Add updates up until 2024-12-04 16:34:49

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@ -498,7 +503,7 @@ leg.; station PM12;
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<materialsCitation id="3B676903FFD38B1DDBF4FE16FB76D246" ID-GBIF-Occurrence="3051754374" collectedFrom="sandy beach and intertidal rocks" collectingDate="27 Nov. 2012, 9 Dec. 2012" collectionCode="MNHN" collectorName="Papua Niugini" country="Papua New Guinea" latitude="-5.135" location="Wonad Island" longLatPrecision="130" longitude="145.82167" pageId="79" pageNumber="80" specimenCode="MNHN-IM-2013-15867" specimenCount="1" stateProvince="Madang">
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(13/11 [5457] mm);
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@ -507,11 +512,9 @@ leg.; station PM12;
,
<geoCoordinate id="EE3B0599FFD38B1DDDBAFE74FCCED1AB" box="[642,781,475,503]" degrees="145" direction="east" minutes="49.3" orientation="longitude" pageId="79" pageNumber="80" precision="92" value="145.82167">145°49.3 E</geoCoordinate>
;
<collectingDate id="EFF5BC76FFD38B1DDC2EFE73FB84D1AA" box="[790,1095,476,503]" isEnumeration="true" pageId="79" pageNumber="80">
<date id="FFB1459EFFD38B1DDC2EFE73FC71D1AA" box="[790,946,476,503]" pageId="79" pageNumber="80" value="2012-11-27">27 Nov. 2012</date>
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<date id="FFB1459EFFD38B1DDC83FE73FB84D1AA" box="[955,1095,476,503]" pageId="79" pageNumber="80" value="2012-12-09">9 Dec. 2012</date>
</collectingDate>
; MNHN expedition Papua Niugini leg.; station PM41;
<collectedFrom id="381320A3FFD38B1DDD31FE50FC4AD245" box="[521,905,511,537]" pageId="79" pageNumber="80">sandy beach and intertidal rocks</collectedFrom>
;

View file

@ -1,108 +1,113 @@
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@ -123,13 +123,13 @@ leg.;
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</paragraph>
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<materialsCitation id="3B68F9693A33FFECFF07FA95FB6B4E17" ID-GBIF-Occurrence="3434743302" collectedFrom="high littoral, fine sand sediments" collectingDate="2006-04-01" collectionCode="CNP-NEM" collectorName="C. Pastor" country="Argentina" latitude="-45.918613" location="ARGENTINA" municipality="San Jorge Gulf" pageId="7" pageNumber="24" specimenCode="CNP-NEM 27259, CNP-NEM 27443" specimenCount="2" specimenCount-male="2" stateProvince="Chubut" typeStatus="paratype">
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</emphasis>
<collectingCountry id="F317B3A43A33FFECFF50FAA8FEEF4E17" box="[189,358,1330,1356]" name="Argentina" pageId="7" pageNumber="24">ARGENTINA</collectingCountry>
<specimenCount id="9D0638BD3A33FFECFE92FAA8FE4D4E17" box="[383,452,1330,1356]" pageId="7" pageNumber="24" type="male">2 ♂♂</specimenCount>
<specimenCount id="9D0638BD3A33FFECFE92FAA8FE4D4E17" box="[383,452,1330,1356]" count="2" pageId="7" pageNumber="24" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype;
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and

View file

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Key to species of
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with simple spicules shorter than 2 abd and subventral precloacal setae present

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<emphasis id="B5D8EABFFFC19E0245F1FA37EE89FA27" bold="true" box="[151,474,1424,1451]" italics="true" pageId="45" pageNumber="46">Pseudoanthidium scapulare</emphasis>
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10 Clypeus flat with a clock-like shape, apical margin hidden by dense fringe of appressed moustache-like hairs..................................................................................................
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group A comprehensive key to the
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group has been presented by
<bibRefCitation id="E33D4B5CFFC19E0246EAF9B7EB01F9A9" author="Litman, J. R. &amp; Fateryga, A. V. &amp; Griswold, T. L. &amp; Aubert, M. &amp; Proshchalykin, M. Yu. &amp; Le Divelec, R. &amp; Burrows, S. &amp; Praz, Ch. J." box="[908,1106,1552,1574]" pageId="45" pageNumber="46" pagination="1287 - 1337" refId="ref30469" refString="Litman, J. R., Fateryga, A. V., Griswold, T. L., Aubert, M., Proshchalykin, M. Yu., Le Divelec, R., Burrows, S. &amp; Praz, Ch. J. (2021) Paraphyly and low levels of genetic divergence in morphologically distinct taxa: revision of the Pseudoanthidium scapulare complex of carder bees (Apoidea: Megachilidae: Anthidiini). Zoological Journal of the Linnean Society, 195, 1287 - 1337. [2022] https: // doi. org / 10.1093 / zoolinnean / zlab 062" type="journal article" year="2021">
Litman
<emphasis id="B5D8EABFFFC19E0246BEF9B6EB59F9A9" box="[984,1034,1552,1574]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
(2021)
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. See also the illustrated review by
<bibRefCitation id="E33D4B5CFFC19E024584F98AEED0F9CF" author="Kasparek, M." box="[226,387,1581,1603]" pageId="45" pageNumber="46" refId="ref29929" refString="Kasparek, M. (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe and Western Turkey. Identification. Distribution. Biology. Chimaira, Frankfurt, 292 pp. [unchanged reprint 2023]" type="book" year="2022">Kasparek (2022)</bibRefCitation>
and the supplement (addition of a new species) by Veerecken (2023).
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<keyLead id="3C5D9598FFC19E0245F1F9ECEACFF9EC" box="[151,1436,1611,1633]" pageId="45" pageNumber="46"> Clypeus not flat, apical margin exposed or only slightly covered with hairs...................................... 11</keyLead>
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6 Depressions of T1T4 with indistinct, diffuse red-brown band; T3T4 with orange maculations.............
<taxonomicName id="40AC4D2EFFC19E02407DFC39EACFFC3E" ID-CoL="8SNNT" authorityName="Pasteels" authorityYear="1969" box="[1307,1436,925,947]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rubellulum">
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<keyLead id="3C5D9598FFC19E0245F1FC1CEACFFC5C" box="[151,1436,955,977]" pageId="45" pageNumber="46"> Depressions of T1T4 black or dark brown; T3T4 with yellow maculations...................................... 7</keyLead>
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7 T1 with yellow lateral spot; scutum and scutellum polished, with shallow, mostly irregularly scattered punctation (interstices up to a few puncture diameters); tegula light brown.............................................
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T1 black; scutum and scutellum with deep, regular punctation (interstices always less than a puncture diameter); tegula dark brown....................................................................................
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<emphasis id="B5D8EABFFFC19E0245F1FE0DEE4BFE48" bold="true" box="[151,280,426,452]" italics="true" pageId="45" pageNumber="46">P. eximium</emphasis>
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2 Apical margin of T5 with small median protrusion; apical margin of scutellum and axillae widely rounded in dorsal view..........................................................................................
<taxonomicName id="40AC4D2EFFC19E02419DFDAAEACFFDAD" ID-CoL="4NP56" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1953" box="[1275,1436,524,546]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="wahrmanicum">
<emphasis id="B5D8EABFFFC19E02419DFDAAEACFFDAD" box="[1275,1436,524,546]" italics="true" pageId="45" pageNumber="46">P. wahrmanicum</emphasis>
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<keyLead id="3C5D9598FFC19E0245F1FD8DEACFFDB3" box="[151,1436,553,575]" pageId="45" pageNumber="46"> Apical margin of T5 straight without median protrusion; scutellum angular posterolaterally in dorsal view.............. 3</keyLead>
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<keyLead id="3C5D9598FFC19E0245F1FDE0EACFFDF5" pageId="45" pageNumber="46">3 Apical margin of T6 with small median protrusion, deeply V-shaped incised; posterior margin of axilla and posterolateral margin of scutellum forming straight line; posterior margin of scutellum rounded in dorsal view...................... 4</keyLead>
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Apical margin of T6 with small ventrad protrusion; posterior margin of axilla and posterolateral margin of scutellum not forming straight line; posterior margin of scutellum straight in dorsal view...............................
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<emphasis id="B5D8EABFFFC19E024055FD38EACFFD38" box="[1331,1436,671,692]" italics="true" pageId="45" pageNumber="46">P. eximium</emphasis>
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4 Scutum black, terga black with pale yellow bands....................................................
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<emphasis id="B5D8EABFFFC19E024023FD1AEACFFD5D" box="[1349,1436,700,722]" italics="true" pageId="45" pageNumber="46">P. enslini</emphasis>
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Scutum yellow and reddish-brown with some black remnants; terga yellow..............................
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5 Omaulus lamellate almost to lower end [
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group]............................................... 6
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<keyLead id="3C5D9598FFC19E0245F1FCB3EACFFCA5" box="[151,1436,788,810]" pageId="45" pageNumber="46"> Omaulus angulate or rounded........................................................................... 8</keyLead>
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The females of
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(both belonging to species group/subgenus
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1 T6 excavated, strongly concave [
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group / Subgen.
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<emphasis id="B5D8EABFFFC19E02464BFEEFECFBFED1" box="[813,936,328,349]" italics="true" pageId="45" pageNumber="46">Exanthidium</emphasis>
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]....................................... 2
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<keyLead id="3C5D9598FFC19E0245F1FEC1EACFFEF7" box="[151,1436,357,379]" pageId="45" pageNumber="46"> T6 not excavated, at most slightly depressed............................................................... 5</keyLead>
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<emphasis id="B5D8EABFFFC29E0145F1FA8CEE4BFAC8" bold="true" box="[151,280,1322,1349]" italics="true" pageId="46" pageNumber="47">P. eximium</emphasis>
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<keyLead id="3C5D9598FFC29E0145F1FAD7EACFFA09" box="[151,1436,1391,1413]" pageId="46" pageNumber="47">4 T6 with hooked or spiniform lateral protrusion............................................................. 5</keyLead>
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<keyLead id="3C5D9598FFC29E0145F1FA2AEACFFA2E" box="[151,1436,1421,1443]" pageId="46" pageNumber="47"> T6 laterally bulging, hooked or spiniform protrusion absent................................................... 7</keyLead>
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5 Base of emargination of T7 pointed (emargination V-shaped); posterior margin of scutellum in dorsal view straight...................................................................................................
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<keyLead id="3C5D9598FFC29E0145F1FA42EACFFA77" box="[151,1436,1509,1531]" pageId="46" pageNumber="47"> Base of emargination straight (emargination U-shaped); posterior margin of scutellum rounded in dorsal view........... 6</keyLead>
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6 Scutum black; yellow band on T1 interrupted in the middle, transition from disc to depression on anterior terga smooth.................................................................................................
<taxonomicName id="40AC4D2EFFC29E014023F987EACFF9B8" ID-CoL="4NP4B" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[1349,1436,1567,1589]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="B5D8EABFFFC29E014023F987EACFF9B8" box="[1349,1436,1567,1589]" italics="true" pageId="46" pageNumber="47">P. enslini</emphasis>
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Scutum with yellow or reddish-brown anterolateral maculation; yellow band on T1 not interrupted; transition from disc to depression step-like..........................................................................
<taxonomicName id="40AC4D2EFFC29E01404EF9FCEACFF9E3" ID-CoL="4NP4D" baseAuthorityName="Pasteels" baseAuthorityYear="1980" box="[1320,1436,1626,1648]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="B5D8EABFFFC29E01404EF9FCEACFF9E3" box="[1320,1436,1626,1648]" italics="true" pageId="46" pageNumber="47">P. guichardi</emphasis>
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7 Emargination of T7 little widening in apical third; T7 without transparent margin; outer faces of femora and tibiae black...............................................................................................
<taxonomicName id="40AC4D2EFFC29E01405CF932EACEF926" ID-CoL="4NP4S" baseAuthorityName="Morawitz" baseAuthorityYear="1894" box="[1338,1437,1685,1706]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pictipes">
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<keyLead id="3C5D9598FFC29E0145F1F914EACFF94B" box="[151,1436,1714,1736]" pageId="46" pageNumber="47"> Emargination of T7 strongly widening in apical third; T7 with transparent margin; outer faces of femora and tibiae yellow... 8</keyLead>
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8 Yellow preoccipital band reaching mandibular base; scutum with yellow anterolateral boomerang-shaped band; apical margin of S2 with median protrusion...............................................................
<taxonomicName id="40AC4D2EFFC29E01406FF94AEACFF88E" authorityName="Kasparek" authorityYear="2021" box="[1289,1436,1773,1794]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="deserticolum">
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Yellow stripe on upper gena not reaching mandibular base and not extending onto vertex; anterolateral marking on scutum absent; apical margin of S2 truncated.........................................................
<taxonomicName id="40AC4D2EFFC29E01419DF88FEACFF8B0" ID-CoL="4NP56" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1953" box="[1275,1436,1831,1853]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="wahrmanicum">
<emphasis id="B5D8EABFFFC29E01419DF88FEACFF8B0" box="[1275,1436,1831,1853]" italics="true" pageId="46" pageNumber="47">P. wahrmanicum</emphasis>
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<keyLead id="3C5D9598FFC29E0145F1F8C4EACFF8FB" box="[151,1436,1890,1912]" pageId="46" pageNumber="47"> Omaulus not lamellate................................................................................ 12</keyLead>
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<paragraph id="871336ADFFC39E0045F1FF30EEFAFF3D" blockId="47.[151,425,150,177]" box="[151,425,150,177]" pageId="47" pageNumber="48">
<heading id="DC5B81C1FFC39E0045F1FF30EEFAFF3D" bold="true" box="[151,425,150,177]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="B5D8EABFFFC39E0045F1FF30EEFAFF3D" bold="true" box="[151,425,150,177]" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="40AC4D2EFFC39E0045F1FF30EE0DFF3D" ID-CoL="4NP4P" authority="(Alfken, 1932)" baseAuthorityName="Alfken" baseAuthorityYear="1932" box="[151,350,150,177]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ochrognathum">P. ochrognathum</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFC39E0045F1FF7CEACFFE0F" pageId="47" pageNumber="48" type="key">
<key id="F33D619CFFC39E0045F1FF7CEACFFE0F" pageId="47" pageNumber="48">
<keyStep id="3C582E08FFC39E0045F1FF7CEACFFEA7" pageId="47" pageNumber="48">
<paragraph id="871336ADFFC39E0045F1FF7CEACFFF7C" blockId="47.[151,1436,219,387]" box="[151,1436,219,241]" pageId="47" pageNumber="48">
<keyLead id="3C5D9598FFC39E0045F1FF7CEACFFF7C" box="[151,1436,219,241]" pageId="47" pageNumber="48">10 T1 with lateral yellow spot; scutum and scutellum with shallow, often scattered punctation......................... 11</keyLead>
</paragraph>
<paragraph id="871336ADFFC39E0045F1FF5EEACFFEA7" blockId="47.[151,1436,219,387]" pageId="47" pageNumber="48">
<keyLead id="3C5D9598FFC39E0045F1FF5EEACFFEA7" pageId="47" pageNumber="48">
T1 black; scutum and scutellum with dense, regular punctation; combs at the base of S5 separated by less than one comb width....................................................................................
<taxonomicName id="40AC4D2EFFC39E004079FEB1EACFFEA7" ID-CoL="4NP42" baseAuthorityName="Benoist" baseAuthorityYear="1950" box="[1311,1436,278,299]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="B5D8EABFFFC39E004079FEB1EACFFEA7" box="[1311,1436,278,299]" italics="true" pageId="47" pageNumber="48">P. beaumonti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3C582E08FFC39E0045F1FE94EACFFE0F" pageId="47" pageNumber="48">
<paragraph id="871336ADFFC39E0045F1FE94EACFFEC4" blockId="47.[151,1436,219,387]" box="[151,1436,307,329]" pageId="47" pageNumber="48">
<keyLead id="3C5D9598FFC39E0045F1FE94EACFFEC4" box="[151,1436,307,329]" pageId="47" pageNumber="48">
11 Ground colour of terga black; median combs at the base of S5 separated by 34 comb widths............
<taxonomicName id="40AC4D2EFFC39E00419CFE93EACFFEC4" ID-CoL="4NP4P" baseAuthorityName="Alfken" baseAuthorityYear="1932" box="[1274,1436,307,329]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFC39E00419CFE93EACFFEC4" box="[1274,1436,307,329]" italics="true" pageId="47" pageNumber="48">P. ochrognathum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="871336ADFFC39E0045F1FEF6EACFFE0F" blockId="47.[151,1436,219,387]" pageId="47" pageNumber="48">
<keyLead id="3C5D9598FFC39E0045F1FEF6EACFFE0F" pageId="47" pageNumber="48">
Tergal discs black, depressions bronze-coloured; median combs at the base of S5 separated by 68 comb widths...................................................................................................
<taxonomicName id="40AC4D2EFFC39E00407DFEC9EACFFE0F" ID-CoL="8SNNT" authorityName="Pasteels" authorityYear="1969" box="[1307,1436,366,387]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rubellulum">
<emphasis id="B5D8EABFFFC39E00407DFEC9EACFFE0F" box="[1307,1436,366,387]" italics="true" pageId="47" pageNumber="48">P. rubellulum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,454 @@
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<mods:title id="DE7319DDF2861B02C5E2A520FD0429C2">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<subSubSection id="CFB66526FFE19E2245F1FF30ECB1FF3D" box="[151,994,151,178]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="871336ADFFE19E2245F1FF30ECB1FF3D" blockId="13.[151,994,151,214]" box="[151,994,151,178]" pageId="13" pageNumber="14">
<heading id="DC5B81C1FFE19E2245F1FF30ECB1FF3D" bold="true" box="[151,994,151,178]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B5D8EABFFFE19E2245F1FF30ECB1FF3D" bold="true" box="[151,994,151,178]" pageId="13" pageNumber="14">
<taxonomicName id="40AC4D2EFFE19E2245F1FF30ED2DFF3D" ID-CoL="8SQNJ" authority="(Friese, 1911)" baseAuthorityName="Friese" baseAuthorityYear="1911" box="[151,638,151,178]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2245F1FF30EE9CFF3D" bold="true" box="[151,463,151,178]" italics="true" pageId="13" pageNumber="14">Pseudoanthidium moricei</emphasis>
(Friese, 1911)
</taxonomicName>
stat. resurrect &amp; comb. nov.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CFB66526FFE19E2245F1FF1CEEB8FF5A" box="[151,491,187,214]" pageId="13" pageNumber="14" type="description">
<paragraph id="871336ADFFE19E2245F1FF1CEEB8FF5A" blockId="13.[151,994,151,214]" box="[151,491,187,214]" pageId="13" pageNumber="14">
(
<figureCitation id="1F972A28FFE19E2245F9FF1CEFBBFF5A" box="[159,232,187,214]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="13" pageNumber="14">Figs 7</figureCitation>
,
<figureCitation id="1F972A28FFE19E224592FF1CEE40FF59" box="[244,275,187,213]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="13" pageNumber="14">20</figureCitation>
,
<figureCitation id="1F972A28FFE19E224478FF1CEE6FFF59" box="[286,316,187,213]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="13" pageNumber="14">21</figureCitation>
,
<figureCitation id="1F972A28FFE19E22442EFF1CEE34FF59" box="[328,359,187,213]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="13" pageNumber="14">22</figureCitation>
,
<figureCitation id="1F972A28FFE19E224414FF1CEEC3FF59" box="[370,400,187,213]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="13" pageNumber="14">23</figureCitation>
,
<figureCitation id="1F972A28FFE19E2244FAFF1CEEE9FF59" box="[412,442,187,213]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="13" pageNumber="14">24</figureCitation>
, 25)
</paragraph>
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<subSubSection id="CFB66526FFE19E2245F1FEA4EC9DFE05" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="871336ADFFE19E2245F1FEA4EDBAFE91" blockId="13.[151,1436,259,394]" box="[151,745,259,285]" pageId="13" pageNumber="14">
<taxonomicName id="40AC4D2EFFE19E2245F1FEA4EEABFE91" authority="Friese, 1911" authorityName="Friese" authorityYear="1911" box="[151,504,259,285]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2245F1FEA4EE3AFE91" box="[151,361,259,285]" italics="true" pageId="13" pageNumber="14">Anthidium moricei</emphasis>
Friese, 1911
</taxonomicName>
.—Female,
<collectingCountry id="FFBB763DFFE19E22471AFEA4EDB6FE91" box="[636,741,259,285]" name="Palestine" pageId="13" pageNumber="14">Palestine</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFE19E2245F1FE80EC59FECD" blockId="13.[151,1436,259,394]" box="[151,778,295,321]" pageId="13" pageNumber="14">
<taxonomicName id="40AC4D2EFFE19E2245F1FE80ED62FECD" authority="(Friese, 1911)" baseAuthorityName="Friese" baseAuthorityYear="1911" box="[151,561,295,321]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2245F1FE80EEC2FECD" box="[151,401,295,321]" italics="true" pageId="13" pageNumber="14">Royanthidium moricei</emphasis>
(Friese, 1911)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFE19E224735FE80EC55FECD" author="Pasteels, J. J." box="[595,774,295,321]" pageId="13" pageNumber="14" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFE19E2245F1FEEBEC9DFE05" blockId="13.[151,1436,259,394]" pageId="13" pageNumber="14">
non
<taxonomicName id="40AC4D2EFFE19E2245AAFEECED35FEE9" authority="(Klug, 1832)" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[204,614,331,357]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2245AAFEECEE98FEE9" box="[204,459,331,357]" italics="true" pageId="13" pageNumber="14">Anthidium melanurum</emphasis>
(Klug, 1832)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFE19E2247E1FEECEC15FEE9" author="Warncke, K." box="[647,838,331,357]" pageId="13" pageNumber="14" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
treated
<taxonomicName id="40AC4D2EFFE19E2246CEFEECEA46FEE9" authority="Friese, 1911" authorityName="Friese" authorityYear="1911" box="[936,1301,331,357]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2246CEFEECEB2EFEE9" box="[936,1149,331,357]" italics="true" pageId="13" pageNumber="14">Anthidium moricei</emphasis>
Friese, 1911
</taxonomicName>
as a junior synonym of
<taxonomicName id="40AC4D2EFFE19E224445FEC8ED3DFE05" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[291,622,367,393]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E224445FEC8ED3DFE05" box="[291,622,367,393]" italics="true" pageId="13" pageNumber="14">Pseudoanthidium melanurum</emphasis>
</taxonomicName>
. Several authors followed him.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE19E2245F1FE10EC53FDEE" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="871336ADFFE19E2245F1FE10EC53FDEE" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
<emphasis id="B5D8EABFFFE19E2245F1FE10EE20FE5D" box="[151,371,439,465]" italics="true" pageId="13" pageNumber="14">Material examined:</emphasis>
<materialsCitation id="37C43CF0FFE19E22441FFE10EA70FE5D" box="[377,1315,439,466]" collectingDate="1909-04-06" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="ZMB" country="Palestine" latitude="38.38" location="Jericho" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="zmb179" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Dushanbe" typeStatus="holotype">
<typeStatus id="5817880FFFE19E22441FFE10EEB5FE5D" box="[377,486,439,465]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FFBB763DFFE19E224488FE10EDD6FE5D" box="[494,645,439,465]" name="Palestine" pageId="13" pageNumber="14">PALESTINE</collectingCountry>
:
<specimenCount id="91AAFD24FFE19E2247E8FE1FEDF0FE5E" box="[654,675,440,466]" count="1" pageId="13" pageNumber="14" type="female"></specimenCount>
,
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,
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,
<geoCoordinate id="E298506AFFE19E224611FE10EC86FE5E" box="[887,981,439,466]" degrees="35.46" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="35.46">35.46°E</geoCoordinate>
,
<date id="F312106DFFE19E2246B9FE10EB14FE5D" box="[991,1095,439,465]" pageId="13" pageNumber="14" value="1909-04-06">
<collectingDate id="E356E985FFE19E2246B9FE10EB14FE5D" box="[991,1095,439,465]" pageId="13" pageNumber="14" value="1909-04-06">6.iv.1909</collectingDate>
</date>
(
<collectionCode id="E1BDAE68FFE19E224132FE10EBC5FE5D" box="[1108,1174,439,465]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="13" pageNumber="14" type="Museum">ZMB</collectionCode>
:
<specimenCode id="D70A9ED6FFE19E2241F9FE10EBA9FE5D" box="[1183,1274,439,465]" pageId="13" pageNumber="14">zmb179</specimenCode>
).—
</materialsCitation>
<materialsCitation id="37C43CF0FFE19E224045FE10EACFFE7A" collectingDate="1989-04-08" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="CMK" collectorName="J. Gusenleitner" country="Jordan" latitude="38.38" location="JORDAN" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="ms0956" specimenCount="1" specimenCount-female="6" specimenCount-male="1" stateProvince="Dushanbe" typeStatus="holotype">
<collectingCountry id="FFBB763DFFE19E224045FE10EACAFE5D" box="[1315,1433,439,465]" name="Jordan" pageId="13" pageNumber="14">JORDAN</collectingCountry>
:
<specimenCount id="91AAFD24FFE19E2245F1FE7CEFEFFE79" box="[151,188,475,501]" count="1" pageId="13" pageNumber="14" type="male">1♂</specimenCount>
,
<locationDeviation id="96A12F8FFFE19E2245AEFE7CEED9FE79" box="[200,394,475,502]" location="Aqaba" pageId="13" pageNumber="14">
<quantity id="40549B48FFE19E2245AEFE7CEE5CFE79" box="[200,271,475,501]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="km" value="80.0">80 km</quantity>
NE Aqaba
</locationDeviation>
(road to Amman) (c.
<geoCoordinate id="E298506AFFE19E22471BFE7CED8DFE79" box="[637,734,475,501]" degrees="30.17" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="555" value="30.17">30.17°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFE19E22478FFE7CEC1BFE7A" box="[745,840,475,502]" degrees="35.42" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="35.42">35.42°E</geoCoordinate>
),
<date id="F312106DFFE19E22463AFE7CEC87FE79" box="[860,980,475,501]" pageId="13" pageNumber="14" value="1989-04-08">
<collectingDate id="E356E985FFE19E22463AFE7CEC87FE79" box="[860,980,475,501]" pageId="13" pageNumber="14" value="1989-04-08">08.iv.1989</collectingDate>
</date>
,
<collectorName id="2A59537BFFE19E224686FE7BEBDBFE79" box="[992,1160,475,502]" pageId="13" pageNumber="14">J. Gusenleitner</collectorName>
leg. (
<collectionCode id="E1BDAE68FFE19E2241A1FE7CEA5FFE79" box="[1223,1292,475,501]" pageId="13" pageNumber="14">CMK</collectionCode>
:
<specimenCode id="D70A9ED6FFE19E224071FE7CEA20FE7A" box="[1303,1395,475,502]" pageId="13" pageNumber="14">ms0956</specimenCode>
).—
</materialsCitation>
<materialsCitation id="37C43CF0FFE19E2245F1FE58EBA7FD95" box="[151,1268,511,538]" collectingDate="1980-06-04" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="CMK" collectorName="Z. Padr" country="Kazakhstan" latitude="38.38" location="Taras" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="ms3870" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Dushanbe" typeStatus="holotype">
<collectingCountry id="FFBB763DFFE19E2245F1FE58EE0AFD95" box="[151,345,511,537]" name="Kazakhstan" pageId="13" pageNumber="14">KAZAKHSTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFE19E224406FE58EED0FD95" box="[352,387,511,537]" count="1" pageId="13" pageNumber="14" type="female">1♀</specimenCount>
,
<location id="82736076FFE19E2244EBFDA7EE9DFD96" LSID="urn:lsid:plazi:treatment:0F0587BBFFE19E2045F1FF30EB88FCFA:82736076FFE19E2244EBFDA7EE9DFD96" box="[397,462,512,538]" country="Kazakhstan" latitude="38.38" longLatPrecision="705" longitude="69.29" name="Taras" pageId="13" pageNumber="14" stateProvince="Dushanbe">Taras</location>
:Akyrtobe(
<geoCoordinate id="E298506AFFE19E22472FFE58EDFAFD95" box="[585,681,511,537]" degrees="42.99" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="555" value="42.99">42.99°N</geoCoordinate>
<geoCoordinate id="E298506AFFE19E2247CBFE58EC5FFD95" box="[685,780,511,537]" degrees="72.09" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="72.09">72.09°E</geoCoordinate>
),
<date id="F312106DFFE19E22467AFE58ECC4FD95" box="[796,919,511,537]" pageId="13" pageNumber="14" value="1980-06-04">
<collectingDate id="E356E985FFE19E22467AFE58ECC4FD95" box="[796,919,511,537]" pageId="13" pageNumber="14" value="1980-06-04">04.vi.1980</collectingDate>
</date>
,
<collectorName id="2A59537BFFE19E2246F9FDA7ECBDFD95" box="[927,1006,511,538]" pageId="13" pageNumber="14">Z. Padr</collectorName>
leg. (
<collectionCode id="E1BDAE68FFE19E224141FE58EB3FFD95" box="[1063,1132,511,537]" pageId="13" pageNumber="14">CMK</collectionCode>
:
<specimenCode id="D70A9ED6FFE19E224112FE58EB9EFD95" box="[1140,1229,511,537]" pageId="13" pageNumber="14">ms3870</specimenCode>
).—
</materialsCitation>
<materialsCitation id="37C43CF0FFE19E224192FE58EE2CFDED" collectingDate="1909-04-06" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="CMK" collectorName="K. Denes &amp; J. Halada &amp; J. Niedl" country="Tajikistan" latitude="38.38" location="Nurek E of Dushanbe" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Dushanbe">
<collectingCountry id="FFBB763DFFE19E224192FE58EACAFD95" box="[1268,1433,511,537]" name="Tajikistan" pageId="13" pageNumber="14">TAJIKISTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFE19E2245F1FD84EFEFFDB1" box="[151,188,547,573]" count="4" pageId="13" pageNumber="14" type="female">4♀</specimenCount>
,
<location id="82736076FFE19E2245A1FD84EEEBFDB1" LSID="urn:lsid:plazi:treatment:0F0587BBFFE19E2045F1FF30EB88FCFA:82736076FFE19E2245A1FD84EEEBFDB1" box="[199,440,547,574]" country="Tajikistan" latitude="38.38" longLatPrecision="705" longitude="69.29" name="Nurek E of Dushanbe" pageId="13" pageNumber="14" stateProvince="Dushanbe">Nurek E of Dushanbe</location>
(
<geoCoordinate id="E298506AFFE19E2244A0FD84ED75FDB1" box="[454,550,547,573]" degrees="38.38" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="555" value="38.38">38.38°N</geoCoordinate>
<geoCoordinate id="E298506AFFE19E22474AFD84EDD8FDB1" box="[556,651,547,573]" degrees="69.29" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="69.29">69.29°E</geoCoordinate>
),
<collectingDate id="E356E985FFE19E2247F8FD84EB01FDB1" box="[670,1106,547,574]" isEnumeration="true" pageId="13" pageNumber="14">
<date id="F312106DFFE19E2247F8FD84EC4BFDB1" box="[670,792,547,573]" pageId="13" pageNumber="14" value="1966-06-09">09.vi.1966</date>
,
<date id="F312106DFFE19E224645FD84ECCFFDB2" box="[803,924,547,574]" pageId="13" pageNumber="14" value="1976-06-25">25.vi.1976</date>
,
<date id="F312106DFFE19E2246C1FD84EB01FDB1" box="[935,1106,547,573]" pageId="13" pageNumber="14" value="1990-06-06" valueMax="1990-06-07" valueMin="1990-06-06">06.07.vi.1990</date>
</collectingDate>
,
<collectorName id="2A59537BFFE19E22413BFD83EB9BFDB1" box="[1117,1224,547,574]" pageId="13" pageNumber="14">K. Deneš</collectorName>
,
<collectorName id="2A59537BFFE19E2241B4FD83EA6FFDB1" box="[1234,1340,547,574]" pageId="13" pageNumber="14">J. Halada</collectorName>
,
<collectorName id="2A59537BFFE19E224020FD83EACFFDB1" box="[1350,1436,547,574]" pageId="13" pageNumber="14">J. Niedl</collectorName>
leg. (
<collectionCode id="E1BDAE68FFE19E2245A9FDE0EE47FDED" box="[207,276,583,609]" pageId="13" pageNumber="14">CMK</collectionCode>
: ms3740,
</materialsCitation>
<materialsCitation id="37C43CF0FFE19E2244E0FDE0EEB6FDED" box="[390,485,583,609]" collectingDate="1909-04-06" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="CMK" collectorName="K. Denes &amp; J. Halada &amp; J. Niedl" country="Tajikistan" latitude="38.38" location="Nurek E of Dushanbe" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="ms3832" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Dushanbe">ms3832,</materialsCitation>
<materialsCitation id="37C43CF0FFE19E22448BFDE0ED1EFDED" box="[493,589,583,609]" collectingDate="1909-04-06" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode="CMK" collectorName="K. Denes &amp; J. Halada &amp; J. Niedl" country="Tajikistan" latitude="38.38" location="Nurek E of Dushanbe" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="ms3869" specimenCount-female="1" stateProvince="Dushanbe">ms3869;</materialsCitation>
<materialsCitation id="37C43CF0FFE19E224732FDE0EC53FDEE" box="[596,768,583,610]" collectingDate="1909-04-06" collectingDateMax="1989-04-08" collectingDateMin="1909-04-06" collectionCode=" OLL" collectorName="K. Denes &amp; J. Halada &amp; J. Niedl" country="Tajikistan" latitude="38.38" location="Nurek E of Dushanbe" longLatPrecision="705" longitude="69.29" pageId="13" pageNumber="14" specimenCode="oll1056" specimenCount-male="1" stateProvince="Dushanbe">
<collectionCode id="E1BDAE68FFE19E224732FDE0EDDCFDED" box="[596,655,583,609]" pageId="13" pageNumber="14">OLL</collectionCode>
: oll1056).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE19E2245A1FDCCEC69FD7D" pageId="13" pageNumber="14" type="multiple">
<paragraph id="871336ADFFE19E2245A1FDCCED3CFD42" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
<bibRefCitation id="E33D4B5CFFE19E2245A1FDCCEE2DFD09" author="Warncke, K." box="[199,382,619,645]" pageId="13" pageNumber="14" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
regarded
<taxonomicName id="40AC4D2EFFE19E224497FDCCED3BFD09" authorityName="Friese" authorityYear="1911" box="[497,616,619,645]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E224497FDCCED3BFD09" box="[497,616,619,645]" italics="true" pageId="13" pageNumber="14">A. moricei</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="40AC4D2EFFE19E2246E4FDCCEB70FD09" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[898,1059,619,645]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2246E4FDCCEB70FD09" box="[898,1059,619,645]" italics="true" pageId="13" pageNumber="14">A. melanurum</emphasis>
</taxonomicName>
, simply noting “matches with
<taxonomicName id="40AC4D2EFFE19E2240E2FDCCEE44FD25" baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2240E2FDCCEE44FD25" italics="true" pageId="13" pageNumber="14">A. melanurum</emphasis>
</taxonomicName>
” (“stimmt … mit
<taxonomicName id="40AC4D2EFFE19E224494FD28EDC6FD25" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[498,661,655,681]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E224494FD28EDC6FD25" box="[498,661,655,681]" italics="true" pageId="13" pageNumber="14">A. melanurum</emphasis>
</taxonomicName>
überein”). However, the two species are so different that it seems unlikely that he ever examined
<taxonomicName id="40AC4D2EFFE19E224491FD13ED3AFD41" authorityName="Friese" authorityYear="1911" box="[503,617,692,717]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E224491FD13ED3AFD41" box="[503,617,692,717]" italics="true" pageId="13" pageNumber="14">P. moricei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="871336ADFFE19E2245A1FD7FEC69FD7D" blockId="13.[151,1437,439,1978]" box="[199,826,727,754]" pageId="13" pageNumber="14">
<emphasis id="B5D8EABFFFE19E2245A1FD7FED58FD7D" box="[199,523,727,754]" italics="true" pageId="13" pageNumber="14">Genetic barcode information</emphasis>
. No information available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE19E2245A1FD5BEDE0FADA" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="871336ADFFE19E2245A1FD5BED10FCD2" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
<emphasis id="B5D8EABFFFE19E2245A1FD5BEE6AFC99" box="[199,313,764,789]" italics="true" pageId="13" pageNumber="14">Diagnosis</emphasis>
. A dark
<taxonomicName id="40AC4D2EFFE19E2244C7FD5CED93FC99" box="[417,704,763,789]" pageId="13" pageNumber="14">
<emphasis id="B5D8EABFFFE19E2244C7FD5CED36FC99" box="[417,613,763,789]" italics="true" pageId="13" pageNumber="14">Pseudoanthidium</emphasis>
species
</taxonomicName>
and one of the two described species with dark (black or dark chestnut-brown) legs. This feature is only shared with
<taxonomicName id="40AC4D2EFFE19E224666FC87ECF2FCB5" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[768,929,799,825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE19E224666FC87ECF2FCB5" box="[768,929,799,825]" italics="true" pageId="13" pageNumber="14">P. obscuratum</emphasis>
</taxonomicName>
, which, however, has a different structure of T6. T6 structurally like
<taxonomicName id="40AC4D2EFFE19E2244C7FCE3ED6EFCD1" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[417,573,835,861]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2244C7FCE3ED6EFCD1" box="[417,573,835,861]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="871336ADFFE19E2245A1FCC0ECA9FB2D" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
The female can easily be distinguished from
<taxonomicName id="40AC4D2EFFE19E2247D8FCCFEC0AFC0D" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[702,857,871,897]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2247D8FCCFEC0AFC0D" box="[702,857,871,897]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
by several features including the absence of yellow bands on the metasomal terga (at most some yellow translucent spots on terga in
<emphasis id="B5D8EABFFFE19E22414AFC2BEBF4FC29" box="[1068,1191,907,933]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="40AC4D2EFFE19E22414AFC2BEBF3FC29" authorityName="Friese" authorityYear="1911" box="[1068,1184,907,933]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">P. moricei</taxonomicName>
)
</emphasis>
, dark legs (tibiae and tarsi at least partly yellow in
<taxonomicName id="40AC4D2EFFE19E2244BEFC17ED20FC45" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[472,627,943,969]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E2244BEFC17ED20FC45" box="[472,627,943,969]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
), presence of lateral bands of white hairs on tergal depressions of T1T5 (no conspicuous pubescence in
<taxonomicName id="40AC4D2EFFE19E22449DFC73EDC5FC61" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[507,662,979,1005]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E22449DFC73EDC5FC61" box="[507,662,979,1005]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
), outer margin of axilla rounded, not extending beyond outer margin of scutum (axilla producing laterally beyond margin of scutum in
<taxonomicName id="40AC4D2EFFE19E224614FC5FEB5EFB9D" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[882,1037,1015,1041]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E224614FC5FEB5EFB9D" box="[882,1037,1015,1041]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
), and punctation on base of clypeus irregular with distance between punctures a puncture diameter (punctation dense and regular in
<taxonomicName id="40AC4D2EFFE19E224188FBBBEADFFBB9" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[1262,1420,1051,1077]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE19E224188FBBBEADFFBB9" box="[1262,1420,1051,1077]" italics="true" pageId="13" pageNumber="14">P. melanurum</emphasis>
</taxonomicName>
). Additionally, the female of
<taxonomicName id="40AC4D2EFFE19E2244AFFBE7ED69FBD5" authorityName="Friese" authorityYear="1911" box="[457,570,1088,1113]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2244AFFBE7ED69FBD5" box="[457,570,1088,1113]" italics="true" pageId="13" pageNumber="14">P. moricei</emphasis>
</taxonomicName>
is distinguished from the female of
<taxonomicName id="40AC4D2EFFE19E2246ADFBE7EB73FBD5" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[971,1056,1088,1113]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE19E2246ADFBE7EB73FBD5" box="[971,1056,1088,1113]" italics="true" pageId="13" pageNumber="14">P. royoi</emphasis>
</taxonomicName>
, by its colouration (overall colour black in
<taxonomicName id="40AC4D2EFFE19E22459BFBC3EE22FBF1" authorityName="Friese" authorityYear="1911" box="[253,369,1124,1149]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E22459BFBC3EE22FBF1" box="[253,369,1124,1149]" italics="true" pageId="13" pageNumber="14">P. moricei</emphasis>
</taxonomicName>
and orange in
<taxonomicName id="40AC4D2EFFE19E224744FBC3ED29FBF1" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[546,634,1124,1149]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE19E224744FBC3ED29FBF1" box="[546,634,1124,1149]" italics="true" pageId="13" pageNumber="14">P. royoi</emphasis>
</taxonomicName>
), and by a much less protruding apical margin of the clypeus and an impunctate margin of the terga (punctures reaching tergal margin in
<taxonomicName id="40AC4D2EFFE19E2246F2FB2FECBAFB2D" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[916,1001,1160,1185]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE19E2246F2FB2FECBAFB2D" box="[916,1001,1160,1185]" italics="true" pageId="13" pageNumber="14">P. royoi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="871336ADFFE19E2245A1FB0CEDE0FADA" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
The male shares with
<taxonomicName id="40AC4D2EFFE19E2244A6FB0BED32FB49" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[448,609,1195,1221]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE19E2244A6FB0BED32FB49" box="[448,609,1195,1221]" italics="true" pageId="13" pageNumber="14">P. obscuratum</emphasis>
</taxonomicName>
the specialized hairs on the sterna, but is distinct, for example, in having stronger tapering mid and hind tibiae compared to
<taxonomicName id="40AC4D2EFFE19E2247A0FB77EC36FB65" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[710,869,1231,1257]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE19E2247A0FB77EC36FB65" box="[710,869,1231,1257]" italics="true" pageId="13" pageNumber="14">P. obscuratum</emphasis>
</taxonomicName>
, a deeper apical emargination of T7, and a stronger lateral projection of T7.The gonostyli are mitten-shaped in
<taxonomicName id="40AC4D2EFFE19E224679FB53ECDDFA81" authorityName="Friese" authorityYear="1911" box="[799,910,1268,1293]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E224679FB53ECDDFA81" box="[799,910,1268,1293]" italics="true" pageId="13" pageNumber="14">P. moricei</emphasis>
</taxonomicName>
, while they are paddle-shaped in
<taxonomicName id="40AC4D2EFFE19E22419EFB53EAC6FA81" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[1272,1429,1267,1293]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE19E22419EFB53EAC6FA81" box="[1272,1429,1267,1293]" italics="true" pageId="13" pageNumber="14">P. obscuratum</emphasis>
</taxonomicName>
. The distance between the anterior and posterior ocellus is about one ocellus diameter in
<taxonomicName id="40AC4D2EFFE19E2241ECFABFEBADFABD" authorityName="Friese" authorityYear="1911" box="[1162,1278,1304,1329]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE19E2241ECFABFEBADFABD" box="[1162,1278,1304,1329]" italics="true" pageId="13" pageNumber="14">P. moricei</emphasis>
</taxonomicName>
, whereas it is about half an ocellus diameter in
<taxonomicName id="40AC4D2EFFE19E22476AFA9BEDFFFAD9" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[524,684,1339,1365]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE19E22476AFA9BEDFFFAD9" box="[524,684,1339,1365]" italics="true" pageId="13" pageNumber="14">P. obscuratum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE19E2045A1FAC7EA3EFCDE" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="871336ADFFE19E2245A1FAC7EDE8F835" blockId="13.[151,1437,439,1978]" pageId="13" pageNumber="14">
<emphasis id="B5D8EABFFFE19E2245A1FAC7EEE1FAF5" box="[199,434,1375,1401]" italics="true" pageId="13" pageNumber="14">Description. Female</emphasis>
(
<figureCitation id="1F972A28FFE19E2244A3FAF8ED48FAF5" box="[453,539,1375,1401]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="13" pageNumber="14">Fig. 7a</figureCitation>
).
<quantity id="40549B48FFE19E224752FAF8ED2FFAF6" box="[564,636,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="mm" value="9.0">9 mm</quantity>
.—
<emphasis id="B5D8EABFFFE19E2247FAFAF8ED8AFAF5" box="[668,729,1375,1401]" italics="true" pageId="13" pageNumber="14">Head</emphasis>
: Clypeus bell-shaped with inwardly curved anterior margin; clypeal apex fully exposed and protruding, thickened, with two erect tubercles on each side (
<figureCitation id="1F972A28FFE19E2241C7FA24EBA7FA11" box="[1185,1268,1411,1437]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="13" pageNumber="14">Fig. 7d</figureCitation>
); apical part of the ridge with short greyish-brown appressed hairs; basal third of clypeus with scattered punctation, with punctures 23 puncture diameter apart; surface polished; punctation denser and finer towards apical part, with subapical area transversely depressed; mandible chestnut brown with five black teeth (contrary to Friese 1911, who wrongly stated only three teeth); apical tooth slightly stronger than inner teeth, which are equally strong; eyes slightly convergent; head black, with a yellow-brown spot in lower paraocular area and a small triangular yellow spot behind the eye; in the
<typeStatus id="5817880FFFE19E2245A6F9FCEE76F9F9" box="[192,293,1627,1653]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, ocelloocular distance
<quantity id="40549B48FFE19E224743F9FCEDD8F9FA" box="[549,651,1627,1654]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="13" pageNumber="14" unit="mm" value="0.78">0.78 mm</quantity>
, ocellooccipital distance 0,
<quantity id="40549B48FFE19E2246DFF9FCEB58F9FA" box="[953,1035,1627,1654]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="13" pageNumber="14" unit="mm" value="91.0">91 mm</quantity>
, i.e. ocellooccipital distance greater than ocelloocular distance; long white pubescence around antennal sockets; head densely punctate.—
<taxonomicName id="40AC4D2EFFE19E22407BF927EAC6F915" box="[1309,1429,1664,1689]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE19E22407BF927EAC6F915" box="[1309,1429,1664,1689]" italics="true" pageId="13" pageNumber="14">Mesosoma</emphasis>
</taxonomicName>
: Black, densely punctate like the head; pronotal lobe strongly lamellate; in profile, scutellum angular laterally and rounded medially; slightly overhanging propodeum; basal zone of propodeum with fine and dense punctation, followed by a zone of large and coarse punctures; propodeal triangle smooth and shining.—
<emphasis id="B5D8EABFFFE19E2241C7F94BEA45F889" box="[1185,1302,1772,1797]" italics="true" pageId="13" pageNumber="14">Metasoma</emphasis>
: Punctation on tergal discs less dense (punctures 12 puncture diameter apart) than on depression; narrow impunctate marginal zone; T6 with brown median protrusion, deeply emarginate medially; punctation on T6 very fine, rugulose; T1 with dense, white brush of bristles laterally; T2T4 with lateral bands of white bristles; T5 with uninterrupted fringe of white hairs; scopa white (
<figureCitation id="1F972A28FFE19E2244DDF8DCED5FF819" box="[443,524,1915,1941]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="13" pageNumber="14">Fig. 7c</figureCitation>
).—
<emphasis id="B5D8EABFFFE19E224751F8DBED3FF819" box="[567,620,1916,1941]" italics="true" pageId="13" pageNumber="14">Legs</emphasis>
: Chestnut-brown with white hairs; yellow bristles on inner face of hind basitarsus.—
<emphasis id="B5D8EABFFFE19E22444FF807EE30F835" box="[297,355,1952,1977]" italics="true" pageId="13" pageNumber="14">Wing</emphasis>
: Slightly brownish infuscated.
</paragraph>
<caption id="D3D36625FFE29E2145F1F827EDD2F86C" ID-DOI="http://doi.org/10.5281/zenodo.14240751" ID-Zenodo-Dep="14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1920,1944]" targetBox="[152,1436,169,1896]" targetPageId="14" targetType="figure">
<paragraph id="871336ADFFE29E2145F1F827EDD2F86C" blockId="14.[151,1437,1920,2016]" pageId="14" pageNumber="15">
<emphasis id="B5D8EABFFFE29E2145F1F827EE46F815" bold="true" box="[151,277,1920,1945]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
<taxonomicName id="40AC4D2EFFE29E21447BF827EDE9F815" authority="(Friese, 1911)" baseAuthorityName="Friese" baseAuthorityYear="1911" box="[285,698,1920,1945]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="moricei">
<emphasis id="B5D8EABFFFE29E21447BF827ED70F814" box="[285,547,1920,1944]" italics="true" pageId="14" pageNumber="15">Pseudoanthidium moricei</emphasis>
(Friese, 1911)
</taxonomicName>
<emphasis id="B5D8EABFFFE29E2147A5F826EC32F815" bold="true" box="[707,865,1921,1945]" pageId="14" pageNumber="15">stat. resurrect.</emphasis>
a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan).
</paragraph>
</caption>
<paragraph id="871336ADFFE39E2045A1FF30EA3FFEB3" blockId="15.[151,1437,151,887]" pageId="15" pageNumber="16">
<emphasis id="B5D8EABFFFE39E2045A1FF30EE7CFF3C" box="[199,303,151,176]" italics="true" pageId="15" pageNumber="16">Variation</emphasis>
. The
<typeStatus id="5817880FFFE39E204416FF30EE81FF3D" box="[368,466,151,177]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
from
<collectingCountry id="FFBB763DFFE39E204770FF30ED28FF3D" box="[534,635,151,177]" name="Palestine" pageId="15" pageNumber="16">Palestine</collectingCountry>
exhibits a black face except for a small yellowish indistinct spot on the lower paraocular area.
<location id="82736076FFE39E2044FFFF1CEEE3FF59" LSID="urn:lsid:plazi:treatment:0F0587BBFFE19E2045F1FF30EB88FCFA:82736076FFE39E2044FFFF1CEEE3FF59" box="[409,432,187,213]" country="Palestine" county="Some" municipality="Central Asian" name="In" pageId="15" pageNumber="16">In</location>
specimens from
<location id="82736076FFE39E204717FF1DEDACFF58" LSID="urn:lsid:plazi:treatment:0F0587BBFFE19E2045F1FF30EB88FCFA:82736076FFE39E204717FF1DEDACFF58" box="[625,767,186,212]" country="Palestine" county="Some" municipality="Central Asian" name="Central Asia" pageId="15" pageNumber="16">Central Asia</location>
, this light area is present as a dirty yellow area in the lower paraocular area.
<location id="82736076FFE39E204432FF79EE38FF74" LSID="urn:lsid:plazi:treatment:0F0587BBFFE19E2045F1FF30EB88FCFA:82736076FFE39E204432FF79EE38FF74" box="[340,363,222,248]" country="Palestine" county="Some" municipality="Central Asian" name="In" pageId="15" pageNumber="16">In</location>
some
<collectingMunicipality id="6777ACD7FFE39E2044D1FF79ED02FF74" box="[439,593,222,248]" pageId="15" pageNumber="16">Central Asian</collectingMunicipality>
specimens, lower half of clypeus yellowish-brown, and mandible yellow-brown.
<collectingCounty id="6E724E21FFE39E20458BFEA6EE7EFE97" box="[237,301,257,283]" pageId="15" pageNumber="16">Some</collectingCounty>
individuals from
<collectingCountry id="FFBB763DFFE39E204492FEA6ED31FE97" box="[500,610,257,283]" name="Tajikistan" pageId="15" pageNumber="16">Tajikistan</collectingCountry>
show remnants of yellow spots of scutellum, axillae, and
<specimenCode id="D70A9ED6FFE39E20418AFEA6EA5FFE97" box="[1260,1292,257,283]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T2</specimenCode>
<specimenCode id="D70A9ED6FFE39E20407AFEA6EA6FFE97" box="[1308,1340,257,283]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T4</specimenCode>
. Ground colour of the tergal integument dark brown instead of black. White hairs on terga restricted to the lateral areas.
</paragraph>
<paragraph id="871336ADFFE39E2045A1FEEEEA3EFCDE" blockId="15.[151,1437,151,887]" pageId="15" pageNumber="16">
<emphasis id="B5D8EABFFFE39E2045A1FEEEEE18FEEE" box="[199,331,329,354]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
.
<emphasis id="B5D8EABFFFE39E204439FEEFEECAFEEE" box="[351,409,328,354]" italics="true" pageId="15" pageNumber="16">Male</emphasis>
(
<figureCitation id="1F972A28FFE39E2044CBFEEFED55FEEE" box="[429,518,328,354]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="15" pageNumber="16">Fig. 7b</figureCitation>
).
<quantity id="40549B48FFE39E204746FEEFED25FEEF" box="[544,630,328,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="10.0">10 mm</quantity>
.—
<emphasis id="B5D8EABFFFE39E2047F1FEEFED87FEEE" box="[663,724,328,354]" italics="true" pageId="15" pageNumber="16">Head</emphasis>
: Black with pale yellow clypeus and lower paraocular area reaching upper level of clypeus; minute yellow spot behind the zenith of the eye; clypeus strongly arcuate upwards; punctation on the upper clypeus sparse and irregular, but lower clypeus densely punctate; face covered by long, dense pubescence, also hiding the apical margin of clypeus; mandible relatively short with three strong teeth; black, sides yellow; distance between anterior and posterior ocellus approximately one ocellus diameter.—
<taxonomicName id="40AC4D2EFFE39E20407BFE70EAC6FE7C" box="[1309,1429,471,496]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE39E20407BFE70EAC6FE7C" box="[1309,1429,471,496]" italics="true" pageId="15" pageNumber="16">Mesosoma</emphasis>
</taxonomicName>
: Black with long white pubescence; pronotal lobe with high lamella difficult to detect under the hairs; scutum with dense punctation, with distance between punctures less than half their diameter; interstices shining; scutellum with angulate outer margin, slightly depressed medially.—
<emphasis id="B5D8EABFFFE39E204641FDE6EC0FFDD6" box="[807,860,577,602]" italics="true" pageId="15" pageNumber="16">Legs</emphasis>
: Black with white pubescence, slender (
<figureCitation id="1F972A28FFE39E204047FDE7EA23FDD6" box="[1313,1392,576,602]" captionStart="FIGURE 7" captionStartId="14.[151,250,1920,1944]" captionTargetBox="[152,1436,169,1896]" captionTargetId="figure-19@14.[152,1436,159,1896]" captionTargetPageId="14" captionText="FIGURE 7. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Habitus dorsal, female (holotype from Palestine); b. Habitus of a male from Jordan; c. Apical terga of the female (holotype from Palestine); d. Head from lateral, female (holotype from Palestine); e. Hind leg (male from Jordan)." figureDoi="http://doi.org/10.5281/zenodo.14240751" httpUri="https://zenodo.org/record/14240751/files/figure.png" pageId="15" pageNumber="16">Fig. 7e</figureCitation>
).—
<emphasis id="B5D8EABFFFE39E2045F1FDC2EE5FFDF2" box="[151,268,613,638]" italics="true" pageId="15" pageNumber="16">Metasoma</emphasis>
: Black with white pubescence especially on apical terga; integument shining; punctures somewhat smaller and denser in marginal zone than on disc; disc little elevated over depression; apical margin impunctate; remnants of a yellow spot close to the midline on T5; T5 with a small lateral bulge; T6 with a small bulge close to the middle and a bigger one laterally; the outer bulge overhangs the apical margin; T7 with a lateral, triangular projection and a deep median emargination; T7 medially depressed; punctation on T6 and T7 coarse and irregular; S3 with undulate apical margin and apically waved hairs; S5 with black basic comb and lateral arm with black comb (approximately 1820 digits).—
<emphasis id="B5D8EABFFFE39E204464FC9FEE3FFCDE" box="[258,364,824,850]" italics="true" pageId="15" pageNumber="16">Genitalia</emphasis>
: Gonostylus much longer than penis valves; penis valve a thin, slightly inward sloping rod.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE39E2045A1FCFBEB88FCFA" box="[199,1243,860,887]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="871336ADFFE39E2045A1FCFBEB88FCFA" blockId="15.[151,1437,151,887]" box="[199,1243,860,887]" pageId="15" pageNumber="16">
<emphasis id="B5D8EABFFFE39E2045A1FCFBEE03FCFA" box="[199,336,860,886]" italics="true" pageId="15" pageNumber="16">Distribution</emphasis>
. Levant (
<collectingCountry id="FFBB763DFFE39E2044DCFCFBED77FCFA" box="[442,548,860,886]" name="Palestine" pageId="15" pageNumber="16">Palestine</collectingCountry>
,
<collectingCountry id="FFBB763DFFE39E204749FCFBED2FFCFA" box="[559,636,860,886]" name="Jordan" pageId="15" pageNumber="16">Jordan</collectingCountry>
) and Central Asia (
<collectingCountry id="FFBB763DFFE39E204635FCFBEC89FCFA" box="[851,986,860,886]" name="Kazakhstan" pageId="15" pageNumber="16">Kazakhstan</collectingCountry>
,
<collectingCountry id="FFBB763DFFE39E204683FCFBEB06FCFA" box="[997,1109,860,886]" name="Tajikistan" pageId="15" pageNumber="16">Tajikistan</collectingCountry>
) (
<figureCitation id="1F972A28FFE39E20410AFCFBEB9DFCFA" box="[1132,1230,860,886]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="15" pageNumber="16">Fig. 27b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="C67C2C1C080FBFBDB5C4F309EEFD5173" ID-DOI="10.11646/zootaxa.5541.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240734" ID-ZooBank="A5D2633B-36A8-42A4-AD21-22041804BE95" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732832365526" checkinUser="plazi" docAuthor="Kasparek, Max" docDate="2024" docId="0F0587BBFFE39E3D45F1F9E4EC0DFECD" docLanguage="en" docName="zootaxa.5541.1.1.pdf" docOrigin="Zootaxa 5541 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5541.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoanthidium obscuratum" docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="F33CFFC3FFEC9E2F4566FFA7EF53FF8C" masterDocTitle="New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="16" updateTime="1733329203150" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="71E58EE0A63A5C2595B6DAE485ACB02F">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<mods:namePart id="B6A79A4E0E8DA8CAAF379D455BDEAD2C">Kasparek, Max</mods:namePart>
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<treatment id="0F0587BBFFE39E3D45F1F9E4EC0DFECD" ID-DOI="http://doi.org/10.5281/zenodo.14248295" ID-Zenodo-Dep="14248295" LSID="urn:lsid:plazi:treatment:0F0587BBFFE39E3D45F1F9E4EC0DFECD" httpUri="http://treatment.plazi.org/id/0F0587BBFFE39E3D45F1F9E4EC0DFECD" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
<subSubSection id="CFB66526FFE39E2045F1F9E4EDB1F9D1" box="[151,738,1603,1630]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="871336ADFFE39E2045F1F9E4EDB1F9D1" blockId="15.[151,738,1603,1666]" box="[151,738,1603,1630]" pageId="15" pageNumber="16">
<heading id="DC5B81C1FFE39E2045F1F9E4EDB1F9D1" bold="true" box="[151,738,1603,1630]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="40AC4D2EFFE39E2045F1F9E4EDB1F9D1" ID-CoL="4NP4N" authority="(Morawitz, 1875)" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[151,738,1603,1630]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE39E2045F1F9E4EDB1F9D1" bold="true" box="[151,738,1603,1630]" pageId="15" pageNumber="16">
<emphasis id="B5D8EABFFFE39E2045F1F9E4ED50F9D2" bold="true" box="[151,515,1603,1630]" italics="true" pageId="15" pageNumber="16">Pseudoanthidium obscuratum</emphasis>
(
<bibRefCitation id="E33D4B5CFFE39E204775F9E4ED8AF9D1" author="Morawitz, F. F." box="[531,729,1603,1629]" pageId="15" pageNumber="16" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz, 1875</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE39E2045F1F9CFEEB8F90E" box="[151,491,1639,1666]" pageId="15" pageNumber="16" type="description">
<paragraph id="871336ADFFE39E2045F1F9CFEEB8F90E" blockId="15.[151,738,1603,1666]" box="[151,491,1639,1666]" pageId="15" pageNumber="16">
(
<figureCitation id="1F972A28FFE39E2045F9F9CFEFBBF90E" box="[159,232,1640,1666]" captionStart="FIGURE 9" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1893]" captionTargetId="figure-19@16.[152,1436,181,1918]" captionTargetPageId="16" captionText="FIGURE 9. Pseudoanthidium obscuratum (Morawitz, 1875). a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan)." figureDoi="http://doi.org/10.5281/zenodo.14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="15" pageNumber="16">Figs 9</figureCitation>
,
<figureCitation id="1F972A28FFE39E204592F9CFEE40F90E" box="[244,275,1640,1666]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="15" pageNumber="16">20</figureCitation>
,
<figureCitation id="1F972A28FFE39E204478F9CFEE6FF90E" box="[286,316,1640,1666]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="15" pageNumber="16">21</figureCitation>
,
<figureCitation id="1F972A28FFE39E20442EF9CFEE34F90E" box="[328,359,1640,1666]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="15" pageNumber="16">22</figureCitation>
,
<figureCitation id="1F972A28FFE39E204414F9CFEEC3F90E" box="[370,400,1640,1666]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="15" pageNumber="16">23</figureCitation>
,
<figureCitation id="1F972A28FFE39E2044FAF9CFEEE9F90E" box="[412,442,1640,1666]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="15" pageNumber="16">24</figureCitation>
, 25)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE39E2045F1F90BECE0F96F" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="871336ADFFE39E2045F1F90BEC44F948" blockId="15.[151,947,1708,1763]" box="[151,791,1708,1732]" pageId="15" pageNumber="16">
<taxonomicName id="40AC4D2EFFE39E2045F1F90BED75F948" ID-CoL="8SQP8" authority="Morawitz, 1875" authorityName="Morawitz" authorityYear="1875" box="[151,550,1708,1732]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE39E2045F1F90BEE2CF948" box="[151,383,1708,1732]" italics="true" pageId="15" pageNumber="16">Anthidium obscuratum</emphasis>
<bibRefCitation id="E33D4B5CFFE39E2044E3F90BED75F948" author="Morawitz, F. F." box="[389,550,1708,1732]" pageId="15" pageNumber="16" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz, 1875</bibRefCitation>
</taxonomicName>
.—Female,
<collectingCountry id="FFBB763DFFE39E2047F8F90BEC40F948" box="[670,787,1708,1732]" name="Uzbekistan" pageId="15" pageNumber="16">Uzbekistan</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFE39E2045F1F96CECE0F96F" blockId="15.[151,947,1708,1763]" box="[151,947,1739,1763]" pageId="15" pageNumber="16">
<taxonomicName id="40AC4D2EFFE39E2045F1F96CED74F96F" ID-CoL="8SQP8" authority="Morawitz, 1875" authorityName="Morawitz" authorityYear="1875" box="[151,551,1739,1763]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE39E2045F1F96CEE2CF96F" box="[151,383,1739,1763]" italics="true" pageId="15" pageNumber="16">Anthidium obscuratum</emphasis>
<bibRefCitation id="E33D4B5CFFE39E2044E3F96CED74F96F" author="Morawitz, F. F." box="[389,551,1739,1763]" pageId="15" pageNumber="16" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz, 1875</bibRefCitation>
</taxonomicName>
.—Male,
<collectingCountry id="FFBB763DFFE39E2047E1F96CEDAAF96F" box="[647,761,1739,1763]" name="Uzbekistan" pageId="15" pageNumber="16">Uzbekistan</collectingCountry>
(
<bibRefCitation id="E33D4B5CFFE39E204660F96CECFBF96F" author="Morawitz, F. F." box="[774,936,1739,1763]" pageId="15" pageNumber="16" pagination="3 - 87" refId="ref30969" refString="Morawitz, F. F. (1894 [1893]) Supplement zur Bienenfauna Turkestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 28, 3 - 87." type="book chapter" year="1894">Morawitz 1894</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="871336ADFFE39E2045F1F8A8EDDFF875" blockId="15.[151,1437,1806,2041]" pageId="15" pageNumber="16">
<emphasis id="B5D8EABFFFE39E2045F1F8A8EEE7F8A4" box="[151,436,1806,1832]" italics="true" pageId="15" pageNumber="16">Type material examined.</emphasis>
<materialsCitation id="37C43CF0FFE39E2044D9F8A8EDDBF875" collectingDate="1871-07-11" collectionCode="ZISP" collectorName="Aleksey Fedtschenko" latitude="39.98" location="Karakasuk" longLatPrecision="699" longitude="71.8" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="5817880FFFE39E2044D9F8A8ED7CF8A5" box="[447,559,1807,1833]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
:
<specimenCount id="91AAFD24FFE39E204722F8A9EDC0F8A4" box="[580,659,1806,1832]" pageId="15" pageNumber="16" type="female">female</specimenCount>
, “Karakasuk”,
<taxonomicName id="40AC4D2EFFE39E204620F8A9EB4BF8A5" authority="Mor." authorityName="Morawitz" authorityYear="1875" box="[838,1048,1806,1833]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE39E204634F8A9EC85F8A4" box="[850,982,1806,1832]" italics="true" pageId="15" pageNumber="16">obscuratum</emphasis>
Mor.
</taxonomicName>
Type”, “Syntypus”, “Lectotypus,
<taxonomicName id="40AC4D2EFFE39E2045F1F896EE85F8C7" authority="Mor." authorityName="Morawitz" authorityYear="1875" box="[151,470,1841,1867]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE39E2045F1F896EECBF8C7" box="[151,408,1841,1867]" italics="true" pageId="15" pageNumber="16">Anthidium obscuratum</emphasis>
Mor.
