Add updates up until 2025-02-07 18:27:38

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<document id="9E1FB7B1CBE49EB3D8EBEEB8FF720E18" ID-DOI="10.3897/zookeys.1226.142678" ID-ZooBank="0B9EFAA1-3377-42F0-B43A-B77F439F88FE" ID-publisher-id="142678" URI-arpha="49D82D8E-47F0-5A5B-A344-09EDC69A48BD" URI-zoobank="http://zoobank.org/0B9EFAA1-3377-42F0-B43A-B77F439F88FE" article-type="research-article" checkinTime="1738952664858" checkinUser="pensoft" docAuthor="Chen, Jia-Heng, Shi, Hong-Liang &amp; Liang, Hong-Bin" docDate="2025" docId="06017AE76FCB5E6A856EC278A60E4487" docLanguage="en" docName="ZooKeys 1226: 183-193" docOrigin="ZooKeys 1226" docSource="https://zookeys.pensoft.net/article/142678/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Megadrypta mirabilis Sciaky &amp; Anichtchenko 2020" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="193" masterDocId="49D82D8E47F05A5BA34409EDC69A48BD" masterDocTitle="Revision of the Chinese endemic genus Megadrypta Sciaky &amp; Anichtchenko, 2020 (Coleoptera, Carabidae, Dryptini), with description of a new species" masterLastPageNumber="193" masterPageNumber="183" pageNumber="183" updateTime="1738953159347" updateUser="ExternalLinkService">
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<mods:title id="A66E0836216F14D5689294BF867366C6">Revision of the Chinese endemic genus Megadrypta Sciaky &amp; Anichtchenko, 2020 (Coleoptera, Carabidae, Dryptini), with description of a new species</mods:title>
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<mods:namePart id="4B61922DBB6C53EC683DDA21044AB16B">Chen, Jia-Heng</mods:namePart>
<mods:nameIdentifier id="7FDEF14EDA5BB4FF2D5E86F0ACBD716D" type="ORCID">0000-0003-2449-8838</mods:nameIdentifier>
<mods:affiliation id="59BC76FD75357A8D2F3C3D6B0E3D2760">College of Forestry, Beijing Forestry University, Beijing, 100083, China</mods:affiliation>
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<mods:namePart id="631C7F144113F15DCBCD5CD38177D775">Shi, Hong-Liang</mods:namePart>
<mods:nameIdentifier id="3EB6E92C29D82D7AD67E7E423D024F6F" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
<mods:affiliation id="EC26F3BA89020BF3CDD7EE3C35639AB0">College of Forestry, Beijing Forestry University, Beijing, 100083, China</mods:affiliation>
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<mods:namePart id="CD78EFFE0EB4078CB0BA6DB0AE54FC26">Liang, Hong-Bin</mods:namePart>
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<mods:affiliation id="2A65DA4841664A164CCE7A71CB8505EA">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:date id="9CFC7B4A75466FB620724FFC66E43324">2025</mods:date>
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<treatment id="06017AE76FCB5E6A856EC278A60E4487" ID-DOI="http://doi.org/10.5281/zenodo.14834508" ID-Zenodo-Dep="14834508" ID-arpha="06017AE7-6FCB-5E6A-856E-C278A60E4487" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487" httpUri="http://treatment.plazi.org/id/06017AE76FCB5E6A856EC278A60E4487">
<subSubSection id="7EAF8494475C95AA1034A67078A83721" type="nomenclature">
<paragraph id="3DB30821E183636BFCF24BA94315330A">
<taxonomicName id="98A653347EA3E9E025360334AB58D8E4" ID-CoL="8MKXP" authority="Sciaky &amp; Anichtchenko, 2020" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="774930FAF0AEA8A78DE0084C76BAD44E" italics="true">Megadrypta mirabilis</emphasis>
<bibRefCitation id="5E4776CD2C8EA7CCEB507BE57BDD27B9" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="1CEE93AC06AC1409AE90B389F87CCA94" type="description">
<paragraph id="9AF970DA573E5E622AF911CC3733A660">
<figureCitation id="BF2327816D86C87542139417D3810114" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Megadrypta spp. A holotype of M. maozhoui sp. nov. B female of M. mirabilis from Hunan. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure2" httpUri="https://binary.pensoft.net/fig/1244300">Figs 2 B</figureCitation>
,
<figureCitation id="2675DE86D9ADB37611220ACC583A6F5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4</figureCitation>
,
<figureCitation id="7F3F6146268C412BACC477FD8C4859F8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mentum of Megadrypta spp. A M. maozhoui sp. nov. B M. mirabilis. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1226.142678.figure6" httpUri="https://binary.pensoft.net/fig/1244304">6 B Chinese vernacular name: 巨逮步甲</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="36164DCCB03C692942AA5CA74C73F35F">
<treatmentCitationGroup id="A096AE2093C55688F5676F7DCBDC5E3E">
<treatmentCitation id="6F6BB8D9CF7AE420B067F2F56C911098" ID-DOI="http://doi.org/10.5281/zenodo.3665229" ID-Zenodo-Dep="3665229" author="Sciaky R &amp; Anichtchenko A" httpUri="http://treatment.plazi.org/id/03D77E1287166B19FF0EFF756042FE6D" page="526" year="2020">
<taxonomicName id="19781B501A3ECDCFED65C351520E479D" ID-CoL="8MKXP" authority="Sciaky &amp; Anichtchenko, 2020: 526" authorityName="Sciaky &amp; Anichtchenko" authorityPageNumber="526" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="071A90FB4C91C2177FC28D5016969B9D" italics="true">Megadrypta mirabilis</emphasis>
<comment id="458CE610048AAA577BB3EF00B3482E86">
<bibRefCitation id="F15606126D81A7CE14E8ED8E2CECC8E3" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020: 526</bibRefCitation>
.
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<subSubSection id="SECID0ESSAE" type="material">
<paragraph id="8404AD70B381A837A1323B83CAA660B8">
<heading id="817C8C7A2FBBA03169BA8FD5839DB513" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="B6218FCF7BB4138BDC72706DEC397599">
<materialsCitation id="6BED4E375903C44EF5B40FFEB00DC904" collectingDate="2007-06-18" collectionCode="IZAS" country="China" elevation="1021" latitude="24.92579" location="Nanling Nature Reserve" longLatPrecision="1" longitude="113.01638" municipality="Ruyuan" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="00E82B6AC2FCD2F4426E0F61EBA95F02" bold="true" italics="true">
<typeStatus id="52BC1256A25BEFFF6B700972A8E6927A" type="holotype">Holotype</typeStatus>
</emphasis>
: China • male, labeled “
<collectingCountry id="A3D3833606A708AECD0527DC441983D7" name="China">CHINA</collectingCountry>
.
<collectingRegion id="BAFA0B52D982924A7765C4D9615FE2A5" country="China" name="Guangdong">Guangdong</collectingRegion>
,
<collectingMunicipality id="7AE097D9B026EA54A28EE825D82E8E8F">Ruyuan</collectingMunicipality>
,
<location id="FD0D9C0CF1D9109E67568CBD259A64DB" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:FD0D9C0CF1D9109E67568CBD259A64DB" country="China" latitude="24.92579" longLatPrecision="1" longitude="113.01638" municipality="Ruyuan" name="Nanling Nature Reserve" stateProvince="Guangdong">Nanling Nature Reserve</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EATAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[113.016380,24.925790]}">
<geoCoordinate id="0D662BA0A3C62B1D405F6CA521D0A09F" degrees="24.92579" direction="north" orientation="latitude" precision="1" value="24.92579">24.92579</geoCoordinate>
,
<geoCoordinate id="F4266FCDC8150FD4DC73EB153001651E" degrees="113.01638" direction="east" orientation="longitude" precision="1" value="113.01638">113.01638</geoCoordinate>
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”; “
<quantity id="B078BB8DBB7D5C4DFCF85CD4CD6EEFE0" metricMagnitude="3" metricUnit="m" metricValue="1.021" unit="m" value="1021.0">
<elevation id="D6CAFA3B08923F590CCCBDB7991601F8" metricMagnitude="3" metricUnit="m" metricValue="1.021" unit="m" value="1021.0">1021 m</elevation>
</quantity>
,
<date id="A20AB61738CF16CB049A504C308DE2D5" value="2007-06-18">
<collectingDate id="7B22CE123004FD8710AA98D1595D2377" value="2007-06-18">2007.6. 18</collectingDate>
</date>
21 Huang Hao ”;
<named-content id="C5CE2AA5BB5618FB8AEB8EC36A9A71F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2093EE0FE670E57CE0E85274CECCB813" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="9834629EB8B3A18ABEADEBFC94C24279" country="China" location="China" specimenCount="1" typeStatus="paratype">
<emphasis id="19D16AA583347DBB02B3383AE08FD497" bold="true" italics="true">
<typeStatus id="652C1CC576C6512AA00268DD3BD305E2" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectingCountry id="B23F5C935FA9E0A9F86C62357507AE4A" name="China">China</collectingCountry>
</materialsCitation>
<materialsCitation id="30C992FF536D2EA7AAD1F2147C6F4186" collectingDate="2007-08-05" collectionCode="IZAS" collectorName="Liu Chunxiang" country="China" location="Cenwanglao Shan Nature Reserve" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="E21798F1451497B41BC86C48BB355EAF" count="1" type="male">1 male</specimenCount>
, labeled “
<collectingCountry id="7E23358F7DDD2481E1684C56C08E4AA8" name="China">CHINA</collectingCountry>
.
<collectingRegion id="F9A9F7BC066142C51DF60FEF56CE1C60" country="China" name="Guangxi">Guangxi Prov.</collectingRegion>
,
<location id="7B32241CBAB0A8E76AD98D7B545AFD77" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:7B32241CBAB0A8E76AD98D7B545AFD77" country="China" name="Cenwanglao Shan Nature Reserve" stateProvince="Guangxi">Cenwanglao Shan Nature Reserve</location>
”; “
<date id="7EEE8A7F2729E0CD5662C810CC0A4F29" value="2007-08-05">
<collectingDate id="56B4B88752543B3910611592FDC28DFE" value="2007-08-05">2007.8. 5</collectingDate>
</date>
day,
<collectorName id="DFA513DFF114663C0B55E57FB8F2B222">Liu Chunxiang</collectorName>
coll. ”;
<named-content id="7E15B28BDD977D1E9D9C7A8699F846E9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="5D9F1D30FC1D150B5994B65DD565E5F1" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQTAE" type="type material">
<paragraph id="B7AD19734044F3CD64DA18C34420AD9F">
<heading id="68BCC387A0FA24B3CAD6C474E25D6B99" reason="title">Non-type materials.</heading>
</paragraph>
<paragraph id="44026FFBFB87A0DD0CB8A433E936B6A4">
<collectingCountry id="8D2A1063C95A04E263767B70F3B6929E" name="China">China</collectingCountry>
<materialsCitation id="7D9C7BA3F7F3D9E08B42FCA72607AE80" collectingDate="2020-07-28" collectionCode="CHYZ" country="China" county="Mt. Mangshan" elevation="1200" location="Chinese" municipality="Jiangjunzhai" specimenCount="1" specimenCount-female="1" stateProvince="Hunan">
<specimenCount id="21B7D445ADA4F72ED67FC7DD4E3411B6" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingCountry id="C45B4B143BD425D7394F7570EFFDF603" name="China">CHINA</collectingCountry>
:
<collectingRegion id="E15E7281A02B7D8020F3DAE3D07CE5FD" country="China" name="Hunan">Hunan</collectingRegion>
,
<collectingCounty id="76EFCD882163070DA3600D5BE5BF829A">Mt. Mangshan</collectingCounty>
,
<collectingMunicipality id="523957E84B2030AB371381FED85A8BCF">Jiangjunzhai</collectingMunicipality>
[in
<location id="79B9E8812DE3E31D018FE85EBEBC34CD" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:79B9E8812DE3E31D018FE85EBEBC34CD" country="China" county="Mt. Mangshan" municipality="Jiangjunzhai" name="Chinese" stateProvince="Hunan">Chinese</location>
],
<date id="83B0181AE7D416378E77A564EB92F635" value="2020-07-28">
<collectingDate id="FCFC4E63B78FB21841523E8B01B756C2" value="2020-07-28">2020. VII- 28</collectingDate>
</date>
,
<quantity id="03093E05D415636D80BBCA9BBDF18E9C" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="8324B96246F5A6C45FD6F44ADE63EA91" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, Leg. Yuzhou Huang ”;
<abbrev id="ABBRID0EWTAE" xlink_title="personal collection of Yu-Zhou Huang">
<collectionCode id="1F57CB85D8ABB8B6B61BA4C19FADD6F6">CHYZ</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="717A5D166B1FA77ED904E90E5B1F6D99" collectingDate="2024-06-01" collectionCode="CCJH" collectorName="Chunfu" country="China" county="Jinxiu County" location="Changtong Township" municipality="Laibin" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi">
<specimenCount id="AF5918D561C6A16BF015C6D7F27FAE15" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingCountry id="1DF463C25E85416FA076F468944DCA62" name="China">CHINA</collectingCountry>
:
<collectingRegion id="E4BF9AF5F6A6942E1AD147859848A621" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingMunicipality id="385CD5A650075F59FEF762C1DD0331E9">Laibin</collectingMunicipality>
,
<collectingCounty id="010CD3D8A41BF8885C526F763F6AF949">Jinxiu County</collectingCounty>
,
<location id="2A4F5327BB9AF2806CFBDCAE17BF25CC" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:2A4F5327BB9AF2806CFBDCAE17BF25CC" country="China" county="Jinxiu County" municipality="Laibin" name="Changtong Township" stateProvince="Guangxi">Changtong Township</location>
,
<location id="E752FA9B32DDF7AC3AB71D2134E16116" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:E752FA9B32DDF7AC3AB71D2134E16116" country="China" county="Jinxiu County" municipality="Laibin" name="Guitian Village" stateProvince="Guangxi">Guitian Village</location>
,
<date id="C06CB1B0A0BD160AB3B690864EF0101C" value="2024-06-01">
<collectingDate id="C241A39E7123B5763BC4C75A086A81B7" value="2024-06-01">2024. VI- 1</collectingDate>
</date>
, leg.
<collectorName id="B37C698952D872820AA695444F4BE777">Chunfu</collectorName>
FENG ”;
<abbrev id="ABBRID0E1TAE" xlink_title="personal collection of Jia-Heng Chen">
<collectionCode id="245BD3D62A2475D488C9A37B70310C18">CCJH</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="68FAA32532A076E1B5C6DF69CBAE81C8" collectionCode="CCJH" collectorName="Chinese" country="China" location="Nanling" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong">
<specimenCount id="AB86DAB474A82D93D18EF75EEF5E2ED3" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingRegion id="B02133BEF14E1ED786B79F4866455ED5" country="China" name="Guangdong">Guangdong</collectingRegion>
,
<location id="3B05544BF854917E4F57E6ED5E7301A6" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:3B05544BF854917E4F57E6ED5E7301A6" country="China" name="Nanling" stateProvince="Guangdong">Nanling</location>
, 2017-8 ” [in
<collectorName id="FEDBF11B12A1ED7F7C0E85327C1D0D0A">Chinese</collectorName>
];
<abbrev id="ABBRID0E5TAE" xlink_title="personal collection of Jia-Heng Chen">
<collectionCode id="25BB9A86127E7D39F1DAE10765930138">CCJH</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="033E5F03FBDD09053B43CCF0DABE0764" collectingDate="2024-05-31" collectionCode="CYXH" country="China" county="Leishan County" elevation="1555" latitude="26.373032" location="Chinese" longLatPrecision="1" longitude="108.19808" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
<specimenCount id="C8F686831D38CEB5DDC97B44B7F19D49" count="1" type="male">1 male</specimenCount>
, labeled “
<collectingCountry id="2685EF4FFB6FCC1E8FE5657A6C024799" name="China">CHINA</collectingCountry>
:
<collectingRegion id="6355B379BBECC92A6A574BF86A080628" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingCounty id="8D573B9619DF09835B86A4AB1BBE0F8C">Leishan County</collectingCounty>
, Mt. Leigongshan, Xiannütang,
<quantity id="682E97FAFAD4C698164829B146324BD3" metricMagnitude="3" metricUnit="m" metricValue="1.555" unit="m" value="1555.0">
<elevation id="7FB17D4CF1071DA23F555AB05BF965D3" metricMagnitude="3" metricUnit="m" metricValue="1.555" unit="m" value="1555.0">1555 m</elevation>
</quantity>
” [in
<location id="D368900ACA7553264667EBD407BFE4C6" LSID="urn:lsid:plazi:treatment:06017AE76FCB5E6A856EC278A60E4487:D368900ACA7553264667EBD407BFE4C6" country="China" county="Leishan County" latitude="26.373032" longLatPrecision="1" longitude="108.19808" name="Chinese" stateProvince="Guizhou">Chinese</location>
]; “
<date id="1217E1F5064C7C916B8B6485F217F968" value="2024-05-31">
<collectingDate id="40CCFAE3D0142D303E61789FAD68BDAB" value="2024-05-31">2024. V. 31</collectingDate>
</date>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFUAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.198085,26.373031]}">
<geoCoordinate id="13840A7C721C9F2D510102867D3B4437" degrees="26.37303146" direction="north" orientation="latitude" precision="1" value="26.373032">26.37303146 N</geoCoordinate>
<geoCoordinate id="6B523E19F20B341D172772681DACFB7F" degrees="108.19808469" direction="east" orientation="longitude" precision="1" value="108.19808">108.19808469 E</geoCoordinate>
</named-content>
, Leg. Ye ”;
<abbrev id="ABBRID0EKUAE" xlink_title="personal collection of Xiao-Han Ye">
<collectionCode id="0FB8938DFE6C57F6487AA22DD9F18CDB">CYXH</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOUAE" type="diagnosis">
<paragraph id="9EDB68305BA9F0AF622B456A05272EBA">
<heading id="9C6FDF4C8E29189B29D09B16AF62EFBE" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2574E21262AF60E8C43563174BAE4168">Dorsal surface nearly black, with very weak bluish luster; femora dark brown to piceous; mouth parts, tibiae, and tarsi sometimes lighter. Mentum with anterior margin straight at middle. Elytra narrowed toward shoulders; elytral striae incised with large punctures (much smaller than interval width). Median lobe of aedeagus in lateral view equally broadened before apical third and then abruptly narrowed; ventral margin strongly curved before apex.</paragraph>
</subSubSection>
<subSubSection id="SECID0ETUAE" type="description">
<paragraph id="BE7333AFFAC9D237D25D5D618BAC69C6">
<heading id="F3CC27E7BF38E600FAB53C2A56F81EB8" reason="title">Supplementary description.</heading>
</paragraph>
<paragraph id="6C9BDB89916DC871BCC72CE815B586B7">
<abbrev id="ABBRID0EZUAE" xlink_title="length of the body">BL</abbrev>
<quantity id="C82AE90E94BDEF6CC5E29DD08C11F18B" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.61" metricValueMin="1.39" unit="mm" value="15.0" valueMax="16.1" valueMin="13.9">13.916.1 mm</quantity>
(15.0 mm in the
<typeStatus id="5B1131BC281FB66BEA49CF3E21F4AA6A" type="holotype">holotype</typeStatus>
); dorsal and ventral surface nearly black, with very weak bluish luster; mouth parts brown; femora dark brown to piceous; tibiae and tarsi sometimes lighter (Fig.
<figureCitation id="5A33F13F673C1860824761ABD7C26F3A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Megadrypta spp. A holotype of M. maozhoui sp. nov. B female of M. mirabilis from Hunan. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure2" httpUri="https://binary.pensoft.net/fig/1244300">2 B</figureCitation>
).
<emphasis id="9F7257416943F5F17BBC31BA6BC93C38" bold="true" italics="true">Head</emphasis>
: labrum trilobate; medial lobe slightly extended anteriorly; anterolateral angles rounded-obtuse, sparsely pubescent near anterior margin; clypeus impunctate and pubescent; vertex densely and coarsely punctate; temporae slightly shorter than eyes, well tumid behind eyes, sparsely and finely pubescent and punctate.
<emphasis id="D9B52A6FD3329C63018D5F39FEC72E33" bold="true" italics="true">Pronotum</emphasis>
:
<abbrev id="ABBRID0EHVAE" xlink_title="length of the pronotum">PL</abbrev>
/
<abbrev id="ABBRID0ELVAE" xlink_title="width of the pronotum">PW</abbrev>
1.621.68; lateral margins gradually sinuate twice, behind anterior angles and before posterior angles; sides flattened from above and a little extended laterally near middle, median region elevated.
<emphasis id="A5F67C79683F2ECC52C70122D425B9F4" bold="true" italics="true">Elytra</emphasis>
:
<abbrev id="ABBRID0ESVAE" xlink_title="length of the elytra">EL</abbrev>
/
<abbrev id="ABBRID0EWVAE" xlink_title="width of the elytra">EW</abbrev>
1.441.50; base with 78 parascutellar pores, strongly narrowed at shoulders; elytral striae incised with large punctures (much smaller than interval width).
<emphasis id="DCFE134934A8B3EE112386182F6512EF" bold="true" italics="true">Male genitalia</emphasis>
: median lobe of aedeagus relatively stout (Fig.
<figureCitation id="8BF7FBB54EB4C52AE32249EE0FCD9350" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4</figureCitation>
), equally thickened before apical third and then abruptly narrowed in lateral view, with ventral margin strongly curved before apical lamella (Fig.
<figureCitation id="2E185BEFDF6B0B59479F3D1930EE8B31" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4 A, D</figureCitation>
); same in dorsal view, with apical lamella short (
<abbrev id="ABBRID0EFWAE" xlink_title="length of the apical lamella of the aedeagus">ALL</abbrev>
/
<abbrev id="ABBRID0EJWAE" xlink_title="width of apical lamella of the aedeagus">ALW</abbrev>
0.460.51), obtuse and rounded; endophallus with a central sclerite, hook-shaped in right lateral view (Fig.
<figureCitation id="466EECC1F7C4E1236C9005EAE8F78C9C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4 A</figureCitation>
), with blurred border in ventral view (Fig.
<figureCitation id="462FF97227E0CDABFB544C3E018D3525" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVWAE" type="distribution">
<paragraph id="6B7252A2599C93E2015236DB3F84101D">
<heading id="EB941F1B26318787C4B4AF5C230BCD23" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="476168FF19C73B1903933E70F807F562">
This species is widely distributed in the Nanling Area of southern
<collectingCountry id="1D7CEDA0A83A4CB3E1BE2E50AD2B48F6" name="China">China</collectingCountry>
(Fig.
<figureCitation id="33B4CBA1512E2D7C9A48ABABB2C850BF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map for Megadrypta spp. Red circles: M. mirabilis Sciaky &amp; Anichtchenko; yellow circles: M. maozhoui sp. nov." figureDoi="10.3897/zookeys.1226.142678.figure1" httpUri="https://binary.pensoft.net/fig/1244299">1</figureCitation>
), where it is currently recorded from four provinces: in southern
<collectingRegion id="FA60D7AA9C98294492EDE7F4F972E73A" country="China" name="Guizhou">Guizhou</collectingRegion>
from Mount Leigongshan (new provincial record); in southern
<collectingRegion id="57B97945F085511BDC8CA91754C6603A" country="China" name="Hunan">Hunan</collectingRegion>
from Mount Mangshan (new provincial record); in north
<collectingRegion id="5532B18C27EABBCDB2D4E9F75F00F119" country="China" name="Guangxi">Guangxi</collectingRegion>
from Mounts Maoershan, Dayaoshan, and Cenwanglaoshan; and in northern
<collectingRegion id="1056ED6F6D2078AA8194898D57B3647B" country="China" name="Guangdong">Guangdong</collectingRegion>
from the Nanling Natural Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="8E1D8468CF50D235CDA5DBEB5B8ABD59" ID-DOI="10.3897/afrinvertebr.66.129611" ID-ZooBank="9A70A477-D862-40E5-A132-3227D96C3ECC" ID-publisher-id="129611" URI-arpha="5630D620-B234-5FD1-B5C0-EDF357AAF7CA" URI-zoobank="http://zoobank.org/9A70A477-D862-40E5-A132-3227D96C3ECC" article-type="research-article" checkinTime="1738952593943" checkinUser="pensoft" docAuthor="Rollinson, Lisa &amp; Cabrero, Allan" docDate="2025" docId="1FE42C3A1274556DB7D05F6D27DCDC85" docLanguage="en" docName="African Invertebrates 66(1): 73-115" docOrigin="African Invertebrates 66 (1)" docSource="https://africaninvertebrates.pensoft.net/article/129611/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Enica adelphe Rollinson &amp; Cabrero, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="115" masterDocId="5630D620B2345FD1B5C0EDF357AAF7CA" masterDocTitle="Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834" masterLastPageNumber="115" masterPageNumber="73" pageNumber="73" updateTime="1738953078235" updateUser="ExternalLinkService">
<mods:mods id="91A33B977FDF8AC85A4E801E14AE6AAB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3FC10A27934679ADB3047A68F39E722B">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
</mods:titleInfo>
<mods:name id="C5E3BC7CA59A9D378664CFB44A50E2BC" type="personal">
<mods:role id="A86F500DA22055563D8AC3A4661C1F60">
<mods:roleTerm id="65514200DAF7C40A2D32F19747FEABA6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="604D97372CB586C62696C53E26239EFF">Rollinson, Lisa</mods:namePart>
<mods:nameIdentifier id="44C32CC8608D092DE4BC679918378431" type="ORCID">https://orcid.org/0009-0007-0152-7073</mods:nameIdentifier>
<mods:affiliation id="6BB3D499BACD8EBB64F8BAAF4038978B">Texas A &amp; M University, College Station, Texas, USA</mods:affiliation>
</mods:name>
<mods:name id="AA53BD70A7AF7F235612413D10399F21" type="personal">
<mods:role id="4D247F398097823648EC1BC8F1C248C8">
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</mods:role>
<mods:namePart id="D851C362C1B7C55BA8FF58CA526CBDF0">Cabrero, Allan</mods:namePart>
<mods:nameIdentifier id="9005B163C6B2EDE94087EEC4BE9D7C3D" type="ORCID">0000-0002-2935-3534</mods:nameIdentifier>
<mods:affiliation id="45C79487E3EC375224830A7349532621">National Museum of Natural History, Washington, D. C., USA</mods:affiliation>
</mods:name>
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<mods:titleInfo id="84745A69538A1CA0429EE65262277CFA">
<mods:title id="BF959CCCFD543E6DFD32A1F8CE522FA5">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="0C29E4B4ECE878DB5CA7804BF6B1E1FA">
<mods:date id="DEF9A6EBD89D4D33A13E52DF4C5334FD">2025</mods:date>
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<mods:number id="A939E835C1F95295E46A75B3EC13CCBF">66</mods:number>
</mods:detail>
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<mods:identifier id="EFF939CE7B1AA8AE065E2F0FC6D10537" type="ZooBank">9A70A477-D862-40E5-A132-3227D96C3ECC</mods:identifier>
</mods:mods>
<treatment id="1FE42C3A1274556DB7D05F6D27DCDC85" ID-DOI="http://doi.org/10.5281/zenodo.14834486" ID-Zenodo-Dep="14834486" ID-arpha="1FE42C3A-1274-556D-B7D0-5F6D27DCDC85" ID-zoobank="https://zoobank.org/4BD142FC-44B9-4C63-9524-FC88187C4E6D" LSID="urn:lsid:plazi:treatment:1FE42C3A1274556DB7D05F6D27DCDC85" LSID-ZBK="urn:lsid:zoobank.org:act:4BD142FC-44B9-4C63-9524-FC88187C4E6D" httpUri="http://treatment.plazi.org/id/1FE42C3A1274556DB7D05F6D27DCDC85">
<subSubSection id="C52A3DB368AD354A030F12B24CBFEC9E" type="nomenclature">
<paragraph id="FA47A9363DDE7097196C4A7342CB39D2">
<taxonomicName id="4A78E570B854942F0A1BC0FC645BC0B2" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="EC3BAD8653C69ADEAECA7A85DEDBA440" italics="true">Enica adelphe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B2B975C95BABBA0F89458A0A7BC760" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CDDAAB84C6B13B37DC3137E7071550AF" type="description">
<paragraph id="A20890B1D1F6A14BE0E787201B98CDB7">
<figureCitation id="EB9480774C4B8D15757707889BCFAF59" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Enica adelphe sp. nov. NMSA - DIP- 89520 ♂, holotype (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica adelphe sp. nov SAM-DIP-A 004969, ♀ (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure3" httpUri="https://binary.pensoft.net/fig/1244176">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIVAE" type="diagnosis">
<paragraph id="D5345088507B98CEEE68B8899B512E98">
<heading id="3625E617E8C54F31B51499DE63036FE7" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B7EE2B7F7E0C5FE81C4FB5D1E18C30EA">The species is distinguished from other species in the genus by the absence of scales on the occiput, two dark elongated spots on the occiput extending from the dorsal to the lateral margin, red cuticle on the abdomen past tergite 4, and the dark brown scales on the metathoracic femur.</paragraph>
<caption id="4C510378359679D3629A0350C6BD250C" ID-DOI="10.3897/afrinvertebr.66.129611.figure3" ID-arpha="9EC04CF9-DAE1-5B5B-BC32-F421A0310566" httpUri="https://binary.pensoft.net/fig/1244176" startId="F3">
<paragraph id="D5FCA87B4363839F81C20D62A7724B58">
<label id="E614A519F09B8255970FBA592E2671B5">Figure 3.</label>
</paragraph>
<paragraph id="B94AEB214B00476CE10DBBEE825963E6">
<taxonomicName id="85DC1024FC2C8C9324C4B116760E6EEA" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="BAA903B295128B30A85040EF760C35AC" italics="true">Enica adelphe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9E1E317CF25ACE217F919B985BB326B8" rank="species">sp. nov.</taxonomicNameLabel>
<named-content id="EBCA86BDE5EC6733161B14919B8B25CF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP- 89520 ♂, holotype (
<emphasis id="C87B2936304B427F25A1067F57832628" bold="true">A C</emphasis>
)
<emphasis id="CD98B716882B7B423B3929A94007CA6B" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="5BBA1BC9FD5101DD977E522C8769AD95" bold="true">B</emphasis>
habitus, lateral view
<emphasis id="57B649AB4D870EF608F1257A2E3A2E8C" bold="true">C</emphasis>
habitus, frontal view;
<taxonomicName id="3BECE807F2BDBAEE02A320B6E3AE5594" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="8E4D9929CC2CE92180C195AAA2456F32" italics="true">Enica adelphe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A313222723D691C9E912BD69E9C9A670" rank="species">sp. nov</taxonomicNameLabel>
SAM-DIP-A 004969, ♀ (
<emphasis id="B96CD5B3974B88BD25989ECC37A426D0" bold="true">D F</emphasis>
)
<emphasis id="F869E3977AB372BDCE4B5DC3308E1B32" bold="true">D</emphasis>
habitus, frontal view
<emphasis id="F695B7E6AE59B9A8F0D2B141D055A7BB" bold="true">E</emphasis>
habitus, dorsal view
<emphasis id="77C015F2845F01AC9D89FE246C7BD8EB" bold="true">F</emphasis>
habitus, lateral view. Scale bars: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEXAE" type="etymology">
<paragraph id="CDA32EAB9AE4CC7117435D8B12CFAA1A">
<heading id="88E963C07112B6FEA076D6BF000C74CF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="404679D999D278FD278F3FC1F333C32D">This species is named in honor of author L. Rollinsons sister, Jessica Rollinson. The specific epithet is Greek for sister.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXAE" type="description">
<paragraph id="87738238527CC5DA761BF808EBD21C35">
<heading id="52A9E9DC7253213E62FB5BAF6E367426" reason="title">Description.</heading>
</paragraph>
<paragraph id="FAB3B4F68984ED622B6035CE6B6406A2">
<emphasis id="DE93B0320A2AA6F43D7571C835F18675" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, with dark brown to black upside down “ V ” spot not touching eyes, males: similar to head, slightly darker than head; frons black setose, with white scales, males no setation near eyes, females setose near eyes only dorsally; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput whitish yellow setose, without scales, dorsal eye margin with dark brown to black spots present, not touching eye margin, extending more than three times as long as wide; gena significantly darkened to reddish brown or black, extremely sparsely setose.
</paragraph>
<paragraph id="5F43ABDA386847B4E147F79CC2A2546B">
<emphasis id="4C759473E378D5B8F4A90FE8E3A014F4" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, black, brown setose.
</paragraph>
<paragraph id="CB70A569AF4027F01208B98D41734DF5">
<emphasis id="F9FC17DB6971993C6819BA2CE2BFDD36" bold="true" italics="true">Antenna</emphasis>
: black; scape creamy white, approximately as long as pedicel, white and black setose laterally and dorsally, setae extending to tip of pedicel; pedicel dark brown, short black setose dorsally; postpedicel black, tapering distally, longer than scape and pedicel combined, black scales dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="00E5DD63F03F3E004CF4C97CC367AE1F">
<emphasis id="B99865B877329AE44835B707D97D7841" bold="true" italics="true">Thorax</emphasis>
: reddish brown and light orangish brown; scutum predominantly black, reddish on margins, grey to brown scales centrally and white scales peripherally; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, light yellow scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, asetose or white scales dorsally; metepimeron silvery grey pubescence, asetose; scutellum red, anterior black spot or red, medial black stripe, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of yellowish-white scales.
</paragraph>
<paragraph id="5A7D23A1975B40930A8CB41760FAF735">
<emphasis id="9E21D0D098A76D26D08B7937FCCBB853" bold="true" italics="true">Leg</emphasis>
: dark brown, covered in scales, some regular setae present; pro coxa dark reddish brown, silvery grey pubescence, white scales, thin white setose, thick black setose; pro femur dark brown, dark brown scales, black setose; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa dark reddish brown, white pubescence, white scales, thick black setose; mes femur dark brown, dark brown scales, black setose; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa dark reddish brown, white pubescent, white scales, thin white setose, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur dark brown, dark brown scales, long and thick black setose; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="19FEF79151830A604727713C8BF6909D">
<emphasis id="EB2D584A4091977145813F9A0AC97616" bold="true" italics="true">Wing</emphasis>
:
<quantity id="0AB6D427A308974D8CB8C05B1048CFDF" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.8" metricValueMin="8.2" unit="mm" value="9.0" valueMax="9.8" valueMin="8.2">8.29.8 mm</quantity>
, microtrichia absent; membrane color pattern males: minimal spots of infuscation in r 2
<subScript id="DAC932C21FF53E0A49EA76E2F0208CCB">+ 3</subScript>
, br, and bm, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="F61D0F0029B14B0BD31225D36A1DA5CF">+ 3</subScript>
, base of m
<subScript id="E95ADD755CCC3A91D9BFB80BC7BC0AF8">2</subScript>
, m-cu, split between M
<subScript id="432DC8599759384C1A6595B273ADF6CA">1</subScript>
+ 2 and M
<subScript id="1AC17916DFF2BCF20879865B06101C6E">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="740DE7E0280273D6A4DC59EDE0EA7742">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="BEEB5D4C22653C4BDF82CAF11CC63018">1</subScript>
open; R
<subScript id="9921F702ED3F4B15CB060FAA3839E7CB">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="882BA4966B1BA11CCD6B635B2C838C4E">4</subScript>
open, proximally R
<subScript id="537A659427C40AF2368B450A95ED31DB">4</subScript>
and R
<subScript id="55AB1B72434742B70F9B8D47B0A7D6D5">5</subScript>
parallel, R
<subScript id="4B0CE231CD9405606364421A9AFEB6A0">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="42EC63BC98143D898195C44C9F364483">5</subScript>
terminating posterior to wing apex; r
<subScript id="E0F54E5F093904645A9244012F7B911F">5</subScript>
open; M
<subScript id="41F672CA87DCFE21D6F10159DF8660C3">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="A4AD4A292C1E44DBA60D17E057A4D4C2">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="26F8F247586C0CA416AD963DF0D826D2">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.
</paragraph>
<paragraph id="3C8CD8EDAB8B7BCC8139D38FE8CD97D2">
<emphasis id="8BC65AF55E679F2A11E87B70A58E6A1D" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, predominantly black anteriorly and red posteriorly; tergites smooth, setae with small sockets only; T 1 predominantly black, no red or a very thin line of red posteriorly, snow white setose, with scales; T 27 predominantly dark reddish brown, dark brown setose, white scales, white scales in lines on posterior margins of T 2 and T 4; S 18 predominantly light reddish-orange, yellow to white scales, thin black setose and long white setose; lateral margin black setae remaining close to abdomen, male T 8 brown scales, black setose.
</paragraph>
<paragraph id="85825F7E3D168362D20C12D774553C9D">
<emphasis id="0D8F171FB99AF76A32B46CA2BD8C8735" bold="true" italics="true">Terminalia</emphasis>
: gonocoxites longer than wide, broader at base, gradually narrowing, fused almost completely with phallus, with short, fine hairs apically; gonostylus with sharp projection; phallus inflated and helmet-like at base, apically separating from gonocoxite, continuing straight apically; lateral aedegal apodeme round with medial concavity; ejaculatory apodeme long, rounded in lateral view, extending significantly past anterior margin of the gonocoxites, ending at start of lateral strut. (Fig.
<figureCitation id="3917DE600E1290D47EE04FC7E82260BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral and ventral view of Enica male terminalia. Terminalia of Enica imitata NMSA - DIP- 90025 (A, C); Terminalia of Enica adelphe sp. nov. NMSA - DIP- 89520 (B, D). Abbreviations: b ph = base of phallus; ej apod = ejaculatory apodeme; goncx = gonocoxite; gonst = gonostylus; lat aed apod = lateral aedeagal apodeme; ph = phallus. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure4" httpUri="https://binary.pensoft.net/fig/1244177">4 B, D</figureCitation>
)
</paragraph>
<caption id="4A93C4FEB20428CDDE4E20C90BD71DA9" ID-DOI="10.3897/afrinvertebr.66.129611.figure4" ID-arpha="85EE03ED-89BC-5FAF-9BF0-0784CEA12BCC" httpUri="https://binary.pensoft.net/fig/1244177" startId="F4">
<paragraph id="5A6ED1738F1FB0A255B8062065074A6A">
<label id="2974B3577A7BEC312A4F43A4FDA56D3F">Figure 4.</label>
</paragraph>
<paragraph id="20C43D1D01A6317F9A3805473524208F">
Lateral and ventral view of
<taxonomicName id="13AA41B8BFFE1DDD5BBA5D9B6AB19B36" authorityName="Macquart" authorityYear="1834" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="3CDD2E7A9D5A103264EF88AE018C49C4" italics="true">Enica</emphasis>
</taxonomicName>
male terminalia. Terminalia of
<taxonomicName id="864FECB3C154CDF506B9D93D084093F3" authority="NMSA" authorityName="NMSA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="imitata">
<emphasis id="450D26E1F37BE74FCBDF4ECD3EC05884" italics="true">Enica imitata</emphasis>
<named-content id="0F94443AB872D57BE87B75FEE3B58C4D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
</taxonomicName>
- DIP- 90025 (
<emphasis id="F98E530E168A6A21A747355C2C53E770" bold="true">A, C</emphasis>
); Terminalia of
<taxonomicName id="07C6586116E26DA1B0E6C4A2802FD4A0" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="DCB86B5A27ECE5D770150DB80CFD1F1F" italics="true">Enica adelphe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B879E65836E702BC5F63B5A4B4DD7DCA" rank="species">sp. nov.</taxonomicNameLabel>
<named-content id="689C630E3F080A65E92396D0EAE83DA1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP- 89520 (
<emphasis id="01BDB1AA0A63305BBA53C07BD0369446" bold="true">B, D</emphasis>
). Abbreviations: b ph = base of phallus; ej apod = ejaculatory apodeme; goncx = gonocoxite; gonst = gonostylus; lat aed apod = lateral aedeagal apodeme; ph = phallus. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EB3AE" type="type locality">
<paragraph id="C29FF938C4B5D35A39ED11231E8197F1">
<heading id="6B515847B810C54F3DA87B3DBF491FE1" reason="title">Type Locality.</heading>
</paragraph>
<paragraph id="BC58227FD83A94A3481D377AB956FD7D">
<collectingCountry id="14A748F903692011393DD96E49DB45C2" name="South Africa">South Africa</collectingCountry>
:
<collectingRegion id="535A6EFA7D4BFB22906624D9AD0FB469" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
: Richtersveld, Grootderm
<quantity id="D25E06356424E1A8BD95F80DC7D21506" metricMagnitude="4" metricUnit="m" metricValue="4.2" unit="km" value="42.0">42 km</quantity>
NE (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK3AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[16.916667,-28.375000]}">
<geoCoordinate id="BB9F03BDF7313C0E6631F8B04C9E5F8B" degrees="28" direction="south" minutes="22" orientation="latitude" precision="15" seconds="30" value="-28.375">28 ° 22 ' 30 &quot; S</geoCoordinate>
,
<geoCoordinate id="EC122F579D263354BAC89E6E9068EAA9" degrees="016" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="16.916666">016 ° 55 ' 00 &quot; E</geoCoordinate>
</named-content>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP3AE" type="material">
<paragraph id="FCC4B81717519D246448A18EF3C5E945">
<heading id="830EA550CEC2630F49CE2A4912AB2138" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="C2AEF249B5FDAB3CBB8341D8E025B02D">
<materialsCitation id="5167E091F076F2B2FBF762DCB35F6330" collectingDate="1989-09-03" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Stuckenberg &amp; Brian" country="South Africa" elevation="200" latitude="-28.375" location="Grootderm" longLatPrecision="20" longitude="16.916666" municipality="Richtersveld" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="FF9764C692025094760E77A0049D81A3" bold="true">
<emphasis id="07F5CE2BC57A89267B7A4046107DF637" bold="true" italics="true">
<typeStatus id="B8B8E7137A7B6428382D59D70127A824" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="659C7A1F6F6F4B4EA06A0897ADD17243" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="CDFBD1A02FB4AD5C5BD73D56264B5142" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="020B09A15F2014509C49D74623E69D06" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="9E5339770076312507C2820EAF8CD61D">Richtersveld</collectingMunicipality>
,
<location id="56A5CE9CD3F875048167000971401EFA" LSID="urn:lsid:plazi:treatment:1FE42C3A1274556DB7D05F6D27DCDC85:56A5CE9CD3F875048167000971401EFA" country="South Africa" latitude="-28.375" longLatPrecision="20" longitude="16.916666" municipality="Richtersveld" name="Grootderm" stateProvince="Northern Cape">Grootderm</location>
<quantity id="47B613E4B4081BA9BD7FF0DFAED09534" metricMagnitude="4" metricUnit="m" metricValue="4.2" unit="km" value="42.0">42 km</quantity>
NE;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E43AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[16.916667,-28.375000]}">
<geoCoordinate id="ECCA6C204D746B42AD2D9B8E6C76277F" degrees="28" direction="south" minutes="22" orientation="latitude" precision="15" seconds="30" value="-28.375">28 ° 22 ' 30 &quot; S</geoCoordinate>
,
<geoCoordinate id="E96E5D00EDB55BEEA7E6E9265982698A" degrees="016" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="16.916666">016 ° 55 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="21AC9922C0ABB5A3D5FAFBBBA09FD12A" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<quantity id="1E342A4481EAAE98FCF9FBFF9FEC1017" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
a. s. l.
</elevation>
;
<date id="5BCDD2E56C85B7ABB8182BA74445EEC0" value="1989-09-03">
<collectingDate id="EEE5ACDCB91FAE0939C6C247EBE2D6BC" value="1989-09-03">03 Sep. 1989</collectingDate>
</date>
;
<collectorName id="7F5D1E5DA9A6F89C242E1FAD28D88F92">Londt</collectorName>
,
<collectorName id="8AB691563BB9CEC6CCB8E04A9E26106E">Jason</collectorName>
,
<collectorName id="E4B50E768A2E0380AF206A0E0E014263">Stuckenberg</collectorName>
,
<collectorName id="6781F3EEE38616D8B02314A0A67980A8">Brian</collectorName>
leg.; eroded sandy valley with flowers;
<named-content id="913CE255DE21400F4AAE6DE50E1FA108" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="C48FC2C37B4C4917B76F939CF7086EA7" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 89520,
<named-content id="5B40839493396B7D269859515E0B997C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="F545963B703792C7625558C048BFDD72" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
<paragraph id="A05514C75F7F731C1A7E9F66843072EF">
<materialsCitation id="C8019C0B8FF55E456BF0A0B991EF17CA" collectingDate="1986-09-09" collectionCode="SAMC" collectorName="Struck, M." country="South Africa" latitude="-28.283054" location="Richtersveld Numees" longLatPrecision="20" longitude="16.966667" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="587F4D3F6CE305009EAC828A7CEC782A" bold="true">
<emphasis id="40C5DDDE38B257AE1EF43620ED704CDC" bold="true" italics="true">
<typeStatus id="B89E7AC96721279968CC9FA4073CA824" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="B7F20E3D3EE0781BDC6E1D3283322314" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="CD66E1F08978BD8B10166188FD0A66E6" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="DA23E83AA05B960B9A6062308B2BF016" count="1" type="female">1 ♀</specimenCount>
<location id="E45D2B06B4BF378F1847B856C7E27F0F" LSID="urn:lsid:plazi:treatment:1FE42C3A1274556DB7D05F6D27DCDC85:E45D2B06B4BF378F1847B856C7E27F0F" country="South Africa" latitude="-28.283054" longLatPrecision="20" longitude="16.966667" name="Richtersveld Numees" stateProvince="Northern Cape">Richtersveld Numees</location>
(= Numees abandoned mine);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET4AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[16.966667,-28.283056]}">
<geoCoordinate id="79E4B017607CD9F61251213A96CA9BF8" degrees="28" direction="south" minutes="16" orientation="latitude" precision="15" seconds="59" value="-28.283054">28 ° 16 ' 59 &quot; S</geoCoordinate>
,
<geoCoordinate id="2DC1407CF9DDA0744DC3644EC04EB3DE" degrees="016" direction="east" minutes="58" orientation="longitude" precision="15" seconds="00" value="16.966667">016 ° 58 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="1AA1EE6D8FBB24614FFA2AEAC7A35294" value="1986-09-09">
<collectingDate id="C13DC0119D912087557407CA89796D65" value="1986-09-09">09 Sep. 1986</collectingDate>
</date>
;
<collectorName id="A67D11334F4AA763844F3FB32A7BE6A5">Struck, M.</collectorName>
leg.; SAM-DIP-A 004969,
<named-content id="6A130C09FD9625173DDCFD139E6FD33A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="455E2FB5D1CFBD895A2C116DF1F2DBF1" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="BCC2E19022149492AAF9637167D24B76" collectingDate="1989-09-03" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Stuckenberg &amp; Brian" country="South Africa" elevation="350" latitude="-28.316668" location="Grootderm" longLatPrecision="20" longitude="16.916666" municipality="Richtersveld" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="paratype">
<specimenCount id="C129DB60C126F69D44BA9370B428020D" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="C11E39834CC22399E0EFB2F50D34C80D">Richtersveld</collectingMunicipality>
,
<location id="E14AD3A5DF6392A4B85290DFB448844C" LSID="urn:lsid:plazi:treatment:1FE42C3A1274556DB7D05F6D27DCDC85:E14AD3A5DF6392A4B85290DFB448844C" country="South Africa" latitude="-28.316668" longLatPrecision="20" longitude="16.916666" municipality="Richtersveld" name="Grootderm" stateProvince="Northern Cape">Grootderm</location>
<quantity id="84B1321E85E157FF3B58C37E98120B25" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
NE;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E64AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[16.916667,-28.316667]}">
<geoCoordinate id="65DE2360101D25F12F2251CE5627683A" degrees="28" direction="south" minutes="19" orientation="latitude" precision="15" seconds="00" value="-28.316668">28 ° 19 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="E464F11863D72281974B23F695C2001A" degrees="016" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="16.916666">016 ° 55 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="E0A0EB73B41110A174A6832562F648DF" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity id="AD354F97F170799FD010D4B052BC36D7" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
a. s. l.
