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<document id="D96DFA9B3C40FFC5AFA089B4416DEE27" ID-DOI="10.1186/s13358-024-00317-6" ID-ISSN="1664-2384" ID-Zenodo-Dep="12005564" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1718654929440" checkinUser="felipe" docAuthor="Montagna, Matteo, Magoga, Giulia &amp; Magnani, Fabio" docDate="2024" docId="03F4A916C864D005C4A5FB3CFBB75E6D" docLanguage="en" docName="SwissJPalaeontol.143.17.pdf" docOrigin="Swiss Journal of Palaeontology (17) 143 (1)" docSource="http://dx.doi.org/10.1186/s13358-024-00317-6" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Merithone Montagna &amp; Magoga 2024, gen. nov." docType="treatment" docVersion="1" lastPageNumber="6" masterDocId="FFCDD16EC861D000C408FFF1FF915D68" masterDocTitle="The Middle Triassic palaeontomofauna of Monte San Giorgio with the description of Merithone laetitiae († Permithonidae) gen. et sp. nov." masterLastPageNumber="10" masterPageNumber="1" pageNumber="6" updateTime="1721678352683" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="830A2A5DCD0711EA14883945769B1D7D">The Middle Triassic palaeontomofauna of Monte San Giorgio with the description of Merithone laetitiae († Permithonidae) gen. et sp. nov.</mods:title>
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<mods:namePart id="C7E1DF7F5823B900C355F1F42D297C73">Montagna, Matteo</mods:namePart>
<mods:affiliation id="CD552164EF6FCCF10E7B0AE77837924C">Department of Agricultural Sciences, University of Naples Federico II, Portici, Italy &amp; Interuniversity Center for Studies on Bioinspired Agro-Environmental Technology (BAT Center), University of Naples Federico II, Portici, Italy</mods:affiliation>
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<mods:namePart id="E81BA48F48A8B56F07B92921ADD2AC53">Magoga, Giulia</mods:namePart>
<mods:affiliation id="B6F762AB3801712B211ADE59B902CA65">Department of Agricultural Sciences, University of Naples Federico II, Portici, Italy</mods:affiliation>
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<mods:namePart id="5937BB66E5521403DB94928D18B56814">Magnani, Fabio</mods:namePart>
<mods:affiliation id="9425849F63BC1C8BD86013E775D43B06">Museo Cantonale di Storia Naturale, Lugano, Switzerland</mods:affiliation>
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<mods:title id="011047882B9943DF684EBE35DC081829">Swiss Journal of Palaeontology</mods:title>
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<mods:part id="C483AE895A3C26C665BDFE430695DC11">
<mods:date id="56E9D3C40BBD8F041B43327DD67F8133">2024</mods:date>
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<mods:number id="DD10DA4E11B8B412B848D6510A6BAB4A">2024-04-29</mods:number>
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<mods:detail id="A369064DD24DEDA217AC21622BCBC36D" type="volume">
<mods:number id="E45DF302E71E31B776C418485474B972">143</mods:number>
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<mods:identifier id="BA298F969D7E414F169481A93A8416AD" type="ISSN">1664-2384</mods:identifier>
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<treatment id="03F4A916C864D005C4A5FB3CFBB75E6D" LSID="urn:lsid:plazi:treatment:03F4A916C864D005C4A5FB3CFBB75E6D" httpUri="http://treatment.plazi.org/id/03F4A916C864D005C4A5FB3CFBB75E6D" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="C3474B8BC864D005C4A5FB3CFD55598D" box="[173,708,1229,1254]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BE21800C864D005C4A5FB3CFD55598D" blockId="5.[151,775,1005,1925]" box="[173,708,1229,1254]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="4C5D6383C864D005C4F1FB3CFDCB598E" authority="Montagna &amp; Magoga" authorityName="Montagna &amp; Magoga" authorityYear="2024" box="[249,602,1229,1254]" class="Insecta" family="Permithonidae" genus="Merithone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B929C412C864D005C4F1FB3CFEF8598D" box="[249,361,1229,1253]" italics="true" pageId="5" pageNumber="6">Merithone</emphasis>
Montagna &amp; Magoga
</taxonomicName>
,
<taxonomicNameLabel id="A21A7969C864D005C66CFB3FFD55598D" box="[612,708,1230,1254]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C4A5FB1FFF78584D" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="8BE21800C864D005C4A5FB1FFF78584D" blockId="5.[151,775,1005,1925]" pageId="5" pageNumber="6">
<typeStatus id="54E6A6A2C864D005C4A5FB1FFF72586E" box="[173,227,1262,1286]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="4C5D6383C864D005C54DFB1CFD86586D" authority="Montagna &amp; Magoga &amp; Magnani, 2024" authorityName="Montagna &amp; Magoga &amp; Magnani" authorityYear="2024" box="[325,535,1261,1285]" class="Insecta" family="Permithonidae" genus="Merithone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="laetitiae" status="sp. nov.">
