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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.32.38161" ID-GBIF-Dataset="23cdd843-223d-4b59-b7c6-307e35e71c07" ID-GBIF-Taxon="159168993" ID-Pensoft-Pub="1314-2615-32-15" ID-Pensoft-UUID="6291936E45B856A1AA18C2B24BDDDC76" ID-ZooBank="731EF530E43D4FB8AAA39B4A0AD1AE09" ModsDocID="1314-2615-32-15" checkinTime="1566303228586" checkinUser="pensoft" docAuthor="Shear, William A. &amp; Steinmann, David B." docDate="2019" docId="2FF79DF1AF0F5DA69F29CB09B3DA0922" docLanguage="en" docName="SubterBiol 32: 15-32" docOrigin="Subterranean Biology 32" docSource="http://dx.doi.org/10.3897/subtbiol.32.38161" docTitle="Coloradesmus aquiliensis Shear 1984, comb. nov." docType="treatment" docVersion="3" id="6291936E45B856A1AA18C2B24BDDDC76" lastPageNumber="21" masterDocId="6291936E45B856A1AA18C2B24BDDDC76" masterDocTitle="Cave millipedes of the United States. XV. Coloradesmus gen. nov. (Diplopoda, Polydesmida, Macrosternodesmidae), and four new species from caves in Colorado, USA" masterLastPageNumber="32" masterPageNumber="15" pageId="6" pageNumber="21" updateTime="1643462002733" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Cave millipedes of the United States. XV. Coloradesmus gen. nov. (Diplopoda, Polydesmida, Macrosternodesmidae), and four new species from caves in Colorado, USA</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shear, William A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Steinmann, David B.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>32</mods:number>
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<mods:start>15</mods:start>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.32.38161</mods:identifier>
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<mods:identifier type="ZooBank">731EF530E43D4FB8AAA39B4A0AD1AE09</mods:identifier>
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<treatment ID-GBIF-Taxon="159168993" LSID="urn:lsid:plazi:treatment:2FF79DF1AF0F5DA69F29CB09B3DA0922" httpUri="http://treatment.plazi.org/id/2FF79DF1AF0F5DA69F29CB09B3DA0922" lastPageNumber="21" pageId="6" pageNumber="21">
<subSubSection pageId="6" pageNumber="21" type="nomenclature">
<paragraph pageId="6" pageNumber="21">
<taxonomicName LSID="2ff79df1-af0f-5da6-9f29-cb09b3da0922" authority="Shear, 1984" authorityName="Shear" authorityYear="1984" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus aquiliensis" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="aquiliensis">Coloradesmus aquiliensis (Shear, 1984)</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="21">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–6" captionStartId="F2" captionText="Figures 1 - 6. Coloradesmus aquiliensis male. 1 Collum, dorsal view 2 midbody ring, dorsal view 3 telson, dorsal view 4 right leg 7, posterior view 5 gonopods, ventral view 6 gonopods, posterior view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures1-6" httpUri="https://binary.pensoft.net/fig/326945" pageId="6" pageNumber="21">Figs 1-6</figureCitation>
,
<figureCitation captionStart="Figures 7–10" captionStartId="F3" captionText="Figures 7 - 10. Coloradesmus species males. 7 C. aquiliensis gonopods, anterior view 8 C. aquiliensis gonopods, lateral view 9 C. hopkinae collum, dorsal view 10 C. hopkinae midbody ring, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures7-10" httpUri="https://binary.pensoft.net/fig/326946" pageId="6" pageNumber="21">7-8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="reference_group">
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Diplopoda" family="Macrosternodesmidae" genus="Speodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speodesmus aquiliensis" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="aquiliensis">Speodesmus aquiliensis</taxonomicName>
Shear, 1984, p. 96.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="types">
<paragraph pageId="6" pageNumber="21">Types.</paragraph>
<paragraph pageId="6" pageNumber="21">Male holotype (American Museum of Natural History) from Fulford Cave, 21 mi SE of Eagle, Eagle Co.: Colorado, collected 6 July 1980 by J. R. Holsinger et al.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="diagnosis">
<paragraph pageId="6" pageNumber="21">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="21">
A larger species of
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus</emphasis>
</taxonomicName>
distinguished by details of the gonopods; in
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus hopkinsae" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="hopkinsae">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus hopkinsae</emphasis>
</taxonomicName>
the prefemoral process is much broader and has a laterodistal tooth.
