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<document id="00738371FF3B7CE51BABE8EC30CEDFD7" ENCODING="UTF-8" ID-CLB-Dataset="276867" ID-DOI="https://doi.org/10.5281/zenodo.291971" ID-GBIF-Dataset="0bb2f033-2b79-4d87-803f-91694067c3f3" ID-ISBN="978-0-9506207-7-0" ID-Zenodo-Dep="291971" checkinTime="1588973351969" checkinUser="admin" docAuthor="Jarvis, Charlie" docDate="2007" docId="C9B62DB508BA29063BC6219F8C1DE2D9" docLanguage="en" docName="ch7.gg.C.xml" docOrigin="Order out of Chaos. Linnaean Plant Types and their Types, London: Linnaean Society of London in association with the Natural History Museum" docTitle="Klasea nudicaulis Fourr." docType="treatment" docVersion="11" lastPageNumber="401" masterDocId="E213C9FA410C8D52CE14FE18C476F6F3" masterDocTitle="Chapter 7: Linnaean Plant Names and their Types (part C)" masterLastPageNumber="473" masterPageNumber="370" pageNumber="401" updateTime="1699548976333" updateUser="ExternalLinkService">
<mods:mods id="C10A1B0D19F4F95EC47329AF264B06EA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C4C6FD3B489FC438F7F257DCB84423E9">
<mods:title id="8310493BFF823D205A2A88706B4145A1">Chapter 7: Linnaean Plant Names and their Types (part C)</mods:title>
</mods:titleInfo>
<mods:name id="E86928D67A1E867DA85E81105FE9831F" type="personal">
<mods:role id="DBE620835A684B19DD276C85F9555DB5">
<mods:roleTerm id="155D1C5FAE280DF3AC56C7B0CCBAD5C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B1926837406136687906328BB37CA3A">Jarvis, Charlie</mods:namePart>
<mods:affiliation id="1405ECB09A29EE0AD6C0E305BAA6A998">Department of Botany, Natural History Museum, Cromwell Road, London, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B99A5C441872DCF891C11B89BEAD9318">text</mods:typeOfResource>
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<mods:dateIssued id="3BA3919D65D98F46A3B3BF58D8E383D5">2007</mods:dateIssued>
<mods:publisher id="93318FE74913BC4E5C81F8627C5EBB9A">Linnaean Society of London in association with the Natural History Museum</mods:publisher>
<mods:place id="BA685B8AA8D16E75DE68D13BC6479FC9">
<mods:placeTerm id="764679C73B8DD69F5DCAE53EFE10A241">London</mods:placeTerm>
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</mods:originInfo>
<mods:titleInfo id="6715B0B3A0546390A19CA4EA2FDE3E44">
<mods:title id="ABA2FBCACE652D4066A9F1C19081517F">Order out of Chaos. Linnaean Plant Types and their Types</mods:title>
</mods:titleInfo>
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<mods:classification id="5C81284E9F863E5B1FA1FE6F6EBC6ACC">book chapter</mods:classification>
<mods:identifier id="03452AC3316169F82D9F17F259D41537" type="DOI">https://doi.org/10.5281/zenodo.291971</mods:identifier>
<mods:identifier id="D6A0D5A12F0591D2C2B02050EC171796" type="ISBN">978-0-9506207-7-0</mods:identifier>
<mods:identifier id="3CB232BE53E73D1C840926E902A56D50" type="Zenodo-Dep">291971</mods:identifier>
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<treatment id="C9B62DB508BA29063BC6219F8C1DE2D9" ID-DOI="http://doi.org/10.5281/zenodo.4379002" ID-GBIF-Taxon="164003922" ID-Zenodo-Dep="4379002" LSID="urn:lsid:plazi:treatment:C9B62DB508BA29063BC6219F8C1DE2D9" httpUri="http://treatment.plazi.org/id/C9B62DB508BA29063BC6219F8C1DE2D9" lastPageNumber="401" pageNumber="401">
<subSubSection id="CD0398C122DAEC191F3EE677F498E530" pageNumber="401" type="reference_group">
<paragraph id="9BDEAE05A3EA541536AF245FAC63DA90" pageNumber="401">
<treatmentCitation id="12E9BC1C873A5E3DC77BED8EB731B37B" author="Linnaeus" pageNumber="1232" year="1759">
<bibCitation id="A412F1DA5584BFCBC5F1E4F0FE4F6844" author="Linnaeus" pagination="1232" title="Centaurea nudicaulis" volumeTitle="Systema Naturae, ed. 10, 2" year="1759">