</taxonomicName>
female (Warncke 1978)” (
<collectionCode id="E1BDAE68FFE39E204665F896EC6DF8C7" box="[771,830,1841,1867]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="16">ZISP</collectionCode>
, examined by photograph). See also
<bibRefCitation id="E33D4B5CFFE39E204186F896EACBF8C7" author="Warncke, K." box="[1248,1432,1841,1867]" pageId="15" pageNumber="16" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
. This is apparently one of the specimens collected by
<collectorName id="2A59537BFFE39E204666F8F3ECA7F8E2" box="[768,1012,1876,1902]" pageId="15" pageNumber="16">Aleksey Fedtschenko</collectorName>
on
<date id="F312106DFFE39E204145F8F3EBEBF8E2" box="[1059,1208,1876,1902]" pageId="15" pageNumber="16" value="1871-07-11">
<collectingDate id="E356E985FFE39E204145F8F3EBEBF8E2" box="[1059,1208,1876,1902]" pageId="15" pageNumber="16" value="1871-07-11">11 July 1871</collectingDate>
</date>
[in Julian calendar, actually 23 July in the Gregorian calendar] (
<bibRefCitation id="E33D4B5CFFE39E204719F8D1EC7CF81D" author="Morawitz, F. F." box="[639,815,1910,1937]" pageId="15" pageNumber="16" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz 1875</bibRefCitation>
, travel itinerary by
<bibRefCitation id="E33D4B5CFFE39E204160F8D1EBDFF81C" author="Baker, D. B." box="[1030,1164,1910,1936]" pageId="15" pageNumber="16" pagination="231 - 252" refId="ref29553" refString="Baker, D. B. (2004) Type material of Hymenoptera described by O. I. Radoszkowsky in the Natural History Museum, London, and the localities of A. P. Fedtschenko's Reise in Turkestan. Mitteilungen aus dem Museum fur Naturkunde Berlin, Deutsche Entomologische Zeitschrift, 51, 231 - 252. https: // doi. org / 10.1002 / mmnd. 4810510207" type="journal article" year="2004">Baker 2004</bibRefCitation>
). The location where the material was collected is indicated as “
<location id="82736076FFE39E20475AF83EEDE8F83F" LSID="urn:lsid:plazi:treatment:0F0587BBFFE39E3D45F1F9E4EC0DFECD:82736076FFE39E20475AF83EEDE8F83F" box="[572,699,1945,1971]" latitude="39.98" longLatPrecision="699" longitude="71.8" name="Karakasuk" pageId="15" pageNumber="16">Karakasuk</location>
”, a mountain pass and rangeland area situated above Schahimardan (Shohimardon) in the
<location id="82736076FFE39E2044E0F81BED71F85A" LSID="urn:lsid:plazi:treatment:0F0587BBFFE39E3D45F1F9E4EC0DFECD:82736076FFE39E2044E0F81BED71F85A" box="[390,546,1980,2006]" latitude="39.98" longLatPrecision="699" longitude="71.8" name="Ferghana area" pageId="15" pageNumber="16">Ferghana area</location>
(approximately
<geoCoordinate id="E298506AFFE39E2047B3F81BEC67F85A" box="[725,820,1980,2006]" degrees="39.98" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="555" value="39.98">39.98°N</geoCoordinate>
<geoCoordinate id="E298506AFFE39E20465EF81BECCBF85A" box="[824,920,1980,2006]" degrees="71.80" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="555" value="71.8">71.80°E</geoCoordinate>
), probably identical with the village “Kokand”. Today, it is an Uzbek enclave in Kyrgyzstan
</materialsCitation>
.
</paragraph>
<caption id="D3D36625FFFC9E3F45F1F830EA20F85F" ID-DOI="http://doi.org/10.5281/zenodo.14240755" ID-Zenodo-Dep="14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1943,1967]" targetBox="[152,1436,181,1893]" targetPageId="16" targetType="figure">
<paragraph id="871336ADFFFC9E3F45F1F830EA20F85F" blockId="16.[151,1436,1943,2004]" pageId="16" pageNumber="17">
<emphasis id="B5D8EABFFFFC9E3F45F1F830EE46F823" bold="true" box="[151,277,1943,1967]" pageId="16" pageNumber="17">FIGURE 9.</emphasis>
<taxonomicName id="40AC4D2EFFFC9E3F4478F830EC42F823" authority="(Morawitz, 1875)" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[286,785,1943,1967]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFFC9E3F4478F830ED03F823" box="[286,592,1943,1967]" italics="true" pageId="16" pageNumber="17">Pseudoanthidium obscuratum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFC9E3F4739F830EC5BF823" author="Morawitz, F. F." box="[607,776,1943,1967]" pageId="16" pageNumber="17" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz, 1875</bibRefCitation>
)
</taxonomicName>
. a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan).
</paragraph>
</caption>
<paragraph id="871336ADFFFD9E3E45A1FF30EA1CFE7E" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
Other material that had beeen attributed to
<typeStatus id="5817880FFFFD9E3E47D8FF30EDBDFF3D" box="[702,750,151,177]" pageId="17" pageNumber="18">type</typeStatus>
material: Female, “Kumar”, “VIII.26” [year? to be read as 26. August, following
<bibRefCitation id="E33D4B5CFFFD9E3E44C0FF1DED1BFF58" author="Friese, H." box="[422,584,186,212]" pageId="17" pageNumber="18" refId="ref29768" refString="Friese, H. (1898) Die Bienen Europa's (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp." type="book" year="1898">Friese (1898)</bibRefCitation>
], “
<taxonomicName id="40AC4D2EFFFD9E3E470BFF1DEC27FF58" authorityName="Morawitz" authorityYear="1875" box="[621,884,186,212]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFFD9E3E470BFF1DEC27FF58" box="[621,884,186,212]" italics="true" pageId="17" pageNumber="18">Anthidium obscuratum</emphasis>
</taxonomicName>
”, “coll. Friese”, “
<typeStatus id="5817880FFFFD9E3E4136FF1DEBFCFF58" box="[1104,1199,186,212]" pageId="17" pageNumber="18">TYPUS</typeStatus>
”, “
<taxonomicName id="40AC4D2EFFFD9E3E41BEFF1DEDD4FF74" authority="Mor. J. Pasteels" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obscuratum" subGenus="Reanthidium">
<emphasis id="B5D8EABFFFFD9E3E41BEFF1DEACFFF58" box="[1240,1436,186,212]" italics="true" pageId="17" pageNumber="18">Pseudoanthidium</emphasis>
(
<emphasis id="B5D8EABFFFFD9E3E45C7FF79EE63FF74" box="[161,304,222,248]" italics="true" pageId="17" pageNumber="18">Reanthidium</emphasis>
)
<emphasis id="B5D8EABFFFFD9E3E4425FF79EE94FF74" box="[323,455,222,248]" italics="true" pageId="17" pageNumber="18">obscuratum</emphasis>
Mor. J. Pasteels
</taxonomicName>
det. 1964”, http://coll.mfn-berlin.de/u/97637a (
<collectionCode id="E1BDAE68FFFD9E3E41CEFF79EBB4FF74" box="[1192,1255,222,248]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="18" type="Museum">ZMB</collectionCode>
).—The locality “Kumar” could not be synonymized with certainty with a modern locality name, but seems to be Vodil, close to Shohimardon (
<geoCoordinate id="E298506AFFFD9E3E4438FE81EEEDFECC" box="[350,446,294,320]" degrees="40.17" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="40.17">40.17°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E44A1FE81ED74FECC" box="[455,551,294,320]" degrees="71.72" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="71.72">71.72°E</geoCoordinate>
).
<bibRefCitation id="E33D4B5CFFFD9E3E475DFE82ED80FECC" author="Friese, H." box="[571,723,293,320]" pageId="17" pageNumber="18" refId="ref29768" refString="Friese, H. (1898) Die Bienen Europa's (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp." type="book" year="1898">Friese (1898)</bibRefCitation>
mentioned that he possesses two typical females from Kumar (“
<taxonomicName id="40AC4D2EFFFD9E3E45CBFEEEEEE3FEEF" authorityName="Morawitz" authorityYear="1875" box="[173,432,329,355]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFFD9E3E45CBFEEEEEE3FEEF" box="[173,432,329,355]" italics="true" pageId="17" pageNumber="18">Anthidium obscuratum</emphasis>
</taxonomicName>
liegt mir in 2 typischen
<specimenCount id="91AAFD24FFFD9E3E47B3FEEDEDB9FEE8" box="[725,746,330,356]" count="1" pageId="17" pageNumber="18" type="female"></specimenCount>
von Kumar (26. August) vor”). However, the kind of label indicates that at least the specimen deposited in
<collectionCode id="E1BDAE68FFFD9E3E47D6FEC9EDBEFE04" box="[688,749,366,392]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="18" type="Museum">ZMB</collectionCode>
today was labelled as
<typeStatus id="5817880FFFFD9E3E468EFEC9EB4BFE04" box="[1000,1048,366,392]" pageId="17" pageNumber="18">type</typeStatus>
by Friese rather than by Morawitz himself. It is known that Friese had a practice of adding
<typeStatus id="5817880FFFFD9E3E467CFE36EC19FE27" box="[794,842,401,427]" pageId="17" pageNumber="18">type</typeStatus>
labels to material he later received and identified as conspecific with previously described taxa (
<bibRefCitation id="E33D4B5CFFFD9E3E47E1FE12ECEDFE43" author="Rasmussen, C. &amp; Ascher, J. S." box="[647,958,436,463]" pageId="17" pageNumber="18" pagination="1 - 118" refId="ref31580" refString="Rasmussen, C. &amp; Ascher, J. S. (2008) Heinrich Friese (1860 - 1948): Names proposed and notes on a pioneer melittologist (Hymenoptera, Anthophila). Zootaxa, 1833 (1), 1 - 118. https: // doi. org / 10.11646 / zootaxa. 1833.1.1" type="journal article" year="2008">Rasmussen &amp; Ascher 2008</bibRefCitation>
). Therefore, Frieses material from Kumar cannot be treated as
<typeStatus id="5817880FFFFD9E3E441AFE7EEEFFFE7F" box="[380,428,473,499]" pageId="17" pageNumber="18">type</typeStatus>
material, although it was probably collected in close proximity to the
<typeStatus id="5817880FFFFD9E3E41D8FE7EEBBDFE7F" box="[1214,1262,473,499]" pageId="17" pageNumber="18">type</typeStatus>
locality.
</paragraph>
<paragraph id="871336ADFFFD9E3E45A1FE5BEDE9FCF7" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1FE5BEEB4FD9A" box="[199,487,508,534]" italics="true" pageId="17" pageNumber="18">Other material examined</emphasis>
(material in
<collectionCode id="E1BDAE68FFFD9E3E47E7FE5BEDE9FD9A" box="[641,698,508,534]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
examined by photographs): KAZAKHASTAN:
<specimenCount id="91AAFD24FFFD9E3E418CFE5BEA5EFD9A" box="[1258,1293,508,534]" count="1" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
,
<specimenCount id="91AAFD24FFFD9E3E4078FE5BEA10FD9A" box="[1310,1347,508,534]" count="2" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
,
<quantity id="40549B48FFFD9E3E4034FE5BEACFFD9A" box="[1362,1436,508,535]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="km" value="35.0">35 km</quantity>
W Almaty (
<geoCoordinate id="E298506AFFFD9E3E447AFD87EE28FDB6" box="[284,379,544,570]" degrees="43.21" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="43.21">43.21°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E44E2FD87EEB0FDB6" box="[388,483,544,570]" degrees="76.44" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="76.44">76.44°E</geoCoordinate>
),
<date id="F312106DFFFD9E3E449EFD87ED2AFDB6" box="[504,633,544,570]" pageId="17" pageNumber="18" value="1981-07-16">16.vii.1981</date>
, M. Kocourek leg. (
<collectionCode id="E1BDAE68FFFD9E3E4638FD87ECF0FDB6" box="[862,931,544,570]" pageId="17" pageNumber="18">CMK</collectionCode>
: ex coll. Kocourek, ms3216, 3219, 3223).—
<collectingCountry id="FFBB763DFFFD9E3E45F1FDE3EE04FDD2" box="[151,343,580,606]" name="Kyrgyzstan" pageId="17" pageNumber="18">KYRGYZSTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFFD9E3E4406FDE3EED7FDD2" box="[352,388,580,606]" count="2" pageId="17" pageNumber="18" type="female">2♀</specimenCount>
,
<specimenCount id="91AAFD24FFFD9E3E44E9FDE3EEE7FDD2" box="[399,436,580,606]" count="4" pageId="17" pageNumber="18" type="male">4♂</specimenCount>
,
<quantity id="40549B48FFFD9E3E44D8FDE4ED50FDD1" box="[446,515,579,606]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="km" value="15.0">15 km</quantity>
<collectionCode id="E1BDAE68FFFD9E3E476EFDE3ED4BFDD2" box="[520,536,580,606]" pageId="17" pageNumber="18">S</collectionCode>
<collectingRegion id="4568F84FFFFD9E3E477AFDE4ED2FFDD1" box="[540,636,579,605]" country="Kyrgyzstan" name="Bishkek Shaary" pageId="17" pageNumber="18">Bishkek</collectingRegion>
: Con-Aryk (
<geoCoordinate id="E298506AFFFD9E3E4665FDE3EC37FDD2" box="[771,868,580,606]" degrees="42.74" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="42.74">42.74°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E460FFDE4EC9BFDD2" box="[873,968,579,606]" degrees="74.57" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="74.57">74.57°E</geoCoordinate>
),
<date id="F312106DFFFD9E3E46BFFDE4EB08FDD1" box="[985,1115,579,605]" pageId="17" pageNumber="18" value="1981-07-08">08.vii.1981</date>
, M. Kocourek leg. (
<collectionCode id="E1BDAE68FFFD9E3E4052FDE3EA2AFDD2" box="[1332,1401,580,606]" pageId="17" pageNumber="18">CMK</collectionCode>
: ex coll. Kocourek, ms3217, 3218, 3220-3222, 3224).—
<specimenCount id="91AAFD24FFFD9E3E4794FDC0EC59FD0D" box="[754,778,615,641]" count="1" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
, ibid.,
<date id="F312106DFFFD9E3E4631FDC0EC9AFD0D" box="[855,969,615,641]" pageId="17" pageNumber="18" value="1983-07-06">6.vii.1983</date>
(
<collectionCode id="E1BDAE68FFFD9E3E46BEFDC0EB47FD0D" box="[984,1044,615,641]" pageId="17" pageNumber="18">OLL</collectionCode>
: oll1058).—
<specimenCount id="91AAFD24FFFD9E3E41CBFDC0EB95FD0E" box="[1197,1222,615,642]" count="1" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
,
<collectingRegion id="4568F84FFFFD9E3E41B7FDC0EA7EFD0D" box="[1233,1325,615,641]" country="Kyrgyzstan" name="Bishkek Shaary" pageId="17" pageNumber="18">Bishkek</collectingRegion>
(
<geoCoordinate id="E298506AFFFD9E3E405AFDC0EACFFD0D" box="[1340,1436,615,641]" degrees="42.88" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="42.88">42.88°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E45F1FD2CEFA5FD2A" box="[151,246,651,678]" degrees="74.56" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="74.56">74.56°E</geoCoordinate>
),
<date id="F312106DFFFD9E3E446BFD2CEED6FD29" box="[269,389,651,677]" pageId="17" pageNumber="18" value="1983-06-30">30.vi.1983</date>
(
<collectionCode id="E1BDAE68FFFD9E3E44F1FD2CEE80FD29" box="[407,467,651,677]" pageId="17" pageNumber="18">OLL</collectionCode>
: oll1059).—
<collectingCountry id="FFBB763DFFFD9E3E473BFD2CEC50FD2A" box="[605,771,651,678]" name="Tajikistan" pageId="17" pageNumber="18">TAJIKISTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFFD9E3E4677FD2CEC67FD29" box="[785,820,651,677]" count="1" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
, Veshab (
<geoCoordinate id="E298506AFFFD9E3E46CEFD2CEB5BFD29" box="[936,1032,651,677]" degrees="39.41" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="39.41">39.41°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E4174FD2CEB23FD29" box="[1042,1136,651,677]" degrees="68.93" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="68.93">68.93°E</geoCoordinate>
),
<date id="F312106DFFFD9E3E41EEFD2CEBBCFD29" box="[1160,1263,651,677]" pageId="17" pageNumber="18" value="1919-08">viii.1919</date>
, Glasunov leg. (
<collectionCode id="E1BDAE68FFFD9E3E45F9FD08EFB7FD45" box="[159,228,687,713]" pageId="17" pageNumber="18">CMK</collectionCode>
: ex coll. F. Morawitz, ms3535).—
<specimenCount id="91AAFD24FFFD9E3E4712FD09EDC3FD45" box="[628,656,686,713]" count="2" pageId="17" pageNumber="18" type="female">2♀</specimenCount>
, Seravschan, Veschab (
<geoCoordinate id="E298506AFFFD9E3E46FFFD08ECAAFD45" box="[921,1017,687,713]" degrees="39.41" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="39.41">39.41°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E4167FD08EB33FD45" box="[1025,1120,687,713]" degrees="68.93" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="68.93">68.93°E</geoCoordinate>
), 1892, Glasunov leg. (coll. F. Morawitz) (
<collectionCode id="E1BDAE68FFFD9E3E4427FD75EED2FD60" box="[321,385,722,748]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
).—
<specimenCount id="91AAFD24FFFD9E3E44CBFD75EEB6FD61" box="[429,485,722,749]" count="1" pageId="17" pageNumber="18" type="generic">1 ex.</specimenCount>
, Varzob district: Khodza-obi-garm,
<collectionCode id="E1BDAE68FFFD9E3E46EDFD75ECC8FD60" box="[907,923,722,748]" pageId="17" pageNumber="18">S</collectionCode>
slope of Gissar Range (
<geoCoordinate id="E298506AFFFD9E3E41D9FD75EA73FD61" box="[1215,1312,722,749]" degrees="38" direction="north" minutes="53" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="38.883335">38°53N</geoCoordinate>
68°
<geoCoordinate id="E298506AFFFD9E3E4032FD75EADCFD60" box="[1364,1423,722,748]" degrees="47" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55555" value="47.0">47°E</geoCoordinate>
),
<date id="F312106DFFFD9E3E45F1FD51EE42FC9C" box="[151,273,758,784]" pageId="17" pageNumber="18" value="1943-09-16">16.ix.1943</date>
, Nikolskaya leg. (
<collectionCode id="E1BDAE68FFFD9E3E4488FD51ED7EFC9C" box="[494,557,758,784]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
).—
<specimenCount id="91AAFD24FFFD9E3E473FFD51ED28FC9C" box="[601,635,758,784]" count="2" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
, Warsant [Varsant], 21.vii [1892 according to the collectors travel itinerary], Glas. [apparently stands for Glasunov], coll. F. Morawitz (
<collectionCode id="E1BDAE68FFFD9E3E46CBFCBDECBCFCB8" box="[941,1007,794,820]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
).—
<specimenCount id="91AAFD24FFFD9E3E417BFCBDEB6AFCB8" box="[1053,1081,794,820]" count="2" pageId="17" pageNumber="18" type="female">2♀</specimenCount>
, Obburdan (
<geoCoordinate id="E298506AFFFD9E3E41AEFCBDEA7BFCB8" box="[1224,1320,794,820]" degrees="40.44" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="40.44">40.44°N</geoCoordinate>
<geoCoordinate id="E298506AFFFD9E3E4057FCBDEAC3FCB8" box="[1329,1424,794,820]" degrees="69.30" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="69.3">69.30°E</geoCoordinate>
), 9.vii. (year?) (
<collectionCode id="E1BDAE68FFFD9E3E4453FC99EE23FCD4" box="[309,368,830,856]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
, also mentioned by
<bibRefCitation id="E33D4B5CFFFD9E3E472BFC9AEDAAFCD4" author="Morawitz, F. F." box="[589,761,829,856]" pageId="17" pageNumber="18" pagination="3 - 87" refId="ref30969" refString="Morawitz, F. F. (1894 [1893]) Supplement zur Bienenfauna Turkestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 28, 3 - 87." type="book chapter" year="1894">Morawitz 1894</bibRefCitation>
and
<bibRefCitation id="E33D4B5CFFFD9E3E4648FC9AECEBFCD4" author="Friese, H." box="[814,952,829,856]" pageId="17" pageNumber="18" refId="ref29768" refString="Friese, H. (1898) Die Bienen Europa's (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp." type="book" year="1898">Friese 1898</bibRefCitation>
).—
<specimenCount id="91AAFD24FFFD9E3E4680FC99EB50FCD4" box="[998,1027,830,856]" count="3" pageId="17" pageNumber="18" type="female">3♀</specimenCount>
, Kumar, 26.viii (year? See above for further material from this locality) (
<collectionCode id="E1BDAE68FFFD9E3E4740FCC6ED32FCF7" box="[550,609,865,891]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
,
<collectionCode id="E1BDAE68FFFD9E3E470AFCC6EDFEFCF7" box="[620,685,865,891]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="18" type="Museum">ZMB</collectionCode>
).
</paragraph>
<paragraph id="871336ADFFFD9E3E45A1FC22ECABFC4F" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1FC22EE2FFC13" box="[199,380,901,927]" italics="true" pageId="17" pageNumber="18">Other material:</emphasis>
MoA (1989) listed
<specimenCount id="91AAFD24FFFD9E3E470CFC21EC47FC13" box="[618,788,901,928]" count="2" pageId="17" pageNumber="18" type="generic">two specimens</specimenCount>
in the Mavromoustakis collection (cf. Mavromoustakis 1939). Additionally,
<specimenCount id="91AAFD24FFFD9E3E4419FC0EED4BFC4F" box="[383,536,937,963]" count="1" pageId="17" pageNumber="18" type="generic">one specimen</specimenCount>
each in
<collectionCode id="E1BDAE68FFFD9E3E4710FC0EED99FC4F" box="[630,714,937,963]" pageId="17" pageNumber="18">HNMB</collectionCode>
and
<collectionCode id="E1BDAE68FFFD9E3E4666FC0EEC6DFC4F" box="[768,830,937,963]" pageId="17" pageNumber="18">SDEI</collectionCode>
(not examined).
</paragraph>
<paragraph id="871336ADFFFD9E3E45A1FC6AEC69FC6A" blockId="17.[151,1437,151,2036]" box="[199,826,972,999]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1FC6AED58FC6B" box="[199,523,972,999]" italics="true" pageId="17" pageNumber="18">Genetic barcode information</emphasis>
. No information available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFD9E3E45A1FC56EE7CFBFA" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="871336ADFFFD9E3E45A1FC56EE7CFBFA" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1FC56EE6AFB86" box="[199,313,1009,1034]" italics="true" pageId="17" pageNumber="18">Diagnosis</emphasis>
. The female has a unique T6 with a lateral, slightly depressed wing-like projection and an undulated apical posterior margin. The male is characterized by laterally thick, impressed projections of T5 and T6 combined with a broad, semi-rectangular emargination of T7. Disc of T6 protruding. The male has specialized hairs on S3 and combs on S5.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFD9E3E45A1FB27EE29F878" pageId="17" pageNumber="18" type="description">
<paragraph id="871336ADFFFD9E3E45A1FB27EDE8FA72" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1FB27EE18FB15" box="[199,331,1152,1177]" italics="true" pageId="17" pageNumber="18">Description</emphasis>
.
<emphasis id="B5D8EABFFFFD9E3E443EFBD8EEFFFB15" box="[344,428,1151,1177]" italics="true" pageId="17" pageNumber="18">Female</emphasis>
(
<figureCitation id="1F972A28FFFD9E3E44DDFBD8ED5FFB15" box="[443,524,1151,1177]" captionStart="FIGURE 9" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1893]" captionTargetId="figure-19@16.[152,1436,181,1918]" captionTargetPageId="16" captionText="FIGURE 9. Pseudoanthidium obscuratum (Morawitz, 1875). a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan)." figureDoi="http://doi.org/10.5281/zenodo.14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="17" pageNumber="18">Fig. 9a</figureCitation>
).
<quantity id="40549B48FFFD9E3E4779FBD8ED2EFB16" box="[543,637,1151,1178]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="17" pageNumber="18" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
.—
<emphasis id="B5D8EABFFFFD9E3E47F8FBD8ED88FB15" box="[670,731,1151,1177]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
: Predominantly black, except for a small triangular spot behind the zenith of the eye; basal area of clypeus scattered punctate with polished interstices, lower half of clypeus densely punctate; apical ridge protruding, polished with mostly 56 tubercles (
<figureCitation id="1F972A28FFFD9E3E46C1FB60ECBCFB6D" box="[935,1007,1223,1249]" captionStart="FIGURE 8" captionStartId="15.[151,250,1461,1485]" captionTargetBox="[151,1436,911,1438]" captionTargetId="figure-380@15.[151,1436,911,1438]" captionTargetPageId="15" captionText="FIGURE 8. Pseudoanthidium moricei (Friese, 1911) stat. resurrect. a. Face of a female from Palestine (holotype); b. Face of a female from Kazakhstan. Note the different extension of light colouration." figureDoi="http://doi.org/10.5281/zenodo.14240753" httpUri="https://zenodo.org/record/14240753/files/figure.png" pageId="17" pageNumber="18">Figs 8</figureCitation>
,
<figureCitation id="1F972A28FFFD9E3E469DFB60EB4AFB6D" box="[1019,1049,1223,1249]" captionStart="FIGURE 9" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1893]" captionTargetId="figure-19@16.[152,1436,181,1918]" captionTargetPageId="16" captionText="FIGURE 9. Pseudoanthidium obscuratum (Morawitz, 1875). a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan)." figureDoi="http://doi.org/10.5281/zenodo.14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="17" pageNumber="18">9c</figureCitation>
,
<figureCitation id="1F972A28FFFD9E3E4142FB60EB17FB6D" box="[1060,1092,1223,1249]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="17" pageNumber="18">20</figureCitation>
); ridge anteriorly covered with dense fringe of short hairs; mandible robust, shining, black and chestnut-brown, with five teeth, separated by acute notches; head densely and finely punctate, with only supraclypeal area shining with scattered punctures; antenna black.—
<taxonomicName id="40AC4D2EFFFD9E3E4591FA94EE3CFAC0" box="[247,367,1331,1356]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFD9E3E4591FA94EE3CFAC0" box="[247,367,1331,1356]" italics="true" pageId="17" pageNumber="18">Mesosoma</emphasis>
</taxonomicName>
: Scutum, scutellum and axilla black; apical margin of scutellum acute-angled in profile, widely rounded in dorsal view; omaulus angulate; mesepisternum with long whitish hairs.—
<emphasis id="B5D8EABFFFFD9E3E411CFAF1EBBCFAE3" box="[1146,1263,1366,1391]" italics="true" pageId="17" pageNumber="18">Metasoma</emphasis>
: Black; lateral yellow spot on T1, mediolateral yellow spots on T3T5, often reduced in size; T6 black; lateral, slightly depressed wing-like projection and an undulated apical posterior margin with a shallow, light brown lamellate lobe (
<figureCitation id="1F972A28FFFD9E3E405DFA3AEADCFA3B" box="[1339,1423,1437,1463]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="17" pageNumber="18">Fig. 22</figureCitation>
); ventral scopa dark grey.—
<emphasis id="B5D8EABFFFFD9E3E44D9FA65EEA7FA57" box="[447,500,1474,1499]" italics="true" pageId="17" pageNumber="18">Legs</emphasis>
: Dark brown to black; pubescence white, with yellow bristles on inner face of hind basitarsus.—
<emphasis id="B5D8EABFFFFD9E3E444FFA42EE30FA72" box="[297,355,1509,1534]" italics="true" pageId="17" pageNumber="18">Wing</emphasis>
: Slightly brownish infuscated.
</paragraph>
<paragraph id="871336ADFFFD9E3E45A1F9AFEE29F878" blockId="17.[151,1437,151,2036]" pageId="17" pageNumber="18">
<emphasis id="B5D8EABFFFFD9E3E45A1F9AFEE52F9AE" box="[199,257,1544,1570]" italics="true" pageId="17" pageNumber="18">Male</emphasis>
(
<figureCitation id="1F972A28FFFD9E3E4475F9AFEE3BF9AE" box="[275,360,1544,1570]" captionStart="FIGURE 9" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1893]" captionTargetId="figure-19@16.[152,1436,181,1918]" captionTargetPageId="16" captionText="FIGURE 9. Pseudoanthidium obscuratum (Morawitz, 1875). a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan)." figureDoi="http://doi.org/10.5281/zenodo.14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="17" pageNumber="18">Fig. 9b</figureCitation>
).
<quantity id="40549B48FFFD9E3E4419F9AFEEBCF9AF" box="[383,495,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="17" pageNumber="18" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
.—
<emphasis id="B5D8EABFFFFD9E3E4769F9AFED1FF9AE" box="[527,588,1544,1570]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
: Black except for a small yellow, triangular spot behind the zenith of the eye; clypeus ivory-coloured, bell-shaped, anterior margin semicircularly curved inwards (
<figureCitation id="1F972A28FFFD9E3E41C6F98BEBA4F9CA" box="[1184,1271,1580,1606]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="17" pageNumber="18">Fig. 21</figureCitation>
); anterior half of clypeus smooth with scattered punctures, while posterior part with dense, fine punctation and long hair; apical margin black and crenulate; mandible short, black or chestnut-brown, with three strong black teeth; lower part of lower paraocular area ivory-coloured, not reaching upper end of clypeus and antennal socket; pubescence pale grey-yellow.—
<taxonomicName id="40AC4D2EFFFD9E3E4426F91BEEEBF959" box="[320,440,1724,1749]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFD9E3E4426F91BEEEBF959" box="[320,440,1724,1749]" italics="true" pageId="17" pageNumber="18">Mesosoma</emphasis>
</taxonomicName>
: Scutum black with regular, fine punctation; pronotal lobe dark brown and lamellate; scutellum, in profile, rounded medially and acute-angled laterally; outer margin of axilla not protruding beyond margin of scutellum.—
<emphasis id="B5D8EABFFFFD9E3E44C7F8A4ED45F890" box="[417,534,1795,1820]" italics="true" pageId="17" pageNumber="18">Metasoma</emphasis>
: Black; T1, sometimes also T2 and T3, with a small lateral yellow spot; (T2) T3-T5 with mediolateral yellow maculae; T5 and T6 with lateral projection, depressed from dorsal; T5 shining with scattered, irregular punctation; T6 mediolaterally bulging; T7 with median bulge and median rectangular emargination (broader than long) (
<figureCitation id="1F972A28FFFD9E3E4745F8CAEDCBF80B" box="[547,664,1901,1927]" captionStart="FIGURE 9" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1893]" captionTargetId="figure-19@16.[152,1436,181,1918]" captionTargetPageId="16" captionText="FIGURE 9. Pseudoanthidium obscuratum (Morawitz, 1875). a. Habitus dorsal, female from Uzbekistan; b. Habitus dorsal, male from Kyrgyzstan; c. Head of the female from Uzbekistan; de. Apical terga of the male (specimen from Kyrgyzstan)." figureDoi="http://doi.org/10.5281/zenodo.14240755" httpUri="https://zenodo.org/record/14240755/files/figure.png" pageId="17" pageNumber="18">Figs 9de</figureCitation>
,
<figureCitation id="1F972A28FFFD9E3E47C3F8C9ED90F804" box="[677,707,1902,1928]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="17" pageNumber="18">21</figureCitation>
,
<figureCitation id="1F972A28FFFD9E3E47B4F8C9EDA1F804" box="[722,754,1902,1928]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="17" pageNumber="18">22</figureCitation>
); lateral margin angulate; S3 with specialized undulate and apically hooked hairs; S5 with black basal comb, and strong black comb with approximately 25 digits apically at long arms.—
<emphasis id="B5D8EABFFFFD9E3E442EF812EEE1F843" box="[328,434,1973,1999]" italics="true" pageId="17" pageNumber="18">Genitalia</emphasis>
: Gonostyli paddle-shaped, tapering towards the apex; penis valve slender, curved in the upper part (
<figureCitation id="1F972A28FFFD9E3E447FF87EEE3EF878" box="[281,365,2009,2036]" captionStart="FIGURE 25" captionStartId="40.[151,250,2009,2033]" captionTargetBox="[176,1412,1240,1996]" captionTargetId="figure-103@40.[176,1412,1240,2001]" captionTargetPageId="40" captionText="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240785" httpUri="https://zenodo.org/record/14240785/files/figure.png" pageId="17" pageNumber="18">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFE9E3D45A1FF30EC0DFECD" pageId="18" pageNumber="19" type="distribution">
<paragraph id="871336ADFFFE9E3D45A1FF30EC0DFECD" blockId="18.[151,1437,150,321]" pageId="18" pageNumber="19">
<emphasis id="B5D8EABFFFFE9E3D45A1FF30EE03FF3D" box="[199,336,151,177]" italics="true" pageId="18" pageNumber="19">Distribution</emphasis>
. In addition to the localities listed under “material examined”, Murao
<emphasis id="B5D8EABFFFFE9E3D4137FF30EBDBFF3D" box="[1105,1160,151,177]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2015) found the species in
<collectingCountry id="FFBB763DFFFE9E3D45D2FF1CEE64FF59" box="[180,311,187,213]" name="Kazakhstan" pageId="18" pageNumber="19">Kazakhstan</collectingCountry>
in Aksu Valley (
<geoCoordinate id="E298506AFFFE9E3D4495FF1CED00FF59" box="[499,595,187,213]" degrees="44.78" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="44.78">44.78°N</geoCoordinate>
<geoCoordinate id="E298506AFFFE9E3D473DFF1CEDEBFF59" box="[603,696,187,213]" degrees="79.10" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="79.1">79.10°E</geoCoordinate>
) and Kashi-Kaindy, Jabagly (
<geoCoordinate id="E298506AFFFE9E3D4163FF1DEB36FF59" box="[1029,1125,186,213]" degrees="42.25" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="42.25">42.25°N</geoCoordinate>
<geoCoordinate id="E298506AFFFE9E3D410BFF1DEB9FFF59" box="[1133,1228,186,213]" degrees="70.25" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="70.25">70.25°E</geoCoordinate>
). Mavromoustakis (1939) reported material from Hissar Valley in
<collectingCountry id="FFBB763DFFFE9E3D47D3FF78EC70FF75" box="[693,803,223,249]" name="Tajikistan" pageId="18" pageNumber="19">Tajikistan</collectingCountry>
(Kvak,
<geoCoordinate id="E298506AFFFE9E3D46E4FF78ECB3FF75" box="[898,992,223,249]" degrees="38.80" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="38.8">38.80°N</geoCoordinate>
<geoCoordinate id="E298506AFFFE9E3D468FFF78EB1BFF75" box="[1001,1096,223,249]" degrees="68.76" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="68.76">68.76°E</geoCoordinate>
; Rujdasht,
<geoCoordinate id="E298506AFFFE9E3D41AFFF78EA7BFF75" box="[1225,1320,223,249]" degrees="38.84" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="38.84">38.84°N</geoCoordinate>
<geoCoordinate id="E298506AFFFE9E3D4057FF78EAC2FF75" box="[1329,1425,223,249]" degrees="68.83" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="68.83">68.83°E</geoCoordinate>
). For
<collectingCountry id="FFBB763DFFFE9E3D45A1FEA4EE1EFE91" box="[199,333,259,285]" name="Kazakhstan" pageId="18" pageNumber="19">Kazakhstan</collectingCountry>
, mentioned by Marikovskaya (1986) without details. Overall, the species was thus reported from
<collectingCountry id="FFBB763DFFFE9E3D45F1FE80EE4EFECD" box="[151,285,295,321]" name="Kazakhstan" pageId="18" pageNumber="19">Kazakhstan</collectingCountry>
,
<collectingCountry id="FFBB763DFFFE9E3D444EFE80EEF8FECD" box="[296,427,295,321]" name="Kyrgyzstan" pageId="18" pageNumber="19">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="FFBB763DFFFE9E3D44D0FE80ED64FECD" box="[438,567,295,321]" name="Uzbekistan" pageId="18" pageNumber="19">Uzbekistan</collectingCountry>
, and
<collectingCountry id="FFBB763DFFFE9E3D4717FE80ED8CFECD" box="[625,735,295,321]" name="Tajikistan" pageId="18" pageNumber="19">Tajikistan</collectingCountry>
(
<figureCitation id="1F972A28FFFE9E3D4789FE80EC02FECD" box="[751,849,295,321]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="18" pageNumber="19">Fig. 26b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,313 @@
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<mods:number id="0AB94A85C2F644192DF2FF918FE1A3CD">1</mods:number>
</mods:detail>
<mods:extent id="D6DD6411224BB01736064DFBAE63E780" unit="page">
<mods:start id="5810C6AF33DC3B7345D74DEEC107EDF8">1</mods:start>
<mods:end id="088F7E1593E49AD6B60EB8EC52EAF213">50</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="BFAC3771D66C9C9DA264D80EA127F775">
<mods:url id="6F810B504C9705534EA5223F40D32F64">http://dx.doi.org/10.11646/zootaxa.5541.1.1</mods:url>
</mods:location>
<mods:classification id="625440A61193336363A5339D9F9D3C0B">journal article</mods:classification>
<mods:identifier id="CEF92F88908F6968C60DEA960C199494" type="DOI">10.11646/zootaxa.5541.1.1</mods:identifier>
<mods:identifier id="7AB6F84EC643769D5C2BA9ACD7E468C9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A673DA2179F01548D329F7067321C72E" type="Zenodo-Dep">14240734</mods:identifier>
<mods:identifier id="B44992418AEEBF335760F526160E74C9" type="ZooBank">A5D2633B-36A8-42A4-AD21-22041804BE95</mods:identifier>
</mods:mods>
<treatment id="0F0587BBFFE59E2545F1FB00EAD4F875" LSID="urn:lsid:plazi:treatment:0F0587BBFFE59E2545F1FB00EAD4F875" httpUri="http://treatment.plazi.org/id/0F0587BBFFE59E2545F1FB00EAD4F875" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="CFB66526FFE59E2645F1FB00EB3DFB4E" box="[151,1134,1191,1218]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="871336ADFFE59E2645F1FB00EB3DFB4E" blockId="9.[151,1134,1191,1255]" box="[151,1134,1191,1218]" pageId="9" pageNumber="10">
<heading id="DC5B81C1FFE59E2645F1FB00EB3DFB4E" bold="true" box="[151,1134,1191,1218]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B5D8EABFFFE59E2645F1FB00EB3DFB4E" bold="true" box="[151,1134,1191,1218]" pageId="9" pageNumber="10">
<taxonomicName id="40AC4D2EFFE59E2645F1FB00EC50FB4E" ID-CoL="8SQBH" authority="(Mavromoustakis, 1954)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[151,771,1191,1218]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE59E2645F1FB00EE9FFB4E" bold="true" box="[151,460,1191,1218]" italics="true" pageId="9" pageNumber="10">Pseudoanthidium ivanovi</emphasis>
(
<bibRefCitation id="E33D4B5CFFE59E2644BDFB0FEDA8FB4E" author="Mavromoustakis, G. A." box="[475,763,1192,1218]" pageId="9" pageNumber="10" pagination="249 - 252" refId="ref30628" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726" type="journal article" year="1954">Mavromoustakis, 1954</bibRefCitation>
)
</taxonomicName>
stat. resurrect. &amp; comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645F1FB6BEE10FB6A" box="[151,323,1228,1255]" pageId="9" pageNumber="10" type="description">
<paragraph id="871336ADFFE59E2645F1FB6BEE10FB6A" blockId="9.[151,1134,1191,1255]" box="[151,323,1228,1255]" pageId="9" pageNumber="10">
(
<figureCitation id="1F972A28FFE59E2645F9FB6BEFBBFB6B" box="[159,232,1228,1255]" captionStart="FIGURE 5" captionStartId="10.[151,250,1621,1645]" captionTargetBox="[151,1435,669,1597]" captionTargetId="figure-257@10.[151,1435,669,1597]" captionTargetPageId="10" captionText="FIGURE 5. Pseudoanthidium ivanovi (Mavromoustakis, 1954) stat. resurrect. Female from Afghanistan. a. Habitus dorsal; b. Apical terga; c. Mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.14240745" httpUri="https://zenodo.org/record/14240745/files/figure.png" pageId="9" pageNumber="10">Figs 5</figureCitation>
,
<figureCitation id="1F972A28FFE59E264592FB6BEE40FB6A" box="[244,275,1228,1254]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="9" pageNumber="10">20</figureCitation>
,
<figureCitation id="1F972A28FFE59E264478FB6BEE68FB6A" box="[286,315,1228,1254]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="9" pageNumber="10">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645F1FAB7EA26FA7D" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="871336ADFFE59E2645F1FAB7EC70FAA4" blockId="9.[151,1437,1296,1413]" box="[151,803,1296,1320]" pageId="9" pageNumber="10">
<taxonomicName id="40AC4D2EFFE59E2645F1FAB7ED11FAA4" ID-CoL="8SQBH" authority="Mavromoustakis, 1954" authorityName="Mavromoustakis" authorityYear="1954" box="[151,578,1296,1320]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE59E2645F1FAB7EE01FAA4" box="[151,338,1296,1320]" italics="true" pageId="9" pageNumber="10">Anthidium ivanovi</emphasis>
<bibRefCitation id="E33D4B5CFFE59E26443EFAB7ED11FAA4" author="Mavromoustakis, G. A." box="[344,578,1296,1320]" pageId="9" pageNumber="10" pagination="249 - 252" refId="ref30628" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726" type="journal article" year="1954">Mavromoustakis, 1954</bibRefCitation>
</taxonomicName>
.—Female,
<collectingCountry id="FFBB763DFFE59E2647DEFAB7EC4CFAA4" box="[696,799,1296,1320]" name="Tajikistan" pageId="9" pageNumber="10">Tajikistan</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFE59E2645F1FA88ECB6FACB" blockId="9.[151,1437,1296,1413]" box="[151,997,1327,1351]" pageId="9" pageNumber="10">
<taxonomicName id="40AC4D2EFFE59E2645F1FA88EC4BFACB" ID-CoL="8SQBH" authority="(Mavromoustakis, 1954)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[151,792,1327,1351]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ivanovi" subGenus="Pseudoanthidium">
<emphasis id="B5D8EABFFFE59E2645F1FA88ED48FACB" box="[151,539,1327,1351]" italics="true" pageId="9" pageNumber="10">Anthidium (Pseudoanthidium) ivanovi</emphasis>
(
<bibRefCitation id="E33D4B5CFFE59E264741FA88EC42FACB" author="Mavromoustakis, G. A." box="[551,785,1327,1351]" pageId="9" pageNumber="10" pagination="249 - 252" refId="ref30628" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726" type="journal article" year="1954">Mavromoustakis, 1954</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFE59E264653FA88ECB2FACB" author="Warncke, K." box="[821,993,1327,1351]" pageId="9" pageNumber="10" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFE59E2645F1FAE9EC1FFA09" blockId="9.[151,1437,1296,1413]" pageId="9" pageNumber="10">
non
<taxonomicName id="40AC4D2EFFE59E2645A4FAE9EDF6FAEA" ID-CoL="4NP4N" authority="(Morawitz, 1875)" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[194,677,1358,1382]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE59E2645A4FAE9EEA3FAEA" box="[194,496,1358,1382]" italics="true" pageId="9" pageNumber="10">Pseudoanthidium obscuratum</emphasis>
(
<bibRefCitation id="E33D4B5CFFE59E26449DFAE9EDCCFAEA" author="Morawitz, F. F." box="[507,671,1358,1382]" pageId="9" pageNumber="10" pagination="1 - 160" refId="ref30848" refString="Morawitz, F. F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1 - 160." type="journal article" year="1875">Morawitz, 1875</bibRefCitation>
)
</taxonomicName>
.—
<taxonomicName id="40AC4D2EFFE59E2647A3FAE9EC78FAE9" ID-CoL="8SQBH" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[709,811,1358,1381]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE59E2647A3FAE9EC78FAE9" box="[709,811,1358,1381]" italics="true" pageId="9" pageNumber="10">A. ivanovi</emphasis>
</taxonomicName>
is regarded as a junior synonym of
<taxonomicName id="40AC4D2EFFE59E2641EFFAE8EA4AFAEA" ID-CoL="4NP4N" baseAuthorityName="Morawitz" baseAuthorityYear="1875" box="[1161,1305,1358,1382]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="obscuratum">
<emphasis id="B5D8EABFFFE59E2641EFFAE8EA4AFAEA" box="[1161,1305,1358,1382]" italics="true" pageId="9" pageNumber="10">P. obscuratum</emphasis>
</taxonomicName>
in secondary sources such as by ITIS (2023) and
<bibRefCitation id="E33D4B5CFFE59E264754FACBEC14FA09" author="Ascher, J. S. &amp; Pickering, J." box="[562,839,1388,1413]" pageId="9" pageNumber="10" refId="ref29495" refString="Ascher, J. S. &amp; Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: http: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species (accessed 1 August 20024)" type="url" year="2024">Ascher &amp; Pickering (2024)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFE59E2645F1FA14EA26FA7D" blockId="9.[151,1437,1459,2025]" pageId="9" pageNumber="10">
The species was not included in Pasteels (1969) revision of the genera of Old World Anthidiini.
<bibRefCitation id="E33D4B5CFFE59E264180FA14EACFFA41" author="Warncke, K." box="[1254,1436,1459,1485]" pageId="9" pageNumber="10" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
listed the species but did not include it in his consecutive numbering of species, nor did he include it in his key.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645A1FA5CEE37F945" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="871336ADFFE59E2645A1FA5CEB07F9B5" blockId="9.[151,1437,1459,2025]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1FA5CEEC8F999" box="[199,411,1531,1557]" italics="true" pageId="9" pageNumber="10">Material examined</emphasis>
:
<materialsCitation id="37C43CF0FFE59E2644CEFA5CEB03F9B5" collectingDate="2011-08-04" collectionCode="CMK" collectorName="O. Pak" country="Afghanistan" elevation="3850" latitude="34.6" location="Hajigak Pass" longLatPrecision="719" longitude="68.05" municipality="Kohi-Bana Mountains" pageId="9" pageNumber="10" specimenCount="1" specimenCount-female="1" stateProvince="Bamiyan">
<collectingCountry id="FFBB763DFFE59E2644CEFA5CED22F999" box="[424,625,1531,1557]" name="Afghanistan" pageId="9" pageNumber="10">AFGHANISTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFE59E26471DFA5CEDCDF999" box="[635,670,1531,1557]" count="1" pageId="9" pageNumber="10" type="female">1♀</specimenCount>
,
<collectingRegion id="4568F84FFFE59E2647CAFA5CEC45F999" box="[684,790,1531,1557]" country="Afghanistan" name="Bamyan" pageId="9" pageNumber="10">Bamiyan</collectingRegion>
:
<locationDeviation id="96A12F8FFFE59E264679FA5CECA8F999" box="[799,1019,1531,1557]" pageId="9" pageNumber="10">
<quantity id="40549B48FFE59E264679FA5CEC35F999" box="[799,870,1531,1557]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="km" value="30.0">30 km</quantity>
SE Bamiyan
</locationDeviation>
,
<collectingMunicipality id="6777ACD7FFE59E264160FA5CEA51F999" box="[1030,1282,1531,1557]" pageId="9" pageNumber="10">Kohi-Bana Mountains</collectingMunicipality>
, near
<location id="82736076FFE59E264022FA5CEF9AF9B5" LSID="urn:lsid:plazi:treatment:0F0587BBFFE59E2545F1FB00EAD4F875:82736076FFE59E264022FA5CEF9AF9B5" country="Afghanistan" latitude="34.6" longLatPrecision="719" longitude="68.05" municipality="Kohi-Bana Mountains" name="Hajigak Pass" pageId="9" pageNumber="10" stateProvince="Bamiyan">Hajigak Pass</location>
(c.
<geoCoordinate id="E298506AFFE59E264591F9B8EE05F9B5" box="[247,342,1567,1593]" degrees="34.60" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="555" value="34.6">34.60°N</geoCoordinate>
<geoCoordinate id="E298506AFFE59E26443BF9B9EEE8F9B5" box="[349,443,1566,1593]" degrees="68.05" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="555" value="68.05">68.05°E</geoCoordinate>
),
<quantity id="40549B48FFE59E2644A9F9B8ED3FF9B5" box="[463,620,1567,1593]" metricMagnitude="3" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="9" pageNumber="10" unit="m" value="3850.0" valueMax="4000.0" valueMin="3700.0">
<elevation id="0C81D19EFFE59E2644A9F9B8ED3FF9B5" box="[463,620,1567,1593]" metricMagnitude="3" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="9" pageNumber="10" unit="m" value="3850.0" valueMax="4000.0" valueMin="3700.0">37004000 m</elevation>
</quantity>
,
<date id="F312106DFFE59E26471EF9B8EC52F9B5" box="[632,769,1567,1593]" pageId="9" pageNumber="10" value="2011-08-04">
<collectingDate id="E356E985FFE59E26471EF9B8EC52F9B5" box="[632,769,1567,1593]" pageId="9" pageNumber="10" value="2011-08-04">04.viii.2011</collectingDate>
</date>
,
<collectorName id="2A59537BFFE59E26466AF9B8EC0AF9B5" box="[780,857,1567,1593]" pageId="9" pageNumber="10">O. Pak</collectorName>
leg. (
<collectionCode id="E1BDAE68FFE59E2646FEF9B8EC8EF9B5" box="[920,989,1567,1593]" pageId="9" pageNumber="10">CMK</collectionCode>
: mk1537)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFE59E2645A1F9E4ED31F90D" blockId="9.[151,1437,1459,2025]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1F9E4EE22F9D1" box="[199,369,1603,1629]" italics="true" pageId="9" pageNumber="10">
<typeStatus id="5817880FFFE59E2645A1F9E4EE22F9D1" box="[199,369,1603,1629]" isOtherTypeStatus="true" pageId="9" pageNumber="10">Other material</typeStatus>
</emphasis>
. The female
<typeStatus id="5817880FFFE59E26476CF9E4ED3FF9D1" box="[522,620,1603,1629]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is deposited in the
<location id="82736076FFE59E264629F9E4EB41F9D1" LSID="urn:lsid:plazi:treatment:0F0587BBFFE59E2545F1FB00EAD4F875:82736076FFE59E264629F9E4EB41F9D1" box="[847,1042,1603,1629]" country="Cyprus" municipality="It" name="Mavromoustakis'" pageId="9" pageNumber="10" stateProvince="Nicosia">Mavromoustakis</location>
collection,
<collectingRegion id="4568F84FFFE59E2641FCF9E4EBA1F9D1" box="[1178,1266,1603,1629]" country="Cyprus" name="Nicosia" pageId="9" pageNumber="10">Nicosia</collectingRegion>
,
<collectingCountry id="FFBB763DFFE59E264199F9E4EA03F9D1" box="[1279,1360,1603,1629]" name="Cyprus" pageId="9" pageNumber="10">Cyprus</collectingCountry>
(MoA 1989).
<collectingMunicipality id="6777ACD7FFE59E264581F9C0EFABF90D" box="[231,248,1639,1665]" pageId="9" pageNumber="10">It</collectingMunicipality>
was not available for this study.
</paragraph>
<paragraph id="871336ADFFE59E2645A1F92CEE37F945" blockId="9.[151,1437,1459,2025]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1F92CED42F929" box="[199,529,1675,1701]" italics="true" pageId="9" pageNumber="10">Genetic barcode information.</emphasis>
Genetic barcode of specimen mk1537 available (
<collectionCode id="E1BDAE68FFE59E264150F92CEBCCF929" box="[1078,1183,1675,1701]" pageId="9" pageNumber="10">ABABQ</collectionCode>
177-24). A BIN has not yet been assigned.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645A1F974EE31F8B9" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="871336ADFFE59E2645A1F974EE31F8B9" blockId="9.[151,1437,1459,2025]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1F974EECEF961" box="[199,413,1747,1773]" italics="true" pageId="9" pageNumber="10">Diagnosis (female)</emphasis>
. Characterized by a narrow, shining flange on each side of the middle of T6 combined with impunctate, polished areas around the ocelli; apical ridge of the clypeus shining, protruding slightly above disc, with shallow tubercles.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2545A1F898EA18FD09" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="871336ADFFE59E2545A1F898EA18FD09" blockId="9.[151,1437,1459,2025]" lastBlockId="10.[151,1436,151,645]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1F898EE18F8D4" box="[199,331,1855,1880]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
.
<emphasis id="B5D8EABFFFE59E26443FF898EEFEF8D5" box="[345,429,1855,1881]" italics="true" pageId="9" pageNumber="10">Female</emphasis>
.
<quantity id="40549B48FFE59E2644DDF898EEA8F8D5" box="[443,507,1855,1881]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="10" unit="mm" value="7.0">7 mm</quantity>
long.—
<emphasis id="B5D8EABFFFE59E264730F898EDC0F8D5" box="[598,659,1855,1881]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
: Black, except for a yellow triangular postocular maculation; clypeus densely punctate, finer towards apical margin; basal area (anterior margin) polished and impunctate, very narrowly at sides and somewhat broadly in the middle (
<figureCitation id="1F972A28FFE59E2647FBF820EDA1F82D" box="[669,754,1927,1953]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="9" pageNumber="10">Fig. 20</figureCitation>
); subapical area transversely depressed, very finely rugosely punctured; apical ridge thickened with some shallow denticles, polished; mandible yellow, but reddish brown towards the margins; with four (possibly five) low, black teeth; supraclypeal area densely punctate except for apical margin; vertex densely punctate except for impunctate, polished area around ocelli; postocular yellow stripe (
<typeStatus id="5817880FFFE69E254076FF30EA26FF3D" box="[1296,1397,151,177]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
) or triangular yellow spot (
<collectingCountry id="FFBB763DFFE69E2544F3FF1CED73FF59" box="[405,544,187,213]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
specimen); vertex length 2.25 x the diameter of the posterior ocellus; face around antennal sockets and gena with long silvery hair.—
<taxonomicName id="40AC4D2EFFE69E2547ADFF78EC10FF74" box="[715,835,223,248]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE69E2547ADFF78EC10FF74" box="[715,835,223,248]" italics="true" pageId="10" pageNumber="11">Mesosoma</emphasis>
</taxonomicName>
: Scutum densely punctate (
<figureCitation id="1F972A28FFE69E254112FF78EB97FF75" box="[1140,1220,223,249]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="10" pageNumber="11">Fig. 4c</figureCitation>
), slightly finer than on vertex; pronotal lobe with high lamella; outer margin of scutellum and axilla widely rounded in dorsal view; apical margin with a very broadly interrupted yellow stripe (
<typeStatus id="5817880FFFE69E254650FE80ECC8FECD" box="[822,923,295,321]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
) or entirely black (
<collectingCountry id="FFBB763DFFE69E254108FE80EBAAFECD" box="[1134,1273,295,321]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
specimen); no axillar suture visible; mesepisternum with long white hair; omaulus widely rounded.—
<emphasis id="B5D8EABFFFE69E25411CFEECEBBCFEE8" box="[1146,1263,331,356]" italics="true" pageId="10" pageNumber="11">Metasoma</emphasis>
: Punctation on T1T5 fine, T1 with impunctate apical margin; punctation on T2T5 practically reaches outer margin; punctation on T6 irregular, somewhat rugulose; T1 with broad yellow stripe broadly interrupted in the middle and attenuated inwards (
<typeStatus id="5817880FFFE69E254599FE10EE35FE5D" box="[255,358,439,465]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
; only short lateral stripe in the
<collectingCountry id="FFBB763DFFE69E2547A6FE10EC1AFE5D" box="[704,841,439,465]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
specimen); T2 with yellow band almost reaching the middle; T3 with yellow band on each side, subcontiguous in the middle; T4T5 with yellow band, notched in the middle; T6 black with median, transverse, interrupted yellow stripe (
<typeStatus id="5817880FFFE69E2546D6FE58EB45FD95" box="[944,1046,511,537]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
, entirely black in the
<collectingCountry id="FFBB763DFFE69E254075FE58EACFFD95" box="[1299,1436,511,537]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
specimen); T6 trapezoid, with a shallow lamellate flange on each side of the middle (
<figureCitation id="1F972A28FFE69E254133FD84EBFEFDB1" box="[1109,1197,547,573]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="10" pageNumber="11">Figs 4b</figureCitation>
,
<figureCitation id="1F972A28FFE69E2541DFFD84EB8AFDB1" box="[1209,1241,547,573]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="10" pageNumber="11">22</figureCitation>
); outer margin of S6 rounded; scopa in the middle blackish brown, lighter at margins.—
<emphasis id="B5D8EABFFFE69E2546C1FDE0EC8FFDEC" box="[935,988,583,608]" italics="true" pageId="10" pageNumber="11">Legs</emphasis>
: Femora black with some yellowish red apically; tibiae bright yellow; tarsi reddish yellow; hind basitarsi with dense light golden hair on inner face.
</paragraph>
</subSubSection>
<caption id="D3D36625FFE69E2545F1F9F2ED10F91D" ID-DOI="http://doi.org/10.5281/zenodo.14240745" ID-Zenodo-Dep="14240745" httpUri="https://zenodo.org/record/14240745/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1621,1645]" targetBox="[151,1435,669,1597]" targetPageId="10" targetType="figure">
<paragraph id="871336ADFFE69E2545F1F9F2ED10F91D" blockId="10.[151,1437,1621,1682]" pageId="10" pageNumber="11">
<emphasis id="B5D8EABFFFE69E2545F1F9F2EE41F9E1" bold="true" box="[151,274,1621,1645]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
<taxonomicName id="40AC4D2EFFE69E25447FF9F2EC49F9E1" authority="(Mavromoustakis, 1954)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[281,794,1621,1645]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE69E25447FF9F2ED48F9E1" box="[281,539,1621,1645]" italics="true" pageId="10" pageNumber="11">Pseudoanthidium ivanovi</emphasis>
(
<bibRefCitation id="E33D4B5CFFE69E254741F9F2EC40F9E1" author="Mavromoustakis, G. A." box="[551,787,1621,1645]" pageId="10" pageNumber="11" pagination="249 - 252" refId="ref30628" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726" type="journal article" year="1954">Mavromoustakis, 1954</bibRefCitation>
)
</taxonomicName>
<emphasis id="B5D8EABFFFE69E254647F9F2ECEEF9E1" bold="true" box="[801,957,1621,1645]" pageId="10" pageNumber="11">stat. resurrect.</emphasis>
Female from Afghanistan. a. Habitus dorsal; b. Apical terga; c.
<taxonomicName id="40AC4D2EFFE69E25445FF9DDEEF4F91D" box="[313,423,1658,1681]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="33" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
in lateral view.
</paragraph>
</caption>
<subSubSection id="CFB66526FFE69E2545A1F967EEC7F8E5" pageId="10" pageNumber="11" type="discussion">
<paragraph id="871336ADFFE69E2545A1F967EBE9F8CA" blockId="10.[151,1437,1727,2042]" pageId="10" pageNumber="11">
<emphasis id="B5D8EABFFFE69E2545A1F967EFAFF955" box="[199,252,1728,1753]" italics="true" pageId="10" pageNumber="11">Note</emphasis>
:
<bibRefCitation id="E33D4B5CFFE69E25446FF918EEEFF955" author="Warncke, K." box="[265,444,1727,1753]" pageId="10" pageNumber="11" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
mentioned that
<taxonomicName id="40AC4D2EFFE69E25470BF967ED8AF955" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[621,729,1728,1753]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE69E25470BF967ED8AF955" box="[621,729,1728,1753]" italics="true" pageId="10" pageNumber="11">P. ivanovi</emphasis>
</taxonomicName>
has only half of the vertex length of
<emphasis id="B5D8EABFFFE69E25410DF967EA5FF955" box="[1131,1292,1727,1753]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="40AC4D2EFFE69E25410DF967EA5BF955" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[1131,1288,1727,1753]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="melanurum">P. melanurum</taxonomicName>
.
</emphasis>
Specifically, the vertex length is 2.25 x the diameter of the posterior ocellus, while it is 3.3 x the diameter of the posterior ocellus in
<emphasis id="B5D8EABFFFE69E2545DEF8AFEE0DF8AD" box="[184,350,1799,1825]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="40AC4D2EFFE69E2545DEF8AFEE09F8AD" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[184,346,1799,1825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="melanurum">P. melanurum</taxonomicName>
.
</emphasis>
However, it is unlikely that
<bibRefCitation id="E33D4B5CFFE69E2547D4F8A0EC39F8AD" author="Warncke, K." box="[690,874,1799,1825]" pageId="10" pageNumber="11" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
examined the
<typeStatus id="5817880FFFE69E25417CF8AFEB19F8AE" box="[1050,1098,1800,1826]" pageId="10" pageNumber="11">type</typeStatus>
material of
<taxonomicName id="40AC4D2EFFE69E2541BBF8AFEA1DF8AD" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[1245,1358,1800,1825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ivanovi">
<emphasis id="B5D8EABFFFE69E2541BBF8AFEA1DF8AD" box="[1245,1358,1800,1825]" italics="true" pageId="10" pageNumber="11">P. ivanovi</emphasis>
</taxonomicName>
in the Mavromoustakis collection, and therefore it is not clear what Warnckes statement is based on.
</paragraph>
<paragraph id="871336ADFFE69E2545A1F8E8EEC7F8E5" blockId="10.[151,1437,1727,2042]" box="[199,404,1871,1898]" pageId="10" pageNumber="11">
<emphasis id="B5D8EABFFFE69E2545A1F8E8EE52F8E5" box="[199,257,1871,1897]" italics="true" pageId="10" pageNumber="11">Male</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE69E2545A1F8D4EAD4F875" pageId="10" pageNumber="11" type="distribution">
<paragraph id="871336ADFFE69E2545A1F8D4EAD4F875" blockId="10.[151,1437,1727,2042]" pageId="10" pageNumber="11">
<emphasis id="B5D8EABFFFE69E2545A1F8D4EE1EF801" box="[199,333,1907,1933]" italics="true" pageId="10" pageNumber="11">Distribution</emphasis>
.
<collectingCountry id="FFBB763DFFE69E25443DF8D4EEB2F801" box="[347,481,1907,1933]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
and
<collectingCountry id="FFBB763DFFE69E25477FF8D4EDD6F801" box="[537,645,1907,1933]" name="Tajikistan" pageId="10" pageNumber="11">Tajikistan</collectingCountry>
(
<figureCitation id="1F972A28FFE69E2547F3F8D4EDA4F801" box="[661,759,1907,1933]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="10" pageNumber="11">Fig. 26b</figureCitation>
). The
<typeStatus id="5817880FFFE69E254658F8D4ECCDF801" box="[830,926,1907,1933]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
was collected at Khorog (Chorog),
<collectingCountry id="FFBB763DFFE69E254056F8D4EACFF801" box="[1328,1436,1907,1933]" name="Tajikistan" pageId="10" pageNumber="11">Tajikistan</collectingCountry>
(
<geoCoordinate id="E298506AFFE69E2545F9F83EEFACF838" box="[159,255,1945,1972]" degrees="37" direction="north" minutes="30" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="37.5">37°30N</geoCoordinate>
,
<geoCoordinate id="E298506AFFE69E25446BF83EEE3AF838" box="[269,361,1945,1972]" degrees="71" direction="east" minutes="33" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="71.55">71°33E</geoCoordinate>
), at
<quantity id="40549B48FFE69E2544FBF830EEAAF83E" box="[413,505,1943,1970]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="11" unit="m" value="2200.0">2,200 m</quantity>
above sea level in the Pamir Mountains at the border with
<collectingCountry id="FFBB763DFFE69E2541C2F830EA7DF83D" box="[1188,1326,1943,1969]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
. The new material was collected at
<quantity id="40549B48FFE69E2544CDF81CED12F85A" box="[427,577,1979,2006]" metricMagnitude="3" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="10" pageNumber="11" unit="m" value="3850.0" valueMax="4000.0" valueMin="3700.0">37004000 m</quantity>
above sea level in the Baba Mountains (Kōh-i Bābā or Kūh-e Bābā), a western extension of the Hindu Kush,
<collectingCountry id="FFBB763DFFE69E2544B9F878ED3AF875" box="[479,617,2015,2041]" name="Afghanistan" pageId="10" pageNumber="11">Afghanistan</collectingCountry>
. It is the highest place where
<taxonomicName id="40AC4D2EFFE69E2546CBF878EB3EF875" authorityName="Friese" authorityYear="1898" box="[941,1133,2015,2041]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE69E2546CBF878EB3EF875" box="[941,1133,2015,2041]" italics="true" pageId="10" pageNumber="11">Pseudoanthidium</emphasis>
</taxonomicName>
has been collected so far.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,313 @@
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<paragraph id="871336ADFFE79E2445F1FEC8ED2FFE05" blockId="11.[151,636,367,430]" box="[151,636,367,394]" pageId="11" pageNumber="12">
<heading id="DC5B81C1FFE79E2445F1FEC8ED2FFE05" bold="true" box="[151,636,367,394]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B5D8EABFFFE79E2445F1FEC8ED2FFE05" bold="true" box="[151,636,367,394]" pageId="11" pageNumber="12">
<taxonomicName id="40AC4D2EFFE79E2445F1FEC8ED44FE06" authorityName="Kasparek" authorityYear="2024" box="[151,535,367,394]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="microrubrum" status="sp. nov.">
<emphasis id="B5D8EABFFFE79E2445F1FEC8ED44FE06" bold="true" box="[151,535,367,394]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium microrubrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFE79E244778FEC8ED2FFE05" box="[542,636,367,393]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445F1FE34EEB8FE22" box="[151,491,403,430]" pageId="11" pageNumber="12" type="description">
<paragraph id="871336ADFFE79E2445F1FE34EEB8FE22" blockId="11.[151,636,367,430]" box="[151,491,403,430]" pageId="11" pageNumber="12">
(
<figureCitation id="1F972A28FFE79E2445F9FE34EFBBFE21" box="[159,232,403,429]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Figs 6</figureCitation>
,
<figureCitation id="1F972A28FFE79E244592FE34EE40FE21" box="[244,275,403,429]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="11" pageNumber="12">20</figureCitation>
,
<figureCitation id="1F972A28FFE79E244478FE34EE6FFE21" box="[286,316,403,429]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="11" pageNumber="12">21</figureCitation>
,
<figureCitation id="1F972A28FFE79E24442EFE34EE34FE21" box="[328,359,403,429]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="11" pageNumber="12">22</figureCitation>
,
<figureCitation id="1F972A28FFE79E244414FE34EEC3FE21" box="[370,400,403,429]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="11" pageNumber="12">23</figureCitation>
,
<figureCitation id="1F972A28FFE79E2444FAFE34EEE9FE21" box="[412,442,403,429]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="11" pageNumber="12">24</figureCitation>
, 25)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445F1FE7CEC9BFD42" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="871336ADFFE79E2445F1FE7CEEA9FDED" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445F1FE7CEFAAFE79" box="[151,249,475,501]" italics="true" pageId="11" pageNumber="12">Material</emphasis>
.