<emphasis id="B929C412C864D005C54DFB1CFD86586D" box="[325,535,1261,1285]" italics="true" pageId="5" pageNumber="6">Merithone laetitiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A7969C864D005C616FB1FFDE0586D" box="[542,625,1262,1285]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
by monotypy (
<figureCitation id="13660485C864D005C497FAFCFF4B584D" box="[159,218,1293,1317]" captionStart="Fig" captionStartId="6.[167,198,1611,1631]" captionTargetBox="[211,1377,236,1597]" captionTargetId="figure-144@6.[211,1377,236,1597]" captionTargetPageId="6" captionText="Fig. 3 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. Scale bar, 1 mm" figureDoi="http://doi.org/10.5281/zenodo.12005584" httpUri="https://zenodo.org/record/12005584/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C4A5FADCFD1F580D" pageId="5" pageNumber="6" type="description">
<paragraph id="8BE21800C864D005C4A5FADCFD1F580D" blockId="5.[151,775,1005,1925]" pageId="5" pageNumber="6">
Code Zoobank: LSID
<uri id="FFCC1402C864D005C5F1FADCFD1F580D" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="FFFB22D5C864D005C49FFABCFD1F580D" box="[151,654,1357,1381]" pageId="5" pageNumber="6">16C0A670-1BE5-44FD-BC6C-0E10C87C3045</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C4A5FA9CFDEA588D" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BE21800C864D005C4A5FA9CFDEA588D" blockId="5.[151,775,1005,1925]" pageId="5" pageNumber="6">
Etymology:
<taxonomicName id="4C5D6383C864D005C538FA9FFEF558ED" authorityName="Rheims &amp; Jäger" authorityYear="2022" box="[304,356,1390,1413]" class="Arachnida" family="Sparassidae" genus="Meri" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C538FA9FFEF558ED" box="[304,356,1390,1413]" italics="true" pageId="5" pageNumber="6">Meri</emphasis>
</taxonomicName>
- named after Meride, the village near the locality where the fossil has been collected; the suffix -
<emphasis id="B929C412C864D005C4A9FA5CFF4C58AD" box="[161,221,1453,1477]" italics="true" pageId="5" pageNumber="6">thone</emphasis>
from the
<typeStatus id="54E6A6A2C864D005C541FA5FFEE958AE" box="[329,376,1454,1478]" pageId="5" pageNumber="6">type</typeStatus>
genus of the family †
<taxonomicName id="4C5D6383C864D005C657FA5CFC9258AD" authorityName="Tillyard" authorityYear="1922" box="[607,771,1453,1477]" class="Insecta" family="Permithonidae" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Permithonidae</taxonomicName>
,
<taxonomicName id="4C5D6383C864D005C49FFA3CFE3E588D" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[151,431,1485,1509]" class="Insecta" family="Permithonidae" genus="Permithone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C49FFA3CFE85588D" box="[151,276,1485,1509]" italics="true" pageId="5" pageNumber="6">Permithone</emphasis>
<bibRefCitation id="EFCC65F1C864D005C512FA3CFE3E588D" author="Tillyard, R. J." box="[282,431,1485,1509]" pageId="5" pageNumber="6" pagination="279 - 292" refId="ref6763" refString="Tillyard, R. J. (1922). Some new Permian insects from Belmont, N. S. W. in the collection of Mr. John Mitchell. Proceedings of the Linnean Society of New South Wales, 47, 279 - 292." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C4A4FA1CFBB75E6D" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BE21800C864D005C4A4FA1CFD665AED" blockId="5.[151,775,1005,1925]" pageId="5" pageNumber="6">
Diagnosis: the general habitus of the new taxon (new genus, new species) comprises a well-developed prothorax, shorter than meso- plus meta- thorax; the prognathous head capsule has prominent compound eyes and filiform antennae; the subequal membranous wings without pterostigma; the fore wings with broad costal area possessing several irregular veins from subcostal vein (SC) to costal margin (some distally bifurcated); the SC merging distally the radius (R), the radial sector (RS) with six primary branches; the medial anterior (MA) not coalesced with RS basally; the CUP apparently not deeply forked; and nygmata not visible prompts the ascription of the fossil to the family of †
<taxonomicName id="4C5D6383C864D005C5BBF89CFD615AED" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[435,752,1901,1925]" class="Insecta" family="Permithonidae" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">
Permithonidae
