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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1072.67935" ID-GBIF-Dataset="0a9353f0-1f24-45f0-8ddc-bd941470a567" ID-PMC="PMC8626413" ID-Pensoft-Pub="1313-2970-1072-1" ID-Pensoft-UUID="DDB04884FF7C52BD9D1814C0EA84E496" ID-PubMed="34899006" ID-ZooBank="9F578799E05F44118E2D378E3C17F3E3" ModsDocID="1313-2970-1072-1" checkinTime="1637381493323" checkinUser="pensoft" docAuthor="Li, Ya, Li, Shuqiang &amp; Lin, Yucheng" docDate="2021" docId="C9D976B150CD5F7D83547E2E41A9F771" docLanguage="en" docName="ZooKeys 1072: 1-47" docOrigin="ZooKeys 1072" docPubDate="2021-11-19" docSource="http://dx.doi.org/10.3897/zookeys.1072.67935" docTitle="Patu qiqi Miller, Griswold &amp; Yin 2009" docType="treatment" docVersion="4" id="DDB04884FF7C52BD9D1814C0EA84E496" lastPageNumber="1" masterDocId="DDB04884FF7C52BD9D1814C0EA84E496" masterDocTitle="Taxonomic study on fourteen symphytognathid species from Asia (Araneae, Symphytognathidae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="1" updateTime="1668151161727" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic study on fourteen symphytognathid species from Asia (Araneae, Symphytognathidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Ya</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1097-6192</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Yucheng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5054-0633</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China &amp; The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<mods:nameIdentifier type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1072</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>47</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1072.67935</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1072.67935</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1072-1</mods:identifier>
<mods:identifier type="ZooBank">9F578799E05F44118E2D378E3C17F3E3</mods:identifier>
<mods:identifier type="Pensoft-UUID">DDB04884FF7C52BD9D1814C0EA84E496</mods:identifier>
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<treatment ID-GBIF-Taxon="190343155" LSID="urn:lsid:plazi:treatment:C9D976B150CD5F7D83547E2E41A9F771" httpUri="http://treatment.plazi.org/id/C9D976B150CD5F7D83547E2E41A9F771" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="C9D976B1-50CD-5F7D-8354-7E2E41A9F771" authority="Miller, Griswold &amp; Yin, 2009" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu qiqi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="qiqi">Patu qiqi Miller, Griswold &amp; Yin, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Patu qiqi A female habitus, dorsal B female habitus, ventral C female habitus, lateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Abbreviations: CD = copulatory ducts; FD = fertilisation ducts; S = spermathecae; Sp = scape. Scale bars: 0.50 (A-C); 0.10 (D-G)." figureDoi="10.3897/zookeys.1072.67935.figure12" httpUri="https://binary.pensoft.net/fig/612454" pageId="0" pageNumber="1">Figures 12</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of fourteen species of genera Patu, Kirinua gen. nov. and Swilda gen. nov. in Asia 1 P. catba sp. nov. 2 P. dakou sp. nov. 3 P. damtao sp. nov. 4 P. jiangzhou sp. nov. 5 P. jidanweishi 6 P. nagarat sp. nov. 7 P. nigeri 8 P. putao sp. nov. 9 P. qiqi 10 P. xiaoxiao 11 K. maguai sp. nov. 12 K. yangshuo sp. nov. 13 S. longtou 14 S. spinathoraxi." figureDoi="10.3897/zookeys.1072.67935.figure23" httpUri="https://binary.pensoft.net/fig/612465" pageId="0" pageNumber="1">, 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu qiqi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="qiqi">Patu qiqi</taxonomicName>
Miller, Griswold &amp; Yin, 2009: 66, figs 65F-H, 67E, F, 73A and B (♂♀).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♀ (CASENT 9029328, HNU) and
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratypes</emphasis>
5♀, 2 juv. (CASENT 9029327, HNU)
<emphasis bold="true" pageId="0" pageNumber="1">China</emphasis>
: Yunnan Province, Gaoligongshan, Nujiang Prefecture, Nujiang State Nature Reserve, Qiqihe, 9.9 air km W of Gongshan (
<geoCoordinate degrees="27.715" direction="north" orientation="latitude" precision="55" value="27.715">27.715°N</geoCoordinate>
,
<geoCoordinate degrees="98.565" direction="east" orientation="longitude" precision="55" value="98.565">98.565°E</geoCoordinate>
; 2000 m alt.), 9-14.VII.2000, H. Yan et al. leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2018-08-13" collectorName="Y. Lin" country="China" county="Gongshan County" elevation="2525" latitude="27.8784" location="Dulongjiang Town" longLatPrecision="1" longitude="98.42274" municipality="Gongshan County" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan Province">
<specimenCount type="female">2♀</specimenCount>
(NHMSU-HA122)
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="China">China</collectingCountry>
</emphasis>
:
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCounty>Gongshan County</collectingCounty>
, at
