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<document id="4B3D54244F4A8084AB56A44EAB4EDB2F" ID-CLB-Dataset="300345" ID-DOI="10.1186/s13358-022-00257-z" ID-GBIF-Dataset="21b570bc-eafa-4448-b23b-61a89ddddd63" ID-ISSN="1664-2384" ID-PMC="PMC9499918" ID-PubMed="36164559" ID-Zenodo-Dep="11998918" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718652706118" checkinUser="felipe" docAuthor="Ferrante, Christophe, Menkveld-Gfeller, Ursula &amp; Cavin, Lionel" docDate="2022" docId="03E487A4F34BFF86139F8B66BDCDFBFD" docLanguage="en" docName="SwissJPalaeontol.141.15.pdf" docOrigin="Swiss Journal of Palaeontology (15) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00257-z" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Libys callolepis Ferrante &amp; Menkveld-Gfeller &amp; Cavin 2022, sp. nov." docType="treatment" docUuid="876BF37D-30B0-4DBD-BEFA-FC66C579129A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="6" masterDocId="FFDDFFDCF34FFF83132A8D4DBF0DFFFE" masterDocTitle="The first Jurassic coelacanth from Switzerland" masterLastPageNumber="20" masterPageNumber="1" pageNumber="5" updateTime="1722275673157" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="23EE8E50BA3F288B37DCC08829F3DEEA">The first Jurassic coelacanth from Switzerland</mods:title>
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<mods:namePart id="4595A22BC4D2818341EB52D08219E574">Ferrante, Christophe</mods:namePart>
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<mods:namePart id="B338F74E12517065F1E7EED7DCF7726E">Menkveld-Gfeller, Ursula</mods:namePart>
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<mods:namePart id="D409E23DF4E091DC2B1A71C4635BCE54">Cavin, Lionel</mods:namePart>
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<treatment id="03E487A4F34BFF86139F8B66BDCDFBFD" ID-DOI="http://doi.org/10.5281/zenodo.13126254" ID-Zenodo-Dep="13126254" LSID="urn:lsid:zoobank.org:act:876BF37D-30B0-4DBD-BEFA-FC66C579129A" httpUri="http://treatment.plazi.org/id/03E487A4F34BFF86139F8B66BDCDFBFD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C3576539F34BFF87139F8B66BE91F9BD" box="[181,412,1579,1603]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BF236B2F34BFF87139F8B66BE91F9BD" blockId="4.[181,412,1579,1603]" box="[181,412,1579,1603]" pageId="4" pageNumber="5">
<heading id="D0BA81DEF34BFF87139F8B66BE91F9BD" box="[181,412,1579,1603]" fontSize="10" level="2" pageId="4" pageNumber="5" reason="7">
<taxonomicName id="4C4D4D31F34BFF87139F8B66BE32F9BD" authority="Ferrante &amp; Menkveld-Gfeller &amp; Cavin, 2022" authorityName="Ferrante &amp; Menkveld-Gfeller &amp; Cavin" authorityYear="2022" box="[181,319,1579,1603]" class="Coelacanthi" family="Mawsoniidae" genus="Libys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coelacanthiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="callolepis" status="sp. nov.">L. callolepis</taxonomicName>
<taxonomicNameLabel id="A20A57DBF34BFF87126F8B66BE91F9BD" box="[325,412,1579,1603]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B939EAA0F34BFF87126F8B66BE98F9BD" bold="true" box="[325,405,1579,1603]" pageId="4" pageNumber="5">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8713BD8B26BCBCFEDD" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BF236B2F34BFF8713BD8B26BE07F97D" blockId="4.[151,775,1643,1891]" box="[151,266,1643,1667]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8B26BE07F97D" bold="true" box="[151,266,1643,1667]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD8BC6BCBCFEDD" blockId="4.[151,775,1643,1891]" lastBlockId="4.[813,1437,235,291]" pageId="4" pageNumber="5">
<taxonomicName id="4C4D4D31F34BFF8713BD8BC6BE2BF95D" box="[151,294,1675,1699]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8BC6BFC2F95D" box="[151,207,1675,1699]" italics="true" pageId="4" pageNumber="5">Libys</emphasis>
species
</taxonomicName>
with the postparietal shield about half the length of the parietonasal shield (the parietonasal is then proportionally shorter than in the
<typeStatus id="54F68810F34BFF8711208B81BD34F91A" box="[522,569,1740,1764]" pageId="4" pageNumber="5">type</typeStatus>
species). Te teeth covering the prearticular are very small, and rounded and smooth. Between 4147 neural arches. Fin rays are slender than in the
<typeStatus id="54F68810F34BFF87126E8A61BE7EF8BA" box="[324,371,1836,1860]" pageId="4" pageNumber="5">type</typeStatus>
species and then not expanded. Te scales are strongly ornamented with irregularly sized and elongated round-to-ovoid ridges disposed along a longitudinal axis.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078C06BBE6FDFD" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF236B2F34BFF8710078C06BB63FE9D" blockId="4.[813,1436,331,419]" box="[813,1134,331,355]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078C06BB63FE9D" bold="true" box="[813,1134,331,355]" pageId="4" pageNumber="5">Measurements and meristic</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078C26BB0FFE5D" blockId="4.[813,1436,331,419]" pageId="4" pageNumber="5">
(TB) Total body length
<quantity id="4CB59B57F34BFF87171A8C26BB86FE7D" box="[1072,1163,363,387]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="mm" value="290.0">290 mm</quantity>
(estimation); (SL) standard length
<quantity id="4CB59B57F34BFF8710888CC6BCF3FE5D" box="[930,1022,395,419]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="5" unit="mm" value="255.0">255 mm</quantity>
.
</paragraph>
<paragraph id="8BF236B2F34BFF8710078C86BBE6FDFD" blockId="4.[813,1437,458,515]" pageId="4" pageNumber="5">d1.f= 10; d2.f =16; pect.f =1822; pelv.f ≥17; ana.f= 2023; cau.f= 15/1416; n.a =4447; h.a= 1820.</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078F66BB90FD3D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BF236B2F34BFF8710078F66BCA4FDBD" blockId="4.[813,1436,555,708]" box="[813,937,555,579]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078F66BCA4FDBD" bold="true" box="[813,937,555,579]" pageId="4" pageNumber="5">Etymology</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078F01BB90FD3D" blockId="4.[813,1436,555,708]" pageId="4" pageNumber="5">
From the ancient Greek ΚΑλός, kalós, (beautiful, nice) and λεπίς, lepís, (scale) in reference to the nicely ornamented scales of the species, which differentiates it from the
<typeStatus id="54F68810F34BFF8717278FE1BB33FD3A" box="[1037,1086,684,708]" pageId="4" pageNumber="5">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078FA6BC87FA3D" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF236B2F34BFF8710078FA6BBC6FCFD" blockId="4.[813,1437,747,995]" box="[813,1227,747,771]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078FA6BBC6FCFD" bold="true" box="[813,1227,747,771]" pageId="4" pageNumber="5">
<typeStatus id="54F68810F34BFF8710078FA6BC94FCFD" box="[813,921,747,771]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
and only known specimen
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078E41BB74FC1D" blockId="4.[813,1437,747,995]" pageId="4" pageNumber="5">
<materialsCitation id="3B253CEFF34BFF8710078E41BB74FC1D" collectionCode="NMBE" country="Switzerland" county="Natural History Museum" location="Most" municipality="Te" pageId="4" pageNumber="5" specimenCode="NMBE 5034072" specimenCount="1" stateProvince="Bern" typeStatus="holotype">
<specimenCode id="DBEB9EC9F34BFF8710078E41BCD0FCDD" box="[813,989,779,803]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE 5034072</specimenCode>
and 5034073, a sub-complete specimen preserved in right lateral view as part and counterpart. Most of the bones, including the scales on the body, are preserved in anatomical position and only the bones of the cheek and the jaw are missing. Te specimen is kept in the collections of the Natural History Museum Bern (Canton of Bern, Switzerland).
</materialsCitation>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078946BB5EFBDD" blockId="4.[813,1437,1035,1155]" box="[813,1107,1035,1059]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078946BB5EFBDD" bold="true" box="[813,1107,1035,1059]" pageId="4" pageNumber="5">
Horizon and
<typeStatus id="54F68810F34BFF8710EE8946BCFAFBDD" box="[964,1015,1035,1059]" pageId="4" pageNumber="5">type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078966BB25FB7D" blockId="4.[813,1437,1035,1155]" pageId="4" pageNumber="5">
Toarcian (Lower Jurassic), Creux de lOurs section, locality of Les Pueys near the Teysachaux summit (
<collectingRegion id="4989F850F34BFF8716078906BC82FB7D" country="Switzerland" name="Fribourg" pageId="4" pageNumber="5">Canton of Fribourg</collectingRegion>
,
<collectingCountry id="F35A7622F34BFF8710B28926BB11FB7D" box="[920,1052,1131,1155]" name="Switzerland" pageId="4" pageNumber="5">Switzerland</collectingCountry>
).
</paragraph>
<paragraph id="8BF236B2F34BFF87100789E6BB09FB3D" blockId="4.[813,1437,1195,1475]" box="[813,1028,1195,1219]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF87100789E6BB09FB3D" bold="true" box="[813,1028,1195,1219]" pageId="4" pageNumber="5">Nomenclatural act</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078981BC87FA3D" blockId="4.[813,1437,1195,1475]" pageId="4" pageNumber="5">
Te present work and its nomenclatural act are registered in ZooBank, the online registration system for the International Commission on Zoological Nomenclature. Te Life Science Identifiers for this publication is “
<uri id="FFDC3AB0F34BFF8710728806BB27FA7D" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:pub:
<uuid id="FFEB0C67F34BFF8717558806BB27FA7D" pageId="4" pageNumber="5">D03A2AC9-51F9-45E6- 8CAC-F06A1526AF2C</uuid>
</uri>
” and can be resolved appending the prefix “http://zoobank.org/” in any standard web browser.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF86100788A7BDCDFBFD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF236B2F34BFF87100788A7BCB6F9FD" blockId="4.[813,1437,1514,1891]" box="[813,955,1514,1539]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF87100788A7BCB6F9FD" bold="true" box="[813,955,1514,1539]" pageId="4" pageNumber="5">Description</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078B40BCA1F9DA" blockId="4.[813,1437,1514,1891]" box="[813,940,1549,1572]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078B40BCA1F9DA" bold="true" box="[813,940,1549,1572]" pageId="4" pageNumber="5">Generalities</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078B61BB84F89D" blockId="4.[813,1437,1514,1891]" pageId="4" pageNumber="5">
Most of the bones of the specimen are preserved on the part (
<collectionCode id="ED5CAE77F34BFF87104C8B06BCBCF99D" box="[870,945,1611,1635]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE</collectionCode>
5034073) (
<figureCitation id="13762A37F34BFF8717028B06BB6EF99D" box="[1064,1123,1611,1635]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
). From the head, most of the dermal bones of the skull roof are preserved. Te bones of the cheek and the lower jaw are missing, revealing then the bones of the palatoquadrate and of the branchial apparatus. Te axial skeleton, the girdles and the fins are almost entirely preserved, except the posterior tip of the supplementary caudal fin lobe. Te scales are well preserved on the entire body in natural position, especially on the counterpart (
<collectionCode id="ED5CAE77F34BFF8710A88A06BCC0F89D" box="[898,973,1867,1891]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE</collectionCode>
5034072) (
<figureCitation id="13762A37F34BFF87176B8A06BB76F89D" box="[1089,1147,1867,1891]" captionStart="Fig" captionStartId="7.[168,199,1783,1803]" captionTargetBox="[152,1438,237,1949]" captionTargetId="figure-14@8.[198,1392,253,1911]" captionTargetPageId="8" captionText="Fig. 4 Skeleton of Libys callolepis sp. nov. on the counterpart (holotype, NMBE 5034072).A Photos with osteological details: 1, articular head of the scapulocoracoid. 2, Scales on the flank immediately beneath the first anterior dorsal fin. 3, Scales of the lateral line showing the ornamental pattern with the larger central tubercles (white arrowheads point, showed only on one scale).4, Scales on the ventral flank from the pelvic to the anal fin. 5, Axial mesomere (white arrowhead) surrounded by some fin rays of the anal fin. 6, Axial mesomeres (white arrowhead) partially covered by sediment in the pelvic fin. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126233" httpUri="https://zenodo.org/record/13126233/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BF236B2F34AFF8613878DA1BE0EFDBD" blockId="5.[151,776,235,579]" pageId="5" pageNumber="6">
Te specimen may represent an adult individual as all the basal plates are fully ossified, which is a feature observed in adult coelacanths (e.g.,
<bibRefCitation id="EFDC4B43F34AFF8612878C66BD4AFEBD" author="Schultze" box="[429,583,299,323]" firstAuthor="Schultze" pageId="5" pageNumber="6" pagination="101 - 108" refId="ref14584" refString="Schultze, H. - P. (1980). Eier legende und lebendgebarende Quastenflosser. Natur und Museum, 110, 101 - 108." type="journal article" year="1980">Schultze, 1980</bibRefCitation>
;
<bibRefCitation id="EFDC4B43F34AFF86117F8C66BFC7FE9D" author="Witzmann" etAl="et al." firstAuthor="Witzmann" pageId="5" pageNumber="6" pagination="309 - 316" refId="ref15023" refString="Witzmann, F., Dorka, M., &amp; Korn, D. (2010). A juvenile Early Carboniferous (Visean) coelacanth from Rosenbeck (Rhenish Mountains, Germany) with derived postcranial characters. Fossil Record, 13, 309 - 316. https: // doi. org / 10.1002 / mmng. 201000004" type="journal article" year="2010">Witzmann et al., 2010</bibRefCitation>
). Tis view is reinforced because the specimen shows some well ossified axial mesomere and the scapulocoracoid (
<figureCitation id="13762A37F34AFF8613B48CC6BFFDFE5D" box="[158,240,395,419]" captionStart="Fig" captionStartId="7.[168,199,1783,1803]" captionTargetBox="[152,1438,237,1949]" captionTargetId="figure-14@8.[198,1392,253,1911]" captionTargetPageId="8" captionText="Fig. 4 Skeleton of Libys callolepis sp. nov. on the counterpart (holotype, NMBE 5034072).A Photos with osteological details: 1, articular head of the scapulocoracoid. 2, Scales on the flank immediately beneath the first anterior dorsal fin. 3, Scales of the lateral line showing the ornamental pattern with the larger central tubercles (white arrowheads point, showed only on one scale).4, Scales on the ventral flank from the pelvic to the anal fin. 5, Axial mesomere (white arrowhead) surrounded by some fin rays of the anal fin. 6, Axial mesomeres (white arrowhead) partially covered by sediment in the pelvic fin. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126233" httpUri="https://zenodo.org/record/13126233/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
<figureCitation id="13762A37F34AFF8613DA8CC6BFF4FE5D" box="[240,249,395,419]" captionStart="Fig" captionStartId="1.[168,199,1175,1195]" captionTargetBox="[152,1437,237,1315]" captionTargetId="figure-295@1.[169,1421,253,1162]" captionTargetPageId="1" captionText="Fig. 1 Maps showing the localization of the fossiliferous outcrops. A Paleogeographical map showing the continent configuration during the Toarcian (modified from Scotese,2014, Map 39) with the distribution of (1) Diplurus longicaudatus, (2) Holophagus gulo, (3) Trachymetopon, (4) Undina (?) barroviensis, (5) Libys callolepis sp. nov. (6) Atacamaia solitaria and (7) Indocoelacanthus robustus. B Paleogeographic map of the NW Tethys Ocean (modified from Fantasia et al., 2019), enlargement of B (red square). C Map of Switzerland with the locality of Les Pueys near the Teysachaux summit (Canton of Fribourg, Switzerland)" figureDoi="http://doi.org/10.5281/zenodo.11998922" httpUri="https://zenodo.org/record/11998922/files/figure.png" pageId="5" pageNumber="6">1</figureCitation>
, A
<figureCitation id="13762A37F34AFF8612308CC6BE2BFE5D" box="[282,294,395,419]" captionStart="Fig" captionStartId="9.[168,199,1857,1877]" captionTargetBox="[152,1438,237,1889]" captionTargetId="figure-11@9.[284,1304,253,1844]" captionTargetPageId="9" captionText="Fig. 5 Skull of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073). A Photos and B outline of the skull" figureDoi="http://doi.org/10.5281/zenodo.11998937" httpUri="https://zenodo.org/record/11998937/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
, A
<figureCitation id="13762A37F34AFF86126F8CC6BE5FFE5D" box="[325,338,395,419]" captionStart="Fig" captionStartId="13.[168,199,1136,1156]" captionTargetBox="[152,1437,237,1136]" captionTargetId="figure-273@13.[168,1420,253,1124]" captionTargetPageId="13" captionText="Fig. 6 Comparison of scales between Libys callolepis sp. nov. and some other Latimeriidae. A Libys callolepis sp. nov. (holotype, NMBE 5034072). Scales of the flank below the neural arches between the basal plates of the anterior and posterior dorsal fin. B Libys polypterus (L. superbus) (BSM AS I 801a). Scales of the flank located between the neural arches and the basal plate of the second dorsal fin. C Undina penicillata (BSM 1870 XIV 22). Scales of the middle flank located on the neural arches below the basal plate of the second dorsal fin. D Trachymetopon (Macropoma) substriolatum (holotype, SMC J27415) from the Kimmeridgian of Cottenham, Cambridgeshire. Scales of the middle flank. It is worth noting that this species was first attributed by Huxley (1866) to the genus Macropoma and latter to Coccoderma by Reis (1888). Forey (1998) kept this species within Coccoderma, but remarked that it shows affinities with Holophagus. Recently,Cavin et al. (2021a) attributed this specimen to the genus of Trachymetopon regarding, among other characteristics, the coarse ornamentation on the dermal bones" figureDoi="http://doi.org/10.5281/zenodo.11998940" httpUri="https://zenodo.org/record/11998940/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
, B). Although a long supplementary lobe of the caudal fin is considered as a juvenile character (e.g., Forey, 1981;
<bibRefCitation id="EFDC4B43F34AFF8612368C86BEB8FE1D" author="Schultze" box="[284,437,459,483]" firstAuthor="Schultze" pageId="5" pageNumber="6" pagination="90 - 91" refId="ref14543" refString="Schultze, H. - P. (1972). Early growth stages in coelacanth fishes. Nature New Biology, 236, 90 - 91. https: // doi. org / 10.1038 / newbio 236090 a 0" type="journal article" year="1972">Schultze, 1972</bibRefCitation>
), the prominent supplementary lobe of the caudal fin (
<figureCitation id="13762A37F34AFF8612A08CA6BECBFDFD" box="[394,454,491,515]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) observed in the specimen of Teysachaux represents rather a generic character of the specimen.
