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<mods:title id="CFE0037157D24C614AD760F3E4C79D9D">Morphology of male and female terminalia and taxonomic revision of the burrower bugs genus Canthophorus (Heteroptera: Cydnidae)</mods:title>
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<treatment id="385C87B6FFD00E7EFEB3FDD8FB78FA21" LSID="urn:lsid:plazi:treatment:385C87B6FFD00E7EFEB3FDD8FB78FA21" httpUri="http://treatment.plazi.org/id/385C87B6FFD00E7EFEB3FDD8FB78FA21" lastPageNumber="35" pageId="34" pageNumber="35">
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<taxonomicName id="77F54D23FFD00E7EFEB3FDD8FD25FD94" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[262,676,613,636]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD00E7EFEB3FDD8FE74FD94" bold="true" box="[262,501,613,636]" italics="true" pageId="34" pageNumber="35">Canthophorus mixtus</emphasis>
<bibRefCitation id="D4644B51FFD00E7EFE4BFDD8FD25FD94" author="Asanova RB" box="[510,676,613,636]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="34" pageNumber="35" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
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(
<figureCitation id="28CE2A25FFD00E7EFF7CFD39FEE6FD73" box="[201,359,644,668]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="34" pageNumber="35">
Figures
<date id="C44B1060FFD00E7EFE96FD39FEBBFD73" box="[291,314,644,667]" pageId="34" pageNumber="35">
3
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,
<figureCitation id="28CE2A25FFD00E7EFEC3FD39FE14FD73" box="[374,405,644,667]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="34" pageNumber="35">
5
<collectionCode id="D6E4AE65FFD00E7EFE36FD39FE14FD73" box="[387,405,644,667]" pageId="34" pageNumber="35">C</collectionCode>
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,
<figureCitation id="28CE2A25FFD00E7EFE11FD39FE6EFD73" box="[420,495,644,667]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="34" pageNumber="35">
6
<collectionCode id="D6E4AE65FFD00E7EFE05FD39FE41FD73" box="[432,448,644,667]" pageId="34" pageNumber="35">T</collectionCode>
, 6
<collectionCode id="D6E4AE65FFD00E7EFE69FD39FE6EFD73" box="[476,495,644,667]" pageId="34" pageNumber="35">U</collectionCode>
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,
<figureCitation id="28CE2A25FFD00E7EFE48FD39FDC7FD73" box="[509,582,644,668]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="34" pageNumber="35">
7
<collectionCode id="D6E4AE65FFD00E7EFDBFFD39FD9FFD73" box="[522,542,644,667]" pageId="34" pageNumber="35">K</collectionCode>
–7
<collectionCode id="D6E4AE65FFD00E7EFD8DFD39FDC7FD73" box="[568,582,644,667]" pageId="34" pageNumber="35">S</collectionCode>
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,
<figureCitation id="28CE2A25FFD00E7EFDE0FD39FD23FD73" box="[597,674,644,668]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="34" pageNumber="35">
8
<collectionCode id="D6E4AE65FFD00E7EFDD4FD39FDF4FD73" box="[609,629,644,667]" pageId="34" pageNumber="35">O</collectionCode>
–8
<collectionCode id="D6E4AE65FFD00E7EFD3AFD39FD23FD73" box="[655,674,644,667]" pageId="34" pageNumber="35">U</collectionCode>
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,
<figureCitation id="28CE2A25FFD00E7EFD04FD39FE4BFD53" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="34" pageNumber="35">
11
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<figureCitation id="28CE2A25FFD00E7EFE6CFD19FD83FD53" box="[473,514,676,699]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="34" pageNumber="35">
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</emphasis>
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<taxonomicName id="77F54D23FFD00E7EFF2BFD73FDD4FD0B" authority="Asanova 1964: 140" authorityName="Asanova" authorityPageNumber="140" authorityYear="1964" box="[158,597,718,739]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD00E7EFF2BFD73FEF4FD0B" box="[158,373,718,739]" italics="true" pageId="34" pageNumber="35">Canthophorus mixtus</emphasis>
<bibRefCitation id="D4644B51FFD00E7EFE33FD73FDD4FD0B" author="Asanova RB" box="[390,597,718,739]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="34" pageNumber="35" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova 1964: 140</bibRefCitation>
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<collectingCountry id="C8E27630FFD00E7EFF76FD55FEB5FD15" box="[195,308,744,765]" name="Kazakhstan" pageId="34" pageNumber="35" value="Kazakhstan">Kazakhstan</collectingCountry>
,
<collectingRegion id="7231F842FFD00E7EFEFCFD55FE73FD15" box="[329,498,744,765]" pageId="34" pageNumber="35">Karaganda Prov</collectingRegion>
.,
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S of
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<collectingMunicipality id="502EACDAFFD00E7EFD11FD55FC8DFD15" box="[676,780,744,765]" pageId="34" pageNumber="35">Zhana-Ark</collectingMunicipality>
[Atasu],
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; lost]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD00E7EFF2BFCE3FBEBFBC0" pageId="34" pageNumber="35" type="description">
<paragraph id="B04A36A0FFD00E7EFF2BFCE3FEA1FC9D" blockId="34.[158,781,193,1939]" box="[158,288,862,885]" pageId="34" pageNumber="35">
<heading id="EB0281CCFFD00E7EFF2BFCE3FEA1FC9D" bold="true" box="[158,288,862,885]" fontSize="10" level="2" pageId="34" pageNumber="35" reason="6">
<emphasis id="8281EAB2FFD00E7EFF2BFCE3FEA1FC9D" bold="true" box="[158,288,862,885]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD00E7EFF2BFC38FF6EFBF4" blockId="34.[158,781,193,1939]" pageId="34" pageNumber="35">Lateral margin of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metapleural evaporatorium with long and curved peritreme. Membrane brownish, opaque.</paragraph>
<paragraph id="B04A36A0FFD00E7EFF15FB88FD39FB64" blockId="34.[158,781,193,1939]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFF15FB88FE9BFBA4" bold="true" box="[160,282,1077,1100]" pageId="34" pageNumber="35">Pygophore</emphasis>
slightly longer than wide, almost as in
<taxonomicName id="77F54D23FFD00E7EFD46FB88FEAFFB84" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFD00E7EFD46FB88FEAFFB84" italics="true" pageId="34" pageNumber="35">C. melanopterus</emphasis>
</taxonomicName>
, but with less deeply concave lateral walls and slightly shorter, not concave dorsal infolding.
</paragraph>
<paragraph id="B04A36A0FFD00E7EFF2BFB14FD71FAC8" blockId="34.[158,781,193,1939]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFF2BFB14FE51FB28" bold="true" box="[158,464,1193,1217]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFF2BFB14FE88FB28" bold="true" box="[158,265,1193,1216]" italics="true" pageId="34" pageNumber="35">Paramere</emphasis>
(
<figureCitation id="28CE2A25FFD00E7EFEAEFB14FE41FB28" box="[283,448,1193,1217]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="34" pageNumber="35">
Figures
<date id="C44B1060FFD00E7EFECCFB14FE11FB28" box="[377,400,1193,1216]" pageId="34" pageNumber="35">
3
<collectionCode id="D6E4AE65FFD00E7EFE33FB14FE11FB28" box="[390,400,1193,1216]" pageId="34" pageNumber="35">I</collectionCode>
</date>
–3
<collectionCode id="D6E4AE65FFD00E7EFE1EFB14FE41FB28" box="[427,448,1193,1216]" pageId="34" pageNumber="35">K</collectionCode>
</figureCitation>
).
</emphasis>
Sensory process rather long, broadly or rather sharply rounded at apex. In specimen from Gyumri, hypophysis strongly curved inwards and sensory process more strongly bent dorsad (
<figureCitation id="28CE2A25FFD00E7EFDC9FAB4FD61FAC8" box="[636,736,1289,1312]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="34" pageNumber="35">
Figure 3
<collectionCode id="D6E4AE65FFD00E7EFD60FAB4FD61FAC8" box="[725,736,1289,1312]" pageId="34" pageNumber="35">J</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="B04A36A0FFD00E7EFF2BFA80FAEDFF11" blockId="34.[158,781,193,1939]" lastBlockId="34.[843,1466,194,1938]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFF2BFA80FE1CFABC" bold="true" box="[158,413,1341,1364]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFF2BFA80FE89FABC" bold="true" box="[158,264,1341,1364]" italics="true" pageId="34" pageNumber="35">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFD00E7EFEAFFA80FE0FFABC" box="[282,398,1341,1364]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="34" pageNumber="35">
Figure 5
<collectionCode id="D6E4AE65FFD00E7EFECEFA80FE0FFABC" box="[379,398,1341,1364]" pageId="34" pageNumber="35">C</collectionCode>
</figureCitation>
).
</emphasis>
Ventral lobe of conjunctiva rather long, slightly protruding beyond apical margin of theca; in ventral view, apically not widened, almost rectangular, at sides of apex with two denticles directed distad and with triangular depression between them (
<figureCitation id="28CE2A25FFD00E7EFD9FFA00FD12FA3C" box="[554,659,1469,1492]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="34" pageNumber="35">
Figure 6
<collectionCode id="D6E4AE65FFD00E7EFD36FA00FD12FA3C" box="[643,659,1469,1492]" pageId="34" pageNumber="35">T</collectionCode>
</figureCitation>
); in lateral view, lobe rather narrow (
<figureCitation id="28CE2A25FFD00E7EFE02FA60FDA2FA1C" box="[439,547,1501,1524]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="34" pageNumber="35">
Figure 6
<collectionCode id="D6E4AE65FFD00E7EFDA5FA60FDA2FA1C" box="[528,547,1501,1524]" pageId="34" pageNumber="35">U</collectionCode>
</figureCitation>
). Apical part of conjunctiva slightly shorter than in
<taxonomicName id="77F54D23FFD00E7EFE5BFA40FDD2F9FC" box="[494,595,1533,1556]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFD00E7EFE5BFA40FDD2F9FC" box="[494,595,1533,1556]" italics="true" pageId="34" pageNumber="35">C. dubius</emphasis>
</taxonomicName>
and
<taxonomicName id="77F54D23FFD00E7EFD3EFA40FF51F9DC" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFD00E7EFD3EFA40FF51F9DC" italics="true" pageId="34" pageNumber="35">C. melanopterus</emphasis>
</taxonomicName>
. Apical lobes rather long, directed dorsolaterad and curved posteriad. Vesica long, without lateral branch. Dorsolateral lobes rather short, straight, directed dorsolaterad and slightly posteriad, with spaced bases and rectangularly rounded apices in lateral view. Anterior walls of dorsolateral lobes subapically with weakly sclerotised areas. Spicule extending from posterior end of basal plate, longer then in all previous species and directed anteromediad by apices. Each spicule at base sharply curved in a C-shape, in distal part smoothly or rather sharply arcuately curved, gradually tapering to pointed, more or less straight or sometimes distinctly curved outwards apex (
<figureCitation id="28CE2A25FFD00E7EFD3BF8C6FCE6FF31" box="[654,871,194,1939]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="34" pageNumber="35">
Figure 7
<collectionCode id="D6E4AE65FFD00E7EFD5EF8C6FD7FF87A" box="[747,766,1915,1938]" pageId="34" pageNumber="35">K</collectionCode>
– 7
<collectionCode id="D6E4AE65FFD00E7EFCECFF7FFCE6FF31" box="[857,871,194,217]" pageId="34" pageNumber="35">S</collectionCode>
</figureCitation>
). Single denticle on anterior end of spicule basal plate short, triangular, pointed and directed anterolaterad.
</paragraph>
<paragraph id="B04A36A0FFD00E7EFCF9FEA9FBEBFBC0" blockId="34.[843,1466,194,1938]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFCF9FEA9FBE1FEC3" bold="true" box="[844,1120,276,299]" pageId="34" pageNumber="35">
<emphasis id="8281EAB2FFD00E7EFCF9FEA9FC42FEC3" bold="true" box="[844,963,276,299]" italics="true" pageId="34" pageNumber="35">Gynatrium</emphasis>
(
<figureCitation id="28CE2A25FFD00E7EFC67FEA9FBD1FEC3" box="[978,1104,276,299]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="34" pageNumber="35">
Figure 11
<collectionCode id="D6E4AE65FFD00E7EFB88FEA9FBD1FEC3" box="[1085,1104,276,299]" pageId="34" pageNumber="35">A</collectionCode>
</figureCitation>
).
</emphasis>
Posterior pouches of gynatrial sac lying anterior to its posterior margin, rarely right on margin, very long and narrow, 4–6 times as long as wide, of the same width almost all over, at extreme apices rounded and slightly widened (
<figureCitation id="28CE2A25FFD00E7EFC01FE29FBA9FE43" box="[948,1064,404,427]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="34" pageNumber="35">
Figure 12
<collectionCode id="D6E4AE65FFD00E7EFBACFE29FBA9FE43" box="[1049,1064,404,427]" pageId="34" pageNumber="35">F</collectionCode>
</figureCitation>
). Basally pouches directed laterad, in middle sharply curved posteriad. Very rarely (in specimens from Azerbaijan, Kars and Altay Territory, Kamen’-na-Obi) pouches entirely straight (
<figureCitation id="28CE2A25FFD00E7EFBF9FE4EFB3CFDE2" box="[1100,1213,499,522]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="34" pageNumber="35">
Figure 11
<collectionCode id="D6E4AE65FFD00E7EFB1EFE4EFB3CFDE2" box="[1195,1213,499,522]" pageId="34" pageNumber="35">B</collectionCode>
</figureCitation>
); in one of the specimens examined (Bashkortostan, Aksenovo), left pouch curved while right one straight. Sometimes inner surfaces of pouches at apex or at place of bend irregularly sclerotised [probably, these areas of sclerotisation are the result of injuring the walls of the posterior pouches by the aedeagal spicules during copulation]. Anterior part of gynatrial sac usually rather large, only slightly smaller than in
<taxonomicName id="77F54D23FFD00E7EFB1CFD6FFA8AFD01" box="[1193,1291,722,745]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFD00E7EFB1CFD6FFA8AFD01" box="[1193,1291,722,745]" italics="true" pageId="34" pageNumber="35">C. dubius</emphasis>
</taxonomicName>
and
<taxonomicName id="77F54D23FFD00E7EFA89FD6FFCFFFCE1" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFD00E7EFA89FD6FFCFFFCE1" italics="true" pageId="34" pageNumber="35">C. melanopterus</emphasis>
</taxonomicName>
. Proximal part of spermathecal duct slightly longer than in these species, but shorter than half of median part as well as distal part of duct; less often median part shortened and proximal part as long as half of its lengths. Proximal tubular part of spermathecal capsule 2–3 times as long as transverse diameter of distal spherical part. First rami usually rather strongly widened anteriorly and more or less rectangularly rounded (
<figureCitation id="28CE2A25FFD00E7EFC18FC6FFBC0FC01" box="[941,1089,977,1001]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="34" pageNumber="35">
Figure 8
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–8
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) at anterior ends, less often evenly or acutely rounded (
<figureCitation id="28CE2A25FFD00E7EFC48FC4CFB0AFBE1" box="[1021,1163,1009,1033]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="34" pageNumber="35">
Figure 8
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, 8
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) or triangular, with elongated inner angle (
<figureCitation id="28CE2A25FFD00E7EFC79FBACFBDDFBC0" box="[972,1116,1041,1064]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="34" pageNumber="35">
Figure 8
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).
</paragraph>
</subSubSection>
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<emphasis id="8281EAB2FFD00E7EFCF9FBD6FC32FB6A" bold="true" box="[844,947,1131,1154]" pageId="34" pageNumber="35">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD00E7EFCF9FB2EFB78FA21" blockId="34.[843,1466,194,1938]" pageId="34" pageNumber="35">
The
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and
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of
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<emphasis id="8281EAB2FFD00E7EFB26FB2EFB7BFB42" box="[1171,1274,1171,1194]" italics="true" pageId="34" pageNumber="35">C. mixtus</emphasis>
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are absent in the
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collection; probably, Asanova left them in the Institute of Zoology in Alma-Ata, and by now they are lost (at least
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have not received any response from Alma-Ata to my inquiries about the
<typeStatus id="6F4E8802FFD00E7EFB34FAAEFB3DFAC2" box="[1153,1212,1299,1322]" pageId="34" pageNumber="35">types</typeStatus>
). Data on the
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series in the paper of
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are inaccurate; viewing the
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collection showed that she usually adduced only one of similar labels, and made many mistakes in dates and numbers of specimens. A significant part of the
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is currently missing in the
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collection.
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</treatment>
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920
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<emphasis id="8281EAB2FFD20E7CFCCEFE84FB5AFEB8" bold="true" box="[891,1243,313,336]" italics="true" pageId="32" pageNumber="33">Canthophorus wagneri wagneri</emphasis>
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(
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Figures 3
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5
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<figureCitation id="28CE2A25FFD20E7CFBAFFEE4FBE6FE98" box="[1050,1127,345,368]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="32" pageNumber="33">
6
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<figureCitation id="28CE2A25FFD20E7CFBC3FEE4FB0CFE98" box="[1142,1165,345,368]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="32" pageNumber="33">
<date id="C44B1060FFD20E7CFBC3FEE4FB0CFE98" box="[1142,1165,345,368]" pageId="32" pageNumber="33">
7
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<figureCitation id="28CE2A25FFD20E7CFB28FEE4FB72FE98" box="[1181,1267,345,368]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="32" pageNumber="33">
8
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10
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<figureCitation id="28CE2A25FFD20E7CFACEFEE4FA29FE98" box="[1403,1448,345,368]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="32" pageNumber="33">
12
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913. Vorob’ ev.”, J
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“Temir-khanshura” [
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(Ter-Minassjan), 1J(
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Additional material.
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[?][date unknown] (
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<specimenCount id="A6F3FD29FFD20E7CFAF2FD51FAE4FCE9" box="[1351,1381,747,769]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFACAFD51FA31FCE9" box="[1407,1456,748,769]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFCF9FCBBFA38FCF3" box="[844,1465,774,796]" collectionCode="ZISP" country="Russia" location="Krasnodar" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Krasnodar">
<collectingRegion id="7231F842FFD20E7CFCF9FCBBFC30FCF3" box="[844,945,774,795]" pageId="32" pageNumber="33">Krasnodar</collectingRegion>
, V.191[?] [collector unknown],
<specimenCount id="A6F3FD29FFD20E7CFA89FCBBFADBFCF4" box="[1340,1370,774,796]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFACAFCBBFA31FCF3" box="[1407,1456,774,795]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFCF9FC9CFC26FC83" collectingDate="1910-07-01" collectingDateMax="1956-07-10" collectingDateMin="1910-07-01" collectionCode="ZISP, ZMUM" collectorName="K. Arnoldi &amp; S. Nikulin" country="Russia" location="Gelendzhik" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="3" stateProvince="Krasnodar">
<location id="B52A607BFFD20E7CFCF9FC9CFC3DFCDE" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCF9FC9CFC3DFCDE" box="[844,956,801,822]" country="Russia" name="Gelendzhik" pageId="32" pageNumber="33" stateProvince="Krasnodar">Gelendzhik</location>
,
<date id="C44B1060FFD20E7CFC7BFC9CFBB6FCDE" box="[974,1079,801,822]" pageId="32" pageNumber="33" value="1910-07-01">
<collectingDate id="D40FE988FFD20E7CFC7BFC9CFBB6FCDE" box="[974,1079,801,822]" pageId="32" pageNumber="33" value="1910-07-01">1.VII.1910</collectingDate>
</date>
(Vorob’ ev),
<specimenCount id="A6F3FD29FFD20E7CFB76FC9CFB63FCDF" box="[1219,1250,801,823]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB4CFC9CFAABFCDE" box="[1273,1322,801,822]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
),
<date id="C44B1060FFD20E7CFAFDFC9CFA3EFCDE" box="[1352,1471,801,822]" pageId="32" pageNumber="33" value="1956-07-10">
<collectingDate id="D40FE988FFD20E7CFAFDFC9CFA3EFCDE" box="[1352,1471,801,822]" pageId="32" pageNumber="33" value="1956-07-10">10.VII.1956</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFCE1FC81FC49FCB9" box="[852,968,828,849]" pageId="32" pageNumber="33">K. Arnoldi</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFC5CFC81FB89FCB9" box="[1001,1032,827,849]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB95FC81FBD4FCB9" box="[1056,1109,828,849]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
),
<date id="C44B1060FFD20E7CFBC4FC81FB77FCB9" box="[1137,1270,827,849]" pageId="32" pageNumber="33" value="1934-08" valueMax="1934-09-31" valueMin="1934-08-00">
<collectingDate id="D40FE988FFD20E7CFBC4FC81FB77FCB9" box="[1137,1270,827,849]" pageId="32" pageNumber="33" value="1934-08" valueMax="1934-09-31" valueMin="1934-08-00">VIII–IX.1934</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFAA5FC81FA01FCB9" box="[1296,1408,828,849]" pageId="32" pageNumber="33">S. Nikulin</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFA14FC81FA3EFCB9" box="[1441,1471,827,849]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFCE1FCEBFC1EFC83" box="[852,927,854,875]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFC77FCEBFA38FC83" box="[962,1465,854,876]" collectingDate="1911-09-18" collectionCode="ZISP" country="Russia" location="Armavir" municipality="Armavir" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Krasnodar">
<collectingMunicipality id="502EACDAFFD20E7CFC77FCEBFB92FC83" box="[962,1043,854,875]" pageId="32" pageNumber="33">Armavir</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFB9AFCEBFB1EFC83" box="[1071,1183,854,875]" pageId="32" pageNumber="33" value="1911-09-18">
<collectingDate id="D40FE988FFD20E7CFB9AFCEBFB1EFC83" box="[1071,1183,854,875]" pageId="32" pageNumber="33" value="1911-09-18">18.IX.1911</collectingDate>
</date>
(Volnukhin),
<specimenCount id="A6F3FD29FFD20E7CFAF1FCEBFAE3FC84" box="[1348,1378,854,876]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFACAFCEBFA31FC83" box="[1407,1456,854,875]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFCF9FCCCFBC4FC48" collectingDate="1913-06-11" collectingDateMax="1924-06-31" collectingDateMin="1913-06-11" collectionCode="ZISP" country="Russia" location="Anapa" municipality="Anapa" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Krasnodar">
<collectingMunicipality id="502EACDAFFD20E7CFCF9FCCCFC0DFC6E" box="[844,908,881,902]" pageId="32" pageNumber="33">Anapa</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFC05FCCCFB9CFC6E" box="[944,1053,881,902]" pageId="32" pageNumber="33" value="1913-06-11">
<collectingDate id="D40FE988FFD20E7CFC05FCCCFB9CFC6E" box="[944,1053,881,902]" pageId="32" pageNumber="33" value="1913-06-11">11.VI.1913</collectingDate>
</date>
(Vorob’ ev),
<specimenCount id="A6F3FD29FFD20E7CFB7EFCCCFB6BFC6E" box="[1227,1258,880,902]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
,
<date id="C44B1060FFD20E7CFAB8FCCCFAFAFC6E" box="[1293,1403,880,902]" pageId="32" pageNumber="33" value="1924-05" valueMax="1924-06-31" valueMin="1924-05-00">
<collectingDate id="D40FE988FFD20E7CFAB8FCCCFAFAFC6E" box="[1293,1403,880,902]" pageId="32" pageNumber="33" value="1924-05" valueMax="1924-06-31" valueMin="1924-05-00">V–VI.1924</collectingDate>
</date>
(M. Bel’ govskiy),
<specimenCount id="A6F3FD29FFD20E7CFC6FFC36FC7BFC49" box="[986,1018,907,929]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBB3FC36FBBFFC48" box="[1030,1086,907,928]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBFAFC36FB19FC53" collectingDate="1916-05-02" collectionCode="ZISP" country="Russia" location="Krasnaya Polyana" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Kiritshenko">
<location id="B52A607BFFD20E7CFBFAFC36FA84FC48" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFBFAFC36FA84FC48" box="[1103,1285,907,928]" country="Russia" name="Krasnaya Polyana" pageId="32" pageNumber="33" stateProvince="Kiritshenko">Krasnaya Polyana</location>
, “07.. ... .. V.” [
<date id="C44B1060FFD20E7CFA1DFC36FC12FC53" pageId="32" pageNumber="33" value="1916-05-02">2. V.1916</date>
] (Kiritshenko),
<specimenCount id="A6F3FD29FFD20E7CFB9AFC1BFBCCFC53" box="[1071,1101,933,955]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBEEFC1BFB10FC53" box="[1115,1169,934,955]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB16FC1BFAAAFC3D" collectingDate="1985-05-23" collectionCode="ZISP" collectorName="B. A. Korotyaev" country="Russia" location="Inal" municipality="Dzhubga" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Kiritshenko">
<locationDeviation id="A1F82F82FFD20E7CFB16FC1BFA91FC53" bearing="315.0" box="[1187,1296,934,955]" distance="10000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFB16FC1BFB60FC53" box="[1187,1249,934,955]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="km" value="10.0">10 km</quantity>
NW
</locationDeviation>
<collectingMunicipality id="502EACDAFFD20E7CFAA2FC1BFAF0FC53" box="[1303,1393,934,955]" pageId="32" pageNumber="33">Dzhubga</collectingMunicipality>
, bay of
<location id="B52A607BFFD20E7CFCF9FC7DFCF5FC3D" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCF9FC7DFCF5FC3D" box="[844,884,960,981]" country="Russia" municipality="Dzhubga" name="Inal" pageId="32" pageNumber="33" stateProvince="Kiritshenko">Inal</location>
,
<date id="C44B1060FFD20E7CFC30FC7DFC69FC3D" box="[901,1000,960,981]" pageId="32" pageNumber="33" value="1985-05-23">
<collectingDate id="D40FE988FFD20E7CFC30FC7DFC69FC3D" box="[901,1000,960,981]" pageId="32" pageNumber="33" value="1985-05-23">23.V.1985</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFC4BFC7DFB25FC3D" box="[1022,1188,960,981]" pageId="32" pageNumber="33">B.A. Korotyaev</collectorName>
), 1J (
<collectionCode id="D6E4AE65FFD20E7CFB5BFC7DFAA5FC3D" box="[1262,1316,960,981]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFA88FC7DFBC0FBE3" collectingDate="1990-05-12" collectionCode="ZISP" collectorName="B. A. Korotyaev" country="Russia" location="Kuban' Riv." municipality="Krasnodar" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Kiritshenko">
<locationDeviation id="A1F82F82FFD20E7CFA88FC7DFA3BFC3D" bearing="112.5" box="[1341,1466,960,981]" distance="26000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFA88FC7DFA00FC3D" box="[1341,1409,960,981]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="32" pageNumber="33" unit="km" value="26.0">26 km</quantity>
ESE
</locationDeviation>
<collectingMunicipality id="502EACDAFFD20E7CFCF9FC66FC32FC18" box="[844,947,987,1008]" pageId="32" pageNumber="33">Krasnodar</collectingMunicipality>
, high bank of
<location id="B52A607BFFD20E7CFBC5FC66FB77FC18" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFBC5FC66FB77FC18" box="[1136,1270,986,1008]" country="Russia" municipality="Krasnodar" name="Kuban' Riv." pageId="32" pageNumber="33" stateProvince="Kiritshenko">Kuban’ Riv.</location>
,
<date id="C44B1060FFD20E7CFAB8FC66FAF1FC18" box="[1293,1392,987,1008]" pageId="32" pageNumber="33" value="1990-05-12">
<collectingDate id="D40FE988FFD20E7CFAB8FC66FAF1FC18" box="[1293,1392,987,1008]" pageId="32" pageNumber="33" value="1990-05-12">12.V.1990</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFA3EFC66FC37FBE3" pageId="32" pageNumber="33">B.A. Korotyaev</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFC7AFC4BFC6CFBE3" box="[975,1005,1013,1035]" count="2" pageId="32" pageNumber="33" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBB6FC4BFBB8FBE3" box="[1027,1081,1014,1035]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBE1FC4BFAC2FBCD" collectionCode="ZISP" country="Russia" location="Belaja" municipality="Belaja" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Republic of Adygeya">
<emphasis id="8281EAB2FFD20E7CFBE1FC4BFAAFFBE3" box="[1108,1326,1014,1035]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFBE1FC4BFAAFFBE3" box="[1108,1326,1014,1035]" pageId="32" pageNumber="33">Republic of Adygeya</collectingRegion>
</emphasis>
: “fl.