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.4" unit="km" value="54.0">54 km</quantity>
of from
<collectingMunicipality>Gongshan County</collectingMunicipality>
to
<location LSID="urn:lsid:plazi:treatment:C9D976B150CD5F7D83547E2E41A9F771:8856CCB45535E926B1C5B4F1DEC8F83A" country="China" county="Gongshan County" latitude="27.8784" longLatPrecision="1" longitude="98.42274" municipality="Gongshan County" name="Dulongjiang Town" stateProvince="Yunnan Province">Dulongjiang Town</location>
, in primary forest, leaf litter (
<geoCoordinate degrees="27.87840" direction="north" orientation="latitude" precision="1" value="27.8784">27.87840°N</geoCoordinate>
,
<geoCoordinate degrees="98.42274" direction="east" orientation="longitude" precision="1" value="98.42274">98.42274°E</geoCoordinate>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.525" unit="m" value="2525.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.525" unit="m" value="2525.0">2525 m</elevation>
</quantity>
alt.),
<collectingDate value="2018-08-13">13.VIII.2018</collectingDate>
,
<collectorName>Y. Lin</collectorName>
et al. leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The male differs from other
<taxonomicName authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Patu</emphasis>
</taxonomicName>
species, with the exception of
<taxonomicName lsidName="P. nigeri" pageId="0" pageNumber="1" rank="species" species="nigeri">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigeri</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. silho" pageId="0" pageNumber="1" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="1">P. silho</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. xiaoxiao" pageId="0" pageNumber="1" rank="species" species="xiaoxiao">
<emphasis italics="true" pageId="0" pageNumber="1">P. xiaoxiao</emphasis>
</taxonomicName>
, by the palp with an unexposed embolus (fig. 73A and B in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
). It differs by having a hooked median apophysis vs. lacking in
<taxonomicName lsidName="P. nigeri" pageId="0" pageNumber="1" rank="species" species="nigeri">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigeri</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. silho" pageId="0" pageNumber="1" rank="species" species="silho">
<emphasis italics="true" pageId="0" pageNumber="1">P. silho</emphasis>
</taxonomicName>
(cf. fig. 73A in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
and fig. 4A and B in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2154.1.3" author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="47 - 68" refId="B11" refString="Lin, Y, Li, S, 2009. First described Patu spiders (Araneae, Symphytognathidae) from Asia. Zootaxa 2154: 47 - 68, DOI: https://doi.org/10.11646/zootaxa.2154.1.3" title="First described Patu spiders (Araneae, Symphytognathidae) from Asia." url="https://doi.org/10.11646/zootaxa.2154.1.3" volume="2154" year="2009">Lin and Li 2009</bibRefCitation>
and fig. 5A and B in
<bibRefCitation author="Saaristo, MI" journalOrPublisher="Phelsuma" pageId="0" pageNumber="1" pagination="53 - 56" refId="B17" refString="Saaristo, MI, 1996. Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles. Phelsuma 4: 53 - 56" title="Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles." volume="4" year="1996">Saaristo 1996</bibRefCitation>
) and it differs from
<taxonomicName lsidName="P. xiaoxiao" pageId="0" pageNumber="1" rank="species" species="xiaoxiao">
<emphasis italics="true" pageId="0" pageNumber="1">P. xiaoxiao</emphasis>
</taxonomicName>
by lacking a tegular process (cf. fig. 73A in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
and Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu xiaoxiao A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: CD = copulatory ducts; Co = conductor; CO = copulatory opening; E = embolus; FD = fertilisation ducts; Fe = femur; MA = median apophysis; Pa = patella; Pl = parmula; S = spermathecae; T = tegulum; Ti = tibia; TP = tegular process. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1072.67935.figure14" httpUri="https://binary.pensoft.net/fig/612456" pageId="0" pageNumber="1">14A</figureCitation>
). The female is most similar to that of
<taxonomicName lsidName="P. nigeri" pageId="0" pageNumber="1" rank="species" species="nigeri">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigeri</emphasis>
</taxonomicName>
in the shape of the epigyne and the configuration of the vulva, but it can be easily distinguished by the indistinct scape and the hidden copulatory openings vs. a tongue-shaped parmula and exposed copulatory openings (cf. Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Patu qiqi A female habitus, dorsal B female habitus, ventral C female habitus, lateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Abbreviations: CD = copulatory ducts; FD = fertilisation ducts; S = spermathecae; Sp = scape. Scale bars: 0.50 (A-C); 0.10 (D-G)." figureDoi="10.3897/zookeys.1072.67935.figure12" httpUri="https://binary.pensoft.net/fig/612454" pageId="0" pageNumber="1">12F-G</figureCitation>
vs. 10F-G).