<taxonomicName id="D05F1027125F8B7046EABD783C935CD1" ID-CoL="S742" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1759" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="nudicaulis">
<emphasis id="A9E89A8C62976E283873C08FBE951378" bold="true" pageNumber="401">Centaurea nudicaulis</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="A6ADD843687979EAA0A5EFD7148DF364">
<emphasis id="7D84A2C9FC257EF185F0E4DF906601DD" italics="true" pageNumber="401">Systema Naturae</emphasis>
, ed. 10, 2
</volumeTitle>
: 1232. 1759
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="152A6CA367C2C49E44CF80F0E44FC438" pageNumber="401" type="distribution">
<paragraph id="9F8E3A75A1DEB275EDED22EBB3F6625A" pageNumber="401">[&quot;Habitat in Italia, rupe victoriae Galloprovinciae.&quot;] Sp. Pl. ed. 2, 2: 1300 (1763). RCN: 6638.</paragraph>
</subSubSection>
<subSubSection id="6A89911661AB17C14323F8787556F5BF" pageNumber="401" type="materials_examined">
<paragraph id="B8EE3A2311670D9DED3473E6C9ACB448" pageNumber="401">Type not designated.</paragraph>
</subSubSection>
<subSubSection id="43D2B3241013D81F1B6DE49E0727F9B2" pageNumber="401" type="materials_examined">
<paragraph id="87E4E14A99BA20C95ED44D528868F92D" pageNumber="401">
Original material:
<materialsCitation id="C007168E6C2400035D2009EC3C6711D1" ID-GBIF-Occurrence="2612072865" collectionCode="LINN" specimenCode="1030.72">
Herb. Linn. No. 1030.72 (
<collectionCode id="DFCFD3FBF24000206F3C0398045E2A52">LINN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3CD50E09166115BA2EFE5962F3CB5D6" pageNumber="401" type="nomenclature">
<paragraph id="78D45EA1CE6BDF549674FFC712353E0E" pageNumber="401">
Current name:
<taxonomicName id="456D52DD3A05051E5A87A8BF018F52A4" ID-CoL="6NKCH" authority="(L.) Fourr." authorityName="Fourr." baseAuthorityName="Linnaeus" baseAuthorityYear="1759" class="Magnoliopsida" family="Asteraceae" genus="Klasea" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="nudicaulis">
<emphasis id="F32868987CA149CF0E847F68DEFEEAEB" italics="true" pageNumber="401">
<emphasis id="DB918B3B95BA647EEF2AA767158C9231" bold="true" italics="true" pageNumber="401">Klasea nudicaulis</emphasis>
</emphasis>
(L.) Fourr.
</taxonomicName>
(
<taxonomicName id="7AB92A6963FCAC3B0CE6EEDB193C18DC" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="586E7EE277D3CBFCEC42CAF3E2992EA0" pageNumber="401" type="discussion">
<paragraph id="5037E0D223D008AB061A56BE09FB7F7C" pageNumber="401">
<emphasis id="33FC926CBC4D76DAFD52BBF103BF859B" italics="true" pageNumber="401">Note:</emphasis>
<normalizedToken id="C81666547EEFE6FA805E0F2582C7E750" originalValue="Cantó">Canto</normalizedToken>
(in
<emphasis id="5482E04F741483F619DB011B29369165" italics="true" pageNumber="401">Lazaroa</emphasis>
6: 23. 1984) designated a Barrelier figure, cited by Linnaeus in
<volumeTitle id="2CED0FDCB937159A0225DB699E53F04B">
<emphasis id="C901D272396B71EE891E20E5913FCA8D" italics="true" pageNumber="401">Species Plantarum</emphasis>
, ed. 2, 2
</volumeTitle>
: 1300 (1763) but not in the protologue in 1759, as type. It is consequently not an original element for the name and, as original material appears to exist (1030.72 LINN),
<normalizedToken id="D11431BF33E35C427604619D2C57DAE0" originalValue="Cantó’s">Canto's</normalizedToken>