<collectingMunicipality id="502EACDAFFD20E7CFAC4FC4BFA2FFBE3" box="[1393,1454,1014,1035]" pageId="32" pageNumber="33">Belaja</collectingMunicipality>
” [date unknown] (Bartholomey),
<specimenCount id="A6F3FD29FFD20E7CFB70FBADFB62FBCE" box="[1221,1251,1040,1062]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFAB3FBADFABDFBCD" box="[1286,1340,1040,1061]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFAD7FBADFC11FBB2" collectingDate="1902-06-14" collectionCode="ZISP" country="Russia" location="Makhoshevskiy" municipality="Maykop" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Republic Of Adygeya">
<collectingMunicipality id="502EACDAFFD20E7CFAD7FBADFA34FBCD" box="[1378,1461,1040,1061]" pageId="32" pageNumber="33">Maykop</collectingMunicipality>
,
<location id="B52A607BFFD20E7CFCFEFB96FC64FBA8" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCFEFB96FC64FBA8" box="[843,997,1067,1088]" country="Russia" municipality="Maykop" name="Makhoshevskiy" pageId="32" pageNumber="33" stateProvince="Republic Of Adygeya">Makhoshevskiy</location>
forest,
<date id="C44B1060FFD20E7CFBE2FB96FB47FBA8" box="[1111,1222,1067,1088]" pageId="32" pageNumber="33" value="1902-06-14">
<collectingDate id="D40FE988FFD20E7CFBE2FB96FB47FBA8" box="[1111,1222,1067,1088]" pageId="32" pageNumber="33" value="1902-06-14">14.VI.1902</collectingDate>
</date>
(Shaposhnikov),
<specimenCount id="A6F3FD29FFD20E7CFA2EFB96FA38FBA8" box="[1435,1465,1066,1088]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFCE1FBF8FC08FBB2" box="[852,905,1093,1114]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFC1AFBF8FC2BFB9D" collectingDate="1932-05-15" collectionCode="ZMUM" country="Russia" location="Maykop" municipality="Maykop" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Republic Of Adygeya">
<collectingMunicipality id="502EACDAFFD20E7CFC1AFBF8FB83FBB2" box="[943,1026,1093,1114]" pageId="32" pageNumber="33">Maykop</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFB94FBF8FB04FBB2" box="[1057,1157,1093,1114]" pageId="32" pageNumber="33" value="1932-05-15">
<collectingDate id="D40FE988FFD20E7CFB94FBF8FB04FBB2" box="[1057,1157,1093,1114]" pageId="32" pageNumber="33" value="1932-05-15">15.V.1932</collectingDate>
</date>
(Shaposhnikov), 1J,
<specimenCount id="A6F3FD29FFD20E7CFA2EFBF8FA38FBB3" box="[1435,1465,1093,1115]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFCE7FBDDFC23FB9D" box="[850,930,1120,1141]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFC6DFBDDFA85FB67" collectingDate="1986-10-04" collectionCode="ZMUM" collectorName="N. Sinichenkova &amp; V. Zherikhin" country="Russia" location="Abadzekhskaya Settlm" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Abadzekhskaya Settlm">
<collectingRegion id="7231F842FFD20E7CFC6DFBDDFB60FB9D" box="[984,1249,1120,1141]" pageId="32" pageNumber="33">Abadzekhskaya Settlm</collectingRegion>
.,
<date id="C44B1060FFD20E7CFAA1FBDDFAEFFB9D" box="[1300,1390,1120,1141]" pageId="32" pageNumber="33" value="1986-10-04">
<collectingDate id="D40FE988FFD20E7CFAA1FBDDFAEFFB9D" box="[1300,1390,1120,1141]" pageId="32" pageNumber="33" value="1986-10-04">4.X.1986</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFA14FBDDFC52FB67" pageId="32" pageNumber="33">N. Sinichenkova</collectorName>
,
<collectorName id="1D005376FFD20E7CFC57FBC7FBE7FB67" box="[994,1126,1146,1167]" pageId="32" pageNumber="33">V. Zherikhin</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFBC9FBC7FB1BFB78" box="[1148,1178,1146,1168]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB18FBC7FB7CFB67" box="[1197,1277,1146,1167]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFAA6FBC7FA34FB42" collectionCode="ZISP" country="Russia" location="Pyatigorsk" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Stavropol' Terr">
<emphasis id="8281EAB2FFD20E7CFAA6FBC7FA2CFB67" box="[1299,1453,1146,1168]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFAA6FBC7FA2CFB67" box="[1299,1453,1146,1168]" pageId="32" pageNumber="33">Stavropol’ Terr</collectingRegion>
</emphasis>
.:
<location id="B52A607BFFD20E7CFCF9FB28FC35FB42" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCF9FB28FC35FB42" box="[844,948,1173,1194]" country="Russia" name="Pyatigorsk" pageId="32" pageNumber="33" stateProvince="Stavropol' Terr">Pyatigorsk</location>
[date and collector unknown],
<specimenCount id="A6F3FD29FFD20E7CFA8EFB28FAD8FB42" box="[1339,1369,1172,1194]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFACDFB28FA2FFB42" box="[1400,1454,1173,1194]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFCF9FB0DFBFDFB37" collectingDate="1913-06-16" collectingDateMax="1950-07-31" collectingDateMin="1913-06-16" collectionCode="ZISP, ZMUM" collectorName="S. Nikulin" country="Russia" location="Essentuki" municipality="Essentuki" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="3" stateProvince="Stavropol' Terr">
<collectingMunicipality id="502EACDAFFD20E7CFCF9FB0DFC2CFB2D" box="[844,941,1200,1221]" pageId="32" pageNumber="33">Essentuki</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFC0CFB0DFBA6FB2D" box="[953,1063,1200,1221]" pageId="32" pageNumber="33" value="1913-06-16">
<collectingDate id="D40FE988FFD20E7CFC0CFB0DFBA6FB2D" box="[953,1063,1200,1221]" pageId="32" pageNumber="33" value="1913-06-16">16.VI.1913</collectingDate>
</date>
(Veshnyakov),
<specimenCount id="A6F3FD29FFD20E7CFB70FB0DFB62FB2D" box="[1221,1251,1199,1221]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB47FB0DFAA9FB2D" box="[1266,1320,1200,1221]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
),
<date id="C44B1060FFD20E7CFA89FB0DFA3BFB2D" box="[1340,1466,1199,1221]" pageId="32" pageNumber="33" value="1950-06" valueMax="1950-07-31" valueMin="1950-06-00">
<collectingDate id="D40FE988FFD20E7CFA89FB0DFA3BFB2D" box="[1340,1466,1199,1221]" pageId="32" pageNumber="33" value="1950-06" valueMax="1950-07-31" valueMin="1950-06-00">VI–VII.1950</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFCE1FB77FC4AFB37" box="[852,971,1226,1247]" pageId="32" pageNumber="33">S. Nikulin</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFC5FFB77FB89FB37" box="[1002,1032,1225,1247]" count="2" pageId="32" pageNumber="33" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB90FB77FBF4FB37" box="[1061,1141,1226,1247]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB20FB77FBBAFAFC" collectingDate="1917-06-15" collectingDateMax="1931-07-10" collectingDateMin="1917-06-15" collectionCode="ZISP, ZMUM" country="Russia" location="Nal' chik" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Kabardino-Balkar Republic">
<emphasis id="8281EAB2FFD20E7CFB20FB77FA32FB37" box="[1173,1459,1226,1247]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFB20FB77FA32FB37" box="[1173,1459,1226,1247]" pageId="32" pageNumber="33">Kabardino-Balkar Republic</collectingRegion>
</emphasis>
:
<location id="B52A607BFFD20E7CFCF9FB58FC23FB12" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCF9FB58FC23FB12" box="[844,930,1252,1274]" country="Russia" name="Nal' chik" pageId="32" pageNumber="33" stateProvince="Kabardino-Balkar Republic">Nal’ chik</location>
,
<date id="C44B1060FFD20E7CFC04FB58FBC5FB12" box="[945,1092,1252,1274]" pageId="32" pageNumber="33" value="1917-06-15" valueMax="1917-06-17" valueMin="1917-06-15">
<collectingDate id="D40FE988FFD20E7CFC04FB58FBC5FB12" box="[945,1092,1252,1274]" pageId="32" pageNumber="33" value="1917-06-15" valueMax="1917-06-17" valueMin="1917-06-15">15–17.VI.1917</collectingDate>
</date>
(Bartenev),
<specimenCount id="A6F3FD29FFD20E7CFB72FB58FB67FB12" box="[1223,1254,1252,1274]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB42FB58FAACFB12" box="[1271,1325,1253,1274]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
),
<date id="C44B1060FFD20E7CFAF6FB58FA38FB12" box="[1347,1465,1253,1274]" pageId="32" pageNumber="33" value="1931-07-10">
<collectingDate id="D40FE988FFD20E7CFAF6FB58FA38FB12" box="[1347,1465,1253,1274]" pageId="32" pageNumber="33" value="1931-07-10">10.VII.1931</collectingDate>
</date>
(Nikulin), 1J (
<collectionCode id="D6E4AE65FFD20E7CFC51FB42FBB5FAFC" box="[996,1076,1279,1300]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBFCFB42FB8BFAC7" collectingDate="1926-07-10" collectionCode="ZISP" country="Russia" location="Kendelen" municipality="Gundelen" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Kabardino-balkar Republic">
“
<collectingMunicipality id="502EACDAFFD20E7CFBE6FB42FB30FAFC" box="[1107,1201,1279,1300]" pageId="32" pageNumber="33">Gundelen</collectingMunicipality>
” [
<location id="B52A607BFFD20E7CFB79FB42FAB1FAFC" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFB79FB42FAB1FAFC" box="[1228,1328,1279,1300]" country="Russia" municipality="Gundelen" name="Kendelen" pageId="32" pageNumber="33" stateProvince="Kabardino-balkar Republic">Kendelen</location>
],
<date id="C44B1060FFD20E7CFAF6FB42FA3BFAFC" box="[1347,1466,1279,1300]" pageId="32" pageNumber="33" value="1926-07-10">
<collectingDate id="D40FE988FFD20E7CFAF6FB42FA3BFAFC" box="[1347,1466,1279,1300]" pageId="32" pageNumber="33" value="1926-07-10">10.VII.1926</collectingDate>
</date>
(Berg),
<specimenCount id="A6F3FD29FFD20E7CFC2DFAA7FC3FFAC7" box="[920,958,1305,1327]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFC78FAA7FB82FAC7" box="[973,1027,1306,1327]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBA3FAA7FBA9FA8C" collectingDate="1886-05-06" collectionCode="ZISP" country="Russia" location="Sapitskaya budka" municipality="Vladikavkaz" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Republic of North Ossetia-Alania">
<emphasis id="8281EAB2FFD20E7CFBA3FAA7FADEFAC7" box="[1046,1375,1306,1327]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFBA3FAA7FADEFAC7" box="[1046,1375,1306,1327]" pageId="32" pageNumber="33">Republic of North Ossetia-Alania</collectingRegion>
</emphasis>
, N spole of Il’ Mt. [
<location id="B52A607BFFD20E7CFC64FA89FB0DFAA1" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFC64FA89FB0DFAA1" box="[977,1164,1332,1353]" country="Russia" municipality="Vladikavkaz" name="Sapitskaya budka" pageId="32" pageNumber="33" stateProvince="Republic of North Ossetia-Alania">Sapitskaya budka</location>
] SE
<collectingMunicipality id="502EACDAFFD20E7CFB65FA89FACCFAA1" box="[1232,1357,1332,1353]" pageId="32" pageNumber="33">Vladikavkaz</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFAD7FA89FA38FAA1" box="[1378,1465,1332,1353]" pageId="32" pageNumber="33" value="1886-05-06">
<collectingDate id="D40FE988FFD20E7CFAD7FA89FA38FAA1" box="[1378,1465,1332,1353]" pageId="32" pageNumber="33" value="1886-05-06">6.V.1886</collectingDate>
</date>
(Ananov),
<specimenCount id="A6F3FD29FFD20E7CFC0FFAF2FC59FA8C" box="[954,984,1358,1380]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFC5EFAF2FBA0FA8C" box="[1003,1057,1359,1380]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB82FAF2FBF9FA96" collectingDate="1901-06-04" collectionCode="ZISP" country="Russia" location="Khasavyurt" municipality="Khasavyurt" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Republic of Dagestan">
<emphasis id="8281EAB2FFD20E7CFB82FAF2FA95FA8C" box="[1079,1300,1359,1380]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFB82FAF2FA95FA8C" box="[1079,1300,1359,1380]" pageId="32" pageNumber="33">Republic of Dagestan</collectingRegion>
</emphasis>
:
<collectingMunicipality id="502EACDAFFD20E7CFA93FAF2FA19FA8C" box="[1318,1432,1359,1380]" pageId="32" pageNumber="33">Khasavyurt</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFA12FAF2FC1AFA96" pageId="32" pageNumber="33" value="1901-06-04">
<collectingDate id="D40FE988FFD20E7CFA12FAF2FC1AFA96" pageId="32" pageNumber="33" value="1901-06-04">4. VI.1901</collectingDate>
</date>
(Bekman),
<specimenCount id="A6F3FD29FFD20E7CFBBAFAD4FBACFA97" box="[1039,1069,1385,1407]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB8EFAD4FBF0FA96" box="[1083,1137,1385,1406]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB37FAD4FBD4FA71" collectingDate="1925-06-08" collectionCode="ZISP" country="Russia" location="Makhachkala" municipality="Makhachkala" pageId="32" pageNumber="33" specimenCount="5" specimenCount-female="5" stateProvince="Kiritshenko">
<collectingMunicipality id="502EACDAFFD20E7CFB37FAD4FA86FA96" box="[1154,1287,1385,1406]" pageId="32" pageNumber="33">Makhachkala</collectingMunicipality>
, 1,2.V.,
<date id="C44B1060FFD20E7CFAE2FAD4FA38FA96" box="[1367,1465,1385,1406]" pageId="32" pageNumber="33" value="1925-06-08">
<collectingDate id="D40FE988FFD20E7CFAE2FAD4FA38FA96" box="[1367,1465,1385,1406]" pageId="32" pageNumber="33" value="1925-06-08">8.VI.1925</collectingDate>
</date>
(Kiritshenko),
<specimenCount id="A6F3FD29FFD20E7CFC51FA39FB83FA72" box="[996,1026,1412,1434]" count="5" pageId="32" pageNumber="33" type="female">5♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBADFA39FBCFFA71" box="[1048,1102,1412,1433]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBDCFA39FC3BFA26" collectingDate="1972-06-03" collectionCode="ZISP" country="Russia" location="Sovetskoe [Khebda] Settlm" municipality="Avarskoe Koysu Riv." pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Kiritshenko">
gorge of
<collectingMunicipality id="502EACDAFFD20E7CFB67FA39FA37FA71" box="[1234,1462,1412,1433]" pageId="32" pageNumber="33">Avarskoe Koysu Riv.</collectingMunicipality>
,
<locationDeviation id="A1F82F82FFD20E7CFCF9FA22FC23FA5C" bearing="180.0" box="[844,930,1439,1460]" distance="14000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFCF9FA22FC0AFA5C" box="[844,907,1439,1460]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="32" pageNumber="33" unit="km" value="14.0">14 km</quantity>
S
</locationDeviation>
<location id="B52A607BFFD20E7CFC1EFA22FB40FA5C" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFC1EFA22FB40FA5C" box="[939,1217,1439,1460]" country="Russia" municipality="Avarskoe Koysu Riv." name="Sovetskoe [Khebda] Settlm" pageId="32" pageNumber="33" stateProvince="Kiritshenko">Sovetskoe [Khebda] Settlm</location>
.,
<date id="C44B1060FFD20E7CFB65FA22FAB2FA5C" box="[1232,1331,1439,1460]" pageId="32" pageNumber="33" value="1972-06-03">
<collectingDate id="D40FE988FFD20E7CFB65FA22FAB2FA5C" box="[1232,1331,1439,1460]" pageId="32" pageNumber="33" value="1972-06-03">3.VI.1972</collectingDate>
</date>
(Kasparyan),
<specimenCount id="A6F3FD29FFD20E7CFCF9FA04FCEBFA27" box="[844,874,1465,1487]" count="2" pageId="32" pageNumber="33" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFCC8FA04FC32FA26" box="[893,947,1465,1486]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFC7FFA04FB41FA01" collectingDate="1983-07-15" collectionCode="ZISP" collectorName="I. M. Kerzhner" country="Russia" location="Sergokala Settlm" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Kiritshenko">
<locationDeviation id="A1F82F82FFD20E7CFC7FFA04FBBCFA26" bearing="225.0" box="[970,1085,1465,1486]" distance="15000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFC7FFA04FB8DFA26" box="[970,1036,1465,1486]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="km" value="15.0">15 km</quantity>
SW
</locationDeviation>
<location id="B52A607BFFD20E7CFBFCFA04FB78FA26" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFBFCFA04FB78FA26" box="[1097,1273,1465,1486]" country="Russia" name="Sergokala Settlm" pageId="32" pageNumber="33" stateProvince="Kiritshenko">Sergokala Settlm</location>
., dry meadow,
<date id="C44B1060FFD20E7CFA2EFA04FC22FA01" pageId="32" pageNumber="33" value="1983-07-15">
<collectingDate id="D40FE988FFD20E7CFA2EFA04FC22FA01" pageId="32" pageNumber="33" value="1983-07-15">15. VII.1983</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFC01FA69FBC2FA01" box="[948,1091,1492,1513]" pageId="32" pageNumber="33">I.M. Kerzhner</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFBE3FA69FBF5FA01" box="[1110,1140,1491,1513]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB36FA69FB38FA01" box="[1155,1209,1492,1513]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB79FA69FBCEF9F6" collectingDate="1983-07-22" collectionCode="ZISP" collectorName="I. M. Kerzhner" country="Russia" elevation="1500" location="Akhty Settlm" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Kiritshenko">
<locationDeviation id="A1F82F82FFD20E7CFB79FA69FAA0FA01" bearing="90.0" box="[1228,1313,1492,1513]" distance="10000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFB79FA69FA8AFA01" box="[1228,1291,1492,1513]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="km" value="10.0">10 km</quantity>
E
</locationDeviation>
<location id="B52A607BFFD20E7CFA9CFA69FA33FA01" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFA9CFA69FA33FA01" box="[1321,1458,1492,1513]" country="Russia" name="Akhty Settlm" pageId="32" pageNumber="33" stateProvince="Kiritshenko">Akhty Settlm</location>
.,
<quantity id="770D9B45FFD20E7CFCF9FA53FC54F9EB" box="[844,981,1518,1540]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">
<elevation id="3BD8D193FFD20E7CFCF9FA53FC54F9EB" box="[844,981,1518,1540]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">1000–2000 m</elevation>
</quantity>
amsl, steppes and meadows,
<date id="C44B1060FFD20E7CFAB2FA53FAFFF9EB" box="[1287,1406,1518,1539]" pageId="32" pageNumber="33" value="1983-07-22">
<collectingDate id="D40FE988FFD20E7CFAB2FA53FAFFF9EB" box="[1287,1406,1518,1539]" pageId="32" pageNumber="33" value="1983-07-22">22.VII.1983</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFA25FA53FC2BF9F6" pageId="32" pageNumber="33">I.M. Kerzhner</collectorName>
), 1J,
<specimenCount id="A6F3FD29FFD20E7CFC53F9B4FB85F9F6" box="[998,1028,1544,1566]" count="2" pageId="32" pageNumber="33" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBA7F9B4FBC9F9F6" box="[1042,1096,1545,1566]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBEFF9B4FA37F9D0" collectingDate="1992-05-21" collectionCode="ZISP" collectorName="M. G. Volkovitsh" country="Russia" location="Sarykum sands" municipality="Makhachkala" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Kumtorkala">
<collectingRegion id="7231F842FFD20E7CFBEFF9B4FB51F9F6" box="[1114,1232,1545,1566]" pageId="32" pageNumber="33">Kumtorkala</collectingRegion>
,
<location id="B52A607BFFD20E7CFB6FF9B4FAF1F9F6" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFB6FF9B4FAF1F9F6" box="[1242,1392,1545,1566]" country="Russia" municipality="Makhachkala" name="Sarykum sands" pageId="32" pageNumber="33" stateProvince="Kumtorkala">Sarykum sands</location>
,
<locationDeviation id="A1F82F82FFD20E7CFACEF9B4FC0BF9D0" bearing="292.5" distance="20000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFACEF9B4FA3BF9F6" box="[1403,1466,1545,1566]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="km" value="20.0">20 km</quantity>
WNW
</locationDeviation>
<collectingMunicipality id="502EACDAFFD20E7CFC24F99EFB97F9D0" box="[913,1046,1571,1592]" pageId="32" pageNumber="33">Makhachkala</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFB94F99EFB04F9D0" box="[1057,1157,1571,1592]" pageId="32" pageNumber="33" value="1992-05-21">
<collectingDate id="D40FE988FFD20E7CFB94F99EFB04F9D0" box="[1057,1157,1571,1592]" pageId="32" pageNumber="33" value="1992-05-21">21.V.1992</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFB21F99EFABAF9D0" box="[1172,1339,1571,1592]" pageId="32" pageNumber="33">M.G. Volkovitsh</collectorName>
), 1J (
<collectionCode id="D6E4AE65FFD20E7CFACCF99EFA2EF9D0" box="[1401,1455,1571,1592]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD20E7CFCF9F983FAC9F9BB" box="[844,1352,1598,1620]" collectingDate="1903-04-23" collectionCode="ZISP" country="Georgia" location="Tiflis" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Tiflis">
<emphasis id="8281EAB2FFD20E7CFCF9F983FC1EF9BB" bold="true" box="[844,927,1598,1619]" pageId="32" pageNumber="33">
<collectingCountry id="C8E27630FFD20E7CFCF9F983FC1EF9BB" box="[844,927,1598,1619]" name="Georgia" pageId="32" pageNumber="33" value="Georgia">Georgia</collectingCountry>
</emphasis>
:
<collectingRegion id="7231F842FFD20E7CFC07F983FC67F9BB" box="[946,998,1598,1620]" country="Georgia" name="Tbilisi" pageId="32" pageNumber="33">Tiflis</collectingRegion>
,
<date id="C44B1060FFD20E7CFC43F983FBE3F9BB" box="[1014,1122,1598,1619]" pageId="32" pageNumber="33" value="1903-04-23">
<collectingDate id="D40FE988FFD20E7CFC43F983FBE3F9BB" box="[1014,1122,1598,1619]" pageId="32" pageNumber="33" value="1903-04-23">23.IV.1903</collectingDate>
</date>
(Satunin),
<specimenCount id="A6F3FD29FFD20E7CFB6FF983FB79F9BC" box="[1242,1272,1598,1620]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFABEF983FAC0F9BB" box="[1291,1345,1598,1619]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD20E7CFAEDF983FBBDF960" collectingDate="1937-08-08" collectionCode="ZISP" country="Armenia" location="Vanadzor" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Lori Prov">
<emphasis id="8281EAB2FFD20E7CFAEDF983FA32F9BB" bold="true" box="[1368,1459,1598,1619]" pageId="32" pageNumber="33">
<collectingCountry id="C8E27630FFD20E7CFAEDF983FA32F9BB" box="[1368,1459,1598,1619]" name="Armenia" pageId="32" pageNumber="33" value="Armenia">Armenia</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD20E7CFCF9F9E5FC3AF985" box="[844,955,1624,1645]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFCF9F9E5FC3AF985" box="[844,955,1624,1645]" pageId="32" pageNumber="33">Lori Prov</collectingRegion>
</emphasis>
., “Kirovakan” [
<location id="B52A607BFFD20E7CFBCFF9E5FB5CF985" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFBCFF9E5FB5CF985" box="[1146,1245,1624,1645]" country="Armenia" name="Vanadzor" pageId="32" pageNumber="33" stateProvince="Lori Prov">Vanadzor</location>
],
<date id="C44B1060FFD20E7CFB4AF9E5FAF0F985" box="[1279,1393,1624,1645]" pageId="32" pageNumber="33" value="1937-08-08">
<collectingDate id="D40FE988FFD20E7CFB4AF9E5FAF0F985" box="[1279,1393,1624,1645]" pageId="32" pageNumber="33" value="1937-08-08">8.VIII.1937</collectingDate>
</date>
(Ter-Minassjan),
<specimenCount id="A6F3FD29FFD20E7CFC7EF9CEFC68F961" box="[971,1001,1651,1673]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFC4BF9CEFBB5F960" box="[1022,1076,1651,1672]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBFBF9CEFBB6F94B" collectingDate="1931-07-01" collectionCode="ZISP" collectorName="A. Koton" country="Armenia" location="Dilizhan" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Tavush Prov">
<emphasis id="8281EAB2FFD20E7CFBFBF9CEFB4CF960" box="[1102,1229,1651,1672]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFBFBF9CEFB4CF960" box="[1102,1229,1651,1672]" pageId="32" pageNumber="33">Tavush Prov</collectingRegion>
</emphasis>
.:
<location id="B52A607BFFD20E7CFB52F9CEFABCF960" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFB52F9CEFABCF960" box="[1255,1341,1651,1672]" country="Armenia" name="Dilizhan" pageId="32" pageNumber="33" stateProvince="Tavush Prov">Dilizhan</location>
,
<date id="C44B1060FFD20E7CFAFAF9CEFA38F960" box="[1359,1465,1651,1672]" pageId="32" pageNumber="33" value="1931-07-01">
<collectingDate id="D40FE988FFD20E7CFAFAF9CEFA38F960" box="[1359,1465,1651,1672]" pageId="32" pageNumber="33" value="1931-07-01">1.VII.1931</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFCE6F933FC37F94B" box="[851,950,1678,1699]" pageId="32" pageNumber="33">A. Koton</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFC7EF933FC68F94B" box="[971,1001,1677,1699]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFC4FF933FBB1F94B" box="[1018,1072,1678,1699]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFBF3F933FC11F955" collectingDate="1955-05-23" collectionCode="ZISP" country="Armenia" location="Idzhevan" municipality="Idzhevan" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Tavush Prov">
<collectingMunicipality id="502EACDAFFD20E7CFBF3F933FB20F94B" box="[1094,1185,1678,1699]" pageId="32" pageNumber="33">Idzhevan</collectingMunicipality>
,
<date id="C44B1060FFD20E7CFB1AF933FA92F94B" box="[1199,1299,1678,1699]" pageId="32" pageNumber="33" value="1955-05-23">
<collectingDate id="D40FE988FFD20E7CFB1AF933FA92F94B" box="[1199,1299,1678,1699]" pageId="32" pageNumber="33" value="1955-05-23">23.V.1955</collectingDate>
</date>
(Loginova),
<specimenCount id="A6F3FD29FFD20E7CFA2EF933FA38F94B" box="[1435,1465,1677,1699]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFCE7F915FC09F955" box="[850,904,1704,1725]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFC15F915FB81F91A" collectingDate="1934-06-13" collectionCode="ZISP" country="Armenia" location="Aragats Mt." municipality="Alagez" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Aragatsotn prov" typeStatus="paratype">
<emphasis id="8281EAB2FFD20E7CFC15F915FBC3F955" box="[928,1090,1704,1725]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFC15F915FBC3F955" box="[928,1090,1704,1725]" pageId="32" pageNumber="33">Aragatsotn prov</collectingRegion>
</emphasis>
., Kipchakh, NW slope of “
<collectingMunicipality id="502EACDAFFD20E7CFADFF915FA2FF955" box="[1386,1454,1704,1725]" pageId="32" pageNumber="33">Alagez</collectingMunicipality>
” [
<location id="B52A607BFFD20E7CFCE4F97EFC51F930" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCE4F97EFC51F930" box="[849,976,1731,1752]" country="Armenia" municipality="Alagez" name="Aragats Mt." pageId="32" pageNumber="33" stateProvince="Aragatsotn prov">Aragats Mt.</location>
],
<date id="C44B1060FFD20E7CFC50F97EFBD5F930" box="[997,1108,1731,1752]" pageId="32" pageNumber="33" value="1934-06-13">
<collectingDate id="D40FE988FFD20E7CFC50F97EFBD5F930" box="[997,1108,1731,1752]" pageId="32" pageNumber="33" value="1934-06-13">13.VI.1934</collectingDate>
</date>
(Ter-Minassjan),
<specimenCount id="A6F3FD29FFD20E7CFABEF97EFAA8F930" box="[1291,1321,1730,1752]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFA8EF97EFAEEF930" box="[1339,1391,1731,1752]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
) [
<typeStatus id="6F4E8802FFD20E7CFA33F97EFCF7F91A" pageId="32" pageNumber="33">paratype</typeStatus>
of
<emphasis id="8281EAB2FFD20E7CFC2DF960FB41F91A" box="[920,1216,1757,1778]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="77F54D23FFD20E7CFC2DF960FC77F91A" authorityName="Asanova" authorityYear="1964" box="[920,1014,1757,1778]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mixtus">C. mixtus</taxonomicName>
];
<materialsCitation id="009D3CFDFFD20E7CFBB8F960FAB7F8E5" collectingDate="1902-05-24" collectionCode="ZISP" country="Armenia" location="Sevan] Lake" municipality="Gokcha" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Gegharkunik Prov">
<collectingRegion id="7231F842FFD20E7CFBB8F960FB41F91A" box="[1037,1216,1757,1778]" pageId="32" pageNumber="33">Gegharkunik Prov</collectingRegion>
., NW shore of “
<collectingMunicipality id="502EACDAFFD20E7CFAD1F960FA2FF91B" box="[1380,1454,1757,1779]" pageId="32" pageNumber="33">Gokchа</collectingMunicipality>
” [
<location id="B52A607BFFD20E7CFCE4F945FC54F8E5" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFCE4F945FC54F8E5" box="[849,981,1784,1805]" country="Armenia" municipality="Gokcha" name="Sevan] Lake" pageId="32" pageNumber="33" stateProvince="Gegharkunik Prov">Sevan] Lake</location>
,
<date id="C44B1060FFD20E7CFC56F945FBC6F8E5" box="[995,1095,1784,1805]" pageId="32" pageNumber="33" value="1902-05-24">
<collectingDate id="D40FE988FFD20E7CFC56F945FBC6F8E5" box="[995,1095,1784,1805]" pageId="32" pageNumber="33" value="1902-05-24">24.V.1902</collectingDate>
</date>
(Elachich),
<specimenCount id="A6F3FD29FFD20E7CFB7DF945FB67F8E5" box="[1224,1254,1783,1805]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB4DF945FAAFF8E5" box="[1272,1326,1784,1805]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
</emphasis>
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFAF0F945FB28F8C0" collectingDate="1948-06-24" collectionCode="ZISP" country="Armenia" location="Erevan" municipality="Erevan" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Gegharkunik Prov">
<emphasis id="8281EAB2FFD20E7CFAF0F945FA0BF8E5" box="[1349,1418,1784,1805]" italics="true" pageId="32" pageNumber="33">
<collectingMunicipality id="502EACDAFFD20E7CFAF0F945FA0BF8E5" box="[1349,1418,1784,1805]" pageId="32" pageNumber="33">Erevan</collectingMunicipality>
</emphasis>
,
<date id="C44B1060FFD20E7CFA2EF945FC1DF8C0" pageId="32" pageNumber="33" value="1948-06-24">
<collectingDate id="D40FE988FFD20E7CFA2EF945FC1DF8C0" pageId="32" pageNumber="33" value="1948-06-24">24. VI.1948</collectingDate>
</date>
(Borchsenius),
<specimenCount id="A6F3FD29FFD20E7CFB88F8AEFBDAF8C0" box="[1085,1115,1810,1832]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBDEF8AEFB20F8C0" box="[1131,1185,1811,1832]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD20E7CFB03F8AEFAAFF8AA" collectingDate="1972-05-31" collectionCode="ZISP" collectorName="E. Sugonjaev" country="Armenia" location="Khosrov Forest" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Ararat Province">
