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<document id="02F844D1262A491122E58B329196BA16" ID-DOI="10.11646/phytotaxa.314.1.15" ID-ISSN="1179-3163" ID-Zenodo-Dep="13686194" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725481839570" checkinUser="felipe" docAuthor="Pham, Van The, Vu, Tien Chinh, Rajapaksha, Ranil, Trinh, Ngoc Bon, Averyanov, Leonid &amp; Nguyen, Thi Lien Thuong" docDate="2017" docId="062087DF1074FFEBFF29FF26FA03459E" docLanguage="en" docName="phytotaxa.314.1.15.pdf" docOrigin="Phytotaxa 314 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.314.1.15" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phyllagathis phamhoangii V. T. Pham, T. C. Vu &amp; Ranil 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="141" masterDocId="FA19FFA71075FFEAFFA1FFD2FF894251" masterDocTitle="Phyllagathis phamhoangii (Sonerileae, Melastomataceae), a new species from central Vietnam" masterLastPageNumber="144" masterPageNumber="140" pageNumber="141" updateTime="1725482351818" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E6C9CF8576BC51D8BE88C1071F91B97E">Phyllagathis phamhoangii (Sonerileae, Melastomataceae), a new species from central Vietnam</mods:title>
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<mods:namePart id="751872A613C28336E2353CA8116B5E87">Pham, Van The</mods:namePart>
<mods:affiliation id="25B2DF500D1E9EC047BFA34AF2D5E2DE">Department of Plant Resource, Institute of Ecology and Biological Resources, Vietnam Academy of Sciences and Technology, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="477AEA5D4276981868ADF7D19C84F9F4" type="email">phamvthe@gmail.com</mods:nameIdentifier>
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<mods:namePart id="904D62B73809455A5AB6234E7E5B58C6">Vu, Tien Chinh</mods:namePart>
<mods:affiliation id="32F6ABA17E5473E2BD53D78B992B291C">Department of Specimen Collection Management, Vietnam National Museum of Nature, Vietnam Academy of Sciences and Technology, Hanoi, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Sciences and Technology, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="A2D147FAA1ACBD23896A36382234FF96">Rajapaksha, Ranil</mods:namePart>
<mods:affiliation id="97779FB5F48A130554C28DAF49FD4F2B">Department of Crop Science, Faculty of Agriculture, University of Peradeniya, Peradeniya, Sri Lanka.</mods:affiliation>
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<mods:namePart id="ABE177198C1743B0209593FF8E491988">Trinh, Ngoc Bon</mods:namePart>
<mods:affiliation id="D85FE4A2E1829D3298873B5F0C4EEDCB">Department of Plant Resource, Silviculture Research Institute, Vietnamese Academy of Forest Sciences, Duc Thang ward, Tu Liem Northern, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="4EB511EDB1CF1900416D574BDBEAABF5">Averyanov, Leonid</mods:namePart>
<mods:affiliation id="A29C6CCA6B430FD77F3A30EB12D3A88C">Komarov Botanical Institute, Russian Academy of Science, Prof. Popov Str. 2, RU- 197376 St. - Petersburg, Russia.</mods:affiliation>
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<mods:namePart id="8B7DABBC83317BD1904675644E3D407E">Nguyen, Thi Lien Thuong</mods:namePart>
<mods:affiliation id="A898390AE47ECE230B74D99FFDD0EAA4">Center of Scientific Research and Practice, Thu Dau Mot University, No 06, Tran Van On street, Phu Hoa ward, Thu Dau Mot city, Binh Duong province, Vietnam.</mods:affiliation>
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<taxonomicName id="49894D4A1074FFEBFF29FF26FD67435E" ID-CoL="7WLZY" authority="V.T.Pham, T.C.Vu &amp; Ranil" authorityName="V. T. Pham, T. C. Vu &amp; Ranil" authorityYear="2017" box="[136,750,244,271]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="phamhoangii" status="sp. nov.">
<emphasis id="BCFDEADB1074FFEBFF29FF26FE33435E" bold="true" box="[136,442,244,271]" italics="true" pageId="1" pageNumber="141">Phyllagathis phamhoangii</emphasis>
V.T.Pham, T.C.Vu &amp; Ranil
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,
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(