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus manitou" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="manitou">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus manitou</emphasis>
</taxonomicName>
differs from
<taxonomicName genus="C." lsidName="C. aquiliensis" pageId="6" pageNumber="21" rank="species" species="aquiliensis">
<emphasis italics="true" pageId="6" pageNumber="21">C. aquiliensis</emphasis>
</taxonomicName>
in the endomerite, which in
<taxonomicName lsidName="manitou" pageId="6" pageNumber="21" rank="species" species="manitou">
<emphasis italics="true" pageId="6" pageNumber="21">manitou</emphasis>
</taxonomicName>
emerges from the acropodite at nearly a right angle, then turns sharply distad.
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus beckleyi" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="beckleyi">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus beckleyi</emphasis>
</taxonomicName>
and
<taxonomicName genus="C." lsidName="C. warneri" pageId="6" pageNumber="21" rank="species" species="warneri">
<emphasis italics="true" pageId="6" pageNumber="21">C. warneri</emphasis>
</taxonomicName>
are only half the length of the other three known species of the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="description">
<paragraph pageId="6" pageNumber="21">Corrections to original description.</paragraph>
<paragraph pageId="6" pageNumber="21">With the characters of the genus. Lengths of a series of specimens from Fulford Cave, the type locality, varied from 4.5-6.2 mm, the longer specimens strongly extended. Setae on collum, metazonites and telson long, acute. In the description of the gonopods, the prefemoral process is erroneously described as bifid and the endomerite is shown in the drawings as attached to the prefemoral process, giving that impression.</paragraph>
<caption doi="10.3897/subtbiol.32.38161.figures1-6" httpUri="https://binary.pensoft.net/fig/326945" pageId="6" pageNumber="21" start="Figures 1–6" startId="F2">
<paragraph pageId="6" pageNumber="21">
Figures 1-6.
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus aquiliensis" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="aquiliensis">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus aquiliensis</emphasis>
</taxonomicName>
male.
<emphasis bold="true" pageId="6" pageNumber="21">1</emphasis>
Collum, dorsal view
<emphasis bold="true" pageId="6" pageNumber="21">2</emphasis>
midbody ring, dorsal view
<emphasis bold="true" pageId="6" pageNumber="21">3</emphasis>
telson, dorsal view
<emphasis bold="true" pageId="6" pageNumber="21">4</emphasis>
right leg 7, posterior view
<emphasis bold="true" pageId="6" pageNumber="21">5</emphasis>
gonopods, ventral view
<emphasis bold="true" pageId="6" pageNumber="21">6</emphasis>
gonopods, posterior view. See text for explanation of labels.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.32.38161.figures7-10" httpUri="https://binary.pensoft.net/fig/326946" pageId="6" pageNumber="21" start="Figures 7–10" startId="F3">
<paragraph pageId="6" pageNumber="21">
Figures 7-10.
<taxonomicName authority="Shear &amp; Steinmann, 2019" authorityName="Shear &amp; Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus" order="Polydesmida" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="21">Coloradesmus</emphasis>
</taxonomicName>
species males.
<emphasis bold="true" pageId="6" pageNumber="21">7</emphasis>
<taxonomicName genus="C." lsidName="C. aquiliensis" pageId="6" pageNumber="21" rank="species" species="aquiliensis">
<emphasis italics="true" pageId="6" pageNumber="21">C. aquiliensis</emphasis>
</taxonomicName>
gonopods, anterior view
<emphasis bold="true" pageId="6" pageNumber="21">8</emphasis>
<taxonomicName genus="C." lsidName="C. aquiliensis" pageId="6" pageNumber="21" rank="species" species="aquiliensis">
<emphasis italics="true" pageId="6" pageNumber="21">C. aquiliensis</emphasis>
</taxonomicName>
gonopods, lateral view
<emphasis bold="true" pageId="6" pageNumber="21">9</emphasis>
<taxonomicName genus="C." lsidName="C. hopkinae" pageId="6" pageNumber="21" rank="species" species="hopkinae">
<emphasis italics="true" pageId="6" pageNumber="21">C. hopkinae</emphasis>
</taxonomicName>
collum, dorsal view
<emphasis bold="true" pageId="6" pageNumber="21">10</emphasis>
<taxonomicName genus="C." lsidName="C. hopkinae" pageId="6" pageNumber="21" rank="species" species="hopkinae">
<emphasis italics="true" pageId="6" pageNumber="21">C. hopkinae</emphasis>
</taxonomicName>
midbody ring, dorsal view. See text for explanation of labels.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="records">
<paragraph pageId="6" pageNumber="21">Records.</paragraph>
<paragraph pageId="6" pageNumber="21">
COLORADO: Eagle Co.: Fulford Cave, 10,000' asl, organic matter in packrat midden,
<geoCoordinate degrees="39" direction="north" minutes="29" orientation="latitude" precision="925" value="39.483334">39°29'N</geoCoordinate>
,
