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<document id="31E742A330B00BE99429C79C8879D9E1" ID-CLB-Dataset="30921" ID-DOI="10.11646/zootaxa.4375.3.9" ID-GBIF-Dataset="ebd24618-d854-4b63-840c-9219f742d58b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1158998" ID-ZooBank="411427AD-0943-4B9E-B482-AA640EE582DA" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1516867533818" checkinUser="plazi" docAuthor="Baňař, Petr &amp; Heiss, Ernst" docDate="2018" docId="CA0F87B6FF9AA576FF1AFA41A2995798" docLanguage="en" docName="zootaxa.4375.3.9.pdf" docOrigin="Zootaxa 4375 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Comorocoris estherineae Baňař &amp; Heiss 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="437" masterDocId="3636FFCEFF9BA572FF8DFE6AA64A5568" masterDocTitle="A new species of Comorocoris from Northern Madagascar (Hemiptera: Heteroptera: Aradidae)" masterLastPageNumber="440" masterPageNumber="433" pageNumber="434" updateTime="1698517122624" updateUser="ExternalLinkService">
<mods:mods id="F28C1661A6056B2B88BD83E4B6291B03" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="131FFF19D9EBCBCBDB8315C9C05D3755">
<mods:title id="D6D5A95686C6B0F91302DAD1F343144D">A new species of Comorocoris from Northern Madagascar (Hemiptera: Heteroptera: Aradidae)</mods:title>
</mods:titleInfo>
<mods:name id="B9D692483A8476813425F827FE3B22A1" type="personal">
<mods:role id="A424C310047A5E81152FC7145A47DEFA">
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<mods:namePart id="32FF0EC5EAB3E7E36602ED3B1D5ECBC6">Baňař, Petr</mods:namePart>
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<mods:namePart id="0CC9336B544091A79745B7641529F563">Heiss, Ernst</mods:namePart>
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<mods:typeOfResource id="9EC1EDE2514D0AB056C79946D4C361C5">text</mods:typeOfResource>
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<mods:titleInfo id="76943F40E6E543FEF95453A6315606BD">
<mods:title id="70244A9C60C69EEC15FD3FF6AD4AC62C">Zootaxa</mods:title>
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<mods:part id="CC536A90B00A335548D524E1B9CD90E3">
<mods:date id="1B6DB471C40389685869DD60D0594975">2018</mods:date>
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<mods:number id="A69549B53EA56C4A8708D50BF373948A">2018-01-25</mods:number>
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<mods:detail id="4FDFEB0361C1979106FC2C5D873D0F52" type="volume">
<mods:number id="DC9468DB140491D70DB534C39E08D24A">4375</mods:number>
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<mods:classification id="AE34F701A6E4D021232B4AD7D392A40B">journal article</mods:classification>
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</mods:mods>
<treatment id="CA0F87B6FF9AA576FF1AFA41A2995798" ID-DOI="http://doi.org/10.5281/zenodo.5954447" ID-GBIF-Taxon="140417866" ID-Zenodo-Dep="5954447" LSID="urn:lsid:plazi:treatment:CA0F87B6FF9AA576FF1AFA41A2995798" httpUri="http://treatment.plazi.org/id/CA0F87B6FF9AA576FF1AFA41A2995798" lastPageId="4" lastPageNumber="437" pageId="1" pageNumber="434">
<subSubSection id="0ABC652BFF9AA573FF1AFA41A7D7510F" pageId="1" pageNumber="434" type="nomenclature">
<paragraph id="421936A0FF9AA573FF1AFA41A463512E" blockId="1.[151,553,1067,1127]" box="[151,553,1067,1094]" pageId="1" pageNumber="434">
<heading id="195181CCFF9AA573FF1AFA41A463512E" bold="true" box="[151,553,1067,1094]" fontSize="11" level="1" pageId="1" pageNumber="434" reason="1">
<emphasis id="70D2EAB2FF9AA573FF1AFA41A463512E" bold="true" box="[151,553,1067,1094]" pageId="1" pageNumber="434">
<taxonomicName id="85A64D23FF9AA573FF1AFA41A78F512D" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[151,453,1067,1093]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF9AA573FF1AFA41A78F512D" bold="true" box="[151,453,1067,1093]" italics="true" pageId="1" pageNumber="434">Comorocoris estherineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BE157C9FF9AA573FE41FA46A463512E" box="[460,553,1068,1094]" pageId="1" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="421936A0FF9AA573FF1AFA24A7D7510F" blockId="1.[151,553,1067,1127]" box="[151,413,1102,1127]" pageId="1" pageNumber="434">
(
<figureCitation id="DA9D2A25FF9AA573FF12FA24A740510F" box="[159,266,1102,1127]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[168,1435,213,1915]" captionTargetId="figure@2.[151,1436,193,1932]" captionTargetPageId="2" captionText="FIGURES 1–7. Comorocoris species, dorsal habitus. 1–2, C. estherineae sp. nov., 1, male holotype, 2, female paratype; 3, 6, C.inexpectatusHeiss &amp; Baňař,2012, 3, male holotype, 6, female paratype; 4, 7, C.glabridorsumHeiss &amp; Baňař,2012, 4, male holotype, 7, female paratype; 5, C. testudiformis Heiss, 1985, female paratype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="1" pageNumber="434">Figs. 1–2</figureCitation>
