1857]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="6" pageNumber="207">Fig. 46</figureCitation>
). Corpus bursae with a very long and slender &quot;neck&quot; (
<figureCitation id="6DB45D87AA180570339DFB00FAC3F8A9" box="[1232,1317,1830,1857]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="6" pageNumber="207">Fig. 45</figureCitation>
); the main part small and oval-shaped, without distinctive pectinations or signum. Accessory sac absent. Ductus spermathecae short and very slender, with about 3 large coils and weakly chitinized, plate-like vesicle.
</paragraph>
<paragraph id="F5304102AA180570378AFBB5FA86F81D" blockId="6.[151,1437,714,2037]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFBB5FEA5F844" bold="true" box="[199,323,1938,1964]" pageId="6" pageNumber="207">Bionomics</emphasis>
(
<figureCitation id="6DB45D87AA180570361EFBB4FE31F845" box="[339,471,1938,1965]" captionStart="FIGURES 53–59" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[160,1421,181,1885]" captionTargetId="figure-18@15.[154,1434,181,1885]" captionTargetPageId="15" captionText="FIGURES 53–59. Bionomics of Astrotischeria andina Diškus &amp; Stonis, sp. nov. 53, 54, habitat, Saqsaywaman, Cusco, Peru, 3,595–3,600 m; 55–57, host plant Baccharis buxifolia (Lam.) Pers., Asteraceae; 58, 59, leaf mines." figureDoi="http://doi.org/10.5281/zenodo.13757559" httpUri="https://zenodo.org/record/13757559/files/figure.png" pageId="6" pageNumber="207">Figs 53–59</figureCitation>
). The host plant is
<taxonomicName id="328F3A81AA18057035E1FBB4FBFAF845" authority="(Lam.) Pers." authorityName="Pers." baseAuthorityName="Lam." box="[684,1052,1939,1965]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="species" species="buxifolia">
<emphasis id="C7FB9D10AA18057035E1FBB4FC6FF845" box="[684,905,1939,1965]" italics="true" pageId="6" pageNumber="207">Baccharis buxifolia</emphasis>
(Lam.) Pers.
</taxonomicName>
(=
<taxonomicName id="328F3A81AA1805703371FBB4FA77F845" authority="Cuatrec." authorityName="Cuatrec." box="[1084,1425,1939,1965]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="C7FB9D10AA1805703371FBB4FAC1F844" box="[1084,1319,1939,1965]" italics="true" pageId="6" pageNumber="207">Baccharis peruviana</emphasis>
Cuatrec.
</taxonomicName>
),
<taxonomicName id="328F3A81AA18057037DAFB90FEF4F839" box="[151,274,1975,2001]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
(identified by Liliana Katinas) (
<figureCitation id="6DB45D87AA1805703538FB90FD10F839" box="[629,758,1974,2001]" captionStart="FIGURES 53–59" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[160,1421,181,1885]" captionTargetId="figure-18@15.[154,1434,181,1885]" captionTargetPageId="15" captionText="FIGURES 53–59. Bionomics of Astrotischeria andina Diškus &amp; Stonis, sp. nov. 53, 54, habitat, Saqsaywaman, Cusco, Peru, 3,595–3,600 m; 55–57, host plant Baccharis buxifolia (Lam.) Pers., Asteraceae; 58, 59, leaf mines." figureDoi="http://doi.org/10.5281/zenodo.13757559" httpUri="https://zenodo.org/record/13757559/files/figure.png" pageId="6" pageNumber="207">Figs 55–57</figureCitation>
). Larvae mine leaves in October. The leaf mine is blotch-like, pale brownish-yellow to brown (
<figureCitation id="6DB45D87AA1805703548FBFCFD61F81D" box="[517,647,2010,2037]" captionStart="FIGURES 53–59" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[160,1421,181,1885]" captionTargetId="figure-18@15.[154,1434,181,1885]" captionTargetPageId="15" captionText="FIGURES 53–59. Bionomics of Astrotischeria andina Diškus &amp; Stonis, sp. nov. 53, 54, habitat, Saqsaywaman, Cusco, Peru, 3,595–3,600 m; 55–57, host plant Baccharis buxifolia (Lam.) Pers., Asteraceae; 58, 59, leaf mines." figureDoi="http://doi.org/10.5281/zenodo.13757559" httpUri="https://zenodo.org/record/13757559/files/figure.png" pageId="6" pageNumber="207">Figs 58, 59</figureCitation>
). Adults occur in November. Otherwise, the biology is unknown.
</paragraph>
</subSubSection>
<caption id="A1F0118AAA19057137DAFACFFD88F8A0" ID-DOI="http://doi.org/10.5281/zenodo.13757533" ID-Zenodo-Dep="13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="7" pageNumber="208" startId="7.[151,264,1768,1792]" targetBox="[212,1375,181,1744]" targetPageId="7" targetType="figure">
<paragraph id="F5304102AA19057137DAFACFFD88F8A0" blockId="7.[151,1436,1768,1864]" pageId="7" pageNumber="208">
<emphasis id="C7FB9D10AA19057137DAFACFFEADF8E8" bold="true" box="[151,331,1768,1792]" pageId="7" pageNumber="208">FIGURES 4–10.</emphasis>
Adults of mountainous
<taxonomicName id="328F3A81AA1905713503FACFFCA9F8E8" box="[590,847,1768,1792]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="undefined-4">
<emphasis id="C7FB9D10AA1905713503FACFFD3BF8E8" box="[590,733,1768,1792]" italics="true" pageId="7" pageNumber="208">Astrotischeria</emphasis>
species. 4
</taxonomicName>
,
<taxonomicName id="328F3A81AA1905713410FACFFB76F8E8" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[861,1168,1768,1792]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="viscacha" status="sp. nov.">
<emphasis id="C7FB9D10AA1905713410FACFFC30F8E8" box="[861,982,1768,1792]" italics="true" pageId="7" pageNumber="208">A. viscacha</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="C7FB9D10AA19057133D3FACFFB15F8E8" bold="true" box="[1182,1267,1768,1792]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="DCC8206BAA19057133D3FACFFB15F8E8" box="[1182,1267,1768,1792]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 2,990 m, male holotype; 5, 6, same, female paratype; 7,
<taxonomicName id="328F3A81AA190571350EFB2BFCBCF8CC" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[579,858,1804,1828]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="andina" status="sp. nov.">
<emphasis id="C7FB9D10AA190571350EFB2BFD41F8CC" box="[579,679,1804,1828]" italics="true" pageId="7" pageNumber="208">A. andina</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="C7FB9D10AA190571342BFB2BFC51F8CC" bold="true" box="[870,951,1804,1828]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="DCC8206BAA190571342BFB2BFC51F8CC" box="[870,951,1804,1828]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN).
</paragraph>
</caption>
<subSubSection id="BD951289AA190571378AFBB5FDE0F839" pageId="7" pageNumber="208" type="distribution">
<paragraph id="F5304102AA190571378AFBB5FDE0F839" blockId="7.[151,1437,1938,2037]" pageId="7" pageNumber="208">
<emphasis id="C7FB9D10AA190571378AFBB5FE87F844" bold="true" box="[199,353,1938,1964]" pageId="7" pageNumber="208">Distribution.</emphasis>
This species is currently known from a single locality in
<collectingCountry id="8D980192AA190571349DFBB4FBE5F845" box="[976,1027,1939,1965]" name="Peru" pageId="7" pageNumber="208">Peru</collectingCountry>
(Saqsaywaman,
<collectingRegion id="374B8FE0AA19057133F2FBB5FAEEF844" box="[1215,1288,1938,1964]" country="Peru" name="Cusco" pageId="7" pageNumber="208">Cusco</collectingRegion>
, at elevations of
<quantity id="3277ECE7AA19057137FBFB91FE86F838" box="[182,352,1974,2001]" metricMagnitude="3" metricUnit="m" metricValue="3.5975" metricValueMax="3.6" metricValueMin="3.595" pageId="7" pageNumber="208" unit="m" value="3597.5" valueMax="3600.0" valueMin="3595.0">
<geoCoordinate id="90BB27C5AA19057137FBFB91FF12F839" box="[182,244,1974,2001]" degrees="3,595" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="3.595">3,595</geoCoordinate>
<geoCoordinate id="90BB27C5AA19057137B9FB91FEA7F839" box="[244,321,1974,2001]" degrees="3,600" direction="west" orientation="longitude" pageId="7" pageNumber="208" precision="55" value="-3.6">–3,600</geoCoordinate>
m
</quantity>
) (
<figureCitation id="6DB45D87AA190571363BFB91FE1FF839" box="[374,505,1974,2001]" captionStart="FIGURES 53–59" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[160,1421,181,1885]" captionTargetId="figure-18@15.[154,1434,181,1885]" captionTargetPageId="15" captionText="FIGURES 53–59. Bionomics of Astrotischeria andina Diškus &amp; Stonis, sp. nov. 53, 54, habitat, Saqsaywaman, Cusco, Peru, 3,595–3,600 m; 55–57, host plant Baccharis buxifolia (Lam.) Pers., Asteraceae; 58, 59, leaf mines." figureDoi="http://doi.org/10.5281/zenodo.13757559" httpUri="https://zenodo.org/record/13757559/files/figure.png" pageId="7" pageNumber="208">Figs 53, 54</figureCitation>
).
</paragraph>
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<subSubSection id="BD951289AA190571378AFBFDFB69F81D" box="[199,1167,2010,2037]" pageId="7" pageNumber="208" type="etymology">
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<emphasis id="C7FB9D10AA190571378AFBFDFEA8F81C" bold="true" box="[199,334,2010,2036]" pageId="7" pageNumber="208">Etymology.</emphasis>
The species is named after the Andes, the mountain ridge where it occurs.
</paragraph>
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243
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<emphasis id="C7FB9D10AA1A057237DAFD6DFE24FE8D" bold="true" box="[151,450,330,357]" italics="true" pageId="4" pageNumber="205">Astrotischeria peruanica</emphasis>
Diškus &amp; Stonis
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http://zoobank.org/NomenclaturalActs/
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(
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,
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,
<figureCitation id="6DB45D87AA1A05723605FDB5FE69FE45" box="[328,399,402,429]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="4" pageNumber="205">48–50</figureCitation>
)
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<emphasis id="C7FB9D10AA1A057237DAFDFCFEA3FE1C" bold="true" box="[151,325,474,501]" pageId="4" pageNumber="205">Type material.</emphasis>
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,
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, near
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, elevation ca.
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on
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<collectingDate id="91759E2AAA1A057235F1FDD8FCC9FDF1" box="[700,815,511,537]" pageId="4" pageNumber="205" value="2008-10-19">19.x.2008</collectingDate>
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, ex pupa
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<collectingDate id="91759E2AAA1A057234ECFDD8FC1EFDF1" box="[929,1016,511,537]" pageId="4" pageNumber="205" value="2008-11">xi.2008</collectingDate>
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, field card no. 4942, leg.
<collectorName id="587A24D4AA1A0572326EFDD8FA7EFDF1" box="[1315,1432,511,537]" pageId="4" pageNumber="205">A. Diškus</collectorName>
, genitalia slide no. AD1180 (
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.
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,
<materialsCitation id="45E74B5FAA1A0572358CFE04FAD7FDD5" box="[705,1329,547,573]" collectingDate="2008-10-19" collectingDateMax="2008-11" collectingDateMin="2008-10-19" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="2110" latitude="-13.162222" location="Urubamba Province" longLatPrecision="21" longitude="-72.540276" municipality="Machu Picchu" pageId="4" pageNumber="205" specimenCount="1" typeStatus="paratype">same label data as holotype, genitalia slide no. AD1172</materialsCitation>
<materialsCitation id="45E74B5FAA1A0572327CFE04FA71FDD5" box="[1329,1431,547,573]" collectionCode="MfN" country="Peru" pageId="4" pageNumber="205" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="E3898A8BAA1A0572327CFE04FAA2FDD5" box="[1329,1348,547,573]" count="1" pageId="4" pageNumber="205" type="male">♂</specimenCount>
(
<collectionCode id="939ED9C7AA1A0572321FFE04FA69FDD5" box="[1362,1423,547,573]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="4" pageNumber="205" type="Museum">MfN</collectionCode>
)
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.
</paragraph>
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<emphasis id="C7FB9D10AA1A0572378AFE61FEA4FD88" bold="true" box="[199,322,582,608]" pageId="4" pageNumber="205">Diagnosis.</emphasis>
Externally, this species resembles other
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<emphasis id="C7FB9D10AA1A05723449FE60FC44FD89" box="[772,930,583,609]" italics="true" pageId="4" pageNumber="205">Astrotischeria</emphasis>
species
</taxonomicName>
with ochre-colored spots. In the male genitalia, it is distinguished by a slender, straight valva; a slender, deeply divided phallus distally; a basally greatly extended uncus; and slender, slightly curved dorsal lobes, characters that differentiate it from all other species within the genus.
</paragraph>
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<emphasis id="C7FB9D10AA1A0572378AFEF0FEE2FD19" bold="true" box="[199,260,727,753]" pageId="4" pageNumber="205">Male</emphasis>
(
<figureCitation id="6DB45D87AA1A0572365EFEF0FEBFFD19" box="[275,345,727,753]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1467,1491]" captionTargetBox="[177,1410,179,1441]" captionTargetId="figure-18@3.[177,1410,179,1441]" captionTargetPageId="3" captionText="FIGURES 1–3. Adults of mountainous Astrotischeria species. 1, A. parapallens Diškus &amp; Stonis, 3,320 m, non-type specimen, a new elevation record; 2, A. peruanica Diškus &amp; Stonis, sp. nov., holotype, 2,110 m; 3, A. montivaga Diškus &amp; Stonis, sp. nov., paratype, 3,320 m (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757529" httpUri="https://zenodo.org/record/13757529/files/figure.png" pageId="4" pageNumber="205">Fig. 2</figureCitation>
). Forewing length 3.0–
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; wingspan
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(n = 2). Head: frons ochre; palpi ochreous cream; pecten very slender, brownish, distally ochre; frontal tuft glossy grey to brown, ochreous distally; collar distinctive, ochre, pale brown distally; antenna slightly longer than half the length of forewing; flagellum glossy brown on upper side, ochre on underside. Thorax grey-brown medially, ochre laterally; tegula dark grey-brown. Forewing dark grey-brown, with large elongated ochre patches medially; fringe grey-brown; fringe line indistinct; forewing underside dark brown, without spots or androconia. Hindwing and its fringe grey-brown to dark grey-brown, without androconia. Legs grey-brown.
</paragraph>
<paragraph id="F5304102AA1A0572378AFFF4FCA6FB49" blockId="4.[151,1437,474,1437]" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A0572378AFFF4FE93FC04" bold="true" box="[199,373,978,1005]" pageId="4" pageNumber="205">Male genitalia</emphasis>
(
<figureCitation id="6DB45D87AA1A057236CBFFF4FDEAFC05" box="[390,524,979,1005]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Figs 16–23</figureCitation>
). Capsule 480–540 µm long, 230–240 µm wide. Uncus comprised of two long lobes strongly widened and thickened basally, partially divided into two lobes: long lateral and short rounded medial (
<figureCitation id="6DB45D87AA1A057237D2F83CFE93FBDD" box="[159,373,1051,1077]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Figs 18, 19, 21, 22</figureCitation>
). Socii large, slender, membranous, covered with tiny spines. Valva divided: ventral lobe (main body) very narrow and straight (
<figureCitation id="6DB45D87AA1A05723540F818FD85FBB1" box="[525,611,1087,1113]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Fig. 16</figureCitation>
); dorsal lobe inwardly slightly sinuous, slender (
<figureCitation id="6DB45D87AA1A057233D8F818FAFBFBB1" box="[1173,1309,1087,1113]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Figs 17, 20</figureCitation>
). Transtilla absent. Anellus slightly thickened laterally with about three chetae on each side (
<figureCitation id="6DB45D87AA1A05723353F844FB46FB95" box="[1054,1184,1123,1149]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Figs 17, 20</figureCitation>
). Phallus (
<figureCitation id="6DB45D87AA1A05723259F844FA72FB95" box="[1300,1428,1123,1149]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Figs 17, 23</figureCitation>
) 255–340 µm long, slender, apically deeply divided (
<figureCitation id="6DB45D87AA1A05723592F8A0FCD2FB49" box="[735,820,1159,1185]" captionStart="FIGURES 16–23" captionStartId="9.[151,264,1886,1910]" captionTargetBox="[172,1406,181,1862]" captionTargetId="figure-18@9.[161,1425,181,1863]" captionTargetPageId="9" captionText="FIGURES 16–23. Male genitalia of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov., 2,110 m. 16–19, holotype, genitalia slide no. AD1180 (MfN); 20–23, paratype, with phallus (23) removed, genitalia slide no. AD1172 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757539" httpUri="https://zenodo.org/record/13757539/files/figure.png" pageId="4" pageNumber="205">Fig. 23</figureCitation>
).
</paragraph>
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<emphasis id="C7FB9D10AA1A0572378AF88CFEF8FB2D" bold="true" box="[199,286,1195,1221]" pageId="4" pageNumber="205">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="F5304102AA1A0572378AF8E9FDCEFAD9" blockId="4.[151,1437,474,1437]" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A0572378AF8E9FEA5FB00" bold="true" box="[199,323,1230,1256]" pageId="4" pageNumber="205">Bionomics</emphasis>
(
<figureCitation id="6DB45D87AA1A05723619F8E8FE3CFB01" box="[340,474,1230,1257]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="4" pageNumber="205">Figs 48–50</figureCitation>
). Host plant is unidentified, possibly
<taxonomicName id="328F3A81AA1A057234CBF8E8FB44FB01" authority="Pers." authorityName="Pers." box="[902,1186,1231,1257]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="species" species="trinervis">
<emphasis id="C7FB9D10AA1A057234CBF8E8FBB9FB00" box="[902,1119,1231,1257]" italics="true" pageId="4" pageNumber="205">Baccharis trinervis</emphasis>
Pers.
</taxonomicName>
,
<taxonomicName id="328F3A81AA1A057233E3F8E8FACFFB01" box="[1198,1321,1231,1257]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
(
<figureCitation id="6DB45D87AA1A05723277F8E8FA76FB01" box="[1338,1424,1231,1257]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="4" pageNumber="205">Fig. 48</figureCitation>
). Larvae mine leaves in October, producing irregular or rounded blotch mines (
<figureCitation id="6DB45D87AA1A057234A8F8D4FB85FAE5" box="[997,1123,1266,1293]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="4" pageNumber="205">Figs 49, 50</figureCitation>
).Adults occur in November. Otherwise, the biology is unknown.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1A0572378AF91DFDFAFA91" pageId="4" pageNumber="205" type="distribution">
<paragraph id="F5304102AA1A0572378AF91DFDFAFA91" blockId="4.[151,1437,474,1437]" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A0572378AF91DFE87FABC" bold="true" box="[199,353,1338,1364]" pageId="4" pageNumber="205">Distribution.</emphasis>
This species is currently known from a single locality in
<collectingCountry id="8D980192AA1A057234ABF91CFBFAFABD" box="[998,1052,1339,1365]" name="Peru" pageId="4" pageNumber="205">Peru</collectingCountry>
, Urubamba Province, near Machu Picchu, at an elevation of
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.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1A0572378AF9A4FBA5FA75" box="[199,1091,1411,1437]" pageId="4" pageNumber="205" type="etymology">
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<emphasis id="C7FB9D10AA1A0572378AF9A4FEA8FA75" bold="true" box="[199,334,1411,1437]" pageId="4" pageNumber="205">Etymology.</emphasis>
The new species is named after
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, the country of its occurrence.
</paragraph>
</subSubSection>
</treatment>
</document>

240
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<mods:number id="5721A1F8E60584518C2BDD9051D59638">2</mods:number>
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<mods:start id="65B720062B8B680C031EDDFFA01FDC59">201</mods:start>
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<mods:location id="8A51F11EFC5D2BFD71D45019BC418D88">
<mods:url id="0C0E1878D6A07FF69D22C1ABAB0AEBB3">http://dx.doi.org/10.11646/zootaxa.5507.2.1</mods:url>
</mods:location>
<mods:classification id="2D531C6C859B877982BE0B790381B706">journal article</mods:classification>
<mods:identifier id="C97147F4EB7273F207E90D1D2B3F8456" type="DOI">10.11646/zootaxa.5507.2.1</mods:identifier>
<mods:identifier id="F06E571D9E43BDB2B372418A49FC88D3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DB4E8D612860BCE90CCA35C2D3B40606" type="Zenodo-Dep">13757527</mods:identifier>
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<treatment id="7D26F014AA1A057337DAF9C1FC17FCD1" ID-DOI="http://doi.org/10.5281/zenodo.13757543" ID-Zenodo-Dep="13757543" LSID="urn:lsid:plazi:treatment:7D26F014AA1A057337DAF9C1FC17FCD1" httpUri="http://treatment.plazi.org/id/7D26F014AA1A057337DAF9C1FC17FCD1" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<subSubSection id="BD951289AA1A057237DAF9C1FBB9F9CD" pageId="4" pageNumber="205" type="nomenclature">
<paragraph id="F5304102AA1A057237DAF9C1FCE5F9E9" blockId="4.[151,1119,1510,1609]" box="[151,771,1510,1537]" pageId="4" pageNumber="205">
<heading id="AE78F66EAA1A057237DAF9C1FCE5F9E9" bold="true" box="[151,771,1510,1537]" fontSize="11" level="1" pageId="4" pageNumber="205" reason="1">
<emphasis id="C7FB9D10AA1A057237DAF9C1FCE5F9E9" bold="true" box="[151,771,1510,1537]" pageId="4" pageNumber="205">
<taxonomicName id="328F3A81AA1A057237DAF9C1FD7EF9E8" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[151,664,1510,1537]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="montivaga" status="sp. nov.">
<emphasis id="C7FB9D10AA1A057237DAF9C1FE21F9E9" bold="true" box="[151,455,1510,1537]" italics="true" pageId="4" pageNumber="205">Astrotischeria montivaga</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicNameLabel id="DCC8206BAA1A057235EBF9C0FCE5F9E9" box="[678,771,1511,1537]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F5304102AA1A057237DAFA2DFBB9F9CD" blockId="4.[151,1119,1510,1609]" box="[151,1119,1546,1573]" pageId="4" pageNumber="205">
http://zoobank.org/NomenclaturalActs/
<uuid id="81297BD7AA1A05723511FA2DFBB9F9CD" box="[604,1119,1546,1573]" pageId="4" pageNumber="205">59EE4B18-6259-4D57-9AC3-29D2E5E4E540</uuid>
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1A057237DAFA08FEA5F9A1" box="[151,323,1583,1609]" pageId="4" pageNumber="205" type="description">
<paragraph id="F5304102AA1A057237DAFA08FEA5F9A1" blockId="4.[151,1119,1510,1609]" box="[151,323,1583,1609]" pageId="4" pageNumber="205">
(
<figureCitation id="6DB45D87AA1A057237D2FA08FF01F9A1" box="[159,231,1583,1609]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1467,1491]" captionTargetBox="[177,1410,179,1441]" captionTargetId="figure-18@3.[177,1410,179,1441]" captionTargetPageId="3" captionText="FIGURES 1–3. Adults of mountainous Astrotischeria species. 1, A. parapallens Diškus &amp; Stonis, 3,320 m, non-type specimen, a new elevation record; 2, A. peruanica Diškus &amp; Stonis, sp. nov., holotype, 2,110 m; 3, A. montivaga Diškus &amp; Stonis, sp. nov., paratype, 3,320 m (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757529" httpUri="https://zenodo.org/record/13757529/files/figure.png" pageId="4" pageNumber="205">Figs 3</figureCitation>
,
<figureCitation id="6DB45D87AA1A057237B9FA08FEDAF9A1" box="[244,316,1583,1609]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="4" pageNumber="205">24–28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1A057237DAFA50FEB0F915" pageId="4" pageNumber="205" type="materials_examined">
<paragraph id="F5304102AA1A057237DAFA50FEB0F915" blockId="4.[151,1437,1654,2041]" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A057237DAFA50FEA2F978" bold="true" box="[151,324,1654,1681]" pageId="4" pageNumber="205">Type material.</emphasis>
<materialsCitation id="45E74B5FAA1A05723607FA50FD2BF931" collectingDate="2008-10-28" collectingDateMax="2008-11" collectingDateMin="2008-10-28" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="3320" latitude="-13.047223" location="Quinua" longLatPrecision="21" longitude="-74.13555" municipality="Huamanga Province" pageId="4" pageNumber="205" specimenCount="1" specimenCount-male="1" stateProvince="Ayacucho" typeStatus="holotype">
<typeStatus id="2A34FFA0AA1A05723607FA50FE51F979" box="[330,439,1655,1681]" pageId="4" pageNumber="205" type="holotype">Holotype</typeStatus>
:
<specimenCount id="E3898A8BAA1A0572368DFA50FE0AF979" box="[448,492,1655,1681]" count="1" pageId="4" pageNumber="205" type="male">1 ♂</specimenCount>
,
<collectingCountry id="8D980192AA1A057236BAFA50FDA7F979" box="[503,577,1655,1681]" name="Peru" pageId="4" pageNumber="205">PERU</collectingCountry>
,
<collectingMunicipality id="1554DB78AA1A05723501FA50FCD7F979" box="[588,817,1655,1681]" pageId="4" pageNumber="205">Huamanga Province</collectingMunicipality>
,
<quantity id="3277ECE7AA1A05723471FA50FC64F979" box="[828,898,1655,1681]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="205" unit="km" value="37.0">37 km</quantity>
NE of
<collectingRegion id="374B8FE0AA1A05723482FA50FBA7F979" box="[975,1089,1655,1681]" country="Peru" name="Ayacucho" pageId="4" pageNumber="205">Ayacucho</collectingRegion>
,
<location id="F05017D9AA1A05723301FA50FB47F979" LSID="urn:lsid:plazi:treatment:7D26F014AA1A057337DAF9C1FC17FCD1:F05017D9AA1A05723301FA50FB47F979" box="[1100,1185,1655,1681]" country="Peru" latitude="-13.047223" longLatPrecision="21" longitude="-74.13555" municipality="Huamanga Province" name="Quinua" pageId="4" pageNumber="205" stateProvince="Ayacucho">Quinua</location>
,
<geoCoordinate id="90BB27C5AA1A057233E6FA51FAF9F979" box="[1195,1311,1654,1681]" degrees="13" direction="south" minutes="2" orientation="latitude" pageId="4" pageNumber="205" precision="15" seconds="50" value="-13.047223">13°2'50&quot;S</geoCoordinate>
,
<geoCoordinate id="90BB27C5AA1A05723267FA50FA71F979" box="[1322,1431,1655,1681]" degrees="74" direction="west" minutes="8" orientation="longitude" pageId="4" pageNumber="205" precision="15" seconds="8" value="-74.13555">74°8'8&quot;W</geoCoordinate>
, elevation ca.
<quantity id="3277ECE7AA1A05723662FABCFE69F95D" box="[303,399,1691,1717]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="4" pageNumber="205" unit="m" value="3320.0">
<elevation id="7EA2A631AA1A05723662FABCFE69F95D" box="[303,399,1691,1717]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="4" pageNumber="205" unit="m" value="3320.0">3,320 m</elevation>
</quantity>
, mining larvae possibly
<collectedFrom id="469302FFAA1A057235E5FABCFC8EF95D" box="[680,872,1691,1717]" pageId="4" pageNumber="205">
on
<taxonomicName id="328F3A81AA1A05723581FABCFC8EF95D" box="[716,872,1691,1717]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="C7FB9D10AA1A05723581FABCFCD8F95D" box="[716,830,1691,1717]" italics="true" pageId="4" pageNumber="205">Baccharis</emphasis>
sp.
</taxonomicName>
</collectedFrom>
,
<date id="813167C2AA1A0572343BFABCFC0EF95D" box="[886,1000,1691,1717]" pageId="4" pageNumber="205" value="2008-10-28">
<collectingDate id="91759E2AAA1A0572343BFABCFC0EF95D" box="[886,1000,1691,1717]" pageId="4" pageNumber="205" value="2008-10-28">28.x.2008</collectingDate>
</date>
, ex pupa
<date id="813167C2AA1A05723315FABCFB56F95D" box="[1112,1200,1691,1717]" pageId="4" pageNumber="205" value="2008-11">
<collectingDate id="91759E2AAA1A05723315FABCFB56F95D" box="[1112,1200,1691,1717]" pageId="4" pageNumber="205" value="2008-11">xi.2008</collectingDate>
</date>
, field card no. 4955, leg.
<collectorName id="587A24D4AA1A0572378AFA98FEDAF931" box="[199,316,1727,1753]" pageId="4" pageNumber="205">A. Diškus</collectorName>
, genitalia slide no. AD1185 (
<collectionCode id="939ED9C7AA1A057235C4FA98FD20F931" box="[649,710,1727,1753]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="4" pageNumber="205" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2A34FFA0AA1A05723594FA98FCA0F931" box="[729,838,1727,1753]" pageId="4" pageNumber="205" type="paratype">Paratypes</typeStatus>
<specimenCount id="E3898A8BAA1A05723403FA98FC9AF931" box="[846,892,1727,1753]" count="2" pageId="4" pageNumber="205" type="male">2 ♂</specimenCount>
,
<materialsCitation id="45E74B5FAA1A057234C7FA98FF27F915" collectingDate="2008-10-28" collectingDateMax="2008-11" collectingDateMin="2008-10-28" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="3320" latitude="-13.047223" location="Quinua" longLatPrecision="21" longitude="-74.13555" municipality="Huamanga Province" pageId="4" pageNumber="205" specimenCount="1" stateProvince="Ayacucho" typeStatus="paratype">same label data as holotype, genitalia slide no. AD</materialsCitation>
<materialsCitation id="45E74B5FAA1A0572378CFAC4FEB4F915" box="[193,338,1763,1789]" collectionCode="MfN" country="Peru" pageId="4" pageNumber="205" specimenCount="746" specimenCount-male="746" stateProvince="Ayacucho" typeStatus="paratype">
<specimenCount id="E3898A8BAA1A0572378CFAC4FF19F915" box="[193,255,1763,1789]" count="746" pageId="4" pageNumber="205" type="male">746♂</specimenCount>
(
<collectionCode id="939ED9C7AA1A05723643FAC4FEADF915" box="[270,331,1763,1789]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="4" pageNumber="205" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1A0573378AFB21FB73FD41" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205" type="diagnosis">
<paragraph id="F5304102AA1A0572378AFB21FC96F865" blockId="4.[151,1437,1654,2041]" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A0572378AFB21FEA4F8C8" bold="true" box="[199,322,1798,1824]" pageId="4" pageNumber="205">Diagnosis.</emphasis>
Externally, this species is characterized by a yellow-ochre forewing with irregular dark brown patches. In the male genitalia, the combination of a very slender and straight valva, a phallus with two wide and double-folded apical lobes, a basally greatly extended uncus, and wide, distally distinctly pointed dorsal lobes distinguishes the new species from all other species of the genus.
</paragraph>
<paragraph id="F5304102AA1A0573378AFBB0FDA4FEA9" blockId="4.[151,1437,1654,2041]" lastBlockId="5.[151,1437,151,825]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<emphasis id="C7FB9D10AA1A0572378AFBB0FEE2F859" bold="true" box="[199,260,1943,1969]" pageId="4" pageNumber="205">Male</emphasis>
(
<figureCitation id="6DB45D87AA1A0572365EFBB0FEBEF859" box="[275,344,1943,1969]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1467,1491]" captionTargetBox="[177,1410,179,1441]" captionTargetId="figure-18@3.[177,1410,179,1441]" captionTargetPageId="3" captionText="FIGURES 1–3. Adults of mountainous Astrotischeria species. 1, A. parapallens Diškus &amp; Stonis, 3,320 m, non-type specimen, a new elevation record; 2, A. peruanica Diškus &amp; Stonis, sp. nov., holotype, 2,110 m; 3, A. montivaga Diškus &amp; Stonis, sp. nov., paratype, 3,320 m (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757529" httpUri="https://zenodo.org/record/13757529/files/figure.png" pageId="4" pageNumber="205">Fig. 3</figureCitation>
). Forewing length ranges from 3.7 to
<quantity id="3277ECE7AA1A0572344EFBB0FCBBF859" box="[771,861,1943,1969]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="205" unit="mm" value="3.9">3.9 mm</quantity>
; wingspan from 8.1 to 8.
<quantity id="3277ECE7AA1A0572333FFBB0FB50F859" box="[1138,1206,1943,1969]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="mm" value="4.0">4 mm</quantity>
(n = 3). Head: frons pale ochre to yellow-ochre; palpi yellowish cream; frontal tuft dark grey proximally, transitioning to yellowish cream distally; collar yellowish cream; antenna distinctly longer than half the length of forewing; flagellum glossy dark grey or annulated with dark grey. Thorax pale yellowish with some grey scales; tegula predominantly grey with some yellowish scales. Forewing yellow-ochre to pale yellowish with irregular blackish brown to dark brown patches; fringe pale grey to grey; fringe line indistinctive or absent; forewing underside grey, without spots or androconia. Hindwing and its fringe pale greyish cream to grey (depending on the angle of view), grey on underside, without androconia. Legs glossy grey.
</paragraph>
<paragraph id="F5304102AA1B0573378AFD6CFE24FDD5" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFD6CFE96FE8C" bold="true" box="[199,368,330,357]" pageId="5" pageNumber="206">Male genitalia</emphasis>
(
<figureCitation id="6DB45D87AA1B05733633FD6CFDE6FE8D" box="[382,512,331,357]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Figs 24–28</figureCitation>
). Capsule measures 575–580 µm in length, 255–500 µm in width. Uncus comprised of two long lobes strongly widened basally, partially divided into two lobes: a long lateral lobe and a short rounded medial lobe (
<figureCitation id="6DB45D87AA1B05733665FDB4FE4FFE45" box="[296,425,402,429]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Figs 24, 25</figureCitation>
). Socii relatively small, membranous, covered with tiny spines. Valva divided: ventral lobe (main body) very narrow and straight (
<figureCitation id="6DB45D87AA1B0573351DFD90FD40FE39" box="[592,678,439,465]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Fig. 24</figureCitation>
); dorsal lobe inwardly curved, wide, and distinctly pointed distally (
<figureCitation id="6DB45D87AA1B057337D2FDFCFEC5FE1D" box="[159,291,474,501]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Figs 24, 25</figureCitation>
). Transtilla absent. Anellus only slightly thickened laterally with about three chetae on each side (
<figureCitation id="6DB45D87AA1B05733222FDFCFF51FDF1" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Fig. 24</figureCitation>
). Phallus (
<figureCitation id="6DB45D87AA1B05733660FDD8FE48FDF1" box="[301,430,511,537]" captionStart="FIGURES 24–28" captionStartId="10.[151,264,1892,1916]" captionTargetBox="[163,1423,181,1868]" captionTargetId="figure-21@10.[163,1424,181,1868]" captionTargetPageId="10" captionText="FIGURES 24–28. Male genitalia of Astrotischeria montivaga Diškus &amp; Stonis, sp. nov., 3,320 m. 24, genitalia capsule with phallus removed, holotype, slide no. AD1185 (MfN); 25, 26, same, paratype, genitalia slide no. AD746 (MfN); 27, same, phallus; 28, same, holotype, genitalia slide no. AD1185 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757545" httpUri="https://zenodo.org/record/13757545/files/figure.png" pageId="5" pageNumber="206">Figs 27, 28</figureCitation>
) measures 535–540 µm in length, slender in basal half, broadly divided into two double-folded lobes in apical half.
</paragraph>
<paragraph id="F5304102AA1B0573378AFE60FE44FD89" blockId="5.[151,1437,151,825]" box="[199,418,583,609]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFE60FEC3FD89" bold="true" box="[199,293,583,609]" pageId="5" pageNumber="206">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="F5304102AA1B0573378AFE4DFB73FD41" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFE4DFEA5FD6C" bold="true" box="[199,323,618,644]" pageId="5" pageNumber="206">Bionomics</emphasis>
. Host plant is unknown, possibly
<taxonomicName id="328F3A81AA1B057335FEFE4CFCC3FD6D" box="[691,805,619,645]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="genus">
<emphasis id="C7FB9D10AA1B057335FEFE4CFCC3FD6D" box="[691,805,619,645]" italics="true" pageId="5" pageNumber="206">Baccharis</emphasis>
</taxonomicName>
L.,
<taxonomicName id="328F3A81AA1B05733406FE4CFC2CFD6D" box="[843,970,619,645]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
. Larvae mine leaves in October, producing irregular blotch-like mines. Adults occur in November. Otherwise, the biology is unknown.
</paragraph>
</subSubSection>
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<emphasis id="C7FB9D10AA1B0573378AFE95FE87FD24" bold="true" box="[199,353,690,716]" pageId="5" pageNumber="206">Distribution.</emphasis>
This species is currently known from a single locality in Quinua,
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NE of
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, Huamanga Province of
<collectingCountry id="8D980192AA1B057336EDFEF0FE30FD19" box="[416,470,727,753]" name="Peru" pageId="5" pageNumber="206">Peru</collectingCountry>
, at an elevation of
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(
<figureCitation id="6DB45D87AA1B05733458FEF0FC8CFD19" box="[789,874,727,753]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="5" pageNumber="206">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1B0573378AFEDCFC17FCD1" pageId="5" pageNumber="206" type="etymology">
<paragraph id="F5304102AA1B0573378AFEDCFC17FCD1" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFEDCFEAEFCFD" bold="true" box="[199,328,763,789]" pageId="5" pageNumber="206">Etymology</emphasis>
. The species name derived from the Latin
<emphasis id="C7FB9D10AA1B0573341AFEDCFC3EFCFC" box="[855,984,763,788]" italics="true" pageId="5" pageNumber="206">montivagus</emphasis>
(mountain-wandering or living in the mountains), in reference to the mountainous distribution of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>

252
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<mods:title id="05F929BA7D5E8BB25A8ACFE179B7A819">How high can trumpet moths occur: documentation of mountainous leaf-mining Tischeriidae, featuring a species from record-high elevations</mods:title>
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<emphasis id="C7FB9D10AA1B057337DAFFA5FE56FC75" bold="true" box="[151,432,898,925]" italics="true" pageId="5" pageNumber="206">Astrotischeria viscacha</emphasis>
Diškus &amp; Stonis
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http://zoobank.org/NomenclaturalActs/
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(
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,
<figureCitation id="6DB45D87AA1B0573365DFFECFEBFFC0D" box="[272,345,971,997]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">29–33</figureCitation>
,
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,
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<emphasis id="C7FB9D10AA1B057337DAF834FEAEFBC4" bold="true" box="[151,328,1042,1069]" pageId="5" pageNumber="206">Type material.</emphasis>
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, elevation ca.
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, mining larvae
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on
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<emphasis id="C7FB9D10AA1B0573362CF810FDADFBB9" box="[353,587,1079,1105]" italics="true" pageId="5" pageNumber="206">Baccharis salicifolia</emphasis>
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</collectedFrom>
,
<date id="813167C2AA1B05733516F810FD28FBB9" box="[603,718,1079,1105]" pageId="5" pageNumber="206" value="2008-10-22">
<collectingDate id="91759E2AAA1B05733516F810FD28FBB9" box="[603,718,1079,1105]" pageId="5" pageNumber="206" value="2008-10-22">22.x.2008</collectingDate>
</date>
, ex pupa
<date id="813167C2AA1B05733470F810FC73FBB9" box="[829,917,1079,1105]" pageId="5" pageNumber="206" value="2008-11">
<collectingDate id="91759E2AAA1B05733470F810FC73FBB9" box="[829,917,1079,1105]" pageId="5" pageNumber="206" value="2008-11">xi.2008</collectingDate>
</date>
, field card no. 4949, leg.
<collectorName id="587A24D4AA1B057333F7F810FAC9FBB9" box="[1210,1327,1079,1105]" pageId="5" pageNumber="206">A. Diškus</collectorName>
, genitalia slide no. AD1177
<specimenCount id="E3898A8BAA1B05733610F87CFE96FB9D" box="[349,368,1115,1141]" count="1" pageId="5" pageNumber="206" type="male">♂</specimenCount>
(
<collectionCode id="939ED9C7AA1B05733632F87CFE5AFB9D" box="[383,444,1115,1141]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="5" pageNumber="206" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2A34FFA0AA1B05733683F87CFDDDFB9D" box="[462,571,1115,1141]" pageId="5" pageNumber="206" type="paratype">Paratypes</typeStatus>
<specimenCount id="E3898A8BAA1B0573350FF87CFD8AFB9D" box="[578,620,1115,1141]" count="2" pageId="5" pageNumber="206" type="female">2 ♀</specimenCount>
,
<materialsCitation id="45E74B5FAA1B05733537F87CFB08FB9D" box="[634,1262,1115,1141]" collectingDate="2008-10-22" collectingDateMax="2008-11" collectingDateMin="2008-10-22" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="2990" latitude="-13.42139" location="Calca Province" longLatPrecision="21" longitude="-71.84334" municipality="Pisac" pageId="5" pageNumber="206" specimenCount="1" typeStatus="paratype">same label data as holotype, genitalia slide no. AD1176</materialsCitation>
<materialsCitation id="45E74B5FAA1B057333A3F87CFAB6FB9D" box="[1262,1360,1115,1141]" collectionCode="MfN" country="Peru" pageId="5" pageNumber="206" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="E3898A8BAA1B057333A3F87CFB1AFB9D" box="[1262,1276,1115,1141]" count="1" pageId="5" pageNumber="206" type="female">♀</specimenCount>
(
<collectionCode id="939ED9C7AA1B05733246F87CFAAEFB9D" box="[1291,1352,1115,1141]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="5" pageNumber="206" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1B0570378AF859FBCFFEA9" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206" type="diagnosis">
<paragraph id="F5304102AA1B0573378AF859FD4FFAED" blockId="5.[151,1437,1042,2041]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AF859FEA4FB70" bold="true" box="[199,322,1150,1176]" pageId="5" pageNumber="206">Diagnosis.</emphasis>
This species exhibits strong sexual dimorphism; males are characterized by irrorated forewings, while females display distinctive cream and dark brown oblique patches on the forewings. In the male genitalia, the combination of a chunky, wide uncus and a slender phallus with unique, T-shaped lateral lobes distinguishes the new species from all other species within the genus.
</paragraph>
<paragraph id="F5304102AA1B0573378AF928FBF6FA35" blockId="5.[151,1437,1042,2041]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AF928FEE2FAC1" bold="true" box="[199,260,1295,1321]" pageId="5" pageNumber="206">Male</emphasis>
(
<figureCitation id="6DB45D87AA1B0573365FF928FEB0FAC1" box="[274,342,1295,1321]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1768,1792]" captionTargetBox="[212,1375,181,1744]" captionTargetId="figure-18@7.[212,1375,181,1744]" captionTargetPageId="7" captionText="FIGURES 4–10. Adults of mountainous Astrotischeria species. 4, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, male holotype; 5, 6, same, female paratype; 7, A. andina Diškus &amp; Stonis, sp. nov., male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="5" pageNumber="206">Fig. 4</figureCitation>
). Forewing length
<quantity id="3277ECE7AA1B05733568F928FD9AFAC1" box="[549,636,1295,1321]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="206" unit="mm" value="4.8">4.8 mm</quantity>
; wingspan
<quantity id="3277ECE7AA1B057335BAF928FCBCFAC1" box="[759,858,1295,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="206" unit="mm" value="10.3">10.3 mm</quantity>
(n = 1). Head: frons glossy grey cream; palpi cream; frontal tuft and collar glossy grey cream; antenna slightly longer than half the length of forewing; flagellum pale brown to dark brown. Thorax and tegulae glossy grey cream. Forewing glossy grey cream, densely irrorated with ochre and numerous dark brown scales, some ochre-tipped; fringe pale ochreous brown; fringe line indistinctive or absent; forewing underside pale brown, without spots or androconia. Hindwing and its fringe pale grey to pale ochreous brown (angle-dependent), without androconia. Legs pale grey-brown.
</paragraph>
<paragraph id="F5304102AA1B0573378AF9C0FAD8F979" blockId="5.[151,1437,1042,2041]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AF9C0FE96F9E8" bold="true" box="[199,368,1510,1537]" pageId="5" pageNumber="206">Male genitalia</emphasis>
(
<figureCitation id="6DB45D87AA1B05733633F9C0FDE6F9E9" box="[382,512,1511,1537]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Figs 29–33</figureCitation>
). Capsule 550 µm long, 250 µm wide. Uncus relatively short and chunky, with wide lateral lobes and short, wide inner lobes (
<figureCitation id="6DB45D87AA1B05733512FA2CFD07F9CD" box="[607,737,1547,1573]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Figs 29, 30</figureCitation>
). Socii relatively large, membranous, covered with tiny spines. Valva divided: ventral lobe (main body) very slender and slightly sinuous (
<figureCitation id="6DB45D87AA1B057334A1FA08FBA7F9A1" box="[1004,1089,1583,1609]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Fig. 31</figureCitation>
); dorsal lobe inwardly curved, strongly chitinized, distinctly pointed distally (
<figureCitation id="6DB45D87AA1B057335EBFA74FD1CF985" box="[678,762,1619,1645]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Fig. 31</figureCitation>
). Transtilla absent. Anellus weakly chitinized, indistinctive. Phallus (
<figureCitation id="6DB45D87AA1B057337B5FA50FE9EF979" box="[248,376,1655,1681]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Figs 32, 33</figureCitation>
) 420 µm long, slender, deeply divided apically, with T-shaped lobes distally (
<figureCitation id="6DB45D87AA1B05733390FA50FAD4F979" box="[1245,1330,1655,1681]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1911,1935]" captionTargetBox="[165,1421,181,1886]" captionTargetId="figure-18@11.[165,1421,181,1887]" captionTargetPageId="11" captionText="FIGURES 29–33. Male genitalia of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, holotype, genitalia slide no. AD1177 (MfN). 29, 30, uncus, 31, genitalia capsule with phallus removed, focused on valvae; 32, 33, phallus." figureDoi="http://doi.org/10.5281/zenodo.13757549" httpUri="https://zenodo.org/record/13757549/files/figure.png" pageId="5" pageNumber="206">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="F5304102AA1B0573378AFABCFD9AF881" blockId="5.[151,1437,1042,2041]" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFABCFEF8F95D" bold="true" box="[199,286,1691,1717]" pageId="5" pageNumber="206">Female</emphasis>
(
<figureCitation id="6DB45D87AA1B05733660FABCFE74F95D" box="[301,402,1690,1717]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1768,1792]" captionTargetBox="[212,1375,181,1744]" captionTargetId="figure-18@7.[212,1375,181,1744]" captionTargetPageId="7" captionText="FIGURES 4–10. Adults of mountainous Astrotischeria species. 4, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, male holotype; 5, 6, same, female paratype; 7, A. andina Diškus &amp; Stonis, sp. nov., male holotype, 3,595–3,600 m; 8, same, male paratype; 9, 10, same, female paratype (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757533" httpUri="https://zenodo.org/record/13757533/files/figure.png" pageId="5" pageNumber="206">Figs 5, 6</figureCitation>
). Forewing length
<quantity id="3277ECE7AA1B05733525FABDFD14F95D" box="[616,754,1690,1717]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="5" pageNumber="206" unit="mm" value="4.4" valueMax="4.5" valueMin="4.3">4.3–4.5 mm</quantity>
; wingspan
<quantity id="3277ECE7AA1B0573343DFABCFC11F95D" box="[880,1015,1691,1717]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="9.7" metricValueMin="9.3" pageId="5" pageNumber="206" unit="mm" value="9.5" valueMax="9.7" valueMin="9.3">9.3–9.7 mm</quantity>
(n = 2). Head: frons ochreous cream; palpi cream; frontal tuft and collar yellow ochre but pale grey-brown distally and laterally; antenna slightly longer than half the length of forewing; flagellum pale, sparsely annulated with brown scales. Thorax and tegulae yellow-ochre. Forewing with oblique yellow-ochre and dark brown patches; fringe pale ochre; fringe line indistinctive or absent; forewing underside pale ochreous brown, without spots or androconia. Hindwing and its fringe grey cream, without androconia. Legs pale grey-brown.
</paragraph>
<paragraph id="F5304102AA1B0570378AFB54FBFBFF11" blockId="5.[151,1437,1042,2041]" lastBlockId="6.[151,1436,151,501]" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206">
<emphasis id="C7FB9D10AA1B0573378AFB54FE68F864" bold="true" box="[199,398,1906,1933]" pageId="5" pageNumber="206">Female genitalia</emphasis>
(
<figureCitation id="6DB45D87AA1B057336D2FB54FDC5F865" box="[415,547,1907,1933]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="5" pageNumber="206">Figs 43, 44</figureCitation>
) 840 µm long. Ovipositor lobes large, flattened, clothed with short, modified setae ('peg setae'); area between ovipositor lobes relatively narrow, with tiny indistinctive papillae and some short setae. Second pair of lobes, lateral and anterior to the ovipositor lobes, three to four times smaller than ovipositor lobes, but bearing setae. Anterior apophyses distinctly shorter than posterior apophyses; prela comprised of two pairs of unique projections (
<figureCitation id="6DB45D87AA180570369BFCB0FDCAFF59" box="[470,556,151,177]" captionStart="FIGURES 43–46" captionStartId="13.[151,264,1899,1923]" captionTargetBox="[159,1424,181,1876]" captionTargetId="figure-18@13.[159,1427,181,1876]" captionTargetPageId="13" captionText="FIGURES 43–46. Female genitalia of mountainous Astrotischeria species. 43, 44, A. viscacha Diškus &amp; Stonis, sp. nov., 2,990 m, paratype, genitalia slide no.AD1176 (MfN); 45, 46, A. andina Diškus &amp; Stonis, sp. nov., 3,595–3,600 m, paratype, genitalia slide no. AD1179 (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757555" httpUri="https://zenodo.org/record/13757555/files/figure.png" pageId="6" pageNumber="207">Fig. 44</figureCitation>
). Corpus bursae with a very long and slender &quot;neck&quot;; the main part small and oval-shaped, without distinctive pectinations or signum. Accessory sac absent. Ductus spermathecae short and very slender, with about 4–5 medium-large or small coils and an indistinctive vesicle.
</paragraph>
<paragraph id="F5304102AA180570378AFD25FBCFFEA9" blockId="6.[151,1436,151,501]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFD25FEA5FEF4" bold="true" box="[199,323,258,284]" pageId="6" pageNumber="207">Bionomics</emphasis>
. Host plant is
<taxonomicName id="328F3A81AA18057036B9FD24FC37FEF5" authority="(Ruiz &amp; Pav.) Pers." authorityName="Pers." baseAuthorityName="Ruiz &amp; Pav." box="[500,977,259,285]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis id="C7FB9D10AA18057036B9FD24FD06FEF5" box="[500,736,259,285]" italics="true" pageId="6" pageNumber="207">Baccharis salicifolia</emphasis>
(Ruiz &amp; Pav.) Pers.
</taxonomicName>
,
<taxonomicName id="328F3A81AA1805703493FD24FBBFFEF5" box="[990,1113,259,285]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
(
<figureCitation id="6DB45D87AA1805703326FD24FB13FEF5" box="[1131,1269,258,285]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="6" pageNumber="207">Figs 51, 52</figureCitation>
). Larvae mine leaves in October; adults occur in November. Otherwise, the biology is unknown.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA180570378AFD6DFE60FE61" pageId="6" pageNumber="207" type="distribution">
<paragraph id="F5304102AA180570378AFD6DFE60FE61" blockId="6.[151,1436,151,501]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFD6DFE87FE8C" bold="true" box="[199,353,330,356]" pageId="6" pageNumber="207">Distribution.</emphasis>
This species is currently known from a single locality in
<collectingCountry id="8D980192AA180570334FFD6CFBD3FE8D" box="[1026,1077,331,357]" name="Peru" pageId="6" pageNumber="207">Peru</collectingCountry>
(Pisac, Calca Province), at the elevation of
<quantity id="3277ECE7AA180570366EFD48FE64FE61" box="[291,386,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.9899999999999998" pageId="6" pageNumber="207" unit="m" value="2990.0">2,990 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA180570378AFDB4FDB5FE1D" pageId="6" pageNumber="207" type="etymology">
<paragraph id="F5304102AA180570378AFDB4FDB5FE1D" blockId="6.[151,1436,151,501]" pageId="6" pageNumber="207">
<emphasis id="C7FB9D10AA180570378AFDB4FEAEFE45" bold="true" box="[199,328,403,429]" pageId="6" pageNumber="207">Etymology</emphasis>
. The species name is derived from the name of the viscacha, rodents of the family
<taxonomicName id="328F3A81AA1805703248FDB4FA7AFE45" authorityName="Bennett" authorityYear="1833" box="[1285,1436,403,429]" class="Mammalia" family="Chinchillidae" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="207" phylum="Chordata" rank="family">Chinchillidae</taxonomicName>
living in the Andes. It refers to the specific chunky uncus found in the male genitalia of the new species, which reminds one of the ears of the viscacha.
