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<mods:title id="83823F039005C764B34CC59430BF5E4A">A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia</mods:title>
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<mods:namePart>Giachino, Pier Mauro</mods:namePart>
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<mods:namePart>Eberhard, Stefan</mods:namePart>
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<treatment ID-GBIF-Taxon="182754070" LSID="urn:lsid:zoobank.org:act:B98AC026-EEAD-43A7-B4B0-CF5121A12836" httpUri="http://treatment.plazi.org/id/B7E825B39D9B599CB651811C3E1B7D3C" lastPageNumber="269" pageId="0" pageNumber="269">
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<taxonomicName LSID="http://zoobank.org/B98AC026-EEAD-43A7-B4B0-CF5121A12836" authority="Giachino & Eberhard & Perina, 2021" authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. nov.">Kimberleytyphlus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="269">gen. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 34, 35" captionStartId="F11" captionText="Figures 34, 35. Kimberleytyphlus carrboydianus gen. et sp. nov., HT ♂ 34 habitus 35 aedeagus in lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1044.58844.figures34-35" httpUri="https://binary.pensoft.net/fig/553770" pageId="0" pageNumber="269">Figs 34-35</figureCitation>
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<treatment id="B7E825B39D9B599CB651811C3E1B7D3C" ID-GBIF-Taxon="182754070" LSID="urn:lsid:zoobank.org:act:B98AC026-EEAD-43A7-B4B0-CF5121A12836" httpUri="http://treatment.plazi.org/id/B7E825B39D9B599CB651811C3E1B7D3C" lastPageNumber="269" pageId="0" pageNumber="269" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae">
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<subSubSection id="942B4F0D1D899CB0E55E3924CEED469A" pageId="0" pageNumber="269" type="nomenclature">
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<paragraph id="5120C0C2E42A388F76531690F6EE3ABD" pageId="0" pageNumber="269">
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<taxonomicName id="60A202171EF9AF4F992C4556AED8E09A" ID-CoL="8MM75" LSID="http://zoobank.org/B98AC026-EEAD-43A7-B4B0-CF5121A12836" authority="Giachino & Eberhard & Perina, 2021" authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. nov.">Kimberleytyphlus</taxonomicName>
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<taxonomicNameLabel id="0DDF2B38CBAADA68321E639A8C460BFA" pageId="0" pageNumber="269">gen. nov.</taxonomicNameLabel>
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<figureCitation id="97DF63D4E2DDE1E21DC5222A4098359F" captionStart="Figures 34, 35" captionStartId="F11" captionText="Figures 34, 35. Kimberleytyphlus carrboydianus gen. et sp. nov., HT ♂ 34 habitus 35 aedeagus in lateral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1044.58844.figures34-35" httpUri="https://binary.pensoft.net/fig/553770" pageId="0" pageNumber="269">Figs 34-35</figureCitation>
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</paragraph>
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<subSubSection pageId="0" pageNumber="269" type="type species">
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<paragraph pageId="0" pageNumber="269">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="269">
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||||
<taxonomicName authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus carrboydianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="carrboydianus">
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||||
<emphasis italics="true" pageId="0" pageNumber="269">Kimberleytyphlus carrboydianus</emphasis>
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||||
<subSubSection id="4BA2FE7F0C33FAFA6556D7A6B986843D" pageId="0" pageNumber="269" type="type species">
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<paragraph id="727BD552214B8D960DC3CCB9031A6B32" pageId="0" pageNumber="269">Type species.</paragraph>
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||||
<paragraph id="BD50B95F558CEED62F3A8D6C5C986363" pageId="0" pageNumber="269">
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||||
<taxonomicName id="589748483637E082B6D4441C52FFC2F3" authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus carrboydianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="carrboydianus">