</elevation>
;
<date id="C6CDA4254D38B2EF8CAA68AA13FAE743" value="1989-09-03">
<collectingDate id="D6B43348CB8C8CA503D299E8496D3580" value="1989-09-03">03 Sep. 1989</collectingDate>
</date>
;
<collectorName id="C5373D8B12CBD658C7FC62575E296AF4">Londt</collectorName>
,
<collectorName id="B38E3470982E8E4F1CA5F68BDFAC0313">Jason</collectorName>
,
<collectorName id="6448D02781AAD2F9603E1CCFC92461D4">Stuckenberg</collectorName>
,
<collectorName id="8487712E4734F2181242E9ACA8BE8F4D">Brian</collectorName>
leg.; sandy valley below a rocky hillside;
<named-content id="CE4179865849F586DCE8F6E3282C17C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="9E5D6620B1994A332152794A38BAFCDF" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 89519,
<named-content id="72ABC72A39CD2B9EDEA6A40343B0363E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="0C233CFC4874DB46ED14245CCAE4907D" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL5AE" type="distribution">
<paragraph id="283F881974ACADC30776962DDF937533">
<heading id="5E2DAF57274166D45C5F989F2E6F3E05" reason="title">Distribution, Biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="635079FDDFF8DAA015825051F1457DF2">
Known from only two localities in the
<collectingRegion id="5AC7C446E63A7F40F4F27564A9D70476" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
in
<collectingCountry id="CE0BC1D306ABD9CDD14ED793ECD0112D" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="125732D1DA899422D2B2937100A99DDE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. imitata and E. adelphe sp. nov. specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure5" httpUri="https://binary.pensoft.net/fig/1244178">5</figureCitation>
). Although it is only known from
<collectingCountry id="5C86D476682501B2FE5E17D3659F8CD3" name="South Africa">South Africa</collectingCountry>
, it is likely that this species extends northward into
<collectingCountry id="74DD6B6F4303721C0B79DEF1939CBBEE" name="Namibia">Namibia</collectingCountry>
. It is only known from
<specimenCount id="E30EF2C7582EF31B00E08D867ED8D945" count="3" type="generic">3 specimens</specimenCount>
, between 2 collecting events in 1986 and 1989. This species is known to occur in the Succulent Karoo biodiversity hotspot. Adult flies have only been collected in September (Table
<tableCitation id="1023FC0FD474CA1B878B9332610F78CB" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="98FA702B322EAC9A9D7AD5E23399FECB" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="930A0D08980D4C44565DFB6FBED76D17" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="16075E363D7B60DF53F53778CB9C1E2A" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
<caption id="5E4B94B558D8B5200BC5567C1D820130" ID-Table-UUID="5E4B94B558D8B5200BC5567C1D820130" httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" startId="T2" targetIsTable="true">
<paragraph id="5F9571E9FA164BA91BB232B5BA76FD17">
<label id="FD78F8FD24EC2755FF912D81D95606F5">Table 2.</label>
</paragraph>
<paragraph id="0F44A7118A513352CFD190FED8839684">
Seasonal imago flight activity for
<taxonomicName id="54AE38C8E6A88AF121D4CA1A2A34D5E2">
<emphasis id="02429897C69B37F64BE6D5ED1AEFA7A5" italics="true">Enica</emphasis>
species
</taxonomicName>
through number of specimens collected for each month.
</paragraph>
</caption>
<paragraph id="7E5F45CC908E7D7F89A43E51B8176026">
<table id="TID0EBFAI" rules="all">
<tbody id="199218984D48A54141E66FE90B98E0B1">
<tr id="28645C10079513CB89527B59F10B2611">
<th id="15DC6995FF91E61B5F2ECA4850DA7320" colspan="1" rowspan="1">Species</th>
<th id="A6C4964D9645D5A91DD910DED6DE18DB" colspan="1" rowspan="1">Jan</th>
<th id="0B0B3D68FDF1A58BF72F9AD78B76221A" colspan="1" rowspan="1">Feb</th>
<th id="5D5AC1ADDE1643A4A92E005727451F0B" colspan="1" rowspan="1">Mar</th>
<th id="2621C5F513FC65AB2CE5F6CA8F4D2F89" colspan="1" rowspan="1">Apr</th>
<th id="8A87DCA5842931D253521415163D22BE" colspan="1" rowspan="1">May</th>
<th id="1B4449F4ED693340C0E4AE069E3847FA" colspan="1" rowspan="1">Jun</th>
<th id="A6D409A6D8AF1535BA9C3542FCB7F0F9" colspan="1" rowspan="1">Jul</th>
<th id="393453C3FC4DC68107DBF6F3F0D367A8" colspan="1" rowspan="1">Aug</th>
<th id="FB355465A0BD8D5250A0CE9829C1F4E0" colspan="1" rowspan="1">Sep</th>
<th id="D0261C940C5E8B2FCE334C0978445D13" colspan="1" rowspan="1">Oct</th>
<th id="331B94A5B0828830988E79F9658B1496" colspan="1" rowspan="1">Nov</th>
<th id="3CD9E055578CDBF94B121F57AB73F914" colspan="1" rowspan="1">Dec</th>
</tr>
<tr id="95F7ED70A12F8B583B756FBF73331A2A">
<td id="7B86799847BCDEA992CAA1BB6E7545A7" colspan="1" rowspan="1">
<taxonomicName id="56E58155988D8B0A396B0DBAB8F8A3CC" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="3AF5E96210F3F596D83656102AE07E21" italics="true">E. adelphe</emphasis>
</taxonomicName>
<emphasis id="EF0079FDA320407978297461A2B861E7" bold="true">
<taxonomicNameLabel id="7830E5BA79768A559766920D8C25660F" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
<td id="306FF42313164552985905EBEE0DBD8B" colspan="1" rowspan="1">-</td>
<td id="750758FA618120896D2B6B9507B72C93" colspan="1" rowspan="1">-</td>
<td id="F29323F213245F053234A240734690EA" colspan="1" rowspan="1">-</td>
<td id="8A2DBC46733900972C458FCFE680D5F0" colspan="1" rowspan="1">-</td>
<td id="A7A98E0D5745BAE6F46ECF869FBC3087" colspan="1" rowspan="1">-</td>
<td id="063DACD109F6B349D101B6AE34ECC230" colspan="1" rowspan="1">-</td>
<td id="66336537BB9237DCCBB21B5EC38CDA0C" colspan="1" rowspan="1">-</td>
<td id="E5113744AC4EE3E117BC89D9E8834E8F" colspan="1" rowspan="1">-</td>
<td id="1181A4C2387861764D6955EED9BA2A17" colspan="1" rowspan="1">
<emphasis id="94BE89B5132DF1D5BDA4C22F5962572D" bold="true">3</emphasis>
</td>
<td id="2CBE8A5D7D49E61C912B0B00A62F0101" colspan="1" rowspan="1">-</td>
<td id="D8411B5FA006E4AAD9CBDCCDD145FCF9" colspan="1" rowspan="1">-</td>
<td id="C747BAB1C9F0DF1CF4C5FD0B7806ACCB" colspan="1" rowspan="1">-</td>
</tr>
<tr id="77AFEE1124F7C9535F248E29365AB258">
<td id="4167A56E19AD638943D4DCE7B73A243E" colspan="1" rowspan="1">
<taxonomicName id="79DA6D31C7D13DA1F13F0651737BA429" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="clavicornis">
<emphasis id="83EA8A18DEBDA2C1F76A6BD75ECA57A6" italics="true">E. clavicornis</emphasis>
</taxonomicName>
</td>
<td id="853EEC8DA944F8FF1A185C68474B68B6" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="128E3928EADA69BE39EA772732F1E668" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="5FD8726983A3E2E2F895B943DF0D3FAC" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="A8234219A17B7E7AC31B0911BB153045" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="BA79D4622D0EA330E52C3C0375A1D982" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4D36D34BED4ED791ACB07387F6597BA0" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="850514237EF3731888F8540AA7BB5651" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="B70E24A1D76C4A8F62B155491ECDA694" colspan="1" isEmpty="true" rowspan="1">-</td>
<emphasis id="2CEB38EF90FF09718E407489D8A40FF1" bold="true">
<td id="5A9D4D20DA0127475570E426E89C8D44" colspan="1" rowspan="1">2</td>
<td id="40E792A8FB13E104E3943DB58151C600" colspan="1" rowspan="1">1</td>
<td id="741160BE2BECC39C5FF1B9EF10AFBB2B" colspan="1" rowspan="1">8</td>
</emphasis>
<td id="86C3B844F0691572597895B5B03A8FD3" colspan="1" rowspan="1">-</td>
</tr>
<tr id="E7FABB05546934039EEE026395D045C4">
<td id="9D8CA9AD1347A66CE1047642CBE2FC42" colspan="1" rowspan="1">
<taxonomicName id="4531CE514F55A0C4779A9E42AA96E13B" authorityName="Hesse" authorityYear="1975" class="Insecta" family="Bombyliidae" genus="Nomalonia" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="eremophila">
<emphasis id="E908F4BFAFE622AD40FBDE7292067ED0" italics="true">E. eremophila</emphasis>
</taxonomicName>
</td>
<emphasis id="CDF88DDD6199F73718A7B36538C338F2" bold="true">
<td id="464AA625D009E855F3BFD8F1CFB5E604" colspan="1" rowspan="1">3</td>
<td id="24C97A482F2AC5430BED91ED12C86BB9" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="49CE7492AD4B424BF00A123601F54136" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="BECA75FB9C519ED1328C41DE2DA3650A" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="95E8C845C165EABC60502E8EACB3C44B" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="B246439A7B6B44BCC7D1E7EF8BDC91EB" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="5760522708C7A4789FDE45591CAB91CF" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="425F812AFABE17EF957A74D713687F58" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="9FB1956E5BF6CA86E540E136FCBD0534" colspan="1" rowspan="1">3</td>
<td id="839477075CE3A69F7D8B83F242D9A541" colspan="1" rowspan="1">19</td>
<td id="64278B297E0542AE54984A36154A643B" colspan="1" rowspan="1">40</td>
<td id="F15E846682C01BFFF1272355B6ADE569" colspan="1" rowspan="1">11</td>
</emphasis>
</tr>
<tr id="47037AAF806F1431C92FA566A3EE688F">
<td id="AD1E197EEC03DD07CE58591BF8345386" colspan="1" rowspan="1">
<taxonomicName id="1BDCE71DBE1C450A2A73A36E77437586" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="henicoides">
<emphasis id="4C843ECFF7B28A3E8769E5D21BD2FA46" italics="true">E. henicoides</emphasis>
</taxonomicName>
</td>
<td id="FD38E1C93ACE4060F702483D969F56AD" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4E33A080E73AD9582EB1A4B25FCBCCD0" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="B87C9213290D91D2A3B73AD48B66BFFE" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="D0A58C877FE03610454C0AAE2D03ECE4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="417D936F9B4F98A6EC2B0860DA6E2340" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="2601C5CE96CB07BFEF54C4E959DC782E" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4FD6B89E01DF009308BE00297C11BFF6" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="65237D21416F52E3FDCD9CD380649148" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="30FDF711CD0692E619A60A7DDACDEAAC" colspan="1" isEmpty="true" rowspan="1">-</td>
<emphasis id="06B1C7B57EB4FA3DC5D737EC3AD97650" bold="true">
<td id="1D33EBCC22605DCC332BF38C5009A314" colspan="1" rowspan="1">2</td>
<td id="8615F805FF348D00CE1EACCC7E0B3371" colspan="1" rowspan="1">3</td>
<td id="8ABA4D2F6DC382285A5BE74795C601A6" colspan="1" rowspan="1">1</td>
</emphasis>
</tr>
<tr id="9F1F3B0A1EEA9730CCF5FDA96E7F4AD8">
<td id="3B672CD4AC0347BA2D175F42FB2826B2" colspan="1" rowspan="1">
<taxonomicName id="2A21723147D0A0162B2682B77295F4EA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="imitata">
<emphasis id="5A6F65FC71BA18EB8A191CE7A38E07F6" italics="true">E. imitata</emphasis>
</taxonomicName>
</td>
<td id="162F717EEDEA57C8BCCF0174D43574F0" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="5303684A3AF6265D022A6353D92B02C7" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="25983F774180F9B929042E9D8D31B9E4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="9853D31881E16517D09369B8968E46D3" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="21D0EBC4866E281ED65872D31900B6A1" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="FFF50800C9B33BB3F2AC3FD075168B6A" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="56ADED44C980C5BB5E5782C9200C0882" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="A79E67FE661F3785B46DC312434CFE3E" colspan="1" isEmpty="true" rowspan="1">-</td>
<emphasis id="8F242785EEC1F02D648EA872ADC6C57F" bold="true">
<td id="DEA086D09B7CD23C510A051E9E8B902F" colspan="1" rowspan="1">1</td>
<td id="1F1D0575E0208BC7A68D96AD82398D50" colspan="1" rowspan="1">2</td>
<td id="A2DE041F44D202B26249F2EEBBEFA543" colspan="1" rowspan="1">2</td>
</emphasis>
<td id="95E013F7686C0DA224816B55D3B86E37" colspan="1" rowspan="1">-</td>
</tr>
<tr id="3BB4992D0FD283E113CFD1BCAEEFC60D">
<td id="A9B2913DC87A1CBBAA55593643B4069D" colspan="1" rowspan="1">
<taxonomicName id="DC1316C3D71EE680F2F80E299A3D82B8" authorityName="Wiedemann" authorityYear="1819" class="Insecta" family="Bombyliidae" genus="Anthrax" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="AACEBA183FAC01760DDCCCD2937DBE98" italics="true">E. longirostris</emphasis>
</taxonomicName>
</td>
<td id="5BF73A67EDBD907B8622A9370B53ABEA" colspan="1" rowspan="1">
<emphasis id="484BB809507A02C13AD819FA8CFFD03A" bold="true">6</emphasis>
</td>
<td id="CFEBA53F5FF8F4C6029D1F5FB7ED8E79" colspan="1" rowspan="1">-</td>
<td id="1E65F3A8336472844CD12B99A946F48B" colspan="1" rowspan="1">-</td>
<td id="0650FF7409BB589CE009AADC65D38501" colspan="1" rowspan="1">-</td>
<td id="1540C61383AC8A96306EE8FF32274283" colspan="1" rowspan="1">-</td>
<td id="68D4413FDE12D8F7FE2D3184A4F2CBA0" colspan="1" rowspan="1">-</td>
<td id="7EC191E9AFDF18591BFEDD6A3564C863" colspan="1" rowspan="1">-</td>
<emphasis id="8FC352BEB7C0BDC38A1CEB7614C454FA" bold="true">
<td id="C599F35B272289C5F1328071D0FA897A" colspan="1" rowspan="1">1</td>
<td id="0317F105CAB864B09D97B12923DDD7A0" colspan="1" rowspan="1">11</td>
<td id="8ECD04D240ABCEA8B442A7C34C1C4256" colspan="1" rowspan="1">86</td>
<td id="F5A3DE06F4EA85819458DBB364EE0A90" colspan="1" rowspan="1">39</td>
<td id="58629446B29DBA573DCC7DFFEAAC0875" colspan="1" rowspan="1">73</td>
</emphasis>
</tr>
<tr id="AD82DE1C986B5D946908C6EC52284F3F">
<td id="2C4A8A997CEB9591CEA6F9473DF7B2E8" colspan="1" rowspan="1">
<taxonomicName id="9F4EACDCDA9A11FDB3BB27D9B079DADD" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sporanthera">
<emphasis id="EAB27BF8F3B206E24859546E1902C9EB" italics="true">E. sporanthera</emphasis>
</taxonomicName>
</td>
<td id="A9B9B11001557FE05C4E1EA3074DC0F9" colspan="1" rowspan="1">-</td>
<td id="46DFC6FB611CA6F6D44A9316B98CC516" colspan="1" rowspan="1">-</td>
<td id="55C0BBDE4A66D785A77C8ED0A98BFEC4" colspan="1" rowspan="1">-</td>
<td id="364FD60AF89BB5F72CB0381331E0E5E2" colspan="1" rowspan="1">-</td>
<td id="22855578E93E9E3DC2FEE4904B8CF886" colspan="1" rowspan="1">-</td>
<td id="B3AD31FE204F83AB45CC707EDD26C00F" colspan="1" rowspan="1">-</td>
<td id="0B9FCCF0B336A98C2E7F9EAEF9197CD0" colspan="1" rowspan="1">-</td>
<td id="5582BD70DB6BFCEB54D86FB23148FF47" colspan="1" rowspan="1">-</td>
<td id="9F4369CFB85B228E06856005F284C403" colspan="1" rowspan="1">
<emphasis id="59FBF18762CB3108E89C633E89C583EF" bold="true">1</emphasis>
</td>
<td id="94C64203682CC8934BFF9DF943365072" colspan="1" rowspan="1">-</td>
<td id="A57EBD2AAC017EAEB33167658593DBC3" colspan="1" rowspan="1">
<emphasis id="835FEB7BDACB93E2A11D9D583C8F1D62" bold="true">3</emphasis>
</td>
<td id="9FCC0279C521D450021DBF65F9C190FF" colspan="1" rowspan="1">-</td>
</tr>
<tr id="D4B144B090C4C5435634CF9400AFA9B8">
<td id="A963AE1F51DAC5CFA66BAEA95D981B61" colspan="1" rowspan="1">
<taxonomicName id="26BC9C4E4562573AA7717486A68527A2" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="syrticola">
<emphasis id="A7C2766919BE8AA69BAC3B2D6B3660AC" italics="true">E. syrticola</emphasis>
</taxonomicName>
</td>
<td id="64220B6DBD9353629943E7372C09CFF6" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="FB101A26505FB410296BFBD996E99AC6" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="7FAEDB2F581CBBCA02DD2B2E939DACA4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="78AF4A9707F82FB7D09D588687509BF7" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="1A12FE8766E463CE5453EE5C83D72CC5" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="03C71667232E536BD10963A9A0369B6A" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="7017AC04CBB36412F711532268C3E20A" colspan="1" isEmpty="true" rowspan="1">-</td>
<emphasis id="8403D20B4204AA4405E7031E600EF92C" bold="true">
<td id="2333871C2911005ABA331E7C6DEE10F1" colspan="1" rowspan="1">1</td>
<td id="30BF499FFCC39EB35FDA6AE18C31926B" colspan="1" rowspan="1">5</td>
<td id="01AA3267002BD84EA6C49C3A3CA94F36" colspan="1" rowspan="1">22</td>
<td id="4F6C7904D810CD4974017B24916D30CB" colspan="1" rowspan="1">2</td>
</emphasis>
<td id="3BF456750691B82F1B1AC7EA40A02561" colspan="1" rowspan="1">-</td>
</tr>
</tbody>
</table>
</paragraph>
<caption id="C104972F5CF313B142C6D362922D7C72" ID-DOI="10.3897/afrinvertebr.66.129611.figure5" ID-arpha="DBAF4E44-F079-51CD-8CF6-FDE946821688" httpUri="https://binary.pensoft.net/fig/1244178" startId="F5">
<paragraph id="950D06609284BDC3ED1233BD75983304">
<label id="A0EEC8EFBBF367DF0CBF966B536DEF37">Figure 5.</label>
</paragraph>
<paragraph id="509AB11B3B01640E3E696D0F893ADDB5">
Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of
<taxonomicName id="2495734E51B2E1391CC6467E9CC1D93D" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="imitata">
<emphasis id="182FBA37BB2C9AA0BF2D6F28E02EDDE1" italics="true">E. imitata</emphasis>
</taxonomicName>
and
<taxonomicName id="5EF75654B67A35BC0659A876DC9874E5" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="adelphe" status="sp. nov.">
<emphasis id="14DB8CA3BD91F0A1070EF2F1DECD0524" italics="true">E. adelphe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="077EC25028CC216CB880C167C836C53C" rank="species">sp. nov.</taxonomicNameLabel>
specimens studied.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,576 @@
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<mods:title id="ABBAE0A44A51AD6307BFDB9BECC08685">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
</mods:titleInfo>
<mods:name id="4ACB3A730714CEE3CA54CA1A1E09071D" type="personal">
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<mods:namePart id="F82199B066C02BDB2BF17390CBB589F5">Rollinson, Lisa</mods:namePart>
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<mods:affiliation id="30E23B737D82A0FEF0CA8F1EFF9139E2">Texas A &amp; M University, College Station, Texas, USA</mods:affiliation>
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<mods:namePart id="4F00F2724D1FE2D6CE4436F986FFA710">Cabrero, Allan</mods:namePart>
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<mods:title id="077D9F1B604C85FEC39B53E00A091C0D">African Invertebrates</mods:title>
</mods:titleInfo>
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<mods:date id="F14F0058661EAEC9987B772918E5C8F4">2025</mods:date>
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<subSubSection id="3739E2EE89B409CAC7B5AA93F59C36E5" type="nomenclature">
<paragraph id="B1117DF3F15F386728AE14EC001532C0">
<taxonomicName id="D9F7AC10F29E5C238414341A88D6DBBF" authority="(Hesse, 1956)" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="clavicornis" status="comb. nov.">
<emphasis id="AE839E570DA2BA10AB723B051067328C" italics="true">Enica clavicornis</emphasis>
(
<bibRefCitation id="DAECAA1BB920D7BCC464D91213588625" author="Hesse AJ" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 27-46. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="5CBF36FC116B3291451EA5B89A044BCE" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4C9ACFDCE9C9B892201293AB99783AB7" type="description">
<paragraph id="BBDC49505E99CEAF31F0D8E0C6D03F26">
<figureCitation id="4C5336B09B5BA959D945C19F444DC756" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Enica clavicornis ♂ NMSA - DIP- 93558 (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica eremophila ♂ NMSA - DIP- 93041 (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure6" httpUri="https://binary.pensoft.net/fig/1244179">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDTAG" type="Taxon depository">
<paragraph id="7772E079B59B21D55E8810A9338A7EC2">
<heading id="E130E867A98F069D175B60862CD6960F" reason="title">Taxon depository.</heading>
</paragraph>
<paragraph id="95244F477E52C79626C809E752E23B2A">
ZooBank:
<ext-link id="9A41BA4BDB1FC3161B79DA35DEB985CA" ext-link-type="uri" xlink_href="https://zoobank.org/A4E281A4-196C-46DE-83F0-93D4E9CF0777" xlink_type="simple">
https://zoobank.org/
<uuid id="520688465FCBF7FD2DFD39F6618E9974">A4E281A4-196C-46DE-83F0-93D4E9CF0777</uuid>
</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOTAG" type="diagnosis">
<paragraph id="7C8B271CF5F10AAC615588D99C199507">
<heading id="1C4C411342E49BE73ACBBE9296BE8FE2" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="32A1810B0EA5A39CBD02A3E8E489D6A5">The species is distinguished from other species in the genus by the light creamy tan flattened setae on the haltere and the club-shaped postpedicel.</paragraph>
<caption id="12B223F00A7A18DD8481CE346F3CBA92" ID-DOI="10.3897/afrinvertebr.66.129611.figure6" ID-arpha="6341EB98-8B3F-59B5-AA4B-47F7B10620BE" httpUri="https://binary.pensoft.net/fig/1244179" startId="F6">
<paragraph id="CFEEA1253FA71C1BF3A65AD0672F14BD">
<label id="C9EA46CE5BFD3C334EC67A8E73CC0A17">Figure 6.</label>
</paragraph>
<paragraph id="6816A7768ED03367414187A067B4D25F">
<taxonomicName id="CB04C45E926C1B443CEFE1AEC1ABD04A" authority="♂ NMSA" authorityName="NMSA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="clavicornis">
<emphasis id="69CE3995A5A1C83B56EB4867CB6BDB3F" italics="true">Enica clavicornis</emphasis>
<named-content id="B336D021CF61B3EDF6DA4EB2585F9313" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
</taxonomicName>
- DIP- 93558 (
<emphasis id="AD0B3FA649F641B2F27001D78A2BE822" bold="true">A C</emphasis>
)
<emphasis id="E014981F1FA5D915861004E1C5E7CE25" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="932565A295B7F55A7B1A56524BD2DF3D" bold="true">B</emphasis>
habitus, lateral view
<emphasis id="8A5BCF04D2841A0F8D8D38C5AF10993A" bold="true">C</emphasis>
habitus, frontal view;
<taxonomicName id="4C8787D47127436C45D5F12863C78599" authority="♂ NMSA" authorityName="NMSA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="eremophila">
<emphasis id="26AB973DB9162B7905092DC16428F999" italics="true">Enica eremophila</emphasis>
<named-content id="7A3C7D7FDFCDEC184CBC300D08D1243B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
</taxonomicName>
- DIP- 93041 (
<emphasis id="94C90BDD52B3D88EFE6FB14B2FA9EAFE" bold="true">D F</emphasis>
)
<emphasis id="C6CDB6329575C86CDDA567E50EA22F55" bold="true">D</emphasis>
habitus, frontal view
<emphasis id="43E460DC5D2F1BDDA1808CA68EEE79D6" bold="true">E</emphasis>
habitus, dorsal view
<emphasis id="6A8DE716365C83405593B49435CB2500" bold="true">F</emphasis>
habitus, lateral view. Scale bars: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOVAG" type="description">
<paragraph id="B4EE83B8A5C4133505C9307F6AB26D5F">
<heading id="164D2AE9F22B333494FC6E83ECA98919" reason="title">Description.</heading>
</paragraph>
<paragraph id="1638823C45E079EEB303490A448672A9">
<emphasis id="71F39581DEC896D44FF2D0B630BFBB76" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, without dark spot, males: similar to head, slightly darker than head; frons black setose, with white scales, broadly setose near eyes, extending ventral to antennae; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly or black and light yellow macrosetose, white or light brown scales posteriorly; occiput whitish yellow setose, brown setose on occiput spots, white scales antero-dorsally, dorsal eye margin with dark brown to black spots not more than three times as long as wide, absent in males; gena creamy white, whitish yellow setose.
</paragraph>
<paragraph id="347744F64A9521E2E80A6D5895C1AD64">
<emphasis id="AE88EEB131E8BAF1DDC08C860B50C7FB" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, light brown to brown, light yellowish-brown setose.
</paragraph>
<paragraph id="08C543FC58E6A124F45B8356CC96D4A8">
<emphasis id="64F0B28E816CC9AD65176EDBD4FE0078" bold="true" italics="true">Antenna</emphasis>
: dark brown; scape creamy white, approximately as long as pedicel, white and black setose laterally and dorsally, setae short, not extending to end of pedicel; pedicel creamy white, short black setose dorsally; postpedicel dark brown, proximal bulb with elongated distal part, longer than scape and pedicel combined, white or light yellow scales dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="7F81BF4992C588C568783B672AC0A951">
<emphasis id="569F7751997B7E9CCD246D59E3794628" bold="true" italics="true">Thorax</emphasis>
: light orangish brown; scutum medially black, red around margins, red sometimes extends in medial line from behind head to center of thorax, yellowish-white to white scales with two submedial longitudinal stripes of white scales; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, light yellow scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, long white flattened setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, long white scales ventrally, black macrosetose and white setose dorsally; anepimeron silvery grey pubescence, long white scales, white setose; katepisternum silvery grey pubescence, long white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, long white scales dorsally; metepimeron silvery grey pubescence, asetose; scutellum light orangish red, slight anterior black spot, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of light brown and white scales, black setose, white scales, black setose or yellowish-white scales.
</paragraph>
<paragraph id="239FCE21E004F5394D6B75059A22BDC7">
<emphasis id="D091E2A7F9D475E7E3136BB5D77F0110" bold="true" italics="true">Leg</emphasis>
: light orangish brown, covered in scales, some regular setae present; pro coxa light orangish brown, silvery grey pubescence, white scales, thin white setose, thick black setose; pro femur light orangish brown, white scales proximally, brown scales distally, thick black setose ventrally; pro tibia light brown, raised short and fine light brown setose ventrally, large black setose dorsally; mes coxa light orangish brown, white pubescence, white scales, thin white setose, thick black setose; mes femur light orangish brown, white scales proximally, brown scales distally, thick black setose ventrally; mes tibia light brown, light brown scales, long black setose, distal tip with long black setae; met coxa light orangish brown, white pubescent, white scales, thin white setose, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur light orangish brown, white scales proximally, brown scales distally, long and thick black setose ventrally; met tibia light brown, light brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with brown scales dorsally, black setose ventrally; mes tarsomere with brown scales dorsally, black setose ventrally; met tarsomere with brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="E25ACF08A23AC0658BA2FE11644E9542">
<emphasis id="A6A50C9A08D1F6E62F4F11E947D0C563" bold="true" italics="true">Wing</emphasis>
:
<quantity id="2723DD39B748695849A0F9BF5E0BB939" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.1" metricValueMin="9.1" unit="mm" value="9.6" valueMax="10.1" valueMin="9.1">9.110.1 mm</quantity>
, microtrichia absent; membrane color pattern males: minimal spots of infuscation in r 2
<subScript id="10CEE6583E9573D069598ACC457E9D25">+ 3</subScript>
, br, and bm, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="04BD69AAE3FBE802405A7F01E4026CC3">+ 3</subScript>
, base of m
<subScript id="78663111CA5D173422A18D994FF09BD1">2</subScript>
, m-cu, split between M
<subScript id="4CD9B7F4A6C2C248F35A0E5435857595">1</subScript>
+ 2 and M
<subScript id="B492548B8A03866837299B55BE5AA633">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="DFFC2E66173D46F6C5958828CD1B53EB">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="3ADA522F8F856C4AB5BD192EE7C0EB7B">1</subScript>
open; R
<subScript id="C2E8CDCBA457B4357AD72BB49188DAA2">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="150BFC36433F857192AC92FCA37310FA">4</subScript>
open, proximally R
<subScript id="D3020712744A170FC7F132095E163094">4</subScript>
and R
<subScript id="8E83D17EC58EB27CF5BBB7AAB4CBC2E3">5</subScript>
parallel, R
<subScript id="19833F70390C9B112DA92D622D426622">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="E582EAA2CDB3166DA021A744321477CE">5</subScript>
terminating posterior to wing apex; r
<subScript id="C7B0B32B5A6B502C2FEC8D97809A7C46">5</subScript>
open; M
<subScript id="DA309B7FE21F922895ACADE14D5CC9F9">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="A744E64779F02DF9904D5A20880CF82A">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="68F9E0E9D4870B624F08EEF5D88D2ED1">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem light yellowish-brown setose.
</paragraph>
<paragraph id="0206A14588AE51C958D7E367C366F4A4">
<emphasis id="BC9659BC3C4B23FF1F0CEA4747E6DC35" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, reddish-orange with a black spot on T 2, middle of T 3; tergites smooth, setae with small sockets only; T 1 reddish yellow to light brown, light yellowish-brown to light whitish-brown setose, with scales; T 2 T 7 predominantly reddish-orange, thin black setose, brown to light yellowish-brown scales, scales uniformly colored throughout; S 18 predominantly light reddish-orange, yellow to white scales, thin black setose and long white setose; lateral margin black setae remaining close to abdomen, male T 8 dull, light yellow scales, black setose.
</paragraph>
<paragraph id="9ED612D35DDDEB2EC395DE50E8F04F5D">
<emphasis id="70DA2D46C567BCC16E1BF55A5D2C410F" bold="true" italics="true">Terminalia</emphasis>
: Male terminalia drawings in
<bibRefCitation id="7C21C6A39375B60C6C4F782ED382060C" author="Hesse" firstAuthor="Hesse" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 2746. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse (1956</bibRefCitation>
, p 45):
<ext-link id="C249425346E8494932A7490C6C52A810" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/40844180" xlink_type="simple">https://www.biodiversitylibrary.org/page/40844180</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVYAG" type="material">
<paragraph id="B3C6A18BB1BA93A8F8D7B58251B4D880">
<heading id="AF8F6EED91856FECF3E7C91930343E46" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="B20A3594B5B344C0D39567F76A295E0D">
<materialsCitation id="2E22D0BB3A8FC273348F20BEB418B46B" collectingDate="1933-11-11" collectionCode="NMSA" collectorName="van Son, G." country="South Africa" latitude="-30.313055" location="Hondeklip-Baai" longLatPrecision="20" longitude="17.275" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="FC8079F43B8BAAE5D5B47EC1D3E9590B" bold="true">
<emphasis id="A75462EA1D0BED41BFA6BB898128231E" bold="true" italics="true">
<typeStatus id="D758C4DDF2FA07144AA0F60BD679D569" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="9CB6A12B74DF7AE6B462BFFD83439D8A" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="EA8409D7E07A8212EF00D40ED00A15D6" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="6CC18281A234C4BB51714459022F81EB" count="1" type="male">1 ♂</specimenCount>
<location id="3F54D8675343FB633688A94D5BC97BBB" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:3F54D8675343FB633688A94D5BC97BBB" country="South Africa" latitude="-30.313055" longLatPrecision="20" longitude="17.275" name="Hondeklip-Baai" stateProvince="Northern Cape">Hondeklip-Baai</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDZAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.275000,-30.313056]}">
<geoCoordinate id="69CAB6CD751D83B2941448AD74546B1D" degrees="30" direction="south" minutes="18" orientation="latitude" precision="15" seconds="47" value="-30.313055">30 ° 18 ' 47 &quot; S</geoCoordinate>
,
<geoCoordinate id="3215715EFE21A672AC8110C8FCDEA0EE" degrees="017" direction="east" minutes="16" orientation="longitude" precision="15" seconds="30" value="17.275">017 ° 16 ' 30 &quot; E</geoCoordinate>
</named-content>
;
<date id="D3D9B56A45CA8E49A7B9466993CF9DBD" value="1933-11-11">
<collectingDate id="687641130249653F05B79E3F82068760" value="1933-11-11">11 Nov. 1933</collectingDate>
</date>
;
<collectorName id="AC0BB258A651D324091199A353343766">van Son, G.</collectorName>
leg.;
<named-content id="8A5F7D75FA0F64E617E55F89A9ACA9EC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="080960FD2BB85D6D73B4A261F2169E8F" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 051430,
<named-content id="CB863CD1A73B2EF4A480E10BD8380CAB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="8B32882077E90E3A107C258CD8B42AB0" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
<paragraph id="E279D84C7AE0728AA85D535D37D5CF4F">
<materialsCitation id="201ECBAB123A4CA907239E8ABB55591E" collectingDate="1950-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-30.388332" location="Wallekraal" longLatPrecision="20" longitude="17.508888" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="CD9E4935900BE791C11ECA97E08A3A22" bold="true">
<emphasis id="AB599388E4DBE1A86C3BC9960C79C0E8" bold="true" italics="true">
<typeStatus id="84BC5F6F4A46F99E0D957410823A04F6" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="DA1E09BE30EBCDCBC6AC1948057C32DE" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="126435DBB5C416C7E61B3BBC9BC49C24" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="1BFCAFCB3D9A4DCECE2BAB9EA7A4FD6C" count="1" type="male">1 ♂</specimenCount>
<location id="E1A7FF48B0539250552E5A96934B6BE3" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:E1A7FF48B0539250552E5A96934B6BE3" country="South Africa" latitude="-30.388332" longLatPrecision="20" longitude="17.508888" name="Wallekraal" stateProvince="Northern Cape">Wallekraal</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZZAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.508889,-30.388333]}">
<geoCoordinate id="A16B62C59693C1319D9561C028D5396C" degrees="30" direction="south" minutes="23" orientation="latitude" precision="15" seconds="18" value="-30.388332">30 ° 23 ' 18 &quot; S</geoCoordinate>
,
<geoCoordinate id="03ACE4332409A3B63B5138AFEACFA6B5" degrees="017" direction="east" minutes="30" orientation="longitude" precision="15" seconds="32" value="17.508888">017 ° 30 ' 32 &quot; E</geoCoordinate>
</named-content>
;
<date id="210689ECFFC9E8910032D07E2E684E59" value="1950-10">
<collectingDate id="DDB4B634AAAD1EC589C3F46C321C3ECD" value="1950-10">Oct. 1950</collectingDate>
</date>
; SAM
<collectorName id="B626C4D443C1B3EE146BAF5D8578FBD2">Museum Staff</collectorName>
leg.; SAM-DIP-A 002151,
<named-content id="4D7F3A2E74A4B23439E3D4F6F340A930" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="5D56FCF34C128F445A131DB95E5270A5" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
<paragraph id="32EB35276AE03944CA02B4ABEA102B84">
<materialsCitation id="AF5B78C366A876D66F64FBFC86F5F89F" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<collectingCountry id="899B39E22D3594726A203FE3CE07CB20" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="76E98635164C1A2E307AD62D21FDBC3C" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="8A1A99901A750BDCADED4F686FE0EF68" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="6755E3D5D0CB0CB6AF1E5E82D82885F1">Elands Bay</collectingMunicipality>
,
<location id="C683F12B783D87059B6DCEBB54F2034F" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:C683F12B783D87059B6DCEBB54F2034F" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EG1AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="FEA3620682B1CF26649CB1ABA25FFC4D" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="495601B052E7AC8E90D8687F224F5526" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="C16D90D786E81F7D5ED45E667D8E87DD" value="1948-11">
<collectingDate id="1BA4B3C2FBD674FBE5300403CEBDD4F7" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="EAE46B272D7624C1651DB6330519D87B">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="F962AF10A7D4C9BC909D338A5C98CFC6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="0B8BBD7D66F8783429A719EF71A3EBE3" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="1D84F3D95EA9F2A9D4F3D79EDD77FF2D" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="B2B7B1F65B774383F7D6B4A0C0D415CF" count="1" type="female">1 ♀</specimenCount>
<collectingMunicipality id="F09BFAD840C4AD0953DC7168323869A5">Elands Bay</collectingMunicipality>
,
<location id="CB13B4B05EDB4E5B5BD81659640896CF" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:CB13B4B05EDB4E5B5BD81659640896CF" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ES1AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="932E487831DC5B147C53EC020E0B26A6" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="DB248DD179EB9F7BCD7B7D9BD0627954" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="29357E532D68F8DF48A193F1DC0A32EC" value="1948-11">
<collectingDate id="B33196DB1A0B2330D8C945ADD8A97C3D" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="27EEEDCACEFCFDC79523B99E63A0A765">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="D0DFE39B446234B7CBBBC2630218621F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E3FF0CCA992E8247EE1B193EBA465A2E" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="19CBB145A4015C198732BF509BC36AEC" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="B472BB00A61D8B4189379AB2E071E09B" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="F2959652E77E52E448E46463E9B9B8C2">Elands Bay</collectingMunicipality>
,
<location id="ABBC5F373093D2E71DA482A12AC79CD4" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:ABBC5F373093D2E71DA482A12AC79CD4" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E51AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="D0A743DD6CDA69EB2DEB2596A0F320EB" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="F1AFA777C4D6E3BD978A8FA75CCA215A" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="BBB548D3B9CC6B4DA9FD0D7E437D600A" value="1948-11">
<collectingDate id="A54565328027140B67861219ABFA2F9E" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="05890AA3886A0D6D1EFFCE65BC568F39">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="8D022EA71E83CF63097C41406AC5BD2B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="065F39938664C7C0E55DF332FEF63522" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="1BBE8D84F5A93E439FA7F56278E7898D" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="7D148A095CDC7F9E5F8299DA543E2876" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="2E539271454A4D9EBDEE5C958C8D70C4">Elands Bay</collectingMunicipality>
,
<location id="1C0F91C88D5AA107324CD19BD93DDBEA" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:1C0F91C88D5AA107324CD19BD93DDBEA" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK2AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="546FEDEE3AE7CC2FBCC93CF22FD0C8B8" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="221898DAC5B9F307544A24172C1F1606" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="295BB06802A73D187EF19416DB3F85A7" value="1948-11">
<collectingDate id="D8478E22091EB5C793E2EC181B10F566" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="E65D7EB07929F3049AA4655396C0391D">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="C149F9D2B4D96B00D56792AB8C7529C6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="7C88B3D87FA4EF1D900432AD3FEC471E" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="551B13AC94E9E00AC46E6705996E7BE2" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="E0B1E0721A0229A4C00E9BD227548618" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="67C876FCBB86FD7DEB0CC9A1C88BF642">Elands Bay</collectingMunicipality>
,
<location id="7F9696D7F8183201284D16B2E08FDAE9" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:7F9696D7F8183201284D16B2E08FDAE9" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EW2AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="E65980DE008399DA68E23C5CFDD5645B" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="BA02EED01B66181DD3E8CCF163CCB45F" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="B0032D7FF996B41A7749A9C17E76E892" value="1948-11">
<collectingDate id="16BA930CDCAF132315874924BE477A1A" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="6F70DDD2D299567441757320E524748C">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="2765B4BCC4391ECAEAD029CBD0075D64" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="DD907C1545D0AE6E64D5063039E04A66" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="73FE1D61AFC9C5993EFF9F635E79E0F0" collectingDate="1948-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="5DE506D3F790AC6CED533FAB8D80D03F" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="8DB441037F87A0D19D5E73CD4D28D674">Elands Bay</collectingMunicipality>
,
<location id="E17B1BDCE15D182223D36FFBAE9C9DBE" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:E17B1BDCE15D182223D36FFBAE9C9DBE" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EC3AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="9EF443011E736FF094538E17D6CC2E3A" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="5310A169160FFC52C2B85806382CA791" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="1C921AA4FF6939C96C7C74EDED11BC9A" value="1948-11">
<collectingDate id="260BC3C3B7CFDC0801DB34678DB25F10" value="1948-11">Nov. 1948</collectingDate>
</date>
; SAM
<collectorName id="C211B27742F7F982476BC701E26B0EC7">Museum Staff</collectorName>
leg.; SAM-DIP-A 002152,
<named-content id="C5A7806D0743BCE73FFD468D9447928B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="1FEFF63026096E64BE10208F47307532" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL3AG" type="material">
<paragraph id="A0901B2701CEB137A7D2AD698818CC0E">
<heading id="4525078CB87B7EEF50A4CAEE974797E0" reason="title">Other material.</heading>
</paragraph>
<paragraph id="F180FCA287B2CFAD3FC7A4482CCA4008">
<materialsCitation id="2D013D3D0C584F956C900B4823A436C3" collectingDate="1933-11-11" collectionCode="SAMC" collectorName="van Son, G." country="South Africa" latitude="-30.313055" location="Hondeklip-Baai" longLatPrecision="20" longitude="17.275" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<collectingCountry id="CCE417ABB5794FECA105B25DC7D4D485" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="2ED697F9937A521495C858893F883B3D" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="4E7A28A6E1F1AA05B212C374BDE6E0A0" count="1" type="female">1 ♀</specimenCount>
<location id="0A9B01DEFF72445E6D5D620628EF70F7" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:0A9B01DEFF72445E6D5D620628EF70F7" country="South Africa" latitude="-30.313055" longLatPrecision="20" longitude="17.275" name="Hondeklip-Baai" stateProvince="Northern Cape">Hondeklip-Baai</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EU3AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.275000,-30.313056]}">
<geoCoordinate id="FD6CC10E4F73EBF4E32458616C915CA1" degrees="30" direction="south" minutes="18" orientation="latitude" precision="15" seconds="47" value="-30.313055">30 ° 18 ' 47 &quot; S</geoCoordinate>
,
<geoCoordinate id="FC5EE836ED1B64D1C528159B1F0A3919" degrees="017" direction="east" minutes="16" orientation="longitude" precision="15" seconds="30" value="17.275">017 ° 16 ' 30 &quot; E</geoCoordinate>
</named-content>
;
<date id="FAC4F56005052C1C26083F9B40DB6685" value="1933-11-11">
<collectingDate id="4A810C2EA813E84E59A77B4DAB1354F8" value="1933-11-11">11 Nov. 1933</collectingDate>
</date>
;
<collectorName id="2C8852888CFE010C44379A2157E72CAD">van Son, G.</collectorName>
leg.; SAM-DIP-A 002150,
<named-content id="A775675BEAD5DBBAC5422DE8FF07FA4E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="4A5A4B76BC1073C9426820E0A94BB8C3" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="A63ABF6845B2E2467823D9534E1C7D2F">
<materialsCitation id="28E40B717B61A0B9572D6CB0B15B86E5" collectingDate="2009-09-29" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Dikow &amp; Torsten" country="South Africa" elevation="35" latitude="-31.595278" location="Lutzville" longLatPrecision="20" longitude="18.381945" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<collectingCountry id="EC59992708BD50C17502B3EEDCB5C20B" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="8A86C5B4FDC62A92892E590F0313D6C0" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="36C6393452337F17B9720C337364D2D9" count="1" type="female">1 ♀</specimenCount>
<location id="D6DD13180F64B823A4DFC2C44738BCEC" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:D6DD13180F64B823A4DFC2C44738BCEC" country="South Africa" latitude="-31.595278" longLatPrecision="20" longitude="18.381945" name="Lutzville" stateProvince="Western Cape">Lutzville</location>
,
<quantity id="84AC6E6C1CA519137A6CF0A3DED39F35" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
NE;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EC4AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.381944,-31.595278]}">
<geoCoordinate id="95FFE8AD181227E8F45690645B97322F" degrees="31" direction="south" minutes="35" orientation="latitude" precision="15" seconds="43" value="-31.595278">31 ° 35 ' 43 &quot; S</geoCoordinate>
,
<geoCoordinate id="E62AF2E212AC7DED5967D9539D39BBBE" degrees="018" direction="east" minutes="22" orientation="longitude" precision="15" seconds="55" value="18.381945">018 ° 22 ' 55 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="2DE4AB47E88CD44208E0C5D8B42D1F98" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">
<quantity id="52C9B8B33E0C3763129F72229E9FF6BA" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">35 m</quantity>
a. s. l.
</elevation>
;
<date id="038D968FF07C2A2AF5D7E32B91C1AB19" value="2009-09-29">
<collectingDate id="BC78314CAD6D68A62FF1E433B62EDA18" value="2009-09-29">29 Sep. 2009</collectingDate>
</date>
;
<collectorName id="3509C0AD471EF448E9043604CDDB774F">Londt</collectorName>
,
<collectorName id="459F7A180D55E6E3118200581F0C6A8E">Jason</collectorName>
,
<collectorName id="476D9F3F20016E5268BD74903D2D0C4D">Dikow</collectorName>
,
<collectorName id="7C99A7101E49B3CC9438D524DD8D4419">Torsten</collectorName>
leg.; red vegetated sand dune, woody fynbos;
<named-content id="CC9E1E4144DCBF792AF105F54D622FD6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="16B276A2BBA5BBEE37358D4B4124FEB2" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 93557,
<named-content id="4FD566D807E180B4CF0ECF59F11A1784" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="45B9E6C15E2AE682B1D4492CBEFA5CFA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="8215EB9231C7D22E449096D9E69B07B1" collectingDate="2009-09-29" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Dikow &amp; Torsten &amp; Red" country="South Africa" elevation="35" latitude="-31.595278" location="Lutzville" longLatPrecision="20" longitude="18.381945" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="2CF216DDD7EF9717156E43101F5B86A4" count="1" type="male">1 ♂</specimenCount>
<location id="C5109C166FC91F602A91361AE98D73BC" LSID="urn:lsid:plazi:treatment:29672236EE265DADBE3A1901AFBD8391:C5109C166FC91F602A91361AE98D73BC" country="South Africa" latitude="-31.595278" longLatPrecision="20" longitude="18.381945" name="Lutzville" stateProvince="Western Cape">Lutzville</location>
,
<quantity id="3230709D1ECE72B3F4368FB858D78E01" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
NE;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ES4AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.381944,-31.595278]}">
<geoCoordinate id="06FA07687DEB67C07994D7F9FBCBF569" degrees="31" direction="south" minutes="35" orientation="latitude" precision="15" seconds="43" value="-31.595278">31 ° 35 ' 43 &quot; S</geoCoordinate>
,
<geoCoordinate id="C4F39152286922B54B7D460BD3A076D0" degrees="018" direction="east" minutes="22" orientation="longitude" precision="15" seconds="55" value="18.381945">018 ° 22 ' 55 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="7C608E1EC9CD099037E0C9A46BC4BDE7" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">
<quantity id="03DF91FB349D49A623BB0D5555C8E177" metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">35 m</quantity>
a. s. l.
