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<mods:title id="28293899A519828946ACFB5BEE6F1EBE">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="C6EEB5042D4A5B4DD42C45100E64A197">Leu, Marc</mods:namePart>
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<mods:namePart id="761E40C8A4C0668879BAD3E4BDC1DCA3">Bucher, Hugo</mods:namePart>
<mods:affiliation id="AFB23232761614CEA930986206B177A1">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="625F4313991751B735331C1755F2A4F9">Ji, Cheng</mods:namePart>
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<mods:date id="C53725D54A034AD5695D66915B401146">2022</mods:date>
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<subSubSection id="C338FF806444FF9CFF1BEB5BD15AF987" box="[151,363,1611,1635]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8B9DAC0B6444FF9CFF1BEB5BD15AF987" blockId="45.[151,363,1611,1667]" box="[151,363,1611,1635]" pageId="45" pageNumber="46">
<heading id="D0D51B676444FF9CFF1BEB5BD15AF987" box="[151,363,1611,1635]" fontSize="9" level="2" pageId="45" pageNumber="46" reason="8">
<taxonomicName id="4C22D7886444FF9CFF1BEB5BD13AF987" authority="Orchard, 2005" authorityName="Orchard" authorityYear="2005" box="[151,267,1611,1635]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFF1BEB5BD13AF987" box="[151,267,1611,1635]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
? n. sp. B
</heading>
</paragraph>
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<subSubSection id="C338FF806444FF9CFF1BEB7BD3A1FC27" pageId="45" pageNumber="46" type="description">
<paragraph id="8B9DAC0B6444FF9CFF1BEB7BD105F967" blockId="45.[151,363,1611,1667]" box="[151,308,1643,1667]" pageId="45" pageNumber="46">
<figureCitation id="1319B08E6444FF9CFF1BEB7BD0CEF967" box="[151,255,1643,1667]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="45" pageNumber="46">Figs. 15N</figureCitation>
;
<figureCitation id="1319B08E6444FF9CFE85EB7BD105F967" box="[265,308,1643,1667]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="45" pageNumber="46">16B</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFF1BEBBBD18CF927" blockId="45.[151,776,1707,1923]" box="[151,445,1707,1731]" pageId="45" pageNumber="46">Number of specimens:&gt;10</paragraph>
<paragraph id="8B9DAC0B6444FF9CFF1BEBDCD213F8A7" blockId="45.[151,776,1707,1923]" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFF1BEBDCD139F907" box="[151,264,1740,1763]" italics="true" pageId="45" pageNumber="46">Diagnosis.</emphasis>
Segminate P
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element with subsymmetrical, posteriorly rounded and anteriorly tapered basal cavity. Large terminal, subtriangular, recurved cusp. Anterior process bearing a few denticles only.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFF1BEA5CD3D3FE47" blockId="45.[151,776,1707,1923]" lastBlockId="45.[813,1437,235,963]" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFF1BEA5CD12AF887" box="[151,283,1868,1891]" italics="true" pageId="45" pageNumber="46">Description.</emphasis>
Te element shows a large cusp at the posterior end. In lateral view, the large, recurved, subtriangular cusp is about as high as the whole unit is long. An anterior process with two or three small upright to reclined denticles is present. Te lower margin is straight to concave. In aboral view, the subsymmetrical basal cavity is sub-rounded at the posterior margin and tapers towards the anterior end.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCCEECBBD4EFFD47" blockId="45.[813,1437,235,963]" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFCCEECBBD396FE27" box="[834,935,427,451]" italics="true" pageId="45" pageNumber="46">Remarks.</emphasis>
Tese P
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elements most closely resemble
<taxonomicName id="4C22D7886444FF9CFAE2ECBCD434FE07" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196444FF9CFAE2ECBCD34CFE07" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
aff.
<emphasis id="B95670196444FF9CFC23ECDBD434FE07" box="[943,1029,459,483]" italics="true" pageId="45" pageNumber="46">discreta</emphasis>
</taxonomicName>
, but differ in having a much shorter anterior process and recurved denticles. Tey also bear some superficial resemblance with the P
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elements of the SpathianAnisian genus
<taxonomicName id="4C22D7886444FF9CFBCBEF3BD48FFDA7" authorityName="Hirschmann" authorityYear="1959" box="[1095,1214,555,579]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFBCBEF3BD48FFDA7" box="[1095,1214,555,579]" italics="true" pageId="45" pageNumber="46">Cornudina</emphasis>
</taxonomicName>
, but differ in their conformation of the basal cavity.
<taxonomicName id="4C22D7886444FF9CFB3CEF5BD54DFD87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1200,1404,587,611]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="unicorna" status="n. gen.">
<emphasis id="B95670196444FF9CFB3CEF5BD54DFD87" box="[1200,1404,587,611]" italics="true" pageId="45" pageNumber="46">Urdyella unicorna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626444FF9CFA04EF5CD36BFD60" pageId="45" pageNumber="46" rank="genus">n. gen.</taxonomicNameLabel>
n. sp. usually has a much thinner and larger, sickleshaped cusp and a rounded basal cavity.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1EFBCD396FC67" blockId="45.[813,1437,235,963]" pageId="45" pageNumber="46">
Because of the absence of a posterior process, some elements, especially arched ones, resemble homologous elements of
<taxonomicName id="4C22D7886444FF9CFC10EFFBD41EFCE7" authorityName="Zhang &amp; Yang" authorityYear="1991" box="[924,1071,747,771]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFC10EFFBD41EFCE7" box="[924,1071,747,771]" italics="true" pageId="45" pageNumber="46">Guangxidella</emphasis>
</taxonomicName>
and could possibly be assigned to that genus, but their basal cavity do not match those of
<taxonomicName id="4C22D7886444FF9CFCC6EE3BD3ECFCA7" authorityName="Zhang &amp; Yang" authorityYear="1991" box="[842,989,811,835]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFCC6EE3BD3ECFCA7" box="[842,989,811,835]" italics="true" pageId="45" pageNumber="46">Guangxidella</emphasis>
</taxonomicName>
and resemble instead that of
<taxonomicName id="4C22D7886444FF9CFAA4EE3BD383FC87" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196444FF9CFAA4EE3BD5ADFCA7" box="[1320,1436,811,835]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
aff.
<emphasis id="B95670196444FF9CFCD0EE5BD383FC87" box="[860,946,843,867]" italics="true" pageId="45" pageNumber="46">discreta</emphasis>
</taxonomicName>
, hence we tentatively retain them within
<taxonomicName id="4C22D7886444FF9CFCA1EE7BD390FC67" authorityName="Orchard" authorityYear="2005" box="[813,929,875,899]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFCA1EE7BD390FC67" box="[813,929,875,899]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1EE9CD3A1FC27" blockId="45.[813,1437,235,963]" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFCA1EE9CD399FC47" box="[813,936,908,931]" italics="true" pageId="45" pageNumber="46">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B6444FF9CFB8FEE9BD479FC47" box="[1027,1096,907,931]" name="China" pageId="45" pageNumber="46">China</collectingCountry>
: Luolou, Smithian age, Laren,
<collectingRegion id="49E662E96444FF9CFCA1EEBBD3BCFC27" box="[813,909,939,963]" country="China" name="Guangxi" pageId="45" pageNumber="46">Guangxi</collectingRegion>
.
</paragraph>
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<taxonomicName id="4C22D7886444FF9CFCA1EEFBD390FBE7" authorityName="Orchard" authorityYear="2005" box="[813,929,1003,1027]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFCA1EEFBD390FBE7" box="[813,929,1003,1027]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
? n. sp. C
</heading>
</paragraph>
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<subSubSection id="C338FF806444FF9FFCA1E91BD0CAFD67" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="8B9DAC0B6444FF9CFCA1E91BD43EFBC7" blockId="45.[813,1039,1003,1059]" box="[813,1039,1035,1059]" pageId="45" pageNumber="46">
<figureCitation id="1319B08E6444FF9CFCA1E91BD3A4FBC7" box="[813,917,1035,1059]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="45" pageNumber="46">Figs. 15O</figureCitation>
;
<figureCitation id="1319B08E6444FF9CFC13E91BD43DFBC7" box="[927,1036,1035,1059]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="45" pageNumber="46">16A, F, M</figureCitation>
.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E95BD5B1FB87" blockId="45.[813,1436,1099,1347]" box="[813,1408,1099,1124]" pageId="45" pageNumber="46">
1989
<taxonomicName id="4C22D7886444FF9CFCEBE95BD464FB87" box="[871,1109,1099,1123]" genus="Cratognathodus" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B95670196444FF9CFCEBE95BD427FB87" box="[871,1046,1099,1123]" italics="true" pageId="45" pageNumber="46">Cratognathodus</emphasis>
sp. A
</taxonomicName>
; Tang, p. 405, pl. 32, fig. 8.
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E97BD3B1FB47" blockId="45.[813,1436,1099,1347]" pageId="45" pageNumber="46">
2013
<taxonomicName id="4C22D7886444FF9CFCE6E97BD4FCFB67" authority="Sweet" authorityName="Sweet" box="[874,1229,1131,1155]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="peculiaris">
<emphasis id="B95670196444FF9CFCE6E97BD432FB67" box="[874,1027,1131,1155]" italics="true" pageId="45" pageNumber="46">Neospathodus</emphasis>
?
<emphasis id="B95670196444FF9CFB9BE97BD4B3FB67" box="[1047,1154,1131,1155]" italics="true" pageId="45" pageNumber="46">peculiaris</emphasis>
Sweet
</taxonomicName>
; Yan et al., p. 516, fig.
<specimenCount id="9D2467826444FF9CFCD8E99BD3B1FB47" box="[852,896,1163,1187]" count="6" pageId="45" pageNumber="46" type="generic">6M.</specimenCount>
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E9BBD342FAE7" blockId="45.[813,1436,1099,1347]" pageId="45" pageNumber="46">
2014
<taxonomicName id="4C22D7886444FF9CFCE0E9BBD43AFB27" box="[876,1035,1195,1219]" family="Gondolellidae" genus="Discretella" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95670196444FF9CFCE0E9BBD3D1FB27" box="[876,992,1195,1219]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
sp.
</taxonomicName>
indet. A; Maekawa &amp; Igo in Shigeta et al., p. 202207, fig. 151.1151.12, 151.16151.18 (only).
</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E81BD4A3FAA7" blockId="45.[813,1436,1099,1347]" pageId="45" pageNumber="46">2014 Genus gen. indet. D; Maekawa &amp; Igo in Shigeta et al., p. 269, figs. 193.10193.12.</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E87BD463FA67" blockId="45.[813,1437,1387,1923]" box="[813,1106,1387,1411]" pageId="45" pageNumber="46">Number of specimens:&gt;10</paragraph>
<paragraph id="8B9DAC0B6444FF9CFCA1E89CD399F9C7" blockId="45.[813,1437,1387,1923]" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFCA1E89CD463FA47" box="[813,1106,1419,1443]" italics="true" pageId="45" pageNumber="46">Diagnosis and description.</emphasis>
Same as
<taxonomicName id="4C22D7886444FF9CFB33E89BD502FA47" authorityName="Orchard" authorityYear="2005" box="[1215,1331,1419,1443]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFB33E89BD502FA47" box="[1215,1331,1419,1443]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
? n. sp. B except that a very small posterior process without denticle is always present and the laterally twisted basal cavity tapers at both ends. Te anterior process may bear up to 5 denticles.
</paragraph>
<paragraph id="8B9DAC0B6444FF9FFCCEEB3BD226FDA7" blockId="45.[813,1437,1387,1923]" lastBlockId="46.[151,776,235,643]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="B95670196444FF9CFCCEEB3BD396F9A7" box="[834,935,1579,1603]" italics="true" pageId="45" pageNumber="46">Remarks.</emphasis>
Te P
<subScript id="17A6AE4E6444FF9CFC60EB27D3C4F9AC" attach="left" box="[1004,1013,1591,1608]" fontSize="7" pageId="45" pageNumber="46">1</subScript>
elements of this species are very similar to those of the coeval
<taxonomicName id="4C22D7886444FF9CFBDDEB5BD4F4F987" authorityName="Orchard" authorityYear="2005" box="[1105,1221,1611,1635]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFBDDEB5BD4F4F987" box="[1105,1221,1611,1635]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
? n. sp. B and both species may be conspecific although the presence of the incipient posterior process and the geometry of the basal cavity suggest they correspond at least to two different morphotypes, possibly due to sexual dimorphism. Tey also share strong similarities with the P
<subScript id="17A6AE4E6444FF9CFB67EBE7D4C5F8EC" attach="left" box="[1259,1268,1783,1800]" fontSize="7" pageId="45" pageNumber="46">1</subScript>
of
<emphasis id="B95670196444FF9CFA91EBFBD5ADF8E7" box="[1309,1436,1771,1795]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C22D7886444FF9CFA91EBFBD5A3F8E7" authorityName="Orchard" authorityYear="2005" box="[1309,1426,1771,1795]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">Discretella</taxonomicName>
?
</emphasis>
n. sp. D, which may be a descendant of
<taxonomicName id="4C22D7886444FF9CFB51EA1BD560F8C7" authorityName="Orchard" authorityYear="2005" box="[1245,1361,1803,1827]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B95670196444FF9CFB51EA1BD560F8C7" box="[1245,1361,1803,1827]" italics="true" pageId="45" pageNumber="46">Discretella</emphasis>
</taxonomicName>
? n. sp. B via reduction/loss of the anterior process, forms like,
<figureCitation id="1319B08E6444FF9CFCA1EA5BD3A1F887" box="[813,912,1867,1891]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="45" pageNumber="46">Fig. 16M</figureCitation>
(and figs. 193.10193.12 of Maekawa and Igo in
<bibRefCitation id="EFB3D1FA6444FF9CFCC5EA7BD427F867" author="Shigeta" box="[841,1046,1899,1923]" etAl="et al." firstAuthor="Shigeta" pageId="45" pageNumber="46" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
) that bear only one or two anterior denticles being intermediary. Te relationship with
<taxonomicName id="4C22D7886447FF9FFD55EDFCD121FEC7" family="Gondolellidae" genus="Discretella" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95670196447FF9FFD55EDFCD0D6FEC7" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
sp.
</taxonomicName>
indet. A Maekawa &amp; Igo is unclear but the P
<subScript id="17A6AE4E6447FF9FFF2AEC27D09EFEAC" attach="left" box="[166,175,311,328]" fontSize="7" pageId="46" pageNumber="47">1</subScript>
of the latter is almost identical, except that the incipient posterior process bears a small triangular denticle. Maekawa &amp; Igo noted also that the anterior process of the P
<subScript id="17A6AE4E6447FF9FFF58EC87D0ECFE4C" attach="left" box="[212,221,407,424]" fontSize="7" pageId="46" pageNumber="47">1</subScript>
element of
<taxonomicName id="4C22D7886447FF9FFEE6EC9BD239FE47" box="[362,520,395,419]" family="Gondolellidae" genus="Discretella" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95670196447FF9FFEE6EC9BD1EFFE47" box="[362,478,395,419]" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
sp.
</taxonomicName>
indet. A (Maekawa &amp; Igo) may also have been reduced over time. Conversely,
<taxonomicName id="4C22D7886447FF9FFF1BECDBD245FE07" authority="(Wang &amp; Wang, 1976)" baseAuthorityName="Wang &amp; Wang" baseAuthorityYear="1976" box="[151,628,459,484]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B95670196447FF9FFF1BECDBD140FE07" box="[151,369,459,483]" italics="true" pageId="46" pageNumber="47">Discretella robustus</emphasis>
(
<bibRefCitation id="EFB3D1FA6447FF9FFE0DECDBD25AFE07" author="Wang &amp; Wang" box="[385,619,459,484]" firstAuthor="Wang" pageId="46" pageNumber="47" pagination="387 - 416" refId="ref47753" refString="Wang, C., &amp; Wang, Z. (1976). Triassic conodonts from the Mount Jolmo Lungma region. In Xizang Scientific Expedition Team of Chinese Academy of Sciences (Ed.), A report of scientific expedition in the Mount Jolmo Lungma Region (1966 - 1968), Paleontology (Vol. 2, pp. 387 - 416). Beijing: Science Press. in Chinese; original title translated." type="journal article" year="1976">Wang &amp; Wang, 1976</bibRefCitation>
)
</taxonomicName>
, which share a similar recurved cusp and a sigmoidal profile in lower view, has more developed anterior and posterior processes and may be their forerunner.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BEF5CD0CAFD67" blockId="46.[151,776,235,643]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF1BEF5CD123FD87" box="[151,274,588,611]" italics="true" pageId="46" pageNumber="47">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B6447FF9FFEE2EF5BD182FD87" box="[366,435,587,611]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
: Luolou, Smithian age, Laren,
<collectingRegion id="49E662E96447FF9FFF1BEF7BD0C6FD67" box="[151,247,619,643]" country="China" name="Guangxi" pageId="46" pageNumber="47">Guangxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806447FF9FFF1BEFBBD15FFD27" box="[151,366,683,707]" pageId="46" pageNumber="47" type="multiple">
<paragraph id="8B9DAC0B6447FF9FFF1BEFBBD15FFD27" blockId="46.[151,366,683,739]" box="[151,366,683,707]" pageId="46" pageNumber="47">
<heading id="D0D51B676447FF9FFF1BEFBBD15FFD27" box="[151,366,683,707]" fontSize="9" level="2" pageId="46" pageNumber="47" reason="8">
<taxonomicName id="4C22D7886447FF9FFF1BEFBBD13AFD27" authorityName="Orchard" authorityYear="2005" box="[151,267,683,707]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="B95670196447FF9FFF1BEFBBD13AFD27" box="[151,267,683,707]" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
</taxonomicName>
? n. sp. D
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806447FF9FFF1BEFDBD26DFAA7" pageId="46" pageNumber="47" type="description">
<paragraph id="8B9DAC0B6447FF9FFF1BEFDBD108FD07" blockId="46.[151,366,683,739]" box="[151,313,715,739]" pageId="46" pageNumber="47">
<figureCitation id="1319B08E6447FF9FFF1BEFDBD0EEFD07" box="[151,223,715,739]" captionStart="Fig" captionStartId="46.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 Cornudininae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AW Urdyella unicorna n. gen. n. sp.; A LAR225, PIMUZ 39345; B LAR225, PIMUZ 39346;C LAR229, PIMUZ 39347; D LAR227, PIMUZ 39348;E LAR229,PIMUZ 39349; F LAR229,PIMUZ 39350; G LAR225,PIMUZ 39351; H LAR229,PIMUZ 39352; I QIA123, PIMUZ 39353; J LAR227,PIMUZ 39354; K QIA123, PIMUZ 39355; L QIA123, PIMUZ 39356; M LAR227,PIMUZ 39357; N LAR225, PIMUZ 39358; O QIA120, PIMUZ 39359; P LAR225, PIMUZ 39360; Q QIA121, PIMUZ 39361; R LAR232, PIMUZ 39362; S LAR232, PIMUZ 39363; T QIA124,PIMUZ 39364; U LAR232, PIMUZ 39365; V QIA123, PIMUZ 39366; W SHA341, PIMUZ 39367.X-AA Urdyella tridenta n. gen. n. sp.; X LAR227, PIMUZ 39341; Y LAR227, PIMUZ 39342; Z LAR227, PIMUZ 39343; AA LAR227, PIMUZ 39344.AB, AC Discretella? n. sp. D; AB QIA124, PIMUZ 39118; AC QIA123, PIMUZ 39119. AD Neostrachanognathus n. sp. A; LAR232, PIMUZ 39183. AE Urdyella n. sp. A; LAR232, PIMUZ 39340" figureDoi="http://doi.org/10.5281/zenodo.13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
AB, AC.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BEE1BD185FCA7" blockId="46.[151,775,779,835]" pageId="46" pageNumber="47">2014 Genus gen. indet. D; Maekawa &amp; Igo in Shigeta et al., p. 269, figs. 193.89.</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BEE7BD246FC67" blockId="46.[151,631,875,931]" box="[151,631,875,899]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF1BEE7BD0CEFC67" box="[151,255,875,899]" italics="true" pageId="46" pageNumber="47">
<typeStatus id="549912A96447FF9FFF1BEE7BD0CAFC67" box="[151,251,875,899]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen illustrated in
<figureCitation id="1319B08E6447FF9FFD8AEE7BD27FFC67" box="[518,590,875,899]" captionStart="Fig" captionStartId="46.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 Cornudininae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AW Urdyella unicorna n. gen. n. sp.; A LAR225, PIMUZ 39345; B LAR225, PIMUZ 39346;C LAR229, PIMUZ 39347; D LAR227, PIMUZ 39348;E LAR229,PIMUZ 39349; F LAR229,PIMUZ 39350; G LAR225,PIMUZ 39351; H LAR229,PIMUZ 39352; I QIA123, PIMUZ 39353; J LAR227,PIMUZ 39354; K QIA123, PIMUZ 39355; L QIA123, PIMUZ 39356; M LAR227,PIMUZ 39357; N LAR225, PIMUZ 39358; O QIA120, PIMUZ 39359; P LAR225, PIMUZ 39360; Q QIA121, PIMUZ 39361; R LAR232, PIMUZ 39362; S LAR232, PIMUZ 39363; T QIA124,PIMUZ 39364; U LAR232, PIMUZ 39365; V QIA123, PIMUZ 39366; W SHA341, PIMUZ 39367.X-AA Urdyella tridenta n. gen. n. sp.; X LAR227, PIMUZ 39341; Y LAR227, PIMUZ 39342; Z LAR227, PIMUZ 39343; AA LAR227, PIMUZ 39344.AB, AC Discretella? n. sp. D; AB QIA124, PIMUZ 39118; AC QIA123, PIMUZ 39119. AD Neostrachanognathus n. sp. A; LAR232, PIMUZ 39183. AE Urdyella n. sp. A; LAR232, PIMUZ 39340" figureDoi="http://doi.org/10.5281/zenodo.13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
AB.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BEE9BD18DFC47" blockId="46.[151,631,875,931]" box="[151,444,907,931]" pageId="46" pageNumber="47">Number of specimens:&gt;20</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BEEDCD222FBE7" blockId="46.[151,776,971,1348]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF1BEEDCD139FC07" box="[151,264,972,995]" italics="true" pageId="46" pageNumber="47">Diagnosis.</emphasis>
Same P
<subScript id="17A6AE4E6447FF9FFEE4EEC7D140FC0C" attach="left" box="[360,369,983,1000]" fontSize="7" pageId="46" pageNumber="47">1</subScript>
element as in
<taxonomicName id="4C22D7886447FF9FFDAEEEDBD2A7FC07" authorityName="Orchard" authorityYear="2005" box="[546,662,971,995]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="B95670196447FF9FFDAEEEDBD2A7FC07" box="[546,662,971,995]" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
</taxonomicName>
? n. sp. C except it bears no anterior process.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BE91BD339FB67" blockId="46.[151,776,971,1348]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF1BE91BD0CDFBC7" box="[151,252,1035,1059]" italics="true" pageId="46" pageNumber="47">Remarks.</emphasis>
As for
<taxonomicName id="4C22D7886447FF9FFED8E91BD1F9FBC7" authorityName="Orchard" authorityYear="2005" box="[340,456,1035,1059]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="B95670196447FF9FFED8E91BD1F9FBC7" box="[340,456,1035,1059]" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
</taxonomicName>
? n. sp. B and C (see above) the generic assignment of these forms is uncertain and may deserve a new genus name, especially given that the present taxa bear neither posterior nor anterior denticles.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFF1BE99CD26DFAA7" blockId="46.[151,776,971,1348]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF1BE99CD123FB47" box="[151,274,1164,1187]" italics="true" pageId="46" pageNumber="47">Occurrence</emphasis>
. Laren road cut, Luolou Formation, Smithian age, Nanpanjiang basin, South
<collectingCountry id="F335EC9B6447FF9FFE6BE9BBD218FB27" box="[487,553,1195,1219]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
(this paper); North-eastern
<collectingCountry id="F335EC9B6447FF9FFF79E9DBD165FB07" box="[245,340,1227,1251]" name="Vietnam" pageId="46" pageNumber="47">Vietnam</collectingCountry>
,
<emphasis id="B95670196447FF9FFEEAE9DBD1CFFB07" box="[358,510,1227,1251]" italics="true" pageId="46" pageNumber="47">Novispathodu</emphasis>
s ex gr.
<taxonomicName id="4C22D7886447FF9FFDEAE9DCD336FB07" authority="Zone" authorityName="Zone" box="[614,775,1228,1251]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196447FF9FFDEAE9DCD28EFB07" box="[614,703,1228,1251]" italics="true" pageId="46" pageNumber="47">waageni</emphasis>
Zone
</taxonomicName>
between
<taxonomicName id="4C22D7886447FF9FFE8CE9FBD229FAE7" authorityName="Chao" authorityYear="1959" box="[256,536,1259,1283]" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rursiradiatus">
<emphasis id="B95670196447FF9FFE8CE9FBD229FAE7" box="[256,536,1259,1283]" italics="true" pageId="46" pageNumber="47">Flemingites rursiradiatus</emphasis>
</taxonomicName>
beds and Urdyceras tulongensis beds and within the
<taxonomicName id="4C22D7886447FF9FFD89E81CD2F4FAC7" authorityName="Hyatt &amp; Smith" authorityYear="1905" box="[517,709,1291,1315]" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="koeneni">
<emphasis id="B95670196447FF9FFD89E81CD2F4FAC7" box="[517,709,1291,1315]" italics="true" pageId="46" pageNumber="47">Owenites koeneni</emphasis>
</taxonomicName>
beds, Bac Tuy Formation (
<bibRefCitation id="EFB3D1FA6447FF9FFE0EE83BD27CFAA7" author="Shigeta" box="[386,589,1323,1347]" etAl="et al." firstAuthor="Shigeta" pageId="46" pageNumber="47" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,245 @@
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<paragraph id="8B9DAC0B6445FF9DFF1BEFDBD236FD07" blockId="44.[151,519,715,771]" box="[151,519,715,739]" pageId="44" pageNumber="45">
<heading id="D0D51B676445FF9DFF1BEFDBD236FD07" box="[151,519,715,739]" fontSize="9" level="2" pageId="44" pageNumber="45" reason="8">
<taxonomicName id="4C22D7886445FF9DFF1BEFDBD236FD07" authority="(Muller, 1956)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[151,519,715,739]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF1BEFDBD159FD07" box="[151,360,715,739]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(
<bibRefCitation id="EFB3D1FA6445FF9DFEF9EFDBD1CFFD07" author="Muller" box="[373,510,715,739]" firstAuthor="Muller" pageId="44" pageNumber="45" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
)
</taxonomicName>
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</subSubSection>
<subSubSection id="C338FF806445FF9DFF1BEFFBD1BEFCE7" box="[151,399,747,771]" pageId="44" pageNumber="45" type="description">
<paragraph id="8B9DAC0B6445FF9DFF1BEFFBD1BEFCE7" blockId="44.[151,519,715,771]" box="[151,399,747,771]" pageId="44" pageNumber="45">
<figureCitation id="1319B08E6445FF9DFF1BEFFBD108FCE7" box="[151,313,747,771]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="44" pageNumber="45">Figs. 15C, G, L</figureCitation>
;
<figureCitation id="1319B08E6445FF9DFECFEFFBD1BDFCE7" box="[323,396,747,771]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="44" pageNumber="45">16D, S</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806445FF9DFF1BEE3BD18DFAC7" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="8B9DAC0B6445FF9DFF1BEE3BD16BFC87" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
*1956
<taxonomicName id="4C22D7886445FF9DFF53EE3BD1C5FCA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[223,500,811,835]" family="Kockelellidae" genus="Ctenognathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta" status="sp. nov.">Ctenognathodus discreta</taxonomicName>
<taxonomicNameLabel id="A265CD626445FF9DFE73EE3CD20CFCA7" box="[511,573,812,835]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
; Müller, pp. 821 822, pl. 95, fig. 28.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BEE7BD0C9FC47" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
1989
<taxonomicName id="4C22D7886445FF9DFF59EE7BD205FC67" authority="(Muller)" baseAuthorityName="Muller" box="[213,564,875,899]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF59EE7BD1FFFC67" box="[213,462,875,899]" italics="true" pageId="44" pageNumber="45">Neospathodus discreta</emphasis>
(Müller)
</taxonomicName>
; Tang, p. 402, pl. 30, fig. 7.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BEEBBD242FC27" blockId="44.[151,775,811,1251]" box="[151,627,939,963]" pageId="44" pageNumber="45">
2005
<taxonomicName id="4C22D7886445FF9DFF5EEEBBD1B4FC27" box="[210,389,939,963]" family="Gondolellidae" genus="Discretella" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B95670196445FF9DFF5EEEBBD177FC27" box="[210,326,939,963]" italics="true" pageId="44" pageNumber="45">Discretella</emphasis>
sp. A
</taxonomicName>
; Orchard, p. 83, fig. 8.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BEEDBD106FBE7" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="AB328B256445FF9DFF1BEEDBD106FBE7" pageId="44" pageNumber="45">
2008
<taxonomicName id="4C22D7886445FF9DFF58EEDBD23BFC07" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[212,522,971,995]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF58EEDBD197FC07" box="[212,422,971,995]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(Müller)
</taxonomicName>
;
<treatmentCitation id="0A838A1A6445FF9DFD9BEEDBD333FC07" author="Orchard, M. J." box="[535,770,971,995]" page="402" pageId="44" pageNumber="45" year="2008">
<bibRefCitation id="EFB3D1FA6445FF9DFD9BEEDBD280FC07" author="Orchard" box="[535,689,971,995]" firstAuthor="Orchard" pageId="44" pageNumber="45" pagination="393 - 412" refId="ref45653" refString="Orchard, M. J. (2008). Lower Triassic conodonts from the Canadian Arctic, their intercalibration with ammonoid-based stages and a comparison with other North American Olenekian faunas. Polar Research, 27 (3), 393 - 412. https: // doi. org / 10.1111 / j. 1751 - 8369.2008.00072. x" type="journal article" year="2008">Orchard 2008</bibRefCitation>
, p. 402
</treatmentCitation>
, figs. 8.18, 8.19.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BE91BD106FBA7" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
2010
<taxonomicName id="4C22D7886445FF9DFF5AE91BD221FBC7" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[214,528,1035,1059]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF5AE91BD19BFBC7" box="[214,426,1035,1059]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(Müller)
</taxonomicName>
; Beranek et al., p. 65, figs. 6.18, 6.19.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BE95BD101FB47" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
2014
<taxonomicName id="4C22D7886445FF9DFF55E95BD22BFB87" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[217,538,1099,1123]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF55E95BD180FB87" box="[217,433,1099,1123]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(Müller)
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., pp. 196202, figs. 141.13141.33, 142145, 146.1146.30.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BE9BBD142FB07" blockId="44.[151,775,811,1251]" pageId="44" pageNumber="45">
2018
<taxonomicName id="4C22D7886445FF9DFF5FE9BBD239FB27" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[211,520,1195,1219]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFF5FE9BBD194FB27" box="[211,421,1195,1219]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(Müller)
</taxonomicName>
; Maekawa in Maekawa et al., p. 23, fig. 14.4.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFF1BE81BD18DFAC7" blockId="44.[151,444,1291,1315]" box="[151,444,1291,1315]" pageId="44" pageNumber="45">Number of specimens:&gt;20</paragraph>
</subSubSection>
<subSubSection id="C338FF806445FF9DFF1BE85DD1F9FA80" box="[151,456,1356,1381]" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8B9DAC0B6445FF9DFF1BE85DD1F9FA80" blockId="44.[151,776,1356,1925]" box="[151,456,1356,1381]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFF1BE85DD139FA80" box="[151,264,1357,1380]" italics="true" pageId="44" pageNumber="45">Diagnosis.</emphasis>
See
<bibRefCitation id="EFB3D1FA6445FF9DFEB5E85CD1F0FA80" author="Muller" box="[313,449,1356,1380]" firstAuthor="Muller" pageId="44" pageNumber="45" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806445FF9DFF1BE87DD153F9E1" pageId="44" pageNumber="45" type="description">
<paragraph id="8B9DAC0B6445FF9DFF1BE87DD153F9E1" blockId="44.[151,776,1356,1925]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFF1BE87DD12AFA60" box="[151,283,1389,1412]" italics="true" pageId="44" pageNumber="45">Description.</emphasis>
Te carminate-to-segminate P
<subScript id="17A6AE4E6445FF9DFDFDE869D24BFA6E" attach="left" box="[625,634,1401,1418]" fontSize="7" pageId="44" pageNumber="45">1</subScript>
element has an upturned and inverted basal margin in the posterior one-third to one-half of the element. Te cusp is usually larger than the other discrete and upright denticles but not conspicuously.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806445FF9DFF20EB1DD397FDC7" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8B9DAC0B6445FF9DFF20EB1DD523FE87" blockId="44.[151,776,1356,1925]" lastBlockId="44.[813,1437,235,547]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFF20EB1DD13CF9C1" box="[172,269,1549,1573]" italics="true" pageId="44" pageNumber="45">Remarks.</emphasis>
Maekawa &amp; Igo (in
<bibRefCitation id="EFB3D1FA6445FF9DFE54EB1DD2A4F9C1" author="Shigeta" box="[472,661,1549,1573]" etAl="et al." firstAuthor="Shigeta" pageId="44" pageNumber="45" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
) described two different morphotypes, A and B, for
<taxonomicName id="4C22D7886445FF9DFDB1EB3CD330F9A0" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[573,769,1580,1604]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFDB1EB3CD330F9A0" box="[573,769,1580,1604]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
</taxonomicName>
. Teir morphotype A would correspond to the
<typeStatus id="549912A96445FF9DFD27EB5CD336F980" box="[683,775,1612,1636]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFB3D1FA6445FF9DFF12EB7CD117F960" author="Muller" box="[158,294,1644,1668]" firstAuthor="Muller" pageId="44" pageNumber="45" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
) and their morphotype B has an upturned posterior margin and a relatively broader and triangular cusp. Although Maekawa &amp; Igo (in
<bibRefCitation id="EFB3D1FA6445FF9DFD84EBBCD2F8F921" author="Shigeta" box="[520,713,1708,1733]" etAl="et al." firstAuthor="Shigeta" pageId="44" pageNumber="45" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
) have recently illustrated numerous specimens, the intraspecific variation of this species remains unclear due to its general scarcity. Based on their illustrations and unpublished material, there seems to be scope for further differentiation. Both morphotypes A and B have a posterior process and hence a tapered posterior margin of the basal cavity. However, some of our specimens do not develop a posterior process and have a rounded posterior margin at the basal cavity. Tose are here assigned to a different species, which for now is kept in open nomenclature (
<taxonomicName id="4C22D7886445FF9DFB91EC5BD535FE87" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[1053,1284,331,355]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFB91EC5BD4BAFE87" box="[1053,1163,331,355]" italics="true" pageId="44" pageNumber="45">Discretella</emphasis>
aff.
<emphasis id="B95670196445FF9DFB38EC5BD535FE87" box="[1204,1284,331,355]" italics="true" pageId="44" pageNumber="45">discreta</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCCEEC7CD397FDC7" blockId="44.[813,1437,235,547]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFCCEEC7CD387FE67" box="[834,950,364,387]" italics="true" pageId="44" pageNumber="45">Occurrence</emphasis>
. Wide global occurrence in the Smithian.
<taxonomicName id="4C22D7886445FF9DFAEBEC7CD348FE47" class="Cephalopoda" family="Prionitidae" genus="Meekoceras" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis id="B95670196445FF9DFAEBEC7CD348FE47" italics="true" pageId="44" pageNumber="45">Meekoceras</emphasis>
</taxonomicName>
beds, Nevada (
<bibRefCitation id="EFB3D1FA6445FF9DFB93EC9BD494FE47" author="Muller" box="[1055,1189,395,419]" firstAuthor="Muller" pageId="44" pageNumber="45" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
).
<taxonomicName id="4C22D7886445FF9DFB37EC9BD525FE27" authority="Zone, Canadian Arctic (Orchard, 2008)" authorityName="Zone, Canadian Arctic (Orchard" authorityYear="2008" class="Cephalopoda" family="Flemingitidae" genus="Euflemingites" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="romunderi">
<emphasis id="B95670196445FF9DFB37EC9BD367FE27" italics="true" pageId="44" pageNumber="45">Euflemingites romunderi</emphasis>
Zone, Canadian Arctic (
<bibRefCitation id="EFB3D1FA6445FF9DFBE2ECBBD53BFE27" author="Orchard" box="[1134,1290,427,451]" firstAuthor="Orchard" pageId="44" pageNumber="45" pagination="393 - 412" refId="ref45653" refString="Orchard, M. J. (2008). Lower Triassic conodonts from the Canadian Arctic, their intercalibration with ammonoid-based stages and a comparison with other North American Olenekian faunas. Polar Research, 27 (3), 393 - 412. https: // doi. org / 10.1111 / j. 1751 - 8369.2008.00072. x" type="journal article" year="2008">Orchard, 2008</bibRefCitation>
)
</taxonomicName>
. Jabal Safra,
<collectingCountry id="F335EC9B6445FF9DFCA1ECDBD341FE07" box="[813,880,459,483]" name="Oman" pageId="44" pageNumber="45">Oman</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6445FF9DFCF3ECDBD429FE07" author="Orchard" box="[895,1048,459,483]" firstAuthor="Orchard" pageId="44" pageNumber="45" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
). North
<collectingCountry id="F335EC9B6445FF9DFBFBECDBD4E1FE07" box="[1143,1232,459,483]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
(Tang, 1989).
<emphasis id="B95670196445FF9DFAF5ECDCD5A7FE07" box="[1401,1430,460,483]" italics="true" pageId="44" pageNumber="45">Nv</emphasis>
. ex. gr.
<taxonomicName id="4C22D7886445FF9DFCF1ECFCD4EBFDE7" authority="Zone, Taho Formation" authorityName="Zone, Taho Formation" box="[893,1242,491,515]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196445FF9DFCF1ECFCD3E3FDE7" box="[893,978,492,515]" italics="true" pageId="44" pageNumber="45">waageni</emphasis>
Zone, Taho Formation
</taxonomicName>
,
<collectingCountry id="F335EC9B6445FF9DFB66ECFCD515FDE7" box="[1258,1316,492,515]" name="Japan" pageId="44" pageNumber="45">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6445FF9DFAB4ECFBD3A8FDC7" author="Maekawa" etAl="et al." firstAuthor="Maekawa" pageId="44" pageNumber="45" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,395 @@
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<paragraph id="8B9DAC0B644DFF95FF1BE91BD240FBC7" blockId="36.[151,625,1035,1091]" box="[151,625,1035,1060]" pageId="36" pageNumber="37">
<heading id="D0D51B67644DFF95FF1BE91BD240FBC7" box="[151,625,1035,1060]" fontSize="9" level="2" pageId="36" pageNumber="37" reason="8">
<taxonomicName id="4C22D788644DFF95FF1BE91BD240FBC7" authority="(Sweet, 1970)" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[151,625,1035,1060]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9567019644DFF95FF1BE91BD108FBC7" box="[151,313,1035,1059]" italics="true" pageId="36" pageNumber="37">Novispathodus</emphasis>
ex gr.
