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<document id="55E1B51990AD6B84C134A93CB71EDE66" ID-DOI="10.3897/phytokeys.248.120387" ID-publisher-id="120387" URI-arpha="9100FB1E-F666-5521-A003-A5A3572B9B53" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1729607794056" checkinUser="pensoft" docAuthor="Gentallan Jr., Renerio P., Sengun, Seda, Bartolome, Michael Cedric B., Quiñones, Kristine Joyce O., Coronado, Nadine B., Borromeo, Teresita H. &amp; Timog, Emmanuel Bonifacio S." docDate="2024" docId="C2018846DDF358CAAC4D14B657D4A714" docLanguage="en" docName="PhytoKeys 248: 1-40" docOrigin="PhytoKeys 248" docSource="https://phytokeys.pensoft.net/article/120387/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Vitex arvensis Gentallan, Sengun &amp; M. B. Bartolome 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="40" masterDocId="9100FB1EF6665521A003A5A3572B9B53" masterDocTitle="The Vitex trifolia complex (Lamiaceae) in the Philippines" masterLastPageNumber="40" masterPageNumber="1" pageNumber="1" updateTime="1729613092210" updateUser="admin">
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<mods:title id="11DA531CDE11A8A26DE846E0393D63DB">The Vitex trifolia complex (Lamiaceae) in the Philippines</mods:title>
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<mods:namePart id="B4D3046B47AF889B3EF722AE42C09F07">Gentallan Jr., Renerio P.</mods:namePart>
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<mods:affiliation id="9D5DED3BD1F40DD3F5D04246EEB70C67">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="BAE8401E7DD5F75288B4BBE629FA0563">Sengun, Seda</mods:namePart>
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<mods:affiliation id="FAD02408A13A04BB4552C96962B1B39E">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="6933D92EB1FFAAE7D8E82CF77573DEFC">Bartolome, Michael Cedric B.</mods:namePart>
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<mods:affiliation id="A73B8F7C16700BD4166E99B2C53B11C9">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="82A37F7BF0DB50746C735FF73FA84BCE">Quiñones, Kristine Joyce O.</mods:namePart>
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<mods:affiliation id="95840EEA53E3BCCCC49BDB431030FEA7">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="17CC5735F12EB94639D34C68FB642194">Coronado, Nadine B.</mods:namePart>
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<mods:affiliation id="50111225F508A0F1F85F78A43B47DAA5">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="956529FDC9CF83FF7F18BE53EDEEBD39">Borromeo, Teresita H.</mods:namePart>
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<mods:affiliation id="5A2E212CFF422B314AE7A81651C662EF">Institute of Crop Science, College of Agriculture and Food Science, University of the Philippines Los Baños, Los Baños, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="4480207FF90E455299D439202CD6391E">Timog, Emmanuel Bonifacio S.</mods:namePart>
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<mods:date id="E583964D1EB5E8E819ECDCE955A018F9">2024</mods:date>
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<treatment id="C2018846DDF358CAAC4D14B657D4A714" ID-arpha="C2018846-DDF3-58CA-AC4D-14B657D4A714" ID-ipni="urn:lsid:ipni.org:names:77350665-1" LSID="urn:lsid:plazi:treatment:C2018846DDF358CAAC4D14B657D4A714" LSID-IPNI="urn:lsid:ipni.org:names:77350665-1" httpUri="http://treatment.plazi.org/id/C2018846DDF358CAAC4D14B657D4A714">
<subSubSection id="2958C238D87458E4FDB9699DD6D8B16E" type="nomenclature">
<paragraph id="1384AE22A3ED6E2F9CF9535C5DB32F6A">
<heading id="32146AFA203EE4F1FDF63E57F32FCF04" reason="label">1.</heading>
<taxonomicName id="BC32BBFC673FFF005AF468C5089ADE91" authority="Gentallan, Sengun &amp; M. B. Bartolome" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis" status="sp. nov.">
<emphasis id="27C16FFCDA308C194C87CD597D66577D" italics="true">Vitex arvensis</emphasis>
Gentallan, Sengun &amp; M. B. Bartolome
</taxonomicName>
<taxonomicNameLabel id="F429B8C8C3ACB3588D81ED14DB5CC8CF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D7BF6BE79B432F24BF21DAB4C737B609" type="description">
<paragraph id="96C1AE84724BF522807FCC69DD10409E">
<figureCitation id="8696966E514C2E91652B5B938C8CB460" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Vitex arvensis A habit B part of inflorescence C side view of flower D calyx open, adaxial E Corolla open F style fruit in G calyx H fruit in cross-section I calyx. Illustration by Daryl Ceribo. Scale bars: 2.5 cm (A, B); 1 cm (C), 1 mm (D, E, F, G, H, I)." figureDoi="10.3897/phytokeys.248.120387.figure7" httpUri="https://binary.pensoft.net/fig/1158185">Figs 7</figureCitation>
,
<figureCitation id="2ABEA1C16487CAD35F6A3C8180431DED" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Morphological characteristics of V. arvensis A branch with inflorescence B sixth fully unfolded leaf C abaxial leaf lamina surface D F flower G inflorescence unit H I mature fruit with and without calyx." figureDoi="10.3897/phytokeys.248.120387.figure8" httpUri="https://binary.pensoft.net/fig/1158186">8</figureCitation>
,
<figureCitation id="3C35285307C9EE10DCBADEDA22319665" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Leaf undersurface pubescence of V. arvensis (40 × magnification). Photos: L. E. Endonela." figureDoi="10.3897/phytokeys.248.120387.figure9" httpUri="https://binary.pensoft.net/fig/1158187">9</figureCitation>
,
<figureCitation id="CDA4B798BE1A8F4F0600B961C6EF4C3B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Natural distribution of Vitex arvensis in the Philippines." figureDoi="10.3897/phytokeys.248.120387.figure10" httpUri="https://binary.pensoft.net/fig/1158188">10</figureCitation>
,
<figureCitation id="F0E9469CB94673419E3DBD041A8A6FDF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype specimen of Vitex arvensis." figureDoi="10.3897/phytokeys.248.120387.figure11" httpUri="https://binary.pensoft.net/fig/1158189">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5CC981F93F47EF9590BEDCF0A4B4E6A2" type="reference_group">
<paragraph id="28CA6002F8B49EB849D010576BF70715">
<treatmentCitationGroup id="D32A3B7CB364932B49729DCE8681FEDE">
<treatmentCitation id="851B663A17F0CE6C6F7223F1A4E9A2CF">
<taxonomicName id="C0890959889FB666C0FDFA5700AF7781" ID-CoL="5BL7L" authority="Blanco" authorityName="Blanco" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="leucoxylon">
<emphasis id="2B6EECE9C1504F2AC2A31021B3567AE8" italics="true">Vitex leucoxylon</emphasis>
Blanco
</taxonomicName>
, Fl. Filip. 516. 1837. nom. illeg.
