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data/03/82/23/0382231AFFD3FFA91398FEC3FC40F8DA.xml
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<mods:identifier id="C93E7331DD45949A46B11A39BFBA6738" type="DOI">10.11646/zootaxa.5514.5.1</mods:identifier>
<mods:identifier id="CBC0381103BE3CC332FB6903BA78C64C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="05D8090EAA4300E6032D967D1BFAB227" type="Zenodo-Dep">13914727</mods:identifier>
<mods:identifier id="F7F00094FD0C054C7E4A16DBE6D6A9D0" type="ZooBank">615D8E3D-9216-4CCC-9A6D-A6AFDA4C6FE3</mods:identifier>
</mods:mods>
<treatment id="03DB4C6DFFCCFFAB24FBF874FEC363B1" LSID="urn:lsid:zoobank.org:act:06118480-8423-403D-8B64-336E3F6C73E6" httpUri="http://treatment.plazi.org/id/03DB4C6DFFCCFFAB24FBF874FEC363B1" lastPageId="23" lastPageNumber="424" pageId="16" pageNumber="417">
<subSubSection id="C368AEF0FFCCFFAC24FBF874FC1E6735" pageId="16" pageNumber="417" type="nomenclature">
<treatment id="03DB4C6DFFCCFFAB24FBF874FEC363B1" ID-DOI="http://doi.org/10.5281/zenodo.13915062" ID-Zenodo-Dep="13915062" LSID="urn:lsid:zoobank.org:act:06118480-8423-403D-8B64-336E3F6C73E6" httpUri="http://treatment.plazi.org/id/03DB4C6DFFCCFFAB24FBF874FEC363B1" lastPageId="23" lastPageNumber="424" pageId="16" pageNumber="417">
<subSubSection id="C368AEF0FFCCFFAC24FBF874FE586751" box="[151,493,1806,1833]" pageId="16" pageNumber="417" type="nomenclature">
<paragraph id="8BCDFD7BFFCCFFAC24FBF874FE586751" blockId="16.[151,939,1806,1905]" box="[151,493,1806,1833]" pageId="16" pageNumber="417">
<heading id="D0854A17FFCCFFAC24FBF874FE586751" bold="true" box="[151,493,1806,1833]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
<emphasis id="B9062169FFCCFFAC24FBF874FE586751" bold="true" box="[151,493,1806,1833]" pageId="16" pageNumber="417">
@ -60,14 +64,14 @@
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFCCFFAC24FBF849FEAA6709" pageId="16" pageNumber="417" type="description">
<paragraph id="8BCDFD7BFFCCFFAC24FBF849FC1E6735" blockId="16.[151,939,1806,1905]" box="[151,939,1843,1869]" pageId="16" pageNumber="417">
<uri id="FFE3F179FFCCFFAC24FBF849FC1E6735" box="[151,939,1843,1869]" pageId="16" pageNumber="417">
urn:lsid:zoobank.org:act:
<uuid id="FFD4C7AEFFCCFFAC25C6F849FC1E6735" box="[426,939,1843,1869]" pageId="16" pageNumber="417">06118480-8423-403D-8B64-336E3F6C73E6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFCCFFAC24FBF82DFEAA6709" box="[151,287,1879,1905]" pageId="16" pageNumber="417" type="description">
<paragraph id="8BCDFD7BFFCCFFAC24FBF82DFEAA6709" blockId="16.[151,939,1806,1905]" box="[151,287,1879,1905]" pageId="16" pageNumber="417">
(
<figureCitation id="1349E1FEFFCCFFAC24F3F82DFEA26709" box="[159,279,1879,1905]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="17.[151,250,1757,1781]" captionStartId-1="18.[151,250,890,914]" captionStartId-2="21.[151,250,1761,1785]" captionTargetBox-0="[158,1436,355,1724]" captionTargetBox-1="[151,1435,181,866]" captionTargetBox-2="[163,1432,191,1737]" captionTargetId-0="figure-89@17.[151,1436,345,1734]" captionTargetId-1="figure-19@18.[151,1435,181,866]" captionTargetId-2="figure-18@21.[155,1432,181,1737]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="21" captionText-0="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." captionText-1="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove." captionText-2="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi-0="http://doi.org/10.5281/zenodo.13914747" figureDoi-1="http://doi.org/10.5281/zenodo.13914749" figureDoi-2="http://doi.org/10.5281/zenodo.13914751" httpUri-0="https://zenodo.org/record/13914747/files/figure.png" httpUri-1="https://zenodo.org/record/13914749/files/figure.png" httpUri-2="https://zenodo.org/record/13914751/files/figure.png" pageId="16" pageNumber="417">Figs. 9–11</figureCitation>
@ -78,20 +82,18 @@ urn:lsid:zoobank.org:act:
<paragraph id="8BCDFD7BFFCCFFAD24FBF8E5FEAF6139" blockId="16.[151,1436,1950,2013]" lastBlockId="17.[151,1437,150,321]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
<emphasis id="B9062169FFCCFFAC24FBF8E5FEAE67C0" bold="true" box="[151,283,1950,1977]" pageId="16" pageNumber="417">Type series</emphasis>
.
<materialsCitation id="3B1AF726FFCCFFAD254BF8E5FC8A60C9" collectingDate="2009-01-15" collectionCode="PNAH" collectorName="R. Barba" country="Cuba" county="Baracoa Municipality" elevation="500" lastPageId="17" lastPageNumber="418" latitude="20.4575" location="Trail" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" pageId="16" pageNumber="417" specimenCode="CZACC 3.4380" specimenCount="1" specimenCount-male="1" stateProvince="Guantanamo" typeStatus="holotype">
<materialsCitation id="3B1AF726FFCCFFAD254BF8E5FC8A60C9" collectingDate="2009-01-15" collectionCode="PNAH" collectorName="R. Barba" country="Cuba" county="Baracoa Municipality" elevation="500" lastPageId="17" lastPageNumber="418" latitude="20.4575" location="Monte Iberia" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" pageId="16" pageNumber="417" specimenCode="CZACC 3.4380" specimenCount="1" specimenCount-male="1" stateProvince="Guantanamo province" typeStatus="holotype">
<typeStatus id="54C943D9FFCCFFAC254BF8E5FE3A67C1" box="[295,399,1951,1977]" pageId="16" pageNumber="417" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7436F2FFCCFFAC25F9F8E5FE1F67C1" box="[405,426,1951,1977]" count="1" pageId="16" pageNumber="417" type="male">♂</specimenCount>
(α) (
<specimenCode id="DBD45500FFCCFFAC25B2F8E5FD2167C1" box="[478,660,1951,1977]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="16" pageNumber="417">CZACC 3.4380</specimenCode>
),
<location id="8EADABA0FFCCFFAC26CAF8E5FD6E67C1" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC26CAF8E5FD6E67C1" box="[678,731,1951,1977]" country="Cuba" county="Baracoa Municipality" latitude="20.4575" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" name="Trail" pageId="16" pageNumber="417" stateProvince="Guantanamo">Trail</location>
<locationDeviation id="9A7FE459FFCCFFAC26CAF8E5FD6E67C1" box="[678,731,1951,1977]" pageId="16" pageNumber="417">Trail</locationDeviation>
to the lagoon of
<location id="8EADABA0FFCCFFAC27FFF8E5FC6967C1" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC27FFF8E5FC6967C1" box="[915,988,1951,1977]" country="Cuba" county="Baracoa Municipality" latitude="20.4575" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" name="Monte" pageId="16" pageNumber="417" stateProvince="Guantanamo">Monte</location>
Iberia,
<collectingMunicipality id="6BA96701FFCCFFAC2043F8E5FF7F67A5" pageId="16" pageNumber="417">Alejandro de Humboldt National Park</collectingMunicipality>
(
<collectionCode id="ED6365BEFFCCFFAC24B7F8B9FE9867A5" box="[219,301,1987,2013]" pageId="16" pageNumber="417">PNAH</collectionCode>
),
<location id="8EADABA0FFCCFFAC27FFF8E5FB9367C1" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC27FFF8E5FB9367C1" box="[915,1062,1951,1977]" country="Cuba" county="Baracoa Municipality" latitude="20.4575" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" name="Monte Iberia" pageId="16" pageNumber="417" stateProvince="Guantanamo province">Monte Iberia</location>
,
<location id="8EADABA0FFCCFFAC2043F8E5FF7F67A5" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCCFFAC2043F8E5FF7F67A5" country="Cuba" county="Baracoa Municipality" latitude="20.4575" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" name="Alejandro de Humboldt National Park" pageId="16" pageNumber="417" stateProvince="Guantanamo province">Alejandro de Humboldt National Park</location>
(PNAH),
<collectingCounty id="62AC85F7FFCCFFAC252DF8B9FD8267A5" box="[321,567,1987,2013]" pageId="16" pageNumber="417">Baracoa Municipality</collectingCounty>
,
<collectingRegion id="49B63399FFCCFFAC262FF8B9FCF567A5" box="[579,832,1987,2013]" country="Cuba" name="Guantanamo" pageId="16" pageNumber="417">Guantánamo province</collectingRegion>
@ -114,15 +116,13 @@ a.s.l.
<collectorName id="268798ADFFCDFFAD26F6FFEDFD4A60C9" box="[666,767,151,177]" pageId="17" pageNumber="418">R. Barba</collectorName>
coll..
</materialsCitation>
<materialsCitation id="3B1AF726FFCDFFAD2725FFEDFE8560AD" collectionCode="CZACC" country="Cuba" pageId="17" pageNumber="418" specimenCode="CZACC 3.4380" specimenCount="1" specimenCount-female="1" stateProvince="Guantanamo" typeStatus="paratype">
<materialsCitation id="3B1AF726FFCDFFAD2725FFEDFE8560AD" collectingDate="2009-01-15" collectionCode="CZACC" collectorName="R. Barba" country="Cuba" county="Baracoa Municipality" elevation="500" latitude="20.4575" location="Monte Iberia" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" pageId="17" pageNumber="418" specimenCode="CZACC 3.4380" specimenCount="1" specimenCount-female="1" stateProvince="Guantanamo" typeStatus="paratype">
<typeStatus id="54C943D9FFCDFFAD2725FFEDFC0F60C9" box="[841,954,151,177]" pageId="17" pageNumber="418" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7436F2FFCDFFAD27AAFFEDFC4760C9" box="[966,1010,151,177]" count="1" pageId="17" pageNumber="418" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBD45500FFCDFFAD206EFFEDFB0F60C9" box="[1026,1210,151,177]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="17" pageNumber="418">CZACC 3.4380</specimenCode>
, in the same vial as the
<typeStatus id="54C943D9FFCDFFAD24ADFFC1FE9D60AD" box="[193,296,187,213]" pageId="17" pageNumber="418" type="holotype">holotype</typeStatus>
)
, in the same vial as the holotype)
</materialsCitation>
;
<materialsCitation id="3B1AF726FFCDFFAD2550FFC1FCDF60AD" box="[316,874,187,213]" collectingDate="2009-01-15" collectionCode="CZACC" collectorName="R. Barba" country="Cuba" county="Baracoa Municipality" elevation="500" latitude="20.4575" location="Trail" longLatPrecision="21" longitude="-74.72639" municipality="Alejandro de Humboldt National Park" pageId="17" pageNumber="418" specimenCode="CZACC 3.3391" specimenCount="1" specimenCount-male="1" stateProvince="Guantanamo" typeStatus="paratype">
@ -132,12 +132,12 @@ coll..