,
<figureCitation id="DA9D2A25FF9AA573FE9BFA25A708510F" box="[278,322,1103,1127]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="1" pageNumber="434">8–9</figureCitation>
,
<figureCitation id="DA9D2A25FF9AA573FEC3FA25A7DF510F" box="[334,405,1103,1127]" captionStart="FIGURES 13–14" captionStartId="4.[151,264,1507,1530]" captionTargetBox="[151,1436,807,1485]" captionTargetId="figure@4.[151,1436,807,1485]" captionTargetPageId="4" captionText="FIGURES 13–14. Comorocoris estherineae sp. nov., male paratype, scanning electron micrographs, pygophore removed. 13, dorsal view; 14, anterior view. Scale bars in mm." httpUri="https://zenodo.org/record/1159004/files/figure.png" pageId="1" pageNumber="434">13–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0ABC652BFF9AA573FF1AFAFCA46C5353" pageId="1" pageNumber="434" type="materials_examined">
<paragraph id="421936A0FF9AA573FF1AFAFCA46C5353" blockId="1.[151,1436,1174,2027]" pageId="1" pageNumber="434">
<emphasis id="70D2EAB2FF9AA573FF1AFAFCA7F351C7" bold="true" box="[151,441,1174,1199]" pageId="1" pageNumber="434">Type material. Holotype</emphasis>
: male, ‘MDA/
<date id="36181060FF9AA573FDEDFAFCA4AC51C7" box="[608,742,1174,1199]" pageId="1" pageNumber="434" value="2015-01-11">Jan.2015/11</date>
<emphasis id="70D2EAB2FF9AA573FD60FAFCA2BA51C5" bold="true" box="[749,1264,1174,1199]" pageId="1" pageNumber="434">N MADAGASCAR / MONTAGNE D‘ AMBRE</emphasis>
~
<quantity id="855E9B45FF9AA573FA88FAFCA30351C7" box="[1285,1353,1174,1199]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="1" pageNumber="434" unit="m" value="945.0">945m</quantity>
, circuit / „
<emphasis id="70D2EAB2FF9AA573FF3EFAD0A75F51BB" bold="true" box="[179,277,1210,1235]" pageId="1" pageNumber="434">Sommet</emphasis>
“, S12°31’28‘‘E49°09’52‘‘
<emphasis id="70D2EAB2FF9AA573FDC7FAD0A41851BB" bold="true" box="[586,594,1210,1235]" pageId="1" pageNumber="434">/</emphasis>
sifting litter+rotten wood, Winkler app. extr.
<emphasis id="70D2EAB2FF9AA573FBD9FAD0A21651BB" bold="true" box="[1108,1116,1210,1235]" pageId="1" pageNumber="434">/</emphasis>
<date id="36181060FF9AA573FBEEFAD0A29C51BB" box="[1123,1238,1210,1235]" pageId="1" pageNumber="434" value="2015-01-14">14.1.2015</date>
, P. Baňař &amp; E.M. Rabotoson lgt.’ [printed] // ‘HOLOTYPE /
<taxonomicName id="85A64D23FF9AA573FDF0FAB5A55A519E" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[637,784,1247,1270]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="70D2EAB2FF9AA573FDF0FAB5A55A519E" box="[637,784,1247,1270]" italics="true" pageId="1" pageNumber="434">Comorocoris</emphasis>
</taxonomicName>
/
<taxonomicName id="85A64D23FF9AA573FCABFAB4A5ED519E" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[806,935,1246,1270]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF9AA573FCABFAB4A5ED519E" box="[806,935,1246,1270]" italics="true" pageId="1" pageNumber="434">estherineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BE157C9FF9AA573FC23FAB4A242519F" box="[942,1032,1246,1271]" pageId="1" pageNumber="434" rank="species">
<emphasis id="70D2EAB2FF9AA573FC23FAB4A242519F" bold="true" box="[942,1032,1246,1271]" pageId="1" pageNumber="434">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Baňař &amp; Heiss det. 2018’ [printed red label] (MMBC).
<emphasis id="70D2EAB2FF9AA573FE0DFB68A7B25073" bold="true" box="[384,504,1282,1307]" pageId="1" pageNumber="434">Paratypes</emphasis>
: 3 ♂, 5 ♀, same locality label as holotype [one male gold-coated for SEM] (2 ♂, 2 ♀ MMBC, 1 ♂, 3 ♀ CEHI); 3 ♂: ‘MDA/
<date id="36181060FF9AA573FDE9FB4CA4A65056" box="[612,748,1318,1342]" pageId="1" pageNumber="434" value="2015-01-08">Jan.2015/08</date>
<emphasis id="70D2EAB2FF9AA573FD7EFB4CA2B95055" bold="true" box="[755,1267,1318,1343]" pageId="1" pageNumber="434">N MADAGASCAR / MONTAGNE D‘ AMBRE</emphasis>
<quantity id="855E9B45FF9AA573FB77FB4CA3035057" box="[1274,1353,1318,1343]" metricMagnitude="3" metricUnit="m" metricValue="1.169" pageId="1" pageNumber="434" unit="m" value="1169.0">1169m</quantity>
, circuit
<emphasis id="70D2EAB2FF9AA573FF1AFB20A6D5500B" bold="true" box="[151,159,1354,1379]" pageId="1" pageNumber="434">/</emphasis>
„
<emphasis id="70D2EAB2FF9AA573FF38FB20A75D500B" bold="true" box="[181,279,1354,1379]" pageId="1" pageNumber="434">Sommet</emphasis>
“, S12°31’52‘‘E49°10’17‘‘
<emphasis id="70D2EAB2FF9AA573FDDEFB20A411500B" bold="true" box="[595,603,1354,1379]" pageId="1" pageNumber="434">/</emphasis>
sifting
<taxonomicName id="85A64D23FF9AA573FD3AFB21A56D500B" box="[695,807,1355,1379]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="434" phylum="Tracheophyta" rank="genus">
<emphasis id="70D2EAB2FF9AA573FD3AFB21A56D500B" box="[695,807,1355,1379]" italics="true" pageId="1" pageNumber="434">Pandanus</emphasis>
</taxonomicName>
litter, Winkler app. extr.