<figureCitation id="16B22A4C1074FFEBFCFCFF24FC17435E" box="[861,926,246,271]" captionStart="FIGURE 1" captionStartId="2.[136,229,1813,1834]" captionTargetBox="[190,1398,171,1784]" captionTargetId="figure-14@2.[190,1398,171,1784]" captionTargetPageId="2" captionText="FIGURE 1. Phyllagathis phamhoangii V.T.Pham,T.C.Vu &amp; Ranil.a. flowering plant in its natural habitat.b. flattened flowering specimen. c. portion of stem. d. leaf adaxial surface. e. leaf abaxial surface.f. leaf blade margin, adaxial surface. g. leaf blade margin, abaxial surface. h, i. inflorescence. j, k. petals adaxial (left) and abaxial (right) views. l, n. removed stamens. m. intact stamens. o. removed antepetalous stamen. p. style. q. style and ovary, longitudinal section. r. flower bud. s. petals, transverse section of flower bud, view from below. t. transverse section of flower bud, view from above. u. inflorescence with flower buds and young fruits. v. young fruit, view from above. w. young fruit transverse section. x. young fruit longitudinal section. y. young seeds. z. scape apex. Scale bars: a, b, d, e = 5cm; c = 1cm; f, g, h, i, j, k, m = 0.5cm; l, n, o, p, q, r, s, t, v, w, s, y = 1 mm; u = 5 mm; z = 2 mm. Photos and design by Pham Van The based on the type, HAL 11790 (LE, VNMN)." figureDoi="http://doi.org/10.5281/zenodo.13686196" httpUri="https://zenodo.org/record/13686196/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
)
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<emphasis id="BCFDEADB1074FFEBFF29FECEFF504360" bold="true" box="[136,217,284,305]" pageId="1" pageNumber="141">Type:—</emphasis>
<materialsCitation id="3EE13C941074FFEBFF79FECEFB3B4328" collectingDate="2011-05-19" collectorName="L. Averyanov &amp; P. K. Loc &amp; N. Q. Hieu &amp; P. V. The &amp; N. T. Vinh" country="Vietnam" county="Dai Loc District" elevation="500" latitude="15.828889" location="Dai Hong town" longLatPrecision="21" longitude="107.95528" municipality="Dai Hong Municipality" pageId="1" pageNumber="141" specimenCount="2" stateProvince="Quang Nam" typeStatus="holotype">
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.
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,
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, environs of
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, on large shady mossy boulders on very steep humid rocky slope in rich secondary broad-leaved evergreen forest,
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.,
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,
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, CPC 3419
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, (
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: HN!;
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: LE!)
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.
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Acaulescent rhizomatous herb, with leaves up to
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tall, sparsely hairy in all parts except petals and stamens, fewer on the leaves.
<emphasis id="BCFDEADB1074FFEBFE87FE1CFE0F43B6" box="[294,390,462,487]" italics="true" pageId="1" pageNumber="141">Rhizome</emphasis>
creeping to suberect, tuberous, orange-brown setose,
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in diameter, with conspicuous leaf scars.
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35, petiolate; petiole
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long; leaf blade cordate to reniform,
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wide,
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long, at base cordate, apex shortly acuminate, margin finely denticulate; lamina with 4 pairs of basal veins and a midrib, membranous, adaxial surface often light green, abaxial surface often light purple with prominent veins.
<emphasis id="BCFDEADB1074FFEBFAB7FDE8FA254002" box="[1302,1452,570,595]" italics="true" pageId="1" pageNumber="141">Inflorescence</emphasis>
a dichasial cyme; peduncle reddish,
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long, cylindrical, usually longer than petioles, with minute glands.
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4-merous, subtended by a few subulate bracts
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long; pedicel ca.
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long, with minute glands.
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campanulate, 4-ridged, about
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long and
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wide, with minute brown glands.
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triangular, about
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long, acute.
<emphasis id="BCFDEADB1074FFEBFE10FD18FE7E40B2" box="[433,503,714,739]" italics="true" pageId="1" pageNumber="141">Petals</emphasis>
pink, broadly obovate to almost round, obtuse or, shortly acuminate, glabrous,
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long,
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wide.