<materialsCitation id="37C43CF0FFE79E244461FE7CECFAFD95" collectionCode="OLL" collectorName="T. Wood" country="Morocco" elevation="1750" latitude="33.244" location="Fes-Meknes" longLatPrecision="6" longitude="4.6762" municipality="Boulemane" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="5817880FFFE79E244461FE7CEE20FE79" box="[263,371,475,501]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
.
<specimenCount id="91AAFD24FFE79E24441BFE7BEEC6FE7A" box="[381,405,476,502]" count="1" pageId="11" pageNumber="12" type="male"></specimenCount>
.
<collectingCountry id="FFBB763DFFE79E2444C6FE7CED61FE79" box="[416,562,475,501]" name="Morocco" pageId="11" pageNumber="12">MOROCCO</collectingCountry>
:
<location id="82736076FFE79E24475AFE7CED9BFE79" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E24475AFE7CED9BFE79" box="[572,712,475,501]" country="Morocco" latitude="33.244" longLatPrecision="6" longitude="4.6762" municipality="Boulemane" name="Fes-Meknes" pageId="11" pageNumber="12">Fès-Meknès</location>
,
<location id="82736076FFE79E2447B4FE7CEC07FE79" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E2447B4FE7CEC07FE79" box="[722,852,475,501]" country="Morocco" latitude="33.244" longLatPrecision="6" longitude="4.6762" municipality="Boulemane" name="Boulemane" pageId="11" pageNumber="12">Boulemane</location>
,
<location id="82736076FFE79E244639FE7CECCDFE7A" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E244639FE7CECCDFE7A" box="[863,926,475,502]" country="Morocco" latitude="33.244" longLatPrecision="6" longitude="4.6762" municipality="Boulemane" name="R 503" pageId="11" pageNumber="12">R503</location>
,
<location id="82736076FFE79E2446CCFE7CEBDBFE7A" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E2446CCFE7CEBDBFE7A" box="[938,1160,475,502]" country="Morocco" latitude="33.244" longLatPrecision="6" longitude="4.6762" municipality="Boulemane" name="SE of Ait Karmosse" pageId="11" pageNumber="12">SE of Ait Karmosse</location>
(
<geoCoordinate id="E298506AFFE79E2441F0FE7CEA40FE79" box="[1174,1299,475,501]" degrees="33.2440" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="33.244">33.2440°N</geoCoordinate>
-
<geoCoordinate id="E298506AFFE79E244047FE7CEAC3FE79" box="[1313,1424,475,501]" degrees="4.6762" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="4.6762">4.6762°E</geoCoordinate>
),
<quantity id="40549B48FFE79E2445F1FE58EFBCFD96" box="[151,239,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="12" unit="m" value="1750.0">
<elevation id="0C81D19EFFE79E2445F1FE58EFBCFD96" box="[151,239,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="12" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<collectorName id="2A59537BFFE79E24459AFDA7EE0FFD95" box="[252,348,511,538]" pageId="11" pageNumber="12">T. Wood</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFE79E24472CFE58EDAAFD95" box="[586,761,511,537]" pageId="11" pageNumber="12">ABABZ002-23</accessionNumber>
(
<collectionCode id="E1BDAE68FFE79E24466FFE58EC16FD95" box="[777,837,511,537]" pageId="11" pageNumber="12">OLL</collectionCode>
: tjw293)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFE79E2446ACFE58EEA5FDED" collectingDate="2022-05-22" collectionCode="OLL" collectorName="T. Wood" country="Morocco" elevation="1900" latitude="33.2966" location="Fes-Meknes" longLatPrecision="6" longitude="4.6896" municipality="Boulemane" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5817880FFFE79E2446ACFE58EB62FD95" box="[970,1073,511,537]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
:
<specimenCount id="91AAFD24FFE79E24415BFE58EB33FD95" box="[1085,1120,511,537]" count="1" pageId="11" pageNumber="12" type="female">1♀</specimenCount>
,
<collectingCountry id="FFBB763DFFE79E244109FE58EA52FD95" box="[1135,1281,511,537]" name="Morocco" pageId="11" pageNumber="12">MOROCCO</collectingCountry>
:
<location id="82736076FFE79E24406AFE58EACBFD95" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E24406AFE58EACBFD95" box="[1292,1432,511,537]" country="Morocco" latitude="33.2966" longLatPrecision="6" longitude="4.6896" municipality="Boulemane" name="Fes-Meknes" pageId="11" pageNumber="12">Fès-Meknès</location>
,
<location id="82736076FFE79E2445F1FD84EE4AFDB1" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E2445F1FD84EE4AFDB1" box="[151,281,547,573]" country="Morocco" latitude="33.2966" longLatPrecision="6" longitude="4.6896" municipality="Boulemane" name="Boulemane" pageId="11" pageNumber="12">Boulemane</location>
,
<location id="82736076FFE79E244443FD84EE37FDB2" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E244443FD84EE37FDB2" box="[293,356,547,574]" country="Morocco" latitude="33.2966" longLatPrecision="6" longitude="4.6896" municipality="Boulemane" name="R 503" pageId="11" pageNumber="12">R503</location>
,
<locationDeviation id="96A12F8FFFE79E244416FD83ED09FDB1" box="[368,602,547,574]" location="Boulemane" pageId="11" pageNumber="12">
<quantity id="40549B48FFE79E244416FD83EEF9FDB1" box="[368,426,547,574]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="km" value="7.0">7 km</quantity>
SE Boulemane
</locationDeviation>
(
<geoCoordinate id="E298506AFFE79E24470EFD84EDB6FDB1" box="[616,741,547,573]" degrees="33.2966" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="33.2966">33.2966°N</geoCoordinate>
-
<geoCoordinate id="E298506AFFE79E244792FD84EC30FDB1" box="[756,867,547,573]" degrees="4.6896" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="4.6896">4.6896°E</geoCoordinate>
),
<quantity id="40549B48FFE79E244611FD84EC9DFDB2" box="[887,974,547,574]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="m" value="1900.0">
<elevation id="0C81D19EFFE79E244611FD84EC9DFDB2" box="[887,974,547,574]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="F312106DFFE79E2446BCFD84EB18FDB1" box="[986,1099,547,573]" pageId="11" pageNumber="12" value="2022-05-22">
<collectingDate id="E356E985FFE79E2446BCFD84EB18FDB1" box="[986,1099,547,573]" pageId="11" pageNumber="12" value="2022-05-22">22.v.2022</collectingDate>
</date>
,
<collectorName id="2A59537BFFE79E244131FD83EBE5FDB1" box="[1111,1206,547,574]" pageId="11" pageNumber="12">T. Wood</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFE79E2445F1FDE0EE15FDED" box="[151,326,583,609]" pageId="11" pageNumber="12">ABABZ003-23</accessionNumber>
(
<collectionCode id="E1BDAE68FFE79E244432FDE0EEC3FDED" box="[340,400,583,609]" pageId="11" pageNumber="12">OLL</collectionCode>
: tjw297)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFE79E2445A1FDCBEC9BFD42" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1FDCBED40FD09" box="[199,531,619,646]" italics="true" pageId="11" pageNumber="12">Genetic barcode information</emphasis>
.
<collectingMunicipality id="6777ACD7FFE79E244743FDCCED02FD09" box="[549,593,619,645]" pageId="11" pageNumber="12">The</collectingMunicipality>
genetic barcode of the
<typeStatus id="5817880FFFE79E24460CFDCCEC9FFD09" box="[874,972,619,645]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(tjw293) and the
<typeStatus id="5817880FFFE79E2441C5FDCBEA50FD0A" box="[1187,1283,620,646]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
(tjw297) are available (
<specimenCode id="D70A9ED6FFE79E24446AFD28EEEDFD25" box="[268,446,655,681]" collectionCode="ABABZ" pageId="11" pageNumber="12">ABABZ002-23</specimenCode>
,
<specimenCode id="D70A9ED6FFE79E2444AFFD28ED0AFD25" box="[457,601,655,681]" pageId="11" pageNumber="12">ABABZ003</specimenCode>
-23).
<location id="82736076FFE79E2447E9FD28EDE8FD25" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E2447E9FD28EDE8FD25" box="[655,699,655,681]" municipality="The" name="The" pageId="11" pageNumber="12">The</location>
BIN is BOLD:
<specimenCode id="D70A9ED6FFE79E24460EFD28EC88FD25" box="[872,987,655,681]" pageId="11" pageNumber="12">AFC0423</specimenCode>
.
<location id="82736076FFE79E244680FD28EB3FFD25" LSID="urn:lsid:plazi:treatment:0F0587BBFFE79E2445F1FEC8EAC2F851:82736076FFE79E244680FD28EB3FFD25" box="[998,1132,655,681]" municipality="The" name="Information" pageId="11" pageNumber="12">Information</location>
on the nearest neighbours, barcode gap, and mean genetic distances are provided under
<taxonomicName id="40AC4D2EFFE79E244626FD13EC92FD41" authorityName="Kasparek" authorityYear="2024" box="[832,961,691,717]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFE79E244626FD13EC92FD41" box="[832,961,691,717]" italics="true" pageId="11" pageNumber="12">P. farsiense</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445A1FD7FEBEAFBB9" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="871336ADFFE79E2445A1FD7FEBEAFBB9" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1FD7FEE6AFD7D" box="[199,313,728,753]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
.A small
<taxonomicName id="40AC4D2EFFE79E2444FEFD70ED0FFD7D" authorityName="Friese" authorityYear="1898" box="[408,604,727,753]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE79E2444FEFD70ED0FFD7D" box="[408,604,727,753]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium</emphasis>
</taxonomicName>
with orange maculation.The female is characterized by a strongly protruding apical ridge of the clypeus, combined with a triangular T6 featuring apical emargination, a fine punctation of scutum and terga, a dark brown scopa, and its diminutive size. Other
<taxonomicName id="40AC4D2EFFE79E244626FCB8EB0FFCB5" box="[832,1116,799,825]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E244626FCB8EB57FCB5" box="[832,1028,799,825]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium</emphasis>
species
</taxonomicName>
with red or orange are much larger (
<taxonomicName id="40AC4D2EFFE79E24458EFCE3EE6DFCD1" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[232,318,836,861]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE79E24458EFCE3EE6DFCD1" box="[232,318,836,861]" italics="true" pageId="11" pageNumber="12">P. royoi</emphasis>
</taxonomicName>
,
<quantity id="40549B48FFE79E24442AFCE4EE94FCD1" box="[332,455,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="11" pageNumber="12" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
) or possess distinctive morphological differences, such as a flat clypeus (
<taxonomicName id="40AC4D2EFFE79E244199FCE3EAC6FCD1" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[1279,1429,836,861]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="canariense">
<emphasis id="B5D8EABFFFE79E244199FCE3EAC6FCD1" box="[1279,1429,836,861]" italics="true" pageId="11" pageNumber="12">P. canariense</emphasis>
</taxonomicName>
,
<taxonomicName id="40AC4D2EFFE79E2445F1FCCFEE53FC0D" authorityName="Vereecken &amp; Litman" authorityYear="2023" box="[151,256,871,897]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jacobii">
<emphasis id="B5D8EABFFFE79E2445F1FCCFEE53FC0D" box="[151,256,871,897]" italics="true" pageId="11" pageNumber="12">P. jacobii</emphasis>
</taxonomicName>
), a bilobed apex of T6 (
<taxonomicName id="40AC4D2EFFE79E24476AFCCFED31FC0D" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[524,610,872,897]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE79E24476AFCCFED31FC0D" box="[524,610,872,897]" italics="true" pageId="11" pageNumber="12">P. royoi</emphasis>
</taxonomicName>
), or coarse punctation of scutum and terga (
<taxonomicName id="40AC4D2EFFE79E24412BFCCFEBB7FC0D" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[1101,1252,871,897]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFE79E24412BFCCFEBB7FC0D" box="[1101,1252,871,897]" italics="true" pageId="11" pageNumber="12">P. puncticolle</emphasis>
</taxonomicName>
). In comparison with other orange or red-brown
<taxonomicName id="40AC4D2EFFE79E24449BFC2CEC48FC29" box="[509,795,907,933]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E24449BFC2CED92FC29" box="[509,705,907,933]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium</emphasis>
species
</taxonomicName>
, the male of this species is distinguished from
<taxonomicName id="40AC4D2EFFE79E244042FC2BEA2AFC29" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[1316,1401,908,933]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE79E244042FC2BEA2AFC29" box="[1316,1401,908,933]" italics="true" pageId="11" pageNumber="12">P. royoi</emphasis>
</taxonomicName>
by its small size (
<quantity id="40549B48FFE79E24445FFC08EE28FC46" box="[313,379,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
vs.
<quantity id="40549B48FFE79E2444CCFC08ED5DFC45" box="[426,526,943,970]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="12" unit="mm" value="11.5">11.5 mm</quantity>
) and a straight apical margin of T6 (protruding with a broad apical emargination in
<taxonomicName id="40AC4D2EFFE79E2445D5FC73EE5BFC61" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[179,264,980,1005]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE79E2445D5FC73EE5BFC61" box="[179,264,980,1005]" italics="true" pageId="11" pageNumber="12">P. royoi</emphasis>
</taxonomicName>
). Additionally, the lower face of this species is orange-red (black in
<emphasis id="B5D8EABFFFE79E244166FC73EBCCFC61" box="[1024,1183,980,1005]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="40AC4D2EFFE79E244166FC73EBCBFC61" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[1024,1176,980,1005]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="canariense">P. canariense</taxonomicName>
;
</emphasis>
dark yellow-orange in
<taxonomicName id="40AC4D2EFFE79E2445F1FC5FEE53FB9D" authorityName="Vereecken &amp; Litman" authorityYear="2023" box="[151,256,1015,1041]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jacobii">
<emphasis id="B5D8EABFFFE79E2445F1FC5FEE53FB9D" box="[151,256,1015,1041]" italics="true" pageId="11" pageNumber="12">P. jacobii</emphasis>
</taxonomicName>
). Regarding
<taxonomicName id="40AC4D2EFFE79E2444F4FC5FED7AFB9D" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[402,553,1015,1041]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFE79E2444F4FC5FED7AFB9D" box="[402,553,1015,1041]" italics="true" pageId="11" pageNumber="12">P. puncticolle</emphasis>
</taxonomicName>
, where females often exhibit red-brown maculation, the maculation of males is typically yellow, and they have a much coarser tergal punctation compared to
<emphasis id="B5D8EABFFFE79E244167FBBBEBEAFBB9" box="[1025,1209,1051,1077]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="40AC4D2EFFE79E244167FBBBEBE6FBB9" authorityName="Kasparek" authorityYear="2024" box="[1025,1205,1051,1077]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="microrubrum">P. microrubrum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445A1FBE7EBB5F8FE" pageId="11" pageNumber="12" type="description">
<paragraph id="871336ADFFE79E2445A1FBE7ED62F985" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1FBE7EE18FBD5" box="[199,331,1088,1113]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
.
<emphasis id="B5D8EABFFFE79E244439FB98EEE0FBD5" box="[351,435,1087,1113]" italics="true" pageId="11" pageNumber="12">Female</emphasis>
(
<figureCitation id="1F972A28FFE79E2444A1FB98ED4DFBD5" box="[455,542,1087,1113]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Fig. 6a</figureCitation>
).
<quantity id="40549B48FFE79E24475EFBE7EDD3FBD6" box="[568,640,1088,1114]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
.—
<emphasis id="B5D8EABFFFE79E2447C7FB98ED8DFBD5" box="[673,734,1087,1113]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: Clypeus orange, with basal margin arched inwards; finely punctate with a basal impunctate area; subapical area transversely depressed, very finely rugosely punctate; apical ridge thickened with some erect denticles; ridge dark red-brown, and polished (
<figureCitation id="1F972A28FFE79E244163FB20EB04FB2D" box="[1029,1111,1159,1185]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Fig. 6d</figureCitation>
); mandible orange, gradually becoming brown towards the teeth; five black teeth, to1 being the strongest, with the others almost equally-sized; teeth separated by acute incisions; lower paraocular area with large orange spot not reaching the antennal socket; broad orange preoccipital band reaching the middle of the eye; hairs on head scattered, dark grey to black.—
<taxonomicName id="40AC4D2EFFE79E2445F1FABFEE5CFABD" box="[151,271,1304,1329]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE79E2445F1FABFEE5CFABD" box="[151,271,1304,1329]" italics="true" pageId="11" pageNumber="12">Mesosoma</emphasis>
</taxonomicName>
: Scutum with dense, shallow punctation and shining interstices; scutellum rounded in profile, little overhanging propodeum; orange; pronotal lobe lamellate; omaulus rounded; pubescence of mesosoma short, dark grey.—
<emphasis id="B5D8EABFFFE79E24458CFAC7EE0CFAF5" box="[234,351,1376,1401]" italics="true" pageId="11" pageNumber="12">Metasoma</emphasis>
: T1 entirely red except for a diffuse black area in the middle (
<figureCitation id="1F972A28FFE79E244176FAF8EB32FAF5" box="[1040,1121,1375,1401]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Fig. 6c</figureCitation>
); T2T6 orange, interrupted by a median black area; T6, in dorsal view, a wide-angular triangle with a median projection with a small apical emargination to accommodate the sting; S1S5 orange, S6 dark grey, triangular; scopal hairs dark brown.—
<emphasis id="B5D8EABFFFE79E244006FA0FEAC6FA4D" box="[1376,1429,1448,1473]" italics="true" pageId="11" pageNumber="12">Legs</emphasis>
: Trochanteres and basal areas of femora black, the rest orange; orange bristles on inner face of hind basitarsus.—
<emphasis id="B5D8EABFFFE79E2445F1FA57EF82F985" box="[151,209,1520,1545]" italics="true" pageId="11" pageNumber="12">Wing</emphasis>
: Moderately brown infuscated.
</paragraph>
<paragraph id="871336ADFFE79E2445A1F9B4EBB5F8FE" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1F9B4EE52F9A1" box="[199,257,1555,1581]" italics="true" pageId="11" pageNumber="12">Male</emphasis>
(
<figureCitation id="1F972A28FFE79E244474F9B4EE34F9A1" box="[274,359,1555,1581]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Fig. 6b</figureCitation>
).
<quantity id="40549B48FFE79E24441BF9B3EE92F9A2" box="[381,449,1556,1582]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
.—
<emphasis id="B5D8EABFFFE79E244484F9B4ED4CF9A1" box="[482,543,1555,1581]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: Black; lower part of paraocular area dirty yellow; minute yellow spot behind the eye; dense, long white pubescence on lower face; mandible reddish-brown with three strong teeth.—
<taxonomicName id="40AC4D2EFFE79E24407BF99FEAC6F9DD" box="[1309,1429,1592,1617]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE79E24407BF99FEAC6F9DD" box="[1309,1429,1592,1617]" italics="true" pageId="11" pageNumber="12">Mesosoma</emphasis>
</taxonomicName>
: Black with dense punctation; pronotal lobe small, black with shallow lamella, hidden by dense pubescence; scutellum and axillae crescent-shaped in dorsal view, slightly convexly bulging; in profile, rounded medially and acute angular laterally; tegula reddish-brown.—
<emphasis id="B5D8EABFFFE79E24477EF903EDDEF931" box="[536,653,1700,1725]" italics="true" pageId="11" pageNumber="12">Metasoma</emphasis>
: T1T6 black with broad lateral orange-brown bands; median space between the bands larger in anterior terga than in posterior terga; depression in area next to the orange-brown bands also orange-brown with a bronze-coloured touch; T7 brownish-yellow, emarginate at apical margin; S1, S2 and base of S3 with felt-like cover of plumose, branched hairs; S3 with a fan-like fringe of corkscrew-like curved and apically hooked hairs; S5 with a submedian pair of black combs and another pair of larger black combs on lateral arms (
<figureCitation id="1F972A28FFE79E244009F894EFE1F8FD" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-19@12.[152,1436,181,1918]" captionTargetPageId="12" captionText="FIGURE 6. Pseudoanthidium microrubrum Kasparek sp. nov. a. Habitus dorsal, female from Morocco; b. Habitus dorsal, male, from Morocco; c. Habitus of the female in oblique view (Morocco); d. Face of the female (Morocco); e. Metasoma of the male in dorsal view (note the presence of specialized hairs and combs) (Morocco)." figureDoi="http://doi.org/10.5281/zenodo.14240749" httpUri="https://zenodo.org/record/14240749/files/figure.png" pageId="11" pageNumber="12">Fig. 6e</figureCitation>
).—
<emphasis id="B5D8EABFFFE79E2445B8F8F0EE1BF8FD" box="[222,328,1879,1905]" italics="true" pageId="11" pageNumber="12">Genitalia</emphasis>
: Gonostyli paddle-shaped; penis valves slender, unfused, but touching at the apex.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445A1F8DBEAC2F851" pageId="11" pageNumber="12" type="etymology">
<paragraph id="871336ADFFE79E2445A1F8DBEDD8F835" blockId="11.[151,1437,475,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1F8DBEEC1F819" box="[199,402,1916,1941]" italics="true" pageId="11" pageNumber="12">Derivatio nominis</emphasis>
. Name combined from “micro” small, and “rubrum” red. The name should indicate that this is the smallest red
<taxonomicName id="40AC4D2EFFE79E24440EF838EDD8F835" box="[360,651,1951,1977]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B5D8EABFFFE79E24440EF838ED7FF835" box="[360,556,1951,1977]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="871336ADFFE79E2445A1F864EAC2F851" blockId="11.[151,1437,475,2014]" box="[199,1425,1987,2014]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445A1F864EE03F851" box="[199,336,1987,2013]" italics="true" pageId="11" pageNumber="12">Distribution</emphasis>
: Only known from the province of Boulemane in the Région Fès Meknès of
<collectingCountry id="FFBB763DFFE79E2441C9F863EA47F852" box="[1199,1300,1988,2014]" name="Morocco" pageId="11" pageNumber="12">Morocco</collectingCountry>
(
<figureCitation id="1F972A28FFE79E244042F864EAD7F851" box="[1316,1412,1987,2013]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="11" pageNumber="12">Fig. 26a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,76 @@
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<subSubSection id="CFB66526FFE79E2445F1FF30EDF7FF3D" box="[151,676,151,178]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="871336ADFFE79E2445F1FF30EDF7FF3D" blockId="11.[151,676,151,213]" box="[151,676,151,178]" pageId="11" pageNumber="12">
<heading id="DC5B81C1FFE79E2445F1FF30EDF7FF3D" bold="true" box="[151,676,151,178]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="40AC4D2EFFE79E2445F1FF30EDF7FF3D" ID-CoL="4NP4L" authority="(Klug, 1832)" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[151,676,151,178]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFE79E2445F1FF30EDF7FF3D" bold="true" box="[151,676,151,178]" pageId="11" pageNumber="12">
<emphasis id="B5D8EABFFFE79E2445F1FF30ED51FF3E" bold="true" box="[151,514,151,178]" italics="true" pageId="11" pageNumber="12">Pseudoanthidium melanurum</emphasis>
(Klug, 1832)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE79E2445F1FF1CEACBFE91" pageId="11" pageNumber="12" type="description">
<paragraph id="871336ADFFE79E2445F1FF1CEEF6FF59" blockId="11.[151,676,151,213]" box="[151,421,187,213]" pageId="11" pageNumber="12">
(
<figureCitation id="1F972A28FFE79E2445F9FF1CEFA4FF59" box="[159,247,187,213]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="11" pageNumber="12">Figs 20</figureCitation>
,
<figureCitation id="1F972A28FFE79E244464FF1CEE73FF59" box="[258,288,187,213]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="11" pageNumber="12">21</figureCitation>
,
<figureCitation id="1F972A28FFE79E24444AFF1CEE18FF59" box="[300,331,187,213]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="11" pageNumber="12">22</figureCitation>
,
<figureCitation id="1F972A28FFE79E244430FF1CEE27FF59" box="[342,372,187,213]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="11" pageNumber="12">23</figureCitation>
,
<figureCitation id="1F972A28FFE79E2444E6FF1CEECEFF59" box="[384,413,187,213]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="11" pageNumber="12">24</figureCitation>
)
</paragraph>
<paragraph id="871336ADFFE79E2445F1FEA4EACBFE91" blockId="11.[151,1432,259,286]" box="[151,1432,259,286]" pageId="11" pageNumber="12">This species shows a conspicuous variation across its distribution area. A detailed study will be published separately.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,328 @@
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<mods:title id="6170E9B5AD7FE14E557635E8F0D350E5">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<treatment id="0F0587BBFFE89E2B45F1FF30EB79F8F0" LSID="urn:lsid:plazi:treatment:0F0587BBFFE89E2B45F1FF30EB79F8F0" httpUri="http://treatment.plazi.org/id/0F0587BBFFE89E2B45F1FF30EB79F8F0" lastPageNumber="5" pageId="4" pageNumber="5">
<subSubSection id="CFB66526FFE89E2B45F1FF30EC41FF3D" box="[151,786,151,178]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="871336ADFFE89E2B45F1FF30EC41FF3D" blockId="4.[151,786,151,214]" box="[151,786,151,178]" pageId="4" pageNumber="5">
<heading id="DC5B81C1FFE89E2B45F1FF30EC41FF3D" bold="true" box="[151,786,151,178]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="40AC4D2EFFE89E2B45F1FF30EC41FF3D" ID-CoL="4NP45" authority="(Mavromoustakis, 1948)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1948" box="[151,786,151,178]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bytinskii">
<emphasis id="B5D8EABFFFE89E2B45F1FF30EC41FF3D" bold="true" box="[151,786,151,178]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45F1FF30EE88FF3E" bold="true" box="[151,475,151,178]" italics="true" pageId="4" pageNumber="5">Pseudoanthidium bytinskii</emphasis>
(
<bibRefCitation id="E33D4B5CFFE89E2B448CFF30EC59FF3D" author="Mavromoustakis, G. A." box="[490,778,151,177]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis, 1948</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE89E2B45F1FF1CEE10FF5A" box="[151,323,187,214]" pageId="4" pageNumber="5" type="description">
<paragraph id="871336ADFFE89E2B45F1FF1CEE10FF5A" blockId="4.[151,786,151,214]" box="[151,323,187,214]" pageId="4" pageNumber="5">
(
<figureCitation id="1F972A28FFE89E2B45F9FF1CEFBBFF59" box="[159,232,187,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-15@5.[151,1435,181,1881]" captionTargetPageId="5" captionText="FIGURE 2. Pseudoanthidium bytinskii (Mavromoustakis, 1948), form Israel. a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female, with bulging mandibles; d. Mandible of the male with depressed basal part; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240738" httpUri="https://zenodo.org/record/14240738/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
,
<figureCitation id="1F972A28FFE89E2B4592FF1CEE6FFF5A" box="[244,316,187,214]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStart-3="FIGURE 23" captionStart-4="FIGURE 24" captionStart-5="FIGURE 25" captionStartId-0="35.[151,250,1943,1967]" captionStartId-1="37.[151,250,1943,1967]" captionStartId-2="38.[151,250,1491,1515]" captionStartId-3="39.[151,250,1918,1942]" captionStartId-4="40.[151,250,1170,1194]" captionStartId-5="40.[151,250,2009,2033]" captionTargetBox-0="[180,1408,181,1912]" captionTargetBox-1="[179,1408,181,1908]" captionTargetBox-2="[409,1188,1275,1417]" captionTargetBox-3="[173,1415,561,1882]" captionTargetBox-4="[307,1281,311,1152]" captionTargetBox-5="[176,1412,1240,1996]" captionTargetId-0="figure-15@35.[180,1408,181,1919]" captionTargetId-1="figure-15@37.[179,1408,181,1919]" captionTargetId-3="figure-169@39.[173,1415,561,1895]" captionTargetId-4="graphics-80@40.[648,939,711,796]" captionTargetId-5="figure-103@40.[176,1412,1240,2001]" captionTargetPageId-0="35" captionTargetPageId-1="37" captionTargetPageId-2="38" captionTargetPageId-3="39" captionTargetPageId-4="40" captionTargetPageId-5="40" captionText-0="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." captionText-1="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." captionText-2="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." captionText-3="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." captionText-4="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." captionText-5="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi-0="http://doi.org/10.5281/zenodo.14240775" figureDoi-1="http://doi.org/10.5281/zenodo.14240779" figureDoi-2="http://doi.org/10.5281/zenodo.14240781" figureDoi-3="http://doi.org/10.5281/zenodo.14240783" figureDoi-4="http://doi.org/10.5281/zenodo.14275912" figureDoi-5="http://doi.org/10.5281/zenodo.14240785" httpUri-0="https://zenodo.org/record/14240775/files/figure.png" httpUri-1="https://zenodo.org/record/14240779/files/figure.png" httpUri-2="https://zenodo.org/record/14240781/files/figure.png" httpUri-3="https://zenodo.org/record/14240783/files/figure.png" httpUri-4="https://zenodo.org/record/14275912/files/figure.png" httpUri-5="https://zenodo.org/record/14240785/files/figure.png" pageId="4" pageNumber="5">2025</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE89E2B45F1FEA7EC3AFEBA" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="871336ADFFE89E2B45F1FEA7EC47FE94" blockId="4.[151,873,256,310]" box="[151,788,256,280]" pageId="4" pageNumber="5">
<taxonomicName id="40AC4D2EFFE89E2B45F1FEA7ED0FFE94" ID-CoL="8SRRC" authority="(Mavromoustakis, 1948)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1948" box="[151,604,256,280]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bytinskii">
<emphasis id="B5D8EABFFFE89E2B45F1FEA7EE0EFE94" box="[151,349,256,280]" italics="true" pageId="4" pageNumber="5">Anthidium bytinskii</emphasis>
(
<bibRefCitation id="E33D4B5CFFE89E2B440CFEA7ED05FE94" author="Mavromoustakis, G. A." box="[362,598,256,280]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis, 1948</bibRefCitation>
)
</taxonomicName>
.—Female,
<collectingCountry id="FFBB763DFFE89E2B47B3FEA7EC42FE94" box="[725,785,256,280]" name="Israel" pageId="4" pageNumber="5">Israel</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFE89E2B45F1FEB9EC3AFEBA" blockId="4.[151,873,256,310]" box="[151,873,286,310]" pageId="4" pageNumber="5">
<taxonomicName id="40AC4D2EFFE89E2B45F1FEB9EC3AFEBA" ID-CoL="4NP45" authority="(Mavromoustakis 1948)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1948" box="[151,873,286,310]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bytinskii" subGenus="Paraanthidiellum">
<emphasis id="B5D8EABFFFE89E2B45F1FEB9ED3CFEBA" box="[151,623,286,310]" italics="true" pageId="4" pageNumber="5">Pseudoanthidium (Paraanthidiellum) bytinskii</emphasis>
(
<bibRefCitation id="E33D4B5CFFE89E2B471AFEB9EC31FEBA" author="Mavromoustakis, G. A." box="[636,866,286,310]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis 1948</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE89E2B45F1FEC0EB79F8F0" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="871336ADFFE89E2B45F1FEC0EC6BFD9A" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45F1FEC0EE21FE0D" box="[151,370,359,385]" italics="true" pageId="4" pageNumber="5">Material examined.</emphasis>
<materialsCitation id="37C43CF0FFE89E2B441EFEC0EA04FE0D" box="[376,1367,359,386]" collectingDate="1988-04-07" collectionCode="CMK" collectorName="K. Guichard" country="Israel" latitude="31.45" location="Ein Gedi" longLatPrecision="729" longitude="35.38" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1">
<collectingCountry id="FFBB763DFFE89E2B441EFEC0EE8DFE0D" box="[376,478,359,385]" name="Israel" pageId="4" pageNumber="5">ISRAEL</collectingCountry>
:
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,
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,
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,
<date id="F312106DFFE89E2B4633FEC0EC9EFE0D" box="[853,973,359,385]" pageId="4" pageNumber="5" value="1988-04-07">
<collectingDate id="E356E985FFE89E2B4633FEC0EC9EFE0D" box="[853,973,359,385]" pageId="4" pageNumber="5" value="1988-04-07">07.iv.1988</collectingDate>
</date>
,
<collectorName id="2A59537BFFE89E2B46BEFECFEB32FE0D" box="[984,1121,359,386]" pageId="4" pageNumber="5">K. Guichard</collectorName>
leg. (
<collectionCode id="E1BDAE68FFE89E2B41F8FEC0EBB0FE0D" box="[1182,1251,359,385]" pageId="4" pageNumber="5">CMK</collectionCode>
: ms2707)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFE89E2B401DFEC0EE31FE40" collectingDate="2018-04-20" collectionCode="OLL" collectorName="M. Halada" country="Israel" latitude="30.13" location="Menuha Junction" longLatPrecision="734" longitude="35.13" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1">
<specimenCount id="91AAFD24FFE89E2B401DFEC0EACBFE0D" box="[1403,1432,359,385]" count="1" pageId="4" pageNumber="5" type="male">1♂</specimenCount>
,
<locationDeviation id="96A12F8FFFE89E2B45F1FE2BEE6CFE2A" box="[151,319,396,423]" location="Eilat" pageId="4" pageNumber="5">
<quantity id="40549B48FFE89E2B45F1FE2BEFB6FE2A" box="[151,229,396,422]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="km" value="100.0">100km</quantity>
N Eilat
</locationDeviation>
,
<location id="82736076FFE89E2B442CFE2BED5DFE2A" LSID="urn:lsid:plazi:treatment:0F0587BBFFE89E2B45F1FF30EB79F8F0:82736076FFE89E2B442CFE2BED5DFE2A" box="[330,526,396,422]" country="Israel" latitude="30.13" longLatPrecision="734" longitude="35.13" name="Menuha Junction" pageId="4" pageNumber="5">Menuha Junction</location>
, R90,
<geoCoordinate id="E298506AFFE89E2B4733FE2AEDE7FE2B" box="[597,692,397,423]" degrees="30.13" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="30.13">30.13°N</geoCoordinate>
<geoCoordinate id="E298506AFFE89E2B47DDFE2BEC4BFE2B" box="[699,792,396,423]" degrees="35.13" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="35.13">35.13°E</geoCoordinate>
,
<date id="F312106DFFE89E2B4642FE2BECCEFE2A" box="[804,925,396,422]" pageId="4" pageNumber="5" value="2018-04-20">
<collectingDate id="E356E985FFE89E2B4642FE2BECCEFE2A" box="[804,925,396,422]" pageId="4" pageNumber="5" value="2018-04-20">20.iv.2018</collectingDate>
</date>
,
<collectorName id="2A59537BFFE89E2B46CEFE2AEB4EFE2A" box="[936,1053,396,423]" pageId="4" pageNumber="5">M. Halada</collectorName>
leg.; COI sequence: ABABX468- 22 (
<collectionCode id="E1BDAE68FFE89E2B45A4FE15EFADFE40" box="[194,254,434,460]" pageId="4" pageNumber="5">OLL</collectionCode>
: oll836)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFE89E2B44EEFE15EE32FE7D" collectingDate="2019-05-04" collectionCode="OLL" collectorName="M. Halada" country="Israel" latitude="30.16" location="Yahel" longLatPrecision="734" longitude="35.15" pageId="4" pageNumber="5" specimenCount="1" specimenCount-female="1">
<specimenCount id="91AAFD24FFE89E2B44EEFE15EEF2FE40" box="[392,417,434,460]" count="1" pageId="4" pageNumber="5" type="female">1♀</specimenCount>
,
<quantity id="40549B48FFE89E2B44CAFE15EEA1FE40" box="[428,498,434,460]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="km" value="10.0">10 km</quantity>
N
<location id="82736076FFE89E2B4774FE15ED06FE40" LSID="urn:lsid:plazi:treatment:0F0587BBFFE89E2B45F1FF30EB79F8F0:82736076FFE89E2B4774FE15ED06FE40" box="[530,597,434,460]" country="Israel" latitude="30.16" longLatPrecision="734" longitude="35.15" name="Yahel" pageId="4" pageNumber="5">Yahel</location>
,
<geoCoordinate id="E298506AFFE89E2B4706FE15EDECFE40" box="[608,703,434,460]" degrees="30.16" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="30.16">30.16°N</geoCoordinate>
<geoCoordinate id="E298506AFFE89E2B47A3FE15EC70FE41" box="[709,803,434,461]" degrees="35.15" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="35.15">35.15°E</geoCoordinate>
,
<date id="F312106DFFE89E2B4648FE15ECCDFE40" box="[814,926,434,460]" pageId="4" pageNumber="5" value="2019-05-04">
<collectingDate id="E356E985FFE89E2B4648FE15ECCDFE40" box="[814,926,434,460]" pageId="4" pageNumber="5" value="2019-05-04">04.v.2019</collectingDate>
</date>
,
<collectorName id="2A59537BFFE89E2B46CFFE15EB4CFE40" box="[937,1055,434,460]" pageId="4" pageNumber="5">M. Halada</collectorName>
leg.; COI sequence: ABABX469- 22 (
<collectionCode id="E1BDAE68FFE89E2B45A0FE70EE51FE7D" box="[198,258,471,497]" pageId="4" pageNumber="5">OLL</collectionCode>
: oll837)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFE89E2B44E5FE70EC67FD9A" collectingDate="2016-03-28" collectionCode="CMK" collectorName="J. Monks" country="Oman" elevation="270" latitude="23.3" location="Al Hajar Mts." longLatPrecision="754" longitude="57.1" municipality="Wadi Al-Arabein" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1">
<collectingCountry id="FFBB763DFFE89E2B44E5FE70EE8CFE7D" box="[387,479,471,497]" name="Oman" pageId="4" pageNumber="5">OMAN</collectingCountry>
:
<specimenCount id="91AAFD24FFE89E2B448BFE70ED42FE7D" box="[493,529,471,497]" count="1" pageId="4" pageNumber="5" type="male">1♂</specimenCount>
,
<location id="82736076FFE89E2B4747FE70ED97FE7E" LSID="urn:lsid:plazi:treatment:0F0587BBFFE89E2B45F1FF30EB79F8F0:82736076FFE89E2B4747FE70ED97FE7E" box="[545,708,471,498]" country="Oman" latitude="23.3" longLatPrecision="754" longitude="57.1" municipality="Wadi Al-Arabein" name="Al Hajar Mts." pageId="4" pageNumber="5">Al Hajar Mts.</location>
nr.
<collectingMunicipality id="6777ACD7FFE89E2B4790FE70ECEFFE7D" box="[758,956,471,497]" pageId="4" pageNumber="5">Wadi Al-Arabein</collectingMunicipality>
,
<geoCoordinate id="E298506AFFE89E2B46ADFE70EB79FE7D" box="[971,1066,471,497]" degrees="23.30" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="23.3">23.30°N</geoCoordinate>
<geoCoordinate id="E298506AFFE89E2B4153FE70EBC0FE7E" box="[1077,1171,471,498]" degrees="57.10" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="57.1">57.10°E</geoCoordinate>
,
<quantity id="40549B48FFE89E2B41C4FE70EBBCFE7E" box="[1186,1263,471,498]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="5" unit="m" value="270.0">
<elevation id="0C81D19EFFE89E2B41C4FE70EBBCFE7E" box="[1186,1263,471,498]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="5" unit="m" value="270.0">270 m</elevation>
</quantity>
,
<date id="F312106DFFE89E2B4199FE70EA28FE7D" box="[1279,1403,471,497]" pageId="4" pageNumber="5" value="2016-03-28">
<collectingDate id="E356E985FFE89E2B4199FE70EA28FE7D" box="[1279,1403,471,497]" pageId="4" pageNumber="5" value="2016-03-28">28.iii.2016</collectingDate>
</date>
,
<collectorName id="2A59537BFFE89E2B40ECFE7FEFB6FD9A" pageId="4" pageNumber="5">J. Monks</collectorName>
leg.; COI sequence: ABABX291-21 (
<collectionCode id="E1BDAE68FFE89E2B47E9FE5AED87FD9B" box="[655,724,509,535]" pageId="4" pageNumber="5">CMK</collectionCode>
: jm017)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1FD85EE57FD0A" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1FD85EE9AFDB0" box="[199,457,546,572]" italics="true" pageId="4" pageNumber="5">Material not examined</emphasis>
.
<materialsCitation id="37C43CF0FFE89E2B44BFFD85EE53FD0A" country="Palestine" location="According" municipality="Ein Gedi" pageId="4" pageNumber="5" specimenCount="2" typeStatus="allotype">
MoA (1989) reported
<specimenCount id="91AAFD24FFE89E2B47B2FD85EC2FFDB0" box="[724,892,546,572]" count="2" pageId="4" pageNumber="5" type="generic">two specimens</specimenCount>
in the Mavromoustakis collection, the type and the “
<typeStatus id="5817880FFFE89E2B45A9FDE0EE62FDED" box="[207,305,583,609]" pageId="4" pageNumber="5" type="allotype">allotype</typeStatus>
”.
<location id="82736076FFE89E2B4421FDE0EEEEFDED" LSID="urn:lsid:plazi:treatment:0F0587BBFFE89E2B45F1FF30EB79F8F0:82736076FFE89E2B4421FDE0EEEEFDED" box="[327,445,583,609]" country="Palestine" municipality="Ein Gedi" name="According" pageId="4" pageNumber="5">According</location>
to
<bibRefCitation id="E33D4B5CFFE89E2B448FFDE0EDAFFDED" author="Mavromoustakis, G. A." box="[489,764,583,609]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis (1948)</bibRefCitation>
, the type location is
<collectingMunicipality id="6777ACD7FFE89E2B4690FDE0EB33FDED" box="[1014,1120,583,609]" pageId="4" pageNumber="5">Ein Gedi</collectingMunicipality>
, located at the border with
<collectingCountry id="FFBB763DFFE89E2B45F1FDCBEE53FD0A" box="[151,256,620,646]" name="Palestine" pageId="4" pageNumber="5">Palestine</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1FD35ED64FC90" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1FD35ED54FD20" box="[199,519,658,684]" italics="true" pageId="4" pageNumber="5">Genetic barcode information</emphasis>
:
<materialsCitation id="37C43CF0FFE89E2B4772FD35ED67FC90" collectionCode="ABABX" collectorName="The &amp; Species ID Tree" country="Oman" location="Oman" pageId="4" pageNumber="5" specimenCode="ABABX291-21, ABABX468-22" specimenCount="3">
DNA sequences were derived from
<specimenCount id="91AAFD24FFE89E2B46FDFD35EB03FD20" box="[923,1104,658,684]" count="3" pageId="4" pageNumber="5" type="generic">three specimens</specimenCount>
, with one from
<collectingCountry id="FFBB763DFFE89E2B419FFD35EA6EFD20" box="[1273,1341,658,684]" name="Oman" pageId="4" pageNumber="5">Oman</collectingCountry>
(jm017) with only 314 bp (
<specimenCode id="D70A9ED6FFE89E2B4405FD10ED4AFD5D" box="[355,537,695,721]" collectionCode="ABABX" pageId="4" pageNumber="5">ABABX291-21</specimenCode>
), but the other two (oll836, oll837) with&gt;600 bp (
<specimenCode id="D70A9ED6FFE89E2B412EFD10EBADFD5D" box="[1096,1278,695,721]" collectionCode="ABABX" pageId="4" pageNumber="5">ABABX468-22</specimenCode>
, ABABX469- 22). A BIN has not yet been assigned.
<collectorName id="2A59537BFFE89E2B4759FD7BED38FD7A" box="[575,619,732,758]" pageId="4" pageNumber="5">The</collectorName>
species is not closely associated with any other
<taxonomicName id="40AC4D2EFFE89E2B41E7FD7BEACFFD7A" box="[1153,1436,732,758]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B41E7FD7BEA16FD7A" box="[1153,1349,732,758]" italics="true" pageId="4" pageNumber="5">Pseudoanthidium</emphasis>
species
</taxonomicName>
in the COI
<collectorName id="2A59537BFFE89E2B4472FCA5EE94FC90" box="[276,455,770,796]" pageId="4" pageNumber="5">Species ID Tree</collectorName>
(
<figureCitation id="1F972A28FFE89E2B44B1FCA5ED78FC90" box="[471,555,770,796]" captionStart="FIGURE 28" captionStartId="43.[151,250,1958,1982]" captionTargetBox="[221,1351,178,1932]" captionTargetId="graphics-20@43.[333,900,178,1905]" captionText="FIGURE 28. Species ID tree as inferred from the DNA sequence of the mitochondrial cytochrome c oxidase subunit 1 gene (COI gene, “barcoding gene”). Bootstrap values above 70% are shown next to the nodes." figureDoi="http://doi.org/10.5281/zenodo.14275914" httpUri="https://zenodo.org/record/14275914/files/figure.png" pageId="4" pageNumber="5">Fig. 28</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1FC8FED08FC00" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1FC8FEE6AFCCD" box="[199,313,808,833]" italics="true" pageId="4" pageNumber="5">Diagnosis</emphasis>
. The species can easily be distinguished from all other species of
<taxonomicName id="40AC4D2EFFE89E2B4122FC80EA5BFCCD" authorityName="Friese" authorityYear="1898" box="[1092,1288,807,833]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE89E2B4122FC80EA5BFCCD" box="[1092,1288,807,833]" italics="true" pageId="4" pageNumber="5">Pseudoanthidium</emphasis>
</taxonomicName>
by its axilla which forms a strong tooth pointed posteriorly. Additionally, the female is distinct from all other species of the genus by its medially swollen mandible.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1FC3FEE24FAFD" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1FC3FEE1EFC3D" box="[199,333,920,945]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
.
<emphasis id="B5D8EABFFFE89E2B4439FC30EEE7FC3D" box="[351,436,919,945]" italics="true" pageId="4" pageNumber="5">Female</emphasis>
(
<figureCitation id="1F972A28FFE89E2B44AEFC30ED4DFC3D" box="[456,542,919,945]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-15@5.[151,1435,181,1881]" captionTargetPageId="5" captionText="FIGURE 2. Pseudoanthidium bytinskii (Mavromoustakis, 1948), form Israel. a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female, with bulging mandibles; d. Mandible of the male with depressed basal part; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240738" httpUri="https://zenodo.org/record/14240738/files/figure.png" pageId="4" pageNumber="5">Fig. 2a</figureCitation>
).
<quantity id="40549B48FFE89E2B4751FC30EDD8FC3E" box="[567,651,919,946]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="mm" value="10.0">10 mm</quantity>
long.—
<emphasis id="B5D8EABFFFE89E2B478AFC30EC7AFC3D" box="[748,809,919,945]" italics="true" pageId="4" pageNumber="5">Head</emphasis>
: Head blocky (as typical for many
<taxonomicName id="40AC4D2EFFE89E2B41B0FC30EFBDFC5A" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B41B0FC30EACFFC3D" box="[1238,1436,919,945]" italics="true" pageId="4" pageNumber="5">Pseudoanthidium</emphasis>
species
</taxonomicName>
); clypeus bell-shaped, finely punctate (
<figureCitation id="1F972A28FFE89E2B47CEFC1BEDAFFC5B" box="[680,764,956,983]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="4" pageNumber="5">Fig. 20</figureCitation>
), with interstices somewhat longer at the base than towards the apex; apical ridge crenulate with a median tubercle larger than others; apical margin largely hidden by white pubescence; mandible yellow with black teeth, with to1 being the strongest, to2 and to3 equally strong, to4- to7 small; to8 the second strongest; mandible with a median bulge on outer face (
<figureCitation id="1F972A28FFE89E2B4133FB8BEB9BFBCA" box="[1109,1224,1068,1094]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-15@5.[151,1435,181,1881]" captionTargetPageId="5" captionText="FIGURE 2. Pseudoanthidium bytinskii (Mavromoustakis, 1948), form Israel. a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female, with bulging mandibles; d. Mandible of the male with depressed basal part; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240738" httpUri="https://zenodo.org/record/14240738/files/figure.png" pageId="4" pageNumber="5">Fig. 2cd</figureCitation>
); upper and lower paraocular area, and supraclypeal area yellow; large yellow spot on the frons between the antennal sockets and the lower ocelli; yellow preoccipital band extending over lower end of gena.—
<taxonomicName id="40AC4D2EFFE89E2B4151FBDFEBE3FB1D" box="[1079,1200,1144,1169]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE89E2B4151FBDFEBE3FB1D" box="[1079,1200,1144,1169]" italics="true" pageId="4" pageNumber="5">Mesosoma</emphasis>
</taxonomicName>
: Scutum black with anterolateral yellow band; large yellow spot on mesepisternum; pronotal lobe yellow with high lamella; apical margin of scutellum widely rounded in dorsal view, lamellate laterally; yellow, interrupted by black maculation in the middle; axilla yellow with a strong tooth pointed posteriorly.—
<emphasis id="B5D8EABFFFE89E2B46D9FB4FECA7FA8D" box="[959,1012,1256,1281]" italics="true" pageId="4" pageNumber="5">Legs</emphasis>
: Yellow.—
<emphasis id="B5D8EABFFFE89E2B4110FB4FEBBFFA8D" box="[1142,1260,1256,1281]" italics="true" pageId="4" pageNumber="5">Metasoma</emphasis>
: Yellow lateral bands covering most of
<collectionCode id="E1BDAE68FFE89E2B44C5FAABEEE4FAAB" box="[419,439,1292,1319]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T</collectionCode>
1
<collectionCode id="E1BDAE68FFE89E2B44B7FAABEEB6FAAB" box="[465,485,1292,1319]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T</collectionCode>
5, contiguous or subcontiguous in the middle;
<collectionCode id="E1BDAE68FFE89E2B468EFAABECA9FAAA" box="[1000,1018,1292,1318]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T</collectionCode>
6 entirely yellow; with an impunctate apical margin; outer margin of
<collectionCode id="E1BDAE68FFE89E2B4492FA95ED55FAC0" box="[500,518,1330,1356]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T</collectionCode>
<quantity id="40549B48FFE89E2B4760FA95ED63FAC0" box="[518,560,1330,1356]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="5" unit="in" value="6.0">6 in</quantity>
dorsal view with a shoulder and a small bilobed extension centrally (
<figureCitation id="1F972A28FFE89E2B405CFA95EADCFAC0" box="[1338,1423,1330,1356]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="4" pageNumber="5">Fig. 22</figureCitation>
); scopal hairs yellow.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1FADBEDBFF96B" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1FADBEE52FA1A" box="[199,257,1404,1430]" italics="true" pageId="4" pageNumber="5">Male</emphasis>
(
<figureCitation id="1F972A28FFE89E2B4468FADBEE33FA1A" box="[270,352,1404,1430]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-15@5.[151,1435,181,1881]" captionTargetPageId="5" captionText="FIGURE 2. Pseudoanthidium bytinskii (Mavromoustakis, 1948), form Israel. a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female, with bulging mandibles; d. Mandible of the male with depressed basal part; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240738" httpUri="https://zenodo.org/record/14240738/files/figure.png" pageId="4" pageNumber="5">Fig. 2b</figureCitation>
).
<quantity id="40549B48FFE89E2B4415FADBEE94FA1B" box="[371,455,1404,1431]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="5" unit="mm" value="8.5">8.5 mm</quantity>
long. The
<specimenCount id="91AAFD24FFE89E2B475CFADBEDEAFA1A" box="[570,697,1404,1430]" count="3" pageId="4" pageNumber="5" type="male">three males</specimenCount>
examined noticeable smaller than the female.—
<emphasis id="B5D8EABFFFE89E2B41ABFADBEA59FA1A" box="[1229,1290,1404,1430]" italics="true" pageId="4" pageNumber="5">Head</emphasis>
: Colouration pattern as in the female; clypeus and paraocular area pale yellow (
<figureCitation id="1F972A28FFE89E2B461DFA05EC9DFA30" box="[891,974,1442,1468]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
), gena and preoccipital area dark yellow; mandible yellow with three strong, black teeth; mandible not bulging as in the female; antennal segment sg1 yellow ventrally, brown dorsally; sg2 brown; sg 3-sg4 yellow-brown, sg5 bicoloured yellow-brown and brown, sg6-sg13 brown.—
<taxonomicName id="40AC4D2EFFE89E2B4464F9B4EE29F9A0" box="[258,378,1555,1580]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE89E2B4464F9B4EE29F9A0" box="[258,378,1555,1580]" italics="true" pageId="4" pageNumber="5">Mesosoma</emphasis>
</taxonomicName>
: As in female.—
<emphasis id="B5D8EABFFFE89E2B4753F9B4ED39F9A0" box="[565,618,1555,1580]" italics="true" pageId="4" pageNumber="5">Legs</emphasis>
: Yellow.—
<emphasis id="B5D8EABFFFE89E2B4781F9B4EC0FF9A0" box="[743,860,1555,1580]" italics="true" pageId="4" pageNumber="5">Metasoma</emphasis>
(
<figureCitation id="1F972A28FFE89E2B460CF9B5ECE8F9A0" box="[874,955,1554,1580]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-15@5.[151,1435,181,1881]" captionTargetPageId="5" captionText="FIGURE 2. Pseudoanthidium bytinskii (Mavromoustakis, 1948), form Israel. a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female, with bulging mandibles; d. Mandible of the male with depressed basal part; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240738" httpUri="https://zenodo.org/record/14240738/files/figure.png" pageId="4" pageNumber="5">Fig. 2e</figureCitation>
): Similar to female; T5T6 entirely yellow; T7 yellow with broad apical emargination (
<figureCitation id="1F972A28FFE89E2B47E6F990EC57F9DD" box="[640,772,1591,1617]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStartId-0="39.[151,250,1918,1942]" captionStartId-1="40.[151,250,1170,1194]" captionTargetBox-0="[173,1415,561,1882]" captionTargetBox-1="[307,1281,311,1152]" captionTargetId-0="figure-169@39.[173,1415,561,1895]" captionTargetId-1="graphics-80@40.[648,939,711,796]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." captionText-1="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi-0="http://doi.org/10.5281/zenodo.14240783" figureDoi-1="http://doi.org/10.5281/zenodo.14275912" httpUri-0="https://zenodo.org/record/14240783/files/figure.png" httpUri-1="https://zenodo.org/record/14275912/files/figure.png" pageId="4" pageNumber="5">Figs 2324</figureCitation>
); S2 with apical fringe of plumose hairs; S3 with modified hairs (undulate hairs basally and long, hooked hairs protruding beyond these basal hairs); S5 with basal comb and projection with black apical comb (approximately 17 more or less equally long digits); S6 laterally with semicircular incisions.—
<emphasis id="B5D8EABFFFE89E2B4478F900EEDBF94D" box="[286,392,1703,1729]" italics="true" pageId="4" pageNumber="5">Genitalia</emphasis>
: Gonostylus approximately parallel-sided, with apex rounded and unnotched; penis valves slender with rounded apex, non-contiguous (
<figureCitation id="1F972A28FFE89E2B47EDF96BED8CF96B" box="[651,735,1740,1767]" captionStart="FIGURE 25" captionStartId="40.[151,250,2009,2033]" captionTargetBox="[176,1412,1240,1996]" captionTargetId="figure-103@40.[176,1412,1240,2001]" captionTargetPageId="40" captionText="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240785" httpUri="https://zenodo.org/record/14240785/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="871336ADFFE89E2B45A1F954EC4BF8DA" blockId="4.[151,1437,359,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1F954EE7CF880" box="[199,303,1779,1804]" italics="true" pageId="4" pageNumber="5">Variation</emphasis>
.
<bibRefCitation id="E33D4B5CFFE89E2B4458F955ED1FF880" author="Mavromoustakis, G. A." box="[318,588,1778,1804]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis (1948)</bibRefCitation>
mentioned three apical and four smaller lower teeth. The female examined here has three large and five smaller lower teeth. Genae described by
<bibRefCitation id="E33D4B5CFFE89E2B46D3F8B0EB97F8BD" author="Mavromoustakis, G. A." box="[949,1220,1815,1841]" pageId="4" pageNumber="5" pagination="428 - 434" refId="ref30580" refString="Mavromoustakis, G. A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428 - 434. https: // doi. org / 10.1080 / 00222934708654652" type="journal article" year="1948">Mavromoustakis (1948)</bibRefCitation>
as broadly yellow, while they are entirely yellow in the examined specimen.
</paragraph>
<paragraph id="871336ADFFE89E2B45A1F8C5EB79F8F0" blockId="4.[151,1437,359,1916]" box="[199,1066,1890,1916]" pageId="4" pageNumber="5">
<emphasis id="B5D8EABFFFE89E2B45A1F8C5EE03F8F0" box="[199,336,1890,1916]" italics="true" pageId="4" pageNumber="5">Distribution</emphasis>
. Levant (
<collectingCountry id="FFBB763DFFE89E2B44DDF8C5EEAFF8F0" box="[443,508,1890,1916]" name="Israel" pageId="4" pageNumber="5">Israel</collectingCountry>
) and Arabian Peninsula (
<collectingCountry id="FFBB763DFFE89E2B4673F8C5EC0EF8F0" box="[789,861,1890,1916]" name="Oman" pageId="4" pageNumber="5">Oman</collectingCountry>
) (new) (
<figureCitation id="1F972A28FFE89E2B46DAF8C5EB4DF8F0" box="[956,1054,1890,1916]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="4" pageNumber="5">Fig. 27a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,348 @@
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<mods:title id="1CC17B31EDE186572EF346D1AF7A97D1">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<subSubSection id="CFB66526FFEB9E2845F1FAE4EC66FAD2" box="[151,821,1347,1374]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="871336ADFFEB9E2845F1FAE4EC66FAD2" blockId="7.[151,821,1347,1410]" box="[151,821,1347,1374]" pageId="7" pageNumber="8">
<heading id="DC5B81C1FFEB9E2845F1FAE4EC66FAD2" bold="true" box="[151,821,1347,1374]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B5D8EABFFFEB9E2845F1FAE4EC66FAD2" bold="true" box="[151,821,1347,1374]" pageId="7" pageNumber="8">
<taxonomicName id="40AC4D2EFFEB9E2845F1FAE4EDF4FAD1" authority="(Friese, 1917)" authorityName="Kasparek" authorityYear="2024" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[151,679,1347,1373]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre" status="comb. nov.">
<emphasis id="B5D8EABFFFEB9E2845F1FAE4EEA5FAD1" bold="true" box="[151,502,1347,1373]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium fulviventre</emphasis>
(
<bibRefCitation id="E33D4B5CFFEB9E284760FAE4EDCCFAD1" author="Friese, H." box="[518,671,1347,1373]" pageId="7" pageNumber="8" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFEB9E2847C8FAE3EC66FAD2" box="[686,821,1348,1374]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845F1FACFEE10FA0E" box="[151,323,1384,1410]" pageId="7" pageNumber="8" type="description">
<paragraph id="871336ADFFEB9E2845F1FACFEE10FA0E" blockId="7.[151,821,1347,1410]" box="[151,323,1384,1410]" pageId="7" pageNumber="8">
(
<figureCitation id="1F972A28FFEB9E2845F9FACFEFBBFA0E" box="[159,232,1384,1410]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
,
<figureCitation id="1F972A28FFEB9E284592FACFEE40FA0E" box="[244,275,1384,1410]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="7" pageNumber="8">20</figureCitation>
,
<figureCitation id="1F972A28FFEB9E284478FACFEE68FA0E" box="[286,315,1384,1410]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="7" pageNumber="8">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845F1FA0DEC08FA69" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="871336ADFFEB9E2845F1FA0DEC3FFA4E" blockId="7.[151,876,1450,1509]" box="[151,876,1450,1474]" pageId="7" pageNumber="8">
<taxonomicName id="40AC4D2EFFEB9E2845F1FA0DEEA6FA4E" ID-CoL="5VCGK" authority="Friese, 1917" authorityName="Friese" authorityYear="1917" box="[151,501,1450,1474]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre">
<emphasis id="B5D8EABFFFEB9E2845F1FA0DEE21FA4E" box="[151,370,1450,1474]" italics="true" pageId="7" pageNumber="8">Anthidium fulviventre</emphasis>
<bibRefCitation id="E33D4B5CFFEB9E28441FFA0DEEA6FA4E" author="Friese, H." box="[377,501,1450,1474]" pageId="7" pageNumber="8" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
</taxonomicName>
.—Female,
<collectingCountry id="FFBB763DFFEB9E28470BFA0DEDE2FA4E" box="[621,689,1450,1474]" name="Russia" pageId="7" pageNumber="8">Russia</collectingCountry>
(Siberia:
<collectingRegion id="4568F84FFFEB9E284672FA0DEC32FA4E" box="[788,865,1450,1474]" country="Russia" name="Irkutsk" pageId="7" pageNumber="8">Irkutsk</collectingRegion>
).
</paragraph>
<paragraph id="871336ADFFEB9E2845F1FA6AEC08FA69" blockId="7.[151,876,1450,1509]" box="[151,859,1485,1509]" pageId="7" pageNumber="8">
<taxonomicName id="40AC4D2EFFEB9E2845F1FA6AEDCBFA69" ID-CoL="5VCGK" authority="Friese, 1917" authorityName="Friese" authorityYear="1917" box="[151,664,1485,1509]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E2845F1FA6AED46FA69" box="[151,533,1485,1509]" italics="true" pageId="7" pageNumber="8">Anthidium (Proanthidium) fulviventre</emphasis>
<bibRefCitation id="E33D4B5CFFEB9E28477AFA6AEDCBFA69" author="Friese, H." box="[540,664,1485,1509]" pageId="7" pageNumber="8" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFEB9E2847D0FA6AEC0BFA69" author="Pasteels, J. J." box="[694,856,1485,1509]" pageId="7" pageNumber="8" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845F1F9B6EA07F973" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="871336ADFFEB9E2845F1F9B6EEF0F9FD" blockId="7.[151,1437,1553,2037]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845F1F9B6EFAAF9A7" box="[151,249,1553,1579]" italics="true" pageId="7" pageNumber="8">Material</emphasis>
:
<materialsCitation id="37C43CF0FFEB9E284461F9B6EEF0F9FD" country="Russia" determinerName="Friese" location="Siberia" pageId="7" pageNumber="8" specimenCount="3" specimenCount-female="3" stateProvince="Irkutsk" typeStatus="holotype">
<collectingCountry id="FFBB763DFFEB9E284461F9B6EE3CF9A7" box="[263,367,1553,1579]" name="Russia" pageId="7" pageNumber="8">RUSSIA</collectingCountry>
:
<typeStatus id="5817880FFFEB9E28441FF9B6EEB7F9A7" box="[377,484,1553,1579]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
,
<specimenCount id="91AAFD24FFEB9E284489F9B6ED6DF9A7" box="[495,574,1553,1579]" count="1" pageId="7" pageNumber="8" type="female">female</specimenCount>
,
<collectingCountry id="FFBB763DFFEB9E28472EF9B6EDE3F9A7" box="[584,688,1553,1579]" name="Russia" pageId="7" pageNumber="8">RUSSIA</collectingCountry>
:
<location id="82736076FFEB9E2847DCF9B6EC5FF9A7" LSID="urn:lsid:plazi:treatment:0F0587BBFFEB9E2645F1FAE4EC14FBD5:82736076FFEB9E2847DCF9B6EC5FF9A7" box="[698,780,1553,1579]" country="Russia" name="Siberia" pageId="7" pageNumber="8" stateProvince="Irkutsk">Siberia</location>
,
<collectingRegion id="4568F84FFFEB9E284671F9B6EC3AF9A7" box="[791,873,1553,1579]" country="Russia" name="Irkutsk" pageId="7" pageNumber="8">Irkutsk</collectingRegion>
, 1897. “
<taxonomicName id="40AC4D2EFFEB9E2846A0F9B6EBEBF9A7" authorityName="Friese" authorityYear="1917" box="[966,1208,1553,1579]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre">
<emphasis id="B5D8EABFFFEB9E2846A0F9B6EBEBF9A7" box="[966,1208,1553,1579]" italics="true" pageId="7" pageNumber="8">Anthidium fulviventre</emphasis>
</taxonomicName>
♀ n. Fr. 1910
<determinerName id="72CFDA69FFEB9E28403EF9B6EACFF9A7" box="[1368,1436,1553,1579]" pageId="7" pageNumber="8">Friese</determinerName>
det.“ / „
<taxonomicName id="40AC4D2EFFEB9E284588F993ED9EF9C2" authority="Fr. J. Pasteels" authorityName="Fr. J. Pasteels" box="[238,717,1588,1615]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre">
<emphasis id="B5D8EABFFFEB9E284588F993ED63F9C2" box="[238,560,1588,1614]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium fulviventre</emphasis>
Fr. J. Pasteels
</taxonomicName>
det. 1964” / “Holotype
<taxonomicName id="40AC4D2EFFEB9E2846BEF993EA47F9C2" authority="Friese" authorityName="Friese" authorityYear="1917" box="[984,1300,1588,1614]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulviventre">
<emphasis id="B5D8EABFFFEB9E2846BEF993EB99F9C2" box="[984,1226,1588,1614]" italics="true" pageId="7" pageNumber="8">Anthidium fulviventre</emphasis>
Friese
</taxonomicName>
♀, det. v. d. Zanden 94” (
<figureCitation id="1F972A28FFEB9E284454F9F0EEC6F9FD" box="[306,405,1623,1649]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="7" pageNumber="8">Figure 4</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="871336ADFFEB9E2845A1F9DCEC35F918" blockId="7.[151,1437,1553,2037]" box="[199,870,1658,1685]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1F9DCED40F919" box="[199,531,1658,1685]" italics="true" pageId="7" pageNumber="8">Genetic barcode information.</emphasis>
No genetic data are available.