</paragraph>
<paragraph id="8BF236B2F34AFF8613BD8F20BEDBFD7A" blockId="5.[151,776,621,1027]" box="[151,470,621,644]" pageId="5" pageNumber="6">
<emphasis id="B939EAA0F34AFF8613BD8F20BEDBFD7A" bold="true" box="[151,470,621,644]" pageId="5" pageNumber="6">Dermal bones of the skull roof</emphasis>
</paragraph>
<paragraph id="8BF236B2F34AFF8613BD8FC1BDCDFBFD" blockId="5.[151,776,621,1027]" pageId="5" pageNumber="6">
Te bones of the skull are only preserved on the part (
<figureCitation id="13762A37F34AFF8613B48FE6BFE8FD3D" box="[158,229,683,707]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
and
<figureCitation id="13762A37F34AFF8612348FE6BE26FD3D" box="[286,299,683,707]" captionStart="Fig" captionStartId="9.[168,199,1857,1877]" captionTargetBox="[152,1438,237,1889]" captionTargetId="figure-11@9.[284,1304,253,1844]" captionTargetPageId="9" captionText="Fig. 5 Skull of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073). A Photos and B outline of the skull" figureDoi="http://doi.org/10.5281/zenodo.11998937" httpUri="https://zenodo.org/record/11998937/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). Te skull roof is divided into a parietonasal and postparietal shields, free from one to the other and separated by the intracranial joint, which appears to be straight. Te postparietal shield appears to be smaller than the parietonasal shield, this last being about 1.45 longer. Although slightly lower, this ratio is however close to the ratios of Jurassic and Cretaceous coelacanths that have typically a parietonasal shield circa 1.5 to 2 times longer than the postparietal shield (e.g.,
<taxonomicName id="4C4D4D31F34AFF8612EF8EE6BD7CFC3D" authorityName="Munster" authorityYear="1842" box="[453,625,939,963]" class="Coelacanthi" family="Mawsoniidae" genus="Libys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coelacanthiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="polypterus">
<emphasis id="B939EAA0F34AFF8612EF8EE6BD7CFC3D" box="[453,625,939,963]" italics="true" pageId="5" pageNumber="6">Libys polypterus</emphasis>
</taxonomicName>
has a ratio of about 1.7). On the Teysachaux specimen, the dermal bones of the skull appear to be smooth and unornamented.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="C3576539F34BFF87139F8E61BEE4FCBD" box="[181,489,811,835]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BF236B2F34BFF87139F8E61BEE4FCBD" blockId="4.[181,489,811,835]" box="[181,489,811,835]" pageId="4" pageNumber="5">
<heading id="D0BA81DEF34BFF87139F8E61BEE4FCBD" box="[181,489,811,835]" fontSize="10" level="2" pageId="4" pageNumber="5" reason="7">
<taxonomicName id="4C4D4D31F34BFF87139F8E61BEE4FCBD" authority="Munster 1842" authorityName="Munster" authorityYear="1842" box="[181,489,811,835]" class="Coelacanthi" family="Mawsoniidae" genus="Libys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coelacanthiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="polypterus">
<emphasis id="B939EAA0F34BFF87139F8E61BE46FCBD" bold="true" box="[181,331,811,835]" italics="true" pageId="4" pageNumber="5">L. polypterus</emphasis>
Münster 1842
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<subSubSection id="C3576539F34BFF8713BD8E26BE29FB5D" pageId="4" pageNumber="5" type="diagnosis">
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<emphasis id="B939EAA0F34BFF8713BD8E26BE07FC7D" bold="true" box="[151,266,875,899]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD8EC6BE29FB5D" blockId="4.[151,775,875,1187]" pageId="4" pageNumber="5">
<taxonomicName id="4C4D4D31F34BFF8713BD8EC6BE24FC5D" box="[151,297,907,931]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8EC6BFDDFC5D" box="[151,208,907,931]" italics="true" pageId="4" pageNumber="5">Libys</emphasis>
species
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with the postparietal shield less than half the length of the parietonasal shield. Te teeth covering the palate and the lower jaw are very small, and most are rounded and bear delicate radiating ridges. 70 neural arches. Fin rays of the posterior dorsal, pectoral, pelvic and anal fins are expanded. Te pectoral fin is relatively long, reaching back to posterior level of pelvic fin. Te scales are covered with a sparse ornament of short ridges.
</paragraph>
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<emphasis id="B939EAA0F34BFF8713BD8986BED4FB1D" bold="true" box="[151,473,1227,1251]" pageId="4" pageNumber="5">Measurements and meristic</emphasis>
</paragraph>
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(SL) Standard length
<quantity id="4CB59B57F34BFF87125489A6BED6FAFD" box="[382,475,1259,1283]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="mm" value="600.0">600 mm</quantity>
.
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD8846BED1FABD" blockId="4.[151,775,1227,1347]" pageId="4" pageNumber="5">d1.f=10; d2.f= 1520; pect.f= 16; pelv.f= 19; ana.f =1820; cau.f= 21/19; n.a=70; h.a= 23.</paragraph>
</subSubSection>
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<emphasis id="B939EAA0F34BFF8713BD8826BE0EFA7D" bold="true" box="[151,259,1387,1411]" pageId="4" pageNumber="5">
<typeStatus id="54F68810F34BFF8713BD8826BE0EFA7D" box="[151,259,1387,1411]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD88C6BED5FA5D" blockId="4.[151,472,1387,1443]" box="[151,472,1419,1443]" pageId="4" pageNumber="5">
<materialsCitation id="3B253CEFF34BFF8713BD88C6BED5FA5D" box="[151,472,1419,1443]" collectionCode="BSM" pageId="4" pageNumber="5" specimenCode="BSM 1870" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBEB9EC9F34BFF8713BD88C6BE04FA5D" box="[151,265,1419,1443]" collectionCode="BSM" country="Germany" httpUri="http://grbio.org/cool/c14k-izbs" name="Berliner Staatisches Museum" pageId="4" pageNumber="5">BSM 1870</specimenCode>
.xrv.502, head only.
</materialsCitation>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD8886BEB3FA1D" blockId="4.[151,654,1483,1539]" box="[151,446,1483,1507]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8886BEB3FA1D" bold="true" box="[151,446,1483,1507]" pageId="4" pageNumber="5">
Horizon and
<typeStatus id="54F68810F34BFF8712058886BE6FFA1D" box="[303,354,1483,1507]" pageId="4" pageNumber="5">type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD88A6BD83F9FD" blockId="4.[151,654,1483,1539]" box="[151,654,1515,1539]" pageId="4" pageNumber="5">
Tithonian (Upper Jurassic),
<collectingRegion id="4989F850F34BFF8712EE88A6BD15F9FD" box="[452,536,1515,1539]" country="Germany" name="Bayern" pageId="4" pageNumber="5">Bavaria</collectingRegion>
,
<collectingCountry id="F35A7622F34BFF87110888A6BD84F9FD" box="[546,649,1515,1539]" name="Germany" pageId="4" pageNumber="5">Germany</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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Münster 1842
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<emphasis id="B939EAA0F34CFF8010078831BCADFA6A" bold="true" box="[813,928,1404,1428]" pageId="3" pageNumber="4">Diagnosis</emphasis>
(emended from Forey, 1998)
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A genus of medium-sized, relatively deep-bodied coelacanths. Te head is nearly as deep as long. Te postparietal shield is much expanded posterolaterally where the supratemporals are particularly large. Te palate is deep and tapers rapidly anteriorly and the symplectic is also long, in keeping with the rather deep head. Te cheek is covered with large thin bones which are, however, well separated from one another. Te postorbital and lachrymojugal are broad and the preopercle is long and straplike. Te opercle is substantially deeper than broad. Te subopercle and the preorbital are absent. Sclerotic ossicles are present. In the lower jaw the angular shows a prominent dorsal expansion and the principal coronoid is developed posterodorsally as a rounded finger-like process. Te most obvious specialization of
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are the sensory canals which open to the surface through a large groove crossed by pillars. Ornamentation is absent from the skull bones. Te shoulder girdle shows a very narrow cleithrum, clavicle and extracleithrum. Te anocleithrum is simple and sigmoid to blade-like. Te pelvic fin is located well behind the level of the anterior dorsal fin and is supported by very narrow pelvic bones. Te rays of the anterior dorsal fin and the caudal fin are ornamented with many prominent denticles. Fin rays of pectoral, ventral, posterior dorsal and anal fins are very closely articulated close to their bases. Te supplementary caudal fin is prominent developing apart from the caudal fin profile. Te lateral line scales carry a large sensory tube which opens through several secondary tubules. An ossified lung is present.
