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<mods:title id="CC10E535D6DBDE8F38245012209B0D06">A new species of genus Psilocladia Warren from India, bridging the boundaries of Africa and Asia (Lepidoptera: Geometridae: Ennominae)</mods:title>
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<mods:namePart id="D17CC8472AD0A8DF4941FC08888FD7DD">Lenka, Rajesh</mods:namePart>
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<mods:namePart id="DEC5A677C3F97ADA1E1BB78DAFC5D2A1">Chatterjee, Pallab</mods:namePart>
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<mods:namePart id="3DE25B7C6A129D3C3B11C758DB4BDC2F">Singh, Navneet</mods:namePart>
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<mods:affiliation id="682135233A6ADE6BE3F1F39F252137B1">Zoological Survey of India, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
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<emphasis id="01FAEAC75C11FFD7E4F594F2A463BDAB" bold="true" box="[151,367,1438,1465]" italics="true" pageId="1" pageNumber="317">Psilocladia indica</emphasis>
M. Singh, Lenka &amp; N. Singh
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(
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,
<figureCitation id="ABB52A505C11FFD7E575948BA4A8BE13" box="[279,420,1511,1537]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="1" pageNumber="317">910, 1213</figureCitation>
)
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<emphasis id="01FAEAC75C11FFD7E4F59742A475BE5A" bold="true" box="[151,377,1582,1608]" pageId="1" pageNumber="317">Material examined</emphasis>
:
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:
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, Alt.
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leg.
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(
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):
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<location id="3651600E5C11FFD7E6D197D3A6A7BECB" LSID="urn:lsid:plazi:treatment:BB2787C35C11FFD1E4F594F2A6F2B9C3:3651600E5C11FFD7E6D197D3A6A7BECB" box="[691,939,1727,1753]" country="India" latitude="27.1041" longLatPrecision="6" longitude="92.1122" municipality="Kalakthang" name="West Kameng district" pageId="1" pageNumber="317" stateProvince="Arunachal Pradesh">West Kameng district</location>
,
<specimenCount id="2588FD5C5C11FFD7E7D497D3A6EDBECB" box="[950,993,1727,1753]" count="1" pageId="1" pageNumber="317" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="D355ACAF5C11FFD7E79297D3A178BECB" box="[1008,1140,1727,1753]" pageId="1" pageNumber="317">Kalakthang</collectingMunicipality>
,
<date id="473010155C11FFD7E0E297D3A1F6BECB" box="[1152,1274,1727,1753]" pageId="1" pageNumber="317" value="2022-06-10">
<collectingDate id="5774E9FD5C11FFD7E0E297D3A1F6BECB" box="[1152,1274,1727,1753]" pageId="1" pageNumber="317" value="2022-06-10">10.vi.2022</collectingDate>
</date>
, Alt.
<quantity id="F4769B305C11FFD7E15A97D2A09BBECB" box="[1336,1431,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="3.453" pageId="1" pageNumber="317" unit="m" value="345.3">
<elevation id="B8A3D1E65C11FFD7E15A97D2A09BBECB" box="[1336,1431,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="3.453" pageId="1" pageNumber="317" unit="m" value="345.3">345.3 m</elevation>
</quantity>
,
<geoCoordinate id="56BA50125C11FFD7E4F5978EA402BEEF" box="[151,270,1762,1789]" degrees="27.1041" direction="north" orientation="latitude" pageId="1" pageNumber="317" precision="5" value="27.1041">27.1041ᵒN</geoCoordinate>
<geoCoordinate id="56BA50125C11FFD7E57A978EA483BEEF" box="[280,399,1762,1789]" degrees="92.1122" direction="east" orientation="longitude" pageId="1" pageNumber="317" precision="5" value="92.1122">92.1122ᵒE</geoCoordinate>
,
<collectorName id="9E7B53035C11FFD7E5FF978FA70CBEEF" box="[413,512,1763,1789]" pageId="1" pageNumber="317">D. Mitra</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="83E63C885C11FFD7E627978FA431BF33" collectingDate="2023-06-12" collectorName="P. Chatterjee" country="India" county="Tenga" elevation="1282" latitude="27.2196" location="Husi" longLatPrecision="6" longitude="92.6098" municipality="Husi" pageId="1" pageNumber="317" specimenCount="2" specimenCount-female="2" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="2588FD5C5C11FFD7E627978FA77EBEEF" box="[581,626,1763,1789]" count="2" pageId="1" pageNumber="317" type="female">2 ♀</specimenCount>
,
<collectingCounty id="DA504E595C11FFD7E6E0978FA7C5BEEF" box="[642,713,1763,1789]" pageId="1" pageNumber="317">Tenga</collectingCounty>
,
<collectingMunicipality id="D355ACAF5C11FFD7E6B5978FA600BEEF" box="[727,780,1763,1789]" pageId="1" pageNumber="317">Husi</collectingMunicipality>
gaon,
<date id="473010155C11FFD7E73C978FA6D4BEEF" box="[862,984,1763,1789]" pageId="1" pageNumber="317" value="2023-06-12">
<collectingDate id="5774E9FD5C11FFD7E73C978FA6D4BEEF" box="[862,984,1763,1789]" pageId="1" pageNumber="317" value="2023-06-12">12.vi.2023</collectingDate>
</date>
, Alt.
<quantity id="F4769B305C11FFD7E078978FA178BEEF" box="[1050,1140,1763,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="1" pageNumber="317" unit="m" value="1282.0">
<elevation id="B8A3D1E65C11FFD7E078978FA178BEEF" box="[1050,1140,1763,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="1" pageNumber="317" unit="m" value="1282.0">1282 m</elevation>
</quantity>
,
<geoCoordinate id="56BA50125C11FFD7E0E0978EA1F6BEEF" box="[1154,1274,1762,1789]" degrees="27.2196" direction="north" orientation="latitude" pageId="1" pageNumber="317" precision="5" value="27.2196">27.2196ᵒN</geoCoordinate>
<geoCoordinate id="56BA50125C11FFD7E166978EA076BEEF" box="[1284,1402,1762,1789]" degrees="92.6098" direction="east" orientation="longitude" pageId="1" pageNumber="317" precision="5" value="92.6098">92.6098ᵒE</geoCoordinate>
,
<collectorName id="9E7B53035C11FFD7E1EB978FA406BF33" pageId="1" pageNumber="317">P. Chatterjee</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="83E63C885C11FFD7E525966BA00ABF33" box="[327,1286,1798,1825]" collectingDate="2023-06-15" collectorName="P. Chatterjee" country="India" county="Tenga" elevation="1624" latitude="27.1995" location="Helipad" longLatPrecision="6" longitude="92.5019" municipality="Helipad" pageId="1" pageNumber="317" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="2588FD5C5C11FFD7E525966BA47CBF33" box="[327,368,1799,1825]" count="1" pageId="1" pageNumber="317" type="female">1 ♀</specimenCount>
,
<collectingCounty id="DA504E595C11FFD7E51F966BA4C8BF33" box="[381,452,1799,1825]" pageId="1" pageNumber="317">Tenga</collectingCounty>
,
<collectingMunicipality id="D355ACAF5C11FFD7E5AC966BA726BF33" box="[462,554,1799,1825]" pageId="1" pageNumber="317">Helipad</collectingMunicipality>
,
<date id="473010155C11FFD7E656966AA7A2BF33" box="[564,686,1798,1825]" pageId="1" pageNumber="317" value="2023-06-15">
<collectingDate id="5774E9FD5C11FFD7E656966AA7A2BF33" box="[564,686,1798,1825]" pageId="1" pageNumber="317" value="2023-06-15">15.vi.2023</collectingDate>
</date>
, Alt.
<quantity id="F4769B305C11FFD7E68A966BA632BF33" box="[744,830,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.624" pageId="1" pageNumber="317" unit="m" value="1624.0">
<elevation id="B8A3D1E65C11FFD7E68A966BA632BF33" box="[744,830,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.624" pageId="1" pageNumber="317" unit="m" value="1624.0">1624 m</elevation>
</quantity>
,
<geoCoordinate id="56BA50125C11FFD7E72B966AA6CCBF33" box="[841,960,1798,1825]" degrees="27.1995" direction="north" orientation="latitude" pageId="1" pageNumber="317" precision="5" value="27.1995">27.1995ᵒN</geoCoordinate>
<geoCoordinate id="56BA50125C11FFD7E7A4966AA131BF33" box="[966,1085,1798,1825]" degrees="92.5019" direction="east" orientation="longitude" pageId="1" pageNumber="317" precision="5" value="92.5019">92.5019ᵒE</geoCoordinate>
,
<collectorName id="9E7B53035C11FFD7E02A966BA1D8BF33" box="[1096,1236,1799,1825]" pageId="1" pageNumber="317">P. Chatterjee</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="83E63C885C11FFD7E173966BA10CBF57" collectingDate="2023-07-24" collectorName="P. Chatterjee" country="India" county="Dirang" elevation="1741" latitude="27.3316" location="Sagar" longLatPrecision="6" longitude="92.2661" municipality="Sagar" pageId="1" pageNumber="317" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="2588FD5C5C11FFD7E173966BA036BF33" box="[1297,1338,1799,1825]" count="1" pageId="1" pageNumber="317" type="female">1 ♀</specimenCount>
,
<collectingCounty id="DA504E595C11FFD7E125966BA094BF33" box="[1351,1432,1799,1825]" pageId="1" pageNumber="317">Dirang</collectingCounty>
,
<collectingMunicipality id="D355ACAF5C11FFD7E4F59647A5DBBF57" box="[151,215,1835,1861]" pageId="1" pageNumber="317">Sagar</collectingMunicipality>
basti,
<date id="473010155C11FFD7E5439647A4AFBF57" box="[289,419,1835,1861]" pageId="1" pageNumber="317" value="2023-07-24">
<collectingDate id="5774E9FD5C11FFD7E5439647A4AFBF57" box="[289,419,1835,1861]" pageId="1" pageNumber="317" value="2023-07-24">24.vii.2023</collectingDate>
</date>
, Alt.