<emphasis id="B9567019644DFF95FEF3E91CD1E9FBC7" box="[383,472,1036,1059]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
(
<bibRefCitation id="EFB3D1FA644DFF95FE69E91BD259FBC7" author="Sweet" box="[485,616,1035,1059]" firstAuthor="Sweet" pageId="36" pageNumber="37" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C338FF80644DFF94FF1BE93BD1F0FE47" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="8B9DAC0B644DFF95FF1BE93BD176FBA7" blockId="36.[151,625,1035,1091]" box="[151,327,1067,1091]" pageId="36" pageNumber="37">
<figureCitation id="1319B08E644DFF95FF1BE93BD176FBA7" box="[151,327,1067,1091]" captionStart="Fig" captionStartId="21.[169,200,1283,1303]" captionTargetBox="[153,1438,238,1422]" captionTargetId="figure-168@21.[229,1363,254,1271]" captionTargetPageId="21" captionText="Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181" figureDoi="http://doi.org/10.5281/zenodo.12000270" httpUri="https://zenodo.org/record/12000270/files/figure.png" pageId="36" pageNumber="37">Fig. 14C, D, F, G</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE97BD17BFB47" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
*1970
<taxonomicName id="4C22D788644DFF95FF51E97BD1E9FB67" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[221,472,1131,1155]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni" status="sp. nov.">
<emphasis id="B9567019644DFF95FF51E97BD1E9FB67" box="[221,472,1131,1155]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62644DFF95FE6CE97CD22DFB67" box="[480,540,1132,1155]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
; Sweet, pp. 260261, pl. 1, figs. 11, 12.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE9BBD0C8FB07" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1977
<taxonomicName id="4C22D788644DFF95FF59E9BBD22CFB27" authority="Sweet" authorityName="Sweet" box="[213,541,1195,1219]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF59E9BBD1E1FB27" box="[213,464,1195,1219]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Goel, p. 1094, pl. 2, figs. 14.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE9FBD1FEFAC7" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1978
<taxonomicName id="4C22D788644DFF95FF69E9FBD27FFAE7" authority="Sweet" authorityName="Sweet" box="[229,590,1259,1283]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF69E9FBD1C0FAE7" box="[229,497,1259,1283]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Weitschat &amp; Lehmann, pl. 14, figs. 1112.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE83BD0E3FA87" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1979
<taxonomicName id="4C22D788644DFF95FF59E83BD22DFAA7" authority="Sweet" authorityName="Sweet" box="[213,540,1323,1347]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF59E83BD1E1FAA7" box="[213,464,1323,1347]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Solien, p. 292, pl. 3, fig. 9.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE87BD199FA47" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1980
<taxonomicName id="4C22D788644DFF95FF5FE87BD227FA67" authority="Sweet" authorityName="Sweet" box="[211,534,1387,1411]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5FE87BD1FDFA67" box="[211,460,1387,1411]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Chhabra &amp; Sahni, pl. 1, figs. 910, 14?, 16, 20?.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE8BBD125FA07" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1982
<taxonomicName id="4C22D788644DFF95FF5BE8BBD212FA27" authority="Sweet" authorityName="Sweet" box="[215,547,1451,1475]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5BE8BBD1E5FA27" box="[215,468,1451,1475]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Koike, p. 39, pl. 6, figs. 2427.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BE8FBD170F9C7" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1983
<taxonomicName id="4C22D788644DFF95FF5AE8FBD211F9E7" authority="Sweet" authorityName="Sweet" box="[214,544,1515,1539]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5AE8FBD1E3F9E7" box="[214,466,1515,1539]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Matsuda, p. 8891, pl. 1, figs. 610.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEB3BD122F987" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1984
<taxonomicName id="4C22D788644DFF95FF5EEB3BD222F9A7" authority="Sweet" authorityName="Sweet" box="[210,531,1579,1603]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5EEB3BD1FBF9A7" box="[210,458,1579,1603]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Berry et al., p. 133, pl. 1, figs. 14.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEB7BD1CFF947" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
1984
<taxonomicName id="4C22D788644DFF95FF5BEB7BD212F967" authority="Sweet" authorityName="Sweet" box="[215,547,1643,1667]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5BEB7BD1E4F967" box="[215,469,1643,1667]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Dagis, p. 24, pl. 7, figs. 25, 710, pl. 8. fig. 7 (only).
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEBBBD2CAF927" blockId="36.[151,775,1131,1923]" box="[151,763,1707,1731]" pageId="36" pageNumber="37">
2004
<taxonomicName id="4C22D788644DFF95FF5EEBBBD222F927" authority="Sweet" authorityName="Sweet" box="[210,531,1707,1731]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF5EEBBBD1FBF927" box="[210,458,1707,1731]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Zhao et al., figs. 1, 3.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEBDBD2A2F8E6" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
2007
<taxonomicName id="4C22D788644DFF95FF59EBDBD22FF907" authority="Sweet" authorityName="Sweet" box="[213,542,1739,1763]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF59EBDBD1E0F907" box="[213,465,1739,1763]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Zhao &amp; Orchard in Zhao et al., pp. 3637, pl. 1, figs. 5A, B, 10A, B.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEA1BD19BF8A7" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
2007
<taxonomicName id="4C22D788644DFF95FF55EA1BD143F8C7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[217,370,1803,1827]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9567019644DFF95FF55EA1BD143F8C7" box="[217,370,1803,1827]" italics="true" pageId="36" pageNumber="37">Neospathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788644DFF95FE42EA1CD249F8C7" authority="Sweet" authorityName="Sweet" box="[462,632,1803,1827]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FE42EA1CD216F8C7" box="[462,551,1804,1827]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
Sweet
</taxonomicName>
; Orchard &amp; Krystyn, plate, figs. 818.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FF1BEA5BD1BEF867" blockId="36.[151,775,1131,1923]" pageId="36" pageNumber="37">
2008
<taxonomicName id="4C22D788644DFF95FF53EA5BD20DF887" authority="Sweet" authorityName="Sweet" box="[223,572,1867,1891]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FF53EA5BD1D4F887" box="[223,485,1867,1891]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Nakrem et al., figs. 5.7, 5.8, 5.11, 5.14.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EDFBD43BFEC7" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2008
<taxonomicName id="4C22D788644DFF95FCEBEDFBD499FEE7" authority="Sweet" authorityName="Sweet" box="[871,1192,235,259]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCEBEDFBD46EFEE7" box="[871,1119,235,259]" italics="true" pageId="36" pageNumber="37">Neospathodus waageni</emphasis>
Sweet
</taxonomicName>
; Orchard, p. 406, pl. 8, figs. 8.1, 8.2, 8.8, 8.9.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EC3BD48DFE87" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2009
<taxonomicName id="4C22D788644DFF95FCE6EC3BD4F8FEA7" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[874,1225,299,323]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCE6EC3BD45CFEA7" box="[874,1133,299,323]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Orchard &amp; Zonneveld, p. 785, figs. 13.113.10, 14, 15.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EC7BD49CFE27" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2009
<taxonomicName id="4C22D788644DFF95FCE6EC7BD432FE67" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[874,1027,363,387]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9567019644DFF95FCE6EC7BD432FE67" box="[874,1027,363,387]" italics="true" pageId="36" pageNumber="37">Neospathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788644DFF95FBDCEC7CD4C4FE67" authority="Sweet" authorityName="Sweet" box="[1104,1269,363,387]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FBDCEC7CD498FE67" box="[1104,1193,364,387]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
Sweet
</taxonomicName>
; Igo in Shigeta et al., p. 194, figs. 152.1, 152.3, 152.1415?, 152.1618, 152.19?, 153.89, 156.9, 156.1419.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1ECDBD38CFDE7" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2010
<taxonomicName id="4C22D788644DFF95FCFCECDBD4ECFE07" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[880,1245,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCFCECDBD44AFE07" box="[880,1147,459,483]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Beranek et al., figs. 6.2223.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EF1BD3B2FD87" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2012a
<taxonomicName id="4C22D788644DFF95FCF5EF1BD4D4FDC7" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[889,1253,523,547]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF5EF1BD4B5FDC7" box="[889,1156,523,547]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Goudemand &amp; Orchard in Goudemand et al., p. 1031, figs. 3C?, D, E, H, N, S.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EF7BD4BEFD47" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2013
<taxonomicName id="4C22D788644DFF95FCF9EF7BD4C3FD67" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[885,1266,619,643]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF9EF7BD4B8FD67" box="[885,1161,619,643]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Zhao et al., figs. 9CC, 10A-H, 11F?, 11P-R?.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EFBBD380FD07" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2014
<taxonomicName id="4C22D788644DFF95FCE7EFBBD445FD27" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[875,1140,683,707]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni" status="subsp. nov.">
<emphasis id="B9567019644DFF95FCE7EFBBD445FD27" box="[875,1140,683,707]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62644DFF95FBF1EFBCD4D0FD27" box="[1149,1249,683,707]" pageId="36" pageNumber="37" rank="subSpecies">n. subsp.</taxonomicNameLabel>
A; Goudemand, figs. 1A1D
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EFFBD3F8FC87" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2014
<taxonomicName id="4C22D788644DFF95FCE1EFFBD422FCE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[877,1043,747,771]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9567019644DFF95FCE1EFFBD422FCE7" box="[877,1043,747,771]" italics="true" pageId="36" pageNumber="37">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788644DFF95FBE4EFFCD515FCE7" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[1128,1316,747,771]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FBE4EFFCD4F2FCE7" box="[1128,1219,748,771]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
(Sweet)
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., p. 244, figs. 174.31174.57, 175 178, 179.1179.3, 179.7179.12, 179.16179.30, 180, 181.1181.27.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EE7BD4E2FC47" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2015
<taxonomicName id="4C22D788644DFF95FCF8EE7BD4C0FC67" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[884,1265,875,899]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF8EE7BD4B9FC67" box="[884,1160,875,899]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., figs. 6.23, 7.11, 8.3, 8.8, 8.9, 8.11, 8.14.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EEBBD34FFC07" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2016
<taxonomicName id="4C22D788644DFF95FCF8EEBBD4C1FC27" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[884,1264,939,963]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF8EEBBD4B6FC27" box="[884,1159,939,963]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Liang et al., fig. 4.8.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1EEFBD393FBA7" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2018
<taxonomicName id="4C22D788644DFF95FCE1EEFBD422FBE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[877,1043,1003,1027]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B9567019644DFF95FCE1EEFBD422FBE7" box="[877,1043,1003,1027]" italics="true" pageId="36" pageNumber="37">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788644DFF95FBE4EEFCD515FBE7" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[1128,1316,1003,1027]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FBE4EEFCD4F2FBE7" box="[1128,1219,1004,1027]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
(Sweet)
</taxonomicName>
; Maekawa in Maekawa et al., figs. 22.3, 22.1011, 22.13?, 23.14 15, 23.18?.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E95BD371FB67" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2018
<taxonomicName id="4C22D788644DFF95FCE6E95BD4E3FB87" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[874,1234,1099,1123]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCE6E95BD442FB87" box="[874,1139,1099,1123]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Lyu et al., figs. 5, 6.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E99BD382FB27" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2019
<taxonomicName id="4C22D788644DFF95FCF4E99BD4CBFB47" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[888,1274,1163,1187]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF4E99BD4BFFB47" box="[888,1166,1163,1187]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Lyu et al., figs. 7.810.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E9DBD44DFAE7" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2019
<taxonomicName id="4C22D788644DFF95FCF4E9DBD4CDFB07" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[888,1276,1227,1251]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCF4E9DBD4A1FB07" box="[888,1168,1227,1251]" italics="true" pageId="36" pageNumber="37">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Souquet &amp; Goudemand, figs. 3, 4a, 4g-
<quantity id="4CDA01EE644DFF95FBEDE9FBD448FAE7" box="[1121,1145,1259,1283]" metricMagnitude="3" metricUnit="kg" metricValue="4.0" pageId="36" pageNumber="37" unit="t" value="4.0">4t</quantity>
.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E81BD470FAA7" blockId="36.[813,1437,235,1347]" pageId="36" pageNumber="37">
2019
<taxonomicName id="4C22D788644DFF95FCE3E81BD53EFAC7" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[879,1295,1291,1315]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644DFF95FCE3E81BD424FAC7" box="[879,1045,1291,1315]" italics="true" pageId="36" pageNumber="37">Novispathodus</emphasis>
aff.
<emphasis id="B9567019644DFF95FBDDE81CD49DFAC7" box="[1105,1196,1292,1315]" italics="true" pageId="36" pageNumber="37">waageni</emphasis>
(Sweet)
</taxonomicName>
; Souquet &amp; Goudemand, figs. 4b
<specimenCount id="9D246782644DFF95FBA8E83BD470FAA7" box="[1060,1089,1323,1347]" count="4" pageId="36" pageNumber="37" type="generic">4f.</specimenCount>
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E87BD460FA67" blockId="36.[813,1105,1387,1411]" box="[813,1105,1387,1411]" pageId="36" pageNumber="37">
<emphasis id="B9567019644DFF95FCA1E87BD422FA67" box="[813,1043,1387,1411]" italics="true" pageId="36" pageNumber="37">Number of specimens</emphasis>
.&gt; 50.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E8BCD451FA27" blockId="36.[813,1437,1451,1891]" box="[813,1120,1451,1475]" pageId="36" pageNumber="37">
<emphasis id="B9567019644DFF95FCA1E8BCD3A6FA27" box="[813,919,1452,1475]" italics="true" pageId="36" pageNumber="37">Diagnosis</emphasis>
. See
<bibRefCitation id="EFB3D1FA644DFF95FC42E8BBD46DFA27" author="Sweet" box="[974,1116,1451,1475]" firstAuthor="Sweet" pageId="36" pageNumber="37" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet (1970)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9DAC0B644DFF95FCA1E8DBD3ECF887" blockId="36.[813,1437,1451,1891]" pageId="36" pageNumber="37">
<emphasis id="B9567019644DFF95FCA1E8DBD3A0FA07" box="[813,913,1483,1507]" italics="true" pageId="36" pageNumber="37">Remarks.</emphasis>
Despite the exclusion of several former morphotypes which have been formally described as separate species, this common species still encompasses a lot of variation in the P
<subScript id="17A6AE4E644DFF95FBB5EB27D473F9AC" attach="left" box="[1081,1090,1591,1608]" fontSize="7" pageId="36" pageNumber="37">1</subScript>
element and may deserve further differentiation in the future. Up to six morphotypes have been recognized by distinct authors, e.g.
<bibRefCitation id="EFB3D1FA644DFF95FAA4EB7BD342F947" author="Zhao" etAl="et al." firstAuthor="Zhao" pageId="36" pageNumber="37" pagination="41 - 43" refId="ref49023" refString="Zhao, L., Orchard, M. J., &amp; Tong, J. (2004). Lower Triassic conodont biostratigraphy and speciation of Neospathodus waageni around the Induan - Olenekian boundary of Chaohu, Anhui Province, China. Albertiana, 29, 41 - 43." type="journal article" year="2004">Zhao et al. (2004)</bibRefCitation>
or
<bibRefCitation id="EFB3D1FA644DFF95FC1BEB9BD4FAF947" author="Orchard and Krystyn" box="[919,1227,1675,1700]" firstAuthor="Orchard" pageId="36" pageNumber="37" pagination="30 - 34" refId="ref45762" refString="Orchard, M. J., &amp; Krystyn, L. (2007). Conodonts from the Induan - Olenekian boundary interval at Mud, Spiti. Albertiana, 35, 30 - 34." type="journal article" year="2007">Orchard and Krystyn (2007)</bibRefCitation>
, and several others later by Goudemand (unpublished Ph.D. thesis, 2011). It is not yet clear whether these various morphotypes may be of any utility and hence whether some of those may deserve assignment to separate species (see for instance
<bibRefCitation id="EFB3D1FA644DFF95FCA1EA3CD3E7F8A7" author="Lyu" box="[813,982,1835,1860]" etAl="et al." firstAuthor="Lyu" pageId="36" pageNumber="37" pagination="824 - 836" refId="ref44419" refString="Lyu, Z., Orchard, M. J., Chen, Z. Q., Zhao, L., Zhang, L., &amp; Zhang, X. (2018). A taxonomic re-assessment of the Novispathodus waageni group and its role in defining the base of the Olenekian (Lower Triassic). Journal of Earth Science, 29 (4), 824 - 836. https: // doi. org / 10.1007 / s 12583 - 018 - 0795 - 3" type="journal article" year="2018">Lyu et al., 2018</bibRefCitation>
for a discussion on the value of
<taxonomicName id="4C22D788644DFF95FAC8EA3CD3FEF887" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="subSpecies" species="waageni" subSpecies="eowaageni">
<emphasis id="B9567019644DFF95FAC8EA3CD3FEF887" italics="true" pageId="36" pageNumber="37">N. waageni eowaageni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9DAC0B644CFF94FF21EDFCD1F0FE47" blockId="37.[151,775,235,419]" pageId="37" pageNumber="38">
<emphasis id="B9567019644CFF94FF21EDFCD11AFEE7" box="[173,299,236,259]" italics="true" pageId="37" pageNumber="38">Occurrence</emphasis>
.
<taxonomicName id="4C22D788644CFF94FEB6EDFCD186FEE7" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[314,439,236,259]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019644CFF94FEB6EDFCD186FEE7" box="[314,439,236,259]" italics="true" pageId="37" pageNumber="38">N. waageni</emphasis>
</taxonomicName>
is a very common species in Smithian rocks worldwide. Its FAD has been proposed as a proxy for defining the base of the Olenekian (see discussions in e.g.
<bibRefCitation id="EFB3D1FA644CFF94FEFEEC5BD20CFE87" author="Goudemand" box="[370,573,331,355]" firstAuthor="Goudemand" pageId="37" pageNumber="38" pagination="49 - 51" refId="ref41034" refString="Goudemand, N. (2014). Note on the conodonts from the Induan - Olenekian boundary. Albertiana, 42, 49 - 51." type="journal article" year="2014">Goudemand 2014</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644CFF94FDC2EC5CD330FE87" author="Lyu" box="[590,769,331,356]" etAl="et al." firstAuthor="Lyu" pageId="37" pageNumber="38" pagination="824 - 836" refId="ref44419" refString="Lyu, Z., Orchard, M. J., Chen, Z. Q., Zhao, L., Zhang, L., &amp; Zhang, X. (2018). A taxonomic re-assessment of the Novispathodus waageni group and its role in defining the base of the Olenekian (Lower Triassic). Journal of Earth Science, 29 (4), 824 - 836. https: // doi. org / 10.1007 / s 12583 - 018 - 0795 - 3" type="journal article" year="2018">Lyu et al., 2018</bibRefCitation>
; Orchard, 2007, 2010;
<bibRefCitation id="EFB3D1FA644CFF94FE19EC7BD28DFE67" author="Orchard &amp; Krystyn" box="[405,700,363,388]" firstAuthor="Orchard" pageId="37" pageNumber="38" pagination="30 - 34" refId="ref45762" refString="Orchard, M. J., &amp; Krystyn, L. (2007). Conodonts from the Induan - Olenekian boundary interval at Mud, Spiti. Albertiana, 35, 30 - 34." type="journal article" year="2007">Orchard &amp; Krystyn, 2007</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644CFF94FD41EC7BD121FE47" author="Zhao" etAl="et al." firstAuthor="Zhao" pageId="37" pageNumber="38" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al., 2007</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644CFF94FE90EC9BD182FE47" author="Shigeta" box="[284,435,395,419]" firstAuthor="Shigeta" pageId="37" pageNumber="38" refId="ref46464" refString="Shigeta, Y., Zakharov, Y. D., Maeda, H., &amp; Popov, A. M. (Eds.). 2009. The Lower Triassic System in the Abrek Bay Area, South Primorye, Russia" type="book" year="2009">Shigeta, 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,358 @@
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Genus
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<taxonomicNameLabel id="A265CD626458FF80FEFDEDFCD1E3FEE7" box="[369,466,236,260]" pageId="49" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
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species.
<taxonomicName id="4C22D7886458FF80FEABEC3BD1DFFEA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[295,494,299,323]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="unicorna" status="sp. nov.">Urdyella unicorna</taxonomicName>
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</paragraph>
<paragraph id="8B9DAC0B6458FF80FF1BEC5CD1EEFE67" blockId="49.[151,775,299,419]" pageId="49" pageNumber="50">
<materialsCitation id="3B4AA6566458FF80FF1BEC5CD1EAFE67" country="China" location="Luolou Formation" municipality="Laren" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B95670196458FF80FF1BEC5CD184FE87" box="[151,437,331,355]" italics="true" pageId="49" pageNumber="50">
<typeStatus id="549912A96458FF80FF1BEC5CD0FAFE87" box="[151,203,332,355]" pageId="49" pageNumber="50">Type</typeStatus>
stratum and locality.
</emphasis>
<location id="8EFDFAD06458FF80FE37EC5BD2B5FE87" LSID="urn:lsid:plazi:treatment:038B1D1D6458FF80FF1BEDFBD4E5FE47:8EFDFAD06458FF80FE37EC5BD2B5FE87" box="[443,644,331,355]" country="China" municipality="Laren" name="Luolou Formation" pageId="49" pageNumber="50" stateProvince="Guangxi">Luolou Formation</location>
,
<collectingMunicipality id="6BF936716458FF80FD03EC5CD2FFFE87" box="[655,718,332,355]" pageId="49" pageNumber="50">Laren</collectingMunicipality>
road cut,
<collectingRegion id="49E662E96458FF80FF4BEC7BD1BDFE67" box="[199,396,363,387]" country="China" name="Guangxi" pageId="49" pageNumber="50">Guangxi Province</collectingRegion>
,
<collectingCountry id="F335EC9B6458FF80FE1AEC7BD1EAFE67" box="[406,475,363,387]" name="China" pageId="49" pageNumber="50">China</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DAC0B6458FF80FF1BEC9BD272FE47" blockId="49.[151,775,299,419]" box="[151,579,395,419]" pageId="49" pageNumber="50">
<emphasis id="B95670196458FF80FF1BEC9BD139FE47" box="[151,264,395,419]" italics="true" pageId="49" pageNumber="50">Etymology</emphasis>
: Named after Severine Urdy.
</paragraph>
<paragraph id="8B9DAC0B6458FF80FF1BECDCD1D2FDA7" blockId="49.[151,775,459,1923]" pageId="49" pageNumber="50">
<emphasis id="B95670196458FF80FF1BECDCD130FE07" box="[151,257,460,483]" italics="true" pageId="49" pageNumber="50">Diagnosis</emphasis>
. Short coniform-to-segminiscaphate P
<subScript id="17A6AE4E6458FF80FD4AECC7D2FEFE0C" attach="left" box="[710,719,471,488]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
element with a long cusp, usually at least twice the length of the adjacent denticle, a very short anterior process, and a broadly excavated basal cavity.
</paragraph>
<paragraph id="8B9DAC0B6458FF80FF21EF5BD2D6FA87" blockId="49.[151,775,459,1923]" pageId="49" pageNumber="50">
<emphasis id="B95670196458FF80FF21EF5BD13CFD87" box="[173,269,587,611]" italics="true" pageId="49" pageNumber="50">Remarks</emphasis>
. Te relationship of this genus to
<taxonomicName id="4C22D7886458FF80FD01EF5BD336FD87" authorityName="Hirschmann" authorityYear="1959" box="[653,775,587,611]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD01EF5BD336FD87" box="[653,775,587,611]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
is uncertain (
<bibRefCitation id="EFB3D1FA6458FF80FEA1EF7BD1E0FD67" author="Orchard" box="[301,465,619,643]" firstAuthor="Orchard" pageId="49" pageNumber="50" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
, 2007). P
<subScript id="17A6AE4E6458FF80FDB0EF67D274FD6C" attach="left" box="[572,581,631,648]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
elements similar to the P
<subScript id="17A6AE4E6458FF80FF7FEF87D0CDFD4C" attach="left" box="[243,252,663,680]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
element of
<taxonomicName id="4C22D7886458FF80FE0AEF9BD1D6FD47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[390,487,651,675]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B95670196458FF80FE0AEF9BD1D6FD47" box="[390,487,651,675]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626458FF80FE7CEF9CD20CFD40" box="[496,573,652,676]" pageId="49" pageNumber="50" rank="genus">n. gen.</taxonomicNameLabel>
are often considered as belonging to
<emphasis id="B95670196458FF80FE1EEFBBD222FD27" box="[402,531,683,707]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C22D7886458FF80FE1EEFBBD23EFD27" authorityName="Hirschmann" authorityYear="1959" box="[402,527,683,707]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">Cornudina</taxonomicName>
.
</emphasis>
Yet, as explained by
<bibRefCitation id="EFB3D1FA6458FF80FF1BEFDBD178FD07" author="Orchard" box="[151,329,715,739]" firstAuthor="Orchard" pageId="49" pageNumber="50" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
, the
<typeStatus id="549912A96458FF80FE0BEFDBD1DBFD07" box="[391,490,715,739]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C22D7886458FF80FD98EFDBD2BFFD07" authorityName="Hirschmann" authorityYear="1959" box="[532,654,715,739]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD98EFDBD2BFFD07" box="[532,654,715,739]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
(
<taxonomicName id="4C22D7886458FF80FD2DEFDCD0C5FCE7" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="breviramulus">
<emphasis id="B95670196458FF80FD2DEFDCD0C5FCE7" italics="true" pageId="49" pageNumber="50">O. breviramulus</emphasis>
</taxonomicName>
) appears to be a P
<subScript id="17A6AE4E6458FF80FE5EEFE7D1EAFCEC" attach="left" box="[466,475,759,776]" fontSize="7" pageId="49" pageNumber="50">2</subScript>
element. Both
<bibRefCitation id="EFB3D1FA6458FF80FD03EFFCD109FCC7" author="Kozur and Mostler" firstAuthor="Kozur" pageId="49" pageNumber="50" pagination="1 - 19" refId="ref43462" refString="Kozur, H., &amp; Mostler, H. (1971). Probleme der Conodonten-forschung in der Trias. Geologische-Palaontologische Mitteilungen Innsbruck, 1 (4), 1 - 19." type="journal article" year="1971">Kozur and Mostler (1971</bibRefCitation>
, p. 11) and Sweet (in
<bibRefCitation id="EFB3D1FA6458FF80FDB7EE1BD330FCC7" author="Clark" box="[571,769,779,803]" etAl="et al." firstAuthor="Clark" pageId="49" pageNumber="50" refId="ref39882" refString="Clark, D. L., Sweet, W. C., Bergstrom, S. M., et al. (1981). Conodonta. Part W, Miscellanea, Supplement 2. In R. A. Robison (Ed.), Treatise on invertebrate paleontology. Boulder: University of Kansas Geological Society of America." type="book" year="1981">Clark et al., 1981</bibRefCitation>
, p. W155) placed
<taxonomicName id="4C22D7886458FF80FEDBEE3BD1E0FCA7" authorityName="Hirschmann" authorityYear="1959" box="[343,465,811,835]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FEDBEE3BD1E0FCA7" box="[343,465,811,835]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
with
<taxonomicName id="4C22D7886458FF80FD9DEE3BD2B9FCA7" authorityName="Hirschmann" authorityYear="1959" box="[529,648,811,835]" genus="Chirodella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD9DEE3BD2B9FCA7" box="[529,648,811,835]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
</taxonomicName>
as a multi-element
<taxonomicName id="4C22D7886458FF80FF75EE5BD141FC87" authorityName="Hirschmann" authorityYear="1959" box="[249,368,843,867]" genus="Chirodella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FF75EE5BD141FC87" box="[249,368,843,867]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
</taxonomicName>
. Te
<typeStatus id="549912A96458FF80FE3DEE5BD225FC87" box="[433,532,843,867]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C22D7886458FF80FDB7EE5BD283FC87" authorityName="Hirschmann" authorityYear="1959" box="[571,690,843,867]" genus="Chirodella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FDB7EE5BD283FC87" box="[571,690,843,867]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
</taxonomicName>
(
<emphasis id="B95670196458FF80FD4FEE5CD1B3FC67" italics="true" pageId="49" pageNumber="50">Metalonchodina triquetra</emphasis>
) is an S
<subScript id="17A6AE4E6458FF80FE6AEE67D1DEFC6C" attach="left" box="[486,495,887,904]" fontSize="7" pageId="49" pageNumber="50">2</subScript>
element. Te
<typeStatus id="549912A96458FF80FD16EE7BD336FC67" box="[666,775,875,899]" pageId="49" pageNumber="50" type="holotype">holotypes</typeStatus>
of both
<taxonomicName id="4C22D7886458FF80FF7AEE9BD141FC47" authorityName="Hirschmann" authorityYear="1959" box="[246,368,907,931]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FF7AEE9BD141FC47" box="[246,368,907,931]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886458FF80FE22EE9BD214FC47" authorityName="Hirschmann" authorityYear="1959" box="[430,549,907,931]" genus="Chirodella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE22EE9BD214FC47" box="[430,549,907,931]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
</taxonomicName>
are from Muschelkalk (Middle Triassic) collections made by
<bibRefCitation id="EFB3D1FA6458FF80FDF9EEBCD333FC27" author="Tatge" box="[629,770,939,964]" firstAuthor="Tatge" pageId="49" pageNumber="50" pagination="129 - 147" refId="ref47382" refString="Tatge, U. (1956). Conodonten aus dem germanischen Muschelkalk. Palaontologische Zeitschrift, 30 (3 - 4), 129 - 147. https: // doi. org / 10.1007 / BF 03041777" type="journal article" year="1956">Tatge (1956)</bibRefCitation>
, who described no elements like those described by
<bibRefCitation id="EFB3D1FA6458FF80FF1BEEFBD17AFBE7" author="Orchard" box="[151,331,1003,1027]" firstAuthor="Orchard" pageId="49" pageNumber="50" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
as P
<subScript id="17A6AE4E6458FF80FE07EEE7D1A5FBEC" attach="left" box="[395,404,1015,1032]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
.
<emphasis id="B95670196458FF80FE21EEFBD219FBE7" box="[429,552,1003,1027]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
sensu formo does not occur in Orchards nor in Koikes Spathian collections of
<taxonomicName id="4C22D7886458FF80FF75E93BD142FBA7" authorityName="Hirschmann" authorityYear="1959" box="[249,371,1067,1091]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FF75E93BD142FBA7" box="[249,371,1067,1091]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
. For this reason,
<bibRefCitation id="EFB3D1FA6458FF80FDCDE93BD2E5FBA7" author="Koike" box="[577,724,1067,1091]" firstAuthor="Koike" pageId="49" pageNumber="50" pagination="113 - 120" refId="ref42835" refString="Koike, T. (1996). Skeletal apparatuses of Triassic conodonts of Cornudia. Prof. Hisayoshi Igo Commemorative Volume on Geology and Paleontology of Japan and Southeast Asia, 113 - 120." type="book chapter" year="1996">Koike (1996)</bibRefCitation>
and
<bibRefCitation id="EFB3D1FA6458FF80FF1BE95BD179FB87" author="Orchard" box="[151,328,1099,1123]" firstAuthor="Orchard" pageId="49" pageNumber="50" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
regarded the two genera as unrelated: Koike reconstructed
<taxonomicName id="4C22D7886458FF80FE00E97BD237FB67" authorityName="Hirschmann" authorityYear="1959" box="[396,518,1131,1155]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE00E97BD237FB67" box="[396,518,1131,1155]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
as an apparatus consisting of P
<subScript id="17A6AE4E6458FF80FE9AE987D12EFB4C" attach="left" box="[278,287,1175,1192]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
and P
<subScript id="17A6AE4E6458FF80FEE6E987D142FB4C" attach="left" box="[362,371,1175,1192]" fontSize="7" pageId="49" pageNumber="50">2</subScript>
elements only (
<taxonomicName id="4C22D7886458FF80FDA7E99BD0E4FB27" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="breviramulis">
<emphasis id="B95670196458FF80FDA7E99BD0E4FB27" italics="true" pageId="49" pageNumber="50">Cornudina breviramulis</emphasis>
</taxonomicName>
) or of P
<subScript id="17A6AE4E6458FF80FEBDE9A7D10BFB2C" attach="left" box="[305,314,1207,1224]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
elements only (
<taxonomicName id="4C22D7886458FF80FE61E9BBD2A4FB27" box="[493,661,1195,1219]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="igoi">
<emphasis id="B95670196458FF80FE61E9BBD2A4FB27" box="[493,661,1195,1219]" italics="true" pageId="49" pageNumber="50">Cornudina igoi</emphasis>
</taxonomicName>
), whereas Orchard reconstructed
<taxonomicName id="4C22D7886458FF80FE29E9DBD22EFB07" authorityName="Hirschmann" authorityYear="1959" box="[421,543,1227,1251]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE29E9DBD22EFB07" box="[421,543,1227,1251]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
? as having an octomembrate apparatus whose P
<subScript id="17A6AE4E6458FF80FE64E9E7D1C0FAEC" attach="left" box="[488,497,1271,1288]" fontSize="7" pageId="49" pageNumber="50">2</subScript>
element vaguely resembles the
<typeStatus id="549912A96458FF80FF75E81BD16DFAC7" box="[249,348,1291,1315]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C22D7886458FF80FE0FE81BD1CCFAC7" authorityName="Hirschmann" authorityYear="1959" box="[387,509,1291,1315]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE0FE81BD1CCFAC7" box="[387,509,1291,1315]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
, but he questioned the very validity of that name for Spathian forms that might be unrelated to the true Middle Triassic
<taxonomicName id="4C22D7886458FF80FDEBE85BD2D0FA87" authorityName="Hirschmann" authorityYear="1959" box="[615,737,1355,1379]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FDEBE85BD2D0FA87" box="[615,737,1355,1379]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B6458FF80FF1BE87BD1A6F9C7" blockId="49.[151,775,459,1923]" pageId="49" pageNumber="50">
<emphasis id="B95670196458FF80FF11E87BD129FA67" box="[157,280,1387,1411]" italics="true" pageId="49" pageNumber="50">Chirodella</emphasis>
sensu formo does not occur in our Smithian collections of
<taxonomicName id="4C22D7886458FF80FEFDE89BD1E3FA47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[369,466,1419,1443]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B95670196458FF80FEFDE89BD1E3FA47" box="[369,466,1419,1443]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626458FF80FE6CE89CD200FA40" box="[480,561,1420,1444]" pageId="49" pageNumber="50" rank="genus">n. gen.</taxonomicNameLabel>
and neither does
<emphasis id="B95670196458FF80FF11E8BBD12AFA27" box="[157,283,1451,1475]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
sensu formo. Hence we suggest that
<taxonomicName id="4C22D7886458FF80FD3AE8BCD0FBFA07" authorityName="Hirschmann" authorityYear="1959" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD3AE8BCD0FBFA07" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
-likeP
<subScript id="17A6AE4E6458FF80FE9EE8C7D12AFA0C" attach="left" box="[274,283,1495,1512]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
elements found in the Smithian, and possibly those found in the Spathian, belong to
<taxonomicName id="4C22D7886458FF80FD0BE8FBD2D9F9E7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[647,744,1515,1539]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B95670196458FF80FD0BE8FBD2D9F9E7" box="[647,744,1515,1539]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626458FF80FD7EE8FCD0F7F9C0" pageId="49" pageNumber="50" rank="genus">n. gen.</taxonomicNameLabel>
not to
<emphasis id="B95670196458FF80FE9AEB1BD1A6F9C7" box="[278,407,1547,1571]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C22D7886458FF80FE9AEB1BD1A2F9C7" authorityName="Hirschmann" authorityYear="1959" box="[278,403,1547,1571]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">Cornudina</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B9DAC0B6458FF80FF21EB3BD4E5FE47" blockId="49.[151,775,459,1923]" lastBlockId="49.[813,1436,235,419]" pageId="49" pageNumber="50">
Based on multi-element considerations, Orchard grouped both
<emphasis id="B95670196458FF80FEDDEB5BD1FEF987" box="[337,463,1611,1635]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
and
<taxonomicName id="4C22D7886458FF80FD96EB5BD29CF987" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[538,685,1611,1635]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD96EB5BD29CF987" box="[538,685,1611,1635]" italics="true" pageId="49" pageNumber="50">Spathicuspus</emphasis>
</taxonomicName>
within the same subfamily
<taxonomicName id="4C22D7886458FF80FEF3EB7BD22CF967" authorityName="Orchard" authorityYear="2005" box="[383,541,1643,1667]" pageId="49" pageNumber="50" rank="subFamily" subFamily="Cornudininae">Cornudininae</taxonomicName>
. Because
<taxonomicName id="4C22D7886458FF80FD1DEB7BD08FF947" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FD1DEB7BD08FF947" italics="true" pageId="49" pageNumber="50">Spathicuspus</emphasis>
</taxonomicName>
seemed to occur first, he hypothesized that
<emphasis id="B95670196458FF80FD58EB9CD0DAF927" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
evolved from
<taxonomicName id="4C22D7886458FF80FE2FEBBBD207F927" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[419,566,1707,1731]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE2FEBBBD207F927" box="[419,566,1707,1731]" italics="true" pageId="49" pageNumber="50">Spathicuspus</emphasis>
</taxonomicName>
near the Lower/ Middle Triassic boundary through overall shortening of the P
<subScript id="17A6AE4E6458FF80FEA4EBE7D100F8EC" attach="left" box="[296,305,1783,1800]" fontSize="7" pageId="49" pageNumber="50">1</subScript>
element. Since
<taxonomicName id="4C22D7886458FF80FE7EEBFBD262F8E7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[498,595,1771,1795]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FE7EEBFBD262F8E7" box="[498,595,1771,1795]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
occurs already in the Smithian, we suggest that
<taxonomicName id="4C22D7886458FF80FDBAEA1BD2F8F8C7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[566,713,1803,1827]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FDBAEA1BD2F8F8C7" box="[566,713,1803,1827]" italics="true" pageId="49" pageNumber="50">Spathicuspus</emphasis>
</taxonomicName>
may have evolved from
<taxonomicName id="4C22D7886458FF80FEFEEA3BD1E2F8A7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[370,467,1835,1859]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FEFEEA3BD1E2F8A7" box="[370,467,1835,1859]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
and the Spathian forms of
<emphasis id="B95670196458FF80FF11EA5BD12AF887" box="[157,283,1867,1891]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
either belong to
<emphasis id="B95670196458FF80FE6FEA5BD27AF887" box="[483,587,1867,1891]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C22D7886458FF80FE6FEA5BD276F887" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[483,583,1867,1891]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">Urdyella</taxonomicName>
,
</emphasis>
derived directly from
<emphasis id="B95670196458FF80FF59EA7BD10DF867" box="[213,316,1899,1923]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C22D7886458FF80FF59EA7BD108F867" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[213,313,1899,1923]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">Urdyella</taxonomicName>
,
</emphasis>
or as assumed by Orchard, evolved from
<taxonomicName id="4C22D7886458FF80FCA1EDFBD3F1FEE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[813,960,235,259]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FCA1EDFBD3F1FEE7" box="[813,960,235,259]" italics="true" pageId="49" pageNumber="50">Spathicuspus</emphasis>
</taxonomicName>
, although the latter hypothesis seems less likely.
<taxonomicName id="4C22D7886458FF80FCF8EC1BD3E4FEC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[884,981,267,291]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FCF8EC1BD3E4FEC7" box="[884,981,267,291]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
itself may have evolved from
<taxonomicName id="4C22D7886458FF80FAA8EC1BD5ADFEC7" authorityName="Orchard" authorityYear="2005" box="[1316,1436,267,291]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FAA8EC1BD5ADFEC7" box="[1316,1436,267,291]" italics="true" pageId="49" pageNumber="50">Discretella</emphasis>
</taxonomicName>
via forms like
<taxonomicName id="4C22D7886458FF80FC57EC3BD462FEA7" authorityName="Orchard" authorityYear="2005" box="[987,1107,299,323]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FC57EC3BD462FEA7" box="[987,1107,299,323]" italics="true" pageId="49" pageNumber="50">Discretella</emphasis>
</taxonomicName>
? n. sp. B.