<bibRefCitation id="EFCC65F1C864D005C654F89CFD615AED" author="Tillyard, R. J." box="[604,752,1901,1925]" pageId="5" pageNumber="6" pagination="279 - 292" refId="ref6763" refString="Tillyard, R. J. (1922). Some new Permian insects from Belmont, N. S. W. in the collection of Mr. John Mitchell. Proceedings of the Linnean Society of New South Wales, 47, 279 - 292." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE21800C864D005C74AFF1CFBB75E6D" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
Comparison: The new genus differs from the previously described genera based on forewing shape and features. The forewings are elongate, more than two times (~ 2.7) as long as wide, not broad and markedly triangular as in
<taxonomicName id="4C5D6383C864D005C725FE9CFBCD5CED" authority="Martynov, 1933" authorityName="Martynov" authorityYear="1933" box="[813,1116,365,390]" class="Insecta" family="Permithonidae" genus="Eopsychops" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C725FE9CFC395CED" box="[813,936,365,389]" italics="true" pageId="5" pageNumber="6">Eopsychops</emphasis>
<bibRefCitation id="EFCC65F1C864D005C7B8FE9FFBCD5CED" author="Martynov, A. V." box="[944,1116,365,390]" pageId="5" pageNumber="6" pagination="23 - 62" refId="ref5581" refString="Martynov, A. V. (1933). Permian fossil insects from the Arkangelsk District. Part I. The order Mecoptera. Trudy Paleozoologicheskogo Instituta Akademii Nauk SSSR, 2, 23 - 62." type="journal article" year="1933">Martynov, 1933</bibRefCitation>
</taxonomicName>
,
<emphasis id="B929C412C864D005C063FE9CFB7E5CED" box="[1131,1263,365,389]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C5D6383C864D005C063FE9CFB7B5CED" authorityName="Tillyard" authorityYear="1922" box="[1131,1258,365,389]" class="Insecta" family="Permithonidae" genus="Permithone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Permithone</taxonomicName>
,
</emphasis>
<taxonomicName id="4C5D6383C864D005C0FFFE9CFC545CCD" authority="Tillyard, 1926" authorityName="Tillyard" authorityYear="1926" class="Insecta" family="Permithonidae" genus="Permopsychops" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C0FFFE9CFA0D5CED" box="[1271,1436,365,389]" italics="true" pageId="5" pageNumber="6">Permopsychops</emphasis>
<bibRefCitation id="EFCC65F1C864D005C725FE7CFC545CCD" author="Tillyard, R. J." box="[813,965,397,421]" pageId="5" pageNumber="6" pagination="265 - 282" refId="ref6811" refString="Tillyard, R. J. (1926). Upper Permian insects of New South Wales. II. The orders Mecoptera, Paramecoptera and Neuroptera. Proceedings of the Linnean Society of New South Wales, 51, 265 - 282." type="journal article" year="1926">Tillyard, 1926</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5D6383C864D005C001FE7CFCF05CAD" authority="Pinto &amp; de Ornellas, 1980" authorityName="Pinto &amp; de Ornellas" authorityYear="1980" class="Insecta" family="Permithonidae" genus="Permipsythone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C001FE7CFB385CCD" box="[1033,1193,397,421]" italics="true" pageId="5" pageNumber="6">Permipsythone</emphasis>
<bibRefCitation id="EFCC65F1C864D005C0BAFE7CFCF05CAD" author="Pinto, I. D. &amp; de Ornellas, L. P." pageId="5" pageNumber="6" pagination="155 - 159" refId="ref6087" refString="Pinto, I. D., &amp; de Ornellas, L. P. (1980). Permian Insects from the Parana Basin, South Brasil II Neuroptera. Pesquisas, 13, 155 - 159." type="journal article" year="1980">Pinto &amp; de Ornellas, 1980</bibRefCitation>
</taxonomicName>
, not oval as in
<taxonomicName id="4C5D6383C864D005C000FE5CFAF75CAD" authority="Prokop et al., 2015" authorityName="Prokop" authorityYear="2015" box="[1032,1382,429,453]" class="Insecta" family="Permithonidae" genus="Lodevosisyra" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C000FE5CFB025CAD" box="[1032,1171,429,453]" italics="true" pageId="5" pageNumber="6">Lodevosisyra</emphasis>
<bibRefCitation id="EFCC65F1C864D005C092FE5CFAF75CAD" author="Prokop, J. &amp; Rodrigues Fernandes, F. &amp; Lapeyrie, J. &amp; Nel, A." box="[1178,1382,429,453]" pageId="5" pageNumber="6" pagination="263 - 270" refId="ref6206" refString="Prokop, J., Rodrigues Fernandes, F., Lapeyrie, J., &amp; Nel, A. (2015). Discovery of the first lacewings (Neuroptera: Permithonidae) from the Guadalupian of the Lodeve Basin (Southern France). Geobios, 48, 263 - 270." type="journal article" year="2015">Prokop et al., 2015</bibRefCitation>
</taxonomicName>
. The costal area is wider in its first half, slightly restricted at the base, but not as strongly as in