<quantity id="F4769B305C11FFD7E5BD9647A73ABF57" box="[479,566,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.741" pageId="1" pageNumber="317" unit="m" value="1741.0">
<elevation id="B8A3D1E65C11FFD7E5BD9647A73ABF57" box="[479,566,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.741" pageId="1" pageNumber="317" unit="m" value="1741.0">1741 m</elevation>
</quantity>
,
<geoCoordinate id="56BA50125C11FFD7E6239646A7B5BF57" box="[577,697,1834,1861]" degrees="27.3316" direction="north" orientation="latitude" pageId="1" pageNumber="317" precision="5" value="27.3316">27.3316ᵒN</geoCoordinate>
<geoCoordinate id="56BA50125C11FFD7E6A29646A63ABF57" box="[704,822,1834,1861]" degrees="92.2661" direction="east" orientation="longitude" pageId="1" pageNumber="317" precision="5" value="92.2661">92.2661ᵒE</geoCoordinate>
,
<collectorName id="9E7B53035C11FFD7E7209647A6C3BF57" box="[834,975,1835,1861]" pageId="1" pageNumber="317">P. Chatterjee</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C11FFD4E4A59622A6E7BAE3" lastPageId="2" lastPageNumber="318" pageId="1" pageNumber="317" type="diagnosis">
<paragraph id="333136D55C11FFD4E4A59622A6BCB8EB" blockId="1.[151,1437,1582,2041]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="318" pageId="1" pageNumber="317">
<emphasis id="01FAEAC75C11FFD7E4A59622A436BF7A" bold="true" box="[199,314,1870,1896]" pageId="1" pageNumber="317">Diagnosis</emphasis>
(
<figureCitation id="ABB52A505C11FFD7E52B9623A4B9BF7B" box="[329,437,1871,1897]" captionStart="FIGURES 14" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[207,1380,191,1909]" captionTargetId="figure-29@3.[196,1391,181,1919]" captionTargetPageId="3" captionText="FIGURES 14. Adults of Psilocladia indica sp. nov.: 1, Male, Holotype (Upper Singchung, Arunachal Pradesh); 2, Female, Paratype (Karadei, Odisha); 3, Male, Paratype (Karadei, Odisha); 4, Female, Paratype (Husigaon, Arunachal Pradesh). a. dorsal sides; b. ventral sides." figureDoi="http://doi.org/10.5281/zenodo.14954390" httpUri="https://zenodo.org/record/14954390/files/figure.png" pageId="1" pageNumber="317">Figs. 14</figureCitation>
): Externally,
<taxonomicName id="F48E4D565C11FFD7E62A9623A7A4BF7B" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[584,680,1871,1897]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C11FFD7E62A9623A7A4BF7B" box="[584,680,1871,1897]" italics="true" pageId="1" pageNumber="317">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C11FFD7E6CC9623A60BBF7B" bold="true" box="[686,775,1871,1897]" pageId="1" pageNumber="317">
<taxonomicNameLabel id="1AC957BC5C11FFD7E6CC9623A60BBF7B" box="[686,775,1871,1897]" pageId="1" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble
<taxonomicName id="F48E4D565C11FFD7E71A9623A6F1BF7B" authorityName="Warren" authorityYear="1898" box="[888,1021,1871,1897]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C11FFD7E71A9623A6F1BF7B" box="[888,1021,1871,1897]" italics="true" pageId="1" pageNumber="317">P. obliquata</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C11FFD7E06E9623A143BF7B" box="[1036,1103,1870,1897]" captionStart="FIGURES 58" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[199,1388,191,1899]" captionTargetId="figure-19@4.[189,1398,181,1920]" captionTargetPageId="4" captionText="FIGURES 58. Adults of Psilocladia spp.: 5, P. obliquata, Male, Holotype (South Africa); 6, P. repudiosa, Male, Holotype (Cameroon); 7, P. diaereta, Male, Holotype (Kenya); 8, P. loxostigma, Male, Holotype (Congo). a. dorsal sides; b. ventral sides (for figures 58 © NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.14954392" httpUri="https://zenodo.org/record/14954392/files/figure.png" pageId="1" pageNumber="317">Fig. 5</figureCitation>
). However, the new species is significantly smaller in size with a forewing length of
<quantity id="F4769B305C11FFD7E68E961FA668BF9F" box="[748,868,1907,1933]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="317" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
compared to approximately
<quantity id="F4769B305C11FFD7E0C3961FA1E2BF9F" box="[1185,1262,1907,1933]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="317" unit="mm" value="18.0">18 mm</quantity>
in
<taxonomicName id="F48E4D565C11FFD7E172961FA099BF9F" authorityName="Warren" authorityYear="1898" box="[1296,1429,1907,1933]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C11FFD7E172961FA099BF9F" box="[1296,1429,1907,1933]" italics="true" pageId="1" pageNumber="317">P. obliquata</emphasis>
</taxonomicName>
.
<taxonomicName id="F48E4D565C11FFD7E4F596FBA46FBFA3" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[151,355,1943,1969]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C11FFD7E4F596FBA46FBFA3" box="[151,355,1943,1969]" italics="true" pageId="1" pageNumber="317">Psilocladia indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C11FFD7E50B96FBA4CDBFA3" bold="true" box="[361,449,1943,1969]" pageId="1" pageNumber="317">
<taxonomicNameLabel id="1AC957BC5C11FFD7E50B96FBA4CDBFA3" box="[361,449,1943,1969]" pageId="1" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinct by the forewing with a broad, obliquely straight medial line from inner margin to lower edge of the cell, then narrower towards the costa with a distinctive inward curve within the cell and a conspicuous dark brown triangular patch on the inner margin; hindwing with a more prominent antemedial line. Whereas in
<taxonomicName id="F48E4D565C12FFD4E57D91FBA4AAB8A3" authorityName="Warren" authorityYear="1898" box="[287,422,151,177]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C12FFD4E57D91FBA4AAB8A3" box="[287,422,151,177]" italics="true" pageId="2" pageNumber="318">P. obliquata</emphasis>
</taxonomicName>
, medial line of forewing is partially double between the inner margin and the lower edge of the cell and subsequently running obliquely straight to the costa; also a postmedial series of dark brown spots on each vein from the costa to CuA2, which is absent in
<taxonomicName id="F48E4D565C12FFD4E68C91B3A643B8EB" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[750,847,223,249]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C12FFD4E68C91B3A643B8EB" box="[750,847,223,249]" italics="true" pageId="2" pageNumber="318">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C12FFD4E73491B3A6BCB8EB" bold="true" box="[854,944,223,249]" pageId="2" pageNumber="318">
<taxonomicNameLabel id="1AC957BC5C12FFD4E73491B3A6BCB8EB" box="[854,944,223,249]" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="333136D55C12FFD4E4A5906FA4ABB99B" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
In male genitalia,
<taxonomicName id="F48E4D565C12FFD4E5F9906FA70CB90F" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[411,512,259,285]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C12FFD4E5F9906FA70CB90F" box="[411,512,259,285]" italics="true" pageId="2" pageNumber="318">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C12FFD4E669906FA764B90F" bold="true" box="[523,616,259,285]" pageId="2" pageNumber="318">
<taxonomicNameLabel id="1AC957BC5C12FFD4E669906FA764B90F" box="[523,616,259,285]" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="ABB52A505C12FFD4E618906FA7FBB90F" box="[634,759,259,285]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Figs. 910</figureCitation>
) is distinct from
<taxonomicName id="F48E4D565C12FFD4E7A1906FA141B90F" authorityName="Warren" authorityYear="1898" box="[963,1101,259,285]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C12FFD4E7A1906FA141B90F" box="[963,1101,259,285]" italics="true" pageId="2" pageNumber="318">P. obliquata</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C12FFD4E002906FA1BFB90F" box="[1120,1203,259,285]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Fig. 11</figureCitation>
) by its symmetrical pair of furca, arising straightly from their bulbous bases, as well as a markedly longer uncus and a shorter medial process of the gnathos. In contrast, male genitalia of
<taxonomicName id="F48E4D565C12FFD4E6939027A675B977" authorityName="Warren" authorityYear="1898" box="[753,889,331,357]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C12FFD4E6939027A675B977" box="[753,889,331,357]" italics="true" pageId="2" pageNumber="318">P. obliquata</emphasis>
</taxonomicName>
bear basely curved, asymmetrical furca: the left one significantly longer.
</paragraph>
<paragraph id="333136D55C12FFD4E4A590FFA733BA0B" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
In the female genitalia (
<figureCitation id="ABB52A505C12FFD4E5AC90FFA759B9BF" box="[462,597,403,429]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Figs. 1213</figureCitation>
),
<taxonomicName id="F48E4D565C12FFD4E60A90FFA7C5B9BF" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[616,713,403,429]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C12FFD4E60A90FFA7C5B9BF" box="[616,713,403,429]" italics="true" pageId="2" pageNumber="318">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C12FFD4E6AD90FFA62BB9BF" bold="true" box="[719,807,403,429]" pageId="2" pageNumber="318">
<taxonomicNameLabel id="1AC957BC5C12FFD4E6AD90FFA62BB9BF" box="[719,807,403,429]" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinct from
<taxonomicName id="F48E4D565C12FFD4E7B990FFA16CB9BF" authorityName="Warren" authorityYear="1898" box="[987,1120,403,429]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C12FFD4E7B990FFA16CB9BF" box="[987,1120,403,429]" italics="true" pageId="2" pageNumber="318">P. obliquata</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C12FFD4E00D90FFA1B3B9BF" box="[1135,1215,403,429]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Fig. 14</figureCitation>
) by its significantly larger signum, which has a small medial concavity and transverse ends terminating in spines, giving it a somewhat groundnut-shaped appearance. In
<taxonomicName id="F48E4D565C12FFD4E67190B7A795B9E7" authorityName="Warren" authorityYear="1898" box="[531,665,475,501]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C12FFD4E67190B7A795B9E7" box="[531,665,475,501]" italics="true" pageId="2" pageNumber="318">P. obliquata</emphasis>
</taxonomicName>
, the signum is oval, distinctly smaller, with a large medial concavity and small spines around its periphery.