</elevation>
;
<date id="F1C5A0F5D28CEA4FFB416993DF62ADB0" value="2009-09-29">
<collectingDate id="5BC0AF3A505FE86B24707AE74C226839" value="2009-09-29">29 Sep. 2009</collectingDate>
</date>
;
<collectorName id="82568E8F25404A9856FD99A3933C1F88">Londt</collectorName>
,
<collectorName id="154950E906E780DE68C4E31381B71ED7">Jason</collectorName>
,
<collectorName id="B6C8FEEFAD7272F86405348E43C5DC10">Dikow</collectorName>
,
<collectorName id="825DB1198B0B739CA5FDDDF781C6C093">Torsten</collectorName>
leg.;
<collectorName id="8FB29B211BF14B45A5016A52BD463D58">Red</collectorName>
vegetated sand dune, woody fynbos;
<named-content id="8D1A46E34A150850CE2DED8D2FA61C47" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="D2242416D10C520206852540A8A47802" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 93558,
<named-content id="5B4A9224385FB8FEF469B95D814D3126" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="92671EB1115BBEF8D1D82062B0285833" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E64AG" type="distribution">
<paragraph id="3882C905F9956C56269B764DC24FBF54">
<heading id="66142DDAF5E79022A963CEE665E8C9BF" reason="title">Distribution, biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="8D7F64F02614287F3D43CE765B34F78A">
Known from localities in the
<collectingRegion id="C9144CEC6515B2CAF1CA49541B40693E" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
and
<collectingRegion id="33C5D7D9DA557E63CADB11A335029233" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
of
<collectingCountry id="1C5A8666C8C35F4561A6F839772AF897" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="5EC2EC5E69933E48CFABC1359A16E364" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. eremophila, E. syrticola and E. clavicornis specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure7" httpUri="https://binary.pensoft.net/fig/1244180">7</figureCitation>
). This species is only known from
<specimenCount id="9A7BE5985AD6D4495A8CC451FF71D95A" count="11" type="generic">11 specimens</specimenCount>
, collected in 4 collecting events spread between 19332009. The species is known to occur in the Succulent Karoo biodiversity hotspot. Adult flies are active from September - November (Table
<tableCitation id="E585520FABBFDC45CA0B6221AE8D52BF" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="7012D27CF5913715F8AA4DFFF1BE77E0" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="FDA2AAB1E81667BF3E7CB63BD95B4D8F" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="CC11C246E473CF50F0D80C860B3C7FC4" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
<caption id="C49EF28653B0DD0729BA30BD1EED3584" ID-DOI="10.3897/afrinvertebr.66.129611.figure7" ID-arpha="9D7F5FCF-8730-5F73-9437-51B350C5B9E5" httpUri="https://binary.pensoft.net/fig/1244180" startId="F7">
<paragraph id="04A2F6B4A7ED25C4650B9DF94345DF5D">
<label id="8F22F53C17BFFAD1099E6AB6E9BBA0D1">Figure 7.</label>
</paragraph>
<paragraph id="2038076CED6563D7267EA9C04AB71EB4">
Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of
<taxonomicName id="8E1FCBC9D2AFECE70BD3D58292041224" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="eremophila">
<emphasis id="54C1F83E8D256F9CAC0AD791F0556ABA" italics="true">E. eremophila</emphasis>
</taxonomicName>
,
<taxonomicName id="F89E91187D38EC2225B91262A21F7542" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="syrticola">
<emphasis id="C3F5B63D0A244E81457A4AFF22C4A1A3" italics="true">E. syrticola</emphasis>
</taxonomicName>
and
<taxonomicName id="81AFFA9A8A8722624E1231660956FFAD" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="clavicornis">
<emphasis id="87846B9E63D70516FBC5018D55D89638" italics="true">E. clavicornis</emphasis>
</taxonomicName>
specimens studied.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<mods:title id="4F2D434E29E8DCC5EFE47C02BC486CCF">Revision of the Chinese endemic genus Megadrypta Sciaky &amp; Anichtchenko, 2020 (Coleoptera, Carabidae, Dryptini), with description of a new species</mods:title>
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<treatment id="44ABD71D63915D5DAE7E651BF4454556" ID-DOI="http://doi.org/10.5281/zenodo.14834504" ID-Zenodo-Dep="14834504" ID-arpha="44ABD71D-6391-5D5D-AE7E-651BF4454556" LSID="urn:lsid:plazi:treatment:44ABD71D63915D5DAE7E651BF4454556" httpUri="http://treatment.plazi.org/id/44ABD71D63915D5DAE7E651BF4454556">
<subSubSection id="4AA96D3960B4E6D9F772D9619EE0B30C" type="nomenclature">
<paragraph id="DA0E6AE375D87B794A9C96AA55CD9583">
<heading id="C55AEB0F20E48E215301324E9C63C874" reason="label">Genus</heading>
<taxonomicName id="78FF0788FD13EE9A39071FEFB6BE68AB" ID-CoL="8LYNZ" authority="Sciaky &amp; Anichtchenko, 2020" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="6BA440606020206A10A188E3A9C55B15" italics="true">Megadrypta</emphasis>
<bibRefCitation id="CE31629D9E16BE694BAC4240BC0BF2BE" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9BD422CB40ECCD2704230F63D400FD37" type="reference_group">
<paragraph id="6EE3F2FAAE29305489A7B8E14FD0A35A">
<treatmentCitationGroup id="4304B64F788F4126E1FE266BB2E7E64A">
<treatmentCitation id="12BC34AD6CC3A57F39C6A4D8FD07BB1F" ID-DOI="http://doi.org/10.5281/zenodo.3665221" ID-Zenodo-Dep="3665221" author="Sciaky R &amp; Anichtchenko A" httpUri="http://treatment.plazi.org/id/03D77E1287116B1DFF0EFAC26453F83B" page="525" year="2020">
<taxonomicName id="E55F657D214BDBD1B4F611BD9A9C6F6B" ID-CoL="8LYNZ" authority="Sciaky &amp; Anichtchenko, 2020: 525" authorityName="Sciaky &amp; Anichtchenko" authorityPageNumber="525" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="81EDB950A6E649C3E94A3776A1C6A5C4" italics="true">Megadrypta</emphasis>
<comment id="B39D7D9D5433D23F220BC3DD7C6B75D8">
<bibRefCitation id="AF1B84BAFC87D3152AFE53E5D600B549" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020: 525</bibRefCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1FAC" type="type species">
<paragraph id="F049032F3022B769ADACCCA0CCDA2D5A">
<heading id="099C90EBD1F210BCD9C39FAF4C7AA185" reason="title">
<typeStatus id="BD710D321FD71CCC3A1B69271A53BD28">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="F2B6EE36816FE629A3C2FD281D281007">
<taxonomicName id="3ADE2C7B0098F57F5B2C341723A83D9C" authority="Sciaky &amp; Anichtchenko, 2020" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="5B3D8C1F8636FF9DA42CE93A52FF2B8F" italics="true">Megadrypta mirabilis</emphasis>
<bibRefCitation id="A8D5C9A5F977C9DAFDA73331170F0AAC" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELGAC" type="description">
<paragraph id="73B4DAF6EF305EB066E56552173BE313">
<heading id="4E98298E9BE66F712019E927197EE7CE" reason="title">Diagnostic characters.</heading>
</paragraph>
<paragraph id="D7E362AECCF74D52CC5326341148A6BA">
Large dryptine ground beetles with
<abbrev id="ABBRID0ERGAC" xlink_title="length of the body">BL</abbrev>
13.017.0 mm. Dorsal surface dark with more or less bluish metallic luster; mouth parts, antennae, and legs reddish or dark brown. All terminal palpomeres securiform in males, slenderer and weakly expanded at apex in females; labrum trilobate, with medial lobe forwardly projecting; antennae long, aligned backward reaching middle of elytra; antennomere 1 nearly as long as sum of antennomeres 27 combined; temporae well developed. Pronotum cylindrical; anterior angles obtuse; lateral margins of pronotum distinctly doubly concave, before middle and after middle; lateral margins of pronotum completely beaded; posterior angles nearly rectangular, apex rounded. Elytra wide; shoulders extremely narrow; with 68 parascutellar pores; striae deeply punctate; intervals slightly convex and pubescent; posterior margins of elytra truncated, slightly sinuate near outer-apical angles. Hind wings reduced. Legs slender, tarsal claws toothless. Male genitalia stout; apical lamella short and wide, apex rounded and thickened; endophallus with more or less distinct central sclerite. Gonocoxite 2 of ovipositor falciform and smooth, without ensiform setae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVGAC" type="Geographical notes">
<paragraph id="DD708F13C7B5B048F986C368C88FAA60">
<heading id="B76B4CAE25D0F20890E5A75EDFAEF6E8" reason="title">Geographical notes</heading>
</paragraph>
<paragraph id="CB0F64B4D86C1D8298AE22036FFBCDF9">
(Fig.
<figureCitation id="F262B19E22DAB87F65DD4A498A5B6E85" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map for Megadrypta spp. Red circles: M. mirabilis Sciaky &amp; Anichtchenko; yellow circles: M. maozhoui sp. nov." figureDoi="10.3897/zookeys.1226.142678.figure1" httpUri="https://binary.pensoft.net/fig/1244299">1</figureCitation>
). Members of this genus were only found in mid-elevation (
<quantity id="7CCE545C9891E423307880D52638BF61" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">10001500 m</quantity>
) habitats in southern
<collectingCountry id="0D37A47D8EA5C1810332875E6EDBB5BE" name="China">China</collectingCountry>
.
<taxonomicName id="EB02EFBD56F8E7A3E3585B40C7138F7D" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="E65F4E6D70DE7A025DDEB0BA017CD1EF" italics="true">Megadrypta mirabilis</emphasis>
</taxonomicName>
is distributed in the Nanling Area including southeastern
<collectingRegion id="723764B543FAB225DBCF25DA2C02D631" country="China" name="Guizhou">Guizhou</collectingRegion>
, northern
<collectingRegion id="3B1011AB051F72B7E468A5EF975234EE" country="China" name="Guangxi">Guangxi</collectingRegion>
, and northern
<collectingRegion id="35536D8B7DBE06D1350A20ACD9984570" country="China" name="Guangdong">Guangdong</collectingRegion>
.
<taxonomicName id="F8249997E061CDCF5BDE54CC22A970EC" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="C441BB4F75D483B11B28EC35C7B1EF77" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9F3F772FB978112F8F3E3177029F8E5" rank="species">sp. nov.</taxonomicNameLabel>
is distributed in southwestern mountains around the
<collectingRegion id="E2DA4EC3674A8B7522B7F0388BB9C9EC" country="China" name="Sichuan">Sichuan</collectingRegion>
Basin, and it is currently recorded from central
<collectingRegion id="07C0977EF3A0E3B4AC831F69F07EBC1A" country="China" name="Sichuan">Sichuan</collectingRegion>
and northeastern
<collectingRegion id="B9E1CC9D9C211625A63976A10251C3A2" country="China" name="Yunnan">Yunnan</collectingRegion>
.
</paragraph>
<caption id="362443FF5475DF88BB26E1DA9DCF1E02" ID-DOI="10.3897/zookeys.1226.142678.figure1" ID-arpha="1A5CF65D-6313-5315-8327-0C33CF99149A" httpUri="https://binary.pensoft.net/fig/1244299" startId="F1">
<paragraph id="89D4DC97CE237DF7A1CE63E9B86A5A7C">
<label id="768C77DAE24C79F7D94F071FB01633E2">Figure 1.</label>
</paragraph>
<paragraph id="E1372DEBE3BD293FA2846A643C8290BE">
Distribution map for
<taxonomicName id="E0011458414A90FF07F0189AAACA8F35" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9D3F60419A145FC68E44AEB5BA9FCB49" italics="true">Megadrypta</emphasis>
spp.
</taxonomicName>
Red circles:
<taxonomicName id="C09DBD1DFA470D7E35E975DCD9652142" authority="Sciaky &amp; Anichtchenko" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="942093C2576A7FFFB544ACFE71666231" italics="true">M. mirabilis</emphasis>
Sciaky &amp; Anichtchenko
</taxonomicName>
; yellow circles:
<taxonomicName id="4789298FFB65FDFF1DC5180F3B9A35CF" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="4EF39BC206C615E4EBAC5D5FB0B9738F" italics="true">M. maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1160A946D39E367FA90F9C4CBE862928" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="858171D31661483CE97AFD8F2F50E997">
<title id="465292BA0BE5373309467A06636CAB7C">
Key to species of
<taxonomicName id="B18B4BD396EE924F86DD4523336A02A0" authority="Sciaky &amp; Anichtchenko, 2020" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="E5D81C894D435D4C692FCE4CA1634E35" italics="true">Megadrypta</emphasis>
<bibRefCitation id="C00CC6930CCE1805F9B53A931EFFA1EA" DOI="10.11646/zootaxa.4731.4.5" author="Sciaky R &amp; Anichtchenko A" issue="4" journalOrPublisher="Zootaxa" pagination="522-530" refId="B2" refString="Sciaky R, Anichtchenko A (2020) Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530. https://doi.org/10.11646/zootaxa.4731.4.5" title="Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)." volume="4731" year="2020">Sciaky &amp; Anichtchenko, 2020</bibRefCitation>
</taxonomicName>
</title>
</paragraph>
<paragraph id="0EB3AC98493E8E1D114ACBA93A9E137D">
<table id="TID0EPFAC" rules="all">
<tbody id="EB7A00F0ADB6579B33EFA41E5611FA2D">
<table-wrap id="F2CCFF052D946875278E49B2E5DFF572" content-type="key" orientation="portrait" position="anchor">
<tr id="A820F26B251506EAFEDDB52E517F4A19">
<td id="93AC90045E427CB940E37CD7D02022D8" colspan="1" rowspan="1">1</td>
<td id="F2AA5A82D87FB816A5F8BE7389611BBF" colspan="1" rowspan="1">
Body very large (
<abbrev id="ABBRID0E2JAC" xlink_title="length of the body">BL</abbrev>
1317 mm); pronotal lateral bead complete; elytron with 68 parascutellar pores; elytra strongly narrowed toward shoulders; hind wings strongly reduced; gonocoxite 2 of ovipositor without setae
</td>
<td id="6A5ED73B68A451AF8F169104830946F2" colspan="1" rowspan="1">
<emphasis id="3D001519354F0794CF094B2B1C35FBE8" bold="true">
2 (
<taxonomicName id="08682864DA478AA64CD3224460F99F47" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="94D18BBAD29DA837D0C9F190E37ECF69" bold="true" italics="true">Megadrypta</emphasis>
</taxonomicName>
)
</emphasis>
</td>
</tr>
<tr id="A511A73EB15D28799A7D26A525CD5300">
<td id="67B4EC8C62011A9D100A347CFC789236" colspan="1" rowspan="1"></td>
<td id="07C0516C512B20BFD71CE02D1B0E371D" colspan="1" rowspan="1">
Body smaller in size (
<abbrev id="ABBRID0EVKAC" xlink_title="length of the body">BL</abbrev>
&lt;14 mm); pronotal lateral bead complete or not; elytron with 14 parascutellar pores; elytra less pronouncedly narrowed toward shoulders; hind wings well developed; gonocoxite 2 of ovipositor setose (except for the genus
<taxonomicName id="34C22A7E3D10555F46169B498AB4DC40" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Nesiodrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0D6C3B26482EC8A443BE8C053CECC665" italics="true">Nesiodrypta</emphasis>
Jeannel, 1949
</taxonomicName>
without setae)
</td>
<td id="772C980CEA8C06F5D5D9062C7A21A1B8" colspan="1" rowspan="1">
<emphasis id="EA09EF5F459EBDDA96B7B7F5E18F29F2" bold="true">
other
<taxonomicName id="7554D1E6D4761A7A4A2B42AB5737A137" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Dryptini">Dryptini</taxonomicName>
genera
</emphasis>
</td>
</tr>
<tr id="ACF2B4D3F16946B1F1C12872708D18EE">
<td id="09395B7064F0A264501D7A61810249E1" colspan="1" rowspan="1">2</td>
<td id="2D11EC73DC7895DF6FBDA42FBAC88A6B" colspan="1" rowspan="1">
Dorsal surface black, with bluish luster very faint and hardly visible; femora dark brown to piceous, tibiae and tarsi sometimes lighter (Fig.
<figureCitation id="2854A83B7C1BD30A368BF05057D15569" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Megadrypta spp. A holotype of M. maozhoui sp. nov. B female of M. mirabilis from Hunan. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure2" httpUri="https://binary.pensoft.net/fig/1244300">2 B</figureCitation>
); mentum with anterior margin straight at middle (Fig.
<figureCitation id="C6CF14E4841306433884542C5EF40E5F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mentum of Megadrypta spp. A M. maozhoui sp. nov. B M. mirabilis. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1226.142678.figure6" httpUri="https://binary.pensoft.net/fig/1244304">6 B</figureCitation>
); elytral striae incised with large punctures (much smaller than interval width); in lateral view, median lobe of aedeagus equally broadened before apical third and then abruptly narrowed, ventral margin strongly curved before apical lamella (Fig.
<figureCitation id="6CC9C2524DC3D9F91DABBEAC55353489" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4 A</figureCitation>
)
</td>
<td id="DA4CD8EC780544EBFC4F7F574EEC5312" colspan="1" rowspan="1">
<taxonomicName id="429C786581E1E84B9657C68E4E8A0E53" authority="Sciaky &amp; Anichtchenko" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="E9B0556E13C2F5F3D6D9F7BA8378CCCA" bold="true">
<emphasis id="811F70D46C571E912F1AD472BB56413F" bold="true" italics="true">M</emphasis>
.
<emphasis id="95E579DCF6448FAC15E5C344A61A5A40" bold="true" italics="true">mirabilis</emphasis>
Sciaky &amp; Anichtchenko
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="6869687C79547D45B355A213F6FF63DA">
<td id="FC444D0CD07CC293E90C1D318B0F1066" colspan="1" rowspan="1"></td>
<td id="195F1BF87B1CA026F2BC18AAE40285B4" colspan="1" rowspan="1">
Dorsal surface dark with distinct bluish metallic luster; legs entirely reddish brown (Fig.
<figureCitation id="CE641A8DDB04F0F95162E81F4F295C41" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Megadrypta spp. A holotype of M. maozhoui sp. nov. B female of M. mirabilis from Hunan. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure2" httpUri="https://binary.pensoft.net/fig/1244300">2 A</figureCitation>
); mentum with anterior margin concave at middle (Fig.
<figureCitation id="AB503CE2ADBC75F06456AD2DF4EFEABB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mentum of Megadrypta spp. A M. maozhoui sp. nov. B M. mirabilis. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1226.142678.figure6" httpUri="https://binary.pensoft.net/fig/1244304">6 A</figureCitation>
); elytral striae interrupted by very coarse foveae (of similar size as interval width) (Fig.
<figureCitation id="12AE5E7D728BE688499C4337D60247D0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3 C</figureCitation>
); in lateral view, median lobe of aedeagus gradually attenuate to apex, ventral margin almost straight before apical lamella (Fig.
<figureCitation id="55BD6ED7C1718E6D746CE0E96AA5FB51" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5 A</figureCitation>
)
</td>
<td id="1D66948226F54BA48F0C62BF0F58D3BD" colspan="1" rowspan="1">
<emphasis id="DDA22C7769C2A7998926E82BBFEE9A4B" bold="true">
<taxonomicName id="151045B0FAC64C078373A4A1A762A530" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="EEC9733834AA4BDC8571C8BE81B318EE" bold="true" italics="true">M</emphasis>
.
<emphasis id="A005DC1D54EA18C8201ACC4C7E89BC56" bold="true" italics="true">maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5B7623BCC8C123D73F2C65F491D1EAF7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,384 @@
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<mods:title id="B16C121AB673BE278BBA48C92ADDC83A">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
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<mods:name id="A1FC7C135EC3BFDA2D13195875DF4EFD" type="personal">
<mods:role id="8A0A83A895AC9CBCFE1437803B31292C">
<mods:roleTerm id="21A458D6B083EFD3EF632B13409FE5F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8059FF89EF63A742338A5754F589346F">Rollinson, Lisa</mods:namePart>
<mods:nameIdentifier id="0DDE42353FAE5D130C57A8AEDC45011D" type="ORCID">https://orcid.org/0009-0007-0152-7073</mods:nameIdentifier>
<mods:affiliation id="882CED09F8B11E26970CA8FD9A1459C2">Texas A &amp; M University, College Station, Texas, USA</mods:affiliation>
</mods:name>
<mods:name id="0310CCBD251ECDA3631ED49F80F08A65" type="personal">
<mods:role id="6EAD7C5ED730FD8D9C6C5EB4C5BFF417">
<mods:roleTerm id="C366659418C0C758F7412F4D9CB4B487">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B2D774F4399E9AA5989CA7024EE53A3">Cabrero, Allan</mods:namePart>
<mods:nameIdentifier id="E12CFED6F1C632A771F7FB2A639052D7" type="ORCID">0000-0002-2935-3534</mods:nameIdentifier>
<mods:affiliation id="CA4BBF9840C808660B1992208DF0A791">National Museum of Natural History, Washington, D. C., USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FA00DE45D5C8B371ED9343528998E126">text</mods:typeOfResource>
<mods:relatedItem id="F0717413EC44E97C91FAB2F7D824F852" type="host">
<mods:titleInfo id="5FD7A72DBBDDF8A5B8080DB71C548CC5">
<mods:title id="30A1F5ABAD29DDEE36251CAA04CCA39E">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="4CD4F89E28B083CBF97EA1C6B053CFB5">
<mods:date id="3498450A46965FE53C27EDC48EA00B10">2025</mods:date>
<mods:detail id="FEF938D84878A9776BFA0C9557F06EF6" type="pubDate">
<mods:number id="F347721492866BEE1A3A99F0F5086666">2025-02-07</mods:number>
</mods:detail>
<mods:detail id="58A2670CCBF1CB9991204B95184CECFA" type="volume">
<mods:number id="7298161EEA1FECA459EA834ED816EA69">66</mods:number>
</mods:detail>
<mods:detail id="E2DF0C4E2D490EB21300C6C9D17DC13F" type="issue">
<mods:number id="D19511EC807799B67351CFCE3A2AB576">1</mods:number>
</mods:detail>
<mods:extent id="4E9AA4E141F80AF63CFE037DCEF2E609" unit="page">
<mods:start id="57D5087934C9EB542EE98386BC5232AD">73</mods:start>
<mods:end id="D1931AB43E923A42B9CEB88775068D45">115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="87434F77A12003015AED2913EA56E4C3">journal article</mods:classification>
<mods:identifier id="FC352A2B918F2408037CAD03C0CE0D88" type="DOI">10.3897/afrinvertebr.66.129611</mods:identifier>
<mods:identifier id="6D3B4E9E6A7BAAC7E1529102EB80C976" type="ZooBank">9A70A477-D862-40E5-A132-3227D96C3ECC</mods:identifier>
</mods:mods>
<treatment id="87F748CF9F83587CA797FD03A5C61C0A" ID-DOI="http://doi.org/10.5281/zenodo.14834494" ID-Zenodo-Dep="14834494" ID-arpha="87F748CF-9F83-587C-A797-FD03A5C61C0A" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A" httpUri="http://treatment.plazi.org/id/87F748CF9F83587CA797FD03A5C61C0A">
<subSubSection id="5418187A27F1D04F89D24EFCE50E7CA0" type="nomenclature">
<paragraph id="73A838ED1C5859F43E6C4A5D64FB4DD5">
<taxonomicName id="CFF82CFECC99D370D394A24DC3DA7FD5" authority="(Hesse, 1956)" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="imitata" status="comb. nov.">
<emphasis id="DD23D236070A17E23BD443C9445BF12A" italics="true">Enica imitata</emphasis>
(
<bibRefCitation id="1DD442220AEE4A6A4967FF2F96F47E0A" author="Hesse AJ" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 27-46. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B9034913F4FD0209E84918FAFC1656" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DBA03216FF6DE1FB2C9A6D92DDCFDCE0" type="description">
<paragraph id="3BE913A65BA52E24C17A695FD3E3FF74">
<figureCitation id="A1D8CE7DA361E8D8076F0EF1E5D9C789" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Enica henicoides ♀ SAM-DIP-A 002170 (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica imitata ♀ NMSA - DIP- 90027 (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure8" httpUri="https://binary.pensoft.net/fig/1244181">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGWAI" type="Taxon depository">
<paragraph id="B4F8D0558AA68B76D3EF934258907145">
<heading id="587BE8D31BA53A0E46304019795567F9" reason="title">Taxon depository.</heading>
</paragraph>
<paragraph id="69C451E046AB0D44E4DFC9CC6AD68163">
ZooBank:
<ext-link id="DDACF68F294DACDC145826EB851715B6" ext-link-type="uri" xlink_href="https://zoobank.org/B9E4599C-A9B0-4162-AB1D-8E79EF2D72B8" xlink_type="simple">
https://zoobank.org/
<uuid id="76AD0E06799EDC3467080BB725976AD2">B9E4599C-A9B0-4162-AB1D-8E79EF2D72B8</uuid>
</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERWAI" type="diagnosis">
<paragraph id="D4226FC804A1173EAB6CAA932B9794D0">
<heading id="7E58BCEA355A3A484790497D33BD95D9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="118A90FA19C8B91784A1BA393DB6B6C4">The species is distinguished from other species in the genus by the white scales present on the occiput, additionally two dark elongated spots can be seen on the dorsal margin of the occiput. Metathoracic femur has white scales ventrally and brown scales dorsally.</paragraph>
</subSubSection>
<subSubSection id="SECID0EWWAI" type="description">
<paragraph id="9AA95F40B36DB36E725E6440752D355D">
<heading id="B59D38DD9822CD93FD656026A31A0E78" reason="title">Description.</heading>
</paragraph>
<paragraph id="3E53632B3D97C96B858245AF6453EB65">
<emphasis id="70144A14C0E536E0814100583DFBEFE3" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, with dark brown to black triangular spot extending to eyes, males: similar to head, slightly darker than head; frons black setose, with white scales, males no setation near eyes, females setose near eyes only dorsally; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput whitish yellow setose, brown setose on occiput spots, white scales antero-dorsally, dorsal eye margin with dark brown to black spots present, touching or almost touching eye margin, never extending ventrally past level of antennae; gena creamy white, extremely sparsely setose.
</paragraph>
<paragraph id="E415127B56455FEC9D0F8281B75D394E">
<emphasis id="096CE6A91F59AEDA6F708DD166B4821E" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, light brown to brown, brown setose.
</paragraph>
<paragraph id="04588B780C99AE6B235A8775DBB20A4B">
<emphasis id="6857D75F1220CCBE13D166AA3EB02A00" bold="true" italics="true">Antenna</emphasis>
: dark brown; scape creamy white, approximately as long as pedicel, black setose dorsally and ventrally, setae extending to tip of pedicel; pedicel reddish brown or dark brown, short black setose dorsally; postpedicel dark brown, tapering distally, longer than scape and pedicel combined, white or light yellow scales dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="533992A0127B136A76E3AF9E15D72287">
<emphasis id="9493BC4CB4460F77693A5D95BC0C47DB" bold="true" italics="true">Thorax</emphasis>
: reddish brown and light orangish brown; scutum predominantly black, reddish on margins, grey to brown scales centrally and white scales peripherally, with white scales in two short anterior stripes sub medially; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, white scales dorsally; metepimeron silvery grey pubescence, asetose; scutellum red, anterior black spot, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of light brown and white scales, black setose or white scales, black setose.
</paragraph>
<paragraph id="DA44BFB10ECDF94452ABAD9FC536C25A">
<emphasis id="F72B95C115F909307BCAB9DA1B4530B4" bold="true" italics="true">Leg</emphasis>
: dark brown, covered in scales, some regular setae present; pro coxa dark reddish brown, silvery grey pubescence, white scales, thick black setose; pro femur dark brown, white scales ventrally, brown scales dorsally, few short black setae; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa dark reddish brown, white pubescence, white scales, thick black setose; mes femur dark brown, white scales ventrally, brown scales dorsally, few short black setae; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa dark reddish brown, white pubescent, white scales, thin white setose, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur dark brown, white scales ventrally, brown scales dorsally, few short black setae; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="FF84672941646C4982C5FFFCDDB36A9C">
<emphasis id="E798173B70D19430A25A923562827A15" bold="true" italics="true">Wing</emphasis>
: 7.89.0 mm, microtrichia absent; membrane color pattern males: slightly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="46D42669B8F2E7C9FE29F3877C7967CD">+ 3</subScript>
, base of m
<subScript id="4D3BFB5C266AF579F74CB6FE4C537E98">2</subScript>
, m-cu, split between M
<subScript id="23C43D5CF4E13942AFBCAFDE659C2819">1</subScript>
+ 2 and M
<subScript id="5DB8E006E9E0D40720784EA088167440">3</subScript>
, middle of br and basal portion of r
<subScript id="DF8A138B8035CBEEE4F1EC377DFB123D">1</subScript>
, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="EAFBB9D7115FF93FE3430B1E0188ABAB">+ 3</subScript>
, base of m
<subScript id="2197D69F662E5D2B9546223F8F6AE746">2</subScript>
, m-cu, split between M
<subScript id="A0CD491CC23263FE9A39C966AD7A0D8A">1</subScript>
+ 2 and M
<subScript id="1843CA063B67B3473C77CF84A87C509B">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="BC16DE2D5CCFEB195AD4F6E1CD9EB15F">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="6C95D4BD84326A5EE541248AB98D805C">1</subScript>
open; R
<subScript id="D4FF9EE9CF9F801FDF04227740F76F33">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="C3BA1F804057DA16259909D873DE5C6F">4</subScript>
open, proximally R
<subScript id="9437ABCA5CFDBB18105B946515130A17">4</subScript>
and R
<subScript id="1FA943C417864D98DFC77A8EDCAC3AF7">5</subScript>
parallel, R
<subScript id="A6F605FA63D92FF029A7A7EECF8AC610">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="CA497859A863EC17BBD47414BD14EF60">5</subScript>
terminating posterior to wing apex; r
<subScript id="5196AD788E2A1C643C55A2C82671A863">5</subScript>
open; M
<subScript id="2821DE3C26E576901BEE103757547ED3">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="A9F3B964C855AA87E84F18DC047DB4B4">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="F22D386C49F5EC334729B61EDD3867AE">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.
</paragraph>
<paragraph id="F5D6EA2B1A6A05E559F1262190571030">
<emphasis id="616322BB1CF7CFC46EAEE8A06CD9411F" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, black; tergites smooth, setae with small sockets only; T 1 predominantly black, no red or a very thin line of red posteriorly, snow white setose, with scales; T 2 T 7 predominantly black, light brown setose, white scales, white scales on spots on posterior margins of T 2 and T 4, single spot medially on T 2; S 18 predominantly black with reddish brown, dense snow white scales, thin black setose and long white setose; lateral margin black setae remaining close to abdomen, male T 8 white scales, black setose.
</paragraph>
<paragraph id="45CFC8BF29494817812BE265D510BD12">
<emphasis id="BD412419C725222309141D8DA22F3757" bold="true" italics="true">Terminalia</emphasis>
: gonocoxites longer than wide, broader at base, convex basally with straight apical third, fused almost completely with phallus, with short, fine hairs apically; gonostylus with sharp projection; phallus inflated and helmet-like at base, apically separating from gonocoxite, slightly curving back towards gonocoxite; lateral aedegal apodeme round, extending laterally; ejaculatory apodeme long, rounded in lateral view, extending significantly past anterior margin of the gonocoxites, ending under lateral strut. (Fig.
<figureCitation id="7DAB352FCCEDE455CD9361C0BC0ED47C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral and ventral view of Enica male terminalia. Terminalia of Enica imitata NMSA - DIP- 90025 (A, C); Terminalia of Enica adelphe sp. nov. NMSA - DIP- 89520 (B, D). Abbreviations: b ph = base of phallus; ej apod = ejaculatory apodeme; goncx = gonocoxite; gonst = gonostylus; lat aed apod = lateral aedeagal apodeme; ph = phallus. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure4" httpUri="https://binary.pensoft.net/fig/1244177">4 A, C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA1AI" type="material">
<paragraph id="0FC1EED8FE535555511F7DD685CCDC99">
<heading id="609116ADAD336B716DEF338E9B998DBF" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="CFDA598F032436BB187FA53A539567F7">
<materialsCitation id="B3EB8C9EDF350D4D3E5C7D9371480F50" collectingDate="1952-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.967224" location="Moordenaars Karoo Lammerfontein" longLatPrecision="20" longitude="20.815556" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="62E4BF22580FAC44CDBFD51168EFC761" bold="true">
<emphasis id="4476C472844B4B10A88A3EFD31C3943A" bold="true" italics="true">
<typeStatus id="03C0FFC70379D854EFA81968E9F282FA" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="6F832CD1F70FA0B2C55EA1A890665CF3" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="07F8430BAA3C69A53B3E95AAD6995466" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="6F5CB5726A585BCE234C8E39C7AF0B99" count="1" type="male">1 ♂</specimenCount>
<location id="1DE6C10EDCD9430DB1034DE92EC90731" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A:1DE6C10EDCD9430DB1034DE92EC90731" country="South Africa" latitude="-32.967224" longLatPrecision="20" longitude="20.815556" name="Moordenaars Karoo Lammerfontein" stateProvince="Western Cape">Moordenaars Karoo Lammerfontein</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EO1AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[20.815556,-32.967222]}">
<geoCoordinate id="C8BD7A91D4B714AC0745C206E61CD7AC" degrees="32" direction="south" minutes="58" orientation="latitude" precision="15" seconds="02" value="-32.967224">32 ° 58 ' 02 &quot; S</geoCoordinate>
,
<geoCoordinate id="78C43287121D224AFE2E0D95B2959CFD" degrees="020" direction="east" minutes="48" orientation="longitude" precision="15" seconds="56" value="20.815556">020 ° 48 ' 56 &quot; E</geoCoordinate>
</named-content>
;
<date id="54FC66EBD4E480BF91278F0BBB3F42F3" value="1952-10">
<collectingDate id="3A955EA78EF9ECF60BA97BE7424A1B8C" value="1952-10">Oct. 1952</collectingDate>
</date>
; SAM
<collectorName id="01D65537E69C0D20A4FB962932CE34DE">Museum Staff</collectorName>
leg.; SAM-DIP-A 002172,
<named-content id="DA0A746CB519D35661352BB1DA73907E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="5B0AEBE0DABFECDE29703969722C88D9" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
<paragraph id="7795CACCCD1292AD6E95CE3A1EE7939E">
<materialsCitation id="63CFA5F880B1E50E3140FE5312D1B362" collectingDate="1952-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.967224" location="Moordenaars Karoo Lammerfontein" longLatPrecision="20" longitude="20.815556" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="CDAA0CC50E2C10E66340F1CBB6EFEB92" bold="true">
<emphasis id="4CC78B6D5800868C2A6072C180E6F1F8" bold="true" italics="true">
<typeStatus id="624B12C33987AE7189179324CDDD9EE0" type="paratype">Paratype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="9A7F57F4CFD4F36C339C8CB886656257" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="E1EB2E370E83181D9389EAC0FC9B11A1" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="5F0B611330DCA8640C92F23EC4971AD6" count="1" type="male">1 ♂</specimenCount>
<location id="C4667E9644D4FE285C95C868090978FC" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A:C4667E9644D4FE285C95C868090978FC" country="South Africa" latitude="-32.967224" longLatPrecision="20" longitude="20.815556" name="Moordenaars Karoo Lammerfontein" stateProvince="Western Cape">Moordenaars Karoo Lammerfontein</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EA2AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[20.815556,-32.967222]}">
<geoCoordinate id="BB91B5FF43DB53CE74AF7B8B6739D620" degrees="32" direction="south" minutes="58" orientation="latitude" precision="15" seconds="02" value="-32.967224">32 ° 58 ' 02 &quot; S</geoCoordinate>
,
<geoCoordinate id="F72966CFD4CD4D9205D4997357AEB6F7" degrees="020" direction="east" minutes="48" orientation="longitude" precision="15" seconds="56" value="20.815556">020 ° 48 ' 56 &quot; E</geoCoordinate>
</named-content>
;
<date id="F9F402C3F02AC8C1D18184BB20D2E6DC" value="1952-10">
<collectingDate id="29ECC9F680420E57BF9BEA21EAB82A65" value="1952-10">Oct. 1952</collectingDate>
</date>
; SAM
<collectorName id="3C3CC0C52FD358896462DE86007F1B18">Museum Staff</collectorName>
leg.; SAM-DIP-A 002172,
<named-content id="6D6752E26D957D29C1CCA73523448E4C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="D3D015A9A8C3CAA2916B082AA3F26C50" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EI2AI" type="material">
<paragraph id="2AAC34920FEC074CD492B9B045D0A5B8">
<heading id="47DF783A5CFDD576ED94EC537C9CA0BA" reason="title">Other material.</heading>
</paragraph>
<paragraph id="BC0D3EC516479406DF419BD439856880">
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<collectingCountry id="8772E4F9953B26966757C5311D7AD143" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="2A84E07383CC783B7B7E64FFA1C23292" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="10CB5EE673659A5E7D31EAF1BE1E115B" count="1" type="female">1 ♀</specimenCount>
<location id="F3565154E6F54D53366329B30F137684" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A:F3565154E6F54D53366329B30F137684" country="South Africa" latitude="-31.569445" longLatPrecision="20" longitude="19.732224" name="Calvinia" stateProvince="Northern Cape">Calvinia</location>
,
<quantity id="15ED935F3A224E279A2E8C616A655CBB" metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
S;
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<geoCoordinate id="B9654ED9729473707555FDFB1C352C48" degrees="31" direction="south" minutes="34" orientation="latitude" precision="15" seconds="10" value="-31.569445">31 ° 34 ' 10 &quot; S</geoCoordinate>
,
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<quantity id="C9CEA0DCCEBF7132015E18A44F269325" metricMagnitude="3" metricUnit="m" metricValue="1.08" unit="m" value="1080.0">1080 m</quantity>
a. s. l.
</elevation>
;
<date id="751794F05E7D635B5646ADBE8ACCBE11" value="2002-09-07">
<collectingDate id="878C7B1BC836EC6A0D2E4D081B96EFBC" value="2002-09-07">07 Sep. 2002</collectingDate>
</date>
;
<collectorName id="8D469ADD9308CFF17759A7CC3776CB9A">Londt</collectorName>
,
<collectorName id="661771954F0F321BE4C3ADAF42B7ADD7">Jason</collectorName>
leg.; sandy roadside;
<named-content id="32532393685E7EFFC25B91616B2CED0A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="F31A76B4180E8164098733E3FEE59640" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 84590,
<named-content id="CE92A3B1FCCE382459B91F87A3E92454" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="5BC993F740DC019CC209BC2D3D550E6E" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="136E41F280212418088C77C507D4B003" collectingDate="1998-11-07" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Karoo Macchia" country="South Africa" elevation="1290" latitude="-32.252777" location="Renoster River Area" longLatPrecision="20" longitude="20.694166" municipality="Sutherland" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape">
<specimenCount id="DE1C37AA2110434FB9993FC498498F99" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="743485C527FE7110463DA668FB322F1F">Sutherland</collectingMunicipality>
,
<quantity id="F686168C349889AFC68241E3FF536486" metricMagnitude="4" metricUnit="m" metricValue="1.8" unit="km" value="18.0">18 km</quantity>
N,
<location id="208B453C85B458682002B55922D339FA" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A:208B453C85B458682002B55922D339FA" country="South Africa" latitude="-32.252777" longLatPrecision="20" longitude="20.694166" municipality="Sutherland" name="Renoster River Area" stateProvince="Northern Cape">Renoster River Area</location>
;
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<geoCoordinate id="9577BD107B8B137DCF476D8A717A0BE8" degrees="32" direction="south" minutes="15" orientation="latitude" precision="15" seconds="10" value="-32.252777">32 ° 15 ' 10 &quot; S</geoCoordinate>
,
<geoCoordinate id="76D98E110E1033473F2F8063875449A0" degrees="020" direction="east" minutes="41" orientation="longitude" precision="15" seconds="39" value="20.694166">020 ° 41 ' 39 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="ACCE9BF0D04E48AAA0543D6860836246" metricMagnitude="3" metricUnit="m" metricValue="1.29" unit="m" value="1290.0">
<quantity id="418BC62A19326B67648D21A9C2F8D916" metricMagnitude="3" metricUnit="m" metricValue="1.29" unit="m" value="1290.0">1290 m</quantity>
a. s. l.
</elevation>
;
<date id="E44B79243320A4747E1DA5CABAA04C56" value="1998-11-07">
<collectingDate id="FB02A8AB252ED56388F539CB236C9ACC" value="1998-11-07">07 Nov. 1998</collectingDate>
</date>
;
<collectorName id="0AD90F5C5F3F67EE46832E1C107F075A">Londt</collectorName>
,
<collectorName id="3ABFAD33368FC23A8634D0E45529E2DD">Jason</collectorName>
leg.;
<collectorName id="73EAF1D1B2DE74B7F4871AB9D3377A93">Karoo Macchia</collectorName>
;
<named-content id="02C63EA814BB6837840766523AF2B558" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="C99E3D379115F928DD1F7B11CFAF16AD" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 90025,
<named-content id="BA544DDB213C45CAFA72D99C9EEA465B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="87D8C5F7C4C66211D2021DB4AB67B3C7" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="89668FFD72C49D8F0B330CFEEC65FA47" collectingDate="1998-11-07" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Karoo Macchia" country="South Africa" elevation="1290" latitude="-32.252777" location="Renoster River Area" longLatPrecision="20" longitude="20.694166" municipality="Sutherland" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="3CB8D08980B7EFB5504605E414E00418" count="1" type="female">1 ♀</specimenCount>
<collectingMunicipality id="267B89BAD84DDBC78B3470F3A21930DA">Sutherland</collectingMunicipality>
,
<quantity id="32A8351C8A21D6294AF6E00F135750E7" metricMagnitude="4" metricUnit="m" metricValue="1.8" unit="km" value="18.0">18 km</quantity>
N,
<location id="5859B20A43D0186A535CF1EF923A4BEA" LSID="urn:lsid:plazi:treatment:87F748CF9F83587CA797FD03A5C61C0A:5859B20A43D0186A535CF1EF923A4BEA" country="South Africa" latitude="-32.252777" longLatPrecision="20" longitude="20.694166" municipality="Sutherland" name="Renoster River Area" stateProvince="Northern Cape">Renoster River Area</location>
;
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<geoCoordinate id="A156C104D9BA87122CAB52C7E7AC35DA" degrees="32" direction="south" minutes="15" orientation="latitude" precision="15" seconds="10" value="-32.252777">32 ° 15 ' 10 &quot; S</geoCoordinate>
,
<geoCoordinate id="A952698997361495C2FF806EC19AD1D9" degrees="020" direction="east" minutes="41" orientation="longitude" precision="15" seconds="39" value="20.694166">020 ° 41 ' 39 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="47FCE2E295FA65189BF2F058294E7314" metricMagnitude="3" metricUnit="m" metricValue="1.29" unit="m" value="1290.0">
<quantity id="CA92B9E355178E82C8FEF9980E89653F" metricMagnitude="3" metricUnit="m" metricValue="1.29" unit="m" value="1290.0">1290 m</quantity>
a. s. l.
</elevation>
;
<date id="1252B344EF65E8B3D1BDF749A44EB452" value="1998-11-07">
<collectingDate id="F734BA56215F835F0887E88D93DB5BDF" value="1998-11-07">07 Nov. 1998</collectingDate>
</date>
;
<collectorName id="8B3EAD3A5727E7068877154894B32834">Londt</collectorName>
,
<collectorName id="F3B2823E2491339EF3E7C7DF651274B9">Jason</collectorName>
leg.;
<collectorName id="4A64405FBABEC6E518548B1465B0A0DB">Karoo Macchia</collectorName>
;
<named-content id="C249B5060497377E38B5DC3642A5B75D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="C0BB68739908F2840D4C7C8ACC3519FE" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 90027,
<named-content id="FEB00630D2C8BB33721A476F9A1E719E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="4DB4FD667B211A34D1C73FD08BB18412" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E53AI" type="distribution">
<paragraph id="B94D645927B7AD96249917DDD481492A">
<heading id="EA69ED01A4D7652FCA670F6A2A48AAFB" reason="title">Distribution, biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="781AE6ECE366EF8F1305D8962AE686FF">
Known from localities in the
<collectingRegion id="46425DEA9D48DA509A38A3701854C1A4" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
and
<collectingRegion id="326877DFDEB87142F5773939C6E83F70" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
of
<collectingCountry id="DC02986E776FAC6E88DCD1A4CF7AA194" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="42F62DE4B959D8498668F000AC9D4987" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. imitata and E. adelphe sp. nov. specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure5" httpUri="https://binary.pensoft.net/fig/1244178">5</figureCitation>
). This species is known from
<specimenCount id="47427011146417DB5F0CB7FB6E3E7430" count="5" type="generic">5 specimens</specimenCount>
, collected in 3 collecting events spread between 19522002. The species is known to occur in the Succulent Karoo and the
<collectingRegion id="552787BBD25A69DBD97560CF6B39EFD8" country="South Africa" name="Western Cape">Cape</collectingRegion>
Floristic Region biodiversity hotspots. Adult flies are active from September - November (Table
<tableCitation id="53BC4F43E1276C85C907FCC81FD07BB3" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="A168BE63DB0250CEB2B9F3A0FAB4A372" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="0373FE70B535B9BE923E93467C2E007A" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="7EC4BB6A2EB33292CAB1913E0C2B63B4" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,301 @@
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<mods:title id="2CE10B34BC37CD119DB6ED3092B6E38F">Two new species in the green lacewing genus Santocellus (Neuroptera, Chrysopidae, Leucochrysini)</mods:title>
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<mods:affiliation id="10606E94D9E00C4085A654C8C31B0817">Department of Entomology &amp; Nematology, University of California, Davis, CA 95616 and Department of Entomology, Comstock Hall, Cornell University, Ithaca, NY, USA</mods:affiliation>
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<paragraph id="8649E6B9BEA22BCAA4BE936BF5E8FC02">
<taxonomicName id="B1708ED18352B6848F158E7CDDAAC6A6" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="helene" status="sp. nov.">
<emphasis id="6C479D55A26BDE4FAB9330559F81E942" italics="true">Santocellus helene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDB5A5B15CDDBAD7CC7E28E42B9C8119" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D759DEDC6005A585AE61819E0D6ECC71" type="description">
<paragraph id="0BF34A4D7C49B6545364A32FE767F718">
<figureCitation id="F4C90161C5DAFDB1A1F31F61332E37EA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings." figureDoi="10.3897/zookeys.1226.140386.figure3" httpUri="https://binary.pensoft.net/fig/1244339">Figs 3 b</figureCitation>
,
<figureCitation id="63C62B3D757F46AA47C7F8C568117E2E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7</figureCitation>
,
<figureCitation id="656BEF4893AD2CA3C9384EB7757AC2A8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci." figureDoi="10.3897/zookeys.1226.140386.figure8" httpUri="https://binary.pensoft.net/fig/1244344">8 b, d, f</figureCitation>
,
<figureCitation id="44D56466F667C079656787EF18080C4E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gonarcal complex of Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a posterolateral b lateral c frontolateral d posteroventral e dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes." figureDoi="10.3897/zookeys.1226.140386.figure9" httpUri="https://binary.pensoft.net/fig/1244345">9</figureCitation>
,
<figureCitation id="5024FB6E7A10A96406599639698CE4EE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976)." figureDoi="10.3897/zookeys.1226.140386.figure10" httpUri="https://binary.pensoft.net/fig/1244346">10 e</figureCitation>
,
<figureCitation id="A9A95FB273C1CA941FB6FDB47AC3BEA1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba." figureDoi="10.3897/zookeys.1226.140386.figure11" httpUri="https://binary.pensoft.net/fig/1244347">11 e</figureCitation>
,
<figureCitation id="1A1AE3ED33ADCACF85A20F1836FD5DBE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species." figureDoi="10.3897/zookeys.1226.140386.figure12" httpUri="https://binary.pensoft.net/fig/1244348">12 e</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPJAE" type="etymology">
<paragraph id="571D0BC8C53B08847B4F2AFA4AEC93CB">
<heading id="2FE9FCC518F4DBFEC83FDDAE8DD7107B" reason="title">Etymology of name.</heading>
</paragraph>
<paragraph id="ACF2A8E1FEA487697A691064333FD4DD">The species is named in honor of Hélène Legrand, wife of Jean Legrand (see Dedication above). The species name is her given name in apposition (restrictive) to the masculine genus name.</paragraph>
</subSubSection>
<subSubSection id="SECID0EUJAE" type="Type specimen">
<paragraph id="4AF464AE583995465434DB81EF16062C">
<heading id="DBCDD8D39856ED929E0A1AF7428A0453" reason="title">
<typeStatus id="69423D4746462D7A6F824C2BEC16C9B1">Type</typeStatus>
specimen.