</paragraph>
<caption doi="10.3897/zookeys.1072.67935.figure12" httpUri="https://binary.pensoft.net/fig/612454" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu qiqi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="qiqi">
<emphasis italics="true" pageId="0" pageNumber="1">Patu qiqi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
female habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
female habitus, ventral
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
female habitus, lateral
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
epigyne, ventral
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
epigyne, lateral
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
vulva, ventral
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
vulva, dorsal. Abbreviations: CD = copulatory ducts; FD = fertilisation ducts; S = spermathecae; Sp = scape. Scale bars: 0.50 (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
); 0.10 (
<emphasis bold="true" pageId="0" pageNumber="1">D-G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
See Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Patu qiqi A female habitus, dorsal B female habitus, ventral C female habitus, lateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Abbreviations: CD = copulatory ducts; FD = fertilisation ducts; S = spermathecae; Sp = scape. Scale bars: 0.50 (A-C); 0.10 (D-G)." figureDoi="10.3897/zookeys.1072.67935.figure12" httpUri="https://binary.pensoft.net/fig/612454" pageId="0" pageNumber="1">12A-G</figureCitation>
and
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="9 - 195" refId="B16" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160Rivera-Quiroz" volume="11" year="2009">Miller et al. (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
China (Yunnan) (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Distribution records of fourteen species of genera Patu, Kirinua gen. nov. and Swilda gen. nov. in Asia 1 P. catba sp. nov. 2 P. dakou sp. nov. 3 P. damtao sp. nov. 4 P. jiangzhou sp. nov. 5 P. jidanweishi 6 P. nagarat sp. nov. 7 P. nigeri 8 P. putao sp. nov. 9 P. qiqi 10 P. xiaoxiao 11 K. maguai sp. nov. 12 K. yangshuo sp. nov. 13 S. longtou 14 S. spinathoraxi." figureDoi="10.3897/zookeys.1072.67935.figure23" httpUri="https://binary.pensoft.net/fig/612465" pageId="0" pageNumber="1">23</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

67
data/C9/D9/C5/C9D9C5FB5616202BB463FDF01EC5F23D.xml Normal file
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@ -0,0 +1,67 @@
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<mods:part id="CFB2C41EC3F071CA52C4EA2B92BF1C4C">
<mods:date id="6B036CBE03B888446B14AAD4937D6E1A">2016</mods:date>
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<taxonomicName id="68E86453CAFDCFF1409A0F76B4F95A55" class="Insecta" family="Ichneumonidae" genus="Homotropus" kingdom="Animalia" lsidName="Homotropus nigricornis" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="nigricornis">Homotropus nigricornis</taxonomicName>
Thomson, 1890
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<paragraph id="4D55D3A656CA20CD9C208298446ED05E" pageId="0" pageNumber="9042">Distribution</paragraph>
<paragraph id="2E4E70DE160E265431ED5998395FA2E8" pageId="0" pageNumber="9042">Ireland</paragraph>
</subSubSection>
<subSubSection id="936B14371CDF20F26D6D78F49343F97D" pageId="0" pageNumber="9042" type="notes">
<paragraph id="03380E3E9900B7C85AAE8A70ADDD9AEB" pageId="0" pageNumber="9042">Notes</paragraph>
<paragraph id="76FB9B426504ACEB697E52C70604C8FA" pageId="0" pageNumber="9042">
<bibRefCitation id="EA850FE919AA8526EFE2B6B585BF95C0" author="Fitton, M. G." journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="9042" pagination="311 - 320" title="A key to the European genera of diplazontine ichneumon-flies, with notes on the British fauna" volume="7" year="1982">Fitton and Rotheray (1982)</bibRefCitation>
doubted whether the few British specimens identified as
<taxonomicName id="C38792EA6AF7EB27708E4D9E50A97CE0" class="Insecta" family="Ichneumonidae" genus="Ethelurgina" kingdom="Animalia" lsidName="Ethelurgina nigricorne" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="nigricorne">nigricorne</taxonomicName>
were distinct from ornatum. One apparently British specimen in BMNH (Capron coll., no other details) is
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(det. Broad and Klopfstein).
<bibRefCitation id="541614DACD4F7604BF23AF00D6A2BB5A" author="Beirne, B. P." journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="9042" pagination="661 - 712" title="British species of Diplazonini (Bassini auctt.) with a study of the genital and postgenital abdominal sclerites in the male (Hym.: Ichneum.)" volume="91" year="1941">Beirne (1941)</bibRefCitation>
records
<taxonomicName id="0E160E7278263945080A9401945C7528" class="Insecta" family="Ichneumonidae" genus="Ethelurgina" kingdom="Animalia" lsidName="Ethelurgina nigricorne" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="nigricorne">nigricorne</taxonomicName>
from Ireland.
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857
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</quantity>
,
<date value="2014-08-20">
<collectingDate value="2014-08-20">20 Aug. 2014</collectingDate>
</date>
,
<collectorName>H. Naka</collectorName>
leg.
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Other specimens</emphasis>
.