<emphasis id="70D2EAB2FF9AA573FBDFFB20A210500B" bold="true" box="[1106,1114,1354,1379]" pageId="1" pageNumber="434">/</emphasis>
<date id="36181060FF9AA573FBE9FB20A283500B" box="[1124,1225,1354,1379]" pageId="1" pageNumber="434" value="2015-01-08">8.1.2015</date>
, P. Baňař &amp; E.M. Rabotoson lgt.’ (1 ♂ MMBC, 2 ♂ CEHI); 1 ♀: ‘MDA/
<date id="36181060FF9AA573FC8AFB04A5DA50EE" box="[775,912,1390,1414]" pageId="1" pageNumber="434" value="2015-01-09">Jan.2015/09</date>
<emphasis id="70D2EAB2FF9AA573FC1AFB04A3D650ED" bold="true" box="[919,1436,1390,1415]" pageId="1" pageNumber="434">N MADAGASCAR / MONTAGNE D‘ AMBRE</emphasis>
~
<quantity id="855E9B45FF9AA573FF28FBF9A6BF50C3" box="[165,245,1427,1451]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="434" unit="m" value="1120.0">1120m</quantity>
, circuit
<emphasis id="70D2EAB2FF9AA573FEDFFBF8A71050C3" bold="true" box="[338,346,1426,1451]" pageId="1" pageNumber="434">/</emphasis>
„
<emphasis id="70D2EAB2FF9AA573FEE2FBF8A79B50C3" bold="true" box="[367,465,1426,1451]" pageId="1" pageNumber="434">Sommet</emphasis>
“, S12°32’14‘‘E49°10’28‘‘
<emphasis id="70D2EAB2FF9AA573FC85FBF8A55A50C3" bold="true" box="[776,784,1426,1451]" pageId="1" pageNumber="434">/</emphasis>
sifting
<taxonomicName id="85A64D23FF9AA573FCE5FBF9A59250C3" box="[872,984,1427,1451]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="434" phylum="Tracheophyta" rank="genus">
<emphasis id="70D2EAB2FF9AA573FCE5FBF9A59250C3" box="[872,984,1427,1451]" italics="true" pageId="1" pageNumber="434">Pandanus</emphasis>
</taxonomicName>
litter, Winkler app. extr.
<emphasis id="70D2EAB2FF9AA573FB75FBF8A34A50C3" bold="true" box="[1272,1280,1426,1451]" pageId="1" pageNumber="434">/</emphasis>
<date id="36181060FF9AA573FA84FBF8A33650C3" box="[1289,1404,1426,1451]" pageId="1" pageNumber="434" value="2015-12-01">12.1.2015</date>
, P. Baňař &amp; E.M. Rabotoson lgt.’ (MMBC); 1 ♂: ‘MDA/
<date id="36181060FF9AA573FC8EFBDCA5C050A6" box="[771,906,1462,1486]" pageId="1" pageNumber="434" value="2015-01-12">Jan.2015/12</date>
<emphasis id="70D2EAB2FF9AA573FC1EFBDCA3D650A5" bold="true" box="[915,1436,1462,1487]" pageId="1" pageNumber="434">N MADAGASCAR / MONTAGNE D‘ AMBRE</emphasis>
~
<quantity id="855E9B45FF9AA573FF29FBB1A6B9509B" box="[164,243,1499,1523]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="434" unit="m" value="1100.0">1100m</quantity>
<emphasis id="70D2EAB2FF9AA573FF76FBB0A749509B" bold="true" box="[251,259,1498,1523]" pageId="1" pageNumber="434">/</emphasis>
sifting litter close to camp,
<date id="36181060FF9AA573FDCEFBB0A4F9509B" box="[579,691,1498,1523]" pageId="1" pageNumber="434" value="2015-01-16">16.1.2015</date>
<emphasis id="70D2EAB2FF9AA573FD36FBB0A489509B" bold="true" box="[699,707,1498,1523]" pageId="1" pageNumber="434">/</emphasis>
Winkler apparatus extraction
<emphasis id="70D2EAB2FF9AA573FB94FBB0A26B509B" bold="true" box="[1049,1057,1498,1523]" pageId="1" pageNumber="434">/</emphasis>
P. Baňař &amp; E.M. Rabotoson lgt.’ (MMBC). All paratypes are provided with a label: ‘PARATYPE /
<taxonomicName id="85A64D23FF9AA573FC03FB95A26B537E" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[910,1057,1535,1558]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="70D2EAB2FF9AA573FC03FB95A26B537E" box="[910,1057,1535,1558]" italics="true" pageId="1" pageNumber="434">Comorocoris</emphasis>
</taxonomicName>
/
<taxonomicName id="85A64D23FF9AA573FBB6FB94A2F6537E" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[1083,1212,1534,1558]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF9AA573FBB6FB94A2F6537E" box="[1083,1212,1534,1558]" italics="true" pageId="1" pageNumber="434">estherineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BE157C9FF9AA573FB48FB94A36B537F" box="[1221,1313,1534,1559]" pageId="1" pageNumber="434" rank="species">
<emphasis id="70D2EAB2FF9AA573FB48FB94A36B537F" bold="true" box="[1221,1313,1534,1559]" pageId="1" pageNumber="434">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Baňař &amp; Heiss det. 2018’ [printed red label].
