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<document id="C1C91E46E2572852AE4809F08AC7A8AC" ID-DOI="10.3897/zookeys.1232.146855" ID-ZooBank="5778FBF6-2A76-4E42-A5A7-FA79CBAF56B9" ID-publisher-id="146855" URI-arpha="D1264EEB-CEDE-596D-9907-97E1726CB3F0" URI-zoobank="http://zoobank.org/5778FBF6-2A76-4E42-A5A7-FA79CBAF56B9" article-type="research-article" checkinTime="1742495071518" checkinUser="pensoft" docAuthor="Ni, Yi, Yu, Kun &amp; Zhang, Junxia" docDate="2025" docId="106A82DCC8FE5F1BBCD2EE4A99B8F0F0" docLanguage="en" docName="ZooKeys 1232: 343-355" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/146855/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Spartaeus siloi Ni, Yu &amp; Zhang, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="355" masterDocId="D1264EEBCEDE596D990797E1726CB3F0" masterDocTitle="Description of three new species of Spartaeini (Araneae, Salticidae, Spartaeinae) from China and Malaysia" masterLastPageNumber="355" masterPageNumber="343" pageNumber="343" updateTime="1742495462206" updateUser="ExternalLinkService">
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<mods:title id="D86C505BC401FD7C3DFED37796C4D147">Description of three new species of Spartaeini (Araneae, Salticidae, Spartaeinae) from China and Malaysia</mods:title>
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<mods:namePart id="06142BC6B31358B5355FEDD5C7C04C6A">Ni, Yi</mods:namePart>
<mods:nameIdentifier id="794B782EFB93596008188C55DB095AA9" type="ORCID">https://orcid.org/0009-0003-3239-3697</mods:nameIdentifier>
<mods:affiliation id="1A8C6DE21C5A776883CA2B45C9243B05">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="55DA12905CD8056784EB9FE8EBA85B8C">Yu, Kun</mods:namePart>
<mods:nameIdentifier id="4128B39C9CC365F98DAC4CAFF2DA4405" type="ORCID">0000-0002-5517-4484</mods:nameIdentifier>
<mods:affiliation id="BC6DE93D3675B09912A579CB536BF51A">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="D8CCB13CE54D63A905BC6FAC7D583C9D">Zhang, Junxia</mods:namePart>
<mods:nameIdentifier id="1D0B53F1E17E11CA97EF1F554D4E4BE6" type="ORCID">0000-0003-2179-3954</mods:nameIdentifier>
<mods:affiliation id="3FF5E7221287F59B6B02EFC5942AF053">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:date id="554E7DD6FEDB9E5B5FA12A6B54C1E497">2025</mods:date>
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<mods:number id="E72E7C1CC5CB78E56EF7AFEFFCBDE549">2025-03-20</mods:number>
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<treatment id="106A82DCC8FE5F1BBCD2EE4A99B8F0F0" ID-DOI="http://doi.org/10.5281/zenodo.15059388" ID-Zenodo-Dep="15059388" ID-arpha="106A82DC-C8FE-5F1B-BCD2-EE4A99B8F0F0" ID-zoobank="https://zoobank.org/531DF00D-0A70-4AB3-B507-89B1EEC6A5AE" LSID="urn:lsid:plazi:treatment:106A82DCC8FE5F1BBCD2EE4A99B8F0F0" LSID-ZBK="urn:lsid:zoobank.org:act:531DF00D-0A70-4AB3-B507-89B1EEC6A5AE" httpUri="http://treatment.plazi.org/id/106A82DCC8FE5F1BBCD2EE4A99B8F0F0">
<subSubSection id="CDA9C95B7848E6611B2FB3D5E48BCBA0" type="nomenclature">
<paragraph id="2F07254A648C7081BF01736859CFC21C">
<taxonomicName id="C675EE30B555E0D2497792988059C774" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="siloi" status="sp. nov.">
<emphasis id="898B1C0AEA590916B2B79E629776C149" italics="true">Spartaeus siloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="85DEC91FFA756D787A650A01A674A7F9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2A8DFC566B69102B0318C6AADAD53C93" type="description">
<paragraph id="D417FB651463B540FAC3D0A98855A5D9">
<figureCitation id="566A9589083DE53BE011E9B81B15FDA3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">Figs 3</figureCitation>
,
<figureCitation id="80FF4DD9F5512A46F8A599F8F4579622" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4</figureCitation>
,
<figureCitation id="D9A6C6F66358D894E206B05E4A1781E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 Common name. 诗雷散蛛</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAJAE" type="type material">
<paragraph id="F905E3DEFAC5C8DFCC9A7ADD1D78AF21">
<heading id="2A1F3989EA1AC1F2AF1B3F4AA85AE50E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="495CF5E0CB3118A1E0F72CE16B8EEFF2">
<materialsCitation id="7EAAA5B926F393E9BFB738AB5F1A1ACF" collectingDate="2024-04-27" collectorName="W. Wang" country="China" county="Daxin County" elevation="211" latitude="22.6931" location="Chongzuo City" longLatPrecision="5" longitude="107.03697" municipality="Autonomous Region" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="C9F7DAFE62327985F60035962D844671" bold="true" italics="true">
<typeStatus id="3550B14CA44B344204A8739CF55470F0" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="B4B55B4C4A952BF09AD390485D6EE8C3" name="China">China</collectingCountry>
<specimenCount id="7CCD581708E98BBF0FA160DF7BFC21C4" count="1" type="male"></specimenCount>
(MHBU-ARA- 00027416);
<collectingRegion id="694FFBF94B172425B6439A82DD720398" country="China" name="Guangxi">Guangxi Zhuang</collectingRegion>
<collectingMunicipality id="F7E60C1FAAE69D9304E3798AC6227B9A">Autonomous Region</collectingMunicipality>
,
<location id="8A77E7296930E95733F19BC6BA833FAE" LSID="urn:lsid:plazi:treatment:106A82DCC8FE5F1BBCD2EE4A99B8F0F0:8A77E7296930E95733F19BC6BA833FAE" country="China" county="Daxin County" latitude="22.6931" longLatPrecision="5" longitude="107.03697" municipality="Autonomous Region" name="Chongzuo City" stateProvince="Guangxi">Chongzuo City</location>
,
<collectingCounty id="9555F1C1181B887BAC35312745878990">Daxin County</collectingCounty>
,
<location id="89EEE07A620F5386126EBBFA1C4A80EE" LSID="urn:lsid:plazi:treatment:106A82DCC8FE5F1BBCD2EE4A99B8F0F0:89EEE07A620F5386126EBBFA1C4A80EE" country="China" county="Daxin County" latitude="22.6931" longLatPrecision="5" longitude="107.03697" municipality="Autonomous Region" name="Longdian" stateProvince="Guangxi">Longdian</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOJAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.036975,22.693100]}">
<geoCoordinate id="C7ECA982DA2D6332E4699BB78DEE6AB5" degrees="22.6931" direction="north" orientation="latitude" precision="5" value="22.6931">22.6931 ° N</geoCoordinate>
,
<geoCoordinate id="B16469BC6E7B792DFAB57272C6514334" degrees="107.036975" direction="east" orientation="longitude" precision="1" value="107.03697">107.036975 ° E</geoCoordinate>
</named-content>
,
<quantity id="52ACF0FF2EFDAA9B7A8B79C2D3C4D123" metricMagnitude="2" metricUnit="m" metricValue="2.11" unit="m" value="211.0">
<elevation id="324F8F822BFDA3EAD6B6328C3E2435D9" metricMagnitude="2" metricUnit="m" metricValue="2.11" unit="m" value="211.0">211 m</elevation>
</quantity>
elev.;
<date id="7E66F7ECA4092EF0541DE1B20A77332B" value="2024-04-27">
<collectingDate id="E03E0D7C0CDBA3C6D1400C8CA1754F5C" value="2024-04-27">27. IV. 2024</collectingDate>
</date>
;
<collectorName id="CA08C26A9CBE7DC3F9B67D07C28F065E">W. Wang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="25A86EA7DD9C03A34200DBDC49CE4FA1" collectingDate="2024-05-12" collectorName="J. Zhang &amp; K. Yu &amp; Z. Yang &amp; Y. Ni &amp; Y. Li" country="China" county="Daxin County" elevation="145" latitude="22.6538" location="Chongzuo City" longLatPrecision="7" longitude="107.0755" municipality="Autonomous Region Autonomous Region" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="32D70C9EB602ED39BC43CADA0BA1C439" bold="true" italics="true">
<typeStatus id="277D00689BF2BC744553BC7AE5158100" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="C538FE6E76614ABF2A69632A82721406" name="China">China</collectingCountry>
<specimenCount id="9C001FB08C0ADA3910025D9EA1E40889" count="1" type="female">1 ♀</specimenCount>
(MHBU-ARA- 00027417); same data as holotype;
<specimenCount id="72E7338FDC01A1C7538582C6E1E7DF45" count="1" type="male">1 ♂</specimenCount>
(MHBU-ARA- 00027538);
<collectingRegion id="193DE8ED0F69B838FA0141714E9E2A57" country="China" name="Guangxi">Guangxi Zhuang</collectingRegion>
<collectingMunicipality id="925425CC51FB17020B5516C108B49043">Autonomous Region Autonomous Region</collectingMunicipality>
,
<location id="33D7C780CFA57871C8F53D8A94EC47EB" LSID="urn:lsid:plazi:treatment:106A82DCC8FE5F1BBCD2EE4A99B8F0F0:33D7C780CFA57871C8F53D8A94EC47EB" country="China" county="Daxin County" latitude="22.6538" longLatPrecision="7" longitude="107.0755" municipality="Autonomous Region Autonomous Region" name="Chongzuo City" stateProvince="Guangxi">Chongzuo City</location>
,
<collectingCounty id="7CBE263E955D7ACDF425FC67B58F35A9">Daxin County</collectingCounty>
,
<location id="68FF94B35B7C67F7B069EF3602386F69" LSID="urn:lsid:plazi:treatment:106A82DCC8FE5F1BBCD2EE4A99B8F0F0:68FF94B35B7C67F7B069EF3602386F69" country="China" county="Daxin County" latitude="22.6538" longLatPrecision="7" longitude="107.0755" municipality="Autonomous Region Autonomous Region" name="Xinkang" stateProvince="Guangxi">Xinkang</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZJAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.075500,22.653800]}">
<geoCoordinate id="7D5FFE9E08A3FA0F60394D90C6D66D1D" degrees="22.6538" direction="north" orientation="latitude" precision="5" value="22.6538">22.6538 ° N</geoCoordinate>
,
<geoCoordinate id="5DA70CB99E088E1E23E0182FCC0D7F4B" degrees="107.0755" direction="east" orientation="longitude" precision="5" value="107.0755">107.0755 ° E</geoCoordinate>
</named-content>
,
<quantity id="B2BFBB7A75F5F54D00BB089D997D9421" metricMagnitude="2" metricUnit="m" metricValue="1.45" unit="m" value="145.0">
<elevation id="D3827E6CF21DEA045FB9038518DAA867" metricMagnitude="2" metricUnit="m" metricValue="1.45" unit="m" value="145.0">145 m</elevation>
</quantity>
elev.;
<date id="CB8489E477A4AE485AC0FBA5919ED217" value="2024-05-12">
<collectingDate id="03A8E9582B20E64C3FCEE0CFA5FCD0E9" value="2024-05-12">12. V. 2024</collectingDate>
</date>
;
<collectorName id="E7DCBC200A93480C46C767ED57D31E40">J. Zhang</collectorName>
,
<collectorName id="F6B6D1AB28B42EF759A4CD46607E8692">K. Yu</collectorName>
,
<collectorName id="9E99445FAE738DABD17EBFC365C658BB">Z. Yang</collectorName>
,
<collectorName id="A6B0CC060974C9D7AC0F643D007D7D9A">Y. Ni</collectorName>
&amp;
<collectorName id="C2CC8347FACD37F346E7DDCBB3D4F250">Y. Li</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5JAE" type="etymology">
<paragraph id="6030347112B827A0FE5C65F000392E39">
<heading id="ABD9826A907BEF961AB91D5551E28B40" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="8C48C95DAAC8932C5E6091B801D917DA">
The specific epithet is derived from “ Si Loi ”, which is a unique artistic style of Zhuang folk songs in Daxin County (the
<typeStatus id="F4BA3CD4758F8FEEB32EACDD35661234">type</typeStatus>
locality). Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDKAE" type="diagnosis">
<paragraph id="998D976256D4D6304FAE6CCC3515A231">
<heading id="C9A648C15489887688F60FB8F503F8C1" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6D8DF1B44AD70320E761C1D73DD03233">
The new species resembles
<taxonomicName id="81F118B4CEB5B4BC1C711707D73A0C01" authority="Song, Chen &amp; Gong, 1991" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="1957A4F7147C5D6A6AE8012E258F78BE" italics="true">S. platnicki</emphasis>
<bibRefCitation id="800AD6F176CAD8E3739957768E1C866E" author="Song DX &amp; Chen ZQ &amp; Gong LS" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="424-427" refId="B6" refString="Song DX, Chen ZQ, Gong LS (1991) A new species of the genus Spartaeus from China (Araneae: Salticidae). Acta Zootaxonomica Sinica 16: 424-427." title="A new species of the genus Spartaeus from China (Araneae: Salticidae)." volume="16" year="1991">
Song, Chen &amp; Gong,
<quantity id="BAC04E899E092347B10CD89F3D4EC9C2" metricMagnitude="1" metricUnit="m" metricValue="5.05714" unit="in" value="1991.0">1991 in</quantity>
</bibRefCitation>
</taxonomicName>
the shape of the palpal bulb and copulatory duct, but it can be distinguished by the following: (1) the length of the embolus is much less than half of the bulb width (Figs
<figureCitation id="04C50A5C6D675A8721803264B3E66FAB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 B</figureCitation>
,
<figureCitation id="1ECA93D6763936E357B77EDE6114C6CC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 B</figureCitation>
; vs. in
<taxonomicName id="131F4B2F416441F20214E2D116E44FD8" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="CD5AE859CFC47F814621D39BCBEFB49A" italics="true">S. platnicki</emphasis>
</taxonomicName>
, the length of the embolus is more than half of bulb width; see
<bibRefCitation id="3DEEACDB4D548A5BBF5092A7EB4E3B43" DOI="10.11646/zootaxa.5389.1.1" author="Lin" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zootaxa" pagination="1-33" refId="B2" refString="Lin Y, Xin Y, Wang C, Li S (2023) On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae). Zootaxa 5389 (1): 133. https://doi.org/10.11646/zootaxa.5389.1.1" title="On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae)." volume="5389" year="2023">Lin et al. 2023</bibRefCitation>
: fig. 18 A); (2) the end of the retrolateral tibial apophysis is relatively straight (Figs
<figureCitation id="F06916C57E946FE91A728345398C43D2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 B</figureCitation>
,
<figureCitation id="1CDCACB50CEFDABCEFE517020CAF250A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 B</figureCitation>
; vs. in
<taxonomicName id="1830ED4FA472D1E3E491B88E84A81F8E" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="EF36AD480CC3FF06FF2ED45309CDF4E6" italics="true">S. platnicki</emphasis>
</taxonomicName>
, the end of the retrolateral tibial apophysis is strongly curved; see
<bibRefCitation id="995F71156880246F7AD102780F6EB90C" DOI="10.11646/zootaxa.5389.1.1" author="Lin" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zootaxa" pagination="1-33" refId="B2" refString="Lin Y, Xin Y, Wang C, Li S (2023) On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae). Zootaxa 5389 (1): 133. https://doi.org/10.11646/zootaxa.5389.1.1" title="On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae)." volume="5389" year="2023">Lin et al. 2023</bibRefCitation>
: fig. 18 A); (3) the ventral intermediate tibial apophysis is rounded at the distal end in ventral view (Figs
<figureCitation id="5EDBF44A505676D4A76178B48C16B266" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 B</figureCitation>
,
<figureCitation id="545D61967503EDCCEFAC26D177051AC0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 B</figureCitation>
; vs. in
<taxonomicName id="92A6997DAE19AEDC9621C609E84D0F2B" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="30DDE0CDFAF071332C8CF1FE5AF7314F" italics="true">S. platnicki</emphasis>
</taxonomicName>
, the ventral intermediate tibial apophysis is sharp at the distal end in ventral view; see
<bibRefCitation id="253DAFD21E6045D6F70505DCBD458DC4" DOI="10.11646/zootaxa.5389.1.1" author="Lin" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zootaxa" pagination="1-33" refId="B2" refString="Lin Y, Xin Y, Wang C, Li S (2023) On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae). Zootaxa 5389 (1): 133. https://doi.org/10.11646/zootaxa.5389.1.1" title="On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae)." volume="5389" year="2023">Lin et al. 2023</bibRefCitation>
: fig. 18 A); and (4) the two copulatory ducts are nearly parallel (Figs
<figureCitation id="388EA461AE6D82DB738AA5A4EA7F26A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 D</figureCitation>
,
<figureCitation id="F6F5942E549AC2B7915B6A3E23154CBE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 D</figureCitation>
; vs. curved posteriorly in
<taxonomicName id="A5611A9130FB49423D4FE4ABF61334EE" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="3E916E21CD26DA18A0BCD13EAC5AF8FD" italics="true">S. platnicki</emphasis>
</taxonomicName>
; see
<bibRefCitation id="DAA069DA732500E149D4969C0B7FF669" DOI="10.11646/zootaxa.5389.1.1" author="Lin" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zootaxa" pagination="1-33" refId="B2" refString="Lin Y, Xin Y, Wang C, Li S (2023) On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae). Zootaxa 5389 (1): 133. https://doi.org/10.11646/zootaxa.5389.1.1" title="On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae)." volume="5389" year="2023">Lin et al. 2023</bibRefCitation>
: fig. 19 A).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQNAE" type="description">
<paragraph id="0AFD84E5C6BEB264C7FDA134BF7E4EEB">
<heading id="49BCA4FD3F874C786D6F50743C8DC698" reason="title">Description.</heading>
</paragraph>
<paragraph id="CC3A4A746C233BB03AD3ACC26126DD12">
<emphasis id="B6876279F44E976BD6154582A455C16A" bold="true">Male.</emphasis>
Measurements of
<typeStatus id="BFF39787E99FB8599D75A2A7BC84F506" type="holotype">holotype</typeStatus>
: total length 7.19. Carapace 3.42 long, 2.37 wide. Abdomen 3.58 long, 1.56 wide. Eye measurements:
<abbrev id="ABBRID0EYNAE" xlink_title="anterior median eye">
<collectionCode id="E381BB9D1C0FDF791515E1DF13978693">AME</collectionCode>
</abbrev>
0.81,
<abbrev id="ABBRID0E3NAE" xlink_title="anterior lateral eye">
<collectionCode id="FEEFBFE46FBBD34A2D3C37D31A3E22FD">ALE</collectionCode>
</abbrev>
0.45,
<abbrev id="ABBRID0EAOAE" xlink_title="posterior median eye">
<collectionCode id="4E2DD834BC52F3CFA11BA05FDE6C8D7E">PME</collectionCode>
</abbrev>
0.30,
<abbrev id="ABBRID0EEOAE" xlink_title="posterior lateral eye">
<collectionCode id="12A169C9DD21F8AA1CEAF9EDEB0A368B">PLE</collectionCode>
</abbrev>
0.30. Leg measurements: I 15.55 (4.43, 1.28, 4.70, 3.7, 1.44), II 9.29 (2.30, 0.91, 2.61, 2.40, 1.07), III 10.05 (2.67, 0.89, 2.38, 2.53, 1.58), IV 13 (3.16, 1.07, 3.39, 4.19, 1.19); leg formula 1432. Chelicera (Fig.
<figureCitation id="E872EE118DB3B979C5DD9EE01168CC05" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 D</figureCitation>
) dark brown, with seven promarginal and eight retromarginal teeth. Carapace (Fig.
<figureCitation id="7BE4EE5D563CA5FBBDE8B88FA4FD6AA2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 B, C</figureCitation>
) yellow, margin and eye surroundings blackish-brown, anterior lateral margin deep black, eye field with black setae, eye surroundings covered by yellow and white setae, fovea dark and slit-like; endites and labium yellowish-brown, with black setae at distal end, endites slender. Legs (Fig.
<figureCitation id="BD50EACC2E92F7A6F6BAD86018F44064" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 B</figureCitation>
) with long and thin spines, coxa and trochanter off-white, femur to tibia with obvious grey stripes on anterior and posterior sides. Abdomen (Fig.
<figureCitation id="C5CDF23E8C4E8425A91DB88A63920943" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 B</figureCitation>
) yellow, black strips interspersed with white setae around margins, broken near spinnerets; dorsal side with tawny patches and black setae; ventral side off-white, with white setae on both sides, center covered with greyish-white scales. Spinnerets (Fig.
<figureCitation id="743EAB5AFF33FAD602ABA8B58363711C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 B</figureCitation>
) white, spinnerets lateral inward white, lateral exterior with dark brown patches, covered with white setae.