</emphasis>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3063628441" collectingDate="1960-07-19" collectionCode="T" collectorName="Ebato" country="Japan" location="To" municipality="Mt Takao" specimenCount="1" specimenCount-female="1" stateProvince="Tokyo" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Mt Takao</collectingMunicipality>
,
<collectingRegion country="Japan" name="Tokyo">Tokyo</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:E4F2C74E12CAE267919D1C01534B923A" country="Japan" municipality="Mt Takao" name="To" stateProvince="Tokyo">To</location>
,
<date value="1960-07-19">
<collectingDate value="1960-07-19">19 Jul. 1960</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ebato</collectorName>
leg. (NSMT-I-L-75536)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628425" collectingDate="1953-05-23" collectionCode="T" collectorName="Ebato" country="Japan" location="Nashimoto" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:491725448E8480809463AC79EB9719D5" country="Japan" name="Nashimoto" stateProvince="Shizuoka">Nashimoto</location>
,
<collectingRegion country="Japan" name="Shizuoka">Shizuoka Pref.</collectingRegion>
,
<date value="1953-05-23">
<collectingDate value="1953-05-23">23 May 1953</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ebato</collectorName>
leg. (NSMT-I-L-75538)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628398" collectingDate="1959-06-05" collectionCode="T" collectorName="Ebato" country="Japan" location="Nashimoto" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
, ditto,
<date value="1959-06-05">
<collectingDate value="1959-06-05">5 Jun. 1959</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ebato</collectorName>
leg. (NSMT-I-L-75537)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628345" collectingDate="1961-06-10" collectionCode="T" collectorName="Ebato" country="Japan" location="Nashimoto" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
, ditto,
<date value="1961-06-10">
<collectingDate value="1961-06-10">10 Jun. 1961</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ebato</collectorName>
leg. (NSMT-I-L-75539)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628325" collectingDate="1966-08-24" collectionCode="T" collectorName="Ebato" country="Japan" location="Nashimoto" specimenCount="2" specimenCount-male="2" stateProvince="Shizuoka" typeStatus="holotype">
<specimenCount type="male">2♂</specimenCount>
, ditto,
<date value="1966-08-24">
<collectingDate value="1966-08-24">24 Aug. 1966</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ebato</collectorName>
leg. (NSMT-I-L-75534, 75535)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628339" collectingDate="1993-06-26" collectorName="A. Sasaki" country="Japan" location="Nukata Town" municipality="Kuragari-Valley" specimenCount="1" specimenCount-female="1" stateProvince="Aichi" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Kuragari-Valley</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:EF5AD67BF2C298B19839A2AE097E20E0" country="Japan" municipality="Kuragari-Valley" name="Nukata Town" stateProvince="Aichi">Nukata Town</location>
,
<collectingRegion country="Japan" name="Aichi">Aichi Pref.</collectingRegion>
,
<date value="1993-06-26">
<collectingDate value="1993-06-26">26 Jun. 1993</collectingDate>
</date>
,
<collectorName>A. Sasaki</collectorName>
leg. (NSMT-I-L-75593)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628373" collectingDate="1993-06-27" collectionCode="T" collectorName="Ikenoue" country="Japan" location="Tokuyama City" municipality="Sugano" specimenCount="1" specimenCount-male="1" stateProvince="Yamaguchi" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Sugano</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:7AACDBE1ACB421AA1BFD2496A192F43C" country="Japan" municipality="Sugano" name="Tokuyama City" stateProvince="Yamaguchi">Tokuyama City</location>
,
<collectingRegion country="Japan" name="Yamaguchi">Yamaguchi Pref.</collectingRegion>
,
<date value="1993-06-27">
<collectingDate value="1993-06-27">27 Jun. 1993</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Ikenoue</collectorName>
leg. (NSMT-I-L-75594)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628312" collectingDate="1992-05-28" collectorName="Y. Yanagita" country="Japan" location="Aira Town" municipality="Shimomyo" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Shimomyo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:95A2AE9F1C2B55E22A5ABD434D173AD3" country="Japan" municipality="Shimomyo" name="Aira Town" stateProvince="Kagoshima">Aira Town</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1992-05-28">
<collectingDate value="1992-05-28">28 May 1992</collectingDate>
</date>
,
<collectorName>Y. Yanagita</collectorName>
leg. (NSMT-I-L-75596)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628411" collectingDate="1992-07-14" collectorName="Y. Yanagita" country="Japan" location="Izumi City" municipality="Kamitsuru" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Kamitsuru</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:E3AB572BD003B439CED0288076968155" country="Japan" municipality="Kamitsuru" name="Izumi City" stateProvince="Kagoshima">Izumi City</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1992-07-14">
<collectingDate value="1992-07-14">14 Jul. 1992</collectingDate>
</date>
,
<collectorName>Y. Yanagita</collectorName>
leg. (NSMT-I-L-75595)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628423" collectingDate="1958-08-05" collectionCode="T" collectorName="I. Yakushima" country="Japan" location="Mt Ishizukadake" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:66AA56C349670FA5ADB685E30A927762" country="Japan" name="Mt Ishizukadake" stateProvince="Kagoshima">Mt Ishizukadake</location>
,
<collectorName>I. Yakushima</collectorName>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1958-08-05">
<collectingDate value="1958-08-05">5 Aug. 1958</collectingDate>
</date>
, B.