</paragraph>
</subSubSection>
<subSubSection id="0ABC652BFF9AA573FF4AF82CA37953A3" pageId="1" pageNumber="434" type="diagnosis">
<paragraph id="421936A0FF9AA573FF4AF82CA37953A3" blockId="1.[151,1436,1174,2027]" pageId="1" pageNumber="434">
<emphasis id="70D2EAB2FF9AA573FF4AF82CA70B5337" bold="true" box="[199,321,1606,1631]" pageId="1" pageNumber="434">Diagnosis.</emphasis>
The general aspect of the new species resembles that of
<taxonomicName id="85A64D23FF9AA573FC69F82DA2D15336" authorityName="Heiss" authorityYear="1985" box="[996,1179,1606,1630]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="testudiformis">
<emphasis id="70D2EAB2FF9AA573FC69F82DA2D15336" box="[996,1179,1606,1630]" italics="true" pageId="1" pageNumber="434">C. testudiformis</emphasis>
</taxonomicName>
; the dorsal surface of thorax and abdomen is densely covered by flat, round granules in both species. However, it is distinguished from the latter species by its larger size, the darker colouration of its body and its pale brown legs, and the shorter antennae (slightly less than 1.5 times as long as width of head). See also the Key to
<taxonomicName id="85A64D23FF9AA573FBCCF8D9A29E53A2" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[1089,1236,1715,1738]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="70D2EAB2FF9AA573FBCCF8D9A29E53A2" box="[1089,1236,1715,1738]" italics="true" pageId="1" pageNumber="434">Comorocoris</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="0ABC652BFF9AA576FF4AF8BCA2AD549D" lastPageId="4" lastPageNumber="437" pageId="1" pageNumber="434" type="description">
<paragraph id="421936A0FF9AA573FF4AF8BCA2D0527B" blockId="1.[151,1436,1174,2027]" pageId="1" pageNumber="434">
<emphasis id="70D2EAB2FF9AA573FF4AF8BCA7135387" bold="true" box="[199,345,1750,1775]" pageId="1" pageNumber="434">Description.</emphasis>
Apterous, dark brown, some specimens almost blackish, antennae and legs pale brown; dorsum of thorax and abdomen bulbously elevated, surface with numerous patches of flat tubercles.
</paragraph>
<paragraph id="421936A0FF9AA573FF4AF975A2195283" blockId="1.[151,1436,1174,2027]" pageId="1" pageNumber="434">
<emphasis id="70D2EAB2FF9AA573FF4AF975A723525E" box="[199,361,1823,1846]" italics="true" pageId="1" pageNumber="434">Measurements</emphasis>
(in mm). Male
<typeStatus id="9D1D8802FF9AA573FD97F974A436525F" box="[538,636,1822,1847]" pageId="1" pageNumber="434" type="holotype">holotype</typeStatus>
(one female
<typeStatus id="9D1D8802FF9AA573FC82F975A53A525F" box="[783,880,1823,1847]" pageId="1" pageNumber="434" type="paratype">paratype</typeStatus>
in brackets). Total body length: 2.96 (3.54); head length (including collar): 0.56 (0.55); head width across eyes: 0.64 (0.67); minimum interocular distance: 0.42 (0.44); length of antennal segments: I: 0.22 (0.24), II: 0.20 (0.22), III: 0.22 (0.26), IV: 0.22 (0.26); pronotum length: 0.47 (0.47), pronotum width: 1.76 (2.04); maximum width of abdomen: 2.00 (2.45), maximum height of abdomen in strictly lateral view, measured from outline of connexivum to dorsalmost point of dorsal outline and to ventralmost point of ventral outline: dorsal part: 1.11 (1.31), ventral part: 0.44 (0.46).
</paragraph>
<caption id="16D96628FF99A570FF1AF9CBA215529C" httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="2" pageNumber="435" targetBox="[168,1435,213,1915]" targetPageId="2">
<paragraph id="421936A0FF99A570FF1AF9CBA215529C" blockId="2.[151,1436,1953,2036]" pageId="2" pageNumber="435">
<emphasis id="70D2EAB2FF99A570FF1AF9CBA77052DF" bold="true" box="[151,314,1953,1976]" pageId="2" pageNumber="435">FIGURES 1–7.</emphasis>
<taxonomicName id="85A64D23FF99A570FECFF9C8A78D52DF" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[322,455,1954,1975]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="70D2EAB2FF99A570FECFF9C8A78D52DF" box="[322,455,1954,1975]" italics="true" pageId="2" pageNumber="435">Comorocoris</emphasis>
</taxonomicName>
species, dorsal habitus. 1–2,
<taxonomicName id="85A64D23FF99A570FD7EF9C8A5CC52DF" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[755,902,1954,1975]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF99A570FD7EF9C8A5CC52DF" box="[755,902,1954,1975]" italics="true" pageId="2" pageNumber="435">C. estherineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BE157C9FF99A570FC01F9C8A59452DF" box="[908,990,1954,1975]" pageId="2" pageNumber="435" rank="species">
<emphasis id="70D2EAB2FF99A570FC01F9C8A59452DF" bold="true" box="[908,990,1954,1975]" pageId="2" pageNumber="435">sp. nov.</emphasis>
</taxonomicNameLabel>
, 1, male holotype, 2, female paratype; 3, 6,
<taxonomicName id="85A64D23FF99A570FF1AF9ABA44752BE" authority="Heiss &amp; Banar, 2012" authorityName="Heiss &amp; Banar" authorityYear="2012" box="[151,525,1984,2006]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="70D2EAB2FF99A570FF1AF9ABA77952BE" box="[151,307,1985,2006]" italics="true" pageId="2" pageNumber="435">C. inexpectatus</emphasis>
Heiss &amp; Baňař, 2012
</taxonomicName>
, 3, male holotype, 6, female paratype; 4, 7,
<taxonomicName id="85A64D23FF99A570FC4BF9ABA30D52BE" authority="Heiss &amp; Banar, 2012" authorityName="Heiss &amp; Banar" authorityYear="2012" box="[966,1351,1984,2006]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="glabridorsum">
<emphasis id="70D2EAB2FF99A570FC4BF9ABA22752BD" box="[966,1133,1984,2006]" italics="true" pageId="2" pageNumber="435">C. glabridorsum</emphasis>
Heiss &amp; Baňař, 2012
</taxonomicName>