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@ -48,7 +49,7 @@ Prout
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; plate 34
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; plate 33 [EX]
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<figureCitation id="136D2A73FF199EEDBCAF14E0FEEF4BA0" box="[223,375,1693,1715]" captionStart="Fig" captionStartId="157.[116,155,1470,1489]" captionTargetBox="[93,1196,210,1439]" captionTargetId="figure-62@157.[93,1197,210,1440]" captionTargetPageId="157" captionText="Fig. 346. Morphology of Scea (³³). A, head of S. dimidiata, lateral view; B, head of S. dimidiata, frontal view; C, head of S. dimidiata, posterior view; D, head of S. auriflamma, lateral view; E, head of S. auriflamma, frontal view; F, head of S. auriflamma, posterior view; G, head of S. discinota, lateral view; H, S. auriflamma wings (illustration by J.S. Miller)." pageId="172" pageNumber="847">Figure 346G</figureCitation>
<figureCitation id="136D2A73FF199EEDBCAF14E0FEEF4BA0" box="[223,375,1693,1715]" captionStart="Fig" captionStartId="157.[116,155,1470,1489]" captionTargetBox="[93,1196,210,1439]" captionTargetId="figure-62@157.[93,1197,210,1440]" captionTargetPageId="157" captionText="Fig. 346. Morphology of Scea (³³). A, head of S. dimidiata, lateral view; B, head of S. dimidiata, frontal view; C, head of S. dimidiata, posterior view; D, head of S. auriflamma, lateral view; E, head of S. auriflamma, frontal view; F, head of S. auriflamma, posterior view; G, head of S. discinota, lateral view; H, S. auriflamma wings (illustration by J.S. Miller)." figureDoi="http://doi.org/10.5281/zenodo.13126165" httpUri="https://zenodo.org/record/13126165/files/figure.png" pageId="172" pageNumber="847">Figure 346G</figureCitation>
; plates 33, 39Q
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Warren
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; plate 33 [EX]
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Druce
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; plate 34
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; plate 33
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<emphasis id="B922EAE4FF279ED3BF3B16D7FC4449D3" box="[843,988,1193,1216]" italics="true" pageId="146" pageNumber="821">Josia fustula</emphasis>
Warren
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<figureCitation id="136D2A73FF279ED3BF6E16B5FB9849CD" box="[798,1024,1224,1246]" captionStart="Fig" captionStartId="128.[150,188,1616,1635]" captionTargetBox="[128,1230,212,1585]" captionTargetId="figure-0@128.[128,1231,210,1586]" captionTargetPageId="128" captionText="Fig. 336. Morphology of Josia. A, J. fustula ³ head, lateral view; B, J. fustula ³ head, lateral view; C, J. ligula ³ head, lateral view; D, J. ligula ³ head, frontal view; E, J. ligula ³ head, posterior view; F, J. mononeura ³ wings; G, J. frigida ♀ wings; H, J. ligula ³ wings; I, J. megaera ³ tegula; J, J. megaera ³ labial palpus (illustration by J.S. Miller)." figureDoi="http://doi.org/10.5281/zenodo.13126142" httpUri="https://zenodo.org/record/13126142/files/figure.png" pageId="146" pageNumber="821">Figure 336A, 336B</figureCitation>
; plate 32
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<emphasis id="B922EAE4FF2C9ED8BC9916E5FEE549BE" box="[233,381,1174,1197]" italics="true" pageId="153" pageNumber="828">Josia turgida</emphasis>
Warren
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; plate 33
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<figureCitation id="136D2A73FF2D9ED9BF3E145BFC484B28" box="[846,976,1574,1596]" captionStart="Fig" captionStartId="134.[150,188,1376,1395]" captionTargetBox="[131,1224,210,1346]" captionTargetId="figure-143@134.[131,1224,209,1346]" captionTargetPageId="134" captionText="Fig. 342. Genitalia of Josia subcuneifera (³ holotype, JSM-1375; ♀ JSM-1712). A, ³ genitalia; B, ³ Tg8; C, ³ St8; D, ♀ genitalia, ventral view; E, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.13126156" httpUri="https://zenodo.org/record/13126156/files/figure.png" pageId="152" pageNumber="827">Figure 342</figureCitation>
; plate 33
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; plates 32, 39M [EX]
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; plate 32
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; plate 28 [EX]
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; plate 26 [EX]
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; plate 26 [EX]
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; plate 26 [EX]
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holds true, the two are distinct.
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Fig. 284. Morphology of
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; plate 31
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; plate 31 [EX]
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@ -206,11 +207,11 @@ with a wing pattern similar to that of
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<figureCitation id="136D2A73FFC89E3CBF91136CFBD94C35" box="[993,1089,273,294]" captionStart="Fig" captionStartId="118.[150,188,1472,1491]" captionTargetBox="[131,1224,210,1442]" captionTargetId="figure-1@118.[131,1224,210,1442]" captionTargetPageId="118" captionText="Fig. 332. Scanning electron micrographs of Scea angustimargo and Notascea obliquaria (³³). A, FW of S. angustimargo (dorsal surface), showing area at anterolateral corner of DC; B, S. angustimargo, showing elongate scales at base of radial sector; C, FW of N. obliquaria (dorsal surface), showing fovea beyond anterolateral corner of DC, between bases of radial sector and R1; D, short, rounded scales within FW fovea of N. obliquaria; E, single round scale from FW fovea of N. obliquaria; F, close-up of rounded scale in E." pageId="125" pageNumber="800">332CF</figureCitation>
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<figureCitation id="136D2A73FFC89E3CB83E136CFBE24C35" box="[1102,1146,273,294]" captionStart="Fig" captionStartId="120.[150,188,1684,1703]" captionTargetBox="[131,1224,210,1653]" captionTargetId="figure-1@120.[131,1224,210,1653]" captionTargetPageId="120" captionText="Fig. 334. Genitalia of Notascea obliquaria (³ JSM-356, ♀ JSM-357). A, ³ genitalia; B, ³ St8; C, aedeagus; D, ³ Tg8; E, ♀ genitalia (illustration by A. Trabka)." pageId="125" pageNumber="800">334</figureCitation>
<figureCitation id="136D2A73FFC89E3CB83E136CFBE24C35" box="[1102,1146,273,294]" captionStart="Fig" captionStartId="120.[150,188,1684,1703]" captionTargetBox="[131,1224,210,1653]" captionTargetId="figure-1@120.[131,1224,210,1653]" captionTargetPageId="120" captionText="Fig. 334. Genitalia of Notascea obliquaria (³ JSM-356, ♀ JSM-357). A, ³ genitalia; B, ³ St8; C, aedeagus; D, ³ Tg8; E, ♀ genitalia (illustration by A. Trabka)." figureDoi="http://doi.org/10.5281/zenodo.13126138" httpUri="https://zenodo.org/record/13126138/files/figure.png" pageId="125" pageNumber="800">334</figureCitation>
; plate 32 [EX]
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View file

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; plate 32
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; plate 31
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; plate 30
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; plate 29 [EX]
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; plate 29 [EX]
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; plate 28
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<mods:namePart id="7933F1BDE3ACE7E0D551A2E5452CDE57">Barclay, Maxwell V. L.</mods:namePart>
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<mods:affiliation id="65DCF9EF0375D69C61ABB2623A355612">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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</mods:role>
<mods:namePart id="8E39A06D199A32251FA012339A3906B8">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="D55107BD44F00C3F0A1A56A8A69172F3">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:title id="B06D5F0385936C55DEA73CE36EE7639E">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="BC826DD461EEFBA4839EC55D6CE3C8D1">
<mods:date id="BE18B9712CFB6C70148A0F2406F0B3FF">2024</mods:date>
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<subSubSection id="01C4A73A28EDED17B48D1DA2C67F18ED" type="nomenclature">
<paragraph id="A53DDF4480AAE6B9197ADA28D7D24B03">
<taxonomicName id="108C69F02DCE82E3A2153597A6C10C49" ID-CoL="49DSX" authority="Pittino, 1987" authorityName="Pittino" authorityYear="1987" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="triestinae" subGenus="Omorgus">
<emphasis id="E267D52625E929429B05027F40D45F12" italics="true">Omorgus</emphasis>
(
<emphasis id="485369B9259A42382A2CC7B40AE2F474" italics="true">Omorgus</emphasis>
)
<emphasis id="36979BFEFF458C7241FEE34AF5987E07" italics="true">triestinae</emphasis>
<bibRefCitation id="3572C92A71A0D12BFDB4E93107281A2A" author="Pittino R" issue="17" journalOrPublisher="Giornale Italiano di Entomologia" pagination="377-397" refId="B84" refString="Pittino R (1987) New Coleoptera Trogidae from South America (XXXII contribution to the knowledge of Coleoptera Scarabaeoidea). Giornale Italiano di Entomologia 3 (17): 377 - 397." title="New Coleoptera Trogidae from South America (XXXII contribution to the knowledge of Coleoptera Scarabaeoidea)." volume="3" year="1987">Pittino, 1987</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D3A5A111B064B36C77AF8174D5F1FB0E" type="reference_group">
<paragraph id="FD2ADE74ABCC2A18595AE6847CE0CB96">
<treatmentCitationGroup id="4BD5D962682706A234F13993D3BC301F">
<treatmentCitation id="3247960B8C093EC47FABA289E2729323" author="Pittino R" page="378" year="1987">
<taxonomicName id="13FBBEA1D9AB50B888E9A81B2F9C9767" ID-CoL="49DSX" authority="Pittino, 1987: 378" authorityName="Pittino" authorityPageNumber="378" authorityYear="1987" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="triestinae">
<emphasis id="06CDEDB9B1C683375C72689E765EF544" italics="true">Omorgus triestinae</emphasis>
<comment id="491EAF130A8162C36AE728137C889A93">
<bibRefCitation id="9BD1C8C041E37351D1789ACDF406F545" author="Pittino R" issue="17" journalOrPublisher="Giornale Italiano di Entomologia" pagination="377-397" refId="B84" refString="Pittino R (1987) New Coleoptera Trogidae from South America (XXXII contribution to the knowledge of Coleoptera Scarabaeoidea). Giornale Italiano di Entomologia 3 (17): 377 - 397." title="New Coleoptera Trogidae from South America (XXXII contribution to the knowledge of Coleoptera Scarabaeoidea)." volume="3" year="1987">Pittino, 1987: 378</bibRefCitation>
;
<treatmentCitation id="61B6580F8BBE69A3454CE3CEB46F91FC" author="Scholtz CH" page="1407" year="1990">
<bibRefCitation id="6E061D8FA0BFBFFDA6B0B34B11011EE3" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1407" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 13911456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1407</bibRefCitation>
</treatmentCitation>
(as new synonym of
<taxonomicName id="9DF74AEB6A2CB82F1FA3730F3C625BDC" ID-CoL="49DSR" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="suberosus">
<emphasis id="49FED66D894DBE6C8B07FC46CE442567" italics="true">Omorgus suberosus</emphasis>
</taxonomicName>
),
<treatmentCitation id="DF8AB62D1CAFF07D57FA712E9277B684" author="Zidek J" page="17" year="2013">
<bibRefCitation id="3464FD9CF1415E3E7954A8E071E77080" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="17" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 138." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 17</bibRefCitation>
</treatmentCitation>
(catalogue as synonym of
<taxonomicName id="B4E1E227CA3C50423FE0B8C5D408E21C" ID-CoL="49DSR" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="suberosus">
<emphasis id="39415A153EC983C42B0866478A347AC6" italics="true">Omorgus suberosus</emphasis>
</taxonomicName>
);
<treatmentCitation id="A83DFD8658940AC0DD1F8030F5AB7417" author="Smith ABT" page="88" year="2017">
<bibRefCitation id="41CDAE6F0708E80993E50A00255A0439" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Revista Chilena de Entomología" page="88" pagination="81-103" refId="B102" refString="Smith ABT (2017) Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution. Revista Chilena de Entomología 43: 81103." title="Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution." volume="43" year="2017">Smith 2017: 88</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="213DE8568F736C699FDAF7C11F931AE8" ID-CoL="49DSR" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="suberosus">
<emphasis id="4493FCEBE1006D433C35025700C2D082" italics="true">Omorgus suberosus</emphasis>
</taxonomicName>
);
<treatmentCitation id="062C9ACF5AECA04F73DD06B602A9D670" author="Zidek J" page="109" year="2017">
<bibRefCitation id="8573FF19FBACE66C10C409A7DF8FD747" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="109" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 109</bibRefCitation>
</treatmentCitation>
(catalogue as synonym of
<taxonomicName id="31FA9B31C2FED23ACF3427DF0BA29035" ID-CoL="49DSR" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="suberosus">
<emphasis id="D42EA40BC422D31BAF7D65E3EDF961D4" italics="true">Omorgus suberosus</emphasis>
</taxonomicName>
);
<treatmentCitation id="99B643C399B6609DCA403E1DDC75AD07" author="Huchet JB &amp; Costa-Silva V" page="559" year="2018">
<bibRefCitation id="1884B9283DFD3CCB4FDBFEF8AE82934D" DOI="10.11646/zootaxa.4524.5.3" author="Huchet JB &amp; Costa-Silva V" firstAuthor="Huchet" issue="5" journalOrPublisher="Zootaxa" page="559" pagination="553-566" refId="B56" refString="Huchet JB, Costa-Silva V (2018) A new species of Polynoncus Burmeister, 1876 from Brazil (Coleoptera: Trogidae). Zootaxa 4524 (5): 553566. https://doi.org/10.11646/zootaxa.4524.5.3" title="A new species of Polynoncus Burmeister, 1876 from Brazil (Coleoptera: Trogidae)." volume="4524" year="2018">Huchet and Costa-Silva 2018: 559</bibRefCitation>
</treatmentCitation>
(reinstated as a valid name);
<treatmentCitation id="CB3D5DA7F7219B84883402DA62054717" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFB7FFF9FE7EF99DFC9E1B66" page="2016" year="2021">
<bibRefCitation id="21F25AD329EB464CA51392D43BC5BB56" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2016" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 2016</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQZCI" type="material">
<paragraph id="6E3E6AC4ADED178434D5DD863A94AB45">
<heading id="988BA57E22CF4084BB71E4DFBD8C8D7D" reason="title">Type specimens examined.</heading>
</paragraph>
<paragraph id="0182D52A975361CC9FBC04A711E187D5">
<materialsCitation id="327BEE233FD97CBB16FFBE6347F2A18B" collectionCode="MNHN" country="Brazil" location="Minas-Geraes" specimenCount="1" specimenCount-male="1" stateProvince="Minas-Geraes" typeStatus="holotype">
<emphasis id="81CB407E8CB254446811919E07582155" bold="true" italics="true">
<typeStatus id="3473E08D507E6E8D69C550702E9C251D" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="4C75EB7CB99E6261C00748257F8357C5" count="1" type="male"></specimenCount>
<named-content id="66EBB191D6E4462CF74D32A6B94EF143" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle, Paris">
<emphasis id="A3153DC55C243183B418B5A0C9EFE8A4" bold="true">
<collectionCode id="4058EE1AFF6F473E3B8B0A0A65909CBA" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</emphasis>
</named-content>
). See
<bibRefCitation id="EC7811D9AE4E3DEC01568E450DE1A11E" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2015" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 2015)</bibRefCitation>
for high quality images of the type specimen.
<emphasis id="CC0C7AA2505BD57E4C63F1EB98FBF604" bold="true">Type locality</emphasis>
: “ Brésil,
<collectingRegion id="AD5DD1D86815CEFCB458E544EC11B596">Minas-Geraes</collectingRegion>
” [
<collectingCountry id="63770BD1896FCA5A0FCF5BF17A1F0F7B" name="Brazil">Brazil</collectingCountry>
].
