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<document id="FC696A7627ACA7D5B75356C0DEDFD586" ID-DOI="10.3897/zookeys.1207.117225" ID-ZooBank="94F18819-5AF5-4100-AB35-AA3C3976EE80" ID-publisher-id="117225" URI-arpha="255F6420-5705-5F50-9167-509E6C76FA1C" URI-zoobank="http://zoobank.org/94F18819-5AF5-4100-AB35-AA3C3976EE80" article-type="research-article" checkinTime="1721676390257" checkinUser="pensoft" docAuthor="Angus, Robert B., Maté, Jason F., Angus, Elizabeth M. & Král, David" docDate="2024" docId="121D8F6875C25860934C05AC5878D13E" docLanguage="en" docName="ZooKeys 1207: 205-299" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/117225/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Aphodius (Liothorax) krelli Angus, Maté, Angus & Král, 2024, sp. nov." docType="treatment" docUuid="40001D30-4534-4350-97CB-5E286675ADC8" docUuidSource="ZooBank" docVersion="1" lastPageNumber="299" masterDocId="255F642057055F509167509E6C76FA1C" masterDocTitle="Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" masterLastPageNumber="299" masterPageNumber="205" pageNumber="205" updateTime="1721676524171" updateUser="pensoft">
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<mods:titleInfo id="A538BABF7ADD0B434DC31DA3F4574621">
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<mods:title id="5D2F697E1A81816DA903B9EE0BC3547E">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:namePart id="2F40C75EF16DA0908496AF7E5577D6C5">Angus, Robert B.</mods:namePart>
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<mods:nameIdentifier id="404CC0B4F4D2ACC5AF6660AD9F32A6D9" type="ORCID">0000-0002-3860-5617</mods:nameIdentifier>
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<mods:affiliation id="5AFBBE9A33AB5010C24AFF91A822940D">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="2696BD9160A6FF430F0EEA07F4F1E326">Maté, Jason F.</mods:namePart>
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<mods:nameIdentifier id="82BDFA1FE74110BBB8363E05724E6046" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
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<mods:affiliation id="5E91CB155EDCD616FF4E8B0AD1D50068">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="6C5D6058D287B52788F0D35EF2A32902">Angus, Elizabeth M.</mods:namePart>
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<mods:affiliation id="CE6C3C1EE25C93C68E5E6D42D48B6BD0">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="44D10A81E1DCFAA744C278567C553747">Král, David</mods:namePart>
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<mods:affiliation id="5E8ABCF49D816C15B2FFE0D27DD73627">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:date id="503AD83FE49C46BA7B9758E36C37554D">2024</mods:date>
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<mods:number id="4108C994D80CBA2523C58AAFBB6284FA">2024-07-22</mods:number>
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<mods:identifier id="4D6A9A88536B0486EE25A8E7331F563F" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<treatment id="121D8F6875C25860934C05AC5878D13E" ID-arpha="121D8F68-75C2-5860-934C-05AC5878D13E" ID-zoobank="https://zoobank.org/40001D30-4534-4350-97CB-5E286675ADC8" LSID="urn:lsid:zoobank.org:act:40001D30-4534-4350-97CB-5E286675ADC8" LSID-ZBK="urn:lsid:zoobank.org:act:40001D30-4534-4350-97CB-5E286675ADC8" httpUri="http://treatment.plazi.org/id/121D8F6875C25860934C05AC5878D13E">
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<paragraph id="C7EBB16733D9A5D818D51ABB3EB60543">
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<taxonomicName id="641D88A6C08774D39EDEFC89BBC07A57" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli" status="sp. nov." subGenus="Liothorax">
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<emphasis id="819F136BA4DF833A6704244BCCBD6CBC" italics="true">Aphodius</emphasis>
|
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(
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<emphasis id="91235B625CD06F888128F306C8EF0025" italics="true">Liothorax</emphasis>
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)
|
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<emphasis id="37F9A3D35341927155E602D2BF810AB6" italics="true">krelli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="996E338E95030A2B1DC0D506F8971022" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="8871EBBBE57B576F765990FFB6D7E957" type="description">
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<figureCitation id="15F1245A112439222FB6E69D60E646DB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 e</figureCitation>
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,
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<figureCitation id="3267C155F91B22554D1E5C2168B8AB49" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 h</figureCitation>
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,
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<figureCitation id="C8957C8D5A8961E4CE6C418CF3AC05C5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 j</figureCitation>
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,
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<figureCitation id="DAE01236610D6BEE8B06FBDF8B6C6F06" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 b</figureCitation>
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||||
,
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||||
<figureCitation id="8BDD8B2036CAB4869C212B8879234528" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 y</figureCitation>
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,
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<figureCitation id="E955F72A8B4D0C31631298E72FF3C92D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">
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<quantity id="A33C1DECECEEDD7F5D4A18638F8B3C18" metricMagnitude="1" metricUnit="m" metricValue="1.8" unit="m" value="18.0">18 m</quantity>
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, n
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<figureCitation id="5D466C37BB4A02CDFE9F61C48AE9932D" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">
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<quantity id="291133A10AE55399C0D3BD1E0C7F0FC3" metricMagnitude="1" metricUnit="m" metricValue="2.3" unit="m" value="23.0">23 m</quantity>
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, o
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,
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<figureCitation id="326DDFAEABA127BDB24EF8CE36560001" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 y</figureCitation>
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,
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<figureCitation id="BBA20CFBC6530EC11419E380C1FDD1FF" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Mitotic chromosomes arranged as karyotypes a – d A. bameuli sp. nov., ♂ paratypes, FR, Corsica, by the Lac de Melo. Autosome 9 polymorphic in a, b homozygous submetacentric in c, d. a, c plain b and d the same nuclei C-banded e – l A. krelli sp. nov., types, IT, Sardinia e, f paratype ♂ with the heterochromatic long arm of autosome 1 weakly developed e plain f the same nucleus C-banded g, h holotype with the heterochromatic long arm of autosome 1 fully developed g plain h the same nucleus C-banded i, j paratype ♂ with both replicates of autosome 7 lacking the heterochromatic long arm i plain j the same nucleus C-banded k, l paratype ♀ with autosome 9 heterozygous for an inversion polymorphism k plain l the same nucleus C-banded m – p A. wilsonae SP m, n holotype ♂ m plain n the same nucleus C-banded o, p paratype ♀ from Balneario de Corconte o plain p the same nucleus C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure33" httpUri="https://binary.pensoft.net/fig/1097522">33 e – l</figureCitation>
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,
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<figureCitation id="09D56CEBDA3F3191A4DC647572CA7263" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Meiotic chromosomes, metaphase I from testis a – d A. niger, GB, Hampshire, New Forest, White Moor a, c plain b, d the same nuclei C-banded e – g A. muscorum, HU, Hortobágyi e plain f, g the same nucleus f plain g C-banded h A. bameuli paratype, FR, Corsica, by Lac de Melo, plain i, j A. krelli IT, Sardinia, paratype i plain j the same nucleus C-banded k, l A. wilsonae, SP, Areños k plain l the same nucleus C-banded. The sex bivalent is indicated by an arrow. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure34" httpUri="https://binary.pensoft.net/fig/1097526">34 i, j</figureCitation>
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</paragraph>
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<subSubSection id="SECID0EFICM" type="type material">
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<paragraph id="67F114377F97BD74A239849EC6E097BC">
|
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<heading id="0950BDA265B593BCF479CEE9899DDBD2" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FCC28C9789031CCCB81381F2B6E71D1A">
|
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<materialsCitation id="D51054BB0BB6093B82C6884CA8F1397D" collectingDate="2012-04-03" collectingDateMax="2012-04-05" collectingDateMin="2012-04-03" collectionCode="NHMUK" collectorName="R. B. & E. M. Angus." location="Badde Salighes" municipality="Nuoro Province" specimenCount="1" specimenCount-male="1" stateProvince="Sardegna" typeStatus="holotype">
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<emphasis id="0E6E2B3B417A9FE0E18BB23EC4E1E193" bold="true" italics="true">
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<typeStatus id="1B0EF07682684EEBF03901B66073E597" type="holotype">Holotype</typeStatus>
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||||
</emphasis>
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||||
<specimenCount id="163606D72D16E5BDF28ABAB3F3960D48" count="1" type="male">♂</specimenCount>
|
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:
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<collectingRegion id="0A09620034D4F8C08C15BE073128C558" country="Italy" name="Sardegna">Sardinia</collectingRegion>
|
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,
|
||||
<collectingMunicipality id="5841B3626D771E7847F8798C3C79D585">Nuoro Province</collectingMunicipality>
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,
|
||||
<location id="EC953F74D22DAD1F8DBB554A48E31C66" LSID="urn:lsid:plazi:treatment:121D8F6875C25860934C05AC5878D13E:EC953F74D22DAD1F8DBB554A48E31C66" municipality="Nuoro Province" name="Badde Salighes" stateProvince="Sardegna">Badde Salighes</location>
|
||||
<date id="38B500A5FFB24E7A0ABF810D0644240D" value="2012-04-03">
|
||||
<collectingDate id="DD609F706B1343E8FE5AC89954D4F727" value="2012-04-03">3. iv. 2012</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="B9674880B717412CC73A6283D3A3FE69">R. B. & E. M. Angus.</collectorName>
|
||||
L 5.0 mm, b
|
||||
<quantity id="87462D546006F10AF4DD537F0819D4E7" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
|
||||
. Chromosome prep. 2,
|
||||
<date id="A3AA49316472248F3BE2EC73D1E85718" value="2012-04-05">
|
||||
<collectingDate id="3F2C14B307CA22C449A0ABB522FBA5EF" value="2012-04-05">5. iv. 2012</collectingDate>
|
||||
</date>
|
||||
(
|
||||
<named-content id="9819E5E7453961BEF745BE01A5EC20E7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="29467308FB2BB1E8888E359E94001E99" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="725EA3807B53AB7C8424D3E57B2A54C4" bold="true" italics="true">
|
||||
<typeStatus id="1742F49EDD14E9D37818601641E50A4E" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="2324B61656AE1BE8DF1FAD6DCF690150" count="4" type="male">4 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="35CEBE76D416412A3BB74EDC6D210F28" count="4" type="female">4 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="DB0A05CD30CB908D065C5DA6F03913FD" type="holotype">holotype</typeStatus>
|
||||
, but
|
||||
<specimenCount id="6593C422FC1EC6FF4A753E17B095AC53" count="1" type="male">1 ♂</specimenCount>
|
||||
taken on
|
||||
<date id="1F7FC6FF1A2D1395387A61486D342656" value="2012-04-01">1. iv. 2012</date>
|
||||
.
|
||||
<specimenCount id="680600D908ACF24DB3E190BC91F14613" count="1" type="male">♂</specimenCount>
|
||||
<typeStatus id="F31A73FF231CD99815F47DB1A807FE86" type="paratype">paratypes</typeStatus>
|
||||
with chromosome preparation data prep. 5:
|
||||
<date id="A8B9369C60C5B5A0C113A66232F319EF" value="2012-04-05">5. iv. 2012</date>
|
||||
and prep. 1:
|
||||
<date id="39FEBC65866652F6D74354B68A6E8738" value="2012-04-10">10. iv. 2012</date>
|
||||
,
|
||||
<specimenCount id="A6E842BBA75D74298E798A1E2B26A5D7" count="1" type="female">♀</specimenCount>
|
||||
<typeStatus id="43B2AE0D4EA6F2CA9271E03D1AA20B89" type="paratype">paratypes</typeStatus>
|
||||
with data preps 3, 4, 6–8:
|
||||
<date id="F134C5021B049F2EF564A18666F7F563" value="2012-04-05">5. iv. 2012</date>
|
||||
(
|
||||
<named-content id="6EFFB0EE7585A0ACB8CDFA007BA30D34" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="D4F89B0A673E332FDE9DA5DDD0C404D3" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="C976BEB38173DC5BD152BEF14535A721" count="1" type="female">1 ♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="5AFADC53684F66C58AE3989A7E7149BC" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<named-content id="4483DF13678C49296298A162AE9E849D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="00A87D9CF41552DCF973534B33D70C69" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
<specimenCount id="8B1C74C57499993158757A59893C65E2" count="1" type="female">1 ♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="D8C7C6C46E845007FB94D68F5F896BC8" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<abbrev id="ABBRID0E6ICM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="A474A0C06227620F844F1EC173275481">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
<specimenCount id="8025F746399CFC9B3F90DA04ADEA68A9" count="1" type="male">1 ♂</specimenCount>
|
||||
, chromosome prep 5,
|
||||
<date id="598E4BCF00B9014A396660CB4AAFB914" value="2012-04-05">5. iv. 2012</date>
|
||||
, data as
|
||||
<typeStatus id="2A548DEB563780CA9A3268278FF66276" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<named-content id="23396BAF9FE425DBE14230C05B4113FD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/denver-museum-nature-science" xlink_title="Denver Museum of Nature and Science">
|
||||
<collectionCode id="FDF6E3733B11E4F629D1A39956990C37" country="USA" httpUri="http://grbio.org/cool/mmzs-jqix" name="Denver Museum of Nature and Science" type="Museum">DMNS</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="E4DF95268DBE3EE3B2DCDD180BD22551" count="1" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<collectingRegion id="B9B07298454A28DA8E708BA73F4253B9" country="Italy" name="Sardegna">Sardinia</collectingRegion>
|
||||
, Sassari, Ciaccia,
|
||||
<date id="7C78D367D6E3323E6666209CBAA9759D" value="2008-07-17">17. VII. 2008</date>
|
||||
, old sheep dung, R. B. Angus (
|
||||
<abbrev id="ABBRID0EHJCM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="6CC010C91FFA95D04463067A7FAC02F2">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
); 2 unsexed,
|
||||
<collectingCountry id="D9740BABBFE439E5D9E0EE5D51522E11" name="Italy">ITALY</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="11A0F91522A223B29B1C6BADF7AC215F" country="Italy" name="Sardegna">Sardegna</collectingRegion>
|
||||
, Nuoro, Altopiano della Campeda,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOJCM" specific-use="{"type":"Point","coordinates":[8.784067,40.354083]}">
|
||||
<geoCoordinate id="C260639FF233ED256AC0B0AFC43B01FA" degrees="40" direction="north" minutes="21.245" orientation="latitude" precision="1" value="40.354084">40 ° 21.245 ' N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="B7F0EFFE14FD60B88D3510371584FE0D" degrees="8" direction="east" minutes="47.044" orientation="longitude" precision="1" value="8.784066">8 ° 47.044 ' E</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<quantity id="497DD8EBEFE0777F4C8EF709D5ED811A" metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">580 m</quantity>
|
||||
,
|
||||
<date id="C375C12F2193E02B4F698EFC04F34687" value="2005-05-18">18. 5. 2005</date>
|
||||
leg. Starke (
|
||||
<abbrev id="ABBRID0ETJCM" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="0C22AB4E737A59CA293EEE9AE54AC072">ABC</collectionCode>
|
||||
</abbrev>
|
||||
); 1 unsexed, [
|
||||
<collectingCountry id="80FC1E3F4A6B7A7FC774D98C726DC37A" name="Italy">ITALY</collectingCountry>
|
||||
], Badde Salighes
|
||||
<collectingRegion id="20B4F23CF726566A206B625B8211C611" country="Italy" name="Sardegna">Sardegna</collectingRegion>
|
||||
[Nuoro Pr],
|
||||
<taxonomicName id="A211247211F7C5FC184630172E973CF9" authority="Carpaneto" authorityName="Carpaneto" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger" subGenus="Nialus">
|
||||
<emphasis id="743CCE14D4D491F9D0BE7252FC7BBAC4" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="E58A81D1B992B80EEE0E31C7457270A7" italics="true">Nialus</emphasis>
|
||||
)
|
||||
<emphasis id="6E5CC08585424CEC3BA43165F7AF487A" italics="true">niger</emphasis>
|
||||
Carpaneto
|
||||
</taxonomicName>
|
||||
det. 1982 (
|
||||
<abbrev id="ABBRID0EFKCM" xlink_title="Muséum d’histiore naturelle, Genève">
|
||||
<collectionCode id="334F831659DE520991829F07E48CC4BE">MNHG</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJKCM" type="Differential diagnosis">
|
||||
<paragraph id="0B3C131C1D5565F96D983692F5ABCF66">
|
||||
<heading id="6372D2B2D8C418EC32D9C7E62D8D8049" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="096526FBD12BC2DECE560EE76722FC84">
|
||||
Sardinian
|
||||
<taxonomicName id="0BE8B6C98F41DC944AE3FB751A66FC38" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli">
|
||||
<emphasis id="ADF21F9D2C886AA78BB73FEE849C712B" italics="true">A. krelli</emphasis>
|
||||
</taxonomicName>
|
||||
differs from Corsican
|
||||
<taxonomicName id="2C8158102563935E27A1A1F7BFF23ADA" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli">
|
||||
<emphasis id="8A4844B3430BF4D36B5806C211A1B747" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
in being less parallel-sided (Fig.
|
||||
<figureCitation id="B164A51CDAC847CE269D87548EF419FA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 e, d</figureCitation>
|
||||
) and with the pronotum more heavily punctate (Fig.
|
||||
<figureCitation id="FBD7E75E6029BF75622BFDD3C236C086" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 h, g</figureCitation>
|
||||
). The karyotypes of the two are very similar in the sizes and shapes of their chromosomes (Fig.
|
||||
<figureCitation id="4D24BBA1F271C2AE139F96DAE69A206D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Mitotic chromosomes arranged as karyotypes a – d A. bameuli sp. nov., ♂ paratypes, FR, Corsica, by the Lac de Melo. Autosome 9 polymorphic in a, b homozygous submetacentric in c, d. a, c plain b and d the same nuclei C-banded e – l A. krelli sp. nov., types, IT, Sardinia e, f paratype ♂ with the heterochromatic long arm of autosome 1 weakly developed e plain f the same nucleus C-banded g, h holotype with the heterochromatic long arm of autosome 1 fully developed g plain h the same nucleus C-banded i, j paratype ♂ with both replicates of autosome 7 lacking the heterochromatic long arm i plain j the same nucleus C-banded k, l paratype ♀ with autosome 9 heterozygous for an inversion polymorphism k plain l the same nucleus C-banded m – p A. wilsonae SP m, n holotype ♂ m plain n the same nucleus C-banded o, p paratype ♀ from Balneario de Corconte o plain p the same nucleus C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure33" httpUri="https://binary.pensoft.net/fig/1097522">33 e – l</figureCitation>
|
||||
) but the long arm of autosome
|
||||
<quantity id="9320381447EAC616D4E0029BBAF332A0" metricMagnitude="-1" metricUnit="m" metricValue="1.778" unit="in" value="7.0">7 in</quantity>
|
||||
<taxonomicName id="7E413221976882F2B29822576C4087CF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli">
|
||||
<emphasis id="5C28B432B9825BCD5358C16ED1A942C4" italics="true">A. krelli</emphasis>
|
||||
</taxonomicName>
|
||||
is heterochromatic and is polymorphic for a deletion – it may be present (Fig.
|
||||
<figureCitation id="A3612F4DF6C4BCD50BE733565EF45A6D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Mitotic chromosomes arranged as karyotypes a – d A. bameuli sp. nov., ♂ paratypes, FR, Corsica, by the Lac de Melo. Autosome 9 polymorphic in a, b homozygous submetacentric in c, d. a, c plain b and d the same nuclei C-banded e – l A. krelli sp. nov., types, IT, Sardinia e, f paratype ♂ with the heterochromatic long arm of autosome 1 weakly developed e plain f the same nucleus C-banded g, h holotype with the heterochromatic long arm of autosome 1 fully developed g plain h the same nucleus C-banded i, j paratype ♂ with both replicates of autosome 7 lacking the heterochromatic long arm i plain j the same nucleus C-banded k, l paratype ♀ with autosome 9 heterozygous for an inversion polymorphism k plain l the same nucleus C-banded m – p A. wilsonae SP m, n holotype ♂ m plain n the same nucleus C-banded o, p paratype ♀ from Balneario de Corconte o plain p the same nucleus C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure33" httpUri="https://binary.pensoft.net/fig/1097522">33 f, i – l</figureCitation>
|
||||
) or absent (Fig.
|
||||
<figureCitation id="068628180DF1442717D88DD3B4838236" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Mitotic chromosomes arranged as karyotypes a – d A. bameuli sp. nov., ♂ paratypes, FR, Corsica, by the Lac de Melo. Autosome 9 polymorphic in a, b homozygous submetacentric in c, d. a, c plain b and d the same nuclei C-banded e – l A. krelli sp. nov., types, IT, Sardinia e, f paratype ♂ with the heterochromatic long arm of autosome 1 weakly developed e plain f the same nucleus C-banded g, h holotype with the heterochromatic long arm of autosome 1 fully developed g plain h the same nucleus C-banded i, j paratype ♂ with both replicates of autosome 7 lacking the heterochromatic long arm i plain j the same nucleus C-banded k, l paratype ♀ with autosome 9 heterozygous for an inversion polymorphism k plain l the same nucleus C-banded m – p A. wilsonae SP m, n holotype ♂ m plain n the same nucleus C-banded o, p paratype ♀ from Balneario de Corconte o plain p the same nucleus C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure33" httpUri="https://binary.pensoft.net/fig/1097522">
|
||||
<quantity id="77320524AD659D4672DDE261AC79584E" metricMagnitude="-2" metricUnit="kg" metricValue="3.3" unit="g" value="33.0">33 g</quantity>
|
||||
, h
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEMCM" type="description">
|
||||
<paragraph id="F1038D55A61AD4D64A77D36C195C3FAF">
|
||||
<heading id="726C60626188C89690A523D61E769150" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="95CCFE391FB58429449704B309AA0574">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="EC721BE84A300D251BD024223490061C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 e</figureCitation>
|
||||
). Length: 4.4–5.0 mm, width: 2.0–
|
||||
<quantity id="BF007B3EF0D84A557C2DF54FE673F8DA" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
|
||||
(
|
||||
<specimenCount id="8ACD994447ADA48296F0FC304D5922DB" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="5347CEAC78F7249AFF0E819A25CE4FC4" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5 mm</quantity>
|
||||
, width 1.7–2.0 mm (
|
||||
<specimenCount id="3A7828CF44559BE23EAF4B422D290965" count="1" type="female">♀</specimenCount>
|
||||
). Head: glossy black with no trace of reticulation, anterior margin vaguely browner. Sculpture of double punctation, the fine punctures separated by ca 2 × their width, the coarser punctures ca 3 × the diameter of the fine ones, distributed over the anterior and lateral parts of the head and on the frons. Frons elevated in a vague hump on disc. Frontoclypeal suture very fine but distinct and complete. Anterior margin of clypeus emarginated medially, the margin rounded either side of this emargination. Anterior and lateral margins of head bordered, the border extending from the back of the genae. Genae distinct, fairly abruptly divergent anteriorly from anterior margin of the eyes, widest point behind middle of genae, in front of which the genal margins curve more gently to the clypeus, which they meet at a slight angle. Epipharynx (Fig.
|
||||
<figureCitation id="EDE2985731A27E1BAB6FA028AD3FF080" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 j</figureCitation>
|
||||
) with anterior margin of clithra strongly emarginate each side of the median tylus, and with a few fine acropariae. Chaetopedia stout and long, four or five each side; chaetopariae closely set, fairly stout, forming a line each side. Prophobae very fine, clustered along the edges of the mesoepitorma; apophobae fine and long, arranged in a line outside the chaetopariae. Antennae and palpi blackish brown. Head and pronotum as in Fig.
|
||||
<figureCitation id="BCB12324D9D5EE8CA476AE7A23BDCD72" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 h</figureCitation>
|
||||
. Pronotum: hemicylindrical, highly arched transversely but very weakly so longitudinally in the mid-line. Lateral margins entirely visible from above. Lateral margin bordered, basal margin with very fine border, this sometimes complete but usually absent from the median 1 / 2 to 1 / 3 of the margin. Surface glossy black with no hint of reticulation but with double punctation. Fine punctures dot-like, extending over the whole surface, coarser punctures ca 4–8 × the diameter of the dots, sparser and tending to be smaller on the disc. Fine punctures separated by ca 4–6 × their diameter, coarse ones by 1.5–2.5 × their diameter.
|
||||
</paragraph>
|
||||
<paragraph id="EC5016A168EE30049F0E1E369E469F05">Scutellum: narrow, pentagonal, sides parallel in basal 1 / 2 then convergent to a blunt point apically. Surface glossy black with a few punctures in basal 1 / 2.</paragraph>
|
||||
<paragraph id="46B7765B2FF420EB79E3101E220B816E">
|
||||
Elytra: elongate, not quite parallel-sided, widest just behind middle, then tapered to bluntly rounded apex. Ground colour of interstices silky black with very fine faint isodiametric reticulation (Fig.
|
||||
<figureCitation id="545A429566806D32CC3D29CC467508A8" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 y</figureCitation>
|
||||
). Striae glossy black. Interstices flat, ca 10 × the width of the striae, with scattered fine punctures separated by ca 4–5 × their width. Sutural interstices weakly raised longitudinally from their external margins to ca 1 / 3 of the way to the suture, then depressed to the suture. Striae narrow, with vertical sides, with a single row of punctures separated by ca 3 × their diameter, the sides of these punctures encroaching into the strial margins. Metaventrite with median diamond-shaped area fairly strongly punctate, the punctures larger than in
|
||||
<taxonomicName id="B6436CB75E547983BC6D6568919BD7E6" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli">
|
||||
<emphasis id="822B2E2A8830D87219F2E1E1A75BD679" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
, in males concave to median impressed furrow, in females often flatter.
|
||||
</paragraph>
|
||||
<paragraph id="D4E59FDE37176ED8CE3742B1DAD2D9EF">
|
||||
Legs: dark brown, longer spur of mid tibiae clearly longer than basal segment of tarsus, although sometimes only slightly so (Fig.
|
||||
<figureCitation id="00B6E0929716330325695E0EB9BA63D7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">
|
||||
<quantity id="5F93C2DEF00239943E29AFD5ABDC44EE" metricMagnitude="1" metricUnit="m" metricValue="1.8" unit="m" value="18.0">18 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
). Post tibiae with apical fringe of spines short and of even-length ventrally, longer, sparser and of uneven lengths dorsally.
|
||||
</paragraph>
|
||||
<paragraph id="DC7047CEF4428F07EA9B02D628D1DA81">
|
||||
Aedeagus: Fig.
|
||||
<figureCitation id="75589FB84A121FFBC9B7D1CC3CA74919" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">
|
||||
<quantity id="6600B2D06691EEAF123A92AEA5EEEA07" metricMagnitude="1" metricUnit="m" metricValue="2.3" unit="m" value="23.0">23 m</quantity>
|
||||
– o
|
||||
</figureCitation>
|
||||
. Length ca
|
||||
<quantity id="196C5D73DB870E1B3EE91465B9D77AB6" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
, length of parameres ca
|
||||
<quantity id="E39B2FA86D8BB14DB3F9D81589E2D41A" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
, of basal piece ca
|
||||
<quantity id="89C427D5929F70918DD591948564FD24" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
|
||||
, of endophallic tooth-field ca
|
||||
<quantity id="98E3BCC86C357D09ACA1D060B6998D49" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
|
||||
. Length of longest endophallic teeth 56–58 μm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUNCM" type="etymology">
|
||||
<paragraph id="E9F90E3EB2FF7F60195359B01E751D4F">
|
||||
<heading id="55A24C6A36912533C2E79896C85E0F4A" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="885798F8F4CCE2FDCEB12DBC6475F55D">
|
||||
This species is named after Dr F. - T. Krell of the Museum of Nature and Science, Denver,
|
||||
<collectingRegion id="09CE3B6DBFF37D50E1C0A4CDDB19A250" country="United States of America" name="Colorado">Colorado</collectingRegion>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZNCM" type="remarks">
|
||||
<paragraph id="D40F841F91371DBFDB1E4C21E4DAD5B1">
|
||||
<heading id="1B2BC53B4C0156D900D6E25A610113C4" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8A5B9E86C4B1E2869B72BFA491593527">
|
||||
<taxonomicName id="985427C330A4ABE78566FFEB0C61F7D2" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli" status="sp. nov.">
|
||||
<emphasis id="6FC0F9D8A77BA07119017BB657430B22" italics="true">Aphodius krelli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="C75C6F0B982FDE120E7A947241C4256D" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is endemic to Sardinia (Fig.
|
||||
<figureCitation id="56D2D3C0FEA84C070F2FBBD28ACB1A3A" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
).
|
||||
<bibRefCitation id="809A285AC67E8BF5A111CE35C1D62FA1" author="Pittino R" firstAuthor="Pittino" issue="7-8" journalOrPublisher="Bolletino della Societá entomologica italiana, Genova" pagination="127-134" refId="B40" refString="Pittino R (2010) Aphodiidae interessanti della regione sardo-corsa (Col. Scarabaeoidea). Bolletino della Societá entomologica italiana, Genova 112 (7–8): 127–134." title="Aphodiidae interessanti della regione sardo-corsa (Col. Scarabaeoidea)." volume="112" year="2010">Pittino (2010)</bibRefCitation>
|
||||
lists various Sardinian localities, including Badde Salighes, but all the analysed material is from Badde Salighes (Fig.
|
||||
<figureCitation id="FEFAA10816380D28B78F1D8BAC41D2AB" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 d</figureCitation>
|
||||
).
|
||||
<taxonomicName id="471A66B3F8BD4A397EDFABCB3BB785BF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli" status="sp. nov.">
|
||||
<emphasis id="D8E7FD730E085D1159F920DA52FF9FAB" italics="true">Aphodius krelli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7514438F92566427230F8F2B3C299613" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is described as a new species despite the fact that its karyotype is similar to that of
|
||||
<taxonomicName id="6B5843BC378435A904F79A3B4775ABC6" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov.">
|
||||
<emphasis id="FD6DC13ECE819655BE694A391F7CBBD8" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D454F5708C4BD4BB5DB737F9C108100A" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
from
|
||||
<collectingCountry id="F2BDF769B25B129AF9AAB6BDD81FB58F" name="France">Corsica</collectingCountry>
|
||||
. In spite of the karyological similarities, it is genetically divergent from
|
||||
<taxonomicName id="B114E32B3ADDE919A22F2B60FA42A9C6" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli">
|
||||
<emphasis id="3221EDCDB462E24098EFB4E3E07472A5" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
, from which it differs by 4.7 %, and instead clusters with
|
||||
<taxonomicName id="8131D10B192A88351A30B377617FF1EE" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens" status="sp. nov.">
|
||||
<emphasis id="78B5F4B9E1EA6E723FB5967B60529E79" italics="true">A. bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="2A34ADA6F1C30D19C7CA7E307FDBBB7C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, forming a strongly supported clade. In addition the morphometric study clusters
|
||||
<taxonomicName id="2B52E78C3865A8A0A309044A995847D5" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli" status="sp. nov.">
|
||||
<emphasis id="3A86F2647CE6C5D74C9D14FF7AB1742F" italics="true">A. krelli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4F190A4F28302DA89CE270336F2C85E4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
between the
|
||||
<taxonomicName id="AD1D9BD293CE2E1A5783C85338958AA6" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="3FA1C4C03F692F6A513D42731049051C" italics="true">niger</emphasis>
|
||||
</taxonomicName>
|
||||
- crown group and the 3 B group, from which it differs in being significantly more convex. Morphologically it differs from
|
||||
<taxonomicName id="ECEFB814BCB28BF95A6090F0E50C2FBD" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov.">
|
||||
<emphasis id="497B9A82A8ABBC55ED69A4398F72142E" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="6F436919608CB685B6BB9E67B10C7B90" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in being generally more heavily sculptured and more rounded. Its habitat is also different – whereas
|
||||
<taxonomicName id="738A8FAE212DAED0148C8FE7C0FF0A3A" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov.">
|
||||
<emphasis id="C00D317BACF78869EAF4B93FA8B71CCE" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="48F74B251FD0C2E3439A18216425A2DA" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is a dung-inhabiting species,
|
||||
<taxonomicName id="D71A6F987AAAA51B59FC17FDCEA717C8" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="krelli" status="sp. nov.">
|
||||
<emphasis id="57394A30E2DA018542A27D321CA53F5F" italics="true">A. krelli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="467718AA623C08A5D2A61D6B6C812151" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, like most
|
||||
<taxonomicName id="C94CB02D5E9CD589B8EB454DFC4BB440" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="09F67281721504AA1EAC0DF0A664DD91" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
, is found at the edge of water. The
|
||||
<typeStatus id="FD144528B007B05B60DBA7E6120F5F84">type</typeStatus>
|
||||
material, brought home alive for laboratory studies, was divided into two lots. One was given organic detritus from where it was collected while the other was given damp fibrous cow-dung of the sort eaten by
|
||||
<taxonomicName id="B8C81A093233E9A86EF40783135FA54A" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov.">
|
||||
<emphasis id="37734413CF941176C65597D35C6AF7CF" italics="true">A. bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E761AD626B50AE58EDDD773E6B1BD458" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
The beetles given detritus continued feeding on the journey home but those given dung did not feed at all.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
294
data/21/73/75/21737574B2615B2FA919858AE9561D34.xml
Normal file
294
data/21/73/75/21737574B2615B2FA919858AE9561D34.xml
Normal file
|
|
@ -0,0 +1,294 @@
|
|||
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<mods:titleInfo id="43673D75C03FCBCC56B3FC912EBE934B">
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<mods:title id="3CB4D93E62061376AEA3C258B92C15D5">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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</mods:titleInfo>
|
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<mods:name id="EC0925C071E341F73E84CD7AD61D7C0B" type="personal">
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<mods:namePart id="1103732C8DB897183C1D351A1078B714">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="73F9F30448BBDD9337888726A8AFC5EC">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:affiliation id="EB9013388E9F05CABE37D9E139C2B2DD">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="D69238CD9B1098370CE46911F7EB6AA9">Angus, Elizabeth M.</mods:namePart>
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<mods:affiliation id="F58D58292F5F82276D9D93690C0D5C9D">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="3813B7E31AD0785DE813F60425B58A8B">Král, David</mods:namePart>
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<mods:affiliation id="7C112B9FB5AC3E8DE248A4B3D70CDEBF">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:date id="976C837AB40B85DE5A58EF019899EE7E">2024</mods:date>
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<mods:identifier id="8D08B52482E1AF03722CCAFE32F44E46" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<taxonomicName id="487207B23A8FC40794174AAA4ECDDF0D" ID-CoL="FGDV" authority="Gusakov, 2004" authorityName="Gusakov" authorityYear="2004" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rusakovi" subGenus="Liothorax">
|
||||
<emphasis id="E2149F00572789DCD9E73BE0448AC148" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="1F6629EE68CF6D1CD90D1968EFC46BCD" italics="true">Liothorax</emphasis>
|
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)
|
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<emphasis id="D9E37088BD828B6F35D392E48F5FDDF7" italics="true">rusakovi</emphasis>
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<bibRefCitation id="EB73F8604AB58890EBE7F346DB7FE6AC" author="Gusakov AA" refId="B19" refString="Gusakov AA (2004) Novye vidy plastinchtousykh zhukov palearcticheskoj fauny. [New species of scarab beetles from the Palaearctic region]. Izdatel'skij dom Ejdos, Kaluga, 1 - 38." year="2004">Gusakov, 2004</bibRefCitation>
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<figureCitation id="1193F73A663D7E07866CFDE8617E90A1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">Figs 2 i</figureCitation>
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,
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<figureCitation id="1261AC824F2FFC02C54056360436A0F1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">
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<quantity id="EBE51462A23D8D515F1A84B54006B860" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
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,
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<figureCitation id="88D723A88A1BBCBFF2C4D9D52360CE9F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 c</figureCitation>
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,
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<figureCitation id="535836C2F2DBD1989FF1BB1033E8920F" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 f</figureCitation>
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,
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<figureCitation id="161AC2BA52679385B3B0E72B4884F003" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">
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<quantity id="C41BCF9A46F3B7D10559EFFE7FB822EF" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</quantity>
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, n
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,
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<figureCitation id="2BD1FF7CC013DEED03C6A8D9C44D5350" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 h, v</figureCitation>
|
||||
,
|
||||
<figureCitation id="C6322BF941514CB3F8137B40418829B7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">
|
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22 p, p ’,
|
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, n
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</paragraph>
|
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|
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<taxonomicName id="3BFBE2C39A38CF1B9C714E4C877A807F" ID-CoL="FGDV" authority="Gusakov, 2004: 6" authorityName="Gusakov" authorityPageNumber="6" authorityYear="2004" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rusakovi" subGenus="Liothorax">
|
||||
<emphasis id="6F2F9141C423322A11CC458ED723E27E" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="49DD720BC28E5F8DD46CF4B8E3ECE609" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="14B89E83A985AA47DF37DB94F8B8CFC5" italics="true">rusakovi</emphasis>
|
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<comment id="2CFF2E27368963F4E3D9887662C6DE4F">
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<bibRefCitation id="D005895BA1507387AFBFCCC6212E2CA3" author="Gusakov AA" refId="B19" refString="Gusakov AA (2004) Novye vidy plastinchtousykh zhukov palearcticheskoj fauny. [New species of scarab beetles from the Palaearctic region]. Izdatel'skij dom Ejdos, Kaluga, 1 - 38." year="2004">Gusakov, 2004: 6</bibRefCitation>
|
||||
.
|
||||
</comment>
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</taxonomicName>
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<paragraph id="C68D0924CA74DF4A2EB002DFBE8EEC5B">
|
||||
<heading id="A6D67A50C6D25E5BB2787F0E9C78EA48" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="2A8CE819E79688CC943661D3B6843477">
|
||||
<specimenCount id="EA5DB39CBB2641B4A59461E5FF899195" count="2" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
|
||||
from the
|
||||
<collectingRegion id="8400CAC2C1E6DD1770BE4053D2D839C7" country="Russia" name="Orenburg">Orenburg</collectingRegion>
|
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oblast’,
|
||||
<collectingCountry id="599DC08D1547DDF816947AAF7FDCCB85" name="Russia">Russia</collectingCountry>
|
||||
(
|
||||
<named-content id="E4FB26A94043F8D46CB42D86168F9C13" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
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<collectionCode id="FFA25036636F3C63C48B0127C47A5B39" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EY4HK" type="material">
|
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<paragraph id="7D475BA7BB8323CCE37441804215D47A">
|
||||
<heading id="58DF04B7BF598803B4971452C507D5D4" reason="title">Additional material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="D1E96CC23E59B389B3444C1FE8298DF6">
|
||||
RU:
|
||||
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|
||||
,
|
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<collectingRegion id="9994252EFDE0C9E7BB76E4CED1242B76" country="Russia" name="Ul'yanovsk">Ulyanovsk</collectingRegion>
|
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,
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<geoCoordinate id="AF04EF649D17D70AEA0A73001875A585" degrees="54.056" direction="north" orientation="latitude" precision="55" value="54.056">54.056 ° N</geoCoordinate>
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,
|
||||
<geoCoordinate id="8071E3DC453CBC785C573AC393B5F34A" degrees="48.519" direction="east" orientation="longitude" precision="55" value="48.519">48.519 ° E</geoCoordinate>
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</named-content>
|
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(
|
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<named-content id="BFB2EF81AA0FB56DC88E38F8B6CE5AD3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
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<collectionCode id="AD0B69580852A96BC9E49CD5672ECBD1" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
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||||
</named-content>
|
||||
);
|
||||
<specimenCount id="C7C0CB5C5E77F323ACF2944957DCB9ED" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="453D72D17CD82CC636E50348DFB9AB64" country="Russia" name="Volgograd">Volgograd</collectingRegion>
|
||||
oblast’,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EN5HK" specific-use="{"type":"Point","coordinates":[44.500000,48.700000]}">
|
||||
<geoCoordinate id="9DD4AE6CDFD61AD93FBC3F4E8377929E" degrees="48.7" direction="north" orientation="latitude" precision="5555" value="48.7">48.7 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="908876044E52D778473C3F11057AD3AD" degrees="44.5" direction="east" orientation="longitude" precision="5555" value="44.5">44.5 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, Fastov. leg. Grebennikov. (
|
||||
<named-content id="98A103EB47901C83DD0E29A1497280A3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="0A1BAAE126EFD7374A9C30F67252F4D9" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="D0C443892BBB8128C480748EEE0C76B1">
|
||||
KZ or UZ
|
||||
<specimenCount id="314B84E8C706A6BD7A918209FF882886" count="1" type="female">1 ♀</specimenCount>
|
||||
“ Syr-Darja. / B. v. Bodemeyer. ” (
|
||||
<named-content id="F028F0277033614851C5EFE15F8B3747" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="7CC30681ED602863D264A1AE9EDA20A4" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E35HK" type="Differential diagnosis">
|
||||
<paragraph id="1EDEF8169C3FAC90841E328357AA3AB6">
|
||||
<heading id="F5A10CB70DA91619AD27A6571350A263" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="707345FE00D77ACB0ABA68753181A4C0">
|
||||
A conspicuously narrow parallel-sided species, comparable with
|
||||
<taxonomicName id="E97D62F61FE77F85ACEC36CB4D95D514" authorityName="Harold" authorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kraatzi">
|
||||
<emphasis id="DE2EDB2D0E7F5AA030E0C7DA4F2BC759" italics="true">A. kraatzi</emphasis>
|
||||
</taxonomicName>
|
||||
but larger, length
|
||||
<quantity id="30C33B3FF2B5B6753E6AD9B8C78A328B" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 mm</quantity>
|
||||
and base of pronotum completely bordered by a fine impressed line, this sometimes weaker medially (Fig.
|
||||
<figureCitation id="DC6C994C0E9538A4929561248170654A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 c</figureCitation>
|
||||
). Protibial spur of male strongly incurved apically (Fig.
|
||||
<figureCitation id="05B217CBF28F6362D976F330933DE6F5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 v</figureCitation>
|
||||
).
|
||||
<specimenCount id="A5BD59A5D9ECE04AA37ABF81A2766590" count="1" type="female">♀</specimenCount>
|
||||
with spermatheca larger (Fig.
|
||||
<figureCitation id="A4A7474F10150B2C1B70FEA0E36685D7" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">
|
||||
<quantity id="611561B709AFAEF9E0289FE561DC355B" metricMagnitude="1" metricUnit="m" metricValue="2.6" unit="m" value="26.0">26 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZ6HK" type="redescription">
|
||||
<paragraph id="FA9496A02D6837A3DF418826E837395C">
|
||||
<heading id="A6AE2A58AF32242A1B0154D232271619" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="751A7F311DCA72D50DE913E0E3E8CE49">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="8462DB43FADC37D1834F6819227EF0CD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 i</figureCitation>
|
||||
). Glossy black, length
|
||||
<quantity id="1486BAAD1461EB669022518EEA2BC2ED" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.7" unit="mm" value="5.1" valueMax="5.5" valueMin="4.7">4.7–5.5 mm</quantity>
|
||||
, width
|
||||
<quantity id="D916F38A5C9DADB96260AAAFAC8DB83F" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||||
. Head fairly closely punctate, sometimes slightly rugose on the frons and anterolateral parts of the clypeus, and occasionally rugose over all except the central part of the clypeal bulge. Double punctation scarcely apparent except when the punctation is sparser on the clypeal bulge. Clypeus emarginated anteriorly, edges of this emargination bluntly rounded (head and pronotum: Fig.
|
||||
<figureCitation id="C8FE247D01F0F4E4491F0D42DF612DB7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">
|
||||
<quantity id="329FA1ACCC274D268075F13774869D0F" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
|
||||
</figureCitation>
|
||||
). Frontoclypeal suture interrupted medially in
|
||||
<specimenCount id="B809864FDFE2A05102E3C2864992CEFB" count="1" type="female">♀</specimenCount>
|
||||
, in
|
||||
<specimenCount id="96C89D77A9396194E36816CCA395361C" count="1" type="male">♂</specimenCount>
|
||||
sometimes with a very weak tubercle, viewed with suitable lighting (Fig.
|
||||
<figureCitation id="B645B86BDE592FC9CFAB4847E9C60362" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 i</figureCitation>
|
||||
). Epipharynx (Fig.
|
||||
<figureCitation id="7730FC8B477D437B1A148A0E724E6705" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 i</figureCitation>
|
||||
) with anterior margin of clithra almost straight either side of the median tylus and the mesoepitorma scarcely darkened. Prophobae rather strong, grouped towards the margins of the mesoepitorma. Chaetopedia three or four each side. Pronotum highly arched transversely, weakly so longitudinally. Entire lateral margins visible from above (Figs
|
||||
<figureCitation id="AB54FD05E608ED674281E1F5F416E462" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 i</figureCitation>
|
||||
,
|
||||
<figureCitation id="69691973E81D60F1516A6D091C9C95E3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">
|
||||
<quantity id="DA2D2B7A1A201990E9D92E0D8A569498" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
|
||||
</figureCitation>
|
||||
). Surface with double punctation, this often close though sometimes sparser on the disc. In some specimens the punctures are deeply impressed, so the pronotal surface does not appear smooth. Basal margin entirely bordered, the border rather heavy (Fig.
|
||||
<figureCitation id="29084C5F72D9C33FD0E6DE78E472E322" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 c</figureCitation>
|
||||
). Elytra noticeably elongate, parallel-sided. Striae narrow but rather deep, strial punctures separated by ca 4 × their diameters. Interstices 6–8 × the width of the striae, with distinct sparse fine punctation and fine reticulation (Fig.
|
||||
<figureCitation id="BF6A3105F53A90246D48407A484CB0B4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 f</figureCitation>
|
||||
). Apex bluntly rounded. Metaventrite (Fig.
|
||||
<figureCitation id="683303A2244539AD8ACAB00696F97466" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">
|
||||
<quantity id="C4060086E179FBAA3BB0C4E67E5EBF90" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
) with median diamond- shaped plate fairly strongly punctate and no obvious sexual dimorphism. Legs with longer spur of mid tibiae slightly longer than the basal segment (Fig.
|
||||
<figureCitation id="C38DC50AFF4B6AA4A61326236D3D2D86" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 h</figureCitation>
|
||||
), and in males the spur of the anterior tibiae is distinctly hooked (Fig.
|
||||
<figureCitation id="A65AAE19066602C189CC7CB9A67F379A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 v</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="012164FF37B7A118BF803037643F5E7F">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="961EA71E795467B562D8E1C679A7CC33" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 p, p</figureCitation>
|
||||
’) length ca
|
||||
<quantity id="1CA0F6241E4FE40981ED3596C08D7E92" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
, parameres relatively short and blunt-ended, somewhat downturned apically. Endophallus with scales but no hairs or bristles.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ESBAM" type="remarks">
|
||||
<paragraph id="60C81B5729902C52EDFBBA9220C6725D">
|
||||
<heading id="19ADDF39C08444C6855FFA32B39F61C7" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4B20F95518D297C576769908998D53D3">
|
||||
Distributed mainly in southern
|
||||
<collectingCountry id="98CB042226EFC702D46B508712C7BFB9" name="Russia">Russia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="EC485BF06482FC3A10B8B5D758E3CD6F" name="Ukraine">Ukraine</collectingCountry>
|
||||
, and
|
||||
<collectingCountry id="14FE5E6BDF4F1E5093D63D123CAE3354" name="Kazakhstan">Kazakhstan</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="36B35940EC1AA454170EF40185DDE9FE" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b, c</figureCitation>
|
||||
).
|
||||
<bibRefCitation id="38DCBDBB0ECD5E56984A544E19C6630A" author="Gusakov AA" firstAuthor="Gusakov" refId="B19" refString="Gusakov AA (2004) Novye vidy plastinchtousykh zhukov palearcticheskoj fauny. [New species of scarab beetles from the Palaearctic region]. Izdatel’skij dom Ejdos, Kaluga, 1–38." year="2004">Gusakov (2004)</bibRefCitation>
|
||||
lists the
|
||||
<typeStatus id="64548D007E0930FABDFE2A45919D1C68" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="A73EB2F838D65BFED7ADA34480F4E851" count="14" type="generic" typeStatus="paratype">14 paratypes</specimenCount>
|
||||
as having been collected in a dried-out rivulet at Ilek,
|
||||
<quantity id="72455446C9D6F934C5AA9EC1A3E404A7" metricMagnitude="5" metricUnit="m" metricValue="1.2" unit="km" value="120.0">120 km</quantity>
|
||||
SSW of Orenburg on the southern Urals, with two further
|
||||
<typeStatus id="F22B18B79F3373D1048C8DEA03E9B1BB" type="paratype">paratypes</typeStatus>
|
||||
from the Rostov and
|
||||
<collectingRegion id="7D3C9217492229605412455EFFC89EF4" country="Ukraine" name="Kharkiv">Kharkov</collectingRegion>
|
||||
districts of
|
||||
<collectingCountry id="972AE048B76F9A8884B6FDFB9663DEAE" name="Ukraine">Ukraine</collectingCountry>
|
||||
and four more from the Temirsky region and Uralsk oblast’ in
|
||||
<collectingCountry id="9EB703242FC9CF4B043F5DF2A43D2C70" name="Kazakhstan">Kazakhstan</collectingCountry>
|
||||
The Ulyanovsk specimen (
|
||||
<named-content id="8C5DA249CABE12F4AB0BEA97D93F792D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="745E977EDC14FDD1D81D7123BAD6E922" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
) is labelled as having been taken in a damp saline area (solonchak).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
387
data/3E/F5/EA/3EF5EA588BCC5C07A9036A941DDCB1D7.xml
Normal file
387
data/3E/F5/EA/3EF5EA588BCC5C07A9036A941DDCB1D7.xml
Normal file
|
|
@ -0,0 +1,387 @@
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<mods:titleInfo id="65C8101C6133C69AB415583CD9BD748F">
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<mods:title id="505915838034A50CF32E85A21E4447D1">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:affiliation id="E424E521E498443076F577EE0821D168">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:affiliation id="BE87C5D8AA095C5E729D257BB4623CF5">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:affiliation id="B3E4BC486FD8E2B56B36F357082E7D62">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<taxonomicName id="D892D45A97BA697318A7EFA78BC878A3" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="6B9E7E7D3C22199741D2A6508CFCFFF5" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="A7830C7C998E54570E60D4816D10BD5D" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="AB80C8B85674BF7B9A79DC5D2B134C39" italics="true">alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="39960CE37534803D23CE79CD1A729EF0" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="F5B96226186D63FCCE554C1E1A99CF3C" type="description">
|
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<paragraph id="5122A4712936D7FE7E8579FDC76277A1">
|
||||
<figureCitation id="BEF4A433321BDECB42D77D2E68BD6144" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">Figs 3 e – g</figureCitation>
|
||||
,
|
||||
<figureCitation id="A557E7BCD0EBB50F5A57076B9AF2A522" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="2C61D37C4DE54D0B67A49ABAFC942B43" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="7B3FB13D97FD2F0CCD05EB10CDE69102" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 k, l</figureCitation>
|
||||
,
|
||||
<figureCitation id="D1BFADCD316532A8F8378978571A9A1C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">
|
||||
<quantity id="E29EAEEDB9ACADE1515FDE17D0E3FEB1" metricMagnitude="4" metricUnit="kg" metricValue="1.8" unit="t" value="18.0">18 t</quantity>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="74D40ED1CCF0B0F9382C8157EE5171A9" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 o</figureCitation>
|
||||
,
|
||||
<figureCitation id="3C89E3104F012D57EC75D3A1D97456A0" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 n, o</figureCitation>
|
||||
,
|
||||
<figureCitation id="3FE7F324238E0EC8584D577668312252" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 e – j 25 d, 26 b’, c’</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ER6CM" type="type material">
|
||||
<paragraph id="99CC3C90151BD3DD2B654A796B3F09F0">
|
||||
<heading id="C81D460E7EF6E6402937151BAF4B8D21" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B49927E8D4A827AC80B500CDA0327449">
|
||||
<materialsCitation id="0FD86841500B845ABBCE33F14BA6AB39" collectingDate="1992-06-18" collectionCode="NHMUK" collectorName="A. Ballerio." country="Turkey" location="Ovitdagi" specimenCount="1" specimenCount-male="1" stateProvince="Rize" typeStatus="holotype">
|
||||
<emphasis id="B50DB534B44A023F6BA129C54735DCC0" bold="true" italics="true">
|
||||
<typeStatus id="1315D8CB88F6418F0F281899F758D730" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="695B7256699F6B90043D2766CAB615AE" count="1" type="male">♂</specimenCount>
|
||||
, “
|
||||
<collectingCountry id="A3B66E4EFAD8BCA44232E610368737AF" name="Turkey">Turchia</collectingCountry>
|
||||
– Vil.
|
||||
<collectingRegion id="71B1EA5D2CAC1F99608507AEF626F03A" country="Turkey" name="Rize">Rize</collectingRegion>
|
||||
,
|
||||
<location id="519FC1F7974D5FC76DE1501B0EB67595" LSID="urn:lsid:plazi:treatment:3EF5EA588BCC5C07A9036A941DDCB1D7:519FC1F7974D5FC76DE1501B0EB67595" country="Turkey" name="Ovitdagi" stateProvince="Rize">Ovitdagi</location>
|
||||
gecidi mt 2600,
|
||||
<date id="6F4313DA2457CBBDC42992EFE2B0BED5" value="1992-06-18">
|
||||
<collectingDate id="2E7A4A2370B0C7DA11CEF6AF9BB4421B" value="1992-06-18">18 - jun- 1992</collectingDate>
|
||||
</date>
|
||||
. leg.
|
||||
<collectorName id="7364B405DADFC58B216C474D4349CF27">
|
||||
<tp_taxon-name id="20EBEA493BC772B1605CA1E679836CC5">
|
||||
<emphasis id="88983157ABADDAB64D8AE882745F4B4C" italics="true">
|
||||
<tp_taxon-name-part id="9D8F8E0B42B623D423803E4AA14A336D" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Ballerio.
|
||||
</collectorName>
|
||||
” (
|
||||
<named-content id="3AE8F6839E5775E673A1C45A8EADC654" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="050F06799A175BBB5CDBBA5F9F677D7E" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="91CF39186A00BC537076BCC7839A3297" bold="true" italics="true">
|
||||
<typeStatus id="C7ED1BC2004A04285EB6136073737BFE" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="CE335098652FF51A3BC19C8294B7AB5F" count="4" type="male">4 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="91C2A60A4D350E55279F88C5E5114316" count="7" type="female">7 ♀♀</specimenCount>
|
||||
, 17 unsexed, data as
|
||||
<typeStatus id="A73636FECA829AA1FE8C4309A68138D1" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
<specimenCount id="51D554CB339546E54689C8145C1FB4EB" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="4048B02C1825E805B24367A3F032CD0F" count="1" type="female">1 ♀</specimenCount>
|
||||
, 1 with mouthparts dissected (
|
||||
<named-content id="A4877366761F4489A6F28F30864E4D64" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="806D64D14403E9DF4E0771E2F187F06F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<specimenCount id="1358381C3BCB74D50CAAC0861A1E8B93" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="7A113594A09B6896084851E33E30767D" count="2" type="female">2 ♀♀</specimenCount>
|
||||
, 14 unsexed (
|
||||
<abbrev id="ABBRID0ENADM" xlink_title="Alberto Ballerio Collection">
|
||||
<collectionCode id="3075AC01E00A6987B219560B73612A98">AB</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
<specimenCount id="22BBCCEFFCB7827AD85A287C7F86516C" count="5" type="female">5 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="93D2DAA365187DCF1E262D64BF23B436" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<abbrev id="ABBRID0ERADM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="7D029A32E5934D6B4B42F9F96038C2C4">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVADM" type="material">
|
||||
<paragraph id="42B43E515FE2B55DA4E6BC7780E9FCBE">
|
||||
<heading id="CE8327344311EFAA1FB72C00717EC673" reason="title">Other material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="918EF982FCB1604DC035C61C43B8C89E">
|
||||
<specimenCount id="BE8CC664B645F67A437B03E22DF19A4C" count="4" type="generic">Four specimens</specimenCount>
|
||||
, unsexed, Ulu Dag b. Bursa As. M. occ. V: 1968 Schweiger,
|
||||
<abbrev id="ABBRID0E2ADM" xlink_title="Muséum d’histiore naturelle, Genève">MNHG</abbrev>
|
||||
, may belong to this species. A male from central
|
||||
<collectingCountry id="DCC692623396835CF2DCC58D300CE738" name="Armenia">Armenia</collectingCountry>
|
||||
, Selim Pass
|
||||
<quantity id="42A2E6A3AB735107EC92C1DD668D161C" metricMagnitude="3" metricUnit="m" metricValue="2.35" unit="m" value="2350.0">2350 m</quantity>
|
||||
, coll. Ziani is shown by its parameres to belong to the
|
||||
<taxonomicName id="22E499DF779267E7C7D033A59AD90798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="51F41304FCF2E6743554A387560D6A95" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
species complex, and it agrees with
|
||||
<taxonomicName id="6FB4B1440E744722495C5B5618C19A12" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti">
|
||||
<emphasis id="98B0A1B018B83D6BE32578E729108730" italics="true">A. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
in its black to blackish-brown appendages and its small size, length
|
||||
<quantity id="EE74B5793897405C580CD6A767FDFB01" metricMagnitude="-3" metricUnit="m" metricValue="3.9" unit="mm" value="3.9">3.9 mm</quantity>
|
||||
. The epipharynx of this specimen has the apophobae less regular but the darkened area is narrowed in its basal 1 / 4.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVBDM" type="Differential diagnosis">
|
||||
<paragraph id="5DE366C76F2C9A8F2C6E500C6C3F1EDF">
|
||||
<heading id="1EEE7307B48224EFC056A695B1A90194" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DFB86B24FB3739CDF157008CB114B486">
|
||||
<taxonomicName id="21868A06EE8A9FE1F275B04572B34FAE" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="E90A02BFD703DFA1AB2B99088E06ED60" italics="true">Aphodius alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="2F7501775719275E6C813C04354D09AE" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is the second
|
||||
<taxonomicName id="4A296D5E9BB602A310F8888C873AD3EC" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="3527E186623443D6EC7A130F218C4FFE" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
group species recognised in
|
||||
<collectingCountry id="2F0C59E5C91725C3632E9F5F199A7C5B" name="Turkey">Turkey</collectingCountry>
|
||||
, and as such requires comparison with
|
||||
<taxonomicName id="324A58A7247878F0240A0238E5AD69C1" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="63EAF9BD8157DC096541AE0453AF8EF0" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
. The most striking distinctions are the size ranges: in
|
||||
<taxonomicName id="923D5BC380AF4007825C1134550F8979" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="FDD9708E9D7EDA1FD2908332CE03DDC0" italics="true">A. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="915FAA9E80C2ED0FC61D01500D3C01FD" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<quantity id="70AD679D520718FF3307F19AB9B4F742" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.1–4.9 mm</quantity>
|
||||
, in
|
||||
<taxonomicName id="CB6F96BC4023E852741A496C8B40B139" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="3F0AAE577A014E0BB4D673F65A522CF0" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
<quantity id="70B807D21D5EC00970F81492015A95C3" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.8" metricValueMin="4.5" unit="mm" value="5.15" valueMax="5.8" valueMin="4.5">4.5–5.8 mm</quantity>
|
||||
and the colour of the appendages: in
|
||||
<taxonomicName id="46BBB866A4ABBA925C5D4E7B9231C3BF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="FB2A96F5D9F2A8F5C7B6190FA965A0FC" italics="true">L. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4C27C572C0E1454E40C9F0EAD9566C9E" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
the legs are blackish brown to very dark brown (Fig.
|
||||
<figureCitation id="39D7FE093FC58429197F3B1BC67D5E7E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e – g</figureCitation>
|
||||
), in
|
||||
<taxonomicName id="217C03F57EB627738E238DB359912DA2" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="385E3BA9A004F9CFC3A379C130FBD3FC" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
mid brown (Fig.
|
||||
<figureCitation id="FB2F4ADF12E95D587E1593CD08EFA421" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 c, d</figureCitation>
|
||||
). The maxillary palpi are metallic black to very dark blackish brown in
|
||||
<taxonomicName id="BB71C4789AD3D29D5A3B147BE536E054" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="662807FD2D8FBB20262B399A135CE839" italics="true">A. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="8AFCD2F06D58C4B3F5323923FE2FE635" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
as against mid brown in
|
||||
<taxonomicName id="DE03F4436DFA088F3160D5C708C0B5C5" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="057AC98BC8DB5EAF1DC6E134E405663F" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
(metallic black in the
|
||||
<collectingRegion id="4B4D1323892A51D8FE3ADD755D53BA3D" country="Turkey" name="Artvin">Artvin</collectingRegion>
|
||||
specimen). In
|
||||
<taxonomicName id="50C153D7604DF1CEFB7CCA0F4571E73F" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="E03CC052D0EF74FFE578CF8EFD869E9C" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
there is a sexual dimorphism in the head: the males with the central area of the clypeus clearly more strongly bulging upwards than in females, and the outer margin of the genae almost continuous with the clypeal margin in the male, but distinctly angled outwards in the female. In
|
||||
<taxonomicName id="EC00DD86A139C1BB813D055BBEF3824E" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="716D71226E7AAA6AF0CF456BC472F6FB" italics="true">A. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="982960E91C075B15885ED6EB060B172E" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
the central area of the clypeus of both sexes bulges upwards strongly, and the outer margin of the genae is almost continuous with that of the clypeus. The epipharynxes of the two species show some clear differences: in
|
||||
<taxonomicName id="88464CDC572B58163CF93BB96463D16D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="0F3B7C4009B458ED0B6A3B8693C5C63D" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
the median darkened area is broadly triangular, widest at its base, there are six or seven chaetopedia each side, and the rows of apophobae are arranged in narrow bands, ca two bristles wide, while in
|
||||
<taxonomicName id="00288B3FEB2CE30E58E3A0F3A27F942B" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="8DF8509920783649F0C104FCAA3F075B" italics="true">A. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="32954D9C017989C0FC0A4BF969A183D9" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
the darkened area is narrowed over its basal 1 / 3, there are four chaetopedia each side and the rows of apophobae are as single lines (Fig.
|
||||
<figureCitation id="B16B844DC71A6E6CC6245A8CFFFC8A0B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 c, d, k, l</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWGDM" type="description">
|
||||
<paragraph id="5304CF7D4551EE4FAD33BC3C91DE1FB9">
|
||||
<heading id="398AA2312AE69334D61F4E3CA301FF24" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="29E3A428707B86630A395FD83B87FDA0">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="9BDF57D6560B83743998BB61CE0C1EC6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e – g</figureCitation>
|
||||
). Length 4.1–4.5 (
|
||||
<specimenCount id="AE36495769E9F097057BBBC8505C230F" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="2C2DFE3F42280956C8CE5A2CD5694004" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.1–4.9 mm</quantity>
|
||||
(
|
||||
<specimenCount id="050432AFCCC3221478453F794032531E" count="1" type="female">♀</specimenCount>
|
||||
), width
|
||||
<quantity id="EDD909A6C46799D5DC3767DA01BD2E8A" metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="1.95" metricValueMin="1.9" unit="mm" value="1.9249999999999998" valueMax="1.95" valueMin="1.9">1.9–1.95 mm</quantity>
|
||||
(
|
||||
<specimenCount id="ABB7746C4372F7CC9B8EBE2B14DF9170" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="722EF87D629545DE34942CEB6FB78615" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" unit="mm" value="1.95" valueMax="2.0" valueMin="1.9">1.9–2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="C5FC0423090E4214629CC7D825F20D3A" count="1" type="female">♀</specimenCount>
|
||||
). Head (Figs
|
||||
<figureCitation id="6DF6A31FF7BDA4ADBC4E8DF546E91D26" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e – g</figureCitation>
|
||||
,
|
||||
<figureCitation id="8B904C9AF467E5E4F892ECE9E512C223" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 c, d</figureCitation>
|
||||
) black, anterior clypeal margin narrowly dark brown, excised medially, angles at sides of excision bluntly rounded. Central area of clypeus bulging upwards in both sexes, punctation fine, moderately dense, with some variation in strength, this not dependant on the sex of the specimen. Frontoclypeal suture fine and straight-transverse over central 1 / 2, with no trace of a median tubercle, then angled forward to meet the sides just anterior to the genae. Lateral margins of genae either continuous with those of clypeus or with a very slight outward angle, without sexual dimorphism. Maxillary palpi glossy blackish brown, antennae dark brown. Epipharynx (Fig.
|
||||
<figureCitation id="C1B0AF4C9B333A193E550DFB43727CD8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 k, l</figureCitation>
|
||||
) with the central tylus strongly projecting anteriorly and the anterior margin of the clithra clearly excised either side. Central darkened sclerotised epitorma triangular in apical 2 / 3, then the sides curved inwards so that the width at the base of the darkened region is ca 3 / 4 of its maximum width. Acropariae virtually absent, chaetopariae well-spaced, four each side. Surface of gymnopedia covered with small tooth-like asperities. Prophobae fairly strong, clustered at the edges of the median sclerotised mesoepitorma, apophobae fine, arranged in a single line outside the chaetopariae.