</treatmentCitation>
, not
<taxonomicName id="5297DB7E09B88B13E9122BEFC3282F6C" ID-CoL="5BL7L" authority="L. f." authorityName="L. f." class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="leucoxylon">
<emphasis id="4B41743027946F79015E5B0B2EF84B98" italics="true">Vitex leucoxylon</emphasis>
L. f.
</taxonomicName>
Type:
<collectingCountry id="C5FF4887DA8CDC7614E1CC95DECF957E" name="Philippines">Philippines</collectingCountry>
,
<collectingRegion id="3D552B3741C4AC90F66831129B731CF1" country="Philippines" name="Rizal">Rizal</collectingRegion>
,
<collectingRegion id="A3BBC833163E81B0B89BF90366ED786E" country="Philippines" name="Antipolo">Antipolo</collectingRegion>
,
<date id="903DB0F6A81CF6E353C67D4F0DC45CD7" value="1914-01-13">13 Jan. 1914</date>
, Merrill Sp. Blanc. 440 (
<typeStatus id="00AEA9B0D3B1C72B425F6F051494935D" type="neotype">neotype</typeStatus>
: L [L. 2768327, designated by
<bibRefCitation id="915681CE3A8244847C393AD0C6AA2FBB" DOI="10.3767/blumea.2024.69.02.01" author="Sengun S &amp; Ngrouille M &amp; Paton A &amp; De Kok RP" etAl="et al." firstAuthor="Sengun" issue="2" journalOrPublisher="Blumea" pagination="93-121" refId="B39" refString="Sengun S, Ngrouille M, Paton A, De Kok RP (2024) Taxonomic Revision of the Vitex trifolia Complex (Lamiaceae). Blumea 69 (2): 93121. https://doi.org/10.3767/blumea.2024.69.02.01" title="Taxonomic Revision of the Vitex trifolia Complex (Lamiaceae)." volume="69" year="2024">Sengun et al. (2024)</bibRefCitation>
]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AA0028834D1A1DA1A27DF22EC0ABEC21">
<treatmentCitationGroup id="818554BBDDAD5EFF65B04F612A668C51">
<treatmentCitation id="89334378C102DCF309792C48F14C396F" author="Moldenke" journal="Phytologia" page="308" score="4.864797368871433" volume="38" year="1978">
<taxonomicName id="3C53C35BD92A93D7EFCE6A4680FF64E9" ID-CoL="7FTD7" authority="Moldenke" authorityName="Moldenke" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="variety" species="negundo" variety="philippinensis">
<emphasis id="A19E6D65B6ACDC302FBA8C54D3E9F9A3" italics="true">Vitex negundo var. philippinensis</emphasis>
Moldenke
</taxonomicName>
, Phytologia 38: 308. 1978
</treatmentCitation>
. Type:
<collectingCountry id="3DFBDB5CACB3F9FCE8364674571A67ED" name="Philippines">Philippines</collectingCountry>
,
<collectingRegion id="65E0CBD3884887C6EBA61B06522CCEDB" country="Philippines" name="Laguna">Laguna</collectingRegion>
, Los Baños, Apr., 1906, A. D. E. Elmer 8125 (
<typeStatus id="A474211FBC03499D677368C90DBBEF67" type="holotype">holotype</typeStatus>
: PNH, destroyed;
<typeStatus id="6C86084E3A2FC3CD43497B8D11DE5E11" type="isotype">isotypes</typeStatus>
: K, NY [NY 00138511]). syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPDI" type="diagnosis">
<paragraph id="571FA13FA365A15BAD05329B67587194">
<heading id="80C26258587F465B3C5242B01F643340" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1BD1ADFE17422810218FEB6DC705BD25">
This new species differs from
<taxonomicName id="FAE5E14E78024F7C1BBF21894132A299" authority="Moldenke" authorityName="Moldenke" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="CE96BB006E596E615F1834227B4F240D" italics="true">Vitex elmeri</emphasis>
Moldenke
</taxonomicName>
by having terminal leaflet which is narrowly elliptic to lanceolate, 4.316.0 by 1.0
<quantity id="46B98C43BE029811D1A4F05106C52A81" metricMagnitude="-2" metricUnit="m" metricValue="3.7" unit="cm" value="3.7">3.7 cm</quantity>
, 2.45.7 times as long as wide, and with abaxial surface moderately pubescent (vs
<taxonomicName id="BF31E26ABAC7A2D2F945CDFC95F022AF" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="5F7E757834C569EDE7B936B7955748F8" italics="true">V. elmeri</emphasis>
</taxonomicName>
terminal leaflet narrowly elliptic, 4.012.0 by
<quantity id="B4D00EF5619E117B3ED9639C3B61D289" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="3.5" metricValueMin="0.7" unit="cm" value="2.1" valueMax="3.5" valueMin="0.7">0.73.5 cm</quantity>
, 2.58.4 times as long as wide); inflorescence with caducous flowers, leaving a clear set of scars on the axis (vs
<taxonomicName id="A2742D631EFFC2071519CBD116B13E1B" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="F042840B4FF11C93F76B16265EE9CE43" italics="true">V. elmeri</emphasis>
</taxonomicName>
inflorescence with persistent flowers); fruit 3.04.9 by 3.0
<quantity id="9E1ABDC90BAA501EAC69EDFFD3213C61" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
(vs
<taxonomicName id="402C6A9C6E224E174EBFDD6D00638D97" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="B1373F43BE22AEE347C528D08AD35B9F" italics="true">V. elmeri</emphasis>
</taxonomicName>
bears 1.83.1 by
<quantity id="E65A2DCB80AF28A6C1E5F4CA07C8F84B" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" unit="mm" value="0.6" valueMax="0.8" valueMin="0.4">0.40.8 mm</quantity>
fruits fully covered by the calyx at maturity).
