diff --git a/data/03/EB/5D/03EB5D28FF8306484DA0FB44FCF18DD2.xml b/data/03/EB/5D/03EB5D28FF8306484DA0FB44FCF18DD2.xml new file mode 100644 index 00000000000..011726e885a --- /dev/null +++ b/data/03/EB/5D/03EB5D28FF8306484DA0FB44FCF18DD2.xml @@ -0,0 +1,373 @@ + + + +Cyclocephala ignota, a New Species of Cyclocephala Dejean, 1821 from Bolivia (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) + + + +Author + +Ratcliffe, Brett C. + + + +Author + +Seidel, Matthias + +text + + +The Coleopterists Bulletin + + +2023 + +2023-03-17 + + +77 + + +1 + + +19 +23 + + + + +http://dx.doi.org/10.1649/0010-065x-77.1.19 + +journal article +304853 +10.1649/0010-065X-77.1.19 +46a65ae6-3993-4152-b3ee-044f1620d6b0 +1938-4394 +13242666 +urn:lsid:zoobank.org:pub:9FDA95A2-78A0-469B-8A8A-EF75A959E49E + + + + + + +Cyclocephala ignota +Ratcliffe and Seidel + +, +new species + + + + + +zoobank.org/ + +urn:lsid:zoobank.org:act: +97A9536A-74EC-4334-92EB-3CB936680056 + +( +Figs. 1–7 +, +11 +) + + + + +Type Material. + +Holotype + +labeled “ +BOLIVIA +: Nor Yungas, / +Caranavi +, + +iii.2010 + +/ local collector // Coll. +Matthias +/ Seidel 2020” and with our red holotype label + +. + +Three ♂ +paratypes +with the same data and with our yellow paratype label + +. +Holotype +deposited in the Naturhistorisches Museum Wien ( +NHMW +), Austria. One paratype deposited in National Museum Prague ( +NMPC +), Czech Republic; one +paratype +deposited in Matthias Seidel Collection ( +MSPC +), Wien, Austria; and one paratype deposited in the B. C. Ratcliffe Collection ( +BCRC +), +Lincoln, NE +, +USA +. + + + + +Diagnosis. +This species will key to couplet 55 for + +Cyclocephala pereirai +Martínez + +( +Figs. 8–10 +) in +Endrödi (1985) +where the protarsi are considered scarcely thickened and with a slightly curved inner claw versus strongly thickened and with a strongly curved inner claw. + +Cyclocephala pereirai + +was described from +Argentina +and remains rare in collections, and its parameres are similar to those of + +C. ignota + +(compare +Figs. 6 +and +10 +). + + +We characterize the male protarsi of our new species as slender and only slightly curved, thus suggesting + +C. ignota + +is closest to + +C. pereirai + +. However, in dorsal view the proportions of the pronotum and elytra are different between the two species (compare +Figs. 1 +and +8 +), and in lateral view + +C. ignota + +is dorsoventrally compressed, whereas + +C. pereirai + +is not (as is typical for most + +Cyclocephala +species + +) (compare +Figs. 2 +and +9 +). In addition, + +C. ignota + +has long, dense setae on the ventral surfaces of the metatibiae and metatarsomeres, and + +C. pereirai + +does not (compare +Figs.1, 3 +, and 5 vs. +Fig. 8 +). Lastly, the color of + +C. ignota + +is light brown, and the color of + +C. pereirai + +is testaceous to reddish brown, although color may be unreliable depending on age of the specimens and method of specimen preservation. + + + + + +Description of +Holotype + +( +Figs. 1–7 +). Male. Length 12.0 mm; width 6.5 mm. Head dark brown, reminder of body light brown. Body dorsoventrally compressed ( +Fig. 2 +). +Head: +Surface completely, densely rugopunctate with long, yellowish brown setae on frons either side of middle. Frontoclypeal region with a slender, transversely biarcuate carinula not reaching lateral margins. Clypeal apex broadly parabolic, distinctly and narrowly reflexed. Interocular width equals 8.0 transverse eye diameters; ocular canthi short. Antenna with 9 antennomeres, club subequal in length to antennomeres 2–6. Mandibles projecting beyond clypeal apex ( +Fig. 1 +). Mentum slightly convex, strongly narrowed to apex, and with long, dense, yellowish brown setae. +Pronotum: +Widest just before middle ( +Fig. 1 +). Surface with punctures moderate in size and density, setigerous; setae long, dense, yellowish brown ( +Fig. 4 +). Base with marginal bead. +Elytra: +Surface weakly roughened and disc with small, dense, shallow, setigerous punctures; setae long, sparse, yellowish brown. Sides weakly, transversely wrinkled; setae along lateral margins dense ( +Fig. 3 +). +Pygidium: +Disc with minute to small, sparse punctures; basal half and lateral angles more densely punctate and with long, sparse, yellowish brown setae ( +Fig. 5 +). In lateral view, surface weakly convex. +Legs: +Protibia tridentate. Protarsomere 5 and median claw slightly enlarged; claw weakly arcuate, slender, apex narrowly cleft. Metatarsus longer than metatibia, both with long, dense, yellowish brown setae on their ventral surfaces ( +Figs. 1, 3, 5 +). +Venter: +Prosternal process, if present, small and obscured by long, dense, yellowish brown setae. Thoracic ventrites and coxae with long, dense, yellowish brown setae ( +Fig. 3 +). +Parameres: +In caudal view, shape subtriangular, apices rounded ( +Figs. 6, 7 +). + + + +Figs. 1–7. +Holotype of + +Cyclocephala ignota + +, + +new species + +. +1) +Dorsal view; +2) +Lateral view; +3) +Ventral view; +4) +Frontal view; +5) +Caudal view; +6) +Parameres, caudal view; +7) +Parameres, lateral view. + + + +Variation. +Male ( +3 paratypes +). Length 12.0–12.5 mm; width 6.2–7.0 mm. The +paratypes +do not significantly differ from the +holotype +. + + + + +Etymology. +The specific epithet is from the Latin +ignotus +, an adjective referring to unknown or strange and used here in reference to the atypical body morphology of this species relative to other species of + +Cyclocephala + +. + + + + +Figs. 8–10. + +Cyclocephala pereirai + +. +8) +Dorsal view; +9) +Lateral view (to show body shape only); +10) +Parameres, caudal view. + + + + +Fig. 11. +Location of type locality of + +Cyclocephala ignota + +, + +new species + +at Caranavi, Caranavi Province in La Paz Department, Bolivia. + + + + +Distribution. + +Cyclocephala ignota + +is known only from Caranavi in the Department of +La Paz +in the Yungas region of +Bolivia +( +Fig. 11 +). + + +Locality Records. +Four specimens +examined. + +BOLIVIA +(3): + +LA PAZ +(3): +Caranavi Department +, +Caranavi +. + + + + +Temporal Distribution. +March (4). + + +Natural History. +Nothing is known of the life history of + +C. ignota + +. Adults of other species of + +Cyclocephala + +are nocturnal and commonly collected at lights. + + + + \ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFC2FFF5FDFF0DF7FC8FDCA1.xml b/data/31/57/E0/3157E00AFFC2FFF5FDFF0DF7FC8FDCA1.xml new file mode 100644 index 00000000000..15dacbadb05 --- /dev/null +++ b/data/31/57/E0/3157E00AFFC2FFF5FDFF0DF7FC8FDCA1.xml @@ -0,0 +1,738 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Zorodictyna silvadavilae + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +3890C5AE-5D35-464E-924E-DFF174ABC154 + + + + + +Figs 24–26 + + + +Zorodictyna +sp. + +CG47 – + +Wheeler +et al. +2017 + +. + + + + + +Diagnosis + + + +Males of + +Zorodictyna silvadavilae + +sp. nov. +are distinguished from those of + +Z. oswaldi +( +Lenz, 1891 +) + +by the retrobasal cymbial extension (CE) that extends at least ¼ the length of papal tibia and thick retrolateral paracymbial process (PP) (vs + +Z. oswaldi + +lacks the PP and has very short CE; +Griswold 1993 +: figs 19–21); males are distinguished from + +Z. almae + +sp. nov. +by having only a small or no crest on embolus, and the retrolateral branch of RTA simple ( +Fig. 25A–C +) (vs + +Z. almae + +has a large crest on the E and the retrolateral branch of RTA is forked apically, +Fig. 23A–C +). Females of + +Z. silvadavilae + +are distinguished from + +Z. oswaldi + +by lacking teeth on lateral lobes (LL) ( +Fig. 25D +) (vs + +Z. oswaldi + +have teeth on LL; +Griswold 1993 +: fig. 24). The females are distinguished from + +Z. almae + +by having an endogyne with copulatory ducts (CD) equal to spermathecae (S) ( +Fig. 25E +) (vs in + +Z. almae + +, the CD are much larger than S, +Fig. 23E +). + + + + + +Etymology + + + +The species name is a matronym in honor of Dr Diana Silva Dávila, who made an extensive expedition to +Madagascar +in 2003, collecting specimens and tissues for the Spiders–AToL (Assembling the Tree of Life) project. + + + + + +Type material + + + + + +Holotype + +MADAGASCAR +– + +Antsiranana Province + +• + +; +Marojejy Reserve +, + +8.4 km +NNW of Manantenina + +; +14°26′ S +, +49°45′ E +; + +700 m +a.s.l. + +; + +10–16 Nov. 1993 + +; +C. Griswold +, +J. Coddington +, +N. Scharff +, +S. Larcher +, +R. Andriamasimanana +leg.; general collecting; CEG008; [habitus and male palp illustrated]; + + + +CASENT_9006231; +CAS +. + + + +Paratypes + + + + +MADAGASCAR +– + +Antsiranana Province + +• +1 ♂ +; same data as for holotype; CASENT_9006231; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9082258; +USNM + +• + +1 ♀ +; same data as for preceding; +S. Larcher +leg.; [endogyne drawn]; CASENT_9006230; +USNM + +. – + + +Toamasina Province + +• +1 ♂ +; +Parc National Masoala +, +Ambohitsitondroina Mt. +, +Ambanizana +; +15°34′20″ S +, +50°00′25″ E +; + +900–1010 m +a.s.l. + +; 5 +Mar + +. + + + +Fig. 24. + +Zorodictyna silvadavilae + +sp. nov. +, male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). +A, C +. Dorsal view. +B, D +. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme. + + + +2003; +Diana Silva Dávila +leg.; raking tree trunks; montane rainforest; DNA vouchers ARACG000047; DSD0024; CASENT_9014998; +CAS +• + +1 ♀ +; same data as for preceding; CASENT_9015400; +CAS + +• + +2 ♀♀ +; +Parc National de Marojejy +, NW; + +810 m +a.s.l. + +; + +Feb. 2002 + +; +S. Goodman +leg.; +pitfall +; [habitus illustrated]; FMNH-INS 0000-056-622; +FMNH + +• + +1 ♀ +; +Mt. Ambanizana +; +15°34′29.9″ S +, +050°00′12.3″ E +; + +750–800 m +a.s.l. + +; + +1 Mar. 2003 + +; +D. Andriamalala +, +D. Silva Dávila +et al. +leg.; rainforest; DSD0017; CASENT_9015400; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +• +1 ♂ +; +R.N.I. Marojejy +, + +11 km +NW of Manantenina + +, +Antranohofa +; + +1225 m +a.s.l. + +; +14°26.2′ S +, +49°44.5′ E +; + +24 Oct.–2 Nov. 1996 + +; +S. Goodman +leg.; undisturbed mossy forest; +pitfalls +at camp #3; +Mad +1996#7; +FMNH + +. + + + + + +Description + + + + +Male +holotype + +(CASENT_9006231; CAS; except as otherwise indicated) + +MEASUREMENTS. Total length: 17.60. Carapace: length 10.50, width 8.30, height 4.50. + +COLORATION ( +Fig. 24A–B +). Dorsal and lateral of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white. + + +STERNUM ( +Fig. 24B +). Oval with slightly sinuous margins; 3.70 wide, 4.50 long. + +CHILUM. 0.18 high, 0.63 wide. +EYES. ALE 0.28; AME 0.23; PLE 0.28; PME 0.25; ALE–AME 0.30; AME–AME 0.40; PLE–PME 0.35; PME–PME 0.15; MOQ: AW 0.58, PW 0.73, L 0.65. +CLYPEUS. With ten long setae. +LEGS. Trochanters I–III shallowly notched, IV entire. Unpaired claws smooth, rudimentary on legs III and IV. Scopulae beneath all tarsi, apical half of metatarsus I and apical third of metatarsus II. + +Leg measurements ( +paratype +CASENT_9014998; CAS) + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I9.703.80
II8.503.707.908.003.9032.00
III7.503.406.007.703.6028.20
IV10.003.208.5012.604.7039.00
+
+ +Spination ( +paratype +CASENT_9014998; CAS) + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl2d3rl4pl1rl1
IIpl3d3rl4pl1rl1pl2d2rl3v2-2-2-2pl4rl3v2-3-2
IIIpl3d3rl4pl1rl1pl2d2rl2v2-2-2pl5rl5v2-1-2-2
IVpl4d3rl3pl1rl1pl2d2rl2v2-2-2pl5rl5v2-1-2-2
+
