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<taxonomicName id="FCC13BF00771FFF8FF47FA8B3C37FADE" ID-CoL="74X3" authority="Montemayor &amp; Carpintero, 2007" authorityName="Montemayor &amp; Carpintero" authorityYear="2007" box="[163,779,1304,1329]" class="Insecta" family="Tingidae" genus="Pterovianaida" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1200" phylum="Arthropoda" rank="genus">
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<emphasis id="09B59C610771FFF8FF47FA8B3E5CFADF" bold="true" box="[163,352,1304,1328]" italics="true" pageId="28" pageNumber="1200">Pterovianaida</emphasis>
<bibRefCitation id="5F503D820771FFF8FE82FA8A3C37FADE" author="Montemayor SI &amp; Carpintero DL" box="[358,779,1305,1329]" pageId="28" pageNumber="1200" pagination="133 - 136" refId="ref25263" refString="Montemayor SI, Carpintero DL. 2007. A new macropterous genus with a new species of Vianaididae (Heteroptera, Tingoidea, Vianaididae) from Peru. Studies on Neotropical Fauna and Environment 42: 133-136." type="journal article" year="2007">Montemayor &amp; Carpintero, 2007</bibRefCitation>
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<emphasis id="09B59C610771FFF8FF47FAC43E26FA83" box="[163,282,1367,1388]" italics="true" pageId="28" pageNumber="1200">Diagnosis:</emphasis>
This genus is characterized by the abundant long hairs on the head, which is remarkably declined, by the presence of paranota constricted in the middle and a conspicuously raised distal part of the pronotum and by the scent gland that is laterally projected with the edges of the tips of both anterior and posterior peritreme branches enlarged.
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<emphasis id="09B59C610771FFF8FF47F9D03E90F9B7" box="[163,428,1603,1624]" italics="true" pageId="28" pageNumber="1200">Redescription: Head:</emphasis>
Triangular in dorsal view, pubescent, with large, long hairs on vertex, once referred as macrochaetae; pedicel subequal to basiflagellomere and to distiflagellomere, each one of these more than twice the length of scape; eyes fully developed, compound; rostrum reaching at least the second abdominal segment.
<emphasis id="09B59C610771FFF8FE7EF9683EC8F8FF" box="[410,500,1787,1808]" italics="true" pageId="28" pageNumber="1200">Thorax:</emphasis>
Posterior lobe strongly raised, pubescent; entirely punctate; collar distinct, punctate; anterior border straight, posterior convex; paranota explanate, smooth, constricted in the middle, between the anterior and posterior lobes.
<emphasis id="09B59C610771FFF8FAC7FBC73A9EFB86" box="[1315,1442,1108,1129]" italics="true" pageId="28" pageNumber="1200">Hemelytra:</emphasis>
Macropterous, clavus and vein-less membrane well-defined; scale-like projections on hemelytra border; punctate on coriaceous parts; membrane with one inner row of punctations; hypocosta broad and finely punctate.
<emphasis id="09B59C610771FFF8FCDFFB7D3CEAFAED" box="[827,982,1261,1283]" italics="true" pageId="28" pageNumber="1200">Scent gland:</emphasis>
Anterior branch gradually projected laterally, tip much more projected than base; tip tilted horizontally, large, conspicuously inclined frontwards; posterior branch usually curved, with enlarged tip.
<emphasis id="09B59C610771FFF8FA8DFAD93A9FFAB0" box="[1385,1443,1354,1375]" italics="true" pageId="28" pageNumber="1200">Legs:</emphasis>
Femora same width of the other segments; tarsi two segment, second segment remarkably longer and many times the length of the first.
<emphasis id="09B59C610771FFF8FB76FA363A3AFA55" box="[1170,1286,1445,1466]" italics="true" pageId="28" pageNumber="1200">Abdomen:</emphasis>
Rectangular, covered with hairs; spiracles located ventrally near lateral margins of abdominal sternites, these straight.
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species:
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.
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<emphasis id="09B59C610771FFF8FCDFF9FA3CF0F991" box="[827,972,1641,1662]" italics="true" pageId="28" pageNumber="1200">Distribution:</emphasis>
Known from
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and
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(
<figureCitation id="A3FA5CF60771FFF8FACFF9FA3AB6F991" box="[1323,1418,1641,1663]" captionStart="Figure 18" captionStartId="15.[146,228,1527,1549]" captionTargetBox="[288,1282,195,1487]" captionTargetId="figure-137@15.[288,1282,195,1487]" captionTargetPageId="15" captionText="Figure 18. Maps of distribution of the extant vianaidines: A, Anommatocoris species; B, Henryvianaida species; C, Pterovianaida species; D, Thaumamannia species." figureDoi="http://doi.org/10.5281/zenodo.14856891" httpUri="https://zenodo.org/record/14856891/files/figure.png" pageId="28" pageNumber="1200">Fig. 18C</figureCitation>
).