</paragraph>
<paragraph id="333136D55C12FFD4E4A5934FA6E7BAE3" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
Furthermore,
<taxonomicName id="F48E4D565C12FFD4E506934FA4C5BA2F" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[356,457,547,573]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C12FFD4E506934FA4C5BA2F" box="[356,457,547,573]" italics="true" pageId="2" pageNumber="318">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C12FFD4E5B1934FA73CBA2F" bold="true" box="[467,560,547,573]" pageId="2" pageNumber="318">
<taxonomicNameLabel id="1AC957BC5C12FFD4E5B1934FA73CBA2F" box="[467,560,547,573]" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinct from its other three congeners,
<taxonomicName id="F48E4D565C12FFD4E043934FA1A3BA2F" authorityName="Prout" authorityYear="1915" box="[1057,1199,547,573]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="repudiosa">
<emphasis id="01FAEAC75C12FFD4E043934FA1A3BA2F" box="[1057,1199,547,573]" italics="true" pageId="2" pageNumber="318">P. repudiosa</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C12FFD4E0A3934FA005BA2F" box="[1217,1289,547,573]" captionStart="FIGURES 58" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[199,1388,191,1899]" captionTargetId="figure-19@4.[189,1398,181,1920]" captionTargetPageId="4" captionText="FIGURES 58. Adults of Psilocladia spp.: 5, P. obliquata, Male, Holotype (South Africa); 6, P. repudiosa, Male, Holotype (Cameroon); 7, P. diaereta, Male, Holotype (Kenya); 8, P. loxostigma, Male, Holotype (Congo). a. dorsal sides; b. ventral sides (for figures 58 © NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.14954392" httpUri="https://zenodo.org/record/14954392/files/figure.png" pageId="2" pageNumber="318">Fig. 6</figureCitation>
),
<taxonomicName id="F48E4D565C12FFD4E143934FA090BA2F" authorityName="Prout" authorityYear="1923" box="[1313,1436,547,573]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C12FFD4E143934FA090BA2F" box="[1313,1436,547,573]" italics="true" pageId="2" pageNumber="318">P. diaereta</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C12FFD4E4FD932BA5EABA73" box="[159,230,583,609]" captionStart="FIGURES 58" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[199,1388,191,1899]" captionTargetId="figure-19@4.[189,1398,181,1920]" captionTargetPageId="4" captionText="FIGURES 58. Adults of Psilocladia spp.: 5, P. obliquata, Male, Holotype (South Africa); 6, P. repudiosa, Male, Holotype (Cameroon); 7, P. diaereta, Male, Holotype (Kenya); 8, P. loxostigma, Male, Holotype (Congo). a. dorsal sides; b. ventral sides (for figures 58 © NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.14954392" httpUri="https://zenodo.org/record/14954392/files/figure.png" pageId="2" pageNumber="318">Fig. 7</figureCitation>
), and
<taxonomicName id="F48E4D565C12FFD4E54C932BA4CBBA73" authorityName="Prout" authorityYear="1915" box="[302,455,583,609]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C12FFD4E54C932BA4CBBA73" box="[302,455,583,609]" italics="true" pageId="2" pageNumber="318">P. loxostigma</emphasis>
</taxonomicName>
(
<figureCitation id="ABB52A505C12FFD4E5BA932BA711BA73" box="[472,541,583,609]" captionStart="FIGURES 58" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[199,1388,191,1899]" captionTargetId="figure-19@4.[189,1398,181,1920]" captionTargetPageId="4" captionText="FIGURES 58. Adults of Psilocladia spp.: 5, P. obliquata, Male, Holotype (South Africa); 6, P. repudiosa, Male, Holotype (Cameroon); 7, P. diaereta, Male, Holotype (Kenya); 8, P. loxostigma, Male, Holotype (Congo). a. dorsal sides; b. ventral sides (for figures 58 © NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.14954392" httpUri="https://zenodo.org/record/14954392/files/figure.png" pageId="2" pageNumber="318">Fig. 8</figureCitation>
), by its dark brown wings that are basally infused with white. In
<taxonomicName id="F48E4D565C12FFD4E16D932BA07EBA73" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[1295,1394,583,609]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C12FFD4E16D932BA07EBA73" box="[1295,1394,583,609]" italics="true" pageId="2" pageNumber="318">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C12FFD4E119932BA5C4BA97" bold="true" pageId="2" pageNumber="318">
<taxonomicNameLabel id="1AC957BC5C12FFD4E119932BA5C4BA97" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the outer margin of the hindwing exhibits reduced crenulation with a protruded apex at Rs. In contrast,
<taxonomicName id="F48E4D565C12FFD4E1EA9307A40ABABB" authorityName="Prout" authorityYear="1915" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="repudiosa">
<emphasis id="01FAEAC75C12FFD4E1EA9307A40ABABB" italics="true" pageId="2" pageNumber="318">P. repudiosa</emphasis>
</taxonomicName>
displays less extended Rs into a tail but has deep crenulations along the outer margin. In
<taxonomicName id="F48E4D565C12FFD4E09693E3A060BABB" authorityName="Prout" authorityYear="1923" box="[1268,1388,655,681]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C12FFD4E09693E3A060BABB" box="[1268,1388,655,681]" italics="true" pageId="2" pageNumber="318">P. diaereta</emphasis>
</taxonomicName>
and
<taxonomicName id="F48E4D565C12FFD4E4F593DFA421BADF" authorityName="Prout" authorityYear="1915" box="[151,301,691,717]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C12FFD4E4F593DFA421BADF" box="[151,301,691,717]" italics="true" pageId="2" pageNumber="318">P. loxostigma</emphasis>
</taxonomicName>
, there is an excision between Rs and M3, which is more pronounced without a medial protrusion in
<taxonomicName id="F48E4D565C12FFD4E1EA93DFA5FFBAE3" authorityName="Prout" authorityYear="1923" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C12FFD4E1EA93DFA5FFBAE3" italics="true" pageId="2" pageNumber="318">P. diaereta</emphasis>
</taxonomicName>
and shallower with a noticeable medial protrusion in
<taxonomicName id="F48E4D565C12FFD4E72C93BBA6E9BAE3" authorityName="Prout" authorityYear="1915" box="[846,997,727,753]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="318" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C12FFD4E72C93BBA6E9BAE3" box="[846,997,727,753]" italics="true" pageId="2" pageNumber="318">P. loxostigma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C12FFD1E4A59396A18FB99B" lastPageId="7" lastPageNumber="323" pageId="2" pageNumber="318" type="description">
<paragraph id="333136D55C12FFD4E4A59396A43FBC27" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
<emphasis id="01FAEAC75C12FFD4E4A59396A457BB06" bold="true" box="[199,347,762,788]" pageId="2" pageNumber="318">Description:</emphasis>
<emphasis id="01FAEAC75C12FFD4E5069397A4ADBB07" box="[356,417,763,789]" italics="true" pageId="2" pageNumber="318">Head</emphasis>
: Antennae simple and lamellate in both sexes. The frons yellow. Labial palpi are porrect and reach beyond the frons, with terminal segment gently drooping. The vertex is covered with long yellow scales. Ocellus well-developed and marked by pale yellow piliform scales.
<emphasis id="01FAEAC75C12FFD4E7F5922FA6E4BB4F" box="[919,1000,835,861]" italics="true" pageId="2" pageNumber="318">Thorax</emphasis>
: The notum covered with smooth pale white scales. Patagia pale brown, while tegulae are white. The pleuron and sternum are pale brown. Legs feature a bunch of hair-pencil scaled basally on the fore tibia. The hind tibia has two pairs of spurs. The forewing length is
<quantity id="F4769B305C12FFD4E4F592C3A41DBBDB" box="[151,273,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="2" pageNumber="318" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
in males and
<quantity id="F4769B305C12FFD4E5D092C3A720BBDB" box="[434,556,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="318" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
in females. Wings have a pale brown ground color, infused with grey scales. The forewing is elongated with a straight costa up to two-thirds then well arched with a subacute apex. The outer margin is more or less straight, the anal margin round and somewhat lobed, and the inner margin is slightly sinuous in the middle.
</paragraph>
<paragraph id="333136D55C12FFD4E4AD9553A5E5BD23" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">Forewing with transverse lines reddish brown; the antemedial line is short obliquely straight running from the inner margin to the edge of the cell; medial line is broad between the inner margin and the lower edge of the cell, then appears slender with an inward curve between the lower edge of the cell and costal margin. There is a broad prominent triangular or trapezoid reddish-brown patch at the inner margin of submarginal area. Sometimes an indistinct postmedial line, dark and highly curved near costal margin, runs obliquely straight to join the reddish-brown triangular or trapezoid patch; the basal part along the inner area is infused with white; cilia are brown.</paragraph>
<paragraph id="333136D55C12FFD4E4A59457A090BE1B" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
Hindwing with costal margin straight, and the outer margin is produced at Rs. The medial and postmedial lines are more or less obliquely straight, with the latter originates from a small, dark brown triangular patch at the costa and slightly curved just before meeting the inner margin; cilia are the same as forewing. Discal spot absents in both wings. The underside of both wings is dark brown but in forewing, the area from basal to the outer margin above M3 is faded. The transverse line is darker. The hindwing has a postmedial line partially doubled from inner margin to CuA2.