</materialsCitation>
</paragraph>
<paragraph id="096D2169FA012A8C91627A14064FB39C">
<emphasis id="82D1AD6AD1857E137BF929C56958AE98" bold="true">
<emphasis id="695DF90B250BD9E0FC81F6A0583257E2" bold="true" italics="true">
<typeStatus id="E74C3E7DAA08AE963E7BDCA797F48D67" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
Same data label of the
<typeStatus id="02C400F8F79799307CD3805C9DCBA00C" type="holotype">holotype</typeStatus>
(11
<named-content id="5046A88DCCACF661BACF6160EF97A649" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle, Paris">
<emphasis id="21AE6E6AD5AE4415C8A4B64D8CB63805" bold="true">
<collectionCode id="F5F4D6C23E97439FBFED8548A4CBBB6E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</emphasis>
</named-content>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES1CI" type="distribution">
<paragraph id="0BBC0C81CB02AA09EA57F7C28FDF9738">
<heading id="3F7EE1110DE6D1BBEE582780EC966254" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="DE9675E7ACDA4268928936467F7354A8">
<collectingCountry id="48FE3860EBB8A5E3E274F8F8801EC230" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="25BE8AD30CBE535F11C066D03BB01BAF" name="Brazil">Brazil</collectingCountry>
, and
<collectingCountry id="51C3FED005F575F6291D22F0CBBDA0FF" name="Paraguay">Paraguay</collectingCountry>
(
<bibRefCitation id="495898007329DA27B020F3F08258D65E" DOI="10.11646/zootaxa.4524.5.3" author="Huchet and Costa-Silva" firstAuthor="Huchet" issue="5" journalOrPublisher="Zootaxa" pagination="553-566" refId="B56" refString="Huchet JB, Costa-Silva V (2018) A new species of Polynoncus Burmeister, 1876 from Brazil (Coleoptera: Trogidae). Zootaxa 4524 (5): 553566. https://doi.org/10.11646/zootaxa.4524.5.3" title="A new species of Polynoncus Burmeister, 1876 from Brazil (Coleoptera: Trogidae)." volume="4524" year="2018">Huchet and Costa-Silva 2018</bibRefCitation>
;
<bibRefCitation id="F636B9AF925F302F8C80302BF702F25D" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,536 @@
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<mods:title id="0BFC938D91CE5320F5193DD7C16CB742">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:namePart id="6E1C9744FA4982E04EAD56D7E83693A9">da Costa-Silva, Vinícius</mods:namePart>
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<mods:affiliation id="4D1C4232CFB681A981738D37F9ECC367">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:namePart id="90E915F597AADCF07A5B17DD4C4F3774">Strümpher, Werner P.</mods:namePart>
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<mods:namePart id="777535A31D6CA8D70EFE2F6C25E76AF4">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="BA989DD623781E3C8AA798223F6D8852" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="2139A2BC532A16FB4AD89CEF8EA5B12D">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="A721AC221D481CA83C8D53E70051E02C" type="personal">
<mods:role id="DBAF5D8C6EFF6B562297B7A6399247A7">
<mods:roleTerm id="B9AD41853EADAF91440AAF1130C5B9D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36B5B2250DC53E22E03F2EA283EB3575">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="E53DF1596E33F168229CA68ADF1B8472" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="94914EE743EFD526A48E5041D3BD6ADB">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="32604D572D852F20286DF53D179B5E49">text</mods:typeOfResource>
<mods:relatedItem id="5193F21D38D53FE21C3C77E540F61EA9" type="host">
<mods:titleInfo id="A29DD0C8CB514C869F6EBF0ABB3ED2D1">
<mods:title id="597DEC026F681F5A3B36A366DE107C6F">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="F6183AA70B0732B338F825F5C695B03A">
<mods:date id="23AD7A44DD40266486E78534D16F807B">2024</mods:date>
<mods:detail id="6C80FBBD8386F58766F3BE8159CE7139" type="pubDate">
<mods:number id="4F0A41F6311C9F9FB87CA836B0541EBC">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="DA409DAF359F9D46897F184312E3F829" type="volume">
<mods:number id="56BC6CFAE9BD85E9C02F7A4ECC811164">74</mods:number>
</mods:detail>
<mods:detail id="AAF147FB1B8DC13D1A23CFA88AE99883" type="issue">
<mods:number id="189B2FD61E23FC7A729A529F6577EC73">1</mods:number>
</mods:detail>
<mods:extent id="AA17C7F4B24060823D7ADDF10783ABAB" unit="page">
<mods:start id="266DBD0C183B33856906C87DEE5E5303">81</mods:start>
<mods:end id="AB19C0A4BD99A5C4646F1C5FB17C2E0A">101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1AE9EAD722DD067CDDC8F01BD98D6B76">journal article</mods:classification>
<mods:identifier id="6985E7AFC8848333E5F22F96E1812186" type="DOI">10.3897/contrib.entomol.74.e126799</mods:identifier>
<mods:identifier id="6B7010622C43E0D88D9751E39ABDEF72" type="ZooBank">339AB68C-DCB1-435F-B88E-2C42D0A7B05E</mods:identifier>
</mods:mods>
<treatment id="444630C0E5CC5052B7AF0E8D509751C1" ID-DOI="http://doi.org/10.5281/zenodo.13123980" ID-Zenodo-Dep="13123980" ID-arpha="444630C0-E5CC-5052-B7AF-0E8D509751C1" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1" httpUri="http://treatment.plazi.org/id/444630C0E5CC5052B7AF0E8D509751C1">
<subSubSection id="469810FBF06D7FBBDF3D1B71087287FD" type="nomenclature">
<paragraph id="58C52711D80F10EB820AC126192F01E3">
<taxonomicName id="DD05640FE0A919B024835E3EF0D1095B" ID-CoL="49DQS" authority="(Harold, 1872)" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei" subGenus="Omorgus">
<emphasis id="5E08BACD149D9ADD72C9555EBAF52FA3" italics="true">Omorgus</emphasis>
(
<emphasis id="48F10D0B906E950909B6F96532D589B0" italics="true">Omorgus</emphasis>
)
<emphasis id="4D20A38DB2D1E84093D35788078214B0" italics="true">candezei</emphasis>
(
<bibRefCitation id="A86C986635B4068593387E2123092C31" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E03755BD876A092FD3A45F55A636B361" type="description">
<paragraph id="77E1D5823670D16CAFE901F2DAC81575">
<figureCitation id="7DB4A1A9D93745F6C50EC6622117AE79" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Trox candezei Harold, 1872 (now Omorgus candezei). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Jonathan Brecko (RBINS). Length of specimen: 10.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure1" httpUri="https://binary.pensoft.net/fig/1103863">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="026D186AA74294334D939BDE4ACE9781" type="reference_group">
<paragraph id="5726E768CCCC343D286E87B51734C206">
<treatmentCitationGroup id="F6CCB68261923F7C7C704C08BCE47890">
<treatmentCitation id="765C24CD58F331D91D6AC43CB653F1F2" author="Harold E" page="113" year="1872">
<taxonomicName id="FD9BAA4F39206041D8C6EC8237359F1A" ID-CoL="59DHV" authority="Harold, 1872: 113" authorityName="Harold" authorityPageNumber="113" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="BD862F72603609B481CB61371553ACF3" italics="true">Trox candezei</emphasis>
<comment id="A8B156E270322D04BF43743F51C9DF55">
<bibRefCitation id="6389C8DE798E986B2E85F31572A86C6A" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872: 113</bibRefCitation>
(original description);
<treatmentCitation id="4E7ACCE2D4640242EA648AA394F56383" author="Burmeister HCC" page="259" year="1876">
<bibRefCitation id="D9683710B5B01BCEAD6573C57E70575C" author="Burmeister HCC" firstAuthor="Burmeister" journalOrPublisher="Stettiner Entomologische Zeitung" page="259" pagination="241-268" refId="B18" refString="Burmeister HCC (1876) Die argentinischen Arten der Gattung Trox F. Stettiner Entomologische Zeitung 37: 241268." title="Die argentinischen Arten der Gattung Trox F." volume="37" year="1876">Burmeister 1876: 259</bibRefCitation>
</treatmentCitation>
(as syn. of
<taxonomicName id="2F5EF1B8590B9F3ACC3E4F9A764D66BE" ID-CoL="59DJ6" authorityName="Blanchard" authorityYear="1847" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="78EDB3B043818F6318C6050B60FCD10C" italics="true">Trox ciliatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="AC724A3A5EF6D5E4A95A7961FD3B8964" author="Bruch C" page="193" year="1911">
<bibRefCitation id="68863C66F4AB21BB3CD480650F1871E3" DOI="10.5962/bhl.title.8799" author="Bruch C" firstAuthor="Bruch" page="193" refId="B16" refString="Bruch C (1911) Catálogo sistemático de los Coleópteros de la República Argentina. Pars IV: Familias Lucanidae, Scarabaeidae (Coproini - Cetonini), Passalidae. Revista del Museo de La Plata, Tomo XVII, 181225. https://doi.org/10.5962/bhl.title.8799" year="1911">Bruch 1911: 193</bibRefCitation>
</treatmentCitation>
(checklist as syn. of
<taxonomicName id="1E2AE85560A2ADF4A0311C79B56B9662" ID-CoL="59DJ6" authorityName="Blanchard" authorityYear="1847" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="C8502587418F712F4B174BFEE321C050" italics="true">Trox ciliatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="1E1BBE9F28AEF600CC624D4259DEB159" author="Arrow GJ" page="55" year="1912">
<bibRefCitation id="5E92DBE31C49B1C472A24A17C630D609" author="Arrow GJ" firstAuthor="Arrow" journalOrPublisher="Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk" page="55" refId="B3" refString="Arrow GJ (1912) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk, 66 pp." title="Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43." year="1912">Arrow 1912: 55</bibRefCitation>
</treatmentCitation>
(catalogue as syn. of
<taxonomicName id="2AE18C940D5F244521CEF2EF396DA97C" ID-CoL="59DJ6" authorityName="Blanchard" authorityYear="1847" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="897DFAA3BBC8F93C6449B5768B8E09F0" italics="true">Trox ciliatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="6C942FA96D0465CC2B8F5D4DCD536007" author="Blackwelder RE" page="218" year="1944">
<bibRefCitation id="130D1A8F9A57534DA7A342B7484BAAF5" DOI="10.5479/si.03629236.185.2" author="Blackwelder RE" firstAuthor="Blackwelder" journalOrPublisher="Bulletin of the United States National Museum" page="218" pagination="218-219" refId="B10" refString="Blackwelder RE (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae. Bulletin of the United States National Museum 185: 218219. https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae." volume="185" year="1944">Blackwelder 1944: 218</bibRefCitation>
</treatmentCitation>
(checklist as syn. of
<taxonomicName id="011D9AC5EBC9249CB1CA141195B64A73" ID-CoL="59DJ6" authorityName="Blanchard" authorityYear="1847" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B54F2557AD87918C20E5674BF3D93218" italics="true">Trox ciliatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="C9BE9FF4F52BDE35C388B2E1E30DBFAB" author="Vaurie P" page="153" year="1962">
<bibRefCitation id="B8BB8B441D8293398AD364D3817B210F" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="153" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 153</bibRefCitation>
</treatmentCitation>
(redescription).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E1DFF55B1D10B8F64A3D9DCDDDE4371D">
<treatmentCitationGroup id="D595E99217335804178E7E10C1EBBC18">
<treatmentCitation id="7264014CB57289904B05316497D5338A" author="Scholtz CH" page="9" year="1982">
<taxonomicName id="7A4D655C54E2009FE950903B0CA67AC5" ID-CoL="59DHV" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei" subGenus="Omorgus">
<emphasis id="87EDB3ED93C1CDD1DEAE54EAB4565C05" italics="true">Trox</emphasis>
(
<emphasis id="FCF57A5613B501731404DDFA3C592579" italics="true">Omorgus</emphasis>
)
<emphasis id="47B85BDD773985F49837B0BDE00EE0CA" italics="true">candezei</emphasis>
</taxonomicName>
<comment id="8377F701BD60FAE3FB654602DD6F23E1">
:
<bibRefCitation id="79F1AAE31BAB3C2953DDEE7B1D1D830D" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="9" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 127." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 9</bibRefCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="ED231AD96FDEC287C2DD0591F005858F">
<treatmentCitationGroup id="F9F7116EB326B943622AAEE717DB95C8">
<treatmentCitation id="A09F3C7C1E0E91674D5F058D38DBBF4E" author="Scholtz CH" page="1406" year="1990">
<taxonomicName id="DAFCBE7E659B267DDF3106F3158772FD" ID-CoL="49DQS" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="049C0E915B94429D92D1688126896961" italics="true">Omorgus candezei</emphasis>
</taxonomicName>
<comment id="8729551516E124AE2CDBC7FB812505C3">
:
<bibRefCitation id="16A00EB0E569A300D9899B9D0B736280" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1406" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 13911456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1406</bibRefCitation>
(redescription);
<treatmentCitation id="DE1CA26AF2C10669C5B7379790AF2202" author="Diéguez VM &amp; Gómez RS" page="94" year="2004">
<bibRefCitation id="3078F9BF5F18859E5DDA05E30F78C331" author="Diéguez VM &amp; Gómez RS" firstAuthor="Dieguez" issue="1-2" journalOrPublisher="Revista de la Sociedad Entomológica de Argentina" page="94" pagination="92-95" refId="B36" refString="Diéguez VM, Gómez RS (2004) Aporte al conocimiento de las Trogidae de la Argentina. Revista de la Sociedad Entomológica de Argentina 63 (12): 9295." title="Aporte al conocimiento de las Trogidae de la Argentina." volume="63" year="2004">Diéguez and Gómez 2004: 94</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="7F99E038A2CFAE8AE1E8D9C43B77444C" author="Deloya C" page="122" year="2005">
<bibRefCitation id="58FD8D4FA52DDC0AB2A91B9C2C4CD395" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="484C07372F1861268C0E7410927A4EE2" author="Gómez RS" page="514" year="2008">
<bibRefCitation id="F57A22137ED4FB4B79F647EAEF6F31A2" author="Gómez RS" editor="Claps LE &amp; Debandi G &amp; Roig-Juñent S" firstAuthor="Gomez" journalOrPublisher="Sociedad Entomológica Argentina" page="514" pagination="509-518" refId="B46" refString="Gómez RS (2008) Trogidae. In: Claps LE, Debandi G, Roig-Juñent S (Eds) Biodiversidad de Artrópodos Argentinos II. Sociedad Entomológica Argentina, 509518." title="Trogidae." volumeTitle="Biodiversidad de Artrópodos Argentinos II." year="2008">Gómez 2008: 514</bibRefCitation>
</treatmentCitation>
(key and notes for Argentinean species);
<treatmentCitation id="A28C3602833D27A21F234829FF119827" author="Zidek J" page="8" year="2013">
<bibRefCitation id="177D6DA52C53330CD2BF1CA1A162CCC5" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="8" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 138." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 8</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="CCD87DF6834675EE2279DF7E5886CABF" author="Zidek J" page="99" year="2017">
<bibRefCitation id="28F671516F917C80482B25F9AA09059A" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="99" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 99</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="994F53114AC647F99A027AC8E8C50145" author="Smith ABT" page="84" year="2017">
<bibRefCitation id="5629B6FD83B5ABEACC6F536246CA3DDF" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Revista Chilena de Entomología" page="84" pagination="81-103" refId="B102" refString="Smith ABT (2017) Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution. Revista Chilena de Entomología 43: 81103." title="Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution." volume="43" year="2017">Smith 2017: 84</bibRefCitation>
</treatmentCitation>
(notes, distribution).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3CAG" type="Type specimen examined">
<paragraph id="65453C0281E67489CE4ABAE4AC9869E6">
<heading id="617BC16DC5E14A20C60255285432066E" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="497C313EF8AC0C836D524B9F597B8083">
<materialsCitation id="9A366A06F06F44CC4342EC3826C102DF" collectionCode="RBINS" collectorName="Candeze" country="Argentina" determinerName="Scholtz" location="Pampas of South America" specimenCount="1" typeStatus="holotype">
<emphasis id="B5FF953C1217510513F12B8A2982AB83" bold="true" italics="true">
<typeStatus id="F4C604B1E5ADFCAE0482742DD9D6F8EA" type="holotype">Holotype</typeStatus>
, by monotypy
</emphasis>
(
<named-content id="6C512C3F3B920E410067BCD80B4D6FA7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/royal-belgian-institute-natural-sciences" xlink_title="Royal Belgian Institute of Natural Sciences">
<emphasis id="6137F3DA84ABE15B49EAFDE514CC7375" bold="true">
<collectionCode id="2C041E5A7211BF1615D4424AA7E2FCA1" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" type="Museum">RBINS</collectionCode>
</emphasis>
</named-content>
Fig.