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<paragraph id="3B7E40730771FFF8FCDFF90D3A9EF884" blockId="28.[827,1442,1693,1899]" pageId="28" pageNumber="1200">
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Discussion:
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was described by
<bibRefCitation id="5F503D820771FFF8FCDFF92F3BF7F93E" author="Montemayor SI &amp; Carpintero DL" box="[827,1227,1724,1746]" pageId="28" pageNumber="1200" pagination="133 - 136" refId="ref25263" refString="Montemayor SI, Carpintero DL. 2007. A new macropterous genus with a new species of Vianaididae (Heteroptera, Tingoidea, Vianaididae) from Peru. Studies on Neotropical Fauna and Environment 42: 133-136." type="journal article" year="2007">Montemayor &amp; Carpintero (2007)</bibRefCitation>
based on a single slide-mounted specimen. Additionally, it is clear the authors were not aware of macropterous species described by
<bibRefCitation id="5F503D820771FFF8FC30F88B3B93F8C1" author="Schuh RT &amp; Cassis G &amp; Guilbert E" box="[980,1199,1816,1838]" pageId="28" pageNumber="1200" pagination="38 - 53" refId="ref25415" refString="Schuh RT, Cassis G, Guilbert E. 2006. Description of the first recent macropterous species of Vianaidinae (Heteroptera: Tingidae) with comments on the phylogenetic relationships of the family within the Cimicomorpha. Journal of the New York Entomological Society 114: 38-53." type="journal article" year="2006">
Schuh
<emphasis id="09B59C610771FFF8FBC0F88A3B62F8C2" box="[1060,1118,1816,1838]" italics="true" pageId="28" pageNumber="1200">et al.</emphasis>
(2006)
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when proposing this genus. However, the genus gained its second species almost ten years after its original description, based on
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<paragraph id="3B7E40730772FFFBFF37FCA13D8BFCA5" blockId="31.[211,695,818,842]" box="[211,695,818,842]" pageId="31" pageNumber="1203">
<heading id="6036F71F0772FFFBFF37FCA13D8BFCA5" box="[211,695,818,842]" centered="true" fontSize="9" level="2" pageId="31" pageNumber="1203" reason="2">
<taxonomicName id="FCC13BF00772FFFBFF37FCA13D8BFCA5" ID-CoL="7VBV" authority="Drake &amp; Davis, 1960" authorityName="Drake &amp; Davis" authorityYear="1960" box="[211,695,818,842]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="genus">
<emphasis id="09B59C610772FFFBFF37FCA13D8BFCA5" bold="true" box="[211,695,818,842]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFF37FCA13E94FCA5" bold="true" box="[211,424,818,842]" italics="true" pageId="31" pageNumber="1203">Thaumamannia</emphasis>
<bibRefCitation id="5F503D820772FFFBFE4BFCA13D8BFCA5" author="Drake CJ &amp; Davis NT" box="[431,695,818,842]" pageId="31" pageNumber="1203" pagination="1 - 100" refId="ref24289" refString="Drake CJ, Davis NT. 1960. The morphology, phylogeny, and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Hemiptera: Heteroptera). Entomologica Americana 39: 1-100." type="journal article" year="1960">Drake &amp; Davis, 1960</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73DB13F80772FFFBFF75FCE33B2EFA85" pageId="31" pageNumber="1203" type="diagnosis">
<paragraph id="3B7E40730772FFFBFF75FCE33D60FBF0" blockId="31.[145,761,880,1055]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFF75FCE33E32FC6A" box="[145,270,880,901]" italics="true" pageId="31" pageNumber="1203">Diagnosis:</emphasis>
Characterized by the inclined head, the presence of explanate paranota and costal area, the relatively small scutellum, conspicuously concave hemelytra, and the laterally projected scent gland, strongly Y-shaped with enlarged tips in both anterior and posterior branches of the peritreme.
</paragraph>
<paragraph id="3B7E40730772FFFBFF75FBAE3B7FFDA4" blockId="31.[145,762,1085,1383]" lastBlockId="31.[809,1426,197,587]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFF75FBAE3E46FBBD" box="[145,378,1085,1106]" italics="true" pageId="31" pageNumber="1203">Redescription: Head:</emphasis>
Declined, usually with most of its length hidden in dorsal view, heavily pubescent; clypeus usually in different colour than head; pedicel subequal to basiflagellomere, both smaller than distiflagellomere and usually twice as big as scape; eyes reduced with none or few ommatidia, triangular in shape; rostrum reaching at least the third abdominal segment.