, 4, male holotype, 7, female paratype; 5,
<taxonomicName id="85A64D23FF99A570FE53F9B5A54A529C" authority="Heiss, 1985" authorityName="Heiss" authorityYear="1985" box="[478,768,2014,2036]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="testudiformis">
<emphasis id="70D2EAB2FF99A570FE53F9B5A4CA529C" box="[478,640,2015,2036]" italics="true" pageId="2" pageNumber="435">C. testudiformis</emphasis>
Heiss, 1985
</taxonomicName>
, female paratype. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="16D96628FF98A571FF1AFA87A4F05029" httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="3" pageNumber="436" targetBox="[155,1409,219,1203]" targetPageId="3">
<paragraph id="421936A0FF98A571FF1AFA87A4F05029" blockId="3.[151,1436,1261,1345]" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF1AFA87A70D506B" bold="true" box="[151,327,1261,1284]" pageId="3" pageNumber="436">FIGURES 8–12.</emphasis>
<taxonomicName id="85A64D23FF98A571FEC3FA85A799506C" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[334,467,1263,1284]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="70D2EAB2FF98A571FEC3FA85A799506C" box="[334,467,1263,1284]" italics="true" pageId="3" pageNumber="436">Comorocoris</emphasis>
</taxonomicName>
species, lateral habitus. 8–9,
<taxonomicName id="85A64D23FF98A571FC8DFA85A5D9506B" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[768,915,1262,1284]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF98A571FC8DFA85A5D9506B" box="[768,915,1262,1284]" italics="true" pageId="3" pageNumber="436">C. estherineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BE157C9FF98A571FC17FA85A5A1506C" box="[922,1003,1263,1284]" pageId="3" pageNumber="436" rank="species">
<emphasis id="70D2EAB2FF98A571FC17FA85A5A1506C" bold="true" box="[922,1003,1263,1284]" pageId="3" pageNumber="436">sp. nov.</emphasis>
</taxonomicNameLabel>
, 8, male holotype, 9, female paratype; 10,
<taxonomicName id="85A64D23FF98A571FF1AFB67A7F7504B" authority="Heiss, 1985" authorityName="Heiss" authorityYear="1985" box="[151,445,1293,1315]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="testudiformis">
<emphasis id="70D2EAB2FF98A571FF1AFB67A770504A" box="[151,314,1293,1314]" italics="true" pageId="3" pageNumber="436">C. testudiformis</emphasis>
Heiss, 1985
</taxonomicName>
, female paratype; 11,
<taxonomicName id="85A64D23FF98A571FD2CFB67A26B504B" authority="Heiss &amp; Banar, 2012" authorityName="Heiss &amp; Banar" authorityYear="2012" box="[673,1057,1293,1315]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="70D2EAB2FF98A571FD2CFB67A575504B" box="[673,831,1293,1315]" italics="true" pageId="3" pageNumber="436">C. inexpectatus</emphasis>
Heiss &amp; Baňař, 2012
</taxonomicName>
, male holotype, 12,
<taxonomicName id="85A64D23FF98A571FB7EFB67A7275029" authority="Heiss &amp; Banar, 2012" authorityName="Heiss &amp; Banar" authorityYear="2012" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="glabridorsum">
<emphasis id="70D2EAB2FF98A571FB7EFB67A3D6504A" box="[1267,1436,1293,1314]" italics="true" pageId="3" pageNumber="436">C. glabridorsum</emphasis>
Heiss &amp; Baňař, 2012
</taxonomicName>
, male holotype. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="421936A0FF98A571FF4AFB04A2F95337" blockId="3.[151,1437,1390,2027]" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF4AFB04A74150EE" box="[199,267,1390,1414]" italics="true" pageId="3" pageNumber="436">Head.</emphasis>
Wider than long, width: length ratio
<quantity id="855E9B45FF98A571FD48FB05A55F50EF" box="[709,789,1390,1415]" metricMagnitude="-2" metricUnit="m" metricValue="2.8956" pageId="3" pageNumber="436" unit="in" value="1.14">1.14 in</quantity>
male,
<quantity id="855E9B45FF98A571FCE8FB05A5FC50EF" box="[869,950,1390,1415]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="3" pageNumber="436" unit="in" value="1.21">1.21 in</quantity>
female; clypeus reaching about middle of antennal segment I, antenniferous lobes very short, wide and blunt; antennae short, 1.43 times as long as width of head in male, 1.46 times in female, segment I strongly bent at base (at level of clypeal apex), thickest, segment II thinner, shortest, tapering toward base, segment III approximately as long as segment I, subcylindrical, thinnest with conspicuous basal petiole, segment IV fusiform; eyes oval, slightly exceeding lateral outline of head; postocular lobes inconspicuous in male, slightly exceeding outer margin of eyes in female, ocular index
<quantity id="855E9B45FF98A571FAC1F849A3D65353" box="[1356,1436,1570,1595]" metricMagnitude="-2" metricUnit="m" metricValue="9.652" pageId="3" pageNumber="436" unit="in" value="3.8">3.80 in</quantity>
male,
<quantity id="855E9B45FF98A571FF56F82DA7635337" box="[219,297,1606,1631]" metricMagnitude="-2" metricUnit="m" metricValue="9.7028" pageId="3" pageNumber="436" unit="in" value="3.82">3.82 in</quantity>
female. Rostrum slightly longer than head, rostral groove wide open posteriorly.
</paragraph>
<paragraph id="421936A0FF98A571FF4AF801A5F153A3" blockId="3.[151,1437,1390,2027]" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF4AF801A77753EA" box="[199,317,1643,1666]" italics="true" pageId="3" pageNumber="436">Pronotum.</emphasis>
Strongly declivous anteriorly, short and wide, ratio length / width 4.33; lateral margins flattened, concave and converging anteriorly, anterior margin straight; disk with six oval callosities bordered by smaller flat round granulation; posterior margin concave medially, straight laterally.