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||||
<emphasis id="08682E3EC3EF6A2CB2CD301ECF6A9E44" italics="true" pageId="0" pageNumber="269">Kimberleytyphlus carrboydianus</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="269" type="diagnosis">
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||||
<paragraph pageId="0" pageNumber="269">Diagnosis.</paragraph>
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||||
<paragraph pageId="0" pageNumber="269">
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||||
<subSubSection id="68D62AE3571A13A6C22FEC1AB298071E" pageId="0" pageNumber="269" type="diagnosis">
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<paragraph id="9D35A2215BF1EE909F6FDD3BAB3CC456" pageId="0" pageNumber="269">Diagnosis.</paragraph>
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||||
<paragraph id="7B9408E803B2AC7CDE97CE9F2E731D7A" pageId="0" pageNumber="269">
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||||
Genus of the "
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||||
<taxonomicName class="Insecta" family="Carabidae" genus="Illaphanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Illaphanus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus">
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||||
<emphasis italics="true" pageId="0" pageNumber="269">Illaphanus</emphasis>
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||||
<taxonomicName id="B48C817485DC97DEB17AC28F3A074E0C" class="Insecta" family="Carabidae" genus="Illaphanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Illaphanus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus">
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||||
<emphasis id="736F3A4C83525F0C9B746952E1F8022C" italics="true" pageId="0" pageNumber="269">Illaphanus</emphasis>
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||||
</taxonomicName>
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phyletic series" (sensu
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<bibRefCitation author="Giachino, PM" editor="Daccordi, M" journalOrPublisher="Monografie del Museo regionale di Scienze naturali, Torino" pageId="0" pageNumber="269" pagination="137 - 238" refId="B12" refString="Giachino, PM, 2005. Revision of the Australian Anillina (Coleoptera, Carabidae, Bembidiini). In: Daccordi, M, Giachino, PM, Eds., Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin. II. Monografie del Museo regionale di Scienze naturali, Torino 42: 137 - 238" title="Revision of the Australian Anillina (Coleoptera, Carabidae, Bembidiini)." volume="42" volumeTitle="Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin. II." year="2005">Giachino 2005</bibRefCitation>
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||||
<bibRefCitation id="7D245146CB52628C16A24B3AD101F5F3" author="Giachino, PM" editor="Daccordi, M" journalOrPublisher="Monografie del Museo regionale di Scienze naturali, Torino" pageId="0" pageNumber="269" pagination="137 - 238" refId="B12" refString="Giachino, PM, 2005. Revision of the Australian Anillina (Coleoptera, Carabidae, Bembidiini). In: Daccordi, M, Giachino, PM, Eds., Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin. II. Monografie del Museo regionale di Scienze naturali, Torino 42: 137 - 238" title="Revision of the Australian Anillina (Coleoptera, Carabidae, Bembidiini)." volume="42" volumeTitle="Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin. II." year="2005">Giachino 2005</bibRefCitation>
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||||
), with species strongly characterised by normal metatrochanters; metafemora non dentate; elytra not reduced at tip; elytral disc without longitudinal grooves and bearing three setae, scaly microsculpture and 9th pore of the umbilicate series in normal position (placed after the 8th one); aedeagus with median lobe subrectilinear, size of basal bulb normal and parameres each bearing one apical seta. Labial tooth absent.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="269" type="description">
|
||||
<paragraph pageId="0" pageNumber="269">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">Species of a medium size (TL mm 1.57-1.59) and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
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||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="269">Head</emphasis>