</paragraph>
</subSubSection>
</treatment>
</document>

287
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<mods:namePart id="F0C0BA1CFE1F12EC14940B8B3690CFCB">Diškus, Arūnas</mods:namePart>
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<mods:affiliation id="B75824E0A7201C9EF00C8C74019A45B5">Vytautas Magnus University, K. Donelaičio g. 58, 44248, Kaunas, Lithuania</mods:affiliation>
<mods:nameIdentifier id="F46DA6959B3697B90C731965444A5A80" type="email">diskus.biotaxonomy@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="A8649C38026AB0CE2E6DDADDC1EC86DA">Remeikis, Andrius</mods:namePart>
<mods:nameIdentifier id="C047AA140FBF6AA021255C57049D980D" type="ORCID">0000-0002-9310-1112</mods:nameIdentifier>
<mods:affiliation id="A373E7E7547DABA78F29DE47E5F9D0A8">State Research Institute Nature Research Centre, Akademijos g. 2, Vilnius 08412, Lithuania</mods:affiliation>
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<mods:namePart id="0AB517E26610D41629813C4A05728549">Orlovskytė, Svetlana</mods:namePart>
<mods:nameIdentifier id="D7DEA3F60765689448BD4FCAD50DA15F" type="ORCID">0000-0002-1643-7712</mods:nameIdentifier>
<mods:affiliation id="5A75A1BC18D9228F4D8515952B2DE3A4">State Research Institute Nature Research Centre, Akademijos g. 2, Vilnius 08412, Lithuania</mods:affiliation>
<mods:nameIdentifier id="A76C3A0E5C9509CB49E8AA1E23961EE9" type="email">s.orlovskyte@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2B16BA7CE2C5108ECEED6B4114EBBD03">Katinas, Liliana</mods:namePart>
<mods:nameIdentifier id="749FACDCD17E3D3AA802BF81A920F679" type="ORCID">0000-0002-6335-5402</mods:nameIdentifier>
<mods:affiliation id="9269AC9F0B9AB1630771A0D16ABC9DB7">Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina</mods:affiliation>
<mods:nameIdentifier id="9E4824C9442CBDBC00CE28DF2DC387EA" type="email">katinas@fcnym.unlp.edu.ar</mods:nameIdentifier>
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<mods:date id="4A92F1C30EE61082725BF92BEF4819B9">2024</mods:date>
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<treatment id="7D26F014AA1C057237DAFA85FA99FF11" ID-DOI="http://doi.org/10.5281/zenodo.13757531" ID-Zenodo-Dep="13757531" LSID="urn:lsid:plazi:treatment:7D26F014AA1C057237DAFA85FA99FF11" httpUri="http://treatment.plazi.org/id/7D26F014AA1C057237DAFA85FA99FF11" lastPageId="4" lastPageNumber="205" pageId="2" pageNumber="203">
<subSubSection id="BD951289AA1C057437DAFA85FBF8F8ED" pageId="2" pageNumber="203" type="nomenclature">
<paragraph id="F5304102AA1C057437DAFA85FF0CF909" blockId="2.[151,1436,1698,1833]" pageId="2" pageNumber="203">
<heading id="AE78F66EAA1C057437DAFA85FF0CF909" bold="true" centered="true" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
<emphasis id="C7FB9D10AA1C057437DAFA85FF0CF909" bold="true" pageId="2" pageNumber="203">
<taxonomicName id="328F3A81AA1C057437DAFA85FCEDF954" ID-CoL="BHY64" authority="Diskus &amp; Stonis, 2021" authorityName="Diskus &amp; Stonis" authorityYear="2021" box="[151,779,1698,1725]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="parapallens">
<emphasis id="C7FB9D10AA1C057437DAFA85FE3DF955" bold="true" box="[151,475,1698,1725]" italics="true" pageId="2" pageNumber="203">Astrotischeria parapallens</emphasis>
Diškus &amp; Stonis, 2021
</taxonomicName>
: photographic documentation and a new elevation record
</emphasis>
</heading>
</paragraph>
<paragraph id="F5304102AA1C057437DAFACCFBF8F8ED" blockId="2.[151,1436,1698,1833]" box="[151,1054,1771,1797]" pageId="2" pageNumber="203">http://zoobank.org/NomenclaturalActs/3ae6711c-34c4-404d-befd-f29c9c048b02</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1C057437DAFB28FE8AF8C1" box="[151,364,1806,1833]" pageId="2" pageNumber="203" type="description">
<paragraph id="F5304102AA1C057437DAFB28FE8AF8C1" blockId="2.[151,1436,1698,1833]" box="[151,364,1806,1833]" pageId="2" pageNumber="203">
(
<figureCitation id="6DB45D87AA1C057437D2FB28FF00F8C1" box="[159,230,1807,1833]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1467,1491]" captionTargetBox="[177,1410,179,1441]" captionTargetId="figure-18@3.[177,1410,179,1441]" captionTargetPageId="3" captionText="FIGURES 1–3. Adults of mountainous Astrotischeria species. 1, A. parapallens Diškus &amp; Stonis, 3,320 m, non-type specimen, a new elevation record; 2, A. peruanica Diškus &amp; Stonis, sp. nov., holotype, 2,110 m; 3, A. montivaga Diškus &amp; Stonis, sp. nov., paratype, 3,320 m (MfN)." figureDoi="http://doi.org/10.5281/zenodo.13757529" httpUri="https://zenodo.org/record/13757529/files/figure.png" pageId="2" pageNumber="203">Figs 1</figureCitation>
,
<figureCitation id="6DB45D87AA1C057437B9FB29FEDDF8C1" box="[244,315,1806,1833]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1873,1897]" captionTargetBox="[160,1427,181,1849]" captionTargetId="figure-21@8.[160,1427,181,1849]" captionTargetPageId="8" captionText="FIGURES 11–15. Male genitalia of Astrotischeria parapallens Diškus &amp; Stonis, 3,320 m (a new elevation record), non-type specimen, genitalia slide no. AD1140 (MfN). 11, genitalia capsule with phallus removed, focused on the ventral view; 12, same, focused on the dorsal view; 13, anellus; 14, phallus; 15, basal processes of valvae." figureDoi="http://doi.org/10.5281/zenodo.13757537" httpUri="https://zenodo.org/record/13757537/files/figure.png" pageId="2" pageNumber="203">11–15</figureCitation>
,
<figureCitation id="6DB45D87AA1C0574360AFB28FE82F8C1" box="[327,356,1807,1833]" captionStart="FIGURES 47–52" captionStartId="14.[151,264,1862,1886]" captionTargetBox="[174,1413,181,1837]" captionTargetId="figure-21@14.[174,1413,181,1838]" captionTargetPageId="14" captionText="FIGURES 47–52. Bionomics of mountainous Astrotischeria species. 47, habitat of A. parapallens Diškus &amp; Stonis and A. montivaga Diškus &amp; Stonis, sp. nov., Ayacucho, Huamanga Province, Peru, 3,320 m; 48, unidentified host plant of A. peruanica Diškus &amp; Stonis, sp. nov., possibly Baccharis trinervis Pers., Asteraceae, 2,110 m; 49, 50, leaf mines with feeding larvae of Astrotischeria peruanica Diškus &amp; Stonis, sp. nov.; 51, 52, Baccharis salicifolia (Ruiz &amp; Pav.) Pers., Asteraceae, a host plant of Astrotischeria viscacha Diškus &amp; Stonis, sp. nov., Pisac, Calca Province, Peru at an elevation of 2,990 m." figureDoi="http://doi.org/10.5281/zenodo.13757557" httpUri="https://zenodo.org/record/13757557/files/figure.png" pageId="2" pageNumber="203">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1C057437DAFB71FD90F851" pageId="2" pageNumber="203" type="materials_examined">
<paragraph id="F5304102AA1C057437DAFB71FD90F851" blockId="2.[151,1437,1878,1977]" pageId="2" pageNumber="203">
<emphasis id="C7FB9D10AA1C057437DAFB71FE62F898" bold="true" box="[151,388,1878,1904]" pageId="2" pageNumber="203">Material examined.</emphasis>
<materialsCitation id="45E74B5FAA1C057436C3FB70FD94F851" collectingDate="2008-10-28" collectingDateMax="2008-11" collectingDateMin="2008-10-28" collectionCode="MfN" collectorName="A. Diskus" country="Peru" elevation="3320" latitude="-13.047223" location="Quinua" longLatPrecision="21" longitude="-74.13555" municipality="Huamanga Province" pageId="2" pageNumber="203" specimenCount="1" specimenCount-male="1" stateProvince="Ayacucho">
<specimenCount id="E3898A8BAA1C057436C3FB70FE58F899" box="[398,446,1879,1905]" count="1" pageId="2" pageNumber="203" type="male">1 ♂</specimenCount>
,
<collectingCountry id="8D980192AA1C05743680FB70FDF1F899" box="[461,535,1879,1905]" name="Peru" pageId="2" pageNumber="203">PERU</collectingCountry>
,
<collectingMunicipality id="1554DB78AA1C0574356BFB70FCF6F899" box="[550,784,1879,1905]" pageId="2" pageNumber="203">Huamanga Province</collectingMunicipality>
,
<quantity id="3277ECE7AA1C05743452FB70FC8FF899" box="[799,873,1879,1905]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="203" unit="km" value="37.0">37 km</quantity>
NE of
<collectingRegion id="374B8FE0AA1C0574348EFB70FBD3F899" box="[963,1077,1879,1905]" country="Peru" name="Ayacucho" pageId="2" pageNumber="203">Ayacucho</collectingRegion>
,
<location id="F05017D9AA1C0574330EFB70FB7EF899" LSID="urn:lsid:plazi:treatment:7D26F014AA1C057237DAFA85FA99FF11:F05017D9AA1C0574330EFB70FB7EF899" box="[1091,1176,1879,1905]" country="Peru" latitude="-13.047223" longLatPrecision="21" longitude="-74.13555" municipality="Huamanga Province" name="Quinua" pageId="2" pageNumber="203" stateProvince="Ayacucho">Quinua</location>
,
<geoCoordinate id="90BB27C5AA1C057433EAFB71FAFDF899" box="[1191,1307,1878,1905]" degrees="13" direction="south" minutes="2" orientation="latitude" pageId="2" pageNumber="203" precision="15" seconds="50" value="-13.047223">13°2'50&quot;S</geoCoordinate>
,
<geoCoordinate id="90BB27C5AA1C05743267FB70FA7EF899" box="[1322,1432,1879,1905]" degrees="74" direction="west" minutes="8" orientation="longitude" pageId="2" pageNumber="203" precision="15" seconds="8" value="-74.13555">74°8'8&quot;W</geoCoordinate>
, elevation ca.
<quantity id="3277ECE7AA1C0574367CFB5CFE74F87D" box="[305,402,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="2" pageNumber="203" unit="m" value="3320.0">
<elevation id="7EA2A631AA1C0574367CFB5CFE74F87D" box="[305,402,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="2" pageNumber="203" unit="m" value="3320.0">3,320 m</elevation>
</quantity>
, mining larvae
<collectedFrom id="469302FFAA1C0574350AFB5CFCECF87D" box="[583,778,1915,1941]" pageId="2" pageNumber="203">
on
<taxonomicName id="328F3A81AA1C05743520FB5CFCECF87D" box="[621,778,1915,1941]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="203" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="C7FB9D10AA1C05743520FB5CFD39F87D" box="[621,735,1915,1941]" italics="true" pageId="2" pageNumber="203">Baccharis</emphasis>
sp.
</taxonomicName>
</collectedFrom>
,
<date id="813167C2AA1C05743454FB5CFC6DF87D" box="[793,907,1915,1941]" pageId="2" pageNumber="203" value="2008-10-28">
<collectingDate id="91759E2AAA1C05743454FB5CFC6DF87D" box="[793,907,1915,1941]" pageId="2" pageNumber="203" value="2008-10-28">28.x.2008</collectingDate>
</date>
, ex pupa
<date id="813167C2AA1C057434B3FB5CFBB3F87D" box="[1022,1109,1915,1941]" pageId="2" pageNumber="203" value="2008-11">
<collectingDate id="91759E2AAA1C057434B3FB5CFBB3F87D" box="[1022,1109,1915,1941]" pageId="2" pageNumber="203" value="2008-11">xi.2008</collectingDate>
</date>
, field card no. 4955, leg.
<collectorName id="587A24D4AA1C057432CCFB5CFF0EF851" pageId="2" pageNumber="203">A. Diškus</collectorName>
, genitalia slide no. AD1140 (
<collectionCode id="939ED9C7AA1C05743560FBB8FD8CF851" box="[557,618,1951,1977]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="2" pageNumber="203" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="A1F0118AAA1D057537DAF99CFE31F9F4" ID-DOI="http://doi.org/10.5281/zenodo.13757529" ID-Zenodo-Dep="13757529" httpUri="https://zenodo.org/record/13757529/files/figure.png" pageId="3" pageNumber="204" startId="3.[151,264,1467,1491]" targetBox="[177,1410,179,1441]" targetPageId="3" targetType="figure">
<paragraph id="F5304102AA1D057537DAF99CFE31F9F4" blockId="3.[151,1437,1467,1564]" pageId="3" pageNumber="204">
<emphasis id="C7FB9D10AA1D057537DAF99CFEDCFA3C" bold="true" box="[151,314,1467,1492]" pageId="3" pageNumber="204">FIGURES 1–3.</emphasis>
Adults of mountainous
<taxonomicName id="328F3A81AA1D05753566F99BFCC7FA3C" box="[555,801,1468,1492]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="C7FB9D10AA1D05753566F99BFD5CFA3C" box="[555,698,1468,1492]" italics="true" pageId="3" pageNumber="204">Astrotischeria</emphasis>
species. 1
</taxonomicName>
,
<taxonomicName id="328F3A81AA1D05753460F99BFB89FA3C" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" box="[813,1135,1468,1492]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="parapallens">
<emphasis id="C7FB9D10AA1D05753460F99BFC27FA3C" box="[813,961,1468,1492]" italics="true" pageId="3" pageNumber="204">A. parapallens</emphasis>
Diškus &amp; Stonis
</taxonomicName>
, 3,320 m, non-type specimen, a new elevation record; 2,
<taxonomicName id="328F3A81AA1D057536E1F9C7FD02FA10" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[428,740,1504,1528]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="peruanica" status="sp. nov.">
<emphasis id="C7FB9D10AA1D057536E1F9C7FDD7FA1F" box="[428,561,1504,1527]" italics="true" pageId="3" pageNumber="204">A. peruanica</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="C7FB9D10AA1D057535BDF9C7FCA4FA10" bold="true" box="[752,834,1504,1528]" pageId="3" pageNumber="204">
<taxonomicNameLabel id="DCC8206BAA1D057535BDF9C7FCA4FA10" box="[752,834,1504,1528]" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, 2,110 m; 3,
<taxonomicName id="328F3A81AA1D0575337BF9C7FA94FA10" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2024" box="[1078,1394,1504,1528]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="montivaga" status="sp. nov.">
<emphasis id="C7FB9D10AA1D0575337BF9C7FB58FA1F" box="[1078,1214,1504,1527]" italics="true" pageId="3" pageNumber="204">A. montivaga</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="C7FB9D10AA1D05753233F9C7FF25F9F4" bold="true" pageId="3" pageNumber="204">
<taxonomicNameLabel id="DCC8206BAA1D05753233F9C7FF25F9F4" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, 3,320 m (MfN).
</paragraph>
</caption>
<subSubSection id="BD951289AA1D0575378AFA49FB96F938" pageId="3" pageNumber="204" type="diagnosis">
<paragraph id="F5304102AA1D0575378AFA49FB96F938" blockId="3.[151,1437,1646,2032]" pageId="3" pageNumber="204">
<emphasis id="C7FB9D10AA1D0575378AFA49FEA4F960" bold="true" box="[199,322,1646,1672]" pageId="3" pageNumber="204">Diagnosis.</emphasis>
Externally, this species can be confused with other densely speckled
<taxonomicName id="328F3A81AA1D05753318FA49FAB5F960" box="[1109,1363,1646,1672]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C7FB9D10AA1D05753318FA49FB15F960" box="[1109,1267,1646,1672]" italics="true" pageId="3" pageNumber="204">Astrotischeria</emphasis>
species.
</taxonomicName>
In the male genitalia,
<taxonomicName id="328F3A81AA1D05753609FAB5FE0CF944" box="[324,490,1682,1708]" class="Insecta" family="Tischeriidae" genus="Astrotischeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="parapallens">
<emphasis id="C7FB9D10AA1D05753609FAB5FE0CF944" box="[324,490,1682,1708]" italics="true" pageId="3" pageNumber="204">A. parapallens</emphasis>
</taxonomicName>
is distinguished by a slender and straight valva, a relatively short and distally widelobed phallus, an anteriorly thickened vinculum, and distally wide, dentate dorsal lobes.
</paragraph>
</subSubSection>
<subSubSection id="BD951289AA1D0575378AFAFDFD84F818" pageId="3" pageNumber="204" type="description">
<paragraph id="F5304102AA1D0575378AFAFDFCF2F86C" blockId="3.[151,1437,1646,2032]" pageId="3" pageNumber="204">
<emphasis id="C7FB9D10AA1D0575378AFAFDFEB7F91C" bold="true" box="[199,337,1754,1780]" pageId="3" pageNumber="204">Description</emphasis>
. The species was described by
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Stonis
<emphasis id="C7FB9D10AA1D05753456FAFCFCB7F91C" box="[795,849,1754,1780]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. (2021b)
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. The newly examined specimen from an altitude of
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, with a forewing length of
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and a wingspan of
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(n = 1) (
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), is larger than previously known specimens (3.9–4.0 mm in forewing length,
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in wingspan). In the male genitalia (
<figureCitation id="6DB45D87AA1D05753227FB05FF39F889" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1873,1897]" captionTargetBox="[160,1427,181,1849]" captionTargetId="figure-21@8.[160,1427,181,1849]" captionTargetPageId="8" captionText="FIGURES 11–15. Male genitalia of Astrotischeria parapallens Diškus &amp; Stonis, 3,320 m (a new elevation record), non-type specimen, genitalia slide no. AD1140 (MfN). 11, genitalia capsule with phallus removed, focused on the ventral view; 12, same, focused on the dorsal view; 13, anellus; 14, phallus; 15, basal processes of valvae." figureDoi="http://doi.org/10.5281/zenodo.13757537" httpUri="https://zenodo.org/record/13757537/files/figure.png" pageId="3" pageNumber="204">Figs 11–15</figureCitation>
), the capsule of the examined new specimen is 580 µm long and 280 µm wide; the phallus (
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) is 390 µm long. For a full description, see
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Stonis
<emphasis id="C7FB9D10AA1D05753538FB4CFD48F86C" box="[629,686,1898,1924]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
(2021b)
</bibRefCitation>
.
</paragraph>
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<emphasis id="C7FB9D10AA1D0575378AFBA9FEADF840" bold="true" box="[199,331,1934,1960]" pageId="3" pageNumber="204">Bionomics.</emphasis>
The host plant is
<taxonomicName id="328F3A81AA1D0575355BFBA9FD6EF840" box="[534,648,1934,1960]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="genus">
<emphasis id="C7FB9D10AA1D0575355BFBA9FD6EF840" box="[534,648,1934,1960]" italics="true" pageId="3" pageNumber="204">Baccharis</emphasis>
</taxonomicName>
L., possibly
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<emphasis id="C7FB9D10AA1D05753451FBA8FC73F840" box="[796,917,1934,1960]" italics="true" pageId="3" pageNumber="204">B. latifolia</emphasis>
(Ruiz &amp; Pav.) Pers.
</taxonomicName>
,
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. Larvae mine leaves in late October. The blotch mine is irregular, elongated, usually pale brownish, with little or no frass (
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Stonis
<emphasis id="C7FB9D10AA1D057537DAFBF0FF2EF818" box="[151,200,2006,2032]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2021b
</bibRefCitation>
). Adults occur in November.
</paragraph>
</subSubSection>
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<emphasis id="C7FB9D10AA1A0572378AFCB1FE87FF58" bold="true" box="[199,353,150,176]" pageId="4" pageNumber="205">Distribution.</emphasis>
Previously, this species was known from a single locality in
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(Muyurina,
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East of
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), at an elevation of about
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(
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Stonis
<emphasis id="C7FB9D10AA1A05723594FC9CFCF6FF3D" box="[729,784,187,213]" italics="true" pageId="4" pageNumber="205">et al.</emphasis>
2021b
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). Now, this species has been recorded in Quinua,
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NE of
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, Huamanga Province of
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, at an elevation of
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(a new elevation record) (
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).
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<emphasis id="1BAFEADEFFD2FFEBFF565697E503FCDE" bold="true" box="[151,314,758,785]" italics="true" pageId="1" pageNumber="479">
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<paragraph id="296436CCFFD2FFEBFF56577BE4D2FCFA" blockId="1.[151,415,758,821]" box="[151,235,795,821]" pageId="1" pageNumber="479">
(
<figureCitation id="B1E02A49FFD2FFEBFF5E577BE4D9FCFA" box="[159,224,795,821]" captionStart="FIGURE 1" captionStartId="1.[152,255,1869,1893]" captionTargetBox="[340,1247,974,1846]" captionTargetId="figure-337@1.[340,1247,974,1846]" captionTargetPageId="1" captionText="FIGURE 1. Garra dohjei¸ ZSI FF 10399, holotype, 99.6 mm SL; A. lateral, B. dorsal, C. ventral. ADBUFM/5090/8, paratype, 87.8 SL, male; D. lateral. E" figureDoi="http://doi.org/10.5281/zenodo.13757922" httpUri="https://zenodo.org/record/13757922/files/figure.png" pageId="1" pageNumber="479">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61C16547FFD2FFE8FF565703E6FAFF36" lastPageId="2" lastPageNumber="480" pageId="1" pageNumber="479" type="materials_examined">
<paragraph id="296436CCFFD2FFEBFF565703E616FC0A" blockId="1.[151,1437,867,965]" pageId="1" pageNumber="479">
<materialsCitation id="99B33C91FFD2FFEBFF565703E616FC0A" collectingDate="2022-03-06" collectorName="Batngenlang Mawlong &amp; N. Mylliem Umlong" country="India" county="Ri-bhoi district" elevation="837" latitude="25.67111" location="Niangdai village" longLatPrecision="21" longitude="91.82916" municipality="Niangdai River" pageId="1" pageNumber="479" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis id="1BAFEADEFFD2FFEBFF565703E534FCB2" bold="true" box="[151,269,867,893]" pageId="1" pageNumber="479">
<typeStatus id="F660886EFFD2FFEBFF565703E531FCB2" box="[151,264,867,893]" pageId="1" pageNumber="479" type="holotype">Holotype</typeStatus>
:
</emphasis>
ZSI FF 10399,
<quantity id="EE239B29FFD2FFEBFE025703E611FCB2" box="[451,552,867,893]" metricMagnitude="-2" metricUnit="m" metricValue="9.959999999999999" pageId="1" pageNumber="479" unit="mm" value="99.6">99.6 mm</quantity>
SL, female;
<collectingCountry id="51CC765CFFD2FFEBFD7A5703E6C0FCB2" box="[699,761,867,893]" name="India" pageId="1" pageNumber="479">India</collectingCountry>
:
<collectingRegion id="EB1FF82EFFD2FFEBFCC55703E7BAFCB2" box="[772,899,867,893]" country="India" name="Meghalaya" pageId="1" pageNumber="479">Meghalaya</collectingRegion>
,
<collectingCounty id="C0054E40FFD2FFEBFC4E5703E004FCB2" box="[911,1085,867,893]" pageId="1" pageNumber="479">Ri-bhoi district</collectingCounty>
,
<collectingMunicipality id="C900ACB6FFD2FFEBFB885704E0CCFCB2" box="[1097,1269,867,895]" pageId="1" pageNumber="479">Ñiangdai River</collectingMunicipality>
near
<location id="2C046017FFD2FFEBFAF75704E4DAFC6E" LSID="urn:lsid:plazi:treatment:A17287DAFFD2FFECFF565697E114FDAE:2C046017FFD2FFEBFAF75704E4DAFC6E" country="India" county="Ri-bhoi district" latitude="25.67111" longLatPrecision="21" longitude="91.82916" municipality="Niangdai River" name="Niangdai village" pageId="1" pageNumber="479" stateProvince="Meghalaya">Ñiangdai village</location>
(Brahmaputra basin),
<geoCoordinate id="4CEF500BFFD2FFEBFE2157E8E65FFC6C" box="[480,614,904,931]" degrees="25" direction="north" minutes="40" orientation="latitude" pageId="1" pageNumber="479" precision="15" seconds="16" value="25.67111">25°40′16″N</geoCoordinate>
&amp;
<geoCoordinate id="4CEF500BFFD2FFEBFD4B57E8E737FC6C" box="[650,782,904,931]" degrees="91" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="479" precision="15" seconds="45" value="91.82916">91°49′45″E</geoCoordinate>
, altitude
<elevation id="A2F6D1FFFFD2FFEBFCB757E7E04AFC6E" box="[886,1139,903,929]" metricMagnitude="2" metricUnit="m" metricValue="8.37" pageId="1" pageNumber="479" unit="m" value="837.0">
<quantity id="EE239B29FFD2FFEBFCB757E7E784FC6E" box="[886,957,903,929]" metricMagnitude="2" metricUnit="m" metricValue="8.37" pageId="1" pageNumber="479" unit="m" value="837.0">837 m</quantity>
above sea level
</elevation>
;
<collectorName id="842E531AFFD2FFEBFBBE57E7E146FC6E" box="[1151,1407,903,929]" pageId="1" pageNumber="479">Batngenlang Mawlong</collectorName>
&amp;
<collectorName id="842E531AFFD2FFEBFF5657CBE5B9FC0A" box="[151,384,939,965]" pageId="1" pageNumber="479">N. Mylliem Umlong</collectorName>
,
<date id="5D65100CFFD2FFEBFE4A57CBE615FC0A" box="[395,556,939,965]" pageId="1" pageNumber="479" value="2022-03-06">
<collectingDate id="4D21E9E4FFD2FFEBFE4A57CBE615FC0A" box="[395,556,939,965]" pageId="1" pageNumber="479" value="2022-03-06">6 March 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="7DA46644FFD2FFEBFF59532DE594F847" ID-DOI="http://doi.org/10.5281/zenodo.13757922" ID-Zenodo-Dep="13757922" httpUri="https://zenodo.org/record/13757922/files/figure.png" pageId="1" pageNumber="479" startId="1.[152,255,1869,1893]" targetBox="[340,1247,974,1846]" targetPageId="1" targetType="figure">
<paragraph id="296436CCFFD2FFEBFF59532DE594F847" blockId="1.[151,1437,1869,1929]" pageId="1" pageNumber="479">
<emphasis id="1BAFEADEFFD2FFEBFF59532DE52EF8AA" bold="true" box="[152,279,1869,1893]" pageId="1" pageNumber="479">FIGURE 1.</emphasis>
<taxonomicName id="EEDB4D4FFFD2FFEBFEDC532DE5CEF8AB" authority="¸ ZSI FF" authorityName="ZSI FF" box="[285,503,1869,1893]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="479" phylum="Chordata" rank="species" species="dohjei">
<emphasis id="1BAFEADEFFD2FFEBFEDC532DE593F8AA" box="[285,426,1869,1893]" italics="true" pageId="1" pageNumber="479">Garra dohjei¸</emphasis>
ZSI FF
</taxonomicName>
10399, holotype, 99.6 mm SL;
<emphasis id="1BAFEADEFFD2FFEBFCF4532DE777F8AA" bold="true" box="[821,846,1869,1893]" pageId="1" pageNumber="479">A.</emphasis>
lateral,
<emphasis id="1BAFEADEFFD2FFEBFC5F532DE78CF8AA" bold="true" box="[926,949,1869,1893]" pageId="1" pageNumber="479">B.</emphasis>
dorsal,
<emphasis id="1BAFEADEFFD2FFEBFBC4532DE024F8AA" bold="true" box="[1029,1053,1869,1893]" pageId="1" pageNumber="479">C.</emphasis>
ventral. ADBUFM/5090/8, paratype, 87.8 SL, male;
<emphasis id="1BAFEADEFFD2FFEBFEF25311E572F846" bold="true" box="[307,331,1905,1929]" pageId="1" pageNumber="479">D.</emphasis>
lateral. E
</paragraph>
</caption>
<paragraph id="296436CCFFD2FFEBFF0653D7E1A5F81D" blockId="1.[151,1436,1975,2038]" box="[199,1436,1975,2002]" pageId="1" pageNumber="479">
<emphasis id="1BAFEADEFFD2FFEBFF0653D7E571F81E" bold="true" box="[199,328,1975,2001]" pageId="1" pageNumber="479">
<typeStatus id="F660886EFFD2FFEBFF0653D7E57AF81E" box="[199,323,1975,2001]" pageId="1" pageNumber="479" type="paratype">Paratypes</typeStatus>
:
</emphasis>
ZSI FF 10400,
<specimenCount id="3FDDFD45FFD2FFEBFE3F53D7E60EF81D" box="[510,567,1975,2002]" count="2" pageId="1" pageNumber="479" type="generic">2 ex.</specimenCount>
,
<materialsCitation id="99B33C91FFD2FFEBFD8553D7E1AEF81D" box="[580,1431,1975,2002]" collectingDate="2022-03-06" collectorName="Batngenlang Mawlong &amp; N. Mylliem Umlong" country="India" county="Ri-bhoi district" elevation="837" latitude="25.67111" location="Niangdai village" longLatPrecision="21" longitude="91.82916" municipality="Niangdai River" pageId="1" pageNumber="479" specimenCode="ADBU-FM/5090" specimenCount="4" stateProvince="Meghalaya" typeStatus="paratype">
<quantity id="EE239B29FFD2FFEBFD8553D7E6D1F81D" box="[580,744,1975,2002]" metricMagnitude="-2" metricUnit="m" metricValue="8.615" metricValueMax="9.65" metricValueMin="7.58" pageId="1" pageNumber="479" unit="mm" value="86.15" valueMax="96.5" valueMin="75.8">75.8–96.5 mm</quantity>
SL, males; data as for holotype.
<specimenCode id="797D9EB7FFD2FFEBFB9C53D7E11AF81D" box="[1117,1315,1975,2002]" collectionCode="ADBU-FM" pageId="1" pageNumber="479">ADBU-FM/5090</specimenCode>
/2–5,
<specimenCount id="3FDDFD45FFD2FFEBFA9F53D7E1AEF81D" box="[1374,1431,1975,2002]" count="4" pageId="1" pageNumber="479" type="generic">4 ex.</specimenCount>
</materialsCitation>
,
</paragraph>
<paragraph id="296436CCFFD2FFE8FF5653BBE6A2FF1A" blockId="1.[151,1436,1975,2038]" lastBlockId="2.[151,1437,150,933]" lastPageId="2" lastPageNumber="480" pageId="1" pageNumber="479">
<materialsCitation id="99B33C91FFD2FFE8FF5653BBE6AEFF1A" collectingDate="2022-03-06" collectorName="Batngenlang Mawlong &amp; N. Mylliem Umlong" country="India" county="Ri-bhoi district" elevation="837" lastPageId="2" lastPageNumber="480" latitude="25.67111" location="Niangdai village" longLatPrecision="21" longitude="91.82916" municipality="Niangdai River" pageId="1" pageNumber="479" specimenCode="ADBU-FM/5090" specimenCount="5" stateProvince="Meghalaya">
53.0–
<quantity id="EE239B29FFD2FFEBFF1653BBE573F839" box="[215,330,2011,2038]" metricMagnitude="-1" metricUnit="m" metricValue="1.152" pageId="1" pageNumber="479" unit="mm" value="115.2">115.2 mm</quantity>
SL, females;
<specimenCode id="797D9EB7FFD2FFEBFE2B53BBE68BF839" box="[490,690,2011,2038]" collectionCode="ADBU-FM" pageId="1" pageNumber="479">ADBU-FM/5090</specimenCode>
/8–11,
<specimenCount id="3FDDFD45FFD2FFEBFD3A53BBE70FF839" box="[763,822,2011,2038]" count="3" pageId="1" pageNumber="479" type="generic">3 ex.</specimenCount>
,
<quantity id="EE239B29FFD2FFEBFC8553BBE7D0F839" box="[836,1001,2011,2038]" metricMagnitude="-2" metricUnit="m" metricValue="6.56" metricValueMax="9.25" metricValueMin="3.87" pageId="1" pageNumber="479" unit="mm" value="65.6" valueMax="92.5" valueMin="38.7">38.7–92.5 mm</quantity>
SL, males; data as for holotype.
<specimenCount id="3FDDFD45FFD2FFE8FAAA53BCE535FF7E" count="2" lastPageId="2" lastPageNumber="480" pageId="1" pageNumber="479" type="generic">Two specimens</specimenCount>
(
<specimenCode id="797D9EB7FFD1FFE8FEDC54F6E5DDFF7E" box="[285,484,150,177]" collectionCode="ADBU-FM" pageId="2" pageNumber="480">ADBU-FM/5090</specimenCode>
/
<quantity id="EE239B29FFD1FFE8FE2C54F6E643FF7E" box="[493,634,150,177]" metricMagnitude="0" metricUnit="m" metricValue="3.1151999999999997" pageId="2" pageNumber="480" unit="mm" value="3115.2">3, 115.2 mm</quantity>
SL, female;
<specimenCode id="797D9EB7FFD1FFE8FCD154F6E7EEFF7E" box="[784,983,150,177]" collectionCode="ADBU-FM" pageId="2" pageNumber="480">ADBU-FM/5090</specimenCode>
/11,
<quantity id="EE239B29FFD1FFE8FBC554F6E052FF7E" box="[1028,1131,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" pageId="2" pageNumber="480" unit="mm" value="92.5">92.5 mm</quantity>
SL, male) were dissected, cleared and stained for biology and osteology
</materialsCitation>
.
</paragraph>
<paragraph id="296436CCFFD1FFE8FF0654BEE6FAFF36" blockId="2.[151,1437,150,933]" box="[199,707,222,249]" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FF0654BEE5BEFF36" bold="true" box="[199,391,222,249]" pageId="2" pageNumber="480">Common name:</emphasis>
‘
<emphasis id="1BAFEADEFFD1FFE8FE5954BFE5D2FF37" box="[408,491,223,249]" italics="true" pageId="2" pageNumber="480">Doh jei</emphasis>
’ (in Khasi dialect).
</paragraph>
</subSubSection>
<subSubSection id="61C16547FFD1FFE8FF065562E7A3FE1E" pageId="2" pageNumber="480" type="diagnosis">
<paragraph id="296436CCFFD1FFE8FF065562E7A3FE1E" blockId="2.[151,1437,150,933]" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FF065562E57BFED3" bold="true" box="[199,322,258,284]" pageId="2" pageNumber="480">Diagnosis.</emphasis>
<emphasis id="1BAFEADEFFD1FFE8FE875563E5E7FED2" box="[326,478,259,285]" italics="true" pageId="2" pageNumber="480">
<taxonomicName id="EEDB4D4FFFD1FFE8FE875563E5E3FED2" box="[326,474,259,285]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="480" phylum="Chordata" rank="species" species="dohjei">Garra dohjei</taxonomicName>
,
</emphasis>
is distinguished from its congeners by having the following combination of characters: a black spot immediately anterior to upper angle of gill opening, 5–6 dark black stripes on caudal peduncle; well-developed transverse lobe with 9–12 low minute tubercles, transverse groove deep; snout rounded with incipient proboscis; long pelvic fin reaching to midway of anus-anal distance when adpressed, surpassing anus; 33–34 lateral line scales, 9–10 predorsal scales, 16 circumpeduncular scales rows, transverse scale rows 4½/3½/4½; 8½ branched dorsal-fin rays, 15 branched pectoral-fin rays; and total vertebrae 33.
</paragraph>
</subSubSection>
<subSubSection id="61C16547FFD1FFECFF0655BAE607FEAA" lastPageId="6" lastPageNumber="484" pageId="2" pageNumber="480" type="description">
<paragraph id="296436CCFFD1FFE8FF0655BAE527FC6A" blockId="2.[151,1437,150,933]" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FF0655BAE568FE3B" bold="true" box="[199,337,474,500]" pageId="2" pageNumber="480">Description</emphasis>
. Morphometric data in
<tableCitation id="64590377FFD1FFE8FDAF55BBE6FEFE3A" box="[622,711,475,501]" captionStart="TABLE1" captionStartId="2.[151,239,980,1004]" captionText="TABLE1. Biometric data of Garra dohjei (n = 11); range and mean ± SD includes holotype. Holotype Paratypes Range Mean S.D. Standard length (mm) 99.6 38.7–115.2 In percents of Standard length Body depth 22.3 15.0–22.8 21.4 1.0 Head length (dorsal) 22.7 15.0–26.6 22.2 0.8 Head length (lateral) 25.6 22.3–29.0 24.4 1.0 Head depth at nape 16.2 13.3–17.9 16.3 0.2" pageId="2" pageNumber="480">Table 1</tableCitation>
. Body elongate; predorsal region cylindrical in cross section, slightly broad laterally, then gently compressing up to dorsal-fin base, ratio of body depth to body width at dorsal-fin origin 0.9–1.4 times, thereafter decreasing towards caudal-fin base. Dorsal profile evenly arched to dorsal-fin origin, thereafter gently sloping towards caudal peduncle. Ventral profile flattened from head to chest, then more or less round up to pelvic-fin origin, and straight from pelvic to caudal-fin base. Head moderately large; moderately depressed, ratio of head width to head depth at nape 0.5–1.2 times with slightly convex interorbital space; ratio of head depth at nape to dorsal head length and to lateral head length 0.6–1.0 and 0.6–0.7 times respectively; ratio of head width at nape to dorsal head length and to lateral head length 0.3–1.1 and 0.3–0.7 times respectively. Snout broadly rounded with transverse lobe covered with 9–12 rounded pits, some fitted with tubercles, demarcated posteriorly by deep transverse groove, 12–13 located on anterior lateral side of nostrils, 10–12 on dorso-anteriormost portion of transverse lobe. Incipient proboscis notch-like in ethmoid region without forming prominent groove anterior to nostrils. Sublachrymal groove shallow. Lateral groove shallow, not connected with rostral cap. Rostral lobe absent.
</paragraph>
<caption id="7DA46644FFD1FFE8FF5657B4E175FAE1" pageId="2" pageNumber="480" startId="2.[151,239,980,1004]" targetType="table">
<paragraph id="296436CCFFD1FFE8FF5657B4E04AFC23" blockId="2.[151,1425,980,2018]" box="[151,1139,980,1005]" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FF5657B4E53AFC23" bold="true" box="[151,259,980,1004]" pageId="2" pageNumber="480">TABLE1.</emphasis>
Biometric data of
<taxonomicName id="EEDB4D4FFFD1FFE8FE0D57B5E661FC23" box="[460,600,980,1004]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="480" phylum="Chordata" rank="species" species="dohjei">
<emphasis id="1BAFEADEFFD1FFE8FE0D57B5E661FC23" box="[460,600,980,1004]" italics="true" pageId="2" pageNumber="480">Garra dohjei</emphasis>
</taxonomicName>
(n = 11); range and mean ± SD includes holotype.
</paragraph>
<paragraph id="296436CCFFD1FFE8FD11579DE7D6FBDA" blockId="2.[151,1425,980,2018]" box="[720,1007,1021,1045]" pageId="2" pageNumber="480">Holotype Paratypes</paragraph>
<paragraph id="296436CCFFD1FFE8FC4C5045E163FBF2" blockId="2.[151,1425,980,2018]" box="[909,1370,1061,1085]" pageId="2" pageNumber="480">Range Mean S.D.</paragraph>
<paragraph id="296436CCFFD1FFE8FF62502DE7C7FBAA" blockId="2.[151,1425,980,2018]" box="[163,1022,1101,1125]" pageId="2" pageNumber="480">Standard length (mm) 99.6 38.7–115.2</paragraph>
<paragraph id="296436CCFFD1FFE8FF625015E5D4FB42" blockId="2.[151,1425,980,2018]" box="[163,493,1141,1165]" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FF625015E5D4FB42" bold="true" box="[163,493,1141,1165]" pageId="2" pageNumber="480">In percents of Standard length</emphasis>
</paragraph>
<paragraph id="296436CCFFD1FFE8FF6250FDE175FB7A" blockId="2.[151,1425,980,2018]" box="[163,1356,1181,1205]" pageId="2" pageNumber="480">Body depth 22.3 15.0–22.8 21.4 1.0</paragraph>
<paragraph id="296436CCFFD1FFE8FF6250A5E175FB11" blockId="2.[151,1425,980,2018]" box="[163,1356,1221,1246]" pageId="2" pageNumber="480">Head length (dorsal) 22.7 15.0–26.6 22.2 0.8</paragraph>
<paragraph id="296436CCFFD1FFE8FF62508EE175FAC9" blockId="2.[151,1425,980,2018]" box="[163,1356,1262,1286]" pageId="2" pageNumber="480">Head length (lateral) 25.6 22.3–29.0 24.4 1.0</paragraph>
<paragraph id="296436CCFFD1FFE8FF625176E175FAE1" blockId="2.[151,1425,980,2018]" box="[163,1356,1302,1326]" pageId="2" pageNumber="480">Head depth at nape 16.2 13.3–17.9 16.3 0.2</paragraph>
</caption>
<paragraph id="296436CCFFD1FFE8FF625123E175FA94" blockId="2.[151,1425,980,2018]" box="[163,1356,1347,1371]" pageId="2" pageNumber="480">Head depth at eye 10.9 9.4–12.9 10.6 0.3</paragraph>
<paragraph id="296436CCFFD1FFE8FF62510BE175FA4C" blockId="2.[151,1425,980,2018]" box="[163,1356,1387,1411]" pageId="2" pageNumber="480">Head width 18.7 15.2–20.1 17.2 1.4</paragraph>
<paragraph id="296436CCFFD1FFE8FF6251F3E175FA64" blockId="2.[151,1425,980,2018]" box="[163,1356,1427,1451]" pageId="2" pageNumber="480">Body width at dorsal-fin origin 19.4 13.7–19.5 18.8 0.6</paragraph>
<paragraph id="296436CCFFD1FFE8FF6251DBE175F875" blockId="2.[151,1425,980,2018]" pageId="2" pageNumber="480">Body width at anal-fin origin 10.1 8.2–11.1 10.4 0.4 Caudal peduncle length 15.7 11.6–16.4 15.0 1.9 Caudal peduncle depth 13.1 10.9–13.8 13.1 0.4 Dorsal-fin length 24.4 19.2–27.4 23.2 1.0 Dorsal-fin base length 17.5 12.4–20.0 16.3 1.1 Anal-fin length 19.4 15.5–20.6 18.3 1.0 Anal-fin base length 8.0 7.1–8.7 7.5 0.5 Pectoral-fin length 22.9 18.7–24.3 21.9 0.9 Pelvic-fin length 20.1 16.2–20.4 19.8 0.7 Predorsal length 49.0 40.2–49.0 47.6 1.2 Prepectoral length 23.2 20.5–25.2 20.9 0.3 Prepelvic length 53.9 48.9–56.2 52.3 1.7 Preanus length 69.3 63.6–75.9 66.8 2.6</paragraph>
<paragraph id="296436CCFFD1FFE9FB9B53AAE62FFD32" blockId="2.[151,1425,980,2018]" lastBlockId="3.[163,1356,420,1451]" lastPageId="3" lastPageNumber="481" pageId="2" pageNumber="480">
<emphasis id="1BAFEADEFFD1FFE8FB9B53AAE1A8F82D" box="[1114,1425,1994,2018]" italics="true" pageId="2" pageNumber="480">......continued on the next page</emphasis>
Snout length 13.8 8.8–14.9 13.0 0.8 Eye diameter 4.6 4.0–7.1 4.3 0.3 Interorbital distance 13.9 9.6–13.9 13.1 0.7 Pulvinus width 8.5 6.2–9.9 8.5 0.5 Pulvinus length 6.4 5.0–7.4 5.8 0.7 Gular disc width 12.6 11.0–14.6 9.5 2.0 Gular disc length 9.8 6.9–11.5 9.5 1.5 Mouth gap width 15.6 12.4–15.6 14.3 1.1
<emphasis id="1BAFEADEFFD0FFE9FF625686E62FFD32" bold="true" box="[163,534,741,766]" pageId="3" pageNumber="481">In percents of head length (dorsal)</emphasis>
</paragraph>
<caption id="7DA46644FFD0FFE9FF6254C2E175FE4C" pageId="3" pageNumber="481" startId="3.[163,247,162,186]" targetType="table">
<paragraph id="296436CCFFD0FFE9FF6254C2E5A8FF75" blockId="3.[163,1370,162,387]" box="[163,401,162,186]" pageId="3" pageNumber="481">
<emphasis id="1BAFEADEFFD0FFE9FF6254C2E529FF75" bold="true" box="[163,272,162,186]" pageId="3" pageNumber="481">TABLE 1.</emphasis>
(Continued)
</paragraph>
<paragraph id="296436CCFFD0FFE9FD1154AAE7D6FF2C" blockId="3.[163,1370,162,387]" box="[720,1007,202,227]" pageId="3" pageNumber="481">Holotype Paratypes</paragraph>
<paragraph id="296436CCFFD0FFE9FC4C5493E175FE4C" blockId="3.[163,1370,162,387]" pageId="3" pageNumber="481">Range Mean S.D. Preanal length 77.4 70.6–79.9 75.1 2.0 Pelvic-anal distance 24.5 22.4–27.1 24.9 1.7 Vent-anal distance 6.2 5.0–6.7 6.0 0.6</paragraph>
</caption>
<paragraph id="296436CCFFD0FFE9FF62576EE620FB8E" blockId="3.[163,1356,420,1451]" pageId="3" pageNumber="481">
Snout length 56.9 52.4–61.0 58.4 2.1 Eye diameter 19.2 18.3–27.6 19.4 0.8 Interorbital distance 58.5 45.9–63.7 59.0 1.9 Pulvinus length 23.6 23.6–34.4 26.1 2.3 Pulvinus width 37.4 30.4–41.0 38.2 1.3 Gular length 44.5 35.7–46.2 44.7 1.6 Gular width 54.2 49.4–59.1 56.0 1.7
<emphasis id="1BAFEADEFFD0FFE9FF625049E620FB8E" bold="true" box="[163,537,1065,1089]" pageId="3" pageNumber="481">In percents of head length (lateral)</emphasis>
</paragraph>
<paragraph id="296436CCFFD0FFE9FF625031E62AFA4D" blockId="3.[163,1356,420,1451]" pageId="3" pageNumber="481">
Snout length 54.0 36.9–57.0 51.7 5.7 Eye diameter 17.9 17.3–24.9 19.7 3.1 Interorbital distance 54.4 40.1–54.4 47.6 5.5 Pulvinus length 24.9 21.2–25.5 23.9 1.7 Pulvinus width 33.4 25.8–37.5 32.5 3.2 Gular length 38.5 29.0–43.8 37.8 4.5 Gular width 49.9 37.1–56.0 49.9 5.2
<emphasis id="1BAFEADEFFD0FFE9FF62510BE62AFA4D" bold="true" box="[163,531,1386,1411]" pageId="3" pageNumber="481">In percents of pelvic-anal distance</emphasis>
</paragraph>
<paragraph id="296436CCFFD0FFE9FF6251F3E175FA64" blockId="3.[163,1356,420,1451]" box="[163,1356,1426,1451]" pageId="3" pageNumber="481">Vent-anal distance 27.5 19.5–27.5 23.8 2.7</paragraph>
<paragraph id="296436CCFFD0FFE9FF0651BAE5FBF895" blockId="3.[151,1437,1498,2026]" pageId="3" pageNumber="481">
Barbels in two pairs; rostral barbel anterolaterally located, shorter than eye diameter (about half of eye diameter); maxillary barbel at corner of mouth, shorter than rostral barbel. Rostral cap well-developed, pendulous and greatly crenulated with wide papillated distal margin, separated from upper jaw by deep groove, laterally continuous with lower lip around corners of mouth (
<figureCitation id="B1E02A49FFD0FFE9FDE05225E65FF990" box="[545,614,1605,1631]" captionStart="FIGURE 2" captionStartId="4.[152,255,1084,1108]" captionTargetBox="[345,1236,381,1061]" captionTargetId="figure-129@4.[345,1242,381,1061]" captionTargetPageId="4" captionText="FIGURE 2. Oromandibular structure of Garra dohjei¸ ZSI FF 10399, holotype, 99.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13757924" httpUri="https://zenodo.org/record/13757924/files/figure.png" pageId="3" pageNumber="481">Fig. 2</figureCitation>
). Upper lip absent; upper jaw fully covered by rostral cap, with thin horny sheath on cutting edge. Lower lip modified to form gular disc. Disc elliptical, shorter than wide, narrower than head width through base of maxillary barbell. Each labellum on anterolateral lobe of lower lip well-developed, papillated with a slight notch on lateral region of lateroposterior flap. Torus fleshy and crescentic skin fold with numerous minute papillae, anterior separated from lower jaw by deep subtoral fold extending along entire length of lower jaw and posteriorly bordered by shallow transverse toral groove with central elliptical pulvinus. Lateral and posterior labrum, surrounding central pulvinus, semi-circular, papillated and distally free; posterior margin reaching vertical of posterior margin of eye.
</paragraph>
<paragraph id="296436CCFFD0FFEEFF065304E161FEAA" blockId="3.[151,1437,1498,2026]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="482" pageId="3" pageNumber="481">Dorsal fin with 2 simple and 8½ branched rays; distal margin concave; origin almost nearly, or at midway between snout tip and caudal-fin base, inserted vertical in advanced from pelvic-fin origin; first branched ray longest, last branched ray not extending vertically to vent. Pectoral fin acuminating with 1 simple and 15 branched rays, reaching midway from posterior pectoral-fin base to pelvic-fin origin when adpressed. Pelvic fin acuminating with 1 simple and 8 branched rays, reaching to midway of anus-anal distance when adpressed, surpassing anus; origin closer to anal-fin origin than to pectoral-fin origin, inserted vertically below to base of third branched dorsal-fin rays; distal margin almost truncate.Anal fin short with 2 simple and 5½ branched rays, first branched ray longest, nearly reaching base of caudal fin when adpressed; origin midway between caudal-fin base and pelvic-fin origin. Anus closer to anal-fin origin than to pelvic-fin origin. Caudal fin forked with 10+9 principal rays; lobes subequal, upper lobe slightly longer than lower, tips of lobes pointed. Caudal peduncle 0.9–1.3 times longer than deep.</paragraph>
<caption id="7DA46644FFD7FFEEFF59505CE06BFB9B" ID-DOI="http://doi.org/10.5281/zenodo.13757924" ID-Zenodo-Dep="13757924" box="[152,1106,1084,1109]" httpUri="https://zenodo.org/record/13757924/files/figure.png" pageId="4" pageNumber="482" startId="4.[152,255,1084,1108]" targetBox="[345,1236,381,1061]" targetPageId="4" targetType="figure">
<paragraph id="296436CCFFD7FFEEFF59505CE06BFB9B" blockId="4.[152,1106,1084,1109]" box="[152,1106,1084,1109]" pageId="4" pageNumber="482">
<emphasis id="1BAFEADEFFD7FFEEFF59505CE521FB9A" bold="true" box="[152,280,1084,1109]" pageId="4" pageNumber="482">FIGURE 2.</emphasis>
Oromandibular structure of
<taxonomicName id="EEDB4D4FFFD7FFEEFDFB505DE72FFB9B" authority="¸ ZSI FF" authorityName="ZSI FF" box="[570,790,1085,1109]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="482" phylum="Chordata" rank="species" species="dohjei">
<emphasis id="1BAFEADEFFD7FFEEFDFB505DE6F1FB9A" box="[570,712,1085,1109]" italics="true" pageId="4" pageNumber="482">Garra dohjei¸</emphasis>
ZSI FF
</taxonomicName>
10399, holotype, 99.6 mm SL.