), same data as the holotype
</materialsCitation>
;
<materialsCitation id="3B1AF726FFCDFFAD271AFFC1FC3F6081" collectingDate="1996-10-01" collectionCode="CZACC" country="Cuba" latitude="20.4306" location="Rio Piloto" longLatPrecision="14" longitude="-74.88555" municipality="Moa" pageId="17" pageNumber="418" specimenCode="CZACC 3.3393" specimenCount="1" specimenCount-female="1" stateProvince="Holguin" typeStatus="paratype">
<materialsCitation id="3B1AF726FFCDFFAD271AFFC1FC3F6081" collectingDate="1996-10-01" collectionCode="CZACC" country="Cuba" latitude="20.4306" location="Rio Piloto" longLatPrecision="14" longitude="-74.88555" municipality="Moa" pageId="17" pageNumber="418" specimenCode="CZACC 3.3393" specimenCount="1" specimenCount-female="1" stateProvince="Holguin province" typeStatus="paratype">
<specimenCount id="9D7436F2FFCDFFAD271AFFC1FC1560AD" box="[886,928,187,213]" count="1" pageId="17" pageNumber="418" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBD45500FFCDFFAD27DCFFC1FBDD60AD" box="[944,1128,187,213]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="17" pageNumber="418">CZACC 3.3393</specimenCode>
),
<location id="8EADABA0FFCDFFAD2010FFC1FB4760AD" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCDFFAD2010FFC1FB4760AD" box="[1148,1266,187,213]" country="Cuba" latitude="20.4306" longLatPrecision="14" longitude="-74.88555" municipality="Moa" name="Rio Piloto" pageId="17" pageNumber="418" stateProvince="Holguin">Río Piloto</location>
<location id="8EADABA0FFCDFFAD2010FFC1FB4760AD" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCDFFAD2010FFC1FB4760AD" box="[1148,1266,187,213]" country="Cuba" latitude="20.4306" longLatPrecision="14" longitude="-74.88555" municipality="Moa" name="Rio Piloto" pageId="17" pageNumber="418" stateProvince="Holguin province">Río Piloto</location>
,
<collectingMunicipality id="6BA96701FFCDFFAD2092FFC1FA8160AD" box="[1278,1332,187,213]" pageId="17" pageNumber="418">Moa</collectingMunicipality>
,
@ -155,12 +155,12 @@ coll..
(no coll.)
</materialsCitation>
;
<materialsCitation id="3B1AF726FFCDFFAD27F5FFA5FEAF6139" collectingDate="1996-10-04" collectionCode="CZACC" collectorName="N. Vinas Davila" country="Cuba" latitude="20.442554" location="Toldo" longLatPrecision="1" longitude="-74.860245" municipality="Moa" pageId="17" pageNumber="418" specimenCode="CZACC 3.3392" specimenCount="1" specimenCount-male="1" stateProvince="Holguin" typeStatus="paratype">
<materialsCitation id="3B1AF726FFCDFFAD27F5FFA5FEAF6139" collectingDate="1996-10-04" collectingMethod="yellow dish trap" collectionCode="CZACC" collectorName="N. Vinas Davila" country="Cuba" latitude="20.442554" location="Toldo" longLatPrecision="1" longitude="-74.860245" municipality="Moa" pageId="17" pageNumber="418" specimenCode="CZACC 3.3392" specimenCount="1" specimenCount-male="1" stateProvince="Holguin province" typeStatus="paratype">
<specimenCount id="9D7436F2FFCDFFAD27F5FFA5FC736081" box="[921,966,223,249]" count="1" pageId="17" pageNumber="418" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBD45500FFCDFFAD27B4FFA5FB266081" box="[984,1171,223,249]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="17" pageNumber="418">CZACC 3.3392</specimenCode>
),
<location id="8EADABA0FFCDFFAD20C5FFA5FB586081" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCDFFAD20C5FFA5FB586081" box="[1193,1261,223,249]" country="Cuba" latitude="20.442554" longLatPrecision="1" longitude="-74.860245" municipality="Moa" name="Toldo" pageId="17" pageNumber="418" stateProvince="Holguin">Toldo</location>
<location id="8EADABA0FFCDFFAD20C5FFA5FB586081" LSID="urn:lsid:plazi:treatment:03DB4C6DFFCCFFAB24FBF874FEC363B1:8EADABA0FFCDFFAD20C5FFA5FB586081" box="[1193,1261,223,249]" country="Cuba" latitude="20.442554" longLatPrecision="1" longitude="-74.860245" municipality="Moa" name="Toldo" pageId="17" pageNumber="418" stateProvince="Holguin province">Toldo</location>
,
<collectingMunicipality id="6BA96701FFCDFFAD2090FFA5FA876081" box="[1276,1330,223,249]" pageId="17" pageNumber="418">Moa</collectingMunicipality>
,
@ -171,8 +171,8 @@ coll..
<geoCoordinate id="EE469BBCFFCDFFAD2535FE79FE466165" box="[345,499,259,285]" degrees="20" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="418" precision="1" seconds="33.2" value="20.442554">20º26'33.2''N</geoCoordinate>
;
<geoCoordinate id="EE469BBCFFCDFFAD266EFE78FD146165" box="[514,673,258,285]" degrees="74" direction="west" minutes="51" orientation="longitude" pageId="17" pageNumber="418" precision="1" seconds="36.9" value="-74.860245">74º51'36.9''W</geoCoordinate>
),
<collectingMethod id="5233856CFFCDFFAD26D4FE79FBB16165" box="[696,1028,259,285]" pageId="17" pageNumber="418">collected in yellow dish trap</collectingMethod>
), collected in
<collectingMethod id="5233856CFFCDFFAD2724FE79FBB16165" box="[840,1028,259,285]" pageId="17" pageNumber="418">yellow dish trap</collectingMethod>
on the ground,
<date id="FFCCDBBBFFCDFFAD20AFFE79FA9D6165" box="[1219,1320,259,285]" pageId="17" pageNumber="418" value="1996-10-04">
<collectingDate id="EF882253FFCDFFAD20AFFE79FA9D6165" box="[1219,1320,259,285]" pageId="17" pageNumber="418" value="1996-10-04">4.x.1996</collectingDate>
@ -399,9 +399,9 @@ male (CZACC 3.4380)
</paragraph>
<paragraph id="8BCDFD7BFFCEFFAE24ABF8CBFE7967B3" blockId="18.[151,1437,1177,1995]" box="[199,460,1969,1995]" pageId="18" pageNumber="419">
<emphasis id="B9062169FFCEFFAE24ABF8CBFEC667B2" box="[199,371,1969,1994]" italics="true" pageId="18" pageNumber="419">Measurements:</emphasis>
<tableCitation id="C6F0C8C0FFCEFFAE2515F8CBFE7967B3" box="[377,460,1969,1995]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." pageId="18" pageNumber="419">Table 3</tableCitation>
<tableCitation id="C6F0C8C0FFCEFFAE2515F8CBFE7967B3" box="[377,460,1969,1995]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionTargetPageId="19" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" pageId="18" pageNumber="419" tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
</paragraph>
<caption id="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" box="[152,1342,151,176]" pageId="19" pageNumber="420" startId="19.[152,244,151,175]" targetType="table">
<caption id="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" ID-Table-UUID="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" box="[152,1342,151,176]" httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" pageId="19" pageNumber="420" startId="19.[152,244,151,175]" targetBox="[163,1410,192,1815]" targetIsTable="true" targetPageId="19" targetType="table">
<paragraph id="8BCDFD7BFFCFFFAF24F4FFEDFA8B60D7" blockId="19.[152,1342,151,176]" box="[152,1342,151,176]" pageId="19" pageNumber="420">
<emphasis id="B9062169FFCFFFAF24F4FFEDFEBD60D7" bold="true" box="[152,264,151,175]" pageId="19" pageNumber="420">TABLE 3</emphasis>
. Measurements (in mm) of the male holotype (H) and paratypes (P) of
@ -416,7 +416,7 @@ male (CZACC 3.4380)
</caption>
<paragraph id="8BCDFD7BFFCFFFAF25C4FFBAFAF7676F" pageId="19" pageNumber="420">
<table id="F9720FDBFFCF004324CFFFBAFA37676F" box="[163,1410,192,1815]" gridcols="6" gridrows="40" pageId="19" pageNumber="420">
<tr id="3542FF39FFCF004324CFFFBAFA37608D" box="[163,1410,192,245]" gridrow="0" pageId="19" pageNumber="420" rowspan-0="1">
<tr id="3542FF39FFCF004324CFFFBAFA37608D" box="[163,1410,192,245]" gridrow="0" isHeader="true" pageId="19" pageNumber="420" rowspan-0="1">
<th id="76939645FFCF00432519FFBAFDA0608D" box="[373,533,192,245]" gridcol="1" gridrow="0" pageId="19" pageNumber="420">H ♂α CZACC 3.4380</th>
<th id="76939645FFCF00432609FFBAFCB3608D" box="[613,774,192,245]" gridcol="2" gridrow="0" pageId="19" pageNumber="420">P ♂α CZACC 3.3392</th>
<th id="76939645FFCF00432724FFBAFC5C608D" box="[840,1001,192,245]" gridcol="3" gridrow="0" pageId="19" pageNumber="420">P ♂β CZACC 3.3391</th>
@ -946,13 +946,13 @@ Female
<typeStatus id="54C943D9FFCBFFAB2544FECDFE2661A9" box="[296,403,439,465]" pageId="23" pageNumber="424" type="paratype">paratype</typeStatus>
</emphasis>
(CZACC 3.4380). Similar to male, but slightly slenderer. Free tergite III with four small tubercles between the pair of spiniform apophyses. Chelicerae small, non hypertelic, without a gap between the cheliceral fingers as in α males; fixed and movable fingers with six distal teeth. Legs III–IV with femur, patella, and tibia slightly thinner than in males; metatarsus III not enlarged. Femora II–IV with a distal spiniform apophysis, as in the male. Coloration pattern as in males. Genital operculum wider than the male. Tarsal formula: 7(3):12(4):8:8. Measurements:
<tableCitation id="C6F0C8C0FFCBFFAB2525FD11FE2B62FD" box="[329,414,619,645]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." pageId="23" pageNumber="424">Table 3</tableCitation>
<tableCitation id="C6F0C8C0FFCBFFAB2525FD11FE2B62FD" box="[329,414,619,645]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionTargetPageId="19" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" pageId="23" pageNumber="424" tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BCDFD7BFFCBFFAB24ABFDF4FA37636D" blockId="23.[151,1437,151,969]" pageId="23" pageNumber="424">
<emphasis id="B9062169FFCBFFAB24ABFDF4FEF262D0" bold="true" box="[199,327,654,680]" pageId="23" pageNumber="424">Variability</emphasis>
. Males (n=3); females (n=2). Measurements in
<tableCitation id="C6F0C8C0FFCBFFAB270CFDF5FC0062D1" box="[864,949,655,681]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." pageId="23" pageNumber="424">Table 3</tableCitation>
<tableCitation id="C6F0C8C0FFCBFFAB270CFDF5FC0062D1" box="[864,949,655,681]" captionStart="TABLE 3" captionStartId="19.[152,244,151,175]" captionTargetPageId="19" captionText="TABLE 3. Measurements (in mm) of the male holotype (H) and paratypes (P) of Caribbiantes barbai sp. nov.." httpUri="http://table.plazi.org/id/DF0DADF3FFCFFFAF24F4FFEDFA8B60D7" pageId="23" pageNumber="424" tableUuid="DF0DADF3FFCFFFAF24F4FFEDFA8B60D7">Table 3</tableCitation>
. Cheliceral dentition: fixed finger with 5–7 distal teeth, movable finger with 6–8 distal teeth. In the swollen region of metatarsus III each studied male had a different number of setae (26, 31, 32). The tarsal formula varies only in leg II: 7(3):12–15(4–6):8:8. Hypertelic chelicerae in only two studied males (α males—including the
<typeStatus id="54C943D9FFCBFFAB2721FD81FC01636D" box="[845,948,763,789]" pageId="23" pageNumber="424" type="holotype">holotype</typeStatus>
);
@ -960,7 +960,7 @@ Female
has small chelicerae (ss male).