|
||||
<subSubSection id="CD338C57BA5A2580DD70F2F1C87299C0" pageId="0" pageNumber="269" type="description">
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||||
<paragraph id="D3352E4F774E141FA425FC314544A55F" pageId="0" pageNumber="269">Description.</paragraph>
|
||||
<paragraph id="2C66EBEA2CF817247725FD0F5EC53D26" pageId="0" pageNumber="269">Species of a medium size (TL mm 1.57-1.59) and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence.</paragraph>
|
||||
<paragraph id="9FFFA0CF2A1EAF4280B14DE8E8B1A80B" pageId="0" pageNumber="269">
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<emphasis id="CD64211B2B8FD755E4496C713B1CC457" bold="true" italics="true" pageId="0" pageNumber="269">Head</emphasis>
|
||||
large but slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with the submentum. Labial tooth absent. Antennae moniliform, without particular features.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="269">Pronotum</emphasis>
|
||||
<paragraph id="F3539CAD8217B338D2F81D8A137EA13E" pageId="0" pageNumber="269">
|
||||
<emphasis id="FA10BA8902EE013DFA6E0B2FCFBEAD27" bold="true" italics="true" pageId="0" pageNumber="269">Pronotum</emphasis>
|
||||
trapezoidal, with sides not sinuate in the basal third, denticulated at the basal third. Basal angles obtuse, sharp, not rounded; basal border slightly narrower than anterior border; presence of two marginal setae, the posterior one placed near basal angles.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="269">Elytra</emphasis>
|
||||
<paragraph id="2E8325C90087D22144DD10E0343AC615" pageId="0" pageNumber="269">
|
||||
<emphasis id="7021505A11B4F146791F6EB23338A8EF" bold="true" italics="true" pageId="0" pageNumber="269">Elytra</emphasis>
|
||||
subrectangular elongate, separately rounded, not truncated and apically only slightly emarginated, convex, without longitudinal grooves. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area and distinctly crenulate up to the level of 8th-9th pores of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu
|
||||
<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Paris (A)" pageId="0" pageNumber="269" pagination="33 - 204" refId="B33" refString="Jeannel, R, 1963. Monographie des " Anillini ", Bembidiides endoges (Coleoptera Trechidae). Memoires du Museum national d'Histoire naturelle, Paris (A) 28: 33 - 204" title="Monographie des " Anillini ", Bembidiides endoges (Coleoptera Trechidae)." volume="28" year="1963">Jeannel 1963</bibRefCitation>
|
||||
<bibRefCitation id="01262A71C4DBEB1A0EB89095843D5F9C" author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Paris (A)" pageId="0" pageNumber="269" pagination="33 - 204" refId="B33" refString="Jeannel, R, 1963. Monographie des " Anillini ", Bembidiides endoges (Coleoptera Trechidae). Memoires du Museum national d'Histoire naturelle, Paris (A) 28: 33 - 204" title="Monographie des " Anillini ", Bembidiides endoges (Coleoptera Trechidae)." volume="28" year="1963">Jeannel 1963</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Giachino, PM" journalOrPublisher="Biodiversity Journal, monograph" pageId="0" pageNumber="269" pagination="1 - 112" refId="B16" refString="Giachino, PM, Vailati, D, 2011. Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, monograph 1: 1 - 112" title="Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini)." volume="1" year="2011">Giachino and Vailati 2011</bibRefCitation>
|
||||
<bibRefCitation id="84FA6D21CADE66ED1BA187DCB9C721C5" author="Giachino, PM" journalOrPublisher="Biodiversity Journal, monograph" pageId="0" pageNumber="269" pagination="1 - 112" refId="B16" refString="Giachino, PM, Vailati, D, 2011. Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, monograph 1: 1 - 112" title="Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini)." volume="1" year="2011">Giachino and Vailati 2011</bibRefCitation>
|
||||
); disc bearing three setae.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="269">Legs</emphasis>
|
||||
<paragraph id="9A0158B0881E272E51D5A927D3973008" pageId="0" pageNumber="269">
|
||||
<emphasis id="B7E642D934E405BACCCC86B1F22D4044" bold="true" italics="true" pageId="0" pageNumber="269">Legs</emphasis>
|
||||
relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal, two slightly dilated protarsomeres, without adhesive phanerae in males.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="269">Aedeagus</emphasis>