choice is not effective.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.869.34380" ID-GBIF-Dataset="0733a684-7888-4526-9449-dee89fe914f3" ID-PMC="PMC6690876" ID-Pensoft-Pub="1313-2970-869-1" ID-Pensoft-UUID="5E1D7C137E42575296C2A10148C7891D" ID-PubMed="31413658" ID-ZooBank="50D49A54A3864738AE1F393B77EC652C" ModsDocID="1313-2970-869-1" checkinTime="1565056329852" checkinUser="pensoft" docAuthor="Jeong, Man-Ki, Soh, Ho Young &amp; Suh, Hae-Lip" docDate="2019" docId="C9B63C59DB6A57749F75A5A34E5E384C" docLanguage="en" docName="ZooKeys 869: 1-18" docOrigin="ZooKeys 869" docSource="http://dx.doi.org/10.3897/zookeys.869.34380" docTitle="Heteromastus namhaensis Jeong, Soh &amp; Suh, 2019, sp. nov." docType="treatment" docUuid="D41E7B49-B712-42A7-8095-94E906ABB121" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="677FFFA5FB24FFE1FF94FF87FFFF7104" masterDocTitle="Three new species of Heteromastus (Annelida, Capitellidae) from Korean waters, with genetic evidence based on two gene markers" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1668167666701" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new species of Heteromastus (Annelida, Capitellidae) from Korean waters, with genetic evidence based on two gene markers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jeong, Man-Ki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soh, Ho Young</mods:namePart>
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<mods:name type="personal">
<mods:role>
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<mods:namePart>Suh, Hae-Lip</mods:namePart>
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<mods:date>2019</mods:date>
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<mods:number>869</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.869.34380</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.869.34380</mods:identifier>
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<treatment ID-GBIF-Taxon="158995535" LSID="urn:lsid:zoobank.org:act:D41E7B49-B712-42A7-8095-94E906ABB121" httpUri="http://treatment.plazi.org/id/C9B63C59DB6A57749F75A5A34E5E384C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName LSID="c9b63c59-db6a-5774-9f75-a5a34e5e384c" class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus namhaensis" order="Annelida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="namhaensis">Heteromastus namhaensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">
Figures 2
<normalizedToken originalValue="A–G">A-G</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Heteromastus namhaensis sp. nov. A abdominal hooded hook in lateral view B posterior end in dorsal view (holotype, NA 00155558). Heteromastus gusipoensis sp. nov. C abdominal hooded hook in frontal view D posterior end in dorsal view (holotype, NA 00155561). Heteromastus koreanus sp. nov. E abdominal hooded hook in frontal view F posterior end in dorsal view (holotype, NA 00155565). Abbreviations: ac, anal cirrus; mf, main fang; neu, neuropod; no, notopod; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure5" httpUri="https://binary.pensoft.net/fig/322703" pageId="0" pageNumber="1">5A, B</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Methylene green staining patterns of Korean three new species A anterior end of H. namhaensis sp. nov., lateral view (paratype, NA 00155560) B anterior end of H. gusipoensis sp. nov., lateral view (using additional specimens from type locality) C anterior end of H. koreanus sp. nov., lateral view (NA 00065689)." figureDoi="10.3897/zookeys.869.34380.figure6" httpUri="https://binary.pensoft.net/fig/322704" pageId="0" pageNumber="1">6A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
: MABIKNA00155558, sex uncertain, Cheongsando,
<geoCoordinate degrees="34" direction="north" minutes="1.662" orientation="latitude" precision="1" value="34.0277">34°1.662'N</geoCoordinate>
,
<geoCoordinate degrees="127" direction="east" minutes="4.272" orientation="longitude" precision="1" value="127.0712">127°4.272'E</geoCoordinate>
, subtidal, sandy mud bottom, 34 m depth, March 2016, coll. Man-Ki Jeong.