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFCBFFAB24ABFC64FEC363B1" pageId="23" pageNumber="424" type="distribution">
<subSubSection id="C368AEF0FFCBFFAB24ABFC64FE1963F9" pageId="23" pageNumber="424" type="distribution">
<paragraph id="8BCDFD7BFFCBFFAB24ABFC64FE1963F9" blockId="23.[151,1437,151,969]" pageId="23" pageNumber="424">
<emphasis id="B9062169FFCBFFAB24ABFC64FED46340" bold="true" box="[199,353,798,824]" pageId="23" pageNumber="424">Distribution.</emphasis>
This species has the eastern-most records for the genus. It is likely a Cuban short-ranged endemic, so far restricted to three localities in Toldo and Monte
@ -969,6 +969,8 @@ Plateaus (Holguín and Guantánamo provinces, respectively) (
<figureCitation id="1349E1FEFFCBFFAB255BFC1DFE1563F9" box="[311,416,871,897]" captionStart="FIGURE 12" captionStartId="22.[151,250,1756,1780]" captionTargetBox="[164,1423,181,1732]" captionTargetId="figure-19@22.[164,1423,181,1732]" captionTargetPageId="22" captionText="FIGURE 12. Records and habitat of Caribbiantes spp. A. Map of Cuba (general and expanded sector as indicated), showing known localities of Caribbiantes cubanus (circles, type locality in red); C. obtusus sp. nov. (triangles, type locality in red) and C. barbai sp. nov. (squares, type locality in red); B. Submontane rain forest in Alejandro de Humboldt National Park, Monte Iberia, type locality of C. barbai sp. nov.; C. Mountains of Topes de Collantes, Sancti Spíritus province, record of C. cubanus and C. obtusus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13914753" httpUri="https://zenodo.org/record/13914753/files/figure.png" pageId="23" pageNumber="424">Fig. 12A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFCBFFAB24ABFCF1FEC363B1" pageId="23" pageNumber="424" type="biology_ecology">
<paragraph id="8BCDFD7BFFCBFFAB24ABFCF1FEC363B1" blockId="23.[151,1437,151,969]" pageId="23" pageNumber="424">
<emphasis id="B9062169FFCBFFAB24ABFCF1FECA63DC" bold="true" box="[199,383,906,933]" pageId="23" pageNumber="424">Natural history</emphasis>
. This species was collected under the bark of living trees, in the sub-montane rainforest at

1393
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<heading id="D0854A17FFDEFFBE24FBFDC8FD8262B4" bold="true" box="[151,567,690,720]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B9062169FFDEFFBE24FBFDC8FD8262B4" bold="true" box="[151,567,690,720]" pageId="2" pageNumber="403">
Genus
<taxonomicName id="4C7286F8FFDEFFBE2482FDC8FD8262B4" ID-CoL="3HHR" authority="Silhavy, 1973" authorityName="Silhavy" authorityYear="1973" box="[238,567,690,720]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE2482FDC8FE3C62B5" bold="true" box="[238,393,690,717]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
<bibRefCitation id="EFE3808AFFDEFFBE25FDFDC9FD8262B4" author="Silhavy, V." box="[401,567,690,720]" pageId="2" pageNumber="403" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý, 1973</bibRefCitation>
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<treatmentCitation id="0AD3DB6AFFDEFFBE24FBFC61FE6A634B" author="Silhavy, V." box="[151,479,795,821]" page="135" pageId="2" pageNumber="403" year="1973">
<taxonomicName id="4C7286F8FFDEFFBE24FBFC61FE6A634B" ID-CoL="3HHR" authority="Silhavy 1973: 135" authorityName="Silhavy" authorityPageNumber="135" authorityYear="1973" box="[151,479,795,821]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE24FBFC61FEAE634B" box="[151,283,795,819]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
<bibRefCitation id="EFE3808AFFDEFFBE254EFC66FE6A634B" author="Silhavy, V." box="[290,479,795,821]" pageId="2" pageNumber="403" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý 1973: 135</bibRefCitation>
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; Kury 2003: 35.
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<emphasis id="B9062169FFDEFFBE24FBFC1BFE9E6303" bold="true" box="[151,299,865,891]" pageId="2" pageNumber="403">
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species
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.
<taxonomicName id="4C7286F8FFDEFFBE2555FC1BFD606303" ID-CoL="7RXXC" authority="Silhavy, 1973" authorityName="Silhavy" authorityYear="1973" box="[313,725,865,894]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="cubanus">
<emphasis id="B9062169FFDEFFBE2555FC1BFD9A6303" box="[313,559,865,891]" italics="true" pageId="2" pageNumber="403">Caribbiantes cubanus</emphasis>
<bibRefCitation id="EFE3808AFFDEFFBE265BFC19FD606303" author="Silhavy, V." box="[567,725,865,894]" pageId="2" pageNumber="403" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý, 1973</bibRefCitation>
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, by original designation.
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<emphasis id="B9062169FFDEFFBE24ABFCFFFE3963E7" bold="true" box="[199,396,901,927]" pageId="2" pageNumber="403">Included species</emphasis>
.
<taxonomicName id="4C7286F8FFDEFFBE25F1FCFFFCF563E7" ID-CoL="7RXXC" authority="Silhavy, 1973" authorityName="Silhavy" authorityYear="1973" box="[413,832,901,930]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="cubanus">
<emphasis id="B9062169FFDEFFBE25F1FCFFFD2363E7" box="[413,662,901,927]" italics="true" pageId="2" pageNumber="403">Caribbiantes cubanus</emphasis>
<bibRefCitation id="EFE3808AFFDEFFBE26F3FCFDFCF563E7" author="Silhavy, V." box="[671,832,901,930]" pageId="2" pageNumber="403" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý, 1973</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7286F8FFDEFFBE2722FCFFFB8063E7" authorityName="Barroso &amp; Pérez-González" authorityYear="2024" box="[846,1077,901,927]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="barbai" status="sp. nov.">
<emphasis id="B9062169FFDEFFBE2722FCFFFB8063E7" box="[846,1077,901,927]" italics="true" pageId="2" pageNumber="403">Caribbiantes barbai</emphasis>
</taxonomicName>
<emphasis id="B9062169FFDEFFBE2052FCFFFB2F63E7" bold="true" box="[1086,1178,901,927]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A2359C12FFDEFFBE2052FCFFFB2F63E7" box="[1086,1178,901,927]" pageId="2" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4C7286F8FFDEFFBE20C7FCFFFA2863E7" authorityName="Barroso &amp; Pérez-González" authorityYear="2024" box="[1195,1437,901,927]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="B9062169FFDEFFBE20C7FCFFFA2863E7" box="[1195,1437,901,927]" italics="true" pageId="2" pageNumber="403">Caribbiantes obtusus</emphasis>
</taxonomicName>
<emphasis id="B9062169FFDEFFBE24FBFCD3FF4463BB" bold="true" box="[151,241,937,963]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A2359C12FFDEFFBE24FBFCD3FF4463BB" box="[151,241,937,963]" pageId="2" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFDEFFBF24ABFCB7FEA461D5" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="8BCDFD7BFFDEFFBE24ABFCB7FA3B665F" blockId="2.[151,1437,865,2008]" pageId="2" pageNumber="403">
<emphasis id="B9062169FFDEFFBE24ABFCB7FE1A639F" bold="true" box="[199,431,973,999]" pageId="2" pageNumber="403">Emended diagnosis</emphasis>
.
<taxonomicName id="4C7286F8FFDEFFBE25D3FCB7FDE4639F" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[447,593,973,999]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE25D3FCB7FDE4639F" box="[447,593,973,999]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
is differentiated from the rest of the
<taxonomicName id="4C7286F8FFDEFFBE2796FCB7FB10639F" ID-CoL="7NM37" authorityName="Roewer" authorityYear="1913" box="[1018,1189,973,999]" class="Arachnida" family="Biantidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="subFamily" subFamily="Stenostygninae">Stenostygninae</taxonomicName>
genera by having the following combination of characters: dorsal scutum sub-rectangular, strong spiniform apophyses on mesotergal areas III and IV of almost equal size, and free tergite III with a pair of robust spiniform apophyses or a pair of tubercles (
<figureCitation id="1349E1FEFFDEFFBE2561FB43FEC5642B" box="[269,368,1081,1107]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="2" pageNumber="403">Figs. 1B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2512FB43FE15642B" box="[382,416,1081,1107]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="2" pageNumber="403">6C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE25C2FB43FE67642B" box="[430,466,1081,1107]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="2" pageNumber="403">9C</figureCitation>
). Tibia of the pedipalps with the ventro-mesal spines equidistant (
<figureCitation id="1349E1FEFFDEFFBE20ACFB43FAA9642B" box="[1216,1308,1081,1107]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[236,1352,193,1907]" captionTargetId="figure-18@5.[224,1364,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Caribbiantes cubanus, male holotype (MCZ IZ-14702), right pedipalp (trochanter to tarsus): A. pedipalp mesal view; B. pedipalp ectal view. Abbreviations: Tr, trochanter; Fe, femur; Pa, patella; Ti, tibia; Ta, tarsus; TC, tarsal claw; S, spine; edS, ectodistal spines; g, gap in the tubercles series. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914731" httpUri="https://zenodo.org/record/13914731/files/figure.png" pageId="2" pageNumber="403">Fig. 2A</figureCitation>
). Chelicera strongly swollen (chelicera hand hypertelic) in α males, whereas β males exhibit small chelicera as in females; no intermediate sizes (grade) where observed between hypertelic and small chelicerae (
<figureCitation id="1349E1FEFFDEFFBE2027FBFBFB1864E3" box="[1099,1197,1153,1179]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="2" pageNumber="403">Figs. 1B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE20D6FBFBFB6964E3" box="[1210,1244,1153,1179]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="2" pageNumber="403">6C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2085FBFBFAB864E3" box="[1257,1293,1153,1179]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="2" pageNumber="403">9C</figureCitation>
). Metatarsus III of “samooidean
<typeStatus id="54C943D9FFDEFFBE251DFBDFFE1364C7" box="[369,422,1189,1215]" pageId="2" pageNumber="403">type</typeStatus>
”,
<emphasis id="B9062169FFDEFFBE25D0FBDCFE6A64C7" box="[444,479,1190,1215]" italics="true" pageId="2" pageNumber="403">i.e.</emphasis>
with a long intrusion of the astragalus by the calcaneus on the ventral surface (
<bibRefCitation id="EFE3808AFFDEFFBE213FFBDFFE35649B" author="Perez-Gonzalez, A. &amp; Sharma, P. P. &amp; Proud, D. N." pageId="2" pageNumber="403" pagination="253 - 260" refId="ref17182" refString="Perez-Gonzalez, A., Sharma, P. P. &amp; Proud, D. N. (2016) Morphological tricks and blessed genitalia: rectifying the family placement of Fijicolana tuberculata (Opiliones: Laniatores: Zalmoxidae). Zootaxa, 4061 (3), 253 - 260. https: // doi. org / 10.11646 / zootaxa. 4061.3.3" type="journal article" year="2016">
Pérez-González
<emphasis id="B9062169FFDEFFBE256BFBB0FE8B649B" box="[263,318,1225,1251]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2016
</bibRefCitation>
). Male sexually dimorphic metatarsus III with a subdistal enlargement, located mainly between the third and fourth distal quarter of the segment; the enlargement has a deep and narrow buttonhole-shaped ventral groove, which bears conspicuous, striated brush-like setae, twisted and distally enlarged in the shape of a long paint brush; the brush-like setae are longitudinally arranged in two alternate rows (
<figureCitation id="1349E1FEFFDEFFBE200AFA4FFB796537" box="[1126,1228,1333,1359]" captionStart="FIGURE 3" captionStartId="6.[151,250,890,914]" captionTargetBox="[151,1435,181,866]" captionTargetId="figure-19@6.[151,1435,181,866]" captionTargetPageId="6" captionText="FIGURE 3. Glandular structure of male metatarsus III in Caribbiantes cubanus. A. Light microscope photo, male (CZACC 3.4372), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E., Scanning electron micrographs, male (CZACC 3.3266): B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note its depth; D. Closer zoom into the groove, white arrow indicates a seta (long paint brush, basally striated and twisted), small arrowheads point to glandular openings (pores) irregularly arranged around the setae; E. Detail of the integument of the ventral groove." figureDoi="http://doi.org/10.5281/zenodo.13914733" httpUri="https://zenodo.org/record/13914733/files/figure.png" pageId="2" pageNumber="403">Figs. 3B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE20B0FA4CFB4A6528" box="[1244,1279,1334,1360]" captionStart="FIGURE 7" captionStartId="14.[151,250,1774,1798]" captionTargetBox="[151,1435,1065,1750]" captionTargetId="figure-443@14.[151,1435,1065,1750]" captionTargetPageId="14" captionText="FIGURE 7. Glandular structure of male metatarsus III in Caribbiantes obtusus sp. nov. (male CZACC 3.4379). A. Light microscope photo, general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs: B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, arrows show setae (long paint brush, basally striated and twisted); E. Integument of the ventral groove, white arrowheads indicate glandular openings (pores) irregularly arranged around the setae." figureDoi="http://doi.org/10.5281/zenodo.13914743" httpUri="https://zenodo.org/record/13914743/files/figure.png" pageId="2" pageNumber="403">7B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2163FA4FFAF46537" box="[1295,1345,1333,1359]" captionStart="FIGURE 10" captionStartId="18.[151,250,890,914]" captionTargetBox="[151,1435,181,866]" captionTargetId="figure-19@18.[151,1435,181,866]" captionTargetPageId="18" captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove." figureDoi="http://doi.org/10.5281/zenodo.13914749" httpUri="https://zenodo.org/record/13914749/files/figure.png" pageId="2" pageNumber="403">10B</figureCitation>
). Tarsal count: 6–7:10–15:7–8:7–8. General shape of the penis tubular with
<emphasis id="B9062169FFDEFFBE27F3FA20FB9E650B" box="[927,1067,1369,1395]" italics="true" pageId="2" pageNumber="403">pars distalis</emphasis>
slightly enlarged (
<figureCitation id="1349E1FEFFDEFFBE216EFA23FAD2650C" box="[1282,1383,1369,1396]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2119FA23FA22650B" box="[1397,1431,1369,1395]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE24FBFA07FF7D65EF" box="[151,200,1405,1431]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11C</figureCitation>
).