</heading>
</paragraph>
<paragraph id="F250875F9B9B14B860ED05BA6B851256">
<emphasis id="157333737F92D7588F00DEF91CDBDBE4" bold="true">
(sex and figures; verbatim label data [specimen location])
<emphasis id="D7856A270EB75DC37CBCA7549D90EDDF" bold="true" italics="true">
<typeStatus id="1412867D22DDDBE3B24527473A7B2A23" type="holotype">Holotype</typeStatus>
</emphasis>
</emphasis>
<materialsCitation id="2BE55428F3B780187C5AE664CFD1252C" collectionCode="CAS" collectorName="F. H. Walz &amp; Phillip A. Adams" country="Bolivia" location="Cochabamba" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
Male (Figs
<figureCitation id="F061A62432A13FF152CF85D9AAE6B9D0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings." figureDoi="10.3897/zookeys.1226.140386.figure3" httpUri="https://binary.pensoft.net/fig/1244339">3 b</figureCitation>
,
<figureCitation id="D943A0230457DC03F8008A6FB151F2B3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7</figureCitation>
<figureCitation id="5821EFD4D57446BAF2D82E8225A1EAFD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gonarcal complex of Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a posterolateral b lateral c frontolateral d posteroventral e dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes." figureDoi="10.3897/zookeys.1226.140386.figure9" httpUri="https://binary.pensoft.net/fig/1244345">9</figureCitation>
) (1)
<collectingCountry id="358CB171FA40A080322A37C687DBA6B9" name="Bolivia">Bolivia</collectingCountry>
/
<collectingRegion id="8F48C27F2798D2CAD722CB972B684016" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
/ Alto Palmar, XI- 60 (2) Bought /
<collectorName id="7F3D24230377684314A18637C2413000">F. H. Walz</collectorName>
(3)
<collectorName id="2F09684D755294CB0E64166F29E880F4">Phillip A. Adams</collectorName>
/ Collection / 1998 bequest to / Calif. Acad. Sci. (4)
<taxonomicName id="AF5127447D67097820444915BEE1BD78" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Gonzaga" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="genus" status="n. sp.">
<emphasis id="C7281C5D51E0570C4BF6A149886AED86" italics="true">Gonzaga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28BDB2834DAFE7C0052294A52AEE0D82" rank="species">n. sp.</taxonomicNameLabel>
; (5) Photos 4-23 - 79 [
<named-content id="11B314C1F1142915895FFA54EFB3E18B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">
<collectionCode id="3370A5750B9D48FA821229D8391B82ED" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</named-content>
]. No other specimens
</materialsCitation>
.
</paragraph>
<caption id="19AED49A43B7DB49EB917A5CCEB48533" ID-DOI="10.3897/zookeys.1226.140386.figure8" ID-arpha="384C7C0F-5919-5F14-A893-B3BDA04D33DB" httpUri="https://binary.pensoft.net/fig/1244344" startId="F8">
<paragraph id="89710B9B4D4DE69D497A8DF691E74B73">
<label id="F3DEE0EAB4DC8350F389A9290B9F1ADD">Figure 8.</label>
</paragraph>
<paragraph id="1E43DFF5B9A092BD982EC2021FAB81F6">
Side-by-side comparisons of male external abdominal features
<emphasis id="6E3E8F58B29744D3B255FB7A194BEC0E" bold="true">a, c, e</emphasis>
<taxonomicName id="55143B8834DAEE640FDAC3411E1C15EF" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="4E9AB0E10370A5193CB8228A52F40EA1" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200E77968EA2B0254D77EC83A1AC084" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Nicaragua, Río San Juan, Bartola (
<named-content id="F3CE677252FD9264997F14B4002FC27B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="0B38A3C3EB4F1C2A3950E4CC871CE40D" bold="true">b, d, f</emphasis>
<taxonomicName id="30A91E418F0F3F6ACDC2BF8CF22A4136" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="helene" status="sp. nov.">
<emphasis id="8C5BD39BC5164585E7DE57BBBB5C3834" italics="true">Santocellus helene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8C7AB57D98FB1A94F6B3361DAAFAF62" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Bolivia, Cochabamba, Alto Palmar (
<named-content id="AE24FE036FBF62A76EB4AB5F47B3343F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="67AD05DCC2F9CE16C28DF30C227EB889" bold="true">a, b</emphasis>
abdominal segments A 6 - terminus, lateral
<emphasis id="BE611BB983895276F832D2A0EF5C4B26" bold="true">c, d</emphasis>
terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae
<emphasis id="9D7356905FC11A01218F2D4FA942582E" bold="true">e, f</emphasis>
callus cerci.
</paragraph>
</caption>
<caption id="9C5AE19C67F61FF685C496594584E271" ID-DOI="10.3897/zookeys.1226.140386.figure9" ID-arpha="93E63636-7959-5E25-8937-6090DB969722" httpUri="https://binary.pensoft.net/fig/1244345" startId="F9">
<paragraph id="3F7C24D51AAC4E4E02B268ABBB91E9A9">
<label id="E166A151C80688AC242EB0BB99BE7963">Figure 9.</label>
</paragraph>
<paragraph id="50C5B8A447D676AA6B2DDC3E933CC499">
Gonarcal complex of
<taxonomicName id="96218A5BD03F9DC12450E9CBB4F0162C" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="helene" status="sp. nov.">
<emphasis id="E02169406F769F608EFE0BD6D6B0D21B" italics="true">Santocellus helene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C43B97565C7B2F00AAFF1571CDC1405" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Bolivia, Cochabamba, Alto Palmar (
<named-content id="6042FE4BE23897B4DC4A4B7CC213F411" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="B3122681DED70D04E12514AA908D338D" bold="true">a</emphasis>
posterolateral
<emphasis id="3E68100670BF20323C27588A1779F9EA" bold="true">b</emphasis>
lateral
<emphasis id="9635871A0159B705B30645677C6E8B46" bold="true">c</emphasis>
frontolateral
<emphasis id="8D5D42BFD0D0541209C6224114ADA9FC" bold="true">d</emphasis>
posteroventral
<emphasis id="203339903512B7BDE59A7B62AEAE41FE" bold="true">e</emphasis>
dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQNAE" type="description">
<paragraph id="14AB1448A252E85AC3D0FE14232426BD">
<heading id="8D49421C2BB06A13E92106F88E17CBF5" reason="title">
Description (based only on male
<typeStatus id="BB3D93D2703A842E6956909865FCB83F" type="holotype">holotype</typeStatus>
).
</heading>
</paragraph>
<paragraph id="7276F4959C1C17937178D2A0BAE7D09E">
<emphasis id="210E0285A2DC6654D6B3B838714B553A" bold="true">Adult.</emphasis>
Medium-sized.
<emphasis id="998CD0152DD1A0276611EAE0986D3EED" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="93374E8AEE12BF4FF28606DF0E54CBE4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7 a d</figureCitation>
):
<quantity id="F8EAE52B185043CB252B9EEC6406284E" metricMagnitude="-3" metricUnit="m" metricValue="1.81" unit="mm" value="1.81">1.81 mm</quantity>
wide (dorsum, including eyes); ratio of head width to eye width = 1.94: 1; sutures not visible. Vertex raised, with small posterior fold. Antenna
<quantity id="200E96C6857D9E807F4CE2AA29574FEC" metricMagnitude="-2" metricUnit="m" metricValue="2.36" unit="mm" value="23.6">23.6 mm</quantity>
long (~ 1.2 times length of forewing); scapes prominent, approximately as long as broad (
<quantity id="0ACFE303A6049742F2679A30072C7D70" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.170.18 mm</quantity>
). Pedicel round, tubular, not modified. Torulus relatively large, extending well below mid-eye level. Frons short to moderate length, with pale setae, lower margin scalloped upward above clypeus (below toruli). Tentorial pits distinct, round, raised. Gena
<quantity id="A817A80EC6218EC7BB0DF353923CBDF5" metricMagnitude="-4" metricUnit="m" metricValue="2.1" unit="mm" value="0.21">0.21 mm</quantity>
long; ratio of genal length to distance between tentorial pits = 0.47: 1. Clypeus quadrate, except dorsal margin broadly curved upward below frons, distal margin relatively straight, pale mesally; surface rounded, slightly raised mesally, with sparse, amber setae of medium length. Labrum tapering anteriorly, with bilobed frontal margin, distinct cleft between lobes moderately deep, lateral margins tapering, long setae basally, shorter setae distally. Labium with submentum slender, rounded ventrally, with round, slightly bulbous tip.
<emphasis id="72CC81BB5A0EC5816CF9EC4D1101FC7F" bold="true" italics="true">Coloration</emphasis>
: Scapes with frontal surface pale, mesal tip with dark mark extending onto mesal base of pedicel; pedicel tan throughout; flagellum largely pale, with basal three segments, distal segments dark brown to black. Lateral margins of vertex with pair of dark brown, curvy longitudinal bands, forming cream-colored circular spot centrally, each band with pale spot adjacent to circular spot, thus forming transverse row of three round cream-colored spots. Frons heavily marked with dark brown to reddish black distally, with broad, dark brown, scalloped band along margin above clypeus, laterally extending almost to mesal margins of eyes, mesally extending very short distance between scapes; area below scapes cream-colored, without markings. Clypeus with slender stripe across midsection, not reaching lateral margins; labrum with slight brownish tinge mesally. Clypeus with dark brown marks laterally, blending into dark distal marks on gena. Venter of head pale, pair of small brown spots near base of mentum and eyes; distal three maxillary palpomeres dark brown to black, basal two probably pale; distal labial palpomere black, basal two pale.
</paragraph>
<paragraph id="4F0A6262128A1AA4A4EEEF3F61524956">
<emphasis id="912AB15C28B6952DD8B29824940D2224" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="213621DA4CDFA66AE589C5A43CD8FF47" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7 a, b, e</figureCitation>
): Prothorax approximately two times wider than long:
<quantity id="9D04434074851BC0C33649989E03A13C" metricMagnitude="-4" metricUnit="m" metricValue="6.4" unit="mm" value="0.64">0.64 mm</quantity>
long,
<quantity id="98F64093C683F0C0D9D2D47BB27F6885" metricMagnitude="-4" metricUnit="m" metricValue="3.1" unit="mm" value="0.31">0.31 mm</quantity>
wide; ratio of length to width = 0.48: 1. Surface with fine, golden setae, most short, some long to very long. Legs slender, with short to medium length, robust, setae, few long setae; tips of tarsi smooth, shiny (black) dorsally, with pair of smooth, small, swellings at terminus (Fig.
<figureCitation id="24F62546CF851FBF749F21D28B54FB31" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7 e</figureCitation>
). Tarsal claw moderately long, narrow, base without dilation but with small mesal spur, two pairs of very long setae distally.
<emphasis id="BC3D94F4ABBB8B6571E830D87D9F27DF" bold="true" italics="true">Coloration</emphasis>
: [Note: cream- or tan-colored areas described here probably green in life.] Three thoracic segments mostly dark brown; pronotum with margins dark brown to black, center cream-colored, mottled with ~ six large dark brown to black patches; venter, pleural region cream to light tan. Mesonotum, metanotum with paired cream and dark brown patches interspersed in roundish pattern. Legs pale, mostly unmarked, with setae mostly golden, few longer setae pale; surface of terminal segment (including base of tarsal claws and pair of small terminal swellings) shiny, black (Fig.
<figureCitation id="F41CA86E7D32EE94BE92C22FDDAD5165" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7 e</figureCitation>
). Tarsal claw moderately long, narrow, with small spur; base slightly enlarged, without distinct dilation.
</paragraph>
<paragraph id="23FFD29C3980564FEE7D05384EF6D2C1">
<emphasis id="A701A78581F13970B9B7FBED4831BFC3" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="4955ABC2EF570EF06647F781EB8226BA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings." figureDoi="10.3897/zookeys.1226.140386.figure3" httpUri="https://binary.pensoft.net/fig/1244339">3 b</figureCitation>
): Forewing (left)
<quantity id="BFCEB1EB61E2A1923A6881A6A20463E3" metricMagnitude="-2" metricUnit="m" metricValue="1.94" unit="mm" value="19.4">19.4 mm</quantity>
long,
<quantity id="C54F3B66C39BFEAB5A14A7748A1F57B0" metricMagnitude="-3" metricUnit="m" metricValue="7.2" unit="mm" value="7.2">7.2 mm</quantity>
wide (maximum). Costal area moderately tall; tallest costal cell (# 10)
<quantity id="6C74E43357E2BA5FBA65F02EA1249DD6" metricMagnitude="-3" metricUnit="m" metricValue="1.7" unit="mm" value="1.7">1.7 mm</quantity>
tall, 0.24 times maximum height of wing. First intramedian cell triangular, stout,
<quantity id="160DC3257249CC745C868EE5A9EAD97D" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
wide, ~ 0.6 times width of third median cell. Origin of first radial crossvein ~
<quantity id="971D42256A0DF3B8031545D3B5C0D16A" metricMagnitude="-4" metricUnit="m" metricValue="3.2" unit="mm" value="0.32">0.32 mm</quantity>
distal to origin of Rs; radial area (between
<collectionCode id="C719CDD00A8F35713281A948F850A624" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
and Rs) with single row of 14 closed cells; tallest radial cell 2.9 times taller than wide. Three b cells, each full sized, slightly longer than tall; first b cell heavily marked on all sides. Two series of gradates; 8 inner gradates, 9 outer gradates; size of cells bounded by gradates fairly uniform, lengthening slightly distally. Four b cells (beneath Psm, after im 2); last b cell wider than tall. Three intracubital cells, distal one open; first cubital crossvein located basal to second mediocubital crossvein; icu 1, icu 2 each shorter than icu 3; icu 1 + icu 2 longer than icu 3. Vein 1 A forked. Terminal margin with approximately 5 simple veinlets, 13 forked veinlets.
</paragraph>
<paragraph id="7A8F9569E124D7A4E6296010E2A0B60A">
Hindwing:
<quantity id="D28B332693AA5936DF9BECAF0E3A17AB" metricMagnitude="-2" metricUnit="m" metricValue="1.72" unit="mm" value="17.2">17.2 mm</quantity>
long,
<quantity id="FAA8937695D57D418BD6D119AC6B4792" metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
wide; 12 radial cells (counted from origin of
<collectionCode id="8F3568207E6390F4587176E90D631EAC" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, not false origin); t cell absent; first b cell beneath RP relatively small, quadrate, heavily marked. Two series of gradates; 6 inner gradates, 8 outer gradates. Three b cells; 4 b cells beyond im 2; 3 intracubital cells, distal one open. Terminal margin with 5 simple basal veinlets, 12 forked basal veinlets.
<emphasis id="53096FDC27509732EFF45080150EFD90" bold="true" italics="true">Coloration</emphasis>
: Wings hyaline, with bold dark brown markings. Forewing: stigma with large dark-brown spot basally, second smaller dark-brown spot distally; veins pale except in marks; penultimate sections of
<collectionCode id="E34F411F5561E0EAB1403E30A113665C" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, Rs, including
<collectionCode id="20542266C330DB337F8B7AD63F56FC7A" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
Rs crossveins dark brown, surrounded by dark brown suffusion; slightly elongate dark brown mark at tip of wing, dark streak across center of wing, through inner gradate veins; dark-brown streak from base of Rs to margin of wing; scattered dark-brown marks along lower basal half of wing edge. Hindwing: stigma with one large dark-brown mark; base of RP, t cell absent; first b cell small, quadrate with dark-brown mark; distal section of cu (on posterior margin of wing) dark brown. Terminal margin with 4 simple veinlets, ~ 14 forked veinlets.
</paragraph>
<paragraph id="EB7A20630DAB1443327BCF71F19EB788">
<emphasis id="EDF99E4544E4AC0903B4384D18848E37" bold="true" italics="true">Male abdomen</emphasis>
(Figs
<figureCitation id="4D3F3617363DA8804677CFF843BE4B7D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci." figureDoi="10.3897/zookeys.1226.140386.figure8" httpUri="https://binary.pensoft.net/fig/1244344">8</figureCitation>
,
<figureCitation id="05D2710803B4A596EAE155E9B92DFCCC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Gonarcal complex of Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a posterolateral b lateral c frontolateral d posteroventral e dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes." figureDoi="10.3897/zookeys.1226.140386.figure9" httpUri="https://binary.pensoft.net/fig/1244345">9</figureCitation>
, slightly teneral). Segments 13 similar in size, proportion to distal segments, only slightly larger than basal segments. Fifth, sixth tergites (
<collectionCode id="63657966F6A9EA7472A3A324F362683C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
5,
<collectionCode id="83197EC629931F0389202E8CF44DFCF6" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
6) slightly taller than seventh tergite (
<collectionCode id="AED9B49C30DC0F78AD0D2116C0D65FFB" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
7). Tergites, sclerites, pleural region with numerous, short to medium length setae, microsetae; spiracles not enlarged; microtholi absent. Abdominal sternites moderately tall (S 6: ratio of length to width = 1.2: 1), with dorsal margins straight, slanting; distal sternites with slightly long, slender setae throughout. Callus cerci almost round to slightly ovate, 0.23
<quantity id="015CC0E5FAC9AA4441DB2B9B47FDBE75" metricMagnitude="-4" metricUnit="m" metricValue="1.6" unit="mm" value="0.16">0.16 mm</quantity>
diameter, with ~ 31 long trichobothria; cupuliform bases of variable size [those within central section larger (~
<quantity id="7250D189B72C4E981BBC9D16E433653C" metricMagnitude="-5" metricUnit="m" metricValue="2.8" unit="mm" value="0.028">0.028 mm</quantity>
) than those on periphery (~
<quantity id="95E73B766833168850FFC699402321F8" metricMagnitude="-5" metricUnit="m" metricValue="1.5" unit="mm" value="0.015">0.015 mm</quantity>
)]. Dorsum of
<collectionCode id="7F5A68B55E48EA2B232EAF334D02104E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct rounded distally, fused mesally with very small to no cleft; no midline suture visible; with long setae, densest along distal margin; dorsal apodeme straight, extending along full length of
<collectionCode id="18159EFE74E04708AF350C713661AD6D" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct, above upper side of callus cerci, with large ventral spur extending downward along basal margin of callus cerci to posterodistal margin of ectoproct. S 8 + 9 fused, apparently without suture, tapering to acute apex (lateral view); S 9 slightly darker than S 8; ventral apodeme extending across dorsal surface of S 8 + 9; S 9 spatulate distally, with terminal membrane prominent, attached across full margin of sternite; membrane bearing single, irregular row of large robust gonocristae laterally, smaller, denser gonocristae mesally.
</paragraph>
<paragraph id="036388C3429E817B975111D8642B97FA">
<emphasis id="AB3912F0A50E0E315941E1BE357E1E7A" bold="true">
<emphasis id="ED3C3D552F3DF47ACF34A750E529A728" bold="true" italics="true">Male genitalia</emphasis>
.
</emphasis>
Tignum, gonapsis, parameres, quadrate hood, pseudopenis, spinellae absent. Gonarcus broad, with gonarcal bridge bearing pair of elongate gonocornua fused laterally forming dome-like plate above mediuncus. Gonarcal arms large, rounded, expanded slightly forward. Mediuncus elongate, in two sections: basal section extending from below gonarcal bridge, with pair of elongate, rod-like internal apodemes covered dorsally by heavy subanal membrane; distal section of mediuncus extending distally from basal section at acute angle, with pair of flange-like arms lateral to mesal beak; gonosaccus beneath mediuncus, stiff, flat laterally, with few gonosetae.
</paragraph>
<paragraph id="D3225B0C1EBD35091EB9205C3527B094">
<emphasis id="BBC6D3F7757DB35EF5A4A4969675BA45" bold="true">
<emphasis id="24C87B363B6259603219BAF8011D7155" bold="true" italics="true">Female abdomen and genitalia</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFQAE" type="variation">
<paragraph id="67BB78E0C017CDAF14A07BA0D4AF9DC3">
<heading id="0E6F9D4D316BA365473050E366184187" reason="title">Variation.</heading>
</paragraph>
<paragraph id="DDCF139CADB99A3E6BEE5EF7233B5896">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EKQAE" type="distribution">
<paragraph id="860FF5910DACA607633B31F598BBC965">
<heading id="47808325AADE87C2D29628F46FA19562" reason="title">Known distribution.</heading>
</paragraph>
<paragraph id="663839B482260F8B52D1CF11FE0AF4DA">
South America.
<collectingCountry id="57F70031800C1FBC5200D58AEFEA9864" name="Bolivia">Bolivia</collectingCountry>
(
<collectingRegion id="C3470CBF701CEC0C56700F9900257720" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,355 @@
<document id="1041AEDA3C2DE707C7E39A4E6AD95A07" ID-DOI="10.3897/afrinvertebr.66.129611" ID-ZooBank="9A70A477-D862-40E5-A132-3227D96C3ECC" ID-publisher-id="129611" URI-arpha="5630D620-B234-5FD1-B5C0-EDF357AAF7CA" URI-zoobank="http://zoobank.org/9A70A477-D862-40E5-A132-3227D96C3ECC" article-type="research-article" checkinTime="1738952593943" checkinUser="pensoft" docAuthor="Rollinson, Lisa &amp; Cabrero, Allan" docDate="2025" docId="A1249FE843185A939BA00B5DF7CBE0B4" docLanguage="en" docName="African Invertebrates 66(1): 73-115" docOrigin="African Invertebrates 66 (1)" docSource="https://africaninvertebrates.pensoft.net/article/129611/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Enica sporanthera Rollinson &amp; Cabrero 2025, comb. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="115" masterDocId="5630D620B2345FD1B5C0EDF357AAF7CA" masterDocTitle="Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834" masterLastPageNumber="115" masterPageNumber="73" pageNumber="73" updateTime="1738953078235" updateUser="ExternalLinkService">
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<mods:title id="DB9AC6FAB3E14C93FC03564F0A289C1B">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
</mods:titleInfo>
<mods:name id="2AB620B928D2C6F999788E4525AF2F45" type="personal">
<mods:role id="7A523AD79B6DA56C126FFE1ED30E08E6">
<mods:roleTerm id="C16030D7E637768F5C455431070C77DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F2BECE6C8A0A9FB9754253EE088847A">Rollinson, Lisa</mods:namePart>
<mods:nameIdentifier id="C7ABA7B36A70258B3BA46BF230F41E60" type="ORCID">https://orcid.org/0009-0007-0152-7073</mods:nameIdentifier>
<mods:affiliation id="AC26B11B0E4F0CAD847BAF85BB93AEDD">Texas A &amp; M University, College Station, Texas, USA</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="D02FE52A3B39D74082473D6ADFCC8D74">Cabrero, Allan</mods:namePart>
<mods:nameIdentifier id="B758752D76BE50ADCC1E4391FB59052E" type="ORCID">0000-0002-2935-3534</mods:nameIdentifier>
<mods:affiliation id="38D7A0F043979E2A0E0DBB9A3A92C45A">National Museum of Natural History, Washington, D. C., USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="463D2847EA86D4C05F780DAF33838454">text</mods:typeOfResource>
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<mods:titleInfo id="E5BE2562E1821E06FA1504634C89A94D">
<mods:title id="69FB3F778A1DB304E2968B624D6D07D5">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="84D29CDCEC09C1BAF3B989969A048381">
<mods:date id="91A4C3B817F9D073AFB1290EB9F6198E">2025</mods:date>
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<mods:number id="F53C22D7D2E297D074353CE8CE42EF24">2025-02-07</mods:number>
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<mods:number id="108D08D0D2F080EB08E441BBE14E37A3">66</mods:number>
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</mods:extent>
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<mods:classification id="FC837A034A6603F61D6EC7D36633169C">journal article</mods:classification>
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</mods:mods>
<treatment id="A1249FE843185A939BA00B5DF7CBE0B4" ID-DOI="http://doi.org/10.5281/zenodo.14834498" ID-Zenodo-Dep="14834498" ID-arpha="A1249FE8-4318-5A93-9BA0-0B5DF7CBE0B4" LSID="urn:lsid:plazi:treatment:A1249FE843185A939BA00B5DF7CBE0B4" httpUri="http://treatment.plazi.org/id/A1249FE843185A939BA00B5DF7CBE0B4">
<subSubSection id="C3B4393388E75D9F8C0661A895EFAC58" type="nomenclature">
<paragraph id="CB6A6B7E9C894766D50BA5F32EA48A1A">
<taxonomicName id="A00E40DB45E8AA0C1421B462E7DC5004" authority="(Hesse, 1956)" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sporanthera" status="comb. nov.">
<emphasis id="5D7758BA7ACE9BB601D1C1433B36E770" italics="true">Enica sporanthera</emphasis>
(
<bibRefCitation id="4ED64DDBCCDA553D017C835392E4A1C1" author="Hesse AJ" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 27-46. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="5C715C4A6BE6CD9B8E52AB17303ACA46" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FA172706A68CFCF71FB1D5349547BADD" type="description">
<paragraph id="77792A3EA3670B84DB57609D6B616576">
<figureCitation id="B228EBE7E369D7AB2FC16E53674544F2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Enica sporanthera ♂ SAM-DIP-A 002174 (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica syrticola ♂ NMSA - DIP- 89516 (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure11" httpUri="https://binary.pensoft.net/fig/1244184">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERWDI" type="Taxon depository">
<paragraph id="995C3DDD1E4EF06D468CC1E3F12BCB5B">
<heading id="567022ADB60F00E7CC1C6AF45B67DCA2" reason="title">Taxon depository.</heading>
</paragraph>
<paragraph id="CB8233087A34907C9FCB66582AABF9C0">
ZooBank:
<ext-link id="E36F6B63E531C3CB09473FAF24B49409" ext-link-type="uri" xlink_href="https://zoobank.org/12C60B29-A951-413E-9E8A-6E73C7434B7A" xlink_type="simple">
https://zoobank.org/
<uuid id="FE548503AEFB4F1EA7622B932B76C56B">12C60B29-A951-413E-9E8A-6E73C7434B7A</uuid>
</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3WDI" type="diagnosis">
<paragraph id="743C2CB509637063631D3EF2EAC310DD">
<heading id="A6F91C2C522AB021D27ED2373E842606" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="40C0B204CAF477431E545FD1FE788236">The species is distinguished from other species in the genus by the elongated black spot on the dorsal occiput in both males and females, which extends across the head.</paragraph>
<caption id="A22F55ED527331795C73D9B74FDB2E38" ID-DOI="10.3897/afrinvertebr.66.129611.figure11" ID-arpha="228898C8-FD75-5391-91E1-E3243F942C57" httpUri="https://binary.pensoft.net/fig/1244184" startId="F11">
<paragraph id="B34DA0E252E7356E07FB36DDFB61033A">
<label id="9C819CE2DE07FA0A60C5CFF98ED8E694">Figure 11.</label>
</paragraph>
<paragraph id="5100EDF9000B8D71E43C0520E63BFCE2">
<taxonomicName id="289B8ACF570C6B9B701C77F79F8F34C5" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sporanthera">
<emphasis id="813AD7870C1D771ED8D7F9B5051CB0F4" italics="true">Enica sporanthera</emphasis>
</taxonomicName>
♂ SAM-DIP-A 002174 (
<emphasis id="5820F7877CA4678609B6B507FC310A77" bold="true">A C</emphasis>
)
<emphasis id="D34C6D4EC480451BE6FF4D2036B9B611" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="A5321130A4FFF7DD1C6A42B4DE0B5D12" bold="true">B</emphasis>
habitus, lateral view
<emphasis id="BB143D7CEBA75C08C890C375A643AB7F" bold="true">C</emphasis>
habitus, frontal view;
<taxonomicName id="6221BEC265660BCE24B005CE1FF7D55E" authority="♂ NMSA" authorityName="NMSA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="syrticola">
<emphasis id="CF0272BFA7C0D317F4B2ACE5443B9CA9" italics="true">Enica syrticola</emphasis>
<named-content id="3CC27EB45140619CA2FE0E63C4157351" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
</taxonomicName>
- DIP- 89516 (
<emphasis id="54790AEB5972B79F754DE2F8E17A6522" bold="true">D F</emphasis>
)
<emphasis id="A7D01F63386C6904DBB23116739AC11C" bold="true">D</emphasis>
habitus, frontal view
<emphasis id="9AD5328D8613F6AF00C68B724BE983AF" bold="true">E</emphasis>
habitus, dorsal view
<emphasis id="0EF8916D4B8ECD272828E1A00484ED14" bold="true">F</emphasis>
habitus, lateral view. Scale bars: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EYYDI" type="description">
<paragraph id="42D04B5F4936BCCE1FA696CD00527AC9">
<heading id="2600AD61E9CF713DECAF4ED19E2723B7" reason="title">Description.</heading>
</paragraph>
<paragraph id="84BDF377DCC252490D5EFAA201EF50C6">
<emphasis id="BEA3565F2AA2ECCCB42B7F2896E5C082" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: similar to head, slightly darker than head, without dark spot, males: similar to head, slightly darker than head; frons black setose, with white scales, males no setation near eyes, females setose near eyes only dorsally; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput collar of dense snow white setose, white scales antero-dorsally, dorsal eye margin with dark brown to black spots present, touching or almost touching eye margin, never extending ventrally past level of antennae; gena creamy white, extremely sparsely setose.
</paragraph>
<paragraph id="93C664B104286AACD034854A28AE882F">
<emphasis id="8C9B47BFF9FA88B58F1671A124D38FEB" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, light brown to brown, light yellowish-brown setose.
</paragraph>
<paragraph id="279B46F603F0E8F9DA8C58D1F7EDC435">
<emphasis id="F6DF801A5DD57F97A529435250951B93" bold="true" italics="true">Antenna</emphasis>
: dark brown; scape creamy white, approximately as long as pedicel, black setose dorsally and ventrally, setae short, not extending to end of pedicel; pedicel brown or creamy white, short black setose dorsally; postpedicel dark brown, tapering distally, longer than scape and pedicel combined, white or light yellow scales dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="25A64604CC4F2F2AAA3755FD367C3DB8">
<emphasis id="ADB3DDAD6E4C45749FD77FAD93803ACE" bold="true" italics="true">Thorax</emphasis>
: light orangish brown; scutum predominantly black, reddish on margins, grey to brown scales centrally and white scales peripherally; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, white scales dorsally; metepimeron silvery grey pubescence, asetose; scutellum red, anterior black spot, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of white scales, black setose.
</paragraph>
<paragraph id="BA25ED02FDD334A98A510946F6204B92">
<emphasis id="CDEFF1DFCFCA2F472E214B1E7028356A" bold="true" italics="true">Leg</emphasis>
: light orangish brown, covered in scales, some regular setae present; pro coxa light orangish brown to brown, silvery grey pubescence, white scales, thin white setose, thick black setose; pro femur dorsally dark brown, ventrally orangish brown, white scales ventrally, brown scales dorsally, few short black setae; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa light orangish brown, white pubescence, white scales, thin white setose, thick black setose; mes femur dorsally dark brown, ventrally orangish brown, white scales ventrally, brown scales dorsally, few short black setae; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa light orangish brown, white pubescent, white scales, thin white setose, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur dorsally dark brown, ventrally orangish brown, white scales ventrally, brown scales dorsally, few short black setae; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="451611210DCE68C744DDFCC883031337">
<emphasis id="5FCF4E2F89FDA124AACE070CA9BABBDA" bold="true" italics="true">Wing</emphasis>
:
<quantity id="E14DDE14CD2E2527167F38067D70FDFF" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="8.9" metricValueMin="8.2" unit="mm" value="8.55" valueMax="8.9" valueMin="8.2">8.28.9 mm</quantity>
, microtrichia absent; membrane color pattern males: minimal spots of infuscation in r 2
<subScript id="2262242BA48EF56D8EE7BA1877E72567">+ 3</subScript>
, br, and bm, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="5949572EF677BCC678340941C9B2DC5D">+ 3</subScript>
, base of m
<subScript id="A5A6B785FDE00BB20206729D11D3F943">2</subScript>
, m-cu, split between M
<subScript id="127F626BD5BA0E25C6EC4FABDF742EB3">1</subScript>
+ 2 and M
<subScript id="BBE6EE1F76357841809659F6744D20FE">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="77923A3C5735EF073C09C3D9B5742C5F">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="A75F0CD3DCEAAF96074D7B20F4592310">1</subScript>
open; R
<subScript id="E107FC46C4E2C023C65447349C98A2E3">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="27D84FD819FDE22E9980C3D583F65DE3">4</subScript>
open, proximally R
<subScript id="D446359C3C3BB4E6F0A50E948F0DDCCB">4</subScript>
and R
<subScript id="FF0B42D3394591FC061175FFB624C478">5</subScript>
parallel, R
<subScript id="7102FE693BC943F407BE0ABEE906A099">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="6E6284B797A519ADC9A45CD53C9C3EF6">5</subScript>
terminating posterior to wing apex; r
<subScript id="EA244E7B23B70715B03C27B2D1AE6C2C">5</subScript>
open; M
<subScript id="4D65EE42A92418ABE2B78151844C9060">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="F9B7E3FC2F3405413779B22441FA270A">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="0C684D78DC4A2213D87D4D8C9A013E62">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.
</paragraph>
<paragraph id="4D44E5C32294CFC65303EDF48AE40676">
<emphasis id="9CAFE7B6A6C594569E02306F5512EFDF" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, predominantly black anteriorly and red posteriorly; tergites smooth, setae with small sockets only; T 1 reddish yellow to light brown, light yellowish-brown to light whitish-brown setose, with scales; T 2 T 7 predominantly reddish-orange, thin black setose, brown to light yellowish-brown and white scales, white scales on spots on posterior margins of T 2, rarely T 3, and T 4, single spot medially on T 2; S 18 predominantly light reddish-orange, yellow to white scales, thin black setose and long white setose; lateral margin many black setae extending away from abdomen, male T 8 white scales, black setose.
</paragraph>
<paragraph id="FB53C74E0AF53D3B334275ACEFD87151">
<emphasis id="F748CDA18CD89F420C634A6A00A40FBB" bold="true" italics="true">Terminalia</emphasis>
: Male terminalia drawings in
<bibRefCitation id="31D25A71B3A9340D00463C4C8FF5F0FD" author="Hesse" firstAuthor="Hesse" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 2746. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse (1956</bibRefCitation>
, p 39):
<ext-link id="41AA9D5D4EA982B12332E002315E25A1" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/40844174" xlink_type="simple">https://www.biodiversitylibrary.org/page/40844174</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E62DI" type="material">
<paragraph id="533E77BDC7952804E95B18F0435EFAE0">
<heading id="5B90C7526AAFBA213F9353E35D0E55C7" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="C6F44F3364559EF3A47833E7BBBEB701">
<materialsCitation id="62352699C736F432923D097390F08089" collectingDate="1931-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-30.150833" location="Bowesdorp Namaqualand" longLatPrecision="20" longitude="17.931944" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="F819B4B7AA74E2751CA38ADBD63BF3D3" bold="true">
<emphasis id="78417986B7F63EE6172784F7CFFEED9B" bold="true" italics="true">
<typeStatus id="411C6EA8366CB48BEAE626674D0142D7" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="F4BCD691A967FAE7E08EB33A596AF464" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="0D59C5A1AC0F33E0990489BC5848DE07" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="BC202178EDE0F34BEF540F4A580880B2" count="1" type="male">1 ♂</specimenCount>
<location id="4F85668101E136E4264C4EBCC6B732AC" LSID="urn:lsid:plazi:treatment:A1249FE843185A939BA00B5DF7CBE0B4:4F85668101E136E4264C4EBCC6B732AC" country="South Africa" latitude="-30.150833" longLatPrecision="20" longitude="17.931944" name="Bowesdorp Namaqualand" stateProvince="Northern Cape">Bowesdorp Namaqualand</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EN3DI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.931944,-30.150833]}">
<geoCoordinate id="8D84934049B12A310AFC5271ED125761" degrees="30" direction="south" minutes="09" orientation="latitude" precision="15" seconds="03" value="-30.150833">30 ° 09 ' 03 &quot; S</geoCoordinate>
,
<geoCoordinate id="F731A4613D4903E241A789D0AED42D40" degrees="017" direction="east" minutes="55" orientation="longitude" precision="15" seconds="55" value="17.931944">017 ° 55 ' 55 &quot; E</geoCoordinate>
</named-content>
;
<date id="B29B85996FB37F3EB20189BAA3DAD72C" value="1931-11">
<collectingDate id="11481E75DEC565AAB58F830DFC91B352" value="1931-11">Nov. 1931</collectingDate>
</date>
; SAM
<collectorName id="031EFF3E549BD3BED7CC13A45877B52E">Museum Staff</collectorName>
leg.; SAM-DIP-A 002173,
<named-content id="274218AEF528FC9380FA694C5F1E14EC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="EBA78E7C63D3BDDF2737B4942663DEDD" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
.
</materialsCitation>
</paragraph>
<paragraph id="E10ADCAE7CB5B0CBB54D83342BB3AC56">
<materialsCitation id="93D8A47A56FF9CA494AAAE379634D64E" collectingDate="1931-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-30.416666" location="Garies Namaqualand" longLatPrecision="20" longitude="17.9" municipality="Klip Vlei" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="4DF46992D9A22B26314E2F76FEE0CEA0" bold="true">
<emphasis id="1A93830AA5C09B182849C13D5F87EE52" bold="true" italics="true">
<typeStatus id="52B0712D40A043001F4F047920D82BA0" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="15FD7556959979E5A3724238630C6AFD" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="0118D1194958BE578C5808315ACDCE72" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="8BCF27EA4BDF8EE67F22E1C463E873B7" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="992A8AD98E981D1C1A062DA3611318B0">Klip Vlei</collectingMunicipality>
,
<location id="4A89FF419FBF861D33B469321E1E609C" LSID="urn:lsid:plazi:treatment:A1249FE843185A939BA00B5DF7CBE0B4:4A89FF419FBF861D33B469321E1E609C" country="South Africa" latitude="-30.416666" longLatPrecision="20" longitude="17.9" municipality="Klip Vlei" name="Garies Namaqualand" stateProvince="Northern Cape">Garies Namaqualand</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EA4DI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.900000,-30.416667]}">
<geoCoordinate id="C68D13E42B211992531CAF470F578BAC" degrees="30" direction="south" minutes="25" orientation="latitude" precision="15" seconds="00" value="-30.416666">30 ° 25 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="C322BA8E271F2E066AAAE57E1D0DCADA" degrees="017" direction="east" minutes="54" orientation="longitude" precision="15" seconds="00" value="17.9">017 ° 54 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="541B443E3F3DA63EA95A1B291011DCED" value="1931-11">
<collectingDate id="67AC3189AFCA22B2EC2CB7F2C8E195EE" value="1931-11">Nov. 1931</collectingDate>
</date>
; SAM
<collectorName id="948BD768118DD53F82E824B778A18E90">Museum Staff</collectorName>
leg.; SAM-DIP-A 002174,
<named-content id="91830F8C52A9E7DC2DD0CEAA01F883D7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="D0583E39D18325E5618DFE2F36443080" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="2C77785D3CA4A2FF52D835AB5CE7A7B6">
<materialsCitation id="3DFDE035CEEAE1E0521D1F0C75A90435" collectingDate="1947-09" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.127777" location="Pakhuis Paas" longLatPrecision="20" longitude="19.041945" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<collectingCountry id="0D67B6AA149AFFABBD39977C8FAE46C4" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="F76341ECE3E1507DC6752559CE5CCC35" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="12C49C337FBE07D6F0ABAFE7EFB35197" count="1" type="male">1 ♂</specimenCount>
<location id="EAFCEAE377F45B3BEEF93618B2EC2624" LSID="urn:lsid:plazi:treatment:A1249FE843185A939BA00B5DF7CBE0B4:EAFCEAE377F45B3BEEF93618B2EC2624" country="South Africa" latitude="-32.127777" longLatPrecision="20" longitude="19.041945" name="Pakhuis Paas" stateProvince="Western Cape">Pakhuis Paas</location>
, east of;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EO4DI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.041944,-32.127778]}">
<geoCoordinate id="C1ADB38C322916E0A228F05D6E8F5BB2" degrees="32" direction="south" minutes="07" orientation="latitude" precision="15" seconds="40" value="-32.127777">32 ° 07 ' 40 &quot; S</geoCoordinate>
,
<geoCoordinate id="0278FB2200359B04888BEA8D8713112F" degrees="019" direction="east" minutes="02" orientation="longitude" precision="15" seconds="31" value="19.041945">019 ° 02 ' 31 &quot; E</geoCoordinate>
</named-content>
;
<date id="6928F6B25599F001857AD919C7D47BA2" value="1947-09">
<collectingDate id="434DEBCD3909776E005ABB539B1A4F8D" value="1947-09">Sep. 1947</collectingDate>
</date>
; SAM
<collectorName id="D3D635578E8938CD405B902F64CDADFF">Museum Staff</collectorName>
leg.; SAM-DIP-A 002175,
<named-content id="76A86AD3A2DD4B56B6C9A50421799D0C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="3E5B257A996A3A4277A032E3759F3D2C" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX4DI" type="material">
<paragraph id="5CAC08D0220DFAF1DB1BC8B8E897EFBE">
<heading id="4AFB13B42012C83ED155528CB35BF954" reason="title">Other material.</heading>
</paragraph>
<paragraph id="593F5C7AFEC2F9282600D8D38767EEFF">
<materialsCitation id="50367987D2F7CE211F37EAE0DE217B81" collectingDate="1931-11" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-30.150833" location="Bowesdorp Namaqualand" longLatPrecision="20" longitude="17.931944" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<collectingCountry id="14BA1A8D79DF729843E7DDA2D274AD3C" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="164580FD3FE5EC24A621429FC8BA9FE8" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="1C51C7AD23CEED8E8EC9F5E4B5D8593A" count="1" type="female">1 ♀</specimenCount>
<location id="4140E342C33D3969E82F0E85C5E714AE" LSID="urn:lsid:plazi:treatment:A1249FE843185A939BA00B5DF7CBE0B4:4140E342C33D3969E82F0E85C5E714AE" country="South Africa" latitude="-30.150833" longLatPrecision="20" longitude="17.931944" name="Bowesdorp Namaqualand" stateProvince="Northern Cape">Bowesdorp Namaqualand</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EA5DI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.931944,-30.150833]}">
<geoCoordinate id="0BCDBA4642A2D3B88E2A2A7B75C6EF39" degrees="30" direction="south" minutes="09" orientation="latitude" precision="15" seconds="03" value="-30.150833">30 ° 09 ' 03 &quot; S</geoCoordinate>
,
<geoCoordinate id="2D513BDD9854660D86B819F76E5591E0" degrees="017" direction="east" minutes="55" orientation="longitude" precision="15" seconds="55" value="17.931944">017 ° 55 ' 55 &quot; E</geoCoordinate>
</named-content>
;
<date id="FF96F7EE37A454D0F02EF9B1EFACA186" value="1931-11">
<collectingDate id="2D2D7F73C5607A84E8CF90CCCEC6817D" value="1931-11">Nov. 1931</collectingDate>
</date>
; SAM
<collectorName id="0282E4B42C027E741B6592A85273C340">Museum Staff</collectorName>
leg.; SAM-DIP-A 002173,
<named-content id="522479A372979FDAE5E3CCF602459EB9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="A146CAB943A61846E57093612360B768" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ5DI" type="distribution">
<paragraph id="1C897114EED77B181EF01EC6E871FA52">
<heading id="C805D94E77319F0FD169A188346BB212" reason="title">Distribution, biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="0545FD455DA7473AF78C536C5D1586FD">
Known from localities in the
<collectingRegion id="24DEF96D385FAFD043A966AA1ADDCE79" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
and
<collectingRegion id="0D8AEE33430BBA0CEB028FEF73B4AABA" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
of
<collectingCountry id="A6CCB9BEF7040568D4A9AC8235D5ED39" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="16290EBA8CF19D96D63952A60A244990" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. longirostris, E. henicoides and E. sporanthera specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure9" httpUri="https://binary.pensoft.net/fig/1244182">9</figureCitation>
). This species is known from
<specimenCount id="0D05091D8459E8036DEC3418A1BF409B" count="4" type="generic">4 specimens</specimenCount>
, collected in three collecting events spread between 19522002. The species is known to occur in the Succulent Karoo and the
<collectingRegion id="8DE7F06E386FDB60B3C639C18BFBB365" country="South Africa" name="Western Cape">Cape</collectingRegion>
Floristic Region biodiversity hotspots. Adult flies are active from September - November (Table
<tableCitation id="0EB634D89F88A480ED16C374BA77B725" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="FE082B4D3C9B85EA1D7517E93626A9E7" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="90428592A89B5E31BAF4A9D67E81D387" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="A99C7BC8C5A79926ECD0CB0756577596" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,409 @@
<document id="854B18D680AD6351825778D272A8BD09" ID-DOI="10.3897/afrinvertebr.66.129611" ID-ZooBank="9A70A477-D862-40E5-A132-3227D96C3ECC" ID-publisher-id="129611" URI-arpha="5630D620-B234-5FD1-B5C0-EDF357AAF7CA" URI-zoobank="http://zoobank.org/9A70A477-D862-40E5-A132-3227D96C3ECC" article-type="research-article" checkinTime="1738952593943" checkinUser="pensoft" docAuthor="Rollinson, Lisa &amp; Cabrero, Allan" docDate="2025" docId="A66900FAAD9A57698F0E3D7E57629120" docLanguage="en" docName="African Invertebrates 66(1): 73-115" docOrigin="African Invertebrates 66 (1)" docSource="https://africaninvertebrates.pensoft.net/article/129611/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Enica henicoides Rollinson &amp; Cabrero 2025, comb. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="115" masterDocId="5630D620B2345FD1B5C0EDF357AAF7CA" masterDocTitle="Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834" masterLastPageNumber="115" masterPageNumber="73" pageNumber="73" updateTime="1738953078235" updateUser="ExternalLinkService">
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<mods:title id="2C3D56CBA31266A95D4229C314D4D176">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="A96E94ECEEFA868D7AF82C050000E3AF">Rollinson, Lisa</mods:namePart>
<mods:nameIdentifier id="9F0244DDF9DEE3CB6F009B84A555F61F" type="ORCID">https://orcid.org/0009-0007-0152-7073</mods:nameIdentifier>
<mods:affiliation id="2EEE29ACD65B4F70A569193FB07BF764">Texas A &amp; M University, College Station, Texas, USA</mods:affiliation>
</mods:name>
<mods:name id="0B513592ACE2AB09A7371CB118405EBD" type="personal">
<mods:role id="48CB57D1B1B78EBB45A60CAD46C72EB7">
<mods:roleTerm id="94C231329D5DC90505635B9BAC9F028A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B2D439339532CBF4FEA611E03DD205C">Cabrero, Allan</mods:namePart>
<mods:nameIdentifier id="8E70641C26593C4B4D3C5FB650FCEB79" type="ORCID">0000-0002-2935-3534</mods:nameIdentifier>
<mods:affiliation id="3D30D7F3BC9E7B009575895FDEA7AFBA">National Museum of Natural History, Washington, D. C., USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0334EB72C98170499EED82584891E59A">text</mods:typeOfResource>
<mods:relatedItem id="EBAA0441F9DBB209EAEBD596290EBCC4" type="host">
<mods:titleInfo id="0C7226DEF2E69E31781F0A712413F11A">
<mods:title id="3B43A8637744C5A1EF518FABD9C1AC6B">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="BEC24DA378B96CC1541E7BD5C1B47D70">
<mods:date id="66EA68E8A298F066C09437AD0FAC3AB9">2025</mods:date>
<mods:detail id="C9619FEBBBF84FF0D80D83048950FEA4" type="pubDate">
<mods:number id="978EE3D73C2A97C42AA9E93DA421478C">2025-02-07</mods:number>
</mods:detail>
<mods:detail id="2587CD5F8F116977B82FA67959D8ADC2" type="volume">
<mods:number id="4240FF49D252695D1E4632FFE0C18EFF">66</mods:number>
</mods:detail>
<mods:detail id="728269F94773EB4CC348267ADCE8856D" type="issue">
<mods:number id="79954AC15266D2CA7C24E90DAE077E6A">1</mods:number>
</mods:detail>
<mods:extent id="DBAC8A269F1D3CF3C10BB46E359ACB9A" unit="page">
<mods:start id="AF6BAC4AF41834C681DEBB6FCF94970D">73</mods:start>
<mods:end id="A534DF274B1664BB10A1EAB83F84B967">115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A3F6D25D044E72B9655F9995032F0F61">journal article</mods:classification>
<mods:identifier id="BCBBECB16BE275DD3445AE13D2F2121D" type="DOI">10.3897/afrinvertebr.66.129611</mods:identifier>
<mods:identifier id="7F2D2383546FBFAAEBE82BD8176EA407" type="ZooBank">9A70A477-D862-40E5-A132-3227D96C3ECC</mods:identifier>
</mods:mods>
<treatment id="A66900FAAD9A57698F0E3D7E57629120" ID-DOI="http://doi.org/10.5281/zenodo.14834492" ID-Zenodo-Dep="14834492" ID-arpha="A66900FA-AD9A-5769-8F0E-3D7E57629120" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120" httpUri="http://treatment.plazi.org/id/A66900FAAD9A57698F0E3D7E57629120">
<subSubSection id="6E71492C5D1511B0BB0BE3A276B88985" type="nomenclature">
<paragraph id="6FD92C91DB92E0BDBE9D7247559EF9A2">
<taxonomicName id="895852EE71640AE7737D5940A3106402" authority="(Hesse, 1956)" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="henicoides" status="comb. nov.">
<emphasis id="6A644CB965D24B7E400A92728B5A89AB" italics="true">Enica henicoides</emphasis>
(
<bibRefCitation id="742822D26398699AC64A2553BD689F23" author="Hesse AJ" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 27-46. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="11DBA94C3E03282CCF39246FACBB3CE7" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D9F6F4B71D5CEB5086A811B07F8F4BB6" type="description">
<paragraph id="6BEAE364CFB27A381625DFBF396BD21A">
<figureCitation id="E99A1BCDB9E0C2302C8D844B8D093A85" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Enica henicoides ♀ SAM-DIP-A 002170 (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica imitata ♀ NMSA - DIP- 90027 (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure8" httpUri="https://binary.pensoft.net/fig/1244181">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKKAI" type="Taxon depository">
<paragraph id="02668F58548B616FF51563A0ADE84E1B">
<heading id="E1B158D323924DB5ED5E3667C665CEF3" reason="title">Taxon depository.</heading>
</paragraph>
<paragraph id="CB2E92541C5A40115882EF99EC22E43D">
ZooBank:
<ext-link id="B4B5E9CBAED1C48A734D2BBB2248D9E4" ext-link-type="uri" xlink_href="https://zoobank.org/3BE08163-D9C1-4B57-B5E6-9B0B0ECC9313" xlink_type="simple">
https://zoobank.org/
<uuid id="FEAE8A149B2081D4AE2FFCCE956539FC">3BE08163-D9C1-4B57-B5E6-9B0B0ECC9313</uuid>
</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVKAI" type="diagnosis">
<paragraph id="7BE2D94C9F169310F1E9E733FE1B0AC5">
<heading id="26E1DF04BF3A9598B2B20EAC94203861" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="16AEC9038AB8AB29634873EA7E11C3CC">The species is distinguished from other species in the genus by the dorsal occiput with whitish yellow setae and small ovoid black spots in females. In males it is distinguished by the combination of a dorsal occiput without markings and a dark brown setose haltere stem.</paragraph>
<caption id="97F9E198863133355CC1FC472D32FD45" ID-DOI="10.3897/afrinvertebr.66.129611.figure8" ID-arpha="6F8F6806-AADC-5350-B73C-1DC33A470F50" httpUri="https://binary.pensoft.net/fig/1244181" startId="F8">
<paragraph id="493DB0701430C3EA6BF1E57443E611FF">
<label id="7DE798229971EA323E838BBF7CA57503">Figure 8.</label>
</paragraph>
<paragraph id="EC1CF85DC2D71831B86CA36F423E43EB">
<taxonomicName id="A67830AEB833B4F2DD757E6A9CDF66A5" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="henicoides">
<emphasis id="511058A118C8BBE811FEDC4362B10178" italics="true">Enica henicoides</emphasis>
</taxonomicName>
♀ SAM-DIP-A 002170 (
<emphasis id="29FB293AD516BD4C8429BF39B5E3A164" bold="true">A C</emphasis>
)
<emphasis id="D02C39464CBD9A95D5F881423BD45231" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="9AC52318C7B689211F2235DC31CD2408" bold="true">B</emphasis>
habitus, lateral view
<emphasis id="44B0902CE0AC1910CF12F46F51E6C3D4" bold="true">C</emphasis>
habitus, frontal view;
<taxonomicName id="0BFFE14D575CA4244F0A6DB129A6B085" authority="♀ NMSA" authorityName="NMSA" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="imitata">
<emphasis id="F32CA4E247A9E6B89CA33C33B8924B7C" italics="true">Enica imitata</emphasis>
<named-content id="051B09B150A40835A7E4E9DCF7058400" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
</taxonomicName>
- DIP- 90027 (
<emphasis id="2F654727B1DD4F0A4FB563EEE2B65EB9" bold="true">D F</emphasis>
)
<emphasis id="E60DED53CB459BA5EEA483190F703719" bold="true">D</emphasis>
habitus, frontal view
<emphasis id="FCAFF6C55FD8FF36269665DFCBD9754B" bold="true">E</emphasis>
habitus, dorsal view
<emphasis id="8DCAB6E650DD0EE9B66E80F8C2B69658" bold="true">F</emphasis>
habitus, lateral view. Scale bars: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERMAI" type="description">
<paragraph id="44EF4E50013272380E8C707E077DC0B4">
<heading id="4BB6BFA8DEE669AA60F654E39D33C727" reason="title">Description.</heading>
</paragraph>
<paragraph id="D40BD9B9F0A32553915BA8090B031E6E">
<emphasis id="6402253EE52A02450B61ED1D4D95EA2C" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, without dark spot, males: similar to head, slightly darker than head; frons black setose, with white scales, males no setation near eyes, females setose near eyes only dorsally, past half way point setose only medially; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput whitish yellow setose, brown setose on occiput spots, white scales antero-dorsally, females: antero-dorsally with dark brown to black spots, usually oval males: antero-dorsally without markings; gena creamy white, extremely sparsely setose.