</paragraph>
</caption>
<paragraph id="296436CCFFD7FFEEFF0650E3E174FAE2" blockId="4.[151,1437,1155,1614]" pageId="4" pageNumber="482">Lateral line complete with 33(2)–34*(9) scales. Transverse scale rows above lateral line scales 4½; between lateral line and pelvic-fin origin 4; between lateral line to anal-fin origin 4½. Circumpeduncular scales 16. Predorsal scales 9 (2) or 10* (9); scales regularly arranged, same size as flank scales. Chest and belly scaled. One long axillary scale at pelvic-fin base, its tip reaching posterior end of pelvic-fin base. Scales between anus and anal-fin origin 4 (11). Dorsal-fin base scales 6 (11). Anal-fin base scales 4 (11), of which last 3–4 connected to anal-fin base.</paragraph>
<paragraph id="296436CCFFD7FFEEFF065157E731FA72" blockId="4.[151,1437,1155,1614]" pageId="4" pageNumber="482">
<emphasis id="1BAFEADEFFD7FFEEFF065157E5F0FA9E" bold="true" box="[199,457,1335,1361]" pageId="4" pageNumber="482">Osteological features.</emphasis>
Total vertebrae 33 (1), abdominal vertebrae 17 (1), caudal vertebrae 16 (1), dorsal fin insertion between 10
<superScript id="DEAE9B84FFD7FFEEFEBF513BE5B2FAA6" attach="left" box="[382,395,1371,1385]" fontSize="6" pageId="4" pageNumber="482">th</superScript>
and 11
<superScript id="DEAE9B84FFD7FFEEFE1B513BE5DEFAA6" attach="left" box="[474,487,1371,1385]" fontSize="6" pageId="4" pageNumber="482">th</superScript>
vertebrae, and anal fin insertion between 21
<superScript id="DEAE9B84FFD7FFEEFC10513BE7E5FAA6" attach="left" box="[977,988,1371,1385]" fontSize="6" pageId="4" pageNumber="482">st</superScript>
and 22
<superScript id="DEAE9B84FFD7FFEEFBEC513BE004FAA6" attach="left" box="[1069,1085,1371,1385]" fontSize="6" pageId="4" pageNumber="482">nd</superScript>
vertebrae. Pelvic fin articulates from lateral rib of 11
<superScript id="DEAE9B84FFD7FFEEFE4C511FE5A3FA42" attach="left" box="[397,410,1407,1421]" fontSize="6" pageId="4" pageNumber="482">th</superScript>
vertebra. First branchial arch with 19 gill rakers viz.,
<quantity id="EE239B29FFD7FFEEFBD7511FE06BFA56" box="[1046,1106,1407,1433]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="4" pageNumber="482" unit="in" value="17.0">17 in</quantity>
hypobranchial and
<quantity id="EE239B29FFD7FFEEFAF8511FE15EFA56" box="[1337,1383,1407,1433]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="482" unit="in" value="2.0">2 in</quantity>
epibranchial. 5
<superScript id="DEAE9B84FFD7FFEEFEDA51C3E511FA7E" attach="left" box="[283,296,1443,1457]" fontSize="6" pageId="4" pageNumber="482">th</superScript>
ceratobranchial with 9 teeth in three rows.
</paragraph>
<paragraph id="296436CCFFD7FFEEFF0651A7E72DF982" blockId="4.[151,1437,1155,1614]" pageId="4" pageNumber="482">
<emphasis id="1BAFEADEFFD7FFEEFF0651A7E58BFA2E" bold="true" box="[199,434,1479,1505]" pageId="4" pageNumber="482">Sexual dimorphism</emphasis>
. Males with slender body, smaller than females with prominent tubercles in transverse lobe (
<figureCitation id="B1E02A49FFD7FFEEFF16518BE522F9CA" box="[215,283,1515,1541]" captionStart="FIGURE 3" captionStartId="4.[152,255,1970,1994]" captionTargetBox="[340,1247,1637,1947]" captionTargetId="figure-379@4.[340,1247,1637,1947]" captionTargetPageId="4" captionText="FIGURE 3. Lateral view of head showing sexual dimorphic characters of Garra dohjei. A. ZSI FF 10399, holotype, 99.6 mm SL with low tubercles; B. ADBU-MF/5090/8, paratype, 87.8 mm SL, male with prominent tubercles." figureDoi="http://doi.org/10.5281/zenodo.13757926" httpUri="https://zenodo.org/record/13757926/files/figure.png" pageId="4" pageNumber="482">Fig. 3</figureCitation>
). Females with wider body than males at dorsal-fin origin. Males distinguished from females in having deeper body (18.2–23.5 % SL vs. 14.7–19.8), narrower body at anal-fin origin (8.8–9.7 % SL vs. 10.0–12.6), and longer preanus distance (69.0–75.4 % SL vs. 63.8–69.9).
</paragraph>
<caption id="7DA46644FFD7FFEEFF5953D2E0A1F821" ID-DOI="http://doi.org/10.5281/zenodo.13757926" ID-Zenodo-Dep="13757926" httpUri="https://zenodo.org/record/13757926/files/figure.png" pageId="4" pageNumber="482" startId="4.[152,255,1970,1994]" targetBox="[340,1247,1637,1947]" targetPageId="4" targetType="figure">
<paragraph id="296436CCFFD7FFEEFF5953D2E0A1F821" blockId="4.[151,1436,1970,2031]" pageId="4" pageNumber="482">
<emphasis id="1BAFEADEFFD7FFEEFF5953D2E521F804" bold="true" box="[152,280,1970,1995]" pageId="4" pageNumber="482">FIGURE 3.</emphasis>
Lateral view of head showing sexual dimorphic characters of
<emphasis id="1BAFEADEFFD7FFEEFC5053D3E024F805" box="[913,1053,1970,1994]" italics="true" pageId="4" pageNumber="482">
<taxonomicName id="EEDB4D4FFFD7FFEEFC5053D3E023F805" box="[913,1050,1970,1994]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="482" phylum="Chordata" rank="species" species="dohjei">Garra dohjei</taxonomicName>
.
</emphasis>
<emphasis id="1BAFEADEFFD7FFEEFBE253D2E002F805" bold="true" box="[1059,1083,1970,1994]" pageId="4" pageNumber="482">A.</emphasis>
ZSI FF 10399, holotype, 99.6 mm SL with low tubercles;
<emphasis id="1BAFEADEFFD7FFEEFE4253B7E5A3F820" bold="true" box="[387,410,2007,2031]" pageId="4" pageNumber="482">B.</emphasis>
ADBU-MF/5090/8, paratype, 87.8 mm SL, male with prominent tubercles.
</paragraph>
</caption>
<caption id="7DA46644FFD6FFEFFF5955D8E7DBFE1F" box="[152,994,440,465]" pageId="5" pageNumber="483" startId="5.[152,255,440,464]" targetType="figure">
<paragraph id="296436CCFFD6FFEFFF5955D8E7DBFE1F" blockId="5.[152,994,440,465]" box="[152,994,440,465]" pageId="5" pageNumber="483">
<emphasis id="1BAFEADEFFD6FFEFFF5955D8E521FE1E" bold="true" box="[152,280,440,465]" pageId="5" pageNumber="483">FIGURE 4.</emphasis>
<emphasis id="1BAFEADEFFD6FFEFFEDE55D9E595FE1E" box="[287,428,441,465]" italics="true" pageId="5" pageNumber="483">
<taxonomicName id="EEDB4D4FFFD6FFEFFEDE55D9E59FFE1E" box="[287,422,441,465]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="483" phylum="Chordata" rank="species" species="dohjei">Garra dohjei</taxonomicName>
¸
</emphasis>
live image of an uncatalogued individual, 90.4 mm SL.
</paragraph>
</caption>
<paragraph id="296436CCFFD6FFEFFF06559FE773FD66" blockId="5.[151,1437,511,681]" pageId="5" pageNumber="483">
<emphasis id="1BAFEADEFFD6FFEFFF06559FE5ABFDD6" bold="true" box="[199,402,511,537]" pageId="5" pageNumber="483">Notes on biology.</emphasis>
One dissected gravid female (ADBU-MF/5090/
<quantity id="EE239B29FFD6FFEFFC74559FE005FDD6" box="[949,1084,511,538]" metricMagnitude="0" metricUnit="m" metricValue="3.1151999999999997" pageId="5" pageNumber="483" unit="mm" value="3115.2">3, 115.2 mm</quantity>
SL,
<quantity id="EE239B29FFD6FFEFFBB0559FE08FFDD6" box="[1137,1206,511,537]" metricMagnitude="-2" metricUnit="kg" metricValue="2.4699999999999998" pageId="5" pageNumber="483" unit="g" value="24.7">24.7 g</quantity>
body wt.) contained about 2533 ova (weight of ovaries
<quantity id="EE239B29FFD6FFEFFDF05643E6B4FDF2" box="[561,653,547,573]" metricMagnitude="-4" metricUnit="kg" metricValue="9.74" pageId="5" pageNumber="483" unit="mg" value="974.0">974 mg</quantity>
). Air bladder physostomous, with two chambers (total length of air bladder 10.8% SL); anterior lobe (55.6% of total length of air bladder) longer than wide (ratio of width to length of anterior chamber 0.6 times); posterior lobe of air bladder 44.3% of total length of air bladder) with a narrow tapering posterior end, ratio of its maximum width to length 0.3 times.
</paragraph>
<caption id="7DA46644FFD6FFEFFF595344E0ECF8F3" ID-DOI="http://doi.org/10.5281/zenodo.13757928" ID-Zenodo-Dep="13757928" box="[152,1237,1828,1852]" httpUri="https://zenodo.org/record/13757928/files/figure.png" pageId="5" pageNumber="483" startId="5.[152,255,1828,1852]" targetBox="[151,1430,705,1801]" targetPageId="5" targetType="figure">
<paragraph id="296436CCFFD6FFEFFF595344E0ECF8F3" blockId="5.[152,1237,1828,1852]" box="[152,1237,1828,1852]" pageId="5" pageNumber="483">
<emphasis id="1BAFEADEFFD6FFEFFF595344E521F8F3" bold="true" box="[152,280,1828,1852]" pageId="5" pageNumber="483">FIGURE 5.</emphasis>
Map depicting the collection locality of
<taxonomicName id="EEDB4D4FFFD6FFEFFD745345E703F8F3" box="[693,826,1828,1852]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="483" phylum="Chordata" rank="species" species="dohjei">
<emphasis id="1BAFEADEFFD6FFEFFD745345E703F8F3" box="[693,826,1828,1852]" italics="true" pageId="5" pageNumber="483">Garra dohjei</emphasis>
</taxonomicName>
. Solid star represents the collection site.
</paragraph>
</caption>
<paragraph id="296436CCFFD6FFECFF06530AE62AFF36" blockId="5.[151,1436,1898,2033]" lastBlockId="6.[151,1437,151,609]" lastPageId="6" lastPageNumber="484" pageId="5" pageNumber="483">
<emphasis id="1BAFEADEFFD6FFEFFF06530AE561F84B" bold="true" box="[199,344,1898,1924]" pageId="5" pageNumber="483">Colouration</emphasis>
. In fresh specimens (
<figureCitation id="B1E02A49FFD6FFEFFD8C530AE6ABF84A" box="[589,658,1898,1925]" captionStart="FIGURE 4" captionStartId="5.[152,255,440,464]" captionText="FIGURE 4. Garra dohjei¸ live image of an uncatalogued individual, 90.4 mm SL." pageId="5" pageNumber="483">Fig. 4</figureCitation>
), head and body dark olivaceous, both dorsally and laterally, orange to pinkish cream on ventral surface of head and thorax. Background colour of each scale on flanks faint dark olivaceous. Each scale with darker margin on free posterior end. A black spot immediately anterior to upper angle of gill opening. Five or six narrow dark black stripes on caudal peduncle region. Eye pupil black with light grey ring. Interradial membrane of anal and caudal fin orange. Lower lobe of caudal-fin rays from caudal fork tinged with black, rays above caudal fork dark olivaceous, distal margin of caudal-fin rays fainted. A faint black blotch at caudal-fin base in live specimens.
</paragraph>
<paragraph id="296436CCFFD5FFECFF065563E607FEAA" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="484">Colour in formalin. In formalin preserved specimens, head, dorsum and side blackish or dark grey. Mouth, chest and abdomen pale yellowish. Distal margin of fins greyish white. Dorsal, caudal, pelvic and pectoral-fins rays dark grey, interradial membrane light grey.</paragraph>
</subSubSection>
<subSubSection id="61C16547FFD5FFECFF06550EE091FDF2" pageId="6" pageNumber="484" type="distribution">
<paragraph id="296436CCFFD5FFECFF06550EE091FDF2" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="484">
<emphasis id="1BAFEADEFFD5FFECFF06550EE5C8FE47" bold="true" box="[199,497,366,393]" pageId="6" pageNumber="484">Distribution and habitat</emphasis>
. This species is known from the Ñiangdai, a small tributary of the Brahmaputra River along its southern basin (
<figureCitation id="B1E02A49FFD5FFECFE3055F3E60AFE62" box="[497,563,402,429]" captionStart="FIGURE 5" captionStartId="5.[152,255,1828,1852]" captionTargetBox="[151,1430,705,1801]" captionTargetId="figure-148@5.[151,1436,705,1805]" captionTargetPageId="5" captionText="FIGURE 5. Map depicting the collection locality of Garra dohjei. Solid star represents the collection site." figureDoi="http://doi.org/10.5281/zenodo.13757928" httpUri="https://zenodo.org/record/13757928/files/figure.png" pageId="6" pageNumber="484">Fig. 5</figureCitation>
) at Ñiangdai village under Ri-bhoi district about
<quantity id="EE239B29FFD5FFECFB9655F2E08BFE62" box="[1111,1202,402,429]" metricMagnitude="4" metricUnit="m" metricValue="2.57" pageId="6" pageNumber="484" unit="km" value="25.7">25.7 km</quantity>
away from Shillong,
<collectingRegion id="EB1FF82EFFD5FFECFF5655D7E52FFE1E" box="[151,278,439,465]" country="India" name="Meghalaya" pageId="6" pageNumber="484">Meghalaya</collectingRegion>
,
<collectingCountry id="51CC765CFFD5FFECFEE055D7E567FE1E" box="[289,350,439,465]" name="India" pageId="6" pageNumber="484">India</collectingCountry>
. The catchment area at the collection site is covered with forests and sparsely arranged cultivated land on both sides. The substrate at the collection site comprised boulders, cobbles, pebbles, gravels, sand, silt and clay (
<figureCitation id="B1E02A49FFD5FFECFF18559FE51BFDD6" box="[217,290,511,537]" captionStart="FIGURE 6" captionStartId="6.[152,247,1336,1360]" captionTargetBox="[340,1247,633,1313]" captionTargetId="figure-235@6.[340,1247,633,1313]" captionTargetPageId="6" captionText="FIGURE 6. Type locality of Garra dohjei, Ñiangdai River near Ñiangdai village, Meghalaya, India. The direction of flow is northwards" figureDoi="http://doi.org/10.5281/zenodo.13757930" httpUri="https://zenodo.org/record/13757930/files/figure.png" pageId="6" pageNumber="484">Fig. 6</figureCitation>
).
<emphasis id="1BAFEADEFFD5FFECFEFA559FE157FDD6" box="[315,1390,511,537]" italics="true" pageId="6" pageNumber="484">
<taxonomicName id="EEDB4D4FFFD5FFECFEFA559FE67AFDD7" baseAuthorityName="McClelland" baseAuthorityYear="1839" box="[315,579,511,536]" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="aequipinnatus">Devario aequipinnatus</taxonomicName>
,
<taxonomicName id="EEDB4D4FFFD5FFECFD93559FE79FFDD6" baseAuthorityName="McClelland" baseAuthorityYear="1839" box="[594,934,511,537]" family="Cyprinidae" genus="Neolissochilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="hexagonolepis">Neolissochilus hexagonolepis</taxonomicName>
,
<taxonomicName id="EEDB4D4FFFD5FFECFC75559FE04FFDD6" baseAuthorityName="Yazdani &amp; Talukdar" baseAuthorityYear="1975" box="[948,1142,511,537]" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="shalynius">Pethia shalynius</taxonomicName>
,
<taxonomicName id="EEDB4D4FFFD5FFECFB45559FE157FDD6" baseAuthorityName="McClelland" baseAuthorityYear="1842" box="[1156,1390,511,537]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="lissorhynchus">Garra lissorhynchus</taxonomicName>
</emphasis>
,
<taxonomicName id="EEDB4D4FFFD5FFECFA40559FE4DBFDF3" baseAuthorityName="McClelland" baseAuthorityYear="1838" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="1BAFEADEFFD5FFECFA40559FE4DBFDF3" italics="true" pageId="6" pageNumber="484">G. nasuta</emphasis>
</taxonomicName>
,
<taxonomicName id="EEDB4D4FFFD5FFECFF315643E5B9FDF3" box="[240,384,547,573]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="1BAFEADEFFD5FFECFF315643E563FDF2" box="[240,346,547,573]" italics="true" pageId="6" pageNumber="484">Schistura</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="EEDB4D4FFFD5FFECFE4E5643E673FDF3" box="[399,586,547,573]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="1BAFEADEFFD5FFECFE4E5643E61DFDF2" box="[399,548,547,573]" italics="true" pageId="6" pageNumber="484">Glyptothorax</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="EEDB4D4FFFD5FFECFD495643E731FDF3" box="[648,776,547,573]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="484" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="1BAFEADEFFD5FFECFD495643E6D8FDF2" box="[648,737,547,573]" italics="true" pageId="6" pageNumber="484">Channa</emphasis>
sp.
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were collected with the new species.
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<emphasis id="1BAFEADEFFD5FFECFF065627E571FDAE" bold="true" box="[199,328,583,609]" pageId="6" pageNumber="484">Etymology</emphasis>
. The specific epithet is derived from ‘
<emphasis id="1BAFEADEFFD5FFECFD345627E771FDAF" box="[757,840,583,609]" italics="true" pageId="6" pageNumber="484">Doh jei</emphasis>
’, the local name of
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<emphasis id="1BAFEADEFFD5FFECFBE95627E057FDAE" box="[1064,1134,583,609]" italics="true" pageId="6" pageNumber="484">Garra</emphasis>
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in Khasi dialect.
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354
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@ -0,0 +1,354 @@
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<figureCitation id="DA872A7EFFC1DD6FFED96232FEA116AF" box="[306,377,1395,1421]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="19.[151,250,1205,1229]" captionStartId-1="20.[151,250,1926,1950]" captionStartId-2="21.[151,250,1926,1950]" captionTargetBox-0="[151,1436,181,1181]" captionTargetBox-1="[151,1436,181,1902]" captionTargetBox-2="[151,1436,181,1902]" captionTargetId-0="figure-17@19.[151,1436,181,1181]" captionTargetId-1="figure-19@20.[151,1436,181,1902]" captionTargetId-2="figure-17@21.[151,1436,181,1902]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 14. Anthracites zebra Hebard, 1922 male titillators in apical (A), dorso-proximal (B) and ventro-distal (C) views. Scale bars: 1 mm." captionText-1="FIGURE 15. Anthracites zebra Hebard, 1922 male (A) and female (B) in their natural habitat at Lake Linao." captionText-2="FIGURE 16. Anthracites zebra Hebard, 1922 females feeding on the flowers of a Melastoma plant at Lake Linao." figureDoi-0="http://doi.org/10.5281/zenodo.13757727" figureDoi-1="http://doi.org/10.5281/zenodo.13757729" figureDoi-2="http://doi.org/10.5281/zenodo.13757733" httpUri-0="https://zenodo.org/record/13757727/files/figure.png" httpUri-1="https://zenodo.org/record/13757729/files/figure.png" httpUri-2="https://zenodo.org/record/13757733/files/figure.png" pageId="18" pageNumber="331">14–16</figureCitation>
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•
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[6°59'N, 125°16'E]; coll.
<collectorName id="EF49532DFFC1DD6FFF7C6163FEC7151E" box="[151,287,1570,1596]" pageId="18" pageNumber="331">E.A. Merna</collectorName>
; (
<collectionCode id="24ADAE3EFFC1DD6FFEDA6163FEA7151E" box="[305,383,1570,1596]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="18" pageNumber="331">ANSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="420336FBFFC1DD6FFF2C6107FF27158A" blockId="18.[151,1437,1533,1992]" pageId="18" pageNumber="331">
<emphasis id="70C8EAE9FFC1DD6FFF2C6107FC1C157D" bold="true" box="[199,964,1605,1632]" pageId="18" pageNumber="331">
Material examined (images and description from
<bibRefCitation id="262D4B0AFFC1DD6FFCFA6107FC64157D" author="Ingrisch, S." box="[785,956,1605,1632]" pageId="18" pageNumber="331" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
)
</emphasis>
•
<materialsCitation id="F2D43CA6FFC1DD6FFC316107FF23158A" collectingDate="1958-07-14" collectingDateMax="1958-07-18" collectingDateMin="1958-07-14" collectionCode="BPBM" collectorName="H. E. Milliron" country="Philippines" elevation="1260" location="Clarin River" municipality="Mindanao" pageId="18" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFC316107FC261542" box="[986,1022,1606,1632]" count="1" pageId="18" pageNumber="331" type="male">1♂</specimenCount>
;
<collectingCountry id="3AAB766BFFC1DD6FFBE06107FB6B1542" box="[1035,1203,1606,1632]" name="Philippines" pageId="18" pageNumber="331">PHILIPPINES</collectingCountry>
,
<collectingMunicipality id="A267AC81FFC1DD6FFB566107FAE91542" box="[1213,1329,1606,1632]" pageId="18" pageNumber="331">Mindanao</collectingMunicipality>
,
<collectingRegion id="8078F819FFC1DD6FFAD76107FECB15A6" country="Philippines" name="Misamis Occidental" pageId="18" pageNumber="331">Misamis Occidental</collectingRegion>
, tributary of
<location id="47636020FFC1DD6FFE59612BFD9B15A6" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFE59612BFD9B15A6" box="[434,579,1642,1668]" country="Philippines" municipality="Mindanao" name="Clarin River" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Clarin River</location>
, elev.
<quantity id="85449B1EFFC1DD6FFD79612BFD3315A6" box="[658,747,1642,1668]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="18" pageNumber="331" unit="m" value="1260.0">
<elevation id="C991D1C8FFC1DD6FFD79612BFD3315A6" box="[658,747,1642,1668]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="18" pageNumber="331" unit="m" value="1260.0">1260 m</elevation>
</quantity>
[8°12'N, 123°42'E];
<date id="3602103BFFC1DD6FFC0E612BFB7415A7" box="[997,1196,1642,1669]" pageId="18" pageNumber="331" value="1958-07-14" valueMax="1958-07-18" valueMin="1958-07-14">
<collectingDate id="2646E9D3FFC1DD6FFC0E612BFB7415A7" box="[997,1196,1642,1669]" pageId="18" pageNumber="331" value="1958-07-14" valueMax="1958-07-18" valueMin="1958-07-14">14–18 July 1958</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFC1DD6FFB13612BFA4015A6" box="[1272,1432,1642,1668]" pageId="18" pageNumber="331">H.E. Milliron</collectorName>
; (
<collectionCode id="24ADAE3EFFC1DD6FFF7461CFFF2B158A" box="[159,243,1678,1704]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="331" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="420336FBFFC1DD6FFF2C61F3FEB314EA" blockId="18.[151,1437,1533,1992]" pageId="18" pageNumber="331">
<emphasis id="70C8EAE9FFC1DD6FFF2C61F3FE3315EE" bold="true" box="[199,491,1714,1740]" pageId="18" pageNumber="331">New material examined.</emphasis>
<materialsCitation id="F2D43CA6FFC1DD6FFE1961F3FBBF15EE" box="[498,1127,1714,1740]" country="Philippines" location="Mount Malindang" pageId="18" pageNumber="331" specimenCount="1" stateProvince="Misamis Occidental">
<collectingCountry id="3AAB766BFFC1DD6FFE1961F3FD4215EE" box="[498,666,1714,1740]" name="Philippines" pageId="18" pageNumber="331">PHILIPPINES</collectingCountry>
,
<collectingRegion id="8078F819FFC1DD6FFD4D61F3FC5415EE" box="[678,908,1714,1740]" country="Philippines" name="Misamis Occidental" pageId="18" pageNumber="331">Misamis Occidental</collectingRegion>
:
<location id="47636020FFC1DD6FFC7D61F3FBBF15EE" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFC7D61F3FBBF15EE" box="[918,1127,1714,1740]" country="Philippines" name="Mount Malindang" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Mount Malindang</location>
</materialsCitation>
: •
<materialsCitation id="F2D43CA6FFC1DD6FFB6861F3FA8B15D2" collectingDate="2024-01-07" collectionCode="ZRC" collectorName="Pandan &amp; M. K. Tan" country="Philippines" latitude="8.2938" location="Linao Lake" longLatPrecision="1" longitude="123.61192" pageId="18" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFB6861F3FB7F15EE" box="[1155,1191,1714,1740]" count="1" pageId="18" pageNumber="331" type="male">1♂</specimenCount>
; along coast of
<location id="47636020FFC1DD6FFAB061F3FF0B15D2" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFAB061F3FF0B15D2" country="Philippines" latitude="8.2938" longLatPrecision="1" longitude="123.61192" name="Linao Lake" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Linao Lake</location>
;
<geoCoordinate id="2788503CFFC1DD6FFF0A6197FE8C15D2" box="[225,340,1750,1776]" degrees="8.29380" direction="north" orientation="latitude" pageId="18" pageNumber="331" precision="1" value="8.2938">N8.29380</geoCoordinate>
,
<geoCoordinate id="2788503CFFC1DD6FFE896197FE3515D2" box="[354,493,1750,1776]" degrees="123.61192" direction="east" orientation="longitude" pageId="18" pageNumber="331" precision="1" value="123.61192">E123.61192</geoCoordinate>
;
<date id="3602103BFFC1DD6FFE106197FD6815D2" box="[507,688,1750,1776]" pageId="18" pageNumber="331" value="2024-01-07">
<collectingDate id="2646E9D3FFC1DD6FFE106197FD6815D2" box="[507,688,1750,1776]" pageId="18" pageNumber="331" value="2024-01-07">7 January 2024</collectingDate>
</date>
, 20h26; on
<collectorName id="EF49532DFFC1DD6FFCD76197FC5615D2" box="[828,910,1750,1776]" pageId="18" pageNumber="331">Pandan</collectorName>
leaf; coll.
<collectorName id="EF49532DFFC1DD6FFBFA6197FB5A15D2" box="[1041,1154,1750,1776]" pageId="18" pageNumber="331">M.K. Tan</collectorName>
; MIN.24.1 (
<collectionCode id="24ADAE3EFFC1DD6FFAF96197FA9415D2" box="[1298,1356,1750,1776]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="331">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFC1DD6FFA9A6197FF39141A" collectingDate="2024-01-07" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" latitude="8.2938" location="Linao Lake" longLatPrecision="1" longitude="123.61192" pageId="18" pageNumber="331" specimenCount="1" specimenCount-female="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFA9A6197FA4E15D2" box="[1393,1430,1750,1776]" count="1" pageId="18" pageNumber="331" type="female">1♀</specimenCount>
; along coast of
<location id="47636020FFC1DD6FFEAF61BBFE131436" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFEAF61BBFE131436" box="[324,459,1786,1812]" country="Philippines" latitude="8.2938" longLatPrecision="1" longitude="123.61192" name="Linao Lake" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Linao Lake</location>
;
<geoCoordinate id="2788503CFFC1DD6FFE3361BBFD931436" box="[472,587,1786,1812]" degrees="8.29380" direction="north" orientation="latitude" pageId="18" pageNumber="331" precision="1" value="8.2938">N8.29380</geoCoordinate>
,
<geoCoordinate id="2788503CFFC1DD6FFDB361BBFD3C1436" box="[600,740,1786,1812]" degrees="123.61192" direction="east" orientation="longitude" pageId="18" pageNumber="331" precision="1" value="123.61192">E123.61192</geoCoordinate>
;
<date id="3602103BFFC1DD6FFD1A61BBFC7D1436" box="[753,933,1786,1812]" pageId="18" pageNumber="331" value="2024-01-07">
<collectingDate id="2646E9D3FFC1DD6FFD1A61BBFC7D1436" box="[753,933,1786,1812]" pageId="18" pageNumber="331" value="2024-01-07">7 January 2024</collectingDate>
</date>
, 20h26; on herb; coll.
<collectorName id="EF49532DFFC1DD6FFB4561BBFAF81436" box="[1198,1312,1786,1812]" pageId="18" pageNumber="331">M.K. Tan</collectorName>
; MIN.24.2 (
<collectionCode id="24ADAE3EFFC1DD6FFF74605FFF01141A" box="[159,217,1822,1848]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="331">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFC1DD6FFF1C605FFDD4147E" collectingDate="2024-01-07" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" latitude="8.29562" location="Linao Lake" longLatPrecision="1" longitude="123.61205" pageId="18" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFF1C605FFEC4141A" box="[247,284,1822,1848]" count="1" pageId="18" pageNumber="331" type="male">1♂</specimenCount>
; along coast of
<location id="47636020FFC1DD6FFE27605FFD88141A" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFE27605FFD88141A" box="[460,592,1822,1848]" country="Philippines" latitude="8.29562" longLatPrecision="1" longitude="123.61205" name="Linao Lake" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Linao Lake</location>
;
<geoCoordinate id="2788503CFFC1DD6FFDB1605FFD15141B" box="[602,717,1822,1849]" degrees="8.29562" direction="north" orientation="latitude" pageId="18" pageNumber="331" precision="1" value="8.29562">N8.29562</geoCoordinate>
,
<geoCoordinate id="2788503CFFC1DD6FFD3C605FFCBB141B" box="[727,867,1822,1849]" degrees="123.61205" direction="east" orientation="longitude" pageId="18" pageNumber="331" precision="1" value="123.61205">E123.61205</geoCoordinate>
;
<date id="3602103BFFC1DD6FFC85605FFBC3141A" box="[878,1051,1822,1848]" pageId="18" pageNumber="331" value="2024-01-07">
<collectingDate id="2646E9D3FFC1DD6FFC85605FFBC3141A" box="[878,1051,1822,1848]" pageId="18" pageNumber="331" value="2024-01-07">7 January 2024</collectingDate>
</date>
, 20h42; on the leaf of a small tree; coll.
<collectorName id="EF49532DFFC1DD6FFF3B6003FE98147E" box="[208,320,1858,1884]" pageId="18" pageNumber="331">M.K. Tan</collectorName>
; MIN.24.3 (
<collectionCode id="24ADAE3EFFC1DD6FFE216003FDDC147E" box="[458,516,1858,1884]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="331">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFC1DD6FFDCD6003FB9514A2" collectingDate="2024-01-07" collectionCode="ZRC" collectorName="K. C. Grumo &amp; A. A. Gono" country="Philippines" elevation="6" latitude="8.30603" location="Linao Lake" longLatPrecision="1" longitude="123.61685" pageId="18" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFDCD6003FD92147E" box="[550,586,1858,1884]" count="1" pageId="18" pageNumber="331" type="male">1♂</specimenCount>
; along coast of
<location id="47636020FFC1DD6FFCEB6003FC5D147E" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFCEB6003FC5D147E" box="[768,901,1858,1884]" country="Philippines" latitude="8.30603" longLatPrecision="1" longitude="123.61685" name="Linao Lake" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Linao Lake</location>
;
<geoCoordinate id="2788503CFFC1DD6FFC7A6003FBDA147E" box="[913,1026,1858,1884]" degrees="8.30603" direction="north" orientation="latitude" pageId="18" pageNumber="331" precision="1" value="8.30603">N8.30603</geoCoordinate>
,
<geoCoordinate id="2788503CFFC1DD6FFBE46003FB42147F" box="[1039,1178,1858,1885]" degrees="123.61685" direction="east" orientation="longitude" pageId="18" pageNumber="331" precision="1" value="123.61685">E123.61685</geoCoordinate>
, 1555.0±
<quantity id="85449B1EFFC1DD6FFAE86003FA93147E" box="[1283,1355,1858,1885]" metricMagnitude="0" metricUnit="m" metricValue="6.3" pageId="18" pageNumber="331" unit="m" value="6.3">
<elevation id="C991D1C8FFC1DD6FFAE86003FA93147E" box="[1283,1355,1858,1885]" metricMagnitude="0" metricUnit="m" metricValue="6.3" pageId="18" pageNumber="331" unit="m" value="6.3">6.3 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFC1DD6FFA656003FEE814A2" pageId="18" pageNumber="331" value="2024-01-07">
<collectingDate id="2646E9D3FFC1DD6FFA656003FEE814A2" pageId="18" pageNumber="331" value="2024-01-07">7 January 2024</collectingDate>
</date>
, 21h26; on fern leaf; coll.
<collectorName id="EF49532DFFC1DD6FFDBC6027FD3A14A2" box="[599,738,1894,1920]" pageId="18" pageNumber="331">K.C. Grumo</collectorName>
&amp;
<collectorName id="EF49532DFFC1DD6FFCEF6027FC5C14A2" box="[772,900,1894,1920]" pageId="18" pageNumber="331">A.A. Gono</collectorName>
; MIN.24.5 (
<collectionCode id="24ADAE3EFFC1DD6FFBE76027FB9E14A2" box="[1036,1094,1894,1920]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="331">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFC1DD6FFB8E6027FEBF14EA" collectingDate="2024-01-08" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" latitude="8.30909" location="Linao Lake" longLatPrecision="1" longitude="123.6125" pageId="18" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Occidental">
<specimenCount id="54BAFD72FFC1DD6FFB8E6027FB5114A2" box="[1125,1161,1894,1920]" count="1" pageId="18" pageNumber="331" type="male">1♂</specimenCount>
; forest near
<location id="47636020FFC1DD6FFAFF6027FA4014A2" LSID="urn:lsid:plazi:treatment:CA1587EDFFC1DD6EFF7C620EFA4815CB:47636020FFC1DD6FFAFF6027FA4014A2" box="[1300,1432,1894,1920]" country="Philippines" latitude="8.30909" longLatPrecision="1" longitude="123.6125" name="Linao Lake" pageId="18" pageNumber="331" stateProvince="Misamis Occidental">Linao Lake</location>
;
<geoCoordinate id="2788503CFFC1DD6FFF7C60CBFED11486" box="[151,265,1930,1956]" degrees="8.30909" direction="north" orientation="latitude" pageId="18" pageNumber="331" precision="1" value="8.30909">N8.30909</geoCoordinate>
,
<geoCoordinate id="2788503CFFC1DD6FFEF360CBFE7B1487" box="[280,419,1930,1957]" degrees="123.61250" direction="east" orientation="longitude" pageId="18" pageNumber="331" precision="1" value="123.6125">E123.61250</geoCoordinate>
, 1651.0±11.7;
<date id="3602103BFFC1DD6FFDBB60CBFCDD1486" box="[592,773,1930,1956]" pageId="18" pageNumber="331" value="2024-01-08">
<collectingDate id="2646E9D3FFC1DD6FFDBB60CBFCDD1486" box="[592,773,1930,1956]" pageId="18" pageNumber="331" value="2024-01-08">8 January 2024</collectingDate>
</date>
, before 19h29; on herb near the ground; coll.
<collectorName id="EF49532DFFC1DD6FFACD60CBFA401486" box="[1318,1432,1930,1956]" pageId="18" pageNumber="331">M.K. Tan</collectorName>
; MIN.24.13 (
<collectionCode id="24ADAE3EFFC1DD6FFEC860EFFE8714EA" box="[291,351,1966,1992]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="331">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="16C36673FFC0DD6EFF7C63F4FE9617D3" ID-DOI="http://doi.org/10.5281/zenodo.13757727" ID-Zenodo-Dep="13757727" httpUri="https://zenodo.org/record/13757727/files/figure.png" pageId="19" pageNumber="332" startId="19.[151,250,1205,1229]" targetBox="[151,1436,181,1181]" targetPageId="19" targetType="figure">
<paragraph id="420336FBFFC0DD6EFF7C63F4FE9617D3" blockId="19.[151,1437,1205,1265]" pageId="19" pageNumber="332">
<emphasis id="70C8EAE9FFC0DD6EFF7C63F4FEFA17EF" bold="true" box="[151,290,1205,1229]" pageId="19" pageNumber="332">FIGURE 14.</emphasis>
<taxonomicName id="85BC4D78FFC0DD6EFEC163F4FDAC17EF" authority="Hebard, 1922" authorityName="Hebard" authorityYear="1922" box="[298,628,1205,1229]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="332" phylum="Arthropoda" rank="species" species="zebra">
<emphasis id="70C8EAE9FFC0DD6EFEC163F4FE0717EF" box="[298,479,1205,1229]" italics="true" pageId="19" pageNumber="332">Anthracites zebra</emphasis>
<bibRefCitation id="262D4B0AFFC0DD6EFE0363F4FDAC17EF" author="Hebard, M." box="[488,628,1205,1229]" pageId="19" pageNumber="332" pagination="121 - 299" refId="ref9228" refString="Hebard, M. (1922) Studies in Malayan, Melanesian and Australian Tettigoniidae (Orthoptera). Proceedings of the Academy of Natural Sciences, Philadelphia, 74, 121 - 299, pls. 11 - 22." type="journal article" year="1922">Hebard, 1922</bibRefCitation>
</taxonomicName>
male titillators in apical (A), dorso-proximal (B) and ventro-distal (C) views. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="0AA66570FFC0DD6EFF2C6205FC88157B" pageId="19" pageNumber="332" type="discussion">
<paragraph id="420336FBFFC0DD6EFF2C6205FE981687" blockId="19.[151,1436,1347,1770]" pageId="19" pageNumber="332">
<emphasis id="70C8EAE9FFC0DD6EFF2C6205FEE3167C" bold="true" box="[199,315,1348,1374]" pageId="19" pageNumber="332">Remarks.</emphasis>
In
<collectingRegion id="8078F819FFC0DD6EFE8B6202FD9B167F" box="[352,579,1347,1373]" country="Philippines" name="Misamis Occidental" pageId="19" pageNumber="332">Misamis Occidental</collectingRegion>
, this species has been reported in the tributaries of Clarin River to the east of Mount Malindang. New material was collected from Lake Linao to the west side of Mount Malindang from recent field sampling.
</paragraph>
<paragraph id="420336FBFFC0DD6EFF2C62EEFC88157B" blockId="19.[151,1436,1347,1770]" pageId="19" pageNumber="332">
There are minor differences in the colour patterns, particularly the dorsum of its femur is not black in the specimens from Mount Malindang, in contrast to the species from Mount Apo and Clarin River. Colour variations were also noted by
<bibRefCitation id="262D4B0AFFC0DD6EFE9162B6FDF11530" author="Ingrisch, S." box="[378,553,1527,1554]" pageId="19" pageNumber="332" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch (2015)</bibRefCitation>
between the specimen from Mount Apo and from Clarin River. However, as we do not observe clear-cut differences in the male abdominal apex and titillators (
<figureCitation id="DA872A7EFFC0DD6EFBDC615AFB511514" box="[1079,1161,1563,1590]" captionStart="FIGURE 14" captionStartId="19.[151,250,1205,1229]" captionTargetBox="[151,1436,181,1181]" captionTargetId="figure-17@19.[151,1436,181,1181]" captionTargetPageId="19" captionText="FIGURE 14. Anthracites zebra Hebard, 1922 male titillators in apical (A), dorso-proximal (B) and ventro-distal (C) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13757727" httpUri="https://zenodo.org/record/13757727/files/figure.png" pageId="19" pageNumber="332">Fig. 14</figureCitation>
), we do not consider the material from different parts of Mindanao as different species.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFC0DD6EFF2C6122FF281583" pageId="19" pageNumber="332" type="distribution">
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<emphasis id="70C8EAE9FFC0DD6EFF2C6122FEB9155F" bold="true" box="[199,353,1635,1661]" pageId="19" pageNumber="332">Distribution.</emphasis>
<collectingRegion id="8078F819FFC0DD6EFE856125FDC7155C" box="[366,543,1635,1662]" country="Philippines" name="Davao del Sur" pageId="19" pageNumber="332">Davao del Sur</collectingRegion>
: Mount Apo;
<collectingRegion id="8078F819FFC0DD6EFD266122FC61155F" box="[717,953,1635,1661]" country="Philippines" name="Misamis Occidental" pageId="19" pageNumber="332">Misamis Occidental</collectingRegion>
: Clarin River, Lake Linao [new locality record].
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFC0DD6EFF2C61EDFA4815CB" pageId="19" pageNumber="332" type="biology_ecology">
<paragraph id="420336FBFFC0DD6EFF2C61EDFA4815CB" blockId="19.[151,1436,1347,1770]" pageId="19" pageNumber="332">
<emphasis id="70C8EAE9FFC0DD6EFF2C61EDFEF415E4" bold="true" box="[199,300,1708,1734]" pageId="19" pageNumber="332">Ecology.</emphasis>
At Lake Linao, this species was quite abundant among shrubs and tall grasses mixed with herbaceous plants (
<figureCitation id="DA872A7EFFC0DD6EFF02618EFEE515C8" box="[233,317,1743,1770]" captionStart="FIGURE 15" captionStartId="20.[151,250,1926,1950]" captionTargetBox="[151,1436,181,1902]" captionTargetId="figure-19@20.[151,1436,181,1902]" captionTargetPageId="20" captionText="FIGURE 15. Anthracites zebra Hebard, 1922 male (A) and female (B) in their natural habitat at Lake Linao." figureDoi="http://doi.org/10.5281/zenodo.13757729" httpUri="https://zenodo.org/record/13757729/files/figure.png" pageId="19" pageNumber="332">Fig. 15</figureCitation>
). At night, adults and juveniles were observed to feed on flowers on numerous occasions (
<figureCitation id="DA872A7EFFC0DD6EFADB618EFA5B15CB" box="[1328,1411,1743,1769]" captionStart="FIGURE 16" captionStartId="21.[151,250,1926,1950]" captionTargetBox="[151,1436,181,1902]" captionTargetId="figure-17@21.[151,1436,181,1902]" captionTargetPageId="21" captionText="FIGURE 16. Anthracites zebra Hebard, 1922 females feeding on the flowers of a Melastoma plant at Lake Linao." figureDoi="http://doi.org/10.5281/zenodo.13757733" httpUri="https://zenodo.org/record/13757733/files/figure.png" pageId="19" pageNumber="332">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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Tan, Bahoy &amp; Sabang
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,
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(
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,
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,
<figureCitation id="DA872A7EFFD1DD7FFED367FAFE9F13F7" box="[312,327,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">3</figureCitation>
,
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)
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<emphasis id="70C8EAE9FFD1DD7FFF7C6642FE58123F" bold="true" box="[151,384,259,285]" pageId="2" pageNumber="315">Material examined.</emphasis>
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<collectingMunicipality id="A267AC81FFD1DD7FFBD36642FB45123F" box="[1080,1181,259,285]" pageId="2" pageNumber="315">Gingoog</collectingMunicipality>
,
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,
<location id="47636020FFD1DD7FFABA6645FED41263" LSID="urn:lsid:plazi:treatment:CA1587EDFFD1DD7BFF7C67D6FCDC113B:47636020FFD1DD7FFABA6645FED41263" country="Philippines" county="Mindanao" latitude="8.435183" longLatPrecision="1" longitude="125.00138" municipality="Gingoog" name="Mount Balatukan" pageId="2" pageNumber="315" stateProvince="Misamis Oriental">Mount Balatukan</location>
;
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,
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.a.s.l;
<date id="3602103BFFD1DD7FFD096666FC4A1263" box="[738,914,295,321]" pageId="2" pageNumber="315" value="2024-03-26">
<collectingDate id="2646E9D3FFD1DD7FFD096666FC4A1263" box="[738,914,295,321]" pageId="2" pageNumber="315" value="2024-03-26">26 March 2024</collectingDate>
</date>
, 16h50; inside primary forest, on a shrub leaf; coll.
<collectorName id="EF49532DFFD1DD7FFF24660AFE8C1247" box="[207,340,331,357]" pageId="2" pageNumber="315">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD1DD7FFE9D660AFDD71247" box="[374,527,331,357]" pageId="2" pageNumber="315">A.M. Sabang</collectorName>
; JS12 (
<collectionCode id="24ADAE3EFFD1DD7FFDB4660AFD7A1247" box="[607,674,331,357]" country="Philippines" name="Philippine National Museum" pageId="2" pageNumber="315">PNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C662EFABD1143" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<materialsCitation id="F2D43CA6FFD1DD7FFF2C662EFAB91143" collectingDate="2024-03-17" collectionCode="PNM" collectorName="D. C Bahoy &amp; A. M. Sabang &amp; D. C. Bahoy &amp; Gingoog City &amp; Barangay Kalagonoy" country="Philippines" county="Mindanao" elevation="1250" latitude="8.742659" location="Barangay Kalagonoy" longLatPrecision="1" longitude="125.0212" municipality="Gingoog City" pageId="2" pageNumber="315" specimenCount="6" specimenCount-male="6" stateProvince="Misamis Oriental" typeStatus="paratype">
<emphasis id="70C8EAE9FFD1DD7FFF2C662EFE5112AF" bold="true" box="[199,393,367,397]" pageId="2" pageNumber="315">
<typeStatus id="9D078859FFD1DD7FFF2C662EFEE712AB" box="[199,319,367,393]" pageId="2" pageNumber="315" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="54BAFD72FFD1DD7FFEBF6630FEA212AF" box="[340,378,369,397]" count="3" pageId="2" pageNumber="315" type="male">3♂</specimenCount>
):
</emphasis>
<collectingCountry id="3AAB766BFFD1DD7FFE7E662EFDE512AB" box="[405,573,367,393]" name="Philippines" pageId="2" pageNumber="315">PHILIPPINES</collectingCountry>
,
<collectingCounty id="AB624E77FFD1DD7FFDA7662EFD1812AB" box="[588,704,367,393]" pageId="2" pageNumber="315">Mindanao</collectingCounty>
,
<collectingRegion id="8078F819FFD1DD7FFD3B662EFC4412AB" box="[720,924,367,393]" country="Philippines" name="Misamis Oriental" pageId="2" pageNumber="315">Misamis Oriental</collectingRegion>
: •
<specimenCount id="54BAFD72FFD1DD7FFC2A662EFC3D12AB" box="[961,997,367,393]" count="1" pageId="2" pageNumber="315" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD1DD7FFC1C662EFB4D12AB" box="[1015,1173,367,393]" pageId="2" pageNumber="315">Gingoog City</collectingMunicipality>
,
<location id="47636020FFD1DD7FFB4E6631FA4012AB" LSID="urn:lsid:plazi:treatment:CA1587EDFFD1DD7BFF7C67D6FCDC113B:47636020FFD1DD7FFB4E6631FA4012AB" box="[1189,1432,367,394]" country="Philippines" county="Mindanao" latitude="8.742659" longLatPrecision="1" longitude="125.0212" municipality="Gingoog City" name="Barangay Kalagonoy" pageId="2" pageNumber="315" stateProvince="Misamis Oriental">Barangay Kalagonoy</location>
,
<location id="47636020FFD1DD7FFF7C66D5FEB8128F" LSID="urn:lsid:plazi:treatment:CA1587EDFFD1DD7BFF7C67D6FCDC113B:47636020FFD1DD7FFF7C66D5FEB8128F" box="[151,352,403,430]" country="Philippines" county="Mindanao" latitude="8.742659" longLatPrecision="1" longitude="125.0212" municipality="Gingoog City" name="Mount Balatukan" pageId="2" pageNumber="315" stateProvince="Misamis Oriental">Mount Balatukan</location>
;
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,
<geoCoordinate id="2788503CFFD1DD7FFE0B66D2FD86128C" box="[480,606,403,430]" degrees="125.0126" direction="east" orientation="longitude" pageId="2" pageNumber="315" precision="5" value="125.0126">E125.0126</geoCoordinate>
,
<quantity id="85449B1EFFD1DD7FFD8066D2FD11128F" box="[619,713,403,430]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="315" unit="m" value="1250.0">
<elevation id="C991D1C8FFD1DD7FFD8066D2FD11128F" box="[619,713,403,430]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="315" unit="m" value="1250.0">1250 m</elevation>
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.a.s.l;
<date id="3602103BFFD1DD7FFCE366D2FC63128F" box="[776,955,403,429]" pageId="2" pageNumber="315" value="2024-03-17">
<collectingDate id="2646E9D3FFD1DD7FFCE366D2FC63128F" box="[776,955,403,429]" pageId="2" pageNumber="315" value="2024-03-17">17 March 2024</collectingDate>
</date>
, 9h36; in an open area of the buffer zone of the foot of the stratovolcano, in between pandan leaf; coll.
<collectorName id="EF49532DFFD1DD7FFCA466F6FC1612F3" box="[847,974,439,465]" pageId="2" pageNumber="315">D.C Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD1DD7FFC1A66F6FB5212F3" box="[1009,1162,439,465]" pageId="2" pageNumber="315">A.M. Sabang</collectorName>
; K20 (MNHN-EO1353) •
<specimenCount id="54BAFD72FFD1DD7FFF41669AFF1712D7" box="[170,207,475,501]" count="1" pageId="2" pageNumber="315" type="male">1♂</specimenCount>
; same locality and date as previous, 1017h; an open area in secondary forest, at the buffer zone of the foot of the stratovolcano, in between pandan leaf; coll.
<collectorName id="EF49532DFFD1DD7FFD0166BEFCAB113B" box="[746,883,511,537]" pageId="2" pageNumber="315">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD1DD7FFC7766BEFBE0113B" box="[924,1080,511,537]" pageId="2" pageNumber="315">A.M. Sabang</collectorName>
; KS5 (MNHN-EO1354) •
<specimenCount id="54BAFD72FFD1DD7FFA9A66BEFA4E113B" box="[1393,1430,511,537]" count="1" pageId="2" pageNumber="315" type="male">1♂</specimenCount>
;
<collectorName id="EF49532DFFD1DD7FFF7C6562FEE8111F" box="[151,304,547,573]" pageId="2" pageNumber="315">Gingoog City</collectorName>
,
<collectorName id="EF49532DFFD1DD7FFED06565FDF1111F" box="[315,553,547,574]" pageId="2" pageNumber="315">Barangay Kalagonoy</collectorName>
,
<location id="47636020FFD1DD7FFDD86565FD21111F" LSID="urn:lsid:plazi:treatment:CA1587EDFFD1DD7BFF7C67D6FCDC113B:47636020FFD1DD7FFDD86565FD21111F" box="[563,761,547,574]" country="Philippines" county="Mindanao" latitude="8.742659" longLatPrecision="1" longitude="125.0212" municipality="Gingoog City" name="Mount Balatukan" pageId="2" pageNumber="315" stateProvince="Misamis Oriental">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD1DD7FFCEF6562FC5C111C" box="[772,900,547,574]" degrees="8.742659" direction="north" orientation="latitude" pageId="2" pageNumber="315" precision="1" value="8.742659">N8.742659</geoCoordinate>
,
<geoCoordinate id="2788503CFFD1DD7FFC656562FBFE111C" box="[910,1062,547,574]" degrees="125.021199" direction="east" orientation="longitude" pageId="2" pageNumber="315" precision="1" value="125.0212">E125.021199</geoCoordinate>
,
<quantity id="85449B1EFFD1DD7FFBDA6562FB54111F" box="[1073,1164,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="315" unit="m" value="1380.0">
<elevation id="C991D1C8FFD1DD7FFBDA6562FB54111F" box="[1073,1164,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="315" unit="m" value="1380.0">1380 m</elevation>
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.a.s.l; an open area inside secondary forest, in between pandan leaves;
<date id="3602103BFFD1DD7FFD606506FCE21143" box="[651,826,583,609]" pageId="2" pageNumber="315" value="2024-03-17">
<collectingDate id="2646E9D3FFD1DD7FFD606506FCE21143" box="[651,826,583,609]" pageId="2" pageNumber="315" value="2024-03-17">17 March 2024</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFD1DD7FFC966506FBDA1143" box="[893,1026,583,609]" pageId="2" pageNumber="315">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD1DD7FFBCF6506FB641143" box="[1060,1212,583,609]" pageId="2" pageNumber="315">A.M. Sabang</collectorName>
; KS31 (
<collectionCode id="24ADAE3EFFD1DD7FFAFD6506FA811143" box="[1302,1369,583,609]" country="Philippines" name="Philippine National Museum" pageId="2" pageNumber="315">PNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD1DD7FFF2C652AFDF0118B" pageId="2" pageNumber="315" type="etymology">
<paragraph id="420336FBFFD1DD7FFF2C652AFDF0118B" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C652AFE9611A7" bold="true" box="[199,334,619,645]" pageId="2" pageNumber="315">Etymology.</emphasis>
The species name refers to the wing-like titillators with their lateral arms strongly sclerotized and plate-like; alatus = winged in Latin.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD1DD7FFF2C65F2FBE31087" pageId="2" pageNumber="315" type="diagnosis">
<paragraph id="420336FBFFD1DD7FFF2C65F2FBE31087" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C65F2FE9A11EF" bold="true" box="[199,322,691,717]" pageId="2" pageNumber="315">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="85BC4D78FFD1DD7FFD7F65F2FBC511EC" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[660,1053,691,718]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="romblon">
<emphasis id="70C8EAE9FFD1DD7FFD7F65F2FCA011EF" box="[660,888,691,717]" italics="true" pageId="2" pageNumber="315">Anthracites romblon</emphasis>
<bibRefCitation id="262D4B0AFFD1DD7FFC9665F2FBC511EC" author="Ingrisch, S." box="[893,1053,691,718]" pageId="2" pageNumber="315" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="8078F819FFD1DD7FFBB665F2FB1D11EF" box="[1117,1221,691,717]" country="Philippines" name="Romblon" pageId="2" pageNumber="315">Romblon</collectingRegion>
Island,
<taxonomicName id="85BC4D78FFD1DD7FFAF065F2FE4311D0" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="70C8EAE9FFD1DD7FFAF065F2FF2C11D3" italics="true" pageId="2" pageNumber="315">Anthracites sinuatus</emphasis>
<bibRefCitation id="262D4B0AFFD1DD7FFF106596FE4311D0" author="Ingrisch, S." box="[251,411,727,754]" pageId="2" pageNumber="315" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="85BC4D78FFD1DD7FFE3B6596FCB111D0" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[464,873,727,754]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="unispinus">
<emphasis id="70C8EAE9FFD1DD7FFE3B6596FD1A11D3" box="[464,706,727,753]" italics="true" pageId="2" pageNumber="315">Anthracites unispinus</emphasis>
<bibRefCitation id="262D4B0AFFD1DD7FFD236596FCB111D0" author="Ingrisch, S." box="[712,873,727,754]" pageId="2" pageNumber="315" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
from Luzon by the face with a pair of black bands; but differs from these species and all other congeners by the pronotum having its anterior margin with two lateral bands, near posterior margin with two large black patches having well-defined margins forming triangles, male tegmen without dark lateral parts, male cercus F-shaped with the apex having two inner processes, male subgenital plate with its posterior margin having dense long setae, and the male titillators strongly sclerotized and wing-like with the lateral arm plate-like (instead of slender with different shapes at the apex).
