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1a. Pale interocular bar present; usually eight (rarely nine) supralabial scales...........................................................................
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1b. Pale interocular bar absent; usually nine (rarely eight) supralabial scales..................................................................
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<emphasis id="18D78C1DFF9CFF8CFCE20E7A4C9A5866" box="[818,1169,1115,1138]" italics="true" pageId="33" pageNumber="34">Sibynophis geminatus geminatus</emphasis>
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<emphasis id="18D78C1DFF9CFF8CFD9B0EAF4ACE58B1" bold="true" box="[587,709,1166,1189]" italics="true" pageId="33" pageNumber="34">Stegonotus</emphasis>
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Represented in the
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by a single species......................
<taxonomicName id="EDA32B8CFF9CFF8CFC6C0E994C9A58DB" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[956,1169,1208,1231]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="18D78C1DFF9CFF8CFC6C0E994C9A58DB" box="[956,1169,1208,1231]" italics="true" pageId="33" pageNumber="34">Stegonotus muelleri</emphasis>
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1a. Tail color distinctly different from body color (in life: tail color red, body color green); hemipenes spinose...........................................
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2a. Body and tail color green (adults and some juveniles in life) or red (some juveniles in life)......................................................
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2b. Body and tail color yellow, gray, or white (in life)...............
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1a. Longitudinal rows of dorsal body scales at midbody 21–23 (males), 21–29 (females); dorsal scales on head 9–16 along a line passing between the supraocular scales (
<figureCitation id="B2984C8AFF9CFF8CFC740CBF4C0C5AA1" box="[932,1031,1694,1717]" captionStart="FIGURE" captionStartId="34.[155,167,832,851]" captionTargetBox="[293,1003,545,819]" captionTargetId="figure-375@34.[293,1003,545,819]" captionTargetPageId="34" captionText="FIGURE 46. Dorsal view of head of (A) Tropidolaemus subannulatus (KU 324585), (B) Tropidolaemus philippensis (KU 334672). Dorsal scales of head along line between supraocular scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512778" httpUri="https://zenodo.org/record/11512778/files/figure.png" pageId="33" pageNumber="34">Fig. 46A</figureCitation>
); third supralabial scale usually separated from subocular scale by one or two scales; color variable, shades of green, blue or green, and blue in females, and green in males, but blue in some Negros Island populations; crossbands on body blue and white, red and white, or blue, red and white; color of postocular stripe variable (females), white or red (males), or red (juveniles)....................................................................
<taxonomicName id="EDA32B8CFF9FFF8FFC8C0B7D4C9A5D67" ID-CoL="7D8Q6" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[860,1169,348,371]" class="Reptilia" family="Viperidae" genus="Tropidolaemus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="subannulatus">
<emphasis id="18D78C1DFF9FFF8FFC8C0B7D4C9A5D67" box="[860,1169,348,371]" italics="true" pageId="34" pageNumber="35">Tropidolaemus subannulatus</emphasis>
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</paragraph>
<paragraph id="2A1C500FFF9FFF8FFFAB0B5D4C9A5E07" blockId="34.[123,1174,220,531]" pageId="34" pageNumber="35">
<keyLead id="9152F33AFF9FFF8FFFAB0B5D4C9A5E07" pageId="34" pageNumber="35">
1b. Longitudinal rows of dorsal body scales at midbody 18–19 (males and females); dorsal scales on head 6–8 along a line between the supraocular scales (
<figureCitation id="B2984C8AFF9FFF8FFCD00BBD4B695DA7" box="[768,866,412,435]" captionStart="FIGURE" captionStartId="34.[155,167,832,851]" captionTargetBox="[293,1003,545,819]" captionTargetId="figure-375@34.[293,1003,545,819]" captionTargetPageId="34" captionText="FIGURE 46. Dorsal view of head of (A) Tropidolaemus subannulatus (KU 324585), (B) Tropidolaemus philippensis (KU 334672). Dorsal scales of head along line between supraocular scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512778" httpUri="https://zenodo.org/record/11512778/files/figure.png" pageId="34" pageNumber="35">Fig. 46B</figureCitation>
); third supralabial scale usually in contact with subocular scale (rarely separated by one scale); in life: background body color turquoise-green (males) or green (females); postocular stripe usually black (rarely white)......................................................
<taxonomicName id="EDA32B8CFF9FFF8FFCBA0BDD4C9A5E07" ID-CoL="59BSC" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[874,1169,508,531]" class="Reptilia" family="Viperidae" genus="Tropidolaemus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="philippensis">
<emphasis id="18D78C1DFF9FFF8FFCBA0BDD4C9A5E07" box="[874,1169,508,531]" italics="true" pageId="34" pageNumber="35">Tropidolaemus philippensis</emphasis>
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<mods:title id="3454081EEA25667879415107C95EA04A">Illustrated Key to the Snakes of the Philippines</mods:title>
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<paragraph id="2A1C500FFF9DFF8DFD960FD54AC15A1F" blockId="32.[508,789,1524,1579]" box="[582,714,1524,1547]" pageId="32" pageNumber="33">
<heading id="7154E763FF9DFF8DFD960FD54AC15A1F" box="[582,714,1524,1547]" centered="true" fontSize="10" level="2" pageId="32" pageNumber="33" reason="4">
<taxonomicName id="EDA32B8CFF9DFF8DFD960FD54AC15A1F" ID-CoL="7846" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[582,714,1524,1547]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFF9DFF8DFD960FD54AC15A1F" bold="true" box="[582,714,1524,1547]" italics="true" pageId="32" pageNumber="33">Rhabdophis</emphasis>
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<subSubSection id="62B90384FF9DFF8DFE2C0C354B1E5A3F" box="[508,789,1556,1579]" pageId="32" pageNumber="33" type="multiple">
<paragraph id="2A1C500FFF9DFF8DFE2C0C354B1E5A3F" blockId="32.[508,789,1524,1579]" box="[508,789,1556,1579]" pageId="32" pageNumber="33">Key to Philippine species.</paragraph>
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<keyLead id="9152F33AFF9DFF8DFFAB0C1E4C9A5A62" pageId="32" pageNumber="33">1a. Dorsal body scales in 15–17 longitudinal rows at midbody; background color of ventral body scales black................................................................. 2</keyLead>
</paragraph>
<paragraph id="2A1C500FFF9DFF8DFFAB0C5E4C9A5AA2" blockId="32.[123,1173,1599,1718]" pageId="32" pageNumber="33">
<keyLead id="9152F33AFF9DFF8DFFAB0C5E4C9A5AA2" pageId="32" pageNumber="33">1b. Dorsal body scales in 17–19 longitudinal rows at midbody; background color of ventral body scales pale.................................................................. 3</keyLead>
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<keyLead id="9152F33AFF9CFF8CFFAB0AFD4C9A5D07" pageId="33" pageNumber="34">
2a. Light lateral body stripe not continuous across angle of jaw, separated from white patch behind eye............................................
<taxonomicName id="EDA32B8CFF9CFF8CFCC00ADD4C9A5D07" ID-CoL="8SB2N" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[784,1169,252,275]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="subSpecies" species="auriculatus" subSpecies="auriculatus">
<emphasis id="18D78C1DFF9CFF8CFCC00ADD4C9A5D07" box="[784,1169,252,275]" italics="true" pageId="33" pageNumber="34">Rhabdophis auriculatus auriculatus</emphasis>
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<keyLead id="9152F33AFF9CFF8CFFAB0B3D4C9A5D47" pageId="33" pageNumber="34">
2b. Light lateral body stripe continuous across angle of jaw to postocular region................................................................
<taxonomicName id="EDA32B8CFF9CFF8CFC920B1D4C9A5D47" ID-CoL="8J9V5" authorityName="Leviton" authorityYear="1970" box="[834,1169,316,339]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="subSpecies" species="auriculatus" subSpecies="myersi">
<emphasis id="18D78C1DFF9CFF8CFC920B1D4C9A5D47" box="[834,1169,316,339]" italics="true" pageId="33" pageNumber="34">Rhabdophis auriculatus myersi</emphasis>
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<keyStep id="915748AAFF9CFF8CFFAB0B7D4C9A5D87" pageId="33" pageNumber="34">
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<keyLead id="9152F33AFF9CFF8CFFAB0B7D4C9A5D67" box="[123,1169,348,371]" pageId="33" pageNumber="34">3a. Dorsal body scales in 17 longitudinal rows at midbody.............................. 4</keyLead>
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<paragraph id="2A1C500FFF9CFF8CFFAB0B5D4C9A5D87" blockId="33.[123,1174,220,403]" box="[123,1169,380,403]" pageId="33" pageNumber="34">
<keyLead id="9152F33AFF9CFF8CFFAB0B5D4C9A5D87" box="[123,1169,380,403]" pageId="33" pageNumber="34">3b. Dorsal body scales in 19 longitudinal rows at midbody.............................. 5</keyLead>
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4a. Pale dorsolateral body stripes present; pale nuchal spot present (except for hypermelanistic individuals from Lubang Island)...............................
<taxonomicName id="EDA32B8CFF9CFF8CFC440BE74C995DC9" ID-CoL="793Y8" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[916,1170,454,477]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="spilogaster">
<emphasis id="18D78C1DFF9CFF8CFC440BE74C995DC9" box="[916,1170,454,477]" italics="true" pageId="33" pageNumber="34">Rhabdophis spilogaster</emphasis>
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<keyLead id="9152F33AFF9CFF8CFFAB0BC74C9A5DE9" box="[123,1169,486,509]" pageId="33" pageNumber="34">
4b. Pale dorsolateral body stripes absent; pale nuchal spot absent........
<taxonomicName id="EDA32B8CFF9CFF8CFC7B0BC74C9A5DE9" ID-CoL="4RW4H" authorityName="Taylor" authorityYear="1922" box="[939,1169,486,509]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="18D78C1DFF9CFF8CFC7B0BC74C9A5DE9" box="[939,1169,486,509]" italics="true" pageId="33" pageNumber="34">Rhabdophis barbouri</emphasis>
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<keyLead id="9152F33AFF9CFF8CFFAB08304C9A5E5C" pageId="33" pageNumber="34">
5a. Distinct white stripe extends across supralabial scales; one anterior temporal scale, in contact with the sixth supralabial scale; usually two preocular scales (
<figureCitation id="B2984C8AFF9CFF8CFCE008104B995E5C" box="[816,914,561,584]" captionStart="FIGURE" captionStartId="33.[155,167,836,855]" captionTargetBox="[282,1014,663,824]" captionTargetId="figure-848@33.[282,1014,663,824]" captionTargetPageId="33" captionText="FIGURE 45. Lateral view of head of (A) Rhabdophis lineatus (KU 326696), (B) Rhabdophis chrysargos (composite drawing from photographs of multiple individuals). Anterior temporal scales (green), sixth supralabial scale (purplish-gray), preocular scales (blue)." figureDoi="http://doi.org/10.5281/zenodo.11512776" httpUri="https://zenodo.org/record/11512776/files/figure.png" pageId="33" pageNumber="34">Fig. 45A</figureCitation>
)..
<taxonomicName id="EDA32B8CFF9CFF8CFC6608104C9A5E5C" ID-CoL="4RW52" baseAuthorityName="Peters" baseAuthorityYear="1861" box="[950,1169,561,584]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="18D78C1DFF9CFF8CFC6608104C9A5E5C" box="[950,1169,561,584]" italics="true" pageId="33" pageNumber="34">Rhabdophis lineatus</emphasis>
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<paragraph id="2A1C500FFF9CFF8CFFAB08704C995E9C" blockId="33.[123,1173,529,648]" pageId="33" pageNumber="34">
<keyLead id="9152F33AFF9CFF8CFFAB08704C995E9C" pageId="33" pageNumber="34">
5b. No distinct white stripe on supralabial scales; two anterior temporal scales, neither in contact with the sixth supralabial scale; one preocular scale (
<figureCitation id="B2984C8AFF9CFF8CFD0008504B3B5E9C" box="[720,816,625,648]" captionStart="FIGURE" captionStartId="33.[155,167,836,855]" captionTargetBox="[282,1014,663,824]" captionTargetId="figure-848@33.[282,1014,663,824]" captionTargetPageId="33" captionText="FIGURE 45. Lateral view of head of (A) Rhabdophis lineatus (KU 326696), (B) Rhabdophis chrysargos (composite drawing from photographs of multiple individuals). Anterior temporal scales (green), sixth supralabial scale (purplish-gray), preocular scales (blue)." figureDoi="http://doi.org/10.5281/zenodo.11512776" httpUri="https://zenodo.org/record/11512776/files/figure.png" pageId="33" pageNumber="34">Fig. 45B</figureCitation>
)......
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1a. Dorsal body scales in 16 or 18 longitudinal rows on anterior third of body; mid-dorsal body scales keeled......................................................
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1b. Dorsal body scales in 14 longitudinal rows on anterior third of body; mid-dorsal body scales smooth.........................................................
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1a. Ocular scale overlaps third supralabial scale (
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)............
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1b. Third supralabial scale overlaps ocular scale (
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)..............................
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2a. Longitudinal scale rows around midbody 24–28...............
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<emphasis id="18D78C1DFF9DFF8DFCA90F364C9A593A" box="[889,1169,1303,1326]" italics="true" pageId="32" pageNumber="33">Ramphotyphlops cumingii</emphasis>
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44. Lateral view of head of (A)
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(FMNH 96520; a redrafting by EH of original image in
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(PBS 2001; a redrafting by EH of original image in
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). Ocular scale (green), third supralabial scale (purplish-gray). Fig. A reproduced and modified with permission of V. Wallach and Journal of Herpetology.
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3a. Width of preocular scale about equal to width of ocular scale; width of rostral scale ~0.75 head width; nasal scale nearly, but not completely, divided...........
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3b. Preocular scale much narrower than ocular scale; width of rostral scale about half the width of head; nasal scale completely divided.........................
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Represented by a single species....................................
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<emphasis id="18D78C1DFFA0FFB0FDBD090F4AA95F51" bold="true" box="[621,674,814,837]" italics="true" pageId="29" pageNumber="30">Naja</emphasis>
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1a. Dorsal body scales in 21–23 longitudinal rows at midbody; dorsal color uniformly light brown or olive......................................................
<taxonomicName id="EDA32B8CFFA0FFB0FC1009BB4C9A5FA5" ID-CoL="45KXX" authorityName="Taylor" authorityYear="1922" box="[960,1169,922,945]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="philippinensis">
<emphasis id="18D78C1DFFA0FFB0FC1009BB4C9A5FA5" box="[960,1169,922,945]" italics="true" pageId="29" pageNumber="30">Naja philippinensis</emphasis>
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<keyLead id="9152F33AFFA0FFB0FFAB099B4C9A5FE5" pageId="29" pageNumber="30">1b. Dorsal body scales in 17–19 longitudinal rows at midbody; dorsal color black or dark brown........................................................................... 2</keyLead>
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2a. A few anterior ventral body scales light yellow, followed by band of black that gradually fades posteriorly; 162–178 ventral body scales..............................
<taxonomicName id="EDA32B8CFFA0FFB0FC0F0E044C9A5828" ID-CoL="45KY2" authorityName="Peters" authorityYear="1861" box="[991,1169,1061,1084]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="samarensis">
<emphasis id="18D78C1DFFA0FFB0FC0F0E044C9A5828" box="[991,1169,1061,1084]" italics="true" pageId="29" pageNumber="30">Naja samarensis</emphasis>
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2b. Ventral body surface dark or light but without a distinctive black band on anterior portion; 178–189 ventral body scales........................................
<taxonomicName id="EDA32B8CFFA0FFB0FC340E474C9A5869" ID-CoL="45KY7" authorityName="Muller" authorityYear="1887" box="[996,1169,1126,1149]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="sumatrana">
<emphasis id="18D78C1DFFA0FFB0FC340E474C9A5869" box="[996,1169,1126,1149]" italics="true" pageId="29" pageNumber="30">Naja sumatrana</emphasis>
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<paragraph id="2A1C500FFFA0FFB0FE2C0E984B1E58C4" blockId="29.[508,789,1176,1232]" box="[508,789,1209,1232]" pageId="29" pageNumber="30">Key to Philippine species.</paragraph>
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<keyLead id="9152F33AFFA0FFB0FFAB0EC54C9A58EF" box="[123,1169,1252,1275]" pageId="29" pageNumber="30">1a. Dorsal body scales in 15 longitudinal rows at midbody.............................. 2</keyLead>
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2a. Light vertebral stripe present; no prominent dorsal body blotches; quadrangular black spots present on ventral body scales....................................
<taxonomicName id="EDA32B8CFFA0FFB0FC120F6E4C9A5972" ID-CoL="49632" authorityName="Gunther" authorityYear="1864" box="[962,1169,1359,1382]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="modestus">
<emphasis id="18D78C1DFFA0FFB0FC120F6E4C9A5972" box="[962,1169,1359,1382]" italics="true" pageId="29" pageNumber="30">Oligodon modestus</emphasis>
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2b. Light vertebral stripe absent; light dorsal body spots or blotches usually present; distinctive markings absent from ventral body scales..........................
<taxonomicName id="EDA32B8CFFA0FFB0FC670FAE4C9A59B2" ID-CoL="4963C" authorityName="Gunther" authorityYear="1873" box="[951,1169,1423,1446]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="notospilus">
<emphasis id="18D78C1DFFA0FFB0FC670FAE4C9A59B2" box="[951,1169,1423,1446]" italics="true" pageId="29" pageNumber="30">Oligodon notospilus</emphasis>
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3a. Number of supralabial scales usually six; number of anterior temporal scales two; no transverse markings on dorsal surface of body...............................
<taxonomicName id="EDA32B8CFFA0FFB0FC6C0FFB4C9A59E5" baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[956,1169,1498,1521]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="meyerinki">
<emphasis id="18D78C1DFFA0FFB0FC6C0FFB4C9A59E5" box="[956,1169,1498,1521]" italics="true" pageId="29" pageNumber="30">Oligodon meyerinki</emphasis>
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4a. Dorsal body blotches&gt; 30, irregularly shaped, forming poorly defined dark crossbands; number of ventral body scales&gt; 180; number of ventral body scales + subcaudal scales&gt; 230 (females; unknown for males).....................................
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<paragraph id="2A1C500FFFA3FFB3FFAB0AFD4C9A5D07" blockId="30.[123,1173,220,275]" pageId="30" pageNumber="31">
<keyLead id="9152F33AFFA3FFB3FFAB0AFD4C9A5D07" pageId="30" pageNumber="31">4b. Dorsal body blotches well-defined, numbering &lt;30; number of ventral body scales &lt;180; number of ventral body scales + subcaudal scales &lt;220.............................. 5</keyLead>
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</keyStep>
<keyStep id="915748AAFFA3FFB3FFAB0B074C9A5DA9" pageId="30" pageNumber="31">
<paragraph id="2A1C500FFFA3FFB3FFAB0B074C9A5D49" blockId="30.[123,1174,294,445]" pageId="30" pageNumber="31">
<keyLead id="9152F33AFFA3FFB3FFAB0B074C9A5D49" pageId="30" pageNumber="31">
5a. Dark dorsal body blotches extend laterally to ventral body scales; two preocular scales (
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); number of ventral body scales + subcaudal scales 211–216.......
<taxonomicName id="EDA32B8CFFA3FFB3FC670B674C9A5D49" ID-CoL="6SKQW" baseAuthorityName="Taylor" baseAuthorityYear="1918" box="[951,1169,326,349]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="18D78C1DFFA3FFB3FC670B674C9A5D49" box="[951,1169,326,349]" italics="true" pageId="30" pageNumber="31">Oligodon maculatus</emphasis>
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<paragraph id="2A1C500FFFA3FFB3FFAB0B474C9A5DA9" blockId="30.[123,1174,294,445]" pageId="30" pageNumber="31">
<keyLead id="9152F33AFFA3FFB3FFAB0B474C9A5DA9" pageId="30" pageNumber="31">
5b. Dark dorsal body blotches saddle-shaped, not extending to ventral body scales; usually one preocular scale (
<figureCitation id="B2984C8AFFA3FFB3FE980BA749A15D89" box="[328,426,390,413]" captionStart="FIGURE" captionStartId="30.[155,167,667,686]" captionTargetBox="[293,1003,454,652]" captionTargetId="figure-542@30.[293,1003,454,652]" captionTargetPageId="30" captionText="FIGURE 41. Lateral view of head of (A) Oligodon maculatus (KU 321699), (B) Oligodon ancorus (KU 346379). Preoc- ular scales (green)." figureDoi="http://doi.org/10.5281/zenodo.11512764" httpUri="https://zenodo.org/record/11512764/files/figure.png" pageId="30" pageNumber="31">Fig. 41B</figureCitation>
); number of ventral body scales + subcaudal scales 183–207...................................................................
<taxonomicName id="EDA32B8CFFA3FFB3FC1F0B874C9A5DA9" ID-CoL="495Z6" baseAuthorityName="Girard" baseAuthorityYear="1858" box="[975,1169,422,445]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="ancorus">
<emphasis id="18D78C1DFFA3FFB3FC1F0B874C9A5DA9" box="[975,1169,422,445]" italics="true" pageId="30" pageNumber="31">Oligodon ancorus</emphasis>
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<emphasis id="18D78C1DFFA1FFB1FDE108574AEB5E99" bold="true" box="[561,736,630,653]" italics="true" pageId="28" pageNumber="29">Malayotyphlops</emphasis>
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1a. Small subocular scale present, in contact with second and third supralabial scales, preocular scale, and ocular scale......................................
<taxonomicName id="EDA32B8CFFA1FFB1FC5808C04C9A5EEC" ID-CoL="88L24" baseAuthorityName="Taylor" baseAuthorityYear="1919" box="[904,1169,737,760]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="manilae">
<emphasis id="18D78C1DFFA1FFB1FC5808C04C9A5EEC" box="[904,1169,737,760]" italics="true" pageId="28" pageNumber="29">Malayotyphlops manilae</emphasis>
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<paragraph id="2A1C500FFFA1FFB1FFAB09204C9A5F0C" blockId="28.[123,1174,705,792]" box="[123,1169,769,792]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB09204C9A5F0C" box="[123,1169,769,792]" pageId="28" pageNumber="29">1b. Small subocular scale absent................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA1FFB1FFAB090A4C9A5F96" pageId="28" pageNumber="29">
<paragraph id="2A1C500FFFA1FFB1FFAB090A4C9A5F56" blockId="28.[123,1173,811,898]" box="[123,1169,811,834]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB090A4C9A5F56" box="[123,1169,811,834]" pageId="28" pageNumber="29">2a. Dorsal body scales in &lt;29 longitudinal rows at midbody............................ 3</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA1FFB1FFAB096A4C9A5F96" blockId="28.[123,1173,811,898]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB096A4C9A5F96" pageId="28" pageNumber="29">2b. Dorsal body scales in ≥ 29 (usually 30) longitudinal rows at midbody; a dark dorsal stripe present with sharply defined lateral edges............................................. 9</keyLead>
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<keyStep id="915748AAFFA1FFB1FFAB09B74C9A5FF9" pageId="28" pageNumber="29">
<paragraph id="2A1C500FFFA1FFB1FFAB09B74C9A5FD9" blockId="28.[123,1169,918,1005]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB09B74C9A5FD9" pageId="28" pageNumber="29">
3a. Third supralabial scale does not extend dorsally to levels of nostrils (
<figureCitation id="B2984C8AFFA1FFB1FC5709B74BE25FB9" box="[903,1001,918,941]" captionStart="FIGURE" captionStartId="28.[155,167,1587,1606]" captionTargetBox="[125,1172,1299,1576]" captionTargetId="figure-886@28.[120,1176,1299,1576]" captionTargetPageId="28" captionText="FIGURE 40. Lateral view of head of (A) Malayotyphlops denrorum (PNM 9813; a redrafting by EH of original image in Wynn et al. 2016), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016), (C) Malayotyphlops castanotus (CAS SUR-27940; a redrafting by EH of original image in Wynn et al. 1993). Postnasal scale (blue; or posterior part of nasal scale, if nasal scale incompletely sdivided), preocular scale (green), second supralabial scale (purplish-gray), third supralabial scale (reddish brown). Figs. A–C reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512762" httpUri="https://zenodo.org/record/11512762/files/figure.png" pageId="28" pageNumber="29">Fig. 40A</figureCitation>
)...................................................................
<taxonomicName id="EDA32B8CFFA1FFB1FCA609974C9A5FD9" ID-CoL="88L23" authorityName="Wynn, Diesmos &amp; Brown" authorityYear="2016" box="[886,1169,950,973]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="denrorum">
<emphasis id="18D78C1DFFA1FFB1FCA609974C9A5FD9" box="[886,1169,950,973]" italics="true" pageId="28" pageNumber="29">Malayotyphlops denrorum</emphasis>
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</paragraph>
<paragraph id="2A1C500FFFA1FFB1FFAB09F74C9A5FF9" blockId="28.[123,1169,918,1005]" box="[123,1169,982,1005]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB09F74C9A5FF9" box="[123,1169,982,1005]" pageId="28" pageNumber="29">
3b. Third supralabial scale extends dorsally to levels of nostrils (
<figureCitation id="B2984C8AFFA1FFB1FCE109F74B995FF9" box="[817,914,982,1005]" captionStart="FIGURE" captionStartId="28.[155,167,1587,1606]" captionTargetBox="[125,1172,1299,1576]" captionTargetId="figure-886@28.[120,1176,1299,1576]" captionTargetPageId="28" captionText="FIGURE 40. Lateral view of head of (A) Malayotyphlops denrorum (PNM 9813; a redrafting by EH of original image in Wynn et al. 2016), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016), (C) Malayotyphlops castanotus (CAS SUR-27940; a redrafting by EH of original image in Wynn et al. 1993). Postnasal scale (blue; or posterior part of nasal scale, if nasal scale incompletely sdivided), preocular scale (green), second supralabial scale (purplish-gray), third supralabial scale (reddish brown). Figs. A–C reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512762" httpUri="https://zenodo.org/record/11512762/files/figure.png" pageId="28" pageNumber="29">Fig. 40B</figureCitation>
)................. 4
</keyLead>
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<keyStep id="915748AAFFA1FFB1FFAB0E214C9A5863" pageId="28" pageNumber="29">
<paragraph id="2A1C500FFFA1FFB1FFAB0E214C9A5823" blockId="28.[123,1173,1024,1143]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB0E214C9A5823" pageId="28" pageNumber="29">
4a. Preocular scale overlaps second supralabial scale, and postnasal scale does not overlap third supralabial scale (
<figureCitation id="B2984C8AFFA1FFB1FE880E0149B25823" box="[344,441,1056,1079]" captionStart="FIGURE" captionStartId="28.[155,167,1587,1606]" captionTargetBox="[125,1172,1299,1576]" captionTargetId="figure-886@28.[120,1176,1299,1576]" captionTargetPageId="28" captionText="FIGURE 40. Lateral view of head of (A) Malayotyphlops denrorum (PNM 9813; a redrafting by EH of original image in Wynn et al. 2016), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016), (C) Malayotyphlops castanotus (CAS SUR-27940; a redrafting by EH of original image in Wynn et al. 1993). Postnasal scale (blue; or posterior part of nasal scale, if nasal scale incompletely sdivided), preocular scale (green), second supralabial scale (purplish-gray), third supralabial scale (reddish brown). Figs. A–C reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512762" httpUri="https://zenodo.org/record/11512762/files/figure.png" pageId="28" pageNumber="29">Fig. 40B</figureCitation>
)..................................................... 5
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA1FFB1FFAB0E614C9A5863" blockId="28.[123,1173,1024,1143]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB0E614C9A5863" pageId="28" pageNumber="29">
4b. Preocular scale does not overlap second supralabial scale and postnasal scale overlaps third supralabial scale (
<figureCitation id="B2984C8AFFA1FFB1FE880E4149B25863" box="[344,441,1120,1143]" captionStart="FIGURE" captionStartId="28.[155,167,1587,1606]" captionTargetBox="[125,1172,1299,1576]" captionTargetId="figure-886@28.[120,1176,1299,1576]" captionTargetPageId="28" captionText="FIGURE 40. Lateral view of head of (A) Malayotyphlops denrorum (PNM 9813; a redrafting by EH of original image in Wynn et al. 2016), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016), (C) Malayotyphlops castanotus (CAS SUR-27940; a redrafting by EH of original image in Wynn et al. 1993). Postnasal scale (blue; or posterior part of nasal scale, if nasal scale incompletely sdivided), preocular scale (green), second supralabial scale (purplish-gray), third supralabial scale (reddish brown). Figs. A–C reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512762" httpUri="https://zenodo.org/record/11512762/files/figure.png" pageId="28" pageNumber="29">Fig. 40C</figureCitation>
)..................................................... 6
</keyLead>
</paragraph>
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<paragraph id="2A1C500FFFA1FFB1FFAB0EAA4C9A58D6" blockId="28.[123,1174,1163,1282]" pageId="28" pageNumber="29">
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5a. Dorsal body scales in 26 longitudinal rows behind head; dorsal stripe ≥ 17 scale rows wide on anterior of body..........................................