<figureCitation id="C2D3919DE41BCB664567C7DD839C9B36" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Trox candezei Harold, 1872 (now Omorgus candezei). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Jonathan Brecko (RBINS). Length of specimen: 10.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure1" httpUri="https://binary.pensoft.net/fig/1103863">1</figureCitation>
). First label [red, typeset]: “ Coll. R. I. Sc. N. B. /
<collectingCountry id="783778DC38742AE8C7C402D254A7C16E" name="Argentina">Argentine</collectingCountry>
/ Pampas / ex coll.
<collectorName id="49689AA0824095DC1D8A0503A1CA4694">Candeze</collectorName>
”. Second label [white]: “ det.
<bibRefCitation id="C77182C6D2A98C500AE4CA230EDDDBE2" author="Harold" firstAuthor="Harold" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold 1872</bibRefCitation>
/ [yellow card, Edgar von Harold`s handwriting] Tr. Candezei / Harold typ. ”. Third label [red, typeset]: “ TYPE ”. Fourth label [white aged with green border]: “ Candezei / Har. / Pampas / Ch. ”. Fifth label [white, printed]: “
<taxonomicName id="06A05EAD1F2385B3574DB01DEA6FA23F" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="5ED1CE23ECD53979AF95EF0E505F5200" italics="true">Omorgus candezei</emphasis>
</taxonomicName>
/ (Harold) / det.
<determinerName id="C89ABE0E0410A6D8A79D99DDDA6695C3">Scholtz</determinerName>
1988 ” (Fig.
<figureCitation id="86637EEF365F9BDC2EB925D566C49CA0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Trox candezei Harold, 1872 (now Omorgus candezei). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Jonathan Brecko (RBINS). Length of specimen: 10.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure1" httpUri="https://binary.pensoft.net/fig/1103863">1 C</figureCitation>
).
<emphasis id="16A18373437B55EEAA5BE08E2462A9BA" bold="true">Type locality</emphasis>
: “ die Pampas von Südamerika ” [the
<location id="D439DD667AA7C147D98E2277DD2EC865" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:D439DD667AA7C147D98E2277DD2EC865" country="Argentina" name="Pampas of South America">Pampas of South America</location>
].
</materialsCitation>
</paragraph>
<caption id="174EBD75E6B09DDB16F5808CD848689C" ID-DOI="10.3897/contrib.entomol.74.e126799.figure1" ID-arpha="D15C0638-8132-54BD-98A1-02782EF2351F" httpUri="https://binary.pensoft.net/fig/1103863" startId="F1">
<paragraph id="9BF49AAC054C56556D65D7DFB86AEF08">
<label id="F8CBDF309BF6D90F1D96416F0251933C">Figure 1.</label>
</paragraph>
<paragraph id="D25DD791110A7E2DC3384570F448AA14">
Holotype of
<taxonomicName id="14911D9820DCA6CB08997729B0A2A203" authority="Harold, 1872" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="8902DA3F7141692C04C1504E7489A5D6" italics="true">Trox candezei</emphasis>
<bibRefCitation id="45B72E2C3F47FFCC1DC8318DF28C47E2" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="F07A56E8E30C1118198F12E102A47D1E" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="1897EE085401364BB64205FFD2359A63" italics="true">Omorgus candezei</emphasis>
</taxonomicName>
).
<emphasis id="5661BAA0249525C5305FF967D48C2668" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="74BD2AC186442D33BEBC9D5F19731B02" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="0726E8605E4EBA317E7359AB0AD9DB8E" bold="true">C</emphasis>
. Labels. Photos by Jonathan Brecko (
<named-content id="7A4191C8DC5C17A62F6BA9465E4895B6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/royal-belgian-institute-natural-sciences" xlink_title="Royal Belgian Institute of Natural Sciences">RBINS</named-content>
). Length of specimen: 10.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESFAG" type="notes">
<paragraph id="17EC04C217520AA5249973F78E2F6ED5">
<heading id="D8B66B9B70151096643AE5370809C3D6" reason="title">Note.</heading>
</paragraph>
<paragraph id="70171FEF9F8A610B0CE530F15BC215FB">
According to
<bibRefCitation id="B9B624FA7B52110A46EABA58D8FEE028" author="Harold E" firstAuthor="Harold" journalOrPublisher="Coleopterologische Hefte" page="114" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold (1872: 114)</bibRefCitation>
, the
<typeStatus id="D9F725ED2375886C56E4886C8C78507A" type="holotype">holotype</typeStatus>
was found in a factory in Verviers (
<collectingCountry id="C143D08C03129A8FE3BD9F728AF3B7C9" name="Belgium">Belgium</collectingCountry>
) in a consignment of sheeps wool originally from the Pampas region (
<collectingCountry id="CA67936EC17BCD63B97C50301BDC6838" name="Argentina">Argentina</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3FAG" type="distribution">
<paragraph id="B183426B4B2FC5B701E438F4319EAC49">
<heading id="6EBE75BFDC1BC7209199E1B6C29E5BCE" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="7A7082F7F3698C4D039138CFCBDE107B">
<collectingCountry id="6F0F92493666EBC44C8C791DCEAB961C" name="Argentina">Argentina</collectingCountry>
(
<bibRefCitation id="D417E53E7FA6A3952C676F7834A0D75E" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962</bibRefCitation>
;
<bibRefCitation id="AC32FC89DEEACD69634C4F4D8DE7CA4E" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 13911456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
;
<bibRefCitation id="AE9163B5E435B381E2104ECA66E52731" author="Smith" firstAuthor="Smith" journalOrPublisher="Revista Chilena de Entomología" pagination="81-103" refId="B102" refString="Smith ABT (2017) Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution. Revista Chilena de Entomología 43: 81103." title="Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution." volume="43" year="2017">Smith 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOGAG" type="type material">
<paragraph id="CD6AA135C770B6C4DD778087D9F1EE31">
<heading id="D3742BD7657EF3A809844A20083A38BC" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="E92D0469BB6CDCEDFA2FC304C0234A86">
<emphasis id="E49B3236A9680856355E46E52A05A6A4" bold="true">
(
<specimenCount id="58473B20B0967B6DA1D7346B60E6274A" count="15" type="generic">15 specimens</specimenCount>
).
</emphasis>
<materialsCitation id="B1EFB25C8DE500C50423EF8C8BA7A805" country="Argentina" location="Buenos Aires" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingCountry id="5203BA7905782860A8D6D92D31A8D184" name="Argentina">ARGENTINA</collectingCountry>
<emphasis id="28DB778653FD409C22F11BDFDAE7AEAA" bold="true">
<collectingRegion id="D57A7E121CE9AE887B0FFCC50ADAF0E8" country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="9B78BE9BDB644EC276C1F2C14EFEFD44" collectionCode="MACN" collectorName="C. Bruch" country="Argentina" location="Cordoba" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1;
<collectorName id="10084889A564F3EE52C27A84178CF30B">C. Bruch</collectorName>
leg.;
<named-content id="24FA02F26570E664C867C1F8C8B9DA1D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales Bernardino Rivadavia">
<collectionCode id="2B1707F4A63060CBFB8BF8F256E7EC19" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
.
<emphasis id="81282263DCA1BAFB51CEE4B06D51E43E" bold="true">
<location id="241F42F4375B025E57927A7745271F84" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:241F42F4375B025E57927A7745271F84" country="Argentina" name="Cordoba" stateProvince="Ciudad Autonoma de Buenos Aires">Córdoba</location>
</emphasis>
</materialsCitation>
<materialsCitation id="3BB7006B33E614BA09E486701388242C" collectionCode="A, MACN" collectorName="Villa &amp; Yacanto &amp; Stevenin" country="Argentina" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1; [
<collectorName id="F9A723E02589E640B0A32C093770B69A">Villa</collectorName>
]
<collectorName id="D206CA554801C6FBE3CC46F88D1C7EE1">Yacanto</collectorName>
, A.
<collectorName id="ADF56476CC5E0386F1B18C4BB18DD675">Stevenin</collectorName>
leg.;
<named-content id="B349B6FDC9750052973C98C2A2F42632" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales Bernardino Rivadavia">
<collectionCode id="D939267AB96AB71B0579FE3DA272AA12" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="400E9E6CEF9099AA5C984676C564A60A" collectingDate="1959-04" collectionCode="CVMD" collectorName="M. J. Viana" country="Argentina" location="Calamuchita" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1;
<location id="26791BB34D728D347F292D4BDFA3E7A4" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:26791BB34D728D347F292D4BDFA3E7A4" country="Argentina" name="Calamuchita" stateProvince="Ciudad Autonoma de Buenos Aires">Calamuchita</location>
, “ El Sauco ”;
<date id="53C3F02DED396311DC12E1B17841D3CF" value="1959-04">
<collectingDate id="912E9C9AA2372A3F60BD383698417C7B" value="1959-04">Apr. 1959</collectingDate>
</date>
,
<collectorName id="F1DDABC22560FF4FE5897B05FA0BF5C1">M. J. Viana</collectorName>
leg.;
<abbrev id="ABBRID0ECHAG" xlink_title="Collection of Víctor Manuel Diéguez [Private Collection]">
<collectionCode id="67C703CCD6EF70E16D39BCCA1DFEF6A8">CVMD</collectionCode>
</abbrev>
</materialsCitation>
.
<materialsCitation id="B28E9E2633C6A57090BB71CE57018889" collectingDate="2010-03-22" collectingDateMax="2010-03-25" collectingDateMin="2010-03-22" collectionCode="CVMD" collectorName="G. Arriagada &amp; R. Garcez" country="Argentina" location="Reserva de Biosfera Nacunan" municipality="Santa Rosa" specimenCount="1" stateProvince="Mendoza">
<emphasis id="A12507FB764C34A1F422CE95F74D46EC" bold="true">
<collectingRegion id="C05CFF5ACA90F258E936F09554258AA5" country="Argentina" name="Mendoza">Mendoza</collectingRegion>
</emphasis>
• 1;
<collectingMunicipality id="D701174E06DAB6E01FCE7CAC4E501CE3">Santa Rosa</collectingMunicipality>
,
<location id="5781EFA50795CF606813E7087BC81EEE" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:5781EFA50795CF606813E7087BC81EEE" country="Argentina" municipality="Santa Rosa" name="Reserva de Biosfera Nacunan" stateProvince="Mendoza">Reserva de Biosfera Ñacuñan</location>
;
<date id="3F901EBF546E138B61554CF432094E03" value="2010-03-22" valueMax="2010-03-25" valueMin="2010-03-22">
<collectingDate id="0973D84AF3952E289FA16AA1E339A634" value="2010-03-22" valueMax="2010-03-25" valueMin="2010-03-22">2225 Mar. 2010</collectingDate>
</date>
;
<collectorName id="29D0855BFBC24025C102E87A0909342C">G. Arriagada</collectorName>
&amp;
<collectorName id="D63AFCD51102F60C4CE5F51A559CBFAF">R. Garcez</collectorName>
leg.;
<abbrev id="ABBRID0EIHAG" xlink_title="Collection of Víctor Manuel Diéguez [Private Collection]">
<collectionCode id="CBE57D2D34B4855076A9786A8B076AFC">CVMD</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="FA0815DCD4B01B1673D1D9B5DDE8686C" collectingDate="2002-12-11" collectingDateMax="2002-12-13" collectingDateMin="2002-12-11" collectionCode="IADIZA" collectorName="V. M. Dieguez" country="Argentina" county="Santa Rosa" location="Rio Negro" municipality="Reserva de Biosfera Nacunan" specimenCount="1" stateProvince="Mendoza">
1;
<collectingCounty id="68FB10F3E01EE485BB2B75D400C040A5">Santa Rosa</collectingCounty>
,
<collectingMunicipality id="8EC30DD85B7EDC5191B2B65D6FCA723C">Reserva de Biosfera Ñacuñan</collectingMunicipality>
;
<date id="5DD92F948B3E8F7EAEA112C11071CB55" value="2002-12-11" valueMax="2002-12-13" valueMin="2002-12-11">
<collectingDate id="1BEF0453628698561252BD207F4EDE98" value="2002-12-11" valueMax="2002-12-13" valueMin="2002-12-11">1113 Dec. 2002</collectingDate>
</date>
;
<collectorName id="2507830F80736032EACA502EF7A7540F">V. M. Diéguez</collectorName>
leg.;
<abbrev id="ABBRID0EMHAG" xlink_title="Instituto Argentino de Investigaciones de Zonas Áridas">
<collectionCode id="86CC39D9F116B2CF22242E4BBF850903">IADIZA</collectionCode>
</abbrev>
.
