Add updates up until 2024-07-11 22:45:03

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<document id="BD845AC2EDDF75A1725B46E4E07980EF" ID-CLB-Dataset="299689" ID-DOI="10.11646/zootaxa.5476.1.15" ID-GBIF-Dataset="139735a4-8b0a-4bce-8ee2-d2172963f007" ID-ISSN="1175-5326" ID-Zenodo-Dep="12724142" ID-ZooBank="6032BF9E-6F66-48F0-B362-04350EA8023E" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1720374744868" checkinUser="plazi" docAuthor="Hsu, Jhih-Wei &amp; Shih, Hsi-Te" docDate="2024" docId="301287C5807197341ABE9386FCC2FCD6" docLanguage="en" docName="zootaxa.5476.1.15.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.15" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptychognathus lipkei N. K. Ng 2010" docType="treatment" docVersion="5" lastPageNumber="161" masterDocId="CC2BFFBD8077973D1A299311FFA9FFEF" masterDocTitle="A new species of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Vanuatu" masterLastPageNumber="165" masterPageNumber="152" pageNumber="158" updateTime="1720711193919" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4C7AF0C35F4592C6ED9AF2BBF3F6FCA6">A new species of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Vanuatu</mods:title>
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<mods:namePart id="B8F38126018BDAC5AB99A66B0FFBCFCB">Hsu, Jhih-Wei</mods:namePart>
<mods:affiliation id="AB5AB49A1D25C64617402D08BC9B09E7">National Chung Hsing University, Department of Life Science. 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:namePart id="44EA9BA81BDA802E4ED68D531FBC09A2">Shih, Hsi-Te</mods:namePart>
<mods:affiliation id="F66F375F93A82944866422BC629F67AF">National Chung Hsing University, Department of Life Science. 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<taxonomicName id="7FBB4D508071973B1ABE9386FE6EFF5D" ID-CoL="6WNM5" authority="Ng, 2010" authorityName="N. K. Ng" authorityYear="2010" box="[151,455,151,178]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="158" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC18071973B1ABE9386FE6EFF5D" bold="true" box="[151,455,151,178]" pageId="6" pageNumber="158">
<emphasis id="8ACFEAC18071973B1ABE9386FEE5FF5D" bold="true" box="[151,332,151,178]" italics="true" pageId="6" pageNumber="158">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC18071973B1BAD9386FE6EFF5D" bold="true" box="[388,455,151,178]" italics="true" pageId="6" pageNumber="158">lipkei</emphasis>
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<paragraph id="B80436D38071973B1ABE93AAFE97FF3A" blockId="6.[151,455,151,214]" box="[151,318,187,214]" pageId="6" pageNumber="158">
(
<figureCitation id="20802A568071973B1AB693AAFEB5FF39" box="[159,284,187,214]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Figs. 1E, F</figureCitation>
,
<figureCitation id="20802A568071973B1B0E93AAFE9CFF3A" box="[295,309,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">3</figureCitation>
)
</paragraph>
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<subSubSection id="F0A165588071973B1ABE9212FF7EFE66" pageId="6" pageNumber="158" type="materials_examined">
<paragraph id="B80436D38071973B1ABE9212FF7EFE66" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">
<emphasis id="8ACFEAC18071973B1ABE9212FE2BFEF2" bold="true" box="[151,386,259,285]" pageId="6" pageNumber="158">Material examined.</emphasis>
<materialsCitation id="08D33C8E8071973B1BA29212FC79FEF2" ID-GBIF-Occurrence="4905142302" box="[395,976,259,286]" collectionCode="ZRC" pageId="6" pageNumber="158" specimenCode="ZRC 2023.0250" specimenCount="1" specimenCount-male="1">
<specimenCount id="AEBDFD5A8071973B1BA29212FE7EFEF2" box="[395,471,259,285]" count="1" pageId="6" pageNumber="158" type="male">1 male</specimenCount>
(11.4 ×
<quantity id="7F439B368071973B18129212FD3FFEF2" box="[571,662,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="158" unit="mm" value="9.6">9.6 mm</quantity>
),
<specimenCode id="E81D9EA88071973B18829212FCCDFEF1" box="[683,868,259,286]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="158">ZRC 2023.0250</specimenCode>
, st. VM4
</materialsCitation>
,
<materialsCitation id="08D33C8E8071973B19F49212FA30FEAE" ID-GBIF-Occurrence="4905142307" collectingDate="2006-09-11" collectionCode="SANTO" collectorName="Marine Biodiversity Survey" country="Vanuatu" location="Espiritu Santo Island" pageId="6" pageNumber="158" specimenCode="SANTO 2006" specimenCount="1" stateProvince="Sanma">
<locationDeviation id="A9B62FF18071973B19F49212FA31FEF2" box="[989,1432,259,286]" firstLocation="Rose Point" pageId="6" pageNumber="158" secondLocation="Nasouli River">between Rose Point and Nasouli River</locationDeviation>
,
<collectingRegion id="7A7FF8318071973B1ABE9236FEFAFEAE" box="[151,339,295,321]" country="Vanuatu" name="Sanma" pageId="6" pageNumber="158">Sanma Province</collectingRegion>
,
<location id="BD6460088071973B1B489236FDFDFEAE" LSID="urn:lsid:plazi:treatment:301287C5807197341ABE9386FCC2FCD6:BD6460088071973B1B489236FDFDFEAE" box="[353,596,295,321]" country="Vanuatu" name="Espiritu Santo Island" pageId="6" pageNumber="158" stateProvince="Sanma">Espiritu Santo Island</location>
,
<collectingCountry id="C0AC76438071973B184B9236FD68FEAE" box="[610,705,295,321]" name="Vanuatu" pageId="6" pageNumber="158">Vanuatu</collectingCountry>
, coll.
<specimenCode id="E81D9EA88071973B19209236FC0EFEAE" box="[777,935,295,321]" collectionCode="SANTO" pageId="6" pageNumber="158">SANTO 2006</specimenCode>
<collectorName id="154E53058071973B19989236FB59FEAE" box="[945,1264,295,321]" pageId="6" pageNumber="158">Marine Biodiversity Survey</collectorName>
,
<date id="CC0510138071973B1ED79236FA30FEAE" box="[1278,1433,295,321]" pageId="6" pageNumber="158" value="2006-09-11">
<collectingDate id="DC41E9FB8071973B1ED79236FA30FEAE" box="[1278,1433,295,321]" pageId="6" pageNumber="158" value="2006-09-11">11 Sep. 2006</collectingDate>
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.
<materialsCitation id="08D33C8E8071973B1ABE925BFF4FFE8B" ID-GBIF-Occurrence="4905142304" box="[151,230,330,356]" country="Vanuatu" location="Others" pageId="6" pageNumber="158" specimenCount="1" stateProvince="Sanma">
<location id="BD6460088071973B1ABE925BFF4FFE8B" LSID="urn:lsid:plazi:treatment:301287C5807197341ABE9386FCC2FCD6:BD6460088071973B1ABE925BFF4FFE8B" box="[151,230,330,356]" country="Vanuatu" name="Others" pageId="6" pageNumber="158" stateProvince="Sanma">Others</location>
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:
<materialsCitation id="08D33C8E8071973B1ADC925AFF7AFE66" ID-GBIF-Occurrence="4905142305" collectingDate="2005-01-03" collectorName="Bea Jonho Stream &amp; Island" country="Solomon Islands" location="Isabel" pageId="6" pageNumber="158" specimenCode="MNHN-B29902" specimenCount="1" specimenCount-male="1" stateProvince="Isabel">
<specimenCount id="AEBDFD5A8071973B1ADC925AFEEDFE8B" box="[245,324,330,357]" count="1" pageId="6" pageNumber="158" type="male">1 male</specimenCount>
(12.8 ×
<quantity id="7F439B368071973B1B86925AFDB2FE8A" box="[431,539,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="158" unit="mm" value="10.7">10.7 mm</quantity>
),
<specimenCode id="E81D9EA88071973B181B925AFD45FE8A" box="[562,748,331,357]" collectionCode="MNHN-B" pageId="6" pageNumber="158">MNHN-B29902</specimenCode>
,
<collectorName id="154E53058071973B18D2925AFC7BFE8A" box="[763,978,330,357]" pageId="6" pageNumber="158">Bea Jonho Stream</collectorName>
,
<collectingRegion id="7A7FF8318071973B19C8925BFB8DFE8B" box="[993,1060,330,356]" country="Solomon Islands" name="Isabel" pageId="6" pageNumber="158">Isabel</collectingRegion>
<collectorName id="154E53058071973B1E06925BFBDFFE8B" box="[1071,1142,330,356]" pageId="6" pageNumber="158">Island</collectorName>
,
<collectingCountry id="C0AC76438071973B1EAC925BFAE1FE8B" box="[1157,1352,330,356]" name="Solomon Islands" pageId="6" pageNumber="158">Solomon Islands</collectingCountry>
,
<date id="CC0510138071973B1F7E925AFF7AFE66" pageId="6" pageNumber="158" value="2005-01-03">
<collectingDate id="DC41E9FB8071973B1F7E925AFF7AFE66" pageId="6" pageNumber="158" value="2005-01-03">3 Jan. 2005</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<subSubSection id="F0A165588071973B1AEE9283FD09FCDA" pageId="6" pageNumber="158" type="diagnosis">
<paragraph id="B80436D38071973B1AEE9283FD09FCDA" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">
<emphasis id="8ACFEAC18071973B1AEE9283FEEBFE43" bold="true" box="[199,322,402,428]" pageId="6" pageNumber="158">Diagnosis.</emphasis>
Carapace (
<figureCitation id="20802A568071973B1BEB9283FDE8FE42" box="[450,577,402,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Figs. 1E, F</figureCitation>
,
<figureCitation id="20802A568071973B18679283FDDBFE43" box="[590,626,402,428]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">3A</figureCitation>
) subquadrate, 1.191.20 times as wide as long; dorsal surface glabrous (except for metabranchial region), postfrontal region indistinct, not separated into any lobes. Front broad; frontal margin concave medially, lined with tiny granules. Anterolateral margins with only 1 distinct, bluntly triangular tooth behind low external orbital tooth. Third maxillipeds (
<figureCitation id="20802A568071973B190492ECFC2AFDF8" box="[813,899,509,535]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3B</figureCitation>
) almost glabrous, exopod 1.25 times as wide as ischium. Chelipeds (
<figureCitation id="20802A568071973B1B579130FE59FDD4" box="[382,496,545,571]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Fig. 1E, F</figureCitation>
) symmetrical; inner surface of palm glabrous; in male, proximal half of immovable finger with tuft of long soft setae on outer surface, extending to palm side, pulvinus absent at base of fingers (
<figureCitation id="20802A568071973B1F469154FF12FD6C" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3C</figureCitation>
). P4 (
<figureCitation id="20802A568071973B1B299179FEF0FD6C" box="[256,345,616,643]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3D</figureCitation>
) relatively long, propodus and dactylus covered with dense short setae on anterior margins and dorsal surface, dense short setae and sparse long setae on posterior margins. P5 (
<figureCitation id="20802A568071973B1E0E919DFBD5FD49" box="[1063,1148,652,678]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3E</figureCitation>
) short; propodus covered with dense short setae and sparse long setae on dorsal surface, anterior margin and distal 1/31/2 of posterior margin; dactylus with dense short setae on margins. Male pleon (
<figureCitation id="20802A568071973B193491C5FCD8FD01" box="[797,881,724,750]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3F</figureCitation>
) narrow; distal margin of telson not concave nor with tuft of setae. G1 (
<figureCitation id="20802A568071973B1BBD91E6FDAAFCFE" box="[404,515,759,785]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3GI</figureCitation>
) slender, distally curved toward dorsal; tip chitinous, with 1 short rounded lobe in lateral view (
<figureCitation id="20802A568071973B1B03900AFE38FCDA" box="[298,401,795,821]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3I, J</figureCitation>
), opened toward lateral.
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807197351AEE902EFC80FE42" lastPageId="8" lastPageNumber="160" pageId="6" pageNumber="158" type="description">
<paragraph id="B80436D38071973B1AEE902EFB91FC07" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">
<emphasis id="8ACFEAC18071973B1AEE902EFEF0FCB6" bold="true" box="[199,345,831,857]" pageId="6" pageNumber="158">Description.</emphasis>
Carapace (
<figureCitation id="20802A568071973B1BF0902EFDFEFCB6" box="[473,599,831,857]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Figs. 1E, F</figureCitation>
,
<figureCitation id="20802A568071973B184D902EFD2EFCB6" box="[612,647,831,857]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">3A</figureCitation>
) subquadrate, 1.191.20 times (n = 2) as wide as long; dorsal surface flattish, finely punctate, glabrous (except for metabranchial region); regions weakly defined, with noticeable groove between epigastric regions, metabranchial region slightly sloping outwards. Front broad, part near orbital regions slightly convex; frontal margin concave medially in dorsal view, weakly divided into indistinct two lobes, lined with small, rounded granules; postfrontal region indistinct, not separated into any lobes.
</paragraph>
<paragraph id="B80436D38071973B1AEE90E0FE3FFB51" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">Supraorbital margins and margins of anterior part in external orbital tooth lined with small granules.Anterolateral margins almost not granulated (except for anterior part in external orbital tooth), with only 1 distinct tooth behind external orbital tooth; external orbital tooth low, bluntly triangular, slightly sloping forward; first tooth blunter than external orbital tooth. Posterolateral margins slightly convergent posteriorly; posterolateral regions sparsely furnished with short, soft setae. Infraorbital ridge consisting of 2225 small, rounded granules. Epistome broad, median part triangular.</paragraph>
<paragraph id="B80436D38071973B1AEE97D9FE22FAA2" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">
Third maxillipeds (
<figureCitation id="20802A568071973B1BB797D9FE5DFB0D" box="[414,500,1224,1250]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3B</figureCitation>
) broad, external surface almost glabrous; merus with oblique shallow groove along mesial margin of external surface, anterolateral angle broadly rounded, slightly sloping laterally; ischium with indistinct shallow groove slightly mesial to midline of external surface; exopod very broad, convex, 1.25 times broader than ischium.