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5b. Dorsal body scales in 28 longitudinal rows behind head; dorsal stripe 15 scale rows wide on anterior of body, narrowing posteriorly.......................
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<emphasis id="18D78C1DFFA1FFB1FCBF0ECA4C9A5916" box="[879,1169,1259,1282]" italics="true" pageId="28" pageNumber="29">Malayotyphlops luzonensis</emphasis>
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40. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFA1FFB1FE3E0C124ADB5A52" authorityName="Wynn, Diesmos &amp; Brown" authorityYear="2016" box="[494,720,1587,1606]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="denrorum">
<emphasis id="18D78C1DFFA1FFB1FE3E0C124ADB5A52" box="[494,720,1587,1606]" italics="true" pageId="28" pageNumber="29">Malayotyphlops denrorum</emphasis>
</taxonomicName>
(PNM 9813; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA1FFB1FFAB0C6C49185A74" author="Wynn" box="[123,275,1613,1632]" etAl="et al." firstAuthor="Wynn" pageId="28" pageNumber="29" pagination="157 - 168" refId="ref36443" refString="WYNN, ADDISON H., ARVIN C. DIESMOS, AND RAFE M. BROWN. 2016. Two new species of Malayotyphlops from the northern Philippines, with redescriptions of Malayotyphlops luzonensis (Taylor) and Malayotyphlops ruber (Boettger). Journal of Herpetology 50 (1): 157 - 168." type="journal article" year="2016">Wynn et al. 2016</bibRefCitation>
), (B)
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<emphasis id="18D78C1DFFA1FFB1FE950C6C4A255A74" box="[325,558,1613,1632]" italics="true" pageId="28" pageNumber="29">Malayotyphlops luzonensis</emphasis>
</taxonomicName>
(CM 2653; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA1FFB1FC240C6C4C875A74" author="Wynn" box="[1012,1164,1613,1632]" etAl="et al." firstAuthor="Wynn" pageId="28" pageNumber="29" pagination="157 - 168" refId="ref36443" refString="WYNN, ADDISON H., ARVIN C. DIESMOS, AND RAFE M. BROWN. 2016. Two new species of Malayotyphlops from the northern Philippines, with redescriptions of Malayotyphlops luzonensis (Taylor) and Malayotyphlops ruber (Boettger). Journal of Herpetology 50 (1): 157 - 168." type="journal article" year="2016">Wynn et al. 2016</bibRefCitation>
), (C)
<taxonomicName id="EDA32B8CFFA1FFB1FF4E0C4649815A6E" baseAuthorityName="Wynn &amp; Leviton" baseAuthorityYear="1993" box="[158,394,1639,1658]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="castanotus">
<emphasis id="18D78C1DFFA1FFB1FF4E0C4649815A6E" box="[158,394,1639,1658]" italics="true" pageId="28" pageNumber="29">Malayotyphlops castanotus</emphasis>
</taxonomicName>
(CAS SUR-27940; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA1FFB1FC4B0C464C3E5A6E" author="Wynn" box="[923,1077,1639,1658]" etAl="et al." firstAuthor="Wynn" pageId="28" pageNumber="29" pagination="34 - 45" refId="ref36388" refString="WYNN, ADDISON H., AND ALAN E. LEVITON. 1993. Two new species of blind snake, genus Typhlops (Reptilia: Squamata: Typhlopidae), from the Philippine Archipelago. Proceedings of the Biological Society of Washington 106 (1): 34 - 45, 3 figs." type="book chapter" year="1993">Wynn et al. 1993</bibRefCitation>
). Postnasal scale (blue; or posterior part of nasal scale, if nasal scale incompletely sdivided), preocular scale (green), second supralabial scale (purplish-gray), third supralabial scale (reddish brown). Figs. A–C reproduced and modified with permission of A. Wynn and Journal of Herpetology.
</paragraph>
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6a. Well-delineated, dark dorsal stripe 9–11 scale rows wide, not narrowing posteriorly..............................................................
<taxonomicName id="EDA32B8CFFA0FFB0FCBC0ADD4C9A5D07" ID-CoL="3XMDL" baseAuthorityName="Wynn &amp; Leviton" baseAuthorityYear="1993" box="[876,1169,252,275]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="castanotus">
<emphasis id="18D78C1DFFA0FFB0FCBC0ADD4C9A5D07" box="[876,1169,252,275]" italics="true" pageId="29" pageNumber="30">Malayotyphlops castanotus</emphasis>
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<keyLead id="9152F33AFFA0FFB0FFAB0B3D4C9A5D47" pageId="29" pageNumber="30">6b. Lateral scale rows of dorsal stripe lighter than (or not as completely pigmented as) medial rows; dorsal stripe narrows posteriorly................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA0FFB0FFAB0B464C9A5DCA" pageId="29" pageNumber="30">
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<keyLead id="9152F33AFFA0FFB0FFAB0B464C9A5D8A" pageId="29" pageNumber="30">
7a. Pale (unpigmented) collar present behind head; dorsal body scales in&gt; 400 transverse rows between head and tail.......................................
<taxonomicName id="EDA32B8CFFA0FFB0FC5C0BA64C9A5D8A" ID-CoL="3XMDM" baseAuthorityName="Wynn &amp; Leviton" baseAuthorityYear="1993" box="[908,1169,391,414]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="collaris">
<emphasis id="18D78C1DFFA0FFB0FC5C0BA64C9A5D8A" box="[908,1169,391,414]" italics="true" pageId="29" pageNumber="30">Malayotyphlops collaris</emphasis>
</taxonomicName>
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<paragraph id="2A1C500FFFA0FFB0FFAB0B864C9A5DCA" blockId="29.[123,1174,359,478]" pageId="29" pageNumber="30">
<keyLead id="9152F33AFFA0FFB0FFAB0B864C9A5DCA" pageId="29" pageNumber="30">7b. No pale collar behind head; dorsal body scales in &lt;400 transverse rows between head and tail........................................................................... 8</keyLead>
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<keyStep id="915748AAFFA0FFB0FFAB0BD34C9A5E3D" pageId="29" pageNumber="30">
<paragraph id="2A1C500FFFA0FFB0FFAB0BD34C9A5E1D" blockId="29.[123,1169,498,553]" box="[123,1169,498,521]" pageId="29" pageNumber="30">
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8a. Dark dorsal body stripe 15 scale rows wide on anterior of body......
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<emphasis id="18D78C1DFFA0FFB0FC730BD34C9A5E1D" box="[931,1169,498,521]" italics="true" pageId="29" pageNumber="30">Malayotyphlops ruber</emphasis>
</taxonomicName>
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<paragraph id="2A1C500FFFA0FFB0FFAB08334C9A5E3D" blockId="29.[123,1169,498,553]" box="[123,1169,530,553]" pageId="29" pageNumber="30">
<keyLead id="9152F33AFFA0FFB0FFAB08334C9A5E3D" box="[123,1169,530,553]" pageId="29" pageNumber="30">
8b. Dark dorsal body stripe 13 scale rows wide on anterior of body......
<taxonomicName id="EDA32B8CFFA0FFB0FC7308334C9A5E3D" ID-CoL="88L22" authorityName="Wynn, Diesmos &amp; Brown" authorityYear="2016" box="[931,1169,530,553]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="andyi">
<emphasis id="18D78C1DFFA0FFB0FC7308334C9A5E3D" box="[931,1169,530,553]" italics="true" pageId="29" pageNumber="30">Malayotyphlops andyi</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="915748AAFFA0FFB0FFAB081C4C9A5EA1" pageId="29" pageNumber="30">
<paragraph id="2A1C500FFFA0FFB0FFAB081C4C9A5E60" blockId="29.[123,1169,573,693]" pageId="29" pageNumber="30">
<keyLead id="9152F33AFFA0FFB0FFAB081C4C9A5E60" pageId="29" pageNumber="30">
9a. Dorsal body scales in&gt; 350 transverse rows between head and tail.........................................................................
<taxonomicName id="EDA32B8CFFA0FFB0FC83087C4C9A5E60" ID-CoL="3XMDK" baseAuthorityName="Taylor" baseAuthorityYear="1917" box="[851,1169,605,628]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="canlaonensis">
<emphasis id="18D78C1DFFA0FFB0FC83087C4C9A5E60" box="[851,1169,605,628]" italics="true" pageId="29" pageNumber="30">Malayotyphlops canlaonensis</emphasis>
</taxonomicName>
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<paragraph id="2A1C500FFFA0FFB0FFAB085F4C9A5EA1" blockId="29.[123,1169,573,693]" pageId="29" pageNumber="30">
<keyLead id="9152F33AFFA0FFB0FFAB085F4C9A5EA1" pageId="29" pageNumber="30">
9b. Dorsal body scales in &lt;350 transverse rows between head and tail...........................................................................
<taxonomicName id="EDA32B8CFFA0FFB0FCBC08BF4C9A5EA1" baseAuthorityName="Gray" baseAuthorityYear="1845" box="[876,1169,670,693]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="ruficaudus">
<emphasis id="18D78C1DFFA0FFB0FCBC08BF4C9A5EA1" box="[876,1169,670,693]" italics="true" pageId="29" pageNumber="30">Malayotyphlops ruficaudus</emphasis>
</taxonomicName>
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Represented in the
<collectingCountry id="52B4109FFFA1FFB1FE9A086249C95E4E" box="[330,450,579,602]" name="Philippines" pageId="28" pageNumber="29">Philippines</collectingCountry>
by a single species.................
<taxonomicName id="EDA32B8CFFA1FFB1FCAF08624C9A5E4E" box="[895,1169,579,602]" class="Reptilia" family="Pythonidae" genus="Malayopython" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="18D78C1DFFA1FFB1FCAF08624C9A5E4E" box="[895,1169,579,602]" italics="true" pageId="28" pageNumber="29">Malayopython reticulatus</emphasis>
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<mods:title id="2B94767A975A69BEC46086C8FD1F2E87">Illustrated Key to the Snakes of the Philippines</mods:title>
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<treatment id="A20AE119FFA2FF8DFDE50BEF4C9A5E07" ID-DOI="http://doi.org/10.5281/zenodo.12728199" ID-GBIF-Taxon="235040688" ID-Zenodo-Dep="12728199" LSID="urn:lsid:plazi:treatment:A20AE119FFA2FF8DFDE50BEF4C9A5E07" httpUri="http://treatment.plazi.org/id/A20AE119FFA2FF8DFDE50BEF4C9A5E07" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<taxonomicName id="EDA32B8CFFA2FFB2FDE50BEF4AD05DF1" ID-CoL="6432H" authority="Jan, 1862" authorityName="Jan" authorityYear="1862" box="[565,731,462,485]" class="Reptilia" family="Colubridae" genus="Pseudorabdion" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFA2FFB2FDE50BEF4AD05DF1" bold="true" box="[565,731,462,485]" italics="true" pageId="31" pageNumber="32">Pseudorabdion</emphasis>
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<paragraph id="2A1C500FFFA2FFB2FE2C0BCF4B1E5E11" blockId="31.[508,789,462,517]" box="[508,789,494,517]" pageId="31" pageNumber="32">Key to Philippine species.</paragraph>
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1a. Loreal scale absent, prefrontal scale in contact with supralabial scales (
<figureCitation id="B2984C8AFFA2FFB2FC4D08384C155E24" box="[925,1054,537,560]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43B–D</figureCitation>
)....... 2
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA2FFB2FFAB08184C9A5E44" blockId="31.[123,1169,537,592]" box="[123,1169,569,592]" pageId="31" pageNumber="32">
<keyLead id="9152F33AFFA2FFB2FFAB08184C9A5E44" box="[123,1169,569,592]" pageId="31" pageNumber="32">
1b. Loreal scale present, prefrontal scale not in contact with supralabial scales (
<figureCitation id="B2984C8AFFA2FFB2FC0108184C395E44" box="[977,1074,569,592]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43A</figureCitation>
)..... 4
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<paragraph id="2A1C500FFFA2FFB2FFAB08424C9A5E8E" blockId="31.[123,1174,611,698]" pageId="31" pageNumber="32">
<keyLead id="9152F33AFFA2FFB2FFAB08424C9A5E8E" pageId="31" pageNumber="32">
2b. Frontal scale does not border eye; supraocular and postocular scales present, fused to each other (
<figureCitation id="B2984C8AFFA2FFB2FF4F08A2492B5E8E" box="[159,288,643,666]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43C–D</figureCitation>
)................................................................ 3
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</paragraph>
<paragraph id="2A1C500FFFA2FFB2FFAB08824C9A5EAE" blockId="31.[123,1174,611,698]" box="[123,1169,675,698]" pageId="31" pageNumber="32">
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2a. Frontal scale borders eye; supraocular scale absent (
<figureCitation id="B2984C8AFFA2FFB2FD0208824B395EAE" box="[722,818,675,698]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43B</figureCitation>
)........................ 6
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<paragraph id="2A1C500FFFA2FFB2FFAB08EF4C9A5F71" blockId="31.[123,1174,718,1029]" pageId="31" pageNumber="32">
<keyLead id="9152F33AFFA2FFB2FFAB08EF4C9A5F71" pageId="31" pageNumber="32">
3a. Parietal scale in contact with fourth and fifth supralabial scales (
<figureCitation id="B2984C8AFFA2FFB2FC8E08EF4BCA5EF1" box="[862,961,718,741]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43C</figureCitation>
); each scale of first dorsal body scale row has dark center and light edges; each ventral body scale dark brown, lighter along posterior edge; supraocular and postocular scales usually fused to eye; ventral body scales 132–144 (males), 144–157 (females); subcaudal scales 22–24 (males), 16–17 (females)....................................................
<taxonomicName id="EDA32B8CFFA2FFB2FC8A096F4C9A5F71" ID-CoL="4P9CZ" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[858,1169,846,869]" class="Reptilia" family="Colubridae" genus="Pseudorabdion" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="oxycephalum">
<emphasis id="18D78C1DFFA2FFB2FC8A096F4C9A5F71" box="[858,1169,846,869]" italics="true" pageId="31" pageNumber="32">Pseudorabdion oxycephalum</emphasis>
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</paragraph>
<paragraph id="2A1C500FFFA2FFB2FFAB094F4C9A5811" blockId="31.[123,1174,718,1029]" pageId="31" pageNumber="32">
<keyLead id="9152F33AFFA2FFB2FFAB094F4C9A5811" pageId="31" pageNumber="32">
3b. Parietal scale in contact with fifth, but not fourth, supralabial scale (
<figureCitation id="B2984C8AFFA2FFB2FC5D094F4BF85F91" box="[909,1011,878,901]" captionStart="FIGURE" captionStartId="31.[155,167,1616,1635]" captionTargetBox="[293,1003,1156,1604]" captionTargetId="figure-599@31.[293,1003,1156,1604]" captionTargetPageId="31" captionText="FIGURE 43. Lateral view of head of (A) Pseudorabdion mcnamarae (KU 327199), (B) Pseudorabdion cf. collaris (KU 315197), (C) Pseudorabdion oxycephalum (KU 324641), (D) Pseudorabdion montanum (KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D)." figureDoi="http://doi.org/10.5281/zenodo.11512770" httpUri="https://zenodo.org/record/11512770/files/figure.png" pageId="31" pageNumber="32">Fig. 43D</figureCitation>
); each scale of first dorsal body scale row has light center and dark edges; each ventral body scale whitish, with dark brown pigment confined to a broad transverse band; supraocular and postocular scales not fused to eye; ventral body scales 146–148 (males), 154–161 (females); subcaudal scales 28 (males), 21–24 (females)..................................
<taxonomicName id="EDA32B8CFFA2FFB2FCA609CF4C9A5811" ID-CoL="4P9CY" authorityName="Leviton &amp; Brown" authorityYear="1959" box="[886,1169,1006,1029]" class="Reptilia" family="Colubridae" genus="Pseudorabdion" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="montanum">
<emphasis id="18D78C1DFFA2FFB2FCA609CF4C9A5811" box="[886,1169,1006,1029]" italics="true" pageId="31" pageNumber="32">Pseudorabdion montanum</emphasis>
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4a. Light nuchal collar usually present; subcaudal scales 26–29 (males), 17–23 (females)............................................................
<taxonomicName id="EDA32B8CFFA2FFB2FCB90E184C9A5844" ID-CoL="4P9CW" baseAuthorityName="Taylor" baseAuthorityYear="1917" box="[873,1169,1081,1104]" class="Reptilia" family="Colubridae" genus="Pseudorabdion" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="mcnamarae">
<emphasis id="18D78C1DFFA2FFB2FCB90E184C9A5844" box="[873,1169,1081,1104]" italics="true" pageId="31" pageNumber="32">Pseudorabdion mcnamarae</emphasis>
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<paragraph id="2A1C500FFFA2FFB2FFAB0E784C9A5864" blockId="31.[123,1169,1049,1136]" box="[123,1169,1113,1136]" pageId="31" pageNumber="32">
<keyLead id="9152F33AFFA2FFB2FFAB0E784C9A5864" box="[123,1169,1113,1136]" pageId="31" pageNumber="32">4b. Light nuchal collar absent; subcaudal scales&gt; 30 (males and females).................. 5</keyLead>
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43. Lateral view of head of (A)
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<emphasis id="18D78C1DFFA2FFB2FE210C714AD55A77" box="[497,734,1616,1635]" italics="true" pageId="31" pageNumber="32">Pseudorabdion mcnamarae</emphasis>
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(KU 327199), (B)
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cf.
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(KU 315197), (C)
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<emphasis id="18D78C1DFFA2FFB2FF3E0C4B49ED5A69" box="[238,486,1642,1661]" italics="true" pageId="31" pageNumber="32">Pseudorabdion oxycephalum</emphasis>
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(KU 324641), (D)
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<emphasis id="18D78C1DFFA2FFB2FD5A0C4B4B605A69" box="[650,875,1642,1661]" italics="true" pageId="31" pageNumber="32">Pseudorabdion montanum</emphasis>
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(KU 305063). Loreal scale (green; A), parietal scale (purplish-gray; C–D), fourth supralabial scale (light gray; C–D), postocular scale (blue; B), frontal scale (reddish brown; B–D), scale formed from fusion of supraocular and postocular scales (yellow; C–D).
</paragraph>
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5a. Dorsal body scales uniform pale brown, each thinly edged with pigmentless border; distal portion of hemipenes calyculate, subcaudal scales 33–35 (females), 40 (males)......................................................................
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5b. Dorsal body scales pale, centers and posterior tips nearly without pigment; anterior edge of each dorsal body scale with brown mottling; large areas of lateral portion of ventral body scales without pigment; distal portion of hemipenes minutely spinose; subcaudal scales 36–39 (males; unknown for females).....................................
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<emphasis id="18D78C1DFF9DFF8DFCAE0BBD4C9A5DA7" box="[894,1169,412,435]" italics="true" pageId="32" pageNumber="33">Pseudorabdion talonuran</emphasis>
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6a. Postocular scale absent (fused to ocular scale); maxillary teeth 10–12...
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<emphasis id="18D78C1DFF9DFF8DFC6C0B9D4C9A5DC7" box="[956,1169,444,467]" italics="true" pageId="32" pageNumber="33">Pseudorabdion ater</emphasis>
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6b. Postocular scale present, not fused to ocular scale (
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); maxillary teeth 22–25.............................................................
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cf.
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85
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Represented in the
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by a single species.............
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<emphasis id="18D78C1DFFA3FFB3FDEC09C14ADE5FE3" bold="true" box="[572,725,992,1015]" italics="true" pageId="30" pageNumber="31">Oxyrhabdium</emphasis>
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1a. Loreal scale separated from or in contact with second supralabial scale; if loreal scale in contact with second supralabial scale, their length of contact is less than half the length of the contact between loreal and postnasal scales (
<figureCitation id="B2984C8AFFA3FFB3FD6C0E4F4B2E5891" box="[700,805,1134,1157]" captionStart="FIGURE" captionStartId="30.[155,167,1682,1701]" captionTargetBox="[298,1002,1466,1671]" captionTargetId="figure-566@30.[293,1003,1466,1671]" captionTargetPageId="30" captionText="FIGURE 42. Lateral view of head of (A) Oxyrhabdium modestum (KU 311301), (B) Oxyrhabdium leporinum leporinum (KU 322335). Loreal scale (green), postnasal scale (blue), second supralabial scale (purplish-gray)." figureDoi="http://doi.org/10.5281/zenodo.11512768" httpUri="https://zenodo.org/record/11512768/files/figure.png" pageId="30" pageNumber="31">Fig. 42A</figureCitation>
); usually eight supralabial scales (rarely seven), with fifth and six (rarely fourth and fifth) supralabial scales bordering eye (
<figureCitation id="B2984C8AFFA3FFB3FF4F0E8F490E58D1" box="[159,261,1198,1221]" captionStart="FIGURE" captionStartId="30.[155,167,1682,1701]" captionTargetBox="[298,1002,1466,1671]" captionTargetId="figure-566@30.[293,1003,1466,1671]" captionTargetPageId="30" captionText="FIGURE 42. Lateral view of head of (A) Oxyrhabdium modestum (KU 311301), (B) Oxyrhabdium leporinum leporinum (KU 322335). Loreal scale (green), postnasal scale (blue), second supralabial scale (purplish-gray)." figureDoi="http://doi.org/10.5281/zenodo.11512768" httpUri="https://zenodo.org/record/11512768/files/figure.png" pageId="30" pageNumber="31">Fig. 42A</figureCitation>
); dorsal surfaces of head, body, and tail brown or reddish-brown in color; juveniles have distinct white nuchal collar, but lack light crossbands on other parts of the body...............................................................
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1b. Length of contact between loreal scale and second supralabial scale similar to length of contact between loreal scale and postnasal scale (
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); seven supralabial scales, with fourth and fifth supralabial scales bordering eye (
<figureCitation id="B2984C8AFFA3FFB3FDFB0F6F4A875971" box="[555,652,1358,1381]" captionStart="FIGURE" captionStartId="30.[155,167,1682,1701]" captionTargetBox="[298,1002,1466,1671]" captionTargetId="figure-566@30.[293,1003,1466,1671]" captionTargetPageId="30" captionText="FIGURE 42. Lateral view of head of (A) Oxyrhabdium modestum (KU 311301), (B) Oxyrhabdium leporinum leporinum (KU 322335). Loreal scale (green), postnasal scale (blue), second supralabial scale (purplish-gray)." figureDoi="http://doi.org/10.5281/zenodo.11512768" httpUri="https://zenodo.org/record/11512768/files/figure.png" pageId="30" pageNumber="31">Fig. 42B</figureCitation>
); dorsal surfaces of head, body, and tail dark gray or olive in color; juveniles have distinct white nuchal collar plus light crossbands on other parts of body and tail; pale crossbands may be faintly visible in adults....................... 2
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42. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFA3FFB3FE200CB34ACA5AB1" baseAuthorityName="Duméril" baseAuthorityYear="1853" box="[496,705,1682,1701]" class="Reptilia" family="Cyclocoridae" genus="Oxyrhabdium" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="modestum">
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(KU 311301), (B)
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(KU 322335). Loreal scale (green), postnasal scale (blue), second supralabial scale (purplish-gray).
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2a. Adult dorsal color pattern uniform (lacking pale crossbands); number of ventral body scales + subcaudal scales 192–230..........................
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<emphasis id="18D78C1DFFA2FFB2FCC50ADD4C9A5D07" box="[789,1169,252,275]" italics="true" pageId="31" pageNumber="32">Oxyrhabdium leporinum leporinum</emphasis>
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2b. Adults and juveniles with narrow white dorsal body crossbands and white nuchal collar; number of ventral body scales + subcaudal scales 221–235...
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<emphasis id="18D78C1DFFA2FFB2FCC10B1D4C9A5D47" box="[785,1169,316,339]" italics="true" pageId="31" pageNumber="32">Oxyrhabdium leporinum visayanum</emphasis>
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Represented by a single species...................................
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1b. Dorsal body scales in 21 longitudinal rows at 25
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<paragraph id="2A1C500FFFA5FFB5FF190ADD4C4C5D07" blockId="24.[201,1095,219,275]" box="[201,1095,252,275]" pageId="24" pageNumber="25">
Key to species reported from or likely to occur in marine waters in the
<collectingCountry id="52B4109FFFA5FFB5FC180ADD4C4F5D07" box="[968,1092,252,275]" name="Philippines" pageId="24" pageNumber="25">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
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1a. Longitudinal scale rows around thickest part of body&gt; 72..........
<taxonomicName id="EDA32B8CFFA5FFB5FC720B074C9A5D29" ID-CoL="8GB7F" baseAuthorityName="Laidlaw" baseAuthorityYear="1901" box="[930,1169,294,317]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="18D78C1DFFA5FFB5FC720B074C9A5D29" box="[930,1169,294,317]" italics="true" pageId="24" pageNumber="25">Hydrophis annandalei</emphasis>
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</keyLead>
</paragraph>
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<keyLead id="9152F33AFFA5FFB5FFAA0B674C9A5D49" box="[122,1169,326,349]" pageId="24" pageNumber="25">1b. Longitudinal scale rows around thickest part of body 25–70.......................... 2</keyLead>
</paragraph>
</keyStep>
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<paragraph id="2A1C500FFFA5FFB5FFAA0B504C9A5DBC" blockId="24.[122,1173,369,456]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0B504C9A5DBC" pageId="24" pageNumber="25">
2a. Rostral scale fragmented into four or five smaller scales (
<figureCitation id="B2984C8AFFA5FFB5FD2C0B504B405D9C" box="[764,843,369,392]" captionStart="FIGURE" captionStartId="24.[155,167,823,842]" captionTargetBox="[464,832,469,813]" captionTargetId="figure-672@24.[464,832,469,813]" captionTargetPageId="24" captionText="FIGURE 35. Rostral scale fragmented into four or five scales in Hydrophis anomalus (specimen number unknown; a redrafting by EH of original image in Rasmussen et al. 2011). Rostral scale (reddish brown). Fig. 35, modified original line drawing reproduced with permission of A. Rasmussen and Zootaxa and Magnolia Press." figureDoi="http://doi.org/10.5281/zenodo.11512752" httpUri="https://zenodo.org/record/11512752/files/figure.png" pageId="24" pageNumber="25">Fig. 35</figureCitation>
); dorsal head scales with thickened edges..................................................
<taxonomicName id="EDA32B8CFFA5FFB5FC620BB04C9A5DBC" ID-CoL="8J66H" baseAuthorityName="Schmidt" baseAuthorityYear="1852" box="[946,1169,401,424]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="anomalus">
<emphasis id="18D78C1DFFA5FFB5FC620BB04C9A5DBC" box="[946,1169,401,424]" italics="true" pageId="24" pageNumber="25">Hydrophis anomalus</emphasis>
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</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA5FFB5FFAA0B904C9A5DDC" blockId="24.[122,1173,369,456]" box="[122,1169,433,456]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0B904C9A5DDC" box="[122,1169,433,456]" pageId="24" pageNumber="25">2b. Rostral scale not fragmented into four or five smaller scales.......................... 3</keyLead>
</paragraph>
</keyStep>
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35. Rostral scale fragmented into four or five scales in
<taxonomicName id="EDA32B8CFFA5FFB5FD1B09164B745F5E" baseAuthorityName="Schmidt" baseAuthorityYear="1852" box="[715,895,823,842]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="anomalus">
<emphasis id="18D78C1DFFA5FFB5FD1B09164B745F5E" box="[715,895,823,842]" italics="true" pageId="24" pageNumber="25">Hydrophis anomalus</emphasis>
</taxonomicName>
(specimen number unknown; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA5FFB5FE6E09714A745F77" author="Rasmussen" box="[446,639,848,867]" etAl="et al." firstAuthor="Rasmussen" pageId="24" pageNumber="25" pagination="1 - 20" refId="ref35158" refString="RASMUSSEN, ARNE R., JOHAN ELMBERG, PETER GRAVLUND, AND IVAN INEICH. 2011. Sea snakes (Serpentes subfamilies Hydrophiinae and Laticaudinae) in Vietnam: a comprehensive checklist and an updated identification key. Zootaxa 2894: 1 - 20." type="journal article" year="2011">Rasmussen et al. 2011</bibRefCitation>
). Rostral scale (reddish brown). Fig. 35, modified original line drawing reproduced with permission of A. Rasmussen and
<emphasis id="18D78C1DFFA5FFB5FDA7094B4AB65F69" box="[631,701,874,893]" italics="true" pageId="24" pageNumber="25">Zootaxa</emphasis>
and Magnolia Press.
</paragraph>
</caption>
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<paragraph id="2A1C500FFFA5FFB5FFAA09BA4C9A5FC6" blockId="24.[122,1174,923,1010]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA09BA4C9A5FC6" pageId="24" pageNumber="25">
3a. Ventral body scales large (&gt; 1/4 body width) on anterior part of body and small on posterior part of body..................................................