<emphasis id="09B59C610772FFFBFD46FB663DC5FAE5" box="[674,761,1269,1290]" italics="true" pageId="31" pageNumber="1203">Thorax:</emphasis>
Posterior lobe slightly raised; coarsely punctate, mostly on collar and posterior lobe, collar indistinct; anterior border usually concave; posterior usually straight; paranota explanate, smooth, usually projected forwards.
<emphasis id="09B59C610772FFFBFCCDFF773C98FF16" box="[809,932,228,249]" italics="true" pageId="31" pageNumber="1203">Hemelytra:</emphasis>
Entirely coriaceous and coleopteroid, ovate, pubescent; remarkably convex, no clavus nor membrane distinct; carina-like vein extending variably from the anterior border, deeply punctate; costal area explanate, with one inner row of punctations only; hypocosta broad and coarsely punctate.
<emphasis id="09B59C610772FFFBFBDEFEEE3BF0FE7D" box="[1082,1228,381,402]" italics="true" pageId="31" pageNumber="1203">Scent gland:</emphasis>
Anterior branch gradually projected laterally, tip much more projected than base; varying in height, conspicuously inclined frontwards; tip tilted horizontally, large; posterior branch usually curved, with enlarged tip.
<emphasis id="09B59C610772FFFBFB18FE6B3A09FDE2" box="[1276,1333,504,525]" italics="true" pageId="31" pageNumber="1203">Legs:</emphasis>
Femora only slightly swollen; tarsi two segment, second segment at least five-times longer.
</paragraph>
<paragraph id="3B7E40730772FFFBFCCDFDF93C88FD71" blockId="31.[809,1423,617,670]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFCCDFDF93CEFFD90" box="[809,979,618,639]" italics="true" pageId="31" pageNumber="1203">
<typeStatus id="E47AFED10772FFFBFCCDFDF93C58FD90" box="[809,868,618,639]" pageId="31" pageNumber="1203">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="FCC13BF00772FFFBFC05FDFA3C8CFD71" authority="Drake &amp; Davis, 1960" authorityName="Drake &amp; Davis" authorityYear="1960" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="09B59C610772FFFBFC05FDFA3A2EFD90" box="[993,1298,617,639]" italics="true" pageId="31" pageNumber="1203">Thaumamannia manni</emphasis>
<bibRefCitation id="5F503D820772FFFBFAFBFDFA3C8CFD71" author="Drake CJ &amp; Davis NT" pageId="31" pageNumber="1203" pagination="1 - 100" refId="ref24289" refString="Drake CJ, Davis NT. 1960. The morphology, phylogeny, and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Hemiptera: Heteroptera). Entomologica Americana 39: 1-100." type="journal article" year="1960">Drake &amp; Davis, 1960</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="3B7E40730772FFFBFCCDFD2F3AB0FD3E" blockId="31.[809,1420,700,722]" box="[809,1420,700,722]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFCCDFD2F3C87FD3E" box="[809,955,700,721]" italics="true" pageId="31" pageNumber="1203">Distribution:</emphasis>
<collectingCountry id="43D600E30772FFFBFC26FD2F3B36FD3D" box="[962,1034,700,722]" name="Brazil" pageId="31" pageNumber="1203">Brazil</collectingCountry>
,
<collectingCountry id="43D600E30772FFFBFBF0FD2F3B58FD3D" box="[1044,1124,700,722]" name="Bolivia" pageId="31" pageNumber="1203">Bolivia</collectingCountry>
and
<collectingCountry id="43D600E30772FFFBFB7AFD2F3A32FD3D" box="[1182,1294,700,722]" name="Suriname" pageId="31" pageNumber="1203">Suriname</collectingCountry>
(
<figureCitation id="A3FA5CF60772FFFBFAF9FD2F3A41FD3E" box="[1309,1405,700,722]" captionStart="Figure 18" captionStartId="15.[146,228,1527,1549]" captionTargetBox="[288,1282,195,1487]" captionTargetId="figure-137@15.[288,1282,195,1487]" captionTargetPageId="15" captionText="Figure 18. Maps of distribution of the extant vianaidines: A, Anommatocoris species; B, Henryvianaida species; C, Pterovianaida species; D, Thaumamannia species." figureDoi="http://doi.org/10.5281/zenodo.14856891" httpUri="https://zenodo.org/record/14856891/files/figure.png" pageId="31" pageNumber="1203">Fig. 18D</figureCitation>
).