</paragraph>
<caption id="D80B79F4127E56617DCA63BB9DED00C9" ID-DOI="10.3897/zookeys.1232.146855.figure3" ID-arpha="18A09ACE-9544-5353-A06F-65229F53275A" httpUri="https://binary.pensoft.net/fig/1286792" startId="F3">
<paragraph id="0CD7D76E7B78FD50ED6D0941C68D441E">
<label id="492372B4DC16602C3C41544BEC7C4F42">Figure 3.</label>
</paragraph>
<paragraph id="B7A286C8F81EF657B9F4CF3876CA495A">
<taxonomicName id="C25742A946E0837C5EE8CA7B422B2E7E" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="siloi" status="sp. nov.">
<emphasis id="2072E0D4FE8086BF9AF0DDAF850DAF43" italics="true">Spartaeus siloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF20D690878FB232C71CCA044C3FA83" rank="species">sp. nov.</taxonomicNameLabel>
, habitus of holotype male (
<emphasis id="D05D393511CD7237A71417DFA1475AE2" bold="true">B, C)</emphasis>
and paratype female (
<emphasis id="5AB754658BE887F57433967D6B7E2846" bold="true">A, E</emphasis>
), cheliceral teeth of holotype male (
<emphasis id="2182070A84583B219B43E981EB63D3B3" bold="true">D</emphasis>
) and paratype female (
<emphasis id="7A7D419505C18EA523516600523E276F" bold="true">F</emphasis>
); in dorsal (
<emphasis id="B33B0EBB0B1C9FDEB044FE7B44F756E3" bold="true">A, B</emphasis>
), front (
<emphasis id="222ABCBB1C7697BECA3A881748AAC6C2" bold="true">C, E</emphasis>
) and back (
<emphasis id="894F34A4E85029B8208B2E5CFA32C01C" bold="true">D, F</emphasis>
) view. Scale bars: 0.1 mm (
<emphasis id="3D01A6DDE85E01B0EB440BE1DAE774CF" bold="true">A, B</emphasis>
); 0.5 mm (
<emphasis id="60F6E721016520843A0B2DD785BDE056" bold="true">C F</emphasis>
).
</paragraph>
</caption>
<caption id="A673B32DD030B4DF80E6405A8EC4C55D" ID-DOI="10.3897/zookeys.1232.146855.figure4" ID-arpha="474F1057-6F0F-5EBB-AC05-29E8E296E63D" httpUri="https://binary.pensoft.net/fig/1286793" startId="F4">
<paragraph id="73D10F6B9E8ACD48F54E541EE0E1725E">
<label id="581BBAE7BA48817F16780973839E9780">Figure 4.</label>
</paragraph>
<paragraph id="9792488410489DBE3105598F3029E8A7">
<taxonomicName id="9F428B43A002E61ABD1EACF93556E620" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="siloi" status="sp. nov.">
<emphasis id="9440403EA0E6192081F6644FB8B3D1AB" italics="true">Spartaeus siloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17AEB0B28B2B4228031995D795712C79" rank="species">sp. nov.</taxonomicNameLabel>
, palp of holotype male (
<emphasis id="03281F4D70BA273E96EE77BB41ECABD6" bold="true">A C</emphasis>
), epigyne (
<emphasis id="A97BAF1822D6526F7CE783C6880E3642" bold="true">D</emphasis>
) and vulva (
<emphasis id="7020A154CA4F91B52ADF13ACAFCE0EE3" bold="true">E</emphasis>
) of paratype female; in prolateral (
<emphasis id="F5245D6F20E82CBD1B0AEFA83C317F52" bold="true">A</emphasis>
), ventral (
<emphasis id="80180807CB246D79291187147DA92E59" bold="true">B, D</emphasis>
), retrolateral (
<emphasis id="4476A07EF29921FC788F36978F6F4492" bold="true">C</emphasis>
) and dorsal (
<emphasis id="E118467A2CDA80AF6AA2ED331E3C8F5F" bold="true">E</emphasis>
) view. Scale bars: 0.2 mm (
<emphasis id="23BD091755C75C71CA97EB92A35FE189" bold="true">A E</emphasis>
).
</paragraph>
</caption>
<caption id="C6185635A4F0AA530C413773C7812B4E" ID-DOI="10.3897/zookeys.1232.146855.figure5" ID-arpha="BAE8DCF4-AAFA-5CDB-80B6-484A7D573BAF" httpUri="https://binary.pensoft.net/fig/1286794" startId="F5">
<paragraph id="33D7E94154649F8AAB3937541BBA5627">
<label id="420831E7D405902E81AF3383506E163E">Figure 5.</label>
</paragraph>
<paragraph id="DF3EA64FF1B800759FA6B5C55AFF99AE">
<taxonomicName id="2AB139CC021AB050110711C070BEAA6F" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="siloi" status="sp. nov.">
<emphasis id="82F51A9434EA14EA32CB9D87F34F1FE0" italics="true">Spartaeus siloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB1DA6118DDF09FC1E761EDBFD165755" rank="species">sp. nov.</taxonomicNameLabel>
, palp of holotype male (
<emphasis id="2FB4FEA6CB63CAB1A221822EC290C128" bold="true">A C</emphasis>
), epigyne (
<emphasis id="EB3F8973B0525FCBD860662F1FCB5826" bold="true">D</emphasis>
) and vulva (
<emphasis id="3B30E5D11F8689346C2DF273A7BB8328" bold="true">E</emphasis>
) of paratype female; in prolateral (
<emphasis id="F62782FB6F84C1E90AC4A59319CBC96E" bold="true">A</emphasis>
), ventral (
<emphasis id="455053E4CA684D4C3CB2F54FB68C2B50" bold="true">B, D</emphasis>
), retrolateral (
<emphasis id="12EFA0C35297DC01DC9B3FF90415C7E4" bold="true">C</emphasis>
) and dorsal (
<emphasis id="5E9DBA1006B156774C7865D10A5D3209" bold="true">E</emphasis>
) view. Scale bars: 0.2 mm (
<emphasis id="9A33F959A6A37DD92F82D8A2B36574BE" bold="true">A E</emphasis>
).
</paragraph>
</caption>
<paragraph id="3FA16C370F4DDA3ACD56E9E2059FDDA6">
Palp (Figs
<figureCitation id="2929923E9AA019802FAD454CE815F3BF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 A C</figureCitation>
,
<figureCitation id="32244FD1A30885503371F9F3B4651667" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 A C</figureCitation>
): embolus thin and short, sheep-horn-like in retrolateral view, base of embolus hidden behind membrane formed by distal haematodocha; tegular oval, surface smooth; tegular apophysis triangular, weakly sclerotized in ventral view; ventral tibial apophysis thick, triangular in ventral view; retrolateral tibial apophysis soft and movable, long and flat in ventral view, with small processes on distal end; dorsal intermediate tibial apophysis blackish-brown, with snake-scale-like processes on outer surface, edge wide and flat; ventral intermediate tibial apophysis yellowish-brown, rounded at distal end in ventral view; distal end of cymbium with grey hair tuft.
</paragraph>
<paragraph id="ADF4E5F9046BDADC74CB0F9369578CBC">
<emphasis id="410D378BDBB9C05A5E1B15C96FD105E8" bold="true">Female.</emphasis>
Measurements of
<typeStatus id="075C7C0D9603C18CD5E58BA9209947A6" type="paratype">paratype</typeStatus>
: total length 6.55. Carapace 2.92 long, 2.31 wide. Abdomen 3.33 long, 2.07 wide. Eye measurements:
<abbrev id="ABBRID0EETAE" xlink_title="anterior median eye">
<collectionCode id="9A5C9FEA821BCA67C6D93D72CE1EC0F4">AME</collectionCode>
</abbrev>
0.68,
<abbrev id="ABBRID0EITAE" xlink_title="anterior lateral eye">
<collectionCode id="C46723F274BFA622CDFC2548175618BD">ALE</collectionCode>
</abbrev>
0.40,
<abbrev id="ABBRID0EMTAE" xlink_title="posterior median eye">
<collectionCode id="B5E53670A713B9B4B705A7E9065C0F68">PME</collectionCode>
</abbrev>
0.28,
<abbrev id="ABBRID0EQTAE" xlink_title="posterior lateral eye">
<collectionCode id="24F6246E703B02B73B1C6FCA46F72CEC">PLE</collectionCode>
</abbrev>
0.39. Leg measurements: I 9.15 (2.72, 1.03, 2.70, 1.84, 0.86), II 7.16 (2.25, 0.76, 1.88, 1.53, 0.74), III 7.45 (2.19, 074, 1.84, 1.83, 0.85), IV 8.68 (2.85, 0.72, 2.67, 1.52, 0.92); leg formula 1432. Chelicera (Fig.
<figureCitation id="F96050D290B080320357B5C3F562E014" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 F</figureCitation>
) brown, with six promarginal and eight retromarginal teeth. Habitus (Fig.
<figureCitation id="2C84C052DE7B9EDFC79254DFEDBD9497" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Spartaeus siloi sp. nov., habitus of holotype male (B, C) and paratype female (A, E), cheliceral teeth of holotype male (D) and paratype female (F); in dorsal (A, B), front (C, E) and back (D, F) view. Scale bars: 0.1 mm (A, B); 0.5 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure3" httpUri="https://binary.pensoft.net/fig/1286792">3 A, E</figureCitation>
) similar to that of male, but different in: carapace paler at margin forming a black band from anterior lateral eyes to rear edge of carapace; with a black band in lower position between the anterior median eyes and the posterior lateral eyes; black strips on lateral dorsal side of abdomen not continuous, but with intermittent small black bands.
</paragraph>
<paragraph id="5C6B2679C6B0BFF1B000E9AFEA221494">
Epigyne (Figs
<figureCitation id="FE743A5FDDC035E1C1688034B1E6202A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure4" httpUri="https://binary.pensoft.net/fig/1286793">4 D, E</figureCitation>
,
<figureCitation id="A07288C3062EEDC2CAE6E27DB097BA38" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spartaeus siloi sp. nov., palp of holotype male (A C), epigyne (D) and vulva (E) of paratype female; in prolateral (A), ventral (B, D), retrolateral (C) and dorsal (E) view. Scale bars: 0.2 mm (A E)." figureDoi="10.3897/zookeys.1232.146855.figure5" httpUri="https://binary.pensoft.net/fig/1286794">5 D, E</figureCitation>
): epigynal plate oval, slightly raised in a mound-like shape; copulatory openings small, slit-shaped in ventral view, anteriorly located; copulatory ducts extending to posterior part, dark and slender; spermathecae reddish-brown, large and kidney-shaped; fertilization ducts short, connecting with spermathecae at posterior part; accessory glands located centrally at inner margins of spermathecae, partly hidden by spermathecae in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGUAE" type="distribution">
<paragraph id="9E254954CBC3CA3DC328CCDD40CBAA82">
<heading id="E600DB73548F57C0E4ADF3A10F05E3F5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A5F69C5615C959F1FD9015CA999CD351">
<collectingCountry id="8101EF3AC263CA889EA1628EF60998B3" name="China">China</collectingCountry>
(
<collectingRegion id="E003CBA68E810195167FB0DF9FC44BA9" country="China" name="Guangxi">Guangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELUAE" type="comments">
<paragraph id="67D32C8F4E2A79F09C0AD85CEE180D6A">
<heading id="26AB242276BE7313A18F87F0124FB9FF" reason="title">Comments.</heading>
</paragraph>
<paragraph id="728156B0DDBFA1418701943325EAA543">
Following a previous study on
<taxonomicName id="1149FA74AFB1B3C40A997F8B944E6382" authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="DC6327B0A2EC0E7E18A23E95BCB9FF71" italics="true">Spartaeus</emphasis>
</taxonomicName>
(
<bibRefCitation id="BBA4C133FF54CFAE92763A81B8A0D37E" DOI="10.11646/zootaxa.4303.3.4" author="Yang" etAl="et al." firstAuthor="Yang" issue="3" journalOrPublisher="Zootaxa" pagination="379-394" refId="B15" refString="Yang SF, Liu W, Liu P, Peng XJ (2017) Five new species of Spartaeus Thorell, 1891 from China (Araneae: Salticidae). Zootaxa 4303 (3): 379394. https://doi.org/10.11646/zootaxa.4303.3.4" title="Five new species of Spartaeus Thorell, 1891 from China (Araneae: Salticidae)." volume="4303" year="2017">Yang et al. 2017</bibRefCitation>
: figs 4, 8, 10), we labelled the weekly sclerotized apophysis derived from the tegulum of male palpal bulb as
<abbrev id="ABBRID0E3UAE" xlink_title="tegular apophysis">
<collectionCode id="46212E24727484A6A5A4D49F4B7DA165" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
. This structure was identified as M
<quantity id="D516A43A2F49E93B9C5539800DC28B0E" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">
<subScript id="6CCBA0255A5AEB1E26730A8CF24B5D26">3</subScript>
in
</quantity>
<bibRefCitation id="C2A6C9E91AC7D0F353BDB0BA74219C2D" DOI="10.5962/bhl.part.15964" author="Wanless FR" firstAuthor="Wanless" journalOrPublisher=")" pagination="135-205" refId="B10" refString="Wanless FR (1984) A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool.) 46: 135205. https://doi.org/10.5962/bhl.part.15964" title="A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool." volume="46" year="1984">Wanless (1984)</bibRefCitation>
. Additionally, the membranous patch of the distal haematodocha at the base of embolus is referred to here as M, aligning with the designation of M
<quantity id="47430B39343D5A029C23512D97446027" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">
<subScript id="51A8C0A38BCDF16D578B3AC011C3479E">1</subScript>
in
</quantity>
<bibRefCitation id="1D2956EBD1C674CF799F64D9501B8BE6" DOI="10.5962/bhl.part.15964" author="Wanless FR" journalOrPublisher=")" pagination="135-205" refId="B10" refString="Wanless FR (1984) A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool.) 46: 135205. https://doi.org/10.5962/bhl.part.15964" title="A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool." volume="46" year="1984">Wanlesss (1984</bibRefCitation>
) study on
<taxonomicName id="257622E3CA50F93E9CEF513F0F6B5A6D" authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="4CE38D8C01760E2A8A24EBDC65B48D85" italics="true">Spartaeus</emphasis>
</taxonomicName>
. In the redescription of
<taxonomicName id="104BDE0F132546CF027A7BF6DA20A350" authority="Song, Chen &amp; Gong, 1991" authorityName="Song, Chen &amp; Gong" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="5F4BF0ECF741C12FDE12495F4A77B630" italics="true">S. platnicki</emphasis>
<bibRefCitation id="564D2EF0FB57B1473A3466A32538BFFE" author="Song DX &amp; Chen ZQ &amp; Gong LS" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="424-427" refId="B6" refString="Song DX, Chen ZQ, Gong LS (1991) A new species of the genus Spartaeus from China (Araneae: Salticidae). Acta Zootaxonomica Sinica 16: 424-427." title="A new species of the genus Spartaeus from China (Araneae: Salticidae)." volume="16" year="1991">Song, Chen &amp; Gong, 1991</bibRefCitation>
</taxonomicName>
, the marked
<abbrev id="ABBRID0E5VAE" xlink_title="tegular apophysis">
<collectionCode id="02E0BC574C38B3C8130523E97B320C3D" country="0" name="Timescale Adventures Research and Interpretive Center">TA</collectionCode>
</abbrev>
(
<bibRefCitation id="B58C8D7A1233A63C304214E683CBECA5" DOI="10.11646/zootaxa.5389.1.1" author="Lin" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zootaxa" pagination="1-33" refId="B2" refString="Lin Y, Xin Y, Wang C, Li S (2023) On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae). Zootaxa 5389 (1): 133. https://doi.org/10.11646/zootaxa.5389.1.1" title="On ten jumping spider species described by Song and Chai (1992) based on type specimens (Araneae, Salticidae)." volume="5389" year="2023">Lin et al. 2023</bibRefCitation>
: fig. 18) is likely mislabelled and probably should be M in this study, considering its position at the base of embolus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,321 @@
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<mods:title id="4C56EABC8DC661B30E0ED53671CFDB6D">A taxonomic synopsis and identification key to the genera of the Myaptex group (Diptera, Asilidae, Asilinae), with description of a remarkable new genus and three new species from the South American Chaco</mods:title>
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<mods:namePart id="1D6DECD61F01562721871801AD802666">Soares, Matheus M. M.</mods:namePart>
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<mods:namePart id="FEF6BF3A6D0A4C76B7625EBEBB6BE154">Camargo, Alexssandro</mods:namePart>
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<mods:affiliation id="E8FC2A00DC2B55616F6C95156062713B">Natural History Museum Vienna, 2 nd Zoological Department. Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:namePart id="193995660E36D3323C6D8D8EE85DE7F2">Lamas, Carlos J. E.</mods:namePart>
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<subSubSection id="B7D99926DD5CD6533DF10BACF53B17E7" type="nomenclature">
<paragraph id="4CA1159D4AEEFEAF70E6064F789AA70C">
<taxonomicName id="FD2DEDD3C1D9628CE9C7AACBAEB9E22F" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="726696EBBD6094FB78A1CA42AA6299AC" italics="true">Cardiasilus dangeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF6E8A839AC78E6340B746E5DD5306CD" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="76F6CF2D28D32BCEA6670378E0047BF0" type="description">
<paragraph id="074970E5EC1153915336758D56315164">
<figureCitation id="28A11D3A921CD6E50BBF3202A7E7241E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">Figs 6</figureCitation>
,
<figureCitation id="D3BF335550B7985921A7F6F45678299F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7</figureCitation>
,
<figureCitation id="1ECAF846E3EB75D0D4EA923575B65988" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8</figureCitation>
,
<figureCitation id="CE665CE26641863E6450808D3D78A075" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6WAG" type="diagnosis">
<paragraph id="EEB31E342D78DE21C0277C67FD01D567">
<heading id="DFA437CF109B840031859288995E31F4" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B434C2C02B6786C6C934DF706F56B417">
The black postalar macrosetae (Fig.
<figureCitation id="6DFD047C69DD832CF3DAEF2FF0725F81" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 B</figureCitation>
), the epandrium strongly excavated at mid-inner dorsal margin (Fig.
<figureCitation id="F2B2EE81795FC49F93AB949FAE06320D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7 A, F, H</figureCitation>
), and the subepandrial sclerite somewhat trapezoidal at middle (Fig.
<figureCitation id="C44D707FF78B9E46185FE43D785EC3DC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8 D</figureCitation>
) should promptly distinguish this species from its congeners.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERXAG" type="description">
<paragraph id="3AFC2ED92D32F3E15A611AC0444B0F19">
<heading id="DD970E500FCD10ABBD145999DA89DF8C" reason="title">Description.</heading>
</paragraph>
<paragraph id="0555867F57A2DEBB12D94C0AEFC2848E">
<emphasis id="D5ADAF4B812C02EFFA052C0D9F8A9F87" bold="true">Male</emphasis>
(Fig.
<figureCitation id="63E1B28F5D1AFA041693B49B9943C2E5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 A</figureCitation>
). Body Length:
<quantity id="26D22FF53A8D104D5F1300645277B0F5" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" unit="mm" value="13.2">13.2 mm</quantity>
; wing length:
<quantity id="23DA0872FB1683D2FCAC0D4530EF3C53" metricMagnitude="-3" metricUnit="m" metricValue="7.1" unit="mm" value="7.1">7.1 mm</quantity>
.
<emphasis id="D4C1DCEC9038AB6E387C0D8F9487F147" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="AFA036C9FDF229786DE91755FD9F6513" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 A D</figureCitation>
). Similar to
<taxonomicName id="D58B5134C64C449596F99703F10C9F09" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="1B46F293E262A16580366D4FC688A6EF" italics="true">C. aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FD48B1AA03A465D7D063F115E9529074" rank="species">sp. nov.</taxonomicNameLabel>
except as noted:
<emphasis id="63C64B547ADED46A365F27259B89E394" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="A6AEA29738E57ECBDE90C64FD9D73B60" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 A, B, D</figureCitation>
) postalar macrosetae black.
<emphasis id="00A3FA374C1220692126463DC7194377" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="852F8CA6A6BF689BB369617FBF28E86F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 B</figureCitation>
).
<emphasis id="8E79B5BAFC7B22E7280F10FADA062802" bold="true">Leg I.</emphasis>
Tibia with posterior row of four long black macrosetae from basal 1 / 4 to apex.
<emphasis id="866F3052B729548CE0145FBC916CE796" bold="true">Leg II.</emphasis>
Femur with anterior row of four strong black macrosetae (basalmost white). Tibia with macrosetae wholly black.
<emphasis id="B671F0FB0566D2A178336EFCDC915323" bold="true">Leg III.</emphasis>
Femur with anteroventral row of short, strong, and sparse black macrosetae (basalmost white).
<emphasis id="973A6B0A9E528940D1301EEAE4AE6DFA" bold="true" italics="true">Terminalia</emphasis>
(Figs
<figureCitation id="DE2DD9DCB4BED895F76B94E0AFD4EAB2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7</figureCitation>
,
<figureCitation id="56FAAE2193E0C3BB06673A3EBB95C585" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8</figureCitation>
). Similar to
<taxonomicName id="6747A4A3939241E0F037327C2E9C6760" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="DE8F74A8F5014FADAB644ED3FAEC98D6" italics="true">C. aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BE6E97392646DDEB886D82A90BAF6B34" rank="species">sp. nov.</taxonomicNameLabel>
, except as noted: Epandrium strongly excavated at mid-inner dorsal margin (Fig.
<figureCitation id="17A75794F8D73C2C2929324B52A1B040" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7 A, F</figureCitation>
); subepandrial sclerite somewhat trapezoidal at its middle (Fig.
<figureCitation id="B8F22DB642E3D4E6C6A0515351C59ED3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8 D</figureCitation>
).