</paragraph>
<paragraph id="7B48B5B6D487C130998ADFF821E01A86">
<emphasis id="934B91AEA467110526E909E9E49E5A0A" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, black, light yellowish-brown setose.
</paragraph>
<paragraph id="7BCCBA006F5B18BB92EEF1F32B1B216A">
<emphasis id="C12E439F4AC57781396493BEE010FCEE" bold="true" italics="true">Antenna</emphasis>
: dark brown; scape creamy white, approximately as long as pedicel, black setose dorsally and ventrally, setae extending to tip of pedicel; pedicel creamy white, short black setose dorsally; postpedicel dark brown, tapering distally, longer than scape and pedicel combined, sparsely white scales and dark setose dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="342E11149E75587487305441BFC607F8">
<emphasis id="105C397B4CD296AF23FD2645E86300E0" bold="true" italics="true">Thorax</emphasis>
: light orangish brown; scutum predominantly black, reddish on margins, light to dark brown scales, two submedial longitudinal stripes with lighter scales; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, white scales dorsally; metepimeron silvery grey pubescence, asetose; scutellum red, anterior black spot, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of yellowish-white scales.
</paragraph>
<paragraph id="E1138D522D996D2B5033645A5C776A96">
<emphasis id="6F49D07B575D1E22EBE290FA00E12608" bold="true" italics="true">Leg</emphasis>
: light orangish brown, covered in scales, some regular setae present; pro coxa light orangish brown, silvery grey pubescence, white scales, thin white setose, thick black setose; pro femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa light orangish brown, white pubescence, white scales, thick black setose; mes femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa light orangish brown, white pubescent, white scales, thin white setose, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="1ED16C49A202FED060685C72BCB974E4">
<emphasis id="72E197C30BD025EEC0B43C47F042343F" bold="true" italics="true">Wing</emphasis>
: 8.99.0 mm, microtrichia absent; membrane color pattern males: as females but less darkly infuscated, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="6DF534E92D849933D53B4BAED322FBA7">+ 3</subScript>
, base of m
<subScript id="DB2ECA59D6498B471E1699544E38A7A2">2</subScript>
, m-cu, split between M
<subScript id="37DC94A567AAE8AABD23E208F24F66A4">1</subScript>
+ 2 and M
<subScript id="F767B0775CCCB9C123A9193C27639FEF">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="0B0A65AA8D93D908325425753B22CF67">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="46050EFBE34EA266A0A638E51D34AB2B">1</subScript>
open; R
<subScript id="5BA523E21CA324DCAF977B4ACE86EC12">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="5E8F1CE0248AF7F9DC11B6377114F18D">4</subScript>
open, proximally R
<subScript id="CE9A8F6E71E394107F757D26B739E179">4</subScript>
and R
<subScript id="9D34C4F170F9512E5D7F14F396068BCD">5</subScript>
parallel, R
<subScript id="C6AF6B735C30058614A864E940860BEF">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="08CDEB074CECC86D8923649FE46B19EE">5</subScript>
terminating posterior to wing apex; r
<subScript id="D7625210C2BEC61FCB561D2552D550AD">5</subScript>
open; M
<subScript id="C99144A6F09395AA67CF51CF08EFCB44">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="85D0952893119273C64BB5E8238CE784">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="B57298E0F5D3F801EBFBEC8981679BF6">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.
</paragraph>
<paragraph id="D923D687DA19AD2E24CB78341FD6B6A9">
<emphasis id="75AFB10A02AD5BD90DC8CE856B509C34" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, predominantly black anteriorly and red posteriorly; tergites smooth, setae with small sockets only; T 1 predominantly reddish-orange, some black or reddish yellow to light brown, light yellowish-brown to light whitish-brown setose, with scales; T 2 T 7 predominantly reddish-orange, thin black setose, brown to light yellowish-brown scales, lighter scales on posterior margins of some tergites; S 18 predominantly light reddish-orange, yellow to white scales, thin black setose and long white setose; lateral margin black setae remaining close to abdomen, male T 8 white scales, black setose.
</paragraph>
<paragraph id="3F9A9658889E35F1E8EA73810AAAF0C3">
<emphasis id="BD5B99B24E16BFCB253258AC66C50C03" bold="true" italics="true">Terminalia</emphasis>
: Male terminalia drawings in
<bibRefCitation id="6AE776414B445C83FB82EF19D5CBAD47" author="Hesse" firstAuthor="Hesse" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 2746. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse (1956</bibRefCitation>
, p 41):
<ext-link id="5C81A62E83ABC741A3FFF4438491B7CA" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/40844180" xlink_type="simple">https://www.biodiversitylibrary.org/page/40844180</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWPAI" type="material">
<paragraph id="3AC4DD79898ABBD84BD19ED36E5E29C3">
<heading id="092D5EEA6A7EB881E32C27AF448C6B43" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="35AB13B9FE7F5F1383A6052043731B66">
<materialsCitation id="B69522FA5456F3D5A7FDF74E7E570BF9" collectingDate="1920-11" collectionCode="NHMUK" collectorName="Turner, R. E." country="South Africa" latitude="-33.36989" location="Ceres" longLatPrecision="1" longitude="19.311861" specimenCode="NHMUK 015136073" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="7AB7AB645993803E8574DE54C4580A0D" bold="true">
<emphasis id="CEBB854EF4DD3A2008B6FBD8D9D73BC1" bold="true" italics="true">
<typeStatus id="79D74641719D795CEEFDBCA9BF77BDA1" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="766DE837EFEB1D56CB8B579CD975163A" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="1AA3D5A0B8C6B99B1C2933CCDB3E84D7" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="9623C55AE4068345D28BBDA39EEF6C1C" count="1" type="male">1 ♂</specimenCount>
<location id="57717EA7D99A80B4C069059A2CFCEBBE" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:57717EA7D99A80B4C069059A2CFCEBBE" country="South Africa" latitude="-33.36989" longLatPrecision="1" longitude="19.311861" name="Ceres" stateProvince="Western Cape">Ceres</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEQAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.311861,-33.369889]}">
<geoCoordinate id="7B8B9AF00B0739D372DB12F692ACDCE7" degrees="33" direction="south" minutes="22" orientation="latitude" precision="1" seconds="11.6" value="-33.36989">33 ° 22 ' 11.6 &quot; S</geoCoordinate>
,
<geoCoordinate id="AE6973881BBA00E4B293BB639D6DEEC6" degrees="19" direction="east" minutes="18" orientation="longitude" precision="1" seconds="42.7" value="19.311861">19 ° 18 ' 42.7 &quot; E</geoCoordinate>
</named-content>
;
<date id="758B100AC8155FD25C8F4DF1415FFA53" value="1920-11">
<collectingDate id="0AF62C84E2479B604C435A15EA5DD52C" value="1920-11">Nov. 1920</collectingDate>
</date>
;
<collectorName id="82AAA5F5465ACE06ED7534AFC5CC5E54">Turner, R. E.</collectorName>
leg.;
<specimenCode id="64E59018DFD095513E54EDEE2D127DF9" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK 015136073</specimenCode>
,
<named-content id="0A7A5C015954E8A46D1D99A7439BE9E1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="48B36F7199167DC0D555D24EDA6E4E55" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
.
</materialsCitation>
</paragraph>
<paragraph id="F16095FBE58AE932AB79E01B6676F128">
<materialsCitation id="4975CDA1719DFB128B679E2C1EF08FDF" collectingDate="1931-11" collectionCode="SAMC" collectorName="Cockerell, D. A." country="South Africa" latitude="-31.383333" location="Nieuwoudtville" longLatPrecision="20" longitude="19.1" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="2E04651B6D2EAAE7998AC340D53B36F6" bold="true">
<emphasis id="5488688933C5D8DE4C785EB24D66F570" bold="true" italics="true">
<typeStatus id="D8AEB75ACE86D45078393CAE2B343EAD" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="256C2C4392AC88D25F0FBBBAB5D8C4A3" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="4CACB2DA9D65EABD28C34E4F05193C96" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="B1B93279B060957D1D1B96A34C5C2C62" count="1" type="female">1 ♀</specimenCount>
<location id="57BDCC2A0DEE678D731781AFAB8E004D" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:57BDCC2A0DEE678D731781AFAB8E004D" country="South Africa" latitude="-31.383333" longLatPrecision="20" longitude="19.1" name="Nieuwoudtville" stateProvince="Northern Cape">Nieuwoudtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXQAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.100000,-31.383333]}">
<geoCoordinate id="5489A81092693A6D03C22F30464E5E44" degrees="31" direction="south" minutes="23" orientation="latitude" precision="15" seconds="00" value="-31.383333">31 ° 23 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="6BDB5072767BDA4310CB8A3ED8D5DB41" degrees="019" direction="east" minutes="06" orientation="longitude" precision="15" seconds="00" value="19.1">019 ° 06 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="2476F673D0806D2E2E042E01C4C19EA7" value="1931-11">
<collectingDate id="5DFADA8FA793E046345DA7FEEA42DAB8" value="1931-11">Nov. 1931</collectingDate>
</date>
;
<collectorName id="9DF01513C446AC5C5DBDAB0DE2D676B1">Cockerell, D. A.</collectorName>
leg.; SAM-DIP-A 002171,
<named-content id="3532B959DD193E340FEE9167D4048C49" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="D51266B69212B86A165B54F106665C9F" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="BFE1684B75ADD7C606D1FC302C9AF97C" collectingDate="1931-11-18" collectingDateMax="1931-11-22" collectingDateMin="1931-11-18" collectionCode="NHMUK" collectorName="Mackie, A." country="South Africa" latitude="-31.383333" location="Nieuwoudtville" longLatPrecision="20" longitude="19.1" specimenCode="NHMUK 015136071" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape" typeStatus="paratype">
<specimenCount id="107F7230CCE0C928DE08EAA6D6B9FCD8" count="1" type="female">1 ♀</specimenCount>
<location id="1062C1E72879EE7A2B61603FCF937179" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:1062C1E72879EE7A2B61603FCF937179" country="South Africa" latitude="-31.383333" longLatPrecision="20" longitude="19.1" name="Nieuwoudtville" stateProvince="Northern Cape">Nieuwoudtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDRAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.100000,-31.383333]}">
<geoCoordinate id="7D7F87527634032BCB4B9437A6F1157F" degrees="31" direction="south" minutes="23" orientation="latitude" precision="15" seconds="00" value="-31.383333">31 ° 23 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="C82DCBCE70D60EA0AEAD2350EA09E91B" degrees="019" direction="east" minutes="06" orientation="longitude" precision="15" seconds="00" value="19.1">019 ° 06 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="E3BE167436C02F3E943853B8942CBCAA" value="1931-11-18" valueMax="1931-11-22" valueMin="1931-11-18">
<collectingDate id="25AA33B99C929A7811EB1C054787FDDB" value="1931-11-18" valueMax="1931-11-22" valueMin="1931-11-18">18-22 Nov. 1931</collectingDate>
</date>
;
<collectorName id="115DCCF2274CF33A1CC120705ABE5D5B">Mackie, A.</collectorName>
leg.;
<specimenCode id="DA8159D52D46479B525B5225223B8B9B" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK 015136071</specimenCode>
,
<named-content id="3468373718C811EB998391514BF0010E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="DB1F56F36D73C6EE9B1CB0A5D9F6822C" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="26CFD3CC2A73249750285ECD9E7F3E5A">
<materialsCitation id="ED989A64387E2EE66A9119FDAE20C17D" collectingDate="1950-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.04778" location="Bulhoek" longLatPrecision="20" longitude="18.824446" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<collectingCountry id="740775B54394456B579D707A09B54731" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="E8E7B2DB378C2C3994BBAAA295987A36" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="7332BF2156674F4D36DB388B4359DD48" count="1" type="female">1 ♀</specimenCount>
<location id="865F8A1B112EE2E701DC73873377E813" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:865F8A1B112EE2E701DC73873377E813" country="South Africa" latitude="-32.04778" longLatPrecision="20" longitude="18.824446" name="Bulhoek" stateProvince="Western Cape">Bulhoek</location>
(= Bulshoek), Klaver - Clanw. (= Klawer - Clanwilliam);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERRAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.824444,-32.047778]}">
<geoCoordinate id="F9B57514E052A43421A46949DA3A3D6D" degrees="32" direction="south" minutes="02" orientation="latitude" precision="15" seconds="52" value="-32.04778">32 ° 02 ' 52 &quot; S</geoCoordinate>
,
<geoCoordinate id="8581797F9F40B08E8DF62384B1552F74" degrees="018" direction="east" minutes="49" orientation="longitude" precision="15" seconds="28" value="18.824446">018 ° 49 ' 28 &quot; E</geoCoordinate>
</named-content>
;
<date id="AD1362F5F2FCC8347B8336E9EB55C0C2" value="1950-10">
<collectingDate id="2052FA7D2D724B99623367B332E18086" value="1950-10">Oct. 1950</collectingDate>
</date>
; SAM
<collectorName id="AC2C5DBA2FCA65F2BCC2EF00157F326A">Museum Staff</collectorName>
leg.; SAM-DIP-A 002170,
<named-content id="AAE16B8C2E232EE1EF7775CA35E8E40D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="6333FB6C29D4FE1F5272C458D03FFFD0" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="5885B09F7B6E97363247339329B67FF1" collectingDate="1950-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.04778" location="Bulhoek" longLatPrecision="20" longitude="18.824446" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="8B941869028C42B6F27DABB2A6430EFC" count="1" type="female">1 ♀</specimenCount>
<location id="6438F57F57437292AFC0303F60DC0AA9" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:6438F57F57437292AFC0303F60DC0AA9" country="South Africa" latitude="-32.04778" longLatPrecision="20" longitude="18.824446" name="Bulhoek" stateProvince="Western Cape">Bulhoek</location>
(= Bulshoek), Klaver - Clanw. (= Klawer - Clanwilliam);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4RAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.824444,-32.047778]}">
<geoCoordinate id="F3F057B8E5353E22FFF16579D4095AF0" degrees="32" direction="south" minutes="02" orientation="latitude" precision="15" seconds="52" value="-32.04778">32 ° 02 ' 52 &quot; S</geoCoordinate>
,
<geoCoordinate id="AD1C8859AA26D4EB5061FB142722645B" degrees="018" direction="east" minutes="49" orientation="longitude" precision="15" seconds="28" value="18.824446">018 ° 49 ' 28 &quot; E</geoCoordinate>
</named-content>
;
<date id="1F6989F2772217A3F5773420D98FAD5F" value="1950-10">
<collectingDate id="DF0057D3965F3F5FC5524B0BC196EAE3" value="1950-10">Oct. 1950</collectingDate>
</date>
; SAM
<collectorName id="6277704F3C04DDA5A11635FF6AC8C7C5">Museum Staff</collectorName>
leg.; SAM-DIP-A 002170,
<named-content id="7A4410EDE3F1F85EA83B2BA115564AE3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E751DE332AA6E63F624E870246B0E1A8" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="4B1B34B957675FBFBDD5EB468D8C229C" collectingDate="1930-12-11" collectionCode="NHMUK" collectorName="Simmons, H. W." country="South Africa" latitude="-33.90678" location="Town" longLatPrecision="1" longitude="19.121944" municipality="Franschhoek" specimenCode="NHMUK 015136072" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="E6AFB58F817F11045C1D745A52AA07AE" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="F2777C2980C2E37B588FDAD1E7A1B72D">Franschhoek</collectingMunicipality>
(= French Hoek),
<quantity id="5E2297C026BB6FA7386AA2CE441985FB" metricMagnitude="4" metricUnit="m" metricValue="6.437376" unit="mi" value="40.0">40 mi</quantity>
from
<collectingRegion id="D3492182A1EE0D0EB850D790104BED01" country="South Africa" name="Western Cape">Cape</collectingRegion>
<location id="75ED75BC682193E8B1610F4A7AA9D4CE" LSID="urn:lsid:plazi:treatment:A66900FAAD9A57698F0E3D7E57629120:75ED75BC682193E8B1610F4A7AA9D4CE" country="South Africa" latitude="-33.90678" longLatPrecision="1" longitude="19.121944" municipality="Franschhoek" name="Town" stateProvince="Western Cape">Town</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJSAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.121944,-33.906778]}">
<geoCoordinate id="A1690E765285065C6E0076864AED2E5D" degrees="33" direction="south" minutes="54" orientation="latitude" precision="1" seconds="24.4" value="-33.90678">33 ° 54 ' 24.4 &quot; S</geoCoordinate>
,
<geoCoordinate id="D8C97C4FE69EC336A2831D3805FE2A6D" degrees="19" direction="east" minutes="07" orientation="longitude" precision="1" seconds="19.0" value="19.121944">19 ° 07 ' 19.0 &quot; E</geoCoordinate>
</named-content>
;
<date id="15366544D494D7EE11513A1E1AB5EDDE" value="1930-12-11">
<collectingDate id="73AC31550D823B7E2CADBFC601A5A6D7" value="1930-12-11">11 Dec. 1930</collectingDate>
</date>
;
<collectorName id="4AC550BD6159471D0DD01706058488B3">Simmons, H. W.</collectorName>
leg.;
<specimenCode id="A34A16536C544711A34277CC77EE0F77" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK 015136072</specimenCode>
,
<named-content id="A9EBF38614CC3E1EBDA0A61C6EFFE435" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="49621B9C57F96D503EE797AA8C335B89" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESSAI" type="distribution">
<paragraph id="BE172B97A0264C491B2D639615928E39">
<heading id="7A539EE70163B8A2EA56E6342B375888" reason="title">Distribution, biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="1B77C3DA709EB12B71311CE78387978B">
Known from localities in the
<collectingRegion id="1A29892E21F498E7864E7BC67EF50E48" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
and
<collectingRegion id="99F592E95C6C4A813C8FE21A88F0FC8D" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
of
<collectingCountry id="9AE2B3B40DD4E6AB5230EBACFE490F53" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="4863009FB4D3FD726151ACD78E3C5E99" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. longirostris, E. henicoides and E. sporanthera specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure9" httpUri="https://binary.pensoft.net/fig/1244182">9</figureCitation>
). This species is only known from
<specimenCount id="35554038D2566DC89DCEEBAB00CEC502" count="6" type="generic">6 specimens</specimenCount>
, collected in 4 collecting events spread between 19201950. The species is known to occur in the Succulent Karoo biodiversity hotspots. Adult flies are active from October December (Table
<tableCitation id="C5E09748272EE4F4281909971DAE7342" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="1BCC559CBB7F7CB676AD46D14C368E4B" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="00F5953F4C8B1C25CF2FEB0ADF4EE8D9" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="066F7B92C0BD0C0B1970E6239419812C" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
<caption id="B1245DB313D3E0D54E2AB441B413CACC" ID-DOI="10.3897/afrinvertebr.66.129611.figure9" ID-arpha="7BB56640-B5B7-57BE-8750-36BCA495CEE2" httpUri="https://binary.pensoft.net/fig/1244182" startId="F9">
<paragraph id="ACB483663AF90B969807F863C86DF507">
<label id="55121ADA3F3466BB64C19CB26874E171">Figure 9.</label>
</paragraph>
<paragraph id="97985B26630B7912D223CCAFC78329E7">
Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of
<taxonomicName id="3B8ED0CC75AB601B92AE23908633CA40" authorityName="Wiedemann" authorityYear="1819" class="Insecta" family="Bombyliidae" genus="Anthrax" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="5E954C3C2611625051A6655C75C673F1" italics="true">E. longirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="1186090F344BD223C95EBB42B7404E57" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="henicoides">
<emphasis id="C4CB85D7DA4BA76FE6012CD95534B0D8" italics="true">E. henicoides</emphasis>
</taxonomicName>
and
<taxonomicName id="767748375D35902232066144B6B84960" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="sporanthera">
<emphasis id="4FF4C51B96667C4453B7F4E0478AAE92" italics="true">E. sporanthera</emphasis>
</taxonomicName>
specimens studied.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,513 @@
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<mods:classification id="BDD0CF07A670A01AC1338B5ECAB9E21B">journal article</mods:classification>
<mods:identifier id="8C3DE2DCF70E74E450DA374CAD057ADB" type="DOI">10.3897/zookeys.1226.142678</mods:identifier>
<mods:identifier id="66BF73ABCE949853A3DE604FA0A9D871" type="ZooBank">0B9EFAA1-3377-42F0-B43A-B77F439F88FE</mods:identifier>
</mods:mods>
<treatment id="BBE478042F23567196C8BC6D11FFE30A" ID-DOI="http://doi.org/10.5281/zenodo.14834506" ID-Zenodo-Dep="14834506" ID-arpha="BBE47804-2F23-5671-96C8-BC6D11FFE30A" ID-zoobank="https://zoobank.org/A513A8DB-BB5D-46C8-9407-8C0DA236DC10" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A" LSID-ZBK="urn:lsid:zoobank.org:act:A513A8DB-BB5D-46C8-9407-8C0DA236DC10" httpUri="http://treatment.plazi.org/id/BBE478042F23567196C8BC6D11FFE30A">
<subSubSection id="F3F36377F81BE8BA993739D897894748" type="nomenclature">
<paragraph id="21D013905F382C2E5D090F7A3F76E06C">
<taxonomicName id="79FC5B5540236D9CC890BB4041C66014" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="51ACF0079A14EC63E6D7795CD1103EB5" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63ACAC88009944441FB13F12D4239299" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="35F80295A780E25FF54BA18C251E6A5A" type="description">
<paragraph id="DCC1F07B4B238AB3E9B9D20AC0CD30FC">
<figureCitation id="0242F32DCB5A59325D675CC636C57228" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Megadrypta spp. A holotype of M. maozhoui sp. nov. B female of M. mirabilis from Hunan. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure2" httpUri="https://binary.pensoft.net/fig/1244300">Figs 2 A</figureCitation>
,
<figureCitation id="6203FB7136CD0B09AF35B16A43360FC0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3</figureCitation>
,
<figureCitation id="9430BAD8E5321DC2ABA74DA62DB8A505" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5</figureCitation>
,
<figureCitation id="26CA220CCDC4957BB7F40D993799F718" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mentum of Megadrypta spp. A M. maozhoui sp. nov. B M. mirabilis. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1226.142678.figure6" httpUri="https://binary.pensoft.net/fig/1244304">6 A</figureCitation>
,
<figureCitation id="D73C8C2A7C3EBADA49775C7B71ECDF9F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Collecton site and habitus of M. maozhoui sp. nov. A Collection site on Mount Tiantaishan, Sichuan (B and C photographed in area of the red circle, D photographed in area of yellow circle) B D habitus (photographed by Mao-Zhou Xu) E habitus, from Mount Zhougongshan (photographed by Yong Wang)." figureDoi="10.3897/zookeys.1226.142678.figure7" httpUri="https://binary.pensoft.net/fig/1244305">7 Chinese vernacular name: 茂洲巨逮步甲</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4PAC" type="type material">
<paragraph id="9B1F7E2B2AFC72AB80B80B330F1E33B8">
<heading id="629448612D437FDB863972003B1CB50A" reason="title">Type materials.</heading>
</paragraph>
<paragraph id="7432417967BF601CA434B43AE896F90F">
<materialsCitation id="B9661FFF9EB1660EFADFD26E1ED2F46D" collectingDate="2024-05-01" collectingDateMax="2024-05-05" collectingDateMin="2024-05-01" collectionCode="IZAS" collectorName="Tianxuan GU" country="China" county="Qionglai" determinerName="Chen" elevation="1246" latitude="30.2674" location="Sandaowan" longLatPrecision="6" longitude="103.1199" municipality="Mt. Tiantaishan" specimenCount="2" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="949ED302077CD812A88FF1D735901C03" bold="true">
<emphasis id="6B1392E4412EC433697EA21D772D4D24" bold="true" italics="true">
<typeStatus id="1F19583D0C992C21B774A4B45EE9B620" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="AD27967B5F447EC11E6AAE7A6D4F4803" name="China">China</collectingCountry>
</emphasis>
• male, labeled “
<collectingRegion id="3D719E972CCA23E4330F2122C07CAC08" country="China" name="Sichuan">Sichuan</collectingRegion>
,
<collectingCounty id="7A922F208624415E5ED51D96C5E19C74">Qionglai</collectingCounty>
,
<collectingMunicipality id="CD3DBCA8BFA166B42D1CD245DF5D6235">Mt. Tiantaishan</collectingMunicipality>
,
<location id="F3351D2145AAF1476E4DCC7E810DD4D0" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A:F3351D2145AAF1476E4DCC7E810DD4D0" country="China" county="Qionglai" latitude="30.2674" longLatPrecision="6" longitude="103.1199" municipality="Mt. Tiantaishan" name="Sandaowan" stateProvince="Sichuan">Sandaowan</location>
,
<quantity id="DCFE25598EB6DFAA143E284A2B696FC9" metricMagnitude="3" metricUnit="m" metricValue="1.246" unit="m" value="1246.0">
<elevation id="5A38C64886D49111B76557F98327FBAE" metricMagnitude="3" metricUnit="m" metricValue="1.246" unit="m" value="1246.0">1246 m</elevation>
</quantity>
,
<date id="EAD60DF07F4DE08CB0696A341C48266A" value="2024-05-01" valueMax="2024-05-05" valueMin="2024-05-01">
<collectingDate id="06DC25463192C663C15874DB6AB4283D" value="2024-05-01" valueMax="2024-05-05" valueMin="2024-05-01">2024. V. 1-5</collectingDate>
</date>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMAAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[103.119900,30.267400]}">
<geoCoordinate id="0868AE828B7787CC0D7F2591A8381F37" degrees="30.2674" direction="north" orientation="latitude" precision="5" value="30.2674">N 30.2674</geoCoordinate>
<geoCoordinate id="04F51A98269A91DE347E16C44DA7492C" degrees="103.1199" direction="east" orientation="longitude" precision="5" value="103.1199">E 103.1199</geoCoordinate>
</named-content>
, Maozhou XU &amp;
<collectorName id="78847960F245CF171B66E6A8B980BCED">Tianxuan GU</collectorName>
leg. ”; “
<typeStatus id="1872769AE7643BFC9115366C049D65E6" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="84282A220E8FF8605E8E5E927DB6577B" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="61935F660B96B08A0062D2BF78B71092" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DE8DA8D0274A15C672BDD912BF06773" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="BF461DB01D9A60AF29BCC2E6B2DBC257">Chen</determinerName>
&amp; Shi. 2024 ” [red label];
<named-content id="B8205C0B03C378C948D18BA9BAD1B7B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="BC4D2BB37A442139F037CA0AEF237690" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="30561CFA04F1ED11B307CCAE926B0EB9" country="China" location="China" specimenCount="1" typeStatus="paratype">
<emphasis id="75F21D3DD66EE0FBA1FE5F250A818B5D" bold="true">
<emphasis id="5D1102479474687E621032F6B85175C9" bold="true" italics="true">
<typeStatus id="7F1A32658A3A7AF3CF123E6C65E94AD9" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="4D538E6F43C5C7B0250E4FAC371C5BF3" name="China">China</collectingCountry>
</emphasis>
</materialsCitation>
<materialsCitation id="2243705A763F5B12026648BDC16CD353" collectingDate="2024-05-30" collectionCode="CCJH" collectorName="Maozhou XU" country="China" county="Sandaowan" elevation="1246" latitude="30.2674" location="Qionglai" longLatPrecision="6" longitude="103.1199" municipality="Mt. Tiantaishan" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="43582480CD5FADAACB3D055017E25D97" count="2" type="male">2 male</specimenCount>
and
<specimenCount id="61E177012CE83039479F17DF4FD285C4" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingRegion id="312F308D35ADF1C8F1AC6BEBC665D107" country="China" name="Sichuan">Sichuan</collectingRegion>
,
<location id="B22B3F58D0C4166692369F5364FF5189" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A:B22B3F58D0C4166692369F5364FF5189" country="China" county="Sandaowan" latitude="30.2674" longLatPrecision="6" longitude="103.1199" municipality="Mt. Tiantaishan" name="Qionglai" stateProvince="Sichuan">Qionglai</location>
,
<collectingMunicipality id="E26D4A093294C18A7DE79023324E05E6">Mt. Tiantaishan</collectingMunicipality>
,
<collectingCounty id="75CAD99C0D66846558BEF85019E2A51C">Sandaowan</collectingCounty>
,
<quantity id="8C90107954A1CB9280AE3D7F628831DE" metricMagnitude="3" metricUnit="m" metricValue="1.246" unit="m" value="1246.0">
<elevation id="C349BD9007B9A1111F8C0F39A467AE84" metricMagnitude="3" metricUnit="m" metricValue="1.246" unit="m" value="1246.0">1246 m</elevation>
</quantity>
,
<date id="722AB68A163C827B08369178BA72FFC4" value="2024-05-30">
<collectingDate id="23D18FAD0DAB067C3EF3CD427926DE1E" value="2024-05-30">2024. V. 30</collectingDate>
</date>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHBAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[103.119900,30.267400]}">
<geoCoordinate id="63D05B519FFF40E6084B2060014C0C8C" degrees="30.2674" direction="north" orientation="latitude" precision="5" value="30.2674">N 30.2674</geoCoordinate>
<geoCoordinate id="39B54DA0914DA5FF7643EC1B5AC2725C" degrees="103.1199" direction="east" orientation="longitude" precision="5" value="103.1199">E 103.1199</geoCoordinate>
</named-content>
,
<collectorName id="4212C7C1D323417880D439422010E3AA">Maozhou XU</collectorName>
leg. ”;
<abbrev id="ABBRID0EMBAE" xlink_title="personal collection of Jia-Heng Chen">
<collectionCode id="A18B2D11073D97D27C9A09411EB15DB3">CCJH</collectionCode>
</abbrev>
.
<emphasis id="13BA7B7DA4053BC525DA5D349E36316D" bold="true">
<collectingCountry id="7D3DE39379538EB8FDC99B15BE82A359" name="China">China</collectingCountry>
</emphasis>
</materialsCitation>
<materialsCitation id="9F54E39814918527669C823B320CA1A4" collectingDate="2024-09-09" collectionCode="CAU" country="China" county="Yiliang County" elevation="1311" latitude="27.8809" location="Zhaotong" longLatPrecision="6" longitude="104.6809" municipality="Aiziping" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="2BB56F82513BBA77FDD25DD7086ECF62" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingRegion id="BB43052DD7C871A7FBED9BF7473019AE" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<location id="5920E889152F1DC78B871B4FFAE8A201" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A:5920E889152F1DC78B871B4FFAE8A201" country="China" county="Yiliang County" latitude="27.8809" longLatPrecision="6" longitude="104.6809" municipality="Aiziping" name="Zhaotong" stateProvince="Yunnan">Zhaotong</location>
,
<collectingCounty id="1863BFA7D294C0A3FF17E82C62BF8C0E">Yiliang County</collectingCounty>
,
<location id="2F5B2806247FEECC5A6FEDE4A68B3EFA" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A:2F5B2806247FEECC5A6FEDE4A68B3EFA" country="China" county="Yiliang County" latitude="27.8809" longLatPrecision="6" longitude="104.6809" municipality="Aiziping" name="Luowang Township" stateProvince="Yunnan">Luowang Township</location>
,
<collectingMunicipality id="4F20DBFDED78C26508542B4C27E9BF02">Aiziping</collectingMunicipality>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVBAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[104.680900,27.880900]}">
<geoCoordinate id="82499BCA2474B364A523CCE3B6615F01" degrees="27.8809" direction="north" orientation="latitude" precision="5" value="27.8809">N 27.8809</geoCoordinate>
<geoCoordinate id="F5BC57E0858F35345B78E682D75694BB" degrees="104.6809" direction="east" orientation="longitude" precision="5" value="104.6809">E 104.6809</geoCoordinate>
</named-content>
,
<quantity id="30AB4E14ED2BB26F58DAFD766DED4503" metricMagnitude="3" metricUnit="m" metricValue="1.311" unit="m" value="1311.0">
<elevation id="2352338D7811C0C93F8060658CE0898B" metricMagnitude="3" metricUnit="m" metricValue="1.311" unit="m" value="1311.0">1311 m</elevation>
</quantity>
,
<date id="5433B30727BFDB47AA3DE7E2F6E5202B" value="2024-09-09">
<collectingDate id="C382C425603CACDADD0E11924A3E753C" value="2024-09-09">2024.9. 9</collectingDate>
</date>
, Anxian SHI leg. ”;
<named-content id="A021334E84F08D1E5F8AB4DA2FE3B9E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/china-agricultural-university" xlink_title="Entomological Museum, China Agricultural University">
<collectionCode id="16CC63A8188C72ABF1F7F98A57B19D52" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" type="">CAU</collectionCode>
</named-content>
.
<emphasis id="D7930A1787D6E49802346C1547744A16" bold="true">
<collectingCountry id="03108483CB74130E0A184C5A165FDAF5" name="China">China</collectingCountry>
</emphasis>
</materialsCitation>
<materialsCitation id="F3220CEA6AB236F1EE5232598FCC155F" collectionCode="IZAS" country="China" location="Ya'an" municipality="Mt. Zhougongshan" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="C1D342D7D89C1506189970D06689B13A" count="1" type="female">1 female</specimenCount>
, labeled “
<collectingRegion id="F0DDCB8BDB7115BD9664B74CCE56EEB3" country="China" name="Sichuan">Sichuan</collectingRegion>
,
<location id="CA55DDCA2369CCE437CAF1E44C7BB8E0" LSID="urn:lsid:plazi:treatment:BBE478042F23567196C8BC6D11FFE30A:CA55DDCA2369CCE437CAF1E44C7BB8E0" country="China" municipality="Mt. Zhougongshan" name="Ya'an" stateProvince="Sichuan">Yaan</location>
,
<collectingMunicipality id="817616CB21C209B73ECBB5C9C10C1003">Mt. Zhougongshan</collectingMunicipality>
, 2018.7, Yong WANG leg. ” [alcohol immersion];
<named-content id="D511BD95977DD5113CF96A28478E149C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="3463396C0888A3EC9750F8E7190B1ACB" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="816E728A0CA76C6E8596144B3A7C2D1B" ID-DOI="10.3897/zookeys.1226.142678.figure2" ID-arpha="9CDC06EB-824D-5DE7-A519-4CDBCF88CC9B" httpUri="https://binary.pensoft.net/fig/1244300" startId="F2">
<paragraph id="53AA8952ECBB62CEA7F7AEB2469E9A75">
<label id="5D320DA2FDBE54DFE25DB1E2ECF4991B">Figure 2.</label>
</paragraph>
<paragraph id="7854F20151F447828F91664B5646DE8A">
Habitus of
<taxonomicName id="C26DAE5EA20A28E6E0847C1975048F21" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="E485984314BD86FA34F4931EC50ADA3E" italics="true">Megadrypta</emphasis>
spp.
<emphasis id="DCCD15EDE847920295C397A99CE6C037" bold="true">A</emphasis>
</taxonomicName>
holotype of
<taxonomicName id="2CD94736B780496D7481543306F8D3AE" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="22761C7805BEDF4739F2F448BB816532" italics="true">M. maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FA4003F68B14DB3497F71E4C030F90E" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="D8A1F3076AF5C1048488AA4377808D68" bold="true">B</emphasis>
female of
<taxonomicName id="DE12D5CCB30CA20758A4D97E0E648AD3" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="D2B33CBD34FBD00154B84E62F477DC71" italics="true">M. mirabilis</emphasis>
</taxonomicName>
from Hunan. Scale bar: 5.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESDAE" type="diagnosis">
<paragraph id="6B15489EAE8EA9B77D48A18EA5412DD5">
<heading id="C0C2DB1353C7646400AD141BCBB28ADA" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="00CBADF026F0CE5632C7C93DB0004803">
Dorsal surface dark with distinct bluish metallic luster, mouth parts, antennae and all parts of legs reddish brown. Mentum with anterior margin concave and shallowly emarginate at middle. Pronotum with lateral margins gradually sinuate twice, behind anterior angles and before posterior angles. Elytra with 78 parascutellar pores, significantly narrowed base and discontinuous striae, interrupted by coarse foveae (of similar size as interval width) (Fig.
<figureCitation id="1528E5FB81D6A95F4F64436862D8823B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3 C</figureCitation>
). Apical lamella of median lobe of aedeagus short (
<abbrev id="ABBRID0E3DAE" xlink_title="length of the apical lamella of the aedeagus">ALL</abbrev>
/
<abbrev id="ABBRID0EAEAE" xlink_title="width of apical lamella of the aedeagus">ALW</abbrev>
0.550.61), obtuse and rounded (Fig.
<figureCitation id="FD8C4D9503948A78E0ADADC91F88D746" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5 B, C</figureCitation>
); median lobe of aedeagus in lateral view stout, gradually attenuate to apex, ventral margin very faintly curved before apical lamella (Fig.
<figureCitation id="F07C88235E3054C6730C0AAD333899D5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5 A, D</figureCitation>
); endophallus with a distinct central sclerite (Fig.
<figureCitation id="AA66091A368D976AFD9F36D59FE81173" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5 C, D</figureCitation>
).