<emphasis id="BCFDEADB1074FFEBFE60FD3CFD974156" box="[449,542,750,775]" italics="true" pageId="1" pageNumber="141">Stamens</emphasis>
8, dimorphic, unequal in length, arranged in 2 whorls; antesepalous stamens 4, filaments
<quantity id="49719B2C1074FFEBFF56FCC0FEE0417A" box="[247,361,786,811]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="1" pageNumber="141" unit="mm" value="7.75" valueMax="8.0" valueMin="7.5">7.58 mm</quantity>
long, glabrous, anthers narrowly ovate, glabrous,
<quantity id="49719B2C1074FFEBFC3BFCC0FB85417A" box="[922,1036,786,811]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="1" pageNumber="141" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
long, light violet, strongly curved on ventral side, connective with an inflated base, ventrally with two laterally compressed lobes, pore 1, apical, more or less oblique on the ventral side; antepetalous stamens 4, filaments
<quantity id="49719B2C1074FFEBFCCBFC88FC364122" box="[874,959,858,883]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="141" unit="mm" value="5.5">5.5 mm</quantity>
long, anthers narrowly ovate,
<quantity id="49719B2C1074FFEBFAB9FC88FAE44122" box="[1304,1389,858,883]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="141" unit="mm" value="2.5">2.5 mm</quantity>
long, yellow, glabrous, connective with a short dorsal spur.
<emphasis id="BCFDEADB1074FFEBFD48FCACFCA641C6" box="[745,815,894,919]" italics="true" pageId="1" pageNumber="141">Ovary</emphasis>
4-locular, ca.
<quantity id="49719B2C1074FFEBFC70FCACFBA141C6" box="[977,1064,894,919]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="141" unit="mm" value="2.5">2.5 mm</quantity>
long, a little longer than one third of hypanthium length, crown consisting of four thick connate lobes.
<emphasis id="BCFDEADB1074FFEBFC26FC70FC4141EA" box="[903,968,930,955]" italics="true" pageId="1" pageNumber="141">Styles</emphasis>
ca.
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long.
<emphasis id="BCFDEADB1074FFEBFBDCFC70FB6D41EA" box="[1149,1252,930,955]" italics="true" pageId="1" pageNumber="141">Capsules</emphasis>
subquadrangular, cup-shaped, about
<quantity id="49719B2C1074FFEBFEF8FC14FE11418E" box="[345,408,966,991]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="141" unit="mm" value="3.0">3 mm</quantity>
wide and
<quantity id="49719B2C1074FFEBFDA9FC14FDED418E" box="[520,612,966,991]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="141" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long (when dry), crown persistent.
<emphasis id="BCFDEADB1074FFEBFC4CFC14FBA4418E" box="[1005,1069,966,991]" italics="true" pageId="1" pageNumber="141">Seeds</emphasis>
numerous, ca. 0.25 ×
<quantity id="49719B2C1074FFEBFA81FC14FAE9418E" box="[1312,1376,966,991]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="141" unit="mm" value="1.0">1 mm</quantity>
(when young), narrowly conical, glabrous.
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CFBDCFBB34676" blockId="1.[136,1455,426,1999]" box="[189,1082,1038,1063]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CFBDCFED64676" bold="true" box="[189,351,1038,1063]" pageId="1" pageNumber="141">Phenology:—</emphasis>
Flowering was observed in April to May. Fruiting started in May.