</paragraph>
<caption id="C907FAF29636EDFDEE62BBFF17BECF9B" ID-DOI="10.3897/zookeys.1232.142494.figure6" ID-arpha="4F939530-A7EB-53BF-9F1D-40B1B58B866A" httpUri="https://binary.pensoft.net/fig/1286740" startId="F6">
<paragraph id="277ACDFF57A70F5093E279B20CD1BDA5">
<label id="986439A776021347CFB953200AFEF16C">Figure 6.</label>
</paragraph>
<paragraph id="9001C237E480682A5659FED05FC117E0">
<taxonomicName id="75F85E5E15E8E938508B22A991E559D7" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="398AD4FDFC62C7AFD8BC83BB3E27D2CA" italics="true">Cardiasilus dangeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A098FF1361ABE63979888AE458B244AF" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype, before dissection)
<emphasis id="A97917A1A7F7D5008DFD83327FCB4CFA" bold="true">A, B</emphasis>
habitus lateral and dorsal, respectively
<emphasis id="CA56F4962BACC36CA247B5A8CFD709D8" bold="true">C</emphasis>
head, anterior view
<emphasis id="17F595B133FA7545F29E2D9C1CE5993F" bold="true">D</emphasis>
head and thorax, lateral view
<emphasis id="43A896070056BD214F3F6E9987EFD628" bold="true">E</emphasis>
abdomen, ventral view
<emphasis id="2C6EFF8122AC9FCC6A888BD6F654513A" bold="true">F</emphasis>
wing.
</paragraph>
</caption>
<caption id="1419901CF6BE452DD91724DEA33E50EE" ID-DOI="10.3897/zookeys.1232.142494.figure7" ID-arpha="8C9544D6-F669-5584-B4C2-6D1844DCCBC0" httpUri="https://binary.pensoft.net/fig/1286741" startId="F7">
<paragraph id="BCE2E0D9D19D05648F26D690E1F72BDC">
<label id="39D902242C58674CEE608A9DFAD6B215">Figure 7.</label>
</paragraph>
<paragraph id="1D205981C83A33BC1D54F1B02AA5FA51">
<taxonomicName id="6A02F27B658F55FA16DF1AF71313EE7B" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="FC7BBBD83E501CCB66056ACDCC953F9C" italics="true">Cardiasilus dangeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B74C615E93C98E470572F47B2994999A" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype)
<emphasis id="98AA0C095B4C6A78794D68F9B945C21F" bold="true">A C</emphasis>
terminalia in dorsal, ventral, and lateral views, respectively
<emphasis id="A72A01F5AEF2FC91DA339D2C26A8CD0A" bold="true">D</emphasis>
tergite 8
<emphasis id="5AD76199E477E1F5DD4B0BDD33245B28" bold="true">E</emphasis>
sternite 8
<emphasis id="2E70055C11EC5232D71B2ECB304AFCB1" bold="true">F, G</emphasis>
dissected terminalia in dorsal and ventral views, respectively
<emphasis id="FB871BE896FEF0A34437E81E660FA6DF" bold="true">H, I</emphasis>
dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium.
</paragraph>
</caption>
<caption id="E2E0482D0268710D54C42F23CF27CDA7" ID-DOI="10.3897/zookeys.1232.142494.figure8" ID-arpha="11B42DDA-D459-5F94-9A7C-98802C33D54B" httpUri="https://binary.pensoft.net/fig/1286742" startId="F8">
<paragraph id="0FCEBAEE66366418CF6327844969ADFB">
<label id="3FDC9851E33CBD53F399BEE8B4A55D1C">Figure 8.</label>
</paragraph>
<paragraph id="A467D76AD4B42353318C6992D2002452">
<taxonomicName id="68356C3B5B5EBB74FFE13C7206A74318" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="1BCC3398B1B7DEA01D614BBBDA5CC1E1" italics="true">Cardiasilus dangeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="164FDFFC4B2DC0C43A466ECD9DA1D531" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype)
<emphasis id="984DC7527E5AC464645723E38343EA30" bold="true">A</emphasis>
inner appendages
<emphasis id="648672B2E7AAA8474E2AD4B36A487266" bold="true">B</emphasis>
gonocoxite and gonostylus
<emphasis id="EFA67CBAF9F39A9843AE5464E144D9B8" bold="true">C</emphasis>
phallus and ejaculatory apodeme
<emphasis id="8346A4553B683B04D3FABD57D0131EB5" bold="true">D</emphasis>
subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus.
</paragraph>
</caption>
<paragraph id="2C434D6BF89FA71780F0662B0755D04F">
<emphasis id="98AF6B1F9E1A1290F55BA5D2740C32EE" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ4AG" type="type material">
<paragraph id="9B9481FB2D08A21E8C2A33D8F21C3B6C">
<heading id="5D596CE72BBB2CBA63741C8B3ADF982F" reason="title">Type examined material.</heading>
</paragraph>
<paragraph id="DE593636113A8918AA6D50DF10CD3823">
<materialsCitation id="70285208388D90E29508CFC375A0575B" collectingDate="2008-01-19" collectingDateMax="2008-01-31" collectingDateMin="2008-01-19" collectionCode="MZUSP, R" country="Brazil" latitude="-21.608334" location="Porto Murtinho" longLatPrecision="21" longitude="-57.817223" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="AE7A6F95C5E79854015D0F125C7B7B7D" bold="true" italics="true">
<typeStatus id="7B99A015931684B92BE5DF1B902E8E28" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="DC47F50C866464F835E9C5FEC47DF78A" count="1" type="male"></specimenCount>
(
<named-content id="582D9C71E71B7FA9E55A29FE075E9FE7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="69134CCEEE45E8B83E27A247E2F29C39" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
) labelled: “
<collectingCountry id="206535C448CDFD71BF02D977C4E2662D" name="Brazil">BRASIL</collectingCountry>
: MS [state of
<collectingRegion id="4F8E365E807008F6F03ED2A8377DEA6B" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
]:
<location id="729B09A6861E50826311FF3169539FA4" LSID="urn:lsid:plazi:treatment:10EDCD7218F854D9A3A0979C657815B3:729B09A6861E50826311FF3169539FA4" country="Brazil" latitude="-21.608334" longLatPrecision="21" longitude="-57.817223" name="Porto Murtinho" stateProvince="Mato Grosso do Sul">Porto Murtinho</location>
| Faz. São Fernando km 12 |
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E34AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.817222,-21.608333]}">
<geoCoordinate id="3959B3312AEC91AC886AEEF775DC2FD3" degrees="21" direction="south" minutes="36" orientation="latitude" precision="15" seconds="30" value="-21.608334">21 ° 36 ' 30 &quot; S</geoCoordinate>
,
<geoCoordinate id="64EB46D67A9FF9EAD201D41659499F86" degrees="57" direction="west" minutes="49" orientation="longitude" precision="15" seconds="02" value="-57.817223">57 ° 49 ' 02 &quot; W</geoCoordinate>
</named-content>
|
<date id="F514DB221FB387B191BFF70F4259573F" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">
<collectingDate id="674131E8582EBBE0B83B253B1DC48925" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">1931. I. 2008</collectingDate>
</date>
| Nihei, S., Figueiredo,
<collectionCode id="8AB7BF14B7961DEFF43CC3C498E95CA4" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. &amp; Almeida, J. | (col.) ” “
<typeStatus id="F4528A4C69AFE24ED1E5E6C99C091EBE" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="F017906F534BD4829A2B0D72AD7A49C3" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi">
<emphasis id="A029B2B6AEC697A8E7038A28C2023754" italics="true">Cardiasilus dangeloi</emphasis>
</taxonomicName>
| Soares, Camargo &amp; Lamas [red label] ”
</materialsCitation>
.
<typeStatus id="B0E8070418F93230DB23A8F59B84D2F2" type="holotype">Holotype</typeStatus>
condition: Good, left postpedicel broken off, abdomen glued to thorax with remains of glue obscuring pruinosity of tergites one to three, terminalia dissected and stored in microvial at the same pin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EM5AG" type="remarks">
<paragraph id="7181E5CC99267A1CB84CCCC74A675D8A">
<heading id="AF68E71F74BB5D2B22D3CDDFC9B3A294" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A83FEA741B89E87E0CA8BBA84D19B9B2">
The new species is remarkably similar to
<taxonomicName id="44EB2953B6883D65F3C78305E23538C4" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="35478D687FF4026370B9EE067CB1FCF5" italics="true">C. aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1762116D81F965B9199E0ED5F443DC8B" rank="species">sp. nov.</taxonomicNameLabel>
, differing only in the color of postalar macrosetae black (Fig.
<figureCitation id="88E1EAABBD0C5A2EFFA023210BD21B55" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 B</figureCitation>
), anterior row of macrosetae on mid femur wholly black, only the basalmost macroseta white (Fig.
<figureCitation id="AF29EBFCBD30A17E215E09FA7FEE1934" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 A</figureCitation>
), epandrium strongly excavated at mid-inner dorsal margin (Fig.
<figureCitation id="04E484F7711EB92D7930B81B02EDA9A4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7 A, F, H</figureCitation>
) and subepandrial sclerite somewhat trapezoidal at middle (Fig.
<figureCitation id="DE5F2C83290ED741F44E73CA441F2533" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8 D</figureCitation>
). In
<taxonomicName id="4DCF8C45E1CD9B96DA272BE527851688" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="724496F3EDAD7ED88B9985E934E18784" italics="true">C. aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4FCA5F471A0DE0168FA0C10DA63DF1D" rank="species">sp. nov.</taxonomicNameLabel>
the postalar macrosetae are yellow (Fig.
<figureCitation id="FD0CCC625676125F8E7E89F71F4757D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
), the anterior row of macrosetae on the mid femur is white, the mid-inner dorsal margin of the epandrium is weakly excavated (Fig.
<figureCitation id="C5552ABB18C268EE0E0D31D12691A47F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 A</figureCitation>
) and the subepandrial sclerite is somewhat diamond-shaped (Fig.
<figureCitation id="1100B0421BDDE58D506162E2341C354D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 D</figureCitation>
). The epandrium strongly excavated medially at inner margin and the lack of a dorsal preapical process also differs
<taxonomicName id="9DEE0E7EE496182B49BEDD1552DF287B" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="A57BA6DEE6A1296841C80AC4734A3A23" italics="true">C. dangeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E923E61BCB2F6E70FA24333BEC6E311C" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="2AF6829CC8BACBC498E48A69ED93E619" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda" status="sp. nov.">
<emphasis id="4E07258DF0599E068B2159F0A75E2ED1" italics="true">C. ruda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="23738B69F129B4789C837333798E8169" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1ABG" type="distribution">
<paragraph id="20048F8652F2BEA71EF513A9EB45814C">
<heading id="9B92FE1D3DC4C86FDEE67058CDDB603D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C1A47C4A8D5291E300231E3E2FBDE290">
<collectingCountry id="53C2CAA1F6910F56D95A51BB0DD83A56" name="Brazil">Brazil</collectingCountry>
(state of
<collectingRegion id="EF448337650E687E71C610AD765E7190" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
) (Fig.
<figureCitation id="A34BC17A601CC0CB08AF00F4F92014C9" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEBBG" type="etymology">
<paragraph id="89397905578EFBBC8864D1B9AA9F1444">
<heading id="AB955DDB9461E2CFF2A22F7798183439" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9FE568A4D0348C8ACF2686CDC9947D48">
The new species is named after Gio DAngelo (
<collectionCode id="4DEC320E3ADECA306267BCA18C166624" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" type="">INPA</collectionCode>
), an artist, photographer, and myrmecologist who is the partner of the first author and deeply passionate about scientific illustration. Gio also kindly created the illustration for Fig.
<figureCitation id="F423C35272E2D233C49057652FE94E36" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Illustration of Cardiasilus dangeloi gen. nov. et sp. nov." figureDoi="10.3897/zookeys.1232.142494.figure20" httpUri="https://binary.pensoft.net/fig/1286754">20</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,277 @@
<document id="F07F2336E2DD0863537F40784D8EE29B" ID-DOI="10.3897/zookeys.1232.146855" ID-ZooBank="5778FBF6-2A76-4E42-A5A7-FA79CBAF56B9" ID-publisher-id="146855" URI-arpha="D1264EEB-CEDE-596D-9907-97E1726CB3F0" URI-zoobank="http://zoobank.org/5778FBF6-2A76-4E42-A5A7-FA79CBAF56B9" article-type="research-article" checkinTime="1742495071518" checkinUser="pensoft" docAuthor="Ni, Yi, Yu, Kun &amp; Zhang, Junxia" docDate="2025" docId="123D83865A4652BDA03676D0A1F27C00" docLanguage="en" docName="ZooKeys 1232: 343-355" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/146855/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Calxattus dengba Ni, Yu &amp; Zhang, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="355" masterDocId="D1264EEBCEDE596D990797E1726CB3F0" masterDocTitle="Description of three new species of Spartaeini (Araneae, Salticidae, Spartaeinae) from China and Malaysia" masterLastPageNumber="355" masterPageNumber="343" pageNumber="343" updateTime="1742495462206" updateUser="ExternalLinkService">
<mods:mods id="B518F6A90F73D6F0878DA3BD0D17D46A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B70A9A224F74801B0B445FEF4E6C84F">
<mods:title id="57561DCDC26A259C186FB041CC78CDE9">Description of three new species of Spartaeini (Araneae, Salticidae, Spartaeinae) from China and Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="F902E49389448830EFA020F8DA849171" type="personal">
<mods:role id="02A4E9779D9F37206FB38CB1E6156002">
<mods:roleTerm id="632E4682D709AC05090BE5695809301B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4F013ECA0477A3ACF341AC7D5DFC603">Ni, Yi</mods:namePart>
<mods:nameIdentifier id="FC1C38724398CA6744657DD621FE9DDD" type="ORCID">https://orcid.org/0009-0003-3239-3697</mods:nameIdentifier>
<mods:affiliation id="E659BC0AC84110197F263313A8E3DFAB">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name id="D9EBE2EA95621D99664125DDE5DA438C" type="personal">
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<mods:roleTerm id="340D5AE474BF205DA5950519BCFF4508">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03B34B205D28AC92FBB3C12C51218189">Yu, Kun</mods:namePart>
<mods:nameIdentifier id="FF373BD2BF6D89CBC4DD834499AF71D1" type="ORCID">0000-0002-5517-4484</mods:nameIdentifier>
<mods:affiliation id="36BA5ED3F3080F70644E23A64F42415E">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name id="A4E4F795F3BE18BAEC082EA512EEDB8F" type="personal">
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<mods:roleTerm id="E3706585087E5FDC20424DB5345D537B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B50B8EDDF4966FC5F654BCDD6B97341">Zhang, Junxia</mods:namePart>
<mods:nameIdentifier id="23BE5978468B03757686751437054FA4" type="ORCID">0000-0003-2179-3954</mods:nameIdentifier>
<mods:affiliation id="4584BFDFCF240C884001C825AE406CF9">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
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<mods:date id="977EC27565CAD1F20F3D92BC14FA819B">2025</mods:date>
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<mods:number id="56A9642D521C91D74E2FC3EAA61DA1FE">2025-03-20</mods:number>
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<mods:identifier id="9EAAFA5FD98A28449361A40B46258E1A" type="DOI">10.3897/zookeys.1232.146855</mods:identifier>
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<treatment id="123D83865A4652BDA03676D0A1F27C00" ID-DOI="http://doi.org/10.5281/zenodo.15059384" ID-Zenodo-Dep="15059384" ID-arpha="123D8386-5A46-52BD-A036-76D0A1F27C00" ID-zoobank="https://zoobank.org/655EF590-7FF4-4ED5-AFEF-52392536DE9F" LSID="urn:lsid:plazi:treatment:123D83865A4652BDA03676D0A1F27C00" LSID-ZBK="urn:lsid:zoobank.org:act:655EF590-7FF4-4ED5-AFEF-52392536DE9F" httpUri="http://treatment.plazi.org/id/123D83865A4652BDA03676D0A1F27C00">
<subSubSection id="6E548E429EAA7D3DAB1921B84716908A" type="nomenclature">
<paragraph id="A9BAF98D08AB74D53B06FDE25D9FBE3D">
<taxonomicName id="4822BB6A8230FEDA385E8753BFF737C3" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Calxattus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="dengba" status="sp. nov.">
<emphasis id="2FBBC92DAF4BCF9ECCCF256AC8ECD600" italics="true">Calxattus dengba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="020A24ED86EF87706D1CDFE236F484AC" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B75AA074FDA16D3D69B7FDA2A88B300F" type="description">
<paragraph id="3FBF0943CB98E7B1DA6E52FAB954328F">
<figureCitation id="EEC785D724EF4655079E48F25B106108" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">Figs 1</figureCitation>
,
<figureCitation id="1AE3A5B1E7F595043E997A76BF04AFAE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Calxattus dengba sp. nov., palp of holotype male (A C); in prolateral (A), ventral (B) and retrolateral (C) view. Scale bars: 0.2 mm (A C)." figureDoi="10.3897/zookeys.1232.146855.figure2" httpUri="https://binary.pensoft.net/fig/1286791">2 Common name. 僜巴岩跳蛛</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERNAC" type="type material">
<paragraph id="0DEC3AB7C7E480BB6FEADE146F268130">
<heading id="A71A0ACFCA505A9D2E9B5461FDFB3BC3" reason="title">Type material.</heading>
</paragraph>
<paragraph id="D842ADC7156DA18E40F581E3435FED8D">
<materialsCitation id="064ABFA28F95FFC50B78AC117EA14D3A" collectingDate="2002-08-09" collectorName="M. Zhu" country="China" county="Zayu County" location="Nyingchi City" municipality="Autonomous Region" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="D603A706BBFF836F094177105052E3AA" bold="true" italics="true">
<typeStatus id="21FB66010623EE0844CAB32F8086CAD0" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="8F130BF2BDFC2CE489936FF55E1195E7" name="China">China</collectingCountry>
<specimenCount id="D5A180EF12F789AFAA5B60EE7885EC78" count="1" type="male"></specimenCount>
(MHBU-ARA- 00027415);
<collectingRegion id="DFD108EDF97325E7BF754E1023E090A5" country="China" name="Xizang">Xizang</collectingRegion>
<collectingMunicipality id="A820F0333A809CDBF91DC4098D082FF5">Autonomous Region</collectingMunicipality>
:
<location id="14AA0AE3D1D7250B3262BDAD2EABC3C3" LSID="urn:lsid:plazi:treatment:123D83865A4652BDA03676D0A1F27C00:14AA0AE3D1D7250B3262BDAD2EABC3C3" country="China" county="Zayu County" municipality="Autonomous Region" name="Nyingchi City" stateProvince="Xizang">Nyingchi City</location>
,
<collectingCounty id="6085A8541C008137DCA40CA040EA6862">Zayu County</collectingCounty>
,
<location id="23EB032A78BF99400BDBB30F97104D30" LSID="urn:lsid:plazi:treatment:123D83865A4652BDA03676D0A1F27C00:23EB032A78BF99400BDBB30F97104D30" country="China" county="Zayu County" municipality="Autonomous Region" name="Xia Zayu Town" stateProvince="Xizang">Xia Zayu Town</location>
;
<date id="26F3F5B325BFD5438B5E9DCDE23CD737" value="2002-08-09">
<collectingDate id="86490FC625B26EFFEDA60A22166DA335" value="2002-08-09">9. VIII. 2002</collectingDate>
</date>
;
<collectorName id="6CFFB3D422A570018160C5DA6CDD0C46">M. Zhu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3NAC" type="etymology">
<paragraph id="A80BC6E0CC32F272D33DCF8A31E5DB1B">
<heading id="6432D33081B62965BE40CC81A868C3ED" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="D3DA25F613EC1DF3F167B26401F68617">
The specific epithet is derived from the Dengba people, who live in Zayu County, the
<typeStatus id="6EADAB27F28E33412980AADC67186C42">type</typeStatus>
locality of the new species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBOAC" type="diagnosis">
<paragraph id="2D5DA0B5EAC1F7DC098FBC4D2A752C3D">
<heading id="74DCFDE4836F10FC2C288F21B2F4D2E1" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F1A98794B79B3E3D705FE59954D9D6C2">
The male of the new species can be distinguished from its sole congeneric species,
<taxonomicName id="5B6A1DDFE8EB4857FD9BDFAEA048D79C" authorityName="Yang, Liu, Liu &amp; Peng" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="49ADDF0C8F003DB8E9A32855F483DF17" italics="true">C. serratus</emphasis>
</taxonomicName>
, by the following: (1) the
<abbrev id="ABBRID0ESOAC" xlink_title="retrolateral tibial apophysis">
<collectionCode id="7A6A44F282CC1D6EBF66BD674EE07B8E">RTA</collectionCode>
</abbrev>
lacks processes on its prolateral and retrolateral edges (Figs
<figureCitation id="8723A7C9FB221E81DE46161CF37B3DD6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 E</figureCitation>
,
<figureCitation id="6C1B753ABAA93C844B7F5AC5ECF143EA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Calxattus dengba sp. nov., palp of holotype male (A C); in prolateral (A), ventral (B) and retrolateral (C) view. Scale bars: 0.2 mm (A C)." figureDoi="10.3897/zookeys.1232.146855.figure2" httpUri="https://binary.pensoft.net/fig/1286791">2 B</figureCitation>
; vs. in
<taxonomicName id="4DB31035FFCCD28C95ED5BF6870340E3" authorityName="Yang, Liu, Liu &amp; Peng" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="BEC0A56530D3BE2BCD58DF91C88FEDD3" italics="true">C. serratus</emphasis>
</taxonomicName>
, the
<abbrev id="ABBRID0EJPAC" xlink_title="retrolateral tibial apophysis">
<collectionCode id="051947781DA30FBD290E52BAE6B8C919">RTA</collectionCode>
</abbrev>
carries median processes on both prolateral and retrolateral edges; see
<bibRefCitation id="BD3B40BC6E07CA90DAC2FFCE2C175EA9" DOI="10.3969/j.issn.1005-9628.2023.01.002" author="Wang" etAl="et al." firstAuthor="Wang" issue="1" journalOrPublisher="Acta Arachnologica Sinica" pagination="8-15" refId="B9" refString="Wang WH, Yu K, Zhang JX (2023) Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species. Acta Arachnologica Sinica 32 (1): 815. https://doi.org/10.3969/j.issn.1005-9628.2023.01.002" title="Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species." volume="32" year="2023">Wang et al. 2023</bibRefCitation>
: figs 5 B, 7 B); (2) the embolus is relatively short, originates from the median portion of the palpal bulb and terminates near the midpoint of the lower edge of the groove-shaped embolic guide (Figs
<figureCitation id="7129DDEC741F704A2C70469C4D6C098F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 E</figureCitation>
,
<figureCitation id="9E19E19D3F2188F1B195F766AD793DA0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Calxattus dengba sp. nov., palp of holotype male (A C); in prolateral (A), ventral (B) and retrolateral (C) view. Scale bars: 0.2 mm (A C)." figureDoi="10.3897/zookeys.1232.146855.figure2" httpUri="https://binary.pensoft.net/fig/1286791">2 B</figureCitation>
; vs. in
<taxonomicName id="6CABD92C69D29C77D98903AB051E1482" authorityName="Yang, Liu, Liu &amp; Peng" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5A757D67C6808EBE8FD5FD5F6877A8F7" italics="true">C. serratus</emphasis>
</taxonomicName>
, the embolus is long, originates from the lower portion of the bulb and terminates through the groove-shaped embolic guide near the functional conductor; see
<bibRefCitation id="E41B416A30B68F8B003BE413A426F75B" DOI="10.3969/j.issn.1005-9628.2023.01.002" author="Wang" etAl="et al." firstAuthor="Wang" issue="1" journalOrPublisher="Acta Arachnologica Sinica" pagination="8-15" refId="B9" refString="Wang WH, Yu K, Zhang JX (2023) Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species. Acta Arachnologica Sinica 32 (1): 815. https://doi.org/10.3969/j.issn.1005-9628.2023.01.002" title="Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species." volume="32" year="2023">Wang et al. 2023</bibRefCitation>
: figs 5 B, 7 B); and (3) the median apophysis is located in the upper half of the retrolateral part of the bulb (Figs
<figureCitation id="D1BFB1A4DA195C094375DAF3488B9E7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 E</figureCitation>
,
<figureCitation id="67E209A011C6ED30B4AAE641BE918265" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Calxattus dengba sp. nov., palp of holotype male (A C); in prolateral (A), ventral (B) and retrolateral (C) view. Scale bars: 0.2 mm (A C)." figureDoi="10.3897/zookeys.1232.146855.figure2" httpUri="https://binary.pensoft.net/fig/1286791">2 B</figureCitation>
; vs. in
<taxonomicName id="AC1E1CEED6A8ACF7E2DDD6F7DEA70571" authorityName="Yang, Liu, Liu &amp; Peng" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Spartaeus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CEF06E71D1CE1FDDA99262C489B6B0B6" italics="true">C. serratus</emphasis>
</taxonomicName>
, the median apophysis is located in the lower half of the retrolateral part of the bulb; see
<bibRefCitation id="092513AAEDB9CE1EC6DA8857EDBA7E89" DOI="10.3969/j.issn.1005-9628.2023.01.002" author="Wang" etAl="et al." firstAuthor="Wang" issue="1" journalOrPublisher="Acta Arachnologica Sinica" pagination="8-15" refId="B9" refString="Wang WH, Yu K, Zhang JX (2023) Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species. Acta Arachnologica Sinica 32 (1): 815. https://doi.org/10.3969/j.issn.1005-9628.2023.01.002" title="Calxattus gen. nov. from China (Araneae: Salticidae: Spartaeinae: Spartaeini), with redescription of the type species." volume="32" year="2023">Wang et al. 2023</bibRefCitation>
: figs 5 B, 7 B).