<taxonomicName id="EDA32B8CFFA5FFB5FC67099A4C9A5FC6" ID-CoL="3N8CM" baseAuthorityName="Schmidt" baseAuthorityYear="1852" box="[951,1169,955,978]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="viperinus">
<emphasis id="18D78C1DFFA5FFB5FC67099A4C9A5FC6" box="[951,1169,955,978]" italics="true" pageId="24" pageNumber="25">Hydrophis viperinus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA5FFB5FFAA09FA4C9A5FE6" blockId="24.[122,1174,923,1010]" box="[122,1169,987,1010]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA09FA4C9A5FE6" box="[122,1169,987,1010]" pageId="24" pageNumber="25">3b. Ventral body scales small on both anterior and posterior parts of body.................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA5FFB5FFAA0E274C9A5869" pageId="24" pageNumber="25">
<paragraph id="2A1C500FFFA5FFB5FFAA0E274C9A5829" blockId="24.[122,1173,1030,1149]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0E274C9A5829" pageId="24" pageNumber="25">
4a. Ventral surface of body uniform yellow or white; dorsal part of body uniform dark brown or black........................................................
<taxonomicName id="EDA32B8CFFA5FFB5FC110E074C9A5829" ID-CoL="3N8BL" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[961,1169,1062,1085]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="platurus">
<emphasis id="18D78C1DFFA5FFB5FC110E074C9A5829" box="[961,1169,1062,1085]" italics="true" pageId="24" pageNumber="25">Hydrophis platurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA5FFB5FFAA0E674C9A5869" blockId="24.[122,1173,1030,1149]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0E674C9A5869" pageId="24" pageNumber="25">4b. Ventral surface of body not uniform yellow or white; dorsal part of body not uniform dark brown or black............................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA5FFB5FFAA0EB14C9A58D3" pageId="24" pageNumber="25">
<paragraph id="2A1C500FFFA5FFB5FFAA0EB14C9A58B3" blockId="24.[122,1169,1168,1223]" box="[122,1169,1168,1191]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0EB14C9A58B3" box="[122,1169,1168,1191]" pageId="24" pageNumber="25">
5a. Mental scale elongate, not triangular (
<figureCitation id="B2984C8AFFA5FFB5FDEA0EB14A9058B3" box="[570,667,1168,1191]" captionStart="FIGURE" captionStartId="24.[155,167,1628,1647]" captionTargetBox="[299,997,1352,1614]" captionTargetId="figure-676@24.[299,997,1352,1614]" captionTargetPageId="24" captionText="FIGURE 36. Ventral view of head of (A) Hydrophis schistosus (specimen number unknown; a redrafting by EH of orig- inal image in Rasmussen et al. 2011), (B) Hydrophis curtus (KU 40063). Mental scale (green), and either elongate (A) or triangular (B). Fig. 36A modified original line drawing reproduced with permission of A. Rasmussen and Zootaxa and Magnolia Press." figureDoi="http://doi.org/10.5281/zenodo.11512754" httpUri="https://zenodo.org/record/11512754/files/figure.png" pageId="24" pageNumber="25">Fig. 36A</figureCitation>
)...................
<taxonomicName id="EDA32B8CFFA5FFB5FC7E0EB14C9A58B3" ID-CoL="3N8BV" box="[942,1169,1168,1191]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schistosus">
<emphasis id="18D78C1DFFA5FFB5FC7E0EB14C9A58B3" box="[942,1169,1168,1191]" italics="true" pageId="24" pageNumber="25">Hydrophis schistosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA5FFB5FFAA0E914C9A58D3" blockId="24.[122,1169,1168,1223]" box="[122,1169,1200,1223]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0E914C9A58D3" box="[122,1169,1200,1223]" pageId="24" pageNumber="25">
5b. Mental scale triangular, not elongate (
<figureCitation id="B2984C8AFFA5FFB5FDEA0E914A9158D3" box="[570,666,1200,1223]" captionStart="FIGURE" captionStartId="24.[155,167,1628,1647]" captionTargetBox="[299,997,1352,1614]" captionTargetId="figure-676@24.[299,997,1352,1614]" captionTargetPageId="24" captionText="FIGURE 36. Ventral view of head of (A) Hydrophis schistosus (specimen number unknown; a redrafting by EH of orig- inal image in Rasmussen et al. 2011), (B) Hydrophis curtus (KU 40063). Mental scale (green), and either elongate (A) or triangular (B). Fig. 36A modified original line drawing reproduced with permission of A. Rasmussen and Zootaxa and Magnolia Press." figureDoi="http://doi.org/10.5281/zenodo.11512754" httpUri="https://zenodo.org/record/11512754/files/figure.png" pageId="24" pageNumber="25">Fig. 36B</figureCitation>
).................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA5FFB5FFAA0EFA4C9A5906" pageId="24" pageNumber="25">
<paragraph id="2A1C500FFFA5FFB5FFAA0EFA4C9A58E6" blockId="24.[122,1169,1243,1298]" box="[122,1169,1243,1266]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0EFA4C9A58E6" box="[122,1169,1243,1266]" pageId="24" pageNumber="25">6a. Enlarged chin shields absent................................................... 7</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA5FFB5FFAA0EDA4C9A5906" blockId="24.[122,1169,1243,1298]" box="[122,1169,1275,1298]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0EDA4C9A5906" box="[122,1169,1275,1298]" pageId="24" pageNumber="25">6b. Enlarged chin shields present, one or two pairs.................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA5FFB4FFAA0F074C9A5CE7" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<paragraph id="2A1C500FFFA5FFB5FFAA0F074C9A5929" blockId="24.[122,1169,1318,1341]" box="[122,1169,1318,1341]" pageId="24" pageNumber="25">
<keyLead id="9152F33AFFA5FFB5FFAA0F074C9A5929" box="[122,1169,1318,1341]" pageId="24" pageNumber="25">
7a. Midbody scales in&gt; 45 longitudinal rows around body................
<taxonomicName id="EDA32B8CFFA5FFB5FC1A0F074C9A5929" ID-CoL="3N8C5" baseAuthorityName="Gray" baseAuthorityYear="1846" box="[970,1169,1318,1341]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="stokesii">
<emphasis id="18D78C1DFFA5FFB5FC1A0F074C9A5929" box="[970,1169,1318,1341]" italics="true" pageId="24" pageNumber="25">Hydrophis stokesii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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36. Ventral view of head of (A)
<taxonomicName id="EDA32B8CFFA5FFB5FE210C7D4AA35A7B" box="[497,680,1628,1647]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schistosus">
<emphasis id="18D78C1DFFA5FFB5FE210C7D4AA35A7B" box="[497,680,1628,1647]" italics="true" pageId="24" pageNumber="25">Hydrophis schistosus</emphasis>
</taxonomicName>
(specimen number unknown; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA5FFB5FF220C5749BD5A9D" author="Rasmussen" box="[242,438,1654,1673]" etAl="et al." firstAuthor="Rasmussen" pageId="24" pageNumber="25" pagination="1 - 20" refId="ref35158" refString="RASMUSSEN, ARNE R., JOHAN ELMBERG, PETER GRAVLUND, AND IVAN INEICH. 2011. Sea snakes (Serpentes subfamilies Hydrophiinae and Laticaudinae) in Vietnam: a comprehensive checklist and an updated identification key. Zootaxa 2894: 1 - 20." type="journal article" year="2011">Rasmussen et al. 2011</bibRefCitation>
), (B)
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<emphasis id="18D78C1DFFA5FFB5FE380C574A765A9D" box="[488,637,1654,1673]" italics="true" pageId="24" pageNumber="25">Hydrophis curtus</emphasis>
</taxonomicName>
(KU 40063). Mental scale (green), and either elongate (A) or triangular (B). Fig. 36A modified original line drawing reproduced with permission of A. Rasmussen and
<emphasis id="18D78C1DFFA5FFB5FBFB0CAE4C7A5AB6" box="[1067,1137,1679,1698]" italics="true" pageId="24" pageNumber="25">Zootaxa</emphasis>
and Magnolia Press.
</paragraph>
</caption>
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<keyLead id="9152F33AFFA4FFB4FFAB0AFD4C9A5CE7" box="[123,1169,220,243]" pageId="25" pageNumber="26">
7b. Midbody scales in &lt;45 longitudinal rows around body.................
<taxonomicName id="EDA32B8CFFA4FFB4FC070AFD4C9A5CE7" ID-CoL="3N88Y" baseAuthorityName="Shaw" baseAuthorityYear="1802" box="[983,1169,220,243]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="curtus">
<emphasis id="18D78C1DFFA4FFB4FC070AFD4C9A5CE7" box="[983,1169,220,243]" italics="true" pageId="25" pageNumber="26">Hydrophis curtus</emphasis>
</taxonomicName>
</keyLead>
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</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB0B274C9A5D29" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB0B274C9A5D09" blockId="25.[123,1169,262,317]" box="[123,1169,262,285]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0B274C9A5D09" box="[123,1169,262,285]" pageId="25" pageNumber="26">
8a. Longitudinal scale rows around thickest part of body ≤ 2...............
<taxonomicName id="EDA32B8CFFA4FFB4FC1E0B274C9A5D09" baseAuthorityName="Gray" baseAuthorityYear="1849" box="[974,1169,262,285]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="jerdoni">
<emphasis id="18D78C1DFFA4FFB4FC1E0B274C9A5D09" box="[974,1169,262,285]" italics="true" pageId="25" pageNumber="26">Hydrophis jerdoni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0B074C9A5D29" blockId="25.[123,1169,262,317]" box="[123,1169,294,317]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0B074C9A5D29" box="[123,1169,294,317]" pageId="25" pageNumber="26">8b. Longitudinal scale rows around thickest part of body ≥ 25........................... 9</keyLead>
</paragraph>
</keyStep>
<paragraph id="2A1C500FFFA4FFB4FFAB0B704C9A5D7C" blockId="25.[123,1169,337,392]" box="[123,1169,337,360]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0B704C9A5D7C" box="[123,1169,337,360]" pageId="25" pageNumber="26">
<keyStep id="915748AAFFA4FFB4FFAB0B704C9A5D7C" box="[123,1169,337,360]" pageId="25" pageNumber="26">
9a. Frontal scale and parietal scales more or less fragmented (
<figureCitation id="B2984C8AFFA4FFB4FCC30B704B7F5D7C" box="[787,884,337,360]" captionStart="FIGURE" captionStartId="25.[155,167,618,637]" captionTargetBox="[293,1003,406,610]" captionTargetId="figure-1094@25.[293,1003,406,610]" captionTargetPageId="25" captionText="FIGURE 37. Dorsal view of head of (A) Hydrophis peronii (CAS 136104; a redrafting by EH of original image in Leviton et al. 2014), (B) Hydrophis melanocephalus (KU 94556). Frontal scales (green), parietal scales (purplish-gray). Fig. A reproduced and modified with permission of A. Leviton and the Calfornia Academy of Sciences." figureDoi="http://doi.org/10.5281/zenodo.11512756" httpUri="https://zenodo.org/record/11512756/files/figure.png" pageId="25" pageNumber="26">Fig. 37A</figureCitation>
)......
<taxonomicName id="EDA32B8CFFA4FFB4FC1E0B704C9A5D7C" ID-CoL="3N8BJ" baseAuthorityName="Dumeril" baseAuthorityYear="1853" box="[974,1169,337,360]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="peronii">
<emphasis id="18D78C1DFFA4FFB4FC1E0B704C9A5D7C" box="[974,1169,337,360]" italics="true" pageId="25" pageNumber="26">Hydrophis peronii</emphasis>
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</keyStep>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0B504C9A5EFC" blockId="25.[123,1169,337,392]" lastBlockId="25.[123,1169,721,776]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0B504C9A5EFC" pageId="25" pageNumber="26">
9b. Frontal scale and parietal scales unfragmented (
<figureCitation id="B2984C8AFFA4FFB4FD740B504B0F5D9C" box="[676,772,369,392]" captionStart="FIGURE" captionStartId="25.[155,167,618,637]" captionTargetBox="[293,1003,406,610]" captionTargetId="figure-1094@25.[293,1003,406,610]" captionTargetPageId="25" captionText="FIGURE 37. Dorsal view of head of (A) Hydrophis peronii (CAS 136104; a redrafting by EH of original image in Leviton et al. 2014), (B) Hydrophis melanocephalus (KU 94556). Frontal scales (green), parietal scales (purplish-gray). Fig. A reproduced and modified with permission of A. Leviton and the Calfornia Academy of Sciences." figureDoi="http://doi.org/10.5281/zenodo.11512756" httpUri="https://zenodo.org/record/11512756/files/figure.png" pageId="25" pageNumber="26">Fig. 37B</figureCitation>
)........................... 10
<keyStep id="915748AAFFA4FFB4FFAB08F04C9A5EFC" box="[123,1169,721,744]" pageId="25" pageNumber="26">10a. Maxillary teeth behind fangs ≥ 9.............................................. 11</keyStep>
</keyLead>
</paragraph>
<caption id="7EDC0087FFA4FFB4FF4B084B4BB05EA4" ID-DOI="http://doi.org/10.5281/zenodo.11512756" ID-Zenodo-Dep="11512756" httpUri="https://zenodo.org/record/11512756/files/figure.png" pageId="25" pageNumber="26" startId="25.[155,167,618,637]" targetBox="[293,1003,406,610]" targetPageId="25" targetType="figure">
<paragraph id="2A1C500FFFA4FFB4FF4B084B4BB05EA4" blockId="25.[122,1174,618,688]" pageId="25" pageNumber="26">
<smallCapsWord id="2CFAC6D3FFA4FFB4FF4B084B48D35E68" baselines="632,632" box="[155,216,618,637]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="25" pageNumber="26">FIGURE</smallCapsWord>
37. Dorsal view of head of (A)
<taxonomicName id="EDA32B8CFFA4FFB4FE3D084B4A825E69" baseAuthorityName="Dumeril" baseAuthorityYear="1853" box="[493,649,618,637]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="peronii">
<emphasis id="18D78C1DFFA4FFB4FE3D084B4A825E69" box="[493,649,618,637]" italics="true" pageId="25" pageNumber="26">Hydrophis peronii</emphasis>
</taxonomicName>
(CAS 136104; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFA4FFB4FBB7084B48F15E83" author="Leviton" etAl="et al." firstAuthor="Leviton" pageId="25" pageNumber="26" refId="ref34267" refString="LEVITON, ALAN E., RAFE M. BROWN, AND CAMERON D. SILER. 2014. The dangerously venomous snakes of the Philippine Archipelago. Pages 473 - 530 in G. C. Williams and T. M. Gosliner, eds., The Coral Triangle: The 2011 Hearst Biodiversity Philippine Expedition. California Academy of Sciences, San Francisco, California, USA." type="book" year="2014">Leviton et al. 2014</bibRefCitation>
), (B)
<taxonomicName id="EDA32B8CFFA4FFB4FEFB08A54A1D5E83" authorityName="Gray" authorityYear="1849" box="[299,534,644,663]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="18D78C1DFFA4FFB4FEFB08A54A1D5E83" box="[299,534,644,663]" italics="true" pageId="25" pageNumber="26">Hydrophis melanocephalus</emphasis>
</taxonomicName>
(KU 94556). Frontal scales (green), parietal scales (purplish-gray). Fig. A reproduced and modified with permission of A. Leviton and the Calfornia Academy of Sciences.
</paragraph>
</caption>
<paragraph id="2A1C500FFFA4FFB4FFAB08D04C9A5F1C" blockId="25.[123,1169,721,776]" box="[123,1169,753,776]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB08D04C9A5F1C" box="[123,1169,753,776]" pageId="25" pageNumber="26">10b. Maxillary teeth behind fangs ≤ 8.............................................. 13</keyLead>
</paragraph>
<keyStep id="915748AAFFA4FFB4FFAB093A4C9A5F86" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB093A4C9A5F46" blockId="25.[123,1174,795,914]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB093A4C9A5F46" pageId="25" pageNumber="26">
11a. Neck width divided by width at widest part of body 0.45–0.56; maxillary teeth behind fangs 14–18....................................................
<taxonomicName id="EDA32B8CFFA4FFB4FC40091A4C9A5F46" ID-CoL="3N88M" baseAuthorityName="Shaw" baseAuthorityYear="1802" box="[912,1169,827,850]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="caerulescens">
<emphasis id="18D78C1DFFA4FFB4FC40091A4C9A5F46" box="[912,1169,827,850]" italics="true" pageId="25" pageNumber="26">Hydrophis caerulescens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB097A4C9A5F86" blockId="25.[123,1174,795,914]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB097A4C9A5F86" pageId="25" pageNumber="26">11b. Neck width divided by width at widest part of body ≥ 0.57; maxillary teeth behind fangs 9–13...................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB09874C9A5809" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB09874C9A5FC9" blockId="25.[123,1169,934,1053]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB09874C9A5FC9" pageId="25" pageNumber="26">
12a. Interspaces between black transverse dorsal bands thin, less than two scale rows wide.....................................................
<taxonomicName id="EDA32B8CFFA4FFB4FCCC09E74BEF5FC9" ID-CoL="3N8B8" authorityName="Smith" authorityYear="1926" box="[796,996,966,989]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="18D78C1DFFA4FFB4FCCC09E74BEF5FC9" box="[796,996,966,989]" italics="true" pageId="25" pageNumber="26">Hydrophis ornatus</emphasis>
</taxonomicName>
or
<taxonomicName id="EDA32B8CFFA4FFB4FBDA09E74C9A5FC9" ID-CoL="3N89X" baseAuthorityName="Gray" baseAuthorityYear="1849" box="[1034,1169,966,989]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="18D78C1DFFA4FFB4FBDA09E74C9A5FC9" box="[1034,1169,966,989]" italics="true" pageId="25" pageNumber="26">H. inornatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB09C74C9A5809" blockId="25.[123,1169,934,1053]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB09C74C9A5809" pageId="25" pageNumber="26">
12b. Interspaces between black transverse dorsal bands broad, more than two scale rows wide..............................................................
<taxonomicName id="EDA32B8CFFA4FFB4FC100E274C9A5809" ID-CoL="6MJRM" authorityName="Smith" authorityYear="1917" box="[960,1169,1030,1053]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="lamberti">
<emphasis id="18D78C1DFFA4FFB4FC100E274C9A5809" box="[960,1169,1030,1053]" italics="true" pageId="25" pageNumber="26">Hydrophis lamberti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB0E114C9A5893" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB0E114C9A5853" blockId="25.[123,1173,1072,1159]" box="[123,1169,1072,1095]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0E114C9A5853" box="[123,1169,1072,1095]" pageId="25" pageNumber="26">13a. Width of neck&gt; 1/2 width of widest part of body; anterior temporal scale one......... 14</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0E714C9A5893" blockId="25.[123,1173,1072,1159]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0E714C9A5893" pageId="25" pageNumber="26">13b. One or both of the following true: (1) width of neck &lt;1/2 width of widest part of body; (2) anterior temporal scales two................................................ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB0EBA4C9A58C6" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB0EBA4C9A58A6" blockId="25.[123,1169,1179,1234]" box="[123,1169,1179,1202]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0EBA4C9A58A6" box="[123,1169,1179,1202]" pageId="25" pageNumber="26">
14a. Longitudinal scale rows around midbody 8–18 more than around neck..
<taxonomicName id="EDA32B8CFFA4FFB4FC130EBA4C9A58A6" ID-CoL="3N88C" baseAuthorityName="Gray" baseAuthorityYear="1849" box="[963,1169,1179,1202]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="belcheri">
<emphasis id="18D78C1DFFA4FFB4FC130EBA4C9A58A6" box="[963,1169,1179,1202]" italics="true" pageId="25" pageNumber="26">Hydrophis belcheri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0E9A4C9A58C6" blockId="25.[123,1169,1179,1234]" box="[123,1169,1211,1234]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0E9A4C9A58C6" box="[123,1169,1211,1234]" pageId="25" pageNumber="26">
14b. Longitudinal scale rows around midbody 4–8 more than around neck...
<taxonomicName id="EDA32B8CFFA4FFB4FC1A0E9A4C9A58C6" ID-CoL="3N8C3" baseAuthorityName="Shaw" baseAuthorityYear="1802" box="[970,1169,1211,1234]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="spiralis">
<emphasis id="18D78C1DFFA4FFB4FC1A0E9A4C9A58C6" box="[970,1169,1211,1234]" italics="true" pageId="25" pageNumber="26">Hydrophis spiralis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB0EC74C9A59C9" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB0EC74C9A5949" blockId="25.[123,1183,1254,1501]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0EC74C9A5949" pageId="25" pageNumber="26">
15a. Head very small, width of neck ≤ 1/3 width of widest part of body; longitudinal scale rows around neck 17–23; ventral body scales of anterior part of body wider than adjacent scale rows; ventral body scales of posterior part of body are divided by a long longitudinal furrow...................................................................
<taxonomicName id="EDA32B8CFFA4FFB4FC190F674C9A5949" ID-CoL="3N89L" baseAuthorityName="Shaw" baseAuthorityYear="1802" box="[969,1169,1350,1373]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="18D78C1DFFA4FFB4FC190F674C9A5949" box="[969,1169,1350,1373]" italics="true" pageId="25" pageNumber="26">Hydrophis gracilis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0F474C9A59C9" blockId="25.[123,1183,1254,1501]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0F474C9A59C9" pageId="25" pageNumber="26">15b. One or more of the following combinations of characters true: (1) head small or moderate in size, width of neck&gt; 1/3 width of widest part of body; (2) longitudinal scale rows around neck ≥ 23; (3) ventral body scales of posterior part of body not divided by a long longitudinal furrow.......................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA4FFB4FFAB0FD14C9A5A86" pageId="25" pageNumber="26">
<paragraph id="2A1C500FFFA4FFB4FFAB0FD14C9A5A33" blockId="25.[123,1173,1520,1575]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0FD14C9A5A33" pageId="25" pageNumber="26">
16a. Longitudinal scale rows around midbody 29–35; ventral body scales 278–325; dorsal surface of head uniformly dark brown to black.............................
<taxonomicName id="EDA32B8CFFA4FFB4FC190C314C9A5A33" ID-CoL="3N88S" baseAuthorityName="Kharin" baseAuthorityYear="1984" box="[969,1169,1552,1575]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="coggeri">
<emphasis id="18D78C1DFFA4FFB4FC190C314C9A5A33" box="[969,1169,1552,1575]" italics="true" pageId="25" pageNumber="26">Hydrophis coggeri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA4FFB4FFAB0C1A4C9A5A86" blockId="25.[123,1174,1595,1682]" pageId="25" pageNumber="26">
<keyLead id="9152F33AFFA4FFB4FFAB0C1A4C9A5A86" pageId="25" pageNumber="26">16b. One or more of the following combinations of characters true: (1) longitudinal scale rows around midbody&gt; 35; (2) ventral body scales&gt; 325; (3) dorsal surface of head has yellow spot behind nostrils and yellow streak behind eye...................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA7FFB7FFAB0AFD4C9A5D07" pageId="26" pageNumber="27">
<paragraph id="2A1C500FFFA7FFB7FFAB0AFD4C9A5CE7" blockId="26.[123,1169,220,275]" box="[123,1169,220,243]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0AFD4C9A5CE7" box="[123,1169,220,243]" pageId="26" pageNumber="27">17a. Anterior temporal scales one................................................. 18</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA7FFB7FFAB0ADD4C9A5D07" blockId="26.[123,1169,220,275]" box="[123,1169,252,275]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0ADD4C9A5D07" box="[123,1169,252,275]" pageId="26" pageNumber="27">17b. Anterior temporal scales two................................................. 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA7FFB7FFAB0B074C9A5D9C" pageId="26" pageNumber="27">
<paragraph id="2A1C500FFFA7FFB7FFAB0B074C9A5D29" blockId="26.[123,1169,294,317]" box="[123,1169,294,317]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0B074C9A5D29" box="[123,1169,294,317]" pageId="26" pageNumber="27">
18a. Ventral body scales &lt;360; maxillary teeth behind fangs 6–8....
<taxonomicName id="EDA32B8CFFA7FFB7FCBD0B074C9A5D29" ID-CoL="6MK3B" authorityName="Gray" authorityYear="1849" box="[877,1169,294,317]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="18D78C1DFFA7FFB7FCBD0B074C9A5D29" box="[877,1169,294,317]" italics="true" pageId="26" pageNumber="27">Hydrophis melanocephalus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA7FFB7FFAB0B704C9A5D9C" blockId="26.[123,1174,337,392]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0B704C9A5D9C" pageId="26" pageNumber="27">18b. One or both of the following true: (1) ventral body scales ≥ 360; (2) maxillary teeth behind fangs &lt;6.................................................................. 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA7FFB7FFAB0BBA4C9A5E06" pageId="26" pageNumber="27">
<paragraph id="2A1C500FFFA7FFB7FFAB0BBA4C9A5DC6" blockId="26.[123,1173,411,530]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0BBA4C9A5DC6" pageId="26" pageNumber="27">
19a. Longitudinal scale rows around midbody 37–45; dorsal body scales on thickest part of body hexagonal or quadrangular........................................
<taxonomicName id="EDA32B8CFFA7FFB7FC1E0B9A4C9A5DC6" ID-CoL="3N88J" authorityName="Gunther" authorityYear="1872" box="[974,1169,443,466]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="brookii">
<emphasis id="18D78C1DFFA7FFB7FC1E0B9A4C9A5DC6" box="[974,1169,443,466]" italics="true" pageId="26" pageNumber="27">Hydrophis brookii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA7FFB7FFAB0BFA4C9A5E06" blockId="26.[123,1173,411,530]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB0BFA4C9A5E06" pageId="26" pageNumber="27">
19b. Longitudinal scale rows around midbody 31–39; dorsal body scales on thickest part of body with rounded or bluntly pointed tips.................................
<taxonomicName id="EDA32B8CFFA7FFB7FC0D0BDA4C9A5E06" ID-CoL="6MJRK" authorityName="Boulenger" authorityYear="1912" box="[989,1169,507,530]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="klossi">
<emphasis id="18D78C1DFFA7FFB7FC0D0BDA4C9A5E06" box="[989,1169,507,530]" italics="true" pageId="26" pageNumber="27">Hydrophis klossi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA7FFB7FFAB08074C9A5EC9" pageId="26" pageNumber="27">
<paragraph id="2A1C500FFFA7FFB7FFAB08074C9A5E69" blockId="26.[123,1174,550,733]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB08074C9A5E69" pageId="26" pageNumber="27">
20a. Width of neck ≤ 1/2 width of widest part of body; longitudinal scale rows around midbody 12–20 more than around neck; anterior surface of body dark with pale oval spots laterally that sometimes connect as crossbands..................................
<taxonomicName id="EDA32B8CFFA7FFB7FC1408474C9A5E69" ID-CoL="6MK3R" authorityName="Gunther" authorityYear="1864" box="[964,1169,614,637]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="atriceps">
<emphasis id="18D78C1DFFA7FFB7FC1408474C9A5E69" box="[964,1169,614,637]" italics="true" pageId="26" pageNumber="27">Hydrophis atriceps</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA7FFB7FFAB08A74C9A5EC9" blockId="26.[123,1174,550,733]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB08A74C9A5EC9" pageId="26" pageNumber="27">20b. One or both of the following true: (1) width of neck&gt; 1/2 width of widest part of body; (2) longitudinal scale rows around midbody &lt;12 more than around neck; (3) anterior and posterior surface of body dark with distinct white annuli............................ 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA7FFB7FFAB08D14C9A5F7C" pageId="26" pageNumber="27">
<paragraph id="2A1C500FFFA7FFB7FFAB08D14C9A5F3C" blockId="26.[123,1174,752,872]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB08D14C9A5F3C" pageId="26" pageNumber="27">
21a. Head yellowish or olive (adults) or black (juveniles); ventral body scales with a black longitudinal stripe that may fade with age; ventral body scales 290–390...
<taxonomicName id="EDA32B8CFFA7FFB7FC5F09304C9A5F3C" ID-CoL="3N88Z" box="[911,1169,785,808]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="cyanocinctus">
<emphasis id="18D78C1DFFA7FFB7FC5F09304C9A5F3C" box="[911,1169,785,808]" italics="true" pageId="26" pageNumber="27">Hydrophis cyanocinctus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA7FFB7FFAB09114C9A5F7C" blockId="26.[123,1174,752,872]" pageId="26" pageNumber="27">
<keyLead id="9152F33AFFA7FFB7FFAB09114C9A5F7C" pageId="26" pageNumber="27">
21b. Head black; ventral body scales usually black; ventral body scales 314–356; [non-marine] endemic to Lake Taal (Luzon)....................................