</paragraph>
<caption id="86967E4252BB229F42EE9AA987D89DFD" ID-DOI="10.3897/phytokeys.248.120387.figure7" ID-arpha="7A0E8029-1B0B-54CC-A4BD-40D50EE893D5" httpUri="https://binary.pensoft.net/fig/1158185" startId="F7">
<paragraph id="A2A41A2B2B8F82091B9A9D59072EE9A5">
<label id="06F58785082358B7C852B8BA96A0830B">Figure 7.</label>
</paragraph>
<paragraph id="269A9C043DB9ABC3EBC4643D4DE92B6E">
<taxonomicName id="627590CD8175D834CE6D43C3ECE1656C" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="D10F0DA5E324EE6792B5A87753D82067" italics="true">Vitex arvensis</emphasis>
</taxonomicName>
<emphasis id="EC3AD8E93F2B9A6F02BB20D2DA17B193" bold="true">A</emphasis>
habit
<emphasis id="4AE220B047297CD6EA9631D0B96A0E43" bold="true">B</emphasis>
part of inflorescence
<emphasis id="40B35023184036E4E4A460452EBE5128" bold="true">C</emphasis>
side view of flower
<emphasis id="012CF9856216AA31CE8CDE04CE4331BE" bold="true">D</emphasis>
calyx open, adaxial
<emphasis id="64520D0A9EA49A119AB5207FC4F38E5E" bold="true">E</emphasis>
Corolla open
<emphasis id="5500A52E3B8FA5442D18ADAF6A250144" bold="true">F</emphasis>
style fruit in
<emphasis id="E7ADD18DC9D138AD8932E6D5B30C6391" bold="true">G</emphasis>
calyx
<emphasis id="0E36BE80BA91C15A2A8F8B371D6362A3" bold="true">H</emphasis>
fruit in cross-section
<emphasis id="FA083CF209381689182F17BCF16C850F" bold="true">I</emphasis>
calyx. Illustration by Daryl Ceribo. Scale bars: 2.5 cm (
<emphasis id="143864550C477F9C63D525F8F60A44EC" bold="true">A, B</emphasis>
); 1 cm (
<emphasis id="D4678E571FAAA2419335D7BB5E712012" bold="true">C</emphasis>
), 1 mm (
<emphasis id="C3254D9C9B6B600C3DA6B64C447E93F8" bold="true">D, E, F, G, H, I</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGSDI" type="material">
<paragraph id="E71000762BEED02E6BB98D9766BCDD7B">
<heading id="A16BB7DCA6D33F30D36817EF1BFBEE38" reason="title">Type.</heading>
</paragraph>
<paragraph id="B5D8E6C7E7051C59F380CBC5753C0363">
<materialsCitation id="9E0EA41F4E6E790B2D8026DEB6942510" collectingDate="2019-02-08" collectionCode="ICROPS" collectorName="R. P. Gentallan &amp; M. B. Bartolome" country="Philippines" latitude="14.159721" location="in the field genebank of the Institute of Crop Science, University of the Philippines Los Banos" longLatPrecision="21" longitude="121.24528" municipality="Los Banos" specimenCount="1" stateProvince="Laguna" typeStatus="holotype">
<collectingCountry id="8A42F8356DD3DEC2327769351AFCC8FA" name="Philippines">Philippines</collectingCountry>
.
<collectingRegion id="655D84BBEDCAD6C6033D378C51E585DF" country="Philippines" name="Laguna">Laguna</collectingRegion>
:
<collectingMunicipality id="63F5358A671D00BBC1855481A1B29C6C">Los Baños</collectingMunicipality>
,
<location id="F7576FBA1E2DE861C63407ED131EEF68" LSID="urn:lsid:plazi:treatment:C2018846DDF358CAAC4D14B657D4A714:F7576FBA1E2DE861C63407ED131EEF68" country="Philippines" latitude="14.159721" longLatPrecision="21" longitude="121.24528" municipality="Los Banos" name="in the field genebank of the Institute of Crop Science, University of the Philippines Los Banos" stateProvince="Laguna">in the field genebank of the Institute of Crop Science, University of the Philippines Los Baños</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPSDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[121.245278,14.159722]}">
<geoCoordinate id="BFFAC30708DDA9840C5A5322D134D121" degrees="14" direction="north" minutes="09" orientation="latitude" precision="15" seconds="35" value="14.159721">14°0935″N</geoCoordinate>
,
<geoCoordinate id="B74C26A4377C4115408E96960243E140" degrees="121" direction="east" minutes="14" orientation="longitude" precision="15" seconds="43" value="121.24528">121°1443″E</geoCoordinate>
</named-content>
,
<date id="D2A5F09E6FF9F91B562A7A83AD4BB8F1" value="2019-02-08">
<collectingDate id="2265D9B14778A1EEE774DBE3755FDBC9" value="2019-02-08">8 Feb. 2019</collectingDate>
</date>
,
<emphasis id="F002BCE7BA59B0E9FF82A209F569A4A6" italics="true">
<collectorName id="D2D06247D9330E1D2FD1AD4D731D63FF">R. P. Gentallan</collectorName>
&amp;
<collectorName id="75E199FC2F2B8799493DECDB36759AF2">M. B. Bartolome</collectorName>
743
</emphasis>
(
<typeStatus id="CEA83F9B17CF6EC12F486AAB70A1C5C8" type="holotype">holotype</typeStatus>
:
<collectionCode id="D5F031D06F22081BFE621B3D1E322890">ICROPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="45FD001F7AE70A696FDC01035257E4C2" ID-DOI="10.3897/phytokeys.248.120387.figure8" ID-arpha="A573D525-2245-52BC-BC05-9C17C6B2EB3C" httpUri="https://binary.pensoft.net/fig/1158186" startId="F8">
<paragraph id="044E0E35F1289AF007A89ECE4C16CC04">
<label id="8501D3DAC10AC67029A9A110F9237926">Figure 8.</label>
</paragraph>
<paragraph id="65CA934C44588D45F387F5E9780FA853">
Morphological characteristics of
<taxonomicName id="5EA4E4C76B0CEE9381B4A9D44D4A6086" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="FF1209B30FC06BE18BCBEEE2BAC9EA5C" italics="true">V. arvensis</emphasis>
</taxonomicName>
<emphasis id="0E94D561172316A33077D195C83E94EB" bold="true">A</emphasis>
branch with inflorescence
<emphasis id="610F36508D5C469386164A4EA3DC06FF" bold="true">B</emphasis>
sixth fully unfolded leaf
<emphasis id="C6F9B7989F42A995789E3A135A8D04A8" bold="true">C</emphasis>
abaxial leaf lamina surface