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. leg. (NSMT-I-L-75607)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628367" collectingDate="2006-10-03" collectionCode="T" collectorName="M. Owada &amp; Fukuda" country="Japan" location="Yakushima" municipality="Nagata" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Nagata</collectingMunicipality>
, I.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:61AC027943D260EADD48183D295966E0" country="Japan" municipality="Nagata" name="Yakushima" stateProvince="Kagoshima">Yakushima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="2006-10-03">
<collectingDate value="2006-10-03">3 Oct. 2006</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
and
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Fukuda</collectorName>
leg. (NSMT-I-L-75606)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628332" collectingDate="1988-10-13" collectorName="Uken, I &amp; M. Owada" country="Japan" location="Amamiohshima" municipality="Chuo-rindo" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Chuo-rindo</collectingMunicipality>
,
<collectorName>Uken, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:A68732C105CDE3A358C7CC7F6C268D97" country="Japan" municipality="Chuo-rindo" name="Amamiohshima" stateProvince="Kagoshima">Amamiohshima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1988-10-13">
<collectingDate value="1988-10-13">13 Oct. 1988</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75541)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628413" collectingDate="2009-04-22" collectorName="M. Owada" country="Japan" location="Amamiohshima" municipality="Chuo-rindo" specimenCount="3" specimenCount-male="3" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">3♂</specimenCount>
, ditto,
<date value="2009-04-22">
<collectingDate value="2009-04-22">22 Apr. 2009</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75609 to 75611)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628310" collectingDate="1988-10-11" collectorName="Naze, I &amp; M. Owada" country="Japan" location="Amamiohshima" municipality="Kinsakubaru" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Kinsakubaru</collectingMunicipality>
,
<collectorName>Naze, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:C0960611D9BFEE952DA19BD0DC36B6B7" country="Japan" municipality="Kinsakubaru" name="Amamiohshima" stateProvince="Kagoshima">Amamiohshima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1988-10-11">
<collectingDate value="1988-10-11">11 Oct. 1988</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75542)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628451" collectingDate="1988-10-12" collectorName="I. Amamiohshima &amp; M. Owada" country="Japan" location="Mt Yuwan-dake" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:5F9F55AFBEE0AF702E7F8CB87DBAEAB4" country="Japan" name="Mt Yuwan-dake" stateProvince="Kagoshima">Mt Yuwan-dake</location>
,
<collectorName>I. Amamiohshima</collectorName>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1988-10-12">
<collectingDate value="1988-10-12">12 Oct. 1988</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75543)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628301" collectingDate="1968-06-25" collectorName="Naze, I &amp; Y. Kishida" country="Japan" location="Amamiohshima" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectorName>Naze, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:E0368D4685CD55AA31F42954AD1D455A" country="Japan" name="Amamiohshima" stateProvince="Kagoshima">Amamiohshima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1968-06-25">
<collectingDate value="1968-06-25">25 Jun. 1968</collectingDate>
</date>
,
<collectorName>Y. Kishida</collectorName>
leg. (NSMT-I-L-75540)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628376" collectingDate="2009-04-21" collectorName="Setouchi, I &amp; M. Owada" country="Japan" location="Amamiohshima" municipality="Shinokawa" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Shinokawa</collectingMunicipality>
,
<collectorName>Setouchi, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:CCEEC5B8F78A8E49788DB4B8C64263CE" country="Japan" municipality="Shinokawa" name="Amamiohshima" stateProvince="Kagoshima">Amamiohshima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="2009-04-21">
<collectingDate value="2009-04-21">21 Apr. 2009</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75608)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628302" collectingDate="1992-10-31" collectorName="Mikyo, I &amp; M. Owada" country="Japan" location="Tokunoshima" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<collectorName>Mikyo, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:0C14701698AF53A13409370C805BDF3F" country="Japan" name="Tokunoshima" stateProvince="Kagoshima">Tokunoshima</location>
,
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<date value="1992-10-31">
<collectingDate value="1992-10-31">31 Oct. 1992</collectingDate>
</date>
,
<collectorName>M. Owada</collectorName>
leg. (NSMT-I-L-75544)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628340" collectingDate="1974-03-30" collectionCode="T" collectorName="I. Okinawajima &amp; Naito" country="Japan" location="Gogayama" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:FA3697A894B1A4323260A3D4A1FD78BC" country="Japan" name="Gogayama" stateProvince="Okinawa">Gogayama</location>
,
<collectorName>I. Okinawajima</collectorName>
,
<collectingRegion country="Japan" name="Okinawa">Okinawa Pref.</collectingRegion>
,
<date value="1974-03-30">
<collectingDate value="1974-03-30">30 Mar. 1974</collectingDate>
</date>
,
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName>Naito</collectorName>