</paragraph>
<paragraph id="871336ADFFEB9E2845A1F939EA07F973" blockId="7.[151,1437,1553,2037]" pageId="7" pageNumber="8">
A rare, overlooked species, known only from a single female.
<bibRefCitation id="E33D4B5CFFEB9E284619F939EBA4F934" author="Proshchalykin, M. Yu. &amp; Fateryga, A. V." box="[895,1271,1694,1720]" pageId="7" pageNumber="8" pagination="295 - 308" refId="ref31500" refString="Proshchalykin, M. Yu. &amp; Fateryga, A. V. (2017) 77. Family Megachilidae. In: Belokobylskij, S. A. &amp; Lelej, A. S. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences Supplement, 6, pp. 295 - 308." type="journal article" year="2017">Proshchalykin &amp; Fateryga (2017)</bibRefCitation>
noted that this species has not been mentioned since its description and requires revision. They placed it into
<taxonomicName id="40AC4D2EFFEB9E2841D4F966EA74F957" authorityName="Fabricius" authorityYear="1804" box="[1202,1319,1729,1755]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFEB9E2841D4F966EA74F957" box="[1202,1319,1729,1755]" italics="true" pageId="7" pageNumber="8">Anthidium</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="40AC4D2EFFEB9E2845F1F943EE61F972" box="[151,306,1764,1790]" pageId="7" pageNumber="8" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E2845F1F943EE61F972" box="[151,306,1764,1790]" italics="true" pageId="7" pageNumber="8">Proanthidium</emphasis>
</taxonomicName>
.
<bibRefCitation id="E33D4B5CFFEB9E284426F943ED97F972" author="Proshkalykin, M. Yu. &amp; Astafurova, Y. V." box="[320,708,1764,1790]" pageId="7" pageNumber="8" pagination="26 - 34" refId="ref31455" refString="Proshkalykin, M. Yu. &amp; Astafurova, Y. V. (2017) The history of study of the Russian bees (Hymenoptera, Anthophila). A. I. Kurentsov's Annual Memorial Meetings, 28, 26 - 34." type="journal article" year="2017">Proshkalykin &amp; Astafurova (2017)</bibRefCitation>
classified it as a species with an unclear taxonomic status.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2745A1F8A0EC94FF59" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="multiple">
<paragraph id="871336ADFFEB9E2745A1F8A0EC94FF59" blockId="7.[151,1437,1553,2037]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Originally described by
<bibRefCitation id="E33D4B5CFFEB9E2844B0F8A0ED3DF8AD" author="Friese, H." box="[470,622,1799,1825]" pageId="7" pageNumber="8" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese (1917)</bibRefCitation>
, J. Pasteels assigned it to
<taxonomicName id="40AC4D2EFFEB9E2846E9F8A0EB00F8AD" authorityName="Friese" authorityYear="1898" box="[911,1107,1799,1825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFEB9E2846E9F8A0EB00F8AD" box="[911,1107,1799,1825]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium</emphasis>
</taxonomicName>
in 1964 based on a specimen label. However, in a later publication (
<bibRefCitation id="E33D4B5CFFEB9E28470EF88DEC5EF8C8" author="Pasteels, J. J." box="[616,781,1834,1860]" pageId="7" pageNumber="8" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels 1969</bibRefCitation>
), he attributed it to
<taxonomicName id="40AC4D2EFFEB9E284162F88DEB2AF8C8" authorityName="Fabricius" authorityYear="1804" box="[1028,1145,1834,1860]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFEB9E284162F88DEB2AF8C8" box="[1028,1145,1834,1860]" italics="true" pageId="7" pageNumber="8">Anthidium</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="40AC4D2EFFEB9E28419DF88DEAC5F8C8" box="[1275,1430,1834,1860]" pageId="7" pageNumber="8" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E28419DF88DEAC5F8C8" box="[1275,1430,1834,1860]" italics="true" pageId="7" pageNumber="8">Proanthidium</emphasis>
</taxonomicName>
.
<taxonomicName id="40AC4D2EFFEB9E2845F1F8E9EE61F8E4" box="[151,306,1870,1896]" pageId="7" pageNumber="8" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E2845F1F8E9EE61F8E4" box="[151,306,1870,1896]" italics="true" pageId="7" pageNumber="8">Proanthidium</emphasis>
</taxonomicName>
is a subgenus which is characterized by a high number of mandibular teeth (1011) in the female. However, examination of the
<typeStatus id="5817880FFFEB9E28448AF8D6ED4FF807" box="[492,540,1905,1931]" pageId="7" pageNumber="8">type</typeStatus>
specimen revealed only five teeth. Additionally, the scutellum is not indented in the middle and lacks lateral projections (teeth), which are typical traits of
<taxonomicName id="40AC4D2EFFEB9E284164F833EBCEF822" box="[1026,1181,1940,1966]" pageId="7" pageNumber="8" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E284164F833EBCEF822" box="[1026,1181,1940,1966]" italics="true" pageId="7" pageNumber="8">Proanthidium</emphasis>
</taxonomicName>
. T6 is semi-trapezoid, as in many
<taxonomicName id="40AC4D2EFFEB9E284442F810ED14F85D" box="[292,583,1975,2001]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E284442F810EEBBF85D" box="[292,488,1975,2001]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium</emphasis>
species
</taxonomicName>
, rather than rounded as in
<taxonomicName id="40AC4D2EFFEB9E2846EEF810EB70F85D" box="[904,1059,1975,2001]" pageId="7" pageNumber="8" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFEB9E2846EEF810EB70F85D" box="[904,1059,1975,2001]" italics="true" pageId="7" pageNumber="8">Proanthidium</emphasis>
</taxonomicName>
. Furthermore, the apical margin of the clypeus is thickened, as in many
<taxonomicName id="40AC4D2EFFEB9E284701F87DECDEF878" box="[615,909,2010,2036]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B5D8EABFFFEB9E284701F87DEC78F878" box="[615,811,2010,2036]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium</emphasis>
species.
</taxonomicName>
The clypeus also exhibits denser and smaller punctures apically, another feature typical for
<taxonomicName id="40AC4D2EFFE49E2747FBFF30EC32FF3D" authorityName="Friese" authorityYear="1898" box="[669,865,151,177]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE49E2747FBFF30EC32FF3D" box="[669,865,151,177]" italics="true" pageId="8" pageNumber="9">Pseudoanthidium</emphasis>
</taxonomicName>
. Therefore, it is suggested to assign this species to
<taxonomicName id="40AC4D2EFFE49E2745F1FF1CEE08FF59" authorityName="Friese" authorityYear="1898" box="[151,347,187,213]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE49E2745F1FF1CEE08FF59" box="[151,347,187,213]" italics="true" pageId="8" pageNumber="9">Pseudoanthidium</emphasis>
</taxonomicName>
rather than to the subgenus
<taxonomicName id="40AC4D2EFFE49E2747F1FF1CEC93FF59" box="[663,960,187,213]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Proanthidium">
<emphasis id="B5D8EABFFFE49E2747F1FF1CEC5FFF59" box="[663,780,187,213]" italics="true" pageId="8" pageNumber="9">Anthidium</emphasis>
(
<emphasis id="B5D8EABFFFE49E27467AFF1CEC93FF59" box="[796,960,187,213]" italics="true" pageId="8" pageNumber="9">Proanthidium)</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="D3D36625FFE49E2745F1F81DEB8CF87A" ID-DOI="http://doi.org/10.5281/zenodo.14240743" ID-Zenodo-Dep="14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1978,2002]" targetBox="[164,1422,265,1943]" targetPageId="8" targetType="figure">
<paragraph id="871336ADFFE49E2745F1F81DEB8CF87A" blockId="8.[151,1437,1978,2038]" pageId="8" pageNumber="9">
<emphasis id="B5D8EABFFFE49E2745F1F81DEE47F85E" bold="true" box="[151,276,1978,2002]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="40AC4D2EFFE49E2744F9F81DEC09F85E" authority="(Friese, 1917)" authorityName="Kasparek" authorityYear="2024" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[415,858,1978,2002]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fulviventre" status="stat. nov.">
<emphasis id="B5D8EABFFFE49E2744F9F81DED90F85E" box="[415,707,1978,2002]" italics="true" pageId="8" pageNumber="9">Pseudoanthidium fulviventre</emphasis>
(
<bibRefCitation id="E33D4B5CFFE49E2747B4F81DEC00F85E" author="Friese, H." box="[722,851,1978,2002]" pageId="8" pageNumber="9" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
)
</taxonomicName>
<emphasis id="B5D8EABFFFE49E274604F81DEC90F85E" bold="true" box="[866,963,1978,2002]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="AEEB57C4FFE49E274604F81DEC90F85E" box="[866,963,1978,2002]" pageId="8" pageNumber="9">stat. nov.</taxonomicNameLabel>
</emphasis>
Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d.
<taxonomicName id="40AC4D2EFFE49E274652F879ECF1F879" box="[820,930,2014,2037]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
from dorsal; e. Habitus lateral.
</paragraph>
</caption>
<subSubSection id="CFB66526FFE59E2645A1FF30EEF0FE65" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="871336ADFFE59E2645A1FF30EEF0FE65" blockId="9.[151,1437,151,1113]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1FF30EECEFF3D" box="[199,413,151,177]" italics="true" pageId="9" pageNumber="10">Diagnosis (female)</emphasis>
: Maculation reddish yellow (yellow in most species of
<taxonomicName id="40AC4D2EFFE59E26416EFF30EB9FFF3D" authorityName="Friese" authorityYear="1898" box="[1032,1228,151,177]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE59E26416EFF30EB9FFF3D" box="[1032,1228,151,177]" italics="true" pageId="9" pageNumber="10">Pseudoanthidium</emphasis>
</taxonomicName>
; yellow-red or red in
<taxonomicName id="40AC4D2EFFE59E2645D5FF1DEE5BFF5F" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[179,264,186,211]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE59E2645D5FF1DEE5BFF5F" box="[179,264,186,211]" italics="true" pageId="9" pageNumber="10">P. royoi</emphasis>
</taxonomicName>
,
<taxonomicName id="40AC4D2EFFE59E264470FF1DEE96FF5F" authorityName="Kasparek" authorityYear="2024" box="[278,453,185,211]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFE59E264470FF1DEE96FF5F" box="[278,453,185,211]" italics="true" pageId="9" pageNumber="10">P. microrubrum</emphasis>
</taxonomicName>
,
<taxonomicName id="40AC4D2EFFE59E2644B5FF1DED39FF5F" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[467,618,185,211]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFE59E2644B5FF1DED39FF5F" box="[467,618,185,211]" italics="true" pageId="9" pageNumber="10">P. puncticolle</emphasis>
</taxonomicName>
[partim; only females?],
<taxonomicName id="40AC4D2EFFE59E2646E2FF1DEB49FF5F" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[900,1050,186,211]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="canariense">
<emphasis id="B5D8EABFFFE59E2646E2FF1DEB49FF5F" box="[900,1050,186,211]" italics="true" pageId="9" pageNumber="10">P. canariense</emphasis>
</taxonomicName>
, and
<taxonomicName id="40AC4D2EFFE59E264130FF1DEBECFF5F" authorityName="Vereecken &amp; Litman" authorityYear="2023" box="[1110,1215,185,211]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobii">
<emphasis id="B5D8EABFFFE59E264130FF1DEBECFF5F" box="[1110,1215,185,211]" italics="true" pageId="9" pageNumber="10">P. jacobii</emphasis>
</taxonomicName>
). It is distinguished from
<taxonomicName id="40AC4D2EFFE59E2645B0FF7AEE7DFF7A" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[214,302,221,246]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE59E2645B0FF7AEE7DFF7A" box="[214,302,221,246]" italics="true" pageId="9" pageNumber="10">P. royoi</emphasis>
</taxonomicName>
by the presence of smooth lateral flanges of T6 (as opposed to the median bilobed protrusion in
<taxonomicName id="40AC4D2EFFE59E2640EEFF7AEF82FE94" baseAuthorityName="Dusmet" baseAuthorityYear="1915" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFE59E2640EEFF7AEF82FE94" italics="true" pageId="9" pageNumber="10">P. royoi</emphasis>
</taxonomicName>
), from
<taxonomicName id="40AC4D2EFFE59E26444AFF58EE8DFE95" authorityName="Kasparek" authorityYear="2024" box="[300,478,255,281]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFE59E26444AFF58EE8DFE95" box="[300,478,255,281]" italics="true" pageId="9" pageNumber="10">P. microrubrum</emphasis>
</taxonomicName>
by its size (being a robust species,
<quantity id="40549B48FFE59E2646E7FF59ECBDFE95" box="[897,1006,254,281]" metricMagnitude="0" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="9" pageNumber="10" unit="m" value="9.75" valueMax="10.0" valueMin="9.5">9.510 m</quantity>
in length, compared to the smaller
<quantity id="40549B48FFE59E2640E8FF58EF90FEB0" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="mm" value="8.0">8 mm</quantity>
long
<taxonomicName id="40AC4D2EFFE59E264462FE85EEE7FEB7" authorityName="Kasparek" authorityYear="2024" box="[260,436,289,315]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFE59E264462FE85EEE7FEB7" box="[260,436,289,315]" italics="true" pageId="9" pageNumber="10">P. microrubrum</emphasis>
</taxonomicName>
), from
<taxonomicName id="40AC4D2EFFE59E26476CFE85EDF0FEB7" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[522,675,289,315]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFE59E26476CFE85EDF0FEB7" box="[522,675,289,315]" italics="true" pageId="9" pageNumber="10">P. puncticolle</emphasis>
</taxonomicName>
by its black, broad-lipped apical ridge of the clypeus (narrow, fine, brown ridge in
<taxonomicName id="40AC4D2EFFE59E26442DFEE2EEB6FED2" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[331,485,324,350]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFE59E26442DFEE2EEB6FED2" box="[331,485,324,350]" italics="true" pageId="9" pageNumber="10">P. puncticolle</emphasis>
</taxonomicName>
), and from
<taxonomicName id="40AC4D2EFFE59E264714FEE2EC58FED2" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[626,779,325,350]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="canariense">
<emphasis id="B5D8EABFFFE59E264714FEE2EC58FED2" box="[626,779,325,350]" italics="true" pageId="9" pageNumber="10">P. canariense</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFE59E264621FEE2ECE0FED2" authorityName="Vereecken &amp; Litman" authorityYear="2023" box="[839,947,324,350]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobii">
<emphasis id="B5D8EABFFFE59E264621FEE2ECE0FED2" box="[839,947,324,350]" italics="true" pageId="9" pageNumber="10">P. jacobii</emphasis>
</taxonomicName>
by the apically protruding clypeus (apical margin retracted and covered with a dense, moustache-like fringe of hairs in
<taxonomicName id="40AC4D2EFFE59E264161FEC0EBCCFE0C" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1954" box="[1031,1183,359,384]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="canariense">
<emphasis id="B5D8EABFFFE59E264161FEC0EBCCFE0C" box="[1031,1183,359,384]" italics="true" pageId="9" pageNumber="10">P. canariense</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFE59E2641BFFEC0EA17FE0D" authorityName="Vereecken &amp; Litman" authorityYear="2023" box="[1241,1348,359,385]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobii">
<emphasis id="B5D8EABFFFE59E2641BFFEC0EA17FE0D" box="[1241,1348,359,385]" italics="true" pageId="9" pageNumber="10">P. jacobii</emphasis>
</taxonomicName>
). Other distinguishing characteristics include the protruding clypeus with a thick apical ridge adorned with low tubercles, the strongly inward-curving anterior margin of the clypeus, and T6 being black with a shallow lamellate flange on each side of the middle.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645A1FE55EEC7FBBA" pageId="9" pageNumber="10" type="description">
<paragraph id="871336ADFFE59E2645A1FE55ED62FB9F" blockId="9.[151,1437,151,1113]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1FE55EE18FD87" box="[199,331,498,523]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
(
<emphasis id="B5D8EABFFFE59E264439FE56EEF9FD87" box="[351,426,497,523]" italics="true" pageId="9" pageNumber="10">female</emphasis>
):
<quantity id="40549B48FFE59E2644A2FE56ED14FD80" box="[452,583,497,524]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="9" pageNumber="10" unit="mm" value="9.75" valueMax="10.0" valueMin="9.5">9.510 mm</quantity>
long.—
<emphasis id="B5D8EABFFFE59E2647C0FE56EDB0FD87" box="[678,739,497,523]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
: Black with a broad preoccipital band extending to the area behind the upper third of the eye; clypeus as long as wide, reddish yellow, with a black apical ridge strongly protruding and adorned with rounded tubercles (
<figureCitation id="1F972A28FFE59E2647DDFD90EC7DFDDD" box="[699,814,567,593]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="9" pageNumber="10">Figs 4bc</figureCitation>
,
<figureCitation id="1F972A28FFE59E26465CFD90EC09FDDD" box="[826,858,567,593]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="9" pageNumber="10">20</figureCitation>
); broad transition zone between the disc and ridge, also black, with fine punctation; basal margin of clypeus curving inward; mandible yellow with five brown, rounded teeth separated by acute notches; supraclypeal area partly impunctate; large reddish yellow maculation present in lower paraocular area; small area around the ocelli impunctate; antenna brown, with the base of the scape light yellowish brown.—
<taxonomicName id="40AC4D2EFFE59E26441EFD65EEA3FD57" box="[376,496,706,731]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFE59E26441EFD65EEA3FD57" box="[376,496,706,731]" italics="true" pageId="9" pageNumber="10">Mesosoma</emphasis>
</taxonomicName>
: Scutum shining, densely punctate, with punctures mostly separated by half their diameters, but up to two diameters in the centre (
<figureCitation id="1F972A28FFE59E2647BCFD43EC62FD72" box="[730,817,740,766]" captionStart="FIGURE 4" captionStartId="8.[151,250,1978,2002]" captionTargetBox="[164,1422,265,1943]" captionTargetId="figure-47@8.[164,1422,265,1961]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Pseudoanthidium fulviventre (Friese, 1917) stat. nov. Female from Russia. a. Apical terga; b. Face. Note the black, straight apical ridge of the clypeus; c. Clypeus; d. Mesosoma from dorsal; e. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240743" httpUri="https://zenodo.org/record/14240743/files/figure.png" pageId="9" pageNumber="10">Fig. 4d</figureCitation>
); scutellum and axillae reddish yellow except for the base; outer margin of scutellum lamellate laterally; scutellum conspicuously overhanging propodeum.—
<emphasis id="B5D8EABFFFE59E264046FCA0EAC6FCAC" box="[1312,1429,775,800]" italics="true" pageId="9" pageNumber="10">Metasoma</emphasis>
: Terga finely punctate, noticeably finer than on scutum; punctation somewhat irregular, with punctures 13 puncture diameters apart in anterior terga; punctation coarser and denser on posterior terga; puncture size variable on all terga; T1T5 with reddish yellow lateral maculations; spots on T1T3 large, but not reaching the middle; T4 with medium-sized spots, and T5 with a very small spot; T6 entirely black, with dense, rugulose punctation; median lobe and small smooth shoulder on each side of the middle (in dorsal view) (
<figureCitation id="1F972A28FFE59E2646A2FC13EB4AFC42" box="[964,1049,948,974]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="9" pageNumber="10">Fig. 22</figureCitation>
); S1S5 reddish yellow, S6 black; ventral scopa yellow to blackish brown.—
<emphasis id="B5D8EABFFFE59E26471AFC70EDE2FC7C" box="[636,689,983,1008]" italics="true" pageId="9" pageNumber="10">Legs</emphasis>
: Reddish yellow with black base (trochanter, anterior femora).—
<emphasis id="B5D8EABFFFE59E2645F1FC5DEF82FB9F" box="[151,209,1018,1043]" italics="true" pageId="9" pageNumber="10">Wing</emphasis>
: Strongly brownish infuscated.
</paragraph>
<paragraph id="871336ADFFE59E2645A1FBBBEEC7FBBA" blockId="9.[151,1437,151,1113]" box="[199,404,1052,1079]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1FBBBEE52FBBA" box="[199,257,1052,1078]" italics="true" pageId="9" pageNumber="10">Male</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFE59E2645A1FB98EC14FBD5" box="[199,839,1087,1113]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="871336ADFFE59E2645A1FB98EC14FBD5" blockId="9.[151,1437,151,1113]" box="[199,839,1087,1113]" pageId="9" pageNumber="10">
<emphasis id="B5D8EABFFFE59E2645A1FB98EE03FBD5" box="[199,336,1087,1113]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
: Only known from
<collectingRegion id="4568F84FFFE59E264749FB98EDD2FBD5" box="[559,641,1087,1113]" country="Russia" name="Irkutsk" pageId="9" pageNumber="10">Irkutsk</collectingRegion>
, Siberia (
<collectingCountry id="FFBB763DFFE59E26478CFB98EC68FBD5" box="[746,827,1087,1113]" name="Russia" pageId="9" pageNumber="10">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,259 @@
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<mods:title id="9B6848264B3C39016007A579F1CE9389">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<mods:namePart id="B51D7956CF9C1C669CA9FE23E938C0C6">Kasparek, Max</mods:namePart>
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<mods:part id="65081BEAB21D1B7C79DEB11C1DFE3850">
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<treatment id="0F0587BBFFEB9E2845F1FF30EDEFFB78" ID-DOI="http://doi.org/10.5281/zenodo.14248289" ID-Zenodo-Dep="14248289" LSID="urn:lsid:plazi:treatment:0F0587BBFFEB9E2845F1FF30EDEFFB78" httpUri="http://treatment.plazi.org/id/0F0587BBFFEB9E2845F1FF30EDEFFB78" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="CFB66526FFEB9E2845F1FF30ED15FF3D" box="[151,582,151,178]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="871336ADFFEB9E2845F1FF30ED15FF3D" blockId="7.[151,582,151,213]" box="[151,582,151,178]" pageId="7" pageNumber="8">
<heading id="DC5B81C1FFEB9E2845F1FF30ED15FF3D" bold="true" box="[151,582,151,178]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B5D8EABFFFEB9E2845F1FF30ED15FF3D" bold="true" box="[151,582,151,178]" pageId="7" pageNumber="8">
<taxonomicName id="40AC4D2EFFEB9E2845F1FF30EEB2FF3D" authorityName="Kasparek" authorityYear="2024" box="[151,481,151,178]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="farsiense" status="sp. nov.">
<emphasis id="B5D8EABFFFEB9E2845F1FF30EEB2FF3D" bold="true" box="[151,481,151,178]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium farsiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFEB9E28448FFF30ED15FF3D" box="[489,582,151,177]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845F1FF1CEE10FF59" box="[151,323,187,213]" pageId="7" pageNumber="8" type="description">
<paragraph id="871336ADFFEB9E2845F1FF1CEE10FF59" blockId="7.[151,582,151,213]" box="[151,323,187,213]" pageId="7" pageNumber="8">
(
<figureCitation id="1F972A28FFEB9E2845F9FF1CEFB4FF59" box="[159,231,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1920,1944]" captionTargetBox="[152,1436,174,1911]" captionTargetId="figure-19@6.[152,1436,174,1911]" captionTargetPageId="6" captionText="FIGURE 3. Pseudoanthidium farsiense Kasparek sp. nov. Female from Iran. a. Habitus dorsal; b. Apical terga and scopa. Note the noticeable long scopal hairs; c. Head, with protruding apical ridge of the clypeus; d. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240740" httpUri="https://zenodo.org/record/14240740/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
,
<figureCitation id="1F972A28FFEB9E284592FF1CEE40FF59" box="[244,275,187,213]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="7" pageNumber="8">20</figureCitation>
,
<figureCitation id="1F972A28FFEB9E284478FF1CEE68FF59" box="[286,315,187,213]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="7" pageNumber="8">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845F1FEA6EEE1FE09" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="871336ADFFEB9E2845F1FEA6EB42FEB2" blockId="7.[151,1437,257,1269]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845F1FEA6EFAAFE97" box="[151,249,257,283]" italics="true" pageId="7" pageNumber="8">Material</emphasis>
.
<materialsCitation id="37C43CF0FFEB9E284461FEA6EB42FEB2" collectingDate="2019-05-04" collectionCode="CMK" collectorName="C. Schmid-Egger" country="Iran" elevation="1677" latitude="29.1855" location="Fars prov." longLatPrecision="6" longitude="54.1389" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Fars prov." typeStatus="holotype">
<typeStatus id="5817880FFFEB9E284461FEA6EE20FE97" box="[263,371,257,283]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
.
<specimenCount id="91AAFD24FFEB9E28441BFEA6EE80FE97" box="[381,467,257,283]" pageId="7" pageNumber="8" type="female">Female</specimenCount>
.
<collectingCountry id="FFBB763DFFEB9E2844BBFEA6ED77FE97" box="[477,548,257,283]" name="Iran" pageId="7" pageNumber="8">IRAN</collectingCountry>
,
<collectingRegion id="4568F84FFFEB9E284756FEA6EDF0FE97" box="[560,675,257,283]" country="Iran" name="Fars" pageId="7" pageNumber="8">Fars prov.</collectingRegion>
,
<locationDeviation id="96A12F8FFFEB9E2847CBFEA6ECF6FE97" box="[685,933,257,284]" pageId="7" pageNumber="8">
<quantity id="40549B48FFEB9E2847CBFEA6EDA8FE97" box="[685,763,257,284]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="km" value="2.5">2.5 km</quantity>
SW Haji Abad
</locationDeviation>
(
<geoCoordinate id="E298506AFFEB9E2846D5FEA6EB7CFE90" box="[947,1071,257,284]" degrees="29.1855" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="29.1855">29.1855°N</geoCoordinate>
<geoCoordinate id="E298506AFFEB9E284150FEA6EBE1FE90" box="[1078,1202,257,284]" degrees="54.1389" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="54.1389">54.1389°E</geoCoordinate>
),
<quantity id="40549B48FFEB9E2841A2FEA6EA48FE97" box="[1220,1307,257,283]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="7" pageNumber="8" unit="m" value="1677.0">
<elevation id="0C81D19EFFEB9E2841A2FEA6EA48FE97" box="[1220,1307,257,283]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="7" pageNumber="8" unit="m" value="1677.0">1677 m</elevation>
</quantity>
,
<date id="F312106DFFEB9E284041FEA6EACBFE97" box="[1319,1432,257,283]" pageId="7" pageNumber="8" value="2019-05-04">
<collectingDate id="E356E985FFEB9E284041FEA6EACBFE97" box="[1319,1432,257,283]" pageId="7" pageNumber="8" value="2019-05-04">04.v.2019</collectingDate>
</date>
,
<collectorName id="2A59537BFFEB9E2845F1FE83EE09FEB2" box="[151,346,292,318]" pageId="7" pageNumber="8">C. Schmid-Egger</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFEB9E284722FE83EDA6FEB3" box="[580,757,292,319]" pageId="7" pageNumber="8">ABABX275-21</accessionNumber>
(
<collectionCode id="E1BDAE68FFEB9E284662FE83EC1AFEB2" box="[772,841,292,318]" pageId="7" pageNumber="8">CMK</collectionCode>
: IR-c04; seg079).
</materialsCitation>
</paragraph>
<paragraph id="871336ADFFEB9E2845A1FEEFEEE1FE09" blockId="7.[151,1437,257,1269]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FEEFED41FEED" box="[199,530,327,354]" italics="true" pageId="7" pageNumber="8">Genetic barcode information.</emphasis>
The barcode of the
<typeStatus id="5817880FFFEB9E284788FEE0EC03FEED" box="[750,848,327,353]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
(seg079) is available under number ABABX275-21. BIN is BOLD:AEL4174.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845A1FE29EC2AFD9D" pageId="7" pageNumber="8" type="description">
<paragraph id="871336ADFFEB9E2845A1FE29EC2AFD9D" blockId="7.[151,1437,257,1269]" pageId="7" pageNumber="8">
In the barcode analysis,
<taxonomicName id="40AC4D2EFFEB9E2844BBFE28EDDCFE24" authorityName="Kasparek" authorityYear="2024" box="[477,655,398,424]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFEB9E2844BBFE28EDDCFE24" box="[477,655,398,424]" italics="true" pageId="7" pageNumber="8">P. microrubrum</emphasis>
</taxonomicName>
turned out to be the nearest neighbour, with a barcode gap of 5.2%. This value is identical with the average distance. The next neighbour is
<taxonomicName id="40AC4D2EFFEB9E2846ADFE15EB17FE47" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[971,1092,433,459]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEB9E2846ADFE15EB17FE47" box="[971,1092,433,459]" italics="true" pageId="7" pageNumber="8">P. alpinum</emphasis>
</taxonomicName>
with a closest distance gap of 5.5% and an average distance 5.8%. DNA sequences of the closely related
<taxonomicName id="40AC4D2EFFEB9E28468FFE72EA03FE62" authority="(Morawitz, 1888)" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[1001,1360,468,494]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFEB9E28468FFE72EBD2FE62" box="[1001,1153,468,494]" italics="true" pageId="7" pageNumber="8">P. puncticolle</emphasis>
(
<bibRefCitation id="E33D4B5CFFEB9E2841F7FE73EA14FE62" author="Morawitz, F. F." box="[1169,1351,468,494]" pageId="7" pageNumber="8" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz, 1888</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFEB9E2840EEFE72EFADFD9D" authorityName="Kasparek" authorityYear="2024" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="syriacum" status="sp. nov.">
<emphasis id="B5D8EABFFFEB9E2840EEFE72EFADFD9D" italics="true" pageId="7" pageNumber="8">P. syriacum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFEB9E284463FE5FEE0FFD9E" box="[261,348,504,530]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
(
<taxonomicName id="40AC4D2EFFEB9E28440DFE5FEEB0FD9D" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[363,483,503,529]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEB9E28440DFE5FEEB0FD9D" box="[363,483,503,529]" italics="true" pageId="7" pageNumber="8">P. alpinum</emphasis>
</taxonomicName>
group, see below) are not available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845A1FDBCEB21FD4D" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="871336ADFFEB9E2845A1FDBCEB21FD4D" blockId="7.[151,1437,257,1269]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FDBCEE6AFDB8" box="[199,313,539,564]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
. T6 semi triangular-shaped (in dorsal view), a feature shared with
<taxonomicName id="40AC4D2EFFEB9E28415FFDBCEB80FDB8" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[1081,1235,538,564]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFEB9E28415FFDBCEB80FDB8" box="[1081,1235,538,564]" italics="true" pageId="7" pageNumber="8">P. puncticolle</emphasis>
</taxonomicName>
,
<taxonomicName id="40AC4D2EFFEB9E284185FDBCEAC6FDB8" authorityName="Kasparek" authorityYear="2024" box="[1251,1429,538,564]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFEB9E284185FDBCEAC6FDB8" box="[1251,1429,538,564]" italics="true" pageId="7" pageNumber="8">P. microrubrum</emphasis>
</taxonomicName>
, and
<taxonomicName id="40AC4D2EFFEB9E2845A0FD98EE14FDD4" authorityName="Kasparek" authorityYear="2024" box="[198,327,575,600]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFEB9E2845A0FD98EE14FDD4" box="[198,327,575,600]" italics="true" pageId="7" pageNumber="8">P. syriacum</emphasis>
</taxonomicName>
. Punctation of terga fine (rough in
<taxonomicName id="40AC4D2EFFEB9E2847ACFD98EC33FDD4" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[714,864,574,600]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFEB9E2847ACFD98EC33FDD4" box="[714,864,574,600]" italics="true" pageId="7" pageNumber="8">P. puncticolle</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFEB9E2846F3FD98EB44FDD4" authorityName="Kasparek" authorityYear="2024" box="[917,1047,575,600]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFEB9E2846F3FD98EB44FDD4" box="[917,1047,575,600]" italics="true" pageId="7" pageNumber="8">P. syriacum</emphasis>
</taxonomicName>
), maculation of integument yellow (red in
<emphasis id="B5D8EABFFFEB9E28458FFDC5EEF1FDF7" box="[233,418,609,635]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="40AC4D2EFFEB9E28458FFDC5EEC8FDF7" authorityName="Kasparek" authorityYear="2024" box="[233,411,609,635]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microrubrum">P. microrubrum</taxonomicName>
;
</emphasis>
partly reddish in
<taxonomicName id="40AC4D2EFFEB9E28470EFDC5EC53FDF7" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[616,768,609,635]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFEB9E28470EFDC5EC53FDF7" box="[616,768,609,635]" italics="true" pageId="7" pageNumber="8">P. puncticolle</emphasis>
</taxonomicName>
), scopal hairs yellow (black in
<taxonomicName id="40AC4D2EFFEB9E284104FDC5EA41FDF7" authorityName="Kasparek" authorityYear="2024" box="[1122,1298,609,635]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFEB9E284104FDC5EA41FDF7" box="[1122,1298,609,635]" italics="true" pageId="7" pageNumber="8">P. microrubrum</emphasis>
</taxonomicName>
), and apical ridge of clypeus little protruding outwards (strongly protruding outwards in
<taxonomicName id="40AC4D2EFFEB9E284172FD22EB94FD12" authorityName="Kasparek" authorityYear="2024" box="[1044,1223,644,670]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFEB9E284172FD22EB94FD12" box="[1044,1223,644,670]" italics="true" pageId="7" pageNumber="8">P. microrubrum</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFEB9E284060FD22EADFFD12" authorityName="Kasparek" authorityYear="2024" box="[1286,1420,645,670]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFEB9E284060FD22EADFFD12" box="[1286,1420,645,670]" italics="true" pageId="7" pageNumber="8">P. syriacum</emphasis>
</taxonomicName>
).
<taxonomicName id="40AC4D2EFFEB9E2845F1FD00EE94FD4D" authorityName="Kasparek" authorityYear="2024" box="[151,455,679,705]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFEB9E2845F1FD00EE94FD4D" box="[151,455,679,705]" italics="true" pageId="7" pageNumber="8">Pseudoanthidium farsiense</emphasis>
</taxonomicName>
is conspicuously smaller than
<taxonomicName id="40AC4D2EFFEB9E284646FD0FECF1FD4D" authorityName="Kasparek" authorityYear="2024" box="[800,930,680,705]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFEB9E284646FD0FECF1FD4D" box="[800,930,680,705]" italics="true" pageId="7" pageNumber="8">P. syriacum</emphasis>
</taxonomicName>
(
<quantity id="40549B48FFEB9E2846D4FD00ECA0FD4E" box="[946,1011,679,706]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="8" unit="mm" value="7.0">7 mm</quantity>
vs.
<quantity id="40549B48FFEB9E284147FD00EB35FD4D" box="[1057,1126,679,705]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="9.0">9 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845A1FD6CEEC7FB22" pageId="7" pageNumber="8" type="description">
<paragraph id="871336ADFFEB9E2845A1FD6CEA3AFB07" blockId="7.[151,1437,257,1269]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FD6CEE18FD68" box="[199,331,715,740]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
(
<emphasis id="B5D8EABFFFEB9E284439FD6DEEF9FD68" box="[351,426,714,740]" italics="true" pageId="7" pageNumber="8">female</emphasis>
).
<quantity id="40549B48FFEB9E2844A2FD6CED54FD69" box="[452,519,715,741]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="8" unit="mm" value="7.0">7 mm</quantity>
long.—
<emphasis id="B5D8EABFFFEB9E284700FD6DEDF0FD68" box="[614,675,714,740]" italics="true" pageId="7" pageNumber="8">Head</emphasis>
: Black except for a bright yellow lower paraocular area, clypeus, and a broad preoccipital band reaching the upper one third of the eye (
<figureCitation id="1F972A28FFEB9E2846ABFD49EB77FC84" box="[973,1060,750,776]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="7" pageNumber="8">Fig. 20</figureCitation>
); clypeus 1.1 times broader than long, with impunctate base and dense punctation across the disc; apical ridge convexly lip-like protruding, with approximately five strong tubercles (
<figureCitation id="1F972A28FFEB9E284748FC93ED2DFCC2" box="[558,638,820,846]" captionStart="FIGURE 3" captionStartId="6.[151,250,1920,1944]" captionTargetBox="[152,1436,174,1911]" captionTargetId="figure-19@6.[152,1436,174,1911]" captionTargetPageId="6" captionText="FIGURE 3. Pseudoanthidium farsiense Kasparek sp. nov. Female from Iran. a. Habitus dorsal; b. Apical terga and scopa. Note the noticeable long scopal hairs; c. Head, with protruding apical ridge of the clypeus; d. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240740" httpUri="https://zenodo.org/record/14240740/files/figure.png" pageId="7" pageNumber="8">Fig. 3c</figureCitation>
); transition zone between disc and apical ridge brown; mandible yellow, gradually changing to brown towards the teeth; five black teeth; antenna black except for reddish-brown sg3- sg5.—
<taxonomicName id="40AC4D2EFFEB9E284587FCDCEE0AFC18" box="[225,345,891,916]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFEB9E284587FCDCEE0AFC18" box="[225,345,891,916]" italics="true" pageId="7" pageNumber="8">Mesosoma</emphasis>
</taxonomicName>
: Scutum with dense punctation and narrow, shining ridges between the punctures; broad yellow lateral bands anteriorly; scutellum overhanging propodeum, with a narrow, lamellate, transparent apical margin; yellow laterally and black medially; axilla yellow with some black remnants proximately; pronotal lobe with high anterolateral lamella; omaulus rounded; mesepisternum with long white pubescence.—
<emphasis id="B5D8EABFFFEB9E284115FC42EBBBFC72" box="[1139,1256,997,1022]" italics="true" pageId="7" pageNumber="8">Metasoma</emphasis>
: Lateral yellow bands on terga, becoming closer towards the apex; punctation denser on depressions than on discs; T6 semi triangular in dorsal view (
<figureCitation id="1F972A28FFEB9E284420FB8DEEC9FBC9" box="[326,410,1066,1093]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
), black with rugulose surface; ventral scopa with long, golden hairs (
<figureCitation id="1F972A28FFEB9E2841C4FB8DEBA6FBC8" box="[1186,1269,1066,1092]" captionStart="FIGURE 3" captionStartId="6.[151,250,1920,1944]" captionTargetBox="[152,1436,174,1911]" captionTargetId="figure-19@6.[152,1436,174,1911]" captionTargetPageId="6" captionText="FIGURE 3. Pseudoanthidium farsiense Kasparek sp. nov. Female from Iran. a. Habitus dorsal; b. Apical terga and scopa. Note the noticeable long scopal hairs; c. Head, with protruding apical ridge of the clypeus; d. Habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.14240740" httpUri="https://zenodo.org/record/14240740/files/figure.png" pageId="7" pageNumber="8">Fig. 3b</figureCitation>
); S6 triangular with grey hairs.—
<emphasis id="B5D8EABFFFEB9E28440CFBE8EECCFBE4" box="[362,415,1103,1128]" italics="true" pageId="7" pageNumber="8">Legs</emphasis>
: Femora black with brown apices; tibiae yellow, inner surface brownish; mid and hind basitarsi blackish; hind basitarsus with short bristles on the inner face.—
<emphasis id="B5D8EABFFFEB9E2846A2FBD5EB5BFB07" box="[964,1032,1138,1163]" italics="true" pageId="7" pageNumber="8">Wings</emphasis>
: Strongly brownish infuscated.
</paragraph>
<paragraph id="871336ADFFEB9E2845A1FB33EEC7FB22" blockId="7.[151,1437,257,1269]" box="[199,404,1172,1199]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FB33EE52FB22" box="[199,257,1172,1198]" italics="true" pageId="7" pageNumber="8">Male</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEB9E2845A1FB1FEDEFFB78" pageId="7" pageNumber="8" type="etymology">
<paragraph id="871336ADFFEB9E2845A1FB1FEAC0FB5D" blockId="7.[151,1437,257,1269]" box="[199,1427,1207,1234]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FB1FEED9FB5D" box="[199,394,1208,1233]" italics="true" pageId="7" pageNumber="8">Derivatio nominis</emphasis>
. Named after
<collectingRegion id="4568F84FFFEB9E284742FB1FED06FB5E" box="[548,597,1208,1234]" country="Iran" name="Fars" pageId="7" pageNumber="8">Fars</collectingRegion>
, the historical region of southern
<collectingCountry id="FFBB763DFFEB9E2846D8FB1FECBFFB5E" box="[958,1004,1208,1234]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
, where the
<typeStatus id="5817880FFFEB9E284100FB1FEBC7FB5E" box="[1126,1172,1208,1234]" pageId="7" pageNumber="8">type</typeStatus>
material was collected.
</paragraph>
<paragraph id="871336ADFFEB9E2845A1FB7DEDEFFB78" blockId="7.[151,1437,257,1269]" box="[199,700,1242,1269]" pageId="7" pageNumber="8">
<emphasis id="B5D8EABFFFEB9E2845A1FB7DEE03FB78" box="[199,336,1242,1268]" italics="true" pageId="7" pageNumber="8">Distribution</emphasis>
.
<collectingCountry id="FFBB763DFFEB9E284438FB7CEED8FB79" box="[350,395,1243,1269]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
(
<collectingRegion id="4568F84FFFEB9E2844FCFB7CED64FB78" box="[410,567,1242,1269]" country="Iran" name="Fars" pageId="7" pageNumber="8">Fars province</collectingRegion>
) (
<figureCitation id="1F972A28FFEB9E28472BFB7DEDFCFB78" box="[589,687,1242,1268]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="7" pageNumber="8">Fig. 27b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,210 @@
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<mods:title id="5E34C3C3B05CF47FF5039AABAD5EC98D">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<mods:title id="63674B169F0C207A892C210A961D9CF6">Zootaxa</mods:title>
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<mods:date id="CCBA4BE95ED56A3852C30ECFECC98F63">2024</mods:date>
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<mods:number id="37A57A3C0FD762BA1312620C0DD4ECB3">2024-11-25</mods:number>
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<mods:number id="9465208458C05FCBB3C17C26E381580F">5541</mods:number>
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<mods:identifier id="BFF4BD5C6C109E6D19C319382B7850CA" type="DOI">10.11646/zootaxa.5541.1.1</mods:identifier>
<mods:identifier id="8C2E58AB73D4AD06D755C1BBE671D4FB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0F0587BBFFEE9E2D45F1FB55EC4BF84B" ID-DOI="http://doi.org/10.5281/zenodo.14248287" ID-Zenodo-Dep="14248287" LSID="urn:lsid:plazi:treatment:0F0587BBFFEE9E2D45F1FB55EC4BF84B" httpUri="http://treatment.plazi.org/id/0F0587BBFFEE9E2D45F1FB55EC4BF84B" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="CFB66526FFEE9E2D45F1FB55EDE4FA80" box="[151,695,1266,1293]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="871336ADFFEE9E2D45F1FB55EDE4FA80" blockId="2.[151,695,1266,1329]" box="[151,695,1266,1293]" pageId="2" pageNumber="3">
<heading id="DC5B81C1FFEE9E2D45F1FB55EDE4FA80" bold="true" box="[151,695,1266,1293]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="40AC4D2EFFEE9E2D45F1FB55EDE4FA80" ID-CoL="4NP3X" authority="(Morawitz, 1873)" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[151,695,1266,1293]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEE9E2D45F1FB55EDE4FA80" bold="true" box="[151,695,1266,1293]" pageId="2" pageNumber="3">
<emphasis id="B5D8EABFFFEE9E2D45F1FB55EE8AFA81" bold="true" box="[151,473,1266,1293]" italics="true" pageId="2" pageNumber="3">Pseudoanthidium alpinum</emphasis>
(
<bibRefCitation id="E33D4B5CFFEE9E2D448EFB55EDFCFA80" author="Morawitz, F. F." box="[488,687,1266,1292]" pageId="2" pageNumber="3" pagination="129 - 189" refId="ref30815" refString="Morawitz, F. F. (1873 [1874]) Die Bienen Daghestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 10, 129 - 189." type="book chapter" year="1873">Morawitz, 1873</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEE9E2D45F1FAB0EEB8FABD" box="[151,491,1303,1329]" pageId="2" pageNumber="3" type="description">
<paragraph id="871336ADFFEE9E2D45F1FAB0EEB8FABD" blockId="2.[151,695,1266,1329]" box="[151,491,1303,1329]" pageId="2" pageNumber="3">
(
<figureCitation id="1F972A28FFEE9E2D45F9FAB0EFB5FABD" box="[159,230,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1843]" captionTargetId="figure-15@3.[151,1435,181,1843]" captionTargetPageId="3" captionText="FIGURE 1. Pseudoanthidium alpinum (Morawitz, 1873). a. Habitus dorsal, female (Greece); b. Habitus dorsal, male (Greece); c. Habitus lateral, female (Romania, holotype of Anthidium serraticeps Friese, 1917, a junior synonym of P. alpinum). d. Apical terga and scopa, female (Kazakhstan). e. Ventral side of the metasoma of the male with felt-like hairs, specialized hairs on S3, and combs on S5 (Spain)." figureDoi="http://doi.org/10.5281/zenodo.14240736" httpUri="https://zenodo.org/record/14240736/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D4592FAB0EE41FABD" box="[244,274,1303,1329]" captionStart="FIGURE 18" captionStartId="31.[151,250,1966,1990]" captionTargetBox="[201,1385,1117,1943]" captionTargetId="figure-37@31.[201,1385,1117,1943]" captionTargetPageId="31" captionText="FIGURE 18. Pseudoanthidium tajikistanicum Kasparek sp. nov. Female from Tajikistan. a. Habitus dorsal; b. Habitus lateral; c. Scutellum and axillae." figureDoi="http://doi.org/10.5281/zenodo.14240771" httpUri="https://zenodo.org/record/14240771/files/figure.png" pageId="2" pageNumber="3">18</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D4478FAB0EE6FFABD" box="[286,316,1303,1329]" captionStart="FIGURE 19" captionStartId="32.[151,250,1582,1606]" captionTargetBox="[151,1435,669,1558]" captionTargetId="figure-197@32.[151,1435,669,1558]" captionTargetPageId="32" captionText="FIGURE 19. Pseudoanthidium variabile (Pasteels, 1980). Female holotype from Oman. a. Habitus dorsal; b. Habitus lateral; c. Metasomal terga." figureDoi="http://doi.org/10.5281/zenodo.14240773" httpUri="https://zenodo.org/record/14240773/files/figure.png" pageId="2" pageNumber="3">19</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D442EFAB0EE34FABD" box="[328,359,1303,1329]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="2" pageNumber="3">20</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D4414FAB0EEC3FABD" box="[370,400,1303,1329]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="2" pageNumber="3">21</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D44FAFAB0EEE8FABD" box="[412,443,1303,1329]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="2" pageNumber="3">22</figureCitation>
,
<figureCitation id="1F972A28FFEE9E2D44A0FAB0EEB0FABD" box="[454,483,1303,1329]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="2" pageNumber="3">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEE9E2D45F1FAFCEB1AF90F" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="871336ADFFEE9E2D45F1FAFCEE5AFA1E" blockId="2.[151,1437,1371,1488]" pageId="2" pageNumber="3">
<taxonomicName id="40AC4D2EFFEE9E2D45F1FAFCED52FAFF" ID-CoL="8SFJ4" authority="Morawitz, 1873" authorityName="Morawitz" authorityYear="1873" box="[151,513,1371,1395]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEE9E2D45F1FAFCEE0AFAFF" box="[151,345,1371,1395]" italics="true" pageId="2" pageNumber="3">Anthidium alpinum</emphasis>
<bibRefCitation id="E33D4B5CFFEE9E2D4439FAFCED52FAFF" author="Morawitz, F. F." box="[351,513,1371,1395]" pageId="2" pageNumber="3" pagination="129 - 189" refId="ref30815" refString="Morawitz, F. F. (1873 [1874]) Die Bienen Daghestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 10, 129 - 189." type="book chapter" year="1873">Morawitz, 1873</bibRefCitation>
</taxonomicName>
.—Male,
<collectingCountry id="FFBB763DFFEE9E2D4706FAFCEDFBFAFF" box="[608,680,1371,1395]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
:
<collectingRegion id="4568F84FFFEE9E2D47D7FAFBEC40FAF8" box="[689,787,1372,1396]" country="Russia" name="Dagestan" pageId="2" pageNumber="3">Dagestan</collectingRegion>
.
<typeStatus id="5817880FFFEE9E2D467DFAFCEC2AFAFF" box="[795,889,1371,1395]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
in ZISP examined by photograph (
<bibRefCitation id="E33D4B5CFFEE9E2D41BEFAFCEFADFA1E" author="Kasparek, M. &amp; Ebmer, A. W." pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer 2023</bibRefCitation>
).
</paragraph>
<paragraph id="871336ADFFEE9E2D45F1FA3FEDD7FA3D" blockId="2.[151,1437,1371,1488]" box="[151,644,1432,1457]" pageId="2" pageNumber="3">
<taxonomicName id="40AC4D2EFFEE9E2D45F1FA3FED54FA3D" ID-CoL="8SFJ7" authority="Mocsary, 1884" authorityName="Mocsary" authorityYear="1884" box="[151,519,1432,1457]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="barbatum">
<emphasis id="B5D8EABFFFEE9E2D45F1FA3FEE39FA3C" box="[151,362,1432,1456]" italics="true" pageId="2" pageNumber="3">Anthidium barbatum</emphasis>
<bibRefCitation id="E33D4B5CFFEE9E2D4416FA3EED54FA3D" author="Mocsary, A." box="[368,519,1433,1457]" pageId="2" pageNumber="3" pagination="241 - 278" refId="ref30778" refString="Mocsary, A. (1884) Species generis Anthidium Fabr. regionis Palaearcticae. Termeszetrajzi Fuzetek Kiadija a Magyar Nemzeti Muzeum [Naturhistorische Hefte herausgegeben vom Ungarischen National-Museum], 8, 241 - 278." type="journal article" year="1884">Mocsáry, 1884</bibRefCitation>
</taxonomicName>
.—
<collectingCountry id="FFBB763DFFEE9E2D4742FA3EEDD3FA3D" box="[548,640,1433,1457]" name="Hungary" pageId="2" pageNumber="3">Hungary</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFEE9E2D45F1FA10EA6AFA43" blockId="2.[151,1437,1371,1488]" box="[151,1337,1463,1488]" pageId="2" pageNumber="3">
<taxonomicName id="40AC4D2EFFEE9E2D45F1FA10EEA9FA43" ID-CoL="5VCJ5" authority="Friese, 1917" authorityName="Friese" authorityYear="1917" box="[151,506,1463,1487]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="serraticeps">
<emphasis id="B5D8EABFFFEE9E2D45F1FA10EE2BFA43" box="[151,376,1463,1487]" italics="true" pageId="2" pageNumber="3">Anthidium serraticeps</emphasis>
<bibRefCitation id="E33D4B5CFFEE9E2D4418FA10EEA9FA43" author="Friese, H." box="[382,506,1463,1487]" pageId="2" pageNumber="3" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
</taxonomicName>
.—
<typeStatus id="5817880FFFEE9E2D477EFA10ED2AFA43" box="[536,633,1463,1487]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
in ZMB examined, synonymy confirmed (
<bibRefCitation id="E33D4B5CFFEE9E2D414DFA10EA7DFA43" author="Kasparek, M. &amp; Ebmer, A. W." box="[1067,1326,1463,1487]" pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer 2023</bibRefCitation>
).
</paragraph>
<paragraph id="871336ADFFEE9E2D45F1FA59EB1AF90F" blockId="2.[151,1437,1533,1991]" pageId="2" pageNumber="3">
<emphasis id="B5D8EABFFFEE9E2D45F1FA59EEB1F99B" box="[151,482,1533,1560]" italics="true" pageId="2" pageNumber="3">Genetic barcode information</emphasis>
. Three DNA barcoding sequences are available (ABABX465-22, ABABX466-22, ABABX467-22) and were analysed by
<bibRefCitation id="E33D4B5CFFEE9E2D4730F986ECD5F9B7" author="Kasparek, M. &amp; Ebmer, A. W." box="[598,902,1569,1595]" pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer (2023)</bibRefCitation>
. The COI sequence allowed for the assignment of the previously unknown female to the male. A BIN has not yet been assigned.
<taxonomicName id="40AC4D2EFFEE9E2D417CF9E2EA13F9D3" ID-CoL="4NP3X" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[1050,1344,1605,1631]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEE9E2D417CF9E2EA13F9D3" box="[1050,1344,1605,1631]" italics="true" pageId="2" pageNumber="3">Pseudoanthidium alpinum</emphasis>
</taxonomicName>
clusters in the Species ID Tree with
<taxonomicName id="40AC4D2EFFEE9E2D44B6F9CDED03F90F" authorityName="Kasparek" authorityYear="2024" box="[464,592,1641,1667]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFEE9E2D44B6F9CDED03F90F" box="[464,592,1641,1667]" italics="true" pageId="2" pageNumber="3">P. farsiense</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFEE9E2D47E1F9CDEC65F90F" authorityName="Kasparek" authorityYear="2024" box="[647,822,1641,1667]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFEE9E2D47E1F9CDEC65F90F" box="[647,822,1641,1667]" italics="true" pageId="2" pageNumber="3">P. microrubrum</emphasis>
</taxonomicName>
(see under
<taxonomicName id="40AC4D2EFFEE9E2D46DEF9CDEB6AF90F" authorityName="Kasparek" authorityYear="2024" box="[952,1081,1641,1667]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFEE9E2D46DEF9CDEB6AF90F" box="[952,1081,1641,1667]" italics="true" pageId="2" pageNumber="3">P. farsiense</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFEE9E2D45A1F929EC4BF84B" pageId="2" pageNumber="3" type="description">
<paragraph id="871336ADFFEE9E2D45A1F929EC1BF947" blockId="2.[151,1437,1533,1991]" pageId="2" pageNumber="3">
<emphasis id="B5D8EABFFFEE9E2D45A1F929EE54F92B" box="[199,263,1678,1703]" italics="true" pageId="2" pageNumber="3">Notes</emphasis>
.
<typeStatus id="5817880FFFEE9E2D4471F929EE1CF924" box="[279,335,1678,1704]" pageId="2" pageNumber="3">Type</typeStatus>
species of the genus
<taxonomicName id="40AC4D2EFFEE9E2D472AF92AEC43F92B" authorityName="Friese" authorityYear="1898" box="[588,784,1677,1703]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFEE9E2D472AF92AEC43F92B" box="[588,784,1677,1703]" italics="true" pageId="2" pageNumber="3">Pseudoanthidium</emphasis>
</taxonomicName>
. Descriptions of both sexes, as well as photographs, are provided by
<bibRefCitation id="E33D4B5CFFEE9E2D4443F916EEB3F947" author="Kasparek, M." box="[293,480,1713,1739]" pageId="2" pageNumber="3" refId="ref29929" refString="Kasparek, M. (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe and Western Turkey. Identification. Distribution. Biology. Chimaira, Frankfurt, 292 pp. [unchanged reprint 2023]" type="book" year="2022">Kasparek (2022)</bibRefCitation>
and
<bibRefCitation id="E33D4B5CFFEE9E2D4771F916EC10F947" author="Kasparek, M. &amp; Ebmer, A. W." box="[535,835,1713,1739]" pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer (2023)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFEE9E2D45A1F971EB04F8F3" blockId="2.[151,1437,1533,1991]" pageId="2" pageNumber="3">
In addition to the records in
<bibRefCitation id="E33D4B5CFFEE9E2D4762F972EC63F963" author="Kasparek, M. &amp; Ebmer, A. W." box="[516,816,1749,1775]" pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer (2023)</bibRefCitation>
, the BOLD database (www.boldsystems.org) lists
<specimenCount id="91AAFD24FFEE9E2D4002F972EE5FF89F" count="3" pageId="2" pageNumber="3" type="generic">three specimens</specimenCount>
in the Naturalis Biodiversity Center, Leiden (
<collectingCountry id="FFBB763DFFEE9E2D4672F95EEC81F89F" box="[788,978,1785,1811]" name="Netherlands" pageId="2" pageNumber="3">The Netherlands</collectingCountry>
) from Iktol [Terskol;
<geoCoordinate id="E298506AFFEE9E2D41ACF95EEA7AF898" box="[1226,1321,1785,1812]" degrees="43.25" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="555" value="43.25">43.25°N</geoCoordinate>
<geoCoordinate id="E298506AFFEE9E2D4057F95EEAC2F898" box="[1329,1425,1785,1812]" degrees="42.51" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="42.51">42.51°E</geoCoordinate>
], Caucasus (
<collectingCountry id="FFBB763DFFEE9E2D4476F8BAEE33F8BB" box="[272,352,1821,1847]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
),
<date id="F312106DFFEE9E2D4415F8BAEEBDF8BB" box="[371,494,1821,1847]" pageId="2" pageNumber="3" value="1927-06-27">27.vi.1927</date>
. Identification has been confirmed by T. J. Wood in 2024. Furthermore,
<bibRefCitation id="E33D4B5CFFEE9E2D4070F8BAEE60F8D7" author="Kasparek, M. &amp; Ebmer, A. W." pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer (2023)</bibRefCitation>
reported a specimen from
<collectingCountry id="FFBB763DFFEE9E2D4738F8E6EDFCF8D7" box="[606,687,1857,1883]" name="Turkey" pageId="2" pageNumber="3">Turkey</collectingCountry>
, whose character traits are somewhat different from
<taxonomicName id="40AC4D2EFFEE9E2D419CF8E5EA22F8D7" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[1274,1393,1857,1883]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEE9E2D419CF8E5EA22F8D7" box="[1274,1393,1857,1883]" italics="true" pageId="2" pageNumber="3">P. alpinum</emphasis>
</taxonomicName>
. As only a single female was available, they provisionally assigned it to “
<taxonomicName id="40AC4D2EFFEE9E2D46F9F8C1EB10F8F3" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[927,1091,1893,1919]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFEE9E2D46F9F8C1ECE6F8F3" box="[927,949,1894,1919]" italics="true" pageId="2" pageNumber="3">P.</emphasis>
aff.
<emphasis id="B5D8EABFFFEE9E2D4681F8C2EB10F8F3" box="[999,1091,1893,1919]" italics="true" pageId="2" pageNumber="3">alpinum</emphasis>
</taxonomicName>
”.
</paragraph>
<paragraph id="871336ADFFEE9E2D45F1F82EEC4BF84B" blockId="2.[151,1437,1533,1991]" pageId="2" pageNumber="3">
<emphasis id="B5D8EABFFFEE9E2D45F1F82EEE7BF82F" box="[151,296,1929,1955]" italics="true" pageId="2" pageNumber="3">Distribution.</emphasis>
Georgia,
<collectingCountry id="FFBB763DFFEE9E2D44C6F82EEEA1F82F" box="[416,498,1929,1955]" name="Greece" pageId="2" pageNumber="3">Greece</collectingCountry>
,
<collectingCountry id="FFBB763DFFEE9E2D4764F82DED36F828" box="[514,613,1930,1956]" name="Hungary" pageId="2" pageNumber="3">Hungary</collectingCountry>
,
<collectingCountry id="FFBB763DFFEE9E2D4713F82EEDE5F82F" box="[629,694,1929,1955]" name="Israel" pageId="2" pageNumber="3">Israel</collectingCountry>
,
<collectingCountry id="FFBB763DFFEE9E2D47A1F82EEC1EF82F" box="[711,845,1929,1955]" name="Kazakhstan" pageId="2" pageNumber="3">Kazakhstan</collectingCountry>
,
<collectingCountry id="FFBB763DFFEE9E2D463AF82EEC97F82F" box="[860,964,1929,1955]" name="Romania" pageId="2" pageNumber="3">Romania</collectingCountry>
,
<collectingCountry id="FFBB763DFFEE9E2D46B3F82EEB73F82F" box="[981,1056,1929,1955]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
(
<collectingRegion id="4568F84FFFEE9E2D4155F82EEBF3F82F" box="[1075,1184,1929,1955]" country="Russia" name="Dagestan" pageId="2" pageNumber="3">Dagestan</collectingRegion>
, Kabardino-Balkarian Republic),
<collectingCountry id="FFBB763DFFEE9E2D4472F80AEE04F84B" box="[276,343,1965,1991]" name="Spain" pageId="2" pageNumber="3">Spain</collectingCountry>
, and
<collectingCountry id="FFBB763DFFEE9E2D44F6F80AEEB3F84B" box="[400,480,1965,1991]" name="Turkey" pageId="2" pageNumber="3">Turkey</collectingCountry>
(
<bibRefCitation id="E33D4B5CFFEE9E2D4489F80AEC5EF84B" author="Kasparek, M. &amp; Ebmer, A. W." box="[495,781,1965,1991]" pageId="2" pageNumber="3" pagination="39 - 50" refId="ref29969" refString="Kasparek, M. &amp; Ebmer, A. W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39 - 50. https: // doi. org / 10.47446 / OSMIA 11.7" type="journal article" year="2023">Kasparek &amp; Ebmer 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,198 @@
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<mods:title id="FD384EFEF4BA049812553B96AB96BCBE">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<treatment id="0F0587BBFFF19E3145F1FB0DEDB5FECD" ID-DOI="http://doi.org/10.5281/zenodo.14248303" ID-Zenodo-Dep="14248303" LSID="urn:lsid:plazi:treatment:0F0587BBFFF19E3145F1FB0DEDB5FECD" httpUri="http://treatment.plazi.org/id/0F0587BBFFF19E3145F1FB0DEDB5FECD" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="CFB66526FFF19E3245F1FB0DED9AFB49" box="[151,713,1194,1221]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="871336ADFFF19E3245F1FB0DED9AFB49" blockId="29.[151,713,1194,1257]" box="[151,713,1194,1221]" pageId="29" pageNumber="30">
<heading id="DC5B81C1FFF19E3245F1FB0DED9AFB49" bold="true" box="[151,713,1194,1221]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B5D8EABFFFF19E3245F1FB0DED9AFB49" bold="true" box="[151,713,1194,1221]" pageId="29" pageNumber="30">
<taxonomicName id="40AC4D2EFFF19E3245F1FB0DED37FB49" authority="Kasparek" authorityName="Kasparek" authorityYear="2024" box="[151,612,1194,1221]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="syriacum" status="sp. nov.">
<emphasis id="B5D8EABFFFF19E3245F1FB0DEEB0FB49" bold="true" box="[151,483,1194,1221]" italics="true" pageId="29" pageNumber="30">Pseudoanthidium syriacum</emphasis>
Kasparek
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFF19E32470DFB0CED9AFB49" box="[619,713,1195,1221]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245F1FB68EE02FB65" box="[151,337,1230,1257]" pageId="29" pageNumber="30" type="description">
<paragraph id="871336ADFFF19E3245F1FB68EE02FB65" blockId="29.[151,713,1194,1257]" box="[151,337,1230,1257]" pageId="29" pageNumber="30">
(
<figureCitation id="1F972A28FFF19E3245F9FB68EFA5FB65" box="[159,246,1230,1257]" captionStart="FIGURE 15" captionStartId="28.[151,250,1003,1027]" captionTargetBox="[243,1344,191,967]" captionTargetId="figure-19@28.[230,1357,181,980]" captionTargetPageId="28" captionText="FIGURE 15. Comparison of body size of Pseudoanthidium royoi and P. melanurum, as expressed by the marginal cell length of the wing. Pseudoanthidium melanurum: female N= 21, male N= 31; P. royoi N=1 for each sex." figureDoi="http://doi.org/10.5281/zenodo.14240765" httpUri="https://zenodo.org/record/14240765/files/figure.png" pageId="29" pageNumber="30">Figs 15</figureCitation>
,
<figureCitation id="1F972A28FFF19E324464FB68EE72FB65" box="[258,289,1231,1257]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="29" pageNumber="30">20</figureCitation>
,
<figureCitation id="1F972A28FFF19E32444AFB68EE1AFB65" box="[300,329,1231,1257]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="29" pageNumber="30">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245F1FAB0EC69FAF5" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="871336ADFFF19E3245F1FAB0ED41FAD9" blockId="29.[151,1437,1302,2013]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245F1FAB0EE22FABD" box="[151,369,1303,1329]" italics="true" pageId="29" pageNumber="30">Material examined.</emphasis>
<materialsCitation id="37C43CF0FFF19E324413FAB0ED41FAD9" collectingDate="1996-05-23" collectionCode="CMK" collectorName="Ma. Halada" country="Syria" latitude="33.75" location="Rankos" longLatPrecision="721" longitude="36.39" municipality="Female" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Dimashq" typeStatus="holotype">
<typeStatus id="5817880FFFF19E324413FAB0EEB2FABD" box="[373,481,1303,1329]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="FFBB763DFFF19E32448EFAB0ED13FABD" box="[488,576,1303,1329]" name="Syria" pageId="29" pageNumber="30">SYRIA</collectingCountry>
:
<collectingMunicipality id="6777ACD7FFF19E32472FFAB0EDCDFABD" box="[585,670,1303,1329]" pageId="29" pageNumber="30">Female</collectingMunicipality>
,
<location id="82736076FFF19E3247C1FAB0EDA8FABD" LSID="urn:lsid:plazi:treatment:0F0587BBFFF19E3145F1FB0DEDB5FECD:82736076FFF19E3247C1FAB0EDA8FABD" box="[679,763,1303,1329]" country="Syria" latitude="33.75" longLatPrecision="721" longitude="36.39" municipality="Female" name="Rankos" pageId="29" pageNumber="30" stateProvince="Dimashq">Rankos</location>
[Rankus],
<locationDeviation id="96A12F8FFFF19E324617FAB0EB15FABD" box="[881,1094,1303,1329]" pageId="29" pageNumber="30">
<quantity id="40549B48FFF19E324617FAB0ECE5FABD" box="[881,950,1303,1329]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">40 km</quantity>
N Damascus
</locationDeviation>
(
<geoCoordinate id="E298506AFFF19E324134FAB1EBE1FABD" box="[1106,1202,1302,1329]" degrees="33.75" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="33.75">33.75°N</geoCoordinate>
<geoCoordinate id="E298506AFFF19E3241D1FAB0EA45FABD" box="[1207,1302,1303,1329]" degrees="36.39" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="36.39">36.39°E</geoCoordinate>
),
<date id="F312106DFFF19E324041FAB0EACBFABD" box="[1319,1432,1303,1329]" pageId="29" pageNumber="30" value="1996-05-23">
<collectingDate id="E356E985FFF19E324041FAB0EACBFABD" box="[1319,1432,1303,1329]" pageId="29" pageNumber="30" value="1996-05-23">23.v.1996</collectingDate>
</date>
,
<collectorName id="2A59537BFFF19E3245F1FA9CEE49FAD9" box="[151,282,1339,1365]" pageId="29" pageNumber="30">Ma. Halada</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF19E32443FFA9CEECDFAD9" box="[345,414,1339,1365]" pageId="29" pageNumber="30">CMK</collectionCode>
: ms3348).