<emphasis id="F028CF8A9B3B9D1BA67C171CA83F7257" bold="true">D F</emphasis>
flower
<emphasis id="0085A007486BC9A0F7BE6BDDAFBCC2E9" bold="true">G</emphasis>
inflorescence unit
<emphasis id="DADDB6D5F122A82C1AB36DCB3A0B999D" bold="true">H I</emphasis>
mature fruit with and without calyx.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E1TDI" type="description">
<paragraph id="FDE3D4B46E3A3B4417535DD42F07DB4D">
<heading id="7A735D1FE4CAEBB420B7D5FABA23EA83" reason="title">Description.</heading>
</paragraph>
<paragraph id="A0D7213127CB4139A6AC270084A04BF3">
Shrub to small tree,
<quantity id="2B3C1DA5B5601C19E6883440057471BD" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">24 m</quantity>
high. Leaves (1 ) 35 - foliolate, reflexed to drooping; terminal leaflet elliptic to lanceolate, 4.316.0 by 1.0
<quantity id="D413ED771DBFA8220B2FA6F85CCBDFEA" metricMagnitude="-2" metricUnit="m" metricValue="3.7" unit="cm" value="3.7">3.7 cm</quantity>
, 3.05.0 times as long as wide, terminal petiolules
<quantity id="F9D37E36C006A5D2D58F3B7F9912CC24" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="2.3" metricValueMin="0.0" unit="cm" value="1.15" valueMax="2.3" valueMin="0.0">02.3 cm</quantity>
long, moderate olive green (137 B) above, grayish yellow green (191 A to 191 B) below; secondary veins 1017 pairs; lateral leaflets 3.914.5 by
<quantity id="7A9300DFAFA8F145B34940013A18C4F0" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="3.4" metricValueMin="0.9" unit="cm" value="2.15" valueMax="3.4" valueMin="0.9">0.93.4 cm</quantity>
, 2.45.7 times as long as wide, lateral petiolules
<quantity id="4116448885809EAAF4EF16CF1916763A" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.5" metricValueMin="0.0" unit="cm" value="1.25" valueMax="2.5" valueMin="0.0">02.5 cm</quantity>
long, secondary veins 811 pairs; basal leaflets when present, 1.98.4 by
<quantity id="6E917A3FC4019D5590B42E0011B4BBA6" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="3.2" metricValueMin="0.4" unit="cm" value="1.8" valueMax="3.2" valueMin="0.4">0.43.2 cm</quantity>
, 2.34.0 times as long as wide, basal petiolules
<quantity id="B5B20ACBCDAA3B68E796BB1C3A1630F2" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="3.2" metricValueMin="0.4" unit="cm" value="1.8" valueMax="3.2" valueMin="0.4">0.43.2 cm</quantity>
; apex short to long acuminate, base attenuate, acute to short acuminate, or rounded, margin entire, but occasionally dentate when young; abaxial leaf surface sparsely to moderately pubescent, hairs whitish; petiole 2.0
<quantity id="76998F568FEFFD3881635144A936C0DA" metricMagnitude="-1" metricUnit="m" metricValue="1.03" unit="cm" value="10.3">10.3 cm</quantity>
long, round to tetragonal in cross-section, puberulent. Inflorescence lax paniculate, axis
<quantity id="64E7D713A83C6B93517AF6491DC1F94E" metricMagnitude="-1" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="2.35" metricValueMin="0.88" unit="cm" value="16.15" valueMax="23.5" valueMin="8.8">8.823.5 cm</quantity>
long, with primary branch at panicle base and along panicle axis, consisting of lateral cymes in lax clusters attached to the panicle axis or primary branch, with the oldest terminal flower often unfertilized at maturity, angular in cross-section, pubescent; bracteole triangular, up to
<quantity id="F5DC8E41A54643FCA8A76A7B8142AF1A" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
long, apex acute, tomentose. Calyx 5 - lobed, 5 - ribbed; lobes 0.21.3 by
<quantity id="0717B81F2A24C1015C2C1D09AEFB2858" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="12.5" metricValueMin="6.0" unit="mm" value="0.925" valueMax="1.25" valueMin="0.6">0.61.25 mm</quantity>
, acute, velutinous; flowering calyx 1.0
<quantity id="62DE288E12AC21D3925C669B19A4D1E8" metricMagnitude="-3" metricUnit="m" metricValue="2.49" unit="mm" value="2.49">2.49 mm</quantity>
long; fruiting calyx cup-shaped, 3.0
<quantity id="3E7B46A204916D08F666013BB4344AF6" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
diameter, covering 1 / 2 to 3 / 4 of the mature fruit. Corolla 5 - lobed, strong violet to light violet (N 88 B to N 88 C), outside covered with appressed hairs; lower lip broadly ovate to orbicular, 2.04.5 by
<quantity id="9BA3A656CC5F318ED697500283BE4F7F" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="4.5" metricValueMin="2.4" unit="mm" value="3.45" valueMax="4.5" valueMin="2.4">2.44.5 mm</quantity>
, apex acute to obtuse, margin entire, often patent, strong violet (N 89 B), yellowish at base, with white and yellowish hairs at corolla mouth; corolla throat inner diameter
<quantity id="0D872F5F20B89619361B6EDC183570A2" metricMagnitude="-3" metricUnit="m" metricValue="2.29" metricValueMax="2.53" metricValueMin="2.05" unit="mm" value="2.29" valueMax="2.53" valueMin="2.05">2.052.53 mm</quantity>
; lateral lobes 1.62.1 by 1.62.0 mm, apex round to acute, often patent to reflexed, light violet (N 88 C); upper lip 2 - lobed, lobes 1.31.7 by
<quantity id="68D64A5B943C7BD107C49DFED4431C4D" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.9" metricValueMin="1.4" unit="mm" value="1.65" valueMax="1.9" valueMin="1.4">1.41.9 mm</quantity>
, apex acute, often patent, light violet (N 88 D); tube infundibular