</paragraph>
<caption id="045FDA44C966FE5067CCE65CAF59247B" ID-DOI="10.3897/zookeys.1232.146855.figure1" ID-arpha="6A3A6BA1-4ADC-5E56-935E-80D98BFB7C96" httpUri="https://binary.pensoft.net/fig/1286790" startId="F1">
<paragraph id="269740502E61B74B94B6531ABDDE935D">
<label id="608FE2E9C43100400629D96F03151297">Figure 1.</label>
</paragraph>
<paragraph id="5ACB3DA0267D480DB8414F583C09B2A3">
<taxonomicName id="653D3A5D9E7527AD9F7C7897DF392644" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Calxattus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="dengba" status="sp. nov.">
<emphasis id="F4F329D0064FDB220490DD1D5DFEFA83" italics="true">Calxattus dengba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8129C31AAA1C7522F86B5D91E7F37163" rank="species">sp. nov.</taxonomicNameLabel>
, habitus of holotype male (
<emphasis id="87500E58A82A279BC74A872BB0F5BD77" bold="true">A, B</emphasis>
), cheliceral teeth (
<emphasis id="6EF1DBA760D35B901FAD120F05FC6ECD" bold="true">C</emphasis>
), palp of holotype male (
<emphasis id="9B0C7E989EFC3E5C5E76D893689E30C0" bold="true">D F</emphasis>
); in dorsal (
<emphasis id="80EA5FA6BB2FC0331E2B60F59E793755" bold="true">A</emphasis>
), front (
<emphasis id="FAD84BFAB1D7BCE0EBA0D82081558DC4" bold="true">B</emphasis>
), prolateral (
<emphasis id="804CFDDD70326B8847FE3B9ACB943BF9" bold="true">D</emphasis>
), ventral (
<emphasis id="F6C49BDC7CEFB5126C12C43648850F12" bold="true">E</emphasis>
), retrolateral (
<emphasis id="2529A6CE992B12856ABA7AEA44D122B5" bold="true">F</emphasis>
) and back (
<emphasis id="CE6F9FBEBD588BB1E71B6621556AD98B" bold="true">C</emphasis>
) view. Scale bars: 1 mm (
<emphasis id="A644AEE89A455931F66CDFCCD096EC3A" bold="true">A, B</emphasis>
); 0.2 mm (
<emphasis id="F31188438C7A605B259E38945DFE0EEE" bold="true">C F</emphasis>
).
</paragraph>
</caption>
<caption id="B85AAAE1EDD537335BEF2B0DD1C3348C" ID-DOI="10.3897/zookeys.1232.146855.figure2" ID-arpha="A8B994C7-640D-54C5-B3EF-2314A816BEEC" httpUri="https://binary.pensoft.net/fig/1286791" startId="F2">
<paragraph id="1DB380294BB8F0B05E6D7EDC64D0FC97">
<label id="12C7F19E0166AED3357A35F510ABF85A">Figure 2.</label>
</paragraph>
<paragraph id="F46B9DFC0D07FFB179A9241DD17BE985">
<taxonomicName id="1D2DD7F192D4C3508993F0894B2CB31E" authorityName="Ni &amp; Yu &amp; Zhang" authorityYear="2025" class="Arachnida" family="Salticidae" genus="Calxattus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="dengba" status="sp. nov.">
<emphasis id="D1080A1515F4BA0461152E926CB94AB0" italics="true">Calxattus dengba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7D2A6F0EB6469B0B7115BB384C40B38" rank="species">sp. nov.</taxonomicNameLabel>
, palp of holotype male (
<emphasis id="9FE7C118A09F51CC32A2CD6823B4753E" bold="true">A C</emphasis>
); in prolateral (
<emphasis id="1CB1CB4A0F12D712A4B0A70C331C4FE5" bold="true">A</emphasis>
), ventral (
<emphasis id="FFA358B7A0F510F78B99C3B5713B5909" bold="true">B</emphasis>
) and retrolateral (
<emphasis id="8E3B18BDFAA17C389F53FF51034CA6A0" bold="true">C</emphasis>
) view. Scale bars: 0.2 mm (
<emphasis id="7797EA3148AF892D0DF46651A182A479" bold="true">A C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQDAE" type="description">
<paragraph id="A6ECC76CEC20B74021FD39A42DE501A6">
<heading id="A7869A708DE98A2A281F104ED014A3DC" reason="title">Description.</heading>
</paragraph>
<paragraph id="90E649E56AEE815F7AB6CA04EAC0FA83">
<emphasis id="FE6549E7D6EE202AC3CF16471376EDE9" bold="true">Male.</emphasis>
Measurements of
<typeStatus id="9D40EDF5FC9C20875E1DF2BAC5098C57" type="holotype">holotype</typeStatus>
: total length 7.26. Carapace 3.52 long, 2.47 wide. Abdomen 3.34 long, 2.28 wide. Eye measurements:
<abbrev id="ABBRID0EYDAE" xlink_title="anterior median eye">
<collectionCode id="A22ADF6BE1C4D2B8E94BE0B1AC402E4D">AME</collectionCode>
</abbrev>
0.63,
<abbrev id="ABBRID0E3DAE" xlink_title="anterior lateral eye">
<collectionCode id="AE96FA4E14C386EC2DBB70CC3EF20B0C">ALE</collectionCode>
</abbrev>
0.26,
<abbrev id="ABBRID0EAEAE" xlink_title="posterior median eye">
<collectionCode id="943078905A54CE2289F1B4B1AD0CDB4E">PME</collectionCode>
</abbrev>
0.43,
<abbrev id="ABBRID0EEEAE" xlink_title="posterior lateral eye">
<collectionCode id="4AE747358B17CE3AF2C34BFF0D64A4B9">PLE</collectionCode>
</abbrev>
0.37. Leg measurements: I 14.18 (3.78, 1.35, 4.47, 2.91, 1.67), II 7.89 (2.80, 0.96, 2.28, 1.35, 0.50), III 7.74 (2.41, 0.87, 1.86, 1.84, 0.76), IV 11.39 (3.07, 0.83, 3.04, 3.27, 1.18); leg formula 1423. Chelicera (Fig.
<figureCitation id="704A6DBA3E0A6E58B7E3063AEDE05095" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 C</figureCitation>
) brown, with six promarginal and seven retromarginal teeth. Carapace (Fig.
<figureCitation id="887D7F7997F179F5ED546FF70B035A81" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 A, B</figureCitation>
) reddish-brown, eye surroundings black, clypeus and eye surroundings covered by sparse white setae, fovea dark and longitudinal slit-like. Endites and labium yellowish-brown, with brown setae at distal end; sternum flat and creamy yellow, covered with white and brown setae. Legs (Fig.
<figureCitation id="DA45108A5C8ECD0BCE01B97776FA884E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 A</figureCitation>
) dark yellow, bearing long and thin spines; leg I extra long. Abdomen (Fig.
<figureCitation id="4B3AC57A8FC3A6C3EEF55F7926EBCACE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 A</figureCitation>
) pale yellow and elongated oval, surface faded; dorsal side with visible grey bands at edge, ending near spinnerets; short yellow setae covered on dorsal lateral sides of anterior half; ventral side with fine or flattened setae in middle part. Spinnerets (Fig.
<figureCitation id="2B02590B4423CA1F0931C2F171A73E54" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Calxattus dengba sp. nov., habitus of holotype male (A, B), cheliceral teeth (C), palp of holotype male (D F); in dorsal (A), front (B), prolateral (D), ventral (E), retrolateral (F) and back (C) view. Scale bars: 1 mm (A, B); 0.2 mm (C F)." figureDoi="10.3897/zookeys.1232.146855.figure1" httpUri="https://binary.pensoft.net/fig/1286790">1 A</figureCitation>
) yellow, covered with white setae.
</paragraph>
<paragraph id="3929F2BC2EA425A353CF18BB6C7AB78B">
Palp (Figs
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,
<figureCitation id="B44E05466FDE79537BD1B57DAF9B449A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Calxattus dengba sp. nov., palp of holotype male (A C); in prolateral (A), ventral (B) and retrolateral (C) view. Scale bars: 0.2 mm (A C)." figureDoi="10.3897/zookeys.1232.146855.figure2" httpUri="https://binary.pensoft.net/fig/1286791">2 A C</figureCitation>
): embolus thread-like, terminating near midpoint of embolic guide in ventral view; median apophysis black, hook-like in ventral view, located in upper half of retrolateral part of bulb, near functional conductor; functional conductor fan-shaped, with plow-like furrows on surface; membranous tegular apophysis transparent, curved in middle portion, originating from tegular apex and covering half of basal embolus; retrolateral and ventral tibial apophyses finger-like in ventral view, surface of retrolateral tibial apophysis with fingerprint-like patterns.
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<paragraph id="78444E3E387F6A3CEF782A3EE1EEC659">
<emphasis id="5B2BA480543241960F93AA576F7F8701" bold="true">Female.</emphasis>
Unknown.
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<paragraph id="E57E38C68C590FA9DFDED67A3B7C4E5F">
<heading id="07D4DEB4535C653655EF4589D21815C4" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="70935F612918CA5FAEDB526FB4AAC628">
<collectingCountry id="16C7D916436E74172BE3075F2E5C59A1" name="China">China</collectingCountry>
(
<collectingRegion id="344A37AFD4C36F8663530C7898F09122" country="China" name="Xizang">Xizang</collectingRegion>
).
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</subSubSection>
</treatment>
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<emphasis id="EE0B52EEB9C8EB9436997B8DB87BACE2" italics="true">Calxattus</emphasis>
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<mods:identifier id="3FF8CB6023C32074EA3E41E6F441599E" type="DOI">10.3897/zookeys.1232.142494</mods:identifier>
<mods:identifier id="31003FCAA5CC79F987B4F59269B6979E" type="ZooBank">67DBBE51-5990-4842-AF0F-141CB0AFA4D2</mods:identifier>
</mods:mods>
<treatment id="3FC0DF610E4E555989ACAE111A6F0E03" ID-arpha="3FC0DF61-0E4E-5559-89AC-AE111A6F0E03" ID-zoobank="https://zoobank.org/CC0B7CD3-B621-45A9-8B0A-1852110EE91C" LSID="urn:lsid:plazi:treatment:3FC0DF610E4E555989ACAE111A6F0E03" LSID-ZBK="urn:lsid:zoobank.org:act:CC0B7CD3-B621-45A9-8B0A-1852110EE91C" httpUri="http://treatment.plazi.org/id/3FC0DF610E4E555989ACAE111A6F0E03">
<subSubSection id="2FC91115582F0961345184E8A9A151D5" type="nomenclature">
<paragraph id="697BB665A25064574673531FB66301A1">
<taxonomicName id="011BB2A897AD05F0310A192BA136566E" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="2525C7FB68059ECD6709B760BA28FEB7" italics="true">Cardiasilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8768A33404420C4B3B1D4A1FDD46BE46" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2CD214E329C02702E2234857256B8F55" type="description">
<paragraph id="84015E6CCC15D7C517E34263C9EA1D96">
<figureCitation id="FC817C8B3C6A52C2FF38F439F5E55E7B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">Figs 2</figureCitation>
,
<figureCitation id="F53F456401FDF1E88DC08FA2BC830BCF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3</figureCitation>
,
<figureCitation id="7863B4D42721F61B81AA32C3DCFF1C9D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4</figureCitation>
,
<figureCitation id="2C33CBA8F4E3420745A5F558746CA153" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cardiasilus aysu sp. nov. (identified female) A, B habitus lateral and dorsal, respectively C E terminalia in dorsal, ventral, and lateral views, respectively F spermathecae. Abbreviations: cerc = cercus; hyprct = hypoproct; lat sc = lateral sclerite of genital fork; st = sternite; tg = tergite." figureDoi="10.3897/zookeys.1232.142494.figure5" httpUri="https://binary.pensoft.net/fig/1286739">5</figureCitation>
,
<figureCitation id="ABE15E10BACB0E50766D3DCAC6AD567D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6</figureCitation>
,
<figureCitation id="FACA558E66A03D082A8F9B5F705E0EBE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7</figureCitation>
,
<figureCitation id="47FABD14570AFCCADBB8A9EFEB2D14E5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cardiasilus dangeloi sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure8" httpUri="https://binary.pensoft.net/fig/1286742">8</figureCitation>
,
<figureCitation id="900DA11A13E9BA11F8E05105EE01FF87" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">9</figureCitation>
,
<figureCitation id="07C26F12A1BDDC680054399812F8A65F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10</figureCitation>
,
<figureCitation id="1BD5B792746BC1865CFC7897B415AD45" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cardiasilus ruda sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure11" httpUri="https://binary.pensoft.net/fig/1286745">11</figureCitation>
,
<figureCitation id="005DC49EF7D73D721B9E37FA417DAE51" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDVAE" type="type species">
<paragraph id="0D5A3218B83F678EEF39996913E6F965">
<heading id="4F6C4CF3E96B0821775F829E421803EC" reason="title">
<typeStatus id="95265BD60EEB908339EE46106E6A802A">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="E6440E283CA48467963A6A0236FADD94">
<taxonomicName id="A4262F846694B31DAAB32BC97AC6572A" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="9ABB177C90D6093AF2B9DE8C48235913" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E52241F548FBEEFD24015EBAE514024" rank="species">sp. nov.</taxonomicNameLabel>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUVAE" type="etymology">
<paragraph id="0E2E40F7FCDF014BDED406FD2F05117C">
<heading id="69E4C23DF083F1EDDF2E1839490CF393" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="48D8557670A1FD24513D6DD53D544AC9">
From the Greek feminine word
<emphasis id="49D29AA8D0857F069DDCF8875B2381B1" italics="true">kardia</emphasis>
= heart +
<emphasis id="0D52F36B0672C924EDB3F44D35C2E24B" italics="true">asilus</emphasis>
= common epithet of robber flies. The name refers to the distinct inflated epandria, somewhat heart ideogram-shaped in dorsal view. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5VAE" type="diagnosis">
<paragraph id="1EFAB5647E9CD41A0B7564FDA4238182">
<heading id="2D77AC277BB02A2AD10EABC61A710D6B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="86018A085AD1501006CEEF8DB7BA4C0A">
<emphasis id="A25D7BF386606A8EC965220DB62483D6" bold="true">
<emphasis id="00485B2247FCE85A8C9F6260DBFF2C25" bold="true" italics="true">Head</emphasis>
.
</emphasis>
Antenna ~ 3 / 4 as long as eye height (Fig.
<figureCitation id="B5A0038056E76918314D1A7A1F7B8106" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A</figureCitation>
). Scape and pedicel subequally long (Fig.
<figureCitation id="3245A8087151124216649C5A848C13BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 E</figureCitation>
). Postpedicel lanceolate, laterally compressed and slightly tapering towards apex, about as long as scape and pedicel combined (Fig.
<figureCitation id="D76D4C4C8A578C2FF48DF9E57ABE5F84" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 E</figureCitation>
). Stylus slightly longer than postpedicel, composed of two elements (Fig.
<figureCitation id="CA3D9FAC1623DEDC471FBFEB4DFBF912" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 E</figureCitation>
). Face wide, slightly narrowing at antennal level and slightly gibbous at lower 1 / 3, mystax restricted to gibbosity (Fig.
<figureCitation id="5098FC9845239F5699EA11DEAB15A9E0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 D</figureCitation>
). Frons with parallel slopes, slightly concave at antennal level, twice wider than higher (Fig.
<figureCitation id="DE3043D6F47C7F6C305BA13BE3D72CF3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 D</figureCitation>
). Palpus one-segmented, short, ~ 1 / 5 length of proboscis. Proboscis ~ 3.5 / 5 as long as eye height (Fig.
<figureCitation id="6F60EE11AB94EC4131F6982214902BF0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
).
<emphasis id="DFD589888637AC48E7FCD286599D4E43" bold="true" italics="true">Thorax</emphasis>
. Acrostichal setae indistinct (Fig.
<figureCitation id="C3698FD225575596F5F9AF285464A7B9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
). Anterior dorsocentral macrosetae absent, scutum with three to four pairs of posterior dorsocentral macrosetae (Fig.
<figureCitation id="5299CCF0F4BF03CCB680D75D5FFB3F26" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
). Scutellum tumid with pair of marginal macrosetae (Fig.
<figureCitation id="F5F3ABAFDA43BA4EB9AECFCC0F66A765" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 B</figureCitation>
). Anatergite bare. Postmetacoxal area membranous.
<emphasis id="43A85A7F6CE126160F1B7FF28639F4B4" bold="true" italics="true">Legs.</emphasis>
Femora swollen.
<emphasis id="62C137E9372A15CEAC0A461580ED554B" bold="true" italics="true">Wing.</emphasis>
Distinctly shorter than abdomen, with bifurcation of veins
<emphasis id="56BD4AEE8BCDD7B454EF84ABE065DB43" italics="true">
<collectionCode id="9DC50229BF672A1102C413268E4EDB57" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="A9F2206723B2BEF163B249F2889F34C3">4</subScript>
</emphasis>
and
<emphasis id="88B1DCD1C83D38E48632395EDBFED597" italics="true">
<collectionCode id="D339BAA7853C806F8E92472EE28511D0" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="70D5421E3825EC3FC2A2AAA6E18B428F">5</subScript>
</emphasis>
placed before apex of discal cell by approx. length of
<emphasis id="57932CB4AF929A3046100ACFC19CE887" italics="true">r-m</emphasis>
cross vein (Fig.