|
||||
<paragraph id="6F0DA6EABBA65DAC207933D5CBAD5F8F" pageId="0" pageNumber="269">
|
||||
<emphasis id="769FFDC1BA265CDB47F9675CFE8D3F3F" bold="true" italics="true" pageId="0" pageNumber="269">Aedeagus</emphasis>
|
||||
relatively small, median lobe long, subrectilinear with basal bulb of normal size. Parameres long, bearing one apical seta. Endophallus without any sclerified phanerae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="269" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="269">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<subSubSection id="4C0FE39479FA0C6496C38102C002E74C" pageId="0" pageNumber="269" type="etymology">
|
||||
<paragraph id="C5EEBB9CDA433799E46E35E37ADFE6AA" pageId="0" pageNumber="269">Etymology.</paragraph>
|
||||
<paragraph id="DC618007E6292CEBAD6EEF49B2910D7D" pageId="0" pageNumber="269">
|
||||
The name combines the name Kimberley (region where the type locality is located) and the Greek suffix -
|
||||
<emphasis italics="true" pageId="0" pageNumber="269">typhlos</emphasis>
|
||||
<emphasis id="F1B23121A12C6395BFD6D6E1E8CB59E5" italics="true" pageId="0" pageNumber="269">typhlos</emphasis>
|
||||
(meaning blind).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="269" type="species included">
|
||||
<paragraph pageId="0" pageNumber="269">Species included.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<subSubSection id="B0790CE2C5D6116BA2026688F5115422" pageId="0" pageNumber="269" type="species included">
|
||||
<paragraph id="F6245075679B252B4243593AB6A27A00" pageId="0" pageNumber="269">Species included.</paragraph>
|
||||
<paragraph id="7DAA2F82E62942597880612AE3D26E28" pageId="0" pageNumber="269">
|
||||
Currently only
|
||||
<taxonomicName lsidName="K. carrboydianus" pageId="0" pageNumber="269" rank="species" species="carrboydianus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="269">K. carrboydianus</emphasis>
|
||||
<taxonomicName id="1B134DAE2C160123DBAE31B0BDBD558C" lsidName="K. carrboydianus" pageId="0" pageNumber="269" rank="species" species="carrboydianus">
|
||||
<emphasis id="12979168E932DD44243FDACACD040AB6" italics="true" pageId="0" pageNumber="269">K. carrboydianus</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. belongs to this genus.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1044.58844.figures34-35" httpUri="https://binary.pensoft.net/fig/553770" pageId="0" pageNumber="269" start="Figures 34, 35" startId="F11">
|
||||
<paragraph pageId="0" pageNumber="269">
|
||||
<emphasis bold="true" pageId="0" pageNumber="269">Figures 34, 35.</emphasis>
|
||||
<taxonomicName authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus carrboydianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="carrboydianus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="269">Kimberleytyphlus carrboydianus</emphasis>
|
||||
<caption id="480A4581903FB92489740DA7B631DDE4" doi="10.3897/zookeys.1044.58844.figures34-35" httpUri="https://binary.pensoft.net/fig/553770" pageId="0" pageNumber="269" start="Figures 34, 35" startId="F11">
|
||||
<paragraph id="25B527E3377B9CBC9E0108C66796F573" pageId="0" pageNumber="269">
|
||||
<emphasis id="A2633595E3A772B618F13EF78D3540B7" bold="true" pageId="0" pageNumber="269">Figures 34, 35.</emphasis>
|
||||
<taxonomicName id="8CE08C2C1E427F6663ED5495AEC4AF5C" authorityName="Giachino & Eberhard & Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Kimberleytyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimberleytyphlus carrboydianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="carrboydianus">
|
||||
<emphasis id="CAD6C15097B1780CADF3A25F6978D386" italics="true" pageId="0" pageNumber="269">Kimberleytyphlus carrboydianus</emphasis>
|
||||
</taxonomicName>
|
||||
gen. et sp. nov., HT ♂
|
||||
<emphasis bold="true" pageId="0" pageNumber="269">34</emphasis>
|
||||
<emphasis id="8AC28727F8034113415F1FFF8B86640B" bold="true" pageId="0" pageNumber="269">34</emphasis>
|
||||
habitus
|
||||
<emphasis bold="true" pageId="0" pageNumber="269">35</emphasis>
|
||||
<emphasis id="D5A6807EF188AEB84E5EEC9BE452E068" bold="true" pageId="0" pageNumber="269">35</emphasis>
|
||||
aedeagus in lateral view. Scale bars: 0.1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
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