<taxonomicName id="4C5D6383C864D005C0B3FE1CFC3E5F4D" authority="Martynov, 1928" authorityName="Martynov" authorityYear="1928" class="Insecta" family="Permithonidae" genus="Palaemerobius" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C0B3FE1CFACA5F6D" box="[1211,1371,493,517]" italics="true" pageId="5" pageNumber="6">Palaemerobius</emphasis>
<bibRefCitation id="EFCC65F1C864D005C16DFE1FFC3E5F4D" author="Martynov, A. V." pageId="5" pageNumber="6" pagination="1 - 118" refId="ref5499" refString="Martynov, A. V. (1928). Permian fossil Insects of North-East Europe. Travaux du Musee Geologique pres l'academie des Sciences de l'URSS, 4, 1 - 118." type="journal article" year="1928">Martynov, 1928</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D6383C864D005C7C9FDFCFA8C5F4D" authority="Martynov, 1931" authorityName="Martynov" authorityYear="1931" box="[961,1309,525,550]" class="Insecta" family="Permithonidae" genus="Permegalomus" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C7C9FDFCFBF15F4D" box="[961,1120,525,549]" italics="true" pageId="5" pageNumber="6">Permegalomus</emphasis>
<bibRefCitation id="EFCC65F1C864D005C064FDFFFA8C5F4D" author="Martynov, A. V." box="[1132,1309,525,550]" pageId="5" pageNumber="6" pagination="149 - 212" refId="ref5533" refString="Martynov, A. V. (1931). O novykh iskopaemykh nasekomykh Tikhikh Gor, Otdel Neoptera, {II New fossil insects from Tikhie Gory, Division Neoptera II}. Trudy Geologicheskogo Muzeya, Akademii Nauk SSSR, 8, 149 - 212." type="journal article" year="1931">Martynov, 1931</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D6383C864D005C127FDFFFCF85F2D" authorityName="Pinto &amp; de Ornellas" authorityYear="1980" class="Insecta" family="Permithonidae" genus="Permipsythone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C127FDFFFCF85F2D" italics="true" pageId="5" pageNumber="6">Permipsythone</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D6383C864D005C7A0FDDCFA9B5F2D" authority="Martynov, 1933" authorityName="Martynov" authorityYear="1933" box="[936,1290,557,582]" class="Insecta" family="Permithonidae" genus="Permithonopsis" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C7A0FDDCFBDE5F2D" box="[936,1103,557,581]" italics="true" pageId="5" pageNumber="6">Permithonopsis</emphasis>
<bibRefCitation id="EFCC65F1C864D005C052FDDFFA9B5F2D" author="Martynov, A. V." box="[1114,1290,557,582]" pageId="5" pageNumber="6" pagination="23 - 62" refId="ref5581" refString="Martynov, A. V. (1933). Permian fossil insects from the Arkangelsk District. Part I. The order Mecoptera. Trudy Paleozoologicheskogo Instituta Akademii Nauk SSSR, 2, 23 - 62." type="journal article" year="1933">Martynov, 1933</bibRefCitation>
</taxonomicName>
; veinlets are present and branched in the proximal part of the wing, but not reticulated as in
<taxonomicName id="4C5D6383C864D005C059FD9FFA075FED" authority="Tillyard, 1926" authorityName="Tillyard" authorityYear="1926" box="[1105,1430,621,645]" class="Insecta" family="Permithonidae" genus="Permorapisma" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C059FD9FFB605FED" box="[1105,1265,622,645]" italics="true" pageId="5" pageNumber="6">Permorapisma</emphasis>
<bibRefCitation id="EFCC65F1C864D005C0F4FD9CFA075FED" author="Tillyard, R. J." box="[1276,1430,621,645]" pageId="5" pageNumber="6" pagination="265 - 282" refId="ref6811" refString="Tillyard, R. J. (1926). Upper Permian insects of New South Wales. II. The orders Mecoptera, Paramecoptera and Neuroptera. Proceedings of the Linnean Society of New South Wales, 51, 265 - 282." type="journal article" year="1926">Tillyard, 1926</bibRefCitation>
</taxonomicName>
. The new genus is close to
<taxonomicName id="4C5D6383C864D005C063FD7CFCF05FAD" authority="Prokop et al., 2015" authorityName="Prokop" authorityYear="2015" class="Insecta" family="Permithonidae" genus="Lodevothone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B929C412C864D005C063FD7CFB655FCD" box="[1131,1268,653,677]" italics="true" pageId="5" pageNumber="6">Lodevothone</emphasis>
<bibRefCitation id="EFCC65F1C864D005C109FD7CFCF05FAD" author="Prokop, J. &amp; Rodrigues Fernandes, F. &amp; Lapeyrie, J. &amp; Nel, A." pageId="5" pageNumber="6" pagination="263 - 270" refId="ref6206" refString="Prokop, J., Rodrigues Fernandes, F., Lapeyrie, J., &amp; Nel, A. (2015). Discovery of the first lacewings (Neuroptera: Permithonidae) from the Guadalupian of the Lodeve Basin (Southern France). Geobios, 48, 263 - 270." type="journal article" year="2015">Prokop et al., 2015</bibRefCitation>
</taxonomicName>
, from which it differs in the presence of branched veinlets and less than seven Radial posterior branches, and to