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CFBE0FAB34772" blockId="1.[136,1455,426,1999]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CFBE0FE62461A" bold="true" box="[189,491,1074,1099]" pageId="1" pageNumber="141">Diagnostic characters:—</emphasis>
The described species is closely allied to
<taxonomicName id="49894D4A1074FFEBFC42FBE0FB28461A" box="[995,1185,1074,1099]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="nanakorniana">
<emphasis id="BCFDEADB1074FFEBFC42FBE0FB28461A" box="[995,1185,1074,1099]" italics="true" pageId="1" pageNumber="141">P. nanakorniana</emphasis>
</taxonomicName>
from
<collectingCountry id="F69E76591074FFEBFB50FBE0FADC461A" box="[1265,1365,1074,1099]" name="Thailand" pageId="1" pageNumber="141">Thailand</collectingCountry>
and
<taxonomicName id="49894D4A1074FFEBFA36FBE0FE8E463E" authorityName="C. Hansen" authorityYear="1990" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="subrotunda">
<emphasis id="BCFDEADB1074FFEBFA36FBE0FE8E463E" italics="true" pageId="1" pageNumber="141">P. subrotunda</emphasis>
</taxonomicName>
from north of
<collectingCountry id="F69E76591074FFEBFE0EFB84FD87463E" box="[431,526,1110,1135]" name="Vietnam" pageId="1" pageNumber="141">Vietnam</collectingCountry>
in habit and some morphologic characters, such as the tuberous rhizome and leaf shape. However, it differs from
<taxonomicName id="49894D4A1074FFEBFE4DFBA8FD2D46C2" box="[492,676,1146,1171]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="nanakorniana">
<emphasis id="BCFDEADB1074FFEBFE4DFBA8FD2D46C2" box="[492,676,1146,1171]" italics="true" pageId="1" pageNumber="141">P. nanakorniana</emphasis>
</taxonomicName>
in the sparse hairs on all of its parts except petals and stamens, much longer petioles, dichasial inflorescences, longer hypanthia, stamens unequal in length, longer petals and ovary. It differs from
<taxonomicName id="49894D4A1074FFEBFEB6FB10FE3A468A" authorityName="C. Hansen" authorityYear="1990" box="[279,435,1218,1243]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="subrotunda">
<emphasis id="BCFDEADB1074FFEBFEB6FB10FE3A468A" box="[279,435,1218,1243]" italics="true" pageId="1" pageNumber="141">P. subrotunda</emphasis>
</taxonomicName>
by the taller plants, smaller leaves with longer petioles, cordate to reniform blades with a shortly acuminate apex, dichasial inflorescences, longer and cylindrical peduncles, longer and narrower hypanthia. The detailed comparison between
<emphasis id="BCFDEADB1074FFEBFDA9FAD8FCF74772" box="[520,894,1290,1315]" italics="true" pageId="1" pageNumber="141">
<taxonomicName id="49894D4A1074FFEBFDA9FAD8FD324772" box="[520,699,1290,1315]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="phamhoangii">P. phamhoangii</taxonomicName>
,
<taxonomicName id="49894D4A1074FFEBFD64FAD8FCF74772" box="[709,894,1290,1315]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="nanakorniana">P. nanakorniana</taxonomicName>
</emphasis>
and
<taxonomicName id="49894D4A1074FFEBFC15FAD8FBC64772" authorityName="C. Hansen" authorityYear="1990" box="[948,1103,1290,1315]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="subrotunda">
<emphasis id="BCFDEADB1074FFEBFC15FAD8FBC64772" box="[948,1103,1290,1315]" italics="true" pageId="1" pageNumber="141">P. subrotunda</emphasis>
</taxonomicName>
are given in
<tableCitation id="C30B03721074FFEBFB41FAD8FABD4772" box="[1248,1332,1290,1315]" captionStart="TABLE 1" captionStartId="3.[136,224,1234,1257]" captionText="TABLE 1. Comparison of diagnostic features of Phyllagathis phamhoangii with P. nanakorniana and P. subrotunda." pageId="1" pageNumber="141">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C69365421074FFEBFF1CFAFCFEEC4482" pageId="1" pageNumber="141" type="distribution">
<paragraph id="8E3636C91074FFEBFF1CFAFCFA0D4716" blockId="1.[136,1455,426,1999]" box="[189,1412,1326,1351]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CFAFCFEFC4716" bold="true" box="[189,373,1326,1351]" pageId="1" pageNumber="141">Distribution:—</emphasis>