+ +MALE PALP ( +Fig. 25A–C +). Tibia with apical, short pronged VTA connected to conical RTA, behind which dorso-retrolateral excavation accommodates retrobasal cymbial extension; cymbium oval with narrowly-rounded apical extremity, without dorsal patch of chemosensitive setae, having stout retrolateral paracymbial process (PP) and retrobasal cymbial extension (CE) extending at least ¼ length of papal tibia; subtegulum large, filling most of cymbial concavity and extending to prolateral cymbial margin, and with probasal broad ridge (SL) that interlocks with tegulum; tegulum rounded with conical process (TL) on prolateral margin, this process interlocking with subtegular ridge; embolus (E) broad and thick, arising proapically, with pointed apex, small crest subapically and curved basomediam process (EBP) arising from embolus base; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) hyaline fan-shaped, narrow at base; paraconductor (Pc) small, cradling inner margin of E. + + + +Fig. 25. + +Zorodictyna silvadavilae + +sp. nov. +, genitalia drawings; A–C += +male holotype (CASENT_9006231; CAS); D–E += +female paratype (CASENT_9006230; USNM). +A +. Left palp, prolateral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Epigyne, ventral view. +E +. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels. + + + + +Female +paratype + +(CASENT_9015400; CAS; except as otherwise indicated) MEASUREMENTS. Total length: 23.00. Carapace: length 11.20, width 8.30, height 5.00. + + +COLOUR ( +Fig. 24C–D +). As in male: dorsal and lateral of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white. + + +STERNUM ( +Fig. 24D +). Oval with sides straight next to leg coxae; 4.10 wide, 4.90 long. + +CHILUM. 0.25 high, 0.75 wide. +EYES. ALE 0.30; AME 0.29; PLE 0.31; PME 0.28; ALE–AME 0.18; AME–AME 0.50; PLE–PME 0.48; PME–PME 0.23; MOQ: AW 0.73, PW 0.83, L 0.23. +CLYPEUS. With 14 long setae. +LEG. Scopulae beneath all tarsi, apical half of metatarsus I and apical third of metatarsus II. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I8.304.307.006.203.1028.90
II7.504.005.605.802.9025.80
III6.203.204.605.003.0022.00
IV8.803.607.0012.403.5033.30
+
+ +Spination GENITALIA. Epigyne ( +paratype +CASENT_9006230; USNM), ( +Fig. 25D +) with quadrangular median lobe (ML) plate transverse with smooth margin; copulatory openings (CO) hidden beneath anterolateral corners of ML; lateral lobes (LL) convex, without lateral teeth or projections but with faint concavities along margin of ML, converging posteriorly and exposing triangular area of median sector (MS). Endogyne ( +Fig. 25E +) with broad, semitransparent copulatory ducts (CD) extending from beneath ML to simple, convex, bean-shaped spermathecae (S), spermathecal head with several small pores; spermathecal bases meeting in center; fertilization ducts (FD) short and slightly curved. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl2d1v2-2-2-2v2-2-2
IIpl2d1v2-2-2-2v2-2-2
IIIpl2d3rl3pl1rl1pl2d2rl2v2-2-2pl4rl4v2-2-1-2
IVpl3d3rl1pl1rl1pl2d2rl2v2-2-2pl4d2rl3v2-2-1-2
+
+ + +Variation + + + +Males +(n = 4) + +Total length = 17.60–24.00, carapace length/width = 1.12–1.28, carapace height/width = 0.44–0.54, PER/carapace width = 0.20–0.22, PER/OAL = 2.50–2.80, PER/AER = 1.35–1.56, OAL/OQL = 1.00– 1.13, OQP/OQA = 1.22–1.46, sternum length/width = 1.22–1.33, palpal coxa length/width = 1.92–2.90, femur I length/carapace width = 1.04–1.23, metatarsus I length/carapace width = 0.98–1.16, femur IV length/carapace width = 1.10–1.31. Male palp with apical RTA tips acute (Marojejy) to blunt (Masoala), EBP hook at embolic base triangular (Marojejy) to crescentic (Masoala), retromedian tegular lock (TL) blunt (Marojejy) to sharply triangular (Masoala). Males from Marojejy have a small subterminal crest on the embolus, which is absent from the male from Masoala. + +Females +(n = 4) + +Total length = 23.00–32.50, carapace length/width = 1.26–1.41, carapace height/width = 0.46–0.60, PER/carapace width = 0.23–0.26, PER/OAL = 2.87–3.41, PER/AER = 1.30–1.43, OAL/OQL = 1.03– 1.06, OQP/OQA = 1.14–1.23, sternum length/width = 1.18–1.24, palpal coxa length/width = 2.32–3.45, femur I length/carapace width = 0.92–1.00, metatarsus I length/carapace width = 0.66–0.75, femur IV length/carapace width = 1.02–1.07. Epigyne with median lobe length 0.70–0.78 times width; distance from median lobe base to epigastric fold 0.29–0.40 times median lobe length. + + +Note on synonym + + + +A specimen of this species (CASENT_9014998) was used as an exemplar in a previously published molecular phylogeny: it was listed as “ + +Zorodictyna +sp. + +CG47” by + +Wheeler +et al. +(2017) + +. + + + +Natural history + + +This species has been collected when running on leaf litter in closed canopy montane rainforest. Individuals have an entire cribellum but we have yet to learn details of their use of silk. +
+ + + +Distribution + + + +Individuals of this species are known only from the montane rainforests on the Marojejy Massif and the Masoala Peninsula in northeastern +Madagascar +( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFCEFFF0FDDD092CFDB7DA9B.xml b/data/31/57/E0/3157E00AFFCEFFF0FDDD092CFDB7DA9B.xml new file mode 100644 index 00000000000..c3f9a6cd0ee --- /dev/null +++ b/data/31/57/E0/3157E00AFFCEFFF0FDDD092CFDB7DA9B.xml @@ -0,0 +1,716 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Zorodictyna almae + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +9F5FEFEF-155C-4C64-862B-3F6F67A52A01 + + + + + +Figs 22–23 +, +26 + + + +Zorodictyna +sp. 1 + +– + +Polotow +et al. +2015 + +. + + + + + +Diagnosis + + + +Males of + +Zorodictyna almae + +sp. nov. +are distinguished from + +Z. oswaldi +( +Lenz, 1891 +) + +by having a retrobasal cymbial extension (CE) that extends at least ¼ the length of the papal tibia and a thick retrobasal paracymbial process (PP) ( +Fig. 23B–C +) (vs + +Z. oswaldi + +lacks the PP and has a very short CE; +Griswold 1993: 19–21 +). It is distinguished from + +Z. silvadavilae + +sp. nov. +in having the male palp with a nearly triangular basal embolic hook, the embolus lacking a crest, and a retromedian longitudinal ridge that extends for most of the tegulum length ( +Fig. 23A–C +) (vs + +Z. silvadavilae + +has slender, crescentic basal embolic hook, the embolus with subapical distal crest, and the tegular retromedian longitudinal ridge that is weak, short, extending for less than one-half tegulum length, +Fig. 25A–C +). Females of + +Z. almae + +are distinguished from + +Z. oswaldi + +by lacking teeth on the lateral lobes (LL) of the epigyne ( +Fig. 23D +) (vs + +Z. oswaldi + +has the LL with teeth; +Griswold 1993 +: fig. 24); females of + +Z. almae + +are distinguished from + +Z. silvadavilae + +by having the lateral lobes of epigyne parallel and the median sector (MS) rectangular ( +Fig. 23D +). The female endogyne has copulatory ducts (CD) larger than spermathecae ( +Fig. 23E +) (vs in + +Z. silvadavilae + +, the LL are converging posteriorly, and the MS is triangular, +Fig. 25D +, and the endogyne has CD and spermathecae equal in size, +Fig. 25E +). + + + + + +Etymology + + + +The species name is a matronym in honor of Alma Saucedo Mejia, participant in the 2011 expedition that produced the female +paratype +(CASENT_9031271) that was a DNA voucher in + +Polotow +et al. +(2015) + +[DNA-DP-Lyc-voucher, Z-1, DP2014d]. She is also the discoverer of many +Madagascar +goblin spiders. + + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Mahajunga Province + +• + +; +R. F. l’Ankarafantsika +, + +5 km +SSW of Ampijaroa + +; +16°20.3′ S +, +46°47.6′ E +; + +160 m +a.s.l. + +; + +4–7 Feb. 1997 + +; +S. Goodman +leg.; CASENT_9082245; +FMNH +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Mahajunga Province + +• +1 ♀ +; same data as for holotype; CASENT_9082245; +FMNH + +• + +1 ♀ +; +Ampijoroa +, +National Park Ankarafantsika +, near +Lake Ravelobe +; 16°17.48′7″ S, 46°48.50′7″ E; + +26–28 Jan. 2009 + +; +C. Griswold +, +A. Saucedo +and +H. Wood +leg.; DNA-DP-Lyc-voucher, + +Zorodictyna +sp. 1 + +, in +Polotow D. +, +Carmichael A. +& +Griswold C.E. +2015; Z-1; DP2014d; CASENT_9031271; +CAS + +• + +1 ♀ +; +Parc National d’Ankarafantsika +, + +Forêt +de Tsimaloto + +, + +18.3 km +46° NE of Tsaramandroso + +; +16°3′41″ S +, +46°8′37″ E +; + +135 m +a.s.l. + +; + +2–8 Apr. 2001 + +; +J.J. Rafanomezantsoa +et al. +leg.; general collecting, day, tropical dry forest; [epigyne illustrated by CEG]; CASENT_9002829; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +– + +Mahajanga Province + +• +1 ♀ +; +Ampijoroa +, +National Park Ankarafantsika +, near +Lake Ravelobe +; +16°17.487′ S +, +46°48.507′ E +; + +26–28 Jan. 2009 + +; +C. Griswold +, +A. Saucedo +and +H. Wood +leg.; CASENT_9031270; +CAS + +. + + + + + +Description + + + + +Male +holotype + +(CASENT_9082245; FMNH) + +MEASUREMENTS. Total length: 21.00. Carapace: length 12.00, width 9.80, height 5.70. + +COLORATION ( +Fig. 22A–B +). Dorsal and lateral sides of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white. + + +STERNUM ( +Fig. 22B +). Oval with slightly sinuous margins; 4.50 wide, 5.60 long. + +CHILUM. 0.25 high, 0.85 wide. +EYES. ALE 0.28; AME 0.28; PLE 0.35; PME 0.30; ALE–AME 0.13; AME–AME 0.70; PLE–PME 0.45; PME–PME 0.28; MOQ: AW 0.73, PW 0.83, L 0.80. +CLYPEUS. With 18 long setae. +LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apices of metatarsi III and IV, and apical halves of metatarsi I and II. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I10.205.1011.5011.106.3044.20
II11.005.009.709.805.0040.50
III9.304.107.609.404.6035.00
IV11.904.7010.9015.206.0048.70
+
+ +Spination MALE PALP ( +Fig. 23 A–C +). Tibia VTA apical, curved and blunt-tipped, RTA triangular, sharply pointed, located behind this dorso-retrolateral excavation to accommodate retrobasal cymbial extension (CE); cymbium oval, apical extremity narrowly rounded, without dorsal patch of chemosensitive setae, with protruding retrolateral paracymbial process (PP), retrobasal cymbial extension (CE) conical, extending at least ¼ the length of papal tibia; subtegulum large, filling most of cymbial concavity and extending to prolateral cymbial margin, with probasal broad ridge (SL) that interlocks with tegulum; tegulum rounded with rectangular process on prolateral margin (TL), which interlocks with subtegular ridge; embolus (E) proapical, broad and thick, with pointed apex, inner margin with twisted longitudinal ridge and triangular basomedian process (EBP) arising from embolus base; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) hyaline, fan shaped, narrow at base; paraconductor (Pc) large, cradling inner margin of E and extending past E apex; retroventral surface of tegulum with deep longitudinal groove that extends for two thirds of tegulum length. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl3d3rl2pl1rl1pl3d3rl3v2-2-2-2pl4rl4v2-2-3
IIpl3d3rl3pl2d3rl2v2-2-2-2pl3rl3v2-2-3
IIIpl3d3rl3pl1rl1pl2d2rl2v2-2-2pl4d1rl4v2-2-2
IVpl3d3rl3pl1rl1pl2d2rl2v2-2-2pl4rl4v1-1-1-2-3
+
+ + +Female +paratype + +(CASENT_9082245; FMNH; except as otherwise indicated) MEASUREMENTS. Total length: 30.00. Carapace: length 14.00, width 11.30, height 4.40. + + +COLOUR ( +Fig. 22C–D +). As in male: dorsal and lateral surfaces of cephalothorax, abdomen and legs unicolorous grey-brown; venter of sternum, mouthparts, coxae and trochanters yellow white. + + +STERNUM ( +Fig. 22D +). Oval with slightly sinuous sides; 4.90 wide, 6.40 long. + +CHILUM. 0.25 high, 1.08 wide. +EYES. ALE 0.33; AME 0.33; PLE 0.31; PME 0.29; ALE–AME 0.20; AME–AME 0.75; PLE–PME 0.55; PME–PME 0.35; MOQ: AW 0.78, PW 0.93, L 0.78. +CLYPEUS. With ten long setae. +LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apex of metatarsus III, apical half of metatarsus II and all of metatarsus I. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I10.305.2011.5011.106.3044.20
II9.504.907.206.803.8032.30
III8.204.205.807.203.8029.20
IV10.904.709.0012.604.8042.00
+
+ +Spination GENITALIA. Epigyne ( +paratype +CASENT_9002829; CAS) ( +Fig. 23D +) transverse quadrangular median lobe (ML) plate with smooth margin; copulatory openings (CO) hidden beneath anterolateral corners of ML; lateral lobes (LL) convex, without lateral teeth or projections but with concavities along lateral margins of ML, LL straight and parallel medially, exposing rectangular area of median sector (MS). Endogyne ( +Fig. 23E +) with broad, simple, dark copulatory ducts (CD) larger than spermathecae (S), spermathecae with one half twist; fertilization ducts (FD) short and straight. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d2v2-2-2-2v2-2-2
IIpl2d2v2-2-2-2v2-2-2
IIIpl3d3rl4pl1rl1pl2d2rl2v2-2-2pl4d1rl4v2-2-2
IVpl1d3rl3pl1rl1pl2d2rl2v2-2-2pl5rl4v2-1-2-2
+