|
||||
</paragraph>
|
||||
<paragraph id="F1C1B9814281FDE4153DA55A80DD7302">
|
||||
Pronotum hemicylindrical, highly arched transversely, scarcely at all longitudinally. Entire lateral margins visible from above. Surface with double punctation, some specimens (male and female) have the punctation heavier with the larger punctures separated by at least twice their diameter, sparser on disc (Figs
|
||||
<figureCitation id="F8098C23F1C79AB3E6331468A7E3DA42" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e, g</figureCitation>
|
||||
,
|
||||
<figureCitation id="2489111E63F203E105ED351096CEB5DD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 d</figureCitation>
|
||||
), in others the punctation is sparser and finer (Figs
|
||||
<figureCitation id="F4B0C75198A0BF0593D3482BF0FB9A63" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 f</figureCitation>
|
||||
,
|
||||
<figureCitation id="7D0167B14F1BF1B5C011AB1F1B9B7A28" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 c, d</figureCitation>
|
||||
). Posterior margin completely but narrowly bordered (Figs
|
||||
<figureCitation id="E4D0CEF25B0E224351976C8C5D7C1B05" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e – g</figureCitation>
|
||||
,
|
||||
<figureCitation id="B0D87DE90B6BC5FA58AAC8CEFE988ADB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 c, d</figureCitation>
|
||||
), the border not widened medially, continuous round the posterior corners. Scutellum pentagonal, elongate, glossy, sometimes with a few punctures medio-basally.
|
||||
</paragraph>
|
||||
<paragraph id="B79992BD7384F4DE12E9AD826CE877F7">
|
||||
Elytra black, interstices flat, 6–8 × width of the striae, with fine isodiametric reticulation and sparse fine punctures. Metaventrite (Fig.
|
||||
<figureCitation id="04833D539F3602C48E72341BB7568228" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 n, o</figureCitation>
|
||||
) rather finely punctured in both sexes, flattened over median diamond-shaped plate, sometimes with an impressed median line. Legs dark blackish brown (Fig.
|
||||
<figureCitation id="3C4EB562C17085537BF7DA5A71C43380" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 e – g</figureCitation>
|
||||
), basal segment of mesotarsi elongate, slightly longer than the longer tibial spur (Fig.
|
||||
<figureCitation id="EDA7E9403B1B58BD6A8B98C0C9E7DE6B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">
|
||||
<quantity id="83068B5F127FB04F321C7F71B5759606" metricMagnitude="4" metricUnit="kg" metricValue="1.8" unit="t" value="18.0">18 t</quantity>
|
||||
</figureCitation>
|
||||
). The legs and metaventrite Fig.
|
||||
<figureCitation id="64D365754A7324E0826123E5AA3F7CC7" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 o</figureCitation>
|
||||
appear unnaturally pale brown due to the intense lighting.
|
||||
</paragraph>
|
||||
<paragraph id="392B081013D9088FF4119AA591292FD8">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="2682E436CB1967CECDB125ADD1FD63BF" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 e – j</figureCitation>
|
||||
) Aedeagus length ca
|
||||
<quantity id="656B9BFECC9D2B5945B83682A6D4FAE7" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
, parameres length ca
|
||||
<quantity id="FC71400D1A2929653DC75D852EDE9188" metricMagnitude="-4" metricUnit="m" metricValue="4.4" unit="mm" value="0.44">0.44 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="6C50E541D7F638D8439D2EC7529A825E" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" unit="mm" value="0.59">0.59 mm</quantity>
|
||||
. Length of tooth-field ca
|
||||
<quantity id="7D9CE1B5F3ECD5B928152DCD0A831C9F" metricMagnitude="-4" metricUnit="m" metricValue="5.8" unit="mm" value="0.58">0.58 mm</quantity>
|
||||
, length of longest teeth ca 54 µm. Parameres in lateral view weakly to moderately downturned apically (Fig.
|
||||
<figureCitation id="5A93AD7BA6F32B153193138AD2FFBDAE" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 h – j</figureCitation>
|
||||
). In dorsoventral view sensory area either not visible (Fig.
|
||||
<figureCitation id="54A09AC5C25C56640DC3F47F96E9B37F" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">
|
||||
<quantity id="9E1CFAB6795603D61AAFDBC10DF8EC07" metricMagnitude="-2" metricUnit="kg" metricValue="2.4" unit="g" value="24.0">24 g</quantity>
|
||||
</figureCitation>
|
||||
, the
|
||||
<typeStatus id="8C049856F7F451256164D6FC415016D9" type="holotype">holotype</typeStatus>
|
||||
), or visible after squashing as a curved band round the paramere apex (Fig.
|
||||
<figureCitation id="CE8DAC32068BBF5CB92580309EB9B0E3" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 e, f, a</figureCitation>
|
||||
<typeStatus id="DC061A582CA331B982E23792EB0F7CB2" type="paratype">paratype</typeStatus>
|
||||
). The parameres struts are clearly less darkened in the
|
||||
<typeStatus id="C3ED8F0C23CB94DD8D2E344F84F4D53B" type="paratype">paratype</typeStatus>
|
||||
, suggesting that the aedeagus is less fully hardened, which would render it more prone to squashing.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EKJDM" type="remarks">
|
||||
<paragraph id="D78BC8D95F4DA79D8B51EFF4C8ACB1FF">
|
||||
<heading id="9E7EF003027DEA623B30DCBB93DFAE29" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="C8C682F2D3BEE5DCF8C343AAA1C8FA12">
|
||||
Apparently widespread on high ground in
|
||||
<collectingCountry id="E060AC2244D234B0B35B9F398B0EB59A" name="Turkey">Anatolia</collectingCountry>
|
||||
and possibly extending into
|
||||
<collectingCountry id="877D6F55DB92605F99340253AAFE0AFE" name="Armenia">Armenia</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="5594852E19A52F9154CCAA6CDC39DCE7" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
). According to
|
||||
<emphasis id="9F9CB61CF1591F889D6D9222ADF24981" italics="true">A.</emphasis>
|
||||
Ballerio (pers. comm.,
|
||||
<date id="30FA95A83A8E03225EAAE26A8578D298" value="2023-06-16">16. vi. 2023</date>
|
||||
) the
|
||||
<taxonomicName id="DFBBD6B94896F1FC381EA794FC147F38" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5EFF5B6A1F4110030EE710BE7264488D" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
were found on snow patches and in puddles at the sides of the road passing through Ovit pass, a typical alpine prairie (Fig.
|
||||
<figureCitation id="4056593B40DE5CF4616775113881D3B3" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 e</figureCitation>
|
||||
). They were mixed with fewer
|
||||
<taxonomicName id="A75F8E62889D33B262DF57C47CF27A42" authority="Pittino & Ballerio, 1994" authorityName="Pittino & Ballerio" authorityYear="1994" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ovitensis" subGenus="Neagolius">
|
||||
<emphasis id="0A869695BB770C0C54AAD264602B252C" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="F91785E5B56D8AA7C65A359853FF2A7E" italics="true">Neagolius</emphasis>
|
||||
)
|
||||
<emphasis id="CBE6FD177B64CA14E83D827F9DE594A8" italics="true">ovitensis</emphasis>
|
||||
Pittino & Ballerio, 1994
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
273
data/3F/6E/65/3F6E65BAE8115765BEA7804ADA8107F4.xml
Normal file
273
data/3F/6E/65/3F6E65BAE8115765BEA7804ADA8107F4.xml
Normal file
|
|
@ -0,0 +1,273 @@
|
|||
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<mods:title id="06FE67B68B83BF069D534E6BD55BF143">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:affiliation id="FA5B30C811E9FF2CEAE69ABB6C81C7FD">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<taxonomicName id="5BB5D02EB8C413E168C71B89D49CE88D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chellala" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="F315DDA433D442307E578A732A2B018F" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="A76B3A1206770CB7683F8E6818141DFB" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="EF8E8CD4222A24C3CB87DFD180420208" italics="true">chellala</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="1AEF8461F690711851C34145C01BEEB7" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
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|
||||
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|
||||
<subSubSection id="22A9E97A00E5E5424F89B73E72956CB1" type="description">
|
||||
<paragraph id="C725C8174CC0EE23802E37853C910899">
|
||||
<figureCitation id="F19DA2E80D15849BA78FB3737B15A539" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">Figs 2 h</figureCitation>
|
||||
,
|
||||
<figureCitation id="E4B3D3DA98153DA34BB2B77F8206E27F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 l</figureCitation>
|
||||
,
|
||||
<figureCitation id="F6C5701194EDA7868F8754DBE2AC4388" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 k</figureCitation>
|
||||
,
|
||||
<figureCitation id="A867FC1E9B1E227EF06AEDA19CB2BADC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 e</figureCitation>
|
||||
,
|
||||
<figureCitation id="E1C17F081FE452B764256D364129E3CD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 e</figureCitation>
|
||||
,
|
||||
<figureCitation id="404AB5D542F0005791B4E62E1C0A2C81" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 d</figureCitation>
|
||||
,
|
||||
<figureCitation id="A79A5782408B245C981442BAAACA5B15" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 e, f</figureCitation>
|
||||
,
|
||||
<figureCitation id="EDEF9F3FD756F0EECA0C6B0F1CEB5B92" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 s – s ” ’, t, u, 25 l, 26 l</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1IHK" type="type material">
|
||||
<paragraph id="A84ED08ADB8B2D56E05F96F253A1FE2E">
|
||||
<heading id="0482244EEB6C4B6FC8F96FC66CB9AD19" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="0701B06167C503AA0BDB1CB5E936D61F">
|
||||
<materialsCitation id="33EB45266113D3EF33958EC6B58B3281" collectionCode="NHMUK" collectorName="De Vauloger. Nevinson" country="Algeria" latitude="33.074" location="Chellala" longLatPrecision="72" longitude="-0.123" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis id="26A2F836DB69E1FFB38EA4922BD82129" bold="true" italics="true">
|
||||
<typeStatus id="3BBCB36D393F92C598CF58C86D22E040" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="8BB44BAC307AE26E88ED655FCC6412AD" count="1" type="male">♂</specimenCount>
|
||||
:
|
||||
<location id="57B5192E548F8D24D53B592FA666FBA9" LSID="urn:lsid:plazi:treatment:3F6E65BAE8115765BEA7804ADA8107F4:57B5192E548F8D24D53B592FA666FBA9" country="Algeria" latitude="33.074" longLatPrecision="72" longitude="-0.123" name="Chellala">Chellala</location>
|
||||
,
|
||||
<collectingCountry id="F1875712377F968448B07DAF2A7373D4" name="Algeria">Algeria</collectingCountry>
|
||||
(
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIJHK" specific-use="{"type":"Point","coordinates":[-0.123000,33.074000]}">
|
||||
<geoCoordinate id="038E5F3BD2715E55A56911E60494A194" degrees="33.074" direction="north" orientation="latitude" precision="55" value="33.074">33.074 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="AB18790A4EC4739AA0A345BEDB134450" degrees="0.123" direction="west" orientation="longitude" precision="55" value="-0.123">0.123 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
)
|
||||
<collectorName id="14ED9386D51B91B29E06501D5674755E">De Vauloger. Nevinson</collectorName>
|
||||
coll. 1918 – 14 (
|
||||
<named-content id="60CE04FEE64D46EF5F47FBA61E8A0FDB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="4E7B2412351083736925EDFB6E01C1A4" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="A4F408F792486881069F93A31B80FEDF" bold="true" italics="true">
|
||||
<typeStatus id="8CD5D0AFAE9C2ED22F5FA0E154BD7313" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="41AF46248527E2F243047E8E1953CA45" count="8" type="male">8 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="ACFA7FB49BB3933A9EE18F63D888DB2B" count="14" type="female">14 ♀♀</specimenCount>
|
||||
, 1 unsexed (lacking head), data as
|
||||
<typeStatus id="3D88423F4EB6E021EB27ED31002B0812" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
<specimenCount id="3A9218F1CF278402EC5B7027517F225F" count="1" type="male">1 ♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="180A45038ACA508AE0F4460317B87E8C" count="2" type="female">2 ♀♀</specimenCount>
|
||||
have the labels “ Prov. D’Alger. Chellala 1895 de Vauloger ”, these presumably the basis of the English labels on the other specimens.
|
||||
<specimenCount id="7ED41302AEEF32CA2173B53C72A2CC42" count="9" type="male">9 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="ED8793E691289B07C6D9C8117EDE322D" count="3" type="female">3 ♀♀</specimenCount>
|
||||
, Taguin (near Zmalet El Emir Abdelkader,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXJHK" specific-use="{"type":"Point","coordinates":[-1.402000,35.221000]}">
|
||||
<geoCoordinate id="A6CBF71C98CB8F4311ACDC6A7AE07FA6" degrees="35.221" direction="north" orientation="latitude" precision="55" value="35.221">35.221 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="4AEBAA549B116FF0F570CC44403D1F00" degrees="1.402" direction="west" orientation="longitude" precision="55" value="-1.402">1.402 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
),
|
||||
<collectingCountry id="23570828DB1B39166837F72FC1CA8137" name="Algeria">Algeria</collectingCountry>
|
||||
, De Vauloger. Nevinson coll. 1918 – 14.
|
||||
<named-content id="67233C5395D96BD6AE39C29B959B0F18" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
|
||||
. As with the Chellala material,
|
||||
<specimenCount id="5B37EC5DE3BA5B23E6D15E311F3162CB" count="2" type="male">2 ♂♂</specimenCount>
|
||||
have the French label “ Prov. D’Alger Taguin 1895 de Vauloger ” (
|
||||
<named-content id="E1C86702825EFE73F0F6BC6FDC168FAE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="11A128589CE50D301507CFED1D4896F5" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEKHK" type="Differential diagnosis">
|
||||
<paragraph id="A5D2812928C14300DD37254E6F6BBBEF">
|
||||
<heading id="D3E79B3BCD4625C57026A39A38D2F016" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="05236B811389FAAD6EE3CF344884C45F">
|
||||
Closely resembling dark specimens of
|
||||
<taxonomicName id="004548EDACD2FA436949F453EC62B989" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="04B3EA8C412C7389A6F865BC33C1F246" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
but basal margin of pronotum (Fig.
|
||||
<figureCitation id="AEB3DE2573C4F1D17D89EE825497DF16" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 e</figureCitation>
|
||||
) more broadly interrupted than in
|
||||
<taxonomicName id="EB291C5951360FDE8A93BA139464AEED" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="096991B3CA3422A30A9EA6AB7D844518" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="49CAF8467DEDF3F61AFEF06F2F5DC1E3" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 c, d</figureCitation>
|
||||
). Endophallus of aedeagus, when extruded, with the scale-patches (Fig.
|
||||
<figureCitation id="8AF399AACED9282FB696BA2CF1FB54AE" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 q</figureCitation>
|
||||
’, s ” ’) clearly different from the dense bristle-field of
|
||||
<taxonomicName id="15189A6731763D05306F3DCAC0EE15EA" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="6DFE270F0286EA4D0A40A0973B2A4049" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="A51421313F7285871FCDC9AD59869999" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 q, q</figureCitation>
|
||||
’).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2LHK" type="description">
|
||||
<paragraph id="1B3C606AA2674FBB37ABC54AFF0156CF">
|
||||
<heading id="97606A4440A1ED931C44B82220E098A8" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8235C3C6CE53375D2128B7283B3C86DB">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="74391BE781B36E924B60DF95698A69D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 h</figureCitation>
|
||||
). Length
|
||||
<quantity id="60287C6B1DF919B1031BA14ECA0DBFE0" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.5" metricValueMin="3.6" unit="mm" value="4.05" valueMax="4.5" valueMin="3.6">3.6–4.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="CE71AF9125C6AEB97EC117A4B27C92E6" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="9980187B4E0C79F0F0126873A076B391" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.5" metricValueMin="3.7" unit="mm" value="4.1" valueMax="4.5" valueMin="3.7">3.7–4.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="887C642114889151D24EA140C0A70FEB" count="1" type="female">♀</specimenCount>
|
||||
), width
|
||||
<quantity id="2AE1674A5A0EC2932CFD3A4FBD0D2296" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="8878E61BEBE123D4B6FF66C9A968DBE5" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="0662941D4F0028ABB2834C3FD9286EDC" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
|
||||
(
|
||||
<specimenCount id="62F9497FC20FC150D2CA85B2CCA9A08B" count="1" type="female">♀</specimenCount>
|
||||
). Black, antennae, maxillary palpi and legs dark brown. Head: glossy black with anterior from in front of the genae dull brown, this colour merging into the general black ground colour. Anterior margin of clypeus emarginate medially, sides of the emargination rounded. Surface glossy with no trace of reticulation but with sparse double punctation, the larger punctures scarce medially. Disc of clypeus raised as a hump. Frontoclypeal suture fine, without tubercles, complete, angled forward just mediad of the inner margin of the eyes, running to anterior end of genae. Genae protruding in front of eyes, rounded, no angle between anterior end of genae and clypeus. Head and pronotum (Fig.
|
||||
<figureCitation id="52FF0A298E24DC73B51CBAFEE18583A2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 l</figureCitation>
|
||||
): epipharynx (Fig.
|
||||
<figureCitation id="0D97BD2DE50ED43F96B4D059E98686D4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 k</figureCitation>
|
||||
) similar to that of
|
||||
<taxonomicName id="B885D278CFAC7D3A39C7D3AD51BFB829" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="ABE7DC0860ED60DAF8FB581DF384885B" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, but, at least in the specimen figured, more heavily sclerotised medially. Pronotum: hemicylindrical, highly arched transversely and weakly so longitudinally in mid-line. Sides of pronotum weakly curved, somewhat convergent anteriorly. Lateral margins in basal 1 / 4 not visible from above, concealed by lateral bulging of pronotal surface. Surface glossy, with no trace of reticulation but with sparse large punctures interspersed with much finer ones, these ca 1 / 5 of the diameter of the larger ones. In some specimens the punctation is a little stronger and closer. Anterior margin of pronotum not bordered, lateral margins bordered from just mediad of anterior angles to base, this border extending (as a very fine impressed line) over the lateral 1 / 3 of the hind margin. Median 1 / 3 of hind margin without any trace of an impressed line (Fig.
|
||||
<figureCitation id="29CB950E6AEA15E40E4BC447F9F45370" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 e</figureCitation>
|
||||
). Scutellum pentagonal, sides parallel in basal 1 / 2, the convergent to blunt apical point. Surface glossy black, impunctate. Elytra black, not quite parallel-sided, widest just behind middle then sides convergent to bluntly rounded apex. Interstices wide and almost flat, ca 5 × the width of the striae, silky black due to fine indistinct reticulation, and with sparse fine punctures, these separated by ca 8 × their diameters. Elytral sculpture: Fig.
|
||||
<figureCitation id="9A7434C92C4C7135A9F937AA71C4269F" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 e</figureCitation>
|
||||
. Striae narrow, vertical-sided, glossy, strial punctures encroaching on their margins. The striae stop short of the elytral apex on the inner 1 / 2 of the elytra, leaving a smooth glossy apical field. Metaventrite (Fig.
|
||||
<figureCitation id="A1888CE1AD78A51869E284146511169C" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 e, f</figureCitation>
|
||||
) fairly finely punctate, median furrow of central diamond-shaped plate flattish. No consistent sexual dimorphism. Legs dull mid- to dark brown. Long spur of mid tibiae clearly longer than the basal segment of the tarsi (Fig.
|
||||
<figureCitation id="77F6C7326A4D60730334740199D7DA61" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 d</figureCitation>
|
||||
). Hind tibiae with apical spines fairly long and unequal on dorsal / external edge, shorter and of even length along ventral / internal edge.
|
||||
</paragraph>
|
||||
<paragraph id="3BAACB1C47981A1C3ADF4C88E1BCBA96">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="F87F9BD579E482D3CA68260E3A0B5271" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 s – s</figureCitation>
|
||||
” ’, t, u) small, length ca
|
||||
<quantity id="DE178F5EE628918E1CC95F57E6FEA6E9" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
|
||||
. Apex of parameres membranous, obliquely truncate, often distorted in dried specimens, even when they are soaked again. Endophallus with scales, some of which are elongate, when not everted indistinguishable from that of
|
||||
<taxonomicName id="1523FA667B4D93DADE4F767F2E19562E" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="22BC229E8A443A7EA26C4DC8711522DE" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="184C82FD3593879842B6BA0E0AB3AE77" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 v, x</figureCitation>
|
||||
). However, the everted endophallus shows a clearly different arrangement of scale-patches (Fig.
|
||||
<figureCitation id="FB62777B230837EB3524E63EBE3858E1" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 q</figureCitation>
|
||||
’, s ” ’). The
|
||||
<specimenCount id="0DFD9531401B3172DFC60676592517D9" count="2" type="generic">two specimens</specimenCount>
|
||||
with the endophallus everted were the only ones where it had become extruded by partial decomposition of the specimen. These endophalli were softened in dilute potassium hydroxide, transferred to alcohol and then critical-point dried. Unfortunately the better of the two (Fig.
|
||||
<figureCitation id="3542201F22F417F30538259CDD778612" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 s – s</figureCitation>
|
||||
” ’) disintegrated when removal from the SEM stub was attempted. For this reason, the specimen with the less good endophallus is chosen as the
|
||||
<typeStatus id="D32FC31B82B88F9C6B9A9B4ED838536B" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EFOHK" type="etymology">
|
||||
<paragraph id="427197FAA1165670F63BE3A3BE40B2E3">
|
||||
<heading id="E7B897DC4587EC1528FBEEC115B781DC" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B50233CCE79728A12F2DC2F2CE1A2D76">
|
||||
The specific name
|
||||
<taxonomicName id="A4338F30605CDE1F3E2B21ABF2EE78DB" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chellala">
|
||||
<emphasis id="F3D2754ED47C819544BD6CB452786E49" italics="true">chellala</emphasis>
|
||||
</taxonomicName>
|
||||
is a feminine noun, the name of the
|
||||
<typeStatus id="DC72230937FDC34EFB713C4E4A73FD8D">type</typeStatus>
|
||||
locality.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVOHK" type="remarks">
|
||||
<paragraph id="288DC21AC72B929A3D38FC4026A3A538">
|
||||
<heading id="C57422AEE22EC6219E6FBEFD88FE7E2C" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="ED00A52D73597C128BC27690320A5870">
|
||||
So far known only from North Africa (Fig.
|
||||
<figureCitation id="087B6E633BF515E0C2DE8C9CA652E1AF" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b</figureCitation>
|
||||
). We have no information about the habitat.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
399
data/40/F7/E1/40F7E142199A525784B70C9E5518C20F.xml
Normal file
399
data/40/F7/E1/40F7E142199A525784B70C9E5518C20F.xml
Normal file
|
|
@ -0,0 +1,399 @@
|
|||
<document id="F5093028FE65AB651B6772100069155B" ID-CLB-Dataset="300069" ID-DOI="10.3897/mycokeys.107.124814" ID-GBIF-Dataset="6bd1ed8a-9466-43f9-9c30-03cef8f291b5" ID-publisher-id="124814" URI-arpha="A492D575-58D9-5C61-8CEC-72ADB271323B" article-type="research-article" checkinTime="1721676058192" checkinUser="pensoft" docAuthor="Aung, Sein Lai Lai, Liu, Feng-Yin, Gou, Ya-Nan, Nwe, Zin Mar, Yu, Zhi-He & Deng, Jian-Xin" docDate="2024" docId="40F7E142199A525784B70C9E5518C20F" docLanguage="en" docName="MycoKeys 107: 125-139" docOrigin="MycoKeys 107" docSource="https://mycokeys.pensoft.net/article/124814/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.mycokeys" docStyleName="PensoftTaxPub.0000.journal_article.mycokeys" docTitle="Alternaria momordicae S. L. L. Aung & J. X. Deng 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="139" masterDocId="A492D57558D95C618CEC72ADB271323B" masterDocTitle="Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China" masterLastPageNumber="139" masterPageNumber="125" pageNumber="125" updateTime="1721676827755" updateUser="ExternalLinkService">
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<mods:mods id="0E819EBC1D6731C1926E879D30CEDBBE" xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo id="7FAF2459B6DB1F0CB319719FE84A22EC">
|
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<mods:title id="03ACB36559B696782A7438ECE099769A">Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="ED50143DACAA4E54274C706DD45373FD" type="personal">
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||||
<mods:role id="70F6F250C0F47C847E353A0B230EC66A">
|
||||
<mods:roleTerm id="A6DB9A0D42DBDB7730B8151E75C5A06C">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="9E77E20448B5273A9B9F03BB08EE2927">Aung, Sein Lai Lai</mods:namePart>
|
||||
<mods:nameIdentifier id="CC67F1BEF4A70483559110B6A921CBFA" type="ORCID">https://orcid.org/0009-0006-2738-5598</mods:nameIdentifier>
|
||||
<mods:affiliation id="F5BE9934B88FDDE2F48504257E4C56B4">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="7CE5B3D7CEB6E56F85BC20AF78BE58C5" type="personal">
|
||||
<mods:role id="9D4DA29F653D139F813C1812929C4449">
|
||||
<mods:roleTerm id="B4AA40940D66AE1B6712018E1F8567DD">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="3171D5B63B349BBF01C601291369F680">Liu, Feng-Yin</mods:namePart>
|
||||
<mods:nameIdentifier id="1FCF744C04F48D297EBE7B0580EBF90D" type="ORCID">0000-0003-3114-603X</mods:nameIdentifier>
|
||||
<mods:affiliation id="9763DCFADDB0CB6A2A99222C8EC93306">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="7D5B7E17B8A750C7A4A5EE93CB1DBAB8" type="personal">
|
||||
<mods:role id="27C86331DF36A9B4C9F90A3B128042F3">
|
||||
<mods:roleTerm id="704EF1BCC5E7FE506DEC11A83858DAE4">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="AD7C204C58818F1F32DE981E3CD736C3">Gou, Ya-Nan</mods:namePart>
|
||||
<mods:nameIdentifier id="DB766DE2884BE1D97B8EBC5324816A67" type="ORCID">https://orcid.org/0009-0000-6376-8306</mods:nameIdentifier>
|
||||
<mods:affiliation id="F5B73F2062EED56E7A7BBE8F742D5CF5">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="251D6F829B99D6A2DA04F569EBEA2335" type="personal">
|
||||
<mods:role id="EE83D4D83B5778EC554A328B5A31DC4B">
|
||||
<mods:roleTerm id="FF8D2BA420BEC967DF01A53D484AA2D4">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="908763301261CC10A0BA8CA2CFDAA2B2">Nwe, Zin Mar</mods:namePart>
|
||||
<mods:nameIdentifier id="935E88586AB3B597324A97D220EF69F8" type="ORCID">https://orcid.org/0009-0000-6376-8306</mods:nameIdentifier>
|
||||
<mods:affiliation id="11259E78CF3E9E84E668A6531F1019CE">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="28A68DF6F85F9E71EE96B795031D3239" type="personal">
|
||||
<mods:role id="19EFF05049EA789765F03C661F065221">
|
||||
<mods:roleTerm id="ABEFE9A7E06645000789A99AB7A66AF8">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="026C7034C1CECC0772501F49684C95E6">Yu, Zhi-He</mods:namePart>
|
||||
<mods:nameIdentifier id="AC0E5A2813EE9C0161C43461AAB5EE60" type="ORCID">0000-0001-9477-4135</mods:nameIdentifier>
|
||||
<mods:affiliation id="49324AC595E63B3B2FDF7A78ED92947F">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="ADC41FDA1D5A3AA84A051A171ACA3F8F" type="personal">
|
||||
<mods:role id="C957D0D5835357DEFFDCFBE18E219796">
|
||||
<mods:roleTerm id="F238C9143FF34F4E385D4F6143EEE2E4">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="6933A73DEF5F79D32688E452770772D6">Deng, Jian-Xin</mods:namePart>
|
||||
<mods:nameIdentifier id="0D44F3B21D91D2FA0C7AB7ADFC18A16C" type="ORCID">0000-0001-7304-5603</mods:nameIdentifier>
|
||||
<mods:affiliation id="3F76D07101FC8FA278E249E567FA1F50">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="E896DCF34728FEDD3006CFFE75CBC877">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="6B9488DD95B209331EE6157930E6337E" type="host">
|
||||
<mods:titleInfo id="10A4D8935D5744A04C17F1451DDFB653">
|
||||
<mods:title id="1E822F0142013DFCB7718425FD3CA8F1">MycoKeys</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="6873082CA5E44AA57EB4133BC9874522">
|
||||
<mods:date id="90CDB057053BEF8F6AA935B541EBA09C">2024</mods:date>
|
||||
<mods:detail id="A139EBDB3B2BAD243D7238BBA985E4E1" type="pubDate">
|
||||
<mods:number id="E2FA33F552CDC988B8081129631016C8">2024-07-22</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="12CBA5273A85852F5221EEA9B7987587" type="volume">
|
||||
<mods:number id="2547FBFB8B01FFE20CF0883C665CB7A8">107</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="D1DE053196A3F6969E8986519314324D" unit="page">
|
||||
<mods:start id="03B8B6C0212D6CB470D402E6472B4596">125</mods:start>
|
||||
<mods:end id="2127BF2172D945B89449F45801BC3F20">139</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="D377B8C7F55DCE49D4007BEBC15241EE">journal article</mods:classification>
|
||||
<mods:identifier id="9A4257DA2C60362A8DB87D2289C2A92A" type="CLB-Dataset">300069</mods:identifier>
|
||||
<mods:identifier id="FE91D872A67106A5CDE4976BA79CC992" type="DOI">10.3897/mycokeys.107.124814</mods:identifier>
|
||||
<mods:identifier id="AB77ABF8768BB35474E797CADA05EB88" type="GBIF-Dataset">6bd1ed8a-9466-43f9-9c30-03cef8f291b5</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="40F7E142199A525784B70C9E5518C20F" ID-DOI="http://doi.org/10.5281/zenodo.12796842" ID-Zenodo-Dep="12796842" ID-arpha="40F7E142-199A-5257-84B7-0C9E5518C20F" ID-mycobank="MycoBank No: 851270" LSID="urn:lsid:plazi:treatment:40F7E142199A525784B70C9E5518C20F" httpUri="http://treatment.plazi.org/id/40F7E142199A525784B70C9E5518C20F">
|
||||
<subSubSection id="A5DD17E4B37E9ECA58D9BFFEC2F8EE3B" type="nomenclature">
|
||||
<paragraph id="6D40111F2AD5EC9597639A08E6827C99">
|
||||
<taxonomicName id="7ED87934BBE3D58DBC7CF7759BEA5295" authority="S. L. L. Aung & J. X. Deng" authorityName="S. L. L. Aung & J. X. Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="6CC777C502B3168CDA6152C03885D100" italics="true">Alternaria momordicae</emphasis>
|
||||
S. L. L. Aung & J. X. Deng
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="031A9ED62F9727778447BD390B6C264C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C6328F135D3A3976B0DF046AC40D6DA7" type="description">
|
||||
<paragraph id="A2A8B5AC22EA4B24E03FC9840C2CB94F">
|
||||
<figureCitation id="F7CF773E091C62E39ACC2B90E3DA8CE3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alternaria momordicae sp. nov. (ex-type YZU 161378) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure3" httpUri="https://binary.pensoft.net/fig/1097808">Fig. 3</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELGAK" type="material">
|
||||
<paragraph id="38F9666809ADFF751F0331E3DBC11D89">
|
||||
<heading id="D88ADD0DF36803EBB7816C6C24C825D2" reason="title">
|
||||
<typeStatus id="43E294F7591089B71FA0BCF5AADC7096">Type</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B1F3DE7FDD5BA46811D8276FA823377B">
|
||||
<collectingCountry id="B3AD9BAB0CE3787814D3F8AB46A44EFC" name="China">China</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="1B9FBDAE270729B7307CBC8BDAE6930C" country="China" name="Sichuan">Sichuan Province</collectingRegion>
|
||||
, Deyang city infected leaves of
|
||||
<taxonomicName id="9A45B626BC5069151AA455435E5E4A0F" class="Magnoliopsida" family="Cucurbitaceae" genus="Momordica" kingdom="Plantae" order="Cucurbitales" phylum="Tracheophyta" rank="species" species="charantia">
|
||||
<emphasis id="9FD81D6A34216C8169A3CC4D46DDAED7" italics="true">Momordica charantia</emphasis>
|
||||
</taxonomicName>
|
||||
. 2016,
|
||||
<collectionCode id="8AD48AEF48A0BF35373738819308402D" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
|
||||
. X Deng, (
|
||||
<abbrev id="ABBRID0E3GAK" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="21803DD7924BD8EBD749BF5346BA8AE6" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
- H- 2016001,
|
||||
<typeStatus id="BBEF90CA9D6BB009809BAC6DA99E5659" type="holotype">holotype</typeStatus>
|
||||
), ex-type culture
|
||||
<abbrev id="ABBRID0EAHAK" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="69FF7C2D74C3DFA970C2D05091FEF64D" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
161378.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEHAK" type="etymology">
|
||||
<paragraph id="58BBABB36CE0333349E7085F1516C275">
|
||||
<heading id="7D192B5EB7872B6249AC7D7BEA1C453C" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="EC385281F96DD9AFF4BE4C5A4B78F5D0">
|
||||
Refers to the host genus,
|
||||
<taxonomicName id="2256480012C3B008F020A96B6A9B133F" class="Magnoliopsida" family="Cucurbitaceae" genus="Momordica" kingdom="Plantae" order="Cucurbitales" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="C0AC7DCD18C16265B81FF20E1F354104" italics="true">Momordica</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERHAK" type="description">
|
||||
<paragraph id="6F93DF0625CA0EADC46E51EF9A6418BB">
|
||||
<heading id="5F7888ADAC224A5380654B58FCF6B28C" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E1FE4080F25B1271711E53DB746FCF31">
|
||||
<emphasis id="7FF1148421ABCA083144670BE63A7AFE" bold="true" italics="true">Colonies</emphasis>
|
||||
on
|
||||
<abbrev id="ABBRID0E3HAK" xlink_title="potato dextrose agar">
|
||||
<collectionCode id="71AF866422B4401293D92C805A63B73C" country="Sri Lanka" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13944" lsid="urn:lsid:biocol.org:col:13944" name="Royal Botanic Gardens" type="Herbarium">PDA</collectionCode>
|
||||
</abbrev>
|
||||
(7 d at 25 °
|
||||
<collectionCode id="18A8D9D240177EAC5DA8A81C9F5A032F" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
) greyish yellow-green, light white at the edge, buff to salmon in reverse, surface compact,
|
||||
<quantity id="E7AC9287566BBDE3C6C40A003985D8E6" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">50–55 mm</quantity>
|
||||
in diam. (Fig.
|
||||
<figureCitation id="B51592A7160513428816761ED5567483" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alternaria momordicae sp. nov. (ex-type YZU 161378) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure3" httpUri="https://binary.pensoft.net/fig/1097808">3 A, B</figureCitation>
|
||||
). On
|
||||
<abbrev id="ABBRID0EEIAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
(7 d at 22 °
|
||||
<collectionCode id="AF432F7AB5DCA91DB7CD21FA63761168" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
),
|
||||
<emphasis id="D168572A1BFB5E1A0AA60BF0E38ABF8D" bold="true" italics="true">conidiophores</emphasis>
|
||||
arising from substrate, simple, straight or flexuous, septate, olivaceous buff to olivaceous, 26.5–93 × 3–4 μm (x ̄ = 59.5 × 3.8 μm, n = 20);
|
||||
<emphasis id="14EF258DB3F766C34EC9CE6A00296CA3" bold="true" italics="true">conidiogenous cells</emphasis>
|
||||
5–10 × 3–5 µm (x ̄ = 7 × 4 µm, n = 20), mono- to polytretic, terminal, determinate, cylindrical, olivaceous buff to olivaceous, smooth, thin-walled, apically doliiform, with 1 conidiogenous locus cicatrized on conidial secession, sometimes swollen near conidiogenous loci;
|
||||
<emphasis id="188CD18755E5269FDEE727DCECDA3EFE" bold="true" italics="true">conidia</emphasis>
|
||||
3–4 units per chain, arising from the apex or near the apex of the conidiophores or terminal hyphae, muriform, long ellipsoid or ovoid to obclavate, sometime inverted club-shaped, 6–42 × 4–34 μm (x ̄ = 32.8 × 13.5 μm, n = 50), 1–5 transverse septa, apical beak 2–19.5 μm long and 1–2 septa (Fig.
|
||||
<figureCitation id="8AE01E9132ED251B1A3F003FF2FA4F00" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alternaria momordicae sp. nov. (ex-type YZU 161378) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure3" httpUri="https://binary.pensoft.net/fig/1097808">3 C, E</figureCitation>
|
||||
); On
|
||||
<abbrev id="ABBRID0EVIAK" xlink_title="V8 juice agar">
|
||||
<collectionCode id="2FB099530D337E901F460978D5A4F67B" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
8
|
||||
<collectionCode id="C51BD945A1EB4B1493031EB3BA52BDC5" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
|
||||
</abbrev>
|
||||
(7 d at 22 °
|
||||
<collectionCode id="BDA7C6A7ECC9AF870D37BDF5C10E91E8" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
),
|
||||
<emphasis id="ACDB6A34DFB1F7AD619CDA2A40BF5056" bold="true" italics="true">conidiophores</emphasis>
|
||||
straight or curved, smooth-walled, olivaceous buff 23–63 (– 208) × 3–5 μm (x ̄ = 64.9 × 4.2 μm, n = 20);
|
||||
<emphasis id="8F8CF92F06695B23CFD71EE212CEE2D2" bold="true" italics="true">conidiogenous cells</emphasis>
|
||||
5–13 × 3–4 µm (x ̄ = 7 × 4 µm, n = 20), mono- to polytretic, terminal, determinate, cylindrical, olivaceous buff, smooth, thin-walled, apically doliiform, with 1 conidiogenous locus cicatrized on conidial secession, sometimes swollen near conidiogenous loci;
|
||||
<emphasis id="7B060FA75E96C6D130F079906F7398CC" bold="true" italics="true">conidia</emphasis>
|
||||
3–4 units per chain, muriform, long ellipsoid or ovoid to obclavate, inverted club-shaped, 24–61 × 10 – 17 μm (x ̄ = 39 × 14.3 μm, n = 50), 1–5 transverse septa with apical beak 3–25.5 μm long and 1–2 septa (Fig.
|
||||
<figureCitation id="18A141C1C4F1E3AF2A4562FFDD35BA65" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alternaria momordicae sp. nov. (ex-type YZU 161378) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure3" httpUri="https://binary.pensoft.net/fig/1097808">3 D, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="F9F3B712F77BB22BAC2078AB4113FEDA" ID-DOI="10.3897/mycokeys.107.124814.figure3" ID-arpha="7C3076CB-3903-5462-9AC4-E3364ADAD3AA" httpUri="https://binary.pensoft.net/fig/1097808" startId="F3">
|
||||
<paragraph id="E669CF2CB1909865B6D1AD6096462245">
|
||||
<label id="3F86C39B08D64ED3C94789CDADCB93B9">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="AC8B0B21563B66D915272E62490441CD">
|
||||
<taxonomicName id="4B64460D6950D23A4FBFC37173DF51FA" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="2E055F49B742F2A302FCD835A5AA49EF" italics="true">Alternaria momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="FA9000E35518B8D66ECD41512E881B5B" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(ex-type
|
||||
<abbrev id="ABBRID0EZJAK" xlink_title="Fungi Herbarium of Yangtze University">YZU</abbrev>
|
||||
161378)
|
||||
<emphasis id="AA93D08CDF87C90C1C145E3941FB609A" bold="true">A, B</emphasis>
|
||||
seven-day-old culture on
|
||||
<abbrev id="ABBRID0E6JAK" xlink_title="potato dextrose agar">PDA</abbrev>
|
||||
<emphasis id="EE70D984E62FB68E6537C56881946B45" bold="true">C, D</emphasis>
|
||||
conidiophores and conidia on
|
||||
<abbrev id="ABBRID0EFKAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
and
|
||||
<abbrev id="ABBRID0EJKAK" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
, respectively
|
||||
<emphasis id="47E64A49341ADFA07616458FA40E212B" bold="true">E, F</emphasis>
|
||||
conidia on
|
||||
<abbrev id="ABBRID0EPKAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
and
|
||||
<abbrev id="ABBRID0ETKAK" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
, respectively. Scale bars: 25 μm (
|
||||
<emphasis id="0621CD7EB29F962510246083F82BB1D4" bold="true">E, F</emphasis>
|
||||
); 50 μm (
|
||||
<emphasis id="2AB92BEFC5838F69512BECF357DDD0DA" bold="true">C, D</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EALAK" type="Additional isolate examined">
|
||||
<paragraph id="B0B6EA870AC060F60ADD8BB0D2FF3B78">
|
||||
<heading id="1538412978B49E6858282B2693F4C522" reason="title">Additional isolate examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5D6CFDACD084B709F0387C34AC8EFCAA">
|
||||
<collectingCountry id="F74C3D07B65B47CBF2134A6FF9DB4E2B" name="China">China</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="5DE7B0D3CDC83C9DF8F947A1B377FF8C" country="China" name="Sichuan">Sichuan Province</collectingRegion>
|
||||
, Deyang city infected leaves of
|
||||
<taxonomicName id="8E47974A54A94B19714A473135B3BAC0" class="Magnoliopsida" family="Cucurbitaceae" genus="Momordica" kingdom="Plantae" order="Cucurbitales" phylum="Tracheophyta" rank="species" species="charantia">
|
||||
<emphasis id="E9AAEF0768C1D996AD692B486CD4A627" italics="true">Momordica charantia</emphasis>
|
||||
</taxonomicName>
|
||||
. 2016,
|
||||
<collectionCode id="16C4980E29866DB7C608C169A0FC984F" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
|
||||
. X Deng, living culture
|
||||
<abbrev id="ABBRID0ERLAK" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="F028FA879068C000151EE8A1A0B46592" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
161379.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVLAK" type="notes">
|
||||
<paragraph id="8990270D8159587C3BEC33A3DF026AEF">
|
||||
<heading id="6CCE2F5B1C307359EAF82BBD8735FFFD" reason="title">Notes.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BE7B0090D4B3990D3BB18C5E84D11B43">
|
||||
After the combined dataset of
|
||||
<abbrev id="ABBRID0E2LAK" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="C5189D2079416F0EAE37A425EAA5A937">ITS</collectionCode>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="18A5ED1D28C010B428B0D32ACC2B33E8" italics="true">
|
||||
<collectionCode id="C7058BA1B955B7D0AD7204B8717F4ECE">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0ECMAK" xlink_title="translation elongation factor 1 alpha">
|
||||
<emphasis id="31FE9A361E6DE2F87DD20A90F9C98180" italics="true">
|
||||
<collectionCode id="590EDF69BD05B3BACD066997F7AB148B" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" type="Herbarium">TEF</collectionCode>
|
||||
1
|
||||
</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<abbrev id="ABBRID0EHMAK" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="E208F3FB9210C77B0F555A328401F7A1" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="AA73217CDB1CA21C076EF958247B4785" italics="true">Alt a 1</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0EOMAK" xlink_title="endopolygalacturonase gene">
|
||||
<emphasis id="9E09FC1C7733A6A38F73FC2EE3E85633" italics="true">EndoPG</emphasis>
|
||||
</abbrev>
|
||||
and OPA 10-2 gene fragments,
|
||||
<taxonomicName id="68D6D7B14A6E80E60930688C917AD545" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="3BCA6C8A3B9281883EC322356BB6142A" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="5F48EFC884FE8E40D0D35DB51AC9E85C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is readily distinguished from its sister species
|
||||
<taxonomicName id="73B249717FDFFD0F01495355EC2734B2" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="baoshanensis">
|
||||
<emphasis id="658BCC6EF83C450E795FDADB2B440E35" italics="true">A. baoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="EA7006C024C25BA81C3C30797B38E2A2" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="6579600B10A61EB3ADD25CB3BCAB4B9F" italics="true">A. jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="FFE99D97647DDE283997B48D277A7344" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="0E0BA4F50F9078025AC6F70CB45D6892" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="1CE91C2E696E521D1719C29A76C1B4A4" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="078F3A0746889F05AB9B4D66F757691A" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="ovoidea">
|
||||
<emphasis id="B8CA40A37112E29D0893E46F575DECFA" italics="true">A. ovoidea</emphasis>
|
||||
</taxonomicName>
|
||||
, (Fig.
|
||||
<figureCitation id="0828EAF785DD7915016DB19E6EB28275" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree of the Alternaria species most related to the new taxa based on maximum likelihood analysis using the combined gene sequences of ITS, GAPDH, TEF 1, RPB 2, Alt a 1, EndoPG and OPA 10-2 which rooted with Alternaria alternantherae (CBS 124392) from sect. Alternantherae. The Bayesian posterior probabilities > 0.60 (PP) and bootstrap support values > 60 (BS) are given at the nodes (PP / BS). The novel species are highlighted in bold. Ex-type isolates are marked with a superscript T and Representative isolates are marked with a superscript R." figureDoi="10.3897/mycokeys.107.124814.figure1" httpUri="https://binary.pensoft.net/fig/1097806">1</figureCitation>
|
||||
). After a nucleotide pairwise comparison as suggested by
|
||||
<bibRefCitation id="7698433835C6205735B799D66BC9DF90" DOI="10.5943/mycosphere/7/11/4" author="Jeewon R & Hyde KD" firstAuthor="Jeewon" issue="11" journalOrPublisher="Mycosphere" pagination="1669-1677" refId="B10" refString="Jeewon R, Hyde KD (2016) Establishing species boundaries and new taxa among fungi: Recommendations to resolve taxonomic ambiguities. Mycosphere 7 (11): 1669–1677. https://doi.org/10.5943/mycosphere/7/11/4" title="Establishing species boundaries and new taxa among fungi: Recommendations to resolve taxonomic ambiguities." volume="7" year="2016">Jeewon and Hyde (2016)</bibRefCitation>
|
||||
, the present species can be readily distinguished from the closet species
|
||||
<taxonomicName id="2DB360DFF7EE828958285CB40F1332A8" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="27326806EC48CF7E07EB494011D337E6" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
and others related a novel species based on any of the
|
||||
<abbrev id="ABBRID0E3OAK" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="AF8CA87D0CB173DD974F0028A2905FC6">ITS</collectionCode>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="FD7A14B5F1763E432CA5D5FF5B22D6F7" italics="true">
|
||||
<collectionCode id="7467E11CE058CB56F77081F4476720C9">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0EDPAK" xlink_title="translation elongation factor 1 alpha">
|
||||
<emphasis id="7DAE88736AE8B33671E325B4CA6CEC6D" italics="true">
|
||||
<collectionCode id="477D5FB35B4EF1393CFBAF91B815C46C" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" type="Herbarium">TEF</collectionCode>
|
||||
1
|
||||
</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<abbrev id="ABBRID0EIPAK" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="46EBE10D863138320EE892B050E3FF05" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="4ABBD97C015916BF64AA425ADE0E2CBD" italics="true">Alt a 1</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0EPPAK" xlink_title="endopolygalacturonase gene">
|
||||
<emphasis id="CA4BD21F932303C7FF3DB5A3EEBC78D3" italics="true">EndoPG</emphasis>
|
||||
</abbrev>
|
||||
, and OPA 10-2 genes, which has 1 bp difference in the
|
||||
<abbrev id="ABBRID0ETPAK" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="2E909702783B1CE037007CCF0EF3552C">ITS</collectionCode>
|
||||
</abbrev>
|
||||
region, 1 bp in
|
||||
<emphasis id="F10C74594F9E563F4AFFCFD56C827ABA" italics="true">
|
||||
<collectionCode id="4AF384174D3BFAF21E2546F06FDC9712">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
, 1 bp in
|
||||
<abbrev id="ABBRID0E1PAK" xlink_title="translation elongation factor 1 alpha">
|
||||
<emphasis id="E2CA215E96207490949F82CE4A9EDEA8" italics="true">
|
||||
<collectionCode id="FB29AE7DEB0512C12DB596AF1E1CFD1F" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" type="Herbarium">TEF</collectionCode>
|
||||
1
|
||||
</emphasis>
|
||||
</abbrev>
|
||||
, 4 bp in
|
||||
<abbrev id="ABBRID0E6PAK" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="5B9E5886D758C957B7BFD63CBFFD094F" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
, 8 bp in
|
||||
<emphasis id="D9E636ECDCC16DD21CAACB3EA01C767A" italics="true">Alt a 1</emphasis>
|
||||
and 10 bp in
|
||||
<abbrev id="ABBRID0EGQAK" xlink_title="endopolygalacturonase gene">
|
||||
<emphasis id="14A7C2AD3E1292E18EA2FAA19685155F" italics="true">EndoPG</emphasis>
|
||||
</abbrev>
|
||||
when compared with
|
||||
<taxonomicName id="66ADB18E9B8D688270304C2AA7136083" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="2E9BB94FCCB1612E32F71E2282EF2AAB" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
and 1 bp in
|
||||
<emphasis id="89D6EBF1D75130C1129F66DECD644F86" italics="true">
|
||||
<collectionCode id="38AD849AC8998F7ADAE724100579B180">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
, 4 bp in
|
||||
<abbrev id="ABBRID0EYQAK" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="75515A856FB782464808487A3D2D6DE9" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
, and 11 bp in OPA 10-2 when compared with
|
||||
<taxonomicName id="B314DE97F5A56FFE13E28DBE05B068DF" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov">
|
||||
<emphasis id="DC7CD07BF7A264B9B8D03DD64A14F05D" italics="true">A. jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="44E436494EBCDE95FAC6A701F26FED59" rank="species">sp. nov</taxonomicNameLabel>
|
||||
.. Morphologically,
|
||||
<taxonomicName id="229183CE0CA315F9DB435A4DBA749C16" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="5A30F723BE7413CC430A0948A3167557" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="3592B80496A40ABF4C690ED082FF4FD3" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
produces conidia on
|
||||
<abbrev id="ABBRID0ESRAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
that are significantly shorter than those on
|
||||
<abbrev id="ABBRID0EWRAK" xlink_title="V8 juice agar">
|
||||
<collectionCode id="C4A9F4324567D8D7E7CC2C41E14DE759" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
8
|
||||
<collectionCode id="EF8B0703811BA1D7AAEEA06039EB1909" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
|
||||
</abbrev>
|
||||
. It can be distinguished from
|
||||
<taxonomicName id="0A1604D11A5209EE8EAE1EB8C8E560FF" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="baoshanensis">
|
||||
<emphasis id="E0A6CD484F839157DF6343A28316CB3C" italics="true">A. baoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="23B7FB3C7BD90EA76FAA3428DCF164AB" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="C37ACBA7BB4C13767663FBF77FAC0027" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="BB11A9A151A464BF2BC6BFE02AA77AD4" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="ovoidea">
|
||||
<emphasis id="E55ECE4467781C502AC5AB1C6BAA4526" italics="true">A. ovoidea</emphasis>
|
||||
</taxonomicName>
|
||||
by producing conidia on a single conidiogenous locus with a wider body and longer beak in a chain of 3–4 units. Additionally, it differs from
|
||||
<taxonomicName id="9CE21858F269FAC87F0779F8584CFFE0" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="FFDC72486E2B67C723ED2C7D37DBB8FA" italics="true">A. jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="B17F1772CFCD91DA7243F211D7CEAA10" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by having a longer beak (Table
|
||||
<tableCitation id="23568F9DFC7AEE13C376C19EDEDCAE27" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Conidial features of the novel Alternaria species proposed here and their closest relatives in section Alternaria." httpUri="http://table.plazi.org/id/0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7" tableUuid="0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7">2</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
389
data/80/4F/B7/804FB70A33F65170BB500EA7C567EA54.xml
Normal file
389
data/80/4F/B7/804FB70A33F65170BB500EA7C567EA54.xml
Normal file
|
|
@ -0,0 +1,389 @@
|
|||
<document id="3375A7FA00049EE74A0651FC1DE3B017" ID-DOI="10.3897/zookeys.1207.117225" ID-ZooBank="94F18819-5AF5-4100-AB35-AA3C3976EE80" ID-publisher-id="117225" URI-arpha="255F6420-5705-5F50-9167-509E6C76FA1C" URI-zoobank="http://zoobank.org/94F18819-5AF5-4100-AB35-AA3C3976EE80" article-type="research-article" checkinTime="1721676390257" checkinUser="pensoft" docAuthor="Angus, Robert B., Maté, Jason F., Angus, Elizabeth M. & Král, David" docDate="2024" docId="804FB70A33F65170BB500EA7C567EA54" docLanguage="en" docName="ZooKeys 1207: 205-299" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/117225/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Aphodius (Liothorax) wilsonae Mate" docType="treatment" docVersion="1" lastPageNumber="299" masterDocId="255F642057055F509167509E6C76FA1C" masterDocTitle="Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" masterLastPageNumber="299" masterPageNumber="205" pageNumber="205" updateTime="1721676524171" updateUser="pensoft">
|
||||
<mods:mods id="297D627E6EDD477201206938CC98001A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="52D49DAD11E46872724442B90B8686E0">
|
||||
<mods:title id="82AE8D8CB641FBF9FCEF8ACBBA6DFF66">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="4265FB6C9548E19162F1A25D9551ED11" type="personal">
|
||||
<mods:role id="1E833BB193FB518EB9A55AB37BD96D52">
|
||||
<mods:roleTerm id="7A6ABDE03178348364C7928556A863F6">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="FBFE7648C6388B7BA7F956EC313616DC">Angus, Robert B.</mods:namePart>
|
||||
<mods:nameIdentifier id="B1E938C7495D47D5411B46E0A5FD6423" type="ORCID">0000-0002-3860-5617</mods:nameIdentifier>
|
||||
<mods:affiliation id="1053BC908C5F2BEACB075C28E9FF5C00">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
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<mods:namePart id="CC61557DFAB743A0E9A7344B158796A1">Maté, Jason F.</mods:namePart>
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<mods:affiliation id="C8ABAF1395AE4968314E5D19C17E8031">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="B5BB1BF91B0BB95B3647C152169F5BBE">Angus, Elizabeth M.</mods:namePart>
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<mods:affiliation id="887D0B34E3E21621E62057266209E045">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="D0DE697DAF0F987B3CDBAEDE95E41A99">Král, David</mods:namePart>
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<mods:affiliation id="0CDBD0598A0C7D4F8938E8F9F5DC74BD">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:typeOfResource id="FE38ABB1210C123F6825133693DD96A0">text</mods:typeOfResource>
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<mods:title id="B631E28CB38E90A6FA3D30D289657D4E">ZooKeys</mods:title>
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<mods:part id="2864889E918870FEC3F1608C9B11627F">
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<mods:date id="8020DF77E1785D46E171F3AD3163DB47">2024</mods:date>
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<mods:number id="407C9096C11A83306C56342DC7D6453A">2024-07-22</mods:number>
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<mods:number id="83D03CAB1B3D177C042E79B41980BDE2">1207</mods:number>
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<mods:identifier id="CBA92DAA2555D32CFEB752CDDB54DCC4" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<mods:identifier id="329E4F63B1670380C625CED508984778" type="ZooBank">94F18819-5AF5-4100-AB35-AA3C3976EE80</mods:identifier>
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<subSubSection id="D505DDAC7A96990349DE2FB24641B6B5" type="nomenclature">
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||||
<taxonomicName id="D633626269A2640D0C1860DF7D474ABA" ID-CoL="FGDW" authority="Mate" authorityName="Mate" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae" subGenus="Liothorax">
|
||||
<emphasis id="B1A27203D92A46725344DA0ED8F677A9" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="E8495800A9FE14FF46FE9E73282C9418" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="C59BF41799D0AE3CD9DC92F96585EF0D" italics="true">wilsonae</emphasis>
|
||||
<bibRefCitation id="FDEA75CF2EC52C8CE0DB238AC5361D52" author="Maté JF & Angus RB" journalOrPublisher="Koleopterologische Rundschau" pagination="325-336" refId="B33" refString="Mate JF, Angus RB (2005) Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky. Koleopterologische Rundschau 75: 325 - 336." title="Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky." volume="75" year="2005">Maté & Angus, 2005</bibRefCitation>
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||||
</taxonomicName>
|
||||
, stat. rest.
|
||||
</paragraph>
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||||
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<subSubSection id="62DB48AA8A4AD6540040370BF457B3FE" type="description">
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<paragraph id="9835F330884C7DD9E2D6407AEDAE3B4B">
|
||||
<figureCitation id="1CF461C60B9B510C7B46ACEE31171650" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">
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Figs
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<quantity id="B103C257A04A4899FE2FF430D2B06414" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" unit="g" value="4.0">4 g</quantity>
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</figureCitation>
|
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,
|
||||
<figureCitation id="F70CF50DB6F6C80ADBFB789664E69FFF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 k</figureCitation>
|
||||
,
|
||||
<figureCitation id="954D19DCDF8714C095B725B050DECD9B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 k, l</figureCitation>
|
||||
,
|
||||
<figureCitation id="0C7A936E694007F0366B7C4398459543" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 h, i</figureCitation>
|
||||
,
|
||||
<figureCitation id="62778F18D061EEF6A541CDFC88F78F12" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="6FC699AF5CB7917202A476F58E499552" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 w</figureCitation>
|
||||
,
|
||||
<figureCitation id="4E3231C349E387DA3A352095D875528B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 o</figureCitation>
|
||||
,
|
||||
<figureCitation id="9073C436F32F98000052DFFB8BC6A112" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 u – w</figureCitation>
|
||||
,
|
||||
<figureCitation id="0F088D65EEB32ED9A742BF82A681AC7E" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">
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<quantity id="615532AF836D2C9F1C9B888A74A8E23B" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" unit="g" value="23.0">23 g</quantity>
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– i
|
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</figureCitation>
|
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,
|
||||
<figureCitation id="2F760D38A13397C1B220565E491D50AC" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 o</figureCitation>
|
||||
,
|
||||
<figureCitation id="3863AB3D10C066E1EA2122986CF0F338" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 d ’, e’, 33 m-p, 34 k, l</figureCitation>
|
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</paragraph>
|
||||
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|
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<treatmentCitation id="A0B243698B8FE69D64C630C8A97B2401" author="Maté JF & Angus RB" page="329" year="2005">
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<taxonomicName id="A506E3232C5F2E1493F52F83B56714B4" ID-CoL="FGDW" authority="Mate" authorityName="Mate" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae" subGenus="Liothorax">
|
||||
<emphasis id="FF1B1D2153386BCB406119D2D362F4E7" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="B9FA71BF3133575AFBDACD395F626BFC" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="4C702AABFE8D8D052B4EAE0DFB535FE7" italics="true">wilsonae</emphasis>
|
||||
<comment id="2A16BF64358825E46655AD93D78F3BE0">
|
||||
<bibRefCitation id="53719F0A9D3B37D14B76E8C6434D2568" author="Maté JF & Angus RB" journalOrPublisher="Koleopterologische Rundschau" pagination="325-336" refId="B33" refString="Mate JF, Angus RB (2005) Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky. Koleopterologische Rundschau 75: 325 - 336." title="Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky." volume="75" year="2005">Maté & Angus, 2005: 329</bibRefCitation>
|
||||
.
|
||||
</comment>
|
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</taxonomicName>
|
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|
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<paragraph id="71168FFAA4CE9A4CC0711129D8C823DE">
|
||||
<heading id="2B7E7C297EE94454610CB32CEF21EB07" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7AA1D0C92B05656E24163393FF8CF264">
|
||||
<materialsCitation id="FF70A635EE65CC27D5C8A945D44D151D" collectingDate="2001-04-26" collectionCode="NHMUK" collectorName="SP Provincia de Burgos & de Corconte" latitude="40.031" location="Balneario de Corconte" longLatPrecision="69" longitude="-3.884" municipality="Provincia de Burgos" specimenCount="1" specimenCount-male="1" stateProvince="Angus" typeStatus="holotype">
|
||||
<emphasis id="F7E9EFFF870B8ACEC5909D4C4E4D8136" bold="true" italics="true">
|
||||
<typeStatus id="2E15133B1666831E07B12B9DE603D04E" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="596F51DBB6FA26404F0D8BADC281834F" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<collectorName id="C427119C4FBE31F3FF2B431FF6745C8E">
|
||||
SP
|
||||
<collectingMunicipality id="51A8C25D0E07F333D13093DC7B0257E2">Provincia de Burgos</collectingMunicipality>
|
||||
</collectorName>
|
||||
,
|
||||
<location id="EA516796F9C68F643F3422FC9E73A395" LSID="urn:lsid:plazi:treatment:804FB70A33F65170BB500EA7C567EA54:EA516796F9C68F643F3422FC9E73A395" latitude="40.031" longLatPrecision="69" longitude="-3.884" municipality="Provincia de Burgos" name="Balneario de Corconte" stateProvince="Angus">
|
||||
Balneario
|
||||
<collectorName id="15D14B2E5ACC86673B7AA7CF4C575311">de Corconte</collectorName>
|
||||
</location>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEODM" specific-use="{"type":"Point","coordinates":[-3.884000,40.031000]}">
|
||||
<geoCoordinate id="839723CA6688673D0276107AF75D6B35" degrees="40.031" direction="north" orientation="latitude" precision="55" value="40.031">40.031 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D47C6219F715694E4114CA6AAFEF6E5E" degrees="3.884" direction="west" orientation="longitude" precision="55" value="-3.884">3.884 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<date id="ADF34598FBB5A9419C95E727EB212312" value="2001-04-26">
|
||||
<collectingDate id="2D7D063EAD28D58D289DE227B73B5485" value="2001-04-26">26. iv. 2001</collectingDate>
|
||||
</date>
|
||||
. leg. R. B.
|
||||
<collectingRegion id="5E43044454EDBE0B9E1E6106FA23763A" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
(
|
||||
<named-content id="0E255E7BAB1C28C4786A5BA63EFE4460" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="4498DFEBA7B64EEDB773F18CFED27463" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="09D9F013E65F877C8AB01C3FBEB6D199" bold="true" italics="true">
|
||||
<typeStatus id="1E519C7A4E623FCA50885052C8A885F6" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: SP: Provincia de
|
||||
<collectingRegion id="CE727BCDE658D8066F91D9A378F97DCD" country="Spain" name="Madrid">Madrid</collectingRegion>
|
||||
, Manzanares el Real, Embalse de Santillana,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETODM" specific-use="{"type":"Point","coordinates":[-3.857000,40.720000]}">
|
||||
<geoCoordinate id="FC9FC00394788CABEA2017E93D7CD6BE" degrees="40.720" direction="north" orientation="latitude" precision="55" value="40.72">40.720 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="16FF4D1AA0F139837DDCE3F9D77102E5" degrees="3.857" direction="west" orientation="longitude" precision="55" value="-3.857">3.857 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<date id="E344390E9F4A823A7C0F2E564256225F" value="2003-04-01">1. iv. 2003</date>
|
||||
, sieving detritus from edge of water, leg. R. B.
|
||||
<collectingRegion id="5B42111867D9F85BD23381651D12718D" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
& G. I. Aradottir,
|
||||
<specimenCount id="443EC6BFD893C6B5A100B512D589EDEF" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="E4E84C2B4A4F1D8103F06FAD6114EB49" count="1" type="female">1 ♀</specimenCount>
|
||||
, 1 unsexed (
|
||||
<named-content id="E0B9C6C2AD3B7A602387DD4282CB0E64" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="D1C5C279F44A3D3C687D2C8A53F329C7" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3ODM" type="material">
|
||||
<paragraph id="F6866CF710E993C8625544EED8C8C3A4">
|
||||
<heading id="DD9CA96319CA759B6129790D18207BA8" reason="title">Other material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="337664288BA408E8F2C32BDC5EC31319">
|
||||
SP Provincia de
|
||||
<collectingRegion id="7AD9DA1866EA050A525B9A7D97A022C5" country="Spain" name="Cantabria">Cantabria</collectingRegion>
|
||||
, Areños,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFPDM" specific-use="{"type":"Point","coordinates":[-4.724000,43.112000]}">
|
||||
<geoCoordinate id="614B244830B7AEF3D4464D2748EB2443" degrees="43.112" direction="north" orientation="latitude" precision="55" value="43.112">43.112 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="DF347A3E6267A63A2E16EBE17C8E34E8" degrees="4.724" direction="west" orientation="longitude" precision="55" value="-4.724">4.724 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<date id="68899A1929D022766898BEAC648870AA" value="2012-06-05">5. vi. 2012</date>
|
||||
. By digging mud under dried-out pool.
|
||||
<specimenCount id="071F7F6C60BFCE5311C5C5628636C2CD" count="1" type="male">1 ♂</specimenCount>
|
||||
. Leg. R. B & E. M.
|
||||
<collectingRegion id="DC49AFEA1BC1ADE398BA1B35A2227E34" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
; Provincia de Burgos, S of Balneario de Corconte, strandline detritus.
|
||||
<date id="F982E3E66E913D2B576214D0FDBE7F27" value="2014-04-01">1. iv. 2014</date>
|
||||
. leg. R. B.