<emphasis id="01FAEAC75C12FFD4E48A9483A45EBE1B" box="[232,338,1519,1545]" italics="true" pageId="2" pageNumber="318">Abdomen</emphasis>
: The dorsum is pale brown, mixed with pale yellow and black scales. The anal tuft is pale yellow.
</paragraph>
<paragraph id="333136D55C12FFD4E4A5977FA4CBBEF3" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
<emphasis id="01FAEAC75C12FFD4E4A5977FA460BE3F" box="[199,364,1555,1581]" italics="true" pageId="2" pageNumber="318">Male genitalia</emphasis>
(
<figureCitation id="ABB52A505C12FFD4E518977FA4FABE3F" box="[378,502,1555,1581]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Figs. 910</figureCitation>
): The uncus is broad at the base, elongated, slender with a pointed apex. The lateral arms of the gnathos are weakly sclerotized, and the median process is short with numerous setal hairs. The valva is simple, broad, with a uniformly convexed ventral margin and an apex that is somewhat acute with slightly pointed apices; the costa is broad, and slightly incurved medially. A pair of symmetrical furca, arise straightly from the bulbous base. The aedeagus is short and slender with an apically acute extension; the vesica is membranous and has a weakly sclerotized patch.
</paragraph>
<paragraph id="333136D55C12FFD4E4A59787A7B2BFAB" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="318">
<emphasis id="01FAEAC75C12FFD4E4A59787A485BF17" box="[199,393,1771,1797]" italics="true" pageId="2" pageNumber="318">Female genitalia</emphasis>
(
<figureCitation id="ABB52A505C12FFD4E5F99787A724BF17" box="[411,552,1771,1797]" captionStart="FIGURES 914" captionStartId="5.[151,264,1905,1929]" captionTargetBox="[215,1374,156,1880]" captionTargetId="figure-29@5.[205,1383,151,1889]" captionTargetPageId="5" captionText="FIGURES 914. Genitalia structure of Psilocladia spp.: 9, P. indica sp. nov., male, HT (Upper Singchung); 10, P. indica sp. nov., male, PT (Karadei); 11, P. obliquata, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12, P. indica sp. nov., female (Karadei); 13, P. indica sp. nov., female (Husigaon); 14, P. obliquata, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus." figureDoi="http://doi.org/10.5281/zenodo.14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="2" pageNumber="318">Figs. 1213</figureCitation>
): Papillae anales are somewhat tapered and setosed. Posterior apophyses are typically long and slender. Anterior apophyses are extremely reduced and acute with pointed tips. The ductus bursae is slender and tubular. Corpus bursae has a broad, tube-like proximal part with moderate sclerotization where the ductus bursae conjoins with the corpus bursae. The posterior part is membranous, bearing a large signum with small medial concavity. The transverse ends of the signum terminate in a spine, giving it a groundnut-shaped appearance, with a few central minute spines scattered inside.
</paragraph>
<caption id="67F1665D5C13FFD5E4F596FBA47BBFE5" ID-DOI="http://doi.org/10.5281/zenodo.14954390" ID-Zenodo-Dep="14954390" httpUri="https://zenodo.org/record/14954390/files/figure.png" pageId="3" pageNumber="319" startId="3.[151,264,1943,1967]" targetBox="[207,1380,191,1909]" targetPageId="3" targetType="figure">
<paragraph id="333136D55C13FFD5E4F596FBA47BBFE5" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="319">
<emphasis id="01FAEAC75C13FFD5E4F596FBA430BFBD" bold="true" box="[151,316,1943,1967]" pageId="3" pageNumber="319">FIGURES 14.</emphasis>
Adults of
<taxonomicName id="F48E4D565C13FFD5E5C996FBA769BFBD" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[427,613,1943,1967]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="319" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C13FFD5E5C996FBA769BFBD" box="[427,613,1943,1967]" italics="true" pageId="3" pageNumber="319">Psilocladia indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C13FFD5E60F96FBA7B3BFBD" bold="true" box="[621,703,1943,1967]" pageId="3" pageNumber="319">
<taxonomicNameLabel id="1AC957BC5C13FFD5E60F96FBA7B3BFBD" box="[621,703,1943,1967]" pageId="3" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 1, Male, Holotype (Upper Singchung, Arunachal Pradesh); 2, Female, Paratype (Karadei, Odisha); 3, Male, Paratype (Karadei, Odisha); 4, Female, Paratype (Husigaon, Arunachal Pradesh). a. dorsal sides; b. ventral sides.
</paragraph>
</caption>
<caption id="67F1665D5C14FFD2E4F596FBA4CABFEA" ID-DOI="http://doi.org/10.5281/zenodo.14954392" ID-Zenodo-Dep="14954392" httpUri="https://zenodo.org/record/14954392/files/figure.png" pageId="4" pageNumber="320" startId="4.[151,264,1943,1967]" targetBox="[199,1388,191,1899]" targetPageId="4" targetType="figure">
<paragraph id="333136D55C14FFD2E4F596FBA4CABFEA" blockId="4.[151,1437,1943,2042]" pageId="4" pageNumber="320">
<emphasis id="01FAEAC75C14FFD2E4F596FBA430BFBD" bold="true" box="[151,316,1943,1967]" pageId="4" pageNumber="320">FIGURES 58.</emphasis>
Adults of
<taxonomicName id="F48E4D565C14FFD2E5CF96F4A75ABFBD" box="[429,598,1944,1968]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="320" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="01FAEAC75C14FFD2E5CF96F4A72CBFA2" box="[429,544,1944,1968]" italics="true" pageId="4" pageNumber="320">Psilocladia</emphasis>
spp.
</taxonomicName>
: 5,
<taxonomicName id="F48E4D565C14FFD2E61F96F4A7F5BFA2" authorityName="Warren" authorityYear="1898" box="[637,761,1944,1968]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="320" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C14FFD2E61F96F4A7F5BFA2" box="[637,761,1944,1968]" italics="true" pageId="4" pageNumber="320">P. obliquata</emphasis>
</taxonomicName>
, Male, Holotype (South Africa); 6,
<taxonomicName id="F48E4D565C14FFD2E00C96F4A1E1BFA2" authorityName="Prout" authorityYear="1915" box="[1134,1261,1944,1968]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="320" phylum="Arthropoda" rank="species" species="repudiosa">
<emphasis id="01FAEAC75C14FFD2E00C96F4A1E1BFA2" box="[1134,1261,1944,1968]" italics="true" pageId="4" pageNumber="320">P. repudiosa</emphasis>
</taxonomicName>
, Male, Holotype (Cameroon); 7,
<taxonomicName id="F48E4D565C14FFD2E55A96D0A4AFBFC6" authorityName="Prout" authorityYear="1923" box="[312,419,1980,2004]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="320" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C14FFD2E55A96D0A4AFBFC6" box="[312,419,1980,2004]" italics="true" pageId="4" pageNumber="320">P. diaereta</emphasis>
</taxonomicName>
, Male, Holotype (Kenya); 8,
<taxonomicName id="F48E4D565C14FFD2E6AD96D0A65ABFC6" authorityName="Prout" authorityYear="1915" box="[719,854,1980,2004]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="320" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C14FFD2E6AD96D0A65ABFC6" box="[719,854,1980,2004]" italics="true" pageId="4" pageNumber="320">P. loxostigma</emphasis>
</taxonomicName>
, Male, Holotype (Congo). a. dorsal sides; b. ventral sides (for figures 58 © NHMUK).
</paragraph>
</caption>
<caption id="67F1665D5C15FFD3E4F5961DA68FBFE4" ID-DOI="http://doi.org/10.5281/zenodo.14954394" ID-Zenodo-Dep="14954394" httpUri="https://zenodo.org/record/14954394/files/figure.png" pageId="5" pageNumber="321" startId="5.[151,264,1905,1929]" targetBox="[215,1374,156,1880]" targetPageId="5" targetType="figure">
<paragraph id="333136D55C15FFD3E4F5961DA68FBFE4" blockId="5.[151,1436,1905,2038]" pageId="5" pageNumber="321">
<emphasis id="01FAEAC75C15FFD3E4F5961DA444BF9B" bold="true" box="[151,328,1905,1929]" pageId="5" pageNumber="321">FIGURES 914.</emphasis>
Genitalia structure of
<taxonomicName id="F48E4D565C15FFD3E652961EA7D4BF9B" box="[560,728,1906,1930]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="01FAEAC75C15FFD3E652961EA7AFBF98" box="[560,675,1906,1930]" italics="true" pageId="5" pageNumber="321">Psilocladia</emphasis>
spp.