+ + +Fig. 22. + +Zorodictyna almae + +sp. nov. +, male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). +A, C +. Dorsal view. +B, D +. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme. + + + + +Fig. 23. + +Zorodictyna almae + +sp. nov. +, genitalia drawings; A–C += +male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS). +A +. Left palp, prolateral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Epigyne, ventral view. +E +. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold. + + + + +Variation + + + +Male + + +Variation unknown: only the single +holotype +has been studied. + + +Female +(n = 3) + +[Female variation from CASENT_9082245, CASENT_9031270 and CASENT_9031271]. Total length = 20.00–30.00, carapace length/width = 1.24–1.30, carapace height/width = 0.39–0.48, PER/carapace width = 0.21–0.24, PER/OAL = 2.90–3.13, PER/AER = 1.35–1.36, OQP/OQA = 1.18–1.23, sternum length/width = 1.20–1.31, palpal coxa length/width = 2.24–2.44, femur I length/carapace width = 0.87– 1.00, metatarsus I length/carapace width = 0.64–0.74, femur IV length/carapace width = 0.96–1.06, palpal tarsus length/carapace width = 0.34–0.41, palpal tarsus length/palpal patella length = 1.46–2.00, palpal tarsus length/palpal tibia length = 1.09–1.57, palpal tarsus length/palpal femur length = 0.76–0.95, palpal tibia length/palpal patella length = 1.08–1.40. + + +Note on synonym + + + +A specimen of this species (CASENT_9031271) was used as an exemplar in a previously published molecular phylogeny, listed as “ + +Zorodictyna +sp. 1 + +” by + +Polotow +et al. +(2015) + +. + + + +Natural history + + + +The species + +Zorodictyna almae + +sp. nov. +is recorded from tropical dry forest.Although they have an entire cribellum we know nothing of their use of cribellate silk. + +
+ + + +Distribution + + + + +Zorodictyna almae + +sp. nov. +is known from National Park Ankarafantsika in Mahajunga Province in northwestern +Madagascar +( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFD4FFE4FD87092DFA8BDCCE.xml b/data/31/57/E0/3157E00AFFD4FFE4FD87092DFA8BDCCE.xml new file mode 100644 index 00000000000..766930f20b1 --- /dev/null +++ b/data/31/57/E0/3157E00AFFD4FFE4FD87092DFA8BDCCE.xml @@ -0,0 +1,387 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Zorascar pasunepomme + +gen. et sp. nov. + + + + +urn:lsid:zoobank.org:act: +CC2504B1-3B59-4DE9-9C27-0457AAE53102 + + + + + +Figs 16E +, +17–21 +, +26 + + + + + +Diagnosis + + + +The female of + +Zorascar pasunepomme + +gen. et sp. nov. +is characterized by the epigyne with a median lobe (ML) that is narrow and deeply concave medially and copulatory openings (CO) exposed laterad of median lobe ( +Figs 16E +, +18A +, +19A +). + + + + + +Etymology + + +The species name, a noun in apposition, refers to the famous painting of René Magritte depicting an apple, named ‘Ceci n’est pas une pomme’, to which the genitalia appear to be reminiscent. + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Antsiranana Province + +• + +; +Bamevika +; +14°18′57.312″ S +, +48°43′26.180″ E +; + +1581 m +a.s.l. + +; + +2 May 2019 + +; +L. Kemp +leg.; hand collected, grassland; DNA RJ4; BINCO_MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247346; +RMCA +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Antsiranana Province + +• +1 juv. +; same data as for holotype; [SEM imaged]; BINCO_MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247347; +RMCA + +• + +1 juv. +; Bamevika; +14°18′58.9″ S +, +48°43′22.9″ E +; + +1557 m +a.s.l. + +; +M. Jocque +leg.; hand collected, forest leaf litter; DNA RJ2; BINCO_ MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247349; +RMCA + +• + +1 juv. +; same data as for preceding; DNA RJ3; BINCO_MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247348; +RMCA + +. + + + + + +Description + + + + +Female +holotype + +(BE_RMCA_ARA.Ara.247346; RMCA) + +MEASUREMENTS. Total length: 6.82. Carapace: length 2.91, width 2.30, height 1.42. + +COLOUR ( +Figs 17 +). Carapace brownish yellow, with two contrasting dark bands of short setae along margin and faint median band between PME and fovea; chelicerae brownish, anteriorly with two thin longitudinal darker stripes; sternum yellowish brown; legs brownish yellow with three darker rings on femora, two on tibiae, distally and proximally; abdomen brownish yellow with spotted pattern on dorsum and sides; spinnerets pale yellow. + + +STERNUM ( +Fig. 17B +). Oval with slightly sinuous sides; 1.21 wide, 1.42 long. + +CHILUM. 0.12 high, 0.20 wide. +EYES. ALE 0.16; AME 0.12; PLE 0.16; PME 0.16; ALE–AME 0.05; AME–AME 0.07; PLE–PME 0.16; PME–PME 0.10; MOQ: AW 0.28, PW 0.39, L 0.36. +CLYPEUS. With five long setae. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I2.101.051.821.611.007.58
II1.680.911.471.400.806.26
III1.750.841.331.610.706.23
IV2.310.912.032.661.209.11
+
+ + +Fig. 17. + +Zorascar pasunepomme + +gen. et sp. nov. +, holotype female (BE_RMCA_ARA.Ara.247346; RMCA), habitus. +A +. Dorsal view. +B +. Ventral view. +C +. Lateral view. Scale bars = 1 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué. + + + +Spination GENITALIA. Epigyne ( +Figs 16E +, +18A +, +19A–B +) with median lobe (ML) narrow and deeply concave medially (MS), copulatory openings (CO) exposed laterad of ML; lateral lobe (LL) with pair of stout basal, blunt, conical projections (LP); spermathecae dark, comma shaped as in transparency. Endogyne ( +Figs 18B–D +, +19C–F +): sperm ducts (CD) short, cylindrical, with three twists; spermathecae (S) reniform, with fairly narrow ventral base, and dorsal, roughly semicircular part, provided with few pores; fertilization ducts (FD) short, flattened and slightly curved. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d1v1v2-2-2-2v2-2-2
IIpl2d1pl3v2-2-2-2v2-2-2
IIIpl3d3rl2pl1rl1pl2d2rl2v2-2-2pl3d1rl3v2-2-2
IVpl3d2rl1pl1rl1pl2d2rl2v2-2-2disp14
+
+ +Male + +Unknown. +
+ + + +Distribution + + + +Known only from the +type +locality in northern +Madagascar +at an altitude of around +1500 m +( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFDBFFEFFD2B0A74FADCDE5C.xml b/data/31/57/E0/3157E00AFFDBFFEFFD2B0A74FADCDE5C.xml new file mode 100644 index 00000000000..70d778f41ee --- /dev/null +++ b/data/31/57/E0/3157E00AFFDBFFEFFD2B0A74FADCDE5C.xml @@ -0,0 +1,396 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + +Genus + +Zorascar + +gen. nov. + + + + +urn:lsid:zoobank.org:act: +F5981AD2-4EC8-412D-B90A-0F6A51FEF784 + + + + + +Figs 13–21 +, +26 + + + + + + +Type +species + + + + + +Zorascar pasunepipe + +gen. et sp. nov. + + + + + +Diagnosis + + + + +Zorascar + +gen. nov. +are ecribellate udubids distinguished from all +Madagascar +udubids (except + +Tabiboka + +gen. nov. +) by having the posterior spinnerets (PMS and PLS) reduced in size, with the PLS less than one half the size of ALS; the male palp ( +Figs 14A–C +; +15A–C +, +16A–B +) with an RTA only, lacking the VTA, the cymbial retrobasal extension (CE) elongate, extending most of length of tibia, without retrolateral paracymbial process; the female epigyne with short median lobe (ML), concave medially and not extending to posterior margin of epigyne, with copulatory openings (CO) exposed ( +Fig. 16C, E +). + +Zorascar + +is further distinguished from + +Uduba +Simon, 1880 + +males by lacking a threadlike embolus and a knob-like process between palpal patella and tibia and from + +Uduba + +females by having compact spermathecae rather than multiple loops. It is distinguished from + +Zorodictyna + +by lacking an entire cribellum or vestige and in lacking extensive scopulae beneath leg metatarsi. + +Zorascar + +may be distinguished from + +Tabiboka + +gen. nov. +by having the male palp with cymbial retrobasal extension (CE) elongate ( + +Tabiboka + +shorter), palpal lacking a VTA ( +Figs 14A–B +, +15A–B +, +16A–B +) ( + +Tabiboka + +with a VTA); female epigyne median lobe (ML) short, concave medially, not extending to posterior margin, copulatory openings (CO) covered ( +Figs 16C, E +, +18A +). + + + + + +Etymology + + + +The genus name is derived from the name ʻ + +Zorodictyna + +ʼ, in combination with the ending of +Madagascar +. The gender is masculine. + + + + + +Description + + + +BODY. Medium size spiders (males: 7.81–8.60; females: 8.50–10.10) with smooth teguments of carapace ( +Figs 13A–D +, +17A, C +), sternum and legs. Carapace longer than wide (L/W: males; 1.21–1.34; females: 1.33–1.34) with sparse cover of dark setae; widest at level of coxae II, narrowed to 0.56 times maximum width in male and 0.70 times maximum width in female (cephalic width measured on posterior tangent of PME); profile convex, highest point located at well-developed fovea. + + +COLORATION ( +Figs 13A–E +, +17A–C +). Sclerites brownish yellow with darker longitudinal pattern on carapace and chelicerae; legs ringed; abdomen brownish yellow with faint complex pattern. + + +EYES ( +Figs 13C–D +, +17A +). In two straight rows: ALE larger than AME, PLE and PME similar to ALE; tapetum of secondary eyes canoe shaped. Clypeus straight, narrow, with long setae. + + +PROSOMA. Chilum double, about twice as wide as high, lateral margin faintly delimited. Chelicerae ( +Figs 13A–E +, +17A–B +): paturon conical with some long setae in center and two faint longitudinal stripes; three teeth on both margins; fangs smooth. Labium ( +Figs 13E +, +17B +) roughly rectangular. Endites ( +Figs 13E +, +17B +, +20D +) roughly rectangular, with retrolateral margins sinuous, distal part slightly wider than proximal part, with distal serrula present, as simple row ( +Fig. 20D +). Sternum ( +Figs 13E +, +17B +) oval, slightly longer than wide, with slightly sinuous margins, and with tiny triangular extensions or precoxal sclerites, not extending between coxae IV. + + +LEGS ( +Figs 13A–B, E +, +17A–C +, +20E–F +). Slender. Formula 4123 or 4132. With three tarsal claws, unpaired claw smooth, rudimentary on legs III and IV, superior claws with six to eight teeth ( +Figs 21A–B +). Without claw tufts, tenent setae or scopulae. Numerous spines, pairs of ventral spines on metatarsus overlapping. Trichobothria on tarsi in three rows, starting close to tip, length variable but not increasing ( +Figs 13F +, +20E–F +); proximal plate of bothria with six ridges ( +Figs 21D–E +). Tarsal organ with circular opening provided with two to three anterior slits, in distal half of tarsus ( +Figs 20E +, +21C–D +). + + +ABDOMEN ( +Figs 13A–B, E +, +17A–C +). Oval, without scuta and cribellum. Colulus: as hairy field. Spinnerets ( +Figs 20A–C +): ALS large, conical, biarticulate, in juvenile provided with two major ampullate and four piriform gland spigots ( +Fig. 20A +); PLS smaller, with three spigots ( +Fig. 20B +); PMS reduced, with three spigots ( +Fig. 20C +). Adult spinnerets much as in + +Tabiboka + +gen. nov. +ALS and PLS, conical, biarticulate; PMS one-segmented; ALS large, PMS and PLS small. Females with two ALS major ampullate gland spigots and field of piriform gland spigots and tartipores; PMS with at least one anterior minor ampullate gland spigot and one to few posterior cylindrical gland spigots and aciniform gland spigots; PLS with one to few cylindrical gland spigots; males lack cylindrical gland spigots and have second major ampullate gland spigot reduced to nubbin. + + +MALE PALP ( +Figs 14–15 +, +16A–B +). Tibia with short RTA inserted medially and sharp triangular prolateral process ( +Figs 14C +, +15C +), without VTA at retroapical margin; cymbium oval with rounded distal extremity, without dorsal patch of short setae, with large pipe shaped posterior extension (CE); bulbus provided with very large retrobasal subtegulum lobe (SL), visible over one third of bulbus, and tegulum rounded with well developed prolateral conical protrusion (corresponding locking lobe, TL) ( +Figs 14E–F +, +15B–C +, +16A +); embolus (E) with large base inserted in center of tegulum, short, conical and folded ( +Figs 14D–F +, +15E–F +); median apophysis (MA) situated at base of E, short, sclerotized with membranous base; conductor (C) relatively small, transverse, hyaline fan-shaped, prolaterally extending from narrow