<emphasis id="8281EAB2FFD20E7CFB03F8AEFAD7F8C0" box="[1206,1366,1811,1832]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFB03F8AEFAD7F8C0" box="[1206,1366,1811,1832]" pageId="32" pageNumber="33">Ararat Province</collectingRegion>
</emphasis>
,
<location id="B52A607BFFD20E7CFAD3F8AEFC0AF8AA" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFAD3F8AEFC0AF8AA" country="Armenia" name="Khosrov Forest" pageId="32" pageNumber="33" stateProvince="Ararat Province">Khosrov Forest</location>
,
<date id="C44B1060FFD20E7CFC28F890FB81F8AA" box="[925,1024,1837,1858]" pageId="32" pageNumber="33" value="1972-05-31">
<collectingDate id="D40FE988FFD20E7CFC28F890FB81F8AA" box="[925,1024,1837,1858]" pageId="32" pageNumber="33" value="1972-05-31">31.V.1972</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFBA3F890FB25F8AA" box="[1046,1188,1837,1858]" pageId="32" pageNumber="33">E. Sugonjaev</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFB08F890FB5AF8AB" box="[1213,1243,1837,1859]" count="2" pageId="32" pageNumber="33" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFB44F890FAA6F8AA" box="[1265,1319,1837,1858]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD20E7CFA8AF890FBDEF89F" collectingDate="1965-07-12" collectionCode="ZMUM" country="Azerbaijan" elevation="1800" location="Altyagach" municipality="Altyagach" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Khizi Distr">
<emphasis id="8281EAB2FFD20E7CFA8AF890FA33F8AA" bold="true" box="[1343,1458,1837,1858]" pageId="32" pageNumber="33">
<collectingCountry id="C8E27630FFD20E7CFA8AF890FA33F8AA" box="[1343,1458,1837,1858]" name="Azerbaijan" pageId="32" pageNumber="33" value="Azerbaijan">Azerbaijan</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD20E7CFCF9F8F5FC4AF8B5" box="[844,971,1864,1885]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFCF9F8F5FC4AF8B5" box="[844,971,1864,1885]" pageId="32" pageNumber="33">Khizi Distr</collectingRegion>
</emphasis>
.,
<collectingMunicipality id="502EACDAFFD20E7CFC46F8F5FBD8F8B5" box="[1011,1113,1864,1885]" pageId="32" pageNumber="33">Altyagach</collectingMunicipality>
,
<quantity id="770D9B45FFD20E7CFBCCF8F5FB56F8B5" box="[1145,1239,1864,1885]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="32" pageNumber="33" unit="m" value="1800.0">
<elevation id="3BD8D193FFD20E7CFBCCF8F5FB56F8B5" box="[1145,1239,1864,1885]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="32" pageNumber="33" unit="m" value="1800.0">1800 m</elevation>
</quantity>
amsl,
<date id="C44B1060FFD20E7CFAF6F8F5FA38F8B5" box="[1347,1465,1864,1885]" pageId="32" pageNumber="33" value="1965-07-12">
<collectingDate id="D40FE988FFD20E7CFAF6F8F5FA38F8B5" box="[1347,1465,1864,1885]" pageId="32" pageNumber="33" value="1965-07-12">12.VII.1965</collectingDate>
</date>
(Gidayatov),
<specimenCount id="A6F3FD29FFD20E7CFC60F8DFFC72F890" box="[981,1011,1890,1912]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFBBDF8DFFBD8F89F" box="[1032,1113,1890,1911]" collectionName="Russia, Moscow, Moscow State University" pageId="32" pageNumber="33">ZMUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD20E7CFBC5F8DFFA37F87A" collectingDate="1997-06-10" collectionCode="ZISP" collectorName="M. Volkovitsh" country="Turkey" location="Oltu" municipality="Sutkas def." pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Erzurum Prov">
<emphasis id="8281EAB2FFD20E7CFBC5F8DFFB3BF89F" bold="true" box="[1136,1210,1890,1911]" pageId="32" pageNumber="33">
<collectingCountry id="C8E27630FFD20E7CFBC5F8DFFB3BF89F" box="[1136,1210,1890,1911]" name="Turkey" pageId="32" pageNumber="33" value="Turkey">Turkey</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD20E7CFB7AF8DFFADEF89F" box="[1231,1375,1890,1911]" italics="true" pageId="32" pageNumber="33">
<collectingRegion id="7231F842FFD20E7CFB7AF8DFFADEF89F" box="[1231,1375,1890,1911]" pageId="32" pageNumber="33">Erzurum Prov</collectingRegion>
</emphasis>
.,
<collectingMunicipality id="502EACDAFFD20E7CFACCF8DFFCF5F87A" pageId="32" pageNumber="33">Sutkas def.</collectingMunicipality>
,
<emphasis id="8281EAB2FFD20E7CFC34F8C0FC0AF87A" box="[897,907,1917,1938]" italics="true" pageId="32" pageNumber="33">c</emphasis>
.
<locationDeviation id="A1F82F82FFD20E7CFC3AF8C0FC5CF87A" bearing="180.0" box="[911,989,1917,1938]" distance="8000.0" pageId="32" pageNumber="33">
<quantity id="770D9B45FFD20E7CFC3AF8C0FC46F87A" box="[911,967,1917,1938]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="32" pageNumber="33" unit="km" value="8.0">8 km</quantity>
S
</locationDeviation>
<location id="B52A607BFFD20E7CFC52F8C0FB95F87A" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD20E7CFC52F8C0FB95F87A" box="[999,1044,1917,1938]" country="Turkey" municipality="Sutkas def." name="Oltu" pageId="32" pageNumber="33" stateProvince="Erzurum Prov">Oltu</location>
,
<date id="C44B1060FFD20E7CFB97F8C0FB10F87A" box="[1058,1169,1917,1938]" pageId="32" pageNumber="33" value="1997-06-10">
<collectingDate id="D40FE988FFD20E7CFB97F8C0FB10F87A" box="[1058,1169,1917,1938]" pageId="32" pageNumber="33" value="1997-06-10">10.VI.1997</collectingDate>
</date>
(
<collectorName id="1D005376FFD20E7CFB16F8C0FAB7F87A" box="[1187,1334,1917,1938]" pageId="32" pageNumber="33">M. Volkovitsh</collectorName>
),
<specimenCount id="A6F3FD29FFD20E7CFAFCF8C0FAE9F87A" box="[1353,1384,1916,1938]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD20E7CFACDF8C0FA2FF87A" box="[1400,1454,1917,1938]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="32" pageNumber="33">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD30E7DFF2BFF7CFE88FF19" collectionCode="ZISP" collectorName="A. V. Xienzopolski" country="Turkey" location="Kazikoparan" pageId="33" pageNumber="34" specimenCount="2" specimenCount-female="2" stateProvince="Igdir Prov">
<emphasis id="8281EAB2FFD30E7DFF2BFF7CFE85FF3E" box="[158,260,193,215]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="7231F842FFD30E7DFF2BFF7CFE85FF3E" box="[158,260,193,215]" pageId="33" pageNumber="34">Iğdır Prov</collectingRegion>
</emphasis>
.,
<location id="B52A607BFFD30E7DFEACFF7CFE15FF3E" LSID="urn:lsid:plazi:treatment:385C87B6FFD20E7DFCCEFE84FC8DF929:B52A607BFFD30E7DFEACFF7CFE15FF3E" box="[281,404,193,214]" country="Turkey" name="Kazikoparan" pageId="33" pageNumber="34" stateProvince="Igdir Prov">Kazikoparan</location>
[date unknown] (
<collectorName id="1D005376FFD30E7DFDF2FF7CFC83FF3E" box="[583,770,193,214]" pageId="33" pageNumber="34">A.V. Xienzopolski</collectorName>
),
<specimenCount id="A6F3FD29FFD30E7DFF2BFF61FF3DFF1A" box="[158,188,220,242]" count="2" pageId="33" pageNumber="34" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD30E7DFF7EFF61FE80FF19" box="[203,257,220,241]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="33" pageNumber="34">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFF2BFEFAFF74FCAC" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="B04A36A0FFD30E7DFF2BFEFAFE8AFEB6" blockId="33.[158,781,193,1849]" box="[158,267,327,350]" pageId="33" pageNumber="34">
<heading id="EB0281CCFFD30E7DFF2BFEFAFE8AFEB6" bold="true" box="[158,267,327,350]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFF2BFEFAFE8AFEB6" bold="true" box="[158,267,327,350]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD30E7DFF2BFED2FF74FCAC" blockId="33.[158,781,193,1849]" pageId="33" pageNumber="34">
Ventral lobe of conjunctiva at extreme apex rather sharply widened and widely rounded in ventral view; dorsal surface of lobe apex bearing short longitudinal sclerotised carina with apical end visible in ventral view and large impressions at sides of carina (
<figureCitation id="28CE2A25FFD30E7DFDE2FE53FD7DFDED" box="[599,764,494,517]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="33" pageNumber="34">
Figure 6
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, 6
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). Spicule at conjunctival dorsolateral lobes (
<figureCitation id="28CE2A25FFD30E7DFD22FDB3FC82FDCD" box="[663,771,526,549]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="33" pageNumber="34">
Figure
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7
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) less widely curved basally and comparatively shorter: denticle on spicule basal plate extending far beyond middle of spicule length; beyond middle, spicule curved arcuately outwards rather weakly; extreme apex of spicule curved inwards relative to its axis so that outer margin of spicule before apex forms obtuse angle. Denticle on spicule basal plate rather short, curved at right angle in middle, with rectangular apex directed mediad.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFF2BFC27FC80FAF0" pageId="33" pageNumber="34" type="discussion">
<paragraph id="B04A36A0FFD30E7DFF2BFC27FE84FC59" blockId="33.[158,781,193,1849]" box="[158,261,922,945]" pageId="33" pageNumber="34">
<heading id="EB0281CCFFD30E7DFF2BFC27FE84FC59" bold="true" box="[158,261,922,945]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFF2BFC27FE84FC59" bold="true" box="[158,261,922,945]" pageId="33" pageNumber="34">Remarks</emphasis>
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</paragraph>
<paragraph id="B04A36A0FFD30E7DFF2BFC7FFC80FAF0" blockId="33.[158,781,193,1849]" pageId="33" pageNumber="34">
According to
<bibRefCitation id="D4644B51FFD30E7DFE81FC7FFE60FC31" author="Asanova RB" box="[308,481,962,985]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="33" pageNumber="34" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova (1964)</bibRefCitation>
,
<specimenCount id="A6F3FD29FFD30E7DFE5AFC7FFDE0FC31" box="[495,609,962,985]" count="4" pageId="33" pageNumber="34" type="male">four males</specimenCount>
from her material differ from all the other males of the species in the shorter spicule, not curved before apex, and in the more massive base of denticle on the anterior end of spicule basal plate. I found in the
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collection only
<specimenCount id="A6F3FD29FFD30E7DFD2AFBFFFC8DFBB1" box="[671,780,1090,1113]" count="2" pageId="33" pageNumber="34" type="male">two males</specimenCount>
from the listed four, and they do not have the distinctions described by Asanova, like all the males examined by me from Armenia. On the other hand,
<bibRefCitation id="D4644B51FFD30E7DFD8AFB1CFF64FB30" author="Kment P &amp; Jindra Z." journalOrPublisher="Acta Entomologica Musei Nationalis Pragae." pageId="33" pageNumber="34" pagination="3 - 16" part="45" refId="ref40164" refString="Kment P, Jindra Z. 2005. New and interesting records of true bugs (Heteroptera) from Turkey, southeastern Europe, Near and Middle East. Acta Entomologica Musei Nationalis Pragae. 45: 3 - 16." title="New and interesting records of true bugs (Heteroptera) from Turkey, southeastern Europe, Near and Middle East" type="journal article" year="2005">Kment and Jindra (2005)</bibRefCitation>
pointed out that the dorsolateral lobes in specimens collected in Turkey have arming almost identical to those figured in Asanova’ s paper for the Armenian specimens.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFF2BFAD3FDFEF9E5" pageId="33" pageNumber="34" type="description">
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<emphasis id="8281EAB2FFD30E7DFF2BFAD3FEC2FA6D" bold="true" box="[158,323,1390,1413]" pageId="33" pageNumber="34">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD30E7DFF2BFA2BFDFEF9E5" blockId="33.[158,781,193,1849]" pageId="33" pageNumber="34">
Body 5.00–
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length, 3.00–
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width; pronotum
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width; head
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width; ocular index 3.18–3.68; length of antennal segments I–V: 0.28–0.38: 0.46–0.67: 0.43–0.87: 0.76–0.97: 0.85–1.06.
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<subSubSection id="F8EF652BFFD30E7DFF2BF9DFFC8DF929" pageId="33" pageNumber="34" type="distribution">
<paragraph id="B04A36A0FFD30E7DFF2BF9DFFEA6F991" blockId="33.[158,781,193,1849]" box="[158,295,1634,1657]" pageId="33" pageNumber="34">
<heading id="EB0281CCFFD30E7DFF2BF9DFFEA6F991" bold="true" box="[158,295,1634,1657]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFF2BF9DFFEA6F991" bold="true" box="[158,295,1634,1657]" pageId="33" pageNumber="34">Distribution</emphasis>
</heading>
</paragraph>
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Russia (Caucasus), Georgia, Armenia, Azerbaijan (
<bibRefCitation id="D4644B51FFD30E7DFD16F937FD7CF949" author="Lis JA" box="[675,765,1674,1697]" editor="Aukema B &amp; Rieger C" journalOrPublisher="Amsterdam: Netherlands Entomological Society" pageId="33" pageNumber="34" pagination="119 - 147" refId="ref40328" refString="Lis JA. 2006. Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs). In: Aukema B and Rieger C, editors. Catalogue of the Heteroptera of the Palaearctic Region Vol. 5, Amsterdam: Netherlands Entomological Society; p. 119 - 147." title="Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs)" type="book chapter" volumeTitle="Catalogue of the Heteroptera of the Palaearctic Region" year="2006">Lis 2006</bibRefCitation>
), Turkey (
<bibRefCitation id="D4644B51FFD30E7DFF46F917FE4CF929" author="Kment P &amp; Jindra Z." box="[243,461,1706,1729]" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae." pageId="33" pageNumber="34" pagination="3 - 16" part="45" refId="ref40164" refString="Kment P, Jindra Z. 2005. New and interesting records of true bugs (Heteroptera) from Turkey, southeastern Europe, Near and Middle East. Acta Entomologica Musei Nationalis Pragae. 45: 3 - 16." title="New and interesting records of true bugs (Heteroptera) from Turkey, southeastern Europe, Near and Middle East" type="journal article" year="2005">Kment &amp; Jindra 2005</bibRefCitation>
), and Iran (
<bibRefCitation id="D4644B51FFD30E7DFD8BF917FD7CF929" author="Ghahari H &amp; Cherot F &amp; Linnavuori RE &amp; Ostovan H." box="[574,765,1706,1729]" journalOrPublisher="ZooKeys." pageId="33" pageNumber="34" pagination="1 - 31" part="26" refId="ref39906" refString="Ghahari H, Cherot F, Linnavuori RE, Ostovan H. 2009. Annotated catalogue of Iranian burrower bugs (Heteroptera, Pentatomoidea, Cydnidae). ZooKeys. 26: 1 - 31." title="Annotated catalogue of Iranian burrower bugs (Heteroptera, Pentatomoidea, Cydnidae)" type="journal article" year="2009">Ghahari et al. 2009</bibRefCitation>
).
</paragraph>
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(
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Figures 6
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,
<figureCitation id="28CE2A25FFD30E7DFD91F89FFDBEF8D1" box="[548,575,1826,1849]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="33" pageNumber="34">
7
<collectionCode id="D6E4AE65FFD30E7DFD84F89FFDBEF8D1" box="[561,575,1826,1849]" pageId="33" pageNumber="34">J</collectionCode>
</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFCF9FF7FFC6AFF31" box="[844,1003,194,217]" pageId="33" pageNumber="34" type="multiple">
<paragraph id="B04A36A0FFD30E7DFCF9FF7FFC6AFF31" blockId="33.[844,1466,194,1850]" box="[844,1003,194,217]" pageId="33" pageNumber="34">
<heading id="EB0281CCFFD30E7DFCF9FF7FFC6AFF31" bold="true" box="[844,1003,194,217]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFCF9FF7FFC6AFF31" bold="true" box="[844,1003,194,217]" pageId="33" pageNumber="34">
<typeStatus id="6F4E8802FFD30E7DFCF9FF7FFC02FF31" box="[844,899,194,217]" pageId="33" pageNumber="34">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFCF9FF57FA36FDB7" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="B04A36A0FFD30E7DFCF9FF57FB3FFEA8" blockId="33.[844,1466,194,1850]" pageId="33" pageNumber="34">
<materialsCitation id="009D3CFDFFD30E7DFCF9FF57FB3AFEA8" collectingDate="1974-05-03" collectionCode="ZISP" collectorName="G. Medvedev" country="Turkmenistan" latitude="38.521137" location="Syunt Mt." longLatPrecision="1" longitude="56.36947" municipality="Magtymguly" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Balkan velayat" typeStatus="holotype">
<typeStatus id="6F4E8802FFD30E7DFCF9FF57FC33FEE9" box="[844,946,234,257]" pageId="33" pageNumber="34">Holotype</typeStatus>
:
<emphasis id="8281EAB2FFD30E7DFC76FF57FBE3FEE9" bold="true" box="[963,1122,234,257]" pageId="33" pageNumber="34">
<collectingCountry id="C8E27630FFD30E7DFC76FF57FBE3FEE9" box="[963,1122,234,257]" name="Turkmenistan" pageId="33" pageNumber="34" value="Turkmenistan">Turkmenistan</collectingCountry>
</emphasis>
,
<emphasis id="8281EAB2FFD30E7DFBC2FF57FA9DFEE9" box="[1143,1308,234,257]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="7231F842FFD30E7DFBC2FF57FA9DFEE9" box="[1143,1308,234,257]" pageId="33" pageNumber="34">Balkan velayat</collectingRegion>
</emphasis>
,
<collectingMunicipality id="502EACDAFFD30E7DFA84FF57FA3BFEE9" box="[1329,1466,234,257]" pageId="33" pageNumber="34">Magtymguly</collectingMunicipality>
etrap, top of
<location id="B52A607BFFD30E7DFC6AFEB4FBCFFEC8" LSID="urn:lsid:plazi:treatment:385C87B6FFD30E7EFF58F8BFFC8CFDCC:B52A607BFFD30E7DFC6AFEB4FBCFFEC8" box="[991,1102,265,288]" country="Turkmenistan" latitude="38.521137" longLatPrecision="1" longitude="56.36947" municipality="Magtymguly" name="Syunt Mt." pageId="33" pageNumber="34" stateProvince="Balkan velayat">Syunt Mt.</location>
[
<geoCoordinate id="D5C15067FFD30E7DFBEAFEB4FB73FEC8" box="[1119,1266,265,289]" degrees="38" direction="north" minutes="31" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="16.1" value="38.521137">
38°31
<emphasis id="8281EAB2FFD30E7DFB14FEB4FB26FEC9" bold="true" box="[1185,1191,265,289]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
16.1
<emphasis id="8281EAB2FFD30E7DFB60FEB4FB5EFEC9" bold="true" box="[1237,1247,265,289]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="D5C15067FFD30E7DFB48FEB4FA0EFEC8" box="[1277,1423,265,289]" degrees="56" direction="east" minutes="22" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="10.1" value="56.36947">
56°22
<emphasis id="8281EAB2FFD30E7DFA88FEB4FAC2FEC9" bold="true" box="[1341,1347,265,289]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
10.1
<emphasis id="8281EAB2FFD30E7DFAC4FEB4FAFAFEC9" bold="true" box="[1393,1403,265,289]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
E
</geoCoordinate>
],
<date id="C44B1060FFD30E7DFA10FEB4FC16FEA8" pageId="33" pageNumber="34" value="1974-05-03">
<collectingDate id="D40FE988FFD30E7DFA10FEB4FC16FEA8" pageId="33" pageNumber="34" value="1974-05-03">3. V.1974</collectingDate>
</date>
(
<collectorName id="1D005376FFD30E7DFC1CFE94FBBEFEA8" box="[937,1087,297,320]" pageId="33" pageNumber="34">G. Medvedev</collectorName>
), J (
<collectionCode id="D6E4AE65FFD30E7DFBC2FE94FB33FEA8" box="[1143,1202,297,320]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="33" pageNumber="34">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B04A36A0FFD30E7DFCC1FEF4FA36FDB7" blockId="33.[844,1466,194,1850]" pageId="33" pageNumber="34">
<materialsCitation id="009D3CFDFFD30E7DFCC1FEF4FB38FE28" collectingDate="1973-07-13" collectionCode="ZISP" country="Turkmenistan" latitude="38.20486" location="Khozly at Sumbar Riv." longLatPrecision="1" longitude="57.091053" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Balkan velayat, Magtymguly etrap" typeStatus="paratype">
<typeStatus id="6F4E8802FFD30E7DFCC1FEF4FC5FFE88" box="[884,990,329,352]" pageId="33" pageNumber="34">Paratypes</typeStatus>
:
<emphasis id="8281EAB2FFD30E7DFBA6FEF4FB33FE88" bold="true" box="[1043,1202,329,352]" pageId="33" pageNumber="34">
<collectingCountry id="C8E27630FFD30E7DFBA6FEF4FB33FE88" box="[1043,1202,329,352]" name="Turkmenistan" pageId="33" pageNumber="34" value="Turkmenistan">Turkmenistan</collectingCountry>
</emphasis>
:
<collectingRegion id="7231F842FFD30E7DFB5FFEF4FBA0FE68" pageId="33" pageNumber="34">
<emphasis id="8281EAB2FFD30E7DFB5FFEF4FA32FE88" box="[1258,1459,329,352]" italics="true" pageId="33" pageNumber="34">Balkan velayat</emphasis>
, Magtymguly etrap
</collectingRegion>
: “Khazly [?]” [ravine
<location id="B52A607BFFD30E7DFAF1FED4FC56FE48" LSID="urn:lsid:plazi:treatment:385C87B6FFD30E7EFF58F8BFFC8CFDCC:B52A607BFFD30E7DFAF1FED4FC56FE48" country="Turkmenistan" latitude="38.20486" longLatPrecision="1" longitude="57.091053" name="Khozly at Sumbar Riv." pageId="33" pageNumber="34" stateProvince="Balkan velayat, Magtymguly etrap">Khozly at Sumbar Riv.</location>
б
<geoCoordinate id="D5C15067FFD30E7DFC45FE34FB00FE48" box="[1008,1153,392,416]" degrees="38" direction="north" minutes="12" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="17.5" value="38.20486">
38°12
<emphasis id="8281EAB2FFD30E7DFB85FE35FBB7FE48" bold="true" box="[1072,1078,392,416]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
17.5
<emphasis id="8281EAB2FFD30E7DFBD1FE35FBEFFE48" bold="true" box="[1124,1134,392,416]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="D5C15067FFD30E7DFB3BFE34FAA1FE48" box="[1166,1312,392,416]" degrees="57" direction="east" minutes="05" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="27.8" value="57.091053">
57°05
<emphasis id="8281EAB2FFD30E7DFB7BFE35FB55FE48" bold="true" box="[1230,1236,392,416]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
27.8
<emphasis id="8281EAB2FFD30E7DFAB7FE35FA8DFE48" bold="true" box="[1282,1292,392,416]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
E
</geoCoordinate>
],
<date id="C44B1060FFD30E7DFA8DFE34FA38FE48" box="[1336,1465,393,416]" pageId="33" pageNumber="34" value="1973-07-13">
<collectingDate id="D40FE988FFD30E7DFA8DFE34FA38FE48" box="[1336,1465,393,416]" pageId="33" pageNumber="34" value="1973-07-13">13.VII.1973</collectingDate>
</date>
[collector unknown], 1J (
<collectionCode id="D6E4AE65FFD30E7DFBC3FE14FB30FE28" box="[1142,1201,425,448]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="33" pageNumber="34">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD30E7DFB7EFE14FB6CFE08" collectingDate="1974-05-03" collectionCode="ZISP" collectorName="G. Medvedev" country="Turkmenistan" location="Syunt Mt." municipality="Syunt Mt." pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Balkan Velayat, Magtymguly Etrap">
top of
<collectingMunicipality id="502EACDAFFD30E7DFAAAFE14FA12FE28" box="[1311,1427,425,448]" pageId="33" pageNumber="34">Syunt Mt.</collectingMunicipality>
,
<date id="C44B1060FFD30E7DFA10FE14FC16FE08" pageId="33" pageNumber="34" value="1974-05-03">
<collectingDate id="D40FE988FFD30E7DFA10FE14FC16FE08" pageId="33" pageNumber="34" value="1974-05-03">3. V.1974</collectingDate>
</date>
(
<collectorName id="1D005376FFD30E7DFC07FE74FBD3FE08" box="[946,1106,457,480]" pageId="33" pageNumber="34">G. Medvedev</collectorName>
),
<specimenCount id="A6F3FD29FFD30E7DFBDAFE74FB11FE08" box="[1135,1168,456,480]" count="1" pageId="33" pageNumber="34" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD30E7DFB1FFE74FB64FE08" box="[1194,1253,457,480]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="33" pageNumber="34">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD30E7DFAB6FE74FBF1FDF7" collectingDate="1974-07-28" collectionCode="ZMUM" country="Turkmenistan" location="Garrygala" municipality="Kopet Dag Mt. Range" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Balkan Velayat, Magtymguly Etrap">
<collectingMunicipality id="502EACDAFFD30E7DFAB6FE74FC13FE17" pageId="33" pageNumber="34">Kopet Dag Mt. Range</collectingMunicipality>
, “Kara-kala” [
<location id="B52A607BFFD30E7DFBE3FE55FB45FE17" LSID="urn:lsid:plazi:treatment:385C87B6FFD30E7EFF58F8BFFC8CFDCC:B52A607BFFD30E7DFBE3FE55FB45FE17" box="[1110,1220,488,511]" country="Turkmenistan" municipality="Kopet Dag Mt. Range" name="Garrygala" pageId="33" pageNumber="34" stateProvince="Balkan Velayat, Magtymguly Etrap">Garrygala</location>
] env.,
<date id="C44B1060FFD30E7DFA8DFE55FA3BFE17" box="[1336,1466,488,511]" pageId="33" pageNumber="34" value="1974-07-28">
<collectingDate id="D40FE988FFD30E7DFA8DFE55FA3BFE17" box="[1336,1466,488,511]" pageId="33" pageNumber="34" value="1974-07-28">28.VII.1974</collectingDate>
</date>
(Aksent’ ev),
<specimenCount id="A6F3FD29FFD30E7DFC69FDB5FC7FFDC8" box="[988,1022,520,544]" count="1" pageId="33" pageNumber="34" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD30E7DFBA4FDB5FBE8FDF7" box="[1041,1129,520,543]" collectionName="Russia, Moscow, Moscow State University" pageId="33" pageNumber="34">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD30E7DFB34FDB5FA35FDB7" collectingDate="1973-07-19" collectionCode="ZISP" country="Turkmenistan" latitude="38.338917" location="Ipay-Kala" longLatPrecision="1" longitude="57.136086" municipality="Baharly" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Ahal velayat">
<emphasis id="8281EAB2FFD30E7DFB34FDB5FA8AFDF7" box="[1153,1291,520,543]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="7231F842FFD30E7DFB34FDB5FA8AFDF7" box="[1153,1291,520,543]" pageId="33" pageNumber="34">Ahal velayat</collectingRegion>
</emphasis>
,
<collectingMunicipality id="502EACDAFFD30E7DFAABFDB5FAF0FDF7" box="[1310,1393,520,543]" pageId="33" pageNumber="34">Baharly</collectingMunicipality>
etrap, gorge urochishche
<location id="B52A607BFFD30E7DFBF9FD95FB34FDD7" LSID="urn:lsid:plazi:treatment:385C87B6FFD30E7EFF58F8BFFC8CFDCC:B52A607BFFD30E7DFBF9FD95FB34FDD7" box="[1100,1205,552,575]" country="Turkmenistan" latitude="38.338917" longLatPrecision="1" longitude="57.136086" municipality="Baharly" name="Ipay-Kala" pageId="33" pageNumber="34" stateProvince="Ahal velayat">Ipay-Kala</location>
[
<geoCoordinate id="D5C15067FFD30E7DFB6BFD95FAF0FDD7" box="[1246,1393,552,576]" degrees="38" direction="north" minutes="20" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="20.1" value="38.338917">
38°20
<emphasis id="8281EAB2FFD30E7DFA95FD95FAA7FDA8" bold="true" box="[1312,1318,552,576]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
20.1
<emphasis id="8281EAB2FFD30E7DFAE0FD95FADEFDA8" bold="true" box="[1365,1375,552,576]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="D5C15067FFD30E7DFA21FD95FC39FDB7" degrees="57" direction="east" minutes="08" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="09.9" value="57.136086">
57° 08
<emphasis id="8281EAB2FFD30E7DFCD3FDFAFCEDFDB7" bold="true" box="[870,876,583,607]" italics="true" pageId="33" pageNumber="34">ʹ</emphasis>
09.9
<emphasis id="8281EAB2FFD30E7DFC2EFDFAFC24FDB7" bold="true" box="[923,933,583,607]" italics="true" pageId="33" pageNumber="34">ʺ</emphasis>
E
</geoCoordinate>
],
<date id="C44B1060FFD30E7DFC79FDF5FBCFFDB7" box="[972,1102,584,607]" pageId="33" pageNumber="34" value="1973-07-19">
<collectingDate id="D40FE988FFD30E7DFC79FDF5FBCFFDB7" box="[972,1102,584,607]" pageId="33" pageNumber="34" value="1973-07-19">19.VII.1973</collectingDate>
</date>
(collector unknown), 1J (
<collectionCode id="D6E4AE65FFD30E7DFAC4FDF5FA2DFDB7" box="[1393,1452,584,607]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="33" pageNumber="34">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFCF9FD08FA2FFB7A" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="B04A36A0FFD30E7DFCF9FD08FC38FD24" blockId="33.[844,1466,194,1850]" box="[844,953,693,716]" pageId="33" pageNumber="34">
<heading id="EB0281CCFFD30E7DFCF9FD08FC38FD24" bold="true" box="[844,953,693,716]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFCF9FD08FC38FD24" bold="true" box="[844,953,693,716]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD30E7DFCF9FD60FA2FFB7A" blockId="33.[844,1466,194,1850]" pageId="33" pageNumber="34">
Ventral lobe of conjunctiva in ventral view slightly widening distally, its apex evenly rounded, with small notch in middle of apical margin; ventral wall of lobe in apical half with narrow desclerotised groove (
<figureCitation id="28CE2A25FFD30E7DFB75FC81FAADFCBB" box="[1216,1324,828,851]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="33" pageNumber="34">
Figure 6
<collectionCode id="D6E4AE65FFD30E7DFAAFFC81FAADFCBB" box="[1306,1324,828,851]" pageId="33" pageNumber="34">R</collectionCode>
</figureCitation>
); dorsal wall of lobe entirely membranous, with barely noticeable carina and very small impressions at its sides (
<figureCitation id="28CE2A25FFD30E7DFA8DFCC1FA2BFC7B" box="[1336,1450,892,915]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="33" pageNumber="34">
Figure 6
<collectionCode id="D6E4AE65FFD30E7DFA29FCC1FA2BFC7B" box="[1436,1450,892,915]" pageId="33" pageNumber="34">S</collectionCode>
</figureCitation>
). Spicule at conjunctival dorsolateral lobes (
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Figure 7
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) rather widely curved at base, long, protruding beyond apex of denticle on spicule basal plate slightly less than half of length; beyond middle, spicule curved strongly and sharply; extreme apex of spicule not curved, obliquely truncated. Denticle on anterior end of spicule basal plate longer than in nominotypical subspecies, more or less spear-shaped at apex, directed dorsolaterad and anteriad.