</paragraph>
<paragraph id="3B7E40730772FFFBFCCDFD633B2EFA85" blockId="31.[809,1425,752,1387]" pageId="31" pageNumber="1203">
<emphasis id="09B59C610772FFFBFCCDFD633C88FCEA" box="[809,948,752,773]" italics="true" pageId="31" pageNumber="1203">Discussion:</emphasis>
The genus was described by
<bibRefCitation id="5F503D820772FFFBFAC3FD633C82FCCB" author="Drake CJ &amp; Davis NT" pageId="31" pageNumber="1203" pagination="1 - 100" refId="ref24289" refString="Drake CJ, Davis NT. 1960. The morphology, phylogeny, and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Hemiptera: Heteroptera). Entomologica Americana 39: 1-100." type="journal article" year="1960">Drake &amp; Davis (1960)</bibRefCitation>
and the second species,
<taxonomicName id="FCC13BF00772FFFBFB00FC9C3B1AFCAD" authority="Van Doesburg (1977)" authorityName="Van Doesburg" authorityYear="1977" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="vanderdrifti">
<emphasis id="09B59C610772FFFBFB00FC9C3AACFCCC" box="[1252,1424,782,804]" italics="true" pageId="31" pageNumber="1203">T. vanderdrifti</emphasis>
<bibRefCitation id="5F503D820772FFFBFCCDFCBE3B1AFCAD" author="Van Doesburg PH" box="[809,1062,813,835]" pageId="31" pageNumber="1203" pagination="187 - 189" refId="ref25604" refString="Van Doesburg PH. 1977. A new species of Thaumamannia from Surinam (Heteroptera, Tingidae, Vianaidinae). Zoologische Mededelingen 52: 187-189." type="journal article" year="1977">Van Doesburg (1977)</bibRefCitation>
</taxonomicName>
was described 17 years later. From these two species, only
<taxonomicName id="FCC13BF00772FFFBFB60FCDF3BD2FC8E" authorityName="Drake &amp; Davis" authorityYear="1960" box="[1156,1262,844,865]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="09B59C610772FFFBFB60FCDF3BD2FC8E" box="[1156,1262,844,865]" italics="true" pageId="31" pageNumber="1203">T. manni</emphasis>
</taxonomicName>
was collected associated with ants (
<bibRefCitation id="5F503D820772FFFBFBF3FCF93BC6FC6F" author="Drake CJ &amp; Davis NT" box="[1047,1274,874,896]" pageId="31" pageNumber="1203" pagination="1 - 100" refId="ref24289" refString="Drake CJ, Davis NT. 1960. The morphology, phylogeny, and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Hemiptera: Heteroptera). Entomologica Americana 39: 1-100." type="journal article" year="1960">Drake &amp; Davis, 1960</bibRefCitation>
). The broadly ovate body, which was considered a strong diagnostic feature of the genus, is now challenged with the description of a new species herein described. However, the conspicuously Y-shaped scent gland remains as an important diagnostic character for the genus, currently composed by a total of four species, including the two species described below.
<taxonomicName id="FCC13BF00772FFFBFBA7FBD23AADFBB9" authorityName="Van Doesburg" authorityYear="1977" box="[1091,1425,1089,1110]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="vanderdrifti">
<emphasis id="09B59C610772FFFBFBA7FBD23AADFBB9" box="[1091,1425,1089,1110]" italics="true" pageId="31" pageNumber="1203">Thaumamannia vanderdrifti</emphasis>
</taxonomicName>
was the first
<taxonomicName id="FCC13BF00772FFFBFC5EFBF33B7EFB99" box="[954,1090,1120,1142]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1210" phylum="Arthropoda" rank="subFamily" subFamily="Vianaidinae">Vianaidinae</taxonomicName>
reported from
<collectingCountry id="43D600E30772FFFBFB0DFBF33A12FB99" box="[1257,1326,1120,1142]" name="Brazil" pageId="31" pageNumber="1203">Brazil</collectingCountry>
(
<bibRefCitation id="5F503D820772FFFBFAD8FBF33C9AFB7C" author="Guidoti M &amp; Barcellos A &amp; Ferreira RL" pageId="31" pageNumber="1203" pagination="99 - 103" refId="ref24764" refString="Guidoti M, Barcellos A, Ferreira RL. 2014. Novelties from a cave lace bug (Heteroptera, Tingidae): first nymphal description with a discussion on the morphological patterns of Vianaidinae immatures. Iheringia 104: 99-103." type="journal article" year="2014">
Guidoti
<emphasis id="09B59C610772FFFBFCCDFBEC3C5EFB7C" box="[809,866,1150,1172]" italics="true" pageId="31" pageNumber="1203">et al.</emphasis>
, 2014
</bibRefCitation>
) and now both
<taxonomicName id="FCC13BF00772FFFBFBBDFBEC3BF7FB7C" authorityName="Guidoti &amp; Montemayor &amp; Campos &amp; Guilbert" authorityYear="2020" box="[1113,1227,1150,1172]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="09B59C610772FFFBFBBDFBEC3BF7FB7C" box="[1113,1227,1150,1172]" italics="true" pageId="31" pageNumber="1203">T. insolita</emphasis>
</taxonomicName>
and
<taxonomicName id="FCC13BF00772FFFBFAE2FBEC3AADFB7B" authorityName="Guidoti &amp; Montemayor &amp; Campos &amp; Guilbert" authorityYear="2020" box="[1286,1425,1151,1172]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="urucuana">
<emphasis id="09B59C610772FFFBFAE2FBEC3AADFB7B" box="[1286,1425,1151,1172]" italics="true" pageId="31" pageNumber="1203">T. urucuana</emphasis>
</taxonomicName>
are also reported from that country.