<emphasis id="B929C412C864D005C770FD1CFBB75E6D" box="[888,1062,749,773]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C5D6383C864D005C770FD1CFBB35E6D" authorityName="Martynov" authorityYear="1933" box="[888,1058,749,773]" class="Insecta" family="Permithonidae" genus="Permithonopsis" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Permithonopsis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

229
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<mods:title id="D502068F2937CBAF39BA290A69CFC85D">The Middle Triassic palaeontomofauna of Monte San Giorgio with the description of Merithone laetitiae († Permithonidae) gen. et sp. nov.</mods:title>
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<mods:namePart id="48AF4A9F45EBB106CE97D39B65EE2FDF">Montagna, Matteo</mods:namePart>
<mods:affiliation id="0B5FC35C40868D2170E8B3E9EE98AB0C">Department of Agricultural Sciences, University of Naples Federico II, Portici, Italy &amp; Interuniversity Center for Studies on Bioinspired Agro-Environmental Technology (BAT Center), University of Naples Federico II, Portici, Italy</mods:affiliation>
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<paragraph id="8BE21800C864D005C74AFCFCFAC65E4E" blockId="5.[813,1437,237,1925]" box="[834,1367,781,806]" pageId="5" pageNumber="6">
<taxonomicName id="4C5D6383C864D005C74AFCFCFB835E4D" authority="Montagna &amp; Magoga &amp; Magnani, 2024" authorityName="Montagna &amp; Magoga &amp; Magnani" authorityYear="2024" box="[834,1042,781,805]" class="Insecta" family="Permithonidae" genus="Merithone" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="laetitiae" status="sp. nov.">
<emphasis id="B929C412C864D005C74AFCFCFB835E4D" box="[834,1042,781,805]" italics="true" pageId="5" pageNumber="6">Merithone laetitiae</emphasis>
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<taxonomicNameLabel id="A21A7969C864D005C011FCFFFBFA5E4D" box="[1049,1131,782,805]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
Montagna &amp; Magoga
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<subSubSection id="C3474B8BC864D005C74AFCDCFAB35E0D" pageId="5" pageNumber="6" type="description">
<paragraph id="8BE21800C864D005C74AFCDCFAB35E0D" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
Code Zoobank: LSID
<uri id="FFCC1402C864D005C086FCDCFAB35E0D" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="FFFB22D5C864D005C725FCBCFAB35E0D" box="[813,1314,845,869]" pageId="5" pageNumber="6">968E793A-F8F8-4B60-ADFE-E0B724A7F1EA</uuid>
</uri>
</paragraph>
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<subSubSection id="C3474B8BC864D005C74AFC9CFB985EAD" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BE21800C864D005C74AFC9CFB985EAD" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
Etimology:
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<emphasis id="B929C412C864D005C7CBFC9CFB8C5EED" box="[963,1053,877,901]" italics="true" pageId="5" pageNumber="6">laetitiae</emphasis>
</taxonomicName>
is derived from the prosoponym Laetitia, the daughter of the first author, to whom the species is dedicated.
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C74AFC3CFAB9596D" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BE21800C864D005C74AFC3CFAB9596D" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
<materialsCitation id="3B35125DC864D005C74AFC3CFAB5596D" country="Switzerland" location="Museo Cantonale di Storia Naturale" municipality="Lugano" pageId="5" pageNumber="6" specimenCount="1" typeStatus="holotype">
Material:
<typeStatus id="54E6A6A2C864D005C7A3FC3CFB9A5E8D" box="[939,1035,973,997]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
specimen MCSN 8679,
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,
<collectingMunicipality id="6B86827AC864D005C04BFC1FFB06596E" box="[1091,1175,1006,1030]" pageId="5" pageNumber="6">Lugano</collectingMunicipality>
,
<collectingCountry id="F34A5890C864D005C0A9FC1CFAB5596D" box="[1185,1316,1005,1029]" name="Switzerland" pageId="5" pageNumber="6">Switzerland</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C74AFBFFFC2D59CD" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BE21800C864D005C74AFBFFFBE5592D" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
<typeStatus id="54E6A6A2C864D005C74AFBFFFCE9594E" box="[834,888,1038,1062]" pageId="5" pageNumber="6">Type</typeStatus>
horizon:
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member of
<collectingMunicipality id="6B86827AC864D005C725FBDCFBE1592D" box="[813,1136,1069,1093]" pageId="5" pageNumber="6">Meride Limestone Formation</collectingMunicipality>
.