<quantity id="853071F751E29A62A984F8B0E349B657" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.7" metricValueMin="3.5" unit="mm" value="4.1" valueMax="4.7" valueMin="3.5">3.54.7 mm</quantity>
long, very pale purple (85 D) to very light purple (85 C). Stamens strongly didynamous, exserted up to 1 / 2 of its length, filaments
<quantity id="5EC7A874E82124B213711565BC25ABC2" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.3" metricValueMin="2.3" unit="mm" value="3.3" valueMax="4.3" valueMin="2.3">2.34.3 mm</quantity>
long, inserted 1 / 2 of the way on the corolla tube. Ovary 0.71.2 by
<quantity id="CCF3E1F8A4DD8FE62FE57A390192C86D" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 mm</quantity>
, globose, glabrous; style
<quantity id="5802EAF0A083D1618D64DF8BA200E648" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="6.6" metricValueMin="3.6" unit="mm" value="5.1" valueMax="6.6" valueMin="3.6">3.66.6 mm</quantity>
long; stigma lobes
<quantity id="13D442A69B744C5DC6EE1FDD6B24D2A2" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" unit="mm" value="0.65" valueMax="0.9" valueMin="0.4">0.40.9 mm</quantity>
long. Fruit brown when mature; dried ellipsoid to ovoid, 3.04.9 by 3.0
<quantity id="48FEB6230888596555E7A74283E0F1F7" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
, apex notched to truncated, glabrous with sparse glands.
</paragraph>
<caption id="A92126B6CE49AB45636DE8D328002BB5" ID-DOI="10.3897/phytokeys.248.120387.figure9" ID-arpha="F248A23D-AF02-5973-BC7E-8CCDCFDE6EDF" httpUri="https://binary.pensoft.net/fig/1158187" startId="F9">
<paragraph id="949EEDD52DE047F6D81B36F62F2122B4">
<label id="51B436C51CBBCA27532F1F611A30116A">Figure 9.</label>
</paragraph>
<paragraph id="A3F8098509FE06DD6F8F4D9F26D853CE">
Leaf undersurface pubescence of
<taxonomicName id="0D1A510589C0925DC3761820FCD150B6" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="0C0C6D152384AC5D8C3B50E8A35AE264" italics="true">V. arvensis</emphasis>
</taxonomicName>
(40 × magnification). Photos: L. E. Endonela.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXUDI" type="phenology">
<paragraph id="1D2DA44E2698AC3972E605F0C2463BC6">
<heading id="63D058543E85385F7CD0DB02FD546151" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="B32E5E1FB00636BB2C72716E73103E6C">Produces fruits and flowers all year round.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3UDI" type="habitat">
<paragraph id="B38F2E0664E2D4CBFF0D3299DFC9C5AB">
<heading id="9EA8AB8DA3187135D9553B74E49E91C1" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="DBE77CF65FEC0223FFB5C154F32424CA">
Growing primarily in farmlands, home gardens under cultivation as medicinal and / or hedge article, but recorded in secondary forests, mixed thickets, along rivers, trails and ridges, open places and in wastelands. Altitude:
<quantity id="4228FC5D0A143E7A82942838532AB98B" metricMagnitude="2" metricUnit="m" metricValue="2.575" metricValueMax="5.0" metricValueMin="0.15" unit="m" value="257.5" valueMax="500.0" valueMin="15.0">
15
<specimenCount id="2143EA5A8330E31D6E13A7D9B5199D5E" count="500" type="generic">500 m.</specimenCount>
</quantity>
</paragraph>
<caption id="215B4AF46B9C9849D3BFF68422A15474" ID-DOI="10.3897/phytokeys.248.120387.figure10" ID-arpha="6539BFD7-8372-5A0F-8DDB-9B4F7C22E0BA" httpUri="https://binary.pensoft.net/fig/1158188" startId="F10">
<paragraph id="F46EBBE4FE05B68820170C5782CCC792">
<label id="F7670FB5186E1AD4ED53A625F4E85D55">Figure 10.</label>
</paragraph>
<paragraph id="D320B7ED0909F800E0D383FE89CAB701">
Natural distribution of
<taxonomicName id="DEF665053C2C5721C4567B3288CA250F" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="7FE05F67E2A8867AD3A47C0F6F05F82A" italics="true">Vitex arvensis</emphasis>
</taxonomicName>
in the Philippines.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EZVDI" type="distribution">
<paragraph id="FA93A763600F768B8795468B9D785381">
<heading id="DF16928C26F85B144BDD6979248E4DF2" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A6F1B1CE05CD53E8F63DCC4154489B0F">
Endemic in the
<collectingCountry id="9B28BED6678CCF58C97229900718246A" name="Philippines">Philippines</collectingCountry>
, distributed primarily from Central to South Luzon but widespread throughout in cultivation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5VDI" type="Vernacular names">
<paragraph id="B1EC305C85334E7E36D4E2F278D3BD08">
<heading id="1C281E261CFBD10A26BF8E92D04ED429" reason="title">Vernacular names.</heading>
</paragraph>
<paragraph id="0D8F7B59848DC3A683BEA0689FBEB151">
lagundi (Tagalog, Bisaya), dangla (Ilokano,
<collectingRegion id="E51A0D07956042D09D63561728C377BE" country="Philippines" name="Zambales">Zambales</collectingRegion>
)
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDWDI" type="conservation status">
<paragraph id="A784588CAE344ED5CEAE49A16729958A">
<heading id="9E9D1967DD81F0D1A214A11C710C6AA6" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="59FC344A704C77B43C6809FB00FC41C7">
IUCN assessment gives the extent of occurrence (
<abbrev id="ABBRID0EJWDI" xlink_title="extent of occurrence">EOO</abbrev>
) and area of occurrence (
<abbrev id="ABBRID0ENWDI" xlink_title="area of occurrence">AOO</abbrev>
), as greater than the threshold for the vulnerable category, thus the status would be least concern (
<abbrev id="ABBRID0ERWDI" xlink_title="least concern">LC</abbrev>
).