<figureCitation id="9C965D9D2B12FAA041DB4A55214B5CE8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 G</figureCitation>
). Distance between apex of veins
<emphasis id="5813624B0FEF764DFB25692935E14BC0" italics="true">
<collectionCode id="6CF1BC3EC4D6F7574281C5E404AB4915" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="11F6D08A5FF87BB169D1A37E3CB8DC4C">4</subScript>
</emphasis>
and
<emphasis id="19EB3C5DC48B0812B8CD5B326B1845E6" italics="true">
<collectionCode id="D0E81B5E90238A031EC5094F446DC4EA" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="ED8202E3D501406A4D8425C246B3D7F5">5</subScript>
</emphasis>
~ 1.52 × longer than distance between apex of veins
<emphasis id="1163948F626D0A3918F4B1BF5F299E00" italics="true">
<collectionCode id="AC5221CC179FF50761E9828BB2B89000" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="F5BD1E6A21C6CAA8F4929369FCAA4E35">5</subScript>
</emphasis>
and
<emphasis id="A419739A04155DA5E8F0C83E3FB9FB63" italics="true">
M
<subScript id="9B2AFE58336B9EBF5A2ABAD14837D626">1</subScript>
</emphasis>
(Fig.
<figureCitation id="72A9381001E2BC7733C051741ED009B6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 G</figureCitation>
). Cells
<emphasis id="B5495955E0EA230BBA0C1CE1AC78CBCC" italics="true">
m
<subScript id="61B091E2FEB60FDF9F1E01DBDC1E7E93">3</subScript>
</emphasis>
and
<emphasis id="40B6D79878C79A9B623A104115201560" italics="true">cua</emphasis>
closed and petiolate before wing margin (Fig.
<figureCitation id="AB0AD769F4C5BF31689974A58B744D91" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 G</figureCitation>
).
<emphasis id="32FF4028968E88A9549E415A06996E8B" bold="true" italics="true">Abdomen</emphasis>
. Abdominal sternites 26 with two to three pairs of pale yellow macrosetae mid-laterally (Figs
<figureCitation id="B2A624BB2B641F0428A6CEFB31600902" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 H</figureCitation>
,
<figureCitation id="D036A46CC2C04D5C06B03D1DEDB87B88" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cardiasilus dangeloi sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure6" httpUri="https://binary.pensoft.net/fig/1286740">6 E</figureCitation>
,
<figureCitation id="63F655F701CB1BF4B4426A659CF434A8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">9 E</figureCitation>
). Sternite 8 with mid-posterior digitiform projection almost as long as sternite 8 length (Figs
<figureCitation id="7613F6C66506EB91158DDE04BE470B94" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 E</figureCitation>
,
<figureCitation id="B678C033793FEA5A928EA20B889D165C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cardiasilus dangeloi sp. nov. (male holotype) A C terminalia in dorsal, ventral, and lateral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure7" httpUri="https://binary.pensoft.net/fig/1286741">7 E</figureCitation>
,
<figureCitation id="342AC3FE5346E3831A7137EAB474DFF6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10 E</figureCitation>
).
<emphasis id="6B1847B948491FA249CBBD1A99790256" bold="true" italics="true">Terminalia</emphasis>
. Epandria inflated laterally and posteriorly, resembling the ideogram of heart in dorsal view. Phallus long and thin, longer than length of hypandrium plus gonocoxite, divided into two prongs along its entire length (i. e., Fig.
<figureCitation id="77B87F27CF0F0560140E122B512F12DB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 A C, F I</figureCitation>
).
</paragraph>
<caption id="474DCA84E0BC6185064E39674A5B070B" ID-DOI="10.3897/zookeys.1232.142494.figure2" ID-arpha="6D757D7A-B81D-5396-80E6-6CA382B23FA8" httpUri="https://binary.pensoft.net/fig/1286736" startId="F2">
<paragraph id="36C9D424B1625BD81D0362F41728BA45">
<label id="F703160CD3074C3D7D4A83258EC26ABC">Figure 2.</label>
</paragraph>
<paragraph id="AFC6F19DB02AE14226020A32BF0E8569">
<taxonomicName id="6238DB0077031A5B66408F76B663157F" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="F084F12C144744E3B615F8B2D307B14F" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C48EA64C1B6D9FA8F5D96DA2F253F1D6" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype)
<emphasis id="F31C3CBC612C59FA4FFE256EBBA6B366" bold="true">A, B</emphasis>
habitus lateral and dorsal, respectively
<emphasis id="76BB1C0D8ADD928EBC99449C7065E6A9" bold="true">C</emphasis>
head and thorax, lateral view
<emphasis id="7BCD6ED638197E87B0CD32CC62FC7B76" bold="true">D</emphasis>
head, anterior view
<emphasis id="9F36F5B7D18834400CD6FB8CCDD86995" bold="true">E, F</emphasis>
antenna, lateral and dorsal views, respectively
<emphasis id="CABB01DB43E4FA384BF9B6F38FBABE38" bold="true">G</emphasis>
wing
<emphasis id="DF4D2CD5671EA76C4E926B0CCC135976" bold="true">H</emphasis>
abdomen, ventral view. Abbreviations:
<emphasis id="AF7E0F46B8E2D1E959A1ABCF8705AA18" italics="true">d</emphasis>
= discal cell;
<emphasis id="7127BD618FED2FAFA005CE1C06B84635" italics="true">m 3</emphasis>
= third medial cell;
<emphasis id="123849275E5A9A8618BF1E9FA7A74B41" italics="true">r 1</emphasis>
= first radial cell;
<emphasis id="B0890670EC3BF3D76CC4E5E5EACCAB5D" italics="true">r 2 + 3</emphasis>
= second + third radial cell;
<emphasis id="AD088EB6F8E32D9C9360C3CF3E02DBC5" italics="true">R 2 + 3</emphasis>
= second branch of radius;
<emphasis id="1E0425E2C24003A2BCFF07B77F941998" italics="true">R 4</emphasis>
= upper branch of third branch of radius;
<emphasis id="FAFC35A2C0110C364BF31640F70B2330" italics="true">R 5</emphasis>
= lower branch of third branch of radius.
</paragraph>
</caption>
<caption id="41BC4386A32990C8C8CE4E54F6B9D823" ID-DOI="10.3897/zookeys.1232.142494.figure3" ID-arpha="446AC9E5-E159-5984-9DFF-A44FF6116E88" httpUri="https://binary.pensoft.net/fig/1286737" startId="F3">
<paragraph id="2C90B67BFC91BADC4500B8F245DCE935">
<label id="C201ED7AD363B58AC4D1DF93D210E3D7">Figure 3.</label>
</paragraph>
<paragraph id="871FFF0766FB15A8AA42818DF4DE02D5">
<taxonomicName id="C2C425CAE82BC7A12B6C71FA5C12C477" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="F6064C23DA16D24E365A0064E52BB021" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="514AD06EFE6FE12859403606931083CA" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="CB24C4AFBF04391B673CB26C2A09A29B" bold="true">A C</emphasis>
male holotype
<emphasis id="0E6EB3AB0ED70E796D21D73AA0D28050" bold="true">D I</emphasis>
male paratype)
<emphasis id="A4ED57BCCEDF1954D1542C0BA62D5733" bold="true">A C</emphasis>
terminalia in dorsal, lateral, and ventral views, respectively
<emphasis id="857160DF34FE8984D381C670A54C82AB" bold="true">D</emphasis>
tergite 8
<emphasis id="80134EE8EC3E2950A4833FA14725B3A7" bold="true">E</emphasis>
sternite 8
<emphasis id="9473E0416BA7D755AC2C765BAFDCFF02" bold="true">F, G</emphasis>
dissected terminalia in dorsal and ventral views, respectively
<emphasis id="2C2B7B9CA91B3E665F040D90394DA276" bold="true">H, I</emphasis>
dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EA4AE" type="remarks">
<paragraph id="52F5D76F182E72C5E991FE3BE9D2E1C5">
<heading id="4C2B839E351E4FD2FED6FEFC07DAEC8B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="756CD5878A051DC1957F7092A5950D23">
<taxonomicName id="FE05CD971B0EA772D8B9F8763380764E" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1C529055F7CDC59E4B6451FAED7D5D02" italics="true">Cardiasilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE4BDF916AE80DD72A0A5C4FC2976E02" rank="genus">gen. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="79583F3954916EF74ECE6C4803AC5FF3" authority="Hull" authorityName="Hull" authorityYear="1962" class="Insecta" family="Asilidae" genus="Myaptex" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="FEFC033A0C8856CC77DD0D2A105F4861" italics="true">Myaptex</emphasis>
Hull
</taxonomicName>
by the inflated male epandria (Fig.
<figureCitation id="E343FBB493F4041BCE77A7F31FBEAA82" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B Terminalia in dorsal and ventral views, respectively C, D terminalia without inner appendages in dorsal and ventral views, respectively E inner appendages, lateral view F gonocoxite and gonostylus, lateral view G cercus and subepandrial sclerite, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure14" httpUri="https://binary.pensoft.net/fig/1286748">14 A D</figureCitation>
), but can be easily distinguished by the following set of characters: postpedicel lanceolate, approx. as long as length of scape and pedicel combined (Fig.
<figureCitation id="3B9C31FD191989E027EAB3CA3E636BFA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 E, F</figureCitation>
); face slightly gibbous at lower 1 / 3 (Fig.
<figureCitation id="C568D0494FD71DC88A2FA1818CB605EA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
); scutum lacking distinct rows of acrostichal setae; anterior dorsocentral setae absent; femora mostly covered with short black setae (Fig.
<figureCitation id="46C46342FD224108B6FA7D7055E14362" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 C</figureCitation>
); male sternite 8 with a long digitiform projection at posterior margin (Figs
<figureCitation id="ABEC5F12FD9BBF89DA8A05BDDBF838E7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 H</figureCitation>
,
<figureCitation id="10FECA99C1C030837342C959D4991B63" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 C, E</figureCitation>
); gonocoxite L-shaped, with rounded apex and covering the gonostylus versus postpedicel oval, shorter than scape and pedicel combined (Fig.
<figureCitation id="7CFC0DF623436662DD9EABDFDEEFB599" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B habitus lateral and dorsal, respectively, C head and thorax, lateral view D head, anterior view E abdomen, lateroventral view F terminalia, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure13" httpUri="https://binary.pensoft.net/fig/1286747">13 C</figureCitation>
); face distinctly gibbous at lower 2 / 3 (Fig.
<figureCitation id="70A4CA6FACA3C6ED107B0A5594D34ADC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B habitus lateral and dorsal, respectively, C head and thorax, lateral view D head, anterior view E abdomen, lateroventral view F terminalia, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure13" httpUri="https://binary.pensoft.net/fig/1286747">13 C</figureCitation>
); scutum with two distinct rows of acrostichal setae (Fig.
<figureCitation id="F24B6D5982302C746E374C4DA5DC06EB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B habitus lateral and dorsal, respectively, C head and thorax, lateral view D head, anterior view E abdomen, lateroventral view F terminalia, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure13" httpUri="https://binary.pensoft.net/fig/1286747">13 A, C</figureCitation>
); three to four pairs of anterior dorsocentral macrosetae; femora mostly covered with long vestiture of white setae (Fig.
<figureCitation id="5B755A02C7F1CB38F0BE070E357F687F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B habitus lateral and dorsal, respectively, C head and thorax, lateral view D head, anterior view E abdomen, lateroventral view F terminalia, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure13" httpUri="https://binary.pensoft.net/fig/1286747">13 A</figureCitation>
); male sternite 8 without projections at posterior margin; gonocoxite squared at base, with a digitiform apicoventral projection acute apically (Fig.
<figureCitation id="CB73BDDFD7FE71CC8C94549FD0E7C4CF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Myaptex brachyptera (Philippi, 1865) (identified male from Chile) A, B Terminalia in dorsal and ventral views, respectively C, D terminalia without inner appendages in dorsal and ventral views, respectively E inner appendages, lateral view F gonocoxite and gonostylus, lateral view G cercus and subepandrial sclerite, dorsal view." figureDoi="10.3897/zookeys.1232.142494.figure14" httpUri="https://binary.pensoft.net/fig/1286748">14 F</figureCitation>
) in
<taxonomicName id="781285E84B9E203AC244EE511DA74412" authorityName="Hull" authorityYear="1962" class="Insecta" family="Asilidae" genus="Myaptex" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="D08A6702FC9AF93FEC667C06AE1047E7" italics="true">Myaptex</emphasis>
</taxonomicName>
. It is also similar to some undescribed
<taxonomicName id="579C7D06C82199F06627E52825965677">
<emphasis id="0E08A3DBB3237B4FD99147C629AADD7B" italics="true">Lecania</emphasis>
Macquart species
</taxonomicName>
, including
<taxonomicName id="8C0383A1F74389B23916EF17FDC812AE" authority="(Wiedemann, 1828)" baseAuthorityName="Wiedemann" baseAuthorityYear="1828" class="Insecta" family="Asilidae" genus="Nerax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="eurylabis">
<emphasis id="84BF5C2F0FB54CB5C93B77FA37EE7541" italics="true">Nerax eurylabis</emphasis>
(Wiedemann, 1828)
</taxonomicName>
a species that belongs in
<taxonomicName id="68C07AE38DDB49A63A3A018C54A938C5" class="Insecta" family="Asilidae" genus="Lecania" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="18981D93CBFFAF9C5B2ABD606F3EB3EB" italics="true">Lecania</emphasis>
</taxonomicName>
(unpublished data) with inflated male epandria, but can be easily distinguished by the scutellum with one pair of apical macrosetae and the abdominal sternites with macrosetae (scutellum bare in
<taxonomicName id="E6EA96410BE9342D6A21C73B701B232A" class="Insecta" family="Asilidae" genus="Lecania" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="9975DCA67828BB88D1AEBBF0F61958EA" italics="true">Lecania</emphasis>
</taxonomicName>
or at most bearing tiny, short setulae and sternites only with sparse setae).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIAAG" type="distribution">
<paragraph id="03796B0BA93121B6C434027B834DFEA5">
<heading id="DD17D943E058F0751DC56F325B97AC1F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="DF681306AF063350975A643ED6DE0A64">
The new genus is known to occur only in the state of
<collectingRegion id="7D7BAFF0DE5637327BA03BD543DE3B2E" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
(Central-West
<collectingCountry id="37CB2209829CBBDEA50694F0AFA53A68" name="Brazil">Brazil</collectingCountry>
) and department of
<collectingRegion id="3C2A5160810F210D6B2013A2BE08A119" country="Paraguay" name="Cordillera">Cordillera</collectingRegion>
(Central-West of the Oriental Region of
<collectingCountry id="DE512295B317343816800D07620FFD61" name="Paraguay">Paraguay</collectingCountry>
), in biomes of Pantanal and Chaco, respectively (Fig.
<figureCitation id="FF04074032361BAFDD68713E8F05C5F0" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,277 @@
<document id="BD2226CE55666A1D8F3CFAF305C13CCF" ID-DOI="10.3897/zookeys.1232.142494" ID-ZooBank="67DBBE51-5990-4842-AF0F-141CB0AFA4D2" ID-publisher-id="142494" URI-arpha="F990D92B-4A07-5A20-84D0-1366BB179EBD" URI-zoobank="http://zoobank.org/67DBBE51-5990-4842-AF0F-141CB0AFA4D2" article-type="research-article" checkinTime="1742495072448" checkinUser="pensoft" docAuthor="Soares, Matheus M. M., Camargo, Alexssandro &amp; Lamas, Carlos J. E." docDate="2025" docId="6FED5ABB577F536FA4CB5F375D3F8B16" docLanguage="en" docName="ZooKeys 1232: 311-341" docOrigin="ZooKeys 1232" docSource="https://zookeys.pensoft.net/article/142494/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cardiasilus ruda Soares, Camargo &amp; Lamas, 2025, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="341" masterDocId="F990D92B4A075A2084D01366BB179EBD" masterDocTitle="A taxonomic synopsis and identification key to the genera of the Myaptex group (Diptera, Asilidae, Asilinae), with description of a remarkable new genus and three new species from the South American Chaco" masterLastPageNumber="341" masterPageNumber="311" pageNumber="311" updateTime="1742495171300" updateUser="pensoft">
<mods:mods id="3C5F0DB25513E4885898BEAE765CB8AB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC15944994EB8CB6E51C6E44A7F736FB">
<mods:title id="9A158725EC87980E768D775AEB6892FF">A taxonomic synopsis and identification key to the genera of the Myaptex group (Diptera, Asilidae, Asilinae), with description of a remarkable new genus and three new species from the South American Chaco</mods:title>
</mods:titleInfo>
<mods:name id="BC484ED1646010CDD26A2C8470BB65AF" type="personal">
<mods:role id="87FFEC1A100726060907BF505505AED3">
<mods:roleTerm id="22C11FC2863620453EAB4601E7E23318">Author</mods:roleTerm>
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<mods:namePart id="B0C9D10B2C5C78A5664E1D7105F22CC5">Soares, Matheus M. M.</mods:namePart>
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<mods:affiliation id="765F82D453AA8CAD1C838A232FDE278F">Museu de Zoologia da Universidade de São Paulo, Laboratório de Diptera, Av. Nazaré, 481, 04263 - 000 - Ipiranga, São Paulo, São Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:name id="C24E8879A95C96CE1F353355B1A7F313" type="personal">
<mods:role id="23D2B1949A0642BE8BF07E9E56ABCE62">
<mods:roleTerm id="E0EE71D008D18BD9DBAAB0DBFC466458">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FCED28EF6B4694EF732A7775FA91E33">Camargo, Alexssandro</mods:namePart>
<mods:nameIdentifier id="99188ABB9EBAD866CEF4D8547CFA0486" type="ORCID">0000-0001-7408-7223</mods:nameIdentifier>
<mods:affiliation id="ECA902153F277041262A937121C6BAE4">Natural History Museum Vienna, 2 nd Zoological Department. Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name id="35076081673FDC13F747C013FD378162" type="personal">
<mods:role id="02FD6E7210A87727F8CEF7919BA8A951">
<mods:roleTerm id="E012F787E2CCAC18D2D25003068B74D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D3E394D9BDB5A72086CD617834CAEB48">Lamas, Carlos J. E.</mods:namePart>
<mods:nameIdentifier id="2D8E40A71128615B64A8107C130AC649" type="ORCID">0000-0002-7750-590X</mods:nameIdentifier>
<mods:affiliation id="7FB4D958941C0961A8FD70FDAE813E5A">Museu de Zoologia da Universidade de São Paulo, Laboratório de Diptera, Av. Nazaré, 481, 04263 - 000 - Ipiranga, São Paulo, São Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="52BFC5FFA7C353A58A02A4E8F8372724">text</mods:typeOfResource>
<mods:relatedItem id="0E6C42D210BD4B5D09A8F20FA9FA7E9B" type="host">
<mods:titleInfo id="00DC784FB837D3035CCF6B2311524C03">
<mods:title id="369363F8D607219618A86FDCE82E6E40">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="0C2DC5D73C043043A09CF8A0EE2EC312">
<mods:date id="FFC3BA497A2E3A0C80581A34A8C1E31F">2025</mods:date>
<mods:detail id="4D3363B43A72259876D06E558B6988E8" type="pubDate">
<mods:number id="4DC40C1A8207EF52F859163CCCE49B19">2025-03-20</mods:number>
</mods:detail>
<mods:detail id="68EF72CA7DB9F064D737DC6C5920BCC1" type="volume">
<mods:number id="A1318E941DA82D4F932A9358A31A3866">1232</mods:number>
</mods:detail>
<mods:extent id="6D35E92E0C49AD2289EA2EAF951B1516" unit="page">
<mods:start id="9ABEDCCB7BAD3893E3953D6580FF26C2">311</mods:start>
<mods:end id="6E18690EDB59C6779F815AC7BD63F5A4">341</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0E0508C5C83F496096F0E52F8FF96A7E">journal article</mods:classification>
<mods:identifier id="9163EE296F9C8865A4A0C92E205447D0" type="DOI">10.3897/zookeys.1232.142494</mods:identifier>
<mods:identifier id="B0F74F8F2CC52B4D6132782F71E89A7D" type="ZooBank">67DBBE51-5990-4842-AF0F-141CB0AFA4D2</mods:identifier>
</mods:mods>
<treatment id="6FED5ABB577F536FA4CB5F375D3F8B16" ID-arpha="6FED5ABB-577F-536F-A4CB-5F375D3F8B16" ID-zoobank="https://zoobank.org/00CEB9B1-E2CF-4C9B-B998-C8305DA907BB" LSID="urn:lsid:plazi:treatment:6FED5ABB577F536FA4CB5F375D3F8B16" LSID-ZBK="urn:lsid:zoobank.org:act:00CEB9B1-E2CF-4C9B-B998-C8305DA907BB" httpUri="http://treatment.plazi.org/id/6FED5ABB577F536FA4CB5F375D3F8B16">
<subSubSection id="11CCDD5ABFC6F573E04922981A9BB2D9" type="nomenclature">
<paragraph id="CF6E8CD10D09C694E3A5D2BDBF89DA93">
<taxonomicName id="9F4A0CD537D3A2A93D368DDD15D32D97" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda" status="sp. nov.">
<emphasis id="24B563EFEE785D04BAC91A5D19FE844C" italics="true">Cardiasilus ruda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCC6307054CA606E869DD078D64B7E8D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F59DD1412BAA447939CDCA5F201AC9B0" type="description">
<paragraph id="490C23CCBB28D969546D42C9A4631A79">
<figureCitation id="06D98AF9D17CB749A0C3A5A5A0D283F8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">Figs 9</figureCitation>
,
<figureCitation id="58ECD38BE4C82C49D5A910F8B76C22A1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10</figureCitation>
,
<figureCitation id="0EC1A21F636B90AF33E7DA9AC1DDF3F6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cardiasilus ruda sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure11" httpUri="https://binary.pensoft.net/fig/1286745">11</figureCitation>
,
<figureCitation id="CE4485937C7E3201EA7DA3A23E25B138" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHDBG" type="diagnosis">
<paragraph id="6BE5BCA5E82B3AAF2EB8BF8456BEDF13">
<heading id="FFA45B3F1037D2BCCD1D8D12710307E7" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4136184542E9C877CE7AD7933957CF39">
The inner margin of epandrium with a short, dorsal, sub-triangular, pre-apical process followed by a short, ventral, sub-rectangular, anteriorly curved, apical process (Fig.