</paragraph>
<paragraph id="8BE21800C864D005C74AFBBFFC2D59CD" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
<typeStatus id="54E6A6A2C864D005C74AFBBFFCE9590E" box="[834,888,1102,1126]" pageId="5" pageNumber="6">Type</typeStatus>
locality: VM12 site (
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45° 53
<emphasis id="B929C412C864D005C0BDFBBDFB51590D" box="[1205,1216,1100,1125]" italics="true" pageId="5" pageNumber="6">′</emphasis>
26
<emphasis id="B929C412C864D005C0EDFBBDFB62590D" box="[1253,1267,1100,1125]" italics="true" pageId="5" pageNumber="6">ʺ</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE697EC7C864D005C12BFBBCFCAF59ED" degrees="8" direction="east" minutes="56" orientation="longitude" pageId="5" pageNumber="6" precision="15" seconds="49" value="8.946944">
8° 56
<emphasis id="B929C412C864D005C155FBBDFAF9590D" box="[1373,1384,1100,1125]" italics="true" pageId="5" pageNumber="6">′</emphasis>
49
<emphasis id="B929C412C864D005C186FBBDFA0D590D" box="[1422,1436,1100,1125]" italics="true" pageId="5" pageNumber="6">ʺ</emphasis>
E
</geoCoordinate>
),
<location id="8E824EDBC864D005C75EFB9CFB9A59ED" LSID="urn:lsid:plazi:treatment:03F4A916C864D007C74AFCFCFE8E5EC2:8E824EDBC864D005C75EFB9CFB9A59ED" box="[854,1035,1133,1157]" country="Switzerland" county="Monte San Giorgio" latitude="45.890556" longLatPrecision="18" longitude="8.946944" municipality="Meride" name="Val Mara valley" pageId="5" pageNumber="6">Val Mara valley</location>
near
<collectingMunicipality id="6B86827AC864D005C05DFB9CFB3559ED" box="[1109,1188,1133,1157]" pageId="5" pageNumber="6">Meride</collectingMunicipality>
(
<collectingCounty id="6283608CC864D005C0B0FB9CFA0959ED" box="[1208,1432,1133,1157]" pageId="5" pageNumber="6">Monte San Giorgio</collectingCounty>
,
<collectingCountry id="F34A5890C864D005C725FB7CFC2059CD" box="[813,945,1165,1189]" name="Switzerland" pageId="5" pageNumber="6">Switzerland</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D005C74AFB5CFAF759AD" box="[834,1382,1197,1222]" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BE21800C864D005C74AFB5CFAF759AD" blockId="5.[813,1437,237,1925]" box="[834,1382,1197,1222]" pageId="5" pageNumber="6">Diagnosis: as for the genus, being the only species.</paragraph>
</subSubSection>
<subSubSection id="C3474B8BC864D007C74AFB3CFE8E5EC2" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="8BE21800C864D005C74AFB3CFC495A6D" blockId="5.[813,1437,237,1925]" pageId="5" pageNumber="6">
Description (
<figureCitation id="13660485C864D005C7D1FB3CFBB9598D" box="[985,1064,1229,1253]" captionStart="Fig" captionStartId="6.[167,198,1611,1631]" captionTargetBox="[211,1377,236,1597]" captionTargetId="figure-144@6.[211,1377,236,1597]" captionTargetPageId="6" captionText="Fig. 3 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. Scale bar, 1 mm" figureDoi="http://doi.org/10.5281/zenodo.12005584" httpUri="https://zenodo.org/record/12005584/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
,
<figureCitation id="13660485C864D005C035FB3CFBDB598D" box="[1085,1098,1229,1253]" captionStart="Fig" captionStartId="7.[167,198,665,685]" captionTargetBox="[176,751,236,653]" captionTargetId="figure-348@7.[176,751,236,653]" captionTargetPageId="7" captionText="Fig. 4 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. head and thorax details. Scale bar, 500 µm" figureDoi="http://doi.org/10.5281/zenodo.12005598" httpUri="https://zenodo.org/record/12005598/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
,
<figureCitation id="13660485C864D005C057FB3CFBFD598D" box="[1119,1132,1229,1253]" captionStart="Fig" captionStartId="7.[167,198,1817,1837]" captionTargetBox="[303,1285,1311,1801]" captionTargetId="figure-367@7.[299,1289,1307,1804]" captionTargetPageId="7" captionText="Fig. 5 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. forewing venation. Scale bar, 1 mm. C: Costa; Sc:Subcosta; R: Radius; M: Media; RS: Radial Sector; MA: Medial Anterior; MP: Medial Posterior; CUA: Cubital Anterior; CUP:Cubital Posterior" figureDoi="http://doi.org/10.5281/zenodo.12005602" httpUri="https://zenodo.org/record/12005602/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). Almost complete specimen of ~
<quantity id="4CA5B5E5C864D005C79AFB1CFC77586D" box="[914,998,1261,1285]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="6" unit="mm" value="6.4">6.4 mm</quantity>
length, preserved in dorso-lateral view. Prognathous head (
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in length) with prominent compound eyes of
<quantity id="4CA5B5E5C864D005C003FADCFBF2582D" box="[1035,1123,1325,1349]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="0.4">0.4 mm</quantity>
diameter; eyes surrounded by short spines; chewing mouthpart with visible mandibles and the short maxillary palps (only the left visible); filiform antennae composed of more than 20 antennomers (
<quantity id="4CA5B5E5C864D005C79CFA5CFC6C58AD" box="[916,1021,1453,1477]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" pageId="5" pageNumber="6" unit="mm" value="2.04">2.04 mm</quantity>