</paragraph>
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<subSubSection id="F8EF652BFFD30E7DFCF9FB55FAACFA6F" pageId="33" pageNumber="34" type="description">
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<heading id="EB0281CCFFD30E7DFCF9FB55FC70FB17" bold="true" box="[844,1009,1256,1279]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="6">
<emphasis id="8281EAB2FFD30E7DFCF9FB55FC70FB17" bold="true" box="[844,1009,1256,1279]" pageId="33" pageNumber="34">Measurements</emphasis>
</heading>
</paragraph>
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Body
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length,
<quantity id="770D9B45FFD30E7DFBC6FAADFB7AFACF" box="[1139,1275,1296,1320]" metricMagnitude="-1" metricUnit="m" metricValue="1.0921999999999998" metricValueMax="1.2319" metricValueMin="0.9525" pageId="33" pageNumber="34" unit="in" value="4.3" valueMax="4.85" valueMin="3.75">3.75–4.85 in</quantity>
width; pronotum
<quantity id="770D9B45FFD30E7DFCF9FA8DFC50FAAF" box="[844,977,1328,1352]" metricMagnitude="-2" metricUnit="m" metricValue="9.652" metricValueMax="10.795" metricValueMin="8.509" pageId="33" pageNumber="34" unit="in" value="3.8" valueMax="4.25" valueMin="3.35">3.35–4.25 in</quantity>
width; head
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width; ocular index 3.41–3.68; length of antennal segments I–V: 0.32–0.39: 0.50–0.63: 0.57–0.70: 0.78–0.92: 0.87–0.98.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD30E7DFCF9FA60FC7DF8D2" pageId="33" pageNumber="34" type="discussion">
<paragraph id="B04A36A0FFD30E7DFCF9FA60FC32FA1C" blockId="33.[844,1466,194,1850]" box="[844,947,1501,1524]" pageId="33" pageNumber="34">
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<emphasis id="8281EAB2FFD30E7DFCF9FA60FC32FA1C" bold="true" box="[844,947,1501,1524]" pageId="33" pageNumber="34">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD30E7DFCF9F9B8FC7DF8D2" blockId="33.[844,1466,194,1850]" pageId="33" pageNumber="34">
<bibRefCitation id="D4644B51FFD30E7DFCF9F9B8FB85F9F4" author="Asanova RB" box="[844,1028,1541,1564]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="33" pageNumber="34" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova (1964)</bibRefCitation>
distinguished one Turkmenian male from the other specimens of
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<emphasis id="8281EAB2FFD30E7DFB17F998FAA1F9D4" box="[1186,1312,1573,1596]" italics="true" pageId="33" pageNumber="34">C. wagneri</emphasis>
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examined in her work by the shape of “the hook of penal appendage”, but she did not decide on an attribution of a taxonomic status to this “deviation” for want of material. The specimen examined by her (“Kara-Kala, Kopet-Dag,
<date id="C44B1060FFD30E7DFA2DF919FC18F933" pageId="33" pageNumber="34" value="1935-05-26">26. V.1935</date>
(Ushatinskaya), 1J”) is currently absent in the
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collection, but its spicule figured in Asanova’ s paper completely corresponds to the diagnosis of my new subspecies.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD00E7EFF2BFF7CFEDFFE48" pageId="34" pageNumber="35" type="etymology">
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<emphasis id="8281EAB2FFD00E7EFF2BFF7CFE99FF30" bold="true" box="[158,280,193,216]" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD00E7EFF2BFF54FEDFFE48" blockId="34.[158,781,193,1939]" pageId="34" pageNumber="35">
The subspecific epithet is derived from the Ancient Greek Ὑρκανία, the name of a satrapy in the Achaemenid Empire at the south-east coast of the Caspian Sea in the territory of present-day Northern Iran and the extreme south-west of Turkmenistan that encloses the
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locality of the subspecies.
</paragraph>
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<emphasis id="8281EAB2FFD00E7EFF2BFE58FEA6FE14" bold="true" box="[158,295,485,508]" pageId="34" pageNumber="35">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD00E7EFF2BFDB0FC8CFDCC" blockId="34.[158,781,193,1939]" box="[158,781,525,548]" pageId="34" pageNumber="35">Figure 15. South-Western Turkmenistan (Western Kopet Dag).</paragraph>
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</treatment>
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<emphasis id="8281EAB2FFD40E7AFF53FEFCFDBCFEB0" bold="true" box="[230,573,321,344]" italics="true" pageId="38" pageNumber="39">Canthophorus niveimarginatus</emphasis>
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(
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Figures 3
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,
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5
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,
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, 6
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,
<figureCitation id="28CE2A25FFD40E7AFE5BFEDCFDBAFE90" box="[494,571,353,376]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="38" pageNumber="39">
7
<collectionCode id="D6E4AE65FFD40E7AFE4EFEDCFD8DFE90" box="[507,524,353,376]" pageId="38" pageNumber="39">T</collectionCode>
, 7
<collectionCode id="D6E4AE65FFD40E7AFD9DFEDCFDBAFE90" box="[552,571,353,376]" pageId="38" pageNumber="39">U</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD40E7AFDFEFEDCFD1CFE90" box="[587,669,353,376]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="38" pageNumber="39">
<date id="C44B1060FFD40E7AFDFEFEDCFDE9FE90" box="[587,616,353,376]" pageId="38" pageNumber="39">
8
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</date>
, 8
<collectionCode id="D6E4AE65FFD40E7AFD36FEDCFD1CFE90" box="[643,669,353,376]" pageId="38" pageNumber="39">W</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD40E7AFD19FEDCFD59FE90" box="[684,728,353,376]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="38" pageNumber="39">
11
<collectionCode id="D6E4AE65FFD40E7AFD70FEDCFD59FE90" box="[709,728,353,376]" pageId="38" pageNumber="39">D</collectionCode>
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)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7AFF2BFE37FE89FE52" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="B04A36A0FFD40E7AFF2BFE37FE89FE52" blockId="38.[158,781,193,1938]" pageId="38" pageNumber="39">
<materialsCitation id="009D3CFDFFD40E7AFF2BFE37FE85FE52" country="Japan" location="Japan" pageId="38" pageNumber="39" specimenCount="1" typeStatus="syntype">
<taxonomicName id="77F54D23FFD40E7AFF2BFE37FDF0FE77" authority="Scott 1874: 289" authorityName="Scott" authorityPageNumber="289" authorityYear="1874" box="[158,625,394,415]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="niveimarginatus">
<emphasis id="8281EAB2FFD40E7AFF2BFE37FE48FE77" box="[158,457,394,415]" italics="true" pageId="38" pageNumber="39">Canthophorus niveimarginatus</emphasis>
<bibRefCitation id="D4644B51FFD40E7AFE67FE37FDF0FE77" author="Scott J." box="[466,625,394,415]" journalOrPublisher="Annals and Magazine of Natural History." pageId="38" pageNumber="39" pagination="289 - 304" part="14" refId="ref40679" refString="Scott J. 1874. On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species. Annals and Magazine of Natural History. 14 (ser. 4): 289 - 304; 360 - 365; 426 - 452." title="On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species" type="journal article" year="1874">Scott 1874: 289</bibRefCitation>
</taxonomicName>
, 295 [
<typeStatus id="6F4E8802FFD40E7AFD05FE37FC88FE77" box="[688,777,394,415]" pageId="38" pageNumber="39">syntypes</typeStatus>
:
<collectingCountry id="C8E27630FFD40E7AFF76FE18FF7FFE52" box="[195,254,421,442]" name="Japan" pageId="38" pageNumber="39" value="Japan">Japan</collectingCountry>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7AFF2BFDBFFD72FA9B" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="B04A36A0FFD40E7AFF2BFDBFFEF4FDF1" blockId="38.[158,781,193,1938]" box="[158,373,514,537]" pageId="38" pageNumber="39">
<heading id="EB0281CCFFD40E7AFF2BFDBFFEF4FDF1" bold="true" box="[158,373,514,537]" fontSize="10" level="2" pageId="38" pageNumber="39" reason="6">
<emphasis id="8281EAB2FFD40E7AFF2BFDBFFEF4FDF1" bold="true" box="[158,373,514,537]" pageId="38" pageNumber="39">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD40E7AFF2BFD9BFD72FA9B" blockId="38.[158,781,193,1938]" pageId="38" pageNumber="39">
<materialsCitation id="009D3CFDFFD40E7AFF2BFD9BFF63FD98" collectingDate="1979-06" collectionCode="ZISP" country="Russia" location="Usynskiy maral sovkhoz" municipality="Ermakovskiy Distr" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Krasnoyarsk Terr">
<emphasis id="8281EAB2FFD40E7AFF2BFD9BFF63FDD3" bold="true" box="[158,226,550,571]" pageId="38" pageNumber="39">
<collectingCountry id="C8E27630FFD40E7AFF2BFD9BFF63FDD3" box="[158,226,550,571]" name="Russia" pageId="38" pageNumber="39" value="Russia">Russia</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD40E7AFF4EFD9BFE30FDD3" box="[251,433,550,571]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="7231F842FFD40E7AFF4EFD9BFE30FDD3" box="[251,433,550,571]" pageId="38" pageNumber="39">Krasnoyarsk Terr</collectingRegion>
</emphasis>
.:
<collectingMunicipality id="502EACDAFFD40E7AFE65FD9BFD16FDD3" box="[464,663,550,571]" pageId="38" pageNumber="39">Ermakovskiy Distr</collectingMunicipality>
.,
<location id="B52A607BFFD40E7AFD05FD9BFEC4FDBE" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFD05FD9BFEC4FDBE" country="Russia" municipality="Ermakovskiy Distr" name="Usynskiy maral sovkhoz" pageId="38" pageNumber="39" stateProvince="Krasnoyarsk Terr">Usynskiy maral sovkhoz</location>
, 22,23.
<date id="C44B1060FFD40E7AFE11FDFCFE75FDBE" box="[420,500,577,598]" pageId="38" pageNumber="39" value="1979-06">VI.1979</date>
(Korotyaev), 1J,
<specimenCount id="A6F3FD29FFD40E7AFD5BFDFCFC8DFDBE" box="[750,780,576,598]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFF10FDE6FF5AFD98" box="[165,219,603,624]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF4FFDE6FE13FD63" collectingDate="1964-06-23" collectionCode="ZISP" collectorName="M. Kozlov" country="Russia" location="Kosh-Agach Settlm" municipality="Altay Republic" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFF4FFDE6FE17FD98" box="[250,406,603,624]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFF4FFDE6FE17FD98" box="[250,406,603,624]" pageId="38" pageNumber="39">Altay Republic</collectingMunicipality>
</emphasis>
:
<location id="B52A607BFFD40E7AFE04FDE6FD02FD98" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFE04FDE6FD02FD98" box="[433,643,603,624]" country="Russia" municipality="Altay Republic" name="Kosh-Agach Settlm" pageId="38" pageNumber="39">Kosh-Agach Settlm</location>
.,
<date id="C44B1060FFD40E7AFD28FDE6FC8AFD98" box="[669,779,603,624]" pageId="38" pageNumber="39" value="1964-06-23">23.VI.1964</date>
(
<collectorName id="1D005376FFD40E7AFF13FDCBFE94FD63" box="[166,277,630,651]" pageId="38" pageNumber="39">M. Kozlov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFE9DFDCBFEC7FD63" box="[296,326,629,651]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFEE0FDCBFE0AFD63" box="[341,395,630,651]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFE2AFDCBFF63FD28" collectingDate="1949-07-23" collectionCode="ZISP" country="Russia" location="Boyarovka Settlm" municipality="Republic of Tyva" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFE2AFDCBFDC4FD63" box="[415,581,630,651]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFE2AFDCBFDC4FD63" box="[415,581,630,651]" pageId="38" pageNumber="39">Republic of Tyva</collectingMunicipality>
</emphasis>
:
<location id="B52A607BFFD40E7AFDE1FDCBFC88FD63" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFDE1FDCBFC88FD63" box="[596,777,630,651]" country="Russia" municipality="Republic of Tyva" name="Boyarovka Settlm" pageId="38" pageNumber="39">Boyarovka Settlm</location>
.
<locationDeviation id="A1F82F82FFD40E7AFF2BFD2DFE06FD4D" box="[158,391,656,677]" pageId="38" pageNumber="39">nr Kaa-khem Settlm.</locationDeviation>
,
<date id="C44B1060FFD40E7AFE2BFD2DFD94FD4D" box="[414,533,656,677]" pageId="38" pageNumber="39" value="1949-07-23">23.VII.1949</date>
(Perevozchikova),
<specimenCount id="A6F3FD29FFD40E7AFD5BFD2DFC8DFD4E" box="[750,780,656,678]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFF10FD16FF5AFD28" box="[165,219,683,704]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF47FD16FC89FD28" box="[242,776,682,704]" collectingDate="1949-08-02" collectionCode="ZISP" country="Russia" location="Kyzyl" pageId="38" pageNumber="39" specimenCount="1">
<location id="B52A607BFFD40E7AFF47FD16FEAEFD28" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF47FD16FEAEFD28" box="[242,303,683,704]" country="Russia" name="Kyzyl" pageId="38" pageNumber="39">Kyzyl</location>
,
<date id="C44B1060FFD40E7AFE8AFD16FE33FD28" box="[319,434,683,704]" pageId="38" pageNumber="39" value="1949-08-02">2.VIII.1949</date>
[collector unknown], 1J (
<collectionCode id="D6E4AE65FFD40E7AFD7EFD16FC80FD28" box="[715,769,683,704]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF2BFD78FE06FD1D" collectingDate="1979-07-13" collectionCode="ZISP" collectorName="B. A. Korotyaev" country="Russia" location="Yenisey" municipality="Kyzyl" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="502EACDAFFD40E7AFF2BFD78FF5DFD32" box="[158,220,709,730]" pageId="38" pageNumber="39">Kyzyl</collectingMunicipality>
, floodplain of
<location id="B52A607BFFD40E7AFE35FD78FE4FFD32" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFE35FD78FE4FFD32" box="[384,462,709,730]" country="Russia" municipality="Kyzyl" name="Yenisey" pageId="38" pageNumber="39">Yenisey</location>
, left bank,
<date id="C44B1060FFD40E7AFDE4FD78FD49FD32" box="[593,712,709,730]" pageId="38" pageNumber="39" value="1979-07-13">13.VII.1979</date>
(
<collectorName id="1D005376FFD40E7AFD6BFD78FE89FD1D" pageId="38" pageNumber="39">B.A. Korotyaev</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFEA9FD5DFEBBFD1D" box="[284,314,735,757]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFEFFFD5DFE01FD1D" box="[330,384,736,757]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFE20FD5DFE07FCC2" collectingDate="1979-07-20" collectionCode="ZISP" collectorName="B. A. Korotyaev" country="Russia" location="Kadyy Riv." municipality="Ovyurskiy Distr" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="502EACDAFFD40E7AFE20FD5DFDB9FD1D" box="[405,568,736,757]" pageId="38" pageNumber="39">Ovyurskiy Distr</collectingMunicipality>
.,
<locationDeviation id="A1F82F82FFD40E7AFDFCFD5DFF66FCE7" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFDFCFD5DFD09FD1D" box="[585,648,736,757]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="39" unit="km" value="20.0">20 km</quantity>
E Torchalyg Settlm.
</locationDeviation>
,
<location id="B52A607BFFD40E7AFF47FD47FEE6FCE7" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF47FD47FEE6FCE7" box="[242,359,762,783]" country="Russia" municipality="Ovyurskiy Distr" name="Kadyy Riv." pageId="38" pageNumber="39">Kadyy Riv.</location>
, steppe and floodplain,
<date id="C44B1060FFD40E7AFDE2FD47FD4CFCE7" box="[599,717,762,783]" pageId="38" pageNumber="39" value="1979-07-20">20.VII.1979</date>
(
<collectorName id="1D005376FFD40E7AFD6BFD47FE89FCC2" pageId="38" pageNumber="39">B.A. Korotyaev</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFEAEFCA8FEB8FCC3" box="[283,313,789,811]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFEFCFCA8FEFEFCC2" box="[329,383,789,810]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFE26FCA8FE1BFCB7" collectingDate="1989-06-10" collectionCode="ZISP" collectorName="O. N. Kabakov" country="Russia" elevation="1000" location="Sherlovaya Mt." municipality="Borzinskiy Distr" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Zabaykal' skiy Terr.">
<collectingRegion id="7231F842FFD40E7AFE26FCA8FDCFFCC2" box="[403,590,789,811]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFE26FCA8FDC9FCC2" box="[403,584,789,811]" italics="true" pageId="38" pageNumber="39">Zabaykal’ skiy Terr</emphasis>
.
</collectingRegion>
:
<collectingMunicipality id="502EACDAFFD40E7AFDE8FCA8FC87FCC2" box="[605,774,789,810]" pageId="38" pageNumber="39">Borzinskiy Distr</collectingMunicipality>
.:
<location id="B52A607BFFD40E7AFF2BFC8DFEB8FCAD" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF2BFC8DFEB8FCAD" box="[158,313,816,837]" country="Russia" municipality="Borzinskiy Distr" name="Sherlovaya Mt." pageId="38" pageNumber="39" stateProvince="Zabaykal' skiy Terr.">Sherlovaya Mt.</location>
,
<emphasis id="8281EAB2FFD40E7AFEF3FC92FED1FCAD" box="[326,336,815,837]" italics="true" pageId="38" pageNumber="39">с</emphasis>
.