base to wide flattened tip ( +Figs 14D, F +, +15D–F +); paraconductor (Pc) as narrow membranous extension inserted between tegulum and base of MA, running backward to MA and ending with fattened fan-shaped tip accompanying E ( +Figs 14D +, +15D–E +, +16A–B +). + + +GENITALIA. Epigyne ( +Figs 16C, E +, +18A +, +19A–B +) with median lobe (ML) as single plate with central roughly triangular depression, delimited by margins forming narrow, elongate arch, lateral lobe (LL) basally with two short but stout, conical and blunt projections (LP). Endogyne ( +Figs 16D +, +18B–D +, +19C–F +) with copulatory ducts (CD) very short, twisted; spermathecae (S) with narrow base and broad roughly reniform dorsal part; no evidence of large Bennett’s Gland pore; fertilization ducts (FD) short, median. + + + +Species included + + + + +Zorascar pasunepipe + +gen. et sp. nov. +( +type +species; + + +); + +Zorascar pasunepomme + +gen. et sp. nov. +( + +). + + + +Affinities + + + +From the molecular analyses ( +Fig. 2 +), + +Zorascar + +nov. gen. +appears closely related to species in the +Udubidae +that have been identified as + +Tabiboka + +gen. nov. + + + + + +Distribution + + + + +Zorascar + +gen. nov. +is known only from highland forest in northwestern +Madagascar +( +Fig. 26 +). + + + + \ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFDDFFE1FD9509B5FEFCDE7A.xml b/data/31/57/E0/3157E00AFFDDFFE1FD9509B5FEFCDE7A.xml new file mode 100644 index 00000000000..bf903f138df --- /dev/null +++ b/data/31/57/E0/3157E00AFFDDFFE1FD9509B5FEFCDE7A.xml @@ -0,0 +1,675 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Zorascar pasunepipe + +gen. et sp. nov. + + + + +urn:lsid:zoobank.org:act: +1C01B17C-DC6C-4BA5-942B-77F90554F3A0 + + + + + +Figs 13–15 +, +16A–D +, +26 + + + + + +Diagnosis + + + +The male of + +Zorascar pasunepipe + +gen. et sp. nov. +is easily recognized by the presence of an enormous and tortuous cymbial retrobasal extension (CE), extending most of the tibial length ( +Figs 14A–C +, +15A–B +, +16A–B +). The female is provided with an epigyne ( +Fig. 16C +) with the median lobe (ML) that is broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially (vs the female epigyne of + +Z. pasunepomme + +gen. et sp. nov. +has a ML that is narrow and deeply concave medially, and with CO exposed laterad of the ML, +Fig. 16E +). + + + + + +Etymology + + + +The species name, a noun in apposition, refers to the famous painting of René Magritte depicting a pipe, named ‘Ceci n’est pas une pipe’, to which the cymbial extension is reminiscent, at least according the artist who made the drawings of + +Zorascar + +gen. nov. +for the present paper. + + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Antsiranana Province + +• + +; +Bamevika +, +Andriakanala Forest +; +14°18′58.932″ S +, +48°43′22.886″ E +; + +1586 m +a.s.l. + +; + +6–11 Feb. 2019 + +; +M. Jocque +leg.; hand collected around camp; DNA RJ5; BINCO_MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247343; +RMCA +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Antsiranana Province + +• +1 juv. +; same data as for holotype; DNA RJ1; BINCO_ MAD_19_0079_2; BE_ +RMCA +_ARA. +Ara. +247344; +RMCA + +• + +1 ♂ +, +1 ♀ +; +Réserve Spéciale Manongarivo +, + +10.8 km +229° SW of Antanambao + +; +13°58′ S +, +48°26′ E +; + +400 m +a.s.l. + +; + +8 Nov. 1998 + +; +B.L. Fisher +leg.; rainforest; EC19 sifted litter; [SEM stub 30:4]; CASENT_9016138; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +– + +Antsiranana Province + +• +1 ♂ +; +Réserve Spéciale Manongarivo +, + +10.8 km +229° SW of Antanambao + +, +13°58′ S +, +48°26′ E +; + +400 m +a.s.l. + +; + +8 Nov. 1998 + +; +B.L. Fisher +leg.; rainforest; EC19 sifted litter; [male palp drawn; male stacked images]; CASENT_9016135; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9016136; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9016137; +CAS + +• + +1 ♀ +; +Réserve Spéciale Manongarivo +, + +14.5 km +220° SW of Antanambao + +; +13°60′ S +, +48°26′ E +; + +1175 m +a.s.l. + +; + +20 Oct. 1998 + +; +B.L. Fisher +leg.; rainforest; EB09 sifted litter; BLF1938; CASENT_9016139; +CAS + +• + +1 ♀ +; same data as for preceding; CASENT_9016141; +CAS + +. + + + + + +Description + + + + +Male +holotype + +(BE_RMCA_ARA.Ara.247343; RMCA) + +MEASUREMENTS. Total length: 7.81. Carapace: length 3.91, width 2.91, height 1.85. + +COLORATION ( +Figs 13A, C, E +). Carapace brownish yellow, with two faint, longitudinal dark bands along margin and between PME and fovea; chelicerae brownish with two faint longitudinal darker stripes; sternum brownish yellow, with faint radiating stripes; legs brownish yellow with three faint darker rings on femora and two on tibias, distally and proximally; abdomen brownish yellow with ill-defined pattern on dorsum and sides; spinnerets pale yellow. + + + +Fig. 13. + +Zorascar pasunepipe + +gen. et sp. nov. +; A, C, E–F += +holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D += +paratype female (CASENT_9016138; CAS). +A–B +. Habitus, dorsal view. +C–D +. Prosoma, dorsal view. +E +. Habitus, ventral view. +F +. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood. + + + +STERNUM ( +Fig. 13E +) oval with slightly sinuous margins; 1.49 wide, 1.70 long. + + + +Fig. 14. + +Zorascar pasunepipe + +gen. et sp. nov. +, holotype male (BE_RMCA_ARA.Ara.247343; RMCA). +A +. Male right palp (mirrored), retrolateral view. +B +. Idem, ventral view. +C +. Idem, dorsal view. +D +. Idem, bulbus, retrolateral view. +E +. Idem, ventral view. +F +. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5mm; D–F = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué. + + +CHILUM. 0.12 high, 0.20 wide. +EYES. ALE 0.18; AME 0.15; PLE 0.18; PME 0.16; ALE–AME 0.05; AME–AME 0.08; PLE–PME 0.18; PME–PME 0.10; MOQ: AW 0.33, PW 0.44, L 0.44. + + +Fig. 15. + +Zorascar pasunepipe + +gen. et sp. nov. +, holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). +A +. Retrolateral view. +B +. Ventral view. +C +. Prolateral view. +D +. Bulbus, retrolateral view. +E +. Bulbus, ventral view. +F +. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5 mm; D–F = 0.2 mm. 3D micro-CT scans by Arnaud Henrard. + + +CLYPEUS. With 14 long setae. +LEGS. Trochanters shallowly notched. Unpaired claws smooth, rudimentary on legs III and IV. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I3.361.333.152.731.0511.62
II3.151.262.102.731.3310.57
III2.450.841.962.731.059.03
IV3.501.263.084.271.4013.51
+
+ +Spination MALE PALP ( +Figs 14–15 +, +16A–B +). RTA short, emerging in center of tibia, with cylindrical base and sharp triangular apex; tibia with dorso-retrolateral excavation bordered by median row of long setae and dorsal margin of RTA, accommodating very large, pipe-shaped retrobasal cymbial extension (CE); cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setaeor retrobasal paracymbial process; subtegulum large, filling posterior third of cymbial concavity, with basal ridge (SL) interacting with the tegulum; tegulum rounded with massive, pointed, prolateral protuberance (TL); embolus (E) inserted on center of tegulum, short, stout at base, with medial groove, lamellar apically and curved retrolaterad; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) translucent, flattened, fan shaped, narrow at base and strongly widened at extremity; paraconductor (Pc) pedicel-like membranous extension originating at base of MA, running backward along MA and ending with flattened fan-shaped tip embracing E. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl2d3rl3pl1pl2d2rl2v2-2-2-2pl2rl2v2-2
IIpl3d3rl4pl1pl2d3rl3v2-2-2-2pl2rl2v2-2-1
IIIpl3d3rl4pl1rl1pl2d2rl2v2-2-2pl3d1rl3v2-2-2
IVpl4d3rl1pl1rl1pl2d2rl2v2-2-2disp14
+
+ +Female +(CASENT_9016138; CAS) + +MEASUREMENTS. Total length: 8.50. Carapace: length 4.80, width 3.60, height 1.70. + +COLOUR ( +Figs 13B, D +). Carapace brownish yellow, dark along margin and with two contrasting dark bands along margin of pars cephalica, short longitudinal line behind each PME and longer faint median band between PME and fovea; chelicerae brownish-yellow, anteriorly with two thin longitudinal darker stripes; sternum yellowish white with faint median dusky oval mark; legs brownish yellow with three darker rings basally, medially and apically on femur and two basally and apically on tibia; abdomen yellow-white with spotted pattern on dorsum and sides, venter with two broken dusky longitudinal bands and dark surrounding spinnerets, spinnerets pale yellow. + +STERNUM. Oval with slightly sinuous sides; 2.10 wide, 2.20 long. +CHILUM. 0.15 high, 0.20 wide. +EYES. ALE 0.13; AME 0.09; PLE 0.16; PME 0.15; ALE–AME 0.08; AME–AME 0.25; PLE–PME 0.18; PME–PME 0.10; MOQ: AW 0.26, PW 0.43, L 0.35. +CLYPEUS. With eight long setae. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I3.401.603.302.601.8012.70
II3.101.402.602.501.6011.20
III3.001.302.102.701.6010.70
IV4.001.403.602.901.6013.50
+
+ +Spination GENITALIA. Epigyne ( +Fig. 16C +) with median lobe (ML) broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially; lateral lobes (LL) rounded and becoming narrowed to a blunt point medially over median sector (MS); lateral processes (LP) small, blunt. Endogyne ( +Fig. 16D +) with narrow, curved copulatory ducts (CD), these hidden beneath the spermathecal heads (S); fertilization ducts (FD) posterior. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d12v2-2-2-2v2-2-2
IId1v2-2-2-2v2-2-2
IIIpl1d2rl1pl2d2rl2v2-2-2pl4rl4v2-2-2
IVpl1d2pl2d2rl2v2-2-2pl4rl4v2-1-2-2
+
+ + +Variation + + + +Male +(n = 5) + + +Total length = 7.81–8.60, carapace length/width = 1.21–1.34, carapace height/width = 0.46–0.56, PER/carapace width = 0.23–0.24, PER/OAL = 2.28–2.44, PER/AER = 1.40–1.44, OAL/ OQL = 1.14–1.25, OQP/OQA = 1.00–1.52, sternum length/width = 1.07–1.16, palpal coxa length/width = 2.00–2.50, femur I length/carapace width = 1.05–1.13, metatarsus I length/carapace width = 0.87–1.12, femur IV length/carapace width = 1.17–1.24, cymbium length/carapace width = 0.39–0.46, cymbium length/palpal patella length = 2.00–2.57, cymbium length/palpal tibia length = 1.60–2.29, cymbium length/palpal femur length = 0.84–1.00, palpal tibia length/palpal patella length 1.00–1.43. Male palp of the +holotype +from Andriakanala Forest and of specimens from Manongarivo. i.e.; CASENT_9016135–138 and CASENT_9016141 are very similar but differ in a few characteristics. The the +holotype +has the RTA more narrowly pointed, the cymbial retrobasal extension longer and more twisted, the tegular prolateral locking process more narrowly triangular and the apex of the MA sharper than those features in males from Manongarivo. + + +Female +(n = 2) + +Total length = 8.50–10.10, carapace length/width = 1.33–1.34, carapace height/ width = 0.47–0.58, PER/AER = 1.37–1.39, OQP/OQA = 1.48–1.62, sternum length/width = 1.05–1.12, palpal coxa length/width = 2.50–2.55, femur I length/carapace width = 0.94–0.97, metatarsus I length/carapace width = 0.72–0.76, palpal tarsus length/carapace width = 0.34–0.36, palpal tarsus length/palpal tibia length = 1.44–2.17, palpal tarsus length/palpal tibia length = 1.30–1.44, palpal tibia length/palpal patella length = 1.11–1.50. Epigyne with median lobe length 1.80–3.33 times width; distance from atrium base to epigastric fold 0.19–0.26 times epigyne length. + + +Natural history + + + +This species has been collected in forest from +400 m +to +1600 m +a.s.l. The function of the remarkable retrobasal cymbial extension on the male palp is unknown. + +
+ + + +Distribution + + + + +Zorascar pasunepipe + +gen. et sp. nov. +is known only from +Antsiranana Province +in northwestern +Madagascar +( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFE5FFE9FDEC092CFEAEDD1E.xml b/data/31/57/E0/3157E00AFFE5FFE9FDEC092CFEAEDD1E.xml new file mode 100644 index 00000000000..756ad3f3636 --- /dev/null +++ b/data/31/57/E0/3157E00AFFE5FFE9FDEC092CFEAEDD1E.xml @@ -0,0 +1,830 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Tabiboka polotowae + +gen. et sp. nov. + + + + +urn:lsid:zoobank.org:act: +5FF0F818-0EC8-428C-86F4-36FF94F62ED7 + + + + + +Figs 10–12 +, +26 + + + +Zorodictyna +spp. 3 + +and 4 – + +Polotow +et al. +2015 + +. + + + + + +Diagnosis + + + +The males of + +Tabiboka polotowae + +gen. et sp. nov. +may be distinguished by an elongate embolus (E) arising retrobasally near 200°, the paraconductor (Pc) visible between embolus and the conductor (C) base, the palpal tibia having blunt retromedian process (RTA) arising near basally and narrow, finger-like ventral apical process (VTA) ( +Figs 11A–C +, +12A–F +) (vs in + +T. milleri + +gen. et sp. nov. +, males have shorter and broader E and MA, a small VTA and greatly elongated RTA, +Fig. 5A–C +; and in + +T. milloti + +gen. et sp. nov. +, the MA is elongate, cruciform, and the VTA is thumb-like, +Fig. 9A–C +). Females of + +T. polotowae + +have epigyne median lobe (ML) rectangular, narrowest in middle or with sides straight ( +Figs 11D +, +12G +) (vs heart-shaped ML in + +T. milleri + +, +Figs 4B +, +5D +, + +7A and + +T + + +. milloti, +Fig. 9D +). + + + + + +Etymology + + + +The species name is a matronym in honor of Dr Daniele Polotow, who studies the phylogenetics and taxonomy of spiders of the OC clade including Lycosoidea. She led the research that in 2015 led to the proposal and diagnosis of the +Udubidae +and the current phylogenetic framework for Lycosoidea interrelationships, and she collected many of the data used in this paper. + + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Antananarivo Province + +• + +; NE outskirts of +Antananarivo +, +Ambohimanga Village +; +18°44′ S +, +47°34′ E +(coordinates approximate); + +27 Jul. 1992 + +; +V. and B. Roth +leg.; VBRoth002; [palp & endogyne drawn; SEM palp]; CASENT_9016117; +CAS +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Antananarivo Province + +• +1 ♀ +; same data as for holotype; CASENT_9016117; +CAS + +• + +1 ♀ +; same data as for preceding; + +1 Nov. 1993 + +; +J. Coddington +, +N. Scharff +, +S. Larcher +, +C. Griswold +, +R. Andriamasimanana +leg.; +Berlese +of fine leaf litter; CEG010; CASENT_9016116; +CAS + +• + +1 ♂ +; + +10 km +NE of Tana + +; +18°45′32″ S +, +48°33′45″ E +; + +1300 m +a.s.l. + +; + +5 August 1992 + +; +V. Roth +, leg.; [habitus photos of male]; +Roth +001; CASENT_9016118; +CAS + +. + + + +Fianarantsoa Province + +• +1 ♀ +; +Parc National Andringitra +, + +34 km +S of Ambalavao + +; +22°08′48.9″ S +, +45°57′03.4″ E +; + +1580 m +a.s.l. + +; + +7 Jan. 2009 + +; +H. Wood +leg.; sifting litter in the day; primary montane rainforest; [SEM stubs 221, 9, 10, 222, 4] [ +Voucher +DP-2014b -“ +Zorodictynidae +sp. 3”, +D. Polotow Lycosoidea Total Evidence Phylogeny +]; HW0813; CASENT_9029890; +CAS + +• + +1 ♀ +; same data as for preceding; [ +Voucher +DP-2014a -“ +Zorodictynidae +sp. 4”, +D. Polotow Lycosoidea Total Evidence Phylogeny +]; CASENT_9029889; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +– + +Antananarivo Province + +• +1 ♀ +; +Station Forestière Angavokely +, + +22 km +E of +Antananarivo + +; +18°56′ S +, +47°45′ E +; + +1300 m +a.s.l. + +; low canopy remnant forest, general collecting; + +8 Sep. 2001 + +; +D. Ubick +, +D. Andriamalala +, +T.Andriambinintsoa +, +J.J. Rafanomezantsoa +leg.; DU-01-2; CASENT_901371; +CAS + +• + +1 ♂ +, +1 ♀ +; same data as for preceding; CASENT_901317; +CAS + +• + +1 ♂ +; +Ambohimanga +; +18°44′ S +, +47°34′ E +; + +1400 m +a.s.l. + +; + +2 Nov. 1993 + +; +C. Griswold +, +S. Larcher +, +N. Scharff +leg.; night collecting; CEG009; CASENT_9016115; +USNM + +• + +1 ♂ +; + +10 km +NE of Tana + +; +18°45′32″ S +, +48°33′45″ E +; + +1300 m +a.s.l. + +; + +5 Aug. 1992 + +; +V. Roth +, leg.; [habitus photos of male]; +Roth +001; CASENT_9016118; +CAS + +. + + + + + +Description + + + + +Male +holotype + +(CASENT_9016117; CAS) + +MEASUREMENTS. Total length: 8.10. Carapace: length 4.50, width 3.40, height 1.90. + +COLORATION ( +Fig. 10A, C +). Carapace brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black on ocular area and surrounding all eyes; chelicerae brownish yellow; sternum yellow, unmarked; legs brownish yellow, with subbasal and subapical dusky annuli on all femora, other segments unmarked; abdomen brownish yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides brownish yellow mottled with dark marks, venter yellow except darker around spinneret base. + + + +Fig. 10. + +Tabiboka polotowae + +gen. et sp. nov. +, male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). +A–B +. Dorsal view. +C–D +. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme. + + + +STERNUM ( +Fig. 10C +). Oval with gently curved margins; 1.70 wide, 2.20 long. + + + +Fig. 11. + +Tabiboka polotowae + +gen. et sp. nov. +, genitalia drawings; A–C += +male holotype (CASENT_9016117; CAS); D–E += +female paratype (CASENT_9016117; CAS). +A +. Left palp, prolateral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Epigyne, ventral view. +E +. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels. + + + + +Fig. 12. + +Tabiboka polotowae + +gen. et sp. nov. +, SEM of male and female genitalia; A–F += +male holotype (CASENT_9016117; CAS); G–H += +female paratype (CASENT_9029889; CAS). +A +. Right palp (mirrored), prolateral, slightly ventral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Idem, dorsal view. +E +. Bulb, ventral view. +F +. RTA, ventral view. +G +. Epigyne, ventral view. +H +. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow. + + +CHILUM. 0.08 high, 0.28 wide. +EYES. ALE 0.11; AME 0.09; PLE 0.13; PME 0.13; ALE–AME 0.05; AME–AME 0.20; PLE–PME 0.33; PME–PME 0.13; MOQ: AW 0.23, PW 0.33, L 0.28. +CLYPEUS. 0.18 high, with four long setae. +LEGS. Trochanters shallowly notched. Unpaired claw smooth, well developed on all legs. Metatarsi lacking subbasal crack. Scopulae absent. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I3.051.503.202.801.8012.35
II3.201.502.502.301.6011.10
III3.001.302.102.601.5010.50
IV3.901.402.304.302.0013.90
+
+ +Spination MALE PALP ( +Figs 11A–C +, +12A–F +). Tibia with narrow finger-shaped VTA at retroapical margin and broad, median RTA arising near tibia base; cymbium oval, without dorsal patch of chemosensitive setae, with broad, sharp, conical retrobasal cymbial extension (CE) that extends to tibial apex; subtegulum large, filling most of cymbial concavity, with basal ridge (SL) at 180° that interacts with tegulum; tegulum oval with broad, basal notch, beneath this, conical lobe (TL) at 180° that locks with subtegulum; embolus (E) arises from retromargin of tegulum, simple, narrow, elongate, extending past apex of tegulum; median apophysis (MA) as sclerotized hook arising from soft base; conductor (C) hyaline, translucent, with narrow base arising near E base and greatly widened at extremity; paraconductor (Pc) as finger-like membranous structure originating between bases of E and C and extending less than half length of E. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d3pl2d1v2-2-2-2pl2rl2v2-2-2
IIpl1d3rl1pl2v2-2-2-2pl3d1rl2v2-2-2
IIIpl2d2rl3pl1rl1pl2d2rl2v2-2-2pl4rl3v2-2-2
IVpl2d3rl1rl1pl2d2rl2v2-2-2pl3rl4v1-1-2-2-2
+
+ + +Female +paratype + +(CASENT_9016116; CAS) + +MEASUREMENTS. Total length: 11.50. Carapace: length 5.20, width 3.60, height 2.00. + +COLOUR ( +Fig. 10B, D +). As in male except ocular area not all black and legs only faintly marked. + + +STERNUM ( +Fig. 10D +). Oval with slightly sinuous margins; 2.10 wide, 2.40 long. + +CHILUM. 0.09 high, 0.33 wide. +EYES. ALE 0.14; AME 0.08; PLE 0.15; PME 0.13; ALE–AME 0.06; AME–AME 0.25; PLE–PME 0.20; PME–PME 0.09; MOQ: AW 0.28, PW 0.38, L 0.28. +CLYPEUS. 0.10 high, with four long setae. +LEGS. Very weak scopula beneath tarsus I, scopulae otherwise absent. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I3.301.602.702.201.6011.40
II3.051.702.302.251.4010.70
III2.801.302.002.301.509.90
IV3.901.603.004.002.0014.50
+
+ +Spination GENITALIA. Epigyne ( +Figs 11D +, +12G +) median lobe (ML) rectangular, elongate, narrowest in middle with apex of ML extending beyond epigastric furrow; lateral lobe (LL) convex, with transverse curved concavity posteriorly, narrow, only 0.25 maximum width of ML, copulatory openings (CO) hidden beneath anterolateral margins of ML. Endogyne ( +Figs 11E +, +12H +): spermathecae (S) simple, oval, length 1.06 times width, with scattered pores anteriorly on spermathecal head. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d1v2-2-2-2v2-2-2
IIpl1d1v2-2-2v2-2-2
IIIpl2d3rl1pl2d1rl2v2-2-2pl5rl3v2-2-3
IVpl1d2rl1pl1d2v1-2-2pl4d2rl1v1-1-2-2-3
+
+ + +Variation + + + +Male +(n = 2) + +Total length = 8.10–8.50, carapace length/width = 1.32–1.34, carapace height/width = 0.40–0.56, PER/OAL = 2.21–2.22, PER/AER = 1.33–1.38, OAL/OQL = 1.22–1.23, OQP/OQA = 1.44–2.00, clypeus height/AME = 1.00–2.33, cheliceral length/clypeus height = 8.57–17.00, sternum length/width = 1.05–1.29, palpal coxa length/width = 2.08–2.17, femur I length/carapace width = 0.90–1.06, metatarsus I length/carapace width = 0.82–0.86, cymbium length/carapace width = 0.37–0.40, cymbium length/palpal patella length = 2.15–2.27, cymbium length/palpal tibia length = 1.75–2.27, cymbium length/palpal femur length = 0.78–1.00, palpal tibia length/palpal patella length = 1.00–1.23. + +Female +(n = 3) + + +Total length = 8.40–17.00, carapace length/width = 1.31–1.46, carapace height/ width = 0.44–0.75, PER/carapace width = 0.25–0.27, PER/OAL = 2.40–2.88, PER/AER = 1.31–1.45, OAL/OQL = 1.20– 1.36, OQP/OQA= 1.36–1.56, clypeus height/AME = 1.33–2.33, cheliceral length/clypeus height = 10.67– 20.00, sternum length/width = 1.14–1.22, palpal coxa length/width = 1.57–2.26, femur I length/carapace width = 0.89–1.32, metatarsus I length/carapace width = 0.61–0.64, femur IV length/carapace width = 1.00–1.18, palpal tarsus length/carapace width = 0.33–0.38, palpal tarsus length/palpal tibia length = 1.91–2.15, palpal tarsus length/palpal tibia length = 1.33–1.75, palpal tarsus length/palpal femur length = 0.88–0.89, palpal tibia length/palpal patella length = 1.09–1.23. Epigyne with median lobe length 1.29– 1.38 times width; median lobe maximum width 1.34–2.10 times minimum width; median lobe width 2.00–5.00 times lateral lobe width; median lobe extends to ( +Fig. 12G +) or beyond ( +Fig. 11D +) epigastric furrow. Spermatheca (each) length 1.06 times width. + + + +Note on synonyms + + + +Specimens of this species have been used as exemplars in a previously published molecular phylogeny: they were listed by + +Polotow +et al. +(2015) + +as “ + +Zorodictyna +sp. 3 + +” (CASENT_9029890; CAS) and “ + +Zorodictyna +sp. 4 + +” (CASENT_9029889; CAS). + + + +Natural history + + + +The species + +Tabiboka polotowae + +gen. et sp. nov. +is recorded from both highly disturbed and intact forest habitats. Many specimens have been collected while concealed in leaf litter. + +
+ + + +Distribution + + + + +Tabiboka polotowae + +gen. et sp. nov. +occurs in the eastern uplands of +Madagascar +from the south to the center ( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFEFFFD6FDF60E9FFA8ED8E6.xml b/data/31/57/E0/3157E00AFFEFFFD6FDF60E9FFA8ED8E6.xml new file mode 100644 index 00000000000..e75dd1bf4c6 --- /dev/null +++ b/data/31/57/E0/3157E00AFFEFFFD6FDF60E9FFA8ED8E6.xml @@ -0,0 +1,647 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Tabiboka milloti + +gen. et sp. nov. + + + + +urn:lsid:zoobank.org:act: +FF163E94-73C4-4D75-AC93-1BEB4FCA536F + + + + +Figs 8–9, 26 + + +Zorodictyna +sp. + +CG45 – + +Wheeler +et al. +2017 + +. + + + + + +Diagnosis + + + +The males of + +Tabiboka milloti + +gen. et sp. nov. +may be distinguished by the palpal tibia with a short, blunt retromedian process (RTA) and broad, thumb-like ventral apical process (VTA) (Figs 8E, G, 9B–C), the slender embolus (E) and the elongate, cruciform median apophysis (MA) with the apex extending to embolus tip (Figs 8F, 9A–B) (vs in + +T. milleri + +gen. et sp. nov. +, the the VTA is smaller and the RTA is greatly elongated, +Fig. 5A–C +, and the E and the MA are shorted and broader; and in + +T. polotowae + +gen. et sp. nov. +, the VTA is finger-like and the MA is shorter, +Fig. 11A–C +). Females of + +T. milloti + +have the epigyne with a heart-shaped median lobe (ML), broad anteriorly (anterior width greater than one-half total width, +Fig. 9D +) (vs in + +T. milleri + +, the ML is narrow anteriorly, anterior width less than one-half total width, +Fig. 5D +; and in + +T. polotowae + +, the ML is rectangular, narrowest in middle or with sides straight, +Fig. 11D +). + + + + + +Etymology + + + +The species name is a patronym in honor of Jacques Millot, who collected the +holotype +of this species and many other interesting arthropods in +Madagascar +during his missions in the mid-20 +th +century. + + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Toamasina Province + +• + +; Périnet (= Andasibe), Res. Analamazaotra, Parc National Andasibe; + +Aug. 1946 + +; +J. Millot +leg.; CASENT_9016063; +MNHN +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Toamasina Province + +• +1 ♀ +; Res. Analamazaotra, +Parc National Andasibe +, road + +23 km +E of Moramanga + +; + +960 m +a.s.l. + +; +18°56′38″ S +, +48°25′03″ E +; + +16–18 Jan. 2003 + +; +C. Griswold +, +D. Silva +and +D. Andriamalala +leg.; general collecting in rainforest; BLF7993; CASENT_9018308; +CAS + +• + +1 ♂ +; same data as for preceding; general collecting at night in rainforest; AToL DNA voucher: ARACG045; BLF7994; CASENT_9005657; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9005261; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +– + +Antananarivo Province + +• +2 ♂♂ +; +Tsinjoarivo forest +, +Ankadivory +; +19°42′56.592″ S +, +47°49′15.0234″ E +; + +1385 m +a.s.l. + +; + +11–14 Jan. 2015 + +; +B.L. Fisher +et al. +leg.; +pitfall traps +in montane rainforest; BLF35447-5; CASENT_9065578; +CAS + +• + +1 ♂ +; same data as for preceding; BLF35447-20; CASENT_9065583; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_906558; +CAS + +. + + + + + +Description + + + + +Male +holotype + +(CASENT_9016063; MNHN) + +MEASUREMENTS. Total length: 7.50. Carapace: length 4.60, width 3.70, height 1.80. +COLORATION (Fig. 8A–B). Carapace pale brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black surrounding all eyes; chelicerae brownish yellow; sternum yellow-white, unmarked; legs brownish yellow, femora, tibiae and metatarsi with faint median and subapical dusky annuli; abdomen pale yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides mottled with dark marks, venter yellow except with darker mottling around spinneret base. +STERNUM. Oval with slightly sinuous margins; 2.00 wide, 2.50 long. +CHILUM. 0.13 high, 0.35 wide. +EYES. ALE 0.33; AME 0.13; PLE 0.15; PME 0.14; ALE–AME 0.05; AME–AME 0.20; PLE–PME 0.13; PME–PME 0.06; MOQ: AW 0.26, PW 0.38, L 0.30. +CLYPEUS. 0.10 high, with six long setae. +LEGS. Trochanters deeply notched. Unpaired claw smooth, rudimentary on legs III and IV. Metatarsi lacking subbasal crack. Scopulae absent. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I4.301.803.903.502.2015.50
II3.801.803.103.001.9013.60
III3.501.502.603.301.8012.70
IV5.301.804.005.402.4018.90
+
+Spination MALE PALP (Figs 8C–G, 9A–C). Tibia with VTA thumb-shaped, RTA broad, median with sharp, ventroapical point (vRTA), blunt retrodorsal projection (dRTA); cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setae, with small, sharp, conical retrobasal cymbial extension (CE) extending to tibial apex; subtegulum large, filling most of cymbial concavity, with transverse ridge (SL) that interacts with tegulum; tegulum oval with large, with rounded retrodistal protuberance and conical promedian lobe (TL) that locks with subtegulum; embolus (E) arises from retromargin of tegulum, short, simple, narrow; median apophysis (MA) strongly sclerotized elongate hook curved 270° prolaterad; conductor (C) hyaline, translucent, flattened, fan-shaped, widened at extremity; paraconductor (Pc) finger-like membranous, originating at base of MA and extending two-thirds length of E. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d3rl1pl2rl1v2-2-2-2pl1-2v2-2-2
IIpl2d3rl2pl2rl2v2-2-2-2pl1-2v2-2-2
IIIpl3d3rl2pl1rl1pl2rl2v2-2-2pl2rl2v2-2-2
IVpl3d3rl2rl1pl2d1rl2v2-2-2pl4d3rl2v1-2-1-2-2
+
+ + +Female +paratype + +(CASENT_9018308; CAS) + +MEASUREMENTS. Total length: 10.20. Carapace: length 5.00, width 3.60, height 1.80. +COLOUR.As in male except contrast between yellow-white background and dusky markings much bolder, especially annuli on posterior femora, tibiae and metatarsi. +STERNUM. Oval with slightly sinuous sides; 2.20 wide, 2.40 long. +CHILUM. 0.13 high, 0.35 wide. +EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.18; ALE–AME 0.08; AME–AME 0.15; PLE–PME 0.18; PME–PME 0.16; MOQ: AW 0.25, PW 0.38, L 0.38. + +Fig. 8 +(on next page). + +Tabiboka milloti + +gen. nov. et sp. nov. +male +holotype +(CASENT_9016063; MNHN). +A +. Habitus, dorsal view. +B +. Carapace, dorsal view. +C +. SEM of right male palp (mirrored), prolateral view. +D +. Idem, ventral view. +E +. Idem, retrolateral view. +F +. Idem, detail of bulb, ventral view. +G +. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = +4.6 mm +); C–E = +0.5 mm +; F–G = +0.1 mm +. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick. + + + +Fig. 9. + +Tabiboka milloti + +gen. nov. et sp. nov. +genitalia drawings. +A–C = +male holotype (CASENT_9016063; MNHN). +D–E = +female paratype (CASENT_9018308; CAS). +A +. Left palp, prolateral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Epigyne, ventral view. +E +. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold. + + +CLYPEUS. 0.20 high, with five long setae. +LEGS. Scopulae absent. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I3.501.703.002.501.4012.10
II3.201.702.502.301.5011.20
III3.201.302.202.701.4010.80
IV4.501.603.404.501.9015.90
+
+ +Spination GENITALIA. Epigyne ( +Fig. 9D +) median lobe (ML) heart-shaped, sides rounded, broad anteriorly, anterior width greater than one-half total width. Endogyne ( +Fig. 9E +) with each spermatheca narrow, longer than broad. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d1v2-2-2-2v2-2-2
IIpl1d1v2-2-2-2v2-2-2
IIIpl1d3rl1pl2d2rl2v2-2-2pl4d2rl3v2-2-2
IVpl1d3rl1pl2d2rl2v2-2-2pl2d5rl3v2-1-2-2
+
+ + +Variation + + + +Male +(n = 3) + + +Total length = 6.90–7.50, carapace length/width = 1.21–1.29, carapace height/width = 0.43–0.59, PER/carapace width = 0.23–0.24, PER/OAL = 2.38–2.44, PER/AER = 1.38–1.39, OAL/OQL = 1.13– 1.23, OQP/OQA= 1.38–1.43, clypeus height/AME = 1.14–1.56, cheliceral length/clypeus height = 9.14– 18.00, sternum length/width = 1.05–1.25, palpal coxa length/width = 2.00–2.80, femur I length/carapace width = 1.11–1.12, metatarsus I length/carapace width = 0.95–1.00, femur IV length/carapace width = 1.23–1.43, cymbium length/carapace width = 0.32–0.35, cymbium length/palpal patella length = 1.20– 1.50, cymbium length/palpal tibia length = 1.09–1.26, cymbium length/palpal femur length = 0.63–0.75, palpal tibia length/palpal patella length 1.10–1.38. Males from Tsinjoarivo and Analamazaotra are marked like the +holotype +but the contrast between the yellow-white background and dusky markings on the cephalothorax, legs and abdomen is much greater. + + +Female + + +Variation unknown; only +one female +has been discovered. + + + +Note on synonym + + + +A specimen of this species (CASENT_9005657) was used as an exemplar in a previously published molecular phylogeny: it was listed as “ + +Zorodictyna +sp. + +CG45” by + +Wheeler +et al. +(2017) + +. + + + +Natural history + + + +The species + +Tabiboka milloti + +gen. et sp. nov. +is only known from rainforest. + +
+ + + +Distribution + + + + +Tabiboka milloti + +gen. et sp. nov. +is known only from mountains in east-central +Madagascar +( +Fig. 26 +). + + +
+
\ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFF4FFC4FD2A0D19FBE0DB55.xml b/data/31/57/E0/3157E00AFFF4FFC4FD2A0D19FBE0DB55.xml new file mode 100644 index 00000000000..d63fcd99e90 --- /dev/null +++ b/data/31/57/E0/3157E00AFFF4FFC4FD2A0D19FBE0DB55.xml @@ -0,0 +1,358 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + +Genus + +Tabiboka + +gen. nov. + + + + +urn:lsid:zoobank.org:act: +E7CAC38D-2089-4115-9764-56F76B3F621F + + + + + +Figs 3–12 +, +26 + + + + + + +Type +species + + + + + +Tabiboka milleri + +gen. et sp. nov. + + + + + +Diagnosis + + + + +Tabiboka + +gen. nov. +are ecribellate udubids where the male has palps with a short cymbial retrobasal extension (CE) extending to or just past tibial apex, lack a retrolateral paracymbial process, and have the tibia with both VTA and RTA ( +Figs 5A–C +, +6A–C +, +9A–C +, +11A–C +, +12A–D +). The female epigyne is provided with a median lobe (ML) that extends to the posterior margin, separating lateral lobes (LL) and covering the copulatory openings (CO) ( +Figs 4B +, +5D +, +7A–B +, +9D +, +11D +, +12G +). The genus is distinguished from + +Uduba +Simon, 1880 + +males by lacking the threadlike embolus and knob-like process between the palpal patella and tibia and from + +Uduba + +females by having compact spermathecae rather than multiple loops. Males differ from + +Zorascar + +gen. nov. +by having the cymbial retrobasal extension short, extending to or just past the tibial apex and the palpal tibia with a VTA and RTA. Females differ from the ones of + +Zorascar + +gen. nov. +by having the epigyne with a median lobe that extends to the posterior margin, separating the lateral lobes and covering the copulatory openings. + +Tabiboka + +is distinguished from + +Zorodictyna +Strand, 1907 + +by lacking an entire cribellum or vestige and lacking an extensive scopulae beneath the leg metatarsi. It is distinguished from all +Madagascar +udubids, except + +Zorascar + +by having the posterior spinnerets (PMS and PLS) reduced in size, and the PLS less than half the size of the ALS. + + + + + +Etymology + + +The genus name is an arbitrary combination of letters. The gender is feminine. + + + + +Description + + + +BODY. Medium size spiders (males: 6.90–12.00; females: 8.40–17.00) with smooth teguments of carapace ( +Figs 3A, C–D, F +, 8A–B, 10A–B), sternum and legs. Carapace piriform, longer than wide (L/W: males: 1.21–1.41; females: 1.26–1.46), with sparse cover of short, dark setae; thoracic fovea present, linear; profile higher in ocular and thoracic area and abruptly sloping posteriorly ( +Fig. 3G–H +). + + +COLORATION ( +Figs 3 +, +4A +, 8A–B, 10). Carapace brownish yellow with darker longitudinal pattern on carapace; chelicerae uniform brownish orange; legs faintly ringed; abdomen brownish yellow with faint complex pale pattern. + + +EYES ( +Figs 3C, F +, 8B, 10A–B). In two straight rows: ALE larger than AME, PLE and PME similar to ALE; tapetum of secondary eyes canoe-shaped, shiny. Clypeus with long setae, height 1.14 to 2.86 diameter of AME in male and +1.11 to 2.40 in +female. + + +PROSOMA. Chilum double, about three times or more as wide as high. Chelicerae ( +Figs 3D, F–H +): paturon conical, with some long setae medially, three teeth on both margins, with retrolateral thick setae at fang base; fang with serrate texture on proximal edge. Labium ( +Figs 3B, E +, +10C–D +) roughly diamond shaped. Endites ( +Figs 3B, E +, +10C–D +) roughly rectangular, with retrolateral margins sinuous, distal part slightly wider than proximal part, with subdistal serrula present as simple row. Sternum ( +Figs 3B, E +, +10C–D +) oval, slightly longer than wide, with slightly curved margins, not extending between coxae IV. + + +LEGS. Slender ( +Figs 3A, H +, +4A +, 8A, 10). Formula 4123. With three tarsal claws; unpaired claw smooth, present on legs I and II, large to small, typically reduced to nubbins on legs III and IV; superior claws with six to eight teeth; without claw tufts or tenent setae; scopulae absent or rarely present (female of + +T. polotowae + +gen. et sp. nov. +). Trichobothria in two or more rows on tarsi, in three or more on metatarsi, starting close to tip, all equal in lengths; distal and proximal plates of bothria well differentiated, proximal plates with transverse ridges. Tarsal organ: tear-drop or keyhole-shaped opening, close to tarsus tip. Spines numerous, with four pairs of ventral spines on tibia I, not overlapping, and with three pairs of ventral spines on metatarsus I. Calamistrum absent. Male tibial crack (autospasy suture) present or absent. Distal borders of leg trochanters with deep (forelegs) or shallow (hind legs) notch ( +Fig. 4C–D +). + + +ABDOMEN ( +Figs 3A–B, D–E, G–H +, 8A, 10). Oval, without scuta and without cribellum. Epiandrous glands absent. Colulus: as triangular hairy lobe. Spinnerets: ALS and PLS, conical, biarticulate; PMS one-segmented; ALS large, PMS and PLS small. Females with two ALS major ampullate gland spigots and field of about 20–30 piriform gland spigots and tartipores; PMS with one anterior minor ampullate gland spigot and large tartipore and one to few posterior cylindrical gland spigots and aciniform gland spigots; PLS with one to few cylindrical gland spigots; males lacking cylindrical gland spigots and second of each pair of both major and minor ampullate gland spigots reduced to nubbins. + + +MALE PALP ( +Figs 5A–C +, +6 +, +9A–C +, +11A–C +). Tibia with large or stout RTA, inserted medially, provided with basal concavity (BC), with short rounded or tapered VTA and with sharp triangular dorso-prolateral process. Cymbium oval with rounded distal extremity, without dorsal patch of short setae, with cymbial retrobasal extension (CE), without retrolateral paracymbial process. Bulbus provided with retrobasal subtegulum locking lobe (SL), and tegulum rounded with well-developed prolateral conical protrusion at base of embolus (corresponding to locking lobe, TL). Embolus (E) with fixed base, originating prolateraly, directed clockwise, long, large and lamellate ( +Figs 5A–C +, +6D–F +) or slender and tapered apically (Figs 8C–F, 9A–B, 11A–C, 12A–C, E). Median apophysis (MA) hook-shaped, situated opposite of embolus; conductor (C) hyaline, well developed, fan-shaped, medially extending from narrow base to wide flattened tip. Paraconductor (Pc) as narrow, finger-like membranous extension inserted between tegulum and base of MA. + + +GENITALIA. Epigyne ( +Figs 4B +, +5D +, +7A–B +, +9D +, +11D +) single, shield-shaped plate (ML) with convex median and lateral folds, lateral sector (LL) convex or with pocket. Endogyne ( +Figs 5E +, +7C–F +, +9E +, +11E +, +12H +): copulatory ducts (CD) very short, twisted; spermatheca (S) two or three lobed, head of spermatheca provided with pores; no evidence of enlarged Bennett’s Gland pore through spermathecal base. + + + +Species included + + + + +Tabiboka milleri + +gen. et sp. nov. +( +type +species; + + +); + +Tabiboka milloti + +gen. et sp. nov. +( + + +); + +Tabiboka polotowae + +gen. et sp. nov. +( + + +). + + + +Affinities + + + +From the molecular analyses ( +Fig. 2 +), + +Tabiboka + +gen. nov. +appears closely related to species in the +Udubidae +that have been identified as + +Zorascar + +gen. nov. + + + + + +Distribution + + + + +Tabiboka + +gen. nov. +are known only from mid-eastern +Madagascar +( +Fig. 26 +). + + + + \ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml b/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml new file mode 100644 index 00000000000..39d7e1667a4 --- /dev/null +++ b/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml @@ -0,0 +1,1720 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + + + +Tabiboka milleri + +gen. et sp. nov. + + + + +urn:lsid:zoobank.org:act: +33C8C4F1-A241-41E2-9B38-1D31AFBF4BE4 + + + + + +Figs 3–7 +, +26 + + +Zorocratidae +gen. Talatakely sp. Talatakely – +Morphbank 2023 +: images [476092–509952] SpiderAToL, +March 3, 2008 +, https://www.morphbank.net/?id=218853 [accessed +20 Oct. 2023 +]. + + + +Zorodictyna +sp. 2 + +– + +Polotow +et al. +2015 + +. + + + +Zorodictyna +sp. + +CG300 – + +Wheeler +et al. +2017 + +. + + + + + +Diagnosis + + + +Males of + +Tabiboka milleri + +gen. et sp. nov. +may be distinguished by the morphology of the palpal tibia with the RTA having an elongate curved spur arising retrobasally and extending distally to the apex of the tibia ( +Figs 5B–C +, +6A–C +) (vs + +T. milloti + +gen. et sp. nov. +and + +T. polotowae + +gen. et sp. nov. +have a short and stout RTA, +Figs 9B–C +, +11B–C +), a short, broad embolus (E) and a small, slender paraconductor (Pc) largely hidden behind the embolus ( +Figs 5A–C +, +6D–F +) (vs in + +T. milloti + +and + +T. polotowae + +, the E is slender and the Pc is situated between the E and the C, Figs 8F, 9B, 11B, 12E). Females of + +T. milleri + +have the epigyne with a broad, swollen, oval median lobe (ML) that extends posteriorly beyond the epigastric furrow, narrow anteriorly (anterior width less than half the total width, +Figs 4B +, +5D +, +7A +) (vs in + +T. milloti + +, the ML is broad anteriorly, +Fig. 9D +; and in + +T. polotowae + +, the ML is medially narrowed, +Figs 11D +, +12G +). + + + + + +Etymology + + + +The species name is a patronym in honor of Jeremy Miller, who collected widely in +Madagascar +and led a team that carried out a pioneering study of entelegyne spider phylogeny ( + +Miller +et al. +2010 + +). + + + + + +Type material + + + + +Holotype + + + + +MADAGASCAR +– + +Fianarantsoa Province + +• + +; +Ranomafana National Park +, +Talatakely +; +21°15′ S +, +47°25′ E +; + +900 m +a.s.l. + +; + +5–7 Dec. 1993 + +; +C. Griswold +, +N. Scharff +, +S. Larcher +and +R. Andriamasamanana +leg.; CASENT_9016094; +CAS +. + + + + +Paratypes + + + + +MADAGASCAR +– + +Fianarantsoa Province + +• +1 ♀ +; same collection data as for holotype; [ +Standard +view preparations and images for AToL-Spiders project; SEM preparations +Box +79-8, 10, 11, 19, +Box +81, 1, 2]; CASENT_9016094; +CAS + +• + +1 ♂ +, +1 ♀ +; +Ranomafana +N.P.; +21°12′ S +, +47°27′ E +; + +18 May 1992 + +; +V. and B. Roth +, +S. Kariko +leg.; general collecting; [habitus photos, stacked images, +H.M. Wood +; + +Apr. 2005 + +]; CASENT_9016098; +CAS + +• + +1♂ +, +1 ♀ +; same data as for preceding; [ + +palp drawn]; CASENT_9016086; +CAS + +• + +1 ♂ +, +1 ♀ +; Parc National +Ranomafana +, +Talatekely forest +, + +42.3 km +58° NE of +Fianarantsoa + +; +21°15′28.0″ S +, +47°25′21.8″ E +; + +1050 m +a.s.l. + +; + +24 Dec. 2005 + +– + +14 Jan. 2006 + +; +H. Wood +, +J. Miller +, +J.J. Rafonomezantsoa +, +E. Rajeriarison +, +V. Andriamananony +leg.; montane rainforest; general collecting; [AToL DNA +Voucher +ARACG0300] [HW009/RNAlater: Mad-052]; CASENT_9024004; +CAS + +• + +1 ♂ +; +Ranomafana +N.P.; +21°12′ S +, +47°27′ E +; + +18 May 1992 + +; +V. and B. Roth +, +S. Kariko +leg.; general collecting; [stacked images, HMW, + +April-2005 + +]; CASENT_9016097; +CAS + +• + +1 ♀ +; Parc National +Ranomafana +, +Vohiparara +, + +3.6 km +W of Ranomafana + +; +21°14.243′ S +, +47°23.842′ E +; + +1150 m +a.s.l. + +; + +13–14 Jan. 2009 + +; +D. Andriamalala +, +C. Griswold +, +G. Hormiga +, +A. Saucedo +, +N. Scharff +and +H. Wood +leg.; primary montane rainforest; general collecting day and night; [ +NS Photo +voucher 893-897; GH photos 5659-5666]; CASENT_9029871; +CAS + +. – + + +Mahajanga Province + +• +1 ♀ +; +Ampijoroa +, +National Park Ankarafantsika +, near +Lake Ravelobe +; +16°17′48″ S +, +46°48′50″ E +; + +26–28 Jan. 2009 + +; +C. Griswold +, +A. Saucedo +, +H. Wood +leg.; [SEM female epigyne + used for DNA voucher DP-2014c - “ +Zorodictyna sp. 2 +”]; CASENT_9035866; +CAS + +. + + + +Other material examined + + + + +MADAGASCAR +– + +Antananarivo Province + +• +1 ♂ +; +Tsinjoarivo Forest +, +Mahatsinjo +; +19°40′47.496″ S +, +47°46′22.835″ E +; + +1654 m +a.s.l. + +; + +26–29 Aug. 2014 + +; +B.L. Fisher +leg.; montane rainforest; BLF35410; CASENT_9065708; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9065709; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9065678; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9065579; +CAS + +• + +4 ♂♂ +, +5 ♀♀ +; same data as for preceding; CASENT_9017817; +CAS + +• + +2 ♀♀ +; +Tsinjoarivo Forest +, +Ankadivory +; +19°43′00″ S +, +47°49′18″ E +; + +1369 m +a.s.l. + +; + +28 Apr.–1 May 2014 + +; +B.L. Fisher +et al. +leg.; montane rainforest, sifted litter; BLF33370; CASENT_9064661; +CAS + +. – + + +Fianarantsoa Province + +• +1 ♀ +; +Forêt d’Ankazomivady +, + +28 km +SSW of Ambositra + +, +4.7 km +SW of +Ambalamanakana +; +20°47′ S +, +47°10′ E +; + +1670 m +a.s.l. + +; + +7–14 Jan. 1998 + +; +S. Goodman +, SG0001 leg.; disturbed upper montane forest; +pitfall traps +#1–3; CASENT_9016064; +FMNH + +• + +9 ♀♀ +; same data as for preceding; CASENT_9016065; +FMNH + +• + +1 ♀ +; same data as for preceding; CASENT_9016066; +FMNH + +• + +9 ♀♀ +; same data as for preceding; CASENT_9016067; +FMNH + +• + +1 ♂ +; same data as for preceding; CASENT_9016068; +FMNH + +• + +1 ♂ +; +Ankazomivady +, + +29 km +SSW of Ambositra + +; +20°47′ S +, +47°10′ E +; + +1700 m +a.s.l. + +; + +7 Jan. 1998 + +; +B.L. Fisher +leg.; forest; EBD10 +pitfall trap +; BLF1592b; CASENT_9016087; +CAS + +• + +1 ♀ +; +Ambatofitorahana +, + +33 km +S of Ambositra + +; +20°49′00.00″ S +, +47°10′59.99″ E +; + +13 Mar. 1994 + +; +A. Pauly +leg.; natural forest; BE_ +RMCA +_ARA. +Ara. +201626; +RMCA + +• + +24 ♂♂ +, +5 ♀♀ +; +Ranomafana National Park +, +Talatakely +; +21°12′ S +, +47°27′ E +(coordinates approximate); + +900 m +a.s.l. + +; + +4–16 Jan. 2001 + +; +D.H. and K.M. Kavanaugh +, +R.L. Brett +, +E. Elsom +and +F. Vargas +leg.; mixed tropical forest; +pitfall traps +#4–16; COL-DHK-2001-001TN; CASENT_9003432; +CAS + +• + +1 ♀ +; same data as for preceding; CASENT_9003534; +CAS + +• + +1 ♀ +; Parc National +Ranomafana +, +Talatakely +; +21°15′ S +, +47°26′ E +; + +13–27 Apr. 1998 + +; +C.E. Griswold +, +D.H. Kavanaugh +, +N.D. Penny +, +M.J. Raherilalao +, +J.S. Ranorianarisoa +, +J. Schweikert +and +D. Ubick +leg.; +pitfall traps +; CASENT_9006115; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9016084; +CAS + +• + +1 ♂ +; Parc National +Ranomafana +, +Talatakely +; +21°15′ S +, +47°26′ E +; + +19–30 Apr. 1998 + +, +C.E. Griswold +, +D.H. Kavanaugh +, +N.D. Penny +, +M.J. Raherilalao +, +J.S. Ranorianarisoa +, +J. Schweikert +, +D. Ubick +leg.;general collecting; CASENT_9016082; +CAS + +• + +2 ♀♀ +; same data as for preceding; CASENT_9016088; +CAS + +• + +6 ♂♂ +; Parc National +Ranomafana +, +Talatakely +, trail F; +21°16′ S +, +47°25′ E +; + +940–985 m +a.s.l. + +; + +1–21 Nov. 1998 + +; +V.F. Lee +and +K.J. Ribardo +leg.; +pitfall trap +; CASENT_9016089; +CAS + +• + +2 ♂♂ +; same data as for preceding; CASENT_9016090; +CAS + +• + +1 ♂ +; same data as for preceding; CASENT_9016093; +CAS + +• + +2 ♀♀ +; Parc National +Ranomafana +, +Vatoharanana +; +21°17′ S +, +47°26′ E +; + +1200 m +a.s.l. + +; + +15–29 Apr. 1998 + +; +C.E. Griswold +, +D.H. Kavanaugh +, +N.D. Penny +, +M.J. Raherilalao +, +J.S. Ranorianarisoa +, +J. Schweikert +, +D. Ubick +leg.; primary forest; CEG-MA-1998-03; CASENT_9016080; +CAS + +• + +1 ♀ +; Parc National +Ranomafana +, +Vatoharanana +, trail K; +21°17′ S +, +47°26′ E +; + +990 m +a.s.l. + +; + +14–19 Nov. 1998 + +; +V.F. Lee +and +K.J. Ribardo +leg.; +pitfall trap +; VFL-KJR-02; CASENT_9016091; +CAS + +• + +1 ♀ +; same data as for preceding; CASENT_9016092; +CAS + +• + +1 ♀ +; Parc National +Ranomafana +, +Vatoharanana River +, + +4.1 km +231° SW of Ranomafana + +; +21°17′24″ S +, +47°26′00″ E +; + +1100 m +a.s.l. + +; + +27–31 Mar. 2003 + +; +Fisher-Griswold Arthropod Team +leg.; montane rainforest; general day collecting; BLF8394; CASENT_9017837; +CAS + +• + +1 ♀ +; Parc National +Ranomafana +, +Vohiparara +Piste Touristique; +21°14′ S +, +47°24′ E +; + +1000 m +a.s.l. + +; + +23 Apr. 1998 + +; +C.E. Griswold +, +D.H. Kavanaugh +, +N.D. Penny +, +M.J. Raherilalao +, +E. Rajeriarison +, +J.S. Ranorianarisoa +, +J. Schweikert +, +D. Ubick +leg.; running in litter; general collecting; CEG-MA-1998; CASENT_9016081; +CAS + +• + +6 ♂♂ +; +Ranomafana National Park +, +Vohiparara +; +21°12′ S +, +47°27′ E +(coordinates approximate); + +1150 m +a.s.l. + +; + +2–22 Jan. 2001 + +; +D.H. and K.M. Kavanaugh +, +R.L. Brett +, +E. Elsom +, +F. Vargas +leg.; mixed tropical forest; general collecting; COL- DHK-2001-004; CASENT_9003479; +CAS + +• + +6 ♀♀ +; Parc National +Ranomafana +, + +2.3 km +N of Vohiparara Village + +; +21°13′ S +, +47°23′ E +; + +1100 m +a.s.l. + +; + +18 Apr. 1998 + +; +C.E. Griswold +, +D.H. Kavanaugh +, +N.D. Penny +, +M.J. Raherilalao +, +E. Rajeriarison +, +J.S. Ranorianarisoa +, +J. Schweikert +, +D. Ubick +leg.; CASENT_9016083; +CAS + +• + +6♀♀ +; same data as for preceding; CASENT_9016085; +CAS + +• + +2 ♀♀ +; +Ranomafana +N.P., +Trail +F; +21°12′ S +, +47°27′ E +(coordinates approximate); + +18 May 1992 + +; +Albert +for +Kariko +, +Roth +leg.; sifting #1; CASENT_9030889; +MCZ + +• + +1 ♀ +; +Ranomafana +N.P., +Maharira +, summit; +21°12′ S +, +47°27′ E +; + +9 Apr. 1992 + +; +Albert +for +Kariko +, +Roth +leg.; cryptic #2; CASENT_9030890; +MCZ + +• + +1 ♂ +; +Ranomafana +N.P., +Trail +M; +21°12′ S +, +47°27′ E +(coordinates approximate); + +15 May 1992 + +; +Albert +for +Kariko +, +Roth +leg.; sifting 1hr. samples #2; CASENT_9030891; +MCZ + +• + +1 ♀ +; +Ranomafana +N.P., +Trail +M; +21°12′ S +, +47°27′ E +(coordinates approximate); + +15 May 1992 + +; +Albert +for +Kariko +, +Roth +leg.; sifting; CASENT_9030892; +MCZ + +• + +3 ♀♀ +; +Ranomafana +N.P., +Trail +B; +21°12′ S +, +47°27′ E +(coordinates approximate); + +825 m +a.s.l. + +; + +19 May 1992 + +; +Albert +for +Kariko +, +Roth +leg.; sifting, #1; CASENT_9030893; +MCZ + +• + +1 ♀ +; +Ranomafana +N.P., +Maharira +, summit; +21°12′ S +, +47°27′ E +(coordinates approximate); + +11 Apr 1992 + +; +Emile +for +Kariko +, +Roth +leg.; sifting #3; CASENT_9030894; +MCZ + +• + +2 ♀♀ +; +Ranomafana +N.P.; +21°12′ S +, +47°27′ E +; + +Apr. 1992 + +; +V. and B. Roth +, +S. Kariko +leg.; forest; general collecting from foliage; Ranomafana-1000-92; CASENT_9016099; +CAS + +• + +1 ♂ +, +1 ♀ +; +Ranomafana +N.P., trail F, secondary forest; +21°12′ S +, +47°27′ E +(coordinates approximate); + +Apr. 1992 + +; +Albert +for +Kariko +, +Roth +leg.; sifting #3; MCZ-VBR-0116; CASENT_9016100; +MCZ + +• + +4 ♂♂ +, +7 ♀♀ +; +Ranomafana +N.P.; +21°12′ S +, +47°27′ E +(coordinates approximate); + +20–31 May 1992 + +; +V. and B. Roth +leg.; +can traps +; MCZ-VBR-02; CASENT_9016101; +MCZ + +• + +1 ♂ +; same data as for preceding; CASENT_9016102; +CAS + +. + + + + +Fig. 3. + +Tabiboka milleri + +gen. et sp. nov. +, A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS). +A, D +. Habitus, dorsal view. +B, E +. Idem, ventral view. +C, F +. Carapace, dorsal view. +G–H +. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood. + + + + +Fig. 4. + +Tabiboka milleri + +gen. et sp. nov. +A +. Female paratype (CASENT_9029871; CAS) from Ranomafana (Photo © N. Scharff). +B +. Female paratype (CASENT_9016098; CAS), epigyne, ventral view. +C +. Female paratype (CASENT_9016094; CAS), trochanter 1, ventral view. +D +. Idem, trochanter 4. (No scales provided). Stacked microphotographs by Charles Griswold. + + + + +Fig. 5. + +Tabiboka milleri + +gen. nov. et sp. nov. +genitalia drawings of male and female paratypes (CASENT_9016086; CAS). +A +. Male left palp, prolateral view. +B +. Idem, ventral view. +C +. Idem, retrolateral view. +D +. Epigyne, ventral view. +E +. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels. + + + + +Fig. 6. + +Tabiboka milleri + +gen.nov.et sp. nov. +SEM of male right palp(mirrored), paratype(CASENT_9016094; CAS). +A +. Palp, ventral, slightly retrolateral view. +B +. Idem, retrolateral view. +C +. Idem, dorsal view. +D +. Bulb, prolateral view. +E +. Idem, ventral view. +F +. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–C = 0.43 mm; D–F = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick. + + + + +Fig. 7. + +Tabiboka milleri + +gen. nov. et sp. nov. +SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). +A +. Epigyne, ventral view. +B +. Idem, ventro-posterior view. +C +. Endogyne, dorsal view. +D +. Idem, slightly anterior view. +E +. Spermatheca, right dorsal view. +F +. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila. + + + + + +Description + + + + +Male +holotype + +(CASENT_9016094; CAS) + +MEASUREMENTS. Total length: 10.50. Carapace: length 6.10, width 4.40, height 2.20. + +COLORATION ( +Fig. 3A–C, G +). Carapace brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black on ocular area and surrounding all eyes; chelicerae brownish yellow; sternum yellow, unmarked; legs brownish yellow, unmarked; abdomen brownish yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides brownish yellow mottled with dark marks, venter yellow except darker around spinneret base. + + +STERNUM ( +Fig. 3B +). Oval with slightly sinuous margins; 2.30 wide, 2.60 long. + +CHILUM. 0.13 high, 0.35 wide. +EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.15; ALE–AME 0.06; AME–AME 0.40; PLE–PME 0.20; PME–PME 0.09; MOQ: AW 0.33, PW 0.48, L 0.35. +CLYPEUS. 0.25 high, with eight long setae. +LEGS. Trochanter I notched, trochanter IV shallowly notched. Unpaired claw smooth, rudimentary on legs III and IV. Scopulae absent. +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I4.702.004.804.702.8019.00
II4.201.903.804.002.2016.10
III4.301.703.504.102.0015.60
IV5.202.004.606.502.5020.80
+
+ +Spination MALE PALP ( +Figs 5A–C +, +6 +). Tibia with slender apical VTA, bifid apical RTA continuous with elongate, slender basal RTA; tibia with dorso-retrolateral excavation (BC) bordered dorsad of RTA, accommodating conical retrobasal cymbial extension; cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setae, with sinuous retrolateral margin, with conical retrobasal cymbial extension (CE) that extends ⅓ of tibial length from tibial apex; subtegulum large, filling most of cymbial concavity, with rounded lobe extending proximad and small, conical lobe (SL) that interacts with the tegulum; tegulum oval with large, rounded retrodistal protuberance and conical promedian lobe (TL) that locks with subtegulum; embolus (E) arises from center of tegulum, short, broad; strongly sclerotized MA forms short hook curved prolaterad; conductor (C) hyaline, translucent, flattened, fan-shaped, narrow at base and strongly widened at extremity; paraconductor (Pc) as finger-like membranous structure originating at base of MA and extending to anterior half of E. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl2d3rl3pl2d2rl2v2-2-2pl3rl3v2-2-3
IIpl3d3rl4pl3d2rl2v2-2-2-2pl5rl4v2-1-1
IIIpl3d3rl4pl1rl1pl2d2rl2v2-2-2pl4rl4v2-2-2
IVpl3d3rl2rl1pl2d2rl2v2-2-2pl4rl4v2-2-2
+
+ + +Female +paratype + +(CASENT_9016094; CAS) + +MEASUREMENTS. Total length: 14.00. Carapace: length 6.80, width 5.20, height 2.70. + +COLOUR ( +Figs 3D–F, H +, +4A +). As in male except abdomen much darker, mostly brown with lighter paracardiac marks and faint posterodorsal chevrons, sides dark, venter yellow brown with transverse dark mottling. + + +STERNUM ( +Fig. 3E +). Oval with slightly sinuous margins; 2.60 wide, 3.00 long. + +CHILUM. 0.13 high, 0.50 wide. +EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.15; ALE–AME 0.06; AME–AME 0.40; PLE–PME 0.25; PME–PME 0.11; MOQ: AW 0.40, PW 0.55, L 0.48. +CLYPEUS. 0.30 high, with eight long setae. + +LEGS. Trochanter I notched ( +Fig. 4C +), trochanter IV shallowly notched ( +Fig. 4D +). Unpaired claw smooth, rudimentary on legs III and IV. Scopulae absent. + +Leg measurements + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMttTotal
I5.102.404.203.601.9017.20
II4.802.403.603.301.8015.90
III4.001.902.904.001.8014.60
IV5.202.204.406.102.3020.20
+
+ +Spination GENITALIA. Epigyne ( +Figs 5D +, +7A–B +) with broad, oval median lobe (ML) that extends posteriad of epigastric furrow; lateral lobes (LL) convex, lacking teeth or concavities; copulatory openings (CO) located beneath anteromedian margin of median lobe and, in many specimens, containing simple plugs. Endogyne ( +Figs 5E +, +7C–F +) with spermathecae (S) dark, smoothly convex, simple or with slight transverse indentations; sperm ducts (CD) anteromedian, short, joining seamlessly with spermathecae; spermathecal heads with 20–30 small pores; fertilization ducts (FD) short, flattened and slightly curved ( +Figs 5E +, +7E +). + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
FPTMt
Ipl1d2v2-2-2-2v2-2-2
IIpl1d1v2-2-2-2v2-2-2
IIIpl2d3rl1pl1rl1pl2d2rl2v2-2-2pl5rl3v2-2-1-2
IVpl1d3rl1pl1rl1pl2d2rl2v2-2-2pl5rl3v2-2-1-2
+
+ + +Variation + + + +Males +(n = 5) + +Male genitalia display slight geographic variation. Males from Ranomafana have very slender retrobasal tibial processes, with length greater than 20 times width, whereas in males from Ankazomivay and Tsinjoarivo these are thick, with length less than 15 times width. The distal RTA may be deeply cleft in males from Ranomafana or only slightly cleft as in males from Ankazomivady. Most individuals have a crack across the subbasal part of the leg tibia just distad of the basalmost ventral spines, but this varies. Within the Ranomafana population some have the tibial leg crack clearly marked whereas in others the crack cannot be distinguished. Total length = 9.60–12.00, carapace length/width = 1.28–1.41, carapace height/width = 0.44–0.59, PER/carapace width = 0.24–0.26, PER/OAL = 2.33–2.65, PER/AER = 1.38– 1.46, OAL/OQL = 1.24–1.42, OQP/OQA = 1.46–1.57. Clypeus height/AME = 1.20–2.86, cheliceral length/clypeus height = 8.00–13.33, sternum length/width = 1.11–1.19, palpal coxa length/width = 2.14–2.50, femur I length/carapace width = 1.04–1.22, metatarsus I length/carapace width = 1.02–1.11, femur IV length/carapace width = 1.09–1.30, cymbium length/carapace width = 0.36–0.42, cymbium length/palpal patella length = 1.45–2.43, cymbium length/palpal tibia length = 1.23–1.90, cymbium length/palpal femur length = 0.76–0.92, palpal tibia length/palpal patella length 1.00–1.57. + +Females +(n = 5) + + +Total length = 11.50–14.00, carapace length/width = 1.26–1.31, carapace height/width = 0.45–0.65, PER/carapace width = 0.25–0.33, PER/OAL = 2.44–2.95, PER/AER = 1.35–1.70, OAL/OQL = 1.13– 1.38, OQP/OQA= 1.33–1.71, clypeus height/AME = 1.60–2.40, cheliceral length/clypeus height = 7.67– 12.50, sternum length/width = 1.14–1.30, palpal coxa length/width = 1.94–2.57, femur I length/carapace width = 0.92–1.10, metatarsus I length/carapace width = 0.62–0.71, femur IV length/ carapace width = 0.92–1.10, palpal tarsus length/carapace width = 0.32–0.37, palpal tarsus length/palpal tibia length = 1.15–1.31, palpal tarsus length/palpal femur length = 0.74–0.86, palpal tibia length/palpal patella length = 1.08–1.37. The oval epigynal median lobe may be evenly curved in lateral outline ( +Fig. 5D +) or be attenuated posteriorly ( +Fig. 4B +). Median lobe length 0.93–1.38 times width, with median lobe extending posteriad of epigastric furrow by 0.11–0.29 of lobe length. Spermathecae may be evenly rounded or have as many as two shallow dorsal constrictions. Spermathecae separated by 0.27–0.35 the total width of the spermathecae. + + + +Note on synonyms + + + +Specimens of this species have been used as exemplars in previous studies including morphological documentation and in published molecular phylogenies. The morphology of the species was documented for the Spiders Assembling the Tree of Life (AToL) study (2001–2009) as the exemplar “ +Zorocratidae +gen. Talatakely sp. Talatakely”, Images [476092–509952] SpiderAToL, +March 3, 2008 +, https://www.morphbank.net/?id=218853 ( +Morphbank 2023 +): CASENT_9016094. Specimens were used as the exemplars “ + +Zorodictyna +sp. 2 + +” by + +Polotow +et al. +(2015) + +(CASENT_9035866) and as “ + +Zorodictyna +sp. + +CG300” by + +Wheeler +et al. +(2017) + +(CASENT_9024004). + + + +Natural history + + +This species is commonly found running on leaf litter in closed-canopy montane rainforest. Individuals were also collected in pitfall traps or under objects. +
+ + + +Distribution + + + + +Tabiboka milleri + +gen. et sp. nov. +are known only from rainforests in +Antananarivo +and +Fianarantsoa +Provinces in southeastern +Madagascar +( +Fig. 26 +). + + +
+
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