<emphasis id="B9062169FFDEFFBE24B2FA07FE2365EF" box="[222,406,1405,1431]" italics="true" pageId="2" pageNumber="403">Capsula interna</emphasis>
composed of a laterally flattened lance-shaped (narrow-leaf)
<emphasis id="B9062169FFDEFFBE2035FA07FB2C65EF" box="[1113,1177,1405,1431]" italics="true" pageId="2" pageNumber="403">stylus</emphasis>
and two basally fused conductors, medially thin and apically widened in laminar ends with the form of lobes or truncate, ventrally projected (
<figureCitation id="1349E1FEFFDEFFBE24F3FABFFEB76598" box="[159,258,1477,1504]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5E</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2563FABFFE8565A7" box="[271,304,1477,1503]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8E</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2552FABFFED865A7" box="[318,365,1477,1503]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11E</figureCitation>
).
<emphasis id="B9062169FFDEFFBE25EFFABFFE7365A7" box="[387,454,1477,1503]" italics="true" pageId="2" pageNumber="403">Stylus</emphasis>
slightly larger than conductors (
<figureCitation id="1349E1FEFFDEFFBE2755FABFFC2E6598" box="[825,923,1477,1504]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5E</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE27C4FABFFC7F65A7" box="[936,970,1477,1503]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8E</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE27BBFABFFBB365A7" box="[983,1030,1477,1503]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11E</figureCitation>
).
<emphasis id="B9062169FFDEFFBE2070FABCFB1F65A7" box="[1052,1194,1477,1503]" italics="true" pageId="2" pageNumber="403">Pars distalis</emphasis>
ventrally with apical process in the form of a low hook with a broad base, or with a rounded tip and two pairs of apical setae: setae of the apical pair small and bifurcate; the three pairs of ventrolateral setae longer and foliar (
<figureCitation id="1349E1FEFFDEFFBE2036F977FB626650" box="[1114,1239,1549,1576]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5D, F</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2089F977FA906650" box="[1253,1317,1549,1576]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8D, F</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE215DF977FA34665F" box="[1329,1409,1549,1575]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11D, F</figureCitation>
).
</paragraph>
<paragraph id="8BCDFD7BFFDEFFBF24ABF94BFEA461D5" blockId="2.[151,1437,865,2008]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<emphasis id="B9062169FFDEFFBE24ABF94BFED86633" bold="true" box="[199,365,1585,1611]" pageId="2" pageNumber="403">Comparisons.</emphasis>
<taxonomicName id="4C7286F8FFDEFFBE251FF94BFDB06633" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[371,517,1585,1611]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE251FF94BFDB06633" box="[371,517,1585,1611]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
is easily differentiated from the rest of the stenostygnine genera. Differences from
<taxonomicName id="4C7286F8FFDEFFBE24FBF92CFE9C6608" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[151,297,1622,1648]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE24FBF92CFE9C6608" box="[151,297,1622,1648]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
include the following:
<taxonomicName id="4C7286F8FFDEFFBE265CF92FFD776617" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[560,706,1621,1647]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE265CF92FFD776617" box="[560,706,1621,1647]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
has mesotergal areas and free tergites armed (only with granules or tubercles in
<taxonomicName id="4C7286F8FFDEFFBE2525F900FE6E66EC" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[329,475,1658,1684]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE2525F900FE6E66EC" box="[329,475,1658,1684]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
); area I is divided by a median longitudinal groove in
<taxonomicName id="4C7286F8FFDEFFBE200AF903FB4D66EB" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[1126,1272,1657,1683]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE200AF903FB4D66EB" box="[1126,1272,1657,1683]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
(undivided in
<taxonomicName id="4C7286F8FFDEFFBE24FBF9E4FE9C66C0" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[151,297,1694,1720]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE24FBF9E4FE9C66C0" box="[151,297,1694,1720]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
) (
<emphasis id="B9062169FFDEFFBE252AF9E7FED766CF" box="[326,354,1693,1719]" italics="true" pageId="2" pageNumber="403">cf.</emphasis>
<figureCitation id="1349E1FEFFDEFFBE2500F9E7FE4066CF" box="[364,501,1693,1719]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="2" pageNumber="403">Figs. 1A, C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2668F9E7FDF866C0" box="[516,589,1693,1720]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="2" pageNumber="403">6A, B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2637F9E7FD1166C0" box="[603,676,1693,1720]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="2" pageNumber="403">9A, B</figureCitation>
, and
<bibRefCitation id="EFE3808AFFDEFFBE2689F9E7FC7866CF" author="Mamani, C. V. &amp; Porto, W. &amp; Iglesias, P. P. &amp; Perez-Gonzalez, A." box="[741,973,1693,1719]" pageId="2" pageNumber="403" pagination="182 - 202" refId="ref17030" refString="Mamani, C. V., Porto, W., Iglesias, P. P. &amp; Perez-Gonzalez, A. (2021) Two new species of Stenostygnus from South America (Opiliones: Biantidae: Stenostygninae). Zootaxa, 4984 (1), 182 - 202. https: // doi. org / 10.11646 / zootaxa. 4984.1.15" type="journal article" year="2021">
Mamani
<emphasis id="B9062169FFDEFFBE2721F9E4FC3D66CF" box="[845,904,1693,1719]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2021
</bibRefCitation>
, figs. 2A, E, 8A, E); and male sexually dimorphic traits of swollen chelicerae and enlarged metatarsus III are not observed in
<taxonomicName id="4C7286F8FFDEFFBE200BF9B8FB4C66A4" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[1127,1273,1730,1756]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE200BF9B8FB4C66A4" box="[1127,1273,1730,1756]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
(
<emphasis id="B9062169FFDEFFBE2167F9BBFA9866A3" box="[1291,1325,1729,1755]" italics="true" pageId="2" pageNumber="403">cf.,</emphasis>
<figureCitation id="1349E1FEFFDEFFBE2159F9BBFA2266A3" box="[1333,1431,1729,1755]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="2" pageNumber="403">Figs. 1B</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE24FBF99FFF0E6687" box="[151,187,1765,1791]" captionStart="FIGURE 3" captionStartId="6.[151,250,890,914]" captionTargetBox="[151,1435,181,866]" captionTargetId="figure-19@6.[151,1435,181,866]" captionTargetPageId="6" captionText="FIGURE 3. Glandular structure of male metatarsus III in Caribbiantes cubanus. A. Light microscope photo, male (CZACC 3.4372), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E., Scanning electron micrographs, male (CZACC 3.3266): B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note its depth; D. Closer zoom into the groove, white arrow indicates a seta (long paint brush, basally striated and twisted), small arrowheads point to glandular openings (pores) irregularly arranged around the setae; E. Detail of the integument of the ventral groove." figureDoi="http://doi.org/10.5281/zenodo.13914733" httpUri="https://zenodo.org/record/13914733/files/figure.png" pageId="2" pageNumber="403">3A</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE24AAF99FFF5C6687" box="[198,233,1765,1791]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="2" pageNumber="403">6C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE249FF99CFEAD6778" box="[243,280,1766,1792]" captionStart="FIGURE 7" captionStartId="14.[151,250,1774,1798]" captionTargetBox="[151,1435,1065,1750]" captionTargetId="figure-443@14.[151,1435,1065,1750]" captionTargetPageId="14" captionText="FIGURE 7. Glandular structure of male metatarsus III in Caribbiantes obtusus sp. nov. (male CZACC 3.4379). A. Light microscope photo, general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs: B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, arrows show setae (long paint brush, basally striated and twisted); E. Integument of the ventral groove, white arrowheads indicate glandular openings (pores) irregularly arranged around the setae." figureDoi="http://doi.org/10.5281/zenodo.13914743" httpUri="https://zenodo.org/record/13914743/files/figure.png" pageId="2" pageNumber="403">7A</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE254EF99FFEF06687" box="[290,325,1765,1791]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="2" pageNumber="403">9C</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2523F99FFECB6687" box="[335,382,1765,1791]" captionStart="FIGURE 10" captionStartId="18.[151,250,890,914]" captionTargetBox="[151,1435,181,866]" captionTargetId="figure-19@18.[151,1435,181,866]" captionTargetPageId="18" captionText="FIGURE 10. Glandular structure of male metatarsus III in Caribbiantes barbai sp. nov. A. Light microscope photo, holotype male (CZACC 3. 4380), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E. Scanning electron micrographs, male (CZACC 3.3392); B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note the depth of the groove; D. Closer zoom into the groove, white arrow shows a seta (long paint brush, basally striated and twisted), small arrowheads indicate glandular openings (pores) regularly arranged around the setae; E. Detail of the integument and pores (arrowheads) of the ventral groove." figureDoi="http://doi.org/10.5281/zenodo.13914749" httpUri="https://zenodo.org/record/13914749/files/figure.png" pageId="2" pageNumber="403">10A</figureCitation>