</taxonomicName>
: 9,
<taxonomicName id="F48E4D565C15FFD3E69F961EA65ABF98" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[765,854,1906,1930]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C15FFD3E69F961EA65ABF98" box="[765,854,1906,1930]" italics="true" pageId="5" pageNumber="321">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C15FFD3E73F961EA6BABF98" bold="true" box="[861,950,1906,1930]" pageId="5" pageNumber="321">
<taxonomicNameLabel id="1AC957BC5C15FFD3E73F961EA6BDBF98" box="[861,945,1906,1930]" pageId="5" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male, HT (Upper Singchung); 10,
<taxonomicName id="F48E4D565C15FFD3E17F961EA07ABF98" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[1309,1398,1906,1930]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C15FFD3E17F961EA07ABF98" box="[1309,1398,1906,1930]" italics="true" pageId="5" pageNumber="321">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C15FFD3E11C961EA5C6BFBC" bold="true" pageId="5" pageNumber="321">
<taxonomicNameLabel id="1AC957BC5C15FFD3E11C961EA5C9BFBC" pageId="5" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male, PT (Karadei); 11,
<taxonomicName id="F48E4D565C15FFD3E5AC96FAA746BFBC" authorityName="Warren" authorityYear="1898" box="[462,586,1942,1966]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C15FFD3E5AC96FAA746BFBC" box="[462,586,1942,1966]" italics="true" pageId="5" pageNumber="321">P. obliquata</emphasis>
</taxonomicName>
, male (Mpumalanga, South Africa) (Image source, Dr. P. Sihvonen); 12,
<taxonomicName id="F48E4D565C15FFD3E12096FAA090BFBC" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[1346,1436,1942,1966]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C15FFD3E12096FAA090BFBC" box="[1346,1436,1942,1966]" italics="true" pageId="5" pageNumber="321">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C15FFD3E4F596D6A5E3BFC0" bold="true" box="[151,239,1978,2002]" pageId="5" pageNumber="321">
<taxonomicNameLabel id="1AC957BC5C15FFD3E4F596D6A5E6BFC0" box="[151,234,1978,2002]" pageId="5" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female (Karadei); 13,
<taxonomicName id="F48E4D565C15FFD3E5B496D6A722BFC0" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[470,558,1978,2002]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C15FFD3E5B496D6A722BFC0" box="[470,558,1978,2002]" italics="true" pageId="5" pageNumber="321">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C15FFD3E65796D6A780BFC0" bold="true" box="[565,652,1978,2002]" pageId="5" pageNumber="321">
<taxonomicNameLabel id="1AC957BC5C15FFD3E65796D6A78BBFC0" box="[565,647,1978,2002]" pageId="5" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female (Husigaon); 14,
<taxonomicName id="F48E4D565C15FFD3E7E696D6A6F2BFC0" authorityName="Warren" authorityYear="1898" box="[900,1022,1978,2002]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="321" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C15FFD3E7E696D6A6F2BFC0" box="[900,1022,1978,2002]" italics="true" pageId="5" pageNumber="321">P. obliquata</emphasis>
</taxonomicName>
, female (KwaZulu-Natal, Bulwer, South Africa) (Image source, Dr. P. Sihvonen). a, genitalia capsule; b, aedeagus.
</paragraph>
</caption>
<caption id="67F1665D5C16FFD0E4F596F8A6CCBFE6" ID-DOI="http://doi.org/10.5281/zenodo.14954396" ID-Zenodo-Dep="14954396" httpUri="https://zenodo.org/record/14954396/files/figure.png" pageId="6" pageNumber="322" startId="6.[151,264,1940,1964]" targetBox="[162,1424,194,1905]" targetPageId="6" targetType="figure">
<paragraph id="333136D55C16FFD0E4F596F8A6CCBFE6" blockId="6.[151,1436,1940,2037]" pageId="6" pageNumber="322">
<emphasis id="01FAEAC75C16FFD0E4F596F8A45BBFBE" bold="true" box="[151,343,1940,1964]" pageId="6" pageNumber="322">FIGURES 1518.</emphasis>
Distribution map and collection habitats: 15, Map showing global distribution of species of
<taxonomicName id="F48E4D565C16FFD0E14B96F8A5EFBFC2" pageId="6" pageNumber="322">
<emphasis id="01FAEAC75C16FFD0E14B96F8A090BFBE" box="[1321,1436,1940,1964]" italics="true" pageId="6" pageNumber="322">Psilocladia</emphasis>
species
</taxonomicName>
; 1618, Collection habitat of
<taxonomicName id="F48E4D565C16FFD0E67196D5A767BFC2" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[531,619,1976,2000]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="322" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C16FFD0E67196D5A767BFC2" box="[531,619,1976,2000]" italics="true" pageId="6" pageNumber="322">P. indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C16FFD0E61096D5A7CFBFC3" bold="true" box="[626,707,1977,2001]" pageId="6" pageNumber="322">
<taxonomicNameLabel id="1AC957BC5C16FFD0E61096D5A7CFBFC3" box="[626,707,1977,2001]" pageId="6" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 16, Karadei, Angul dist., Odisha; 17, Upper Singchung, West Kameng dist., Arunachal Pradesh; 18, Husigaon, West Kameng dist., Arunachal Pradesh.
</paragraph>
</caption>
<paragraph id="333136D55C17FFD1E4A591FAA18FB99B" blockId="7.[151,1436,150,465]" pageId="7" pageNumber="323">
<emphasis id="01FAEAC75C17FFD1E4A591FAA441B8A2" bold="true" box="[199,333,150,176]" pageId="7" pageNumber="323">Bionomics:</emphasis>
<taxonomicName id="F48E4D565C17FFD1E53191FBA713B8A3" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[339,543,151,177]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C17FFD1E53191FBA713B8A3" box="[339,543,151,177]" italics="true" pageId="7" pageNumber="323">Psilocladia indica</emphasis>
</taxonomicName>
<emphasis id="01FAEAC75C17FFD1E64491FBA773B8A3" bold="true" box="[550,639,151,177]" pageId="7" pageNumber="323">
<taxonomicNameLabel id="1AC957BC5C17FFD1E64491FBA773B8A3" box="[550,639,151,177]" pageId="7" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is found in a variety of ecosystems (
<figureCitation id="ABB52A505C17FFD1E07791FBA196B8A3" box="[1045,1178,150,177]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1940,1964]" captionTargetBox="[162,1424,194,1905]" captionTargetId="figure-19@6.[151,1436,181,1916]" captionTargetPageId="6" captionText="FIGURES 1518. Distribution map and collection habitats: 15, Map showing global distribution of species of Psilocladia species; 1618, Collection habitat of P. indica sp. nov.: 16, Karadei, Angul dist., Odisha; 17, Upper Singchung, West Kameng dist., Arunachal Pradesh; 18, Husigaon, West Kameng dist., Arunachal Pradesh." figureDoi="http://doi.org/10.5281/zenodo.14954396" httpUri="https://zenodo.org/record/14954396/files/figure.png" pageId="7" pageNumber="323">Figs. 1518</figureCitation>
) ranging from lowland tropical forests to wet temperate forests, at altitude range up to
<quantity id="F4769B305C17FFD1E70F91D7A6CBB8C7" box="[877,967,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="323" unit="m" value="1800.0">1800 m</quantity>
. These habitats are dominated by several prominent plant families, including
<taxonomicName id="F48E4D565C17FFD1E67E91B3A785B8EB" box="[540,649,223,249]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E6F391B3A60DB8EB" box="[657,769,223,249]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E76B91B3A6A0B8EB" box="[777,940,223,249]" class="Magnoliopsida" family="Passifloraceae" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Passifloraceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E7D191B3A14BB8EB" box="[947,1095,223,249]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Brassicaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E02D91B3A1C7B8EB" box="[1103,1227,223,249]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E0B191B3A094B8EB" box="[1235,1432,223,249]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E4F5906FA40EB90F" box="[151,258,259,285]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E56F906FA487B90F" box="[269,395,259,285]" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
,
<taxonomicName id="F48E4D565C17FFD1E5F7906FA71FB90F" box="[405,531,259,285]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
, and
<taxonomicName id="F48E4D565C17FFD1E62F906FA7D2B90F" box="[589,734,259,285]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="323" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
. The Eastern Ghats (
<collectingCountry id="4B9976455C17FFD1E7A7906FA10EB90F" box="[965,1026,259,285]" name="India" pageId="7" pageNumber="323">India</collectingCountry>
) too are markedly different from the habitats of its congeners in the Afrotropic. African species of
<taxonomicName id="F48E4D565C17FFD1E75E904BA6B0B953" authorityName="Warren" authorityYear="1898" box="[828,956,295,321]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C17FFD1E75E904BA6B0B953" box="[828,956,295,321]" italics="true" pageId="7" pageNumber="323">Psilocladia</emphasis>
</taxonomicName>
inhabit diverse ecosystems, such as coastal forests to grasslands (
<taxonomicName id="F48E4D565C17FFD1E5EB9027A703B977" authorityName="Warren" authorityYear="1898" box="[393,527,331,357]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C17FFD1E5EB9027A703B977" box="[393,527,331,357]" italics="true" pageId="7" pageNumber="323">P. obliquata</emphasis>
</taxonomicName>
) (
<bibRefCitation id="571F4B245C17FFD1E6449027A7D8B977" author="Mapcarta" box="[550,724,331,357]" pageId="7" pageNumber="323" refId="ref4271" refString="Mapcarta (2024) Open map of the world. Available from: https://www.mapcarta.com (accessed 17 May 2024)" type="url" year="2024">Mapcarta 2024</bibRefCitation>
), tropical rainforests (
<taxonomicName id="F48E4D565C17FFD1E7AB9027A153B977" authorityName="Prout" authorityYear="1915" box="[969,1119,331,357]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C17FFD1E7AB9027A153B977" box="[969,1119,331,357]" italics="true" pageId="7" pageNumber="323">P. loxostigma</emphasis>
</taxonomicName>
and
<taxonomicName id="F48E4D565C17FFD1E0F69027A012B977" authorityName="Prout" authorityYear="1915" box="[1172,1310,331,357]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="repudiosa">
<emphasis id="01FAEAC75C17FFD1E0F69027A012B977" box="[1172,1310,331,357]" italics="true" pageId="7" pageNumber="323">P. repudiosa</emphasis>
</taxonomicName>
) (
<bibRefCitation id="571F4B245C17FFD1E1579027A5F9B99B" author="Linder, H. P. &amp; Helen, M. de &amp; Klerk &amp; Born, J. &amp; Burgess, N. D. &amp; Fjeldsa, J. &amp; Rahbek, C." pageId="7" pageNumber="323" pagination="1189 - 1205" refId="ref4191" refString="Linder, H. P., Helen, M. de Klerk, Born, J., Burgess, N. D., Fjeldsa, J. &amp; Rahbek, C. (2012) The partitioning of Africa: statistically defined biogeographical regions in sub-Saharan Africa. Journal of Biogeography, 39 (7), 1189-1205. https://doi.org/10.1111/j.1365-2699.2012.02728.x" type="journal article" year="2012">