</paragraph>
<caption id="C85C1853BA9D70D59E63ED51CAA3BE6A" ID-DOI="10.3897/zookeys.1226.142678.figure3" ID-arpha="EE7AE5F5-03C2-53F6-8895-9B1B36DAA4C4" httpUri="https://binary.pensoft.net/fig/1244301" startId="F3">
<paragraph id="E50C31A59F5081F29D0709A67DA40780">
<label id="44627CD4734F6CB36301A981EFA5DCA7">Figure 3.</label>
</paragraph>
<paragraph id="BB8E47A0A7EF1A1CC8F6A0F27730D6C1">
Holotype of
<taxonomicName id="44177315BDEBBA2072D6EA4A8264CADC" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="CE78CB46AE333C0BBBB7D03C6F01C815" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2CDC6A5DC174A4375A153DED6272CD0" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="CB2472BC8B647A58F7F782639EE45FB4" bold="true">A</emphasis>
head in dorsal view
<emphasis id="9553B31C01963ECE2A3E92C2589008B6" bold="true">B</emphasis>
pronotum
<emphasis id="2CAE190CC1CC3BF001DDCFD92E5AA853" bold="true">C</emphasis>
basal elytra
<emphasis id="BAFFD1A6C61C97B52EC38FE516864716" bold="true">D</emphasis>
abdomen. Scale bars: 1.0 mm (
<emphasis id="3EFAA0A1E9387346BBADD215839C9543" bold="true">A C</emphasis>
); 2.0 mm (
<emphasis id="5B733E998D0D4D70F2CB0A7C9BACD70C" bold="true">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EUFAE" type="comparisons">
<paragraph id="50F99FCE5127BAE2157C8148FA49838D">
<heading id="9383EE1730CA665255CA4A9120F1114D" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="8EC23D10DE83D605B353B8074C258AA7">
<taxonomicName id="126E39478A5ED65E02682967350E657D" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="78A7336535DF32A82BC9E6D5029E4EEA" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5E40BB7F1C694805ED877D9D9CA561A2" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from the
<typeStatus id="2491DF61A52BAAC0D108428484046154">type</typeStatus>
species of the genus by the combination of following characters: (1) dorsum with distinct bluish metallic luster (versus dark without or with very faint metallic luster in
<taxonomicName id="0A17F12E1F0A7D974C7556C1186DCB1F" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="93AFD1E8F43FA94D14F0E164076469F2" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (2) legs entirely reddish brown, always much paler than that of the latter species; (3) mentum with anterior margin concaved and shallowly emarginate at middle (versus straight and without emargination in
<taxonomicName id="4BF2036AE9ED3E8F2F29CFA02A0CADCF" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="CAAF5E5B085100CBAEF25A2645B16FCD" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (4) pronotum narrower
<abbrev id="ABBRID0E2GAE" xlink_title="length of the pronotum">PL</abbrev>
/
<abbrev id="ABBRID0E6GAE" xlink_title="width of the pronotum">PW</abbrev>
1.761.88 (versus
<abbrev id="ABBRID0EDHAE" xlink_title="length of the pronotum">PL</abbrev>
/
<abbrev id="ABBRID0EHHAE" xlink_title="width of the pronotum">PW</abbrev>
<quantity id="0D4A0D10A6870B002CD7B4B5128505D7" metricMagnitude="-2" metricUnit="m" metricValue="4.191" metricValueMax="4.2672" metricValueMin="4.1148" unit="in" value="1.65" valueMax="1.68" valueMin="1.62">1.621.68 in</quantity>
<taxonomicName id="EB14DC2C415DCBFD9320669F9BB0E6F4" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="1869031AB991AE2911FF53796ADA93D1" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (5) elytra more elongated,
<abbrev id="ABBRID0EWHAE" xlink_title="length of the elytra">EL</abbrev>
/
<abbrev id="ABBRID0E1HAE" xlink_title="width of the elytra">EW</abbrev>
1.561.68 (versus
<abbrev id="ABBRID0E5HAE" xlink_title="length of the elytra">EL</abbrev>
/
<abbrev id="ABBRID0ECIAE" xlink_title="width of the elytra">EW</abbrev>
<quantity id="CEEA3C4BB11B669CC575B8D9A471E56C" metricMagnitude="-2" metricUnit="m" metricValue="3.7338" metricValueMax="3.81" metricValueMin="3.6576" unit="in" value="1.47" valueMax="1.5" valueMin="1.44">1.441.50 in</quantity>
<taxonomicName id="4AA21D2C826EAFD84EE5727B93450371" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="0362EF5D033D0430BEC8EB8D62EB050B" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (6) elytral striae discontinuous, interrupted by coarse foveae (versus striae entire with normal punctures in
<taxonomicName id="B4D2A05F596ACD16461DB146DB638D1E" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4D93610B4237443479C834B92CB93C29" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (7) median lobe of aedeagus in lateral view gradually attenuate to apex, ventral margin nearly straight before apex (versus median lobe evenly thickened before apical third and then abruptly narrowed, ventral margin distinctly curved before apex in
<taxonomicName id="34BCC1B434D637E53A52DAD9CFE9B376" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="DEA535466100A00DE193A0323EC30EC4" italics="true">M. mirabilis</emphasis>
</taxonomicName>
); (8) apical lamella of aedeagus with apex obtuse and rounded in dorsal view (versus a little narrower in the latter species); (9) central sclerite of endophallus more distinctly sclerotized with clear border in ventral view (versus less sclerotized with ambiguous border in the latter species).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHJAE" type="description">
<paragraph id="EC012292C918924D3138DB9BA64D48B2">
<heading id="4885B9E517D39CFB7DFA8156DE9E7F47" reason="title">Description.</heading>
</paragraph>
<paragraph id="F133E5EF0078F7BF8CB4991E792ED6C5">
<abbrev id="ABBRID0ENJAE" xlink_title="length of the body">BL</abbrev>
<quantity id="7964FD164E40838BA37CCB8F3F094EE2" metricMagnitude="-2" metricUnit="m" metricValue="1.455" metricValueMax="1.55" metricValueMin="1.3599999999999999" unit="mm" value="14.55" valueMax="15.5" valueMin="13.6">13.615.5 mm</quantity>
(
<quantity id="7FDA07C3E2091D11FE7441A7D336B836" metricMagnitude="-2" metricUnit="m" metricValue="1.38" unit="mm" value="13.8">13.8 mm</quantity>
in
<typeStatus id="C7A184F1FDF231D14D2CF53C64C1079E" type="holotype">holotype</typeStatus>
); dorsal and ventral surface with distinct bluish metallic luster; palpi, antennae and all parts of legs reddish brown, mandibles dark brown.
<emphasis id="6F2082A3A10403294D0801349587A20B" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="727BC1D050B154E1D48F6E2B74CC96F4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3 A</figureCitation>
): mandibles long, about five times as long as labrum, hooked at apex; labrum trilobate, medial lobe slightly extended anteriorly, anterolateral angles rounded-obtuse, sparsely pubescent near anterior margin; clypeus impunctate and pubescent; vertex densely and coarsely punctate; temporae slightly shorter than eyes, well tumid behind eyes, superficially pubescent and punctate; antennae long, a little exceeding midpoint of elytra.
<emphasis id="35CCC762EB45BA40EC6F20A1A4A0E0B1" bold="true" italics="true">Pronotum</emphasis>
(Fig.
<figureCitation id="397A086322742889AA625D6737398E4C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3 B</figureCitation>
) cylindrical (
<abbrev id="ABBRID0E6JAE" xlink_title="length of the pronotum">PL</abbrev>
/
<abbrev id="ABBRID0EDKAE" xlink_title="width of the pronotum">PW</abbrev>
1.761.88), disc pubescent and coarsely punctate; anterior angles obtuse; lateral margins distinctly doubly sinuate behind anterior angles and before posterior angles; lateral margins entirely beaded; posterior angles nearly rectangular, apex rounded; median line shallow but distinct; sides of pronotum flattened from above and a little laterally extended near middle, median region elevated.
<emphasis id="299A4C897C29176EF766851111C43314" bold="true" italics="true">Elytra</emphasis>
(Fig.
<figureCitation id="37BB3CF2473A88FAFC7005EC208015EB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Megadrypta maozhoui sp. nov. A head in dorsal view B pronotum C basal elytra D abdomen. Scale bars: 1.0 mm (A C); 2.0 mm (D)." figureDoi="10.3897/zookeys.1226.142678.figure3" httpUri="https://binary.pensoft.net/fig/1244301">3 C</figureCitation>
) wide (
<abbrev id="ABBRID0EOKAE" xlink_title="length of the elytra">EL</abbrev>
/
<abbrev id="ABBRID0ESKAE" xlink_title="width of the elytra">EW</abbrev>
1.561.68), widest on apical third; with 78 parascutellar pores; base strongly narrowed; intervals slightly convex and evenly pubescent; striae discontinuous, interrupted by coarse foveae, the foveae much wider than parascutellar pores, in similar size of interval width; posterior margins truncated, slightly sinuate near outer-apical angles; outer-apical angles obtuse, sutural angles acute angled.
<emphasis id="5A69D92E48C77DF954C003D1C7635687" bold="true" italics="true">Male genitalia</emphasis>
(Fig.
<figureCitation id="840028B66B387BFE69CFC839C3499224" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5</figureCitation>
): median lobe of aedeagus relatively stout, gradually attenuate to apex; in lateral view, median lobe widest near base, ventral margin very weakly curved before apical lamella (Fig.
<figureCitation id="3BC110FE18C65D4D71E30A5C38C708B5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Median lobe of aedeagus, a paratype of Megadrypta maozhoui sp. nov. from Tiantaishan, Sichuan A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure5" httpUri="https://binary.pensoft.net/fig/1244303">5 A D</figureCitation>
), apical lamella slightly thickened apically; median lobe in dorsal view (Fig.
<figureCitation id="8D85A2AD308AE467068D7DA667B87284" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Median lobe of aedeagus, Megadrypta mirabilis. A right lateral view B ventral view C dorsal view D left lateral view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1226.142678.figure4" httpUri="https://binary.pensoft.net/fig/1244302">4 B, C</figureCitation>
), with short (
<abbrev id="ABBRID0EFLAE" xlink_title="length of the apical lamella of the aedeagus">ALL</abbrev>
/
<abbrev id="ABBRID0EJLAE" xlink_title="width of apical lamella of the aedeagus">ALW</abbrev>
0.550.61), obtuse and rounded apical lamella; endophallus with a distinct central sclerite, the central sclerite axe-shaped, with well-defined border in ventral view.
</paragraph>
<caption id="FD28946F07FA99A740442C637F20852F" ID-DOI="10.3897/zookeys.1226.142678.figure4" ID-arpha="3E436279-15CB-5D20-80A0-A3C2076373FB" httpUri="https://binary.pensoft.net/fig/1244302" startId="F4">
<paragraph id="5849742868A474DA37CC2CC6B2D14501">
<label id="A554C0C88FA25CAC58A995AD4379C2C0">Figure 4.</label>
</paragraph>
<paragraph id="47A97EF644E4FCB450734BAC62FA483C">
Median lobe of aedeagus,
<taxonomicName id="D064DE58F36A7D887608594A6E3673D7" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="110D0E4C6ECA14217722E04AE21F6016" italics="true">Megadrypta mirabilis</emphasis>
</taxonomicName>
.
<emphasis id="DD486D820A5E184CF5EA8F3F33EB3C05" bold="true">A</emphasis>
right lateral view
<emphasis id="19E8B852F42FA2F9338453A7918E06B0" bold="true">B</emphasis>
ventral view
<emphasis id="E9743A4406A9D6CAFBF2BDEC07ED44FD" bold="true">C</emphasis>
dorsal view
<emphasis id="B4A1EEA09ED6D7CCF19498EABCE69755" bold="true">D</emphasis>
left lateral view. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="E1244A22420F1498BEA634B072B11C17" ID-DOI="10.3897/zookeys.1226.142678.figure5" ID-arpha="2A3A444B-BA1B-5BA6-A624-8A843C98AD2D" httpUri="https://binary.pensoft.net/fig/1244303" startId="F5">
<paragraph id="846479D04FC38520B5F40A98ECC48F8D">
<label id="3653B001D874839E46F0A42D47BCFF2A">Figure 5.</label>
</paragraph>
<paragraph id="38570DE2E4FFB93EDC8DE9656D1AF77B">
Median lobe of aedeagus, a paratype of
<taxonomicName id="B2CFAAFDE364AAF96973876363F305E8" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="D039C318827452D85B12A0BB44EC97E1" italics="true">Megadrypta maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74892F14824387D8DBA670AC6F16346E" rank="species">sp. nov.</taxonomicNameLabel>
from Tiantaishan, Sichuan
<emphasis id="CE9BB5B69A770F550AEFF00F80214CAB" bold="true">A</emphasis>
right lateral view
<emphasis id="44836DEC5152676CFB4701F5942B51E0" bold="true">B</emphasis>
ventral view
<emphasis id="3A4A447F5228ACCBDC961874D8075E21" bold="true">C</emphasis>
dorsal view
<emphasis id="22A66A637719B54C1AC4674E05276BAD" bold="true">D</emphasis>
left lateral view. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="A6BA45AA032D6D5EC8EA10B4BA1A8237" ID-DOI="10.3897/zookeys.1226.142678.figure6" ID-arpha="A210621C-F7C8-56BD-9C3C-0549E3C84866" httpUri="https://binary.pensoft.net/fig/1244304" startId="F6">
<paragraph id="D42FE785F115D1A91401DF26D4FE8E6F">
<label id="7B41C08F9DEC0D35D28B6A83E36B5A42">Figure 6.</label>
</paragraph>
<paragraph id="77EFF37DB8439C697AC5CCA554198FBF">
Mentum of
<taxonomicName id="350C9591808970C78307309E75C43314" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="17A830D95735476464FAAC07886D658A" italics="true">Megadrypta</emphasis>
spp.
<emphasis id="4BB0287F78EA8C8B630F15FF1D79D225" bold="true">A</emphasis>
</taxonomicName>
<taxonomicName id="DAABC1459EAB07944871633F69B2C3E1" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="7FA1D5157112964F28D889DAA75B6A95" italics="true">M. maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="898C24159837FAB6A837CA4A384049CB" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0018EFEA6C4359B9305643C3D9263232" bold="true">B</emphasis>
<taxonomicName id="1E417C0E1D56A837A69D34D5F3291C4F" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="03E2489F1C94FBE83559AC1AA7477E7C" italics="true">M. mirabilis</emphasis>
</taxonomicName>
. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="8EFD43D0D7E41682896B2D5768C4B6A3" ID-DOI="10.3897/zookeys.1226.142678.figure7" ID-arpha="EB6F1937-ADB1-5090-8243-1BE1979F9181" httpUri="https://binary.pensoft.net/fig/1244305" startId="F7">
<paragraph id="3EBA00C23B8CD6FA41106F0226FFC970">
<label id="921682A3C95D269285FA33F3C84550E5">Figure 7.</label>
</paragraph>
<paragraph id="B4C731D65698B06A6AC6DEBDFAF915CA">
Collecton site and habitus of
<taxonomicName id="F2E556D270AB811D0888BB4106305B08" authorityName="Chen &amp; Shi &amp; Liang" authorityYear="2025" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maozhoui" status="sp. nov.">
<emphasis id="945F550FADE4A12DF1CF8CA72AF9E870" italics="true">M. maozhoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEB478CBA9666D2628FB68C4A3A39921" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="2FD00753F4B226EFED51E75EAB401636" bold="true">A</emphasis>
Collection site on Mount Tiantaishan, Sichuan (
<emphasis id="71F57FA5968D3FCF3A5B29C751F8754E" bold="true">B</emphasis>
and
<emphasis id="84D71D02B5CF6ADD6B2FFC4CBF74ACA1" bold="true">C</emphasis>
photographed in area of the red circle,
<emphasis id="917E7A8E93DB51813352AD3F0A1B5664" bold="true">D</emphasis>
photographed in area of yellow circle)
<emphasis id="7EA7EB67995C70FE79AB4ABA51D8881C" bold="true">B D</emphasis>
habitus (photographed by Mao-Zhou Xu)
<emphasis id="E3D90F5787455AED05C19AB91298369C" bold="true">E</emphasis>
habitus, from Mount Zhougongshan (photographed by Yong Wang).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E6PAE" type="etymology">
<paragraph id="A5E857C488549E9CFDD95177E26CF552">
<heading id="BBEA344B3AA7C8D653412A620A4F2EA4" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A2C81186B064F623BEBAC33E4C2C2E88">
This species is named after Mr Mao-Zhou Xu, who collected and donated most of the
<typeStatus id="ADBAC7A4D0490EB58FCFBAE250C90ADD">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEQAE" type="distribution">
<paragraph id="AA5D23BAD94029561CF53C1647FEB171">
<heading id="D0283B5012B22E2FFBB1E02C87B4BF00" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="360AE493BAA8131ADE5AEF707C1543CF">
This species is known from three localities in two provinces of southwestern
<collectingCountry id="7B9E68FB021BC5D34A7715049F315F6E" name="China">China</collectingCountry>
: Mount Tiantaishan and Mount Zhougongshan of central
<collectingRegion id="6F7DECF8E923047DA304582AD4EA8769" country="China" name="Sichuan">Sichuan</collectingRegion>
, and Luowang Township of northeastern
<collectingRegion id="7B1CFCA4DF5FCE951312A8C05950F8BD" country="China" name="Yunnan">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJQAE" type="habitat">
<paragraph id="DFAA3FEFFB507FDCDA027A33D54625FF">
<heading id="49099223E6CE733DC8D803E800D2E6BD" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="2248D5AC70A792AE3528C64DF3AE6B2D">
(Fig.
<figureCitation id="48D8662750C18C99D331A31ECA957316" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Collecton site and habitus of M. maozhoui sp. nov. A Collection site on Mount Tiantaishan, Sichuan (B and C photographed in area of the red circle, D photographed in area of yellow circle) B D habitus (photographed by Mao-Zhou Xu) E habitus, from Mount Zhougongshan (photographed by Yong Wang)." figureDoi="10.3897/zookeys.1226.142678.figure7" httpUri="https://binary.pensoft.net/fig/1244305">7</figureCitation>
) All specimens were collected on low vegetation under latifoliate forest at night.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,585 @@
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<mods:title id="90E34C05F3BD6101E7E63637509C4AD0">Two new species in the green lacewing genus Santocellus (Neuroptera, Chrysopidae, Leucochrysini)</mods:title>
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<mods:namePart id="5CA90AFCB8757D95D047B706B68A663F">Tauber, Catherine A.</mods:namePart>
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<mods:affiliation id="72E57077758D519D7D41BC327C7ED965">Department of Entomology &amp; Nematology, University of California, Davis, CA 95616 and Department of Entomology, Comstock Hall, Cornell University, Ithaca, NY, USA</mods:affiliation>
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<subSubSection id="DBE9565DEF932B101B590F11D58465AF" type="nomenclature">
<paragraph id="9066EBD974A367813818303B35D06AA4">
<taxonomicName id="B03B7132EE5C92FEDC9A5F9EF37AF857" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="3A80432EEFE5EEEEEE6CD63B6F88A233" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B4F6F5CE76D67BB98D1D7EA2687792" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FC74D98A27C768066CFAFBF80E34FB09" type="description">
<paragraph id="B5517025F0AB36266992C2380BCDC830">
<figureCitation id="DE2579DB79115987C2D57DE4FB8B197D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a frontal b head, prothorax, dorsal c head, frontolateral d ventral. Width across head (dorsal, including eyes), 1.85 mm." figureDoi="10.3897/zookeys.1226.140386.figure1" httpUri="https://binary.pensoft.net/fig/1244337">Figs 1</figureCitation>
,
<figureCitation id="090C10FAD6C9208CE47AC6A8B548EA7E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External features of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a head, thorax, dorsal b body, lateral c abdomen, dorsal d terminus, lateral e protarsal claw, lateral. Width across head (including eyes), 1.85 mm; diameter of callus cerci, ~ 0.2 mm." figureDoi="10.3897/zookeys.1226.140386.figure2" httpUri="https://binary.pensoft.net/fig/1244338">2</figureCitation>
,
<figureCitation id="8DE9E85F9B44BA4F7D0BFB0F865EDF9E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings." figureDoi="10.3897/zookeys.1226.140386.figure3" httpUri="https://binary.pensoft.net/fig/1244339">3 a</figureCitation>
,
<figureCitation id="2B958DF27E187E0BBB99218AA74A309B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male abdomen of Santocellus legrandi, sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a dorsum of abdominal segments A 2 - terminus b lateral view of abdominal segments A 6 - terminus c dorsum of abdominal segments A 7 to terminus with gonarcus everted d lateral view of abdominal terminus. Length of T 8 along dorsal margin, 0.68 mm. Abbreviations: c. c. callus cerci; d. ap. dorsal apodeme of T 9 + ectoproct; gcr field of gonocristae on terminal membrane of S 9; gs gonarcus; S 8 + 9 fused eighth and ninth sternites; T 3, T 7, T 8 third, seventh, and eighth tergites; T 9 + e fused ninth tergite and ectoproct; v. ap. ventral apodeme of S 8 + 9." figureDoi="10.3897/zookeys.1226.140386.figure4" httpUri="https://binary.pensoft.net/fig/1244340">4</figureCitation>
,
<figureCitation id="4B5D90A011779EFBA5EF00DECEA0B055" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a gonarcal complex, dorsal b same, lateral c same, frontal d same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus." figureDoi="10.3897/zookeys.1226.140386.figure5" httpUri="https://binary.pensoft.net/fig/1244341">5</figureCitation>
,
<figureCitation id="BC060C3B7C2E1F7C9B1B41342BF61543" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cleared abdominal terminus of female Santocellus legrandi sp. nov., allotype, Costa Rica, San José (SDMC) a segments A 7 to terminus, lateral b genitalia, lateral c subgenitale, distal view d spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum." figureDoi="10.3897/zookeys.1226.140386.figure6" httpUri="https://binary.pensoft.net/fig/1244342">6</figureCitation>
,
<figureCitation id="FBF13E44CEA430F6810EBBD5DC0C6B15" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci." figureDoi="10.3897/zookeys.1226.140386.figure8" httpUri="https://binary.pensoft.net/fig/1244344">8 a, c, e</figureCitation>
,
<figureCitation id="5578AE46746128DE1ECEE9154EEC8B72" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976)." figureDoi="10.3897/zookeys.1226.140386.figure10" httpUri="https://binary.pensoft.net/fig/1244346">10 d</figureCitation>
,
<figureCitation id="899F98B87608AE427CFB929241C7134C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba." figureDoi="10.3897/zookeys.1226.140386.figure11" httpUri="https://binary.pensoft.net/fig/1244347">11 d</figureCitation>
,
<figureCitation id="D547514C26050D6C22515FF96C6749F3" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species." figureDoi="10.3897/zookeys.1226.140386.figure12" httpUri="https://binary.pensoft.net/fig/1244348">12 d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAJAC" type="etymology">
<paragraph id="490313A9356FD31ED350A73D0EADA30F">
<heading id="E313C1A4A3D0623477FCA080643387CA" reason="title">Etymology of name.</heading>
</paragraph>
<paragraph id="BEFC8B235F4A9316B612FA89FC3DF265">The species carries the surname of Jean Legrand (see Dedication above). Jean had a special love for art (especially painting); it is thus fitting that his name is associated with a beautiful lacewing that has stunning wing markings. The species name is a singular noun in the genitive case following a masculine genus name.</paragraph>
</subSubSection>
<subSubSection id="SECID0EFJAC" type="Type specimens">
<paragraph id="8F71C39D187BF228DA140D63F7D53C76">
<heading id="4774B0D64884F867D41F089AAE0B3DA4" reason="title">
<typeStatus id="07B55DA0FE3CE71C0A32FAD248B68693">Type</typeStatus>
specimens.
</heading>
</paragraph>
<paragraph id="289E9CC10EF1ADCEB2EF8466E155472E">
<emphasis id="A3230220A5CD63A02EF9113D69E9D724" bold="true">
(sex and figures; verbatim label data [specimen location])
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</emphasis>
</emphasis>
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<collectingDate id="D8C5140F1A74C5838113AFE664E12AA3" value="2001-07-31">
31
<superScript id="22A64D6D9DE3A9EA99D5275280537B7A">st</superScript>
July 2001
</collectingDate>
</date>
/
<collectorName id="F54BA1653C9964E9D273CB93C58A131D">G. C. McGavin Coll.</collectorName>
(2)
<collectorName id="0D2259B13100D01A6BCC92BEB3D58903">G. C. McGavin Coll.</collectorName>
/
<collectorName id="E7D37F747699DCF3D638D99F9B54EB90">Pres. G. C. McGavin</collectorName>
/ OUM- 2002-006 [
<named-content id="B4DD8A8C3040BD6FD92503549E3C615E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oxford-university-museum-natural-history" xlink_title="Oxford University Museum of Natural History">
<collectionCode id="B4B5E4631592609EC06B0285468BD220">OUMNH</collectionCode>
</named-content>
].
<emphasis id="98641DEE3EE8F2514FD76BEAD79AD8C5" bold="true" italics="true">
<typeStatus id="61B248FEC841C5AA3966EA35687DD9C9" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
<materialsCitation id="9561ECEA64CDE18537ACF031C6487207" collectionCode="T, CAS" collectorName="H. Haldell &amp; Phillip A. Adams" country="Honduras" county="Male." location="Barraneo" municipality="Rio Paulaya" specimenCount="1" typeStatus="holotype">
<collectingCounty id="3B35528829C3292740E293BDF2FD8ECC">Male.</collectingCounty>
(1)
<collectingCountry id="B37BD4296B6A611D56770C05A7DCAE2E" name="Honduras">Honduras</collectingCountry>
1923 /
<collectingMunicipality id="DF104567DF1E1C9F7F045A44F5133E98">Rio Paulaya</collectingMunicipality>
: /
<location id="351A18DC94037E93D4225C15E597C3FD" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:351A18DC94037E93D4225C15E597C3FD" country="Honduras" county="Male." municipality="Rio Paulaya" name="Barraneo">Barraneo</location>
, iv- 17 / 303
<collectionCode id="29DD89F32240FB424954D43980AB4BEA" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="2849DCBF177AD3F72541CF9AC8132146">H. Haldell</collectorName>
(2)
<collectorName id="87B9CCD25CF3CDE5E80D56EF267EC382">Phillip A. Adams</collectorName>
/ Collection / 1998 bequest to / Calif. Acad. Sci. [
<named-content id="164A7D4E5B884FDFABAD14A03E1715CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">
<collectionCode id="7A7A798F0EBF0D688ADE9A2093B100AC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</named-content>
].
<emphasis id="2634896BD6FA1D2169536DCBA0F89835" bold="true" italics="true">
<typeStatus id="A31DBCDA871159901B751803CE129CF8" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
<materialsCitation id="8605F33FABDF7200E03B972BA7A26F87" collectionCode="NHMUK" country="Panama" determinerName="P. Adams" location="Male." specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<location id="9C0AA9CAEBE17FF15DFEAF2388991DA2" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:9C0AA9CAEBE17FF15DFEAF2388991DA2" country="Panama" name="Male." stateProvince="Chiriqui">Male.</location>
(1)
<collectingRegion id="8425BE03ED463F176036238B702E74AB" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
(2) prob.
<taxonomicName id="B788781130F5E676D0CE361CCCE85DD3" authority="Esb-Pet" authorityName="Esben-Petersen" authorityYear="1933" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="risi">
<emphasis id="63CFD0253392547F86F878C95D9AFD8E" italics="true">Leucochrysa risi</emphasis>
Esb-Pet
</taxonomicName>
, det.
<determinerName id="955AD9F7FD7D4ACD6A296E60C3E0B54B">P. Adams</determinerName>
80 [
<named-content id="977FDD385968FF99A6CF828CE4115A77" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="550035719C94667BA333A7BBB4ED3FD4" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
; collection site probably in
<collectingCountry id="F1A815FF60DCB6F74E48101F14BC3957" name="Panama">Panama</collectingCountry>
; abdomen in poor condition; pieces of exoskeleton in attached micro-vial]
</materialsCitation>
.
</paragraph>
<caption id="AAF3940259EC062C9BAE8A1273343157" ID-DOI="10.3897/zookeys.1226.140386.figure1" ID-arpha="9AA5B356-D858-5732-A8CF-D11F85882453" httpUri="https://binary.pensoft.net/fig/1244337" startId="F1">
<paragraph id="0AE193A56488191AC37C8AA9598843C3">
<label id="26DEB691047246B84E92C10A9B96FCF6">Figure 1.</label>
</paragraph>
<paragraph id="FC78C7706BC58385AE7B86E4A839E493">
Head of
<taxonomicName id="F7993254D7B06C69DAA83DBA7C974890" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="80F93E2FF548B9120AC632A294C8A630" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D139ECE1EF0733C9503F67252A2AB739" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, male, Nicaragua, Río San Juan, Bartola (
<named-content id="14D06B31D9318BCF4221EEA92EE4E4FC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="069434616D592935150EFB96A01E68C1" bold="true">a</emphasis>
frontal
<emphasis id="8E909897B99363AD53BC45B288DF9A9B" bold="true">b</emphasis>
head, prothorax, dorsal
<emphasis id="E1DE1E721116295B26462C70F9404012" bold="true">c</emphasis>
head, frontolateral
<emphasis id="CEDE74AD1EAC465D7736EB6E8B191C6E" bold="true">d</emphasis>
ventral. Width across head (dorsal, including eyes), 1.85 mm.
</paragraph>
</caption>
<caption id="5EED22BE4AB03F40B4316C91FB0D9A49" ID-DOI="10.3897/zookeys.1226.140386.figure2" ID-arpha="145706FC-F4DC-5FC7-AE0B-3A5B98E9BAD3" httpUri="https://binary.pensoft.net/fig/1244338" startId="F2">
<paragraph id="55E9C2AF8985744914031CC1097A5484">
<label id="AC4796FFE8E7F2E79B52B7BB5C935BA2">Figure 2.</label>
</paragraph>
<paragraph id="EB3629D4F001EA3877284FB38BAEB09C">
External features of
<taxonomicName id="A4D45978987A8F318D454D87CFE5C1C3" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="12B69359D9204170C1E3A3B06FCF039B" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="778B0F9F55DA675BA720174CF815BDAB" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, male, Nicaragua, Río San Juan, Bartola (
<named-content id="3A8795954DFDD2D0AB89EF0F77FFCD32" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="05CFD906C7FE87B083D4ED109C01406C" bold="true">a</emphasis>
head, thorax, dorsal
<emphasis id="5C0A94054E161902771FAF7AA70510E0" bold="true">b</emphasis>
body, lateral
<emphasis id="408B646775BB4A6DE46ABA63CCF86AAB" bold="true">c</emphasis>
abdomen, dorsal
<emphasis id="F57919CD73C4F8CAD2A723937E90E949" bold="true">d</emphasis>
terminus, lateral
<emphasis id="A0E4DED0BF866AA79B4046B304A78A61" bold="true">e</emphasis>
protarsal claw, lateral. Width across head (including eyes), 1.85 mm; diameter of callus cerci, ~ 0.2 mm.
</paragraph>
</caption>
<caption id="DCDB17CFCA185732190381CDA449D989" ID-DOI="10.3897/zookeys.1226.140386.figure3" ID-arpha="CB3DBA85-94B0-5E6A-AD0C-C849C888619B" httpUri="https://binary.pensoft.net/fig/1244339" startId="F3">
<paragraph id="2FDF055213D0E0F18646AEE8436B0212">
<label id="88E3DD0A0EBEB7348BE609F5D8C33E52">Figure 3.</label>
</paragraph>
<paragraph id="CFB669186DE0D33281ED14235AD005C8">
Wings of two new
<taxonomicName id="8F3852A1D23AC8052104263ADC2EF47D">
<emphasis id="4F5434C8873DD8AE5C127ABFF28A9F89" italics="true">Santocellus</emphasis>
species
</taxonomicName>
<emphasis id="F7A5B57D0318A801E58D3D034F99CE1E" bold="true">a</emphasis>
<taxonomicName id="9A8F8DAB86A2607331DCCE4E32EAC015" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="A84AC9E580FFDAABA398C37B76276744" italics="true">S. legrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="982C0464D2EB966977D741E6720E2F41" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male, Nicaragua, Río San Juan, Bartola (
<named-content id="B40C41083F57F77D0DEC7B9A3980539D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
); forewing length, 18.7 mm
<emphasis id="3B4F88146371B83BAC217612805CF2C4" bold="true">b</emphasis>
<taxonomicName id="0033D9CB0F13371216E091FB68F020B3" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="helene" status="sp. nov.">
<emphasis id="088065A733878FFE08D8939AEA720896" italics="true">S. helene</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5FB5E24EFC4C66AFF0AEC14FA4B3A9AA" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male, Bolivia, Cochabamba (
<named-content id="3C82920FDD04D3C33119F3AA5A7E371D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings.
</paragraph>
</caption>
<caption id="488D0FFE89D00DF708C1FBF32023F667" ID-DOI="10.3897/zookeys.1226.140386.figure4" ID-arpha="A3F1271A-64B4-519A-A2FF-B46537295441" httpUri="https://binary.pensoft.net/fig/1244340" startId="F4">
<paragraph id="63468E638660F2456DE9E32DF38CDA58">
<label id="DDD0106FFFA632CABF294AEE45A8F9DB">Figure 4.</label>
</paragraph>
<paragraph id="196A80CE7830A6AE3539435B82459197">
Male abdomen of
<taxonomicName id="8CB48F53E24E5A882BD15C9A0EA070EE" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="59A7E6ECEA1B27C2A0259C45ACBA1EEE" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="7E4352A8327D917FA2E186CA7DC72600" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Nicaragua, Río San Juan, Bartola (
<named-content id="96680238429E30C2B4AF27082E0909F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="45B6EB1A56DA067931BE7E62CB093834" bold="true">a</emphasis>
dorsum of abdominal segments A 2 - terminus
<emphasis id="62C5F40D787EE6F20A91EA66B3FA26C7" bold="true">b</emphasis>
lateral view of abdominal segments A 6 - terminus
<emphasis id="5406F3382E6BA15AC989ED5D3CC24057" bold="true">c</emphasis>
dorsum of abdominal segments A 7 to terminus with gonarcus everted
<emphasis id="0C47530ED9C93DA689810EE2C1FEBD96" bold="true">d</emphasis>
lateral view of abdominal terminus. Length of T 8 along dorsal margin, 0.68 mm. Abbreviations: c. c. callus cerci; d. ap. dorsal apodeme of T 9 + ectoproct; gcr field of gonocristae on terminal membrane of S 9; gs gonarcus; S 8 + 9 fused eighth and ninth sternites; T 3, T 7, T 8 third, seventh, and eighth tergites; T 9 + e fused ninth tergite and ectoproct; v. ap. ventral apodeme of S 8 + 9.
</paragraph>
</caption>
<caption id="3F77222CFCAC6332B395152B1DAB1B9F" ID-DOI="10.3897/zookeys.1226.140386.figure5" ID-arpha="EF0E83CB-D3E2-51A9-9880-82191AEE7AA8" httpUri="https://binary.pensoft.net/fig/1244341" startId="F5">
<paragraph id="95A8F415BA8F0D5AD8512E634F8FE1B5">
<label id="CE0AF1CCBC5B6C570E68723ACF77B38A">Figure 5.</label>
</paragraph>
<paragraph id="617A969D2EF0DDFB91FB557B6EA4DE7A">
Male genitalia of
<taxonomicName id="741E251A2C8CE58A811AD711C3F468BD" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="CA9F71144945B4F69EB0F33F14292A9F" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="038AA51847290063E6D2D7CD8A2BAFA4" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Nicaragua, Río San Juan, Bartola (
<named-content id="5C55BC61F58BB6C6594842D696813747" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="1EAE126943813766C150CAEF107644B9" bold="true">a</emphasis>
gonarcal complex, dorsal
<emphasis id="3F2DEAF9B18DF6170D20425B28516918" bold="true">b</emphasis>
same, lateral
<emphasis id="339BFB3078B9FA84851BB61A7551DCDF" bold="true">c</emphasis>
same, frontal
<emphasis id="C0C213A10A536DDA4FCEF826199685BA" bold="true">d</emphasis>
same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus.
</paragraph>
</caption>
<caption id="573A5D01036F24997DC7B0E8D7D41057" ID-DOI="10.3897/zookeys.1226.140386.figure6" ID-arpha="BE95E231-2577-5ED8-90CA-D2902A2D7288" httpUri="https://binary.pensoft.net/fig/1244342" startId="F6">
<paragraph id="EAD5C6CB3124DA2C8CA9D52D615BCDB2">
<label id="049764046063BD382506580883DB9E1A">Figure 6.</label>
</paragraph>
<paragraph id="5DA78ED5A55BE6B2EBC4FA16EA060EED">
Cleared abdominal terminus of female
<taxonomicName id="C33A2174EDEF54F0AAAEE81662E81014" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
<emphasis id="09C03E83FBCF189EF22954F409CDF1B6" italics="true">Santocellus legrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F33C19E01C7B144A0B8E9B499D694F25" rank="species">sp. nov.</taxonomicNameLabel>
, allotype, Costa Rica, San José (
<named-content id="E1936CC4F5914E236864C92355DA9B9D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/san-diego-natural-history-museum" xlink_title="San Diego Natural History Museum">SDMC</named-content>
)
<emphasis id="96437A462AC07464135F91074E1DE66A" bold="true">a</emphasis>
segments A 7 to terminus, lateral
<emphasis id="E94B30CA372297B8898FC0F162202E3A" bold="true">b</emphasis>
genitalia, lateral
<emphasis id="B47A8A3AFAF3F191EC344049D4E22B8F" bold="true">c</emphasis>
subgenitale, distal view
<emphasis id="3F03AAA2B4AEF94D7EDF1FBE8E919E8D" bold="true">d</emphasis>
spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVCAE" type="description">
<paragraph id="4F5EA65E6BC15D9ACE20471B3F738A2C">
<heading id="F5EC7A11576127A840708CFFE8048BB1" reason="title">Description.</heading>
</paragraph>
<paragraph id="5A2F31ABB36E738AFE11F37F59683D3E">
<emphasis id="3B535E632A65A355CDF830E4D038357B" bold="true">Adult.</emphasis>
Medium-sized.
<emphasis id="51125A14FAA81F22A97CE4798620C672" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="779EAF1E0C8F4E0B268A2C0D60C63215" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a frontal b head, prothorax, dorsal c head, frontolateral d ventral. Width across head (dorsal, including eyes), 1.85 mm." figureDoi="10.3897/zookeys.1226.140386.figure1" httpUri="https://binary.pensoft.net/fig/1244337">1</figureCitation>
): Dorsum, including eyes 1.70 (F) 1.85 (M) mm wide; ratio of head width to eye width = 1.71 (M) 1.86 (F): 1; sutures not visible. Vertex raised, with small posterior fold. Antenna long (
<quantity id="57F3CB3C86EEE07313230A9AF6BBC94A" metricMagnitude="-2" metricUnit="m" metricValue="2.8" unit="mm" value="28.0">28 mm</quantity>
,
<emphasis id="0ECC3D346CF4B8FD2D9DC4FB85766114" italics="true">n</emphasis>
= 1), ~ 1.5 times length of forewing; scape large, with approximately equal length and width; length of flagellomeres (beyond # 5) ~ 1.61.8 times width, with setae in four rings; labrum emarginate. Torulus relatively large, extending slightly below mid-eye level. Frons with short to moderate length, pale setae, lower margin scalloped upward above clypeus (below toruli). Tentorial pits distinct, round, raised. Gena
<quantity id="985EAAC6BB8A9C833729448C3F68C0A9" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
long; ratio of genal length to distance between tentorial pits = 0.43: 1. Clypeus quadrate, except dorsal margin broadly curved upward below frons, distal margin relatively straight, pale mesally, dorsal surface rounded, slightly raised mesally, with sparse, short to medium length amber setae. Labrum tapering anteriorly, with bilobed frontal margin, distinct cleft between lobes moderately deep, lateral margins tapering, long setae basally, shorter setae distally. Labium with submentum slender, rounded ventrally, with rounded, slightly bulbous tip.
<emphasis id="17A77296916FFBDFEB56B1A28F9BDA61" bold="true" italics="true">Coloration</emphasis>
: Markings on vertex, frons dark brown to black with tinge of deep red; broad, scalloped stripe across frons; vertex with large pentagonal mark, cream area within; clypeus with slender stripe across midsection, not reaching margins; labrum with dark reddish brown tinge mesally. Antenna with scapes pale frontally, dark brown dorsally, frontolaterally; pedicel cream with tinge of red distally; basal three flagellomeres dark brown to black frontally, medially; remaining flagellomeres pale. Maxilla with distal three palpomeres dark brown to black, basal two palpomeres pale with black markings laterally. Labium pale, with distal two palpomeres black, basal one pale.
</paragraph>
<paragraph id="9A8BC0AD7C5F9F29FA91C0DDB375F49A">
<emphasis id="D0CD0264F4BFEFE3C05503839C7608F6" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="60AABD3721C8D0DCA9C9F493D887E1CC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External features of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a head, thorax, dorsal b body, lateral c abdomen, dorsal d terminus, lateral e protarsal claw, lateral. Width across head (including eyes), 1.85 mm; diameter of callus cerci, ~ 0.2 mm." figureDoi="10.3897/zookeys.1226.140386.figure2" httpUri="https://binary.pensoft.net/fig/1244338">2 a</figureCitation>
): Prothorax wider than long:
<quantity id="343CC4F684D055E973BCB184F08CB00C" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.3" metricValueMin="8.6" unit="mm" value="0.895" valueMax="0.93" valueMin="0.86">0.860.93 mm</quantity>
long,
<quantity id="36A4B404A17D14890186E9BBD3476ED2" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.27" metricValueMin="1.11" unit="mm" value="1.19" valueMax="1.27" valueMin="1.11">1.111.27 mm</quantity>
wide, ratio of length to width = 0.69: 0.78; setae scattered, long to medium length, thin, golden. Legs robust, of moderate length, with dense, short, golden setae; tarsal claws without basal dilation, but with two prongs.
<emphasis id="1164F2D14F58A0298EE4EEAC0D003B13" bold="true" italics="true">Coloration</emphasis>
: Pronotum cream mesally, dark brown laterally; anterior brown section broadly quadrate, extending about halfway across dorsal surface, with central section bearing cream spot; posterior section cream mesally, with brown stripe laterally. Mesonotum dark brown laterally, with patches of cream mesally on prescutum, scutum. Scutellum with large cream spot. Metanotum entirely dark brown dorsally. Thoracic pleuron, venter pale to light tan. Legs pale, mostly unmarked, with short to medium length, golden setae, few long, pale setae; terminal segment (base of tarsal claws) smooth, shiny, black dorsally, with pair of small (indistinct) swellings, two pairs of moderately long setae at tip (Fig.
<figureCitation id="8FAD11B261396FABD412CC369F5210BB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm." figureDoi="10.3897/zookeys.1226.140386.figure7" httpUri="https://binary.pensoft.net/fig/1244343">7 e</figureCitation>
). Tarsal claw moderately long, narrow distally, with single small spur; base slightly enlarged, without distinct dilation.
</paragraph>
<caption id="58B953CD7B3467F577B2D938A387D038" ID-DOI="10.3897/zookeys.1226.140386.figure7" ID-arpha="7CFF7022-2B48-5F71-BBDD-7EB3D9120226" httpUri="https://binary.pensoft.net/fig/1244343" startId="F7">
<paragraph id="DD954FBEA2E9743270A29F9F949F9B89">
<label id="CBF42682A2CA2DE7B1D2253086B19B83">Figure 7.</label>
</paragraph>
<paragraph id="A0C84561F56966858C907B976355B775">
Head and thoracic region of
<taxonomicName id="2B3D4A860544B0DA8F5BAB1A89A214E8" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="helene" status="sp. nov.">
<emphasis id="E0577F31D156A9B7D122F9B5250ACA2F" italics="true">Santocellus helene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="957878F1EE4E2DDB488B76618E18B9E7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male, Bolivia, Cochabamba, Alto Palmar (
<named-content id="BF8519144165995B810E8A48E2324B77" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">CAS</named-content>
)
<emphasis id="8BF4CB4203EEB801FAB0D84648E55DEE" bold="true">a</emphasis>
upper body (head, thorax) dorsal
<emphasis id="60F5E844692CDDCDB2E7C121CC02050C" bold="true">b</emphasis>
head, prothorax, lateral
<emphasis id="3C90B3ABCA15F3974EBE60CD3C0E3DE6" bold="true">c</emphasis>
head, frontal
<emphasis id="F9B9307B60FAC6FE4E7C45DFC6949233" bold="true">d</emphasis>
head, ventral
<emphasis id="7C95A1B64B1A38C9B694ADC75E2BF7A9" bold="true">e</emphasis>
tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm.
</paragraph>
</caption>
<paragraph id="6F43A93C114AA6FEC4D33C4458CFD4E0">
<emphasis id="EC32B8E7DE506C7F32A850681ACC38DE" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="906D168C8D438DCF16635BA729A0A1B3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings." figureDoi="10.3897/zookeys.1226.140386.figure3" httpUri="https://binary.pensoft.net/fig/1244339">3 a</figureCitation>
): Measurements: male left wing (
<emphasis id="FDD1B76F73856D33F2B60BF693A5DAEA" italics="true">n</emphasis>
= 12); female, right wing (
<emphasis id="A40C8EBEF95689ACC8D9FF983B9C16AD" italics="true">n</emphasis>
= 1). Forewing:
<quantity id="5E097E6789BAD4B8A29DDAB1A370C32B" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.8699999999999999" metricValueMin="1.73" unit="mm" value="18.0" valueMax="18.7" valueMin="17.3">17.318.7 mm</quantity>
long,
<quantity id="88D3146301DFBFE69E17E4E5B05CB5E8" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.9" metricValueMin="6.2" unit="mm" value="6.55" valueMax="6.9" valueMin="6.2">6.26.9 mm</quantity>
wide (maximum). Costal area moderately tall; tallest costal cell (# 9, 10)
<quantity id="16FF8B33F6ABED835C547B7CD4400B80" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
tall, 0.230.24 times maximum height of wing. First intramedian cell triangular, elongate,
<quantity id="EB9C36F0C99F5BB45F68C73458E69AA0" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
wide, ~ 0.70.8 times width of third median cell. Origin of first radial crossvein ~
<quantity id="15C872C5BE707FCDFC05BC56991A55E8" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
distal to origin of Rs; radial area (between
<collectionCode id="E2DE29BAF8F23A080726581916591D9B" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
and Rs) with single row of 13 or 14 closed cells; tallest radial cell
<quantity id="0FFC46DC4025D399483E29186E15C0A8" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
, 0.770.80 times taller than tallest costal crossvein. Three b cells, each full sized, slightly longer than tall; first b cell heavily marked on all sides. Two series of gradates; 810 inner gradates, 7 or 8 outer gradates; size of cells bounded by gradates fairly uniform, lengthening then shortening distally. Four to 5 b cells; last b cell wider than tall. Three intracubital cells, distal one open; first cubital crossvein located slightly basal to first mediocubital crossvein; icu 1, icu 2 each shorter than icu 3; icu 1 + icu 2 considerably longer than icu 3. Vein 1 A forked. Terminal margin with approximately 5 simple veinlets, 13 forked veinlets. Hindwing: 16.717.0. mm long, 5.0
<quantity id="8F53B92D7E5148C7C4470167F3C2459D" metricMagnitude="-3" metricUnit="m" metricValue="5.1" unit="mm" value="5.1">5.1 mm</quantity>
wide. Eleven to 14 radial cells (counted from origin of
<collectionCode id="53F91EC7A56F2DAC8D0E8C4A10EB079B" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, not false origin); t cell absent. Two series of gradates; 59 inner gradates, 7 outer gradates. Two to 4 b cells; 4 or 5 b cells beyond im 2. Terminal margin with 7 or 8 simple basal veinlets, 911 forked basal veinlets.
<emphasis id="3F420515D68630F7822919BFE04A57D8" bold="true" italics="true">Coloration</emphasis>
: Wing surface hyaline, with bold, dark brown markings. Forewing: Stigma with moderately large, dark brown spot basally, second, smaller dark brown spot distally; veins pale except when surrounded by dark marks: e. g., distal part of
<collectionCode id="028F424E60B21DBF25FFA8225D2A6C10" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
and Rs dark brown with surrounding area dark brown; slightly elongate dark brown mark at tip of wing; two dark spots in approximate center of wing; scattered dark brown marks along basal 2 / 3 of lower edge of wing; veins dark when within dark spots. Hindwing: stigma with one large dark brown mark; base of RP, basal b cell dark, within dark brown mark; distal section of cu (upper fork) dark; posterior margin of wing with dark brown section, including suffusion at intersections with marginal veinlets.