</paragraph>
<caption id="9BB3D5263DCE05B0C94A73F196001115" ID-DOI="10.3897/phytokeys.248.120387.figure11" ID-arpha="69631B1A-3E95-5274-93DD-80E196D83954" httpUri="https://binary.pensoft.net/fig/1158189" startId="F11">
<paragraph id="B213D6AFD1A1F7F1E4AEDAEC4A806C00">
<label id="EA9DBF60743BC3497C2E59D1AF007812">Figure 11.</label>
</paragraph>
<paragraph id="0AE4CD553CE13C6AE8309A817E6D1695">
Holotype specimen of
<taxonomicName id="D439460C37FC0B4CF3FEDE86A4B573A7" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="C22645F60C00C7D501680AF5F6666B34" italics="true">Vitex arvensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ENXDI" type="notes">
<paragraph id="AD1B4CBC15C5AE2A0EA2FC3F64342693">
<heading id="7B102426336DD3BAFE30F6EC9604F8BC" reason="title">Notes.</heading>
</paragraph>
<paragraph id="8A550BED2A5E36BCEA7A4D02D23C5F3A">
1. This taxon has never been recognized at the species level before. The specific epithet
<taxonomicName id="48CB0C4FEDA72E3668F8575E966DAA53" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="2D289DC535EFFFF8F71437DD82D3199E" italics="true">arvensis</emphasis>
</taxonomicName>
provides insight into its usually cultivated nature since it is primarily cultivated on a commercial scale, and in home gardens as a medicinal plant.
</paragraph>
<paragraph id="526ED35A061ED624CF03FF069B2C127D">
2. This is a putative natural interspecific hybrid between
<taxonomicName id="B8F83A9E57C29F31DBCE8FEB572D2ECA" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="8CD494D146A97B701FCECFA593293A8C" italics="true">V. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4CB49DE30597CF807AC6B47E13C89D56" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="23C488A345778D7EAD8A5B59F5454E69" italics="true">V. elmeri</emphasis>
</taxonomicName>
. Although now distributed across the
<collectingCountry id="2CD37E98C3BAEA0B747BBFB4B749C975" name="Philippines">Philippines</collectingCountry>
, it was primarily distributed in areas where
<taxonomicName id="98D7EBC5BF7141F1E5375C9F92D9906F" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="D4A4985EDD3A64ADA0005A1BE92CFCC5" italics="true">V. elmeri</emphasis>
</taxonomicName>
and
<taxonomicName id="08D70F58F46E490E8F54CAB90B020A6B" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="BF0287907D367FD321F318AD1450D8D9" italics="true">V. bicolor</emphasis>
</taxonomicName>
overlap, from South to Central Luzon. This species bears resemblance to both parents:
<taxonomicName id="C6A2743124786171F5E2BA4C9DFADA22" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="elmeri">
<emphasis id="50408FE280AC362946BBD1096E352FAF" italics="true">V. elmeri</emphasis>
</taxonomicName>
in terms of the yellow corolla ridge color, presence of villous abaxial secondary veins, sparse pubescence on the lamina undersurface, and having dentate leaves when young; and
<taxonomicName id="8B57765FF62C4585A052E5C1B79382C8" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="102976002DAE18557CE97809B13F2127" italics="true">V. bicolor</emphasis>
</taxonomicName>
in terms of lax inflorescence units, flower color, and leaf size.
</paragraph>
<paragraph id="184439CB039FE08A9F7940BF65862C70">
3. Aside from bearing caducous flowers, this new species further differs from
<taxonomicName id="F0302BEEB59C804BFB52B38EB01FFDD2" authority="Willd." authorityName="Willd." class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="7BD785BB6887B52B41A47B426737061C" italics="true">Vitex bicolor</emphasis>
Willd.
</taxonomicName>
by having the mature ovoid fruits covered by the calyx by 1 / 23 / 4 along its length which dries brown, while
<taxonomicName id="0641F2BC6BB6D493A8F4FF987B71604C" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="D1ECEB53D4258E78FFDB88FDC5654FCD" italics="true">V. bicolor</emphasis>
</taxonomicName>
has larger (5.25.8 by
<quantity id="1E9D98A72ECB6F065F1F4CBEDCE3A2BB" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.6" metricValueMin="4.3" unit="mm" value="4.949999999999999" valueMax="5.6" valueMin="4.3">4.35.6 mm</quantity>
) globose fruits covered by the calyx to only up to 1 / 5 along its length which dries black.