</paragraph>
<paragraph id="421936A0FF98A571FF4AF8BDA77F525F" blockId="3.[151,1437,1390,2027]" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF4AF8BDA71B5386" box="[199,337,1751,1774]" italics="true" pageId="3" pageNumber="436">Mesonotum.</emphasis>
Five times as wide as long, roundly elevated at middle (
<figureCitation id="DA9D2A25FF98A571FB8EF8BCA23F5387" box="[1027,1141,1750,1775]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="3" pageNumber="436">Figs. 8–9</figureCitation>
), surface with flat round granulation of different size and elongate smooth callosities laterad of median elevation along anterior margin (
<figureCitation id="DA9D2A25FF98A571FF12F974A762525F" box="[159,296,1822,1847]" captionStart="FIGURES 13–14" captionStartId="4.[151,264,1507,1530]" captionTargetBox="[151,1436,807,1485]" captionTargetId="figure@4.[151,1436,807,1485]" captionTargetPageId="4" captionText="FIGURES 13–14. Comorocoris estherineae sp. nov., male paratype, scanning electron micrographs, pygophore removed. 13, dorsal view; 14, anterior view. Scale bars in mm." httpUri="https://zenodo.org/record/1159004/files/figure.png" pageId="3" pageNumber="436">Figs. 13–14</figureCitation>
).
</paragraph>
<paragraph id="421936A0FF98A571FF4AF929A7F452CA" blockId="3.[151,1437,1390,2027]" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF4AF929A7045232" box="[199,334,1859,1882]" italics="true" pageId="3" pageNumber="436">Metanotum.</emphasis>
Fused to mesonotum and mtg I+II with a round median elevation, the fusion line not clearly discernible; structure of surface as of mesonotum, the two anterolateral curved callosities are larger (
<figureCitation id="DA9D2A25FF98A571FA81F90CA3DE5217" box="[1292,1428,1894,1919]" captionStart="FIGURES 13–14" captionStartId="4.[151,264,1507,1530]" captionTargetBox="[151,1436,807,1485]" captionTargetId="figure@4.[151,1436,807,1485]" captionTargetPageId="4" captionText="FIGURES 13–14. Comorocoris estherineae sp. nov., male paratype, scanning electron micrographs, pygophore removed. 13, dorsal view; 14, anterior view. Scale bars in mm." httpUri="https://zenodo.org/record/1159004/files/figure.png" pageId="3" pageNumber="436">Figs. 13–14</figureCitation>
) than those on mesonotum.
</paragraph>
<paragraph id="421936A0FF98A576FF4AF9C4A2EC5475" blockId="3.[151,1437,1390,2027]" lastBlockId="4.[151,1436,151,752]" lastPageId="4" lastPageNumber="437" pageId="3" pageNumber="436">
<emphasis id="70D2EAB2FF98A571FF4AF9C4A77252AE" box="[199,312,1966,1990]" italics="true" pageId="3" pageNumber="436">Abdomen.</emphasis>
Mtg I+II fused to each other, medially roundly elevated; surface with flat granulation and a larger oval callosity laterally; posteriorly strongly sloping and fused to mtg III–VII, the bisinuate posterior margin delimiting the smaller flat granulation of mtg I+II surface and the larger one of mtg III–VII (
<figureCitation id="DA9D2A25FF9FA576FB3FFEFDA2B555D8" box="[1202,1279,151,176]" captionStart="FIGURES 13–14" captionStartId="4.[151,264,1507,1530]" captionTargetBox="[151,1436,807,1485]" captionTargetId="figure@4.[151,1436,807,1485]" captionTargetPageId="4" captionText="FIGURES 13–14. Comorocoris estherineae sp. nov., male paratype, scanning electron micrographs, pygophore removed. 13, dorsal view; 14, anterior view. Scale bars in mm." httpUri="https://zenodo.org/record/1159004/files/figure.png" pageId="4" pageNumber="437">Fig.13</figureCitation>
); deltg II–VII flat, laterally expanded, lateral margins slightly sinuate; spiracles II–VII sublateral to lateral placed on rim of laterally reflexed vltg II–VII, visible from above, VIII terminal on ptg VIII. Abdominal scent gland scars present on posterior margin of mtg III (conspicuous), IV (reduced) and on mtg V (strongly reduced).
</paragraph>
<paragraph id="421936A0FF9FA576FF4AFF4DA54A5428" blockId="4.[151,1436,151,752]" box="[199,768,295,320]" pageId="4" pageNumber="437">
Ventral structures as in other
<taxonomicName id="85A64D23FF9FA576FD83FF42A4EB5457" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[526,673,296,319]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="70D2EAB2FF9FA576FD83FF42A4EB5457" box="[526,673,296,319]" italics="true" pageId="4" pageNumber="437">Comorocoris</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="421936A0FF9FA576FF4AFF27A70054E0" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF4AFF27A749540C" box="[199,259,333,356]" italics="true" pageId="4" pageNumber="437">Legs.</emphasis>
Pale brown, unarmed, slender, sparsely covered with short, semi-erect setae. Tibiae straight with preapical comb.
</paragraph>
<paragraph id="421936A0FF9FA576FF4AFFFEA4B354A7" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF4AFFFEA73854C4" box="[199,370,404,428]" italics="true" pageId="4" pageNumber="437">Male genitalia.</emphasis>
Exposed posterior part of pygophore oval, strongly declivous, surface transversely rugose. The
<typeStatus id="9D1D8802FF9FA576FF1AFFDDA6B354B8" box="[151,249,439,464]" pageId="4" pageNumber="437" type="holotype">holotype</typeStatus>
was not dissected for the study of parameres.
</paragraph>
<paragraph id="421936A0FF9FA576FF1AFFB6A2AD549D" blockId="4.[151,1436,151,752]" box="[151,1255,476,501]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF1AFFB6A73C549C" box="[151,374,476,500]" italics="true" pageId="4" pageNumber="437">Sexual dimorphism.</emphasis>
General structures similar in both sexes, female of larger body size and width.