|
||||
<collectingRegion id="3C75201BBEAC5FE9C1307788E05EF96C" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="69D44C25AECB04CD0A8BD50BC6EA419E" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<named-content id="44C45A8436C156C7704D677B226B863A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="87086F4BD6F0C89CCA40E2ACCC3E1B63" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
); Provincia de Álava, Vittoria au Mt Gorbea,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERPDM" specific-use="{"type":"Point","coordinates":[-2.782000,34.634000]}">
|
||||
<geoCoordinate id="D11B5AE8BC0932C5004BBD4EDE6BE0A6" degrees="34.634" direction="north" orientation="latitude" precision="55" value="34.634">34.634 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="689E1994BBCD3ABA2486960098DE75F1" degrees="2.782" direction="west" orientation="longitude" precision="55" value="-2.782">2.782 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<date id="2E7B4931ADAA8E396F23B8AB8CACCFA3" value="1879-06">Juin 1879</date>
|
||||
.
|
||||
<specimenCount id="67C573C39D2923D7E7283F51566E0EEF" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EWPDM" xlink_title="Muséum d’histiore naturelle, Genève">
|
||||
<collectionCode id="73519E68157BE81E643BE3A885E3B680">MNHG</collectionCode>
|
||||
</abbrev>
|
||||
); Provincia de
|
||||
<collectingRegion id="F308F246C7D6A9A256D9AF0EF21501DC" country="Spain" name="Madrid">Madrid</collectingRegion>
|
||||
, Puerto de la Morcuera,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4PDM" specific-use="{"type":"Point","coordinates":[-3.399000,40.828000]}">
|
||||
<geoCoordinate id="1C3139CA0BB3ED95B783A78F37EE22AE" degrees="40.828" direction="north" orientation="latitude" precision="55" value="40.828">40.828 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EA2076BFA653CB531C2AC3F504598B8A" degrees="3.399" direction="west" orientation="longitude" precision="55" value="-3.399">3.399 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
. 15.1
|
||||
<date id="14A798966E64AD850501211FA1636D17" value="1997-05">v. 1997</date>
|
||||
. 2, unsexed (
|
||||
<named-content id="533E5F699A00F2997CBBE44EF0D76545" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="9F57021E2FB08BDE56DCBCBA342D80E0" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="D62844A6E389A46B978E8D4AE075B5F4">
|
||||
P: Monchique, Algarve,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELQDM" specific-use="{"type":"Point","coordinates":[-8.557000,37.317000]}">
|
||||
<geoCoordinate id="E04D6C67E99250C00BC51736E430F96E" degrees="37.317" direction="north" orientation="latitude" precision="55" value="37.317">37.317 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="6FCF9050A840A451C8DA7561415B09C0" degrees="8.557" direction="west" orientation="longitude" precision="55" value="-8.557">8.557 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="5EAF0CC8C2C70EA65420D265B5F7F157" value="1910-05-06" valueMax="1910-05-13" valueMin="1910-05-06">6–13. v. 1910</date>
|
||||
, leg. K.
|
||||
<collectingCountry id="325652BB40314D6C0DF8C4F880580F93" name="Jordan">Jordan</collectingCountry>
|
||||
.
|
||||
<specimenCount id="509F1409F1C102FF8A83B944975C6C3F" count="3" type="male">3 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="3A4A29C547036C53BD986907E57D55C2" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="B5041832D85A39A8017EF0C56F2974E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="49C4977620F1C7897892EB7A482B085C" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
); Serra de Estrela,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXQDM" specific-use="{"type":"Point","coordinates":[-7.708000,40.326000]}">
|
||||
<geoCoordinate id="FBA874C57C71B21CD659263E2762A447" degrees="40.326" direction="north" orientation="latitude" precision="55" value="40.326">40.326 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="CFC46883EBEA9D998470F1E03E6B6315" degrees="7.708" direction="west" orientation="longitude" precision="55" value="-7.708">7.708 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
. leg. H. Fery.
|
||||
<specimenCount id="18D53FC0E150FA7751DC0418C2874E95" count="2" type="male">2 ♂♂</specimenCount>
|
||||
, 1 unsexed (
|
||||
<abbrev id="ABBRID0E3QDM" xlink_title="Zoologische Staatsammlung München">
|
||||
<collectionCode id="288234EC96C18B3EE524CE3653CA2B56" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EARDM" type="Differential diagnosis">
|
||||
<paragraph id="FB4A18637327067EADFFCB5FE07C1745">
|
||||
<heading id="E56BE20AB41865952AE2FC412B97641E" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DD9C23D0E7F141FFC2C1CB0933167234">
|
||||
Endophallic teeth conspicuously shorter than in all other species, ca as high as long. Length of longest teeth ca 35 μm (Fig.
|
||||
<figureCitation id="67DE15C5DD45219D2CF95936ABF85CBC" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">
|
||||
<quantity id="74535DBEB2531EFEA7A6F0EC73B9BE1E" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" unit="g" value="23.0">23 g</quantity>
|
||||
– i
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="2754882719023E4032DA01227BFDB866" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 o</figureCitation>
|
||||
), as against at least 40 μm in other species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EORDM" type="redescription">
|
||||
<paragraph id="A279E5A62BF604E68893749732388A51">
|
||||
<heading id="340F64D293B37238147FC266BFBB3BC4" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8EB0090C09A4844C79408550DBB97609">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="47551F9E04864C73E03303A0D42EBB69" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">
|
||||
<quantity id="1B44E8FBA09C35CDB0507C034997F7A3" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" unit="g" value="4.0">4 g</quantity>
|
||||
</figureCitation>
|
||||
). Length
|
||||
<quantity id="091B415A686E504E64EDA0AB8B0EFEA7" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.8" metricValueMin="3.3" unit="mm" value="4.05" valueMax="4.8" valueMin="3.3">3.3–4.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="153F169C1187E294496B2615E0EBFCDE" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="E0AC1B4A303E2C1EC9798EF3363E0567" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" unit="mm" value="4.4" valueMax="5.0" valueMin="3.8">3.8–5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="C06E330B2E747B3C6AE1299712C0D57D" count="1" type="female">♀</specimenCount>
|
||||
), width
|
||||
<quantity id="1862760067BDAF33DDA1CDD7F0261D5E" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.6–2.1 mm</quantity>
|
||||
(
|
||||
<specimenCount id="44F1F3BBC70FC6849EC04E9A01AC99C0" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="7F01EC86660EE09314EE586BD00ECDD0" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7–2.2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="4CC7BBBDF12673BFD2D86809ABAB6E32" count="1" type="female">♀</specimenCount>
|
||||
). Black without obvious paler edges to head, antennae, palpi, and legs black to very dark brown. Head with the frontoclypeal suture very fine, often more or less effaced, laterally almost always so, and without tubercles. Clypeus bulging upwards medially, with double punctation, this finer on disc. Epipharynx (Fig.
|
||||
<figureCitation id="D6C4510C5FB532D625EC4B25F46884B5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 h, i</figureCitation>
|
||||
) with anterior margin of clithra strongly emarginate either side of the projecting median tylus, with a few very fine acropariae. Chaetopedia well-developed, stout, 6–8 each side. Chaetopariae shorter and thinner than chaetopedia, forming a closely-set line each side. Apophobae forming a line of very fine hairs outside the chaetopariae, prophobae lying in small inconspicuous groups near the sides of the mesoepitorma.
|
||||
</paragraph>
|
||||
<paragraph id="54E221F5A9D9F4DE8FF6C05982370391">
|
||||
Head and pronotum (Fig.
|
||||
<figureCitation id="FD365D8C9F0F1C909040B538E1966C7E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 k, l</figureCitation>
|
||||
): pronotum highly arched transversely and generally distinctly arched longitudinally, giving a somewhat domed appearance, the hemicylindrical appearance less usual than in other
|
||||
<taxonomicName id="C6C9240C5595AF7373EC856BE92B89A2" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="D5075272543D73E7800FE1B166BB01CD" italics="true">Liothorax</emphasis>
|
||||
species.
|
||||
</taxonomicName>
|
||||
Surface of pronotum bulging outwards laterally so that the lateral margins are sometimes not visible from above in their basal 1 / 3 s. Surface with double punctation, the strength of this variable, from rather fine (Fig.
|
||||
<figureCitation id="4C4A6BADD5A3375B0804F00725D83A78" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 k</figureCitation>
|
||||
) to distinctly coarser.
|
||||
</paragraph>
|
||||
<paragraph id="A994D6E22306B5FEA82A87BC36E64AD3">Scutellum elongate, pentagonal, glossy with a few punctures medially.</paragraph>
|
||||
<paragraph id="0CF1D869D5082367441BF51FD7FB8ED0">
|
||||
Elytra black, interstices with very fine reticulation (Fig.
|
||||
<figureCitation id="6685AFF3673CBD8EA7FDD17903FA57CB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 w</figureCitation>
|
||||
) and widely separated fine punctures, ca 8 × the width of the striae. Striae glossy, vertical-sided, with punctures separated by ca 2 × their diameter and encroaching on the vertical sides of the striae (Fig.
|
||||
<figureCitation id="D8B359B87289ED53B0274E831534A036" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 w</figureCitation>
|
||||
). Metaventrite (Fig.
|
||||
<figureCitation id="C60394BA5BED50B1F1B3F04BBA4E2826" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 u – w</figureCitation>
|
||||
) with central diamond-shaped area bearing sparse fine or moderate punctures and often small patches of reticulation. No obvious sexual dimorphism. Legs dark brown, longer spur of mesotibiae clearly slightly longer than basal segment of mesotarsus (Fig.
|
||||
<figureCitation id="7014CF3C6DEAE576B8A1BA7BBB9569A9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 o</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="74A93D9069B4C0DF61A361423863998A">
|
||||
Aedeagus (Figs
|
||||
<figureCitation id="4C918F3CA927747075CD7079208C56DE" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">
|
||||
<quantity id="242D8B5E964FC21EAE2CED80F87AD7D7" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" unit="g" value="23.0">23 g</quantity>
|
||||
– i
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="35F974FE2C7D0C317F765CC72C024321" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 o</figureCitation>
|
||||
). Length ca
|
||||
<quantity id="FB65098B340687820B8A84F0227903B7" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
, paramere length ca
|
||||
<quantity id="61F9F2608C1193C33175382BFD249C3D" metricMagnitude="-4" metricUnit="m" metricValue="4.3" unit="mm" value="0.43">0.43 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="D525ED26A31C2B43E34D54E18CDB2977" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" unit="mm" value="0.69">0.69 mm</quantity>
|
||||
, length of endophallic tooth-field ca
|
||||
<quantity id="705E78B04847E69DF80DAF2037535117" metricMagnitude="-4" metricUnit="m" metricValue="3.7" unit="mm" value="0.37">0.37 mm</quantity>
|
||||
, length of longest teeth ca 35 μm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EKTDM" type="remarks">
|
||||
<paragraph id="5CA9860423F5DDACE7503FC7979F3A49">
|
||||
<heading id="F7D623301C0D0CAD21DAF04EBD7F6A55" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="AD3D6214B6D283CAB02E67656B117340">
|
||||
<taxonomicName id="236789B3D23AB74EC6808C850A1B8DA6" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae">
|
||||
<emphasis id="4CEEFD0EFA25E88BC4DFDDBABE41D0D0" italics="true">Aphodius wilsonae</emphasis>
|
||||
</taxonomicName>
|
||||
stat. rest. is widely distributed mainly on higher ground in the northern half of
|
||||
<collectingCountry id="75810F8FABAC5FF0A64FFC406B9DB326" name="Spain">Spain</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="3AA9064D564DE8B4F198DFC1C6B909D1" name="Portugal">Portugal</collectingCountry>
|
||||
, but with some material from the Algarve (this location needs to be verified by future sampling; Fig.
|
||||
<figureCitation id="0714839A7DEA0F2380FFF70C9319B30C" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
). It is generally taken in the spring and early summer, either in detritus at the edges of water bodies (Fig.
|
||||
<figureCitation id="39C5D506D72CA7139A8DB1760ABE8C4D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 f</figureCitation>
|
||||
), or at the roots of vegetation where pools have dried out. It is sometimes very abundant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
375
data/88/7F/D0/887FD09C04295E11A580FB578A3EFEB5.xml
Normal file
375
data/88/7F/D0/887FD09C04295E11A580FB578A3EFEB5.xml
Normal file
|
|
@ -0,0 +1,375 @@
|
|||
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<mods:titleInfo id="050D53A53945D8FD639A0B80AA83DC69">
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<mods:title id="E918D78631CE482F00D1489888320B24">Bivalves of superfamily Galeommatoidea (Mollusca, Bivalvia) from western South Africa, with observations on commensal relationships and habitats</mods:title>
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<mods:affiliation id="93CD98342FE624B4A9C1107F5D483961">Santa Barbara Museum of Natural History, 2259 Puesta del Sol Road, Santa Barbara, CA 93105, USA</mods:affiliation>
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<mods:affiliation id="E9A7164B821471D88943C0FD72E288FD">Department of Biological Sciences, University of Cape Town, Private Bag X 3, Rondebosch 7701, South Africa</mods:affiliation>
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<mods:affiliation id="367E7346BB7791DC6DA8A03708697E49">Sea Change Trust, Cape Town, Western Cape, South Africa & Department of Botany and Zoology, Stellenbosch University, Private Bag X 1, Matieland, 7602, South Africa</mods:affiliation>
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<mods:affiliation id="9A5EA93923B650FDBCC9BCBDA4AA8E12">Department of Ecology and Evolutionary Biology, University of Colorado Boulder, 1900 Pleasant Street, Boulder, CO 80309, USA</mods:affiliation>
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<mods:namePart id="D08A1F68A137A6F8098F091AB447D65C">Li, Jingchun</mods:namePart>
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|
||||
<mods:start id="DD3BA763AF4E995C34457BFE143CC72B">301</mods:start>
|
||||
<mods:end id="E001A35030CCBC114D3476ECD2E102C2">323</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="7A5768BA18FF20AF476C8A6A53BAAF07">journal article</mods:classification>
|
||||
<mods:identifier id="CAA982B0FA4AACDC5FD2F53F7FEEB01B" type="DOI">10.3897/zookeys.1207.124517</mods:identifier>
|
||||
<mods:identifier id="8D8220BB2CC71F71812FD984E7F39505" type="ZooBank">682C6C4D-D59A-4D51-9D2B-9ABDEB841CD6</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="887FD09C04295E11A580FB578A3EFEB5" ID-DOI="http://doi.org/10.5281/zenodo.12796864" ID-Zenodo-Dep="12796864" ID-arpha="887FD09C-0429-5E11-A580-FB578A3EFEB5" ID-zoobank="https://zoobank.org/25211096-71D8-41CA-A5CF-DADEDA4CD18B" LSID="urn:lsid:plazi:treatment:887FD09C04295E11A580FB578A3EFEB5" LSID-ZBK="urn:lsid:zoobank.org:act:25211096-71D8-41CA-A5CF-DADEDA4CD18B" httpUri="http://treatment.plazi.org/id/887FD09C04295E11A580FB578A3EFEB5">
|
||||
<subSubSection id="CB10AFBA4365C46D97419302FEBAAD12" type="nomenclature">
|
||||
<paragraph id="5A7629B2ABEDBFF29AFCEEFEC55C6518">
|
||||
<taxonomicName id="741605C8E91C15C6EC35645A0C3A986B" authorityName="Valentich-Scott & Griffiths & Landschoff & Li & Li" authorityYear="2024" class="Bivalvia" family="Montacutidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="species" species="ducentiunus" status="sp. nov.">
|
||||
<emphasis id="D5B46C53603FA8E5926A5BB533AA11ED" italics="true">Brachiomya ducentiunus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4367838668BBF9B05D29B05B299C52E8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="DAD96713AFD5766C642A619CAA01B428" type="description">
|
||||
<paragraph id="8D2A430874F1DB4BF20E4DD7BB3DABD6">
|
||||
<figureCitation id="1FF0E477ADD4F3333550B45D26C31628" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Brachiomya ducentiunus sp. nov. A, B holotype, SBMNH 713162, length 2.50 mm, height 1.75 mm A exterior of right valve B exterior of left valve C – H paratypes, SBMNH 666970, crystallization on exterior and interior of shell is from dried bleach, not a sculpture element C exterior of right valve, length 2.17 mm, height 1.38 mm D exterior of left valve, length 2.40 mm, height 1.56 mm E interior of left valve, length 1.87 mm, height 1.19 mm F interior of right valve, length 2.42 mm, height 1.67 mm G hinge of right valve H hinge of left valve." figureDoi="10.3897/zookeys.1207.124517.figure1" httpUri="https://binary.pensoft.net/fig/1097710">Figs 1 A – H</figureCitation>
|
||||
,
|
||||
<figureCitation id="B0CD8737F0F8BE73E01E290B7B1E46F1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Brachiomya ducentiunus new species, living animals A, B crawling on hard substrate C overview of specimens crawling on the aboral surface of urchin Spatagobrissus mirabilis D crawling on urchin spine with mantle and foot extended E, F extended mantle between urchin spines." figureDoi="10.3897/zookeys.1207.124517.figure2" httpUri="https://binary.pensoft.net/fig/1097711">2 A – F</figureCitation>
|
||||
<xref id="1339AFC80E62C6233AB105EE1DF356FA" ref-type="supplementary-material" rid="S2">, Suppl. material 2</xref>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6PAE" type="type locality">
|
||||
<paragraph id="CEBC2A9F4F36A7F6C8BC5AEFCEED3344">
|
||||
<heading id="830AA1D2F402F570B4F1E23E8917AF6D" reason="title">Type locality.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="3E31FAF0199B41EA4DB525FE8AEDA790">
|
||||
Miller’s Point Lagoon, in False Bay,
|
||||
<collectingRegion id="2D94A137E0F59BC02CAA280425FBFBCE" country="South Africa" name="Western Cape">Western Cape Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="8532C17C5F25CCCC2701D3CE19C53CAA" name="South Africa">South Africa</collectingCountry>
|
||||
;
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIQAE" specific-use="{"type":"Point","coordinates":[18.477000,-34.231000]}">
|
||||
<geoCoordinate id="A6591618CE50F522F57147CA2EA9BFFB" degrees="34.231" direction="south" orientation="latitude" precision="55" value="-34.231">34.231 ° S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="692354F6FB9EF341C4F04FD2E4423D91" degrees="18.477" direction="east" orientation="longitude" precision="55" value="18.477">18.477 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
;
|
||||
<quantity id="0A86D978ABEEBA4B4024B3E4A7ADC023" metricMagnitude="0" metricUnit="m" metricValue="3.0" unit="m" value="3.0">3 m</quantity>
|
||||
; attached to spines, or crawling amongst spines of
|
||||
<taxonomicName id="7737D43A3D488DE68DB9AE640DE04080" authority="(Clark, 1923)" baseAuthorityName="Clark" baseAuthorityYear="1923" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" phylum="Echinodermata" rank="species" species="mirabilis">
|
||||
<emphasis id="45891FAAD0CAB564F129A0B93A9310D7" italics="true">Spatagobrissus mirabilis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="C19E815CF848753AE42A4F4D0A2B14AF" author="Clark HL" journalOrPublisher="Annals of the South African Museum" pagination="221-435" refId="B8" refString="Clark HL (1923) Echinoderm fauna of South Africa. Annals of the South African Museum 13: 221 - 435. [pls. 8 - 23]" title="Echinoderm fauna of South Africa." volume="13" year="1923">Clark, 1923</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, collected by Charles Griffiths,
|
||||
<date id="46A6C51D4356BF69C0ABCF42079BDEF4" value="2018-07">July 2018</date>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EYQAE" type="type material">
|
||||
<paragraph id="FE36A743AFE6BD76B95AB256E4A1D5FC">
|
||||
<heading id="A7587728C12232C1888AB5132A873072" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="90B75098F8015F64C437A39A510D1A63">
|
||||
<materialsCitation id="5FAA1D7DE9CBC23285603024BB186358" collectingDate="2016-06-09" collectionCode="SBMNH, UCM" collectorName="Jannes Landschoff & Craig Foster" specimenCode="SBMNH 713162, SAMC-A 096817, SBMNH 666970, UCM 60476" specimenCount="21" typeStatus="holotype">
|
||||
<emphasis id="5B521984558C2D6930B4AB75ECC5CDD4" bold="true" italics="true">
|
||||
<typeStatus id="6D11E0FCBD0655E746B308CF18DC9F99" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="C485E016BF38453F7944705C15EE3111" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Brachiomya ducentiunus sp. nov. A, B holotype, SBMNH 713162, length 2.50 mm, height 1.75 mm A exterior of right valve B exterior of left valve C – H paratypes, SBMNH 666970, crystallization on exterior and interior of shell is from dried bleach, not a sculpture element C exterior of right valve, length 2.17 mm, height 1.38 mm D exterior of left valve, length 2.40 mm, height 1.56 mm E interior of left valve, length 1.87 mm, height 1.19 mm F interior of right valve, length 2.42 mm, height 1.67 mm G hinge of right valve H hinge of left valve." figureDoi="10.3897/zookeys.1207.124517.figure1" httpUri="https://binary.pensoft.net/fig/1097710">1 A, B</figureCitation>
|
||||
),
|
||||
<specimenCode id="F804D995549686E0525E2E45F9657E00" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" type="Museum">
|
||||
<named-content id="F9365F79F2B4E6A4FD57D258DA1B99A3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/santa-barbara-museum-natural-history" xlink_title="Santa Barbara Museum of Natural History">SBMNH</named-content>
|
||||
713162
|
||||
</specimenCode>
|
||||
, length
|
||||
<quantity id="8DB849E5CA385548ECCFD796A8DA546F" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.50 mm</quantity>
|
||||
, height
|
||||
<quantity id="EB71333F3F35B97390D7F3A844AFFC83" metricMagnitude="-3" metricUnit="m" metricValue="1.75" unit="mm" value="1.75">1.75 mm</quantity>
|
||||
, preserved in 70 % EtOH.
|
||||
<specimenCount id="F73E1515B4E804E2CF8F4609D61882C3" count="13" type="generic" typeStatus="paratype">
|
||||
13
|
||||
<emphasis id="40EF574DA4405F21BC306C9079A87CB5" bold="true" italics="true">Paratypes</emphasis>
|
||||
</specimenCount>
|
||||
,
|
||||
<specimenCode id="1A0D9466D59CA5691904DB7D1B4AD9A8" collectionCode="SAMC-A">SAMC-A 096817</specimenCode>
|
||||
, same locality as holotype, largest specimen length
|
||||
<quantity id="EC8ABA572B9A28B93010E7B9752EC6C7" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
|
||||
, height
|
||||
<quantity id="AC5E0251C31DD95EC5070EB226D9E878" metricMagnitude="-3" metricUnit="m" metricValue="1.7" unit="mm" value="1.7">1.7 mm</quantity>
|
||||
, preserved in 70 % EtOH, collected by
|
||||
<collectorName id="E973DE7C1EF1F9B6021338489B1A9A2E">Jannes Landschoff</collectorName>
|
||||
and
|
||||
<collectorName id="13273A68B16184940F7B8431DC2EEC4E">Craig Foster</collectorName>
|
||||
,
|
||||
<date id="2412CCD1125DAF569D785D63CB3F21A6" value="2016-06-09">
|
||||
<collectingDate id="C0E7913E175E7037E37B5B2FA3330A0D" value="2016-06-09">9 June 2016</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<specimenCount id="A2083305C750AF8B0D992FCC34719273" count="7" type="generic" typeStatus="paratype">
|
||||
7
|
||||
<emphasis id="40D08409D435681342ACE800FC0E72AA" bold="true" italics="true">Paratypes</emphasis>
|
||||
</specimenCount>
|
||||
,
|
||||
<specimenCode id="4B2831F0F54BF53684885313B4ABE226" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" type="Museum">
|
||||
<named-content id="DBA9AD58778B5B26A8CC1797D3E84F3F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/santa-barbara-museum-natural-history" xlink_title="Santa Barbara Museum of Natural History">SBMNH</named-content>
|
||||
666970
|
||||
</specimenCode>
|
||||
, same locality and collector as
|
||||
<typeStatus id="BFE3BC79BC3793415B06D0588B7A3710" type="holotype">holotype</typeStatus>
|
||||
(Fig.
|
||||
<figureCitation id="2CA3B8551FE429AD5E3448CD08133F24" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Brachiomya ducentiunus sp. nov. A, B holotype, SBMNH 713162, length 2.50 mm, height 1.75 mm A exterior of right valve B exterior of left valve C – H paratypes, SBMNH 666970, crystallization on exterior and interior of shell is from dried bleach, not a sculpture element C exterior of right valve, length 2.17 mm, height 1.38 mm D exterior of left valve, length 2.40 mm, height 1.56 mm E interior of left valve, length 1.87 mm, height 1.19 mm F interior of right valve, length 2.42 mm, height 1.67 mm G hinge of right valve H hinge of left valve." figureDoi="10.3897/zookeys.1207.124517.figure1" httpUri="https://binary.pensoft.net/fig/1097710">1 C – H</figureCitation>
|
||||
), dried specimens mounted on SEM stub; length
|
||||
<quantity id="143A30F1CCF09DB4210AAD7649DA0580" metricMagnitude="-3" metricUnit="m" metricValue="2.17" unit="mm" value="2.17">2.17 mm</quantity>
|
||||
, height
|
||||
<quantity id="31ACE22AD3303430D88F69BFD27A533E" metricMagnitude="-3" metricUnit="m" metricValue="1.38" unit="mm" value="1.38">1.38 mm</quantity>
|
||||
; length
|
||||
<quantity id="763AAB09F7C9CEB3D749C97205941E2A" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.40 mm</quantity>
|
||||
, height
|
||||
<quantity id="BC29A0C4678F777CD04F2787D48118E5" metricMagnitude="-3" metricUnit="m" metricValue="1.56" unit="mm" value="1.56">1.56 mm</quantity>
|
||||
; length
|
||||
<quantity id="C72101940B4DAD31E5F15D56A479CBE6" metricMagnitude="-3" metricUnit="m" metricValue="1.87" unit="mm" value="1.87">1.87 mm</quantity>
|
||||
, height
|
||||
<quantity id="128C1946AC005B50EE5E4E05B8E19561" metricMagnitude="-3" metricUnit="m" metricValue="1.19" unit="mm" value="1.19">1.19 mm</quantity>
|
||||
; length
|
||||
<quantity id="D525C899457ED0E7264F3F4E4349BB96" metricMagnitude="-3" metricUnit="m" metricValue="2.42" unit="mm" value="2.42">2.42 mm</quantity>
|
||||
, height
|
||||
<quantity id="4B85AC2C00E2865B296D46518E02D470" metricMagnitude="-3" metricUnit="m" metricValue="1.67" unit="mm" value="1.67">1.67 mm</quantity>
|
||||
.
|
||||
<specimenCount id="A3E53FAB5B7A86232A41603DC0A17C2D" count="1" type="generic" typeStatus="paratype">
|
||||
1
|
||||
<emphasis id="83F3E0EAC397B9FB133B0A0984E2BE79" bold="true" italics="true">Paratype</emphasis>
|
||||
</specimenCount>
|
||||
,
|
||||
<specimenCode id="AAD411FFD35C094C906726BA4A0E7E99" collectionCode="UCM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" type="University or college">
|
||||
<named-content id="3BD993CAF87BB22DF4CABEA8E2788D85" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-colorado-museum-natural-history" xlink_title="Museum of Natural History, University of Colorado">UCM</named-content>
|
||||
60476
|
||||
</specimenCode>
|
||||
; length
|
||||
<quantity id="DDE5B2DF41F3991C9ACE4D363A825374" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
|
||||
, height 1.0 mm.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<caption id="AA2E03021661FF5A4BBD893D3CD074AB" ID-DOI="10.3897/zookeys.1207.124517.figure1" ID-arpha="2455F526-63FA-5D2A-953E-8995C8018E91" httpUri="https://binary.pensoft.net/fig/1097710" startId="F1">
|
||||
<paragraph id="48256D27FB4D5863E52F7821A419EA37">
|
||||
<label id="9F925F917AA448754D34ABCEEADC53C4">Figure 1.</label>
|
||||
</paragraph>
|
||||
<paragraph id="A7621A808686F80E690BF64052F74D26">
|
||||
<taxonomicName id="E982F473D5E4677CBA05E19E2AF4644B" authorityName="Valentich-Scott & Griffiths & Landschoff & Li & Li" authorityYear="2024" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="species" species="ducentiunus" status="sp. nov.">
|
||||
<emphasis id="69ECACBFE51444E79C7C7E9C3BD8AC55" italics="true">Brachiomya ducentiunus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="BEB27E9B3F9608C5220780A9A4C5C5E3" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="2B198C0902069929EB6E388DAB8C8735" bold="true">A, B</emphasis>
|
||||
holotype,
|
||||
<named-content id="FEDC10A632CF4F085EDD5A4FF6047C09" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/santa-barbara-museum-natural-history" xlink_title="Santa Barbara Museum of Natural History">SBMNH</named-content>
|
||||
713162, length 2.50 mm, height 1.75 mm
|
||||
<emphasis id="8787A9DC584134B039A0D1B5CABA4BEB" bold="true">A</emphasis>
|
||||
exterior of right valve
|
||||
<emphasis id="9D957541B78D7E44BC7D711BEB5E0D4F" bold="true">B</emphasis>
|
||||
exterior of left valve
|
||||
<emphasis id="5C57DDED8E6885746D59BC36FBFAD00A" bold="true">C – H</emphasis>
|
||||
paratypes,
|
||||
<named-content id="4EC110B744578BD54D5D9DFE87880BE6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/santa-barbara-museum-natural-history" xlink_title="Santa Barbara Museum of Natural History">SBMNH</named-content>
|
||||
666970, crystallization on exterior and interior of shell is from dried bleach, not a sculpture element
|
||||
<emphasis id="2E8AE2E8F004B7F28924D6D92C380B18" bold="true">C</emphasis>
|
||||
exterior of right valve, length 2.17 mm, height 1.38 mm
|
||||
<emphasis id="3D2E01EC3020AC0D31996F85E05A91A9" bold="true">D</emphasis>
|
||||
exterior of left valve, length 2.40 mm, height 1.56 mm
|
||||
<emphasis id="3EF832461E6B18F3F992967C6CC65FC2" bold="true">E</emphasis>
|
||||
interior of left valve, length 1.87 mm, height 1.19 mm
|
||||
<emphasis id="2324AFD2B4A0447090DD951741500FD7" bold="true">F</emphasis>
|
||||
interior of right valve, length 2.42 mm, height 1.67 mm
|
||||
<emphasis id="CC93B876B6A7B3BA3B4B4E1ECB68F63F" bold="true">G</emphasis>
|
||||
hinge of right valve
|
||||
<emphasis id="D84D8EBD4D366D03C6324A4A3270D2A7" bold="true">H</emphasis>
|
||||
hinge of left valve.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUTAE" type="description">
|
||||
<paragraph id="A7F7B4C10581B73EA65298DEC7C76AB7">
|
||||
<heading id="59339E61C77D1A342F3007C474B7CF86" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="9EB268C9A7EE0952F143576F840E8002">
|
||||
Shell extremely thin, fragile, moderately inflated, translucent; inequilateral, slightly longer anteriorly; ovate-elongate; anterior end obliquely truncate in larger specimens; posterior end broadly rounded; ventral margin straight, slightly invaginated in some; dorsal margin gently sloping from umbos; shell margins weakly gaping; prodissoconch well defined, umbonate, smooth, subcircular; prodissoconch length ~ 350 μm; external sculpture of commarginal striae, with few widely spaced radial striae, especially anteriorly; umbos low, wide; hinge plate extremely narrow, with one minute pseudocardinal in each valve; ligament internal, very short. Length up to
|
||||
<quantity id="DB27F931AFC07DA01CC9AA7331D98726" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.7 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="E1C3B81CBEBA03B19C1990F48653EEBF">
|
||||
<emphasis id="3EB27DF3294BA669F48A5B9DCDF72D97" bold="true">
|
||||
<emphasis id="E54C37DAB2397E8A4C5F2C852B32F670" bold="true" italics="true">Mantle</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Large, reflected, covering ~ 95 % of outer shell surface when fully extended, but not fully covering umbos; mantle can be almost completely retracted into the shell; reflected portion with low papillae; mantle near shell margin with longer tentacles; anterior end with large cowl, serrate on end; cowl can be greatly extended (Fig.
|
||||
<figureCitation id="E9896CF7330925AEDC60F5ECBB36B884" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Brachiomya ducentiunus new species, living animals A, B crawling on hard substrate C overview of specimens crawling on the aboral surface of urchin Spatagobrissus mirabilis D crawling on urchin spine with mantle and foot extended E, F extended mantle between urchin spines." figureDoi="10.3897/zookeys.1207.124517.figure2" httpUri="https://binary.pensoft.net/fig/1097711">2 E, F</figureCitation>
|
||||
; Suppl. material
|
||||
<xref id="E5856445988297EA9A5AF302EC48E528" ref-type="supplementary-material" rid="S2">2</xref>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="08AD0390FC606B4F430DABF1212267BB" ID-DOI="10.3897/zookeys.1207.124517.figure2" ID-arpha="7286E384-B002-532B-907B-608525F859A7" httpUri="https://binary.pensoft.net/fig/1097711" startId="F2">
|
||||
<paragraph id="989667C2C678E755581CD55F43BB476C">
|
||||
<label id="6F257B41385E34B0412ACF224EE85068">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="88EB9E5AF7CEF4964ED7C81404CD95A6">
|
||||
<taxonomicName id="981104C7166B7D79F8634DE0A6DF6E16" authorityName="Valentich-Scott & Griffiths & Landschoff & Li & Li" authorityYear="2024" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="species" species="ducentiunus" status="sp. nov.">
|
||||
<emphasis id="D47E987C452E2B229D673C8E0D9D9469" italics="true">Brachiomya ducentiunus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="3B9C5FBECD235A4F6F0C88D4FB9BE5E2" rank="species">new species</taxonomicNameLabel>
|
||||
, living animals
|
||||
<emphasis id="A95DDC93444AA647B5E50C64A5425EBD" bold="true">A, B</emphasis>
|
||||
crawling on hard substrate
|
||||
<emphasis id="6547B31A6621078FEAAC11EB2936BF95" bold="true">C</emphasis>
|
||||
overview of specimens crawling on the aboral surface of urchin
|
||||
<taxonomicName id="B0CF5C5DD5C27A7FBCCE9BC722E85F86" baseAuthorityName="Clark" baseAuthorityYear="1923" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" phylum="Echinodermata" rank="species" species="mirabilis">
|
||||
<emphasis id="CB310E001F452A20FA9B9C5B605852FA" italics="true">Spatagobrissus mirabilis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="1C17FB3B15231AA3AC01657D4B1D7F7D" bold="true">D</emphasis>
|
||||
crawling on urchin spine with mantle and foot extended
|
||||
<emphasis id="0157E1BD337484F5BDA648983087CD6D" bold="true">E, F</emphasis>
|
||||
extended mantle between urchin spines.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="56C94BC8614F86852DA1DAAB8C1184FD">
|
||||
<emphasis id="99F37389DD9F48E7D328BA5F55B7842B" bold="true">
|
||||
<emphasis id="12770513F5D77AC00F75299071DEB7B3" bold="true" italics="true">Foot</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Of moderate size, ~ the length of the shell when fully extended, vermiform, with slight heel. The species is an active crawler, and can also attach to the host by byssal threads. The foot has been observed to frequently wrap around the urchin spines as the bivalve crawls.
|
||||
</paragraph>
|
||||
<paragraph id="79E82573415970488AA9B3CD1BB0C707">
|
||||
<emphasis id="D9E03C5BE4FF46A24DCB6C5F15600279" bold="true" italics="true">Ctenidia</emphasis>
|
||||
. One demibranch on each side, comprised of ~ 30 narrowly spaced filaments in larger specimens.
|
||||
</paragraph>
|
||||
<paragraph id="A46DC2146DE8C995E95104A4A15C634F">
|
||||
<emphasis id="B93A8100EE2B338093B372D237205336" bold="true" italics="true">Brooding</emphasis>
|
||||
. Up to ten shelled juvenile specimens observed brooding in the dorsal portion of ctenidia in mature specimens.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EGWAE" type="distribution">
|
||||
<paragraph id="9FA58F8DF3CFAF8B55FD3997784F0E6B">
|
||||
<heading id="147C26E3FA38507DB4EA9A20BE0CEF0B" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="527E11917D53EBE71AB51864F37B7E70">
|
||||
Only known from the
|
||||
<typeStatus id="C5C0C565A403285343E5E758C3F6CD34">type</typeStatus>
|
||||
locality in False Bay,
|
||||
<collectingCountry id="217EC338266CD7387CEE54C112F42D13" name="South Africa">South Africa</collectingCountry>
|
||||
, and only found attached to the echinoid
|
||||
<taxonomicName id="70EB13AFDB6DBEEA598D5B3632F4C424" baseAuthorityName="Clark" baseAuthorityYear="1923" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" phylum="Echinodermata" rank="species" species="mirabilis">
|
||||
<emphasis id="1D2808C574A3967C6D49B3D2C242A6CF" italics="true">Spatagobrissus mirabilis</emphasis>
|
||||
</taxonomicName>
|
||||
; not observed free-living.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EXWAE" type="Commensal relationship and habitat">
|
||||
<paragraph id="DB26CBCE1579AF8A77B6A1E1F5B3FF5F">
|
||||
<heading id="5881ABEC55C241713AF57A6CE25F3190" reason="title">Commensal relationship and habitat.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="C7DE14A16221920E688EBA0E4EE089FD">
|
||||
Found crawling on the oral surface of the heart urchin
|
||||
<taxonomicName id="DBD839A627E7BE716176B3F05F1CB5EC" baseAuthorityName="Clark" baseAuthorityYear="1923" class="Echinoidea" family="Maretiidae" genus="Spatagobrissus" kingdom="Animalia" order="Spatangoida" phylum="Echinodermata" rank="species" species="mirabilis">
|
||||
<emphasis id="6E8D23AB86B9A76D2650D0ACFECDAEC5" italics="true">Spatagobrissus mirabilis</emphasis>
|
||||
</taxonomicName>
|
||||
. This host species was found to be living in a specialized microhabitat of coarse gravel and half-buried cobbles or boulders (at least at the
|
||||
<typeStatus id="4BC3C336DACDC355D1FDB923468F1350">type</typeStatus>
|
||||
locality associated with kelp forests). At the
|
||||
<typeStatus id="5CC71174CB3D5E18F016DBD9491AAF37">type</typeStatus>
|
||||
locality in 2018, of 10 sampled heart urchins, all had associated
|
||||
<taxonomicName id="A7352DD64E16952D113BE160868E9AC6" authorityName="Jespersen, Lutzen & C. Nielsen" authorityYear="2004" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="4D0FBDB58A95E847DB485AA069FD8DF3" italics="true">Brachiomya</emphasis>
|
||||
</taxonomicName>
|
||||
on their oral surface. Densities of
|
||||
<taxonomicName id="054AD1877348C713CFD2DBB487D6A4B3" authorityName="Jespersen, Lutzen & C. Nielsen" authorityYear="2004" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="23ADA83C2308BFE1B61F6F20C2E2A899" italics="true">Brachiomya</emphasis>
|
||||
</taxonomicName>
|
||||
ranged from 38 to
|
||||
<specimenCount id="7A93C655DB9CE69898CDA46BEC8A4677" count="172" type="generic">172 specimens</specimenCount>
|
||||
on a single host. Two other commensal species were also recorded on these same urchins, a small but very common unidentified amphipod of family
|
||||
<taxonomicName id="7056260BE39FF74D7F208E1F5E30EE40" authorityName="Dana" authorityYear="1849" class="Malacostraca" family="Lysianassidae" kingdom="Animalia" order="Amphipoda" phylum="Arthropoda" rank="family">Lysianassidae</taxonomicName>
|
||||
, and a large, scale worm (family
|
||||
<taxonomicName id="2AA74D6D5A5F27BD1C534F6D3E7DE196" authorityName="Kinberg" authorityYear="1856" class="Polychaeta" family="Polynoidae" kingdom="Animalia" order="Phyllodocida" phylum="Annelida" rank="family">Polynoidae</taxonomicName>
|
||||
), of which only a few specimens were found. The amphipod and polychaete species also both appear to be new to science.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EAYAE" type="Discovery">
|
||||
<paragraph id="A40A2252B212F85FEC3381D0BE56F418">
|
||||
<heading id="702E844EEEC4BCE510A023462BE5E93F" reason="title">Discovery.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="0BFCA301C8894FCE22ABA2A5549CB3D0">
|
||||
Initially discovered via free-diving in 2016 at the
|
||||
<typeStatus id="A043191279417DA2D8BA44E775DF667F">type</typeStatus>
|
||||
locality, collected by Jannes Landschoff and Craig Foster.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EFYAE" type="etymology">
|
||||
<paragraph id="B8E6976EC9CE02C7E46EBDA4265D02C3">
|
||||
<heading id="96FC15A183E881ED86EEEB8908592945" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="89249CE989AA947BBCBE77DFD842E1F7">
|
||||
The name
|
||||
<taxonomicName id="89A97D5468B9315158B34B3CD92FCB90" authorityName="Valentich-Scott & Griffiths & Landschoff & Li & Li" authorityYear="2024" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="species" species="ducentiunus">
|
||||
<emphasis id="A7A1693E0B54C25711A5204069747D8A" italics="true">ducentiunus</emphasis>
|
||||
</taxonomicName>
|
||||
is from Latin, meaning “ 201. ” The species was discovered while preparing and working on the ‘ 1001 Seaforest Species’ project, a research and storytelling program aimed at increasing awareness of regional kelp bed ecosystems colloquially referred to as ‘ the Great African Seaforest’ (see
|
||||
<ext-link id="B481CA4B5AB52776CDAC1942642D6FB6" ext-link-type="uri" xlink_href="http://www.seachangeproject.com" xlink_type="simple">www. seachangeproject. com</ext-link>
|
||||
). The number 201 was chosen as a unique identifier for the 1001 program, with the goal to link each hundredths species to a species described as new to science.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1YAE" type="comparisons">
|
||||
<paragraph id="FAC18A5FC339A8C8E9FB99ABB1910842">
|
||||
<heading id="2FE45C466E30CEC37D6E2D9B60D0AA7D" reason="title">Comparisons.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="C74C84B7D056DF75A22E5D277B8849FE">
|
||||
The Pacific and Asian
|
||||
<taxonomicName id="D2529E3AD8A96B4A120B5002D1B05047" baseAuthorityName="Pilsbry" baseAuthorityYear="1921" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" kingdom="Animalia" order="Galeommatida" phylum="Mollusca" rank="species" species="stigmatica">
|
||||
<emphasis id="CD12E7C733ABE1143055E235AF95D295" italics="true">Brachiomya stigmatica</emphasis>
|
||||
</taxonomicName>
|
||||
, which is the only other known species in the genus, is more evenly rounded anteriorly, has a strong rust-colored stripe medially, lacks radial striae, and has more developed teeth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
394
data/89/FA/22/89FA22F1B78356FA82264DE0E8904C5A.xml
Normal file
394
data/89/FA/22/89FA22F1B78356FA82264DE0E8904C5A.xml
Normal file
|
|
@ -0,0 +1,394 @@
|
|||
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<mods:title id="696CF86E88F9728838D8AAD8C8352F22">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:affiliation id="392F3F0DBBBE0FC73A1EBF211846A6C6">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="F1F2EE3DFD85ED2F7FA0B6ADFB9CBE5B">Král, David</mods:namePart>
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<mods:part id="4CB11A6BBCD5A0B53CD4593872B465F3">
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<treatment id="89FA22F1B78356FA82264DE0E8904C5A" ID-arpha="89FA22F1-B783-56FA-8226-4DE0E8904C5A" ID-zoobank="https://zoobank.org/F188AEBA-0E15-4EB4-B1AD-A1A49DBBA81E" LSID="urn:lsid:zoobank.org:act:F188AEBA-0E15-4EB4-B1AD-A1A49DBBA81E" LSID-ZBK="urn:lsid:zoobank.org:act:F188AEBA-0E15-4EB4-B1AD-A1A49DBBA81E" httpUri="http://treatment.plazi.org/id/89FA22F1B78356FA82264DE0E8904C5A">
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<paragraph id="4959C07983DF1FE881D804D9BF449603">
|
||||
<taxonomicName id="B650BFEF165B5001E765EE453332C972" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="968D6E73D9B2E2D69A6892A2222A3AB9" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="57EB2D55DFC398710C38B41ABF9C3414" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="3466D8BBB6369566B0476ECA7FA60B61" italics="true">bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="0EA0C270AF86279608E3F09561E43864" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="6403A1BE65E19CF35B9F9ECEF50AC4E0" type="description">
|
||||
<paragraph id="8CFBFA991E2DCB0A6CD2D4E16D82D7CA">
|
||||
<figureCitation id="8F6708A7C71C43CC4149C0BE978E65EB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 d</figureCitation>
|
||||
,
|
||||
<figureCitation id="44936AB42AD96FA83FADC6FB3BA5CECB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">
|
||||
<quantity id="D5A84FBDF1AF89E41E5E24D5E6DA3507" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" unit="g" value="6.0">6 g</quantity>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="B1871492A722AA84CB544BC97B348A8A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">
|
||||
<quantity id="617EB15DCFE09BA72DF53C41C191EEE3" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" unit="g" value="8.0">8 g</quantity>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="3C29AE976F2489902B310A0721B5CEAD" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 a, x</figureCitation>
|
||||
,
|
||||
<figureCitation id="53293AD60F94646BD92854CDF856B522" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 k, l, u 21 p – r, 23 j – l, 24 b, 25 f, 26 w, x, 33 a – d, 34 h</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EV5BM" type="type material">
|
||||
<paragraph id="9AE38723201A86E22DB1BD8786FD8AD7">
|
||||
<heading id="166C74E0940632B1E798043F298D22FC" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="9A56CD9692A413D209ABBA4E3AA48DF5">
|
||||
<materialsCitation id="54DAA3DC35D3F4B447BAAFBFFE04DDFF" collectingDate="2011-06-19" collectingDateMax="2011-06-29" collectingDateMin="2011-06-19" collectionCode="NHMUK" collectorName="R. B. and E. M. Angus. Chromosome & R. B. Angus" country="France" location="Haute-Corse" municipality="Lac de Melo. Cow" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis id="2E8342FE13B733CB680209EDC34D2F76" bold="true" italics="true">
|
||||
<typeStatus id="E23B17E21750685EE4D7B14636AA1085" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="FFF2D1D8C068CC996CD1A65E31829824" count="1" type="male">♂</specimenCount>
|
||||
:
|
||||
<collectingCountry id="5867A7D5C0C2ACC3E7B4F4C5435DC275" name="France">Corsica</collectingCountry>
|
||||
,
|
||||
<location id="7DB68E8E4B2C6DFAA8DF08FA66ED76BE" LSID="urn:lsid:plazi:treatment:89FA22F1B78356FA82264DE0E8904C5A:7DB68E8E4B2C6DFAA8DF08FA66ED76BE" country="France" municipality="Lac de Melo. Cow" name="Haute-Corse">Haute-Corse</location>
|
||||
, pelouse (lawn) grassland by the
|
||||
<collectingMunicipality id="2142CD4E87CFFD2D6D8B83E6FC0C8C96">Lac de Melo. Cow</collectingMunicipality>
|
||||
dung.
|
||||
<date id="F8BE2E364D9BEB8738241480319158CC" value="2011-06-19">
|
||||
<collectingDate id="1F9C6CADDFA7BF0B23A1F316A2479333" value="2011-06-19">19. vi. 2011</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="94033126B81FD91387F4D419D2250500">R. B. and E. M. Angus. Chromosome</collectorName>
|
||||
prep. 2,
|
||||
<date id="DA8CB012F6344FDC597FB68E62010807" value="2011-06-29">
|
||||
<collectingDate id="FC4C56D50E00EF804154F7F7B694DA92" value="2011-06-29">29. vi. 2011</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="AB2756052D9CFB37B5DC2EE9D9746911">R. B. Angus</collectorName>
|
||||
(
|
||||
<named-content id="C371B60534EBE59C89DEDC10B829729A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="5E72E48A9C68CA4F7DB2EC0EB3563838" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="2424EA28795847691986AE98D9DD3D7E" bold="true" italics="true">
|
||||
<typeStatus id="69C713278DD77AEBCCC661C15B5BE22E" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="5ACD977FC7259EB323C75C8DB2E3CAC1" count="2" type="male">2 ♂♂</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="D6966975E2FCFA1A51BEB1C845019777" type="holotype">holotype</typeStatus>
|
||||
, 1 labelled Chromosome prep. 1,
|
||||
<date id="FDBA8C344812055B484B10A6CAC11CAD" value="2011-06-29">29. vi. 2011</date>
|
||||
. R. B. Angus;
|
||||
<specimenCount id="DD0AD4380A2657CBA92C225281176A8A" count="4" type="male">4 ♂♂</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="4BEBB98CA3060C5BE4657893B9FEAE55" type="holotype">holotype</typeStatus>
|
||||
but
|
||||
<date id="DA9509F449212D93D37AC14BAF17B113" value="2011-06-24">24. vi. 2011</date>
|
||||
;
|
||||
<specimenCount id="C06D52B4572F62B3D99A5B505AD565DF" count="14" type="female">14 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="65A4C1DD5F7CD6704BBD2998A6021FA7" type="holotype">holotype</typeStatus>
|
||||
, including Chromosome preps 1–4,
|
||||
<date id="2BFBD21AD6E1138F73EDC87DDE7C4124" value="2011-06-28">28. vi. 2011</date>
|
||||
, 6–9,
|
||||
<date id="98707D8DB485C5232F362986320A8824" value="2011-06-28">28. vi. 2011</date>
|
||||
and 1–3,
|
||||
<date id="9FDC870F38ADC5FF87E96978622968F4" value="2011-07-01">1. vii. 2011</date>
|
||||
, R. B. Angus;
|
||||
<specimenCount id="E9CDB222CB5A13E895FE02823D573FD0" count="1" type="female">1 ♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="42798F9E575BB16CB389186BE6D6152C" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<named-content id="CD6E06F015BDB9EB8FA764DC9BDD6ACD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="DB003CBB94BF6A562A3B052C94F92936" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="C3477FB3727D803F15ECC6536CC203A2" count="11" type="female">11 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="D60F4E6FCE9E7A0EEACC324C3B2641C1" type="holotype">holotype</typeStatus>
|
||||
but
|
||||
<date id="B69BF0570BF405569AB3A95483EE01A1" value="2011-06-24">24. vi. 2011</date>
|
||||
;
|
||||
<specimenCount id="943265CA87A6FD9647018FAE9CBDD24A" count="2" type="male">2 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="3D1F5E58E6CF5B25098E5B910D481F52" count="5" type="female">5 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="4C13255A76EC43EBAA09CE8030EBDAA9" type="holotype">holotype</typeStatus>
|
||||
but
|
||||
<date id="ADAFD364FD3568DDD7259E961AA91889" value="2009-06-30" valueMax="2009-07-01" valueMin="2009-06-30">30. vi and 1. vii. 2009</date>
|
||||
, R. B. and E. M. Angus.
|
||||
<specimenCount id="C02828300C7BAC47EAA10451424D7772" count="1" type="male">♂</specimenCount>
|
||||
Chromosome preps 1 and 2,
|
||||
<date id="AD7F6A934827065178FD1FCB0713D555" value="2009-07-06">6. vii. 2009</date>
|
||||
,
|
||||
<specimenCount id="C598B1B3519A290F0A93D60F4987611F" count="1" type="female">♀</specimenCount>
|
||||
Chromosome preps 3–5,
|
||||
<date id="57B9BFF3784878DAECAE78586EFE7B87" value="2009-07-05">5. vii. 2009</date>
|
||||
(
|
||||
<named-content id="7BD3DD69FCEA5768EC5894ABA60EE055" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="5F67E80A8C1E3CF5ACF13B3C676C9660" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<specimenCount id="607D71270EF9FE875584F4DAE5EBFCF8" count="1" type="male">1 ♂</specimenCount>
|
||||
data as previous (
|
||||
<named-content id="EA49EBB95C4D5EB7C1B483D98BDF6FF9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="205BEC133A35C52D039622A09BD3647D" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="1314E4F5C1CF9C7D40531485C2F0C41B" count="1" type="male">1 ♂</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="1D1D4A21FE4A62508A1BCCBBD48A0624" type="holotype">holotype</typeStatus>
|
||||
but
|
||||
<date id="A69894FA1956FCD17DD9F660DD29E6BA" value="2011-06-24">24. vi. 2011</date>
|
||||
, in F. Krell collection (
|
||||
<named-content id="BED8FFD216BD97D43C0BE1FAAF3535CE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/denver-museum-nature-science" xlink_title="Denver Museum of Nature and Science">
|
||||
<collectionCode id="D8F55307E46D8C21307488FD8DD46B58" country="USA" httpUri="http://grbio.org/cool/mmzs-jqix" name="Denver Museum of Nature and Science" type="Museum">DMNS</collectionCode>
|
||||
</named-content>
|
||||
). 3
|
||||
<collectingCountry id="95CCD003A97C9753F5B73DB2A25924EC" name="France">Corsica</collectingCountry>
|
||||
, Haute-Corse, pelouse (lawn) grassland by the Lac de Melo. Cow dung.
|
||||
<date id="188222486DA01615E9B64CA08E8595E6" value="2011-06-19">19. vi. 2011</date>
|
||||
. R. B. and E. M. Angus (sequenced) (
|
||||
<abbrev id="ABBRID0EX6BM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="B0157977E412DCDC92357D887DB9F669">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
<specimenCount id="2F3C28969F21246B4E34054F88AF2177" count="7" type="male">7 ♂♂</specimenCount>
|
||||
, 23 unsexed, “
|
||||
<date id="15A252BA882FC27530947AE6DCFCFA81" value="1983-07-18">18. vii. 1983</date>
|
||||
.
|
||||
<collectingCountry id="14777FE6113052ACFDC81BEE15F4F20B" name="France">Corse</collectingCountry>
|
||||
. Lac de Nino
|
||||
<quantity id="309BAC91DCB2E30E03DE749C736DE8FB" metricMagnitude="3" metricUnit="m" metricValue="1.7429999999999999" unit="m" value="1743.0">1743 m</quantity>
|
||||
. Rinderkot. Leg. H. Fery ”. In H. Fery collection (
|
||||
<abbrev id="ABBRID0E26BM" xlink_title="Zoologische Staatsammlung München">
|
||||
<collectionCode id="BF8DEB9F0318402DCBAB84589E4AEB22" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||||
</abbrev>
|
||||
); 2 unsexed, “
|
||||
<collectingCountry id="1091A8ACEE73E1C8733A0E94F8525BFF" name="France Corse">France Corse</collectingCountry>
|
||||
Nino-See
|
||||
<quantity id="DC8C36F577A371B9B1F644270A939F4E" metricMagnitude="3" metricUnit="m" metricValue="1.73" unit="m" value="1730.0">1730 m</quantity>
|
||||
.
|
||||
<date id="51D2465215A7FB00C21478AD3B79462D" value="1987-07-12">12. vii. 1987</date>
|
||||
. Leg H. Hirschfelder. ” In H. Fery collection (
|
||||
<abbrev id="ABBRID0E66BM" xlink_title="Zoologische Staatsammlung München">
|
||||
<collectionCode id="0BFCD459F332D240AC770B33574A8037" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||||
</abbrev>
|
||||
). In
|
||||
<abbrev id="ABBRID0EDACM" xlink_title="Carsten Zorn Collection">CZC</abbrev>
|
||||
and
|
||||
<abbrev id="ABBRID0EHACM" xlink_title="Jason Maté Collection">JFMC</abbrev>
|
||||
:
|
||||
<specimenCount id="210F40C694A3A3BBE9060AEFB3C43745" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="62AE284BEEEFDE66458FF136613E0D24" count="1" type="female">1 ♀</specimenCount>
|
||||
, 1 unsexed
|
||||
<collectingCountry id="1C482FBB53E1DB4027528A2A008A8C49" name="France">France</collectingCountry>
|
||||
:
|
||||
<collectingCountry id="2D3720496E399297D7F1FE527BBB09C5" name="France">Corsica</collectingCountry>
|
||||
Plateau d´Alzo
|
||||
<quantity id="86DAE67C28B3DF6545A2B6247141007D" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</quantity>
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOACM" specific-use="{"type":"Point","coordinates":[9.066667,42.266667]}">
|
||||
<geoCoordinate id="8A4AAF98CAD71C4A531E672F70F914BC" degrees="42" direction="north" minutes="16" orientation="latitude" precision="925" value="42.266666">42 ° 16 ' N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="987FF2F5F367A00DC59C7D27B93A33C4" degrees="9" direction="east" minutes="04" orientation="longitude" precision="925" value="9.066667">9 ° 04 ' E</geoCoordinate>
|
||||
</named-content>
|
||||
<date id="7AB20C154BEA24D187BA2D4148F0C3BC" value="1994-05-21">21 / V / 1994</date>
|
||||
leg. C. Zorn.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ETACM" type="Differential diagnosis">
|
||||
<paragraph id="061ACE1AAF8FD0FE495CE10EEFD2DD68">
|
||||
<heading id="565DEDE9A0C52D37472CA1E72CED94B9" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="2745DEE00EEBE543B1CE8C536DA5D486">
|
||||
The parallel-sided, hemicylindrical, lightly sculptured appearance (Fig.
|
||||
<figureCitation id="6E46BAE3CACD1EBF17D37CAEB2852826" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 d</figureCitation>
|
||||
) is distinctive. The pronotal sides bulge outwards laterally so that the lateral margins are not visible from above in basal 1 / 2 of the pronotum. The lateral margins themselves are parallel, only very slightly convergent anteriorly. The general shape of the beetles and their pronotal configuration resembles those of
|
||||
<taxonomicName id="2028C1DD3B56E3F25DA7D9D6C3183BB5" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="E7ADD62A6E2A8F00185A8756D61B8679" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="3CB7CBD5FF4209AC72ABBA0D01DBDFB5" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||||
<emphasis id="4EFDF6A0A06B6758846E1A974DA9ADB5" italics="true">A. muscorum</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="1DF90DCBBC6CBFA362E5A62AF0C236D0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 a, b</figureCitation>
|
||||
), but these species are more heavily sculptured.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EXBCM" type="description">
|
||||
<paragraph id="E0EAEF268E87277A3998D74DACC190A0">
|
||||
<heading id="C824E3E39CE90EF98277E07A47B804FB" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F3317C57AA3A8A42E59056486C963280">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="FEE4555D87FBD424E4D5F7D4C558543E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 d</figureCitation>
|
||||
). Length: 4.0–
|
||||
<quantity id="A313EA87083E6AF73D6A3F74E172C2DA" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
(
|
||||
<specimenCount id="35F9AE2E0CABB9CDDC91797028125DEF" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="9E9999E353EFB0075B18206E9D20B62F" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.8" metricValueMin="3.9" unit="mm" value="4.35" valueMax="4.8" valueMin="3.9">3.9–4.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="F55874CFD3561FD3969E6A72A960F7EC" count="1" type="female">♀</specimenCount>
|
||||
); width:
|
||||
<quantity id="A4B63D0B5E113E130BCF66A412F00FA4" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 mm</quantity>
|
||||
(
|
||||
<specimenCount id="A8D1470FCB9208A26EAB08622ACE72E1" count="1" type="male">♂</specimenCount>
|
||||
), 1.6–2.0 mm (
|
||||
<specimenCount id="2A7EBF870F5049D878F1FAC9872FBC27" count="1" type="female">♀</specimenCount>
|
||||
). Glossy black, tarsi, tibial spurs dark brown, femora and tibiae dark brown with black-bronze reflections. Head: dorsal surface domed, flatter behind frontoclypeal suture. Medial area of clypeus more demarcated as a dome than in
|
||||
<taxonomicName id="DEA9EC82F577594CAFCEEB083C244F7F" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="C1A7D29721A98FA1CCE6AA405EB4D852" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
. Frontoclypeal suture distinct, completely non-tuberculate but with distinct smooth elevated areas each side of the median straight section and a weaker elevation medially. Genae rounded, strongly protuberant laterally in front of eyes. Clypeus with rounded median emargination, angles either side of this rounded. Clypeus and genae with strong raised margin, this slightly brownish. Surface strongly and more or less evenly punctuate, the punctures separated by ca 1.5 × their diameter, and finer medially. Antennae and palpi more or less black. Epipharynx (Fig.
|
||||
<figureCitation id="52AC326A77BDA746E4C6692F6E389453" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">
|
||||
<quantity id="EAEBD66EBE1371E1EC2FEDA92AB2AEB8" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" unit="g" value="8.0">8 g</quantity>
|
||||
</figureCitation>
|
||||
) clithra evenly excised either side of the median tylus, chaetopedia with 4–6 spines, these slightly less robust than those of
|
||||
<taxonomicName id="DDF9C0EC3FFC8B4FE7D293845F10319F" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="7E52B8EC8CFDFAF9CDBCB1C1EEC4EB1C" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
and spines of chaetopariae as long as or longer than chaetopedia. Head and pronotum as in Fig.
|
||||
<figureCitation id="364271476D6DDA0E0F64AA94F545A12F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">
|
||||
<quantity id="1211D65ABC767BE0CAD62D13766FFCD5" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" unit="g" value="6.0">6 g</quantity>
|
||||
</figureCitation>
|
||||
. Pronotum hemicylindrical, highly arched transversely but more or less flat longitudinally, lateral margins more or less parallel, but very slightly convergent anteriorly, weakly and very evenly curved. Pronotal surface bulging outwards from the general curvature over the basal 1 / 2 of the pronotum, ca 1 / 4 of the way in from each side, so that the lateral margins are not visible from above in the basal 1 / 2 of the pronotum (Fig.
|
||||
<figureCitation id="60EA80914716C4185B2CE0D85EB8399A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">
|
||||
<quantity id="CF73718325527C122DF31F2C42DE5FBF" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" unit="g" value="6.0">6 g</quantity>
|
||||
</figureCitation>
|
||||
). Lateral margins with distinct raised border that continued very finely over the lateral ca 1 / 3 of the basal margin, at least the median 1 / 3 of which lacks any border (Fig.
|
||||
<figureCitation id="F32F7555E400DAACEA5888172EDBB6B5" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 a</figureCitation>
|
||||
). Anterior margin without any trace of a raised border. Surface with double punctation of variable strength, but in general weaker than in
|
||||
<taxonomicName id="FE95C479DE598DD7627AD2DA38E4E1A1" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix" status="sp. nov.">
|
||||
<emphasis id="FDC71393DD7270E6B79D3EB4B1379972" italics="true">L. felix</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D2411335298822BA40D1A2485E9082F4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, though the strength of the punctation of the two species overlaps. Larger punctures separated by 2–4 × their diameter, sometimes very strongly impressed and with the pronotal surface depressed immediately round their edges, but sometimes more moderate and with the pronotal surface evenly curved around them. Finer punctures dot-like, in some specimens separated from one another by ca 2 × their diameter but in other specimens much sparser, separated by 4–6 × their diameter. Pronotal surface between the punctures with a silky sheen and a very fine, generally indistinct, small, isodiametric reticulation.
|
||||
</paragraph>
|
||||
<paragraph id="26C6CD871374BB43A99067D22078D31C">Scutellum elongate, pentagonal, ca 10 % of elytral length, glossy black with brownish lateral and apical edges, and with sparse punctures medially.</paragraph>
|
||||
<paragraph id="73E09E2071EA542F86B8A4C64D85888C">
|
||||
Elytra glossy black but the interstices slightly duller than head and pronotum, appearing slightly silky and with a very fine indistinct isodiametric reticulation sometimes visible. Striae narrow (ca 1 / 5 the width of the interstices), vertical sided and with punctures separated by ca 2 × their diameter. In the
|
||||
<specimenCount id="0EA91AF07413CD3DD9F4297C84C9EEA7" count="2" type="male">two ♂♂</specimenCount>
|
||||
the punctures bulge into the interstices but in the
|
||||
<specimenCount id="6DCC87180443B8A16F7C1E07B643B4ED" count="3" type="female">3 ♀♀</specimenCount>
|
||||
they hardly deflect the strial margins. Lateral margins of the elytra distinct, at base strongly upcurved in front of the humeral bulges; stronger basally and at apex ca as wide as stria 2. Interstices 4 × as wide as striae, with fine sparse punctation (Fig.
|
||||
<figureCitation id="4B24064C869771F6C728DDD596CA4B97" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 x</figureCitation>
|
||||
), which is a bit stronger in the humeral area in front of abbreviated stria 9.
|
||||
</paragraph>
|
||||
<paragraph id="379FA43D1BD9AE0DE6311419AE5012D3">
|
||||
Legs (Fig.
|
||||
<figureCitation id="30F30DE96B535FA7AE18CCCCCA2A5981" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 k, l, u</figureCitation>
|
||||
) dark brown with bronze-black reflections and tarsi and tibial spurs brown, rather long and slender, basal segment of hind tarsi as long as segments 2 + 3 + 1 / 2 of segment 4. Longer spur of mid tibiae longer than basal segment of mid tarsi.
|
||||
</paragraph>
|
||||
<paragraph id="C460019F3A050CEEAA1CDB930BDE004F">
|
||||
Metaventrite (Fig.
|
||||
<figureCitation id="120F81C68582DD24CF76FA3BACCA98AF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 p – r</figureCitation>
|
||||
): median diamond-shaped area rather lightly punctate, without pubescence, its mid-line either almost indistinguishable or variably depressed, sometimes forming a distinct longitudinal furrow up to 1 / 3 as wide as the median point of the diamond. There is no sexual dimorphism.
|
||||
</paragraph>
|
||||
<paragraph id="B4661395D225C344450409AB2F28F5FC">
|
||||
Aedeagus (Figs
|
||||
<figureCitation id="BEA2F09A891C362E2E5725F3AEE78771" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 j – l</figureCitation>
|
||||
,
|
||||
<figureCitation id="DB22D8751563BAFC6F2BA598636F560F" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 b</figureCitation>
|
||||
) Length ca
|
||||
<quantity id="E78DB371FA703A7D129EE01FEAAFFA61" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
|
||||
, paramere length ca
|
||||
<quantity id="9C916F487194415B1017A9B0FD9D9C24" metricMagnitude="-4" metricUnit="m" metricValue="4.5" unit="mm" value="0.45">0.45 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="38DB86EFF5CA9C40EDBCAA58F5B071D0" metricMagnitude="-4" metricUnit="m" metricValue="7.6" unit="mm" value="0.76">0.76 mm</quantity>
|
||||
, length of endophallic tooth-field ca
|
||||
<quantity id="04B066F541091C4B6FBD7C7704C7433A" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
|
||||
. Length of longest teeth on endophallus ca 44 μm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPECM" type="etymology">
|
||||
<paragraph id="36D10CACC28CCE33C57A12EF63725968">
|
||||
<heading id="46AC9B4F265D219B94B3FA03B12B5ABD" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6DB05FD8B36BCC6181CC1B0FFCA701B8">
|
||||
Named after our good friend Dr Franck Bameul of Bordeaux, who accompanied RBA and EMA to the Lac de Melo on the
|
||||
<date id="226CF6D3585FA59F6B9490E347826E23" value="2011-06-24">24. vi. 2011</date>
|
||||
and was with us as we all endured a prolonged and spectacular thunderstorm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUECM" type="remarks">
|
||||
<paragraph id="B320965FB184781B720FD6E74ED76FE7">
|
||||
<heading id="E3E436D25EBF685318D98F00E25488BC" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="293667F02D48C75C705C197BFFF87968">
|
||||
<taxonomicName id="282CB37553EA2F6F009B4F9B4095D987" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bameuli" status="sp. nov.">
|
||||
<emphasis id="7B63306F7872CA6C70D55300051EE389" italics="true">Aphodius bameuli</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4F14CDFA02A7063EEE0227DCCBED0416" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is endemic to
|
||||
<collectingCountry id="D7A637D0E165A1CD05B66CD7D61DE7C2" name="France">Corsica</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="E8B9969B8D887E279734220616464487" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
) and, as far as we know, is the only
|
||||
<taxonomicName id="E22776BB7C86436050F6C9BDF565F0B8" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="D8DB11AA50C2C31C7A060A76CB0A6CEA" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
definitely associated with dung. On the Melo lawns (Fig.
|
||||
<figureCitation id="E18F2E52A42470F7AB3E779302B5129A" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 e</figureCitation>
|
||||
) it occurred in cow pats, avoiding the more liquid regions towards the centre of the dung. It ate the dung, which filled the guts of specimens used for chromosome work. Hans Fery (pers. comm.
|
||||
<date id="EA5EC6FEB08F8FB170B12D50528FBAED" value="2015-09">ix. 2015</date>
|
||||
) told us that when he collected it by the Lac de Nino the water was very high, covering the lakeside lawns but that the cow pats protruded above the water surface and that is where the beetles were found. RBA and EMA failed to find it in the cow dung on the lawns in the valley of Pozzi (Ghisoni), further south.
|
||||
</paragraph>
|
||||
<paragraph id="9D0C2A31922EB0F98DF5FB140F82EC6B">It is an elongate, parallel-sided, and lightly sculptured species.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
363
data/90/E9/1B/90E91BD08620595391E3513E8E367030.xml
Normal file
363
data/90/E9/1B/90E91BD08620595391E3513E8E367030.xml
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|
|
@ -0,0 +1,363 @@
|
|||
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<mods:titleInfo id="5E48F271C5CC3E415DC8EA4B502CE942">
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<mods:title id="EC3C60787BC192E100016EA17EE59BCE">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:namePart id="0932EB82EB8BF2858A112982827E033F">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="3CA590021356A9E2945E12319E5E4565">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="F5BA9182771B46B6B2726A22D48F44F2">Maté, Jason F.</mods:namePart>
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<mods:nameIdentifier id="A6BAE4C87C507F4C005262B095B351F8" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
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<mods:affiliation id="A20F54E24B35418CC54F774F4A74292F">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="CE5D24138DA0C9306D604D8BF5BF5BB5">Angus, Elizabeth M.</mods:namePart>
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<mods:affiliation id="E26CEB65EC5BD329FFA5D803DC3DC250">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="AEECDC4BEBCABED5E0358404C787C161">Král, David</mods:namePart>
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<mods:affiliation id="E17C072F0610524314C2612B41F15B16">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:title id="0338869EAAFCF78F37DEF5EA91ABFD4E">ZooKeys</mods:title>
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<mods:number id="F3B4978CB6C0D52A876BF8CCB62CBFF5">1207</mods:number>
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<mods:classification id="19FABD6F94C2EF9D93FEDB1F653A3D73">journal article</mods:classification>
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</mods:mods>
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<treatment id="90E91BD08620595391E3513E8E367030" ID-arpha="90E91BD0-8620-5953-91E3-513E8E367030" LSID="urn:lsid:plazi:treatment:90E91BD08620595391E3513E8E367030" httpUri="http://treatment.plazi.org/id/90E91BD08620595391E3513E8E367030">
|
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<subSubSection id="65870A866783F32ADA66A6D9A98A318A" type="nomenclature">
|
||||
<paragraph id="E70137CC5BF2FE8B7FC55D097F481220">
|
||||
<taxonomicName id="C554E8567A6DD4E8A07692972197545C" ID-CoL="FG45" authority="(Balthasar, 1952)" baseAuthorityName="Balthasar" baseAuthorityYear="1952" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis" subGenus="Liothorax">
|
||||
<emphasis id="447EFBD0193C92F64D52D64B3195DB08" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="EEE9C133BDF044ABE53FC7B6B529BDB2" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="1196F35DD12AAE5391E7046F57B53F39" italics="true">isikdagensis</emphasis>
|
||||
(Balthasar, 1952)
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="BA112B749AE69C243E82076D38DDDBEF" type="description">
|
||||
<paragraph id="0B6CDEE4A66429F70F8336D0849D0A44">
|
||||
<figureCitation id="046D26F8D40F5FC3EE5BE9D6A88AA0D7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">Figs 3 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="7B68DD6BC0C2E0B02DA38FBCA57332ED" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="12725CC6E85CAD2D4A3BF59A097D451C" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 e</figureCitation>
|
||||
,
|
||||
<figureCitation id="AA7A6DF2AB7D2735B3F6662F49227392" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 s</figureCitation>
|
||||
,
|
||||
<figureCitation id="3A814B47EA927A72D7A396EC3E0B4E59" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 c</figureCitation>
|
||||
,
|
||||
<figureCitation id="0F43D90E8E7A3F278C2836F27880E414" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 c ’</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="7392C51AB88B47091AC5DB27DF8CCD27" type="reference_group">
|
||||
<paragraph id="10B4259CDFE212CD1F0D4F32F316BA24">
|
||||
<treatmentCitationGroup id="CD373184827A41F93ACD177D27E874C2">
|
||||
<treatmentCitation id="35C9CB826B776D2733031223BFE80948">
|
||||
<taxonomicName id="720C1CE90DDAB2CA0A6F5CF1BA567FAC" ID-CoL="FG45" authority="Balthasar, 1952: 22" authorityName="Balthasar" authorityPageNumber="22" authorityYear="1952" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis" subGenus="Ataeniomorphus">
|
||||
<emphasis id="6F76EF614D038491E83962839EC8F49B" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="5850481B11339F610F4A1D1C3A9C0F43" italics="true">Ataeniomorphus</emphasis>
|
||||
)
|
||||
<emphasis id="3A10E2C73F01F394D73DCD042DC40FEB" italics="true">isikdagensis</emphasis>
|
||||
Balthasar, 1952: 22
|
||||
</taxonomicName>
|
||||
.