<taxonomicName id="EDA32B8CFFA7FFB7FC1609704C9A5F7C" ID-CoL="3N8BZ" authorityName="Garman" authorityYear="1881" box="[966,1169,849,872]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="semperi">
<emphasis id="18D78C1DFFA7FFB7FC1609704C9A5F7C" box="[966,1169,849,872]" italics="true" pageId="26" pageNumber="27">Hydrophis semperi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

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<document id="EC367E9B1747A7B2DA0686682B919FC8" ID-CLB-Dataset="299708" ID-DOI="10.5281/zenodo.11512680" ID-GBIF-Dataset="ccd739a4-10c3-4700-b27e-459bd82f1676" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512680" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1717720566437" checkinUser="felipe" docAuthor="Weinell, Jeffrey L., Hooper, Errol, Leviton, Alan E. &amp; Brown, Rafe M." docDate="2019" docId="A20AE119FFA6FFB1FD8908F14C9A5DE9" docLanguage="en" docName="ProcCalifAcadSci.66.1.1-49.pdf" docOrigin="Proceedings of the California Academy of Sciences 66 (1)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Lycodon" docType="key" docVersion="6" lastPageNumber="29" masterDocId="5E339961FFBDFFADFFD00A21480B5C14" masterDocTitle="Illustrated Key to the Snakes of the Philippines" masterLastPageNumber="49" masterPageNumber="1" pageNumber="28" updateTime="1720732226891" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0FEA5183467397331F35697FD6652EF6">Illustrated Key to the Snakes of the Philippines</mods:title>
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<mods:affiliation id="E1241F08190239A5433E7BC3CA7AD462">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:affiliation id="70680420A96C9D31765A912C8C9DBCEE">Herpetology Division, Institute of Biodiversity Science &amp; Sustainability, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118. * Corresponding author; Email: jweine 2 @ gmail. com</mods:affiliation>
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<treatment id="A20AE119FFA6FFB1FD8908F14C9A5DE9" ID-DOI="http://doi.org/10.5281/zenodo.12728165" ID-GBIF-Taxon="235040686" ID-Zenodo-Dep="12728165" LSID="urn:lsid:plazi:treatment:A20AE119FFA6FFB1FD8908F14C9A5DE9" httpUri="http://treatment.plazi.org/id/A20AE119FFA6FFB1FD8908F14C9A5DE9" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<taxonomicName id="EDA32B8CFFA6FFB6FD8908F14ABC5EF3" ID-CoL="5HKD" authority="Fitzinger, 1826" box="[601,695,720,743]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFA6FFB6FD8908F14ABC5EF3" bold="true" box="[601,695,720,743]" italics="true" pageId="27" pageNumber="28">Lycodon</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FE2B08D04B1E5F1C" blockId="27.[507,789,720,776]" box="[507,789,753,776]" pageId="27" pageNumber="28">Key to Philippine species.</paragraph>
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1a. Ventral body scales and subcaudal scales strongly keeled; dorsal body scales in 15 longitudinal rows at midbody..............................................
<taxonomicName id="EDA32B8CFFA6FFB6FC66091A4C9B5F46" ID-CoL="8H6BD" baseAuthorityName="Griffin" baseAuthorityYear="1909" box="[950,1168,827,850]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="philippinus">
<emphasis id="18D78C1DFFA6FFB6FC66091A4C9B5F46" box="[950,1168,827,850]" italics="true" pageId="27" pageNumber="28">Lycodon philippinus</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FFAA097A4C9B5F86" blockId="27.[122,1174,795,914]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA097A4C9B5F86" pageId="27" pageNumber="28">1b. Ventral body scales and subcaudal scales smooth; dorsal body scales in 17 longitudinal rows at midbody.................................................................... 2</keyLead>
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</keyStep>
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<paragraph id="2A1C500FFFA6FFB6FFAA09874C9B5FA9" blockId="27.[122,1168,934,989]" box="[122,1168,934,957]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA09874C9B5FA9" box="[122,1168,934,957]" pageId="27" pageNumber="28">
2a. Subcaudal scales in&gt; 130 pairs......................................
<taxonomicName id="EDA32B8CFFA6FFB6FC2309874C9B5FA9" ID-CoL="3WL2F" authorityName="Gaulke" authorityYear="2002" box="[1011,1168,934,957]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="fausti">
<emphasis id="18D78C1DFFA6FFB6FC2309874C9B5FA9" box="[1011,1168,934,957]" italics="true" pageId="27" pageNumber="28">Lycodon fausti</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FFAA09E74C9B5FC9" blockId="27.[122,1168,934,989]" box="[122,1168,966,989]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA09E74C9B5FC9" box="[122,1168,966,989]" pageId="27" pageNumber="28">2b. Subcaudal scales in &lt;130 pairs................................................. 3</keyLead>
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</keyStep>
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<paragraph id="2A1C500FFFA6FFB6FFAA09D14C9B5833" blockId="27.[122,1173,1008,1095]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA09D14C9B5833" pageId="27" pageNumber="28">
3a. Some dorsal body scale rows keeled along the entire length of the body (keels may be faint or absent from scales that have lost their outermost layer)....................
<taxonomicName id="EDA32B8CFFA6FFB6FC200E314C9B5833" ID-CoL="8H5G4" authorityName="Leviton" authorityYear="1955" box="[1008,1168,1040,1063]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="sealei">
<emphasis id="18D78C1DFFA6FFB6FC200E314C9B5833" box="[1008,1168,1040,1063]" italics="true" pageId="27" pageNumber="28">Lycodon sealei</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FFAA0E114C9B5853" blockId="27.[122,1173,1008,1095]" box="[122,1168,1072,1095]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0E114C9B5853" box="[122,1168,1072,1095]" pageId="27" pageNumber="28">3b. Dorsal body scales either all smooth, or, keeled on posterior part of body............... 4</keyLead>
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<keyStep id="915748AAFFA6FFB6FFAA0E7A4C9B5886" pageId="27" pageNumber="28">
<paragraph id="2A1C500FFFA6FFB6FFAA0E7A4C9B5866" blockId="27.[122,1168,1115,1170]" box="[122,1168,1115,1138]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0E7A4C9B5866" box="[122,1168,1115,1138]" pageId="27" pageNumber="28">4a. Subcaudal scales in&gt; 100 pairs................................................. 5</keyLead>
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<paragraph id="2A1C500FFFA6FFB6FFAA0E5A4C9B5886" blockId="27.[122,1168,1115,1170]" box="[122,1168,1147,1170]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0E5A4C9B5886" box="[122,1168,1147,1170]" pageId="27" pageNumber="28">
4b. Subcaudal scales in &lt;100 pairs..................................
<taxonomicName id="EDA32B8CFFA6FFB6FC120E5A4C9B5886" ID-CoL="3WKZY" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[962,1168,1147,1170]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="capucinus">
<emphasis id="18D78C1DFFA6FFB6FC120E5A4C9B5886" box="[962,1168,1147,1170]" italics="true" pageId="27" pageNumber="28">Lycodon capucinus</emphasis>
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<keyStep id="915748AAFFA6FFB6FFAA0E874C9B5909" pageId="27" pageNumber="28">
<paragraph id="2A1C500FFFA6FFB6FFAA0E874C9B58C9" blockId="27.[122,1174,1190,1309]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0E874C9B58C9" pageId="27" pageNumber="28">5a. Dorsal part of body without light crossbands; ventral surface of body does not have dark markings, except for a large spot in lateral corner of each ventral body scale.................. 6</keyLead>
</paragraph>
<paragraph id="2A1C500FFFA6FFB6FFAA0EC74C9B5909" blockId="27.[122,1174,1190,1309]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0EC74C9B5909" pageId="27" pageNumber="28">5b. Dorsal surface of body with light crossbands (at least the anterior part of body); ventral surface of body or tail has dark dots, spots, and/or transverse bands........................... 7</keyLead>
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<keyStep id="915748AAFFA6FFB6FFAA0F114C9B5973" pageId="27" pageNumber="28">
<paragraph id="2A1C500FFFA6FFB6FFAA0F114C9B5953" blockId="27.[122,1168,1328,1383]" box="[122,1168,1328,1351]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0F114C9B5953" box="[122,1168,1328,1351]" pageId="27" pageNumber="28">
6a. Ventral body scales&gt; 200; vertebral scales&gt; 205.......................
<taxonomicName id="EDA32B8CFFA6FFB6FC340F114C9B5953" ID-CoL="3WKZJ" authorityName="Ota &amp; Ross" authorityYear="1994" box="[996,1168,1328,1351]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="18D78C1DFFA6FFB6FC340F114C9B5953" box="[996,1168,1328,1351]" italics="true" pageId="27" pageNumber="28">Lycodon alcalai</emphasis>
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</paragraph>
<paragraph id="2A1C500FFFA6FFB6FFAA0F714C9B5973" blockId="27.[122,1168,1328,1383]" box="[122,1168,1360,1383]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0F714C9B5973" box="[122,1168,1360,1383]" pageId="27" pageNumber="28">
6b. Ventral body scales &lt;200; vertebral scales &lt;205..................
<taxonomicName id="EDA32B8CFFA6FFB6FC750F714C9B5973" authorityName="Ota &amp; Ross" authorityYear="1994" box="[933,1168,1360,1383]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="chrysopteros">
<emphasis id="18D78C1DFFA6FFB6FC750F714C9B5973" box="[933,1168,1360,1383]" italics="true" pageId="27" pageNumber="28">Lycodon chrysopteros</emphasis>
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<keyStep id="915748AAFFA6FFB6FFAA0F5A4C9B5A06" pageId="27" pageNumber="28">
<paragraph id="2A1C500FFFA6FFB6FFAA0F5A4C9B59A6" blockId="27.[122,1174,1403,1554]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0F5A4C9B59A6" pageId="27" pageNumber="28">
7a. Crossbands absent from tail and posterior part od body; ventral bands absent; number of infralabial scales in contact with anterior and posterior chin shields five........
<taxonomicName id="EDA32B8CFFA6FFB6FC190FBA4C9B59A6" ID-CoL="3WL47" authorityName="Ota &amp; Ross" authorityYear="1994" box="[969,1168,1435,1458]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="solivagus">
<emphasis id="18D78C1DFFA6FFB6FC190FBA4C9B59A6" box="[969,1168,1435,1458]" italics="true" pageId="27" pageNumber="28">Lycodon solivagus</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FFAA0F9A4C9B5A06" blockId="27.[122,1174,1403,1554]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0F9A4C9B5A06" pageId="27" pageNumber="28">7b. Dorsal crossbands present throughout length of body and tail; additionally, at least one of the following combinations of characters true: (1) number of infralabial scales in contact with anterior and posterior chin shields six; (2) ventral bands present........................... 8</keyLead>
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<keyStep id="915748AAFFA6FFB6FFAA0C044C9B5A88" pageId="27" pageNumber="28">
<paragraph id="2A1C500FFFA6FFB6FFAA0C044C9B5A48" blockId="27.[122,1174,1573,1692]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0C044C9B5A48" pageId="27" pageNumber="28">
8a. Dorsal light crossbands&gt; 50 on body,&gt; 27 on tail; tail length 36–45% (usually&gt; 38%) of snout-vent length.....................................................
<taxonomicName id="EDA32B8CFFA6FFB6FC050C644C9B5A48" ID-CoL="3WL3H" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[981,1168,1605,1628]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="18D78C1DFFA6FFB6FC050C644C9B5A48" box="[981,1168,1605,1628]" italics="true" pageId="27" pageNumber="28">Lycodon muelleri</emphasis>
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<paragraph id="2A1C500FFFA6FFB6FFAA0C444C9B5A88" blockId="27.[122,1174,1573,1692]" pageId="27" pageNumber="28">
<keyLead id="9152F33AFFA6FFB6FFAA0C444C9B5A88" pageId="27" pageNumber="28">8b. Dorsal light crossbands &lt;30 on body, &lt;25 on tail; tail length 33–39% (usually &lt;38%) of snout-vent length............................................................. 9</keyLead>
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<keyStep id="915748AAFFA1FFB1FFAB0AFD4C9A5D07" pageId="28" pageNumber="29">
<paragraph id="2A1C500FFFA1FFB1FFAB0AFD4C9A5CE7" blockId="28.[123,1169,220,275]" box="[123,1169,220,243]" pageId="28" pageNumber="29">
<keyLead id="9152F33AFFA1FFB1FFAB0AFD4C9A5CE7" box="[123,1169,220,243]" pageId="28" pageNumber="29">
9a. Dorsal light crossbands&gt; 20 on body,&gt; 15 on tail....................
<taxonomicName id="EDA32B8CFFA1FFB1FC030AFD4C9A5CE7" ID-CoL="3WKZV" authorityName="Ota &amp; Ross" authorityYear="1994" box="[979,1169,220,243]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="bibonius">
<emphasis id="18D78C1DFFA1FFB1FC030AFD4C9A5CE7" box="[979,1169,220,243]" italics="true" pageId="28" pageNumber="29">Lycodon bibonius</emphasis>
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10a. Body and tail with dark rings completely encircling the body; number of supralabial scales usually ten (fourth through sixth border eye); dorsal body scales reducing to 15 longitudinal rows at approximately ventral 119...................................
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10b. Dorsal part of body and tail with dark crossbands often extending onto lateral edges of ventral body scales, but not completely encircling the body; number of supralabial scales 8–9 (fourth and fifth border eye); dorsal body scales reduce to 15 longitudinal rows between ventral body scales 131–145.............................................
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1a. Nasal scales fused to internasal scales anterior to rostral (
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); dorsal surface of body with four brown longitudinal stripes..................................
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1b. Nasal scales not fused to internasal scales; dorsal surface of body uniformly colored above, without longitudinal stripes........................................
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1a. Scales in 19 longitudinal rows at midbody; two prefrontal scales (
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)........................................................................
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1b. Scales in 21–25 longitudinal rows at midbody; three prefrontal scales (
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2a. Rostral scale not divided horizontally; upper lip yellow; ventral body scales 213–245.................................................................
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2b. Rostral scale divided horizontally; upper lip brown; ventral body scales 195–205...................................................................
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<paragraph id="2A1C500FFFA8FFB8FE390ADD4B235D07" blockId="21.[489,808,219,275]" box="[489,808,252,275]" pageId="21" pageNumber="22">Key to Philippine subspecies.</paragraph>
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1a. Lateral body scales speckled (black with a bright central fleck); distinct transverse bands only in young animals; in the
<collectingCountry id="52B4109FFFA8FFB8FE730B674A155D49" box="[419,542,326,349]" name="Philippines" pageId="21" pageNumber="22">Philippines</collectingCountry>
, only known from Sibutu Island (
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Archipelago)........................................................
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<emphasis id="18D78C1DFFA8FFB8FC9A0B474C9A5D69" box="[842,1169,358,381]" italics="true" pageId="21" pageNumber="22">Chrysopelea paradisi paradisi</emphasis>
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1b. Lateral body scales mostly light; color pattern of dorsal body surface variable, but transverse bands almost always present; occurs throughout the
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, but not known from Sabtang Island (
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Island group, off northern Luzon)..........
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<emphasis id="18D78C1DFFA8FFB8FCEC0BE74C9A5DC9" box="[828,1169,454,477]" italics="true" pageId="21" pageNumber="22">Chrysopelea paradisi variabilis</emphasis>
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<paragraph id="2A1C500FFFA8FFB8FE7808384B625E24" blockId="21.[424,873,505,560]" box="[424,873,537,560]" pageId="21" pageNumber="22">Key to Philippine species and subspecies.</paragraph>
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<keyLead id="9152F33AFFA8FFB8FFAB08654C9A5E8F" pageId="21" pageNumber="22">1a. Number of subcaudal scales 87–107; number of ventral body scales + subcaudal scales &lt;330; adults lack distinctive black-edged white bars on side of body; juveniles usually with whitish crossbands present............................................................ 2</keyLead>
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1b. Number of subcaudal scales 102–114; number of ventral body scales + subcaudal scales&gt; 325; adults and juveniles with a series of short black-edged white bars on side of body.................................................................
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<emphasis id="18D78C1DFFA8FFB8FCAE08C54C9A5EEF" box="[894,1169,740,763]" italics="true" pageId="21" pageNumber="22">Coelognathus philippinus</emphasis>
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2a. Adults with tail much lighter than posterior portion of body..........................................................................
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<emphasis id="18D78C1DFFA8FFB8FCDD090F4C9A5F51" box="[781,1169,814,837]" italics="true" pageId="21" pageNumber="22">Coelognathus erythrurus erythrurus</emphasis>
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3a. Adults uniform light brown to reddish brown throughout the length of body and tail; no darkening posteriorly................................
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<emphasis id="18D78C1DFFA8FFB8FCD409984C9A5FC4" box="[772,1169,953,976]" italics="true" pageId="21" pageNumber="22">Coelognathus erythrurus manillensis</emphasis>
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3b. Adults become darker posteriorly along length of body and tail....................................................................
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<paragraph id="2A1C500FFFA8FFB8FE780E6D4B625877" blockId="21.[424,873,1067,1123]" box="[424,873,1100,1123]" pageId="21" pageNumber="22">Key to Philippine species and subspecies.</paragraph>
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1a. Ventral body scales&gt; 137; hemipenes narrow, elongate with minute spines ornamenting walls; usually eight supralabial scales, three bordering eye; usually two anterior temporal scales (
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); prominent white spots present along lateral edge of ventral body scales; many dark, usually triangularly-shaped, blotches on ventral body scales............................. 2
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1b. Ventral body scales &lt;136; hemipenes robust, inner walls uniformly spinose, spines of moderate size; supralabial scales 7–8, two or three bordering eye; anterior temporal scale usually one (
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); white spots along lateral edge of ventral body scales, if present, obscure; few or no dark triangular blotches on ventral body scales..................................... 3
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32. Lateral view of head of (A)
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(KU 344152). Anterior temporal scales (green).
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2a. Tail length/total length
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(males),
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(females); subcaudals 52–59 (males), 42–48 (females); white spots along lateral edge of ventral body scales prominent................................................................
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<emphasis id="18D78C1DFFABFFBBFCB00B3C4C9A5D20" box="[864,1169,285,308]" italics="true" pageId="22" pageNumber="23">Cyclocorus lineatus lineatus</emphasis>
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2b. Tail length/total length
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(males),
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(females); subcaudal scales 42–53 (males), 33–44 (females); white spots along lateral edge of ventral body scales very small.........................................................
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<emphasis id="18D78C1DFFABFFBBFCBA0B5F4C9A5D81" box="[874,1169,382,405]" italics="true" pageId="22" pageNumber="23">Cyclocorus lineatus alcalai</emphasis>
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3a. Usually seven supralabial scales, two bordering eye..........
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<emphasis id="18D78C1DFFABFFBBFC840B8B4C9A5DD5" box="[852,1169,426,449]" italics="true" pageId="22" pageNumber="23">Cyclocorus nuchalis nuchalis</emphasis>
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3b. Usually eight supralabial scales, three bordering eye..........
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<emphasis id="18D78C1DFFABFFBBFCB70BEB4C9A5DF5" box="[871,1169,458,481]" italics="true" pageId="22" pageNumber="23">Cyclocorus nuchalis taylori</emphasis>
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<emphasis id="18D78C1DFFA9FFB9FD800E224ACA580E" bold="true" box="[592,705,1027,1050]" italics="true" pageId="20" pageNumber="21">Calliophis</emphasis>
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<paragraph id="2A1C500FFFA9FFB9FE2C0E024B1E582E" blockId="20.[508,789,1027,1082]" box="[508,789,1059,1082]" pageId="20" pageNumber="21">Key to Philippine species.</paragraph>
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1a. Dorsal body surface with black and white crossbands, rather than longitudinal stripes; head black dorsally; tail red dorsally....................................
<taxonomicName id="EDA32B8CFFA9FFB9FC020E4F4C9A5891" ID-CoL="8GPFF" authorityName="Brown, Smart, Leviton &amp; Smith" authorityYear="2018" box="[978,1169,1134,1157]" class="Reptilia" family="Elapidae" genus="Calliophis" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="salitan">
<emphasis id="18D78C1DFFA9FFB9FC020E4F4C9A5891" box="[978,1169,1134,1157]" italics="true" pageId="20" pageNumber="21">Calliophis salitan</emphasis>
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<keyLead id="9152F33AFFA9FFB9FFAB0EAF4C9A58B1" box="[123,1169,1166,1189]" pageId="20" pageNumber="21">1b. Dorsal body surface with pale longitudinal stripes, rather than crossbands............... 2</keyLead>
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2a. Black bands on ventral body surface not in contact with black of lateral body surface; distinct white longitudinal stripe present on lateral body surface, occupying entire first, or parts of first and second, dorsal body scale rows...............................
<taxonomicName id="EDA32B8CFFA9FFB9FC6C0ED94C9A591B" baseAuthorityName="Peters" baseAuthorityYear="1881" box="[956,1169,1272,1295]" class="Reptilia" family="Elapidae" genus="Calliophis" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bilineata">
<emphasis id="18D78C1DFFA9FFB9FC6C0ED94C9A591B" box="[956,1169,1272,1295]" italics="true" pageId="20" pageNumber="21">Calliophis bilineata</emphasis>
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<keyLead id="9152F33AFFA9FFB9FFAB0F394C9A595B" pageId="20" pageNumber="21">2b. Black bands on ventral body surface in contact with black of lateral body surface; distinct white stripe absent from first or second dorsal body scale rows............................. 3</keyLead>
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3a. Cream-colored bands on ventral body surface do not extend onto side of body above first dorsal body scale row.............................................
<taxonomicName id="EDA32B8CFFA9FFB9FC690FA24C9A598E" ID-CoL="8H66R" baseAuthorityName="Steindachner" baseAuthorityYear="1891" box="[953,1169,1411,1434]" class="Reptilia" family="Elapidae" genus="Calliophis" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="suluensis">
<emphasis id="18D78C1DFFA9FFB9FC690FA24C9A598E" box="[953,1169,1411,1434]" italics="true" pageId="20" pageNumber="21">Calliophis suluensis</emphasis>
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3b. Cream-colored bands on ventral body surface extend onto side of body, often to fourth dorsal body scale row...............................................
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<emphasis id="18D78C1DFFA9FFB9FC7D0FE24C9A59CE" box="[941,1169,1475,1498]" italics="true" pageId="20" pageNumber="21">Calliophis philippina</emphasis>
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<emphasis id="18D78C1DFFA9FFB9FD860FD14AB05A13" bold="true" box="[598,699,1520,1543]" italics="true" pageId="20" pageNumber="21">Cerberus</emphasis>
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1a. Dorsal body scales in 27–31 longitudinal rows at midbody; supralabial scales and other scales of head have a strongly papillate texture...........................
<taxonomicName id="EDA32B8CFFA9FFB9FC670C7A4C9A5A66" ID-CoL="69KJ9" authorityName="Boulenger" authorityYear="1896" box="[951,1169,1627,1650]" class="Reptilia" family="Homalopsidae" genus="Cerberus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microlepis">
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1b. Dorsal body scales in &lt;28 longitudinal rows at midbody; supralabial scales and other scales on head smooth.................................................
<taxonomicName id="EDA32B8CFFA9FFB9FC670CBA4C9A5AA6" ID-CoL="6LYJN" box="[951,1169,1691,1714]" class="Reptilia" family="Homalopsidae" genus="Cerberus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="schneideri">
<emphasis id="18D78C1DFFA9FFB9FC670CBA4C9A5AA6" box="[951,1169,1691,1714]" italics="true" pageId="20" pageNumber="21">Cerberus schneideri</emphasis>
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<keyLead id="9152F33AFFAAFFBAFFAB098B4C9A5FF5" pageId="23" pageNumber="24">1a. Temporal region of head heavily pigmented with melanin; number of white dorsal body bands usually &lt;60................................................................. 2</keyLead>
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1b. Temporal region of head not heavily pigmented with melanin; number of white dorsal body bands usually&gt; 60.......................................
<taxonomicName id="EDA32B8CFFAAFFBAFCA60E2B4C9A5835" ID-CoL="88C3R" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[886,1169,1034,1057]" class="Reptilia" family="Elapidae" genus="Hemibungarus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="gemianulis">
<emphasis id="18D78C1DFFAAFFBAFCA60E2B4C9A5835" box="[886,1169,1034,1057]" italics="true" pageId="23" pageNumber="24">Hemibungarus gemianulis</emphasis>
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2a. Number of ventral body scales 223–233 (males), 252–259 (females); black ventral body bands not divided by thin white bands.............................
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<emphasis id="18D78C1DFFAAFFBAFCA90E744C9A5878" box="[889,1169,1109,1132]" italics="true" pageId="23" pageNumber="24">Hemibungarus calligaster</emphasis>
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2b. Number of ventral body scales &lt;223 (males; unknown for females); black ventral body bands divided by thin white band..................................
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<emphasis id="18D78C1DFFAAFFBAFC590EB44C9A58B8" box="[905,1169,1173,1196]" italics="true" pageId="23" pageNumber="24">Hemibungarus mcclungi</emphasis>
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<emphasis id="18D78C1DFFAAFFBAFDEC0EE34AD858CD" bold="true" box="[572,723,1218,1241]" italics="true" pageId="23" pageNumber="24">Hologerrhum</emphasis>
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1a. Posterior border of parietal scales unnotched (
<figureCitation id="B2984C8AFFAAFFBAFD480F2C4AF65930" box="[664,765,1293,1316]" captionStart="FIGURE" captionStartId="23.[155,167,1657,1676]" captionTargetBox="[293,1003,1425,1650]" captionTargetId="figure-516@23.[293,1003,1425,1650]" captionTargetPageId="23" captionText="FIGURE 34. Dorsal view of head of (A) Hologerrhum dermali (CMNH 5075), (B) Hologerrhum philippinum (KU 330056). Parietal scales (reddish brown); black arrows point to posterior margin of parietal scale, showing unnotched pari- etal pattern (A), or notched parietal pattern (B)." figureDoi="http://doi.org/10.5281/zenodo.11512750" httpUri="https://zenodo.org/record/11512750/files/figure.png" pageId="23" pageNumber="24">Fig. 34A</figureCitation>
); dark midventral stripe present; dark midlabial stripe present.......................................
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1b. Posterior border of parietal scales notched (
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); dark midventral stripe absent; dark midlabial stripe absent........................................
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1a. Loreal scale absent (
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); number of ventral body scales I77–188............................................................................
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1b. Loreal scale present (
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); number of ventral body scales 188–199...........................................................................
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1a. Dorsal body scales in 15 longitudinal rows at midbody; vertebral scales enlarged relative to other dorsal body scales......................................
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3a. Yellow stripe present on neck (in life); endemic to
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Archipelago............................................................................
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3b. Yellow stripe absent from neck (in life); not known from
<collectingRegion id="E8679EEDFFABFFBBFCDD09A44B355F88" box="[781,830,901,924]" country="Philippines" name="Sulu" pageId="22" pageNumber="23">Sulu</collectingRegion>
Archipelago.......................................................................
<taxonomicName id="EDA32B8CFFABFFBBFC5509874C9A5FA9" ID-CoL="6CJTN" authorityName="Griffin" authorityYear="1909" box="[901,1169,934,957]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="18D78C1DFFABFFBBFC5509874C9A5FA9" box="[901,1169,934,957]" italics="true" pageId="22" pageNumber="23">Dendrelaphis fuliginosus</emphasis>
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4a. Number of black longitudinal stripes at midbody eight; known from
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, Balabac, and
<location id="2F7C06D4FFABFFBBFF4709D3495F581D" LSID="urn:lsid:plazi:treatment:A20AE119FFABFFBBFDEE0BDF4C9A594D:2F7C06D4FFABFFBBFF4709D3495F581D" box="[151,340,1010,1033]" name="Busuanga Islands" pageId="22" pageNumber="23" stateProvince="Palawan">Busuanga Islands</location>
............................................
<taxonomicName id="EDA32B8CFFABFFBBFC7A09D34C9A581D" ID-CoL="34M3T" authorityName="Van Rooijen &amp; Vogel" authorityYear="2012" box="[938,1169,1010,1033]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="levitoni">
<emphasis id="18D78C1DFFABFFBBFC7A09D34C9A581D" box="[938,1169,1010,1033]" italics="true" pageId="22" pageNumber="23">Dendrelaphis levitoni</emphasis>
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4b. Fewer than eight black longitudinal stripes at midbody; not from
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, Balabac, and
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............................................................. 5
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5a. Thin black stripe usually present along border of ventral scales and first row of dorsal body scales; second narrow black stripe may or may not be present on anterior one-fifth of body along border of second and third dorsal body scale rows; additional dark body stripes absent...........................................................
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5b. Distinct black stripe present along border of ventral scales and first longitudinal row of dorsal body scales, and along border of second and third dorsal body scale rows; additional distinct dark stripes present on posterior third of body, along border of fourth and fifth, and fifth and sixth dorsal body scale rows.................................
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1a. Internasal scales absent (fused to prefrontal scales) (
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); dorsal body scales in 13 longitudinal rows at midbody..........................................
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1b. Internasal scales present (distinct from prefrontal scales) (
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); dorsal body scales in 15 longitudinal rows at midbody..................................