<emphasis bold="true" pageId="0" pageNumber="1">Paratypes</emphasis>
(two specimens): MABIKNA00155560, Yeosu,
<geoCoordinate degrees="34" direction="north" minutes="41.569" orientation="latitude" precision="1" value="34.692818">34°41.569'N</geoCoordinate>
,
<geoCoordinate degrees="127" direction="east" minutes="51.848" orientation="longitude" precision="1" value="127.864136">127°51.848'E</geoCoordinate>
, subtidal, sandy mud bottom, 15 m depth, June 2018; MABIK NA00155559, Jejudo,
<geoCoordinate degrees="33" direction="north" minutes="16.699" orientation="latitude" precision="1" value="33.278316">33°16.699'N</geoCoordinate>
,
<geoCoordinate degrees="127" direction="east" minutes="16.230" orientation="longitude" precision="1" value="127.2705">127°16.230'E</geoCoordinate>
, subtidal, sandy mud bottom, 54 m depth, April 2018. Additional 6 specimens from type locality on SEM stub.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Abdominal hooks with four rows of teeth, three teeth in basal row, three in second and third row, and four to six in superior row. Genital pores present in intersegmental furrows between chaetigers 7-8, 8-9, 9-10, and 10-11. Hemispheric notopodial lobes present on posterior abdominal segments.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype entire, about 60 mm long, 0.9 mm wide for 98 chaetigers (terminal part missing). Paratypes range from 19-41 mm in length, 0.5-0.8 mm width for 41-95 chaetigers. Body thread-like, rounded dorsally, flattened ventrally, widest in anterior thoracic chaetigers, and tapering from abdomen to pygidium. Color brownish yellow in alcohol.</paragraph>
<paragraph pageId="0" pageNumber="1">
Prostomium conical, with short and hemispherical palpode; nuchal organs not seen, eyespots absent (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
). Everted proboscis with numerous small papillae (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A</figureCitation>
). Peristomium uni-annulated and slightly longer than first thoracic chaetiger (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Thorax with 11 chaetigers (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
). Thoracic segments biannulated, with shallow intra- and intersegmental grooves (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
). Anterior five thoracic segments slightly expanded (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
). First chaetiger biramous, with three or four bi-limbated capillaries; chaetigers 2-5 with six to 14 capillaries per fascicle in both parapodia; chaetigers 6-11 with five to 12 long-shafted hooded hooks per fascicle (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B, F</figureCitation>
); thoracic hooks with indistinct node on shaft and at least six teeth in three rows above the main fang (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2F</figureCitation>
). Notopodia located dorso-laterally, dorsally located in last few thoracic segments; neuropodia located in lateral positions (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
). Lateral organs present between noto- and neuropodia of all thoracic chaetigers, nearer to notopodia in chaetigers 5-11; sometimes indistinct on first thoracic chaetigers (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A</figureCitation>
). Genital pores present in intersegmental furrows of between chaetigers 7-8, 8-9, 9-10, and 10-11 (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Transition between thorax and abdomen distinguished by changes in ultrastructure of chaetae and shape of segment (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
); abdominal segments multi-annulated, gradually longer posteriorly, with short-shafted hooded hooks in posterior parapodial lobes; thoracic chaetigers usually bi-annulated, wider than long, with long-shafted hooded hooks in center of segment (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Abdominal parapodial lobes located in posterior end of each segment, well separated from each other, and gradually developed posteriorly (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">
Fig. 2
<normalizedToken originalValue="A–D">A-D</normalizedToken>
</figureCitation>
). Abdominal notopodia separated, mid-dorsal on anterior few segments, becoming dorsolateral in following abdominal region, with six to eight hooded hooks per fascicle, having dorso-posteriorly protruded and hemispheric lobes from chaetiger 90 to end of body (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">
Figs 2