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZUCM" type="type material">
|
||||
<paragraph id="5D0EF45012580B3990E0697B8F2997D3">
|
||||
<heading id="6726B70D99ABE42CC0FC961038264374" reason="title">Type material studied.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BB7F798045D787A27FD8550ACA7FDF7F">
|
||||
<materialsCitation id="EF28DA710CC1A263502346F10624200B" collectorName="Dr V. Balthasar" location="Isik" municipality="Camlidere" specimenCount="9" specimenCount-female="1" specimenCount-male="1" stateProvince="Mus" typeStatus="holotype">
|
||||
<emphasis id="423CF0CE2C50FE639E4D9516BCADA597" bold="true" italics="true">
|
||||
<typeStatus id="8A94FFD84F3F8B8572B220722D2C628D" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(labelled as
|
||||
<specimenCount id="4D193358DA2CD70882308EF70841435B" count="1" type="male">♂</specimenCount>
|
||||
, undissected), with the locality data as given above and a label “
|
||||
<taxonomicName id="6BB092FE05C6467B56674BB2D1D89541" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis" status="sp. nov." subGenus="Ataeniomorphus">
|
||||
<emphasis id="A642C34A37A99342DD401232285851BB" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="4328D360184CD2DEB0CA2D81344F09EE" italics="true">Ataeniomorphus</emphasis>
|
||||
)
|
||||
<emphasis id="78EF67471ADA8F19E87A5FB99D4ABE25" italics="true">isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A967790916D502BB927B59EF53D34866" rank="species">n. sp.</taxonomicNameLabel>
|
||||
,
|
||||
<collectorName id="AF33324972641E260CB3B905B43870D6">Dr V. Balthasar</collectorName>
|
||||
”. At present
|
||||
<taxonomicName id="A996C372296EA80D0182FC569BBD6F3D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="746A2DA0B3B329B7F548BB7CA67037B5" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
is known only from the type series, the
|
||||
<typeStatus id="6E80FC2F7935944BF8F31A50F8B03D8E" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="5D572F79642AF16F90E61DB1463C073E" count="7" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
|
||||
, one labelled as
|
||||
<specimenCount id="FCBD9B4F9AD2ADBBF1C6294BF8F505CC" count="1" type="female">♀</specimenCount>
|
||||
<typeStatus id="E2381FD3862414B7A359EAB2C67F4676" type="allotype">allotype</typeStatus>
|
||||
, with the locality data
|
||||
<collectingMunicipality id="9879105057DF7B400404895CB7D8EA3B">Çamlidere</collectingMunicipality>
|
||||
,
|
||||
<location id="4D5BFDA2887820A73DC4637905FFB81F" LSID="urn:lsid:plazi:treatment:90E91BD08620595391E3513E8E367030:4D5BFDA2887820A73DC4637905FFB81F" municipality="Camlidere" name="Isik" stateProvince="Mus">Isik</location>
|
||||
d., Anat. 23. vi. 47. Exp. N.
|
||||
<collectingRegion id="7F157FFF61ACB2B5AC36431B1A5198DF" country="Turkey" name="Mus">Mus</collectingRegion>
|
||||
. ČSR
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="BE303158334CCBD703BB45445461F821" bold="true" italics="true">
|
||||
<typeStatus id="7DE4ED20A2EDD894D8A6E87EF804E2D3" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: 1 labelled as
|
||||
<specimenCount id="2E06F8CA041CFE06D33DDE6E71A5890A" count="1" type="female">♀</specimenCount>
|
||||
<typeStatus id="3466DF1A73BE2C63D94DC1CEA7724E9B" type="allotype">allotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="07814658E95FDF3FEB7872627B197C34" count="6" type="generic" typeStatus="paratype">6 paratypes</specimenCount>
|
||||
, noted by Balthasar as
|
||||
<specimenCount id="8C25C7AD20A9E8C0965D10EA2F33B92A" count="4" type="male">4 ♂♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="97CEE925A6AC2D6066668651A88BACFA" count="2" type="female">2 ♀♀</specimenCount>
|
||||
. Thus
|
||||
<specimenCount id="7D308C0AB075E0F24EE6F040A4E04151" count="1" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||||
is unaccounted for. Balthasar does not appear to have dissected any of the types (though a male
|
||||
<typeStatus id="9DED1AA143240D53B4BF8992F7686A5E" type="paratype">paratype</typeStatus>
|
||||
had been dissected) and apparently distinguished the sexes on the degree of clypeal bulging and the degree to which the genae were angled out at their junction with the clypeus. Both these characters are clear in two of the
|
||||
<typeStatus id="30021B42B51A0A1451D17B9947EE2DA7" type="paratype">paratypes</typeStatus>
|
||||
(Figs
|
||||
<figureCitation id="06CA743060C276A9BED4ACB9EDECE50F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="1202AB8CB6AF63C4B60F8BE330ECC26D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 a, b</figureCitation>
|
||||
) the largest of the
|
||||
<specimenCount id="56201888B1A7EBC8CD451E81B79110AF" count="1" type="male">♂♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="1DB8600E577F14C9E487E73527A0C7EF" count="1" type="female">♀♀</specimenCount>
|
||||
, but not in two other
|
||||
<specimenCount id="EC1B547577F0BAD4CE6C1485CDA66A5A" count="1" type="female">♀♀</specimenCount>
|
||||
where the reduced clypeal bulging is clear, but not the outward angling of the genae. Assuming
|
||||
<specimenCount id="BBBF47B151851352D89461334684EC58" count="1" type="male">1 ♂</specimenCount>
|
||||
<typeStatus id="6B7A0374D628F720E89CD31DE62BFBB7" type="paratype">paratype</typeStatus>
|
||||
is elsewhere, the
|
||||
<specimenCount id="D9E829B451F21FBDB72D0E9698820459" count="6" type="generic" typeStatus="paratype">6 paratypes</specimenCount>
|
||||
noted by Balthasar comprise
|
||||
<specimenCount id="8C0B486FE70E3ADB0078C1895BCBEF43" count="2" type="male">2 ♂♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="4507FC8BA3F77D2A4DF5384451104313" count="4" type="female">4 ♀♀</specimenCount>
|
||||
. It may be that Balthasar inadvertently switched the numbers of males and females. Balthasar gives the length range as
|
||||
<quantity id="B3D7E7B834D76D47F340910BEF692388" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
|
||||
. Our measurements are based on stacked images.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EIWCM" type="material">
|
||||
<paragraph id="BE306F09FA458E21BD628ACA57FADB71">
|
||||
<heading id="C2DED4A75BEDD3178606553B361CD515" reason="title">Additional material studied.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1BBA6A5E1081228316BA14E4C54DAC23">
|
||||
<specimenCount id="7A786A7B32D4DA89583B1583F55706A1" count="1" type="male">One male</specimenCount>
|
||||
,
|
||||
<collectingCountry id="CF2B2D52C3D9569E61AA3FD8DEA53EFF" name="Turkey">Turkey</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="26E73072647D7599479C62D24D37B5D9" country="Turkey" name="Artvin">Artvin</collectingRegion>
|
||||
Çam gec.
|
||||
<quantity id="EAC041A1EEAB82ACAE99875CADE685BA" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
|
||||
, coll. Ziani, shown by its aedeagus (Fig.
|
||||
<figureCitation id="071A9BA02BA1E45DEDB9A7B286A11619" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 k</figureCitation>
|
||||
) to belong to the
|
||||
<taxonomicName id="57FE9ED5E618EFFF46ACA82AB2A10331" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="2F895BA5DBC75658C1287E755872E62D" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
section, may belong here. Its length is
|
||||
<quantity id="69D0BDB5138587884D1F500DE29C4265" metricMagnitude="-3" metricUnit="m" metricValue="4.64" unit="mm" value="4.64">4.64 mm</quantity>
|
||||
, width
|
||||
<quantity id="0E9B5120ECF0BCEA1D4C5D2FFEAFD3DC" metricMagnitude="-3" metricUnit="m" metricValue="1.89" unit="mm" value="1.89">1.89 mm</quantity>
|
||||
, so it is within the size range of
|
||||
<taxonomicName id="218607D55AC722333F812E987439B684" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="C1F4B5FF443F9C6D09C15AC6D12A62D4" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
, with which it also agrees in the brown, not blackish, legs, though the maxillary palpi are black. The aedeagus (Fig.
|
||||
<figureCitation id="753241223E80037D30962151ABCB9DF6" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 k</figureCitation>
|
||||
) in lateral view is shown to have the parameres only weakly downturned apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMXCM" type="Differential diagnosis">
|
||||
<paragraph id="69AD11E53C6A39C87AC8ACFBF41FD48E">
|
||||
<heading id="2DC56BBD81BF97EB453AF886B0E1C9E6" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5E49BE662BDBD676DB977A194EC90D7E">
|
||||
The larger of the two known Turkish species, identified as a member of the
|
||||
<taxonomicName id="15E9D0743D528C38C3F387E573D8D24A" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="0D30D1B7162523DAE9D894A4CF1FCC90" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
group by the large recurved endophallic teeth (Fig.
|
||||
<figureCitation id="407AD802C3EE4415666E49EB196B08FB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 c</figureCitation>
|
||||
). Maxillary palpi mid-brown. Epipharynx (Fig.
|
||||
<figureCitation id="9244C527A36BFFF7E5ABF8D8CB433A86" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 c, d</figureCitation>
|
||||
) with apophobae arranged in narrow bands ca two bristles wide.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EFYCM" type="redescription">
|
||||
<paragraph id="C4101C7610A523CDB22D63A48A2398C2">
|
||||
<heading id="078ABB4D66740572B03A8CE4F5310D44" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B263EFF543BA8D7C1F98BBB9798EB490">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="7427FC59478206DB0C770D77B167EDB9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 c, d</figureCitation>
|
||||
). Length
|
||||
<quantity id="24015F88F25050D687D8FFE07E527D9D" metricMagnitude="-3" metricUnit="m" metricValue="5.199999999999999" metricValueMax="5.8" metricValueMin="4.6" unit="mm" value="5.199999999999999" valueMax="5.8" valueMin="4.6">4.6–5.8 mm</quantity>
|
||||
, width
|
||||
<quantity id="AE81103C7E8E7D3D887919ECF87E4E15" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" unit="mm" value="2.2" valueMax="2.5" valueMin="1.9">1.9–2.5 mm</quantity>
|
||||
. Head black, browner towards the anterior and lateral margins, no tubercle on frontoclypeal suture. Frontoclypeal suture fine and straight-transverse over middle 1 / 3, at either end of this either effaced or angled forward to reach the edge of the head at the junction of the clypeus and genae. Outer margin of genae more or continuous with that of the frons in male, distinctly angled outwards in female (Fig.
|
||||
<figureCitation id="70B666A3C473CBFFF82784BBC899DC5E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 a, b</figureCitation>
|
||||
). Clypeus moderately bulging upwards medially in male, weakly so in the female, anterior margin broadly excised medially, the sides of this excision bluntly rounded. Punctation rather weak, sometimes stronger and tending to be rugose towards the anterior and lateral margins. Epipharynx (Fig.
|
||||
<figureCitation id="9E490632650ADF8EEF320D02A0D115FA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 c, d</figureCitation>
|
||||
) with the central tylus strongly projecting and the anterior margin of the clithra clearly excised either side. Central darkened sclerotised epitorma broadly triangular, the whole median darkened region appearing widest at base (Fig.
|
||||
<figureCitation id="98E7DA591186D5621A614BB91AE5A28C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 c</figureCitation>
|
||||
) or narrowed basally (Fig.
|
||||
<figureCitation id="FC771C9AE1316CD233B2F2BBCA331ECC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 d</figureCitation>
|
||||
). Acropariae virtually absent, chaetopedia strong, six or seven each side. Surface of gymnopedia covered with small tooth-like asperities. Prophobae quite strong, clustered at the edges of the sclerotised mesoepitorma, apophobae fine, arranged in a narrow slightly irregular band ca two bristles wide, outside the chaetopariae. Antennae and palpi mid brown. Pronotum (Fig.
|
||||
<figureCitation id="C25423DCD44A182E29678CBDE77BCF6F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 a, b</figureCitation>
|
||||
) hemicylindrical, highly arched transversely, scarcely at all longitudinally. Entire lateral margins visible from above. Surface with double punctation, male
|
||||
<typeStatus id="02A018866EB6C8AB5CF5BEAEA3762176" type="paratype">paratype</typeStatus>
|
||||
with the larger punctures separated by at least twice their diameter, sparser on disc. The female
|
||||
<typeStatus id="88649D10CBBB6672BE52DA6C8A599A78" type="paratype">paratype</typeStatus>
|
||||
has the punctation sparser and finer. Posterior margin completely bordered (Figs
|
||||
<figureCitation id="702DA8AD29CEF728E398011B54641992" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="46931B52D279A36FC80AF5C9EB1B6E4C" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 e</figureCitation>
|
||||
), the border wider medially, narrowed but continuous round the posterior corners.
|
||||
</paragraph>
|
||||
<paragraph id="10631723F142A4676A4349897EAB130B">
|
||||
Scutellum pentagonal, elongate, glossy, sometimes with a few punctures medio-basally. Elytra black, interstices flat, 6–8 × width of the striae, with fine isodiametric reticulation and sparse fine punctures (Fig.
|
||||
<figureCitation id="725972FD53ED04BFF502164AF70B87E4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 k</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="631889B8977A66318A107D10C47D0DA3">
|
||||
Metaventrite (Fig.
|
||||
<figureCitation id="871AE950431FDA7F870B94F1AAED9FA0" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 x, y</figureCitation>
|
||||
) moderately punctured, flattened over median diamond-shaped plate, sometimes with an impressed median line. Legs mid brown, basal segment of mesotarsi elongate, either clearly longer than the longer tibial spur, or ca the same length (Fig.
|
||||
<figureCitation id="30C018496FC5D387E2AFFCCC4193D087" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 s</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="C8EFBE2EF44E345BCC729B54978E5878">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="01E0D139A1602ED31D36516920578D53" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 c</figureCitation>
|
||||
): total length ca
|
||||
<quantity id="7CE8CC7BB786602D91B02B4A76923B98" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
, parameres length ca
|
||||
<quantity id="74FC1186AEA0F6043EE1FA802356C0FB" metricMagnitude="-4" metricUnit="m" metricValue="4.8" unit="mm" value="0.48">0.48 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="8E9300CB50F60B15AA693075E5C835EA" metricMagnitude="-4" metricUnit="m" metricValue="6.4" unit="mm" value="0.64">0.64 mm</quantity>
|
||||
, length of endophallic tooth-field ca
|
||||
<quantity id="91E600F7FBFF2B727401475E532AE754" metricMagnitude="-4" metricUnit="m" metricValue="7.0" unit="mm" value="0.7">0.70 mm</quantity>
|
||||
. Length of longest tooth on endophallus (measured from photograph) ca 60 μm. Parameres with darkened sclerotised strut running from near the outer margin at the base to the inner margin at apex, where the apical sensory pads run round the apex from the tip of the oblique strut to the outer edge, almost to its outer apical angle.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EB1CM" type="remarks">
|
||||
<paragraph id="CC042D1326BB4F75CD2B6F6E413595FF">
|
||||
<heading id="9B48E4566BF01789E42E4E0665947759" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F2F8C327E5C30BA3691A778DA70C7737">
|
||||
This species is identified as a member of the
|
||||
<taxonomicName id="817D0097029215FB202011CFC1A2B017" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="676B74CEB995CA8BE542568F83CC3A95" italics="true">Aphodius niger</emphasis>
|
||||
</taxonomicName>
|
||||
group by the large, recurved teeth on its endophallus (Fig.
|
||||
<figureCitation id="DC4E7BEE825E62A593856EC44300BFF7" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 c</figureCitation>
|
||||
), and the parameres with the apical sensory pads running round the apex from the tip of the oblique strut to the outer edge. This may be the result of collapse of only weakly downturned apically parameres as a result of drying.
|
||||
</paragraph>
|
||||
<paragraph id="D599BCEDD39251148C7396BE205BA022">
|
||||
<bibRefCitation id="1713F349D25B96E3435B922B2A904040" author="Dellacasa G & Dellacasa M & Ziani S" issue="2000" journalOrPublisher="Caffi ”" pagination="3-11" refId="B14" refString="Dellacasa G, Dellacasa M, Ziani S (2001 b) Taxonomic position of Aphodius turbatus Baudi, 1870 and systematic redefinition of Aphodius isikdagensis Balthasar, 1952. Rivista del Museo civico de Scienze naturali “ E. Caffi ” 20 (2000): 3–11." title="Taxonomic position of Aphodius turbatus Baudi, 1870 and systematic redefinition of Aphodius isikdagensis Balthasar, 1952. Rivista del Museo civico de Scienze naturali “ E." volume="20">Dellacasa et al. (2001 b</bibRefCitation>
|
||||
) regarded this species as conspecific with
|
||||
<taxonomicName id="9BAF8269649B5F709F21721E88BF49BE" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="2635B0E03E560A4066FFFA2127D4EB0F" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, at that time known as
|
||||
<taxonomicName id="253BEA198EC6DDCC6C9410DB777CCD07" authority="(Petrovitz, 1962)" baseAuthorityName="Petrovitz" baseAuthorityYear="1962" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ressli">
|
||||
<emphasis id="2D7DA012EF922AB69FC29C246FF84B7D" italics="true">A. ressli</emphasis>
|
||||
(Petrovitz, 1962)
|
||||
</taxonomicName>
|
||||
. They figure the aedeagus (without the endophallus in its sac), but the parameres do not really match those of the
|
||||
<typeStatus id="C223727FEDBE7C8D7A29B7BF73A06F7C" type="paratype">paratype</typeStatus>
|
||||
shown in Fig.
|
||||
<figureCitation id="DA23056DB56FF7056D34F3532CAA1715" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 c</figureCitation>
|
||||
.
|
||||
<bibRefCitation id="0BCFC28765A5E5BFD90FB48E4D5EA79F" DOI="10.11646/zootaxa.1407.1.4" author="Dellacasa M & Gordon RD & Dellacasa G" etAl="et al." firstAuthor="Dellacasa" issue="1" journalOrPublisher="Zootaxa" pagination="23-42" refId="B15" refString="Dellacasa M, Gordon RD, Dellacasa G (2007) Systematic review to the genus Liothorax (Coleoptera: Scarabaeoidea: Scarabaeidae: Aphodiinae). Zootaxa 1407 (1): 23–42. https://doi.org/10.11646/zootaxa.1407.1.4" title="Systematic review to the genus Liothorax (Coleoptera: Scarabaeoidea: Scarabaeidae: Aphodiinae)." volume="1407" year="2007">Dellacasa et al. (2007)</bibRefCitation>
|
||||
, in their revision of the world species of
|
||||
<taxonomicName id="C131E87D87035FFA27A71E19A2E6879A" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="14268C79627AF7183A5836DB7DD244D3" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
, show the same figure of the aedeagus.
|
||||
</paragraph>
|
||||
<paragraph id="04E608122C1DD26BCAD95070FCBABEBC">
|
||||
At present
|
||||
<taxonomicName id="6E311865A92CF1F2C13EAC45A3C1735D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="F8D12E37E4955A5E4D8E81125F139350" italics="true">A. isikdagensis</emphasis>
|
||||
</taxonomicName>
|
||||
is known only from the type series, the
|
||||
<typeStatus id="0B614ADFAE7C90F1825061CE5DA5EB07" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="6404200BFBCAC9E4DDFC7A8558516789" count="7" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
|
||||
, one labelled as
|
||||
<specimenCount id="B27F34FEC773D249842E8BCC7C94A71A" count="1" type="female">♀</specimenCount>
|
||||
<typeStatus id="7AF031E333B61836DC15E3D147E61FEC" type="allotype">allotype</typeStatus>
|
||||
, with the locality data Çamlidere, Isik d., Anat. 23. vi. 47. Exp. N. Mus. ČSR (Fig.
|
||||
<figureCitation id="4134AC00C8BC18038BCB74D5D18034BF" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b, c</figureCitation>
|
||||
). Other Turkish material (Vilayet Rize, Ovitdagi gecidi m 2600, coll. Balerio; Kizildag Gecidi
|
||||
<quantity id="5A7A9893E4EEB61C55B8AEABA5EAC400" metricMagnitude="3" metricUnit="m" metricValue="2.29" unit="m" value="2290.0">2290 m</quantity>
|
||||
, coll. Král; Artvin Çam gec.
|
||||
<quantity id="F95215763A44DA8D778B9DA4EADBADB9" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
|
||||
, coll. Ziani) is shown by its parameres to belong to the
|
||||
<taxonomicName id="E364B0EBE62049FBB79B4889BB446103" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="8EA378BBBFD432A88FD63DA8B6576351" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
species complex, and this is also true of a male from central
|
||||
<collectingCountry id="540EC808FBF74B743FE8B7DEF185131B" name="Armenia">Armenia</collectingCountry>
|
||||
, Selim Pass
|
||||
<quantity id="A5758093EC8399247F3F277A5E6A007B" metricMagnitude="3" metricUnit="m" metricValue="2.35" unit="m" value="2350.0">2350 m</quantity>
|
||||
, coll. Ziani. See discussion of
|
||||
<taxonomicName id="3291983BDBFA3FEC2DF578AD6C3774C7" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
|
||||
<emphasis id="7945C74F6F0711B2280C9E73F9685826" italics="true">L. alberti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="0B07AE4313E22AEFF2F68D98E9C70AE8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
772
data/98/01/C7/9801C79367F45A07A4759B4C8A071C7B.xml
Normal file
772
data/98/01/C7/9801C79367F45A07A4759B4C8A071C7B.xml
Normal file
|
|
@ -0,0 +1,772 @@
|
|||
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<mods:mods id="225F23FB0878B74D8F5B0ED162928D11" xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo id="95F3D2346130ED4A2356446D262CD81C">
|
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<mods:title id="730378DEF26F8312B1246B8B47865F00">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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|
||||
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|
||||
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|
||||
<mods:affiliation id="6FB007088EA5738BA55B045D53402684">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="27F223C486DE2EEA55D9E4B8D6A42F19" type="personal">
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</mods:role>
|
||||
<mods:namePart id="8EEC229DF355058F508AE8EA632601A8">Maté, Jason F.</mods:namePart>
|
||||
<mods:nameIdentifier id="D68D5D1531903947C118E85E73BBB63A" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
|
||||
<mods:affiliation id="92CD57DAA9C89A537956E80AC9899E32">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="5EEBE0B2DD21DF71E3DECD92869A0C46">Angus, Elizabeth M.</mods:namePart>
|
||||
<mods:affiliation id="0AE878F8701E761FFBB9EE4534F912D2">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="CC4585EB3427E6C3D114207EFB7E3808" type="personal">
|
||||
<mods:role id="23C9E8718A7B788BA906391996685279">
|
||||
<mods:roleTerm id="96B68CFD8ACFEE17574852618C72D920">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="8F4C0B777588D78F3DC8C38536EAA345">Král, David</mods:namePart>
|
||||
<mods:nameIdentifier id="3E593CCD96752CEA839827162E8F2347" type="ORCID">0000-0002-7930-1927</mods:nameIdentifier>
|
||||
<mods:affiliation id="E9CA8DF85875597592BFA37D3A47759A">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="F53F40922DB32406A18E54A0B3FEC37A">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="BB0EF940FB62297A298FAA1FD726270D" type="host">
|
||||
<mods:titleInfo id="FD42F497EB25F164B331C9EB93BF7502">
|
||||
<mods:title id="4F76FB3BDF83FF054AB78B144FCD0DE1">ZooKeys</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="5BA9B64D4C23A2F94825691378A81F9D">
|
||||
<mods:date id="4D5E558BDDABBC3BD11AE3F7E49A8E1C">2024</mods:date>
|
||||
<mods:detail id="FFA39097306F0958A0E11E2CCC990FC6" type="pubDate">
|
||||
<mods:number id="C3E2ED56D05D250130212CC59532E948">2024-07-22</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="B88DDD5FDFD3CB4B4AF87B90B4E01A29" type="volume">
|
||||
<mods:number id="AFFAB4DE0E408F59816EDA47C560B9D3">1207</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="991DB573FABDBB3CA37C44B134D3BB34" unit="page">
|
||||
<mods:start id="34E067DFBA12AE2DA7273B881AA74B16">205</mods:start>
|
||||
<mods:end id="A44D479CFAC0AF95EC66906E6BA283D2">299</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="04CE53A52D09D76FCFCBDB4E278E3F00">journal article</mods:classification>
|
||||
<mods:identifier id="E12A700328175EF5E2266BEE22556444" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
|
||||
<mods:identifier id="B716A3F800AD66AFE8A96E93B9993262" type="ZooBank">94F18819-5AF5-4100-AB35-AA3C3976EE80</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="9801C79367F45A07A4759B4C8A071C7B" ID-arpha="9801C793-67F4-5A07-A475-9B4C8A071C7B" LSID="urn:lsid:plazi:treatment:9801C79367F45A07A4759B4C8A071C7B" httpUri="http://treatment.plazi.org/id/9801C79367F45A07A4759B4C8A071C7B">
|
||||
<subSubSection id="A72AE012EF2F8465C061F71F6ED51173" type="nomenclature">
|
||||
<paragraph id="C23258EF463F4AE24252E89F6088A982">
|
||||
<taxonomicName id="526C77F6426E99D7F6A88D4801EC0D62" ID-CoL="FHWT" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="plagiatus" subGenus="Liothorax" subSpecies="plagiatus">
|
||||
<emphasis id="62FD405483DB96E6C567CF290827E923" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="186CBBF2D24D634B5A9170D21A124D4C" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="C9AA8EF954397BE92E1661546284E519" italics="true">plagiatus plagiatus</emphasis>
|
||||
(Linnaeus, 1767)
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="A70C3ECEF08589F9E6C963FF14DA1BDD" type="description">
|
||||
<paragraph id="533DE92FF486CA044AC4837AE4A00CE9">
|
||||
<figureCitation id="212566A040CABB5B2E32EEFE0BD9C78A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">Figs 2 a – c</figureCitation>
|
||||
,
|
||||
<figureCitation id="5813272837698AF8AFCDBE80F24552A1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 a – d</figureCitation>
|
||||
,
|
||||
<figureCitation id="947429E49463BD2A27FB2933B621FD30" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 a – f</figureCitation>
|
||||
,
|
||||
<figureCitation id="1720D02C088243A220486509730E3262" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="13EE5D1B96D0BE638C2B484ACDBDF9F2" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 a – c ”, 18 a, 20 a, b, 22 h – l, 25 i, j, 26 a – g, 30 a, b</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVXEK" type="type material">
|
||||
<paragraph id="6FCF914CCDC99E1B88FC434556BCB6F0">
|
||||
<heading id="F857EE02BC72F65C05FA11AC5C30E30A" reason="title">
|
||||
<typeStatus id="3785D7C5B99FC1523F0FA7A7BDE718F1">Type</typeStatus>
|
||||
material examined.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="2C15DB183DD74D140EE01A204C7073F7">
|
||||
<taxonomicName id="8ADF88D5DA2D93E7D7F9637E0D663EC0" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="A11EBF790CB5F8FA0D2DA0169323670D" italics="true">Scarabaeus plagiatus</emphasis>
|
||||
Linnaeus, 1767
|
||||
</taxonomicName>
|
||||
(image examined on
|
||||
<ext-link id="66E5551D1DB1F36F5E7591DF5069EEAA" ext-link-type="uri" xlink_href="http://linnean-online.org/20869/#?s=0%26cv=0" xlink_type="simple">linnean-online. org / 20869 / #? s = 0 & cv = 0</ext-link>
|
||||
). Designated
|
||||
<typeStatus id="212C8D145FF225B7CEC6D9E80500BFDA" type="lectotype">lectotype</typeStatus>
|
||||
by
|
||||
<bibRefCitation id="7BE82B2EB7E8387814E7775A86BCDB60" author="Krell F-T" firstAuthor="Krell" issue="4" journalOrPublisher="The Linnean" pagination="13-20" refId="B26" refString="Krell F-T (1995) Type catalogue of some Scarabaeoidea species described by Linnaeus. The Linnean 11 (4): 13–20." title="Type catalogue of some Scarabaeoidea species described by Linnaeus." volume="11" year="1995">Krell (1995)</bibRefCitation>
|
||||
(
|
||||
<abbrev id="ABBRID0EPYEK" xlink_title="The Linnean Society, Burlington House">
|
||||
<collectionCode id="0A9BE4EC8C422C728396C7FFDD0C02D0">LSCL</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="A2CBB43043D5F4DE792B5E9860F751B5">
|
||||
<taxonomicName id="1D3046F2FCF980672CD9C7C8522F3A49" authority="Endrodi, 1955" authorityName="Endrodi" authorityYear="1955" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hungaricus">
|
||||
<emphasis id="4A4F352180DA20B2720B6EF4035ECC63" italics="true">Aphodius hungaricus</emphasis>
|
||||
Endrődi, 1955
|
||||
</taxonomicName>
|
||||
.
|
||||
<typeStatus id="7D7CCFFC4A377B53C960B0781138CEC2">Type</typeStatus>
|
||||
series examined (
|
||||
<collectionCode id="22F515C5A07F2BF920CCA27105EC8305" country="Hungary" httpUri="http://grbio.org/cool/9j47-dvjw" name="Museum of Natural Sciences" type="">MNSB</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="952509C61317CB86587F5519E56C0E55">
|
||||
<taxonomicName id="FD40206D5598C95501437BB18FD47D0D" authority="Balthasar, 1938" authorityName="Balthasar" authorityYear="1938" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jakutorum">
|
||||
<emphasis id="6C33C8200B31E2C7B39411F55E614785" italics="true">Aphodius jakutorum</emphasis>
|
||||
<materialsCitation id="2DF81FB19DA9C0BD7F34E23AA2BB6547" collectionCode="NMP" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
Balthasar, 1938.
|
||||
<typeStatus id="D4EE62D92039B51D12B16DF668043B85" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="439203ECA72CBC11842EE26047C86E2B" count="1" type="male">♂</specimenCount>
|
||||
(
|
||||
<named-content id="DE314996A2486176F4209C73275E3819" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="E8FB0936466D0855E62F5E622F57DF73" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</materialsCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERZEK" type="material">
|
||||
<paragraph id="F33BE684DBA0B6F3785F51A1F9BE9ACD">
|
||||
<heading id="B9C2EBA1A5F4E294B36883FBE0CE9AF8" reason="title">Additional material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="156BA312C03AAC42F425479E0CA19008">
|
||||
SP: Aiguamolls de l’Empordà, Urbanización Ampuriabrava,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1ZEK" specific-use="{"type":"Point","coordinates":[3.089000,42.225000]}">
|
||||
<geoCoordinate id="4601F9AA9368DB82B5B563F213930929" degrees="42.225" direction="north" orientation="latitude" precision="55" value="42.225">42.225 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="A012D2E592C966F118D6335EB9C3A848" degrees="3.089" direction="east" orientation="longitude" precision="55" value="3.089">3.089 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. spec 5445,
|
||||
<specimenCount id="F2BA62C4A8A83796C480F65001FF4FB8" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="CF4552A02F3D1C3DEC1137B9904FBB2F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales">
|
||||
<collectionCode id="AABCEC43AF60384B6A5A9732A3DF9BD8" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="547BACED1B7FBA5F6E9080245219FA65">
|
||||
GB:
|
||||
<collectingRegion id="9248DBB64FDB427A2C019AC4FC515DAB" country="United Kingdom" name="Norfolk">Norfolk</collectingRegion>
|
||||
, Hunstanton,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EI1EK" specific-use="{"type":"Point","coordinates":[-0.516800,52.963800]}">
|
||||
<geoCoordinate id="998200635F4BFF26F9A28F82FEB524CC" degrees="52.9638" direction="north" orientation="latitude" precision="5" value="52.9638">52.9638 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="F13216D79AF01019F45D0F9CE102899D" degrees="0.5168" direction="west" orientation="longitude" precision="5" value="-0.5168">0.5168 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
. At edge of lake after flood.
|
||||
<date id="E8E2665BBB7F518FDD9DCA49DE18140C" value="2001-10-27">27. x. 2001</date>
|
||||
, R. B.
|
||||
<collectingRegion id="3A538600F444556DD8925C27DBC4DA50" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="9A5B5DA1C78C83DBFC5E093517B81914" count="6" type="male">6 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="61BC81ECC85E2ECF5BB0C59D0190BB42" count="4" type="female">4 ♀♀</specimenCount>
|
||||
, 5 unsexed (
|
||||
<named-content id="B480CC737AB91A12A21EF25C946B254A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="F75FA283217F1C7D263E14E2B6F12F5F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<collectingRegion id="DD3D7E5086C8C973BE97B039AEFC26F4" country="United Kingdom" name="Dorset">Dorset</collectingRegion>
|
||||
, Studland Heath,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EU1EK" specific-use="{"type":"Point","coordinates":[1.954000,50.455000]}">
|
||||
<geoCoordinate id="F8972D3E9688493AA68EE5BE1648EEC7" degrees="50.455" direction="north" orientation="latitude" precision="55" value="50.455">50.455 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="67ACFB388C8AA6DB8A8E3BA4A47A8C44" degrees="1.954" direction="east" orientation="longitude" precision="55" value="1.954">1.954 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Buried in sand by damp path.
|
||||
<date id="970A621396117387169F51F44C1828EF" value="2002-05-17">17. v. 2002</date>
|
||||
, C. J. Wilson & R. B.
|
||||
<collectingRegion id="52C69979CB2C229EDE57A8CCC823B235" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="E572327056CC7736E49545D4F2B6A08C" count="3" type="male">3 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="6CDD6FD8DDEB774C629D689B09C19897" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="F19A33B275E8B8F3644A4B1EF33AB27B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="B88FD682C49F0E15044602E453935816" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0B13F797B0178AEE8074056FF792400D">
|
||||
LI:
|
||||
<collectingCountry id="26982937658CF1233E396875CCD3F9CA" name="Lithuania">Litauen</collectingCountry>
|
||||
, Nowe Święciany,
|
||||
<date id="63666EB0C9FF3D8168948D9643583ADC" value="1916-06-04" valueMax="1916-06-20" valueMin="1916-06-04">4–20.6. 1916</date>
|
||||
, leg. P. Salchert, 1 unsexed; Livind, Smolwy, Sudl, Dünaburg,
|
||||
<date id="083F77FD30B68C6ADC975595B4B59597" value="1916-06-16">16. 6. 1916</date>
|
||||
, leg. Selchert, 1 unsexed (
|
||||
<abbrev id="ABBRID0E61EK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="78B3BABD41FA19431DAC719B5C2167DA">ABC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="48A4FFE781D840B578B04EFE6AF17BC4">
|
||||
HU:
|
||||
<collectingCountry id="116D447F8E4F186CECB568C3161B34A1" name="Hungary">Hungria</collectingCountry>
|
||||
,
|
||||
<taxonomicName id="66BA8C1FD7D4CC3A65542531117A8E70" authorityName="Dalla Torre" authorityYear="1879" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="variety" species="plagiatus" subGenus="Liothorax" variety="immaculatus">
|
||||
<emphasis id="E874E7582C1DFE8A1B5C1F44DE5B71A9" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="A2C59066B24DA00601248D2F281D793C" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="1036ADFEFC920766DBC6B98AC8166B13" italics="true">plagiatus var immaculatus</emphasis>
|
||||
</taxonomicName>
|
||||
, 201539 (
|
||||
<named-content id="9D0E7820704EA1586A6EC8A356FC143F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales">
|
||||
<collectionCode id="A98F35AF2E9534540E85FCD8BA94EAF6" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="85AE316A9CDAA2EA924DD2380F131B13">
|
||||
CZ: Dobré Pole,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED3EK" specific-use="{"type":"Point","coordinates":[16.533000,48.824000]}">
|
||||
<geoCoordinate id="0871F60070DAF3C4882F0A0F9EA04B2B" degrees="48.824" direction="north" orientation="latitude" precision="55" value="48.824">48.824 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="18B44D43608885380F6D1F990E373796" degrees="16.533" direction="east" orientation="longitude" precision="55" value="16.533">16.533 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<date id="8F08B95A058A7C44A00E9D7AB24AB211" value="2011-04-30">30. iv. 2011</date>
|
||||
, Martinů Ivo.
|
||||
<specimenCount id="D38EBD2C677628736BF3EC57B7612C95" count="1" type="male">1 ♂</specimenCount>
|
||||
, DNA (1) (62) JFM, 4 unsexed (
|
||||
<named-content id="A9322CF1189C7713E081BECCE32912E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="7AC023C5FDE25272AEE76E5D3FAB16F3" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
</paragraph>
|
||||
<paragraph id="3B816A1C99217D97B4DFB66E2B763AE6">
|
||||
SK: E of Chlaba,
|
||||
<date id="32D684531307508E066DDB027B5ADDAF" value="2014-06-11">11. vi. 2014</date>
|
||||
, sandy place S of railway,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ER3EK" specific-use="{"type":"Point","coordinates":[18.846000,47.823000]}">
|
||||
<geoCoordinate id="FD72914ABFEA1BD1818B928A5DA16025" degrees="47.823" direction="north" orientation="latitude" precision="55" value="47.823">47.823 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="95171DA9C7820D4784065A82B5035071" degrees="18.846" direction="east" orientation="longitude" precision="55" value="18.846">18.846 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<quantity id="8CD6519D258ED44A361C8EAC832B4291" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="107.0">107 m</quantity>
|
||||
a. s. l., at light, D. Král lg.
|
||||
<specimenCount id="3BB303705094856750C2A13A20E2BAB5" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EW3EK" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="CDABB5F80FA6305A58E17AE8FE1F35BE">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="CA29279D904649C941916D8F6F5F75C1">
|
||||
RU: Pavlovsk Park,
|
||||
<collectingRegion id="0449BF99B1A2CC6A974076D0EB2556D8" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E63EK" specific-use="{"type":"Point","coordinates":[30.476000,59.694000]}">
|
||||
<geoCoordinate id="358A6CD141CA5D9DAFDCB330F879AA64" degrees="59.694" direction="north" orientation="latitude" precision="55" value="59.694">59.694 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="C3BEDF74334657E31E028D90ED920EEA" degrees="30.476" direction="east" orientation="longitude" precision="55" value="30.476">30.476 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, at edge of snowmelt pool.
|
||||
<date id="4D9BE7FEE982720058B6F8BAEFBC9324" value="1982-05">v. 1982</date>
|
||||
, R. B.
|
||||
<collectingRegion id="5ED082C347D00835CBEC5088FD2B69DA" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="0631D8404886857E966096D081D73846" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<named-content id="FA10FCF8E583D17429568FBED31BBD6D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="0D340A94F87BED7E8E518C4544C40A9B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<collectingRegion id="A2EF7EE803E71AB8AFBD94337CBFC3A7" country="Russia" name="Yaroslavl'">Yaroslavl</collectingRegion>
|
||||
, Berditsino,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EL4EK" specific-use="{"type":"Point","coordinates":[40.110800,57.454000]}">
|
||||
<geoCoordinate id="8D46A12DBD6A38EB67DCC13864C060EA" degrees="57.454" direction="north" orientation="latitude" precision="55" value="57.454">57.454 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="E7C0E56FC422D4D5185DD1E2516FF4F5" degrees="40.1108" direction="east" orientation="longitude" precision="5" value="40.1108">40.1108 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. leg. Yakovlev.
|
||||
<specimenCount id="66E6FB7253CA277F8699BFA036B14D31" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<named-content id="02C5A6E3BBAA61A88BD236C41368DC1F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="A5D6D51405B510C4451B187B4BB62818" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<collectingRegion id="6703F44C9DAFDF40D833D49472E5D11C" country="Russia" name="Novosibirsk">Novosibirsk</collectingRegion>
|
||||
oblast’, Sorochicko env.
|
||||
<specimenCount id="FB20F8306EB939D2F8644072B0AC9082" count="116" type="generic">
|
||||
<quantity id="11286A7A274821206AFCED72CD045FF4" metricMagnitude="2" metricUnit="m" metricValue="1.16" unit="m" value="116.0">116 m</quantity>
|
||||
.
|
||||
</specimenCount>
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX4EK" specific-use="{"type":"Point","coordinates":[77.887000,53.285000]}">
|
||||
<geoCoordinate id="2C2F46560496F3D4D4476F875ABC10B5" degrees="53.285" direction="north" orientation="latitude" precision="55" value="53.285">53.285 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="C00037751E237D30DD5CB011064211CA" degrees="77.887" direction="east" orientation="longitude" precision="55" value="77.887">77.887 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Waterside litter.
|
||||
<date id="F20CDF50431035F0712AE2D638868336" value="2012-05-09">9. v. 2012</date>
|
||||
. D. J. Mann & J. Cooter.
|
||||
<specimenCount id="48D75D4E6FFE266A855D2B690C362CFE" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="45955B96A0B1E0753240EA783881D877" count="2" type="female">2 ♀♀</specimenCount>
|
||||
(
|
||||
<named-content id="4523A5862C5BCBFD7D993D555B2EF93A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oxford-university-museum-natural-history" xlink_title="Oxford University Museum">
|
||||
<collectionCode id="64B1A4AD2A1856CAF9CE5557883E27E7" country="United Kingdom" httpUri="http://grbio.org/cool/g6yf-wft0" name="Oxford University Museum of Natural History" type="">OUM</collectionCode>
|
||||
</named-content>
|
||||
); Tuapse, Chernomorsk oblast’,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED5EK" specific-use="{"type":"Point","coordinates":[39.090000,44.098000]}">
|
||||
<geoCoordinate id="8B2E5B84D6A2ABEC9DACCF47B570BBC6" degrees="44.098" direction="north" orientation="latitude" precision="55" value="44.098">44.098 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="E9D0E148C8E0F68C46964A11EEB47880" degrees="39.090" direction="east" orientation="longitude" precision="55" value="39.09">39.090 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. 1, unsexed (
|
||||
<named-content id="287FF2626819C40C9B14B101E4A9B748" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="EFBD348E3B6901CD2E9583357F61F7D9" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
); Minusinsk distr. Shushenskoe,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP5EK" specific-use="{"type":"Point","coordinates":[91.946000,53.324000]}">
|
||||
<geoCoordinate id="1BB581F0E2643F8C4897147DEA1279ED" degrees="53.324" direction="north" orientation="latitude" precision="55" value="53.324">53.324 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="BEA8A8E8683FD333BEF094F3BBBF5C46" degrees="91.946" direction="east" orientation="longitude" precision="55" value="91.946">91.946 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. 1, unsexed (
|
||||
<named-content id="7E066B7C5CE27909705B7203DA462BC1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="DB52B5CC5D166CB6C9DC70869FA205BF" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
); Chuyskaya steppe, Kosh Agach,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E25EK" specific-use="{"type":"Point","coordinates":[88.657000,49.991000]}">
|
||||
<geoCoordinate id="1143BBEB42BF91FA61E0477ED2DCD938" degrees="49.991" direction="north" orientation="latitude" precision="55" value="49.991">49.991 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="033CC7D29823DEA11EDEC16C4D14D1C8" degrees="88.657" direction="east" orientation="longitude" precision="55" value="88.657">88.657 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="528586390A29355965E4B27CE31AD8F4" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="E92E340DF17EE044FEE6AC133344EE21" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="0C70E526332B240B587E38461F191DE2" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
); Bunbui, Kansk neighbourhood, Yenisei government,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH6EK" specific-use="{"type":"Point","coordinates":[99.025000,56.382000]}">
|
||||
<geoCoordinate id="6271C2DDE13214B931E5334ECE984A35" degrees="56.382" direction="north" orientation="latitude" precision="55" value="56.382">56.382 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="FAACB973A7FD0A95CB717C1043F929F8" degrees="99.025" direction="east" orientation="longitude" precision="55" value="99.025">99.025 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, 1 unsexed, black (
|
||||
<named-content id="4A557B6B14E0309CB8398376832F00CA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="137370BA774EC7ABFBDBED2B3BE5221C" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
); Transbaikal, Berezovka near Ulan Ude,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET6EK" specific-use="{"type":"Point","coordinates":[107.240000,51.932000]}">
|
||||
<geoCoordinate id="64FBDF256189A9FD6F0A4664E131A3F2" degrees="51.9320" direction="north" orientation="latitude" precision="5" value="51.932">51.9320 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D9CB6C489FBA631FE0669DA5DCD40DB4" degrees="107.2400" direction="east" orientation="longitude" precision="5" value="107.24">107.2400 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. 1, unsexed (
|
||||
<abbrev id="ABBRID0EY6EK" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="C11F16A0078242438B4CB885A529D262">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="6705E93386435DF929519F7F1D65862C">
|
||||
TR: Bozkale near
|
||||
<collectingRegion id="8663DBC1E58141A1E5D7A0CDDD3EC24A" country="Turkey" name="Kars">Kars</collectingRegion>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBAFK" specific-use="{"type":"Point","coordinates":[42.980000,40.587000]}">
|
||||
<geoCoordinate id="8BFBFFAA91FDFE757601463881B887A1" degrees="40.587" direction="north" orientation="latitude" precision="55" value="40.587">40.587 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="125D795BBF0367D30DAC4476D4573DB4" degrees="42.980" direction="east" orientation="longitude" precision="55" value="42.98">42.980 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
(
|
||||
<named-content id="B8D26ABE75809636DE77EC7DAAD702F0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="889ACE3300D0692F26CEC080773686ED" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="07FD5E42DCB4D6490A9E323C651E8379">
|
||||
IN: Shiraz, Lake Maharlou,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPAFK" specific-use="{"type":"Point","coordinates":[52.759000,29.450000]}">
|
||||
<geoCoordinate id="41783077078F942057CA9CED104F7477" degrees="29.450" direction="north" orientation="latitude" precision="55" value="29.45">29.450 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="636BA22FBA41BCB5A15AF6D99349EBF6" degrees="52.759" direction="east" orientation="longitude" precision="55" value="52.759">52.759 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<specimenCount id="3B5A9C7766D44CDA0DB3C9072538F5BB" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<collectionCode id="3EF11BBDBC366CCE166C32515CCEC516" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" type="">MNHP</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="D7AB0EBA9930CD20AB26E072BDDEDBAF">
|
||||
KZ: Yanvartsevo, right bank of the Ural River,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZAFK" specific-use="{"type":"Point","coordinates":[52.239000,51.424000]}">
|
||||
<geoCoordinate id="D27A8964E4F279002E8CFE4CB80E6502" degrees="51.424" direction="north" orientation="latitude" precision="55" value="51.424">51.424 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="6CF3EFF4DA719E525A1B33B66DFD5F0F" degrees="52.239" direction="east" orientation="longitude" precision="55" value="52.239">52.239 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, leg.
|
||||
<tp_taxon-name id="02AE0C2FDA49E6FD0017ED215EE52D30">
|
||||
<emphasis id="02F3EAEDE8FC65649019E0FA3F953DF8" italics="true">
|
||||
<tp_taxon-name-part id="62DEB3F0B0BFD97C28EEC7A0CBD53CA3" reg="Liothorax" taxon-name-part-type="genus">L.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Arnoldi, 1 unsexed, black (
|
||||
<named-content id="46F1652D96FA3BB397A051B377559D28" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="72DBE53D4F4A29695A1AAB60D9A85AB0" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
); Koksengir S of Zhana-arka, Karaganda.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELBFK" specific-use="{"type":"Point","coordinates":[79.440000,49.440000]}">
|
||||
<geoCoordinate id="A8BA259E7539FAFBFC0FA65CE369B38F" degrees="49.44" direction="north" orientation="latitude" precision="555" value="49.44">49.44 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="903B0229AEAFC6512901D133EF923566" degrees="79.44" direction="east" orientation="longitude" precision="555" value="79.44">79.44 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. leg. Logonova,
|
||||
<date id="5E11B041FE6311FAD707DE5AEB1A8F9B" value="1958-05-22">22. v. 1958</date>
|
||||
.
|
||||
<specimenCount id="6F39454CB2C9D47383DB6832C02A3B68" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="5E8F1F0935919DAE611ABBB22CB60E93" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="769586F17C29D0AE88BC7A5090F3F895" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="47D48AE7E316D44C09F2C973405CF9D9">
|
||||
MG: 40 KM Bayanbulag
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZBFK" specific-use="{"type":"Point","coordinates":[98.583000,45.083000]}">
|
||||
<geoCoordinate id="8322CDEADEF27FF0687CB4EDD7995A21" degrees="45.083" direction="north" orientation="latitude" precision="55" value="45.083">45.083 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="191B0156A524BB040CD03E8524B91001" degrees="98.583" direction="east" orientation="longitude" precision="55" value="98.583">98.583 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="27B8D7CEAA4ABBB325F50E02C72F4C76" value="2005-07-16">16. vii. 2005</date>
|
||||
, leg. A, Mikyška.
|
||||
<specimenCount id="922A34A478857AFD5AF2DB3C84D83B8C" count="1" type="male">1 ♂</specimenCount>
|
||||
, elytra with red flashes (
|
||||
<named-content id="992B1023BD5A144672848A5284F7E4FA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="90FB151D5C34C3D7382ADF1D04AB5F02" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<collectingCountry id="601AF1223462354D27A4C0A4A38E006E" name="Mongolia">Mongolia</collectingCountry>
|
||||
west, 50 KM SW Uliastay. 16. vii, 2005, leg.
|
||||
<tp_taxon-name id="3DFFF48DD678D33B5419C56A55F70AF3">
|
||||
<emphasis id="E24B36D3260D971C84B13D18269BA6B3" italics="true">
|
||||
<tp_taxon-name-part id="E606C6FB7720455A3CC3ACADF9E75EC5" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Mikyška,
|
||||
<specimenCount id="42DB241C9037F00E52CB6786A4429848" count="1" type="male">1 ♂</specimenCount>
|
||||
, elytra with red flashes (
|
||||
<named-content id="42E8AD845DE66F362190E4D8C8648A86" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="9E9A9DBA18D1F4613B474E20F2F03497" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
); Right bank of R. Kerulen (Herlen) near the Prikhity mountains. 1 unsexed, black (
|
||||
<named-content id="474FE20433BC624DDC2C7B0E6FF4B9A7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="5BB409AD095255EBAED7385A6062D9FD" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="820670F97E9109FA568A9B40FD73412B">
|
||||
<collectingCountry id="3CC582485DD631DB61B458D2A2AFA612" name="Switzerland">CH</collectingCountry>
|
||||
: “ Pekin ”, Fry coll.
|
||||
<specimenCount id="2192E104EA7E80BE61CAE1D0C28F2B62" count="2" type="male">2 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="50B94B655CA1FCCCBF8908C8558EA013" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="0DB33992E8CF73F40296E904DE248953" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="AAAF5E4B4E2D48978947EA56A499F9F4" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWCFK" type="Differential diagnosis">
|
||||
<paragraph id="76402A1C9A43659BDF5A1D639F92BA3B">
|
||||
<heading id="8FF060F70B0D729D3F1BE837ADAF4D91" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="96174F63F7AE369E775CBAFA5BBF46B8">
|
||||
<taxonomicName id="08F6E75E91B8CFCCFE5C7B098EC306B7" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="6E78D3EEF82F5E2C35C496104FD1123D" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
is recognisable by the basal segment of the mesotarsi being shorter than the longer mesotibial spur, and the heavily punctured metaventrite, especially in males. The aedeagus is distinctive, with the parameres not decurved apically and the endophallus lacking recurved teeth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EHDFK" type="redescription">
|
||||
<paragraph id="10FD4C9E8D09922979D11BE3E2F61DEC">
|
||||
<heading id="C0AAD5CDD31BDB36E3618276A53217C8" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="889E6C2DB21AA135E77E31B32FB967D4">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="B88EDDE617CC292B9D1AEAD1D9679D86" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 a – c</figureCitation>
|
||||
). Length:
|
||||
<quantity id="48345F3E0C95AADA8C74DB69F8E9BAE9" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.9" metricValueMin="3.6" unit="mm" value="4.25" valueMax="4.9" valueMin="3.6">3.6–4.9 mm</quantity>
|
||||
(
|
||||
<specimenCount id="FF185457F0869A7278061ADC668C32DC" count="1" type="male">♂</specimenCount>
|
||||
), 4.0–
|
||||
<quantity id="F0F6A316876B0C15E028770B94E9E9D9" metricMagnitude="-3" metricUnit="m" metricValue="4.8" unit="mm" value="4.8">4.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="B0FB5A5C1BFDFF26C2D8154A6D24EE34" count="1" type="female">♀</specimenCount>
|
||||
); width: 1.6–2.0 mm (
|
||||
<specimenCount id="6CE9CE71668E8AE7CDD138B4F981F85C" count="1" type="male">♂</specimenCount>
|
||||
), 1.8–2.0 mm (
|
||||
<specimenCount id="CED26F390D99DE117D7D00A80176B5FC" count="1" type="female">♀</specimenCount>
|
||||
). Black, head, pronotum, and scutellum glossy (pronotum sometimes less so) (Fig.
|
||||
<figureCitation id="1E4CBEA32471755573E877C2B09392D5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 a – d</figureCitation>
|
||||
), elytra black, sometimes with a dull red streak over the middle part (Fig.
|
||||
<figureCitation id="9D33D2CF2F4B0658CB69D4F5A640F469" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 b</figureCitation>
|
||||
), interstices with very fine reticulation often giving a silky grey sheen.
|
||||
</paragraph>
|
||||
<paragraph id="142D01D3FC8093F8A5FE8AC6E40B8BC0">
|
||||
Head glossy black with a diffuse dull brown area along the outer margins of the clypeus and genae, Frons with moderately strong punctation, the punctures separated by ca 1.5 × their diameters, and with some sparse very fine punctures between them. Frontoclypeal suture often very fine, straight over median 2 / 3, then angled obliquely forward at each side, reaching the margin at the front of the genae. Occasionally the suture is more distinct. Clypeus bulging upwards medially, this bulge with fine sparse punctation which becomes coarser and rather rugose anterolaterally. Most European material has this rugose area fairly extensive and well-developed, but a male from
|
||||
<collectingRegion id="4733959A11BBD1C639DAC765D568A57D" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
(
|
||||
<collectingCountry id="C681A8299D4A784E74A23760F9CD95E7" name="Russia">Russia</collectingCountry>
|
||||
) has the rugosity weaker and less extensive, and in material from the Tibetan Plateau (Qinghai and Gansu) the rugosity is further reduced (Fig.
|
||||
<figureCitation id="D7DD36B6725697F2EF19DA9716BBB0B5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 a – d</figureCitation>
|
||||
). The genae protrude laterally at an angle of ca 45 ˚ in front of eyes, then curve inwards to meet the frontoclypeal suture ca 1.5 × eye-diameter in front of eyes. Clypeal margin obtusely angled outward from this point, then curved inwards to the median excision. Genae laterally with a narrow raised margin, this continued across the clypeus, becoming weaker medially. Antennae and maxillary palpi black to blackish brown. Epipharynx (Fig.
|
||||
<figureCitation id="97186EB2C3FD1F2F235D018E7463542D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 a – f</figureCitation>
|
||||
) with the clithra generally only weakly sinuate either side of the tylus, but more strongly so in one Chinese specimen (Fig.
|
||||
<figureCitation id="6F0E80EB5AE1187FF69529C046F7402C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 f</figureCitation>
|
||||
). Chaetopedia 2–6.
|
||||
</paragraph>
|
||||
<paragraph id="A936DB186AA674D01FFCF182E24B5263">
|
||||
Pronotum glossy black, hemicylindrical, highly arched transversely, weakly so longitudinally. Lateral area of surface bulging outwards, almost obscuring the lateral margins (viewed from above) in basal 1 / 2. Surface with double punctation, the larger punctures heavy, twice the diameter of those on the head, separated from one another by 1–4 × their diameter, and with very fine dot-like punctures scattered among them. The larger punctures are very sparse on the disc, denser laterally. Basal margin of pronotum varying from being entirely bordered to having the middle 1 / 2 unbordered (Fig.
|
||||
<figureCitation id="695993E3333671A6D1AE3DE4333C7BD0" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 a, b</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="AB0761720C600558B7BBEE9E902FFD18">Scutellum pentagonal, elongate, the sides more or less parallel in basal 1 / 2, then convergent apically to rounded apex. Surface glossy black with a few very fine punctures medially</paragraph>
|
||||
<paragraph id="E7060000CB2F2C86174BDF0A798786CD">
|
||||
Elytra black, sometimes with an oblique reddish streak over their middle parts. Interstices generally with a greyish silky sheen, contrasting with the glossy striae, but sometimes glossy black, as in the
|
||||
<typeStatus id="6AA668332F9C61AD3C56133815198DA7">type</typeStatus>
|
||||
material of
|
||||
<taxonomicName id="06A9A8FB912D7B98EBFBA70312C1EEB1" authority="(Endrodi, 1995) (MNSB)" baseAuthorityName="Endrodi" baseAuthorityYear="1995" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hungaricus">
|
||||
<emphasis id="BFE6564B4DB19864D5AFA784462A938F" italics="true">L. hungaricus</emphasis>
|
||||
(Endrődi, 1995) (MNSB)
|
||||
</taxonomicName>
|
||||
, the
|
||||
<collectingRegion id="BFA2C9056DD167DA1DD4ACA087F4430B" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
specimen already mentioned, and in material from the Tibetan Plateau. The interstices have a fine isodiametric reticulation, easily seen in most European and Mongolian material, but less so in the
|
||||
<collectingRegion id="C2C225CC0CC2DF6A66C5083E4ADBBF07" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
specimen, and almost invisible in the Tibetan material where the elytra appear very highly polished. Nevertheless, if the elytra are viewed with strong light and at high magnification the reticulation is always present (Fig.