<taxonomicName id="4C22D7886458FF80FB56EC3BD563FEA7" authorityName="Orchard" authorityYear="2005" box="[1242,1362,299,323]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FB56EC3BD563FEA7" box="[1242,1362,299,323]" italics="true" pageId="49" pageNumber="50">Discretella</emphasis>
</taxonomicName>
has a much shorter cusp and a larger anterior process. Furthermore, the basal cavity in
<taxonomicName id="4C22D7886458FF80FBFBEC7BD4E9FE67" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1143,1240,363,387]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FBFBEC7BD4E9FE67" box="[1143,1240,363,387]" italics="true" pageId="49" pageNumber="50">Urdyella</emphasis>
</taxonomicName>
is flatter and not as broadly excavated as in
<taxonomicName id="4C22D7886458FF80FBD8EC9BD4FFFE47" authorityName="Hirschmann" authorityYear="1959" box="[1108,1230,395,419]" genus="Cornudina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B95670196458FF80FBD8EC9BD4FFFE47" box="[1108,1230,395,419]" italics="true" pageId="49" pageNumber="50">Cornudina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,168 +0,0 @@
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<subSubSection id="C338FF806459FF81FF1BE93BD14EFBA7" box="[151,383,1067,1091]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8B9DAC0B6459FF81FF1BE93BD14EFBA7" blockId="48.[151,383,1067,1123]" box="[151,383,1067,1091]" pageId="48" pageNumber="49">
<heading id="D0D51B676459FF81FF1BE93BD14EFBA7" box="[151,383,1067,1091]" fontSize="9" level="2" pageId="48" pageNumber="49" reason="8">
<taxonomicName id="4C22D7886459FF81FF1BE93BD117FBA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,294,1067,1091]" family="Gondolellidae" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="48" pageNumber="49" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196459FF81FF1BE93BD117FBA7" box="[151,294,1067,1091]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626459FF81FEA1E93CD157FBA7" box="[301,358,1068,1091]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
A
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<subSubSection id="C338FF806459FF81FF1BE95BD2E5F887" pageId="48" pageNumber="49" type="description">
<paragraph id="8B9DAC0B6459FF81FF1BE95BD122FB87" blockId="48.[151,383,1067,1123]" box="[151,275,1099,1123]" pageId="48" pageNumber="49">
<figureCitation id="1319B08E6459FF81FF1BE95BD122FB87" box="[151,275,1099,1123]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="48" pageNumber="49">Fig. 13M, P</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE99BD137FB27" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2005
<taxonomicName id="4C22D7886459FF81FF59E99BD239FB47" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[213,520,1163,1187]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF59E99BD18AFB47" box="[213,443,1163,1187]" italics="true" pageId="48" pageNumber="49">Neospathodus spathi</emphasis>
Sweet
</taxonomicName>
; Gaetani et al., p. 288, pl. 1, fig. 2
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE9DBD101FAE7" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2015
<taxonomicName id="4C22D7886459FF81FF54E9DBD227FB07" authority="(Sweet)" baseAuthorityName="Sweet" box="[216,534,1227,1251]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF54E9DBD186FB07" box="[216,439,1227,1251]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., p. 112, fig. 8.168.17.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE81BD117FAA7" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2015
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<emphasis id="B95670196459FF81FF54E81BD186FAC7" box="[216,439,1291,1315]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Lehrmann et al., p. 123, fig. 5.28.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE85BD0C2FA67" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2016
<taxonomicName id="4C22D7886459FF81FF54E85BD224FA87" authority="(Sweet)" baseAuthorityName="Sweet" box="[216,533,1355,1379]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF54E85BD187FA87" box="[216,438,1355,1379]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Liang et al., p. 385, fig. 4.10.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE8BBD18CFA27" blockId="48.[151,776,1451,1891]" box="[151,445,1451,1475]" pageId="48" pageNumber="49">Number of specimens:&gt;10</paragraph>
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<emphasis id="B95670196459FF81FF1BE8DCD12AFA07" box="[151,283,1484,1507]" italics="true" pageId="48" pageNumber="49">Description.</emphasis>
Short segminate P1 element with large, terminal, reclined or recurved cusp. Only two anterior denticles. Basal cavity is drop-shaped.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF21EB3BD196F907" blockId="48.[151,776,1451,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF21EB3BD120F9A7" box="[173,273,1579,1603]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Very similar to
<taxonomicName id="4C22D7886459FF81FE5EEB3BD250F9A7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[466,609,1579,1603]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FE5EEB3BD250F9A7" box="[466,609,1579,1603]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
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? n. sp. B (see below), but much shorter and with a larger basal cavity. Similar elements have been illustrated from other Chinese sections by
<bibRefCitation id="EFB3D1FA6459FF81FEC1EB9BD23EF947" author="Chen" box="[333,527,1675,1699]" etAl="et al." firstAuthor="Chen" pageId="48" pageNumber="49" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
and
<bibRefCitation id="EFB3D1FA6459FF81FDC8EB9BD336F947" author="Liang" box="[580,775,1675,1699]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al. (2016)</bibRefCitation>
and hence this species may be useful as an index fossil for intrabasinal correlations.
</paragraph>
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<emphasis id="B95670196459FF81FF21EBFCD119F8E7" box="[173,296,1772,1795]" italics="true" pageId="48" pageNumber="49">Occurrence</emphasis>
. South
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: Luolou Formation, Spathian, Nanpanjiang basin, Mingtang section, (
<bibRefCitation id="EFB3D1FA6459FF81FDC8EA1BD2CFF8C7" author="Liang" box="[580,766,1803,1827]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al., 2016</bibRefCitation>
) Upper Guandao section, (
<bibRefCitation id="EFB3D1FA6459FF81FE3BEA3BD29DF8A7" author="Lehrmann" box="[439,684,1835,1859]" etAl="et al." firstAuthor="Lehrmann" pageId="48" pageNumber="49" pagination="117 - 135" refId="ref43576" refString="Lehrmann, D. J., Stepchinski, L., Altiner, D., Orchard, M. J., Montgomery, P., Enos, P., Ellwood, B. B., Bowring, S. A., Ramezani, J., Wang, H., Wei, J., Yu, M., Griffiths, J. D., Minzoni, M., Schaal, E. K., Li, X., Meyer, K. M., &amp; Payne, J. (2015). An integrated biostratigraphy (conodonts and foraminifers) and chronostratigraphy (paleomagnetic reversals, magnetic susceptibility, elemental chemistry, carbon isotopes and geochronology) for the Permian-Upper Triassic strata of Guandao section, Nanpanjiang Basin, south China. Journal of Asian Earth Sciences, 108, 117 - 135. https: // doi. org / 10.1016 / j. jseaes. 2015.04. 030" type="journal article" year="2015">Lehrmann et al., 2015</bibRefCitation>
),
<taxonomicName id="4C22D7886459FF81FD48EA3CD15EF887" baseAuthorityName="Bender" baseAuthorityYear="1970" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196459FF81FD48EA3CD15EF887" italics="true" pageId="48" pageNumber="49">Triassospathodus homeri</emphasis>
</taxonomicName>
zone, Jiarong (
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).
</paragraph>
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Genus
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SPATHICUSPUS
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species.
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<taxonomicName id="4C22D7886459FF81FEA6EE1BD0F1FCA7" authority="Sweet, 1970, pp. 257 - 258" authorityName="Sweet" authorityPageNumber="257 - 258" authorityYear="1970" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FEA6EE1BD23FFCC7" box="[298,526,779,803]" italics="true" pageId="48" pageNumber="49">Neospathodus spathi</emphasis>
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, pp. 257 258
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, pl. 1, fig. 5.
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<typeStatus id="549912A96459FF81FF1BEE5CD0FAFC87" box="[151,203,844,867]" pageId="48" pageNumber="49">Type</typeStatus>
stratum and locality.
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,
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.
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<emphasis id="B95670196459FF81FF1BEE9BD0CDFC47" box="[151,252,907,931]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Based on his reconstruction of the multi-element apparatus of Neospathodus
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spathi
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considered these taxa deserved assignment not only to a new genus, but also to a new subfamily.
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<mods:title id="A6608A60B073A72E6E8B29D3E00C461B">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="5D97B6B8DB40BC6B16C85E829D60686E">Leu, Marc</mods:namePart>
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<paragraph id="8B9DAC0B6459FF81FF1BEFDBD20CFD07" blockId="48.[151,628,683,739]" box="[151,573,715,739]" pageId="48" pageNumber="49">
<heading id="D0D51B676459FF81FF1BEFDBD20CFD07" box="[151,573,715,739]" fontSize="36" level="2" pageId="48" pageNumber="49" reason="3">
Genus
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SPATHICUSPUS
<bibRefCitation id="EFB3D1FA6459FF81FE2BEFDBD20CFD07" author="Orchard" box="[423,573,715,739]" firstAuthor="Orchard" pageId="48" pageNumber="49" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard 2005</bibRefCitation>
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<subSubSection id="C338FF806459FF81FF1BEE1CD275FBE7" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8B9DAC0B6459FF81FF1BEE1CD171FCA7" blockId="48.[151,775,779,1028]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF1BEE1CD112FCC7" box="[151,291,780,803]" italics="true" pageId="48" pageNumber="49">
<typeStatus id="549912A96459FF81FF1BEE1CD0FAFCC7" box="[151,203,780,803]" pageId="48" pageNumber="49">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C22D7886459FF81FEA6EE1BD0F1FCA7" authority="Sweet, 1970, pp. 257 - 258" authorityName="Sweet" authorityPageNumber="257 - 258" authorityYear="1970" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FEA6EE1BD23FFCC7" box="[298,526,779,803]" italics="true" pageId="48" pageNumber="49">Neospathodus spathi</emphasis>
<bibRefCitation id="EFB3D1FA6459FF81FD99EE1BD2ABFCC7" author="Sweet" box="[533,666,779,803]" firstAuthor="Sweet" pageId="48" pageNumber="49" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
, pp. 257 258
</taxonomicName>
, pl. 1, fig. 5.
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<materialsCitation id="3B4AA6566459FF81FF1BEE5CD1E5FC67" country="Pakistan" county="Narmia" location="Mittiwali Member" municipality="Mianwali Formation" pageId="48" pageNumber="49" specimenCount="1" typeStatus="holotype">
<emphasis id="B95670196459FF81FF1BEE5CD18FFC87" box="[151,446,843,867]" italics="true" pageId="48" pageNumber="49">
<typeStatus id="549912A96459FF81FF1BEE5CD0FAFC87" box="[151,203,844,867]" pageId="48" pageNumber="49">Type</typeStatus>
stratum and locality.
</emphasis>
<location id="8EFDFAD06459FF81FE4BEE5BD2A2FC87" LSID="urn:lsid:plazi:treatment:038B1D1D6459FF81FF1BEFDBD5A6F887:8EFDFAD06459FF81FE4BEE5BD2A2FC87" box="[455,659,843,867]" country="Pakistan" county="Narmia" municipality="Mianwali Formation" name="Mittiwali Member" pageId="48" pageNumber="49">Mittiwali Member</location>
,
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,
<collectingCounty id="62FCD4876459FF81FE9AEE7BD15DFC67" box="[278,364,875,899]" pageId="48" pageNumber="49">Narmia</collectingCounty>
,
<collectingCountry id="F335EC9B6459FF81FEF9EE7BD1E5FC67" box="[373,468,875,899]" name="Pakistan" pageId="48" pageNumber="49">Pakistan</collectingCountry>
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.
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<emphasis id="B95670196459FF81FF1BEE9BD0CDFC47" box="[151,252,907,931]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Based on his reconstruction of the multi-element apparatus of Neospathodus
<taxonomicName id="4C22D7886459FF81FD99EEBBD336FC27" authority="Orchard (2005)" authorityName="Orchard" authorityYear="2005" box="[533,775,939,963]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
spathi
<bibRefCitation id="EFB3D1FA6459FF81FDD2EEBBD336FC27" author="Orchard" box="[606,775,939,963]" firstAuthor="Orchard" pageId="48" pageNumber="49" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
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considered these taxa deserved assignment not only to a new genus, but also to a new subfamily.
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<heading id="D0D51B676459FF81FF1BE93BD14EFBA7" box="[151,383,1067,1091]" fontSize="9" level="2" pageId="48" pageNumber="49" reason="8">
<taxonomicName id="4C22D7886459FF81FF1BE93BD117FBA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,294,1067,1091]" family="Gondolellidae" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="48" pageNumber="49" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196459FF81FF1BE93BD117FBA7" box="[151,294,1067,1091]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626459FF81FEA1E93CD157FBA7" box="[301,358,1068,1091]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
A
</heading>
</paragraph>
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<subSubSection id="C338FF806459FF81FF1BE95BD2E5F887" pageId="48" pageNumber="49" type="description">
<paragraph id="8B9DAC0B6459FF81FF1BE95BD122FB87" blockId="48.[151,383,1067,1123]" box="[151,275,1099,1123]" pageId="48" pageNumber="49">
<figureCitation id="1319B08E6459FF81FF1BE95BD122FB87" box="[151,275,1099,1123]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="48" pageNumber="49">Fig. 13M, P</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE99BD137FB27" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2005
<taxonomicName id="4C22D7886459FF81FF59E99BD239FB47" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[213,520,1163,1187]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF59E99BD18AFB47" box="[213,443,1163,1187]" italics="true" pageId="48" pageNumber="49">Neospathodus spathi</emphasis>
Sweet
</taxonomicName>
; Gaetani et al., p. 288, pl. 1, fig. 2
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE9DBD101FAE7" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2015
<taxonomicName id="4C22D7886459FF81FF54E9DBD227FB07" authority="(Sweet)" baseAuthorityName="Sweet" box="[216,534,1227,1251]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF54E9DBD186FB07" box="[216,439,1227,1251]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., p. 112, fig. 8.168.17.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE81BD117FAA7" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2015
<taxonomicName id="4C22D7886459FF81FF54E81BD226FAC7" authority="(Sweet)" baseAuthorityName="Sweet" box="[216,535,1291,1315]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF54E81BD186FAC7" box="[216,439,1291,1315]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Lehrmann et al., p. 123, fig. 5.28.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE85BD0C2FA67" blockId="48.[151,775,1163,1411]" pageId="48" pageNumber="49">
2016
<taxonomicName id="4C22D7886459FF81FF54E85BD224FA87" authority="(Sweet)" baseAuthorityName="Sweet" box="[216,533,1355,1379]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FF54E85BD187FA87" box="[216,438,1355,1379]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Liang et al., p. 385, fig. 4.10.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE8BBD18CFA27" blockId="48.[151,776,1451,1891]" box="[151,445,1451,1475]" pageId="48" pageNumber="49">Number of specimens:&gt;10</paragraph>
<paragraph id="8B9DAC0B6459FF81FF1BE8DCD23BF9C7" blockId="48.[151,776,1451,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF1BE8DCD12AFA07" box="[151,283,1484,1507]" italics="true" pageId="48" pageNumber="49">Description.</emphasis>
Short segminate P1 element with large, terminal, reclined or recurved cusp. Only two anterior denticles. Basal cavity is drop-shaped.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF21EB3BD196F907" blockId="48.[151,776,1451,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF21EB3BD120F9A7" box="[173,273,1579,1603]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Very similar to
<taxonomicName id="4C22D7886459FF81FE5EEB3BD250F9A7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[466,609,1579,1603]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FE5EEB3BD250F9A7" box="[466,609,1579,1603]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
? n. sp. B (see below), but much shorter and with a larger basal cavity. Similar elements have been illustrated from other Chinese sections by
<bibRefCitation id="EFB3D1FA6459FF81FEC1EB9BD23EF947" author="Chen" box="[333,527,1675,1699]" etAl="et al." firstAuthor="Chen" pageId="48" pageNumber="49" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
and
<bibRefCitation id="EFB3D1FA6459FF81FDC8EB9BD336F947" author="Liang" box="[580,775,1675,1699]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al. (2016)</bibRefCitation>
and hence this species may be useful as an index fossil for intrabasinal correlations.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FF21EBFCD2E5F887" blockId="48.[151,776,1451,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF21EBFCD119F8E7" box="[173,296,1772,1795]" italics="true" pageId="48" pageNumber="49">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B6459FF81FEF1EBFBD1F3F8E7" box="[381,450,1771,1795]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
: Luolou Formation, Spathian, Nanpanjiang basin, Mingtang section, (
<bibRefCitation id="EFB3D1FA6459FF81FDC8EA1BD2CFF8C7" author="Liang" box="[580,766,1803,1827]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al., 2016</bibRefCitation>
) Upper Guandao section, (
<bibRefCitation id="EFB3D1FA6459FF81FE3BEA3BD29DF8A7" author="Lehrmann" box="[439,684,1835,1859]" etAl="et al." firstAuthor="Lehrmann" pageId="48" pageNumber="49" pagination="117 - 135" refId="ref43576" refString="Lehrmann, D. J., Stepchinski, L., Altiner, D., Orchard, M. J., Montgomery, P., Enos, P., Ellwood, B. B., Bowring, S. A., Ramezani, J., Wang, H., Wei, J., Yu, M., Griffiths, J. D., Minzoni, M., Schaal, E. K., Li, X., Meyer, K. M., &amp; Payne, J. (2015). An integrated biostratigraphy (conodonts and foraminifers) and chronostratigraphy (paleomagnetic reversals, magnetic susceptibility, elemental chemistry, carbon isotopes and geochronology) for the Permian-Upper Triassic strata of Guandao section, Nanpanjiang Basin, south China. Journal of Asian Earth Sciences, 108, 117 - 135. https: // doi. org / 10.1016 / j. jseaes. 2015.04. 030" type="journal article" year="2015">Lehrmann et al., 2015</bibRefCitation>
),
<taxonomicName id="4C22D7886459FF81FD48EA3CD15EF887" baseAuthorityName="Bender" baseAuthorityYear="1970" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196459FF81FD48EA3CD15EF887" italics="true" pageId="48" pageNumber="49">Triassospathodus homeri</emphasis>
</taxonomicName>
zone, Jiarong (
<bibRefCitation id="EFB3D1FA6459FF81FD9CEA5BD2F7F887" author="Chen" box="[528,710,1867,1891]" etAl="et al." firstAuthor="Chen" pageId="48" pageNumber="49" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338FF806459FF81FCA1EDFBD42AFEE7" box="[813,1051,235,259]" pageId="48" pageNumber="49" type="multiple">
<paragraph id="8B9DAC0B6459FF81FCA1EDFBD42AFEE7" blockId="48.[813,1199,235,291]" box="[813,1051,235,259]" pageId="48" pageNumber="49">
<heading id="D0D51B676459FF81FCA1EDFBD42AFEE7" box="[813,1051,235,259]" fontSize="9" level="2" pageId="48" pageNumber="49" reason="8">
<taxonomicName id="4C22D7886459FF81FCA1EDFBD38DFEE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[813,956,235,259]" family="Gondolellidae" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FCA1EDFBD38DFEE7" box="[813,956,235,259]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
? n. sp. B
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806459FF81FCA1EC1BD365FB07" pageId="48" pageNumber="49" type="description">
<paragraph id="8B9DAC0B6459FF81FCA1EC1BD49EFEC7" blockId="48.[813,1199,235,291]" box="[813,1199,267,291]" pageId="48" pageNumber="49">
<figureCitation id="1319B08E6459FF81FCA1EC1BD38CFEC7" box="[813,957,267,291]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="48" pageNumber="49">Fig. 13JL, O</figureCitation>
(N, R juvenile forms?)
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1EC5BD557FE87" blockId="48.[813,1437,331,547]" box="[813,1382,331,355]" pageId="48" pageNumber="49">
2005
<taxonomicName id="4C22D7886459FF81FCEBEC5BD404FE87" box="[871,1077,331,355]" genus="Spathicuspus" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B95670196459FF81FCEBEC5BD3C7FE87" box="[871,1014,331,355]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
sp. A
</taxonomicName>
; Orchard, p.77, fig. 2, Nr. A.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1EC7BD34AFE47" blockId="48.[813,1437,331,547]" pageId="48" pageNumber="49">
2016
<taxonomicName id="4C22D7886459FF81FCE1EC7BD49BFE67" authority="(Sweet)" baseAuthorityName="Sweet" box="[877,1194,363,387]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCE1EC7BD47AFE67" box="[877,1099,363,387]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Liang et al., p. 385, fig. 4.9.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1ECBBD36CFE07" blockId="48.[813,1437,331,547]" pageId="48" pageNumber="49">
2019
<taxonomicName id="4C22D7886459FF81FCE6ECBBD493FE27" authority="(Sweet)" baseAuthorityName="Sweet" box="[874,1186,427,451]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCE6ECBBD477FE27" box="[874,1094,427,451]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., fig. 7, nr. 68.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1ECFBD342FDC7" blockId="48.[813,1437,331,547]" pageId="48" pageNumber="49">
2021
<taxonomicName id="4C22D7886459FF81FCE1ECFBD49AFDE7" authority="(Sweet)" baseAuthorityName="Sweet" box="[877,1195,491,515]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCE1ECFBD47DFDE7" box="[877,1100,491,515]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., fig. 7.8 (only).
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1EF5BD460FD87" blockId="48.[813,1437,587,1251]" box="[813,1105,587,611]" pageId="48" pageNumber="49">Number of specimens:&gt;30</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1EF7CD537FD07" blockId="48.[813,1437,587,1251]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCA1EF7CD381FD67" box="[813,944,620,643]" italics="true" pageId="48" pageNumber="49">Description.</emphasis>
Slender segminate P
<subScript id="17A6AE4E6459FF81FB15EF67D493FD6C" attach="left" box="[1177,1186,631,648]" fontSize="7" pageId="48" pageNumber="49">1</subScript>
element with a terminal, reclined or recurved cusp that is usually broader than adjacent denticles, a shallow, posteriorly rounded basal cavity and three to seven anterior denticles.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCCEEFFBD5A3FBA7" blockId="48.[813,1437,587,1251]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCCEEFFBD396FCE7" box="[834,935,747,771]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Despite their overall resemblance with
<taxonomicName id="4C22D7886459FF81FCA1EE1BD43BFCC7" box="[813,1034,779,803]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCA1EE1BD43BFCC7" box="[813,1034,779,803]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
</taxonomicName>
these elements are either too long and bear too many denticles or their cusp is not broad and blunt enough to be assigned to that species. Some of them may not belong to
<taxonomicName id="4C22D7886459FF81FBE0EE7BD4CAFC67" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1132,1275,875,899]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FBE0EE7BD4CAFC67" box="[1132,1275,875,899]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
at all. Similar and apparently coeval elements were illustrated by
<bibRefCitation id="EFB3D1FA6459FF81FAD3EE9BD382FC27" author="Liang" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al. (2016)</bibRefCitation>
and
<bibRefCitation id="EFB3D1FA6459FF81FC62EEBBD489FC27" author="Chen" box="[1006,1208,939,963]" etAl="et al." firstAuthor="Chen" pageId="48" pageNumber="49" refId="ref39612" refString="Chen, Y., Richoz, S., Krystyn, L., &amp; Zhang, Z. (2019). Quantitative stratigraphic correlation of Tethyan conodonts across the Smithian-Spathian (Early Triassic) extinction event. Earth-Science Reviews. https: // doi. org / 10. 1016 / j. earscirev. 2019.03. 004" type="book" year="2019">Chen et al. (2019)</bibRefCitation>
from other sections in
<collectingCountry id="F335EC9B6459FF81FCC6EEDBD3BDFC07" box="[842,908,971,995]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
and
<collectingCountry id="F335EC9B6459FF81FC49EEDBD43CFC07" box="[965,1037,971,995]" name="Oman" pageId="48" pageNumber="49">Oman</collectingCountry>
. Te element illustrated by
<bibRefCitation id="EFB3D1FA6459FF81FACCEEDBD342FBE7" author="Orchard" firstAuthor="Orchard" pageId="48" pageNumber="49" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
has similar aspect ratio and denticulation as for instance the specimen we illustrated in
<figureCitation id="1319B08E6459FF81FB53E91BD50CFBC7" box="[1247,1341,1035,1059]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="48" pageNumber="49">Fig. 13O</figureCitation>
, but it is also much younger and may be only superficially similar.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1E95CD5B2FB27" blockId="48.[813,1437,587,1251]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCA1E95CD399FB87" box="[813,936,1100,1123]" italics="true" pageId="48" pageNumber="49">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B6459FF81FB89E95BD47BFB87" box="[1029,1098,1099,1123]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
: Luolou Formation, Spathian, Mingtang section, Nanpanjiang basin (
<bibRefCitation id="EFB3D1FA6459FF81FB5FE97BD5BCFB67" author="Liang" box="[1235,1421,1131,1155]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al., 2016</bibRefCitation>
), Upper Guandao section (
<bibRefCitation id="EFB3D1FA6459FF81FBC3E99BD4C0FB47" author="Orchard" box="[1103,1265,1163,1187]" firstAuthor="Orchard" pageId="48" pageNumber="49" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
).
<collectingCountry id="F335EC9B6459FF81FA80E99BD565FB47" box="[1292,1364,1163,1187]" name="Oman" pageId="48" pageNumber="49">Oman</collectingCountry>
: Wadi Bani Khalid section, Spathian UAZ
<subScript id="17A6AE4E6459FF81FB24E9A8D480FB2C" attach="left" box="[1192,1201,1208,1224]" fontSize="7" pageId="48" pageNumber="49">7</subScript>
(
<bibRefCitation id="EFB3D1FA6459FF81FB33E9BBD544FB27" author="Chen" box="[1215,1397,1195,1219]" etAl="et al." firstAuthor="Chen" pageId="48" pageNumber="49" refId="ref39612" refString="Chen, Y., Richoz, S., Krystyn, L., &amp; Zhang, Z. (2019). Quantitative stratigraphic correlation of Tethyan conodonts across the Smithian-Spathian (Early Triassic) extinction event. Earth-Science Reviews. https: // doi. org / 10. 1016 / j. earscirev. 2019.03. 004" type="book" year="2019">Chen et al., 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1E9DBD365FB07" blockId="48.[813,1437,587,1251]" box="[813,852,1227,1251]" pageId="48" pageNumber="49">F13</paragraph>
</subSubSection>
<subSubSection id="C338FF806459FF81FCA1E81BD42CFAC7" box="[813,1053,1291,1315]" pageId="48" pageNumber="49" type="multiple">
<paragraph id="8B9DAC0B6459FF81FCA1E81BD42CFAC7" blockId="48.[813,1053,1291,1347]" box="[813,1053,1291,1315]" pageId="48" pageNumber="49">
<heading id="D0D51B676459FF81FCA1E81BD42CFAC7" box="[813,1053,1291,1315]" fontSize="9" level="2" pageId="48" pageNumber="49" reason="8">
<taxonomicName id="4C22D7886459FF81FCA1E81BD38DFAC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[813,956,1291,1315]" family="Gondolellidae" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FCA1E81BD38DFAC7" box="[813,956,1291,1315]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
? n. sp. C
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806459FF81FCA1E83BD5A6F887" pageId="48" pageNumber="49" type="description">
<paragraph id="8B9DAC0B6459FF81FCA1E83BD3B8FAA7" blockId="48.[813,1053,1291,1347]" box="[813,905,1323,1347]" pageId="48" pageNumber="49">
<figureCitation id="1319B08E6459FF81FCA1E83BD3B8FAA7" box="[813,905,1323,1347]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="48" pageNumber="49">Fig. 13Q</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1E87BD3B9FA47" blockId="48.[813,1436,1387,1507]" pageId="48" pageNumber="49">
2016
<taxonomicName id="4C22D7886459FF81FCE1E87BD49BFA67" authority="(Sweet)" baseAuthorityName="Sweet" box="[877,1194,1387,1411]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCE1E87BD47AFA67" box="[877,1099,1387,1411]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Liang et al., p. 385, fig. 4.11.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1E8BBD342FA07" blockId="48.[813,1436,1387,1507]" pageId="48" pageNumber="49">
2021
<taxonomicName id="4C22D7886459FF81FCE1E8BBD49AFA27" authority="(Sweet)" baseAuthorityName="Sweet" box="[877,1195,1451,1475]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCE1E8BBD47DFA27" box="[877,1100,1451,1475]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
; Chen et al., fig. 7.5 (only).
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCA1EB1BD475F9C7" blockId="48.[813,1437,1547,1891]" box="[813,1092,1547,1571]" pageId="48" pageNumber="49">Number of specimens:&gt;5</paragraph>
<paragraph id="8B9DAC0B6459FF81FCCEEB3CD3B8F947" blockId="48.[813,1437,1547,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCCEEB3CD3F7F9A7" box="[834,966,1580,1603]" italics="true" pageId="48" pageNumber="49">Description.</emphasis>
Small P1 element with a rounded basal cavity, a large, reclined and broad cusp, a small posterior denticle and a couple of declining denticles to the anterior.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCCEEBBBD3C5F8C7" blockId="48.[813,1437,1547,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCCEEBBBD396F927" box="[834,935,1707,1731]" italics="true" pageId="48" pageNumber="49">Remarks.</emphasis>
Although the relationship of this species to
<taxonomicName id="4C22D7886459FF81FCA1EBDBD437F907" box="[813,1030,1739,1763]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B95670196459FF81FCA1EBDBD437F907" box="[813,1030,1739,1763]" italics="true" pageId="48" pageNumber="49">Spathicuspus spathi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886459FF81FBB1EBDBD4FDF907" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1085,1228,1739,1763]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196459FF81FBB1EBDBD4FDF907" box="[1085,1228,1739,1763]" italics="true" pageId="48" pageNumber="49">Spathicuspus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626459FF81FB58EBDCD53FF907" box="[1236,1294,1740,1763]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
A is unclear, its shape is intermediate. It differs in the presence of a posterior denticle.
</paragraph>
<paragraph id="8B9DAC0B6459FF81FCCEEA3CD5A6F887" blockId="48.[813,1437,1547,1891]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FCCEEA3CD38CF8A7" box="[834,957,1836,1859]" italics="true" pageId="48" pageNumber="49">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B6459FF81FB9EEA3BD466F8A7" box="[1042,1111,1835,1859]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
: Luolou Formation, Spathian, Mingtang section, Nanpanjiang basin (
<bibRefCitation id="EFB3D1FA6459FF81FB5CEA5BD5B8F887" author="Liang" box="[1232,1417,1867,1891]" etAl="et al." firstAuthor="Liang" pageId="48" pageNumber="49" pagination="381 - 393" refId="ref44082" refString="Liang, L., Tong, J., Song, H., Song, T., Tian, L., Song, H., &amp; Qiu, H. (2016). Lower- Middle Triassic conodont biostratigraphy of the Mingtang section, Nanpanjiang Basin, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 381 - 393. https: // doi. org / 10.1016 / j. palaeo. 2016.07. 027" type="journal article" year="2016">Liang et al., 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,283 @@
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<heading id="D0D51B67645DFF85FF1BEB7BD257F967" box="[151,614,1643,1667]" fontSize="36" level="2" pageId="52" pageNumber="53" reason="3">
Genus
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HADRODONTINA
<bibRefCitation id="EFB3D1FA645DFF85FE4BEB7BD257F967" author="Staesche" box="[455,614,1643,1667]" firstAuthor="Staesche" pageId="52" pageNumber="53" refId="ref46867" refString="Staesche, U. 1964. Conodonten aus dem Skyth von Siidtirol. Neues Jahrbuch fiir Geologie und Palaontologie." type="book" year="1964">Staesche, 1964</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645DFF85FF1BEBBCD4CAFAC7" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8B9DAC0B645DFF85FF1BEBBCD28DF927" blockId="52.[151,775,1707,1795]" box="[151,700,1707,1731]" pageId="52" pageNumber="53">
<emphasis id="B9567019645DFF85FF1BEBBCD110F927" box="[151,289,1708,1731]" italics="true" pageId="52" pageNumber="53">
<typeStatus id="549912A9645DFF85FF1BEBBCD0FAF927" box="[151,203,1708,1731]" pageId="52" pageNumber="53">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C22D788645DFF85FEABEBBBD289F927" authority="Staesche, 1964" authorityName="Staesche" authorityYear="1964" box="[295,696,1707,1731]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="anceps">
<emphasis id="B9567019645DFF85FEABEBBBD220F927" box="[295,529,1707,1731]" italics="true" pageId="52" pageNumber="53">Hadrodontina anceps</emphasis>
<bibRefCitation id="EFB3D1FA645DFF85FD9BEBBBD289F927" author="Staesche" box="[535,696,1707,1731]" firstAuthor="Staesche" pageId="52" pageNumber="53" refId="ref46867" refString="Staesche, U. 1964. Conodonten aus dem Skyth von Siidtirol. Neues Jahrbuch fiir Geologie und Palaontologie." type="book" year="1964">Staesche, 1964</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B645DFF85FF1BEBDCD165F8E7" blockId="52.[151,775,1707,1795]" pageId="52" pageNumber="53">
<materialsCitation id="3B4AA656645DFF85FF1BEBDCD161F8E7" country="Italy" county="South Tirol" location="Campiller" municipality="Skyth" pageId="52" pageNumber="53" specimenCount="1" typeStatus="holotype">
<emphasis id="B9567019645DFF85FF1BEBDCD1FDF907" box="[151,460,1739,1763]" italics="true" pageId="52" pageNumber="53">
<typeStatus id="549912A9645DFF85FF1BEBDCD0FAF907" box="[151,203,1740,1763]" pageId="52" pageNumber="53">Type</typeStatus>
stratum and locality.
</emphasis>
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member,
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,
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,
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.
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<emphasis id="B9567019645DFF85FF1BEA3BD0C4F8A7" box="[151,245,1835,1859]" italics="true" pageId="52" pageNumber="53">Remarks</emphasis>
.
<collectingRegion id="49E662E9645DFF85FE8FEA3BD173F8A6" box="[259,322,1835,1858]" country="Slovenia" name="Celje" pageId="52" pageNumber="53">Tere</collectingRegion>
is still an ongoing debate about the phylogenetic relationships of
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<emphasis id="B9567019645DFF85FE36EA5BD27DF887" box="[442,588,1867,1891]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C22D788645DFF85FDD0EA5BD08AF867" authorityName="Zhang, Aldridge &amp; Donoghue" authorityYear="1997" family="Ellisoniidae" genus="Parapachycladina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FDD0EA5BD08AF867" italics="true" pageId="52" pageNumber="53">Parapachycladina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788645DFF85FF79EA7BD1A1F867" authorityName="Staesche" authorityYear="1964" box="[245,400,1899,1923]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FF79EA7BD1A1F867" box="[245,400,1899,1923]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB3D1FA645DFF85FE13EA7BD21FF867" author="Sweet" box="[415,558,1899,1923]" firstAuthor="Sweet" pageId="52" pageNumber="53" refId="ref47251" refString="Sweet, W. C. (1988). The Conodonta: morphology, taxonomy, paleoecology, and evolutionary history of a long-extinct animal phylum (Vol. 10). Clarendon Press." type="book" year="1988">Sweet (1988)</bibRefCitation>
in his prioniodinid phylogeny considered
<taxonomicName id="4C22D788645DFF85FB93EDFBD480FEE7" box="[1055,1201,235,259]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FB93EDFBD480FEE7" box="[1055,1201,235,259]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788645DFF85FB69EDFBD5B1FEE7" authorityName="Staesche" authorityYear="1964" box="[1253,1408,235,259]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FB69EDFBD5B1FEE7" box="[1253,1408,235,259]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
as sister taxa. Some species of
<taxonomicName id="4C22D788645DFF85FBE4EC1BD4CBFEC7" box="[1128,1274,267,291]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FBE4EC1BD4CBFEC7" box="[1128,1274,267,291]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
were assigned to a new genus
<taxonomicName id="4C22D788645DFF85FC74EC3BD48CFEA7" authorityName="Zhang, Aldridge &amp; Donoghue" authorityYear="1997" box="[1016,1213,299,323]" family="Ellisoniidae" genus="Parapachycladina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FC74EC3BD48CFEA7" box="[1016,1213,299,323]" italics="true" pageId="52" pageNumber="53">Parapachycladina</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB3D1FA645DFF85FB7AEC3BD342FE87" author="Shunxin" etAl="et al." firstAuthor="Shunxin" pageId="52" pageNumber="53" pagination="65 - 72" refId="ref46531" refString="Shunxin, Z., Aldridge, R. J., &amp; Donoghue, P. C. (1997). An early Triassic conodont with periodic growth? Journal of Micropalaeontology, 16 (1), 65 - 72. https: // doi. org / 10.1144 / jm. 16.1. 65" type="journal article" year="1997">Shunxin et al. (1997)</bibRefCitation>
, but this view is not widely accepted. Orchard (2007) observed that
<taxonomicName id="4C22D788645DFF85FB96EC7BD537FE67" authorityName="Staesche" authorityYear="1964" box="[1050,1286,363,387]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="anceps">
<emphasis id="B9567019645DFF85FB96EC7BD537FE67" box="[1050,1286,363,387]" italics="true" pageId="52" pageNumber="53">Hadrodontina anceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C22D788645DFF85FA98EC7BD3B8FE47" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="triassica">
<emphasis id="B9567019645DFF85FA98EC7BD542FE67" box="[1300,1395,363,387]" italics="true" pageId="52" pageNumber="53">Ellisonia</emphasis>
aff.