and
<bibRefCitation id="EFE3808AFFDEFFBE25DEF99FFD396687" author="Mamani, C. V. &amp; Porto, W. &amp; Iglesias, P. P. &amp; Perez-Gonzalez, A." box="[434,652,1765,1791]" pageId="2" pageNumber="403" pagination="182 - 202" refId="ref17030" refString="Mamani, C. V., Porto, W., Iglesias, P. P. &amp; Perez-Gonzalez, A. (2021) Two new species of Stenostygnus from South America (Opiliones: Biantidae: Stenostygninae). Zootaxa, 4984 (1), 182 - 202. https: // doi. org / 10.11646 / zootaxa. 4984.1.15" type="journal article" year="2021">
Mamani
<emphasis id="B9062169FFDEFFBE2679F99CFDF96687" box="[533,588,1765,1791]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2021
</bibRefCitation>
, figs. 6C, 12C, E). Furthermore,
<taxonomicName id="4C7286F8FFDEFFBE2798F99FFB336687" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[1012,1158,1765,1791]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE2798F99FFB336687" box="[1012,1158,1765,1791]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
has
<emphasis id="B9062169FFDEFFBE20DBF99CFAF56687" box="[1207,1344,1765,1791]" italics="true" pageId="2" pageNumber="403">pars distalis</emphasis>
swollen and convex,
<emphasis id="B9062169FFDEFFBE2546F870FEC7675B" box="[298,370,1802,1827]" italics="true" pageId="2" pageNumber="403">versus</emphasis>
a
<emphasis id="B9062169FFDEFFBE25FCF870FDA9675B" box="[400,540,1801,1827]" italics="true" pageId="2" pageNumber="403">pars distalis</emphasis>
swollen but ending in a
<emphasis id="B9062169FFDEFFBE2750F873FC59675B" box="[828,1004,1801,1827]" italics="true" pageId="2" pageNumber="403">lamina apicalis</emphasis>
medially divided in
<taxonomicName id="4C7286F8FFDEFFBE20B0F870FADB675C" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[1244,1390,1802,1828]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE20B0F870FADB675C" box="[1244,1390,1802,1828]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
(
<emphasis id="B9062169FFDEFFBE21EDF873FA28675B" box="[1409,1437,1801,1827]" italics="true" pageId="2" pageNumber="403">cf.</emphasis>
<figureCitation id="1349E1FEFFDEFFBE24FBF857FEA9673F" box="[151,284,1837,1864]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5C, G</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2544F857FEDA673F" box="[296,367,1837,1863]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8C, G</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2517F857FE7E673F" box="[379,459,1837,1863]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11C, G</figureCitation>
and
<bibRefCitation id="EFE3808AFFDEFFBE266FF857FD56673F" author="Mamani, C. V. &amp; Porto, W. &amp; Iglesias, P. P. &amp; Perez-Gonzalez, A." box="[515,739,1837,1863]" pageId="2" pageNumber="403" pagination="182 - 202" refId="ref17030" refString="Mamani, C. V., Porto, W., Iglesias, P. P. &amp; Perez-Gonzalez, A. (2021) Two new species of Stenostygnus from South America (Opiliones: Biantidae: Stenostygninae). Zootaxa, 4984 (1), 182 - 202. https: // doi. org / 10.11646 / zootaxa. 4984.1.15" type="journal article" year="2021">
Mamani
<emphasis id="B9062169FFDEFFBE2604F854FD14673F" box="[616,673,1837,1863]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2021
</bibRefCitation>
, figs. 5A, 11A;
<bibRefCitation id="EFE3808AFFDEFFBE27FBF857FB3C6730" author="Pinto-da-Rocha, R." box="[919,1161,1837,1864]" pageId="2" pageNumber="403" pagination="194 - 198" refId="ref17095" refString="Pinto-da-Rocha, R. (1995) Redescription of Stenostygnus pusio Simon and synonymy of Caribbiantinae with Stenostygninae (Opiliones: Laniatores, Biantidae). Journal of Arachnology, 23, 194 - 198." type="journal article" year="1995">Pinto-da-Rocha 1995</bibRefCitation>
, fig. 5);
<emphasis id="B9062169FFDEFFBE2085F857FA29673F" box="[1257,1436,1837,1863]" italics="true" pageId="2" pageNumber="403">capsula externa</emphasis>
is modified into a rigid
<emphasis id="B9062169FFDEFFBE25C3F82BFD946713" box="[431,545,1873,1899]" italics="true" pageId="2" pageNumber="403">stragulum</emphasis>
, that does not fully invert its position when the penis is everted,
<emphasis id="B9062169FFDEFFBE214CF828FADD6713" box="[1312,1384,1874,1899]" italics="true" pageId="2" pageNumber="403">versus</emphasis>
two
<emphasis id="B9062169FFDEFFBE24FBF80FFF4A67F7" box="[151,255,1909,1935]" italics="true" pageId="2" pageNumber="403">titillators</emphasis>
with apical borders dorsally projected in
<taxonomicName id="4C7286F8FFDEFFBE26BAF80CFCDD67E8" ID-CoL="7MWT" authorityName="Simon" authorityYear="1879" box="[726,872,1910,1936]" class="Arachnida" family="Biantidae" genus="Stenostygnus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE26BAF80CFCDD67E8" box="[726,872,1910,1936]" italics="true" pageId="2" pageNumber="403">Stenostygnus</emphasis>
</taxonomicName>
(
<emphasis id="B9062169FFDEFFBE2716F80FFC2367F7" box="[890,918,1909,1935]" italics="true" pageId="2" pageNumber="403">cf.</emphasis>
<figureCitation id="1349E1FEFFDEFFBE27F0F80FFB9667E8" box="[924,1059,1909,1936]" captionStart="FIGURE 5" captionStartId="10.[151,250,1441,1465]" captionTargetBox="[190,1407,181,1416]" captionTargetId="figure-19@10.[180,1407,181,1416]" captionTargetPageId="10" captionText="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="2" pageNumber="403">Figs. 5G, H</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2043F80FFBDE67E8" box="[1071,1131,1909,1936]" captionStart="FIGURE 8" captionStartId="15.[151,250,1835,1859]" captionTargetBox="[197,1401,189,1812]" captionTargetId="figure-18@15.[186,1401,181,1812]" captionTargetPageId="15" captionText="FIGURE 8. Caribbiantes obtusus sp. nov., entire (C) and distal part (A–B, D–J) of penis. A–F: Drawings with optical microscope, male holotype (CZACC 3.3384), A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron microscopy male CZACC 3.4379). G. Dorsal view; H. Apical view (apical portion of a conductor colored in pink); I. Ventrolateral view; J. Ventral view. Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914745" httpUri="https://zenodo.org/record/13914745/files/figure.png" pageId="2" pageNumber="403">8G, I</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2014F80FFB7467E8" box="[1144,1217,1909,1936]" captionStart="FIGURE 11" captionStartId="21.[151,250,1761,1785]" captionTargetBox="[163,1432,191,1737]" captionTargetId="figure-18@21.[155,1432,181,1737]" captionTargetPageId="21" captionText="FIGURE 11. Caribbiantes barbai sp. nov., entire (C) anddistal part (A–B, D–J) of penis,A–F.Drawings with optical microscope, male holotype (CZACC 3.4380): A. Dorsal view; B. Dorsal view, expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view, expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm. G–J. Scanning electron micrographs (male CZACC 3.3392): G. Dorsal view, expanded; H. Apical view, expanded; I. Left lateral view, expanded; J. Ventral view, expanded.Abbreviations: AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi="http://doi.org/10.5281/zenodo.13914751" httpUri="https://zenodo.org/record/13914751/files/figure.png" pageId="2" pageNumber="403">11G, I</figureCitation>
, and
<bibRefCitation id="EFE3808AFFDEFFBE2092F80FFF6767CB" author="Mamani, C. V. &amp; Porto, W. &amp; Iglesias, P. P. &amp; Perez-Gonzalez, A." pageId="2" pageNumber="403" pagination="182 - 202" refId="ref17030" refString="Mamani, C. V., Porto, W., Iglesias, P. P. &amp; Perez-Gonzalez, A. (2021) Two new species of Stenostygnus from South America (Opiliones: Biantidae: Stenostygninae). Zootaxa, 4984 (1), 182 - 202. https: // doi. org / 10.11646 / zootaxa. 4984.1.15" type="journal article" year="2021">
Mamani
<emphasis id="B9062169FFDEFFBE2108F80CFA2867F7" box="[1380,1437,1909,1935]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2021
</bibRefCitation>
, figs. 5A, 11A;
<bibRefCitation id="EFE3808AFFDEFFBE25EBF8E3FDCF67CC" author="Pinto-da-Rocha, R." box="[391,634,1945,1972]" pageId="2" pageNumber="403" pagination="194 - 198" refId="ref17095" refString="Pinto-da-Rocha, R. (1995) Redescription of Stenostygnus pusio Simon and synonymy of Caribbiantinae with Stenostygninae (Opiliones: Laniatores, Biantidae). Journal of Arachnology, 23, 194 - 198." type="journal article" year="1995">Pinto-da-Rocha 1995</bibRefCitation>
, fig. 5).