<figureCitation id="265CAB5A6FBCE514558A5DB70BDF9409" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10 A, F, G</figureCitation>
) separates this species from its congeners.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERDBG" type="description">
<paragraph id="76F0678FC0D5FAE7E66D0636F99AFBC0">
<heading id="7CFD38A4169B8E1512F0D54C1970E641" reason="title">Description.</heading>
</paragraph>
<paragraph id="0CBC5AD0B2F9E24463460D7207134D2C">
<emphasis id="35D985DAD9CB2B060FBC724777E99C53" bold="true">Male</emphasis>
(Fig.
<figureCitation id="08FA692341C8D7380CD07D8257383AC1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">9 A</figureCitation>
). Body Length:
<quantity id="16E0404717BB4D763B6F5CAED85945E9" metricMagnitude="-2" metricUnit="m" metricValue="1.03" unit="mm" value="10.3">10.3 mm</quantity>
; wing length:
<quantity id="FDE34A46336987B233C7C046DE4B8E4D" metricMagnitude="-3" metricUnit="m" metricValue="5.7" unit="mm" value="5.7">5.7 mm</quantity>
.
<emphasis id="6D374287061279FDE67D758426AD4C63" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="835B66051E651E6435B5FDEE3AA17D27" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">9 A D</figureCitation>
). Similar to
<taxonomicName id="71A7FC3B70FA7E30EADAE4386FB87870" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="E25BF3A18E03E275E1270CA221D731E5" italics="true">C. aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4929E76BC052F4A31109B8F309D695E7" rank="species">sp. nov.</taxonomicNameLabel>
except as noted:
<emphasis id="2810FEA62477D7941C51190AC57E7B39" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="1A6E2A88ED846BC932397FA8B949AD2F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cardiasilus ruda sp. nov. (male holotype, before dissection) A, B habitus lateral and dorsal, respectively C head, anterior view D head and thorax, lateral view E abdomen, ventral view F wing." figureDoi="10.3897/zookeys.1232.142494.figure9" httpUri="https://binary.pensoft.net/fig/1286743">9 A, B, D</figureCitation>
). Two pairs of dorsocentral macrosetae.
<emphasis id="7769A6839AC08AE12349E42AE9A883A0" bold="true" italics="true">Terminalia</emphasis>
(Figs
<figureCitation id="A541C240ACFD383DF7C12E0DEC05E2FB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10</figureCitation>
,
<figureCitation id="16D773B3CB1D6B042DCAECC431C7F361" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cardiasilus ruda sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure11" httpUri="https://binary.pensoft.net/fig/1286745">11</figureCitation>
). Epandrium with a short, dorsal, sub-triangular, pre-apical process followed by a short, ventral, sub-rectangular, anteriorly curved, apical process (Fig.
<figureCitation id="F704C7FAA30671BF98BFF15FF9275468" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10 A, F, G</figureCitation>
). Subepandrial sclerite goblet-shaped (Fig.
<figureCitation id="41379A7B68CD284C60873AB38D5E77D1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cardiasilus ruda sp. nov. (male holotype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite. Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure11" httpUri="https://binary.pensoft.net/fig/1286745">11 D</figureCitation>
).
</paragraph>
<caption id="D637B9358E5ED4CDAEE0BA335FEDD124" ID-DOI="10.3897/zookeys.1232.142494.figure9" ID-arpha="7FF862B3-0E34-5DDE-B1C2-EB95E329AF36" httpUri="https://binary.pensoft.net/fig/1286743" startId="F9">
<paragraph id="070897841AD3E68FC70416B52701E692">
<label id="BCE748F3CC8C55B34018E8DB3A49E7DB">Figure 9.</label>
</paragraph>
<paragraph id="52728EFD1CA8A1142026EF78EB2C5133">
<taxonomicName id="96764E40FDB0BD15086EF31E9628B336" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda" status="sp. nov.">
<emphasis id="58D36F4DC99B991CAA3AB5B765459970" italics="true">Cardiasilus ruda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F04ED1075C982D1C3B9EC293634E461" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype, before dissection)
<emphasis id="04F6E1248C4DDDC6BDFE05E0AE9FE2A9" bold="true">A, B</emphasis>
habitus lateral and dorsal, respectively
<emphasis id="4480A6FFC4210FE0D09ECB8DCDA7F95F" bold="true">C</emphasis>
head, anterior view
<emphasis id="A38026F4A2B520F47CF18871B49C3EFB" bold="true">D</emphasis>
head and thorax, lateral view
<emphasis id="D0D419A8C58FADD5017ED11B5FBE2AB0" bold="true">E</emphasis>
abdomen, ventral view
<emphasis id="8DC71827C13A9E34B908B5EEAD097FA6" bold="true">F</emphasis>
wing.
</paragraph>
</caption>
<caption id="B7716F479B93B8C6A23B639A4A19271B" ID-DOI="10.3897/zookeys.1232.142494.figure10" ID-arpha="74655C58-3DA0-5E43-BC3A-46A2BF084DE7" httpUri="https://binary.pensoft.net/fig/1286744" startId="F10">
<paragraph id="888375859811FA3034A19B6C2BA5E5E9">
<label id="AC40856E148E54340CC9606C13C2ABE1">Figure 10.</label>
</paragraph>
<paragraph id="70FE7786DBFAF90F781D53D8A7548DFA">
<taxonomicName id="22032235DAFF6349C09F72C4C03A3136" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda" status="sp. nov.">
<emphasis id="973378DB51CF0CA1E8D9CE487A9CE320" italics="true">Cardiasilus ruda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32AC8DFACF2D7EF777B89406E7B03A3F" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype)
<emphasis id="A70FACE4EEDAF51B14D191A68C9C7612" bold="true">A C</emphasis>
terminalia in dorsal, lateral, and ventral views, respectively
<emphasis id="7B50B02E4CAD2EACB88FD39A73DFB27E" bold="true">D</emphasis>
tergite 8,
<emphasis id="F8EBD489A761E967C4E11DB5E3935AEE" bold="true">E</emphasis>
sternite 8
<emphasis id="57E5A94B18039C606D867DAD66C850F8" bold="true">F, G</emphasis>
dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium.
</paragraph>
</caption>
<caption id="44AD496444F5D2D98F72D7EAD189F994" ID-DOI="10.3897/zookeys.1232.142494.figure11" ID-arpha="9A7F9731-99B4-5F1A-AAE3-82CC086E1FCC" httpUri="https://binary.pensoft.net/fig/1286745" startId="F11">
<paragraph id="4124F8070DDC5C815803483B2876261F">
<label id="7134EE867146A321BFDA4839A45B1E89">Figure 11.</label>
</paragraph>
<paragraph id="7E86852E563672E8D05741DEEAAB583F">
<taxonomicName id="9DFB43300EE775AEBBF404EE4010DAAC" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda" status="sp. nov.">
<emphasis id="04C7F6681C765FEC325A4A46F45D7A0C" italics="true">Cardiasilus ruda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8C776E45ECE64D427D4E238CD7C8C9" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype)
<emphasis id="077874599DF01FA5E276D71C52123CAE" bold="true">A</emphasis>
inner appendages
<emphasis id="7473034FC9D0ABC6941AA8B2CD05ED42" bold="true">B</emphasis>
gonocoxite and gonostylus
<emphasis id="17C6C5605E651945082AB9E6FAE3FC2A" bold="true">C</emphasis>
phallus and ejaculatory apodeme
<emphasis id="0062477636143C5BE2E8EF004B8B18C6" bold="true">D</emphasis>
subepandrial sclerite. Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus.
</paragraph>
</caption>
<paragraph id="121F008E63F93121BCCD841AF320F7D7">
<emphasis id="3DB0B1F314F94B22390C2B64E486332B" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPIBG" type="type material">
<paragraph id="F766861D5AA69D03D9CD09185E3D9D70">
<heading id="73EC6C661CD537CC302E0C41630C0147" reason="title">Type examined material.</heading>
</paragraph>
<paragraph id="1B36F062E3958005865FB9B03800CF87">
<materialsCitation id="1DCF2C364DDB68880FE920E9E00C06E3" collectingDate="1971-12-25" collectionCode="MZUSP" country="Paraguay" latitude="-25.502556" location="Colonia Pirareta" longLatPrecision="1" longitude="-56.98336" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="19804E29CF391A4A4152F13A02FDBD33" bold="true" italics="true">
<typeStatus id="CDCAE27BCD4E7FF6C3EC8D3CEB914ED8" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="0C676A5E4B47DFE704923A302F2A8FE7" count="1" type="male"></specimenCount>
(
<named-content id="0792B3E106603D2CA11EADE61387FF5B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="33F838A65E6366BA09E72B9A5F69C2D6" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
) labelled: “
<collectingCountry id="90B122098F09B48C77C881704BC2CA98" name="Paraguay">PARAGUAI</collectingCountry>
|
<location id="5CAF46E3412BD529596F878CE23B7F21" LSID="urn:lsid:plazi:treatment:6FED5ABB577F536FA4CB5F375D3F8B16:5CAF46E3412BD529596F878CE23B7F21" country="Paraguay" latitude="-25.502556" longLatPrecision="1" longitude="-56.98336" name="Colonia Pirareta">Colônia Piraretá</location>
[ca
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECJBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-56.983361,-25.502556]}">
<geoCoordinate id="35141D18E808A5D7126C610B96FEDECD" degrees="25" direction="south" minutes="30" orientation="latitude" precision="1" seconds="09.2" value="-25.502556">25 ° 30 ' 09.2 &quot; S</geoCoordinate>
,
<geoCoordinate id="21148AC5E9F0DCE8819E1598DDF0E621" degrees="56" direction="west" minutes="59" orientation="longitude" precision="1" seconds="00.1" value="-56.98336">56 ° 59 ' 00.1 &quot; W</geoCoordinate>
</named-content>
] |
<date id="6B635D8A4BB8639B77FCB0F0E95B8A53" value="1971-12-25">
<collectingDate id="1A38AB18A7318B44C637C68DFADCA540" value="1971-12-25">25. xii. 1971</collectingDate>
</date>
” “
<typeStatus id="2917A882C263C3956C45AD98A8FD8035" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="F896F8949F5AEC88C164D39F4C49C7C4" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ruda">
<emphasis id="120897CAD4EB0AD459D69FA0E54D891C" italics="true">Cardiasilus ruda</emphasis>
</taxonomicName>
| Soares, Camargo &amp; Lamas [red label] ”
</materialsCitation>
.
<typeStatus id="82F6C1214672413A959C9AA95303E451" type="holotype">Holotype</typeStatus>
condition: Both postpedicel and mid legs broken off, terminalia dissected and stored in microvial at the same pin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESJBG" type="remarks">
<paragraph id="EA5DF7A7E15E8AAD17319B9E4F075638">
<heading id="B7717ADDBA96BE1DEE9E83CC92558A12" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="EBBCC87BAC157C4B97B960439224EDF5">
The new species is remarkably similar to
<taxonomicName id="2AAF0065626B4A0ED0F5273D2610FBB4" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="4B76A9E8DD9440F3683B45A12CB72A93" italics="true">C. aysu</emphasis>
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<taxonomicNameLabel id="F31ED9274DBCDFBD8A81BD2E2064734F" rank="species">sp. nov.</taxonomicNameLabel>
, differing only in the shape of epandrium with a short, dorsal, sub-triangular, pre-apical process followed by a short, ventral, sub-rectangular, anteriorly curved, apical process (Fig.
<figureCitation id="6BE042FD7E9A40CA91D58272508550EA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10 A, F, G</figureCitation>
). The epandrium lacks such processes at dorsal inner margin in
<taxonomicName id="380EF732392BB979495089BBFA09FFA1" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="D40E3D9BBD2D57CB2292837A7F8ACB32" italics="true">C. aysu</emphasis>
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<taxonomicNameLabel id="CFBAD585EC9A745D571BE3EE294ED3FA" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="B4CFC24E60FFC32584C982B6A02574DE" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dangeloi" status="sp. nov.">
<emphasis id="50713834C9E47F50DACF87DA869C0BC6" italics="true">C. dangeloi</emphasis>
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<taxonomicNameLabel id="27525818E2D4902EB83A3B1A1AF0E97C" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4KBG" type="distribution">
<paragraph id="F2061562D88AE37CDFF8880B12405F39">
<heading id="498D40E5203152F092068CD65512EA7E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="889CB19B3047B860854BA488BD6CA914">
<collectingCountry id="19CE163CF31E862BCF7A9E2F5CAF881A" name="Paraguay">Paraguay</collectingCountry>
(department of
<collectingRegion id="058F489DC2DCED591304D1C5E28E21C9" country="Paraguay" name="Cordillera">Cordillera</collectingRegion>
) (Fig.
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).
</paragraph>
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<subSubSection id="SECID0EHLBG" type="etymology">
<paragraph id="91A6FF925EC43D66E507AACC63762EDF">
<heading id="BA49AFA7564B61499E11D3D36CA2D24C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="DD0820677EA87DD077A8D9173C0BDEC3">
Rudá is the deity of love in the pantheon of deities of Tupi-Guarani culture. It alludes to the male epandria that resembles an ideogram of a heart (Fig.
<figureCitation id="35327595B43EEA07B9DADCDA0C8B65A6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cardiasilus ruda sp. nov. (male holotype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8, E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; dp epand = dorsal process of epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; tg 8 = tergite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure10" httpUri="https://binary.pensoft.net/fig/1286744">10 A</figureCitation>
). Treated as a noun in apposition.
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<comment id="56E9219283E8187B7E3D8AB26ABA2183">
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species.
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<taxonomicName id="370EAE41BBD8A0CEE97768AE52CF8D87" authority="Thorell, 1878" authorityName="Thorell" authorityYear="1878" class="Insecta" family="Ichneumonidae" genus="Boethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="spinimanus">
<emphasis id="B21AB9ACEA979624BD06A8379A1DF48A" italics="true">Boethus spinimanus</emphasis>
<bibRefCitation id="72E19888C2610B1427CFDFF837482DE5" author="Thorell T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pagination="5-317" refId="B7" refString="Thorell T (1878) Studi sui ragni Malesi e Papuani. II. Ragni di Amboina raccolti Prof. O. Beccari. Annali del Museo Civico di Storia Naturale di Genova 13: 5-317." title="Studi sui ragni Malesi e Papuani. II. Ragni di Amboina raccolti Prof. O. Beccari." volume="13" year="1878">Thorell, 1878</bibRefCitation>
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<subSubSection id="641CAC8A3FB000EF2C09FE5E7C8A62FC" type="reference_group">
<paragraph id="58714A3B184FF67B7E246918B666E98B">
<treatmentCitationGroup id="C9F8D66217D4BC7A3C6E6A226A98674F">
<treatmentCitation id="6107AA158DCCE94692211FA88480182E" author="Wanless FR" page="155" year="1984">
<taxonomicName id="17012321830FDB7B64465189AFE05121" ID-CoL="7SSH" authority="Wanless, 1984: 155" authorityName="Wanless" authorityPageNumber="155" authorityYear="1984" class="Arachnida" family="Salticidae" genus="Taraxella" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="B47A64778BE956C7390DACFA806617B3" italics="true">Taraxella</emphasis>
<comment id="C8D49101C1DBC14155C82376700958C1">
<bibRefCitation id="E34134F0F4B10874C6E28EE6157D25A3" DOI="10.5962/bhl.part.15964" author="Wanless FR" journalOrPublisher=")" pagination="135-205" refId="B10" refString="Wanless FR (1984) A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool.) 46: 135-205. https://doi.org/10.5962/bhl.part.15964" title="A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool." volume="46" year="1984">Wanless, 1984: 155</bibRefCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQXAE" type="type species">
<paragraph id="7635A46988DB00CBA7043E0C56014E17">
<heading id="C76740C991311C7C4BD1286DAB1700D1" reason="title">
<typeStatus id="18FA2C0095EE1FB20447E2034F91F2E3">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="E27BD96C3EB5F64A0AEC771778F36D10">
<taxonomicName id="266F3A22889C49FE074548D93DFCDAE2" authority="Wanless, 1984" authorityName="Wanless" authorityYear="1984" class="Arachnida" family="Salticidae" genus="Taraxella" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="F00D140C1D059E9E7FD9AE3D8530BB7E" italics="true">Taraxella solitaria</emphasis>
<bibRefCitation id="C2E3EE9C8601D86BA734746155814272" DOI="10.5962/bhl.part.15964" author="Wanless FR" journalOrPublisher=")" pagination="135-205" refId="B10" refString="Wanless FR (1984) A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool.) 46: 135-205. https://doi.org/10.5962/bhl.part.15964" title="A review of the spider subfamily Spartaeinae nom. n. (Araneae: Salticidae) with descriptions of six new genera. Bulletin of the British Museum of Natural History (Zool." volume="46" year="1984">Wanless, 1984</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,587 @@
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<mods:title id="3721CB5919B2508E80244C954462A241">A taxonomic synopsis and identification key to the genera of the Myaptex group (Diptera, Asilidae, Asilinae), with description of a remarkable new genus and three new species from the South American Chaco</mods:title>
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<mods:namePart id="AAD8BB82AB49CF78D9FDDDA8BC359267">Soares, Matheus M. M.</mods:namePart>
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<subSubSection id="B619F16A193A11CC35821926428AC0C3" type="nomenclature">
<paragraph id="F4DCEFB328F6194BF18DAE153F6B8F11">
<taxonomicName id="356C3102F80FF67CBD3916F45BDC5EF6" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="6FEA52220DC6B886A4EF79526F610326" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5880B1F53775B51359803C7544650184" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="273CB3F8E939A10C42E5514318D7C596" type="description">
<paragraph id="8F83B2A9AA1AB651352D70AF180BAC2F">
<figureCitation id="F9A02DAF955C7E7FF16FFBE03435B2D2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">Figs 2</figureCitation>
,
<figureCitation id="FFFC4A4D6185DFF562F9CC5D22324B2E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3</figureCitation>
,
<figureCitation id="1D9C8870D927557C81B8AC7220CA1B10" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4</figureCitation>
,
<figureCitation id="3EF3969ABA3F97868253C8194A4362F0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cardiasilus aysu sp. nov. (identified female) A, B habitus lateral and dorsal, respectively C E terminalia in dorsal, ventral, and lateral views, respectively F spermathecae. Abbreviations: cerc = cercus; hyprct = hypoproct; lat sc = lateral sclerite of genital fork; st = sternite; tg = tergite." figureDoi="10.3897/zookeys.1232.142494.figure5" httpUri="https://binary.pensoft.net/fig/1286739">5</figureCitation>
,
<figureCitation id="10013EDD68DD9A47BEBAD31B79605F74" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3HAG" type="diagnosis">
<paragraph id="75B0C14AF8B6F9DD39CF315677F9B668">
<heading id="C17BB39309D2FAD2F396D3E61C6D6B11" reason="title">Diagnosis</heading>
</paragraph>
<paragraph id="E9FEBEA3478D995C79B7223F3737988E">
<emphasis id="13C42EDC54598EAE6C1C9B9D99EB4EC9" bold="true">(males).</emphasis>
The new species can be easily distinguished from the congenerics by the yellow postalar macrosetae (Fig.
<figureCitation id="EBFC81D44BE3F6597BC1F92C1713C759" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 B, C</figureCitation>
), inner dorsal margin of epandrium weakly excavated, and lacking processes (Fig.
<figureCitation id="56F51156076E9BE7B1A06846F0639A22" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 A, F, H</figureCitation>
) and subepandrial sclerite somewhat diamond-shaped near its middle (Fig.
<figureCitation id="B55F93C0BD37F6620084463813864BF8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 D</figureCitation>
).