Linder
<emphasis id="01FAEAC75C17FFD1E1EA9027A5B8B99B" italics="true" pageId="7" pageNumber="323">et al.</emphasis>
2012
</bibRefCitation>
) and urban areas with forest patches (
<taxonomicName id="F48E4D565C17FFD1E6FC9003A619B99B" authorityName="Prout" authorityYear="1923" box="[670,789,367,393]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C17FFD1E6FC9003A619B99B" box="[670,789,367,393]" italics="true" pageId="7" pageNumber="323">P. diaereta</emphasis>
</taxonomicName>
) (
<bibRefCitation id="571F4B245C17FFD1E74F9003A172B99B" author="Kenya Forest Services" box="[813,1150,367,393]" pageId="7" pageNumber="323" refId="ref4104" refString="Kenya Forest Services (2024) Official website of Kenya forest service, Kenya. Available from: https://www.kenyaforestservices.org (accessed 17 May 2024)" type="url" year="2024">
<collectingCountry id="4B9976455C17FFD1E74F9003A674B99B" box="[813,888,367,393]" name="Kenya" pageId="7" pageNumber="323">Kenya</collectingCountry>
forest services (2024))
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C17FFD1E4A590FEA090B9BF" box="[199,1436,402,429]" pageId="7" pageNumber="323" type="distribution">
<paragraph id="333136D55C17FFD1E4A590FEA090B9BF" blockId="7.[151,1436,150,465]" box="[199,1436,402,429]" pageId="7" pageNumber="323">
<emphasis id="01FAEAC75C17FFD1E4A590FEA454B9BE" bold="true" box="[199,344,402,428]" pageId="7" pageNumber="323">Distribution:</emphasis>
The new species is distributed in the Eastern Ghats (
<collectingRegion id="F14AF8375C17FFD1E71D90FFA6C0B9BF" box="[895,972,403,429]" country="India" name="Odisha" pageId="7" pageNumber="323">Odisha</collectingRegion>
) and Himalayas (
<collectingRegion id="F14AF8375C17FFD1E0E090FFA042B9BF" box="[1154,1358,403,429]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="323">Arunachal Pradesh</collectingRegion>
),
<collectingCountry id="4B9976455C17FFD1E13D90FFA094B9BF" box="[1375,1432,403,429]" name="India" pageId="7" pageNumber="323">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C17FFD1E4A590DBA6F2B9C3" box="[199,1022,439,465]" pageId="7" pageNumber="323" type="etymology">
<paragraph id="333136D55C17FFD1E4A590DBA6F2B9C3" blockId="7.[151,1436,150,465]" box="[199,1022,439,465]" pageId="7" pageNumber="323">
<emphasis id="01FAEAC75C17FFD1E4A590DBA45EB9C3" bold="true" box="[199,338,439,465]" pageId="7" pageNumber="323">Etymology:</emphasis>
The name of the species represents country of origin India.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B03AF58B80433093415D0BABB6F45B01">A new species of genus Psilocladia Warren from India, bridging the boundaries of Africa and Asia (Lepidoptera: Geometridae: Ennominae)</mods:title>
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<mods:affiliation id="E3CE8BD9B37619097EADD38355750BFA">Zoological Survey of India, New Alipore, Kolkata- 700053, West Bengal, India &amp; Department of Zoology &amp; Environmental Sciences, Punjabi University, Patiala- 147002, Punjab, India</mods:affiliation>
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<mods:affiliation id="8689FA25DE6E9FD56FA9CDF132FB875E">Zoological Survey of India, New Alipore, Kolkata- 700053, West Bengal, India &amp; Department of Zoology, University of Calcutta, Kolkata- 700019, West Bengal, India</mods:affiliation>
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<mods:namePart id="FA6155CAB55FFCC8F37A4030E818286D">Chatterjee, Pallab</mods:namePart>
<mods:affiliation id="D8A8F34D907D95C6B1CCEC1D985766E7">Zoological Survey of India, New Alipore, Kolkata- 700053, West Bengal, India &amp; Department of Zoology, University of Calcutta, Kolkata- 700019, West Bengal, India</mods:affiliation>
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<heading id="687981B95C11FFD7E4F5932AA72ABA72" bold="true" box="[151,550,582,609]" fontSize="11" level="1" pageId="1" pageNumber="317" reason="1">
<emphasis id="01FAEAC75C11FFD7E4F5932AA72ABA72" bold="true" box="[151,550,582,609]" pageId="1" pageNumber="317">
Genus
<taxonomicName id="F48E4D565C11FFD7E48C932AA72ABA72" ID-CoL="92L9S" authority="Warren, 1898" authorityName="Warren" authorityYear="1898" box="[238,550,582,609]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E48C932AA478BA73" bold="true" box="[238,372,582,609]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
<bibRefCitation id="571F4B245C11FFD7E519932BA72ABA72" author="Warren, W." box="[379,550,582,609]" pageId="1" pageNumber="317" pagination="5 - 41" refId="ref4869" refString="Warren, W. (1898) New species and genera of the families Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old- World Regions. Novitates Zoologicae, 5, 5-41. https://doi.org/10.5962/bhl.part.1646" type="journal article" year="1898">Warren, 1898</bibRefCitation>
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<taxonomicName id="F48E4D565C11FFD7E4F593E7A490BAB1" ID-CoL="92L9S" authority="Warren, 1898" authorityName="Warren" authorityYear="1898" box="[151,412,651,675]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E4F593E7A406BAB1" box="[151,266,651,675]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
<bibRefCitation id="571F4B245C11FFD7E57293E0A490BAB1" author="Warren, W." box="[272,412,651,675]" pageId="1" pageNumber="317" pagination="5 - 41" refId="ref4869" refString="Warren, W. (1898) New species and genera of the families Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old- World Regions. Novitates Zoologicae, 5, 5-41. https://doi.org/10.5962/bhl.part.1646" type="journal article" year="1898">Warren, 1898</bibRefCitation>
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,
<emphasis id="01FAEAC75C11FFD7E5C493E0A72CBAB1" box="[422,544,651,675]" italics="true" pageId="1" pageNumber="317">Novit. Zool.</emphasis>
5: 40.
</paragraph>
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<paragraph id="333136D55C11FFD7E4F593BDA084BAF9" blockId="1.[151,1437,721,1287]" box="[151,1416,721,748]" pageId="1" pageNumber="317">
<emphasis id="01FAEAC75C11FFD7E4F593BDA5CFBAF9" bold="true" box="[151,195,721,747]" pageId="1" pageNumber="317">TS:</emphasis>
<taxonomicName id="F48E4D565C11FFD7E4A893BDA751BAF9" ID-CoL="4PGX3" authority="Warren, 1898" authorityName="Warren" authorityYear="1898" box="[202,605,721,748]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C11FFD7E4A893BDA4B7BAF9" box="[202,443,721,747]" italics="true" pageId="1" pageNumber="317">Psilocladia obliquata</emphasis>
<bibRefCitation id="571F4B245C11FFD7E5A093BEA751BAF9" author="Warren, W." box="[450,605,721,748]" pageId="1" pageNumber="317" pagination="5 - 41" refId="ref4869" refString="Warren, W. (1898) New species and genera of the families Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old- World Regions. Novitates Zoologicae, 5, 5-41. https://doi.org/10.5962/bhl.part.1646" type="journal article" year="1898">Warren, 1898</bibRefCitation>
</taxonomicName>
,
<emphasis id="01FAEAC75C11FFD7E60A93BEA7E1BAF9" box="[616,749,721,747]" italics="true" pageId="1" pageNumber="317">Novit. Zool.</emphasis>
5: 40, by monotypy (
<emphasis id="01FAEAC75C11FFD7E7BD93BDA103BAF9" bold="true" box="[991,1039,721,747]" pageId="1" pageNumber="317">TL:</emphasis>
Northdene, Natal [
<collectingCountry id="4B9976455C11FFD7E08193BDA075BAF9" box="[1251,1401,721,747]" name="South Africa" pageId="1" pageNumber="317">South Africa</collectingCountry>
])
</paragraph>
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<subSubSection id="7B94655E5C11FFD7E4A59399A720BCAD" pageId="1" pageNumber="317" type="diagnosis">
<paragraph id="333136D55C11FFD7E4A59399A5FDBC19" blockId="1.[151,1437,721,1287]" pageId="1" pageNumber="317">
<emphasis id="01FAEAC75C11FFD7E4A59399A448BB1D" bold="true" box="[199,324,757,783]" pageId="1" pageNumber="317">Diagnosis:</emphasis>
The genus
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<emphasis id="01FAEAC75C11FFD7E5AE9399A740BB1D" box="[460,588,757,783]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
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is closely related to an African genus
<taxonomicName id="F48E4D565C11FFD7E065939AA014BB02" authority="Warren, 1895" authorityName="Warren" authorityYear="1895" box="[1031,1304,757,784]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E065939AA17EBB1D" box="[1031,1138,758,783]" italics="true" pageId="1" pageNumber="317">Xenimpia</emphasis>
<bibRefCitation id="571F4B245C11FFD7E019939AA014BB02" author="Warren, W." box="[1147,1304,757,784]" pageId="1" pageNumber="317" pagination="82 - 159" refId="ref4842" refString="Warren, W. (1895) New species and genera of Geometridae in the Tring Museum. Novitates Zoologicae, 2, 82-159." type="journal article" year="1895">Warren, 1895</bibRefCitation>
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, but can be diagnosed by its simple and lamellate antennae in both sexes. In male genitalia,
<taxonomicName id="F48E4D565C11FFD7E0249275A1CABB21" authorityName="Warren" authorityYear="1898" box="[1094,1222,793,819]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E0249275A1CABB21" box="[1094,1222,793,819]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
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features a simple, elongated valva without any sclerotized structures, a pair of elongated furca (symmetrical or asymmetrical) with bulbous base, a slender and elongated aedeagus with a weakly sclerotized patch on vesica. The female genitalia exhibit a distinct large signum. In contrast, the females of
<taxonomicName id="F48E4D565C11FFD7E77792EAA68CBB8D" authorityName="Warren" authorityYear="1895" box="[789,896,902,927]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E77792EAA68CBB8D" box="[789,896,902,927]" italics="true" pageId="1" pageNumber="317">Xenimpia</emphasis>
</taxonomicName>
have simple antennae and males have pectinated antennae. The male genitalia of
<taxonomicName id="F48E4D565C11FFD7E66B92C6A778BBD1" authorityName="Warren" authorityYear="1895" box="[521,628,938,963]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E66B92C6A778BBD1" box="[521,628,938,963]" italics="true" pageId="1" pageNumber="317">Xenimpia</emphasis>
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display a short valva with sclerotized structures; furca absent; a small and broad aedeagus with one or two long cornuti in vesica. Female genitalia display a sclerotized patch but lacks signum.