The species is only known from
<collectingRegion id="4C4DF82B1074FFEBFD43FAFCFCE34716" box="[738,874,1326,1351]" country="Vietnam" name="Quang Nam" pageId="1" pageNumber="141">Quang Nam</collectingRegion>
and
<collectingRegion id="4C4DF82B1074FFEBFC00FAFCFBA04716" box="[929,1065,1326,1351]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="141">Quang Binh</collectingRegion>
Provinces of
<collectingCountry id="F69E76591074FFEBFB62FAFCFAAB4716" box="[1219,1314,1326,1351]" name="Vietnam" pageId="1" pageNumber="141">Vietnam</collectingCountry>
(
<figureCitation id="16B22A4C1074FFEBFA90FAFCFAFE4716" box="[1329,1399,1326,1351]" captionStart="FIGURE 2" captionStartId="3.[136,229,1128,1149]" captionTargetBox="[151,1435,190,1103]" captionTargetId="figure-17@3.[151,1436,190,1103]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Phyllagathis phamhoangii V.T.Pham, T.C.Vu &amp; Ranil. Locations in Quang Binh and Quang Nam provinces are marked by black rounds." figureDoi="http://doi.org/10.5281/zenodo.13686198" httpUri="https://zenodo.org/record/13686198/files/figure.png" pageId="1" pageNumber="141">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CFA80FF5F47E2" blockId="1.[136,1455,426,1999]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CFA80FE25473A" bold="true" box="[189,428,1362,1387]" pageId="1" pageNumber="141">Habitat and ecology</emphasis>
:
<emphasis id="BCFDEADB1074FFEBFE15FA80FE59473A" bold="true" box="[436,464,1362,1387]" pageId="1" pageNumber="141"></emphasis>
The new species grows in rich secondary broad-leaved evergreen forests at an elevations of
<quantity id="49719B2C1074FFEBFF09FAA4FEA047DE" box="[168,297,1398,1423]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="141" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</quantity>
as a lithophytic herb on large wet mossy boulders and cliffs composed of eroded sandstone on steep slopes.
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CFA6CFEEC4482" blockId="1.[136,1455,426,1999]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CFA6CFE584786" bold="true" box="[189,465,1470,1495]" pageId="1" pageNumber="141">Conservation status:—</emphasis>
<taxonomicName id="49894D4A1074FFEBFE70FA6CFD724786" box="[465,763,1470,1495]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="phamhoangii">
<emphasis id="BCFDEADB1074FFEBFE70FA6CFD724786" box="[465,763,1470,1495]" italics="true" pageId="1" pageNumber="141">Phyllagathis phamhoangii</emphasis>
</taxonomicName>
was recorded only from two small subpopulations in
<collectingRegion id="4C4DF82B1074FFEBFAC0FA6CFF3747AA" country="Vietnam" name="Quang Nam" pageId="1" pageNumber="141">Quang Nam</collectingRegion>
and
<collectingRegion id="4C4DF82B1074FFEBFF59FA30FE0847AA" box="[248,385,1506,1531]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="141">Quang Binh</collectingRegion>
provinces of
<collectingCountry id="F69E76591074FFEBFDBFFA30FD0847AA" box="[542,641,1506,1531]" name="Vietnam" pageId="1" pageNumber="141">Vietnam</collectingCountry>
. Though it is locally abundant, existing populations are facing a risk of extinction in the wild, since the areas where this species is found do not belong to any protected forest. The habitats are highly disturbed by local people for collecting timber, fire-wood and non-timber forest products. In addition, the new species has beautiful plants and flowers, and may be used as ornamental and collected by local people in the future. The estimated total known area of its distribution is less than
<quantity id="49719B2C1074FFEBFCAFF9A0FC0E44DA" box="[782,903,1650,1675]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="141" unit="km" value="20.0">20.000 km</quantity>
<superScript id="79FC9B811074FFEBFC26F9A2FC06442F" attach="left" box="[903,911,1648,1662]" fontSize="6" pageId="1" pageNumber="141">2</superScript>
, in habitats that are seriously threatened. Based on our information, we propose a conservation status of