<emphasis id="B9567019645DFF85FCA1EC9CD3B8FE47" box="[813,905,396,419]" italics="true" pageId="52" pageNumber="53">triassica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788645DFF85FC4BEC9BD4FEFE47" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[967,1231,395,419]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="species" species="peculiaris">
<emphasis id="B9567019645DFF85FC4BEC9BD4FEFE47" box="[967,1231,395,419]" italics="true" pageId="52" pageNumber="53">Pachycladina peculiaris</emphasis>
</taxonomicName>
appear to constitute a natural group, although they are currently assigned to different genera. Based on their cladistics analysis,
<bibRefCitation id="EFB3D1FA645DFF85FCA1ECFBD402FDE7" author="Donoghue" box="[813,1075,491,515]" etAl="et al." firstAuthor="Donoghue" pageId="52" pageNumber="53" pagination="119 - 153" refId="ref40086" refString="Donoghue, P. C., Purnell, M. A., Aldridge, R. J., &amp; Zhang, S. (2008). The interrelationships of ' complex'conodonts (Vertebrata). Journal of Systematic Palaeontology, 6 (2), 119 - 153. https: // doi. org / 10.1017 / S 14772019070022 34" type="journal article" year="2008">Donoghue et al. (2008)</bibRefCitation>
concluded that
<taxonomicName id="4C22D788645DFF85FB62ECFBD5B1FDE7" box="[1262,1408,491,515]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FB62ECFBD5B1FDE7" box="[1262,1408,491,515]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
is either a sister taxon to
<taxonomicName id="4C22D788645DFF85FBA3EF1BD4BFFDC7" authorityName="Muller" authorityYear="1956" box="[1071,1166,523,547]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FBA3EF1BD4BFFDC7" box="[1071,1166,523,547]" italics="true" pageId="52" pageNumber="53">Ellisonia</emphasis>
</taxonomicName>
or stays unresolved in a polytomy with
<emphasis id="B9567019645DFF85FC56EF3BD4D4FDA7" box="[986,1253,555,579]" italics="true" pageId="52" pageNumber="53">
<taxonomicName id="4C22D788645DFF85FC56EF3BD40AFDA7" authorityName="Muller" authorityYear="1956" box="[986,1083,555,579]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">Ellisonia</taxonomicName>
,
<taxonomicName id="4C22D788645DFF85FBC6EF3BD4D4FDA7" authorityName="Staesche" authorityYear="1964" box="[1098,1253,555,579]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">Hadrodontina</taxonomicName>
</emphasis>
and
<taxonomicName id="4C22D788645DFF85FAA8EF3BD5A7FDA7" authorityName="Clark" authorityYear="1959" box="[1316,1430,555,579]" genus="Furnishius" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FAA8EF3BD5A7FDA7" box="[1316,1430,555,579]" italics="true" pageId="52" pageNumber="53">Furnishius</emphasis>
</taxonomicName>
. Based on his multi-element apparatus reconstructions,
<bibRefCitation id="EFB3D1FA645DFF85FCA1EF7BD38CFD67" author="Koike" box="[813,957,619,643]" firstAuthor="Koike" pageId="52" pageNumber="53" pagination="161 - 175" refId="ref42946" refString="Koike, T. (2016). Multielement conodont apparatuses of the Ellisonidae from Japan. Paleontological Research, 20 (3), 161 - 175. https: // doi. org / 10.2517 / 2016 PR 007" type="journal article" year="2016">Koike (2016)</bibRefCitation>
included
<taxonomicName id="4C22D788645DFF85FBBDEF7BD4FDFD67" authorityName="Staesche" authorityYear="1964" box="[1073,1228,619,643]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FBBDEF7BD4FDFD67" box="[1073,1228,619,643]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788645DFF85FA86EF7BD5ADFD67" box="[1290,1436,619,643]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FA86EF7BD5ADFD67" box="[1290,1436,619,643]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
within the subfamily
<taxonomicName id="4C22D788645DFF85FB97EF9BD56FFD47" authority="(Koike, 2016)" baseAuthorityName="Koike" baseAuthorityYear="2016" box="[1051,1374,651,675]" family="Ellisoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Hadrodontinae">
Hadrodontinae (
<bibRefCitation id="EFB3D1FA645DFF85FB5DEF9BD565FD47" author="Koike" box="[1233,1364,651,675]" firstAuthor="Koike" pageId="52" pageNumber="53" pagination="161 - 175" refId="ref42946" refString="Koike, T. (2016). Multielement conodont apparatuses of the Ellisonidae from Japan. Paleontological Research, 20 (3), 161 - 175. https: // doi. org / 10.2517 / 2016 PR 007" type="journal article" year="2016">Koike, 2016</bibRefCitation>
)
</taxonomicName>
, supporting the original view of
<bibRefCitation id="EFB3D1FA645DFF85FBD2EFBBD4DDFD27" author="Sweet" box="[1118,1260,683,707]" firstAuthor="Sweet" pageId="52" pageNumber="53" refId="ref47251" refString="Sweet, W. C. (1988). The Conodonta: morphology, taxonomy, paleoecology, and evolutionary history of a long-extinct animal phylum (Vol. 10). Clarendon Press." type="book" year="1988">Sweet (1988)</bibRefCitation>
. We follow here this suprageneric classification and include
<taxonomicName id="4C22D788645DFF85FAA8EFDBD368FCE7" authorityName="Staesche" authorityYear="1964" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FAA8EFDBD368FCE7" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788645DFF85FC1CEFFBD413FCE7" box="[912,1058,747,771]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FC1CEFFBD413FCE7" box="[912,1058,747,771]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
within the subfamily
<taxonomicName id="4C22D788645DFF85FA9FEFFBD367FCC7" baseAuthorityName="Koike" baseAuthorityYear="2016" family="Ellisoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Hadrodontinae">Hadrodontinae</taxonomicName>
. More recently,
<bibRefCitation id="EFB3D1FA645DFF85FB85EE1BD48EFCC7" author="Sun" box="[1033,1215,779,803]" etAl="et al." firstAuthor="Sun" pageId="52" pageNumber="53" refId="ref47103" refString="Sun, Z., Liu, S., Ji, C., Jiang, D., &amp; Zhou, M. (2020). Synchrotron-aided reconstruction of the prioniodinin multielement conodont apparatus (Hadrodontina) from the Lower Triassic of China. Palaeogeography, Palaeoclimatology, Palaeoecology, 560, 109913. https: // doi. org / 10.1016 / j. palaeo. 2020. 109913" type="journal volume" year="2020">Sun et al. (2020)</bibRefCitation>
published 3 natural assemblages of
<taxonomicName id="4C22D788645DFF85FC57EE3BD4C1FCA7" authorityName="Staesche" authorityYear="1964" box="[987,1264,811,835]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645DFF85FC57EE3BD4C1FCA7" box="[987,1264,811,835]" italics="true" pageId="52" pageNumber="53">Hadrodontina aequabilis</emphasis>
</taxonomicName>
and confirmed the suprageneric classification of
<bibRefCitation id="EFB3D1FA645DFF85FB1AEE5BD513FC87" author="Koike" box="[1174,1314,843,867]" firstAuthor="Koike" pageId="52" pageNumber="53" pagination="161 - 175" refId="ref42946" refString="Koike, T. (2016). Multielement conodont apparatuses of the Ellisonidae from Japan. Paleontological Research, 20 (3), 161 - 175. https: // doi. org / 10.2517 / 2016 PR 007" type="journal article" year="2016">Koike (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCCEEE7CD4CAFAC7" blockId="52.[813,1437,235,1315]" pageId="52" pageNumber="53">
In P
<subScript id="17A6AE4E645DFF85FCF8EE67D34CFC6C" attach="left" box="[884,893,887,904]" fontSize="7" pageId="52" pageNumber="53">1</subScript>
elements, what most distinguishes
<taxonomicName id="4C22D788645DFF85FAADEE7BD360FC47" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FAADEE7BD360FC47" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
from
<taxonomicName id="4C22D788645DFF85FC10EE9BD40AFC47" authorityName="Staesche" authorityYear="1964" box="[924,1083,907,931]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FC10EE9BD40AFC47" box="[924,1083,907,931]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
is the basal configuration: the inverted basal attachment surface of
<taxonomicName id="4C22D788645DFF85FA8AEEBBD5ADFC27" box="[1286,1436,939,963]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FA8AEEBBD5ADFC27" box="[1286,1436,939,963]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
occupies the entire lower side plus one lateral side of the carina, whereas in
<taxonomicName id="4C22D788645DFF85FBDDEEFBD4C1FBE7" authorityName="Staesche" authorityYear="1964" box="[1105,1264,1003,1027]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FBDDEEFBD4C1FBE7" box="[1105,1264,1003,1027]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
, a basal cavity with a deep basal groove is usually formed on the lower side and the attachment surface rarely extends over the mid-part of the keel, if at all. Te denticles of
<taxonomicName id="4C22D788645DFF85FCA1E97BD3F2FB67" box="[813,963,1131,1155]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645DFF85FCA1E97BD3F2FB67" box="[813,963,1131,1155]" italics="true" pageId="52" pageNumber="53">Pachycladina</emphasis>
</taxonomicName>
are also less numerous but much larger in relative size than those of
<taxonomicName id="4C22D788645DFF85FB0FE99BD513FB47" authorityName="Staesche" authorityYear="1964" box="[1155,1314,1163,1187]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B9567019645DFF85FB0FE99BD513FB47" box="[1155,1314,1163,1187]" italics="true" pageId="52" pageNumber="53">Hadrodontina</emphasis>
</taxonomicName>
. Based on multi-element reconstructions however,
<bibRefCitation id="EFB3D1FA645DFF85FA8BE9BBD5ACFB27" author="Koike" box="[1287,1437,1195,1219]" firstAuthor="Koike" pageId="52" pageNumber="53" pagination="161 - 175" refId="ref42946" refString="Koike, T. (2016). Multielement conodont apparatuses of the Ellisonidae from Japan. Paleontological Research, 20 (3), 161 - 175. https: // doi. org / 10.2517 / 2016 PR 007" type="journal article" year="2016">Koike (2016)</bibRefCitation>
showed that
<taxonomicName id="4C22D788645DFF85FC44E9DBD4E9FB07" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[968,1240,1227,1251]" genus="Pachycladina" kingdom="Fungi" pageId="52" pageNumber="53" phylum="Ascomycota" rank="species" species="peculiaris">
<emphasis id="B9567019645DFF85FC44E9DBD4E9FB07" box="[968,1240,1227,1251]" italics="true" pageId="52" pageNumber="53">Pachycladina peculiaris</emphasis>
</taxonomicName>
, as well as
<taxonomicName id="4C22D788645DFF85FAE1E9DBD4D5FAE7" authority="(Koike et al., 2004)" baseAuthorityName="Koike" baseAuthorityYear="2004" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="triassica">
<emphasis id="B9567019645DFF85FAE1E9DBD359FAE7" italics="true" pageId="52" pageNumber="53">Ellisonia</emphasis>
aff.
<emphasis id="B9567019645DFF85FC12E9FCD3CCFAE7" box="[926,1021,1260,1283]" italics="true" pageId="52" pageNumber="53">triassica</emphasis>
(
<bibRefCitation id="EFB3D1FA645DFF85FB82E9FBD4EBFAE7" author="Koike" box="[1038,1242,1259,1283]" etAl="et al." firstAuthor="Koike" pageId="52" pageNumber="53" pagination="241 - 253" refId="ref42987" refString="Koike, T., Yamakita, S., &amp; Kadota, N. (2004). A natural assemblage of Ellisonia sp. cf. E. triassica MUller (Vertebrata: Conodonta) from the uppermost Permian in the Suzuka Mountains, central Japan. Paleontological Research, 8 (4), 241 - 253. https: // doi. org / 10.2517 / prpsj. 8.241" type="journal article" year="2004">Koike et al., 2004</bibRefCitation>
)
</taxonomicName>
, should be synonymized with
<emphasis id="B9567019645DFF85FC56E81BD4CAFAC7" box="[986,1275,1291,1315]" italics="true" pageId="52" pageNumber="53">
<taxonomicName id="4C22D788645DFF85FC56E81BD4C6FAC7" authorityName="Staesche" authorityYear="1964" box="[986,1271,1291,1315]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="aequabilis">Hadrodontina aequabilis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,560 @@
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<treatment id="038B1D1D645DFF86FCA1E85BD2E2FBC7" LSID="urn:lsid:plazi:treatment:038B1D1D645DFF86FCA1E85BD2E2FBC7" httpUri="http://treatment.plazi.org/id/038B1D1D645DFF86FCA1E85BD2E2FBC7" lastPageId="55" lastPageNumber="56" pageId="52" pageNumber="53">
<subSubSection id="C338FF80645DFF85FCA1E85BD4D3FA87" box="[813,1250,1355,1379]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8B9DAC0B645DFF85FCA1E85BD4D3FA87" blockId="52.[813,1250,1355,1480]" box="[813,1250,1355,1379]" pageId="52" pageNumber="53">
<heading id="D0D51B67645DFF85FCA1E85BD4D3FA87" box="[813,1250,1355,1379]" fontSize="9" level="2" pageId="52" pageNumber="53" reason="8">
<taxonomicName id="4C22D788645DFF85FCA1E85BD4D3FA87" authority="Staesche, 1964" authorityName="Staesche" authorityYear="1964" box="[813,1250,1355,1379]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645DFF85FCA1E85BD40FFA87" box="[813,1086,1355,1379]" italics="true" pageId="52" pageNumber="53">Hadrodontina aequabilis</emphasis>
<bibRefCitation id="EFB3D1FA645DFF85FBC8E85BD4D3FA87" author="Staesche" box="[1092,1250,1355,1379]" firstAuthor="Staesche" pageId="52" pageNumber="53" refId="ref46867" refString="Staesche, U. 1964. Conodonten aus dem Skyth von Siidtirol. Neues Jahrbuch fiir Geologie und Palaontologie." type="book" year="1964">Staesche, 1964</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645DFF85FCA1E87CD398F9E7" pageId="52" pageNumber="53" type="description">
<paragraph id="8B9DAC0B645DFF85FCA1E87CD484FA67" blockId="52.[813,1250,1355,1480]" box="[813,1205,1387,1416]" pageId="52" pageNumber="53">
P
<subScript id="17A6AE4E645DFF85FCB7E867D375FA6C" attach="left" box="[827,836,1399,1416]" fontSize="7" pageId="52" pageNumber="53">1</subScript>
elements:
<figureCitation id="1319B08E645DFF85FC35E87BD472FA67" box="[953,1091,1387,1411]" captionStart="Fig" captionStartId="53.[169,200,1777,1797]" captionTargetBox="[153,1439,238,1914]" captionTargetId="figure-11@53.[229,1363,254,1764]" captionTargetPageId="53" captionText="Fig. 27 Hadrodontinae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to beP1 elements if not specifically identified otherwise. A, Hadrodontina aequabilis (S2 element) (Staesche);SHA344, PIMUZ 39141. BE, Hadrodontina aequabilis (P2 element) (Staesche); B LAR214, PIMUZ 39142;C QIA13, PIMUZ 39143; D LIL500,PIMUZ 39144,E SHA344,PIMUZ 39145. FM Hadrodontina aequabilis (Staesche); F LIL502, PIMUZ 39146; G LIL503, PIMUZ 39147; H QIA134, PIMUZ 39148; I QIA134,PIMUZ 39149; J QIA 133, PIMUZ 39150; K LIL502,PIMUZ 39151; L LIL501,PIMUZ 39152; M LIL501, PIMUZ 39153" figureDoi="http://doi.org/10.5281/zenodo.12000474" httpUri="https://zenodo.org/record/12000474/files/figure.png" pageId="52" pageNumber="53">Figs. 27FM</figureCitation>
;
<figureCitation id="1319B08E645DFF85FBC1E87BD484FA67" box="[1101,1205,1387,1411]" captionStart="Fig" captionStartId="54.[169,200,825,845]" captionTargetBox="[153,1438,238,911]" captionTargetId="figure-393@54.[229,1363,254,813]" captionTargetPageId="54" captionText="Fig. 28 Hadrodontinae from Shanggang.Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, C, E Hadrodontina aequabilis (Staesche); A SHA 345C, PIMUZ 39154; C SHA345C,PIMUZ 39155; E SHA344C, PIMUZ 39156. B, D Hadrodontina aequabilis (P2 element) (Staesche); B SHA345C, PIMUZ 39157; D SHA345C, PIMUZ 39158" figureDoi="http://doi.org/10.5281/zenodo.12000480" httpUri="https://zenodo.org/record/12000480/files/figure.png" pageId="52" pageNumber="53">28A, C, E</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1E89CD4BEFA47" blockId="52.[813,1250,1355,1480]" box="[813,1167,1419,1448]" pageId="52" pageNumber="53">
P
<figureCitation id="1319B08E645DFF85FCB7E887D375FA4C" box="[827,836,1431,1448]" captionStart="Fig" captionStartId="3.[169,200,1622,1642]" captionTargetBox="[153,1439,238,1680]" captionTargetId="figure-95@3.[288,1304,254,1610]" captionTargetPageId="3" captionText="Fig. 2 Pictures from four of the five studied sections in the Nanpanjiang basin. A Qiakong, B Shanggang, C Laren (Kuang Guodun for scale), D Lilong (Hugo Bucher for scale)" figureDoi="http://doi.org/10.5281/zenodo.12000106" httpUri="https://zenodo.org/record/12000106/files/figure.png" pageId="52" pageNumber="53">
<subScript id="17A6AE4E645DFF85FCB7E887D375FA4C" attach="left" box="[827,836,1431,1448]" fontSize="7" pageId="52" pageNumber="53">2</subScript>
</figureCitation>
elements:
<figureCitation id="1319B08E645DFF85FC35E89BD40AFA47" box="[953,1083,1419,1443]" captionStart="Fig" captionStartId="53.[169,200,1777,1797]" captionTargetBox="[153,1439,238,1914]" captionTargetId="figure-11@53.[229,1363,254,1764]" captionTargetPageId="53" captionText="Fig. 27 Hadrodontinae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to beP1 elements if not specifically identified otherwise. A, Hadrodontina aequabilis (S2 element) (Staesche);SHA344, PIMUZ 39141. BE, Hadrodontina aequabilis (P2 element) (Staesche); B LAR214, PIMUZ 39142;C QIA13, PIMUZ 39143; D LIL500,PIMUZ 39144,E SHA344,PIMUZ 39145. FM Hadrodontina aequabilis (Staesche); F LIL502, PIMUZ 39146; G LIL503, PIMUZ 39147; H QIA134, PIMUZ 39148; I QIA134,PIMUZ 39149; J QIA 133, PIMUZ 39150; K LIL502,PIMUZ 39151; L LIL501,PIMUZ 39152; M LIL501, PIMUZ 39153" figureDoi="http://doi.org/10.5281/zenodo.12000474" httpUri="https://zenodo.org/record/12000474/files/figure.png" pageId="52" pageNumber="53">Figs. 27BE</figureCitation>
;
<figureCitation id="1319B08E645DFF85FBCAE89BD4BEFA47" box="[1094,1167,1419,1443]" captionStart="Fig" captionStartId="54.[169,200,825,845]" captionTargetBox="[153,1438,238,911]" captionTargetId="figure-393@54.[229,1363,254,813]" captionTargetPageId="54" captionText="Fig. 28 Hadrodontinae from Shanggang.Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, C, E Hadrodontina aequabilis (Staesche); A SHA 345C, PIMUZ 39154; C SHA345C,PIMUZ 39155; E SHA344C, PIMUZ 39156. B, D Hadrodontina aequabilis (P2 element) (Staesche); B SHA345C, PIMUZ 39157; D SHA345C, PIMUZ 39158" figureDoi="http://doi.org/10.5281/zenodo.12000480" httpUri="https://zenodo.org/record/12000480/files/figure.png" pageId="52" pageNumber="53">28B, D</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1E8BBD423FA27" blockId="52.[813,1250,1355,1480]" box="[813,1042,1451,1480]" pageId="52" pageNumber="53">
S
<figureCitation id="1319B08E645DFF85FCB6E8A7D372FA2C" box="[826,835,1463,1480]" captionStart="Fig" captionStartId="3.[169,200,1622,1642]" captionTargetBox="[153,1439,238,1680]" captionTargetId="figure-95@3.[288,1304,254,1610]" captionTargetPageId="3" captionText="Fig. 2 Pictures from four of the five studied sections in the Nanpanjiang basin. A Qiakong, B Shanggang, C Laren (Kuang Guodun for scale), D Lilong (Hugo Bucher for scale)" figureDoi="http://doi.org/10.5281/zenodo.12000106" httpUri="https://zenodo.org/record/12000106/files/figure.png" pageId="52" pageNumber="53">
<subScript id="17A6AE4E645DFF85FCB6E8A7D372FA2C" attach="left" box="[826,835,1463,1480]" fontSize="7" pageId="52" pageNumber="53">2</subScript>
</figureCitation>
elements:
<figureCitation id="1319B08E645DFF85FC34E8BBD423FA27" box="[952,1042,1451,1475]" captionStart="Fig" captionStartId="53.[169,200,1777,1797]" captionTargetBox="[153,1439,238,1914]" captionTargetId="figure-11@53.[229,1363,254,1764]" captionTargetPageId="53" captionText="Fig. 27 Hadrodontinae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to beP1 elements if not specifically identified otherwise. A, Hadrodontina aequabilis (S2 element) (Staesche);SHA344, PIMUZ 39141. BE, Hadrodontina aequabilis (P2 element) (Staesche); B LAR214, PIMUZ 39142;C QIA13, PIMUZ 39143; D LIL500,PIMUZ 39144,E SHA344,PIMUZ 39145. FM Hadrodontina aequabilis (Staesche); F LIL502, PIMUZ 39146; G LIL503, PIMUZ 39147; H QIA134, PIMUZ 39148; I QIA134,PIMUZ 39149; J QIA 133, PIMUZ 39150; K LIL502,PIMUZ 39151; L LIL501,PIMUZ 39152; M LIL501, PIMUZ 39153" figureDoi="http://doi.org/10.5281/zenodo.12000474" httpUri="https://zenodo.org/record/12000474/files/figure.png" pageId="52" pageNumber="53">Fig. 27A</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1E8FCD398F9E7" blockId="52.[813,1436,1515,1891]" box="[813,937,1515,1544]" pageId="52" pageNumber="53">
P
<figureCitation id="1319B08E645DFF85FCB7E8E7D375F9EC" box="[827,836,1527,1544]" captionStart="Fig" captionStartId="1.[831,862,1715,1735]" captionTargetBox="[815,1439,914,1715]" captionTargetId="figure-695@1.[843,1410,930,1703]" captionTargetPageId="1" captionText="Fig. 1 A Locations of the studied sections in the Nanpanjiang Basin (Luolou platform) (modified from Bagherpour et al., 2017). a—flood alluvial facies, b—shallow water siliclastic deposit, c—carbonate platform,d—slope,e—basin. Sections: 1—Qiakong, 2—Laren, 3—Shanggang, 4—Lilong, 5—Youping cascade.B Simplified palaeogeographical map of the Early Triassic (modified after the PANALESIS plate tectonic model of Vérard, 2019) South China indicated with a star" figureDoi="http://doi.org/10.5281/zenodo.12000098" httpUri="https://zenodo.org/record/12000098/files/figure.png" pageId="52" pageNumber="53">
<subScript id="17A6AE4E645DFF85FCB7E8E7D375F9EC" attach="left" box="[827,836,1527,1544]" fontSize="7" pageId="52" pageNumber="53">1</subScript>
</figureCitation>
element:
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645DFF87FCA1EB1BD397FB46" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="8B9DAC0B645DFF85FCA1EB1BD39BF9A7" blockId="52.[813,1436,1515,1891]" pageId="52" pageNumber="53">
*1964
<taxonomicName id="4C22D788645DFF85FCF9EB1BD4BDF9C7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[885,1164,1547,1571]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="aequabilis" status="sp. nov.">Hadrodontina aequabilis</taxonomicName>
<taxonomicNameLabel id="A265CD62645DFF85FB19EB1CD4E2F9C7" box="[1173,1235,1548,1571]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
; Staesche, p. 275, figs. 43, 44.
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1EB5BD380F967" blockId="52.[813,1436,1515,1891]" pageId="52" pageNumber="53">
1984
<taxonomicName id="4C22D788645DFF85FCE5EB5CD457F987" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[873,1126,1612,1635]" genus="Sweetocristatus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="unicus" status="sp. nov.">
<emphasis id="B9567019645DFF85FCE5EB5CD457F987" box="[873,1126,1612,1635]" italics="true" pageId="52" pageNumber="53">Sweetocristatus unicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645DFF85FBE1EB5CD49BF987" box="[1133,1194,1612,1635]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
; Dagis, pp. 3738, pl. X, figs. 69.
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1EB9BD391F927" blockId="52.[813,1436,1515,1891]" pageId="52" pageNumber="53">
1990
<taxonomicName id="4C22D788645DFF85FCE1EB9BD44AF947" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[877,1147,1675,1699]" genus="Pachycladina" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="B9567019645DFF85FCE1EB9BD44AF947" box="[877,1147,1675,1699]" italics="true" pageId="52" pageNumber="53">Pachycladina peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645DFF85FB0AEB9CD4F4F947" box="[1158,1221,1676,1699]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
; Shunxin (Zhang), pl. 2, fig. 4
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1EBDBD463F8E7" blockId="52.[813,1436,1515,1891]" pageId="52" pageNumber="53">
1991
<taxonomicName id="4C22D788645DFF85FCE7EBDBD446F907" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[875,1143,1739,1763]" genus="Pachycladina" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="B9567019645DFF85FCE7EBDBD446F907" box="[875,1143,1739,1763]" italics="true" pageId="52" pageNumber="53">Pachycladina peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645DFF85FBF3EBDCD48CF907" box="[1151,1213,1740,1763]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
; Zhang in Zhang &amp; Yang, p. 40, pl. 3, figs. 1, 2.
</paragraph>
<paragraph id="8B9DAC0B645DFF85FCA1EA1BD3A0F887" blockId="52.[813,1436,1515,1891]" pageId="52" pageNumber="53">
1997
<taxonomicName id="4C22D788645DFF85FCF4EA1BD517F8C7" authority="Zhang" authorityName="Zhang" box="[888,1318,1803,1827]" family="Ellisoniidae" genus="Parapachycladina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="peculiaris">
<emphasis id="B9567019645DFF85FCF4EA1BD4F7F8C7" box="[888,1222,1803,1827]" italics="true" pageId="52" pageNumber="53">Parapachycladina peculiaris</emphasis>
Zhang
</taxonomicName>
; Shunxin et al., pp. 6569, pl. 1, figs. 12, pl. 2, figs. 12, pl. 3 figs. 12.
</paragraph>
<caption id="DF5DFC83645CFF84FF25EBE1D34DF88B" ID-DOI="http://doi.org/10.5281/zenodo.12000474" ID-Zenodo-Dep="12000474" httpUri="https://zenodo.org/record/12000474/files/figure.png" pageId="53" pageNumber="54" startId="53.[169,200,1777,1797]" targetBox="[153,1439,238,1914]" targetPageId="53" targetType="figure">
<paragraph id="8B9DAC0B645CFF84FF25EBE1D34DF88B" blockId="53.[169,1416,1775,1903]" pageId="53" pageNumber="54">
<emphasis id="B9567019645CFF84FF25EBE1D0D9F8E1" bold="true" box="[169,232,1777,1797]" pageId="53" pageNumber="54">Fig. 27</emphasis>
<taxonomicName id="4C22D788645CFF84FF7EEBE1D142F8E1" baseAuthorityName="Koike" baseAuthorityYear="2016" box="[242,371,1777,1797]" family="Ellisoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Hadrodontinae">Hadrodontinae</taxonomicName>
from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to beP
<subScript id="17A6AE4E645CFF84FF43EA05D0E7F8C7" attach="left" box="[207,214,1813,1827]" fontSize="6" pageId="53" pageNumber="54">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B9567019645CFF84FDFFEA1BD2B0F8FB" bold="true" box="[627,641,1803,1823]" pageId="53" pageNumber="54">A</emphasis>
,
<taxonomicName id="4C22D788645CFF84FD05EA1BD362F8FB" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[649,851,1803,1823]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645CFF84FD05EA1BD362F8FB" box="[649,851,1803,1823]" italics="true" pageId="53" pageNumber="54">Hadrodontina aequabilis</emphasis>
</taxonomicName>
(S
<subScript id="17A6AE4E645CFF84FCE4EA05D35EF8C7" attach="left" box="[872,879,1813,1827]" fontSize="6" pageId="53" pageNumber="54">2</subScript>
element) (Staesche);SHA344, PIMUZ 39141.
<emphasis id="B9567019645CFF84FB66EA1BD53DF8FB" bold="true" box="[1258,1292,1803,1823]" pageId="53" pageNumber="54">BE</emphasis>
,
<taxonomicName id="4C22D788645CFF84FA98EA1BD0CDF8DE" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645CFF84FA98EA1BD0CDF8DE" italics="true" pageId="53" pageNumber="54">Hadrodontina aequabilis</emphasis>
</taxonomicName>
(P
<subScript id="17A6AE4E645CFF84FE9EEA20D128F8DA" attach="left" box="[274,281,1840,1854]" fontSize="6" pageId="53" pageNumber="54">2</subScript>
element) (Staesche);
<emphasis id="B9567019645CFF84FE42EA36D1EBF8DE" bold="true" box="[462,474,1830,1850]" pageId="53" pageNumber="54">B</emphasis>
LAR214, PIMUZ 39142;
<emphasis id="B9567019645CFF84FD2DEA36D29FF8DE" bold="true" box="[673,686,1830,1850]" pageId="53" pageNumber="54">C</emphasis>
QIA13, PIMUZ 39143;
<emphasis id="B9567019645CFF84FCE5EA36D349F8DE" bold="true" box="[873,888,1830,1850]" pageId="53" pageNumber="54">D</emphasis>
LIL500,PIMUZ 39144,
<emphasis id="B9567019645CFF84FBB9EA36D471F8DE" bold="true" box="[1077,1088,1830,1850]" pageId="53" pageNumber="54">E</emphasis>
SHA344,PIMUZ 39145.
<emphasis id="B9567019645CFF84FA86EA36D503F8DE" bold="true" box="[1290,1330,1830,1850]" pageId="53" pageNumber="54">FM</emphasis>
<taxonomicName id="4C22D788645CFF84FF25EA51D1E0F8B1" authority="(Staesche)" baseAuthorityName="Staesche" box="[169,465,1857,1877]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645CFF84FF25EA51D145F8B1" box="[169,372,1857,1877]" italics="true" pageId="53" pageNumber="54">Hadrodontina aequabilis</emphasis>
(Staesche)
</taxonomicName>
;
<emphasis id="B9567019645CFF84FE54EA51D1D2F8B1" bold="true" box="[472,483,1857,1877]" pageId="53" pageNumber="54">F</emphasis>
LIL502, PIMUZ 39146;
<emphasis id="B9567019645CFF84FD2DEA51D29EF8B1" bold="true" box="[673,687,1857,1877]" pageId="53" pageNumber="54">G</emphasis>
LIL503, PIMUZ 39147;
<emphasis id="B9567019645CFF84FCE1EA51D34AF8B1" bold="true" box="[877,891,1857,1877]" pageId="53" pageNumber="54">H</emphasis>
QIA134, PIMUZ 39148;
<emphasis id="B9567019645CFF84FBCCEA51D477F8B1" bold="true" box="[1088,1094,1857,1877]" pageId="53" pageNumber="54">I</emphasis>
QIA134,PIMUZ 39149;
<emphasis id="B9567019645CFF84FA87EA51D522F8B1" bold="true" box="[1291,1299,1857,1877]" pageId="53" pageNumber="54">J</emphasis>
QIA 133, PIMUZ 39150;
<emphasis id="B9567019645CFF84FEAFEA4BD11EF88B" bold="true" box="[291,303,1883,1903]" pageId="53" pageNumber="54">K</emphasis>
LIL502,PIMUZ 39151;
<emphasis id="B9567019645CFF84FE61EA4BD1C9F88B" bold="true" box="[493,504,1883,1903]" pageId="53" pageNumber="54">L</emphasis>
LIL501,PIMUZ 39152;
<emphasis id="B9567019645CFF84FD39EA4BD2F6F88B" bold="true" box="[693,711,1883,1903]" pageId="53" pageNumber="54">M</emphasis>
LIL501, PIMUZ 39153
</paragraph>
</caption>
<caption id="DF5DFC83645FFF87FF25EE29D497FC67" ID-DOI="http://doi.org/10.5281/zenodo.12000480" ID-Zenodo-Dep="12000480" httpUri="https://zenodo.org/record/12000480/files/figure.png" pageId="54" pageNumber="55" startId="54.[169,200,825,845]" targetBox="[153,1438,238,911]" targetPageId="54" targetType="figure">
<paragraph id="8B9DAC0B645FFF87FF25EE29D497FC67" blockId="54.[169,1366,824,902]" pageId="54" pageNumber="55">
<emphasis id="B9567019645FFF87FF25EE29D0D9FCA9" bold="true" box="[169,232,825,845]" pageId="54" pageNumber="55">Fig. 28</emphasis>
<taxonomicName id="4C22D788645FFF87FF7EEE29D142FCA9" baseAuthorityName="Koike" baseAuthorityYear="2016" box="[242,371,825,845]" family="Ellisoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Hadrodontinae">Hadrodontinae</taxonomicName>
from Shanggang.Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E645FFF87FB48EE53D4FAFCB5" attach="left" box="[1220,1227,835,849]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B9567019645FFF87FE31EE44D1FAFC8C" bold="true" box="[445,459,852,872]" pageId="54" pageNumber="55">A</emphasis>
,
<emphasis id="B9567019645FFF87FE5FEE44D1D1FC8C" bold="true" box="[467,480,852,872]" pageId="54" pageNumber="55">C</emphasis>
,
<emphasis id="B9567019645FFF87FE64EE44D1C2FC8C" bold="true" box="[488,499,852,872]" pageId="54" pageNumber="55">E</emphasis>
<taxonomicName id="4C22D788645FFF87FE7BEE44D32EFC8C" authority="(Staesche)" baseAuthorityName="Staesche" box="[503,799,852,872]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FE7BEE44D2F3FC8C" box="[503,706,852,872]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
(Staesche)
</taxonomicName>
;
<emphasis id="B9567019645FFF87FCAAEE44D305FC8C" bold="true" box="[806,820,852,872]" pageId="54" pageNumber="55">A</emphasis>
SHA 345C, PIMUZ 39154;
<emphasis id="B9567019645FFF87FB83EE44D42DFC8C" bold="true" box="[1039,1052,852,872]" pageId="54" pageNumber="55">C</emphasis>
SHA345C,PIMUZ 39155;
<emphasis id="B9567019645FFF87FB7DEE44D4CDFC8C" bold="true" box="[1265,1276,852,872]" pageId="54" pageNumber="55">E</emphasis>
SHA344C, PIMUZ 39156.
<emphasis id="B9567019645FFF87FEAFEE7FD11EFC67" bold="true" box="[291,303,879,899]" pageId="54" pageNumber="55">B</emphasis>
,
<emphasis id="B9567019645FFF87FEB4EE7FD176FC67" bold="true" box="[312,327,879,899]" pageId="54" pageNumber="55">D</emphasis>
<taxonomicName id="4C22D788645FFF87FEC7EE7FD224FC67" box="[331,533,879,899]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FEC7EE7FD224FC67" box="[331,533,879,899]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
</taxonomicName>
(P
<subScript id="17A6AE4E645FFF87FDA7EE68D203FC62" attach="left" box="[555,562,888,902]" fontSize="6" pageId="54" pageNumber="55">2</subScript>
element) (Staesche);
<emphasis id="B9567019645FFF87FD64EE7FD2C5FC67" bold="true" box="[744,756,879,899]" pageId="54" pageNumber="55">B</emphasis>
SHA345C, PIMUZ 39157;
<emphasis id="B9567019645FFF87FC46EE7FD3E8FC67" bold="true" box="[970,985,879,899]" pageId="54" pageNumber="55">D</emphasis>
SHA345C, PIMUZ 39158
</paragraph>
</caption>
<paragraph id="8B9DAC0B645FFF87FF1BEEDAD152FBE6" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2004
<taxonomicName id="4C22D788645FFF87FF54EEDAD289FC06" authority="Muller, 1956" authorityName="Muller" authorityYear="1956" box="[216,696,970,994]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019645FFF87FF54EEDAD10BFC06" box="[216,314,970,994]" italics="true" pageId="54" pageNumber="55">Ellisonia</emphasis>
sp. aff.
<emphasis id="B9567019645FFF87FE16EEDBD22BFC06" box="[410,538,971,994]" italics="true" pageId="54" pageNumber="55">E. triassica</emphasis>
<bibRefCitation id="EFB3D1FA645FFF87FDA9EEDAD289FC06" author="Muller" box="[549,696,970,994]" firstAuthor="Muller" pageId="54" pageNumber="55" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
</taxonomicName>
; Koike et al., figs. 8.7, 8.8.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE91AD150FBA6" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2009
<taxonomicName id="4C22D788645FFF87FF5FE91AD210FBC6" authority="Sweet" authorityName="Muller" authorityYear="1956" box="[211,545,1034,1058]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FF5FE91AD104FBC6" box="[211,309,1034,1058]" italics="true" pageId="54" pageNumber="55">Ellisonia</emphasis>
? cf.
<emphasis id="B9567019645FFF87FEEBE91AD1E4FBC6" box="[359,469,1034,1058]" italics="true" pageId="54" pageNumber="55">peculiaris</emphasis>
Sweet
</taxonomicName>
; Igo in Shigeta et al., p. 182, fig. 152.22.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE95AD0C4FB66" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2013
<taxonomicName id="4C22D788645FFF87FF5AE95AD205FB86" authority="Clark" authorityName="Clark" box="[214,564,1098,1122]" genus="Parachirognathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="geiseri">
<emphasis id="B9567019645FFF87FF5AE95AD1DBFB86" box="[214,490,1098,1122]" italics="true" pageId="54" pageNumber="55">Parachirognathus geiseri</emphasis>
Clark
</taxonomicName>
; Yan et al., p. 516, fig. 6 FF.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE99AD1A7FB26" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2015
<taxonomicName id="4C22D788645FFF87FF5AE99AD28CFB46" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" box="[214,701,1162,1186]" genus="Parachirognathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="peculiaris">
<emphasis id="B9567019645FFF87FF5AE99AD222FB46" box="[214,531,1162,1186]" italics="true" pageId="54" pageNumber="55">Parachirognathus peculiaris</emphasis>
Zhang &amp; Guo
</taxonomicName>
; Chen et al., figs. 8.2021, 24.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE9DAD1AFFAC6" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2015
<taxonomicName id="4C22D788645FFF87FF58E9DBD228FB06" authority="Dagis" authorityName="Dagis" box="[212,537,1226,1250]" genus="Sweetocristatus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="unicus">
<emphasis id="B9567019645FFF87FF58E9DBD1E1FB06" box="[212,464,1227,1250]" italics="true" pageId="54" pageNumber="55">Sweetocristatus unicus</emphasis>
Dagis
</taxonomicName>
; Chen et al., fig.
<date id="FF9C8ACB645FFF87FD5EE9DAD0FCFAE6" pageId="54" pageNumber="55" value="2016-08-23">8.23. 2016</date>
<taxonomicName id="4C22D788645FFF87FF52E9FAD247FAE6" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[222,630,1258,1282]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FF52E9FAD230FAE6" box="[222,513,1258,1282]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Koike, pp. 164167, fig. 2, nr. 13.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE83AD2B9FA86" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2018
<taxonomicName id="4C22D788645FFF87FF56E83AD258FAA6" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[218,617,1322,1346]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FF56E83AD1C8FAA6" box="[218,505,1322,1346]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Maekawa in Maekawa et al., p. 18, figs. 13.3?13.4 (only).
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE87AD125FA26" blockId="54.[151,775,970,1474]" pageId="54" pageNumber="55">
2020
<taxonomicName id="4C22D788645FFF87FF51E87AD242FA66" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[221,627,1386,1410]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FF51E87AD1CEFA66" box="[221,511,1386,1410]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Sun et al., figs. 17 (natural assemblage of the
<taxonomicName id="4C22D788645FFF87FDE4E89AD336FA46" authorityName="Staesche" authorityYear="1964" box="[616,775,1418,1442]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FDE4E89AD336FA46" box="[616,775,1418,1442]" italics="true" pageId="54" pageNumber="55">Hadrodontina</emphasis>
</taxonomicName>
apparatus).
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BE8FBD125F9E6" blockId="54.[151,775,1514,1826]" box="[151,276,1514,1543]" pageId="54" pageNumber="55">
P
<subScript id="17A6AE4E645FFF87FF2AE8E6D09EF9E3" attach="left" box="[166,175,1526,1543]" fontSize="7" pageId="54" pageNumber="55">2</subScript>
element:
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BEB1AD0D3F9A6" blockId="54.[151,775,1514,1826]" pageId="54" pageNumber="55">
*1964
<taxonomicName id="4C22D788645FFF87FF53EB1AD1C8F9C6" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[223,505,1546,1570]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis" status="sp. nov.">
<emphasis id="B9567019645FFF87FF53EB1AD1C8F9C6" box="[223,505,1546,1570]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645FFF87FD8EEB1BD20EF9C6" box="[514,575,1547,1570]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
; Staesche, p. 275, fig. 44.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BEB5AD126F966" blockId="54.[151,775,1514,1826]" pageId="54" pageNumber="55">
2004
<taxonomicName id="4C22D788645FFF87FF58EB5AD16DF986" box="[212,348,1610,1634]" family="Ellisoniidae" genus="Ellisonia" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019645FFF87FF58EB5AD107F986" box="[212,310,1610,1634]" italics="true" pageId="54" pageNumber="55">Ellisonia</emphasis>
sp.
</taxonomicName>
aff.
<taxonomicName id="4C22D788645FFF87FE00EB5BD26DF986" authority="Muller" authorityName="Muller" authorityYear="1956" box="[396,604,1610,1634]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="triassica">
<emphasis id="B9567019645FFF87FE00EB5BD236F986" box="[396,519,1611,1634]" italics="true" pageId="54" pageNumber="55">E. triassica</emphasis>
Müller
</taxonomicName>
; Koike et al., p. 247, fig. 8.6
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BEB9AD1B9F926" blockId="54.[151,775,1514,1826]" pageId="54" pageNumber="55">
2016
<taxonomicName id="4C22D788645FFF87FF5AEB9AD26FF946" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[214,606,1674,1698]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FF5AEB9AD1C0F946" box="[214,497,1674,1698]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Koike, p. 165, figs. 2.4 (P2 element).