</paragraph>
</subSubSection>
<subSubSection id="0ABC652BFF9FA576FF4AFF95A29257A5" pageId="4" pageNumber="437" type="etymology">
<paragraph id="421936A0FF9FA576FF4AFF95A76E5708" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF4AFF95A7045770" bold="true" box="[199,334,511,536]" pageId="4" pageNumber="437">Etymology.</emphasis>
Dedicated to the first author’s good friend and colleague, Marie Estherine Rabotoson (
<collectingRegion id="8062F842FF9FA576FF13FC4EA7785755" box="[158,306,548,573]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="437">Fianarantsoa</collectingRegion>
,
<collectingCountry id="3AB17630FF9FA576FEC8FC4EA7845755" box="[325,462,548,573]" name="Madagascar" pageId="4" pageNumber="437">Madagascar</collectingCountry>
) who collected part of the
<typeStatus id="9D1D8802FF9FA576FCA4FC4FA5135755" box="[809,857,549,573]" pageId="4" pageNumber="437">type</typeStatus>
series and participated in several field trips in
<collectingCountry id="3AB17630FF9FA576FF1AFC2DA76B5708" box="[151,289,583,608]" name="Madagascar" pageId="4" pageNumber="437">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="421936A0FF9FA576FF4AFC06A29257A5" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF4AFC06A7B357ED" bold="true" box="[199,505,620,645]" pageId="4" pageNumber="437">Collecting circumstances.</emphasis>
All known specimens were collected by sifting of mountain evergreen rain forest leaf litter in Montagne d’Ambre National Park in Northern
<collectingCountry id="3AB17630FF9FA576FCFCFCE5A5B157C0" box="[881,1019,655,680]" name="Madagascar" pageId="4" pageNumber="437">Madagascar</collectingCountry>
. Sifted samples were subsequently extracted in Winkler apparatus during two or three days (
<figureCitation id="DA9D2A25FF9FA576FC9BFCDEA52357A5" box="[790,873,692,717]" captionStart="FIGURES 15–16. 15" captionStartId="5.[151,264,1926,1949]" captionTargetBox="[151,1436,303,1905]" captionTargetId="figure@5.[151,1436,303,1905]" captionTargetPageId="5" captionText="FIGURES 15–16. 15, Microhabitat of part of type series of Comorocoris estherineae sp. nov. in Montagne d’Ambre National Park; 16, extraction of sifted material using Winkler apparatus in Montagne d’Ambre National Park." httpUri="https://zenodo.org/record/1159006/files/figure.png" pageId="4" pageNumber="437">Fig. 16</figureCitation>
), mixed several times every day.
</paragraph>
</subSubSection>
<subSubSection id="0ABC652BFF9FA576FF4AFCBDA2995798" box="[199,1235,727,752]" pageId="4" pageNumber="437" type="distribution">
<paragraph id="421936A0FF9FA576FF4AFCBDA2995798" blockId="4.[151,1436,151,752]" box="[199,1235,727,752]" pageId="4" pageNumber="437">
<emphasis id="70D2EAB2FF9FA576FF4AFCBDA72A5798" bold="true" box="[199,352,727,752]" pageId="4" pageNumber="437">Distribution.</emphasis>
Known only from Montagne d’Ambre National Park in Northern
<collectingCountry id="3AB17630FF9FA576FBCBFCBDA29A5798" box="[1094,1232,727,752]" name="Madagascar" pageId="4" pageNumber="437">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="293034CE0D1D89D731D2D4B24C63657E">A new species of Comorocoris from Northern Madagascar (Hemiptera: Heteroptera: Aradidae)</mods:title>
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Key to
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<emphasis id="70D2EAB2FF9DA574FF7DF938A7C65204" bold="true" box="[240,396,1874,1900]" italics="true" pageId="6" pageNumber="439">Comorocoris</emphasis>
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species
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1 Body colour brown to dark brown, whole surface of thorax and abdomen reticulate and strongly granulose, leaving only few larger smooth callosities on lateral sclerites of thoracic segments and mtg III–IV; abdomen very high (strongly convex) in lateral view (
<figureCitation id="DA9D2A25FF9DA574FECBF9B9A7E75281" box="[326,429,2003,2025]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="6" pageNumber="439">Figs. 8–10</figureCitation>
)................................................................................. 2
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- Body colour black, only pro- and mesonotum with more extended flat round granulation, surface of metanotum and abdomen smooth with isolated patches of granules; abdomen lower from lateral view (
<figureCitation id="DA9D2A25FF9CA575FC4DFEDEA27855A2" box="[960,1074,180,202]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="7" pageNumber="440">Figs. 11–12</figureCitation>
)........................... 3
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2 Smaller species, body length
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in male,
<quantity id="855E9B45FF9CA575FD3CFEBBA54D558F" box="[689,775,209,231]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="440" unit="mm" value="3.1">3.10 mm</quantity>
in female, maximum abdomen width
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in male,
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females, antennae about twice as long as width of head, colouration reddish-brown (
<figureCitation id="DA9D2A25FF9CA575FBBDFE85A23C546D" box="[1072,1142,239,261]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[168,1435,213,1915]" captionTargetId="figure@2.[151,1436,193,1932]" captionTargetPageId="2" captionText="FIGURES 1–7. Comorocoris species, dorsal habitus. 1–2, C. estherineae sp. nov., 1, male holotype, 2, female paratype; 3, 6, C.inexpectatusHeiss &amp; Baňař,2012, 3, male holotype, 6, female paratype; 4, 7, C.glabridorsumHeiss &amp; Baňař,2012, 4, male holotype, 7, female paratype; 5, C. testudiformis Heiss, 1985, female paratype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="7" pageNumber="440">Figs. 5</figureCitation>
,
<figureCitation id="DA9D2A25FF9CA575FB09FE85A2D5546D" box="[1156,1183,239,261]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="7" pageNumber="440">10</figureCitation>
); Moheli Island, Comoro Archipelago...................................................................