<emphasis id="3EE6BF7DB530E007F9CE73C317D62F57" bold="true">
<location id="4435EB90E1768BFD6BD3A60D056D4153" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:4435EB90E1768BFD6BD3A60D056D4153" country="Argentina" county="Santa Rosa" municipality="Reserva de Biosfera Nacunan" name="Rio Negro" stateProvince="Mendoza">Río Negro</location>
</emphasis>
</materialsCitation>
<materialsCitation id="6FE48DA564B53DAE14E7CDF3D1124E1E" collectingDate="1963-12" collectionCode="CMNC" collectorName="Bachmann" country="Argentina" location="Rio Negro" municipality="Rio Colorado" specimenCount="1" stateProvince="Mendoza">
1;
<collectingMunicipality id="64A37262C5AAD9E5FC712DDDDBE1B1FF">Río Colorado</collectingMunicipality>
;
<date id="83FF04A63C4E778A673216EF0C3C16FE" value="1963-12">
<collectingDate id="F49B12FC238188294B5F5FB19FD29C37" value="1963-12">Dec. 1963</collectingDate>
</date>
;
<collectorName id="A10E1D6A9380F1D86FA08CB2E43AE431">Bachmann</collectorName>
leg.;
<named-content id="5FE8EA9EFAE11339E4F2F765C1318A21" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="524B5C359D2868FDF6020708890E4054">CMNC</collectionCode>
</named-content>
.
<emphasis id="82DFF1F9774B31CCA899378F0AD7D75F" bold="true">
<location id="8B520E6AFB2865D2AE1A2B58F88542D2" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:8B520E6AFB2865D2AE1A2B58F88542D2" country="Argentina" municipality="Rio Colorado" name="Rio Negro" stateProvince="Mendoza">Río Negro</location>
</emphasis>
</materialsCitation>
<materialsCitation id="13283FB8113AE53C7CEECD8C37DB852D" collectingDate="1829-09" collectionCode="MNHN" collectorName="D'Orbigny" country="Argentina" location="Village de Patagones" specimenCount="1" stateProvince="Mendoza">
1;
<location id="4331AB6D6EBBCD416837FB7A6CB443F4" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:4331AB6D6EBBCD416837FB7A6CB443F4" country="Argentina" name="Village de Patagones" stateProvince="Mendoza">Village de Patagones</location>
[= Viedma];
<date id="4CB03879B4D7C47FB72AD1E4CFEAE210" value="1829-09">
<collectingDate id="E7656F0BE32428E641A6544011D19EA0" value="1829-09">Set. 1829</collectingDate>
</date>
;
<collectorName id="269C4C0CB86A1C6F553F58BA07E73D41">DOrbigny</collectorName>
leg.;
<named-content id="5D7ACCA996D39DB4FCC5893FEF2393A5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle, Paris">
<collectionCode id="E663E619D68A8AB930D32E8574D04D24" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="F8664BB5A9B996E6807FDAB52A9D45B7" collectingDate="1982-01-18" collectingDateMax="1982-01-26" collectingDateMin="1982-01-18" collectionCode="A, CMNC" collectorName="Howden" country="Argentina" location="18 km S Ariona" specimenCount="1" stateProvince="San Luis">
<emphasis id="7BF816D627B48C65CFC85D27217807C2" bold="true">
<collectingRegion id="1FB92443F3B35520491FBB4DE73CAD3C" country="Argentina" name="San Luis">San Luis</collectingRegion>
</emphasis>
• 2;
<location id="471693409D7227194408CFF2C8E7BD2B" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:471693409D7227194408CFF2C8E7BD2B" country="Argentina" name="18 km S Ariona" stateProvince="San Luis">
<quantity id="41FC80C56B3D8428408F1F885E4530FC" metricMagnitude="4" metricUnit="m" metricValue="1.8" unit="km" value="18.0">18 km</quantity>
S Ariona
</location>
;
<date id="4A76FBF7EDC6C776E7719329426E3DD6" value="1982-01-18" valueMax="1982-01-26" valueMin="1982-01-18">
<collectingDate id="8F297DB27B6275898404276877924CC3" value="1982-01-18" valueMax="1982-01-26" valueMin="1982-01-18">1826 Jan. 1982</collectingDate>
</date>
; H. &amp; A.
<collectorName id="5CB6DAD0FC65729E1F025CB654B06BDC">Howden</collectorName>
leg.;
<named-content id="832B575403BCB89912D7001C155FCEFB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="9ABC777A0314D4C68A68CEC41A7AB9DD">CMNC</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="F70CB95D61B8534DA6799E8CB3671B6F" collectingDate="1980-02" collectionCode="CVMD" collectorName="M. J. Viana" country="Argentina" location="Arizona" specimenCount="1" stateProvince="San Luis">
1;
<location id="75AF3D422A9DCE0E6AF040A6F99DA3E7" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:75AF3D422A9DCE0E6AF040A6F99DA3E7" country="Argentina" name="Arizona" stateProvince="San Luis">Arizona</location>
;
<date id="F05F9BB6EB644F5BD2D69D7ABBF91891" value="1980-02">
<collectingDate id="C994D52047437FC6C709A4E57AC0191B" value="1980-02">Fev. 1980</collectingDate>
</date>
;
<collectorName id="47A59459748ECCE2CC5FD3BEAAE63091">M. J. Viana</collectorName>
leg.;
<abbrev id="ABBRID0ECIAG" xlink_title="Collection of Víctor Manuel Diéguez [Private Collection]">
<collectionCode id="A2AFD9CD4A27D79FB72559314D462C1F">CVMD</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="A274DCB5B07797CBC2942323CBCC92E0" collectingDate="1995-11-24" collectionCode="CVMD" collectorName="Flores &amp; Roig" country="Argentina" location="Quines" specimenCount="1" stateProvince="San Luis">
1;
<location id="B154E436967F3C0DD95BE54469781964" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:B154E436967F3C0DD95BE54469781964" country="Argentina" name="Quines" stateProvince="San Luis">Quines</location>
;
<date id="39435019650BF75DD0091C20EB7B64F2" value="1995-11-24">
<collectingDate id="B15FB8CCBF40713116D7B5CFF8752DF4" value="1995-11-24">24 Nov. 1995</collectingDate>
</date>
;
<collectorName id="5D8D971E0AD08ED04EF9FD49F8806E2C">Flores</collectorName>
&amp;
<collectorName id="4AC19B8E90C6D45F0E9A3EE269349FDE">Roig</collectorName>
leg.;
<abbrev id="ABBRID0EGIAG" xlink_title="Collection of Víctor Manuel Diéguez [Private Collection]">
<collectionCode id="6BE2472ABCF065974A192ED88D99D579">CVMD</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="B0E1FF4B366DD350AF7B56F47DC25B96" collectingDate="1914-02" collectionCode="CVMD" collectorName="M. J. Viana" country="Argentina" location="San Geronimo" specimenCount="1" stateProvince="San Luis">
1;
<location id="011CE4E673FFDB6F82419D6527586F87" LSID="urn:lsid:plazi:treatment:444630C0E5CC5052B7AF0E8D509751C1:011CE4E673FFDB6F82419D6527586F87" country="Argentina" name="San Geronimo" stateProvince="San Luis">San Geronimo</location>
;
<date id="AA42A1F47A39171C2E94986907FF6A35" value="1914-02">
<collectingDate id="22531FCD64CA0E5BCA542F6D1BAF6DC0" value="1914-02">Fev. 1914</collectingDate>
</date>
;
<collectorName id="030F2FEAE80631A9C2E121E0D0C886B5">M. J. Viana</collectorName>
leg.;
<abbrev id="ABBRID0EKIAG" xlink_title="Collection of Víctor Manuel Diéguez [Private Collection]">
<collectionCode id="F9A7634576465ED7193C8FB1F6282DDC">CVMD</collectionCode>
</abbrev>
</materialsCitation>
.
<materialsCitation id="6435343ECAADAB80402799E0F5A44D09" collectionCode="MACN" collectorName="Weiser" country="Argentina" location="Santa Fe" specimenCount="1" stateProvince="Santa Fe">
<emphasis id="D33239A3730AF4ED48D71E2C72FA18B4" bold="true">
<collectingRegion id="E3B09BD21E873FB704A5CD1EEE61B4F4" country="Argentina" name="Santa Fe">Santa Fe</collectingRegion>
</emphasis>
• 1;
<collectorName id="BB78CDE66724EAD462E1FAF8271B4442">Weiser</collectorName>
leg.;
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<collectingRegion id="9C6B780C2F84A23C4E03CFFCEBE552DC" country="Argentina" name="Santiago del Estero">Santiago del Estero</collectingRegion>
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• 1;
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;
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<collectingDate id="84395719367B2E4C7E7B3A33332BD2A9" value="1957-03-22" valueMax="1957-05-22" valueMin="1957-03-22">22 Mar. 22 May 1957</collectingDate>
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,
<collectorName id="8EEC05C59E8F015AD1ADC0249DAC25C2">Golbach</collectorName>
leg.;
<named-content id="48725D4C585D5EF934AEF1808257588F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/fundaci%C3%B3n-miguel-lillo-0" xlink_title="Instituto Fundación Miguel Lillo">
<collectionCode id="204561847B131B24CC9A07BF1AC67AEE">IFML</collectionCode>
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<materialsCitation id="6844783451F4017EA088A8FFA61816CF" collectionCode="A, MACN" collectorName="Breyer" country="Argentina" location="Tucuman" specimenCount="1" stateProvince="Tucuman">
<emphasis id="8339E2AC74E5875A626F2421F0A2F51C" bold="true">
<collectingRegion id="BC444F9CF022244C9175EBECB0CAFE1E" country="Argentina" name="Tucuman">Tucumán</collectingRegion>
</emphasis>
• 1; A.
<collectorName id="5B5F77AB85510F11DAA99CB125DECD71">Breyer</collectorName>
leg.;
<named-content id="F9B33F12A4658DF6EA2CE2E8D36380B4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales Bernardino Rivadavia">
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</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,65 +1,65 @@
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<taxonomicName id="E6F03CB9D69B63833C6FEFACEDA10633" ID-CoL="49DRX" authority="Scholtz, 1990: 1412" authorityName="Scholtz" authorityPageNumber="1412" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles" subGenus="Omorgus">
<emphasis id="29989CB8B0F530B4BED6FDA63F6B21B4" italics="true">Omorgus</emphasis>
(
<emphasis id="34465DDFB9BDC6CDC18819440EFB82F3" italics="true">Omorgus</emphasis>
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<treatmentCitation id="2E1396F1009F96BAF4D47D9F66150BF5" author="Deloya C" page="122" year="2005">
<taxonomicName id="297CD42063B579CA7E7E416F48B1C8DF" ID-CoL="49DRX" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles">
<emphasis id="B02A5F40C251584226707F1949FC9AA2" italics="true">Omorgus nocheles</emphasis>
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:
<treatmentCitation id="26398C8E4500FB53489290072C720403" author="Deloya C" page="122" year="2005">
<bibRefCitation id="019FB8B70E7E0645990AECF49F78E9E8" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
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(checklist).
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@ -135,7 +133,7 @@
<heading id="2F319CD9CB9FB7D8760340BEFB3EF481" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="0A5731EBBB8CFA8207A8B52CE655063D">
<materialsCitation id="7291451BD4827CB7DAE4CFA73ACFFF80" collectingDate="1956-10-24" collectingDateMax="1956-10-25" collectingDateMin="1956-10-24" collectionCode="LACM" collectorName="Second &amp; Clarke Scholtz's &amp; Third &amp; Fig." country="Argentina" location="First" specimenCode="ENT 160322" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<materialsCitation id="7291451BD4827CB7DAE4CFA73ACFFF80" collectingDate="1956-10-24" collectingDateMax="1956-10-25" collectingDateMin="1956-10-24" collectionCode="LACM" collectorName="Second &amp; Clarke Scholtz's &amp; Third &amp; Fig." country="Argentina" location="Rio Negro" specimenCode="ENT 160322" specimenCount="1" specimenCount-male="1" stateProvince="El Bolson" typeStatus="holotype">
<emphasis id="0B50EF78F5E8B1E03F46096976EC4B04" bold="true" italics="true">
<typeStatus id="660A141DC886A0C85EC26312BC6644B4" type="holotype">Holotype</typeStatus>
, by original designation
@ -147,19 +145,13 @@
<collectionCode id="B1D213A8D05EF8A8583AB8BA86D061E9" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</emphasis>
</named-content>
).
<location id="C2A6B3AFB6CDB0303D8532C99DA8782B" LSID="urn:lsid:plazi:treatment:A182B442C86E55DAB2E2411DCF6DE621:C2A6B3AFB6CDB0303D8532C99DA8782B" country="Argentina" name="First">First</location>
label [white, typeset]: “ El Bolson, Rio Negro, /
). First label [white, typeset]: “ El Bolson, Rio Negro, /
<collectingCountry id="0A69F7ED15E70A0F71E9B86EEFA25CF1" name="Argentina">ARGENTINA</collectingCountry>
/
<date id="5ABB321AEE08BE1668763432AC0827B9" value="1956-10-24" valueMax="1956-10-25" valueMin="1956-10-24">
<collectingDate id="6F5236B17B110BF484B7DF76D8BFEAB7" value="1956-10-24" valueMax="1956-10-25" valueMin="1956-10-24">Oct. 2425, 1956</collectingDate>
</date>
/ Andor Kovacs ”.
<collectorName id="1DF7F26EC4465A8EFC8C9C67B69A72ED">Second</collectorName>
label [white with red border,
<collectorName id="7D2F02D73F78F71CBA1070B4B41E0803">Clarke Scholtzs</collectorName>
handwriting]: “
/ Andor Kovacs ”. Second label [white with red border, Clarke Scholtzs handwriting]: “
<taxonomicName id="CEB0BCE44FE5BB094405CB6AD243F028" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="2E0CAEC95657233AAED691E54B8A4D2D" italics="true">Omorgus</emphasis>
</taxonomicName>
@ -167,21 +159,22 @@ handwriting]: “
<taxonomicName id="4E8A56E8388BD30EAD000661297D3D36" authorityName="Scholtz, 1990:" authorityYear="1412" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles">
<emphasis id="1DAC39DE40BEC947955EDA1EE51E4557" italics="true">nocheles</emphasis>
</taxonomicName>
/ 1988 / Scholtz /
<typeStatus id="876BBB90B734ABBB03ABB6843C4624E0" type="holotype">HOLOTYPE</typeStatus>
”.