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BEBDAD120F8C6" blockId="54.[151,775,1514,1826]" pageId="54" pageNumber="55">
2020
<taxonomicName id="4C22D788645FFF87FF52EBDAD25FF906" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[222,622,1738,1762]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FF52EBDAD1CAF906" box="[222,507,1738,1762]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Sun et al., figs. 17 (natural assemblage of the
<taxonomicName id="4C22D788645FFF87FDE0EBFAD336F8E6" authorityName="Staesche" authorityYear="1964" box="[620,775,1770,1794]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FDE0EBFAD336F8E6" box="[620,775,1770,1794]" italics="true" pageId="54" pageNumber="55">Hadrodontina</emphasis>
</taxonomicName>
apparatus).
</paragraph>
<paragraph id="8B9DAC0B645FFF87FF1BEA5AD399FBE6" blockId="54.[151,274,1866,1895]" lastBlockId="54.[813,1437,970,1186]" pageId="54" pageNumber="55">
S
<subScript id="17A6AE4E645FFF87FF29EA46D09FF883" attach="left" box="[165,174,1878,1895]" fontSize="7" pageId="54" pageNumber="55">2</subScript>
element. 2004
<taxonomicName id="4C22D788645FFF87FCE6EEDAD3DEFC06" box="[874,1007,970,994]" family="Ellisoniidae" genus="Ellisonia" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019645FFF87FCE6EEDAD3F8FC06" box="[874,969,970,994]" italics="true" pageId="54" pageNumber="55">Ellisonia</emphasis>
sp.
</taxonomicName>
aff.
<taxonomicName id="4C22D788645FFF87FBADEEDBD4C1FC06" authority="Muller" authorityName="Muller" authorityYear="1956" box="[1057,1264,970,994]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="triassica">
<emphasis id="B9567019645FFF87FBADEEDBD4AAFC06" box="[1057,1179,971,994]" italics="true" pageId="54" pageNumber="55">E. triassica</emphasis>
Müller
</taxonomicName>
; Koike et al., p. 247, fig. 8.4
</paragraph>
<paragraph id="8B9DAC0B645FFF87FCA1E91AD39EFBA6" blockId="54.[813,1437,970,1186]" pageId="54" pageNumber="55">
2016
<taxonomicName id="4C22D788645FFF87FCE1E91AD4DEFBC6" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[877,1263,1034,1058]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FCE1E91AD4B2FBC6" box="[877,1155,1034,1058]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Koike, p. 165, figs. 2.911.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FCA1E95AD397FB46" blockId="54.[813,1437,970,1186]" pageId="54" pageNumber="55">
2020
<taxonomicName id="4C22D788645FFF87FCFFE95AD532FB86" authority="Staesche" authorityName="Staesche" authorityYear="1964" box="[883,1283,1098,1122]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FCFFE95AD4A0FB86" box="[883,1169,1098,1122]" italics="true" pageId="54" pageNumber="55">Hadrodontina aequabilis</emphasis>
Staesche
</taxonomicName>
; Sun et al., figs. 17 (natural assemblage of the
<taxonomicName id="4C22D788645FFF87FA8DE97AD5ADFB66" authorityName="Staesche" authorityYear="1964" box="[1281,1436,1130,1154]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FA8DE97AD5ADFB66" box="[1281,1436,1130,1154]" italics="true" pageId="54" pageNumber="55">Hadrodontina</emphasis>
</taxonomicName>
apparatus).
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645FFF87FCA1E9DAD579FA27" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="8B9DAC0B645FFF87FCA1E9DAD3B4FAE6" blockId="54.[813,1436,1226,1282]" pageId="54" pageNumber="55">
<emphasis id="B9567019645FFF87FCA1E9DAD3A5FB06" box="[813,916,1226,1250]" italics="true" pageId="54" pageNumber="55">Material.</emphasis>
P
<subScript id="17A6AE4E645FFF87FC21E9C6D387FB03" attach="left" box="[941,950,1238,1255]" fontSize="7" pageId="54" pageNumber="55">1</subScript>
, more than 40; P
<subScript id="17A6AE4E645FFF87FBF3E9C6D4B9FB03" attach="left" box="[1151,1160,1238,1255]" fontSize="7" pageId="54" pageNumber="55">2</subScript>
, more than 40; S
<subScript id="17A6AE4E645FFF87FAC3E9C6D569FB03" attach="left" box="[1359,1368,1238,1255]" fontSize="7" pageId="54" pageNumber="55">2</subScript>
more than 10.
</paragraph>
<paragraph id="8B9DAC0B645FFF87FCA1E83AD579FA27" blockId="54.[813,1437,1322,1890]" pageId="54" pageNumber="55">
<emphasis id="B9567019645FFF87FCA1E83AD3C1FAA6" box="[813,1008,1322,1346]" italics="true" pageId="54" pageNumber="55">Revised diagnosis.</emphasis>
Robust to slender angulate P
<subScript id="17A6AE4E645FFF87FAA3E826D509FAA3" attach="left" box="[1327,1336,1334,1351]" fontSize="7" pageId="54" pageNumber="55">1</subScript>
element. Te 68 (usually 7) denticles are rather short, radiating, conical, with subtriangular free ends and they are increasingly reclined towards the posterior. Te concave lower side is occupied by a large and wide groove.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645FFF86FCCEE8DAD2E2FBC7" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8B9DAC0B645FFF86FCCEE8DAD332FD87" blockId="54.[813,1437,1322,1890]" lastBlockId="55.[151,775,235,1059]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis id="B9567019645FFF87FCCEE8DAD391FA06" box="[834,928,1482,1506]" italics="true" pageId="54" pageNumber="55">Remarks</emphasis>
. Te observed variation in the denticulation of the P
<subScript id="17A6AE4E645FFF87FC04E8E6D3A0F9E3" attach="left" box="[904,913,1526,1543]" fontSize="7" pageId="54" pageNumber="55">1</subScript>
and P
<subScript id="17A6AE4E645FFF87FC51E8E6D3D7F9E3" attach="left" box="[989,998,1526,1543]" fontSize="7" pageId="54" pageNumber="55">2</subScript>
elements suggests there may be scope for further specific differentiation: in some elements the height of the denticles changes smoothly along the unit (
<figureCitation id="1319B08E645FFF87FCB8EB5AD3A5F986" box="[820,916,1610,1634]" captionStart="Fig" captionStartId="54.[169,200,825,845]" captionTargetBox="[153,1438,238,911]" captionTargetId="figure-393@54.[229,1363,254,813]" captionTargetPageId="54" captionText="Fig. 28 Hadrodontinae from Shanggang.Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, C, E Hadrodontina aequabilis (Staesche); A SHA 345C, PIMUZ 39154; C SHA345C,PIMUZ 39155; E SHA344C, PIMUZ 39156. B, D Hadrodontina aequabilis (P2 element) (Staesche); B SHA345C, PIMUZ 39157; D SHA345C, PIMUZ 39158" figureDoi="http://doi.org/10.5281/zenodo.12000480" httpUri="https://zenodo.org/record/12000480/files/figure.png" pageId="54" pageNumber="55">Fig. 28B</figureCitation>
), whereas in others the denticles are alternatively high and low (
<figureCitation id="1319B08E645FFF87FBAFEB7AD4B8F966" box="[1059,1161,1642,1666]" captionStart="Fig" captionStartId="54.[169,200,825,845]" captionTargetBox="[153,1438,238,911]" captionTargetId="figure-393@54.[229,1363,254,813]" captionTargetPageId="54" captionText="Fig. 28 Hadrodontinae from Shanggang.Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, C, E Hadrodontina aequabilis (Staesche); A SHA 345C, PIMUZ 39154; C SHA345C,PIMUZ 39155; E SHA344C, PIMUZ 39156. B, D Hadrodontina aequabilis (P2 element) (Staesche); B SHA345C, PIMUZ 39157; D SHA345C, PIMUZ 39158" figureDoi="http://doi.org/10.5281/zenodo.12000480" httpUri="https://zenodo.org/record/12000480/files/figure.png" pageId="54" pageNumber="55">Fig. 28D</figureCitation>
).
<bibRefCitation id="EFB3D1FA645FFF87FB2EEB7AD503F966" author="Dagis" box="[1186,1330,1642,1666]" firstAuthor="Dagis" pageId="54" pageNumber="55" pagination="1 - 69" refId="ref39946" refString="Dagis, A. A. (1984). Early Triassic conodonts of northern Middle Siberia. Academy of Sciences of the USSR, Siberian Branch. Institute of Geology and Geophysics Transactions, 554, 1 - 69." type="journal article" year="1984">Dagis (1984)</bibRefCitation>
assigned some specimens (here considered as junior synonyms of
<taxonomicName id="4C22D788645FFF87FCA1EBBBD3FDF926" authorityName="Staesche" authorityYear="1964" box="[813,972,1706,1730]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645FFF87FCA1EBBBD3FDF926" box="[813,972,1706,1730]" italics="true" pageId="54" pageNumber="55">Ha. aequabilis</emphasis>
</taxonomicName>
) to
<taxonomicName id="4C22D788645FFF87FC70EBBBD4C3F926" box="[1020,1266,1707,1730]" genus="Sweetocristatus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="unicus">
<emphasis id="B9567019645FFF87FC70EBBBD4C3F926" box="[1020,1266,1707,1730]" italics="true" pageId="54" pageNumber="55">Sweetocristatus unicus</emphasis>
</taxonomicName>
. Yet, the genus
<taxonomicName id="4C22D788645FFF87FCA1EBDBD465F906" authority="Szaniawski" authorityName="Szaniawski" authorityYear="1979" box="[813,1108,1738,1762]" genus="Sweetocristatus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FCA1EBDBD3E5F906" box="[813,980,1739,1762]" italics="true" pageId="54" pageNumber="55">Sweetocristatus</emphasis>
Szaniawski
</taxonomicName>
(in
<bibRefCitation id="EFB3D1FA645FFF87FB08EBDAD350F8E6" author="Szaniawski &amp; Malkowski" firstAuthor="Szaniawski" pageId="54" pageNumber="55" pagination="231 - 264" refId="ref47343" refString="Szaniawski, H., &amp; Malkowski, K. (1979). Conodonts from the Kapp Starostin Formation (Permian) of Spitsbergen. Acta Palaeontologica Polonica, 24 (2), 231 - 264." type="journal article" year="1979">Szaniawski &amp; Malkowski, 1979</bibRefCitation>
) was established for Upper Artinskian to Lower Guadalupian (Permian) P
<subScript id="17A6AE4E645FFF87FBCDEA06D47BF8C3" attach="left" box="[1089,1098,1814,1831]" fontSize="7" pageId="54" pageNumber="55">1</subScript>
elements that, although superficially similar to homologous elements of
<taxonomicName id="4C22D788645FFF87FA8DEA3AD5ADF8A6" authorityName="Staesche" authorityYear="1964" box="[1281,1436,1834,1858]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9567019645FFF87FA8DEA3AD5ADF8A6" box="[1281,1436,1834,1858]" italics="true" pageId="54" pageNumber="55">Hadrodontina</emphasis>
</taxonomicName>
and
<emphasis id="B9567019645FFF87FCD3EA5AD3C6F886" box="[863,1015,1866,1890]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="4C22D788645FFF87FCD3EA5AD3C2F886" box="[863,1011,1866,1890]" genus="Pachycladina" kingdom="Fungi" pageId="54" pageNumber="55" phylum="Ascomycota" rank="genus">Pachycladina</taxonomicName>
,
</emphasis>
bear a more developed, higher cusp, a higher carina and have a more elongated process. Te cusp in
<taxonomicName id="4C22D788645EFF86FF7EEC1BD160FEC7" authorityName="Muller" authorityYear="1956" box="[242,337,267,291]" family="Ellisoniidae" genus="Ellisonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="B9567019645EFF86FF7EEC1BD160FEC7" box="[242,337,267,291]" italics="true" pageId="55" pageNumber="56">Ellisonia</emphasis>
</taxonomicName>
P
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elements is more conspicuous than in those of
<taxonomicName id="4C22D788645EFF86FE92EC3BD188FEA7" authorityName="Staesche" authorityYear="1964" box="[286,441,299,323]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="B9567019645EFF86FE92EC3BD188FEA7" box="[286,441,299,323]" italics="true" pageId="55" pageNumber="56">Hadrodontina</emphasis>
</taxonomicName>
or
<taxonomicName id="4C22D788645EFF86FE6AEC3BD249FEA7" box="[486,632,299,323]" genus="Pachycladina" kingdom="Fungi" pageId="55" pageNumber="56" phylum="Ascomycota" rank="genus">
<emphasis id="B9567019645EFF86FE6AEC3BD249FEA7" box="[486,632,299,323]" italics="true" pageId="55" pageNumber="56">Pachycladina</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB3D1FA645EFF86FD05EC3BD0ECFE87" author="Chen" etAl="et al." firstAuthor="Chen" pageId="55" pageNumber="56" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
illustrated coeval specimens from Jiarong they assigned to either
<taxonomicName id="4C22D788645EFF86FEEBEC7CD26FFE67" box="[359,606,364,387]" genus="Sweetocristatus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="unicus">
<emphasis id="B9567019645EFF86FEEBEC7CD26FFE67" box="[359,606,364,387]" italics="true" pageId="55" pageNumber="56">Sweetocristatus unicus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C22D788645EFF86FD05EC7BD162FE47" genus="Parachirognathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="peculiaris">
<emphasis id="B9567019645EFF86FD05EC7BD162FE47" italics="true" pageId="55" pageNumber="56">Parachirognathus peculiaris</emphasis>
</taxonomicName>
(see their figs. 8.20 and 8.23) but we fail to distinguish them from
<taxonomicName id="4C22D788645EFF86FE3CECBBD2F3FE27" authorityName="Staesche" authorityYear="1964" box="[432,706,427,451]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645EFF86FE3CECBBD2F3FE27" box="[432,706,427,451]" italics="true" pageId="55" pageNumber="56">Hadrodontina aequabilis</emphasis>
</taxonomicName>
. With three natural assemblages of
<taxonomicName id="4C22D788645EFF86FE62ECDBD336FE07" authorityName="Staesche" authorityYear="1964" box="[494,775,459,483]" family="Ellisoniidae" genus="Hadrodontina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="aequabilis">
<emphasis id="B9567019645EFF86FE62ECDBD336FE07" box="[494,775,459,483]" italics="true" pageId="55" pageNumber="56">Hadrodontina aequabilis</emphasis>
</taxonomicName>
from the late Smithian Helongshan Formation, South
<collectingCountry id="F335EC9B645EFF86FF1BEF1BD0EAFDC7" box="[151,219,523,547]" name="China" pageId="55" pageNumber="56">China</collectingCountry>
,
<bibRefCitation id="EFB3D1FA645EFF86FF65EF1BD193FDC7" author="Sun" box="[233,418,523,547]" etAl="et al." firstAuthor="Sun" pageId="55" pageNumber="56" refId="ref47103" refString="Sun, Z., Liu, S., Ji, C., Jiang, D., &amp; Zhou, M. (2020). Synchrotron-aided reconstruction of the prioniodinin multielement conodont apparatus (Hadrodontina) from the Lower Triassic of China. Palaeogeography, Palaeoclimatology, Palaeoecology, 560, 109913. https: // doi. org / 10.1016 / j. palaeo. 2020. 109913" type="journal volume" year="2020">Sun et al. (2020)</bibRefCitation>
observations fits with our findings; the angulate shape and no distinct cusp in the P
<subScript id="17A6AE4E645EFF86FD72EF27D336FDAC" attach="left" box="[766,775,567,584]" fontSize="7" pageId="55" pageNumber="56">1</subScript>
element and the distally twisting shape of the P
<subScript id="17A6AE4E645EFF86FD19EF47D2AFFD8C" attach="left" box="[661,670,599,616]" fontSize="7" pageId="55" pageNumber="56">2</subScript>
element.
</paragraph>
<paragraph id="8B9DAC0B645EFF86FF21EF7CD2E2FBC7" blockId="55.[151,775,235,1059]" pageId="55" pageNumber="56">
<emphasis id="B9567019645EFF86FF21EF7CD119FD67" box="[173,296,620,643]" italics="true" pageId="55" pageNumber="56">Occurrence</emphasis>
.
<collectingCountry id="F335EC9B645EFF86FEC8EF7BD1BDFD67" box="[324,396,619,643]" name="Russia" pageId="55" pageNumber="56">Russia</collectingCountry>
; Smithian
<taxonomicName id="4C22D788645EFF86FD93EF7BD2F6FD67" box="[543,711,619,643]" class="Cephalopoda" family="Hedenstroemiidae" genus="Hedenstroemia" kingdom="Animalia" pageId="55" pageNumber="21" phylum="Mollusca" rank="genus">Hedenstroemia</taxonomicName>
and tardus zone, Siberia (
<bibRefCitation id="EFB3D1FA645EFF86FE1CEF9BD221FD47" author="Dagis" box="[400,528,651,675]" firstAuthor="Dagis" pageId="55" pageNumber="56" pagination="1 - 69" refId="ref39946" refString="Dagis, A. A. (1984). Early Triassic conodonts of northern Middle Siberia. Academy of Sciences of the USSR, Siberian Branch. Institute of Geology and Geophysics Transactions, 554, 1 - 69." type="journal article" year="1984">Dagis 1984</bibRefCitation>
), Zhitkov Formation, Smithian age, Abrek Bay area, South Primorye (
<bibRefCitation id="EFB3D1FA645EFF86FD23EFBBD0FAFD07" author="Shigeta" firstAuthor="Shigeta" pageId="55" pageNumber="56" refId="ref46464" refString="Shigeta, Y., Zakharov, Y. D., Maeda, H., &amp; Popov, A. M. (Eds.). 2009. The Lower Triassic System in the Abrek Bay Area, South Primorye, Russia" type="book" year="2009">Shigeta, 2009</bibRefCitation>
).
<collectingCountry id="F335EC9B645EFF86FF6DEFDBD114FD07" box="[225,293,715,739]" name="China" pageId="55" pageNumber="56">China</collectingCountry>
; Beisi Formation, Taiping, Pingguo Western
<collectingRegion id="49E662E9645EFF86FF1BEFFBD16CFCE7" box="[151,349,747,771]" country="China" name="Guangxi" pageId="55" pageNumber="56">Guangxi Province</collectingRegion>
(
<bibRefCitation id="EFB3D1FA645EFF86FEE0EFFBD278FCE7" author="Shunxin" box="[364,585,747,771]" etAl="et al." firstAuthor="Shunxin" pageId="55" pageNumber="56" pagination="65 - 72" refId="ref46531" refString="Shunxin, Z., Aldridge, R. J., &amp; Donoghue, P. C. (1997). An early Triassic conodont with periodic growth? Journal of Micropalaeontology, 16 (1), 65 - 72. https: // doi. org / 10.1144 / jm. 16.1. 65" type="journal article" year="1997">Shunxin et al., 1997</bibRefCitation>
), Luolou Formation, Smithian age, Jiarong and Bianyang, Nanpanjiang Basin, southern
<collectingRegion id="49E662E9645EFF86FED9EE3BD210FCA7" box="[341,545,811,835]" country="China" name="Guizhou" pageId="55" pageNumber="56">Guizhou Province</collectingRegion>
(
<bibRefCitation id="EFB3D1FA645EFF86FDBAEE3BD331FCA7" author="Chen" box="[566,768,811,835]" etAl="et al." firstAuthor="Chen" pageId="55" pageNumber="56" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA645EFF86FF1BEE5CD175FC87" author="Yan" box="[151,324,843,867]" etAl="et al." firstAuthor="Yan" pageId="55" pageNumber="56" pagination="509 - 522" refId="ref48484" refString="Yan, C., Wang, L., Jiang, H., Wignall, P. B., Sun, Y., Chen, Y., &amp; Lai, X. (2013). Uppermost Permian to Lower Triassic conodonts at Bianyang section, Guizhou province, South China. Palaios, 28 (8), 509 - 522. https: // doi. org / 10.2110 / palo. 2012. p 12 - 077 r" type="journal article" year="2013">Yan et al., 2013</bibRefCitation>
), Late Smithian Helongshan Formation, Chaohu,
<collectingRegion id="49E662E9645EFF86FF71EE7BD19AFC67" box="[253,427,875,899]" country="China" name="Anhui" pageId="55" pageNumber="56">Anhui Province</collectingRegion>
(
<bibRefCitation id="EFB3D1FA645EFF86FE30EE7BD25AFC67" author="Sun" box="[444,619,875,899]" etAl="et al." firstAuthor="Sun" pageId="55" pageNumber="56" refId="ref47103" refString="Sun, Z., Liu, S., Ji, C., Jiang, D., &amp; Zhou, M. (2020). Synchrotron-aided reconstruction of the prioniodinin multielement conodont apparatus (Hadrodontina) from the Lower Triassic of China. Palaeogeography, Palaeoclimatology, Palaeoecology, 560, 109913. https: // doi. org / 10.1016 / j. palaeo. 2020. 109913" type="journal volume" year="2020">Sun et al., 2020</bibRefCitation>
).
<collectingCountry id="F335EC9B645EFF86FD08EE7CD2F2FC67" box="[644,707,876,899]" name="Japan" pageId="55" pageNumber="56">Japan</collectingCountry>
: Taho Formation, Shirokawa-cho, Higashiuwa-gun,
<collectingRegion id="49E662E9645EFF86FD4DEE9BD139FC27" country="Japan" name="Ehime" pageId="55" pageNumber="56">Ehime Prefecture</collectingRegion>
(below SmithianSpathian boundary) (
<bibRefCitation id="EFB3D1FA645EFF86FD4AEEBBD0FAFC07" author="Koike" firstAuthor="Koike" pageId="55" pageNumber="56" pagination="161 - 175" refId="ref42946" refString="Koike, T. (2016). Multielement conodont apparatuses of the Ellisonidae from Japan. Paleontological Research, 20 (3), 161 - 175. https: // doi. org / 10.2517 / 2016 PR 007" type="journal article" year="2016">Koike 2016</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA645EFF86FF53EEDBD19FFC07" author="Koike" box="[223,430,971,995]" etAl="et al." firstAuthor="Koike" pageId="55" pageNumber="56" pagination="241 - 253" refId="ref42987" refString="Koike, T., Yamakita, S., &amp; Kadota, N. (2004). A natural assemblage of Ellisonia sp. cf. E. triassica MUller (Vertebrata: Conodonta) from the uppermost Permian in the Suzuka Mountains, central Japan. Paleontological Research, 8 (4), 241 - 253. https: // doi. org / 10.2517 / prpsj. 8.241" type="journal article" year="2004">Koike et al., 2004</bibRefCitation>
). Europe: Campiller member, Werfen Formation, Skyth, South Tirol,
<collectingCountry id="F335EC9B645EFF86FDD2EEFBD2BFFBE7" box="[606,654,1003,1027]" name="Italy" pageId="55" pageNumber="56">Italy</collectingCountry>
(
<bibRefCitation id="EFB3D1FA645EFF86FD2DEEFBD0FAFBC7" author="Staesche" firstAuthor="Staesche" pageId="55" pageNumber="56" refId="ref46867" refString="Staesche, U. 1964. Conodonten aus dem Skyth von Siidtirol. Neues Jahrbuch fiir Geologie und Palaontologie." type="book" year="1964">Staesche, 1964</bibRefCitation>
),
<collectingCountry id="F335EC9B645EFF86FF6CE91BD10DFBC7" box="[224,316,1035,1059]" name="Slovenia" pageId="55" pageNumber="56">Slovenia</collectingCountry>
(
<bibRefCitation id="EFB3D1FA645EFF86FEC5E91BD2F5FBC7" author="Kolar-Jurkovsek &amp; Jurkovsek" box="[329,708,1035,1059]" firstAuthor="Kolar-Jurkovsek" pageId="55" pageNumber="56" refId="ref43188" refString="Kolar-Jurkovsek, T., &amp; Jurkovsek, B. (2019). Konodonti Slovenije / Conodonts of Slovenia (p. 260). Ljubljana: Geoloski zavod Slovenije." type="book" year="2019">Kolar-Jurkovšek &amp; Jurkovšek, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,524 @@
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<mods:namePart id="8631F3C285F59403AED5691418AF6DAE">Vennemann, Torsten</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="5DA9776A78856408FFBF719F0F08A473">Bagherpour, Borhan</mods:namePart>
</mods:name>
<mods:name id="254D8E8BC5F358B09DD5ED228BC3C804" type="personal">
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</mods:role>
<mods:namePart id="109E85E2A3A460D050719CF359FE075B">Ji, Cheng</mods:namePart>
</mods:name>
<mods:name id="11D1B60DB979A60401D05B726B80134C" type="personal">
<mods:role id="F09B9B22F64B457BA61A1770EF4536B4">
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</mods:role>
<mods:namePart id="ACB0D5ACEBB52E5FA0192A9E12B2D437">Brosse, Morgane</mods:namePart>
</mods:name>
<mods:name id="D4206500F382F53459F57C02D19B989A" type="personal">
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</mods:role>
<mods:namePart id="2515E28DB694648158CB95CECDCF822D">Goudemand, Nicolas</mods:namePart>
</mods:name>
<mods:typeOfResource id="4527C1C28F8EAF6159A2D42993EB7AF2">text</mods:typeOfResource>
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<mods:titleInfo id="3DD0C420318079E9E0DD7A689D412AAE">
<mods:title id="FFAACA339BFBB4827BB9529CB00B4918">Swiss Journal of Palaeontology</mods:title>
</mods:titleInfo>
<mods:part id="44676B4CA3FC5F61C1866185A0DD6324">
<mods:date id="705A2B45C93B229921C165D24A5DBEE2">2022</mods:date>
<mods:detail id="E1DD3D6B20D842985365E16F44B2E7F3" type="series">
<mods:title id="F306C5E2EEE473E8CE61F856EBAB5DDD">19</mods:title>
</mods:detail>
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<mods:number id="113482595B5D803F582A12833ACCDBE9">2022-11-22</mods:number>
</mods:detail>
<mods:detail id="7132210B390708E4CBCE9D84072983B4" type="volume">
<mods:number id="F58BC11E9FE80C2D88C358C826FF63C6">141</mods:number>
</mods:detail>
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<mods:number id="E7716616C9985C06E2717E56E4DE828A">1</mods:number>
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<mods:start id="3D7A8D53F8A5CF7A048F04864BEF92DF">1</mods:start>
<mods:end id="D8DDCDD4318A42F31E5440555E3F3EE8">61</mods:end>
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<mods:url id="17E69DCD7956F9AD46EBAD36B63AF722">http://dx.doi.org/10.1186/s13358-022-00259-x</mods:url>
</mods:location>
<mods:classification id="B2013282B01CB260ED5ABC374070134A">journal article</mods:classification>
<mods:identifier id="29C3B9973CA9791C3FBA6EEA49FB000B" type="DOI">10.1186/s13358-022-00259-x</mods:identifier>
<mods:identifier id="01FFC497A8211E6F3327478BD2D9E7B0" type="ISSN">1664-2384</mods:identifier>
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<mods:identifier id="9144E1421D243B2FAA0707AD8FA14563" type="Zenodo-Dep">12000082</mods:identifier>
</mods:mods>
<treatment id="038B1D1D6473FFADFF1BEF9BD2A8FDA7" LSID="urn:lsid:plazi:treatment:038B1D1D6473FFADFF1BEF9BD2A8FDA7" httpUri="http://treatment.plazi.org/id/038B1D1D6473FFADFF1BEF9BD2A8FDA7" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C338FF806473FFABFF1BEF9BD0E4FD27" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B9DAC0B6473FFABFF1BEF9BD0E4FD27" blockId="26.[151,775,651,739]" pageId="26" pageNumber="27">
<taxonomicName id="4C22D7886473FFABFF1BEF9BD0E4FD27" authority="(Zhao &amp; Orchard, 2007)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF1BEF9BD223FD47" box="[151,530,651,675]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(
<bibRefCitation id="EFB3D1FA6473FFABFDA2EF9BD0FAFD27" author="Zhao &amp; Orchard" firstAuthor="Zhao" pageId="26" pageNumber="27" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao &amp; Orchard, 2007</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806473FFABFF1BEFDBD2B3FD07" box="[151,642,715,739]" pageId="26" pageNumber="27" type="description">
<paragraph id="8B9DAC0B6473FFABFF1BEFDBD2B3FD07" blockId="26.[151,775,651,739]" box="[151,642,715,739]" pageId="26" pageNumber="27">
<figureCitation id="1319B08E6473FFABFF1BEFDBD1F4FD07" box="[151,453,715,739]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.13127510" httpUri="https://zenodo.org/record/13127510/files/figure.png" pageId="26" pageNumber="27">Fig. 17DF; MP, R, S, U, X</figureCitation>
, AA, AB, AD, AF
</paragraph>
</subSubSection>
<subSubSection id="C338FF806473FFABFF1BEE1BD47EFEE7" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="8B9DAC0B6473FFABFF1BEE1BD269FCA7" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
*2007
<taxonomicName id="4C22D7886473FFABFF6FEE1BD279FCC7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[227,584,779,803]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis" status="sp. nov.">
<emphasis id="B95670196473FFABFF6FEE1BD279FCC7" box="[227,584,779,803]" italics="true" pageId="26" pageNumber="27">Neospathodus pingdingshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFDDBEE1CD2A8FCC7" box="[599,665,780,803]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
; Zhao &amp; Orchard, Zhao et al., p.36, pl. 1, fig. 4AC.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BEE5BD2EAFC47" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2012a
<taxonomicName id="4C22D7886473FFABFF79EE5BD0CFFC67" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF79EE5BD249FC87" box="[245,632,843,867]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Goudemand &amp; Orchard in Goudemand et al., p. 10301031, figs. 2B, F, G, IJ, M, P, Q, AD, 3T-U, 6.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BEEBBD1C4FC07" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2013
<taxonomicName id="4C22D7886473FFABFF59EEBBD332FC27" authority="Zhao &amp; Orchard" authorityName="Zhao &amp; Orchard" box="[213,771,939,963]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF59EEBBD20DFC27" box="[213,572,939,963]" italics="true" pageId="26" pageNumber="27">Neospathodus pingdingshanensis</emphasis>
Zhao &amp; Orchard
</taxonomicName>
; Chen et al., p 825, fig. 3.10, 3.12.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BEEFBD27CFBC7" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2013
<taxonomicName id="4C22D7886473FFABFF50EEFBD1EFFBE7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[220,478,1003,1027]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni" status="subsp. nov.">
<emphasis id="B95670196473FFABFF50EEFBD1EFFBE7" box="[220,478,1003,1027]" italics="true" pageId="26" pageNumber="27">Neospathodus waageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFE62EEFBD241FBE7" box="[494,624,1003,1027]" pageId="26" pageNumber="27" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
A; Metcalfe et al., p. 1144, figs. 9.19.5, 9.7, 9.8, 9.10.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE93BD15EFB67" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2014
<taxonomicName id="4C22D7886473FFABFF60E93BD0CFFB87" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF60E93BD243FBA7" box="[236,626,1067,1091]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Maekawa &amp; Igo in Shigeta, p. 239240, figs. 171.13171.31.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE99BD2DAFB27" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2015
<taxonomicName id="4C22D7886473FFABFF60E99BD0CFFB27" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF60E99BD243FB47" box="[236,626,1163,1187]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Chen et al., p. 111, 112, figs. 7.17.4, 8.5, 8.6.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE9DBD2A3FAE7" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2016
<taxonomicName id="4C22D7886473FFABFF5BE9DBD148FB07" authorityName="Orchard" authorityYear="2005" box="[215,377,1227,1251]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFF5BE9DBD148FB07" box="[215,377,1227,1251]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
ex. gr.
<taxonomicName id="4C22D7886473FFABFE5BE9DBD0CFFAE7" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFE5BE9DBD2AAFB07" box="[471,667,1227,1251]" italics="true" pageId="26" pageNumber="27">pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Komatsu et al., p. 69, figs. 5.4a5.5c.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE81BD249FAA7" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2016
<taxonomicName id="4C22D7886473FFABFF52E81BD208FAC7" authority="Koike" authorityName="Koike" authorityYear="1982" box="[222,569,1291,1315]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B95670196473FFABFF52E81BD1D7FAC7" box="[222,486,1291,1315]" italics="true" pageId="26" pageNumber="27">Neospathodus robustus</emphasis>
Koike
</taxonomicName>
; Chen &amp; Kolar-Jurkovšek in Chen et al., p. 93, fig. 9.5 (only).
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE85BD1FAFA47" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2018
<taxonomicName id="4C22D7886473FFABFF60E85BD0CFFA67" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF60E85BD243FA87" box="[236,626,1355,1379]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Maekawa in Maekawa et al., p. 3637, figs. 20.220.18, 21.121.13.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE8BBD1CEFA07" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2019
<taxonomicName id="4C22D7886473FFABFF60E8BBD0CFFA07" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF60E8BBD243FA27" box="[236,626,1451,1475]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Chen et al., fig. 3, nr. 8.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF1BE8FBD259F9C7" blockId="26.[151,775,779,1571]" pageId="26" pageNumber="27">
2019
<taxonomicName id="4C22D7886473FFABFF60E8FBD0CFF9C7" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF60E8FBD243F9E7" box="[236,626,1515,1539]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Liu et al., p. 13, pl. 3 fig. 5 (only).
</paragraph>
<paragraph id="8B9DAC0B6473FFABFCA1EDFBD47EFEE7" blockId="26.[813,1103,235,259]" box="[813,1103,235,259]" pageId="26" pageNumber="27">
<emphasis id="B95670196473FFABFCA1EDFBD3A5FEE7" box="[813,916,235,259]" italics="true" pageId="26" pageNumber="27">Material.</emphasis>
&gt;
<specimenCount id="9D2467826473FFABFC24EDFBD47DFEE7" box="[936,1100,235,259]" count="100" pageId="26" pageNumber="27" type="generic">100 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806473FFABFCA1EC3DD52CFDA0" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8B9DAC0B6473FFABFCA1EC3DD52CFDA0" blockId="26.[813,1438,300,1605]" pageId="26" pageNumber="27">
<emphasis id="B95670196473FFABFCA1EC3DD3ABFEA0" box="[813,922,301,324]" italics="true" pageId="26" pageNumber="27">Diagnosis</emphasis>
(Zhao and Orchard, in
<bibRefCitation id="EFB3D1FA6473FFABFB4BEC3CD5A7FEA0" author="Zhao" box="[1223,1430,300,324]" etAl="et al." firstAuthor="Zhao" pageId="26" pageNumber="27" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al., 2007</bibRefCitation>
; emended by Goudemand, in
<bibRefCitation id="EFB3D1FA6473FFABFBFFEC5CD5BCFE80" author="Goudemand" box="[1139,1421,332,356]" etAl="et al." firstAuthor="Goudemand" pageId="26" pageNumber="27" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
). Small segminate P
<subScript id="17A6AE4E6473FFABFC77EC69D435FE6E" attach="left" box="[1019,1028,377,394]" fontSize="7" pageId="26" pageNumber="27">1</subScript>
elements characterized by a length/ height ratio in the range of 1.322.34, and about 49 robust, wide, mostly fused, and distinctively posteriorly recurved denticles. In lateral view, the basal margin is usually straight. A large, broadly expanded oval to sub-rounded basal cavity is upturned on the inner margin and flat to downturned on the outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806473FFADFCA1EF5CD2A8FDA7" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8B9DAC0B6473FFABFCA1EF5CD3F8F9A1" blockId="26.[813,1438,300,1605]" pageId="26" pageNumber="27">
<emphasis id="B95670196473FFABFCA1EF5CD3A0FD80" box="[813,913,588,612]" italics="true" pageId="26" pageNumber="27">Remarks.</emphasis>
Goudemand (in
<bibRefCitation id="EFB3D1FA6473FFABFBEAEF5CD5A5FD80" author="Goudemand" box="[1126,1428,588,612]" etAl="et al." firstAuthor="Goudemand" pageId="26" pageNumber="27" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
) revised the original diagnosis by noticing that the basal margin is not necessarily straight and therefore, the most strikingly difference to
<taxonomicName id="4C22D7886473FFABFBBCEFBDD487FD20" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1072,1206,685,708]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196473FFABFBBCEFBDD487FD20" box="[1072,1206,685,708]" italics="true" pageId="26" pageNumber="27">Nv. waageni</emphasis>
</taxonomicName>
is the denticulation: the denticles axes are distinctively recurved posteriorly whereas in
<taxonomicName id="4C22D7886473FFABFC22EFFDD405FCE0" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[942,1076,749,772]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196473FFABFC22EFFDD405FCE0" box="[942,1076,749,772]" italics="true" pageId="26" pageNumber="27">Nv. waageni</emphasis>
</taxonomicName>
and
<emphasis id="B95670196473FFABFBFCEFFDD537FCE0" box="[1136,1286,748,772]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="4C22D7886473FFABFBFCEFFDD530FCE0" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[1136,1281,748,772]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="abruptus">Nv. abruptus</taxonomicName>
,
</emphasis>
the denticles are straight, inclined or radiating. Te two or three denticles anterior of the cusp are often clearly asymmetrical and the posterior edge of the element is much shorter than the anterior one. In most sections worldwide,
<taxonomicName id="4C22D7886473FFABFAF5EE7DD3C0FC41" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFAF5EE7DD3C0FC41" italics="true" pageId="26" pageNumber="27">Nv. pingdingshanensis</emphasis>
</taxonomicName>
first occurs within the positive δ
<superScript id="7C5701436473FFABFADBEE98D558FC7D" attach="both" box="[1367,1385,904,921]" fontSize="7" pageId="26" pageNumber="27">13</superScript>
<subScript id="17A6AE4E6473FFABFAE5EE9DD5ADFC4E" attach="left" box="[1385,1436,909,938]" fontSize="7" pageId="26" pageNumber="27">Ccarb</subScript>
excursion of the latest Smithian and may extend to the earliest Spathian (
<bibRefCitation id="EFB3D1FA6473FFABFC70EEDDD522FC01" author="Goudemand" box="[1020,1299,973,997]" etAl="et al." firstAuthor="Goudemand" pageId="26" pageNumber="27" pagination="169 - 178" refId="ref41269" refString="Goudemand, N., Romano, C., Leu, M., Bucher, H., Trotter, J. A., &amp; Williams, I. S. (2019). Dynamic interplay between climate and marine biodiversity upheavals during the early Triassic Smithian-Spathian biotic crisis. Earth-Science Reviews, 195, 169 - 178. https: // doi. org / 10.1016 / j. earscirev. 2019.01. 013" type="journal article" year="2019">Goudemand et al., 2019</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA6473FFABFAAAEEDDD350FBE1" author="Leu" etAl="et al." firstAuthor="Leu" pageId="26" pageNumber="27" pagination="52 - 67" refId="ref43752" refString="Leu, M., Bucher, H., &amp; Goudemand, N. (2019). Clade-dependent size response of conodonts to environmental changes during the late Smithian extinction. Earth-Science Reviews, 195, 52 - 67. https: // doi. org / 10.1016 / j. earscirev. 2018.11. 003" type="journal article" year="2019">Leu et al., 2019</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA6473FFABFCE2EEFDD403FBE1" author="Zhang" box="[878,1074,1005,1029]" etAl="et al." firstAuthor="Zhang" pageId="26" pageNumber="27" refId="ref48781" refString="Zhang, L., Orchard, M. J., Brayard, A., Algeo, T. J., Zhao, L., Chen, Z. Q., &amp; Lyu, Z. (2019). The Smithian / Spathian boundary (late Early Triassic): A review" type="book" year="2019">Zhang et al., 2019</bibRefCitation>
). Some elements within our rich material resemble
<taxonomicName id="4C22D7886473FFABFB80E91DD534FBC1" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[1036,1285,1037,1061]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFB80E91DD534FBC1" box="[1036,1285,1037,1061]" italics="true" pageId="26" pageNumber="27">Ns. Pingdingshanensis</emphasis>
</taxonomicName>
, but appear to have a relatively small basal cavity, more posteriorly recurved denticles or more numerous denticles that what was previously described for this species. Tese elements (
<figureCitation id="1319B08E6473FFABFCB8E99CD3C2FB40" box="[820,1011,1164,1189]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.13127510" httpUri="https://zenodo.org/record/13127510/files/figure.png" pageId="26" pageNumber="27">Figs. 17C, J, K, T</figureCitation>
;
<figureCitation id="1319B08E6473FFABFC73E99DD452FB40" box="[1023,1123,1165,1189]" captionStart="Fig" captionStartId="28.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@29.[228,1362,254,1763]" captionTargetPageId="29" captionText="Fig. 18 Novispathodinae from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, B, D, F, LN, Q, S, T Novispathodus praebrevissimus n. sp.; A LIL506,PIMUZ 39274; B LAR207, PIMUZ 39275; D LIL508, PIMUZ 39276; F LIL507, PIMUZ 39277;L LIL507, PIMUZ 39278; M LIL509, PIMUZ 39279; N LIL507,PIMUZ 39280; Q LIL507, PIMUZ 39281; S LIL507, PIMUZ 39282; T LIL507, PIMUZ 39283. C, O, R Novispathodus?praebrevissimus n. sp.; C LAR204,PIMUZ 39284; O LIL507, PIMUZ 39285; R LIL507, PIMUZ 39286. E, I Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard); E LIL507,PIMUZ 39232, I LIL507, PIMUZ 39233. G Triassospathodus aff. symmetricus (Orchard);LIL507, PIMUZ 39309. H Novispathodus ex gr. abruptus (Orchard); LIL507, PIMUZ 39212. J, K Novispathodus praebrevissimus (juvenile) n. sp.; J LIL507, PIMUZ 39287; K LIL507,PIMUZ 39288. P Novispathodus robustispinus (Zhao &amp; Orchard); LIL507,PIMUZ 39290" figureDoi="http://doi.org/10.5281/zenodo.13127512" httpUri="https://zenodo.org/record/13127512/files/figure.png" pageId="26" pageNumber="27">18E, IK</figureCitation>
;
<figureCitation id="1319B08E6473FFABFBFCE99DD4F3FB41" box="[1136,1218,1165,1189]" captionStart="Fig" captionStartId="30.[169,200,1700,1720]" captionTargetBox="[153,1439,238,1918]" captionTargetId="figure-16@30.[229,1363,254,1687]" captionTargetPageId="30" captionText="Fig. 19 Novispathodinae from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A Novispathodus cf.brevissimus (Orchard); LIL515C,PIMUZ 39199. B, F Novispathodus brevissimus (Orchard); B LIL515B, PIMUZ 39200; F BAN1, PIMUZ 39201. C Triassospathodus homeri (Bender); LIL515D,PIMUZ 39317. D, H Novispathodus?brevissimus (Orchard); D LIL515D,PIMUZ 39202; H LIL514A, PIMUZ 39203. E Triassospathodus cf.homeri (Bender); LIL515C, PIMUZ 39318. G, K, OQ Novispathodus n. sp. A; G LIL504,PIMUZ 39251; K LIL504,PIMUZ 39252; O LIL505, PIMUZ 39253; P LIL505, PIMUZ 39254;Q LIL505,PIMUZ 39255. I, L, M, Triassospathodus symmetricus (Orchard); I QIA141, PIMUZ 39336; L LIL512, PIMUZ 39337; M LIL514A, PIMUZ 39338. J Novispathodus ex gr.pingdingshanensis (Zhao &amp; Orchard); QIA138, PIMUZ 39237. N Triassospathodus cf. symmetricus; LIL514B, PIMUZ 39339.R Novispathodus praebrevissimus n.sp; LIL505, PIMUZ 39289" figureDoi="http://doi.org/10.5281/zenodo.12000347" httpUri="https://zenodo.org/record/12000347/files/figure.png" pageId="26" pageNumber="27">19O, Q</figureCitation>
;
<figureCitation id="1319B08E6473FFABFB43E99DD551FB41" box="[1231,1376,1165,1189]" captionStart="Fig" captionStartId="31.[169,200,1715,1735]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-16@32.[229,1363,254,1763]" captionTargetPageId="32" captionText="Fig. 20 Novispathodinae from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AG, KM, Q, X Novispathodus ex gr. abruptus (Orchard); A LAR212, PIMUZ 39213; B LAR207, PIMUZ 39214; C LAR205,PIMUZ 39215; D LAR202, PIMUZ 39216;E LAR202, PIMUZ 39217; F LIL506, PIMUZ 39218; G LAR210, PIMUZ 39219; K LIL505,PIMUZ 39220; L LIL506, PIMUZ 39221; M LIL505,PIMUZ 39222; Q LAR203,PIMUZ 39223; X QIA136,PIMUZ 39224. H, V Novispathodus sp. indet.; H QIA136, PIMUZ 39291; V, QIA134, PIMUZ 39292. I, J, P, AA Novispathodus n. sp. Z; I QIA138, PIMUZ 39247;J QIA138,PIMUZ 39248; P LIL506, PIMUZ 39249; AA QIA136, PIMUZ 39250. N, O, S Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard); N LIL506, PIMUZ 39234; O LIL505, PIMUZ 39235; S QIA136, PIMUZ 39236. R sp.indet.; LIL511, PIMUZ 39296. T, Y, Z Novispathodus n. sp. A; T QIA134,PIMUZ 39256,Y QIA135, PIMUZ 39257; Z QIA134, PIMUZ 39258. U Novispathodus expansus (Zhao &amp; Orchard); QIA134, PIMUZ 39242. W Novispathodus gryphus n. sp.; QIA135, PIMUZ 39246" figureDoi="http://doi.org/10.5281/zenodo.13127514" httpUri="https://zenodo.org/record/13127514/files/figure.png" pageId="26" pageNumber="27">20H, NO, S</figureCitation>
; here assigned to
<emphasis id="B95670196473FFABFC23E9BDD3F8FB20" box="[943,969,1197,1220]" italics="true" pageId="26" pageNumber="27">N.</emphasis>
ex gr.