<normalizedToken originalValue="A–D">A-D</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Heteromastus namhaensis sp. nov. A abdominal hooded hook in lateral view B posterior end in dorsal view (holotype, NA 00155558). Heteromastus gusipoensis sp. nov. C abdominal hooded hook in frontal view D posterior end in dorsal view (holotype, NA 00155561). Heteromastus koreanus sp. nov. E abdominal hooded hook in frontal view F posterior end in dorsal view (holotype, NA 00155565). Abbreviations: ac, anal cirrus; mf, main fang; neu, neuropod; no, notopod; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure5" httpUri="https://binary.pensoft.net/fig/322703" pageId="0" pageNumber="1">5B</figureCitation>
). Abdominal neuropodia well separated, with 10-12 hooded hooks per fascicle, having slightly protruded lobes in posterior abdomen; neuropodial lobes less developed than notopodial lobes (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Figs 2C, D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Heteromastus namhaensis sp. nov. A abdominal hooded hook in lateral view B posterior end in dorsal view (holotype, NA 00155558). Heteromastus gusipoensis sp. nov. C abdominal hooded hook in frontal view D posterior end in dorsal view (holotype, NA 00155561). Heteromastus koreanus sp. nov. E abdominal hooded hook in frontal view F posterior end in dorsal view (holotype, NA 00155565). Abbreviations: ac, anal cirrus; mf, main fang; neu, neuropod; no, notopod; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure5" httpUri="https://binary.pensoft.net/fig/322703" pageId="0" pageNumber="1">5B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Hooded hooks with main fang extending slightly beyond hoods. Abdominal hooks with distinct node on shaft and four rows of small teeth above main fang; three teeth in basal row, three in second and third row, and four to six in superior row (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Figs 2E, G</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Heteromastus namhaensis sp. nov. A abdominal hooded hook in lateral view B posterior end in dorsal view (holotype, NA 00155558). Heteromastus gusipoensis sp. nov. C abdominal hooded hook in frontal view D posterior end in dorsal view (holotype, NA 00155561). Heteromastus koreanus sp. nov. E abdominal hooded hook in frontal view F posterior end in dorsal view (holotype, NA 00155565). Abbreviations: ac, anal cirrus; mf, main fang; neu, neuropod; no, notopod; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure5" httpUri="https://binary.pensoft.net/fig/322703" pageId="0" pageNumber="1">5A</figureCitation>
). Pygidium with digitate anal cirrus (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heteromastus namhaensis sp. nov. A anterior end, left lateral view (holotype, NA 00155558) B same, dorsal view C posterior abdominal segments, right lateral view (holotype, NA 00155558) D posterior end, dorsal view (holotype, NA 00155558) E abdominal short-shafted hook, frontal view. F thoracic long-shafted hook, lateral view G abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1">Figs 2D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Heteromastus namhaensis sp. nov. A abdominal hooded hook in lateral view B posterior end in dorsal view (holotype, NA 00155558). Heteromastus gusipoensis sp. nov. C abdominal hooded hook in frontal view D posterior end in dorsal view (holotype, NA 00155561). Heteromastus koreanus sp. nov. E abdominal hooded hook in frontal view F posterior end in dorsal view (holotype, NA 00155565). Abbreviations: ac, anal cirrus; mf, main fang; neu, neuropod; no, notopod; pyg, pygidium." figureDoi="10.3897/zookeys.869.34380.figure5" httpUri="https://binary.pensoft.net/fig/322703" pageId="0" pageNumber="1">5B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.869.34380.figure2" httpUri="https://binary.pensoft.net/fig/322700" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
Figure 2.
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus namhaensis" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus namhaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior end, left lateral view (holotype, NA00155558)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
same, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior abdominal segments, right lateral view (holotype, NA00155558)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
posterior end, dorsal view (holotype, NA00155558)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal short-shafted hook, frontal view.