<taxonomicName id="FCC13BF00772FFFBFB31FB0E3C41FB3E" authorityName="Guidoti &amp; Montemayor &amp; Campos &amp; Guilbert" authorityYear="2020" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="09B59C610772FFFBFB31FB0E3C41FB3E" italics="true" pageId="31" pageNumber="1203">Thaumamannia insolita</emphasis>
</taxonomicName>
and
<specimenCount id="2DC78BFA0772FFFBFC55FB2F3B79FB3D" box="[945,1093,1212,1234]" count="1" pageId="31" pageNumber="1203" type="generic">one specimen</specimenCount>
of
<taxonomicName id="FCC13BF00772FFFBFB80FB2F3BD6FB3E" authorityName="Guidoti &amp; Montemayor &amp; Campos &amp; Guilbert" authorityYear="2020" box="[1124,1258,1212,1233]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="urucuana">
<emphasis id="09B59C610772FFFBFB80FB2F3BD6FB3E" box="[1124,1258,1212,1233]" italics="true" pageId="31" pageNumber="1203">T. urucuana</emphasis>
</taxonomicName>
had its locality data reported as lost and we only know the state where they came from (Pará State,
<collectingCountry id="43D600E30772FFFBFB92FB6A3BFCFAE0" box="[1142,1216,1273,1295]" name="Brazil" pageId="31" pageNumber="1203">Brazil</collectingCountry>
). The only species with immature forms described is
<taxonomicName id="FCC13BF00772FFFBFB46FA8B3A7EFAC2" authorityName="Van Doesburg" authorityYear="1977" box="[1186,1346,1304,1325]" class="Insecta" family="Tingidae" genus="Thaumamannia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1203" phylum="Arthropoda" rank="species" species="vanderdrifti">
<emphasis id="09B59C610772FFFBFB46FA8B3A7EFAC2" box="[1186,1346,1304,1325]" italics="true" pageId="31" pageNumber="1203">T. vanderdrifti</emphasis>
</taxonomicName>
, which had its fifth instar described and analysed in SEM (
<bibRefCitation id="5F503D820772FFFBFCD5FAC63B38FA85" author="Guidoti M &amp; Barcellos A &amp; Ferreira RL" box="[817,1028,1365,1387]" pageId="31" pageNumber="1203" pagination="99 - 103" refId="ref24764" refString="Guidoti M, Barcellos A, Ferreira RL. 2014. Novelties from a cave lace bug (Heteroptera, Tingidae): first nymphal description with a discussion on the morphological patterns of Vianaidinae immatures. Iheringia 104: 99-103." type="journal article" year="2014">
Guidoti
<emphasis id="09B59C610772FFFBFC68FAC53CFEFA85" box="[908,962,1365,1387]" italics="true" pageId="31" pageNumber="1203">et al.</emphasis>
, 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
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</document>

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@ -0,0 +1,179 @@
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<heading id="01097011FFB6FFEF39B62DE1ACFDF947" box="[223,718,1740,1765]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="1322" reason="2">
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<taxonomicName id="9DFEBCFEFFB6FFEF385F2DE1AC79F947" authority="PATERSON" authorityName="PATERSON" authorityYear="2020" box="[310,586,1740,1765]" class="Mammalia" family="Mustelidae" genus="Corumictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus" status="gen. nov.">
<smallCapsWord id="5CA751A1FFB6FFEF385F2DE1AFFFF940" baselines="1758,1758" box="[310,460,1740,1764]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Corumictis" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF385F2DE1AFFFF940" bold="true" box="[310,460,1740,1764]" italics="true" pageId="4" pageNumber="1322">CORUMICTIS</emphasis>
</smallCapsWord>
<smallCapsWord id="5CA751A1FFB6FFEF38BA2DE1AC79F947" baselines="1759,1759" box="[467,586,1740,1765]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Paterson" pageId="4" pageNumber="1322">PATERSON</smallCapsWord>
</taxonomicName>
,
<emphasis id="688A1B6FFFB6FFEF3B3E2DFDACFDF947" bold="true" box="[599,718,1740,1764]" pageId="4" pageNumber="1322">
<taxonomicNameLabel id="73B9A614FFB6FFEF3B3E2DFDACFDF947" box="[599,718,1740,1764]" pageId="4" pageNumber="1322" rank="genus">
<smallCapsWord id="5CA751A1FFB6FFEF3B3E2DFDACBBF940" baselines="1758" box="[599,648,1744,1763]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="gen" pageId="4" pageNumber="1322">GEN</smallCapsWord>
.