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD1DD7BFF2C64EEFDD512D7" lastPageId="6" lastPageNumber="319" pageId="2" pageNumber="315" type="description">
<paragraph id="420336FBFFD1DD7FFF2C64EEFCEE10EB" blockId="2.[151,1437,259,1978]" box="[199,822,943,970]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C64EEFE8110EB" bold="true" box="[199,345,943,969]" pageId="2" pageNumber="315">Description.</emphasis>
Habitus of male as shown in
<figureCitation id="DA872A7EFFD1DD7FFD4E64EEFCD810EB" box="[677,768,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="2" pageNumber="315">Figs 1A</figureCitation>
,
<figureCitation id="DA872A7EFFD1DD7FFCE764EEFCEA10EB" box="[780,818,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="2" pageNumber="315">2A</figureCitation>
.
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C6492FDFF1717" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C6492FED310CF" box="[199,267,979,1005]" italics="true" pageId="2" pageNumber="315">Head.</emphasis>
Fastigium verticis compressed laterally, in dorsal view conical with faint fine longitudinal groove (
<figureCitation id="DA872A7EFFD1DD7FFA846492FF651733" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3A</figureCitation>
); apex truncated, shorter than scapus (
<figureCitation id="DA872A7EFFD1DD7FFD9F64B6FD151733" box="[628,717,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3B</figureCitation>
). Lateral ocelli anterior to median one. Frons shining with few shallowly impressed dots (
<figureCitation id="DA872A7EFFD1DD7FFE29635AFDC21717" box="[450,538,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C637EFE881613" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C637EFEF8177B" box="[199,288,1087,1113]" italics="true" pageId="2" pageNumber="315">Thorax.</emphasis>
Pronotum smooth, disc broadly rounded into paranota, apical area rounded and faintly shouldered; anterior transverse sulcus short and broadly rounded, posterior transverse sulcus longer and angularly rounded; anterior margin truncated and straight; posterior margin subtruncated (
<figureCitation id="DA872A7EFFD1DD7FFC3A63C6FBE81783" box="[977,1072,1159,1185]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3A</figureCitation>
). Paranota with ventral margin concave, posterior margin obliquely rising posteriorly and slightly concave (
<figureCitation id="DA872A7EFFD1DD7FFC1063EAFB8C17E7" box="[1019,1108,1195,1221]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3B</figureCitation>
). Acoustic spiracle large and oval, completely covered by paranota. Prosternum with two long acute spines pointing nearly ventrad. Mesosternal and metasternal lobes conical with acute apices. Internal and external tympanum on anterior tibia slightly swollen with narrow slit.
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C627DFA8A16E3" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C627DFEDC1677" box="[199,260,1340,1365]" italics="true" pageId="2" pageNumber="315">Legs.</emphasis>
Anterior coxa with spine long and slightly curved. Femora with the following number of spines on ventral margins: anterior femur 6–8 external, 5–6 internal; middle femur 6–7 external, 3 internal; posterior femur 8–9 external, 1–2 internal. Knee lobes of anterior and middle femora obtuse or subacute externally and spinose internally; knee of posterior femur bispinose on both sides, longer than those of anterior and middle femora.
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C628DFB6C1528" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C628DFE7F16C7" box="[199,423,1483,1509]" italics="true" pageId="2" pageNumber="315">Tegmina and wings.</emphasis>
Tegmen micropterous, mirror area fully covered by pronotal disc (
<figureCitation id="DA872A7EFFD1DD7FFB79628AFB3416C7" box="[1170,1260,1483,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3A</figureCitation>
), reaching apex of third abdominal tergite, with apex subtruncated; hind wing present, but covered by tegmen.
</paragraph>
<paragraph id="420336FBFFD1DD7FFF2C6152FA91149B" blockId="2.[151,1437,259,1978]" pageId="2" pageNumber="315">
<emphasis id="70C8EAE9FFD1DD7FFF2C6152FEE1150F" box="[199,313,1555,1581]" italics="true" pageId="2" pageNumber="315">Abdomen.</emphasis>
Tenth abdominal tergite transverse, distinctly broader than long, at posterior margin truncated with two very short lateral lobules; posterior margin of lateral lobules raised and having fine setae (
<figureCitation id="DA872A7EFFD1DD7FFB206176FAFF1573" box="[1227,1319,1591,1617]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3D</figureCitation>
). Epiproct broadly tongue shaped, basal half having longitudinal groove at middle (
<figureCitation id="DA872A7EFFD1DD7FFC54611AFBCE1557" box="[959,1046,1627,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3E</figureCitation>
). Cercus short and stout, F-shaped, compressed, distal end roundly truncated (
<figureCitation id="DA872A7EFFD1DD7FFD83613EFD1815BB" box="[616,704,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3D</figureCitation>
); at apex with two inner processes pointing inwards (perpendicular to length of cercus) (
<figureCitation id="DA872A7EFFD1DD7FFE9461E2FE01159F" box="[383,473,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3D</figureCitation>
); dorso-distal process conical, larger with apex having small sclerotized tooth, ventroproximal process parallel to and slightly shorter than dorso-distal process, cylindrical, slenderer, with apex obtuse (
<figureCitation id="DA872A7EFFD1DD7FFF7461AAFEC61427" box="[159,286,1771,1797]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Figs 3D–G</figureCitation>
). Subgenital plate about as long as wide; with anterior margin concave; apical end angularly emarginated, posterior margin at middle truncated with dense long setae (
<figureCitation id="DA872A7EFFD1DD7FFCAE604EFC08140B" box="[837,976,1807,1833]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Figs 3F, 3G</figureCitation>
); margin around bases of styli on dorsal side swollen and with triangular internal tooth of subacute sclerotized apex (
<figureCitation id="DA872A7EFFD1DD7FFC016072FB99146F" box="[1002,1089,1843,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3E</figureCitation>
). Styli stout with apex rounded (
<figureCitation id="DA872A7EFFD1DD7FFF746016FF2B1453" box="[159,243,1879,1905]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="2" pageNumber="315">Fig. 3F</figureCitation>
). Titillators (
<figureCitation id="DA872A7EFFD1DD7FFE6A6016FE1B1453" box="[385,451,1879,1905]" captionStart="FIGURE 4" captionStartId="6.[151,250,1662,1686]" captionTargetBox="[166,1414,561,1614]" captionTargetId="figure-175@6.[151,1436,561,1639]" captionTargetPageId="6" captionText="FIGURE 4. Anthracites alatus sp. nov. ♂ titillators in ventro-proximal (A), dorso-distal (B) and ventro-distal (C) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13757707" httpUri="https://zenodo.org/record/13757707/files/figure.png" pageId="2" pageNumber="315">Fig. 4</figureCitation>
) separate, strongly sclerotized, wing-like; lateral arm plate-like, internal-half very broad, external-third stoutly tongue shaped; apical end broadly rounded, pointing dorso-distally, along margin slightly folded; base with proximal end clog-shaped; provided with two pairs of lateral sclerites weakly sclerotized.
</paragraph>
<caption id="16C36673FFD0DD7EFF7C60C9FABF14E6" ID-DOI="http://doi.org/10.5281/zenodo.13757699" ID-Zenodo-Dep="13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="3" pageNumber="316" startId="3.[151,250,1928,1952]" targetBox="[178,1410,181,1904]" targetPageId="3" targetType="figure">
<paragraph id="420336FBFFD0DD7EFF7C60C9FABF14E6" blockId="3.[151,1436,1928,1988]" pageId="3" pageNumber="316">
<emphasis id="70C8EAE9FFD0DD7EFF7C60C9FEC01482" bold="true" box="[151,280,1928,1952]" pageId="3" pageNumber="316">FIGURE 1.</emphasis>
Dorsal habitus of ♂
<taxonomicName id="85BC4D78FFD0DD7EFDEE60C9FD1B1482" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[517,707,1928,1952]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
<emphasis id="70C8EAE9FFD0DD7EFDEE60C9FD1B1482" box="[517,707,1928,1952]" italics="true" pageId="3" pageNumber="316">Anthracites alatus</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFD0DD7EFD2560C9FCFB1482" bold="true" box="[718,803,1928,1952]" pageId="3" pageNumber="316">
<taxonomicNameLabel id="6BFB5792FFD0DD7EFD2560C9FCFB1482" box="[718,803,1928,1952]" pageId="3" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="85BC4D78FFD0DD7EFC8F60C9FB311482" authority="Dohrn, 1905" authorityName="Dohrn" authorityYear="1905" box="[868,1257,1928,1952]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="70C8EAE9FFD0DD7EFC8F60C9FB811482" box="[868,1113,1928,1952]" italics="true" pageId="3" pageNumber="316">Anthracites geniculatus</emphasis>
<bibRefCitation id="262D4B0AFFD0DD7EFB8F60C9FB311482" author="Dohrn, H. W. L." box="[1124,1257,1928,1952]" pageId="3" pageNumber="316" pagination="237 - 246" refId="ref9191" refString="Dohrn, H. W. L. (1905) Orthopterologisches aus dem Stettiner Museum I. Neue und ungenugend bekannte Conocephaliden des indo-malayischen Gebietes. Stettiner Entomologische Zeitung, 66, 237 - 246." type="journal article" year="1905">Dohrn, 1905</bibRefCitation>
</taxonomicName>
(B),
<taxonomicName id="85BC4D78FFD0DD7EFACC60C9FE7114E6" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFD0DD7EFACC60C9FEC814E6" italics="true" pageId="3" pageNumber="316">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFD0DD7EFEFC60EDFE7114E6" author="Ingrisch, S." box="[279,425,1964,1988]" pageId="3" pageNumber="316" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
(C),
<taxonomicName id="85BC4D78FFD0DD7EFE3560EDFD5F14E6" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[478,647,1964,1988]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="sebu" status="sp. nov.">
<emphasis id="70C8EAE9FFD0DD7EFE3560EDFD5F14E6" box="[478,647,1964,1988]" italics="true" pageId="3" pageNumber="316">Anthracites sebu</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFD0DD7EFD6560EDFD0614E6" bold="true" box="[654,734,1964,1988]" pageId="3" pageNumber="316">
<taxonomicNameLabel id="6BFB5792FFD0DD7EFD6560EDFD0614E6" box="[654,734,1964,1988]" pageId="3" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(D) and
<taxonomicName id="85BC4D78FFD0DD7EFCD260EDFBA514E6" authority="Hebard, 1922" authorityName="Hebard" authorityYear="1922" box="[825,1149,1964,1988]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="zebra">
<emphasis id="70C8EAE9FFD0DD7EFCD260EDFC3414E6" box="[825,1004,1964,1988]" italics="true" pageId="3" pageNumber="316">Anthracites zebra</emphasis>
<bibRefCitation id="262D4B0AFFD0DD7EFC1860EDFBA514E6" author="Hebard, M." box="[1011,1149,1964,1988]" pageId="3" pageNumber="316" pagination="121 - 299" refId="ref9228" refString="Hebard, M. (1922) Studies in Malayan, Melanesian and Australian Tettigoniidae (Orthoptera). Proceedings of the Academy of Natural Sciences, Philadelphia, 74, 121 - 299, pls. 11 - 22." type="journal article" year="1922">Hebard, 1922</bibRefCitation>
</taxonomicName>
(E). Scale bars: 5 mm.
</paragraph>
</caption>
<caption id="16C36673FFD7DD79FF7C6044FABF1463" ID-DOI="http://doi.org/10.5281/zenodo.13757701" ID-Zenodo-Dep="13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="4" pageNumber="317" startId="4.[151,250,1797,1821]" targetBox="[178,1409,181,1774]" targetPageId="4" targetType="figure">
<paragraph id="420336FBFFD7DD79FF7C6044FABF1463" blockId="4.[151,1436,1797,1858]" pageId="4" pageNumber="317">
<emphasis id="70C8EAE9FFD7DD79FF7C6044FECA143F" bold="true" box="[151,274,1797,1821]" pageId="4" pageNumber="317">FIGURE 2</emphasis>
Lateral habitus of ♂
<taxonomicName id="85BC4D78FFD7DD79FDEF6047FD1A143C" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[516,706,1798,1822]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
<emphasis id="70C8EAE9FFD7DD79FDEF6047FD1A143C" box="[516,706,1798,1822]" italics="true" pageId="4" pageNumber="317">Anthracites alatus</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFD7DD79FD266047FCFB143C" bold="true" box="[717,803,1798,1822]" pageId="4" pageNumber="317">
<taxonomicNameLabel id="6BFB5792FFD7DD79FD266047FCFB143C" box="[717,803,1798,1822]" pageId="4" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="85BC4D78FFD7DD79FC886047FB31143F" authority="Dohrn, 1905" authorityName="Dohrn" authorityYear="1905" box="[867,1257,1797,1822]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="70C8EAE9FFD7DD79FC886047FB80143C" box="[867,1112,1798,1822]" italics="true" pageId="4" pageNumber="317">Anthracites geniculatus</emphasis>
<bibRefCitation id="262D4B0AFFD7DD79FB8F6047FB31143F" author="Dohrn, H. W. L." box="[1124,1257,1797,1822]" pageId="4" pageNumber="317" pagination="237 - 246" refId="ref9191" refString="Dohrn, H. W. L. (1905) Orthopterologisches aus dem Stettiner Museum I. Neue und ungenugend bekannte Conocephaliden des indo-malayischen Gebietes. Stettiner Entomologische Zeitung, 66, 237 - 246." type="journal article" year="1905">Dohrn, 1905</bibRefCitation>
</taxonomicName>
(B),
<taxonomicName id="85BC4D78FFD7DD79FACC6047FE711463" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFD7DD79FACC6047FEC81460" italics="true" pageId="4" pageNumber="317">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFD7DD79FEFC606BFE711463" author="Ingrisch, S." box="[279,425,1833,1858]" pageId="4" pageNumber="317" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
(C),
<taxonomicName id="85BC4D78FFD7DD79FE35606BFD5F1460" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[478,647,1834,1858]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="sebu" status="sp. nov.">
<emphasis id="70C8EAE9FFD7DD79FE35606BFD5F1460" box="[478,647,1834,1858]" italics="true" pageId="4" pageNumber="317">Anthracites sebu</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFD7DD79FD65606BFD061460" bold="true" box="[654,734,1834,1858]" pageId="4" pageNumber="317">
<taxonomicNameLabel id="6BFB5792FFD7DD79FD65606BFD061460" box="[654,734,1834,1858]" pageId="4" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(D) and
<taxonomicName id="85BC4D78FFD7DD79FCD2606BFBA51460" authority="Hebard, 1922" authorityName="Hebard" authorityYear="1922" box="[825,1149,1834,1858]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="zebra">
<emphasis id="70C8EAE9FFD7DD79FCD2606BFC341460" box="[825,1004,1834,1858]" italics="true" pageId="4" pageNumber="317">Anthracites zebra</emphasis>
<bibRefCitation id="262D4B0AFFD7DD79FC18606BFBA51460" author="Hebard, M." box="[1011,1149,1834,1858]" pageId="4" pageNumber="317" pagination="121 - 299" refId="ref9228" refString="Hebard, M. (1922) Studies in Malayan, Melanesian and Australian Tettigoniidae (Orthoptera). Proceedings of the Academy of Natural Sciences, Philadelphia, 74, 121 - 299, pls. 11 - 22." type="journal article" year="1922">Hebard, 1922</bibRefCitation>
</taxonomicName>
(E). Scale bars: 5 mm.
</paragraph>
</caption>
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<emphasis id="70C8EAE9FFD6DD78FF7C61C5FECB15BE" bold="true" box="[151,275,1668,1692]" pageId="5" pageNumber="318">FIGURE 3.</emphasis>
<taxonomicName id="85BC4D78FFD6DD78FEF161C5FE0C15BE" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[282,468,1668,1692]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
<emphasis id="70C8EAE9FFD6DD78FEF161C5FE0C15BE" box="[282,468,1668,1692]" italics="true" pageId="5" pageNumber="318">Anthracites alatus</emphasis>
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<emphasis id="70C8EAE9FFD6DD78FE3061C5FDF415BE" bold="true" box="[475,556,1668,1692]" pageId="5" pageNumber="318">
<taxonomicNameLabel id="6BFB5792FFD6DD78FE3061C5FDF415BE" box="[475,556,1668,1692]" pageId="5" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
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♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G).
</paragraph>
</caption>
<paragraph id="420336FBFFD6DD78FF2C6077FE461472" blockId="5.[151,1437,1846,2017]" box="[199,414,1846,1872]" pageId="5" pageNumber="318">Female. Unknown.</paragraph>
<paragraph id="420336FBFFD6DD7BFF2C601BFC1A12F3" blockId="5.[151,1437,1846,2017]" lastBlockId="6.[151,1436,151,537]" lastPageId="6" lastPageNumber="319" pageId="5" pageNumber="318">
Colouration. Body generally pale yellow when preserved (
<figureCitation id="DA872A7EFFD6DD78FC8B601BFC671456" box="[864,959,1882,1908]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="5" pageNumber="318">Figs 1A</figureCitation>
,
<figureCitation id="DA872A7EFFD6DD78FC27601BFC2A1456" box="[972,1010,1882,1908]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="5" pageNumber="318">2A</figureCitation>
), green when alive. Face with pair of black bands from inner margin of antennal scapus to outer base of mandibles, widening ventrad (
<figureCitation id="DA872A7EFFD6DD78FB31603FFAEA14BA" box="[1242,1330,1918,1944]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="5" pageNumber="318">Fig. 3C</figureCitation>
). Median ocellus cream coloured. Scapus with basal-half and dorso-inner side black, otherwise pale yellow; antennae yellow (
<figureCitation id="DA872A7EFFD6DD78FF746087FEC614C2" box="[159,286,1990,2016]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="5" pageNumber="318">Figs 3A–C</figureCitation>
). Maxillary palpi with segments pale; mandible black, clypeus with pair of black bands continuing from frons (
<figureCitation id="DA872A7EFFD5DD7BFF0B67D6FEEF1393" box="[224,311,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3B</figureCitation>
). Pronotum pale yellow, anterior margin with two lateral bands, near posterior margin with two large black patches having well-defined margins shaping like triangles (
<figureCitation id="DA872A7EFFD5DD7BFC6E67FAFBCC13F7" box="[901,1044,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Figs 3A, 3B</figureCitation>
). Tegmen with lateral field at base bright yellow, otherwise veins pale yellow and cells generally black (
<figureCitation id="DA872A7EFFD5DD7BFC7F679EFC3213DB" box="[916,1002,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3B</figureCitation>
); dorsal field mostly pale yellow, some cells dark brown (
<figureCitation id="DA872A7EFFD5DD7BFE8E6642FE18123F" box="[357,448,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3A</figureCitation>
). Anterior and middle femora unicolourous pale yellow; fore and middle tibiae similar, but fore tibia having distal part of tympanum with small black dorsum spot and distally slightly darker (
<figureCitation id="DA872A7EFFD5DD7BFADD6666FA571263" box="[1334,1423,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3B</figureCitation>
). Posterior femur pale yellow, knees slightly darker. Posterior tibia pale yellow with spines brown. Tarsus cream coloured with distal margin black. Abdominal tergite pale yellow, with posterior margin having lateral bands that become narrower towards apical segments (
<figureCitation id="DA872A7EFFD5DD7BFD6266D2FD3C128F" box="[649,740,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3D</figureCitation>
). Male tenth abdominal tergite, epiproct and cercus generally orange yellow (
<figureCitation id="DA872A7EFFD5DD7BFEAD66F6FE7812F3" box="[326,416,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3D</figureCitation>
). Sternites and subgenital plate yellow (
<figureCitation id="DA872A7EFFD5DD7BFCB466F6FC6D12F3" box="[863,949,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1668,1692]" captionTargetBox="[151,1432,181,1644]" captionTargetId="figure-17@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 3. Anthracites alatus sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (with titillator, t, intact) (E), ventral (F) and ventro-posterior (G) views.The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757705" httpUri="https://zenodo.org/record/13757705/files/figure.png" pageId="6" pageNumber="319">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFD5DD7BFF2C669AFDD512D7" blockId="6.[151,1436,151,537]" box="[199,525,475,501]" pageId="6" pageNumber="319">
<emphasis id="70C8EAE9FFD5DD7BFF2C669AFEA612D7" bold="true" box="[199,382,475,501]" pageId="6" pageNumber="319">Measurements.</emphasis>
See
<tableCitation id="0F3E0340FFD5DD7BFE58669AFDDF12D7" box="[435,519,475,501]" captionStart="TABLE 1" captionStartId="22.[151,239,152,176]" captionText="TABLE 1. Measurements of Anthracites, in mm." pageId="6" pageNumber="319">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD5DD7BFF2C66BFFCDC113B" box="[199,772,510,537]" pageId="6" pageNumber="319" type="distribution">
<paragraph id="420336FBFFD5DD7BFF2C66BFFCDC113B" blockId="6.[151,1436,151,537]" box="[199,772,510,537]" pageId="6" pageNumber="319">
<emphasis id="70C8EAE9FFD5DD7BFF2C66BFFEB9113A" bold="true" box="[199,353,510,536]" pageId="6" pageNumber="319">Distribution.</emphasis>
<collectingRegion id="8078F819FFD5DD7BFE8C66BEFDF7113B" box="[359,559,511,537]" country="Philippines" name="Misamis Oriental" pageId="6" pageNumber="319">Misamis Oriental</collectingRegion>
: Mount Balatukan.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="00E4F0B5437CF35C882C1D90B4A6F126">A taxonomic review on the katydids of the genus Anthracites Redtenbacher, 1891 (Tettigoniidae: Conocephalinae: Agraeciini) from Mindanao, Philippines</mods:title>
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<figureCitation id="DA872A7EFFD4DD7AFEBA651CFEB81155" box="[337,352,605,631]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[190,1382,199,1916]" captionTargetId="figure-17@9.[181,1407,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Anthracites geniculatus Dohrn, 1905 ♂ titillators in dorsal (A), dorso-distal (B, C), dorso-proximal (D), ventral (E), ventro-distal (F) and lateral (G) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13757711" httpUri="https://zenodo.org/record/13757711/files/figure.png" pageId="7" pageNumber="320">6</figureCitation>
)
</paragraph>
</subSubSection>
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<treatmentCitationGroup id="62AC11D5FFD4DD7AFF7C65DAFB5D1191" box="[151,1157,667,692]" pageId="7" pageNumber="320">
<treatmentCitation id="C31D10EAFFD4DD7AFF7C65DAFDE71191" author="Dohrn, H. W. L." box="[151,575,667,691]" page="241" pageId="7" pageNumber="320" year="1905">
<taxonomicName id="85BC4D78FFD4DD7AFF7C65DAFDE71191" ID-CoL="5VGQV" authority="Dohrn, 1905: 241" authorityName="Dohrn" authorityPageNumber="241" authorityYear="1905" box="[151,575,667,691]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="320" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="70C8EAE9FFD4DD7AFF7C65DAFE5F1191" box="[151,391,667,691]" italics="true" pageId="7" pageNumber="320">Anthracites geniculatus</emphasis>
<bibRefCitation id="262D4B0AFFD4DD7AFE6665DAFDE71191" author="Dohrn, H. W. L." box="[397,575,667,691]" pageId="7" pageNumber="320" pagination="237 - 246" refId="ref9191" refString="Dohrn, H. W. L. (1905) Orthopterologisches aus dem Stettiner Museum I. Neue und ungenugend bekannte Conocephaliden des indo-malayischen Gebietes. Stettiner Entomologische Zeitung, 66, 237 - 246." type="journal article" year="1905">Dohrn, 1905: 241</bibRefCitation>
</taxonomicName>
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—
<treatmentCitation id="C31D10EAFFD4DD7AFDB165DAFCD11191" author="Kirby, W. F." box="[602,777,667,691]" page="263" pageId="7" pageNumber="320" year="1906">
<bibRefCitation id="262D4B0AFFD4DD7AFDB165DAFCD11191" author="Kirby, W. F." box="[602,777,667,691]" pageId="7" pageNumber="320" pagination="1 - 562" refId="ref9378" refString="Kirby, W. F. (1906) Orthoptera Saltatoria. Part I. (Achetidae et Phasgonuridae). In: A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae). Vol. 2. British Museum (Natural History), London, pp. i - viii + 1 - 562." type="book chapter" year="1906">Kirby, 1906: 263</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C31D10EAFFD4DD7AFCFF65DDFC621191" author="Karny, H. H." box="[788,954,667,692]" page="25" pageId="7" pageNumber="320" year="1912">
<bibRefCitation id="262D4B0AFFD4DD7AFCFF65DDFC621191" author="Karny, H. H." box="[788,954,667,692]" pageId="7" pageNumber="320" refId="ref9326" refString="Karny, H. H. (1912) Orthoptera Fam. Locustidae. Subfam. Agraeciinae. In: Wytsman, P. (Ed.), Genera Insectorum. Vol. 141. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 47." type="book" year="1912">Karny, 1912: 25</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C31D10EAFFD4DD7AFC2E65DAFB591191" author="Ingrisch, S." box="[965,1153,667,691]" page="79" pageId="7" pageNumber="320" year="2015">
<bibRefCitation id="262D4B0AFFD4DD7AFC2E65DAFB591191" author="Ingrisch, S." box="[965,1153,667,691]" pageId="7" pageNumber="320" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015: 79</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD4DD7AFF7C659BFEB2171B" pageId="7" pageNumber="320" type="materials_examined">
<paragraph id="420336FBFFD4DD7AFF7C659BFCB0101F" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF7C659BFC7511D6" bold="true" box="[151,941,730,757]" pageId="7" pageNumber="320">
Material examined (images and description from
<bibRefCitation id="262D4B0AFFD4DD7AFD1A659BFC7811D6" author="Ingrisch, S." box="[753,928,730,756]" pageId="7" pageNumber="320" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
).
</emphasis>
<materialsCitation id="F2D43CA6FFD4DD7AFC5D659AFCB0101F" collectingDate="1959-11-13" collectionCode="BPBM" collectorName="L. W. Quate &amp; C. M. Yoshimoto" country="Philippines" county="San Francisco" location="Mindanao" municipality="Other" pageId="7" pageNumber="320" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<emphasis id="70C8EAE9FFD4DD7AFC5D659AFB8511DA" bold="true" box="[950,1117,731,760]" pageId="7" pageNumber="320">
<typeStatus id="9D078859FFD4DD7AFC5D659AFBFF11D7" box="[950,1063,731,757]" pageId="7" pageNumber="320" type="holotype">Holotype</typeStatus>
: •
<specimenCount id="54BAFD72FFD4DD7AFBA3659DFB8511DA" box="[1096,1117,732,760]" count="1" pageId="7" pageNumber="320" type="female">♀</specimenCount>
</emphasis>
;
<collectingCountry id="3AAB766BFFD4DD7AFB85659AFACE11D7" box="[1134,1302,731,757]" name="Philippines" pageId="7" pageNumber="320">PHILIPPINES</collectingCountry>
,
<location id="47636020FFD4DD7AFAC8659AFA4011D7" LSID="urn:lsid:plazi:treatment:CA1587EDFFD4DD75FF7C6578FA44123F:47636020FFD4DD7AFAC8659AFA4011D7" box="[1315,1432,731,757]" country="Philippines" county="San Francisco" municipality="Other" name="Mindanao" pageId="7" pageNumber="320">Mindanao</location>
; MZPW.
<emphasis id="70C8EAE9FFD4DD7AFF1765BFFE95103A" bold="true" box="[252,333,766,792]" pageId="7" pageNumber="320">
<collectingMunicipality id="A267AC81FFD4DD7AFF1765BFFE90103A" box="[252,328,766,792]" pageId="7" pageNumber="320">Other</collectingMunicipality>
:
</emphasis>
•
<specimenCount id="54BAFD72FFD4DD7AFE9B65BEFE4B103E" box="[368,403,767,796]" count="1" pageId="7" pageNumber="320" type="female">
1
<emphasis id="70C8EAE9FFD4DD7AFE956441FE4B103E" bold="true" box="[382,403,768,796]" pageId="7" pageNumber="320">♀</emphasis>
</specimenCount>
;
<collectingCountry id="3AAB766BFFD4DD7AFE4C65BEFD97103B" box="[423,591,767,793]" name="Philippines" pageId="7" pageNumber="320">PHILIPPINES</collectingCountry>
, Mindanao. Agusan,
<collectingCounty id="AB624E77FFD4DD7AFCA765BEFC2B103B" box="[844,1011,767,793]" pageId="7" pageNumber="320">San Francisco</collectingCounty>
,
<quantity id="85449B1EFFD4DD7AFBE865BEFB88103B" box="[1027,1104,767,793]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="320" unit="km" value="10.0">10 km</quantity>
SE, [8°29'N, 125°59'E];
<date id="3602103BFFD4DD7AFA6B65BEFE97101F" pageId="7" pageNumber="320" value="1959-11-13">
<collectingDate id="2646E9D3FFD4DD7AFA6B65BEFE97101F" pageId="7" pageNumber="320" value="1959-11-13">13 November 1959</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFD4DD7AFE786462FDCC101F" box="[403,532,803,829]" pageId="7" pageNumber="320">L.W. Quate</collectorName>
&amp;
<collectorName id="EF49532DFFD4DD7AFDD36462FD2C101F" box="[568,756,803,829]" pageId="7" pageNumber="320">C.M. Yoshimoto</collectorName>
; (
<collectionCode id="24ADAE3EFFD4DD7AFCE36462FC84101F" box="[776,860,803,829]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="320" type="Museum">BPBM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="420336FBFFD4DD7AFF2C6406FCE3108B" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C6406FE301042" bold="true" box="[199,488,838,865]" pageId="7" pageNumber="320">New material examined.</emphasis>
<materialsCitation id="F2D43CA6FFD4DD7AFE046406FCEF108B" collectingDate="2022-09-23" collectionCode="PNM" collectorName="M. Salvador &amp; Salvador, J. A &amp; Salvador, M &amp; Astillo, J &amp; Sabacajan, J. P &amp; Sabacajan, E" country="Philippines" county="Mindanao" elevation="134" latitude="8.468" location="Mount Magdiwata" longLatPrecision="77" longitude="125.973" municipality="San Francisco" pageId="7" pageNumber="320" specimenCount="135" specimenCount-male="1" stateProvince="Agusan del Sur">
<collectingCountry id="3AAB766BFFD4DD7AFE046406FD4F1043" box="[495,663,839,865]" name="Philippines" pageId="7" pageNumber="320">PHILIPPINES</collectingCountry>
,
<collectingCounty id="AB624E77FFD4DD7AFD4A6406FCCD1043" box="[673,789,839,865]" pageId="7" pageNumber="320">Mindanao</collectingCounty>
,
<collectingRegion id="8078F819FFD4DD7AFCF46406FC171043" box="[799,975,839,865]" country="Philippines" name="Agusan del Sur" pageId="7" pageNumber="320">Agusan del Sur</collectingRegion>
: •
<specimenCount id="54BAFD72FFD4DD7AFC026406FBD61043" box="[1001,1038,839,865]" count="1" pageId="7" pageNumber="320" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD4DD7AFBF16406FB631043" box="[1050,1211,839,865]" pageId="7" pageNumber="320">San Francisco</collectingMunicipality>
,
<location id="47636020FFD4DD7AFB2E6406FA411043" LSID="urn:lsid:plazi:treatment:CA1587EDFFD4DD75FF7C6578FA44123F:47636020FFD4DD7AFB2E6406FA411043" box="[1221,1433,839,865]" country="Philippines" county="Mindanao" latitude="8.468" longLatPrecision="77" longitude="125.973" municipality="San Francisco" name="Mount Magdiwata" pageId="7" pageNumber="320" stateProvince="Agusan del Sur">Mount Magdiwata</location>
;
<geoCoordinate id="2788503CFFD4DD7AFF7C642AFF3510A7" box="[151,237,875,901]" degrees="8.468" direction="north" orientation="latitude" pageId="7" pageNumber="320" precision="55" value="8.468">N8.468</geoCoordinate>
,
<geoCoordinate id="2788503CFFD4DD7AFF13642BFEBF10A7" box="[248,359,874,901]" degrees="125.973" direction="east" orientation="longitude" pageId="7" pageNumber="320" precision="55" value="125.973">E125.973</geoCoordinate>
,
<specimenCount id="54BAFD72FFD4DD7AFE99642AFE1D10A7" box="[370,453,875,901]" count="134" pageId="7" pageNumber="320" type="generic">
<quantity id="85449B1EFFD4DD7AFE99642AFE1810A7" box="[370,448,875,901]" metricMagnitude="2" metricUnit="m" metricValue="1.34" pageId="7" pageNumber="320" unit="m" value="134.0">
<elevation id="C991D1C8FFD4DD7AFE99642AFE1810A7" box="[370,448,875,901]" metricMagnitude="2" metricUnit="m" metricValue="1.34" pageId="7" pageNumber="320" unit="m" value="134.0">134 m</elevation>
</quantity>
.
</specimenCount>
a.s.l.;
<date id="3602103BFFD4DD7AFDE8642AFD3910A7" box="[515,737,875,901]" pageId="7" pageNumber="320" value="2022-09-23">
<collectingDate id="2646E9D3FFD4DD7AFDE8642AFD3910A7" box="[515,737,875,901]" pageId="7" pageNumber="320" value="2022-09-23">23 September 2022</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFD4DD7AFCC8642AFC7710A7" box="[803,943,875,901]" pageId="7" pageNumber="320">M. Salvador</collectorName>
, J.A.
<collectorName id="EF49532DFFD4DD7AFC04642AFBA310A7" box="[1007,1147,875,901]" pageId="7" pageNumber="320">Salvador, M</collectorName>
.
<collectorName id="EF49532DFFD4DD7AFB6F642AFB3510A7" box="[1156,1261,875,901]" pageId="7" pageNumber="320">Astillo, J</collectorName>
.
<collectorName id="EF49532DFFD4DD7AFB1C642AFA4110A7" box="[1271,1433,875,901]" pageId="7" pageNumber="320">Sabacajan, J.P</collectorName>
.
<collectorName id="EF49532DFFD4DD7AFF7C64CEFEF3108B" box="[151,299,911,937]" pageId="7" pageNumber="320">Sabacajan, E</collectorName>
. Astillo III; 11JAS287, TRORT138-23 (
<collectionCode id="24ADAE3EFFD4DD7AFD0764CEFCF7108B" box="[748,815,911,937]" country="Philippines" name="Philippine National Museum" pageId="7" pageNumber="320">PNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="420336FBFFD4DD7AFF2C64F2FEB2171B" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<materialsCitation id="F2D43CA6FFD4DD7AFF2C64F2FCC310EF" box="[199,795,947,973]" country="Philippines" location="Mindanao" pageId="7" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Surigao del Sur">
<collectingCountry id="3AAB766BFFD4DD7AFF2C64F2FEB710EF" box="[199,367,947,973]" name="Philippines" pageId="7" pageNumber="320">PHILIPPINES</collectingCountry>
,
<location id="47636020FFD4DD7AFE6964F2FE2E10EF" LSID="urn:lsid:plazi:treatment:CA1587EDFFD4DD75FF7C6578FA44123F:47636020FFD4DD7AFE6964F2FE2E10EF" box="[386,502,947,973]" country="Philippines" name="Mindanao" pageId="7" pageNumber="320" stateProvince="Surigao del Sur">Mindanao</location>
,
<collectingRegion id="8078F819FFD4DD7AFDE264F2FD1310EF" box="[521,715,947,973]" country="Philippines" name="Surigao del Sur" pageId="7" pageNumber="320">Surigao del Sur</collectingRegion>
: •
<specimenCount id="54BAFD72FFD4DD7AFD1C64F2FCC310EF" box="[759,795,947,973]" count="1" pageId="7" pageNumber="320" type="male">1♂</specimenCount>
</materialsCitation>
;
<materialsCitation id="F2D43CA6FFD4DD7AFCDB64F2FDD410D3" country="Philippines" elevation="20" latitude="8.8625" location="Barangay Mat-e" longLatPrecision="1" longitude="126.29889" municipality="Cagwait" pageId="7" pageNumber="320" specimenCount="20">
<collectingMunicipality id="A267AC81FFD4DD7AFCDB64F2FC4810EF" box="[816,912,947,973]" pageId="7" pageNumber="320">Cagwait</collectingMunicipality>
,
<location id="47636020FFD4DD7AFC4864F2FBBC10EF" LSID="urn:lsid:plazi:treatment:CA1587EDFFD4DD75FF7C6578FA44123F:47636020FFD4DD7AFC4864F2FBBC10EF" box="[931,1124,947,973]" country="Philippines" latitude="8.8625" longLatPrecision="1" longitude="126.29889" municipality="Cagwait" name="Barangay Mat-e" pageId="7" pageNumber="320">Barangay Mat-e</location>
; 8°51’45”N, 126°17’56”E [
<geoCoordinate id="2788503CFFD4DD7AFF756497FECB10D3" box="[158,275,982,1009]" degrees="8.86250" direction="north" orientation="latitude" pageId="7" pageNumber="320" precision="1" value="8.8625">N8.86250</geoCoordinate>
,
<geoCoordinate id="2788503CFFD4DD7AFECA6496FE7710D3" box="[289,431,983,1009]" degrees="126.29889" direction="east" orientation="longitude" pageId="7" pageNumber="320" precision="1" value="126.29889">E126.29889</geoCoordinate>
],
<specimenCount id="54BAFD72FFD4DD7AFE2F6496FDD410D3" box="[452,524,983,1009]" count="20" pageId="7" pageNumber="320" type="generic">
<quantity id="85449B1EFFD4DD7AFE2F6496FDDF10D3" box="[452,519,983,1009]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="320" unit="m" value="20.0">
<elevation id="C991D1C8FFD4DD7AFE2F6496FDDF10D3" box="[452,519,983,1009]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="320" unit="m" value="20.0">20 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
<materialsCitation id="F2D43CA6FFD4DD7AFDE76496FB8D10D3" box="[524,1109,983,1009]" collectionCode="ZRC" collectorName="K. C. Grumo &amp; A. A. Gono" country="Philippines" pageId="7" pageNumber="320" specimenCount="1" specimenCount-female="1">
a.s.l.; coll.
<collectorName id="EF49532DFFD4DD7AFD636496FCCF10D3" box="[648,791,983,1009]" pageId="7" pageNumber="320">K.C. Grumo</collectorName>
&amp;
<collectorName id="EF49532DFFD4DD7AFCAB6496FC6710D3" box="[832,959,983,1009]" pageId="7" pageNumber="320">A.A. Gono</collectorName>
(
<collectionCode id="24ADAE3EFFD4DD7AFC3A6496FBD310D3" box="[977,1035,983,1009]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="320">ZRC</collectionCode>
) •
<specimenCount id="54BAFD72FFD4DD7AFBDA6496FB8D10D3" box="[1073,1109,983,1009]" count="1" pageId="7" pageNumber="320" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="F2D43CA6FFD4DD7AFB8D6496FCB51737" country="Philippines" elevation="110" latitude="8.85806" location="Barangay San Roque" longLatPrecision="1" longitude="126.00028" municipality="San Miguel" pageId="7" pageNumber="320" specimenCount="110">
<collectingMunicipality id="A267AC81FFD4DD7AFB8D6496FB3610D3" box="[1126,1262,983,1009]" pageId="7" pageNumber="320">San Miguel</collectingMunicipality>
,
<location id="47636020FFD4DD7AFB166496FF3C1737" LSID="urn:lsid:plazi:treatment:CA1587EDFFD4DD75FF7C6578FA44123F:47636020FFD4DD7AFB166496FF3C1737" country="Philippines" latitude="8.85806" longLatPrecision="1" longitude="126.00028" municipality="San Miguel" name="Barangay San Roque" pageId="7" pageNumber="320">Barangay San Roque</location>
; 8°51’29”N, 126°0’1”E [
<geoCoordinate id="2788503CFFD4DD7AFE1664BBFDAA1737" box="[509,626,1018,1045]" degrees="8.85806" direction="north" orientation="latitude" pageId="7" pageNumber="320" precision="1" value="8.85806">N8.85806</geoCoordinate>
,
<geoCoordinate id="2788503CFFD4DD7AFD9764BAFCD21737" box="[636,778,1019,1045]" degrees="126.00028" direction="east" orientation="longitude" pageId="7" pageNumber="320" precision="1" value="126.00028">E126.00028</geoCoordinate>
],
<specimenCount id="54BAFD72FFD4DD7AFCF764BAFCB51737" box="[796,877,1019,1045]" count="110" pageId="7" pageNumber="320" type="generic">
<quantity id="85449B1EFFD4DD7AFCF764BAFCB01737" box="[796,872,1019,1045]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="320" unit="m" value="110.0">
<elevation id="C991D1C8FFD4DD7AFCF764BAFCB01737" box="[796,872,1019,1045]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="320" unit="m" value="110.0">110 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
<materialsCitation id="F2D43CA6FFD4DD7AFC8664BAFEBE171B" collectingDate="2023-08-30" collectionCode="ZRC" collectorName="K. C. Grumo &amp; A. A. Gono" country="Philippines" pageId="7" pageNumber="320" specimenCount="1">
a.s.l.;
<date id="3602103BFFD4DD7AFC4164BAFB861737" box="[938,1118,1019,1045]" pageId="7" pageNumber="320" value="2023-08-30">
<collectingDate id="2646E9D3FFD4DD7AFC4164BAFB861737" box="[938,1118,1019,1045]" pageId="7" pageNumber="320" value="2023-08-30">30 August 2023</collectingDate>
</date>
, 21h50; coll.
<collectorName id="EF49532DFFD4DD7AFB1E64BAFAA71737" box="[1269,1407,1019,1045]" pageId="7" pageNumber="320">K.C. Grumo</collectorName>
&amp;
<collectorName id="EF49532DFFD4DD7AFF7C635EFECB171B" box="[151,275,1055,1081]" pageId="7" pageNumber="320">A.A. Gono</collectorName>
(
<collectionCode id="24ADAE3EFFD4DD7AFEC9635EFE86171B" box="[290,350,1055,1081]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="320">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD4DD7AFF2C6302FB741683" pageId="7" pageNumber="320" type="discussion">
<paragraph id="420336FBFFD4DD7AFF2C6302FBDD17A3" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C6302FEE3177F" bold="true" box="[199,315,1091,1117]" pageId="7" pageNumber="320">Remarks.</emphasis>
Male previously unknown. Our female from
<collectingRegion id="8078F819FFD4DD7AFCA26302FC15177F" box="[841,973,1091,1117]" country="El Salvador" name="San Miguel" pageId="7" pageNumber="320">San Miguel</collectingRegion>
resembles the
<typeStatus id="9D078859FFD4DD7AFB906302FB38177F" box="[1147,1248,1091,1117]" pageId="7" pageNumber="320" type="holotype">holotype</typeStatus>
, and the general colour patterns of the collected males from nearby also resemble the
<typeStatus id="9D078859FFD4DD7AFC706326FBD917A3" box="[923,1025,1127,1153]" pageId="7" pageNumber="320" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="420336FBFFD4DD7AFF2C63CBFB741683" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C63CBFE781786" bold="true" box="[199,416,1162,1188]" pageId="7" pageNumber="320">Revised diagnosis.</emphasis>
In addition to the diagnosis given in
<bibRefCitation id="262D4B0AFFD4DD7AFCD063CAFC331787" author="Ingrisch, S." box="[827,1003,1162,1189]" pageId="7" pageNumber="320" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch (2015)</bibRefCitation>
: the male of this species is very similar to
<taxonomicName id="85BC4D78FFD4DD7AFF5D63EEFCBF17EB" authority="Tan, Baroga-Barbecho &amp; Yap, 2018" authorityName="Tan, Baroga-Barbecho &amp; Yap" authorityYear="2018" box="[182,871,1199,1225]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="320" phylum="Arthropoda" rank="species" species="furvuseques">
<emphasis id="70C8EAE9FFD4DD7AFF5D63EEFE1E17EB" box="[182,454,1199,1225]" italics="true" pageId="7" pageNumber="320">Anthracites furvuseques</emphasis>
<bibRefCitation id="262D4B0AFFD4DD7AFE2563EEFCBF17EB" author="Tan, M. K. &amp; Ingrisch, S. &amp; Robillard, T. &amp; Baroga-Barbecho, J. B. &amp; Yap, S. A." box="[462,871,1199,1225]" pageId="7" pageNumber="320" pagination="331 - 348" refId="ref9438" refString="Tan, M. K., Ingrisch, S., Robillard, T., Baroga-Barbecho, J. B. &amp; Yap, S. A. (2018) New taxa and notes on spine-headed katydids (Orthoptera: Conocephalinae: Agraeciini) from the Philippines. Zootaxa, 4462 (3), 331 - 348. https: // doi. org / 10.11646 / zootaxa. 4462.3.2" type="journal article" year="2018">Tan, Baroga-Barbecho &amp; Yap, 2018</bibRefCitation>
</taxonomicName>
from Siargao Island in the colour patterns on the legs and the cercus; but differs by the head and pronotum more red brown than black and the posterior femur dorsum not black, the cercus with the two internal teeth long and slender (instead of conical), subgenital plate black with dorsad-pointing lateral processes (instead of cream coloured and without the processes), and the shape of its titillators. The male is also very similar to
<taxonomicName id="85BC4D78FFD4DD7AFD60627EFC54167B" authorityName="Ingrisch" authorityYear="2015" box="[651,908,1343,1369]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="320" phylum="Arthropoda" rank="species" species="flagellatus">
<emphasis id="70C8EAE9FFD4DD7AFD60627EFC54167B" box="[651,908,1343,1369]" italics="true" pageId="7" pageNumber="320">Anthracites flagellatus</emphasis>
</taxonomicName>
from
<collectingRegion id="8078F819FFD4DD7AFC3C627EFB45167B" box="[983,1181,1343,1369]" country="Philippines" name="Misamis Oriental" pageId="7" pageNumber="320">Misamis Oriental</collectingRegion>
in colour patterns (as described in
<bibRefCitation id="262D4B0AFFD4DD7AFECC6222FE16165F" author="Ingrisch, S." box="[295,462,1378,1405]" pageId="7" pageNumber="320" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
); but differs by the subgenital plate having triangular dorsad-pointing lateral processes with subobtuse apices (instead of spine-like with acute apices), and the shape of its titillators.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD4DD75FF2C62EBFDD513DB" lastPageId="8" lastPageNumber="321" pageId="7" pageNumber="320" type="description">
<paragraph id="420336FBFFD4DD7AFF2C62EBFCF516E7" blockId="7.[151,1437,730,2017]" box="[199,813,1450,1477]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C62EBFEAD16E6" bold="true" box="[199,373,1450,1476]" pageId="7" pageNumber="320">Redescription.</emphasis>
Male habitus as shown in
<figureCitation id="DA872A7EFFD4DD7AFD7462EAFD2116E7" box="[671,761,1451,1477]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="7" pageNumber="320">Figs 1B</figureCitation>
,
<figureCitation id="DA872A7EFFD4DD7AFCEE62EAFCF116E7" box="[773,809,1451,1477]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="7" pageNumber="320">2B</figureCitation>
.
</paragraph>
<paragraph id="420336FBFFD4DD7AFF2C628EFC7616CB" blockId="7.[151,1437,730,2017]" box="[199,942,1486,1513]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C628EFED316CB" box="[199,267,1487,1513]" italics="true" pageId="7" pageNumber="320">Head.</emphasis>
Head and pronotum not different from female (
<figureCitation id="DA872A7EFFD4DD7AFCCA628EFC7A16CB" box="[801,930,1486,1513]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Figs 5A–C</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFD4DD7AFF2C62B2FE651577" blockId="7.[151,1437,730,2017]" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C62B2FE68152E" box="[199,432,1523,1549]" italics="true" pageId="7" pageNumber="320">Tegmina and wings.</emphasis>
Tegmen micropterous, mirror area fully covered by pronotal disc (
<figureCitation id="DA872A7EFFD4DD7AFB2D62B2FAFB152F" box="[1222,1315,1522,1549]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Fig. 5A</figureCitation>
), reaching middle of second abdominal tergite, with apex subtruncated; infumated, mostly cream-coloured with numerous black spots (
<figureCitation id="DA872A7EFFD4DD7AFEC8617AFE681577" box="[291,432,1594,1621]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Figs 5A, 5B</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFD4DD75FF2C611EFE7B13F7" blockId="7.[151,1437,730,2017]" lastBlockId="8.[151,1436,151,285]" lastPageId="8" lastPageNumber="321" pageId="7" pageNumber="320">
<emphasis id="70C8EAE9FFD4DD7AFF2C611EFEE1155B" box="[199,313,1631,1657]" italics="true" pageId="7" pageNumber="320">Abdomen.</emphasis>
Tenth abdominal tergite narrows posteriorly, at posterior margin truncated and bent ventrad, at middle emarginated and having setae (
<figureCitation id="DA872A7EFFD4DD7AFE1461C2FD4915BF" box="[511,657,1666,1693]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Figs 5D, 5E</figureCitation>
). Epiproct tongue shaped (
<figureCitation id="DA872A7EFFD4DD7AFC2061C2FBFD15BF" box="[971,1061,1666,1693]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Fig. 5E</figureCitation>
). Cercus very stout, rectangular, distinctly dorso-laterally compressed; externo-distal part slightly conical with apex obtuse; at apex with two inner processes pointing inwards in distal half; dorsal process long and slender, cylindrical but with basal half slightly broader and slightly curved inward, at apex forming small acute tooth; ventral process shorter, slightly surpassing middle of dorso-internal process, more cylindrical, slenderer, at apex subacute (
<figureCitation id="DA872A7EFFD4DD7AFBE36052FB51140F" box="[1032,1161,1810,1837]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Figs 5D–G</figureCitation>
). Subgenital plate longer than wide; with anterior margin concave; gently converging lateral margins; posterior margin narrow and deeply excised, posterior margin between bases of styli terminating into tube-shaped process and almost reaching apex of styli (
<figureCitation id="DA872A7EFFD4DD7AFF3D603EFEF614BB" box="[214,302,1918,1945]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Fig. 5D</figureCitation>
); margin around bases of styli on dorsal side swollen and with triangular internal tooth (
<figureCitation id="DA872A7EFFD4DD7AFB10603EFA8D14BB" box="[1275,1365,1918,1945]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Fig. 5G</figureCitation>
). Styli stout and long with apex rounded (
<figureCitation id="DA872A7EFFD4DD7AFDCC60E2FD72149F" box="[551,682,1954,1981]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="7" pageNumber="320">Figs 5D–G</figureCitation>
). Titillators (
<figureCitation id="DA872A7EFFD4DD7AFCD760E2FC5A149F" box="[828,898,1955,1981]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[190,1382,199,1916]" captionTargetId="figure-17@9.[181,1407,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Anthracites geniculatus Dohrn, 1905 ♂ titillators in dorsal (A), dorso-distal (B, C), dorso-proximal (D), ventral (E), ventro-distal (F) and lateral (G) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13757711" httpUri="https://zenodo.org/record/13757711/files/figure.png" pageId="7" pageNumber="320">Fig. 6</figureCitation>
) separate, with two large and long lateral arms pointing posteriorly; lateral arm in dorsal and ventral views tapers into subacute apex, each with small stout ventral tooth with subacute apex; base typical of genus, at proximal end stout; stiffened along rim and provided with two pairs of lateral sclerites.