|
||||
<figureCitation id="BE34E7CAE7C224DEE948C0A8B36D649C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 a – c</figureCitation>
|
||||
). Scanning electron microscopy (Fig.
|
||||
<figureCitation id="07985D98E13738BE5966432B7539F0E5" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 a</figureCitation>
|
||||
”, c ”) confirms that this reticulation is on the elytral surface rather than being buried under the epicuticle. In most specimens the fine punctures of the interstices are very faint, though with some variation, but in the Tibetan material they are considerably stronger (Fig.
|
||||
<figureCitation id="C71C5F0E2ADFA3B8681BA13163631DC3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 a, a</figureCitation>
|
||||
’, a ”, c, c’, c ”). Interstices flat, ca 8–10 × wider than the striae. Striae narrow, vertical-sided, their bottoms glossy. Strial punctures in single rows, small, separated by ca 5 × their diameter, and scarcely indenting the strial margins. Lateral margins of elytra gently rounded and slightly divergent over their basal 1 / 2, then tapered to the bluntly rounded apex.
|
||||
</paragraph>
|
||||
<paragraph id="A991977F52FEDE1D8A0591774721FE8C">
|
||||
Metaventrite median diamond-shaped plate in males flat and heavily punctate, the punctures bearing distinct yellowish setae (Fig.
|
||||
<figureCitation id="29C83EB442E512DC64807B7B86394F7E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 a</figureCitation>
|
||||
) which may be lost, especially if an attempt is made to clean dead specimens. In females the median plate is less flat, generally depressed medially and the punctation is finer, ca 2 / 3 the strength of that in males, and the setae are more frequently lost, even in living material (Fig.
|
||||
<figureCitation id="16CB56E252F0040072B2B18FB5BE0E0C" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 b</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="34A85B4C154CDFD5ABD58F37E390FD1C">
|
||||
Legs dark brown to black, tarsi dark brown, and tibial spurs a slightly paler brown. Basal segment of mid tarsi clearly shorter than the longer tibial spur (Fig.
|
||||
<figureCitation id="4A004AD0B90823672BD380CCC46D00DB" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 a</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="F82675C496A7D3CE648BBFCFEE7E8AA9">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="82210F90C511480D33B65E432414A0E5" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 a – c, h – k</figureCitation>
|
||||
) ca
|
||||
<quantity id="EC3BA52D719F08E05DA13929ACB4D020" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
long in
|
||||
<emphasis id="D8769F252D07225368418B8649B41A02" italics="true">
|
||||
A. p.
|
||||
<taxonomicName id="96FF7860A3264FBEADA4EC5691963834" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">plagiatus</taxonomicName>
|
||||
</emphasis>
|
||||
,
|
||||
<quantity id="AE6F70CC8136CA6665744FF8688C990B" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" unit="mm" value="1.325" valueMax="1.4" valueMin="1.25">1.25–1.4 mm</quantity>
|
||||
in
|
||||
<emphasis id="7B58B74E830B03574A83B32733E059B6" italics="true">A. p. sinoplagiatus</emphasis>
|
||||
ssp. nov., the parameres widened apically and the endophallus bearing fields of small scales but no hairs or bristles. The expanded apical area of the parameres is more abruptly widened in Chinese material, giving a square-ended appearance (Fig.
|
||||
<figureCitation id="C5034DE5E2D37C224B3D5361C8ECE46B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 i, j, k</figureCitation>
|
||||
), than in European where it is more oblique (Fig.
|
||||
<figureCitation id="43DCFE594EBB716029780DD33CD7384D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 h</figureCitation>
|
||||
) and the
|
||||
<typeStatus id="DA9A50FD3FB410D6ACEA82F996CFB05F" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="3364CC538175BDAEA134B891AD321BB0" authority="(Balthasar, 1938)" baseAuthorityName="Balthasar" baseAuthorityYear="1938" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jakutorum">
|
||||
<emphasis id="B081233902BC984DB873F7FA5512475C" italics="true">A. jakutorum</emphasis>
|
||||
(Balthasar, 1938)
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="1E9D67DC0D7E9ED67681355F72520276" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 l</figureCitation>
|
||||
) appears to match European material in this character. However, some caution is needed as this part of the parameres tends to shrivel when dried and exposed and often does not fully expand when wetted out. Note that the aedeagus of the
|
||||
<collectingRegion id="E2CD05AE2A92978434ECE39371B0F5B3" country="China" name="Beijing">Peking</collectingRegion>
|
||||
specimens shown in Fig.
|
||||
<figureCitation id="86333BD6BAE8628146BE41CE83799891" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 i</figureCitation>
|
||||
is ca
|
||||
<quantity id="1AE046523C8E68514F4CC9AD1880F323" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
long, matching
|
||||
<emphasis id="2244EB26ED0CC440ECE67402097048BA" italics="true">
|
||||
A. p.
|
||||
<taxonomicName id="FC4E436B2D12AF967002F6E2B70B64BC" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">plagiatus</taxonomicName>
|
||||
</emphasis>
|
||||
rather than
|
||||
<emphasis id="58B14F13D95807A4FE505E1A8EF19A65" italics="true">A. p. sinoplagiatus</emphasis>
|
||||
ssp. nov.
|
||||
</paragraph>
|
||||
<paragraph id="08FF14271A922D36DEFC866300541ECA">
|
||||
Spermatheca (Fig.
|
||||
<figureCitation id="F9E53FE8BADECDB2432AA599112BE252" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 a – g</figureCitation>
|
||||
) ca
|
||||
<quantity id="C45AC8FE105F1766E5A4D4EC41F7A05D" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" unit="mm" value="0.36">0.36 mm</quantity>
|
||||
long in
|
||||
<emphasis id="14009A7D72A460C6E0C7F6FF8CCA95DA" italics="true">
|
||||
A. p.
|
||||
<taxonomicName id="87D51D70B016290E16D357FD46D4F608" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">plagiatus</taxonomicName>
|
||||
</emphasis>
|
||||
, but clearly longer in
|
||||
<emphasis id="F3B7804C6DED2A3E43CE967B64FD2910" italics="true">A. p. sinoplagiatus</emphasis>
|
||||
ssp. nov., length ca
|
||||
<quantity id="C1A4B0CB6D69F27B17C9CA1BF06BC3FA" metricMagnitude="-4" metricUnit="m" metricValue="4.2" unit="mm" value="0.42">0.42 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EYJFK" type="remarks">
|
||||
<paragraph id="DF05687F014E43CB8D99FA1668ACEC2B">
|
||||
<heading id="994EA350BA45EEC40183FA74F1D24232" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="642CC249FBD238250DB2BD8514670133">
|
||||
<taxonomicName id="967F2310E78B3E39948ECE5D5C294DD9" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="0805AF8531F29095DA0F4CA6DF8E088E" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
is the most widely distributed of all the Palaearctic
|
||||
<taxonomicName id="9ADFC004751EE07597E862E8264F79FA" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9C50A046FEB768251FE9809966598595" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
, with its range extending from
|
||||
<collectingCountry id="197F66B1CDF1EBBE64172B8C6F04C58E" name="United Kingdom">England</collectingCountry>
|
||||
in the west to east Siberia (Yakutia) and
|
||||
<collectingCountry id="F23B1A21356C93E58C2DD09A6E4D6372" name="China">China</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="7E5B37521B0E4650E83403AE2A2846FA" country="China" name="Qinghai">Qinghai</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="C0784566CCC51ED70ECBFD8C1107A5CB" country="China" name="Gansu">Gansu</collectingRegion>
|
||||
, and the
|
||||
<collectingRegion id="701E31DF614C4812D6485FD3680D3AA2" country="China" name="Beijing">Beijing</collectingRegion>
|
||||
area) in the east (Fig.
|
||||
<figureCitation id="C25F87E325AD3348DFF0AF95D82D889A" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b, c</figureCitation>
|
||||
). The southern limits of its range are unclear. There are no confirmed records from the Iberian Peninsula and the Catalogue of Palaearctic
|
||||
<taxonomicName id="3EFADC8785145AEDA77044993FBEAC62" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||||
gives no records from
|
||||
<collectingCountry id="045AF5A16CB42A94E74EB51E9EF5FB2B" name="Italy">Italy</collectingCountry>
|
||||
. Records from the Balkans require confirmation because of confusion with
|
||||
<taxonomicName id="81849B93E03717B30BF2C124AF8731C4" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="4B5291FC67147EED3F590576A137ABCC" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
. All material from
|
||||
<collectingCountry id="BF78B1471E7577E74B418B94622934A1" name="Syria">Syria</collectingCountry>
|
||||
examined in this study has been determined to be
|
||||
<taxonomicName id="545DC35E49D4B9A0B458669C4BB6A3C5" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="discoides">
|
||||
<emphasis id="70402D12D1791D5556CFF6B1B24F5E76" italics="true">L. discoides</emphasis>
|
||||
</taxonomicName>
|
||||
and not
|
||||
<taxonomicName id="C491BBB1ADDFF1F6B31D3CF32C6866B1" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="F1D40CC38813A2E87053C46F85CB7071" italics="true">L. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
as reported by
|
||||
<bibRefCitation id="6B8A6CCBC65349B5BE8DBA51E6F20ECC" DOI="10.3406/linly.1992.11007" author="Baraud J" firstAuthor="Baraud" journalOrPublisher="Faune de France, France et Régions Limitrophes. Société Linnéenne de Lyon 78" refId="B6" refString="Baraud J (1992) Coléoptères Scarabaeoidea d’Europe. Faune de France, France et Régions Limitrophes. Société Linnéenne de Lyon 78, 856 pp. [11 pls] https://doi.org/10.3406/linly.1992.11007" title="Coléoptères Scarabaeoidea d’Europe." year="1992">Baraud (1992)</bibRefCitation>
|
||||
. The only (doubtful) record of
|
||||
<taxonomicName id="1AA87E14C09F8839121AB3D560BC8266" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="9255D98BAD9ADC8C6AC4F9911DB6EBC8" italics="true">L. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
from the Middle East is a female from
|
||||
<collectingCountry id="B04E42E7D52355CE9A59D8F24439A722" name="Iran">Iran</collectingCountry>
|
||||
(Shiraz) found in the collections of the Paris Museum by one of the authors (
|
||||
<collectionCode id="B6F2F141F865712163C116EBBA79F52D">JFM</collectionCode>
|
||||
) which was collected during the 1965 Mission Franco-Iranienne and was identified by Baraud as
|
||||
<taxonomicName id="8A569076D77CF89CD3FB636A9C6C62BF" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="DDC77CDB876D38A14E627CE15A94EF30" italics="true">L. niger</emphasis>
|
||||
</taxonomicName>
|
||||
. This specimen was not mentioned by him in his Coléoptères Scarabaeoidea d´Europe (1992) nor, to our knowledge, by any other author, and, apart from
|
||||
<taxonomicName id="C60F9A045C14266CBB8E729794F0149E" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kraatzi">
|
||||
<emphasis id="5BF9FEE7BF6E40AD02D53CD633A33127" italics="true">A. kraatzi</emphasis>
|
||||
</taxonomicName>
|
||||
from Ashooradeh Island is the only specimen of
|
||||
<taxonomicName id="13205669C13F096AE22ED1F380DE13FB" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="35109E26AF31055065BF38FEBC1F9F7B" italics="true">Liothorax</emphasis>
|
||||
</taxonomicName>
|
||||
known for
|
||||
<collectingCountry id="6ED5AEDBD49CB90822D860464C1C8EC5" name="Iran">Iran</collectingCountry>
|
||||
. This specimen was examined in 2004 and tentatively identified as a form of
|
||||
<taxonomicName id="9F73DC8E2F0851DE897C743DCA291D9F" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae">
|
||||
<emphasis id="9BDFE21B43197E0621C197D51F1B68B2" italics="true">wilsonae</emphasis>
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName id="2D5D9F425E42364A1F7B06B554C934E2" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="B63715A8F92789EC9CD35092F9283EBA" italics="true">plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
based on a strongly punctured metaventrite, shortened tarsi, and a surface finish notably different from
|
||||
<taxonomicName id="D26175DA2554862034272DA9ED7F27F5" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="502ED64D4DF81C52A20E200E55853788" italics="true">niger</emphasis>
|
||||
</taxonomicName>
|
||||
, but due to its divergent morphology was not included in the
|
||||
<typeStatus id="10660C73A3B5DE9B710DBC76AF9FBDAD">type</typeStatus>
|
||||
series. In light of the work done since, we suspect that it is a member of the
|
||||
<taxonomicName id="6A2A08F573186309705F908BFED1B0E6" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="8042FBA6A811FBCAA764A6EDFF954828" italics="true">plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
group, but requiring additional material, in particular males. As such this record has been labelled on the distribution maps (Fig.
|
||||
<figureCitation id="37C9C4CE3E0260871E4F63A9964FF48D" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 c</figureCitation>
|
||||
) with a black question mark and a black square.
|
||||
</paragraph>
|
||||
<paragraph id="E46384766D3114CEFF2FD5DD476FE2F9">
|
||||
One of the peculiarities of
|
||||
<taxonomicName id="DA3F297B85D972ADD2ED9D3DFE5722D3" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="D95264389BE7D8A7838D9144983C8B37" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
is the existence of two distinct colour forms, one plain black and the other, exemplified by the
|
||||
<typeStatus id="7CB8933687E790CDA0F19E4E9D26360E" type="lectotype">lectotype</typeStatus>
|
||||
presumably from
|
||||
<collectingCountry id="209B741C986992C0240F03A056A2414C" name="Sweden">Sweden</collectingCountry>
|
||||
, with a red longitudinal-oblique flash over the central area of each elytron. One of us (
|
||||
<collectionCode id="554C26E6024F9B3E17107C5A4A673666">RBA</collectionCode>
|
||||
) has noted the red-streaked form in
|
||||
<collectingCountry id="DEF095BDA37E0906A5C6864E184E4FA3" name="United Kingdom">England</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="6F0886064E5E37CA2C4560D0DF6E625F" name="Ukraine">Ukraine</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="A1823C11DBDB46B19DA844D65FA2A080" country="Ukraine" name="Kyiv">Kiev</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="38217D656B436101D8AC49631CC75562" country="Ukraine" name="Odesa">Odessa</collectingRegion>
|
||||
), and
|
||||
<collectingCountry id="CC06AEF9DF361807E837266CF777994F" name="Mongolia">Mongolia</collectingCountry>
|
||||
. The red form appears to be absent from the
|
||||
<collectingCountry id="7E133B2FAF4B9C7E67A503CE81C1E1BD" name="Czech Republic">Czech Republic</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="CDE14E070234BA1EE495E9B1E37EFFCE" name="Hungary">Hungary</collectingCountry>
|
||||
as well as from
|
||||
<collectingCountry id="1A60BF204998E772F44FC9268A06FCE2" name="Russia">Russia</collectingCountry>
|
||||
. As far as is known, this red-streaked form does not occur in
|
||||
<taxonomicName id="1DEFB87370D2926BCA2572044F59FE61" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="plagiatus" subSpecies="sinoplagiatus">
|
||||
<emphasis id="8B1999AA6D6A8820A5774A09D8A7C2DC" italics="true">A. plagiatus sinoplagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="A753F8B8F470CAF61CF6E90743EEEEC4">
|
||||
<taxonomicName id="5FA399638C5276764E49C79B3F7A877B" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="A1D1F13A506A480522297802A4524493" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
is normally found at the roots of vegetation in damp places and is often found washed up among strandline detritus at the edge of pools (Fig.
|
||||
<figureCitation id="AD3A43427E0AFD61DAE779FE2F0E72A9" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 a, c</figureCitation>
|
||||
). It is generally associated with salinity, as in the
|
||||
<collectingRegion id="2FBABD7F1983E15C2451B763EE71BF44" country="China" name="Qinghai">Qinghai</collectingRegion>
|
||||
pool shown in Fig.
|
||||
<figureCitation id="591851F51A1A08B93EA1D1BC37CBB937" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 c</figureCitation>
|
||||
, but there did not appear to be any trace of salinity in Pavlovsk Park near
|
||||
<collectingRegion id="41ADB38D3F05F2D0BFE532CDEB91431B" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
where RBA collected
|
||||
<specimenCount id="FA75AAA1E56B08D88A0D1DDD97539D33" count="1" type="generic">one specimen</specimenCount>
|
||||
at the edge of a snowmelt pool in
|
||||
<date id="3061CB7C0C7B721EC30B3A4D25EA4B93" value="1982-04">April 1982</date>
|
||||
(Fig.
|
||||
<figureCitation id="A8E53BB166D26E2E1B4F413EDAF9D8E8" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 a</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
281
data/A3/AA/B4/A3AAB4F7DFCF5000804CA4F8B4FB44E5.xml
Normal file
281
data/A3/AA/B4/A3AAB4F7DFCF5000804CA4F8B4FB44E5.xml
Normal file
|
|
@ -0,0 +1,281 @@
|
|||
<document id="06A0BE83DE374B4B6630C5E44BD778C2" ID-DOI="10.3897/zookeys.1207.117225" ID-ZooBank="94F18819-5AF5-4100-AB35-AA3C3976EE80" ID-publisher-id="117225" URI-arpha="255F6420-5705-5F50-9167-509E6C76FA1C" URI-zoobank="http://zoobank.org/94F18819-5AF5-4100-AB35-AA3C3976EE80" article-type="research-article" checkinTime="1721676390257" checkinUser="pensoft" docAuthor="Angus, Robert B., Maté, Jason F., Angus, Elizabeth M. & Král, David" docDate="2024" docId="A3AAB4F7DFCF5000804CA4F8B4FB44E5" docLanguage="en" docName="ZooKeys 1207: 205-299" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/117225/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Aphodius (Liothorax) plagiatus subsp. sinoplagiatus Angus, Maté, Angus & Král, 2024, subsp. nov." docType="treatment" docUuid="4ADB8359-223B-4AFA-B942-510A6EF9C63C" docUuidSource="ZooBank" docVersion="1" lastPageNumber="299" masterDocId="255F642057055F509167509E6C76FA1C" masterDocTitle="Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" masterLastPageNumber="299" masterPageNumber="205" pageNumber="205" updateTime="1721676524171" updateUser="pensoft">
|
||||
<mods:mods id="E0769A223B2D97997F8B2E3FB175E140" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="7BE562801989636CD206C8D2154B1A13">
|
||||
<mods:title id="2B6381DE4C7638C122E34BD27E55862E">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="BC4A52556332A3A9145AD44892F1C6D2" type="personal">
|
||||
<mods:role id="FE46F03ED3DA51A767FCC7F9FDFC395F">
|
||||
<mods:roleTerm id="02861E59500DDA402BFECDCE69F29B70">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="66523042919E349386A8486DD8E7F55F">Angus, Robert B.</mods:namePart>
|
||||
<mods:nameIdentifier id="15D7300CC8B7B828406DC2F46F642950" type="ORCID">0000-0002-3860-5617</mods:nameIdentifier>
|
||||
<mods:affiliation id="F54D7C07D820694802510C06BF664B67">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="C7CDEA0DB7843B8F7BB34F9F401F35C8" type="personal">
|
||||
<mods:role id="7C6915F1A6EDE79047945C64E629A18A">
|
||||
<mods:roleTerm id="C372E0C559E2EAB376F36554BF496BBA">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="6EF04F29E3BC4AAA1AE6FD1F0C079408">Maté, Jason F.</mods:namePart>
|
||||
<mods:nameIdentifier id="AABCE4E1FA3AC0250673BC839099F0DA" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
|
||||
<mods:affiliation id="89C4544C1664FE9E45580172BDFD8ABF">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="BF5BD88C173AE96C724A78DFE3460F92" type="personal">
|
||||
<mods:role id="9CB5D96F4CC9A8722C61B179CE57D3C2">
|
||||
<mods:roleTerm id="93D888703D9DB911689AC9B422B5668C">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="92ACFE5C0BB54A4841CB67E33416D41B">Angus, Elizabeth M.</mods:namePart>
|
||||
<mods:affiliation id="82030174195F6D0C365D8F4261D44EE2">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="15A3041571ADE45C78A89134C7811FA4" type="personal">
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|
||||
<mods:roleTerm id="6B79ACA2F0B493E72C64358790F8890E">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="4B2237A290B2D04A915CF992F79087BF">Král, David</mods:namePart>
|
||||
<mods:nameIdentifier id="68975E18440370E426E3B5D09EEB7B0C" type="ORCID">0000-0002-7930-1927</mods:nameIdentifier>
|
||||
<mods:affiliation id="36D05A68E7D07FED72395E0C19C55738">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="956561EC86BBABBFB2B6A5A7980C2B9B">text</mods:typeOfResource>
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||||
<mods:title id="FDAAC4F23CA1585A2EC027E44F578CD0">ZooKeys</mods:title>
|
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</mods:titleInfo>
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<mods:part id="90694123FD18302395E589F65AA18B98">
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<mods:date id="1DB17646602845AAFF9A5591C65C76A9">2024</mods:date>
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||||
<mods:detail id="105499DD83E6B22996CFCE25BD726F1C" type="pubDate">
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<mods:number id="6AE12C9E81796CDF224311CFDC5D8D4B">2024-07-22</mods:number>
|
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</mods:detail>
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||||
<mods:detail id="2217942084768D974524AC8157FC66F2" type="volume">
|
||||
<mods:number id="42E97C1722CB6791AF24F9410FE1229F">1207</mods:number>
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||||
</mods:detail>
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<mods:extent id="65375345ECE4AC3674C594B5145B9BF0" unit="page">
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<mods:start id="358BE85CB8E88F3465D0986E27DB4A6D">205</mods:start>
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<mods:end id="1173A37CDF46B7F13D2D12A16A5F52F2">299</mods:end>
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<mods:classification id="CB5E190A79F3A664F2A23242AD0D69F6">journal article</mods:classification>
|
||||
<mods:identifier id="AC08BAFABB0613C34C2AEBB8E32EF327" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
|
||||
<mods:identifier id="B0DA3DE29A9BC1A609E26E91C522192C" type="ZooBank">94F18819-5AF5-4100-AB35-AA3C3976EE80</mods:identifier>
|
||||
</mods:mods>
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||||
<treatment id="A3AAB4F7DFCF5000804CA4F8B4FB44E5" ID-arpha="A3AAB4F7-DFCF-5000-804C-A4F8B4FB44E5" ID-zoobank="https://zoobank.org/4ADB8359-223B-4AFA-B942-510A6EF9C63C" LSID="urn:lsid:zoobank.org:act:4ADB8359-223B-4AFA-B942-510A6EF9C63C" LSID-ZBK="urn:lsid:zoobank.org:act:4ADB8359-223B-4AFA-B942-510A6EF9C63C" httpUri="http://treatment.plazi.org/id/A3AAB4F7DFCF5000804CA4F8B4FB44E5">
|
||||
<subSubSection id="F3900B8D59A07F698C82B9E70F63AE0E" type="nomenclature">
|
||||
<paragraph id="523BFE8E11EB7483B0A79B80106FB327">
|
||||
<taxonomicName id="943FE3A5817D5BE9F1C36332CC1A6C06" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="plagiatus" status="subsp. nov." subGenus="Liothorax" subSpecies="sinoplagiatus">
|
||||
<emphasis id="5C720D500779E5C12FE677C03AB20068" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="BC91979B8EB2347543E0C14F038A33CF" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="6B2E5F77FFD7456F8B67599E9C9B0B83" italics="true">plagiatus sinoplagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="925C79CC89E2BCF2AF0AA370E5313363" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C39BF202A0E11FF408D651CCD8F2C7B4" type="description">
|
||||
<paragraph id="4F61C834B410379BC878069D83760A75">
|
||||
<figureCitation id="F02DBE427D5997920B1F7CE80EEEED21" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">Figs 22 j, k</figureCitation>
|
||||
,
|
||||
<figureCitation id="A426DA97BAAADAD129363EB3D2609BBB" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 f, g</figureCitation>
|
||||
,
|
||||
<figureCitation id="F561098015CDEC129D04A5C7F89CD942" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Mitotic chromosomes, arranged as karyotypes a, b A. plagiatus plagiatus, ♂, GB, Dorset, Studland a plain (Giemsa stained) b the same nucleus C-banded c A. plagiatus sinoplagiatus, ♂, CH, Qinghai, Gangca, plain d, e A. rutilipennis, ♂, CY, Zakaki Marshes d plain e C-banded f – h A. kraatzi ♀, SK f, g nucleus without B-chromosomes f plain g C-banded h, i nucleus with 1 B-chromosome h plain i C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure30" httpUri="https://binary.pensoft.net/fig/1097512">30 c</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EDSFK" type="type material">
|
||||
<paragraph id="7379DD66FF9974A9B3349FBD7162EF0A">
|
||||
<heading id="DBA5C2DB9FA1584D7D4FEADDEAF79164" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="19F1F24CFAD90A05745F718B84A3AFB7">
|
||||
<emphasis id="0AD21DAB8C2FA2DF90BA9CD225498100" bold="true" italics="true">
|
||||
<typeStatus id="E6A18E563F1BA37D1BDDAEEED8D4B231" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="5C0464B64A87ECBD113BD680B740D6BC" count="1" type="female">♀</specimenCount>
|
||||
Amdo / 1884 / Przevalsky (printed label), Zaidam / Burch Budde / 1-11 - 000 //
|
||||
<taxonomicName id="36FD34BE9B213175E270FB9316660F1F" authorityName="Hellwig" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="843129B493034F1C13AF1885D93D98CD" italics="true">Aphodius</emphasis>
|
||||
</taxonomicName>
|
||||
/
|
||||
<taxonomicName id="BDD80E204BA46AD55E2D34E9429C19D9" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="13936BD8EB913D8A5D72ACC38362C80D" italics="true">plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
/ var. (handwritten) (
|
||||
<named-content id="38B712C00BB28FA9779ADEB5DD7974ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="E3ADA83DE9ABBA11542276C29094397D" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<emphasis id="8AE35A20D01366AAE8A008B08111606D" bold="true" italics="true">
|
||||
<typeStatus id="C83CE96446BE81609C7F30FE67F8B43D" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="40BCBF9B4739F36D2791973F18EFD5D4" count="1" type="male">1 ♂</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="6229643AD034BED61B8B3426C7699A91" type="holotype">holotype</typeStatus>
|
||||
(originally on the same pin) (
|
||||
<named-content id="70F3884383D972E7E5D5C5385FD353AD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="C7C8542B0B00AD5FA78FEE02B9389AEA" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="12CF7A7121B0EDDA03C467F49DBF5171" count="1" type="male">1 ♂</specimenCount>
|
||||
<collectingCountry id="9F64361AD3E634472F0E2E5D6BE0460E" name="China">China</collectingCountry>
|
||||
<collectingRegion id="BD4E52941EF285B7D8D1E10EDFF0852A" country="China" name="Qinghai">Qinghai</collectingRegion>
|
||||
/ N.
|
||||
<collectingRegion id="6DDDB4EE9A610B75CCAB9E8F61C7ACC0" country="China" name="Qinghai">Qinghai</collectingRegion>
|
||||
Hu / Gangca. Roadside / pool Printed label and a further label with the same data in Chinese.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENTFK" specific-use="{"type":"Point","coordinates":[100.183333,37.300000]}">
|
||||
<geoCoordinate id="23AC045431B87C9ECF55FFB520187D8E" degrees="37" direction="north" minutes="18" orientation="latitude" precision="925" value="37.3">37 ° 18 ' N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="FC67FFFCF4C0D30861F1141D658D4790" degrees="100" direction="east" minutes="11" orientation="longitude" precision="925" value="100.183334">100 ° 11 ' E</geoCoordinate>
|
||||
</named-content>
|
||||
/
|
||||
<quantity id="444DA29544508F5CDDC789D8785B629E" metricMagnitude="3" metricUnit="m" metricValue="3.37" unit="m" value="3370.0">3370 m</quantity>
|
||||
.
|
||||
<date id="16409C0F433FBFA91AEA9B91BDBA23CF" value="2013-06-05">5. vi. 2013</date>
|
||||
, / R. B. Angus, F.
|
||||
<emphasis id="A5559540E6C56E814471285AE10F655D" italics="true">L.</emphasis>
|
||||
Jia & Y. Zhang. //
|
||||
<taxonomicName id="D037DD274F347BF8A6C1141048E77850" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="560F5BA39726AF66F5A8DF925EE4EDEF" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<emphasis id="8B874E87A59D2B88FD0E621639563FB1" italics="true">L.</emphasis>
|
||||
) / Chromosome prep. /
|
||||
<date id="C9040CDB34298F92D9A888ABF7A13A69" value="2013-06-06">6. vi 2013</date>
|
||||
. / R. B. Angus // JFM 130815017 (printed labels) (
|
||||
<named-content id="7642F3AA97AD546374D417E5360FA8F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="E2E92351DB9225B8CD38064DD282DD77" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="9B2B211D3A5DCB46C3FCDD506B411140" count="1" type="female">1 ♀</specimenCount>
|
||||
Cina-Gansu / Dingxi / 7.8 – 12.8 1996 / E. Kučera leg. // Ex coll. David Král / (
|
||||
<named-content id="52D10F2179DB0D795DDA15431B5F150C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="967EC81A3A42C16B75BD9623ED9B8018" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERUFK" type="Differential diagnosis">
|
||||
<paragraph id="25B84F13BE6EAA75B4608C31FB5B91D8">
|
||||
<heading id="B264AD76B3A84D5F8E63FE3F8566B5B7" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6B9521E2940E0ADADEE838CE845EB387">
|
||||
<taxonomicName id="73AB40FB91D65C9EB33216A46CD648AC" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="plagiatus" subGenus="Liothorax" subSpecies="sinoplagiatus">
|
||||
<emphasis id="7337959046EAE8C9A1761BB393FCD29D" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="86196DDE203A99387148CEA258F78A9E" italics="true">L.</emphasis>
|
||||
)
|
||||
<emphasis id="172B9981F1BE614AE965419B7C1F09DC" italics="true">plagiatus sinoplagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<emphasis id="3E41BE76A945E61EF98339129000EC21" italics="true">A.</emphasis>
|
||||
(
|
||||
<emphasis id="9D765A87A7078860CE26B57D4AC300DD" italics="true">L.</emphasis>
|
||||
)
|
||||
<emphasis id="F0D474F9961E5C8815DCCAA2DB23A869" italics="true">
|
||||
p.
|
||||
<taxonomicName id="479BB12D26FC86A337E625FDFCC0B4E6" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">plagiatus</taxonomicName>
|
||||
</emphasis>
|
||||
by its very glossy varnished appearance, the slightly larger aedeagus (length ca
|
||||
<quantity id="0BE755D39CC787179D1BA01DCB00DE65" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" unit="mm" value="1.325" valueMax="1.4" valueMin="1.25">1.25–1.4 mm</quantity>
|
||||
) with abruptly widened paramere apices (Fig.
|
||||
<figureCitation id="B72C08303B1ADF8CA95EE97B557436AB" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 j, k</figureCitation>
|
||||
), and above all by the strikingly larger spermatheca, length ca
|
||||
<quantity id="8A14428D79C32B57D305726106548719" metricMagnitude="-4" metricUnit="m" metricValue="4.2" unit="mm" value="0.42">0.42 mm</quantity>
|
||||
(Fig.
|
||||
<figureCitation id="5811DF12545D38B361E1C20995A3E5D0" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 f, g</figureCitation>
|
||||
), as against ca
|
||||
<quantity id="8AFF2CABE2F7EFF42AD79BBC0A102C1D" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" unit="mm" value="0.36">0.36 mm</quantity>
|
||||
in
|
||||
<emphasis id="7231B79F3B9099B94B3D575758AB5651" italics="true">A.</emphasis>
|
||||
(
|
||||
<emphasis id="03AC0FEAC0E5B5DEC9E5C8BBD0E5786C" italics="true">L.</emphasis>
|
||||
)
|
||||
<emphasis id="DF53545A5E7FC68A0F372F9CA2C87F96" italics="true">
|
||||
p.
|
||||
<taxonomicName id="8A486A808F5D546BC8A14B5242B0CC94" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">plagiatus</taxonomicName>
|
||||
</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="FB65997455F875E22D6CC24ADA5C66F5" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 a – e</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZWFK" type="description">
|
||||
<paragraph id="81DF013856FAF0052DBA4E763B1251C0">
|
||||
<heading id="28DF32911B1160953EF6973AE4607E69" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6428AB8B7D69FA5C6A419ECD1FAEF631">
|
||||
Length:
|
||||
<quantity id="9752B0BF84F920E7A76F5EB790B6E63D" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
|
||||
(
|
||||
<specimenCount id="699CE42633D8681D797D248D96E918C3" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="05CC96A641E83BD1491421D56C43B1D5" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" unit="mm" value="4.35" valueMax="4.6" valueMin="4.1">4.1–4.6 mm</quantity>
|
||||
(
|
||||
<specimenCount id="7F7CFDBB2C33BDE099A67D8CF2C05404" count="1" type="female">♀</specimenCount>
|
||||
), width 1.8–2.0 mm (
|
||||
<specimenCount id="780BE43821C73B64B7CB5CFB868550EF" count="1" type="male">♂</specimenCount>
|
||||
), 1.8–2.0 mm. Head: glossy black, sparsely double punctate, clypeus bulging upwards, frontoclypeal suture without tubercles, straight over middle ¾, effaced laterally. Outer angles of anterior emargination rounded.
|
||||
</paragraph>
|
||||
<paragraph id="9000CE5EDE0C072CE9C4CB15B51431D5">
|
||||
Pronotum: highly arched transversely, lateral margins not visible from above medially. Surface sparsely double punctate, the larger punctures slightly stronger than on head, the
|
||||
<collectingRegion id="956A67389DF65AE437FFA58A282CD96D" country="China" name="Gansu">Gansu</collectingRegion>
|
||||
male more heavily punctate. Basal margin largely unbordered, the raised margin extending ca ¼ of the way in from the hind angles.
|
||||
</paragraph>
|
||||
<paragraph id="9D84D38E940203B78EFBA7C57381AF6B">
|
||||
Elytra black, highly polished (varnished), as scutellum. Metaventrite (
|
||||
<specimenCount id="4CCD90F6D1F9B38E25B4E7B30E573D5D" count="1" type="male">♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="567C9B53E5A5F7A021C66F69395CC849" count="1" type="female">♀</specimenCount>
|
||||
) as in the nominal subspecies. Legs mid- to dark brown, longer mesotibial spur longer than basal segment of mesotarsus.
|
||||
</paragraph>
|
||||
<paragraph id="0855E31A03A3A32FC518F848A4164EBF">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="A0086BCEC72358EA7551DB065CE3DD3B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 j, k</figureCitation>
|
||||
) length
|
||||
<quantity id="76623A5699F0A0F78697D12E2E828805" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" unit="mm" value="1.325" valueMax="1.4" valueMin="1.25">1.25–1.4 mm</quantity>
|
||||
, outer apical membranous part abruptly widened.
|
||||
</paragraph>
|
||||
<paragraph id="64FC6E1AA60D9B47CD2D8A3213836F46">
|
||||
Spermatheca (Fig.
|
||||
<figureCitation id="CD4D87FE4F77839EA3959D7DCE717E87" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 f, g</figureCitation>
|
||||
) length ca
|
||||
<quantity id="26B37D861A82CCF555DB79C234EE63C0" metricMagnitude="-4" metricUnit="m" metricValue="4.2" unit="mm" value="0.42">0.42 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMXFK" type="etymology">
|
||||
<paragraph id="A5626687ADEDACE50300BBE4168779DD">
|
||||
<heading id="67766590265D9C6430B9EA02A8B30EA1" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F0FD78DC728DD70B7FA41867C9E93538">
|
||||
<tp_taxon-name id="7925B7AAA867168FB214110777F8F674">
|
||||
<tp_taxon-name-part id="B20D12F628F79851F76A588C4A392B4F" reg="sinoplagiatus" taxon-name-part-type="species">
|
||||
<emphasis id="8FE67E8F8BC803F57B04E178314CFB25" italics="true">sinoplagiatus</emphasis>
|
||||
</tp_taxon-name-part>
|
||||
</tp_taxon-name>
|
||||
is an adjective indicating its Chinese distribution.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3XFK" type="remarks">
|
||||
<paragraph id="D7E0A73A0CD6E915D21252DA23F97199">
|
||||
<heading id="C10219843217532D132F1C700CFC9F4B" reason="title">Remark.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FF90B36045E225EDC79CABBA07A41053">
|
||||
The aedeagus of
|
||||
<specimenCount id="886BF0249D3790E7D073D2376BFC7107" count="2" type="male">2 ♂♂</specimenCount>
|
||||
from near
|
||||
<collectingRegion id="A824FA9308D6A80FF92C466902CD6D94" country="China" name="Beijing">Beijing</collectingRegion>
|
||||
(Fig.
|
||||
<figureCitation id="B505C59173A98AD6BA0AD804CF0967DE" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 i</figureCitation>
|
||||
) resembles that of
|
||||
<tp_taxon-name id="65D73E59852151815043F2355EBF1DA6">
|
||||
<emphasis id="323C81C6E616BB306428D57704EA9EE1" italics="true">
|
||||
<tp_taxon-name-part id="87667D60D3FFCAB0E04049995B365602" reg="Liothorax" taxon-name-part-type="genus">L.</tp_taxon-name-part>
|
||||
<tp_taxon-name-part id="1AB61DEC0C8BB4CF170276EA297B00DC" reg="plagiatus" taxon-name-part-type="species">p.</tp_taxon-name-part>
|
||||
<tp_taxon-name-part id="94EF90F4AA3E20679D8E664110BDF6A2" reg="sinoplagiatus" taxon-name-part-type="subspecies">sinoplagiatus</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
in its widened paramere apices, but is smaller, length ca
|
||||
<quantity id="207DDD92AF9AEA0DF0E79584DD6A91A9" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
437
data/AF/8B/C1/AF8BC118B8C0561788F1062974FE8977.xml
Normal file
437
data/AF/8B/C1/AF8BC118B8C0561788F1062974FE8977.xml
Normal file
|
|
@ -0,0 +1,437 @@
|
|||
<document id="29F71CD27973CA21C42DEF78FD2D618E" ID-DOI="10.3897/zookeys.1207.117225" ID-ZooBank="94F18819-5AF5-4100-AB35-AA3C3976EE80" ID-publisher-id="117225" URI-arpha="255F6420-5705-5F50-9167-509E6C76FA1C" URI-zoobank="http://zoobank.org/94F18819-5AF5-4100-AB35-AA3C3976EE80" article-type="research-article" checkinTime="1721676390257" checkinUser="pensoft" docAuthor="Angus, Robert B., Maté, Jason F., Angus, Elizabeth M. & Král, David" docDate="2024" docId="AF8BC118B8C0561788F1062974FE8977" docLanguage="en" docName="ZooKeys 1207: 205-299" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/117225/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Aphodius (Liothorax) niger Illiger 1798" docType="treatment" docVersion="1" lastPageNumber="299" masterDocId="255F642057055F509167509E6C76FA1C" masterDocTitle="Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" masterLastPageNumber="299" masterPageNumber="205" pageNumber="205" updateTime="1721676524171" updateUser="pensoft">
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<mods:mods id="D90B918E9F00780964799FEBB83D2BCA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6B71869391574E0CD159C3418C40275D">
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<mods:title id="D664EC5BD14D5B4A6026246C1CBA7E77">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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||||
</mods:titleInfo>
|
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|
||||
<mods:nameIdentifier id="E7B765CCA30845D728C7C6828B42542D" type="ORCID">0000-0002-3860-5617</mods:nameIdentifier>
|
||||
<mods:affiliation id="27AEC6345E1A81C41F3BE6B1C324F165">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="91C355F5106F2759893F1FFF1B56C4B1" type="personal">
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|
||||
<mods:nameIdentifier id="2EBC07ED95046A565B5DE6E3072DB4A6" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
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<mods:affiliation id="6917CC5673D466E342D16D6194CC70E9">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="804B5874E7887FE03D1BF7F6957AD558">Angus, Elizabeth M.</mods:namePart>
|
||||
<mods:affiliation id="D743A369AAE424AF0A56CB5033DB065D">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="A5C9AF67B579B15AF1D0130F2BC95776">Král, David</mods:namePart>
|
||||
<mods:nameIdentifier id="1F167585EE4948D547CF64918765652D" type="ORCID">0000-0002-7930-1927</mods:nameIdentifier>
|
||||
<mods:affiliation id="FEB24633AE35E7B01A8406875E691523">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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</mods:name>
|
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<mods:typeOfResource id="FB09D3D421660255726AB3BD653AD4EA">text</mods:typeOfResource>
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<mods:title id="9534218F9093AE9328FDB19184351C51">ZooKeys</mods:title>
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<mods:part id="338F51BE800D547234E48873AA8EDF4B">
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<mods:date id="1547D7FBDB9864C5168F4451A9315EF4">2024</mods:date>
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<mods:number id="0B012C824D57085E1B91CD419507D447">2024-07-22</mods:number>
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<mods:identifier id="9D7A4F1ACB9F37BCC670C103195C6EF4" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<subSubSection id="49920B18E44093F7C9BAF599154232EA" type="nomenclature">
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<paragraph id="DB5B6F3793CAE578B049A1AC336BE7F2">
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||||
<taxonomicName id="773866007EEFD13CBDCB9E4E1316C614" ID-CoL="FHT7" authority="Illiger, 1798" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger" subGenus="Liothorax">
|
||||
<emphasis id="B97B60B7333329ABCC7940DB9B73DB4F" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="F91B6456CB6B78F0D8B7A9C83A1F7B63" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="F4C6B0B4F20044A3E16F763419AD0092" italics="true">niger</emphasis>
|
||||
Illiger, 1798
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
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<subSubSection id="C640E24A96B8FF582FA0BF6700D961CF" type="description">
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<paragraph id="3A2F7AD28A74C46413520F3B636D99C5">
|
||||
<figureCitation id="220E7885C20228A07F80BA3110FB0937" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 a</figureCitation>
|
||||
,
|
||||
<figureCitation id="2C5A8F22DF1E3EE87F8F9D79EA6D1A31" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 e</figureCitation>
|
||||
,
|
||||
<figureCitation id="46BB9A5CB2F93CED57130CE98AC78EC1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 a – c</figureCitation>
|
||||
,
|
||||
<figureCitation id="A8C027A70F1FA78FC104E49D74D82CAF" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." figureDoi="10.3897/zookeys.1207.117225.figure15" httpUri="https://binary.pensoft.net/fig/1097476">15 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="35A1AB950621997C67EE0CFA782AFBF7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 l, m</figureCitation>
|
||||
,
|
||||
<figureCitation id="1DB8D17A6B941208B8E8C84D53CF26D7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 i</figureCitation>
|
||||
,
|
||||
<figureCitation id="E8F97FFC1EAAA0A6FF5956D766819515" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Elytral apices a A. (L.) niger, SV, Södermanland, Tullgarn b A. (Nialus) varians, FR, left elytron c A. (Calamosternus) granarius, CY, right elytron d A. (C.) hyxos, CY, left elytron e A. (Agoliinus) lapponum, GB, Cumbria, both elytra f A. (Agrilinus) constans, IT, Sardinia, left elytron. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure19" httpUri="https://binary.pensoft.net/fig/1097484">19 a</figureCitation>
|
||||
,
|
||||
<figureCitation id="E89A1C2A68A5A981FCA3B0B359A2EEB3" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 a – c</figureCitation>
|
||||
,
|
||||
<figureCitation id="7D0AF7DA82361512ADC2A9B0E58DA45C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 a – f</figureCitation>
|
||||
,
|
||||
<figureCitation id="0CD05893EEF6F4CD0E160C81725A303B" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 a</figureCitation>
|
||||
,
|
||||
<figureCitation id="C9D3F6584339539A9D44F0C5472E945D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure25" httpUri="https://binary.pensoft.net/fig/1097496">25 a</figureCitation>
|
||||
,
|
||||
<figureCitation id="6A6AC5EB2508669DEBEE8D8A901AE1EE" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">
|
||||
<quantity id="E4755080BC079D5F42C7074B65F0111B" metricMagnitude="4" metricUnit="kg" metricValue="2.6" unit="t" value="26.0">26 t</quantity>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="097A555D11CA87F369754439F1908927" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitotic chromosomes, arranged as karyotypes a, b A. niger, ♂, SV, Tullgarn a plain b the same nucleus C-banded (1 replicate of autosome 5 missing) c A. niger ♀, SV, Tullgarn, C-banded d, e A. niger, ♂♂, GB, Hampshire, New Forest, C-banded e with 3 B-chromosomes f, g A. muscorum, ♂, HU, Hortobágyi f plain g the same nucleus C-banded h A. muscorum, ♂, HU, Hortobágyi, C-banded i, j A. muscorum, ♀, HU, Kisújszállás i plain j C-banded k, l A. muscorum, ♀, HU, Kisújszállás k plain l the same nucleus C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure31" httpUri="https://binary.pensoft.net/fig/1097515">31 a, b, d, e</figureCitation>
|
||||
,
|
||||
<figureCitation id="C9131A7AAEE25CB50DDD1AFD0CE263DA" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Meiotic chromosomes, metaphase I from testis a – d A. niger, GB, Hampshire, New Forest, White Moor a, c plain b, d the same nuclei C-banded e – g A. muscorum, HU, Hortobágyi e plain f, g the same nucleus f plain g C-banded h A. bameuli paratype, FR, Corsica, by Lac de Melo, plain i, j A. krelli IT, Sardinia, paratype i plain j the same nucleus C-banded k, l A. wilsonae, SP, Areños k plain l the same nucleus C-banded. The sex bivalent is indicated by an arrow. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure34" httpUri="https://binary.pensoft.net/fig/1097526">34 a – d</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
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|
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<paragraph id="1491460A10BCDB27ABB453606AD20DE5">
|
||||
<treatmentCitationGroup id="8C985D4E02EE03EB6FF5D6D17A09D9C5">
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<treatmentCitation id="8F0701C1BBED69804E27E9F606D3B886">
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<taxonomicName id="4E0E2DB354D3ADFF2AFC4FBC59EE3AB9" ID-CoL="FHT7" authority="Illiger, 1798: 24" authorityName="Illiger" authorityPageNumber="24" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="A1D89CFDBDEB8E637E998CB0B26ED929" italics="true">Aphodius niger</emphasis>
|
||||
Illiger, 1798: 24
|
||||
</taxonomicName>
|
||||
. Conserved as valid name: International Commission for Zoological Nomenclature Opinion 2009 (2005: 45).
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERHAM" type="type material">
|
||||
<paragraph id="8C4673081A4C6A8AB8474F49BC3A6C37">
|
||||
<heading id="D6E7AD72E274EC1D2B9EB55C6AB5447B" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1E5B9A35DC23479DB7912B4474581B6E">
|
||||
<materialsCitation id="213FF3365280D9359CCF7FB51AB06DED" collectionCode="ZMB" specimenCount="1" typeStatus="lectotype">
|
||||
<emphasis id="F25A69A0193055CCA4240FA10D7661B9" bold="true" italics="true">
|
||||
<typeStatus id="86F416316612102089DA959023133D1A" type="lectotype">Lectotype</typeStatus>
|
||||
</emphasis>
|
||||
, unsexed, “ Suec ”, designated by
|
||||
<bibRefCitation id="4F0A8120B7743E579F9F5CDDFF910416" author="Krell F-T & Mann DJ & Angus RB & Maté JF" etAl="et al." firstAuthor="Krell" issue="2" journalOrPublisher="Bulletin of Zoological Nomenclature" pagination="127-131" refId="B27" refString="Krell F-T, Mann DJ, Angus RB, Maté JF (2003) Case 3254 Aphodius niger Illiger, 1798 (Insecta, Coleoptera): Proposed conservation of the specific name. Bulletin of Zoological Nomenclature 60 (2): 127–131." title="Case 3254 Aphodius niger Illiger, 1798 (Insecta, Coleoptera): Proposed conservation of the specific name." volume="60" year="2003">Krell et al. (2003)</bibRefCitation>
|
||||
(
|
||||
<abbrev id="ABBRID0EAIAM" xlink_title="Humboldt Museum">
|
||||
<collectionCode id="90A5AF045D57F53D91A81D1DD0614C25" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEIAM" type="material">
|
||||
<paragraph id="CD968201A0D2E7F94E5678073DA49473">
|
||||
<heading id="FA08EB652F0FA6CDFCF3A0B74CEEB7F4" reason="title">Additional material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B38CFE09D22C877C52A268F3633310C7">
|
||||
SV:
|
||||
<collectingRegion id="6AAC84F658A8134A8B793D4AEF9A9B4B" country="Sweden" name="Sodermanland">Sodermanland</collectingRegion>
|
||||
, Holo, Tulgarn, Nasuden.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENIAM" specific-use="{"type":"Point","coordinates":[17.620000,58.950000]}">
|
||||
<geoCoordinate id="A82BA3EC82A78FF1510C7EF261C10056" degrees="58.95" direction="north" orientation="latitude" precision="555" value="58.95">58.95 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="97FF730A282DC0C3786A8647096D3872" degrees="17.62" direction="east" orientation="longitude" precision="555" value="17.62">17.62 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Trampled bare organic mud.
|
||||
<date id="CE5A738E070F805D9685EBFBAAC3F315" value="2011-05-15">15. v. 2011</date>
|
||||
. Hans-Erik & Livia Wanntorp.
|
||||
<specimenCount id="DCA5CA9F90C331EEF4444B3A656E0EEB" count="4" type="male">4 ♂♂</specimenCount>
|
||||
, chromosome preps 2:
|
||||
<date id="1C84CD0D9208D46AE11025732463CD6A" value="2001-05-24">24. v. 2001</date>
|
||||
, 1:
|
||||
<date id="CA2A7F0B30B977CBA756B2822C6135D9" value="2011-05-25">25. v. 2011</date>
|
||||
, 2:
|
||||
<date id="68E671D1DC8A0A7F50D3BC92ADADD0F0" value="2011-05-25">25. v. 2011</date>
|
||||
, 1:
|
||||
<date id="120AB96A9234774F75CB8CFE50DC221E" value="2011-05-31">31. v. 2011</date>
|
||||
, R. B.
|
||||
<collectingRegion id="C8077C25C0C8ED0F79FE5D11BD1C29AE" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
;
|
||||
<specimenCount id="F674A0B5E707EBE48FF29C50C82901DF" count="7" type="female">7 ♀♀</specimenCount>
|
||||
, chromosome preps 3:
|
||||
<date id="1D711F19B833BE035B40AD32BF1420BA" value="2011-05-24">24. v. 2011</date>
|
||||
, 4:
|
||||
<date id="645047CBBC03A4BE60AAEBC275391ACD" value="2011-05-24">24. v. 2011</date>
|
||||
, 9:
|
||||
<date id="DB7FC9FE5279CD8C1B882666F10A3BFC" value="2011-05-25">25. v. 2011</date>
|
||||
, 2:
|
||||
<date id="5825ECE91922AEED9D13C57D9D6BC532" value="2011-05-31">31. v. 2011</date>
|
||||
, 3:
|
||||
<date id="A3BC13BC80D18D22AFE1FC0F43B87B6F" value="2011-05-31">31. v. 2011</date>
|
||||
, 4:
|
||||
<date id="DAE909C83A5FE054CFDA260F673CB9AB" value="2015-05-31">31. v. 2015</date>
|
||||
, 5:
|
||||
<date id="18225223DEDAC16456D62954BA5B97D3" value="2011-05-31">31. v. 2011</date>
|
||||
, R. B.
|
||||
<collectingRegion id="74A5E96FCE665FD3B89D99197EE86E69" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
(
|
||||
<named-content id="F1027E7393285F463647335A9F371325" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="720CAAE6CC05BDD9C77B6CBCE41BE37F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="65CF96CCF242F0049B62A22464ACAAED" count="1" type="male">1 ♂</specimenCount>
|
||||
same data, chromosome prep. 2:
|
||||
<date id="2B886B3D64EC4ED1FE08A51DDC04A686" value="2011-05-24">24. v. 2011</date>
|
||||
(
|
||||
<named-content id="D7B692B9238CE9DEAEC514457F92693A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="FF6CD3379F3998C39D0A154425BAAFF9" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0480F8677DA123747B36C820C5F107D6">
|
||||
GB:
|
||||
<collectingRegion id="9722F7FA8B4DF797CF310130DAA1F6E2" country="United Kingdom" name="Hampshire">Hampshire</collectingRegion>
|
||||
, New Forest. Long Slade Bottom.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6IAM" specific-use="{"type":"Point","coordinates":[-1.619000,50.801000]}">
|
||||
<geoCoordinate id="05ED48EE86E904138A6C80CE7B17B0B5" degrees="50.801" direction="north" orientation="latitude" precision="55" value="50.801">50.801 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="C48770F04278479FDA466131D720666A" degrees="1.619" direction="west" orientation="longitude" precision="55" value="-1.619">1.619 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
. Temporary pool after heavy rain.
|
||||
<date id="34C8D5623525208937D041CF5426B472" value="2002-05-30">30. v. 2002</date>
|
||||
, R. B.
|
||||
<collectingRegion id="422121C30CE6E235949BFABE4FB80271" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="9D88CC7E7685A02D1B3E52D8AEBC5A5A" count="4" type="male">4 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="6175475EC479F0AFBA9AA4AA69F961A3" count="4" type="female">4 ♀♀</specimenCount>
|
||||
, 5 unsexed. Chromosome preps 1–3: 31. v., 2: 1. vi, 4:
|
||||
<date id="1D74AA6DC4180348377C97144541A388" value="2011-06-01">1. vi. 2011</date>
|
||||
, R. B.
|
||||
<collectingRegion id="0D3A0E90293BEE4E139AE0D9AEA2C75F" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
(
|
||||
<named-content id="9A260F78F8C6DF6C8DBB548F9AE5DB18" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="84EB037B3D6A830B56D32227D47D8CC3" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="F6DBA67248F607EDD6185B892A5C0630" count="1" type="male">1 ♂</specimenCount>
|
||||
, same data. (
|
||||
<named-content id="70905218842D272EF445C3646736F933" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="24283CA7EC558CEC10FDB364CB797B44" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
); New Forest, White Moor Pond.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPJAM" specific-use="{"type":"Point","coordinates":[-1.607000,50.820000]}">
|
||||
<geoCoordinate id="D1536232211E47E39AF5D00043CF403F" degrees="50.820" direction="north" orientation="latitude" precision="55" value="50.82">50.820 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="46EF09FFAD31D4B034BE2EFF6417FE45" degrees="1.607" direction="west" orientation="longitude" precision="55" value="-1.607">1.607 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="D51B178E25D5629CEAF2DECC4930632E" value="2015-10-02">2. x. 2015</date>
|
||||
,
|
||||
<date id="F85ABD2EB0C4F9AF92EEBF25F39CD16C" value="2016-05-02">2. v. 2016</date>
|
||||
, R. B.
|
||||
<collectingRegion id="ACD7740E8E5E160333A0BDF58B81B524" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="66EBCDEBE6CFFC2C443F0A8D853EC9C9" count="8" type="male">8 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="1B2AC82ACBB3A2A29299AE28E0AC8300" count="7" type="female">7 ♀♀</specimenCount>
|
||||
, 3 unsexed. Chromosome preps: 3:
|
||||
<date id="1C4E761A17CD83D946F1EA2BB21E8F79" value="2015-10">x. 2015</date>
|
||||
, 5:
|
||||
<date id="071DDF65553EE1DF7D7533750DE306C8" value="2016-05">v. 2016</date>
|
||||
. New Forest, Balmer Lawn.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXJAM" specific-use="{"type":"Point","coordinates":[-1.567000,50.828000]}">
|
||||
<geoCoordinate id="09ADC8DD03A4525E9C44FFA9A3136CE0" degrees="50.828" direction="north" orientation="latitude" precision="55" value="50.828">50.828 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="69387DA92148F51C32A3BED1244037E1" degrees="1.567" direction="west" orientation="longitude" precision="55" value="-1.567">1.567 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
. At edges of pond and ditch.
|
||||
<date id="87C7638D1D97DA6F8313F5AEBC121C34" value="2002-05">V. 2002</date>
|
||||
. R. B.
|
||||
<collectingRegion id="C38026BF2024ECD7F3C311BFA90E1AB4" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
(
|
||||
<named-content id="830FD7AA73B99736DB0840EEA92D9454" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="8998A8F82D1E77E021AB639001803771" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). Inkpen Common, Berkshire
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDKAM" specific-use="{"type":"Point","coordinates":[-1.453000,51.375000]}">
|
||||
<geoCoordinate id="565497BBB6C216B26C0E9324EB3AD6CC" degrees="51.375" direction="north" orientation="latitude" precision="55" value="51.375">51.375 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="AF1479C0B9BEB21C26DF36F350798869" degrees="1.453" direction="west" orientation="longitude" precision="55" value="-1.453">1.453 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
(fide
|
||||
<bibRefCitation id="826798CA12790F010725F3613F08D1A3" author="Mann DJ & Garvey L" firstAuthor="Mann" issue="3" journalOrPublisher="Coleopterist" pagination="138-142" refId="B31" refString="Mann DJ, Garvey L (2014) Aphodius (Liothorax) niger (Illiger) (Scarabaeidae: Aphodiinae) inside and outside the New Forest. Coleopterist 23 (3): 138–142." title="Aphodius (Liothorax) niger (Illiger) (Scarabaeidae: Aphodiinae) inside and outside the New Forest." volume="23" year="2014">Mann and Garvey (2014)</bibRefCitation>
|
||||
, not examined) 1 unsexed (
|
||||
<named-content id="1F513F612843479FFF0E482AB88A2C1D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oxford-university-museum-natural-history" xlink_title="Oxford University Museum">
|
||||
<collectionCode id="6DB108FD22AF434568397A2ACC852CCE" country="United Kingdom" httpUri="http://grbio.org/cool/g6yf-wft0" name="Oxford University Museum of Natural History" type="">OUM</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0FF665D631719665AFBF6A28BF38294B">
|
||||
RU:
|
||||
<collectingRegion id="BC1FC490C4F9EB7D4F400A3AA8DF44BA" country="Russia" name="Yaroslavl'">Yaroslavl</collectingRegion>
|
||||
oblast’, Berditsino,
|
||||
<geoCoordinate id="E7C6CB9DBB559DF25D9B0DCCE7BCBEDF" degrees="57.454" direction="north" orientation="latitude" precision="55" value="57.454">57.454 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="47E64EA5914C3FBA12DDFEA0A1E1291C" degrees="40.1108" direction="east" orientation="longitude" precision="5" value="40.1108">40.1108 ° E</geoCoordinate>
|
||||
. leg. Yakovlev.
|
||||
<specimenCount id="19EB0375C08180E1FF16CCC1EAF9AE3F" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<named-content id="EF6B5EFCBA9B7E79922FFE6247B340C0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||||
<collectionCode id="463947FF1F1A91DF12BB50378B7CD71A" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWKAM" type="Differential diagnosis">
|
||||
<paragraph id="8B3C07DD2FDA8129F59FE1C0C97BAE5C">
|
||||
<heading id="19A4F054291FD95BA278F7F378F8177B" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FD8492977465EA3A018C5C7CE4C26E00">
|
||||
<taxonomicName id="48527262B01AEDC7E5BE2BC2331ECFA1" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="0AB315111688D1711A50AC0D50FEE83A" italics="true">Aphodius niger</emphasis>
|
||||
</taxonomicName>
|
||||
is immediately distinguished from all other
|
||||
<taxonomicName id="89DF6286ACDC1AE75605D18E082B8025">
|
||||
<emphasis id="8E4E13319C691D3B81C6CA5BC930ED69" italics="true">Liothorax</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(and all
|
||||
<taxonomicName id="6DF04FC7B7C3EE966C5BE83D20875F79" authorityName="Leach" authorityYear="1815" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Aphodiinae">Aphodiinae</taxonomicName>
|
||||
whose chromosomes are known) by its long, almost entirely euchromatic, Y chromosome (Fig.
|
||||
<figureCitation id="79FF85C641D389E21B965B24AFF3BA9B" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitotic chromosomes, arranged as karyotypes a, b A. niger, ♂, SV, Tullgarn a plain b the same nucleus C-banded (1 replicate of autosome 5 missing) c A. niger ♀, SV, Tullgarn, C-banded d, e A. niger, ♂♂, GB, Hampshire, New Forest, C-banded e with 3 B-chromosomes f, g A. muscorum, ♂, HU, Hortobágyi f plain g the same nucleus C-banded h A. muscorum, ♂, HU, Hortobágyi, C-banded i, j A. muscorum, ♀, HU, Kisújszállás i plain j C-banded k, l A. muscorum, ♀, HU, Kisújszállás k plain l the same nucleus C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure31" httpUri="https://binary.pensoft.net/fig/1097515">31 a, b, d, e</figureCitation>
|
||||
). The basal segment of the mesotarsus is always clearly longer than the longer mesotibial spur (Fig.
|
||||
<figureCitation id="104556791C34007426DFD7188DE4AA93" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 i</figureCitation>
|
||||
). Elytra glossy black, though interstices have fine reticulation. Central part of metaventrite without reticulation (Fig.
|
||||
<figureCitation id="86C1042F3494E0C0319D1C4B0D970269" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 a – d</figureCitation>
|
||||
). Endophallic teeth clearly longer than high. Length of longest endophallic teeth at least 40 μm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6LAM" type="redescription">
|
||||
<paragraph id="C9702EE6C78F4F20FE3B4F59BA9167DC">
|
||||
<heading id="CB613ECB9021B0A69573A6B1C79981B7" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FAC89330199CA16CED67EB378846D1FF">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="639452F0AA4CDFE272A8F9A55D7C15E8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 a</figureCitation>
|
||||
). Length
|
||||
<quantity id="AB918F647DF53B757F9BDCEE51982DF1" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="6180AB73245FF5450A5A65E975F65323" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="86180F5E246B0BB9C411C7549D2F702E" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.0" metricValueMin="4.1" unit="mm" value="4.55" valueMax="5.0" valueMin="4.1">4.1–5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="AFCDCE607BF1AA373ACAF9A566217194" count="1" type="female">♀</specimenCount>
|
||||
), width
|
||||
<quantity id="6FC2E7286462D77A8B6342D7E3C0FEA0" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.8–2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="DF7D4BFAAE097B756F599673C7D4F9EC" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="2C4EB9827DE60A4BB7F37D759F023CBE" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="07ACC0ABE57056484470789B21FFCEC1" count="1" type="female">♀</specimenCount>
|
||||
) (Swedish material),
|
||||
<quantity id="DC3F56186B6F5A75A78BC867D72A192E" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="B9BBE306EC324955DEB8FFB66C18086F" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="5E234F6FC9C7B8CB09AC1243DD8148CB" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.1" metricValueMin="4.4" unit="mm" value="4.75" valueMax="5.1" valueMin="4.4">4.4–5.1 mm</quantity>
|
||||
(
|
||||
<specimenCount id="BBEBC0EBC36EDB761E962F8FB9018B3A" count="1" type="female">♀</specimenCount>
|
||||
), width 2.0–
|
||||
<quantity id="799BB1D7E11680D30CC36161EC74FAAC" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="AC52931AF4886EB6FE8AD5CB1B2E6BF3" count="1" type="male">♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="6C9A8DC45EF8F3353155F1BFB8326C75" count="1" type="female">♀</specimenCount>
|
||||
) (English Material).
|
||||
</paragraph>
|
||||
<paragraph id="D77ABE4D3855D3A11FD88A5EF30D88BA">
|
||||
Head black, without obvious brownish margin. Anterior margin of clypeus excised over middle 1 / 3, the lateral edges of this excision bluntly rounded. Frontoclypeal suture very fine, without any trace of a tubercle, straight-transverse over middle 1 / 2, angled forward to run to the clypeo-genal junction laterally. The lateral parts of the suture may be almost completely effaced. Clypeus raised in a rounded bulge medially. Surface moderately strongly punctate with double punctation. Larger punctures separated by ca their own diameter over most of the head, but sparser on clypeal bulge. Antennae and palpi blackish brown. Epipharynx (Fig.
|
||||
<figureCitation id="45BA84A89AB9A766FFECE537462DDB5A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 a – c</figureCitation>
|
||||
) with the anterior margin of the clithra strongly excised either side of the median tylus and with a few fine acropariae. Chaetopedia stout, 3–6 each side. Chaetopariae ca as stout as chaetopedia, forming a close-set line each side. Apophobae very fine, arranged in a line outside that of the chaetopariae; prophobae vine, quite long, arranged against the sides of the mesoepitorma. Pronotum (Fig.