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
thoracic long-shafted hook, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
abdominal short-shafted hook, lateral view. Abbreviations: ac, anal cirrus; cc, capillary chaetae; Ch, chaetiger; gp, genital pore; hh, hooded hooks; lo, lateral organ; mf, main fang; neu, neuropod; no, notopod; per, peristomium; pro, prostomium; prob, proboscis; pyg, pygidium.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="methyl green staining pattern">
<paragraph pageId="0" pageNumber="1">Methyl green staining pattern.</paragraph>
<paragraph pageId="0" pageNumber="1">
Prostomium, peristomium and thoracic chaetigers 1-2 not stained (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Methylene green staining patterns of Korean three new species A anterior end of H. namhaensis sp. nov., lateral view (paratype, NA 00155560) B anterior end of H. gusipoensis sp. nov., lateral view (using additional specimens from type locality) C anterior end of H. koreanus sp. nov., lateral view (NA 00065689)." figureDoi="10.3897/zookeys.869.34380.figure6" httpUri="https://binary.pensoft.net/fig/322704" pageId="0" pageNumber="1">Fig. 6A</figureCitation>
). Thoracic chaetigers 3-11 stained blue; chaetigers 3-8 stained dark blue; chaetigers 9-11 stained light blue; post-chaetal region of chaetiger 11 not stained (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Methylene green staining patterns of Korean three new species A anterior end of H. namhaensis sp. nov., lateral view (paratype, NA 00155560) B anterior end of H. gusipoensis sp. nov., lateral view (using additional specimens from type locality) C anterior end of H. koreanus sp. nov., lateral view (NA 00065689)." figureDoi="10.3897/zookeys.869.34380.figure6" httpUri="https://binary.pensoft.net/fig/322704" pageId="0" pageNumber="1">Fig. 6A</figureCitation>
). Abdominal region without any distinct staining pattern.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species is named for its wide distribution in Namhae (=Korean name of southern sea of Korea).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Subtidal areas (15-54 m) near southern part of Korea (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of study area with main collecting locations indicated. Abbreviations (district names): AH, Anheung; CS, Cheongsando; GY, Gwangyang; JJ, Jejudo; MA, Muan; SC, Seochun; YG, Yeonggwang; YS, Yeosu." figureDoi="10.3897/zookeys.869.34380.figure1" httpUri="https://binary.pensoft.net/fig/322699" pageId="0" pageNumber="1">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus namhaensis" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus namhaensis</emphasis>
</taxonomicName>
was sampled from soft sediments in March of 2016 (10 ind./m2), April of 2018 (40 ind./m2), and June of 2018 (20 ind./m2). The most well-developed individual (having over 100 segments) was obtained in March and eggs in the coelom were 87-94
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter. Surface sediment of the station was mainly composed of sandy mud with fragmented shells.
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Leiochrides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiochrides yokjidoensis" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="yokjidoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Leiochrides yokjidoensis</emphasis>
</taxonomicName>
Jeong, Wi &amp; Suh, 2017 co-occurred in Jejudo of Korea (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.685.12700" author="Jeong, MK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="91 - 103" refId="B19" refString="Jeong, MK, Wi, JH, Suh, HL, 2017a. A new species of Leiochrides from the Korean subtidal waters with notes on the taxonomic status of the genus Pseudomastus (Annelida, Capitellidae). . ZooKeys 685: 91 - 103" title="A new species of Leiochrides from the Korean subtidal waters with notes on the taxonomic status of the genus Pseudomastus (Annelida, Capitellidae)." url="https://doi.org/10.3897/zookeys.685.12700" volume="685" year="2017 a">Jeong et al. 2017a</bibRefCitation>
;
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of study area with main collecting locations indicated. Abbreviations (district names): AH, Anheung; CS, Cheongsando; GY, Gwangyang; JJ, Jejudo; MA, Muan; SC, Seochun; YG, Yeonggwang; YS, Yeosu." figureDoi="10.3897/zookeys.869.34380.figure1" httpUri="https://binary.pensoft.net/fig/322699" pageId="0" pageNumber="1">Fig. 1</figureCitation>
). The salinity range among sampling locations was about 31-32.5 PSU.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus namhaensis" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus namhaensis</emphasis>
</taxonomicName>
resembles
<taxonomicName lsidName="H. filiformis" pageId="0" pageNumber="1" rank="species" species="filiformis">
<emphasis italics="true" pageId="0" pageNumber="1">H. filiformis</emphasis>
</taxonomicName>
sensu Hutchings &amp; Rainer, 1982 in the absence of distinct eyespots on prostomium, three teeth in basal row above the main fang of abdominal hooks, and the presence of posteriorly extended abdominal notopodial lobes (Table 2). However, they differ in the shape of notopodial lobes in posterior abdomen (hemispheric protrusion in
<taxonomicName lsidName="H. namhaensis" pageId="0" pageNumber="1" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. namhaensis</emphasis>
</taxonomicName>
vs broadly-based and rounded lamellae in
<taxonomicName lsidName="H. filiformis" pageId="0" pageNumber="1" rank="species" species="filiformis">
<emphasis italics="true" pageId="0" pageNumber="1">H. filiformis</emphasis>
</taxonomicName>
sensu Hutchings &amp; Rainer, 1982), the different dental structure of abdominal hooks (Table 2).