</paragraph>
<paragraph id="420336FBFFDBDD75FF2C679EFDD513DB" blockId="8.[151,1436,151,285]" box="[199,525,223,249]" pageId="8" pageNumber="321">
<emphasis id="70C8EAE9FFDBDD75FF2C679EFEA613DB" bold="true" box="[199,382,223,249]" pageId="8" pageNumber="321">Measurements.</emphasis>
See
<tableCitation id="0F3E0340FFDBDD75FE58679EFDDF13DB" box="[435,519,223,249]" captionStart="TABLE 1" captionStartId="22.[151,239,152,176]" captionText="TABLE 1. Measurements of Anthracites, in mm." pageId="8" pageNumber="321">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFDBDD75FF2C6643FA44123F" box="[199,1436,258,285]" pageId="8" pageNumber="321" type="distribution">
<paragraph id="420336FBFFDBDD75FF2C6643FA44123F" blockId="8.[151,1436,151,285]" box="[199,1436,258,285]" pageId="8" pageNumber="321">
<emphasis id="70C8EAE9FFDBDD75FF2C6643FEB9123E" bold="true" box="[199,353,258,284]" pageId="8" pageNumber="321">Distribution.</emphasis>
<collectingRegion id="8078F819FFDBDD75FE8C6642FDC0123F" box="[359,536,259,285]" country="Philippines" name="Agusan del Sur" pageId="8" pageNumber="321">Agusan del Sur</collectingRegion>
: San Francisco;
<collectingRegion id="8078F819FFDBDD75FD3B6642FCA6123F" box="[720,894,259,285]" country="Philippines" name="Surigao del Sur" pageId="8" pageNumber="321">Surigao del Sur</collectingRegion>
[new locality record]:
<collectingRegion id="8078F819FFDBDD75FB956642FAD8123F" box="[1150,1280,259,285]" country="El Salvador" name="San Miguel" pageId="8" pageNumber="321">San Miguel</collectingRegion>
and Cagwait.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="576D97C90E93311C766378A44794DF89">A taxonomic review on the katydids of the genus Anthracites Redtenbacher, 1891 (Tettigoniidae: Conocephalinae: Agraeciini) from Mindanao, Philippines</mods:title>
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<mods:namePart id="E8BB196791EC1A90CB08E6F02BB69982">Sabang, Aira Mae M.</mods:namePart>
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<mods:namePart id="3A4508F06B5D166F40C81C6065EE891E">Bahoy, Daphne Cayle M.</mods:namePart>
<mods:affiliation id="AF4B67D8711E256CCC85F5617CC61BB0">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:namePart id="9BDE7478B9917BF6AE5C4EB99211EA8E">Grumo, Kim C.</mods:namePart>
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<mods:affiliation id="8A1259F9C14DE20EE313AC8C86A8CBD1">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:affiliation id="4976C817E50594D9E690BBC77DD4E385">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:name id="6982FFDC7623E2C6726C2EF63933EBAC" type="personal">
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<mods:affiliation id="05FBE28EC1640BE8C0D7EEC74CB819E4">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:name id="F014EF2F0CC2B7D3B5B43F431E60EA7D" type="personal">
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<mods:affiliation id="A319C0B4EC9074D5994135B71A27708B">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France</mods:affiliation>
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<emphasis id="70C8EAE9FFD5DD7BFF7C604DFE721405" bold="true" box="[151,426,1804,1831]" italics="true" pageId="6" pageNumber="319">Anthracites flagellatus</emphasis>
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<emphasis id="70C8EAE9FFD5DD7BFF7C6010FEA5144B" box="[151,381,1873,1897]" italics="true" pageId="6" pageNumber="319">Anthracites flagellatus</emphasis>
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Type material examined (images and description from
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).
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: •
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[
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SW Gingoog
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], [8°41'N, 124°55'E];
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; coll.
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; (
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).
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<emphasis id="70C8EAE9FFD4DD7AFF2C67D6FE3F1392" bold="true" box="[199,487,150,177]" pageId="7" pageNumber="320">New material examined.</emphasis>
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,
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: •
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;
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.
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a.s.l; in an open area of riparian forest,
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away from the stream, over leaf of the shrub; coll.
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&amp;
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)
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•
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<collectingMunicipality id="A267AC81FFD4DD7AFC5A679EFBCB13DB" box="[945,1043,223,249]" pageId="7" pageNumber="320">Gingoog</collectingMunicipality>
City
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;
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, 09h36; open area of a secondary forest, buffer zone of the foot of the stratovolcano, found in between pandan leaves; coll.
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&amp;
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; KS1 (
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and KS2 (MNHN-EO1355) •
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, 12h41; open area of a secondary forest, buffer zone of the foot of the stratovolcano, found in between pandan leaves; coll.
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&amp;
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; KS17 (
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.
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<emphasis id="70C8EAE9FFD4DD7AFF2C66F7FEB912F2" bold="true" box="[199,353,438,464]" pageId="7" pageNumber="320">Distribution.</emphasis>
<collectingRegion id="8078F819FFD4DD7AFE9B66F6FDE712F3" box="[368,575,439,465]" country="Philippines" name="Misamis Oriental" pageId="7" pageNumber="320">Misamis Oriental</collectingRegion>
: Mount Empagatao, Hindangon, Minalwang (municipality of Claveria), Minubanan, Mount Kibungol; Mount Balatukan [new locality record].
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<figureCitation id="DA872A7EFFD9DD77FEED67FAFEF113F7" box="[262,297,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="10" pageNumber="323">2C</figureCitation>
,
<figureCitation id="DA872A7EFFD9DD77FEDE67FAFEB713F7" box="[309,367,187,213]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="10.[151,250,1945,1969]" captionStartId-1="11.[151,250,1135,1159]" captionStartId-2="12.[151,250,1926,1950]" captionStartId-3="13.[151,250,1924,1948]" captionTargetBox-0="[158,1430,434,1921]" captionTargetBox-1="[172,1414,202,1093]" captionTargetBox-2="[151,1436,181,1902]" captionTargetBox-3="[151,1436,181,1900]" captionTargetId-0="figure-62@10.[158,1430,434,1921]" captionTargetId-1="figure-17@11.[151,1436,181,1112]" captionTargetId-2="figure-19@12.[151,1436,181,1902]" captionTargetId-3="figure-17@13.[151,1436,181,1900]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." captionText-1="FIGURE 8. Anthracites pyramidalis Ingrisch, 2015 ♂ titillators in ventro-distal (A), ventral (B) and dorso-proximal (C) views. Scale bar: 1 mm." captionText-2="FIGURE 9. Anthracites pyramidalis Ingrisch, 2015 males in their natural habitat at Claveria." captionText-3="FIGURE 10. Anthracites pyramidalis Ingrisch, 2015 females in their natural habitat at Claveria." figureDoi-0="http://doi.org/10.5281/zenodo.13757713" figureDoi-1="http://doi.org/10.5281/zenodo.13757715" figureDoi-2="http://doi.org/10.5281/zenodo.13757717" figureDoi-3="http://doi.org/10.5281/zenodo.13757719" httpUri-0="https://zenodo.org/record/13757713/files/figure.png" httpUri-1="https://zenodo.org/record/13757715/files/figure.png" httpUri-2="https://zenodo.org/record/13757717/files/figure.png" httpUri-3="https://zenodo.org/record/13757719/files/figure.png" pageId="10" pageNumber="323">7–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD9DD77FF7C67B8FD921233" box="[151,586,249,273]" pageId="10" pageNumber="323" type="reference_group">
<paragraph id="420336FBFFD9DD77FF7C67B8FD921233" blockId="10.[151,586,249,273]" box="[151,586,249,273]" pageId="10" pageNumber="323">
<treatmentCitationGroup id="62AC11D5FFD9DD77FF7C67B8FD921233" box="[151,586,249,273]" pageId="10" pageNumber="323">
<treatmentCitation id="C31D10EAFFD9DD77FF7C67B8FD921233" author="Ingrisch, S." box="[151,586,249,273]" page="83" pageId="10" pageNumber="323" year="2015">
<taxonomicName id="85BC4D78FFD9DD77FF7C67B8FD921233" ID-CoL="5VGR6" authority="Ingrisch, 2015: 83" authorityName="Ingrisch" authorityPageNumber="83" authorityYear="2015" box="[151,586,249,273]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="323" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFD9DD77FF7C67B8FE531233" box="[151,395,249,273]" italics="true" pageId="10" pageNumber="323">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFD9DD77FE7967B8FD921233" author="Ingrisch, S." box="[402,586,249,273]" pageId="10" pageNumber="323" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015: 83</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFD9DD73FF7C6678FC8912EC" lastPageId="14" lastPageNumber="327" pageId="10" pageNumber="323" type="materials_examined">
<paragraph id="420336FBFFD9DD77FF7C6678FE5F12B9" blockId="10.[151,1436,312,411]" pageId="10" pageNumber="323">
<emphasis id="70C8EAE9FFD9DD77FF7C6678FBC21270" bold="true" box="[151,1050,312,339]" pageId="10" pageNumber="323">
Type material examined (images and description from
<bibRefCitation id="262D4B0AFFD9DD77FCB36679FBD41270" author="Ingrisch, S." box="[856,1036,312,338]" pageId="10" pageNumber="323" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
).
</emphasis>
<materialsCitation id="F2D43CA6FFD9DD77FBC36678FE5F12B9" collectingDate="1967-05-01" collectingDateMax="1967-05-05" collectingDateMin="1967-05-01" collectionCode="BPBM" collectorName="H. M. Torrevillas" country="Philippines" elevation="1500" location="Mount Balatukan" municipality="Mindanao" pageId="10" pageNumber="323" specimenCount="1" specimenCount-female="1" stateProvince="Misamis Oriental" typeStatus="holotype">
<emphasis id="70C8EAE9FFD9DD77FBC36678FB461271" bold="true" box="[1064,1182,313,339]" pageId="10" pageNumber="323">
<typeStatus id="9D078859FFD9DD77FBC36678FB411271" box="[1064,1177,313,339]" pageId="10" pageNumber="323" type="holotype">Holotype</typeStatus>
:
</emphasis>
•
<emphasis id="70C8EAE9FFD9DD77FB2E667BFB021274" bold="true" box="[1221,1242,314,342]" pageId="10" pageNumber="323">
<specimenCount id="54BAFD72FFD9DD77FB2E667BFB021274" box="[1221,1242,314,342]" count="1" pageId="10" pageNumber="323" type="female">♀</specimenCount>
</emphasis>
;
<collectingCountry id="3AAB766BFFD9DD77FB1B6678FA401271" box="[1264,1432,313,339]" name="Philippines" pageId="10" pageNumber="323">PHILIPPINES</collectingCountry>
,
<collectingMunicipality id="A267AC81FFD9DD77FF7C661DFED31254" box="[151,267,348,374]" pageId="10" pageNumber="323">Mindanao</collectingMunicipality>
,
<collectingRegion id="8078F819FFD9DD77FEFC661DFE061254" box="[279,478,348,374]" country="Philippines" name="Misamis Oriental" pageId="10" pageNumber="323">Misamis Oriental</collectingRegion>
,
<location id="47636020FFD9DD77FE02661CFD751254" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD9DD77FE02661CFD751254" box="[489,685,348,375]" country="Philippines" municipality="Mindanao" name="Mount Balatukan" pageId="10" pageNumber="323" stateProvince="Misamis Oriental">Mount Balatukan</location>
[8°46'N, 124°59'E], elev.
<quantity id="85449B1EFFD9DD77FC38661CFBA91255" box="[979,1137,349,375]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="10" pageNumber="323" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">
<elevation id="C991D1C8FFD9DD77FC38661CFBA91255" box="[979,1137,349,375]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="10" pageNumber="323" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">1000–2000 m</elevation>
</quantity>
;
<date id="3602103BFFD9DD77FB96661DFAC61254" box="[1149,1310,348,375]" pageId="10" pageNumber="323" value="1967-05-01" valueMax="1967-05-05" valueMin="1967-05-01">
<collectingDate id="2646E9D3FFD9DD77FB96661DFAC61254" box="[1149,1310,348,375]" pageId="10" pageNumber="323" value="1967-05-01" valueMax="1967-05-05" valueMin="1967-05-01">1–5 May1967</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFD9DD77FA8A661CFECC12B8" pageId="10" pageNumber="323">H.M. Torrevillas</collectorName>
; (
<collectionCode id="24ADAE3EFFD9DD77FECC66C0FEA312B9" box="[295,379,385,411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="10" pageNumber="323" type="Museum">BPBM</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="16C36673FFD9DD77FF7C60D8FC4614D8" ID-DOI="http://doi.org/10.5281/zenodo.13757713" ID-Zenodo-Dep="13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="10" pageNumber="323" startId="10.[151,250,1945,1969]" targetBox="[158,1430,434,1921]" targetPageId="10" targetType="figure">
<paragraph id="420336FBFFD9DD77FF7C60D8FC4614D8" blockId="10.[151,1436,1945,2042]" pageId="10" pageNumber="323">
<emphasis id="70C8EAE9FFD9DD77FF7C60D8FECA1490" bold="true" box="[151,274,1945,1970]" pageId="10" pageNumber="323">FIGURE 7.</emphasis>
<taxonomicName id="85BC4D78FFD9DD77FEF360D8FD7C1493" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[280,676,1945,1969]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="323" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFD9DD77FEF360D8FDD41493" box="[280,524,1945,1969]" italics="true" pageId="10" pageNumber="323">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFD9DD77FDF960D8FD7C1493" author="Ingrisch, S." box="[530,676,1945,1969]" pageId="10" pageNumber="323" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G).
</paragraph>
</caption>
<caption id="16C36673FFD8DD76FF7C632EFE9C1789" ID-DOI="http://doi.org/10.5281/zenodo.13757715" ID-Zenodo-Dep="13757715" httpUri="https://zenodo.org/record/13757715/files/figure.png" pageId="11" pageNumber="324" startId="11.[151,250,1135,1159]" targetBox="[172,1414,202,1093]" targetPageId="11" targetType="figure">
<paragraph id="420336FBFFD8DD76FF7C632EFE9C1789" blockId="11.[151,1437,1135,1196]" pageId="11" pageNumber="324">
<emphasis id="70C8EAE9FFD8DD76FF7C632EFECA17A5" bold="true" box="[151,274,1135,1159]" pageId="11" pageNumber="324">FIGURE 8.</emphasis>
<taxonomicName id="85BC4D78FFD8DD76FEF36331FD7C17A5" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[280,676,1135,1160]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="324" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFD8DD76FEF36331FDD417AA" box="[280,524,1136,1160]" italics="true" pageId="11" pageNumber="324">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFD8DD76FDF96331FD7C17A5" author="Ingrisch, S." box="[530,676,1135,1160]" pageId="11" pageNumber="324" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
♂ titillators in ventro-distal (A), ventral (B) and dorso-proximal (C) views. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="420336FBFFD8DD73FF2C63B1FE491241" blockId="11.[151,1437,1264,2041]" lastBlockId="14.[151,1437,151,1562]" lastPageId="14" lastPageNumber="327" pageId="11" pageNumber="324">
<emphasis id="70C8EAE9FFD8DD76FF2C63B1FE3F1628" bold="true" box="[199,487,1264,1290]" pageId="11" pageNumber="324">New material examined.</emphasis>
<materialsCitation id="F2D43CA6FFD8DD76FE0663B1FC3C1628" box="[493,996,1264,1290]" country="Philippines" location="Mindanao" pageId="11" pageNumber="324" specimenCount="1" stateProvince="Misamis Oriental">
<collectingCountry id="3AAB766BFFD8DD76FE0663B1FD4D1628" box="[493,661,1264,1290]" name="Philippines" pageId="11" pageNumber="324">PHILIPPINES</collectingCountry>
,
<location id="47636020FFD8DD76FD7463B1FCCB1628" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FD7463B1FCCB1628" box="[671,787,1264,1290]" country="Philippines" name="Mindanao" pageId="11" pageNumber="324" stateProvince="Misamis Oriental">Mindanao</location>
,
<collectingRegion id="8078F819FFD8DD76FCF663B1FC3C1628" box="[797,996,1264,1290]" country="Philippines" name="Misamis Oriental" pageId="11" pageNumber="324">Misamis Oriental</collectingRegion>
</materialsCitation>
: •
<materialsCitation id="F2D43CA6FFD8DD76FC1663B1FA54160C" collectingDate="2023-02-13" country="Philippines" elevation="6" latitude="8.70113" location="Mount Sumagaya" longLatPrecision="1" longitude="125.01298" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FC1663B1FBF91628" box="[1021,1057,1264,1290]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FBC663B1FB571628" box="[1069,1167,1264,1290]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, near
<location id="47636020FFD8DD76FB2463B0FA401628" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FB2463B0FA401628" box="[1231,1432,1264,1291]" country="Philippines" latitude="8.70113" longLatPrecision="1" longitude="125.01298" municipality="Claveria" name="Mount Sumagaya" pageId="11" pageNumber="324">Mount Sumagaya</location>
;
<geoCoordinate id="2788503CFFD8DD76FF7C6255FED1160C" box="[151,265,1300,1326]" degrees="8.70113" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.70113">N8.70113</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FEF86255FE46160D" box="[275,414,1300,1327]" degrees="125.01298" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.01298">E125.01298</geoCoordinate>
, 1259.0±
<quantity id="85449B1EFFD8DD76FDEE6255FD93160C" box="[517,587,1300,1327]" metricMagnitude="0" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="324" unit="m" value="5.8">
<elevation id="C991D1C8FFD8DD76FDEE6255FD93160C" box="[517,587,1300,1327]" metricMagnitude="0" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="324" unit="m" value="5.8">5.8 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FD666255FC8E160C" box="[653,854,1300,1326]" pageId="11" pageNumber="324" value="2023-02-13">
<collectingDate id="2646E9D3FFD8DD76FD666255FC8E160C" box="[653,854,1300,1326]" pageId="11" pageNumber="324" value="2023-02-13">13 February 2023</collectingDate>
</date>
, 19h56; mossy forest; coll. local guide; MIN.23.11
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FF7C6279FC951654" collectingDate="2023-02-13" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" latitude="8.70109" location="Mount Sumagaya" longLatPrecision="1" longitude="125.01304" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-female="1">
<specimenCount id="54BAFD72FFD8DD76FF7C6279FF631670" box="[151,187,1336,1362]" count="1" pageId="11" pageNumber="324" type="female">1♀</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FF236279FEF21670" box="[200,298,1336,1362]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, near
<location id="47636020FFD8DD76FE826279FDEA1670" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FE826279FDEA1670" box="[361,562,1336,1362]" country="Philippines" latitude="8.70109" longLatPrecision="1" longitude="125.01304" municipality="Claveria" name="Mount Sumagaya" pageId="11" pageNumber="324">Mount Sumagaya</location>
;
<geoCoordinate id="2788503CFFD8DD76FDD76279FD751670" box="[572,685,1336,1362]" degrees="8.70109" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.70109">N8.70109</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FD536276FC9B1670" box="[696,835,1335,1362]" degrees="125.01304" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.01304">E125.01304</geoCoordinate>
, 1258.0±5.0 m.a.s.l.;
<date id="3602103BFFD8DD76FBDB6279FB2F1670" box="[1072,1271,1336,1362]" pageId="11" pageNumber="324" value="2023-02-13">
<collectingDate id="2646E9D3FFD8DD76FBDB6279FB2F1670" box="[1072,1271,1336,1362]" pageId="11" pageNumber="324" value="2023-02-13">13 February 2023</collectingDate>
</date>
, 20h00; mossy forest, on foliage of shrub; coll.
<collectorName id="EF49532DFFD8DD76FDEF621DFDAB1654" box="[516,627,1372,1398]" pageId="11" pageNumber="324">M.K. Tan</collectorName>
; MIN.23.14 (
<collectionCode id="24ADAE3EFFD8DD76FCE0621DFC9D1654" box="[779,837,1372,1398]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="324">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FC8E621DFDEE16BB" collectingDate="2023-02-13" collectorName="J. A. Salvador" country="Philippines" location="Mount Sumagaya" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FC8E621DFC521654" box="[869,906,1372,1398]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FC73621AFC221657" box="[920,1018,1371,1397]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, near
<location id="47636020FFD8DD76FBD6621DFADF1654" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FBD6621DFADF1654" box="[1085,1287,1372,1398]" country="Philippines" municipality="Claveria" name="Mount Sumagaya" pageId="11" pageNumber="324">Mount Sumagaya</location>
;
<date id="3602103BFFD8DD76FAF8621DFF0A16BB" pageId="11" pageNumber="324" value="2023-02-13">
<collectingDate id="2646E9D3FFD8DD76FAF8621DFF0A16BB" pageId="11" pageNumber="324" value="2023-02-13">13 February 2023</collectingDate>
</date>
; coll.
<collectorName id="EF49532DFFD8DD76FEFF623EFE7716BB" box="[276,431,1407,1433]" pageId="11" pageNumber="324">J.A. Salvador</collectorName>
; MIN.23.19
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FDA7623EFA54169F" collectingDate="2023-02-15" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" elevation="7" latitude="8.70567" location="Mount Balutukan" longLatPrecision="1" longitude="125.00653" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-female="1">
<specimenCount id="54BAFD72FFD8DD76FDA7623EFDA816BB" box="[588,624,1407,1433]" count="1" pageId="11" pageNumber="324" type="female">1♀</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FD96623EFD0716BB" box="[637,735,1407,1433]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, foot of
<location id="47636020FFD8DD76FCD262C1FBD916BB" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FCD262C1FBD916BB" box="[825,1025,1407,1434]" country="Philippines" latitude="8.70567" longLatPrecision="1" longitude="125.00653" municipality="Claveria" name="Mount Balutukan" pageId="11" pageNumber="324">Mount Balutukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FBE0623EFBA516B8" box="[1035,1149,1407,1434]" degrees="8.70567" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.70567">N8.70567</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FB63623EFACB16B8" box="[1160,1299,1407,1434]" degrees="125.00653" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.00653">E125.00653</geoCoordinate>
, 1208.0±
<quantity id="85449B1EFFD8DD76FA91623EFF6C169F" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="324" unit="m" value="6.5">
<elevation id="C991D1C8FFD8DD76FA91623EFF6C169F" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="324" unit="m" value="6.5">6.5 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FF1C62E2FE19169F" box="[247,449,1443,1470]" pageId="11" pageNumber="324" value="2023-02-15">
<collectingDate id="2646E9D3FFD8DD76FF1C62E2FE19169F" box="[247,449,1443,1470]" pageId="11" pageNumber="324" value="2023-02-15">15 February 2023</collectingDate>
</date>
, 20h; open area outside mossy forest, on shrub leaf; coll.
<collectorName id="EF49532DFFD8DD76FBAE62E2FB6C169F" box="[1093,1204,1443,1469]" pageId="11" pageNumber="324">M.K. Tan</collectorName>
; MIN.23.53 (
<collectionCode id="24ADAE3EFFD8DD76FAA162E2FA5C169F" box="[1354,1412,1443,1469]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="324">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FF7C6286FC031526" collectingDate="2023-02-15" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" latitude="8.7045" location="Mount Balutukan" longLatPrecision="1" longitude="125.00521" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FF7C6286FF6416C3" box="[151,188,1479,1505]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FF2C6286FEF116C3" box="[199,297,1479,1505]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, foot of
<location id="47636020FFD8DD76FE6A6286FD9016C3" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FE6A6286FD9016C3" box="[385,584,1479,1505]" country="Philippines" latitude="8.7045" longLatPrecision="1" longitude="125.00521" municipality="Claveria" name="Mount Balutukan" pageId="11" pageNumber="324">Mount Balutukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FDBA6287FD1C16C3" box="[593,708,1478,1505]" degrees="8.70450" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.7045">N8.70450</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FD266287FC8F16C3" box="[717,855,1478,1505]" degrees="125.00521" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.00521">E125.00521</geoCoordinate>
, 1197.0±6.0 m.a.s.l.;
<date id="3602103BFFD8DD76FBA86287FAD116C3" box="[1091,1289,1478,1505]" pageId="11" pageNumber="324" value="2023-02-15">
<collectingDate id="2646E9D3FFD8DD76FBA86287FAD116C3" box="[1091,1289,1478,1505]" pageId="11" pageNumber="324" value="2023-02-15">15 February 2023</collectingDate>
</date>
, 21h06; open area outside mossy forest, on fern leaf; coll.
<collectorName id="EF49532DFFD8DD76FD7A62AAFCD91527" box="[657,769,1515,1541]" pageId="11" pageNumber="324">M.K. Tan</collectorName>
; MIN.23.58 (
<collectionCode id="24ADAE3EFFD8DD76FC7262ABFC0B1526" box="[921,979,1514,1540]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="324">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FC1F62ABFEBB156E" collectingDate="2023-02-16" collectionCode="ZRC" collectorName="M. K. Tan" country="Philippines" elevation="6" latitude="8.70142" location="Mount Sumagaya" longLatPrecision="1" longitude="125.00961" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FC1F62ABFBC11526" box="[1012,1049,1514,1540]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FBCC62ABFB511526" box="[1063,1161,1514,1540]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
, near
<location id="47636020FFD8DD76FB2662AAFA401526" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FB2662AAFA401526" box="[1229,1432,1514,1541]" country="Philippines" latitude="8.70142" longLatPrecision="1" longitude="125.00961" municipality="Claveria" name="Mount Sumagaya" pageId="11" pageNumber="324">Mount Sumagaya</location>
;
<geoCoordinate id="2788503CFFD8DD76FF7C614FFED2150A" box="[151,266,1550,1576]" degrees="8.70142" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.70142">N8.70142</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FEFC614FFE79150B" box="[279,417,1550,1577]" degrees="125.00961" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.00961">E125.00961</geoCoordinate>
, 1194.0±
<quantity id="85449B1EFFD8DD76FDE0614FFD8B150A" box="[523,595,1550,1576]" metricMagnitude="0" metricUnit="m" metricValue="6.3" pageId="11" pageNumber="324" unit="m" value="6.3">
<elevation id="C991D1C8FFD8DD76FDE0614FFD8B150A" box="[523,595,1550,1576]" metricMagnitude="0" metricUnit="m" metricValue="6.3" pageId="11" pageNumber="324" unit="m" value="6.3">6.3 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FD73614FFCBE150A" box="[664,870,1550,1576]" pageId="11" pageNumber="324" value="2023-02-16">
<collectingDate id="2646E9D3FFD8DD76FD73614FFCBE150A" box="[664,870,1550,1576]" pageId="11" pageNumber="324" value="2023-02-16">16 February 2023</collectingDate>
</date>
, 19h30; mossy forest edge; coll.
<collectorName id="EF49532DFFD8DD76FB0F614EFA8A150B" box="[1252,1362,1551,1577]" pageId="11" pageNumber="324">M.K. Tan</collectorName>
<emphasis id="70C8EAE9FFD8DD76FAB1614EFA4D150A" box="[1370,1429,1550,1576]" italics="true" pageId="11" pageNumber="324">et al.</emphasis>
; MIN.23.64 (
<collectionCode id="24ADAE3EFFD8DD76FEC96173FE84156E" box="[290,348,1586,1612]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="324">ZRC</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FE926173FA831552" collectingDate="2023-07-07" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" elevation="1133" latitude="8.739639" location="Mount Balatukan" longLatPrecision="1" longitude="125.0291" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FE926173FE47156E" box="[377,415,1586,1612]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
: buffer zone of the foot of
<location id="47636020FFD8DD76FD2D6173FC54156E" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FD2D6173FC54156E" box="[710,908,1586,1612]" country="Philippines" latitude="8.739639" longLatPrecision="1" longitude="125.0291" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FC7C6173FBCF156E" box="[919,1047,1586,1612]" degrees="8.739639" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.739639">N8.739639</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FBCA6173FB62156F" box="[1057,1210,1586,1613]" degrees="125.029097" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.0291">E125.029097</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FB2F6173FAC7156E" box="[1220,1311,1586,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.133" pageId="11" pageNumber="324" unit="m" value="1133.0">
<elevation id="C991D1C8FFD8DD76FB2F6173FAC7156E" box="[1220,1311,1586,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.133" pageId="11" pageNumber="324" unit="m" value="1133.0">1133 m</elevation>
</quantity>
.a.s.l;
<date id="3602103BFFD8DD76FAB26173FF0A1552" pageId="11" pageNumber="324" value="2023-07-07">
<collectingDate id="2646E9D3FFD8DD76FAB26173FF0A1552" pageId="11" pageNumber="324" value="2023-07-07">7 July 2023</collectingDate>
</date>
; open area in secondary forest, along the trail, on shrub leaf; coll.
<collectorName id="EF49532DFFD8DD76FC446117FBEC154D" box="[943,1076,1621,1648]" pageId="11" pageNumber="324">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FBBE6117FB34154D" box="[1109,1260,1621,1648]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; 5 (
<collectionCode id="24ADAE3EFFD8DD76FAF86117FA8B1552" box="[1299,1363,1622,1648]" country="Philippines" name="Philippine National Museum" pageId="11" pageNumber="324">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FA9A6117FB991595" collectingDate="2023-07-10" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" elevation="1441" latitude="8.743741" location="Mount Balatukan" longLatPrecision="1" longitude="125.019615" municipality="Mount Balatukan" pageId="11" pageNumber="324" specimenCount="1" specimenCount-female="1">
<specimenCount id="54BAFD72FFD8DD76FA9A6117FA4E1552" box="[1393,1430,1622,1648]" count="1" pageId="11" pageNumber="324" type="female">1♀</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FF7C613BFE8715B1" box="[151,351,1657,1684]" pageId="11" pageNumber="324">Mount Balatukan</collectingMunicipality>
;
<geoCoordinate id="2788503CFFD8DD76FE806138FE3215B1" box="[363,490,1657,1683]" degrees="8.743741" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.743741">N8.743741</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FE1C6138FD4815B6" box="[503,656,1657,1684]" degrees="125.019616" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.019615">E125.019616</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FD776138FD2015B1" box="[668,760,1657,1683]" metricMagnitude="3" metricUnit="m" metricValue="1.441" pageId="11" pageNumber="324" unit="m" value="1441.0">
<elevation id="C991D1C8FFD8DD76FD776138FD2015B1" box="[668,760,1657,1683]" metricMagnitude="3" metricUnit="m" metricValue="1.441" pageId="11" pageNumber="324" unit="m" value="1441.0">1441 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FCD76138FC0915B1" box="[828,977,1657,1683]" pageId="11" pageNumber="324" value="2023-07-10">
<collectingDate id="2646E9D3FFD8DD76FCD76138FC0915B1" box="[828,977,1657,1683]" pageId="11" pageNumber="324" value="2023-07-10">10 July 2023</collectingDate>
</date>
; on an open area inside the mossy forest of
<location id="47636020FFD8DD76FF5D61DFFEA41595" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FF5D61DFFEA41595" box="[182,380,1693,1720]" country="Philippines" latitude="8.743741" longLatPrecision="1" longitude="125.019615" municipality="Mount Balatukan" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
, on a shrub leaf; coll.
<collectorName id="EF49532DFFD8DD76FD9E61DCFD2B1595" box="[629,755,1693,1719]" pageId="11" pageNumber="324">D.C.Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FCFD61DCFC761595" box="[790,942,1693,1719]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; K15 (
<collectionCode id="24ADAE3EFFD8DD76FC1261DCFBE11595" box="[1017,1081,1693,1719]" country="Philippines" name="Philippine National Museum" pageId="11" pageNumber="324">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FBB261DCFA8015DD" collectingDate="2024-03-17" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Gingoog City" elevation="1200" latitude="8.4417" location="Mount Balatukan" longLatPrecision="7" longitude="125.0129" municipality="Barangay Kalagonoy" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FBB261DCFBA71595" box="[1113,1151,1693,1719]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
:
<collectingCounty id="AB624E77FFD8DD76FB6061DCFAFD1595" box="[1163,1317,1693,1719]" pageId="11" pageNumber="324">Gingoog City</collectingCounty>
,
<collectingMunicipality id="A267AC81FFD8DD76FADA61DFFECB15F9" pageId="11" pageNumber="324">Barangay Kalagonoy</collectingMunicipality>
,
<location id="47636020FFD8DD76FECA6180FE3215F9" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FECA6180FE3215F9" box="[289,490,1729,1755]" country="Philippines" county="Gingoog City" latitude="8.4417" longLatPrecision="7" longitude="125.0129" municipality="Barangay Kalagonoy" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FE136180FD8315F9" box="[504,603,1729,1755]" degrees="8.4417" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="5" value="8.4417">N8.4417</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FD826180FD3F15FE" box="[617,743,1729,1756]" degrees="125.0129" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="5" value="125.0129">E125.0129</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FD1F6180FC8A15F9" box="[756,850,1729,1755]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="324" unit="m" value="1200.0">
<elevation id="C991D1C8FFD8DD76FD1F6180FC8A15F9" box="[756,850,1729,1755]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="324" unit="m" value="1200.0">1200 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FC7C6180FB9215F9" box="[919,1098,1729,1755]" pageId="11" pageNumber="324" value="2024-03-17">
<collectingDate id="2646E9D3FFD8DD76FC7C6180FB9215F9" box="[919,1098,1729,1755]" pageId="11" pageNumber="324" value="2024-03-17">17 March 2024</collectingDate>
</date>
, 09h50; along the trail of the buffer zone of
<location id="47636020FFD8DD76FEAB61A4FDDE15DC" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FEAB61A4FDDE15DC" box="[320,518,1764,1791]" country="Philippines" county="Gingoog City" latitude="8.4417" longLatPrecision="7" longitude="125.0129" municipality="Barangay Kalagonoy" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
, in between pandan leaves; coll.
<collectorName id="EF49532DFFD8DD76FC9561A4FBDC15DC" box="[894,1028,1764,1791]" pageId="11" pageNumber="324">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FBC261A4FB1A15DC" box="[1065,1218,1764,1791]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; KS3 (
<collectionCode id="24ADAE3EFFD8DD76FAFB61A4FA8815DD" box="[1296,1360,1765,1791]" country="Philippines" name="Philippine National Museum" pageId="11" pageNumber="324">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FA9A61A4FBEF1464" collectionCode="PNM" collectorName="March &amp; D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Gingoog City" elevation="1223" latitude="8.44168" location="Mount Balatukan" longLatPrecision="1" longitude="125.01279" municipality="Barangay Kalagonoy" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FA9A61A4FA4F15DD" box="[1393,1431,1765,1791]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
:
<collectingCounty id="AB624E77FFD8DD76FF7C6049FEEC1400" box="[151,308,1800,1826]" pageId="11" pageNumber="324">Gingoog City</collectingCounty>
,
<collectingMunicipality id="A267AC81FFD8DD76FEA96048FDEB1400" box="[322,563,1800,1827]" pageId="11" pageNumber="324">Barangay Kalagonoy</collectingMunicipality>
,
<location id="47636020FFD8DD76FDAA6048FCD31400" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FDAA6048FCD31400" box="[577,779,1800,1827]" country="Philippines" county="Gingoog City" latitude="8.44168" longLatPrecision="1" longitude="125.01279" municipality="Barangay Kalagonoy" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FCF26049FC521400" box="[793,906,1800,1826]" degrees="8.44168" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.44168">N8.44168</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FC726049FBFC1401" box="[921,1060,1800,1827]" degrees="125.01279" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.01279">E125.01279</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FBD96049FB481400" box="[1074,1168,1800,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.223" pageId="11" pageNumber="324" unit="m" value="1223.0">
<elevation id="C991D1C8FFD8DD76FBD96049FB481400" box="[1074,1168,1800,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.223" pageId="11" pageNumber="324" unit="m" value="1223.0">1223 m</elevation>
</quantity>
.a.s.l.; 17
<collectorName id="EF49532DFFD8DD76FB176049FA9D1400" box="[1276,1349,1800,1826]" pageId="11" pageNumber="324">March</collectorName>
20h24; found in between pandan leaves; coll.
<collectorName id="EF49532DFFD8DD76FDB8606DFD021464" box="[595,730,1836,1862]" pageId="11" pageNumber="324">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FCEA606DFC431464" box="[769,923,1836,1862]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; KB9 (
<collectionCode id="24ADAE3EFFD8DD76FC04606DFBF71464" box="[1007,1071,1836,1862]" country="Philippines" name="Philippine National Museum" pageId="11" pageNumber="324">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FBB8606DFB5C14AC" collectionCode="PNM" collectorName="July &amp; D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Gingoog City" elevation="1460" latitude="8.743797" location="Mount Balatukan" longLatPrecision="1" longitude="125.01893" municipality="Barangay Kalagonoy" pageId="11" pageNumber="324" specimenCount="1" specimenCount-female="1">
<specimenCount id="54BAFD72FFD8DD76FBB8606DFBA01464" box="[1107,1144,1836,1862]" count="1" pageId="11" pageNumber="324" type="female">1♀</specimenCount>
:
<collectingCounty id="AB624E77FFD8DD76FB6C606DFAFB1464" box="[1159,1315,1836,1862]" pageId="11" pageNumber="324">Gingoog City</collectingCounty>
,
<collectingMunicipality id="A267AC81FFD8DD76FADA606DFECB1448" pageId="11" pageNumber="324">Barangay Kalagonoy</collectingMunicipality>
,
<location id="47636020FFD8DD76FEF56011FE3D1448" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FEF56011FE3D1448" box="[286,485,1872,1898]" country="Philippines" county="Gingoog City" latitude="8.743797" longLatPrecision="1" longitude="125.01893" municipality="Barangay Kalagonoy" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FE1B6011FDA91448" box="[496,625,1872,1898]" degrees="8.743797" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.743797">N8.743797</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FD90600EFCCC1448" box="[635,788,1871,1898]" degrees="125.018932" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.01893">E125.018932</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FCF46011FCA31448" box="[799,891,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="11" pageNumber="324" unit="m" value="1460.0">
<elevation id="C991D1C8FFD8DD76FCF46011FCA31448" box="[799,891,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="11" pageNumber="324" unit="m" value="1460.0">1460 m</elevation>
</quantity>
.a.s.l.; 10
<collectorName id="EF49532DFFD8DD76FC0A6011FBC81448" box="[993,1040,1872,1898]" pageId="11" pageNumber="324">July</collectorName>
20h23; along the trail of the mossy forest of
<location id="47636020FFD8DD76FF176035FE1A14AF" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FF176035FE1A14AF" box="[252,450,1907,1934]" country="Philippines" county="Gingoog City" latitude="8.743797" longLatPrecision="1" longitude="125.01893" municipality="Barangay Kalagonoy" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
, found on a fern; coll.
<collectorName id="EF49532DFFD8DD76FD2B6035FC9D14AF" box="[704,837,1907,1934]" pageId="11" pageNumber="324">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FC8C6035FC2714AF" box="[871,1023,1907,1934]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; K6 (
<collectionCode id="24ADAE3EFFD8DD76FBD06035FBA314AC" box="[1083,1147,1908,1934]" country="Philippines" name="Philippine National Museum" pageId="11" pageNumber="324">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD76FB706035FB7D14F7" collectingDate="2024-03-24" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Gingoog City" elevation="1310" latitude="8.4334" location="Mount Balatukan" longLatPrecision="54" longitude="125.016" municipality="Jamboree Site" pageId="11" pageNumber="324" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FB706035FB6614AC" box="[1179,1214,1908,1934]" count="1" pageId="11" pageNumber="324" type="female">1♀</specimenCount>
,
<specimenCount id="54BAFD72FFD8DD76FB276035FB2914AC" box="[1228,1265,1908,1934]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingCounty id="AB624E77FFD8DD76FB156032FA4014AF" box="[1278,1432,1907,1933]" pageId="11" pageNumber="324">Gingoog City</collectingCounty>
,
<collectingMunicipality id="A267AC81FFD8DD76FF7C60D6FEE11493" box="[151,313,1943,1969]" pageId="11" pageNumber="324">Jamboree Site</collectingMunicipality>
,
<location id="47636020FFD8DD76FEAC60D9FDD71493" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FEAC60D9FDD71493" box="[327,527,1943,1970]" country="Philippines" county="Gingoog City" latitude="8.4334" longLatPrecision="54" longitude="125.016" municipality="Jamboree Site" name="Mount Balatukan" pageId="11" pageNumber="324">Mount Balatukan</location>
;
<geoCoordinate id="2788503CFFD8DD76FDF760D6FD581493" box="[540,640,1943,1969]" degrees="8.4334" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="5" value="8.4334">N8.4334</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FD6660D6FD231490" box="[653,763,1943,1970]" degrees="125.016" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="55" value="125.016">E125.016</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FCE260D6FCBF1493" box="[777,871,1943,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="11" pageNumber="324" unit="m" value="1310.0">
<elevation id="C991D1C8FFD8DD76FCE260D6FCBF1493" box="[777,871,1943,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="11" pageNumber="324" unit="m" value="1310.0">1310 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FC4060D6FB841493" box="[939,1116,1943,1969]" pageId="11" pageNumber="324" value="2024-03-24">
<collectingDate id="2646E9D3FFD8DD76FC4060D6FB841493" box="[939,1116,1943,1969]" pageId="11" pageNumber="324" value="2024-03-24">24 March 2024</collectingDate>
</date>
, 09h25; along the campsite, found on leaf of the shrub, coll.
<collectorName id="EF49532DFFD8DD76FE0460FAFDA814F7" box="[495,624,1979,2005]" pageId="11" pageNumber="324">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFD8DD76FD6460FAFCFA14F7" box="[655,802,1979,2005]" pageId="11" pageNumber="324">A.M. Sabang</collectorName>
; JB11/J13 and JB8/J14 (both PNM)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFD8DD73FB5160FAFAFD1393" collectingDate="2024-04-22" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" elevation="1232" lastPageId="14" lastPageNumber="327" latitude="8.697233" location="Mount Sumagaya-Lumot" longLatPrecision="1" longitude="125.02436" municipality="Claveria" pageId="11" pageNumber="324" specimenCount="1" specimenCount-male="1">
<specimenCount id="54BAFD72FFD8DD76FB5160FAFB0714F7" box="[1210,1247,1979,2005]" count="1" pageId="11" pageNumber="324" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFD8DD76FB0160FAFA9314F7" box="[1258,1355,1979,2005]" pageId="11" pageNumber="324">Claveria</collectingMunicipality>
; foot of
<location id="47636020FFD8DD76FF7C609EFE6814DB" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFD8DD76FF7C609EFE6814DB" box="[151,432,2015,2041]" country="Philippines" latitude="8.697233" longLatPrecision="1" longitude="125.02436" municipality="Claveria" name="Mount Sumagaya-Lumot" pageId="11" pageNumber="324">Mount Sumagaya-Lumot</location>
;
<geoCoordinate id="2788503CFFD8DD76FE57609FFDE214DA" box="[444,570,2014,2040]" degrees="8.697233" direction="north" orientation="latitude" pageId="11" pageNumber="324" precision="1" value="8.697233">N8.697233</geoCoordinate>
,
<geoCoordinate id="2788503CFFD8DD76FDAE609FFD0514DB" box="[581,733,2014,2041]" degrees="125.024364" direction="east" orientation="longitude" pageId="11" pageNumber="324" precision="1" value="125.02436">E125.024364</geoCoordinate>
,
<quantity id="85449B1EFFD8DD76FD03609EFC9C14DB" box="[744,836,2015,2041]" metricMagnitude="3" metricUnit="m" metricValue="1.232" pageId="11" pageNumber="324" unit="m" value="1232.0">
<elevation id="C991D1C8FFD8DD76FD03609EFC9C14DB" box="[744,836,2015,2041]" metricMagnitude="3" metricUnit="m" metricValue="1.232" pageId="11" pageNumber="324" unit="m" value="1232.0">1232 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFD8DD76FC6C609EFBFD14DB" box="[903,1061,2015,2041]" pageId="11" pageNumber="324" value="2024-04-22">
<collectingDate id="2646E9D3FFD8DD76FC6C609EFBFD14DB" box="[903,1061,2015,2041]" pageId="11" pageNumber="324" value="2024-04-22">22 April 2024</collectingDate>
</date>
, 21h18; open area in a secondary forest,
<quantity id="85449B1EFFDDDD73FF0D67D6FEF71393" box="[230,303,151,177]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="327" unit="m" value="130.0">
<elevation id="C991D1C8FFDDDD73FF0D67D6FEF71393" box="[230,303,151,177]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="327" unit="m" value="130.0">130 m</elevation>
</quantity>
from a river, found on the leaf of a shrub; coll.
<collectorName id="EF49532DFFDDDD73FCBF67D6FC021393" box="[852,986,151,177]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FC1467D6FB401393" box="[1023,1176,151,177]" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; JS4 (
<collectionCode id="24ADAE3EFFDDDD73FB3667D6FAC51393" box="[1245,1309,151,177]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFDDDD73FAD467D6FF3F123E" collectingDate="2024-04-23" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" elevation="1253" latitude="8.69605" location="Mount Sumagaya-Lumot" longLatPrecision="1" longitude="125.025444" municipality="Claveria" pageId="14" pageNumber="327" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="54BAFD72FFDDDD73FAD467D6FABD1393" box="[1343,1381,151,177]" count="2" pageId="14" pageNumber="327" type="male">2♂</specimenCount>
,
<specimenCount id="54BAFD72FFDDDD73FA9A67D6FA4E1393" box="[1393,1430,151,177]" count="1" pageId="14" pageNumber="327" type="female">1♀</specimenCount>
;
<collectingMunicipality id="A267AC81FFDDDD73FF7C67FAFF2113F7" box="[151,249,187,213]" pageId="14" pageNumber="327">Claveria</collectingMunicipality>
; foot of
<location id="47636020FFDDDD73FE8B67FAFD5913F7" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFDDDD73FE8B67FAFD5913F7" box="[352,641,187,213]" country="Philippines" latitude="8.69605" longLatPrecision="1" longitude="125.025444" municipality="Claveria" name="Mount Sumagaya-Lumot" pageId="14" pageNumber="327">Mount Sumagaya-Lumot</location>
;
<geoCoordinate id="2788503CFFDDDD73FD6467FBFCD913F7" box="[655,769,186,213]" degrees="8.69605" direction="north" orientation="latitude" pageId="14" pageNumber="327" precision="1" value="8.69605">N8.69605</geoCoordinate>
,
<geoCoordinate id="2788503CFFDDDD73FCFB67FBFC7113F7" box="[784,937,186,213]" degrees="125.025447" direction="east" orientation="longitude" pageId="14" pageNumber="327" precision="1" value="125.025444">E125.025447</geoCoordinate>
,
<quantity id="85449B1EFFDDDD73FC5C67FBFBCE13F7" box="[951,1046,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.253" pageId="14" pageNumber="327" unit="m" value="1253.0">
<elevation id="C991D1C8FFDDDD73FC5C67FBFBCE13F7" box="[951,1046,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.253" pageId="14" pageNumber="327" unit="m" value="1253.0">1253 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFDDDD73FBB067FAFAD813F7" box="[1115,1280,187,213]" pageId="14" pageNumber="327" value="2024-04-23">
<collectingDate id="2646E9D3FFDDDD73FBB067FAFAD813F7" box="[1115,1280,187,213]" pageId="14" pageNumber="327" value="2024-04-23">23 April 2024</collectingDate>
</date>
, 20h15; open area in a secondary forest,
<quantity id="85449B1EFFDDDD73FE2F679FFE2513DB" box="[452,509,222,249]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="327" unit="m" value="45.0">
<elevation id="C991D1C8FFDDDD73FE2F679FFE2513DB" box="[452,509,222,249]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="327" unit="m" value="45.0">45 m</elevation>
</quantity>
from a river, found on the leaf of a shrub; coll.
<collectorName id="EF49532DFFDDDD73FBF8679EFB4013DB" box="[1043,1176,223,249]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FB51679EFA8B13DB" box="[1210,1363,223,249]" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; SB24 (
<collectionCode id="24ADAE3EFFDDDD73FF746643FF07123E" box="[159,223,258,284]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFDDDD73FEEF6643FE551241" collectingDate="2024-04-25" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" elevation="1172" latitude="8.700011" location="Mount Sumagaya-Lumot" longLatPrecision="1" longitude="125.005775" municipality="Claveria" pageId="14" pageNumber="327" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="54BAFD72FFDDDD73FEEF6643FEF2123E" box="[260,298,258,284]" count="2" pageId="14" pageNumber="327" type="male">2♂</specimenCount>
,
<specimenCount id="54BAFD72FFDDDD73FED36643FE84123E" box="[312,348,258,284]" count="1" pageId="14" pageNumber="327" type="female">1♀</specimenCount>
;
<collectingMunicipality id="A267AC81FFDDDD73FE866643FE17123E" box="[365,463,258,284]" pageId="14" pageNumber="327">Claveria</collectingMunicipality>
; foot of
<location id="47636020FFDDDD73FDDD6642FC8F123E" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFDDDD73FDDD6642FC8F123E" box="[566,855,258,285]" country="Philippines" latitude="8.700011" longLatPrecision="1" longitude="125.005775" municipality="Claveria" name="Mount Sumagaya-Lumot" pageId="14" pageNumber="327">Mount Sumagaya-Lumot</location>
;
<geoCoordinate id="2788503CFFDDDD73FC8E6643FC3C123E" box="[869,996,258,284]" degrees="8.700011" direction="north" orientation="latitude" pageId="14" pageNumber="327" precision="1" value="8.700011">N8.700011</geoCoordinate>
,
<geoCoordinate id="2788503CFFDDDD73FC196643FB53123F" box="[1010,1163,258,285]" degrees="125.005772" direction="east" orientation="longitude" pageId="14" pageNumber="327" precision="1" value="125.005775">E125.005772</geoCoordinate>
,
<quantity id="85449B1EFFDDDD73FB726643FB2F123E" box="[1177,1271,258,284]" metricMagnitude="3" metricUnit="m" metricValue="1.172" pageId="14" pageNumber="327" unit="m" value="1172.0">
<elevation id="C991D1C8FFDDDD73FB726643FB2F123E" box="[1177,1271,258,284]" metricMagnitude="3" metricUnit="m" metricValue="1.172" pageId="14" pageNumber="327" unit="m" value="1172.0">1172 m</elevation>
</quantity>
.a.s.l.;
<date id="3602103BFFDDDD73FAD66643FF0A1262" pageId="14" pageNumber="327" value="2024-04-25">
<collectingDate id="2646E9D3FFDDDD73FAD66643FF0A1262" pageId="14" pageNumber="327" value="2024-04-25">25 April 2024</collectingDate>
</date>
, 19h32; along the trail to forest patch,
<quantity id="85449B1EFFDDDD73FD976667FD6D1262" box="[636,693,294,320]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="327" unit="m" value="20.0">
<elevation id="C991D1C8FFDDDD73FD976667FD6D1262" box="[636,693,294,320]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="327" unit="m" value="20.0">20 m</elevation>
</quantity>
from a river, found on the leaf of a grass; coll.
<collectorName id="EF49532DFFDDDD73FB576667FA98121D" box="[1212,1344,293,320]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FA8A6667FF351241" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; SS37 (
<collectionCode id="24ADAE3EFFDDDD73FEA96608FE5D1241" box="[322,389,329,355]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="16C36673FFDFDD71FF7C60C7FB9314BC" ID-DOI="http://doi.org/10.5281/zenodo.13757717" ID-Zenodo-Dep="13757717" box="[151,1099,1926,1950]" httpUri="https://zenodo.org/record/13757717/files/figure.png" pageId="12" pageNumber="325" startId="12.[151,250,1926,1950]" targetBox="[151,1436,181,1902]" targetPageId="12" targetType="figure">
<paragraph id="420336FBFFDFDD71FF7C60C7FB9314BC" blockId="12.[151,1099,1926,1950]" box="[151,1099,1926,1950]" pageId="12" pageNumber="325">
<emphasis id="70C8EAE9FFDFDD71FF7C60C7FECB14BC" bold="true" box="[151,275,1926,1950]" pageId="12" pageNumber="325">FIGURE 9.</emphasis>
<taxonomicName id="85BC4D78FFDFDD71FEF260C7FD7F14BC" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[281,679,1926,1950]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="325" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFDFDD71FEF260C7FDD614BC" box="[281,526,1926,1950]" italics="true" pageId="12" pageNumber="325">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFDFDD71FDFF60C7FD7F14BC" author="Ingrisch, S." box="[532,679,1926,1950]" pageId="12" pageNumber="325" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
males in their natural habitat at Claveria.