<collectorName id="D91AAF562281C8F03486105022E1F6CA">Third</collectorName>
label [white, printed]: “ [QR Code] /
/ 1988 / Scholtz / HOLOTYPE ”. Third label [white, printed]: “ [QR Code] /
<named-content id="FEEE47DB5F8AC4E956F4B67397E4C6E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Los Angeles County Museum of Natural History">
<collectionCode id="371EE81E9F28FDBF33ABD98AAFDA0655" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</named-content>
<specimenCode id="ED39F1A6817E9BD789909C3E25CA69B4" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">ENT 160322</specimenCode>
” (
<collectorName id="6FF2D71C4E8B973D349C2BC441DCEAD1">Fig.</collectorName>
” (Fig.
<figureCitation id="E3CFCD420D7512307ED4E5531A788F0D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Omorgus nocheles Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Aedeagus in dorsal view; D. Aedeagus in lateral view; E. Labels. Photos by Brian Brown (LACM). Length of specimen: 15.516.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure5" httpUri="https://binary.pensoft.net/fig/1103867">5 E</figureCitation>
).
<emphasis id="1B6A793ADDCE4BC43968060F57ED6724" bold="true">Type locality</emphasis>
: “ Argentina, El Bolsón, Río Negro ”.
: “
<collectingCountry id="208CEC9249AD0CED1235E162D91BF78A">Argentina</collectingCountry>
,
<collectingRegion id="D579A93BF9A73911F1FF14825D35EE6E">El Bolsón</collectingRegion>
,
<location id="5C577A1F25662EAB142D59A5AA54B48A" LSID="urn:lsid:plazi:treatment:A182B442C86E55DAB2E2411DCF6DE621:5C577A1F25662EAB142D59A5AA54B48A" country="Argentina" name="Rio Negro" stateProvince="El Bolson">Río Negro</location>
”.
</materialsCitation>
</paragraph>
<caption id="C7A5662C9FF94DB618D54A6100D4E422" ID-DOI="10.3897/contrib.entomol.74.e126799.figure5" ID-arpha="6FFC447E-F115-5C50-B1A4-1C2D0634977F" httpUri="https://binary.pensoft.net/fig/1103867" startId="F5">

View file

@ -0,0 +1,224 @@
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<mods:title id="92FCBF92F267CC6F611ADF8B385FE5A8">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<treatment id="A7BB9E96A795587296D1BFBFE26C500D" ID-DOI="http://doi.org/10.5281/zenodo.13123974" ID-Zenodo-Dep="13123974" ID-arpha="A7BB9E96-A795-5872-96D1-BFBFE26C500D" LSID="urn:lsid:plazi:treatment:A7BB9E96A795587296D1BFBFE26C500D" httpUri="http://treatment.plazi.org/id/A7BB9E96A795587296D1BFBFE26C500D">
<subSubSection id="896AD0F1A87BC3F7CC3CEF5FFAFE573E" type="nomenclature">
<paragraph id="0AF870508A7255243594B50B7E134556">
<taxonomicName id="E8DA01A125180F0495637D342EE090E8" ID-CoL="49DQP" authority="(Harold, 1872)" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="badeni" subGenus="Omorgus">
<emphasis id="1B3B125F618CE905B4F1CA8AB3731129" italics="true">Omorgus</emphasis>
(
<emphasis id="07C911419D72D9BA57558B659A03453D" italics="true">Omorgus</emphasis>
)
<emphasis id="BCCE07E11CBD0D38C1C5DCB5809CEA71" italics="true">badeni</emphasis>
(
<bibRefCitation id="F17F58E7BB09E2EEBFA5906A0F02233B" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8580E72B48448F85E558D16BEB7E6156" type="reference_group">
<paragraph id="688C3F049FEF938AF1FBCD5AF1B7B41F">
<treatmentCitationGroup id="74BF1D1DE1461EADF8C0C05C1E72E86A">
<treatmentCitation id="0A6F9067DFBC98C529E696B254D185E7" author="Harold E" page="83" year="1872">
<taxonomicName id="FA19350B70F6A188C2096BD06B972D1B" ID-CoL="59DH7" authority="Harold, 1872: 83" authorityName="Harold" authorityPageNumber="83" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="badeni">
<emphasis id="6286A814EDEFFE16B5848097C7A8197F" italics="true">Trox badeni</emphasis>
<comment id="27A2332EF519278689B97DF186F1F694">
<bibRefCitation id="74026A32A9B671DBB19AEF646C8B2201" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872: 83</bibRefCitation>
(original description);
<treatmentCitation id="4EB68CF1782FEB164489B429CBBBE6F1" author="Arrow GJ" page="54" year="1912">
<bibRefCitation id="B822C188CCD95A3756749ABB86E495E4" author="Arrow GJ" firstAuthor="Arrow" journalOrPublisher="Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk" page="54" refId="B3" refString="Arrow GJ (1912) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk, 66 pp." title="Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43." year="1912">Arrow 1912: 54</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="DF37FA8ADA34A7B204D203032CF96E7E" author="Vaurie P" page="147" year="1962">
<bibRefCitation id="CBD8D39EF1557E106988A5CE43552F06" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="147" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 147</bibRefCitation>
</treatmentCitation>
(as syn. of
<taxonomicName id="27218C3EABCD76DE2A248E4D4A9E32CF" ID-CoL="49DQQ" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei">
<emphasis id="5DF9EBA1447C316A55DF8DBA4BAF5669" italics="true">Omorgus borrei</emphasis>
</taxonomicName>
);
<treatmentCitation id="244CD4A236556E9C75EB97E0061F4532" author="Scholtz CH" page="9" year="1982">
<bibRefCitation id="D54EFD52DCB3E7C25AE1B9806005F015" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="9" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 127." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 9</bibRefCitation>
</treatmentCitation>
(as syn. of
<taxonomicName id="169D3B42F87DB6AE1A9FEAB03B532801" ID-CoL="49DQQ" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei">
<emphasis id="036F43FE584F687D5757DAD0889337F9" italics="true">Omorgus borrei</emphasis>
</taxonomicName>
).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="1EE796BCF00A1696B474CBEC22CE127E">
<treatmentCitationGroup id="F2FEFEB10889CD54F15ADC5BB442D70F">
<treatmentCitation id="36DC2CB2BB869A901B07ACB4373F9F4E" author="Scholtz CH" page="1416" year="1990">
<taxonomicName id="61A09DFACDB1FA000CB66D0E6F18F6A2" ID-CoL="49DQP" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="badeni" subGenus="Omorgus">
<emphasis id="D32E6F8EA74146CED0F9B07ACF7BC9D5" italics="true">Omorgus</emphasis>
(
<emphasis id="5B851125EE7A30D2BE10E75F7D346414" italics="true">Omorgus</emphasis>
)
<emphasis id="4F9EFF7B0A50FE2FEFFE421D3594839D" italics="true">badeni</emphasis>
</taxonomicName>
<comment id="1017C30E903B8D0B98C1235D119C521D">
:
<bibRefCitation id="8C13D2BF6C719ED3E185E23FEA9A60A6" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1416" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 13911456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1416</bibRefCitation>
(revalidated, redescribed and
<typeStatus id="32B6A488349D919F15AF3C01174C3BAE" type="lectotype">lectotype</typeStatus>
designated);
<treatmentCitation id="D47ECB2F7B10B30D2936E60F9A7DF974" author="Zidek J" page="7" year="2013">
<bibRefCitation id="B554AC7551B9B7D5AB357FE2B279209C" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="7" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 138." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 7</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="FDD111320833F2147A33F73DA458A5E1" author="Zidek J" page="98" year="2017">
<bibRefCitation id="96719BD23EBF5621DC9D91C742C52972" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="98" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 98</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="F957BEF3E33628D8F1443D04600B317C" ID-DOI="http://doi.org/10.5281/zenodo.10532089" ID-Zenodo-Dep="10532089" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFA6FFEEFE72FE32FB0518E5" page="2001" year="2021">
<bibRefCitation id="33554252635FCBB6B2E1A078C5F720B3" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2001" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 2001</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="95B9635B4BC160C54D5A8B0FF6CA7338">
<treatmentCitationGroup id="FAB455C3F036675C070C955DC4C0913D">
<treatmentCitation id="30643D144FFF22484DB0B694F6F71256" author="Morrone JJ" page="57" year="2001">
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<emphasis id="339E891BF90B702F97AA8A1656F15D07" italics="true">Omorgus badeni</emphasis>
</taxonomicName>
<comment id="6D9A945A5114834D9A2818969F362FDA">
:
<bibRefCitation id="95D734582D978F7882549A6936002B02" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Caldasia" page="57" pagination="43-76" refId="B71" refString="Morrone JJ (2001) Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism. Caldasia 23 (1): 4376." title="Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism." volume="23" year="2001">Morrone 2001: 57</bibRefCitation>
(mention, biogeography);
<treatmentCitation id="764D74B4585864F52DFF3C2FA857311C" author="Deloya C" page="122" year="2005">
<bibRefCitation id="9D58B3705D83608C2907B069376044BD" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="3C8F2704C25393E27BEE07147AD19E2F" author="Morrone JJ" page="57" year="2014">
<bibRefCitation id="EA968246B64BA2DBB3830C49CE61A5BC" DOI="10.11646/zootaxa.3782.1.1" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Zootaxa" page="57" pagination="1-110" refId="B72" refString="Morrone JJ (2014) Biogeographical regionalisation of the Netropical region. Zootaxa 3782 (1): 1110. https://doi.org/10.11646/zootaxa.3782.1.1" title="Biogeographical regionalisation of the Netropical region." volume="3782" year="2014">Morrone 2014: 57</bibRefCitation>
</treatmentCitation>
(mention, biogeography).
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EMSAE" type="Type specimen examined">
<paragraph id="CF6795ACC0B8B54195FE1F70E05603B3">
<heading id="5264E66FA55AD73FF1E16FD6937DECB9" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="34E3CCF14C4DD6100F67DBC21F95BAD0">
<materialsCitation id="6C9EA7160A962839614CF0AD951E813C" collectionCode="RBINS" country="Brazil" location="See" specimenCount="1" stateProvince="Ceara" typeStatus="lectotype">
<emphasis id="61EE383023A2D6CC029A559B616B6B15" bold="true" italics="true">
<typeStatus id="3E6F60049337F8A90DDB89E79C8A4069" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<named-content id="98241D71F244FEC4008D5255EAFAF91B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/royal-belgian-institute-natural-sciences" xlink_title="Royal Belgian Institute of Natural Sciences">
<emphasis id="F1B1B16F60F1C9A3A97EEDC276F0A603" bold="true">
<collectionCode id="7BC5FDCB8EC9123E4BBC6269EDDF4483" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" type="Museum">RBINS</collectionCode>
</emphasis>
</named-content>
).
<location id="AF75D2CB57EAACDE0276B2D545E45BBF" LSID="urn:lsid:plazi:treatment:A7BB9E96A795587296D1BFBFE26C500D:AF75D2CB57EAACDE0276B2D545E45BBF" country="Brazil" name="See" stateProvince="Ceara">See</location>
<bibRefCitation id="3E684A1DD84B0C7BBFCBCBDC7DB2BAFD" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2001" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 2001)</bibRefCitation>
for high quality images of the type specimen.
<emphasis id="B122E4C53768A10C0B1EDCB57F3165EC" bold="true">Type locality</emphasis>
: “
<collectingRegion id="F9EAA8023517413CD6F891E4CEFA5DA6" country="Brazil" name="Ceara">Ceará</collectingRegion>
,
<collectingCountry id="6E74ED7050D707FEE7728AC456A5E757" name="Brazil">Brazil</collectingCountry>
”.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0ECTAE" type="distribution">
<paragraph id="075E4252ADB90898771EF3B63CC724F7">
<heading id="841FEA6EE7886C269DC5AC613E9C51BF" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="288A6F51CEDA1BB8D3096360F8FAA045">
<collectingCountry id="D863206B4CC8CEC80220BC7D40316D7A" name="Brazil">Brazil</collectingCountry>
. For details of distribution, see
<bibRefCitation id="9A4545EB5EC38C0AD1FC35359969BBC7" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (3132): 19932024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021)</bibRefCitation>
.