</paragraph>
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Superficially,
<taxonomicName id="F48E4D565C11FFD7E5049579A4EABC3D" authorityName="Warren" authorityYear="1898" box="[358,486,1045,1071]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E5049579A4EABC3D" box="[358,486,1045,1071]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
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also resembles
<taxonomicName id="F48E4D565C11FFD7E6FF9579A6CCBC3D" authority="Warren, 1894" authorityName="Warren" authorityYear="1894" box="[669,960,1045,1072]" class="Insecta" family="Geometridae" genus="Zanclopera" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E6FF9579A612BC3D" box="[669,798,1045,1071]" italics="true" pageId="1" pageNumber="317">Zanclopera</emphasis>
<bibRefCitation id="571F4B245C11FFD7E745957AA6CCBC3D" author="Warren, W." box="[807,960,1045,1072]" pageId="1" pageNumber="317" pagination="366 - 466" refId="ref4805" refString="Warren, W. (1894) New genera and species of Geometridae. Novitates Zoologicae, 1, 366-466. https://doi.org/10.5962/bhl.part.24566" type="journal article" year="1894">Warren, 1894</bibRefCitation>
</taxonomicName>
and
<emphasis id="01FAEAC75C11FFD7E7989579A18CBC3D" box="[1018,1152,1045,1071]" italics="true" pageId="1" pageNumber="317">Triginoptila</emphasis>
<bibRefCitation id="571F4B245C11FFD7E0EB957AA029BC3D" author="Warren, W." box="[1161,1317,1045,1072]" pageId="1" pageNumber="317" pagination="366 - 466" refId="ref4805" refString="Warren, W. (1894) New genera and species of Geometridae. Novitates Zoologicae, 1, 366-466. https://doi.org/10.5962/bhl.part.24566" type="journal article" year="1894">Warren, 1894</bibRefCitation>
, currently, junior synonyms of an Indian genus
<taxonomicName id="F48E4D565C11FFD7E64A9555A623BC41" authority="Moore, 1868" authorityName="Moore" authorityYear="1868" box="[552,815,1081,1108]" class="Insecta" family="Geometridae" genus="Krananda" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E64A9555A796BC41" box="[552,666,1081,1107]" italics="true" pageId="1" pageNumber="317">Krananda</emphasis>
<bibRefCitation id="571F4B245C11FFD7E6C29556A623BC41" author="Moore, F." box="[672,815,1081,1108]" pageId="1" pageNumber="317" pagination="612 - 686" refId="ref4298" refString="Moore, F. (1868) On the Lepidopterous Insects of Bengal. Proceedings of the Zoological Society of London, 1867 (3), 612-686, pls. 32 - 33." type="journal article" year="1868">Moore, 1868</bibRefCitation>
</taxonomicName>
but
<taxonomicName id="F48E4D565C11FFD7E73C9555A6D2BC41" authorityName="Warren" authorityYear="1898" box="[862,990,1081,1107]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E73C9555A6D2BC41" box="[862,990,1081,1107]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
</taxonomicName>
can be easily diagnosed from
<taxonomicName id="F48E4D565C11FFD7E1499555A091BC41" authorityName="Moore" authorityYear="1868" box="[1323,1437,1081,1107]" class="Insecta" family="Geometridae" genus="Krananda" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E1499555A091BC41" box="[1323,1437,1081,1107]" italics="true" pageId="1" pageNumber="317">Krananda</emphasis>
</taxonomicName>
by its antennal morphology (
<bibRefCitation id="571F4B245C11FFD7E5809531A776BC65" author="Warren, W." box="[482,634,1117,1143]" pageId="1" pageNumber="317" pagination="5 - 41" refId="ref4869" refString="Warren, W. (1898) New species and genera of the families Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old- World Regions. Novitates Zoologicae, 5, 5-41. https://doi.org/10.5962/bhl.part.1646" type="journal article" year="1898">Warren 1898</bibRefCitation>
;
<bibRefCitation id="571F4B245C11FFD7E6EA9531A641BC65" author="Jiang, N. &amp; Li, X. &amp; Hausmann, A. &amp; Cheng, R. &amp; Xue, D. &amp; Han, H." box="[648,845,1117,1144]" pageId="1" pageNumber="317" pagination="427 - 441" refId="ref4028" refString="Jiang, N., Li, X., Hausmann, A., Cheng, R., Xue, D. &amp; Han, H. (2017) A molecular phylogeny of the Palaearctic and Oriental members of the tribe Boarmiini (Lepidoptera: Geometridae: Ennominae). Invertebrate Systematics, 31 (4), 427-441. https://doi.org/10.1071/IS17005" type="journal article" year="2017">
Jiang
<emphasis id="01FAEAC75C11FFD7E6AF9532A60CBC65" box="[717,768,1117,1143]" italics="true" pageId="1" pageNumber="317">et al</emphasis>
. 2017
</bibRefCitation>
). Besides this,
<taxonomicName id="F48E4D565C11FFD7E7999531A177BC65" authorityName="Warren" authorityYear="1898" box="[1019,1147,1117,1143]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E7999531A177BC65" box="[1019,1147,1117,1143]" italics="true" pageId="1" pageNumber="317">Psilocladia</emphasis>
</taxonomicName>
and
<taxonomicName id="F48E4D565C11FFD7E0D49531A024BC65" authorityName="Moore" authorityYear="1868" box="[1206,1320,1117,1143]" class="Insecta" family="Geometridae" genus="Krananda" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C11FFD7E0D49531A024BC65" box="[1206,1320,1117,1143]" italics="true" pageId="1" pageNumber="317">Krananda</emphasis>
</taxonomicName>
belong to different tribes, i.e.,
<taxonomicName id="F48E4D565C11FFD7E51595EDA709BC89" box="[375,517,1153,1179]" pageId="1" pageNumber="316" rank="tribe" tribe="Gonodontini">Gonodontini</taxonomicName>
and
<taxonomicName id="F48E4D565C11FFD7E65B95EDA7A1BC89" box="[569,685,1153,1179]" class="Insecta" family="Geometridae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="324" phylum="Arthropoda" rank="tribe" tribe="Boarmiini">Boarmiini</taxonomicName>
, respectively, which exhibit distinct characteristics (
<bibRefCitation id="571F4B245C11FFD7E08C95EDA090BC89" author="Holloway, J. D." box="[1262,1436,1153,1179]" pageId="1" pageNumber="317" refId="ref3986" refString="Holloway, J. D. (1993 [1994]) The Moths of Borneo. Part 11. Family Geometridae, Subfamily Ennominae. Malayan Nature Journal, Southdene Sdn Bhd., 309 pp., 19 pls." type="book" year="1993">Holloway 1993</bibRefCitation>
[1994];
<bibRefCitation id="571F4B245C11FFD7E49395C9A72CBCAD" author="Murillo-Ramos, L. &amp; Brehm, G. &amp; Sihvonen, P. &amp; Hausmann, A. &amp; Holm, S. &amp; Reza Ghanavi, H. &amp; Ounap, E. &amp; Truuverk, A. &amp; Staude, H. &amp; Friedrich, E. &amp; Tammaru, T. &amp; Wahlberg, N." box="[241,544,1189,1216]" pageId="1" pageNumber="317" refId="ref4335" refString="Murillo-Ramos, L., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Reza Ghanavi, H., Ounap, E., Truuverk, A., Staude, H., Friedrich, E., Tammaru, T. &amp; Wahlberg, N. (2019) A comprehensive molecular phylogeny of Geometridae (lepidoptera) with a focus on enigmatic small subfamilies. PeerJ, 7, e 7386. https://doi.org/10.7717/peerj.7386" type="journal volume" year="2019">
Murillo-Ramos
<emphasis id="01FAEAC75C11FFD7E5C695CAA4D9BCAD" box="[420,469,1189,1215]" italics="true" pageId="1" pageNumber="317">et al</emphasis>
. 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C11FFD7E4A595A5A720BD15" pageId="1" pageNumber="317" type="distribution">
<paragraph id="333136D55C11FFD7E4A595A5A720BD15" blockId="1.[151,1437,721,1287]" pageId="1" pageNumber="317">
<emphasis id="01FAEAC75C11FFD7E4A595A5A46FBCF1" bold="true" box="[199,355,1225,1251]" pageId="1" pageNumber="317">Distribution:</emphasis>
<collectingCountry id="4B9976455C11FFD7E50F95A5A70CBCF1" box="[365,512,1225,1251]" name="South Africa" pageId="1" pageNumber="317">South Africa</collectingCountry>
,
<collectingCountry id="4B9976455C11FFD7E66D95A5A76BBCF1" box="[527,615,1225,1251]" name="Malawi" pageId="1" pageNumber="317">Malawi</collectingCountry>
,
<collectingCountry id="4B9976455C11FFD7E61795A5A7CEBCF1" box="[629,706,1225,1251]" name="Democratic Republic of the Congo" pageId="1" pageNumber="317">Congo</collectingCountry>
,
<collectingCountry id="4B9976455C11FFD7E6B295A6A610BCF6" box="[720,796,1226,1252]" name="Kenya" pageId="1" pageNumber="317">Kenya</collectingCountry>
,
<collectingCountry id="4B9976455C11FFD7E74895A5A693BCF1" box="[810,927,1225,1251]" name="Cameroon" pageId="1" pageNumber="317">Cameroon</collectingCountry>
(
<bibRefCitation id="571F4B245C11FFD7E7D295A5A145BCF1" author="Warren, W." box="[944,1097,1225,1251]" pageId="1" pageNumber="317" pagination="5 - 41" refId="ref4869" refString="Warren, W. (1898) New species and genera of the families Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old- World Regions. Novitates Zoologicae, 5, 5-41. https://doi.org/10.5962/bhl.part.1646" type="journal article" year="1898">Warren 1898</bibRefCitation>
;