</paragraph>
</caption>
<caption id="16C36673FFDEDD70FF7C60C5FBB314BE" ID-DOI="http://doi.org/10.5281/zenodo.13757719" ID-Zenodo-Dep="13757719" box="[151,1131,1924,1948]" httpUri="https://zenodo.org/record/13757719/files/figure.png" pageId="13" pageNumber="326" startId="13.[151,250,1924,1948]" targetBox="[151,1436,181,1900]" targetPageId="13" targetType="figure">
<paragraph id="420336FBFFDEDD70FF7C60C5FBB314BE" blockId="13.[151,1131,1924,1948]" box="[151,1131,1924,1948]" pageId="13" pageNumber="326">
<emphasis id="70C8EAE9FFDEDD70FF7C60C5FEF814BE" bold="true" box="[151,288,1924,1948]" pageId="13" pageNumber="326">FIGURE 10.</emphasis>
<taxonomicName id="85BC4D78FFDEDD70FECD60C5FD6B14BE" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[294,691,1924,1948]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="326" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFDEDD70FECD60C5FDC214BE" box="[294,538,1924,1948]" italics="true" pageId="13" pageNumber="326">Anthracites pyramidalis</emphasis>
<bibRefCitation id="262D4B0AFFDEDD70FDCA60C5FD6B14BE" author="Ingrisch, S." box="[545,691,1924,1948]" pageId="13" pageNumber="326" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
females in their natural habitat at Claveria.
</paragraph>
</caption>
<paragraph id="420336FBFFDDDD73FF2C662CFC8912EC" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<materialsCitation id="F2D43CA6FFDDDD73FF2C662CFC9512EC" collectionCode="PNM" collectorName="January &amp; D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Mindanao" elevation="5" latitude="6.183222" location="Mount Busa" longLatPrecision="1" longitude="124.68278" municipality="Lake" pageId="14" pageNumber="327" specimenCount="6" specimenCount-male="1" stateProvince="South Cotabato">
<collectingCountry id="3AAB766BFFDDDD73FF2C662CFEB712A5" box="[199,367,365,391]" name="Philippines" pageId="14" pageNumber="327">PHILIPPINES</collectingCountry>
,
<collectingCounty id="AB624E77FFDDDD73FE94662DFE2B12A4" box="[383,499,364,390]" pageId="14" pageNumber="327">Mindanao</collectingCounty>
,
<collectingRegion id="8078F819FFDDDD73FDEF662DFD6312A4" box="[516,699,364,390]" country="Philippines" name="South Cotabato" pageId="14" pageNumber="327">South Cotabato</collectingRegion>
: •
<specimenCount id="54BAFD72FFDDDD73FD0B662CFCDC12A5" box="[736,772,365,391]" count="1" pageId="14" pageNumber="327" type="male">1♂</specimenCount>
;
<collectingMunicipality id="A267AC81FFDDDD73FCFC662DFC9712A4" box="[791,847,364,390]" pageId="14" pageNumber="327">Lake</collectingMunicipality>
<collectingRegion id="8078F819FFDDDD73FCB0662DFC4E12A4" box="[859,918,364,390]" country="Philippines" name="Cebu" pageId="14" pageNumber="327">Sebu</collectingRegion>
, foot of the northern slope of
<location id="47636020FFDDDD73FAED662CFA4F12A5" LSID="urn:lsid:plazi:treatment:CA1587EDFFD9DD73FF7C67D6FC4B1538:47636020FFDDDD73FAED662CFA4F12A5" box="[1286,1431,365,391]" country="Philippines" county="Mindanao" latitude="6.183222" longLatPrecision="1" longitude="124.68278" municipality="Lake" name="Mount Busa" pageId="14" pageNumber="327" stateProvince="South Cotabato">Mount Busa</location>
;
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,
<geoCoordinate id="2788503CFFDDDD73FECF66D1FE651288" box="[292,445,400,426]" degrees="124.682778" direction="east" orientation="longitude" pageId="14" pageNumber="327" precision="1" value="124.68278">E124.682778</geoCoordinate>
, 1041±
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<elevation id="C991D1C8FFDDDD73FDF966D1FD9C1288" box="[530,580,400,427]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="327" unit="m" value="5.0">5 m</elevation>
</quantity>
.
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a.s.l.; 30
<collectorName id="EF49532DFFDDDD73FD4666D0FCDD1289" box="[685,773,401,427]" pageId="14" pageNumber="327">January</collectorName>
, 2024, 13h57; open area in secondary forest, found on the leaf of a shrub; coll.
<collectorName id="EF49532DFFDDDD73FE9466F5FDDC12EF" box="[383,516,435,462]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FDCD66F5FD6612EF" box="[550,702,435,462]" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; AI5 (
<collectionCode id="24ADAE3EFFDDDD73FCE966F5FC9D12EC" box="[770,837,436,462]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFDDDD73FF2C6696FA4D11CB" pageId="14" pageNumber="327" type="discussion">
<paragraph id="420336FBFFDDDD73FF2C6696FC041136" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C6696FEE312D3" bold="true" box="[199,315,471,497]" pageId="14" pageNumber="327">Remarks.</emphasis>
The species was described based on a female
<typeStatus id="9D078859FFDDDD73FCA66696FC7712D3" box="[845,943,471,497]" pageId="14" pageNumber="327" type="holotype">holotype</typeStatus>
and a male nymph from Mount Balatukan. New material collected from Claveria allows us to describe the adult male.
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C655FFA4D11CB" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C655FFE7A111A" bold="true" box="[199,418,542,568]" pageId="14" pageNumber="327">Revised diagnosis.</emphasis>
In additions to the diagnosis given in
<bibRefCitation id="262D4B0AFFDDDD73FCBB655FFBDA111B" author="Ingrisch, S." box="[848,1026,542,569]" pageId="14" pageNumber="327" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch (2015)</bibRefCitation>
: the species is similar to
<taxonomicName id="85BC4D78FFDDDD73FAF0655FFE4F117E" authority="Ingrisch 2015" authorityName="Ingrisch" authorityYear="2015" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="327" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="70C8EAE9FFDDDD73FAF0655FFF2C1179" italics="true" pageId="14" pageNumber="327">Anthracites sinuatus</emphasis>
<bibRefCitation id="262D4B0AFFDDDD73FF176500FE4F117E" author="Ingrisch, S." box="[252,407,577,604]" pageId="14" pageNumber="327" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch 2015</bibRefCitation>
</taxonomicName>
from Luzon Island and
<taxonomicName id="85BC4D78FFDDDD73FD4C6500FBEF117E" authority="Ingrisch, 2015" authorityName="Ingrisch" authorityYear="2015" box="[679,1079,577,604]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="327" phylum="Arthropoda" rank="species" species="romblon">
<emphasis id="70C8EAE9FFDDDD73FD4C6500FC561179" box="[679,910,577,603]" italics="true" pageId="14" pageNumber="327">Anthracites romblon</emphasis>
<bibRefCitation id="262D4B0AFFDDDD73FC7E6500FBEF117E" author="Ingrisch, S." box="[917,1079,577,604]" pageId="14" pageNumber="327" pagination="1 - 308" refId="ref9269" refString="Ingrisch, S. (2015) A revision of the Axylus group of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) and of some other species formerly included in Nicsara or Anthracites. Zootaxa, 4046 (1), 1 - 308. https: // doi. org / 10.11646 / zootaxa. 4046.1.1" type="journal article" year="2015">Ingrisch, 2015</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="8078F819FFDDDD73FB976500FB3C1179" box="[1148,1252,577,603]" country="Philippines" name="Romblon" pageId="14" pageNumber="327">Romblon</collectingRegion>
Island in the Xshaped titillators and in the apex of the lateral extension; but the species differs from both species by the face having a black band that widen ventrad (instead of a pair of black bands), abdominal tergites with lateral bands (instead of median band in
<taxonomicName id="85BC4D78FFDDDD73FEAC65EDFDF411E4" authorityName="Ingrisch" authorityYear="2015" box="[327,556,684,710]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="327" phylum="Arthropoda" rank="species" species="romblon">
<emphasis id="70C8EAE9FFDDDD73FEAC65EDFDF411E4" box="[327,556,684,710]" italics="true" pageId="14" pageNumber="327">Anthracites romblon</emphasis>
</taxonomicName>
) and the cercus having a dorsal distal process very long, falcate with acute apex and a shorter nipple-like ventral process, the titillators with only a pair of lateral sclerites (instead of two or none).
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFDDDD73FF2C65B3FDD516A9" pageId="14" pageNumber="327" type="description">
<paragraph id="420336FBFFDDDD73FF2C65B3FCF5102F" blockId="14.[151,1437,151,1562]" box="[199,813,754,781]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C65B3FEAD102E" bold="true" box="[199,373,754,780]" pageId="14" pageNumber="327">Redescription.</emphasis>
Male habitus as shown in
<figureCitation id="DA872A7EFFDDDD73FD7465B2FD20102F" box="[671,760,755,781]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="14" pageNumber="327">Figs 1C</figureCitation>
,
<figureCitation id="DA872A7EFFDDDD73FCEE65B2FCF1102F" box="[773,809,755,781]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="14" pageNumber="327">2C</figureCitation>
.
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C6457FB8A1013" blockId="14.[151,1437,151,1562]" box="[199,1106,790,817]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C6457FE701012" box="[199,424,790,816]" italics="true" pageId="14" pageNumber="327">Head and pronotum</emphasis>
. Head and pronotum not different from female (
<figureCitation id="DA872A7EFFDDDD73FC2E6457FB9E1013" box="[965,1094,790,817]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[151,1436,309,1823]" captionTargetId="figure-62@8.[151,1436,309,1823]" captionTargetPageId="8" captionText="FIGURE 5. Anthracites geniculatus Dohrn, 1905 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), posterior (E), dorso-lateral (F) and dorso-posterior (G) views. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757709" httpUri="https://zenodo.org/record/13757709/files/figure.png" pageId="14" pageNumber="327">Figs 5A–C</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C647BFCD910B9" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C647BFE7F1071" box="[199,423,825,851]" italics="true" pageId="14" pageNumber="327">Tegmina and wings.</emphasis>
Tegmen micropterous, mirror area partly covered by pronotal disc (
<figureCitation id="DA872A7EFFDDDD73FB7D6478FB371076" box="[1174,1263,825,852]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Fig. 7A</figureCitation>
), reaching base of second abdominal tergite, with apex subtruncated; infumated (except mirror area), veins orange yellow with cells black, lateral field veins cream coloured (
<figureCitation id="DA872A7EFFDDDD73FD8C64C1FD2D10B9" box="[615,757,896,923]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Figs 7A, 7B</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C64E5FCED1645" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C64E5FEE1109C" box="[199,313,932,958]" italics="true" pageId="14" pageNumber="327">Abdomen.</emphasis>
Tenth abdominal tergite with posterior margin emarginated at middle, laterally produced into two stout triangular lobes with truncated apices; posterior end at middle depressed forming groove (
<figureCitation id="DA872A7EFFDDDD73FB736486FAFF10C0" box="[1176,1319,967,994]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Figs 7D, 7E</figureCitation>
). Epiproct tongue shaped, at middle depressed (
<figureCitation id="DA872A7EFFDDDD73FDD564AAFD4D1727" box="[574,661,1003,1029]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Fig. 7F</figureCitation>
). Cercus stout, slightly curved inwards, dorso-laterally compressed; externo-distal part stoutly conical with subacute apex; at distal end with two inner processes pointing inwards; dorsal process very long (nearly as long as base of cercus), falcate and slightly curved inwards, apex with acute spine; ventral process shorter, reaching middle of dorsal process, elongated nipple-like with apex obtuse (
<figureCitation id="DA872A7EFFDDDD73FAD86314FF7217B1" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Figs 7D– F</figureCitation>
). Subgenital plate about as long as wide, with anterior margin concave, gently converging lateral margins, apical margin roundly excised (
<figureCitation id="DA872A7EFFDDDD73FE4563DDFDD01794" box="[430,520,1180,1206]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Fig. 7G</figureCitation>
); margin around bases of styli on dorsal side swollen and with cylindrical internal tooth of obtuse apex (
<figureCitation id="DA872A7EFFDDDD73FE786381FE3217F8" box="[403,490,1216,1242]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Fig. 7F</figureCitation>
). Styli stout with apex obtuse (
<figureCitation id="DA872A7EFFDDDD73FCBA6381FC7017F8" box="[849,936,1216,1242]" captionStart="FIGURE 7" captionStartId="10.[151,250,1945,1969]" captionTargetBox="[158,1430,434,1921]" captionTargetId="figure-62@10.[158,1430,434,1921]" captionTargetPageId="10" captionText="FIGURE 7. Anthracites pyramidalis Ingrisch, 2015 ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), dorso-posterior (E), ventro-posterior (F) and ventral (G) views. The arrow points to internal tooth of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–G)." figureDoi="http://doi.org/10.5281/zenodo.13757713" httpUri="https://zenodo.org/record/13757713/files/figure.png" pageId="14" pageNumber="327">Fig. 7F</figureCitation>
). Titillators (
<figureCitation id="DA872A7EFFDDDD73FBD66381FB5917F8" box="[1085,1153,1216,1242]" captionStart="FIGURE 8" captionStartId="11.[151,250,1135,1159]" captionTargetBox="[172,1414,202,1093]" captionTargetId="figure-17@11.[151,1436,181,1112]" captionTargetPageId="11" captionText="FIGURE 8. Anthracites pyramidalis Ingrisch, 2015 ♂ titillators in ventro-distal (A), ventral (B) and dorso-proximal (C) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13757715" httpUri="https://zenodo.org/record/13757715/files/figure.png" pageId="14" pageNumber="327">Fig. 8</figureCitation>
) separate, X-shaped and wing-like; lateral arm pointing ventro-distally; lateral arm slender forming ‘skeleton of the wing’, gently narrowing into apex faintly hammer-head, apex with stout apical and proximal teeth; proximal of lateral arm with flattened semi-sclerotized hyalinous plate forming ‘wing’; base at proximal end slender and tapering into subacute apex; provided with two pairs of lateral sclerites, roundly conical.
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C6230FDD516A9" blockId="14.[151,1437,151,1562]" box="[199,525,1393,1419]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C6230FEA616A9" bold="true" box="[199,382,1393,1419]" pageId="14" pageNumber="327">Measurements.</emphasis>
See
<tableCitation id="0F3E0340FFDDDD73FE586230FDDF16A9" box="[435,519,1393,1419]" captionStart="TABLE 1" captionStartId="22.[151,239,152,176]" captionText="TABLE 1. Measurements of Anthracites, in mm." pageId="14" pageNumber="327">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFDDDD73FF2C62D5FE9316F0" pageId="14" pageNumber="327" type="distribution">
<paragraph id="420336FBFFDDDD73FF2C62D5FE9316F0" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C62D5FEB9168C" bold="true" box="[199,353,1428,1454]" pageId="14" pageNumber="327">Distribution.</emphasis>
<collectingRegion id="8078F819FFDDDD73FE8C62D5FDF5168C" box="[359,557,1428,1454]" country="Philippines" name="Misamis Oriental" pageId="14" pageNumber="327">Misamis Oriental</collectingRegion>
: Mount Balatukan, Mount Sumagaya-Lumot;
<collectingRegion id="8078F819FFDDDD73FBC562D5FB07168C" box="[1070,1247,1428,1454]" country="Philippines" name="South Cotabato" pageId="14" pageNumber="327">South Cotabato</collectingRegion>
: Lake
<collectingRegion id="8078F819FFDDDD73FACD62D5FA86168C" box="[1318,1374,1428,1454]" country="Philippines" name="Cebu" pageId="14" pageNumber="327">Sebu</collectingRegion>
[new locality record].
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFDDDD73FF2C629DFC4B1538" pageId="14" pageNumber="327" type="biology_ecology">
<paragraph id="420336FBFFDDDD73FF2C629DFC4B1538" blockId="14.[151,1437,151,1562]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C629DFEF416D4" bold="true" box="[199,300,1500,1526]" pageId="14" pageNumber="327">Ecology.</emphasis>
This species was found to be abundant in open area outside or near mossy forest (
<figureCitation id="DA872A7EFFDDDD73FB09629AFAF416D4" box="[1250,1324,1499,1526]" captionStart="FIGURE 9" captionStartId="12.[151,250,1926,1950]" captionTargetBox="[151,1436,181,1902]" captionTargetId="figure-19@12.[151,1436,181,1902]" captionTargetPageId="12" captionText="FIGURE 9. Anthracites pyramidalis Ingrisch, 2015 males in their natural habitat at Claveria." figureDoi="http://doi.org/10.5281/zenodo.13757717" httpUri="https://zenodo.org/record/13757717/files/figure.png" pageId="14" pageNumber="327">Figs 9</figureCitation>
,
<figureCitation id="DA872A7EFFDDDD73FAD1629DFA8116D4" box="[1338,1369,1500,1526]" captionStart="FIGURE 10" captionStartId="13.[151,250,1924,1948]" captionTargetBox="[151,1436,181,1900]" captionTargetId="figure-17@13.[151,1436,181,1900]" captionTargetPageId="13" captionText="FIGURE 10. Anthracites pyramidalis Ingrisch, 2015 females in their natural habitat at Claveria." figureDoi="http://doi.org/10.5281/zenodo.13757719" httpUri="https://zenodo.org/record/13757719/files/figure.png" pageId="14" pageNumber="327">10</figureCitation>
). It is sometimes also found inside the forest among foliage of small trees.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DD76756C035E9155543A1DAC96FBEF7E">A taxonomic review on the katydids of the genus Anthracites Redtenbacher, 1891 (Tettigoniidae: Conocephalinae: Agraeciini) from Mindanao, Philippines</mods:title>
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Tan, Bahoy &amp; Sabang
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(
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<figureCitation id="DA872A7EFFDDDD73FED361C3FEA615BE" box="[312,382,1666,1692]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="15.[151,250,1822,1846]" captionStartId-1="16.[151,250,1181,1205]" captionStartId-2="17.[151,250,1793,1817]" captionTargetBox-0="[151,1435,181,1798]" captionTargetBox-1="[177,1429,205,1153]" captionTargetBox-2="[151,1436,181,1757]" captionTargetId-0="figure-17@15.[151,1436,181,1798]" captionTargetId-1="figure-19@16.[151,1436,181,1157]" captionTargetId-2="figure-17@17.[151,1436,181,1769]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." captionText-1="FIGURE 12. Anthracites sebu sp. nov. ♂ titillators in apical (A), ventro-proximal (B), dorso-proximal (C) and ventro-distal (D) views. Scale bars: 1 mm" captionText-2="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi-0="http://doi.org/10.5281/zenodo.13757721" figureDoi-1="http://doi.org/10.5281/zenodo.13757723" figureDoi-2="http://doi.org/10.5281/zenodo.13757725" httpUri-0="https://zenodo.org/record/13757721/files/figure.png" httpUri-1="https://zenodo.org/record/13757723/files/figure.png" httpUri-2="https://zenodo.org/record/13757725/files/figure.png" pageId="14" pageNumber="327">11–13</figureCitation>
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</date>
, 16h09; open area inside secondary forest,
<quantity id="85449B1EFFDDDD73FC7E604BFC151407" box="[917,973,1802,1829]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="327" unit="m" value="15.0">
<elevation id="C991D1C8FFDDDD73FC7E604BFC151407" box="[917,973,1802,1829]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="327" unit="m" value="15.0">15 m</elevation>
</quantity>
away from a stream; found on the leaf of a shrub [
<taxonomicName id="85BC4D78FFDDDD73FF12606FFE65146A" box="[249,445,1838,1864]" class="Magnoliopsida" family="Apocynaceae" genus="Catharanthus" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="327" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="70C8EAE9FFDDDD73FF12606FFE4B146A" box="[249,403,1838,1864]" italics="true" pageId="14" pageNumber="327">Catharanthus</emphasis>
sp.
</taxonomicName>
]; coll.
<collectorName id="EF49532DFFDDDD73FDE1606FFD57146A" box="[522,655,1838,1864]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FD5A606FFC92146A" box="[689,842,1838,1864]" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; AI5.6 (
<collectionCode id="24ADAE3EFFDDDD73FC48606FFC3E146A" box="[931,998,1838,1864]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="420336FBFFDDDD73FF2C6010FEB614D8" blockId="14.[151,1436,1731,2042]" pageId="14" pageNumber="327">
<emphasis id="70C8EAE9FFDDDD73FF2C6010FE61144D" bold="true" box="[199,441,1873,1903]" pageId="14" pageNumber="327">
<typeStatus id="9D078859FFDDDD73FF2C6010FEE71449" box="[199,319,1873,1899]" pageId="14" pageNumber="327" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="54BAFD72FFDDDD73FEBA6012FEAC144D" box="[337,372,1875,1903]" count="1" pageId="14" pageNumber="327" type="male">1♂</specimenCount>
,
<specimenCount id="54BAFD72FFDDDD73FE686012FE70144D" box="[387,424,1875,1903]" count="1" pageId="14" pageNumber="327" type="female">1♀</specimenCount>
):
</emphasis>
•
<materialsCitation id="F2D43CA6FFDDDD73FE386010FB3C1449" box="[467,1252,1873,1900]" collectingDate="2024-01-30" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Mindanao" elevation="4" latitude="6.182333" location="Barangay Lamlakak" longLatPrecision="1" longitude="124.68108" municipality="Lake" pageId="14" pageNumber="327" specimenCount="1" specimenCount-female="1" stateProvince="South Cotabato" typeStatus="paratype">
<specimenCount id="54BAFD72FFDDDD73FE386010FE201449" box="[467,504,1873,1899]" count="1" pageId="14" pageNumber="327" type="female">1♀</specimenCount>
; same locality as holotype;
<geoCoordinate id="2788503CFFDDDD73FCDC6010FC6F1449" box="[823,951,1873,1899]" degrees="6.182333" direction="north" orientation="latitude" pageId="14" pageNumber="327" precision="1" value="6.182333">N6.182333</geoCoordinate>
,
<geoCoordinate id="2788503CFFDDDD73FC286010FB851449" box="[963,1117,1873,1899]" degrees="124.681083" direction="east" orientation="longitude" pageId="14" pageNumber="327" precision="1" value="124.68108">E124.681083</geoCoordinate>
, 1077±
<quantity id="85449B1EFFDDDD73FB5B6010FB3C1449" box="[1200,1252,1873,1899]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="327" unit="m" value="4.0">
<specimenCount id="54BAFD72FFDDDD73FB5B6010FB3C1449" box="[1200,1252,1873,1899]" count="4" pageId="14" pageNumber="327" type="generic">
<elevation id="C991D1C8FFDDDD73FB5B6010FB3C1449" box="[1200,1252,1873,1899]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="327" unit="m" value="4.0">4 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="F2D43CA6FFDDDD73FB086010FF3F1490" collectingDate="2024-01-30" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" pageId="14" pageNumber="327" specimenCount="1" stateProvince="South Cotabato" typeStatus="paratype">
.a.s.l;
<date id="3602103BFFDDDD73FACA6010FF0A14AD" pageId="14" pageNumber="327" value="2024-01-30">
<collectingDate id="2646E9D3FFDDDD73FACA6010FF0A14AD" pageId="14" pageNumber="327" value="2024-01-30">30 January 2024</collectingDate>
</date>
, 16h18; found inside a coiled dead leaf of abaca plant (
<taxonomicName id="85BC4D78FFDDDD73FCD36037FC1014AD" box="[824,968,1909,1935]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="14" pageNumber="327" phylum="Tracheophyta" rank="species" species="textilis">
<emphasis id="70C8EAE9FFDDDD73FCD36037FC1014AD" box="[824,968,1909,1935]" italics="true" pageId="14" pageNumber="327">Musa textilis</emphasis>
</taxonomicName>
); coll.
<collectorName id="EF49532DFFDDDD73FBFC6034FB4414AD" box="[1047,1180,1909,1935]" pageId="14" pageNumber="327">D.C. Bahoy</collectorName>
&amp;
<collectorName id="EF49532DFFDDDD73FB566034FA8D14AD" box="[1213,1365,1909,1935]" pageId="14" pageNumber="327">A.M. Sabang</collectorName>
; LS15 (
<collectionCode id="24ADAE3EFFDDDD73FF7460D9FF071490" box="[159,223,1944,1970]" country="Philippines" name="Philippine National Museum" pageId="14" pageNumber="327">PNM</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="F2D43CA6FFDDDD73FF1560D9FBDB1490" box="[254,1027,1944,1971]" collectingDate="2024-01-30" collectionCode="PNM" collectorName="D. C. Bahoy &amp; A. M. Sabang" country="Philippines" county="Mindanao" elevation="3" latitude="6.185531" location="Barangay Lamlakak" longLatPrecision="1" longitude="124.68407" municipality="Lake" pageId="14" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="South Cotabato" typeStatus="paratype">
<specimenCount id="54BAFD72FFDDDD73FF1560D9FEFB1490" box="[254,291,1944,1970]" count="1" pageId="14" pageNumber="327" type="male">1♂</specimenCount>
; same locality as holotype;
<geoCoordinate id="2788503CFFDDDD73FDB060D9FD021491" box="[603,730,1944,1971]" degrees="6.185531" direction="north" orientation="latitude" pageId="14" pageNumber="327" precision="1" value="6.185531">N6.185531</geoCoordinate>
,
<geoCoordinate id="2788503CFFDDDD73FD0D60D9FCA71490" box="[742,895,1944,1970]" degrees="124.684064" direction="east" orientation="longitude" pageId="14" pageNumber="327" precision="1" value="124.68407">E124.684064</geoCoordinate>
, 1052±
<quantity id="85449B1EFFDDDD73FC3860D9FBDB1490" box="[979,1027,1944,1971]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="327" unit="m" value="3.0">
<specimenCount id="54BAFD72FFDDDD73FC3860D9FBDB1490" box="[979,1027,1944,1971]" count="3" pageId="14" pageNumber="327" type="generic">
<elevation id="C991D1C8FFDDDD73FC3860D9FBDB1490" box="[979,1027,1944,1971]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="327" unit="m" value="3.0">3 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
.a.s.l;
<date id="3602103BFFDDDD73FBD460D9FB231490" box="[1087,1275,1944,1970]" pageId="14" pageNumber="327" value="2024-02-02">2 February 2024</date>
, 20h08; found on open area
<quantity id="85449B1EFFDDDD73FEC160FDFEBA14F4" box="[298,354,1980,2006]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="327" unit="km" value="1.0">1 km</quantity>
from a small agricultural site; found on the leaf of shrub; coll. D.C. Bahoy &amp; A.M. Sabang; L57A (MNHN-EO1356).
</paragraph>
</subSubSection>
<caption id="16C36673FFDCDD72FF7C605FFDC6145C" ID-DOI="http://doi.org/10.5281/zenodo.13757721" ID-Zenodo-Dep="13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="15" pageNumber="328" startId="15.[151,250,1822,1846]" targetBox="[151,1435,181,1798]" targetPageId="15" targetType="figure">
<paragraph id="420336FBFFDCDD72FF7C605FFDC6145C" blockId="15.[151,1437,1822,1918]" pageId="15" pageNumber="328">
<emphasis id="70C8EAE9FFDCDD72FF7C605FFEC71414" bold="true" box="[151,287,1822,1846]" pageId="15" pageNumber="328">FIGURE 11.</emphasis>
<taxonomicName id="85BC4D78FFDCDD72FECD605FFE081414" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[294,464,1822,1846]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="328" phylum="Arthropoda" rank="species" species="sebu" status="sp. nov.">
<emphasis id="70C8EAE9FFDCDD72FECD605FFE081414" box="[294,464,1822,1846]" italics="true" pageId="15" pageNumber="328">Anthracites sebu</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFDCDD72FE3C605FFDF11414" bold="true" box="[471,553,1822,1846]" pageId="15" pageNumber="328">
<taxonomicNameLabel id="6BFB5792FFDCDD72FE3C605FFDF11414" box="[471,553,1822,1846]" pageId="15" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F).
</paragraph>
</caption>
<caption id="16C36673FFC3DD6DFF7C63DCFE6F17FB" ID-DOI="http://doi.org/10.5281/zenodo.13757723" ID-Zenodo-Dep="13757723" httpUri="https://zenodo.org/record/13757723/files/figure.png" pageId="16" pageNumber="329" startId="16.[151,250,1181,1205]" targetBox="[177,1429,205,1153]" targetPageId="16" targetType="figure">
<paragraph id="420336FBFFC3DD6DFF7C63DCFE6F17FB" blockId="16.[151,1436,1181,1241]" pageId="16" pageNumber="329">
<emphasis id="70C8EAE9FFC3DD6DFF7C63DCFEF91797" bold="true" box="[151,289,1181,1205]" pageId="16" pageNumber="329">FIGURE 12.</emphasis>
<taxonomicName id="85BC4D78FFC3DD6DFEC263DCFE0C1797" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[297,468,1181,1205]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="329" phylum="Arthropoda" rank="species" species="sebu" status="sp. nov.">
<emphasis id="70C8EAE9FFC3DD6DFEC263DCFE0C1797" box="[297,468,1181,1205]" italics="true" pageId="16" pageNumber="329">Anthracites sebu</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFC3DD6DFE3763DCFDF61797" bold="true" box="[476,558,1181,1205]" pageId="16" pageNumber="329">
<taxonomicNameLabel id="6BFB5792FFC3DD6DFE3763DCFDF61797" box="[476,558,1181,1205]" pageId="16" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ titillators in apical (A), ventro-proximal (B), dorso-proximal (C) and ventro-distal (D) views. Scale bars: 1 mm
</paragraph>
</caption>
<subSubSection id="0AA66570FFC3DD6DFF2C626AFB671667" box="[199,1215,1323,1349]" pageId="16" pageNumber="329" type="etymology">
<paragraph id="420336FBFFC3DD6DFF2C626AFB671667" blockId="16.[151,1437,1323,2023]" box="[199,1215,1323,1349]" pageId="16" pageNumber="329">
<emphasis id="70C8EAE9FFC3DD6DFF2C626AFE961667" bold="true" box="[199,334,1323,1349]" pageId="16" pageNumber="329">Etymology.</emphasis>
This species is named after its
<typeStatus id="9D078859FFC3DD6DFD47626AFD041667" box="[684,732,1323,1349]" pageId="16" pageNumber="329">type</typeStatus>
locality, in Lake
<collectingRegion id="8078F819FFC3DD6DFC4B626AFC031667" box="[928,987,1323,1349]" country="Philippines" name="Cebu" pageId="16" pageNumber="329">Sebu</collectingRegion>
; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFC3DD6DFF2C620FFD3015CC" pageId="16" pageNumber="329" type="diagnosis">
<paragraph id="420336FBFFC3DD6DFF2C620FFCD41538" blockId="16.[151,1437,1323,2023]" pageId="16" pageNumber="329">
<emphasis id="70C8EAE9FFC3DD6DFF2C620FFE9A164A" bold="true" box="[199,322,1358,1384]" pageId="16" pageNumber="329">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="85BC4D78FFC3DD6DFD70620FFC70164A" authorityName="Ingrisch" authorityYear="2015" box="[667,936,1358,1384]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="329" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFC3DD6DFD70620FFC70164A" box="[667,936,1358,1384]" italics="true" pageId="16" pageNumber="329">Anthracites pyramidalis</emphasis>
</taxonomicName>
by the general colouration, face with a broad black band from fastigium verticis to base of clypeus, widening ventrad and its titillators X-shaped; but differs by the pronotum unicolourous orange without black spots at hind margin, male tegmen also without dark cells, male cercus with a stout distal internal process not surpassing apex of cercus, constricted just before apex and with sharp tooth at tip (instead of two processes, one of which long and falcate), and the titillators having lateral arms distinctly having a stout hook (instead of a hammer-headed apex).
</paragraph>
<paragraph id="420336FBFFC3DD6DFF2C6162FDE7158A" blockId="16.[151,1437,1323,2023]" pageId="16" pageNumber="329">
The species is also similar to
<taxonomicName id="85BC4D78FFC3DD6DFDC66162FCC0151F" authorityName="Ingrisch" authorityYear="2015" box="[557,792,1571,1597]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="329" phylum="Arthropoda" rank="species" species="romblon">
<emphasis id="70C8EAE9FFC3DD6DFDC66162FCC0151F" box="[557,792,1571,1597]" italics="true" pageId="16" pageNumber="329">Anthracites romblon</emphasis>
</taxonomicName>
by its male cercus with a stout distal internal process constricted just before apex and with a sharp tooth at tip, but differs by the process smaller, slenderer and not surpassing apex of cercus. It also differs by the absence of dark patterns on pronotum and the tergites having lateral black bands (instead of median ones).
</paragraph>
<paragraph id="420336FBFFC3DD6DFF2C61F0FD3015CC" blockId="16.[151,1437,1323,2023]" pageId="16" pageNumber="329">
The female also differs from
<taxonomicName id="85BC4D78FFC3DD6DFDE061F0FCC015E9" authorityName="Ingrisch" authorityYear="2015" box="[523,792,1713,1739]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="329" phylum="Arthropoda" rank="species" species="pyramidalis">
<emphasis id="70C8EAE9FFC3DD6DFDE061F0FCC015E9" box="[523,792,1713,1739]" italics="true" pageId="16" pageNumber="329">Anthracites pyramidalis</emphasis>
</taxonomicName>
by the subgenital plate transverse and at the posterior end broadly bilobed and its lateral margins not upcurved.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFC3DD6FFF2C61B9FDD517FB" lastPageId="18" lastPageNumber="331" pageId="16" pageNumber="329" type="description">
<paragraph id="420336FBFFC3DD6DFF2C61B9FCEE14C5" blockId="16.[151,1437,1323,2023]" pageId="16" pageNumber="329">
<emphasis id="70C8EAE9FFC3DD6DFF2C61B9FE811430" bold="true" box="[199,345,1784,1810]" pageId="16" pageNumber="329">Description.</emphasis>
Male habitus as shown in
<figureCitation id="DA872A7EFFC3DD6DFD7B61B9FD351430" box="[656,749,1784,1810]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="16" pageNumber="329">Figs 1D</figureCitation>
,
<figureCitation id="DA872A7EFFC3DD6DFD1761B9FCFA1430" box="[764,802,1784,1810]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="16" pageNumber="329">2D</figureCitation>
. Morphology typical of genus (
<figureCitation id="DA872A7EFFC3DD6DFB6061B9FAC21430" box="[1163,1306,1784,1810]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="16" pageNumber="329">Figs 11A–C</figureCitation>
), similar to
<taxonomicName id="85BC4D78FFC3DD6DFF7C605AFEBE1417" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[151,358,1819,1845]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="329" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
<emphasis id="70C8EAE9FFC3DD6DFF7C605AFEBE1417" box="[151,358,1819,1845]" italics="true" pageId="16" pageNumber="329">Anthracites alatus</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFC3DD6DFE9B605DFE141414" bold="true" box="[368,460,1820,1846]" pageId="16" pageNumber="329">
<taxonomicNameLabel id="6BFB5792FFC3DD6DFE9B605DFE141414" box="[368,460,1820,1846]" pageId="16" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Thoracic spiracle large and oval, nearly completely covered by paranota (
<figureCitation id="DA872A7EFFC3DD6DFACC605AFA571414" box="[1319,1423,1819,1846]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="16" pageNumber="329">Fig. 11B</figureCitation>
). Tegmen micropterous, mirror area fully covered by pronotal disc, reaching base of fourth abdominal tergite, with apex roundly truncated (
<figureCitation id="DA872A7EFFC3DD6DFE436023FDD7145F" box="[424,527,1890,1917]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="16" pageNumber="329">Fig. 11A</figureCitation>
). Femora with the following number of spines on ventral margins: anterior femur 7 external, 6 internal; middle femur 6–7 external, 2–3 internal; posterior femur 10–11 external, 2–3 internal. Knee lobes of anterior and middle femora obtuse externally and spinose internally; knee of posterior femur bispinose on both sides, longer than those of anterior and middle femora.
</paragraph>
<caption id="16C36673FFC2DD6CFF7C6040FED41443" ID-DOI="http://doi.org/10.5281/zenodo.13757725" ID-Zenodo-Dep="13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="17" pageNumber="330" startId="17.[151,250,1793,1817]" targetBox="[151,1436,181,1757]" targetPageId="17" targetType="figure">
<paragraph id="420336FBFFC2DD6CFF7C6040FED41443" blockId="17.[151,1436,1793,1889]" pageId="17" pageNumber="330">
<emphasis id="70C8EAE9FFC2DD6CFF7C6040FEFB143B" bold="true" box="[151,291,1793,1817]" pageId="17" pageNumber="330">FIGURE 13.</emphasis>
<taxonomicName id="85BC4D78FFC2DD6CFEC76040FE00143B" authorityName="Sabang &amp; Bahoy &amp; Grumo &amp; Gono &amp; Salvador &amp; Rivera &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[300,472,1793,1817]" class="Insecta" family="Tettigoniidae" genus="Anthracites" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="330" phylum="Arthropoda" rank="species" species="sebu" status="sp. nov.">
<emphasis id="70C8EAE9FFC2DD6CFEC76040FE00143B" box="[300,472,1793,1817]" italics="true" pageId="17" pageNumber="330">Anthracites sebu</emphasis>
</taxonomicName>
<emphasis id="70C8EAE9FFC2DD6CFE0A6040FDED143B" bold="true" box="[481,565,1793,1817]" pageId="17" pageNumber="330">
<taxonomicNameLabel id="6BFB5792FFC2DD6CFE0A6040FDED143B" box="[481,565,1793,1817]" pageId="17" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D).
</paragraph>
</caption>
<paragraph id="420336FBFFC1DD6FFF2C67D6FC271178" blockId="18.[151,1437,151,1277]" pageId="18" pageNumber="331">
<emphasis id="70C8EAE9FFC1DD6FFF2C67D6FEE11393" box="[199,313,151,177]" italics="true" pageId="18" pageNumber="331">Abdomen.</emphasis>
Tenth abdominal tergite transverse, distinctly wider than long, narrows towards apex; at posterior margin truncated, faintly bulbous laterally and emarginated in middle, without setae (
<figureCitation id="DA872A7EFFC1DD6FFB8E67FBFB1613F6" box="[1125,1230,186,212]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11D</figureCitation>
); lateral lobe near posterior end faintly folded (
<figureCitation id="DA872A7EFFC1DD6FFE0E679FFD9313DA" box="[485,587,222,248]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11E</figureCitation>
). Epiproct broadly tongue shaped, at base with middle slightly depressed forming a groove (
<figureCitation id="DA872A7EFFC1DD6FFE9A6640FE0F1239" box="[369,471,257,283]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11E</figureCitation>
). Cercus short and stout, dorso-laterally compressed; distal end triangular subacute; with stout distal internal process triangular, pointing inward, constricted just before apex and with sharp tooth at tip (
<figureCitation id="DA872A7EFFC1DD6FFF2D6609FEA81240" box="[198,368,328,354]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Figs 11D, 11E</figureCitation>
). Subgenital plate about as long as wide; with anterior margin concave; apical end angularly emarginated with dense setae, posterior margin at middle produced into long cylindrical process with acute apex, process reaching middle of emargination (
<figureCitation id="DA872A7EFFC1DD6FFD9F66CEFD00128B" box="[628,728,399,425]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11F</figureCitation>
); margin around bases of styli on dorsal side swollen and with acute internal tooth (
<figureCitation id="DA872A7EFFC1DD6FFE6C66F3FE3512EF" box="[391,493,434,461]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11E</figureCitation>
). Styli stout and long with apex rounded (
<figureCitation id="DA872A7EFFC1DD6FFC3266F3FBE612EF" box="[985,1086,434,461]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11F</figureCitation>
). Titillators (
<figureCitation id="DA872A7EFFC1DD6FFB3866F3FAFF12EF" box="[1235,1319,434,461]" captionStart="FIGURE 12" captionStartId="16.[151,250,1181,1205]" captionTargetBox="[177,1429,205,1153]" captionTargetId="figure-19@16.[151,1436,181,1157]" captionTargetPageId="16" captionText="FIGURE 12. Anthracites sebu sp. nov. ♂ titillators in apical (A), ventro-proximal (B), dorso-proximal (C) and ventro-distal (D) views. Scale bars: 1 mm" figureDoi="http://doi.org/10.5281/zenodo.13757723" httpUri="https://zenodo.org/record/13757723/files/figure.png" pageId="18" pageNumber="331">Fig. 12</figureCitation>
) separate, X-shaped and wing-like; lateral arm pointing ventrally; lateral arm slender forming the ‘skeleton of the wing’, apical end forming strongly sclerotized and stout hook with subacute apex; proximal of lateral arm with flattened semi-sclerotized hyalinous plate forming the ‘wing’; base with proximal end tongue shaped having rounded apex; provided with two pairs of lateral sclerites, small, slender and slightly curved.
</paragraph>
<paragraph id="420336FBFFC1DD6FFF2C6525FAE111CA" blockId="18.[151,1437,151,1277]" pageId="18" pageNumber="331">
Female (
<figureCitation id="DA872A7EFFC1DD6FFEC26525FE0A115C" box="[297,466,612,638]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Figs 13A, 13B</figureCitation>
). Tenth abdominal tergite short and transverse, posterior end angularly emarginated in middle; forming two broadly rounded lobes; with longitudinal median groove (
<figureCitation id="DA872A7EFFC1DD6FFC1265C6FB851183" box="[1017,1117,647,673]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Fig. 13C</figureCitation>
). Epiproct transverse, almost as wide as long, broadly tongue shaped (
<figureCitation id="DA872A7EFFC1DD6FFDBF65EAFD6111E7" box="[596,697,683,709]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Fig. 13C</figureCitation>
). Cercus with apex pointing (
<figureCitation id="DA872A7EFFC1DD6FFC1065EAFBB811E7" box="[1019,1120,683,709]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Fig. 13C</figureCitation>
). Subgenital plate transverse and at posterior end broadly bilobed (
<figureCitation id="DA872A7EFFC1DD6FFDDC658FFD4711CA" box="[567,671,718,744]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Fig. 13D</figureCitation>
). Ovipositor long sabre-shaped; margins smooth (
<figureCitation id="DA872A7EFFC1DD6FFB23658FFAF411CA" box="[1224,1324,718,744]" captionStart="FIGURE 13" captionStartId="17.[151,250,1793,1817]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-17@17.[151,1436,181,1769]" captionTargetPageId="17" captionText="FIGURE 13. Anthracites sebu sp. nov. ♀ habitus in lateral (A) and dorsal (B) views, abdominal apex in dorsal view (C), subgenital plate in ventral view (D), abdominal apex (including ovipositor) in lateral view (E). Scale bars: 5 mm (A, B, E), 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.13757725" httpUri="https://zenodo.org/record/13757725/files/figure.png" pageId="18" pageNumber="331">Fig. 13E</figureCitation>
).
</paragraph>
<paragraph id="420336FBFFC1DD6FFF2C65B3FB8B1797" blockId="18.[151,1437,151,1277]" pageId="18" pageNumber="331">
Colouration. Body orange (
<figureCitation id="DA872A7EFFC1DD6FFE1D65B3FD89102E" box="[502,593,754,780]" captionStart="FIGURE 1" captionStartId="3.[151,250,1928,1952]" captionTargetBox="[178,1410,181,1904]" captionTargetId="figure-17@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757699" httpUri="https://zenodo.org/record/13757699/files/figure.png" pageId="18" pageNumber="331">Figs 1D</figureCitation>
,
<figureCitation id="DA872A7EFFC1DD6FFDB665B3FD5B102E" box="[605,643,754,780]" captionStart="FIGURE 2" captionStartId="4.[151,250,1797,1821]" captionTargetBox="[178,1409,181,1774]" captionTargetId="figure-19@4.[151,1436,181,1774]" captionTargetPageId="4" captionText="FIGURE 2 Lateral habitus of ♂ Anthracites alatus sp. nov. (A), Anthracites geniculatus Dohrn, 1905 (B), Anthracites pyramidalis Ingrisch, 2015 (C), Anthracites sebu sp. nov. (D) and Anthracites zebra Hebard, 1922 (E). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13757701" httpUri="https://zenodo.org/record/13757701/files/figure.png" pageId="18" pageNumber="331">2D</figureCitation>
). Face with broad black band from fastigium verticis to base of clypeus, widening ventrad and including median ocellus pale (
<figureCitation id="DA872A7EFFC1DD6FFD196454FC80100D" box="[754,856,789,815]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11C</figureCitation>
). Scapus ventrally and on inner side black, dorsally and on outer side pale; antennae orange (
<figureCitation id="DA872A7EFFC1DD6FFD836479FD2E1071" box="[616,758,824,851]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Figs 11A–C</figureCitation>
). Maxillary palpi with segments cream coloured (
<figureCitation id="DA872A7EFFC1DD6FFAC36479FA561071" box="[1320,1422,824,851]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11B</figureCitation>
); mandible black; clypeus basal third black, otherwise yellow orange; labrum yellow orange (
<figureCitation id="DA872A7EFFC1DD6FFB5A641DFAC01054" box="[1201,1304,860,886]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11C</figureCitation>
). Pronotum completely orange yellow without any black patterns; margins of pronotal lateral lobe slightly darker (
<figureCitation id="DA872A7EFFC1DD6FFAC0643EFF10109F" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Figs 11A, 11B</figureCitation>
). Tegmen with lateral field having veins cream coloured and some cells dark (
<figureCitation id="DA872A7EFFC1DD6FFBC064E2FB48109F" box="[1067,1168,931,957]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11B</figureCitation>
); dorsal field with veins and cells unicolourous orange (
<figureCitation id="DA872A7EFFC1DD6FFE176487FDBD10C3" box="[508,613,966,993]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11A</figureCitation>
). Anterior and middle femora unicolourous orange; anterior tibia orange yellow with two black spots, smaller one basal of tympanum and more elongated one distal of tympanum, ventral spines black; middle tibiae orange yellow (
<figureCitation id="DA872A7EFFC1DD6FFD94634CFD3E1705" box="[639,742,1037,1063]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11B</figureCitation>
). Posterior femur orange, knees faintly darker. Posterior tibia pale yellow with spines brown. Tarsus cream coloured with distal margin black. Abdominal tergite pale yellow, with posterior margin having lateral bands that become narrower towards apical segments (
<figureCitation id="DA872A7EFFC1DD6FFB4E6315FAD5174C" box="[1189,1293,1108,1134]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11D</figureCitation>
). Male tenth abdominal tergite, epiproct and cercus generally orange (
<figureCitation id="DA872A7EFFC1DD6FFCE56339FCAC17B0" box="[782,884,1144,1170]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11D</figureCitation>
). Sternites and subgenital plate yellow (
<figureCitation id="DA872A7EFFC1DD6FFAC56339FA4817B0" box="[1326,1424,1144,1170]" captionStart="FIGURE 11" captionStartId="15.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@15.[151,1436,181,1798]" captionTargetPageId="15" captionText="FIGURE 11. Anthracites sebu sp. nov. ♂: anterior half of body in dorsal (A) and lateral (B) views; face in anterior view (C); abdominal apex in dorsal (D), ventral (E) and posterior (F) views. The arrow points to medial process of the subgenital plate. Scale bars: 5 mm (A, B), 2 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.13757721" httpUri="https://zenodo.org/record/13757721/files/figure.png" pageId="18" pageNumber="331">Fig. 11F</figureCitation>
). Ovipositor with valves basally cream coloured, otherwise red brown, at apex darkest.
</paragraph>
<paragraph id="420336FBFFC1DD6FFF2C63FEFDD517FB" blockId="18.[151,1437,151,1277]" box="[199,525,1215,1241]" pageId="18" pageNumber="331">
<emphasis id="70C8EAE9FFC1DD6FFF2C63FEFEA617FB" bold="true" box="[199,382,1215,1241]" pageId="18" pageNumber="331">Measurements.</emphasis>
See
<tableCitation id="0F3E0340FFC1DD6FFE5863FEFDDF17FB" box="[435,519,1215,1241]" captionStart="TABLE 1" captionStartId="22.[151,239,152,176]" captionText="TABLE 1. Measurements of Anthracites, in mm." pageId="18" pageNumber="331">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA66570FFC1DD6FFF2C63A2FD7A17DF" box="[199,674,1251,1277]" pageId="18" pageNumber="331" type="distribution">
<paragraph id="420336FBFFC1DD6FFF2C63A2FD7A17DF" blockId="18.[151,1437,151,1277]" box="[199,674,1251,1277]" pageId="18" pageNumber="331">
<emphasis id="70C8EAE9FFC1DD6FFF2C63A2FEB917DF" bold="true" box="[199,353,1251,1277]" pageId="18" pageNumber="331">Distribution.</emphasis>
<collectingRegion id="8078F819FFC1DD6FFE8C63A2FDC217DF" box="[359,538,1251,1277]" country="Philippines" name="South Cotabato" pageId="18" pageNumber="331">South Cotabato</collectingRegion>
: Lake
<collectingRegion id="8078F819FFC1DD6FFD8863A2FD4717DF" box="[611,671,1251,1277]" country="Philippines" name="Cebu" pageId="18" pageNumber="331">Sebu</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

310
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@ -0,0 +1,310 @@
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<mods:title id="83DAB3196B70AA6F2192BB134AE69547">Two new species of Leptolaimus (Nematoda: Plectida) from Chinese sea area</mods:title>
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<mods:namePart id="A5A9EB66F7494BA4910EC752B684DAB7">Gu, Huimin</mods:namePart>
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<mods:namePart id="FDC90F9EFC5AC440DECB91D309C46B32">Sun, Lingyun</mods:namePart>
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<mods:affiliation id="FE3FBA27BF8DACA67C7F16052B2878B4">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
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<mods:namePart id="8C0DAA2E095995C917E56393CF92C984">Huang, Yong</mods:namePart>
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<subSubSection id="3BA20DFEFF8A347958F0FA78FDEE9DBC" box="[151,514,1482,1509]" pageId="5" pageNumber="389" type="nomenclature">
<paragraph id="73075E75FF8A347958F0FA78FDEE9DBC" blockId="5.[151,514,1482,1545]" box="[151,514,1482,1509]" pageId="5" pageNumber="389">
<heading id="284FE919FF8A347958F0FA78FDEE9DBC" bold="true" box="[151,514,1482,1509]" fontSize="11" level="1" pageId="5" pageNumber="389" reason="1">
<emphasis id="41CC8267FF8A347958F0FA78FDEE9DBC" bold="true" box="[151,514,1482,1509]" pageId="5" pageNumber="389">
<taxonomicName id="B4B825F6FF8A347958F0FA78FE709DBC" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[151,412,1482,1509]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF8A347958F0FA78FE709DBC" bold="true" box="[151,412,1482,1509]" italics="true" pageId="5" pageNumber="389">Leptolaimus hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AFF3F1CFF8A347959C3FA79FDEE9DBC" box="[420,514,1483,1509]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347958F0FA5DFE639E50" box="[151,399,1519,1545]" pageId="5" pageNumber="389" type="description">
<paragraph id="73075E75FF8A347958F0FA5DFE639E50" blockId="5.[151,514,1482,1545]" box="[151,399,1519,1545]" pageId="5" pageNumber="389">
(
<figureCitation id="EB8342F0FF8A347958F8FA5DFEC49E50" box="[159,296,1519,1545]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1891,1915]" captionStartId-1="7.[151,250,1891,1915]" captionTargetBox-0="[247,1341,197,1851]" captionTargetBox-1="[200,1387,181,1866]" captionTargetId-0="figure-18@6.[231,1357,181,1867]" captionTargetId-1="figure-36@7.[200,1387,181,1866]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. Drawings of Leptolaimus hamatus sp. nov. (A) entire view of male holotype; (B) entire view of female paratype, showing reproductive system; (C) anterior end of male holotype, showing cephalic setae, amphidial fovea, cuticle pore and cuticle alae; (D) cloacal region of male holotype, showing spicule, gubernaculum and precloacal supplements. Scale bars: A, 20 µm; B, 30 µm; C, D 10 µm." captionText-1="FIGURE 4. Micrographs of Leptolaimus hamatus sp. nov. (A) anterior end of male holotype, showing cephalic setae, amphidial fovea and first body pore (arrow); (B) posterior region of male 2 paratype, showing tubular supplements; (C) posterior end of male holotype, showing tubular supplements, spicule and gubernaculum; (D) middle region of female paratype, showing vulva (arrow1), ovaries (arrow 2) and eggs (arrow 3). Scale bars: A, C, D 10 µm; B 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13757773" figureDoi-1="http://doi.org/10.5281/zenodo.13757775" httpUri-0="https://zenodo.org/record/13757773/files/figure.png" httpUri-1="https://zenodo.org/record/13757775/files/figure.png" pageId="5" pageNumber="389">Figures 3–4</figureCitation>
,
<tableCitation id="3E3A6BCEFF8A34795954FA5DFE689E50" box="[307,388,1519,1545]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionText="TABLE 1. Individual measurements of Leptolaimus gracilis sp. nov. and L. hamatus sp. nov. (in µm except a, b, c, c′ and V%) a, ratio of body length to maximum body diameter; b, ratio of body length to pharynx length; c, ratio of body length to tail length; c′, ratio of tail length to cloacal or anus body diameter; V%, position of vulva from anterior end expressed as a percentage of total body length; -, no data." pageId="5" pageNumber="389">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347958F0F985FA989F5C" pageId="5" pageNumber="389" type="materials_examined">
<paragraph id="73075E75FF8A347958F0F985FBE49EC0" blockId="5.[151,1436,1590,1905]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958F0F985FDCC9E09" bold="true" box="[151,544,1590,1617]" pageId="5" pageNumber="389">
<typeStatus id="AC03E0D7FF8A347958F0F985FEE89E08" box="[151,260,1591,1617]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
and
<typeStatus id="AC03E0D7FF8A34795926F985FE409E08" box="[321,428,1591,1617]" pageId="5" pageNumber="389" type="paratype">paratype</typeStatus>
material.