<taxonomicName id="4C22D7886473FFABFB9EE9BCD4E7FB20" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[1042,1238,1196,1220]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFB9EE9BCD4E7FB20" box="[1042,1238,1196,1220]" italics="true" pageId="26" pageNumber="27">pingdingshanensis</emphasis>
</taxonomicName>
) may deserve differentiation in the future. Two elements illustrated by Metcalfe et al., (2013, figs. 9.6, 9.9, p. 1144) as
<taxonomicName id="4C22D7886473FFABFAACE9FDD385FAC0" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni" status="subsp. nov.">
<emphasis id="B95670196473FFABFAACE9FDD385FAC0" italics="true" pageId="26" pageNumber="27">Neospathodus waageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFC31E81DD406FAC0" box="[957,1079,1293,1317]" pageId="26" pageNumber="27" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
A display a higher anterior end and more recurved denticles than the other elements they included in
<taxonomicName id="4C22D7886473FFABFC7AE85DD4C2FA80" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1014,1267,1357,1381]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni" status="subsp. nov.">
<emphasis id="B95670196473FFABFC7AE85DD4C2FA80" box="[1014,1267,1357,1381]" italics="true" pageId="26" pageNumber="27">Neospathodus waageni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFB73E85DD54DFA80" box="[1279,1404,1357,1381]" pageId="26" pageNumber="27" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
A, which we synonymized with
<taxonomicName id="4C22D7886473FFABFBF9E87DD554FA61" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[1141,1381,1389,1413]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFBF9E87DD554FA61" box="[1141,1381,1389,1413]" italics="true" pageId="26" pageNumber="27">Ns. pingdingshanensis</emphasis>
</taxonomicName>
: elements like these two elements do not seem to fit in
<taxonomicName id="4C22D7886473FFABFAF6E89DD3C0FA21" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFAF6E89DD3C0FA21" italics="true" pageId="26" pageNumber="27">Ns. pingdingshanensis</emphasis>
</taxonomicName>
and may deserve assignment to a new species. Some further elements resembling
<taxonomicName id="4C22D7886473FFABFA80E8DDD3A5F9E1" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFA80E8DDD3A5F9E1" italics="true" pageId="26" pageNumber="27">Ns. pingdingshanensis</emphasis>
</taxonomicName>
but distinctively shorter than
<taxonomicName id="4C22D7886473FFABFB56E8FDD350F9C0" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFB56E8FDD350F9C0" italics="true" pageId="26" pageNumber="27">Ns. pingdingshanensis</emphasis>
</taxonomicName>
are herein assigned to a new species (
<taxonomicName id="4C22D7886473FFABFA8DEB1DD5B0F9C1" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1281,1409,1549,1573]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gryphus" status="n. sp.">
<emphasis id="B95670196473FFABFA8DEB1DD5B0F9C1" box="[1281,1409,1549,1573]" italics="true" pageId="26" pageNumber="27">Nv. gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFA04EB1DD37AF9A0" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
see below).
</paragraph>
<subSection id="E2ADB5E06473FFABFF25EB74D1B2F99C" box="[169,387,1636,1656]" pageId="26" pageNumber="27" type="multiple">
<paragraph id="8B9DAC0B6473FFABFF25EB74D1B2F99C" blockId="26.[169,387,1636,1656]" box="[169,387,1636,1656]" pageId="26" pageNumber="27">(See figure on next page.)</paragraph>
</subSection>
<caption id="DF5DFC836473FFABFF25EB6FD2C9F867" ID-DOI="http://doi.org/10.5281/zenodo.13127510" ID-Zenodo-Dep="13127510" httpUri="https://zenodo.org/record/13127510/files/figure.png" pageId="26" pageNumber="27" startId="26.[169,200,1663,1683]" targetBox="[152,1438,238,1791]" targetPageId="27">
<paragraph id="8B9DAC0B6473FFABFF25EB6FD2C9F867" blockId="26.[169,1414,1662,1927]" pageId="26" pageNumber="27">
<figureCitation id="1319B08E6473FFABFF25EB6FD0D9F977" box="[169,232,1663,1683]" captionStart="Fig" captionStartId="27.[169,200,1766,1786]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000312" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="26" pageNumber="27">
<emphasis id="B95670196473FFABFF25EB6FD0D9F977" bold="true" box="[169,232,1663,1683]" pageId="26" pageNumber="27">Fig. 17</emphasis>
</figureCitation>
<taxonomicName id="4C22D7886473FFABFF7EEB6FD1B2F977" authorityName="Orchard" authorityYear="2005" box="[242,387,1663,1683]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E6473FFABFF69EBB3D0DDF955" attach="left" box="[229,236,1699,1713]" fontSize="6" pageId="26" pageNumber="27">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B95670196473FFABFD05EB8AD29FF94A" bold="true" box="[649,686,1690,1710]" pageId="26" pageNumber="27">AC</emphasis>
,
<emphasis id="B95670196473FFABFD3AEB8AD28FF94A" bold="true" box="[694,702,1690,1710]" pageId="26" pageNumber="27">J</emphasis>
,
<emphasis id="B95670196473FFABFD4BEB8AD2E3F94A" bold="true" box="[711,722,1690,1710]" pageId="26" pageNumber="27">T</emphasis>
,
<emphasis id="B95670196473FFABFD57EB8AD2C7F94A" bold="true" box="[731,758,1690,1710]" pageId="26" pageNumber="27">AA</emphasis>
<taxonomicName id="4C22D7886473FFABFD77EB8AD347F94A" authorityName="Orchard" authorityYear="2005" box="[763,886,1690,1710]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFD77EB8AD347F94A" box="[763,886,1690,1710]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886473FFABFC27EB8AD4EAF94A" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[939,1243,1690,1710]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFC27EB8AD475F94A" box="[939,1092,1690,1710]" italics="true" pageId="26" pageNumber="27">pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
;
<emphasis id="B95670196473FFABFB6EEB8AD4C1F94A" bold="true" box="[1250,1264,1690,1710]" pageId="26" pageNumber="27">A</emphasis>
, QIA138,PIMUZ 39226;
<emphasis id="B95670196473FFABFF6AEBA4D0C3F92C" bold="true" box="[230,242,1716,1736]" pageId="26" pageNumber="27">B</emphasis>
QIA138, PIMUZ 39227;
<emphasis id="B95670196473FFABFE3BEBA4D1F5F92C" bold="true" box="[439,452,1716,1736]" pageId="26" pageNumber="27">C</emphasis>
LAR202, PIMUZ 39228;
<emphasis id="B95670196473FFABFD07EBA4D2A2F92C" bold="true" box="[651,659,1716,1736]" pageId="26" pageNumber="27">J</emphasis>
QIA136, PIMUZ 39229;
<emphasis id="B95670196473FFABFCD4EBA4D352F92C" bold="true" box="[856,867,1716,1736]" pageId="26" pageNumber="27">T</emphasis>
LIL504, PIMUZ 39230;
<emphasis id="B95670196473FFABFBADEBA4D40DF92C" bold="true" box="[1057,1084,1716,1736]" pageId="26" pageNumber="27">AA</emphasis>
LIL508,PIMUZ 39231.
<emphasis id="B95670196473FFABFB76EBA4D52FF92C" bold="true" box="[1274,1310,1716,1736]" pageId="26" pageNumber="27">DF</emphasis>
,
<emphasis id="B95670196473FFABFAABEBA4D57EF92C" bold="true" box="[1319,1359,1716,1736]" pageId="26" pageNumber="27">MP</emphasis>
,
<emphasis id="B95670196473FFABFAD4EBA4D555F92C" bold="true" box="[1368,1380,1716,1736]" pageId="26" pageNumber="27">R</emphasis>
,
<emphasis id="B95670196473FFABFAE0EBA4D546F92C" bold="true" box="[1388,1399,1716,1736]" pageId="26" pageNumber="27">S</emphasis>
,
<emphasis id="B95670196473FFABFF25EBDFD086F907" bold="true" box="[169,183,1743,1763]" pageId="26" pageNumber="27">U</emphasis>
,
<emphasis id="B95670196473FFABFF4CEBDFD0FCF907" bold="true" box="[192,205,1743,1763]" pageId="26" pageNumber="27">X</emphasis>
,
<emphasis id="B95670196473FFABFF59EBDFD0DEF907" bold="true" box="[213,239,1743,1763]" pageId="26" pageNumber="27">AB</emphasis>
,
<emphasis id="B95670196473FFABFF7BEBDFD122F907" bold="true" box="[247,275,1743,1763]" pageId="26" pageNumber="27">AD</emphasis>
<taxonomicName id="4C22D7886473FFABFE94EBDFD2F7F907" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[280,710,1743,1763]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFE94EBDFD21EF907" box="[280,559,1743,1763]" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
;
<emphasis id="B95670196473FFABFD42EBDFD2ECF907" bold="true" box="[718,733,1743,1763]" pageId="26" pageNumber="27">D</emphasis>
QIA135, PIMUZ 39259;
<emphasis id="B95670196473FFABFC2EEBDFD39CF907" bold="true" box="[930,941,1743,1763]" pageId="26" pageNumber="27">E</emphasis>
LIL508,PIMUZ 39260;
<emphasis id="B95670196473FFABFBE6EBDFD444F907" bold="true" box="[1130,1141,1743,1763]" pageId="26" pageNumber="27">F</emphasis>
LIL506, PIMUZ 39261;
<emphasis id="B95670196473FFABFABFEBDFD574F907" bold="true" box="[1331,1349,1743,1763]" pageId="26" pageNumber="27">M</emphasis>
LIL506, PIMUZ 39262;
<emphasis id="B95670196473FFABFEAFEBFAD100F91A" bold="true" box="[291,305,1770,1790]" pageId="26" pageNumber="27">N</emphasis>
LIL506,PIMUZ 39263;
<emphasis id="B95670196473FFABFE63EBFAD1CFF91A" bold="true" box="[495,510,1770,1790]" pageId="26" pageNumber="27">O</emphasis>
LIL507, PIMUZ 39264;
<emphasis id="B95670196473FFABFD30EBFAD2F9F91A" bold="true" box="[700,712,1770,1790]" pageId="26" pageNumber="27">P</emphasis>
LIL506, PIMUZ 39265;
<emphasis id="B95670196473FFABFC09EBFAD3A0F91A" bold="true" box="[901,913,1770,1790]" pageId="26" pageNumber="27">R</emphasis>
LIL508,PIMUZ 39266;
<emphasis id="B95670196473FFABFBC3EBFAD46BF91A" bold="true" box="[1103,1114,1770,1790]" pageId="26" pageNumber="27">S</emphasis>
LAR204,PIMUZ 39267;
<emphasis id="B95670196473FFABFAADEBFAD51EF91A" bold="true" box="[1313,1327,1770,1790]" pageId="26" pageNumber="27">U</emphasis>
LIL507, PIMUZ 39268;
<emphasis id="B95670196473FFABFEAFEA14D101F8FC" bold="true" box="[291,304,1796,1816]" pageId="26" pageNumber="27">X</emphasis>
LIL508, PIMUZ 39269;
<emphasis id="B95670196473FFABFE61EA14D236F8FC" bold="true" box="[493,519,1796,1816]" pageId="26" pageNumber="27">AB</emphasis>
LIL508, PIMUZ 39270;
<emphasis id="B95670196473FFABFD49EA14D2D0F8FC" bold="true" box="[709,737,1796,1816]" pageId="26" pageNumber="27">AD</emphasis>
LIL508, PIMUZ 39271.
<emphasis id="B95670196473FFABFC13EA14D39CF8FC" bold="true" box="[927,941,1796,1816]" pageId="26" pageNumber="27">G</emphasis>
,
<emphasis id="B95670196473FFABFC39EA14D38AF8FC" bold="true" box="[949,955,1796,1816]" pageId="26" pageNumber="27">I</emphasis>
,
<emphasis id="B95670196473FFABFC4FEA14D3E3F8FC" bold="true" box="[963,978,1796,1816]" pageId="26" pageNumber="27">Q</emphasis>
<taxonomicName id="4C22D7886473FFABFC5BEA14D463F8FC" authorityName="Orchard" authorityYear="2005" box="[983,1106,1796,1816]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFC5BEA14D463F8FC" box="[983,1106,1796,1816]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C22D7886473FFABFBE1EA14D486F8FC" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1133,1207,1796,1816]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gryphus" status="sp. nov.">
?
<emphasis id="B95670196473FFABFBF9EA14D486F8FC" box="[1141,1207,1796,1816]" italics="true" pageId="26" pageNumber="27">gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFB30EA14D4DBF8FC" box="[1212,1258,1796,1816]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196473FFABFB63EA14D4CCF8FC" bold="true" box="[1263,1277,1796,1816]" pageId="26" pageNumber="27">G</emphasis>
QIA136, PIMUZ 39204;
<emphasis id="B95670196473FFABFF6AEA0FD0DDF8D7" bold="true" box="[230,236,1823,1843]" pageId="26" pageNumber="27">I</emphasis>
QIA136, PIMUZ 39205;
<emphasis id="B95670196473FFABFE3DEA0FD1F1F8D7" bold="true" box="[433,448,1823,1843]" pageId="26" pageNumber="27">Q</emphasis>
QIA135, PIMUZ 39206.
<emphasis id="B95670196473FFABFD09EA0FD2A2F8D7" bold="true" box="[645,659,1823,1843]" pageId="26" pageNumber="27">H</emphasis>
,
<emphasis id="B95670196473FFABFD10EA0FD299F8D7" bold="true" box="[668,680,1823,1843]" pageId="26" pageNumber="27">K</emphasis>
,
<emphasis id="B95670196473FFABFD3DEA0FD28DF8D7" bold="true" box="[689,700,1823,1843]" pageId="26" pageNumber="27">L</emphasis>
<taxonomicName id="4C22D7886473FFABFD4CEA0FD3B1F8D7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[704,896,1823,1843]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gryphus" status="sp. nov.">
<emphasis id="B95670196473FFABFD4CEA0FD3B1F8D7" box="[704,896,1823,1843]" italics="true" pageId="26" pageNumber="27">Novispathodus gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFC09EA0FD383F8D7" box="[901,946,1823,1843]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196473FFABFC34EA0FD3F7F8D7" bold="true" box="[952,966,1823,1843]" pageId="26" pageNumber="27">H</emphasis>
QIA135, PIMUZ 39243;
<emphasis id="B95670196473FFABFB00EA0FD4A9F8D7" bold="true" box="[1164,1176,1823,1843]" pageId="26" pageNumber="27">K</emphasis>
LIL506, PIMUZ 39244;
<emphasis id="B95670196473FFABFADAEA0FD550F8D7" bold="true" box="[1366,1377,1823,1843]" pageId="26" pageNumber="27">L</emphasis>
LIL506,PIMUZ 39245.
<emphasis id="B95670196473FFABFEEEEA2AD15EF8AA" bold="true" box="[354,367,1850,1870]" pageId="26" pageNumber="27">
<collectionCode id="ED3334CE6473FFABFEEEEA2AD15EF8AA" box="[354,367,1850,1870]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="27" type="Museum">V</collectionCode>
</emphasis>
<taxonomicName id="4C22D7886473FFABFEF8EA2AD245F8AA" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[372,628,1850,1870]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="praebrevissimus" status="sp. nov.">
<emphasis id="B95670196473FFABFEF8EA2AD245F8AA" box="[372,628,1850,1870]" italics="true" pageId="26" pageNumber="27">Novispathodus praebrevissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626473FFABFDF4EA2AD297F8AA" box="[632,678,1850,1870]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
; LIL507, PIMUZ 39273.
<emphasis id="B95670196473FFABFCE9EA2AD349F8AA" bold="true" box="[869,888,1850,1870]" pageId="26" pageNumber="27">W</emphasis>
,
<emphasis id="B95670196473FFABFC0CEA2AD3BDF8AA" bold="true" box="[896,908,1850,1870]" pageId="26" pageNumber="27">Y</emphasis>
,
<emphasis id="B95670196473FFABFC18EA2AD391F8AA" bold="true" box="[916,928,1850,1870]" pageId="26" pageNumber="27">Z</emphasis>
,
<emphasis id="B95670196473FFABFC25EA2AD3F3F8AA" bold="true" box="[937,962,1850,1870]" pageId="26" pageNumber="27">AE</emphasis>
<taxonomicName id="4C22D7886473FFABFC4AEA2AD470F8AA" authorityName="Orchard" authorityYear="2005" box="[966,1089,1850,1870]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFC4AEA2AD470F8AA" box="[966,1089,1850,1870]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886473FFABFBFBEA2AD527F8AA" authority="(Orchard)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[1143,1302,1850,1870]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196473FFABFBFBEA2AD4F0F8AA" box="[1143,1217,1850,1870]" italics="true" pageId="26" pageNumber="27">abruptus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196473FFABFA91EA2AD501F8AA" bold="true" box="[1309,1328,1850,1870]" pageId="26" pageNumber="27">W</emphasis>
LIL505, PIMUZ 39208;
<emphasis id="B95670196473FFABFEAFEA44D11EF88C" bold="true" box="[291,303,1876,1896]" pageId="26" pageNumber="27">Y</emphasis>
SHA346,PIMUZ 39209;
<emphasis id="B95670196473FFABFE75EA44D234F88C" bold="true" box="[505,517,1876,1896]" pageId="26" pageNumber="27">Z</emphasis>
LIL506, PIMUZ 39210;
<emphasis id="B95670196473FFABFD4FEA44D2EDF88C" bold="true" box="[707,732,1876,1896]" pageId="26" pageNumber="27">AE</emphasis>
SHA346, PIMUZ 39211.
<emphasis id="B95670196473FFABFC29EA44D38EF88C" bold="true" box="[933,959,1876,1896]" pageId="26" pageNumber="27">AC</emphasis>
sp. indet.; LIL504, PIMUZ 39295.
<emphasis id="B95670196473FFABFB5DEA44D4D8F88C" bold="true" box="[1233,1257,1876,1896]" pageId="26" pageNumber="27">AF</emphasis>
<taxonomicName id="4C22D7886473FFABFB62EA44D173F867" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFB62EA44D173F867" italics="true" pageId="26" pageNumber="27">Novispathodus pingdingshanensis</emphasis>
</taxonomicName>
(P
<subScript id="17A6AE4E6473FFABFEDBEA69D16FF863" attach="left" box="[343,350,1913,1927]" fontSize="6" pageId="26" pageNumber="27">1</subScript>
cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272
</paragraph>
</caption>
<caption id="DF5DFC836472FFAAFF25EBF6D1C8F91E" ID-DOI="http://doi.org/10.5281/zenodo.12000312" ID-Zenodo-Dep="12000312" box="[169,505,1766,1786]" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="27" pageNumber="28" startId="27.[169,200,1766,1786]" targetBox="[152,1438,238,1791]" targetPageId="27" targetType="figure">
<paragraph id="8B9DAC0B6472FFAAFF25EBF6D1C8F91E" blockId="27.[169,505,1766,1786]" box="[169,505,1766,1786]" pageId="27" pageNumber="28">
<emphasis id="B95670196472FFAAFF25EBF6D0D6F91E" bold="true" box="[169,231,1766,1786]" pageId="27" pageNumber="28">Fig. 17</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<paragraph id="8B9DAC0B6475FFADFF21EDFCD2A8FDA7" blockId="28.[151,775,235,1667]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF21EDFCD119FEE7" box="[173,296,236,259]" italics="true" pageId="28" pageNumber="29">Occurrence</emphasis>
. Worldwide occurrence.
<collectingCountry id="F335EC9B6475FFADFDE5EDFBD29FFEE7" box="[617,686,235,259]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
: Jinya/ Waili area,
<collectingRegion id="49E662E96475FFADFE94EC1BD144FEC7" box="[280,373,267,291]" country="China" name="Guangxi" pageId="28" pageNumber="29">Guangxi</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6475FFADFE0AEC1BD290FEC7" author="Goudemand" box="[390,673,267,291]" etAl="et al." firstAuthor="Goudemand" pageId="28" pageNumber="29" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
) Jiarong, southern
<collectingRegion id="49E662E96475FFADFE8CEC3BD16EFEA7" box="[256,351,299,323]" country="China" name="Guizhou" pageId="28" pageNumber="29">Guizhou</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6475FFADFEE1EC3BD215FEA7" author="Chen" box="[365,548,299,323]" etAl="et al." firstAuthor="Chen" pageId="28" pageNumber="29" pagination="823 - 826" refId="ref39673" refString="Chen, Y., Twitchett, R. J., Jiang, H., Richoz, S., Lai, X., Yan, C., Sun, Y., Liu, X., &amp; Wang, L. (2013). Size variation of conodonts during the Smithian-Spathian (Early Triassic) global warming event. Geology, 41 (8), 823 - 826. https: // doi. org / 10.1130 / G 34171.1" type="journal article" year="2013">Chen et al., 2013</bibRefCitation>
,
<bibRefCitation id="EFB3D1FA6475FFADFDBDEC3BD254FEA7" author="Chen" box="[561,613,299,323]" firstAuthor="Chen" pageId="28" pageNumber="29" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">2015</bibRefCitation>
). Anshun Fm., Qingyan section,
<collectingRegion id="49E662E96475FFADFEDBEC5BD187FE87" box="[343,438,331,355]" country="China" name="Guizhou" pageId="28" pageNumber="29">Guizhou</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6475FFADFE48EC5BD263FE87" author="Ji" box="[452,594,331,355]" etAl="et al." firstAuthor="Ji" pageId="28" pageNumber="29" pagination="213 - 223" refId="ref42283" refString="Ji, W., Tong, J., Zhao, L., Zhou, S., &amp; Chen, J. (2011). Lower - middle Triassic conodont biostratigraphy of the Qingyan section, Guizhou province, Southwest China. Palaeogeography, Palaeoclimatology, Palaeoecology, 308 (1 - 2), 213 - 223. https: // doi. org / 10.1016 / j. palaeo. 2010.08. 020" type="journal article" year="2011">Ji et al., 2011</bibRefCitation>
). Chaohu (Zhao et al., 2007, 2008). Daxiakou,
<collectingRegion id="49E662E96475FFADFE6AEC7BD218FE67" box="[486,553,363,387]" country="China" name="Hubei" pageId="28" pageNumber="29">Hubei</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6475FFADFDB6EC7BD2C9FE67" author="Zhao" box="[570,760,363,387]" etAl="et al." firstAuthor="Zhao" pageId="28" pageNumber="29" pagination="523 - 540" refId="ref48953" refString="Zhao, L., Chen, Y., Chen, Z. Q., &amp; Cao, L. (2013). Uppermost Permian to Lower Triassic Conodont Zonation from Three Gorges Area, South China. Palaios, 28 (8), 523 - 540. https: // doi. org / 10.2110 / palo. 2012. p 12 - 107 r" type="journal article" year="2013">Zhao et al., 2013</bibRefCitation>
).
<collectingCountry id="F335EC9B6475FFADFF1BEC9BD0C6FE47" box="[151,247,395,419]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
: Bac Tuy Fm.,
<taxonomicName id="4C22D7886475FFADFE3AEC9BD2E2FE47" authorityName="Brayard &amp; Bucher" authorityYear="2008" box="[438,723,395,419]" class="Cephalopoda" family="Xenoceltitidae" genus="Xenoceltites" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="variocostatus">
<emphasis id="B95670196475FFADFE3AEC9BD2E2FE47" box="[438,723,395,419]" italics="true" pageId="28" pageNumber="29">Xenoceltites variocostatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886475FFADFF1BECBBD0C1FE27" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,240,427,451]" class="Cephalopoda" family="Columbitidae" genus="Tirolites" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="B95670196475FFADFF1BECBBD0C1FE27" box="[151,240,427,451]" italics="true" pageId="28" pageNumber="29">Tirolites</emphasis>
</taxonomicName>
beds, (
<bibRefCitation id="EFB3D1FA6475FFADFEB3ECBBD22FFE27" author="Komatsu" box="[319,542,427,451]" etAl="et al." firstAuthor="Komatsu" pageId="28" pageNumber="29" pagination="65 - 74" refId="ref43220" refString="Komatsu, T., Takashima, R., Shigeta, Y., Maekawa, T., Tran, H. D., Cong, T. D., Sakata, S., Dinh, H. D., &amp; Takahashi, O. (2016). Carbon isotopic excursions and detailed ammonoid and conodont biostratigraphies around Smithian-Spathian boundary in the Bac Thuy Formation, Vietnam. Palaeogeography, Palaeoclimatology, Palaeoecology, 454, 65 - 74. https: // doi. org / 10.1016 / j. palaeo. 2016.04. 017" type="journal article" year="2016">Komatsu et al., 2016</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA6475FFADFDA0ECBBD2C9FE27" author="Shigeta" box="[556,760,427,451]" etAl="et al." firstAuthor="Shigeta" pageId="28" pageNumber="29" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
).
<collectingCountry id="F335EC9B6475FFADFF1BECDBD0DCFE07" box="[151,237,459,483]" name="Canada" pageId="28" pageNumber="29">Canada</collectingCountry>
:
<taxonomicName id="4C22D7886475FFADFE8DECDBD22FFE07" baseAuthorityName="Kozur &amp; Mostler" baseAuthorityYear="1976" box="[257,542,459,483]" family="Gondolellidae" genus="Scythogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="mosheri">
<emphasis id="B95670196475FFADFE8DECDBD22FFE07" box="[257,542,459,483]" italics="true" pageId="28" pageNumber="29">Scythogondolella mosheri</emphasis>
</taxonomicName>
zone, Wapiti Lake (
<bibRefCitation id="EFB3D1FA6475FFADFF13ECFBD1E4FDE7" author="Orchard &amp; Zonneveld" box="[159,469,491,515]" firstAuthor="Orchard" pageId="28" pageNumber="29" refId="ref45906" refString="Orchard, M. J., &amp; Zonneveld, J. P. (2009). The Lower Triassic Sulphur Moun-" type="book" year="2009">Orchard &amp; Zonneveld, 2009</bibRefCitation>
).
<collectingCountry id="F335EC9B6475FFADFE67ECFBD260FDE7" box="[491,593,491,515]" name="Australia" pageId="28" pageNumber="29">Australia</collectingCountry>
: Hovea Member of Kockatea Shale, upper part of
<taxonomicName id="4C22D7886475FFADFD87EF1BD336FDC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[523,775,523,547]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196475FFADFD87EF1BD336FDC7" box="[523,775,523,547]" italics="true" pageId="28" pageNumber="29">Neospathodus waageni</emphasis>
</taxonomicName>
Zone, Smithian substage (
<bibRefCitation id="EFB3D1FA6475FFADFE3CEF3BD2BAFDA7" author="Metcalfe" box="[432,651,555,579]" etAl="et al." firstAuthor="Metcalfe" pageId="28" pageNumber="29" pagination="1136 - 1150" refId="ref45079" refString="Metcalfe, I., Nicoll, R. S., Willink, R., Ladjavadi, M., &amp; Grice, K. (2013). Early Triassic (Induan - Olenekian) conodont biostratigraphy, global anoxia, carbon isotope excursions and environmental perturbations: New data from Western Australian Gondwana. Gondwana Research, 23 (3), 1136 - 1150. https: // doi. org / 10.1016 / j. gr. 2012.07. 002" type="journal article" year="2013">Metcalfe et al., 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,585 @@
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<paragraph id="8B9DAC0B6475FFADFCA1EE7BD51AFC67" blockId="28.[813,1437,235,1667]" box="[813,1323,875,900]" pageId="28" pageNumber="29">
<taxonomicName id="4C22D7886475FFADFCA1EE7BD51AFC67" authority="(Orchard, 1995)" authorityName="Orchard" authorityYear="2005" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[813,1323,875,900]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="B95670196475FFADFCA1EE7BD3FEFC67" box="[813,975,875,899]" italics="true" pageId="28" pageNumber="29">Novispathodus</emphasis>
ex gr.
<emphasis id="B95670196475FFADFB98EE7BD447FC67" box="[1044,1142,875,899]" italics="true" pageId="28" pageNumber="29">abruptus</emphasis>
(
<bibRefCitation id="EFB3D1FA6475FFADFB08EE7BD513FC67" author="Orchard" box="[1156,1314,875,899]" firstAuthor="Orchard" pageId="28" pageNumber="29" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
)
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<subSubSection id="C338FF806475FFADFCA1EE9BD555FC47" box="[813,1380,907,931]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9DAC0B6475FFADFCA1EE9BD555FC47" blockId="28.[813,1437,235,1667]" box="[813,1380,907,931]" pageId="28" pageNumber="29">
<figureCitation id="1319B08E6475FFADFCA1EE9BD3A1FC47" box="[813,912,907,931]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="28" pageNumber="29">Figs. 16E</figureCitation>
;
<figureCitation id="1319B08E6475FFADFC16EE9BD434FC47" box="[922,1029,907,931]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.13127510" httpUri="https://zenodo.org/record/13127510/files/figure.png" pageId="28" pageNumber="29">17W, Y, Z</figureCitation>
, AE; 18H; 20AG, K, L, M, Q, X
</paragraph>
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<paragraph id="8B9DAC0B6475FFADFCA1EEDBD544FC07" blockId="28.[813,1437,235,1667]" box="[813,1397,971,995]" pageId="28" pageNumber="29">
1981
<taxonomicName id="4C22D7886475FFADFCEBEEDBD49BFC07" authority="Bender" authorityName="Bender" box="[871,1194,971,995]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196475FFADFCEBEEDBD462FC07" box="[871,1107,971,995]" italics="true" pageId="28" pageNumber="29">Neospathodus homeri</emphasis>
Bender
</taxonomicName>
; Koike, pl. 1, fig. 5.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1EEFBD37CFBC7" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
1984
<taxonomicName id="4C22D7886475FFADFCE4EEFBD473FBE7" box="[872,1090,1003,1027]" family="Gondolellidae" genus="Neospathodus" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B95670196475FFADFCE4EEFBD430FBE7" box="[872,1025,1003,1027]" italics="true" pageId="28" pageNumber="29">Neospathodus</emphasis>
sp. A
</taxonomicName>
; Hatleberg &amp; Clark, pl. 3, fig. 8, 21.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E93BD44BFB87" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
*1995
<taxonomicName id="4C22D7886475FFADFCFBE93BD44EFBA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[887,1151,1067,1091]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus" status="sp. nov.">
<emphasis id="B95670196475FFADFCFBE93BD44EFBA7" box="[887,1151,1067,1091]" italics="true" pageId="28" pageNumber="29">Neospathodus abruptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFB01E93CD4FFFBA7" box="[1165,1230,1068,1091]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
; Orchard, p. 118, 119, figs. 3.163.19, 3.233.26.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E97BD396FB47" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
2005
<taxonomicName id="4C22D7886475FFADFCE5E97BD4DAFB67" authority="(Orchard)" baseAuthorityName="Orchard" box="[873,1259,1131,1155]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFCE5E97BD445FB67" box="[873,1140,1131,1155]" italics="true" pageId="28" pageNumber="29">Novispathodus abruptus</emphasis>
(Orchard)
</taxonomicName>
; Orchard, p. 90, text-fig. 16.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E9BBD4FDFB07" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
2009
<taxonomicName id="4C22D7886475FFADFCFEE9BBD536FB27" authority="(Orchard)" baseAuthorityName="Orchard" box="[882,1287,1195,1219]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFCFEE9BBD4B7FB27" box="[882,1158,1195,1219]" italics="true" pageId="28" pageNumber="29">Novispathodus abruptus</emphasis>
(Orchard)
</taxonomicName>
; Orchard &amp; Zonneveld, p. 784, fig. 15 parts 3437.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E9FBD529FAC7" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
2012a
<taxonomicName id="4C22D7886475FFADFCF8E9FBD428FAE7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[884,1049,1259,1283]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus" status="sp. nov.">Novispathodus</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFB93E9FCD443FAE7" box="[1055,1138,1260,1283]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
A; Goudemand &amp; Orchard in Goudemand et al., p. 1031, figs. 2A, R?, Z?.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E83BD48DFA87" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
2012a
<taxonomicName id="4C22D7886475FFADFCF8E83BD428FAA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[884,1049,1323,1347]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus" status="sp. nov.">Novispathodus</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFBACE83CD442FAA7" box="[1056,1139,1324,1347]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
B; Goudemand &amp; Orchard in Goudemand et al., p. 1031, fig. 3V.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFCA1E87BD3CBFA27" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
2018
<taxonomicName id="4C22D7886475FFADFCE3E87BD4CEFA67" authority="(Orchard)" baseAuthorityName="Orchard" box="[879,1279,1387,1411]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFCE3E87BD4B0FA67" box="[879,1153,1387,1411]" italics="true" pageId="28" pageNumber="29">Novispathodus abruptus</emphasis>
(Orchard)
</taxonomicName>
; Maekawa in Maekawa et al., p. 33, figs., 18.118.3, 18.20, 18.23, 18.24, 18.26, 18.27 (only).