<smallCapsWord id="5CA751A1FFB6FFEF3BFF2DFDACF4F940" baselines="1758" box="[662,711,1744,1763]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="4" pageNumber="1322">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5A41C77DFFB6FFEF39CA2DD9AF6EF88F" blockId="4.[163,732,1780,1836]" pageId="4" pageNumber="1322">
zoobank.org:act:
<uuid id="2E58FDA8FFB6FFEF380F2DD9AF6EF88F" pageId="4" pageNumber="1322">099E3F36-EAD9-4C9D-B547- 1176692E7C97</uuid>
</paragraph>
</subSubSection>
<subSubSection id="12E494F6FFB6FFEF39CA2C7EABB3F923" pageId="4" pageNumber="1322" type="materials_examined">
<paragraph id="5A41C77DFFB6FFEF39CA2C7EAB7FFDAD" blockId="4.[163,703,1874,1896]" lastBlockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF39CA2C7EAF01F8CB" box="[163,306,1875,1896]" italics="true" pageId="4" pageNumber="1322">
<typeStatus id="854579DFFFB6FFEF39CA2C7EAEEBF8CB" box="[163,216,1875,1896]" pageId="4" pageNumber="1322">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="9DFEBCFEFFB6FFEF38502C7EAC8CF8C4" authority="Paterson et al." authorityName="Paterson" box="[313,703,1874,1896]" class="Mammalia" family="Mustelidae" genus="Corumictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="species" species="wolsani">
<emphasis id="688A1B6FFFB6FFEF38502C7EAC26F8C4" box="[313,533,1874,1896]" italics="true" pageId="4" pageNumber="1322">Corumictis wolsani</emphasis>
Paterson
<emphasis id="688A1B6FFFB6FFEF3BEE2C7EAC8CF8C4" box="[647,703,1874,1896]" italics="true" pageId="4" pageNumber="1322">et al.</emphasis>
</taxonomicName>
<emphasis id="688A1B6FFFB6FFEF3A522BEBADF0FF78" box="[827,963,198,219]" italics="true" pageId="4" pageNumber="1322">Diagnosis:</emphasis>
Mustelid of small size with paired temporal crests. P4 large with anteriorly placed, conical protocone. M1 possesses a large parastylar shelf and lacks a postprotocrista. Suprameatal fossa is deep with medial arm nearly reaching level of lateral arm. Alisphenoid canal is absent. Postlateral sulcus on brain is absent. Lateral basioccipital embayment is present. Prominent nasofrontal swellings are present. The dentary is shallow and narrow. An m1 with open trigonid, reduced and posteriorly displaced metaconid, elongated paraconid and blade-like hypoconid.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A52293BAB97FD2B" blockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF3A52293BAA0DFD88" box="[827,1086,534,555]" italics="true" pageId="4" pageNumber="1322">Differential diagnosis:</emphasis>
Differs from
<taxonomicName id="9DFEBCFEFFB6FFEF3DB5293BAB0FFD88" authorityName="Pomel" authorityYear="1846" box="[1244,1340,534,555]" class="Mammalia" family="Mustelidae" genus="Plesictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus">
<emphasis id="688A1B6FFFB6FFEF3DB5293BAB0FFD88" box="[1244,1340,534,555]" italics="true" pageId="4" pageNumber="1322">Plesictis</emphasis>
</taxonomicName>
in being smaller in size, the presence of prominent nasofrontal swellings, deep suprameatal fossa, more gracile dentary, shortened/narrow talonid on m1, smaller m1 entoconid.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A3A29BCAB45FC81" blockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
Differs from
<taxonomicName id="9DFEBCFEFFB6FFEF3A9829BCAA47FD05" authorityName="Lange" authorityYear="1969" box="[1009,1140,657,678]" class="Mammalia" family="Mustelidae" genus="Mustelictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus">
<emphasis id="688A1B6FFFB6FFEF3A9829BCAA47FD05" box="[1009,1140,657,678]" italics="true" pageId="4" pageNumber="1322">Mustelictis</emphasis>
</taxonomicName>