in length) (
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). Well-developed prothorax (
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) but shorter than meso- plus metathorax; sub-rectangular, partially crushed mesothorax (length =
<quantity id="4CA5B5E5C864D005C79BF9FCFC7D5B4D" box="[915,1004,1549,1573]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="mm" value="0.8">0.8 mm</quantity>
, maximum width =
<quantity id="4CA5B5E5C864D005C0C6F9FCFAA55B4D" box="[1230,1332,1549,1573]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="6" unit="mm" value="0.95">0.95 mm</quantity>
) covered with long setae especially visible on the anterior ridge (
<figureCitation id="13660485C864D005C73CF9BCFCE05B0D" box="[820,881,1613,1637]" captionStart="Fig" captionStartId="7.[167,198,665,685]" captionTargetBox="[176,751,236,653]" captionTargetId="figure-348@7.[176,751,236,653]" captionTargetPageId="7" captionText="Fig. 4 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. head and thorax details. Scale bar, 500 µm" figureDoi="http://doi.org/10.5281/zenodo.12005598" httpUri="https://zenodo.org/record/12005598/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
). Metathorax of
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in length and
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of width, respectively. Forelegs visible, one almost complete (coxa, trochanter, femur of
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, tibiae of
<quantity id="4CA5B5E5C864D005C13BF97CFA095BCD" box="[1331,1432,1677,1701]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="6" unit="mm" value="0.91">0.91 mm</quantity>
, and tarsus of
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;
<figureCitation id="13660485C864D005C03BF95CFBE15BAD" box="[1075,1136,1709,1733]" captionStart="Fig" captionStartId="7.[167,198,665,685]" captionTargetBox="[176,751,236,653]" captionTargetId="figure-348@7.[176,751,236,653]" captionTargetPageId="7" captionText="Fig. 4 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. head and thorax details. Scale bar, 500 µm" figureDoi="http://doi.org/10.5281/zenodo.12005598" httpUri="https://zenodo.org/record/12005598/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
), while only tibiae and tarsus of the other are visible. Abdomen of
<quantity id="4CA5B5E5C864D005C0EBF93CFAA45B8D" box="[1251,1333,1741,1765]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="6" unit="mm" value="1.3">1.3 mm</quantity>
in length partially visible.
</paragraph>
<paragraph id="8BE21800C864D006C74AF8FFFC605A13" blockId="5.[813,1437,237,1925]" lastBlockId="6.[813,1436,1699,1915]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B929C412C864D005C74AF8FFFC155A4D" box="[834,900,1806,1829]" italics="true" pageId="5" pageNumber="6">Wings</emphasis>
(
<figureCitation id="13660485C864D005C792F8FCFC765A4D" box="[922,999,1805,1829]" captionStart="Fig" captionStartId="6.[167,198,1611,1631]" captionTargetBox="[211,1377,236,1597]" captionTargetId="figure-144@6.[211,1377,236,1597]" captionTargetPageId="6" captionText="Fig. 3 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. Scale bar, 1 mm" figureDoi="http://doi.org/10.5281/zenodo.12005584" httpUri="https://zenodo.org/record/12005584/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
,
<figureCitation id="13660485C864D005C7F3F8FCFB995A4D" box="[1019,1032,1805,1829]" captionStart="Fig" captionStartId="7.[167,198,1817,1837]" captionTargetBox="[303,1285,1311,1801]" captionTargetId="figure-367@7.[299,1289,1307,1804]" captionTargetPageId="7" captionText="Fig. 5 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. forewing venation. Scale bar, 1 mm. C: Costa; Sc:Subcosta; R: Radius; M: Media; RS: Radial Sector; MA: Medial Anterior; MP: Medial Posterior; CUA: Cubital Anterior; CUP:Cubital Posterior" figureDoi="http://doi.org/10.5281/zenodo.12005602" httpUri="https://zenodo.org/record/12005602/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). Two pairs of subequal membranous wings without pterostigma and with many veins, long setae present on veins, except for the costal (C) vein, where setae are shorter and denser. Forewings elongate, ~ 2.7 times as long as wide (length:
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; maximum width of inter radial area:
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), with a distally rounded shape; moderately broad costal area (until its mid; maximum width
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), then gently decreasing distally until the joining with subcostal (SC) vein, and proximally at the wing base. Costal area with numerous veinlets that, in the proximal part of the wing, are bifurcated. SC vein almost straight, terminating distally on R; few cross veins between R and RS, the latter with six primary branches. Medial (M) veins not coalesced with RS basally, the former is separated in medial anterior (MA) and medial posterior (MP). Cubital anterior 1 distally forked, cubital posterior simple. Anal veins not clearly visible.