leg. (NSMT-I-L-75612)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628477" collectingDate="1980-08-16" collectionCode="R" collectorName="I. Okinawajima &amp; Sato" country="Japan" location="Chinen-son" municipality="Seifuautaki" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Seifuautaki</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:C3695E4581ED60611BA0FB404FA27B3B" country="Japan" municipality="Seifuautaki" name="Chinen-son" stateProvince="Okinawa">Chinen-son</location>
,
<collectorName>I. Okinawajima</collectorName>
,
<collectingRegion country="Japan" name="Okinawa">Okinawa Pref.</collectingRegion>
,
<date value="1980-08-16">
<collectingDate value="1980-08-16">16 Aug. 1980</collectingDate>
</date>
,
<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName>Sato</collectorName>
leg. (NSMT-I-L-75613)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628354" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="R" collectorName="I. Okinawajima &amp; Sato" country="Japan" location="Chinen-son" municipality="Seifuautaki" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount type="male">1♂</specimenCount>
, same data as for preceding (NSMT-I-L-75614)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063628466" collectingDate="2001-04-21" collectorName="Kunigami-son, I &amp; A. Sasaki" country="Japan" location="Okinawajima" municipality="Okuni-bashi" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount type="female">1♀</specimenCount>
,
<normalizedToken originalValue="Ôkuni-bashi">
<collectingMunicipality>Okuni-bashi</collectingMunicipality>
</normalizedToken>
,
<collectorName>Kunigami-son, I</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:FFD09F19B6EB83421417EAE4AF3C2180" country="Japan" municipality="Okuni-bashi" name="Okinawajima" stateProvince="Okinawa">Okinawajima</location>
,
<collectingRegion country="Japan" name="Okinawa">Okinawa Pref.</collectingRegion>
,
<date value="2001-04-21">
<collectingDate value="2001-04-21">21 Apr. 2001</collectingDate>
</date>
,
<collectorName>A. Sasaki</collectorName>
leg.
</materialsCitation>
(NSMT-I-L-75615).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="etymology">
<paragraph pageId="0" pageNumber="171">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="171">
The specific epithet refers to the darker wing color in comparison to that of
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
, and the habitat of this species is a shaded place.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="diagnosis">
<paragraph pageId="0" pageNumber="171">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="171">
This new species is similar to
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="N. quadrimaculalis" pageId="0" pageNumber="171" rank="species" species="quadrimaculalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. quadrimaculalis</emphasis>
</taxonomicName>
, also distributed in
<collectingCountry name="Japan">Japan</collectingCountry>
, but it can be distinguished by the following characters: length of forewing 12.0-13.0 mm; vertex with erect, dull-orange scales; subdiscal white spot of forewing rounded, small, and blurry; base of discal cell of hindwing identical to ground color; dorsal margin of valva of male genitalia slightly incurved subapically; anterior apophysis of female genitalia straight and narrow; signum of female genitalia nearly elliptical, larger than in
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
(diameter
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.1500000000000001" metricValueMax="1.4000000000000001" metricValueMin="0.8999999999999999" unit="mm" value="0.115" valueMax="0.14" valueMin="0.09">0.09-0.14 mm</quantity>
). This species is also similar to
<taxonomicName lsidName="N. occultalis" pageId="0" pageNumber="171" rank="species" species="occultalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. occultalis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="N. bispina" pageId="0" pageNumber="171" rank="species" species="bispina">
<emphasis italics="true" pageId="0" pageNumber="171">N. bispina</emphasis>
</taxonomicName>
distributed in
<collectingCountry name="China">China</collectingCountry>
, but
<taxonomicName lsidName="N. occultalis" pageId="0" pageNumber="171" rank="species" species="occultalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. occultalis</emphasis>
</taxonomicName>
has the following differences: subdiscal white spot of forewing narrowed or elongated, tuba analis of male genitalia sclerotized, gnathos of male genitalia elongated and narrow at the base;
<taxonomicName lsidName="N. bispina" pageId="0" pageNumber="171" rank="species" species="bispina">
<emphasis italics="true" pageId="0" pageNumber="171">N. bispina</emphasis>
</taxonomicName>
exhibits the following differences: gnathos of male genitalia absent, phallus of male genitalia with a hook-shaped cornutus, corpus bursae of female genitalia with two thorn-like signa. From
<taxonomicName lsidName="N. hortulatoides" pageId="0" pageNumber="171" rank="species" species="hortulatoides">
<emphasis italics="true" pageId="0" pageNumber="171">N. hortulatoides</emphasis>
</taxonomicName>
, the new species can be easily distinguished by the wing maculation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="description">
<paragraph pageId="0" pageNumber="171">Description</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nagiella adults, dorsal view A N. tristalis sp. nov. male B N. tristalis sp. nov. female C N. inferior male D N. inferior female E N. quadrimaculalis male F N. quadrimaculalis female. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1023.60934.figure2" httpUri="https://binary.pensoft.net/fig/519189" pageId="0" pageNumber="171">2A, B</figureCitation>
).