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<mods:title id="9FF8C7B804D853CECCA863B2E1D04FEA">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="083EC9468388B667BDD727C04B84A6AF">
<mods:date id="9F516C028E5D5E2A59719147BA15C629">2019</mods:date>
<mods:detail id="E36219AF108F137F998366ACBC2BDEE7" type="pubDate">
<mods:number id="FC7EF1F1B60A35DD9608DEA0A8011286">2019-09-30</mods:number>
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<mods:detail id="4D94A8BACBF0F0D05617CA50DD6F3D63" type="volume">
<mods:number id="DBB672F1CD70AAAB12891A90C883FF23">66</mods:number>
</mods:detail>
<mods:detail id="2F4405F3BB2B40C8ED3A3BBA3B6AB212" type="issue">
<mods:number id="6BCEEEA8368E7F758D72C82805C13F99">1</mods:number>
</mods:detail>
<mods:extent id="5CCE8D555DC725F6E7275D8AA68E8653" unit="page">
<mods:start id="1E8CD36C9043B135B00FF57D956C5476">1</mods:start>
<mods:end id="DBD378F996DF1A0FAAC5FE1BE6BBCFA4">49</mods:end>
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<mods:classification id="D244BF712633AE54FD779375FDD08156">journal article</mods:classification>
<mods:identifier id="7067000654A42DE03ED0A72D9782256E" type="CLB-Dataset">299708</mods:identifier>
<mods:identifier id="E9D633C069265F73C1259796B68FC6A9" type="DOI">10.5281/zenodo.11512680</mods:identifier>
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<mods:identifier id="10A3CCF8D51F612E58C697910DB5BE92" type="Zenodo-Dep">11512680</mods:identifier>
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<treatment id="A20AE119FFACFFBFFD9808ED4C9A5E98" ID-DOI="http://doi.org/10.5281/zenodo.12728090" ID-GBIF-Taxon="235040675" ID-Zenodo-Dep="12728090" LSID="urn:lsid:plazi:treatment:A20AE119FFACFFBFFD9808ED4C9A5E98" httpUri="http://treatment.plazi.org/id/A20AE119FFACFFBFFD9808ED4C9A5E98" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="62B90384FFACFFBCFD9808ED4AC25EF7" box="[584,713,716,739]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="2A1C500FFFACFFBCFD9808ED4AC25EF7" blockId="17.[511,786,716,771]" box="[584,713,716,739]" pageId="17" pageNumber="18">
<taxonomicName id="EDA32B8CFFACFFBCFD9808ED4AC25EF7" ID-CoL="87CG5" ID-ENA="169863" authority="Oppel, 1811" box="[584,713,716,739]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="subFamily" subFamily="Colubrinae">
<emphasis id="18D78C1DFFACFFBCFD9808ED4AC25EF7" bold="true" box="[584,713,716,739]" pageId="17" pageNumber="18">Colubrinae</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="62B90384FFACFFBCFE2F08CD4B195F17" box="[511,786,748,771]" pageId="17" pageNumber="18" type="multiple">
<paragraph id="2A1C500FFFACFFBCFE2F08CD4B195F17" blockId="17.[511,786,716,771]" box="[511,786,748,771]" pageId="17" pageNumber="18">Key to Philippine genera.</paragraph>
</subSubSection>
<subSubSection id="62B90384FFACFFBFFFAB09364C9A5E98" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="key">
<key id="5E32073EFFACFFBFFFAB09364C9A5E98" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<keyStep id="915748AAFFACFFBCFFAB09364C9A5F5A" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB09364C9A5F3A" blockId="17.[123,1169,791,846]" box="[123,1169,791,814]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB09364C9A5F3A" box="[123,1169,791,814]" pageId="17" pageNumber="18">1a. Dorsal body scales in 19–27 longitudinal rows at midbody........................... 2</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB09164C9A5F5A" blockId="17.[123,1169,791,846]" box="[123,1169,823,846]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB09164C9A5F5A" box="[123,1169,823,846]" pageId="17" pageNumber="18">1b. Dorsal body scales in 14–18 longitudinal rows at midbody........................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFACFFBCFFAB09404C9A5F8C" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB09404C9A5F6C" blockId="17.[123,1169,865,920]" box="[123,1169,865,888]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB09404C9A5F6C" box="[123,1169,865,888]" pageId="17" pageNumber="18">
2a. Pupil strongly vertically elliptical (
<figureCitation id="B2984C8AFFACFFBCFDC909404A715F6C" box="[537,634,865,888]" captionStart="FIGURE" captionStartId="17.[155,167,1128,1147]" captionTargetBox="[302,1003,932,1104]" captionTargetId="figure-757@17.[293,1003,932,1112]" captionTargetPageId="17" captionText="FIGURE 28. Lateral view of head of (A) Boiga angulata (KU 343853), (B) Coelognathus erythrurus manillensis (KU 335195). Note vertically elliptical (A) versus rounded pupil (B)." figureDoi="http://doi.org/10.5281/zenodo.11512738" httpUri="https://zenodo.org/record/11512738/files/figure.png" pageId="17" pageNumber="18">Fig. 28A</figureCitation>
)............................
<taxonomicName id="EDA32B8CFFACFFBCFBD209404C495F6C" authorityName="Fitzinger" authorityYear="1826" box="[1026,1090,865,888]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFBD209404C495F6C" box="[1026,1090,865,888]" italics="true" pageId="17" pageNumber="18">Boiga</emphasis>
</taxonomicName>
(p. 20)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB09A04C9A5F8C" blockId="17.[123,1169,865,920]" box="[123,1169,897,920]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB09A04C9A5F8C" box="[123,1169,897,920]" pageId="17" pageNumber="18">
2b. Pupil circular or slightly vertically elliptical (
<figureCitation id="B2984C8AFFACFFBCFD5909A04AE15F8C" box="[649,746,897,920]" captionStart="FIGURE" captionStartId="17.[155,167,1128,1147]" captionTargetBox="[302,1003,932,1104]" captionTargetId="figure-757@17.[293,1003,932,1112]" captionTargetPageId="17" captionText="FIGURE 28. Lateral view of head of (A) Boiga angulata (KU 343853), (B) Coelognathus erythrurus manillensis (KU 335195). Note vertically elliptical (A) versus rounded pupil (B)." figureDoi="http://doi.org/10.5281/zenodo.11512738" httpUri="https://zenodo.org/record/11512738/files/figure.png" pageId="17" pageNumber="18">Fig. 28B</figureCitation>
).............................. 3
</keyLead>
</paragraph>
</keyStep>
<caption id="7EDC0087FFACFFBCFF4B0E494AA85881" ID-DOI="http://doi.org/10.5281/zenodo.11512738" ID-Zenodo-Dep="11512738" httpUri="https://zenodo.org/record/11512738/files/figure.png" pageId="17" pageNumber="18" startId="17.[155,167,1128,1147]" targetBox="[302,1003,932,1104]" targetPageId="17" targetType="figure">
<paragraph id="2A1C500FFFACFFBCFF4B0E494AA85881" blockId="17.[123,1173,1128,1173]" pageId="17" pageNumber="18">
<smallCapsWord id="2CFAC6D3FFACFFBCFF4B0E4948D3586E" baselines="1142,1142" box="[155,216,1128,1147]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="17" pageNumber="18">FIGURE</smallCapsWord>
28. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFACFFBCFE290E494A74586F" box="[505,639,1128,1147]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angulata">
<emphasis id="18D78C1DFFACFFBCFE290E494A74586F" box="[505,639,1128,1147]" italics="true" pageId="17" pageNumber="18">Boiga angulata</emphasis>
</taxonomicName>
(KU 343853), (B)
<taxonomicName id="EDA32B8CFFACFFBCFCF90E494C63586F" authorityName="Jan" authorityYear="1863" box="[809,1128,1128,1147]" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="subSpecies" species="erythrurus" subSpecies="manillensis">
<emphasis id="18D78C1DFFACFFBCFCF90E494C63586F" box="[809,1128,1128,1147]" italics="true" pageId="17" pageNumber="18">Coelognathus erythrurus manillensis</emphasis>
</taxonomicName>
(KU 335195). Note vertically elliptical (A) versus rounded pupil (B).
</paragraph>
</caption>
<keyStep id="915748AAFFACFFBCFFAB0E9D4C9A5927" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB0E9D4C9A58E7" blockId="17.[123,1174,1212,1331]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0E9D4C9A58E7" pageId="17" pageNumber="18">
3a. Dorsal body scales in 23–27 longitudinal rows at midbody; body emerald green in life, blue after preservation in alcohol......................................
<taxonomicName id="EDA32B8CFFACFFBCFC130EFD4C4958E7" baseAuthorityName="Dowling" baseAuthorityYear="1958" box="[963,1090,1244,1267]" class="Reptilia" family="Colubridae" genus="Gonyosoma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFC130EFD4C4958E7" box="[963,1090,1244,1267]" italics="true" pageId="17" pageNumber="18">Gonyosoma</emphasis>
</taxonomicName>
(p. 24)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB0EDD4C9A5927" blockId="17.[123,1174,1212,1331]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0EDD4C9A5927" pageId="17" pageNumber="18">
3b. Dorsal body scales in 21 longitudinal rows at midbody; body not emerald green in life, not blue after preservation in alcohol....................................
<taxonomicName id="EDA32B8CFFACFFBCFC7E0F3D4C495927" baseAuthorityName="Helfenberger" baseAuthorityYear="2001" box="[942,1090,1308,1331]" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFC7E0F3D4C495927" box="[942,1090,1308,1331]" italics="true" pageId="17" pageNumber="18">Coelognathus</emphasis>
</taxonomicName>
(p. 22)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFACFFBCFFAB0F674C9A59A9" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB0F674C9A5969" blockId="17.[123,1174,1350,1469]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0F674C9A5969" pageId="17" pageNumber="18">
4a. Dorsal body scales in 15–17 longitudinal rows that do not reduce posteriorly; subcaudal scales 27–54 pairs.....................................................
<taxonomicName id="EDA32B8CFFACFFBCFC0D0F474C495969" box="[989,1090,1382,1405]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFC0D0F474C495969" box="[989,1090,1382,1405]" italics="true" pageId="17" pageNumber="18">Oligodon</emphasis>
</taxonomicName>
(p. 30)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB0FA74C9A59A9" blockId="17.[123,1174,1350,1469]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0FA74C9A59A9" pageId="17" pageNumber="18">4b. Dorsal body scales may or may not reduce posteriorly; if dorsal body scales do not reduce posteriorly, then subcaudal scales in&gt; 100 pairs.................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFACFFBCFFAB0FF04C9A5A7C" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB0FF04C9A5A3C" blockId="17.[123,1173,1489,1640]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0FF04C9A5A3C" pageId="17" pageNumber="18">
5a. Lateral edge of supraocular scale robust, protruding over eye rather than curving around eye (
<figureCitation id="B2984C8AFFACFFBCFF4F0FD0490F5A1C" box="[159,260,1521,1544]" captionStart="FIGURE" captionStartId="18.[155,167,426,445]" captionTargetBox="[293,999,216,411]" captionTargetId="figure-695@18.[293,1003,216,411]" captionTargetPageId="18" captionText="FIGURE 29. Lateral view of head of (A) Ptyas luzonensis (KU 306605), (B) Stegonotus muelleri (KU 344097). Supraocular scale (reddish brown) of Ptyas robust, protruding over eye." figureDoi="http://doi.org/10.5281/zenodo.11512740" httpUri="https://zenodo.org/record/11512740/files/figure.png" pageId="17" pageNumber="18">Fig. 29A</figureCitation>
); dorsal body scales usually in 14, 16, or 18 longitudinal rows at midbody.........................................................................
<taxonomicName id="EDA32B8CFFACFFBCFBD70C304C495A3C" authorityName="Fitzinger" authorityYear="1843" box="[1031,1090,1553,1576]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFBD70C304C495A3C" box="[1031,1090,1553,1576]" italics="true" pageId="17" pageNumber="18">Ptyas</emphasis>
</taxonomicName>
(p. 33)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB0C104C9A5A7C" blockId="17.[123,1173,1489,1640]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0C104C9A5A7C" pageId="17" pageNumber="18">
5b. Lateral edge of supraocular scale not robust, and curves with contour of eye (
<figureCitation id="B2984C8AFFACFFBCFC0E0C104C4B5A5C" box="[990,1088,1585,1608]" captionStart="FIGURE" captionStartId="18.[155,167,426,445]" captionTargetBox="[293,999,216,411]" captionTargetId="figure-695@18.[293,1003,216,411]" captionTargetPageId="18" captionText="FIGURE 29. Lateral view of head of (A) Ptyas luzonensis (KU 306605), (B) Stegonotus muelleri (KU 344097). Supraocular scale (reddish brown) of Ptyas robust, protruding over eye." figureDoi="http://doi.org/10.5281/zenodo.11512740" httpUri="https://zenodo.org/record/11512740/files/figure.png" pageId="17" pageNumber="18">Fig. 29B</figureCitation>
); dorsal body scales usually in 15 or 17 longitudinal rows at midbody......................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFACFFBCFFAB0C5D4C9A5AA7" pageId="17" pageNumber="18">
<paragraph id="2A1C500FFFACFFBCFFAB0C5D4C9A5A87" blockId="17.[123,1169,1660,1715]" box="[123,1169,1660,1683]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0C5D4C9A5A87" box="[123,1169,1660,1683]" pageId="17" pageNumber="18">
6a. Loreal scale absent; dorsal body scales in 15 longitudinal rows at midbody..
<taxonomicName id="EDA32B8CFFACFFBCFC350C5D4C495A87" authorityName="Fitzinger" authorityYear="1843" box="[997,1090,1660,1683]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFACFFBCFC350C5D4C495A87" box="[997,1090,1660,1683]" italics="true" pageId="17" pageNumber="18">Liopeltis</emphasis>
</taxonomicName>
(p. 27)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFACFFBCFFAB0CBD4C9A5AA7" blockId="17.[123,1169,1660,1715]" box="[123,1169,1692,1715]" pageId="17" pageNumber="18">
<keyLead id="9152F33AFFACFFBCFFAB0CBD4C9A5AA7" box="[123,1169,1692,1715]" pageId="17" pageNumber="18">6b. Loreal scale present; dorsal body scales in 15–17 longitudinal scale rows at midbody..... 7</keyLead>
</paragraph>
</keyStep>
<caption id="7EDC0087FFAFFFBFFF4B0B8B4A945DC2" ID-DOI="http://doi.org/10.5281/zenodo.11512740" ID-Zenodo-Dep="11512740" httpUri="https://zenodo.org/record/11512740/files/figure.png" pageId="18" pageNumber="19" startId="18.[155,167,426,445]" targetBox="[293,999,216,411]" targetPageId="18" targetType="figure">
<paragraph id="2A1C500FFFAFFFBFFF4B0B8B4A945DC2" blockId="18.[123,1173,426,470]" pageId="18" pageNumber="19">
<smallCapsWord id="2CFAC6D3FFAFFFBFFF4B0B8B48D35DAF" baselines="440,439" box="[155,216,426,445]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="18" pageNumber="19">FIGURE</smallCapsWord>
29. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFAFFFBFFE3B0B8B4A725DA9" baseAuthorityName="Gunther" baseAuthorityYear="1873" box="[491,633,426,445]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="luzonensis">
<emphasis id="18D78C1DFFAFFFBFFE3B0B8B4A725DA9" box="[491,633,426,445]" italics="true" pageId="18" pageNumber="19">Ptyas luzonensis</emphasis>
</taxonomicName>
(KU 306605), (B)
<taxonomicName id="EDA32B8CFFAFFFBFFCCD0B8B4BCD5DA9" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[797,966,426,445]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="18D78C1DFFAFFFBFFCCD0B8B4BCD5DA9" box="[797,966,426,445]" italics="true" pageId="18" pageNumber="19">Stegonotus muelleri</emphasis>
</taxonomicName>
(KU 344097). Supraocular scale (reddish brown) of
<taxonomicName id="EDA32B8CFFAFFFBFFEA90BE249A35DC2" authorityName="Fitzinger" authorityYear="1843" box="[377,424,451,470]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFAFFFBFFEA90BE249A35DC2" box="[377,424,451,470]" italics="true" pageId="18" pageNumber="19">Ptyas</emphasis>
</taxonomicName>
robust, protruding over eye.
</paragraph>
</caption>
<keyStep id="915748AAFFAFFFBFFFAB0BD44C9A5E98" pageId="18" pageNumber="19">
<paragraph id="2A1C500FFFAFFFBFFFAB0BD44C9A5E58" blockId="18.[123,1174,501,652]" pageId="18" pageNumber="19">
<keyLead id="9152F33AFFAFFFBFFFAB0BD44C9A5E58" pageId="18" pageNumber="19">
7a. One or more of the following combinations of characters true: (1) dorsal body scales in 17 longitudinal rows at midbody, and 184–216 smooth ventral body scales; (2) dorsal body scales in 15 longitudinal rows at midbody and ventral body scales keeled.........
<taxonomicName id="EDA32B8CFFAFFFBFFC3708144C495E58" box="[999,1090,565,588]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFAFFFBFFC3708144C495E58" box="[999,1090,565,588]" italics="true" pageId="18" pageNumber="19">Lycodon</emphasis>
</taxonomicName>
(p. 28)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFAFFFBFFFAB08744C9A5E98" blockId="18.[123,1174,501,652]" pageId="18" pageNumber="19">
<keyLead id="9152F33AFFAFFFBFFFAB08744C9A5E98" pageId="18" pageNumber="19">
7b. Dorsal body scales in 17 longitudinal rows at midbody, 220–232 smooth ventral body scales.............................................................
<taxonomicName id="EDA32B8CFFAFFFBFFC1D08544C495E98" authorityName="A.M.C.Duméril, Bibron &amp; A.H.A.Duméril" authorityYear="1854" box="[973,1090,629,652]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFAFFFBFFC1D08544C495E98" box="[973,1090,629,652]" italics="true" pageId="18" pageNumber="19">Stegonotus</emphasis>
</taxonomicName>
(p. 34)
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

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<document id="1ACE05816345FDDAAA524B4732B71F98" ID-CLB-Dataset="299708" ID-DOI="10.5281/zenodo.11512680" ID-GBIF-Dataset="ccd739a4-10c3-4700-b27e-459bd82f1676" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512680" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1717720566437" checkinUser="felipe" docAuthor="Weinell, Jeffrey L., Hooper, Errol, Leviton, Alan E. &amp; Brown, Rafe M." docDate="2019" docId="A20AE119FFADFFBDFDEC0B5D4C9A5A77" docLanguage="en" docName="ProcCalifAcadSci.66.1.1-49.pdf" docOrigin="Proceedings of the California Academy of Sciences 66 (1)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Ahaetulliinae" docType="key" docVersion="4" lastPageNumber="17" masterDocId="5E339961FFBDFFADFFD00A21480B5C14" masterDocTitle="Illustrated Key to the Snakes of the Philippines" masterLastPageNumber="49" masterPageNumber="1" pageNumber="17" updateTime="1720732226891" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="838E0C5C40A557428B9D5B213BB37175">Illustrated Key to the Snakes of the Philippines</mods:title>
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<mods:namePart id="DE69EA6BA886594F8941F1CD9112F9A8">Weinell, Jeffrey L.</mods:namePart>
<mods:affiliation id="C6359C5F99C3A492B75C531B0D2EF290">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:namePart id="8BD11C200B7C673CD27FCEFCAC60980C">Hooper, Errol</mods:namePart>
<mods:affiliation id="521CC2F2C9A5591B8A7CAEB2E3189D03">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:namePart id="2C2702900E3999090E4556F9DF99BCB0">Leviton, Alan E.</mods:namePart>
<mods:affiliation id="97305DC0D92853480534F45FF44CF69A">Herpetology Division, Institute of Biodiversity Science &amp; Sustainability, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118. * Corresponding author; Email: jweine 2 @ gmail. com</mods:affiliation>
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<mods:namePart id="32873486B7EE06A46F7446C2E5ED0E17">Brown, Rafe M.</mods:namePart>
<mods:affiliation id="CDDB9AA2ECD89346CC99861D9A1E7C5A">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:date id="D9DC5E64B2D49E48732F3491F279E6CB">2019</mods:date>
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<treatment id="A20AE119FFADFFBDFDEC0B5D4C9A5A77" ID-DOI="http://doi.org/10.5281/zenodo.12728084" ID-GBIF-Taxon="235040677" ID-Zenodo-Dep="12728084" LSID="urn:lsid:plazi:treatment:A20AE119FFADFFBDFDEC0B5D4C9A5A77" httpUri="http://treatment.plazi.org/id/A20AE119FFADFFBDFDEC0B5D4C9A5A77" lastPageNumber="17" pageId="16" pageNumber="17">
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<taxonomicName id="EDA32B8CFFADFFBDFDEC0B5D4ADE5D87" authority="Figueroa &amp; McKelvy &amp; Grismer &amp; Bell &amp; Lailvaux, 2016" box="[572,725,380,403]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subFamily" subFamily="Ahaetulliinae">
<emphasis id="18D78C1DFFADFFBDFDEC0B5D4ADE5D87" bold="true" box="[572,725,380,403]" pageId="16" pageNumber="17">Ahaetulliinae</emphasis>
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<paragraph id="2A1C500FFFADFFBDFE2F0BBD4B195DA7" blockId="16.[511,786,380,435]" box="[511,786,412,435]" pageId="16" pageNumber="17">Key to Philippine genera.</paragraph>
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<key id="5E32073EFFADFFBDFFAB0BE64C9A5A77" pageId="16" pageNumber="17">
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<paragraph id="2A1C500FFFADFFBDFFAB0BE64C9A5DEA" blockId="16.[123,1174,455,574]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB0BE64C9A5DEA" pageId="16" pageNumber="17">
1a. Pupil horizontally elongated and rectangular (
<figureCitation id="B2984C8AFFADFFBDFD440BE64AF15DCA" box="[660,762,455,478]" captionStart="FIGURE" captionStartId="16.[155,167,746,765]" captionTargetBox="[293,1003,587,732]" captionTargetId="figure-449@16.[293,1003,587,732]" captionTargetPageId="16" captionText="FIGURE 25. Lateral view of head of (A) Ahaetulla prasina preocularis (KU 347854), (B) Dryophiops rubescens (KU 328516). Note horizontally elongate pupil in Ahaetulla versus rounded pupil in Dryophiops." figureDoi="http://doi.org/10.5281/zenodo.11512732" httpUri="https://zenodo.org/record/11512732/files/figure.png" pageId="16" pageNumber="17">Fig. 25A</figureCitation>
); ventral body scales laterally hinged, and their posterior edges unnotched (
<figureCitation id="B2984C8AFFADFFBDFDF60BC64A835DEA" box="[550,648,487,510]" captionStart="FIGURE" captionStartId="16.[155,167,1203,1222]" captionTargetBox="[270,1031,810,1185]" captionTargetId="figure-480@16.[265,1031,810,1190]" captionTargetPageId="16" captionText="FIGURE 26. Ventral body scales of (A) Ahaetulla prasina preocularis (KU 349849): ventral body scales hinged, with posterior edge unnotched, (B) Chrysopelea paradisi variabilis (KU 337271): ventral body scales hinged, and posterior edge notched." figureDoi="http://doi.org/10.5281/zenodo.11512734" httpUri="https://zenodo.org/record/11512734/files/figure.png" pageId="16" pageNumber="17">Fig. 26A</figureCitation>
)........................
<taxonomicName id="EDA32B8CFFADFFBDFC0A0BC64C495DEA" baseAuthorityName="Gaulke" baseAuthorityYear="1994" box="[986,1090,487,510]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFADFFBDFC0A0BC64C495DEA" box="[986,1090,487,510]" italics="true" pageId="16" pageNumber="17">Ahaetulla</emphasis>
</taxonomicName>
(p. 19)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFADFFBDFFAB08264C9A5E2A" blockId="16.[123,1174,455,574]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB08264C9A5E2A" pageId="16" pageNumber="17">
1b. Pupils may be circular or slightly horizontally elongate (
<figureCitation id="B2984C8AFFADFFBDFCD008264B695E0A" box="[768,866,519,542]" captionStart="FIGURE" captionStartId="16.[155,167,746,765]" captionTargetBox="[293,1003,587,732]" captionTargetId="figure-449@16.[293,1003,587,732]" captionTargetPageId="16" captionText="FIGURE 25. Lateral view of head of (A) Ahaetulla prasina preocularis (KU 347854), (B) Dryophiops rubescens (KU 328516). Note horizontally elongate pupil in Ahaetulla versus rounded pupil in Dryophiops." figureDoi="http://doi.org/10.5281/zenodo.11512732" httpUri="https://zenodo.org/record/11512732/files/figure.png" pageId="16" pageNumber="17">Fig. 25B</figureCitation>
); ventral body scales laterally hinged, and their posterior edges notched
<figureCitation id="B2984C8AFFADFFBDFDA608064ADE5E2A" box="[630,725,551,574]" captionStart="FIGURE" captionStartId="16.[155,167,1203,1222]" captionTargetBox="[270,1031,810,1185]" captionTargetId="figure-480@16.[265,1031,810,1190]" captionTargetPageId="16" captionText="FIGURE 26. Ventral body scales of (A) Ahaetulla prasina preocularis (KU 349849): ventral body scales hinged, with posterior edge unnotched, (B) Chrysopelea paradisi variabilis (KU 337271): ventral body scales hinged, and posterior edge notched." figureDoi="http://doi.org/10.5281/zenodo.11512734" httpUri="https://zenodo.org/record/11512734/files/figure.png" pageId="16" pageNumber="17">Fig. 26B</figureCitation>
)............................... 2
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<paragraph id="2A1C500FFFADFFBDFF4B08CB4BCB5F02" blockId="16.[123,1173,746,790]" pageId="16" pageNumber="17">
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25. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFADFFBDFDC108CB4B105EE9" box="[529,795,746,765]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="prasina" subSpecies="preocularis">
<emphasis id="18D78C1DFFADFFBDFDC108CB4B105EE9" box="[529,795,746,765]" italics="true" pageId="16" pageNumber="17">Ahaetulla prasina preocularis</emphasis>
</taxonomicName>
(KU 347854), (B)
<taxonomicName id="EDA32B8CFFADFFBDFC0108CB4C9E5EE9" baseAuthorityName="Gray" baseAuthorityYear="1834" box="[977,1173,746,765]" class="Reptilia" family="Colubridae" genus="Dryophiops" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="rubescens">
<emphasis id="18D78C1DFFADFFBDFC0108CB4C9E5EE9" box="[977,1173,746,765]" italics="true" pageId="16" pageNumber="17">Dryophiops rubescens</emphasis>
</taxonomicName>
(KU 328516). Note horizontally elongate pupil in
<taxonomicName id="EDA32B8CFFADFFBDFDFB09224A755F02" baseAuthorityName="Gaulke" baseAuthorityYear="1994" box="[555,638,771,790]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFADFFBDFDFB09224A755F02" box="[555,638,771,790]" italics="true" pageId="16" pageNumber="17">Ahaetulla</emphasis>
</taxonomicName>
versus rounded pupil in
<taxonomicName id="EDA32B8CFFADFFBDFC8509224BB65F02" authorityName="Boulenger" authorityYear="1896" box="[853,957,771,790]" class="Reptilia" family="Colubridae" genus="Dryophiops" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">Dryophiops</taxonomicName>
.
</paragraph>
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<paragraph id="2A1C500FFFADFFBDFF4B0E9248CE58ED" blockId="16.[123,1173,1203,1273]" pageId="16" pageNumber="17">
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26. Ventral body scales of (A)
<taxonomicName id="EDA32B8CFFADFFBDFE3F0E924AF858D2" box="[495,755,1203,1222]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="prasina" subSpecies="preocularis">
<emphasis id="18D78C1DFFADFFBDFE3F0E924AF858D2" box="[495,755,1203,1222]" italics="true" pageId="16" pageNumber="17">Ahaetulla prasina preocularis</emphasis>
</taxonomicName>
(KU 349849): ventral body scales hinged, with posterior edge unnotched, (B)
<taxonomicName id="EDA32B8CFFADFFBDFEAD0EEC4A8758F4" box="[381,652,1229,1248]" class="Reptilia" family="Colubridae" genus="Chrysopelea" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="paradisi" subSpecies="variabilis">
<emphasis id="18D78C1DFFADFFBDFEAD0EEC4A8758F4" box="[381,652,1229,1248]" italics="true" pageId="16" pageNumber="17">Chrysopelea paradisi variabilis</emphasis>
</taxonomicName>
(KU 337271): ventral body scales hinged, and posterior edge notched.
</paragraph>
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<keyStep id="915748AAFFADFFBDFFAB0F004C9A598C" pageId="16" pageNumber="17">
<paragraph id="2A1C500FFFADFFBDFFAB0F004C9A594C" blockId="16.[123,1173,1313,1432]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB0F004C9A594C" pageId="16" pageNumber="17">
2a. Dorsal body scales in 17 longitudinal rows at midbody; last ventral body scale (i.e., the scale immediately before the precloacal scale) divided.....................
<taxonomicName id="EDA32B8CFFADFFBDFC6B0F604C49594C" authorityName="H.Boie" authorityYear="1826" box="[955,1090,1345,1368]" class="Reptilia" family="Colubridae" genus="Chrysopelea" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFADFFBDFC6B0F604C49594C" box="[955,1090,1345,1368]" italics="true" pageId="16" pageNumber="17">Chrysopelea</emphasis>
</taxonomicName>
(p. 22)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFADFFBDFFAB0F404C9A598C" blockId="16.[123,1173,1313,1432]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB0F404C9A598C" pageId="16" pageNumber="17">2b. Dorsal body scales in 13–15 longitudinal rows at midbody; last ventral body scale undivided........................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFADFFBDFFAB0F8D4C9A5A77" pageId="16" pageNumber="17">
<paragraph id="2A1C500FFFADFFBDFFAB0F8D4C9A59F7" blockId="16.[123,1174,1452,1635]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB0F8D4C9A59F7" pageId="16" pageNumber="17">
3a. Dorsal body scales in 15 longitudinal rows at midbody; vertebral scales similar in size to other dorsal body scales; black postoocular stripe absent....................