<taxonomicName id="49894D4A1074FFEBFD5AF944FC2344FE" box="[763,938,1686,1711]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="141" phylum="Tracheophyta" rank="species" species="phamhoangii">
<emphasis id="BCFDEADB1074FFEBFD5AF944FC2344FE" box="[763,938,1686,1711]" italics="true" pageId="1" pageNumber="141">P. phamhoangii</emphasis>
</taxonomicName>
as Vulnerable (VU) under IUCN criteria B1 a b(iii) (
<bibRefCitation id="EA184B381074FFEBFF6EF968FED04482" author="IUCN Standards &amp; Petitions Subcommittee" box="[207,345,1722,1747]" pageId="1" pageNumber="141" refId="ref2787" refString="IUCN Standards and Petitions Subcommittee (2017) Guidelines for Using the IUCN Red List Categories and Criteria, Version 13. Prepared by the Standards and Petitions Subcommittee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 24 June 2017)" type="url" year="2017">IUCN 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C69365421074FFEBFF1CF90CFA03459E" pageId="1" pageNumber="141" type="etymology">
<paragraph id="8E3636C91074FFEBFF1CF90CFF66456E" blockId="1.[136,1455,426,1999]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CF90CFEED44A6" bold="true" box="[189,356,1758,1783]" pageId="1" pageNumber="141">Etymology:—</emphasis>
The species is named after Prof. Pham Hoang Ho, who spent most of his life studying the flora of
<collectingCountry id="F69E76591074FFEBFF29F8D0FF62454A" box="[136,235,1794,1819]" name="Vietnam" pageId="1" pageNumber="141">Vietnam</collectingCountry>
. He has published many books on the flora which are useful for students, teachers and researchers all over
<collectingCountry id="F69E76591074FFEBFF29F8F4FF62456E" box="[136,235,1830,1855]" name="Vietnam" pageId="1" pageNumber="141">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CF898FCE14532" blockId="1.[136,1455,426,1999]" box="[189,872,1861,1891]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CF898FE1C4532" bold="true" box="[189,405,1866,1891]" pageId="1" pageNumber="141">Vernacular name:</emphasis>
—Vietnamese: Me nguồn phạm hoàng hộ
</paragraph>
<paragraph id="8E3636C91074FFEBFF1CF8BCFA03459E" blockId="1.[136,1455,426,1999]" pageId="1" pageNumber="141">
<emphasis id="BCFDEADB1074FFEBFF1CF8BCFD5C45D6" bold="true" box="[189,725,1902,1927]" pageId="1" pageNumber="141">
Additional specimen examined:—
<collectingCountry id="F69E76591074FFEBFDEFF8BCFD5C45D6" box="[590,725,1902,1927]" name="Vietnam" pageId="1" pageNumber="141">VIETNAM</collectingCountry>
</emphasis>
.
<collectingRegion id="4C4DF82B1074FFEBFD40F8BCFC5845D6" box="[737,977,1902,1927]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="141">Quang Binh province</collectingRegion>
, Minh Hoa District, Dan Hoa Municipality, Giang Man Mountain, Vietnam-Laotian border, about
<quantity id="49719B2C1074FFEBFD4CF840FCAE45FA" box="[749,807,1938,1963]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="141" unit="km" value="6.0">6 km</quantity>
to N of Cha Lo border station, ca.
<quantity id="49719B2C1074FFEBFB17F840FA8445FA" box="[1206,1293,1938,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="141" unit="m" value="1100.0">1100 m</quantity>
,
<geoCoordinate id="EBBD500E1074FFEBFAB8F840FA2E45FA" box="[1305,1447,1938,1963]" degrees="17" direction="north" minutes="41" orientation="latitude" pageId="1" pageNumber="141" precision="15" seconds="09" value="17.685833">17º4109N</geoCoordinate>
,
<geoCoordinate id="EBBD500E1074FFEBFF29F864FEA9459E" box="[136,288,1974,1999]" degrees="105" direction="east" minutes="45" orientation="longitude" pageId="1" pageNumber="141" precision="15" seconds="54" value="105.765">105º4554E</geoCoordinate>
,
<date id="FA3710091074FFEBFE8DF864FE43459E" box="[300,458,1974,1999]" pageId="1" pageNumber="141" value="2008-04-21">21 April 2008</date>
,
<emphasis id="BCFDEADB1074FFEBFE74F864FBEC459E" box="[469,1125,1974,1999]" italics="true" pageId="1" pageNumber="141">L. Averyanov, P.K. Loc, N.T. Vinh, N.S. Khang, HAL 11790</emphasis>
(HN!, HNU!, MO!, LE!).
</paragraph>
</subSubSection>
</treatment>
</document>