</paragraph>
<caption id="9EC5A0C41297245EB4639F2E38BF65B9" ID-DOI="10.3897/zookeys.1232.142494.figure4" ID-arpha="03FC1287-B4CB-54C0-A405-50EE5520BF98" httpUri="https://binary.pensoft.net/fig/1286738" startId="F4">
<paragraph id="F9E0DC06B3187DA6CCE7CE2ACAA0FE6D">
<label id="C519318D98C87C9A1FD674979889BE3B">Figure 4.</label>
</paragraph>
<paragraph id="6529F61625ABF50BF3F72DCC2F57E045">
<taxonomicName id="0B7FA18D86B0BF50FB3C589AA3F27AB6" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="FE1CF48DF59C527CEA559A09AAB39900" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88E3FDF4E4AA61393D5E5999DA6910A6" rank="species">sp. nov.</taxonomicNameLabel>
(male paratype)
<emphasis id="07924ADFF5B074C3CCA70D1B7F18138D" bold="true">A</emphasis>
inner appendages
<emphasis id="802D7039E5EC60878E03781C3E40A56A" bold="true">B</emphasis>
gonocoxite and gonostylus
<emphasis id="69874835BCE26705D39F731A3778A532" bold="true">C</emphasis>
phallus and ejaculatory apodeme
<emphasis id="0DA633D0ECDFE297B931A3D67B50F4D8" bold="true">D</emphasis>
subepandrial sclerite
<emphasis id="3007BE1E771EBC5BD4E9096F31EA6882" bold="true">E</emphasis>
hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESJAG" type="description">
<paragraph id="A97E44B6F7A3DA5E6216C755A772D05F">
<heading id="CFB14BEFA7F3ED8F76EB81FC76B90DA0" reason="title">Description.</heading>
</paragraph>
<paragraph id="BD8571E703DDAC119F32AB576C7865DF">
<emphasis id="C83CADEDF2312148BB353879C5A4561E" bold="true">
Male
<typeStatus id="D71510F57D7D3F2569480975005255AC" type="holotype">holotype</typeStatus>
</emphasis>
(Fig.
<figureCitation id="65935CDBEB50AE73863639E8ADA7FFFD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A</figureCitation>
). Body Length
<quantity id="54F2B74B70BECA5FE51D9154A1915CD0" metricMagnitude="-2" metricUnit="m" metricValue="1.23" unit="mm" value="12.3">12.3 mm</quantity>
; wing length
<quantity id="C346EFBC3C673A244CAF4C5D2F9A290C" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
.
<emphasis id="E254928D2489D0D83008104ADEB4A771" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="FA047F2C774AEF698A99FAC3FDA68016" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A F</figureCitation>
). Scape and pedicel orange-yellow, slightly darker at apex; postpedicel dark brown, lanceolate; stylus dark brown, bare, first element very short, second element long and abruptly narrowed at apex. Face golden pruinose, except gibbosity with mixed silvery and golden pruinosity; mystax with few upper slender black macrosetae and strong golden macrosetae below, extending along oral margin. Frons, golden pruinose, with long black orbital macrosetae. Ocellar tubercle dark brown, with one to two pairs of proclinate long and slender ocellar macrosetae; vertex golden pruinose, almost bare of major setae, only with few slender black setae posteriorly; upper-most four to five postocular macrosetae black, remaining macrosetae yellowish-white. Postcranium golden pruinose, with long, slender, and dense white lower occipital setae. Palpus dark brown, with few short white setae. Proboscis black, ventral surface with long white setae, apex with short white setae.
<emphasis id="E1A8C89DE7CE6777650AD80DA6026707" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="259B8FDF51AEC50137BDDF4F5BEC2938" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A C</figureCitation>
). Antepronotum with few strong golden macrosetae, covered with golden pruinosity and sparse short and white setae. Scutum wholly covered with short vestiture of black setae and golden pruinosity, except notopleural, supra-alar, and acrostichal area close to scutellum with short white setae, in anterior view with U-shaped dark brown median stripe along acrostichal area, followed by L-shaped dark brown paramedian stripe along dorsocentral and intra-alar areas, with both stripes extending slightly beyond transverse suture. Three to four pairs of black, postsutural dorsocentral macrosetae, two black notopleural macrosetae, one black supra-alar macroseta, two yellow postalar macrosetae (anterior one shorter). Scutellum with one pair of apical pale yellow macrosetae, dorsal surface covered with short white setae. Pleuron mostly silvery pruinose, except anepisternum and katepisternum with golden pruinosity anteriorly, anepisternum, katepisternum, anepimeron and meron with few long white setae posteriorly, katatergite with vertical row of yellow macrosetae.
<emphasis id="B9A19B6CF3E434DFB8F13DAA4BEE568F" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="F30ED1D597C5E4171D43AF87ECB48873" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A</figureCitation>
). Mostly yellow, except all coxae with dense silvery pruinosity, all femora dorsally with short black setae, apex of mid and hind femora dorsally, apex of all tibiae, apex of all tarsomeres one to four and all tarsomeres five dark brown. Legs with white or golden macrosetae, except when as noted.
<emphasis id="22F46A70F2C8D925C5A9DE046FAC57A0" bold="true">Leg I.</emphasis>
Coxa with long and dense white macrosetae anteriorly. Femur with one posterior black macroseta at middle, ventral surface with long setae at basal 1 / 2. Tibia with one anterodorsal black macroseta at basal 1 / 4, three dorsal black macrosetae, one near middle and two at apical 1 / 2, one antero, one dorsal, one posterodorsal, and one ventral black macrosetae at apex, two anterior long macrosetae at apex (~ 2 × longer than remaining apical macrosetae), posterior row of four long macrosetae from basal 1 / 4 to apex, three ventral long black setae near middle. Basitarsus with one antero and one posteroventral short black macrosetae at apex, one posteroventral long macroseta near base, tarsomeres one to three with crown of long macrosetae at apex: one antero and one posteroventral, one antero and one posterodorsal.
<emphasis id="AA6A53104042E311667763EDAF268D7F" bold="true">Leg II.</emphasis>
Coxa with long and slender macrosetae at apical 1 / 2, some of them reaching lateral surface. Ventral surface of femur with short vestiture of white setae and five to six long ventral macrosetae at basal 1 / 2, one anterodorsal apical short black macroseta, one posterodorsal preapical short macroseta, anterior row of four macrosetae from base to apex, anteroventral row of short intermixed black and white macrosetae. Tibia with two to three short dorsal black macrosetae at apical 1 / 2, one ventral long black macroseta at middle, ventral row of short and slender black setae from basal 2 / 4 to apex, one posterior long and slender black seta at basal 1 / 3, two ventral strong macrosetae, one black at middle and one white preapical, one posterior strong macroseta at apical 1 / 3, crown of long intermixed black and white macrosetae at apex: one antero and one posteroventral, one antero and one posterodorsal, one anterior and one posterior. Tarsus with chaetotaxy similar to fore tarsus.
<emphasis id="8EC93CEACD42397EB5B131AE3FE6A245" bold="true">Leg III.</emphasis>
Coxa laterally with two posterior macrosetae. Femur with two to three anterior macrosetae, anteroventral row of short macrosetae, posteroventral row of long and slender setae at basal 1 / 2, few long and slender posterodorsal setae near base, one anterodorsal black preapical macroseta, one dorsal preapical macroseta. Tibia with one antero and one posterodorsal short black macrosetae near base, two long anterodorsal black macrosetae: one near middle and one at apical 1 / 2, two long anteroventral black macrosetae at apical 1 / 2, crown of black macrosetae at apex: one antero and one posteroventral, one ventral, one anterior and one dorsal. Tarsus with chaetotaxy similar to fore and mid tarsus.
<emphasis id="A321500AD1C5F19FF373DFFB2B1507D5" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="45F7E963E81E7C8454A8C6827B715D13" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 G</figureCitation>
). Hyaline, veins brown, orangish at base and Sc. Membrane with sparse dark brown microtrichia at apex of cells
<emphasis id="899DB92593AF4738D551D4324FAE0E54" italics="true">
r
<subScript id="B55DC49191C89D211A9B028D633583A9">1</subScript>
</emphasis>
,
<emphasis id="7226F8AA7CDB7A07059798432DC84378" italics="true">
r
<subScript id="7D4FF400E08CF7C75E0D92F986688B59">2 + 3</subScript>
</emphasis>
,
<emphasis id="CF2D8D90898ECC107CEB6DAEC94C4C8A" italics="true">
r
<subScript id="1EA52552C1368141B1A678A297A063A7">4</subScript>
</emphasis>
and bordering the veins
<emphasis id="E0C41D4E71200457BFAAB3FEE06B4750" italics="true">
<collectionCode id="3FF5C9DF32F918F9AEE5AE6F4C8D23B0" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="4A1759B9BBFA863AC1D0887AF30BBE22">4</subScript>
</emphasis>
and
<emphasis id="3B0F815790F004FEE296DFCBF30A67B2" italics="true">
<collectionCode id="7422554EBDE4FD127716280BE1C938A0" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="8FD0E37C8EA99AAFFE1A95F365D5405F">5</subScript>
</emphasis>
. Halter: yellow.
<emphasis id="979DF8AB585A9CE86B44BB0E8DD9F9AA" bold="true" italics="true">Abdomen</emphasis>
(Fig.
<figureCitation id="00123E1B1AB8614B5FBF10B24549998E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cardiasilus aysu sp. nov. (male holotype) A, B habitus lateral and dorsal, respectively C head and thorax, lateral view D head, anterior view E, F antenna, lateral and dorsal views, respectively G wing H abdomen, ventral view. Abbreviations: d = discal cell; m 3 = third medial cell; r 1 = first radial cell; r 2 + 3 = second + third radial cell; R 2 + 3 = second branch of radius; R 4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius." figureDoi="10.3897/zookeys.1232.142494.figure2" httpUri="https://binary.pensoft.net/fig/1286736">2 A, B, H</figureCitation>
). Mostly brown, becoming orange from segments five to eight, densely covered with golden pruinosity, except lateral margins of tergites one to four, silvery pruinose. Posterior margin of tergite one with six to seven long black macrosetae, lateral margin with seven to eight white lateral marginal macrosetae, lateral margins of tergites two to six with two strong white lateral marginal macrosetae, tergites mostly clothed by short vestiture of black setae, becoming white laterally. Sternites one to four densely silvery pruinose, sternites five to eight mostly orange, with weak silvery pruinosity, sternites two to six with two to three pairs of pale yellow macrosetae mid-laterally and clothed with short, sparse, white setae.
<emphasis id="AE17462B84272C7D7E7F9601429C1336" bold="true" italics="true">Terminalia</emphasis>
(Figs
<figureCitation id="D6F4B6F4503DE7AE3731D4AF86B2BCC9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3</figureCitation>
,
<figureCitation id="833CF21370824EC527AF7EFAA9C85068" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4</figureCitation>
). Orange-brown. Tergite eight somewhat saddle-shaped, narrowing at middle of anterior and posterior margins, posterior 1 / 2 with two to three rows of short setae, longer at posterior corners (Fig.
<figureCitation id="16166EA775303B103F36FC37041D04B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 D</figureCitation>
). Sternite eight with short and slender white setae at posterior 1 / 3, with long digitiform projection at middle of posterior margin, ~ 2 / 3 as long as sternite eight length, with conspicuous yellow setae (Fig.
<figureCitation id="A82445F3AEC07E5A207726DF713190B9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 E</figureCitation>
). Epandrium inflated laterally and posteriorly, resembling the ideogram of a heart in dorsal view, lacking inner and apical projections dorsally, inner ventral margin with a short preapical dentiform process, mostly with short vestiture of brownish setae, except apex with slightly longer yellowish setae (Fig.
<figureCitation id="F7030CA83D1A2043F82C974222E27EC9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 A C, F I</figureCitation>
). Cercus short, digitiform, laterally compressed, covered with short setae (Fig.
<figureCitation id="FA69A6841BC9A11923F337F453B03BFB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cardiasilus aysu sp. nov. (A C male holotype D I male paratype) A C terminalia in dorsal, lateral, and ventral views, respectively D tergite 8 E sternite 8 F, G dissected terminalia in dorsal and ventral views, respectively H, I dissected terminalia without inner appendages in dorsal and ventral views, respectively. Abbreviations: cerc = cercus; epand = epandrium; goncx = gonocoxite; sbepand scl = subepandrial sclerite; st 8 = sternite 8; vp epand = ventral process of epandrium." figureDoi="10.3897/zookeys.1232.142494.figure3" httpUri="https://binary.pensoft.net/fig/1286737">3 A</figureCitation>
). Subepandrial sclerite long and narrowing towards apex, with a median subtriangular lateral process (somewhat diamond-shaped near its middle), almost at the same level, internally, with a pair of short, tooth-like processes directed anteriorly, and with a short dentiform preapical process, apex dorsoventrally flattened and covered with short setae (Fig.
<figureCitation id="949F803D14EB7DF7D16D6024275419AB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 D</figureCitation>
). Hypandrium subrectangular, with a short concavity at posterior margin, covered with short setae at posterior 1 / 2 (Fig.
<figureCitation id="84DF6104A827B19B5089D6B2C28F54D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 E</figureCitation>
). Gonocoxite L-shaped, pointed at base and rounded at apex, with slightly preapical concavity at dorsal margin, few short setae at basal 2 / 3 of external surface (Fig.
<figureCitation id="4D2BE0AC87E61BB1FA23ADAE0DC93E26" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 A, B</figureCitation>
). Gonostylus ~ 1 / 2 as long as gonocoxite, somewhat digitiform, with ventral indentation at apical 1 / 3, apex rounded (Fig.
<figureCitation id="7DB89FB538ACFC94202CA9C9F4A0D753" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 A, B</figureCitation>
). Ejaculatory apodeme fan-shaped (Fig.
<figureCitation id="1D2C496D48D33B20CCA5483B93AC808A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 C</figureCitation>
). Phallus long and thin, longer than length of hypandrium plus gonocoxite, divided into two prongs along the entire length (Fig.
<figureCitation id="6A440FE3B0E13B936502DAD0DA1E1904" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cardiasilus aysu sp. nov. (male paratype) A inner appendages B gonocoxite and gonostylus C phallus and ejaculatory apodeme D subepandrial sclerite E hypandrium. Abbreviations: ej apod = ejaculatory apodeme; hypd = hypandrium; goncx = gonocoxite; gonst = gonostylus; ph = phallus." figureDoi="10.3897/zookeys.1232.142494.figure4" httpUri="https://binary.pensoft.net/fig/1286738">4 C</figureCitation>
).
</paragraph>
<paragraph id="061A483DD3BB147DCC5735313CE25C0F">
<emphasis id="285246B557097A564DB1F7B9EF784E0A" bold="true">Female</emphasis>
(Fig.
<figureCitation id="D056DDC29F67DEE6A15C16392E9E815B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cardiasilus aysu sp. nov. (identified female) A, B habitus lateral and dorsal, respectively C E terminalia in dorsal, ventral, and lateral views, respectively F spermathecae. Abbreviations: cerc = cercus; hyprct = hypoproct; lat sc = lateral sclerite of genital fork; st = sternite; tg = tergite." figureDoi="10.3897/zookeys.1232.142494.figure5" httpUri="https://binary.pensoft.net/fig/1286739">5</figureCitation>
). Similar to male, except as noted: Body length
<quantity id="8B0DE909DAEA0EFC0C32E4D1E481DDF0" metricMagnitude="-2" metricUnit="m" metricValue="1.35" unit="mm" value="13.5">13.5 mm</quantity>
; wing length
<quantity id="40B45D7A3EEF5DF87115AB27E12FB8EF" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
. Abdomen wholly brown, only the segment seven and the basal 1 / 2 of ovipositor sometimes orange, tergite seven with dark brown macrosetae posteriorly, sternite seven with white macrosetae laterally and posteriorly.
<emphasis id="18C09CF267E4FEA823A9A513F301D3E5" bold="true" italics="true">Terminalia</emphasis>
(Fig.
<figureCitation id="20C977024880D770D8DEAD2FD19E4610" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cardiasilus aysu sp. nov. (identified female) A, B habitus lateral and dorsal, respectively C E terminalia in dorsal, ventral, and lateral views, respectively F spermathecae. Abbreviations: cerc = cercus; hyprct = hypoproct; lat sc = lateral sclerite of genital fork; st = sternite; tg = tergite." figureDoi="10.3897/zookeys.1232.142494.figure5" httpUri="https://binary.pensoft.net/fig/1286739">5 C F</figureCitation>
). Laterally compressed from middle of tergite and sternite eight, long and slender, almost two times the length of tergite seven, covered with short black setae, longer before the opening of the genital fork, apex of sternite eight curved ventrally, weakly sclerotized, bare, and strongly striated. Tergite 9 + 10 slightly longer than cercus, mostly bare and shiny, only covered with few short and sparse white setae. Cercus short, digitiform, covered with short and slender white setae. Hypoproct short, V-shaped. Two sclerotized and rugose spermathecae present, membranous at base.
</paragraph>
<caption id="BE06C7A30A6094800C5E6A5C078F7BF0" ID-DOI="10.3897/zookeys.1232.142494.figure5" ID-arpha="A05C8DE9-05C8-55FF-BFC0-7A6CE7CD904D" httpUri="https://binary.pensoft.net/fig/1286739" startId="F5">
<paragraph id="812F0A4E336E94084CDBD1D739C8DA03">
<label id="D0DAE8BF884C5A5AFBE2E83A9FB6CCCF">Figure 5.</label>
</paragraph>
<paragraph id="D07886BE79385A111CB1019CB0D43FEB">
<taxonomicName id="3F45CDACCCDA67CA40DE9283E172FBEE" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu" status="sp. nov.">
<emphasis id="303289430F057587ECB77953A099C178" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C9AC11C6DEAED802977CA911F2D59F6" rank="species">sp. nov.</taxonomicNameLabel>
(identified female)
<emphasis id="8C70EE353584E87913CE7A7DC5F6C9D4" bold="true">A, B</emphasis>
habitus lateral and dorsal, respectively
<emphasis id="406E48EEAF9140197CCBEEC4F080F659" bold="true">C E</emphasis>
terminalia in dorsal, ventral, and lateral views, respectively
<emphasis id="46630433CB9AC698B1B05E86B559132C" bold="true">F</emphasis>
spermathecae. Abbreviations: cerc = cercus; hyprct = hypoproct; lat sc = lateral sclerite of genital fork; st = sternite; tg = tergite.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E2OAG" type="type material">
<paragraph id="9945709CDB852D59198BDFADB0D64A8A">
<heading id="D713CC1E56C2EE665B43500E782189B9" reason="title">Type examined material.</heading>
</paragraph>
<paragraph id="F436453B4EEF01B6B9BB84BFA43F2161">
<materialsCitation id="61066D95C69E67D098285E9EEB8BB7A9" collectingDate="2013-12-11" collectionCode="MZUSP" country="Brazil" latitude="-21.697777" location="Trilha Fazenda Campo Florido" longLatPrecision="1" longitude="-57.76586" municipality="Porto Murtinho" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="E34D37A2590EC8EA67B4AAB677E2D458" bold="true" italics="true">
<typeStatus id="2802220BF6B8C737DC4643E2530BF68E" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="E1592267EC147DD02B9E89DC7EF213A9" count="1" type="male"></specimenCount>
(
<named-content id="1EE3EECE90D17D0FE75CFCEE0ACA29AC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="36EF2998BDAC9A062E0762682007DC43" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
) labelled: “
<collectingCountry id="7482CD8F01000337F6C1727958A16866" name="Brazil">Brasil</collectingCountry>
: MS [state of
<collectingRegion id="4D89FB7E1CABA65F55A0386F972DF062" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
]:
<collectingMunicipality id="F1ED3EFA0C2E8A2CC0E077F9F33452B0">Porto Murtinho</collectingMunicipality>
|
<location id="F43E1A8CE69FBAA4E44872DB0AB7A4C2" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:F43E1A8CE69FBAA4E44872DB0AB7A4C2" country="Brazil" latitude="-21.697777" longLatPrecision="1" longitude="-57.76586" municipality="Porto Murtinho" name="Trilha Fazenda Campo Florido" stateProvince="Mato Grosso do Sul">Trilha Fazenda Campo Florido</location>
|
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOPAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.765861,-21.697778]}">
<geoCoordinate id="2234AB6ABD517D0A6A8308148469076D" degrees="21" direction="south" minutes="41" orientation="latitude" precision="1" seconds="52, 0" value="-21.697777">21 ° 41 ' 52, 0 &quot; S</geoCoordinate>
,
<geoCoordinate id="C5C790ADCEAC982283CD6E8D080D397A" degrees="57" direction="west" minutes="45" orientation="longitude" precision="1" seconds="57, 1" value="-57.76586">57 ° 45 ' 57, 1 &quot; W</geoCoordinate>
</named-content>
| Ativa |
<date id="4E8D4102C3A95E532185D0215AC295FD" value="2013-12-11">
<collectingDate id="BA97828B15B3419A019AB4D976900283" value="2013-12-11">11. xii. 2013</collectingDate>
</date>
| Lamas &amp; eq. cols. | SISBIOTA | CNPq / FAPESP ” “
<typeStatus id="8737D37D613D761EA92D255311776C4B" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="C8CD671BD31786B645FB3BC61F8DE3F3" authorityName="Soares &amp; Camargo &amp; Lamas" authorityYear="2025" class="Insecta" family="Asilidae" genus="Cardiasilus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="aysu">
<emphasis id="72010CEFB8EE72A6A831D7F268F011B9" italics="true">Cardiasilus aysu</emphasis>
</taxonomicName>
| Soares, Camargo &amp; Lamas [red label] ”
</materialsCitation>
.
<typeStatus id="0CF554395B40FEBAFFD207729BC976C9" type="holotype">Holotype</typeStatus>
condition: Good, not dissected.