<taxonomicName id="EDA32B8CFFADFFBDFC140FED4C4959F7" authorityName="Boulenger" authorityYear="1896" box="[964,1090,1484,1507]" class="Reptilia" family="Colubridae" genus="Dryophiops" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFADFFBDFC140FED4C4959F7" box="[964,1090,1484,1507]" italics="true" pageId="16" pageNumber="17">Dryophiops</emphasis>
</taxonomicName>
(p. 23)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFADFFBDFFAB0FCD4C9A5A77" blockId="16.[123,1174,1452,1635]" pageId="16" pageNumber="17">
<keyLead id="9152F33AFFADFFBDFFAB0FCD4C9A5A77" pageId="16" pageNumber="17">
3b. One of the following combinations of characters true: (1) dorsal body scales in 13 longitudinal rows at midbody, vertebral scales similar in size to other dorsal body scales; (2) dorsal body scales in 15 longitudinal rows at midbody, vertebral scales much larger than other dorsal body scales, and a black postocular stripe present........................
<taxonomicName id="EDA32B8CFFADFFBDFC620C6D4C495A77" authorityName="Boulenger" authorityYear="1890" box="[946,1090,1612,1635]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFADFFBDFC620C6D4C495A77" box="[946,1090,1612,1635]" italics="true" pageId="16" pageNumber="17">Dendrelaphis</emphasis>
</taxonomicName>
(p. 23)
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1a. Mental scale not in contact with anterior chin shields (
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1b. Mental scale in contact with anterior chin shields (
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31. Ventral view of head of (A)
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<emphasis id="18D78C1DFFA9FFB9FE220B9D4AC85DDB" box="[498,707,444,463]" italics="true" pageId="20" pageNumber="21">Calamaria palavanensis</emphasis>
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(KU 309445), (B)
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<emphasis id="18D78C1DFFA9FFB9FCBA0B9D4C135DDB" box="[874,1048,444,463]" italics="true" pageId="20" pageNumber="21">Calamaria gervaisii</emphasis>
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(KU 307983).
</paragraph>
<paragraph id="2A1C500FFFA9FFB9FFAB0BF44B7C5DFC" blockId="20.[123,1175,444,488]" box="[123,887,469,488]" pageId="20" pageNumber="21">Mental scale (blue), anterior chin shields (reddish brown), first infralabial scales (green).</paragraph>
</caption>
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2b. Number of pair of subcaudal scales&gt;40 (males),&gt; 30 (females)...........................................................
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<emphasis id="18D78C1DFFA9FFB9FDA9083A4AE65E26" box="[633,749,539,562]" italics="true" pageId="20" pageNumber="21">Calamaria</emphasis>
sp.
</taxonomicName>
Mindoro (Weinell et al.,
<taxonomicName id="EDA32B8CFFA9FFB9FBF4083A4C815E26" box="[1060,1162,539,562]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="41" phylum="Chordata" rank="species" species="review">in review</taxonomicName>
)
</keyLead>
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</keyStep>
<keyStep id="915748AAFFA9FFB9FFAB08674C9A5E69" pageId="20" pageNumber="21">
<paragraph id="2A1C500FFFA9FFB9FFAB08674C9A5E49" blockId="20.[123,1169,582,637]" box="[123,1169,582,605]" pageId="20" pageNumber="21">
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3a. Number of pairs of subcaudal scales ≤ 21 (males), ≤ 14 (females)......
<taxonomicName id="EDA32B8CFFA9FFB9FC6508674C9A5E49" ID-CoL="69752" box="[949,1169,582,605]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="virgulata">
<emphasis id="18D78C1DFFA9FFB9FC6508674C9A5E49" box="[949,1169,582,605]" italics="true" pageId="20" pageNumber="21">Calamaria virgulata</emphasis>
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<paragraph id="2A1C500FFFA9FFB9FFAB08474C9A5E69" blockId="20.[123,1169,582,637]" box="[123,1169,614,637]" pageId="20" pageNumber="21">
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3b. Number of pairs of subcaudal scales 23–25 (males), 16–20 (females)
<taxonomicName id="EDA32B8CFFA9FFB9FC5B08474C9A5E69" ID-CoL="884PJ" box="[907,1169,614,637]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="palavanensis">
<emphasis id="18D78C1DFFA9FFB9FC5B08474C9A5E69" box="[907,1169,614,637]" italics="true" pageId="20" pageNumber="21">Calamaria palavanensis</emphasis>
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<keyStep id="915748AAFFA9FFB9FFAB08B14C9A5ED3" pageId="20" pageNumber="21">
<paragraph id="2A1C500FFFA9FFB9FFAB08B14C9A5EB3" blockId="20.[123,1169,656,743]" box="[123,1169,656,679]" pageId="20" pageNumber="21">
<keyLead id="9152F33AFFA9FFB9FFAB08B14C9A5EB3" box="[123,1169,656,679]" pageId="20" pageNumber="21">
4a. Ventral surface of body with dark bands.......................
<taxonomicName id="EDA32B8CFFA9FFB9FC5808B14C9A5EB3" ID-CoL="PD5Y" box="[904,1169,656,679]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="18D78C1DFFA9FFB9FC5808B14C9A5EB3" box="[904,1169,656,679]" italics="true" pageId="20" pageNumber="21">Calamaria lumbricoidea</emphasis>
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<keyLead id="9152F33AFFA9FFB9FFAB08914C9A5ED3" box="[123,1169,688,711]" pageId="20" pageNumber="21">4b. Ventral surface of body without dark bands....................................... 5</keyLead>
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<paragraph id="2A1C500FFFA9FFB9FFAB08F14C9A5EF3" blockId="20.[123,1169,656,743]" box="[123,1169,720,743]" pageId="20" pageNumber="21">
<keyLead id="9152F33AFFA9FFB9FFAB08F14C9A5EF3" box="[123,1169,720,743]" pageId="20" pageNumber="21">
5a. Dorsal surface behind head has 2–6 dark, black-edged crossbands.....
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<emphasis id="18D78C1DFFA9FFB9FC6208F14C9A5EF3" box="[946,1169,720,743]" italics="true" pageId="20" pageNumber="21">Calamaria bitorques</emphasis>
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<keyLead id="9152F33AFFA9FFB9FFAB08DA4C9A5F06" box="[123,1169,763,786]" pageId="20" pageNumber="21">5b. Dorsal surface behind head lacks dark, black-edged crossbands....................... 6</keyLead>
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<paragraph id="2A1C500FFFA9FFB9FFAB09074C9A5F29" blockId="20.[123,1169,806,861]" box="[123,1169,806,829]" pageId="20" pageNumber="21">
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6a. Diameter of eye less than eye-mouth distance......................
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<emphasis id="18D78C1DFFA9FFB9FC6709074C9A5F29" box="[951,1169,806,829]" italics="true" pageId="20" pageNumber="21">Calamaria joloensis</emphasis>
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<keyLead id="9152F33AFFA9FFB9FFAB09674C9A5F49" box="[123,1169,838,861]" pageId="20" pageNumber="21">6b. Diameter of eye equal to or greater than eye-mouth distance......................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFA9FFB9FFAB09514C9A5FF3" pageId="20" pageNumber="21">
<paragraph id="2A1C500FFFA9FFB9FFAB09514C9A5FB3" blockId="20.[123,1173,880,999]" pageId="20" pageNumber="21">
<keyLead id="9152F33AFFA9FFB9FFAB09514C9A5FB3" pageId="20" pageNumber="21">
7a. Each dorsal body scale with a light network; a continuous light stripe on first row of dorsal body scales present............................................
<taxonomicName id="EDA32B8CFFA9FFB9FC6409B14C9A5FB3" ID-CoL="PD8P" authorityName="Taylor" authorityYear="1922" box="[948,1169,912,935]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="suluensis">
<emphasis id="18D78C1DFFA9FFB9FC6409B14C9A5FB3" box="[948,1169,912,935]" italics="true" pageId="20" pageNumber="21">Calamaria suluensis</emphasis>
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</paragraph>
<paragraph id="2A1C500FFFA9FFB9FFAB09914C9A5FF3" blockId="20.[123,1173,880,999]" pageId="20" pageNumber="21">
<keyLead id="9152F33AFFA9FFB9FFAB09914C9A5FF3" pageId="20" pageNumber="21">
7b. Each dorsal body scale above first row yellowish with a dark network; a dark-edged, interrupted, light stripe on first row of dorsal body scales usually present.....
<taxonomicName id="EDA32B8CFFA9FFB9FC6909F14C9A5FF3" ID-CoL="PD4K" box="[953,1169,976,999]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="gervaisii">
<emphasis id="18D78C1DFFA9FFB9FC6909F14C9A5FF3" box="[953,1169,976,999]" italics="true" pageId="20" pageNumber="21">Calamaria gervaisii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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</treatment>
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Represented in the
<collectingCountry id="52B4109FFFAEFFBEFE9A0B2749C95D09" box="[330,450,262,285]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
by a single species......................
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2a. Dorsal body scales in 23 longitudinal rows at midbody; three anterior temporal scales; eighth infralabial scale anterior to center of eye (
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2b. Dorsal body scales in 19 longitudinal rows at midbody; two anterior temporal scales; eighth infralabial scale posterior to center of eye (
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30. Ventral-lateral view of head of (A)
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<emphasis id="18D78C1DFFAEFFBEFDF809F44AA35FFC" box="[552,680,981,1000]" italics="true" pageId="19" pageNumber="20">Boiga cynodon</emphasis>
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(KU 328496), (B)
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<emphasis id="18D78C1DFFAEFFBEFC9B09F44BD05FFC" box="[843,987,981,1000]" italics="true" pageId="19" pageNumber="20">Boiga philippina</emphasis>
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(KU 304855). Eighth infralabial scale (purplish-gray).
</paragraph>
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3a. Number of preocular scales two; number of ventral body scales &lt;250.....
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<emphasis id="18D78C1DFFAEFFBEFC0C0E0A4C9A5856" box="[988,1169,1067,1090]" italics="true" pageId="19" pageNumber="20">Boiga philippina</emphasis>
</taxonomicName>
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<keyLead id="9152F33AFFAEFFBEFFAB0E6A4C9A5876" box="[123,1169,1099,1122]" pageId="19" pageNumber="20">3b. Number of preocular scales one; number of ventral body scales&gt; 250.................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFAEFFBEFFAB0E544C9A58B8" pageId="19" pageNumber="20">
<paragraph id="2A1C500FFFAEFFBEFFAB0E544C9A5898" blockId="19.[123,1169,1141,1196]" box="[123,1169,1141,1164]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0E544C9A5898" box="[123,1169,1141,1164]" pageId="19" pageNumber="20">
4a. Number of dark dorsal body crossbands ~40, each widening laterally.......
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<emphasis id="18D78C1DFFAEFFBEFC3B0E544C9A5898" box="[1003,1169,1141,1164]" italics="true" pageId="19" pageNumber="20">Boiga angulata</emphasis>
</taxonomicName>
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</paragraph>
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<keyLead id="9152F33AFFAEFFBEFFAB0EB44C9A58B8" box="[123,1169,1173,1196]" pageId="19" pageNumber="20">
4b. Number of dark dorsal body crossbands ~70, not widening laterally........
<taxonomicName id="EDA32B8CFFAEFFBEFC3D0EB44C9A58B8" ID-CoL="5WNHQ" authorityName="Taylor" authorityYear="1923" box="[1005,1169,1173,1196]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="schultzei">
<emphasis id="18D78C1DFFAEFFBEFC3D0EB44C9A58B8" box="[1005,1169,1173,1196]" italics="true" pageId="19" pageNumber="20">Boiga schultzei</emphasis>
</taxonomicName>
</keyLead>
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<paragraph id="2A1C500FFFAEFFBEFFAB0EE14C9A58E3" blockId="19.[123,1173,1216,1303]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0EE14C9A58E3" pageId="19" pageNumber="20">
5a. Dorsal color black, with most scales possessing a yellow or white speckle; yellow or white not arranged into distinct crossbands............................
<taxonomicName id="EDA32B8CFFAEFFBEFCA10EC14C9A58E3" ID-CoL="5DNMM" authorityName="Gaulke, Demegillo &amp; Vogel" authorityYear="2005" box="[881,1169,1248,1271]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="dendrophila" subSpecies="levitoni">
<emphasis id="18D78C1DFFAEFFBEFCA10EC14C9A58E3" box="[881,1169,1248,1271]" italics="true" pageId="19" pageNumber="20">Boiga dendrophila levitoni</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="2A1C500FFFAEFFBEFFAB0F214C9A5903" blockId="19.[123,1173,1216,1303]" box="[123,1169,1280,1303]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0F214C9A5903" box="[123,1169,1280,1303]" pageId="19" pageNumber="20">5b. Dorsal color black, with yellow or white crossbands on body and tail.................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFAEFFBEFFAB0F0A4C9A5996" pageId="19" pageNumber="20">
<paragraph id="2A1C500FFFAEFFBEFFAB0F0A4C9A5976" blockId="19.[123,1169,1323,1410]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0F0A4C9A5976" pageId="19" pageNumber="20">
6a. Yellow or white crossbands ≥ 2 scale rows wide, becoming wider laterally..................................................................
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<emphasis id="18D78C1DFFAEFFBEFC990F6A4C9A5976" box="[841,1169,1355,1378]" italics="true" pageId="19" pageNumber="20">Boiga dendrophila latifasciata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFAEFFBEFFAB0F4A4C9A5996" blockId="19.[123,1169,1323,1410]" box="[123,1169,1387,1410]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0F4A4C9A5996" box="[123,1169,1387,1410]" pageId="19" pageNumber="20">6b. Yellow or white crossbands &lt;2 scale rows wide that do not widen laterally............. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFAEFFBEFFAB0FB44C9A59F8" pageId="19" pageNumber="20">
<paragraph id="2A1C500FFFAEFFBEFFAB0FB44C9A59B8" blockId="19.[123,1173,1429,1516]" box="[123,1169,1429,1452]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0FB44C9A59B8" box="[123,1169,1429,1452]" pageId="19" pageNumber="20">
7a. Interspaces between narrow light crossbands black..........
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<emphasis id="18D78C1DFFAEFFBEFC990FB44C9A59B8" box="[841,1169,1429,1452]" italics="true" pageId="19" pageNumber="20">Boiga dendrophila multicincta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFAEFFBEFFAB0F944C9A59F8" blockId="19.[123,1173,1429,1516]" pageId="19" pageNumber="20">
<keyLead id="9152F33AFFAEFFBEFFAB0F944C9A59F8" pageId="19" pageNumber="20">
7b. Interspaces between narrow light crossbands, which are edged with black, bluish gray to bluish brown...............................................
<taxonomicName id="EDA32B8CFFAEFFBEFC8B0FF44C9A59F8" ID-CoL="5DNMJ" authorityName="Taylor" authorityYear="1922" box="[859,1169,1493,1516]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="dendrophila" subSpecies="divergens">
<emphasis id="18D78C1DFFAEFFBEFC8B0FF44C9A59F8" box="[859,1169,1493,1516]" italics="true" pageId="19" pageNumber="20">Boiga dendrophila divergens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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</treatment>
</document>

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<emphasis id="18D78C1DFFAFFFBFFD820FF04AB559FC" bold="true" box="[594,702,1489,1512]" italics="true" pageId="18" pageNumber="19">Ahaetulla</emphasis>
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<paragraph id="2A1C500FFFAFFFBFFE780FD04B625A1C" blockId="18.[424,873,1489,1544]" box="[424,873,1521,1544]" pageId="18" pageNumber="19">Key to Philippine species and subspecies.</paragraph>
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<key id="5E32073EFFAFFFBFFFAB0C3D4C9A5A8A" pageId="18" pageNumber="19">
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1a. Precloacal scale not divided; ≥ 2 preocular scales...........
<taxonomicName id="EDA32B8CFFAFFFBFFC9F0C3D4C9A5A27" ID-CoL="5DFKD" box="[847,1169,1564,1587]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="prasina" subSpecies="preocularis">
<emphasis id="18D78C1DFFAFFFBFFC9F0C3D4C9A5A27" box="[847,1169,1564,1587]" italics="true" pageId="18" pageNumber="19">Ahaetulla prasina preocularis</emphasis>
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<keyLead id="9152F33AFFAFFFBFFFAB0C1D4C9A5A47" box="[123,1169,1596,1619]" pageId="18" pageNumber="19">1b. Precloacal scale divided; one preocular scale...................................... 2</keyLead>
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2a. Usually two loreal scales; endemic to
<collectingRegion id="E8679EEDFFAFFFBFFDEC0C464A665A6A" box="[572,621,1639,1662]" country="Philippines" name="Sulu" pageId="18" pageNumber="19">Sulu</collectingRegion>
Archipelago........
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<emphasis id="18D78C1DFFAFFFBFFCB70C464C9A5A6A" box="[871,1169,1639,1662]" italics="true" pageId="18" pageNumber="19">Ahaetulla prasina suluensis</emphasis>
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2b. Usually one loreal scale; endemic to
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Represented in the
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by a single species..................
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1b. More than 10 supralabial scales.................................
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2a. One of the following combinations of characters true: (1) dorsal body scales in 15–17 longitudinal rows at midbody and background color of ventral body scales black; (2) dorsal body scales in 19 longitudinal rows at midbody and background color of ventral body scales pale (but not pinkish) in life................................................
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2b. One of the following combinations of characters true: (1) dorsal body scales in 17 longitudinal rows at midbody and background color of ventral body scales pale; (2) dorsal body scales in 19 longitudinal rows at midbody and background color of ventral body scales pinkish in life..............................................................
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Represented by a single genus......................................
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1a. Supralabial scales do not border eye; frontal scale fragmented into multiple smaller scales; internasal scales paired, positioned posterior to a pair of nasal scales; nasal scales in contact with each other medially (
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1b. One or more supralabial scales usually border eye; frontal scale not fragmented; internasal scale single, not paired, and is positioned medially between a pair of nasal scales; nasal scales not in contact with each other medially.....................................
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<mods:namePart id="F69BDBB2D8B1F06E5DECD9C5B785D746">Hooper, Errol</mods:namePart>
<mods:affiliation id="42B13B14E9167D1C0B480CDD88E790B9">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
</mods:name>
<mods:name id="506B5145BAE0328A7D6E065B856D3FAD" type="personal">
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<mods:namePart id="743D9739FF0F489842E17AAAC633D1FE">Leviton, Alan E.</mods:namePart>
<mods:affiliation id="389CEE18AC6FC0F6E0671A1C309F5705">Herpetology Division, Institute of Biodiversity Science &amp; Sustainability, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118. * Corresponding author; Email: jweine 2 @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="210B3460A413DD95C04AD3592AE118F9" type="personal">
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<mods:namePart id="FC7B134E85F82286D0C028359D43E388">Brown, Rafe M.</mods:namePart>
<mods:affiliation id="D7A2A5885A17487E80B45883B4AED7BE">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:typeOfResource id="F5EBF66BAB930F1DDC0FA6A1CB5C21D3">text</mods:typeOfResource>
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<mods:titleInfo id="30F5E9855A3509143646000F01CBB0B8">
<mods:title id="9E46E591C08B7CA772B340D3486A91E4">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="D2A3479E72FE2843A51612E9E410E0D2">
<mods:date id="781CDD5E2B827C180322C3192B018C82">2019</mods:date>
<mods:detail id="2B2036042CB1A12F99735DFCD0A74804" type="pubDate">
<mods:number id="3B44B7124AE4F5B6F4CBE75559CD8DE0">2019-09-30</mods:number>
</mods:detail>
<mods:detail id="AAB3708E9926BDE981CF4DAD06865A6E" type="volume">
<mods:number id="F328F7658D6D49C17DABBC2591BE7D4C">66</mods:number>
</mods:detail>
<mods:detail id="5898D513B2B9F1BD676C78A85DCC6887" type="issue">
<mods:number id="BACD2BDC6B1977755EF050F28484F22F">1</mods:number>
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<mods:extent id="E8A70730D6A188CFC3C1B521C89D9355" unit="page">
<mods:start id="70C97007AFDC5EFBB34502C3956C4DC0">1</mods:start>
<mods:end id="CCD4C548D146B49DBAF72F8B114187CF">49</mods:end>
</mods:extent>
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<mods:classification id="8D8B088C22CE98BA30BDF726C5F5DC55">journal article</mods:classification>
<mods:identifier id="DF7AC7D1E182F65E2873EBDF8A681263" type="CLB-Dataset">299708</mods:identifier>
<mods:identifier id="58F30CEE8EC90307EB454E0A270C2F57" type="DOI">10.5281/zenodo.11512680</mods:identifier>
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<mods:identifier id="FC41591E717D6747E4B4D3AF640C20E1" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="576869E9B6C4A6CC19E5E8D7AC3ADEE8" type="Zenodo-Dep">11512680</mods:identifier>
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<treatment id="A20AE119FFB0FFA3FDFF08C84C9A5FE0" ID-DOI="http://doi.org/10.5281/zenodo.12728070" ID-GBIF-Taxon="235040627" ID-Zenodo-Dep="12728070" LSID="urn:lsid:plazi:treatment:A20AE119FFB0FFA3FDFF08C84C9A5FE0" httpUri="http://treatment.plazi.org/id/A20AE119FFB0FFA3FDFF08C84C9A5FE0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="62B90384FFB0FFA0FDFF08C84AEB5F14" box="[559,736,745,768]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="2A1C500FFFB0FFA0FDFF08C84AEB5F14" blockId="13.[511,786,745,800]" box="[559,736,745,768]" pageId="13" pageNumber="14">
<taxonomicName id="EDA32B8CFFB0FFA0FDFF08C84AEB5F14" ID-CoL="BRL" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[559,736,745,768]" class="Reptilia" family="Lamprophiidae" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="family">
<emphasis id="18D78C1DFFB0FFA0FDFF08C84AEB5F14" bold="true" box="[559,736,745,768]" pageId="13" pageNumber="14">Lamprophiidae</emphasis>
</taxonomicName>
</paragraph>
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<subSubSection id="62B90384FFB0FFA0FE2F09284B195F34" box="[511,786,777,800]" pageId="13" pageNumber="14" type="multiple">
<paragraph id="2A1C500FFFB0FFA0FE2F09284B195F34" blockId="13.[511,786,745,800]" box="[511,786,777,800]" pageId="13" pageNumber="14">Key to Philippine genera.</paragraph>
</subSubSection>
<subSubSection id="62B90384FFB0FFA3FFAB09154C9A5FE0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="key">
<key id="5E32073EFFB0FFA3FFAB09154C9A5FE0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<keyStep id="915748AAFFB0FFA0FFAB09154C9A5F7F" pageId="13" pageNumber="14">
<paragraph id="2A1C500FFFB0FFA0FFAB09154C9A5F5F" blockId="13.[123,1174,820,1003]" box="[123,1169,820,843]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB09154C9A5F5F" box="[123,1169,820,843]" pageId="13" pageNumber="14">
1a. Nasal scale large, round and undivided (
<figureCitation id="B2984C8AFFB0FFA0FD8309154ABF5F5F" box="[595,692,820,843]" captionStart="FIGURE" captionStartId="10.[156,168,1348,1367]" captionTargetBox="[313,1004,1049,1338]" captionTargetId="figure-241@10.[294,1004,1049,1338]" captionText="FIGURE 13. Lateral view of head of (A) Psammodynastes pulverulentus (KU 329688), (B) Cyclocorus lineatus alcalai (KU 324539). Nasal scale (green) undivided (A) or divided (B)." figureDoi="http://doi.org/10.5281/zenodo.12725259" httpUri="https://zenodo.org/record/12725259/files/figure.png" pageId="13" pageNumber="14">Fig. 13A</figureCitation>
)...............
<taxonomicName id="EDA32B8CFFB0FFA0FC5A09154C495F5F" ID-CoL="6YB4" authorityName="Günther" authorityYear="1858" box="[906,1090,820,843]" class="Reptilia" family="Pseudaspididae" genus="Psammodynastes" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB0FFA0FC5A09154C495F5F" box="[906,1090,820,843]" italics="true" pageId="13" pageNumber="14">Psammodynastes</emphasis>
</taxonomicName>
(p. 31)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB0FFA0FFAB09754C9A5F7F" blockId="13.[123,1174,820,1003]" box="[123,1169,852,875]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB09754C9A5F7F" box="[123,1169,852,875]" pageId="13" pageNumber="14">
1b. Nasal scale divided (
<figureCitation id="B2984C8AFFB0FFA0FE53097549E85F7F" box="[387,483,852,875]" captionStart="FIGURE" captionStartId="10.[156,168,1348,1367]" captionTargetBox="[313,1004,1049,1338]" captionTargetId="figure-241@10.[294,1004,1049,1338]" captionText="FIGURE 13. Lateral view of head of (A) Psammodynastes pulverulentus (KU 329688), (B) Cyclocorus lineatus alcalai (KU 324539). Nasal scale (green) undivided (A) or divided (B)." figureDoi="http://doi.org/10.5281/zenodo.12725259" httpUri="https://zenodo.org/record/12725259/files/figure.png" pageId="13" pageNumber="14">Fig. 13B</figureCitation>
)................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFB0FFA0FFAB09554C9A5FFF" pageId="13" pageNumber="14">
<paragraph id="2A1C500FFFB0FFA0FFAB09554C9A5FBF" blockId="13.[123,1174,820,1003]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB09554C9A5FBF" pageId="13" pageNumber="14">
2a. Snout broad (
<figureCitation id="B2984C8AFFB0FFA0FEE7095549915F9F" box="[311,410,884,907]" captionStart="FIGURE" captionStartId="13.[155,167,1247,1266]" captionTargetBox="[311,995,1015,1230]" captionTargetId="figure-832@13.[293,1003,1015,1236]" captionTargetPageId="13" captionText="FIGURE 18. Dorsal head shape and scalation of (A) Oxyrhabdium modestum (KU 311301): narrow snout, (B) Hologerrhum philippinum (KU 330056): broad snout." figureDoi="http://doi.org/10.5281/zenodo.11512716" httpUri="https://zenodo.org/record/11512716/files/figure.png" pageId="13" pageNumber="14">Fig. 18B</figureCitation>
); anterior chin shields smaller than or similar in size to posterior chin shields; subcaudal scales unpaired (
<figureCitation id="B2984C8AFFB0FFA0FDC609B54A7D5FBF" box="[534,630,916,939]" captionStart="FIGURE" captionStartId="10.[156,168,1676,1695]" captionTargetBox="[316,980,1403,1659]" captionTargetId="figure-269@10.[293,1003,1400,1667]" captionText="FIGURE 14. Arrangement of subcaudal scales, (A) Opisthotropis typica (KU 327424): subcaudal scales paired; (B) Cyclocorus lineatus alcalai (KU 324539): subcaudal scales single, not paired." figureDoi="http://doi.org/10.5281/zenodo.12725261" httpUri="https://zenodo.org/record/12725261/files/figure.png" pageId="13" pageNumber="14">Fig. 14B</figureCitation>
)...................................... 3
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB0FFA0FFAB09954C9A5FFF" blockId="13.[123,1174,820,1003]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB09954C9A5FFF" pageId="13" pageNumber="14">
2b. Snout narrow (
<figureCitation id="B2984C8AFFB0FFA0FE86099549CB5FDF" box="[342,448,948,971]" captionStart="FIGURE" captionStartId="13.[155,167,1247,1266]" captionTargetBox="[311,995,1015,1230]" captionTargetId="figure-832@13.[293,1003,1015,1236]" captionTargetPageId="13" captionText="FIGURE 18. Dorsal head shape and scalation of (A) Oxyrhabdium modestum (KU 311301): narrow snout, (B) Hologerrhum philippinum (KU 330056): broad snout." figureDoi="http://doi.org/10.5281/zenodo.11512716" httpUri="https://zenodo.org/record/11512716/files/figure.png" pageId="13" pageNumber="14">Fig. 18A</figureCitation>
); anterior chin shields much larger than posterior chin shields; subcaudal scales paired or unpaired (
<figureCitation id="B2984C8AFFB0FFA0FDF509F54A795FFF" box="[549,626,980,1003]" captionStart="FIGURE" captionStartId="10.[156,168,1676,1695]" captionTargetBox="[316,980,1403,1659]" captionTargetId="figure-269@10.[293,1003,1400,1667]" captionText="FIGURE 14. Arrangement of subcaudal scales, (A) Opisthotropis typica (KU 327424): subcaudal scales paired; (B) Cyclocorus lineatus alcalai (KU 324539): subcaudal scales single, not paired." figureDoi="http://doi.org/10.5281/zenodo.12725261" httpUri="https://zenodo.org/record/12725261/files/figure.png" pageId="13" pageNumber="14">Fig. 14</figureCitation>
)....................................... 4
</keyLead>
</paragraph>
</keyStep>
<caption id="7EDC0087FFB0FFA0FF4B0EFE4A0F591F" ID-DOI="http://doi.org/10.5281/zenodo.11512716" ID-Zenodo-Dep="11512716" httpUri="https://zenodo.org/record/11512716/files/figure.png" pageId="13" pageNumber="14" startId="13.[155,167,1247,1266]" targetBox="[311,995,1015,1230]" targetPageId="13" targetType="figure">
<paragraph id="2A1C500FFFB0FFA0FF4B0EFE4A0F591F" blockId="13.[123,1173,1247,1291]" pageId="13" pageNumber="14">
<smallCapsWord id="2CFAC6D3FFB0FFA0FF4B0EFE48D358E4" baselines="1261,1260" box="[155,216,1247,1266]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="13" pageNumber="14">FIGURE</smallCapsWord>
18. Dorsal head shape and scalation of (A)
<taxonomicName id="EDA32B8CFFB0FFA0FD840EFE4B2D58E6" baseAuthorityName="Duméril" baseAuthorityYear="1853" box="[596,806,1247,1266]" class="Reptilia" family="Cyclocoridae" genus="Oxyrhabdium" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="modestum">
<emphasis id="18D78C1DFFB0FFA0FD840EFE4B2D58E6" box="[596,806,1247,1266]" italics="true" pageId="13" pageNumber="14">Oxyrhabdium modestum</emphasis>
</taxonomicName>
(KU 311301): narrow snout, (B)
<taxonomicName id="EDA32B8CFFB0FFA0FB970EFE491F591F" authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Cyclocoridae" genus="Hologerrhum" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="philippinum">
<emphasis id="18D78C1DFFB0FFA0FB970EFE491F591F" italics="true" pageId="13" pageNumber="14">Hologerrhum philippinum</emphasis>
</taxonomicName>
(KU 330056): broad snout.