<quantity id="770D9B45FFD40E7AFEE6FC8DFE23FCAD" box="[339,418,816,837]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">
<elevation id="3BD8D193FFD40E7AFEE6FC8DFE23FCAD" box="[339,418,816,837]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">1000 m</elevation>
</quantity>
amsl, forest-steppe,
<date id="C44B1060FFD40E7AFDC0FC8DFD62FCAD" box="[629,739,816,837]" pageId="38" pageNumber="39" value="1989-06-10">10.VI.1989</date>
(
<collectorName id="1D005376FFD40E7AFD41FC8DFE9BFCB7" pageId="38" pageNumber="39">O. N. Kabakov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFE9AFCF7FECCFC88" box="[303,333,842,864]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFEE8FCF7FE12FCB7" box="[349,403,842,863]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFE1DFCF7FC89FC92" collectingDate="1989-07-14" collectionCode="ZISP" collectorName="O. N. Kabakov" country="Russia" elevation="1050" location="Tsagan-Oluy Settlm" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<location id="B52A607BFFD40E7AFE1DFCF7FDF3FCB7" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFE1DFCF7FDF3FCB7" box="[424,626,842,863]" country="Russia" name="Tsagan-Oluy Settlm" pageId="38" pageNumber="39">Tsagan-Oluy Settlm</location>
.,
<quantity id="770D9B45FFD40E7AFD36FCF7FC8DFCB7" box="[643,780,842,864]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="38" pageNumber="39" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">
<elevation id="3BD8D193FFD40E7AFD36FCF7FC8DFCB7" box="[643,780,842,864]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="38" pageNumber="39" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</elevation>
</quantity>
amsl, forest-steppe,
<date id="C44B1060FFD40E7AFEDCFCD8FE61FC92" box="[361,480,869,890]" pageId="38" pageNumber="39" value="1989-07-14">14.VII.1989</date>
(
<collectorName id="1D005376FFD40E7AFE47FCD8FD06FC92" box="[498,647,869,890]" pageId="38" pageNumber="39">O.N. Kabakov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD29FCD8FD3BFC92" box="[668,698,868,890]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFD7EFCD8FC80FC92" box="[715,769,869,890]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF2BFCC2FC88FC47" collectingDate="1925-05-29" collectingDateMax="1927-07-10" collectingDateMin="1925-05-29" collectionCode="ZISP" country="Russia" location="Khanka Lake" municipality="Kamen' - Rybolov" pageId="38" pageNumber="39" specimenCount="4" specimenCount-female="4" stateProvince="Primorskiy Terr">
<emphasis id="8281EAB2FFD40E7AFF2BFCC2FEB9FC7C" box="[158,312,895,916]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="7231F842FFD40E7AFF2BFCC2FEB9FC7C" box="[158,312,895,916]" pageId="38" pageNumber="39">Primorskiy Terr</collectingRegion>
</emphasis>
.:
<collectingMunicipality id="502EACDAFFD40E7AFEFFFCC2FE75FC7C" box="[330,500,895,917]" pageId="38" pageNumber="39">Kamen’-Rybolov</collectingMunicipality>
,
<location id="B52A607BFFD40E7AFE4BFCC2FD03FC7C" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFE4BFCC2FD03FC7C" box="[510,642,895,916]" country="Russia" municipality="Kamen' - Rybolov" name="Khanka Lake" pageId="38" pageNumber="39" stateProvince="Primorskiy Terr">Khanka Lake</location>
, 24,
<date id="C44B1060FFD40E7AFD1FFCC2FF35FC47" pageId="38" pageNumber="39" value="1925-05-29" valueMax="1925-05-30" valueMin="1925-05-29">25,29–30. V.</date>
1908 (
<location id="B52A607BFFD40E7AFF47FC27FECFFC47" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF47FC27FECFFC47" box="[242,334,922,943]" country="Russia" municipality="Kamen' - Rybolov" name="Cherskiy" pageId="38" pageNumber="39" stateProvince="Primorskiy Terr">Cherskiy</location>
),
<specimenCount id="A6F3FD29FFD40E7AFED7FC27FE01FC47" box="[354,384,921,943]" count="3" pageId="38" pageNumber="39" type="female">3♀</specimenCount>
,
<date id="C44B1060FFD40E7AFE3AFC27FD84FC47" box="[399,517,922,943]" pageId="38" pageNumber="39" value="1927-07-10">10.VII.1927</date>
(
<location id="B52A607BFFD40E7AFDA1FC27FD0AFC47" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFDA1FC27FD0AFC47" box="[532,651,922,943]" country="Russia" municipality="Kamen' - Rybolov" name="Stackelberg" pageId="38" pageNumber="39" stateProvince="Primorskiy Terr">Stackelberg</location>
),
<specimenCount id="A6F3FD29FFD40E7AFD2BFC27FD3DFC47" box="[670,700,921,943]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFD7EFC27FC80FC47" box="[715,769,922,943]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD40E7AFF2BFC09FC89FC21" box="[158,776,948,970]" collectionCode="ZISP" collectorName="I. M. Kerzhner" country="Mongolia" location="Mongolia" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFF2BFC09FE81FC21" bold="true" box="[158,256,948,969]" pageId="38" pageNumber="39">
<collectingCountry id="C8E27630FFD40E7AFF2BFC09FE81FC21" box="[158,256,948,969]" name="Mongolia" pageId="38" pageNumber="39" value="Mongolia">Mongolia</collectingCountry>
</emphasis>
: [without locality] 1976 (
<collectorName id="1D005376FFD40E7AFE4AFC09FD0CFC21" box="[511,653,948,969]" pageId="38" pageNumber="39">I.M. Kerzhner</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD2AFC09FD3CFC22" box="[671,701,948,970]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFD7EFC09FC80FC21" box="[715,769,948,969]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF2BFC72FF63FC17" collectingDate="1968-08-30" collectionCode="ZISP" country="Mongolia" location="Ulaangom" municipality="Uvs Prov" pageId="38" pageNumber="39" specimenCount="1">
<emphasis id="8281EAB2FFD40E7AFF2BFC72FF7FFC0C" box="[158,254,975,996]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFF2BFC72FF7FFC0C" box="[158,254,975,996]" pageId="38" pageNumber="39">Uvs Prov</collectingMunicipality>
</emphasis>
:
<locationDeviation id="A1F82F82FFD40E7AFEA7FC72FEF1FC0C" bearing="180.0" box="[274,368,975,996]" distance="15000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFEA7FC72FED7FC0C" box="[274,342,975,996]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="km" value="15.0">15 km</quantity>
S
</locationDeviation>
<location id="B52A607BFFD40E7AFEC8FC72FE65FC0C" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFEC8FC72FE65FC0C" box="[381,484,975,996]" country="Mongolia" municipality="Uvs Prov" name="Ulaangom" pageId="38" pageNumber="39">Ulaangom</location>
,
<date id="C44B1060FFD40E7AFE41FC72FDF2FC0C" box="[500,627,975,996]" pageId="38" pageNumber="39" value="1968-08-30">30.VIII.1968</date>
(Arnoldi), 1J (
<collectionCode id="D6E4AE65FFD40E7AFF10FC57FF5AFC17" box="[165,219,1002,1023]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF5AFC57FE09FBF1" collectingDate="1975-07-22" collectionCode="ZISP" collectorName="E. L. Gurjeva" country="Mongolia" location="Shine-Ider" municipality="Khovsgol Prov" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFF5AFC57FE00FC17" box="[239,385,1002,1023]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFF5AFC57FE00FC17" box="[239,385,1002,1023]" pageId="38" pageNumber="39">Khövsgöl Prov</collectingMunicipality>
</emphasis>
.,
<locationDeviation id="A1F82F82FFD40E7AFE22FC57FE70FC17" bearing="0.0" box="[407,497,1002,1023]" distance="17000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFE22FC57FE57FC17" box="[407,470,1002,1023]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="38" pageNumber="39" unit="km" value="17.0">17 km</quantity>
N
</locationDeviation>
<location id="B52A607BFFD40E7AFE4EFC57FDE2FC17" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFE4EFC57FDE2FC17" box="[507,611,1002,1023]" country="Mongolia" municipality="Khovsgol Prov" name="Shine-Ider" pageId="38" pageNumber="39">Shine-Ider</location>
,
<date id="C44B1060FFD40E7AFDDAFC57FD67FC17" box="[623,742,1002,1023]" pageId="38" pageNumber="39" value="1975-07-22">22.VII.1975</date>
(
<collectorName id="1D005376FFD40E7AFD42FC57FE8DFBF1" pageId="38" pageNumber="39">E. L. Gurjeva</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFEABFBB9FEBDFBF2" box="[286,316,1028,1050]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFEFFFBB9FE01FBF1" box="[330,384,1028,1049]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFE26FBB9FDB9FBDC" collectingDate="1975-07-23" collectionCode="ZISP" collectorName="E. P. Nartshuk; M. A. Kozlov" country="Mongolia" location="Muren" pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="2">
<locationDeviation id="A1F82F82FFD40E7AFE26FBB9FD8BFBF1" bearing="202.5" box="[403,522,1028,1049]" distance="25000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFE26FBB9FE50FBF1" box="[403,465,1028,1049]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="km" value="25.0">25 km</quantity>
SSW
</locationDeviation>
<location id="B52A607BFFD40E7AFDA7FBB9FDD4FBF1" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFDA7FBB9FDD4FBF1" box="[530,597,1028,1049]" country="Mongolia" name="Muren" pageId="38" pageNumber="39">Muren</location>
,
<date id="C44B1060FFD40E7AFDD5FBB9FD56FBF1" box="[608,727,1028,1049]" pageId="38" pageNumber="39" value="1975-07-23">23.VII.1975</date>
(
<collectorName id="1D005376FFD40E7AFD53FBB9FE0CFBDC" pageId="38" pageNumber="39">E.P. Nartshuk; M.A. Kozlov</collectorName>
), 1J,
<specimenCount id="A6F3FD29FFD40E7AFE78FBA2FE6AFBDC" box="[461,491,1054,1076]" count="2" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFE4EFBA2FDB0FBDC" box="[507,561,1055,1076]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFDF1FBA2FDCCFBA6" collectingDate="1983-07-26" collectionCode="ZMUM" collectorName="I. Ovchinnikov" country="Mongolia" location="Khankh Settlm" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<location id="B52A607BFFD40E7AFDF1FBA2FD5EFBDC" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFDF1FBA2FD5EFBDC" box="[580,735,1055,1076]" country="Mongolia" name="Khankh Settlm" pageId="38" pageNumber="39">Khankh Settlm</location>
.,
<date id="C44B1060FFD40E7AFD5BFBA2FF77FBA6" pageId="38" pageNumber="39" value="1983-07-26">26. VII.1983</date>
(
<collectorName id="1D005376FFD40E7AFEB8FB84FE2AFBA6" box="[269,427,1081,1102]" pageId="38" pageNumber="39">I. Ovchinnikov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFE76FB84FE60FBA7" box="[451,481,1081,1103]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFE43FB84FDC7FBA6" box="[502,582,1081,1102]" collectionName="Russia, Moscow, Moscow State University" pageId="38" pageNumber="39">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFDEBFB84FC89FB81" collectingDate="1967-07-29" collectionCode="ZISP" country="Mongolia" location="Chulut" municipality="Arkhangai Prov" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFDEBFB84FD7EFBA6" box="[606,767,1081,1102]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFDEBFB84FD7EFBA6" box="[606,767,1081,1102]" pageId="38" pageNumber="39">Arkhangai Prov</collectingMunicipality>
</emphasis>
.:
<locationDeviation id="A1F82F82FFD40E7AFF2BFBE9FF79FB81" bearing="90.0" box="[158,248,1108,1129]" distance="35000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFF2BFBE9FF61FB81" box="[158,224,1108,1129]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="39" unit="km" value="35.0">35 km</quantity>
E
</locationDeviation>
somon
<location id="B52A607BFFD40E7AFEFBFBE9FE13FB81" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFEFBFBE9FE13FB81" box="[334,402,1108,1129]" country="Mongolia" municipality="Arkhangai Prov" name="Chulut" pageId="38" pageNumber="39">Chulut</location>
,
<date id="C44B1060FFD40E7AFE15FBE9FD97FB81" box="[416,534,1108,1129]" pageId="38" pageNumber="39" value="1967-07-29">29.VII.1967</date>
(Kerzhner),
<specimenCount id="A6F3FD29FFD40E7AFD2EFBE9FD3BFB81" box="[667,698,1107,1129]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFD7EFBE9FC80FB81" box="[715,769,1108,1129]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF2BFBD3FF63FB76" collectingDate="1975-06-28" collectionCode="ZISP" collectorName="A. F. Emeljanov" country="Mongolia" location="Stariy Tsakhir" pageId="38" pageNumber="39" specimenCount="1">
<locationDeviation id="A1F82F82FFD40E7AFF2BFBD3FE8EFB6B" bearing="225.0" box="[158,271,1134,1155]" distance="12000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFF2BFBD3FF61FB6B" box="[158,224,1134,1155]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="38" pageNumber="39" unit="km" value="12.0">12 km</quantity>
SW
</locationDeviation>
<location id="B52A607BFFD40E7AFEACFBD3FE29FB6B" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFEACFBD3FE29FB6B" box="[281,424,1134,1155]" country="Mongolia" name="Stariy Tsakhir" pageId="38" pageNumber="39">Stariy Tsakhir</location>
,
<date id="C44B1060FFD40E7AFE03FBD3FDA4FB6B" box="[438,549,1134,1155]" pageId="38" pageNumber="39" value="1975-06-28">28.VI.1975</date>
(
<collectorName id="1D005376FFD40E7AFD82FBD3FD58FB6B" box="[567,729,1134,1155]" pageId="38" pageNumber="39">A.F. Emeljanov</collectorName>
), 1J (
<collectionCode id="D6E4AE65FFD40E7AFF10FB34FF5AFB76" box="[165,219,1161,1182]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF44FB34FD36FB51" collectingDate="1975-06-28" collectionCode="ZISP" collectorName="E. L. Gurjeva" country="Mongolia" location="Ikh-Tamir" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<locationDeviation id="A1F82F82FFD40E7AFF44FB34FEE7FB76" bearing="315.0" box="[241,358,1161,1182]" distance="30000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFF44FB34FEB3FB76" box="[241,306,1161,1182]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="39" unit="km" value="30.0">30 km</quantity>
NW
</locationDeviation>
<location id="B52A607BFFD40E7AFEC5FB34FE52FB76" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFEC5FB34FE52FB76" box="[368,467,1161,1182]" country="Mongolia" name="Ikh-Tamir" pageId="38" pageNumber="39">Ikh-Tamir</location>
, stony low bunchgrass steppe, under stones,
<date id="C44B1060FFD40E7AFE9BFB19FE1DFB51" box="[302,412,1188,1209]" pageId="38" pageNumber="39" value="1975-06-28">28.VI.1975</date>
(
<collectorName id="1D005376FFD40E7AFE1AFB19FDB7FB51" box="[431,566,1188,1209]" pageId="38" pageNumber="39">E.L. Gurjeva</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFDFFFB19FDE9FB51" box="[586,616,1187,1209]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFDCFFB19FD31FB51" box="[634,688,1188,1209]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFD73FB19FF63FB06" collectingDate="1975-07-07" collectionCode="ZISP" collectorName="E. L. Gurjeva" country="Mongolia" location="Bugat" municipality="Bulgan Prov" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFD73FB19FF4AFB3B" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFD73FB19FF4AFB3B" pageId="38" pageNumber="39">Bulgan Prov</collectingMunicipality>
</emphasis>
.:
<locationDeviation id="A1F82F82FFD40E7AFF53FB03FEEDFB3B" bearing="22.5" box="[230,364,1214,1235]" distance="10000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFF53FB03FEADFB3B" box="[230,300,1214,1235]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="km" value="10.0">10 km</quantity>
NNE
</locationDeviation>
<location id="B52A607BFFD40E7AFECEFB03FE36FB3B" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFECEFB03FE36FB3B" box="[379,439,1214,1235]" country="Mongolia" municipality="Bulgan Prov" name="Bugat" pageId="38" pageNumber="39">Bugat</location>
,
<date id="C44B1060FFD40E7AFE7FFB03FDB5FB3B" box="[458,564,1214,1235]" pageId="38" pageNumber="39" value="1975-07-07">7.VII.1975</date>
(
<collectorName id="1D005376FFD40E7AFDFFFB03FD54FB3B" box="[586,725,1214,1235]" pageId="38" pageNumber="39">E.L. Gurjeva</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD5BFB03FC8DFB3C" box="[750,780,1214,1236]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFF10FB64FF5AFB06" box="[165,219,1241,1262]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF45FB64FDB2FAE0" collectingDate="1976-08-06" collectingDateMax="1976-08-09" collectingDateMin="1976-08-06" collectionCode="ZISP" collectorName="E. L. Gurjeva" country="Mongolia" location="Salkhit Mt." municipality="Numre-chiyn-Gol Riv." pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="2">
<collectingMunicipality id="502EACDAFFD40E7AFF45FB64FE50FB06" box="[240,465,1241,1262]" pageId="38" pageNumber="39">Numre-chiyn-Gol Riv.</collectingMunicipality>
,
<locationDeviation id="A1F82F82FFD40E7AFE6BFB64FDC5FB06" bearing="135.0" box="[478,580,1241,1262]" distance="32000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFE6BFB64FD9FFB06" box="[478,542,1241,1262]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="39" unit="km" value="32.0">32 km</quantity>
SE
</locationDeviation>
<location id="B52A607BFFD40E7AFDFBFB64FD40FB06" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFDFBFB64FD40FB06" box="[590,705,1241,1262]" country="Mongolia" municipality="Numre-chiyn-Gol Riv." name="Salkhit Mt." pageId="38" pageNumber="39">Salkhit Mt.</location>
,
<date id="C44B1060FFD40E7AFD7BFB64FC89FB06" box="[718,776,1241,1262]" pageId="38" pageNumber="39">15.VI</date>
,
<date id="C44B1060FFD40E7AFF2BFB4EFEA8FAE0" box="[158,297,1267,1289]" pageId="38" pageNumber="39" value="1976-08-06" valueMax="1976-08-09" valueMin="1976-08-06">6–9.VIII.1976</date>
(
<collectorName id="1D005376FFD40E7AFE82FB4EFE3BFAE0" box="[311,442,1267,1288]" pageId="38" pageNumber="39">E.L. Gurjeva</collectorName>
),
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(
<collectionCode id="D6E4AE65FFD40E7AFE40FB4EFDAAFAE0" box="[501,555,1267,1288]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFD88FB4EFD19FACB" collectingDate="1975-05-30" collectionCode="ZISP" collectorName="E. Sugonjaev" country="Mongolia" location="Darkhan" municipality="Selenge Prov" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD40E7AFD88FB4EFD3DFAE0" box="[573,700,1267,1288]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFD88FB4EFD3DFAE0" box="[573,700,1267,1288]" pageId="38" pageNumber="39">Selenge Prov</collectingMunicipality>
</emphasis>
.:
<locationDeviation id="A1F82F82FFD40E7AFD7AFB4EFF2AFACB" bearing="180.0" distance="25000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFD7AFB4EFC8DFAE0" box="[719,780,1267,1288]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="km" value="25.0">25 km</quantity>
S
</locationDeviation>
<location id="B52A607BFFD40E7AFF00FAB3FE8AFACB" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF00FAB3FE8AFACB" box="[181,267,1294,1315]" country="Mongolia" municipality="Selenge Prov" name="Darkhan" pageId="38" pageNumber="39">Darkhan</location>
,
<date id="C44B1060FFD40E7AFEACFAB3FEFCFACB" box="[281,381,1294,1315]" pageId="38" pageNumber="39" value="1975-05-30">30.V.1975</date>
(
<collectorName id="1D005376FFD40E7AFE3BFAB3FD96FACB" box="[398,535,1294,1315]" pageId="38" pageNumber="39">E. Sugonjaev</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD9EFAB3FDC8FACB" box="[555,585,1293,1315]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFDEEFAB3FD11FACB" box="[603,656,1294,1315]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFD10FAB3FF63FAB0" collectingDate="1975-07-27" collectionCode="ZISP" collectorName="M. A. Kozlov" country="Mongolia" location="Ingetiyn-Gol Riv." municipality="Selenge" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
10 км NE
<collectingMunicipality id="502EACDAFFD40E7AFF2BFA95FF6CFAD5" box="[158,237,1320,1341]" pageId="38" pageNumber="39">Selenge</collectingMunicipality>
,
<location id="B52A607BFFD40E7AFF48FA95FE33FAD5" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFF48FA95FE33FAD5" box="[253,434,1320,1341]" country="Mongolia" municipality="Selenge" name="Ingetiyn-Gol Riv." pageId="38" pageNumber="39">Ingetiyn-Gol Riv.</location>
,
<date id="C44B1060FFD40E7AFE74FA95FDB6FAD5" box="[449,567,1320,1341]" pageId="38" pageNumber="39" value="1975-07-27">27.VII.1975</date>
(
<collectorName id="1D005376FFD40E7AFDF9FA95FD56FAD5" box="[588,727,1320,1341]" pageId="38" pageNumber="39">M.A. Kozlov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD5BFA95FC8DFAD6" box="[750,780,1320,1342]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFF10FAFEFF5AFAB0" box="[165,219,1347,1368]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFF5AFAFEFD6CFA9B" collectingDate="1976-06" collectorName="M. A. Kozlov" country="Mongolia" location="- Nur Lake" municipality="Derkhin-Tsagan-Obo Riv." pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="2" stateProvince="Dornod Prov">
<emphasis id="8281EAB2FFD40E7AFF5AFAFEFEF0FAB0" box="[239,369,1347,1368]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="7231F842FFD40E7AFF5AFAFEFEF0FAB0" box="[239,369,1347,1368]" pageId="38" pageNumber="39">Dornod Prov</collectingRegion>
</emphasis>
.,
<collectingMunicipality id="502EACDAFFD40E7AFE33FAFEFD09FAB0" box="[390,648,1347,1368]" pageId="38" pageNumber="39">Derkhin-Tsagan-Obo Riv.</collectingMunicipality>
,
<locationDeviation id="A1F82F82FFD40E7AFD21FAFEFC8DFAB0" bearing="67.5" box="[660,780,1347,1368]" distance="60000.0" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFD21FAFEFD52FAB0" box="[660,723,1347,1368]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="38" pageNumber="39" unit="km" value="60.0">60 km</quantity>
ENE
</locationDeviation>
Bayan-Burd[
<location id="B52A607BFFD40E7AFEA9FAE3FE06FA9B" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7AFF53FEFCFA99FC1A:B52A607BFFD40E7AFEA9FAE3FE06FA9B" box="[284,391,1374,1395]" country="Mongolia" municipality="Derkhin-Tsagan-Obo Riv." name="- Nur Lake" pageId="38" pageNumber="39" stateProvince="Dornod Prov">-Nur Lake</location>
], 12,13.
<date id="C44B1060FFD40E7AFE61FAE3FDA7FA9B" box="[468,550,1374,1395]" pageId="38" pageNumber="39" value="1976-06">VI.1976</date>
(
<collectorName id="1D005376FFD40E7AFD80FAE3FD3DFA9B" box="[565,700,1374,1395]" pageId="38" pageNumber="39">M.A. Kozlov</collectorName>
),
<specimenCount id="A6F3FD29FFD40E7AFD7BFAE3FD6CFA9B" box="[718,749,1373,1395]" count="2" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7AFF2BFA78FAACFD17" pageId="38" pageNumber="39" type="description">
<paragraph id="B04A36A0FFD40E7AFF2BFA78FEA1FA34" blockId="38.[158,781,193,1938]" box="[158,288,1477,1500]" pageId="38" pageNumber="39">
<heading id="EB0281CCFFD40E7AFF2BFA78FEA1FA34" bold="true" box="[158,288,1477,1500]" fontSize="10" level="2" pageId="38" pageNumber="39" reason="6">
<emphasis id="8281EAB2FFD40E7AFF2BFA78FEA1FA34" bold="true" box="[158,288,1477,1500]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD40E7AFF2BFA50FEE5F94B" blockId="38.[158,781,193,1938]" pageId="38" pageNumber="39">Distinguished from all other species by metapleural evaporatorium with short and almost straight peritreme slightly protruding beyond middle of metapleuron. Lateral margins of pronotum, corium, abdominal venter and connexivum with yellowish narrow stripe. Membrane brownish, opaque.</paragraph>
<paragraph id="B04A36A0FFD40E7AFF15F901FE4DF93B" blockId="38.[158,781,193,1938]" box="[160,460,1724,1747]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFF15F901FE9BF93B" bold="true" box="[160,282,1724,1747]" pageId="38" pageNumber="39">Pygophore</emphasis>
as in
<taxonomicName id="77F54D23FFD40E7AFEEAF901FE44F93B" authorityName="Asanova" authorityYear="1964" box="[351,453,1724,1747]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD40E7AFEEAF901FE44F93B" box="[351,453,1724,1747]" italics="true" pageId="38" pageNumber="39">C. mixtus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFD40E7AFF15F951FE52F8CA" blockId="38.[158,781,193,1938]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFF15F951FE8EF8EB" bold="true" box="[160,271,1772,1795]" pageId="38" pageNumber="39">Paramere</emphasis>
with rather long and widely rounded apically sensory process (
<figureCitation id="28CE2A25FFD40E7AFEEDF8B6FE42F8CA" box="[344,451,1803,1826]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="38" pageNumber="39">
Figure 3
<collectionCode id="D6E4AE65FFD40E7AFE04F8B6FE42F8CA" box="[433,451,1803,1826]" pageId="38" pageNumber="39">L</collectionCode>
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).
</paragraph>
<paragraph id="B04A36A0FFD40E7AFF15F886FBA2FEB0" blockId="38.[158,781,193,1938]" lastBlockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFF15F886FE8CF8BA" bold="true" box="[160,269,1851,1874]" pageId="38" pageNumber="39">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFD40E7AFE94F886FE13F8BA" box="[289,402,1851,1874]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="38" pageNumber="39">
Figure 5
<collectionCode id="D6E4AE65FFD40E7AFECBF886FE13F8BA" box="[382,402,1851,1874]" pageId="38" pageNumber="39">D</collectionCode>
</figureCitation>
) similar to that of
<taxonomicName id="77F54D23FFD40E7AFDC4F886FD5BF8BA" authorityName="Asanova" authorityYear="1964" box="[625,730,1851,1874]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD40E7AFDC4F886FD5BF8BA" box="[625,730,1851,1874]" italics="true" pageId="38" pageNumber="39">C. mixtus</emphasis>
</taxonomicName>
and distinguished by somewhat longer dorsolateral lobes of conjunctiva and ventral lobe having convex lateral margins subapically and ventral wall convex along median line at apex in ventral view (
<figureCitation id="28CE2A25FFD40E7AFB3BFF5CFB7CFF10" box="[1166,1277,225,248]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="38" pageNumber="39">
Figure
<date id="C44B1060FFD40E7AFB68FF5CFB7CFF10" box="[1245,1277,225,248]" pageId="38" pageNumber="39">
6
<collectionCode id="D6E4AE65FFD40E7AFB5CFF5CFB7CFF10" box="[1257,1277,225,248]" pageId="38" pageNumber="39">V</collectionCode>
</date>
</figureCitation>
); in lateral view, lobe relatively wider and oblique apically (
<figureCitation id="28CE2A25FFD40E7AFA86FEBCFA2BFEF0" box="[1331,1450,257,280]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="38" pageNumber="39">
Figure 3
<collectionCode id="D6E4AE65FFD40E7AFA3AFEBCFA2BFEF0" box="[1423,1450,257,280]" pageId="38" pageNumber="39">W</collectionCode>
</figureCitation>
). Spicule at dorsolateral lobes weakly (
<figureCitation id="28CE2A25FFD40E7AFB62FE9CFAC1FED0" box="[1239,1344,289,312]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="38" pageNumber="39">
Figure 7
<collectionCode id="D6E4AE65FFD40E7AFA9AFE9CFAC1FED0" box="[1327,1344,289,312]" pageId="38" pageNumber="39">T</collectionCode>
</figureCitation>
) or sharply (
<figureCitation id="28CE2A25FFD40E7AFCE1FEFCFC43FEB0" box="[852,962,321,344]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="38" pageNumber="39">
Figure 7
<collectionCode id="D6E4AE65FFD40E7AFC1BFEFCFC43FEB0" box="[942,962,321,344]" pageId="38" pageNumber="39">U</collectionCode>
</figureCitation>
) curved.
</paragraph>
<paragraph id="B04A36A0FFD40E7AFCFBFECCFB98FDEF" blockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFCFBFECCFC4DFE60" bold="true" box="[846,972,369,392]" pageId="38" pageNumber="39">Gynatrium</emphasis>
almost as in
<taxonomicName id="77F54D23FFD40E7AFBE2FECCFB3DFE60" authorityName="Asanova" authorityYear="1964" box="[1111,1212,369,392]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD40E7AFBE2FECCFB3DFE60" box="[1111,1212,369,392]" italics="true" pageId="38" pageNumber="39">C. mixtus</emphasis>
</taxonomicName>
, with somewhat shorter posterior pouches (
<figureCitation id="28CE2A25FFD40E7AFBAEFE2CFB17FE40" box="[1051,1174,401,424]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="38" pageNumber="39">
Figure 11
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). First rami rather wide or narrow anteriorly, more or less obliquely truncated at anterior end, with rounded (
<figureCitation id="28CE2A25FFD40E7AFB25FE6CFA85FE00" box="[1168,1284,465,488]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="38" pageNumber="39">
Figure
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8
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) or acute inner angle (
<figureCitation id="28CE2A25FFD40E7AFC23FE4DFB8BFDEF" box="[918,1034,496,519]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="38" pageNumber="39">
Figure 8
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).
</paragraph>
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<emphasis id="8281EAB2FFD40E7AFCF9FDDCFC70FD90" bold="true" box="[844,1009,609,632]" pageId="38" pageNumber="39">Measurements</emphasis>
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Body 5.60–8.00 in length,
<quantity id="770D9B45FFD40E7AFBC6FD34FB7AFD48" box="[1139,1275,648,672]" metricMagnitude="-1" metricUnit="m" metricValue="1.09855" metricValueMax="1.3462" metricValueMin="0.8509" pageId="38" pageNumber="39" unit="in" value="4.325" valueMax="5.3" valueMin="3.35">3.35–5.30 in</quantity>
width; pronotum
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width; head
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width; ocular index 3.39–4.47; length of antennal segments I–V: 0.29–0.36: 0.53–0.63: 0.48–0.73: 0.69–0.94: 0.78–1.06.