</paragraph>
<paragraph id="B80436D38071973B1AEE9646FB99F9A8" blockId="6.[151,1437,259,2037]" pageId="6" pageNumber="158">
Chelipeds (
<figureCitation id="20802A568071973B1B6F9646FE1FFA9E" box="[326,438,1367,1393]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Fig. 1E, F</figureCitation>
) symmetrical both in male and female, stronger in male. Merus without spines, dorsal and inner margins with dense soft setae, ventral margins glabrous. Carpus almost glabrous on surface, only with several tiny granules; inner distal angle blunt, without any spines. In male, palm (
<figureCitation id="20802A568071973B19FB968FFB81FA57" box="[978,1064,1438,1464]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3C</figureCitation>
) finely punctate, without distinct granules on outer surface; inner surface glabrous, slightly convex medially. Movable finger (dactylus) approximately as long as palm, cutting edge with 6 or 7 smaller and 2 or 3 larger teeth, large teeth blunter; immovable finger slightly shorter than movable finger, cutting edge with 4 or 5 large blunt teeth; proximal half of immovable finger with tuft of long soft setae, extending to palm, without pulvinus at base of fingers.
</paragraph>
<paragraph id="B80436D3807197351AEE9540FF50FF3A" blockId="6.[151,1437,259,2037]" lastBlockId="8.[151,1437,150,717]" lastPageId="8" lastPageNumber="160" pageId="6" pageNumber="158">
Ambulatory legs (
<figureCitation id="20802A568071973B1BBB9540FDA7F984" box="[402,526,1617,1643]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="6" pageNumber="158">Figs. 1E, F</figureCitation>
,
<figureCitation id="20802A568071973B18309540FDF2F984" box="[537,603,1617,1643]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">3D, E</figureCitation>
) slender, P3 and P4 equal in length, longest. Meri without spines, but with long soft setae on proximal 2/3 of anterior margin; posterior margins with sparse long setae and dense short setae; lateral dorsal surface with dense short setae. Carpi of P2 and P3 with dense short setae on distal 3/44/5 anterior margins and dorsal surface; carpi and propodi with dense short setae and sparse long setae on ventral surfaces; propodi sparsely setose on posterior margins; propodi and dactyli with dense short setae on anterior margins. Carpi of P4 and P5 with dense short setae on distal 2/33/4 anterior margins, with sparse long setae on posterior margins. P4 (
<figureCitation id="20802A568071973B1AEF9436FE89F8AE" box="[198,288,1831,1857]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3D</figureCitation>
) relatively long; merus ca. 3.6 as long as wide (ZRC 2023.0250); propodus ca. 2.3 as long as wide (ZRC 2023.0250); propodus and dactylus covered with dense short setae on anterior margins and dorsal surface, with dense short setae and sparse long setae on posterior margins; dactylus ca. 1.1 times as long as propodus (ZRC 2023.0250). P5 (
<figureCitation id="20802A568071973B1B7D9483FE00F842" box="[340,425,1938,1965]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="6" pageNumber="158">Fig. 3E</figureCitation>
) short; merus ca. 2.7 as long as wide (ZRC 2023.0250); propodus ca. 1.3 as long as wide (ZRC 2023.0250), covered with dense short setae (distal part denser) and sparse long setae on anterior margins and dorsal surface near anterior margin, with dense short setae and sparse long setae on distal 1/31/2 posterior margin; dactylus ca. 0.7 times as long as propodus (ZRC 2023.0250), with dense short setae on anterior and posterior margins.
</paragraph>
<caption id="ECC4665B8070973A1ABE9467FF71F839" ID-DOI="http://doi.org/10.5281/zenodo.12724154" ID-Zenodo-Dep="12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="7" pageNumber="159" startId="7.[151,250,1910,1934]" targetBox="[159,1428,189,1878]" targetPageId="7" targetType="figure">
<paragraph id="B80436D38070973A1ABE9467FF71F839" blockId="7.[151,1437,1910,2006]" pageId="7" pageNumber="159">
<emphasis id="8ACFEAC18070973A1ABE9467FEA7F861" bold="true" box="[151,270,1910,1934]" pageId="7" pageNumber="159">FIGURE 3</emphasis>
.
<taxonomicName id="7FBB4D508070973A1B359467FDB5F861" authorityName="N. K. Ng" authorityYear="2010" box="[284,540,1910,1934]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="159" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC18070973A1B359467FE1AF861" box="[284,435,1910,1934]" italics="true" pageId="7" pageNumber="159">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC18070973A1BCD9467FDB5F861" box="[484,540,1910,1934]" italics="true" pageId="7" pageNumber="159">lipkei</emphasis>
</taxonomicName>
. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view).
</paragraph>
</caption>
<paragraph id="B80436D3807F97351AEE93CEFED6FEAE" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="160">
Male pleon (
<figureCitation id="20802A56807F97351B7193CEFE04FF16" box="[344,429,223,249]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3F</figureCitation>
) narrow; external surface smooth, without any granules; lateral margins lined with short setae; telson tongue-shaped, longer and narrower than sixth pleonal segment, distal margin of telson not concave, without tuft of setae.
</paragraph>
<paragraph id="B80436D3807F97351AEE925AFBC6FE66" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="160">
Male G1 (
<figureCitation id="20802A56807F97351B11925AFE00FE8A" box="[312,425,331,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3GJ</figureCitation>
) slender, slightly curved, distally curved toward dorsal; tip chitinous, with 1 short rounded lobe in lateral view (
<figureCitation id="20802A56807F97351B57927EFE4CFE66" box="[382,485,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3I, J</figureCitation>
), opened toward lateral; G2 shorter than 1/4 length of G1.
</paragraph>
<paragraph id="B80436D3807F97351AEE9283FC80FE42" blockId="8.[151,1437,150,717]" box="[199,809,402,429]" pageId="8" pageNumber="160">
<emphasis id="8ACFEAC1807F97351AEE9283FF5EFE43" bold="true" box="[199,247,402,428]" pageId="8" pageNumber="160">Size</emphasis>
. Largest male 12.8 ×
<quantity id="7F439B36807F97351BC59282FDF9FE42" box="[492,592,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="8" pageNumber="160" unit="mm" value="10.7">10.7 mm</quantity>
(MNHN-B29902).
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807F97351AEE92A7FBAFFE3E" box="[199,1030,438,465]" pageId="8" pageNumber="160" type="distribution">
<paragraph id="B80436D3807F97351AEE92A7FBAFFE3E" blockId="8.[151,1437,150,717]" box="[199,1030,438,465]" pageId="8" pageNumber="160">
<emphasis id="8ACFEAC1807F97351AEE92A7FEC8FE3F" bold="true" box="[199,353,438,464]" pageId="8" pageNumber="160">Distribution.</emphasis>
At present, only known from
<collectingCountry id="C0AC7643807F9735189892A6FCA4FE3E" box="[689,781,439,465]" name="Vanuatu" pageId="8" pageNumber="160">Vanuatu</collectingCountry>
and
<collectingCountry id="C0AC7643807F9735196A92A6FBABFE3E" box="[835,1026,439,465]" name="Solomon Islands" pageId="8" pageNumber="160">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807F97341AEE92CAFCC2FCD6" lastPageId="9" lastPageNumber="161" pageId="8" pageNumber="160" type="discussion">
<paragraph id="B80436D3807F97351AEE92CAFEFEFD22" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="160">
<emphasis id="8ACFEAC1807F97351AEE92CAFE92FE1A" bold="true" box="[199,315,475,501]" pageId="8" pageNumber="160">Remarks.</emphasis>
The morphology of the
<specimenCount id="AEBDFD5A807F9735186092CAFD19FE1A" box="[585,688,475,501]" count="2" pageId="8" pageNumber="160" type="male">two male</specimenCount>
specimens of this species agrees with the description of
<taxonomicName id="7FBB4D50807F97351F0E92CAFA29FE1A" authorityName="N. K. Ng" authorityYear="2010" box="[1319,1408,475,501]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351F0E92CAFA29FE1A" box="[1319,1408,475,501]" italics="true" pageId="8" pageNumber="160">P. lipkei</emphasis>
</taxonomicName>
in N.K.
<bibRefCitation id="DC2A4B22807F97351AFD92EEFEEEFDF6" author="Ng, N. K." box="[212,327,511,537]" pageId="8" pageNumber="160" pagination="547 - 560" refId="ref9566" refString="Ng, N. K. (2010) A new species of Ptychognathus Stimpson, 1858, from Cebu Island, Philippines (Decapoda, Brachyura, Varunidae). Crustaceana Monographs, 14, 547 - 560. https: // doi. org / 10.1163 / 9789047427759 _ 038" type="journal article" year="2010">Ng (2010)</bibRefCitation>
and
<bibRefCitation id="DC2A4B22807F97351B5392EEFDFDFDF6" author="Komai, T. &amp; Maenosono, T. &amp; Saeki, T. &amp; Naruse, T." box="[378,596,511,537]" pageId="8" pageNumber="160" pagination="58 - 76" refId="ref9179" refString="Komai, T., Maenosono, T., Saeki, T. &amp; Naruse, T. (2021) Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae). Zootaxa, 5048 (1), 58 - 76. https: // doi. org / 10.11646 / zootaxa. 5048.1.3" type="journal article" year="2021">
Komai
<emphasis id="8ACFEAC1807F97351BE492EEFDADFDF6" box="[461,516,511,537]" italics="true" pageId="8" pageNumber="160">et al.</emphasis>
(2021)
</bibRefCitation>
in most characters (e.g., carapace, maxilliped, ambulatory legs, G1), except the immovable fingers of male chelipeds, which covered with tuft of long soft setae on the proximal half (
<figureCitation id="20802A56807F97351F179132FA3AFDD2" box="[1342,1427,547,573]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3C</figureCitation>
) [vs. immovable finger glabrous in
<taxonomicName id="7FBB4D50807F973518309156FDDBFD8E" authorityName="N. K. Ng" authorityYear="2010" box="[537,626,583,609]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F973518309156FDDBFD8E" box="[537,626,583,609]" italics="true" pageId="8" pageNumber="160">P. lipkei</emphasis>
</taxonomicName>
(N.K.
<bibRefCitation id="DC2A4B22807F973518E99156FC8CFD8E" author="Ng, N. K." box="[704,805,583,609]" pageId="8" pageNumber="160" pagination="547 - 560" refId="ref9566" refString="Ng, N. K. (2010) A new species of Ptychognathus Stimpson, 1858, from Cebu Island, Philippines (Decapoda, Brachyura, Varunidae). Crustaceana Monographs, 14, 547 - 560. https: // doi. org / 10.1163 / 9789047427759 _ 038" type="journal article" year="2010">Ng 2010</bibRefCitation>
: fig. 2G;
<bibRefCitation id="DC2A4B22807F973519A49156FBF3FD8E" author="Komai, T. &amp; Maenosono, T. &amp; Saeki, T. &amp; Naruse, T." box="[909,1114,583,609]" pageId="8" pageNumber="160" pagination="58 - 76" refId="ref9179" refString="Komai, T., Maenosono, T., Saeki, T. &amp; Naruse, T. (2021) Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae). Zootaxa, 5048 (1), 58 - 76. https: // doi. org / 10.11646 / zootaxa. 5048.1.3" type="journal article" year="2021">
Komai
<emphasis id="8ACFEAC1807F973519C99156FBB1FD8E" box="[992,1048,583,609]" italics="true" pageId="8" pageNumber="160">et al.</emphasis>
2021
</bibRefCitation>
: fig. 3B)]. Even the apparent difference on male chelipeds, however, this species and
<taxonomicName id="7FBB4D50807F9735193C917AFCC6FD6A" authorityName="N. K. Ng" authorityYear="2010" box="[789,879,619,645]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F9735193C917AFCC6FD6A" box="[789,879,619,645]" italics="true" pageId="8" pageNumber="160">P. lipkei</emphasis>
</taxonomicName>
form a well-supported clade in COI tree (
<figureCitation id="20802A56807F97351F66917AFA3DFD6A" box="[1359,1428,619,645]" captionStart="FIGURE 4" captionStartId="10.[151,250,1316,1340]" captionTargetBox="[184,1404,188,1285]" captionTargetId="figure-17@10.[177,1411,181,1292]" captionTargetPageId="10" captionText="FIGURE 4. A neighbor-joining tree for species of Ptychognathus, based on the cytochrome c oxidase subunit I (COI) gene. Probability values at the nodes represent support values. Only values&gt; 50% are shown. For haplotype names, see Table 1." figureDoi="http://doi.org/10.5281/zenodo.12724156" httpUri="https://zenodo.org/record/12724156/files/figure.png" pageId="8" pageNumber="160">Fig. 4</figureCitation>
) with very low distances within the clade (
<tableCitation id="F5390368807F97351858919EFD63FD46" box="[625,714,655,681]" captionStart="TABLE 2" captionStartId="8.[151,239,764,788]" captionText="TABLE 2. Matrix of percentage pairwise nucleotide divergence with Kimura 2-parameter (K2P) distances (upper) and" pageId="8" pageNumber="160">Table 2</tableCitation>
). As a result, we tentatively refer to this species as
<taxonomicName id="7FBB4D50807F97351F3A919EFA35FD46" authorityName="N. K. Ng" authorityYear="2010" box="[1299,1436,655,681]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351F3A919EFA80FD47" box="[1299,1321,655,680]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351F77919EFA35FD46" box="[1374,1436,655,681]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
(see Discussion).
</paragraph>
<caption id="ECC4665B807F97351ABE91EDFA34FCFB" box="[151,1437,764,789]" pageId="8" pageNumber="160" startId="8.[151,239,764,788]" targetType="table">
<paragraph id="B80436D3807F97351ABE91EDFA34FCFB" blockId="8.[151,1437,764,861]" box="[151,1437,764,789]" pageId="8" pageNumber="160">
<emphasis id="8ACFEAC1807F97351ABE91EDFEACFCFB" bold="true" box="[151,261,764,788]" pageId="8" pageNumber="160">TABLE 2</emphasis>
. Matrix of percentage pairwise nucleotide divergence with Kimura 2-parameter (K2P) distances (upper) and
</paragraph>
</caption>
<tableNote id="455D375D807F97351ABE9030FB82FCB3" pageId="8" pageNumber="160" targetBox="[161,1420,875,1195]" targetPageId="8">
<paragraph id="B80436D3807F97351ABE9030FA35FCD7" blockId="8.[151,1437,764,861]" box="[151,1436,800,825]" pageId="8" pageNumber="160">mean numbers of differences (lower) based on the cytochrome c oxidase subunit I (COI) within and between species of</paragraph>
<paragraph id="B80436D3807F97351ABE9055FB82FCB3" blockId="8.[151,1437,764,861]" box="[151,1067,836,861]" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351ABE9055FE9FFCB3" authorityName="Stimpson" authorityYear="1858" box="[151,310,836,860]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="8ACFEAC1807F97351ABE9055FE9FFCB3" box="[151,310,836,860]" italics="true" pageId="8" pageNumber="160">Ptychognathus</emphasis>
</taxonomicName>
(see Table 1 and Fig. 5). Values of the range are shown in parentheses.
</paragraph>
</tableNote>
<paragraph id="B80436D3807F97351B4A907DFACEFB44" pageId="8" pageNumber="160">
<table id="CABBC473807F68C21A88907AFA25FB44" box="[161,1420,875,1195]" gridcols="8" gridrows="7" pageId="8" pageNumber="160">
<tr id="068B3491807F68C21A88907AFA25FC4F" box="[161,1420,875,928]" gridrow="0" pageId="8" pageNumber="160" rowspan-0="1">
<th id="455A5DED807F68C21B71907AFE4EFC4F" box="[344,487,875,928]" gridcol="1" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351B4A907DFE75FC6C" authorityName="Hsu &amp; Shih" authorityYear="2020" box="[355,476,876,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="stimpsoni">
<emphasis id="8ACFEAC1807F97351B4A907DFE75FC6C" box="[355,476,876,899]" italics="true" pageId="8" pageNumber="160">P. stimpsoni</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21825907AFDDDFC4F" box="[524,628,875,928]" gridcol="2" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351825907DFDDDFC6C" authorityName="Heller" authorityYear="1865" box="[524,628,875,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="8ACFEAC1807F97351825907DFDDDFC6C" box="[524,628,875,899]" italics="true" pageId="8" pageNumber="160">P. pusillus</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C218B1907AFD49FC4F" box="[664,736,875,928]" gridcol="3" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F9735189A907DFD49FC4F" authorityName="Hsu &amp; Shih" authorityYear="2024" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="amikee">
<emphasis id="8ACFEAC1807F9735189A907DFD49FC4F" italics="true" pageId="8" pageNumber="160">P. amikee</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21923907AFCC9FC4F" box="[778,864,875,928]" gridcol="4" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351923907DFCC9FC6C" authorityName="Hsu, Shih &amp; Li" authorityYear="2022" box="[778,864,875,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="8ACFEAC1807F97351923907DFCC9FC6C" box="[778,864,875,899]" italics="true" pageId="8" pageNumber="160">P. sakaii</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C219A1907AFBBEFC4F" box="[904,1047,875,928]" gridcol="5" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F973519B1907DFBAEFC6C" authorityName="Osawa &amp; NK Ng" authorityYear="2006" box="[920,1031,875,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="8ACFEAC1807F973519B1907DFBAEFC6C" box="[920,1031,875,899]" italics="true" pageId="8" pageNumber="160">P. insolitus</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21E11907AFB7DFC4F" box="[1080,1236,875,928]" gridcol="6" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351E11907DFB7DFC6C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1080,1236,875,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="8ACFEAC1807F97351E11907DFBE5FC6C" box="[1080,1100,876,899]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351E53907AFB7DFC6C" box="[1146,1236,875,899]" italics="true" pageId="8" pageNumber="160">barbatus</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21EDD907AFA25FC4F" box="[1268,1420,875,928]" gridcol="7" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351EDD907DFA25FC6C" authorityName="Sakai" authorityYear="1955" box="[1268,1420,875,899]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="hachijoensis">
<emphasis id="8ACFEAC1807F97351EDD907DFA25FC6C" box="[1268,1420,875,899]" italics="true" pageId="8" pageNumber="160">P. hachijoensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="068B3491807F68C21A8890BFFA25FC29" box="[161,1420,942,966]" gridrow="1" pageId="8" pageNumber="160" rowspan-3="1">
<th id="455A5DED807F68C21A8890BFFEB3FC29" box="[161,282,942,966]" gridcol="0" gridrow="1" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351A8890BEFEABFC29" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[161,258,942,966]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="amikee">
<emphasis id="8ACFEAC1807F97351A8890BEFEABFC29" box="[161,258,942,966]" italics="true" pageId="8" pageNumber="160">P. amikee</emphasis>
</taxonomicName>
</th>
<td id="455A5DED807F68C21B7190BFFE4EFC29" box="[344,487,942,966]" gridcol="1" gridrow="1" pageId="8" pageNumber="160">11.23</td>
<td id="455A5DED807F68C2182590BFFDDDFC29" box="[524,628,942,966]" gridcol="2" gridrow="1" pageId="8" pageNumber="160">10.85</td>
<td id="455A5DED807F68C2192390BFFCC9FC29" box="[778,864,942,966]" gridcol="4" gridrow="1" pageId="8" pageNumber="160">11.83</td>
<td id="455A5DED807F68C219A190BFFBBEFC29" box="[904,1047,942,966]" gridcol="5" gridrow="1" pageId="8" pageNumber="160">15.38</td>
<td id="455A5DED807F68C21E1190BFFB7DFC29" box="[1080,1236,942,966]" gridcol="6" gridrow="1" pageId="8" pageNumber="160">13.18</td>
<td id="455A5DED807F68C21EDD90BFFA25FC29" box="[1268,1420,942,966]" gridcol="7" gridrow="1" pageId="8" pageNumber="160">14.05</td>
</tr>
<tr id="068B3491807F68C21A8890DAFA25FC0C" box="[161,1420,971,995]" gridrow="2" pageId="8" pageNumber="160" rowspan-0="1" rowspan-3="1">
<td id="455A5DED807F68C21B7190DAFE4EFC0C" box="[344,487,971,995]" gridcol="1" gridrow="2" pageId="8" pageNumber="160">(11.0511.42)</td>
<td id="455A5DED807F68C2182590DAFDDDFC0C" box="[524,628,971,995]" gridcol="2" gridrow="2" pageId="8" pageNumber="160">(-)</td>
<td id="455A5DED807F68C2192390DAFCC9FC0C" box="[778,864,971,995]" gridcol="4" gridrow="2" pageId="8" pageNumber="160">(-)</td>
<td id="455A5DED807F68C219A190DAFBBEFC0C" box="[904,1047,971,995]" gridcol="5" gridrow="2" pageId="8" pageNumber="160">(15.2815.48)</td>
<td id="455A5DED807F68C21E1190DAFB7DFC0C" box="[1080,1236,971,995]" gridcol="6" gridrow="2" pageId="8" pageNumber="160">(12.9813.37)</td>
<td id="455A5DED807F68C21EDD90DAFA25FC0C" box="[1268,1420,971,995]" gridcol="7" gridrow="2" pageId="8" pageNumber="160">(13.9514.34)</td>
</tr>
<tr id="068B3491807F68C21A8890E1FA25FBC9" box="[161,1420,1008,1062]" gridrow="3" pageId="8" pageNumber="160" rowspan-0="1" rowspan-3="1">
<td id="455A5DED807F68C21B7190E1FE4EFBC9" box="[344,487,1008,1062]" gridcol="1" gridrow="3" pageId="8" pageNumber="160">68 (6769)</td>
<td id="455A5DED807F68C2182590E1FDDDFBC9" box="[524,628,1008,1062]" gridcol="2" gridrow="3" pageId="8" pageNumber="160">65 (-)</td>
<td id="455A5DED807F68C2192390E1FCC9FBC9" box="[778,864,1008,1062]" gridcol="4" gridrow="3" pageId="8" pageNumber="160">71 (-)</td>
<td id="455A5DED807F68C219A190E1FBBEFBC9" box="[904,1047,1008,1062]" gridcol="5" gridrow="3" pageId="8" pageNumber="160">89.5 (8990)</td>
<td id="455A5DED807F68C21E1190E1FB7DFBC9" box="[1080,1236,1008,1062]" gridcol="6" gridrow="3" pageId="8" pageNumber="160">78 (7779)</td>
<td id="455A5DED807F68C21EDD90E1FA25FBC9" box="[1268,1420,1008,1062]" gridcol="7" gridrow="3" pageId="8" pageNumber="160">82.5 (8284)</td>
</tr>
<tr id="068B3491807F68C21A889722FA25FBA4" box="[161,1420,1075,1099]" gridrow="4" pageId="8" pageNumber="160">
<th id="455A5DED807F68C21A889722FEB3FBA4" box="[161,282,1075,1099]" gridcol="0" gridrow="4" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351A889725FEB3FBA4" authorityName="N. K. Ng" authorityYear="2010" box="[161,282,1075,1099]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351A889725FF1CFBA4" box="[161,181,1076,1099]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351ACB9722FEB3FBA4" box="[226,282,1075,1099]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
</th>
<td id="455A5DED807F68C21B719722FE4EFBA4" box="[344,487,1075,1099]" gridcol="1" gridrow="4" pageId="8" pageNumber="160">14.78</td>
<td id="455A5DED807F68C218259722FDDDFBA4" box="[524,628,1075,1099]" gridcol="2" gridrow="4" pageId="8" pageNumber="160">13.65</td>
<td id="455A5DED807F68C218B19722FD49FBA4" box="[664,736,1075,1099]" gridcol="3" gridrow="4" pageId="8" pageNumber="160">15.19</td>
<td id="455A5DED807F68C219239722FCC9FBA4" box="[778,864,1075,1099]" gridcol="4" gridrow="4" pageId="8" pageNumber="160">13.22</td>
<td id="455A5DED807F68C219A19722FBBEFBA4" box="[904,1047,1075,1099]" gridcol="5" gridrow="4" pageId="8" pageNumber="160">16.07</td>
<td id="455A5DED807F68C21E119722FB7DFBA4" box="[1080,1236,1075,1099]" gridcol="6" gridrow="4" pageId="8" pageNumber="160">14.93</td>
<td id="455A5DED807F68C21EDD9722FA25FBA4" box="[1268,1420,1075,1099]" gridcol="7" gridrow="4" pageId="8" pageNumber="160">13.58</td>
</tr>
<tr id="068B3491807F68C21A889741FA25FB87" box="[161,1420,1104,1128]" gridrow="5" pageId="8" pageNumber="160" rowspan-0="1">
<td id="455A5DED807F68C21B719741FE4EFB87" box="[344,487,1104,1128]" gridcol="1" gridrow="5" pageId="8" pageNumber="160">(14.7314.92)</td>
<td id="455A5DED807F68C218259741FDDDFB87" box="[524,628,1104,1128]" gridcol="2" gridrow="5" pageId="8" pageNumber="160">(-)</td>
<td id="455A5DED807F68C218B19741FD49FB87" box="[664,736,1104,1128]" gridcol="3" gridrow="5" pageId="8" pageNumber="160">(-)</td>
<td id="455A5DED807F68C219239741FCC9FB87" box="[778,864,1104,1128]" gridcol="4" gridrow="5" pageId="8" pageNumber="160">(-)</td>
<td id="455A5DED807F68C219A19741FBBEFB87" box="[904,1047,1104,1128]" gridcol="5" gridrow="5" pageId="8" pageNumber="160">(15.9716.17)</td>
<td id="455A5DED807F68C21E119741FB7DFB87" box="[1080,1236,1104,1128]" gridcol="6" gridrow="5" pageId="8" pageNumber="160">(14.9115.1)</td>
<td id="455A5DED807F68C21EDD9741FA25FB87" box="[1268,1420,1104,1128]" gridcol="7" gridrow="5" pageId="8" pageNumber="160">(13.3813.78)</td>
</tr>
<tr id="068B3491807F68C21A889764FA25FB44" box="[161,1420,1141,1195]" gridrow="6" pageId="8" pageNumber="160" rowspan-0="1">
<td id="455A5DED807F68C21B719764FE4EFB44" box="[344,487,1141,1195]" gridcol="1" gridrow="6" pageId="8" pageNumber="160">87.3 (8788)</td>
<td id="455A5DED807F68C218259764FDDDFB44" box="[524,628,1141,1195]" gridcol="2" gridrow="6" pageId="8" pageNumber="160">81 (-)</td>
<td id="455A5DED807F68C218B19764FD49FB44" box="[664,736,1141,1195]" gridcol="3" gridrow="6" pageId="8" pageNumber="160">89 (-)</td>
<td id="455A5DED807F68C219239764FCC9FB44" box="[778,864,1141,1195]" gridcol="4" gridrow="6" pageId="8" pageNumber="160">79 (-)</td>
<td id="455A5DED807F68C219A19764FBBEFB44" box="[904,1047,1141,1195]" gridcol="5" gridrow="6" pageId="8" pageNumber="160">93.5 (9394)</td>
<td id="455A5DED807F68C21E119764FB7DFB44" box="[1080,1236,1141,1195]" gridcol="6" gridrow="6" pageId="8" pageNumber="160">88.1 (8889)</td>
<td id="455A5DED807F68C21EDD9764FA25FB44" box="[1268,1420,1141,1195]" gridcol="7" gridrow="6" pageId="8" pageNumber="160">81 (8082)</td>
</tr>
</table>
</paragraph>
<paragraph id="B80436D3807F97351B4997CFFAC1F9ED" pageId="8" pageNumber="160">
<table id="CABBC473807F68C21A8897CCFA21F9ED" box="[161,1416,1245,1538]" gridcols="7" gridrows="5" pageId="8" pageNumber="160">
<tr id="068B3491807F68C21A8897CCFA21FB1A" box="[161,1416,1245,1269]" gridrow="0" pageId="8" pageNumber="160" rowspan-0="1">
<th id="455A5DED807F68C21B4997CCFE7FFB1A" box="[352,470,1245,1269]" gridcol="1" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351B4997CFFE7FFB1A" authorityName="T Sakai" authorityYear="1939" box="[352,470,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="takahasii">
<emphasis id="8ACFEAC1807F97351B4997CFFE7FFB1A" box="[352,470,1245,1269]" italics="true" pageId="8" pageNumber="160">P. takahasii</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C2183C97CCFD26FB1A" box="[533,655,1245,1269]" gridcol="2" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F9735183C97CFFD26FB1A" authorityName="N. K. Ng" authorityYear="2010" box="[533,655,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F9735183C97CFFD80FB1A" box="[533,553,1246,1269]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F9735187E97CCFD26FB1A" box="[599,655,1245,1269]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C218E797CCFC86FB1A" box="[718,815,1245,1269]" gridcol="3" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F973518E797CFFC86FB1A" authorityName="De Man" authorityYear="1892" box="[718,815,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="8ACFEAC1807F973518E797CFFC86FB1A" box="[718,815,1245,1269]" italics="true" pageId="8" pageNumber="160">P. pilosus</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C2195397CCFC75FB1A" box="[890,988,1245,1269]" gridcol="4" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F9735195397CFFC75FB1A" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1868" box="[890,988,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="riedelii">
<emphasis id="8ACFEAC1807F9735195397CFFC75FB1A" box="[890,988,1245,1269]" italics="true" pageId="8" pageNumber="160">P. riedelii</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21E0797CCFB17FB1A" box="[1070,1214,1245,1269]" gridcol="5" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351E1E97CFFB1DFB1A" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1079,1204,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="altimanus">
<emphasis id="8ACFEAC1807F97351E1E97CFFB1DFB1A" box="[1079,1204,1245,1269]" italics="true" pageId="8" pageNumber="160">P. altimanus</emphasis>
</taxonomicName>
</th>
<th id="455A5DED807F68C21ED097CCFA21FB1A" box="[1273,1416,1245,1269]" gridcol="6" gridrow="0" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351F3197CFFAC0FB1A" authorityName="Hsu &amp; Shih" authorityYear="2020" box="[1304,1385,1245,1269]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="makii">
<emphasis id="8ACFEAC1807F97351F3197CFFAC0FB1A" box="[1304,1385,1245,1269]" italics="true" pageId="8" pageNumber="160">P. makii</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="068B3491807F68C21A889614FA21FAD4" box="[161,1416,1285,1339]" gridrow="1" pageId="8" pageNumber="160">
<th id="455A5DED807F68C21A889614FEB3FAD4" box="[161,282,1285,1339]" gridcol="0" gridrow="1" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351A889617FEABFAF2" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[161,258,1285,1309]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="amikee">
<emphasis id="8ACFEAC1807F97351A889617FEABFAF2" box="[161,258,1285,1309]" italics="true" pageId="8" pageNumber="160">P. amikee</emphasis>
</taxonomicName>
</th>
<td id="455A5DED807F68C21B499614FE7FFAD4" box="[352,470,1285,1339]" gridcol="1" gridrow="1" pageId="8" pageNumber="160">13.29 (-)</td>
<td id="455A5DED807F68C2183C9614FD26FAD4" box="[533,655,1285,1339]" gridcol="2" gridrow="1" pageId="8" pageNumber="160">15.19 (-)</td>
<td id="455A5DED807F68C218E79614FC86FAD4" box="[718,815,1285,1339]" gridcol="3" gridrow="1" pageId="8" pageNumber="160">16.33 (-)</td>
<td id="455A5DED807F68C219539614FC75FAD4" box="[890,988,1285,1339]" gridcol="4" gridrow="1" pageId="8" pageNumber="160">18.55 (-)</td>
<td id="455A5DED807F68C21E079614FB17FAD4" box="[1070,1214,1285,1339]" gridcol="5" gridrow="1" pageId="8" pageNumber="160">18.75 (18.5518.96)</td>
<td id="455A5DED807F68C21ED09614FA21FAD4" box="[1273,1416,1285,1339]" gridcol="6" gridrow="1" pageId="8" pageNumber="160">16.26 (16.1716.36)</td>
</tr>
<tr id="068B3491807F68C21A889659FA21FA92" box="[161,1416,1352,1405]" gridrow="2" pageId="8" pageNumber="160" rowspan-0="1">
<td id="455A5DED807F68C21B499659FE7FFA92" box="[352,470,1352,1405]" gridcol="1" gridrow="2" pageId="8" pageNumber="160">79 (-)</td>
<td id="455A5DED807F68C2183C9659FD26FA92" box="[533,655,1352,1405]" gridcol="2" gridrow="2" pageId="8" pageNumber="160">89 (-)</td>
<td id="455A5DED807F68C218E79659FC86FA92" box="[718,815,1352,1405]" gridcol="3" gridrow="2" pageId="8" pageNumber="160">96 (-)</td>
<td id="455A5DED807F68C219539659FC75FA92" box="[890,988,1352,1405]" gridcol="4" gridrow="2" pageId="8" pageNumber="160">107 (-)</td>
<td id="455A5DED807F68C21E079659FB17FA92" box="[1070,1214,1352,1405]" gridcol="5" gridrow="2" pageId="8" pageNumber="160">108 (107109)</td>
<td id="455A5DED807F68C21ED09659FA21FA92" box="[1273,1416,1352,1405]" gridcol="6" gridrow="2" pageId="8" pageNumber="160">95.5 (9596)</td>
</tr>
<tr id="068B3491807F68C21A88969BFA21FA2F" box="[161,1416,1418,1472]" gridrow="3" pageId="8" pageNumber="160" rowspan-2="1">
<th id="455A5DED807F68C21A88969BFEB3FA2F" box="[161,282,1418,1472]" gridcol="0" gridrow="3" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351A88969AFEB3FA4D" authorityName="N. K. Ng" authorityYear="2010" box="[161,282,1418,1442]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351A88969AFF1CFA4D" box="[161,181,1419,1442]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351ACB969BFEB3FA4D" box="[226,282,1418,1442]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
</th>
<td id="455A5DED807F68C21B49969BFE7FFA2F" box="[352,470,1418,1472]" gridcol="1" gridrow="3" pageId="8" pageNumber="160">13.79 (-)</td>
<td id="455A5DED807F68C218E7969BFC86FA2F" box="[718,815,1418,1472]" gridcol="3" gridrow="3" pageId="8" pageNumber="160">14.86 (-)</td>
<td id="455A5DED807F68C21953969BFC75FA2F" box="[890,988,1418,1472]" gridcol="4" gridrow="3" pageId="8" pageNumber="160">15.23 (-)</td>
<td id="455A5DED807F68C21E07969BFB17FA2F" box="[1070,1214,1418,1472]" gridcol="5" gridrow="3" pageId="8" pageNumber="160">14.83 (14.6415.02)</td>
<td id="455A5DED807F68C21ED0969BFA21FA2F" box="[1273,1416,1418,1472]" gridcol="6" gridrow="3" pageId="8" pageNumber="160">15.64 (15.5415.74)</td>
</tr>
<tr id="068B3491807F68C21A8896DCFA21F9ED" box="[161,1416,1485,1538]" gridrow="4" pageId="8" pageNumber="160" rowspan-0="1" rowspan-2="1">
<td id="455A5DED807F68C21B4996DCFE7FF9ED" box="[352,470,1485,1538]" gridcol="1" gridrow="4" pageId="8" pageNumber="160">82 (-)</td>
<td id="455A5DED807F68C218E796DCFC86F9ED" box="[718,815,1485,1538]" gridcol="3" gridrow="4" pageId="8" pageNumber="160">88 (-)</td>
<td id="455A5DED807F68C2195396DCFC75F9ED" box="[890,988,1485,1538]" gridcol="4" gridrow="4" pageId="8" pageNumber="160">90 (-)</td>
<td id="455A5DED807F68C21E0796DCFB17F9ED" box="[1070,1214,1485,1538]" gridcol="5" gridrow="4" pageId="8" pageNumber="160">88 (8789)</td>
<td id="455A5DED807F68C21ED096DCFA21F9ED" box="[1273,1416,1485,1538]" gridcol="6" gridrow="4" pageId="8" pageNumber="160">92.5 (9293)</td>
</tr>
</table>
</paragraph>
<paragraph id="B80436D3807F97351AEE9521FDC6F959" blockId="8.[151,1437,1584,2043]" pageId="8" pageNumber="160">
<taxonomicName id="7FBB4D50807F97351AEE9521FE48F9A5" authorityName="N. K. Ng" authorityYear="2010" box="[199,481,1584,1610]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351AEE9521FEC4F9A5" box="[199,365,1584,1610]" italics="true" pageId="8" pageNumber="160">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351B8A9521FE48F9A5" box="[419,481,1584,1610]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="7FBB4D50807F973518859520FC60F9A4" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[684,969,1584,1611]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="8ACFEAC1807F973518859520FCB8F9A5" box="[684,785,1584,1610]" italics="true" pageId="8" pageNumber="160">P. glaber</emphasis>
Stimpson,
<quantity id="7F439B36807F973519B89521FC60F9A4" box="[913,969,1584,1611]" metricMagnitude="1" metricUnit="m" metricValue="4.71932" pageId="8" pageNumber="160" unit="in" value="1858.0">1858</quantity>
</taxonomicName>
in the anterolateral teeth of carapace and the setae distribution on ambulatory legs, but the proximal half of outer surface of immovable finger is covered with long dense soft setae, which extends to the palm side in
<taxonomicName id="7FBB4D50807F973519089568FC02F97D" authorityName="N. K. Ng" authorityYear="2010" box="[801,939,1656,1682]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F973519089568FC9EF97D" box="[801,823,1657,1682]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F973519449569FC02F97D" box="[877,939,1656,1682]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
(
<figureCitation id="20802A56807F973519949569FBBDF97D" box="[957,1044,1656,1682]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3C</figureCitation>
) [vs. immovable finger is entirely glabrous (
<bibRefCitation id="DC2A4B22807F97351B2F958DFE7FF959" author="Komai, T. &amp; Maenosono, T. &amp; Saeki, T. &amp; Naruse, T." box="[262,470,1692,1719]" pageId="8" pageNumber="160" pagination="58 - 76" refId="ref9179" refString="Komai, T., Maenosono, T., Saeki, T. &amp; Naruse, T. (2021) Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae). Zootaxa, 5048 (1), 58 - 76. https: // doi. org / 10.11646 / zootaxa. 5048.1.3" type="journal article" year="2021">
Komai
<emphasis id="8ACFEAC1807F97351B72958CFE25F959" box="[347,396,1692,1718]" italics="true" pageId="8" pageNumber="160">et al</emphasis>
. 2021
</bibRefCitation>
: figs. 1, 3B)].
</paragraph>
<paragraph id="B80436D3807F97351AEE95D0FB0DF85D" blockId="8.[151,1437,1584,2043]" pageId="8" pageNumber="160">
Among other species of
<taxonomicName id="7FBB4D50807F97351BFF95D1FDD5F935" authorityName="Stimpson" authorityYear="1858" box="[470,636,1728,1754]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="8ACFEAC1807F97351BFF95D1FDD5F935" box="[470,636,1728,1754]" italics="true" pageId="8" pageNumber="160">Ptychognathus</emphasis>
</taxonomicName>
,
<taxonomicName id="7FBB4D50807F973518A095D0FCA7F935" authorityName="N. K. Ng" authorityYear="2010" box="[649,782,1728,1754]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F973518A095D0FD37F935" box="[649,670,1729,1754]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F973518F995D1FCA7F935" box="[720,782,1728,1754]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
is remarkably similar to
<taxonomicName id="7FBB4D50807F97351E0B95D0FBC4F935" authorityName="Sakai" authorityYear="1939" box="[1058,1133,1728,1754]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="ishii">
<emphasis id="8ACFEAC1807F97351E0B95D0FBC4F935" box="[1058,1133,1728,1754]" italics="true" pageId="8" pageNumber="160">P. ishii</emphasis>
</taxonomicName>
in the immovable finger of male cheliped with a tuft of long setae on outer surface and the ambulatory legs covered with dense short setae on the dorsal surface. However,
<taxonomicName id="7FBB4D50807F97351BF39418FDF7F8CD" authorityName="N. K. Ng" authorityYear="2010" box="[474,606,1800,1826]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351BF39418FE59F8CD" box="[474,496,1801,1826]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F973518099419FDF7F8CD" box="[544,606,1800,1826]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="7FBB4D50807F973519A39418FC7DF8CD" authorityName="Sakai" authorityYear="1939" box="[906,980,1800,1826]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="ishii">
<emphasis id="8ACFEAC1807F973519A39418FC7DF8CD" box="[906,980,1800,1826]" italics="true" pageId="8" pageNumber="160">P. ishii</emphasis>
</taxonomicName>
by the long soft setae only distributed on the proximal half of immovable finger (
<figureCitation id="20802A56807F9735187C943DFD02F8A9" box="[597,683,1836,1862]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3C</figureCitation>
) [vs. long setae present on that part, as well as the movable fingers and the margin of palm (
<bibRefCitation id="DC2A4B22807F97351B989441FD9CF885" author="Sakai, T." box="[433,565,1872,1898]" pageId="8" pageNumber="160" pagination="42 - 111" refId="ref10226" refString="Sakai, T. (1939) n. k. In: Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo Co., Ltd, Tokyo, pp. 365 - 741, pls. 42 - 111." type="book chapter" year="1939">Sakai 1939</bibRefCitation>
: text-fig. 114b)]; and only the parts near the margins of carpi and propodi of ambulatory legs covered with dense short setae and sparse long setae (
<figureCitation id="20802A56807F973519869465FB8FF861" box="[943,1062,1908,1934]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="8" pageNumber="160">Fig. 3D, E</figureCitation>
) [vs. all surfaces of the carpi and propodi covered with dense short setae, but without long setae (
<bibRefCitation id="DC2A4B22807F973519499489FB99F85D" author="Komai, T. &amp; Maenosono, T. &amp; Saeki, T. &amp; Naruse, T." box="[864,1072,1944,1970]" pageId="8" pageNumber="160" pagination="58 - 76" refId="ref9179" refString="Komai, T., Maenosono, T., Saeki, T. &amp; Naruse, T. (2021) Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae). Zootaxa, 5048 (1), 58 - 76. https: // doi. org / 10.11646 / zootaxa. 5048.1.3" type="journal article" year="2021">
Komai
<emphasis id="8ACFEAC1807F9735199C9488FC44F85D" box="[949,1005,1944,1970]" italics="true" pageId="8" pageNumber="160">et al.</emphasis>
2021
</bibRefCitation>
: fig. 9A)].
</paragraph>
<paragraph id="B80436D3807F97341AEE94ACFEFBFF3A" blockId="8.[151,1437,1584,2043]" lastBlockId="9.[151,1437,151,825]" lastPageId="9" lastPageNumber="161" pageId="8" pageNumber="160">
In addition,
<taxonomicName id="7FBB4D50807F97351B7094ACFE40F839" authorityName="N. K. Ng" authorityYear="2010" box="[345,489,1980,2006]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807F97351B7094ACFEC7F839" box="[345,366,1981,2006]" italics="true" pageId="8" pageNumber="160">P.</emphasis>
aff.
<emphasis id="8ACFEAC1807F97351B8294ADFE40F839" box="[427,489,1980,2006]" italics="true" pageId="8" pageNumber="160">lipkei</emphasis>
</taxonomicName>
is also morphologically similar to
<taxonomicName id="7FBB4D50807F973519A594ACFBADF839" authorityName="Heller" authorityYear="1865" box="[908,1028,1980,2006]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="8ACFEAC1807F973519A594ACFBADF839" box="[908,1028,1980,2006]" italics="true" pageId="8" pageNumber="160">P. pusillus</emphasis>
</taxonomicName>
,
<taxonomicName id="7FBB4D50807F97351E3E94ACFBD3F839" authorityName="Hsu, Shih &amp; Li" authorityYear="2022" box="[1047,1146,1980,2006]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="8ACFEAC1807F97351E3E94ACFBD3F839" box="[1047,1146,1980,2006]" italics="true" pageId="8" pageNumber="160">P. sakaii</emphasis>
</taxonomicName>
, and
<taxonomicName id="7FBB4D50807F97351EE894ACFAE5F839" authorityName="Hsu &amp; Shih" authorityYear="2020" box="[1217,1356,1981,2006]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="stimpsoni">
<emphasis id="8ACFEAC1807F97351EE894ACFAE5F839" box="[1217,1356,1981,2006]" italics="true" pageId="8" pageNumber="160">P. stimpsoni</emphasis>
</taxonomicName>
in the chelipedal fingers with dense long setae on outer surface and the male telson without a distinct tuft of soft setae on distal margin, but can be distinguished by the characters of carapace, distribution of long setae on male chelae, and ambulatory legs.
</paragraph>
<paragraph id="B80436D3807E97341AEE93CEFB61FE66" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="161">
<taxonomicName id="7FBB4D50807E97341AEE93CEFE4FFF16" authorityName="N. K. Ng" authorityYear="2010" box="[199,486,223,249]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807E97341AEE93CEFEC4FF16" box="[199,365,223,249]" italics="true" pageId="9" pageNumber="161">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC1807E97341B8193CEFE4FFF16" box="[424,486,223,249]" italics="true" pageId="9" pageNumber="161">lipkei</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="7FBB4D50807E973418D493CEFCDAFF16" authorityName="Heller" authorityYear="1865" box="[765,883,223,249]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="8ACFEAC1807E973418D493CEFCDAFF16" box="[765,883,223,249]" italics="true" pageId="9" pageNumber="161">P. pusillus</emphasis>
</taxonomicName>
by the distribution of setae on male chelipedal fingers and ambulatory legs, including the long soft setae present only on the proximal half of immovable fingers (
<figureCitation id="20802A56807E97341AB69236FF5EFEAE" box="[159,247,295,321]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3C</figureCitation>
) [vs. setae present on both proximal halves of immovable and movable fingers (
<bibRefCitation id="DC2A4B22807E97341EBC9236FADBFEAE" author="De Man, J. G." box="[1173,1394,294,321]" pageId="9" pageNumber="161" pagination="537 - 550" refId="ref8643" refString="De Man, J. G. (1905) On species of Crustacea of the genera Ptychognathus Stimps. and Palaemon Fabr. from Christmas Island. Proceedings of the Zoological Society of London, 1905, 537 - 550." type="book chapter" year="1905">De Man 1905: 539</bibRefCitation>
, pl. 17(5)]; and the carpi of ambulatory legs covered with dense short setae on the anterior margins (
<figureCitation id="20802A56807E97341EC1925AFACBFE8A" box="[1256,1378,331,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3D, E</figureCitation>
) [vs. anterior margins of the carpi glabrous (
<bibRefCitation id="DC2A4B22807E97341865927EFCBDFE66" author="De Man, J. G." box="[588,788,366,393]" pageId="9" pageNumber="161" pagination="75 - 218" refId="ref8568" refString="De Man, J. G. (1895) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 9, 75 - 218. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man 1895: 99</bibRefCitation>
, fig. 22,
<bibRefCitation id="DC2A4B22807E9734195E927EFBE4FE66" author="De Man, J. G." box="[887,1101,366,393]" pageId="9" pageNumber="161" pagination="537 - 550" refId="ref8643" refString="De Man, J. G. (1905) On species of Crustacea of the genera Ptychognathus Stimps. and Palaemon Fabr. from Christmas Island. Proceedings of the Zoological Society of London, 1905, 537 - 550." type="book chapter" year="1905">De Man 1905: 539</bibRefCitation>
, pl. 17(1)].
</paragraph>
<paragraph id="B80436D3807E97341AEE9282FCB7FD6A" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="161">
<taxonomicName id="7FBB4D50807E97341AEE9282FE4FFE42" authorityName="N. K. Ng" authorityYear="2010" box="[199,486,403,429]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807E97341AEE9282FEC4FE42" box="[199,365,403,429]" italics="true" pageId="9" pageNumber="161">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC1807E97341B819282FE4FFE42" box="[424,486,403,429]" italics="true" pageId="9" pageNumber="161">lipkei</emphasis>
</taxonomicName>
and
<taxonomicName id="7FBB4D50807E9734180A9282FD2CFE42" authorityName="Hsu, Shih &amp; Li" authorityYear="2022" box="[547,645,403,429]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="8ACFEAC1807E9734180A9282FD2CFE42" box="[547,645,403,429]" italics="true" pageId="9" pageNumber="161">P. sakaii</emphasis>
</taxonomicName>
can be differentiated by the characters of carapace, male chelipedal fingers, and ambulatory legs, including the postfrontal region lacking any lobes, and the anterolateral margin excluding the orbital tooth with only one distinct tooth (
<figureCitation id="20802A56807E9734192692CAFCC1FE1A" box="[783,872,475,501]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3A</figureCitation>
) [vs. obscure lobes separated by shallow grooves, and the anterolateral margin with two indistinct teeth (
<bibRefCitation id="DC2A4B22807E973418C692EEFC02FDF6" author="Hsu, J. - W. &amp; Shih, H. - T. &amp; Li, J. - J." box="[751,939,511,537]" pageId="9" pageNumber="161" refId="ref8935" refString="Hsu, J. - W., Shih, H. - T. &amp; Li, J. - J. (2022 b) Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan. Nauplius, 30, e 2022002. https: // doi. org / 10.1590 / 2358 - 2936 e 2022002" type="journal volume" year="2022">
Hsu
<emphasis id="8ACFEAC1807E9734190D92EEFCF2FDF6" box="[804,859,511,537]" italics="true" pageId="9" pageNumber="161">et al.</emphasis>
2022b
</bibRefCitation>
: fig. 2A)]; long soft setae present only on the proximal half of immovable finger (
<figureCitation id="20802A56807E973418039132FDD6FDD2" box="[554,639,547,573]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3C</figureCitation>
) [vs. setae present on both proximal halves of immovable and movable fingers (
<bibRefCitation id="DC2A4B22807E97341ADD9156FE1DFD8E" author="Hsu, J. - W. &amp; Shih, H. - T. &amp; Li, J. - J." box="[244,436,583,609]" pageId="9" pageNumber="161" refId="ref8935" refString="Hsu, J. - W., Shih, H. - T. &amp; Li, J. - J. (2022 b) Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan. Nauplius, 30, e 2022002. https: // doi. org / 10.1590 / 2358 - 2936 e 2022002" type="journal volume" year="2022">
Hsu
<emphasis id="8ACFEAC1807E97341B039156FECAFD8E" box="[298,355,583,609]" italics="true" pageId="9" pageNumber="161">et al.</emphasis>
2022b
</bibRefCitation>
: fig. 2D)]; and anterior margins of ambulatory carpi not compressed distally (
<figureCitation id="20802A56807E97341F349156FA3DFD8E" box="[1309,1428,583,609]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3D, E</figureCitation>
) [vs. distinctly compressed (
<bibRefCitation id="DC2A4B22807E97341BE3917AFD21FD6A" author="Hsu, J. - W. &amp; Shih, H. - T. &amp; Li, J. - J." box="[458,648,619,645]" pageId="9" pageNumber="161" refId="ref8935" refString="Hsu, J. - W., Shih, H. - T. &amp; Li, J. - J. (2022 b) Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan. Nauplius, 30, e 2022002. https: // doi. org / 10.1590 / 2358 - 2936 e 2022002" type="journal volume" year="2022">
Hsu
<emphasis id="8ACFEAC1807E97341BD6917AFD91FD6A" box="[511,568,619,645]" italics="true" pageId="9" pageNumber="161">et al.</emphasis>
2022b
</bibRefCitation>
: fig. 2G, H)].
</paragraph>
<paragraph id="B80436D3807E97341AEE919EFCC2FCD6" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="161">
<taxonomicName id="7FBB4D50807E97341AEE919EFE48FD46" authorityName="N. K. Ng" authorityYear="2010" box="[199,481,655,681]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC1807E97341AEE919EFEC4FD46" box="[199,365,655,681]" italics="true" pageId="9" pageNumber="161">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC1807E97341B8A919EFE48FD46" box="[419,481,655,681]" italics="true" pageId="9" pageNumber="161">lipkei</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="7FBB4D50807E97341885919EFC9AFD47" authorityName="Hsu &amp; Shih" authorityYear="2020" box="[684,819,655,680]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="stimpsoni">
<emphasis id="8ACFEAC1807E97341885919EFC9AFD47" box="[684,819,655,680]" italics="true" pageId="9" pageNumber="161">P. stimpsoni</emphasis>
</taxonomicName>
but can be distinguished by the characters of the male chelipedal fingers and ambulatory legs, including only the proximal half part of immovable finger covered with long dense soft setae (
<figureCitation id="20802A56807E97341B7191C6FE07FD1E" box="[344,430,727,753]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3C</figureCitation>
) [vs. proximal halves of both movable and immovable fingers with long dense soft setae (
<bibRefCitation id="DC2A4B22807E97341AB691EAFEC1FCFA" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[159,360,763,789]" pageId="9" pageNumber="161" pagination="249 - 265" refId="ref10460" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">Hsu &amp; Shih 2020</bibRefCitation>
: fig. 4E)]; and anterior margins and surfaces of ambulatory carpi and propodi bearing numerous setae (
<figureCitation id="20802A56807E97341AF7900EFEFCFCD6" box="[222,341,799,825]" captionStart="FIGURE 3" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[159,1428,189,1878]" captionTargetId="figure-22@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ptychognathus aff. lipkei. AJ, male (11.4 × 9.6 mm, ZRC 2023.0250). A, carapace; B, left third maxilliped; C, outer view of left male cheliped; D, left P4; E, left P5; F, male pleon; GJ, male right G1 (G, H, dorsal view; I, J, dorsolateral view)." figureDoi="http://doi.org/10.5281/zenodo.12724154" httpUri="https://zenodo.org/record/12724154/files/figure.png" pageId="9" pageNumber="161">Fig. 3D, E</figureCitation>
) [vs. less setae (
<bibRefCitation id="DC2A4B22807E97341825900EFD7CFCD6" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[524,725,799,825]" pageId="9" pageNumber="161" pagination="249 - 265" refId="ref10460" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">Hsu &amp; Shih 2020</bibRefCitation>
: fig. 4G, H)].
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="301287C5807597391ABE973CFBAFF856" ID-DOI="http://doi.org/10.5281/zenodo.12724146" ID-GBIF-Taxon="235037595" ID-Zenodo-Dep="12724146" LSID="urn:lsid:plazi:treatment:301287C5807597391ABE973CFBAFF856" httpUri="http://treatment.plazi.org/id/301287C5807597391ABE973CFBAFF856" lastPageId="4" lastPageNumber="156" pageId="2" pageNumber="154">
<subSubSection id="F0A165588075973F1ABE973CFDA6FBA7" box="[151,527,1069,1096]" pageId="2" pageNumber="154" type="nomenclature">
<paragraph id="B80436D38075973F1ABE973CFDA6FBA7" blockId="2.[151,527,1069,1132]" box="[151,527,1069,1096]" pageId="2" pageNumber="154">
<heading id="E34C81BF8075973F1ABE973CFDA6FBA7" bold="true" box="[151,527,1069,1096]" fontSize="11" level="1" pageId="2" pageNumber="154" reason="1">
<emphasis id="8ACFEAC18075973F1ABE973CFDA6FBA7" bold="true" box="[151,527,1069,1096]" pageId="2" pageNumber="154">
<taxonomicName id="7FBB4D508075973F1ABE973CFE00FBA7" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[151,425,1069,1096]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="154" phylum="Arthropoda" rank="species" species="amikee" status="sp. nov.">
<emphasis id="8ACFEAC18075973F1ABE973CFE00FBA7" bold="true" box="[151,425,1069,1096]" italics="true" pageId="2" pageNumber="154">Ptychognathus amikee</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91FC57BA8075973F1B98973FFDA6FBA7" box="[433,527,1070,1096]" pageId="2" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F0A165588075973F1ABE9743FEE0FB83" box="[151,329,1106,1132]" pageId="2" pageNumber="154" type="description">
<paragraph id="B80436D38075973F1ABE9743FEE0FB83" blockId="2.[151,527,1069,1132]" box="[151,329,1106,1132]" pageId="2" pageNumber="154">
(
<figureCitation id="20802A568075973F1AB69743FE8CFB83" box="[159,293,1106,1132]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="2" pageNumber="154">Figs. 1AD</figureCitation>
,
<figureCitation id="20802A568075973F1B189743FEE9FB83" box="[305,320,1106,1132]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0A165588075973F1ABE978BFC13FAAB" pageId="2" pageNumber="154" type="materials_examined">
<paragraph id="B80436D38075973F1ABE978BFE42FB12" blockId="2.[151,1437,1177,2032]" pageId="2" pageNumber="154">
<emphasis id="8ACFEAC18075973F1ABE978BFEE0FB5C" bold="true" box="[151,329,1177,1204]" pageId="2" pageNumber="154">Type material.</emphasis>
<materialsCitation id="08D33C8E8075973F1B7A978BFDB2FB36" ID-GBIF-Occurrence="4905142306" collectingDate="2005-07-18" collectorName="Marquet" country="Vanuatu" location="Kirilou River" municipality="Gaua Island" pageId="2" pageNumber="154" specimenCode="MNHN-B29852" specimenCount="1" typeStatus="holotype">
<emphasis id="8ACFEAC18075973F1B7A978BFE13FB5B" box="[339,442,1178,1204]" italics="true" pageId="2" pageNumber="154">
<typeStatus id="670088718075973F1B7A978BFE13FB5B" box="[339,442,1178,1204]" pageId="2" pageNumber="154" type="holotype">Holotype</typeStatus>
</emphasis>
: male (15.8 ×
<quantity id="7F439B368075973F1844978BFD71FB5B" box="[621,728,1178,1204]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="154" unit="mm" value="13.6">13.6 mm</quantity>
),
<specimenCode id="E81D9EA88075973F18C6978BFC00FB5A" box="[751,937,1178,1205]" collectionCode="MNHN-B" pageId="2" pageNumber="154">MNHN-B29852</specimenCode>
,
<location id="BD6460088075973F1991978BFBFDFB5B" LSID="urn:lsid:plazi:treatment:301287C5807597391ABE973CFBAFF856:BD6460088075973F1991978BFBFDFB5B" box="[952,1108,1178,1204]" country="Vanuatu" municipality="Gaua Island" name="Kirilou River" pageId="2" pageNumber="154">Kirilou River</location>
,
<collectingMunicipality id="5860ACA98075973F1E4B978BFB46FB5B" box="[1122,1263,1178,1204]" pageId="2" pageNumber="154">Gaua Island</collectingMunicipality>
,
<collectingCountry id="C0AC76438075973F1ED7978BFAF4FB5B" box="[1278,1373,1178,1204]" name="Vanuatu" pageId="2" pageNumber="154">Vanuatu</collectingCountry>
, coll.
<collectorName id="154E53058075973F1ABE97AFFF5FFB37" box="[151,246,1214,1240]" pageId="2" pageNumber="154">Marquet</collectorName>
and Keith,
<date id="CC0510138075973F1BA897AFFDB2FB36" box="[385,539,1214,1241]" pageId="2" pageNumber="154" value="2005-07-18">
<collectingDate id="DC41E9FB8075973F1BA897AFFDB2FB36" box="[385,539,1214,1241]" pageId="2" pageNumber="154" value="2005-07-18">18 July 2005</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="08D33C8E8075973F180E97AEFE41FB12" ID-GBIF-Occurrence="4905142303" collectingDate="2005-07-20" country="Vanuatu" location="Lembot River" municipality="Gaua Island" pageId="2" pageNumber="154" specimenCode="MNHN-B29853" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="8ACFEAC18075973F180E97AEFD27FB37" box="[551,654,1215,1240]" italics="true" pageId="2" pageNumber="154">
<typeStatus id="670088718075973F180E97AEFD27FB37" box="[551,654,1215,1240]" pageId="2" pageNumber="154" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="AEBDFD5A8075973F18B697AFFCAAFB37" box="[671,771,1214,1240]" count="1" pageId="2" pageNumber="154" type="female">1 female</specimenCount>
(17.4 ×
<quantity id="7F439B368075973F194397AFFC7CFB37" box="[874,981,1214,1240]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="154" unit="mm" value="14.6">14.6 mm</quantity>
),
<specimenCode id="E81D9EA88075973F19C397AFFB0DFB36" box="[1002,1188,1214,1241]" collectionCode="MNHN-B" pageId="2" pageNumber="154">MNHN-B29853</specimenCode>
,
<location id="BD6460088075973F1E9A97AFFAFAFB37" LSID="urn:lsid:plazi:treatment:301287C5807597391ABE973CFBAFF856:BD6460088075973F1E9A97AFFAFAFB37" box="[1203,1363,1214,1240]" country="Vanuatu" municipality="Gaua Island" name="Lembot River" pageId="2" pageNumber="154">Lembot River</location>
,
<collectingMunicipality id="5860ACA98075973F1F4897AFFF77FB13" pageId="2" pageNumber="154">Gaua Island</collectingMunicipality>
,
<collectingCountry id="C0AC76438075973F1AC097F3FEE1FB13" box="[233,328,1250,1276]" name="Vanuatu" pageId="2" pageNumber="154">Vanuatu</collectingCountry>
,
<date id="CC0510138075973F1B7A97F3FE41FB12" box="[339,488,1250,1277]" pageId="2" pageNumber="154" value="2005-07-20">
<collectingDate id="DC41E9FB8075973F1B7A97F3FE41FB12" box="[339,488,1250,1277]" pageId="2" pageNumber="154" value="2005-07-20">20 July 2005</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="B80436D38075973F1AEE9617FC13FAAB" blockId="2.[151,1437,1177,2032]" pageId="2" pageNumber="154">
<emphasis id="8ACFEAC18075973F1AEE9617FE0AFACF" box="[199,419,1286,1312]" italics="true" pageId="2" pageNumber="154">Additional material</emphasis>
:
<materialsCitation id="08D33C8E8075973F1B989617FC1FFAAB" ID-GBIF-Occurrence="4905142301" collectingDate="2006-10-14" collectionCode="ZRC, SANTO" collectorName="SE Baldwin Cove" country="Vanuatu" location="Espiritu Santo Island" pageId="2" pageNumber="154" specimenCode="ZRC 2021.0350, SANTO 2006" specimenCount="1" specimenCount-male="1">
<specimenCount id="AEBDFD5A8075973F1B989617FE52FACF" box="[433,507,1286,1312]" count="1" pageId="2" pageNumber="154" type="male">1 male</specimenCount>
(16.1 × 14.0),
<specimenCode id="E81D9EA88075973F18899617FCFEFACE" box="[672,855,1286,1313]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="154">ZRC 2021.0350</specimenCode>
, st. VM67,
<collectorName id="154E53058075973F19FE9617FB08FACF" box="[983,1185,1286,1312]" pageId="2" pageNumber="154">SE Baldwin Cove</collectorName>
,
<location id="BD6460088075973F1E859617FA31FACF" LSID="urn:lsid:plazi:treatment:301287C5807597391ABE973CFBAFF856:BD6460088075973F1E859617FA31FACF" box="[1196,1432,1286,1312]" country="Vanuatu" name="Espiritu Santo Island" pageId="2" pageNumber="154">Espiritu Santo Island</location>
,
<collectingCountry id="C0AC76438075973F1ABE963BFF5FFAAB" box="[151,246,1322,1348]" name="Vanuatu" pageId="2" pageNumber="154">Vanuatu</collectingCountry>
, coll.
<specimenCode id="E81D9EA88075973F1B10963BFE7CFAAB" box="[313,469,1322,1348]" collectionCode="SANTO" pageId="2" pageNumber="154">SANTO 2006</specimenCode>
Marine Biodiversity Survey,
<date id="CC0510138075973F1908963BFC1FFAAB" box="[801,950,1322,1348]" pageId="2" pageNumber="154" value="2006-10-14">
<collectingDate id="DC41E9FB8075973F1908963BFC1FFAAB" box="[801,950,1322,1348]" pageId="2" pageNumber="154" value="2006-10-14">14 Oct. 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F0A165588075973F1AEE965CFAADF8F7" pageId="2" pageNumber="154" type="diagnosis">
<paragraph id="B80436D38075973F1AEE965CFAADF8F7" blockId="2.[151,1437,1177,2032]" pageId="2" pageNumber="154">
<emphasis id="8ACFEAC18075973F1AEE965CFEEBFA88" bold="true" box="[199,322,1357,1383]" pageId="2" pageNumber="154">Diagnosis.</emphasis>
Carapace (
<figureCitation id="20802A568075973F1BE9965FFDECFA87" box="[448,581,1358,1384]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="2" pageNumber="154">Figs. 1A, C</figureCitation>
,
<figureCitation id="20802A568075973F1878965FFDDCFA87" box="[593,629,1358,1384]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">2A</figureCitation>
) subquadrate, 1.161.19 times as wide as long; dorsal surface glabrous, hepatic region covered with small granules; postfrontal region distinct, separated into two lobes by shallow grooves; front broad, frontal margin almost straight, lined with tiny, rounded granules; anterolateral margins with 2 indistinct teeth behind broad, bluntly triangular external orbital tooth. Third maxillipeds (
<figureCitation id="20802A568075973F1E3C96ABFBC5FA3B" box="[1045,1132,1466,1492]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2B</figureCitation>
) with sparse setae; exopod almost equal in width to ischium. Chelipeds (
<figureCitation id="20802A568075973F18BE96CFFC8CFA17" box="[663,805,1502,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="2" pageNumber="154">Figs. 1 AD</figureCitation>
,
<figureCitation id="20802A568075973F191996CFFCDDFA17" box="[816,884,1502,1528]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">2C, D</figureCitation>
) symmetrical; inner surface of palm glabrous; in male, proximal half of fingers with tufts of long dense soft setae on outer surface, slightly extending to palm side; pulvinus present at base of fingers in male but absent in female (
<figureCitation id="20802A568075973F194F9537FC49F9AF" box="[870,992,1574,1600]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2C, D</figureCitation>
). P4 (
<figureCitation id="20802A568075973F1E099537FBDDF9AF" box="[1056,1140,1574,1600]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2E</figureCitation>
) relatively long; propodus with dense short and sparse long setae on distal 1/31/2 of posterior margins; dactylus with dense short setae on almost entire, posterior margin. P5 (
<figureCitation id="20802A568075973F181D957FFD20F967" box="[564,649,1646,1672]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2F</figureCitation>
) relatively short; propodus covered with dense short and sparse long setae on distal 1/41/3 of posterior margins; dactylus with dense short setae each on anterior and posterior margins. Male pleon (
<figureCitation id="20802A568075973F1B0395A7FE2AF93F" box="[298,387,1718,1744]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2G</figureCitation>
) narrow, distal margin of telson not concave nor with tuft of setae. G1 (
<figureCitation id="20802A568075973F1EEF95A7FA9DF93F" box="[1222,1332,1718,1744]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2IL</figureCitation>
) slender, distally curved toward dorsolateral; tip chitinous, with 2 short rounded lobes in lateral view (
<figureCitation id="20802A568075973F1E9395CBFA9CF91B" box="[1210,1333,1754,1780]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2K, L</figureCitation>
), opened toward lateral and mesial, respectively. Vulvae (
<figureCitation id="20802A568075973F189895EFFCA4F8F7" box="[689,781,1790,1816]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">Fig. 2M</figureCitation>
) with 2 short, rounded sternal vulvar covers.
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807597391AEE9430FB78FBFE" lastPageId="4" lastPageNumber="156" pageId="2" pageNumber="154" type="description">
<paragraph id="B80436D38075973F1AEE9430FD75F81F" blockId="2.[151,1437,1177,2032]" pageId="2" pageNumber="154">
<emphasis id="8ACFEAC18075973F1AEE9430FEF0F8D4" bold="true" box="[199,345,1825,1851]" pageId="2" pageNumber="154">Description.</emphasis>
Carapace (
<figureCitation id="20802A568075973F1BF39433FDC8F8D3" box="[474,609,1826,1852]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="2" pageNumber="154">Figs. 1A, C</figureCitation>
,
<figureCitation id="20802A568075973F18469433FD3AF8D3" box="[623,659,1826,1852]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="2" pageNumber="154">2A</figureCitation>
) subquadrate, 1.161.19 times (n=2) as wide as long; dorsal surface flattish, finely punctate, glabrous; regions weakly defined, with noticeable groove between epigastric regions; hepatic region (especially part near anterolateral margin) covered with small granules; metabranchial region sloping outwards. Front broad, slightly convex near orbital regions; frontal margin almost straight, not divided into any distinct lobes, lined with small, rounded granules; postfrontal region distinct, separated into 2 lobes by shallow grooves, margin of lobe lined with minute granules.
</paragraph>
<paragraph id="B80436D38074973E1AEE9386FC6CFEF2" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="155">Supraorbital margins lined with small granules. Anterolateral margins granulated, with 2 indistinct teeth behind external orbital tooth; external orbital tooth largest, broad, bluntly triangular, slightly sloping forward. Posterolateral margins slightly convergent posteriorly; posterolateral regions almost glabrous. Infraorbital ridge consisting of several small, rounded granules. Epistome broad, median part triangular.</paragraph>
<paragraph id="B80436D38074973E1AEE9236FD13FE42" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="155">
Third maxillipeds (
<figureCitation id="20802A568074973E1B8B9236FE53FEAE" box="[418,506,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="3" pageNumber="155">Fig. 2B</figureCitation>
) broad, with sparse short setae (numerous on exopod) on external surface; merus with oblique shallow groove along mesial margin of external surface, anterolateral angle broadly rounded, slightly sloping laterally; ischium with an indistinct shallow groove slightly mesial to midline of external surface; exopod broad, convex, almost equal in width to ischium.
</paragraph>
<caption id="ECC4665B8074973E1ABE95BAFE5AF8E3" ID-DOI="http://doi.org/10.5281/zenodo.12724144" ID-Zenodo-Dep="12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="3" pageNumber="155" startId="3.[151,250,1707,1731]" targetBox="[151,1436,450,1681]" targetPageId="3" targetType="figure">
<paragraph id="B80436D38074973E1ABE95BAFE5AF8E3" blockId="3.[151,1436,1707,1804]" pageId="3" pageNumber="155">
<emphasis id="8ACFEAC18074973E1ABE95BAFEB9F92B" bold="true" box="[151,272,1707,1732]" pageId="3" pageNumber="155">FIGURE 1</emphasis>
. AD,
<taxonomicName id="7FBB4D508074973E1B4995BDFDE1F92B" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[352,584,1708,1732]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="155" phylum="Arthropoda" rank="species" species="amikee" status="sp. nov.">
<emphasis id="8ACFEAC18074973E1B4995BDFDE1F92B" box="[352,584,1708,1732]" italics="true" pageId="3" pageNumber="155">Ptychognathus amikee</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91FC57BA8074973E187B95BDFD0EF92C" box="[594,679,1708,1732]" pageId="3" pageNumber="155" rank="species">
<emphasis id="8ACFEAC18074973E187B95BDFD08F92B" bold="true" box="[594,673,1708,1732]" pageId="3" pageNumber="155">sp. nov</emphasis>
.
</taxonomicNameLabel>
; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF,
<taxonomicName id="7FBB4D508074973E188995C1FC36F907" authorityName="N. K. Ng" authorityYear="2010" box="[672,927,1744,1768]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="155" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="8ACFEAC18074973E188995C1FC9EF907" box="[672,823,1744,1768]" italics="true" pageId="3" pageNumber="155">Ptychognathus</emphasis>
aff.
<emphasis id="8ACFEAC18074973E194E95C1FC36F907" box="[871,927,1744,1768]" italics="true" pageId="3" pageNumber="155">lipkei</emphasis>
</taxonomicName>
, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view.
</paragraph>
</caption>
<paragraph id="B80436D3807497391AEE942BFC76FEF2" blockId="3.[151,1437,1850,2020]" lastBlockId="4.[151,1437,150,1977]" lastPageId="4" lastPageNumber="156" pageId="3" pageNumber="155">
Chelipeds (
<figureCitation id="20802A568074973E1B63942BFE77F8BB" box="[330,478,1850,1876]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="3" pageNumber="155">Figs. 1 AD</figureCitation>
,
<figureCitation id="20802A568074973E1BC5942BFD9AF8BB" box="[492,563,1850,1876]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="3" pageNumber="155">2C, D</figureCitation>
) symmetrical in male and female, stronger in male. Merus without spines, dorsal and inner margins proximally with dense soft setae, ventral margins glabrous. Carpus almost glabrous on surface, only with several tiny granules; inner distal angle blunt, without any spines. In male, palm (
<figureCitation id="20802A568074973E1EDF9493FAE2F873" box="[1270,1355,1922,1948]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="3" pageNumber="155">Fig. 2C</figureCitation>
) finely punctate, without distinct granules on outer surface; inner surface glabrous, slightly convex medially. Movable finger (dactylus) approximately as long as palm, cutting edge with 810 small blunt teeth; immovable and movable fingers almost equal in length, cutting edge with 4 or 5 large blunt teeth; proximal halves of movable and immovable fingers with long dense soft setae, slightly extending to palm side, pulvinus present at base of fingers. In female, outer surface of palm (
<figureCitation id="20802A56807397391B8893CEFE52FF16" box="[417,507,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2D</figureCitation>
) glabrous, with prominent ridge consisting of small granules running parallel to ventral margin; inner surface almost glabrous, only with sparse short setae.
</paragraph>
<paragraph id="B80436D3807397391AEE9236FBF3FD1E" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
Ambulatory legs (
<figureCitation id="20802A56807397391BBB9236FDBEFEAE" box="[402,535,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="4" pageNumber="156">Figs. 1AD</figureCitation>
) slender, P3 and P4 equal in length, longest; setation not different between male and female. Meri without spines, but with long soft setae on proximal half of anterior margin; posterior margins almost glabrous; lateral dorsal surfaces glabrous. P2 relatively short; propodus covered with dense short setae on anterior margin and ventral surface; dorsal surface glabrous, anterior margins of distal half covered with dense short and sparse long setae; dactylus covered with rows of short setae on margins and surface. P3 and P4 (
<figureCitation id="20802A56807397391ECC92A6FAC0FE3E" box="[1253,1385,439,465]" captionStart="FIGURE 1" captionStartId="3.[151,250,1707,1731]" captionTargetBox="[151,1436,450,1681]" captionTargetId="figure-126@3.[151,1436,450,1681]" captionTargetPageId="3" captionText="FIGURE 1. AD, Ptychognathus amikee sp. nov.; AB, male holotype (15.8 × 13.6 mm, MNHN-B29852); CD, female paratype (17.4 × 14.6 mm, MNHN-B29853); EF, Ptychognathus aff. lipkei, male (11.4 × 9.6 mm, ZRC 2023.0250); A, C, E, dorsal view; B, D, F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.12724144" httpUri="https://zenodo.org/record/12724144/files/figure.png" pageId="4" pageNumber="156">Figs. 1AD</figureCitation>
,
<figureCitation id="20802A56807397391F5D92A6FA3DFE3E" box="[1396,1428,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">2E</figureCitation>
) relatively long; meri ca. 2.7 as long as wide (
<typeStatus id="670088718073973918BB92CAFD53FE1A" box="[658,762,475,501]" pageId="4" pageNumber="156" type="holotype">holotype</typeStatus>
); carpi glabrous on margins and surfaces; propodi ca. 1.9 as long as wide (
<typeStatus id="67008871807397391B0692EEFE3EFDF6" box="[303,407,511,537]" pageId="4" pageNumber="156" type="holotype">holotype</typeStatus>
), glabrous on anterior margin and dorsal surface, part near posterior margin of ventral surface covered with dense short setae in P3 but almost glabrous in P4, posterior margins with dense short and sparse long setae on distal 1/31/2; dactyli equal to propodi in length, with dense short setae on anterior and posterior margins. P5 (
<figureCitation id="20802A56807397391AEC917AFEB3FD6A" box="[197,282,619,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2F</figureCitation>
) relatively short; merus ca. 2.6 as long as wide (
<typeStatus id="6700887180739739196F917AFC07FD6A" box="[838,942,619,645]" pageId="4" pageNumber="156" type="holotype">holotype</typeStatus>
); carpus ca. 1.3 as long as wide (
<typeStatus id="67008871807397391F01917AFA39FD6A" box="[1320,1424,619,645]" pageId="4" pageNumber="156" type="holotype">holotype</typeStatus>
), glabrous on margins and surfaces; propodus covered with dense short setae near distal of dorsal surface (denser on distal part), posterior margin with dense short setae and sparse long setae on distal 1/41/3; dactylus ca. 0.8 times as long as propodus (
<typeStatus id="67008871807397391B4C91C6FE64FD1E" box="[357,461,727,753]" pageId="4" pageNumber="156" type="holotype">holotype</typeStatus>
), with dense short setae on anterior and posterior margins.
</paragraph>
<paragraph id="B80436D3807397391AEE91EAFADDFC6E" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
Male pleon (
<figureCitation id="20802A56807397391B7191EAFE19FCFA" box="[344,432,763,789]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2G</figureCitation>
) narrow; external surface smooth, without any granules; lateral margins lined with short setae; telson tongue-shaped, significantly longer and narrower than sixth pleonal segment, distal margin of telson not concave, without tuft of setae. Female pleon (
<figureCitation id="20802A568073973918FA9052FC85FCB2" box="[723,812,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2H</figureCitation>
) wide; external surface smooth, without any granules; lateral margins lined with short setae; telson bluntly triangular, almost equal to sixth pleonal segment in length.
</paragraph>
<paragraph id="B80436D3807397391AEE909AFF77FC02" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
Male G1 (
<figureCitation id="20802A56807397391B17909AFE05FC4A" box="[318,428,907,933]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2IL</figureCitation>
) slender, slightly curved, distally curved toward dorsolateral; tip chitinous, with 2 short rounded lobes in lateral view (
<figureCitation id="20802A56807397391BCC90BEFDF2FC26" box="[485,603,943,969]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2K, L</figureCitation>
), opened toward lateral and mesial, respectively; G2 shorter than 1/4 length of G1.
</paragraph>
<paragraph id="B80436D3807397391AEE90E6FB78FBFE" blockId="4.[151,1437,150,1977]" box="[199,1233,1015,1041]" pageId="4" pageNumber="156">
Female vulvae (
<figureCitation id="20802A56807397391B5290E6FE7EFBFE" box="[379,471,1015,1041]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2M</figureCitation>
) with 2 short, rounded sternal vulvar covers; sunken on mesial part.
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807397391AEE970AFAF6FB4E" pageId="4" pageNumber="156" type="etymology">
<paragraph id="B80436D3807397391AEE970AFDEEFB92" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
<emphasis id="8ACFEAC1807397391AEE970AFEE7FBDA" bold="true" box="[199,334,1051,1077]" pageId="4" pageNumber="156">Etymology.</emphasis>
The new species is named after the late Dr. Ngan Kee Ng for her contribution to the taxonomy of brachyurans, especially the family
<taxonomicName id="7FBB4D5080739739183C972EFD2EFBB6" authorityName="H. Milne Edwards" authorityYear="1853" box="[533,647,1087,1113]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
(cf.
<bibRefCitation id="DC2A4B2280739739189F972EFC2DFBB6" author="Lee, B. Y. &amp; Jaafar, Z." box="[694,900,1087,1113]" pageId="4" pageNumber="156" pagination="1255 - 1264" refId="ref9268" refString="Lee, B. Y. &amp; Jaafar, Z. (2022) In memoriam Ng Ngan Kee (1966 - 2022). Crustaceana, 95, 1255 - 1264. https: // doi. org / 10.1163 / 15685403 - bja 10229" type="journal article" year="2022">Lee &amp; Jaafar 2022</bibRefCitation>
). The name
<taxonomicName id="7FBB4D50807397391E2F972EFBCFFBB6" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1030,1126,1087,1113]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee">“amikee</taxonomicName>
” is an arbitrary combination of friend (amicus or amica) and Kee.
</paragraph>
<paragraph id="B80436D3807397391AEE9797FAF6FB4E" blockId="4.[151,1437,150,1977]" box="[199,1375,1158,1185]" pageId="4" pageNumber="156">
<emphasis id="8ACFEAC1807397391AEE9797FF57FB4F" bold="true" box="[199,254,1158,1184]" pageId="4" pageNumber="156">Size.</emphasis>
Largest male 15.8 ×
<quantity id="7F439B36807397391BC59796FDF9FB4E" box="[492,592,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="4" pageNumber="156" unit="mm" value="13.6">13.6 mm</quantity>
(MNHN-B29852); largest female 17.4 ×
<quantity id="7F439B36807397391E0A9796FB2EFB4E" box="[1059,1159,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="4" pageNumber="156" unit="mm" value="14.6">14.6 mm</quantity>
(MNHN-B29853).
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807397391AEE97BBFCBDFB2A" box="[199,788,1194,1221]" pageId="4" pageNumber="156" type="distribution">
<paragraph id="B80436D3807397391AEE97BBFCBDFB2A" blockId="4.[151,1437,150,1977]" box="[199,788,1194,1221]" pageId="4" pageNumber="156">
<emphasis id="8ACFEAC1807397391AEE97BBFEC8FB2B" bold="true" box="[199,353,1194,1220]" pageId="4" pageNumber="156">Distribution.</emphasis>
At present, only known from
<collectingCountry id="C0AC764380739739189897BAFCB8FB2A" box="[689,785,1195,1221]" name="Vanuatu" pageId="4" pageNumber="156">Vanuatu</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F0A16558807397391AEE97DEFBAFF856" pageId="4" pageNumber="156" type="discussion">
<paragraph id="B80436D3807397391AEE97DEFB12FA96" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
<emphasis id="8ACFEAC1807397391AEE97DEFE92FB06" bold="true" box="[199,315,1231,1257]" pageId="4" pageNumber="156">Remarks.</emphasis>
<taxonomicName id="7FBB4D50807397391B6B97DEFD96FB06" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[322,575,1231,1257]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee" status="sp. nov.">
<emphasis id="8ACFEAC1807397391B6B97DEFD96FB06" box="[322,575,1231,1257]" italics="true" pageId="4" pageNumber="156">Ptychognathus amikee</emphasis>
</taxonomicName>
<emphasis id="8ACFEAC180739739186E97DEFD08FB06" bold="true" box="[583,673,1231,1257]" pageId="4" pageNumber="156">
<taxonomicNameLabel id="91FC57BA80739739186E97DEFD08FB06" box="[583,673,1231,1257]" pageId="4" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to
<taxonomicName id="7FBB4D508073973919DC97DEFB58FB06" authority="Roux, 1917" authorityName="Roux" authorityYear="1917" box="[1013,1265,1231,1257]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="8ACFEAC18073973919DC97DEFBCAFB06" box="[1013,1123,1231,1257]" italics="true" pageId="4" pageNumber="156">P. demani</emphasis>
<bibRefCitation id="DC2A4B22807397391E4297DEFB58FB06" author="Roux, J." box="[1131,1265,1231,1257]" pageId="4" pageNumber="156" pagination="589 - 621" refId="ref10192" refString="Roux, J. (1917) Crustaces. Resultats de l'expedition scientifique neerlandaise a la Nouvelle Guinea. Nova Guinea, No 5, 589 - 621 pls. 27 + 28." type="journal article" year="1917">Roux, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7FBB4D50807397391ED597DEFEE4FAE2" authority="(De Man, 1879)" baseAuthorityName="De Man" baseAuthorityYear="1879" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="8ACFEAC1807397391ED597DEFA35FB06" box="[1276,1436,1231,1257]" italics="true" pageId="4" pageNumber="156">P. intermedius</emphasis>
(
<bibRefCitation id="DC2A4B22807397391AB697E2FEECFAE2" author="De Man, J. G." box="[159,325,1267,1293]" pageId="4" pageNumber="156" pagination="53 - 73" refId="ref8467" refString="De Man, J. G. (1879) On some new or imperfectly known Podophthalmous Crustacea of the Leyden Museum. Notes Leyden Museum, 1, 53 - 73." type="journal article" year="1879">De Man, 1879</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="7FBB4D50807397391BA197E2FCB7FAE2" authority="(A. Milne-Edwards, 1868)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1868" box="[392,798,1267,1293]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="pilipes">
<emphasis id="8ACFEAC1807397391BA197E2FE46FAE2" box="[392,495,1267,1293]" italics="true" pageId="4" pageNumber="156">P. pilipes</emphasis>
(A.
<bibRefCitation id="DC2A4B2280739739180897E2FCBFFAE2" author="Milne-Edwards, A." box="[545,790,1267,1293]" pageId="4" pageNumber="156" pagination="173 - 185" refId="ref9464" refString="Milne-Edwards, A. (1868) Etudes zoologiques sur quelques Crustaces des iles Celebes provenant envoi de M. Riedel. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4, 173 - 185. https: // doi. org / 10.5962 / bhl. title. 10260" type="journal article" year="1868">Milne-Edwards, 1868</bibRefCitation>
)
</taxonomicName>
in the blunt anterolateral teeth and the postfrontal region of carapace with separated into two distinct lobes, but can be distinguished by the other features of carapace, and the distribution of long setae on male chelipedal palm and fingers, and ambulatory legs in both sexes. However, the character of G1 among those species compared is very similar and not useful to separate them.
</paragraph>
<paragraph id="B80436D3807397391AEE9692FDE3F8EA" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
<taxonomicName id="7FBB4D50807397391AEE9692FE6CFA72" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[199,453,1411,1437]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee" status="sp. nov.">
<emphasis id="8ACFEAC1807397391AEE9692FE6CFA72" box="[199,453,1411,1437]" italics="true" pageId="4" pageNumber="156">Ptychognathus amikee</emphasis>
</taxonomicName>
<emphasis id="8ACFEAC1807397391BE79692FD80FA72" bold="true" box="[462,553,1411,1437]" pageId="4" pageNumber="156">
<taxonomicNameLabel id="91FC57BA807397391BE79692FD80FA72" box="[462,553,1411,1437]" pageId="4" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and other three species differ in male chelipedal palm and immovable finger, and ambulatory legs in both sexes. The outer surface of male palm is nearly smooth, and the proximal halves of movable and immovable fingers of the cheliped are covered with dense long setae in
<taxonomicName id="7FBB4D50807397391EAC96DAFB5AFA0A" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1157,1267,1483,1509]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee">
<emphasis id="8ACFEAC1807397391EAC96DAFB5AFA0A" box="[1157,1267,1483,1509]" italics="true" pageId="4" pageNumber="156">P. amikee</emphasis>
</taxonomicName>
(
<figureCitation id="20802A56807397391F2F96DAFAC9FA0A" box="[1286,1376,1483,1509]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2C</figureCitation>
) [vs. setae present only on the proximal half of immovable finger in
<taxonomicName id="7FBB4D5080739739194296FEFC70F9E6" authorityName="Roux" authorityYear="1917" box="[875,985,1519,1545]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="8ACFEAC180739739194296FEFC70F9E6" box="[875,985,1519,1545]" italics="true" pageId="4" pageNumber="156">P. demani</emphasis>
</taxonomicName>
(
<bibRefCitation id="DC2A4B228073973919C396FEFB0FF9E6" author="Roux, J." box="[1002,1190,1518,1545]" pageId="4" pageNumber="156" pagination="589 - 621" refId="ref10192" refString="Roux, J. (1917) Crustaces. Resultats de l'expedition scientifique neerlandaise a la Nouvelle Guinea. Nova Guinea, No 5, 589 - 621 pls. 27 + 28." type="journal article" year="1917">Roux 1917: 615</bibRefCitation>
, pl. 18 fig. 20); small granules present and no dense setae in
<taxonomicName id="7FBB4D508073973918639502FD40F9C2" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[586,745,1555,1581]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="8ACFEAC18073973918639502FD40F9C2" box="[586,745,1555,1581]" italics="true" pageId="4" pageNumber="156">P. intermedius</emphasis>
</taxonomicName>
(
<bibRefCitation id="DC2A4B228073973918D19502FC68F9C2" author="De Man, J. G." box="[760,961,1555,1581]" pageId="4" pageNumber="156" pagination="53 - 73" refId="ref8467" refString="De Man, J. G. (1879) On some new or imperfectly known Podophthalmous Crustacea of the Leyden Museum. Notes Leyden Museum, 1, 53 - 73." type="journal article" year="1879">De Man 1879: 69</bibRefCitation>
;
<bibRefCitation id="DC2A4B228073973919E49502FBC2F9C2" author="De Man, J. G." box="[973,1131,1555,1581]" pageId="4" pageNumber="156" pagination="15 - 29" refId="ref8500" refString="De Man, J. G. (1892) Decapoden des indischen Archipels. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Vol. 2. Brill, Leiden, pp. 265 - 527, pls. 15 - 29. https: // doi. org / 10.5962 / bhl. title. 10298" type="book chapter" year="1892">De Man 1892</bibRefCitation>
: tab. 19, fig. 10a); granules and glabrous in
<taxonomicName id="7FBB4D50807397391B629526FE1BF9BE" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1868" box="[331,434,1591,1617]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="pilipes">
<emphasis id="8ACFEAC1807397391B629526FE1BF9BE" box="[331,434,1591,1617]" italics="true" pageId="4" pageNumber="156">P. pilipes</emphasis>
</taxonomicName>
(A.
<bibRefCitation id="DC2A4B22807397391BCD9526FCBDF9BE" author="Milne-Edwards, A." box="[484,788,1591,1617]" pageId="4" pageNumber="156" pagination="173 - 185" refId="ref9464" refString="Milne-Edwards, A. (1868) Etudes zoologiques sur quelques Crustaces des iles Celebes provenant envoi de M. Riedel. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4, 173 - 185. https: // doi. org / 10.5962 / bhl. title. 10260" type="journal article" year="1868">Milne-Edwards, 1868: 184</bibRefCitation>
, pl. 27 (9))]. The anterior margins of carpi and propodi of ambulatory legs in both sexes are almost glabrous and only distal part of P5 carpus has dense short setae in
<taxonomicName id="7FBB4D50807397391F1A954AFA35F99A" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1331,1436,1627,1653]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee">
<emphasis id="8ACFEAC1807397391F1A954AFA35F99A" box="[1331,1436,1627,1653]" italics="true" pageId="4" pageNumber="156">P. amikee</emphasis>
</taxonomicName>
(
<figureCitation id="20802A56807397391AB6956EFEBAF976" box="[159,275,1663,1689]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2E, F</figureCitation>
) [vs. anterior margins of carpi and propodi with dense short setae in
<taxonomicName id="7FBB4D50807397391E33956EFB21F976" authorityName="Roux" authorityYear="1917" box="[1050,1160,1663,1689]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="8ACFEAC1807397391E33956EFB21F976" box="[1050,1160,1663,1689]" italics="true" pageId="4" pageNumber="156">P. demani</emphasis>
</taxonomicName>
(
<bibRefCitation id="DC2A4B22807397391EBE956EFAF8F976" author="Roux, J." box="[1175,1361,1662,1689]" pageId="4" pageNumber="156" pagination="589 - 621" refId="ref10192" refString="Roux, J. (1917) Crustaces. Resultats de l'expedition scientifique neerlandaise a la Nouvelle Guinea. Nova Guinea, No 5, 589 - 621 pls. 27 + 28." type="journal article" year="1917">Roux 1917: 615</bibRefCitation>
, pl. 18 fig. 18); anterior and posterior margins of propodi with dense long setae in
<taxonomicName id="7FBB4D508073973919D895B2FB3BF952" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[1009,1170,1699,1725]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="8ACFEAC18073973919D895B2FB3BF952" box="[1009,1170,1699,1725]" italics="true" pageId="4" pageNumber="156">P. intermedius</emphasis>
</taxonomicName>
(
<bibRefCitation id="DC2A4B22807397391E8B95B2FAC6F952" author="De Man, J. G." box="[1186,1391,1699,1725]" pageId="4" pageNumber="156" pagination="53 - 73" refId="ref8467" refString="De Man, J. G. (1879) On some new or imperfectly known Podophthalmous Crustacea of the Leyden Museum. Notes Leyden Museum, 1, 53 - 73." type="journal article" year="1879">De Man 1879: 70</bibRefCitation>
;
<bibRefCitation id="DC2A4B22807397391F5595B2FEEFF90E" author="De Man, J. G." pageId="4" pageNumber="156" pagination="15 - 29" refId="ref8500" refString="De Man, J. G. (1892) Decapoden des indischen Archipels. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Vol. 2. Brill, Leiden, pp. 265 - 527, pls. 15 - 29. https: // doi. org / 10.5962 / bhl. title. 10298" type="book chapter" year="1892">De Man 1892: 323</bibRefCitation>
); anterior and posterior margins of carpi, propodi and dactyli with dense long setae in
<taxonomicName id="7FBB4D50807397391F2295D6FAD8F90E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1868" box="[1291,1393,1735,1761]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="pilipes">
<emphasis id="8ACFEAC1807397391F2295D6FAD8F90E" box="[1291,1393,1735,1761]" italics="true" pageId="4" pageNumber="156">P. pilipes</emphasis>
</taxonomicName>
(A.
<bibRefCitation id="DC2A4B22807397391ABE95FAFE6FF8EA" author="Milne-Edwards, A." box="[151,454,1771,1797]" pageId="4" pageNumber="156" pagination="173 - 185" refId="ref9464" refString="Milne-Edwards, A. (1868) Etudes zoologiques sur quelques Crustaces des iles Celebes provenant envoi de M. Riedel. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4, 173 - 185. https: // doi. org / 10.5962 / bhl. title. 10260" type="journal article" year="1868">Milne-Edwards, 1868: 184</bibRefCitation>
, pl. 27 (6)].
</paragraph>
<paragraph id="B80436D3807397391AEE941EFBAFF856" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="156">
This new species is also similar to
<taxonomicName id="7FBB4D50807397391863941EFD63F8C6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[586,714,1807,1833]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="8ACFEAC1807397391863941EFD63F8C6" box="[586,714,1807,1833]" italics="true" pageId="4" pageNumber="156">P. barbatus</emphasis>
</taxonomicName>
,
<taxonomicName id="7FBB4D508073973918FE941EFCD7F8C6" authorityName="Sakai" authorityYear="1955" box="[727,894,1807,1833]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="hachijoensis">
<emphasis id="8ACFEAC18073973918FE941EFCD7F8C6" box="[727,894,1807,1833]" italics="true" pageId="4" pageNumber="156">P. hachijoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="7FBB4D508073973919A5941EFC57F8C6" authorityName="Heller" authorityYear="1865" box="[908,1022,1807,1833]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="8ACFEAC18073973919A5941EFC57F8C6" box="[908,1022,1807,1833]" italics="true" pageId="4" pageNumber="156">P. pusillus</emphasis>
</taxonomicName>
,
<taxonomicName id="7FBB4D50807397391E25941EFBC3F8C6" authorityName="Hsu, Shih &amp; Li" authorityYear="2022" box="[1036,1130,1807,1833]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="8ACFEAC1807397391E25941EFBC3F8C6" box="[1036,1130,1807,1833]" italics="true" pageId="4" pageNumber="156">P. sakaii</emphasis>
</taxonomicName>
, and
<taxonomicName id="7FBB4D50807397391E8F941EFA84F8C7" authorityName="Hsu &amp; Shih" authorityYear="2020" box="[1190,1325,1807,1832]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="stimpsoni">
<emphasis id="8ACFEAC1807397391E8F941EFA84F8C7" box="[1190,1325,1807,1832]" italics="true" pageId="4" pageNumber="156">P. stimpsoni</emphasis>
</taxonomicName>
in having similar distribution of setae in male chelipedal palm and fingers, i.e., proximal halves of both fingers with long setae, but can be separated by the frontal margin of carapace; it is almost straight in
<taxonomicName id="7FBB4D50807397391E7C9446FB68F89E" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1109,1217,1879,1905]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="156" phylum="Arthropoda" rank="species" species="amikee" status="sp. nov.">
<emphasis id="8ACFEAC1807397391E7C9446FB68F89E" box="[1109,1217,1879,1905]" italics="true" pageId="4" pageNumber="156">P. amikee</emphasis>
</taxonomicName>
<emphasis id="8ACFEAC1807397391EE09446FA8DF89E" bold="true" box="[1225,1316,1879,1905]" pageId="4" pageNumber="156">
<taxonomicNameLabel id="91FC57BA807397391EE09446FA8DF89E" box="[1225,1316,1879,1905]" pageId="4" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="20802A56807397391F1D9446FA26F89E" box="[1332,1423,1879,1905]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[181,1405,189,1879]" captionTargetId="figure-22@5.[173,1414,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Ptychognathus amikee sp. nov. AC, EG, IL, male holotype (15.8 × 13.6 mm, MNHN-B29852); D, H, M, female paratype (17.4 × 14.6 mm, MNHN-B29853). A, carapace; B, left third maxilliped; C, outer view of male left cheliped; D, outer view of female left cheliped; E, right P4; F, right P5; G, male pleon; H, female pleon; IL, male right G1 (I, J, dorsal view; K, L, dorsolateral view); M, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724150" httpUri="https://zenodo.org/record/12724150/files/figure.png" pageId="4" pageNumber="156">Fig. 2A</figureCitation>
), instead of concave medially in the other species [A.
<bibRefCitation id="DC2A4B228073973918C2946AFBBCF87A" author="Milne-Edwards, A." box="[747,1045,1915,1941]" pageId="4" pageNumber="156" pagination="155 - 332" refId="ref9515" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie, II. Nouvelles Archives du Museum d'Histoire Naturelle, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards 1873: 136</bibRefCitation>
, pl. 17(4);
<bibRefCitation id="DC2A4B22807397391EBD946AFADAF87A" author="De Man, J. G." box="[1172,1395,1914,1941]" pageId="4" pageNumber="156" pagination="537 - 550" refId="ref8643" refString="De Man, J. G. (1905) On species of Crustacea of the genera Ptychognathus Stimps. and Palaemon Fabr. from Christmas Island. Proceedings of the Zoological Society of London, 1905, 537 - 550." type="book chapter" year="1905">De Man, 1905: 539</bibRefCitation>
, pl. 17(12);
<bibRefCitation id="DC2A4B22807397391AD6948EFE6FF856" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[255,454,1951,1977]" pageId="4" pageNumber="156" pagination="249 - 265" refId="ref10460" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">Hsu &amp; Shih 2020</bibRefCitation>
: figs. 2C, E, 4A;
<bibRefCitation id="DC2A4B228073973918AE948EFCECF856" author="Hsu, J. - W. &amp; Shih, H. - T. &amp; Li, J. - J." box="[647,837,1951,1977]" pageId="4" pageNumber="156" refId="ref8935" refString="Hsu, J. - W., Shih, H. - T. &amp; Li, J. - J. (2022 b) Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan. Nauplius, 30, e 2022002. https: // doi. org / 10.1590 / 2358 - 2936 e 2022002" type="journal volume" year="2022">
Hsu
<emphasis id="8ACFEAC1807397391895948EFD5DF856" box="[700,756,1951,1977]" italics="true" pageId="4" pageNumber="156">et al.</emphasis>
2022b
</bibRefCitation>
: figs. 1, 2A, 3A].
</paragraph>
</subSubSection>
</treatment>
</document>