|
||||
<figureCitation id="A7955718A7ED8A2E6138BF67D86D1115" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 e</figureCitation>
|
||||
) hemicylindrical, black, highly arched transversely, weakly so longitudinally, lateral parts of surface bulging outwards so that the lateral margins are often not visible from above in their basal 1 / 3. Surface glossy black, with double punctation, the larger punctures separated by 1–2 × their diameter, sparser medially and sometimes petering out in anterior 1 / 8 where only the fine punctures are present. Scutellum elongate, pentagonal, glossy black with a few sparse punctures. Elytra black, interstices flat, 6–8 × the width of the striae, finely reticulate and with sparse fine punctures (Fig.
|
||||
<figureCitation id="32D810C09A80118CCB649F6D4D3088F9" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 l, m</figureCitation>
|
||||
), the reticulation stronger towards the apex (Fig.
|
||||
<figureCitation id="DEB47EF21CD4A715D0C29B2DC29B5B86" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Elytral apices a A. (L.) niger, SV, Södermanland, Tullgarn b A. (Nialus) varians, FR, left elytron c A. (Calamosternus) granarius, CY, right elytron d A. (C.) hyxos, CY, left elytron e A. (Agoliinus) lapponum, GB, Cumbria, both elytra f A. (Agrilinus) constans, IT, Sardinia, left elytron. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure19" httpUri="https://binary.pensoft.net/fig/1097484">19 a</figureCitation>
|
||||
). Striae glossy, without reticulation, with a single row of punctures separated by ca 2 × their diameter and excising the strial margins. Metaventrite moderately punctate, the median plate not reticulate, sometimes distinctly concave in males Fig.
|
||||
<figureCitation id="ACB77BE4B86F24FFF6974F19F9127312" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 a</figureCitation>
|
||||
but sometimes almost flat with a median impressed line, as in females (Fig.
|
||||
<figureCitation id="A2A9AEA476A2C5743B2282756E7BCCC5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 b, c</figureCitation>
|
||||
). Legs blackish brown, mesotarsi always with basal segment longer than longer mesotibial spur (Fig.
|
||||
<figureCitation id="F3920F2A0B655A11C7DE016A1F71CA52" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 i</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="AAFF7B9DAA835F382B006D40EAD45A24">
|
||||
Aedeagus (Figs
|
||||
<figureCitation id="C0ED862E2D8BA8F3652FF34A649AC89C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 a – f</figureCitation>
|
||||
,
|
||||
<figureCitation id="FCCDD66E059CF1FC51A2647F7AFC93A1" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 a</figureCitation>
|
||||
) length ca
|
||||
<quantity id="6F12E5DA7F83CE443AFBFE110028E05D" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
|
||||
, paramere length ca
|
||||
<quantity id="6B84CF9A26136AFDE6290B46D9672A90" metricMagnitude="-4" metricUnit="m" metricValue="4.8" unit="mm" value="0.48">0.48 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="4C80F36694A53D5A9F34031B30BF6179" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" unit="mm" value="0.69">0.69 mm</quantity>
|
||||
, tooth-field length ca
|
||||
<quantity id="F867F7775126E2E69ACC2169D93A7BF0" metricMagnitude="-4" metricUnit="m" metricValue="6.7" unit="mm" value="0.67">0.67 mm</quantity>
|
||||
(measurements from the cleared preparation; Fig.
|
||||
<figureCitation id="900B5636D8F35DCF57EA81AA422B2B18" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">24 a</figureCitation>
|
||||
). Length of longest teeth on endophallus 54–65 μm. Parameres with the sclerotised strut running from the base just mediad of the outer lateral margin to the inner apical angle. Apical soft pad of parameres rounded apically, not widened.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVNAM" type="remarks">
|
||||
<paragraph id="3C0B048FEC058E83DE0194F32D8AA5AC">
|
||||
<heading id="6065637E25049C31C7F588CAFDAED703" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BB19D14C64B78E9CC7F64736F76A0DB2">
|
||||
Chromosomally verified material is from the New Forest,
|
||||
<collectingCountry id="48F944AC107B59D664992452DD4FAA91" name="United Kingdom">England</collectingCountry>
|
||||
and Södermannland,
|
||||
<collectingCountry id="3204A0A4B8AD94DC03DD28CFDEC0510A" name="Sweden">Sweden</collectingCountry>
|
||||
. Material from Berditsino,
|
||||
<collectingCountry id="F3D6116D81DB64C1AAEBE308C310B531" name="Russia">Russia</collectingCountry>
|
||||
(
|
||||
<named-content id="139FF33B8244C9E63F24AC816F77DEDF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||||
<collectionCode id="3E3E90B7534DC8F65575E1DE7DCA7759" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||||
</named-content>
|
||||
) is taken to belong to this species. In the New Forest
|
||||
<taxonomicName id="E6A2DA8934E4B57282EECE3C93EEDB63" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="81E58BA875DA299C4BCE3C5CCBA8C09B" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
is typically taken at the edges of pools and ditches in April-May, but when large numbers are taken it is almost always due to inwash from the surrounding grassland after heavy rain (Fig.
|
||||
<figureCitation id="4843AAAFA2F35CF435D7A5DA0FA3D9BB" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 a</figureCitation>
|
||||
). In Britain, outside of the New Forest,
|
||||
<taxonomicName id="0319229C7AB21B19A248F5C4095598E7" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="1443E80F540700170202D7E10DB5F983" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
has recently been found at Inkpen Common near Hungerford in Berkshire (
|
||||
<bibRefCitation id="9D99333060349A9E2C76D1043DDF0A2A" author="Mann and Garvey" firstAuthor="Mann" issue="3" journalOrPublisher="Coleopterist" pagination="138-142" refId="B31" refString="Mann DJ, Garvey L (2014) Aphodius (Liothorax) niger (Illiger) (Scarabaeidae: Aphodiinae) inside and outside the New Forest. Coleopterist 23 (3): 138–142." title="Aphodius (Liothorax) niger (Illiger) (Scarabaeidae: Aphodiinae) inside and outside the New Forest." volume="23" year="2014">Mann and Garvey 2014</bibRefCitation>
|
||||
). In the Swedish locality the beetles were burrowing in trampled organic mud (H. - E. Wanntorp, pers. comm.,
|
||||
<date id="666FA303C9EA507D6EAC8AFF3BA6BE13" value="2011-04">April 2011</date>
|
||||
) (Fig.
|
||||
<figureCitation id="8AAED65C8A8F11EBD0035402BA322F35" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 c</figureCitation>
|
||||
). Very occasionally
|
||||
<taxonomicName id="D0CBD20F5B17ED378534DA597502BF9A" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="E39696AD35ABBEB633BD0D167731D7CE" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
is taken in dung, as by J. Bergsten in the New Forest in 1999, and sometimes in dry areas of the Swedish island of Öland (H. Lundqvist, pers. comm.,
|
||||
<date id="10029805B3E4665A6C510A85BCC41FB6" value="2011-04">April 2011</date>
|
||||
). These are rare, random occurrences and may sometimes result from the beetle taking shelter from drought. One further feature of the occurrence of
|
||||
<taxonomicName id="25844280BC3D774588E82C1286712541" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="29670FC19070E7195736D719A47C6B1F" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
is worthy of comment – it shows dramatic fluctuations in abundance. Thus, in 2002 it was very abundant in its New Forest localities whereas more recently RBA has taken it only in low numbers. Similarly, H. - E. Wanntorp describes the species as being in very high numbers in the locality where he usually only took a few specimens (Fig.
|
||||
<figureCitation id="3D0E405B95D288BD176781F5C1878820" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
380
data/B1/B5/BB/B1B5BBB23CEA5A6DA84DB2710D8655A0.xml
Normal file
380
data/B1/B5/BB/B1B5BBB23CEA5A6DA84DB2710D8655A0.xml
Normal file
|
|
@ -0,0 +1,380 @@
|
|||
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<mods:mods id="85F2C76FCA5BD7A7B393479C1A9712C6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4410331FFB6BFF0D3990EF527F4A867D">
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<mods:title id="9ED7FCE451E5BF220AF4E6EA31DD3251">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:namePart id="983B2036F400D9F2DF0DAFDB9571E670">Angus, Robert B.</mods:namePart>
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||||
<mods:affiliation id="3D70AE1E0EE2EF7C7802650235E39453">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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</mods:name>
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<mods:name id="3B13BFB0257EFAF520B58B2A797AEF20" type="personal">
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</mods:role>
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<mods:namePart id="93F77504A0CB684F2DA95892CD52EA53">Maté, Jason F.</mods:namePart>
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<mods:nameIdentifier id="AF539358D2907DB8A29BCD3B32D36EFD" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
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<mods:affiliation id="55E21D41E7D74AA7065700C0DAA9E84B">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="DF0811DB592347E001E22EE3942B6CB5">Angus, Elizabeth M.</mods:namePart>
|
||||
<mods:affiliation id="586A3CAD463BF88F48C6C2AFBDE2BCE4">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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|
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</mods:role>
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<mods:namePart id="FEE10A2ED7744F2FFAB9F22972FAFB9A">Král, David</mods:namePart>
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<mods:nameIdentifier id="6ACAEC968D41F949BA18A4FB7E12DD75" type="ORCID">0000-0002-7930-1927</mods:nameIdentifier>
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<mods:affiliation id="8E48D08C4527F9B83D83BC3A52DFC685">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:typeOfResource id="1065C6FD9F559643498E4D4D83DEDAEC">text</mods:typeOfResource>
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<mods:title id="D8EE6C36829FDCE18A5B2846EEB054CF">ZooKeys</mods:title>
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<mods:part id="01D2CAE607E52C3BE2FA3D2DA274C7A8">
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<mods:date id="54D1278E39B968552FE192C2403B3B95">2024</mods:date>
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<mods:number id="CF4CC7C0C720CB5ED348492F18D1E92F">2024-07-22</mods:number>
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<mods:number id="E37BA74DBB872CAA98CD500AB4B509DB">1207</mods:number>
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<mods:classification id="B28DC31851CF7ECEA1ECDB9919D17410">journal article</mods:classification>
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<mods:identifier id="DE525BD0DDFA92338AAB4C8F29271C90" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<mods:identifier id="2FA7F29DE8F8635EDDEC7E958FDCCC8E" type="ZooBank">94F18819-5AF5-4100-AB35-AA3C3976EE80</mods:identifier>
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</mods:mods>
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<treatment id="B1B5BBB23CEA5A6DA84DB2710D8655A0" ID-arpha="B1B5BBB2-3CEA-5A6D-A84D-B2710D8655A0" ID-zoobank="https://zoobank.org/ABCB9508-F963-4C3D-ACDA-CEB4642AC551" LSID="urn:lsid:zoobank.org:act:ABCB9508-F963-4C3D-ACDA-CEB4642AC551" LSID-ZBK="urn:lsid:zoobank.org:act:ABCB9508-F963-4C3D-ACDA-CEB4642AC551" httpUri="http://treatment.plazi.org/id/B1B5BBB23CEA5A6DA84DB2710D8655A0">
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<subSubSection id="5385B384AED1B5ABD7B88743B1AA203F" type="nomenclature">
|
||||
<paragraph id="DC1120B40286F6450C588D6961992DEC">
|
||||
<taxonomicName id="1B5952DA83EAA4B8540F3606923B1683" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rodrigoi" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="771F21D46FF0208F3D6A4BA4E9542B22" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="FD5304D590B4BCE8CAB437C8D7C207DA" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="F655003013EDA5BEFC80332810EBA69E" italics="true">rodrigoi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7D18414CFEA4C1CBED72B3E2EAAD4E64" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="E041410F6F234BEF70C579C0CAD5EE19" type="description">
|
||||
<paragraph id="2F18E95703239B8362A9471C56763D13">
|
||||
<figureCitation id="DC611BECEF342B3D8BA19A0E1CE41929" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">Figs 2 d</figureCitation>
|
||||
,
|
||||
<figureCitation id="37661991BDBD078474404DF10F8643D1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">
|
||||
<quantity id="3135E59B1C5D6EF9057191153687EFF0" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" unit="g" value="7.0">7 g</quantity>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="467B7D81FA8D8761140A454361A4893B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure11" httpUri="https://binary.pensoft.net/fig/1097471">11 f</figureCitation>
|
||||
,
|
||||
<figureCitation id="844040A88E7C0BC392B364F182A7E4AA" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 j</figureCitation>
|
||||
,
|
||||
<figureCitation id="9379C7AC7D4022A5239C603CE1B65FDD" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">
|
||||
<quantity id="B134DA3D685C3E0505B74EFA6979EFC0" metricMagnitude="1" metricUnit="m" metricValue="2.2" unit="m" value="22.0">22 m</quantity>
|
||||
, m ’, m ”, 25 k
|
||||
</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ES1FK" type="type material">
|
||||
<paragraph id="59E922941FE33729E896EC389E229DED">
|
||||
<heading id="0B76EE2E0E1CAF7363DD3E468646ACB2" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B18DBB0D62DB6952B8643E47A3AD14E6">
|
||||
<materialsCitation id="B8AE761E4E715CA53376A85F1B7672C6" collectionCode="MNCN" collectorName="Aranjuez F. Moruder & C. M. Veiga" country="Spain" location="Madrid" specimenCount="1" specimenCount-male="1" stateProvince="Madrid" typeStatus="holotype">
|
||||
<emphasis id="0322BD78175813985539E668A71BD8FF" bold="true" italics="true">
|
||||
<typeStatus id="22EA0DCAAC3C4AD9D66025CC0F692A1A" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="C92DFDC535A14A9359380AE0F435A498" count="1" type="male">♂</specimenCount>
|
||||
:
|
||||
<collectingCountry id="3AAE767A58B963B76A94A21D5157D0E7" name="Spain">Spain</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="96952ABE99B5F50AC47AFD2D8DF10437" country="Spain" name="Madrid">Madrid</collectingRegion>
|
||||
<collectorName id="282D60B2270531FFB8F42D3EB6D0D393">Aranjuez F. Morüder</collectorName>
|
||||
//
|
||||
<taxonomicName id="609D329FCF7E55C8C343D181A4A65C7A" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="E8A54CFE0A36049BF4D2AE64E1350137" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
det
|
||||
<collectorName id="20367AB3E4209254313E87B1FAF2CC4E">C. M. Veiga</collectorName>
|
||||
1990 (
|
||||
<named-content id="39AB718D1408756F6EBBDE51E7BBE634" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales">
|
||||
<collectionCode id="65C5084AFF5602F40C60B0F063967EE6" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="31DD86998A15A3D3E039A646EBD6C6A8" bold="true" italics="true">
|
||||
<typeStatus id="F6E6A8B49E34D89D6FD81699EF37E325" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: 4 ex
|
||||
<specimenCount id="01EF37244DD7C6ED2B7674233E0B514F" count="1" type="female">♀</specimenCount>
|
||||
Aranjuez F. Morüder // DA
|
||||
<specimenCount id="44CA59D7347035998BEE88FAA4B23704" count="251" type="generic">251 F.</specimenCount>
|
||||
J. Cabrero // DG
|
||||
<specimenCount id="44CAA14A8E8F50A5A7698BBFF6C29C90" count="251" type="generic">251 F.</specimenCount>
|
||||
J. Cabrero // AP Bucal
|
||||
<specimenCount id="CD0A57E41DA6B9915A6932BB0495EC1F" count="206" type="generic">206 F.</specimenCount>
|
||||
J. Cabrero //
|
||||
<taxonomicName id="F9513B6B4F22E26C159E4B372CE05AA6" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="2CBD4A97BD6DF2A5694B523DF99C3541" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
det C. M. Veiga 1990; 1 ex;
|
||||
<specimenCount id="A7BDBAA42F1AB91A38457B4411EE7348" count="1" type="male">♂</specimenCount>
|
||||
Quero Prov Toledo Lauffer // DG
|
||||
<specimenCount id="07ADEE863AC263AA16B75B3DFB5AEED6" count="178" type="generic">178 F.</specimenCount>
|
||||
J. Cabrero // DA
|
||||
<specimenCount id="422376D418D259779F32E6048ADF9F29" count="249" type="generic">249 F.</specimenCount>
|
||||
J. Cabrero //
|
||||
<taxonomicName id="8D6083E8C5F691E535C55DA79DC14F8C" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="E39E7C2E3417CE68D4345334E13722D6" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
det C. M. Veiga 1990; 1 ex;
|
||||
<specimenCount id="257293AB7BF929FF85EA7F6DA8805DE5" count="1" type="female">♀</specimenCount>
|
||||
Quero
|
||||
<date id="F9AEB5B943081564C8BC7C7E7EFC9E27" value="1908-05">v. 1908</date>
|
||||
Molina // AP Bucal
|
||||
<specimenCount id="52F57CC95780EA37ADC9BF9E2D284CE5" count="56" type="generic">56 F.</specimenCount>
|
||||
J. Cabrero // DG
|
||||
<specimenCount id="EAA1992571100169A1F312CF10C885C5" count="174" type="generic">174 F.</specimenCount>
|
||||
J. Cabrero //
|
||||
<taxonomicName id="5FEA1D685381253A75B0975333606CBB" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="7D584545EDA5D0001F9143AB6CDEFD26" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
det C. M. Veiga 1990; 1 ex;
|
||||
<specimenCount id="F7A012C854EEDAFEE8D28C57B961C177" count="1" type="male">♂</specimenCount>
|
||||
Villacañas (
|
||||
<collectionCode id="6F80B71D0E9067FA5DBE9E4755E05866">TOLEDO</collectionCode>
|
||||
) C. Bolívar // Hoyer C. M. Veiga // AP Bucal
|
||||
<specimenCount id="C61E2626F3CBE2AC7C95AE6EACAFC215" count="205" type="generic">205 F.</specimenCount>
|
||||
J. Cabrero // DA
|
||||
<specimenCount id="34B0770621D39EEC9DD273B6AB39C5C1" count="250" type="generic">250 F.</specimenCount>
|
||||
J. Cabrero //
|
||||
<taxonomicName id="C8C822AA05F6FB948D4A3987257E4335" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="2B2FBA449241D1082E31BFC14D032624" italics="true">Aphodius plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
det C. M. Veiga 1990
|
||||
<specimenCount id="A5B9AF77EB737411AAB7065F8CDD8778" count="1" type="generic">1 ex.</specimenCount>
|
||||
(
|
||||
<named-content id="9F77F9B69489939E283770B0F97F01EC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales">
|
||||
<collectionCode id="93E89019AF7FDCB8DCB3B094B340360F" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E13FK" type="Differential diagnosis">
|
||||
<paragraph id="6520EF123F6A6F7A517BAA7F32982C40">
|
||||
<heading id="81DCC505AD870E2A87383410468F0378" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="70ABDAD8BD6C484AA8BC29B95332F789">
|
||||
<taxonomicName id="870E59C2A32356D6216B46E4FFDD30AB" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rodrigoi" subGenus="Liothorax">
|
||||
<emphasis id="F6D1B5C124D86E7106FFDF26E7B8F3E1" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="6949E7C246B0F30F86DEDEB599A2FC3B" italics="true">L.</emphasis>
|
||||
)
|
||||
<emphasis id="BA8EA39EA0DBC6A590F3FF99058FFE5F" italics="true">rodrigoi</emphasis>
|
||||
</taxonomicName>
|
||||
resembles a small highly-polished somewhat rounded
|
||||
<taxonomicName id="A49EB5000EC5FB29DD201A3902B22F34" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus" subGenus="Liothorax">
|
||||
<emphasis id="3267535D1D8857699B324CF71ADA0D1B" italics="true">A.</emphasis>
|
||||
(
|
||||
<emphasis id="F951D10B95AC3265E764C2443E444A12" italics="true">L.</emphasis>
|
||||
)
|
||||
<emphasis id="1671583B56655318674BDDDFBCEF4D69" italics="true">plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
. The metaventrite is slightly less strongly punctate and with the median diamond-shaped area depressed to mid-line. Aedeagus: (Fig.
|
||||
<figureCitation id="82F3CC5CF0AEE047BADB1B083D0CF01E" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">
|
||||
<quantity id="6C5FBF2802010D0AE5719EE7D4A7F82B" metricMagnitude="1" metricUnit="m" metricValue="2.2" unit="m" value="22.0">22 m</quantity>
|
||||
, m
|
||||
</figureCitation>
|
||||
’, m ”) general size and shape of parameres similar to
|
||||
<taxonomicName id="EF4C03BE34E4D50BB5A4F7F69381404D" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="2E996699B045F83FAA4631A21A0AB498" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
, not turned downwards apically, but with the following differences: apices of parameres less produced laterally, parameres narrower and more converging towards apex, in lateral view parameres gradually curved towards apex and narrowest towards apex compared to
|
||||
<taxonomicName id="459751B9CAAFB91AE1B66A3F8FDAF549" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="C7F37B18DDF2EB257ADF887FCF4F541A" italics="true">L. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
which is narrowest 1 / 3
|
||||
<superScript id="4FF678868513E26FE28111E808522085">rd</superScript>
|
||||
towards apex.
|
||||
</paragraph>
|
||||
<paragraph id="89AA4A8C70BC0B5AA6ECB63A063A7DB4">
|
||||
The apical segment of the maxillary palpi is slightly longer (10 %) than the 2
|
||||
<superScript id="5A18A0DB46BEE28952F7B1272365C021">nd</superScript>
|
||||
segment: terminal segment slightly longer (10 %) than 2
|
||||
<superScript id="A993319971107355F0C464A99BBF5022">nd</superScript>
|
||||
segment (Fig.
|
||||
<figureCitation id="8FD8026E9589A7AD01F0C56642E85A98" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure11" httpUri="https://binary.pensoft.net/fig/1097471">11 f</figureCitation>
|
||||
), as against 15–20 % longer in
|
||||
<taxonomicName id="ACA18A4A1E88EF93153AEB0338E3FB07" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="F09C7094C484CEC089D5DE6106D8AF12" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="03CF754B0EE6AA5E994BF8BEE49CEC20" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure11" httpUri="https://binary.pensoft.net/fig/1097471">11 d, e</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ER6FK" type="description">
|
||||
<paragraph id="54C12EF8EBAB317708B1C351A092B684">
|
||||
<heading id="5EF481081CE61A1CA1BAC658AB4BF8EC" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BD94D593EE34F91366AB26F845FE3645">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="CC204A9CD469E05A10C57D5CF54C7536" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 d</figureCitation>
|
||||
). Length: 3.5–4.0 mm (
|
||||
<specimenCount id="F2BD82768B9B5367B9D43D87BA6D6669" count="1" type="male">♂</specimenCount>
|
||||
) (HT
|
||||
<quantity id="3626AEE522002F29B0A9537CD79C931A" metricMagnitude="-3" metricUnit="m" metricValue="3.75" unit="mm" value="3.75">3.75 mm</quantity>
|
||||
), 4.0–
|
||||
<quantity id="C9F27369654FB775705AB57615C6C302" metricMagnitude="-3" metricUnit="m" metricValue="4.23" unit="mm" value="4.23">4.23 mm</quantity>
|
||||
(
|
||||
<specimenCount id="5DA6D71A30E6D15228567C397D47F002" count="1" type="female">♀</specimenCount>
|
||||
); width:
|
||||
<quantity id="3A27BD37D273A158E85963D7FF308EFC" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
|
||||
(
|
||||
<specimenCount id="8B927A478CB42000C7ABFBA1B44F05B7" count="1" type="male">♂</specimenCount>
|
||||
and
|
||||
<specimenCount id="F2A99710F19DB7E5F6CE9C0D544CE459" count="1" type="female">♀</specimenCount>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="491F949BFF3147CED392E7FD970DA5D4">Glossy black throughout except clypeal and elytral apical edges which are fuscous red. Legs and maxillary palps reddish yellow to maroon, antennae tan except for last three segments which are darker and clothed in off-white setae.</paragraph>
|
||||
<paragraph id="E079F309AA02F10A271B4914EC730F90">Head: frons strongly convex, clypeus flat to slightly concave. Frontoclypeal suture weak, widely interrupted in the middle. Clypeal edge truncated to slightly emarginate, impressed medially; side angles rounded with edge widely elevated. Sides of head almost straight and continuously merging with genae. Genae produced beyond eyes, rounded, strongly bordered and with some short yellowish setae.</paragraph>
|
||||
<paragraph id="443371449C3A4CF9DE5CC6A5C3D376AB">Surface shiny with residual reticulation and strongly punctured. Punctation double, the larger punctures regularly dispersed on the clypeal sides and frons, absent from genae and anterior clypeus. Larger punctures 2–3 × diameter of the smaller ones and strongly impressed, particularly on the sides where the surface almost appears wrinkled and closely spaced (1–1.5 × their diameter). Anterior edge of clypeus very finely and regularly punctured, completely devoid of larger punctures.</paragraph>
|
||||
<paragraph id="D77093798E4746BFE84F776A18CABEE0">
|
||||
Maxillary palp: terminal segment slightly longer (10 %) than 2
|
||||
<superScript id="461434AB3C29AA9A393E77922E46F765">nd</superScript>
|
||||
segment (Fig.
|
||||
<figureCitation id="6F903366A7FE2B446AB414C9C0CD1DF9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure11" httpUri="https://binary.pensoft.net/fig/1097471">11 f</figureCitation>
|
||||
) (in
|
||||
<taxonomicName id="D6423B395B6772C0CF8AA8775C8DD7BC" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="953DD611598CE1F28A49FE614FD6CDE2" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
the terminal segment is longer, 15–20 % longer than the 2
|
||||
<superScript id="689C739A670957AB2BD4310892071685">nd</superScript>
|
||||
segment) (Fig.
|
||||
<figureCitation id="A350BEFC5BEFC4FE5B5039FE59D09859" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure11" httpUri="https://binary.pensoft.net/fig/1097471">11 d, e</figureCitation>
|
||||
). Galeal patch armed with five or six strong galeal chaetae (
|
||||
<quantity id="C7CE559065023CB3FB5FCBB386392DF0" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7–9 in</quantity>
|
||||
<taxonomicName id="0C8CC057C6278664E8B8191038018CFA" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="70FBC471F7EC223431304AFC56CBB45C" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
). Galea smaller, similar in dimensions to the Tibetan ssp. of
|
||||
<taxonomicName id="0DCB9501BDD7A7106C6DDA2F8EB7C66D" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="B2EA40F763543F3F74BFB9C5E7733601" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
but not to European populations in which it is larger.
|
||||
</paragraph>
|
||||
<paragraph id="7192D43331DE27927177A5816122D74C">
|
||||
Epipharynx (Fig.
|
||||
<figureCitation id="3B32FAAE72242AB59EDE723F7A9749D3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">
|
||||
<quantity id="6E15C5D8126F3F030E91A356CF6633F7" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" unit="g" value="7.0">7 g</quantity>
|
||||
</figureCitation>
|
||||
) corypha small and only slightly produced (somewhat similar to the Tibetan populations of
|
||||
<taxonomicName id="E4E0B991252F3A4A869CF73546141317" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="A391DD15474829B6BE9C7DEBE9B27985" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
), with the chaetae (celtes) much reduced in size. Fenestrae on zygum (angustofenestrae) and chaetae (heli) less numerous than in
|
||||
<taxonomicName id="8B567ECDA665C435FFC45341C0F8DB53" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="97ACEDC49C03BDED4A19A89E3B22E165" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
and limited to the edge against pedia. Chaetopaedia as in
|
||||
<taxonomicName id="128A0497EDDFF63BE58DACBBF0C828DF" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="6F7AD68B69174446712364D119D4E637" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
. Chaetopariae noticeably stronger than in
|
||||
<taxonomicName id="5D90C03EDFD87C71CF2E25B17FE2C318" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="E6F139EAEBE309B5E49A5E5A685E8CC2" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="764F4D5C6009A8E23E862E6722A327A7">Pronotum subquadrate, sides slightly rounded, subparallel, and widest towards base. Regularly convex longitudinally and transversely. Base completely and distinctly bordered, border strong but fine. Anterior edge not bordered at all. Lateral margins visible dorsally in the apical 1 / 3 only. Sides strongly bordered throughout, anteriorly going around corner up to the middle of the eye. Lateral margins with short yellowish hairs barely visible dorsally in apical 1 / 2.</paragraph>
|
||||
<paragraph id="189027017C758A1A2DCBF14A57FB08F7">Surface of pronotum black and shiny, not at all alutaceous with only some residual superficial shagreen. Punctation double, larger punctures regularly distributed throughout but densest on the sides and anterolateral corners. Large punctures flat bottomed and umbilicated. Diameter 4–5 × that of the smaller ones and spaced 1–3 × their diameter.</paragraph>
|
||||
<paragraph id="BDD6BEDF45514CEABA6AF9B127DC4B4D">Scutellum elongate, sides rounded, convergent throughout (only slightly pentagonal), length 1 / 11 elytral length. Glossy black, unpunctured, basally impressed, disk convex.</paragraph>
|
||||
<paragraph id="CD6F3F716ECE465B6CF22E3BFEC35325">Elytra black, intervals convex, surface strongly glossy and slightly alutaceous under high magnification. Intervals with double row of very fine and faint punctures. Striae fine, 1 / 8 × width of intervals, with sides crisp and right-angled. Regularly punctured, punctures wider than the striae and separated by 2 × diameter (first 2 striae) or 3–4 × (remainder). Elytral epipleura strong, gradually convergent towards apex, and at humeri forming a small but distinct tooth.</paragraph>
|
||||
<paragraph id="5BD0D5D21371508855789EA323D0E7ED">Underside black with yellowish pubescence on the abdomen.</paragraph>
|
||||
<paragraph id="148D97758E573F070C7584DBAFC508EB">
|
||||
Metaventrite (Fig.
|
||||
<figureCitation id="95CC17440156EC30ECF23537A76D4067" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 j</figureCitation>
|
||||
): median diamond-shaped area rather strongly punctate, glabrous, its mid-line distinctly impressed. Surface regularly heavily punctate, the punctures with one end somewhat pointed, shiny, and not alutaceous. Sexual dimorphism subtle, female flatter than male.
|
||||
</paragraph>
|
||||
<paragraph id="37898C52ECC2C0D1E2DE31B3BEA53CEE">Legs reddish to dark brown, tarsi and tibial spurs red to tan, rather long and slender. Protarsal spur regularly acuminate and curved in both sexes, reaching to apex of second tarsal segment. Metatarsal segments short. Basotarsomere slightly shorter than upper metatibial spine as long as segments 2 + 3. Fimbrial setae short and of unequal length. Longer spur of mid tibiae as long as first two tarsal segments.</paragraph>
|
||||
<paragraph id="37BA7D9D85C732EDCA293AF9A12C8D34">
|
||||
Aedeagus: (Fig.
|
||||
<figureCitation id="6DE4514E19B36A65BAF16A8AAE20C3F4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">
|
||||
<quantity id="37A12A671C7CBEEDBCB5236344C17D6E" metricMagnitude="1" metricUnit="m" metricValue="2.2" unit="m" value="22.0">22 m</quantity>
|
||||
, m
|
||||
</figureCitation>
|
||||
’, m ”) general size and shape of parameres similar to
|
||||
<taxonomicName id="B34106A11EA1E843AA649E8EB20BCCFA" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="0E6E5E8547DF0CBD18296D724A837907" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
, not turned downwards apically, but with the following differences: apices of parameres less produced laterally, parameres narrower and more convergent towards apex, in lateral view parameres gradually curved towards apex and narrowest nearer apex (ca ¼ of the paramere length as against ca 1 /
|
||||
<quantity id="10D9710D33EF0D3CC202AAC96AD5646A" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
|
||||
<taxonomicName id="41DC0B059A5B734CC1C62A51D7C2E6D2" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="BCDB1D5C7396532A87D9CC5AB8028B74" italics="true">L. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="3FAD34640553755B899CDC67DBC35AFB">
|
||||
Endophallus typical of other members of the
|
||||
<taxonomicName id="C7B77BAEFE806FFEB9BC6FF18846D0CB" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="744111CAE4EDA6B93B1F49FFF1218C4F" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
species group, with two patches of elongate triangular scales or bristles. Those in basal patch longer and more elongate than any member of the
|
||||
<taxonomicName id="22C9374F5A07C03C5FA2446EC0C42CE2" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="31A04875637B3FE1DF62F09573DE0DF3" italics="true">plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
group (19–26 µm in
|
||||
<taxonomicName id="0F2EE07D4B5189F94F99AC3956BB24CF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rodrigoi">
|
||||
<emphasis id="67BCCDB76F503AD15199B470F2DBA317" italics="true">A. rodrigoi</emphasis>
|
||||
</taxonomicName>
|
||||
vs 14 – 21 µm), similar to those of
|
||||
<taxonomicName id="8C8B4658C6CB8AFEB530E6841F1DAE66" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae">
|
||||
<emphasis id="B33B1673A1C80ACB2A4DDC96D5EA214A" italics="true">A. wilsonae</emphasis>
|
||||
</taxonomicName>
|
||||
(22–32 µm), those in the apical patch like
|
||||
<taxonomicName id="8449A37EB2F81BDC3A9945628191D3C8" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="73058F2050CFDDC0197FBB82A15A6297" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6FGK" type="etymology">
|
||||
<paragraph id="46DDC995833154A2A250DA137768C202">
|
||||
<heading id="5FB1F9AA07EB45552C537AF99FECCF1D" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="12EF449AD568FA99AFAEA38BD7D47F38">The name is derived from the surname of the name Joaquín Rodrigo Vidre, a Spanish composer whose most famous work, “ Concierto de Aranjuez ”, refers to the geographical area from which the species hails.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEGGK" type="remarks">
|
||||
<paragraph id="59855D54C0E97BB8FB35ED65B132C528">
|
||||
<heading id="F7934A4242EF3D82F0189C7A23E83838" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F4801F30A28584DCD993BBB2BE17F4F4">
|
||||
All the known specimens are from the south of
|
||||
<collectingRegion id="7BB668D3511883D6DD89616C460A9FFE" country="Spain" name="Madrid">Madrid</collectingRegion>
|
||||
(Aranjuez) and the adjacent areas of Toledo (Fig.
|
||||
<figureCitation id="DA0AEFFEDDC01A38EBE5229B8B65018E" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b</figureCitation>
|
||||
) The localities are associated with temporary endorheic saline lagoons and lakes (Fig.
|
||||
<figureCitation id="48391D11FF56D8AA590C051E270215CA" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 b</figureCitation>
|
||||
) One of the authors (
|
||||
<collectionCode id="745E618BDE7D28DCB5022F653A7535EA">JFM</collectionCode>
|
||||
) made repeated visits to several of the localities during late winter over several years (January and April of 2015–2017), but no specimens were found. Although it is possible that the species was missed due to its ecology or time of emergence, it is also possible that the species is very rare or has been extirpated from much of its former distribution. The known localities are part of an extensive system of aquifer-fed saline lagoons encompassing much of La Mancha and collectively known as “ La Mancha Húmeda ” (Wet La Mancha). Many of these lagoons are degraded or lost to agriculture but almost 1 / 2, including several visited for this study, are protected and in apparently good ecological condition (
|
||||
<bibRefCitation id="8422A49DDB0EA97508C0DA85ED49387D" DOI="10.13140/2.1.4445.3764" author="Florin-Beltran" editor="García E" firstAuthor="Florin-Beltran" journalOrPublisher="Instituto Provincial de Investigaciones y Estudios Toledanos, Diputación Provincial de Toledo, Toledo" pagination="65-92" refId="B18" refString="Florín-Beltrán M (2001) La Mancha Húmeda: Valores ecológicos y problemática conservacionista. In: García E (Ed.) Ecología y Conservación de la Mancha Húmeda. Instituto Provincial de Investigaciones y Estudios Toledanos, Diputación Provincial de Toledo, Toledo, 65–92. https://doi.org/10.13140/2.1.4445.3764" title="La Mancha Húmeda: Valores ecológicos y problemática conservacionista." volumeTitle="Ecología y Conservación de la Mancha Húmeda." year="2001">Florín-Beltrán 2001</bibRefCitation>
|
||||
). Hence, it would be hasty to declare this interesting Iberian endemic extinct without carrying out extensive sampling across the area and in particular within the protected lagoons which might still host it.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
389
data/C8/50/01/C85001FB61315D4FBAF4B413BEB88D37.xml
Normal file
389
data/C8/50/01/C85001FB61315D4FBAF4B413BEB88D37.xml
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|
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|
||||
<paragraph id="39A6D143C3DD16D164EA2441F0FE701C">
|
||||
<taxonomicName id="8EE300AE0D0BD973E6EFD375C5F10971" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="BAEC7452CCB7C9502443BFAEB38E248A" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="09427E5CCB2719AD2BF681BAE03120C2" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="0A4D5608D00F606B0843C56ACEA5C2F8" italics="true">bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="F9BCE3AE76ACDBDC71D150CB7F60E582" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="432C79F235C6A4E17ABC227586F484B2" type="description">
|
||||
<paragraph id="A8C8052DC7EEDD928E56729365352CA7">
|
||||
<figureCitation id="BEAAA872B3001CD405892E66515F61AB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 f</figureCitation>
|
||||
,
|
||||
<figureCitation id="A7A14D3B0552400EEFFD72CC3DEEC590" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 j</figureCitation>
|
||||
,
|
||||
<figureCitation id="6F9AC1E870B30A360DDBC72C3F4FD5AD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 f, g</figureCitation>
|
||||
,
|
||||
<figureCitation id="C70C0CB392C8B4AE417390EE0BB603A1" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 d</figureCitation>
|
||||
,
|
||||
<figureCitation id="1B9FB4662EF7FB09058596A6F4DFC516" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 u</figureCitation>
|
||||
,
|
||||
<figureCitation id="A027A9067D1DCC980A2EC863C21473F6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 r</figureCitation>
|
||||
,
|
||||
<figureCitation id="6B14B33BBF8D533C88975BBEB18B5FA5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 l, m, p – t</figureCitation>
|
||||
,
|
||||
<figureCitation id="E7AB88BC26D387045C2499543D6F2E9F" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">
|
||||
<quantity id="73CAB3EEF85AA44615090AC48C854EBE" metricMagnitude="1" metricUnit="m" metricValue="2.4" unit="m" value="24.0">24 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="33836FAE71639DC2604285C5DAC8FB59" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure25" httpUri="https://binary.pensoft.net/fig/1097496">25 e</figureCitation>
|
||||
,
|
||||
<figureCitation id="2B127843399A765C9374240239BD5AF9" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 z</figureCitation>
|
||||
,
|
||||
<figureCitation id="A76B37E2FD63D2AC3781344A13F4CE4A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure32" httpUri="https://binary.pensoft.net/fig/1097519">32 f – i</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMWBM" type="type material">
|
||||
<paragraph id="DC70A84FFFA3D90FDCEBDE2FF7B9B4FB">
|
||||
<heading id="496C2D6C985578C780F00D8C1E322E48" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="70CCFBA9EF93536A2C4E785A1E015A0C">
|
||||
<materialsCitation id="E08A46708E92AD7AC95CD9A636396189" collectingDate="2013-05-01" collectingDateMax="2013-05-08" collectingDateMin="2013-05-01" collectionCode="NHMUK" county="Parco delle Madonie. Piano Battaglia. At" location="Length" municipality="Chromosome" specimenCount="1" specimenCount-male="1" stateProvince="Sicilia" typeStatus="holotype">
|
||||
<emphasis id="7BBA742FD57259764C518BADF4BB8540" bold="true" italics="true">
|
||||
<typeStatus id="9417278462E4CE370173600CBB0F21B2" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="2B3DE2CC4304DC2FE0FFBAB8468AB3F7" count="1" type="male">♂</specimenCount>
|
||||
:
|
||||
<collectingRegion id="01DD026644A4FFE78F22F415B31C2378" country="Italy" name="Sicilia">Sicily</collectingRegion>
|
||||
.
|
||||
<collectingCounty id="1B8E128862A17496B63BDAF87182F969">Parco delle Madonie. Piano Battaglia. At</collectingCounty>
|
||||
edges of pool.
|
||||
<date id="46188ABD1EF5A5F0BFCD6666D0499012" value="2013-05-01">
|
||||
<collectingDate id="571F3AE25CD2684810D771C7AD7459FE" value="2013-05-01">1. v. 2013</collectingDate>
|
||||
</date>
|
||||
. R. B. & E. M.
|
||||
<collectingRegion id="5B794D8234B6F2214A985F549003F784" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<collectingMunicipality id="8C437AE6C838957C63FB9FFF89010CA4">Chromosome</collectingMunicipality>
|
||||
prep. 7:
|
||||
<date id="CC06E0C0EBD4D1531B5E6DAF50E69AEC" value="2013-05-08">
|
||||
<collectingDate id="924F938686BF0C85AEE2C6AC4CB786C2" value="2013-05-08">8. v. 2013</collectingDate>
|
||||
</date>
|
||||
(
|
||||
<named-content id="E48C89D6012DF7B70192DAFD4805E938" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="52115F1FA3E23477FACAF2BA38476719" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<location id="681D15CB41CD59616C2A1F5DF41AD19B" LSID="urn:lsid:plazi:treatment:C85001FB61315D4FBAF4B413BEB88D37:681D15CB41CD59616C2A1F5DF41AD19B" county="Parco delle Madonie. Piano Battaglia. At" municipality="Chromosome" name="Length" stateProvince="Sicilia">Length</location>
|
||||
<quantity id="DEFF1337DAEF1CD90C21DCBDDF6BB05F" metricMagnitude="-3" metricUnit="m" metricValue="3.9" unit="mm" value="3.9">3.9 mm</quantity>
|
||||
, width
|
||||
<quantity id="6EC832E8CE050CAE018E5C366B94ED4A" metricMagnitude="-3" metricUnit="m" metricValue="1.8" unit="mm" value="1.8">1.8 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="81685B0D9AB6FEF7EAF71D1D6DBFE926" bold="true" italics="true">
|
||||
<typeStatus id="8049777A5218059105E34EC974AE81BB" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="C0487373FD2E37AD5212B02450C4A1A7" count="7" type="male">7 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="C5A0D6CC060F460A92CAAB290FDBBB83" count="8" type="female">8 ♀♀</specimenCount>
|
||||
, same data as
|
||||
<typeStatus id="BD7716BB59BB840C23D7E65195730FE4" type="holotype">holotype</typeStatus>
|
||||
. Chromosome preparations 1–8,
|
||||
<date id="2A208068EB06E98F227E24D0810A5965" value="2012-05-07">7. v. 2012</date>
|
||||
, 1–6 and 8,
|
||||
<date id="64124BF3169D1FA890467BE3BA8BE4F8" value="2012-05-08">8. v. 2012</date>
|
||||
(
|
||||
<named-content id="9A171C189A0769DF28EA237D5E50F087" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="623238A2C08E8A4C713C63BD7A38A16A" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<specimenCount id="8FFBDDAB3FA2B56F3F5128A3FDA78277" count="7" type="male">7 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="23D5C79232C389099B04C516041E1541" count="5" type="female">5 ♀♀</specimenCount>
|
||||
, same data as
|
||||
<typeStatus id="E9411F74509E9A4D9AB031FC3CB64623" type="holotype">holotype</typeStatus>
|
||||
but
|
||||
<date id="859C907D934A4B0F23117F7E7A4B2544" value="2018-04-23">23. iv. 2018</date>
|
||||
(
|
||||
<named-content id="B5A3B5FF71199EE8A199D4B0A0F484C1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="D2EB0392DA2D314E1650D7B03C114650" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<specimenCount id="64730093F714D3AD5FDEBDE6DCB5C581" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="E26D1307F6897E6680783AD8B98BA7FE" count="1" type="female">1 ♀</specimenCount>
|
||||
, same data (
|
||||
<named-content id="A7C9CE4EB99ED6147D01D6FA1464990F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="04BE62B58EA8269DE46C8B385587D0AA" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="46F9D22015FF0A9F4AAC6732A7314D0B" count="12" type="male">12 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="4226C80FCAEC47830E4EA1B7C73FDA4E" count="6" type="female">6 ♀♀</specimenCount>
|
||||
,
|
||||
<collectingRegion id="55447F5A86BF2163A178A6081FBCF8D9" country="Italy" name="Sicilia">Sicily</collectingRegion>
|
||||
, Nebrodi, Monte Soro,
|
||||
<quantity id="AA6A7F48C99EB80FC058A5413654370D" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</quantity>
|
||||
a. s. l.
|
||||
<date id="F62144BF21CC3C2BD027C9D9B79FAE37" value="2018-04-25">25. iv. 2018</date>
|
||||
. R. B. & E. M.
|
||||
<collectingRegion id="23AC9705924D58A93F72DA2EEF8A3999" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
. Chromosome preparations 1–4,
|
||||
<date id="D56C5C7400272848209AB743728DDCDB" value="2018-04-27">27. iv. 2018</date>
|
||||
, 1–5,
|
||||
<date id="67FCA8772D74BFBDA7F769DE69BA7EC3" value="2018-05-01">1. v. 2018</date>
|
||||
, 1–5,
|
||||
<date id="D83B1E38A07EAA0DF59C959B66F1FA3B" value="2018-05-02">2. v. 2018</date>
|
||||
, 1–4,
|
||||
<date id="C3DE9F45D1451D2F9531C5D7A5084403" value="2015-05-10">10. v. 2015</date>
|
||||
(
|
||||
<named-content id="3A921F931BE84A81B7BDC0A378D0EFEB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="1C0A202894EFF285B54BAAE912BB5970" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<specimenCount id="9A035F10A4BEB4F044A2AC6D1C7A4D43" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="89E894C2044F7C90D94C5B78B6DC8D70" country="Italy" name="Sicilia">Sicily</collectingRegion>
|
||||
, Nebrodi, leg F. Krell (
|
||||
<named-content id="A00E9976C42246191B49B27646CD8650" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/denver-museum-nature-science" xlink_title="Denver Museum of Nature and Science">
|
||||
<collectionCode id="9AB1F7287C27AF08B791B8DCA32447D0" country="USA" httpUri="http://grbio.org/cool/mmzs-jqix" name="Denver Museum of Nature and Science" type="Museum">DMNS</collectionCode>
|
||||
</named-content>
|
||||
). 3, data as
|
||||
<typeStatus id="8240067360602526F20EC4F7499B0519" type="holotype">holotype</typeStatus>
|
||||
(sequenced) (
|
||||
<abbrev id="ABBRID0ESXBM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="70AEFB2CD8332FA9BEFE3D87E3235627">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
); 1
|
||||
<collectingRegion id="6DA1F8F89871ECE122EB99E0B50EE236" country="Italy" name="Sicilia">SICILY</collectingRegion>
|
||||
, Nebrodi, Monte Soro,
|
||||
<quantity id="34FF4F014225F6EEAE7571C71D909CB9" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</quantity>
|
||||
a. s. l.
|
||||
<date id="40228545EE4F14C4FCD6179A5D2BA33A" value="2018-04-25">25. iv. 2018</date>
|
||||
. R. B. & E. M.
|
||||
<collectingRegion id="D44F3734170A84A490CB1515EC4F2B49" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
. Chromosome preparation 4,
|
||||
<date id="99044DDB4EE992A1915A9F8EBEB2710B" value="2018-04-27">27. iv. 2018</date>
|
||||
(Sequenced) (
|
||||
<abbrev id="ABBRID0EWXBM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="EF67E4C7C50555A536890FB0E1CD827E">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="0463D7A18711F457D5BFDC541F013EA8">
|
||||
<specimenCount id="3F48E620E049E281E5571A02484798B0" count="3" type="male">3 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="B881718B45088A27DF7D202557E8DBBB" count="4" type="female">4 ♀♀</specimenCount>
|
||||
, 37 unsexed:
|
||||
<collectingCountry id="6A18A4C9B7666E66A5C94B1B87C20B1C" name="Italy">ITALY</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="0C366A389AC48A636CEFF17EF62CA35A" country="Italy" name="Sicilia">Sicily</collectingRegion>
|
||||
, Monte Soro, Tumpelufer, welkes Laub.
|
||||
<quantity id="ED1623708802CC9A01D3E00C8E7770CC" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</quantity>
|
||||
NN
|
||||
<date id="392ACC3D39B3A75796159F388E776E24" value="1993-05-30">30. 5. 1993</date>
|
||||
leg. U Schaffrath (
|
||||
<abbrev id="ABBRID0E3XBM" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="7A562CFDCE1821402E674FDBE508E443">ABC</collectionCode>
|
||||
</abbrev>
|
||||
&
|
||||
<abbrev id="ABBRID0EAYBM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="68B60527ED91AE86A2820564C84265F1">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EEYBM" type="Differential diagnosis">
|
||||
<paragraph id="F15D3B1F04C777B3411CBD27E1B5F4C7">
|
||||
<heading id="C08BB7A2AEA4A4EC0FAD8931C8253F97" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="3545DA4D208249D6D480102D661832CB">
|
||||
Most similar to
|
||||
<taxonomicName id="CC064332337EC58193B8BF9F8E67246A" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="89861B477CD3E0CD5D5A63A4475FB9FC" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
, from which it is shown to be different because of its clearly larger X chromosome (Fig.
|
||||
<figureCitation id="A9006F3EC4680B49ABEF5B87AA78FBEF" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure32" httpUri="https://binary.pensoft.net/fig/1097519">32 f – i, a – e</figureCitation>
|
||||
). The slightly smaller aedeagus, with the endophallic tooth-field clearly shorter than the parameres (Figs
|
||||
<figureCitation id="92F0F55921C42C3B65F13F44CEAF0B71" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 p – t</figureCitation>
|
||||
,
|
||||
<figureCitation id="33C84B46B54808690F1E0C08D656ACEC" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">
|
||||
<quantity id="AB73F06E9394F60659D14E2B7689A896" metricMagnitude="1" metricUnit="m" metricValue="2.4" unit="m" value="24.0">24 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
) is also distinctive.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBZBM" type="description">
|
||||
<paragraph id="844ADD4190500D7C07506242D02A9A1D">
|
||||
<heading id="FDBA3D7BE3D75F4DAA4BE0EDB7C9D2BA" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="45B9E7805DF26D0CF4E75CE68264C43B">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="4B56E50032E6BAC51059E65472D6226F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 f</figureCitation>
|
||||
). Length:
|
||||
<quantity id="FF1FAF0C76F124A554B3E32775C9EA14" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.4" metricValueMin="3.6" unit="mm" value="4.0" valueMax="4.4" valueMin="3.6">3.6–4.4 mm</quantity>
|
||||
(
|
||||
<specimenCount id="B47BEB33A3D65B0BD5E3881A0652E9AA" count="1" type="male">♂</specimenCount>
|
||||
), 4.0–
|
||||
<quantity id="D8D5BEB12728E3874D30D9730BECA157" metricMagnitude="-3" metricUnit="m" metricValue="4.8" unit="mm" value="4.8">4.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="7F4086D6A037B61A998AD291ACF503E6" count="1" type="female">♀</specimenCount>
|
||||
), width:
|
||||
<quantity id="898B344F191282D0B8362D9C479A94B1" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.95" metricValueMin="1.65" unit="mm" value="1.7999999999999998" valueMax="1.95" valueMin="1.65">1.65–1.95 mm</quantity>
|
||||
(
|
||||
<specimenCount id="178BC854EB7F4B15E4EC285592D45837" count="1" type="male">♂</specimenCount>
|
||||
), 1.7–2.0 mm (
|
||||
<specimenCount id="CDEC9303BC04992532B5EB90B7A48799" count="1" type="female">♀</specimenCount>
|
||||
). Glossy black, apical segments plus claws of tarsi dark brown.
|
||||
</paragraph>
|
||||
<paragraph id="57D2BD27DEA0B62E11CDBAE2E8C182FA">
|
||||
Head with no trace of reticulation but with sparse double punctures, the punctures separated by ca 2 × the diameter of the larger punctures. Frontoclypeal suture indistinct but present, without tubercles but slightly raised at ends of median section, before turning to run to the anterior margins of the genae. Anterior margin of clypeus emarginated medially, rounded at sides of the emargination. Antennae and palpi black. Epipharynx (Fig.
|
||||
<figureCitation id="0F8AEE15B70981F2A42F2294B0DA6D32" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 f, g</figureCitation>
|
||||
) with anterior margin of clithra emarginate each side of projecting median tylus, with a few fine acropariae. Chaetopedia 4 or 5 each side, stout. Chaetopariae forming a single line each side, closely set, stout. Apophobae forming a narrow band of fine setae outside the chaetopariae, prophobae in a narrow field each side of the mesoepitorma. Head and pronotum (Fig.
|
||||
<figureCitation id="BA9FF2121FBCD1E30AE7768A0049C2BC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 j</figureCitation>
|
||||
): pronotum: surface either smooth and glossy as head or slightly dull, with double punctation, size of the large punctures stronger than on head, but variable, these punctures separated by ca 2 × their diameter. Small punctures tiny dots. Mid-line impunctate or with very fine punctures. Pronotum more or less hemicylindrical, highly arched transversely, in mid-line weakly arched longitudinally. Sides of pronotum bulging outwards in basal 1 / 3, so that the basal 1 / 3 of the lateral margins is not visible from above. Lateral margins completely bordered, this border extending along lateral 1 / 3 of basal margin as a very fine impressed line. Median 1 / 3 of basal margin without an impressed border, as in the
|
||||
<taxonomicName id="7F45E86FF7175FB8A08366D5172918C7" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae">
|
||||
<emphasis id="B7BD64FE2ADED0CF0CFB754462DC24EA" italics="true">A. wilsonae</emphasis>
|
||||
</taxonomicName>
|
||||
shown in Fig.
|
||||
<figureCitation id="719548E24193C9F265C14E7C8AAD30DE" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 d</figureCitation>
|
||||
. Scutellum pentagonal, sides parallel in basal 1 / 2, then converging to blunt apical point. Elytra: sides either almost parallel over basal 2 / 3 or more distinctly rounded to widest point 2 / 3 of the way from the base, bluntly rounded to apex. Interstices flat, ca 6 × the width of striae, glossy black, sometimes with very fine reticulation, especially over basal 1 / 4 (Fig.
|
||||
<figureCitation id="ECCB4DA2FC5E4D747B9CAD7B898AB7FB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 u</figureCitation>
|
||||
). Sutural interstices weakly raised to suture. Striae narrow, shallow, the strial punctures scarcely impinging on their margins. Median diamond-shaped plate of metaventrite with sparse fairly fine punctures, sometimes flattened in males or with a median depression in females (Fig.
|
||||
<figureCitation id="9676EA38057E90641CDAF1D194659916" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 l, m</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="F1E89886013AE066F1BED6ED879D6524">
|
||||
Legs: Long spur of mid tibiae clearly longer than basal segment of tarsi, exceeding it by a distance approximately equal to the width of the basal margin of the second tarsal segment (Fig.
|
||||
<figureCitation id="CE6C5FB72D21882DBFDACBC9B9339C62" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 r</figureCitation>
|
||||
). Spines on apical margin of posterior tarsi short and even on outer / ventral face, longer, sparser, and more irregular on inner / dorsal face.
|
||||
</paragraph>
|
||||
<paragraph id="3C54182318F93D302CC42485408D10F8">
|
||||
Aedeagus (Figs
|
||||
<figureCitation id="BBAFC5B0718E0E758FE8B359505BED88" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 p – t</figureCitation>
|
||||
,
|
||||
<figureCitation id="DA4D03F625049D2D8C15637F43C323E1" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure24" httpUri="https://binary.pensoft.net/fig/1097494">
|
||||
<quantity id="2F175D9485E57FCD0F59E637911C9699" metricMagnitude="1" metricUnit="m" metricValue="2.4" unit="m" value="24.0">24 m</quantity>
|
||||
, n
|
||||
</figureCitation>
|
||||
) Length ca
|
||||
<quantity id="D26123A00F56CA564E8DC3BE92D8B2C6" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
|
||||
, paramere length ca
|
||||
<quantity id="270FDC45E9D37239CFE3FA78D4A81484" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43–0.47 mm</quantity>
|
||||
, basal piece length ca
|
||||
<quantity id="B4BA374DD0042B6B4AF1066113112632" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
|
||||
, tooth field length ca
|
||||
<quantity id="38592FB64459A8E7ACAE951FFCE5F6B9" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
|
||||
. This is one of the few species in which the length of the endophallic tooth-field is less than that of the parameres. Longest teeth on endophallus ca 47 μm long.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E31BM" type="etymology">
|
||||
<paragraph id="F8F1CF82FBEABBA4B2BD41A84B27C144">
|
||||
<heading id="EBE4313E3274B34B5F98C975DCDF3E4C" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E82023A50607D9826D6433AFE9A140D4">
|
||||
<taxonomicName id="A3066E71CE44DB4DEE2191F09168331D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens">
|
||||
<emphasis id="AA577DA42E66ADED4AF2A95E2EB8D8A6" italics="true">bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
– Latin, waging war, adjective, named for the
|
||||
<typeStatus id="DABEF0879C4E1343581748E392C5F880">type</typeStatus>
|
||||
locality, Piano Battaglia (the battlefield).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EM2BM" type="remarks">
|
||||
<paragraph id="415D1AD5D310FE2C39F3DB0891B540F8">
|
||||
<heading id="BD3FE0DCD2C283F0D9E75C3882E810A5" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="89489D87E4B45E6AB8282ADBAE6F02DE">
|
||||
So far known only from
|
||||
<collectingRegion id="47F044D9F84C480C07F2023641979228" country="Italy" name="Sicilia">Sicily</collectingRegion>
|
||||
(Fig.
|
||||
<figureCitation id="3D03B8AE96896A5A2C1B53633C566430" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
). Many of the
|
||||
<typeStatus id="BC7EEFF6613ACD8968ABA3E72C8A457C">types</typeStatus>
|
||||
were taken by swishing the water at the edges of the pool on the Piano Battaglia, washing the beetles from the banks so that they floated on the water. This pool dries out completely in the summer. The Monte Soro specimens were floating and clinging to plants in a shallow pool, following recent rain (Fig.
|
||||
<figureCitation id="919A61DB8380AEDBBFC04D72F401CC4D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">
|
||||
<quantity id="24CC5644593ABD234CFF37C7E1431419" metricMagnitude="-2" metricUnit="kg" metricValue="2.7" unit="g" value="27.0">27 g</quantity>
|
||||
</figureCitation>
|
||||
). We have seen specimens from
|
||||
<collectingRegion id="990EAD83AAB701486E1FFA7EF5B3278B" country="Italy" name="Calabria">Calabria</collectingRegion>
|
||||
(Sta Christina, Cippo di Garibaldi) which are apparently morphologically identical but have not been included in the
|
||||
<typeStatus id="A01961F843180F1922611F0A999B9F7C">type</typeStatus>
|
||||
series due to the scarcity of material and lack of karyological or molecular data. RBA has recently received living material from
|
||||
<collectingRegion id="95FC303B6A2082D036D4FDC407C48960" country="Italy" name="Calabria">Calabria</collectingRegion>
|
||||
: Villaggio Mancusa,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E42BM" specific-use="{"type":"Point","coordinates":[16.640850,39.107270]}">
|
||||
<geoCoordinate id="451F97B239513F6B3D2889F95A6C9DEB" degrees="39.10727" direction="north" orientation="latitude" precision="1" value="39.10727">39.10727 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="F9A5684CF17BAE0FA781056183C9623B" degrees="16.64085" direction="east" orientation="longitude" precision="1" value="16.64085">16.64085 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, from which chromosomes indicating
|
||||
<taxonomicName id="97537084ACFB9D0FBA7CE4789B09B0EF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="F3D63C0111516AF3A3387347B70B82D5" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
, not
|
||||
<taxonomicName id="C386355B956BF8A40E64F72CFCBD0053" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens">
|
||||
<emphasis id="2DDB1B651E7C4401B471B4D6E715ED70" italics="true">A. bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
have been obtained. These data will be published in due course.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
554
data/D4/E3/DD/D4E3DDD826E95A6C969E6CC03CAF933F.xml
Normal file
554
data/D4/E3/DD/D4E3DDD826E95A6C969E6CC03CAF933F.xml
Normal file
|
|
@ -0,0 +1,554 @@
|
|||
<document id="6D8D0E2F38EF1B9EF1077B87E38813E8" ID-CLB-Dataset="300069" ID-DOI="10.3897/mycokeys.107.124814" ID-GBIF-Dataset="6bd1ed8a-9466-43f9-9c30-03cef8f291b5" ID-publisher-id="124814" URI-arpha="A492D575-58D9-5C61-8CEC-72ADB271323B" article-type="research-article" checkinTime="1721676058192" checkinUser="pensoft" docAuthor="Aung, Sein Lai Lai, Liu, Feng-Yin, Gou, Ya-Nan, Nwe, Zin Mar, Yu, Zhi-He & Deng, Jian-Xin" docDate="2024" docId="D4E3DDD826E95A6C969E6CC03CAF933F" docLanguage="en" docName="MycoKeys 107: 125-139" docOrigin="MycoKeys 107" docSource="https://mycokeys.pensoft.net/article/124814/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.mycokeys" docStyleName="PensoftTaxPub.0000.journal_article.mycokeys" docTitle="Alternaria jingzhouensis S. L. L. Aung & J. X. Deng 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="139" masterDocId="A492D57558D95C618CEC72ADB271323B" masterDocTitle="Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China" masterLastPageNumber="139" masterPageNumber="125" pageNumber="125" updateTime="1721676827755" updateUser="ExternalLinkService">
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<mods:mods id="DA7D544816DA4351D695A75908C0FA63" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="52D1AAA1BD40368A7BF3C4120CB74156">
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<mods:title id="008696889904614BBF4F46FDF95AA772">Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China</mods:title>
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<mods:namePart id="8A38B333EF7AFA567EAA29CF33E51854">Aung, Sein Lai Lai</mods:namePart>
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<mods:affiliation id="EB3C42E80FD0C0C0D1CB48C675AAD255">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:namePart id="1CC12996593700308BB1689B01E01F9E">Liu, Feng-Yin</mods:namePart>
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<mods:nameIdentifier id="4A375D6E1B1AC76698989DFF57B56982" type="ORCID">0000-0003-3114-603X</mods:nameIdentifier>
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<mods:affiliation id="28A5132D581A7D740AFA0E1D3C5E33B4">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:namePart id="3F2917A2E93B7C19241D7D0EE1D01D57">Gou, Ya-Nan</mods:namePart>
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<mods:nameIdentifier id="9102B148E5A7EFBC748C0FFC330368F0" type="ORCID">https://orcid.org/0009-0000-6376-8306</mods:nameIdentifier>
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<mods:affiliation id="A23EA5BBC0ADCF9FB3947185B4BF0C67">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:namePart id="A45D40CF491995758756A3C397B173BD">Nwe, Zin Mar</mods:namePart>
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<mods:nameIdentifier id="2539BB2A4B917472DDFAE4E564215891" type="ORCID">https://orcid.org/0009-0000-6376-8306</mods:nameIdentifier>
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<mods:affiliation id="7DB1C440FCBF07EB68A30D557F4C90E6">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:name id="701F1703256495BA38C3076F05EDFC62" type="personal">
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</mods:role>
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<mods:namePart id="E251D76A6184D90CE1C6F285B1D2ACC1">Yu, Zhi-He</mods:namePart>
|
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<mods:nameIdentifier id="FE94F8F71C233B93A6D98BA938585C68" type="ORCID">0000-0001-9477-4135</mods:nameIdentifier>
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<mods:affiliation id="4E3634CB33517312CB8903EBFD9634FB">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:namePart id="A0D509CA0FE1328759DD8623E2646115">Deng, Jian-Xin</mods:namePart>
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<mods:nameIdentifier id="DB3A6386CEA0D3C50F54F69E9D25C603" type="ORCID">0000-0001-7304-5603</mods:nameIdentifier>
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<mods:affiliation id="6CA9BD8FF6576AD0900100CBEB263500">Department of Plant Protection, College of Agriculture, Yangtze University, Jingzhou 434025, China</mods:affiliation>
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<mods:date id="7551EE47B8B5DF05D709359C0E7F579B">2024</mods:date>
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<mods:classification id="FD75C03D4F1C30D30F907D16EB76C346">journal article</mods:classification>
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<treatment id="D4E3DDD826E95A6C969E6CC03CAF933F" ID-DOI="http://doi.org/10.5281/zenodo.12796838" ID-Zenodo-Dep="12796838" ID-arpha="D4E3DDD8-26E9-5A6C-969E-6CC03CAF933F" ID-mycobank="MycoBank No: 851272" LSID="urn:lsid:plazi:treatment:D4E3DDD826E95A6C969E6CC03CAF933F" httpUri="http://treatment.plazi.org/id/D4E3DDD826E95A6C969E6CC03CAF933F">
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||||
<subSubSection id="4969536085390B4D83D90C98A6F8CEB0" type="nomenclature">
|
||||
<paragraph id="25565E5949316D96F0D873F76A1DCE1F">
|
||||
<taxonomicName id="AAF79C08F799EEA6A63903D887E2167A" authority="S. L. L. Aung & J. X. Deng" authorityName="S. L. L. Aung & J. X. Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="7EF3A1DB12C7BA90E5594104A4914534" italics="true">Alternaria jingzhouensis</emphasis>
|
||||
S. L. L. Aung & J. X. Deng
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="391D720D08F2D08566A138B6BF28228B" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="8BF0E7CB7D69FF94C5348847FB318C70" type="description">
|
||||
<paragraph id="D5F2339A63B3062C96D0C6A5B760CD3D">
|
||||
<figureCitation id="C0A677DF6CE6EE943F8586F83EE6F691" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alternaria jingzhouensis sp. nov. (ex-type YZU 221144) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure2" httpUri="https://binary.pensoft.net/fig/1097807">Fig. 2</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6JDI" type="material">
|
||||
<paragraph id="874CACA6190BAA981A3FC1A0299878F3">
|
||||
<heading id="1BEBB01CDBBBD8D9EF77D67F19D7EB18" reason="title">
|
||||
<typeStatus id="9733015216EE818CE7992C6FF81E52B5">Type</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5518B5A4AC72D513138CCCA37CE5B36D">
|
||||
<collectingCountry id="F53970F57DB6FD5326C2632D10E0FCAB" name="China">China</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="AEE3C1DEB3AEAE20F4603DFEAAB85B12" country="China" name="Hubei">Hubei Province</collectingRegion>
|
||||
, Jingzhou city, Yangtze University (west campus) on infected leaves of
|
||||
<taxonomicName id="BF656B7DFA9313B631330F2B8497F2DC" class="Magnoliopsida" family="Cucurbitaceae" genus="Citrullus" kingdom="Plantae" order="Cucurbitales" phylum="Tracheophyta" rank="species" species="lanatus">
|
||||
<emphasis id="BB230CD499F2996BD15C15BF39C839E8" italics="true">Citrullus lanatus</emphasis>
|
||||
</taxonomicName>
|
||||
2022,
|
||||
<collectionCode id="9E05FC3EEA298764013E25F0F650955E" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" type="Herbarium">F</collectionCode>
|
||||
.