</emphasis>
<specimenCount id="65BE95FCFF8A34795A4FF985FD5E9E08" box="[552,690,1591,1617]" count="3" pageId="5" pageNumber="389" type="male">Three males</specimenCount>
and
<specimenCount id="65BE95FCFF8A34795A8CF985FC989E08" box="[747,884,1591,1617]" count="2" pageId="5" pageNumber="389" type="female">two females</specimenCount>
were obtained and measured.
<typeStatus id="AC03E0D7FF8A34795CAAF985FAD99E08" box="[1229,1333,1591,1617]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
(male 1) on slide 3800-1-2-5.
<typeStatus id="AC03E0D7FF8A347959E3F9E9FE0A9E2C" box="[388,486,1627,1653]" pageId="5" pageNumber="389" type="paratype">Paratype</typeStatus>
1 (male 2) on slide 3825-2-2-5,
<typeStatus id="AC03E0D7FF8A34795B31F9E9FC5A9E2C" box="[854,950,1627,1653]" pageId="5" pageNumber="389" type="paratype">paratype</typeStatus>
2 (male 3) on slide 3800-1-2-5,
<typeStatus id="AC03E0D7FF8A34795D41F9E9FA6A9E2C" box="[1318,1414,1627,1653]" pageId="5" pageNumber="389" type="paratype">paratype</typeStatus>
3 (female 1) on slide 3800-1-2-5, and
<typeStatus id="AC03E0D7FF8A34795A4BF9CDFD609EC0" box="[556,652,1663,1689]" pageId="5" pageNumber="389" type="paratype">paratype</typeStatus>
4 (female 2) on slide 3800-1-2-5.
</paragraph>
<paragraph id="73075E75FF8A347958A0F911FA989F5C" blockId="5.[151,1436,1590,1905]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958A0F911FDEC9EE5" bold="true" box="[199,512,1698,1725]" pageId="5" pageNumber="389">Type locality and habitat.</emphasis>
<typeStatus id="AC03E0D7FF8A34795A6DF911FD9E9EE4" box="[522,626,1699,1725]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
and most
<typeStatus id="AC03E0D7FF8A34795A88F911FCB69EE4" box="[751,858,1699,1725]" pageId="5" pageNumber="389" type="paratype">paratypes</typeStatus>
were collected from sea floor muddy sediment at Station 3800-1 (
<geoCoordinate id="168C38B2FF8A34795936F97AFE419EB8" box="[337,429,1735,1763]" degrees="38" direction="north" minutes="0" orientation="latitude" pageId="5" pageNumber="389" precision="925" value="38.0">38°0′ N</geoCoordinate>
,
<geoCoordinate id="168C38B2FF8A347959DCF97AFDC39EB8" box="[443,559,1735,1763]" degrees="122" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="389" precision="925" value="122.5">122°30′ E</geoCoordinate>
, water depth
<quantity id="B440F390FF8A34795AABF975FCE49EB8" box="[716,776,1735,1761]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="389" unit="m" value="46.0">46 m</quantity>
) in the Yellow Sea, and
<specimenCount id="65BE95FCFF8A34795C40F975FB559EB8" box="[1063,1209,1735,1761]" count="1" pageId="5" pageNumber="389" type="generic" typeStatus="paratype">one paratype</specimenCount>
was collected from subtidal muddy sediment at Station 3825-2 (
<geoCoordinate id="168C38B2FF8A34795AEFF95EFD1C9F5C" box="[648,752,1771,1799]" degrees="38" direction="north" minutes="25" orientation="latitude" pageId="5" pageNumber="389" precision="925" value="38.416668">38°25′ N</geoCoordinate>
,
<geoCoordinate id="168C38B2FF8A34795A9BF95EFC819F5C" box="[764,877,1771,1799]" degrees="122" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="389" precision="925" value="122.5">122°30′ E</geoCoordinate>
, water depth
<quantity id="B440F390FF8A34795C64F958FBD19F5C" box="[1027,1085,1770,1797]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="389" unit="m" value="50.0">50 m</quantity>
) in the Yellow Sea in 2008.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347958A0F8BDFE889F14" pageId="5" pageNumber="389" type="etymology">
<paragraph id="73075E75FF8A347958A0F8BDFE889F14" blockId="5.[151,1436,1590,1905]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958A0F8BDFEA29F70" bold="true" box="[199,334,1807,1833]" pageId="5" pageNumber="389">Etymology.</emphasis>
The species epithet comes from the Latin word
<taxonomicName id="B4B825F6FF8A34795B0DF8BDFC259F70" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[874,969,1807,1833]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="hamatus">
<emphasis id="41CC8267FF8A34795B0DF8BDFC259F70" box="[874,969,1807,1833]" italics="true" pageId="5" pageNumber="389">hamatus</emphasis>
</taxonomicName>
, referring to procloacal supplements with hooked distal end.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347458A0F8E5FC649CB0" lastPageId="8" lastPageNumber="392" pageId="5" pageNumber="389" type="description">
<paragraph id="73075E75FF8A347958A0F8E5FC889F28" blockId="5.[151,1436,1590,1905]" box="[199,868,1879,1905]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958A0F8E5FE929F28" bold="true" box="[199,382,1879,1905]" pageId="5" pageNumber="389">Measurements.</emphasis>
All measurement data are given in
<tableCitation id="3E3A6BCEFF8A34795B6DF8E5FCB29F28" box="[778,862,1879,1905]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionText="TABLE 1. Individual measurements of Leptolaimus gracilis sp. nov. and L. hamatus sp. nov. (in µm except a, b, c, c′ and V%) a, ratio of body length to maximum body diameter; b, ratio of body length to pharynx length; c, ratio of body length to tail length; c′, ratio of tail length to cloacal or anus body diameter; V%, position of vulva from anterior end expressed as a percentage of total body length; -, no data." pageId="5" pageNumber="389">Table 1</tableCitation>
.
</paragraph>
<caption id="27C70EFDFF89347A58F0F8D1FE5F9FB1" ID-DOI="http://doi.org/10.5281/zenodo.13757773" ID-Zenodo-Dep="13757773" httpUri="https://zenodo.org/record/13757773/files/figure.png" pageId="6" pageNumber="390" startId="6.[151,250,1891,1915]" targetBox="[247,1341,197,1851]" targetPageId="6" targetType="figure">
<paragraph id="73075E75FF89347A58F0F8D1FE5F9FB1" blockId="6.[151,1437,1891,2024]" pageId="6" pageNumber="390">
<emphasis id="41CC8267FF89347A58F0F8D1FEFF9F22" bold="true" box="[151,275,1891,1915]" pageId="6" pageNumber="390">FIGURE 3.</emphasis>
Drawings of
<taxonomicName id="B4B825F6FF89347A59C7F8D1FD969F22" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[416,634,1891,1915]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="6" pageNumber="390" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF89347A59C7F8D1FD969F22" box="[416,634,1891,1915]" italics="true" pageId="6" pageNumber="390">Leptolaimus hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF89347A5AE7F8D6FD3D9F25" bold="true" box="[640,721,1892,1916]" pageId="6" pageNumber="390">
<taxonomicNameLabel id="5AFF3F1CFF89347A5AE7F8D6FD3D9F25" box="[640,721,1892,1916]" pageId="6" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) entire view of male holotype; (B) entire view of female paratype, showing reproductive system; (C) anterior end of male holotype, showing cephalic setae, amphidial fovea, cuticle pore and cuticle alae; (D) cloacal region of male holotype, showing spicule, gubernaculum and precloacal supplements. Scale bars: A, 20 µm; B, 30 µm; C, D 10 µm.
</paragraph>
</caption>
<caption id="27C70EFDFF88347B58F0F8D1FC199FB1" ID-DOI="http://doi.org/10.5281/zenodo.13757775" ID-Zenodo-Dep="13757775" httpUri="https://zenodo.org/record/13757775/files/figure.png" pageId="7" pageNumber="391" startId="7.[151,250,1891,1915]" targetBox="[200,1387,181,1866]" targetPageId="7" targetType="figure">
<paragraph id="73075E75FF88347B58F0F8D1FC199FB1" blockId="7.[151,1436,1891,2024]" pageId="7" pageNumber="391">
<emphasis id="41CC8267FF88347B58F0F8D1FEE09F22" bold="true" box="[151,268,1891,1915]" pageId="7" pageNumber="391">FIGURE 4</emphasis>
. Micrographs of
<taxonomicName id="B4B825F6FF88347B59DFF8D1FD7C9F22" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[440,656,1891,1915]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="7" pageNumber="391" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF88347B59DFF8D1FD7C9F22" box="[440,656,1891,1915]" italics="true" pageId="7" pageNumber="391">Leptolaimus hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF88347B5AF2F8D1FD089F22" bold="true" box="[661,740,1891,1915]" pageId="7" pageNumber="391">
<taxonomicNameLabel id="5AFF3F1CFF88347B5AF2F8D1FD089F22" box="[661,740,1891,1915]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) anterior end of male holotype, showing cephalic setae, amphidial fovea and first body pore (arrow); (B) posterior region of male 2 paratype, showing tubular supplements; (C) posterior end of male holotype, showing tubular supplements, spicule and gubernaculum; (D) middle region of female paratype, showing vulva (arrow1), ovaries (arrow 2) and eggs (arrow 3). Scale bars: A, C, D 10 µm; B 20 µm.
</paragraph>
</caption>
<paragraph id="73075E75FF87347458F0FF25FEDF98E8" blockId="8.[151,307,151,177]" box="[151,307,151,177]" pageId="8" pageNumber="392">
<emphasis id="41CC8267FF87347458F0FF25FEDF98E8" bold="true" box="[151,307,151,177]" pageId="8" pageNumber="392">Descriptions</emphasis>
</paragraph>
<paragraph id="73075E75FF87347458F0FF6DFF3398A0" blockId="8.[151,1437,223,1006]" box="[151,223,223,249]" pageId="8" pageNumber="392">
<emphasis id="41CC8267FF87347458F0FF6DFF3398A0" bold="true" box="[151,223,223,249]" pageId="8" pageNumber="392">Males</emphasis>
</paragraph>
<paragraph id="73075E75FF87347458F0FEB1FEFC9A94" blockId="8.[151,1437,223,1006]" pageId="8" pageNumber="392">Body slender, tapering anteriorly in pharyngeal region and posteriorly on tail. Cuticle annulated, annuli about 2.0 µm wide at mid-body, without ornamentation. Lateral field present, consists of single alae (two incisures), 3 µm wide at mid-body; originating from anterior third of pharyngeal length, and extending posteriorly to the middle of tail. Numerous body pores present with short and stout setae in sublateral position at both sides of lateral field, and scattered over the body from the pharynx to the proximal part of tail. The first body pore located 22–26 µm from anterior end. Labial region rounded, offset from body contour by a constriction. Inner and outer labial sensilla indistinct. Cephalic setae short, 2–2.5 µm long. Subcephalic sensilla and ocelli absent. Amphidial fovea round, located at level of just post-mid of stoma. Nerve ring surrounding pharynx at level of pre-mid of pharynx, about 40% pharyngeal length from anterior end of the body. Secretory-excretory system not seen. Buccal cavity uniformly tubular. Pharynx cylindrical, with distinct oval basal bulb. Cardia cylindrical. Tail conico-cylindrical with a slightly swollen tip. A pair of precloacal subventral setae 3–4 µm long, 5 µm anterior to cloaca. Two to three pairs of subventral and two pairs of sundorsal caudal setae 3–4 µm long. Three caudal glands present. Spinneret functional.</paragraph>
<paragraph id="73075E75FF87347458A0FD65FECB9BB7" blockId="8.[151,1437,223,1006]" pageId="8" pageNumber="392">Reproductive system diorchic; anterior testis outstretched, posterior one reflexed. Spicules paired, knife-like, with broad proximal half part and narrow curved distal half part. Gubernaculum plate-like, with a straight dorsal apophysis. Accessory apparatus composed of one midventral precloacal papilliform sensillum located on anterior cloacal lip, six midventral tubular supplements extending for 120–130 µm from cloaca towards anterior end; alveolar supplements absent. Tubular supplements slightly curved, rounded proximal ends and hooked distal ends, 11–14 µm long, weakly S-shaped. Posteriormost tubular supplement 21 µm anterior to cloaca, anterior to spicules, the space between adjacent supplements increasing from the posteriormost supplement to the anteriormost one, varying from 13 to 23 µm.</paragraph>
<paragraph id="73075E75FF87347458F0FBA9FF159C6C" blockId="8.[151,1437,1051,2014]" box="[151,249,1051,1077]" pageId="8" pageNumber="392">
<emphasis id="41CC8267FF87347458F0FBA9FF159C6C" bold="true" box="[151,249,1051,1077]" pageId="8" pageNumber="392">Females</emphasis>
</paragraph>
<paragraph id="73075E75FF87347458F0FB8DFC649CB0" blockId="8.[151,1437,1051,2014]" pageId="8" pageNumber="392">Similar to males in most morphological characters, except tail longer (c′&gt; 7.5) without setae, and supplements absent. Reproductive system didelphic, with two opposed, reflexed ovaries, the anterior ovary located to the right side of the intestine, and the posterior one located to the left side of the intestine. Oviduct a narrow tube. Spermathecae indistinct. Vagina straight, 0.3 times vulval body diameter long. Vulva right ventrosublateral, located in the middle of the body, 48–50% of the total body length from the anterior end.</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF87347458A0FB41FF399D98" pageId="8" pageNumber="392" type="diagnosis">
<paragraph id="73075E75FF87347458A0FB41FF399D98" blockId="8.[151,1437,1051,2014]" pageId="8" pageNumber="392">
<emphasis id="41CC8267FF87347458A0FB41FEAE9D54" bold="true" box="[199,322,1267,1293]" pageId="8" pageNumber="392">Diagnosis.</emphasis>
<taxonomicName id="B4B825F6FF873474592DFB41FDD19D54" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[330,573,1267,1293]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF873474592DFB41FDD19D54" box="[330,573,1267,1293]" italics="true" pageId="8" pageNumber="392">Leptolaimus hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8734745A21FB41FD4E9D54" bold="true" box="[582,674,1267,1293]" pageId="8" pageNumber="392">
<taxonomicNameLabel id="5AFF3F1CFF8734745A21FB41FD4E9D54" box="[582,674,1267,1293]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by body 624–690 µm long; rounded labial region offset from body contour; cephalic setae 2–3 µm long; amphidial fovea located 12–13 µm from anterior end; first body pore located 22–26 µm from anterior end; males with six tubular and without alveolar supplements, tubular supplements hooked distally; spicules slightly curved, knife-like and 22–24 µm long; gubernaculum with a straight dorsal apophysis; tail conico-cylindrical; females without supplements, vulva situated at right subventral side of body.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF87347558A0FA79FB8099D0" lastPageId="9" lastPageNumber="393" pageId="8" pageNumber="392" type="discussion">
<paragraph id="73075E75FF87347558A0FA79FB8099D0" blockId="8.[151,1437,1051,2014]" lastBlockId="9.[151,1437,150,393]" lastPageId="9" lastPageNumber="393" pageId="8" pageNumber="392">
<emphasis id="41CC8267FF87347458A0FA79FED79DBC" bold="true" box="[199,315,1483,1509]" pageId="8" pageNumber="392">Remarks.</emphasis>
<taxonomicName id="B4B825F6FF8734745925FA79FDDF9DBC" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[322,563,1483,1509]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF8734745925FA79FDDF9DBC" box="[322,563,1483,1509]" italics="true" pageId="8" pageNumber="392">Leptolaimus hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8734745A5EFA79FD7F9DBC" bold="true" box="[569,659,1483,1509]" pageId="8" pageNumber="392">
<taxonomicNameLabel id="5AFF3F1CFF8734745A5EFA79FD7F9DBC" box="[569,659,1483,1509]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="B4B825F6FF8734745B42FA7EFB1F9DBC" authority="Holovachov &amp; Bostrom, 2013" authorityName="Holovachov &amp; Bostrom" authorityYear="2013" box="[805,1267,1483,1510]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="octavus">
<emphasis id="41CC8267FF8734745B42FA7EFC759DBC" box="[805,921,1484,1509]" italics="true" pageId="8" pageNumber="392">L. octavus</emphasis>
Holovachov &amp; Boström, 2013
</taxonomicName>
and
<taxonomicName id="B4B825F6FF8734745D4EFA7EFCA09E53" authority="(Schuurmans-Stekhoven &amp; De Coninck, 1933) Gerlach, 1958" authorityName="Gerlach" authorityYear="1958" baseAuthorityName="Schuurmans-Stekhoven &amp; De Coninck" baseAuthorityYear="1933" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="elegans">
<emphasis id="41CC8267FF8734745D4EFA7EFA719DBC" box="[1321,1437,1483,1509]" italics="true" pageId="8" pageNumber="392">L. elegans</emphasis>
(Schuurmans-Stekhoven &amp; De Coninck, 1933) Gerlach,
<quantity id="B440F390FF8734745B73FA5DFCA09E53" box="[788,844,1519,1546]" metricMagnitude="1" metricUnit="m" metricValue="4.973319999999999" pageId="8" pageNumber="392" unit="in" value="1958.0">1958</quantity>
</taxonomicName>
in tubular supplements weakly S-shaped, but differs from
<taxonomicName id="B4B825F6FF87347458B2F9A6FEA59E74" authorityName="Holovachov &amp; Bostrom" authorityYear="2013" box="[213,329,1556,1581]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="octavus">
<emphasis id="41CC8267FF87347458B2F9A6FEA59E74" box="[213,329,1556,1581]" italics="true" pageId="8" pageNumber="392">L. octavus</emphasis>
</taxonomicName>
by supplements hooked distally (
<emphasis id="41CC8267FF8734745AA2F9A6FD309E74" box="[709,732,1556,1581]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
strongly cuticularized distally with dentate tips in
<taxonomicName id="B4B825F6FF8734745D7FF9A6FA609E74" authorityName="Holovachov &amp; Bostrom" authorityYear="2013" box="[1304,1420,1556,1581]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="octavus">
<emphasis id="41CC8267FF8734745D7FF9A6FA609E74" box="[1304,1420,1556,1581]" italics="true" pageId="8" pageNumber="392">L. octavus</emphasis>
</taxonomicName>
), spicules knife-like (
<emphasis id="41CC8267FF873474591FF98AFE639E08" box="[376,399,1592,1617]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
slender), vulva right-subventral (
<emphasis id="41CC8267FF8734745B6CF98AFCCE9E08" box="[779,802,1592,1617]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
midventral). Additionally
<taxonomicName id="B4B825F6FF8734745C37F98AFB299E08" authorityName="Holovachov &amp; Bostrom" authorityYear="2013" box="[1104,1221,1592,1617]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="octavus">
<emphasis id="41CC8267FF8734745C37F98AFB299E08" box="[1104,1221,1592,1617]" italics="true" pageId="8" pageNumber="392">L. octavus</emphasis>
</taxonomicName>
is the only species within
<taxonomicName id="B4B825F6FF8734745880F9E9FE9D9E2C" authorityName="de Man" authorityYear="1876" box="[231,369,1627,1653]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="genus">
<emphasis id="41CC8267FF8734745880F9E9FE9D9E2C" box="[231,369,1627,1653]" italics="true" pageId="8" pageNumber="392">Leptolaimus</emphasis>
</taxonomicName>
in having a bursa-like expansion of the lateral field along the proximal part of the tail of both females and males. The new species differs from
<taxonomicName id="B4B825F6FF8734745AAEF932FCD29EC0" authorityName="Gerlach" authorityYear="1958" baseAuthorityName="Schuurmans-Stekhoven &amp; De Coninck" baseAuthorityYear="1933" box="[713,830,1663,1689]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="elegans">
<emphasis id="41CC8267FF8734745AAEF932FCD29EC0" box="[713,830,1663,1689]" italics="true" pageId="8" pageNumber="392">L. elegans</emphasis>
</taxonomicName>
by different shape of supplements (hook-like distally with point tips
<emphasis id="41CC8267FF873474592AF916FE889EE4" box="[333,356,1700,1725]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
knot-like distally with dentate tips), and different shape of gubernaculum (plate-like with a straight dorsal apophysis
<emphasis id="41CC8267FF87347459A6F97AFE349EB8" box="[449,472,1736,1761]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
triangular with a pointed dorsal apophysis). The new species is also similar to
<taxonomicName id="B4B825F6FF8734745DE1F97AFE399F5C" authority="Platt, 1973" authorityName="Platt" authorityYear="1973" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="septempapillatus">
<emphasis id="41CC8267FF8734745DE1F97AFEB99F5C" italics="true" pageId="8" pageNumber="392">L. septempapillatus</emphasis>
<bibRefCitation id="17292384FF873474593BF959FE399F5C" author="Platt, H. M." box="[348,469,1771,1797]" pageId="8" pageNumber="392" pagination="295 - 321" refId="ref5328" refString="Platt, H. M. (1973) Freeliving marine nematodes from Strangford Lough, Northern Ireland. Cahiers de Biologie Marine, 14 (3), 295 - 321." type="journal article" year="1973">
Platt,
<quantity id="B440F390FF87347459FAF959FE399F5C" box="[413,469,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="5.011419999999999" pageId="8" pageNumber="392" unit="in" value="1973.0">1973</quantity>
</bibRefCitation>
</taxonomicName>
in having knife-like spicules and similar number of tubular supplements, but it differs from
<taxonomicName id="B4B825F6FF87347458B0F8A2FE599F70" authorityName="Platt" authorityYear="1973" box="[215,437,1807,1833]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="septempapillatus">
<emphasis id="41CC8267FF87347458B0F8A2FE599F70" box="[215,437,1807,1833]" italics="true" pageId="8" pageNumber="392">L. septempapillatus</emphasis>
</taxonomicName>
by supplements hook-like distally without cuticular collars around their distal ends (
<emphasis id="41CC8267FF8734745DE2F8A2FA709F70" box="[1413,1436,1808,1833]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
knot-like distally with dentate tips and with cuticular collars around their distal ends), and vulva right ventrosublateral (
<emphasis id="41CC8267FF87347458C6F8EAFF549F28" box="[161,184,1880,1905]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
midventral in the latter species). The new species is also similar to
<taxonomicName id="B4B825F6FF8734745BD3F8EAFA709F28" authority="Villares &amp; Pastor de Ward, 2012" authorityName="Villares &amp; Pastor de Ward" authorityYear="2012" box="[948,1436,1879,1906]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="gabinoi">
<emphasis id="41CC8267FF8734745BD3F8EAFBC49F28" box="[948,1064,1879,1905]" italics="true" pageId="8" pageNumber="392">L. gabinoi</emphasis>
Villares &amp; Pastor de Ward, 2012
</taxonomicName>
and
<taxonomicName id="B4B825F6FF87347458AFF8CEFE109FCC" authority="de Man, 1876" authorityName="de Man" authorityYear="1876" box="[200,508,1915,1942]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="papilliger">
<emphasis id="41CC8267FF87347458AFF8CEFEB99FCC" box="[200,341,1915,1941]" italics="true" pageId="8" pageNumber="392">L. papilliger</emphasis>
de Man,
<quantity id="B440F390FF87347459A3F8C9FE109FCC" box="[452,508,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="4.76504" pageId="8" pageNumber="392" unit="in" value="1876.0">1876</quantity>
</taxonomicName>
in having tips of supplements hook-like, but it differs from
<taxonomicName id="B4B825F6FF8734745CCAF8CEFACE9FCC" authorityName="Villares &amp; Pastor de Ward" authorityYear="2012" box="[1197,1314,1915,1941]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="gabinoi">
<emphasis id="41CC8267FF8734745CCAF8CEFACE9FCC" box="[1197,1314,1915,1941]" italics="true" pageId="8" pageNumber="392">L. gabinoi</emphasis>
</taxonomicName>
by shorter cephalic setae (2–3 µm
<emphasis id="41CC8267FF87347459CAF812FE289FE0" box="[429,452,1952,1977]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
5–6 µm long) and more numerous tubular precloacal supplements (6
<emphasis id="41CC8267FF8734745C97F812FAEB9FE0" box="[1264,1287,1952,1977]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
3); it differs from
<taxonomicName id="B4B825F6FF87347458B1F876FE8F9F84" authorityName="de Man" authorityYear="1876" box="[214,355,1987,2013]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="8" pageNumber="392" phylum="Nematoda" rank="species" species="papilliger">
<emphasis id="41CC8267FF87347458B1F876FE8F9F84" box="[214,355,1987,2013]" italics="true" pageId="8" pageNumber="392">L. papilliger</emphasis>
</taxonomicName>
by setiform cephalic setae (
<emphasis id="41CC8267FF8734745AC1F876FD519F84" box="[678,701,1988,2013]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
papilliform cephalic setae) and lack of alveolar supplements (
<emphasis id="41CC8267FF8734745DE2F876FA709F84" box="[1413,1436,1988,2013]" italics="true" pageId="8" pageNumber="392">vs</emphasis>
presence of 12–15 alveolar supplements in
<taxonomicName id="B4B825F6FF8634755A18FF25FCE798E8" authorityName="de Man" authorityYear="1876" box="[639,779,151,177]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="9" pageNumber="393" phylum="Nematoda" rank="species" species="papilliger">
<emphasis id="41CC8267FF8634755A18FF25FCE798E8" box="[639,779,151,177]" italics="true" pageId="9" pageNumber="393">L. papilliger</emphasis>
</taxonomicName>
). The new species furthermore differs from
<taxonomicName id="B4B825F6FF8634755D67FF25FA9898E8" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[1280,1396,151,177]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="9" pageNumber="393" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8634755D67FF25FA9898E8" box="[1280,1396,151,177]" italics="true" pageId="9" pageNumber="393">L. gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8634755D1CFF25FF24988C" bold="true" pageId="9" pageNumber="393">
<taxonomicNameLabel id="5AFF3F1CFF8634755D1CFF25FF24988C" pageId="9" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by shorter cephalic setae (2-3 µm
<emphasis id="41CC8267FF8634755A29FF09FD89988D" box="[590,613,187,212]" italics="true" pageId="9" pageNumber="393">vs</emphasis>
5–6 µm long), conico-cylindrical tail (
<emphasis id="41CC8267FF8634755C7BFF09FBDF988D" box="[1052,1075,187,212]" italics="true" pageId="9" pageNumber="393">vs</emphasis>
elongated conical tail), spicules knife-like, and six weakly S-shaped precloacal supplements with hooked distal ends (
<emphasis id="41CC8267FF8634755C02FF6DFB9098A1" box="[1125,1148,223,248]" italics="true" pageId="9" pageNumber="393">vs</emphasis>
shorter arcuated spicules and eight sickle-shaped supplements). The new species distinguishes from other known species by combination of characters of cephalic setae 2–3 µm long, males with six tubular precloacal supplements with hooked distal ends, without alveolar supplement, spicules knife-like, 22–24 µm long, gubernaculum with a straight dorso-caudal apophysis, females without supplements, vulva right subventral, tail conico-cylindrical.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="F2FC716FCFCB64AD7BE42C1B8036C6E7" ID-DOI="10.11646/zootaxa.5507.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757765" ID-ZooBank="732EF0D1-7CDA-4366-B164-156D4E2E8E7F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726221687458" checkinUser="plazi" docAuthor="Gu, Huimin, Sun, Lingyun &amp; Huang, Yong" docDate="2024" docId="FB11EF63FF8E347958F0FA14FECB9D20" docLanguage="en" docName="zootaxa.5507.2.9.pdf" docOrigin="Zootaxa 5507 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.2.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptolaimus gracilis Gu, Sun &amp; Huang, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="389" masterDocId="0728971BFF8F347C5867FFB2FFEC9859" masterDocTitle="Two new species of Leptolaimus (Nematoda: Plectida) from Chinese sea area" masterLastPageNumber="394" masterPageNumber="384" pageNumber="385" updateTime="1726222090683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CF595BEC00064DCDBE01E8B7D8213757">Two new species of Leptolaimus (Nematoda: Plectida) from Chinese sea area</mods:title>
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<mods:namePart id="C31742A54BB7247D3850D66B65C3D108">Gu, Huimin</mods:namePart>
<mods:nameIdentifier id="3A136CDDCDD4264D6ED8132000956161" type="ORCID">0009-0008-5820-4678</mods:nameIdentifier>
<mods:affiliation id="5403FB60A0EBC8151013D68DD0369EBE">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
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<mods:namePart id="3AB89A94D2E3999E8C99B5F87F68E4BC">Sun, Lingyun</mods:namePart>
<mods:nameIdentifier id="0C72F6FFFF286BDCA2464B7BA09914E8" type="ORCID">0009-0005-3553-3645</mods:nameIdentifier>
<mods:affiliation id="97E5385ED267FF77D2694629BE4CBD1F">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
<mods:nameIdentifier id="9D92B59AA4646602644E13336BE5CC1B" type="email">981779432@qq.com</mods:nameIdentifier>
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<mods:namePart id="CEAEFEFD6872513A210A916BF35BB455">Huang, Yong</mods:namePart>
<mods:affiliation id="280A92F8E5262680EDFA41EBA397BD13">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
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<mods:date id="DCECB1AB77DA0AAC53AFFF30BAAA6DC0">2024</mods:date>
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<treatment id="FB11EF63FF8E347958F0FA14FECB9D20" LSID="urn:lsid:plazi:treatment:FB11EF63FF8E347958F0FA14FECB9D20" httpUri="http://treatment.plazi.org/id/FB11EF63FF8E347958F0FA14FECB9D20" lastPageId="5" lastPageNumber="389" pageId="1" pageNumber="385">
<subSubSection id="3BA20DFEFF8E347D58F0FA14FE1F9D98" box="[151,499,1446,1473]" pageId="1" pageNumber="385" type="nomenclature">
<paragraph id="73075E75FF8E347D58F0FA14FE1F9D98" blockId="1.[151,499,1446,1509]" box="[151,499,1446,1473]" pageId="1" pageNumber="385">
<heading id="284FE919FF8E347D58F0FA14FE1F9D98" bold="true" box="[151,499,1446,1473]" fontSize="11" level="1" pageId="1" pageNumber="385" reason="1">
<emphasis id="41CC8267FF8E347D58F0FA14FE1F9D98" bold="true" box="[151,499,1446,1473]" pageId="1" pageNumber="385">
<taxonomicName id="B4B825F6FF8E347D58F0FA14FE629D98" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[151,398,1446,1473]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="1" pageNumber="385" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8E347D58F0FA14FE629D98" bold="true" box="[151,398,1446,1473]" italics="true" pageId="1" pageNumber="385">Leptolaimus gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AFF3F1CFF8E347D59F2FA15FE1F9D98" box="[405,499,1447,1473]" pageId="1" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="3BA20DFEFF8E347D58F0FA79FE639DBC" box="[151,399,1483,1509]" pageId="1" pageNumber="385" type="description">
<paragraph id="73075E75FF8E347D58F0FA79FE639DBC" blockId="1.[151,499,1446,1509]" box="[151,399,1483,1509]" pageId="1" pageNumber="385">
(
<figureCitation id="EB8342F0FF8E347D58F8FA79FEC49DBC" box="[159,296,1483,1509]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1908,1932]" captionStartId-1="3.[151,250,1856,1880]" captionTargetBox-0="[203,1384,185,1877]" captionTargetBox-1="[256,1331,181,1832]" captionTargetId-0="figure-18@2.[197,1390,179,1883]" captionTargetId-1="figure-36@3.[256,1331,181,1832]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Drawings of Leptolaimus gracilis sp. nov. (A) entire view of male holotype; (B) anterior end of male holotype, showing cephalic setae, buccal cavity and amphidial fovea; (C) entire view of female paratype, showing reproductive system; (D) cloacal region of male holotype, showing spicule, gubernaculum and precloacal supplement. Scale bars: A, C 20 µm; B, D 10 µm." captionText-1="FIGURE 2. Micrographs of Leptolaimus gracilis sp. nov. (A) pharygeal region of male holotype, showing posterior bulb; (B) anterior end of male holotype, showing buccal cavity (arrows); (C) anterior end of female paratype, showing cephalic setae, amphid and buccal cavity (arrow); (D) precloacal region of male 2 paratype, showing precloacal supplements; (E) spicule and gubernaculum of male holotype; (F) middle portion of female paratype, showing vulva (arrow 1), ovary (arrow 2) and egg (arrow 3). Scale bars:10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13757767" figureDoi-1="http://doi.org/10.5281/zenodo.13757769" httpUri-0="https://zenodo.org/record/13757767/files/figure.png" httpUri-1="https://zenodo.org/record/13757769/files/figure.png" pageId="1" pageNumber="385">Figures 1–2</figureCitation>
,
<tableCitation id="3E3A6BCEFF8E347D5954FA79FE689DBC" box="[307,388,1483,1509]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionText="TABLE 1. Individual measurements of Leptolaimus gracilis sp. nov. and L. hamatus sp. nov. (in µm except a, b, c, c′ and V%) a, ratio of body length to maximum body diameter; b, ratio of body length to pharynx length; c, ratio of body length to tail length; c′, ratio of tail length to cloacal or anus body diameter; V%, position of vulva from anterior end expressed as a percentage of total body length; -, no data." pageId="1" pageNumber="385">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8E347D58F0F9A1FB1A9EC0" pageId="1" pageNumber="385" type="materials_examined">
<paragraph id="73075E75FF8E347D58F0F9A1FAD19E08" blockId="1.[151,1436,1554,1797]" pageId="1" pageNumber="385">
<emphasis id="41CC8267FF8E347D58F0F9A1FDF09E75" bold="true" box="[151,540,1554,1581]" pageId="1" pageNumber="385">
<typeStatus id="AC03E0D7FF8E347D58F0F9A1FEE89E74" box="[151,260,1555,1581]" pageId="1" pageNumber="385" type="holotype">Holotype</typeStatus>
and
<typeStatus id="AC03E0D7FF8E347D5959F9A1FE459E74" box="[318,425,1555,1581]" pageId="1" pageNumber="385" type="paratype">paratype</typeStatus>
material.
</emphasis>
<specimenCount id="65BE95FCFF8E347D5A45F9A1FD479E74" box="[546,683,1555,1581]" count="3" pageId="1" pageNumber="385" type="male">Three males</specimenCount>
and
<specimenCount id="65BE95FCFF8E347D5A86F9A1FCB09E74" box="[737,860,1555,1581]" count="1" pageId="1" pageNumber="385" type="female">one female</specimenCount>
were obtained and measured.
<typeStatus id="AC03E0D7FF8E347D5CC9F9A1FAFA9E74" box="[1198,1302,1555,1581]" pageId="1" pageNumber="385" type="holotype">Holotype</typeStatus>
(male 1) on slide SPG2-5-3.
<typeStatus id="AC03E0D7FF8E347D5936F985FE5F9E08" box="[337,435,1591,1617]" pageId="1" pageNumber="385" type="paratype">Paratype</typeStatus>
1 (male 2),
<typeStatus id="AC03E0D7FF8E347D5A5CF985FD779E08" box="[571,667,1591,1617]" pageId="1" pageNumber="385" type="paratype">paratype</typeStatus>
2 (male 3) and
<typeStatus id="AC03E0D7FF8E347D5B2BF985FC409E08" box="[844,940,1591,1617]" pageId="1" pageNumber="385" type="paratype">paratype</typeStatus>
3 (female 1) all on slide SPG3-5-7.
</paragraph>
<paragraph id="73075E75FF8E347D58A0F9E9FB1A9EC0" blockId="1.[151,1436,1554,1797]" pageId="1" pageNumber="385">
<emphasis id="41CC8267FF8E347D58A0F9E9FE179E2D" bold="true" box="[199,507,1626,1653]" pageId="1" pageNumber="385">Type locality and habitat.</emphasis>
<materialsCitation id="C3D05428FF8E347D5A64F9E9FB1F9EC0" collectingDate="2023-05" country="China" county="Sea" latitude="27.862501" location="Dongtou Island of Wenzhou City" longLatPrecision="20" longitude="121.151115" municipality="East" pageId="1" pageNumber="385" specimenCount="2" typeStatus="holotype">
<typeStatus id="AC03E0D7FF8E347D5A64F9E9FD879E2C" box="[515,619,1627,1653]" pageId="1" pageNumber="385" type="holotype">Holotype</typeStatus>
and
<typeStatus id="AC03E0D7FF8E347D5AC3F9E9FCE39E2C" box="[676,783,1627,1653]" pageId="1" pageNumber="385" type="paratype">paratypes</typeStatus>
were all collected from intertidal fine sandy sediments at
<location id="766708AEFF8E347D58F0F9CDFDE59EC0" LSID="urn:lsid:plazi:treatment:FB11EF63FF8E347958F0FA14FECB9D20:766708AEFF8E347D58F0F9CDFDE59EC0" box="[151,521,1663,1689]" country="China" county="Sea" latitude="27.862501" longLatPrecision="20" longitude="121.151115" municipality="East" name="Dongtou Island of Wenzhou City" pageId="1" pageNumber="385">Dongtou Island of Wenzhou City</location>
along the
<collectingMunicipality id="9363C40FFF8E347D5A18F9CDFD439EC0" box="[639,687,1663,1689]" pageId="1" pageNumber="385">East</collectingMunicipality>
<collectingCountry id="0BAF1EE5FF8E347D5AD1F9CDFD159EC0" box="[694,761,1663,1689]" name="China" pageId="1" pageNumber="385">China</collectingCountry>
<collectingCounty id="9A6626F9FF8E347D5B67F9CDFCC49EC0" box="[768,808,1663,1689]" pageId="1" pageNumber="385">Sea</collectingCounty>
(
<geoCoordinate id="168C38B2FF8E347D5B5FF932FC249EC0" box="[824,968,1663,1691]" degrees="27" direction="north" minutes="51" orientation="latitude" pageId="1" pageNumber="385" precision="15" seconds="45" value="27.862501">27°51′45″ N</geoCoordinate>
;
<geoCoordinate id="168C38B2FF8E347D5BB3F932FBBC9EC0" box="[980,1104,1663,1691]" degrees="121" direction="east" minutes="9" orientation="longitude" pageId="1" pageNumber="385" precision="15" seconds="4" value="121.151115">121°9′4″ E</geoCoordinate>
) in
<date id="070678B5FF8E347D5C1BF9CDFB1F9EC0" box="[1148,1267,1663,1689]" pageId="1" pageNumber="385" value="2023-05">
<collectingDate id="1742815DFF8E347D5C1BF9CDFB1F9EC0" box="[1148,1267,1663,1689]" pageId="1" pageNumber="385" value="2023-05">May 2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8E347D58A0F911FF399EB8" pageId="1" pageNumber="385" type="etymology">
<paragraph id="73075E75FF8E347D58A0F911FF399EB8" blockId="1.[151,1436,1554,1797]" pageId="1" pageNumber="385">
<emphasis id="41CC8267FF8E347D58A0F911FEA29EE4" bold="true" box="[199,334,1699,1725]" pageId="1" pageNumber="385">Etymology.</emphasis>
The species epithet comes from the Latin word
<taxonomicName id="B4B825F6FF8E347D5B08F911FC299EE4" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[879,965,1699,1725]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="1" pageNumber="385" phylum="Nematoda" rank="species" species="gracilis">
<emphasis id="41CC8267FF8E347D5B08F911FC299EE4" box="[879,965,1699,1725]" italics="true" pageId="1" pageNumber="385">gracilis</emphasis>
</taxonomicName>
, referring to the new species with slender body.
</paragraph>
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<subSubSection id="3BA20DFEFF8E347958A0F959FD3F99AC" lastPageId="5" lastPageNumber="389" pageId="1" pageNumber="385" type="description">
<paragraph id="73075E75FF8E347D58A0F959FC889F5C" blockId="1.[151,1436,1554,1797]" box="[199,868,1771,1797]" pageId="1" pageNumber="385">
<emphasis id="41CC8267FF8E347D58A0F959FE929F5C" bold="true" box="[199,382,1771,1797]" pageId="1" pageNumber="385">Measurements.</emphasis>
All measurement data are given in
<tableCitation id="3E3A6BCEFF8E347D5B6DF959FCB29F5C" box="[778,862,1771,1797]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionText="TABLE 1. Individual measurements of Leptolaimus gracilis sp. nov. and L. hamatus sp. nov. (in µm except a, b, c, c′ and V%) a, ratio of body length to maximum body diameter; b, ratio of body length to pharynx length; c, ratio of body length to tail length; c′, ratio of tail length to cloacal or anus body diameter; V%, position of vulva from anterior end expressed as a percentage of total body length; -, no data." pageId="1" pageNumber="385">Table 1</tableCitation>
.
</paragraph>
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<paragraph id="73075E75FF8D347E58F0F8C6FF0C9FA0" blockId="2.[151,1437,1908,2041]" pageId="2" pageNumber="386">
<emphasis id="41CC8267FF8D347E58F0F8C6FEF99FD5" bold="true" box="[151,277,1908,1932]" pageId="2" pageNumber="386">FIGURE 1.</emphasis>
Drawings of
<taxonomicName id="B4B825F6FF8D347E59C4F8C7FD9A9FD4" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[419,630,1909,1933]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="2" pageNumber="386" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8D347E59C4F8C7FD9A9FD4" box="[419,630,1909,1933]" italics="true" pageId="2" pageNumber="386">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8D347E5A19F8C7FD3C9FD4" bold="true" box="[638,720,1909,1933]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="5AFF3F1CFF8D347E5A19F8C7FD3C9FD4" box="[638,720,1909,1933]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) entire view of male holotype; (B) anterior end of male holotype, showing cephalic setae, buccal cavity and amphidial fovea; (C) entire view of female paratype, showing reproductive system; (D) cloacal region of male holotype, showing spicule, gubernaculum and precloacal supplement. Scale bars: A, C 20 µm; B, D 10 µm.
</paragraph>
</caption>
<caption id="27C70EFDFF8C347F58F0F8F2FE569FB0" ID-DOI="http://doi.org/10.5281/zenodo.13757769" ID-Zenodo-Dep="13757769" httpUri="https://zenodo.org/record/13757769/files/figure.png" pageId="3" pageNumber="387" startId="3.[151,250,1856,1880]" targetBox="[256,1331,181,1832]" targetPageId="3" targetType="figure">
<paragraph id="73075E75FF8C347F58F0F8F2FE569FB0" blockId="3.[151,1437,1856,2025]" pageId="3" pageNumber="387">
<emphasis id="41CC8267FF8C347F58F0F8F2FEE19F01" bold="true" box="[151,269,1856,1880]" pageId="3" pageNumber="387">FIGURE 2</emphasis>
. Micrographs of
<taxonomicName id="B4B825F6FF8C347F59D9F8F2FD639F01" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[446,655,1856,1880]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="3" pageNumber="387" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8C347F59D9F8F2FD639F01" box="[446,655,1856,1880]" italics="true" pageId="3" pageNumber="387">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8C347F5AF1F8F2FD0B9F01" bold="true" box="[662,743,1856,1880]" pageId="3" pageNumber="387">
<taxonomicNameLabel id="5AFF3F1CFF8C347F5AF1F8F2FD0B9F01" box="[662,743,1856,1880]" pageId="3" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) pharygeal region of male holotype, showing posterior bulb; (B) anterior end of male holotype, showing buccal cavity (arrows); (C) anterior end of female paratype, showing cephalic setae, amphid and buccal cavity (arrow); (D) precloacal region of male 2 paratype, showing precloacal supplements; (E) spicule and gubernaculum of male holotype; (F) middle portion of female paratype, showing vulva (arrow 1), ovary (arrow 2) and egg (arrow 3). Scale bars:10 µm.
</paragraph>
</caption>
<caption id="27C70EFDFF8B347858F0FF25FD609942" pageId="4" pageNumber="388" startId="4.[151,239,151,175]" targetType="table">
<paragraph id="73075E75FF8B347858F0FF25FD609942" blockId="4.[151,1437,150,284]" pageId="4" pageNumber="388">
<emphasis id="41CC8267FF8B347858F0FF25FEE598F6" bold="true" box="[151,265,151,175]" pageId="4" pageNumber="388">TABLE 1.</emphasis>
Individual measurements of
<taxonomicName id="B4B825F6FF8B34785A5AFF25FCF498F6" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[573,792,151,175]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="4" pageNumber="388" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8B34785A5AFF25FCF498F6" box="[573,792,151,175]" italics="true" pageId="4" pageNumber="388">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8B34785B79FF2AFC9F98E9" bold="true" box="[798,883,152,176]" pageId="4" pageNumber="388">
<taxonomicNameLabel id="5AFF3F1CFF8B34785B79FF2AFC9F98E9" box="[798,883,152,176]" pageId="4" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="B4B825F6FF8B34785BC2FF2AFBF698F6" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[933,1050,151,175]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="4" pageNumber="388" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF8B34785BC2FF2AFBF698F6" box="[933,1050,151,175]" italics="true" pageId="4" pageNumber="388">L. hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8B34785C47FF2AFB9998E9" bold="true" box="[1056,1141,152,176]" pageId="4" pageNumber="388">
<taxonomicNameLabel id="5AFF3F1CFF8B34785C47FF2AFB9998E9" box="[1056,1141,152,176]" pageId="4" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in µm except a, b, c, c′ and V%) a, ratio of body length to maximum body diameter; b, ratio of body length to pharynx length; c, ratio of body length to tail length; c′, ratio of tail length to cloacal or anus body diameter; V%, position of vulva from anterior end expressed as a percentage of total body length; -, no data.