</paragraph>
<paragraph id="C84FC717A737BA9E269B8BA1729500B2">
<emphasis id="BE8C963C406D298B0C100FE77BAD1A0C" bold="true" italics="true">Male abdomen</emphasis>
(Figs
<figureCitation id="D621F1C95CFACD9DB9E2F560837B82A1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male abdomen of Santocellus legrandi, sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a dorsum of abdominal segments A 2 - terminus b lateral view of abdominal segments A 6 - terminus c dorsum of abdominal segments A 7 to terminus with gonarcus everted d lateral view of abdominal terminus. Length of T 8 along dorsal margin, 0.68 mm. Abbreviations: c. c. callus cerci; d. ap. dorsal apodeme of T 9 + ectoproct; gcr field of gonocristae on terminal membrane of S 9; gs gonarcus; S 8 + 9 fused eighth and ninth sternites; T 3, T 7, T 8 third, seventh, and eighth tergites; T 9 + e fused ninth tergite and ectoproct; v. ap. ventral apodeme of S 8 + 9." figureDoi="10.3897/zookeys.1226.140386.figure4" httpUri="https://binary.pensoft.net/fig/1244340">4</figureCitation>
,
<figureCitation id="204606CA70373F5D44ABA8B636ACA30F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a gonarcal complex, dorsal b same, lateral c same, frontal d same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus." figureDoi="10.3897/zookeys.1226.140386.figure5" httpUri="https://binary.pensoft.net/fig/1244341">5</figureCitation>
). Segments 13 not particularly long or slender; distal segments only slightly larger than basal segments (male and female). Tergites shallow, with slightly rounded margins; each tergite with distinctive brown marking having white center spot. Tergites, sclerites, pleural region with numerous, short to medium length setae, microsetae. Abdominal sternites moderately tall (S 6: ratio of length to width = 1.3: 1), with dorsal margins straight, slanting, spiracles not enlarged, without microtholi; S 9 with slightly long, robust setae distally, without microtholi. Callus cerci almost round (0.23
<quantity id="176F318EA7709FFD78E7DBE0FAFB1BB0" metricMagnitude="-4" metricUnit="m" metricValue="1.9" unit="mm" value="0.19">0.19 mm</quantity>
diameter), with 3335 long trichobothria; cupuliform bases of variable size [those within central section larger (~
<quantity id="05885583441D5A279ACE82A12688E4AE" metricMagnitude="-5" metricUnit="m" metricValue="2.6" unit="mm" value="0.026">0.026 mm</quantity>
) than those on periphery (~
<quantity id="826DBE146FADCA94A0A175826A99C797" metricMagnitude="-5" metricUnit="m" metricValue="1.4" unit="mm" value="0.014">0.014 mm</quantity>
)]. Dorsum of
<collectionCode id="A3AE5728F438E7FA85A462838FA7848A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct rounded distally, fused mesally, with very small to no cleft; no midline suture visible; with long setae, densest along distal margin; dorsal apodeme simple, straight, short, extending around proximal side of callus cerci. S 8 + 9 fused without suture, tapering to acute apex (lateral view); S 9 slightly darker than S 8; ventral apodeme extending across dorsal surface of S 8 + 9; distal margin of S 8 + 9 spatulate, with terminal membrane prominent; membrane bearing moderately heavy gonocristae in two broad (somewhat irregular) lateral patches, single, small mesal patch of several (~ 3) gonocristae.
<emphasis id="33BFF024980E9FE3BE057463F7416161" bold="true" italics="true">Coloration</emphasis>
: Dorsum of each segment pale with large black central mark; marks on
<collectionCode id="C5B365F7E9E9EC717DE97BA9C5862801" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
2
<collectionCode id="CB0E7DC28C4C97FB0903973234EAFD15" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
8 with prominent white spot in center of black area; mark on
<collectionCode id="D5440B1E1D64F2EB6B1FFC3848AEB8F2" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 without white spot; venter, pleuron white to cream. Setae golden, long; dorsal half of callus cerci light cream colored; ventral half black.
</paragraph>
<paragraph id="1C81746651BA58F8327099E28725F055">
<emphasis id="72F774EA6B01371ABDE698DDC7B08092" bold="true">
<emphasis id="1A8641650A14F842F945ECAE822436A7" bold="true" italics="true">Male genitalia</emphasis>
.
</emphasis>
Tignum, gonapsis, parameres, quadrate hood, pseudopenis, spinellae absent; S 8 + 9 fused, elongate, not extending beyond
<collectionCode id="0A4BB43DD13EC3CDDB6BEE76A070122A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct. Gonarcus broad, with very small gonocornua on frontolateral margin of gonarcal bridge, with relatively large, rounded gonarcal arms expanded forward slightly; mediuncus in two sections; basal section extending from below gonarcal bridge, with pair of internal apodemes dorsomesally, with dorsal surface membranous, broad basally, tapering distally; distal section of mediuncus extending perpendicularly from tip of basal section, distally with flange-like lateral arms partially surrounding mesal beak; gonosaccus beneath mediuncus, stiff, flat, laterally with a few gonosetae on chalazae.
</paragraph>
<paragraph id="CFC8D9F243F0F1AC9EA424043D7029A4">
<emphasis id="FDECD8E62DADDF433C2C5697EE78EB00" bold="true" italics="true">Female abdomen</emphasis>
(Fig.
<figureCitation id="925930F224076A468F1CAF0B559D31CD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cleared abdominal terminus of female Santocellus legrandi sp. nov., allotype, Costa Rica, San José (SDMC) a segments A 7 to terminus, lateral b genitalia, lateral c subgenitale, distal view d spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum." figureDoi="10.3897/zookeys.1226.140386.figure6" httpUri="https://binary.pensoft.net/fig/1244342">6</figureCitation>
;
<emphasis id="A67C427CC359E82C749FAA50A7BC0478" italics="true">n</emphasis>
= 2, both slightly teneral). Callus cerci round,
<quantity id="15FC2DEE3EEA4F149A8456A2A09C4B55" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
in diameter, with 3637 trichobothria of variable length; cupuliform bases also of variable diameter (
<quantity id="2A740DE877CCF0EC5DB9C77411922457" metricMagnitude="-5" metricUnit="m" metricValue="1.95" metricValueMax="2.9" metricValueMin="1.0" unit="mm" value="0.0195" valueMax="0.029" valueMin="0.01">
<geoCoordinate id="5340EC007A45DB9A52E8AC85FFED58B1" degrees="0.010" direction="north" orientation="latitude" precision="55" value="0.01">0.010</geoCoordinate>
<geoCoordinate id="BA8D89DAD9FF1E332EB0C6E3BDE15420" degrees="0.029" direction="west" orientation="longitude" precision="55" value="-0.029">0.029</geoCoordinate>
mm
</quantity>
). Dorsum of
<collectionCode id="F681F6FD6A4E180131AD2E3D276BA1E9" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct rounded posteriorly, tall (lateral view), extending approximately to lateral margin of gonapophysis lateralis. Sternite 7 with dorsal margin relatively straight; distal margin blunt, not particularly heavy, with setae mostly missing. Gonapophyses laterales slender, occupying approximately half of abdominal posterior margin; interior membranous area not greatly expanded or inflatable.
<collectionCode id="02C4B14EC64728C0760288ED6C03D50E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct truncate posteriorly, extending slightly below gonapophyses laterales (lateral view). Gonapophyses laterales occupying ~ 6070 % of posterior margin of abdomen.
</paragraph>
<paragraph id="5AD597955ADCA869F03B44DD9B67E11B">
<emphasis id="E90FB19B6CB2620B0B08E21B4A459064" bold="true">
<emphasis id="0CCABDB1FA0E5E2C414AA58BF6924D6D" bold="true" italics="true">Female genitalia</emphasis>
.
</emphasis>
Entire genital structure compact; spermathecal complex, bursal duct, base of subgenitale all contained below bursa copulatrix. Subgenitale broadly based, with distinct quadrate knob distally; distal section of knob bilobed, with broad, flat trough between lobes; lower frontal region of subgenitale with elongate, downward-projecting, snout-like protuberance (knob). Spermatheca round, doughnut shaped, with broad invagination, distal end with relatively large tapering, sclerotized, coiled and curved tubule (velum), with slit along most or full length, apparently opening to bursa; velum ~
<quantity id="9DFE066B383AB0E0AB52B7971906C83B" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
long, ~
<quantity id="C1093EF73509252C627CD23AE7DC37B5" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
wide at widest point,
<quantity id="7FCE60957664F21D4CC6F141E787E596" metricMagnitude="-5" metricUnit="m" metricValue="4.0" unit="mm" value="0.04">0.04 mm</quantity>
wide near middle. Spermatheca doughnut shaped,
<quantity id="85D2B1D12E4CC15981DA960704033CAA" metricMagnitude="-4" metricUnit="m" metricValue="3.8" unit="mm" value="0.38">0.38 mm</quantity>
in diameter,
<quantity id="4ADD240AB361B7218A436A9FE79038F1" metricMagnitude="-4" metricUnit="m" metricValue="1.6" unit="mm" value="0.16">0.16 mm</quantity>
deep, with invagination in center,
<quantity id="1F9AF4103C476DCD9949564C55F5AF8D" metricMagnitude="-4" metricUnit="m" metricValue="1.9" unit="mm" value="0.19">0.19 mm</quantity>
in diameter,
<quantity id="4D36603C99F525C6A09884F30C6F7611" metricMagnitude="-4" metricUnit="m" metricValue="1.1" unit="mm" value="0.11">0.11 mm</quantity>
deep. Spermathecal duct very faint. Bursa copulatrix folded, pleated, small, barely covering dorsum of spermatheca, with distal end tapered, attached directly to sclerotized spermathecal velum, without bursal duct; bursal glands not found. Colleterial complex, transverse sclerite not discerned.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAHAE" type="variation">
<paragraph id="D7F99A8C105947282565A8C41D1E5701">
<heading id="542B1C5DBCF52D666AB13371D3D41B1A" reason="title">Variation.</heading>
</paragraph>
<paragraph id="136B91833D6374DDF786C9B4A43B3DC3">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EFHAE" type="distribution">
<paragraph id="871227205FE428A67EAA7B53168868BF">
<heading id="A4BB431D459ABB4E8F183A0346716C5D" reason="title">Known distribution.</heading>
</paragraph>
<paragraph id="C7BD0F4725125158B267E4AAA33E8960">
Central America.
<collectingCountry id="33A8798D132A21C7297A4D74135C67A6" name="Belize">Belize</collectingCountry>
(
<collectingRegion id="FAF6B80D93C1A7EB820C7D8FC36C6395" country="Belize" name="Cayo">Cayo</collectingRegion>
),
<collectingCountry id="46D65A6062C929824ADC2088F5903711" name="Costa Rica">Costa Rica</collectingCountry>
(
<collectingRegion id="2FDF1D64EC46F3AD9DB99E7F44B509D8" country="Costa Rica" name="San Jose">San Jose</collectingRegion>
),
<collectingCountry id="DD93232CA7F50ECE09E07E5394141B47" name="Honduras">Honduras</collectingCountry>
(Colón, prob.),
<collectingCountry id="5424A8DCA844C76F946D980D7654A731" name="Nicaragua">Nicaragua</collectingCountry>
(
<collectingRegion id="2DCB313DE7C425B33419152CD9FE8DDB" country="Nicaragua" name="Rio San Juan">Rio San Juan</collectingRegion>
),
<collectingCountry id="114B9B4EA9B7C58C128C2025164AAD59" name="Panama">Panama</collectingCountry>
(
<collectingRegion id="1EF4B3071ADD525540B27C183BE6B981" country="Panama" name="Chiriqui">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,776 @@
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<mods:title id="CCEF778BC8071C2872580A68E6FCFAD2">Species discovery in Southern African bee flies (Diptera, Bombyliidae): A new species in the revised genus Enica Macquart, 1834</mods:title>
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<mods:name id="53778D1FBA09EDB727416DDAA8506070" type="personal">
<mods:role id="7BC8745FC4349203A9DCA15040C0539E">
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<mods:namePart id="A9661825B3E26853CB61938B38DA20F6">Rollinson, Lisa</mods:namePart>
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</mods:role>
<mods:namePart id="70265BCC647DBE87D8D8E666D825CDCE">Cabrero, Allan</mods:namePart>
<mods:nameIdentifier id="6A75344AE65E516B37A8FEF1400EA00E" type="ORCID">0000-0002-2935-3534</mods:nameIdentifier>
<mods:affiliation id="E7F98927078596E3AF3118FD15D2B403">National Museum of Natural History, Washington, D. C., USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8CC1BC0EEA93E5AF9D3FC7FD63567345">text</mods:typeOfResource>
<mods:relatedItem id="6F177688166DD71B773570192F0E8CAB" type="host">
<mods:titleInfo id="EC20B227A88BF6A7C3406369F3EA3353">
<mods:title id="2DA4400C517B3CBA7522FF3C4DFC34B6">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="494B9A1AEC6D83DF82A2A6BD91572D77">
<mods:date id="D2E90064F5F4BA8B48D3F4C167F59B00">2025</mods:date>
<mods:detail id="94B6E5B3C1762D7F88A242A6A2B08624" type="pubDate">
<mods:number id="E18E762B0FE0E92C5679AE722095094A">2025-02-07</mods:number>
</mods:detail>
<mods:detail id="9E8B93669598DAB755DE1EAD0F1802EC" type="volume">
<mods:number id="B2A0F99110CD0B8CFF585E9FCCFFDB57">66</mods:number>
</mods:detail>
<mods:detail id="03FE616DCC940C8A7C0651EE68FE630C" type="issue">
<mods:number id="8C6F5B33D07C6849D7A96ABF0C567526">1</mods:number>
</mods:detail>
<mods:extent id="56F44EB58781BDFEE72001DEA10A379F" unit="page">
<mods:start id="6A4185505C88F8F44683EC8FAA618960">73</mods:start>
<mods:end id="BCFE6282A5D0F2B33F61B00895C96C76">115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="51D2F82C3C01B0373CDCFEB8C16E00D8">journal article</mods:classification>
<mods:identifier id="82A7F3E244BF54DBB095BFDA14FCF28F" type="DOI">10.3897/afrinvertebr.66.129611</mods:identifier>
<mods:identifier id="53B17AC70BF6D7DFF0A4BDFB50641439" type="ZooBank">9A70A477-D862-40E5-A132-3227D96C3ECC</mods:identifier>
</mods:mods>
<treatment id="FD3D8AE9EE005C6FAD8B7889F23A0E3C" ID-DOI="http://doi.org/10.5281/zenodo.14834500" ID-Zenodo-Dep="14834500" ID-arpha="FD3D8AE9-EE00-5C6F-AD8B-7889F23A0E3C" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C" httpUri="http://treatment.plazi.org/id/FD3D8AE9EE005C6FAD8B7889F23A0E3C">
<subSubSection id="D49FE80E4739D9FF0D9ED384880C8B72" type="nomenclature">
<paragraph id="84DB800C68BEA9838953071AC856C928">
<taxonomicName id="E5CC0B1254CAEF1275CD3C2125CBBADB" authority="(Hesse, 1956)" authorityName="Rollinson &amp; Cabrero" authorityYear="2025" baseAuthorityName="Hesse" baseAuthorityYear="1956" class="Insecta" family="Bombyliidae" genus="Enica" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="syrticola" status="comb. nov.">
<emphasis id="4BCF46AC1B6D89A9C266AF6CE8E79EF6" italics="true">Enica syrticola</emphasis>
(
<bibRefCitation id="48A63ACBDA2C0A08A91D75B72E08A90A" author="Hesse AJ" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 27-46. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="078D2FB41F27B8D73CB1F12B0B97BB14" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FB1588B1E86D85E14C4EBACABE42A75B" type="description">
<paragraph id="B9822117BD05D4B11D9E76075E88DD0A">
<figureCitation id="CC0E5A6DBC0085D4206D6F413C46597A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Enica sporanthera ♂ SAM-DIP-A 002174 (A C) A habitus, dorsal view B habitus, lateral view C habitus, frontal view; Enica syrticola ♂ NMSA - DIP- 89516 (D F) D habitus, frontal view E habitus, dorsal view F habitus, lateral view. Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.66.129611.figure11" httpUri="https://binary.pensoft.net/fig/1244184">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQAAK" type="Taxon depository">
<paragraph id="6A54666511243E82F032EE3AAF95F2F2">
<heading id="07D12E29B558496E7F5D7ED52C0B63BD" reason="title">Taxon depository.</heading>
</paragraph>
<paragraph id="709C54B5CE2C4B7E1416315A8DFE7F5F">
ZooBank:
<ext-link id="15E0C79022C884D89DF34D6F187E573F" ext-link-type="uri" xlink_href="https://zoobank.org/5BF9C251-F8F2-45D7-9665-DFF5AD9387D2" xlink_type="simple">
https://zoobank.org/
<uuid id="EFB28F9D3CE86298A52A205C14EF5631">5BF9C251-F8F2-45D7-9665-DFF5AD9387D2</uuid>
</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2AAK" type="diagnosis">
<paragraph id="34570C06F864856DF308160D38D2228D">
<heading id="A99255B2F1F553ED7ADD07EA5108703E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="0E9E2980AE5AEE66C18D7AE6FA0C4E65">The species is distinguished from other species in the genus by the dark brown to black postcranium, which covers the dorsal and posterior surface of the head.</paragraph>
</subSubSection>
<subSubSection id="SECID0EABAK" type="description">
<paragraph id="86731E284B43127AC26546F285FE0985">
<heading id="1E0704283B0513634D129D146006FDD3" reason="title">Description.</heading>
</paragraph>
<paragraph id="6E1CA6629CC8AD7DAB3AA23FC7CC7B36">
<emphasis id="9B10F9FB0DE0BC3468C8499BC67BC740" bold="true" italics="true">Head</emphasis>
: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, with dark brown to black triangular spot extending to eyes, males: reddish-orange, much darker than head; frons black setose, with white scales, males no setation near eyes, females setose near eyes only dorsally; ocellar tubercle slightly raised, dark reddish-brown to black, black macrosetose, white or light brown scales posteriorly; occiput light and dark brown setose, white scales antero-dorsally, entire back of occiput dark brown to black; gena significantly darkened to reddish brown or black, extremely sparsely setose.
</paragraph>
<paragraph id="35DBA6F2BC5CCB0BD05A955265254E72">
<emphasis id="550A628BF52EA6BC9F05961398537E5E" bold="true" italics="true">Proboscis and maxillary palpus</emphasis>
: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, brown, brown setose.
</paragraph>
<paragraph id="B66F0A692703CBC4BD18DAF0E66BAAD1">
<emphasis id="C16F2C5A527F7E86E64ECA2D842A254E" bold="true" italics="true">Antenna</emphasis>
: black; scape creamy white or reddish brown, approximately as long as pedicel, black setose dorsally and ventrally, setae extending to tip of pedicel; pedicel black, short black setose dorsally; postpedicel black, cylindrical (same diameter throughout), longer than scape and pedicel combined, black scales dorsally; stylus reduced, only apical seta-like sensory element present, situated sub apically in cavity on postpedicel.
</paragraph>
<paragraph id="6839AC63FD560BED18024D98BA9F7CFB">
<emphasis id="D07E01609A0023E0A2E89351B01197AC" bold="true" italics="true">Thorax</emphasis>
: reddish brown and light orangish brown; scutum predominantly black, reddish on margins, dark scales centrally and white scales peripherally; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, long white flattened setose, black macrosetose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, white scales dorsally; metepimeron silvery grey pubescence, white setose dorsally; apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of light brown and white scales, black setose.
</paragraph>
<paragraph id="9D686AB98A618A15C2B7753B3ECF28CA">
<emphasis id="873ABE2CFE4484AAC1D60D7B18E5166E" bold="true" italics="true">Leg</emphasis>
: light orangish brown to dark brown, covered in scales, some regular setae present; pro coxa dark reddish brown, silvery grey pubescence, dull whitish yellow scales, thick black setose; pro femur dorsally dark brown, ventrally orangish brown, dark brown scales, black setose; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa dark reddish brown, white pubescence, dull whitish yellow scales, thick black setose; mes femur dorsally dark brown, ventrally orangish brown, dark brown scales, black setose; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa dark reddish brown, white pubescent, dull whitish yellow scales, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur dorsally dark brown, ventrally orangish brown, dark brown scales, brown scales dorsally basally, long and thick black setose; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.
</paragraph>
<paragraph id="8F5DD4427F2E640D1AB7BD4A9AA5A5E6">
<emphasis id="40DAF22895FDB7908A751790E19347BB" bold="true" italics="true">Wing</emphasis>
:
<quantity id="DF81A1CFF6038745FD672FE087E56ACA" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.14" metricValueMin="0.99" unit="mm" value="10.65" valueMax="11.4" valueMin="9.9">9.911.4 mm</quantity>
, microtrichia absent; membrane color pattern males: predominantly brown colored except for white tinted base of wing, areas around crossveins r-r, r-m, m-cu, base of cells r 2
<subScript id="583C7345513F16526103B9CAF53C0EB4">+ 3</subScript>
, center of br, proximal part of r
<subScript id="71E30F90F3E5F01936D69458B6946797">1</subScript>
, base of veins M
<subScript id="53EF0CF4FC547D895ACD5AE242E3DF56">2</subScript>
, split between M
<subScript id="E076DFAED2B66C5CB18E795A1E70DB8D">1</subScript>
+ 2 and M
<subScript id="2B6068204D6C7E77FA10AD3C93B3CF2F">3</subScript>
, females: darkly infuscated except for white tinted base of wing, r-r, r-m, base of r 2
<subScript id="73BB508561806A6CB0E50ED069533308">+ 3</subScript>
, base of m
<subScript id="459A2F643951DC40C96F712DDB644D96">2</subScript>
, m-cu, split between M
<subScript id="1BE5DD643CCC735102C8900A282D7D64">1</subScript>
+ 2 and M
<subScript id="A33620A278190F33EC3BC7E6F0795CA9">3</subScript>
; costal vein dark brown short setose, whitish clear setose proximally; R
<subScript id="1B8783BF07F8B52863DFB82B61C604D2">2 + 3</subScript>
distally distinctly arching anteriorly, r
<subScript id="3E30815EB4368467F80710E01C7486EB">1</subScript>
open; R
<subScript id="6EA2387B56784A776CD40921766FE8DF">4</subScript>
terminating anterior to wing apex, distinctly arching anteriorly; cell r
<subScript id="CCEA4DB2AE27668090FE1C8836B655B3">4</subScript>
open, proximally R
<subScript id="17B401246740C5FD3C99F24BD1FE55D2">4</subScript>
and R
<subScript id="982C1789E56E63A6B5E7C3FE2F95A944">5</subScript>
parallel, R
<subScript id="55A75359E6B406BB7791118054434C09">4</subScript>
strongly diverging anteriorly in distal 2 / 3; R
<subScript id="79E4CE1455E5FB755E3D14E3CC2CD208">5</subScript>
terminating posterior to wing apex; r
<subScript id="F9BA1B49337B730BA95CC5C743CC622A">5</subScript>
open; M
<subScript id="3BA40464E635041AAF0707110377BFF9">1</subScript>
terminating posterior to wing apex; cell d closed by base of M
<subScript id="7804FEC2D0B0FC431CCB012CB6ACAA67">2</subScript>
and m-m, r-m situated in distal ¼; m
<subScript id="77490C145B435D590D0C120277504AEA">3</subScript>
open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.
</paragraph>
<paragraph id="D8857A464DA7C6E3D1584433A8A65E4E">
<emphasis id="691BDB0B17F6B4C8137B426968F6215A" bold="true" italics="true">Abdomen</emphasis>
: shape ovate, predominantly black with at most a thin reddish brown area on posterior edge of each tergite; tergites smooth, setae with small sockets only; T 1 predominantly reddish-orange, some black, light yellowish-brown to light whitish-brown setose, scales; T 2 T 7 predominantly black, thin black setose, brown to light yellowish-brown and white scales, white scales on spots on posterior margins of T 25; S 18 predominantly light reddish-orange, yellow to white scales, thin black setose and long white setose; lateral margin black setae remaining close to abdomen, male T 8 white scales, black setose.
</paragraph>
<paragraph id="F6AE15F256D26B7979BE1A7D2C1517FC">
<emphasis id="73753D824041EE37F1AF90E16E5A041B" bold="true" italics="true">Terminalia</emphasis>
: Male terminalia drawings in
<bibRefCitation id="692DC0458E2953C7F1B37BDA8F53AA6E" author="Hesse" firstAuthor="Hesse" journalOrPublisher="South African Museum" pagination="27-46" refId="B8" refString="Hesse AJ (1956) Annals of the South African Museum. South African Museum 35: 2746. https://www.biodiversitylibrary.org/page/40844208" title="Annals of the South African Museum." url="https://www.biodiversitylibrary.org/page/40844208" volume="35" year="1956">Hesse (1956</bibRefCitation>
, p 43):
<ext-link id="13C0E683CA9F2F1E6096ECD97C082EF8" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/40844178" xlink_type="simple">https://www.biodiversitylibrary.org/page/40844178</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPEAK" type="material">
<paragraph id="A3E26549808595E65588A075C8DA558B">
<heading id="93C8999EF4570106410E85AC6D612179" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="3AFCC83F6271B1338B8622EF434AC28C">
<materialsCitation id="3F3E85416E2B56BEF75BA87761151BA9" collectingDate="1917-11-20" collectionCode="NMSA" collectorName="Roberts, A." country="South Africa" latitude="-32.09" location="Lamberts Bay" longLatPrecision="20" longitude="18.313055" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="308EB1C7FC70450E8F915F69C339D3D4" bold="true">
<emphasis id="547E95B1ECB349C9834BF22A36EC68EE" bold="true" italics="true">
<typeStatus id="1D5DBF87120BDF8F5A3785C0607D3660" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="11EA370B9A8732878DA155D59D12C7D2" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="3246560E79EDA6A4853E83C80D8F5884" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="642EC9D0AD2831047196C8807A535F63" count="1" type="male">1 ♂</specimenCount>
<location id="37101FD4CD65226AC9EE4D87F2F6BC23" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:37101FD4CD65226AC9EE4D87F2F6BC23" country="South Africa" latitude="-32.09" longLatPrecision="20" longitude="18.313055" name="Lamberts Bay" stateProvince="Western Cape">Lamberts Bay</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4EAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.313056,-32.090000]}">
<geoCoordinate id="46D49E1D306BC3E5A4DC954D43C05D39" degrees="32" direction="south" minutes="05" orientation="latitude" precision="15" seconds="24" value="-32.09">32 ° 05 ' 24 &quot; S</geoCoordinate>
,
<geoCoordinate id="6E1150EBB99A7947DC43AD86FBC4A85B" degrees="018" direction="east" minutes="18" orientation="longitude" precision="15" seconds="47" value="18.313055">018 ° 18 ' 47 &quot; E</geoCoordinate>
</named-content>
;
<date id="975D2B379B610A696869EDC9D84304A3" value="1917-11-20">
<collectingDate id="BEF33630801CD801D07DA406EEEF4DFB" value="1917-11-20">20 Nov. 1917</collectingDate>
</date>
;
<collectorName id="231E1CFDD05BD0767364B6FC13EB6441">Roberts, A.</collectorName>
leg.;
<named-content id="98F29D166315701FBF51DAA6AC7DA0A2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="8D3A668AD5FBEC24B9A855BA2B90DBB5" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 048211,
<named-content id="E3E1C80F3A3C5A45EE02F1C685F37A1D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="3D651108BA5D7B7FB00FFB53BFF8BCBB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
.
</materialsCitation>
</paragraph>
<paragraph id="10DF2AEAA1328BD1138073ACA7D3157D">
<materialsCitation id="EF5CADD39C48247F5361F05FF5F28433" collectingDate="1947-09" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-31.616667" location="Calvinia" longLatPrecision="20" longitude="19.366667" municipality="Augusfontein" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="F559889C40E40E2C93F7E8E377AEE200" bold="true">
<emphasis id="2455C0EAA9D59E67007C732A3C8A8950" bold="true" italics="true">
<typeStatus id="D1E127A14BC68F3F9F14516560CD805B" type="paratype">Paratype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="78AEE3A266DD0FD417EDD22DCC52C13F" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="0FDC6F6A9D78BE1AD9CEF6379C8614B6" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="75D69900ADC260E897E7A3CC041126FE" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="C76C429D7DA846A70B77CE91281D72A4">Augusfontein</collectingMunicipality>
(= Augustfontein),
<location id="54FEDC24CFCD46EAB02BEAB3848F54DE" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:54FEDC24CFCD46EAB02BEAB3848F54DE" country="South Africa" latitude="-31.616667" longLatPrecision="20" longitude="19.366667" municipality="Augusfontein" name="Calvinia" stateProvince="Northern Cape">Calvinia</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EUFAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.366667,-31.616667]}">
<geoCoordinate id="DC82757E2EB427B3B254D5F76072C02D" degrees="31" direction="south" minutes="37" orientation="latitude" precision="15" seconds="00" value="-31.616667">31 ° 37 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="6662089AE8F97E81F50D69409D8731B0" degrees="019" direction="east" minutes="22" orientation="longitude" precision="15" seconds="00" value="19.366667">019 ° 22 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="399F20C5166E223302F8B3A8AC544803" value="1947-09">
<collectingDate id="CA3C65C213F21101DF6E4D7237BCF91E" value="1947-09">Sep. 1947</collectingDate>
</date>
; SAM
<collectorName id="478C1FEB3F5D3ACB004530E47743732D">Museum Staff</collectorName>
leg.; SAM-DIP-A 002179,
<named-content id="5BD0FEECF399339373DE57028AE1E32A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="F5F5D57F1EEB0F002AB322653F4D26B4" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="EAA715D8B157E7097ABD02CC0125F41A">
<materialsCitation id="C3741647B02CB428CEDF7DD8AB47B508" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<collectingCountry id="4A96759F35755B4558197CCE8809C629" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="FAEAC97BF1358C2EC6CDD1F43A50D1B9" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="4DD3FF8CE1A758E416E40693861E9FD5" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="FDD078024BCE1C3695A57BDCF8488949">Elands Bay</collectingMunicipality>
,
<location id="FC194429CB03D9284CCA57619C090043" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:FC194429CB03D9284CCA57619C090043" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
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<geoCoordinate id="36BF4316D599482BB417733CB08A247A" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="1DF76CA90282AA0EC6FF31D481D97942" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="B9AF217590D2669F6FD4FFD3B1A54F30" value="1947-10">
<collectingDate id="3CC4E758349FDB47AD0F476CDBE2775D" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="1D531798C1CBBDDA05E430585436DC66">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="B4E25DF94D9536114DCC43B7C9FFEE68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="2E2C3420C4EB1B060ED226E5CF848408" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="EEFF776000C3D7E07363E26C84F24113" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="5E0E6733CF844A6CBF683F69F51888A1" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="163644C28BAE3D0A07D92B03226E9F29">Elands Bay</collectingMunicipality>
,
<location id="3CBF29DEB07602C423E9D7062AB9B325" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:3CBF29DEB07602C423E9D7062AB9B325" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOGAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="EFB628D8CF79A29144E051A7C7C10908" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="2126CEA973135BED286063134CD39F14" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="CDB939AA224D7435C77F4CED47C0DEF7" value="1947-10">
<collectingDate id="1353EAD3B122CA166D444FC9CD265DA6" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="954A8FEF9702910D77738546671BBB2E">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="5F2452FD83D0CCA7BD1FA80898421D96" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="4C02CD612D3D0027549E3B6138316B3B" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="ADAA5C1466B5E81F9BFB2D8DCB20331C" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="F275E5C60A14670BAC92616DF70FDB79" count="1" type="female">1 ♀</specimenCount>
<collectingMunicipality id="1E746598FFB0ECBFF3D94D0A8897C4FA">Elands Bay</collectingMunicipality>
,
<location id="009C4CC56BF823F9723BA6009BBD4002" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:009C4CC56BF823F9723BA6009BBD4002" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1GAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="D0F3E7ECD32A66171C730D2D3FEE30BA" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="DADCE62840C6BD77184F8E9F00DDC87A" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="C951C3B988C0061C3AF7F90945FF2316" value="1947-10">
<collectingDate id="386F0084008AA7A9EEC6C5FE6DD69BBE" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="813C2F7FCD8868048D88735891FE7AED">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="8F1E3551A4E9224618D4B112CB4FB00F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E8D3FCA774FE0EF3DC11E98495CAE244" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="630ABA286530BC5B8B4E1F520196193C" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="4FB9A98ABF650B044D671A874ACC7735" count="1" type="female">1 ♀</specimenCount>
<collectingMunicipality id="C41E905ED0E43690CE59F6975C3581C9">Elands Bay</collectingMunicipality>
,
<location id="263C314C09EE998CB4720056E221D817" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:263C314C09EE998CB4720056E221D817" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGHAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="15A09EA6AEA15DAA58D3BA0A064BEA00" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="AD4DE4FCE98312BF5B163B7429446B7E" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="913AC980D491435EEC0ADF57ED4EF2F6" value="1947-10">
<collectingDate id="243DA7EEDD03DF85B74562C5ED97AC85" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="621D67E39D2E151D44E5938B8FBD1178">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="66D1092B175C4E9978D2121E2BD63500" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="AF75A13447E2736607FA18902A19DF32" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="F5321794109CCF513B4EECEF47E3B92C" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="0D8DD72B0CDA82AEA134BC2600430B30" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="EC96C964B24E133FB1D77B930BC19E2E">Elands Bay</collectingMunicipality>
,
<location id="489BA045782531ADF070FDE18B2C9545" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:489BA045782531ADF070FDE18B2C9545" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESHAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="B89E15104D83C5D2E4D08DCCDB7E622C" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="8D4668676F6C426BF925FBC45E7B0055" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="F0EC0290F178ADA32AA3092CE51B6418" value="1947-10">
<collectingDate id="7AA76BA5FC9E158C3C865748BAECB38F" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="721B13755CED01F29DC777CB762AD897">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="0F2377172946DA363E7D3F828F2D9E0A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="706141391912E42C05EC888F60DAED94" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="5D087AE26C072D9BC794892E0E5F6E16" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="CB3A896C5CCAC2DF75B293A180B1EF0C" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="CF79F5E4D80F996E2272895918AE70F3">Elands Bay</collectingMunicipality>
,
<location id="31BFD5688B6B676C85E3875A9EF59B1B" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:31BFD5688B6B676C85E3875A9EF59B1B" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5HAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="526CC15ACA48305217F4344195B852AF" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="DE642761774D1B0FF842B59E9CB1F80C" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="173AD88B245738C10B8F17233B9A50C9" value="1947-10">
<collectingDate id="702C97B3F765B78B890B5123169000F7" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="0E5178DA2A98BB2AC304D09E33E64B07">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="A8FF68AAD4E5828068867F6BB4BE222E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E4B5EAC707F7753BA2302A47A5FB1A64" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHIAK" type="material">
<paragraph id="DECB766160311BDFEE379166C0DD00D1">
<heading id="9F8392A3B77A50E87EF94565EDC1D7F1" reason="title">Other material.</heading>
</paragraph>
<paragraph id="2A00894BF84701C902294A5383FC8EAB">
<materialsCitation id="EFD920D5AF16353BF73B2F55E94B21FD" collectingDate="1947-09" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" county="Augusfontein" latitude="-31.616667" location="Calvinia" longLatPrecision="20" longitude="19.366667" municipality="Augustfontein" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape">
<collectingCountry id="41C503B2BE13E87223945C43F1A19186" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="5436B425CEB69EFAD8B83E7F49F4BA9E" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
<specimenCount id="F29F9F28181C6457D70093B9C2D12702" count="1" type="male">1 ♂</specimenCount>
<collectingCounty id="FD4170932368C9105C36549A186F0E64">Augusfontein</collectingCounty>
(=
<collectingMunicipality id="EA7B591FDD4D5FEEE0F5A96007419F3F">Augustfontein</collectingMunicipality>
),
<location id="6A756039644FA86D1CA5E135159BA95F" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:6A756039644FA86D1CA5E135159BA95F" country="South Africa" county="Augusfontein" latitude="-31.616667" longLatPrecision="20" longitude="19.366667" municipality="Augustfontein" name="Calvinia" stateProvince="Northern Cape">Calvinia</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQIAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[19.366667,-31.616667]}">
<geoCoordinate id="D4FC733E77FC1281E801737274A69ABA" degrees="31" direction="south" minutes="37" orientation="latitude" precision="15" seconds="00" value="-31.616667">31 ° 37 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="B9FF07E454B657C0F8BF4CC7035802AD" degrees="019" direction="east" minutes="22" orientation="longitude" precision="15" seconds="00" value="19.366667">019 ° 22 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<date id="C5D560F5350ABD5E6379A429F8F91DCA" value="1947-09">
<collectingDate id="34B8A88E4ED8F6A0EA4F4ED0CDE18E90" value="1947-09">Sep. 1947</collectingDate>
</date>
; SAM
<collectorName id="DD9EEDC220E8FA3B1E4F52B9FC9029AF">Museum Staff</collectorName>
leg.; SAM-DIP-A 002179,
<named-content id="97073527D248CFEA63AC85CED1C6D429" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="92B8F319CE95FB4FC695546B02157B18" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="2BE2B627ED09DD3CB53161A8DBB90554" collectingDate="2012-09-16" collectionCode="NMSA" collectorName="Londt &amp; Jason" country="South Africa" elevation="185" latitude="-29.31111" location="Port Nolloth" longLatPrecision="20" longitude="17.108889" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="9195709ADCBCD69E9ADFE288C3732BEC" count="1" type="female">1 ♀</specimenCount>
<location id="B04EE1990D0233E126E300D9C3323587" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:B04EE1990D0233E126E300D9C3323587" country="South Africa" latitude="-29.31111" longLatPrecision="20" longitude="17.108889" name="Port Nolloth" stateProvince="Northern Cape">Port Nolloth</location>
,
<quantity id="3E23B1AF644E6D2C7F99771243748A94" metricMagnitude="4" metricUnit="m" metricValue="2.3" unit="km" value="23.0">23 km</quantity>
E;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3IAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.108889,-29.311111]}">
<geoCoordinate id="48657CAEB1F89B65D2A5CF603F084BA9" degrees="29" direction="south" minutes="18" orientation="latitude" precision="15" seconds="40" value="-29.31111">29 ° 18 ' 40 &quot; S</geoCoordinate>
,
<geoCoordinate id="87AE6DDBE24C92C1B3199AEB25114928" degrees="017" direction="east" minutes="06" orientation="longitude" precision="15" seconds="32" value="17.108889">017 ° 06 ' 32 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="8D1A4390951A67F3BBE35642252B433F" metricMagnitude="2" metricUnit="m" metricValue="1.85" unit="m" value="185.0">
<quantity id="CEB065EC9770CD543AFFB64B7D3B708B" metricMagnitude="2" metricUnit="m" metricValue="1.85" unit="m" value="185.0">185 m</quantity>
a. s. l.
</elevation>
;
<date id="2F4B2D51F07DAD5EA847732A081EF59A" value="2012-09-16">
<collectingDate id="DB7D624963FE529CF1130F682F613726" value="2012-09-16">16 Sep. 2012</collectingDate>
</date>
;
<collectorName id="B4444A6FC77705F98CBAF26B40A64CB1">Londt</collectorName>
,
<collectorName id="C9C4FFBF76F2D8A3E744761B093E97BD">Jason</collectorName>
leg.; sandy road verge;
<named-content id="B902D5A09F030A7DBD03B6BC72E26A94" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="C6A0E3E96F0B03A3BA69F03A747795AE" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 206183,
<named-content id="D06EA541E0EEA06FEBC4DFE1733E334F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="2805A7CFC7AAB38E281DC2F6F5DCB11F" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="E58B97832D2FC0BA2B90F3C2DA6961B8" collectingDate="1989-08-31" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Stuckenberg &amp; Brian" country="South Africa" elevation="50" latitude="-28.808332" location="Richtersveld" longLatPrecision="20" longitude="16.633333" municipality="Alexander Bay" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="12B937A94C4F609E6AE0A8C9915999F7" count="1" type="female">1 ♀</specimenCount>
<collectingMunicipality id="FC9EF0CC6D30F50175A7E809CE5E45DD">Alexander Bay</collectingMunicipality>
,
<quantity id="68F41185D27FF237D0F1E7D2079CA5A2" metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
SE,
<location id="1249A91483B38A291AB4FA251C75AAE9" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:1249A91483B38A291AB4FA251C75AAE9" country="South Africa" latitude="-28.808332" longLatPrecision="20" longitude="16.633333" municipality="Alexander Bay" name="Richtersveld" stateProvince="Northern Cape">Richtersveld</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMJAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[16.633333,-28.808333]}">
<geoCoordinate id="B499D4742FF47E4ACB4CC966A68428CB" degrees="28" direction="south" minutes="48" orientation="latitude" precision="15" seconds="30" value="-28.808332">28 ° 48 ' 30 &quot; S</geoCoordinate>
,
<geoCoordinate id="93E9B1A3228C8CBC79A7DAC70B65F2FE" degrees="016" direction="east" minutes="38" orientation="longitude" precision="15" seconds="00" value="16.633333">016 ° 38 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="F106FE0EEF34AC2E32022241879A7851" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<quantity id="4EE1718B47033C6DC9471198C1435496" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
a. s. l.
</elevation>
;
<date id="86F5A5FC79037E94D07C7771C517C5FE" value="1989-08-31">
<collectingDate id="37BDD7EAC6AEDD25800FAC0127FA8A26" value="1989-08-31">31 Aug. 1989</collectingDate>
</date>
;
<collectorName id="0DF7DED8FB611BDCFFB3604891909E2D">Londt</collectorName>
,
<collectorName id="1F705AD9853DF415AFC2D3E7476AA844">Jason</collectorName>
,
<collectorName id="FD4F8FFD6F2F3BB05DE761F49F1D386E">Stuckenberg</collectorName>
,
<collectorName id="EDD5B70544AA80B5E176FE58BC387AE0">Brian</collectorName>
leg.; costal strandveld, flowers;
<named-content id="F7EB209CA195866C9AEE9EA07D236B83" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="BDEB8E8DD12FFEBEA63722130CC44FBB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 042858,
<named-content id="F9E8AB9763A28895F2BFFC8DDFFCAAB3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="5EA0FA6528DCE25F7A639EFE5787C6BD" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="82AE1809CF2B4078295B0040143E9FED" collectingDate="1989-09-03" collectionCode="NMSA" collectorName="Londt &amp; Jason &amp; Stuckenberg &amp; Brian" country="South Africa" county="Richtersveld" elevation="350" latitude="-28.316668" location="Sandy valley" longLatPrecision="20" longitude="16.916666" municipality="Grootderm" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="3F35C9598F3CFF29B7684476DFF83697" count="1" type="female">1 ♀</specimenCount>
<collectingCounty id="1DE5601DFC78B894E3A8591C0CB6157D">Richtersveld</collectingCounty>
,
<collectingMunicipality id="4258DE16A64CF25A1F0104B59BC92521">Grootderm</collectingMunicipality>
<quantity id="9F0A3CA886077479B81C348D9A965C22" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
NE;
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<geoCoordinate id="3399FFE0EF794C186FA11046D35DA1E3" degrees="28" direction="south" minutes="19" orientation="latitude" precision="15" seconds="00" value="-28.316668">28 ° 19 ' 00 &quot; S</geoCoordinate>
,
<geoCoordinate id="E9EB0DB028420355F8188569021527A7" degrees="16" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="16.916666">16 ° 55 ' 00 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="C47575A5601FF9C3A9D2E7BE19E16F50" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity id="4FC6D86BB0C6F86FA11F286B927AE474" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
a. s. l.