<taxonomicName id="4C7286F8FFDEFFBE26B6F8E3FCD967CB" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[730,876,1945,1971]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE26B6F8E3FCD967CB" box="[730,876,1945,1971]" italics="true" pageId="2" pageNumber="403">Caribbiantes</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C7286F8FFDEFFBE20CDF8E3FA2967CB" ID-CoL="62G5L" authority="Silhavy, 1973" authorityName="Silhavy" authorityYear="1973" box="[1185,1436,1945,1974]" class="Arachnida" family="Biantidae" genus="Bidoma" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE20CDF8E3FB4D67CB" box="[1185,1272,1945,1971]" italics="true" pageId="2" pageNumber="403">Bidoma</emphasis>
<bibRefCitation id="EFE3808AFFDEFFBE216CF8E1FA2967CB" author="Silhavy, V." box="[1280,1436,1945,1974]" pageId="2" pageNumber="403" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý, 1973</bibRefCitation>
</taxonomicName>
by the presence of a longitudinal median groove in area I (
<emphasis id="B9062169FFDEFFBE2748F8C4FCD967AF" box="[804,876,1982,2007]" italics="true" pageId="2" pageNumber="403">versus</emphasis>
area I undivided in
<taxonomicName id="4C7286F8FFDEFFBE2027F8C7FB1767AF" ID-CoL="62G5L" authorityName="Silhavy" authorityYear="1973" box="[1099,1186,1981,2007]" class="Arachnida" family="Biantidae" genus="Bidoma" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDEFFBE2027F8C7FB1767AF" box="[1099,1186,1981,2007]" italics="true" pageId="2" pageNumber="403">Bidoma</emphasis>
</taxonomicName>
) (
<emphasis id="B9062169FFDEFFBE20D0F8C7FB6D67AF" box="[1212,1240,1981,2007]" italics="true" pageId="2" pageNumber="403">cf.</emphasis>
<figureCitation id="1349E1FEFFDEFFBE20B2F8C7FAF567AF" box="[1246,1344,1981,2007]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="2" pageNumber="403">Figs. 1A</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2120F8C7FAC467AF" box="[1356,1393,1981,2007]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="2" pageNumber="403">6A</figureCitation>
,
<figureCitation id="1349E1FEFFDEFFBE2110F8C7FA2967AF" box="[1404,1436,1981,2007]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="2" pageNumber="403">9A</figureCitation>
and
<figureCitation id="1349E1FEFFDFFFBF24AAFFEDFF4F60C9" box="[198,250,151,177]" captionStart="FIGURE 14" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,181,1128]" captionTargetId="figure-18@25.[151,1435,181,1128]" captionTargetPageId="25" captionText="FIGURE 14. Antillean stenostygnines, external morphology: dorsal habitus. A. Caribbiantes cubanus, male (CZACC 3.3380); B. Manahunca bielawskii, male (CZACC 3.4352, neotype); C. Heterolacurbs ovalis, male (FSCA); D. Negreaella rioindiocubanicola, male (CZACC 3.4376); E. Galibrotus carlotanus, male (CZACC 3.3394); F. Decuella cf. cubaorientalis male (CZACC 3.3398); G. Bidoma indivisum male (MCZ IZ-14741, holotype); H. Vestitecola haitensis Šilhavý, 1973, female (MCZ IZ-14736, holotype). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914757" httpUri="https://zenodo.org/record/13914757/files/figure.png" pageId="3" pageNumber="404">14G</figureCitation>
). It also differs from
<taxonomicName id="4C7286F8FFDFFFBF2588FFEDFD8E60C9" ID-CoL="62G5L" authorityName="Silhavy" authorityYear="1973" box="[484,571,151,177]" class="Arachnida" family="Biantidae" genus="Bidoma" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF2588FFEDFD8E60C9" box="[484,571,151,177]" italics="true" pageId="3" pageNumber="404">Bidoma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7286F8FFDFFFBF261DFFEDFC3060C9" ID-CoL="63DZN" authority="Avram, 1977" authorityName="Avram" authorityYear="1977" box="[625,901,151,177]" class="Arachnida" family="Biantidae" genus="Negreaella" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF261DFFEDFD5960C9" box="[625,748,151,177]" italics="true" pageId="3" pageNumber="404">Negreaella</emphasis>
<bibRefCitation id="EFE3808AFFDFFFBF269DFFEDFC3060C9" author="Avram, S." box="[753,901,151,177]" pageId="3" pageNumber="404" pagination="123 - 136" refId="ref16820" refString="Avram, S. (1977) Recherches sur les opilionides de Cuba. III. Genres et especes nouveaux de Caribbiantinae (Biantidae: Gonyleptomorphi). Resultats des Expeditions Biospeologiques. Cubano-Roumaines a Cuba, 2, 123 - 136." type="journal article" year="1977">Avram, 1977</bibRefCitation>
</taxonomicName>
, by the absence of a tubercle on the eye mounds next to one opposite tubercle located on the anterior margin of area I. In addition,
<taxonomicName id="4C7286F8FFDFFFBF2032FFC1FB6C60AD" ID-CoL="63DZN" authorityName="Avram" authorityYear="1977" box="[1118,1241,187,213]" class="Arachnida" family="Biantidae" genus="Negreaella" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF2032FFC1FB6C60AD" box="[1118,1241,187,213]" italics="true" pageId="3" pageNumber="404">Negreaella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7286F8FFDFFFBF217AFFC1FA2960AD" ID-CoL="5KSD" authorityName="Silhavy" authorityYear="1973" box="[1302,1436,187,213]" class="Arachnida" family="Biantidae" genus="Manahunca" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF217AFFC1FA2960AD" box="[1302,1436,187,213]" italics="true" pageId="3" pageNumber="404">Manahunca</emphasis>
</taxonomicName>
are distinguishable from
<taxonomicName id="4C7286F8FFDFFFBF25D4FFA5FDFF6081" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[440,586,223,249]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF25D4FFA5FDFF6081" box="[440,586,223,249]" italics="true" pageId="3" pageNumber="404">Caribbiantes</emphasis>
</taxonomicName>
by the absence of a strong armature on mesotergal areas; they only bear some granules or tubercles, respectively (
<figureCitation id="1349E1FEFFDFFFBF2607FE79FD786165" box="[619,717,259,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1783,1807]" captionTargetBox="[161,1436,185,1749]" captionTargetId="figure-19@4.[151,1436,181,1759]" captionTargetPageId="4" captionText="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="3" pageNumber="404">Figs. 1B</figureCitation>
,
<figureCitation id="1349E1FEFFDFFFBF26B6FE79FD486165" box="[730,765,259,285]" captionStart="FIGURE 6" captionStartId="13.[151,250,1598,1622]" captionTargetBox="[161,1436,181,1564]" captionTargetId="figure-18@13.[151,1436,181,1574]" captionTargetPageId="13" captionText="FIGURE 6. Caribbiantes obtusus sp. nov., habitus of male holotype (CZACC 3.3384): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914739" httpUri="https://zenodo.org/record/13914739/files/figure.png" pageId="3" pageNumber="404">6C</figureCitation>
,
<figureCitation id="1349E1FEFFDFFFBF2765FE79FC986165" box="[777,813,259,285]" captionStart="FIGURE 9" captionStartId="17.[151,250,1757,1781]" captionTargetBox="[158,1436,355,1724]" captionTargetId="figure-89@17.[151,1436,345,1734]" captionTargetPageId="17" captionText="FIGURE 9. Caribbiantes barbai sp. nov., habitus of male holotype (CZACC 3.4380): A–B. Dorsal view; C–D. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914747" httpUri="https://zenodo.org/record/13914747/files/figure.png" pageId="3" pageNumber="404">9C</figureCitation>
;
<figureCitation id="1349E1FEFFDFFFBF2755FE79FC386165" box="[825,909,259,285]" captionStart="FIGURE 14" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,181,1128]" captionTargetId="figure-18@25.[151,1435,181,1128]" captionTargetPageId="25" captionText="FIGURE 14. Antillean stenostygnines, external morphology: dorsal habitus. A. Caribbiantes cubanus, male (CZACC 3.3380); B. Manahunca bielawskii, male (CZACC 3.4352, neotype); C. Heterolacurbs ovalis, male (FSCA); D. Negreaella rioindiocubanicola, male (CZACC 3.4376); E. Galibrotus carlotanus, male (CZACC 3.3394); F. Decuella cf. cubaorientalis male (CZACC 3.3398); G. Bidoma indivisum male (MCZ IZ-14741, holotype); H. Vestitecola haitensis Šilhavý, 1973, female (MCZ IZ-14736, holotype). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914757" httpUri="https://zenodo.org/record/13914757/files/figure.png" pageId="3" pageNumber="404">14D, B</figureCitation>
).
<taxonomicName id="4C7286F8FFDFFFBF27CFFE79FB806165" ID-CoL="3HHR" authorityName="Silhavy" authorityYear="1973" box="[931,1077,259,285]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF27CFFE79FB806165" box="[931,1077,259,285]" italics="true" pageId="3" pageNumber="404">Caribbiantes</emphasis>
</taxonomicName>
presents two pairs of spiniform apophyses with almost the same length on area III and IV, unlike
<taxonomicName id="4C7286F8FFDFFFBF271CFE5DFB3F6139" ID-CoL="4LW3" authority="Silhavy, 1973" authorityName="Silhavy" authorityYear="1973" box="[880,1162,295,323]" class="Arachnida" family="Biantidae" genus="Galibrotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF271CFE5DFC5D6139" box="[880,1000,295,321]" italics="true" pageId="3" pageNumber="404">Galibrotus</emphasis>
<bibRefCitation id="EFE3808AFFDFFFBF2783FE52FB3F6139" author="Silhavy, V." box="[1007,1162,295,323]" pageId="3" pageNumber="404" pagination="110 - 143" refId="ref17300" refString="Silhavy, V. (1973) Two new systematic groups of gonyleptomorphid phalangids from the Antillean-Caribbean Region. Agoristenidae fam. n. and Caribbiantinae subfam. n. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 37 (2), 110 - 143." type="journal article" year="1973">Šilhavý, 1973</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7286F8FFDFFFBF20F4FE5DFA2D6139" ID-CoL="42F7" authority="Avram, 1977" authorityName="Avram" authorityYear="1977" box="[1176,1432,295,321]" class="Arachnida" family="Biantidae" genus="Decuella" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF20F4FE5DFB486139" box="[1176,1277,295,321]" italics="true" pageId="3" pageNumber="404">Decuella</emphasis>
<bibRefCitation id="EFE3808AFFDFFFBF2168FE5DFA2D6139" author="Avram, S." box="[1284,1432,295,321]" pageId="3" pageNumber="404" pagination="123 - 136" refId="ref16820" refString="Avram, S. (1977) Recherches sur les opilionides de Cuba. III. Genres et especes nouveaux de Caribbiantinae (Biantidae: Gonyleptomorphi). Resultats des Expeditions Biospeologiques. Cubano-Roumaines a Cuba, 2, 123 - 136." type="journal article" year="1977">Avram, 1977</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C7286F8FFDFFFBF24ABFE31FDBA611D" ID-CoL="4WGT" authority="Roewer, 1912" authorityName="Roewer" authorityYear="1912" box="[199,527,331,357]" class="Arachnida" family="Biantidae" genus="Heterolacurbs" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF24ABFE31FEDD611D" box="[199,360,331,357]" italics="true" pageId="3" pageNumber="404">Heterolacurbs</emphasis>
<bibRefCitation id="EFE3808AFFDFFFBF2503FE31FDBA611D" author="Roewer, C. F." box="[367,527,331,357]" pageId="3" pageNumber="404" pagination="1 - 242" refId="ref17247" refString="Roewer, C. F. (1912) Die Familien der Assamiden und Phalangodiden der Opiliones-Laniatores. (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv fur Naturgeschichte, 78 A (3), 1 - 242." type="journal article" year="1912">Roewer, 1912</bibRefCitation>
</taxonomicName>
, which have small tubercles on area III and a pair of strong spiniform apophyses on area IV; differentially,
<taxonomicName id="4C7286F8FFDFFFBF25D8FE15FDE061F1" ID-CoL="4WGT" authorityName="Roewer" authorityYear="1912" box="[436,597,367,393]" class="Arachnida" family="Biantidae" genus="Heterolacurbs" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B9062169FFDFFFBF25D8FE15FDE061F1" box="[436,597,367,393]" italics="true" pageId="3" pageNumber="404">Heterolacurbs</emphasis>
</taxonomicName>
possesses a single spiniform apophysis in free tergite III (
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<paragraph id="8BCDFD7BFFDFFFBF24FBFD11FEA762FD" blockId="3.[151,954,546,645]" box="[151,274,618,645]" pageId="3" pageNumber="404">
(
<figureCitation id="1349E1FEFFDFFFBF24F3FD11FEBC62FD" box="[159,265,618,645]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,250,1783,1807]" captionStartId-1="5.[151,250,1943,1967]" captionStartId-2="6.[151,250,890,914]" captionStartId-3="8.[151,250,1590,1614]" captionStartId-4="10.[151,250,1441,1465]" captionTargetBox-0="[161,1436,185,1749]" captionTargetBox-1="[236,1352,193,1907]" captionTargetBox-2="[151,1435,181,866]" captionTargetBox-3="[151,1436,309,1567]" captionTargetBox-4="[190,1407,181,1416]" captionTargetId-0="figure-19@4.[151,1436,181,1759]" captionTargetId-1="figure-18@5.[224,1364,181,1919]" captionTargetId-2="figure-19@6.[151,1435,181,866]" captionTargetId-3="figure-89@8.[151,1436,309,1567]" captionTargetId-4="figure-19@10.[180,1407,181,1416]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="10" captionText-0="FIGURE 1. Caribbiantes cubanus, habitus: A–B, holotype, α male (MCZ IZ-14702). A. dorsal view; B. lateral view; C–D, β male (CZACC 3.3380). C. dorsal view; D. lateral view. Scale bars = 1 mm." captionText-1="FIGURE 2. Caribbiantes cubanus, male holotype (MCZ IZ-14702), right pedipalp (trochanter to tarsus): A. pedipalp mesal view; B. pedipalp ectal view. Abbreviations: Tr, trochanter; Fe, femur; Pa, patella; Ti, tibia; Ta, tarsus; TC, tarsal claw; S, spine; edS, ectodistal spines; g, gap in the tubercles series. Scale bars = 1 mm." captionText-2="FIGURE 3. Glandular structure of male metatarsus III in Caribbiantes cubanus. A. Light microscope photo, male (CZACC 3.4372), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E., Scanning electron micrographs, male (CZACC 3.3266): B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note its depth; D. Closer zoom into the groove, white arrow indicates a seta (long paint brush, basally striated and twisted), small arrowheads point to glandular openings (pores) irregularly arranged around the setae; E. Detail of the integument of the ventral groove." captionText-3="FIGURE 4. Caribbiantes cubanus, male (CZACC 3. 4372), tarsus III, ventral view of the two tarsomeres of the distitarsus and claws, showing the bipartite scopula. Abbreviations: Cl, claw; Sc, scopula. Scale bars = 1 mm." captionText-4="FIGURE 5. Caribbiantes cubanus, entire (C) and distal part (A–B, D–I) of penis,A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view.Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae." figureDoi-0="http://doi.org/10.5281/zenodo.13914729" figureDoi-1="http://doi.org/10.5281/zenodo.13914731" figureDoi-2="http://doi.org/10.5281/zenodo.13914733" figureDoi-3="http://doi.org/10.5281/zenodo.13914735" figureDoi-4="http://doi.org/10.5281/zenodo.13914737" httpUri-0="https://zenodo.org/record/13914729/files/figure.png" httpUri-1="https://zenodo.org/record/13914731/files/figure.png" httpUri-2="https://zenodo.org/record/13914733/files/figure.png" httpUri-3="https://zenodo.org/record/13914735/files/figure.png" httpUri-4="https://zenodo.org/record/13914737/files/figure.png" pageId="3" pageNumber="404">Figs. 1–5</figureCitation>
@ -91,28 +95,35 @@ urn:lsid:zoobank.org:act:
<paragraph id="8BCDFD7BFFDFFFBF24FBFD8FFF7C632F" blockId="3.[151,1437,757,1936]" pageId="3" pageNumber="404">
<emphasis id="B9062169FFDFFFBF24FBFD8FFEA86377" bold="true" box="[151,285,757,783]" pageId="3" pageNumber="404">Type series</emphasis>
.
