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<document id="494489A10036265D4DF5FCC6C23366B3" ID-DOI="10.1093/zoolinnean/zlae027" ID-ISSN="0024-4082" ID-Zenodo-Dep="14803321" ID-ZooBank="1953944-42BC-4858-81DA-3D800E2E0F54" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1738154448734" checkinUser="plazi" docAuthor="Zhang, Guoyi, Naggs, Fred, Andrus, Peter S. &amp; Wade, Christopher M." docDate="2025" docId="7013A309FFC3FFD7FEDCFE5FECA76239" docLanguage="en" docName="zlae027.pdf" docOrigin="Zoological Journal of the Linnean Society 203" docSource="https://doi.org/10.1093/zoolinnean/zlae027" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Pseudobuliminus Gredler 1886" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="8C2ADB71FFCBFFDFFFFFFFF3EC53601D" masterDocTitle="Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae" masterLastPageNumber="13" masterPageNumber="1" pageNumber="9" updateTime="1738685198826" updateUser="ExternalLinkService">
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<mods:title id="E7751391A6E7B3B5CA9F36D2F013879F">Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae</mods:title>
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<mods:namePart id="0FE8B4BB7CE2E9E1FFB35710845122E2">Zhang, Guoyi</mods:namePart>
<mods:affiliation id="508B0B9FC53FDC739C27FEB2AE483584">School of Life Sciences, University of Nottingham, Nottingham NG 7 2 RD, UK</mods:affiliation>
<mods:nameIdentifier id="E65A57396B2C317B26306977ED7B4CB5" type="email">starsareintherose@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D7E10F6566219F732A4CF94074E1FCEC">Naggs, Fred</mods:namePart>
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<mods:namePart id="2649A2312A913AC5C2D4D5E0B7D295D4">Andrus, Peter S.</mods:namePart>
<mods:affiliation id="58584B6723FC3D6B9AE4349AF19345CC">School of Life Sciences, University of Nottingham, Nottingham NG 7 2 RD, UK</mods:affiliation>
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<mods:namePart id="0A4BEAA715F4877BCA46F6A19420D990">Wade, Christopher M.</mods:namePart>
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<mods:part id="07E0291F0A8443D2D13BCDE675D92392">
<mods:date id="DAB7154F6D478AE1BFDFA8DFD3CDAEAA">2025</mods:date>
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<treatment id="7013A309FFC3FFD7FEDCFE5FECA76239" ID-DOI="http://doi.org/10.5281/zenodo.14764133" ID-Zenodo-Dep="14764133" LSID="urn:lsid:plazi:treatment:7013A309FFC3FFD7FEDCFE5FECA76239" httpUri="http://treatment.plazi.org/id/7013A309FFC3FFD7FEDCFE5FECA76239" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="B0A04194FFC3FFD7FEDCFE5FEE3561DB" box="[291,614,428,455]" pageId="8" pageNumber="9" type="nomenclature">
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<heading id="A34DA573FFC3FFD7FEDCFE5FEE3561DB" bold="true" box="[291,614,428,455]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="3FBA699CFFC3FFD7FEDCFE5FEE3561DB" ID-CoL="7PDRK" authority="Gredler, 1886" authorityName="Gredler" authorityYear="1886" box="[291,614,428,455]" class="Gastropoda" family="Bradybaenidae" genus="Pseudobuliminus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="CACECE0DFFC3FFD7FEDCFE5FEE3561DB" bold="true" box="[291,614,428,455]" pageId="8" pageNumber="9">
<emphasis id="CACECE0DFFC3FFD7FEDCFE5FED9D61DA" bold="true" box="[291,462,428,455]" italics="true" pageId="8" pageNumber="9">Pseudobuliminus</emphasis>
Gredler, 1886
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B0A04194FFC3FFD7FF7EFE1EECA76239" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="F805121FFFC3FFD7FF7EFE1EECA76239" blockId="8.[129,778,493,548]" pageId="8" pageNumber="9">
<emphasis id="CACECE0DFFC3FFD7FF7EFE1EED546219" box="[129,263,493,516]" italics="true" pageId="8" pageNumber="9">