</paragraph>
</caption>
<keyStep id="915748AAFFB0FFA0FFAB0F0F4C9A5A71" pageId="13" pageNumber="14">
<paragraph id="2A1C500FFFB0FFA0FFAB0F0F4C9A59B1" blockId="13.[123,1174,1326,1637]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB0F0F4C9A59B1" pageId="13" pageNumber="14">
3a. Maxilla not strongly angled; no maxillary diastema, although anterior and posterior maxillary teeth are much larger than intervening teeth; posterior maxillary tooth grooved; five infralabial scales in contact with chin shields, and fourth infralabial scale broadly in contact with the anterior and posterior chin shields for about an equal length (
<figureCitation id="B2984C8AFFB0FFA0FD2D0FAF4B5559B1" box="[765,862,1422,1445]" captionStart="FIGURE" captionStartId="14.[154,166,451,470]" captionTargetBox="[124,1166,216,439]" captionTargetId="figure-209@14.[119,1174,216,439]" captionTargetPageId="14" captionText="FIGURE 19. Ventral view of head of (A) Hologerrhum philippinum (KU 330065): five infralabial scales in contact with chin shields; anterior chin shields broadly contact infralabial scales 1–4; posterior chin shields broadly in contact with fourth and fifth infralabial scales; (B) Cyclocorus lineatus lineatus (KU 346571): five infralabial scales in contact with chin shields; anterior chin shields broadly in contact with infralabial scales 1–4; posterior chin shields broadly in contact with fifth infralabial scale; (C) Cyclocorus nuchalis nuchalis (KU 327765): four infralabial scales in contact with chin shields, anterior chin shields broadly in contact with infralabial scales 1–3; posterior chin shields broadly in contact with fourth infralabial scale." figureDoi="http://doi.org/10.5281/zenodo.11512718" httpUri="https://zenodo.org/record/11512718/files/figure.png" pageId="13" pageNumber="14">Fig. 19A</figureCitation>
).....
<taxonomicName id="EDA32B8CFFB0FFA0FC610FAF4C4959B1" ID-CoL="4XYW" box="[945,1090,1422,1445]" class="Reptilia" family="Cyclocoridae" genus="Hologerrhum" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB0FFA0FC610FAF4C4959B1" box="[945,1090,1422,1445]" italics="true" pageId="13" pageNumber="14">Hologerrhum</emphasis>
</taxonomicName>
(p. 24)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB0FFA0FFAB0F8F4C9A5A71" blockId="13.[123,1174,1326,1637]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB0F8F4C9A5A71" pageId="13" pageNumber="14">
3b. Maxilla strongly angled; large diastema between anterior and posterior maxillary teeth; posterior maxillary teeth not grooved; one of the following is also true: (1) five infralabial scales in contact with chin shields, and fourth infralabial scale either not in contact with or barely in contact with posterior chin shield (
<figureCitation id="B2984C8AFFB0FFA0FE270C2F4A505A31" box="[503,603,1550,1573]" captionStart="FIGURE" captionStartId="14.[154,166,451,470]" captionTargetBox="[124,1166,216,439]" captionTargetId="figure-209@14.[119,1174,216,439]" captionTargetPageId="14" captionText="FIGURE 19. Ventral view of head of (A) Hologerrhum philippinum (KU 330065): five infralabial scales in contact with chin shields; anterior chin shields broadly contact infralabial scales 1–4; posterior chin shields broadly in contact with fourth and fifth infralabial scales; (B) Cyclocorus lineatus lineatus (KU 346571): five infralabial scales in contact with chin shields; anterior chin shields broadly in contact with infralabial scales 1–4; posterior chin shields broadly in contact with fifth infralabial scale; (C) Cyclocorus nuchalis nuchalis (KU 327765): four infralabial scales in contact with chin shields, anterior chin shields broadly in contact with infralabial scales 1–3; posterior chin shields broadly in contact with fourth infralabial scale." figureDoi="http://doi.org/10.5281/zenodo.11512718" httpUri="https://zenodo.org/record/11512718/files/figure.png" pageId="13" pageNumber="14">Fig. 19B</figureCitation>
), or (2) four infralabial scales in contact with chin shields, and third infralabial scale either not in contact with or barely in contact with posterior chin shield (
<figureCitation id="B2984C8AFFB0FFA0FECC0C6F49775A71" box="[284,380,1614,1637]" captionStart="FIGURE" captionStartId="14.[154,166,451,470]" captionTargetBox="[124,1166,216,439]" captionTargetId="figure-209@14.[119,1174,216,439]" captionTargetPageId="14" captionText="FIGURE 19. Ventral view of head of (A) Hologerrhum philippinum (KU 330065): five infralabial scales in contact with chin shields; anterior chin shields broadly contact infralabial scales 1–4; posterior chin shields broadly in contact with fourth and fifth infralabial scales; (B) Cyclocorus lineatus lineatus (KU 346571): five infralabial scales in contact with chin shields; anterior chin shields broadly in contact with infralabial scales 1–4; posterior chin shields broadly in contact with fifth infralabial scale; (C) Cyclocorus nuchalis nuchalis (KU 327765): four infralabial scales in contact with chin shields, anterior chin shields broadly in contact with infralabial scales 1–3; posterior chin shields broadly in contact with fourth infralabial scale." figureDoi="http://doi.org/10.5281/zenodo.11512718" httpUri="https://zenodo.org/record/11512718/files/figure.png" pageId="13" pageNumber="14">Fig. 19C</figureCitation>
)...........................................
<taxonomicName id="EDA32B8CFFB0FFA0FC190C6F4C495A71" ID-CoL="3XW7" authorityName="A.M.C.Duméril" authorityYear="1853" box="[969,1090,1614,1637]" class="Reptilia" family="Cyclocoridae" genus="Cyclocorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB0FFA0FC190C6F4C495A71" box="[969,1090,1614,1637]" italics="true" pageId="13" pageNumber="14">Cyclocorus</emphasis>
</taxonomicName>
(p. 22)
</keyLead>
</paragraph>
</keyStep>
<paragraph id="2A1C500FFFB0FFA0FFAB0C584C9A5A84" blockId="13.[123,1169,1657,1712]" box="[123,1169,1657,1680]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB0C584C9A5A84" box="[123,1169,1657,1680]" pageId="13" pageNumber="14">
<keyStep id="915748AAFFB0FFA0FFAB0C584C9A5A84" box="[123,1169,1657,1680]" pageId="13" pageNumber="14">4a. Five supralabial scales; subcaudal scales unpaired................ Unnamed genus (p. 35)</keyStep>
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB0FFA3FFAB0CB84C9A5FC0" blockId="13.[123,1169,1657,1712]" lastBlockId="14.[123,1169,957,1012]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA3FFAB0CB84C9A5FC0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
4b. Six or more supralabial scales; subcaudal scales paired.............................. 5
<keyStep id="915748AAFFB3FFA3FFAB099C4C9A5FC0" box="[123,1169,957,980]" pageId="14" pageNumber="15">
5a. Parietal scales in contact with supralabial scales (
<figureCitation id="B2984C8AFFB3FFA3FD64099C4B1D5FC0" box="[692,790,957,980]" captionStart="FIGURE" captionStartId="14.[153,165,877,896]" captionTargetBox="[290,1000,641,862]" captionTargetId="figure-319@14.[290,1000,641,874]" captionTargetPageId="14" captionText="FIGURE 20. Lateral view of head of (A) Myersophis alpestris (KU 308684), (B) Oxyrhabdium modestum (KU 311301). Parietal scale (blue), temporal scales (green), posterior supralabial scales (purplish-gray)." figureDoi="http://doi.org/10.5281/zenodo.11512720" httpUri="https://zenodo.org/record/11512720/files/figure.png" pageId="14" pageNumber="15">Fig. 20A</figureCitation>
)............
<taxonomicName id="EDA32B8CFFB3FFA3FC16099C4C495FC0" ID-CoL="5W96" authorityName="Taylor" authorityYear="1963" box="[966,1090,957,980]" class="Reptilia" family="Cyclocoridae" genus="Myersophis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB3FFA3FC16099C4C495FC0" box="[966,1090,957,980]" italics="true" pageId="14" pageNumber="15">Myersophis</emphasis>
</taxonomicName>
(p. 30)
</keyStep>
</keyLead>
</paragraph>
<caption id="7EDC0087FFB3FFA3FF4A0BE249035E64" ID-DOI="http://doi.org/10.5281/zenodo.11512718" ID-Zenodo-Dep="11512718" httpUri="https://zenodo.org/record/11512718/files/figure.png" pageId="14" pageNumber="15" startId="14.[154,166,451,470]" targetBox="[124,1166,216,439]" targetPageId="14" targetType="figure">
<paragraph id="2A1C500FFFB3FFA3FF4A0BE249035E64" blockId="14.[121,1172,451,624]" pageId="14" pageNumber="15">
<smallCapsWord id="2CFAC6D3FFB3FFA3FF4A0BE248DC5DC1" baselines="465,465" box="[154,215,451,470]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="14" pageNumber="15">FIGURE</smallCapsWord>
19. Ventral view of head of (A)
<taxonomicName id="EDA32B8CFFB3FFA3FE200BE24ADA5DC2" authorityName="Gunther" authorityYear="1858" box="[496,721,451,470]" class="Reptilia" family="Cyclocoridae" genus="Hologerrhum" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="philippinum">
<emphasis id="18D78C1DFFB3FFA3FE200BE24ADA5DC2" box="[496,721,451,470]" italics="true" pageId="14" pageNumber="15">Hologerrhum philippinum</emphasis>
</taxonomicName>
(KU 330065): five infralabial scales in contact with chin shields; anterior chin shields broadly contact infralabial scales 1–4; posterior chin shields broadly in contact with fourth and fifth infralabial scales; (B)
<taxonomicName id="EDA32B8CFFB3FFA3FE6A0BD64AA75E1E" baseAuthorityName="Reinhardt" baseAuthorityYear="1843" box="[442,684,503,522]" class="Reptilia" family="Cyclocoridae" genus="Cyclocorus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="lineatus" subSpecies="lineatus">
<emphasis id="18D78C1DFFB3FFA3FE6A0BD64AA75E1E" box="[442,684,503,522]" italics="true" pageId="14" pageNumber="15">Cyclocorus lineatus lineatus</emphasis>
</taxonomicName>
(KU 346571): five infralabial scales in contact with chin shields; anterior chin shields broadly in contact with infralabial scales 1–4; posterior chin shields broadly in contact with fifth infralabial scale; (C)
<taxonomicName id="EDA32B8CFFB3FFA3FE8C080B4A515E29" authorityName="Taylor" authorityYear="1923" box="[348,602,554,573]" class="Reptilia" family="Cyclocoridae" genus="Cyclocorus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="nuchalis" subSpecies="nuchalis">
<emphasis id="18D78C1DFFB3FFA3FE8C080B4A515E29" box="[348,602,554,573]" italics="true" pageId="14" pageNumber="15">Cyclocorus nuchalis nuchalis</emphasis>
</taxonomicName>
(KU 327765): four infralabial scales in contact with chin shields, anterior chin shields broadly in contact with infralabial scales 1–3; posterior chin shields broadly in contact with fourth infralabial scale.
</paragraph>
</caption>
<caption id="7EDC0087FFB3FFA3FF49094C4B7C5F8E" ID-DOI="http://doi.org/10.5281/zenodo.11512720" ID-Zenodo-Dep="11512720" httpUri="https://zenodo.org/record/11512720/files/figure.png" pageId="14" pageNumber="15" startId="14.[153,165,877,896]" targetBox="[290,1000,641,862]" targetPageId="14" targetType="figure">
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<smallCapsWord id="2CFAC6D3FFB3FFA3FF49094C48DD5F6B" baselines="891,891" box="[153,214,877,896]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="14" pageNumber="15">FIGURE</smallCapsWord>
20. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFB3FFA3FE3D094C4A945F94" authorityName="Taylor" authorityYear="1963" box="[493,671,877,896]" class="Reptilia" family="Cyclocoridae" genus="Myersophis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alpestris">
<emphasis id="18D78C1DFFB3FFA3FE3D094C4A945F94" box="[493,671,877,896]" italics="true" pageId="14" pageNumber="15">Myersophis alpestris</emphasis>
</taxonomicName>
(KU 308684), (B)
<taxonomicName id="EDA32B8CFFB3FFA3FC95094C4C1D5F94" baseAuthorityName="Duméril" baseAuthorityYear="1853" box="[837,1046,877,896]" class="Reptilia" family="Cyclocoridae" genus="Oxyrhabdium" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="modestum">
<emphasis id="18D78C1DFFB3FFA3FC95094C4C1D5F94" box="[837,1046,877,896]" italics="true" pageId="14" pageNumber="15">Oxyrhabdium modestum</emphasis>
</taxonomicName>
(KU 311301).
</paragraph>
<paragraph id="2A1C500FFFB3FFA3FFA809A64B7C5F8E" blockId="14.[120,1172,877,922]" box="[120,887,903,922]" pageId="14" pageNumber="15">Parietal scale (blue), temporal scales (green), posterior supralabial scales (purplish-gray).</paragraph>
</caption>
<paragraph id="2A1C500FFFB3FFA3FFAB09FC4C9A5FE0" blockId="14.[123,1169,957,1012]" box="[123,1169,989,1012]" pageId="14" pageNumber="15">
<keyLead id="9152F33AFFB3FFA3FFAB09FC4C9A5FE0" box="[123,1169,989,1012]" pageId="14" pageNumber="15">
5b. Parietal scales not in contact with supralabial scales (
<figureCitation id="B2984C8AFFB3FFA3FD0F09FC4B345FE0" box="[735,831,989,1012]" captionStart="FIGURE" captionStartId="14.[153,165,877,896]" captionTargetBox="[290,1000,641,862]" captionTargetId="figure-319@14.[290,1000,641,874]" captionTargetPageId="14" captionText="FIGURE 20. Lateral view of head of (A) Myersophis alpestris (KU 308684), (B) Oxyrhabdium modestum (KU 311301). Parietal scale (blue), temporal scales (green), posterior supralabial scales (purplish-gray)." figureDoi="http://doi.org/10.5281/zenodo.11512720" httpUri="https://zenodo.org/record/11512720/files/figure.png" pageId="14" pageNumber="15">Fig. 20B</figureCitation>
).......
<taxonomicName id="EDA32B8CFFB3FFA3FC6009FC4C495FE0" ID-CoL="6BGK" authorityName="Boulenger" authorityYear="1893" box="[944,1090,989,1012]" class="Reptilia" family="Cyclocoridae" genus="Oxyrhabdium" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB3FFA3FC6009FC4C495FE0" box="[944,1090,989,1012]" italics="true" pageId="14" pageNumber="15">Oxyrhabdium</emphasis>
</taxonomicName>
(p. 31)
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(terrestrial species)
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<key id="5E32073EFFB1FFA0FFAB09F74C9A5D47" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<paragraph id="2A1C500FFFB1FFA1FFAB09F74C9A5819" blockId="12.[123,1174,982,1266]" pageId="12" pageNumber="13">
<keyLead id="9152F33AFFB1FFA1FFAB09F74C9A5819" pageId="12" pageNumber="13">
1a. Three postocular scales (
<figureCitation id="B2984C8AFFB1FFA1FE1409F74A465FF9" box="[452,589,982,1005]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="12" pageNumber="13">Fig. 17A–B</figureCitation>
); lateral body scales in oblique or horizontal rows (
<figureCitation id="B2984C8AFFB1FFA1FF4F09D748E65819" box="[159,237,1014,1037]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="12" pageNumber="13">Fig. 15</figureCitation>
).................................................................... 2
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB1FFA1FFAB0E374C9A5859" blockId="12.[123,1174,982,1266]" pageId="12" pageNumber="13">
<keyLead id="9152F33AFFB1FFA1FFAB0E374C9A5859" pageId="12" pageNumber="13">
1b. Two postocular scales (
<figureCitation id="B2984C8AFFB1FFA1FE4C0E374A145839" box="[412,543,1046,1069]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="12" pageNumber="13">Fig. 17C–D</figureCitation>
); lateral body scales in horizontal rows (
<figureCitation id="B2984C8AFFB1FFA1FC1F0E374C245839" box="[975,1071,1046,1069]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="12" pageNumber="13">Fig. 15B</figureCitation>
); number of longitudinal rows of dorsal body scales not reducing posteriorly..................... 3
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<keyStep id="915748AAFFB1FFA1FFAB0E7A4C9A58E6" pageId="12" pageNumber="13">
<paragraph id="2A1C500FFFB1FFA1FFAB0E7A4C9A5886" blockId="12.[123,1174,982,1266]" pageId="12" pageNumber="13">
<keyLead id="9152F33AFFB1FFA1FFAB0E7A4C9A5886" pageId="12" pageNumber="13">
2a. Dorsal body scales in 17–25 longitudinal rows at midbody; postnasal scale vertically elongate, separated from or only narrowly in contact with prefrontal scale (
<figureCitation id="B2984C8AFFB1FFA1FC870E5A4BB35886" box="[855,952,1147,1170]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="12" pageNumber="13">Fig. 17A</figureCitation>
).....
<taxonomicName id="EDA32B8CFFB1FFA1FBDE0E5A4C495886" baseAuthorityName="Wuster and Thorpe" baseAuthorityYear="1996" box="[1038,1090,1147,1170]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB1FFA1FBDE0E5A4C495886" box="[1038,1090,1147,1170]" italics="true" pageId="12" pageNumber="13">Naja</emphasis>
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(p. 30)
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<paragraph id="2A1C500FFFB1FFA1FFAB0EBA4C9A58E6" blockId="12.[123,1174,982,1266]" pageId="12" pageNumber="13">
<keyLead id="9152F33AFFB1FFA1FFAB0EBA4C9A58E6" pageId="12" pageNumber="13">
2b. Dorsal body scales in 15 longitudinal rows throughout length of body; postnasal scale triangular, about as long as tall, and broadly in contact with prefrontal scale (
<figureCitation id="B2984C8AFFB1FFA1FC580E9A4BEC58C6" box="[904,999,1211,1234]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="12" pageNumber="13">Fig. 17B</figureCitation>
).......................................................................
<taxonomicName id="EDA32B8CFFB1FFA1FC620EFA4C4958E6" authorityName="Günther" authorityYear="1864" box="[946,1090,1243,1266]" class="Reptilia" family="Elapidae" genus="Ophiophagus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB1FFA1FC620EFA4C4958E6" box="[946,1090,1243,1266]" italics="true" pageId="12" pageNumber="13">Ophiophagus</emphasis>
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(p. 31)
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17. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFB1FFA1FDDD0CAE4A945AB6" authorityName="Peters" authorityYear="1861" box="[525,671,1679,1698]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="samarensis">
<emphasis id="18D78C1DFFB1FFA1FDDD0CAE4A945AB6" box="[525,671,1679,1698]" italics="true" pageId="12" pageNumber="13">Naja samarensis</emphasis>
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(KU 326653), (B)
<taxonomicName id="EDA32B8CFFB1FFA1FC840CAE4C1B5AB6" baseAuthorityName="Cantor" baseAuthorityYear="1836" box="[852,1040,1679,1698]" class="Reptilia" family="Elapidae" genus="Ophiophagus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hannah">
<emphasis id="18D78C1DFFB1FFA1FC840CAE4C1B5AB6" box="[852,1040,1679,1698]" italics="true" pageId="12" pageNumber="13">Ophiophagus hannah</emphasis>
</taxonomicName>
(KU 321813), (C)
<taxonomicName id="EDA32B8CFFB1FFA1FF4F0C89497E5AAF" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[159,373,1704,1723]" class="Reptilia" family="Elapidae" genus="Hemibungarus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="mcclungi">
<emphasis id="18D78C1DFFB1FFA1FF4F0C89497E5AAF" box="[159,373,1704,1723]" italics="true" pageId="12" pageNumber="13">Hemibungarus mcclungi</emphasis>
</taxonomicName>
(KU 313898), (D)
<taxonomicName id="EDA32B8CFFB1FFA1FDF60C894AD45AAF" box="[550,735,1704,1723]" class="Reptilia" family="Elapidae" genus="Calliophis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="philippina">
<emphasis id="18D78C1DFFB1FFA1FDF60C894AD45AAF" box="[550,735,1704,1723]" italics="true" pageId="12" pageNumber="13">Calliophis philippina</emphasis>
</taxonomicName>
(KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown).
</paragraph>
</caption>
<keyStep id="915748AAFFB0FFA0FFAB0AFD4C9A5D47" pageId="13" pageNumber="14">
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<keyLead id="9152F33AFFB0FFA0FFAB0AFD4C9A5D07" pageId="13" pageNumber="14">
3a. Two anterior temporal scales (
<figureCitation id="B2984C8AFFB0FFA0FE330AFD4A4E5CE7" box="[483,581,220,243]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="13" pageNumber="14">Fig. 17C</figureCitation>
); dorsal body scales in 15 longitudinal rows throughout length of body..............................................
<taxonomicName id="EDA32B8CFFB0FFA0FC730ADD4C495D07" box="[931,1090,252,275]" class="Reptilia" family="Elapidae" genus="Hemibungarus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB0FFA0FC730ADD4C495D07" box="[931,1090,252,275]" italics="true" pageId="13" pageNumber="14">Hemibungarus</emphasis>
</taxonomicName>
(p. 24)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB0FFA0FFAB0B3D4C9A5D47" blockId="13.[123,1173,220,339]" pageId="13" pageNumber="14">
<keyLead id="9152F33AFFB0FFA0FFAB0B3D4C9A5D47" pageId="13" pageNumber="14">
3b. One anterior temporal scale (
<figureCitation id="B2984C8AFFB0FFA0FE0C0B3D4A345D27" box="[476,575,284,307]" captionStart="FIGURE" captionStartId="12.[155,167,1679,1698]" captionTargetBox="[293,1003,1273,1669]" captionTargetId="figure-711@12.[293,1003,1273,1669]" captionTargetPageId="12" captionText="FIGURE 17. Lateral view of head of (A) Naja samarensis (KU 326653), (B) Ophiophagus hannah (KU 321813), (C) Hemibungarus mcclungi (KU 313898), (D) Calliophis philippina (KU 327218). Postocular scales (green), anterior temporal scales (purplish-gray), postnasal scales (blue), prefrontal scales (reddish brown)." figureDoi="http://doi.org/10.5281/zenodo.11512714" httpUri="https://zenodo.org/record/11512714/files/figure.png" pageId="13" pageNumber="14">Fig. 17D</figureCitation>
); dorsal body scales in 13 longitudinal rows throughout length of body..................................................
<taxonomicName id="EDA32B8CFFB0FFA0FC030B1D4C495D47" authorityName="Gray" authorityYear="1835" box="[979,1090,316,339]" class="Reptilia" family="Elapidae" genus="Calliophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB0FFA0FC030B1D4C495D47" box="[979,1090,316,339]" italics="true" pageId="13" pageNumber="14">Calliophis</emphasis>
</taxonomicName>
(p. 21)
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<mods:title id="07212224D54F59CC5B2EC7BE47FAFA25">Illustrated Key to the Snakes of the Philippines</mods:title>
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<treatment id="A20AE119FFB1FFA1FE250AFA4C9A5E5C" ID-DOI="http://doi.org/10.5281/zenodo.12728062" ID-GBIF-Taxon="235040623" ID-Zenodo-Dep="12728062" LSID="urn:lsid:plazi:treatment:A20AE119FFB1FFA1FE250AFA4C9A5E5C" httpUri="http://treatment.plazi.org/id/A20AE119FFB1FFA1FE250AFA4C9A5E5C" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="62B90384FFB1FFA1FE250AFA4B115CE6" box="[501,794,219,242]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="2A1C500FFFB1FFA1FE250AFA4B115CE6" blockId="12.[501,794,219,275]" box="[501,794,219,242]" pageId="12" pageNumber="13">
<emphasis id="18D78C1DFFB1FFA1FE250AFA4B115CE6" bold="true" box="[501,794,219,242]" pageId="12" pageNumber="13">
<taxonomicName id="EDA32B8CFFB1FFA1FE250AFA4A515CE6" ID-CoL="9LQ" authority="Boie, 1827" authorityName="F.Boie" authorityYear="1827" box="[501,602,219,242]" class="Reptilia" family="Elapidae" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="family">Elapidae</taxonomicName>
(marine species)
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<paragraph id="2A1C500FFFB1FFA1FE2F0ADD4B195D07" blockId="12.[501,794,219,275]" box="[511,786,252,275]" pageId="12" pageNumber="13">Key to Philippine genera.</paragraph>
</subSubSection>
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<key id="5E32073EFFB1FFA1FFAB0B074C9A5E5C" pageId="12" pageNumber="13">
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1a. Nasal scales separated by internasal scales; width of ventral body scales&gt; 1/3 body width.................................................................
<taxonomicName id="EDA32B8CFFB1FFA1FC060B674C495D49" baseAuthorityName="Kharin" baseAuthorityYear="2005" box="[982,1090,326,349]" class="Reptilia" family="Elapidae" genus="Laticauda" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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(p. 27)
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<keyLead id="9152F33AFFB1FFA1FFAB0B474C9A5D89" pageId="12" pageNumber="13">1b. Internasal scales absent, nasal scales in contact with each other medially; width of ventral body scales variable............................................................... 2</keyLead>
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2a. Width of ventral body scales&gt; 1/3 body width........................
<taxonomicName id="EDA32B8CFFB1FFA1FC070B8A4C495DD6" authorityName="Lacepède" authorityYear="1804" box="[983,1090,427,450]" class="Reptilia" family="Elapidae" genus="Aipysurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB1FFA1FC070B8A4C495DD6" box="[983,1090,427,450]" italics="true" pageId="12" pageNumber="13">Aipysurus</emphasis>
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(p. 20)
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<keyLead id="9152F33AFFB1FFA1FFAB0BEA4C9A5E16" pageId="12" pageNumber="13">2b. Width of ventral body scales &lt;1/4 body width and often narrower than adjacent lateral body scales...................................................................... 3</keyLead>
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<keyLead id="9152F33AFFB1FFA1FFAB08304C9A5E3C" box="[123,1169,529,552]" pageId="12" pageNumber="13">
3a. Three supralabial scales, second very elongate (
<figureCitation id="B2984C8AFFB1FFA1FD7308304AFB5E3C" box="[675,752,529,552]" captionStart="FIGURE" captionStartId="12.[155,167,820,839]" captionTargetBox="[486,810,590,814]" captionTargetId="figure-659@12.[486,810,589,816]" captionText="FIGURE 16. Lateral view of head of Emydocephalus annulatus (specimen number unknown; a redrafting by EH of original image in Rasmussen et al. 2011). Second supralabial scale (reddish brown). Fig. 16, modified original line drawing reproduced with permission of A. Rasmussen and Zootaxa and Magnolia Press." figureDoi="http://doi.org/10.5281/zenodo.12725263" httpUri="https://zenodo.org/record/12725263/files/figure.png" pageId="12" pageNumber="13">Fig. 16</figureCitation>
)............
<taxonomicName id="EDA32B8CFFB1FFA1FC4908304C495E3C" authorityName="Krefft" authorityYear="1869" box="[921,1090,529,552]" class="Reptilia" family="Elapidae" genus="Emydocephalus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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(p. 24)
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3b. More than three supralabial scales.................................
<taxonomicName id="EDA32B8CFFB1FFA1FC0208104C495E5C" authorityName="Latreille" authorityYear="1801" box="[978,1090,561,584]" class="Reptilia" family="Elapidae" genus="Hydrophis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB1FFA1FC0208104C495E5C" box="[978,1090,561,584]" italics="true" pageId="12" pageNumber="13">Hydrophis</emphasis>
</taxonomicName>
(p. 24)
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1a. Some scales on head strongly keeled (
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); second supralabial scale not in contact with scale forming anterior border of heat sensing pit...................