<emphasis italics="true" pageId="0" pageNumber="171">Head</emphasis>
</emphasis>
: frons brownish grey, smooth. Vertex with erect, dull-orange scales. Labial palpus upturned, dorso-laterally dark brown, ventro-mesally pale white. Antenna dark brown; flagellum filiform with golden cilia ca 1/4 the diameter of flagellum in male.
</paragraph>
<caption doi="10.3897/zookeys.1023.60934.figure2" httpUri="https://binary.pensoft.net/fig/519189" pageId="0" pageNumber="171" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">Figure 2.</emphasis>
<taxonomicName authorityName="Munroe" authorityYear="1976" class="Insecta" family="Crambidae" genus="Nagiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagiella" order="Lepidoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="171">Nagiella</emphasis>
</taxonomicName>
adults, dorsal view
<emphasis bold="true" pageId="0" pageNumber="171">A</emphasis>
<taxonomicName lsidName="N. tristalis" pageId="0" pageNumber="171" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. tristalis</emphasis>
</taxonomicName>
sp. nov. male
<emphasis bold="true" pageId="0" pageNumber="171">B</emphasis>
<taxonomicName lsidName="N. tristalis" pageId="0" pageNumber="171" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. tristalis</emphasis>
</taxonomicName>
sp. nov. female
<emphasis bold="true" pageId="0" pageNumber="171">C</emphasis>
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
male
<emphasis bold="true" pageId="0" pageNumber="171">D</emphasis>
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
female
<emphasis bold="true" pageId="0" pageNumber="171">E</emphasis>
<taxonomicName lsidName="N. quadrimaculalis" pageId="0" pageNumber="171" rank="species" species="quadrimaculalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. quadrimaculalis</emphasis>
</taxonomicName>
male
<emphasis bold="true" pageId="0" pageNumber="171">F</emphasis>
<taxonomicName lsidName="N. quadrimaculalis" pageId="0" pageNumber="171" rank="species" species="quadrimaculalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. quadrimaculalis</emphasis>
</taxonomicName>
female. Scale bars:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Thorax and abdomen</emphasis>
:
</emphasis>
dorsally brownish grey; patagium and tegula with ochreous brown. Ventrally milky white.
</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Wings</emphasis>
:
</emphasis>
length of forewing 12.0-13.0 mm. Ground color of both wings brownish grey, with a large comma-shaped white spot at end of discal cell (the bases of
<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
5 to M3), that of the hindwing somewhat small; cilia concolorous with ground color; postmedial line obscure. Subdiscal white spot of forewing rounded, small and blurry. Base of discal cell of hindwing concolorous with ground color.
</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" italics="true" pageId="0" pageNumber="171">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Nagiella tristalis sp. nov. A whole genitalia B gnathos, magnified view C apex of valva, magnified view." figureDoi="10.3897/zookeys.1023.60934.figure3" httpUri="https://binary.pensoft.net/fig/519190" pageId="0" pageNumber="171">3A-C</figureCitation>
): uncus short, subtrapezoid, blunt on posterior margin. Gnathos nearly triangular, short and small, apex blunt. Tuba analis elongate, not sclerotized, length ca 0.6 times that of valva. Transtilla subtriangular, completely connected medially. Saccus short, anterior margin rounded. Valva somewhat narrow, length ca 3.3 times that of width, dorsal margin slightly incurved subapically; costa more or less inflated with several setae at apex; clasper large, down curved, the apex blunt. Phallus cylindrical, without cornutus.