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<emphasis id="8281EAB2FFD40E7AFCF9FCEEFC54FC82" bold="true" box="[844,981,851,874]" pageId="38" pageNumber="39">Distribution</emphasis>
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Russia (Western and Eastern Siberia, Far East), Kazakhstan, Mongolia, China (North, Central and Southwest Territories) Japan, the Korean Peninsula, east of India, and north of Vietnam (
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332
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<emphasis id="8281EAB2FFD40E7AFC13FB8FFB34FBA1" bold="true" box="[934,1205,1074,1097]" italics="true" pageId="38" pageNumber="39">Canthophorus coeruleus</emphasis>
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Figures 3
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,
<figureCitation id="28CE2A25FFD40E7AFB03FBEFFA85FB81" box="[1206,1284,1106,1129]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="38" pageNumber="39">
<date id="C44B1060FFD40E7AFB03FBEFFB56FB81" box="[1206,1239,1106,1129]" pageId="38" pageNumber="39">
8
<collectionCode id="D6E4AE65FFD40E7AFB76FBEFFB56FB81" box="[1219,1239,1106,1129]" pageId="38" pageNumber="39">X</collectionCode>
</date>
, 8
<collectionCode id="D6E4AE65FFD40E7AFB47FBEFFA85FB81" box="[1266,1284,1106,1129]" pageId="38" pageNumber="39">Y</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD40E7AFAA1FBEFFABEFB81" box="[1300,1343,1106,1129]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="38" pageNumber="39">
11
<collectionCode id="D6E4AE65FFD40E7AFA98FBEFFABEFB81" box="[1325,1343,1106,1129]" pageId="38" pageNumber="39">E</collectionCode>
</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7AFCF9FBC1FABDFB2E" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="B04A36A0FFD40E7AFCF9FBC1FC68FB43" blockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">
<materialsCitation id="009D3CFDFFD40E7AFCF9FBC1FC64FB43" country="Turkmenistan" location="Kopet Dag" pageId="38" pageNumber="39" specimenCount="1" typeStatus="syntype">
<taxonomicName id="77F54D23FFD40E7AFCF9FBC1FB39FB79" authority="Reuter 1902: 144" authorityName="Reuter" authorityPageNumber="144" authorityYear="1902" box="[844,1208,1148,1169]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="8281EAB2FFD40E7AFCF9FBC1FC7AFB79" box="[844,1019,1148,1169]" italics="true" pageId="38" pageNumber="39">Sehirus coeruleus</emphasis>
<bibRefCitation id="D4644B51FFD40E7AFBB2FBC1FB39FB79" author="Reuter OM" box="[1031,1208,1148,1169]" journalOrPublisher="Ofversigt af Finska Vetenskaps-Societetens Forhandlingar." pageId="38" pageNumber="39" pagination="141 - 188" part="44" refId="ref40588" refString="Reuter OM. 1902. Miscellanea Hemipterologica. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar. 44: 141 - 188." title="Miscellanea Hemipterologica" type="journal article" year="1902">Reuter 1902: 144</bibRefCitation>
</taxonomicName>
[
<typeStatus id="6F4E8802FFD40E7AFB7CFBC1FAA3FB79" box="[1225,1314,1148,1169]" pageId="38" pageNumber="39">syntypes</typeStatus>
:
<collectingCountry id="C8E27630FFD40E7AFA84FBC1FA37FB79" box="[1329,1462,1148,1169]" name="Turkmenistan" pageId="38" pageNumber="39" value="Turkmenistan">Turkmenistan</collectingCountry>
,
<location id="B52A607BFFD40E7AFCC4FB2BFC5EFB43" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7BFC13FB8FFE54FBC7:B52A607BFFD40E7AFCC4FB2BFC5EFB43" box="[881,991,1174,1195]" country="Turkmenistan" name="Kopet Dag" pageId="38" pageNumber="39">Kopet Dag</location>
]
</materialsCitation>
.
</paragraph>
<paragraph id="B04A36A0FFD40E7AFCF9FB0CFABDFB2E" blockId="38.[844,1466,194,1922]" box="[844,1340,1201,1222]" pageId="38" pageNumber="39">
<taxonomicName id="77F54D23FFD40E7AFCF9FB0CFBB9FB2E" baseAuthorityName="Reuter" baseAuthorityYear="1902" box="[844,1080,1201,1222]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="8281EAB2FFD40E7AFCF9FB0CFBB9FB2E" box="[844,1080,1201,1222]" italics="true" pageId="38" pageNumber="39">Canthophorus coeruleus</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFD40E7AFBF2FB0CFA87FB2E" author="Asanova RB" box="[1095,1286,1201,1222]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="38" pageNumber="39" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova 1964: 140</bibRefCitation>
, 143.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7AFCF9FABAFAE2F9FD" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="B04A36A0FFD40E7AFCF9FABAFBA3FAF6" blockId="38.[844,1466,194,1922]" box="[844,1058,1287,1310]" pageId="38" pageNumber="39">
<heading id="EB0281CCFFD40E7AFCF9FABAFBA3FAF6" bold="true" box="[844,1058,1287,1310]" fontSize="10" level="2" pageId="38" pageNumber="39" reason="6">
<emphasis id="8281EAB2FFD40E7AFCF9FABAFBA3FAF6" bold="true" box="[844,1058,1287,1310]" pageId="38" pageNumber="39">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD40E7AFCF9FA91FAE2F9FD" blockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">
<materialsCitation id="009D3CFDFFD40E7AFCF9FA91FBDEFAB3" collectionCode="ZISP" collectorName="O. M. Reuter" country="Turkmenistan" location="Ashgabat" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Ashgabat">
<emphasis id="8281EAB2FFD40E7AFCF9FA91FC5AFAA9" bold="true" box="[844,987,1324,1345]" pageId="38" pageNumber="39">
<collectingCountry id="C8E27630FFD40E7AFCF9FA91FC5AFAA9" box="[844,987,1324,1345]" name="Turkmenistan" pageId="38" pageNumber="39" value="Turkmenistan">Turkmenistan</collectingCountry>
</emphasis>
:
<collectingRegion id="7231F842FFD40E7AFC5CFA91FBC4FAA9" box="[1001,1093,1324,1345]" country="Turkmenistan" name="Ashgabat" pageId="38" pageNumber="39">Ashgabat</collectingRegion>
, [date and collector unknown], “
<collectorName id="1D005376FFD40E7AFA32FA91FC0AFAB3" pageId="38" pageNumber="39">O.M. Reuter</collectorName>
det.”,
<specimenCount id="A6F3FD29FFD40E7AFC5DFAFBFB87FAB4" box="[1000,1030,1350,1372]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFB97FAFBFBD6FAB3" box="[1058,1111,1350,1371]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD40E7AFBC2FAFBFAAEFA78" collectingDate="1908-06" collectionCode="ZISP" collectorName="B. Grigorjev" country="Uzbekistan" location="Namangan Fl. [umen] Padschaata Toste" pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="2" stateProvince="Namangan Prov">
<emphasis id="8281EAB2FFD40E7AFBC2FAFBFB69FAB3" bold="true" box="[1143,1256,1350,1371]" pageId="38" pageNumber="39">
<collectingCountry id="C8E27630FFD40E7AFBC2FAFBFB69FAB3" box="[1143,1256,1350,1371]" name="Uzbekistan" pageId="38" pageNumber="39" value="Uzbekistan">Uzbekistan</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD40E7AFAB6FAFBFA2CFAB3" box="[1283,1453,1350,1371]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="7231F842FFD40E7AFAB6FAFBFA2CFAB3" box="[1283,1453,1350,1371]" pageId="38" pageNumber="39">Namangan Prov</collectingRegion>
</emphasis>
., “Fergana sept[entrionalis].,
<location id="B52A607BFFD40E7AFBDCFADCFCFEFA78" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7BFC13FB8FFE54FBC7:B52A607BFFD40E7AFBDCFADCFCFEFA78" country="Uzbekistan" name="Namangan Fl. [umen] Padschaata Toste" pageId="38" pageNumber="39" stateProvince="Namangan Prov">Namangan Fl.[umen] Padschaata Toste</location>
” 15,18.
<date id="C44B1060FFD40E7AFC65FAC6FB9EFA78" box="[976,1055,1403,1424]" pageId="38" pageNumber="39" value="1908-06">
<collectingDate id="D40FE988FFD40E7AFC65FAC6FB9EFA78" box="[976,1055,1403,1424]" pageId="38" pageNumber="39" value="1908-06">VI.1908</collectingDate>
</date>
(
<collectorName id="1D005376FFD40E7AFB86FAC6FB30FA78" box="[1075,1201,1403,1424]" pageId="38" pageNumber="39">B. Grigorjev</collectorName>
)
<specimenCount id="A6F3FD29FFD40E7AFB76FAC6FB60FA79" box="[1219,1249,1403,1425]" count="2" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFB46FAC6FAA9FA78" box="[1267,1320,1403,1424]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFD40E7AFA8BFAC6FA37FA2E" collectingDate="1937-09-06" collectionCode="ZISP" country="Kyrgyzstan" location="Ak-terek" municipality="Fergana Range" pageId="38" pageNumber="39" specimenCount="3" specimenCount-female="3" stateProvince="Jalal-Abad Prov">
<emphasis id="8281EAB2FFD40E7AFA8BFAC6FA32FA78" bold="true" box="[1342,1459,1403,1424]" pageId="38" pageNumber="39">
<collectingCountry id="C8E27630FFD40E7AFA8BFAC6FA32FA78" box="[1342,1459,1403,1424]" name="Kyrgyzstan" pageId="38" pageNumber="39" value="Kyrgyzstan">Kyrgyzstan</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD40E7AFCF9FA2BFC69FA43" box="[844,1000,1430,1451]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="7231F842FFD40E7AFCF9FA2BFC69FA43" box="[844,1000,1430,1451]" pageId="38" pageNumber="39">Jalal-Abad Prov</collectingRegion>
</emphasis>
.,
<collectingMunicipality id="502EACDAFFD40E7AFC4FFA2BFB0DFA43" box="[1018,1164,1430,1451]" pageId="38" pageNumber="39">Fergana Range</collectingMunicipality>
, urochishche
<location id="B52A607BFFD40E7AFAA5FA2BFAE7FA43" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7BFC13FB8FFE54FBC7:B52A607BFFD40E7AFAA5FA2BFAE7FA43" box="[1296,1382,1430,1451]" country="Kyrgyzstan" municipality="Fergana Range" name="Ak-terek" pageId="38" pageNumber="39" stateProvince="Jalal-Abad Prov">Ak-terek</location>
,
<locationDeviation id="A1F82F82FFD40E7AFAC5FA2BFC01FA2E" pageId="38" pageNumber="39">
<quantity id="770D9B45FFD40E7AFAC5FA2BFA20FA43" box="[1392,1441,1430,1451]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="km" value="5.0">5 km</quantity>
N Gava
</locationDeviation>
, 5,6,8,16.VIII.,
<date id="C44B1060FFD40E7AFB94FA0CFB03FA2E" box="[1057,1154,1457,1478]" pageId="38" pageNumber="39" value="1937-09-06">
<collectingDate id="D40FE988FFD40E7AFB94FA0CFB03FA2E" box="[1057,1154,1457,1478]" pageId="38" pageNumber="39" value="1937-09-06">6.IX.1937</collectingDate>
</date>
(Kiritshenko), 4J,
<specimenCount id="A6F3FD29FFD40E7AFAFFFA0CFAE9FA2E" box="[1354,1384,1456,1478]" count="3" pageId="38" pageNumber="39" type="female">3♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFACFFA0CFA2EFA2E" box="[1402,1455,1457,1478]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFCF9FA76FB29FA13" collectingDate="1909-05-26" collectionCode="ZISP" collectorName="A. Jacobson" country="Kyrgyzstan" location="Osh Prov" municipality="Osh Prov" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Jalal-abad Prov">
<emphasis id="8281EAB2FFD40E7AFCF9FA76FC24FA08" box="[844,933,1483,1504]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="502EACDAFFD40E7AFCF9FA76FC24FA08" box="[844,933,1483,1504]" pageId="38" pageNumber="39">Osh Prov</collectingMunicipality>
</emphasis>
.:
<locationDeviation id="A1F82F82FFD40E7AFC0DFA76FAA5FA08" box="[952,1316,1483,1505]" pageId="38" pageNumber="39">from “Lyangor” [Lyangar] to Gul’ cha</locationDeviation>
, 1800 [m asl?],
<date id="C44B1060FFD40E7AFCF9FA5BFC2FFA13" box="[844,942,1510,1531]" pageId="38" pageNumber="39" value="1909-05-26">
<collectingDate id="D40FE988FFD40E7AFCF9FA5BFC2FFA13" box="[844,942,1510,1531]" pageId="38" pageNumber="39" value="1909-05-26">26.V.1909</collectingDate>
</date>
(
<collectorName id="1D005376FFD40E7AFC09FA5BFBB2FA13" box="[956,1075,1510,1531]" pageId="38" pageNumber="39">A. Jacobson</collectorName>
)
<specimenCount id="A6F3FD29FFD40E7AFBF5FA5BFBDFFA13" box="[1088,1118,1509,1531]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFBDEFA5BFB21FA13" box="[1131,1184,1510,1531]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD40E7AFB07FA5BFAE1F9FD" collectingDate="1995-07-10" collectionCode="ZISP" collectorName="A. Koval" country="Kyrgyzstan" elevation="3200" location="Berk-Suu Riv." pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="2" stateProvince="Zaalaisky Range">
<collectingRegion id="7231F842FFD40E7AFB07FA5BFAD5FA13" box="[1202,1364,1510,1531]" pageId="38" pageNumber="39">Zaalaisky Range</collectingRegion>
,
<location id="B52A607BFFD40E7AFAEBFA5BFCF7F9FD" LSID="urn:lsid:plazi:treatment:385C87B6FFD40E7BFC13FB8FFE54FBC7:B52A607BFFD40E7AFAEBFA5BFCF7F9FD" country="Kyrgyzstan" name="Berk-Suu Riv." pageId="38" pageNumber="39" stateProvince="Zaalaisky Range">Berk-Suu Riv.</location>
,
<quantity id="770D9B45FFD40E7AFC35F9BDFC4CF9FD" box="[896,973,1536,1557]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="39" unit="m" value="3200.0">
<elevation id="3BD8D193FFD40E7AFC35F9BDFC4CF9FD" box="[896,973,1536,1557]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="39" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="C44B1060FFD40E7AFC6DF9BDFBCCF9FD" box="[984,1101,1536,1557]" pageId="38" pageNumber="39" value="1995-07-10">
<collectingDate id="D40FE988FFD40E7AFC6DF9BDFBCCF9FD" box="[984,1101,1536,1557]" pageId="38" pageNumber="39" value="1995-07-10">10.VII.1995</collectingDate>
</date>
(
<collectorName id="1D005376FFD40E7AFBE9F9BDFB39F9FD" box="[1116,1208,1536,1557]" pageId="38" pageNumber="39">A. Koval</collectorName>
), 1J,
<specimenCount id="A6F3FD29FFD40E7AFB42F9BDFA94F9FE" box="[1271,1301,1536,1558]" count="2" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD40E7AFA96F9BDFAD9F9FD" box="[1315,1368,1536,1557]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="38" pageNumber="39">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD40E7BFCF9F9D2FDFEFC6B" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="B04A36A0FFD40E7AFCF9F9D2FC4FF96E" blockId="38.[844,1466,194,1922]" box="[844,974,1647,1670]" pageId="38" pageNumber="39">
<heading id="EB0281CCFFD40E7AFCF9F9D2FC4FF96E" bold="true" box="[844,974,1647,1670]" fontSize="10" level="2" pageId="38" pageNumber="39" reason="6">
<emphasis id="8281EAB2FFD40E7AFCF9F9D2FC4FF96E" bold="true" box="[844,974,1647,1670]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD40E7AFCF9F92AFA81F8C5" blockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">Distinguished from all other species by lateral margins of pronotum, corium, abdominal venter and connexivum without light stripes and spots, unicolour with other parts of body. Metapleural evaporatorium with long and curved peritreme. Membrane brownish, opaque.</paragraph>
<paragraph id="B04A36A0FFD40E7AFCFBF8F6FB94F86A" blockId="38.[844,1466,194,1922]" pageId="38" pageNumber="39">
<emphasis id="8281EAB2FFD40E7AFCFBF8F6FC49F88A" bold="true" box="[846,968,1867,1890]" pageId="38" pageNumber="39">Pygophore</emphasis>
as in
<taxonomicName id="77F54D23FFD40E7AFBA8F8F6FB6EF88A" authorityName="Scott" authorityYear="1874" box="[1053,1263,1867,1890]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="niveimarginatus">
<emphasis id="8281EAB2FFD40E7AFBA8F8F6FB6EF88A" box="[1053,1263,1867,1890]" italics="true" pageId="38" pageNumber="39">C. niveimarginatus</emphasis>
</taxonomicName>
but with slightly longer dorsal wall.
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF15FF7CFEE2FEF0" blockId="39.[158,781,193,1868]" pageId="39" pageNumber="40">
<emphasis id="8281EAB2FFD50E7BFF15FF7CFE8EFF30" bold="true" box="[160,271,193,216]" pageId="39" pageNumber="40">Paramere</emphasis>
. Sensory process more strongly curved dorsad than in all previous species, rather long, widely rounded at apex (
<figureCitation id="28CE2A25FFD50E7BFF54FEBCFED2FEF0" box="[225,339,257,280]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="39" pageNumber="40">
Figure 3
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).
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF15FE8CFE3EFDEF" blockId="39.[158,781,193,1868]" pageId="39" pageNumber="40">
<emphasis id="8281EAB2FFD50E7BFF15FE8CFE8CFEA0" bold="true" box="[160,269,305,328]" pageId="39" pageNumber="40">Aedeagus</emphasis>
almost as in
<taxonomicName id="77F54D23FFD50E7BFE2CFE8CFDE2FEA0" authorityName="Scott" authorityYear="1874" box="[409,611,305,328]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="niveimarginatus">
<emphasis id="8281EAB2FFD50E7BFE2CFE8CFDE2FEA0" box="[409,611,305,328]" italics="true" pageId="39" pageNumber="40">C. niveimarginatus</emphasis>
</taxonomicName>
. Ventral lobe in ventral view slightly narrower, with almost parallel lateral margins, very slightly convex distal to middle and slightly converging apically; extreme apex of lobe short and narrow (
<figureCitation id="28CE2A25FFD50E7BFF62FE0CFEC2FE20" box="[215,323,433,456]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="39" pageNumber="40">
Figure
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6
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); ventral wall flattened over entire surface; in lateral view, lobe rather narrow, with rectangularly rounded apex (
<figureCitation id="28CE2A25FFD50E7BFEF4FE4DFE2EFDEF" box="[321,431,496,519]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="39" pageNumber="40">
Figure 6
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).
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF15FD9DFDA8FD7F" blockId="39.[158,781,193,1868]" pageId="39" pageNumber="40">
<emphasis id="8281EAB2FFD50E7BFF15FD9DFE9FFDDF" bold="true" box="[160,286,544,567]" pageId="39" pageNumber="40">Gynatrium</emphasis>
as in
<taxonomicName id="77F54D23FFD50E7BFEEBFD9DFE42FDDF" authorityName="Asanova" authorityYear="1964" box="[350,451,544,567]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD50E7BFEEBFD9DFE42FDDF" box="[350,451,544,567]" italics="true" pageId="39" pageNumber="40">C. mixtus</emphasis>
</taxonomicName>
, with somewhat smaller apical part of gynatrial sac (
<figureCitation id="28CE2A25FFD50E7BFE3BFDFDFD86FDBF" box="[398,519,576,599]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="39" pageNumber="40">
Figure 11
<collectionCode id="D6E4AE65FFD50E7BFE40FDFDFD86FDBF" box="[501,519,576,599]" pageId="39" pageNumber="40">E</collectionCode>
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). First rami moderately wide anteriorly and acutely (
<figureCitation id="28CE2A25FFD50E7BFE68FDDDFDCFFD9F" box="[477,590,608,631]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="39" pageNumber="40">
Figure
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8
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) or rectangularly (
<figureCitation id="28CE2A25FFD50E7BFF12FD3DFE94FD7F" box="[167,277,640,663]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="39" pageNumber="40">
Figure 8
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) rounded at anterior end.
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF2BFD59FEC2FD13" blockId="39.[158,781,193,1868]" box="[158,323,740,763]" pageId="39" pageNumber="40">
<heading id="EB0281CCFFD50E7BFF2BFD59FEC2FD13" bold="true" box="[158,323,740,763]" fontSize="10" level="2" pageId="39" pageNumber="40" reason="6">
<emphasis id="8281EAB2FFD50E7BFF2BFD59FEC2FD13" bold="true" box="[158,323,740,763]" pageId="39" pageNumber="40">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF2BFCB1FDFEFC6B" blockId="39.[158,781,193,1868]" pageId="39" pageNumber="40">
Body
<quantity id="770D9B45FFD50E7BFF57FCB1FEEBFCCB" box="[226,362,780,804]" metricMagnitude="-1" metricUnit="m" metricValue="1.89865" metricValueMax="2.1336" metricValueMin="1.6637" pageId="39" pageNumber="40" unit="in" value="7.475" valueMax="8.4" valueMin="6.55">6.55–8.40 in</quantity>
length,
<quantity id="770D9B45FFD50E7BFE73FCB1FDCFFCCB" box="[454,590,780,804]" metricMagnitude="-1" metricUnit="m" metricValue="1.1303" metricValueMax="1.2827" metricValueMin="0.9779" pageId="39" pageNumber="40" unit="in" value="4.45" valueMax="5.05" valueMin="3.85">3.85–5.05 in</quantity>
width; pronotum
<quantity id="770D9B45FFD50E7BFF2BFC91FEA5FCAB" box="[158,292,811,835]" metricMagnitude="-1" metricUnit="m" metricValue="1.02235" metricValueMax="1.143" metricValueMin="0.9017" pageId="39" pageNumber="40" unit="in" value="4.025" valueMax="4.5" valueMin="3.55">3.55–4.50 in</quantity>
width; head
<quantity id="770D9B45FFD50E7BFE05FC91FDB7FCAB" box="[432,566,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="4.3942" metricValueMax="4.8006" metricValueMin="3.9878" pageId="39" pageNumber="40" unit="in" value="1.73" valueMax="1.89" valueMin="1.57">1.57–1.89 in</quantity>
width; ocular index 3.74–3.95; length of antennal segments I–V: 0.35–0.46: 0.64–0.83: 0.63–0.77: 0.90–1.09: 0.94–1.06.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD50E7BFF2BFC6DFE54FBC7" pageId="39" pageNumber="40" type="distribution">
<paragraph id="B04A36A0FFD50E7BFF2BFC6DFEA6FC0F" blockId="39.[158,781,193,1868]" box="[158,295,976,999]" pageId="39" pageNumber="40">
<heading id="EB0281CCFFD50E7BFF2BFC6DFEA6FC0F" bold="true" box="[158,295,976,999]" fontSize="10" level="2" pageId="39" pageNumber="40" reason="6">
<emphasis id="8281EAB2FFD50E7BFF2BFC6DFEA6FC0F" bold="true" box="[158,295,976,999]" pageId="39" pageNumber="40">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD50E7BFF2BFC45FE54FBC7" blockId="39.[158,781,193,1868]" pageId="39" pageNumber="40">
Iran (
<bibRefCitation id="D4644B51FFD50E7BFF57FC45FECDFBE7" author="Lis JA" box="[226,332,1016,1039]" editor="Aukema B &amp; Rieger C" journalOrPublisher="Amsterdam: Netherlands Entomological Society" pageId="39" pageNumber="40" pagination="119 - 147" refId="ref40328" refString="Lis JA. 2006. Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs). In: Aukema B and Rieger C, editors. Catalogue of the Heteroptera of the Palaearctic Region Vol. 5, Amsterdam: Netherlands Entomological Society; p. 119 - 147." title="Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs)" type="book chapter" volumeTitle="Catalogue of the Heteroptera of the Palaearctic Region" year="2006">Lis 2006</bibRefCitation>
;
<bibRefCitation id="D4644B51FFD50E7BFED3FC45FDD3FBE7" author="Ghahari H &amp; Cherot F &amp; Linnavuori RE &amp; Ostovan H." box="[358,594,1016,1039]" journalOrPublisher="ZooKeys." pageId="39" pageNumber="40" pagination="1 - 31" part="26" refId="ref39906" refString="Ghahari H, Cherot F, Linnavuori RE, Ostovan H. 2009. Annotated catalogue of Iranian burrower bugs (Heteroptera, Pentatomoidea, Cydnidae). ZooKeys. 26: 1 - 31." title="Annotated catalogue of Iranian burrower bugs (Heteroptera, Pentatomoidea, Cydnidae)" type="journal article" year="2009">Ghahari et al. 2009</bibRefCitation>
), Turkmenistan, Uzbekistan, and Kyrgyzstan.
</paragraph>
</subSubSection>
</treatment>
</document>

281
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<mods:identifier id="D3D07CBC980ED8E93152A964E74C798E" type="DOI">10.1080/00379271.2018.1467788</mods:identifier>
<mods:identifier id="B9E48A07C1160CEEC39512BE5A913697" type="Zenodo-Dep">14927017</mods:identifier>
</mods:mods>
<treatment id="385C87B6FFD50E7BFC26FF7FFA2CFD13" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFC26FF7FFA2CFD13" httpUri="http://treatment.plazi.org/id/385C87B6FFD50E7BFC26FF7FFA2CFD13" lastPageNumber="40" pageId="39" pageNumber="40">
<subSubSection id="F8EF652BFFD50E7BFC26FF7FFA0AFF10" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="B04A36A0FFD50E7BFC26FF7FFA0AFF10" blockId="39.[844,1467,194,1868]" pageId="39" pageNumber="40">
<emphasis id="8281EAB2FFD50E7BFC26FF7FFAF2FF31" bold="true" box="[915,1395,194,217]" pageId="39" pageNumber="40">
<taxonomicName id="77F54D23FFD50E7BFC26FF7FFA89FF31" authority="(Walker, 1867)" authorityName="Gapon" authorityYear="2018" baseAuthorityName="Walker" baseAuthorityYear="1867" box="[915,1288,194,217]" class="Insecta" family="Cydnidae" genus="Adomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="aeneus" status="comb. nov.">
<emphasis id="8281EAB2FFD50E7BFC26FF7FFBD6FF31" bold="true" box="[915,1111,194,217]" italics="true" pageId="39" pageNumber="40">Adomerus aeneus</emphasis>
(
<bibRefCitation id="D4644B51FFD50E7BFBD2FF7FFB7FFF31" author="Walker F." box="[1127,1278,194,217]" journalOrPublisher="London: British Museum (Natural History)" pageId="39" pageNumber="40" refId="ref41010" refString="Walker F. 1867. Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Vol. 1. London: British Museum (Natural History)." title="Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Vol. 1" type="book" year="1867">Walker, 1867</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="99B257C9FFD50E7BFAA5FF7FFAF2FF31" box="[1296,1395,194,217]" pageId="39" pageNumber="40" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
<emphasis id="8281EAB2FFD50E7BFCCFFF5CFA0AFF10" bold="true" box="[890,1419,225,249]" pageId="39" pageNumber="40">
(
<figureCitation id="28CE2A25FFD50E7BFC37FF5CFBB1FF10" box="[898,1072,225,248]" captionStart="Figure 2" captionStartId="6.[158,225,1725,1746]" captionTargetBox="[209,1426,182,1704]" captionTargetId="figure-102@6.[209,1426,182,1704]" captionTargetPageId="6" captionText="Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927021" httpUri="https://zenodo.org/record/14927021/files/figure.png" pageId="39" pageNumber="40">
Figures 2
<collectionCode id="D6E4AE65FFD50E7BFC58FF5CFB81FF10" box="[1005,1024,225,248]" pageId="39" pageNumber="40">C</collectionCode>
, 2
<collectionCode id="D6E4AE65FFD50E7BFBA8FF5CFBB1FF10" box="[1053,1072,225,248]" pageId="39" pageNumber="40">D</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD50E7BFBF5FF5CFB10FF10" box="[1088,1169,225,248]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="39" pageNumber="40">
3
<collectionCode id="D6E4AE65FFD50E7BFBF9FF5CFBDEFF10" box="[1100,1119,225,248]" pageId="39" pageNumber="40">N</collectionCode>
, 3
<collectionCode id="D6E4AE65FFD50E7BFBC9FF5CFB10FF10" box="[1148,1169,225,248]" pageId="39" pageNumber="40">O</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD50E7BFB14FF5CFA88FF10" box="[1185,1289,225,249]" captionStart="Figure 18" captionStartId="40.[158,225,1842,1863]" captionTargetBox="[204,1423,1115,1811]" captionTargetId="figure-555@40.[195,1431,1105,1819]" captionTargetPageId="40" captionText="Figure 18. Aedeagus of Adomerus aeneus (Walker, 1867) n. comb. A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927053" httpUri="https://zenodo.org/record/14927053/files/figure.png" pageId="39" pageNumber="40">
18
<collectionCode id="D6E4AE65FFD50E7BFB0EFF5CFB4EFF10" box="[1211,1231,225,248]" pageId="39" pageNumber="40">A</collectionCode>
–18
<collectionCode id="D6E4AE65FFD50E7BFB43FF5CFA88FF10" box="[1270,1289,225,248]" pageId="39" pageNumber="40">C</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFD50E7BFAACFF5CFA03FF10" box="[1305,1410,225,248]" captionStart="Figure 20" captionStartId="42.[158,225,1723,1744]" captionTargetBox="[207,1405,176,1698]" captionTargetId="figure-99@42.[196,1426,168,1706]" captionTargetPageId="42" captionText="Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927057" httpUri="https://zenodo.org/record/14927057/files/figure.png" pageId="39" pageNumber="40">
20
<collectionCode id="D6E4AE65FFD50E7BFA81FF5CFAC6FF10" box="[1332,1351,225,248]" pageId="39" pageNumber="40">A</collectionCode>
, 20
<collectionCode id="D6E4AE65FFD50E7BFAC4FF5CFA03FF10" box="[1393,1410,225,248]" pageId="39" pageNumber="40">B</collectionCode>
</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD50E7BFCF9FEB6FAE9FE62" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="B04A36A0FFD50E7BFCF9FEB6FAF1FEC8" blockId="39.[844,1467,194,1868]" box="[844,1392,267,288]" pageId="39" pageNumber="40">
<taxonomicName id="77F54D23FFD50E7BFCF9FEB6FBF3FEC8" authority="Walker (1867)" authorityName="Walker" authorityYear="1867" box="[844,1138,267,288]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="aeneus">
<emphasis id="8281EAB2FFD50E7BFCF9FEB6FC5CFEC8" box="[844,989,267,288]" italics="true" pageId="39" pageNumber="40">Sehirus aeneus</emphasis>
<bibRefCitation id="D4644B51FFD50E7BFC50FEB6FBF3FEC8" author="Walker F." box="[997,1138,267,288]" journalOrPublisher="London: British Museum (Natural History)" pageId="39" pageNumber="40" refId="ref41010" refString="Walker F. 1867. Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Vol. 1. London: British Museum (Natural History)." title="Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Vol. 1" type="book" year="1867">Walker (1867)</bibRefCitation>
</taxonomicName>
: 169 [
<typeStatus id="6F4E8802FFD50E7BFB1AFEB6FA89FEC8" box="[1199,1288,267,288]" pageId="39" pageNumber="40">holotype</typeStatus>
: Madeira].