<geoCoordinate degrees="106" direction="west" minutes="33" orientation="longitude" precision="925" value="-106.55">106°33'W</geoCoordinate>
, 22 June 1999, 7 males, 13 females (topotypes);
<normalizedToken originalValue="Devil’s">Devil's</normalizedToken>
Den Cave, 11,500' (3506 m) asl, 35°F, under stones and logs in dark zone,
<geoCoordinate degrees="39" direction="north" minutes="30" orientation="latitude" precision="925" value="39.5">39°30'N</geoCoordinate>
,
<geoCoordinate degrees="106" direction="west" minutes="37" orientation="longitude" precision="925" value="-106.61667">106°37'W</geoCoordinate>
, 25 August 1999, 18 males, 7 females, juveniles;
<normalizedToken originalValue="Herbie’s">Herbie's</normalizedToken>
Deli Cave, 9200' asl, under rocks in dark zone,
<geoCoordinate degrees="40" direction="north" minutes="01" orientation="latitude" precision="925" value="40.016666">40°01'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="west" minutes="40" orientation="longitude" precision="925" value="-105.666664">105°40'W</geoCoordinate>
, 31 August 2003, mf; 15 August 2007, 2 males; Lime Creek Cave, 9200' (2804 m) asl,
<geoCoordinate degrees="40" direction="north" minutes="00" orientation="latitude" precision="925" value="40.0">40°00'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="west" minutes="40" orientation="longitude" precision="925" value="-105.666664">105°40'W</geoCoordinate>
, 5 September 2001, 2 males.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:titleInfo>
<mods:title>A review of the subfamily Acaenitinae Foerster, 1869 (Hymenoptera, Ichneumonidae) from Ukrainian Carpathians</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Varga, Alexander</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>1</mods:number>
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<taxonomicName authority="Gravenhorst, 1829" authorityName="Gravenhorst" authorityYear="1829" class="Insecta" family="Ichneumonidae" genus="Arotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arotes" order="Hymenoptera" pageId="0" pageNumber="1008" phylum="Arthropoda" rank="genus">Arotes Gravenhorst, 1829</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="1008" type="reference_group">
<paragraph pageId="0" pageNumber="1008">
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Arotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arotes" order="Hymenoptera" pageId="0" pageNumber="1008" phylum="Arthropoda" rank="genus">Arotes</taxonomicName>
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Arotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arotes albicinctus" order="Hymenoptera" pageId="0" pageNumber="1008" phylum="Arthropoda" rank="species" species="albicinctus">Arotes albicinctus</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="1008">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="1008">This genus is characterized by the combination of the following characters: clypeus transverse and basally flat, with transverse ridge, supra-antennal area with crest between antennal sockets, propodeum with well definded carinae, claws of all tarsi with appressed acute tooth, fore wing with areolet absent, intercubitus distal to vein 2m-cu, first metasomal tergite with white long setae on lateral and ventral parts.</paragraph>
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<subSubSection pageId="0" pageNumber="90015" type="nomenclature">
<paragraph pageId="0" pageNumber="90015">
<taxonomicName LSID="2FF7CA1F-E23D-5821-8B00-F9CA88434BF7" authority="(Floch &amp; Abonnenc, 1944)" baseAuthorityName="Floch &amp; Abonnenc" baseAuthorityYear="1944" class="Insecta" family="Psychodidae" genus="Psathyromyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psathyromyia inflata" order="Diptera" pageId="0" pageNumber="90015" phylum="Arthropoda" rank="species" species="inflata">Psathyromyia inflata (Floch &amp; Abonnenc, 1944)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="90015" type="distribution">
<paragraph pageId="0" pageNumber="90015">Distribution</paragraph>
<paragraph pageId="0" pageNumber="90015">Porto Velho</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="90015" type="notes">
<paragraph pageId="0" pageNumber="90015">Notes</paragraph>
<paragraph pageId="0" pageNumber="90015">
<bibRefCitation author="Galardo, Allan Kardec Ribeiro" journalOrPublisher="Revista da Sociedade Brasileira de Medicina Tropical" pageId="0" pageNumber="90015" pagination="265 - 271" refId="B7910410" refString="Galardo, Allan Kardec Ribeiro, Galardo, Clicia Denis, Silveira, Guilherme Abbad, Ribeiro, Kaio Augusto Nabas, Hijjar, Andrea Valadao, Oliveira, Liliane Leite, Dos Santos, Thiago Vasconcelos, 2015. Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil. Revista da Sociedade Brasileira de Medicina Tropical 48 (3): 265 - 271" title="Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil." volume="48" year="2015">Galardo et al. 2015</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>