<typeStatus id="2701ACBDFFC3FFD7FF7EFE1EECE26219" box="[129,177,493,516]" pageId="8" pageNumber="9">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="3FBA699CFFC3FFD7FEE8FE1EEEF36218" authority="Heude, 1882" authorityName="Heude" authorityYear="1882" box="[279,672,493,517]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="pseudobuliminus">
<emphasis id="CACECE0DFFC3FFD7FEE8FE1EEE536218" box="[279,512,493,517]" italics="true" pageId="8" pageNumber="9">Helix pseudobuliminus</emphasis>
Heude, 1882
</taxonomicName>
, absolute tautonymy.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="722D56CA28AE6BB7629D6F386A8E3E49" ID-DOI="10.1093/zoolinnean/zlae027" ID-ISSN="0024-4082" ID-ZooBank="1953944-42BC-4858-81DA-3D800E2E0F54" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738154448734" checkinUser="plazi" docAuthor="Zhang, Guoyi, Naggs, Fred, Andrus, Peter S. &amp; Wade, Christopher M." docDate="2025" docId="7013A309FFC3FFD7FEDCFE5FEF5A62E2" docLanguage="en" docName="zlae027.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203" docSource="https://doi.org/10.1093/zoolinnean/zlae027" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Pseudobuliminus Gredler 1886" docType="treatment" docVersion="1" masterDocId="8C2ADB71FFCBFFDFFFFFFFF3EC53601D" masterDocTitle="Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae" updateTime="1738154701448" updateUser="GgImagineBatch">
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<mods:title id="982A24757D8927A24AD07000DED38864">Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae</mods:title>
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<mods:nameIdentifier id="E605F131080DFBC8D36720B31BC17543" type="email">starsareintherose@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AB42B12E7FC0558BA12A32681898F722">Andrus, Peter S.</mods:namePart>
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<treatment id="7013A309FFC3FFD7FEDCFE5FEF5A62E2" LSID="urn:lsid:plazi:treatment:7013A309FFC3FFD7FEDCFE5FEF5A62E2" httpUri="http://treatment.plazi.org/id/7013A309FFC3FFD7FEDCFE5FEF5A62E2" pageId="8">
<subSubSection id="B0A04194FFC3FFD7FEDCFE5FEE3561DB" box="[291,614,428,455]" pageId="8" type="nomenclature">
<paragraph id="F805121FFFC3FFD7FEDCFE5FEE3561DB" blockId="8.[291,614,428,455]" box="[291,614,428,455]" pageId="8">
<heading id="A34DA573FFC3FFD7FEDCFE5FEE3561DB" bold="true" box="[291,614,428,455]" centered="true" fontSize="11" level="2" pageId="8" reason="2">
<taxonomicName id="3FBA699CFFC3FFD7FEDCFE5FEE3561DB" ID-CoL="7PDRK" authority="Gredler, 1886" authorityName="Gredler" authorityYear="1886" box="[291,614,428,455]" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" kingdom="Animalia" order="Stylommatophora" pageId="8" phylum="Mollusca" rank="genus">
<emphasis id="CACECE0DFFC3FFD7FEDCFE5FEE3561DB" bold="true" box="[291,614,428,455]" pageId="8">
<emphasis id="CACECE0DFFC3FFD7FEDCFE5FED9D61DA" bold="true" box="[291,462,428,455]" italics="true" pageId="8">Pseudobuliminus</emphasis>
Gredler, 1886
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B0A04194FFC3FFD7FF7EFE1EEF5A62E2" pageId="8" type="materials_examined">
<paragraph id="F805121FFFC3FFD7FF7EFE1EECA76239" blockId="8.[129,778,493,548]" pageId="8">
<emphasis id="CACECE0DFFC3FFD7FF7EFE1EED546219" box="[129,263,493,516]" italics="true" pageId="8">
<typeStatus id="2701ACBDFFC3FFD7FF7EFE1EECE26219" box="[129,177,493,516]" pageId="8">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="3FBA699CFFC3FFD7FEE8FE1EEEF36218" authority="Heude, 1882" authorityName="Heude" authorityYear="1882" box="[279,672,493,517]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="8" phylum="Mollusca" rank="species" species="pseudobuliminus">
<emphasis id="CACECE0DFFC3FFD7FEE8FE1EEE536218" box="[279,512,493,517]" italics="true" pageId="8">Helix pseudobuliminus</emphasis>
Heude, 1882
</taxonomicName>
, absolute tautonymy.