</paragraph>
<paragraph id="80E1B7CFF63CBFD4AFD67F3A96A2A65A">
4. Unlike
<taxonomicName id="070A5D1C9EBA5455CBFF32080822EC9C" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="275D1C0FB325C10750EE9C767B5FEF97" italics="true">V. bicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="405FF3DF621946E654644A9F3DEEC7F3" authorityName="Gentallan, Sengun &amp; M. B. Bartolome" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="arvensis">
<emphasis id="0CE04579934DE9F004FF6F34A93085B1" italics="true">V. arvensis</emphasis>
</taxonomicName>
was observed to have a lower germination rate of seeds, with virtually non-existent seedlings under the canopy. However, with interventions, these can be germinated ex-situ, albeit with a low success rate.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ER2DI" type="Documented uses (in the Philippines)">
<paragraph id="7587ECAC240C0E145B6B515BFA3F3056">
<heading id="EA5471FA997E5E178114107DA4F4A113" reason="title">
Documented uses (in the
<collectingCountry id="66AFD59CDC3A3EE61D9D5D049CB62A00" name="Philippines">Philippines</collectingCountry>
).
</heading>
</paragraph>
<paragraph id="3BE5AA006958EE5D6DEFD8860AD49DC1">Leaves heated and applied as external patch or as liniment for fever, fruits and leaves used as an insecticide; juice extracted from leaves by grinding ingested for cough relief; decoction or boiling of leaves for fever and cough.</paragraph>
</subSubSection>
<subSubSection id="SECID0EW2DI" type="material">
<paragraph id="080A24EA976303F645AAFDF554242305">
<heading id="71C8DB344BCC229E117173DBB692917C" reason="title">Specimens examined.</heading>
</paragraph>
<paragraph id="BF6244BAD13EAE448493B0DEA94DA047">
<emphasis id="1C22CAF4A37760337BBDD1192EBD4E16" bold="true">
<collectingCountry id="FB41ED2055C052A4AA5F9251D2A528F6" name="Philippines">Philippines</collectingCountry>
.
<collectingRegion id="FA9246E36A7075C5B7A3891B38DA96C8" country="Philippines" name="Cagayan">Cagayan</collectingRegion>
</emphasis>
: Peñablanca, San Roque,
<date id="0693131E2613C2ACC22B176D126203F0" value="1917-05">May 1917</date>
, Adduru 173 (US 03805856); Vendivil &amp; Fernando 125366 (PNH); Peñablanca, Callao,
<date id="034243606D4CE668429576791D725A2B" value="1915-04">Apr 1915</date>
, Castillo 22747 (US 03805843); Tuao,
<date id="243871C9C9A0C8064BC657FCEEC7D99D" value="1979-05">May 1979</date>
, Sr. M. Rocero 165079 (PNH);
<emphasis id="CBA1B211936CDB7FAAE2AF9A4539F424" bold="true">
<collectingRegion id="34CDBB2DBB2A813072E1043A2755A24C" country="Philippines" name="Zambales">Zambales</collectingRegion>
</emphasis>
: Botolan,
<date id="A976F7E9B4D669210055E0DF515BCB08" value="1903-05">May 1903</date>
, Merrill 2917 (US 03805807); Botolan, Villar,
<date id="46DB40B7356A3F3AF1E42790CEB80570" value="1947-11">Nov 1947</date>
, R. B. Fox 4928 (PNH);
<emphasis id="5B8B32165AED4EA324E7D64A216FCB73" bold="true">
<collectingRegion id="932970FC603473B83FEFF90255D3E8DE" country="Philippines" name="Bataan">Bataan</collectingRegion>
</emphasis>
: Dinalupihan,
<date id="55AFD06B52F900EB438FEC0143EE39A2" value="1903-02">Feb 1903</date>
, Merrill 1503 (US 03805792);
<emphasis id="7CBCC4FE94AA405E371FB8BD399C8B21" bold="true">
<collectingRegion id="01AF7BC2821C08475F2D6D382C615661" country="Philippines" name="Pampanga">Pampanga</collectingRegion>
</emphasis>
: Arayat, Vidal 1648 (L. 2768325);
<emphasis id="25F52441584CA487AFFB67F098353710" bold="true">
<collectingRegion id="98EA5B81999697278A46566854FBDCC0" country="Philippines" name="Bulacan">Bulacan</collectingRegion>
</emphasis>
: San Miguel,
<quantity id="A1AE80FB36E29B25E76C602F1105524F" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
alt., B. Fegan 113564 (PNH); Doña Remedios Trinidad, Kabayanan,
<date id="66174DB120768AB260D243F06510D755" value="1994-12">Dec 1994</date>
, H. Garcia et al. 15334 (L. 4212458); Norzagaray, San Lorenzo,
<date id="41BDA064B7BB605C4A0081B08DBC6950" value="1996-03">Mar 1996</date>
, E. Barbon &amp; H. Garcia 22403 (L. 4216246);
<emphasis id="DD5142DAB94455EF49D4416989DAFF9F" bold="true">
<collectingRegion id="1A3304702240C7D2BBDD127B55D65FFD" country="Philippines" name="Manila">Manila</collectingRegion>
</emphasis>
:
<collectingRegion id="AEE0C93FF95C8D2C80AE089EF34F4045" country="Philippines" name="Manila">Manila</collectingRegion>
,
<date id="B2ABC6E96BA877EC45BFAA3730A0D599" value="1903-10">Oct 1903</date>
, Merrill 3429 (US 03805814);
<date id="53C5D5522CC6841AE9B7ABFD73D5C185" value="1910-09">Sep 1910</date>
, M. Ramos (E 01121257); near
<collectingRegion id="3E318ADCC596C5B83EDF38192BD2FBCA" country="Philippines" name="Manila">Manila</collectingRegion>
, Capt. Wilkes s. n. (US 00830006); Sampaloc, Balic-balic,
<date id="A40FA8A386D25DAF568B659CE93B7CD6" value="1890-08">Aug 1890</date>
, Loher 4433 (US 03805840);