</paragraph>
<caption doi="10.3897/zookeys.1023.60934.figure3" httpUri="https://binary.pensoft.net/fig/519190" pageId="0" pageNumber="171" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">Figure 3.</emphasis>
Male genitalia of
<taxonomicName authorityName="Matsui &amp; Naka" authorityYear="2021" class="Insecta" family="Crambidae" genus="Nagiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagiella tristalis" order="Lepidoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">Nagiella tristalis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="171">A</emphasis>
whole genitalia
<emphasis bold="true" pageId="0" pageNumber="171">B</emphasis>
gnathos, magnified view
<emphasis bold="true" pageId="0" pageNumber="171">C</emphasis>
apex of valva, magnified view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" italics="true" pageId="0" pageNumber="171">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of Nagiella tristalis sp. nov. A whole genitalia B anterior apophysis, magnified view C signum, magnified view." figureDoi="10.3897/zookeys.1023.60934.figure6" httpUri="https://binary.pensoft.net/fig/519193" pageId="0" pageNumber="171">6A-C</figureCitation>
): ovipositor lobe oblong, setose. Anterior apophysis ca 2 times length of posterior apophysis, narrow, almost straight, with triangular expansion near base. Antrum trapezoid, sclerotized at collar. Ductus bursae almost equal in length to corpus bursae, membranous with wrinkles, slightly narrowed at posterior end. Corpus bursae pear-shaped, membranous; signum nearly elliptical, sparsely covered with denticles.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="biology_ecology">
<paragraph pageId="0" pageNumber="171">Biology.</paragraph>
<paragraph pageId="0" pageNumber="171">
In Honshu,
<collectingCountry name="Japan">Japan</collectingCountry>
, adults are found in May to September, and they are considered bivoltine. They appear to be hardly attracted to light. Larvae feed on
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Rubus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rubus buergeri" order="Rosales" pageId="0" pageNumber="171" phylum="Angiospermae" rank="species" species="buergeri">
<emphasis italics="true" pageId="0" pageNumber="171">Rubus buergeri</emphasis>
</taxonomicName>
and the middle instar larvae overwinter in its leaves.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="distribution">
<paragraph pageId="0" pageNumber="171">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="171">
<materialsCitation ID-GBIF-Occurrence="3063628434" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Ryukyu Islands" specimenCount="1" stateProvince="Tokyo">
<collectingCountry name="Japan">Japan</collectingCountry>
: Honshu (
<collectingRegion country="Japan" name="Tokyo">Tokyo</collectingRegion>
,
<collectingRegion country="Japan" name="Shizuoka">Shizuoka</collectingRegion>
,
<collectingRegion country="Japan" name="Aichi">Aichi</collectingRegion>
,
<collectingRegion country="Japan" name="Tottori">Tottori</collectingRegion>
,
<collectingRegion country="Japan" name="Yamaguchi">Yamaguchi</collectingRegion>
), Kyushu (
<collectingRegion country="Japan" name="Kagoshima">Kagoshima</collectingRegion>
),
<location LSID="urn:lsid:plazi:treatment:C9D9CE9CFFB3503B98FD90FDE60D40F4:257B89266A0408FA7256E4ABAE97B023" country="Japan" name="Ryukyu Islands" stateProvince="Tokyo">Ryukyu Islands</location>
(Yakushima, Amamioshima, Tokunoshima, Okinawajima)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="remarks">
<paragraph pageId="0" pageNumber="171">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="171">
The shapes of the uncus and gnathos show intraspecific variations, i.e., in several specimens, the posterior margin of the uncus is slightly notched medially, and the projection of the gnathos is smaller than that shown on Figure
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Nagiella tristalis sp. nov. A whole genitalia B gnathos, magnified view C apex of valva, magnified view." figureDoi="10.3897/zookeys.1023.60934.figure3" httpUri="https://binary.pensoft.net/fig/519190" pageId="0" pageNumber="171">3B</figureCitation>
. Also, in
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
, the shape of the gnathos varies similarly as in
<taxonomicName lsidName="N. tristalis" pageId="0" pageNumber="171" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. tristalis</emphasis>
</taxonomicName>
. Therefore, these characters are unsuitable for diagnosis of
<taxonomicName lsidName="N. tristalis" pageId="0" pageNumber="171" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. tristalis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
. Similarly,
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.679.11960" author="Ullah, M" journalOrPublisher="ZooKeys" pageId="0" pageNumber="171" pagination="65 - 76" refId="B38" refString="Ullah, M, Yang, Z, Qiao, P, Zhang, Y, 2017. A new cryptic species of Nagiella Munroe from China revealed by DNA barcodes and morphological evidence (Lepidoptera, Crambidae, Spilomelinae). ZooKeys 679: 65 - 76, DOI: https://doi.org/10.3897/zookeys.679.11960" title="A new cryptic species of Nagiella Munroe from China revealed by DNA barcodes and morphological evidence (Lepidoptera, Crambidae, Spilomelinae)." url="https://doi.org/10.3897/zookeys.679.11960" volume="679" year="2017">Ullah et al. (2017)</bibRefCitation>
proposed the shape of the claspers as a diagnostic character for distinguishing between
<taxonomicName lsidName="N. occultalis" pageId="0" pageNumber="171" rank="species" species="occultalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. occultalis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="N. quadrimaculalis" pageId="0" pageNumber="171" rank="species" species="quadrimaculalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. quadrimaculalis</emphasis>
</taxonomicName>
, but we could not find any clear difference among
<taxonomicName lsidName="N. tristalis" pageId="0" pageNumber="171" rank="species" species="tristalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. tristalis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="N. inferior" pageId="0" pageNumber="171" rank="species" species="inferior">
<emphasis italics="true" pageId="0" pageNumber="171">N. inferior</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="N. quadrimaculalis" pageId="0" pageNumber="171" rank="species" species="quadrimaculalis">
<emphasis italics="true" pageId="0" pageNumber="171">N. quadrimaculalis</emphasis>
</taxonomicName>
, because of the overlap in intraspecific variations.
</paragraph>
</subSubSection>
</treatment>
</document>