|
||||
<collectionCode id="BA8B72A3EDFB0D540264C909E7A8C980" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" type="Herbarium">Y</collectionCode>
|
||||
Liu, (
|
||||
<abbrev id="ABBRID0EQKDI" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="617C8F502692D21C75EA399683413533" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
- H- 2022030,
|
||||
<typeStatus id="C7886FF5CF01376C719F85237AEFC591" type="holotype">holotype</typeStatus>
|
||||
), ex-type culture
|
||||
<abbrev id="ABBRID0EUKDI" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="DC68AC234C38B2F710AE3B892C94E116" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
221144.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EYKDI" type="etymology">
|
||||
<paragraph id="01F7D9039BAD0C25B978CD02608B2898">
|
||||
<heading id="B5E10EA985607F4669BCEE0CF567B4BC" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="C22146E45C5B7491C80D451F9E9A5FC9">
|
||||
Named after the collecting locality, Jingzhou (
|
||||
<collectingRegion id="0AA476247AD1DFE617F9713E6140D2CA" country="China" name="Hubei">Hubei</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="A4A3BAF5387063A45676056A3BA0F0DC" name="China">China</collectingCountry>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E4KDI" type="description">
|
||||
<paragraph id="7F5790295D9DEC31810159D0CB21F027">
|
||||
<heading id="7493C17B9290980C7BA5FA8F4BB44919" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5C6C6AFB811184A1627FE336B0B34AC5">
|
||||
<emphasis id="CD69191E950AB2B71A7ADB19175F3356" bold="true" italics="true">Colonies</emphasis>
|
||||
on
|
||||
<abbrev id="ABBRID0EILDI" xlink_title="potato dextrose agar">
|
||||
<collectionCode id="CE2FEE9D8E6B8947EB1B9253307502F1" country="Sri Lanka" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13944" lsid="urn:lsid:biocol.org:col:13944" name="Royal Botanic Gardens" type="Herbarium">PDA</collectionCode>
|
||||
</abbrev>
|
||||
(7 d at 25 °
|
||||
<collectionCode id="6FBB9B24F4EB9D54E53FB6B3D181EEF7" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
) pale luteous to amber in the center, white at the edges, light to moderate rosy buff or pale saffron in reverse, cottony surface and
|
||||
<quantity id="58CDB5310173E19A7B62BE182AC7EB4B" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" unit="mm" value="50.5" valueMax="52.0" valueMin="49.0">49–52 mm</quantity>
|
||||
in diam., at 25 °
|
||||
<collectionCode id="A94F4996E3E13A1875BB58965224CFE2" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
for 7 days (Fig.
|
||||
<figureCitation id="7A8CCAD95EA9D4C1E13E7EADF4B9C99B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alternaria jingzhouensis sp. nov. (ex-type YZU 221144) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure2" httpUri="https://binary.pensoft.net/fig/1097807">2 A, B</figureCitation>
|
||||
). On
|
||||
<abbrev id="ABBRID0EQLDI" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
(7 d at 22 °
|
||||
<collectionCode id="68E74BBB192BAE2014D3B76303F90CAC" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
),
|
||||
<emphasis id="95D1EB821B8C068856460B036AC3D51A" bold="true" italics="true">conidiophores</emphasis>
|
||||
arising from substrate, simple, straight or flexuous, light to olivaceous buff, 41–99 (– 151) × 3.5–5 μm (x ̄ = 73 × 4.4 µm, n = 20),
|
||||
<emphasis id="8461B9A70BF6F8DA2309EC16409A5B99" bold="true" italics="true">conidiogenous cells</emphasis>
|
||||
5–11 × 3–6 µm (x ̄ = 8 × 4 µm, n = 20), mono- to polytretic, terminal, determinate, cylindrical, olivaceous buff, smooth, thin-walled, apically doliiform, with 1 conidiogenous locus cicatrized on conidial secession, sometimes swollen near conidiogenous loci;
|
||||
<emphasis id="0090B3C95E0F668CD7FA368FF440D3ED" bold="true" italics="true">conidia</emphasis>
|
||||
3–5 units per chain, arising from the apex or near the apex of the conidiophores or terminal hyphae, muriform, ellipsoidal, flask-shaped, rostrate, beaked, 28–51 × 11–21 μm (x ̄ = 38 × 16.4, n = 50), with 1–4 transverse septa with 0–2 branching (Fig.
|
||||
<figureCitation id="89DD4D75A16F971225B9C84587097D71" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alternaria jingzhouensis sp. nov. (ex-type YZU 221144) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure2" httpUri="https://binary.pensoft.net/fig/1097807">2 C, E</figureCitation>
|
||||
); On
|
||||
<abbrev id="ABBRID0EBMDI" xlink_title="V8 juice agar">
|
||||
<collectionCode id="67292DF7282886E3E46A4850969788C2" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
8
|
||||
<collectionCode id="C016989F26E1244FC217899205BA58CF" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
|
||||
</abbrev>
|
||||
(7 d at 22 °
|
||||
<collectionCode id="D8E7685CFC6357ACE0F1BC4E36BA0BA3" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||||
),
|
||||
<emphasis id="8390A6340CAE5966670CD44641737850" bold="true" italics="true">conidiophores</emphasis>
|
||||
40–94 × 4–7 μm (x ̄ = 58 × 5, n = 20), simple, straight or flexuous, light to olivaceous buff;
|
||||
<emphasis id="40FB37A728B279E336D93820EB31035A" bold="true" italics="true">conidiogenous cells</emphasis>
|
||||
5–13 × 3–6 µm (x ̄ = 8 × 4 µm, n = 20), mono- to polytretic, terminal, determinate, cylindrical, olivaceous buff, smooth, thin-walled, apically doliiform, with 1 conidiogenous locus, sometimes swollen near conidiogenous loci cicatrized on conidial secession;
|
||||
<emphasis id="5ADB70570BC59DF32795BB42744D5A47" bold="true" italics="true">conidia</emphasis>
|
||||
3–5 units per chain, arising from the apex or near the apex of the conidiophores or terminal hyphae, muriform, ellipsoidal, flask-shaped, rostrate, beaked, 22–51 × 3–16 μm (x ̄ = 33.9 × 13.2, n = 50), 1–6 transverse septa with 0–2 branching (Fig.
|
||||
<figureCitation id="446723CCEB2410D411A2BD707003A380" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alternaria jingzhouensis sp. nov. (ex-type YZU 221144) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." figureDoi="10.3897/mycokeys.107.124814.figure2" httpUri="https://binary.pensoft.net/fig/1097807">2 D, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="6D441801B31A6E60292DA4E3CF4641AB" ID-DOI="10.3897/mycokeys.107.124814.figure2" ID-arpha="94156C3A-D24D-54FF-B8FC-C6326685893A" httpUri="https://binary.pensoft.net/fig/1097807" startId="F2">
|
||||
<paragraph id="2073A0E8AA0A7C019AEDD344570E07A4">
|
||||
<label id="D2FCC65298D86BE02EAA49656F561F05">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="EA02602552724175C4637A083EF8433F">
|
||||
<taxonomicName id="B89431ACBAB4031FDB9CF9C6AA36D115" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="BE1BA6DF956DD257531FF9C6CFBA66C5" italics="true">Alternaria jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="1C273E04157AEE856AE5C7FCB1ED9D4C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(ex-type
|
||||
<abbrev id="ABBRID0EFNDI" xlink_title="Fungi Herbarium of Yangtze University">YZU</abbrev>
|
||||
221144)
|
||||
<emphasis id="4F30F5D005295322AE9EB88E74DB3AE6" bold="true">A, B</emphasis>
|
||||
seven-day-old culture on
|
||||
<abbrev id="ABBRID0ELNDI" xlink_title="potato dextrose agar">PDA</abbrev>
|
||||
<emphasis id="3A4632A98EA124B068EBA29C45267F53" bold="true">C, D</emphasis>
|
||||
conidiophores and conidia on
|
||||
<abbrev id="ABBRID0ERNDI" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
and
|
||||
<abbrev id="ABBRID0EVNDI" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
, respectively
|
||||
<emphasis id="E3EA3E0F6E521442CD6787FE656495BA" bold="true">E, F</emphasis>
|
||||
conidia on
|
||||
<abbrev id="ABBRID0E2NDI" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
and
|
||||
<abbrev id="ABBRID0E6NDI" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
, respectively. Scale bars: 25 μm (
|
||||
<emphasis id="D18B87C235446C4CC830C86756425BC7" bold="true">E, F</emphasis>
|
||||
); 50 μm (
|
||||
<emphasis id="765D22A3787FBDD79EE003379CF1802D" bold="true">C, D</emphasis>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMODI" type="Additional isolate examined">
|
||||
<paragraph id="433A73404A5344E65A7F88A1402DAEC0">
|
||||
<heading id="E42718ECD0E10D28517EB732D4FF91C6" reason="title">Additional isolate examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA9A6EEEA46F4AC3D833F883F2D21939">
|
||||
<collectingCountry id="F5B8135EE29F70F4801390C2B1598AD8" name="China">China</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="99DB4DDED159ED72E34A0042A153C335" country="China" name="Hubei">Hubei Province</collectingRegion>
|
||||
, Jingzhou city, Yangtze University (west campus) on infected leaves of
|
||||
<taxonomicName id="D2DB7300D125A639793B06AE028551C5" class="Magnoliopsida" family="Cucurbitaceae" genus="Citrullus" kingdom="Plantae" order="Cucurbitales" phylum="Tracheophyta" rank="species" species="lanatus">
|
||||
<emphasis id="F76C845BE215BDFBCCA958BFC5776951" italics="true">Citrullus lanatus</emphasis>
|
||||
</taxonomicName>
|
||||
2022,
|
||||
<collectionCode id="9631C8F89BE547631ECE54CD33E98D59" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" type="Herbarium">F</collectionCode>
|
||||
.
|
||||
<collectionCode id="3E116D3D710542B11CF0B16012B67836" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" type="Herbarium">Y</collectionCode>
|
||||
Liu, living culture
|
||||
<abbrev id="ABBRID0E4ODI" xlink_title="Fungi Herbarium of Yangtze University">
|
||||
<collectionCode id="2784D4ACF290ECF041BE579E001A5F01" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14820" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
|
||||
</abbrev>
|
||||
221145.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBPDI" type="notes">
|
||||
<paragraph id="2D72CA925F1569B1BE47A849A025F4D8">
|
||||
<heading id="83A52777261D3323A4F13FF6B968DC91" reason="title">Notes.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DD5435996D801819528291258A7259A7">
|
||||
Phylogenetically,
|
||||
<taxonomicName id="3B77BABC85FC6883C09327900CFCD476" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="28FEC32B8FE94F86FDDD9A742427C888" italics="true">A. jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7CD360A386625CF15BF20FCD05D72472" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is different from its sister species
|
||||
<taxonomicName id="F121A568C6EDAC1659518692A721FAF6" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="baoshanensis">
|
||||
<emphasis id="FE67C8533468D4B7BDD4282A03853A48" italics="true">A. baoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C99E16FD05DAF2E05F1855B407693B66" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="2BEDAA3FDB85C705F33CD2218B8FCAA4" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="168FB69A4BFF9AAEE04D5D76EE246F2A" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="A237E12ACF6F8010E6D650839D9584D6" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E92C425AE8E156815E97CC44A07DE41F" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="0D7D72E8872B0442427520B111F2F1B6" class="Magnoliopsida" family="Asteraceae" genus="Atractylodes" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="orobanches">
|
||||
<emphasis id="35FA109C4F57037C4A334EA1C65086C1" italics="true">A. orobanches</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="97FEF4963AB0A2CA5D0BFA1E6F9C714C" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="ovoidea">
|
||||
<emphasis id="649983AD561480EF466E54D5CCA3D205" italics="true">A. ovoidea</emphasis>
|
||||
</taxonomicName>
|
||||
based on sequences derived from seven genes (Fig.
|
||||
<figureCitation id="D4DDD5F2FF3239867BB8CDC91FC34DFD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree of the Alternaria species most related to the new taxa based on maximum likelihood analysis using the combined gene sequences of ITS, GAPDH, TEF 1, RPB 2, Alt a 1, EndoPG and OPA 10-2 which rooted with Alternaria alternantherae (CBS 124392) from sect. Alternantherae. The Bayesian posterior probabilities > 0.60 (PP) and bootstrap support values > 60 (BS) are given at the nodes (PP / BS). The novel species are highlighted in bold. Ex-type isolates are marked with a superscript T and Representative isolates are marked with a superscript R." figureDoi="10.3897/mycokeys.107.124814.figure1" httpUri="https://binary.pensoft.net/fig/1097806">1</figureCitation>
|
||||
). After conducting a nucleotide pairwise comparison as recommended by
|
||||
<bibRefCitation id="4965B1F725FED11DDA1ED9CA556610C3" DOI="10.5943/mycosphere/7/11/4" author="Jeewon R & Hyde KD" firstAuthor="Jeewon" issue="11" journalOrPublisher="Mycosphere" pagination="1669-1677" refId="B10" refString="Jeewon R, Hyde KD (2016) Establishing species boundaries and new taxa among fungi: Recommendations to resolve taxonomic ambiguities. Mycosphere 7 (11): 1669–1677. https://doi.org/10.5943/mycosphere/7/11/4" title="Establishing species boundaries and new taxa among fungi: Recommendations to resolve taxonomic ambiguities." volume="7" year="2016">Jeewon and Hyde (2016)</bibRefCitation>
|
||||
, the present species can be readily distinguished from the closet species
|
||||
<taxonomicName id="6B84C53133E0ED60724AB65A1F4AB0B8" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="E69F96ABF348761E758FE6126394C4FC" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="CE72A0B486A2D797059954735707E50B" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="77938E20C179B0B9B079659F66D82BDC" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="8583421CB890CDC028457A70C166AE99" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="E371EA139EED5FE25DAD32C87E952FC3" class="Magnoliopsida" family="Asteraceae" genus="Atractylodes" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="orobanches">
|
||||
<emphasis id="4A10EE74D0322A85431DE123BABA3FE0" italics="true">A. orobanches</emphasis>
|
||||
</taxonomicName>
|
||||
constructed on any of the
|
||||
<abbrev id="ABBRID0ESSDI" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="DEFA13F6F6A11EE2F352F45D39908CF1">ITS</collectionCode>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="60CCC50EBAA2B427B49ED9763FF76904" italics="true">
|
||||
<collectionCode id="A46A31A05DE731D481F4F0A77EFF43BB">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0EZSDI" xlink_title="translation elongation factor 1 alpha">
|
||||
<emphasis id="6E78B6BB5CE3FE1D60E0AF9879D5650F" italics="true">
|
||||
<collectionCode id="5410A3515044CE658EB4644863C236DB" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" type="Herbarium">TEF</collectionCode>
|
||||
1
|
||||
</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<abbrev id="ABBRID0E5SDI" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="2E7FBA9996830EB72FDE523C8ED384AC" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
,
|
||||
<emphasis id="2BFD0FCABC11D37566611A9DAE740B1A" italics="true">Alt a 1</emphasis>
|
||||
,
|
||||
<abbrev id="ABBRID0EFTDI" xlink_title="endopolygalacturonase gene">
|
||||
<emphasis id="401B07453845D9A62ACE5E55615C46D6" italics="true">EndoPG</emphasis>
|
||||
</abbrev>
|
||||
, and OPA 10-2 genes, which has 1 bp difference in the
|
||||
<abbrev id="ABBRID0EJTDI" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="C4CCA455AEDCC4E9A63CD9AA47EBA75D">ITS</collectionCode>
|
||||
</abbrev>
|
||||
region, 1 bp in
|
||||
<emphasis id="F3B8B98D14DDF8CF5AC7D53F7CE97FEA" italics="true">
|
||||
<collectionCode id="6BFFEC82CE6267DACC04C9D5FA459289">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
, 1 bp in
|
||||
<abbrev id="ABBRID0EQTDI" xlink_title="translation elongation factor 1 alpha">
|
||||
<emphasis id="C7844EC051A3BCAAB628D81806677D0F" italics="true">
|
||||
<collectionCode id="95FCC204C845155480C2CE700ECBA2E8" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" type="Herbarium">TEF</collectionCode>
|
||||
1
|
||||
</emphasis>
|
||||
</abbrev>
|
||||
, 7 pb in
|
||||
<abbrev id="ABBRID0EVTDI" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="C7D0C1C978493B79FA54AD2A985E7664" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
, 9 bp in
|
||||
<emphasis id="DC1A34B0F2B9A99BC31A078E3DAC770E" italics="true">Alt a 1</emphasis>
|
||||
, 10 bp in
|
||||
<abbrev id="ABBRID0E3TDI" xlink_title="endopolygalacturonase gene">
|
||||
<emphasis id="325921C23DDCFFF9E3A9C0584DF80BBF" italics="true">EndoPG</emphasis>
|
||||
</abbrev>
|
||||
, and 4 bp in OPA 10-2 when compared with
|
||||
<taxonomicName id="E6F34529E5029E2C966661CD2E3C760A" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="CCE60D368C7BEC54CE9AF0F29B47D420" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
, 1 bp in
|
||||
<emphasis id="7A730F1D8ABB087BDF720BD47F3DEE8D" italics="true">
|
||||
<collectionCode id="D7BDB1353D0FD18D8A051069D39A136E">GAPDH</collectionCode>
|
||||
</emphasis>
|
||||
, 4 bp in
|
||||
<abbrev id="ABBRID0EOUDI" xlink_title="RNA polymerase second largest subunit">
|
||||
<emphasis id="F40DB0B331C4581C06E632D640250907" italics="true">RPB 2</emphasis>
|
||||
</abbrev>
|
||||
, and 11 bp in OPA 10-2 when compared with
|
||||
<taxonomicName id="CEAB2B1DC483705F1CE20AE542C7E9B3" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="ABA58B38C6B795559BA7CC4421AB12D7" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="716983F27C0DEBE67572EF355389E03E" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and 49 bp differences in the
|
||||
<abbrev id="ABBRID0E4UDI" xlink_title="internal transcribed spacer rDNA region">
|
||||
<collectionCode id="81D9C4EC2C1BA47992E42B99489407C7">ITS</collectionCode>
|
||||
</abbrev>
|
||||
region when compared with sister species
|
||||
<taxonomicName id="93CB6E806757A404E85ED4C4820C2903" class="Magnoliopsida" family="Asteraceae" genus="Atractylodes" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="orobanches">
|
||||
<emphasis id="273B4F02B03ED8EDDD5C7BBF3B26E48A" italics="true">A. orobanches</emphasis>
|
||||
</taxonomicName>
|
||||
. Morphologically, the species is distinct from
|
||||
<taxonomicName id="76F9D524A2ED40ABAD35E46CDA8CBEC2" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="baoshanensis">
|
||||
<emphasis id="091ACEC65D140F6A03E133C7A0D0DD2E" italics="true">A. baoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="8467FE42B354D12DEC47C99D80234428" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="839B535F0A925C0F30931F3733165964" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="8352CE2324F27BD74DADCEC373F83FC9" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="ovoidea">
|
||||
<emphasis id="9CA74A48CEEF394A686B77F84BD0C581" italics="true">A. ovoidea</emphasis>
|
||||
</taxonomicName>
|
||||
as it produces conidia on a simple conidiogenous locus with occasionally longer beaks in a chain of 3–5 units, and from
|
||||
<taxonomicName id="A01944CAC4C2FC401F42D987496B1AD6" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="98386F8F7A966757ED8041037DD00728" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="968B8B9C20655BF2C11647C64168DFA4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by having shorter beaks (Table
|
||||
<tableCitation id="BCFE0A7ACAF3A99BD17CCE2E7A932203" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Conidial features of the novel Alternaria species proposed here and their closest relatives in section Alternaria." httpUri="http://table.plazi.org/id/0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7" tableUuid="0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7">2</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7" ID-Table-UUID="0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7" httpUri="http://table.plazi.org/id/0BFCE5554CB7F1D4EEDB4F7B1E5BF3A7" startId="T2">
|
||||
<paragraph id="33194AAE0C0802FFD0CDB469F0F29533">
|
||||
<label id="B908DFDACE5ECCDA2F5646830EA2C000">Table 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="040693AFC1FDBA12612EC67CDAFA1E5D">
|
||||
Conidial features of the novel
|
||||
<taxonomicName id="B799662B380ADDED92604C679B584A28">
|
||||
<emphasis id="2823B83BAFDB8A533C4106A7CA7ECFF4" italics="true">Alternaria</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
proposed here and their closest relatives in section
|
||||
<taxonomicName id="3CB6D4A72E2AA68FA4DBC23E72C0AF04" rank="section" section="Alternaria">
|
||||
<emphasis id="A375ADA658B34BA8123EAB1047CC1992" italics="true">Alternaria</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8A47D81D9A6E567F4751425127CE89">
|
||||
<table id="TID0EIYAK" rules="all">
|
||||
<tbody id="439F2D542BD1FD180351737F75792271">
|
||||
<tr id="73D90177BCEF61B4A4409886BB32A838">
|
||||
<th id="C75568AF04856C29AC4563A9530D7A5E" colspan="1" rowspan="2">Species</th>
|
||||
<th id="C8906E987C8330181325655C1A394A4A" colspan="4" rowspan="1">Conidia</th>
|
||||
<th id="9972C346605307E8C8998050CF780B6B" colspan="1" rowspan="2">Conidia per chain</th>
|
||||
<th id="2F0C83349B5F952ACD07A700C208DC14" colspan="1" rowspan="2">Medium</th>
|
||||
<th id="366C88384F90A6DBEE094A04087380BD" colspan="1" rowspan="2">Reference</th>
|
||||
</tr>
|
||||
<tr id="18D97C681F3A702ED9282367A707CC52">
|
||||
<th id="C6979E57D28631803A9DE91414F34C94" colspan="1" rowspan="1">Shape</th>
|
||||
<th id="46994035CD26A447613AF68FE1CDAA8F" colspan="1" rowspan="1">Body (µm)</th>
|
||||
<th id="D4AB5FA157EEE948EF125724D42FD22A" colspan="1" rowspan="1">Beak (µm)</th>
|
||||
<th id="28625672555045A0FA9A55FC119D3307" colspan="1" rowspan="1">Septa</th>
|
||||
</tr>
|
||||
<tr id="4EF723F482F4EB821795A1F624C06DDD">
|
||||
<td id="E74E5A24874F179A805E67671A5E4FE9" colspan="1" rowspan="1">
|
||||
<taxonomicName id="0146C9B15EF0ABE3C739222D2245CD89" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="baoshanensis">
|
||||
<emphasis id="8C3BF29B61867925BD5F1149F1666437" italics="true">A. baoshanensis</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
<td id="1835336673D3AAF76CCB7B3EAFB52BAA" colspan="1" rowspan="1">Subglobose to ellipsoidal, or subcylindrical to obpyriform</td>
|
||||
<td id="D753F593EDD568EEFEA19518BDC9C44B" colspan="1" rowspan="1">25–60 × 12–22</td>
|
||||
<td id="584109B784CDFB4EC7F663E555CD336C" colspan="1" rowspan="1">Short beak</td>
|
||||
<td id="5FF8DC3433D6F6D319E828C34B2B69E5" colspan="1" rowspan="1">3–6</td>
|
||||
<td id="03135F13DF2DF1D8D8FF72FFF2C7CB9D" colspan="1" rowspan="1">1–3</td>
|
||||
<td id="15E3CC6709EDCAA68988B19811EB35DD" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EZZDI" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
</td>
|
||||
<td id="794AFD31F6878BB08D201D7FEA4398F3" colspan="1" rowspan="1">
|
||||
<bibRefCitation id="4228E43567A5E821ECF39DE15A521512" DOI="10.48130/SIF-2023-0001" author="Li JF & Jiang HB & Jeewon R & Hongsanan S & Bhat DJ & Tang SM & Lumyong S & Mortimer PE & Xu JC & Camporesi E & Bulgakov TS & Zhao GJ & Suwannarach N & Phookamsak R" etAl="et al." firstAuthor="Li" issue="1" journalOrPublisher="Studies in Fungi" pagination="1-61" refId="B16" refString="Li JF, Jiang HB, Jeewon R, Hongsanan S, Bhat DJ, Tang SM, Lumyong S, Mortimer PE, Xu JC, Camporesi E, Bulgakov TS, Zhao GJ, Suwannarach N, Phookamsak R (2023) Alternaria: Update on species limits, evolution, multi-locus phylogeny, and classification. Studies in Fungi 8 (1): 1–61. https://doi.org/10.48130/SIF-2023-0001" title="Alternaria: Update on species limits, evolution, multi-locus phylogeny, and classification." volume="8" year="2023">Li et al. (2023)</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<emphasis id="511FCE47C6A7933DED71CBE79B1CB3B0" bold="true">
|
||||
<tr id="A9AA903782DCAA1D1A81D7B5670FD40E">
|
||||
<td id="D7162ADC014CAE4514F12C3EC8BBDDD2" colspan="1" rowspan="2">
|
||||
<taxonomicName id="DBB51EA0B1C81A62B92D0D530FC2A0E7" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="jingzhouensis" status="sp. nov.">
|
||||
<emphasis id="CF782F4A008E0D321B3454FD52301008" bold="true" italics="true">A. jingzhouensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="BCA4DEBC79C5815B11F4219C29F465A8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</td>
|
||||
<td id="DA4D9A76096F6A817C3E552B45114DB3" colspan="1" rowspan="2">Ellipsoidal, flask-shaped, rostrate, beaked</td>
|
||||
<td id="3EBB596B94D22EBE76CB5F9FA98E5DBD" colspan="1" rowspan="1">28–51 × 11–21</td>
|
||||
<td id="ABB72DF73BE421C2FBA5C46C0E7E71D9" colspan="1" rowspan="1">2–7 (– 15)</td>
|
||||
<td id="305E170C05B1A1C2099A58E741A9FF48" colspan="1" rowspan="1">1–4</td>
|
||||
<td id="9BB1AE5155B72FD48E39EBE0919BC94F" colspan="1" rowspan="1">3–5</td>
|
||||
<td id="2901FFF13D64AD97CD9F5878BC35DF33" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0E22DI" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
</td>
|
||||
<td id="D234E880BDD3DDECEEDF52861FA9E9C2" colspan="1" rowspan="1">Present study</td>
|
||||
</tr>
|
||||
<tr id="7AB5BCF27AFCE74596497644CED2D83A">
|
||||
<td id="1B1932294EAE8C9AA6EAF3BB7BE8BD4B" colspan="1" rowspan="1">22–51 × 3–16</td>
|
||||
<td id="D27C5C5C8FBA6A8DA58A72F6731CAF66" colspan="1" rowspan="1">3–7</td>
|
||||
<td id="5EB47A9F801B074F066AC17C6D1B17E0" colspan="1" rowspan="1">1–6</td>
|
||||
<td id="77609DB5E3DB1B541BEC2AB58D0E2911" colspan="1" rowspan="1">3–5</td>
|
||||
<td id="0CFB4ED4E8A8FA0528811AEC38CDE3DA" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0ED4DI" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
</td>
|
||||
<td id="60C8808C4789D956377A342395E7CA7F" colspan="1" rowspan="1">Present study</td>
|
||||
</tr>
|
||||
</emphasis>
|
||||
<tr id="85587A1F476D53B6EF9CFFA4203F9BB9">
|
||||
<td id="84E18EC417B62254A8863B59FD46521A" colspan="1" rowspan="1">
|
||||
<taxonomicName id="1FAEC112BB90D9591506C91C997E571A" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="koreana">
|
||||
<emphasis id="2A2250FB9F5897A7B744F74B3291B16C" italics="true">A. koreana</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
<td id="05AAF39E7C0275853319A224958516FE" colspan="1" rowspan="1">Obovate to long ellipsoid</td>
|
||||
<td id="D8B905F121819D75FF898F04A0FD4C9B" colspan="1" rowspan="1">12.9–61.2 × 8.6 – 20.7</td>
|
||||
<td id="9EF50E0656F5148B5684F82ADE9B2A05" colspan="1" rowspan="1">4.5–9.1</td>
|
||||
<td id="C2661214D73276C2C921AA12D6466ADA" colspan="1" rowspan="1">2–8</td>
|
||||
<td id="1F3B2E947D4CDCAC7A6A83FE8E57490E" colspan="1" rowspan="1">1–2</td>
|
||||
<td id="EBA502E3BF02EAE751BEACD1114EFF65" colspan="1" rowspan="1">SNA</td>
|
||||
<td id="8BB99FC52D5313817F69649BE74517D7" colspan="1" rowspan="1">
|
||||
<bibRefCitation id="F824BD18CB8458BD515A18AEEA9CA036" DOI="10.1007/s11033-021-06887-9" author="Romain BBND & Hassan O & Kim JS & Chang T" etAl="et al." firstAuthor="Romain" issue="1" journalOrPublisher="Molecular Biology Reports" pagination="413-420" refId="B32" refString="Romain BBND, Hassan O, Kim JS, Chang T (2022) Alternaria koreana sp. nov., a new pathogen isolated from leaf spot of ovate-leaf Atractylodes in South Korea. Molecular Biology Reports 49 (1): 413–420. https://doi.org/10.1007/s11033-021-06887-9" title="Alternaria koreana sp. nov., a new pathogen isolated from leaf spot of ovate-leaf Atractylodes in South Korea." volume="49" year="2022">Romain et al. (2022)</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<emphasis id="70E1364F5C5953A5B6E791DEA263EF28" bold="true">
|
||||
<tr id="F2E9EBEC9A2DEA657A5304EAB46C17A5">
|
||||
<td id="9C995CE4994E4B0721538E5435428C5F" colspan="1" rowspan="2">
|
||||
<taxonomicName id="BCBC82EFC68BC4F0E73181E7207822AE" authorityName="Aung & Liu & Gou & Nwe & Yu & Deng" authorityYear="2024" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="momordicae" status="sp. nov.">
|
||||
<emphasis id="4F8B10DD7B6AEA02F4D672B9B35E8DE6" bold="true" italics="true">A. momordicae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="9065A39950CAF1356741FF743437D740" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</td>
|
||||
<td id="5FCAAF1F6A67066A753D5B65AF190F98" colspan="1" rowspan="2">Obclavate, inverted club-shaped</td>
|
||||
<td id="5BDC429DEDD8CA8C8D3254AA05D8935F" colspan="1" rowspan="1">6–42 × 4–34</td>
|
||||
<td id="58E8614D2F206954D92490C54B08C0E3" colspan="1" rowspan="1">2–19.5</td>
|
||||
<td id="3025115AD467198ACBC653AED797D647" colspan="1" rowspan="1">1–5</td>
|
||||
<td id="2277949B456EB3636ADE309C65102CDC" colspan="1" rowspan="1">3–4</td>
|
||||
<td id="1646B33581E8FFE261A4C9A96871857A" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EOAAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
</td>
|
||||
<td id="DFB1516B9476950886E630C00DA15A96" colspan="1" rowspan="1">Present study</td>
|
||||
</tr>
|
||||
<tr id="4B4352978CB442BF6530901D68CD888E">
|
||||
<td id="047D73C2FBEAD67E823118BFF16D42A6" colspan="1" rowspan="1">24–61 × 10–17</td>
|
||||
<td id="3A7A36658DC5DB9A9EC83A923CC01540" colspan="1" rowspan="1">3–25.5</td>
|
||||
<emphasis id="911FB79C74786825B5719F3F7E187F1B" bold="true">
|
||||
<td id="45FA45F1E6CE89674FE5E42E392B1DB7" colspan="1" rowspan="1">1–5</td>
|
||||
<td id="7F0C84D9304BDC622EC477B2339EDA58" colspan="1" rowspan="1">3–4</td>
|
||||
<td id="C31CF08C9738C3B6167E6E2799ABAFCF" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EWBAK" xlink_title="V8 juice agar">V 8 A</abbrev>
|
||||
</td>
|
||||
<td id="A2F9CBBCB655E8C1A8E156A716417B47" colspan="1" rowspan="1">Present study</td>
|
||||
</emphasis>
|
||||
</tr>
|
||||
</emphasis>
|
||||
<tr id="E01FC2EF12C74643871342654E8E05F8">
|
||||
<td id="3211CB1EB1352A6583AB49D19B2DAE9F" colspan="1" rowspan="1">
|
||||
<taxonomicName id="CA33C403B72A5EB5E5F13F1A14EB1E85" class="Magnoliopsida" family="Asteraceae" genus="Atractylodes" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="orobanches">
|
||||
<emphasis id="740C4C593BCFAA92097A4E9BE2B1C517" italics="true">A. orobanches</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
<td id="F29101C200B501E2D1EB2C61B81DBAA2" colspan="1" rowspan="1">Obclavate to ovoid</td>
|
||||
<td id="509CEF884E9E27E1A7DDB3A95702AE0D" colspan="1" rowspan="1">20–50 × 10–20</td>
|
||||
<td id="D7DC4AA7C1EA6E7B1F7BC6D0AD4D84C7" colspan="1" rowspan="1">–</td>
|
||||
<td id="7CBE7B34CE7290389B13747DB79E8172" colspan="1" rowspan="1">3 – 6</td>
|
||||
<td id="4F3AE5DA3E8EA47E2B99DF19738A2990" colspan="1" rowspan="1">1–2</td>
|
||||
<td id="940C503117077008CAE6F96E600F3749" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0ECDAK" xlink_title="potato carrot agar">PCA</abbrev>
|
||||
</td>
|
||||
<td id="C8202C5257508D1D05684116058FC6DA" colspan="1" rowspan="1">
|
||||
<bibRefCitation id="1E5EBBF02664E134FE60140B3248AFC5" DOI="10.48130/SIF-2023-0001" author="Li JF & Jiang HB & Jeewon R & Hongsanan S & Bhat DJ & Tang SM & Lumyong S & Mortimer PE & Xu JC & Camporesi E & Bulgakov TS & Zhao GJ & Suwannarach N & Phookamsak R" etAl="et al." firstAuthor="Li" issue="1" journalOrPublisher="Studies in Fungi" pagination="1-61" refId="B16" refString="Li JF, Jiang HB, Jeewon R, Hongsanan S, Bhat DJ, Tang SM, Lumyong S, Mortimer PE, Xu JC, Camporesi E, Bulgakov TS, Zhao GJ, Suwannarach N, Phookamsak R (2023) Alternaria: Update on species limits, evolution, multi-locus phylogeny, and classification. Studies in Fungi 8 (1): 1–61. https://doi.org/10.48130/SIF-2023-0001" title="Alternaria: Update on species limits, evolution, multi-locus phylogeny, and classification." volume="8" year="2023">Li et al. (2023)</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="24A581DE4A7D24EAA6E39135E9A2BF9C">
|
||||
<td id="FAB13CFF0753F82B1FF28ADCCB0AA2E4" colspan="1" rowspan="1">
|
||||
<taxonomicName id="EC7CCBF37848BFAA55986267A2EBDF61" class="Dothideomycetes" family="Pleosporaceae" genus="Alternaria" kingdom="Fungi" order="Pleosporales" phylum="Ascomycota" rank="species" species="ovoidea">
|
||||
<emphasis id="C88C373CD6174D3EA9C74DB46B8CB088" italics="true">A. ovoidea</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
<td id="E0A3AF20DFE6133EF5527CA1354911BD" colspan="1" rowspan="1">Ovoid</td>
|
||||
<td id="AA5FB7D276934B85F3A615F690F2B73E" colspan="1" rowspan="1">48–65 × 15.5–30</td>
|
||||
<td id="BC2D058B54737FB095195470D8D45E95" colspan="1" rowspan="1">–</td>
|
||||
<td id="2169C78E8B0C8D23012A398CB4858F40" colspan="1" rowspan="1">1 – 3</td>
|
||||
<td id="47343124D3F4766B476132392BD0D140" colspan="1" rowspan="1">1</td>
|
||||
<td id="5DBC6BFAC754EDEB810F4DB0699D501A" colspan="1" rowspan="1">
|
||||
<abbrev id="ABBRID0EREAK" xlink_title="potato dextrose agar">PDA</abbrev>
|
||||
</td>
|
||||
<td id="57334429C44CA04AB383AD44D18D4372" colspan="1" rowspan="1">
|
||||
<bibRefCitation id="EBEF1FB1B3C5139BED9D0DA950314772" DOI="10.3390/jof8090898" author="Li J & Phookamsak R & Jiang H & Bhat DJ & Camporesi E & Lumyong S & Kumla J & Hongsanan S & Mortimer PE & Xu J & Suwannarach N" etAl="et al." firstAuthor="Li" issue="9" journalOrPublisher="Journal of Fungi (Basel, Switzerland)" pagination="898" refId="B15" refString="Li J, Phookamsak R, Jiang H, Bhat DJ, Camporesi E, Lumyong S, Kumla J, Hongsanan S, Mortimer PE, Xu J, Suwannarach N (2022) Additions to the Inventory of the Genus Alternaria Section Alternaria (Pleosporaceae, Pleosporales) in Italy. Journal of Fungi (Basel, Switzerland) 8 (9): 898. https://doi.org/10.3390/jof8090898" title="Additions to the Inventory of the Genus Alternaria Section Alternaria (Pleosporaceae, Pleosporales) in Italy." volume="8" year="2022">Li et al. (2022)</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
</tbody>
|
||||
</table>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
325
data/EE/13/BC/EE13BC0C504859B79CEF09240294C48E.xml
Normal file
325
data/EE/13/BC/EE13BC0C504859B79CEF09240294C48E.xml
Normal file
|
|
@ -0,0 +1,325 @@
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<mods:title id="B0DAD5A1991F9362EF711F19E88D9840">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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|
||||
<taxonomicName id="6CD09F04ED124F4F2E047222B78D8016" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix" status="sp. nov." subGenus="Liothorax">
|
||||
<emphasis id="6E590B798425374593BD0C303B991FB0" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="8446465333A1D8279CD1743C44B648DE" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="13959198A5847E37B4DFF6C8C4CC478A" italics="true">felix</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4471DF7F9B0970A869598C0660F0782E" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
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|
||||
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|
||||
<subSubSection id="13BE7BA26DA41D65C9AABF4220768280" type="description">
|
||||
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|
||||
<figureCitation id="8CBC2968100DB34DA90283160399203F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 c</figureCitation>
|
||||
,
|
||||
<figureCitation id="4D34186ADC42CB5BEF7F43DDD1B95C07" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 i</figureCitation>
|
||||
,
|
||||
<figureCitation id="A99659BF222CD4061DA6CDAAA932C7F0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 e, f</figureCitation>
|
||||
,
|
||||
<figureCitation id="03FBF4C77632454E7D59497C82D1A5F5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." figureDoi="10.3897/zookeys.1207.117225.figure15" httpUri="https://binary.pensoft.net/fig/1097476">15 d</figureCitation>
|
||||
,
|
||||
<figureCitation id="F32EF1C9CF23624DACCC80A8FAE44FB9" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 v</figureCitation>
|
||||
,
|
||||
<figureCitation id="17C8A3392557CDAD442AA09559B83DA0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 p, q</figureCitation>
|
||||
,
|
||||
<figureCitation id="8E54F16E4C572C77A4D2B91F2C81E738" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 f, k</figureCitation>
|
||||
,
|
||||
<figureCitation id="C3EB25802787DE3C3CDAF25D14086F52" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 v, w</figureCitation>
|
||||
,
|
||||
<figureCitation id="435E03EA3436437BCE5EB311266FEF57" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure25" httpUri="https://binary.pensoft.net/fig/1097496">25 c</figureCitation>
|
||||
,
|
||||
<figureCitation id="82AC3D3C27E6CC959C2FAC3A69BCC149" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 v</figureCitation>
|
||||
,
|
||||
<figureCitation id="57401DE1B7AD6783E03A59F59B612217" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure32" httpUri="https://binary.pensoft.net/fig/1097519">32 a – e</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3LBM" type="type material">
|
||||
<paragraph id="EAB8FD6502F57ECF37F5DB3C9D50BCFF">
|
||||
<heading id="BAC715BF834AC8AA41E0D73AA2E6FBBA" reason="title">Type material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="D4EDD0DD27988C29FA95201A50104297">
|
||||
<materialsCitation id="F00E29269E0C70EF7FBB0BF6B571FCED" collectingDate="2009-06-01" collectingDateMax="2009-06-07" collectingDateMin="2009-06-01" collectionCode="NHMUK" collectorName="R. B. Angus. Chromosome & R. B. Angus" country="Italy" location="Abruzzo" specimenCount="1" specimenCount-male="1" stateProvince="Abruzzo" typeStatus="holotype">
|
||||
<emphasis id="4B301194B6143A46B6A69936DDDB0C6C" bold="true" italics="true">
|
||||
<typeStatus id="B89CA2FB2DC0ADAB255F3899FB4257C6" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount id="DEF8EDA4A73A58409330EB2A61F8DD60" count="1" type="male">♂</specimenCount>
|
||||
:
|
||||
<collectingCountry id="44E38418A8013DCED2ED80EEC2206F3D" name="Italy">Italy</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="F16B4AFD55DA6E072923D4815044D269" country="Italy" name="Abruzzo">Abruzzo</collectingRegion>
|
||||
, Campo Felice. Washed into pool.
|
||||
<date id="EBA4CA7716AC088A69274283E23E4DD8" value="2009-06-01">
|
||||
<collectingDate id="7D2B455DD0C8A0F67094FF29D3E4B4ED" value="2009-06-01">1. vi. 2009</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="E996AF74FCC7D9A696AB59A8D794B65E">R. B. Angus. Chromosome</collectorName>
|
||||
prep. 4,
|
||||
<date id="41D9147BD79E21B4C183CAAA7D32C757" value="2009-06-07">
|
||||
<collectingDate id="E818C13AAC6854AFCE7DEFED0645AB4F" value="2009-06-07">7. vi. 2009</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="94E177A815F03D7824525860EC7F3E18">R. B. Angus</collectorName>
|
||||
(
|
||||
<named-content id="EF2012647E82B6513BBFF9F560DD8B31" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="5B0B958B719928D1F4CAF0825DA886E7" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<emphasis id="C0CF04D022CDF9FB161B6E612C44D87D" bold="true" italics="true">
|
||||
<typeStatus id="0810869A274E8F82A17B098A49B4C471" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="4A0372080B09155B2614A38220580705" count="1" type="male">1 ♂</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="1F1AF7271AF499F92FCD6991B8CF025C" type="holotype">holotype</typeStatus>
|
||||
, chromosome prep. 3,
|
||||
<date id="1B7BF0F27A7AA7A564A3566BDCF70818" value="2009-06-07">7. vi. 2009</date>
|
||||
. R. B. Angus;
|
||||
<specimenCount id="3696F591C89B9D554C2D10C7FAE56C1D" count="3" type="female">3 ♀♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="CB8D4DAD671909DD0A073E3E4CEF088F" type="holotype">holotype</typeStatus>
|
||||
, chromosome preps 1 – 3,
|
||||
<date id="9E74D42AA0363BFCF57A05855EDAAFE4" value="2009-06-09">9. vi. 2009</date>
|
||||
. R. B. Angus (
|
||||
<named-content id="D33527ACF2C425A0DE34C7F2AD266CF2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="F956A4E7ED08675CFBDA4F2116B38157" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
);
|
||||
<specimenCount id="B11828413EA56D19E441724DD7142DB7" count="1" type="female">1 ♀</specimenCount>
|
||||
, data as
|
||||
<typeStatus id="0A69FA80FB2D9D01CEB3018C7F9E136F" type="holotype">holotype</typeStatus>
|
||||
,
|
||||
<date id="86C486D5FFDF775C4851DE54EBEDB2D6" value="2009-06-01">1. vi. 2009</date>
|
||||
(sequenced) (
|
||||
<abbrev id="ABBRID0ESMBM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="9CB2080B3D1421442827D26B75BCCD51">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
<collectingRegion id="1BEA89C06CC19E17E2DA8A370DC0729C" country="Italy" name="Basilicata">Basilicata</collectingRegion>
|
||||
, M. Sirino (
|
||||
<collectionCode id="EFE74399C624DE29B3B041D3724CDB97">PZ</collectionCode>
|
||||
)
|
||||
<date id="EB8E4F0D6F51CCBBF3CFB330E366505E" value="1997-11">XI. 1997</date>
|
||||
<quantity id="827450D25D5AD7CDA416A0D306F346C8" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400–1700 m</quantity>
|
||||
Leg. F. Angelini (
|
||||
<abbrev id="ABBRID0EWMBM" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="D1EC72BE4BC8DCEC72E6D12A73001325">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1MBM" type="Differential diagnosis">
|
||||
<paragraph id="72D449939F13A4E77789A8A34AEFFB66">
|
||||
<heading id="5065E843A75101DCA60B34C6867F6907" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7F2BF7C24C8A7515FDA3A96C6560D4C6">
|
||||
<taxonomicName id="091C72770811D99D4790119A815B1C8E" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix" status="sp. nov.">
|
||||
<emphasis id="56F95B398A011096B3D408657944D328" italics="true">Aphodius felix</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="3CFD49801E4B7B47CEC609D47EECE7D8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
was the first new species to be recognised, after
|
||||
<taxonomicName id="A3949685F0A6F554E0FF2B8A4C699B54" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wilsonae">
|
||||
<emphasis id="3DFDC79C2A9807EEE8B052E48213EE98" italics="true">A. wilsonae</emphasis>
|
||||
</taxonomicName>
|
||||
, initially because of its small y chromosome. The endophallic teeth are large, as in
|
||||
<taxonomicName id="CEA3F7FF66B62D16C52EAA5E6933CC32" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="5C2C21B9C826F0FF20F673B1C517BDA8" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
, and the basal segment of the mesotarsi is not longer than the longer mesotibial spur. The pronotum is fairly strongly punctate and the sides bulge outwards so that the lateral margins are not visible from above either all the basal 1 / 2 or visible again in the basal 1 / 6 (Fig.
|
||||
<figureCitation id="61888955940D765E0D70EDD63D1C1EE2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 i</figureCitation>
|
||||
), a character shared with Sicilian
|
||||
<taxonomicName id="EC6E8FF7EF373E5B6CF63400D0A97CDD" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens">
|
||||
<emphasis id="19B73CCAA5FB912B08C35CA7B0033707" italics="true">A. bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="77573D98B5E3E67A82C58A8BF4360709" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 j</figureCitation>
|
||||
). Chromosomally, these two species are clearly distinguished by their X chromosomes, the X being clearly the longest in the nucleus in
|
||||
<emphasis id="869E17152D76F62B8BC0697583EBF2EF" italics="true">A.</emphasis>
|
||||
(
|
||||
<taxonomicName id="38FF58B2E350F1D931AAC2F0C7617E1F" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens">
|
||||
<emphasis id="C23CFD64F6C29BD465B4540351B919F8" italics="true">bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
Fig.
|
||||
<figureCitation id="352E6EE4D8D346C85BA64A7A0A8B9B07" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure32" httpUri="https://binary.pensoft.net/fig/1097519">32 f – i</figureCitation>
|
||||
), but clearly shorter than autosome pairs
|
||||
<quantity id="062591A68A98C7E73BC55CDB4ADE7D15" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
|
||||
<taxonomicName id="5FAAA54A1E304EB89E8781AC5C5C2B91" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="061CB072A48AB1D1260DB93F0FB900AA" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="DCF4D682CDCC996A80D0CF3CCEC74155" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure32" httpUri="https://binary.pensoft.net/fig/1097519">32 a – e</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EYPBM" type="redescription">
|
||||
<paragraph id="4156E12CFFC461857BAE066FF4C04615">
|
||||
<heading id="E365DCEA08A19DD2C7105DC0DA788242" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7951728FB5FCF6A86953477AFA5A4DDA">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="8AC19B6EE1B999D23CCAFFCFED9BCBC3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 c</figureCitation>
|
||||
). Length:
|
||||
<quantity id="C91B944DEBF671EC407B2FF5A0C942FE" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
|
||||
(
|
||||
<specimenCount id="294D289BB9203E430FD41C13757AFF6D" count="1" type="male">♂</specimenCount>
|
||||
), 4.2–5.0 mm (
|
||||
<specimenCount id="005117C61465A983BF8A2E5941B3E64E" count="1" type="female">♀</specimenCount>
|
||||
); width: 1.9–2.0 mm (
|
||||
<specimenCount id="6CDCC66639097EA090C9579D5464CD4A" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="D771A1919A8291A0831F5C813F5BB0A6" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
|
||||
(
|
||||
<specimenCount id="4815FBAF090290B74B1AB4AF661388F8" count="1" type="female">♀</specimenCount>
|
||||
). Glossy black, tarsi, and tibial spurs dark brown, tibiae with a hint of brown-bronze reflections.
|
||||
</paragraph>
|
||||
<paragraph id="164CF78367117F5F26001D4A6A4849AC">
|
||||
Head with dorsal surface domed, flatter behind frontoclypeal suture. Frontoclypeal suture distinct, completely non-tuberculate but with smooth elevated areas each side of the median straight section and a weaker elevation medially. Genae rounded, strongly protuberant laterally in front of eyes. Clypeus with rounded median emargination, angles either side of this rounded. Clypeus and genae with strong raised margin, this slightly brownish. Surface strongly and more or less evenly punctuate, the punctures separated by ca 1.5 × their diameter, and finer medially. Antennae and palpi more or less black. Epipharynx (Fig.
|
||||
<figureCitation id="B7772740423C980263D3FEECD8EBADA7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 e, f</figureCitation>
|
||||
) clithra evenly excised either side of the median tylus, chaetopedia with four or five rather stout spines and spines of chaetopariae mainly shorter than chaetopedia. Head and pronotum as in Fig.
|
||||
<figureCitation id="F350FFCCBD22A7E311CB2BD7A5E285AC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 i</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="084D1132E745DF0777AD8C64E4DAF755">
|
||||
Pronotum hemicylindrical, highly arched transversely but more or less flat longitudinally, lateral margins more or less parallel, but slightly convergent anteriorly, weakly and very evenly curved. Pronotal surface bulging outwards from the general curvature over all or part of the basal 1 / 2 of the pronotum, ca 1 / 4 of the way in from each side, so that the lateral margins are not visible from above either in all of the basal 1 / 2 of the pronotum, or visible in the basal 1 / 6 (Fig.
|
||||
<figureCitation id="3C9C054CDC9DAC38894344C20B8890FC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 i</figureCitation>
|
||||
). Lateral margins with distinct raised border, this continued very finely over the lateral 1 / 3 of the basal margin, the median 1 / 3 of which normally lacks any border Occasionally the median 1 / 4 of the basal margin is bordered but on either side of this is an unbordered section amounting to one 1 / 4 of the length of the basal margin (Fig.
|
||||
<figureCitation id="995EFF35BDD8A4094E9944A7C8175764" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." figureDoi="10.3897/zookeys.1207.117225.figure15" httpUri="https://binary.pensoft.net/fig/1097476">15 d</figureCitation>
|
||||
). Anterior margin without any trace of a raised border. Surface with double punctation of variable strength. Larger punctures separated by 2–4 × their diameter, sometimes very strongly impressed and with the pronotal surface depressed immediately round their edges, but sometimes more moderate and with the pronotal surface evenly curved around them. Finer punctures dot-like, in some specimens separated from one another by ca 2 × their diameter but in other specimens much sparser, separated by 4–6 × their diameter. Pronotal surface between the punctures smooth, glossy. Scutellum: elongate, pentagonal, ca 10 % of elytral length, glossy black with brownish lateral and apical edges, and with sparse punctures medially. Elytra: glossy black but the interstices slightly duller than head and pronotum and with fine isodiametric reticulation (visible at × 40 magnification with bright diffuse illumination) (Fig.
|
||||
<figureCitation id="1C552BAAE34DA90E1B3C34110816D804" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." figureDoi="10.3897/zookeys.1207.117225.figure16" httpUri="https://binary.pensoft.net/fig/1097478">16 v</figureCitation>
|
||||
). Striae narrow (ca 1 / 5 the width of the interstices), vertical sided and with punctures separated by ca 2 × their diameter. In the
|
||||
<specimenCount id="95BB9D40D9EBCD6BADF3DE4C632926A3" count="2" type="male">two ♂♂</specimenCount>
|
||||
the punctures bulge into the interstices but in the
|
||||
<specimenCount id="4C3304A0CA8D0A858ACDD3B9ABDB5D5B" count="3" type="female">three ♀♀</specimenCount>
|
||||
they hardly deflect the strial margins. Lateral margins of the elytra distinct, at base strongly upcurved in front of the humeral bulges; stronger basally and at apex ca as wide as stria 2. Interstices 4 × as wide as striae, with fine sparse punctation, this a bit stronger in the humeral area in front of abbreviated stria 9.
|
||||
</paragraph>
|
||||
<paragraph id="73A468D9D516651C556367ECDBBB15D8">
|
||||
Metaventrite: median diamond-shaped plate fairly strongly punctate, concave to the depressed mid-line (
|
||||
<specimenCount id="8E8FA00134CCF16F3E55F2CF745A25FC" count="2" type="male">2 ♂♂</specimenCount>
|
||||
) (Fig.
|
||||
<figureCitation id="2DB11054A8E5366892813992B031AE5C" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 f, k</figureCitation>
|
||||
), flat with faint median impressed line to median concavity over middle 1 / 3 of the diamond-shaped area (
|
||||
<specimenCount id="40C4ED9019CEE4B8AC4042E5ED3AAD78" count="3" type="female">3 ♀♀</specimenCount>
|
||||
). Legs: black with dark brown tarsi and tibial spurs, rather long and slender, basal segment of hind tarsi as long as segments 2 + 3 + 1 / 2 of segment 4. Longer spur of mid tibiae longer than basal segment of mid tarsi, though sometimes the segment is almost as long as the spur (Fig.
|
||||
<figureCitation id="4CD98156FF181EA8009B4A7ECEB220EA" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 p, q</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4CB7F104A909D7B2F785A7631CD38422">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="0E1FC38732B74662027D0F5CD4E413E1" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 v, w</figureCitation>
|
||||
): size and shape of parameres as
|
||||
<taxonomicName id="1ED1CC5E3C7DADF9949E7DFB47B41339" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="4C2DEF92F2A99E9293FA20A9D673500B" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
, and spines of endophallus strong as in
|
||||
<taxonomicName id="B67E9D2A7E917F23D5C9B40B56005F8F" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||||
<emphasis id="63503324E3830E1A3A28610F82250DDC" italics="true">A. niger</emphasis>
|
||||
</taxonomicName>
|
||||
. Longest teeth on endophallus 45 μm long.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EASBM" type="etymology">
|
||||
<paragraph id="6DB3C565680E7A9412896953CD70D978">
|
||||
<heading id="6A15D3B2CE01CFE2BB8FD51915AE5C53" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="CFE5CF4799D32A894FEE810F79D9BB86">
|
||||
<taxonomicName id="9EF2E4D566440F70233D535FBBEFDFAF" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="79970CE9DC165ED1BA3C5A46DB70FAD2" italics="true">felix</emphasis>
|
||||
</taxonomicName>
|
||||
– Latin, happy, named from the
|
||||
<typeStatus id="0C5862F312C30C0CDDA8A9B1E5F0A275">type</typeStatus>
|
||||
locality, Campo Felice.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EQSBM" type="remarks">
|
||||
<paragraph id="F1790E9E4517E300B8CCF43C4CBF552C">
|
||||
<heading id="B5FB9EBC2FD444653DA7D300A7F5EA8E" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="85883D89F862E276B85DE37857BB0AD8">
|
||||
As yet this species is known only from the
|
||||
<typeStatus id="023BD21B8AC4A4C80D6C54B22945D8ED">type</typeStatus>
|
||||
series. The beetles were taken at the edge of a pool below a grassy slope on Campo Felice (Fig.
|
||||
<figureCitation id="82A8A0526DEB9EDCBFC40324627B9454" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||||
) in the
|
||||
<collectingRegion id="8EB2562E5AA9CC9DB8B94117CFC4238E" country="Italy" name="Abruzzo">Abruzzo</collectingRegion>
|
||||
mountains (Fig.
|
||||
<figureCitation id="EAB2ED609A9BEDFA829D246F39EC0A0E" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 f</figureCitation>
|
||||
) after a day of continuous heavy rain. A return visit a day or so later, after better weather, yielded no further specimens, so that it seems that this species lives at the roots of the grass and other herbs rather than in the mud at the water’s edge. Specimens from northern and central
|
||||
<collectingCountry id="38E23B0AD8CA369A68309EBD51A6D024" name="Italy">Italy</collectingCountry>
|
||||
mentioned in
|
||||
<bibRefCitation id="2D0336BEB00934D29CAF481DCD7F3D07" author="Maté JF & Angus RB" firstAuthor="Mate" journalOrPublisher="Koleopterologische Rundschau" pagination="325-336" refId="B33" refString="Maté JF, Angus RB (2005) Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky. Koleopterologische Rundschau 75: 325–336." title="Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky." volume="75" year="2005">Maté and Angus (2005)</bibRefCitation>
|
||||
are most likely
|
||||
<taxonomicName id="223489B334DCAACF56FBCD41A0A5213C" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="felix">
|
||||
<emphasis id="875B000880F351F53222267F8FDF22D1" italics="true">A. felix</emphasis>
|
||||
</taxonomicName>
|
||||
. See Remarks after
|
||||
<taxonomicName id="B249F9CB9DB7129E2628A77530E2682E" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bellumgerens">
|
||||
<emphasis id="D19E55E88FB6739D729F111ED90EE787" italics="true">A. bellumgerens</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
596
data/F7/A4/18/F7A418809EFA50C998DBBED70B4A40A9.xml
Normal file
596
data/F7/A4/18/F7A418809EFA50C998DBBED70B4A40A9.xml
Normal file
|
|
@ -0,0 +1,596 @@
|
|||
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<mods:mods id="019D4EC0EF9089AB5E1AC0283098C191" xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo id="689E4616FC0758764708D1DFA87B61A5">
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<mods:title id="63E29660DB1984CA34100BB771C828AC">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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|
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<mods:name id="EC4FF0487168E08AD99446E07D5957CC" type="personal">
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<mods:role id="30F54F82221D8E4245AAA9D82387B0D1">
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|
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<mods:nameIdentifier id="43797B71167F7DC5A05069BB2D6ED4D8" type="ORCID">0000-0002-3860-5617</mods:nameIdentifier>
|
||||
<mods:affiliation id="E5F2C02FD5616097C3069B4B9A21EA4B">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
|
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</mods:name>
|
||||
<mods:name id="4DCAC8AB771E36C7565BD8BF5139C5B0" type="personal">
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<mods:namePart id="029D398B17FA152467C7ECADCAD1DEF9">Maté, Jason F.</mods:namePart>
|
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<mods:nameIdentifier id="9ABAB783E15C5E62E0FA69736BF7C7CA" type="ORCID">0000-0002-2505-7113</mods:nameIdentifier>
|
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<mods:affiliation id="72CA1EC3AC320D4654BFFF89951B9779">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="B8090F20C381A42EE629BF256BE4F011">Angus, Elizabeth M.</mods:namePart>
|
||||
<mods:affiliation id="29139F832EB8E375BCEB2AD87BEC00DE">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
|
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</mods:name>
|
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</mods:role>
|
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<mods:namePart id="E7AB5E9F144F6E77D5014D4132D97BAC">Král, David</mods:namePart>
|
||||
<mods:nameIdentifier id="768BE5A699E32A022942142CD9049B58" type="ORCID">0000-0002-7930-1927</mods:nameIdentifier>
|
||||
<mods:affiliation id="228DDC62E1AD531FA5590E42E5C12C6F">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:title id="8684C5459A8BE89AA0A146969B9737EC">ZooKeys</mods:title>
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<mods:part id="9EB62D189DCD906C4323E691C2407820">
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<mods:date id="298C90BAA3F3170A52BCE0D770E1C6BA">2024</mods:date>
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<mods:number id="09F09267D5E85594558755BE8E008D92">2024-07-22</mods:number>
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<mods:number id="0A1C0D8F4CA7E72550C33CCC1106735D">1207</mods:number>
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<mods:start id="BD8758668E4B5F4B2FB042BABE1AE251">205</mods:start>
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<mods:identifier id="A8220655B298CBEAD8D1E7775E707AFE" type="DOI">10.3897/zookeys.1207.117225</mods:identifier>
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<mods:identifier id="D12A551D9BACABFF2B5EE433280748FC" type="ZooBank">94F18819-5AF5-4100-AB35-AA3C3976EE80</mods:identifier>
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<treatment id="F7A418809EFA50C998DBBED70B4A40A9" ID-arpha="F7A41880-9EFA-50C9-98DB-BED70B4A40A9" LSID="urn:lsid:plazi:treatment:F7A418809EFA50C998DBBED70B4A40A9" httpUri="http://treatment.plazi.org/id/F7A418809EFA50C998DBBED70B4A40A9">
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<subSubSection id="8EF979D66B7C26550E695B9270CC44DE" type="nomenclature">
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<taxonomicName id="A9EA3E9FD4406026B044FD9CA9F03ECA" ID-CoL="FHZY" authority="Baudi di Selve, 1870" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis" subGenus="Liothorax">
|
||||
<emphasis id="75528E78C8F978F6E8BE59A0F9A7CC59" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="3EDDF0EBB21174A2A48E12D2C9EFC7ED" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="F8191FA9FFF3204F3F31F02053B4943B" italics="true">rutilipennis</emphasis>
|
||||
<bibRefCitation id="2CB5F05DDFF4C8D14B7A475393B099A7" DOI="10.1002/mmnd.18710150113" author="Baudi di Selve F" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="49-90" refId="B7" refString="Baudi di Selve F (1870) Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia. Berliner Entomologische Zeitschrift 14: 49 - 90. https: // doi. org / 10.1002 / mmnd. 18710150113" title="Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia." volume="14" year="1870">Baudi di Selve, 1870</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, stat. rest.
|
||||
</paragraph>
|
||||
</subSubSection>
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<subSubSection id="BD77DFDB786FC7323D63E8A0692100AC" type="description">
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<paragraph id="D0204410C7D951AA387263326B776AC4">
|
||||
<figureCitation id="BA010CDC3DB9D1F344119A1FA4618038" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">Figs 2 e – g</figureCitation>
|
||||
,
|
||||
<figureCitation id="F717FCE7148778DF5FDB3F2125C4C4D1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 i – k</figureCitation>
|
||||
,
|
||||
<figureCitation id="2FA8B566E7F52CD7CF769724E4B125D2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 j</figureCitation>
|
||||
,
|
||||
<figureCitation id="BF1C9F4CF035E0D1590352CF3E29D7DA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="4031853E658EBA1FA338B4F7915047D7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 c</figureCitation>
|
||||
,
|
||||
<figureCitation id="1E9BEF9310AD7AFFDC930ECC2D30A274" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 c, d</figureCitation>
|
||||
,
|
||||
<figureCitation id="618A2044DA17900B07D679A5059DADDA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">
|
||||
22 q, q ’, r, v, w,
|
||||
<quantity id="420EC430157008BCB3FB873949E74024" metricMagnitude="1" metricUnit="m" metricValue="2.5" unit="m" value="25.0">25 m</quantity>
|
||||
, 26 j, k, 30 d, e
|
||||
</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="055CFAD718260EF125D6F67DE6DF31E4" type="reference_group">
|
||||
<paragraph id="2B0A0BED274A82EE1251980CA8B32C11">
|
||||
<treatmentCitationGroup id="059E69539B93B97F86B9605FDC87F004">
|
||||
<treatmentCitation id="569225BA5447844EADCDE5B5E95D550C" author="Baudi di Selve F" page="64" year="1870">
|
||||
<taxonomicName id="D37F6D0223A6E176002CF1578D612439" ID-CoL="FHZY" authority="Baudi di Selve, 1870: 64" authorityName="Baudi di Selve" authorityPageNumber="64" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="variety" species="plagiatus" subGenus="Liothorax" variety="rutilipennis">
|
||||
<emphasis id="D1BD155BBBE7524534CE0CFBFF723BFB" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="EC44A30AA433A5CB5B970E34BDB260D1" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="1EB3DD834ECB6024610EBE903A558639" italics="true">plagiatus var rutilipennis</emphasis>
|
||||
<comment id="02D48255C144CD578948A363E8DA9D7E">
|
||||
<bibRefCitation id="E61695F72406C1A1253089B941BE4405" DOI="10.1002/mmnd.18710150113" author="Baudi di Selve F" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="49-90" refId="B7" refString="Baudi di Selve F (1870) Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia. Berliner Entomologische Zeitschrift 14: 49 - 90. https: // doi. org / 10.1002 / mmnd. 18710150113" title="Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia." volume="14" year="1870">Baudi di Selve, 1870: 64</bibRefCitation>
|
||||
.
|
||||
</comment>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="54A36869AC3DCCD0B39C9871208CF745">
|
||||
<treatmentCitationGroup id="3D909031C3DA015A8041533602E6C758">
|
||||
<treatmentCitation id="0F9A9809A72D6BF7EA09657092A55922">
|
||||
<taxonomicName id="577148B6CCE5A1E7672DBF843DCC0D8A" ID-CoL="FG46" authority="Petrovitz, 1962: 126" authorityName="Petrovitz" authorityPageNumber="126" authorityYear="1962" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ressli">
|
||||
<emphasis id="D19864974FE556A4B08F6FF3E2A53955" italics="true">Aphodius ressli</emphasis>
|
||||
Petrovitz, 1962: 126
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="A902237D155D00665EB98C984FD2F81C">
|
||||
<treatmentCitationGroup id="C25D543F95DB96A9A639950A9E166411">
|
||||
<treatmentCitation id="77EBC2E0729E557F6CF89CD362D78CCD">
|
||||
<taxonomicName id="D1DACD0BF1E40D314915F69B666DD112" ID-CoL="FG44" authority="Balthasar, 1971: 57" authorityName="Balthasar" authorityPageNumber="57" authorityYear="1971" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cypricola">
|
||||
<emphasis id="37668F2386C97723564F594ABFDCF21E" italics="true">Aphodius cypricola</emphasis>
|
||||
Balthasar, 1971: 57
|
||||
</taxonomicName>
|
||||
.