</paragraph>
<caption id="DF224888C867D006C4AFF9BAFCA85B37" ID-DOI="http://doi.org/10.5281/zenodo.12005584" ID-Zenodo-Dep="12005584" box="[167,825,1611,1631]" httpUri="https://zenodo.org/record/12005584/files/figure.png" pageId="6" pageNumber="7" startId="6.[167,198,1611,1631]" targetBox="[211,1377,236,1597]" targetPageId="6" targetType="figure">
<paragraph id="8BE21800C867D006C4AFF9BAFCA85B37" blockId="6.[167,825,1611,1631]" box="[167,825,1611,1631]" pageId="6" pageNumber="7">
<emphasis id="B929C412C867D006C4AFF9BAFF4B5B37" bold="true" box="[167,218,1611,1631]" pageId="6" pageNumber="7">Fig. 3</emphasis>
<taxonomicName id="4C5D6383C867D006C4ECF9BAFEEC5B37" authorityName="Montagna &amp; Magoga &amp; Magnani" authorityYear="2024" box="[228,381,1611,1631]" class="Insecta" family="Permithonidae" genus="Merithone" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laetitiae">
<emphasis id="B929C412C867D006C4ECF9BAFEEC5B37" box="[228,381,1611,1631]" italics="true" pageId="6" pageNumber="7">Merithone laetitiae</emphasis>
</taxonomicName>
(†
<taxonomicName id="4C5D6383C867D006C598F9BAFD815B37" authorityName="Tillyard" authorityYear="1922" box="[400,528,1611,1631]" class="Insecta" family="Permithonidae" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Permithonidae</taxonomicName>
) gen. nov., sp. nov. Scale bar, 1 mm
</paragraph>
</caption>
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<paragraph id="8BE21800C866D007C4AFFD68FE4D5FA0" blockId="7.[167,734,665,712]" pageId="7" pageNumber="8">
<emphasis id="B929C412C866D007C4AFFD68FF4B5FC5" bold="true" box="[167,218,665,685]" pageId="7" pageNumber="8">Fig. 4</emphasis>
<taxonomicName id="4C5D6383C866D007C4ECFD68FEEC5FC5" authorityName="Montagna &amp; Magoga &amp; Magnani" authorityYear="2024" box="[228,381,665,685]" class="Insecta" family="Permithonidae" genus="Merithone" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="laetitiae">
<emphasis id="B929C412C866D007C4ECFD68FEEC5FC5" box="[228,381,665,685]" italics="true" pageId="7" pageNumber="8">Merithone laetitiae</emphasis>
</taxonomicName>
(†
<taxonomicName id="4C5D6383C866D007C598FD68FD815FC5" authorityName="Tillyard" authorityYear="1922" box="[400,528,665,685]" class="Insecta" family="Permithonidae" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Permithonidae</taxonomicName>
) gen. nov., sp. nov. head and thorax details. Scale bar, 500 µm
</paragraph>
</caption>
<paragraph id="8BE21800C866D007C4A5FCE3FE8E5EC2" blockId="7.[151,775,786,938]" pageId="7" pageNumber="8">
Hind wings are visible and subequal to forewings (
<figureCitation id="13660485C866D007C497FCC3FF4F5E22" box="[159,222,818,842]" captionStart="Fig" captionStartId="6.[167,198,1611,1631]" captionTargetBox="[211,1377,236,1597]" captionTargetId="figure-144@6.[211,1377,236,1597]" captionTargetPageId="6" captionText="Fig. 3 Merithone laetitiae (†Permithonidae) gen. nov., sp. nov. Scale bar, 1 mm" figureDoi="http://doi.org/10.5281/zenodo.12005584" httpUri="https://zenodo.org/record/12005584/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
), as demonstrated by the right hind wing almost completely overlapping the corresponding forewing, the left hind wing is crumpled, and venations are only partially visible.
</paragraph>
</subSubSection>
</treatment>
</document>