<collectingRegion id="8918E2EFE0E5D1300916568EA3DC3E10" country="Philippines" name="Mandaluyong">Mandaluyong</collectingRegion>
, Malamig, G. Edaño 3783 (PNH);
<emphasis id="E02B2C6F41B036FE6138C55C1D9E83BF" bold="true">
<collectingRegion id="B71CD1CF94F8A30E4C67364FCD10060E" country="Philippines" name="Cavite">Cavite</collectingRegion>
</emphasis>
: Mendez, Mangubat 1343 (US 03805791);
<emphasis id="07FDF13D0A347C89ABBB9294CF926558" bold="true">
<collectingRegion id="6159A458468B2A5F39BE6BC144F8DF2E" country="Philippines" name="Laguna">Laguna</collectingRegion>
</emphasis>
: Los Baños,
<date id="643DA9D4E6295920F191B7ACE71F5E6C" value="1903-12">Dec 1903</date>
, Hallier 4055 b (L. 2775187);
<date id="4BE3BDDA9887891618F052BEE405EB15" value="1906-04">Apr 1906</date>
, Elmer 8125 (E 01121256);
<date id="300C1C18638F33CC421F2FEF15DA7DD8" value="1909-09">Sep 1909</date>
, Rosenbluth &amp; Tamesis (L. 2768326);
<quantity id="3669C4DEBB33278A22DABECF916B853B" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</quantity>
alt.,
<date id="EC52F344B26CB02B91435FC4A9B09539" value="1911-01">Jan 1911</date>
, Holman 99 (US 03805805, US 03805808);
<date id="B6E4A7330E64AB7606ED712F6664EA82" value="1917-07">Jul 1917</date>
, Elmer 18119 (US 03805788, L. 2768218); Makiling National Park,
<quantity id="DA62FCB3854A046EF1F6EEE160DA36C5" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
alt.,
<date id="41284650D85DA22C65AA821F5571BD7D" value="1945-10">Oct 1945</date>
, M. D. Sulit 8311 (PNH);
<date id="A7D2050D818D94B37660FEE14FE91342" value="1963-03">Mar 1963</date>
, D. R. Mendoza 92452 (PNH); San Antonio,
<date id="387EE7574911D0670F9125908F42550D" value="1948-03">Mar 1948</date>
, E. Canicosa 9707 (PNH); College,
<quantity id="2FD81C80AD1F3CB4BA981AAEEA824D35" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
alt.,
<date id="A09293F788FD22DDCCFF946F7931685D" value="1954-09">Sep 1954</date>
. J. V. Pancho 32919 (PNH);
<emphasis id="183859CDEADFE27C470DC343A58B9865" bold="true">
<collectingRegion id="C84816EFB521A94CA5442D2826B8C566" country="Philippines" name="Batangas">Batangas</collectingRegion>
</emphasis>
: Mataas na Kahoy, Arañez 14 (L. 2768323);
<emphasis id="6FC66104AE29B89B5C1EA631E2B7556A" bold="true">
<collectingRegion id="CCDF591097080DD337E4A3E4592643DC" country="Philippines" name="Rizal">Rizal</collectingRegion>
</emphasis>
:
<collectingRegion id="42E7864749CF08321FCCB33D25D2EB0D" country="Philippines" name="Marikina">Marikina</collectingRegion>
, Barangka, G. Edaño 36210 (PNH);
<collectingRegion id="DAE0DDAAB3C5874CA7107E36397079C6" country="Philippines" name="Antipolo">Antipolo</collectingRegion>
,
<quantity id="2A7189CFD0F9FB7CDA0CCEFBED7FEECC" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
alt.,
<date id="AC07E908F002069983FD7EBFE2177955" value="1903-03">Mar 1903</date>
, Merrill 1636 (US 03805812);
<date id="E4407C3851182013D87F4976179F7898" value="1913-01">Jan 1913</date>
Merr. Sp. Blanc. 440 (L. 2768327);
<collectingRegion id="1864FBCB3AB9620C3F1A199FD1AE6534" country="Philippines" name="Antipolo">Antipolo</collectingRegion>
, Boso-boso, Loher 4432 (US 03805810); San Mateo,
<date id="4445D600D0353F24F8EADC56305AD013" value="1904-05">May 1904</date>
, Aherns collector 102 (US 03805809); Taytay,
<date id="27A396EF289341B0BAA1D8A562D4DBAC" value="1953-02">Feb 1953</date>
, Lorena 18 (L. 0248373); Tanay,
<date id="B639BD4E8162D835C9CB3FFE90BA06A9" value="1903-05">May 1903</date>
, Merrill 2320 (US 03805790);
<emphasis id="A0EF0FF04D27AFFCA3A5B4627F016734" bold="true">
<collectingRegion id="C27954B54144E8BCDCD51BC3A40347D5" country="Philippines" name="Quezon">Quezon</collectingRegion>
</emphasis>
: Tayabas,
<date id="54754E0CC33D6BDEF32FE2C9158E2E21" value="1916-05">May 1916</date>
, Cailipan 25637 (US 03805789); Infanta,
<date id="565743CFF7923085326C892DB2813D1A" value="1904-09">Sep 1904</date>
, Whitford 853 (US 03805784);
<emphasis id="BDCE6DCCFB08CF2722D45F76F2F083A7" bold="true">
<collectingRegion id="ED0F455DD4BC8BCC09256D38E8FA678C" country="Philippines" name="Camarines Sur">Camarines Sur</collectingRegion>
</emphasis>
: Pasacao, Dalupaon, 1901, Ahern 255 (BO 1616593, US 03805779),
<collectingRegion id="F30DED3CD8B9521947FDF6D9C9AFDD73" country="Philippines" name="Naga">Naga</collectingRegion>
City, Mt. Isarog,
<date id="87E858AB7A99189F5607FDE1F327BFE9" value="1992-10">Oct 1992</date>
, Barbon et al. 8377 (L. 3930320);
<emphasis id="CDAEB973FD6A9395C99025BF723AB98B" bold="true">Agusan Norte</emphasis>
:
<collectingRegion id="A57A855A1A8DB662D47D5EC89805EC9A" country="Philippines" name="Butuan">Butuan City</collectingRegion>
, Tungao,
<date id="C0B1031116D425D93C26A65CB2313448" value="1991-05">May 1991</date>
, Barbon et al. 1917 (L. 4212463).
</paragraph>
</subSubSection>
</treatment>
</document>