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="0DFC395CEBCEFE28C417B1CFAD005600">
|
||||
<treatmentCitationGroup id="8CC49774A19607B66746BD61E8171AD4">
|
||||
<treatmentCitation id="4A880178F47D70FFF5E91ABC3C3E8302">
|
||||
<taxonomicName id="64726A43F95CCA85A4F7F98AC351B386" ID-CoL="FG45" authority="Balthasar 1952" authorityName="Balthasar" authorityYear="1952" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="isikdagensis">
|
||||
<emphasis id="2B904273ED4DF35AC49D6B49BC971B5E" italics="true">Aphodius isikdagensis</emphasis>
|
||||
<comment id="328FA52D1D3407CBF94F09C90080FA6E">
|
||||
Balthasar 1952, sensu
|
||||
<treatmentCitation id="B146240DD59D96DF809067D717616F07" author="Dellacasa G & Dellacasa M & Ziani S" page="6" year="">
|
||||
<bibRefCitation id="DB6FF994E0633356334ECF46D51B82B8" author="Dellacasa G & Dellacasa M & Ziani S" issue="2000" journalOrPublisher="Caffi ”" pagination="3-11" refId="B14" refString="Dellacasa G, Dellacasa M, Ziani S (2001 b) Taxonomic position of Aphodius turbatus Baudi, 1870 and systematic redefinition of Aphodius isikdagensis Balthasar, 1952. Rivista del Museo civico de Scienze naturali “ E. Caffi ” 20 (2000): 3–11." title="Taxonomic position of Aphodius turbatus Baudi, 1870 and systematic redefinition of Aphodius isikdagensis Balthasar, 1952. Rivista del Museo civico de Scienze naturali “ E." volume="20">Dellacasa et al. 2001 b</bibRefCitation>
|
||||
: 6
|
||||
</treatmentCitation>
|
||||
(misidentification).
|
||||
</comment>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6ZGK" type="notes">
|
||||
<paragraph id="ED99FCAA427883BC011BAEB5358E5A5A">
|
||||
<heading id="8C3CCF9039041DC8234AC7A48B61D337" reason="title">Note.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5081799E06910877CF007A3974D94250">
|
||||
<bibRefCitation id="7314E19EEA6EA61F2623967B1E2CE7AB" DOI="10.1002/mmnd.18710150113" author="Baudi di Selve F" firstAuthor="Baudi di Selve" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="49-90" refId="B7" refString="Baudi di Selve F (1870) Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia. Berliner Entomologische Zeitschrift 14: 49–90. https://doi.org/10.1002/mmnd.18710150113" title="Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia." volume="14" year="1870">Baudi di Selve (1870)</bibRefCitation>
|
||||
described a variety of
|
||||
<taxonomicName id="D57B1FA72CEA9DBCF78ADBE537C02B4E" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="1DE8A51EE935B72C83C9161525226BDE" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
in which the elytra were brownish bronze (“ brunneo-subaeneis ”) and said that this variety was less rare (on
|
||||
<collectingCountry id="21BBE46D544A80D47E0EBE3F114D668F" name="Cyprus">Cyprus</collectingCountry>
|
||||
) than the black form. He added that, among other things, the mesosternum (sic) was more delicately and less densely punctate than in European
|
||||
<taxonomicName id="9B4E8CA1844D2A2F1D109A6D407D6ED8" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="51731A4A8B8171F370B2BE1511E65641" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation id="4D7287A595E5D0487490BBBC262E0F18" DOI="10.11646/zootaxa.1407.1.4" author="Dellacasa M & Gordon RD & Dellacasa G" etAl="et al." firstAuthor="Dellacasa" issue="1" journalOrPublisher="Zootaxa" pagination="23-42" refId="B15" refString="Dellacasa M, Gordon RD, Dellacasa G (2007) Systematic review to the genus Liothorax (Coleoptera: Scarabaeoidea: Scarabaeidae: Aphodiinae). Zootaxa 1407 (1): 23–42. https://doi.org/10.11646/zootaxa.1407.1.4" title="Systematic review to the genus Liothorax (Coleoptera: Scarabaeoidea: Scarabaeidae: Aphodiinae)." volume="1407" year="2007">Dellacasa et al. (2007)</bibRefCitation>
|
||||
stated that the name
|
||||
<taxonomicName id="22BE832F292B84A8735C3835FEC75BC8" authority="Baudi di Selve" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="66B33EF6D847015BA507AA7EB8903580" italics="true">A. rutilipennis</emphasis>
|
||||
Baudi di Selve
|
||||
</taxonomicName>
|
||||
was unavailable as it was simply a colour variant, but that would only be the case if the name was “ infrasubspecific ”. For names published before 1961, the International Code of Zoological Nomenclature (1999, 4
|
||||
<superScript id="3A3B07C54A0BA21DED98D4A70FA740A3">th</superScript>
|
||||
edition), states that a name “ is subspecific if first published before 1961 and the author expressly used one of the terms “ variety ” or “ form ” (including the use of the terms “ var. ”, “ forma ”, “ v ” and “ f ”), unless the author also expressly gave it infrasubspecific rank, or the content of the work unambiguously reveals that the name was published for an infrasubspecific entity ”. Baudi makes no reference to infrasubspecific status, nor does his description imply that it was such. This name is therefore available and is the oldest known name for this species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EQ2GK" type="type material">
|
||||
<paragraph id="D8F0A2CF835607111579A17603EF121E">
|
||||
<heading id="A380C53D2B19439E1C75088C0518E174" reason="title">
|
||||
<typeStatus id="D27B2372AF6758B5E34FA6AE0D563878">Type</typeStatus>
|
||||
material examined.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E60DE03C638D682D4B084A9AAC4DC3A1">
|
||||
<taxonomicName id="8CFC955DB07485D5F699BEA48267990A" authority="Baudi di Selve, 1870" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="variety" species="plagiatus" variety="rutilipennis">
|
||||
<emphasis id="FADA367711F832B0C888D9C5908E5BFC" italics="true">Aphodius plagiatus var rutilipennis</emphasis>
|
||||
<bibRefCitation id="8569E17689C0FD144D2A98353E024727" DOI="10.1002/mmnd.18710150113" author="Baudi di Selve F" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="49-90" refId="B7" refString="Baudi di Selve F (1870) Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia. Berliner Entomologische Zeitschrift 14: 49 - 90. https: // doi. org / 10.1002 / mmnd. 18710150113" title="Coleoptorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio de Europaeis notis quibusdam additis. Pars tertia." volume="14" year="1870">Baudi di Selve, 1870</bibRefCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<typeStatus id="B59239E50CDC633C356C767450BDB930">Type</typeStatus>
|
||||
material in the Museo Civico di Storia Naturale Giacomo Doria, Genova (MCSNGDG) – not examined (not available for study).
|
||||
</paragraph>
|
||||
<paragraph id="B8430B88835683F6D2978F1D825F2416">
|
||||
<taxonomicName id="AD98F9CAD0AC721B770518FACBA768FA" authority="Petrovitz, 1962" authorityName="Petrovitz" authorityYear="1962" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ressli">
|
||||
<emphasis id="9BC450F5685384C11FFE28587D675CD1" italics="true">Aphodius ressli</emphasis>
|
||||
<materialsCitation id="C829099B85BA8D9A12EB926572F7264F" collectionCode="MNHG" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||||
Petrovitz, 1962.
|
||||
<typeStatus id="34C10340072ABF3A83C2EAFAA619118A" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="86ACA88AC500C860A7A8D6A0D5A15639" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EU3GK" xlink_title="Muséum d’histiore naturelle, Genève">
|
||||
<collectionCode id="5C63316EE89220FE40306C68B5AB80B3">MNHG</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
</materialsCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph id="60A0ED7568E7E13EA64FD9ACC63C1109">
|
||||
<taxonomicName id="0EB327B1C3AD10F5A9EB44545E24451B" authority="Balthasar, 1971" authorityName="Balthasar" authorityYear="1971" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cypricola">
|
||||
<emphasis id="880F284483AA7987ED172F79B6323976" italics="true">Aphodius cypricola</emphasis>
|
||||
<materialsCitation id="7910104B90A38ACE894F6A897FCC0E6D" collectionCode="NMP" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
Balthasar, 1971.
|
||||
<typeStatus id="951E58D77891F9EEAA1422ACF862DF35" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="3B0CA3EA1A6561C7CD86484AADA4370C" count="1" type="male">♂</specimenCount>
|
||||
(
|
||||
<named-content id="6821BD1E61772F1B14470ED58221E7CC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="27952CF93FB72DF677ABF92291F619E6" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</materialsCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJ4GK" type="material">
|
||||
<paragraph id="658446029439CCCB028BDB8B5B41CC73">
|
||||
<heading id="B3406A54492E4820ECE0C82F642C76EF" reason="title">Additional material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1891897A8BCB5AB4EBD966F53896CEAE">
|
||||
GR: Corfu,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ES4GK" specific-use="{"type":"Point","coordinates":[19.800000,39.600000]}">
|
||||
<geoCoordinate id="AC640454E783CD29754FC658707B69E0" degrees="39.6" direction="north" orientation="latitude" precision="5555" value="39.6">39.6 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="0DFE1251E1B9F78618559D0C37D581EB" degrees="19.8" direction="east" orientation="longitude" precision="5555" value="19.8">19.8 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Reitter, Nevinson coll.
|
||||
<specimenCount id="37E778601A69995953D2C1F70FCA0983" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<named-content id="644285C697FC5C96B829E3B915D7F019" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="CDAD52C9F3B24BB38FDDDEF770CA6D59" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). Evia, Chalcis,
|
||||
<geoCoordinate id="C3A0A560BE397119BC9E4FB4B0BA513C" degrees="36.463" direction="north" orientation="latitude" precision="55" value="36.463">36.463 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="87A1A44882E977F1625BC7DEDE2BC4B4" degrees="26.635" direction="east" orientation="longitude" precision="55" value="26.635">26.635 ° E</geoCoordinate>
|
||||
, 1932, leg. J. Fodor. 1 unsexed (
|
||||
<abbrev id="ABBRID0E24GK" xlink_title="Hungarian Museum of Natural History">
|
||||
<collectionCode id="072CB363DFFB65EAEDA950B8E949075D">MNHB</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
<collectingCountry id="53B2D5509850D7765D3F9DC10CB6CA2A" name="Macedonia">Macedonia</collectingCountry>
|
||||
, Langassa Göll.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EC5GK" specific-use="{"type":"Point","coordinates":[32.147000,40.464000]}">
|
||||
<geoCoordinate id="D6F01A651763C9EB1961859AE5DEFB0B" degrees="40.464" direction="north" orientation="latitude" precision="55" value="40.464">40.464 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="662855330C4277752534CEBAB15BF6E1" degrees="32.147" direction="east" orientation="longitude" precision="55" value="32.147">32.147 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="25268B8A76D38905F1CABAB002C91CC3" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EH5GK" xlink_title="Hungarian Museum of Natural History">
|
||||
<collectionCode id="B4D52CE33E49650AB42A0ECA159D1E9A">MNHB</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="E0AE4C9E6F4638A9BBF1B32BFA418842">
|
||||
AL:
|
||||
<collectingCountry id="CA375C8DF7FDFEE4383C8490FBED567F" name="Albania">Albania</collectingCountry>
|
||||
mer. Butrint,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQ5GK" specific-use="{"type":"Point","coordinates":[20.021000,39.754000]}">
|
||||
<geoCoordinate id="828F46FA764CC18C7C117F72B94D42AD" degrees="39.754" direction="north" orientation="latitude" precision="55" value="39.754">39.754 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="0A9CFA45DF0E66F0A85C22D29B44C61D" degrees="20.021" direction="east" orientation="longitude" precision="55" value="20.021">20.021 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Smetana, 1958, l
|
||||
<specimenCount id="33F44A0F48DB3793F96C0BA55F03C1AB" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<named-content id="963A504D05AB2AC261ADDF6C3E85ECD4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="348FC2416FD9D790A5E0300C24B73E8C" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="D764FB5DECFF71919F69790A601C3A70">
|
||||
TR: Besika (Beşik) Bay,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E55GK" specific-use="{"type":"Point","coordinates":[26.459000,40.274000]}">
|
||||
<geoCoordinate id="8A3C7249B9A233C5BDF2125504121C3D" degrees="40.274" direction="north" orientation="latitude" precision="55" value="40.274">40.274 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D9D10D161B326B5339B4B91530978841" degrees="26.459" direction="east" orientation="longitude" precision="55" value="26.459">26.459 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="AA61D0A022554DE7F5504612CA0C113C" count="2" type="male">2 ♂♂</specimenCount>
|
||||
, 5 unsexed. Ex C. G. Champion (
|
||||
<named-content id="F30DEABAE23FB662BC2110C9324CFE6D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="33684D2335A03600D49F45277147EB0D" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
<collectingCountry id="D93C2863B49F4939B86E4111BC441DB0" name="Turkey">Turkey</collectingCountry>
|
||||
mer. Devecusagi, Yumurtalik env.,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK6GK" specific-use="{"type":"Point","coordinates":[36.039000,36.884000]}">
|
||||
<geoCoordinate id="1DF17BA8FA4830BA8FF4434700612FC9" degrees="36.884" direction="north" orientation="latitude" precision="55" value="36.884">36.884 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="5924F3938D024865B569495D18AC58AF" degrees="36.039" direction="east" orientation="longitude" precision="55" value="36.039">36.039 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="7C7CB93282E2DA66FB6DB7026A9238DA" value="1992-04-14">14. iv. 1992</date>
|
||||
. leg. O. Hovorka. 1 unsexed (
|
||||
<named-content id="75243D1139928BFE7AE2AC0893D7492C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="A29E82E6F8D3D780C820C304B059A2D2" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
) prov.
|
||||
<collectingRegion id="EF30587B1370BD93619351C9A3AB5BFF" country="Turkey" name="Antalya">Antalya</collectingRegion>
|
||||
, Manavgat.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EW6GK" specific-use="{"type":"Point","coordinates":[31.437000,36.763000]}">
|
||||
<geoCoordinate id="B552E24B607EF51B4756FC28F12A26C2" degrees="36.763" direction="north" orientation="latitude" precision="55" value="36.763">36.763 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="C76CF180CD6CDD5615A3F8695379AF2F" degrees="31.437" direction="east" orientation="longitude" precision="55" value="31.437">31.437 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. leg.
|
||||
<tp_taxon-name id="7AB478F6A961A6B5BB281FC2A19A361B">
|
||||
<emphasis id="8C08207B22E5047D4AA05C538304C4A6" italics="true">
|
||||
<tp_taxon-name-part id="CA8565D56740953DC740C96968950475" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Bellmann
|
||||
<date id="F75832CFC1740352C92A36FEE1BE15D3" value="2000-03-13">13. iii. 2000</date>
|
||||
.
|
||||
<specimenCount id="587CB966A182BABE01403078316114DD" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EBAHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="46F1F579BC912801AD303547B281ECCA">ABC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
<collectingRegion id="8AC44297B76D167B041EC6632A7DE871" country="Turkey" name="Antalya">Antalya</collectingRegion>
|
||||
, Manavgat.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIAHK" specific-use="{"type":"Point","coordinates":[31.437000,36.763000]}">
|
||||
<geoCoordinate id="C960B381AABEEB5C789CFFAC6CFEF0B5" degrees="36.763" direction="north" orientation="latitude" precision="55" value="36.763">36.763 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1478172AC5C2D499715A549EF7D0462F" degrees="31.437" direction="east" orientation="longitude" precision="55" value="31.437">31.437 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. leg.
|
||||
<tp_taxon-name id="372FB9A1893E2AAEA290984E2195AE6F">
|
||||
<emphasis id="AA95D3A38A411873532B3A1C9766B75E" italics="true">
|
||||
<tp_taxon-name-part id="71CDE2878DE2148ADA59FC856F2C1F00" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Bellmann
|
||||
<date id="DAEA818AB874521DB3A1F42678593B4D" value="2000-03-13">13. iii. 2000</date>
|
||||
.
|
||||
<specimenCount id="3DE7B1D695ACFFF3DE6D84A414263084" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0ETAHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="4FC1E6C45D5DACC6505117AC7C252977">ABC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
<collectingRegion id="626A11DD8139DEE78204A0F67DC70455" country="Turkey" name="Antalya">Antalya</collectingRegion>
|
||||
, 5 Km nördi, Sagirin, Köprülu kanyon,
|
||||
<date id="A1B0A2AE04B53F389933629BF68D8CDB" value="2002-03-19">19. 03. 2002</date>
|
||||
, leg.
|
||||
<tp_taxon-name id="5017D2FA5B5F1ACB608EFC68422537F0">
|
||||
<emphasis id="123A4212ABE9A4206FBCC4F7A3645AA4" italics="true">
|
||||
<tp_taxon-name-part id="78487CE0C35AD372DF2C8099C9B5A00B" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Bellmann, 1 unsexed (
|
||||
<abbrev id="ABBRID0E4AHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="87D2DCAB8B7149B8381CA154B08F7E63">ABC</collectionCode>
|
||||
</abbrev>
|
||||
); same data,
|
||||
<date id="BEB694D3D6970FF9C92051AEC7C7B7E3" value="2002-03-22">22. 03. 2002</date>
|
||||
, 1 unsexed (
|
||||
<abbrev id="ABBRID0EBBHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="4EA0B3A71900826CC71B2D0715574148">ABC</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
<collectingRegion id="7BFC121739F9AAE0F1EB11808F1D476C" country="Turkey" name="Antalya">Antalya</collectingRegion>
|
||||
, Akseki,
|
||||
<quantity id="686DE92115D2A7F374CC7C0468BE9697" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
|
||||
a. s. l.,
|
||||
<date id="938C9C3164D6AD3CC18CEFE22377A795" value="2002-03-19">19 March 2002</date>
|
||||
, leg. Bellmann 1 unsexed (
|
||||
<abbrev id="ABBRID0EFBHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="EF031CAADBE1A21C7B4942464A1BE674">ABC</collectionCode>
|
||||
</abbrev>
|
||||
); Alanya, Konakli;
|
||||
<date id="1CAFED8812DD85C6042E4E0BAF11006A" value="2000-03-13">13. 03. 2000</date>
|
||||
leg.
|
||||
<tp_taxon-name id="BB6146CAC2895510DC6189940F8D1FFD">
|
||||
<emphasis id="1327A74FF4738825E0CACC6595806824" italics="true">
|
||||
<tp_taxon-name-part id="9824171D85E1AC3F572E8341738EC879" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Bellmann, 1 unsexed (
|
||||
<abbrev id="ABBRID0EPBHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="C2F9138EA658C5ECA4098725B1C722F1">ABC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
<collectingRegion id="80C33DD70BAC74008088AF1831480B28" country="Turkey" name="Istanbul">Istanbul</collectingRegion>
|
||||
Halkali
|
||||
<date id="D2BE108043C89ED1D0E16A190E2FFA99" value="1968-07-28">28. vii. 1968</date>
|
||||
Cl. Besuchet, 1 unsexed (
|
||||
<collectionCode id="64D993B1AFE62DEB849562A814F0AE3C" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="B3F29A7C309642B0246FE0E0546C5A0A">
|
||||
CY:
|
||||
<collectingCountry id="1C2F5C52AB658EAA3663C448A0AC1D7D" name="Akrotiri">Akrotiri</collectingCountry>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYBHK" specific-use="{"type":"Point","coordinates":[32.991000,34.634000]}">
|
||||
<geoCoordinate id="854DDF764C799D36CF5AFF4BA88F2B9F" degrees="34.634" direction="north" orientation="latitude" precision="55" value="34.634">34.634 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1B259D3F06A0C81D422AAD3A6FB4F786" degrees="32.991" direction="east" orientation="longitude" precision="55" value="32.991">32.991 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. G. Mavromoustakis
|
||||
<specimenCount id="9032FE64FC36F26EB21C075204AB2C75" count="2" type="male">2 ♂♂</specimenCount>
|
||||
(
|
||||
<named-content id="CD178FDB4E7341FEF8999448C0CA0720" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="23A762EC733D8BE6D1842172C0D5BBF9" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
); Limassol distr. Zakaki marshes, puddle in lorry park at edge of reedbed,
|
||||
<date id="8EA66747182367B51071C2A4FCAA100A" value="2005-04-03">3. iv. 2005</date>
|
||||
. R. B. Angus.
|
||||
<specimenCount id="D3F286D6C122A513F0E185845C74E492" count="3" type="male">3 ♂♂</specimenCount>
|
||||
, chromosome prep. 4:
|
||||
<date id="3A89635547E8AF80437BA6F5E9B5137B" value="2005-04-07">7. iv. 2005</date>
|
||||
R. B Angus, DNA extracts 110 & 91, J. F. Maté 2013,
|
||||
<specimenCount id="CFCEC999C26E9C12E1C890AC677969D1" count="1" type="female">1 ♀</specimenCount>
|
||||
, chromosome prep. 5:
|
||||
<date id="FFBBE74A3B78AA7640F475C2D8453EFD" value="2005-05-07">7. v. 2005</date>
|
||||
(
|
||||
<named-content id="F9209C6D9DCF6BE50CF61F43275E6EBB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="B24A2EC8FE20FDAA2678FBDAC3CBFB8D" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
); “
|
||||
<collectingCountry id="103B3A294795A23142543C3E0420A033" name="Iceland">Insel</collectingCountry>
|
||||
Cypern ” ex coll, Fodor.
|
||||
<specimenCount id="8853A9F3EDD4E2511F79F41BE38EE983" count="2" type="male">2 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="3F773FF105B018566A7E5D133F76D821" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EFCHK" xlink_title="Hungarian Museum of Natural History">
|
||||
<collectionCode id="EB27DF08DC20D9AA40F731F9E3DFBA61">MNHB</collectionCode>
|
||||
</abbrev>
|
||||
);
|
||||
<collectingCountry id="C3FD62A3E10C8566911EBB86BC82F9A4" name="Cyprus">Zypern</collectingCountry>
|
||||
, Sotira (Salzee),
|
||||
<date id="AF75416F5DEC5B0A3774F4DA860F075E" value="2010-04-06">6. 4. 2010</date>
|
||||
, W. Ziegler leg., 1 unsexed (
|
||||
<abbrev id="ABBRID0EJCHK" xlink_title="Axel Bellmann Collection">
|
||||
<collectionCode id="31BBAB2E5CAD136E7B06BEA28D2D1345">ABC</collectionCode>
|
||||
</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ENCHK" type="Differential diagnosis">
|
||||
<paragraph id="034FA55AF23A88CDD2196A57CB6DE9C4">
|
||||
<heading id="928B6B565B26C74C9D9D19CB1C4DA1D3" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E3CAE39D01A11E5D2A6D1AE9DC74356D">
|
||||
A small black
|
||||
<taxonomicName id="9E3948A88CE3927F3CEDD2B010F46B4A" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="E3878A9976E5BD806C12F6466CCF1064" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
group species, occasionally with dull reddish elytra, immediately distinguished from
|
||||
<taxonomicName id="555DE4B6A7B49B63EB58ABB90352C8B5" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="plagiatus">
|
||||
<emphasis id="E8156B080F7C9D5F9DE067E23AD084E9" italics="true">A. plagiatus</emphasis>
|
||||
</taxonomicName>
|
||||
by its clearly more lightly sculptured metaventrite and the bristles on the endophallus. Black specimens are distinguished from
|
||||
<taxonomicName id="E9BE7E1F55FD252F9C2F734F2F978346" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chellala" status="sp. nov.">
|
||||
<emphasis id="58CDAAD426A342C0B1E9BDD979E7D0DF" italics="true">A. chellala</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="54CA98A22A7D9EA76D6A908F42243EA0" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by the narrower gap in the basal border of the pronotum.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUDHK" type="redescription">
|
||||
<paragraph id="E6EE3A6FAAABDED470AADD6A81EC512E">
|
||||
<heading id="E144A01995313A7EB167AF29F98108DF" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="132F4495F4508C7F4A26B08795F0C521">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="AD40ED417EFEBB08164A2ED42E9760C5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure2" httpUri="https://binary.pensoft.net/fig/1097460">2 e – g</figureCitation>
|
||||
). Head and pronotum (Fig.
|
||||
<figureCitation id="8C1299C105CA22E2D26E430107B9BDC5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 i – k</figureCitation>
|
||||
). Length:
|
||||
<quantity id="E057E0D5805BB76A66F53500EAAB677B" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" unit="mm" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1–4.1 mm</quantity>
|
||||
(
|
||||
<specimenCount id="2DED5523E4CE9C9342551CA19FB3CEB6" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="AA5EA892A0ACB0114BD58193B523D2CD" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.9–4.2 mm</quantity>
|
||||
(
|
||||
<specimenCount id="B784D9E4F359B95E36BC5346B3FA8D28" count="1" type="female">♀</specimenCount>
|
||||
); width:
|
||||
<quantity id="A3051D0705977352D28CB6CB824E14DA" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
|
||||
(
|
||||
<specimenCount id="A059C720701C18D4C5D02EDD1AB65455" count="1" type="male">♂</specimenCount>
|
||||
),
|
||||
<quantity id="C0AE1288ACFABA97409C38FEA786BB74" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 mm</quantity>
|
||||
(
|
||||
<specimenCount id="343007AD879B8D84D655BC5EA733E5D1" count="1" type="female">♀</specimenCount>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8988C341474036AC0040AE013B1BD74D">
|
||||
Head black, anterolateral margins of clypeus and genae with a vague dull yellowish brown strip, and actual margins narrowly raised. Frontoclypeal suture more or less straight over median 1 / 3 but middle part effaced, laterally angled to anterior edge of genae, but this part scarcely traceable. Frons with moderate double punctation which is effaced on the middle 1 / 4. Disc of clypeus bulges upwards, this bulge with only very fine sparse dot-like punctation, surface away from the bulge with moderate double punctation and the surface becoming weakly wrinkled towards the anterior and lateral margins. Front of clypeus weakly excised over middle 1 / 3, recurved either side of this excision. Antennae and maxillary palpi dull yellowish brown antennal clubs and apical segments of palpi darker. Epipharynx (Fig.
|
||||
<figureCitation id="69F6BFAAC91AFEAE37AFF1BCA7D8780B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure7" httpUri="https://binary.pensoft.net/fig/1097466">7 j</figureCitation>
|
||||
) with the anterior margin if the clithra excised either side of the median tylus, and the prophobae and apophobae quite extensively developed and with 3 or 4 chaetopedia each side.
|
||||
</paragraph>
|
||||
<paragraph id="55F4F2B6FCCF87D51630B54E70E20357">
|
||||
Pronotum hemicylindrical, highly arched transversely, weakly so longitudinally. Lateral margins bordered but basal 1 / 2 not visible from above because of the outwardly bulging lateral parts of the pronotal surface in this area. Posterior margin of the pronotum usually with an almost continuous border, this with a small median interruption whose width is approximately the same as the diameter of the larger pronotal punctures (Fig.
|
||||
<figureCitation id="1E19B64CA6F0537A2CCCE3755FD290DF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 c</figureCitation>
|
||||
). In the
|
||||
<typeStatus id="4B7BA0EF92C2A7632A03B6C58810D0F6" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="DD48C30C46F50285F69B3998FFEF0DE1" authorityName="Balthasar" authorityYear="1971" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cypricola">
|
||||
<emphasis id="2DFC91EC12F547611E9671B4046C92BA" italics="true">A. cypricola</emphasis>
|
||||
</taxonomicName>
|
||||
and a female from Greek
|
||||
<collectingCountry id="F6232BAD76A8078716A0E362A231E340" name="Macedonia">Macedonia</collectingCountry>
|
||||
(Langasa-Göl, now Lagkadas near Thessaloniki) the median gap in the basal border of the pronotum is rather wider (Fig.
|
||||
<figureCitation id="2B4643CFB9C23FDC4F1D9D8069F9FAFC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." figureDoi="10.3897/zookeys.1207.117225.figure13" httpUri="https://binary.pensoft.net/fig/1097473">13 d</figureCitation>
|
||||
), but the aedeagal characters are the same as in other material. Pronotal surface with moderate double punctation, the larger punctures separated by 1–2 × their diameter, becoming sparser medially. Smaller punctures dot-like and all punctures absent from a median longitudinal strip.
|
||||
</paragraph>
|
||||
<paragraph id="4E714988EA46854ECF66154FE3B5D6F5">Scutellum elongate, pentagonal, glossy reddish brown with a few punctures medio basally.</paragraph>
|
||||
<paragraph id="E19D93C9D4BED6620AA37B4E61420160">Elytra maroon-bronze or black, interstices with a silky sheen due to the fine reticulation on their surfaces. Striae narrow, shallow and glossy, with punctures separated by ca 2 × their diameter and indenting the strial margins. Interstices ca 6 × as wide as the striae, flat and with a very fine reticulation and scattered very fine punctures separated from one another by ca 1 / 3 of the width of the interstices. Lateral margins of elytra slightly rounded to widest point just behind the middle, then rounded to the blunt apex.</paragraph>
|
||||
<paragraph id="49BA05C7CD2B7AA5C55B00E6CFD864A2">
|
||||
Metaventrite: (Fig.
|
||||
<figureCitation id="D9FD1018601061C5F0BED488A7295630" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 c, d</figureCitation>
|
||||
) grey-bronze with maroon reflections, median diamond-shaped plate depressed medially. Surface reticulate, the reticulation effaced in the middle of the median depression. Punctation fine and sparse, becoming stronger over the outer part of the median depression, but absent from its centre.
|
||||
</paragraph>
|
||||
<paragraph id="681424464CC1A8D600A7D02D665BE3B2">
|
||||
Legs: mid brown, basal segment of mid tarsi clearly shorter than the longer tibial spur (Fig.
|
||||
<figureCitation id="EEB7486EAA8987D2FAB142E627360B36" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 c</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="2EF6667ED42E1F3E33A712A4130F4406">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="73DCFCE319D13E7DDB21351109E9A9D1" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 q, q</figureCitation>
|
||||
’ r, v, w) small, length ca
|
||||
<quantity id="F02AF612044E7825CB7D5ADC32B7011C" metricMagnitude="-4" metricUnit="m" metricValue="7.0" unit="mm" value="0.7">0.7 mm</quantity>
|
||||
with the endophallus bearing a dense field of bristles dorsally and the apices of the parameres membranous, expanded and obliquely truncate.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERFHK" type="remarks">
|
||||
<paragraph id="D3FA97BF2701EEC0779AA7331970741F">
|
||||
<heading id="A6B94EAE7C94E088A38DE6538430FF79" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="03C09E4B7C8D944BA425B3BC205DB511">
|
||||
<taxonomicName id="CF35248205D3DB8002D3E9FFF6B57327" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="0E6F6F772F3E02271E41D170AED9336D" italics="true">Aphodius rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
is distributed in mainly lowland coastal areas of southeast Europe and Asiatic
|
||||
<collectingCountry id="6DC5A4352BA0F19B8FFB655836CA12F2" name="Turkey">Turkey</collectingCountry>
|
||||
(
|
||||
<collectingCountry id="3BB472E736CAC30BD105E5601D984C2A" name="Turkey">Anatolia</collectingCountry>
|
||||
) from Corfu and southern
|
||||
<collectingCountry id="608B78A2C8B22173F913E27A342EC13E" name="Albania">Albania</collectingCountry>
|
||||
(Butrint) in the west, via Evia and
|
||||
<collectingCountry id="BB6CC57E60D8C27CEE33817852EBE3E8" name="Macedonia">Macedonia</collectingCountry>
|
||||
(
|
||||
<collectingCountry id="E2DF3E32885F02D147FE7A6EBB92D83D" name="Greece">Greece</collectingCountry>
|
||||
), Besika Bay (
|
||||
<collectingCountry id="6BF0D48FD18E04EB035BC56D94920AE5" name="Turkey">Turkey</collectingCountry>
|
||||
),
|
||||
<collectingCountry id="289D646D84624DA3A7D0627C4C30A2FD" name="Cyprus">Cyprus</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="3DEF8FDC2E462850318F841445EAF758" country="Cyprus" name="Lemesos">Limassol</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="B11C37C67F851CE3A8FF36E1B0C886F8" country="Cyprus" name="Larnaka">Larnaca</collectingRegion>
|
||||
districts) to
|
||||
<collectingCountry id="F14A26BFA37DBFC87F9412F71F5C488C" name="Turkey">Hatay</collectingCountry>
|
||||
(Iskenderun) (
|
||||
<collectingCountry id="ACA4BB2C5E16C2F49E4F41475E03442F" name="Turkey">Turkey</collectingCountry>
|
||||
: the
|
||||
<typeStatus id="B5744E3BE6F7CD8E2579C24EB1A92A39">type</typeStatus>
|
||||
locality of
|
||||
<taxonomicName id="DCCB3889BF78BFBDE230DE43EFC39A07" authorityName="Petrovitz" authorityYear="1962" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ressli">
|
||||
<emphasis id="75D634EB0074FDE723E2B22717843585" italics="true">L. ressli</emphasis>
|
||||
</taxonomicName>
|
||||
) in the east (Fig.
|
||||
<figureCitation id="57F875C89578A4BC8F0BD1E73EE802FF" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b, c</figureCitation>
|
||||
) On
|
||||
<collectingCountry id="6B71F0AD58DA1928545FC31D397ECEFD" name="Cyprus">Cyprus</collectingCountry>
|
||||
it appears to be mainly winter-active. RBA collected it at the edge of a muddy pool in the Zakaki marshes (
|
||||
<collectingRegion id="87457BFFDA0F588E8D18E390467763E5" country="Cyprus" name="Lemesos">Limassol</collectingRegion>
|
||||
) and C. Makris (
|
||||
<collectingRegion id="534FD5D9CC0E19808CC0488346A4AF5F" country="Cyprus" name="Lemesos">Limassol</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="9BE0933B1017A1E27B24A26DFEB72C2B" name="Cyprus">Cyprus</collectingCountry>
|
||||
) collected it from under a discarded piece of carpet lying on the mud in the same area (pers. comm. to RBA,
|
||||
<date id="49850FCF369D6010898A56601863557E" value="1985-04">April 1985</date>
|
||||
). This whole area is slightly saline.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
449
data/FE/60/5F/FE605F66CB5B5078A68A185AA9D27B99.xml
Normal file
449
data/FE/60/5F/FE605F66CB5B5078A68A185AA9D27B99.xml
Normal file
|
|
@ -0,0 +1,449 @@
|
|||
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<mods:title id="598B60FA099B92D80771380E6DB1D99A">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:affiliation id="743A2CEEDAD295C18A5E0347DD2E2A88">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="8A892366F625A37CB64FED7A6186E1D9">Maté, Jason F.</mods:namePart>
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<treatment id="FE605F66CB5B5078A68A185AA9D27B99" ID-arpha="FE605F66-CB5B-5078-A68A-185AA9D27B99" LSID="urn:lsid:plazi:treatment:FE605F66CB5B5078A68A185AA9D27B99" httpUri="http://treatment.plazi.org/id/FE605F66CB5B5078A68A185AA9D27B99">
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|
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<paragraph id="4E2701444DCF5FD5C5810742887C185D">
|
||||
<taxonomicName id="E37B13F6D9A358005E62BE3B963428E6" ID-CoL="FHNB" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kraatzi" subGenus="Liothorax">
|
||||
<emphasis id="5FA3ADD07326FFD0C8CAFE386DA6A040" italics="true">Aphodius</emphasis>
|
||||
(
|
||||
<emphasis id="9903061F9C7BFF1B67A29FD74C6F6882" italics="true">Liothorax</emphasis>
|
||||
)
|
||||
<emphasis id="C07DF7E907425AF42E101855079CC778" italics="true">kraatzi</emphasis>
|
||||
Harold, 1868
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="308909421ABF3D8ADCBEC15E6AF6E26B" type="description">
|
||||
<paragraph id="DF146E504DA309D5221630ECECADD89E">
|
||||
<figureCitation id="FB5DB74A271A88D550F46489EB65E584" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">Figs 3 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="1151C5159A10F8BAD444DB79AF5598A9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 o, p</figureCitation>
|
||||
,
|
||||
<figureCitation id="0F917F6AF9E4B3607EA4DF7B4FF52F8C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 a, b</figureCitation>
|
||||
,
|
||||
<figureCitation id="24DCDE90EFF0510F43BD1466843AF9F5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 b</figureCitation>
|
||||
,
|
||||
<figureCitation id="A713B1C2BEEEC8DB5B794CCBDB7446AC" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 j</figureCitation>
|
||||
,
|
||||
<figureCitation id="EDAB0735FF6E2DCBB034B9C4FF22BAD5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 f, g</figureCitation>
|
||||
,
|
||||
<figureCitation id="4AB7724E27C123CA7183509D8C11310A" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">
|
||||
<quantity id="BDFF8E6FF4FCDEBDDAA5770563199876" metricMagnitude="-2" metricUnit="kg" metricValue="2.0" unit="g" value="20.0">20 g</quantity>
|
||||
– i
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="F952FF442D45AF9A22624EC3075DF53D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 y, z</figureCitation>
|
||||
,
|
||||
<figureCitation id="46ED5DEBB991ABF3081312CC7576559D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure25" httpUri="https://binary.pensoft.net/fig/1097496">25 h</figureCitation>
|
||||
,
|
||||
<figureCitation id="8BC86E19ED566A829B8FA4F70DCFFC50" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 o – s</figureCitation>
|
||||
,
|
||||
<figureCitation id="D17A55A12D6E9ACB72FF8D38F0296682" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Mitotic chromosomes, arranged as karyotypes a, b A. plagiatus plagiatus, ♂, GB, Dorset, Studland a plain (Giemsa stained) b the same nucleus C-banded c A. plagiatus sinoplagiatus, ♂, CH, Qinghai, Gangca, plain d, e A. rutilipennis, ♂, CY, Zakaki Marshes d plain e C-banded f – h A. kraatzi ♀, SK f, g nucleus without B-chromosomes f plain g C-banded h, i nucleus with 1 B-chromosome h plain i C-banded. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure30" httpUri="https://binary.pensoft.net/fig/1097512">30 f – h</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="7E1ED65566B4750FA4B11602EEA9703C" type="reference_group">
|
||||
<paragraph id="CBC9725C9A48D379DDF28F79D60BE657">
|
||||
<treatmentCitationGroup id="DCEE18BAB343598B153F7BEE28C74E34">
|
||||
<treatmentCitation id="86BC6FC6DD35A7CAD91D5AAFC72D5686">
|
||||
<taxonomicName id="80E2637818641D302F3FC3EFE35E9EE4" ID-CoL="FHNB" authority="Harold, 1868: 84" authorityName="Harold" authorityPageNumber="84" authorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kraatzi">
|
||||
<emphasis id="CF326A057DA8AF8CDCAD8AA7BE8FD0A7" italics="true">Aphodius kraatzi</emphasis>
|
||||
Harold, 1868: 84
|
||||
</taxonomicName>
|
||||
.
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E4RHK" type="type material">
|
||||
<paragraph id="98FDB2775B4E5E153B9B5DBF8435AD64">
|
||||
<heading id="9D0AB71AA5D9AAECC459B5C21CB97395" reason="title">
|
||||
<typeStatus id="F81A7F8627F16254366096CF188AD64F">Type</typeStatus>
|
||||
material examined.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7CD7783F1A54BC37C3CDB4C675281232">
|
||||
(
|
||||
<collectionCode id="FB0ED07699BFD152FFB3A7B7C660487E" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" type="">MNHP</collectionCode>
|
||||
), seen but not studied in detail by DK.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ECSHK" type="material">
|
||||
<paragraph id="8AF677B7E0DA74F6C4C6F7C0E578FD33">
|
||||
<heading id="1FB66BBBE172924FD97A040879B84E83" reason="title">Additional material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B0B7489E1212ADE3D462B3CA0F43394A">
|
||||
SK: S.
|
||||
<collectingCountry id="287D2E3C30B6170D613ED1211BACF6DC" name="Slovakia">Slovakia</collectingCountry>
|
||||
, E. of Chlaba, sandy place S of the railway.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELSHK" specific-use="{"type":"Point","coordinates":[18.844000,47.842000]}">
|
||||
<geoCoordinate id="B3BF83E81BA9E3C6345F3F77F3B62AF8" degrees="47.842" direction="north" orientation="latitude" precision="55" value="47.842">47.842 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="7ADAF94DB43D2FA651884AB46A8091D5" degrees="18.844" direction="east" orientation="longitude" precision="55" value="18.844">18.844 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<quantity id="DDFE159F3900E536855099D4D8F07ED3" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="107.0">107 m</quantity>
|
||||
a. s. l ..
|
||||
<date id="8D63F5286963E3E73A51DB25711033B7" value="2014-06-11">11. vi. 2014</date>
|
||||
. leg. D. Král.
|
||||
<specimenCount id="D1FDD0E3523AEFF4702142E0D1762983" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="E69994D686ACD885A1C6746B88E38C05" count="5" type="female">5 ♀♀</specimenCount>
|
||||
(
|
||||
<named-content id="88A3B3305C4ABFA63143AAD2AE1D67CF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="81E172C975A1E0479E89A62E0551F8BA" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). Martovce.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXSHK" specific-use="{"type":"Point","coordinates":[18.127000,47.858000]}">
|
||||
<geoCoordinate id="E76CF1D83D6D9CDF41696BEE63BAA77C" degrees="47.858" direction="north" orientation="latitude" precision="55" value="47.858">47.858 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="A5BE5CDDBD6814AC45D858E22C7BFAA0" degrees="18.127" direction="east" orientation="longitude" precision="55" value="18.127">18.127 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="0710ECC837444B08E5F0443C61005ECF" count="2" type="female">2 ♀♀</specimenCount>
|
||||
(
|
||||
<named-content id="ADA454298038593FED4C90D647B686FC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="5C7A60ED89FD114F08C592C5F1F74DC4" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). Velké Kosíhy.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDTHK" specific-use="{"type":"Point","coordinates":[18.127000,47.734000]}">
|
||||
<geoCoordinate id="A689D3EF934FD6FD84CB95C56A60AD36" degrees="47.734" direction="north" orientation="latitude" precision="55" value="47.734">47.734 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="3C52338A820818AE07EF5B3118F1FBA4" degrees="18.127" direction="east" orientation="longitude" precision="55" value="18.127">18.127 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="A9CF97730924DD6EE6E7C06FB6653C99" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="0E96AAE647AD1C9D40B99263786A3045" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="DE04A1E818168DC07F08A5D803F01EEA" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). Virt.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPTHK" specific-use="{"type":"Point","coordinates":[18.324000,47.742000]}">
|
||||
<geoCoordinate id="C2B2B2A1044D4637C1E77310DFA1A6F1" degrees="47.742" direction="north" orientation="latitude" precision="55" value="47.742">47.742 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="CBE839C3EE15C8D11BCDBEF86FCAF890" degrees="18.324" direction="east" orientation="longitude" precision="55" value="18.324">18.324 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="CFF53D2B918E9A7859654D93A7B7B6F3" value="2012-07-31" valueMax="2012-08-02" valueMin="2012-07-31">31. vii – 2. viii. 2012</date>
|
||||
. Martinů Ivo leg. 2, unsexed (
|
||||
<named-content id="1951BDF67E65A52B5AD8C626FB9F6354" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="939E5EB973F063AD6651E6F1B114C920" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="BA55095629866530C868B72F349F2CE2">
|
||||
GR: Lesbos, Petra.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4THK" specific-use="{"type":"Point","coordinates":[26.181000,39.399000]}">
|
||||
<geoCoordinate id="D0F1F1215691675572183B61BD1BB255" degrees="39.399" direction="north" orientation="latitude" precision="55" value="39.399">39.399 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D3D90163358E279E178ED29D1472570F" degrees="26.181" direction="east" orientation="longitude" precision="55" value="26.181">26.181 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. 2, unsexed (
|
||||
<named-content id="923C542B9BA1BC0628416F7743566B0D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||||
<collectionCode id="82FD612440749B972CF345C903AD056A" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="EDE48FCF5C4E7303E29956654B40A245">
|
||||
RU:
|
||||
<collectingRegion id="8FCBCC539582FCCE946EFC1294251529" country="Russia" name="Astrakhan'">Astrakhan obl.</collectingRegion>
|
||||
Dosang.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELUHK" specific-use="{"type":"Point","coordinates":[47.916000,46.904000]}">
|
||||
<geoCoordinate id="B5323964ED418E70CC6B10C10D8E9FCC" degrees="46.904" direction="north" orientation="latitude" precision="55" value="46.904">46.904 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="10AC8A5AA7325053946967FEA85CD50B" degrees="47.916" direction="east" orientation="longitude" precision="55" value="47.916">47.916 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Cattle dung.
|
||||
<tp_taxon-name id="8F9317EDC125F891CBB97EBFD7381ACF">
|
||||
<emphasis id="F394D55085B07315C11D029C8F0FA145" italics="true">
|
||||
<tp_taxon-name-part id="6B5FE2114916FD60D7E7D743E64977C9" reg="Aphodius" taxon-name-part-type="genus">A.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Frolov &
|
||||
<tp_taxon-name id="C0675F0275FE7B609B914527866ED244">
|
||||
<emphasis id="CC863F53EF06334D131A56817B3AA084" italics="true">
|
||||
<tp_taxon-name-part id="DDCAECA1EEED55023F553E3E3AD0293C" reg="Liothorax" taxon-name-part-type="genus">L.</tp_taxon-name-part>
|
||||
</emphasis>
|
||||
</tp_taxon-name>
|
||||
Akhmetova.
|
||||
<specimenCount id="C2AB77FCB0F47DA78BBDEDEB9EF275C4" count="2" type="female">2 ♀♀</specimenCount>
|
||||
, 4 unsexed,
|
||||
<date id="975C8081788E789AAADD267BE3548FB0" value="2007-05-23">23. v. 2007</date>
|
||||
,
|
||||
<specimenCount id="5E5CAE3A98FAA03307E1C04AA5650D20" count="1" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<date id="08F4967AC6A261C25CE1B46438E52E26" value="2004-05-01" valueMax="2004-05-03" valueMin="2004-05-01">1–3. v. 2004</date>
|
||||
(
|
||||
<collectionCode id="22C80DC7651D4881DFD231EFA7E4B0C4" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" type="">NHML</collectionCode>
|
||||
).
|
||||
<collectingRegion id="3865F29747F4E9EEB0C39F4431BFB4AF" country="Russia" name="Novosibirsk">Novosibirsk obl.</collectingRegion>
|
||||
Karasuk.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6UHK" specific-use="{"type":"Point","coordinates":[78.000000,53.500000]}">
|
||||
<geoCoordinate id="CD48B589EB8E0ADB7A7FEFC221CB23F2" degrees="53.5" direction="north" orientation="latitude" precision="5555" value="53.5">53.5 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="C8DE368595089762C2BCC60BD69F7B01" degrees="78" direction="east" orientation="longitude" precision="55555" value="78.0">78 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<date id="A61D4BCDA9FC5E74637AC6E7BD2737B5" value="1982-06" valueMax="1982-07-31" valueMin="1982-06-00">vi – vii. 1982</date>
|
||||
. R. B.
|
||||
<collectingRegion id="FE67FE2D95E00A39AD887553A9EA8955" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||||
.
|
||||
<specimenCount id="0E1D277F8622CBF8DC85B0A0A96C338D" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="2219780279844A0872D7ACF1BC2D7E69" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="4BB516409753FF30E5C645718CAF7554" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
). SE Siberia, S Chita reg.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELVHK" specific-use="{"type":"Point","coordinates":[115.663446,50.071166]}">
|
||||
<geoCoordinate id="8EC6EE5C7DDEE109F9A6666B2CFF6BA4" degrees="50.071166" direction="north" orientation="latitude" precision="1" value="50.071167">50.071166 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="961D5356F65706575337378480CB9EDA" degrees="115.663446" direction="east" orientation="longitude" precision="1" value="115.663445">115.663446 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. Zun Tore Lake.
|
||||
<date id="85D385A6A21F9929419AB6AA7313AB8C" value="2002-05-30">30. v. 2002</date>
|
||||
.
|
||||
<specimenCount id="E9C694E98A3B69A7CE5D77FE6175C9AD" count="1" type="male">1 ♂</specimenCount>
|
||||
(
|
||||
<abbrev id="ABBRID0EQVHK" xlink_title="Jason Maté Collection">
|
||||
<collectionCode id="6FAE812174EA10D58632BE2450B46760">JFMC</collectionCode>
|
||||
</abbrev>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="4EB6CAF947CCC6C68B2DB8CEB7611FEC">
|
||||
IN “ O. Ashur-ade ” (Ashooradeh Island).
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZVHK" specific-use="{"type":"Point","coordinates":[53.960000,36.910000]}">
|
||||
<geoCoordinate id="BC48117C9541A39F2F69E92189A3B77F" degrees="36.91" direction="north" orientation="latitude" precision="555" value="36.91">36.91 ° N</geoCoordinate>
|
||||
, 53.96 ° Е
|
||||
</named-content>
|
||||
. Solovkin
|
||||
<date id="FBAB4DE8484AEF4E07F0306B1308016F" value="1913-05-01">1. v. 1913</date>
|
||||
. 4, unsexed (
|
||||
<named-content id="D4733EA31A013CA6A8E71B418F37E785" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||||
<collectionCode id="7A68548F456559D7F5070DA6CB659E45" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||||
</named-content>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="556CCAE0BF5E75D27660123211F912BE">
|
||||
KZ: Dzhulek, Orenburg – Tashkent railway, Syr Darja (Taken as Zholek near Baigakum,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHWHK" specific-use="{"type":"Point","coordinates":[66.466000,44.314000]}">
|
||||
<geoCoordinate id="251294105C9FE1EDB78DA80ED439FE11" degrees="44.314" direction="north" orientation="latitude" precision="55" value="44.314">44.314 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="9CC8DEF6EDAFCF90DB8DAFFC4E95AAC6" degrees="66.466" direction="east" orientation="longitude" precision="55" value="66.466">66.466 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
),
|
||||
<specimenCount id="ADF532ECE48FAC439FFB62C629970069" count="3" type="male">3 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="2647C25440899FB4593415F035A13EE0" count="3" type="female">3 ♀♀</specimenCount>
|
||||
, 9, unsexed (
|
||||
<collectionCode id="EF79DA431F3C785F89622D24F8B8F733" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" type="">NHML</collectionCode>
|
||||
). Kozakhit, Almaty obl. Altyn Emel NP Ul; ken Kalkan.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPWHK" specific-use="{"type":"Point","coordinates":[78.749000,43.874000]}">
|
||||
<geoCoordinate id="ED1CD7377ACA9917C414F40FE6182CB7" degrees="43.874" direction="north" orientation="latitude" precision="55" value="43.874">43.874 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1600DC370CD04351E799200CA70ECE2C" degrees="78.749" direction="east" orientation="longitude" precision="55" value="78.749">78.749 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
. N bank of R. Ile, at light,
|
||||
<date id="8A1827ECB3D49305538C58F5DD3E16B5" value="2012-05-05">5. v. 2012</date>
|
||||
. D. J. Mann & J. Cooter.
|
||||
<specimenCount id="9A308E3B97A1A68065202D845ADA49E4" count="1" type="male">1 ♂</specimenCount>
|
||||
, 3 unsexed (
|
||||
<named-content id="5073199689F8F5648A1589944E7F5836" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oxford-university-museum-natural-history" xlink_title="Oxford University Museum">
|
||||
<collectionCode id="B5060F498B3064F27B4EFF12EA1E2E26" country="United Kingdom" httpUri="http://grbio.org/cool/g6yf-wft0" name="Oxford University Museum of Natural History" type="">OUM</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="F77792C69E471F15444885438D4B0A8F">
|
||||
UZ: Kyzylkum desert Baymachan distr.
|
||||
<date id="3C98F33043C097278F1EA0359388E22B" value="1995-05-11">11. v. 1995</date>
|
||||
. 2, undissected (
|
||||
<named-content id="176BF0EE6794E7D7CEB1E8233172F26C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||||
<collectionCode id="62306480BAFBBDCC937DBBFC9EE5AD39" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="62CF1D3E87C12ED8DAA6478089573B46">
|
||||
AF:
|
||||
<collectingRegion id="D622E930693E35CA8E6D7B9D9015B0A2" country="Afghanistan" name="Kabul">Kabul</collectingRegion>
|
||||
.
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDXHK" specific-use="{"type":"Point","coordinates":[69.213000,34.548000]}">
|
||||
<geoCoordinate id="4DC734C2546C2E680967FC2AFB6521AD" degrees="34.548" direction="north" orientation="latitude" precision="55" value="34.548">34.548 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D35DB5D2C99399407F6C8E0A22673AC7" degrees="69.213" direction="east" orientation="longitude" precision="55" value="69.213">69.213 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
.
|
||||
<specimenCount id="F267F863662666C83876DA70D37AC5C1" count="1" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="24C833E431C27DC6BB4088517AB7F9ED" count="1" type="female">1 ♀</specimenCount>
|
||||
(
|
||||
<named-content id="B848937323773ADC51F4A16486CEFCAD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||||
<collectionCode id="84A6304A59AB1515AE6BC9472B3055D4" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<specimenCount id="6EC1A2AA978061602BF9F7A241892DC7" count="1" type="female">1 ♀</specimenCount>
|
||||
, 1 unsexed (
|
||||
<named-content id="170FAF2EB95EF222B64709383FDCF65A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||||
<collectionCode id="CFD691641330FB51123F56369DD01395" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||||
</named-content>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EQXHK" type="Differential diagnosis">
|
||||
<paragraph id="CC9EFC0ACC95EC47AD1B3EA44905607E">
|
||||
<heading id="6101F0AC5DBEF4ADDD7F38D8BE17A3EA" reason="title">Differential diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="14563AA9282B3E49BBDE0C39E24AB7EF">
|
||||
A conspicuously narrow elongate black beetle, with the scutellum wide, generally wider than elytral interstice 2. Elytral interstices weakly raised, not flat as in most other species. Smaller, length
|
||||
<quantity id="97DF53D92867B252C147AD9990ED294E" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.8" metricValueMin="3.7" unit="mm" value="4.25" valueMax="4.8" valueMin="3.7">3.7–4.8 mm</quantity>
|
||||
. Basal line of pronotum completely effaced over middle 1 / 3 (Fig.
|
||||
<figureCitation id="DF9850CF4F4BADAC1354ED9916D040C8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 b</figureCitation>
|
||||
).
|
||||
<specimenCount id="B2EC25F384D3BBBB3D53AAC870B74D31" count="1" type="female">♀</specimenCount>
|
||||
with spermatheca smaller (Fig.
|
||||
<figureCitation id="771FFAF155E229D4A529DFF6DD3FF3AA" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 o – s</figureCitation>
|
||||
), length ca
|
||||
<quantity id="4208F7B54352FA1BDB31E08907419E8F" metricMagnitude="-4" metricUnit="m" metricValue="2.5" unit="mm" value="0.25">0.25 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E5XHK" type="redescription">
|
||||
<paragraph id="A74AAC52E153AF0F13C80B7EC94B65FC">
|
||||
<heading id="8791578C176E44B47FA742C2DC0F06A5" reason="title">Redescription.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8890F5AD58B33AF7334786CB8E37D72C">
|
||||
General appearance (Fig.
|
||||
<figureCitation id="E50317DFDB524B52CBAB6AC92D4CCDB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure3" httpUri="https://binary.pensoft.net/fig/1097461">3 a, b</figureCitation>
|
||||
). Black, elongate, parallel-sided, and conspicuously narrow. Length:
|
||||
<quantity id="E583AC71C77AD9F74DCFE77A7DCE12A5" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.8" metricValueMin="3.4" unit="mm" value="4.1" valueMax="4.8" valueMin="3.4">3.4–4.8 mm</quantity>
|
||||
, width
|
||||
<quantity id="9F2868A6C7955C2CC19F18075E82A0A0" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="0B1BA6F78C2E8D123D198401717E5690">
|
||||
Head black with anterior and lateral margins of the clypeus narrowly brown. Anterior margin of clypeus emarginated, the sides of this emargination bluntly rounded. Central part of clypeus bulging upwards, frontoclypeal suture occasionally slightly raised medially but without any trace of a real tubercle, straight or slightly curved backwards over middle 1 / 2, angled forward to run to the junction of the clypeus and genae laterally. Punctation double, varying from close and strong to rather sparse, generally weaker and sparser in the central area of the frons and becoming rugose anteriorly and laterally (head and pronotum: Fig.
|
||||
<figureCitation id="07F29B2C30B2B209DD8817C2529A005D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure5" httpUri="https://binary.pensoft.net/fig/1097463">5 o, p</figureCitation>
|
||||
) Maxillary palpi dark brown, the apical segment blackish. Antennae brown, the clubs darker. Epipharynx (Fig.
|
||||
<figureCitation id="6376152DFC492F2B04309E4A6042FEC5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" figureDoi="10.3897/zookeys.1207.117225.figure8" httpUri="https://binary.pensoft.net/fig/1097467">8 a, b</figureCitation>
|
||||
) with the anterior margin of the clithra almost straight either side of the median tylus and the acropariae very well-developed. The prophobae are well-developed on each side of the mesoepitorma and there are three or four chaetopedia on each side. Pronotum black, hemicylindrical, highly arched transversely, only weakly so longitudinally. Entire lateral margins visible from above. Surface with double punctation of variable strength, generally weaker medially. Hind margin bordered only in lateral 1 / 4 s (Fig.
|
||||
<figureCitation id="867AF28D3D740EDDA7A4D200D80AA3C2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." figureDoi="10.3897/zookeys.1207.117225.figure14" httpUri="https://binary.pensoft.net/fig/1097474">14 b</figureCitation>
|
||||
). Scutellum rather wide, generally wider than elytral interstice 2, pentagonal, moderately elongate, sides parallel in basal 1 / 2, then angled to apical convergence. Metaventrite variable. Median diamond-shaped plate sometimes excavated and quite strongly punctured (Fig.
|
||||
<figureCitation id="A977C29A9B59E296A68A12D118F0C2A7" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">
|
||||
<quantity id="B8A9165438A0E02CECE5F953233E6D0E" metricMagnitude="-2" metricUnit="kg" metricValue="2.0" unit="g" value="20.0">20 g</quantity>
|
||||
</figureCitation>
|
||||
), or flatter and more finely punctured (Fig.
|
||||
<figureCitation id="FDEC87643DBF5B4BE66690EED1230832" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure20" httpUri="https://binary.pensoft.net/fig/1097486">20 h, i</figureCitation>
|
||||
). Legs brown, longer spur of mid tibiae clearly longer than the basal segment of the tarsus (Fig.
|
||||
<figureCitation id="9087A2AD568C66203E3B2665E3CB38F0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 f, g</figureCitation>
|
||||
). Elytra glossy black, interstices very weakly convex, finely reticulate and with sparse fine punctation, and 6–8 × width of the striae (Fig.
|
||||
<figureCitation id="C5D89C27AE5316EE1A6FC9E9AE63E36E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 j</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4A1617988CB7C4D5B78A29226BBB4450">
|
||||
Aedeagus (Fig.
|
||||
<figureCitation id="BAAF63FFC98CC5B8DAA2D5AD7F6C9240" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." figureDoi="10.3897/zookeys.1207.117225.figure22" httpUri="https://binary.pensoft.net/fig/1097490">22 y, z</figureCitation>
|
||||
) ca
|
||||
<quantity id="769FC470A7946970C788291455FA4E7C" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
|
||||
long, apical 1 / 3 of parameres slightly more expanded than in
|
||||
<taxonomicName id="27021A1BF1D5A8127521AA5FA4C008CF" authorityName="Baudi di Selve" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rutilipennis">
|
||||
<emphasis id="15469EE3C8E1E0BED4BAEAD6A7C553F0" italics="true">A. rutilipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="0ECA7A546F4F443B2A8C92F2D594AF8B" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chellala" status="sp. nov.">
|
||||
<emphasis id="03083459BCD3238056D47C2A6B940A51" italics="true">A. chellala</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4CF7E8EE2EACA9DD47BC5219B189B569" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, its outer margin more rounded. Endophallus with fine bristles.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EC1HK" type="remarks">
|
||||
<paragraph id="2663B71FBD34588A773730CB0AF949EA">
|
||||
<heading id="F75C72B933997229D0B2FB7A99DB3739" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="72EE77DF86CF91A5DA78F8F9E2BB8206">
|
||||
<taxonomicName id="EBA2BEFD7C335C22FDB746F6E1EB7CF0" authorityName="Harold" authorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kraatzi">
|
||||
<emphasis id="5C0BE16F025C2543D9E7C0CAD59C0B4C" italics="true">Aphodius kraatzi</emphasis>
|
||||
</taxonomicName>
|
||||
is widely distributed in the drier regions of the western and central Palaearctic, from southern
|
||||
<collectingCountry id="7F69243BD230C50E0E7A7B0CDD4B60A0" name="Slovakia">Slovakia</collectingCountry>
|
||||
to
|
||||
<collectingCountry id="097D454D10C2FDB2268CD04925CE6060" name="Kazakhstan">Kazakhstan</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="C0EFE2A7989F93731F41098714E38857" name="Uzbekistan">Uzbekistan</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="406223483E435888E24B9FDE61D9E321" name="Afghanistan">Afghanistan</collectingCountry>
|
||||
, and the Chita region of Transbaikal in
|
||||
<collectingCountry id="A0FEFAFD1A447DB1E89A21ED669F1E97" name="Russia">Russia</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="D8F031A9A2CC12B1DF65955E57C809A0" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 b, c</figureCitation>
|
||||
). Most specimens are taken at light but Andrey Frolov (
|
||||
<collectingRegion id="B739C17F6DBC9D46B470371767A893A4" country="Russia" name="Saint Petersburg">St Petersburg</collectingRegion>
|
||||
) mentions that it can sometimes be found at the roots of grass and other herbaceous plants, as in the area of the
|
||||
<collectingRegion id="46B2740450DC28A533C8176C9B21DE92" country="Russia" name="Astrakhan'">Astrakhan</collectingRegion>
|
||||
oblast’ (
|
||||
<collectingCountry id="F2E3362DD88B7E780F3ADBF11CAB0E3D" name="Russia">Russia</collectingCountry>
|
||||
) shown in Fig.
|
||||
<figureCitation id="FCF6911D658B37B79A43A6F47FAB541A" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." figureDoi="10.3897/zookeys.1207.117225.figure28" httpUri="https://binary.pensoft.net/fig/1097505">28 d</figureCitation>
|
||||
. He adds that material flying to light often arrives from the direction away from neighbouring water (pers. comm. to RBA,
|
||||
<date id="D10B2D8DAE704FE05617A89999D38B62" value="2015-04">April 2015</date>
|
||||
). In 1982 RBA took
|
||||
<specimenCount id="32A8F14B1BD00B7C60659EAF32651F17" count="1" type="generic">one specimen</specimenCount>
|
||||
in a water-filled ditch at Karasuk on the Kulundinskaya Steppe in Western Siberia.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue