Add updates up until 2024-10-30 20:41:04

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<document id="762D4F9F393CB3AA1AAA8AE6398F50B8" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630395207EF3EFE4F7219070D" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius inachoides" docType="treatment" docVersion="4" lastPageNumber="100" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="100" updateTime="1730320830895" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="E3D9551135BDD2E3B1829B99CE02A01E">
<mods:title id="EC2302C7EBEAC1A8147CB831AA8FF042">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="8CBB8B22853A084AF8F4ECEA3C1B4423">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="F7864DBFEDA70A2BE3AD149D942A3ECC">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="7C281931C22313D2A739047BF7EAF691" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="052A1BFF7C0A18A1DD152A1E39C4B093">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="A645844A2CA8445EEE7DFF3C0397E304" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="0E13F3152AB0A06996D54ACED7C57AF7">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="1A30D52614CEC55386BB32619712A01A" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="0E98F9CCEFAC3707B353FF1F77C2B21F">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="6319315F82363337E682BA705A8CCD50">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="AFB662DBB2A0487C63339C791422E85D" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="B37CE898C2AC6B8308F757BBEF334C1A">European Journal of Taxonomy</mods:title>
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<mods:date id="C9248441CD9786E97C8CC1EF497B7A43">2024</mods:date>
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<mods:number id="AB9DA00CA0F4DF92BF96FA861F0E772D">2024-10-11</mods:number>
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<mods:number id="D18B045BDF781F3B605CC9F50DAD2329">959</mods:number>
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<subSubSection id="DDE394FB30395207EF3EFE4F7307033C" box="[553,1035,488,515]" pageId="99" pageNumber="100" type="nomenclature">
<paragraph id="9546C77030395207EF3EFE4F7307033C" blockId="99.[553,1035,488,550]" box="[553,1035,488,515]" pageId="99" pageNumber="100">
<heading id="CE0E701C30395207EF3EFE4F7307033C" box="[553,1035,488,515]" centered="true" fontSize="11" level="2" pageId="99" pageNumber="100" reason="2">
<taxonomicName id="52F9BCF330395207EF3EFE4F7307033C" ID-CoL="35NC3" authority="(Felsche, 1901)" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[553,1035,488,515]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="inachoides">
<emphasis id="A78D1B6230395207EF3EFE4F7441033C" bold="true" box="[553,845,488,515]" italics="true" pageId="99" pageNumber="100">Dichotomius inachoides</emphasis>
(
<bibRefCitation id="F168BA8130395207EE4BFE4F730E033C" author="Felsche C." box="[860,1026,488,515]" pageId="99" pageNumber="100" pagination="135 - 155" refId="ref118967" refString="Felsche C. 1901. Beschreibungen coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift 45 (1): 135 - 155. Available from https: // www. biodiversitylibrary. org / page / 33086609 [accessed 16 Feb. 2024]." type="journal article" year="1901">Felsche, 1901</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DDE394FB30395207EFF1FDAB74430319" box="[742,847,524,550]" pageId="99" pageNumber="100" type="description">
<paragraph id="9546C77030395207EFF1FDAB74430319" blockId="99.[553,1035,488,550]" box="[742,847,524,550]" pageId="99" pageNumber="100">
<figureCitation id="0DC2DBF530395207EFF1FDAB74430319" box="[742,847,524,550]" captionStart="Fig" captionStartId="100.[189,232,1963,1989]" captionTargetBox="[269,1303,278,1896]" captionTargetId="figure-17@100.[258,1329,265,1922]" captionTargetPageId="100" captionText="Fig. 40. Species distribution. A. Dichotomius inachoides (Felsche, 1901). B. Dichotomius latilobatus Boilly &amp; Vaz-de-Mello, 2021. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945575" httpUri="https://zenodo.org/record/13945575/files/figure.png" pageId="99" pageNumber="100">Fig. 40A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30395207EDAAFDE974780267" pageId="99" pageNumber="100" type="reference_group">
<paragraph id="9546C77030395207EDAAFDE9746903B4" blockId="99.[189,1399,589,651]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="B5E9E05E30395207EDAAFDE9746903B4" pageId="99" pageNumber="100">
<treatmentCitation id="1458E16130395207EDAAFDE975880358" author="Felsche C." box="[189,644,589,615]" page="144" pageId="99" pageNumber="100" year="1901">
<taxonomicName id="52F9BCF330395207EDAAFDE975880358" ID-CoL="6VLCK" authority="Felsche, 1901: 144" authorityName="Felsche" authorityPageNumber="144" authorityYear="1901" box="[189,644,589,615]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="inachoides">
<emphasis id="A78D1B6230395207EDAAFDE976910358" box="[189,413,589,615]" italics="true" pageId="99" pageNumber="100">Pinotus inachoides</emphasis>
<bibRefCitation id="F168BA8130395207ECB2FDEA75880358" author="Felsche C." box="[421,644,589,615]" pageId="99" pageNumber="100" pagination="135 - 155" refId="ref118967" refString="Felsche C. 1901. Beschreibungen coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift 45 (1): 135 - 155. Available from https: // www. biodiversitylibrary. org / page / 33086609 [accessed 16 Feb. 2024]." type="journal article" year="1901">Felsche, 1901: 144</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030395207E92CFDEA73B80358" box="[1083,1204,589,615]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230395207E9D4FDEA72100358" box="[1219,1308,589,615]" country="Venezuela" name="Merida" pageId="99" pageNumber="100">Mérida</collectingRegion>
. Name-bearing type:
<typeStatus id="4A4279D230395207EC9BFDD776FF03B5" box="[396,499,624,650]" pageId="99" pageNumber="100" type="holotype">holotype</typeStatus>
(SMTD), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030395207EDAAFD15739C03F3" blockId="99.[189,1399,690,857]" box="[189,1168,690,716]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="B5E9E05E30395207EDAAFD15739C03F3" box="[189,1168,690,716]" pageId="99" pageNumber="100">
<emphasis id="A78D1B6230395207EDAAFD1576BF03F4" box="[189,435,690,716]" italics="true" pageId="99" pageNumber="100">
<taxonomicName id="52F9BCF330395207EDAAFD15769103F3" ID-CoL="6VLCK" authorityName="Felsche" authorityYear="1901" box="[189,413,690,716]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="inachoides">Pinotus inachoides</taxonomicName>
</emphasis>
<treatmentCitation id="1458E16130395207ECADFD15757503F3" author="Gillet J. J. E." box="[442,633,690,716]" page="61" pageId="99" pageNumber="100" year="1911">
<bibRefCitation id="F168BA8130395207ECADFD15757503F3" author="Gillet J. J. E." box="[442,633,690,716]" pageId="99" pageNumber="100" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 61</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130395207EE2FFD15734603F3" author="Blackwelder R. E." box="[824,1098,690,716]" page="207" pageId="99" pageNumber="100" year="1944">
<bibRefCitation id="F168BA8130395207EE2FFD15734603F3" author="Blackwelder R. E." box="[824,1098,690,716]" pageId="99" pageNumber="100" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030395207EDAAFD71737703D0" blockId="99.[189,1399,690,857]" box="[189,1147,725,751]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="B5E9E05E30395207EDAAFD71737703D0" box="[189,1147,725,751]" pageId="99" pageNumber="100">
<taxonomicName id="52F9BCF330395207EDAAFD71754603D0" ID-CoL="6VLCK" authorityName="Felsche" authorityYear="1901" box="[189,586,725,751]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="inachoides" subGenus="Cephagonus">
<emphasis id="A78D1B6230395207EDAAFD71761B03D0" box="[189,279,726,751]" italics="true" pageId="99" pageNumber="100">Pinotus</emphasis>
(
<emphasis id="A78D1B6230395207EC3FFD7276B703D0" box="[296,443,725,751]" italics="true" pageId="99" pageNumber="100">Cephagonus</emphasis>
)
<emphasis id="A78D1B6230395207ECDCFD72754603D0" box="[459,586,725,751]" italics="true" pageId="99" pageNumber="100">inachoides</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130395207EF7FFD72747603D0" author="Luederwaldt H." box="[616,890,725,751]" page="109" pageId="99" pageNumber="100" year="1929">
<bibRefCitation id="F168BA8130395207EF7FFD72747603D0" author="Luederwaldt H." box="[616,890,725,751]" pageId="99" pageNumber="100" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 109</bibRefCitation>
</treatmentCitation>
(key, characteristics).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030395207EDAAFD5F74780267" blockId="99.[189,1399,690,857]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="B5E9E05E30395207EDAAFD5F74780267" pageId="99" pageNumber="100">
<taxonomicName id="52F9BCF330395207EDAAFD5F75CA022D" ID-CoL="35NC3" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[189,710,760,786]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="inachoides" subGenus="Luederwaldtinia">
<emphasis id="A78D1B6230395207EDAAFD5F765F022D" box="[189,339,760,786]" italics="true" pageId="99" pageNumber="100">Dichotomius</emphasis>
(
<emphasis id="A78D1B6230395207EC7BFD5F7522022D" box="[364,558,760,786]" italics="true" pageId="99" pageNumber="100">Luederwaldtinia</emphasis>
)
<emphasis id="A78D1B6230395207EF50FD5F75CA022D" box="[583,710,760,786]" italics="true" pageId="99" pageNumber="100">inachoides</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130395207EFE3FD5E74A2022D" author="Roze J. A." box="[756,942,760,787]" page="44" pageId="99" pageNumber="100" year="1955">
<bibRefCitation id="F168BA8130395207EFE3FD5E74A2022D" author="Roze J. A." box="[756,942,760,787]" pageId="99" pageNumber="100" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
</treatmentCitation>
(cited, checklist for
<collectingCountry id="EDEE87E030395207E9D7FD5F7232022D" box="[1216,1342,760,786]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E16130395207EDFDFCBC75360209" author="Vulcano M. A. &amp; Pereira F. S." box="[234,570,795,822]" page="585" pageId="99" pageNumber="100" year="1967">
<bibRefCitation id="F168BA8130395207EDFDFCBC75360209" author="Vulcano M. A. &amp; Pereira F. S." box="[234,570,795,822]" pageId="99" pageNumber="100" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 585</bibRefCitation>
</treatmentCitation>
(key for the Amazon). —
<treatmentCitation id="1458E16130395207EE73FCBC732A020A" author="Krajcik M." box="[868,1062,795,821]" page="91" pageId="99" pageNumber="100" year="2012">
<bibRefCitation id="F168BA8130395207EE73FCBC732A020A" author="Krajcik M." box="[868,1062,795,821]" pageId="99" pageNumber="100" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 91</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130395207E985FCBC75190266" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." page="33" pageId="99" pageNumber="100" year="2021">
<bibRefCitation id="F168BA8130395207E985FCBC75190266" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." pageId="99" pageNumber="100" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello 2021: 7, 33</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30395207EDAAFC27767D05A3" pageId="99" pageNumber="100" type="materials_examined">
<paragraph id="9546C77030395207EDAAFC2776A602A5" blockId="99.[189,426,896,922]" box="[189,426,896,922]" pageId="99" pageNumber="100">
<heading id="CE0E701C30395207EDAAFC2776A602A5" bold="true" box="[189,426,896,922]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A78D1B6230395207EDAAFC2776A602A5" bold="true" box="[189,426,896,922]" pageId="99" pageNumber="100">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030395207EDAAFC08767D05A3" blockId="99.[189,1399,942,1180]" pageId="99" pageNumber="100">
<materialsCitation id="2591CD2D30395207EDAAFC0873F802F6" box="[189,1268,942,969]" collectingDate="2004-07-06" collectionCode="CEMT" collectorName="D. Garcia" country="Venezuela" elevation="50" location="Cuyagua" pageId="99" pageNumber="100" specimenCount="1" stateProvince="Aragua">
<collectingCountry id="EDEE87E030395207EDAAFC08766702F6" box="[189,363,943,969]" name="Venezuela" pageId="99" pageNumber="100">VENEZUELA</collectingCountry>
<emphasis id="A78D1B6230395207EC90FC0976EA02F7" bold="true" box="[391,486,942,968]" pageId="99" pageNumber="100">
<collectingRegion id="573D099230395207EC90FC0976EA02F7" box="[391,486,942,968]" country="Venezuela" name="Aragua" pageId="99" pageNumber="100">Aragua</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930395207ECEAFC08755E02F6" box="[509,594,943,969]" count="1" pageId="99" pageNumber="100" type="generic">1 spec.</specimenCount>
;
<location id="902691AB30395207EF49FC0875C602F6" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207EF49FC0875C602F6" box="[606,714,943,969]" country="Venezuela" name="Cuyagua" pageId="99" pageNumber="100" stateProvince="Aragua">Cuyagua</location>
;
<elevation id="1ED4204330395207EFC2FC09745C02F7" box="[725,848,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="99" pageNumber="100" unit="m" value="50.0">
<quantity id="52016A9530395207EFC2FC09741D02F6" box="[725,785,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="99" pageNumber="100" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030395207EE49FC0974EE02F6" box="[862,994,942,969]" pageId="99" pageNumber="100" value="2004-07-06">
<collectingDate id="F103185830395207EE49FC0974EE02F6" box="[862,994,942,969]" pageId="99" pageNumber="100" value="2004-07-06">6 Jul. 2004</collectingDate>
</date>
;
<collectorName id="380CA2A630395207EEF9FC08736C02F6" box="[1006,1120,943,969]" pageId="99" pageNumber="100">D. García</collectorName>
leg.;
<collectionCode id="F3E85FB530395207E9B6FC0873F802F6" box="[1185,1268,943,969]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30395207E802FC09758A0530" collectingDate="2015-09-03" collectingDateMax="2015-09-04" collectingDateMin="2015-09-03" collectionCode="CEMT" collectorName="Sector El Cacao &amp; R. Acconcia &amp; J. Gamez" country="Venezuela" county="Barinitas" elevation="1080" location="Andean foothills" municipality="Paguey" pageId="99" pageNumber="100" specimenCount="7" stateProvince="Barinas">
<emphasis id="A78D1B6230395207E802FC09727B02F7" bold="true" box="[1301,1399,942,968]" pageId="99" pageNumber="100">
<collectingRegion id="573D099230395207E802FC09727B02F7" box="[1301,1399,942,968]" country="Venezuela" name="Barinas" pageId="99" pageNumber="100">Barinas</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930395207EDD8FC75762502D3" box="[207,297,978,1004]" pageId="99" pageNumber="100">7 specs</specimenCount>
;
<collectingRegion id="573D099230395207EC21FC75769C02D3" box="[310,400,978,1004]" country="Venezuela" name="Bolivar" pageId="99" pageNumber="100">Bolívar</collectingRegion>
,
<collectingCounty id="7C27BFFC30395207EC8BFC75750502D3" box="[412,521,978,1004]" pageId="99" pageNumber="100">Barinitas</collectingCounty>
,
<collectingMunicipality id="75225D0A30395207EF01FC75756202D3" box="[534,622,978,1004]" pageId="99" pageNumber="100">Paguey</collectingMunicipality>
,
<collectorName id="380CA2A630395207EF6DFC75743702D3" box="[634,827,978,1004]" pageId="99" pageNumber="100">Sector El Cacao</collectorName>
,
<location id="902691AB30395207EE51FC75730302D3" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207EE51FC75730302D3" box="[838,1039,978,1004]" country="Venezuela" county="Barinitas" municipality="Paguey" name="Andean foothills" pageId="99" pageNumber="100" stateProvince="Barinas">Andean foothills</location>
;
<elevation id="1ED4204330395207E90CFC7573BA02D3" box="[1051,1206,978,1004]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="99" pageNumber="100" unit="m" value="1080.0">
<quantity id="52016A9530395207E90CFC75737902D3" box="[1051,1141,978,1004]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="99" pageNumber="100" unit="m" value="1080.0">1080 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030395207E9D3FC75727E02D3" box="[1220,1394,977,1004]" pageId="99" pageNumber="100" value="2015-09-03" valueMax="2015-09-04" valueMin="2015-09-03">
<collectingDate id="F103185830395207E9D3FC75727E02D3" box="[1220,1394,977,1004]" pageId="99" pageNumber="100" value="2015-09-03" valueMax="2015-09-04" valueMin="2015-09-03">34 Sep. 2015</collectingDate>
</date>
;
<collectorName id="380CA2A630395207EDAAFC5276400530" box="[189,332,1013,1039]" pageId="99" pageNumber="100">R. Acconcia</collectorName>
and
<collectorName id="380CA2A630395207EC92FC5276FD0530" box="[389,497,1013,1039]" pageId="99" pageNumber="100">J. Gámez</collectorName>
leg.;
<collectionCode id="F3E85FB530395207EF24FC52758A0530" box="[563,646,1013,1039]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30395207EFB0FC527667050D" collectingDate="1987-11" collectionCode="CEMT" collectorName="Joffre B." country="Venezuela" location="Monte Zerpa" pageId="99" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Merida">
<emphasis id="A78D1B6230395207EFB0FC5274080530" bold="true" box="[679,772,1013,1039]" pageId="99" pageNumber="100">
<collectingRegion id="573D099230395207EFB0FC5274080530" box="[679,772,1013,1039]" country="Venezuela" name="Merida" pageId="99" pageNumber="100">Merida</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930395207EE0BFC5274470530" box="[796,843,1013,1039]" count="1" pageId="99" pageNumber="100" type="male">1 ♂</specimenCount>
;
<location id="902691AB30395207EE40FC5274FE0530" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207EE40FC5274FE0530" box="[855,1010,1013,1039]" country="Venezuela" name="Monte Zerpa" pageId="99" pageNumber="100" stateProvince="Merida">Monte Zerpa</location>
,
<location id="902691AB30395207EEEAFC5273920530" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207EEEAFC5273920530" box="[1021,1182,1013,1039]" country="Venezuela" name="La Hechicera" pageId="99" pageNumber="100" stateProvince="Merida">La Hechicera</location>
;
<date id="E147E1B030395207E9BDFC52722B0530" box="[1194,1319,1013,1039]" pageId="99" pageNumber="100" value="1987-11">
<collectingDate id="F103185830395207E9BDFC52722B0530" box="[1194,1319,1013,1039]" pageId="99" pageNumber="100" value="1987-11">Nov. 1987</collectingDate>
</date>
;
<collectorName id="380CA2A630395207E824FC5277D4050C" pageId="99" pageNumber="100">Joffre B.</collectorName>
leg.;
<collectionCode id="F3E85FB530395207EC0DFBBF7667050D" box="[282,363,1048,1074]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30395207EC94FBBF737B050D" box="[387,1143,1048,1075]" collectingDate="1986-06" collectionCode="La Hechicera, CEMT" collectorName="Joffre B." country="Venezuela" location="Santa Rosa" pageId="99" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="Merida">
<specimenCount id="83FF0CF930395207EC94FBBF76BC050C" box="[387,432,1048,1075]" count="1" pageId="99" pageNumber="100" type="female">1 ♀</specimenCount>
;
<location id="902691AB30395207ECA8FBBF7549050C" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207ECA8FBBF7549050C" box="[447,581,1048,1075]" country="Venezuela" name="Santa Rosa" pageId="99" pageNumber="100" stateProvince="Merida">Santa Rosa</location>
,
<collectionCode id="F3E85FB530395207EF47FBBE75FD050D" box="[592,753,1048,1075]" pageId="99" pageNumber="100">La Hechicera</collectionCode>
;
<date id="E147E1B030395207EFEAFBBE747E050D" box="[765,882,1048,1075]" pageId="99" pageNumber="100" value="1986-06">
<collectingDate id="F103185830395207EFEAFBBE747E050D" box="[765,882,1048,1075]" pageId="99" pageNumber="100" value="1986-06">Jun. 1986</collectingDate>
</date>
;
<collectorName id="380CA2A630395207EE69FBBF74E8050C" box="[894,996,1048,1075]" pageId="99" pageNumber="100">Joffre B.</collectorName>
leg.;
<collectionCode id="F3E85FB530395207E931FBBF737B050D" box="[1062,1143,1048,1074]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30395207E998FBBF75780546" collectedFrom="human faeces" collectingDate="2009-07-08" collectionCode="CEMT" collectorName="D. Mora &amp; P. Colmenares &amp; M. Cordova &amp; M. Nunez" country="Venezuela" elevation="1999" latitude="8.6723585" location="Jaji" longLatPrecision="1" longitude="-71.41072" pageId="99" pageNumber="100" specimenCount="2" stateProvince="Sucre">
<specimenCount id="83FF0CF930395207E998FBBF73E4050C" box="[1167,1256,1048,1075]" pageId="99" pageNumber="100">2 specs</specimenCount>
;
<collectingRegion id="573D099230395207E9E3FBBF7236050D" box="[1268,1338,1048,1074]" country="Venezuela" name="Sucre" pageId="99" pageNumber="100">Sucre</collectingRegion>
,
<location id="902691AB30395207E851FBBF727F050D" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207E851FBBF727F050D" box="[1350,1395,1048,1074]" country="Venezuela" latitude="8.6723585" longLatPrecision="1" longitude="-71.41072" name="Jaji" pageId="99" pageNumber="100" stateProvince="Sucre">Jají</location>
;
<geoCoordinate id="F0CDA1B730395207EDAAFB9D76670569" box="[189,363,1082,1110]" degrees="8" direction="north" minutes="40" orientation="latitude" pageId="99" pageNumber="100" precision="1" seconds="20.49" value="8.6723585">8°4020.49″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730395207EC6EFB9D75360569" box="[377,570,1082,1110]" degrees="71" direction="west" minutes="24" orientation="longitude" pageId="99" pageNumber="100" precision="1" seconds="38.59" value="-71.41072">71°2438.59″ W</geoCoordinate>
;
<elevation id="1ED4204330395207EF5FFB9C75E9056A" box="[584,741,1083,1110]" metricMagnitude="3" metricUnit="m" metricValue="1.999" pageId="99" pageNumber="100" unit="m" value="1999.0">
<quantity id="52016A9530395207EF5FFB9C75AF0569" box="[584,675,1083,1110]" metricMagnitude="3" metricUnit="m" metricValue="1.999" pageId="99" pageNumber="100" unit="m" value="1999.0">1999 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030395207EFE2FB9C7472056A" box="[757,894,1083,1109]" pageId="99" pageNumber="100" value="2009-07-08">
<collectingDate id="F103185830395207EFE2FB9C7472056A" box="[757,894,1083,1109]" pageId="99" pageNumber="100" value="2009-07-08">8 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630395207EE9BFB9B74F90569" box="[908,1013,1084,1110]" pageId="99" pageNumber="100">D. Mora</collectorName>
,
<collectorName id="380CA2A630395207E915FB9B73A2056A" box="[1026,1198,1083,1110]" pageId="99" pageNumber="100">P. Colmenares</collectorName>
,
<collectorName id="380CA2A630395207E9ABFB9C724F056A" box="[1212,1347,1083,1109]" pageId="99" pageNumber="100">M.Córdova</collectorName>
and
<collectorName id="380CA2A630395207EDAAFBF8763E0546" box="[189,306,1119,1145]" pageId="99" pageNumber="100">M. Nuñez</collectorName>
leg.;
<collectedFrom id="26E5848D30395207EC65FBF9751A0547" box="[370,534,1118,1144]" pageId="99" pageNumber="100">human faeces</collectedFrom>
;
<collectionCode id="F3E85FB530395207EF36FBF875780546" box="[545,628,1119,1145]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30395207EF84FBF9766105A3" collectingDate="1985-07-15" collectionCode="CEMT" collectorName="Joffre B." country="Venezuela" location="La Flautera Palmira" pageId="99" pageNumber="100" specimenCount="2" specimenCount-female="2" stateProvince="Tachira">
<emphasis id="A78D1B6230395207EF84FBF975FA0546" bold="true" box="[659,758,1118,1145]" pageId="99" pageNumber="100">
<collectingRegion id="573D099230395207EF84FBF975FA0546" box="[659,758,1118,1145]" country="Venezuela" name="Tachira" pageId="99" pageNumber="100">Táchira</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930395207EE1AFBF8745C0546" box="[781,848,1119,1145]" count="2" pageId="99" pageNumber="100" type="female">2 ♀♀</specimenCount>
; “
<location id="902691AB30395207EE7FFBF873590547" LSID="urn:lsid:plazi:treatment:1D50766630395207EF3EFE4F7219070D:902691AB30395207EE7FFBF873590547" box="[872,1109,1118,1145]" country="Venezuela" name="La Flautera Palmira" pageId="99" pageNumber="100" stateProvince="Tachira">La Flautera Palmira</location>
” (?);
<date id="E147E1B030395207E982FBF9722B0546" box="[1173,1319,1118,1145]" pageId="99" pageNumber="100" value="1985-07-15">
<collectingDate id="F103185830395207E982FBF9722B0546" box="[1173,1319,1118,1145]" pageId="99" pageNumber="100" value="1985-07-15">15 Jul. 1985</collectingDate>
</date>
;
<collectorName id="380CA2A630395207E824FBF977D405A3" pageId="99" pageNumber="100">Joffre B.</collectorName>
leg.;
<collectionCode id="F3E85FB530395207EC0DFB25766105A3" box="[282,365,1154,1180]" pageId="99" pageNumber="100">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30395207EDAAFB6474FB0443" pageId="99" pageNumber="100" type="distribution">
<paragraph id="9546C77030395207EDAAFB64765A05E2" blockId="99.[189,342,1219,1245]" box="[189,342,1219,1245]" pageId="99" pageNumber="100">
<heading id="CE0E701C30395207EDAAFB64765A05E2" bold="true" box="[189,342,1219,1245]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A78D1B6230395207EDAAFB64765A05E2" bold="true" box="[189,342,1219,1245]" pageId="99" pageNumber="100">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030395207EDAAFB5576E60433" blockId="99.[189,490,1266,1292]" box="[189,490,1266,1292]" pageId="99" pageNumber="100">
<collectingCountry id="EDEE87E030395207EDAAFB55763D0433" box="[189,305,1266,1292]" name="Colombia" pageId="99" pageNumber="100">Colombia</collectingCountry>
and
<collectingCountry id="EDEE87E030395207EC7EFB5576EA0433" box="[361,486,1266,1292]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030395207EDABFA9476E10472" blockId="99.[188,493,1331,1357]" box="[188,493,1331,1357]" pageId="99" pageNumber="100">
<heading id="CE0E701C30395207EDABFA9476E10472" bold="true" box="[188,493,1331,1357]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A78D1B6230395207EDABFA9476E10472" bold="true" box="[188,493,1331,1357]" pageId="99" pageNumber="100">
Subregions of
<collectingCountry id="EDEE87E030395207EC78FA9476E10472" box="[367,493,1331,1357]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030395207EDAAFAC574FB0443" blockId="99.[189,1015,1378,1404]" box="[189,1015,1378,1404]" pageId="99" pageNumber="100">
Coastal mainland, Plains, Andes mountains and
<collectingRegion id="573D099230395207EFE5FAC5745E0443" box="[754,850,1378,1404]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="99" pageNumber="100">Oriental</collectingRegion>