<bibRefCitation id="B59F5F6B70554426C231608DF5C8ABE9" author="Morrone" firstAuthor="Morrone" issue="1" journalOrPublisher="Caldasia" pagination="43-76" refId="B71" refString="Morrone JJ (2001) Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism. Caldasia 23 (1): 4376." title="Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism." volume="23" year="2001">Morrone (2001</bibRefCitation>
,
<bibRefCitation id="976145B529AA2C6720E07ACFAA357A57" DOI="10.11646/zootaxa.3782.1.1" author="Morrone" firstAuthor="Morrone" issue="1" journalOrPublisher="Zootaxa" pagination="1-110" refId="B72" refString="Morrone JJ (2014) Biogeographical regionalisation of the Netropical region. Zootaxa 3782 (1): 1110. https://doi.org/10.11646/zootaxa.3782.1.1" title="Biogeographical regionalisation of the Netropical region." volume="3782" year="2014">2014</bibRefCitation>
) report this species from “ Northern
<collectingCountry id="13F6A348D63F670562B9C060279A28FE" name="Colombia">Colombia</collectingCountry>
and northwestern (sic!) ”, without specific locality. Although plausible, so far, no specimen of
<taxonomicName id="AC24386B10F56D750A9D6949803DDBB6" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="badeni" subGenus="Omorgus">
<emphasis id="E51206645C3F0F74A0318217AB41FC72" italics="true">Omorgus</emphasis>
(
<emphasis id="3902140C0E835D22DD2B438EB94A5EC3" italics="true">O.</emphasis>
)
<emphasis id="82E6AAD39DE668CFBA20A208B9BC179F" italics="true">badeni</emphasis>
</taxonomicName>
from
<collectingCountry id="9B9CED2EFDAA55329395FD4FA814CE05" name="Colombia">Colombia</collectingCountry>
has been examined by the authors.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,65 +1,65 @@
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<treatment id="ADD0D40BE88359D1A6767E32E3629EC7" ID-DOI="http://doi.org/10.5281/zenodo.13123984" ID-Zenodo-Dep="13123984" ID-arpha="ADD0D40B-E883-59D1-A676-7E32E3629EC7" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7" httpUri="http://treatment.plazi.org/id/ADD0D40BE88359D1A6767E32E3629EC7">
<subSubSection id="D6569D0108F90FA878365638FFA27BA8" type="nomenclature">
@ -83,7 +83,7 @@
<paragraph id="E9585441FC5061247706F91166720FDF">
<treatmentCitationGroup id="3C1BD2968F8481D1473A05AEC202FA26">
<treatmentCitation id="76CB5D0DD6839A26568827687AFCD6E9" author="Scholtz CH" page="1411" year="1990">
<taxonomicName id="6F4D785E9221EA175BD93EA69607BB66" ID-CoL="49DRH" authority="Scholtz 1990: 1411" authorityName="Scholtz 1990:" authorityPageNumber="1411" authorityYear="1411" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus" subGenus="Omorgus">
<taxonomicName id="6F4D785E9221EA175BD93EA69607BB66" ID-CoL="49DRH" authority="Scholtz 1990: 1411" authorityName="Scholtz" authorityPageNumber="1411" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus" subGenus="Omorgus">
<emphasis id="9A0CF510202397D9A9BFA953F36CEEF0" italics="true">Omorgus</emphasis>
(
<emphasis id="120ABD90D7A963121CC583EDAAAFC07A" italics="true">Omorgus</emphasis>
@ -107,15 +107,13 @@
</paragraph>
<paragraph id="907FFEBB0011282C2BAB97DABA882EE8">
<treatmentCitationGroup id="F5FE837D3DDF72F44846AED267A8BB83">
<treatmentCitation id="B3525DFD19B6B4C39D5FA853CAF52FA1">
<taxonomicName id="E18CB328D234437901F829293BD7ED23" ID-CoL="49DRH" authorityName="Scholtz 1990:" authorityYear="1411" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus">
<treatmentCitation id="B3525DFD19B6B4C39D5FA853CAF52FA1" author="Morrone JJ" page="63" year="2001">
<taxonomicName id="E18CB328D234437901F829293BD7ED23" ID-CoL="49DRH" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus">
<emphasis id="71C53E8C952551DA15A2A2C8BF629274" italics="true">Omorgus indigenus</emphasis>
</taxonomicName>
<comment id="6757F5DC9BF6F13C3CB63C35FAFC4279">
:
<treatmentCitation id="920A67688DFD082579609BF4C4C1C9F6" author="Morrone JJ" page="63" year="2001">
<bibRefCitation id="14108E38F62E9726345CD44ED0114D5E" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Caldasia" page="63" pagination="43-76" refId="B71" refString="Morrone JJ (2001) Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism. Caldasia 23 (1): 4376." title="Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism." volume="23" year="2001">Morrone 2001: 63</bibRefCitation>
</treatmentCitation>
(mention);
<treatmentCitation id="69E5A0D8DBC70E6332AA103C8F7E61C7" author="Deloya C" page="122" year="2005">
<bibRefCitation id="53978BDDBB9B20D070F3BB1C8D8BFEE4" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
@ -135,7 +133,7 @@
<heading id="3DA9EBBB345194AF9D48345D65FA9991" reason="title">Type specimens examined.</heading>
</paragraph>
<paragraph id="F9FE2DCD22A9547407828556EBCB8B64">
<materialsCitation id="88A57FBD0E761A3E8456DD0F416D6241" collectionCode="CMNC" collectorName="C. H. Scholtz" country="Ecuador" location="Bahia Manzanilla" municipality="Bahia Manzanilla" specimenCount="1" specimenCount-male="1" stateProvince="Galapagos" typeStatus="holotype">
<materialsCitation id="88A57FBD0E761A3E8456DD0F416D6241" collectingMethod="carrion traps" collectionCode="CMNC" collectorName="C. H. Scholtz" country="Ecuador" location="Bahia Manzanilla" municipality="Espanola Is." specimenCount="1" specimenCount-male="1" stateProvince="Galapagos" typeStatus="holotype">
<emphasis id="921F2720E1DCB131D4C2138382E5651D" bold="true" italics="true">
<typeStatus id="9F312ABA8FF82EF70B8C9500B866E002" type="holotype">Holotype</typeStatus>
, by original designation
@ -147,9 +145,7 @@
<collectionCode id="6B25FC5E5690CF7BA2BBB48257CC2F75">CMNC</collectionCode>
</emphasis>
</named-content>
). First label [white, typeset]: “ ECU: Galap; Espanola /
<location id="6660461765F2935488AD19334C4DDFE9" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:6660461765F2935488AD19334C4DDFE9" country="Ecuador" municipality="Bahia Manzanilla" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
/ 510. VI. 85, S &amp; J Peck / Prospis grove behind / beach,
). First label [white, typeset]: “ ECU: Galap; Espanola / Bahia Manzanilla / 510. VI. 85, S &amp; J Peck / Prospis grove behind / beach,
<collectingMethod id="F980E25BA8E0A7CDEE7CEDF06C7D894B">carrion traps</collectingMethod>
”. Second label [white with black border, printed]: “ [QR code] / Canadian Museum of / Musée canadien de la / NATURE / CMNCEN 00011596 ”. Third label [white with red border, Clarke Scholtzs handwriting]:
<taxonomicName id="BBB7C4FC3011C49ECC4A8C607AA902EC" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
@ -157,13 +153,13 @@
</taxonomicName>
/ indigenous /
<collectorName id="D778236E2D5D2118CB6C6778CF6F4770">C. H. Scholtz</collectorName>
1988 /
<typeStatus id="5E5BF3ADF8E49D30F1F91F24F5F78385" type="holotype">HOLOTYPE</typeStatus>
”.
1988 / HOLOTYPE ”.
<emphasis id="556357735B7F51055D12AB51B557240D" bold="true">Type locality</emphasis>
: “
<collectingMunicipality id="BC2C37A6666B2D0FC6604C19C0F2FE05">Bahia Manzanilla</collectingMunicipality>
, Española Is.,
<location id="E06B3C5126D1F31CF8D815417E87AD70" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:E06B3C5126D1F31CF8D815417E87AD70" country="Ecuador" municipality="Espanola Is." name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
,
<collectingMunicipality id="87F155EDAEDAE195962C585118AA5072">Española Is.</collectingMunicipality>
,
<collectingRegion id="3F8B059B424DF3B6200649B6B9D475AD" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
” [
<collectingCountry id="A22B443ABB804748B77B5B32EEB26877" name="Ecuador">Ecuador</collectingCountry>
@ -196,7 +192,7 @@ Labels. Photos by François Génier (
</paragraph>
</caption>
<paragraph id="5DC3A163E6C012956E454402D0482C71">
<materialsCitation id="8C220CD440F25A5EF6438ADAE0608C8F" country="Ecuador" location="Galapagos" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
<materialsCitation id="90EBF45ADC150B6C1A4041850F835591" collectedFrom="Prosopis grove behind beach" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectingMethod="carrion traps" collectionCode="CMNC" collectorName="S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" municipality="Espanola Is." specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
<emphasis id="6FE4887AD17D8B2B91B5D3F3A6A04C8B" bold="true" italics="true">
<typeStatus id="A1004E3D6BDDBF47E9D38B0A0498B26E" type="paratype">Paratypes</typeStatus>
examined
@ -207,16 +203,13 @@ examined
<emphasis id="8A58CEE7BA30B113E7556FB89791BA0A" bold="true">
<collectingRegion id="21D49FDAEFE54C6F7EBC5E3B8B5C768A" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="0FC5DF0230B647B7E5B1CC027A25228F" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="CMNC" collectorName="Espanola Is. &amp; Prosopis &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
15;
<collectorName id="CA86002861929AEE1A73DD3A7D4DC637">Española Is.</collectorName>
• 15;
<collectingMunicipality id="7C13990CE5E1E17A88732B764759523B">Española Is.</collectingMunicipality>
,
<location id="231F03F65B72598682A0CC00B5670B30" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:231F03F65B72598682A0CC00B5670B30" country="Ecuador" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
<location id="231F03F65B72598682A0CC00B5670B30" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:231F03F65B72598682A0CC00B5670B30" country="Ecuador" municipality="Espanola Is." name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<collectedFrom id="C76D39C942929318C33C54889B3B859D">Prosopis grove behind beach</collectedFrom>
;
<collectorName id="C36DE96F24C1BEFEB57A2D7CA28867DD">Prosopis</collectorName>
grove behind beach;
<collectingMethod id="A22090C2F9A540311667884F14FED301">carrion traps</collectingMethod>
;
<date id="F72110ED87B4E181D1970CF23A377C4A" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">
@ -230,7 +223,7 @@ leg.;
</named-content>
</materialsCitation>
<materialsCitation id="DB749C411A534F2B888CE41F004CE320" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="NHMUK" collectorName="Espanola Is. &amp; Prosopis &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
<materialsCitation id="DB749C411A534F2B888CE41F004CE320" collectedFrom="Prosopis grove behind beach" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectingMethod="carrion traps" collectionCode="NHMUK" collectorName="Espanola Is. &amp; Prosopis &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" municipality="Espanola Is." specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
2; same collection data;
<named-content id="96A8BEA9D5E6BC626A4B77B4B14474EF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="FF12F6FE37C8D1C8AE5F4271B3325695" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
@ -261,19 +254,16 @@ Endemic to
<specimenCount id="C44DAAC69A8046B97A25CA345B52BCE7" count="24" type="generic">24 specimens</specimenCount>
).
</emphasis>
<materialsCitation id="34B020451435EE41CB728958F03B0515" country="Ecuador" location="Galapagos" specimenCount="1" stateProvince="Galapagos">
<materialsCitation id="C8D68A6B343E27EA4EC860F5B90BA877" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="CMNC" collectorName="S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" municipality="Espanola Is." specimenCount="1" stateProvince="Galapagos">
<collectingCountry id="34A9A47F13665BCD98FE6EB18B353214" name="Ecuador">ECUADOR</collectingCountry>
<emphasis id="B38EA99FBA0B50A24F2B7102F7BEDCB2" bold="true">
<collectingRegion id="C478DDC8F764C50328E3B0C52A64E8FD" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="0AFCB62069FBB10F6EDA59BE264FF3E4" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos">
3;
<collectorName id="1130970AA8D2C3827D467416FE67288E">Española Is.</collectorName>
• 3;
<collectingMunicipality id="5ACDA4B3A034C798757522DB3FAD4D86">Española Is.</collectingMunicipality>
,
<location id="D427AFB25F630DD51B4E13CF67D76FE2" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:D427AFB25F630DD51B4E13CF67D76FE2" country="Ecuador" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
<location id="D427AFB25F630DD51B4E13CF67D76FE2" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:D427AFB25F630DD51B4E13CF67D76FE2" country="Ecuador" municipality="Espanola Is." name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<date id="71327C9CFE1BEB8E570C2732E797C412" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">
<collectingDate id="C84A906A2FC3781AC1F1A5E4C843A407" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">510 Jun. 1985</collectingDate>
@ -286,13 +276,13 @@ leg.;
</named-content>
</materialsCitation>
<materialsCitation id="09E966FDB46D10AAA453AE8F195BAFCB" collectingDate="1985-04-25" collectingDateMax="1985-05-02" collectingDateMin="1985-04-25" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. Peck" country="Ecuador" location="Bahia Manzanilla" municipality="Galapagos" specimenCount="1" stateProvince="Galapagos">
<materialsCitation id="09E966FDB46D10AAA453AE8F195BAFCB" collectingDate="1985-04-25" collectingDateMax="1985-05-02" collectingDateMin="1985-04-25" collectionCode="CMNC" collectorName="S. Peck" country="Ecuador" county="Galapagos" location="Bahia Manzanilla" municipality="Espanola Is." specimenCount="1" stateProvince="Galapagos">
15;
<collectingMunicipality id="B9F740F940ED5D7F842C9F79B4EC7112">Galapagos</collectingMunicipality>
<collectingCounty id="80A29A4DB7669E5B27C7553F8EB8D184">Galapagos</collectingCounty>
,
<collectorName id="5E2FF14EFEEBDCAEEC0E16765E9DB1BF">Española Is.</collectorName>
<collectingMunicipality id="5487EE453F58E968BD88F1EAB81E5267">Española Is.</collectingMunicipality>
,
<location id="37B408DB98959D55DFCC78A20DC9067C" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:37B408DB98959D55DFCC78A20DC9067C" country="Ecuador" municipality="Galapagos" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
<location id="37B408DB98959D55DFCC78A20DC9067C" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:37B408DB98959D55DFCC78A20DC9067C" country="Ecuador" county="Galapagos" municipality="Espanola Is." name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<date id="CCA2D3C61F56CFF8423F304844D9F9F1" value="1985-04-25" valueMax="1985-05-02" valueMin="1985-04-25">
<collectingDate id="0739CB44711063D86BC86D6BB74E7438" value="1985-04-25" valueMax="1985-05-02" valueMin="1985-04-25">25 Apr. 2 May 1985</collectingDate>
@ -305,13 +295,13 @@ leg.;
</named-content>
</materialsCitation>
<materialsCitation id="554F6F69D632899ACAFCEA06AB4805A5" collectingDate="1992-06-02" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. Peck" country="Ecuador" location="Pta. Suarez" municipality="Galapagos" specimenCount="1" stateProvince="Galapagos">
<materialsCitation id="554F6F69D632899ACAFCEA06AB4805A5" collectingDate="1992-06-02" collectionCode="CMNC" collectorName="S. Peck" country="Ecuador" county="Galapagos" location="Pta. Suarez" municipality="Espanola Is." specimenCount="1" stateProvince="Galapagos">
6;
<collectingMunicipality id="39C7FD0A8316B68CDBDB9EBEAAA36CA5">Galapagos</collectingMunicipality>
<collectingCounty id="4C8D106E5615465A1A0AEB5A1A2FFFA1">Galapagos</collectingCounty>
,
<collectorName id="6A7B41509819FB44DF3D5141CAE8BBFF">Española Is.</collectorName>
<collectingMunicipality id="95335BBA637F2B4D3CFD51F2B290C383">Española Is.</collectingMunicipality>
,
<location id="518CC963CCF79E9A5C46D8C0067692F4" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:518CC963CCF79E9A5C46D8C0067692F4" country="Ecuador" municipality="Galapagos" name="Pta. Suarez" stateProvince="Galapagos">Pta. Suarez</location>
<location id="518CC963CCF79E9A5C46D8C0067692F4" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:518CC963CCF79E9A5C46D8C0067692F4" country="Ecuador" county="Galapagos" municipality="Espanola Is." name="Pta. Suarez" stateProvince="Galapagos">Pta. Suarez</location>
;
<date id="D542D485DA0B5E5172803572B11A8A33" value="1992-06-02">
<collectingDate id="E90B840E9A032EC60E36C642E0656B9F" value="1992-06-02">2 Jun. 1992</collectingDate>

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