<materialsCitation id="3B1AF726FFDFFFBF2541FD8FFF7C632F" collectingDate="1936-07-01" collectingDateMax="1936-07-02" collectingDateMin="1936-07-01" collectionCode="MCZ" collectorName="Darlington" country="Cuba" location="Soledad" pageId="3" pageNumber="404" specimenCode="IZ-14702, IZ-38833" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Cienfuegos" typeStatus="holotype">
<materialsCitation id="3B1AF726FFDFFFBF2541FD8FFC836377" box="[301,822,757,784]" collectingDate="1936-07-01" collectingDateMax="1936-07-02" collectingDateMin="1936-07-01" collectionCode="MCZ" collectorName="Darlington" country="Cuba" location="Soledad" pageId="3" pageNumber="404" specimenCode="IZ-14702" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cienfuegos" typeStatus="holotype">
<typeStatus id="54C943D9FFDFFFBF2541FD8FFE206377" box="[301,405,757,783]" pageId="3" pageNumber="404" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7436F2FFDFFFBF25F1FD8CFE076368" box="[413,434,758,784]" count="1" pageId="3" pageNumber="404" type="male">♂</specimenCount>
(α) (
<collectionCode id="ED6365BEFFDFFFBF2580FD8FFD9E6377" box="[492,555,757,783]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="404" type="Museum">MCZ</collectionCode>
<collectionCode id="ED6365BEFFDFFFBF2580FD8FFD9E6377" box="[492,555,757,783]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="404" type="Museum">MCZ</collectionCode>
<specimenCode id="DBD45500FFDFFFBF265FFD8FFD2B6377" box="[563,670,757,783]" collectionCode="IZ-" pageId="3" pageNumber="404">IZ-14702</specimenCode>
) (examined);
) (examined);
</materialsCitation>
<materialsCitation id="3B1AF726FFDFFFBF2752FD8CFA8D6377" box="[830,1336,757,784]" collectingDate="1936-07-01" collectingDateMax="1936-07-02" collectingDateMin="1936-07-01" collectionCode="MCZ" collectorName="Darlington" country="Cuba" location="Soledad" pageId="3" pageNumber="404" specimenCode="IZ-38833" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Cienfuegos" typeStatus="paratype">
<typeStatus id="54C943D9FFDFFFBF2752FD8CFC1B6368" box="[830,942,758,784]" pageId="3" pageNumber="404" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D7436F2FFDFFFBF27D6FD8FFC506368" box="[954,997,757,784]" count="1" pageId="3" pageNumber="404" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED6365BEFFDFFFBF2799FD8FFB816377" box="[1013,1076,757,783]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="404" type="Museum">MCZ</collectionCode>
<collectionCode id="ED6365BEFFDFFFBF2799FD8FFB816377" box="[1013,1076,757,783]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="404" type="Museum">MCZ</collectionCode>
<specimenCode id="DBD45500FFDFFFBF2050FD8FFB126377" box="[1084,1191,757,783]" collectionCode="IZ-" pageId="3" pageNumber="404">IZ-38833</specimenCode>
) (examined) and
) (examined)
</materialsCitation>
<materialsCitation id="3B1AF726FFDFFFBF212CFD8FFF7C632F" collectingDate="1936-07-01" collectingDateMax="1936-07-02" collectingDateMin="1936-07-01" collectionCode="MCZ" collectorName="Darlington" country="Cuba" location="Soledad" pageId="3" pageNumber="404" specimenCode="IZ-14702, IZ-38833" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cienfuegos Province" typeStatus="holotype">
and
<specimenCount id="9D7436F2FFDFFFBF211DFD8FFA296368" box="[1393,1436,757,784]" count="1" pageId="3" pageNumber="404" type="male">1 ♂</specimenCount>
(not examined; see remarks).
<collectingCountry id="F365BDEBFFDFFFBF258EFC63FD95634B" box="[482,544,793,819]" name="Cuba" pageId="3" pageNumber="404">Cuba</collectingCountry>
,
<collectingRegion id="49B63399FFDFFFBF2647FC63FCAD634B" box="[555,792,793,819]" country="Cuba" name="Cienfuegos" pageId="3" pageNumber="404">Cienfuegos Province</collectingRegion>
,
<location id="8EADABA0FFDFFFBF274FFC63FCC8634B" LSID="urn:lsid:plazi:treatment:03DB4C6DFFDFFFB624FBFD58FA3E6711:8EADABA0FFDFFFBF274FFC63FCC8634B" box="[803,893,793,819]" country="Cuba" name="Soledad" pageId="3" pageNumber="404" stateProvince="Cienfuegos">Soledad</location>
(currently Pepito Tey),
<location id="8EADABA0FFDFFFBF274FFC63FCC8634B" LSID="urn:lsid:plazi:treatment:03DB4C6DFFDFFFB624FBFD58FA3E6711:8EADABA0FFDFFFBF274FFC63FCC8634B" box="[803,893,793,819]" country="Cuba" name="Soledad" pageId="3" pageNumber="404" stateProvince="Cienfuegos Province">Soledad</location>
(currently
<location id="8EADABA0FFDFFFBF2795FC63FBC0634B" LSID="urn:lsid:plazi:treatment:03DB4C6DFFDFFFB624FBFD58FA3E6711:8EADABA0FFDFFFBF2795FC63FBC0634B" box="[1017,1141,793,819]" country="Cuba" name="Pepito Tey" pageId="3" pageNumber="404" stateProvince="Cienfuegos Province">Pepito Tey</location>
),
<date id="FFCCDBBBFFDFFFBF20E4FC63FAA2634B" box="[1160,1303,793,819]" pageId="3" pageNumber="404" value="1936-07-01" valueMax="1936-07-02" valueMin="1936-07-01">
<collectingDate id="EF882253FFDFFFBF20E4FC63FAA2634B" box="[1160,1303,793,819]" pageId="3" pageNumber="404" value="1936-07-01" valueMax="1936-07-02" valueMin="1936-07-01">1–2.vii.1936</collectingDate>
</date>
@ -122,7 +133,7 @@ coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFDFFFBA24ABFC1BFDE765D7" lastPageId="6" lastPageNumber="407" pageId="3" pageNumber="404" type="discussion">
<subSubSection id="C368AEF0FFDFFFBF24ABFC1BFE826457" pageId="3" pageNumber="404" type="discussion">
<paragraph id="8BCDFD7BFFDFFFBF24ABFC1BFE826457" blockId="3.[151,1437,757,1936]" pageId="3" pageNumber="404">
<emphasis id="B9062169FFDFFFBF24ABFC1BFE8E6303" bold="true" box="[199,315,865,891]" pageId="3" pageNumber="404">Remarks.</emphasis>
The
@ -133,6 +144,8 @@ The
<typeStatus id="54C943D9FFDFFFBF26D1FC8BFC976473" box="[701,802,1009,1035]" pageId="3" pageNumber="404" type="holotype">holotype</typeStatus>
, mentioned in the original description, was not found in the collection.
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFDFFFBF24ABFB43FDCC67F7" pageId="3" pageNumber="404" type="materials_examined">
<paragraph id="8BCDFD7BFFDFFFBF24ABFB43FDCC67F7" blockId="3.[151,1437,757,1936]" pageId="3" pageNumber="404">
<emphasis id="B9062169FFDFFFBF24ABFB43FE46642B" bold="true" box="[199,499,1081,1107]" pageId="3" pageNumber="404">Other material examined</emphasis>
.
@ -329,6 +342,7 @@ and
<emphasis id="B9062169FFDFFFBF258EF80FFDDC67F7" bold="true" box="[482,617,1909,1935]" pageId="3" pageNumber="404">new record</emphasis>
).
</paragraph>
</subSubSection>
<caption id="DF0DADF3FFD8FFB824FBF98DFC27674B" ID-DOI="http://doi.org/10.5281/zenodo.13914729" ID-Zenodo-Dep="13914729" httpUri="https://zenodo.org/record/13914729/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1783,1807]" targetBox="[161,1436,185,1749]" targetPageId="4" targetType="figure">
<paragraph id="8BCDFD7BFFD8FFB824FBF98DFC27674B" blockId="4.[151,1436,1783,1843]" pageId="4" pageNumber="405">
<emphasis id="B9062169FFD8FFB824FBF98DFEA16777" bold="true" box="[151,276,1783,1807]" pageId="4" pageNumber="405">FIGURE 1.</emphasis>
@ -359,6 +373,7 @@ Glandular structure of male metatarsus III in
A. Light microscope photo, male (CZACC 3.4372), general view of the whole metatarsus III (divided into four quarters to facilitate comparisons); B–E., Scanning electron micrographs, male (CZACC 3.3266): B. View of the glandular swelling, dashed line delineates the form of the ventral, buttonhole-like groove; C. Close-up of the groove delimited in B, note its depth; D. Closer zoom into the groove, white arrow indicates a seta (long paint brush, basally striated and twisted), small arrowheads point to glandular openings (pores) irregularly arranged around the setae; E. Detail of the integument of the ventral groove.