<collectingRegion id="573D099230395207EE4DFAC574910443" box="[858,925,1378,1404]" country="New Zealand" name="West Coast" pageId="99" pageNumber="100">Coast</collectingRegion>
Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30395207EDAAFA047219070D" pageId="99" pageNumber="100" type="reference_group">
<paragraph id="9546C77030395207EDAAFA0476A80482" blockId="99.[189,420,1443,1469]" box="[189,420,1443,1469]" pageId="99" pageNumber="100">
<heading id="CE0E701C30395207EDAAFA0476A80482" bold="true" box="[189,420,1443,1469]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A78D1B6230395207EDAAFA0476A80482" bold="true" box="[189,420,1443,1469]" pageId="99" pageNumber="100">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030395207EDAAFA757219070D" blockId="99.[189,1399,1489,1587]" pageId="99" pageNumber="100">
<bibRefCitation id="F168BA8130395207EDAAFA75768404D4" author="Gillet J. J. E." box="[189,392,1489,1516]" pageId="99" pageNumber="100" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E030395207EC89FA75751204D3" box="[414,542,1490,1516]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130395207EF76FA75747304D3" author="Luederwaldt H." box="[609,895,1490,1516]" pageId="99" pageNumber="100" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 109</bibRefCitation>
(
<collectingCountry id="EDEE87E030395207EE82FA75731904D3" box="[917,1045,1490,1516]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130395207E94FFA75727A04D3" author="Blackwelder R. E." box="[1112,1398,1490,1516]" pageId="99" pageNumber="100" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
(
<collectingCountry id="EDEE87E030395207EDD2FA5276490730" box="[197,325,1525,1551]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130395207EC60FA5275C8072F" author="Vulcano M. A. &amp; Pereira F. S." box="[375,708,1525,1552]" pageId="99" pageNumber="100" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 585</bibRefCitation>
(
<collectingCountry id="EDEE87E030395207EFC7FA52745C0730" box="[720,848,1525,1551]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130395207EE95FA5273480730" author="Krajcik M." box="[898,1092,1525,1551]" pageId="99" pageNumber="100" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 91</bibRefCitation>
(
<collectingCountry id="EDEE87E030395207E946FA5273DD0730" box="[1105,1233,1525,1551]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130395207E814FA527536070C" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." pageId="99" pageNumber="100" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello 2021: 7</bibRefCitation>
, 33 (
<collectingCountry id="EDEE87E030395207EF64F9BF75FE070D" box="[627,754,1560,1586]" name="Venezuela" pageId="99" pageNumber="100">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230395207EFEBF9BF745A070D" box="[764,854,1560,1586]" country="Venezuela" name="Aragua" pageId="99" pageNumber="100">Aragua</collectingRegion>
;
<collectingRegion id="573D099230395207EE76F9BF74B2070D" box="[865,958,1560,1586]" country="Venezuela" name="Barinas" pageId="99" pageNumber="100">Barinas</collectingRegion>
;
<collectingRegion id="573D099230395207EEDDF9BF732F070D" box="[970,1059,1560,1586]" country="Venezuela" name="Merida" pageId="99" pageNumber="100">Mérida</collectingRegion>
;
<collectingRegion id="573D099230395207E938F9BF737E070D" box="[1071,1138,1560,1586]" country="Venezuela" name="Sucre" pageId="99" pageNumber="100">Sucre</collectingRegion>
and
<collectingRegion id="573D099230395207E9BDF9BF7205070D" box="[1194,1289,1560,1586]" country="Venezuela" name="Tachira" pageId="99" pageNumber="100">Táchira</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>