</materialsCitation>
</paragraph>
<paragraph id="871336ADFFF19E3245A1FAF8EC69FAF5" blockId="29.[151,1437,1302,2013]" box="[199,826,1375,1401]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FAF8ED58FAF5" box="[199,523,1375,1401]" italics="true" pageId="29" pageNumber="30">Genetic barcode information</emphasis>
. No information available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245A1FA24EBFEF9DD" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="871336ADFFF19E3245A1FA24EBFEF9DD" blockId="29.[151,1437,1302,2013]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FA24EEFAFA11" box="[199,425,1411,1437]" italics="true" pageId="29" pageNumber="30">Diagnosis (female).</emphasis>
Species with extensive yellow colouration. Black apical ridge of the clypeus strongly protruding with nine irregularly-size tubercles.T6 crescent-shaped with a small median emargination to accommodate the sting. The most similar species is
<taxonomicName id="40AC4D2EFFF19E324751FA6CEDE4FA69" authorityName="Kasparek" authorityYear="2024" box="[567,695,1483,1509]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFF19E324751FA6CEDE4FA69" box="[567,695,1483,1509]" italics="true" pageId="29" pageNumber="30">P. farsiense</emphasis>
</taxonomicName>
, which possesses five tubercles on the clypeal ridge, decreasing in size from the middle to the lateral side, while
<taxonomicName id="40AC4D2EFFF19E3247EAFA48EC5EF984" authorityName="Kasparek" authorityYear="2024" box="[652,781,1519,1544]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFF19E3247EAFA48EC5EF984" box="[652,781,1519,1544]" italics="true" pageId="29" pageNumber="30">P. syriacum</emphasis>
</taxonomicName>
has nine tubercles of irregular and differing sizes. A yellow preoccipital band extends to the malar area in
<taxonomicName id="40AC4D2EFFF19E3247C4F9B4EC76F9A0" authorityName="Kasparek" authorityYear="2024" box="[674,805,1555,1580]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFF19E3247C4F9B4EC76F9A0" box="[674,805,1555,1580]" italics="true" pageId="29" pageNumber="30">P. syriacum</emphasis>
</taxonomicName>
, whereas it reaches only the upper third of the eye in
<taxonomicName id="40AC4D2EFFF19E3240EEF9B4EFAFF9DD" authorityName="Kasparek" authorityYear="2024" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFF19E3240EEF9B4EFAFF9DD" italics="true" pageId="29" pageNumber="30">P. farsiense</emphasis>
</taxonomicName>
. Tergal punctation is conspicuously coarser in
<taxonomicName id="40AC4D2EFFF19E32466CF990ECDEF9DC" authorityName="Kasparek" authorityYear="2024" box="[778,909,1591,1616]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="B5D8EABFFFF19E32466CF990ECDEF9DC" box="[778,909,1591,1616]" italics="true" pageId="29" pageNumber="30">P. syriacum</emphasis>
</taxonomicName>
compared to
<taxonomicName id="40AC4D2EFFF19E324143F990EBF5F9DD" authorityName="Kasparek" authorityYear="2024" box="[1061,1190,1591,1617]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="farsiense">
<emphasis id="B5D8EABFFFF19E324143F990EBF5F9DD" box="[1061,1190,1591,1617]" italics="true" pageId="29" pageNumber="30">P. farsiense</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3145A1F9FCEEC7FF75" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="871336ADFFF19E3145A1F9FCEB9EFF59" blockId="29.[151,1437,1302,2013]" lastBlockId="30.[151,1437,150,321]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1F9FCEE18F9F8" box="[199,331,1627,1652]" italics="true" pageId="29" pageNumber="30">Description</emphasis>
.
<emphasis id="B5D8EABFFFF19E324406F9FCEEE7F9F9" box="[352,436,1627,1653]" italics="true" pageId="29" pageNumber="30">Female</emphasis>
(
<figureCitation id="1F972A28FFF19E3244ADF9FCED76F9F9" box="[459,549,1627,1653]" captionStart="FIGURE 17" captionStartId="31.[151,245,1048,1072]" captionTargetBox="[201,1386,151,1032]" captionTargetId="figure-15@31.[201,1386,151,1032]" captionTargetPageId="31" captionText="FIGURE 17. Pseudoanthidium syriacum Kasparek sp. nov. Female from Syria. a. Habitus dorsal; b. Habitus lateral; c. Apical terga." figureDoi="http://doi.org/10.5281/zenodo.14240769" httpUri="https://zenodo.org/record/14240769/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
).
<quantity id="40549B48FFF19E324727F9FCEDD8F9F9" box="[577,651,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="29" pageNumber="30" unit="mm" value="9.0">9 mm</quantity>
.—
<emphasis id="B5D8EABFFFF19E3247CDF9FCEDBBF9F9" box="[683,744,1627,1653]" italics="true" pageId="29" pageNumber="30">Head</emphasis>
: Head blocky, typical of many
<taxonomicName id="40AC4D2EFFF19E324114F9FCEACEF9F9" box="[1138,1437,1627,1653]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B5D8EABFFFF19E324114F9FCEA65F9F9" box="[1138,1334,1627,1653]" italics="true" pageId="29" pageNumber="30">Pseudoanthidium</emphasis>
species.
</taxonomicName>
Extensive yellow colouration on lower and upper paraocular areas up to the zenith of the eye, on the clypeus, and a yellow preoccipital band reaching the malar area; clypeus broader than long (approximately 1.1 times wider than long); basal area of clypeus impunctate and polished; preapical area depressed with dense punctation; apical ridge strongly protruding and black, featuring about nine blunt tubercles, with the median one being the most prominent (
<figureCitation id="1F972A28FFF19E3245F9F8A8EFA7F8A5" box="[159,244,1807,1833]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="29" pageNumber="30">Fig. 20</figureCitation>
); mandible elongate, with five black teeth decreasing in size from distal to proximal.—
<taxonomicName id="40AC4D2EFFF19E3241A6F8A8EA6BF8A4" box="[1216,1336,1807,1832]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF19E3241A6F8A8EA6BF8A4" box="[1216,1336,1807,1832]" italics="true" pageId="29" pageNumber="30">Mesosoma</emphasis>
</taxonomicName>
: Scutum black, with boomerang-shaped anterolateral yellow band; scutellum and axillae yellow except for scutellar base; outer margin angulate, laterally with a row of transparent cells, depressed in the middle; one yellow spot each on mesepisternum and metepisternum; omaulus angular; punctation on scutum dense, partly honeycomb-like with narrow, shining interstices.—
<emphasis id="B5D8EABFFFF19E324481F838ED0FF834" box="[487,604,1951,1976]" italics="true" pageId="29" pageNumber="30">Metasoma</emphasis>
: Broad yellow bands on T1T6, almost reaching the middle but further apart in proximal terga; punctation coarse, punctures on disc large (
<figureCitation id="1F972A28FFF19E3246E9F864ECB0F851" box="[911,995,1987,2013]" captionStart="FIGURE 17" captionStartId="31.[151,245,1048,1072]" captionTargetBox="[201,1386,151,1032]" captionTargetId="figure-15@31.[201,1386,151,1032]" captionTargetPageId="31" captionText="FIGURE 17. Pseudoanthidium syriacum Kasparek sp. nov. Female from Syria. a. Habitus dorsal; b. Habitus lateral; c. Apical terga." figureDoi="http://doi.org/10.5281/zenodo.14240769" httpUri="https://zenodo.org/record/14240769/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
); T6 crescent-shaped with a very small median emargination to accommodate the sting (
<figureCitation id="1F972A28FFF29E3147D5FF30EC45FF3D" box="[691,790,150,177]" captionStart="FIGURE 15" captionStartId="28.[151,250,1003,1027]" captionTargetBox="[243,1344,191,967]" captionTargetId="figure-19@28.[230,1357,181,980]" captionTargetPageId="28" captionText="FIGURE 15. Comparison of body size of Pseudoanthidium royoi and P. melanurum, as expressed by the marginal cell length of the wing. Pseudoanthidium melanurum: female N= 21, male N= 31; P. royoi N=1 for each sex." figureDoi="http://doi.org/10.5281/zenodo.14240765" httpUri="https://zenodo.org/record/14240765/files/figure.png" pageId="30" pageNumber="31">Figs 15c</figureCitation>
,
<figureCitation id="1F972A28FFF29E314646FF30EC13FF3D" box="[800,832,151,177]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="30" pageNumber="31">22</figureCitation>
); scopal hairs golden.—
<emphasis id="B5D8EABFFFF29E31412BFF30EBD1FF3C" box="[1101,1154,151,176]" italics="true" pageId="30" pageNumber="31">Legs</emphasis>
: Yellow, expect for black trochanteres and basal femora.—
<emphasis id="B5D8EABFFFF29E31476FFF1CED10FF58" box="[521,579,187,212]" italics="true" pageId="30" pageNumber="31">Wing</emphasis>
: Forewing slightly infuscated, particularly on distal parts.
</paragraph>
<paragraph id="871336ADFFF29E3145A1FF78EEC7FF75" blockId="30.[151,1437,150,321]" box="[199,404,223,249]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FF78EE5BFF75" box="[199,264,223,249]" italics="true" pageId="30" pageNumber="31">Male.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FEA4EBECFE91" box="[199,1215,259,285]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="871336ADFFF29E3145A1FEA4EBECFE91" blockId="30.[151,1437,150,321]" box="[199,1215,259,285]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FEA4EEFBFE90" box="[199,424,259,284]" italics="true" pageId="30" pageNumber="31">Derivation nominis.</emphasis>
Adjective. Named after
<collectingCountry id="FFBB763DFFF29E3147DCFEA4EDABFE91" box="[698,760,259,285]" name="Syria" pageId="30" pageNumber="31">Syria</collectingCountry>
, where the
<typeStatus id="5817880FFFF29E314611FEA4ECF4FE91" box="[887,935,259,285]" pageId="30" pageNumber="31">type</typeStatus>
specimen was collected.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FE80EDB5FECD" box="[199,742,295,321]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="871336ADFFF29E3145A1FE80EDB5FECD" blockId="30.[151,1437,150,321]" box="[199,742,295,321]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FE80EE03FECD" box="[199,336,295,321]" italics="true" pageId="30" pageNumber="31">Distribution</emphasis>
. Only known from
<collectingCountry id="FFBB763DFFF29E314748FE80ED3AFECD" box="[558,617,295,321]" name="Syria" pageId="30" pageNumber="31">Syria</collectingCountry>
(
<figureCitation id="1F972A28FFF29E31471EFE80ED8AFECD" box="[632,729,295,321]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="30" pageNumber="31">Fig. 27a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,234 @@
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<mods:title id="DEB12096A42F967D39B702D71329F54F">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<subSubSection id="CFB66526FFF29E3145F1FA49EDF6F984" box="[151,677,1518,1545]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="871336ADFFF29E3145F1FA49EDF6F984" blockId="30.[151,677,1518,1581]" box="[151,677,1518,1545]" pageId="30" pageNumber="31">
<heading id="DC5B81C1FFF29E3145F1FA49EDF6F984" bold="true" box="[151,677,1518,1545]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="40AC4D2EFFF29E3145F1FA49EDF6F984" ID-CoL="4NP55" authority="(Pasteels, 1980)" baseAuthorityName="Pasteels" baseAuthorityYear="1980" box="[151,677,1518,1545]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="variabile">
<emphasis id="B5D8EABFFFF29E3145F1FA49EDF6F984" bold="true" box="[151,677,1518,1545]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145F1FA49EE8DF985" bold="true" box="[151,478,1518,1545]" italics="true" pageId="30" pageNumber="31">Pseudoanthidium variabile</emphasis>
(Pasteels, 1980)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145F1F9B4EE02F9A1" box="[151,337,1555,1581]" pageId="30" pageNumber="31" type="description">
<paragraph id="871336ADFFF29E3145F1F9B4EE02F9A1" blockId="30.[151,677,1518,1581]" box="[151,337,1555,1581]" pageId="30" pageNumber="31">
(
<figureCitation id="1F972A28FFF29E3145F9F9B4EFA5F9A1" box="[159,246,1555,1581]" captionStart="FIGURE 17" captionStartId="31.[151,245,1048,1072]" captionTargetBox="[201,1386,151,1032]" captionTargetId="figure-15@31.[201,1386,151,1032]" captionTargetPageId="31" captionText="FIGURE 17. Pseudoanthidium syriacum Kasparek sp. nov. Female from Syria. a. Habitus dorsal; b. Habitus lateral; c. Apical terga." figureDoi="http://doi.org/10.5281/zenodo.14240769" httpUri="https://zenodo.org/record/14240769/files/figure.png" pageId="30" pageNumber="31">Figs 17</figureCitation>
,
<figureCitation id="1F972A28FFF29E314464F9B4EE72F9A1" box="[258,289,1555,1581]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="30" pageNumber="31">20</figureCitation>
,
<figureCitation id="1F972A28FFF29E31444AF9B4EE1AF9A1" box="[300,329,1555,1581]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="30" pageNumber="31">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145F1F9F0EB64F902" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="871336ADFFF29E3145F1F9F0ED9DF9E3" blockId="30.[151,1079,1623,1678]" box="[151,718,1623,1647]" pageId="30" pageNumber="31">
<taxonomicName id="40AC4D2EFFF29E3145F1F9F0ED4CF9E3" authority="Pasteels, 1980" authorityName="Pasteels" authorityYear="1980" box="[151,543,1623,1647]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="variabile">
<emphasis id="B5D8EABFFFF29E3145F1F9F0EED4F9E3" box="[151,391,1623,1647]" italics="true" pageId="30" pageNumber="31">Royanthidium variabile</emphasis>
Pasteels, 1980
</taxonomicName>
.—
<collectingCountry id="FFBB763DFFF29E31475BF9F0ED2DF9E3" box="[573,638,1623,1647]" name="Oman" pageId="30" pageNumber="31">Oman</collectingCountry>
, female
</paragraph>
<paragraph id="871336ADFFF29E3145F1F9D1EB64F902" blockId="30.[151,1079,1623,1678]" box="[151,1079,1654,1678]" pageId="30" pageNumber="31">
<taxonomicName id="40AC4D2EFFF29E3145F1F9D1EDB6F902" authority="Pasteels, 1980" authorityName="Pasteels" authorityYear="1980" box="[151,741,1654,1678]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="variabile" subGenus="Royanthidium">
<emphasis id="B5D8EABFFFF29E3145F1F9D1ED1CF902" box="[151,591,1654,1678]" italics="true" pageId="30" pageNumber="31">Pseudoanthidium (Royanthidium) variabile</emphasis>
Pasteels, 1980
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFF29E314665F9D1EB67F902" author="Michener, Ch. D. &amp; Griswold, T. L." box="[771,1076,1654,1678]" pageId="30" pageNumber="31" pagination="299 - 327" refId="ref30709" refString="Michener, Ch. D. &amp; Griswold, T. L. (1994) The classification of Old World Anthidiini (Hymenoptera, Megachilidae). The University of Kansas Science Bulletin, 55, 299 - 327." type="journal article" year="1994">Michener &amp; Griswold (1994)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E0F45F1F91BEC69FF3D" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="871336ADFFF29E3145F1F91BEBBBF976" blockId="30.[151,1437,1724,1859]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145F1F91BEE3FF95A" box="[151,364,1724,1750]" italics="true" pageId="30" pageNumber="31">Material examined</emphasis>
.
<materialsCitation id="37C43CF0FFF29E31441DF91BED17F976" collectingDate="1977-10-10" collectionCode="NHMUK" collectorName="K. M. Guichard" country="Oman" latitude="17.22" location="Dhofar Ayun Pools" longLatPrecision="767" longitude="53.9" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Zufar" typeStatus="holotype">
<collectingCountry id="FFBB763DFFF29E31441DF91BEE86F95A" box="[379,469,1724,1750]" name="Oman" pageId="30" pageNumber="31">OMAN</collectingCountry>
:
<specimenCount id="91AAFD24FFF29E314487F91AEEA5F95B" box="[481,502,1725,1751]" count="1" pageId="30" pageNumber="31" type="female"></specimenCount>
<typeStatus id="5817880FFFF29E314498F91BED30F95A" box="[510,611,1724,1750]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
,
<location id="82736076FFF29E314709F91BEC15F95A" LSID="urn:lsid:plazi:treatment:0F0587BBFFF29E0F45F1FA49EDBCFDED:82736076FFF29E314709F91BEC15F95A" box="[623,838,1724,1750]" country="Oman" latitude="17.22" longLatPrecision="767" longitude="53.9" name="Dhofar Ayun Pools" pageId="30" pageNumber="31" stateProvince="Zufar">Dhofar Ayun Pools</location>
(
<geoCoordinate id="E298506AFFF29E314630F91BECEAF95A" box="[854,953,1724,1750]" degrees="17.22" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="17.22">17.22°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFF29E3146A3F91BEB77F95B" box="[965,1060,1724,1751]" degrees="53.90" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="53.9">53.90°E</geoCoordinate>
),
<date id="F312106DFFF29E31415FF91BEBF8F95A" box="[1081,1195,1724,1750]" pageId="30" pageNumber="31" value="1977-10-10">
<collectingDate id="E356E985FFF29E31415FF91BEBF8F95A" box="[1081,1195,1724,1750]" pageId="30" pageNumber="31" value="1977-10-10">10.x.1977</collectingDate>
</date>
,
<collectorName id="2A59537BFFF29E3141DEF91AEA38F95A" box="[1208,1387,1724,1751]" pageId="30" pageNumber="31">K. M. Guichard</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF29E3145F9F947EE5EF976" box="[159,269,1760,1786]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
, B.M. Type Hym 17a.3169)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFF29E314700F947EBB7F976" box="[614,1252,1760,1787]" collectingDate="1977-10-10" collectionCode="NHMUK" collectorName="K. M. Guichard &amp; B. M. Type Hym" country="Oman" latitude="17.22" location="OMAN" longLatPrecision="767" longitude="53.9" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Zufar" typeStatus="paratype">
<collectingCountry id="FFBB763DFFF29E314700F947ED90F976" box="[614,707,1760,1786]" name="Oman" pageId="30" pageNumber="31">OMAN</collectingCountry>
:
<specimenCount id="91AAFD24FFF29E3147A8F947EDA2F976" box="[718,753,1760,1786]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
<typeStatus id="5817880FFFF29E31479EF946EC08F977" box="[760,859,1761,1787]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
, same data as holotype (
<collectionCode id="E1BDAE68FFF29E31410AF947EB8EF976" box="[1132,1245,1760,1786]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFF29E3145A1F8A3ECAFF8CE" blockId="30.[151,1437,1724,1859]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1F8A3EE3DF892" box="[199,366,1796,1822]" italics="true" pageId="30" pageNumber="31">Other material</emphasis>
.
<specimenCount id="91AAFD24FFF29E31441AF8A3EECDF892" box="[380,414,1796,1822]" count="2" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
(incl.
<specimenCount id="91AAFD24FFF29E314480F8A2ED29F892" box="[486,634,1796,1823]" count="1" pageId="30" pageNumber="31" type="generic" typeStatus="paratype">one paratype</specimenCount>
),
<location id="82736076FFF29E3147EBF8A3ECDCF892" LSID="urn:lsid:plazi:treatment:0F0587BBFFF29E0F45F1FA49EDBCFDED:82736076FFF29E3147EBF8A3ECDCF892" box="[653,911,1796,1822]" latitude="17.38" longLatPrecision="767" longitude="54.05" municipality="North Slope" name="Qara Hills North Slope" pageId="30" pageNumber="31">
Qara Hills
<collectingMunicipality id="6777ACD7FFF29E314661F8A3ECDCF892" box="[775,911,1796,1822]" pageId="30" pageNumber="31">North Slope</collectingMunicipality>
</location>
[
<locationDeviation id="96A12F8FFFF29E3146FAF8A2EB7FF892" box="[924,1068,1796,1823]" location="Salalah" pageId="30" pageNumber="31">N of Salalah</locationDeviation>
, exact location not identified] (c.
<geoCoordinate id="E298506AFFF29E3145F1F88FEFABF8CE" box="[151,248,1832,1858]" degrees="17.38" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="17.38">17.38°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFF29E314462F88FEE31F8CF" box="[260,354,1832,1859]" degrees="54.05" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="54.05">54.05°E</geoCoordinate>
),
<date id="F312106DFFF29E314410F88FEEBAF8CE" box="[374,489,1832,1858]" pageId="30" pageNumber="31" value="1977-10-22">
<collectingDate id="E356E985FFF29E314410F88FEEBAF8CE" box="[374,489,1832,1858]" pageId="30" pageNumber="31" value="1977-10-22">22.x.1977</collectingDate>
</date>
,
<collectorName id="2A59537BFFF29E314492F88EEDF6F8CE" box="[500,677,1832,1859]" pageId="30" pageNumber="31">K. M. Guichard</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF29E314783F88FEC68F8CE" box="[741,827,1832,1858]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="30" pageNumber="31" type="Museum">RBINS</collectionCode>
) (
<collectorName id="2A59537BFFF29E314636F88FECFFF8CE" box="[848,940,1832,1858]" pageId="30" pageNumber="31">Pasteels</collectorName>
1980).
</paragraph>
<caption id="D3D36625FFF39E3045F1FBBFEAC0FBBD" ID-DOI="http://doi.org/10.5281/zenodo.14240769" ID-Zenodo-Dep="14240769" box="[151,1427,1048,1073]" httpUri="https://zenodo.org/record/14240769/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,245,1048,1072]" targetBox="[201,1386,151,1032]" targetPageId="31" targetType="figure">
<paragraph id="871336ADFFF39E3045F1FBBFEAC0FBBD" blockId="31.[151,1427,1048,1073]" box="[151,1427,1048,1073]" pageId="31" pageNumber="32">
<emphasis id="B5D8EABFFFF39E3045F1FBBFEE4AFBBC" bold="true" box="[151,281,1048,1072]" pageId="31" pageNumber="32">FIGURE 17.</emphasis>
<taxonomicName id="40AC4D2EFFF39E304479FBBFEDD4FBBC" authority="Kasparek" authorityName="Kasparek" authorityYear="2024" box="[287,647,1048,1072]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="syriacum" status="sp. nov.">
<emphasis id="B5D8EABFFFF39E304479FBBFED76FBBC" box="[287,549,1048,1072]" italics="true" pageId="31" pageNumber="32">Pseudoanthidium syriacum</emphasis>
Kasparek
</taxonomicName>
<emphasis id="B5D8EABFFFF39E3047EBFBBFED8AFBBC" bold="true" box="[653,729,1048,1072]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="AEEB57C4FFF39E3047EBFBBFED8AFBBC" box="[653,729,1048,1072]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female from Syria. a. Habitus dorsal; b. Habitus lateral; c. Apical terga.
</paragraph>
</caption>
<caption id="D3D36625FFF39E3045F1F809EEC1F866" ID-DOI="http://doi.org/10.5281/zenodo.14240771" ID-Zenodo-Dep="14240771" httpUri="https://zenodo.org/record/14240771/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1966,1990]" targetBox="[201,1385,1117,1943]" targetPageId="31" targetType="figure">
<paragraph id="871336ADFFF39E3045F1F809EEC1F866" blockId="31.[151,1436,1966,2026]" pageId="31" pageNumber="32">
<emphasis id="B5D8EABFFFF39E3045F1F809EE73F84A" bold="true" box="[151,288,1966,1990]" pageId="31" pageNumber="32">FIGURE 18.</emphasis>
<taxonomicName id="40AC4D2EFFF39E304441F809ED86F84A" authority="Kasparek" authorityName="Kasparek" authorityYear="2024" box="[295,725,1966,1990]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tajikistanicum" status="sp. nov.">
<emphasis id="B5D8EABFFFF39E304441F809ED3CF84A" box="[295,623,1966,1990]" italics="true" pageId="31" pageNumber="32">Pseudoanthidium tajikistanicum</emphasis>
Kasparek
</taxonomicName>
<emphasis id="B5D8EABFFFF39E3047BAF808EC7FF84B" bold="true" box="[732,812,1967,1991]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="AEEB57C4FFF39E3047BAF808EC7FF84B" box="[732,812,1967,1991]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female from Tajikistan. a. Habitus dorsal; b. Habitus lateral; c. Scutellum and axillae.
</paragraph>
</caption>
<paragraph id="871336ADFFCC9E0F45A1FF30EC69FF3D" blockId="32.[151,1437,151,609]" box="[199,826,151,177]" pageId="32" pageNumber="33">
<emphasis id="B5D8EABFFFCC9E0F45A1FF30ED58FF3D" box="[199,523,151,177]" italics="true" pageId="32" pageNumber="33">Genetic barcode information</emphasis>
. No information available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFCC9E0F45A1FF1CEED6FDB1" pageId="32" pageNumber="33" type="description">
<paragraph id="871336ADFFCC9E0F45A1FF1CEBE5FD95" blockId="32.[151,1437,151,609]" pageId="32" pageNumber="33">
<emphasis id="B5D8EABFFFCC9E0F45A1FF1CEE18FF58" box="[199,331,187,212]" italics="true" pageId="32" pageNumber="33">Description</emphasis>
:
<emphasis id="B5D8EABFFFCC9E0F4438FF1CEEE1FF59" box="[350,434,187,213]" italics="true" pageId="32" pageNumber="33">Female</emphasis>
(
<figureCitation id="1F972A28FFCC9E0F44A3FF1CED48FF59" box="[453,539,187,213]" captionStart="FIGURE 19" captionStartId="32.[151,250,1582,1606]" captionTargetBox="[151,1435,669,1558]" captionTargetId="figure-197@32.[151,1435,669,1558]" captionTargetPageId="32" captionText="FIGURE 19. Pseudoanthidium variabile (Pasteels, 1980). Female holotype from Oman. a. Habitus dorsal; b. Habitus lateral; c. Metasomal terga." figureDoi="http://doi.org/10.5281/zenodo.14240773" httpUri="https://zenodo.org/record/14240773/files/figure.png" pageId="32" pageNumber="33">Fig. 19</figureCitation>
).
<quantity id="40549B48FFCC9E0F4755FF1CED28FF59" box="[563,635,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="mm" value="7.0">7 mm</quantity>
. Colouration of maculations on the terga variable, ranging from redorange (
<typeStatus id="5817880FFFCC9E0F4593FF78EE7BFF75" box="[245,296,223,249]" pageId="32" pageNumber="33">type</typeStatus>
) to golden yellow.—
<emphasis id="B5D8EABFFFCC9E0F477AFF78ED0AFF75" box="[540,601,223,249]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
: Paraocular area, clypeus, preoccipital region, and gena orange; anterior edge of clypeus almost straight, with subapical region depressed; apical ridge brown, with five tubercles (
<figureCitation id="1F972A28FFCC9E0F405CFEA4EADCFE91" box="[1338,1423,259,285]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="32" pageNumber="33">Fig. 20</figureCitation>
); mandible yellow to orange-brown, featuring six more or less equally strong black teeth; postocello-preoccipital distance equal to interocellar distance.—
<taxonomicName id="40AC4D2EFFCC9E0F4700FEECED8DFEE8" box="[614,734,331,356]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFCC9E0F4700FEECED8DFEE8" box="[614,734,331,356]" italics="true" pageId="32" pageNumber="33">Mesosoma</emphasis>
</taxonomicName>
: Scutum with narrow orange anterolateral stripe; axilla and a broad band on posterior scutellum orange; scutellum and axillae crescent-shaped in dorsal view, without a median emargination.—
<emphasis id="B5D8EABFFFCC9E0F442BFE34EE91FE20" box="[333,450,403,428]" italics="true" pageId="32" pageNumber="33">Metasoma</emphasis>
: T1T6 orange with a black median wedge; tergal punctation noticeable finer than on scutum; T6 inversely trapezoid in dorsal view with median bilobed projection (
<figureCitation id="1F972A28FFCC9E0F417CFE10EB2DFE5D" box="[1050,1150,439,465]" captionStart="FIGURE 18" captionStartId="31.[151,250,1966,1990]" captionTargetBox="[201,1385,1117,1943]" captionTargetId="figure-37@31.[201,1385,1117,1943]" captionTargetPageId="31" captionText="FIGURE 18. Pseudoanthidium tajikistanicum Kasparek sp. nov. Female from Tajikistan. a. Habitus dorsal; b. Habitus lateral; c. Scutellum and axillae." figureDoi="http://doi.org/10.5281/zenodo.14240771" httpUri="https://zenodo.org/record/14240771/files/figure.png" pageId="32" pageNumber="33">Figs 18c</figureCitation>
,
<figureCitation id="1F972A28FFCC9E0F41EDFE10EBF8FE5D" box="[1163,1195,439,465]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="32" pageNumber="33">22</figureCitation>
); apical margin dark; scopa black with lighter hairs at the margin.—
<emphasis id="B5D8EABFFFCC9E0F47D0FE7CEDB8FE78" box="[694,747,475,500]" italics="true" pageId="32" pageNumber="33">Legs</emphasis>
: Trochanters and basal tibiae black; legs otherwise orange; orange bristles on inner face of hind basitarsus.—
<emphasis id="B5D8EABFFFCC9E0F47A2FE58EDADFD94" box="[708,766,511,536]" italics="true" pageId="32" pageNumber="33">Wing</emphasis>
: Brown infuscated, especially apically.
</paragraph>
<paragraph id="871336ADFFCC9E0F45A1FD84EED6FDB1" blockId="32.[151,1437,151,609]" box="[199,389,547,573]" pageId="32" pageNumber="33">
<emphasis id="B5D8EABFFFCC9E0F45A1FD84EE52FDB1" box="[199,257,547,573]" italics="true" pageId="32" pageNumber="33">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFCC9E0F45A1FDE0EDBCFDED" box="[199,751,583,609]" pageId="32" pageNumber="33" type="description">
<paragraph id="871336ADFFCC9E0F45A1FDE0EDBCFDED" blockId="32.[151,1437,151,609]" box="[199,751,583,609]" pageId="32" pageNumber="33">
<emphasis id="B5D8EABFFFCC9E0F45A1FDE0EE0BFDED" box="[199,344,583,609]" italics="true" pageId="32" pageNumber="33">Distribution.</emphasis>
Only known from
<collectingCountry id="FFBB763DFFCC9E0F4748FDE0ED21FDED" box="[558,626,583,609]" name="Oman" pageId="32" pageNumber="33">Oman</collectingCountry>
(
<figureCitation id="1F972A28FFCC9E0F47E7FDE0EDB0FDED" box="[641,739,583,609]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="32" pageNumber="33">Fig. 27a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,209 @@
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<mods:title id="2BB9EA4107CD8390E0DD91358B14312D">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<mods:identifier id="24EE33BF95CB2BF00487B6F868F0329C" type="ZooBank">A5D2633B-36A8-42A4-AD21-22041804BE95</mods:identifier>
</mods:mods>
<treatment id="0F0587BBFFF29E3145F1FE35EC49FAF5" ID-DOI="http://doi.org/10.5281/zenodo.14248305" ID-Zenodo-Dep="14248305" LSID="urn:lsid:plazi:treatment:0F0587BBFFF29E3145F1FE35EC49FAF5" httpUri="http://treatment.plazi.org/id/0F0587BBFFF29E3145F1FE35EC49FAF5" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="CFB66526FFF29E3145F1FE35EC54FE21" box="[151,775,402,429]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="871336ADFFF29E3145F1FE35EC54FE21" blockId="30.[151,775,402,465]" box="[151,775,402,429]" pageId="30" pageNumber="31">
<heading id="DC5B81C1FFF29E3145F1FE35EC54FE21" bold="true" box="[151,775,402,429]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="B5D8EABFFFF29E3145F1FE35EC54FE21" bold="true" box="[151,775,402,429]" pageId="30" pageNumber="31">
<taxonomicName id="40AC4D2EFFF29E3145F1FE35EDF1FE21" authority="Kasparek" authorityName="Kasparek" authorityYear="2024" box="[151,674,402,429]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tajikistanicum" status="sp. nov.">
<emphasis id="B5D8EABFFFF29E3145F1FE35ED72FE21" bold="true" box="[151,545,402,429]" italics="true" pageId="30" pageNumber="31">Pseudoanthidium tajikistanicum</emphasis>
Kasparek
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFF29E3147CFFE34EC54FE21" box="[681,775,403,429]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145F1FE10EE02FE5D" box="[151,337,439,465]" pageId="30" pageNumber="31" type="description">
<paragraph id="871336ADFFF29E3145F1FE10EE02FE5D" blockId="30.[151,775,402,465]" box="[151,337,439,465]" pageId="30" pageNumber="31">
(
<figureCitation id="1F972A28FFF29E3145F9FE10EFA5FE5D" box="[159,246,439,465]" captionStart="FIGURE 16" captionStartId="28.[151,250,1941,1965]" captionTargetBox="[151,1435,1124,1918]" captionTargetId="figure-60@28.[151,1435,1124,1918]" captionTargetPageId="28" captionText="FIGURE 16. Pseudoanthidium rubellulum Pasteels, 1969. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view." figureDoi="http://doi.org/10.5281/zenodo.14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="30" pageNumber="31">Figs 16</figureCitation>
,
<figureCitation id="1F972A28FFF29E314464FE10EE72FE5D" box="[258,289,439,465]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="30" pageNumber="31">20</figureCitation>
,
<figureCitation id="1F972A28FFF29E31444AFE10EE1AFE5D" box="[300,329,439,465]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="30" pageNumber="31">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145F1FE58ECA7FD41" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="871336ADFFF29E3145F1FE58EC06FD09" blockId="30.[151,1437,510,1401]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145F1FE58EE27FD95" box="[151,372,511,537]" italics="true" pageId="30" pageNumber="31">Material examined.</emphasis>
<materialsCitation id="37C43CF0FFF29E31441DFE58EDD0FDB1" collectingDate="2003-06-09" collectionCode="CMK" collectorName="E. Ivanova" country="Tajikistan" latitude="38.09" location="Chimbulkak" longLatPrecision="706" longitude="68.8" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Dushanbe" typeStatus="holotype">
<typeStatus id="5817880FFFF29E31441DFE58EEB5FD95" box="[379,486,511,537]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
,
<specimenCount id="91AAFD24FFF29E314495FE58ED11FD95" box="[499,578,511,537]" pageId="30" pageNumber="31" type="female">female</specimenCount>
,
<collectingCountry id="FFBB763DFFF29E31472BFE58EDBEFD95" box="[589,749,511,537]" name="Tajikistan" pageId="30" pageNumber="31">TAJIKISTAN</collectingCountry>
:
<locationDeviation id="96A12F8FFFF29E31479FFE59EC80FD95" box="[761,979,510,537]" pageId="30" pageNumber="31">
<quantity id="40549B48FFF29E31479FFE59EC12FD95" box="[761,833,510,537]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="50.0">50 km</quantity>
S Dushanbe
</locationDeviation>
,
<location id="82736076FFF29E3146B9FE58EB3AFD95" LSID="urn:lsid:plazi:treatment:0F0587BBFFF29E3145F1FE35EC49FAF5:82736076FFF29E3146B9FE58EB3AFD95" box="[991,1129,511,537]" country="Tajikistan" latitude="38.09" longLatPrecision="706" longitude="68.8" name="Chimbulkak" pageId="30" pageNumber="31" stateProvince="Dushanbe">Chimbulkak</location>
vill. (c.
<geoCoordinate id="E298506AFFF29E3141ACFE58EA7AFD95" box="[1226,1321,511,537]" degrees="38.09" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="38.09">38.09°N</geoCoordinate>
<geoCoordinate id="E298506AFFF29E314057FE58EAC3FD95" box="[1329,1424,511,537]" degrees="68.80" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="68.8">68.80°E</geoCoordinate>
),
<date id="F312106DFFF29E3145F1FD84EE41FDB1" box="[151,274,547,573]" pageId="30" pageNumber="31" value="2003-06-09">
<collectingDate id="E356E985FFF29E3145F1FD84EE41FDB1" box="[151,274,547,573]" pageId="30" pageNumber="31" value="2003-06-09">09.vi.2003</collectingDate>
</date>
,
<collectorName id="2A59537BFFF29E31447DFD84EEC0FDB1" box="[283,403,547,573]" pageId="30" pageNumber="31">E. Ivanova</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF29E3144A9FD84ED47FDB1" box="[463,532,547,573]" pageId="30" pageNumber="31">CMK</collectionCode>
: mk1548)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFF29E3147C2FD85ED77FDED" collectingDate="2003-06-09" collectionCode="CMK" collectorName="E. Ivanova" country="Tajikistan" latitude="38.09" location="Chimbulkak" longLatPrecision="706" longitude="68.8" pageId="30" pageNumber="31" specimenCount="55" stateProvince="Dushanbe" typeStatus="paratype">
<typeStatus id="5817880FFFF29E3147C2FD85EC44FDB1" box="[676,791,546,573]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
: 60♀ ex., same data as holotype (CMK: mk1538mk1547, mk1549mk1557) (
<specimenCount id="91AAFD24FFF29E314413FDE1EEE9FDED" box="[373,442,582,609]" count="55" pageId="30" pageNumber="31" type="generic">55 ex.</specimenCount>
in
<collectionCode id="E1BDAE68FFF29E3144BBFDE0ED73FDED" box="[477,544,583,609]" pageId="30" pageNumber="31">CMK</collectionCode>
;
</materialsCitation>
<materialsCitation id="37C43CF0FFF29E31474DFDE0ED9BFDED" box="[555,712,583,609]" collectingDate="2003-06-09" collectionCode="Simone Flaminio" collectorName="E. Ivanova" country="Tajikistan" latitude="38.09" location="Chimbulkak" longLatPrecision="706" longitude="68.8" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Dushanbe" typeStatus="paratype">
<specimenCount id="91AAFD24FFF29E31474DFDE0ED32FDED" box="[555,609,583,609]" count="1" pageId="30" pageNumber="31" type="generic">1 ex.</specimenCount>
in
<collectionCode id="E1BDAE68FFF29E3147E2FDE0ED97FDED" box="[644,708,583,609]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="30" pageNumber="31" type="Museum">ZMB</collectionCode>
;
</materialsCitation>
<materialsCitation id="37C43CF0FFF29E3147A8FDE0EB7BFDED" box="[718,1064,583,609]" collectingDate="2003-06-09" collectionCode="Simone Flaminio" collectorName="E. Ivanova" country="Tajikistan" latitude="38.09" location="Chimbulkak" longLatPrecision="706" longitude="68.8" pageId="30" pageNumber="31" specimenCount="3" stateProvince="Dushanbe" typeStatus="paratype">
<specimenCount id="91AAFD24FFF29E3147A8FDE0EC56FDED" box="[718,773,583,609]" count="3" pageId="30" pageNumber="31" type="generic">3 ex.</specimenCount>
in coll.
<collectionCode id="E1BDAE68FFF29E314639FDE0EB77FDED" box="[863,1060,583,609]" pageId="30" pageNumber="31">Simone Flaminio</collectionCode>
;
</materialsCitation>
<materialsCitation id="37C43CF0FFF29E314148FDE0EF83FD09" collectingDate="2003-06-09" collectionCode="A. Fateryga" collectorName="O. Pak" country="Tajikistan" latitude="38.09" location="Chimbulkak" longLatPrecision="706" longitude="68.8" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Dushanbe" typeStatus="paratype">
<specimenCount id="91AAFD24FFF29E314148FDE0EB36FDED" box="[1070,1125,583,609]" count="1" pageId="30" pageNumber="31" type="generic">1 ex.</specimenCount>
in coll.
<collectionCode id="E1BDAE68FFF29E3141DBFDE0EA12FDED" box="[1213,1345,583,609]" pageId="30" pageNumber="31">A. Fateryga</collectionCode>
(
<collectorName id="2A59537BFFF29E314036FDE0EACFFDED" box="[1360,1436,583,609]" pageId="30" pageNumber="31">O. Pak</collectorName>
leg.))
</materialsCitation>
.—
<typeStatus id="5817880FFFF29E314597FDCCEECEFD09" box="[241,413,619,645]" isOtherTypeStatus="true" pageId="30" pageNumber="31">Other material</typeStatus>
:
<specimenCount id="91AAFD24FFF29E3144C1FDCCEE99FD09" box="[423,458,619,645]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
, same data as holotype (damaged).
</paragraph>
<paragraph id="871336ADFFF29E3145A1FD28EC69FD25" blockId="30.[151,1437,510,1401]" box="[199,826,655,681]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FD28ED58FD25" box="[199,523,655,681]" italics="true" pageId="30" pageNumber="31">Genetic barcode information</emphasis>
. No information available.
</paragraph>
<paragraph id="871336ADFFF29E3145A1FD14ECA7FD41" blockId="30.[151,1437,510,1401]" box="[199,1012,691,717]" pageId="30" pageNumber="31">Efforts to extract DNA from the material for analysis were unsuccessful.</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FD70ED4EFCD1" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="871336ADFFF29E3145A1FD70ED4EFCD1" blockId="30.[151,1437,510,1401]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FD70EE6AFD7C" box="[199,313,727,752]" italics="true" pageId="30" pageNumber="31">Diagnosis</emphasis>
. Characterized by a relatively long scutellum, where the outer edge forms an inverted trapezoid with the outer edge of the axillae. This distinctive shape of the scutellum/axillae is not observed in any other Palaearctic
<taxonomicName id="40AC4D2EFFF29E3145F1FCB8EEEFFCB5" box="[151,444,799,825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B5D8EABFFFF29E3145F1FCB8EE08FCB5" box="[151,347,799,825]" italics="true" pageId="30" pageNumber="31">Pseudoanthidium</emphasis>
species.
</taxonomicName>
Other characteristic traits include coarse tergal punctation, akin to
<taxonomicName id="40AC4D2EFFF29E3141D1FCB8EA03FCB5" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[1207,1360,799,825]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF29E3141D1FCB8EA03FCB5" box="[1207,1360,799,825]" italics="true" pageId="30" pageNumber="31">P. puncticolle</emphasis>
</taxonomicName>
, and a narrow apical ridge of the clypeus.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FCC0EEC7FABD" pageId="30" pageNumber="31" type="description">
<paragraph id="871336ADFFF29E3145A1FCC0EC18FA81" blockId="30.[151,1437,510,1401]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FCC0EE18FC0C" box="[199,331,871,896]" italics="true" pageId="30" pageNumber="31">Description</emphasis>
.
<emphasis id="B5D8EABFFFF29E314431FCC0EEF8FC0D" box="[343,427,871,897]" italics="true" pageId="30" pageNumber="31">Female</emphasis>
(
<figureCitation id="1F972A28FFF29E3144DEFCC0ED5AFC0D" box="[440,521,871,897]" captionStart="FIGURE 18" captionStartId="31.[151,250,1966,1990]" captionTargetBox="[201,1385,1117,1943]" captionTargetId="figure-37@31.[201,1385,1117,1943]" captionTargetPageId="31" captionText="FIGURE 18. Pseudoanthidium tajikistanicum Kasparek sp. nov. Female from Tajikistan. a. Habitus dorsal; b. Habitus lateral; c. Scutellum and axillae." figureDoi="http://doi.org/10.5281/zenodo.14240771" httpUri="https://zenodo.org/record/14240771/files/figure.png" pageId="30" pageNumber="31">Fig. 18</figureCitation>
).
<quantity id="40549B48FFF29E31477DFCC0ED08FC0D" box="[539,603,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="31" unit="mm" value="9.0">9 mm</quantity>
.—
<emphasis id="B5D8EABFFFF29E31471AFCC0EDEAFC0D" box="[636,697,871,897]" italics="true" pageId="30" pageNumber="31">Head</emphasis>
: Black with elongated yellow spot behind the eye; lower paraocular area and occasionally also the lateral parts of the clypeus with a dirty yellow hue (
<figureCitation id="1F972A28FFF29E31414EFC2CEB2FFC29" box="[1064,1148,907,933]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="30" pageNumber="31">Fig. 20</figureCitation>
); anterior edge of clypeus somewhat arcuate inwards, almost straight; coarse punctation in the anterior and fine punctation in the posterior half of the clypeus; clypeal ridge protruding, shining; slightly concave, with an uneven surface of the ridge but lacking distinct tubercles.—
<taxonomicName id="40AC4D2EFFF29E31441FFC50EEA2FB9C" box="[377,497,1015,1040]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF29E31441FFC50EEA2FB9C" box="[377,497,1015,1040]" italics="true" pageId="30" pageNumber="31">Mesosoma</emphasis>
</taxonomicName>
: Scutum curved upward and shining; punctures spaced up to one puncture diameter apart; outer edge of scutellum and axillae forming an inverse trapezoid, with a broad pale yellow band on outer side and interrupted in the middle; surface polished, with punctures larger and further apart than those on scutum (
<figureCitation id="1F972A28FFF29E3145F9FBC4EE53FBF1" box="[159,256,1123,1149]" captionStart="FIGURE 18" captionStartId="31.[151,250,1966,1990]" captionTargetBox="[201,1385,1117,1943]" captionTargetId="figure-37@31.[201,1385,1117,1943]" captionTargetPageId="31" captionText="FIGURE 18. Pseudoanthidium tajikistanicum Kasparek sp. nov. Female from Tajikistan. a. Habitus dorsal; b. Habitus lateral; c. Scutellum and axillae." figureDoi="http://doi.org/10.5281/zenodo.14240771" httpUri="https://zenodo.org/record/14240771/files/figure.png" pageId="30" pageNumber="31">Fig. 18c</figureCitation>
); pronotal lobe lamellate.—
<emphasis id="B5D8EABFFFF29E31475CFBC4EDFCFBF0" box="[570,687,1123,1148]" italics="true" pageId="30" pageNumber="31">Metasoma</emphasis>
: Oval ivory spots present on T1T5; further apart from the middle on anterior terga than on posterior ones; punctation coarse, with punctures separated from each other up to two puncture diameters; T6 semi-trapezoid in dorsal view, with a grey median apical protrusion; punctation of T6 dense and rugulose (
<figureCitation id="1F972A28FFF29E31445DFB68EEC1FB65" box="[315,402,1231,1257]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
); S6 triangular; scopal hairs grey.—
<emphasis id="B5D8EABFFFF29E314657FB68EC35FB64" box="[817,870,1231,1256]" italics="true" pageId="30" pageNumber="31">Legs</emphasis>
: Trochanteres and basal femora black, remaining reddish brown; outer face of mid and hind basitarsi dark grey.
</paragraph>
<paragraph id="871336ADFFF29E3145A1FAB0EEC7FABD" blockId="30.[151,1437,510,1401]" box="[199,404,1303,1329]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FAB0EE52FABD" box="[199,257,1303,1329]" italics="true" pageId="30" pageNumber="31">Male</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FA9CEBEAFAD9" box="[199,1209,1339,1365]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="871336ADFFF29E3145A1FA9CEBEAFAD9" blockId="30.[151,1437,510,1401]" box="[199,1209,1339,1365]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FA9CEEC0FAD8" box="[199,403,1339,1364]" italics="true" pageId="30" pageNumber="31">Derivatio nominis</emphasis>
. Adjective. Named after
<collectingCountry id="FFBB763DFFF29E3147CDFA9CEC4FFAD9" box="[683,796,1339,1365]" name="Tajikistan" pageId="30" pageNumber="31">Tajikistan</collectingCountry>
, where the
<typeStatus id="5817880FFFF29E3146FAFA9CEC9FFAD9" box="[924,972,1339,1365]" pageId="30" pageNumber="31">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF29E3145A1FAF8EC49FAF5" box="[199,794,1375,1401]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="871336ADFFF29E3145A1FAF8EC49FAF5" blockId="30.[151,1437,510,1401]" box="[199,794,1375,1401]" pageId="30" pageNumber="31">
<emphasis id="B5D8EABFFFF29E3145A1FAF8EE03FAF5" box="[199,336,1375,1401]" italics="true" pageId="30" pageNumber="31">Distribution</emphasis>
: Only known from
<collectingCountry id="FFBB763DFFF29E314748FAF8EDCFFAF5" box="[558,668,1375,1401]" name="Tajikistan" pageId="30" pageNumber="31">Tajikistan</collectingCountry>
(
<figureCitation id="1F972A28FFF29E3147CDFAF8EC5DFAF5" box="[683,782,1375,1401]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="30" pageNumber="31">Fig. 26b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E2FECDE1CC84915837E6E480FF9F77F1" ID-DOI="10.11646/zootaxa.5541.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240734" ID-ZooBank="A5D2633B-36A8-42A4-AD21-22041804BE95" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732832365526" checkinUser="plazi" docAuthor="Kasparek, Max" docDate="2024" docId="0F0587BBFFF49E3645F1FB69EDC5F889" docLanguage="en" docName="zootaxa.5541.1.1.pdf" docOrigin="Zootaxa 5541 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5541.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoanthidium royoi" docType="treatment" docVersion="5" lastPageNumber="26" masterDocId="F33CFFC3FFEC9E2F4566FFA7EF53FF8C" masterDocTitle="New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="25" updateTime="1733329203150" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="40A6F7051722B67B6BB32109AC7C5077">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<mods:namePart id="4349CB77976C94FDD485CA5AA5FB1A63">Kasparek, Max</mods:namePart>
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<mods:date id="31AA5955E68A948E6D9F02058FF5C388">2024</mods:date>
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<treatment id="0F0587BBFFF49E3645F1FB69EDC5F889" LSID="urn:lsid:plazi:treatment:0F0587BBFFF49E3645F1FB69EDC5F889" httpUri="http://treatment.plazi.org/id/0F0587BBFFF49E3645F1FB69EDC5F889" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="CFB66526FFF49E3745F1FB69EC89FB65" box="[151,986,1230,1257]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="871336ADFFF49E3745F1FB69EC89FB65" blockId="24.[151,986,1230,1293]" box="[151,986,1230,1257]" pageId="24" pageNumber="25">
<heading id="DC5B81C1FFF49E3745F1FB69EC89FB65" bold="true" box="[151,986,1230,1257]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B5D8EABFFFF49E3745F1FB69EC89FB65" bold="true" box="[151,986,1230,1257]" pageId="24" pageNumber="25">
<taxonomicName id="40AC4D2EFFF49E3745F1FB69ED25FB64" ID-CoL="8SRFZ" authority="(Dusmet, 1915)" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[151,630,1230,1257]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF49E3745F1FB69EEE1FB65" bold="true" box="[151,434,1230,1257]" italics="true" pageId="24" pageNumber="25">Pseudoanthidium royoi</emphasis>
(Dusmet, 1915)
</taxonomicName>
stat. resurrect &amp; comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745F1FB54EE9CFA81" box="[151,463,1267,1293]" pageId="24" pageNumber="25" type="description">
<paragraph id="871336ADFFF49E3745F1FB54EE9CFA81" blockId="24.[151,986,1230,1293]" box="[151,463,1267,1293]" pageId="24" pageNumber="25">
(
<figureCitation id="1F972A28FFF49E3745F9FB54EFA5FA81" box="[159,246,1267,1293]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="24" pageNumber="25">Figs 13</figureCitation>
,
<figureCitation id="1F972A28FFF49E374464FB54EE72FA81" box="[258,289,1267,1293]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="24" pageNumber="25">20</figureCitation>
,
<figureCitation id="1F972A28FFF49E37444AFB54EE19FA81" box="[300,330,1267,1293]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="24" pageNumber="25">21</figureCitation>
,
<figureCitation id="1F972A28FFF49E374430FB54EE26FA81" box="[342,373,1267,1293]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="24" pageNumber="25">22</figureCitation>
,
<figureCitation id="1F972A28FFF49E3744E6FB54EECDFA81" box="[384,414,1267,1293]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="24" pageNumber="25">23</figureCitation>
,
<figureCitation id="1F972A28FFF49E3744CCFB54EE94FA81" box="[426,455,1267,1293]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="24" pageNumber="25">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745F1FA90EC2CFA01" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="871336ADFFF49E3745F1FA90ED81FAC3" blockId="24.[151,895,1335,1421]" box="[151,722,1335,1359]" pageId="24" pageNumber="25">
<taxonomicName id="40AC4D2EFFF49E3745F1FA90EE80FAC3" authority="Dusmet, 1915" authorityName="Dusmet" authorityYear="1915" box="[151,467,1335,1359]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF49E3745F1FA90EE6FFAC3" box="[151,316,1335,1359]" italics="true" pageId="24" pageNumber="25">Anthidium royoi</emphasis>
Dusmet, 1915
</taxonomicName>
. Male, female,
<collectingCountry id="FFBB763DFFF49E374709FA9FED9DFAC3" box="[623,718,1336,1359]" name="Morocco" pageId="24" pageNumber="25">Morocco</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFF49E3745F1FAF1ED9BFAE2" blockId="24.[151,895,1335,1421]" box="[151,712,1366,1390]" pageId="24" pageNumber="25">
<taxonomicName id="40AC4D2EFFF49E3745F1FAF1ED50FAE2" authority="(Dusmet, 1915)" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[151,515,1366,1390]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF49E3745F1FAF1EE32FAE1" box="[151,353,1366,1390]" italics="true" pageId="24" pageNumber="25">Royanthidium royoi</emphasis>
(Dusmet, 1915)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFF49E374744FAF1ED97FAE2" author="Pasteels, J. J." box="[546,708,1366,1390]" pageId="24" pageNumber="25" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFF49E3745F1FAD2EC2CFA01" blockId="24.[151,895,1335,1421]" box="[151,895,1396,1421]" pageId="24" pageNumber="25">
non-
<taxonomicName id="40AC4D2EFFF49E3745A0FAD3ED72FA01" authority="Klug, 1832" authorityName="Klug" authorityYear="1832" box="[198,545,1396,1421]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF49E3745A0FAD3EEFAFA00" box="[198,425,1396,1420]" italics="true" pageId="24" pageNumber="25">Anthidium melanurum</emphasis>
Klug, 1832
</taxonomicName>
.—Synonymy by
<bibRefCitation id="E33D4B5CFFF49E3747B2FAD3EC28FA01" author="Warncke, K." box="[724,891,1396,1421]" pageId="24" pageNumber="25" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745F1FA1CECA0F905" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="871336ADFFF49E3745F1FA1CED37FA75" blockId="24.[151,1437,1466,2033]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745F1FA1CEE25FA59" box="[151,374,1467,1493]" italics="true" pageId="24" pageNumber="25">Material examined:</emphasis>
<collectingCountry id="FFBB763DFFF49E374418FA1CED43FA59" box="[382,528,1467,1493]" name="Morocco" pageId="24" pageNumber="25">MOROCCO</collectingCountry>
:
<specimenCount id="91AAFD24FFF49E37477DFA1CED6DFA59" box="[539,574,1467,1493]" count="1" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
,
<specimenCount id="91AAFD24FFF49E37472BFA1CED22FA59" box="[589,625,1467,1493]" count="1" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
,
<location id="82736076FFF49E374719FA1CEDBFFA59" LSID="urn:lsid:plazi:treatment:0F0587BBFFF49E3645F1FB69EDC5F889:82736076FFF49E374719FA1CEDBFFA59" box="[639,748,1467,1493]" country="Morocco" latitude="31.491667" longLatPrecision="20" longitude="9.7502775" name="Essaouira" pageId="24" pageNumber="25">Essaouira</location>
(
<geoCoordinate id="E298506AFFF49E37479AFA1CECDFFA59" box="[764,908,1467,1493]" degrees="31" direction="north" minutes="29" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="30" value="31.491667">31°2930N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFF49E3746C4FA1DEB7CFA59" box="[930,1071,1466,1493]" degrees="09" direction="east" minutes="45" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="01" value="9.7502775">09°4501E</geoCoordinate>
),
<quantity id="40549B48FFF49E374123FA1CEBD2FA59" box="[1093,1153,1467,1493]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="m" value="18.0">
<elevation id="0C81D19EFFF49E374123FA1CEBD2FA59" box="[1093,1153,1467,1493]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="m" value="18.0">18 m</elevation>
</quantity>
,
<date id="F312106DFFF49E3741E8FA1DEA55FA59" box="[1166,1286,1466,1493]" pageId="24" pageNumber="25" value="2015-04-28">
<collectingDate id="E356E985FFF49E3741E8FA1DEA55FA59" box="[1166,1286,1466,1493]" pageId="24" pageNumber="25" value="2015-04-28">28.iv.2015</collectingDate>
</date>
,
<collectorName id="2A59537BFFF49E374074FA1CEA38FA59" box="[1298,1387,1467,1493]" pageId="24" pageNumber="25">V. Soon</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF49E3745F9FA78EF88FA75" box="[159,219,1503,1529]" pageId="24" pageNumber="25">TUZ</collectionCode>
:
<specimenCode id="D70A9ED6FFF49E374583FA78EE21FA75" box="[229,370,1503,1529]" collectionCode="TUZ" pageId="24" pageNumber="25">TUZ036633</specimenCode>
, -6647;
<specimenCode id="D70A9ED6FFF49E3744ABFA78ED0BFA75" box="[461,600,1503,1529]" collectionCode="TUZ" pageId="24" pageNumber="25">TUZ037-38</specimenCode>
).
</paragraph>
<paragraph id="871336ADFFF49E3745A1F9A4EA02F991" blockId="24.[151,1437,1466,2033]" box="[199,1361,1538,1565]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1F9A4EE24F991" box="[199,375,1539,1565]" italics="true" pageId="24" pageNumber="25">Other material:</emphasis>
<collectingCountry id="FFBB763DFFF49E374419F9A4ED42F991" box="[383,529,1539,1565]" name="Morocco" pageId="24" pageNumber="25">MOROCCO</collectingCountry>
:
<specimenCount id="91AAFD24FFF49E37477DF9A4ED16F991" box="[539,581,1539,1565]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
,
<specimenCount id="91AAFD24FFF49E374735F9A4ED2AF991" box="[595,633,1539,1565]" count="2" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
,
<location id="82736076FFF49E3747E2F9A4EC65F991" LSID="urn:lsid:plazi:treatment:0F0587BBFFF49E3645F1FB69EDC5F889:82736076FFF49E3747E2F9A4EC65F991" box="[644,822,1539,1565]" country="Morocco" latitude="31.49" longLatPrecision="729" longitude="9.78" name="Mogador Island" pageId="24" pageNumber="25">Mogador Island</location>
(
<geoCoordinate id="E298506AFFF49E374623F9A4ECF6F991" box="[837,933,1539,1565]" degrees="31.49" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="31.49">31.49°N</geoCoordinate>
-
<geoCoordinate id="E298506AFFF49E3746D2F9A4EB46F991" box="[948,1045,1539,1565]" degrees="09.78" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="555" value="9.78">09.78°E</geoCoordinate>
),
<date id="F312106DFFF49E37414EF9A4EBC9F991" box="[1064,1178,1539,1565]" pageId="24" pageNumber="25" value="1907-05">
<collectingDate id="E356E985FFF49E37414EF9A4EBC9F991" box="[1064,1178,1539,1565]" pageId="24" pageNumber="25" value="1907-05">May 1907</collectingDate>
</date>
(Dusmet 1915).
</paragraph>
<paragraph id="871336ADFFF49E3745A1F980ECA0F905" blockId="24.[151,1437,1466,2033]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1F980ED46F9CD" box="[199,533,1575,1601]" italics="true" pageId="24" pageNumber="25">Genetic barcode information.</emphasis>
The DNA sequence of the barcoding gene of a female and a male was obtained from material collected from a location close to the
<typeStatus id="5817880FFFF49E374788F9ECEC4DF9E9" box="[750,798,1611,1637]" pageId="24" pageNumber="25">type</typeStatus>
locality in
<collectingCountry id="FFBB763DFFF49E3746C5F9ECEB5BF9E9" box="[931,1032,1611,1637]" name="Morocco" pageId="24" pageNumber="25">Morocco</collectingCountry>
(
<specimenCode id="D70A9ED6FFF49E37417FF9ECEB9DF9E9" box="[1049,1230,1611,1637]" collectionCode="ABABY" pageId="24" pageNumber="25">ABABY004-23</specimenCode>
, ABABY005-23). They were assigned to BOLD:ACF9972, the same BIN as for
<emphasis id="B5D8EABFFFF49E374637F9C8ECA0F905" box="[849,1011,1647,1673]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="40AC4D2EFFF49E374637F9C8ECBCF905" authorityName="Klug" authorityYear="1832" box="[849,1007,1647,1673]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="melanurum">P. melanurum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745A1F934EACAF895" pageId="24" pageNumber="25" type="discussion">
<paragraph id="871336ADFFF49E3745A1F934EACAF895" blockId="24.[151,1437,1466,2033]" pageId="24" pageNumber="25">
In a species ID tree as inferred from the barcoding gene,
<taxonomicName id="40AC4D2EFFF49E37460CF934EC91F920" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[874,962,1683,1708]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF49E37460CF934EC91F920" box="[874,962,1683,1708]" italics="true" pageId="24" pageNumber="25">P. royoi</emphasis>
</taxonomicName>
emerged as s distinct group, sister to
<taxonomicName id="40AC4D2EFFF49E3740E2F934EE46F95D" authorityName="Klug" authorityYear="1832" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF49E3740E2F934EE46F95D" italics="true" pageId="24" pageNumber="25">A. melanurum</emphasis>
</taxonomicName>
(
<figureCitation id="1F972A28FFF49E374440F910EE28F95D" box="[294,379,1719,1745]" captionStart="FIGURE 14" captionStartId="27.[151,250,1328,1352]" captionTargetBox="[151,1436,156,1315]" captionTargetId="graphics-22@27.[156,1095,160,1301]" captionText="FIGURE 14. Maximum Likelihood species ID tree as inferred from the DNA sequence of the mitochondrial cytochrome c oxidase subunit 1 barcoding gene. The tree shows a selection of three closely related species (A. melanurum, A. rhombiferum, and A. royoi), rooted on P. octodentatum as outgroup." figureDoi="http://doi.org/10.5281/zenodo.14275816" httpUri="https://zenodo.org/record/14275816/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
). The ML bootstrap value for the node separating these two groups (based on 1000 bootstrap replicates) was 100% (
<figureCitation id="1F972A28FFF49E3744FEF97CEEBEF979" box="[408,493,1755,1781]" captionStart="FIGURE 14" captionStartId="27.[151,250,1328,1352]" captionTargetBox="[151,1436,156,1315]" captionTargetId="graphics-22@27.[156,1095,160,1301]" captionText="FIGURE 14. Maximum Likelihood species ID tree as inferred from the DNA sequence of the mitochondrial cytochrome c oxidase subunit 1 barcoding gene. The tree shows a selection of three closely related species (A. melanurum, A. rhombiferum, and A. royoi), rooted on P. octodentatum as outgroup." figureDoi="http://doi.org/10.5281/zenodo.14275816" httpUri="https://zenodo.org/record/14275816/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
). For
<taxonomicName id="40AC4D2EFFF49E374754F97CEDDBF978" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[562,648,1755,1780]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF49E374754F97CEDDBF978" box="[562,648,1755,1780]" italics="true" pageId="24" pageNumber="25">P. royoi</emphasis>
</taxonomicName>
, the intraspecfic distance was 0% (identical haplotypes), while it was 0.07% for
<taxonomicName id="40AC4D2EFFF49E37446BF958EEF5F895" authorityName="Klug" authorityYear="1832" box="[269,422,1791,1817]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF49E37446BF958EEF5F895" box="[269,422,1791,1817]" italics="true" pageId="24" pageNumber="25">P. melanurum</emphasis>
</taxonomicName>
. The mean genetic distance between these two species was 2.27%, the barcode gap 2.18%.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3645A1F884EDE0FCB5" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="871336ADFFF49E3645A1F884EE4DFEE9" blockId="24.[151,1437,1466,2033]" lastBlockId="25.[151,1437,151,1797]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1F884EE18F8B0" box="[199,331,1827,1852]" italics="true" pageId="24" pageNumber="25">Description</emphasis>
.