</paragraph>
<paragraph id="F805121FFFC3FFD7FF27FDB0EEE26243" blockId="8.[129,778,579,767]" box="[216,689,579,606]" pageId="8">
<emphasis id="CACECE0DFFC3FFD7FF27FDB0EEE26243" bold="true" box="[216,689,579,606]" pageId="8">
<taxonomicName id="3FBA699CFFC3FFD7FF27FDB0EE336243" authority="Zhang" authorityName="Zhang" authorityYear="2025" box="[216,608,579,606]" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" kingdom="Animalia" order="Stylommatophora" pageId="8" phylum="Mollusca" rank="species" species="dongyiicus" status="sp. nov.">
<emphasis id="CACECE0DFFC3FFD7FF27FDB0EE446243" bold="true" box="[216,535,579,606]" italics="true" pageId="8">Pseudobuliminus dongyiicus</emphasis>
Zhang
</taxonomicName>
<taxonomicNameLabel id="D1FD7376FFC3FFD7FD9AFDB7EEE26243" box="[613,689,580,606]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="F805121FFFC3FFD7FF7EFD99ED6862BC" blockId="8.[129,778,579,767]" pageId="8">
[
<uri id="8C2B1E1DFFC3FFD7FF78FD99ED6562BC" pageId="8">
urn:lsid:zoobank.org:act:
<uuid id="8C1C28CAFFC3FFD7FE5AFD99ED6562BC" pageId="8">D44A4F43-6A43-4C3B-95F8- 74C4221B5ABD</uuid>
</uri>
]
</paragraph>
<paragraph id="F805121FFFC3FFD7FF7EFD5AEF5A62E2" blockId="8.[129,778,579,767]" pageId="8">
The taxonomic description of
<taxonomicName id="3FBA699CFFC3FFD7FE33FD5AEE8B62DC" authorityName="Zhang" authorityYear="2025" box="[460,728,681,705]" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" kingdom="Animalia" order="Stylommatophora" pageId="8" phylum="Mollusca" rank="species" species="dongyiicus">
<emphasis id="CACECE0DFFC3FFD7FE33FD5AEE8B62DC" box="[460,728,681,705]" italics="true" pageId="8">Pseudobuliminus dongyiicus</emphasis>
</taxonomicName>
can be found in the Supporting Information, with photographs of
<taxonomicName id="3FBA699CFFC3FFD7FF7EFD14EDD162E2" authorityName="Zhang" authorityYear="2025" box="[129,386,743,767]" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" kingdom="Animalia" order="Stylommatophora" pageId="8" phylum="Mollusca" rank="species" species="dongyiicus">
<emphasis id="CACECE0DFFC3FFD7FF7EFD14EDD162E2" box="[129,386,743,767]" italics="true" pageId="8">Pseudobuliminus dongyiicus</emphasis>
</taxonomicName>
in theSupporting Information (Fig. S3).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="CACECE0DFFC3FFD7FF27FDB0EE446243" bold="true" box="[216,535,579,606]" italics="true" pageId="8" pageNumber="9">Pseudobuliminus dongyiicus</emphasis>
Zhang
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[
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urn:lsid:zoobank.org:act:
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The taxonomic description of
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can be found in the Supporting Information, with photographs of
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<emphasis id="CACECE0DFFC3FFD7FF7EFD14EDD162E2" box="[129,386,743,767]" italics="true" pageId="8" pageNumber="9">Pseudobuliminus dongyiicus</emphasis>
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in theSupporting Information (Fig. S3).
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The taxonomic description of
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<mods:title id="1F7B5E3B87551C595027B5F67956B9A8">Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae</mods:title>
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<mods:date id="5CC50405E91649480DCBE0E50656657D">2025</mods:date>
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Zhang
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[
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species:
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Crosse &amp; Debeaux, 1863
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, original designation.
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