</paragraph>
</caption>
<subSubSection id="C368AEF0FFDAFFBA24ABFBE3FDE765D7" pageId="6" pageNumber="407" type="diagnosis">
<paragraph id="8BCDFD7BFFDAFFBA24ABFBE3FDE765D7" blockId="6.[151,1437,1176,1563]" pageId="6" pageNumber="407">
<emphasis id="B9062169FFDAFFBA24ABFBE3FE0064CA" bold="true" box="[199,437,1176,1203]" pageId="6" pageNumber="407">Emended diagnosis.</emphasis>
<taxonomicName id="4C7286F8FFDAFFBA25D7FBE3FD0464CB" authorityName="Silhavy" authorityYear="1973" box="[443,689,1177,1203]" class="Arachnida" family="Biantidae" genus="Caribbiantes" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="cubanus">
@ -405,10 +420,10 @@ and
<paragraph id="8BCDFD7BFFDAFFBA24ABF97BFE676663" blockId="6.[151,1437,1176,1563]" box="[199,466,1537,1563]" pageId="6" pageNumber="407">
<emphasis id="B9062169FFDAFFBA24ABF97BFEDC6662" box="[199,361,1537,1562]" italics="true" pageId="6" pageNumber="407">Measurements</emphasis>
:
<tableCitation id="C6F0C8C0FFDAFFBA2514F97BFE796663" box="[376,460,1537,1563]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." pageId="6" pageNumber="407">Table 1</tableCitation>
<tableCitation id="C6F0C8C0FFDAFFBA2514F97BFE796663" box="[376,460,1537,1563]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." httpUri="http://table.plazi.org/id/DF0DADF3FFDAFFBA24F4F930FE3C66FE" pageId="6" pageNumber="407" tableUuid="DF0DADF3FFDAFFBA24F4F930FE3C66FE">Table 1</tableCitation>
.
</paragraph>
<caption id="DF0DADF3FFDAFFBA24F4F930FE3C66FE" pageId="6" pageNumber="407" startId="6.[152,244,1610,1634]" targetType="table">
<caption id="DF0DADF3FFDAFFBA24F4F930FE3C66FE" ID-Table-UUID="DF0DADF3FFDAFFBA24F4F930FE3C66FE" httpUri="http://table.plazi.org/id/DF0DADF3FFDAFFBA24F4F930FE3C66FE" pageId="6" pageNumber="407" startId="6.[152,244,1610,1634]" targetBox="[163,1402,1687,1971]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BCDFD7BFFDAFFBA24F4F930FE3C66FE" blockId="6.[151,1436,1610,1670]" pageId="6" pageNumber="407">
<emphasis id="B9062169FFDAFFBA24F4F930FEBD661A" bold="true" box="[152,264,1610,1634]" pageId="6" pageNumber="407">TABLE 1</emphasis>
. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of
@ -420,7 +435,7 @@ and
</caption>
<paragraph id="8BCDFD7BFFDAFFBA251AF9EDFAD567CB" pageId="6" pageNumber="407">
<table id="F9720FDBFFDA004324CFF9EDFACF67CB" box="[163,1402,1687,1971]" colsContinueIn="7.[163,1402,203,1655]" gridcols="9" gridrows="6" pageId="6" pageNumber="407">
<tr id="3542FF39FFDA004324CFF9EDFACF6691" box="[163,1402,1687,1769]" gridrow="0" pageId="6" pageNumber="407" rowspan-0="1">
<tr id="3542FF39FFDA004324CFF9EDFACF6691" box="[163,1402,1687,1769]" gridrow="0" isHeader="true" pageId="6" pageNumber="407" rowspan-0="1">
<th id="76939645FFDA00432504F9EDFE0B6691" box="[360,446,1687,1769]" gridcol="1" gridrow="0" pageId="6" pageNumber="407">H ♂α MCZ IZ 14702</th>
<th id="76939645FFDA00432583F9EDFDF66691" box="[495,579,1687,1769]" gridcol="2" gridrow="0" pageId="6" pageNumber="407">♂α CZACC 3.3379</th>
<th id="76939645FFDA0043261AF9EDFD7F6691" box="[630,714,1687,1769]" gridcol="3" gridrow="0" pageId="6" pageNumber="407">♂α CZACC 3.3381</th>
@ -492,7 +507,7 @@ and
</caption>
<paragraph id="8BCDFD7BFFDBFFBB251AFFB6FAD5660F" pageId="7" pageNumber="408">
<table id="F9720FDBFFDB004324CFFFB1FACF660F" box="[163,1402,203,1655]" colsContinueFrom="6.[163,1402,1687,1971]" gridcols="9" gridrows="35" pageId="7" pageNumber="408">
<tr id="3542FF39FFDB004324CFFFB1FACF6166" box="[163,1402,203,286]" gridrow="0" pageId="7" pageNumber="408" rowspan-0="1">
<tr id="3542FF39FFDB004324CFFFB1FACF6166" box="[163,1402,203,286]" gridrow="0" isHeader="true" pageId="7" pageNumber="408" rowspan-0="1">
<th id="76939645FFDB00432504FFB1FE0B6166" box="[360,446,203,286]" gridcol="1" gridrow="0" pageId="7" pageNumber="408">H ♂α MCZ IZ 14702</th>
<th id="76939645FFDB00432583FFB1FDF66166" box="[495,579,203,286]" gridcol="2" gridrow="0" pageId="7" pageNumber="408">♂α CZACC 3.3379</th>
<th id="76939645FFDB0043261AFFB1FD7F6166" box="[630,714,203,286]" gridcol="3" gridrow="0" pageId="7" pageNumber="408">♂α CZACC 3.3381</th>
@ -1024,13 +1039,13 @@ Female
<typeStatus id="54C943D9FFD5FFB52545F969FE216655" box="[297,404,1555,1581]" pageId="9" pageNumber="410" type="paratype">paratype</typeStatus>
</emphasis>
(MCZ IZ-38833). Overall habitus similar to the male, differing in mesotergal areas less granulated and metatarsus III not enlarged; unlike α males, cheliceral hands are not hypertelic; cheliceral fingers do not leave a gap between them, and the fixed finger lacks any basal tubercle. Legs III–IV with femur, patella and tibia slightly slimmer than in males. Color pattern as in males, but a little paler. Tarsal formula: 7(3):11/12(4):7:7. Measurements:
<tableCitation id="C6F0C8C0FFD5FFB52525F9D9FE2866C5" box="[329,413,1699,1725]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." pageId="9" pageNumber="410">Table 1</tableCitation>
<tableCitation id="C6F0C8C0FFD5FFB52525F9D9FE2866C5" box="[329,413,1699,1725]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." httpUri="http://table.plazi.org/id/DF0DADF3FFDAFFBA24F4F930FE3C66FE" pageId="9" pageNumber="410" tableUuid="DF0DADF3FFDAFFBA24F4F930FE3C66FE">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BCDFD7BFFD5FFB524ABF9BCFB816709" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="410">
<emphasis id="B9062169FFD5FFB524ABF9BCFEF26698" bold="true" box="[199,327,1734,1760]" pageId="9" pageNumber="410">Variability</emphasis>
. Males (n=18); females (n=25). Measurements in a subsample of selected specimens:
<tableCitation id="C6F0C8C0FFD5FFB52152F9BDFA236699" box="[1342,1430,1735,1761]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." pageId="9" pageNumber="410">Table 1</tableCitation>
<tableCitation id="C6F0C8C0FFD5FFB52152F9BDFA236699" box="[1342,1430,1735,1761]" captionStart="TABLE 1" captionStartId="6.[152,244,1610,1634]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the male holotype (H), a female paratype (P) and selected additional specimens of Caribbiantes cubanus." httpUri="http://table.plazi.org/id/DF0DADF3FFDAFFBA24F4F930FE3C66FE" pageId="9" pageNumber="410" tableUuid="DF0DADF3FFDAFFBA24F4F930FE3C66FE">Table 1</tableCitation>
. Chelicerae: fixed finger with 6 to 7 distal teeth; distal teeth on movable finger may vary from 6 to 8. Metatarsus III in males: the number of setae in the ventral groove of the swollen sector vary from 10 to 15. Tarsal formula: number of segments in leg II varies from 10 to 13. Hypertelic chelicerae observed in nine out of
<specimenCount id="9D7436F2FFD5FFB520D5F849FAAE6735" box="[1209,1307,1843,1869]" count="18" pageId="9" pageNumber="410" type="male">18 males</specimenCount>
(α males— including the
@ -1040,7 +1055,7 @@ Female
have small chelicerae (ss males).
</paragraph>
</subSubSection>
<subSubSection id="C368AEF0FFD5FFB624ABF800FA3E6711" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="distribution">
<subSubSection id="C368AEF0FFD5FFB524ABF800FADA67ED" box="[199,1391,1914,1941]" pageId="9" pageNumber="410" type="distribution">
<paragraph id="8BCDFD7BFFD5FFB524ABF800FADA67ED" blockId="9.[151,1437,151,1941]" box="[199,1391,1914,1941]" pageId="9" pageNumber="410">
<emphasis id="B9062169FFD5FFB524ABF800FED467EC" bold="true" box="[199,353,1914,1940]" pageId="9" pageNumber="410">Distribution.</emphasis>
Known only from the Cuban provinces
@ -1053,6 +1068,7 @@ and
<figureCitation id="1349E1FEFFD5FFB52179F801FAD767ED" box="[1301,1378,1915,1941]" captionStart="FIGURE 12" captionStartId="22.[151,250,1756,1780]" captionTargetBox="[164,1423,181,1732]" captionTargetId="figure-19@22.[164,1423,181,1732]" captionTargetPageId="22" captionText="FIGURE 12. Records and habitat of Caribbiantes spp. A. Map of Cuba (general and expanded sector as indicated), showing known localities of Caribbiantes cubanus (circles, type locality in red); C. obtusus sp. nov. (triangles, type locality in red) and C. barbai sp. nov. (squares, type locality in red); B. Submontane rain forest in Alejandro de Humboldt National Park, Monte Iberia, type locality of C. barbai sp. nov.; C. Mountains of Topes de Collantes, Sancti Spíritus province, record of C. cubanus and C. obtusus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13914753" httpUri="https://zenodo.org/record/13914753/files/figure.png" pageId="9" pageNumber="410">Fig.12</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0DADF3FFD6FFB624FBFADBFAD56630" ID-DOI="http://doi.org/10.5281/zenodo.13914737" ID-Zenodo-Dep="13914737" httpUri="https://zenodo.org/record/13914737/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,250,1441,1465]" targetBox="[190,1407,181,1416]" targetPageId="10" targetType="figure">
<paragraph id="8BCDFD7BFFD6FFB624FBFADBFAD56630" blockId="10.[151,1437,1441,1609]" pageId="10" pageNumber="411">
<emphasis id="B9062169FFD6FFB624FBFADBFEA765C1" bold="true" box="[151,274,1441,1465]" pageId="10" pageNumber="411">FIGURE 5.</emphasis>
@ -1062,6 +1078,7 @@ and
, entire (C) and distal part (A–B, D–I) of penis, A–F. Drawings with optical microscope (male CZACC 3.3379): A. Dorsal view; B. Dorsal view expanded; C. Ventral and lateral views (entire penis); D. Left lateral view; E. Left lateral view expanded (with enlarged detail); F. Ventral view. Scale bars = 0.1 mm; G–I. Scanning electron micrographs (male CZACC 3.3266): G. Dorsal view; H. Left lateral view; I. Ventral view. Abbreviations:AP, apical process; AS, apical setae; CI, capsula interna; Clf, cleft; Co, conductors; Fo, follis; Led, ledge; Str, stragulum; Sty, stylus; VLS, ventrolateral setae.
</paragraph>
</caption>
<subSubSection id="C368AEF0FFD6FFB624ABF9E1FA3E6711" pageId="10" pageNumber="411" type="biology_ecology">
<paragraph id="8BCDFD7BFFD6FFB624ABF9E1FA3E6711" blockId="10.[151,1436,1691,1898]" pageId="10" pageNumber="411">
<emphasis id="B9062169FFD6FFB624ABF9E1FECB66CD" bold="true" box="[199,382,1691,1717]" pageId="10" pageNumber="411">Natural history</emphasis>
. The specimens were mainly collected under stones. In some localities of the Natural Protected Landscape of Topes de Collantes, a mountainous zone in the