<emphasis id="B5D8EABFFFF49E37443AF884EEE3F8B1" box="[348,432,1827,1853]" italics="true" pageId="24" pageNumber="25">Female</emphasis>
(
<figureCitation id="1F972A28FFF49E3744A7F884ED70F8B1" box="[449,547,1827,1853]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="24" pageNumber="25">Fig. 13a</figureCitation>
).
<quantity id="40549B48FFF49E37475FF885EDCCF8B1" box="[569,671,1826,1853]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="24" pageNumber="25" unit="mm" value="11.5">11.5 mm</quantity>
(
<quantity id="40549B48FFF49E3747D6F884EC51F8B1" box="[688,770,1827,1853]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="mm" value="12.0">12 mm</quantity>
for
<typeStatus id="5817880FFFF49E374652F884EC37F8B1" box="[820,868,1827,1853]" pageId="24" pageNumber="25">type</typeStatus>
specimen (Dusmet 1915)).—
<emphasis id="B5D8EABFFFF49E3741DEF884EBA6F8B1" box="[1208,1269,1827,1853]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
: Clypeus with strongly protruding apical ridge; ridge lip-like, black, with six strong tubercles; punctation of clypeus basally scattered, denser and finer toward the apex (
<figureCitation id="1F972A28FFF49E3747F3F8CCEDBFF809" box="[661,748,1899,1925]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="24" pageNumber="25">Fig. 20</figureCitation>
); mandible short, dark orange, with four obtuse black teeth; clypeus and lower paraocular area orange-red; broad orange-red preoccipital band reaching the middle of the eye; golden-red pubescence especially on frons, preoccipital area and gena.—
<emphasis id="B5D8EABFFFF49E3746A3F814EB71F841" box="[965,1058,1971,1997]" italics="true" pageId="24" pageNumber="25">Antenna</emphasis>
: sg1 (scape) and sg3sg6 orange-red, sg2 black, sg7 orange-brown bicoloured, sg8sg13 brown.—
<taxonomicName id="40AC4D2EFFF49E3746E7F870ECAAF87C" box="[897,1017,2007,2032]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF49E3746E7F870ECAAF87C" box="[897,1017,2007,2032]" italics="true" pageId="24" pageNumber="25">Mesosoma</emphasis>
</taxonomicName>
: Scutum with dense, fine punctation with thin interstices (
<figureCitation id="1F972A28FFF59E3644E6FF30EEB3FF3D" box="[384,480,151,177]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="25" pageNumber="26">Fig. 13d</figureCitation>
); mesepisternum with minute orange spot; pronotal lobe orange-red with high anterior lamella; scutellum slightly swollen, widely rounded in dorsal view with a shallow median emargination of apical margin; axilla protruding laterally; axilla and scutellum posterolaterally orange-red.—
<emphasis id="B5D8EABFFFF59E36412AFF78EB92FF74" box="[1100,1217,223,248]" italics="true" pageId="25" pageNumber="26">Metasoma</emphasis>
: T1T4 with broad, lateral orange-red bands that come closer in the middle distally; T5 with inconspicuous, dark median stripe; T6 orange-red, with broad shoulders in dorsal view, and a bi-lobed median protrusion (
<figureCitation id="1F972A28FFF59E364124FE80EBC4FECD" box="[1090,1175,295,321]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="25" pageNumber="26">Fig. 22</figureCitation>
); scopa yellow.—
<emphasis id="B5D8EABFFFF59E364006FE80EAC6FECC" box="[1376,1429,295,320]" italics="true" pageId="25" pageNumber="26">Legs</emphasis>
: Orange-red.
</paragraph>
<paragraph id="871336ADFFF59E3645A1FEC8EDE0FCB5" blockId="25.[151,1437,151,1797]" pageId="25" pageNumber="26">
<emphasis id="B5D8EABFFFF59E3645A1FEC8EE52FE05" box="[199,257,367,393]" italics="true" pageId="25" pageNumber="26">Male</emphasis>
(
<figureCitation id="1F972A28FFF59E364470FEC8EE2DFE05" box="[278,382,367,393]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="25" pageNumber="26">Fig. 13b</figureCitation>
).
<quantity id="40549B48FFF59E3644FFFEC9ED51FE05" box="[409,514,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="25" pageNumber="26" unit="mm" value="11.5">11.5 mm</quantity>
(
<quantity id="40549B48FFF59E364771FEC8EDC5FE05" box="[535,662,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="25" pageNumber="26" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
for
<typeStatus id="5817880FFFF59E3647B7FEC8EC52FE05" box="[721,769,367,393]" pageId="25" pageNumber="26">type</typeStatus>
specimens (Dusmet 1915)).—
<emphasis id="B5D8EABFFFF59E36410AFEC8EBFAFE05" box="[1132,1193,367,393]" italics="true" pageId="25" pageNumber="26">Head</emphasis>
: Clypeus and lower paraocular area dark yellow; orange-red preoccipital band, broader behind the eye than in the middle (
<figureCitation id="1F972A28FFF59E36405EFE34EADDFE21" box="[1336,1422,403,429]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
); mandible short, black at base and reddish medially; three black teeth (strong apical tooth and two smaller blunt teeth); punctation of clypeus fine, scattered at base and dense apically; apical margin black and crenulated.—
<emphasis id="B5D8EABFFFF59E3645F1FE58EFA7FD95" box="[151,244,511,537]" italics="true" pageId="25" pageNumber="26">Antenna</emphasis>
: Scape (sg1) orange-red, sg2 black, sg3sg5 orange-red, sg6sg13 black.—
<taxonomicName id="40AC4D2EFFF59E364130FE58EB9DFD94" box="[1110,1230,511,536]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF59E364130FE58EB9DFD94" box="[1110,1230,511,536]" italics="true" pageId="25" pageNumber="26">Mesosoma</emphasis>
</taxonomicName>
: Shape of scutum as in the female; scutellum black, axilla with minute orange-red spot; pronotal lobal dark brown with high lamella as in female.—
<emphasis id="B5D8EABFFFF59E364423FDE0EEE9FDEC" box="[325,442,583,608]" italics="true" pageId="25" pageNumber="26">Metasoma</emphasis>
: T1T5 similar to the female with broad orange-red lateral bands, closer to the middle posteriorly than anteriorly; T6 orange-red with inconspicuous longitudinal dark midline; T6 laterally bulged; apical margin with a strong, semi-transparent brownish lamella on each side of the middle; T7 orange-red with a deep median emargination (as deep as broad); lateral margin angular (
<figureCitation id="1F972A28FFF59E36460FFD14ECECFD41" box="[873,959,691,717]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="25" pageNumber="26">Figs 23</figureCitation>
,
<figureCitation id="1F972A28FFF59E3646ADFD14ECB8FD41" box="[971,1003,691,717]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="25" pageNumber="26">24</figureCitation>
). S1 with a strong median longitudinal carina; apical margin of S4 shallowly emarginate; S6 with truncate apex, lateral margin sharp-edged; central part shining, with hairs laterally (
<figureCitation id="1F972A28FFF59E36448FFD5CED1AFC99" box="[489,585,763,789]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="25" pageNumber="26">Fig. 13e</figureCitation>
).—
<emphasis id="B5D8EABFFFF59E364710FD5CEDB3FC99" box="[630,736,763,789]" italics="true" pageId="25" pageNumber="26">Genitalia</emphasis>
: Gonostyli broadly paddle-shaped, somewhat twisted; penis valve strong with acute, arcuate apex (
<figureCitation id="1F972A28FFF59E364720FCB8EDF6FCB5" box="[582,677,799,825]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[152,1436,182,1918]" captionTargetId="figure-19@26.[152,1436,181,1918]" captionTargetPageId="26" captionText="FIGURE 13. Pseudoanthidium royoi (Dusmet, 1915) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Habitus lateral, male; d. Dorsal side of mesosoma, male; e. Apical sterna of the male." figureDoi="http://doi.org/10.5281/zenodo.14240763" httpUri="https://zenodo.org/record/14240763/files/figure.png" pageId="25" pageNumber="26">Fig. 13e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF59E3645A1FCE4EE27FB2D" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="871336ADFFF59E3645A1FCE4EE27FB2D" blockId="25.[151,1437,151,1797]" pageId="25" pageNumber="26">
<emphasis id="B5D8EABFFFF59E3645A1FCE4EEE9FCD1" box="[199,442,835,861]" italics="true" pageId="25" pageNumber="26">Differential diagnosis</emphasis>
.
<taxonomicName id="40AC4D2EFFF59E3644AEFCE4ED9EFCD0" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[456,717,835,861]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E3644AEFCE4ED9EFCD0" box="[456,717,835,861]" italics="true" pageId="25" pageNumber="26">Pseudoanthidium royoi</emphasis>
</taxonomicName>
is structurally very similar to
<taxonomicName id="40AC4D2EFFF59E36417DFCE4EBE5FCD1" authorityName="Klug" authorityYear="1832" box="[1051,1206,835,861]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E36417DFCE4EBE5FCD1" box="[1051,1206,835,861]" italics="true" pageId="25" pageNumber="26">P. melanurum</emphasis>
</taxonomicName>
, but is distinguished by its colour, colouring pattern and morphometric features.
<taxonomicName id="40AC4D2EFFF59E36462BFCC0EB06FC0C" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[845,1109,871,897]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E36462BFCC0EB06FC0C" box="[845,1109,871,897]" italics="true" pageId="25" pageNumber="26">Pseudoanthidium royoi</emphasis>
</taxonomicName>
is significantly more robust and larger than
<taxonomicName id="40AC4D2EFFF59E364420FC2CEEB1FC29" authorityName="Klug" authorityYear="1832" box="[326,482,907,933]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E364420FC2CEEB1FC29" box="[326,482,907,933]" italics="true" pageId="25" pageNumber="26">P. melanurum</emphasis>
</taxonomicName>
. No size overlap was found in the length of the marginal cell of the wing, which was used as an indicator for body size (
<figureCitation id="1F972A28FFF59E364741FC08ED2FFC45" box="[551,636,942,969]" captionStart="FIGURE 15" captionStartId="28.[151,250,1003,1027]" captionTargetBox="[243,1344,191,967]" captionTargetId="figure-19@28.[230,1357,181,980]" captionTargetPageId="28" captionText="FIGURE 15. Comparison of body size of Pseudoanthidium royoi and P. melanurum, as expressed by the marginal cell length of the wing. Pseudoanthidium melanurum: female N= 21, male N= 31; P. royoi N=1 for each sex." figureDoi="http://doi.org/10.5281/zenodo.14240765" httpUri="https://zenodo.org/record/14240765/files/figure.png" pageId="25" pageNumber="26">Fig. 15</figureCitation>
). All body maculations are orange-red instead of yellow, and hairs are golden-yellow instead of dirty-white. The female is further characterized by a broad orange-yellow preoccipital band reaching almost to the lower end of the eye, while
<taxonomicName id="40AC4D2EFFF59E364669FC50ECFCFB9D" authorityName="Klug" authorityYear="1832" box="[783,943,1015,1041]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E364669FC50ECFCFB9D" box="[783,943,1015,1041]" italics="true" pageId="25" pageNumber="26">A. melanurum</emphasis>
</taxonomicName>
has only yellow spots behind the upper end of the eye. The males are further distinguished by a broad orange-red preoccipitial band extending between the upper end of the eye (only yellow spots in
<taxonomicName id="40AC4D2EFFF59E3647EFFB98EC79FBD5" authorityName="Klug" authorityYear="1832" box="[649,810,1087,1113]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E3647EFFB98EC79FBD5" box="[649,810,1087,1113]" italics="true" pageId="25" pageNumber="26">A. melanurum</emphasis>
</taxonomicName>
). The antennal scape is light orange-yellow in male
<taxonomicName id="40AC4D2EFFF59E3640EEFB98EF82FBF0" baseAuthorityName="Dusmet" baseAuthorityYear="1915" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E3640EEFB98EF82FBF0" italics="true" pageId="25" pageNumber="26">P. royoi</emphasis>
</taxonomicName>
, while it is black in
<taxonomicName id="40AC4D2EFFF59E3644D8FBC4ED09FBF1" authorityName="Klug" authorityYear="1832" box="[446,602,1123,1149]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E3644D8FBC4ED09FBF1" box="[446,602,1123,1149]" italics="true" pageId="25" pageNumber="26">P. melanurum</emphasis>
</taxonomicName>
. Furthermore, the two species are distinguished by the DNA sequence of the barcoding gene.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF59E3645A1FB0CECE2FA11" pageId="25" pageNumber="26" type="discussion">
<paragraph id="871336ADFFF59E3645A1FB0CECE2FA11" blockId="25.[151,1437,151,1797]" pageId="25" pageNumber="26">
<emphasis id="B5D8EABFFFF59E3645A1FB0CEE2AFB48" box="[199,377,1195,1220]" italics="true" pageId="25" pageNumber="26">Taxonomic note</emphasis>
. Dusmet (1915) described
<taxonomicName id="40AC4D2EFFF59E3647C9FB0CEC35FB48" authorityName="Dusmet" authorityYear="1915" box="[687,870,1195,1221]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E3647C9FB0CEC35FB48" box="[687,870,1195,1221]" italics="true" pageId="25" pageNumber="26">Anthidium royoi</emphasis>
</taxonomicName>
based on a female and
<specimenCount id="91AAFD24FFF59E364112FB0CEBBBFB49" box="[1140,1256,1195,1221]" count="2" pageId="25" pageNumber="26" type="male">two males</specimenCount>
from the island of
<collectingCountry id="FFBB763DFFF59E3645D1FB68EE4DFB65" box="[183,286,1231,1257]" name="Morocco" pageId="25" pageNumber="26">Mogador</collectingCountry>
in south-western
<collectingCountry id="FFBB763DFFF59E36448DFB68ED07FB65" box="[491,596,1231,1257]" name="Morocco" pageId="25" pageNumber="26">Morocco</collectingCountry>
. He compared these specimens with striking orange-red maculations with
<taxonomicName id="40AC4D2EFFF59E3645F1FB54ED2FFA81" authority="(Fabricius, 1787)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[151,636,1267,1293]" class="Insecta" family="Megachilidae" genus="Rhodanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sticticum">
<emphasis id="B5D8EABFFFF59E3645F1FB54EEE3FA80" box="[151,432,1267,1293]" italics="true" pageId="25" pageNumber="26">Rhodanthidium sticticum</emphasis>
(Fabricius, 1787)
</taxonomicName>
. Therefore, it is surprising that
<bibRefCitation id="E33D4B5CFFF59E3646BBFB54EBC1FA81" author="Warncke, K." box="[989,1170,1267,1293]" pageId="25" pageNumber="26" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
noted that he could not recognize differences between
<taxonomicName id="40AC4D2EFFF59E36449FFAB0ED07FABC" authorityName="Dusmet" authorityYear="1915" box="[505,596,1303,1329]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E36449FFAB0ED07FABC" box="[505,596,1303,1329]" italics="true" pageId="25" pageNumber="26">A. royoi</emphasis>
</taxonomicName>
and the black and yellow
<taxonomicName id="40AC4D2EFFF59E3646E1FAB0EB7BFABD" authorityName="Klug" authorityYear="1832" box="[903,1064,1303,1329]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E3646E1FAB0EB7BFABD" box="[903,1064,1303,1329]" italics="true" pageId="25" pageNumber="26">A. melanurum</emphasis>
</taxonomicName>
, and regarded
<taxonomicName id="40AC4D2EFFF59E3641B2FAB0EA63FABC" authorityName="Dusmet" authorityYear="1915" box="[1236,1328,1303,1329]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E3641B2FAB0EA63FABC" box="[1236,1328,1303,1329]" italics="true" pageId="25" pageNumber="26">A. royoi</emphasis>
</taxonomicName>
as junior synonym of
<taxonomicName id="40AC4D2EFFF59E364440FA9CEE94FAD9" authorityName="Klug" authorityYear="1832" box="[294,455,1339,1365]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E364440FA9CEE94FAD9" box="[294,455,1339,1365]" italics="true" pageId="25" pageNumber="26">A. melanurum</emphasis>
</taxonomicName>
.
<bibRefCitation id="E33D4B5CFFF59E3644B0FA9CEDC7FAD9" author="Kasparek, M." box="[470,660,1339,1365]" pageId="25" pageNumber="26" refId="ref29929" refString="Kasparek, M. (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe and Western Turkey. Identification. Distribution. Biology. Chimaira, Frankfurt, 292 pp. [unchanged reprint 2023]" type="book" year="2022">Kasparek (2022)</bibRefCitation>
noted that it is difficult to recognise a similarity between this taxon and
<taxonomicName id="40AC4D2EFFF59E3645A1FAF8EE34FAF5" authorityName="Klug" authorityYear="1832" box="[199,359,1375,1401]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E3645A1FAF8EE34FAF5" box="[199,359,1375,1401]" italics="true" pageId="25" pageNumber="26">A. melanurum</emphasis>
</taxonomicName>
from Dusmets description. New material from
<collectingCountry id="FFBB763DFFF59E3646E3FAF8ECB9FAF5" box="[901,1002,1375,1401]" name="Morocco" pageId="25" pageNumber="26">Morocco</collectingCountry>
has now made it possible to recognise
<taxonomicName id="40AC4D2EFFF59E3645F1FA24EE1EFA10" authorityName="Dusmet" authorityYear="1915" box="[151,333,1411,1437]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E3645F1FA24EE1EFA10" box="[151,333,1411,1437]" italics="true" pageId="25" pageNumber="26">Anthidium royoi</emphasis>
</taxonomicName>
as a valid taxon and to transfer it to
<taxonomicName id="40AC4D2EFFF59E364780FA24ECF9FA11" authorityName="Friese" authorityYear="1898" box="[742,938,1411,1437]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF59E364780FA24ECF9FA11" box="[742,938,1411,1437]" italics="true" pageId="25" pageNumber="26">Pseudoanthidium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF59E3645A1FA00EE42F9A1" pageId="25" pageNumber="26" type="distribution">
<paragraph id="871336ADFFF59E3645A1FA00EE42F9A1" blockId="25.[151,1437,151,1797]" pageId="25" pageNumber="26">
<emphasis id="B5D8EABFFFF59E3645A1FA00EE03FA4D" box="[199,336,1447,1473]" italics="true" pageId="25" pageNumber="26">Distribution</emphasis>
. Endemic to
<collectingCountry id="FFBB763DFFF59E364489FA00ED07FA4D" box="[495,596,1447,1473]" name="Morocco" pageId="25" pageNumber="26">Morocco</collectingCountry>
(
<figureCitation id="1F972A28FFF59E364701FA00ED98FA4D" box="[615,715,1447,1473]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="25" pageNumber="26">Fig. 26a</figureCitation>
). Only known from the Moroccan Atlantic island of
<collectingCountry id="FFBB763DFFF59E364050FA00EACEFA4D" box="[1334,1437,1447,1473]" name="Morocco" pageId="25" pageNumber="26">Mogador</collectingCountry>
<quantity id="40549B48FFF59E3645F1FA6DEFB5FA69" box="[151,230,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="km" value="1.5">1.5 km</quantity>
off the coast of
<collectingCountry id="FFBB763DFFF59E3644C6FA6CED5EFA69" box="[416,525,1483,1509]" name="Morocco" pageId="25" pageNumber="26">Essaouira</collectingCountry>
(
<geoCoordinate id="E298506AFFF59E36477BFA6BED2CFA6B" box="[541,639,1484,1511]" degrees="31" direction="north" minutes="29" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="31.483334">31°29N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFF59E3647F5FA6BEDB0FA6B" box="[659,739,1484,1511]" degrees="9" direction="east" minutes="47" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="9.783334">9°47E</geoCoordinate>
), from where it was collected in 1907 (Dusmet 1915) and the nearby mainland of Essaoiura (
<geoCoordinate id="E298506AFFF59E364490FA48EDD5F985" box="[502,646,1519,1545]" degrees="31" direction="north" minutes="29" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="30" value="31.491667">31°2930N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFF59E3647FDFA49EC7BF985" box="[667,808,1518,1545]" degrees="09" direction="east" minutes="45" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="01" value="9.7502775">09°4501E</geoCoordinate>
), where V. Soon rediscovered it in 2015, more than 100 years later.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF59E3645A1F990EDC5F889" pageId="25" pageNumber="26" type="discussion">
<paragraph id="871336ADFFF59E3645A1F990EDC5F889" blockId="25.[151,1437,151,1797]" pageId="25" pageNumber="26">
<emphasis id="B5D8EABFFFF59E3645A1F990EDE7F9DC" box="[199,692,1591,1617]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="40AC4D2EFFF59E3645A1F990ED44F9DD" ID-CoL="4NP4L" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[199,535,1591,1617]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" sensu="stricto" species="melanurum">Pseudoanthidium melanurum</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFF59E364744F990EDE7F9DC" box="[546,692,1591,1616]" pageId="25" pageNumber="26" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
(i.e. based on records other than
<taxonomicName id="40AC4D2EFFF59E36412EF990EBF0F9DC" ID-CoL="8SRFZ" authorityName="Dusmet" authorityYear="1915" box="[1096,1187,1591,1616]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E36412EF990EBF0F9DC" box="[1096,1187,1591,1616]" italics="true" pageId="25" pageNumber="26">P. royoi</emphasis>
</taxonomicName>
) has so far not been recorded in
<collectingCountry id="FFBB763DFFF59E364447F9FCEED9F9F9" box="[289,394,1627,1653]" name="Morocco" pageId="25" pageNumber="26">Morocco</collectingCountry>
. Paglianios (1994) mention of a personal record lacks supporting detail, and the reference to the Marrakesh-Safi administrative region by
<bibRefCitation id="E33D4B5CFFF59E3647D1F9D8ECE0F915" author="Lhomme, P. &amp; Michez, D. &amp; Christmann, S. &amp; Scheuchl, E. &amp; El Abdouni, I. &amp; Hamroud, L. &amp; Ihsane, O. &amp; Sentil, A. &amp; Smaili, M. C. &amp; Schwarz, M. &amp; Dathe, H. &amp; Straka, J. &amp; Pauly, A. &amp; Schmid-Egger, Ch. &amp; Patiny, S. &amp; Muller, A. &amp; Praz, Ch. &amp; Risch, S. &amp; Kasparek, M. &amp; Kuhlmann, M. &amp; Wood, Th. J. &amp; Bogusch, P. &amp; Ascher, J. &amp; Rasmont, R." box="[695,947,1663,1689]" pageId="25" pageNumber="26" pagination="1 - 159" refId="ref30246" refString="Lhomme, P., Michez, D., Christmann, S., Scheuchl, E., El Abdouni, I., Hamroud, L., Ihsane, O., Sentil, A., Smaili, M. C., Schwarz, M., Dathe, H., Straka, J., Pauly, A., Schmid-Egger, Ch., Patiny, S., Muller, A., Praz, Ch., Risch, S., Kasparek, M., Kuhlmann, M., Wood, Th. J., Bogusch, P., Ascher, J. &amp; Rasmont, R. (2020) The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa, 4892 (1), 1 - 159. https: // doi. org / 10.11646 / zootaxa. 4892.1.1" type="journal article" year="2020">
Lhomme
<emphasis id="B5D8EABFFFF59E364643F9D8EC33F915" box="[805,864,1663,1689]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
(2020)
</bibRefCitation>
is based on Dusmet (1915) (e.g.
<taxonomicName id="40AC4D2EFFF59E364052F9D8EADFF914" ID-CoL="8SRFZ" authorityName="Dusmet" authorityYear="1915" box="[1332,1420,1663,1688]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E364052F9D8EADFF914" box="[1332,1420,1663,1688]" italics="true" pageId="25" pageNumber="26">P. royoi</emphasis>
</taxonomicName>
). The location of a historical record of
<taxonomicName id="40AC4D2EFFF59E36472EF904EDB5F931" ID-CoL="4NP4L" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[584,742,1699,1725]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E36472EF904EDB5F931" box="[584,742,1699,1725]" italics="true" pageId="25" pageNumber="26">P. melanurum</emphasis>
</taxonomicName>
(as
<taxonomicName id="40AC4D2EFFF59E36467EF904EB75F931" ID-CoL="8SPDF" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[792,1062,1698,1725]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="dilobum">
<emphasis id="B5D8EABFFFF59E36467EF904ECC6F931" box="[792,917,1699,1725]" italics="true" pageId="25" pageNumber="26">A. dilobum</emphasis>
<bibRefCitation id="E33D4B5CFFF59E3646F8F904EB75F931" author="Perez, J." box="[926,1062,1698,1725]" pageId="25" pageNumber="26" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
</taxonomicName>
) reported from
<collectingCountry id="FFBB763DFFF59E3641BAF904EA63F931" box="[1244,1328,1699,1725]" name="Algeria" pageId="25" pageNumber="26">Algeria</collectingCountry>
has been questioned by
<bibRefCitation id="E33D4B5CFFF59E36445BF960EEA5F96D" author="Warncke, K." box="[317,502,1735,1761]" pageId="25" pageNumber="26" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
. The nearest population of
<taxonomicName id="40AC4D2EFFF59E364649F960EC9EF96D" ID-CoL="4NP4L" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[815,973,1735,1761]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF59E364649F960EC9EF96D" box="[815,973,1735,1761]" italics="true" pageId="25" pageNumber="26">P. melanurum</emphasis>
</taxonomicName>
thus seems to be in
<collectingCountry id="FFBB763DFFF59E3641DCF960EBADF96D" box="[1210,1278,1735,1761]" name="Georgia" pageId="25" pageNumber="26">Iberia</collectingCountry>
, with
<taxonomicName id="40AC4D2EFFF59E364023F960EACFF96C" ID-CoL="8SRFZ" authorityName="Dusmet" authorityYear="1915" box="[1349,1436,1735,1760]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF59E364023F960EACFF96C" box="[1349,1436,1735,1760]" italics="true" pageId="25" pageNumber="26">P. royoi</emphasis>
</taxonomicName>
appearing to represent an isolated population.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,263 @@
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<subSubSection id="CFB66526FFF79E3445F1FA78EDE3FA75" box="[151,688,1503,1530]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="871336ADFFF79E3445F1FA78EDE3FA75" blockId="27.[151,688,1503,1565]" box="[151,688,1503,1530]" pageId="27" pageNumber="28">
<heading id="DC5B81C1FFF79E3445F1FA78EDE3FA75" bold="true" box="[151,688,1503,1530]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<taxonomicName id="40AC4D2EFFF79E3445F1FA78EDE3FA75" ID-CoL="8SNNT" authority="Pasteels, 1969" authorityName="Pasteels" authorityYear="1969" box="[151,688,1503,1530]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rubellulum">
<emphasis id="B5D8EABFFFF79E3445F1FA78EDE3FA75" bold="true" box="[151,688,1503,1530]" pageId="27" pageNumber="28">
<emphasis id="B5D8EABFFFF79E3445F1FA78EEA8FA76" bold="true" box="[151,507,1503,1530]" italics="true" pageId="27" pageNumber="28">Pseudoanthidium rubellulum</emphasis>
<bibRefCitation id="E33D4B5CFFF79E344765FA47EDE3FA75" author="Pasteels, J. J." box="[515,688,1503,1530]" pageId="27" pageNumber="28" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF79E3445F1F9A4EFA9F991" box="[151,250,1539,1565]" pageId="27" pageNumber="28" type="description">
<paragraph id="871336ADFFF79E3445F1F9A4EFA9F991" blockId="27.[151,688,1503,1565]" box="[151,250,1539,1565]" pageId="27" pageNumber="28">
(
<figureCitation id="1F972A28FFF79E3445F9F9A4EFA2F991" box="[159,241,1539,1565]" captionStart="FIGURE 16" captionStartId="28.[151,250,1941,1965]" captionTargetBox="[151,1435,1124,1918]" captionTargetId="figure-60@28.[151,1435,1124,1918]" captionTargetPageId="28" captionText="FIGURE 16. Pseudoanthidium rubellulum Pasteels, 1969. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view." figureDoi="http://doi.org/10.5281/zenodo.14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="27" pageNumber="28">Fig. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF79E3445F1F9E6ED2DF9F4" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="871336ADFFF79E3445F1F9E6EDB4F9D5" blockId="27.[151,743,1601,1656]" box="[151,743,1601,1625]" pageId="27" pageNumber="28">
<taxonomicName id="40AC4D2EFFF79E3445F1F9E6EDB0F9D5" ID-CoL="8SNNT" authority="Pasteels, 1969" authorityName="Pasteels" authorityYear="1969" box="[151,739,1601,1625]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rubellulum" subGenus="Carinellum">
<emphasis id="B5D8EABFFFF79E3445F1F9E6ED1BF9D5" box="[151,584,1601,1625]" italics="true" pageId="27" pageNumber="28">Pseudoanthidium (Carinellum) rubellulum</emphasis>
<bibRefCitation id="E33D4B5CFFF79E344729F9E6EDB0F9D5" author="Pasteels, J. J." box="[591,739,1601,1625]" pageId="27" pageNumber="28" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels, 1969</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="871336ADFFF79E3445F1F9C7ED2DF9F4" blockId="27.[151,743,1601,1656]" box="[151,638,1632,1656]" pageId="27" pageNumber="28">
<taxonomicName id="40AC4D2EFFF79E3445F1F9C7ED76F9F4" ID-CoL="8SNWQ" authority="Alfken, 1933" authorityName="Alfken" authorityYear="1933" box="[151,549,1632,1656]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFF79E3445F1F9C7EEC9F9F4" box="[151,410,1632,1656]" italics="true" pageId="27" pageNumber="28">Anthidium ochrognathum</emphasis>
Alfken, 1933
</taxonomicName>
(partim)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF79E3445F1F938EC16F951" pageId="27" pageNumber="28" type="discussion">
<paragraph id="871336ADFFF79E3445F1F938EC16F951" blockId="27.[151,1437,1695,2043]" pageId="27" pageNumber="28">
The species status of this taxon has recently been re-established by Kasparek
<emphasis id="B5D8EABFFFF79E344175F907EB1EF935" box="[1043,1101,1695,1721]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
(2023). In the meantime, the hitherto undescribed male was found and is described herein.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF79E3445A1F940EE37F877" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="871336ADFFF79E3445A1F940EC57F8A9" blockId="27.[151,1437,1695,2043]" pageId="27" pageNumber="28">
<emphasis id="B5D8EABFFFF79E3445A1F940EECDF88D" box="[199,414,1767,1793]" italics="true" pageId="27" pageNumber="28">Material examined</emphasis>
(new material not listed in Kasparek
<emphasis id="B5D8EABFFFF79E344635F94FECDEF88D" box="[851,909,1767,1793]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2023):
<collectingCountry id="FFBB763DFFF79E34468FF940EB1CF88D" box="[1001,1103,1767,1793]" name="Israel" pageId="27" pageNumber="28">ISRAEL</collectingCountry>
:
<specimenCount id="91AAFD24FFF79E34413AF940EBDCF88D" box="[1116,1167,1767,1793]" count="83" pageId="27" pageNumber="28" type="female">83♀</specimenCount>
,
<specimenCount id="91AAFD24FFF79E3441FBF940EB91F88D" box="[1181,1218,1767,1793]" count="8" pageId="27" pageNumber="28" type="male">8♂</specimenCount>
,
<quantity id="40549B48FFF79E3441B7F940EA49F88D" box="[1233,1306,1767,1793]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="km" value="10.0">10 km</quantity>
N
<location id="82736076FFF79E344026F940EACFF88D" LSID="urn:lsid:plazi:treatment:0F0587BBFFF79E3245F1FA78EC26FBD5:82736076FFF79E344026F940EACFF88D" box="[1344,1436,1767,1793]" country="Israel" latitude="30.94" longLatPrecision="731" longitude="34.4" name="Nizzana" pageId="27" pageNumber="28">Nizzana</location>
(
<geoCoordinate id="E298506AFFF79E3445F9F8ACEFACF8A9" box="[159,255,1803,1829]" degrees="30.94" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="30.94">30.94°N</geoCoordinate>
<geoCoordinate id="E298506AFFF79E344460F8ACEE36F8A9" box="[262,357,1803,1829]" degrees="34.40" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="34.4">34.40°E</geoCoordinate>
),
<date id="F312106DFFF79E34441FF8ADEEA2F8A8" box="[377,497,1802,1828]" pageId="27" pageNumber="28" value="2023-04-16">
<collectingDate id="E356E985FFF79E34441FF8ADEEA2F8A8" box="[377,497,1802,1828]" pageId="27" pageNumber="28" value="2023-04-16">16.iv.2023</collectingDate>
</date>
,
<collectorName id="2A59537BFFF79E34449BF8ACED21F8A8" box="[509,626,1802,1829]" pageId="27" pageNumber="28">M. Halada</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF79E3447D4F8ACEDABF8A9" box="[690,760,1803,1829]" pageId="27" pageNumber="28">CMK</collectionCode>
).
</paragraph>
<paragraph id="871336ADFFF79E3445A1F888EE37F877" blockId="27.[151,1437,1695,2043]" pageId="27" pageNumber="28">
<emphasis id="B5D8EABFFFF79E3445A1F888ED4DF8C4" box="[199,542,1838,1865]" italics="true" pageId="27" pageNumber="28">Genetic barcode information.</emphasis>
COI sequences were obtained for
<specimenCount id="91AAFD24FFF79E3446A4F888EB1CF8C4" box="[962,1103,1838,1865]" count="2" pageId="27" pageNumber="28" type="female">two females</specimenCount>
(one sequence published in Kasparek
<emphasis id="B5D8EABFFFF79E344461F8F4EE6CF8E0" box="[263,319,1874,1900]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2023; one published for the first time here: ABABX473-22 and ABABZ022-23) and
<specimenCount id="91AAFD24FFF79E34419EF8F5EA0EF8E0" box="[1272,1373,1874,1900]" count="1" pageId="27" pageNumber="28" type="male">one male</specimenCount>
(new here: ABABZ021-23). The sequence of the male was almost identical to that of the female documented in Kasparek
<emphasis id="B5D8EABFFFF79E3445F1F83DEF82F83F" box="[151,209,1945,1971]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
(2023), thereby confirming the identity of the male of this species. The barcoding gene was represented in these
<specimenCount id="91AAFD24FFF79E3445BDF81AEEC1F85B" box="[219,402,1981,2007]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
by two different haplotypes, resulting in an intragroup distance of 0.0024%. The BIN is BOLD:AEO7128.
</paragraph>
</subSubSection>
<caption id="D3D36625FFF09E3345F1FC4CEB25FBAB" ID-DOI="http://doi.org/10.5281/zenodo.14240765" ID-Zenodo-Dep="14240765" httpUri="https://zenodo.org/record/14240765/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1003,1027]" targetBox="[243,1344,191,967]" targetPageId="28" targetType="figure">
<paragraph id="871336ADFFF09E3345F1FC4CEB25FBAB" blockId="28.[151,1436,1003,1064]" pageId="28" pageNumber="29">
<emphasis id="B5D8EABFFFF09E3345F1FC4CEE73FB88" bold="true" box="[151,288,1003,1028]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
Comparison of body size of
<taxonomicName id="40AC4D2EFFF09E33472CFC4BEC64FB8F" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[586,823,1004,1028]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF09E33472CFC4BEC64FB8F" box="[586,823,1004,1028]" italics="true" pageId="28" pageNumber="29">Pseudoanthidium royoi</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF09E33460CFC4BECA5FB88" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[874,1014,1004,1028]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF09E33460CFC4BECA5FB88" box="[874,1014,1004,1028]" italics="true" pageId="28" pageNumber="29">P. melanurum</emphasis>
</taxonomicName>
, as expressed by the marginal cell length of the wing.
<taxonomicName id="40AC4D2EFFF09E334471FBB7ED11FBA4" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[279,578,1040,1064]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFF09E334471FBB7ED11FBA4" box="[279,578,1040,1064]" italics="true" pageId="28" pageNumber="29">Pseudoanthidium melanurum</emphasis>
</taxonomicName>
: female N= 21, male N= 31;
<taxonomicName id="40AC4D2EFFF09E33460BFBB7ECE8FBAB" baseAuthorityName="Dusmet" baseAuthorityYear="1915" box="[877,955,1040,1063]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="royoi">
<emphasis id="B5D8EABFFFF09E33460BFBB7ECE8FBAB" box="[877,955,1040,1063]" italics="true" pageId="28" pageNumber="29">P. royoi</emphasis>
</taxonomicName>
N=1 for each sex.
</paragraph>
</caption>
<caption id="D3D36625FFF09E3345F1F832ED06F85D" ID-DOI="http://doi.org/10.5281/zenodo.14240767" ID-Zenodo-Dep="14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1941,1965]" targetBox="[151,1435,1124,1918]" targetPageId="28" targetType="figure">
<paragraph id="871336ADFFF09E3345F1F832ED06F85D" blockId="28.[151,1437,1941,2001]" pageId="28" pageNumber="29">
<emphasis id="B5D8EABFFFF09E3345F1F832EE71F821" bold="true" box="[151,290,1941,1965]" pageId="28" pageNumber="29">FIGURE 16.</emphasis>
<taxonomicName id="40AC4D2EFFF09E33444DF832EDA7F821" authority="Pasteels, 1969" authorityName="Pasteels" authorityYear="1969" box="[299,756,1941,1965]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rubellulum">
<emphasis id="B5D8EABFFFF09E33444DF832ED07F821" box="[299,596,1941,1965]" italics="true" pageId="28" pageNumber="29">Pseudoanthidium rubellulum</emphasis>
<bibRefCitation id="E33D4B5CFFF09E33473BF832EDA7F821" author="Pasteels, J. J." box="[605,756,1941,1965]" pageId="28" pageNumber="29" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels, 1969</bibRefCitation>
</taxonomicName>
. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view.
</paragraph>
</caption>
<subSubSection id="CFB66526FFF19E3245A1FF30EC1CFE91" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="871336ADFFF19E3245A1FF30EC1CFE91" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FF30EE6AFF3C" box="[199,313,151,176]" italics="true" pageId="29" pageNumber="30">Diagnosis</emphasis>
(male). Omaulus lamellate to lower end and pronotal lobe with high lamella as in
<taxonomicName id="40AC4D2EFFF19E3241B8FF30EACFFF3D" authorityName="Alfken" authorityYear="1933" box="[1246,1436,151,177]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFF19E3241B8FF30EACFFF3D" box="[1246,1436,151,177]" italics="true" pageId="29" pageNumber="30">P. ochrognathum</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF19E3245AFFF1CEE0EFF59" baseAuthorityName="Benoist" baseAuthorityYear="1950" box="[201,349,187,213]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="B5D8EABFFFF19E3245AFFF1CEE0EFF59" box="[201,349,187,213]" italics="true" pageId="29" pageNumber="30">P. beaumonti</emphasis>
</taxonomicName>
; inner edge of malar area rounded (carinate in
<taxonomicName id="40AC4D2EFFF19E3246E5FF1CEB10FF59" authorityName="Alfken" authorityYear="1933" box="[899,1091,187,213]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFF19E3246E5FF1CEB10FF59" box="[899,1091,187,213]" italics="true" pageId="29" pageNumber="30">P. ochrognathum</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF19E324119FF1CEA40FF59" baseAuthorityName="Benoist" baseAuthorityYear="1950" box="[1151,1299,187,213]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="B5D8EABFFFF19E324119FF1CEA40FF59" box="[1151,1299,187,213]" italics="true" pageId="29" pageNumber="30">P. beaumonti</emphasis>
</taxonomicName>
); mid-basal area of the clypeus impunctate to shallowly scattered punctate (clypeus evenly punctate in
<taxonomicName id="40AC4D2EFFF19E3241F5FF78EA03FF75" authorityName="Alfken" authorityYear="1933" box="[1171,1360,223,249]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFF19E3241F5FF78EA03FF75" box="[1171,1360,223,249]" italics="true" pageId="29" pageNumber="30">P. ochrognathum</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF19E3240EEFF78EE5EFE91" baseAuthorityName="Benoist" baseAuthorityYear="1950" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="B5D8EABFFFF19E3240EEFF78EE5EFE91" italics="true" pageId="29" pageNumber="30">P. beaumonti</emphasis>
</taxonomicName>
); depressions of T1T5 brownish bronze-coloured.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245A1FE80EDCCFC29" pageId="29" pageNumber="30" type="description">
<paragraph id="871336ADFFF19E3245A1FE80EDBEFECD" blockId="29.[151,1437,151,1113]" box="[199,749,295,321]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FE80EE18FECC" box="[199,331,295,320]" italics="true" pageId="29" pageNumber="30">Description</emphasis>
.
<emphasis id="B5D8EABFFFF19E32443FFE80EEFEFECD" box="[345,429,295,321]" italics="true" pageId="29" pageNumber="30">Female</emphasis>
. See Kasparek
<emphasis id="B5D8EABFFFF19E32473DFE80EDC7FECD" box="[603,660,295,321]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
(2023).
</paragraph>
<paragraph id="871336ADFFF19E3245A1FEECEDCCFC29" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FEECEE52FEE9" box="[199,257,331,357]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
(
<figureCitation id="1F972A28FFF19E324477FEECEEE3FEE9" box="[273,432,331,357]" captionStart="FIGURE 16" captionStartId="28.[151,250,1941,1965]" captionTargetBox="[151,1435,1124,1918]" captionTargetId="figure-60@28.[151,1435,1124,1918]" captionTargetPageId="28" captionText="FIGURE 16. Pseudoanthidium rubellulum Pasteels, 1969. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view." figureDoi="http://doi.org/10.5281/zenodo.14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="29" pageNumber="30">Figs 16a, 16b</figureCitation>
).
<quantity id="40549B48FFF19E3244A3FEEDED4DFEE9" box="[453,542,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="29" pageNumber="30" unit="mm" value="5.5">5.5 mm</quantity>
.—
<emphasis id="B5D8EABFFFF19E324759FEECED2FFEE9" box="[575,636,331,357]" italics="true" pageId="29" pageNumber="30">Head</emphasis>
: Black, clypeus and a postocular spot pale ivory-yellow; occasionally, a spot also present on the lower paraocular area; marginal areas of clypeus sometimes irregularly black; mid-basal area of the clypeus impunctate to shallowly scattered punctate; apical part of clypeus slightly curved inwards (
<figureCitation id="1F972A28FFF19E3245F9FE10EE56FE5D" box="[159,261,439,465]" captionStart="FIGURE 16" captionStartId="28.[151,250,1941,1965]" captionTargetBox="[151,1435,1124,1918]" captionTargetId="figure-60@28.[151,1435,1124,1918]" captionTargetPageId="28" captionText="FIGURE 16. Pseudoanthidium rubellulum Pasteels, 1969. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view." figureDoi="http://doi.org/10.5281/zenodo.14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="29" pageNumber="30">Fig. 16d</figureCitation>
); mandible pale yellow, with three strong black teeth; transitional area towards teeth brown; antenna light brown, except for sg1sg3, which are dark; long white pubescence particularly around antennal sockets, also covering clypeus.—
<taxonomicName id="40AC4D2EFFF19E32441CFE58EEA1FD94" box="[378,498,511,536]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF19E32441CFE58EEA1FD94" box="[378,498,511,536]" italics="true" pageId="29" pageNumber="30">Mesosoma</emphasis>
</taxonomicName>
: Scutum mostly black, with a narrow anterior yellow stripe observed in 2 out of
<specimenCount id="91AAFD24FFF19E3240E8FE58EE5CFDB1" count="8" pageId="29" pageNumber="30" type="generic">8 specimens</specimenCount>
; shining, with fine punctation, where punctures are mostly 12 puncture diameters apart; posterior margin of scutellum and axilla sharp-edged; punctation often less regular than on scutum; scutellum and axilla occasionally with yellow marginal stripe, but often absent; pronotal lobe with high antero-lateral lamella.—
<emphasis id="B5D8EABFFFF19E324183FDCCEA09FD08" box="[1253,1370,619,644]" italics="true" pageId="29" pageNumber="30">Metasoma</emphasis>
: Pale yellow lateral spot on T1; elongated spots on T2T5, not reaching the middle; T6 black anteriorly and reddish brown posteriorly; T7 light yellow-brown with shallow apical emargination; depressions and smooth, impunctate apical margins of T1T6 brownish bronze-coloured. S3 with apically hooked and corkscrew-like hairs; S5 with a pair of small black combs on the apical margin (approximately 6.5 comb widths apart from each other) and a pair of larger combs at the apex of lateral arms (
<figureCitation id="1F972A28FFF19E324738FCB8EDE8FCB5" box="[606,699,799,825]" captionStart="FIGURE 16" captionStartId="28.[151,250,1941,1965]" captionTargetBox="[151,1435,1124,1918]" captionTargetId="figure-60@28.[151,1435,1124,1918]" captionTargetPageId="28" captionText="FIGURE 16. Pseudoanthidium rubellulum Pasteels, 1969. Male from Israel. a. Habitus dorsal; b. Habitus lateral; c. Apical sterna from ventral; d. Head in frontal view." figureDoi="http://doi.org/10.5281/zenodo.14240767" httpUri="https://zenodo.org/record/14240767/files/figure.png" pageId="29" pageNumber="30">Fig. 16c</figureCitation>
).—
<emphasis id="B5D8EABFFFF19E324781FCB8EC4FFCB4" box="[743,796,799,824]" italics="true" pageId="29" pageNumber="30">Legs</emphasis>
: Femora reddish-brown apically, tibiae and tarsi reddish-brown, sometimes with yellow, brown or black spots.—
<emphasis id="B5D8EABFFFF19E32466BFCE4EC24FCD1" box="[781,887,835,861]" italics="true" pageId="29" pageNumber="30">Genitalia</emphasis>
: Male genitalia very similar to
<taxonomicName id="40AC4D2EFFF19E3241BCFCE4EA3FFCD1" baseAuthorityName="Benoist" baseAuthorityYear="1950" box="[1242,1388,835,861]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="B5D8EABFFFF19E3241BCFCE4EA3FFCD1" box="[1242,1388,835,861]" italics="true" pageId="29" pageNumber="30">P. beaumonti</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF19E3245F1FCC0EE07FC0D" authorityName="Alfken" authorityYear="1933" box="[151,340,871,897]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ochrognathum">
<emphasis id="B5D8EABFFFF19E3245F1FCC0EE07FC0D" box="[151,340,871,897]" italics="true" pageId="29" pageNumber="30">P. ochrognathum</emphasis>
</taxonomicName>
(see fig.
<quantity id="40549B48FFF19E3244DAFCC1EEB5FC0D" box="[444,486,870,897]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="30" unit="in" value="5.0">5 in</quantity>
Kasparek
<emphasis id="B5D8EABFFFF19E32473BFCC0EDC6FC0D" box="[605,661,871,897]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2023), with long, apically unnotched gonostyli; penis valves unfused and at some distance apart, touching at the tip.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245A1FC08EED3FBB9" pageId="29" pageNumber="30" type="discussion">
<paragraph id="871336ADFFF19E3245A1FC08EED3FBB9" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FC08EE7AFC45" box="[199,297,943,969]" italics="true" pageId="29" pageNumber="30">Remarks</emphasis>
: There is a strong colour dimorphism: While females are characterised by orange-red and red-brown maculation, males have pale yellow (ivory-coloured) maculae. The affiliation of the two sexes to the same species was confirmed by genetic barcoding (see above). Such sexual colour dimorphism is not known from other members of
<taxonomicName id="40AC4D2EFFF19E3245D0FBBCEE29FBB9" authorityName="Friese" authorityYear="1898" box="[182,378,1051,1077]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF19E3245D0FBBCEE29FBB9" box="[182,378,1051,1077]" italics="true" pageId="29" pageNumber="30">Pseudoanthidium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF19E3245A1FB98EC26FBD5" box="[199,885,1087,1113]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="871336ADFFF19E3245A1FB98EC26FBD5" blockId="29.[151,1437,151,1113]" box="[199,885,1087,1113]" pageId="29" pageNumber="30">
<emphasis id="B5D8EABFFFF19E3245A1FB98EE03FBD5" box="[199,336,1087,1113]" italics="true" pageId="29" pageNumber="30">Distribution</emphasis>
. Only known from
<collectingCountry id="FFBB763DFFF19E324748FB98ED3FFBD5" box="[558,620,1087,1113]" name="Israel" pageId="29" pageNumber="30">Israel</collectingCountry>
(Kasparek
<emphasis id="B5D8EABFFFF19E32478BFB98EC4DFBD5" box="[749,798,1087,1113]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2023).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,338 @@
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<mods:title id="EF9FA31F850717C2232580132DEF5006">New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)</mods:title>
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<treatment id="0F0587BBFFF89E3945F1FBC5EDE3FD35" ID-DOI="http://doi.org/10.5281/zenodo.14248297" ID-Zenodo-Dep="14248297" LSID="urn:lsid:plazi:treatment:0F0587BBFFF89E3945F1FBC5EDE3FD35" httpUri="http://treatment.plazi.org/id/0F0587BBFFF89E3945F1FBC5EDE3FD35" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<subSubSection id="CFB66526FFF89E3B45F1FBC5ED8BFBF0" box="[151,728,1122,1149]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="871336ADFFF89E3B45F1FBC5ED8BFBF0" blockId="20.[151,728,1122,1185]" box="[151,728,1122,1149]" pageId="20" pageNumber="21">
<heading id="DC5B81C1FFF89E3B45F1FBC5ED8BFBF0" bold="true" box="[151,728,1122,1149]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="40AC4D2EFFF89E3B45F1FBC5ED8BFBF0" ID-CoL="4NP4T" authority="(Morawitz, 1888)" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[151,728,1122,1149]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF89E3B45F1FBC5ED8BFBF0" bold="true" box="[151,728,1122,1149]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45F1FBC5EEA9FBF1" bold="true" box="[151,506,1122,1149]" italics="true" pageId="20" pageNumber="21">Pseudoanthidium puncticolle</emphasis>
(
<bibRefCitation id="E33D4B5CFFF89E3B476FFBC5ED83FBF0" author="Morawitz, F. F." box="[521,720,1122,1148]" pageId="20" pageNumber="21" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz, 1888</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3B45F1FB20EEABFB2D" box="[151,504,1158,1185]" pageId="20" pageNumber="21" type="description">
<paragraph id="871336ADFFF89E3B45F1FB20EEABFB2D" blockId="20.[151,728,1122,1185]" box="[151,504,1158,1185]" pageId="20" pageNumber="21">
(
<figureCitation id="1F972A28FFF89E3B45F9FB20EFA6FB2D" box="[159,245,1159,1185]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B4467FB20EE73FB2D" box="[257,288,1159,1185]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B444DFB20EE1AFB2D" box="[299,329,1159,1185]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="20" pageNumber="21">21</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B4433FB20EE27FB2D" box="[341,372,1159,1185]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="20" pageNumber="21">22</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B4419FB20EECEFB2D" box="[383,413,1159,1185]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="20" pageNumber="21">23</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B44CFFB20EE94FB2D" box="[425,455,1159,1185]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="20" pageNumber="21">24</figureCitation>
, 25)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3B45F1FB6CEB56FAAC" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="871336ADFFF89E3B45F1FB6CEC34FB6F" blockId="20.[151,1029,1227,1312]" box="[151,871,1227,1251]" pageId="20" pageNumber="21">
<taxonomicName id="40AC4D2EFFF89E3B45F1FB6CED4CFB6F" ID-CoL="8SS76" authority="Morawitz, 1888" authorityName="Morawitz" authorityYear="1888" box="[151,543,1227,1251]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF89E3B45F1FB6CEE2BFB6F" box="[151,376,1227,1251]" italics="true" pageId="20" pageNumber="21">Anthidium puncticolle</emphasis>
<bibRefCitation id="E33D4B5CFFF89E3B4418FB6CED4CFB6F" author="Morawitz, F. F." box="[382,543,1227,1251]" pageId="20" pageNumber="21" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz, 1888</bibRefCitation>
</taxonomicName>
.—Female, male.
<collectingCountry id="FFBB763DFFF89E3B47B2FB6CEC30FB6F" box="[724,867,1227,1251]" name="Turkmenistan" pageId="20" pageNumber="21">Turkmenistan</collectingCountry>
.
</paragraph>
<paragraph id="871336ADFFF89E3B45F1FB4DEC4AFA8E" blockId="20.[151,1029,1227,1312]" box="[151,793,1258,1282]" pageId="20" pageNumber="21">
<taxonomicName id="40AC4D2EFFF89E3B45F1FB4DED34FA8E" ID-CoL="4NP4T" authority="Morawitz, 1888" authorityName="Morawitz" authorityYear="1888" box="[151,615,1258,1282]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF89E3B45F1FB4DEE93FA8E" box="[151,448,1258,1282]" italics="true" pageId="20" pageNumber="21">Pseudoanthidium puncticolle</emphasis>
<bibRefCitation id="E33D4B5CFFF89E3B44A0FB4DED34FA8E" author="Morawitz, F. F." box="[454,615,1258,1282]" pageId="20" pageNumber="21" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz, 1888</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFF89E3B47E2FB4DEC46FA8E" author="Popov, V. B." box="[644,789,1258,1282]" pageId="20" pageNumber="21" pagination="315 - 318" refId="ref31377" refString="Popov, V. B. (1950) Generic groupings of the mid-Asian bees of the subgenus Anthiniinae (Hymenoptera, Megachilidae). Doklady Akademii Nauk SSSR, 70, 315 - 318." type="journal article" year="1950">Popov (1950)</bibRefCitation>
.
</paragraph>
<paragraph id="871336ADFFF89E3B45F1FAAFEB56FAAC" blockId="20.[151,1029,1227,1312]" box="[151,1029,1288,1312]" pageId="20" pageNumber="21">
<taxonomicName id="40AC4D2EFFF89E3B45F1FAAFEC13FAAC" ID-CoL="4NP4T" authority="(Morawitz, 1888)" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[151,832,1288,1312]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticolle" subGenus="Paraanthidiellum">
<emphasis id="B5D8EABFFFF89E3B45F1FAAFEDD9FAAC" box="[151,650,1288,1312]" italics="true" pageId="20" pageNumber="21">Pseudoanthidium (Paraanthidiellum) puncticolle</emphasis>
(
<bibRefCitation id="E33D4B5CFFF89E3B47F1FAAFEC6AFAAC" author="Morawitz, F. F." box="[663,825,1288,1312]" pageId="20" pageNumber="21" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz, 1888</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFF89E3B4639FAAFEB52FAAC" author="Pasteels, J. J." box="[863,1025,1288,1312]" pageId="20" pageNumber="21" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3B45F1FAE8EC69F9E9" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="871336ADFFF89E3B45F1FAE8EB64FA59" blockId="20.[151,1437,1358,2033]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45F1FAE8EE21FAE5" box="[151,370,1359,1385]" italics="true" pageId="20" pageNumber="21">Material examined</emphasis>
.
<collectingCountry id="FFBB763DFFF89E3B44E1FAE8ED35FAE5" box="[391,614,1359,1385]" name="Turkmenistan" pageId="20" pageNumber="21">TURKMENISTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFF89E3B4711FAE8EDC9FAE5" box="[631,666,1359,1385]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<location id="82736076FFF89E3B47C9FAE8EC77FAE5" LSID="urn:lsid:plazi:treatment:0F0587BBFFF89E3945F1FBC5EDE3FD35:82736076FFF89E3B47C9FAE8EC77FAE5" box="[687,804,1359,1385]" country="Turkmenistan" latitude="38.43" longLatPrecision="705" longitude="56.28" name="Kara-Kala" pageId="20" pageNumber="21">Kara-Kala</location>
[= Garrygala] (
<geoCoordinate id="E298506AFFF89E3B468EFAE8EB14FAE5" box="[1000,1095,1359,1385]" degrees="38.43" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="38.43">38.43°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B4130FAE9EBE6FAE5" box="[1110,1205,1358,1385]" degrees="56.28" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="56.28">56.28°E</geoCoordinate>
),
<date id="F312106DFFF89E3B41A9FAE9EA6DFAE5" box="[1231,1342,1358,1385]" pageId="20" pageNumber="21" value="1952-05-23">
<collectingDate id="E356E985FFF89E3B41A9FAE9EA6DFAE5" box="[1231,1342,1358,1385]" pageId="20" pageNumber="21" value="1952-05-23">23.v.1952</collectingDate>
</date>
(
<collectionCode id="E1BDAE68FFF89E3B4035FAE8EACBFAE5" box="[1363,1432,1359,1385]" pageId="20" pageNumber="21">CMK</collectionCode>
: ms3704).—
<specimenCount id="91AAFD24FFF89E3B444CFAD4EE12FA01" box="[298,321,1395,1421]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<quantity id="40549B48FFF89E3B4436FAD4EED1FA01" box="[336,386,1395,1421]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="km" value="6.0">6km</quantity>
<collectionCode id="E1BDAE68FFF89E3B44EAFAD4EECFFA01" box="[396,412,1395,1421]" pageId="20" pageNumber="21">S</collectionCode>
<location id="82736076FFF89E3B44C3FAD4ED49FA01" LSID="urn:lsid:plazi:treatment:0F0587BBFFF89E3945F1FBC5EDE3FD35:82736076FFF89E3B44C3FAD4ED49FA01" box="[421,538,1395,1421]" country="Turkmenistan" latitude="38.38" longLatPrecision="705" longitude="56.3" name="Kara-Kala" pageId="20" pageNumber="21">Kara-Kala</location>
(
<geoCoordinate id="E298506AFFF89E3B474AFAD4EDD8FA01" box="[556,651,1395,1421]" degrees="38.38" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="38.38">38.38°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B47F0FAD5EDA6FA01" box="[662,757,1394,1421]" degrees="56.30" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="56.3">56.30°E</geoCoordinate>
),
<date id="F312106DFFF89E3B466DFAD5EC29FA01" box="[779,890,1394,1421]" pageId="20" pageNumber="21" value="1952-05-25">
<collectingDate id="E356E985FFF89E3B466DFAD5EC29FA01" box="[779,890,1394,1421]" pageId="20" pageNumber="21" value="1952-05-25">25.v.1952</collectingDate>
</date>
(
<collectionCode id="E1BDAE68FFF89E3B46EDFAD4EC83FA01" box="[907,976,1395,1421]" pageId="20" pageNumber="21">CMK</collectionCode>
: ms3705).—
<specimenCount id="91AAFD24FFF89E3B410AFAD5EBDAFA01" box="[1132,1161,1394,1421]" count="2" pageId="20" pageNumber="21" type="female">2♀</specimenCount>
,
<specimenCount id="91AAFD24FFF89E3B41F1FAD4EBE8FA01" box="[1175,1211,1395,1421]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<location id="82736076FFF89E3B41ADFAD4EACFFA01" LSID="urn:lsid:plazi:treatment:0F0587BBFFF89E3945F1FBC5EDE3FD35:82736076FFF89E3B41ADFAD4EACFFA01" box="[1227,1436,1395,1421]" country="Turkmenistan" latitude="37.96" longLatPrecision="707" longitude="58.2" name="10 km W Asgabat" pageId="20" pageNumber="21" stateProvince="Ashgabat">
<quantity id="40549B48FFF89E3B41ADFAD4EA46FA01" box="[1227,1301,1395,1421]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="10.0">10 km</quantity>
W
<collectingRegion id="4568F84FFFF89E3B4026FAD4EACFFA01" box="[1344,1436,1395,1421]" country="Turkmenistan" name="Ashgabat" pageId="20" pageNumber="21">Aşgabat</collectingRegion>
</location>
[=
<collectingRegion id="4568F84FFFF89E3B45DFFA30EE61FA3D" box="[185,306,1431,1457]" country="Turkmenistan" name="Ashgabat" pageId="20" pageNumber="21">Aschgabat</collectingRegion>
] (
<geoCoordinate id="E298506AFFF89E3B442CFA30EEF9FA3D" box="[330,426,1431,1457]" degrees="37.96" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="37.96">37.96°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B44D2FA31ED47FA3D" box="[436,532,1430,1457]" degrees="58.20" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="58.2">58.20°E</geoCoordinate>
),
<date id="F312106DFFF89E3B474CFA30EDC8FA3D" box="[554,667,1431,1457]" pageId="20" pageNumber="21" value="1993-05-14">
<collectingDate id="E356E985FFF89E3B474CFA30EDC8FA3D" box="[554,667,1431,1457]" pageId="20" pageNumber="21" value="1993-05-14">14.v.1993</collectingDate>
</date>
,
<collectorName id="2A59537BFFF89E3B47CCFA30EC46FA3D" box="[682,789,1431,1457]" pageId="20" pageNumber="21">J. Halada</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF89E3B463CFA30ECCCFA3D" box="[858,927,1431,1457]" pageId="20" pageNumber="21">CMK</collectionCode>
: ms3742-44).—
<specimenCount id="91AAFD24FFF89E3B4106FA30EB2EFA3D" box="[1120,1149,1431,1457]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<location id="82736076FFF89E3B41EDFA30EA4BFA3D" LSID="urn:lsid:plazi:treatment:0F0587BBFFF89E3945F1FBC5EDE3FD35:82736076FFF89E3B41EDFA30EA4BFA3D" box="[1163,1304,1431,1457]" country="Turkmenistan" latitude="37.69" longLatPrecision="708" longitude="66.83" name="Tzarschanga" pageId="20" pageNumber="21">Tzarschanga</location>
<locationDeviation id="96A12F8FFFF89E3B4045FA31EFA0FA59" location="Karlyuk" pageId="20" pageNumber="21">
<quantity id="40549B48FFF89E3B4045FA31EA3EFA3D" box="[1315,1389,1430,1457]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="21" unit="km" value="45.0">45 km</quantity>
NE Karlyuk
</locationDeviation>
[= Garlyk] (
<geoCoordinate id="E298506AFFF89E3B44E6FA1CEEB3FA59" box="[384,480,1467,1493]" degrees="37.69" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="37.69">37.69°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B4481FA1CED15FA59" box="[487,582,1467,1493]" degrees="66.83" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="66.83">66.83°E</geoCoordinate>
),
<date id="F312106DFFF89E3B473CFA1CED98FA59" box="[602,715,1467,1493]" pageId="20" pageNumber="21" value="1993-05-06">
<collectingDate id="E356E985FFF89E3B473CFA1CED98FA59" box="[602,715,1467,1493]" pageId="20" pageNumber="21" value="1993-05-06">06.v.1993</collectingDate>
</date>
,
<collectorName id="2A59537BFFF89E3B47B0FA1CEC6DFA59" box="[726,830,1467,1493]" pageId="20" pageNumber="21">K. Deneš</collectorName>
leg. (
<collectionCode id="E1BDAE68FFF89E3B461BFA1CEC91FA59" box="[893,962,1467,1493]" pageId="20" pageNumber="21">CMK</collectionCode>
: ms3745).
</paragraph>
<paragraph id="871336ADFFF89E3B45A1FA78ECACF9CD" blockId="20.[151,1437,1358,2033]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1FA78ED76FA75" box="[199,549,1503,1529]" italics="true" pageId="20" pageNumber="21">Other material (not examined).</emphasis>
<collectingCountry id="FFBB763DFFF89E3B474CFA78EC5AFA75" box="[554,777,1503,1529]" name="Turkmenistan" pageId="20" pageNumber="21">TURKMENISTAN</collectingCountry>
:
<specimenCount id="91AAFD24FFF89E3B4675FA78EC65FA75" box="[787,822,1503,1529]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<specimenCount id="91AAFD24FFF89E3B4622FA78EC3BFA75" box="[836,872,1503,1529]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
, Tschikischljar [= Chikishlyar] (
<geoCoordinate id="E298506AFFF89E3B41B7FA79EA62FA75" box="[1233,1329,1502,1529]" degrees="37.57" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="37.57">37.57°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B405EFA79EAC6FA75" box="[1336,1429,1502,1529]" degrees="53.90" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="53.9">53.90°E</geoCoordinate>
) (
<typeStatus id="5817880FFFF89E3B45F9F9A4EF82F991" box="[159,209,1539,1565]" pageId="20" pageNumber="21">type</typeStatus>
locality) (
<bibRefCitation id="E33D4B5CFFF89E3B4421F9A4EEAAF991" author="Morawitz, F. F." box="[327,505,1539,1565]" pageId="20" pageNumber="21" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz 1888</bibRefCitation>
;
<bibRefCitation id="E33D4B5CFFF89E3B4761F9A4EDDEF991" author="Friese, H." box="[519,653,1539,1565]" pageId="20" pageNumber="21" refId="ref29768" refString="Friese, H. (1898) Die Bienen Europa's (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp." type="book" year="1898">Friese 1898</bibRefCitation>
).—
<collectingCountry id="FFBB763DFFF89E3B47D1F9A4EC52F991" box="[695,769,1539,1565]" name="Iran" pageId="20" pageNumber="21">IRAN</collectingCountry>
: Pass E of Neyriz (
<geoCoordinate id="E298506AFFF89E3B46B9F9A4EB6CF991" box="[991,1087,1539,1565]" degrees="29.23" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="29.23">29.23°N</geoCoordinate>
<geoCoordinate id="E298506AFFF89E3B412EF9A5EBF4F991" box="[1096,1191,1538,1565]" degrees="54.41" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="54.41">54.41°E</geoCoordinate>
),
<date id="F312106DFFF89E3B41DAF9A4EA7EF991" box="[1212,1325,1539,1565]" pageId="20" pageNumber="21" value="1978-05-18">18.v.1978</date>
,
<bibRefCitation id="E33D4B5CFFF89E3B405FF9A4EFB0F9CD" author="Warncke, K." pageId="20" pageNumber="21" pagination="171 - 196" refId="ref31857" refString="Warncke, K. (1982) Beitrag zur Bienenfauna des Iran. 15. Die Gattung Anthidium F. Bollettino del Museo Civico di Storia Naturale di Venezia, 32 (1981), 171 - 196." type="journal article" year="1982">Warncke (1982)</bibRefCitation>
.—
<quantity id="40549B48FFF89E3B4460F981EE1FF9CD" box="[262,332,1574,1601]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="km" value="15.0">15 km</quantity>
SE Sarvestan (
<geoCoordinate id="E298506AFFF89E3B4490F980ED05F9CD" box="[502,598,1575,1601]" degrees="29.19" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="29.19">29.19°N</geoCoordinate>
53.35E),
<date id="F312106DFFF89E3B47A8F980EC6FF9CD" box="[718,828,1575,1601]" pageId="20" pageNumber="21" value="1978-05-17">17.v.1978</date>
(
<bibRefCitation id="E33D4B5CFFF89E3B462CF980ECA1F9CD" author="Warncke, K." box="[842,1010,1575,1601]" pageId="20" pageNumber="21" pagination="171 - 196" refId="ref31857" refString="Warncke, K. (1982) Beitrag zur Bienenfauna des Iran. 15. Die Gattung Anthidium F. Bollettino del Museo Civico di Storia Naturale di Venezia, 32 (1981), 171 - 196." type="journal article" year="1982">Warncke 1982</bibRefCitation>
).