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1b. Scales on head smooth (
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); second supralabial scale in contact with scale forming anterior border of heat sensing pit...................................
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1a. Posterior origin of nasal suture is preocular scale (
<figureCitation id="B2984C8AFFB3FFA3FD6E0F344B2B5938" box="[702,800,1301,1324]" captionStart="FIGURE" captionStartId="14.[157,169,1617,1636]" captionTargetBox="[294,1004,1365,1602]" captionTargetId="figure-348@14.[294,1004,1365,1602]" captionTargetPageId="14" captionText="FIGURE 21. Lateral view of head of (A) Indotyphlops braminus (specimen number unknown; redrawn from Taylor 1922a), (B) Ramphotyphlops cumingii (EHT R-99; a redrafting by EH of original image in Taylor 1919). Second supral- abial scale (purplish-gray); preocular scale (green); nspo = nasal suture posterior origin scale." figureDoi="http://doi.org/10.5281/zenodo.11512722" httpUri="https://zenodo.org/record/11512722/files/figure.png" pageId="14" pageNumber="15">Fig. 21A</figureCitation>
)..........
<taxonomicName id="EDA32B8CFFB3FFA3FC670F344C495938" ID-CoL="54D6" authorityName="Hedges, Marion, Lipp, Marin &amp; Vidal" authorityYear="2014" box="[951,1090,1301,1324]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB3FFA3FC670F344C495938" box="[951,1090,1301,1324]" italics="true" pageId="14" pageNumber="15">Indotyphlops</emphasis>
</taxonomicName>
(p. 27)
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</paragraph>
<paragraph id="2A1C500FFFB3FFA2FFAB0F144C9A5847" blockId="14.[123,1169,1301,1356]" lastBlockId="15.[123,1169,1084,1139]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<keyLead id="9152F33AFFB3FFA2FFAB0F144C9A5847" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
1b. Posterior origin of nasal suture is the second supralabial scale (
<figureCitation id="B2984C8AFFB3FFA3FC9A0F144BA25958" box="[842,937,1333,1356]" captionStart="FIGURE" captionStartId="14.[157,169,1617,1636]" captionTargetBox="[294,1004,1365,1602]" captionTargetId="figure-348@14.[294,1004,1365,1602]" captionTargetPageId="14" captionText="FIGURE 21. Lateral view of head of (A) Indotyphlops braminus (specimen number unknown; redrawn from Taylor 1922a), (B) Ramphotyphlops cumingii (EHT R-99; a redrafting by EH of original image in Taylor 1919). Second supral- abial scale (purplish-gray); preocular scale (green); nspo = nasal suture posterior origin scale." figureDoi="http://doi.org/10.5281/zenodo.11512722" httpUri="https://zenodo.org/record/11512722/files/figure.png" pageId="14" pageNumber="15">Fig. 21B</figureCitation>
)............... 2
<keyStep id="915748AAFFB2FFA2FFAB0E1D4C9A5847" box="[123,1169,1084,1107]" pageId="15" pageNumber="16">
2a. Prefrontal scale much wider than rostral scale (
<figureCitation id="B2984C8AFFB2FFA2FD4C0E1D4AF55847" box="[668,766,1084,1107]" captionStart="FIGURE" captionStartId="15.[155,167,516,535]" captionTargetBox="[139,1141,216,503]" captionTargetId="figure-390@15.[120,1176,216,503]" captionTargetPageId="15" captionText="FIGURE 22. Dorsal view of head of (A) Acutotyphlops banaorum (FMNH 262249; a redrafting by EH of original image in Wallach et al. 2007), (B) Malayotyphlops ruber (SMF 16616; a redrafting by EH of original image in Wynn et al. 2016), (C) Ramphotyphlops suluensis (PBS 2001; a redrafting by EH of original image in Taylor 1918). Prefrontal scale (green), rostral scale (purplish-gray). Fig. A reproduced and modified with permission of V. Wallach and Journal of Herpetology. Fig. B reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512724" httpUri="https://zenodo.org/record/11512724/files/figure.png" pageId="15" pageNumber="16">Fig. 22A</figureCitation>
)............
<taxonomicName id="EDA32B8CFFB2FFA2FC790E1D4C495847" ID-CoL="NW8" box="[937,1090,1084,1107]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB2FFA2FC790E1D4C495847" box="[937,1090,1084,1107]" italics="true" pageId="15" pageNumber="16">Acutotyphlops</emphasis>
</taxonomicName>
(p. 19)
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</paragraph>
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21. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFB3FFA3FDD60C704AC45A70" baseAuthorityName="Daudin" baseAuthorityYear="1803" box="[518,719,1617,1636]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="braminus">
<emphasis id="18D78C1DFFB3FFA3FDD60C704AC45A70" box="[518,719,1617,1636]" italics="true" pageId="14" pageNumber="15">Indotyphlops braminus</emphasis>
</taxonomicName>
(specimen number unknown; redrawn from
<bibRefCitation id="4E322DFEFFB3FFA3FB8E0C7048B85A6A" author="Taylor" firstAuthor="Taylor" pageId="14" pageNumber="15" refId="ref35611" refString="TAYLOR, EDWARD H. 1922 a. The Snakes of the Philippine Islands. Bureau of Science, Manila, Philippines. 312 pp., 32 text-figs., 63 tables, 37 pls." type="book" year="1922" yearSuffix="a">Taylor 1922a</bibRefCitation>
), (B)
<taxonomicName id="EDA32B8CFFB3FFA3FF360C4A49CC5A6A" baseAuthorityName="Gray" baseAuthorityYear="1845" box="[230,455,1643,1662]" class="Reptilia" family="Typhlopidae" genus="Ramphotyphlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="18D78C1DFFB3FFA3FF360C4A49CC5A6A" box="[230,455,1643,1662]" italics="true" pageId="14" pageNumber="15">Ramphotyphlops cumingii</emphasis>
</taxonomicName>
(EHT R-99; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB3FFA3FC490C4A4C0E5A6A" author="Taylor" box="[921,1029,1643,1662]" firstAuthor="Taylor" pageId="14" pageNumber="15" pagination="105 - 125" refId="ref35585" refString="TAYLOR, EDWARD H. 1919. New or rare Philippine reptiles. Philippine Journal of Science 14 (1): 105 - 125." type="journal article" year="1919">Taylor 1919</bibRefCitation>
). Second supralabial scale (purplish-gray); preocular scale (green); nspo = nasal suture posterior origin scale.
</paragraph>
</caption>
<caption id="7EDC0087FFB2FFA2FF4B08254B745E69" ID-DOI="http://doi.org/10.5281/zenodo.11512724" ID-Zenodo-Dep="11512724" httpUri="https://zenodo.org/record/11512724/files/figure.png" pageId="15" pageNumber="16" startId="15.[155,167,516,535]" targetBox="[139,1141,216,503]" targetPageId="15" targetType="figure">
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22. Dorsal view of head of (A)
<taxonomicName id="EDA32B8CFFB2FFA2FE3A08254ACA5E03" box="[490,705,516,535]" class="Reptilia" family="Typhlopidae" genus="Acutotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="banaorum">
<emphasis id="18D78C1DFFB2FFA2FE3A08254ACA5E03" box="[490,705,516,535]" italics="true" pageId="15" pageNumber="16">Acutotyphlops banaorum</emphasis>
</taxonomicName>
(FMNH 262249; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB2FFA2FF40083C49335E24" author="Wallach" box="[144,312,541,560]" etAl="et al." firstAuthor="Wallach" pageId="15" pageNumber="16" pagination="690 - 702" refId="ref36078" refString="WALLACH, VAN, RAFE M. BROWN, A. C. DIESMOS, AND G. V. A. GEE. 2007. An enigmatic new species of blind snake from Luzon Island, northern Philippines, with a synopsis of the genus Acutotyphlops (Serpentes: Typhlopidae). Journal of Herpetology 41 (4): 690 - 702, 3 figs., 1 table." type="book chapter" year="2007">Wallach et al. 2007</bibRefCitation>
), (B)
<taxonomicName id="EDA32B8CFFB2FFA2FEB8083C4A2D5E24" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[360,550,541,560]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="ruber">
<emphasis id="18D78C1DFFB2FFA2FEB8083C4A2D5E24" box="[360,550,541,560]" italics="true" pageId="15" pageNumber="16">Malayotyphlops ruber</emphasis>
</taxonomicName>
(SMF 16616; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB2FFA2FC26083C4C805E24" author="Wynn" box="[1014,1163,541,560]" etAl="et al." firstAuthor="Wynn" pageId="15" pageNumber="16" pagination="157 - 168" refId="ref36443" refString="WYNN, ADDISON H., ARVIN C. DIESMOS, AND RAFE M. BROWN. 2016. Two new species of Malayotyphlops from the northern Philippines, with redescriptions of Malayotyphlops luzonensis (Taylor) and Malayotyphlops ruber (Boettger). Journal of Herpetology 50 (1): 157 - 168." type="journal article" year="2016">Wynn et al. 2016</bibRefCitation>
), (C)
<taxonomicName id="EDA32B8CFFB2FFA2FF4D0816498A5E5E" authorityName=", PBS" authorityYear="2001" box="[157,385,567,586]" class="Reptilia" family="Typhlopidae" genus="Ramphotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="suluensis">
<emphasis id="18D78C1DFFB2FFA2FF4D0816498A5E5E" box="[157,385,567,586]" italics="true" pageId="15" pageNumber="16">Ramphotyphlops suluensis</emphasis>
</taxonomicName>
(PBS 2001; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB2FFA2FC9A08164BBD5E5E" author="Taylor" box="[842,950,567,586]" firstAuthor="Taylor" pageId="15" pageNumber="16" pagination="233 - 267" refId="ref35559" refString="TAYLOR, EDWARD H. 1918. Reptiles of the Sulu Archipelago. Philippine Journal of Science 13 (15): 233 - 267." type="journal article" year="1918">Taylor 1918</bibRefCitation>
). Prefrontal scale (green), rostral scale (purplish-gray). Fig. A reproduced and modified with permission of V. Wallach and
<emphasis id="18D78C1DFFB2FFA2FC1508714C845E77" box="[965,1167,592,611]" italics="true" pageId="15" pageNumber="16">Journal of Herpetology</emphasis>
. Fig. B reproduced and modified with permission of A. Wynn and
<emphasis id="18D78C1DFFB2FFA2FD61084B4B705E69" box="[689,891,618,637]" italics="true" pageId="15" pageNumber="16">Journal of Herpetology</emphasis>
.
</paragraph>
</caption>
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23. Lateral view of head of (A)
<taxonomicName id="EDA32B8CFFB2FFA2FE3F09864ABD5FAE" baseAuthorityName="Wallach" baseAuthorityYear="1993" box="[495,694,935,954]" class="Reptilia" family="Typhlopidae" genus="Ramphotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="marxi">
<emphasis id="18D78C1DFFB2FFA2FE3F09864ABD5FAE" box="[495,694,935,954]" italics="true" pageId="15" pageNumber="16">Ramphotyphlops marxi</emphasis>
</taxonomicName>
(FMNH 96520; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB2FFA2FFAB09E048FE5FC0" author="Wallach" box="[123,245,961,980]" firstAuthor="Wallach" pageId="15" pageNumber="16" pagination="263 - 278" refId="ref36033" refString="WALLACH, VAN. 1993. A new species of blind snake, Typhlops marxi, from the Philippines (Serpentes: Typhlopidae). Raffles Bulletin of Zoology 41 (2): 263 - 278, 2 figs., 2 tables." type="book chapter" year="1993">Wallach 1993</bibRefCitation>
), (B)
<taxonomicName id="EDA32B8CFFB2FFA2FEFC09E04A1D5FC0" baseAuthorityName="Taylor" baseAuthorityYear="1919" box="[300,534,961,980]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="luzonensis">
<emphasis id="18D78C1DFFB2FFA2FEFC09E04A1D5FC0" box="[300,534,961,980]" italics="true" pageId="15" pageNumber="16">Malayotyphlops luzonensis</emphasis>
</taxonomicName>
(CM 2653; a redrafting by EH of original image in
<bibRefCitation id="4E322DFEFFB2FFA2FC3F09E04C875FC0" author="Wynn" box="[1007,1164,961,980]" etAl="et al." firstAuthor="Wynn" pageId="15" pageNumber="16" pagination="157 - 168" refId="ref36443" refString="WYNN, ADDISON H., ARVIN C. DIESMOS, AND RAFE M. BROWN. 2016. Two new species of Malayotyphlops from the northern Philippines, with redescriptions of Malayotyphlops luzonensis (Taylor) and Malayotyphlops ruber (Boettger). Journal of Herpetology 50 (1): 157 - 168." type="journal article" year="2016">Wynn et al. 2016</bibRefCitation>
). Rostral scale (blue); arrow indicates either (A) beaked shape of rostral scale, or (B) rounded shape of rostral scale. Fig. A reproduced and modified with permission of V. Wallach and
<emphasis id="18D78C1DFFB2FFA2FD5609D54B445813" box="[646,847,1012,1031]" italics="true" pageId="15" pageNumber="16">Journal of Herpetology</emphasis>
. Fig. B reproduced and modified with permission of A. Wynn and Journal of Herpetology.
</paragraph>
</caption>
<paragraph id="2A1C500FFFB2FFA2FFAB0E7D4C9A5867" blockId="15.[123,1169,1084,1139]" box="[123,1169,1116,1139]" pageId="15" pageNumber="16">
<keyLead id="9152F33AFFB2FFA2FFAB0E7D4C9A5867" box="[123,1169,1116,1139]" pageId="15" pageNumber="16">
2b. Prefrontal scale narrower than rostral scale (
<figureCitation id="B2984C8AFFB2FFA2FDAF0E7D4AF45867" box="[639,767,1116,1139]" captionStart="FIGURE" captionStartId="15.[155,167,516,535]" captionTargetBox="[139,1141,216,503]" captionTargetId="figure-390@15.[120,1176,216,503]" captionTargetPageId="15" captionText="FIGURE 22. Dorsal view of head of (A) Acutotyphlops banaorum (FMNH 262249; a redrafting by EH of original image in Wallach et al. 2007), (B) Malayotyphlops ruber (SMF 16616; a redrafting by EH of original image in Wynn et al. 2016), (C) Ramphotyphlops suluensis (PBS 2001; a redrafting by EH of original image in Taylor 1918). Prefrontal scale (green), rostral scale (purplish-gray). Fig. A reproduced and modified with permission of V. Wallach and Journal of Herpetology. Fig. B reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512724" httpUri="https://zenodo.org/record/11512724/files/figure.png" pageId="15" pageNumber="16">Fig. 22B–C</figureCitation>
)............................ 3
</keyLead>
</paragraph>
<keyStep id="915748AAFFB2FFA2FFAB0EA74C9A58A9" pageId="15" pageNumber="16">
<paragraph id="2A1C500FFFB2FFA2FFAB0EA74C9A5889" blockId="15.[123,1169,1158,1213]" box="[123,1169,1158,1181]" pageId="15" pageNumber="16">
<keyLead id="9152F33AFFB2FFA2FFAB0EA74C9A5889" box="[123,1169,1158,1181]" pageId="15" pageNumber="16">
3a. Rostral scale beaked (
<figureCitation id="B2984C8AFFB2FFA2FE5E0EA749E45889" box="[398,495,1158,1181]" captionStart="FIGURE" captionStartId="15.[155,167,935,954]" captionTargetBox="[307,1002,660,922]" captionTargetId="figure-269@15.[293,1002,660,922]" captionTargetPageId="15" captionText="FIGURE 23. Lateral view of head of (A) Ramphotyphlops marxi (FMNH 96520; a redrafting by EH of original image in Wallach 1993), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016). Rostral scale (blue); arrow indicates either (A) beaked shape of rostral scale, or (B) rounded shape of rostral scale. Fig. A reproduced and modified with permission of V. Wallach and Journal of Herpetology. Fig. B reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512726" httpUri="https://zenodo.org/record/11512726/files/figure.png" pageId="15" pageNumber="16">Fig. 23A</figureCitation>
)..............................
<taxonomicName id="EDA32B8CFFB2FFA2FC5F0EA74C495889" ID-CoL="76ZL" authorityName="Fitzinger" authorityYear="1843" box="[911,1090,1158,1181]" class="Reptilia" family="Typhlopidae" genus="Ramphotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB2FFA2FC5F0EA74C495889" box="[911,1090,1158,1181]" italics="true" pageId="15" pageNumber="16">Ramphotyphlops</emphasis>
</taxonomicName>
(p. 33)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB2FFA2FFAB0E874C9A58A9" blockId="15.[123,1169,1158,1213]" box="[123,1169,1190,1213]" pageId="15" pageNumber="16">
<keyLead id="9152F33AFFB2FFA2FFAB0E874C9A58A9" box="[123,1169,1190,1213]" pageId="15" pageNumber="16">
3b. Rostral scale rounded (
<figureCitation id="B2984C8AFFB2FFA2FE4B0E8749F058A9" box="[411,507,1190,1213]" captionStart="FIGURE" captionStartId="15.[155,167,935,954]" captionTargetBox="[307,1002,660,922]" captionTargetId="figure-269@15.[293,1002,660,922]" captionTargetPageId="15" captionText="FIGURE 23. Lateral view of head of (A) Ramphotyphlops marxi (FMNH 96520; a redrafting by EH of original image in Wallach 1993), (B) Malayotyphlops luzonensis (CM 2653; a redrafting by EH of original image in Wynn et al. 2016). Rostral scale (blue); arrow indicates either (A) beaked shape of rostral scale, or (B) rounded shape of rostral scale. Fig. A reproduced and modified with permission of V. Wallach and Journal of Herpetology. Fig. B reproduced and modified with permission of A. Wynn and Journal of Herpetology." figureDoi="http://doi.org/10.5281/zenodo.11512726" httpUri="https://zenodo.org/record/11512726/files/figure.png" pageId="15" pageNumber="16">Fig. 23B</figureCitation>
)..............................
<taxonomicName id="EDA32B8CFFB2FFA2FC460E874C4958A9" ID-CoL="5KHG" authorityName="Hedges, Marion, Lipp, Marin &amp; Vidal" authorityYear="2014" box="[918,1090,1190,1213]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB2FFA2FC460E874C4958A9" box="[918,1090,1190,1213]" italics="true" pageId="15" pageNumber="16">Malayotyphlops</emphasis>
</taxonomicName>
(p. 29)
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<paragraph id="2A1C500FFFB6FFA6FE35082E4B275E32" blockId="11.[485,812,495,550]" box="[485,812,527,550]" pageId="11" pageNumber="12">Key to Philippine subfamilies.</paragraph>
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<key id="5E32073EFFB6FFA6FFAB081B4C9A5AB2" pageId="11" pageNumber="12">
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<keyLead id="9152F33AFFB6FFA6FFAB081B4C9A5E65" pageId="11" pageNumber="12">
1a. Temporal scales absent; parietal scales in contact with supralabial scales (
<figureCitation id="B2984C8AFFB6FFA6FC69081B4C105E45" box="[953,1051,570,593]" captionStart="FIGURE" captionStartId="10.[157,169,993,1012]" captionTargetBox="[294,1004,793,980]" captionTargetId="figure-213@10.[294,1004,792,980]" captionTargetPageId="10" captionText="FIGURE 12. Lateral view of head of (A) Calamaria gervaisii (KU 322329), (B) Lycodon muelleri (KU 327575). Temporal scales (green), supralabial scales (purplish-gray), parietal scale (blue)." figureDoi="http://doi.org/10.5281/zenodo.12725257" httpUri="https://zenodo.org/record/12725257/files/figure.png" pageId="11" pageNumber="12">Fig. 12A</figureCitation>
)................................................................... Calamariinae (p. 18)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB085B4C9A5E85" blockId="11.[123,1169,570,657]" box="[123,1169,634,657]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB085B4C9A5E85" box="[123,1169,634,657]" pageId="11" pageNumber="12">
1b. Temporal scales present; parietal scales not in contact with supralabial scales (
<figureCitation id="B2984C8AFFB6FFA6FC3D085B4C465E85" box="[1005,1101,634,657]" captionStart="FIGURE" captionStartId="10.[157,169,993,1012]" captionTargetBox="[294,1004,793,980]" captionTargetId="figure-213@10.[294,1004,792,980]" captionTargetPageId="10" captionText="FIGURE 12. Lateral view of head of (A) Calamaria gervaisii (KU 322329), (B) Lycodon muelleri (KU 327575). Temporal scales (green), supralabial scales (purplish-gray), parietal scale (blue)." figureDoi="http://doi.org/10.5281/zenodo.12725257" httpUri="https://zenodo.org/record/12725257/files/figure.png" pageId="11" pageNumber="12">Fig. 12B</figureCitation>
)... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFB6FFA6FFAB08854C9A5F6F" pageId="11" pageNumber="12">
<paragraph id="2A1C500FFFB6FFA6FFAB08854C9A5F0F" blockId="11.[123,1174,676,891]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB08854C9A5F0F" pageId="11" pageNumber="12">
2a. Supralabial scales either do not border eye, or supralabial scales 4–6 border eye; if supralabial scales 4–6 border eye, then dorsal body scales reduce from 19→17 or from 17→15 longitudinal rows posteriorly, and lateral body scales (except possibly first row) keeled (
<figureCitation id="B2984C8AFFB6FFA6FC1508C54C1C5EEF" box="[965,1047,740,763]" captionStart="FIGURE" captionStartId="7.[155,167,853,872]" captionTargetBox="[119,1176,587,845]" captionTargetId="figure-735@7.[119,1176,587,845]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal body scales: (A) spinose (Acrochordus granulatus; KU 302951), (B) keeled (Opisthotropis typica; KU 327424), (C) smooth (Oligodon maculatus; KU 321699)." figureDoi="http://doi.org/10.5281/zenodo.11512690" httpUri="https://zenodo.org/record/11512690/files/figure.png" pageId="11" pageNumber="12">Fig. 3B</figureCitation>
)...................................................................... Natricinae (p. 19)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB09054C9A5F6F" blockId="11.[123,1174,676,891]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB09054C9A5F6F" pageId="11" pageNumber="12">
2b. One or more supralabial scales border eye; if supralabial scales 4–6 border eye and dorsal body scales reduce from 19→17 or 17→15 longitudinal rows posteriorly, then lateral body scales are smooth (
<figureCitation id="B2984C8AFFB6FFA6FE71094549F85F6F" box="[417,499,868,891]" captionStart="FIGURE" captionStartId="7.[155,167,853,872]" captionTargetBox="[119,1176,587,845]" captionTargetId="figure-735@7.[119,1176,587,845]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal body scales: (A) spinose (Acrochordus granulatus; KU 302951), (B) keeled (Opisthotropis typica; KU 327424), (C) smooth (Oligodon maculatus; KU 321699)." figureDoi="http://doi.org/10.5281/zenodo.11512690" httpUri="https://zenodo.org/record/11512690/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
)................................................ 3
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<paragraph id="2A1C500FFFB6FFA6FFAB09AE4C9A5FF2" blockId="11.[123,1174,911,1190]" pageId="11" pageNumber="12">
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3a. Lateral body scales in oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FD8009AE4AB35FB2" box="[592,696,911,934]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A</figureCitation>
); dorsal body scales reduce from 17→13, 15→13, 15→11, or 13→11 longitudinal scale rows posteriorly; third supralabial scale does not border eye................................................... Ahaetulliinae (p. 17)
</keyLead>
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<paragraph id="2A1C500FFFB6FFA6FFAB09CE4C9A58B2" blockId="11.[123,1174,911,1190]" pageId="11" pageNumber="12">
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3b. One of the following combinations of characters true: (1) lateral body scales in horizontal, not oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FEE20E2E49985832" box="[306,403,1039,1062]" captionStart="FIGURE" captionStartId="10.[156,168,1348,1367]" captionTargetBox="[313,1004,1049,1338]" captionTargetId="figure-241@10.[294,1004,1049,1338]" captionText="FIGURE 13. Lateral view of head of (A) Psammodynastes pulverulentus (KU 329688), (B) Cyclocorus lineatus alcalai (KU 324539). Nasal scale (green) undivided (A) or divided (B)." figureDoi="http://doi.org/10.5281/zenodo.12725259" httpUri="https://zenodo.org/record/12725259/files/figure.png" pageId="11" pageNumber="12">Fig. 13B</figureCitation>
), and dorsal body scales in 13 longitudinal rows at midbody; (2) lateral body scales in oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FE0B0E0E4A355852" box="[475,574,1071,1094]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A</figureCitation>
) and dorsal body scales in 15–17 longitudinal rows at midbody, not reducing to fewer than 15 rows before vent; (3) lateral body scales in oblique or horizontal rows (compare
<figureCitation id="B2984C8AFFB6FFA6FE670E4E4A375892" box="[439,572,1135,1158]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A, B</figureCitation>
), and dorsal body scales in 19–23 longitudinal rows at midbody.................................................................... 4
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15. Lateral body scales of (A)
<taxonomicName id="EDA32B8CFFB6FFA6FE230FAA4AB3598A" authorityName="Vogel &amp; Van Rooijen" authorityYear="2008" box="[499,696,1419,1438]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="marenae">
<emphasis id="18D78C1DFFB6FFA6FE230FAA4AB3598A" box="[499,696,1419,1438]" italics="true" pageId="11" pageNumber="12">Dendrelaphis marenae</emphasis>
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(KU 314131): oblique rows, (B)
<taxonomicName id="EDA32B8CFFB6FFA6FC350FAA4C9D598A" baseAuthorityName="Taylor" baseAuthorityYear="1918" box="[997,1174,1419,1438]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="18D78C1DFFB6FFA6FC350FAA4C9D598A" box="[997,1174,1419,1438]" italics="true" pageId="11" pageNumber="12">Oligodon maculatus</emphasis>
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(KU 321699): horizontal rows.
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<keyStep id="915748AAFFB6FFA6FFAB0FEE4C9A5AB2" pageId="11" pageNumber="12">
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<keyLead id="9152F33AFFB6FFA6FFAB0FEE4C9A5A32" pageId="11" pageNumber="12">4a. Dorsal body scales in 17 smooth longitudinal rows throughout length of body; number of ventral body scales 144–185; subcaudal scales arranged in 89–145 pairs; numerous small teeth, 30–50, on each of maxillary and dentary bones.................... Sibynophiinae (p. 19)</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB0C0E4C9A5AB2" blockId="11.[123,1173,1487,1702]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB0C0E4C9A5AB2" pageId="11" pageNumber="12">4b. Dorsal body scales in 15–23 smooth or keeled longitudinal rows at midbody; if dorsal body scales in 17 smooth longitudinal rows throughout length of body, then subcaudal scales arranged in &lt;80 or&gt; 190 pairs; fewer than 30 teeth on each of maxillary and dentary bones..................................................................... Colubrinae (p. 18)</keyLead>
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<treatment id="A20AE119FFB6FFA6FDE30BB34C9A5DC7" ID-DOI="http://doi.org/10.5281/zenodo.12728052" ID-GBIF-Taxon="235040631" ID-Zenodo-Dep="12728052" LSID="urn:lsid:plazi:treatment:A20AE119FFB6FFA6FDE30BB34C9A5DC7" httpUri="http://treatment.plazi.org/id/A20AE119FFB6FFA6FDE30BB34C9A5DC7" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="62B90384FFB6FFA6FDE30BB34AD65DBD" box="[563,733,402,425]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="2A1C500FFFB6FFA6FDE30BB34AD65DBD" blockId="11.[563,733,402,425]" box="[563,733,402,425]" pageId="11" pageNumber="12">
<heading id="7154E763FFB6FFA6FDE30BB34AD65DBD" bold="true" box="[563,733,402,425]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="8">
<taxonomicName id="EDA32B8CFFB6FFA6FDE30BB34AD65DBD" ID-CoL="5YH" authority="Bonaparte, 1831" authorityName="Bonaparte" authorityYear="1831" box="[563,733,402,425]" class="Reptilia" family="Acrochordidae" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="family">
<emphasis id="18D78C1DFFB6FFA6FDE30BB34AD65DBD" bold="true" box="[563,733,402,425]" pageId="11" pageNumber="12">Acrochordidae</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62B90384FFB6FFA6FFAB0B9D4C9A5DC7" box="[123,1169,444,467]" pageId="11" pageNumber="12" type="taxon_list">
<paragraph id="2A1C500FFFB6FFA6FFAB0B9D4C9A5DC7" blockId="11.[123,1169,444,467]" box="[123,1169,444,467]" pageId="11" pageNumber="12">
Represented by a single genus.....................................
<taxonomicName id="EDA32B8CFFB6FFA6FC6A0B9D4C495DC7" authorityName="Hornstedt" authorityYear="1787" box="[954,1090,444,467]" class="Reptilia" family="Acrochordidae" genus="Acrochordus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="18D78C1DFFB6FFA6FC6A0B9D4C495DC7" box="[954,1090,444,467]" italics="true" pageId="11" pageNumber="12">Acrochordus</emphasis>
</taxonomicName>
(p. 19)
</paragraph>
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</treatment>
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