</elevation>
;
<date id="DE89084F1804123A38E98DB30AC6F3EA" value="1989-09-03">
<collectingDate id="13BD7F9504E7E582ABBB3A150B89D562" value="1989-09-03">03 Sep. 1989</collectingDate>
</date>
;
<collectorName id="E05FBD53578C0FC4F26C4E6F5611988C">Londt</collectorName>
,
<collectorName id="C6BFD13D2904AE99E7A136A988E8D5C1">Jason</collectorName>
,
<collectorName id="85D4D795609B601DB70D15DD0EDE7081">Stuckenberg</collectorName>
,
<collectorName id="E3E56A2125765C91646D976306C34411">Brian</collectorName>
leg.;
<location id="3540A2D04EFD99BE8849CF35DECAF0E4" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:3540A2D04EFD99BE8849CF35DECAF0E4" country="South Africa" county="Richtersveld" latitude="-28.316668" longLatPrecision="20" longitude="16.916666" municipality="Grootderm" name="Sandy valley" stateProvince="Northern Cape">Sandy valley</location>
below a rocky hillside;
<named-content id="71C67377D7348C9DAD8AD1313A999A6B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="ECE4643A1BD1B9307C6AC465C7A433DD" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 042859,
<named-content id="EE5760D19B050EE8CEB74E0E645F2BCA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="A2258CB6DAF71BB0A9B1760EAAB7653D" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="CBD9B557510A4C41299D976FDAC143F4" collectingDate="1950-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-30.388332" location="Wallekraal" longLatPrecision="20" longitude="17.508888" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Northern Cape">
<specimenCount id="F1D488FFBED740FC23F7905DD71BE583" count="3" type="female">3 ♀</specimenCount>
<specimenCount id="F19448D7A8699C03A1B419B750C8464E" count="3" type="male">3 ♂</specimenCount>
<location id="F1E54ED7B4F2B8E14081C2C38F96AC31" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:F1E54ED7B4F2B8E14081C2C38F96AC31" country="South Africa" latitude="-30.388332" longLatPrecision="20" longitude="17.508888" name="Wallekraal" stateProvince="Northern Cape">Wallekraal</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMKAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[17.508889,-30.388333]}">
<geoCoordinate id="DC9F5BF28BCEBE703EF4ABE221DE66C2" degrees="30" direction="south" minutes="23" orientation="latitude" precision="15" seconds="18" value="-30.388332">30 ° 23 ' 18 &quot; S</geoCoordinate>
,
<geoCoordinate id="81F16BA595D5ED2A591D763E1B006AE0" degrees="017" direction="east" minutes="30" orientation="longitude" precision="15" seconds="32" value="17.508888">017 ° 30 ' 32 &quot; E</geoCoordinate>
</named-content>
;
<date id="7E79B45449759D06B803F4E3F8FE443A" value="1950-10">
<collectingDate id="D1C4C2DE9706147B8F04B457C3CCB1BB" value="1950-10">Oct. 1950</collectingDate>
</date>
; SAM
<collectorName id="D3B562370B96C8F6B73D34E1179F5685">Museum Staff</collectorName>
leg.; SAM-DIP-A 002180,
<named-content id="ECB26D6B51718B3017C1E9C4CBE525E5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E2AEE69CF2AC27978293BD4A4466567B" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="B5A25A07A23DC70A7B0EA3B735B43FB8">
<materialsCitation id="7A8C8981F31098D3E0150B8C23E6F85E" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Western Cape">
<collectingCountry id="7726CE29477917E1AAE1AE63F259F396" name="South Africa">South Africa</collectingCountry>
<collectingRegion id="1A04E11DCE96C5F50634826046BD4CDA" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
<specimenCount id="D37996B3F954592FEF72A43EFA79230B" count="1" type="female">1 ♀</specimenCount>
<specimenCount id="0B2568031D79CB9E321BF9A7D82EE2F5" count="5" type="male">5 ♂</specimenCount>
<collectingMunicipality id="A0A1E62AAA35CA731F9CBBA73086FDF1">Elands Bay</collectingMunicipality>
,
<location id="5E3D2559CEE7CEA5D53A3D9A6EE8A669" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:5E3D2559CEE7CEA5D53A3D9A6EE8A669" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1KAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="2209C761791BDAA00B621035A7ACC073" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="A3FAEA6568021F295D194CCA1260E120" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="31C396B7E5DF3830E7EDEC6D7676564F" value="1947-10">
<collectingDate id="9BB0B14450EB0D7CE360725413CAC709" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="F1B6F145C0759866EA8C3811981677CD">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="B13C8FE162DF104840ACD62EBEB98FB6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="49D040A591F3C4A56565C7509B1C4AFD" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="C41AAB27F72788DC5CC6828A553359FC" collectingDate="1947-10" collectionCode="SAMC" collectorName="Museum Staff" country="South Africa" latitude="-32.31222" location="Leipoldtville" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="5943FF67D3A50B816CBDF9338A562800" count="1" type="male">1 ♂</specimenCount>
<collectingMunicipality id="33D34DA37DB145F838BC12F3370A6F7E">Elands Bay</collectingMunicipality>
,
<location id="230D2F74A0606921D0191D37DE57A0F1" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:230D2F74A0606921D0191D37DE57A0F1" country="South Africa" latitude="-32.31222" longLatPrecision="20" longitude="18.341389" municipality="Eland's Bay" name="Leipoldtville" stateProvince="Western Cape">Leipoldtville</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGLAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.341389,-32.312222]}">
<geoCoordinate id="E152EE5BE6B846B44BA91ECC493FF653" degrees="32" direction="south" minutes="18" orientation="latitude" precision="15" seconds="44" value="-32.31222">32 ° 18 ' 44 &quot; S</geoCoordinate>
,
<geoCoordinate id="D25F3AF8EAF7EBBF57605DE6C8E32708" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="29" value="18.341389">018 ° 20 ' 29 &quot; E</geoCoordinate>
</named-content>
;
<date id="B8DEB8C1605A46BE019C555D14BAA49C" value="1947-10">
<collectingDate id="9F6C189478135332FC51409EBDD21873" value="1947-10">Oct. 1947</collectingDate>
</date>
; SAM
<collectorName id="5977688BF142E75C66CC72EBB526E505">Museum Staff</collectorName>
leg.; SAM-DIP-A 002177,
<named-content id="D60A9C24CCDC0816F65BD30E45338D13" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="0ED2899604FD16FBEDC6196956E6BB77" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="CA712966B76F380B5CCCBEBFEC0563A9" collectingDate="1967-10-05" collectionCode="SAMC" collectorName="Jakkals R &amp; Gess, F." country="South Africa" latitude="-32.085556" location="Strandveld" longLatPrecision="20" longitude="18.316666" municipality="Jakkalsrivier" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="DD1AA6BDC20187024FA9EE2192CA05E7" count="1" type="male">1 ♂</specimenCount>
<collectorName id="37ECC426B7848A4911CBA92D09C88C31">Jakkals R</collectorName>
(=
<collectingMunicipality id="671C0DD3717013BBEEE47A8B4F287256">Jakkalsrivier</collectingMunicipality>
)
<location id="BABADDA9158DADBC597423181F4DE237" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:BABADDA9158DADBC597423181F4DE237" country="South Africa" latitude="-32.085556" longLatPrecision="20" longitude="18.316666" municipality="Jakkalsrivier" name="Strandveld" stateProvince="Western Cape">Strandveld</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESLAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.316667,-32.085556]}">
<geoCoordinate id="BB781C6783B85C815358EA0BD2A8DB42" degrees="32" direction="south" minutes="05" orientation="latitude" precision="15" seconds="08" value="-32.085556">32 ° 05 ' 08 &quot; S</geoCoordinate>
,
<geoCoordinate id="47A0FB38DA9188B7CC1F321FEAD9C494" degrees="018" direction="east" minutes="18" orientation="longitude" precision="15" seconds="60" value="18.316666">018 ° 18 ' 60 &quot; E</geoCoordinate>
</named-content>
;
<date id="AB58DF307B4F120FCAA79408EBFADA94" value="1967-10-05">
<collectingDate id="58C01328137F1B0C315455DB2BF062DF" value="1967-10-05">05 Oct. 1967</collectingDate>
</date>
;
<collectorName id="69302FC79A643A9A96B4548E5EDE86E2">Gess, F.</collectorName>
leg.; SAM-DIP-A 016204,
<named-content id="8E3ADDA1538F4337B491B517BDCB3A1D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="E710D40E8212AB123D452914142CE217" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="F65A84E26C3815796B420A2501FEAA90" collectingDate="1977-10-09" collectionCode="NMSA" collectorName="Miller, R. M." country="South Africa" latitude="-32.78611" location="Laaiplek" longLatPrecision="20" longitude="18.16111" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="2EF3ED0F27240A94646964EFCB3D76AE" count="1" type="male">1 ♂</specimenCount>
<location id="8FAF83E6DC06260DCF1F66A485D6105C" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:8FAF83E6DC06260DCF1F66A485D6105C" country="South Africa" latitude="-32.78611" longLatPrecision="20" longitude="18.16111" name="Laaiplek" stateProvince="Western Cape">Laaiplek</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5LAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.161111,-32.786111]}">
<geoCoordinate id="0F2AF24E8DF35300CE754DA87F83D5DF" degrees="32" direction="south" minutes="47" orientation="latitude" precision="15" seconds="10" value="-32.78611">32 ° 47 ' 10 &quot; S</geoCoordinate>
,
<geoCoordinate id="91EC9A8CF3CF28B1FC4AF32CA19EC74D" degrees="018" direction="east" minutes="09" orientation="longitude" precision="15" seconds="40" value="18.16111">018 ° 09 ' 40 &quot; E</geoCoordinate>
</named-content>
;
<date id="5A79A626C38B0BDC4068AB53E92D2706" value="1977-10-09">
<collectingDate id="9E6B6026F74B6113134F3337678E8300" value="1977-10-09">09 Oct. 1977</collectingDate>
</date>
;
<collectorName id="1AEB0A0F6729090DD2A6D24AD0E4F0B9">Miller, R. M.</collectorName>
leg.;
<named-content id="2644D779E194962B8DB401CA382AAC9E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="4699A53DC430ECA935395C78CDC5B920" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 89516,
<named-content id="7767AD12444929632F5642AE3429C14E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="7B0FDC937A5D0D9DD9889D18760002C8" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="E6F4CE8B6974B3626987FCB07DD7360A" collectingDate="1917-11-20" collectionCode="SAMC" collectorName="Roberts, A." country="South Africa" latitude="-32.09" location="Lamberts Bay" longLatPrecision="20" longitude="18.313055" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="FE8F84216B3B2B504A47C1C841152D62" count="1" type="female">1 ♀</specimenCount>
<location id="025F004A8A3FEFD6A5445884AF647527" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:025F004A8A3FEFD6A5445884AF647527" country="South Africa" latitude="-32.09" longLatPrecision="20" longitude="18.313055" name="Lamberts Bay" stateProvince="Western Cape">Lamberts Bay</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOMAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.313056,-32.090000]}">
<geoCoordinate id="A7505EE6D1A8F51B705D2AC800F4F7C3" degrees="32" direction="south" minutes="05" orientation="latitude" precision="15" seconds="24" value="-32.09">32 ° 05 ' 24 &quot; S</geoCoordinate>
,
<geoCoordinate id="ABA9FD22A11F36F64131C2753FADCC21" degrees="018" direction="east" minutes="18" orientation="longitude" precision="15" seconds="47" value="18.313055">018 ° 18 ' 47 &quot; E</geoCoordinate>
</named-content>
;
<date id="4A5F25492A1EA2A7E4C6CD773CDCF1F0" value="1917-11-20">
<collectingDate id="D2DE53FAF2A749C2DE320B4B5CC1080B" value="1917-11-20">20 Nov. 1917</collectingDate>
</date>
;
<collectorName id="18C9A25A9F24A5B542891396A467824F">Roberts, A.</collectorName>
leg.; SAM-DIP-A 002176,
<named-content id="9E7F5F0121E2CC2B6BD0E4C2D3EFE38A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-african-museum" xlink_title="Iziko South African Museum">
<collectionCode id="756BD9CCC22EC857330AE2B5F85081FD" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" type="Museum">SAMC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="4843C6448BA7668F8F90D4535DD9FBEC" collectingDate="2019-10-03" collectorName="Cabrero &amp; Deschodt &amp; Flowering &amp; Coll. Cabrero" country="South Africa" elevation="2" latitude="-32.58972" location="Rocherpan Nature Reserve" longLatPrecision="20" longitude="18.336668" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="B989E5AB5ADE13A1FB3FAC9CB9C24D76" count="1" type="male">1 ♂</specimenCount>
<location id="54A9EB8D098D87FCFFF623F599BA96A6" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:54A9EB8D098D87FCFFF623F599BA96A6" country="South Africa" latitude="-32.58972" longLatPrecision="20" longitude="18.336668" name="Rocherpan Nature Reserve" stateProvince="Western Cape">Rocherpan Nature Reserve</location>
Trail;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1MAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.336667,-32.589722]}">
<geoCoordinate id="D93C96630399D91D9B2D3A0A3B399CFC" degrees="32" direction="south" minutes="35" orientation="latitude" precision="15" seconds="23" value="-32.58972">32 ° 35 ' 23 &quot; S</geoCoordinate>
,
<geoCoordinate id="786442B05263BD95E6AFE8478A6CC589" degrees="018" direction="east" minutes="20" orientation="longitude" precision="15" seconds="12" value="18.336668">018 ° 20 ' 12 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="66AC6104EA155E03477CA40904EA1226" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">
<quantity id="F21BB2906BD41659C2DAAD62549F12A2" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">2 m</quantity>
a. s. l.
</elevation>
;
<date id="94F560F7B510DD1B612FD19328B2C368" value="2019-10-03">
<collectingDate id="9F1850D9D97778A37046777FD0122C7D" value="2019-10-03">03 Oct. 2019</collectingDate>
</date>
;
<collectorName id="45D6B5980C8D41C3E9F2C4FA9C83E62C">Cabrero</collectorName>
, A.,
<collectorName id="7142E34C54310C959A25D0F710D2A8E0">Deschodt</collectorName>
, C. leg.;
<collectorName id="317C81BD92042EDC3A7084A1FC01D680">Flowering</collectorName>
mesembs on dunes along coast;,
<collectorName id="1D73A27F4407AAAEA66C09217EB7B947">Coll. Cabrero</collectorName>
</materialsCitation>
<materialsCitation id="B727652063F74364F76FCF0CE925C181" collectingDate="1972-09-10" collectionCode="NMSA" collectorName="Irwin, M. E." country="South Africa" elevation="122" latitude="-31.514444" location="Vanrhynsdorp" longLatPrecision="20" longitude="18.719168" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="88EEBE19EBF752B03E830728D52CE1CE" count="1" type="male">1 ♂</specimenCount>
<location id="7B7D7E235EDDE3801A5508DD189ABE24" LSID="urn:lsid:plazi:treatment:FD3D8AE9EE005C6FAD8B7889F23A0E3C:7B7D7E235EDDE3801A5508DD189ABE24" country="South Africa" latitude="-31.514444" longLatPrecision="20" longitude="18.719168" name="Vanrhynsdorp" stateProvince="Western Cape">Vanrhynsdorp</location>
,
<quantity id="89063ACBA43BF914A7B8387DD49E17A4" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" unit="mi" value="7.0">7 mi.</quantity>
N;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECNAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[18.719167,-31.514444]}">
<geoCoordinate id="CAD4996ED0AB1BE5F1912F631E786F37" degrees="31" direction="south" minutes="30" orientation="latitude" precision="15" seconds="52" value="-31.514444">31 ° 30 ' 52 &quot; S</geoCoordinate>
,
<geoCoordinate id="DA3500CB6024CA89D6846ECCFB005FFB" degrees="018" direction="east" minutes="43" orientation="longitude" precision="15" seconds="09" value="18.719168">018 ° 43 ' 09 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="1430631B522A11D86F0A10F94CF51299" metricMagnitude="2" metricUnit="m" metricValue="1.22" unit="m" value="122.0">
<quantity id="61289335F5150574D50A2FB0BFFD6AF7" metricMagnitude="2" metricUnit="m" metricValue="1.22" unit="m" value="122.0">122 m</quantity>
a. s. l.
</elevation>
;
<date id="F36BCD55BBCF542A0C3EC18F58F2B632" value="1972-09-10">
<collectingDate id="A2EF84448F088376EEF0C9BB470FE185" value="1972-09-10">10 Sep. 1972</collectingDate>
</date>
;
<collectorName id="711C8573D375473712BEEA21367C55FA">Irwin, M. E.</collectorName>
leg.; red dunes;
<named-content id="A4ED9A8BE6B5AF275DE22FC8C8993236" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="4FBF15053665E2082BBDD04715A24CA2" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
- DIP- 050202,
<named-content id="DAE384EF65DDDF6020B5080A8B4D2D50" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="F551F85DF9C22F7C2900CE9B8B4B261A" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPNAK" type="distribution">
<paragraph id="DCF003215AA5AE21807823C700E1B02A">
<heading id="19A334641B7C005B71637883B95C04BC" reason="title">Distribution, biodiversity hotspots, phenology, and biology.</heading>
</paragraph>
<paragraph id="4E08B897C7B06C6624670CCE96D99817">
Known from localities in the
<collectingRegion id="0CC0E39CA62587F0D66003C8C0BF7070" country="South Africa" name="Northern Cape">Northern Cape</collectingRegion>
and
<collectingRegion id="4ED1F2ADCFDD1A02CC70F31A7522689D" country="South Africa" name="Western Cape">Western Cape</collectingRegion>
of
<collectingCountry id="2777BF055EA3F55B20EA841947A14C1E" name="South Africa">South Africa</collectingCountry>
(Fig.
<figureCitation id="FB78B22725B14A19FABA6E8D49C2E42C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of E. eremophila, E. syrticola and E. clavicornis specimens studied." figureDoi="10.3897/afrinvertebr.66.129611.figure7" httpUri="https://binary.pensoft.net/fig/1244180">7</figureCitation>
). This species is known from
<specimenCount id="197186225F28A3572B4EC081D8CAB03A" count="30" type="generic">30 specimens</specimenCount>
, collected in 11 collecting events spread between 19172019. The species is known to occur in the Succulent Karoo and
<collectingRegion id="7303DDB0EA82830DE46FF99F63E57820" country="South Africa" name="Western Cape">Cape</collectingRegion>
Floristic Region biodiversity hotspots. Adult flies are active from August - November (Table
<tableCitation id="8F34765ADE32A41FA44FB58EBCC81EFE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity for Enica species through number of specimens collected for each month." httpUri="http://table.plazi.org/id/5E4B94B558D8B5200BC5567C1D820130" tableUuid="5E4B94B558D8B5200BC5567C1D820130">2</tableCitation>
). Adult flies are presumed pollinators, as other species in the genus have been observed visiting flowers (
<ext-link id="D5A80013381720B6CC5BEF7B66AC3330" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/64518819" xlink_type="simple">https://www.inaturalist.org/observations/64518819</ext-link>
). Larvae are unknown but presumed parasitoids as other
<taxonomicName id="67E8FD7C3623203C37C773A499C81A43" authorityName="Westwood" authorityYear="1838" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Bombyliidae</taxonomicName>
, hosts are unknown (
<bibRefCitation id="348B3D7F79A9531D127F75B3E0B13544" DOI="10.1111/j.1095-8312.1997.tb01490.x" author="Yeates and Greathead" firstAuthor="Yeates" issue="2" journalOrPublisher="Linnean Society of London" pagination="149-185" refId="B25" refString="Yeates DK, Greathead D (1997) The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society. Linnean Society of London 60 (2): 149185. https://doi.org/10.1111/j.1095-8312.1997.tb01490.x" title="The evolutionary pattern of host use in the Bombyliidae (Diptera): A diverse family of parasitoid flies. Biological Journal of the Linnean Society." volume="60" year="1997">Yeates and Greathead 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,473 @@
<document id="A947D71B05244EE343D41C51B6FA722C" ID-DOI="10.3897/phytokeys.252.133501" ID-publisher-id="133501" URI-arpha="431329AC-CE65-508C-968C-16F6A14AAE4B" article-type="research-article" checkinTime="1738952585623" checkinUser="pensoft" docAuthor="Xiong, Chi, Fu, Xiao-Ying, Tan, Ke, Huang, Ya, Zhang, Hong-Jing, Wang, Yi-Chen, Yang, Yu-Bing &amp; Yi, Si-Rong" docDate="2025" docId="FEB62E9DC5B9557D8C1ABD9AFEDB2703" docLanguage="en" docName="PhytoKeys 252: 25-40" docOrigin="PhytoKeys 252" docSource="https://phytokeys.pensoft.net/article/133501/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Gastrochilus obovatifolius C. Xiong, X. Y. Fu &amp; S. R. Yi 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="40" masterDocId="431329ACCE65508C968C16F6A14AAE4B" masterDocTitle="Gastrochilus obovatifolius (Orchidaceae, Aeridinae), a new species from the Daba Mountains of Chongqing, China" masterLastPageNumber="40" masterPageNumber="25" pageNumber="25" updateTime="1738952948316" updateUser="ExternalLinkService">
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<mods:title id="7958866AFC351113B055A78F30F49F37">Gastrochilus obovatifolius (Orchidaceae, Aeridinae), a new species from the Daba Mountains of Chongqing, China</mods:title>
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<mods:affiliation id="D232F4C2E35FC67E95D18F27050C963A">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="CC4A754606F990C936FC4D64A845C113">Fu, Xiao-Ying</mods:namePart>
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<mods:affiliation id="2675A158ABBC8127E9EAEBCBBE566F6D">College of Life Sciences, Guangxi Normal University, Guilin 541006, Guangxi, China</mods:affiliation>
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<mods:affiliation id="C94CCE70B55032CF8D601A20C4AA6D2E">Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, 541006, Guangxi, China</mods:affiliation>
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<mods:namePart id="EEB9A5096EA6FE54FB2F4663A5C58B87">Huang, Ya</mods:namePart>
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<mods:affiliation id="AE6D54C5898CA82D61122C9E5F14A1B0">Chongqing Key Laboratory of Development and Utilization of Genuine Medicinal Materials in Three Gorges Reservoir Area, Chongqing Three Gorges Medical College, Wanzhou, Chongqing, 404120, China</mods:affiliation>
</mods:name>
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<mods:namePart id="E66B62E24B5841FA5781C3D67B86D788">Zhang, Hong-Jing</mods:namePart>
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<mods:affiliation id="09927E8D2C23B65CA9EAF07198E3D548">Chongqing Key Laboratory of Development and Utilization of Genuine Medicinal Materials in Three Gorges Reservoir Area, Chongqing Three Gorges Medical College, Wanzhou, Chongqing, 404120, China</mods:affiliation>
</mods:name>
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<mods:affiliation id="D9C5F2BDB94F28706D364BC56093FB7A">Chongqing Key Laboratory of Development and Utilization of Genuine Medicinal Materials in Three Gorges Reservoir Area, Chongqing Three Gorges Medical College, Wanzhou, Chongqing, 404120, China</mods:affiliation>
</mods:name>
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<mods:date id="94500052A6931E0F3DD9D7B348605503">2025</mods:date>
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<treatment id="FEB62E9DC5B9557D8C1ABD9AFEDB2703" ID-DOI="http://doi.org/10.5281/zenodo.14834479" ID-Zenodo-Dep="14834479" ID-arpha="FEB62E9D-C5B9-557D-8C1A-BD9AFEDB2703" ID-ipni="urn:lsid:ipni.org:names:77356371-1" LSID="urn:lsid:plazi:treatment:FEB62E9DC5B9557D8C1ABD9AFEDB2703" LSID-IPNI="urn:lsid:ipni.org:names:77356371-1" httpUri="http://treatment.plazi.org/id/FEB62E9DC5B9557D8C1ABD9AFEDB2703">
<subSubSection id="98FE18B5C2E1577CE2BF422F6DC6DD4C" type="nomenclature">
<paragraph id="CB06B5AC6E0D93A0678E6D35C1CA4E3F">
<taxonomicName id="3D9F16E08148D54C731F415737490588" authority="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius" status="sp. nov.">
<emphasis id="614A1B02DA613ADBD69EB71705B8F51D" italics="true">Gastrochilus obovatifolius</emphasis>
C. Xiong, X. Y. Fu &amp; S. R. Yi
</taxonomicName>
<taxonomicNameLabel id="8D8B193639BC09F4BCE31074EC441361" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="16B7E8A8841ED4A672BBC786960F486B" type="description">
<paragraph id="F16C94EE5E1C615B5E6A84E26505CD79">
<figureCitation id="20BA8CB16E898B9008891EDAA4065E84" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gastrochilus obovatifolius C. Xiong, X. Y. Fu &amp; S. R. Yi, sp. nov. in vivo. A flowering plant B inflorescence C leaves (C 1 C 3: front view, C 4: back view, C 5: close-up of leaf apex) D flower (front view) E flower (back view) F dissected flower (F 1: dorsal sepal; F 2: lateral sepals; F 3: petals; F 4: labellum) G flower (side view) H flower (top view) I pollinarium." figureDoi="10.3897/phytokeys.252.133501.figure3" httpUri="https://binary.pensoft.net/fig/1244759">Figs 3</figureCitation>
,
<figureCitation id="C692EC385B4A78AD16081E642FA6AFC9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Gastrochilus obovatifolius C. Xiong, X. Y. Fu &amp; S. R. Yi, sp. nov., an overview A habit B stem apex C leaves (C 1: front view, C 2: back view, C 3: side view) D flower (side view) E flower (front view) F flower (back view) G dissected flower (G 1: dorsal sepal; G 2: lateral sepals; G 3: petals; G 4: labellum) H pollinarium and anther cap." figureDoi="10.3897/phytokeys.252.133501.figure4" httpUri="https://binary.pensoft.net/fig/1244760">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1VAE" type="diagnosis">
<paragraph id="E46264278901DD50E40B660F2E47B649">
<heading id="3D51DB599888B6E94366BCC58061D299" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="975C60EB879903C09E2850DC4703E7F6">
<taxonomicName id="661F71059C5E26FD012B2C003C408E41" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius">
<emphasis id="FEA2CCDE616A09A47E302CD906DC39BB" italics="true">Gastrochilus obovatifolius</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="D6CD1FA03BFDECABF438877DFE3803E6" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="balangshanensis">
<emphasis id="6B0C5D46A945DF19C34694F9262139DC" italics="true">G. balangshanensis</emphasis>
</taxonomicName>
., but differs by the longer stem (35 vs.
<quantity id="36FCF7C0863E64A42B21F7E266D56D70" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="cm" value="2.5" valueMax="3.5" valueMin="1.5">1.53.5 cm</quantity>
), obovate leaves (vs. nearly elliptic), and smaller sepals (ca. 5.0 × 3.04.0 vs. 5.66.4 ×
<quantity id="5173A6B455D97365AFCEC592687C529B" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.85.2 mm</quantity>
) and hypochile (4.04.5 × 4.04.2 vs. 6.08.0 ×
<quantity id="0F302BC8022990F1D6E797C98D26F2C7" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.5" metricValueMin="5.8" unit="mm" value="6.65" valueMax="7.5" valueMin="5.8">5.87.5 mm</quantity>
).
</paragraph>
<caption id="E9447AA0DBE8C97F89E4A7FAB6257635" ID-DOI="10.3897/phytokeys.252.133501.figure3" ID-arpha="E79308C2-E32B-59B7-9D0E-F8EF29891F80" httpUri="https://binary.pensoft.net/fig/1244759" startId="F3">
<paragraph id="90C50B97BA49A490C37AC36C0A536967">
<label id="6CF2F6D50C743EC6AB71DF307282947E">Figure 3.</label>
</paragraph>
<paragraph id="C78FA12454AEC49E7A812B9AFCFBF8AB">
<taxonomicName id="5C2CE1F9C80C42CDFA2F212FE0FB64F8" authority="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius" status="sp. nov.">
<emphasis id="F3C3091CF6749C9C66BAD31F40415CE8" italics="true">Gastrochilus obovatifolius</emphasis>
C. Xiong, X. Y. Fu &amp; S. R. Yi
</taxonomicName>
,
<taxonomicNameLabel id="3E0910CE365C145CBD2A689B38CDADD9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B73104B6FA9CBCAE0F3F1C10224A7387" italics="true">in vivo</emphasis>
.
<emphasis id="3B3E0C6F71B4AF9378C5914F841787EC" bold="true">A</emphasis>
flowering plant
<emphasis id="F03C8CE1E1B315A99DF3AC5F26865A8A" bold="true">B</emphasis>
inflorescence
<emphasis id="2543E3408AD22C9F0A2366C63F651F42" bold="true">C</emphasis>
leaves (C 1 C 3: front view, C 4: back view, C 5: close-up of leaf apex)
<emphasis id="0097FBFE4E7565AC4FFFC6C6A9B45CB2" bold="true">D</emphasis>
flower (front view)
<emphasis id="33B4FBCED8E120448F9FCC36CA1F154A" bold="true">E</emphasis>
flower (back view)
<emphasis id="FEAF51D9B4165A956E94132ADA914F0D" bold="true">F</emphasis>
dissected flower (F 1: dorsal sepal; F 2: lateral sepals; F 3: petals; F 4: labellum)
<emphasis id="09BB314377457185D30AF62EDC98EE32" bold="true">G</emphasis>
flower (side view)
<emphasis id="415805CC417CD238A196C512CFD68009" bold="true">H</emphasis>
flower (top view)
<emphasis id="89DADCC84A9FB38E8EC004E3FCC66C10" bold="true">I</emphasis>
pollinarium.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ECYAE" type="material">
<paragraph id="72348A1BBD2D7E9ED0061A67A8C4D90A">
<heading id="7E7B63DC88BDF4E2677712ABC43D121A" reason="title">
<typeStatus id="FE279C686034F375E0FE05C8C582FFF5">Type</typeStatus>
.
</heading>
</paragraph>
<paragraph id="2272E86DC72FD7D44583CFD090F73DD0">
<materialsCitation id="EAA627A792B00430F6341FD046D64295" collectingDate="2024-03-30" collectionCode="IBK, CGMC" country="China" county="Chengkou County" elevation="1650" latitude="31.7" location="Lizishuping" longLatPrecision="1214" longitude="109.183334" municipality="Dong'an Town" specimenCount="2" stateProvince="Chongqing" typeStatus="holotype">
<collectingCountry id="673D5AB3FAACE82D6D8C99B572A5855A" name="China">China</collectingCountry>
<collectingRegion id="7079644233A6C227BC308F2DD3C11467" country="China" name="Chongqing">Chongqing</collectingRegion>
:
<collectingCounty id="336D47277A5052EF46908F51F5AC8D82">Chengkou County</collectingCounty>
(城口县),
<collectingMunicipality id="84FB66DE1C20186A4CE5EE30F72C08C3">Dongan Town</collectingMunicipality>
(东安镇),
<location id="D8938CA99215EF53898F1461D9E1AC1D" LSID="urn:lsid:plazi:treatment:FEB62E9DC5B9557D8C1ABD9AFEDB2703:D8938CA99215EF53898F1461D9E1AC1D" country="China" county="Chengkou County" latitude="31.7" longLatPrecision="1214" longitude="109.183334" municipality="Dong'an Town" name="Lizishuping" stateProvince="Chongqing">Lizishuping</location>
(栗子树坪),
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELYAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.183333,31.700000]}">
<geoCoordinate id="16D5555B039BE5DE6A95C5203186ADB9" degrees="31" direction="north" minutes="42" orientation="latitude" precision="925" value="31.7">31 ° 42 N</geoCoordinate>
,
<geoCoordinate id="6249EAE99287FE70A8C013EFB459560E" degrees="109" direction="east" minutes="11" orientation="longitude" precision="925" value="109.183334">109 ° 11 E</geoCoordinate>
</named-content>
, alt. ca.
<quantity id="ABBDE147B14D36122C9C529C9561B17B" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">
<elevation id="EF349E98685A1E6453AA46841A0F52A0" metricMagnitude="3" metricUnit="m" metricValue="1.65" unit="m" value="1650.0">1650 m</elevation>
</quantity>
,
<date id="3FD6B7D9C26C34B2C79A39AB72E991EB" value="2024-03-30">
<collectingDate id="FD44342C116E14CE1031841D2977791E" value="2024-03-30">30 March 2024</collectingDate>
</date>
,
<emphasis id="96E2F45DF1A594B7CC3C03DDED97A520" italics="true">Si-Rong Yi et al. YSR 2703</emphasis>
(
<typeStatus id="5AC5BE993B0FAFCDA3B226AD138270C5" type="holotype">holotype</typeStatus>
<collectionCode id="0EE291452AA8AD9D9831F84050A00665" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13150" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
!,
<typeStatus id="FAE310CE5FC792D19B44BAD1D4855D8C" type="isotype">isotype</typeStatus>
<collectionCode id="6DC5F205F3FC2D5C2E8B4C539B6290AA">CGMC</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="15F4A89F0DCC78CAEA1BB351191CEE1F" ID-DOI="10.3897/phytokeys.252.133501.figure4" ID-arpha="DFE02B4F-8192-5F41-914E-A06ADC1D4474" httpUri="https://binary.pensoft.net/fig/1244760" startId="F4">
<paragraph id="AE1993ED36BFF665852601DE48B47468">
<label id="684E6558DF8A67B53D2766D244E593D2">Figure 4.</label>
</paragraph>
<paragraph id="F8375C62251E380FD7B962EEA7E5C450">
<taxonomicName id="CA027F218686761198127609C869F910" authority="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius" status="sp. nov.">
<emphasis id="EC08609278E8DF038B8AC9D798204F40" italics="true">Gastrochilus obovatifolius</emphasis>
C. Xiong, X. Y. Fu &amp; S. R. Yi
</taxonomicName>
,
<taxonomicNameLabel id="D9704C53158ABB1BA509187C688B8F0C" rank="species">sp. nov.</taxonomicNameLabel>
, an overview
<emphasis id="F9CA93FEA12032DD5768B964A938695E" bold="true">A</emphasis>
habit
<emphasis id="D6814E8FBE5D993FB0F2968002B6CB33" bold="true">B</emphasis>
stem apex
<emphasis id="65CD7184473ECC36AACCD573F774E701" bold="true">C</emphasis>
leaves (C 1: front view, C 2: back view, C 3: side view)
<emphasis id="2D494B029C3B508F44C45A6FE1A9053E" bold="true">D</emphasis>
flower (side view)
<emphasis id="B24B4903EFCEFB2A17D403BD58C4A8C9" bold="true">E</emphasis>
flower (front view)
<emphasis id="11ACAC58248FE5C86EC32AA3C27622AB" bold="true">F</emphasis>
flower (back view)
<emphasis id="99254390ADB862855055DC36FEE36BE0" bold="true">G</emphasis>
dissected flower (G 1: dorsal sepal; G 2: lateral sepals; G 3: petals; G 4: labellum)
<emphasis id="7F10209269FFA3E59139723C3AFFFCB5" bold="true">H</emphasis>
pollinarium and anther cap.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E1ZAE" type="description">
<paragraph id="034AF8189FB0D625792694F69F00B457">
<heading id="C0F1B11D5411D6E344F891B1C9D12DFE" reason="title">Description.</heading>
</paragraph>
<paragraph id="0C9C450DB8E3CF20B676E309BA9E27DD">
<emphasis id="BA7D5529CC05C05E23B2BD06E957A020" bold="true" italics="true">Epiphytic herbs</emphasis>
, monopodial, pendent, with a short stem, 3.05.0 cm long, leafy.
<emphasis id="7496A275E65F7186966BB77367B4CBA5" bold="true" italics="true">Roots</emphasis>
vermiform, slender, 4.06.0 cm long and ca.
<quantity id="D272F2037F421E64F39696C24EF4725A" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
in diameter.
<emphasis id="8B86DB1F7C52E21A62FE5429E0A0210B" bold="true" italics="true">Stems</emphasis>
short, branched, green, glabrous, up to
<quantity id="DAAA9E7D2E76D9F9DCA7432ECB888D8E" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="cm" value="5.0">5 cm</quantity>
long and ca. 2.0 mm in diameter, covered with sheathing leaf bases; sheaths with purplish-red spots; internodes
<quantity id="6AEC94E46D201C6F8375C04654C98132" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.21.8 mm</quantity>
long.
<emphasis id="F0A8CD1C6FAE89591EB95FA69D4AC991" bold="true" italics="true">Leaves</emphasis>
alternate, distichous, obovate, fleshy, 1.41.6 ×
<quantity id="B68DD13BAA4808F25EAE3A5ED7B1FD69" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="cm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 cm</quantity>
, margin entire, apex acute, obscurely serrate; young leaves yellow-green with a few purplish-red spots, mature leaves green with hardly any purplish-red spots.
<emphasis id="F109B6AED593170A5B23EB8C2F7740D5" bold="true" italics="true">Inflorescences</emphasis>
12 racemes, usually arising from the axil of an upper leaf, 2.0
<quantity id="8B3DFCF2FE4DBD9AA576E67EDA0D9F19" metricMagnitude="-2" metricUnit="m" metricValue="2.5" unit="cm" value="2.5">2.5 cm</quantity>
long, 12 - flowered; peduncle
<quantity id="4BA7588A9FD19BAB48D4304A8582AE1D" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 cm</quantity>
long; bracts 1.02.0 mm long.
<emphasis id="0810AB9A3E905A7BED0AC3C87150BE05" bold="true" italics="true">Flowers</emphasis>
1.21.4 × 1.0
<quantity id="7B65C4B17BEA8C531727C40116797EEF" metricMagnitude="-2" metricUnit="m" metricValue="1.2" unit="cm" value="1.2">1.2 cm</quantity>
, yellow-green, with dark purple stripes on petals and sepals, raised abaxially along the midrib; pedicel and ovary yellowish-green with purple-red spots,
<quantity id="96B9B780B37F010C225BBB2EC936FE7B" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="cm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 cm</quantity>
long.
<emphasis id="835674617C8348E235A8CA3FC8EB8570" bold="true" italics="true">Dorsal sepal</emphasis>
elliptic, concave, ca. 5.0 × 4.0 mm, apex obtuse;
<emphasis id="91F1199BC91507444C71275603CA182E" bold="true" italics="true">lateral sepals</emphasis>
similar to dorsal sepal, slightly oblique, ca. 5.0 × 3.0 mm, apex obtuse; petals oblong, concave, ca. 6.0 × 3.0 mm, apex obtuse.
<emphasis id="A850D11547D3B6A43AD1AF4707FE0179" bold="true" italics="true">Labellum</emphasis>
epichile reniform, yellow-green with purplish-red spots, 1.01.2 ×
<quantity id="3E7BF10D439427288E38C49DD0D7F9A0" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="cm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 cm</quantity>
, revolute, margin erose, smooth and glabrous above, median patch, thickened, with 2 low ridges, dark purple; hypochile sub-hemispherical, yellow-green, mouth with lateral purplish-red markings, sac with purplish-red spots on the underside, obtuse at the apex, 4.0
<quantity id="0DBD334D8485F43BC2CB57B4BA31A1A5" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
tall, 4.0
<quantity id="07058C2D4DCE315676A953E9E3AA76B2" metricMagnitude="-3" metricUnit="m" metricValue="4.2" unit="mm" value="4.2">4.2 mm</quantity>
in diameter, dorsally compressed, slightly bent outward.
<emphasis id="77A1BD02FC12DD666B99B6753718C2D7" bold="true" italics="true">Column</emphasis>
stout, ca. 2.0 ×
<quantity id="38DE0213F30F03B157C84D49CE993774" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
; rostellum bilobed; anther cap galeate with curved emarginate beak, ca. 1.0 ×
<quantity id="4031D75B36FB867243CF55D4DF439405" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
; pollinia 2, ca. 0.8 ×
<quantity id="DDEB7706900216A026F13BD864879E22" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
, yellow, nearly spherical, entire, with a depression in the center; stipe elongate, ca.
<quantity id="DB2391F526E97FAB616A83CE83197621" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
long, viscidium yellow, elliptic, ca. 0.8 ×
<quantity id="D5C4CB0DA4F38FC4A786E9F1FA235273" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
.
<emphasis id="7B1C7B23C08450A3B389350EDF1FE067" bold="true" italics="true">Capsule</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ED2AE" type="phenology">
<paragraph id="253FAB1CD0A57E1C2190C3E2125DABE4">
<heading id="0C0E7D21D0C779F140F942D54A6A3462" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="09BA586B8E7C8413B5B833A996CBA71E">Flowering from March to April.</paragraph>
</subSubSection>
<subSubSection id="SECID0EI2AE" type="etymology">
<paragraph id="30AFC191878C78C723E2FC4657488D74">
<heading id="5832644404647CE2EAFD7A5C269E2AFE" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="00A62F2693A83F72351BA59F770E3DFB">
The specific epithet
<taxonomicName id="D8BF7F49D5C36AF8CA82BA9928836B4E" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius">
<emphasis id="D132CEE81C591D08F87A58CC5FEBD98C" italics="true">obovatifolius</emphasis>
</taxonomicName>
refers to the highly distinctive obovate leaves. The suggested Chinese common name is “ dào luǎn yè pén jù lán (倒卵叶盆距兰) ”.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY2AE" type="distribution">
<paragraph id="584C66308D4303CD979B26ABB596740B">
<heading id="A89B2B7D0F5B329F1D687E43AC7B07CB" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="68A98B2AEB45129DC926ABBBA6031A32">
The new species has only been recorded in Chengkou County, northeast
<collectingRegion id="6FD9A375226A9D89A5E25C38A0ED8490" country="China" name="Chongqing">Chongqing</collectingRegion>
Municipality, bordering
<collectingRegion id="48FA55ADC285FD9E60A7B50ED35B6A30" country="China" name="Shaanxi">Shaanxi Province</collectingRegion>
,
<collectingCountry id="741CB0A66C5AC5D3F0BDD2362459224D" name="China">China</collectingCountry>
(Fig.
<figureCitation id="CB1284CFE2E9A65A8B3F651AED2CA605" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution map of Gastrochilus obovatifolius and three related species of G. sect. Microphylli in southwestern China and adjoining areas." figureDoi="10.3897/phytokeys.252.133501.figure6" httpUri="https://binary.pensoft.net/fig/1244762">6</figureCitation>
). It grows as a trunk epiphyte on
<taxonomicName id="E56DEB73D3793812B4789A1EEB1D9EE7" authority="Seemen" authorityName="Seemen" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="engleriana">
<emphasis id="FF6A8B3E6D8BDC00528A4C2399779F25" italics="true">Quercus engleriana</emphasis>
Seemen
</taxonomicName>
, in evergreen broad-leaved forest at an elevation of
<quantity id="9C2595B6490E07E20258EC26E1C009CF" metricMagnitude="3" metricUnit="m" metricValue="1.635" metricValueMax="1.65" metricValueMin="1.6199999999999999" unit="m" value="1635.0" valueMax="1650.0" valueMin="1620.0">16201650 m</quantity>
a. s. l. (Fig.
<figureCitation id="5C10603C4424240DB9DFAB2474558E1C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Gastrochilus obovatifolius, habitat and plants in situ. A broad-leaved temperate forest in the Daba Mountains of Chongqing B a woodland trail lined with Fargesia spathacea, Quercus engleriana, Pieris japonica, Rhododendron adenopodum, and Zanthoxylum dimorphophyllum C G. obovatifolius growing as a trunk epiphyte on Q. engleriana." figureDoi="10.3897/phytokeys.252.133501.figure5" httpUri="https://binary.pensoft.net/fig/1244761">5</figureCitation>
). Besides
<taxonomicName id="724CED60FB4BEAB82980C5C294555534" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="engleriana">
<emphasis id="873691DE81E43E90E6934C350D9B75E9" italics="true">Q. engleriana</emphasis>
</taxonomicName>
, the most frequent co-occurring angiosperm species include
<taxonomicName id="C6F566A14E94535E4454070178158CFD" authority="Franch." authorityName="Franch." class="Liliopsida" family="Poaceae" genus="Fargesia" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="spathacea">
<emphasis id="FFCD6D8E9B11FFD94B5BDDA8D0E1B069" italics="true">Fargesia spathacea</emphasis>
Franch.
</taxonomicName>
,
<taxonomicName id="01AE61F30F710B3DC50977C0AE739887" authority="(Oliv.) Steward" authorityName="Steward" baseAuthorityName="Oliv." class="Magnoliopsida" family="Ranunculaceae" genus="Hepatica" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="species" species="henryi">
<emphasis id="4D93ADE71E5A33981D353532D0C3DFF8" italics="true">Hepatica henryi</emphasis>
(Oliv.) Steward
</taxonomicName>
,
<taxonomicName id="3071BB23343DE4027E0366DDFD7611E4" authority="(Thunb.) D. Don ex G. Don" authorityName="G. Don" baseAuthorityName="Thunb." class="Magnoliopsida" family="Ericaceae" genus="Pieris" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="95A2432FD679B96225AC91E35FB5D630" italics="true">Pieris japonica</emphasis>
(Thunb.) D. Don ex G. Don
</taxonomicName>
,
<taxonomicName id="8CC15A4C502F78F269E11A706F520C05" authority="Franch." authorityName="Franch." class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="adenopodum">
<emphasis id="82D44D42873B3636A0F4D66057DD99F8" italics="true">Rhododendron adenopodum</emphasis>
Franch.
</taxonomicName>
, and
<taxonomicName id="78C20D6AD991D0F63F66245508B507D9" authority="Hemsl." authorityName="Hemsl." class="Magnoliopsida" family="Rutaceae" genus="Zanthoxylum" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="dimorphophyllum">
<emphasis id="E2B5908D4184770FEA2403222F09DD82" italics="true">Zanthoxylum dimorphophyllum</emphasis>
Hemsl.
</taxonomicName>
</paragraph>
<caption id="127BCA7B42F97429B6B716033184FE99" ID-DOI="10.3897/phytokeys.252.133501.figure5" ID-arpha="2AA4BC01-BBB6-5AD1-BE0A-AD2D493D2CE8" httpUri="https://binary.pensoft.net/fig/1244761" startId="F5">
<paragraph id="B919330D219F5985BAFE9EB1217B696F">
<label id="22A0BE36B0E83B8FD1FC930F954FF349">Figure 5.</label>
</paragraph>
<paragraph id="C4D3EC336C38D1B97CF262470EA63F23">
<taxonomicName id="80F8966CFDD2AAE2C02A6485221F2963" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius">
<emphasis id="E8B1D32AEABE558A53D29EC5EA31292F" italics="true">Gastrochilus obovatifolius</emphasis>
</taxonomicName>
, habitat and plants in situ.
<emphasis id="C3BA8819BB231F5B8F001D73AE69E303" bold="true">A</emphasis>
broad-leaved temperate forest in the Daba Mountains of Chongqing
<emphasis id="C4DEAB50A903421149E083CDD2C206DB" bold="true">B</emphasis>
a woodland trail lined with
<taxonomicName id="1A2D84FD093BA782D65D4F7635CD807E" class="Liliopsida" family="Poaceae" genus="Fargesia" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="spathacea">
<emphasis id="8880EC0FAD10B986682C73EC32AC4D3C" italics="true">Fargesia spathacea</emphasis>
</taxonomicName>
,
<taxonomicName id="DCCBCCD26153CB10AD0C84A64EB60CC3" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="engleriana">
<emphasis id="B01D59F7E6608EA22334238BA2BC4E23" italics="true">Quercus engleriana</emphasis>
</taxonomicName>
,
<taxonomicName id="C0B9E5D6ECDB7A140AD332800E6C759B" class="Magnoliopsida" family="Ericaceae" genus="Pieris" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="060B71B9C03C6481FE2A00C256E1DBEF" italics="true">Pieris japonica</emphasis>
</taxonomicName>
,
<taxonomicName id="62DB811BFFE96CBAB2A84A00ED6FB8B8" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="adenopodum">
<emphasis id="186BC7466C09A7D19B2E7CDB8CC1FE32" italics="true">Rhododendron adenopodum</emphasis>
</taxonomicName>
, and
<taxonomicName id="7F8DAF84036E610BEB42E900F59CFE84" class="Magnoliopsida" family="Rutaceae" genus="Zanthoxylum" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="dimorphophyllum">
<emphasis id="D13EA06D4AEB8531C4ECBA9ADB1A6626" italics="true">Zanthoxylum dimorphophyllum</emphasis>
</taxonomicName>
<emphasis id="919FAE4951EF5140727AF6EF0B499952" bold="true">C</emphasis>
<taxonomicName id="38BB8504E597BFD9F70088BF598F54B3" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius">
<emphasis id="B4E30EC9F710577FF7479F96A5514734" italics="true">G. obovatifolius</emphasis>
</taxonomicName>
growing as a trunk epiphyte on
<taxonomicName id="768C29AF673B9A38757C03CE16CBEC29" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="engleriana">
<emphasis id="79E839288DFE0116CD3363D1EA95FD05" italics="true">Q. engleriana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="78DB9F89DE6CEB43F12A0542074AE3E6" ID-DOI="10.3897/phytokeys.252.133501.figure6" ID-arpha="AAA05ABE-27CA-5F83-A213-53FBC55B3D4C" httpUri="https://binary.pensoft.net/fig/1244762" startId="F6">
<paragraph id="EA924267E8828726241F3962DF2697EB">
<label id="3BAA9D7B9A61BE1D455E53133BFDF932">Figure 6.</label>
</paragraph>
<paragraph id="D427A3474AAAC5C362DAEE2735C8B8E3">
Distribution map of
<taxonomicName id="7ED9EFA7ECE631C824DC7F3ECFEF963C" authorityName="C. Xiong, X. Y. Fu &amp; S. R. Yi" authorityYear="2025" class="Liliopsida" family="Orchidaceae" genus="Gastrochilus" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="obovatifolius">
<emphasis id="75D64DB3ABC9CC231AC2FD9E2E5B3DB8" italics="true">Gastrochilus obovatifolius</emphasis>
</taxonomicName>
and three related species of
<emphasis id="0ABAA8A4C825A2B2FAA0D81C606EB078" italics="true">G. sect. Microphylli</emphasis>
in southwestern China and adjoining areas.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEDAG" type="material">
<paragraph id="E5DFCA0AAE8910DC7A6AE023B1845863">
<heading id="155228EF7E3FE0065E43FF9918C7688F" reason="title">
Additional specimens examined (
<typeStatus id="EE1616EA70B60DC3AA7B7C6BD46DB803" type="paratype">paratypes</typeStatus>
).
</heading>
</paragraph>
<paragraph id="A68A3EA87E3DA0F479786155C1CA1C14">
<materialsCitation id="AD0E7F88D0DC102DFB2A21CF4F76A4D0" collectingDate="2024-04-08" collectionCode="IBK, CGMC" country="China" county="Chengkou County" elevation="1620" latitude="31.7" location="Dongjiachang" longLatPrecision="1214" longitude="109.2" municipality="Dong'an Town" specimenCount="1" stateProvince="Chongqing">
<collectingCountry id="9F71D25A345373B33552C24070D2F9FA" name="China">China</collectingCountry>
<collectingRegion id="EA42803E3983ED1D267FF337942ADF52" country="China" name="Chongqing">Chongqing</collectingRegion>
:
<collectingCounty id="6BEDF95704CD3EC16F15FFAD60685DD6">Chengkou County</collectingCounty>
(城口县),
<collectingMunicipality id="9A84EE25DE0A0B4F4718747792CE89A7">Dongan Town</collectingMunicipality>
(东安镇),
<location id="BE446E2BD3488BEB808EF9DA9AFC510C" LSID="urn:lsid:plazi:treatment:FEB62E9DC5B9557D8C1ABD9AFEDB2703:BE446E2BD3488BEB808EF9DA9AFC510C" country="China" county="Chengkou County" latitude="31.7" longLatPrecision="1214" longitude="109.2" municipality="Dong'an Town" name="Dongjiachang" stateProvince="Chongqing">Dongjiachang</location>
(董家厂),
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENDAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.200000,31.700000]}">
<geoCoordinate id="4A14602C1E8F6F4FB0B172B023128F70" degrees="31" direction="north" minutes="42" orientation="latitude" precision="925" value="31.7">31 ° 42 N</geoCoordinate>
,
<geoCoordinate id="6D56F61E44674DF8291264B2A5A96C68" degrees="109" direction="east" minutes="12" orientation="longitude" precision="925" value="109.2">109 ° 12 E</geoCoordinate>
</named-content>
, alt. ca.
<quantity id="0E7F7093043657BF590F40D9FEEC4682" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">
<elevation id="A885E22D07CE8FC903E22BD2998F30FB" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">1620 m</elevation>
</quantity>
,
<date id="A778540DC6C5EA0FF84C6EC7691398D9" value="2024-04-08">
<collectingDate id="B59C433B0E52A12EA84791C787758CC8" value="2024-04-08">8 April 2024</collectingDate>
</date>
,
<emphasis id="5D5EDE261C4F5918A6A0127F791B52A8" italics="true">Si-Rong Yi et al. YSR 2733</emphasis>
(
<collectionCode id="01FE10DF581AAE6830695E17A0E03559" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13150" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" type="Herbarium">IBK</collectionCode>
!,
<collectionCode id="72591B23A7E64A14F2CA70B546CFEE80">CGMC</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>