</paragraph>
<paragraph id="B04A36A0FFD50E7BFCF9FE98FB8BFEBD" blockId="39.[844,1467,194,1868]" pageId="39" pageNumber="40">
<taxonomicName id="77F54D23FFD50E7BFCF9FE98FB32FED2" authority="Horvath (1899)" authorityName="Horvath" authorityYear="1899" box="[844,1203,293,314]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="8281EAB2FFD50E7BFCF9FE98FB8BFED2" box="[844,1034,293,314]" italics="true" pageId="39" pageNumber="40">Sehirus fuscipennis</emphasis>
<bibRefCitation id="D4644B51FFD50E7BFBA2FE98FB32FED2" author="Horvath G." box="[1047,1203,293,314]" journalOrPublisher="Revue d' Entomologie." pageId="39" pageNumber="40" pagination="82 - 84" part="18" refId="ref40073" refString="Horvath G. 1899. Note sur le Sehirus maculipes et espe`ces affines. Revue d' Entomologie. 18: 82 - 84." title="Note sur le Sehirus maculipes et espe`ces affines" type="journal article" year="1899">Horváth (1899)</bibRefCitation>
</taxonomicName>
: 83 (syn. by
<bibRefCitation id="D4644B51FFD50E7BFAFFFE98FC7CFEBD" author="Aukema B &amp; Constant J." journalOrPublisher="Entomologica Americana." pageId="39" pageNumber="40" pagination="42 - 46" part="122" refId="ref39601" refString="Aukema B, Constant J. 2016. The taxonomic status of Sehirus aeneus Walker, 1867 (Hemiptera: Heteroptera: Cydnidae: Sehirinae). Entomologica Americana. 122 (1 - 2): 42 - 46." title="The taxonomic status of Sehirus aeneus Walker, 1867 (Hemiptera: Heteroptera: Cydnidae: Sehirinae)" type="journal article" year="2016">Aukema &amp; Constant 2016</bibRefCitation>
).
</paragraph>
<paragraph id="B04A36A0FFD50E7BFCF9FEE6FA8BFE98" blockId="39.[844,1467,194,1868]" box="[844,1290,347,368]" pageId="39" pageNumber="40">
<taxonomicName id="77F54D23FFD50E7BFCF9FEE6FBC7FE98" baseAuthorityName="Horváth" baseAuthorityYear="1899" box="[844,1094,347,368]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="8281EAB2FFD50E7BFCF9FEE6FBC7FE98" box="[844,1094,347,368]" italics="true" pageId="39" pageNumber="40">Canthophorus fuscipennis</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFD50E7BFBE0FEE6FB5EFE98" author="Fuente JADL" box="[1109,1247,347,368]" journalOrPublisher="Boletin de la Real Sociedad Espanola de Historia Natural, seccion Biologica." pageId="39" pageNumber="40" pagination="33 - 78" part="70" refId="ref39716" refString="Fuente JADL. 1972. Revision de los Pentatomidos ibericos. Familia Cydnidae Billberg, 1820. Boletin de la Real Sociedad Espanola de Historia Natural, seccion Biologica. 70: 33 - 78. Spanish." title="Revision de los Pentatomidos ibericos. Familia Cydnidae Billberg, 1820" type="journal article" year="1972">Fuente (1972)</bibRefCitation>
: 64.
</paragraph>
<paragraph id="B04A36A0FFD50E7BFCF9FEC8FAE9FE62" blockId="39.[844,1467,194,1868]" box="[844,1384,373,394]" pageId="39" pageNumber="40">
<taxonomicName id="77F54D23FFD50E7BFCF9FEC8FB9FFE62" authorityName="Gapon" authorityYear="2018" baseAuthorityName="Walker" baseAuthorityYear="1867" box="[844,1054,373,394]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="aeneus">
<emphasis id="8281EAB2FFD50E7BFCF9FEC8FB9FFE62" box="[844,1054,373,394]" italics="true" pageId="39" pageNumber="40">Canthophorus aeneus</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFD50E7BFB99FEC8FABCFE62" author="Aukema B &amp; Constant J." box="[1068,1341,373,394]" journalOrPublisher="Entomologica Americana." pageId="39" pageNumber="40" pagination="42 - 46" part="122" refId="ref39601" refString="Aukema B, Constant J. 2016. The taxonomic status of Sehirus aeneus Walker, 1867 (Hemiptera: Heteroptera: Cydnidae: Sehirinae). Entomologica Americana. 122 (1 - 2): 42 - 46." title="The taxonomic status of Sehirus aeneus Walker, 1867 (Hemiptera: Heteroptera: Cydnidae: Sehirinae)" type="journal article" year="2016">Aukema &amp; Constant (2016)</bibRefCitation>
: 42.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD50E7BFCF9FE6EFBA3FE02" box="[844,1058,467,490]" pageId="39" pageNumber="40" type="multiple">
<paragraph id="B04A36A0FFD50E7BFCF9FE6EFBA3FE02" blockId="39.[844,1467,194,1868]" box="[844,1058,467,490]" pageId="39" pageNumber="40">
<heading id="EB0281CCFFD50E7BFCF9FE6EFBA3FE02" bold="true" box="[844,1058,467,490]" fontSize="10" level="2" pageId="39" pageNumber="40" reason="6">
<emphasis id="8281EAB2FFD50E7BFCF9FE6EFBA3FE02" bold="true" box="[844,1058,467,490]" pageId="39" pageNumber="40">Material examined</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD50E7BFCF9FE4AFA2CFD13" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="B04A36A0FFD50E7BFCF9FE4AFA2CFD13" blockId="39.[844,1467,194,1868]" pageId="39" pageNumber="40">
<materialsCitation id="009D3CFDFFD50E7BFCF9FE4AFC24FDA9" collectionCode="ZISP, MNHN" country="France" location="Banyuls-sur-Mer, Pyrenees-Orientales" municipality="Languedoc-Roussillon-Midi-Pyrenees" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD50E7BFCF9FE4AFC13FDE4" bold="true" box="[844,914,503,524]" pageId="39" pageNumber="40">
<collectingCountry id="C8E27630FFD50E7BFCF9FE4AFC13FDE4" box="[844,914,503,524]" name="France" pageId="39" pageNumber="40" value="France">France</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD50E7BFC10FE4AFA84FDE4" box="[933,1285,503,524]" italics="true" pageId="39" pageNumber="40">
<collectingMunicipality id="502EACDAFFD50E7BFC10FE4AFA84FDE4" box="[933,1285,503,524]" pageId="39" pageNumber="40">Languedoc-Roussillon-Midi-Pyrénées</collectingMunicipality>
</emphasis>
, “Banyuls Pyrén. or.” [
<location id="B52A607BFFD50E7BFC30FDAFFB7BFDCF" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFC26FF7FFA2CFD13:B52A607BFFD50E7BFC30FDAFFB7BFDCF" box="[901,1274,530,551]" country="France" municipality="Languedoc-Roussillon-Midi-Pyrenees" name="Banyuls-sur-Mer, Pyrenees-Orientales" pageId="39" pageNumber="40">Banyuls-sur-Mer, Pyrénées-Orientales</location>
], 1J (
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),
<specimenCount id="A6F3FD29FFD50E7BFA29FDAFFA38FDCF" box="[1436,1465,529,551]" count="1" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD50E7BFCE7FD91FC1EFDA9" box="[850,927,556,577]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="39" pageNumber="40">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFC0DFD91FACDFDB4" collectingDate="1980-06-30" collectionCode="ZISP" collectorName="J. Balazuc" country="France" location="Ile du Petit Maletras" municipality="Saint-Just-d' Ardeche" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Auvergne-Rhone-Alpes">
<emphasis id="8281EAB2FFD50E7BFC0DFD91FB0FFDA9" box="[952,1166,556,577]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="7231F842FFD50E7BFC0DFD91FB0FFDA9" box="[952,1166,556,577]" pageId="39" pageNumber="40">Auvergne-Rhône-Alpes</collectingRegion>
</emphasis>
,
<collectingMunicipality id="502EACDAFFD50E7BFB17FD91FAE8FDA9" box="[1186,1385,556,578]" pageId="39" pageNumber="40">Saint-Just-d’ Ardèche</collectingMunicipality>
,
<location id="B52A607BFFD50E7BFACEFD91FC54FDB4" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFC26FF7FFA2CFD13:B52A607BFFD50E7BFACEFD91FC54FDB4" country="France" municipality="Saint-Just-d' Ardeche" name="Ile du Petit Maletras" pageId="39" pageNumber="40" stateProvince="Auvergne-Rhone-Alpes">Île du Petit Maletras</location>
,
<date id="C44B1060FFD50E7BFC51FDFAFBCFFDB4" box="[996,1102,583,604]" pageId="39" pageNumber="40" value="1980-06-30">
<collectingDate id="D40FE988FFD50E7BFC51FDFAFBCFFDB4" box="[996,1102,583,604]" pageId="39" pageNumber="40" value="1980-06-30">30.VI.1980</collectingDate>
</date>
(
<collectorName id="1D005376FFD50E7BFBD7FDFAFB4AFDB4" box="[1122,1227,583,604]" pageId="39" pageNumber="40">J. Balazuc</collectorName>
),
<specimenCount id="A6F3FD29FFD50E7BFB55FDFAFB7FFDB4" box="[1248,1278,582,604]" count="1" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD50E7BFAA4FDFAFAC4FDB4" box="[1297,1349,583,604]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="39" pageNumber="40">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFAEEFDFAFAD5FD9E" collectingDate="1980-06-12" collectionCode="AMC" country="France" location="Gemenos" municipality="Provence-Alpes-Cote d' Azur" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFD50E7BFAEEFDFAFC78FD9E" italics="true" pageId="39" pageNumber="40">
<collectingMunicipality id="502EACDAFFD50E7BFAEEFDFAFC78FD9E" pageId="39" pageNumber="40">Provence-Alpes-Côte d’ Azur</collectingMunicipality>
</emphasis>
,
<location id="B52A607BFFD50E7BFBBDFDDCFBE2FD9E" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFC26FF7FFA2CFD13:B52A607BFFD50E7BFBBDFDDCFBE2FD9E" box="[1032,1123,609,630]" country="France" municipality="Provence-Alpes-Cote d' Azur" name="Gemenos" pageId="39" pageNumber="40">Gémenos</location>
,
<date id="C44B1060FFD50E7BFBDBFDDCFB5BFD9E" box="[1134,1242,609,630]" pageId="39" pageNumber="40" value="1980-06-12">
<collectingDate id="D40FE988FFD50E7BFBDBFDDCFB5BFD9E" box="[1134,1242,609,630]" pageId="39" pageNumber="40" value="1980-06-12">12.VI.1980</collectingDate>
</date>
,
<specimenCount id="A6F3FD29FFD50E7BFB53FDDCFA85FD9F" box="[1254,1284,609,631]" count="1" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD50E7BFAA6FDDCFACCFD9E" box="[1299,1357,609,630]" collectionName="AMC" pageId="39" pageNumber="40">AMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFAD5FDDCFA37FD79" collectingDate="1989-06-20" collectionCode="ZISP" collectorName="J. Coffin" country="France" county="Courthezon" location="Les Palluds" municipality="Vaucluse" pageId="39" pageNumber="40" specimenCount="1">
<collectingMunicipality id="502EACDAFFD50E7BFAD5FDDCFA37FD9E" box="[1376,1462,609,630]" pageId="39" pageNumber="40">Vaucluse</collectingMunicipality>
,
<collectingCounty id="592B4E2CFFD50E7BFCF9FDC1FC3AFD79" box="[844,955,636,657]" pageId="39" pageNumber="40">Courthezon</collectingCounty>
,
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<figureCitation id="28CE2A25FFD50E7BFC3DFCE1FBAEFC9B" box="[904,1071,860,883]" captionStart="Figure 2" captionStartId="6.[158,225,1725,1746]" captionTargetBox="[209,1426,182,1704]" captionTargetId="figure-102@6.[209,1426,182,1704]" captionTargetPageId="6" captionText="Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927021" httpUri="https://zenodo.org/record/14927021/files/figure.png" pageId="39" pageNumber="40">
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<figureCitation id="28CE2A25FFD50E7BFB8BFCE1FB0AFC9B" box="[1086,1163,860,883]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="39" pageNumber="40">
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19
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[
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:
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; lost].
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<emphasis id="8281EAB2FFD50E7BFCF9FC1DFC55FC5D" box="[844,980,928,949]" italics="true" pageId="39" pageNumber="40">Canthophorus</emphasis>
(
<emphasis id="8281EAB2FFD50E7BFC5AFC1DFBF6FC5D" box="[1007,1143,928,949]" italics="true" pageId="39" pageNumber="40">Canthophorus</emphasis>
)
<emphasis id="8281EAB2FFD50E7BFB27FC1DFB73FC5D" box="[1170,1266,928,949]" italics="true" pageId="39" pageNumber="40">maculipes</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFD50E7BFAB9FC1DFC4BFC38" author="Mulsant E &amp; Rey C." journalOrPublisher="Paris: Savy &amp; Deyrolle" pageId="39" pageNumber="40" refId="ref40479" refString="Mulsant E, Rey C. 1866. Histoire naturelle des punaises de France. Pentatomides. Paris: Savy &amp; Deyrolle." title="Histoire naturelle des punaises de France. Pentatomides" type="book" year="1866">Mulsant &amp; Rey 1866: 65</bibRefCitation>
.
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<emphasis id="8281EAB2FFD50E7BFCF9FBA5FBA3FBC7" bold="true" box="[844,1058,1048,1071]" pageId="39" pageNumber="40">Material examined</emphasis>
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:
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,
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,
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, 1J (
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); 1898 (Ern. André), 1J,
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(
<collectionCode id="D6E4AE65FFD50E7BFB52FBEAFAB3FB84" box="[1255,1330,1111,1132]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="39" pageNumber="40">MNHN</collectionCode>
),
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(
<collectionCode id="D6E4AE65FFD50E7BFAC9FBEAFA2EFB84" box="[1404,1455,1111,1132]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="39" pageNumber="40">ZISP</collectionCode>
)
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;
<materialsCitation id="009D3CFDFFD50E7BFCF9FBCFFB6FFB6F" box="[844,1262,1137,1159]" collectionCode="ZISP" country="France" location="d' Oisans" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Auvergne-Rhone-Alpes">
<emphasis id="8281EAB2FFD50E7BFCF9FBCFFB9CFB6F" box="[844,1053,1138,1159]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="7231F842FFD50E7BFCF9FBCFFB9CFB6F" box="[844,1053,1138,1159]" pageId="39" pageNumber="40">Auvergne-Rhône-Alpes</collectingRegion>
</emphasis>
,
<location id="B52A607BFFD50E7BFB9FFBCFFBFCFB6F" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFCE6FC81FB6DFACE:B52A607BFFD50E7BFB9FFBCFFBFCFB6F" box="[1066,1149,1137,1159]" country="France" name="d' Oisans" pageId="39" pageNumber="40" stateProvince="Auvergne-Rhone-Alpes">d’ Oisans</location>
, 1J (
<collectionCode id="D6E4AE65FFD50E7BFB01FBCFFB66FB6F" box="[1204,1255,1138,1159]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="39" pageNumber="40">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFB4CFBCFFBADFB54" collectingDate="1984-08-07" collectionCode="ZISP" collectorName="A. Matocq" country="France" elevation="2109" location="Vars" municipality="Alpes-de-Haute-Provence" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Provence-Alpes-Cote d' Azur">
<emphasis id="8281EAB2FFD50E7BFB4CFBCFFC08FB49" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="7231F842FFD50E7BFB4CFBCFFC08FB49" pageId="39" pageNumber="40">Provence-Alpes-Côte d’ Azur</collectingRegion>
</emphasis>
:
<collectingMunicipality id="502EACDAFFD50E7BFC20FB31FB03FB49" box="[917,1154,1164,1185]" pageId="39" pageNumber="40">Alpes-de-Haute-Provence</collectingMunicipality>
, col de
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,
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<collectingDate id="D40FE988FFD50E7BFAFBFB31FA3BFB49" box="[1358,1466,1164,1185]" pageId="39" pageNumber="40" value="1984-08-07">7.VIII.1984</collectingDate>
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(
<collectorName id="1D005376FFD50E7BFCE6FB1AFC39FB54" box="[851,952,1191,1212]" pageId="39" pageNumber="40">A. Matocq</collectorName>
), 1J (
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)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFB80FB1AFAD3FB54" box="[1077,1362,1190,1212]" collectionCode="MNHN" country="France" location="Draguignan" municipality="Var" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Provence-alpes-cote D' Azur">
<collectingMunicipality id="502EACDAFFD50E7BFB80FB1AFBD7FB54" box="[1077,1110,1191,1212]" pageId="39" pageNumber="40">Var</collectingMunicipality>
,
<location id="B52A607BFFD50E7BFBD5FB1AFB4FFB54" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFCE6FC81FB6DFACE:B52A607BFFD50E7BFBD5FB1AFB4FFB54" box="[1120,1230,1191,1212]" country="France" municipality="Var" name="Draguignan" pageId="39" pageNumber="40" stateProvince="Provence-alpes-cote D' Azur">Draguignan</location>
, 1J (
<collectionCode id="D6E4AE65FFD50E7BFAB4FB1AFACDFB54" box="[1281,1356,1191,1212]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="39" pageNumber="40">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFAE9FB1AFB23FB19" collectingDate="1989-06-12" collectionCode="AMC" collectorName="J. Hamon" country="France" county="Leucate" location="La Gardiole" municipality="Aude" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Occitanie">
<emphasis id="8281EAB2FFD50E7BFAE9FB1AFA35FB54" box="[1372,1460,1191,1212]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="7231F842FFD50E7BFAE9FB1AFA35FB54" box="[1372,1460,1191,1212]" pageId="39" pageNumber="40">Occitanie</collectingRegion>
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,
<collectingMunicipality id="502EACDAFFD50E7BFCF9FB7CFC01FB3E" box="[844,896,1217,1238]" pageId="39" pageNumber="40">Aude</collectingMunicipality>
,
<collectingCounty id="592B4E2CFFD50E7BFC3FFB7CFC55FB3E" box="[906,980,1217,1238]" pageId="39" pageNumber="40">Leucate</collectingCounty>
, vigne en friche
<location id="B52A607BFFD50E7BFBDBFB7CFB5DFB3E" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFCE6FC81FB6DFACE:B52A607BFFD50E7BFBDBFB7CFB5DFB3E" box="[1134,1244,1217,1238]" country="France" county="Leucate" municipality="Aude" name="La Gardiole" pageId="39" pageNumber="40" stateProvince="Occitanie">La Gardiole</location>
piège jaune-orange,
<date id="C44B1060FFD50E7BFA29FB7CFC16FB19" pageId="39" pageNumber="40" value="1989-06-12">
<collectingDate id="D40FE988FFD50E7BFA29FB7CFC16FB19" pageId="39" pageNumber="40" value="1989-06-12">12. VI.1989</collectingDate>
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(
<collectorName id="1D005376FFD50E7BFC10FB61FC7FFB19" box="[933,1022,1244,1265]" pageId="39" pageNumber="40">J. Hamon</collectorName>
rec.),
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(
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)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFD50E7BFB19FB61FB69FACE" collectingDate="1990-06-06" collectionCode="ZISP" collectorName="J. Pericart, A. Matocq" country="Spain" location="Torres de Albarracin" municipality="Teruel" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Aragon">
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<collectingCountry id="C8E27630FFD50E7BFB19FB61FB65FB19" box="[1196,1252,1244,1265]" name="Spain" pageId="39" pageNumber="40" value="Spain">Spain</collectingCountry>
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:
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<collectingRegion id="7231F842FFD50E7BFB45FB61FAB3FB19" box="[1264,1330,1244,1265]" country="Spain" name="Aragon" pageId="39" pageNumber="40">Aragon</collectingRegion>
/
<collectingMunicipality id="502EACDAFFD50E7BFA89FB61FAF2FB19" box="[1340,1395,1244,1265]" pageId="39" pageNumber="40">Teruel</collectingMunicipality>
</emphasis>
,
<location id="B52A607BFFD50E7BFA35FB61FC4AFAE4" LSID="urn:lsid:plazi:treatment:385C87B6FFD50E7BFCE6FC81FB6DFACE:B52A607BFFD50E7BFA35FB61FC4AFAE4" country="Spain" municipality="Teruel" name="Torres de Albarracin" pageId="39" pageNumber="40" stateProvince="Aragon">Torres de Albarracin</location>
,
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<quantity id="770D9B45FFD50E7BFC61FB4AFB85FAE4" box="[980,1028,1271,1292]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="39" pageNumber="40" unit="km" value="4.0">4 km</quantity>
au nord sur rivière Guadalaviar
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,
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<collectingDate id="D40FE988FFD50E7BFA84FB4AFA0FFAE4" box="[1329,1422,1271,1292]" pageId="39" pageNumber="40" value="1990-06-06">6.VI.1990</collectingDate>
</date>
(Ph. Magnien,
<collectorName id="1D005376FFD50E7BFC1CFAACFBF5FACE" box="[937,1140,1297,1318]" pageId="39" pageNumber="40">J. Péricart, A. Matocq</collectorName>
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(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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451
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<heading id="EB0281CCFFD60E78FCF9FADEFAC4FA72" bold="true" fontSize="10" level="2" pageId="36" pageNumber="37" reason="4">
<emphasis id="8281EAB2FFD60E78FCF9FADEFAC4FA72" bold="true" pageId="36" pageNumber="37">
<emphasis id="8281EAB2FFD60E78FCF9FADEFA3BFA92" bold="true" box="[844,1466,1379,1402]" pageId="36" pageNumber="37">
<taxonomicName id="77F54D23FFD60E78FCF9FADEFADCFA92" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Kiritshenko" baseAuthorityYear="1964" box="[844,1373,1379,1402]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="mixtus" status="stat. nov." subSpecies="hissaricus">
<emphasis id="8281EAB2FFD60E78FCF9FADEFB30FA92" bold="true" box="[844,1201,1379,1402]" italics="true" pageId="36" pageNumber="37">Canthophorus mixtus hissaricus</emphasis>
<bibRefCitation id="D4644B51FFD60E78FB0DFADEFADCFA92" author="Asanova RB" box="[1208,1373,1379,1402]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="36" pageNumber="37" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="99B257C9FFD60E78FAD8FADEFA3BFA92" box="[1389,1466,1379,1402]" pageId="36" pageNumber="37">n. stat.</taxonomicNameLabel>
</emphasis>
<emphasis id="8281EAB2FFD60E78FC74FA3EFAC4FA72" bold="true" box="[961,1349,1411,1434]" pageId="36" pageNumber="37">
(
<figureCitation id="28CE2A25FFD60E78FC7CFA3EFBC9FA72" box="[969,1096,1411,1434]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="36" pageNumber="37">
Figures 3
<collectionCode id="D6E4AE65FFD60E78FB86FA3EFBC9FA72" box="[1075,1096,1411,1434]" pageId="36" pageNumber="37">K</collectionCode>
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,
<figureCitation id="28CE2A25FFD60E78FBEDFA3EFB22FA72" box="[1112,1187,1411,1434]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="36" pageNumber="37">
7
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, 7
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,
<figureCitation id="28CE2A25FFD60E78FB06FA3EFA81FA72" box="[1203,1280,1411,1434]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="36" pageNumber="37">
8
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, 8
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</figureCitation>
,
<figureCitation id="28CE2A25FFD60E78FABAFA3EFABDFA72" box="[1295,1340,1411,1434]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="36" pageNumber="37">
11
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)
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD60E78FCF9FA11FA89FA1E" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="B04A36A0FFD60E78FCF9FA11FB93FA34" blockId="36.[844,1466,193,1865]" pageId="36" pageNumber="37">
<emphasis id="8281EAB2FFD60E78FCF9FA11FBC4FA29" box="[844,1093,1452,1473]" italics="true" pageId="36" pageNumber="37">
Canthophorns
<taxonomicName id="77F54D23FFD60E78FC50FA11FAC4FA29" authority="Kiritshenko 1951: 189" authorityName="Kiritshenko" authorityPageNumber="189" authorityYear="1951" box="[997,1349,1452,1473]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hissaricus">
hissaricus
<bibRefCitation id="D4644B51FFD60E78FBE3FA11FAC4FA29" author="Kiritshenko AN" box="[1110,1349,1452,1473]" editor="Pavlovskij E N &amp; Shadin V I" journalOrPublisher="Moskva-Leningrad: Akademiya Nauk SSSR" pageId="36" pageNumber="37" pagination="181 - 197" refId="ref40097" refString="Kiritshenko AN. 1951. True bugs and cicads. In: Pavlovskij E N, and Shadin V I, editors. Kondara Gorge (Experience of a biological monograph). Moskva-Leningrad: Akademiya Nauk SSSR; p. 181 - 197. Russian." title="True bugs and cicads" type="book chapter" year="1951">Kiritshenko 1951: 189</bibRefCitation>
</taxonomicName>
</emphasis>
;
<bibRefCitation id="D4644B51FFD60E78FAEFFA11FA38FA29" author="Kiritshenko AN" box="[1370,1465,1452,1473]" journalOrPublisher="Dushanbe: Institute of Zoology and Parasitology. Russian" pageId="36" pageNumber="37" refId="ref40144" refString="Kiritshenko AN. 1964. Bugs (Hemiptera-Heteroptera). Dushanbe: Institute of Zoology and Parasitology. Russian." title="Bugs (Hemiptera-Heteroptera)" type="book" year="1964">1964: 61</bibRefCitation>
[nomen nudum].