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus namhaensis" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus namhaensis</emphasis>
</taxonomicName>
is also easily distinguished from Korean former record of
<taxonomicName lsidName="H. filiformis" pageId="0" pageNumber="1" rank="species" species="filiformis">
<emphasis italics="true" pageId="0" pageNumber="1">H. filiformis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.12651/JSR.2016.5.3.264" author="Choi, HK" journalOrPublisher="Journal of Species Research" pageId="0" pageNumber="1" pagination="264 - 271" refId="B5" refString="Choi, HK, Yoon, SM, 2016. First record of Heteromastusfiliformis (Polychaeta: Capitellidae) from Korean waters, with a taxonomic note on the genus. . Journal of Species Research 5: 264 - 271" title="First record of Heteromastusfiliformis (Polychaeta: Capitellidae) from Korean waters, with a taxonomic note on the genus." url="https://doi.org/10.12651/JSR.2016.5.3.264" volume="5" year="2016">Choi and Yoon 2016</bibRefCitation>
) by the presence of hemispheric abdominal parapodial lobes and the absence of eyespots in
<taxonomicName lsidName="H. namhaensis" pageId="0" pageNumber="1" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. namhaensis</emphasis>
</taxonomicName>
. In particular, the hemispheric notopodial lobe of
<taxonomicName lsidName="H. namhaensis" pageId="0" pageNumber="1" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. namhaensis</emphasis>
</taxonomicName>
is a unique feature in the genus.
</paragraph>
<caption ID-Table-UUID="7BAA7B4AB18991098668BA5D6AB39083" httpUri="http://table.plazi.org/id/7BAA7B4AB18991098668BA5D6AB39083" pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Table 2. Morphological comparison between
<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus</emphasis>
</taxonomicName>
species of this study and their closely related species. A: absent; P: present; Ch: chaetiger.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Species</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Eyespots</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Dental structure of abdominal hooks</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Notopodial lobes in posterior abdomen</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Methyl green staining pattern</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Habitat (locality)</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. namhaensis" pageId="0" pageNumber="1" rank="species" species="namhaensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. namhaensis</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">A</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4 rows (3/3/3/4-6)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Hemispheric notopodial lobes dorso-posteriorly extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Ch 3-11 blue, abdomen not stained</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Subtidal, 36 m, sandy mud with shell fragments (Korea)</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. gusipoensis" pageId="0" pageNumber="1" rank="species" species="gusipoensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. gusipoensis</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">A</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4 rows (3/3/4/2)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Not extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Ch 3-10 with blue speckles, median part of each segment stained densely</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Intertidal, 0-1 m, sandy mud (Korea)</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="1" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="1">H. koreanus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3 rows (2/3/4)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Rounded notopodial lobes posteriorly extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Ch 6-11 green, Ch 11 dark, abdomen not stained</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Intertidal, estuarine, 0-1 m, sandy mud (Korea)</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. filiformis" pageId="0" pageNumber="1" rank="species" species="filiformis">
<emphasis italics="true" pageId="0" pageNumber="1">H. filiformis</emphasis>
</taxonomicName>
sensu Hutchings &amp; Rainer, 1982
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">A</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
3 rows (3
<normalizedToken originalValue="–4/4–5/4–">-4/4-5/4-</normalizedToken>
6)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Broadly-based and rounded notopodial lobes posteriorly extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Intertidal (Mediterranean)</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. filiformis" pageId="0" pageNumber="1" rank="species" species="filiformis">
<emphasis italics="true" pageId="0" pageNumber="1">H. filiformis</emphasis>
</taxonomicName>
sensu Choi &amp; Yoon, 2016
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3-4 teeth in 3 rows</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Rounded notopodial lobes posteriorly extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Ch 1 &amp; Ch 3-11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Intertidal (Korea)</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="H. tohbaiensis" pageId="0" pageNumber="1" rank="species" species="tohbaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. tohbaiensis</emphasis>
</taxonomicName>
Yabe &amp; Mawatari, 1998
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Variable (&gt;11)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Not extended</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Lake, low salinity, fine mud (Japan)</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

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<bibRefCitation pageId="0" pageNumber="10084">Economou-Amilli et al. 1984</bibRefCitation>
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