in being smaller in size, bearing temporal crests that do not meet, the presence of prominent nasofrontal swellings, alisphenoid canal absent, deep suprameatal fossa, P4 lacking internal shelf, M2 single-rooted, and less prominent m1 entoconid.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A3A2807AAD1FC18" blockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
Differs from
<taxonomicName id="9DFEBCFEFFB6FFEF3A9A2807AA9EFC9C" authorityName="Pohle" authorityYear="1917" box="[1011,1197,810,831]" class="Mammalia" family="Procyonidae" genus="Pseudobassaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus">
<emphasis id="688A1B6FFFB6FFEF3A9A2807AA9EFC9C" box="[1011,1197,810,831]" italics="true" pageId="4" pageNumber="1322">Pseudobassaris</emphasis>
</taxonomicName>
in being smaller in size, the presence of prominent nasofrontal swellings, alisphenoid canal absent, P4 protocone anterior to paracone, M1 parastyle projects labially (not anterolabially) and M2 single-rooted.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A3A28E9AB35FBF7" blockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
Differs from
<taxonomicName id="9DFEBCFEFFB6FFEF3A9428E9AA4DFC7A" authorityName="Riggs" authorityYear="1942" box="[1021,1150,964,985]" class="Mammalia" family="Mustelidae" genus="Promartes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus">
<emphasis id="688A1B6FFFB6FFEF3A9428E9AA4DFC7A" box="[1021,1150,964,985]" italics="true" pageId="4" pageNumber="1322">Promartes</emphasis>
</taxonomicName>
in being smaller size, temporal crests do not meet, prominent nasofrontal swellings present, less prominent supraorbital processes, deep suprameatal fossa, reduced mastoid processes, M1 broader anteroposteriorly.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A3A2F70ABA8FB4D" blockId="4.[827,1444,197,1262]" pageId="4" pageNumber="1322">
Differs from
<taxonomicName id="9DFEBCFEFFB6FFEF3A852F70AA5DFBD1" box="[1004,1134,1117,1138]" class="Mammalia" family="Mustelidae" genus="Zodiolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="genus">
<emphasis id="688A1B6FFFB6FFEF3A852F70AA5DFBD1" box="[1004,1134,1117,1138]" italics="true" pageId="4" pageNumber="1322">Zodiolestes</emphasis>
</taxonomicName>
in: smaller size, temporal crests do not meet, prominent nasofrontal swellings present, less prominent supraorbital processes, lack of tubular external auditory meatus, reduced mastoid processes, unreduced M1 parastyle, M2 single-rooted.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A522E3BAB5DFAC9" blockId="4.[827,1444,1302,1386]" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF3A522E3BAA33FA8F" box="[827,1024,1302,1324]" italics="true" pageId="4" pageNumber="1322">Included species:</emphasis>
<taxonomicName id="9DFEBCFEFFB6FFEF3D6E2E3BABAEFA8F" authority="(Viret, 1929)" authorityName="Paterson &amp; Samuels &amp; Rybczynski &amp; Ryan &amp; Maddin" authorityYear="2020" baseAuthorityName="Viret" baseAuthorityYear="1929" box="[1031,1437,1302,1324]" class="Mammalia" family="Mustelidae" genus="Corumictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="species" species="julieni" status="comb. nov.">
<emphasis id="688A1B6FFFB6FFEF3D6E2E3BAAC6FA8F" bold="true" box="[1031,1269,1302,1324]" italics="true" pageId="4" pageNumber="1322">Corumictis julieni</emphasis>
(
<bibRefCitation id="3E6FBA8CFFB6FFEF3C6C2E3BABA7FA8F" author="Viret J" box="[1285,1428,1302,1324]" pageId="4" pageNumber="1322" pagination="1 - 328" refId="ref15158" refString="Viret J. 1929. Les faunes des mammiferes de l'Oligocene Superieur de la limagne Bourbonnaise. Les Annales de l'Universite de Lyon 47: 1-328." type="journal article" year="1929">Viret, 1929</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="73B9A614FFB6FFEF3A522E18ADF6FAE9" box="[827,965,1333,1354]" pageId="4" pageNumber="1322" rank="species">comb. nov.</taxonomicNameLabel>