</paragraph>
<paragraph id="871336ADFFF89E3B45A1F9ECEC69F9E9" blockId="20.[151,1437,1358,2033]" box="[199,826,1611,1637]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1F9ECED58F9E9" box="[199,523,1611,1637]" italics="true" pageId="20" pageNumber="21">Genetic barcode information</emphasis>
. No information available.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3B45A1F9C8EB24F895" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="871336ADFFF89E3B45A1F9C8EB24F895" blockId="20.[151,1437,1358,2033]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1F9C8EE6AF904" box="[199,313,1647,1672]" italics="true" pageId="20" pageNumber="21">Diagnosis</emphasis>
. The female is characterised by rich yellow and/or orange maculation combined with coarse punctation of terga and a thin, little protruding apical ridge of the yellow or orange clypeus, having three fine tubercles on each side. In contrast, other species either lack a protruding clypeus (as in
<taxonomicName id="40AC4D2EFFF89E3B46CAF910EB74F95D" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1948" box="[940,1063,1719,1745]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bytinskii">
<emphasis id="B5D8EABFFFF89E3B46CAF910EB74F95D" box="[940,1063,1719,1745]" italics="true" pageId="20" pageNumber="21">P. bytinskii</emphasis>
</taxonomicName>
) or have a protruding, thickened apical ridge. The males share the felt-like pubescence on S1S3 only with
<taxonomicName id="40AC4D2EFFF89E3B46B2F97CEB18F979" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[980,1099,1755,1781]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFF89E3B46B2F97CEB18F979" box="[980,1099,1755,1781]" italics="true" pageId="20" pageNumber="21">P. alpinum</emphasis>
</taxonomicName>
and
<taxonomicName id="40AC4D2EFFF89E3B41E7F97CEA63F979" authorityName="Kasparek" authorityYear="2024" box="[1153,1328,1755,1781]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="microrubrum">
<emphasis id="B5D8EABFFFF89E3B41E7F97CEA63F979" box="[1153,1328,1755,1781]" italics="true" pageId="20" pageNumber="21">P. microrubrum</emphasis>
</taxonomicName>
. While
<taxonomicName id="40AC4D2EFFF89E3B40EEF97CEE40F895" baseAuthorityName="Morawitz" baseAuthorityYear="1888" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF89E3B40EEF97CEE40F895" italics="true" pageId="20" pageNumber="21">P. puncticolle</emphasis>
</taxonomicName>
has a light clypeus and a light paraocular area, the face of
<taxonomicName id="40AC4D2EFFF89E3B46C3F958EB4EF895" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[933,1053,1791,1817]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFF89E3B46C3F958EB4EF895" box="[933,1053,1791,1817]" italics="true" pageId="20" pageNumber="21">P. alpinum</emphasis>
</taxonomicName>
is dark.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3945A1F884ED41FD1B" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
<paragraph id="871336ADFFF89E3945A1F884EB4DFE0D" blockId="20.[151,1437,1358,2033]" lastBlockId="22.[151,1437,151,697]" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1F884EE18F8B0" box="[199,331,1827,1852]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
.
<emphasis id="B5D8EABFFFF89E3B4431F884EEF8F8B1" box="[343,427,1827,1853]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
(
<figureCitation id="1F972A28FFF89E3B44DEF884ED46F8B1" box="[440,533,1827,1853]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="20" pageNumber="21">Fig. 11a</figureCitation>
).
<quantity id="40549B48FFF89E3B4741F884ED3BF8B1" box="[551,616,1827,1853]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="21" unit="mm" value="7.0">7 mm</quantity>
.—
<emphasis id="B5D8EABFFFF89E3B47EEF884ED96F8B1" box="[648,709,1827,1853]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
: Black with light maculation; upper and lower paraocular area and clypeus yellow or orange (
<figureCitation id="1F972A28FFF89E3B44DAF8E0ED5CF8ED" box="[444,527,1863,1889]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="20" pageNumber="21">Fig. 20</figureCitation>
); yellow preoccipital band reaching the lower end of the eye; basal area of clypeus and along a mid-line sparsely punctate; subapical clypeus depressed with very fine/rugulose punctation; apical ridge of clypeus little protruding, light brown, with 45 shallow tubercles (
<figureCitation id="1F972A28FFF89E3B46C7F828EB52F825" box="[929,1025,1935,1961]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="20" pageNumber="21">Fig. 11c</figureCitation>
); mandible yellow, becoming brown towards the teeth; five black teeth; to1 und to2 the largest, to3 and to4 smaller, to5 larger and stronger; punctation of head moderately dense with punctures mostly 0.51.0 puncture diameter apart.—
<taxonomicName id="40AC4D2EFFF89E3B415FF870EBE0F87C" box="[1081,1203,2007,2032]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF89E3B415FF870EBE0F87C" box="[1081,1203,2007,2032]" italics="true" pageId="20" pageNumber="21">Mesosoma</emphasis>
</taxonomicName>
: Scutum black with broad anterior yellow-orange band (
<figureCitation id="1F972A28FFFA9E39474AFF30EDDFFF3D" box="[556,652,151,177]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="22" pageNumber="23">Fig. 11d</figureCitation>
); scutum shining; punctation as on head; scutellum and axilla orange; scutellum in profile rounded in the middle and acute-angled laterally; punctures on scutellum and axillae larger than on scutum; pronotal lobe with high lamella; omaulus rounded.—
<emphasis id="B5D8EABFFFFA9E3946CFFF7AEB4DFF7A" box="[937,1054,221,246]" italics="true" pageId="22" pageNumber="23">Metasoma</emphasis>
: T1T6 each with a broad orange or yellow lateral band; punctation coarse, with punctures mostly 0.5 puncture diameters apart; broad shining impunctate apical margin; T6 wide-angled triangular in dorsal view, with short grey protruding apex (
<figureCitation id="1F972A28FFFA9E394078FE86EA21FEB7" box="[1310,1394,289,315]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="22" pageNumber="23">Fig. 22</figureCitation>
).—
<emphasis id="B5D8EABFFFFA9E3945F1FEE2EF9FFED2" box="[151,204,325,350]" italics="true" pageId="22" pageNumber="23">Legs</emphasis>
: Trochanters and basal parts of tibiae black; otherwise orange-yellow; hind basitarsi greyish or black on outer face and with short bristles on inner face.—
<emphasis id="B5D8EABFFFFA9E3947E7FEC0ED96FE0C" box="[641,709,359,384]" italics="true" pageId="22" pageNumber="23">Wings</emphasis>
: Slightly brownish infuscated.
</paragraph>
<caption id="D3D36625FFF99E3A45F1F830ED38F87B" ID-DOI="http://doi.org/10.5281/zenodo.14240759" ID-Zenodo-Dep="14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1943,1967]" targetBox="[152,1436,181,1918]" targetPageId="21" targetType="figure">
<paragraph id="871336ADFFF99E3A45F1F830ED38F87B" blockId="21.[151,1437,1943,2039]" pageId="21" pageNumber="22">
<emphasis id="B5D8EABFFFF99E3A45F1F830EE4AF823" bold="true" box="[151,281,1943,1967]" pageId="21" pageNumber="22">FIGURE 11</emphasis>
.
<taxonomicName id="40AC4D2EFFF99E3A4441F830EC57F823" authority="(Morawitz 1888)" baseAuthorityName="Morawitz" baseAuthorityYear="1888" box="[295,772,1943,1967]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="puncticolle">
<emphasis id="B5D8EABFFFF99E3A4441F830ED03F823" box="[295,592,1943,1967]" italics="true" pageId="21" pageNumber="22">Pseudoanthidium puncticolle</emphasis>
(
<bibRefCitation id="E33D4B5CFFF99E3A4738F830EDAEF823" author="Morawitz, F. F." box="[606,765,1943,1967]" pageId="21" pageNumber="22" pagination="224 - 302" refId="ref30939" refString="Morawitz, F. F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva), 22, 224 - 302." type="journal article" year="1888">Morawitz 1888</bibRefCitation>
)
</taxonomicName>
(a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5.
</paragraph>
</caption>
<paragraph id="871336ADFFFA9E3945A1FE2EED41FD1B" blockId="22.[151,1437,151,697]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945A1FE2EEE52FE2F" box="[199,257,393,419]" italics="true" pageId="22" pageNumber="23">Male</emphasis>
(
<figureCitation id="1F972A28FFFA9E394476FE2EEE23FE2F" box="[272,368,393,419]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="22" pageNumber="23">Fig. 11b</figureCitation>
).
<quantity id="40549B48FFFA9E3944E3FE2EEE94FE28" box="[389,455,393,420]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="mm" value="8.0">8 mm</quantity>
.—
<emphasis id="B5D8EABFFFFA9E39448EFE2EED76FE2F" box="[488,549,393,419]" italics="true" pageId="22" pageNumber="23">Head</emphasis>
: Face largely covered by long white pubescence; clypeus and lower paraocular area ivory-white (
<figureCitation id="1F972A28FFFA9E39443BFE0BEEFCFE4A" box="[349,431,428,454]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="22" pageNumber="23">Fig. 21</figureCitation>
); mandible yellowish with three strong dark teeth.—
<emphasis id="B5D8EABFFFFA9E39469EFE0AEB2BFE4A" box="[1016,1144,429,454]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="40AC4D2EFFFA9E39469EFE0AEB20FE4A" box="[1016,1139,429,454]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Scutum black with coarse punctation (
<figureCitation id="1F972A28FFFA9E39447AFE68EE28FE65" box="[284,379,463,489]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[152,1436,181,1918]" captionTargetId="figure-15@21.[152,1436,181,1918]" captionTargetPageId="21" captionText="FIGURE 11. Pseudoanthidium puncticolle (Morawitz 1888) (a.e. from Turkmenistan). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Head of the female; d. Dorsal view of the mesosoma, male; e. Sterna of the male with specialized undulated and apically hooked hairs on S3 and combs on S5." figureDoi="http://doi.org/10.5281/zenodo.14240759" httpUri="https://zenodo.org/record/14240759/files/figure.png" pageId="22" pageNumber="23">Fig. 11e</figureCitation>
); interstices shining; pronotal lobe dark with high lamella; scutellum shallowly curved upward and depressed in the middle.—
<emphasis id="B5D8EABFFFFA9E394499FE55ED27FD87" box="[511,628,498,523]" italics="true" pageId="22" pageNumber="23">Metasoma</emphasis>
: T1T4 each with a lateral yellow spot (increasing in size fromT1 to T3, smaller in T4); T5T6 black; T7 short, yellow, membranous; S1S3 with dese, felt-like pubescence; S3 with specialized, undulate and apically hooked hairs; S5 with a light, yellowish comb at the apex of a lateral arm (
<figureCitation id="1F972A28FFFA9E39400CFD90EFE6FDFF" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="22" pageNumber="23">Figs 23</figureCitation>
,
<figureCitation id="1F972A28FFFA9E3945A6FDFEEF8DFDFF" box="[192,222,601,627]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="22" pageNumber="23">24</figureCitation>
).—
<emphasis id="B5D8EABFFFFA9E39446FFDFEEE20FDFF" box="[265,371,601,627]" italics="true" pageId="22" pageNumber="23">Genitalia</emphasis>
: Gonostyli paddle-shaped, not notched apically; penis valves separated from each other, the tips bent towards each other (
<figureCitation id="1F972A28FFFA9E3944D7FDDBED55FD1B" box="[433,518,636,663]" captionStart="FIGURE 25" captionStartId="40.[151,250,2009,2033]" captionTargetBox="[176,1412,1240,1996]" captionTargetId="figure-103@40.[176,1412,1240,2001]" captionTargetPageId="40" captionText="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240785" httpUri="https://zenodo.org/record/14240785/files/figure.png" pageId="22" pageNumber="23">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFA9E3945A1FD38EDE3FD35" box="[199,688,671,697]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="871336ADFFFA9E3945A1FD38EDE3FD35" blockId="22.[151,1437,151,697]" box="[199,688,671,697]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945A1FD38EE0BFD35" box="[199,344,671,697]" italics="true" pageId="22" pageNumber="23">Distribution.</emphasis>
<collectingCountry id="FFBB763DFFFA9E394438FD38EEDDFD35" box="[350,398,671,697]" name="Iran" pageId="22" pageNumber="23">Iran</collectingCountry>
,
<collectingCountry id="FFBB763DFFFA9E3944FFFD38ED67FD35" box="[409,564,671,697]" name="Turkmenistan" pageId="22" pageNumber="23">Turkmenistan</collectingCountry>
(
<figureCitation id="1F972A28FFFA9E394725FD38EDF7FD35" box="[579,676,671,697]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="22" pageNumber="23">Fig. 27a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,547 @@
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<paragraph id="871336ADFFFA9E3945F1FC4AED95FB8B" blockId="22.[151,710,1005,1067]" box="[151,710,1005,1032]" pageId="22" pageNumber="23">
<heading id="DC5B81C1FFFA9E3945F1FC4AED95FB8B" bold="true" box="[151,710,1005,1032]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="40AC4D2EFFFA9E3945F1FC4AED95FB8B" ID-CoL="4NP4V" authority="(Friese, 1917)" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[151,710,1005,1032]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhombiferum">
<emphasis id="B5D8EABFFFFA9E3945F1FC4AED95FB8B" bold="true" box="[151,710,1005,1032]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945F1FC4AED46FB84" bold="true" box="[151,533,1005,1032]" italics="true" pageId="22" pageNumber="23">Pseudoanthidium rhombiferum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFA9E394743FC4AEDEDFB8B" author="Friese, H." box="[549,702,1005,1031]" pageId="22" pageNumber="23" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="CFB66526FFFA9E3945F1FBB7EEAAFBA7" box="[151,505,1040,1067]" pageId="22" pageNumber="23" type="description">
<paragraph id="871336ADFFFA9E3945F1FBB7EEAAFBA7" blockId="22.[151,710,1005,1067]" box="[151,505,1040,1067]" pageId="22" pageNumber="23">
(
<figureCitation id="1F972A28FFFA9E3945F9FBB7EFA5FBA6" box="[159,246,1040,1066]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="22" pageNumber="23">Figs 12</figureCitation>
,
<figureCitation id="1F972A28FFFA9E394464FBB7EE72FBA6" box="[258,289,1040,1066]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="22" pageNumber="23">20</figureCitation>
,
<figureCitation id="1F972A28FFFA9E39444AFBB7EE19FBA6" box="[300,330,1040,1066]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="22" pageNumber="23">21</figureCitation>
,
<figureCitation id="1F972A28FFFA9E394430FBB7EE26FBA6" box="[342,373,1040,1066]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="22" pageNumber="23">22</figureCitation>
,
<figureCitation id="1F972A28FFFA9E3944E6FBB7EECDFBA6" box="[384,414,1040,1066]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="22" pageNumber="23">23</figureCitation>
,
<figureCitation id="1F972A28FFFA9E3944CCFBB7EE9BFBA6" box="[426,456,1040,1066]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="22" pageNumber="23">24</figureCitation>
, 25)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFA9E3945F1FBF0ECB9FB1E" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="871336ADFFFA9E3945F1FBF0EDFFFBE3" blockId="22.[151,1002,1111,1170]" box="[151,684,1111,1135]" pageId="22" pageNumber="23">
<taxonomicName id="40AC4D2EFFFA9E3945F1FBF0ED5CFBE3" ID-CoL="8SRVJ" authority="Friese, 1917" authorityName="Friese" authorityYear="1917" box="[151,527,1111,1135]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhombiferum">
<emphasis id="B5D8EABFFFFA9E3945F1FBF0EED8FBE3" box="[151,395,1111,1135]" italics="true" pageId="22" pageNumber="23">Anthidium rhombiferum</emphasis>
<bibRefCitation id="E33D4B5CFFFA9E3944F7FBF0ED5CFBE3" author="Friese, H." box="[401,527,1111,1135]" pageId="22" pageNumber="23" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
</taxonomicName>
.—
<collectingCountry id="FFBB763DFFFA9E39474BFBF0ED3AFBE3" box="[557,617,1111,1135]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
, male.
</paragraph>
<paragraph id="871336ADFFFA9E3945F1FBDDECB9FB1E" blockId="22.[151,1002,1111,1170]" box="[151,1002,1146,1170]" pageId="22" pageNumber="23">
<taxonomicName id="40AC4D2EFFFA9E3945F1FBDDEC7DFB1E" ID-CoL="4NP4V" authority="(Friese, 1917)" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[151,814,1146,1170]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhombiferum" subGenus="Pseudoanthidium">
<emphasis id="B5D8EABFFFFA9E3945F1FBDDEE1BFB1E" box="[151,328,1146,1170]" italics="true" pageId="22" pageNumber="23">Pseudoanthidium</emphasis>
(
<emphasis id="B5D8EABFFFFA9E394431FBDDED5BFB1E" box="[343,520,1146,1170]" italics="true" pageId="22" pageNumber="23">Pseudoanthidium</emphasis>
)
<emphasis id="B5D8EABFFFFA9E394771FBDDEDC8FB1E" box="[535,667,1146,1170]" italics="true" pageId="22" pageNumber="23">rhombiferum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFA9E3947CFFBDDEC74FB1E" author="Friese, H." box="[681,807,1146,1170]" pageId="22" pageNumber="23" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFFA9E39462AFBDDECB9FB1E" author="Pasteels, J. J." box="[844,1002,1146,1170]" pageId="22" pageNumber="23" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFA9E3945F1FB1AEE25F87A" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="871336ADFFFA9E3945F1FB1AEDBCF983" blockId="22.[151,1437,1213,2038]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945F1FB1AEE3DFB5B" box="[151,366,1213,1239]" italics="true" pageId="22" pageNumber="23">Material examined</emphasis>
.
<materialsCitation id="37C43CF0FFFA9E3944E6FB1AEC4FFB76" collectingDate="1905-03-13" collectionCode="ZMB" collectorName="Enslin" country="Israel" latitude="32.04" location="Jaffa" longLatPrecision="727" longitude="34.76" municipality="Syrien" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Tel Aviv-Jaffa" typeStatus="holotype">
<collectingCountry id="FFBB763DFFFA9E3944E6FB1AEEB5FB5B" box="[384,486,1213,1239]" name="Israel" pageId="22" pageNumber="23">ISRAEL</collectingCountry>
:
<specimenCount id="91AAFD24FFFA9E394495FB1AED4BFB5B" box="[499,536,1213,1239]" count="1" pageId="22" pageNumber="23" type="male">1♂</specimenCount>
,
<typeStatus id="5817880FFFFA9E394741FB1AEDDFFB5B" box="[551,652,1213,1239]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
, “
<collectingMunicipality id="6777ACD7FFFA9E3947C3FB1AEDA2FB5B" box="[677,753,1213,1239]" pageId="22" pageNumber="23">Syrien</collectingMunicipality>
(
<location id="82736076FFFA9E394664FB1AEC6DFB5B" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E394664FB1AEC6DFB5B" box="[770,830,1213,1239]" country="Israel" latitude="32.04" longLatPrecision="727" longitude="34.76" municipality="Syrien" name="Jaffa" pageId="22" pageNumber="23" stateProvince="Tel Aviv-Jaffa">Jaffa</location>
)“ [
<collectingRegion id="4568F84FFFFA9E394639FB1AEB50FB5B" box="[863,1027,1213,1239]" country="Israel" name="Tel Aviv" pageId="22" pageNumber="23">Tel Aviv-Jaffa</collectingRegion>
(c.
<geoCoordinate id="E298506AFFFA9E394152FB1AEBC0FB5B" box="[1076,1171,1213,1239]" degrees="32.04" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.04">32.04°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E3941FBFB1AEBACFB5B" box="[1181,1279,1213,1239]" degrees="34.76" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="34.76">34.76°E</geoCoordinate>
)],
<date id="F312106DFFFA9E39407AFB1AEACBFB54" box="[1308,1432,1213,1240]" pageId="22" pageNumber="23" value="1905-03-13">
<collectingDate id="E356E985FFFA9E39407AFB1AEACBFB54" box="[1308,1432,1213,1240]" pageId="22" pageNumber="23" value="1905-03-13">13.iii.1905</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E3945F1FB47EF8CFB76" box="[151,223,1248,1274]" pageId="22" pageNumber="23">Enslin</collectorName>
leg.; see also description by
<bibRefCitation id="E33D4B5CFFFA9E39474FFB47EDECFB76" author="Friese, H." box="[553,703,1248,1274]" pageId="22" pageNumber="23" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese (1917)</bibRefCitation>
(
<collectionCode id="E1BDAE68FFFA9E3947B6FB47EC47FB76" box="[720,788,1248,1274]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFFA9E394627FB47EB20FA91" collectingDate="2018-04-27" collectionCode="OLL" collectorName="M. Halada" country="Israel" latitude="33.1" location="Ramot Naftali" longLatPrecision="724" longitude="35.55" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="91AAFD24FFFA9E394627FB47EC0CFB76" box="[833,863,1248,1274]" count="1" pageId="22" pageNumber="23" type="male">1♂</specimenCount>
,
<location id="82736076FFFA9E39460AFB46EB5CFB76" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E39460AFB46EB5CFB76" box="[876,1039,1248,1275]" country="Israel" latitude="33.1" longLatPrecision="724" longitude="35.55" name="Ramot Naftali" pageId="22" pageNumber="23">Ramot Naftali</location>
,
<locationDeviation id="96A12F8FFFFA9E39417DFB47EA75FB76" box="[1051,1318,1248,1274]" location="Kiryat Shmona" pageId="22" pageNumber="23">
<quantity id="40549B48FFFA9E39417DFB47EB08FB76" box="[1051,1115,1248,1274]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="km" value="10.0">10km</quantity>
S Kiryat Shmona
</locationDeviation>
(
<geoCoordinate id="E298506AFFFA9E394053FB47EACBFB76" box="[1333,1432,1248,1274]" degrees="33.10" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="33.1">33.10°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E3945F1FAA5EFA5FA91" box="[151,246,1282,1309]" degrees="35.55" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.55">35.55°E</geoCoordinate>
),
<date id="F312106DFFFA9E394468FAA4EED5FA91" box="[270,390,1283,1309]" pageId="22" pageNumber="23" value="2018-04-27">
<collectingDate id="E356E985FFFA9E394468FAA4EED5FA91" box="[270,390,1283,1309]" pageId="22" pageNumber="23" value="2018-04-27">27.iv.2018</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E3944F0FAA4ED5CFA91" box="[406,527,1283,1309]" pageId="22" pageNumber="23">M. Halada</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFFA9E39466FFAA4ECE8FA91" box="[777,955,1283,1309]" pageId="22" pageNumber="23">ABABX470-22</accessionNumber>
(
<collectionCode id="E1BDAE68FFFA9E3946A8FAA4EB59FA91" box="[974,1034,1283,1309]" pageId="22" pageNumber="23">OLL</collectionCode>
: oll900)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFFA9E3941FFFAA4EBBFFAB3" collectingDate="2023-04-14" collectionCode="CMK" collectorName="M. Halada" country="Israel" latitude="30.79" location="Ezuz S of Nizzana" longLatPrecision="732" longitude="34.47" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="91AAFD24FFFA9E3941FFFAA4EBE1FA91" box="[1177,1202,1283,1309]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
,
<location id="82736076FFFA9E3941A7FAA4EACFFA91" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E3941A7FAA4EACFFA91" box="[1217,1436,1283,1309]" country="Israel" latitude="30.79" longLatPrecision="732" longitude="34.47" name="Ezuz S of Nizzana" pageId="22" pageNumber="23">Ezuz S of Nizzana</location>
(
<geoCoordinate id="E298506AFFFA9E3945F9FA82EFACFAB3" box="[159,255,1317,1343]" degrees="30.79" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="30.79">30.79°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E39446FFA82EE3BFAB3" box="[265,360,1317,1343]" degrees="34.47" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="34.47">34.47°E</geoCoordinate>
),
<date id="F312106DFFFA9E394418FA82EEA4FAB3" box="[382,503,1317,1343]" pageId="22" pageNumber="23" value="2023-04-14">
<collectingDate id="E356E985FFFA9E394418FA82EEA4FAB3" box="[382,503,1317,1343]" pageId="22" pageNumber="23" value="2023-04-14">14.iv.2023</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394763FA81ED2DFAB3" box="[517,638,1317,1344]" pageId="22" pageNumber="23">M. Halada</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFFA9E394612FA82EB71FAB3" box="[884,1058,1317,1343]" pageId="22" pageNumber="23">ABABZ043-23</accessionNumber>
(
<collectionCode id="E1BDAE68FFFA9E394152FA82EB2AFAB3" box="[1076,1145,1317,1343]" pageId="22" pageNumber="23">CMK</collectionCode>
: mk1232)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFFA9E39406BFA82EBF9FAEE" collectingDate="2023-05-09" collectionCode="TUZ" collectorName="V. Soon" country="Lebanon" elevation="300" latitude="0.55" location="Mies Castle" longitude="35.36" municipality="Gov." pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" stateProvince="Nabatieh" typeStatus="holotype">
<collectingCountry id="FFBB763DFFFA9E39406BFA82EACAFAB3" box="[1293,1433,1317,1343]" name="Lebanon" pageId="22" pageNumber="23">LEBANON</collectingCountry>
:
<specimenCount id="91AAFD24FFFA9E3945F1FAEFEFE9FAEE" box="[151,186,1352,1378]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
,
<collectingRegion id="4568F84FFFFA9E3945ACFAEFEE7CFAEE" box="[202,303,1352,1378]" country="Lebanon" name="Nabatiye" pageId="22" pageNumber="23">Nabatieh</collectingRegion>
Gov.,
<location id="82736076FFFA9E39441BFAEFED57FAEE" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E39441BFAEFED57FAEE" box="[381,516,1352,1378]" country="Lebanon" longitude="35.36" municipality="Gov." name="Mies Castle" pageId="22" pageNumber="23" stateProvince="Nabatieh">Mies Castle</location>
(33.36N, 35.36°E),
<quantity id="40549B48FFFA9E394791FAEFEC11FAEF" box="[759,834,1352,1379]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="m" value="300.0">
<elevation id="0C81D19EFFFA9E394791FAEFEC11FAEF" box="[759,834,1352,1379]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="F312106DFFFA9E394636FAEFEC92FAEE" box="[848,961,1352,1378]" pageId="22" pageNumber="23" value="2023-05-09">
<collectingDate id="E356E985FFFA9E394636FAEFEC92FAEE" box="[848,961,1352,1378]" pageId="22" pageNumber="23" value="2023-05-09">09.v.2023</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E3946A8FAEEEB7BFAEE" box="[974,1064,1352,1379]" pageId="22" pageNumber="23">V. Soon</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFA9E39410CFAEFEBF1FAEE" box="[1130,1186,1352,1378]" pageId="22" pageNumber="23">TUZ</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFFA9E3941ADFAEFEFB8FA2B" collectingDate="2019-05-08" collectionCode="OLL" collectorName="M. Halada" country="Palestine" latitude="32.43" location="Nablus" longLatPrecision="726" longitude="35.27" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="FFBB763DFFFA9E3941ADFAEFEA34FAEE" box="[1227,1383,1352,1378]" name="Palestine" pageId="22" pageNumber="23">PALESTINE</collectingCountry>
:
<specimenCount id="91AAFD24FFFA9E394014FAEFEAC4FAEE" box="[1394,1431,1352,1378]" count="1" pageId="22" pageNumber="23" type="male">1♂</specimenCount>
,
<location id="82736076FFFA9E3945F1FACCEFBAFA09" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E3945F1FACCEFBAFA09" box="[151,233,1387,1413]" country="Palestine" latitude="32.43" longLatPrecision="726" longitude="35.27" name="Nablus" pageId="22" pageNumber="23">Nablus</location>
,
<location id="82736076FFFA9E394590FACDEE96FA09" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E394590FACDEE96FA09" box="[246,453,1386,1413]" country="Palestine" latitude="32.43" longLatPrecision="726" longitude="35.27" name="R 57 xR 60 junction" pageId="22" pageNumber="23">R57xR60 junction</location>
(
<geoCoordinate id="E298506AFFFA9E3944B3FACCED6BFA09" box="[469,568,1387,1413]" degrees="32.43" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.43">32.43°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E394722FACDEDF0FA09" box="[580,675,1386,1413]" degrees="35.27" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.27">35.27°E</geoCoordinate>
),
<date id="F312106DFFFA9E3947DEFACCEC7AFA09" box="[696,809,1387,1413]" pageId="22" pageNumber="23" value="2019-05-08">
<collectingDate id="E356E985FFFA9E3947DEFACCEC7AFA09" box="[696,809,1387,1413]" pageId="22" pageNumber="23" value="2019-05-08">08.v.2019</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394653FACCECFFFA09" box="[821,940,1387,1413]" pageId="22" pageNumber="23">M. Halada</collectorName>
leg.; COI sequence:
<accessionNumber id="98FFAB4EFFFA9E3941FDFACCEA1EFA09" box="[1179,1357,1387,1413]" pageId="22" pageNumber="23">ABABX472-22</accessionNumber>
(
<collectionCode id="E1BDAE68FFFA9E39403BFACCEACAFA09" box="[1373,1433,1387,1413]" pageId="22" pageNumber="23">OLL</collectionCode>
: oll894)
</materialsCitation>
;
<materialsCitation id="37C43CF0FFFA9E39459FFA2AEBB4FA2B" box="[249,1255,1421,1448]" collectingDate="1934-04-26" collectionCode="ZMB" collectorName="Dr. Enslin &amp; R. Stich" country="Palestine" latitude="32.18" location="Nablus" longLatPrecision="727" longitude="35.3" pageId="22" pageNumber="23" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="holotype">
<specimenCount id="91AAFD24FFFA9E39459FFA2AEE4FFA2B" box="[249,284,1421,1447]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
,
<specimenCount id="91AAFD24FFFA9E39444BFA2AEE02FA2B" box="[301,337,1421,1447]" count="4" pageId="22" pageNumber="23" type="male">4♂</specimenCount>
,
<location id="82736076FFFA9E394406FA2AEEFCFA2B" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E394406FA2AEEFCFA2B" box="[352,431,1421,1447]" country="Palestine" latitude="32.18" longLatPrecision="727" longitude="35.3" name="Nablus" pageId="22" pageNumber="23">Nablus</location>
(
<geoCoordinate id="E298506AFFFA9E3944A7FA2AED70FA2B" box="[449,547,1421,1447]" degrees="32.18" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.18">32.18°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E394757FA2AEDC3FA24" box="[561,656,1421,1448]" degrees="35.30" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.3">35.30°E</geoCoordinate>
), 19.-
<date id="F312106DFFFA9E3947ACFA2AEC01FA2B" box="[714,850,1421,1447]" pageId="22" pageNumber="23" value="1934-04-26">
<collectingDate id="E356E985FFFA9E3947ACFA2AEC01FA2B" box="[714,850,1421,1447]" pageId="22" pageNumber="23" value="1934-04-26">26.04.1934</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394606FA29EC87FA2B" box="[864,980,1421,1448]" pageId="22" pageNumber="23">Dr. Enslin</collectorName>
&amp;
<collectorName id="2A59537BFFFA9E39469AFA29EB0AFA2B" box="[1020,1113,1421,1448]" pageId="22" pageNumber="23">R. Stich</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFA9E3941FBFA2AEB8CFA2B" box="[1181,1247,1421,1447]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="37C43CF0FFFA9E394193FA2AEC6BFA46" collectingDate="1927-04-28" collectionCode="ZMB" collectorName="Dr. Enslin" country="Palestine" latitude="31.82" location="Wadi Kelt" longLatPrecision="729" longitude="35.4" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="91AAFD24FFFA9E394193FA2AEA49FA2B" box="[1269,1306,1421,1447]" count="1" pageId="22" pageNumber="23" type="male">1♂</specimenCount>
,
<location id="82736076FFFA9E39404EFA2AEACFFA2B" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E39404EFA2AEACFFA2B" box="[1320,1436,1421,1447]" country="Palestine" latitude="31.82" longLatPrecision="729" longitude="35.4" name="Wadi Kelt" pageId="22" pageNumber="23">Wadi Kelt</location>
(
<geoCoordinate id="E298506AFFFA9E3945F9FA17EE51FA46" box="[159,258,1456,1482]" degrees="31.82" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="31.82">31.82°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E39446BFA17EE3FFA47" box="[269,364,1456,1483]" degrees="35.40" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.4">35.40°E</geoCoordinate>
), 20.
<date id="F312106DFFFA9E3944C8FA17ED61FA46" box="[430,562,1456,1482]" pageId="22" pageNumber="23" value="1927-04-28">
<collectingDate id="E356E985FFFA9E3944C8FA17ED61FA46" box="[430,562,1456,1482]" pageId="22" pageNumber="23" value="1927-04-28">28.04.1927</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E39475BFA16EDFCFA46" box="[573,687,1456,1483]" pageId="22" pageNumber="23">Dr. Enslin</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFA9E394788FA17EC63FA46" box="[750,816,1456,1482]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="37C43CF0FFFA9E39463FFA17ECFAFA61" collectingDate="1996-05-15" collectingDateMax="1996-05-17" collectingDateMin="1996-05-15" collectionCode="CMK" collectorName="Mi. Halada" country="Syria" latitude="32.44" location="Dibbin" longLatPrecision="726" longitude="36.56" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="FFBB763DFFFA9E39463FFA17ECE2FA46" box="[857,945,1456,1482]" name="Syria" pageId="22" pageNumber="23">SYRIA</collectingCountry>
:
<specimenCount id="91AAFD24FFFA9E3946DAFA17EC8CFA46" box="[956,991,1456,1482]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
,
<location id="82736076FFFA9E39468BFA17EB68FA46" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E39468BFA17EB68FA46" box="[1005,1083,1456,1482]" country="Syria" latitude="32.44" longLatPrecision="726" longitude="36.56" name="Dibbin" pageId="22" pageNumber="23">Dibbin</location>
[= Dibin],
<locationDeviation id="96A12F8FFFFA9E3941DCFA17EACEFA46" box="[1210,1437,1456,1482]" location="Suwayda" pageId="22" pageNumber="23">
<quantity id="40549B48FFFA9E3941DCFA17EBA9FA46" box="[1210,1274,1456,1482]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="km" value="30.0">30km</quantity>
S of Suwayda
</locationDeviation>
(
<geoCoordinate id="E298506AFFFA9E3945F9FA74EE51FA61" box="[159,258,1491,1517]" degrees="32.44" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.44">32.44°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E394468FA75EE3EFA61" box="[270,365,1490,1517]" degrees="36.56" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="36.56">36.56°E</geoCoordinate>
),
<date id="F312106DFFFA9E3944E7FA75ED71FA61" box="[385,546,1490,1517]" pageId="22" pageNumber="23" value="1996-05-15" valueMax="1996-05-17" valueMin="1996-05-15">
<collectingDate id="E356E985FFFA9E3944E7FA75ED71FA61" box="[385,546,1490,1517]" pageId="22" pageNumber="23" value="1996-05-15" valueMax="1996-05-17" valueMin="1996-05-15">15.17.v.1996</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394749FA74EDFDFA61" box="[559,686,1491,1517]" pageId="22" pageNumber="23">Mi. Halada</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFA9E394789FA74EC67FA61" box="[751,820,1491,1517]" pageId="22" pageNumber="23">CMK</collectionCode>
: ms4235)
</materialsCitation>
.—
<materialsCitation id="37C43CF0FFFA9E3946ABFA75EDB8F983" collectingDate="2004-04-15" collectionCode="CMK" collectorName="G. Reder" country="Syria" latitude="34.0" location="Ebla" longLatPrecision="721" longitude="37.7" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Halab" typeStatus="holotype">
<specimenCount id="91AAFD24FFFA9E3946ABFA75ECB9FA61" box="[973,1002,1490,1517]" count="1" pageId="22" pageNumber="23" type="male">1♂</specimenCount>
,
<location id="82736076FFFA9E394691FA74EB7EFA61" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E394691FA74EB7EFA61" box="[1015,1069,1491,1517]" country="Syria" latitude="34.0" longLatPrecision="721" longitude="37.7" name="Ebla" pageId="22" pageNumber="23" stateProvince="Halab">Ebla</location>
, c.
<locationDeviation id="96A12F8FFFFA9E394132FA75EA76FA61" box="[1108,1317,1490,1517]" location="Aleppo" pageId="22" pageNumber="23">
<quantity id="40549B48FFFA9E394132FA75EBCFFA61" box="[1108,1180,1490,1517]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="km" value="50.0">50 km</quantity>
SW Aleppo
</locationDeviation>
(
<geoCoordinate id="E298506AFFFA9E394053FA74EACBFA61" box="[1333,1432,1491,1517]" degrees="34.00" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="34.0">34.00°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E3945F1FA52EFA5F983" box="[151,246,1525,1551]" degrees="37.70" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="37.7">37.70°E</geoCoordinate>
),
<date id="F312106DFFFA9E39446CFA52EED1F99C" box="[266,386,1525,1552]" pageId="22" pageNumber="23" value="2004-04-15">
<collectingDate id="E356E985FFFA9E39446CFA52EED1F99C" box="[266,386,1525,1552]" pageId="22" pageNumber="23" value="2004-04-15">15.iv.2004</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E3944E8FA52EEA0F983" box="[398,499,1525,1551]" pageId="22" pageNumber="23">G. Reder</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFA9E394754FA52ED24F983" box="[562,631,1525,1551]" pageId="22" pageNumber="23">CMK</collectionCode>
: gred014)
</materialsCitation>
.
</paragraph>
<paragraph id="871336ADFFFA9E3945A1F9BFEBBBF931" blockId="22.[151,1437,1213,2038]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945A1F9BFEE20F9BE" box="[199,371,1560,1586]" italics="true" pageId="22" pageNumber="23">Other material</emphasis>
. MoA (1989) listed
<specimenCount id="91AAFD24FFFA9E394700F9BFEDF2F9BF" box="[614,673,1560,1587]" count="4" pageId="22" pageNumber="23" type="generic">4 ex.</specimenCount>
for the Mavromoustakis collection. Alfken (1935) reported
<specimenCount id="91AAFD24FFFA9E394004F9BFEACEF9BF" box="[1378,1437,1560,1587]" count="8" pageId="22" pageNumber="23" type="generic">8 ex.</specimenCount>
from
<collectorName id="2A59537BFFFA9E3945B0F99CEE76F9D9" box="[214,293,1595,1621]" pageId="22" pageNumber="23">Nablus</collectorName>
(
<geoCoordinate id="E298506AFFFA9E394453F99CEECBF9D9" box="[309,408,1595,1621]" degrees="32.43" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.43">32.43°N</geoCoordinate>
,
<geoCoordinate id="E298506AFFFA9E3944C2F99DED52F9D9" box="[420,513,1594,1621]" degrees="35.27" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.27">35.27°E</geoCoordinate>
) in 1934. Alfken (1938) reported a female from Kirjat Anawim (
<collectingCountry id="FFBB763DFFFA9E394187F99CEA71F9D9" box="[1249,1314,1595,1621]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
) (
<geoCoordinate id="E298506AFFFA9E39405DF99CEACFF9D9" box="[1339,1436,1595,1621]" degrees="31" direction="north" minutes="48" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="31.8">31°48N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E3945F1F9FAEFABF9F4" box="[151,248,1629,1656]" degrees="35" direction="east" minutes="07" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="35.116665">35°07E</geoCoordinate>
).
<bibRefCitation id="E33D4B5CFFFA9E394468F9FAEE97F9FB" author="Warncke, K." box="[270,452,1629,1655]" pageId="22" pageNumber="23" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
found the species in south-eastern
<collectingCountry id="FFBB763DFFFA9E39463DF9FAECF8F9FB" box="[859,939,1629,1655]" name="Turkey" pageId="22" pageNumber="23">Turkey</collectingCountry>
(
<quantity id="40549B48FFFA9E3946DDF9FAEB55F9FB" box="[955,1030,1629,1655]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="23" unit="km" value="40.0">40 km</quantity>
E
<collectingRegion id="4568F84FFFFA9E39414FF9FAEB0CF9FB" box="[1065,1119,1629,1655]" country="Turkey" name="Sanliurfa" pageId="22" pageNumber="23">Urfa</collectingRegion>
,
<geoCoordinate id="E298506AFFFA9E39410BF9FAEB98F9FB" box="[1133,1227,1629,1655]" degrees="37.16" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="37.16">37.16°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E3941B2F9FAEA62F9F4" box="[1236,1329,1629,1656]" degrees="39.25" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="39.25">39.25°E</geoCoordinate>
) and
<collectorName id="2A59537BFFFA9E394012F9F9EE49F916" pageId="22" pageNumber="23">van der Zanden</collectorName>
(1998) reported material from Repetek in
<collectingCountry id="FFBB763DFFFA9E394669F927ECF9F916" box="[783,938,1664,1690]" name="Turkmenistan" pageId="22" pageNumber="23">Turkmenistan</collectingCountry>
(
<geoCoordinate id="E298506AFFFA9E3946DAF927EB4FF917" box="[956,1052,1664,1691]" degrees="38.56" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="38.56">38.56°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E394141F927EBD7F916" box="[1063,1156,1664,1690]" degrees="63.17" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="63.17">63.17°E</geoCoordinate>
) and
<collectingCountry id="FFBB763DFFFA9E3941AFF927EA7DF916" box="[1225,1326,1664,1690]" name="Palestine" pageId="22" pageNumber="23">Palestine</collectingCountry>
from the collection of the Zoological Museum Moscow. The record from
<collectingCountry id="FFBB763DFFFA9E394601F904EB51F931" box="[871,1026,1699,1725]" name="Turkmenistan" pageId="22" pageNumber="23">Turkmenistan</collectingCountry>
needs confirmation.
</paragraph>
<paragraph id="871336ADFFFA9E3945A1F962EB6CF8C4" blockId="22.[151,1437,1213,2038]" pageId="22" pageNumber="23">
<location id="82736076FFFA9E3945A1F962EFA0F953" LSID="urn:lsid:plazi:treatment:0F0587BBFFFA9E3745F1FC4AEA1AFBF1:82736076FFFA9E3945A1F962EFA0F953" box="[199,243,1733,1759]" country="Israel" county="Mt. Meiron" latitude="32.99" longLatPrecision="725" longitude="35.4" municipality="Hatzor Haglilit" name="The" pageId="22" pageNumber="23">The</location>
BOLD database provided records for
<collectingCountry id="FFBB763DFFFA9E3947FDF962ED8DF953" box="[667,734,1733,1759]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
:
<specimenCount id="91AAFD24FFFA9E39478EF962EC4CF96C" box="[744,799,1733,1760]" count="2" pageId="22" pageNumber="23" type="generic">2 ex.</specimenCount>
,
<locationDeviation id="96A12F8FFFFA9E39464DF962EB2FF953" box="[811,1148,1733,1760]" location="Tzomet Mahanaim" pageId="22" pageNumber="23">
<quantity id="40549B48FFFA9E39464DF962EC30F953" box="[811,867,1733,1759]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="km" value="2.0">2 km</quantity>
W of Tzomet Mahanaim
</locationDeviation>
,
<collectingMunicipality id="6777ACD7FFFA9E3941E0F961EA7CF953" box="[1158,1327,1733,1760]" pageId="22" pageNumber="23">Hatzor Haglilit</collectingMunicipality>
(
<geoCoordinate id="E298506AFFFA9E39405AF962EACFF953" box="[1340,1436,1733,1759]" degrees="32.97" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.97">32.97°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E3945F1F94FEFA5F88F" box="[151,246,1768,1795]" degrees="35.38" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.38">35.38°E</geoCoordinate>
),
<date id="F312106DFFFA9E39446EF94FEE39F88E" box="[264,362,1768,1794]" pageId="22" pageNumber="23" value="1998-05-07">
<collectingDate id="E356E985FFFA9E39446EF94FEE39F88E" box="[264,362,1768,1794]" pageId="22" pageNumber="23" value="1998-05-07">7.v.1998</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394413F94EEE8DF88E" box="[373,478,1768,1795]" pageId="22" pageNumber="23">L. Packer</collectorName>
leg., COI sequences: BOFWI055-11 and BOFWI058-11.—
<specimenCount id="91AAFD24FFFA9E39411FF94FEBF8F88F" box="[1145,1195,1768,1795]" count="1" pageId="22" pageNumber="23" type="generic">1 ex.</specimenCount>
, Ramot, Naftali, Hwy 886 (
<geoCoordinate id="E298506AFFFA9E3945B4F8ACEE61F8A9" box="[210,306,1803,1829]" degrees="33.09" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="33.09">33.09°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E39445AF8ADEEC8F8A9" box="[316,411,1802,1829]" degrees="35.55" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.55">35.55°E</geoCoordinate>
),
<date id="F312106DFFFA9E3944D7F8ACED72F8A9" box="[433,545,1803,1829]" pageId="22" pageNumber="23" value="1998-05-14">
<collectingDate id="E356E985FFFA9E3944D7F8ACED72F8A9" box="[433,545,1803,1829]" pageId="22" pageNumber="23" value="1998-05-14">14.v.1998</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E394756F8ACEDCEF8A9" box="[560,669,1803,1829]" pageId="22" pageNumber="23">L. Packer</collectorName>
leg., COI sequence: BOFWI056-11.—
<specimenCount id="91AAFD24FFFA9E394138F8ADEBCAF8A9" box="[1118,1177,1802,1829]" count="2" pageId="22" pageNumber="23" type="generic">2 ex.</specimenCount>
,
<collectingCounty id="6E724E21FFFA9E3941C1F8ACEA78F8A9" box="[1191,1323,1803,1829]" pageId="22" pageNumber="23">Mt. Meiron</collectingCounty>
(
<geoCoordinate id="E298506AFFFA9E39405AF8ACEACFF8A9" box="[1340,1436,1803,1829]" degrees="32.99" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="32.99">32.99°N</geoCoordinate>
<geoCoordinate id="E298506AFFFA9E3945F1F88AEFA5F8C4" box="[151,246,1837,1864]" degrees="35.40" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="35.4">35.40°E</geoCoordinate>
),
<date id="F312106DFFFA9E39446CF88AEE25F8CB" box="[266,374,1837,1863]" pageId="22" pageNumber="23" value="1998-06-02">
<collectingDate id="E356E985FFFA9E39446CF88AEE25F8CB" box="[266,374,1837,1863]" pageId="22" pageNumber="23" value="1998-06-02">2.vi.1998</collectingDate>
</date>
,
<collectorName id="2A59537BFFFA9E3944E4F889EEBEF8CB" box="[386,493,1837,1864]" pageId="22" pageNumber="23">L. Packer</collectorName>
leg.; COI sequence: BOFWI057-11, BOFWI059-11.
</paragraph>
<paragraph id="871336ADFFFA9E3945A1F8F7EE25F87A" blockId="22.[151,1437,1213,2038]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945A1F8F7ED4BF8E6" box="[199,536,1872,1898]" italics="true" pageId="22" pageNumber="23">Genetic barcode information.</emphasis>
The DNA sequence of the barcoding unit of the COI gene was obtained from
<specimenCount id="91AAFD24FFFA9E3945F1F8D4EE1FF801" box="[151,332,1907,1933]" count="3" pageId="22" pageNumber="23" type="generic">three specimens</specimenCount>
from
<collectingCountry id="FFBB763DFFFA9E3944F5F8D4EE82F801" box="[403,465,1907,1933]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
(
<specimenCode id="D70A9ED6FFFA9E394487F8D4EDC5F801" box="[481,662,1907,1933]" collectionCode="ABABX" pageId="22" pageNumber="23">ABABX470-22</specimenCode>
, ABABX472-22, ABABZ043-23) and the BOLD database provided further five sequences collected by
<collectorName id="2A59537BFFFA9E394743F831EDC3F823" box="[549,656,1941,1968]" pageId="22" pageNumber="23">L. Packer</collectorName>
from
<collectingCountry id="FFBB763DFFFA9E3947B3F832EC40F823" box="[725,787,1941,1967]" name="Israel" pageId="22" pageNumber="23">Israel</collectingCountry>
(
<specimenCode id="D70A9ED6FFFA9E394645F832EC9CF83C" box="[803,975,1941,1968]" collectionCode="BOFWI" pageId="22" pageNumber="23">BOFWI055-11</specimenCode>
, …, BOFWI059-11).
<collectorName id="2A59537BFFFA9E3941A2F832EBA3F823" box="[1220,1264,1941,1967]" pageId="22" pageNumber="23">The</collectorName>
BIN is BOLD: ABU8653.
<collectorName id="2A59537BFFFA9E394471F81FEE10F85E" box="[279,323,1976,2002]" pageId="22" pageNumber="23">The</collectorName>
closest neighbour is
<taxonomicName id="40AC4D2EFFFA9E39474DF81EED96F85E" baseAuthorityName="Mocsary" baseAuthorityYear="1884" box="[555,709,1976,2002]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis id="B5D8EABFFFFA9E39474DF81EED96F85E" box="[555,709,1976,2002]" italics="true" pageId="22" pageNumber="23">P. reticulatum</emphasis>
</taxonomicName>
with a high barcode gap of 11.53% and a high average intergroup distance of 12.10%.
</paragraph>
</subSubSection>
<caption id="D3D36625FFFB9E3845F1F8D3EE99F874" ID-DOI="http://doi.org/10.5281/zenodo.14240761" ID-Zenodo-Dep="14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1908,1932]" targetBox="[152,1436,153,1890]" targetPageId="23" targetType="figure">
<paragraph id="871336ADFFFB9E3845F1F8D3EE99F874" blockId="23.[151,1436,1908,2040]" pageId="23" pageNumber="24">
<emphasis id="B5D8EABFFFFB9E3845F1F8D3EE72F800" bold="true" box="[151,289,1908,1932]" pageId="23" pageNumber="24">FIGURE 12.</emphasis>
<taxonomicName id="40AC4D2EFFFB9E38444EF8D3EDADF800" authority="(Friese, 1917)" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[296,766,1908,1932]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rhombiferum">
<emphasis id="B5D8EABFFFFB9E38444EF8D3ED36F800" box="[296,613,1908,1932]" italics="true" pageId="23" pageNumber="24">Pseudoanthidium rhombiferum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFB9E384715F8D3EDA4F800" author="Friese, H." box="[627,759,1908,1932]" pageId="23" pageNumber="24" pagination="49 - 60" refId="ref29819" refString="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Palaarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49 - 60." type="journal article" year="1917">Friese, 1917</bibRefCitation>
)
</taxonomicName>
. a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine).
</paragraph>
</caption>
<subSubSection id="CFB66526FFF49E3745A1FF30EF84FECD" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="871336ADFFF49E3745A1FF30EF84FECD" blockId="24.[151,1437,150,1149]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1FF30EE6AFF3C" box="[199,313,151,176]" italics="true" pageId="24" pageNumber="25">Diagnosis</emphasis>
. The female is small (
<quantity id="40549B48FFF49E374750FF31EDDCFF3D" box="[566,655,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="24" pageNumber="25" unit="mm" value="6.5">6.5 mm</quantity>
) with a black clypeus and a strongly protruding apical ridge with six tubercles; dark species with femora apically, tibiae and tarsi red-brown; most similar species is
<taxonomicName id="40AC4D2EFFF49E3741ACFF1CEA11FF59" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[1226,1346,187,213]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFF49E3741ACFF1CEA11FF59" box="[1226,1346,187,213]" italics="true" pageId="24" pageNumber="25">P. alpinum</emphasis>
</taxonomicName>
, which, however, has a convexly protruding apical ridge of the clypeus (straight in
<taxonomicName id="40AC4D2EFFF49E37469BFF78EBFEFF75" baseAuthorityName="Friese" baseAuthorityYear="1917" box="[1021,1197,223,249]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rhombiferum">
<emphasis id="B5D8EABFFFF49E37469BFF78EBFEFF75" box="[1021,1197,223,249]" italics="true" pageId="24" pageNumber="25">P. rhombiferum</emphasis>
</taxonomicName>
). The male T6, with two uneven lateral projections and depressions, is unique in the genus, making it impossible to confuse it with any other.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745A1FEECEC2EFBD5" pageId="24" pageNumber="25" type="description">
<paragraph id="871336ADFFF49E3745A1FEECED6AFD25" blockId="24.[151,1437,150,1149]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1FEECEE18FEE8" box="[199,331,331,356]" italics="true" pageId="24" pageNumber="25">Description</emphasis>
.
<emphasis id="B5D8EABFFFF49E37443CFEECEEFDFEE9" box="[346,430,331,357]" italics="true" pageId="24" pageNumber="25">Female</emphasis>
(
<figureCitation id="1F972A28FFF49E3744D9FEECED73FEE9" box="[447,544,331,357]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="24" pageNumber="25">Fig. 12a</figureCitation>
).
<quantity id="40549B48FFF49E374753FEEDEDF9FEE9" box="[565,682,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="24" pageNumber="25" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
.—
<emphasis id="B5D8EABFFFF49E3747ACFEECEC54FEE9" box="[714,775,331,357]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
: Black with a triangular spot behind the zenith of the eye; anterior half of the clypeus scattered punctate with polished interstices, posterior half with fine, rugulose punctation; apical ridge strongly protruding, smooth surface with three shallow tubercles on each side (
<figureCitation id="1F972A28FFF49E3741A2FE34EA4FFE21" box="[1220,1308,403,429]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="24" pageNumber="25">Fig. 20</figureCitation>
); mandible chestnut-brown, six black teeth (to2to5 almost equally strong); sometimes with yellow spot in lower paraocular area; antenna brown.—
<taxonomicName id="40AC4D2EFFF49E3744C4FE7CED49FE78" box="[418,538,475,500]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF49E3744C4FE7CED49FE78" box="[418,538,475,500]" italics="true" pageId="24" pageNumber="25">Mesosoma</emphasis>
</taxonomicName>
: Black; dense and fine punctation; omaulus rounded; low lamella on pronotal lobe; scutellum black, in
<specimenCount id="91AAFD24FFF49E3744CFFE58ED13FD95" box="[425,576,511,537]" count="1" pageId="24" pageNumber="25" type="generic">one specimen</specimenCount>
with narrow yellow stripe.—
<emphasis id="B5D8EABFFFF49E3746E4FE58ECA4FD94" box="[898,1015,511,536]" italics="true" pageId="24" pageNumber="25">Metasoma</emphasis>
: T1T5 with ivory-coloured to yellow lateral oval spots, getting closer to the middle towards the apical terga; interstices shining; broad, polished apical margin; rugulose surface of T6, laterally bulging, apically with shallow grey protrusion and a small emargination to accommodate the sting; scopal hairs dark grey (
<figureCitation id="1F972A28FFF49E3747C1FDCCEC54FD09" box="[679,775,619,645]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="24" pageNumber="25">Fig. 12d</figureCitation>
,
<figureCitation id="1F972A28FFF49E374674FDCCEC61FD09" box="[786,818,619,645]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="24" pageNumber="25">22</figureCitation>
); S6 triangular.—
<emphasis id="B5D8EABFFFF49E374691FDCCEB7FFD08" box="[1015,1068,619,644]" italics="true" pageId="24" pageNumber="25">Legs</emphasis>
: Femora apically, tibiae and tarsi red-brown; hind basitarsus dark grey.
</paragraph>
<paragraph id="871336ADFFF49E3745A1FD14EC2EFBD5" blockId="24.[151,1437,150,1149]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1FD14EE52FD41" box="[199,257,691,717]" italics="true" pageId="24" pageNumber="25">Male</emphasis>
(
<figureCitation id="1F972A28FFF49E374477FD14EE27FD41" box="[273,372,691,717]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="24" pageNumber="25">Fig. 12b</figureCitation>
).
<quantity id="40549B48FFF49E3744EEFD14EE9FFD41" box="[392,460,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="mm" value="7.0">7 mm</quantity>
.—
<emphasis id="B5D8EABFFFF49E37448AFD14ED7AFD41" box="[492,553,691,717]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
: Black with lower paraocular area and clypeus ivory-white and a small ivory-white spot behind the eye; basal area of clypeus scattered punctate, with punctures denser and smaller towards apex; apical margin smooth and straight, covered by long white hairs (
<figureCitation id="1F972A28FFF49E3746C3FD5CECABFC99" box="[933,1016,763,789]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="24" pageNumber="25">Fig. 21</figureCitation>
); mandible chestnut-brown, polished, with three black teeth.—
<emphasis id="B5D8EABFFFF49E3744D0FCB8ED40FCB5" box="[438,531,799,825]" italics="true" pageId="24" pageNumber="25">Antenna</emphasis>
: sg1sg3 (sg4) black; sg5sg13 light brown.—
<taxonomicName id="40AC4D2EFFF49E37415CFCB8EBE1FCB4" box="[1082,1202,799,824]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF49E37415CFCB8EBE1FCB4" box="[1082,1202,799,824]" italics="true" pageId="24" pageNumber="25">Mesosoma</emphasis>
</taxonomicName>
: Scutum black with fine punctation; omaulus rounded; pronotal lobe inconspicuous with a low lamella; scutellum and axillae crescent-shaped in dorsal view; medially rounded and laterally angulate in profile.—
<emphasis id="B5D8EABFFFF49E37468FFCC0EB0DFC0C" box="[1001,1118,871,896]" italics="true" pageId="24" pageNumber="25">Metasoma</emphasis>
: T1T5 black, shining, with lateral pale yellow elongate spots, increasing in size towards the posterior; T6 deformed with a lateral and submedian projection, and two pocket-like depressions laterally; T7 very small, blunt, largely hidden under T6 (
<figureCitation id="1F972A28FFF49E37406EFC08EA3FFC45" box="[1288,1388,943,969]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="24" pageNumber="25">Figs 12e</figureCitation>
,
<figureCitation id="1F972A28FFF49E37401FFC08EAC4FC45" box="[1401,1431,943,969]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="24" pageNumber="25">23</figureCitation>
,
<figureCitation id="1F972A28FFF49E3745F1FC74EFE4FC61" box="[151,183,979,1005]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="24" pageNumber="25">24</figureCitation>
); S3 with a fan-like fringe of undulated and apically hooked hairs; S4 protruding with a median emargination; S5 with a submedian pair of small black combs (approximately 2.5 comb widths apart) and a pair of larger combs on the apex of lateral arms (
<figureCitation id="1F972A28FFF49E3744B7FBBCED78FBB9" box="[465,555,1051,1077]" captionStart="FIGURE 12" captionStartId="23.[151,250,1908,1932]" captionTargetBox="[152,1436,153,1890]" captionTargetId="figure-15@23.[152,1436,153,1890]" captionTargetPageId="23" captionText="FIGURE 12. Pseudoanthidium rhombiferum (Friese, 1917). a. Habitus dorsal, female from Lebanon; b. Habitus dorsal, male holotype from Syria; c. Head of a female from Palestine; d. Apical terga of the female; e. Apical terga of the male (from Palestine); f. Apical sterna of the male with specialized undulated and apically hooked hairs on S3 and strong black combs on S5 (specimen from Palestine)." figureDoi="http://doi.org/10.5281/zenodo.14240761" httpUri="https://zenodo.org/record/14240761/files/figure.png" pageId="24" pageNumber="25">Fig. 12f</figureCitation>
).—
<emphasis id="B5D8EABFFFF49E374731FBBCEDDFFBB8" box="[599,652,1051,1076]" italics="true" pageId="24" pageNumber="25">Legs</emphasis>
: Dark brown and black with dirty-yellow and reddish-brown spots.—
<emphasis id="B5D8EABFFFF49E3745F1FB98EE52FBD5" box="[151,257,1087,1113]" italics="true" pageId="24" pageNumber="25">Genitalia</emphasis>
: The male gonostyli are robust and pincer-like (
<figureCitation id="1F972A28FFF49E37467AFB98EC23FBD5" box="[796,880,1086,1113]" captionStart="FIGURE 25" captionStartId="40.[151,250,2009,2033]" captionTargetBox="[176,1412,1240,1996]" captionTargetId="figure-103@40.[176,1412,1240,2001]" captionTargetPageId="40" captionText="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240785" httpUri="https://zenodo.org/record/14240785/files/figure.png" pageId="24" pageNumber="25">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF49E3745A1FBC4EA1AFBF1" box="[199,1353,1123,1149]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="871336ADFFF49E3745A1FBC4EA1AFBF1" blockId="24.[151,1437,150,1149]" box="[199,1353,1123,1149]" pageId="24" pageNumber="25">
<emphasis id="B5D8EABFFFF49E3745A1FBC4EE03FBF1" box="[199,336,1123,1149]" italics="true" pageId="24" pageNumber="25">Distribution</emphasis>
.