<bibRefCitation id="571F4B245C11FFD7E03595A6A1D5BCF6" author="Prout L. B." box="[1111,1241,1225,1252]" pageId="1" pageNumber="317" pagination="311 - 385" refId="ref4524" refString="Prout L. B. (1915) New genera and species of African Geometridae. Novitates Zoologicae, A Journal of Zoology in connection with the Tring Museum, 22, 311-385. https://doi.org/10.5962/bhl.part.1923" type="journal article" year="1915">Prout 1915</bibRefCitation>
,
<bibRefCitation id="571F4B245C11FFD7E08595A5A02EBCF1" author="Prout, L. B." box="[1255,1314,1225,1251]" pageId="1" pageNumber="317" pagination="191 - 215" refId="ref4574" refString="Prout, L. B. (1923) New Geometridae in the Tring Museum. Novitates Zoologicae, 30, 191-215. https://doi.org/10.5962/bhl.part.28985" type="journal article" year="1923">1923</bibRefCitation>
;
<bibRefCitation id="571F4B245C11FFD7E15295A5A43FBD15" author="Staude, H. &amp; Murphy, R." pageId="1" pageNumber="317" pagination="32 - 48" refId="ref4759" refString="Staude, H. &amp; Murphy, R. (2011) The geometrid fauna (Lepidoptera: Geometridae) of Mount Mulanje, Malawi and the importance of Mount Mulanje as an area of conservation priority. Metamorphosis, 2011, 32-48." type="journal article" year="2011">Staude &amp; Murphy 2011</bibRefCitation>
),
<collectingCountry id="4B9976455C11FFD7E5259581A48DBD15" box="[327,385,1261,1287]" name="India" pageId="1" pageNumber="317">India</collectingCountry>
(
<emphasis id="01FAEAC75C11FFD7E5F39581A717BD15" bold="true" box="[401,539,1261,1287]" pageId="1" pageNumber="317">New record</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C17FFD1E4F5934EA6A9BA2F" box="[151,933,546,573]" pageId="7" pageNumber="323" type="multiple">
<paragraph id="333136D55C17FFD1E4F5934EA6A9BA2F" blockId="7.[151,933,546,573]" box="[151,933,546,573]" pageId="7" pageNumber="323">
<heading id="687981B95C17FFD1E4F5934EA6A9BA2F" bold="true" box="[151,933,546,573]" fontSize="11" level="1" pageId="7" pageNumber="323" reason="1">
<emphasis id="01FAEAC75C17FFD1E4F5934EA6A9BA2F" bold="true" box="[151,933,546,573]" pageId="7" pageNumber="323">
Identification key to the currently known species of
<taxonomicName id="F48E4D565C17FFD1E77D934EA6A9BA2F" authorityName="Warren" authorityYear="1898" box="[799,933,546,573]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="01FAEAC75C17FFD1E77D934EA6A9BA2F" bold="true" box="[799,933,546,573]" italics="true" pageId="7" pageNumber="323">Psilocladia</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7B94655E5C17FFD1E4F59304A090BBAD" pageId="7" pageNumber="323" type="key">
<key id="471F61E45C17FFD1E4F59304A090BBAD" pageId="7" pageNumber="323">
<keyStep id="887A2E705C17FFD1E4F59304A090BA88" pageId="7" pageNumber="323">
<paragraph id="333136D55C17FFD1E4F59304A090BA6E" blockId="7.[151,1437,615,960]" box="[151,1436,615,638]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F59304A090BA6E" box="[151,1436,615,638]" pageId="7" pageNumber="323">
1. In forewing, outer marginal area characterized by fulvous area with a distinct small light subapical spot.................
<emphasis id="01FAEAC75C17FFD1E1F2930BA090BA6E" bold="true" box="[1424,1436,615,636]" pageId="7" pageNumber="323">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="333136D55C17FFD1E4F593E9A090BA88" blockId="7.[151,1437,615,960]" box="[151,1436,645,667]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F593E9A090BA88" box="[151,1436,645,667]" pageId="7" pageNumber="323">
• In forewing, outer marginal area without any fulvous area and light subapical spot..................................
<emphasis id="01FAEAC75C17FFD1E1F293E9A090BA88" bold="true" box="[1424,1436,645,666]" pageId="7" pageNumber="323">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="887A2E705C17FFD1E4F593CEA090BB02" pageId="7" pageNumber="323">
<paragraph id="333136D55C17FFD1E4F593CEA090BAC7" blockId="7.[151,1437,615,960]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F593CEA090BAC7" pageId="7" pageNumber="323">
2. In forewing, an elongated pale subapical spot present; Hindwing with deeper excision on termen near apex between Rs and M3, with no protrusion in the middle.................................................................
<taxonomicName id="F48E4D565C17FFD1E15793ACA090BAC7" authorityName="Prout" authorityYear="1923" box="[1333,1436,703,725]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="diaereta">
<emphasis id="01FAEAC75C17FFD1E15793ACA090BAC7" bold="true" box="[1333,1436,703,725]" italics="true" pageId="7" pageNumber="323">P. diaereta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="333136D55C17FFD1E4F593B1A090BB02" blockId="7.[151,1437,615,960]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F593B1A090BB02" pageId="7" pageNumber="323">
• In forewing, a small, pale subapical spot present; Hindwing with less excision on termen near apex with a slight protrusion in the middle on vein M1.......................................................................
<taxonomicName id="F48E4D565C17FFD1E17B9397A090BB02" authorityName="Prout" authorityYear="1915" box="[1305,1436,762,784]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="loxostigma">
<emphasis id="01FAEAC75C17FFD1E17B9397A090BB02" bold="true" box="[1305,1436,762,784]" italics="true" pageId="7" pageNumber="323">P. loxostigma</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="887A2E705C17FFD1E4F59274A090BB58" pageId="7" pageNumber="323">
<paragraph id="333136D55C17FFD1E4F59274A090BB3E" blockId="7.[151,1437,615,960]" box="[151,1436,791,814]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F59274A090BB3E" box="[151,1436,791,814]" pageId="7" pageNumber="323">
3. In hindwing, postmedial line dark brown, distinct and clearly defined............................................
<emphasis id="01FAEAC75C17FFD1E1F2927BA090BB3E" bold="true" box="[1424,1436,791,812]" pageId="7" pageNumber="323">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="333136D55C17FFD1E4F59259A090BB58" blockId="7.[151,1437,615,960]" box="[151,1436,820,843]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F59259A090BB58" box="[151,1436,820,843]" pageId="7" pageNumber="323">
• In hindwing, postmedial line represented by series of dark brown spots on each veins......................
<taxonomicName id="F48E4D565C17FFD1E1419259A090BB58" authorityName="Prout" authorityYear="1915" box="[1315,1436,820,842]" class="Insecta" family="Geometridae" genus="Xenimpia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="repudiosa">
<emphasis id="01FAEAC75C17FFD1E1419259A090BB58" bold="true" box="[1315,1436,820,842]" italics="true" pageId="7" pageNumber="323">P. repudiosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="887A2E705C17FFD1E4F5923EA090BBAD" pageId="7" pageNumber="323">
<paragraph id="333136D55C17FFD1E4F5923EA090BB97" blockId="7.[151,1437,615,960]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F5923EA090BB97" pageId="7" pageNumber="323">
4. In forewing, medial line partially doubled between the inner margin to lower edge of the cell, then continues as single, obliquely straight towards the costal margin...............................................................
<taxonomicName id="F48E4D565C17FFD1E145921CA090BB97" authorityName="Warren" authorityYear="1898" box="[1319,1436,879,901]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="obliquata">
<emphasis id="01FAEAC75C17FFD1E145921CA090BB97" bold="true" box="[1319,1436,879,901]" italics="true" pageId="7" pageNumber="323">P. obliquata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="333136D55C17FFD1E4F592E1A090BBAD" blockId="7.[151,1437,615,960]" pageId="7" pageNumber="323">
<keyLead id="887F95E05C17FFD1E4F592E1A090BBAD" pageId="7" pageNumber="323">
• In forewing, medial line broad, not doubled between the inner margin to lower edge of cell, then is display an inward curve towards the costal margin..................................................................
<emphasis id="01FAEAC75C17FFD1E09792C7A090BBAD" bold="true" box="[1269,1436,938,960]" pageId="7" pageNumber="323">
<taxonomicName id="F48E4D565C17FFD1E09792C7A045BBD2" authorityName="Singh &amp; Lenka &amp; Chatterjee &amp; Singh" authorityYear="2025" box="[1269,1353,938,960]" class="Insecta" family="Geometridae" genus="Psilocladia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="323" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
<emphasis id="01FAEAC75C17FFD1E09792C7A045BBD2" bold="true" box="[1269,1353,938,960]" italics="true" pageId="7" pageNumber="323">P. indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AC957BC5C17FFD1E13292C6A090BBAD" box="[1360,1436,938,959]" pageId="7" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>