<materialsCitation id="EE1B29C5220D4E39E39B950CB4B035D7" collectingDate="2011-12-11" collectingDateMax="2012-01-25" collectingDateMin="2011-12-11" collectionCode="MZUSP, NHMW" collectorName="Lamas &amp; Nihei &amp; Fazenda Retiro Conceicao &amp; Trilha &amp; da Espinhadeira &amp; Malaise" country="Brazil" latitude="-21.685055" location="Coleta" longLatPrecision="1" longitude="-57.776585" municipality="Porto Murtinho" specimenCount="8" specimenCount-male="8" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
<emphasis id="2DE352A92657060B2A9A5021B64D2A22" bold="true" italics="true">
<typeStatus id="4183811050A45F6C7B05FB5124ADAD5F" type="paratype">Paratypes</typeStatus>
</emphasis>
: • same data as holotype (
<specimenCount id="2E37C9DE56B62B8819625C7633DD0DCD" count="2" type="male">2 ♂</specimenCount>
, one dissected,
<named-content id="CF90BF447DBF28607A746E77980353F8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="E9C2F89E9470F646C79A7295914C01CC" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
); same data, except:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.702833,-21.637519]}">
<geoCoordinate id="8490EE369F6F81F813236D8555C5C0C6" degrees="21" direction="south" minutes="38" orientation="latitude" precision="1" seconds="15, 07" value="-21.63752">21 ° 38 ' 15, 07 &quot; S</geoCoordinate>
,
<geoCoordinate id="BFC90DF3AE01AE1DC05AE7E904830C33" degrees="57" direction="west" minutes="42" orientation="longitude" precision="1" seconds="10, 2" value="-57.702835">57 ° 42 ' 10, 2 &quot; W</geoCoordinate>
</named-content>
|
<location id="A6F9BD1B44BF030B5961631EE9C061E2" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:A6F9BD1B44BF030B5961631EE9C061E2" latitude="-21.685055" longLatPrecision="1" longitude="-57.776585" name="Coleta">Coleta</location>
manual (rede) |
<date id="D268A67C19762519D13027550B556533" value="2011-12-12">
<collectingDate id="755CED3233CFFABA91B96F2740084CAC" value="2011-12-12">12. xii. 2011</collectingDate>
</date>
|
<collectorName id="4FD414B9852FE605FB275CD9D06EA10E">Lamas</collectorName>
,
<collectorName id="713DDFC056A8268A180277B9DD8360BC">Nihei</collectorName>
&amp; eq. col. (
<specimenCount id="71E13FC8462DA49E456932B5C5AF2517" count="1" type="male">1 ♂</specimenCount>
,
<named-content id="AB7E0C6B7465D729FFE79564A563C00C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="F5207E07FD1434A5600095C2BB716AEF" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
); same data, except:
<collectorName id="35BADC1392E3E41C734AFF3A186455FD">Fazenda Retiro Conceição</collectorName>
<collectorName id="15F1BFE2425F6F1D17FBE24146AA4F92">Trilha</collectorName>
|
<collectorName id="D6E77D0A0F9AC26157A8B8E2267B4126">da Espinhadeira</collectorName>
|
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.778472,-21.683250]}">
<geoCoordinate id="9CA1571792795739B0E86358610FA1BD" degrees="21" direction="south" minutes="40" orientation="latitude" precision="1" seconds="59, 7" value="-21.683249">21 ° 40 ' 59, 7 &quot; S</geoCoordinate>
,
<geoCoordinate id="6184A22E0973E433352393A94B574500" degrees="57" direction="west" minutes="46" orientation="longitude" precision="1" seconds="42, 5" value="-57.778473">57 ° 46 ' 42, 5 &quot; W</geoCoordinate>
</named-content>
|
<collectorName id="EF228C3ECDC58B011DA805506A615EC0">Malaise</collectorName>
31 |
<date id="B972CC56D44737C7DB8D050BD47D3C4F" value="2012-01-10" valueMax="2012-01-25" valueMin="2012-01-10">
<collectingDate id="2FF4D4C2C938FDD2D8895D14FE316325" value="2012-01-10" valueMax="2012-01-25" valueMin="2012-01-10">1025. i. 2012</collectingDate>
</date>
|
<collectorName id="6BD3448E18AFB7AE79FC8DFD2197E416">Lamas</collectorName>
,
<collectorName id="9F34D761B70E728304592EEF4DC9D5AA">Nihei</collectorName>
&amp; eq. col. (
<specimenCount id="6C7FEB76378290DDE4875CF429081B19" count="2" type="male">2 ♂</specimenCount>
,
<named-content id="4B01DC9F747341FA715DB0DF0D1CD6B0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="4DA8C51D505F835A765DCB6A6CC99158" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
); same data, except:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.776583,-21.685056]}">
<geoCoordinate id="CA0AB6D2421CDE946799E204288F120D" degrees="21" direction="south" minutes="41" orientation="latitude" precision="1" seconds="06.2" value="-21.685055">21 ° 41 ' 06.2 &quot; S</geoCoordinate>
,
<geoCoordinate id="628A5799EE67F36FCFCF8B577DDE3E1E" degrees="57" direction="west" minutes="46" orientation="longitude" precision="1" seconds="35.7" value="-57.776585">57 ° 46 ' 35.7 &quot; W</geoCoordinate>
</named-content>
| coleta manual (rede) |
<date id="8DD4F4BA6E39F23F30E6EB0BA67BA8D5" value="2011-12-11">
<collectingDate id="C6E93ED9346168B1EC8F5552366DC03D" value="2011-12-11">11. xii. 2011</collectingDate>
</date>
(
<specimenCount id="EC557391C2F60FCEF36BDE6AF1CC2641" count="1" type="male">1 ♂</specimenCount>
,
<named-content id="29D1B777FF7C92E007EA6D0C80CB0D91" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="740C68212F9F6ECED2073DC18D263D4B" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
;
<specimenCount id="CF94C250BB75BAB4CD699418AD726FA5" count="2" type="male">2 ♂</specimenCount>
,
<named-content id="311B973815BD471BC287EA8D0219534F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Natural History Museum Vienna">
<collectionCode id="BF1979F63A4DC56A81FC61D4ED232B55" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
; •
<materialsCitation id="494537B320CF720F1BEDC36AFC6E8B31" collectingDate="2008-01-19" collectingDateMax="2008-01-31" collectingDateMin="2008-01-19" collectionCode="R, MZUSP, NHMW" collectorName="Nihei &amp; Figueiredo &amp; Almeida" country="Brazil" county="Pirizal" latitude="-21.551388" location="Porto Murtinho" longLatPrecision="21" longitude="-57.759724" municipality="Estrada" specimenCount="4" specimenCount-male="4" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
<location id="AB541F2EDFD2330D26242C3965D84E32" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:AB541F2EDFD2330D26242C3965D84E32" country="Brazil" county="Pirizal" latitude="-21.551388" longLatPrecision="21" longitude="-57.759724" municipality="Estrada" name="Porto Murtinho" stateProvince="Mato Grosso do Sul">Porto Murtinho</location>
(norte) |
<collectingMunicipality id="165B46FDE31B3D7492BA2AC2016FA4CE">Estrada</collectingMunicipality>
para
<collectingCounty id="2FBB70B854EDC6F71D46A22CD286F317">Pirizal</collectingCounty>
, km 18 |
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.759722,-21.551389]}">
<geoCoordinate id="4BE32E16118AD262E2911FEA50F29C18" degrees="21" direction="south" minutes="33" orientation="latitude" precision="15" seconds="05" value="-21.551388">21 ° 33 ' 05 &quot; S</geoCoordinate>
,
<geoCoordinate id="274A2B3CEFA460E73E995BA0A9822C94" degrees="57" direction="west" minutes="45" orientation="longitude" precision="15" seconds="35" value="-57.759724">57 ° 45 ' 35 &quot; W</geoCoordinate>
</named-content>
|
<date id="41DE58B88B7956F71F1ADCACBF785133" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">
<collectingDate id="65CBEB38B50570CA36DFC25F94623749" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">1931. i. 2008</collectingDate>
</date>
|
<collectorName id="91E3ADAF64C2AF3A23990CA2F74BD1AD">Nihei</collectorName>
, S.;
<collectorName id="3643C551D068FE166020C31E072AF601">Figueiredo</collectorName>
,
<collectionCode id="C98ED529CC8F84353EE506797700702C" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. &amp;
<collectorName id="2828224701C3DEE625EE382634964420">Almeida</collectorName>
, J. (col.) (
<specimenCount id="F19760E44926661E6E22879BAD0FA263" count="2" type="male">2 ♂</specimenCount>
,
<named-content id="638D7BE928AA3479673CD9148324320B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="30EB4D1BBC76B99FBE71E6E2B089CA4D" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
;
<specimenCount id="5635AE238CB53C0C54077D99AACF513A" count="2" type="male">2 ♂</specimenCount>
,
<named-content id="703B36BB10B06DAE1828C4104B8A5FAC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Natural History Museum Vienna">
<collectionCode id="8F918702A4BEAC2619C087C83F330DAE" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
; •
<materialsCitation id="685B8A97E6FD84ABCB20057B05762200" collectingDate="2008-01-21" collectingDateMax="2008-01-30" collectingDateMin="2008-01-21" collectionCode="MZUSP" collectorName="F. A. Esteves" country="Brazil" county="Pirizal" latitude="-21.551388" location="Porto Murtinho" longLatPrecision="21" longitude="-57.759724" municipality="Estrada" specimenCount="3" specimenCount-male="3" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
same data, except:
<date id="7264C9B7788905DF008767AF288D724D" value="2008-01-21" valueMax="2008-01-30" valueMin="2008-01-21">
<collectingDate id="7E77B8F6785A6173CC3BC5466940C950" value="2008-01-21" valueMax="2008-01-30" valueMin="2008-01-21">2130. i. 2008</collectingDate>
</date>
(
<specimenCount id="CCC33098BCF409643BDAE6FF9EF57CBB" count="2" type="male">2 ♂</specimenCount>
, one dissected,
<named-content id="8C053C37A78986F25877D843216C6DA8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="D69611543CD8149F647FD0EFAFC1F6EF" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
); same data, except:
<date id="177D9B0B154AFF5C8E0034A02A534674" value="2008-01-24">
<collectingDate id="4CA2A9DEFF1C3B5A84CC3590F8B9E6A8" value="2008-01-24">24. i. 2008</collectingDate>
</date>
|
<collectorName id="29E56847C3ACFA0F5B55D285C4387445">F. A. Esteves</collectorName>
col. (
<specimenCount id="59A4F3B8CC02DECA2E5C2B6D65D711A2" count="1" type="male">1 ♂</specimenCount>
,
<named-content id="7988375A04A78C0C9E38F8E5BA15E243" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="1EFF270377093A3E2225F38BA64B84A9" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
)
</materialsCitation>
; •
<materialsCitation id="14169029073617F45AF3C4200C58BF7E" collectingDate="2008-01-19" collectingDateMax="2008-01-31" collectingDateMin="2008-01-19" collectionCode="R, MZUSP" collectorName="Nihei &amp; Figueiredo &amp; Almeida" country="Brazil" county="Camalotes" latitude="-21.707779" location="Faz. Sao Fernando" longLatPrecision="21" longitude="-57.583332" municipality="Arredores do Hotel" specimenCount="2" specimenCount-male="2" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
same data, except:
<location id="BCDBCC9B18C2C800DE7EB70B38E1A4DD" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:BCDBCC9B18C2C800DE7EB70B38E1A4DD" county="Camalotes" latitude="-21.707779" longLatPrecision="21" longitude="-57.583332" municipality="Arredores do Hotel" name="Faz. Sao Fernando">Faz. São Fernando</location>
, km 12 |
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERSAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.817222,-21.608333]}">
<geoCoordinate id="18592D8689EAB98CEA5DF976B9C9C152" degrees="21" direction="south" minutes="36" orientation="latitude" precision="15" seconds="30" value="-21.608334">21 ° 36 ' 30 &quot; S</geoCoordinate>
,
<geoCoordinate id="9CE1247AD299B835F02AF838EFB1AB15" degrees="57" direction="west" minutes="49" orientation="longitude" precision="15" seconds="02" value="-57.817223">57 ° 49 ' 02 &quot; W</geoCoordinate>
</named-content>
|
<date id="83E0A447CC6CE18A0DC4ADBA79B61C69" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">
<collectingDate id="C52AE0B3D50BD0D94F137D5C1C7F4B50" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">1931. i. 2008</collectingDate>
</date>
|
<collectorName id="56ED2081EF8A07591C2CBC00162BD5A8">Nihei</collectorName>
, S.;
<collectorName id="512BEF9A553D7619D162F55EFBCCE913">Figueiredo</collectorName>
,
<collectionCode id="2627784C427C8182F1E1B8BEE23572E7" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. &amp;
<collectorName id="E5B21692C18264D659C06AC3D09B936C">Almeida</collectorName>
, J. | (col.) (
<specimenCount id="1FCABA027E4582328D3AAB4A462F9FC3" count="1" type="male">1 ♂</specimenCount>
,
<named-content id="9EA19B5F89D2883D7A24B4B51BBC2105" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="98A6C0597CDC6CD55CB2D6833E61C4E3" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
); same data, except:
<collectingMunicipality id="87F5E773B1A28CD699FCBB843D539493">Arredores do Hotel</collectingMunicipality>
dos
<collectingCounty id="968A420FEFA36FF796D8627A61C4E553">Camalotes</collectingCounty>
|
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5SAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.583333,-21.707778]}">
<geoCoordinate id="C5FEC8602F178C22182A2732CE673829" degrees="21" direction="south" minutes="42" orientation="latitude" precision="15" seconds="28" value="-21.707779">21 ° 42 ' 28 &quot; S</geoCoordinate>
,
<geoCoordinate id="319AF038D4D91CBDB541A5BA1FB2DA8D" degrees="57" direction="west" minutes="35" orientation="longitude" precision="15" seconds="00" value="-57.583332">57 ° 35 ' 00 &quot; W</geoCoordinate>
</named-content>
|
<date id="9CE488C6E57E3388578C6D54D2E35957" value="2008-01-21" valueMax="2008-01-30" valueMin="2008-01-21">
<collectingDate id="C1437F1F406210C7E2C6ABB6B66631A2" value="2008-01-21" valueMax="2008-01-30" valueMin="2008-01-21">2130. i. 2008</collectingDate>
</date>
(
<specimenCount id="466063E50A540C8AF8EF9723C2BE64D8" count="1" type="male">1 ♂</specimenCount>
,
<named-content id="346527D70C65124A74F0227372666D92" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="DBA8F6700D160F9F6E6BB11B68F4D2A3" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EITAG" type="material">
<paragraph id="989FC7F48E65A67F0590EBDAB1E28651">
<heading id="EB6F1A837A7FCA6B2565DC8EB2121FA6" reason="title">Additional examined material.</heading>
</paragraph>
<paragraph id="B2C2757F32F59E485FC18A8EB87909FD">
<materialsCitation id="BDABB7452015F13882D75D1077E09624" collectingDate="2008-01-19" collectingDateMax="2008-01-31" collectingDateMin="2008-01-19" collectorName="Nihei, S" country="Brazil" county="Porto Murtinho" latitude="-21.551388" location="Pirizal" longLatPrecision="21" longitude="-57.759724" municipality="Estrada" specimenCount="1" stateProvince="Mato Grosso do Sul">
<emphasis id="B7D391190BEBBF5CF7A7F028CD34CECD" bold="true">
<collectingCountry id="001ECCBEA8946BD38531F64E386CB747" name="Brazil">Brazil</collectingCountry>
</emphasis>
, MS [state of
<collectingRegion id="2DDF3C13D8B6D5633C88778166C0F2E5" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
]:
<collectingCounty id="43EBFB6C4F560405DFE6289F1E8BD646">Porto Murtinho</collectingCounty>
(norte),
<collectingMunicipality id="242945D7DFC2374B731D074C6378F6CD">Estrada</collectingMunicipality>
para
<location id="2968147D9898D095400963DD9F695BEB" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:2968147D9898D095400963DD9F695BEB" country="Brazil" county="Porto Murtinho" latitude="-21.551388" longLatPrecision="21" longitude="-57.759724" municipality="Estrada" name="Pirizal" stateProvince="Mato Grosso do Sul">Pirizal</location>
, km 18,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETTAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.759722,-21.551389]}">
<geoCoordinate id="C3724B46089C428473707B39DA4F4C6F" degrees="21" direction="south" minutes="33" orientation="latitude" precision="15" seconds="05" value="-21.551388">21 ° 33 ' 05 &quot; S</geoCoordinate>
,
<geoCoordinate id="E34DE7B85C1140C8C9B31E29E1EB8616" degrees="57" direction="west" minutes="45" orientation="longitude" precision="15" seconds="35" value="-57.759724">57 ° 45 ' 35 &quot; W</geoCoordinate>
</named-content>
,
<date id="AA856D69E5206EABDE9225B6C58175D0" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">
<collectingDate id="0CA967F4CC6BE8B1CE44639D732C094F" value="2008-01-19" valueMax="2008-01-31" valueMin="2008-01-19">1931. i. 2008</collectingDate>
</date>
,
<collectorName id="78B7286D31B61ED0EFECBE9CC6CC3B13">Nihei, S</collectorName>
</materialsCitation>
.; •
<materialsCitation id="957E8CEE6F80D71EC248D408D0977B45" collectingDate="2013-12-11" collectingDateMax="2008-01-31" collectingDateMin="2008-01-19" collectionCode="R, MZUSP, NHMW" collectorName="Almeida, J." country="Brazil" county="Porto Murtinho" latitude="-21.697777" location="Trilha Fazenda Campo Florido" longLatPrecision="1" longitude="-57.76586" municipality="Figueiredo" specimenCount="8" specimenCount-female="8" stateProvince="Mato Grosso do Sul">
<collectingMunicipality id="E4A00D8F656329F28898655D060BFA01">Figueiredo</collectingMunicipality>
,
<collectionCode id="1AE1BDE646CC0322F8C8E33E283C41EE" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. &amp;
<collectorName id="D3EE3001696C5120B9AE6EED349918C7">Almeida, J.</collectorName>
(col.) (
<specimenCount id="47B5556A3DA3A3B93DD6BB7663732942" count="2" type="female">2 ♀</specimenCount>
,
<named-content id="C4FEA3EF3141658A71CE869A66B3156E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="C0EED220AC217A37B254F85B6F6102F7" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
;
<specimenCount id="203BC63535BA83D240DC023D3A8ACB0D" count="2" type="female">2 ♀</specimenCount>
,
<named-content id="4D1BE82304F97A6EBE955CCD25F6578D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Natural History Museum Vienna">
<collectionCode id="1B84AC5D5414353FF774357921DC640F" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
); same data, except:
<location id="FBC177C1A3B0D04EDB750FFE9B1FAA2E" LSID="urn:lsid:plazi:treatment:DC34FA99F5D658A59A0373012AB2BA86:FBC177C1A3B0D04EDB750FFE9B1FAA2E" country="Brazil" latitude="-21.697777" longLatPrecision="1" longitude="-57.76586" municipality="Figueiredo" name="Trilha Fazenda Campo Florido">Trilha Fazenda Campo Florido</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFUAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-57.765861,-21.697778]}">
<geoCoordinate id="90AB9EEAE8B62D4E2FB97E82AE776910" degrees="21" direction="south" minutes="41" orientation="latitude" precision="1" seconds="52, 0" value="-21.697777">21 ° 41 ' 52, 0 &quot; S</geoCoordinate>
,
<geoCoordinate id="777388EA134430747771CE05F83646BE" degrees="57" direction="west" minutes="45" orientation="longitude" precision="1" seconds="57, 1" value="-57.76586">57 ° 45 ' 57, 1 &quot; W</geoCoordinate>
</named-content>
, Ativa,
<date id="54E9427241CC37B4B71DCC1D2EB5C5DA" value="2013-12-11">
<collectingDate id="18B20C58ED9A5C03780DB663F343CCB7" value="2013-12-11">11. xii. 2013</collectingDate>
</date>
, Lamas &amp; eq. cols. (
<specimenCount id="01C3A8A39C4242D8DE93BB0FE8F7E5DD" count="4" type="female">4 ♀</specimenCount>
, two dissected,
<named-content id="A73490FBDBCFBD9ADAEE7E7A793BD36A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<collectionCode id="0F3E84E5C1BB9C9992656178BB2A08CB" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPUAG" type="remarks">
<paragraph id="9C8DE58373D7FD2613DE9C2A3458E2F9">
<heading id="C569C4CBB6D2567F98A11671EE466012" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="BCC444E9CE3955E8D0F4EE57D7D1DEA2">The new species is easily recognized by the characters presented in the key and diagnosis (see above).</paragraph>
</subSubSection>
<subSubSection id="SECID0EUUAG" type="distribution">
<paragraph id="1275FAA82A95CDBE573FE5A04406E48D">
<heading id="638C10C35DC51D7223D1A3F98DB08311" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0F551D73A0E1511AC2278C72E0B4AAFC">
<collectingCountry id="A8F0048CB3BC194049216AEC786509DF" name="Brazil">Brazil</collectingCountry>
(state of
<collectingRegion id="855C29259461275E98B59C45DE91923A" country="Brazil" name="Mato Grosso do Sul">Mato Grosso do Sul</collectingRegion>
) (Fig.
<figureCitation id="C0CD0E7563A0FE848EFBFE08C93CE84E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Known geographical distribution of Cardiasilus gen. nov. species." figureDoi="10.3897/zookeys.1232.142494.figure19" httpUri="https://binary.pensoft.net/fig/1286753">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5UAG" type="etymology">
<paragraph id="6214AE925B2D9A5BE52FB6E15F1AA190">
<heading id="57C21B3BE372ED01D6D9BCBBC288871B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E7D220B291F8D7F3E0293CA1A6CC999D">
From the Tupi-guarani
<emphasis id="2C38709FF02EF361A0A58BB2C24C1706" italics="true">aysú</emphasis>
= love, referring to the male terminalia, which resembles an ideogram of a heart in dorsal view. The species name is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>