<collectingCountry id="FFBB763DFFF49E374438FBC4EECCFBF1" box="[350,415,1123,1149]" name="Israel" pageId="24" pageNumber="25">Israel</collectingCountry>
,
<collectingCountry id="FFBB763DFFF49E3744CCFBC4ED5CFBF1" box="[426,527,1123,1149]" name="Lebanon" pageId="24" pageNumber="25">Lebanon</collectingCountry>
,
<collectingCountry id="FFBB763DFFF49E37477CFBC4EDD1FBF1" box="[538,642,1123,1149]" name="Palestine" pageId="24" pageNumber="25">Palestine</collectingCountry>
, and
<collectingCountry id="FFBB763DFFF49E3747DBFBC4EDABFBF1" box="[701,760,1123,1149]" name="Syria" pageId="24" pageNumber="25">Syria</collectingCountry>
(
<figureCitation id="1F972A28FFF49E374661FBC4EC3AFBF1" box="[775,873,1123,1149]" captionStart="FIGURE 27" captionStartId="42.[151,250,1691,1715]" captionTargetBox="[151,1435,273,1667]" captionTargetId="figure-71@42.[151,1435,273,1667]" captionTargetPageId="42" captionText="FIGURE 27. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium bytinskii, P. puncticolle, P. syriacum sp. nov., and P. variabile; b. Pseudoanthidium farsiense sp. nov., P. moricei, and P. rhombiferum." figureDoi="http://doi.org/10.5281/zenodo.14240790" httpUri="https://zenodo.org/record/14240790/files/figure.png" pageId="24" pageNumber="25">Fig. 27b</figureCitation>
). Confirmation required for
<collectingCountry id="FFBB763DFFF49E3741C0FBC4EA17FBF1" box="[1190,1348,1123,1149]" name="Turkmenistan" pageId="24" pageNumber="25">Turkmenistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B5D8EABFFFFA9E3945F1FD5AED83FC9B" bold="true" box="[151,720,764,792]" pageId="22" pageNumber="23">
<emphasis id="B5D8EABFFFFA9E3945F1FD5AED53FC94" bold="true" box="[151,512,765,792]" italics="true" pageId="22" pageNumber="23">Pseudoanthidium reticulatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFA9E394769FD5BED94FC9B" author="Mocsary, A." box="[527,711,764,791]" pageId="22" pageNumber="23" pagination="241 - 278" refId="ref30778" refString="Mocsary, A. (1884) Species generis Anthidium Fabr. regionis Palaearcticae. Termeszetrajzi Fuzetek Kiadija a Magyar Nemzeti Muzeum [Naturhistorische Hefte herausgegeben vom Ungarischen National-Museum], 8, 241 - 278." type="journal article" year="1884">Mocsáry, 1884</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFA9E3945F1FC86ED29FC2F" pageId="22" pageNumber="23" type="description">
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(
<figureCitation id="1F972A28FFFA9E3945F9FC86EFB5FCB7" box="[159,230,801,827]" captionStart="FIGURE 1" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1843]" captionTargetId="figure-15@3.[151,1435,181,1843]" captionTargetPageId="3" captionText="FIGURE 1. Pseudoanthidium alpinum (Morawitz, 1873). a. Habitus dorsal, female (Greece); b. Habitus dorsal, male (Greece); c. Habitus lateral, female (Romania, holotype of Anthidium serraticeps Friese, 1917, a junior synonym of P. alpinum). d. Apical terga and scopa, female (Kazakhstan). e. Ventral side of the metasoma of the male with felt-like hairs, specialized hairs on S3, and combs on S5 (Spain)." figureDoi="http://doi.org/10.5281/zenodo.14240736" httpUri="https://zenodo.org/record/14240736/files/figure.png" pageId="22" pageNumber="23">Figs 1</figureCitation>
,
<figureCitation id="1F972A28FFFA9E394592FC86EE40FCB7" box="[244,275,801,827]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="22" pageNumber="23">20</figureCitation>
,
<figureCitation id="1F972A28FFFA9E394478FC86EE6FFCB7" box="[286,316,801,827]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="22" pageNumber="23">21</figureCitation>
,
<figureCitation id="1F972A28FFFA9E39442EFC86EE34FCB7" box="[328,359,801,827]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="22" pageNumber="23">22</figureCitation>
,
<figureCitation id="1F972A28FFFA9E394414FC86EEC3FCB7" box="[370,400,801,827]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="22" pageNumber="23">23</figureCitation>
,
<figureCitation id="1F972A28FFFA9E3944FAFC86EEE9FCB7" box="[412,442,801,827]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="22" pageNumber="23">24</figureCitation>
, 25)
</paragraph>
<paragraph id="871336ADFFFA9E3945F1FCC0ED29FC2F" blockId="22.[151,1437,871,931]" pageId="22" pageNumber="23">This species shows a conspicuous variation across its distribution area in the Mediterranean and beyond. A more detailed study will be published separately.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,633 @@
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<heading id="DC5B81C1FFFE9E3D45F1FE35ED91FE20" bold="true" box="[151,706,402,429]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FE35ED91FE20" ID-CoL="8SFXP" authority="(Perez, 1895)" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[151,706,402,429]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum">
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<emphasis id="B5D8EABFFFFE9E3D45F1FE35ED4AFE21" bold="true" box="[151,537,402,429]" italics="true" pageId="18" pageNumber="19">Pseudoanthidium octodentatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFE9E3D474EFE35EDE9FE20" author="Perez, J." box="[552,698,402,429]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
)
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<paragraph id="871336ADFFFE9E3D45F1FE10EEAAFE5D" blockId="18.[151,706,402,465]" box="[151,505,438,465]" pageId="18" pageNumber="19">
(
<figureCitation id="1F972A28FFFE9E3D45F9FE10EFA5FE5D" box="[159,246,439,465]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="18" pageNumber="19">Figs 10</figureCitation>
,
<figureCitation id="1F972A28FFFE9E3D4464FE10EE72FE5D" box="[258,289,439,465]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="18" pageNumber="19">20</figureCitation>
,
<figureCitation id="1F972A28FFFE9E3D444AFE10EE19FE5D" box="[300,330,439,465]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="18" pageNumber="19">21</figureCitation>
,
<figureCitation id="1F972A28FFFE9E3D4430FE10EE26FE5D" box="[342,373,439,465]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="18" pageNumber="19">22</figureCitation>
,
<figureCitation id="1F972A28FFFE9E3D44E6FE10EECDFE5D" box="[384,414,439,465]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="18" pageNumber="19">23</figureCitation>
,
<figureCitation id="1F972A28FFFE9E3D44CCFE10EE9BFE5D" box="[426,456,439,465]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="18" pageNumber="19">24</figureCitation>
, 25)
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFE9E3D45F1FE5CEC66FD21" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="871336ADFFFE9E3D45F1FE5CEDE9FD9F" blockId="18.[151,1005,507,685]" box="[151,698,507,531]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FE5CED5CFD9F" ID-CoL="8SG4W" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[151,527,507,531]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum">
<emphasis id="B5D8EABFFFFE9E3D45F1FE5CEEC2FD9F" box="[151,401,507,531]" italics="true" pageId="18" pageNumber="19">Anthidium octodentatum</emphasis>
<bibRefCitation id="E33D4B5CFFFE9E3D44F1FE5CED5CFD9F" author="Perez, J." box="[407,527,507,531]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
</taxonomicName>
.—Male,
<collectingCountry id="FFBB763DFFFE9E3D4708FE5CEDE9FD9F" box="[622,698,507,531]" name="Algeria" pageId="18" pageNumber="19">Algeria</collectingCountry>
</paragraph>
<paragraph id="871336ADFFFE9E3D45F1FDBDEC29FDBE" blockId="18.[151,1005,507,685]" box="[151,890,538,562]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FDBDED1EFDBE" ID-CoL="8SNMJ" authority="Dusmet y Alonso, 1915" authorityName="Dusmet y Alonso" authorityYear="1915" box="[151,589,538,562]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capapei">
<emphasis id="B5D8EABFFFFE9E3D45F1FDBDEE04FDBD" box="[151,343,538,562]" italics="true" pageId="18" pageNumber="19">Anthidium capapei</emphasis>
Dusmet y
<bibRefCitation id="E33D4B5CFFFE9E3D44A2FDBDED1EFDBE" author="Alonso, J. M." box="[452,589,538,562]" pageId="18" pageNumber="19" pagination="293 - 334" refId="ref29722" refString="Dusmet y Alonso, J. M. (1915) Apidos de Marruecos de los gen. Anthidium, Nomada, Melecta, Crocisa, Coelioxys y Phiarus. Memorias de la Real Sociedad Espanola de Historia Natural, 8, 293 - 334." type="journal article" year="1915">Alonso, 1915</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="FFBB763DFFFE9E3D473CFDBDEDEAFDBE" box="[602,697,538,562]" name="Morocco" pageId="18" pageNumber="19">Morocco</collectingCountry>
).—
<bibRefCitation id="E33D4B5CFFFE9E3D47BBFDBDEC29FDBE" author="Pasteels, J. J." box="[733,890,538,562]" pageId="18" pageNumber="19" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
</paragraph>
<paragraph id="871336ADFFFE9E3D45F1FD9FEDA7FDDC" blockId="18.[151,1005,507,685]" box="[151,756,568,593]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FD9FED64FDDC" ID-CoL="8S728" authority="(Perez, 1895)" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[151,567,568,593]" class="Insecta" family="Megachilidae" genus="Neanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum">
<emphasis id="B5D8EABFFFFE9E3D45F1FD9FEEF9FDDC" box="[151,426,568,592]" italics="true" pageId="18" pageNumber="19">Neanthidium octodentatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFE9E3D44DEFD9EED63FDDC" author="Perez, J." box="[440,560,568,593]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFFE9E3D4733FD9FEDA7FDDC" author="Pasteels, J. J." box="[597,756,568,592]" pageId="18" pageNumber="19" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
</paragraph>
<paragraph id="871336ADFFFE9E3D45F1FDF0ECD8FDE3" blockId="18.[151,1005,507,685]" box="[151,907,599,623]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FDF0ED97FDE3" ID-CoL="8SG4W" authority="(Perez, 1895)" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[151,708,599,623]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum" subGenus="Mesanthidium">
<emphasis id="B5D8EABFFFFE9E3D45F1FDF0EE52FDE3" box="[151,257,599,623]" italics="true" pageId="18" pageNumber="19">Anthidium</emphasis>
(
<emphasis id="B5D8EABFFFFE9E3D4476FDF0EEF2FDE3" box="[272,417,599,623]" italics="true" pageId="18" pageNumber="19">Mesanthidium</emphasis>
)
<emphasis id="B5D8EABFFFFE9E3D44C9FDF0ED6AFDE3" box="[431,569,599,623]" italics="true" pageId="18" pageNumber="19">octodentatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFE9E3D4720FDF0EDEDFDE3" author="Perez, J." box="[582,702,599,623]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFFE9E3D4784FDF0ECD8FDE3" author="Warncke, K." box="[738,907,599,623]" pageId="18" pageNumber="19" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
</paragraph>
<paragraph id="871336ADFFFE9E3D45F1FDD1ECBEFD02" blockId="18.[151,1005,507,685]" box="[151,1005,630,654]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FDD1EDACFD02" ID-CoL="8SFXP" authority="(Perez, 1895)" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[151,767,630,654]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum" subGenus="Neanthidium">
<emphasis id="B5D8EABFFFFE9E3D45F1FDD1EE1BFD02" box="[151,328,630,654]" italics="true" pageId="18" pageNumber="19">Pseudoanthidium</emphasis>
(
<emphasis id="B5D8EABFFFFE9E3D4431FDD1EE89FD02" box="[343,474,630,654]" italics="true" pageId="18" pageNumber="19">Neanthidium</emphasis>
)
<emphasis id="B5D8EABFFFFE9E3D448FFDD1ED20FD02" box="[489,627,630,654]" italics="true" pageId="18" pageNumber="19">octodentatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFE9E3D47E6FDD1EDAAFD02" author="Perez, J." box="[640,761,630,654]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="E33D4B5CFFFE9E3D467BFDD1ECBEFD02" author="Litman, J. R. &amp; Griswold, T. &amp; Danforth, B. N." box="[797,1005,630,654]" pageId="18" pageNumber="19" pagination="183 - 198" refId="ref30407" refString="Litman, J. R., Griswold, T. &amp; Danforth, B. N. (2016) Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution, 100, 183 - 198. https: // doi. org / 10.1016 / j. ympev. 2016.03.018" type="journal article" year="2016">
Litman
<emphasis id="B5D8EABFFFFE9E3D4617FDD0ECF7FD02" box="[881,932,630,654]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2016)
</bibRefCitation>
</paragraph>
<paragraph id="871336ADFFFE9E3D45F1FD33EC66FD21" blockId="18.[151,1005,507,685]" box="[151,821,660,685]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45F1FD33ED1DFD21" ID-CoL="8SPF4" authority="Pasteels, 1981" authorityName="Pasteels" authorityYear="1981" box="[151,590,660,685]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicoloripenne">
<emphasis id="B5D8EABFFFFE9E3D45F1FD33EEEAFD20" box="[151,441,660,684]" italics="true" pageId="18" pageNumber="19">Royanthidium bicoloripenne</emphasis>
Pasteels, 1981
</taxonomicName>
.—
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,
<emphasis id="B5D8EABFFFFE9E3D47B1FD32EC66FD21" bold="true" box="[727,821,661,685]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="AEEB57C4FFFE9E3D47B1FD32EC66FD21" box="[727,821,661,685]" pageId="18" pageNumber="19">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<collectingCounty id="6E724E21FFFE9E3D45F1FD7CEFA5FD79" box="[151,246,731,757]" pageId="18" pageNumber="19">Material</collectingCounty>
examined (all
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):
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,
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(
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,
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leg. (
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,
<date id="F312106DFFFE9E3D46E0FD58EB52FC95" box="[902,1025,767,793]" pageId="18" pageNumber="19" value="1986-03-26">
<collectingDate id="E356E985FFFE9E3D46E0FD58EB52FC95" box="[902,1025,767,793]" pageId="18" pageNumber="19" value="1986-03-26">26.iii.1986</collectingDate>
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,
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leg. (
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: ms2464).—
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,
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N Foum Zguid
</locationDeviation>
,
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, -
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,
<date id="F312106DFFFE9E3D47EAFC84EC5BFCB1" box="[652,776,803,829]" pageId="18" pageNumber="19" value="1986-03-30">
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</date>
,
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leg. (
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: ms2465).—
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,
<locationDeviation id="96A12F8FFFFE9E3D418AFC84EF8EFCED" location="Foum Zguid" pageId="18" pageNumber="19">
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N Foum Zguid
</locationDeviation>
,
<geoCoordinate id="E298506AFFFE9E3D458CFCE0EE13FCED" box="[234,320,839,865]" degrees="30.63" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="30.63">30.63N</geoCoordinate>
, -
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,
<date id="F312106DFFFE9E3D44CFFCE0ED76FCED" box="[425,549,839,865]" pageId="18" pageNumber="19" value="1986-03-31">
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,
<collectorName id="2A59537BFFFE9E3D4754FCE0EDE8FCED" box="[562,699,839,865]" pageId="18" pageNumber="19">M. Schwarz</collectorName>
leg. (
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: ms2466).—
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, Draa-Taf.,
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.
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leg. (coll.
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: tjw316).—
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, Draa-Taf.,
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,
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.
<collectorName id="2A59537BFFFE9E3D467BFC28EC0CFC25" box="[797,863,911,937]" pageId="18" pageNumber="19">Wood</collectorName>
leg.; COI sequence: ABABZ089-23 (coll.
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: tjw317).—
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,
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<location id="82736076FFFE9E3D448FFC14ED04FC41" LSID="urn:lsid:plazi:treatment:0F0587BBFFFE9E3B45F1FE35EADAFB9D:82736076FFFE9E3D448FFC14ED04FC41" box="[489,599,947,973]" country="Morocco" county="Material" latitude="30.9" longLatPrecision="731" longitude="6.59" municipality="Tizi-n-Tinififft" name="Tafraoute" pageId="18" pageNumber="19" stateProvince="Souss-Massa-Draa">Tafraoute</location>
,
<location id="82736076FFFE9E3D4702FC14EC51FC41" LSID="urn:lsid:plazi:treatment:0F0587BBFFFE9E3B45F1FE35EADAFB9D:82736076FFFE9E3D4702FC14EC51FC41" box="[612,770,947,973]" country="Morocco" county="Material" latitude="30.9" longLatPrecision="731" longitude="6.59" municipality="Tizi-n-Tinififft" name="Azrou Ouado" pageId="18" pageNumber="19" stateProvince="Souss-Massa-Draa">Azrou Ouado</location>
,
<quantity id="40549B48FFFE9E3D4677FC14EC10FC41" box="[785,835,947,973]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="km" value="2.0">2km</quantity>
W,
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, -
<geoCoordinate id="E298506AFFFE9E3D4685FC14EB79FC41" box="[995,1066,947,973]" degrees="9.07" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="9.07">9.07E</geoCoordinate>
,
<date id="F312106DFFFE9E3D415FFC14EBE6FC41" box="[1081,1205,947,973]" pageId="18" pageNumber="19" value="2022-03-13">
<collectingDate id="E356E985FFFE9E3D415FFC14EBE6FC41" box="[1081,1205,947,973]" pageId="18" pageNumber="19" value="2022-03-13">13.iii.2022</collectingDate>
</date>
,
<collectionCode id="E1BDAE68FFFE9E3D41A5FC14EB86FC41" box="[1219,1237,947,973]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
.
<collectorName id="2A59537BFFFE9E3D4185FC14EA76FC41" box="[1251,1317,947,973]" pageId="18" pageNumber="19">Wood</collectorName>
leg.; COI sequence: ABABZ088-23 (coll.
<collectorName id="2A59537BFFFE9E3D4764FC70ED1AFC7D" box="[514,585,983,1009]" pageId="18" pageNumber="19">Wood</collectorName>
: tjw318).—
<specimenCount id="91AAFD24FFFE9E3D47B0FC70EDA0FC7D" box="[726,755,983,1009]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
, Draa-Taf.,
<collectorName id="2A59537BFFFE9E3D461FFC70ECB5FC7D" box="[889,998,982,1009]" pageId="18" pageNumber="19">Midelt, R</collectorName>
503,
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<quantity id="40549B48FFFE9E3D417BFC70EB1DFC7D" box="[1053,1102,983,1009]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="km" value="7.0">7km</quantity>
NE Ait Ben Yacoub
</locationDeviation>
,
<geoCoordinate id="E298506AFFFE9E3D4024FC70EACBFC7D" box="[1346,1432,983,1009]" degrees="33.30" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="33.3">33.30N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFFE9E3D45F9FC5CEFB6FB99" box="[159,229,1019,1045]" degrees="4.69" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="4.69">4.69E</geoCoordinate>
,
<date id="F312106DFFFE9E3D4594FC5CEE30FB99" box="[242,355,1019,1045]" pageId="18" pageNumber="19" value="2022-05-21">
<collectingDate id="E356E985FFFE9E3D4594FC5CEE30FB99" box="[242,355,1019,1045]" pageId="18" pageNumber="19" value="2022-05-21">21.v.2022</collectingDate>
</date>
,
<collectionCode id="E1BDAE68FFFE9E3D4409FC5CEED2FB99" box="[367,385,1019,1045]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
.
<collectorName id="2A59537BFFFE9E3D44EBFC5CEE9CFB99" box="[397,463,1019,1045]" pageId="18" pageNumber="19">Wood</collectorName>
leg.; COI sequence: ABABZ090-23 (coll.
<collectorName id="2A59537BFFFE9E3D46DEFC5CECACFB99" box="[952,1023,1019,1045]" pageId="18" pageNumber="19">Wood</collectorName>
: tjw319).—
<specimenCount id="91AAFD24FFFE9E3D41EBFC5CEBF9FB99" box="[1165,1194,1019,1045]" count="1" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectingRegion id="4568F84FFFFE9E3D41D1FC5CEACBFB99" box="[1207,1432,1019,1045]" country="Morocco" name="Souss-Massa-Draa" pageId="18" pageNumber="19">Souss-Massa-Daraâ</collectingRegion>
,
<location id="82736076FFFE9E3D45F1FBB8EEC1FBB5" LSID="urn:lsid:plazi:treatment:0F0587BBFFFE9E3B45F1FE35EADAFB9D:82736076FFFE9E3D45F1FBB8EEC1FBB5" box="[151,402,1055,1081]" country="Morocco" county="Material" latitude="30.9" longLatPrecision="731" longitude="6.59" municipality="Tizi-n-Tinififft" name="Biougra-Imi Mqouren" pageId="18" pageNumber="19" stateProvince="Souss-Massa-Draa">Biougra-Imi Mqouren</location>
,
<locationDeviation id="96A12F8FFFFE9E3D44FAFBB9EDC7FBB5" box="[412,660,1054,1081]" location="Imi Mqourn" pageId="18" pageNumber="19">
<quantity id="40549B48FFFE9E3D44FAFBB9EEB0FBB5" box="[412,483,1054,1081]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="km" value="1.5">1.5km</quantity>
W Imi Mqourn
</locationDeviation>
,
<geoCoordinate id="E298506AFFFE9E3D47F9FBB8EDA6FBB5" box="[671,757,1055,1081]" degrees="30.17" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="30.17">30.17N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFFE9E3D466EFBB9EC1DFBB5" box="[776,846,1054,1081]" degrees="9.25" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="9.25">9.25E</geoCoordinate>
,
<date id="F312106DFFFE9E3D463FFBB8EC81FBB5" box="[857,978,1055,1081]" pageId="18" pageNumber="19" value="2014-04-20">
<collectingDate id="E356E985FFFE9E3D463FFBB8EC81FBB5" box="[857,978,1055,1081]" pageId="18" pageNumber="19" value="2014-04-20">20.iv.2014</collectingDate>
</date>
,
<collectorName id="2A59537BFFFE9E3D46BBFBB8EB64FBB5" box="[989,1079,1055,1081]" pageId="18" pageNumber="19">R. Prosi</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFE9E3D4113FBB8EBE9FBB5" box="[1141,1210,1055,1081]" pageId="18" pageNumber="19">CMK</collectionCode>
: rpr016).—
<specimenCount id="91AAFD24FFFE9E3D402FFBB8EA33FBB5" box="[1353,1376,1055,1081]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
, K17
<location id="82736076FFFE9E3D45F1FBE4EE11FBD1" LSID="urn:lsid:plazi:treatment:0F0587BBFFFE9E3B45F1FE35EADAFB9D:82736076FFFE9E3D45F1FBE4EE11FBD1" box="[151,322,1091,1117]" country="Morocco" county="Material" latitude="30.9" longLatPrecision="731" longitude="6.59" municipality="Tizi-n-Tinififft" name="Quarz-Zag Rd." pageId="18" pageNumber="19" stateProvince="Souss-Massa-Draa">Quarz-Zag Rd.</location>
,
<geoCoordinate id="E298506AFFFE9E3D4429FBE4EEF6FBD1" box="[335,421,1091,1117]" degrees="30.90" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="30.9">30.90N</geoCoordinate>
, -
<geoCoordinate id="E298506AFFFE9E3D44DEFBE5EEADFBD1" box="[440,510,1090,1117]" degrees="6.59" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="555" value="6.59">6.59E</geoCoordinate>
,
<date id="F312106DFFFE9E3D476DFBE5EDD7FBD1" box="[523,644,1090,1117]" pageId="18" pageNumber="19" value="1985-06-01">
<collectingDate id="E356E985FFFE9E3D476DFBE5EDD7FBD1" box="[523,644,1090,1117]" pageId="18" pageNumber="19" value="1985-06-01">01.vi.1985</collectingDate>
</date>
,
<collectorName id="2A59537BFFFE9E3D47F7FBE4EC4FFBD1" box="[657,796,1091,1117]" pageId="18" pageNumber="19">K. Guichard</collectorName>
(
<collectionCode id="E1BDAE68FFFE9E3D464AFBE4ECCFFBD1" box="[812,924,1091,1117]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="19" type="Museum">NHMUK</collectionCode>
: nhmuk004).—
<typeStatus id="5817880FFFFE9E3D412FFBE4EBE0FBD1" box="[1097,1203,1091,1117]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="40AC4D2EFFFE9E3D41BCFBE4EAC5FBD1" authorityName="Pasteels" authorityYear="1981" box="[1242,1430,1091,1117]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicoloripenne">
<emphasis id="B5D8EABFFFFE9E3D41BCFBE4EAC5FBD1" box="[1242,1430,1091,1117]" italics="true" pageId="18" pageNumber="19">R. bicoloripenne</emphasis>
</taxonomicName>
,
<specimenCount id="91AAFD24FFFE9E3D45F1FBC0EFE9FB0D" box="[151,186,1127,1153]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
, Agadir,
<date id="F312106DFFFE9E3D4447FBC0EEC8FB0D" box="[289,411,1127,1153]" pageId="18" pageNumber="19" value="1974-06-19">
<collectingDate id="E356E985FFFE9E3D4447FBC0EEC8FB0D" box="[289,411,1127,1153]" pageId="18" pageNumber="19" value="1974-06-19">19.vi.1974</collectingDate>
</date>
,
<collectorName id="2A59537BFFFE9E3D44C1FBC0ED62FB0D" box="[423,561,1127,1153]" pageId="18" pageNumber="19">K. Guichard</collectorName>
&amp;
<collectorName id="2A59537BFFFE9E3D4733FBC0EDD6FB0D" box="[597,645,1127,1153]" pageId="18" pageNumber="19">Else</collectorName>
leg. (
<collectionCode id="E1BDAE68FFFE9E3D47A2FBC0EC67FB0D" box="[708,820,1127,1153]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="19" type="Museum">NHMUK</collectionCode>
:
<collectorName id="2A59537BFFFE9E3D4659FBC0ECA6FB0D" box="[831,1013,1127,1153]" pageId="18" pageNumber="19">B.M. Type Hym</collectorName>
17a.317,
<specimenCode id="D70A9ED6FFFE9E3D4102FBC0EA04FB0D" box="[1124,1367,1126,1153]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="19" type="Museum">NHMUK 014026652</specimenCode>
).
</paragraph>
<paragraph id="871336ADFFFE9E3D45A1FB2CEA27FA9D" blockId="18.[151,1437,730,1945]" pageId="18" pageNumber="19">
<emphasis id="B5D8EABFFFFE9E3D45A1FB2CEE95FB29" box="[199,454,1163,1189]" italics="true" pageId="18" pageNumber="19">Material not examined</emphasis>
:
<collectingCountry id="FFBB763DFFFE9E3D44B2FB2CED00FB29" box="[468,595,1163,1189]" name="Algeria" pageId="18" pageNumber="19">ALGERIA</collectingCountry>
: Takaina (
<typeStatus id="5817880FFFFE9E3D47A2FB2CEDA5FB29" box="[708,758,1163,1189]" pageId="18" pageNumber="19">type</typeStatus>
locality; locality not identified; locality attributed to Algeria by
<bibRefCitation id="E33D4B5CFFFE9E3D45D8FB08EE1BFB45" author="Friese, H." box="[190,328,1199,1225]" pageId="18" pageNumber="19" refId="ref29768" refString="Friese, H. (1898) Die Bienen Europa's (Apidae europaeae) nach ihren Gattungen, Arten und Varietaten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitare Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp." type="book" year="1898">Friese 1898</bibRefCitation>
),
<date id="F312106DFFFE9E3D4439FB08EEBAFB45" box="[351,489,1198,1225]" pageId="18" pageNumber="19" value="1895-03">March 1895</date>
.—
<collectingCountry id="FFBB763DFFFE9E3D476CFB09EDCCFB45" box="[522,671,1198,1225]" name="Morocco" pageId="18" pageNumber="19">MOROCCO</collectingCountry>
: Marraquesh [Marrakesh],
<date id="F312106DFFFE9E3D46BFFB08EB17FB45" box="[985,1092,1199,1225]" pageId="18" pageNumber="19" value="1907-04-01">1.iv.1907</date>
, Dusmet y
<bibRefCitation id="E33D4B5CFFFE9E3D41ADFB08EA22FB45" author="Alonso, J. M." box="[1227,1393,1198,1225]" pageId="18" pageNumber="19" pagination="293 - 334" refId="ref29722" refString="Dusmet y Alonso, J. M. (1915) Apidos de Marruecos de los gen. Anthidium, Nomada, Melecta, Crocisa, Coelioxys y Phiarus. Memorias de la Real Sociedad Espanola de Historia Natural, 8, 293 - 334." type="journal article" year="1915">Alonso (1915)</bibRefCitation>
(as
<taxonomicName id="40AC4D2EFFFE9E3D45F1FB74EE3CFB60" authorityName="Dusmet y Alonso" authorityYear="1915" box="[151,367,1235,1261]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capapei">
<emphasis id="B5D8EABFFFFE9E3D45F1FB74EE3CFB60" box="[151,367,1235,1261]" italics="true" pageId="18" pageNumber="19">Anthidium capapei</emphasis>
</taxonomicName>
). Agadir,
<date id="F312106DFFFE9E3D4482FB74ED0FFB61" box="[484,604,1235,1261]" pageId="18" pageNumber="19" value="1974-06-18">18.vi.1974</date>
(Pasteels 1981) (as
<taxonomicName id="40AC4D2EFFFE9E3D462FFB74EB50FB61" authorityName="Kasparek" authorityYear="2024" box="[841,1027,1235,1261]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicoloripenne" status="sp. n.">
<emphasis id="B5D8EABFFFFE9E3D462FFB74EB50FB61" box="[841,1027,1235,1261]" italics="true" pageId="18" pageNumber="19">P. bicoloripenne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEEB57C4FFFE9E3D416AFB74EB1EFB61" box="[1036,1101,1235,1261]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
). Igerm [Igherm],
<date id="F312106DFFFE9E3D4040FB74EACEFB61" box="[1318,1437,1235,1261]" pageId="18" pageNumber="19" value="2003-04-28">28.iv.2003</date>
(
<bibRefCitation id="E33D4B5CFFFE9E3D45F9FB50EE22FA9D" author="Litman, J. R. &amp; Griswold, T. &amp; Danforth, B. N." box="[159,369,1271,1297]" pageId="18" pageNumber="19" pagination="183 - 198" refId="ref30407" refString="Litman, J. R., Griswold, T. &amp; Danforth, B. N. (2016) Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution, 100, 183 - 198. https: // doi. org / 10.1016 / j. ympev. 2016.03.018" type="journal article" year="2016">
Litman
<emphasis id="B5D8EABFFFFE9E3D459EFB50EE62FA9D" box="[248,305,1271,1297]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2016
</bibRefCitation>
).—
<collectingCountry id="FFBB763DFFFE9E3D44FCFB50EEA1FA9D" box="[410,498,1271,1297]" name="Libya" pageId="18" pageNumber="19">LIBYA</collectingCountry>
: Jebel Soda [Jabal as Sawdā], Wadi Ghodaifa, 03.-
<date id="F312106DFFFE9E3D414BFB51EBE1FA9D" box="[1069,1202,1270,1297]" pageId="18" pageNumber="19" value="1952-04-03">04.03.1952</date>
(
<bibRefCitation id="E33D4B5CFFFE9E3D41A6FB50EA34FA9D" author="Warncke, K." box="[1216,1383,1271,1297]" pageId="18" pageNumber="19" pagination="119 - 209" refId="ref31828" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke 1980</bibRefCitation>
).
</paragraph>
<paragraph id="871336ADFFFE9E3D45A1FABCEE7AFA2D" blockId="18.[151,1437,730,1945]" pageId="18" pageNumber="19">
A record from
<collectingCountry id="FFBB763DFFFE9E3D4408FABCEE83FAB9" box="[366,464,1307,1333]" name="Lebanon" pageId="18" pageNumber="19">Lebanon</collectingCountry>
(
<bibRefCitation id="E33D4B5CFFFE9E3D44B9FABCEDE7FAB9" author="Ornosa, C. &amp; Ortiz-Sanchez, F. J. &amp; Torres, F." box="[479,692,1307,1333]" pageId="18" pageNumber="19" pagination="61 - 86" refId="ref31152" refString="Ornosa, C., Ortiz-Sanchez, F. J. &amp; Torres, F. (2008) Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Graellsia, 64, 61 - 86. https: // doi. org / 10.3989 / graellsia. 2008. v 64. i 1.55" type="journal article" year="2008">
Ornosa
<emphasis id="B5D8EABFFFFE9E3D475FFABCED21FAB9" box="[569,626,1307,1333]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2008
</bibRefCitation>
; cited by
<bibRefCitation id="E33D4B5CFFFE9E3D4646FABCEB10FAB9" author="Ascher, J. S. &amp; Pickering, J." box="[800,1091,1307,1333]" pageId="18" pageNumber="19" refId="ref29495" refString="Ascher, J. S. &amp; Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: http: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species (accessed 1 August 20024)" type="url" year="2024">Ascher &amp; Pickering 2024</bibRefCitation>
;
<bibRefCitation id="E33D4B5CFFFE9E3D4128FABCEA61FAB9" author="Boustani, M. &amp; Dathe, H. &amp; Ghisbain, G. &amp; Kasparek, M. &amp; Michez, D. &amp; Nemer, N. &amp; Pauly, A. &amp; Risch, S. &amp; Terzo, M. &amp; Van Achter, X. &amp; Wood, T. &amp; Rasmont, P." box="[1102,1330,1307,1333]" pageId="18" pageNumber="19" pagination="1 - 146" refId="ref29624" refString="Boustani, M., Dathe, H., Ghisbain, G., Kasparek, M., Michez, D., Nemer, N., Pauly, A., Risch, S., Terzo, M., Van Achter, X., Wood, T. &amp; Rasmont, P. (2021) The bees of Lebanon (Hymenoptera: Apoidea: Anthophila). Zootaxa, 4976 (1), 1 - 146. https: // doi. org / 10.11646 / zootaxa. 4976.1.1" type="journal article" year="2021">
Boustani
<emphasis id="B5D8EABFFFFE9E3D41DCFABCEBA1FAB9" box="[1210,1266,1307,1333]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2021
</bibRefCitation>
) is likely not correct. The species was described from a locality presumably situated in
<collectingCountry id="FFBB763DFFFE9E3D4170FA98EB3DFAD5" box="[1046,1134,1343,1369]" name="Algeria" pageId="18" pageNumber="19">Algeria</collectingCountry>
.
<bibRefCitation id="E33D4B5CFFFE9E3D411CFA98EA5FFAD5" author="Perez, J." box="[1146,1292,1342,1369]" pageId="18" pageNumber="19" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez (1895)</bibRefCitation>
noted in his description that he would provide information on the
<typeStatus id="5817880FFFFE9E3D4665FAC4EC60FAF1" box="[771,819,1379,1405]" pageId="18" pageNumber="19">type</typeStatus>
locality in a later publication, which, however, never materialized.
</paragraph>
<paragraph id="871336ADFFFE9E3D45A1FA0CEB06F9BD" blockId="18.[151,1437,730,1945]" pageId="18" pageNumber="19">
<emphasis id="B5D8EABFFFFE9E3D45A1FA0CED40FA49" box="[199,531,1451,1477]" italics="true" pageId="18" pageNumber="19">Genetic barcode information</emphasis>
. The barcode sequence of
<specimenCount id="91AAFD24FFFE9E3D4635FA0CEB58FA49" box="[851,1035,1451,1477]" count="3" pageId="18" pageNumber="19" type="generic">three specimens</specimenCount>
from
<collectingCountry id="FFBB763DFFFE9E3D4131FA0CEBEFFA49" box="[1111,1212,1451,1477]" name="Morocco" pageId="18" pageNumber="19">Morocco</collectingCountry>
(tjw017-019) were obtained (
<collectionCode id="E1BDAE68FFFE9E3D446EFA68EE3FFA65" box="[264,364,1487,1513]" pageId="18" pageNumber="19">ABABZ</collectionCode>
088-23, ABABZ089-23, ABABZ090-23). They were assigned to BIN BOLD:AFT8832. In the COI Species ID Tree (
<figureCitation id="1F972A28FFFE9E3D44F6FA54EEB0F981" box="[400,483,1523,1549]" captionStart="FIGURE 28" captionStartId="43.[151,250,1958,1982]" captionTargetBox="[221,1351,178,1932]" captionTargetId="graphics-20@43.[333,900,178,1905]" captionText="FIGURE 28. Species ID tree as inferred from the DNA sequence of the mitochondrial cytochrome c oxidase subunit 1 gene (COI gene, “barcoding gene”). Bootstrap values above 70% are shown next to the nodes." figureDoi="http://doi.org/10.5281/zenodo.14275914" httpUri="https://zenodo.org/record/14275914/files/figure.png" pageId="18" pageNumber="19">Fig. 28</figureCitation>
), nearest neighbours are
<taxonomicName id="40AC4D2EFFFE9E3D479CFA54ECC6F981" baseAuthorityName="Mocsary" baseAuthorityYear="1884" box="[762,917,1523,1549]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis id="B5D8EABFFFFE9E3D479CFA54ECC6F981" box="[762,917,1523,1549]" italics="true" pageId="18" pageNumber="19">P. reticulatum</emphasis>
</taxonomicName>
and the members of the
<taxonomicName id="40AC4D2EFFFE9E3D41CFFA54EA17F981" baseAuthorityName="Klug" baseAuthorityYear="1832" box="[1193,1348,1523,1549]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melanurum">
<emphasis id="B5D8EABFFFFE9E3D41CFFA54EA17F981" box="[1193,1348,1523,1549]" italics="true" pageId="18" pageNumber="19">P. melanurum</emphasis>
</taxonomicName>
and the
<taxonomicName id="40AC4D2EFFFE9E3D45F1F9B0EE5CF9BD" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[151,271,1559,1585]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="alpinum">
<emphasis id="B5D8EABFFFFE9E3D45F1F9B0EE5CF9BD" box="[151,271,1559,1585]" italics="true" pageId="18" pageNumber="19">P. alpinum</emphasis>
</taxonomicName>
groups, but these relationships are not supported by high bootstrap values.
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFFE9E3D45A1F99CEC19F815" pageId="18" pageNumber="19" type="multiple">
<paragraph id="871336ADFFFE9E3D45A1F99CEDA9F94D" blockId="18.[151,1437,730,1945]" pageId="18" pageNumber="19">
<taxonomicName id="40AC4D2EFFFE9E3D45A1F99CEE0BF9D9" ID-CoL="8S227" authorityName="Pasteels" authorityYear="1969" box="[199,344,1595,1621]" class="Insecta" family="Megachilidae" genus="Neanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFE9E3D45A1F99CEE0BF9D9" box="[199,344,1595,1621]" italics="true" pageId="18" pageNumber="19">Neanthidium</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="E33D4B5CFFFE9E3D475EF99CEDB7F9D9" author="Pasteels, J. J." box="[568,740,1595,1621]" pageId="18" pageNumber="19" pagination="3 - 148" refId="ref31256" refString="Pasteels, J. J. (1969) La systematique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l'Ancien Monde. Memoires de la Societe Royale d'Entomologie de Belgique, 31, 3 - 148." type="journal article" year="1969">Pasteels (1969)</bibRefCitation>
as a monotypic genus to accommodate this species. Michener (1997, 2007) recognized the genus, but
<bibRefCitation id="E33D4B5CFFFE9E3D4739F9F8EC15F9F5" author="Litman, J. R. &amp; Griswold, T. &amp; Danforth, B. N." box="[607,838,1631,1657]" pageId="18" pageNumber="19" pagination="183 - 198" refId="ref30407" refString="Litman, J. R., Griswold, T. &amp; Danforth, B. N. (2016) Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution, 100, 183 - 198. https: // doi. org / 10.1016 / j. ympev. 2016.03.018" type="journal article" year="2016">
Litman
<emphasis id="B5D8EABFFFFE9E3D47DFF9F8EDA0F9F5" box="[697,755,1631,1657]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2016)
</bibRefCitation>
relocated it to the genus
<taxonomicName id="40AC4D2EFFFE9E3D410DF9F8EA7CF9F5" ID-CoL="6Z7N" authorityName="Friese" authorityYear="1898" box="[1131,1327,1631,1657]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFE9E3D410DF9F8EA7CF9F5" box="[1131,1327,1631,1657]" italics="true" pageId="18" pageNumber="19">Pseudoanthidium</emphasis>
</taxonomicName>
based on a genetic analysis.
<taxonomicName id="40AC4D2EFFFE9E3D4409F924EDE2F911" ID-CoL="8SPF4" authorityName="Kasparek" authorityYear="2024" box="[367,689,1667,1693]" class="Insecta" family="Megachilidae" genus="Royanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicoloripenne">
<emphasis id="B5D8EABFFFFE9E3D4409F924EDE2F911" box="[367,689,1667,1693]" italics="true" pageId="18" pageNumber="19">Royanthidium bicoloripenne</emphasis>
</taxonomicName>
is herein attributed here to
<taxonomicName id="40AC4D2EFFFE9E3D468BF924EBF5F911" ID-CoL="8SFXP" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[1005,1190,1667,1693]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octodentatum">
<emphasis id="B5D8EABFFFFE9E3D468BF924EBF5F911" box="[1005,1190,1667,1693]" italics="true" pageId="18" pageNumber="19">R. octodentatum</emphasis>
</taxonomicName>
as a junior synonym, based on the examination of the
<typeStatus id="5817880FFFFE9E3D4765F900ED36F94D" box="[515,613,1703,1729]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
in NHMUK.
</paragraph>
<paragraph id="871336ADFFFE9E3D45A1F96CEC19F815" blockId="18.[151,1437,730,1945]" pageId="18" pageNumber="19">
<emphasis id="B5D8EABFFFFE9E3D45A1F96CEE35F969" box="[199,358,1739,1765]" italics="true" pageId="18" pageNumber="19">Identification.</emphasis>
The male is distinguished from all other males of the genus (and from all other males of the Palaearctic Anthidiini) by having eight teeth or spines on T6T7.T6 and T7 each have a blunt spine next to the middle and an acute lateral tooth. The female is distinguished from the other Palaearctic members of
<taxonomicName id="40AC4D2EFFFE9E3D41D3F8B4EA2AF8A1" ID-CoL="6Z7N" authorityName="Friese" authorityYear="1898" box="[1205,1401,1811,1837]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFE9E3D41D3F8B4EA2AF8A1" box="[1205,1401,1811,1837]" italics="true" pageId="18" pageNumber="19">Pseudoanthidium</emphasis>
</taxonomicName>
by its truncate clypeus with small black tubercles at its apical margin, and a median tubercle larger than the others. In both sexes, the species is further characterized by its elongate mandible. There are only a few other Palaearctic members of
<taxonomicName id="40AC4D2EFFFE9E3D4445F8D8EEB4F815" ID-CoL="6Z7N" authorityName="Friese" authorityYear="1898" box="[291,487,1919,1945]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFFE9E3D4445F8D8EEB4F815" box="[291,487,1919,1945]" italics="true" pageId="18" pageNumber="19">Pseudoanthidium</emphasis>
</taxonomicName>
with mandibles almost as long.
</paragraph>
</subSubSection>
<caption id="D3D36625FFFF9E3C45F1F830EB63F85F" ID-DOI="http://doi.org/10.5281/zenodo.14240757" ID-Zenodo-Dep="14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1943,1967]" targetBox="[153,1436,181,1918]" targetPageId="19" targetType="figure">
<paragraph id="871336ADFFFF9E3C45F1F830EB63F85F" blockId="19.[151,1437,1943,2004]" pageId="19" pageNumber="20">
<emphasis id="B5D8EABFFFFF9E3C45F1F830EE4CF823" bold="true" box="[151,287,1943,1967]" pageId="19" pageNumber="20">FIGURE 10.</emphasis>
<taxonomicName id="40AC4D2EFFFF9E3C4440F830EDA6F823" authority="(Perez, 1895)" baseAuthorityName="Perez" baseAuthorityYear="1895" box="[294,757,1943,1967]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="octodentatum">
<emphasis id="B5D8EABFFFFF9E3C4440F830ED34F823" box="[294,615,1943,1967]" italics="true" pageId="19" pageNumber="20">Pseudoanthidium octodentatum</emphasis>
(
<bibRefCitation id="E33D4B5CFFFF9E3C4712F830EDBDF823" author="Perez, J." box="[628,750,1943,1967]" pageId="19" pageNumber="20" refId="ref31351" refString="Perez, J. (1895) Especes nouvelles de melliferes de Barbarie (diagnoses preliminaires). G. Gounouihou, Bordeaux, 64 pp" type="book" year="1895">Pérez, 1895</bibRefCitation>
)
</taxonomicName>
(a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male
</paragraph>
</caption>
<subSubSection id="CFB66526FFF89E3B45A1FF30EA35FC45" pageId="20" pageNumber="21" type="description">
<paragraph id="871336ADFFF89E3B45A1FF30EE8EFDED" blockId="20.[151,1437,151,1041]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1FF30EE18FF3C" box="[199,331,151,176]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
.
<emphasis id="B5D8EABFFFF89E3B4406FF30EEE7FF3D" box="[352,436,151,177]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
(
<figureCitation id="1F972A28FFF89E3B44AFFF30ED63FF3D" box="[457,560,151,177]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="20" pageNumber="21">Fig. 10a</figureCitation>
).
<quantity id="40549B48FFF89E3B472CFF30EDF2FF3D" box="[586,673,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="mm" value="10.0">10 mm</quantity>
.—
<emphasis id="B5D8EABFFFF89E3B47A4FF30EDACFF3D" box="[706,767,151,177]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
: Clypeus and lower paraocular area yellow, with upper paraocular area with a yellow narrow stripe along the orbit; gena and preoccipital area predominantly dark yellow to orange, interrupted in the middle; clypeus broader than long, finely punctate throughout; apical margin black, largely hidden under long white pubescence (
<figureCitation id="1F972A28FFF89E3B47FCFEA4EDBCFE91" box="[666,751,259,285]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[180,1408,181,1912]" captionTargetId="figure-15@35.[180,1408,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Face of the female of Palaearctic species of Pseudoanthidium (continued on the following page)." figureDoi="http://doi.org/10.5281/zenodo.14240775" httpUri="https://zenodo.org/record/14240775/files/figure.png" pageId="20" pageNumber="21">Fig. 20</figureCitation>
); margin crenulated with the median tubercle larger than the others; mandible yellow with six black teeth; teeth to1to3 equally long, to4 being the smallest, followed by to5, with to6 being the largest. Proximal antennal segments (sg1sg7) orange, outer segments brown with fine orange tinge.—
<taxonomicName id="40AC4D2EFFF89E3B4594FEC8EE39FE04" box="[242,362,367,392]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF89E3B4594FEC8EE39FE04" box="[242,362,367,392]" italics="true" pageId="20" pageNumber="21">Mesosoma</emphasis>
</taxonomicName>
: Scutum finely punctate and black with broad anterior yellow band; pronotal lobe with high lamella, outer side yellow, inner side black; scutellum and axillae broadly V-shaped with outer side yellow and anterior side black; outer margin rounded in profile.—
<emphasis id="B5D8EABFFFF89E3B4672FE10EC0BFE5C" box="[788,856,439,464]" italics="true" pageId="20" pageNumber="21">Wings</emphasis>
: Brownish infuscated.—
<emphasis id="B5D8EABFFFF89E3B411FFE10EBBDFE5C" box="[1145,1262,439,464]" italics="true" pageId="20" pageNumber="21">Metasoma</emphasis>
: Broad yellow bands on T1T6, extending closer to the midline in the distal terga; T6 with a small lateral projection pointed posteriorly; lamella with a median emargination for the sting (
<figureCitation id="1F972A28FFF89E3B463FFE58ECEDFD95" box="[857,958,511,537]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="20" pageNumber="21">Figs 10c</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B46ADFE58ECB9FD95" box="[971,1002,511,537]" captionStart="FIGURE 22" captionStartId="38.[151,250,1491,1515]" captionTargetBox="[409,1188,1275,1417]" captionTargetPageId="38" captionText="FIGURE 22. Apical terga of females of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240781" httpUri="https://zenodo.org/record/14240781/files/figure.png" pageId="20" pageNumber="21">22</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B4690FE58EB46FD95" box="[1014,1045,511,537]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="20" pageNumber="21">23</figureCitation>
); marginal zones of terga punctate with very narrow impunctate outer margin.—
<emphasis id="B5D8EABFFFF89E3B47FAFD84ED82FDB0" box="[668,721,547,572]" italics="true" pageId="20" pageNumber="21">Legs</emphasis>
: Femora yellow with some black at the bases and on the inner faces; tibiae and tarsi yellow.
</paragraph>
<paragraph id="871336ADFFF89E3B45A1FDCCEA35FC45" blockId="20.[151,1437,151,1041]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1FDCCEE52FD09" box="[199,257,619,645]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
(
<figureCitation id="1F972A28FFF89E3B4476FDCCEE21FD09" box="[272,370,619,645]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="20" pageNumber="21">Fig. 10b</figureCitation>
).
<quantity id="40549B48FFF89E3B44E0FDCCED52FD09" box="[390,513,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="20" pageNumber="21" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
.—
<emphasis id="B5D8EABFFFF89E3B4744FDCCED0CFD09" box="[546,607,619,645]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
: Clypeus, lower part of supraclypeal area and upper and lower paraocular area yellow (
<figureCitation id="1F972A28FFF89E3B4448FD28EED7FD25" box="[302,388,655,681]" captionStart="FIGURE 21" captionStartId="37.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-15@37.[179,1408,181,1919]" captionTargetPageId="37" captionText="FIGURE 21. Faces of the male of Palaearctic species of the genus Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240779" httpUri="https://zenodo.org/record/14240779/files/figure.png" pageId="20" pageNumber="21">Fig. 21</figureCitation>
); mandible long with three strong black teeth, the lowest tooth being the strongest; yellow maculation on gena and in preoccipital area mostly less extended than in the female, often not reaching the mandibular articulation, and the interruption in the preoccipital area generally wider than in the female.—
<taxonomicName id="40AC4D2EFFF89E3B41D9FD70EA64FD7C" box="[1215,1335,727,752]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B5D8EABFFFF89E3B41D9FD70EA64FD7C" box="[1215,1335,727,752]" italics="true" pageId="20" pageNumber="21">Mesosoma</emphasis>
</taxonomicName>
: Scutum black with fine punctation, occasionally some yellow anteriorly; pronotal lobe as in female; scutellum and axilla as in female, but with less extensive yellow colouration.—
<emphasis id="B5D8EABFFFF89E3B4799FCB8EC67FCB4" box="[767,820,799,824]" italics="true" pageId="20" pageNumber="21">Legs</emphasis>
:As in the female.—
<emphasis id="B5D8EABFFFF89E3B4170FCB8EBD8FCB4" box="[1046,1163,799,824]" italics="true" pageId="20" pageNumber="21">Metasoma</emphasis>
: T1T5 as in the female, T6 entirely yellow or with narrow black midline; T6 and T7 each with a blunt spine next to the middle and an acute lateral tooth (
<figureCitation id="1F972A28FFF89E3B4448FCC0EEF8FC0D" box="[302,427,871,897]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="20" pageNumber="21">Fig. 10c, d</figureCitation>
). S1 with rounded longitudinal elevation along the midline; S2 with subapical transversal carina; apical margin of S4S5 widely concavely emarginate; apical margin of S6 convexly rounded (
<figureCitation id="1F972A28FFF89E3B406CFC2CEA3EFC29" box="[1290,1389,907,933]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[153,1436,181,1918]" captionTargetId="figure-15@19.[152,1436,181,1918]" captionTargetPageId="19" captionText="FIGURE 10. Pseudoanthidium octodentatum (Pérez, 1895) (a.e. from Morocco). a. Habitus dorsal, female; b. Habitus dorsal, male; c. Apical terga of the female; d. Apical terga of the male; e. Apical sterna of the male" figureDoi="http://doi.org/10.5281/zenodo.14240757" httpUri="https://zenodo.org/record/14240757/files/figure.png" pageId="20" pageNumber="21">Figs 10e</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B401FFC2CEAC4FC29" box="[1401,1431,907,933]" captionStart="FIGURE 23" captionStartId="39.[151,250,1918,1942]" captionTargetBox="[173,1415,561,1882]" captionTargetId="figure-169@39.[173,1415,561,1895]" captionTargetPageId="39" captionText="FIGURE 23. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14240783" httpUri="https://zenodo.org/record/14240783/files/figure.png" pageId="20" pageNumber="21">23</figureCitation>
,
<figureCitation id="1F972A28FFF89E3B45F1FC08EFE6FC45" box="[151,181,943,969]" captionStart="FIGURE 24" captionStartId="40.[151,250,1170,1194]" captionTargetBox="[307,1281,311,1152]" captionTargetId="graphics-80@40.[648,939,711,796]" captionTargetPageId="40" captionText="FIGURE 24. Apical terga of males of Palaearctic Pseudoanthidium species." figureDoi="http://doi.org/10.5281/zenodo.14275912" httpUri="https://zenodo.org/record/14275912/files/figure.png" pageId="20" pageNumber="21">24</figureCitation>
).—
<emphasis id="B5D8EABFFFF89E3B4586FC08EE19FC45" box="[224,330,943,969]" italics="true" pageId="20" pageNumber="21">Genitalia</emphasis>
: Gonostyli tubular; penis valve tapering towards the apex with acutely arcuate apex (
<figureCitation id="1F972A28FFF89E3B4060FC08EA09FC45" box="[1286,1370,942,969]" captionStart="FIGURE 25" captionStartId="40.[151,250,2009,2033]" captionTargetBox="[176,1412,1240,1996]" captionTargetId="figure-103@40.[176,1412,1240,2001]" captionTargetPageId="40" captionText="FIGURE 25. Male genitalia of selected Palaearctic species of Pseudoanthidium." figureDoi="http://doi.org/10.5281/zenodo.14240785" httpUri="https://zenodo.org/record/14240785/files/figure.png" pageId="20" pageNumber="21">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFB66526FFF89E3B45A1FC74EADAFB9D" pageId="20" pageNumber="21" type="distribution">
<paragraph id="871336ADFFF89E3B45A1FC74EADAFB9D" blockId="20.[151,1437,151,1041]" pageId="20" pageNumber="21">
<emphasis id="B5D8EABFFFF89E3B45A1FC74EE03FC61" box="[199,336,979,1005]" italics="true" pageId="20" pageNumber="21">Distribution</emphasis>
.
<collectingCountry id="FFBB763DFFF89E3B4406FC74EE9BFC61" box="[352,456,979,1005]" name="Morocco" pageId="20" pageNumber="21">Morocco</collectingCountry>
,
<collectingCountry id="FFBB763DFFF89E3B44B5FC74ED79FC61" box="[467,554,979,1005]" name="Algeria" pageId="20" pageNumber="21">Algeria</collectingCountry>
, and
<collectingCountry id="FFBB763DFFF89E3B470EFC74EDFAFC61" box="[616,681,979,1005]" name="Libya" pageId="20" pageNumber="21">Libya</collectingCountry>
(
<figureCitation id="1F972A28FFF89E3B47DDFC74EC4EFC61" box="[699,797,979,1005]" captionStart="FIGURE 26" captionStartId="41.[151,250,1633,1657]" captionTargetBox="[151,1435,181,1609]" captionTargetId="figure-15@41.[151,1435,181,1609]" captionTargetPageId="41" captionText="FIGURE 26. Distribution of selected species of Pseudoanthidium. a. Pseudoanthidium microrubrum sp. nov., P. octodentatum, and P. royoi stat. resurrect. Since the collection site of P. octodentatum in Algeria is not known, a random location was selected; b. Pseudoanthidium ivanovi stat. resurrect., P. obscuratum, and P. tajikistanicum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240787" httpUri="https://zenodo.org/record/14240787/files/figure.png" pageId="20" pageNumber="21">Fig. 26a</figureCitation>
).
<bibRefCitation id="E33D4B5CFFF89E3B4655FC74EB49FC61" author="Ornosa, C. &amp; Ortiz-Sanchez, F. J. &amp; Torres, F." box="[819,1050,979,1005]" pageId="20" pageNumber="21" pagination="61 - 86" refId="ref31152" refString="Ornosa, C., Ortiz-Sanchez, F. J. &amp; Torres, F. (2008) Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Graellsia, 64, 61 - 86. https: // doi. org / 10.3989 / graellsia. 2008. v 64. i 1.55" type="journal article" year="2008">
Ornosa
<emphasis id="B5D8EABFFFF89E3B46EAFC74EC94FC61" box="[908,967,979,1005]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
(2008)
</bibRefCitation>
also mentioned
<collectingCountry id="FFBB763DFFF89E3B41BEFC74EA69FC61" box="[1240,1338,979,1005]" name="Lebanon" pageId="20" pageNumber="21">Lebanon</collectingCountry>
and this was taken over by
<bibRefCitation id="E33D4B5CFFF89E3B440EFC50EDC8FB9D" author="Ascher, J. S. &amp; Pickering, J." box="[360,667,1015,1041]" pageId="20" pageNumber="21" refId="ref29495" refString="Ascher, J. S. &amp; Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: http: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species (accessed 1 August 20024)" type="url" year="2024">Ascher &amp; Pickering (2024)</bibRefCitation>
. However, confirmation is required (see also
<bibRefCitation id="E33D4B5CFFF89E3B41F1FC50EA2FFB9D" author="Boustani, M. &amp; Dathe, H. &amp; Ghisbain, G. &amp; Kasparek, M. &amp; Michez, D. &amp; Nemer, N. &amp; Pauly, A. &amp; Risch, S. &amp; Terzo, M. &amp; Van Achter, X. &amp; Wood, T. &amp; Rasmont, P." box="[1175,1404,1015,1041]" pageId="20" pageNumber="21" pagination="1 - 146" refId="ref29624" refString="Boustani, M., Dathe, H., Ghisbain, G., Kasparek, M., Michez, D., Nemer, N., Pauly, A., Risch, S., Terzo, M., Van Achter, X., Wood, T. &amp; Rasmont, P. (2021) The bees of Lebanon (Hymenoptera: Apoidea: Anthophila). Zootaxa, 4976 (1), 1 - 146. https: // doi. org / 10.11646 / zootaxa. 4976.1.1" type="journal article" year="2021">
Boustani
<emphasis id="B5D8EABFFFF89E3B4067FC50EA69FB9D" box="[1281,1338,1015,1041]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,51 +1,59 @@
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@ -178,7 +186,7 @@ Craib
1931: 279 (
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).
<materialsCitation id="0D913C93FFEFE4751AB7FE3CECA8C5FD" ID-GBIF-Occurrence="3050815338" collectingDate="1930-02-21" collectionCode="K, BM" collectorName="A. F. G. Kerr" country="Thailand" location="Yanyao" municipality="Surat" pageId="22" pageNumber="23" specimenCode="K000761880, K001129592, BM000945616" specimenCount="2" typeStatus="holotype">
<materialsCitation id="0D913C93FFEFE4751AB7FE3CEB07C581" ID-GBIF-Occurrence="3050815338" box="[234,1068,404,430]" collectingDate="1930-02-21" collectionCode="K, BM" collectorName="A. F. G. Kerr" country="Thailand" location="Yanyao" municipality="Surat" pageId="22" pageNumber="23" specimenCode="K000761880, K001129592, BM000945616" specimenCount="2" typeStatus="holotype">
<emphasis id="8F8DEADCFFEFE4751AB7FE3CEE0CC581" bold="true" box="[234,295,404,430]" pageId="22" pageNumber="23">Type</emphasis>
:
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@ -195,7 +203,9 @@ Craib
<collectorName id="100C5318FFEFE4751915FE3CECF9C581" box="[840,978,404,430]" pageId="22" pageNumber="23">A.F.G. Kerr</collectorName>
18175
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;
;
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:
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@ -203,7 +213,9 @@ Craib
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]! [
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]! (2 sheets);
]! (2 sheets);
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<materialsCitation id="0D913C93FFEFE4751875FE10ECA8C5FD" ID-GBIF-Occurrence="3050815338" box="[552,899,439,466]" collectingDate="1930-02-21" collectionCode="K, BM" collectorName="A. F. G. Kerr" country="Thailand" location="Yanyao" municipality="Surat" pageId="22" pageNumber="23" specimenCode="K000761880, K001129592, BM000945616" specimenCount="2" typeStatus="holotype">
<typeStatus id="6242886CFFEFE4751875FE10EDAAC5FD" box="[552,641,440,466]" pageId="22" pageNumber="23" type="isotype">isotype</typeStatus>
:
<collectionCode id="DBE8AE0BFFEFE47518D1FE1FED90C5FD" box="[652,699,439,466]" country="United Kingdom" name="Bristol Museum" pageId="22" pageNumber="23">BM</collectionCode>

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@ -1,110 +1,119 @@
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3869688" ID-GBIF-Taxon="164400359" ID-Zenodo-Dep="3869688" LSID="urn:lsid:plazi:treatment:F442953B2E41EF5EFF1BFEA5FDC97FCF" httpUri="http://treatment.plazi.org/id/F442953B2E41EF5EFF1BFEA5FDC97FCF" lastPageNumber="61" pageId="60" pageNumber="61">
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<emphasis box="[189,204,333,359]" italics="true" pageId="60" pageNumber="61">*</emphasis>
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<treatment id="F442953B2E41EF5EFF1BFEA5FDC97FCF" ID-DOI="http://doi.org/10.5281/zenodo.3869688" ID-GBIF-Taxon="164400359" ID-Zenodo-Dep="3869688" LSID="urn:lsid:plazi:treatment:F442953B2E41EF5EFF1BFEA5FDC97FCF" httpUri="http://treatment.plazi.org/id/F442953B2E41EF5EFF1BFEA5FDC97FCF" lastPageNumber="61" pageId="60" pageNumber="61" scope_class="Actinopterygii" scope_family="Scophthalmidae" scope_order="Pleuronectiformes" scope_phylum="Chordata" scope_superClass="Gnathostomata">
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<emphasis box="[242,503,333,359]" italics="true" pageId="60" pageNumber="61">Lepidorhombus boscii</emphasis>
<taxonomicName id="BBEB5FAE2E41EF5EFF54FEA5FD297F4E" ID-CoL="6PBSP" ID-ENA="179372" authority="(Risso, 1810)" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[242,668,333,359]" class="Actinopterygii" family="Scophthalmidae" genus="Lepidorhombus" kingdom="Animalia" order="Pleuronectiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="boscii">
<emphasis id="4E9FF83F2E41EF5EFF54FEA5FE427F4E" box="[242,503,333,359]" italics="true" pageId="60" pageNumber="61">Lepidorhombus boscii</emphasis>
(Risso, 1810)
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Fourspotted megrim; Areeiro-de-quatro-manchas
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,
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<emphasis id="4E9FF83F2E41EF5EFA85FEA1FAE27F4D" bold="true" box="[1315,1367,329,356]" pageId="60" pageNumber="61">②③</emphasis>
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The species
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<emphasis box="[375,468,366,390]" italics="true" pageId="60" pageNumber="61">L. boscii</emphasis>
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<emphasis id="4E9FF83F2E41EF5EFED1FE87FE617FAF" box="[375,468,366,390]" italics="true" pageId="60" pageNumber="61">L. boscii</emphasis>
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is recorded off
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(Josephine Bank) and to the northeast of the
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(157 NM southwest of the Georgiy Zima Seamount), both outside their respective EEZs.
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is recorded off Madeira (Josephine Bank) and to the northeast of the
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(157 NM southwest of the
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), both outside their respective EEZs.
<collectionCode id="1AFABCE82E41EF5EFBCFFE67FB087F8F" box="[1129,1213,399,422]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="60" pageNumber="61">MNHN</collectionCode>
, No. 1956-0007,
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<geoCoordinate id="19DF42EA2E41EF5EFF54FE46FEFF7FEF" box="[242,330,430,454]" degrees="20.0" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="5555" value="-20.0">20.0º W</geoCoordinate>
,
<geoCoordinate box="[342,427,430,454]" degrees="40.0" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="5555" value="40.0">40.0º N</geoCoordinate>
;
<collectingRegion box="[440,524,431,454]" country="Ireland" name="Monaghan" pageId="60" pageNumber="61">MNHN</collectingRegion>
<geoCoordinate id="19DF42EA2E41EF5EFEF0FE46FE1E7FEF" box="[342,427,430,454]" degrees="40.0" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="5555" value="40.0">40.0º N</geoCoordinate>
;
</materialsCitation>
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<collectionCode id="1AFABCE82E41EF5EFE1EFE47FDB97FEF" box="[440,524,431,454]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="60" pageNumber="61">MNHN</collectionCode>
, No. 1956-0008,
<geoCoordinate box="[714,802,430,454]" degrees="20.0" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="5555" value="-20.0">20.0º W</geoCoordinate>
<geoCoordinate id="19DF42EA2E41EF5EFD6CFE46FC977FEF" box="[714,802,430,454]" degrees="20.0" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="5555" value="-20.0">20.0º W</geoCoordinate>
,
<geoCoordinate box="[814,899,430,454]" degrees="40.0" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="5555" value="40.0">40.0º N</geoCoordinate>
, both specimens captured in
<date box="[1215,1328,430,454]" pageId="60" pageNumber="61" value="1955-06">June 1955</date>
; FISH 1556599 (
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<geoCoordinate id="19DF42EA2E41EF5EFC88FE46FC367FEF" box="[814,899,430,454]" degrees="40.0" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="5555" value="40.0">40.0º N</geoCoordinate>
,
<specimenCount id="6AEDEFA42E41EF5EFC29FE46FB837FEF" box="[911,1078,430,454]" pageId="60" pageNumber="61">both specimens</specimenCount>
captured in
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<collectingDate id="1811FB052E41EF5EFB19FE47FA857FEF" box="[1215,1328,430,454]" pageId="60" pageNumber="61" value="1955-06">June 1955</collectingDate>
</date>
;
<specimenCode id="2C4D8C562E41EF5EFA9BFE46FEFA7FCF" pageId="60" pageNumber="61">FISH 1556599</specimenCode>
(
<bibRefCitation id="187A59DC2E41EF5EFEFBFE26FDC57FCF" author="Shcherbachev" box="[349,624,462,486]" etAl="et al." firstAuthor="Shcherbachev" pageId="60" pageNumber="61" pagination="110 - 125" refId="ref43908" refString="Shcherbachev Y. N., Kukuev E. I. &amp; Shlibanov V. I. 1985. Composition of the benthic and demersal ichthyocenoses of the submarine mountains in the southern part of the North Atlantic Range. Jounal of Ichthyology 25: 110 - 125." type="journal article" year="1985">
Shcherbachev
<emphasis box="[507,561,462,486]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
<emphasis id="4E9FF83F2E41EF5EFE5DFE27FD847FCF" box="[507,561,462,486]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
1985
</bibRefCitation>
).

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@ -1,107 +1,112 @@
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*
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Roule, 1921
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Longfin escolar;
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New record for Portuguese mainland waters; the first specimen of
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New record for Portuguese mainland waters; the first
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) was caught on the 15
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of
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(
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TL and
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), using a bottom trawl (
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,
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) and other specimens were collected in commercial fisheries in
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. Bold Systems Sample ID MLFPI62, available on the Barcode of Life Data Systems (BOLD; www.barcodinglife.org, under the project titled “Fish of
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and
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[MLFPI]”).
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. Bold Systems Sample ID
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, available on the Barcode of Life Data Systems (BOLD; www.barcodinglife.org, under the project titled “Fish of Portugal and Italy [
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]”).
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<mods:title id="C67B32F10C7E4D3535010C99B89C3645">Updated checklist of marine fishes (Chordata: Craniata) from Portugal and the proposed extension of the Portuguese continental shelf</mods:title>
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<emphasis id="4E9FF83F2E48EF57FF54F95EFE0978F9" box="[242,444,1718,1744]" italics="true" pageId="53" pageNumber="54">Lycenchelys alba</emphasis>
(
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)
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@ -75,25 +75,20 @@
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<emphasis id="4E9FF83F2E48EF57FF54F931FE8878D9" box="[242,317,1752,1776]" italics="true" pageId="53" pageNumber="54">L. alba</emphasis>
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is reported off the
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, in the
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(
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, in the PECS area (56 NM southeast of the
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).
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collected in
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as
collected in 1883 as
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<emphasis id="4E9FF83F2E48EF57FE6DF910FDD47939" box="[459,609,1784,1808]" italics="true" pageId="53" pageNumber="54">Lycodes albus</emphasis>
<bibRefCitation id="187A59DC2E48EF57FDC1F910FD4A7939" author="Vaillant L. L. &amp; Milne-Edwards A." box="[615,767,1784,1808]" pageId="53" pageNumber="54" refId="ref44481" refString="Vaillant L. L. &amp; Milne-Edwards A. 1888. Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883: Poissons. G. Masson, Paris." type="journal volume" year="1888">Vaillant, 1888</bibRefCitation>
<bibRefCitation id="187A59DC2E48EF57FDC1F910FD4A7939" author="Vaillant" box="[615,767,1784,1808]" firstAuthor="Vaillant" pageId="53" pageNumber="54" refId="ref44481" refString="Vaillant L. L. &amp; Milne-Edwards A. 1888. Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883: Poissons. G. Masson, Paris." type="journal volume" year="1888">Vaillant, 1888</bibRefCitation>
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,
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@ -113,7 +108,6 @@ is a synonym of
.
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