(Basionym:
<taxonomicName id="9DFEBCFEFFB6FFEF3D322E18ABAFFAE9" authority="Viret, 1929" authorityName="Viret" authorityYear="1929" box="[1115,1436,1333,1355]" class="Mammalia" family="Mustelidae" genus="Plesictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="species" species="julieni">
<emphasis id="688A1B6FFFB6FFEF3D322E18AB22FAE9" box="[1115,1297,1333,1354]" italics="true" pageId="4" pageNumber="1322">Plesictis julieni</emphasis>
<bibRefCitation id="3E6FBA8CFFB6FFEF3C702E18ABAFFAE9" author="Viret J" box="[1305,1436,1333,1355]" pageId="4" pageNumber="1322" pagination="1 - 328" refId="ref15158" refString="Viret J. 1929. Les faunes des mammiferes de l'Oligocene Superieur de la limagne Bourbonnaise. Les Annales de l'Universite de Lyon 47: 1-328." type="journal article" year="1929">Viret, 1929</bibRefCitation>
</taxonomicName>
;
<emphasis id="688A1B6FFFB6FFEF3A522E79AAA8FACA" bold="true" box="[827,1179,1363,1385]" pageId="4" pageNumber="1322">
<taxonomicName id="9DFEBCFEFFB6FFEF3A522E79AA02FACA" authorityName="Paterson &amp; Samuels &amp; Rybczynski &amp; Ryan &amp; Maddin" authorityYear="2020" box="[827,1073,1363,1385]" class="Mammalia" family="Mustelidae" genus="Corumictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="1322" phylum="Chordata" rank="species" species="wolsani" status="sp. nov.">
<emphasis id="688A1B6FFFB6FFEF3A522E79AA02FACA" bold="true" box="[827,1073,1363,1385]" italics="true" pageId="4" pageNumber="1322">Corumictis wolsani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="73B9A614FFB6FFEF3D572E79AAA8FACA" box="[1086,1179,1364,1385]" pageId="4" pageNumber="1322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described below).
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A522EBFAA45FA46" blockId="4.[827,1442,1426,1510]" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF3A522EBFADFCFA04" box="[827,975,1426,1447]" italics="true" pageId="4" pageNumber="1322">Distribution</emphasis>
: Early and Late Oligocene (Early and Late Arikareean, Ar1Ar3) of
<collectingRegion id="983A099FFFB6FFEF3DC62E9CAB3AFA65" box="[1199,1289,1457,1478]" country="United States of America" name="Oregon" pageId="4" pageNumber="1322">Oregon</collectingRegion>
,
<collectingCountry id="22E987EDFFB6FFEF3C7E2E9CAB62FA65" box="[1303,1361,1457,1478]" name="United States of America" pageId="4" pageNumber="1322">USA</collectingCountry>
; Early Miocene (
<collectionCode id="3CEF5FB8FFB6FFEF3ACE2EFDADE0FA46" box="[935,979,1488,1509]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="1322">MN</collectionCode>
2a) of
<collectingCountry id="22E987EDFFB6FFEF3D492EFDAA41FA46" box="[1056,1138,1488,1509]" name="France" pageId="4" pageNumber="1322">France</collectingCountry>
.
</paragraph>
<paragraph id="5A41C77DFFB6FFEF3A522D23ABB3F923" blockId="4.[827,1442,1550,1664]" pageId="4" pageNumber="1322">
<emphasis id="688A1B6FFFB6FFEF3A522D23AD8AF980" box="[827,953,1550,1571]" italics="true" pageId="4" pageNumber="1322">Etymology</emphasis>
: From the Latin
<emphasis id="688A1B6FFFB6FFEF3DE62D23AB3BF987" box="[1167,1288,1550,1572]" italics="true" pageId="4" pageNumber="1322">ad Corum</emphasis>
, north-west, referring to the dispersal of mustelids to and from the north-western portion of North America, and the Greek ικτις,
<emphasis id="688A1B6FFFB6FFEF3AAC2D46ADC7F923" box="[965,1012,1643,1664]" italics="true" pageId="4" pageNumber="1322">ictis</emphasis>
, a weasel-like carnivorous animal.
</paragraph>
</subSubSection>
</treatment>
</document>