</paragraph>
</caption>
<paragraph id="73075E75FF8B347858F8FEF2FA9C9CAB" pageId="4" pageNumber="388">
<table id="01B8ACD5FF8BCB8358F8FE9FFA909CAB" box="[159,1404,301,1266]" gridcols="10" gridrows="24" pageId="4" pageNumber="388">
<tr id="CD885C37FF8BCB8358F8FE9FFA909930" box="[159,1404,301,361]" gridrow="0" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FE9FFE039930" box="[159,495,301,361]" gridcol="0" gridrow="0" pageId="4" pageNumber="388">Characters</th>
<th id="8E59354BFF8BCB835AE3FE9FFC2D9930" box="[644,961,301,361]" colspan="4" colspanRight="3" gridcol="1" gridrow="0" pageId="4" pageNumber="388">
<taxonomicName id="B4B825F6FF8B34785AE3FE9FFCA5991B" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[644,841,301,322]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="4" pageNumber="388" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8B34785AE3FE9FFCA5991B" box="[644,841,301,322]" italics="true" pageId="4" pageNumber="388">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8B34785B37FE9FFC70991B" bold="true" box="[848,924,301,322]" pageId="4" pageNumber="388">
<taxonomicNameLabel id="5AFF3F1CFF8B34785B37FE9FFC70991B" box="[848,924,301,322]" pageId="4" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂1 ♂2 ♂3 ♀1
</th>
<th id="8E59354BFF8BCB835B8DFE9FFAC89930" box="[1002,1316,301,361]" colspan="4" colspanRight="3" gridcol="5" gridrow="0" pageId="4" pageNumber="388">
<taxonomicName id="B4B825F6FF8B34785B8DFE9FFB5B991B" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[1002,1207,301,322]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="4" pageNumber="388" phylum="Nematoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="41CC8267FF8B34785B8DFE9FFB5B991B" box="[1002,1207,301,322]" italics="true" pageId="4" pageNumber="388">Leptolaimus hamatus</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8B34785CD9FE9FFAE6991B" bold="true" box="[1214,1290,301,322]" pageId="4" pageNumber="388">
<taxonomicNameLabel id="5AFF3F1CFF8B34785CD9FE9FFAE6991B" box="[1214,1290,301,322]" pageId="4" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂1 ♂2 ♂3 ♀1
</th>
<th id="8E59354BFF8BCB835D35FE9FFA909930" box="[1362,1404,301,361]" gridcol="9" gridrow="0" pageId="4" pageNumber="388">♀2</th>
</tr>
<tr id="CD885C37FF8BCB8358F8FEC9FA9099C8" box="[159,1404,379,401]" gridrow="1" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FEC9FE0399C8" box="[159,495,379,401]" gridcol="0" gridrow="1" pageId="4" pageNumber="388">Total body length</th>
<td id="8E59354BFF8BCB835AE3FEC9FD4299C8" box="[644,686,379,401]" gridcol="1" gridrow="1" pageId="4" pageNumber="388">980</td>
<td id="8E59354BFF8BCB835A85FEC9FCE099C8" box="[738,780,379,401]" gridcol="2" gridrow="1" pageId="4" pageNumber="388">978</td>
<td id="8E59354BFF8BCB835B25FEC9FC9E99C8" box="[834,882,379,401]" gridcol="3" gridrow="1" pageId="4" pageNumber="388">1001</td>
<td id="8E59354BFF8BCB835BF0FEC9FC2D99C8" box="[919,961,379,401]" gridcol="4" gridrow="1" pageId="4" pageNumber="388">936</td>
<td id="8E59354BFF8BCB835B8DFEC9FBF899C8" box="[1002,1044,379,401]" gridcol="5" gridrow="1" pageId="4" pageNumber="388">690</td>
<td id="8E59354BFF8BCB835C23FEC9FB8299C8" box="[1092,1134,379,401]" gridcol="6" gridrow="1" pageId="4" pageNumber="388">642</td>
<td id="8E59354BFF8BCB835CF8FEC9FB2599C8" box="[1183,1225,379,401]" gridcol="7" gridrow="1" pageId="4" pageNumber="388">624</td>
<td id="8E59354BFF8BCB835C9DFEC9FAC899C8" box="[1274,1316,379,401]" gridcol="8" gridrow="1" pageId="4" pageNumber="388">700</td>
<td id="8E59354BFF8BCB835D35FEC9FA9099C8" box="[1362,1404,379,401]" gridcol="9" gridrow="1" pageId="4" pageNumber="388">682</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FE10FA9099E0" box="[159,1404,418,441]" gridrow="2" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FE10FE0399E0" box="[159,495,418,441]" gridcol="0" gridrow="2" pageId="4" pageNumber="388">Maximum body diameter</th>
<td id="8E59354BFF8BCB835AE3FE10FD4299E0" box="[644,686,418,441]" gridcol="1" gridrow="2" pageId="4" pageNumber="388">14.7</td>
<td id="8E59354BFF8BCB835A85FE10FCE099E0" box="[738,780,418,441]" gridcol="2" gridrow="2" pageId="4" pageNumber="388">12.8</td>
<td id="8E59354BFF8BCB835B25FE10FC9E99E0" box="[834,882,418,441]" gridcol="3" gridrow="2" pageId="4" pageNumber="388">13.3</td>
<td id="8E59354BFF8BCB835BF0FE10FC2D99E0" box="[919,961,418,441]" gridcol="4" gridrow="2" pageId="4" pageNumber="388">14.5</td>
<td id="8E59354BFF8BCB835B8DFE10FBF899E0" box="[1002,1044,418,441]" gridcol="5" gridrow="2" pageId="4" pageNumber="388">20</td>
<td id="8E59354BFF8BCB835C23FE10FB8299E0" box="[1092,1134,418,441]" gridcol="6" gridrow="2" pageId="4" pageNumber="388">21</td>
<td id="8E59354BFF8BCB835CF8FE10FB2599E0" box="[1183,1225,418,441]" gridcol="7" gridrow="2" pageId="4" pageNumber="388">19.5</td>
<td id="8E59354BFF8BCB835C9DFE10FAC899E0" box="[1274,1316,418,441]" gridcol="8" gridrow="2" pageId="4" pageNumber="388">24</td>
<td id="8E59354BFF8BCB835D35FE10FA9099E0" box="[1362,1404,418,441]" gridcol="9" gridrow="2" pageId="4" pageNumber="388">20</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FE78FA9099B9" box="[159,1404,458,480]" gridrow="3" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FE78FE0399B9" box="[159,495,458,480]" gridcol="0" gridrow="3" pageId="4" pageNumber="388">Head diameter</th>
<td id="8E59354BFF8BCB835AE3FE78FD4299B9" box="[644,686,458,480]" gridcol="1" gridrow="3" pageId="4" pageNumber="388">5</td>
<td id="8E59354BFF8BCB835A85FE78FCE099B9" box="[738,780,458,480]" gridcol="2" gridrow="3" pageId="4" pageNumber="388">4.5</td>
<td id="8E59354BFF8BCB835B25FE78FC9E99B9" box="[834,882,458,480]" gridcol="3" gridrow="3" pageId="4" pageNumber="388">4.5</td>
<td id="8E59354BFF8BCB835BF0FE78FC2D99B9" box="[919,961,458,480]" gridcol="4" gridrow="3" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835B8DFE78FBF899B9" box="[1002,1044,458,480]" gridcol="5" gridrow="3" pageId="4" pageNumber="388">5</td>
<td id="8E59354BFF8BCB835C23FE78FB8299B9" box="[1092,1134,458,480]" gridcol="6" gridrow="3" pageId="4" pageNumber="388">4.5</td>
<td id="8E59354BFF8BCB835CF8FE78FB2599B9" box="[1183,1225,458,480]" gridcol="7" gridrow="3" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835C9DFE78FAC899B9" box="[1274,1316,458,480]" gridcol="8" gridrow="3" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835D35FE78FA9099B9" box="[1362,1404,458,480]" gridcol="9" gridrow="3" pageId="4" pageNumber="388">4</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FE43FA909A5E" box="[159,1404,497,519]" gridrow="4" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FE43FE039A5E" box="[159,495,497,519]" gridcol="0" gridrow="4" pageId="4" pageNumber="388">Length of cephalic setae</th>
<td id="8E59354BFF8BCB835AE3FE43FD429A5E" box="[644,686,497,519]" gridcol="1" gridrow="4" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835A85FE43FCE09A5E" box="[738,780,497,519]" gridcol="2" gridrow="4" pageId="4" pageNumber="388">5</td>
<td id="8E59354BFF8BCB835B25FE43FC9E9A5E" box="[834,882,497,519]" gridcol="3" gridrow="4" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835BF0FE43FC2D9A5E" box="[919,961,497,519]" gridcol="4" gridrow="4" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835B8DFE43FBF89A5E" box="[1002,1044,497,519]" gridcol="5" gridrow="4" pageId="4" pageNumber="388">2.5</td>
<td id="8E59354BFF8BCB835C23FE43FB829A5E" box="[1092,1134,497,519]" gridcol="6" gridrow="4" pageId="4" pageNumber="388">2</td>
<td id="8E59354BFF8BCB835CF8FE43FB259A5E" box="[1183,1225,497,519]" gridcol="7" gridrow="4" pageId="4" pageNumber="388">2</td>
<td id="8E59354BFF8BCB835C9DFE43FAC89A5E" box="[1274,1316,497,519]" gridcol="8" gridrow="4" pageId="4" pageNumber="388">3</td>
<td id="8E59354BFF8BCB835D35FE43FA909A5E" box="[1362,1404,497,519]" gridcol="9" gridrow="4" pageId="4" pageNumber="388">2.5</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FDABFA909A76" box="[159,1404,537,559]" gridrow="5" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FDABFE039A76" box="[159,495,537,559]" gridcol="0" gridrow="5" pageId="4" pageNumber="388">Length of buccal cavity</th>
<td id="8E59354BFF8BCB835AE3FDABFD429A76" box="[644,686,537,559]" gridcol="1" gridrow="5" pageId="4" pageNumber="388">21</td>
<td id="8E59354BFF8BCB835A85FDABFCE09A76" box="[738,780,537,559]" gridcol="2" gridrow="5" pageId="4" pageNumber="388">22</td>
<td id="8E59354BFF8BCB835B25FDABFC9E9A76" box="[834,882,537,559]" gridcol="3" gridrow="5" pageId="4" pageNumber="388">20</td>
<td id="8E59354BFF8BCB835BF0FDABFC2D9A76" box="[919,961,537,559]" gridcol="4" gridrow="5" pageId="4" pageNumber="388">19</td>
<td id="8E59354BFF8BCB835B8DFDABFBF89A76" box="[1002,1044,537,559]" gridcol="5" gridrow="5" pageId="4" pageNumber="388">24</td>
<td id="8E59354BFF8BCB835C23FDABFB829A76" box="[1092,1134,537,559]" gridcol="6" gridrow="5" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835CF8FDABFB259A76" box="[1183,1225,537,559]" gridcol="7" gridrow="5" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835C9DFDABFAC89A76" box="[1274,1316,537,559]" gridcol="8" gridrow="5" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835D35FDABFA909A76" box="[1362,1404,537,559]" gridcol="9" gridrow="5" pageId="4" pageNumber="388">16</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FDF2FA909A0F" box="[159,1404,576,598]" gridrow="6" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FDF2FE039A0F" box="[159,495,576,598]" gridcol="0" gridrow="6" pageId="4" pageNumber="388">Diameter of amphidial fovea</th>
<td id="8E59354BFF8BCB835AE3FDF2FD429A0F" box="[644,686,576,598]" gridcol="1" gridrow="6" pageId="4" pageNumber="388">4.5</td>
<td id="8E59354BFF8BCB835A85FDF2FCE09A0F" box="[738,780,576,598]" gridcol="2" gridrow="6" pageId="4" pageNumber="388">3.5</td>
<td id="8E59354BFF8BCB835B25FDF2FC9E9A0F" box="[834,882,576,598]" gridcol="3" gridrow="6" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835BF0FDF2FC2D9A0F" box="[919,961,576,598]" gridcol="4" gridrow="6" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835B8DFDF2FBF89A0F" box="[1002,1044,576,598]" gridcol="5" gridrow="6" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835C23FDF2FB829A0F" box="[1092,1134,576,598]" gridcol="6" gridrow="6" pageId="4" pageNumber="388">4</td>
<td id="8E59354BFF8BCB835CF8FDF2FB259A0F" box="[1183,1225,576,598]" gridcol="7" gridrow="6" pageId="4" pageNumber="388">3.5</td>
<td id="8E59354BFF8BCB835C9DFDF2FAC89A0F" box="[1274,1316,576,598]" gridcol="8" gridrow="6" pageId="4" pageNumber="388">3</td>
<td id="8E59354BFF8BCB835D35FDF2FA909A0F" box="[1362,1404,576,598]" gridcol="9" gridrow="6" pageId="4" pageNumber="388">3</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FDD5FA909A24" box="[159,1404,615,637]" gridrow="7" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FDD5FE039A24" box="[159,495,615,637]" gridcol="0" gridrow="7" pageId="4" pageNumber="388">Body diameter at amphidial fovea</th>
<td id="8E59354BFF8BCB835AE3FDD5FD429A24" box="[644,686,615,637]" gridcol="1" gridrow="7" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835A85FDD5FCE09A24" box="[738,780,615,637]" gridcol="2" gridrow="7" pageId="4" pageNumber="388">7.5</td>
<td id="8E59354BFF8BCB835B25FDD5FC9E9A24" box="[834,882,615,637]" gridcol="3" gridrow="7" pageId="4" pageNumber="388">7.5</td>
<td id="8E59354BFF8BCB835BF0FDD5FC2D9A24" box="[919,961,615,637]" gridcol="4" gridrow="7" pageId="4" pageNumber="388">7.5</td>
<td id="8E59354BFF8BCB835B8DFDD5FBF89A24" box="[1002,1044,615,637]" gridcol="5" gridrow="7" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835C23FDD5FB829A24" box="[1092,1134,615,637]" gridcol="6" gridrow="7" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835CF8FDD5FB259A24" box="[1183,1225,615,637]" gridcol="7" gridrow="7" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835C9DFDD5FAC89A24" box="[1274,1316,615,637]" gridcol="8" gridrow="7" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835D35FDD5FA909A24" box="[1362,1404,615,637]" gridcol="9" gridrow="7" pageId="4" pageNumber="388">8</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FD3CFA909AFC" box="[159,1404,654,677]" gridrow="8" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FD3CFE039AFC" box="[159,495,654,677]" gridcol="0" gridrow="8" pageId="4" pageNumber="388">Amphidial fovea from anterior end</th>
<td id="8E59354BFF8BCB835AE3FD3CFD429AFC" box="[644,686,654,677]" gridcol="1" gridrow="8" pageId="4" pageNumber="388">14</td>
<td id="8E59354BFF8BCB835A85FD3CFCE09AFC" box="[738,780,654,677]" gridcol="2" gridrow="8" pageId="4" pageNumber="388">14</td>
<td id="8E59354BFF8BCB835B25FD3CFC9E9AFC" box="[834,882,654,677]" gridcol="3" gridrow="8" pageId="4" pageNumber="388">15</td>
<td id="8E59354BFF8BCB835BF0FD3CFC2D9AFC" box="[919,961,654,677]" gridcol="4" gridrow="8" pageId="4" pageNumber="388">11</td>
<td id="8E59354BFF8BCB835B8DFD3CFBF89AFC" box="[1002,1044,654,677]" gridcol="5" gridrow="8" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835C23FD3CFB829AFC" box="[1092,1134,654,677]" gridcol="6" gridrow="8" pageId="4" pageNumber="388">12</td>
<td id="8E59354BFF8BCB835CF8FD3CFB259AFC" box="[1183,1225,654,677]" gridcol="7" gridrow="8" pageId="4" pageNumber="388">12.5</td>
<td id="8E59354BFF8BCB835C9DFD3CFAC89AFC" box="[1274,1316,654,677]" gridcol="8" gridrow="8" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835D35FD3CFA909AFC" box="[1362,1404,654,677]" gridcol="9" gridrow="8" pageId="4" pageNumber="388">12</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FD04FA909A95" box="[159,1404,694,716]" gridrow="9" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FD04FE039A95" box="[159,495,694,716]" gridcol="0" gridrow="9" pageId="4" pageNumber="388">Length of pharynx</th>
<td id="8E59354BFF8BCB835AE3FD04FD429A95" box="[644,686,694,716]" gridcol="1" gridrow="9" pageId="4" pageNumber="388">137</td>
<td id="8E59354BFF8BCB835A85FD04FCE09A95" box="[738,780,694,716]" gridcol="2" gridrow="9" pageId="4" pageNumber="388">143</td>
<td id="8E59354BFF8BCB835B25FD04FC9E9A95" box="[834,882,694,716]" gridcol="3" gridrow="9" pageId="4" pageNumber="388">150</td>
<td id="8E59354BFF8BCB835BF0FD04FC2D9A95" box="[919,961,694,716]" gridcol="4" gridrow="9" pageId="4" pageNumber="388">128</td>
<td id="8E59354BFF8BCB835B8DFD04FBF89A95" box="[1002,1044,694,716]" gridcol="5" gridrow="9" pageId="4" pageNumber="388">136</td>
<td id="8E59354BFF8BCB835C23FD04FB829A95" box="[1092,1134,694,716]" gridcol="6" gridrow="9" pageId="4" pageNumber="388">145</td>
<td id="8E59354BFF8BCB835CF8FD04FB259A95" box="[1183,1225,694,716]" gridcol="7" gridrow="9" pageId="4" pageNumber="388">128</td>
<td id="8E59354BFF8BCB835C9DFD04FAC89A95" box="[1274,1316,694,716]" gridcol="8" gridrow="9" pageId="4" pageNumber="388">139</td>
<td id="8E59354BFF8BCB835D35FD04FA909A95" box="[1362,1404,694,716]" gridcol="9" gridrow="9" pageId="4" pageNumber="388">134</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FD6FFA909AAA" box="[159,1404,733,755]" gridrow="10" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FD6FFE039AAA" box="[159,495,733,755]" gridcol="0" gridrow="10" pageId="4" pageNumber="388">Body diameter at pharyngeal base</th>
<td id="8E59354BFF8BCB835AE3FD6FFD429AAA" box="[644,686,733,755]" gridcol="1" gridrow="10" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835A85FD6FFCE09AAA" box="[738,780,733,755]" gridcol="2" gridrow="10" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835B25FD6FFC9E9AAA" box="[834,882,733,755]" gridcol="3" gridrow="10" pageId="4" pageNumber="388">12</td>
<td id="8E59354BFF8BCB835BF0FD6FFC2D9AAA" box="[919,961,733,755]" gridcol="4" gridrow="10" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835B8DFD6FFBF89AAA" box="[1002,1044,733,755]" gridcol="5" gridrow="10" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835C23FD6FFB829AAA" box="[1092,1134,733,755]" gridcol="6" gridrow="10" pageId="4" pageNumber="388">19</td>
<td id="8E59354BFF8BCB835CF8FD6FFB259AAA" box="[1183,1225,733,755]" gridcol="7" gridrow="10" pageId="4" pageNumber="388">18.5</td>
<td id="8E59354BFF8BCB835C9DFD6FFAC89AAA" box="[1274,1316,733,755]" gridcol="8" gridrow="10" pageId="4" pageNumber="388">21</td>
<td id="8E59354BFF8BCB835D35FD6FFA909AAA" box="[1362,1404,733,755]" gridcol="9" gridrow="10" pageId="4" pageNumber="388">19</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FCB6FA909B43" box="[159,1404,772,794]" gridrow="11" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FCB6FE039B43" box="[159,495,772,794]" gridcol="0" gridrow="11" pageId="4" pageNumber="388">Length of cardia</th>
<td id="8E59354BFF8BCB835AE3FCB6FD429B43" box="[644,686,772,794]" gridcol="1" gridrow="11" pageId="4" pageNumber="388">6.5</td>
<td id="8E59354BFF8BCB835A85FCB6FCE09B43" box="[738,780,772,794]" gridcol="2" gridrow="11" pageId="4" pageNumber="388">7</td>
<td id="8E59354BFF8BCB835B25FCB6FC9E9B43" box="[834,882,772,794]" gridcol="3" gridrow="11" pageId="4" pageNumber="388">10</td>
<td id="8E59354BFF8BCB835BF0FCB6FC2D9B43" box="[919,961,772,794]" gridcol="4" gridrow="11" pageId="4" pageNumber="388">7</td>
<td id="8E59354BFF8BCB835B8DFCB6FBF89B43" box="[1002,1044,772,794]" gridcol="5" gridrow="11" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835C23FCB6FB829B43" box="[1092,1134,772,794]" gridcol="6" gridrow="11" pageId="4" pageNumber="388">9</td>
<td id="8E59354BFF8BCB835CF8FCB6FB259B43" box="[1183,1225,772,794]" gridcol="7" gridrow="11" pageId="4" pageNumber="388">7</td>
<td id="8E59354BFF8BCB835C9DFCB6FAC89B43" box="[1274,1316,772,794]" gridcol="8" gridrow="11" pageId="4" pageNumber="388">9</td>
<td id="8E59354BFF8BCB835D35FCB6FA909B43" box="[1362,1404,772,794]" gridcol="9" gridrow="11" pageId="4" pageNumber="388">8.5</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FC9EFA909B1B" box="[159,1404,812,834]" gridrow="12" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FC9EFE039B1B" box="[159,495,812,834]" gridcol="0" gridrow="12" pageId="4" pageNumber="388">Spicules length along arc</th>
<td id="8E59354BFF8BCB835AE3FC9EFD429B1B" box="[644,686,812,834]" gridcol="1" gridrow="12" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835A85FC9EFCE09B1B" box="[738,780,812,834]" gridcol="2" gridrow="12" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835B25FC9EFC9E9B1B" box="[834,882,812,834]" gridcol="3" gridrow="12" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835BF0FC9EFC2D9B1B" box="[919,961,812,834]" gridcol="4" gridrow="12" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B8DFC9EFBF89B1B" box="[1002,1044,812,834]" gridcol="5" gridrow="12" pageId="4" pageNumber="388">22</td>
<td id="8E59354BFF8BCB835C23FC9EFB829B1B" box="[1092,1134,812,834]" gridcol="6" gridrow="12" pageId="4" pageNumber="388">22</td>
<td id="8E59354BFF8BCB835CF8FC9EFB259B1B" box="[1183,1225,812,834]" gridcol="7" gridrow="12" pageId="4" pageNumber="388">24</td>
<td id="8E59354BFF8BCB835C9DFC9EFAC89B1B" box="[1274,1316,812,834]" gridcol="8" gridrow="12" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835D35FC9EFA909B1B" box="[1362,1404,812,834]" gridcol="9" gridrow="12" pageId="4" pageNumber="388">-</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FCE1FA909B30" box="[159,1404,851,873]" gridrow="13" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FCE1FE039B30" box="[159,495,851,873]" gridcol="0" gridrow="13" pageId="4" pageNumber="388">Length of gubernaculum</th>
<td id="8E59354BFF8BCB835AE3FCE1FD429B30" box="[644,686,851,873]" gridcol="1" gridrow="13" pageId="4" pageNumber="388">12</td>
<td id="8E59354BFF8BCB835A85FCE1FCE09B30" box="[738,780,851,873]" gridcol="2" gridrow="13" pageId="4" pageNumber="388">13</td>
<td id="8E59354BFF8BCB835B25FCE1FC9E9B30" box="[834,882,851,873]" gridcol="3" gridrow="13" pageId="4" pageNumber="388">11</td>
<td id="8E59354BFF8BCB835BF0FCE1FC2D9B30" box="[919,961,851,873]" gridcol="4" gridrow="13" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B8DFCE1FBF89B30" box="[1002,1044,851,873]" gridcol="5" gridrow="13" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835C23FCE1FB829B30" box="[1092,1134,851,873]" gridcol="6" gridrow="13" pageId="4" pageNumber="388">9</td>
<td id="8E59354BFF8BCB835CF8FCE1FB259B30" box="[1183,1225,851,873]" gridcol="7" gridrow="13" pageId="4" pageNumber="388">11</td>
<td id="8E59354BFF8BCB835C9DFCE1FAC89B30" box="[1274,1316,851,873]" gridcol="8" gridrow="13" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835D35FCE1FA909B30" box="[1362,1404,851,873]" gridcol="9" gridrow="13" pageId="4" pageNumber="388">-</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FCC8FA909BC9" box="[159,1404,890,912]" gridrow="14" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FCC8FE039BC9" box="[159,495,890,912]" gridcol="0" gridrow="14" pageId="4" pageNumber="388">Number of precloacal supplements</th>
<td id="8E59354BFF8BCB835AE3FCC8FD429BC9" box="[644,686,890,912]" gridcol="1" gridrow="14" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835A85FCC8FCE09BC9" box="[738,780,890,912]" gridcol="2" gridrow="14" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835B25FCC8FC9E9BC9" box="[834,882,890,912]" gridcol="3" gridrow="14" pageId="4" pageNumber="388">8</td>
<td id="8E59354BFF8BCB835BF0FCC8FC2D9BC9" box="[919,961,890,912]" gridcol="4" gridrow="14" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B8DFCC8FBF89BC9" box="[1002,1044,890,912]" gridcol="5" gridrow="14" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835C23FCC8FB829BC9" box="[1092,1134,890,912]" gridcol="6" gridrow="14" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835CF8FCC8FB259BC9" box="[1183,1225,890,912]" gridcol="7" gridrow="14" pageId="4" pageNumber="388">6</td>
<td id="8E59354BFF8BCB835C9DFCC8FAC89BC9" box="[1274,1316,890,912]" gridcol="8" gridrow="14" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835D35FCC8FA909BC9" box="[1362,1404,890,912]" gridcol="9" gridrow="14" pageId="4" pageNumber="388">-</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FC10FA909BE1" box="[159,1404,930,952]" gridrow="15" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FC10FE039BE1" box="[159,495,930,952]" gridcol="0" gridrow="15" pageId="4" pageNumber="388">Vulva from anterior end</th>
<td id="8E59354BFF8BCB835AE3FC10FD429BE1" box="[644,686,930,952]" gridcol="1" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835A85FC10FCE09BE1" box="[738,780,930,952]" gridcol="2" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B25FC10FC9E9BE1" box="[834,882,930,952]" gridcol="3" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835BF0FC10FC2D9BE1" box="[919,961,930,952]" gridcol="4" gridrow="15" pageId="4" pageNumber="388">456</td>
<td id="8E59354BFF8BCB835B8DFC10FBF89BE1" box="[1002,1044,930,952]" gridcol="5" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C23FC10FB829BE1" box="[1092,1134,930,952]" gridcol="6" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835CF8FC10FB259BE1" box="[1183,1225,930,952]" gridcol="7" gridrow="15" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C9DFC10FAC89BE1" box="[1274,1316,930,952]" gridcol="8" gridrow="15" pageId="4" pageNumber="388">334</td>
<td id="8E59354BFF8BCB835D35FC10FA909BE1" box="[1362,1404,930,952]" gridcol="9" gridrow="15" pageId="4" pageNumber="388">341</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FC7BFA909B86" box="[159,1404,969,991]" gridrow="16" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FC7BFE039B86" box="[159,495,969,991]" gridcol="0" gridrow="16" pageId="4" pageNumber="388">Body diameter at vulva</th>
<td id="8E59354BFF8BCB835AE3FC7BFD429B86" box="[644,686,969,991]" gridcol="1" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835A85FC7BFCE09B86" box="[738,780,969,991]" gridcol="2" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B25FC7BFC9E9B86" box="[834,882,969,991]" gridcol="3" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835BF0FC7BFC2D9B86" box="[919,961,969,991]" gridcol="4" gridrow="16" pageId="4" pageNumber="388">14</td>
<td id="8E59354BFF8BCB835B8DFC7BFBF89B86" box="[1002,1044,969,991]" gridcol="5" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C23FC7BFB829B86" box="[1092,1134,969,991]" gridcol="6" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835CF8FC7BFB259B86" box="[1183,1225,969,991]" gridcol="7" gridrow="16" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C9DFC7BFAC89B86" box="[1274,1316,969,991]" gridcol="8" gridrow="16" pageId="4" pageNumber="388">24</td>
<td id="8E59354BFF8BCB835D35FC7BFA909B86" box="[1362,1404,969,991]" gridcol="9" gridrow="16" pageId="4" pageNumber="388">20</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FC42FA909C5F" box="[159,1404,1008,1030]" gridrow="17" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FC42FE039C5F" box="[159,495,1008,1030]" gridcol="0" gridrow="17" pageId="4" pageNumber="388">V%</th>
<td id="8E59354BFF8BCB835AE3FC42FD429C5F" box="[644,686,1008,1030]" gridcol="1" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835A85FC42FCE09C5F" box="[738,780,1008,1030]" gridcol="2" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835B25FC42FC9E9C5F" box="[834,882,1008,1030]" gridcol="3" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835BF0FC42FC2D9C5F" box="[919,961,1008,1030]" gridcol="4" gridrow="17" pageId="4" pageNumber="388">49</td>
<td id="8E59354BFF8BCB835B8DFC42FBF89C5F" box="[1002,1044,1008,1030]" gridcol="5" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C23FC42FB829C5F" box="[1092,1134,1008,1030]" gridcol="6" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835CF8FC42FB259C5F" box="[1183,1225,1008,1030]" gridcol="7" gridrow="17" pageId="4" pageNumber="388">-</td>
<td id="8E59354BFF8BCB835C9DFC42FAC89C5F" box="[1274,1316,1008,1030]" gridcol="8" gridrow="17" pageId="4" pageNumber="388">48</td>
<td id="8E59354BFF8BCB835D35FC42FA909C5F" box="[1362,1404,1008,1030]" gridcol="9" gridrow="17" pageId="4" pageNumber="388">50</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FBAAFA909C77" box="[159,1404,1048,1070]" gridrow="18" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FBAAFE039C77" box="[159,495,1048,1070]" gridcol="0" gridrow="18" pageId="4" pageNumber="388">Body diameter at cloaca or anus</th>
<td id="8E59354BFF8BCB835AE3FBAAFD429C77" box="[644,686,1048,1070]" gridcol="1" gridrow="18" pageId="4" pageNumber="388">14.4</td>
<td id="8E59354BFF8BCB835A85FBAAFCE09C77" box="[738,780,1048,1070]" gridcol="2" gridrow="18" pageId="4" pageNumber="388">12.5</td>
<td id="8E59354BFF8BCB835B25FBAAFC9E9C77" box="[834,882,1048,1070]" gridcol="3" gridrow="18" pageId="4" pageNumber="388">12.5</td>
<td id="8E59354BFF8BCB835BF0FBAAFC2D9C77" box="[919,961,1048,1070]" gridcol="4" gridrow="18" pageId="4" pageNumber="388">9.5</td>
<td id="8E59354BFF8BCB835B8DFBAAFBF89C77" box="[1002,1044,1048,1070]" gridcol="5" gridrow="18" pageId="4" pageNumber="388">17</td>
<td id="8E59354BFF8BCB835C23FBAAFB829C77" box="[1092,1134,1048,1070]" gridcol="6" gridrow="18" pageId="4" pageNumber="388">18</td>
<td id="8E59354BFF8BCB835CF8FBAAFB259C77" box="[1183,1225,1048,1070]" gridcol="7" gridrow="18" pageId="4" pageNumber="388">17</td>
<td id="8E59354BFF8BCB835C9DFBAAFAC89C77" box="[1274,1316,1048,1070]" gridcol="8" gridrow="18" pageId="4" pageNumber="388">15</td>
<td id="8E59354BFF8BCB835D35FBAAFA909C77" box="[1362,1404,1048,1070]" gridcol="9" gridrow="18" pageId="4" pageNumber="388">14</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FB8DFA909C0C" box="[159,1404,1087,1109]" gridrow="19" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FB8DFE039C0C" box="[159,495,1087,1109]" gridcol="0" gridrow="19" pageId="4" pageNumber="388">Tail length</th>
<td id="8E59354BFF8BCB835AE3FB8DFD429C0C" box="[644,686,1087,1109]" gridcol="1" gridrow="19" pageId="4" pageNumber="388">105</td>
<td id="8E59354BFF8BCB835A85FB8DFCE09C0C" box="[738,780,1087,1109]" gridcol="2" gridrow="19" pageId="4" pageNumber="388">111</td>
<td id="8E59354BFF8BCB835B25FB8DFC9E9C0C" box="[834,882,1087,1109]" gridcol="3" gridrow="19" pageId="4" pageNumber="388">84</td>
<td id="8E59354BFF8BCB835BF0FB8DFC2D9C0C" box="[919,961,1087,1109]" gridcol="4" gridrow="19" pageId="4" pageNumber="388">116</td>
<td id="8E59354BFF8BCB835B8DFB8DFBF89C0C" box="[1002,1044,1087,1109]" gridcol="5" gridrow="19" pageId="4" pageNumber="388">110</td>
<td id="8E59354BFF8BCB835C23FB8DFB829C0C" box="[1092,1134,1087,1109]" gridcol="6" gridrow="19" pageId="4" pageNumber="388">102</td>
<td id="8E59354BFF8BCB835CF8FB8DFB259C0C" box="[1183,1225,1087,1109]" gridcol="7" gridrow="19" pageId="4" pageNumber="388">101</td>
<td id="8E59354BFF8BCB835C9DFB8DFAC89C0C" box="[1274,1316,1087,1109]" gridcol="8" gridrow="19" pageId="4" pageNumber="388">118</td>
<td id="8E59354BFF8BCB835D35FB8DFA909C0C" box="[1362,1404,1087,1109]" gridcol="9" gridrow="19" pageId="4" pageNumber="388">106</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FBD4FA909C25" box="[159,1404,1126,1148]" gridrow="20" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FBD4FE039C25" box="[159,495,1126,1148]" gridcol="0" gridrow="20" pageId="4" pageNumber="388">a</th>
<td id="8E59354BFF8BCB835AE3FBD4FD429C25" box="[644,686,1126,1148]" gridcol="1" gridrow="20" pageId="4" pageNumber="388">66.7</td>
<td id="8E59354BFF8BCB835A85FBD4FCE09C25" box="[738,780,1126,1148]" gridcol="2" gridrow="20" pageId="4" pageNumber="388">76.4</td>
<td id="8E59354BFF8BCB835B25FBD4FC9E9C25" box="[834,882,1126,1148]" gridcol="3" gridrow="20" pageId="4" pageNumber="388">75.2</td>
<td id="8E59354BFF8BCB835BF0FBD4FC2D9C25" box="[919,961,1126,1148]" gridcol="4" gridrow="20" pageId="4" pageNumber="388">64.6</td>
<td id="8E59354BFF8BCB835B8DFBD4FBF89C25" box="[1002,1044,1126,1148]" gridcol="5" gridrow="20" pageId="4" pageNumber="388">34.5</td>
<td id="8E59354BFF8BCB835C23FBD4FB829C25" box="[1092,1134,1126,1148]" gridcol="6" gridrow="20" pageId="4" pageNumber="388">30.6</td>
<td id="8E59354BFF8BCB835CF8FBD4FB259C25" box="[1183,1225,1126,1148]" gridcol="7" gridrow="20" pageId="4" pageNumber="388">33.1</td>
<td id="8E59354BFF8BCB835C9DFBD4FAC89C25" box="[1274,1316,1126,1148]" gridcol="8" gridrow="20" pageId="4" pageNumber="388">29.2</td>
<td id="8E59354BFF8BCB835D35FBD4FA909C25" box="[1362,1404,1126,1148]" gridcol="9" gridrow="20" pageId="4" pageNumber="388">34.1</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FB3CFA909CFD" box="[159,1404,1166,1188]" gridrow="21" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FB3CFE039CFD" box="[159,495,1166,1188]" gridcol="0" gridrow="21" pageId="4" pageNumber="388">b</th>
<td id="8E59354BFF8BCB835AE3FB3CFD429CFD" box="[644,686,1166,1188]" gridcol="1" gridrow="21" pageId="4" pageNumber="388">7.1</td>
<td id="8E59354BFF8BCB835A85FB3CFCE09CFD" box="[738,780,1166,1188]" gridcol="2" gridrow="21" pageId="4" pageNumber="388">6.9</td>
<td id="8E59354BFF8BCB835B25FB3CFC9E9CFD" box="[834,882,1166,1188]" gridcol="3" gridrow="21" pageId="4" pageNumber="388">6.7</td>
<td id="8E59354BFF8BCB835BF0FB3CFC2D9CFD" box="[919,961,1166,1188]" gridcol="4" gridrow="21" pageId="4" pageNumber="388">7.3</td>
<td id="8E59354BFF8BCB835B8DFB3CFBF89CFD" box="[1002,1044,1166,1188]" gridcol="5" gridrow="21" pageId="4" pageNumber="388">5.1</td>
<td id="8E59354BFF8BCB835C23FB3CFB829CFD" box="[1092,1134,1166,1188]" gridcol="6" gridrow="21" pageId="4" pageNumber="388">4.4</td>
<td id="8E59354BFF8BCB835CF8FB3CFB259CFD" box="[1183,1225,1166,1188]" gridcol="7" gridrow="21" pageId="4" pageNumber="388">5.0</td>
<td id="8E59354BFF8BCB835C9DFB3CFAC89CFD" box="[1274,1316,1166,1188]" gridcol="8" gridrow="21" pageId="4" pageNumber="388">5.0</td>
<td id="8E59354BFF8BCB835D35FB3CFA909CFD" box="[1362,1404,1166,1188]" gridcol="9" gridrow="21" pageId="4" pageNumber="388">5.1</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FB07FA909C92" box="[159,1404,1205,1227]" gridrow="22" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FB07FE039C92" box="[159,495,1205,1227]" gridcol="0" gridrow="22" pageId="4" pageNumber="388">c</th>
<td id="8E59354BFF8BCB835AE3FB07FD429C92" box="[644,686,1205,1227]" gridcol="1" gridrow="22" pageId="4" pageNumber="388">9.3</td>
<td id="8E59354BFF8BCB835A85FB07FCE09C92" box="[738,780,1205,1227]" gridcol="2" gridrow="22" pageId="4" pageNumber="388">8.8</td>
<td id="8E59354BFF8BCB835B25FB07FC9E9C92" box="[834,882,1205,1227]" gridcol="3" gridrow="22" pageId="4" pageNumber="388">11.9</td>
<td id="8E59354BFF8BCB835BF0FB07FC2D9C92" box="[919,961,1205,1227]" gridcol="4" gridrow="22" pageId="4" pageNumber="388">8.0</td>
<td id="8E59354BFF8BCB835B8DFB07FBF89C92" box="[1002,1044,1205,1227]" gridcol="5" gridrow="22" pageId="4" pageNumber="388">6.3</td>
<td id="8E59354BFF8BCB835C23FB07FB829C92" box="[1092,1134,1205,1227]" gridcol="6" gridrow="22" pageId="4" pageNumber="388">6.3</td>
<td id="8E59354BFF8BCB835CF8FB07FB259C92" box="[1183,1225,1205,1227]" gridcol="7" gridrow="22" pageId="4" pageNumber="388">6.4</td>
<td id="8E59354BFF8BCB835C9DFB07FAC89C92" box="[1274,1316,1205,1227]" gridcol="8" gridrow="22" pageId="4" pageNumber="388">5.9</td>
<td id="8E59354BFF8BCB835D35FB07FA909C92" box="[1362,1404,1205,1227]" gridcol="9" gridrow="22" pageId="4" pageNumber="388">6.4</td>
</tr>
<tr id="CD885C37FF8BCB8358F8FB69FA909CAB" box="[159,1404,1243,1266]" gridrow="23" pageId="4" pageNumber="388">
<th id="8E59354BFF8BCB8358F8FB69FE039CAB" box="[159,495,1243,1266]" gridcol="0" gridrow="23" pageId="4" pageNumber="388">c′</th>
<td id="8E59354BFF8BCB835AE3FB69FD429CAB" box="[644,686,1243,1266]" gridcol="1" gridrow="23" pageId="4" pageNumber="388">7.3</td>
<td id="8E59354BFF8BCB835A85FB69FCE09CAB" box="[738,780,1243,1266]" gridcol="2" gridrow="23" pageId="4" pageNumber="388">8.9</td>
<td id="8E59354BFF8BCB835B25FB69FC9E9CAB" box="[834,882,1243,1266]" gridcol="3" gridrow="23" pageId="4" pageNumber="388">6.2</td>
<td id="8E59354BFF8BCB835BF0FB69FC2D9CAB" box="[919,961,1243,1266]" gridcol="4" gridrow="23" pageId="4" pageNumber="388">12.2</td>
<td id="8E59354BFF8BCB835B8DFB69FBF89CAB" box="[1002,1044,1243,1266]" gridcol="5" gridrow="23" pageId="4" pageNumber="388">6.5</td>
<td id="8E59354BFF8BCB835C23FB69FB829CAB" box="[1092,1134,1243,1266]" gridcol="6" gridrow="23" pageId="4" pageNumber="388">5.7</td>
<td id="8E59354BFF8BCB835CF8FB69FB259CAB" box="[1183,1225,1243,1266]" gridcol="7" gridrow="23" pageId="4" pageNumber="388">5.9</td>
<td id="8E59354BFF8BCB835C9DFB69FAC89CAB" box="[1274,1316,1243,1266]" gridcol="8" gridrow="23" pageId="4" pageNumber="388">7.7</td>
<td id="8E59354BFF8BCB835D35FB69FA909CAB" box="[1362,1404,1243,1266]" gridcol="9" gridrow="23" pageId="4" pageNumber="388">7.6</td>
</tr>
</table>
</paragraph>
<paragraph id="73075E75FF8B347858F0FAF4FEDF9D39" blockId="4.[151,307,1350,1376]" box="[151,307,1350,1376]" pageId="4" pageNumber="388">
<emphasis id="41CC8267FF8B347858F0FAF4FEDF9D39" bold="true" box="[151,307,1350,1376]" pageId="4" pageNumber="388">Descriptions</emphasis>
</paragraph>
<paragraph id="73075E75FF8B347858F0FA3CFF339DF1" blockId="4.[151,1437,1422,2025]" box="[151,223,1422,1448]" pageId="4" pageNumber="388">
<emphasis id="41CC8267FF8B347858F0FA3CFF339DF1" bold="true" box="[151,223,1422,1448]" pageId="4" pageNumber="388">Males</emphasis>
</paragraph>
<paragraph id="73075E75FF8B347858F0FA00FE6F9F49" blockId="4.[151,1437,1422,2025]" pageId="4" pageNumber="388">Body very slender. Cuticle annulated, annuli about 2.0 µm wide at mid-body, without ornamentation. Lateral field and body pores indistinct. Setae along pharyngeal region not seen. Labial region rounded, offset from body contour by a constriction. Inner and outer labial sensilla indistinct. Cephalic sensilla setiform, equal to 1.2–1.5 times of labial region diameter. Subcephalic and cervical sensilla and ocelli absent. Amphidial fovea unispiral, round in outline, 47–56% corresponding body diameter, located at level of post-mid of stoma. Nerve ring surrounding pharynx at level of middle of pharynx, i.e. 78–94 µm from anterior end of the body. Secretory-excretory system not seen. Buccal cavity uniformly tubular, 4.2–4.9 head diameter deep. Pharynx cylindrical, with distinct oval basal bulb. Cardia cylindrical, its posterior part embedded in intestine. Tail elongate conical, gradually narrowing distally. Three to four pairs of subventral and two pairs of subdorsal caudal setae 4–5 µm long. Three caudal glands present. Spinneret functional.</paragraph>
<paragraph id="73075E75FF8B347958A0F8A8FD42988C" blockId="4.[151,1437,1422,2025]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388">
Reproductive system diorchic, anterior testis outstretched, posterior one reflexed. Spicules paired, symmetrical, arcuated with expanded proximal end and tapered distal end. Gubernaculum with a straight dorso-caudal apophysis. Accessory apparatus composed of eight almost evenly spaced midventral tubular supplements extending for 211– 251 µm from cloaca towards anterior end, without alveolar supplements. Tubular supplements sickle-shaped, with rounded proximal ends and pointed distal ends, 8–8.5 µm long. Posteriormost tubular supplement 21–36 µm anterior to cloaca, and anterior to spicules. The spacing of adjacent supplements in the
<specimenCount id="65BE95FCFF8B34785B98F87CFB9F9FB1" box="[1023,1139,1998,2024]" count="3" pageId="4" pageNumber="388" type="male">three male</specimenCount>
specimens is almost equal (27–30 µm, 22–24 µm and 22–29 µm, respectively) except that of the most anterior two supplements being slightly larger (37 µm, 34 µm and 31 µm, respectively).
</paragraph>
<paragraph id="73075E75FF8A347958F0FEB1FF029944" blockId="5.[151,1437,259,1401]" box="[151,238,259,285]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958F0FEB1FF029944" bold="true" box="[151,238,259,285]" pageId="5" pageNumber="389">Female</emphasis>
</paragraph>
<paragraph id="73075E75FF8A347958F0FE95FD3F99AC" blockId="5.[151,1437,259,1401]" pageId="5" pageNumber="389">
Similar to males in most morphological characters, except tail longer and slender (c′=12.2 vs
<quantity id="B440F390FF8A34795CB3FE95FAA49918" box="[1236,1352,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.9177" metricValueMax="2.2606" metricValueMin="1.5748" pageId="5" pageNumber="389" unit="in" value="7.55" valueMax="8.9" valueMin="6.2">6.2–8.9 in</quantity>
males) without setae, amphidial fovea located in more anterior position, and supplements absent. Reproductive system didelphic, with two opposed, reflexed ovaries, the anterior ovary located to the right side of the intestine, and the posterior one located to the left side of the intestine. Oviduct a narrow tube. Spermathecae indistinct. Uterus a wide and short tube. Vagina straight, 0.2 times vulval body diameter long. Vulva midventral, located in the mid-body, 49% of the total body length from the anterior end.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347958A0FE4CFC5C9AF0" pageId="5" pageNumber="389" type="diagnosis">
<paragraph id="73075E75FF8A347958A0FE4CFC5C9AF0" blockId="5.[151,1437,259,1401]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958A0FE4CFEAE9A41" bold="true" box="[199,322,510,536]" pageId="5" pageNumber="389">Diagnosis.</emphasis>
<taxonomicName id="B4B825F6FF8A3479592EFE4DFDDE9A40" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[329,562,511,537]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8A3479592EFE4DFDDE9A40" box="[329,562,511,537]" italics="true" pageId="5" pageNumber="389">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8A34795A5DFE4DFD799A40" bold="true" box="[570,661,511,537]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="5AFF3F1CFF8A34795A5DFE4DFD799A40" box="[570,661,511,537]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by slender body 936–1001 µm long; rounded labial region offset from body contour; cephalic setae 5–6 µm long; amphidial fovea located 2.8–3.3 times of head diameter from anterior end; males with eight tubular and without alveolar supplements, tubular supplements sickle-shaped with pointed tips; spicules arcuated and about 18 µm long; gubernaculum with a straight dorso-caudal apophysis; female without supplements, vulva situated at midventral side of body.
</paragraph>
</subSubSection>
<subSubSection id="3BA20DFEFF8A347958A0FD01FECB9D20" pageId="5" pageNumber="389" type="discussion">
<paragraph id="73075E75FF8A347958A0FD01FECB9D20" blockId="5.[151,1437,259,1401]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A347958A0FD01FED79A94" bold="true" box="[199,315,691,717]" pageId="5" pageNumber="389">Remarks.</emphasis>
<taxonomicName id="B4B825F6FF8A34795925FD01FDC59A94" authorityName="Gu &amp; Sun &amp; Huang" authorityYear="2024" box="[322,553,691,717]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="41CC8267FF8A34795925FD01FDC59A94" box="[322,553,691,717]" italics="true" pageId="5" pageNumber="389">Leptolaimus gracilis</emphasis>
</taxonomicName>
<emphasis id="41CC8267FF8A34795A48FD01FD649A94" bold="true" box="[559,648,691,717]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="5AFF3F1CFF8A34795A48FD01FD649A94" box="[559,648,691,717]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the group of
<taxonomicName id="B4B825F6FF8A34795BF0FD01FB959A94" box="[919,1145,691,717]" pageId="5" pageNumber="389">
<emphasis id="41CC8267FF8A34795BF0FD01FBCD9A94" box="[919,1057,691,717]" italics="true" pageId="5" pageNumber="389">Leptolaimus</emphasis>
species
</taxonomicName>
in which males have only tubular precloacal supplements and females have no supplements. The new species is similar to
<taxonomicName id="B4B825F6FF8A34795CA2FD65FEA59B4C" authority="Lorenzen, 1972" authorityName="Lorenzen" authorityYear="1972" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="macer">
<emphasis id="41CC8267FF8A34795CA2FD65FA709AA9" box="[1221,1436,727,753]" italics="true" pageId="5" pageNumber="389">Leptolaimus macer</emphasis>
Lorenzen,
<quantity id="B440F390FF8A34795976FD49FEA59B4C" box="[273,329,763,789]" metricMagnitude="1" metricUnit="m" metricValue="5.00888" pageId="5" pageNumber="389" unit="in" value="1972.0">1972</quantity>
</taxonomicName>
in having sickle-shaped tubular precloacal supplements, but differs by longer body length (936– 1001 µm
<emphasis id="41CC8267FF8A34795965FCADFEF59B61" box="[258,281,799,824]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
698–790 µm), larger amphidial fovea (3.5–4.5 µm
<emphasis id="41CC8267FF8A34795B3EFCADFC9C9B61" box="[857,880,799,824]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
2 µm in diameter in males), straight dorso-caudal gubernacular apophysis (
<emphasis id="41CC8267FF8A347959D6FCF1FE249B05" box="[433,456,835,860]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
arcuated dorsal apophysis), males with eight tubular supplements with pointed tips and the spacing of adjacent supplements almost equal except that of the most anterior two supplements (
<emphasis id="41CC8267FF8A34795C8DFCD5FAED9BD9" box="[1258,1281,871,896]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
seven tubular supplements with truncated tips, and there is a distinct gap between the second and the third supplements), female body slender (a=64.6
<emphasis id="41CC8267FF8A347959F6FC1DFE449B91" box="[401,424,943,968]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
36.8–43.6), vulva situated at midventral side of body
<emphasis id="41CC8267FF8A34795C71FC1DFBC19B91" box="[1046,1069,943,968]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
right-ventrosublateral side in
<taxonomicName id="B4B825F6FF8A34795DE1FC1DFF319BB5" authorityName="Lorenzen" authorityYear="1972" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="macer">
<emphasis id="41CC8267FF8A34795DE1FC1DFF319BB5" italics="true" pageId="5" pageNumber="389">L. macer</emphasis>
</taxonomicName>
. In body size and number of precloacal supplements, the new species is also similar to
<taxonomicName id="B4B825F6FF8A34795CEDFC61FD419C48" authority="(Schuurmans-Stekhoven &amp; De Coninck, 1933) Gerlach, 1958" authorityName="Gerlach" authorityYear="1958" baseAuthorityName="Schuurmans-Stekhoven &amp; De Coninck" baseAuthorityYear="1933" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="elegans">
<emphasis id="41CC8267FF8A34795CEDFC61FB179BB4" box="[1162,1275,979,1005]" italics="true" pageId="5" pageNumber="389">L. elegans</emphasis>
(Schuurmans-Stekhoven &amp; De Coninck, 1933) Gerlach, 1958
</taxonomicName>
and
<taxonomicName id="B4B825F6FF8A34795A83FC45FBAF9C48" authority="Platt, 1973" authorityName="Platt" authorityYear="1973" box="[740,1091,1015,1041]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="septempapillatus">
<emphasis id="41CC8267FF8A34795A83FC45FC539C48" box="[740,959,1015,1041]" italics="true" pageId="5" pageNumber="389">L. septempapillatus</emphasis>
<bibRefCitation id="17292384FF8A34795BA1FC45FBAF9C48" author="Platt, H. M." box="[966,1091,1015,1041]" pageId="5" pageNumber="389" pagination="295 - 321" refId="ref5328" refString="Platt, H. M. (1973) Freeliving marine nematodes from Strangford Lough, Northern Ireland. Cahiers de Biologie Marine, 14 (3), 295 - 321." type="journal article" year="1973">Platt, 1973</bibRefCitation>
</taxonomicName>
, but differs from
<taxonomicName id="B4B825F6FF8A34795D61FC45FA969C48" authorityName="Gerlach" authorityYear="1958" baseAuthorityName="Schuurmans-Stekhoven &amp; De Coninck" baseAuthorityYear="1933" box="[1286,1402,1015,1041]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="elegans">
<emphasis id="41CC8267FF8A34795D61FC45FA969C48" box="[1286,1402,1015,1041]" italics="true" pageId="5" pageNumber="389">L. elegans</emphasis>
</taxonomicName>
by spicules slender, arcuated and cephalated proximally
<emphasis id="41CC8267FF8A34795A8EFBA9FCEC9C6D" box="[745,768,1051,1076]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
spicules slightly bent, knife-like; eight tubular supplements sickle-shaped with pointed distal ends
<emphasis id="41CC8267FF8A34795A3FFB8DFD839C01" box="[600,623,1087,1112]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
5–8 tubular supplements weakly S-shaped with dentate tips. The new species differs from
<taxonomicName id="B4B825F6FF8A347959E3FBD1FD8F9C24" authorityName="Platt" authorityYear="1973" box="[388,611,1123,1149]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="5" pageNumber="389" phylum="Nematoda" rank="species" species="septempapillatus">
<emphasis id="41CC8267FF8A347959E3FBD1FD8F9C24" box="[388,611,1123,1149]" italics="true" pageId="5" pageNumber="389">L. septempapillatus</emphasis>
</taxonomicName>
by spicules 18 µm long, slender and cephalated proximally
<emphasis id="41CC8267FF8A34795D46FBD1FAD49C25" box="[1313,1336,1123,1148]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
spicules 22–26 µm long, knife-like; gubernaculum small with dorso-caudal apophyses
<emphasis id="41CC8267FF8A34795C62FB35FBF09CF9" box="[1029,1052,1159,1184]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
rod-like without apophyses; eight tubular supplements sickle-shaped without cuticular collars around their distal ends
<emphasis id="41CC8267FF8A34795C36FB19FB849C9D" box="[1105,1128,1195,1220]" italics="true" pageId="5" pageNumber="389">vs</emphasis>
seven tubular supplements straight and cephalated proximally and knot-like distally with dentate tips and with cuticular collars around their distal ends. The new species can be identified from other known species by combination of characters of cephalic setae 5–6 µm long, males with eight almost equal spaced sickle-shaped supplements, without alveolar supplement, gubernaculum with a straight dorso-caudal apophysis; female without supplements, vulva situated at midventral side of body.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="8CFCCE1C58171EB405B3BE0676F44817" ID-DOI="10.11646/zootaxa.5507.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757765" ID-ZooBank="732EF0D1-7CDA-4366-B164-156D4E2E8E7F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726221687458" checkinUser="plazi" docAuthor="Gu, Huimin, Sun, Lingyun &amp; Huang, Yong" docDate="2024" docId="FB11EF63FF8E347D58F0FB8CFD979D0C" docLanguage="en" docName="zootaxa.5507.2.9.pdf" docOrigin="Zootaxa 5507 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.2.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptolaimus de Man 1876" docType="treatment" docVersion="2" lastPageNumber="385" masterDocId="0728971BFF8F347C5867FFB2FFEC9859" masterDocTitle="Two new species of Leptolaimus (Nematoda: Plectida) from Chinese sea area" masterLastPageNumber="394" masterPageNumber="384" pageNumber="385" updateTime="1726222090683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="521F52F699131070605B09944F5622E3">Two new species of Leptolaimus (Nematoda: Plectida) from Chinese sea area</mods:title>
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<mods:namePart id="32BAD50F88BFC1D6B1A6CB839A53027F">Gu, Huimin</mods:namePart>
<mods:nameIdentifier id="4ECD041DD7247EA9BB8C02A2AF533AF3" type="ORCID">0009-0008-5820-4678</mods:nameIdentifier>
<mods:affiliation id="7DD5791CF627E0595843D72F521C7182">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
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<mods:namePart id="522630C719B57C191D85B6FB90EE9E6A">Sun, Lingyun</mods:namePart>
<mods:nameIdentifier id="0934B687D655179C7E1E9227A50151BD" type="ORCID">0009-0005-3553-3645</mods:nameIdentifier>
<mods:affiliation id="83A877BFAF063E68718FB91CFFF8EA62">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
<mods:nameIdentifier id="5991CEF3548C57B209E25C93E91F818A" type="email">981779432@qq.com</mods:nameIdentifier>
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<mods:namePart id="D804F27EC494EF75C3C565ED9B4FDA6B">Huang, Yong</mods:namePart>
<mods:affiliation id="6BD234DDC60305816F6A0D8EAA476E50">College of Life Sciences, Liaocheng University, Liaocheng 252059, P. R. China</mods:affiliation>
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<mods:title id="2EC055DC805C4B87E6DCAB20AF88AB33">Zootaxa</mods:title>
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<mods:date id="70CECB6A4D04041B07A23909FEA0DD66">2024</mods:date>
<mods:detail id="07CBB71E4729B0CF13D829873E4D41F1" type="pubDate">
<mods:number id="0DA28F8BD6E72BF1214B5CE5D4FD3602">2024-09-12</mods:number>
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<mods:detail id="A23F228E61FECF39D846EFCCEB4F9336" type="volume">
<mods:number id="164549AB6BB740445E9BF4A5B19DFB23">5507</mods:number>
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<mods:url id="348D38652E748C43DADA099AFB73E720">http://dx.doi.org/10.11646/zootaxa.5507.2.9</mods:url>
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<mods:identifier id="A7DB6F352CEE9CD02DF0C2DB3CA28A6A" type="DOI">10.11646/zootaxa.5507.2.9</mods:identifier>
<mods:identifier id="6ACC5AA07B4571F6DE1D146A83673D63" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C3704DB1EB7E5CFDF477C8440BD87FC5" type="Zenodo-Dep">13757765</mods:identifier>
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<treatment id="FB11EF63FF8E347D58F0FB8CFD979D0C" ID-DOI="http://doi.org/10.5281/zenodo.13757777" ID-Zenodo-Dep="13757777" LSID="urn:lsid:plazi:treatment:FB11EF63FF8E347D58F0FB8CFD979D0C" httpUri="http://treatment.plazi.org/id/FB11EF63FF8E347D58F0FB8CFD979D0C" lastPageNumber="385" pageId="1" pageNumber="385">
<subSubSection id="3BA20DFEFF8E347D58F0FB8CFDD99C01" box="[151,565,1086,1113]" pageId="1" pageNumber="385" type="nomenclature">
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Genus
<taxonomicName id="B4B825F6FF8E347D5889FB8CFDD99C01" ID-CoL="87GVG" authority="de Man, 1876" authorityName="de Man" authorityYear="1876" box="[238,565,1086,1113]" class="Chromadorea" family="Leptolaimidae" genus="Leptolaimus" kingdom="Animalia" order="Plectida" pageId="1" pageNumber="385" phylum="Nematoda" rank="genus">
<emphasis id="41CC8267FF8E347D5889FB8CFE689C00" bold="true" box="[238,388,1086,1113]" italics="true" pageId="1" pageNumber="385">Leptolaimus</emphasis>
de Man, 1876
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<emphasis id="41CC8267FF8E347D58F0FB34FEFE9CF9" bold="true" box="[151,274,1158,1184]" pageId="1" pageNumber="385">Diagnosis.</emphasis>
(Modified from Holovachov 2014) Body slender, annulated cuticle usually with lateral alae; cephalic sensilla papilliform or setiform; amphidial fovea ventrally unispiral (circular in outline); tubular buccal cavity elongated; males reproductive system diorchic, may having both tubular and alveolar precloacal supplements, one or no supplement at all; the number of supplements varying from zero to 16 for tubular and zero to 56 for alveolar; females with two opposed and reflexed ovaries, vulva midventral or subventral; alveolar or tubular supplements may be present in females of some species.
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