<taxonomicName id="85A64D23FF9CA575FB06FF67A3D6544A" authority="Heiss, 1985" authorityName="Heiss" authorityYear="1985" box="[1163,1436,268,290]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="testudiformis">
<emphasis id="70D2EAB2FF9CA575FB06FF67A36E544A" box="[1163,1316,269,290]" italics="true" pageId="7" pageNumber="440">C. testudiformis</emphasis>
<bibRefCitation id="26374B51FF9CA575FAA6FF66A3D6544A" author="Heiss" box="[1323,1436,268,290]" pageId="7" pageNumber="440" refString="Heiss, E. (1985) Eine neue aptere Aradidengattung aus Afrika (Heteroptera, Aradidae, Carventinae). Revue Zoologique Africaine, 99, 147 - 152." type="journal article" year="1985">Heiss, 1985</bibRefCitation>
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- Larger species, body length
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in male,
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in female, maximum abdomen width 2.00 mm in male,
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females, antennae short, slightly less than 1.5 times long as width of head, colouration dark brown to blackish (
<figureCitation id="DA9D2A25FF9CA575FB31FF2DA3525435" box="[1212,1304,327,349]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[168,1435,213,1915]" captionTargetId="figure@2.[151,1436,193,1932]" captionTargetPageId="2" captionText="FIGURES 1–7. Comorocoris species, dorsal habitus. 1–2, C. estherineae sp. nov., 1, male holotype, 2, female paratype; 3, 6, C.inexpectatusHeiss &amp; Baňař,2012, 3, male holotype, 6, female paratype; 4, 7, C.glabridorsumHeiss &amp; Baňař,2012, 4, male holotype, 7, female paratype; 5, C. testudiformis Heiss, 1985, female paratype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="7" pageNumber="440">Figs. 1–2</figureCitation>
,
<figureCitation id="DA9D2A25FF9CA575FAAFFF2DA3005435" box="[1314,1354,327,349]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="7" pageNumber="440">8–9</figureCitation>
); Northern Madagascar....................................................................
<taxonomicName id="85A64D23FF9CA575FB33FF0FA3035412" authority="Baňař &amp; Heiss, 2018" authorityName="Baňař &amp; Heiss" authorityYear="2018" box="[1214,1353,357,378]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="estherineae" status="sp. nov.">
<emphasis id="70D2EAB2FF9CA575FB33FF0FA3035412" box="[1214,1353,357,378]" italics="true" pageId="7" pageNumber="440">C. estherineae</emphasis>
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<emphasis id="70D2EAB2FF9CA575FAC2FF0EA3D65412" bold="true" box="[1359,1436,356,378]" pageId="7" pageNumber="440">sp. nov.</emphasis>
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3 Smaller species, body length 3.00–
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, surface submat, sutures separating meso-, metanotum, mtgI and mtg II respectively not interrupted and obliterating on median smooth plate (
<figureCitation id="DA9D2A25FF9CA575FCCCFFF5A5D754DD" box="[833,925,415,437]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[168,1435,213,1915]" captionTargetId="figure@2.[151,1436,193,1932]" captionTargetPageId="2" captionText="FIGURES 1–7. Comorocoris species, dorsal habitus. 1–2, C. estherineae sp. nov., 1, male holotype, 2, female paratype; 3, 6, C.inexpectatusHeiss &amp; Baňař,2012, 3, male holotype, 6, female paratype; 4, 7, C.glabridorsumHeiss &amp; Baňař,2012, 4, male holotype, 7, female paratype; 5, C. testudiformis Heiss, 1985, female paratype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="7" pageNumber="440">Figs. 3, 6</figureCitation>
,
<figureCitation id="DA9D2A25FF9CA575FC2AFFF5A58A54DD" box="[935,960,415,437]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1261,1284]" captionTargetBox="[155,1409,219,1203]" captionTargetId="figure@3.[151,1436,193,1241]" captionTargetPageId="3" captionText="FIGURES 8–12. Comorocoris species, lateral habitus. 8–9, C. estherineae sp. nov., 8, male holotype, 9, female paratype; 10, C. testudiformis Heiss, 1985, female paratype; 11, C. inexpectatus Heiss &amp; Baňař, 2012, male holotype, 12, C. glabridorsum Heiss &amp; Baňař, 2012, male holotype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159002/files/figure.png" pageId="7" pageNumber="440">11</figureCitation>
); Central Madagascar............................................................................................
<taxonomicName id="85A64D23FF9CA575FBB4FFD7A3D654BA" authority="Heiss &amp; Banar, 2012" authorityName="Heiss &amp; Banar" authorityYear="2012" box="[1081,1436,444,466]" class="Insecta" family="Aradidae" genus="Comorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="70D2EAB2FF9CA575FBB4FFD7A28654BA" box="[1081,1228,445,466]" italics="true" pageId="7" pageNumber="440">C. inexpectatus</emphasis>
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- Larger species, body length
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, surface glabrous, sutures separating meso-, metanotum, mtgI and mtg II respectively interrupted and obliterated on median smooth plate (
<figureCitation id="DA9D2A25FF9CA575FC99FF9DA53A5764" box="[788,880,503,525]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[168,1435,213,1915]" captionTargetId="figure@2.[151,1436,193,1932]" captionTargetPageId="2" captionText="FIGURES 1–7. Comorocoris species, dorsal habitus. 1–2, C. estherineae sp. nov., 1, male holotype, 2, female paratype; 3, 6, C.inexpectatusHeiss &amp; Baňař,2012, 3, male holotype, 6, female paratype; 4, 7, C.glabridorsumHeiss &amp; Baňař,2012, 4, male holotype, 7, female paratype; 5, C. testudiformis Heiss, 1985, female paratype.Scale bar 1 mm." httpUri="https://zenodo.org/record/1159000/files/figure.png" pageId="7" pageNumber="440">Figs. 4, 7</figureCitation>
,
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); Central Madagascar...............................................................................................
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Alive, Fig.
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.
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