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFCA1E8FBD474F9E7" box="[813,1093,1515,1539]" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B9DAC0B6475FFADFCA1E8FBD474F9E7" blockId="28.[813,1437,235,1667]" box="[813,1093,1515,1539]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFCA1E8FBD3BFF9E7" box="[813,910,1515,1539]" italics="true" pageId="28" pageNumber="29">Material</emphasis>
.&gt;
<specimenCount id="9D2467826475FFADFC27E8FBD470F9E7" box="[939,1089,1515,1539]" count="50" pageId="28" pageNumber="29" type="generic">50 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFCA1EB3CD4B5F9A7" box="[813,1156,1579,1603]" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B9DAC0B6475FFADFCA1EB3CD4B5F9A7" blockId="28.[813,1437,235,1667]" box="[813,1156,1579,1603]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFCA1EB3CD3A6F9A7" box="[813,919,1580,1603]" italics="true" pageId="28" pageNumber="29">Diagnosis</emphasis>
. As in
<bibRefCitation id="EFB3D1FA6475FFADFC6DEB3BD44FF9A7" author="Orchard" box="[993,1150,1579,1603]" firstAuthor="Orchard" pageId="28" pageNumber="29" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFAEFCA1EB5BD0D0FA07" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B9DAC0B6475FFAEFCA1EB5BD2F4FE47" blockId="28.[813,1437,235,1667]" lastBlockId="31.[151,775,235,1507]" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFCA1EB5BD3BAF987" box="[813,907,1611,1635]" italics="true" pageId="28" pageNumber="29">Remarks</emphasis>
. Here we consider
<taxonomicName id="4C22D7886475FFADFBD4EB5CD4D2F987" box="[1112,1251,1611,1635]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFBD4EB5CD4D2F987" box="[1112,1251,1611,1635]" italics="true" pageId="28" pageNumber="29">Nv. abruptus</emphasis>
</taxonomicName>
in a broad sense, including elements that correspond to
<taxonomicName id="4C22D7886475FFADFB42EB7CD568F967" box="[1230,1369,1643,1667]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFB42EB7CD568F967" box="[1230,1369,1643,1667]" italics="true" pageId="28" pageNumber="29">Nv. abruptus</emphasis>
</taxonomicName>
sensu stricto together with forms that have been suggested to deserve assignment to separate species, such as
<emphasis id="B95670196476FFAEFD36EC1CD2E8FEC7" box="[698,729,268,291]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
. sp. nov. A and
<emphasis id="B95670196476FFAEFE97EC3CD10BFEA7" box="[283,314,300,323]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
. sp. nov. B Goudemand and Orchard (in
<bibRefCitation id="EFB3D1FA6476FFAEFF1BEC5BD185FE87" author="Goudemand" box="[151,436,331,355]" etAl="et al." firstAuthor="Goudemand" pageId="31" pageNumber="32" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
). We consider the most diagnostic feature of
<emphasis id="B95670196476FFAEFED9EC7CD145FE67" box="[341,372,364,387]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
. ex gr.
<taxonomicName id="4C22D7886476FFAEFE4BEC7BD218FE67" box="[455,553,363,387]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFE4BEC7BD218FE67" box="[455,553,363,387]" italics="true" pageId="31" pageNumber="32">abruptus</emphasis>
</taxonomicName>
is the terminal 13 progressively smaller denticles at the posterior end.
</paragraph>
<subSection id="E2ADB5E06475FFADFF25EB8AD1B2F94A" box="[169,387,1690,1710]" pageId="28" pageNumber="29" type="multiple">
<paragraph id="8B9DAC0B6475FFADFF25EB8AD1B2F94A" blockId="28.[169,387,1690,1710]" box="[169,387,1690,1710]" pageId="28" pageNumber="29">(See figure on next page.)</paragraph>
</subSection>
<caption id="DF5DFC836475FFADFF25EBA5D16BF860" ID-DOI="http://doi.org/10.5281/zenodo.13127512" ID-Zenodo-Dep="13127512" httpUri="https://zenodo.org/record/13127512/files/figure.png" pageId="28" pageNumber="29" startId="28.[169,200,1717,1737]" targetBox="[152,1438,238,1801]" targetPageId="29">
<paragraph id="8B9DAC0B6475FFADFF25EBA5D16BF860" blockId="28.[169,1379,1716,1924]" pageId="28" pageNumber="29">
<figureCitation id="1319B08E6475FFADFF25EBA5D0D9F92D" box="[169,232,1717,1737]" captionStart="Fig" captionStartId="29.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@29.[228,1362,254,1763]" captionTargetPageId="29" captionText="Fig. 18 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000320" httpUri="https://zenodo.org/record/12000320/files/figure.png" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF25EBA5D0D9F92D" bold="true" box="[169,232,1717,1737]" pageId="28" pageNumber="29">Fig. 18</emphasis>
</figureCitation>
<taxonomicName id="4C22D7886475FFADFF7BEBA5D1B9F92D" authorityName="Orchard" authorityYear="2005" box="[247,392,1717,1737]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E6475FFADFA88EBAFD53AF929" attach="left" box="[1284,1291,1727,1741]" fontSize="6" pageId="28" pageNumber="29">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B95670196475FFADFE61EBC0D1CAF900" bold="true" box="[493,507,1744,1764]" pageId="28" pageNumber="29">A</emphasis>
,
<emphasis id="B95670196475FFADFD8FEBC0D23EF900" bold="true" box="[515,527,1744,1764]" pageId="28" pageNumber="29">B</emphasis>
,
<emphasis id="B95670196475FFADFD94EBC0D216F900" bold="true" box="[536,551,1744,1764]" pageId="28" pageNumber="29">D</emphasis>
,
<emphasis id="B95670196475FFADFDA3EBC0D20BF900" bold="true" box="[559,570,1744,1764]" pageId="28" pageNumber="29">F</emphasis>
,
<emphasis id="B95670196475FFADFDCEEBC0D254F900" bold="true" box="[578,613,1744,1764]" pageId="28" pageNumber="29">LN</emphasis>
,
<emphasis id="B95670196475FFADFDE2EBC0D24CF900" bold="true" box="[622,637,1744,1764]" pageId="28" pageNumber="29">Q</emphasis>
,
<emphasis id="B95670196475FFADFD0AEBC0D2A0F900" bold="true" box="[646,657,1744,1764]" pageId="28" pageNumber="29">S</emphasis>
,
<emphasis id="B95670196475FFADFD15EBC0D295F900" bold="true" box="[665,676,1744,1764]" pageId="28" pageNumber="29">T</emphasis>
<taxonomicName id="4C22D7886475FFADFD25EBC0D398F900" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[681,937,1744,1764]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="praebrevissimus" status="sp. nov.">
<emphasis id="B95670196475FFADFD25EBC0D398F900" box="[681,937,1744,1764]" italics="true" pageId="28" pageNumber="29">Novispathodus praebrevissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFC22EBC0D3EDF900" box="[942,988,1744,1764]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196475FFADFC6DEBC0D3DEF900" bold="true" box="[993,1007,1744,1764]" pageId="28" pageNumber="29">A</emphasis>
LIL506,PIMUZ 39274;
<emphasis id="B95670196475FFADFB20EBC0D489F900" bold="true" box="[1196,1208,1744,1764]" pageId="28" pageNumber="29">B</emphasis>
LAR207, PIMUZ 39275;
<emphasis id="B95670196475FFADFF6AEBFAD0C4F91A" bold="true" box="[230,245,1770,1790]" pageId="28" pageNumber="29">D</emphasis>
LIL508, PIMUZ 39276;
<emphasis id="B95670196475FFADFE3EEBFAD18CF91A" bold="true" box="[434,445,1770,1790]" pageId="28" pageNumber="29">F</emphasis>
LIL507, PIMUZ 39277;
<emphasis id="B95670196475FFADFDF6EBFAD2B4F91A" bold="true" box="[634,645,1770,1790]" pageId="28" pageNumber="29">L</emphasis>
LIL507, PIMUZ 39278;
<emphasis id="B95670196475FFADFCCFEBFAD364F91A" bold="true" box="[835,853,1770,1790]" pageId="28" pageNumber="29">M</emphasis>
LIL509, PIMUZ 39279;
<emphasis id="B95670196475FFADFB9EEBFAD411F91A" bold="true" box="[1042,1056,1770,1790]" pageId="28" pageNumber="29">N</emphasis>
LIL507,PIMUZ 39280;
<emphasis id="B95670196475FFADFB52EBFAD4DCF91A" bold="true" box="[1246,1261,1770,1790]" pageId="28" pageNumber="29">Q</emphasis>
LIL507, PIMUZ 39281;
<emphasis id="B95670196475FFADFEAFEA15D11FF8FD" bold="true" box="[291,302,1797,1817]" pageId="28" pageNumber="29">S</emphasis>
LIL507, PIMUZ 39282;
<emphasis id="B95670196475FFADFE60EA15D1C6F8FD" bold="true" box="[492,503,1797,1817]" pageId="28" pageNumber="29">T</emphasis>
LIL507, PIMUZ 39283.
<emphasis id="B95670196475FFADFD39EA15D2F3F8FD" bold="true" box="[693,706,1797,1817]" pageId="28" pageNumber="29">C</emphasis>
,
<emphasis id="B95670196475FFADFD45EA15D2E9F8FD" bold="true" box="[713,728,1797,1817]" pageId="28" pageNumber="29">O</emphasis>
,
<emphasis id="B95670196475FFADFD6DEA15D2DCF8FD" bold="true" box="[737,749,1797,1817]" pageId="28" pageNumber="29">R</emphasis>
<taxonomicName id="4C22D7886475FFADFD7EEA15D3CAF8FD" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[754,1019,1797,1817]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="praebrevissimus" status="sp. nov.">
<emphasis id="B95670196475FFADFD7EEA15D35CF8FD" box="[754,877,1797,1817]" italics="true" pageId="28" pageNumber="29">Novispathodus</emphasis>
?
<emphasis id="B95670196475FFADFCF5EA15D3CAF8FD" box="[889,1019,1797,1817]" italics="true" pageId="28" pageNumber="29">praebrevissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFC73EA15D41CF8FD" box="[1023,1069,1797,1817]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196475FFADFBBFEA15D471F8FD" bold="true" box="[1075,1088,1797,1817]" pageId="28" pageNumber="29">C</emphasis>
LAR204,PIMUZ 39284;
<emphasis id="B95670196475FFADFA8AEA15D524F8FD" bold="true" box="[1286,1301,1797,1817]" pageId="28" pageNumber="29">O</emphasis>
LIL507, PIMUZ 39285;
<emphasis id="B95670196475FFADFEAFEA30D11EF8D0" bold="true" box="[291,303,1824,1844]" pageId="28" pageNumber="29">R</emphasis>
LIL507, PIMUZ 39286.
<emphasis id="B95670196475FFADFE61EA30D1C9F8D0" bold="true" box="[493,504,1824,1844]" pageId="28" pageNumber="29">E</emphasis>
,
<emphasis id="B95670196475FFADFD8CEA30D237F8D0" bold="true" box="[512,518,1824,1844]" pageId="28" pageNumber="29">I</emphasis>
<taxonomicName id="4C22D7886475FFADFD86EA30D2B4F8D0" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[522,645,1824,1844]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="B95670196475FFADFD86EA30D2B4F8D0" box="[522,645,1824,1844]" italics="true" pageId="28" pageNumber="29">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886475FFADFD37EA30D3DAF8D0" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[699,1003,1824,1844]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196475FFADFD37EA30D365F8D0" box="[699,852,1824,1844]" italics="true" pageId="28" pageNumber="29">pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
;
<emphasis id="B95670196475FFADFC7EEA30D3CCF8D0" bold="true" box="[1010,1021,1824,1844]" pageId="28" pageNumber="29">E</emphasis>
LIL507,PIMUZ 39232,
<emphasis id="B95670196475FFADFB37EA30D4F0F8D0" bold="true" box="[1211,1217,1824,1844]" pageId="28" pageNumber="29">I</emphasis>
LIL507, PIMUZ 39233.
<emphasis id="B95670196475FFADFF6AEA2AD0C5F8AA" bold="true" box="[230,244,1850,1870]" pageId="28" pageNumber="29">G</emphasis>
<taxonomicName id="4C22D7886475FFADFF74EA2AD255F8AA" authority="(Orchard)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[248,612,1850,1870]" genus="Triassospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196475FFADFF74EA2AD1B4F8AA" box="[248,389,1850,1870]" italics="true" pageId="28" pageNumber="29">Triassospathodus</emphasis>
aff.
<emphasis id="B95670196475FFADFE2BEA2AD23FF8AA" box="[423,526,1850,1870]" italics="true" pageId="28" pageNumber="29">symmetricus</emphasis>
(Orchard)
</taxonomicName>
;LIL507, PIMUZ 39309.
<emphasis id="B95670196475FFADFCA8EA2AD303F8AA" bold="true" box="[804,818,1850,1870]" pageId="28" pageNumber="29">H</emphasis>
<taxonomicName id="4C22D7886475FFADFCBBEA2AD383F8AA" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[823,946,1850,1870]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="B95670196475FFADFCBBEA2AD383F8AA" box="[823,946,1850,1870]" italics="true" pageId="28" pageNumber="29">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886475FFADFC6BEA2AD4B6F8AA" authority="(Orchard)" baseAuthorityName="Orchard" box="[999,1159,1850,1870]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196475FFADFC6BEA2AD400F8AA" box="[999,1073,1850,1870]" italics="true" pageId="28" pageNumber="29">abruptus</emphasis>
(Orchard)
</taxonomicName>
; LIL507, PIMUZ 39212.
<emphasis id="B95670196475FFADFACBEA2AD57EF8AA" bold="true" box="[1351,1359,1850,1870]" pageId="28" pageNumber="29">J</emphasis>
,
<emphasis id="B95670196475FFADFADBEA2AD552F8AA" bold="true" box="[1367,1379,1850,1870]" pageId="28" pageNumber="29">K</emphasis>
<taxonomicName id="4C22D7886475FFADFF25EA45D19BF88D" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[169,426,1877,1897]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="praebrevissimus">
<emphasis id="B95670196475FFADFF25EA45D19BF88D" box="[169,426,1877,1897]" italics="true" pageId="28" pageNumber="29">Novispathodus praebrevissimus</emphasis>
</taxonomicName>
(juvenile) n. sp.;
<emphasis id="B95670196475FFADFDB9EA45D20CF88D" bold="true" box="[565,573,1877,1897]" pageId="28" pageNumber="29">J</emphasis>
LIL507, PIMUZ 39287;
<emphasis id="B95670196475FFADFD77EA45D336F88D" bold="true" box="[763,775,1877,1897]" pageId="28" pageNumber="29">K</emphasis>
LIL507,PIMUZ 39288.
<emphasis id="B95670196475FFADFC49EA45D3E0F88D" bold="true" box="[965,977,1877,1897]" pageId="28" pageNumber="29">P</emphasis>
<taxonomicName id="4C22D7886475FFADFC5AEA45D566F88D" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" box="[982,1367,1877,1897]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="robustispinus">
<emphasis id="B95670196475FFADFC5AEA45D4F1F88D" box="[982,1216,1877,1897]" italics="true" pageId="28" pageNumber="29">Novispathodus robustispinus</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; LIL507,PIMUZ 39290
</paragraph>
</caption>
<caption id="DF5DFC836474FFACFF25EBE0D1C8F8E0" ID-DOI="http://doi.org/10.5281/zenodo.12000320" ID-Zenodo-Dep="12000320" box="[169,505,1776,1796]" httpUri="https://zenodo.org/record/12000320/files/figure.png" pageId="29" pageNumber="30" startId="29.[169,200,1776,1796]" targetBox="[152,1438,238,1801]" targetPageId="29" targetType="figure">
<paragraph id="8B9DAC0B6474FFACFF25EBE0D1C8F8E0" blockId="29.[169,505,1776,1796]" box="[169,505,1776,1796]" pageId="29" pageNumber="30">
<emphasis id="B95670196474FFACFF25EBE0D0D6F8E0" bold="true" box="[169,231,1776,1796]" pageId="29" pageNumber="30">Fig. 18</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<caption id="DF5DFC836477FFAFFF25EBB4D23AF896" ID-DOI="http://doi.org/10.5281/zenodo.12000347" ID-Zenodo-Dep="12000347" httpUri="https://zenodo.org/record/12000347/files/figure.png" pageId="30" pageNumber="31" startId="30.[169,200,1700,1720]" targetBox="[153,1439,238,1918]" targetPageId="30" targetType="figure">
<paragraph id="8B9DAC0B6477FFAFFF25EBB4D23AF896" blockId="30.[169,1388,1699,1906]" pageId="30" pageNumber="31">
<emphasis id="B95670196477FFAFFF25EBB4D0D9F95C" bold="true" box="[169,232,1700,1720]" pageId="30" pageNumber="31">Fig. 19</emphasis>
<taxonomicName id="4C22D7886477FFAFFF7EEBB4D1B2F95C" authorityName="Orchard" authorityYear="2005" box="[242,387,1700,1720]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E6477FFAFFADCEBBDD566F95F" attach="left" box="[1360,1367,1709,1723]" fontSize="6" pageId="30" pageNumber="31">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B95670196477FFAFFDCDEBAED27EF936" bold="true" box="[577,591,1726,1746]" pageId="30" pageNumber="31">A</emphasis>
<taxonomicName id="4C22D7886477FFAFFDDFEBAED3ACF936" authority="(Orchard)" baseAuthorityName="Orchard" box="[595,925,1726,1746]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196477FFAFFDDFEBAED2FFF936" box="[595,718,1726,1746]" italics="true" pageId="30" pageNumber="31">Novispathodus</emphasis>
cf.
<emphasis id="B95670196477FFAFFD65EBAED377F936" box="[745,838,1726,1746]" italics="true" pageId="30" pageNumber="31">brevissimus</emphasis>
(Orchard)
</taxonomicName>
; LIL515C,PIMUZ 39199.
<emphasis id="B95670196477FFAFFBE5EBAED444F936" bold="true" box="[1129,1141,1726,1746]" pageId="30" pageNumber="31">B</emphasis>
,
<emphasis id="B95670196477FFAFFBF2EBAED4B8F936" bold="true" box="[1150,1161,1726,1746]" pageId="30" pageNumber="31">F</emphasis>
<taxonomicName id="4C22D7886477FFAFFB01EBAED0CAF909" authority="(Orchard)" baseAuthorityName="Orchard" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196477FFAFFB01EBAED558F936" box="[1165,1385,1726,1746]" italics="true" pageId="30" pageNumber="31">Novispathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196477FFAFFE8DEBC9D13CF909" bold="true" box="[257,269,1753,1773]" pageId="30" pageNumber="31">B</emphasis>
LIL515B, PIMUZ 39200;
<emphasis id="B95670196477FFAFFE5AEBC9D1D0F909" bold="true" box="[470,481,1753,1773]" pageId="30" pageNumber="31">F</emphasis>
BAN1, PIMUZ 39201.
<emphasis id="B95670196477FFAFFD1BEBC9D295F909" bold="true" box="[663,676,1753,1773]" pageId="30" pageNumber="31">C</emphasis>
<taxonomicName id="4C22D7886477FFAFFD24EBC9D3F0F909" authority="(Bender)" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[680,961,1753,1773]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196477FFAFFD24EBC9D342F909" box="[680,883,1753,1773]" italics="true" pageId="30" pageNumber="31">Triassospathodus homeri</emphasis>
(Bender)
</taxonomicName>
; LIL515D,PIMUZ 39317.
<emphasis id="B95670196477FFAFFB02EBC9D4ACF909" bold="true" box="[1166,1181,1753,1773]" pageId="30" pageNumber="31">D</emphasis>
,
<emphasis id="B95670196477FFAFFB29EBC9D482F909" bold="true" box="[1189,1203,1753,1773]" pageId="30" pageNumber="31">H</emphasis>
<taxonomicName id="4C22D7886477FFAFFB34EBC9D154F8EC" authority="(Orchard)" baseAuthorityName="Orchard" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196477FFAFFB34EBC9D502F909" box="[1208,1331,1753,1773]" italics="true" pageId="30" pageNumber="31">Novispathodus</emphasis>
?
<emphasis id="B95670196477FFAFFF3DEBE4D13FF8EC" box="[177,270,1780,1800]" italics="true" pageId="30" pageNumber="31">brevissimus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196477FFAFFEE0EBE4D14AF8EC" bold="true" box="[364,379,1780,1800]" pageId="30" pageNumber="31">D</emphasis>
LIL515D,PIMUZ 39202;
<emphasis id="B95670196477FFAFFDC9EBE4D262F8EC" bold="true" box="[581,595,1780,1800]" pageId="30" pageNumber="31">H</emphasis>
LIL514A, PIMUZ 39203.
<emphasis id="B95670196477FFAFFC92EBE4D318F8EC" bold="true" box="[798,809,1780,1800]" pageId="30" pageNumber="31">E</emphasis>
<taxonomicName id="4C22D7886477FFAFFCA1EBE4D46FF8EC" authority="(Bender)" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[813,1118,1780,1800]" genus="Triassospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196477FFAFFCA1EBE4D38BF8EC" box="[813,954,1780,1800]" italics="true" pageId="30" pageNumber="31">Triassospathodus</emphasis>
cf.
<emphasis id="B95670196477FFAFFC59EBE4D43EF8EC" box="[981,1039,1780,1800]" italics="true" pageId="30" pageNumber="31">homeri</emphasis>
(Bender)
</taxonomicName>
; LIL515C, PIMUZ 39318.
<emphasis id="B95670196477FFAFFAA6EBE4D509F8EC" bold="true" box="[1322,1336,1780,1800]" pageId="30" pageNumber="31">G</emphasis>
,
<emphasis id="B95670196477FFAFFACDEBE4D57CF8EC" bold="true" box="[1345,1357,1780,1800]" pageId="30" pageNumber="31">K</emphasis>
,
<emphasis id="B95670196477FFAFFF25EA1ED0E3F8C6" bold="true" box="[169,210,1806,1826]" pageId="30" pageNumber="31">OQ</emphasis>
<taxonomicName id="4C22D7886477FFAFFF5BEA1ED163F8C6" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[215,338,1806,1826]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196477FFAFFF5BEA1ED163F8C6" box="[215,338,1806,1826]" italics="true" pageId="30" pageNumber="31">Novispathodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626477FFAFFEDAEA1ED1B3F8C6" box="[342,386,1806,1826]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
A;
<emphasis id="B95670196477FFAFFE16EA1ED199F8C6" bold="true" box="[410,424,1806,1826]" pageId="30" pageNumber="31">G</emphasis>
LIL504,PIMUZ 39251;
<emphasis id="B95670196477FFAFFDEAEA1ED243F8C6" bold="true" box="[614,626,1806,1826]" pageId="30" pageNumber="31">K</emphasis>
LIL504,PIMUZ 39252;
<emphasis id="B95670196477FFAFFCBCEA1ED30EF8C6" bold="true" box="[816,831,1806,1826]" pageId="30" pageNumber="31">O</emphasis>
LIL505, PIMUZ 39253;
<emphasis id="B95670196477FFAFFC71EA1ED438F8C6" bold="true" box="[1021,1033,1806,1826]" pageId="30" pageNumber="31">P</emphasis>
LIL505, PIMUZ 39254;
<emphasis id="B95670196477FFAFFB4AEA1ED4E4F8C6" bold="true" box="[1222,1237,1806,1826]" pageId="30" pageNumber="31">Q</emphasis>
LIL505,PIMUZ 39255.
<emphasis id="B95670196477FFAFFF6AEA39D0DDF8D9" bold="true" box="[230,236,1833,1853]" pageId="30" pageNumber="31">I</emphasis>
,
<emphasis id="B95670196477FFAFFF78EA39D0CEF8D9" bold="true" box="[244,255,1833,1853]" pageId="30" pageNumber="31">L</emphasis>
,
<emphasis id="B95670196477FFAFFE8BEA39D128F8D9" bold="true" box="[263,281,1833,1853]" pageId="30" pageNumber="31">M</emphasis>
,
<taxonomicName id="4C22D7886477FFAFFEADEA39D25FF8D9" authority="(Orchard)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[289,622,1833,1853]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196477FFAFFEADEA39D229F8D9" box="[289,536,1833,1853]" italics="true" pageId="30" pageNumber="31">Triassospathodus symmetricus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196477FFAFFDF9EA39D24AF8D9" bold="true" box="[629,635,1833,1853]" pageId="30" pageNumber="31">I</emphasis>
QIA141, PIMUZ 39336;
<emphasis id="B95670196477FFAFFCCCEA39D37AF8D9" bold="true" box="[832,843,1833,1853]" pageId="30" pageNumber="31">L</emphasis>
LIL512, PIMUZ 39337;
<emphasis id="B95670196477FFAFFB84EA39D42BF8D9" bold="true" box="[1032,1050,1833,1853]" pageId="30" pageNumber="31">M</emphasis>
LIL514A, PIMUZ 39338.
<emphasis id="B95670196477FFAFFB68EA39D4DDF8D9" bold="true" box="[1252,1260,1833,1853]" pageId="30" pageNumber="31">J</emphasis>
<taxonomicName id="4C22D7886477FFAFFB7DEA39D55DF8D9" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1265,1388,1833,1853]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B95670196477FFAFFB7DEA39D55DF8D9" box="[1265,1388,1833,1853]" italics="true" pageId="30" pageNumber="31">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886477FFAFFF56EA54D23BF8BC" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[218,522,1860,1880]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196477FFAFFF56EA54D142F8BC" box="[218,371,1860,1880]" italics="true" pageId="30" pageNumber="31">pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; QIA138, PIMUZ 39237.
<emphasis id="B95670196477FFAFFD5FEA54D2D0F8BC" bold="true" box="[723,737,1860,1880]" pageId="30" pageNumber="31">N</emphasis>
<taxonomicName id="4C22D7886477FFAFFD6AEA54D3C4F8BC" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[742,1013,1860,1880]" genus="Triassospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196477FFAFFD6AEA54D342F8BC" box="[742,883,1860,1880]" italics="true" pageId="30" pageNumber="31">Triassospathodus</emphasis>
cf.
<emphasis id="B95670196477FFAFFC02EA54D3C4F8BC" box="[910,1013,1860,1880]" italics="true" pageId="30" pageNumber="31">symmetricus</emphasis>
</taxonomicName>
; LIL514B, PIMUZ 39339.
<emphasis id="B95670196477FFAFFB4CEA54D4FDF8BC" bold="true" box="[1216,1228,1860,1880]" pageId="30" pageNumber="31">R</emphasis>
<taxonomicName id="4C22D7886477FFAFFB5DEA54D11AF896" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="praebrevissimus">
<emphasis id="B95670196477FFAFFB5DEA54D11AF896" italics="true" pageId="30" pageNumber="31">Novispathodus praebrevissimus</emphasis>
</taxonomicName>
n.sp; LIL505, PIMUZ 39289
</paragraph>
</caption>
<paragraph id="8B9DAC0B6476FFAEFF1BECBCD335FAE7" blockId="31.[151,775,235,1507]" pageId="31" pageNumber="32">
Te P
<subScript id="17A6AE4E6476FFAEFF55ECA7D0D3FE2C" attach="left" box="[217,226,439,456]" fontSize="7" pageId="31" pageNumber="32">1</subScript>
element of this species is less robust, more rectangular with more fused denticles than that of
<taxonomicName id="4C22D7886476FFAEFD3CECDCD0FEFDE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="crassatus">
<emphasis id="B95670196476FFAEFD3CECDCD2F2FE07" box="[688,707,460,483]" italics="true" pageId="31" pageNumber="32">Ic</emphasis>
.
<emphasis id="B95670196476FFAEFD5EECDCD0FEFDE7" italics="true" pageId="31" pageNumber="32">crassatus</emphasis>
</taxonomicName>
. Its morphology also recalls that of the homologous element in
<taxonomicName id="4C22D7886476FFAEFE9BEF1CD238FDC7" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[279,521,523,547]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFE9BEF1CD107FDC7" box="[279,310,524,547]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
.
<emphasis id="B95670196476FFAEFECAEF1BD238FDC7" box="[326,521,523,547]" italics="true" pageId="31" pageNumber="32">pingdingshanensis</emphasis>
</taxonomicName>
, but its basal cavity is relatively smaller and its denticles are not as posteriorly recurved nor usually as broad as in the latter. Te P
<subScript id="17A6AE4E6476FFAEFD42EF47D2E6FD8C" attach="left" box="[718,727,599,616]" fontSize="7" pageId="31" pageNumber="32">1</subScript>
element of
<taxonomicName id="4C22D7886476FFAEFF70EF7CD19CFD67" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[252,429,620,643]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196476FFAEFF70EF7CD19CFD67" box="[252,429,620,643]" italics="true" pageId="31" pageNumber="32">Tr. symmetricus</emphasis>
</taxonomicName>
has more posteriorly reclined denticles, it may have a small terminal denticle but not several of increasingly smaller size. Yet, the distinction between
<taxonomicName id="4C22D7886476FFAEFF70EFDCD19BFD07" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[252,426,716,739]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196476FFAEFF70EFDCD19BFD07" box="[252,426,716,739]" italics="true" pageId="31" pageNumber="32">Tr. symmetricus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFE69EFDCD242FD07" box="[485,627,715,739]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFE69EFDCD242FD07" box="[485,627,715,739]" italics="true" pageId="31" pageNumber="32">Nv. abruptus</emphasis>
</taxonomicName>
may be confusing. Te P
<subScript id="17A6AE4E6476FFAEFEA8EFE7D11CFCEC" attach="left" box="[292,301,759,776]" fontSize="7" pageId="31" pageNumber="32">1</subScript>
element of
<taxonomicName id="4C22D7886476FFAEFE27EFFCD22CFCE7" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[427,541,747,771]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196476FFAEFE27EFFCD1F5FCE7" box="[427,452,748,771]" italics="true" pageId="31" pageNumber="32">Tr</emphasis>
.
<emphasis id="B95670196476FFAEFE5CEFFBD22CFCE7" box="[464,541,747,771]" italics="true" pageId="31" pageNumber="32">homeri</emphasis>
</taxonomicName>
has a more elongated basal cavity and a more developed, posteriorly reclined and laterally deflected process than in both
<taxonomicName id="4C22D7886476FFAEFD07EE3CD0E5FC87" baseAuthorityName="Orchard" baseAuthorityYear="1995" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196476FFAEFD07EE3CD0E5FC87" italics="true" pageId="31" pageNumber="32">Tr. symmetricus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFE99EE5CD197FC87" box="[277,422,843,867]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFE99EE5CD197FC87" box="[277,422,843,867]" italics="true" pageId="31" pageNumber="32">Nv. abruptus</emphasis>
</taxonomicName>
. Goudemand and Orchard (in
<bibRefCitation id="EFB3D1FA6476FFAEFF1BEE7BD182FC67" author="Goudemand" box="[151,435,875,899]" etAl="et al." firstAuthor="Goudemand" pageId="31" pageNumber="32" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
) implicitly suggested that elements like those of their
<emphasis id="B95670196476FFAEFE3CEE9CD1FEFC47" box="[432,463,908,931]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
. sp. nov. B, where the small terminal denticles are not increasingly smaller but of equal (small) height instead, may be transitional between
<taxonomicName id="4C22D7886476FFAEFF1BEEFCD112FBE7" box="[151,291,1003,1027]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFF1BEEFCD112FBE7" box="[151,291,1003,1027]" italics="true" pageId="31" pageNumber="32">Nv. abruptus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFED6EEFCD1FBFBE7" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[346,458,1003,1027]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196476FFAEFED6EEFCD1FBFBE7" box="[346,458,1003,1027]" italics="true" pageId="31" pageNumber="32">Tr. homeri</emphasis>
</taxonomicName>
(compare the elements illustrated in figs. 2.9 and 3.17 of
<bibRefCitation id="EFB3D1FA6476FFAEFE6FE91BD24DFBC7" author="Orchard" box="[483,636,1035,1059]" firstAuthor="Orchard" pageId="31" pageNumber="32" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard 1995</bibRefCitation>
, assigned to
<taxonomicName id="4C22D7886476FFAEFF1BE93CD13BFBA7" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[151,266,1067,1091]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196476FFAEFF1BE93CD13BFBA7" box="[151,266,1067,1091]" italics="true" pageId="31" pageNumber="32">Tr. homeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFECFE93CD1E4FBA7" box="[323,469,1067,1091]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFECFE93CD1E4FBA7" box="[323,469,1067,1091]" italics="true" pageId="31" pageNumber="32">Nv. Abruptus</emphasis>
</taxonomicName>
, respectively). Note further that, based on the material from Tsoteng, it is likely that suchP
<subScript id="17A6AE4E6476FFAEFF53E967D0D9FB6C" attach="left" box="[223,232,1143,1160]" fontSize="7" pageId="31" pageNumber="32">1</subScript>
elements were still associated with a
<taxonomicName id="4C22D7886476FFAEFD0FE97BD08CFB47" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B95670196476FFAEFD0FE97BD08CFB47" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
apparatus. Tis suggests that P
<subScript id="17A6AE4E6476FFAEFD9BE987D211FB4C" attach="left" box="[535,544,1175,1192]" fontSize="7" pageId="31" pageNumber="32">1</subScript>
elements like that of
<taxonomicName id="4C22D7886476FFAEFF1BE9BCD138FB27" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[151,265,1195,1219]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B95670196476FFAEFF1BE9BCD138FB27" box="[151,265,1195,1219]" italics="true" pageId="31" pageNumber="32">Tr. homeri</emphasis>
</taxonomicName>
may have evolved before the more substantial modifications of the rest of the apparatus implied by the difference between
<taxonomicName id="4C22D7886476FFAEFEE7E9FBD23DFAE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[363,524,1259,1283]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B95670196476FFAEFEE7E9FBD23DFAE7" box="[363,524,1259,1283]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFDCDE9FBD2CFFAE7" box="[577,766,1259,1283]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B95670196476FFAEFDCDE9FBD2CFFAE7" box="[577,766,1259,1283]" italics="true" pageId="31" pageNumber="32">Triassospathodus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFF1BE81CD0D0FA07" blockId="31.[151,775,235,1507]" pageId="31" pageNumber="32">
<emphasis id="B95670196476FFAEFF1BE81CD123FAC7" box="[151,274,1292,1315]" italics="true" pageId="31" pageNumber="32">Occurrence</emphasis>
. Worldwide occurrence in latest Smithian and early Spathian rocks.
<taxonomicName id="4C22D7886476FFAEFE62E83BD25EFAA7" ID-CoL="88NX" authorityName="Spath" authorityYear="1930" box="[494,623,1323,1347]" class="Cephalopoda" family="Xenoceltitidae" genus="Xenoceltites" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B95670196476FFAEFE62E83BD25EFAA7" box="[494,623,1323,1347]" italics="true" pageId="31" pageNumber="32">Xenoceltites</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFD22E83BD336FAA7" ID-CoL="7X7S" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[686,775,1323,1347]" class="Cephalopoda" family="Columbitidae" genus="Tirolites" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B95670196476FFAEFD22E83BD336FAA7" box="[686,775,1323,1347]" italics="true" pageId="31" pageNumber="32">Tirolites</emphasis>
</taxonomicName>
beds within
<taxonomicName id="4C22D7886476FFAEFEAAE85CD22BFA87" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[294,538,1355,1379]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFEAAE85CD174FA87" box="[294,325,1356,1379]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
.
<emphasis id="B95670196476FFAEFEDAE85BD22BFA87" box="[342,538,1355,1379]" italics="true" pageId="31" pageNumber="32">pingdingshanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886476FFAEFDD5E85CD336FA87" box="[601,775,1355,1379]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196476FFAEFDD5E85CD249FA87" box="[601,632,1356,1379]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
.
<emphasis id="B95670196476FFAEFD05E85BD336FA87" box="[649,775,1355,1379]" italics="true" pageId="31" pageNumber="32">brevissimus</emphasis>
</taxonomicName>
zones,
<collectingCountry id="F335EC9B6476FFAEFF68E87CD111FA67" box="[228,288,1388,1411]" name="Japan" pageId="31" pageNumber="32">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6476FFAEFEBCE87BD182FA67" author="Koike" box="[304,435,1387,1411]" firstAuthor="Koike" pageId="31" pageNumber="32" pagination="53 - 69" refId="ref42668" refString="Koike, T. (1981). Biostratigraphy of Triassic conodonts in Japan. Science Reports of the Yokohama National University, Section 2, (38), 53 - 69." type="journal article" year="1981">Koike, 1981</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA6476FFAEFE4EE87BD29BFA67" author="Maekawa" box="[450,682,1387,1411]" etAl="et al." firstAuthor="Maekawa" pageId="31" pageNumber="32" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
).
<collectingCountry id="F335EC9B6476FFAEFD4EE87BD336FA67" box="[706,775,1387,1411]" name="Oman" pageId="31" pageNumber="32">Oman</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6476FFAEFF13E89BD172FA47" author="Orchard" box="[159,323,1419,1443]" firstAuthor="Orchard" pageId="31" pageNumber="32" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
). British Columbia (
<bibRefCitation id="EFB3D1FA6476FFAEFDBAE89BD120FA27" author="Orchard &amp; Zonneveld" firstAuthor="Orchard" pageId="31" pageNumber="32" refId="ref45906" refString="Orchard, M. J., &amp; Zonneveld, J. P. (2009). The Lower Triassic Sulphur Moun-" type="book" year="2009">Orchard &amp; Zonneveld, 2009</bibRefCitation>
), South
<collectingCountry id="F335EC9B6476FFAEFEE3E8BBD182FA27" box="[367,435,1451,1475]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
;
<bibRefCitation id="EFB3D1FA6476FFAEFE32E8BBD2E3FA27" author="Goudemand" box="[446,722,1451,1475]" etAl="et al." firstAuthor="Goudemand" pageId="31" pageNumber="32" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012" yearSuffix="b">Goudemand et al., 2012b</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,194 @@
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<treatment id="6900A32CD76D39419915FDBE1D907AE1" ID-DOI="http://doi.org/10.5281/zenodo.13128477" ID-Zenodo-Dep="13128477" LSID="urn:lsid:plazi:treatment:6900A32CD76D39419915FDBE1D907AE1" httpUri="http://treatment.plazi.org/id/6900A32CD76D39419915FDBE1D907AE1" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="A9B341B1D76D394E9915FDBE1A8F7961" box="[813,1180,587,611]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="E116123AD76D394E9915FDBE1A8F7961" blockId="7.[813,1180,587,611]" box="[813,1180,587,611]" pageId="7" pageNumber="8">
<heading id="BA5EA556D76D394E9915FDBE1A8F7961" box="[813,1180,587,611]" fontSize="9" level="2" pageId="7" pageNumber="8" reason="8">
<taxonomicName id="26A969B9D76D394E9915FDBE1A1A7960" authority="Mojsisovics, 1902" authorityName="Mojsisovics" box="[813,1033,587,611]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76D394E9915FDBE1A1A7960" bold="true" box="[813,1033,587,611]" italics="true" pageId="7" pageNumber="8">Syringonautilus sp.</emphasis>
</taxonomicName>
<figureCitation id="79920EBFD76D394E9E37FDBE1A8B7961" box="[1039,1176,587,611]" captionStart="Fig" captionStartId="7.[169,200,1876,1896]" captionTargetBox="[153,1438,1299,1934]" captionTargetId="figure-264@7.[229,1363,1315,1863]" captionTargetPageId="7" captionText="Fig. 5 Syringonautilus sp., PIMUZ 37903, P. 902/Mirigioli (Monte San Giorgio), Middle Besano Formation (bed 104), uppermost Anisian: a lateral view, left side, b oral view, c ventral view" figureDoi="http://doi.org/10.5281/zenodo.12000891" httpUri="https://zenodo.org/record/12000891/files/figure.png" pageId="7" pageNumber="8">Figure 5 ac</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76D394E9915FD7E1A487801" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="E116123AD76D394E9915FD7E1A487801" blockId="7.[813,1437,651,771]" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E9915FD7E1D8379A1" bold="true" box="[813,912,651,675]" pageId="7" pageNumber="8">Material</emphasis>
Specimen
<collectionCode id="87B88AFFD76D394E9E31FD791A4E79A1" box="[1033,1117,652,675]" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="7" pageNumber="8">PIMUZ</collectionCode>
37903, partially preserved as internal mold (about the last half-whorl, fractured and slightly deformed), partially as external mold from which a silicon cast was produced.
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76D39419915FCDE1C857AC1" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="E116123AD76D394E9915FCDE1DD37881" blockId="7.[813,1436,811,996]" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E9915FCDE1DCB7841" bold="true" box="[813,984,811,835]" pageId="7" pageNumber="8">Measurements</emphasis>
At maximum dm (deformed) of 44.5 mm other dimensions are: wh = 23.5 mm, uw =
<quantity id="2651BFDFD76D394E9F06FCBE1B8B7861" box="[1342,1432,843,867]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="10.0">10 mm</quantity>
, ww=
<quantity id="2651BFDFD76D394E9955FC9E1DAF7881" box="[877,956,875,899]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="25.0">25 mm</quantity>
.
</paragraph>
<paragraph id="E116123AD76D394E997AFC7E1AA678E1" blockId="7.[813,1436,811,996]" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E997AFC7E1A1378A1" bold="true" box="[834,1024,907,931]" pageId="7" pageNumber="8">Locality and age</emphasis>
Point 902/ Mirigioli (Meride,
<collectingRegion id="236DDCD8D76D394E9F73FC7E1D6478C1" country="Switzerland" name="Ticino" pageId="7" pageNumber="8">Canton Ticino</collectingRegion>
). Middle Besano Formation, Illyrian, Secedensis Zone (bed 104, light grey dolomite).
</paragraph>
<caption id="B5D642B2D76D394E9A91FC241F7D7F18" ID-DOI="http://doi.org/10.5281/zenodo.12000879" ID-Zenodo-Dep="12000879" httpUri="https://zenodo.org/record/12000879/files/figure.png" pageId="7" pageNumber="8" startId="7.[169,200,977,997]" targetBox="[153,777,238,1062]" targetPageId="7" targetType="figure">
<paragraph id="E116123AD76D394E9A91FC241F7D7F18" blockId="7.[169,755,977,1050]" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E9A91FC241EC878E7" bold="true" box="[169,219,977,997]" pageId="7" pageNumber="8">Fig. 4</emphasis>
<taxonomicName id="26A969B9D76D394E9ADEFC241FB678E7" box="[230,421,977,997]" class="Cephalopoda" family="Ceratitidae" genus="Stoppaniceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Ceratitida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="artinii">
<emphasis id="D3DDCE28D76D394E9ADEFC241F4A78E7" box="[230,345,977,997]" italics="true" pageId="7" pageNumber="8">Stoppaniceras</emphasis>
cf.
<emphasis id="D3DDCE28D76D394E9B4DFC241FB678E7" box="[373,421,977,997]" italics="true" pageId="7" pageNumber="8">artinii</emphasis>
</taxonomicName>
(
<bibRefCitation id="85386FCBD76D394E9B96FC241C3478E7" author="Airaghi, C." box="[430,551,977,997]" pageId="7" pageNumber="8" pagination="1048 - 1050" refId="ref6429" refString="Airaghi, C. (1912 a). Ammoniti degli scisti bituminosi di Besano in Lombardia. Bollettino Della Societa Geologica Italiana, 1911 (30), 1048 - 1050." type="journal article" year="1912">Airaghi, 1912a</bibRefCitation>
,
<bibRefCitation id="85386FCBD76D394E9817FC241C7678E7" author="Airaghi, C." box="[559,613,977,997]" pageId="7" pageNumber="8" pagination="1 - 30" refId="ref6462" refString="Airaghi, C. (1912 b). I molluschi degli Scisti Bituminosi di Besano in Lombardia. Atti Della Societa Italiana Di Scienze Naturali, 51 (1), 1 - 30." type="journal article" year="1912">1912b</bibRefCitation>
) associated with PIMUZ 29942. Image obtained producing positive silicon cast from negative external mold
</paragraph>
</caption>
<paragraph id="E116123AD76D39419915FBFE1C857AC1" blockId="7.[813,1437,1035,1219]" lastBlockId="8.[151,775,235,451]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E9915FBFE1DA57F21" bold="true" box="[813,950,1035,1059]" pageId="7" pageNumber="8">Description</emphasis>
Te specimen consists of about the last whorl, lacking its inner whorls. It shows a rapidly expanding smooth conch, with whorls deeply embracing. Te whorl section is more or less elliptical. Te anterior whorl section is higher than wide, well rounded and with the maximum width at the umbilical shoulders. Te umbilical shoulder passes to the inclined umbilical wall forming a continuous curve. Te flanks are convex, but slightly flattened, converging externally. In the last half-whorl, the ventrolateral shoulder becomes angularly rounded and corresponds to the maximum width of whorl. Te venter is convex and rounded, slightly flattened. Te other characters are not observable.
</paragraph>
</subSubSection>
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<paragraph id="E116123AD76D394E9A91F8A11FDD7C80" blockId="7.[169,1421,1876,1922]" pageId="7" pageNumber="8">
<emphasis id="D3DDCE28D76D394E9A91F8A11ECF7C6A" bold="true" box="[169,220,1876,1896]" pageId="7" pageNumber="8">Fig. 5</emphasis>
<taxonomicName id="26A969B9D76D394E9ADEF8A11F977C6A" box="[230,388,1876,1896]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="D3DDCE28D76D394E9ADEF8A11F767C6A" box="[230,357,1876,1896]" italics="true" pageId="7" pageNumber="8">Syringonautilus</emphasis>
sp.
</taxonomicName>
, PIMUZ 37903, P. 902/Mirigioli (Monte San Giorgio), Middle Besano Formation (bed 104), uppermost Anisian:
<emphasis id="D3DDCE28D76D394E9F20F8A11B307C6A" bold="true" box="[1304,1315,1876,1896]" pageId="7" pageNumber="8">a</emphasis>
lateral view, left side,
<emphasis id="D3DDCE28D76D394E9ACCF89B1F137C80" bold="true" box="[244,256,1902,1922]" pageId="7" pageNumber="8">b</emphasis>
oral view,
<emphasis id="D3DDCE28D76D394E9B61F89B1F707C80" bold="true" box="[345,355,1902,1922]" pageId="7" pageNumber="8">c</emphasis>
ventral view
</paragraph>
</caption>
<subSubSection id="A9B341B1D76239419AAFFE1E1D907AE1" pageId="8" pageNumber="9" type="discussion">
<paragraph id="E116123AD76239419AAFFE1E1D907AE1" blockId="8.[151,776,491,1027]" lastBlockId="8.[813,1436,235,483]" pageId="8" pageNumber="9">
<emphasis id="D3DDCE28D76239419AAFFE1E1F067901" bold="true" box="[151,277,491,515]" pageId="8" pageNumber="9">Discussion</emphasis>
Tis poorly preserved specimen shows similarities with species of
<taxonomicName id="26A969B9D76239419BACFDFE1C2D7921" authorityName="Mojsisovics" authorityYear="1902" box="[404,574,523,547]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419BACFDFE1C2D7921" box="[404,574,523,547]" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="85386FCBD762394198B9FDFE1ED87941" author="von Mojsisovics, E." pageId="8" pageNumber="9" pagination="1 - 322" refId="ref9428" refString="von Mojsisovics, E. (1882). Die Cephalopoden der Mediterranen Trias-Provinz. Abhandlungen Der Kaiserlich K o niglichen Geologischen Reichsanstalt, 10, 1 - 322." type="journal article" year="1882">Mojsisovics, 1882</bibRefCitation>
; Pieroni, 2020) from the Middle Triassic of the Alps. Following
<bibRefCitation id="85386FCBD76239419B33FDBE1FBD7961" author="Kummel, B." box="[267,430,587,611]" pageId="8" pageNumber="9" pagination="1 - 104" refId="ref7798" refString="Kummel, B. (1953). American Triassic Coiled Nautiloids, a study of the classification and evolution of late Paleozoic and early Mesozoic cephalopods, and description of new species. Geological Survey Professional Paper US Geological Survey, 250, 1 - 104." type="journal article" year="1953">Kummel (1953</bibRefCitation>
, p. 60),
<taxonomicName id="26A969B9D76239419833FDBE1CA67961" authorityName="Mojsisovics" authorityYear="1902" box="[523,693,587,611]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419833FDBE1CA67961" box="[523,693,587,611]" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="26A969B9D76239419B4BFD991FE97981" authorityName="Hyatt" authorityYear="1894" box="[371,506,620,643]" class="Cephalopoda" family="Syringonautilidae" genus="Syringoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419B4BFD991FE97981" box="[371,506,620,643]" italics="true" pageId="8" pageNumber="9">Syringoceras</emphasis>
</taxonomicName>
only by the position of its siphuncle: while in
<taxonomicName id="26A969B9D76239419BB7FD7E1C2A79A1" authorityName="Mojsisovics" authorityYear="1902" box="[399,569,651,675]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419BB7FD7E1C2A79A1" box="[399,569,651,675]" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
</taxonomicName>
it is subcentral, in
<taxonomicName id="26A969B9D76239419AAFFD591F0D79C1" authorityName="Hyatt" authorityYear="1894" box="[151,286,684,707]" class="Cephalopoda" family="Syringonautilidae" genus="Syringoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419AAFFD591F0D79C1" box="[151,286,684,707]" italics="true" pageId="8" pageNumber="9">Syringoceras</emphasis>
</taxonomicName>
it is nearly marginal. However, this character is still unknown in many species, and it is not visible in the present specimen. Te whorl section of
<taxonomicName id="26A969B9D762394198B8FD191D147801" authorityName="Hyatt" authorityYear="1894" box="[640,775,748,771]" class="Cephalopoda" family="Syringonautilidae" genus="Syringoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D762394198B8FD191D147801" box="[640,775,748,771]" italics="true" pageId="8" pageNumber="9">Syringoceras</emphasis>
</taxonomicName>
is usually slightly compressed (
<bibRefCitation id="85386FCBD7623941983DFCFE1CBE7821" author="Kummel, B." box="[517,685,779,803]" pageId="8" pageNumber="9" pagination="1 - 104" refId="ref7798" refString="Kummel, B. (1953). American Triassic Coiled Nautiloids, a study of the classification and evolution of late Paleozoic and early Mesozoic cephalopods, and description of new species. Geological Survey Professional Paper US Geological Survey, 250, 1 - 104." type="journal article" year="1953">Kummel, 1953</bibRefCitation>
, p. 61) and more expanded in
<taxonomicName id="26A969B9D76239419BA3FCDE1C567841" authorityName="Mojsisovics" authorityYear="1902" box="[411,581,811,835]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76239419BA3FCDE1C567841" box="[411,581,811,835]" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
</taxonomicName>
as in the present specimen. Among the species belonging to
<taxonomicName id="26A969B9D762394198BBFCB91ED57881" authorityName="Mojsisovics" authorityYear="1902" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D762394198BBFCB91ED57881" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
</taxonomicName>
documented in literature,
<taxonomicName id="26A969B9D76239419BC5FC9E1EA278A1" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bosnensis">
<emphasis id="D3DDCE28D76239419BC5FC9E1EA278A1" italics="true" pageId="8" pageNumber="9">Syringonautilus bosnensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="85386FCBD76239419AFAFC7E1F5A78A1" author="Hauer, F. R. von" box="[194,329,907,931]" pageId="8" pageNumber="9" pagination="251 - 296" refId="ref7370" refString="Hauer, F. R. von (1892). Beitrage zur Kenntnis der Cephalopoden aus der Trias von Bosnien. I. Neue Funde aus dem Muschelkalk von Han Bulog bei Sarajevo. Denkschriften der Kaiserlichen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, 59, 251 - 296" type="journal article" year="1892">Hauer, 1892</bibRefCitation>
, p. 6 [254], pl. 1,
<figureCitation id="79920EBFD7623941982EFC7E1C6F78A1" box="[534,636,907,931]" captionStart="Fig" captionStartId="6.[169,200,1760,1780]" captionTargetBox="[153,1438,238,1871]" captionTargetId="figure-16@6.[285,1305,254,1748]" captionTargetPageId="6" captionText="Fig. 3 Germanonautilus aff. ellipticus Parnes, 1986, PIMUZ 29942,Valle Stelle/Tre Fontane (Monte San Giorgio), Middle Besano Formation (bed 73), Secedensis Zone, Illyrian: a original specimen partially included in the matrix, lateral view, b ventral view,associated with an external mold of Stoppaniceras cf. artinii, c plaster cast replica of the same specimen, lateral view, d plaster cast replica of the same specimen,ventral view, e plaster cast replica of the same specimen, ventrolateral view, f drawing of the ventral view showing the hypothetical whorl section (in grey the flank areas)" figureDoi="http://doi.org/10.5281/zenodo.12000854" httpUri="https://zenodo.org/record/12000854/files/figure.png" pageId="8" pageNumber="9">Fig. 3ac</figureCitation>
; Trinodosus Zone of Han Bulog,
<collectingCountry id="99BE52AAD76239419BBDFC5E1FC378C1" box="[389,464,939,963]" name="Bosnia and Herzegovina" pageId="8" pageNumber="9">Bosnia</collectingCountry>
) shares the shape of whorl section and the umbilical width with the specimen here described. Te whorls of
<taxonomicName id="26A969B9D76239419BC7FC1E1A177A01" authority="(Mojsisovics, 1882)" baseAuthorityName="Mojsisovics" baseAuthorityYear="1882" box="[511,1028,235,1027]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="lilianus">
<emphasis id="D3DDCE28D76239419BC7FC1E1D147F01" box="[511,775,1003,1027]" italics="true" pageId="8" pageNumber="9">Syringonautilus lilianus</emphasis>
(
<bibRefCitation id="85386FCBD7623941990CFF1E1DE87A01" author="von Mojsisovics, E." box="[820,1019,235,259]" pageId="8" pageNumber="9" pagination="1 - 322" refId="ref9428" refString="von Mojsisovics, E. (1882). Die Cephalopoden der Mediterranen Trias-Provinz. Abhandlungen Der Kaiserlich K o niglichen Geologischen Reichsanstalt, 10, 1 - 322." type="journal article" year="1882">Mojsisovics, 1882</bibRefCitation>
)
</taxonomicName>
are less rapidly expanding and the conch has a wider umbilicus and circular to subquadrate whorl sections.
<taxonomicName id="26A969B9D76239419E32FEDE1DB17A61" authority="(Mojsisovics, 1882)" baseAuthorityName="Mojsisovics" baseAuthorityYear="1882" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="lilianus">
<emphasis id="D3DDCE28D76239419E32FEDE1AA77A41" box="[1034,1204,299,323]" italics="true" pageId="8" pageNumber="9">Syringonautilus</emphasis>
cf.
<emphasis id="D3DDCE28D76239419EE4FEDE1B3D7A41" box="[1244,1326,299,323]" italics="true" pageId="8" pageNumber="9">lilianus</emphasis>
(
<bibRefCitation id="85386FCBD76239419F03FEDE1D8B7A61" author="von Mojsisovics, E." pageId="8" pageNumber="9" pagination="1 - 322" refId="ref9428" refString="von Mojsisovics, E. (1882). Die Cephalopoden der Mediterranen Trias-Provinz. Abhandlungen Der Kaiserlich K o niglichen Geologischen Reichsanstalt, 10, 1 - 322." type="journal article" year="1882">Mojsisovics, 1882</bibRefCitation>
)
</taxonomicName>
from Rasa di Varese (Varese, Lombardy, Rasa Dolomite, San Salvatore Dolomite, Reitzi Zone sup.) was described by
<bibRefCitation id="85386FCBD76239419986FE7E1A437AA1" author="Pieroni, V." box="[958,1104,395,419]" pageId="8" pageNumber="9" refId="ref8236" refString="Pieroni, V. (2011). La Rasa di Varese e i suoi fossili. Pietro Macchione (ed.), (pp. 157)" type="journal volume" year="2011">Pieroni (2011</bibRefCitation>
, p. 57, pl. 2 cefalopodi,
<figureCitation id="79920EBFD76239419F6AFE7E1B9D7AA1" box="[1362,1422,395,419]" captionStart="Fig" captionStartId="6.[169,200,1760,1780]" captionTargetBox="[153,1438,238,1871]" captionTargetId="figure-16@6.[285,1305,254,1748]" captionTargetPageId="6" captionText="Fig. 3 Germanonautilus aff. ellipticus Parnes, 1986, PIMUZ 29942,Valle Stelle/Tre Fontane (Monte San Giorgio), Middle Besano Formation (bed 73), Secedensis Zone, Illyrian: a original specimen partially included in the matrix, lateral view, b ventral view,associated with an external mold of Stoppaniceras cf. artinii, c plaster cast replica of the same specimen, lateral view, d plaster cast replica of the same specimen,ventral view, e plaster cast replica of the same specimen, ventrolateral view, f drawing of the ventral view showing the hypothetical whorl section (in grey the flank areas)" figureDoi="http://doi.org/10.5281/zenodo.12000854" httpUri="https://zenodo.org/record/12000854/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
). It shows a more subquadrate whorl section and is more evolute.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,219 @@
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<treatment id="6900A32CD76F394E9915FEBE1B457921" ID-DOI="http://doi.org/10.5281/zenodo.13128474" ID-Zenodo-Dep="13128474" LSID="urn:lsid:plazi:treatment:6900A32CD76F394E9915FEBE1B457921" httpUri="http://treatment.plazi.org/id/6900A32CD76F394E9915FEBE1B457921" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="A9B341B1D76F394C9915FEBE1B5F7A61" box="[813,1356,331,355]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="E116123AD76F394C9915FEBE1B5F7A61" blockId="5.[813,1436,331,387]" box="[813,1356,331,355]" pageId="5" pageNumber="6">
<taxonomicName id="26A969B9D76F394C9915FEBE1B5F7A61" authority="Parnes, 1986" authorityName="Parnes" authorityYear="1986" box="[813,1356,331,355]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="D3DDCE28D76F394C9915FEBE1B5F7A61" bold="true" box="[813,1356,331,355]" italics="true" pageId="5" pageNumber="6">
Germanonautilus aff. ellipticus
<bibRefCitation id="85386FCBD76F394C9E89FEBE1B5F7A61" author="Parnes, A." box="[1201,1356,331,355]" pageId="5" pageNumber="6" pagination="1 - 59" refId="ref8115" refString="Parnes, A. (1986). Middle Triassic cephalopods from the Negev (Israel) and Sinai (Egypt). Geological Survey of Israel, Bulletin, 79, 1 - 59." type="journal article" year="1986">Parnes, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76F394C9F62FEBE1D447A81" pageId="5" pageNumber="6" type="description">
<paragraph id="E116123AD76F394C9F62FEBE1D447A81" blockId="5.[813,1436,331,387]" pageId="5" pageNumber="6">
<figureCitation id="79920EBFD76F394C9F62FEBE1B8F7A61" box="[1370,1436,331,355]" captionStart="Fig" captionStartId="6.[169,200,1760,1780]" captionTargetBox="[153,1438,238,1871]" captionTargetId="figure-16@6.[285,1305,254,1748]" captionTargetPageId="6" captionText="Fig. 3 Germanonautilus aff. ellipticus Parnes, 1986, PIMUZ 29942,Valle Stelle/Tre Fontane (Monte San Giorgio), Middle Besano Formation (bed 73), Secedensis Zone, Illyrian: a original specimen partially included in the matrix, lateral view, b ventral view,associated with an external mold of Stoppaniceras cf. artinii, c plaster cast replica of the same specimen, lateral view, d plaster cast replica of the same specimen,ventral view, e plaster cast replica of the same specimen, ventrolateral view, f drawing of the ventral view showing the hypothetical whorl section (in grey the flank areas)" figureDoi="http://doi.org/10.5281/zenodo.12000854" httpUri="https://zenodo.org/record/12000854/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
<heading id="BA5EA556D76F394C9915FE9E1D447A81" box="[813,855,363,387]" fontSize="10" level="2" pageId="5" pageNumber="6" reason="7">af.</heading>
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76F394C9915FE5E1B407941" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="E116123AD76F394C9915FE5E1B407941" blockId="5.[813,1436,427,579]" pageId="5" pageNumber="6">
<emphasis id="D3DDCE28D76F394C9915FE5E1D837AC1" bold="true" box="[813,912,427,451]" pageId="5" pageNumber="6">Material</emphasis>
Specimen
<collectionCode id="87B88AFFD76F394C9E2BFE591A747AC1" box="[1043,1127,428,451]" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="5" pageNumber="6">PIMUZ</collectionCode>
29942, a poorly preserved natural mold of the living chamber, filled by carbonate sediment; with slightly deformed coiling, partially included in the matrix. Te original shell is dissolved, with its morphology preserved on the ventral area.
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76F394C9915FD9E1B7F7E61" pageId="5" pageNumber="6" type="description">
<paragraph id="E116123AD76F394C9915FD9E1AFB79E1" blockId="5.[813,1437,619,739]" pageId="5" pageNumber="6">
<emphasis id="D3DDCE28D76F394C9915FD9E1DCB7981" bold="true" box="[813,984,619,643]" pageId="5" pageNumber="6">Measurements</emphasis>
At maximum dm (deformed) of
<quantity id="2651BFDFD76F394C9F75FD9E1B8F7981" box="[1357,1436,619,643]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="6" unit="mm" value="95.0">95 mm</quantity>
other dimensions are: wh =
<quantity id="2651BFDFD76F394C9EB0FD7E1AF579A1" box="[1160,1254,651,675]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="40.0">40 mm</quantity>
, uw =
<quantity id="2651BFDFD76F394C9F03FD7E1B8B79A1" box="[1339,1432,651,675]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="6" unit="mm" value="33.0">33 mm</quantity>
, ww=
<quantity id="2651BFDFD76F394C9955FD5E1DD179C1" box="[877,962,683,707]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="6" unit="mm" value="75.0">75 mm</quantity>
; at dm of
<quantity id="2651BFDFD76F394C9E7DFD5E1A8B79C1" box="[1093,1176,683,707]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="70.0">70 mm</quantity>
other dimensions are: wh =
<quantity id="2651BFDFD76F394C9950FD3E1DA479E1" box="[872,951,715,739]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="6" unit="mm" value="34.0">34 mm</quantity>
, uw =
<quantity id="2651BFDFD76F394C99C4FD3E1A5879E1" box="[1020,1099,715,739]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="6" unit="mm" value="24.0">24 mm</quantity>
, ww =
<quantity id="2651BFDFD76F394C9EADFD3E1AF679E1" box="[1173,1253,715,739]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="6" unit="mm" value="69.0">69 mm</quantity>
.
</paragraph>
<paragraph id="E116123AD76F394C9915FCFE1D6E78E1" blockId="5.[813,1437,779,995]" pageId="5" pageNumber="6">
<emphasis id="D3DDCE28D76F394C9915FCFE1DE17821" bold="true" box="[813,1010,779,803]" pageId="5" pageNumber="6">Locality and age</emphasis>
Valle Stelle (Meride,
<collectingRegion id="236DDCD8D76F394C9ED4FCFE1B817821" box="[1260,1426,779,803]" country="Switzerland" name="Ticino" pageId="5" pageNumber="6">Canton Ticino</collectingRegion>
). Middle Besano Formation, Illyrian, Secedensis Zone (recovered from a laminated bituminous dolostone in a gallery of the former mine; correlating to bed 73). In association with an imprint of an ammonoid (
<figureCitation id="79920EBFD76F394C9F27FC7E1B4978A1" box="[1311,1370,907,931]" captionStart="Fig" captionStartId="7.[169,200,977,997]" captionTargetBox="[153,777,238,1062]" captionTargetId="figure-235@7.[200,729,254,964]" captionTargetPageId="7" captionText="Fig. 4 Stoppaniceras cf. artinii (Airaghi, 1912a, 1912b) associated with PIMUZ 29942. Image obtained producing positive silicon cast from negative external mold" figureDoi="http://doi.org/10.5281/zenodo.12000879" httpUri="https://zenodo.org/record/12000879/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
) classified here as
<taxonomicName id="26A969B9D76F394C99E8FC591ACD78C1" box="[976,1246,939,963]" class="Cephalopoda" family="Ceratitidae" genus="Stoppaniceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Ceratitida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="artinii">
<emphasis id="D3DDCE28D76F394C99E8FC591A7B78C1" box="[976,1128,940,963]" italics="true" pageId="5" pageNumber="6">Stoppaniceras</emphasis>
cf.
<emphasis id="D3DDCE28D76F394C9EA1FC591ACD78C1" box="[1177,1246,940,963]" italics="true" pageId="5" pageNumber="6">artinii</emphasis>
</taxonomicName>
(
<bibRefCitation id="85386FCBD76F394C9EC9FC5E1B8678C1" author="Airaghi, C." box="[1265,1429,939,963]" pageId="5" pageNumber="6" pagination="1048 - 1050" refId="ref6429" refString="Airaghi, C. (1912 a). Ammoniti degli scisti bituminosi di Besano in Lombardia. Bollettino Della Societa Geologica Italiana, 1911 (30), 1048 - 1050." type="journal article" year="1912">Airaghi, 1912a</bibRefCitation>
,
<bibRefCitation id="85386FCBD76F394C9915FC3E1D7C78E1" author="Airaghi, C." box="[813,879,971,995]" pageId="5" pageNumber="6" pagination="1 - 30" refId="ref6462" refString="Airaghi, C. (1912 b). I molluschi degli Scisti Bituminosi di Besano in Lombardia. Atti Della Societa Italiana Di Scienze Naturali, 51 (1), 1 - 30." type="journal article" year="1912">1912b</bibRefCitation>
).
</paragraph>
<paragraph id="E116123AD76F394C9915FBFE1B7F7E61" blockId="5.[813,1437,1034,1379]" pageId="5" pageNumber="6">
<emphasis id="D3DDCE28D76F394C9915FBFE1DA57F21" bold="true" box="[813,950,1035,1059]" pageId="5" pageNumber="6">Description</emphasis>
Evolute shell (uw/dm = 0.34), consisting of about the last whorl. Te umbilicus is broad, without any preserved structure. Siphuncle and suture line are not visible. Te whorl section, wider than high, is subelliptical, with the maximum width at the umbilical shoulders. Te ventral area is convex, widely arched, with more or less rounded distinct ventrolateral shoulders. Te flanks are rounded and convex. Te umbilical shoulder passes to the inclined umbilical wall forming a continuous curve. Te growth lines are observable only on the ventral area and form a wide and deep hyponomic sinus.
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76F394E9915FA7E1D8E7AC1" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="discussion">
<paragraph id="E116123AD76F394E9915FA7E1D8E7AC1" blockId="5.[813,1437,1419,1827]" lastBlockId="7.[813,1437,235,548]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<emphasis id="D3DDCE28D76F394C9915FA7E1DB87EA1" bold="true" box="[813,939,1419,1443]" pageId="5" pageNumber="6">Discussion</emphasis>
Te genus
<taxonomicName id="26A969B9D76F394C9E14FA7E1AFE7EA1" authorityName="Mojsisovics" authorityYear="1902" box="[1068,1261,1419,1443]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76F394C9E14FA7E1AFE7EA1" box="[1068,1261,1419,1443]" italics="true" pageId="5" pageNumber="6">Germanonautilus</emphasis>
</taxonomicName>
is usually characterized by a narrow umbilicus and trapezoidal whorl sections, with more or less flattened or depressed venter. Te specimen described here shows similarities in particular with
<taxonomicName id="26A969B9D76F394C99E0F9FE1B857D21" authority="Parnes, 1986" authorityName="Parnes" authorityYear="1986" box="[984,1430,1547,1571]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="D3DDCE28D76F394C99E0F9FE1B117D21" box="[984,1282,1547,1571]" italics="true" pageId="5" pageNumber="6">Germanonautilus ellipticus</emphasis>
<bibRefCitation id="85386FCBD76F394C9F32F9F91B857D21" author="Parnes, A." box="[1290,1430,1547,1571]" pageId="5" pageNumber="6" pagination="1 - 59" refId="ref8115" refString="Parnes, A. (1986). Middle Triassic cephalopods from the Negev (Israel) and Sinai (Egypt). Geological Survey of Israel, Bulletin, 79, 1 - 59." type="journal article" year="1986">Parnes, 1986</bibRefCitation>
</taxonomicName>
. Tis species was described by
<bibRefCitation id="85386FCBD76F394C9EA9F9D91B3E7D41" author="Parnes, A." box="[1169,1325,1579,1603]" pageId="5" pageNumber="6" pagination="1 - 59" refId="ref8115" refString="Parnes, A. (1986). Middle Triassic cephalopods from the Negev (Israel) and Sinai (Egypt). Geological Survey of Israel, Bulletin, 79, 1 - 59." type="journal article" year="1986">Parnes (1986)</bibRefCitation>
from the Middle Triassic of
<collectingCountry id="99BE52AAD76F394C9E3AF9BE1A2E7D61" box="[1026,1085,1611,1635]" name="Israel" pageId="5" pageNumber="6">Israel</collectingCountry>
(Negev, Saharonim Formation, Lower Ladinian). It has a relatively wide umbilicus, a well-rounded elliptical whorl section, which is wider than high, and a regularly arched convex venter. Te present specimen is more evolute (uw/dm= 0.34 at dm =
<quantity id="2651BFDFD76F394C9F7FF93E1B867DE1" box="[1351,1429,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="70.0">70 mm</quantity>
) than
<taxonomicName id="26A969B9D76F394C9940F91E1AA67C01" box="[888,1205,1771,1795]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="D3DDCE28D76F394C9940F91E1AA67C01" box="[888,1205,1771,1795]" italics="true" pageId="5" pageNumber="6">Germanonautilus ellipticus</emphasis>
</taxonomicName>
(uw/dm = 0.24 at dm =
<quantity id="2651BFDFD76F394C9954F8FE1DD37C21" box="[876,960,1803,1827]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="mm" value="80.0">80 mm</quantity>
,
<bibRefCitation id="85386FCBD76F394C99E8F8F91A497C21" author="Parnes, A." box="[976,1114,1803,1827]" pageId="5" pageNumber="6" pagination="1 - 59" refId="ref8115" refString="Parnes, A. (1986). Middle Triassic cephalopods from the Negev (Israel) and Sinai (Egypt). Geological Survey of Israel, Bulletin, 79, 1 - 59." type="journal article" year="1986">Parnes 1986</bibRefCitation>
showing a wider umbilicus and a last whorl only slightly embracing the former volution. Up to now, forms similar to
<taxonomicName id="26A969B9D76D394E982FFB601C887FAE" box="[535,667,1172,1196]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="D3DDCE28D76D394E982FFB601C887FAE" box="[535,667,1172,1196]" italics="true" pageId="7" pageNumber="8">G. ellipticus</emphasis>
</taxonomicName>
were not documented in Southern Alps.
<taxonomicName id="26A969B9D76D394E9BC9FB411DBD7A01" authority="Smith, 1927" authorityName="Smith" authorityYear="1927" box="[497,942,235,1228]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="brooksi">
<emphasis id="D3DDCE28D76D394E9BC9FB411D147FCE" box="[497,775,1204,1228]" italics="true" pageId="7" pageNumber="8">Germanonautilus brooksi</emphasis>
<bibRefCitation id="85386FCBD76D394E9915FF1E1DBD7A01" author="Smith, J. P." box="[813,942,235,259]" pageId="7" pageNumber="8" refId="ref9130" refString="Smith, J. P. (1927). Upper Triassic marine invertebrate faunas of North America: U. S. Geological Survay, Professional Paper, 141, (pp. 135)" type="journal volume" year="1927">Smith, 1927</bibRefCitation>
</taxonomicName>
from Alaska (Carnian) described by
<bibRefCitation id="85386FCBD76D394E9F06FF1E1D607A21" author="Kummel, B." pageId="7" pageNumber="8" pagination="1 - 104" refId="ref7798" refString="Kummel, B. (1953). American Triassic Coiled Nautiloids, a study of the classification and evolution of late Paleozoic and early Mesozoic cephalopods, and description of new species. Geological Survey Professional Paper US Geological Survey, 250, 1 - 104." type="journal article" year="1953">Kummel (1953)</bibRefCitation>
, shows a well-rounded elliptical whorl section and a wide umbilicus. Following
<bibRefCitation id="85386FCBD76D394E9E4AFEDE1B047A41" author="Kummel, B." box="[1138,1303,299,323]" pageId="7" pageNumber="8" pagination="1 - 104" refId="ref7798" refString="Kummel, B. (1953). American Triassic Coiled Nautiloids, a study of the classification and evolution of late Paleozoic and early Mesozoic cephalopods, and description of new species. Geological Survey Professional Paper US Geological Survey, 250, 1 - 104." type="journal article" year="1953">Kummel (1953</bibRefCitation>
, p. 29). the specimens described originally by Smith are small conchs probably corresponding to immature growth stages, with deeply embracing whorls and more distinct umbilical shoulders.
</paragraph>
</subSubSection>
<caption id="B5D642B2D76C394F9A91F9151B907C46" ID-DOI="http://doi.org/10.5281/zenodo.12000854" ID-Zenodo-Dep="12000854" httpUri="https://zenodo.org/record/12000854/files/figure.png" pageId="6" pageNumber="7" startId="6.[169,200,1760,1780]" targetBox="[153,1438,238,1871]" targetPageId="6" targetType="figure">
<paragraph id="E116123AD76C394F9A91F9151B437DF6" blockId="6.[169,1411,1760,1860]" box="[169,1360,1760,1780]" pageId="6" pageNumber="7">
<emphasis id="D3DDCE28D76C394F9A91F9151EC87DF6" bold="true" box="[169,219,1760,1780]" pageId="6" pageNumber="7">Fig. 3</emphasis>
<taxonomicName id="26A969B9D76C394F9ADEF9151C5F7DF6" authority="Parnes, 1986" authorityName="Parnes" authorityYear="1986" box="[230,588,1760,1780]" class="Cephalopoda" family="Tainoceratidae" genus="Germanonautilus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="D3DDCE28D76C394F9ADEF9151F667DF6" box="[230,373,1760,1780]" italics="true" pageId="6" pageNumber="7">Germanonautilus</emphasis>
aff.
<emphasis id="D3DDCE28D76C394F9BAFF9151FCD7DF6" box="[407,478,1760,1780]" italics="true" pageId="6" pageNumber="7">ellipticus</emphasis>
<bibRefCitation id="85386FCBD76C394F9BDAF9151C5F7DF6" author="Parnes, A." box="[482,588,1760,1780]" pageId="6" pageNumber="7" pagination="1 - 59" refId="ref8115" refString="Parnes, A. (1986). Middle Triassic cephalopods from the Negev (Israel) and Sinai (Egypt). Geological Survey of Israel, Bulletin, 79, 1 - 59." type="journal article" year="1986">Parnes, 1986</bibRefCitation>
</taxonomicName>
, PIMUZ 29942,Valle Stelle/Tre Fontane (Monte San Giorgio), Middle Besano Formation (bed
</paragraph>
<paragraph id="E116123AD76C394F9A91F90E1B907C46" blockId="6.[169,1411,1760,1860]" pageId="6" pageNumber="7">
73), Secedensis Zone, Illyrian:
<emphasis id="D3DDCE28D76C394F9B9BF90E1FBD7C0D" bold="true" box="[419,430,1787,1807]" pageId="6" pageNumber="7">a</emphasis>
original specimen partially included in the matrix, lateral view,
<emphasis id="D3DDCE28D76C394F99FAF90E1DDD7C0D" bold="true" box="[962,974,1787,1807]" pageId="6" pageNumber="7">b</emphasis>
ventral view,associated with an external mold of
<taxonomicName id="26A969B9D76C394F9A91F8E31F7B7C28" box="[169,360,1814,1834]" class="Cephalopoda" family="Ceratitidae" genus="Stoppaniceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Ceratitida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="artinii">
<emphasis id="D3DDCE28D76C394F9A91F8E31F0F7C28" box="[169,284,1814,1834]" italics="true" pageId="6" pageNumber="7">Stoppaniceras</emphasis>
cf.
<emphasis id="D3DDCE28D76C394F9B00F8E31F7B7C28" box="[312,360,1814,1834]" italics="true" pageId="6" pageNumber="7">artinii</emphasis>
</taxonomicName>
,
<emphasis id="D3DDCE28D76C394F9B48F8E31F697C28" bold="true" box="[368,378,1814,1834]" pageId="6" pageNumber="7">c</emphasis>
plaster cast replica of the same specimen, lateral view,
<emphasis id="D3DDCE28D76C394F9974F8E31D4B7C28" bold="true" box="[844,856,1814,1834]" pageId="6" pageNumber="7">d</emphasis>
plaster cast replica of the same specimen,ventral view,
<emphasis id="D3DDCE28D76C394F9F09F8E31B2F7C28" bold="true" box="[1329,1340,1814,1834]" pageId="6" pageNumber="7">e</emphasis>
plaster cast replica of the same specimen, ventrolateral view,
<emphasis id="D3DDCE28D76C394F9848F8C51C647C46" bold="true" box="[624,631,1840,1860]" pageId="6" pageNumber="7">f</emphasis>
drawing of the ventral view showing the hypothetical whorl section (in grey the flank areas)
</paragraph>
</caption>
<subSubSection id="A9B341B1D76D394E997AFE3E1AEC7901" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="E116123AD76D394E997AFE3E1B057AE1" blockId="7.[813,1437,235,548]" box="[834,1302,459,483]" pageId="7" pageNumber="8">
Family
<taxonomicName id="26A969B9D76D394E99A9FE3E1B057AE1" authority="Mojsisovics, 1902" authorityName="Mojsisovics" authorityYear="1902" box="[913,1302,459,483]" class="Cephalopoda" family="Syringonautilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="family">
Syringonautilidae
<bibRefCitation id="85386FCBD76D394E9E6EFE3E1B057AE1" author="von Mojsisovics, E." box="[1110,1302,459,483]" pageId="7" pageNumber="8" pagination="173 - 356" refId="ref9458" refString="von Mojsisovics, E. (1902). Das Gebirge um Hallstatt. I Abteilung. Die Cephalopoden Der Hallstatter Kalke, Abhandlungen Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, Supplement-Heft., 6 (1), 173 - 356." type="journal article" year="1902">Mojsisovics, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="E116123AD76D394E997AFE1E1AEC7901" blockId="7.[813,1437,235,548]" box="[834,1279,491,515]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="26A969B9D76D394E99B7FE1E1AEC7901" authority="Mojsisovics, 1902" authorityName="Mojsisovics" authorityYear="1902" box="[911,1279,491,515]" class="Cephalopoda" family="Syringonautilidae" genus="Syringonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="D3DDCE28D76D394E99B7FE1E1A2A7901" box="[911,1081,491,515]" italics="true" pageId="7" pageNumber="8">Syringonautilus</emphasis>
<bibRefCitation id="85386FCBD76D394E9E07FE1E1AEC7901" author="von Mojsisovics, E." box="[1087,1279,491,515]" pageId="7" pageNumber="8" pagination="173 - 356" refId="ref9458" refString="von Mojsisovics, E. (1902). Das Gebirge um Hallstatt. I Abteilung. Die Cephalopoden Der Hallstatter Kalke, Abhandlungen Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, Supplement-Heft., 6 (1), 173 - 356." type="journal article" year="1902">Mojsisovics, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A9B341B1D76D394E997AFDF91B457921" box="[834,1366,523,548]" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="E116123AD76D394E997AFDF91B457921" blockId="7.[813,1437,235,548]" box="[834,1366,523,548]" pageId="7" pageNumber="8">
<typeStatus id="3E12AC98D76D394E997AFDF91D6B7926" box="[834,888,524,548]" pageId="7" pageNumber="8">Type</typeStatus>
species.
<taxonomicName id="26A969B9D76D394E99E1FDFE1B457921" authority="Mojsisovics, 1882" authorityName="Mojsisovics" authorityYear="1882" box="[985,1366,523,547]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="lilianus">
<emphasis id="D3DDCE28D76D394E99E1FDFE1A837921" box="[985,1168,523,547]" italics="true" pageId="7" pageNumber="8">Nautilus lilianus</emphasis>
<bibRefCitation id="85386FCBD76D394E9EAEFDFE1B457921" author="von Mojsisovics, E." box="[1174,1366,523,547]" pageId="7" pageNumber="8" pagination="1 - 322" refId="ref9428" refString="von Mojsisovics, E. (1882). Die Cephalopoden der Mediterranen Trias-Provinz. Abhandlungen Der Kaiserlich K o niglichen Geologischen Reichsanstalt, 10, 1 - 322." type="journal article" year="1882">Mojsisovics, 1882</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>