</paragraph>
<paragraph id="B04A36A0FFD60E78FCF9FA5CFA89FA1E" blockId="36.[844,1466,193,1865]" box="[844,1288,1505,1526]" pageId="36" pageNumber="37">
<taxonomicName id="77F54D23FFD60E78FCF9FA5CFA85FA1E" authority="Asanova 1964: 141" authorityName="Asanova" authorityPageNumber="141" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[844,1284,1505,1526]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD60E78FCF9FA5CFBBDFA1E" box="[844,1084,1505,1526]" italics="true" pageId="36" pageNumber="37">Canthophorus hissaricus</emphasis>
<bibRefCitation id="D4644B51FFD60E78FBF1FA5CFA85FA1E" author="Asanova RB" box="[1092,1284,1505,1526]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="36" pageNumber="37" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova 1964: 141</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD60E78FCF9F986FBA3F9BA" box="[844,1058,1595,1618]" pageId="36" pageNumber="37" type="multiple">
<paragraph id="B04A36A0FFD60E78FCF9F986FBA3F9BA" blockId="36.[844,1466,193,1865]" box="[844,1058,1595,1618]" pageId="36" pageNumber="37">
<heading id="EB0281CCFFD60E78FCF9F986FBA3F9BA" bold="true" box="[844,1058,1595,1618]" fontSize="10" level="2" pageId="36" pageNumber="37" reason="6">
<emphasis id="8281EAB2FFD60E78FCF9F986FBA3F9BA" bold="true" box="[844,1058,1595,1618]" pageId="36" pageNumber="37">Material examined</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD60E78FCF9F9E2FAFAF8A1" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="B04A36A0FFD60E78FCF9F9E2FA3BF941" blockId="36.[844,1466,193,1865]" pageId="36" pageNumber="37">
<materialsCitation id="009D3CFDFFD60E78FCF9F9E2FA36F941" collectingDate="1947-09-10" collectionCode="ZISP" country="Tajikistan" elevation="2600" latitude="38.98461" location="Gissar Range" longLatPrecision="1" longitude="68.79158" municipality="Ruydasht" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Varzob Distr" typeStatus="holotype">
<typeStatus id="6F4E8802FFD60E78FCF9F9E2FC29F99C" box="[844,936,1631,1652]" pageId="36" pageNumber="37">Holotype</typeStatus>
:
<emphasis id="8281EAB2FFD60E78FC09F9E2FBA3F99C" bold="true" box="[956,1058,1631,1652]" pageId="36" pageNumber="37">
<collectingCountry id="C8E27630FFD60E78FC09F9E2FBA3F99C" box="[956,1058,1631,1652]" name="Tajikistan" pageId="36" pageNumber="37" value="Tajikistan">Tajikistan</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD60E78FB8FF9E2FB3CF99C" box="[1082,1213,1631,1652]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="7231F842FFD60E78FB8FF9E2FB3CF99C" box="[1082,1213,1631,1652]" pageId="36" pageNumber="37">Varzob Distr</collectingRegion>
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., urochishche
<collectingMunicipality id="502EACDAFFD60E78FAEAF9E2FA38F99C" box="[1375,1465,1631,1652]" pageId="36" pageNumber="37">Ruydasht</collectingMunicipality>
[
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<quantity id="770D9B45FFD60E78FCE4F9C7FC12F967" box="[849,915,1658,1679]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="37" unit="km" value="40.0">40 km</quantity>
N Dushanbe
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,
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38°59
<emphasis id="8281EAB2FFD60E78FBD5F9C4FBE4F967" bold="true" box="[1120,1125,1657,1679]" italics="true" pageId="36" pageNumber="37">ʹ</emphasis>
04.6
<emphasis id="8281EAB2FFD60E78FB3AF9C4FB19F967" bold="true" box="[1167,1176,1657,1679]" italics="true" pageId="36" pageNumber="37">ʺ</emphasis>
N
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<geoCoordinate id="D5C15067FFD60E78FB06F9C7FAB7F967" box="[1203,1334,1657,1679]" degrees="68" direction="east" minutes="47" orientation="longitude" pageId="36" pageNumber="37" precision="1" seconds="29.7" value="68.79158">
68°47
<emphasis id="8281EAB2FFD60E78FB59F9C4FB70F967" bold="true" box="[1260,1265,1657,1679]" italics="true" pageId="36" pageNumber="37">ʹ</emphasis>
29.7
<emphasis id="8281EAB2FFD60E78FAAEF9C4FAA5F967" bold="true" box="[1307,1316,1657,1679]" italics="true" pageId="36" pageNumber="37">ʺ</emphasis>
E
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], S slopes of
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,
<quantity id="770D9B45FFD60E78FC62F929FB9EF941" box="[983,1055,1684,1705]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="36" pageNumber="37" unit="m" value="2600.0">
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amsl,
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<collectingDate id="D40FE988FFD60E78FBEBF929FB4BF941" box="[1118,1226,1684,1705]" pageId="36" pageNumber="37" value="1947-09-10">10.IX.1947</collectingDate>
</date>
(Kiritshenko), J (
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)
</materialsCitation>
.
</paragraph>
<paragraph id="B04A36A0FFD60E78FCFBF979FA2BF931" blockId="36.[844,1466,193,1865]" box="[846,1450,1732,1754]" pageId="36" pageNumber="37">
Paratypes: The same date as for the holotype, 8J,
<specimenCount id="A6F3FD29FFD60E78FA88F979FADAF932" box="[1341,1371,1732,1754]" count="2" pageId="36" pageNumber="37" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFD60E78FADCF979FA1FF931" box="[1385,1438,1732,1753]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="36" pageNumber="37">ZISP</collectionCode>
).
</paragraph>
<paragraph id="B04A36A0FFD60E78FCC6F959FAFAF8A1" blockId="36.[844,1466,193,1865]" pageId="36" pageNumber="37">
<materialsCitation id="009D3CFDFFD60E78FCC6F959FAF6F8A1" collectingDate="1964-09-01" collectionCode="ZISP" collectorName="V. F. Zaitzev" country="Tajikistan" county="Khozretishi Range" elevation="2500" location="Iokundzh Riv." pageId="36" pageNumber="37" specimenCount="1" stateProvince="Gorno-Badakhshan Autonomous Region">
Additional material.
<emphasis id="8281EAB2FFD60E78FBDAF959FB54F911" bold="true" box="[1135,1237,1764,1785]" pageId="36" pageNumber="37">
<collectingCountry id="C8E27630FFD60E78FBDAF959FB54F911" box="[1135,1237,1764,1785]" name="Tajikistan" pageId="36" pageNumber="37" value="Tajikistan">Tajikistan</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFD60E78FB48F959FB97F8FC" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="7231F842FFD60E78FB48F959FB97F8FC" pageId="36" pageNumber="37">Gorno-Badakhshan Autonomous Region</collectingRegion>
</emphasis>
,
<collectingCounty id="592B4E2CFFD60E78FB99F942FB68F8FC" box="[1068,1257,1791,1812]" pageId="36" pageNumber="37">Khozretishi Range</collectingCounty>
, middle reaches of
<location id="B52A607BFFD60E78FCF9F8A4FC66F8C6" LSID="urn:lsid:plazi:treatment:385C87B6FFD60E7AFCF9FADEFD6AFEE8:B52A607BFFD60E78FCF9F8A4FC66F8C6" box="[844,999,1817,1838]" country="Tajikistan" county="Khozretishi Range" name="Iokundzh Riv." pageId="36" pageNumber="37" stateProvince="Gorno-Badakhshan Autonomous Region">Iokundzh Riv.</location>
,
<locationDeviation id="A1F82F82FFD60E78FBB5F8A4FA34F8C6" box="[1024,1461,1817,1839]" pageId="36" pageNumber="37">
<quantity id="770D9B45FFD60E78FBB5F8A4FBCCF8C6" box="[1024,1101,1817,1838]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="36" pageNumber="37" unit="km" value="75.0">75 km</quantity>
NE “Mushtabad” [Muminabad]
</locationDeviation>
,
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<elevation id="3BD8D193FFD60E78FCF9F889FC50F8A1" box="[844,977,1843,1865]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="36" pageNumber="37" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">2000–3000 m</elevation>
</quantity>
amsl,
<date id="C44B1060FFD60E78FBA1F889FBF5F8A1" box="[1044,1140,1844,1865]" pageId="36" pageNumber="37" value="1964-09-01">
<collectingDate id="D40FE988FFD60E78FBA1F889FBF5F8A1" box="[1044,1140,1844,1865]" pageId="36" pageNumber="37" value="1964-09-01">1.IX.1964</collectingDate>
</date>
(
<collectorName id="1D005376FFD60E78FB36F889FB7DF8A1" box="[1155,1276,1844,1865]" pageId="36" pageNumber="37">V.F. Zaitzev</collectorName>
), 1J (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD70E79FF2BFF7CFB13FCDE" pageId="37" pageNumber="38" type="discussion">
<paragraph id="B04A36A0FFD70E79FF2BFF7CFD89FF30" blockId="37.[158,781,193,1314]" box="[158,520,193,216]" pageId="37" pageNumber="38">
<heading id="EB0281CCFFD70E79FF2BFF7CFD89FF30" bold="true" box="[158,520,193,216]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="6">
<emphasis id="8281EAB2FFD70E79FF2BFF7CFD89FF30" bold="true" box="[158,520,193,216]" pageId="37" pageNumber="38">Change in the taxonomic status</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD70E79FF2BFF54FD58FC56" blockId="37.[158,781,193,1314]" pageId="37" pageNumber="38">
<taxonomicName id="77F54D23FFD70E79FF2BFF54FEAAFEE8" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[158,299,233,256]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FF2BFF54FEAAFEE8" box="[158,299,233,256]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
was described on the basis of the two characters distinguishing this species from the nearest
<taxonomicName id="77F54D23FFD70E79FD09FEB4FF3CFEA8" authorityName="Asanova" authorityYear="1964" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FD09FEB4FF3CFEA8" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
, namely the smaller body size and the more strongly curved spicule at the dorsolateral lobes of conjunctiva (
<figureCitation id="28CE2A25FFD70E79FF12FED4FE9BFE68" box="[167,282,361,384]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
<collectionCode id="D6E4AE65FFD70E79FEB2FED4FE9BFE68" box="[263,282,361,384]" pageId="37" pageNumber="38">R</collectionCode>
</figureCitation>
). Body dimensions were given as follows. Males: body
<quantity id="770D9B45FFD70E79FE9BFE34FE37FE48" box="[302,438,392,416]" metricMagnitude="-1" metricUnit="m" metricValue="1.1049" metricValueMax="1.1176" metricValueMin="1.0921999999999998" pageId="37" pageNumber="38" unit="in" value="4.35" valueMax="4.4" valueMin="4.3">4.30–4.40 in</quantity>
length, pronotum
<quantity id="770D9B45FFD70E79FD36FE34FC8AFE48" box="[643,779,392,416]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="37" pageNumber="38" unit="in" value="2.4" valueMax="2.5" valueMin="2.3">2.30–2.50 in</quantity>
width, abdomen
<quantity id="770D9B45FFD70E79FEE7FE14FE58FE28" box="[338,473,424,448]" metricMagnitude="-2" metricUnit="m" metricValue="6.731" metricValueMax="7.111999999999999" metricValueMin="6.35" pageId="37" pageNumber="38" unit="in" value="2.65" valueMax="2.8" valueMin="2.5">2.50–2.80 in</quantity>
width. Females: body 5.80– 6.00 in length, pronotum 3.00–
<quantity id="770D9B45FFD70E79FE4AFE74FDCFFE08" box="[511,590,457,480]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="37" pageNumber="38" unit="in" value="3.2">3.20 in</quantity>
width, abdomen
<quantity id="770D9B45FFD70E79FF2BFE55FEAAFE17" box="[158,299,488,512]" metricMagnitude="-2" metricUnit="m" metricValue="9.144" metricValueMax="9.398" metricValueMin="8.89" pageId="37" pageNumber="38" unit="in" value="3.6" valueMax="3.7" valueMin="3.5">3.50–3.70 in</quantity>
width. In the re-examination of the
<typeStatus id="6F4E8802FFD70E79FD6BFE55FC8AFE17" box="[734,779,488,511]" pageId="37" pageNumber="38">type</typeStatus>
series, the following measurements were obtained. Males: body
<quantity id="770D9B45FFD70E79FE89FD95FE4DFDD7" box="[316,460,552,576]" metricMagnitude="-1" metricUnit="m" metricValue="1.2255500000000001" metricValueMax="1.3335000000000001" metricValueMin="1.1176000000000001" pageId="37" pageNumber="38" unit="in" value="4.825" valueMax="5.25" valueMin="4.4">4.40–5.25 in</quantity>
length, body
<quantity id="770D9B45FFD70E79FDC9FD95FC8AFDD7" box="[636,779,552,576]" metricMagnitude="-2" metricUnit="m" metricValue="7.5565" metricValueMax="8.128" metricValueMin="6.985" pageId="37" pageNumber="38" unit="in" value="2.975" valueMax="3.2" valueMin="2.75">2.75–3.20 in</quantity>
width, pronotum
<quantity id="770D9B45FFD70E79FED8FDF5FE7CFDB7" box="[365,509,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="6.6675" metricValueMax="6.985" metricValueMin="6.35" pageId="37" pageNumber="38" unit="in" value="2.625" valueMax="2.75" valueMin="2.5">2.50–2.75 in</quantity>
width. Females: body 5.88–6.00 in length, body 3.65–4.00 in width, pronotum
<quantity id="770D9B45FFD70E79FF2BFD35FEA6FD77" box="[158,295,647,671]" metricMagnitude="-2" metricUnit="m" metricValue="8.4455" metricValueMax="8.636" metricValueMin="8.255" pageId="37" pageNumber="38" unit="in" value="3.325" valueMax="3.4" valueMin="3.25">3.25–3.40 in</quantity>
width. Minimal values of these dimensions do not overlap with corresponding minimal values for males of
<taxonomicName id="77F54D23FFD70E79FEBCFD75FEF3FD37" authorityName="Asanova" authorityYear="1964" box="[265,370,712,735]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FEBCFD75FEF3FD37" box="[265,370,712,735]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
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but differ very slightly (by
<quantity id="770D9B45FFD70E79FD03FD75FC8DFD37" box="[694,780,712,735]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="mm" value="0.1">0.1 mm</quantity>
for body length and width and
<quantity id="770D9B45FFD70E79FDBCFD55FDECFD17" box="[521,621,744,767]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="38" unit="mm" value="0.05">0.05 mm</quantity>
for pronotum width), so they can be considered as ultimate values in the variability of the latter species (
<figureCitation id="28CE2A25FFD70E79FDA3FC9AFD04FCD6" box="[534,645,807,830]" captionStart="Figure 17" captionStartId="37.[158,225,1830,1851]" captionTargetBox="[211,1424,1405,1787]" captionTargetId="figure-667@37.[205,1431,1396,1793]" captionTargetPageId="37" captionText="Figure 17. Comparison of body measurements in Canthophorus mixtus mixtus Asanova, 1964 (1, 2) and C. mixtus hissaricus Asanova, 1964 n. stat. (3). 2, specimens from Trans-Ili and Alatau Khozretishi Range. BL – body length; BW – body width; PW – pronotum width." figureDoi="http://doi.org/10.5281/zenodo.14927051" httpUri="https://zenodo.org/record/14927051/files/figure.png" pageId="37" pageNumber="38">Figure 17</figureCitation>
), especially since only
<specimenCount id="A6F3FD29FFD70E79FEADFCFAFEF8FCB6" box="[280,377,839,862]" count="10" pageId="37" pageNumber="38" type="male">10 males</specimenCount>
of
<taxonomicName id="77F54D23FFD70E79FE16FCFAFDB0FCB6" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[419,561,839,862]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FE16FCFAFDB0FCB6" box="[419,561,839,862]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
are known. Ranges of the body size in females of
<taxonomicName id="77F54D23FFD70E79FE68FCDAFDE6FC96" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[477,615,871,894]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FE68FCDAFDE6FC96" box="[477,615,871,894]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
fall into ranges of
<taxonomicName id="77F54D23FFD70E79FF08FC3AFEA2FC76" authorityName="Asanova" authorityYear="1964" box="[189,291,903,926]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FF08FC3AFEA2FC76" box="[189,291,903,926]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
(
<figureCitation id="28CE2A25FFD70E79FE81FC3AFE1DFC76" box="[308,412,903,926]" captionStart="Figure 17" captionStartId="37.[158,225,1830,1851]" captionTargetBox="[211,1424,1405,1787]" captionTargetId="figure-667@37.[205,1431,1396,1793]" captionTargetPageId="37" captionText="Figure 17. Comparison of body measurements in Canthophorus mixtus mixtus Asanova, 1964 (1, 2) and C. mixtus hissaricus Asanova, 1964 n. stat. (3). 2, specimens from Trans-Ili and Alatau Khozretishi Range. BL – body length; BW – body width; PW – pronotum width." figureDoi="http://doi.org/10.5281/zenodo.14927051" httpUri="https://zenodo.org/record/14927051/files/figure.png" pageId="37" pageNumber="38">Figure 17</figureCitation>
), although specimens of the latter species with extreme small sizes are very infrequent.
</paragraph>
<paragraph id="B04A36A0FFD70E79FF73FC71FA25FD9F" blockId="37.[158,781,193,1314]" lastBlockId="37.[844,1466,194,1312]" pageId="37" pageNumber="38">
There are no significant structural differences between
<taxonomicName id="77F54D23FFD70E79FEBEFC51FE25FBEB" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[267,420,1004,1027]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FEBEFC51FE25FBEB" box="[267,420,1004,1027]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
and
<taxonomicName id="77F54D23FFD70E79FE46FC51FDE4FBEB" authorityName="Asanova" authorityYear="1964" box="[499,613,1004,1027]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FE46FC51FDE4FBEB" box="[499,613,1004,1027]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
. The sharply curved spicule at dorsolateral lobes of conjunctiva occurs, although rarely, in specimens of
<taxonomicName id="77F54D23FFD70E79FD2FFB91FC8DFBAB" authorityName="Asanova" authorityYear="1964" box="[666,780,1068,1091]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FD2FFB91FC8DFBAB" box="[666,780,1068,1091]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
from different parts of the natural range, and there is a gradual transitional row from the smoothly and slightly bent spicules to the sharply and strongly curved ones (
<figureCitation id="28CE2A25FFD70E79FF12FB16FEC3FB2A" box="[167,322,1195,1219]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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–7
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). In the
<specimenCount id="A6F3FD29FFD70E79FE1FFB16FDDBFB2A" box="[426,602,1195,1218]" count="3" pageId="37" pageNumber="38" type="generic">three specimens</specimenCount>
examined from the Khozretishi Range, the spicule is curved distinctly but to different degrees: in the male, corresponding to the dimensions of
<taxonomicName id="77F54D23FFD70E79FECFFAB6FD8CFACA" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[378,525,1291,1314]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FECFFAB6FD8CFACA" box="[378,525,1291,1314]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
(body
<quantity id="770D9B45FFD70E79FDDDFAB6FD3AFACA" box="[616,699,1291,1314]" metricMagnitude="-1" metricUnit="m" metricValue="1.1683999999999999" pageId="37" pageNumber="38" unit="in" value="4.6">4.60 in</quantity>
length and
<quantity id="770D9B45FFD70E79FCC9FF7FFC4BFF31" box="[892,970,194,217]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="37" pageNumber="38" unit="in" value="3.1">3.10 in</quantity>
width, pronotum
<quantity id="770D9B45FFD70E79FB21FF7FFB63FF31" box="[1172,1250,194,217]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" pageId="37" pageNumber="38" unit="in" value="2.7">2.70 in</quantity>
width), the spicule is strongly and sharply curved (
<figureCitation id="28CE2A25FFD70E79FB0CFF5FFAA6FF11" box="[1209,1319,226,249]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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</figureCitation>
); among the other and larger males of
<taxonomicName id="77F54D23FFD70E79FBC6FEBCFB5CFEF0" authorityName="Asanova" authorityYear="1964" box="[1139,1245,257,280]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FBC6FEBCFB5CFEF0" box="[1139,1245,257,280]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
, one (body
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length and
<quantity id="770D9B45FFD70E79FC79FE9CFB9DFED0" box="[972,1052,289,312]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="37" pageNumber="38" unit="in" value="3.4">3.40 in</quantity>
width, pronotum
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width) has the spicule curved less strongly, but rather sharply (
<figureCitation id="28CE2A25FFD70E79FCE0FEDCFC46FE90" box="[853,967,353,376]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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); in the third male (body
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length and
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width, pronotum
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width), the spicule is curved even more weakly and more smoothly (
<figureCitation id="28CE2A25FFD70E79FCE0FE7CFC46FE30" box="[853,967,449,472]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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). In the male of
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<emphasis id="8281EAB2FFD70E79FB27FE7CFB7CFE30" box="[1170,1277,449,472]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
from the Trans-Ili Alatau (body
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length and
<quantity id="770D9B45FFD70E79FABAFE5CFAE5FE10" box="[1295,1380,481,504]" metricMagnitude="-2" metricUnit="m" metricValue="8.509" pageId="37" pageNumber="38" unit="in" value="3.35">3.35 in</quantity>
width, pronotum
<quantity id="770D9B45FFD70E79FC0AFDBCFB8FFDF0" box="[959,1038,513,536]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="37" pageNumber="38" unit="in" value="2.8">2.80 in</quantity>
width), the spicule is curved strongly and quite sharply (
<figureCitation id="28CE2A25FFD70E79FB9AFD9DFB24FDDF" box="[1071,1189,544,567]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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). The female from the same locality has the following dimensions: body
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length and
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width, pronotum
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width.
</paragraph>
<paragraph id="B04A36A0FFD70E79FCC1FD3DFB13FCDE" blockId="37.[844,1466,194,1312]" pageId="37" pageNumber="38">
Based on the absence of a hiatus in reliable structural characters and on the partial overlapping of the dimensional characters, but with regard to the rare occurrence of
<taxonomicName id="77F54D23FFD70E79FC4CFD62FBE9FD1E" authorityName="Asanova" authorityYear="1964" box="[1017,1128,735,758]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FC4CFD62FBE9FD1E" box="[1017,1128,735,758]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
specimens corresponding to the diagnosis of
<taxonomicName id="77F54D23FFD70E79FBBFFD42FB1AFCFE" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Asanova" baseAuthorityYear="1964" box="[1034,1179,767,790]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hissaricus">
<emphasis id="8281EAB2FFD70E79FBBFFD42FB1AFCFE" box="[1034,1179,767,790]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
, I consider
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<emphasis id="8281EAB2FFD70E79FA9DFD42FA38FCFE" box="[1320,1465,767,790]" italics="true" pageId="37" pageNumber="38">C. hissaricus</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="77F54D23FFD70E79FB94FCA2FB0AFCDE" authorityName="Asanova" authorityYear="1964" box="[1057,1163,799,822]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD70E79FB94FCA2FB0AFCDE" box="[1057,1163,799,822]" italics="true" pageId="37" pageNumber="38">C. mixtus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD70E79FCF9FC31FA30FBA2" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="B04A36A0FFD70E79FCF9FC31FC38FC4B" blockId="37.[844,1466,194,1312]" box="[844,953,908,931]" pageId="37" pageNumber="38">
<heading id="EB0281CCFFD70E79FCF9FC31FC38FC4B" bold="true" box="[844,953,908,931]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="6">
<emphasis id="8281EAB2FFD70E79FCF9FC31FC38FC4B" bold="true" box="[844,953,908,931]" pageId="37" pageNumber="38">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD70E79FCF9FC09FA30FBA2" blockId="37.[844,1466,194,1312]" pageId="37" pageNumber="38">
Males smaller: body not more than
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length and not more than
<quantity id="770D9B45FFD70E79FC75FC69FB8AFC03" box="[960,1035,980,1003]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="37" pageNumber="38" unit="in" value="3.2">3.20 in</quantity>
width, pronotum not more than
<quantity id="770D9B45FFD70E79FADBFC69FA38FC03" box="[1390,1465,980,1003]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" pageId="37" pageNumber="38" unit="in" value="2.75">2.75 in</quantity>
width. Spicule at dorsolateral lobes of conjunctiva always sharply curved beyond middle (
<figureCitation id="28CE2A25FFD70E79FB1EFBA9FAC7FBC3" box="[1195,1350,1044,1067]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="37" pageNumber="38">
Figure 7
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, 7
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</figureCitation>
). Anterior part of gynatrial sac relatively rather small (
<figureCitation id="28CE2A25FFD70E79FA9DFB8EFA20FBA2" box="[1320,1441,1075,1098]" captionStart="Figure 11" captionStartId="17.[158,225,1769,1790]" captionTargetBox="[198,1415,169,1738]" captionTargetId="figure-81@17.[196,1427,168,1751]" captionTargetPageId="17" captionText="Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="37" pageNumber="38">
Figure 11
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).
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFD70E79FCF9FB1DFBF7FAF7" pageId="37" pageNumber="38" type="description">
<paragraph id="B04A36A0FFD70E79FCF9FB1DFBEFFB5F" blockId="37.[844,1466,194,1312]" box="[844,1134,1184,1207]" pageId="37" pageNumber="38">
<heading id="EB0281CCFFD70E79FCF9FB1DFBEFFB5F" bold="true" box="[844,1134,1184,1207]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="6">
<emphasis id="8281EAB2FFD70E79FCF9FB1DFBEFFB5F" bold="true" box="[844,1134,1184,1207]" pageId="37" pageNumber="38">Additional measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD70E79FCF9FB75FBF7FAF7" blockId="37.[844,1466,194,1312]" pageId="37" pageNumber="38">
Head
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width; ocular index 3.47–4.38; length of antennal segments I–V: 0.28–0.33: 0.42–0.55: 0.46– 0.53: 0.69–0.71: 0.79–0.87.
</paragraph>
</subSubSection>
<caption id="E48A6628FFD70E79FF2BF89BFF5AF898" ID-DOI="http://doi.org/10.5281/zenodo.14927051" ID-Zenodo-Dep="14927051" httpUri="https://zenodo.org/record/14927051/files/figure.png" pageId="37" pageNumber="38" startId="37.[158,225,1830,1851]" targetBox="[211,1424,1405,1787]" targetPageId="37" targetType="figure">
<paragraph id="B04A36A0FFD70E79FF2BF89BFF5AF898" blockId="37.[158,1466,1830,1904]" pageId="37" pageNumber="38">
<emphasis id="8281EAB2FFD70E79FF2BF89BFE84F8D3" bold="true" box="[158,261,1830,1851]" pageId="37" pageNumber="38">Figure 17.</emphasis>
Comparison of body measurements in
<taxonomicName id="77F54D23FFD70E79FD38F89BFBB3F8D3" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[653,1074,1830,1851]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="mixtus" subSpecies="mixtus">
<emphasis id="8281EAB2FFD70E79FD38F89BFC1CF8D3" box="[653,925,1830,1851]" italics="true" pageId="37" pageNumber="38">Canthophorus mixtus mixtus</emphasis>
<bibRefCitation id="D4644B51FFD70E79FC16F89BFBB3F8D3" author="Asanova RB" box="[931,1074,1830,1851]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="37" pageNumber="38" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
(1, 2) and
<taxonomicName id="77F54D23FFD70E79FB2FF89BFF4FF8BD" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Kiritshenko" baseAuthorityYear="1964" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="mixtus" status="n. stat." subSpecies="hissaricus">
<emphasis id="8281EAB2FFD70E79FB2FF89BFADBF8D3" box="[1178,1370,1830,1851]" italics="true" pageId="37" pageNumber="38">C. mixtus hissaricus</emphasis>
<bibRefCitation id="D4644B51FFD70E79FAD5F89BFF4FF8BD" author="Asanova RB" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="37" pageNumber="38" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="99B257C9FFD70E79FF62F8FDFE9CF8BD" box="[215,285,1856,1877]" pageId="37" pageNumber="38">
<emphasis id="8281EAB2FFD70E79FF62F8FDFE99F8BD" bold="true" box="[215,280,1856,1877]" pageId="37" pageNumber="38">n. stat</emphasis>
.
</taxonomicNameLabel>
(3). 2, specimens from Trans-Ili and Alatau Khozretishi Range. BL – body length; BW – body width; PW – pronotum width.
</paragraph>
</caption>
<subSubSection id="F8EF652BFFD40E7AFF2BFF7CFD6AFEE8" box="[158,747,193,256]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="B04A36A0FFD40E7AFF2BFF7CFEA6FF30" blockId="38.[158,781,193,1938]" box="[158,295,193,216]" pageId="38" pageNumber="39">
<heading id="EB0281CCFFD40E7AFF2BFF7CFEA6FF30" bold="true" box="[158,295,193,216]" fontSize="10" level="2" pageId="38" pageNumber="39" reason="6">
<emphasis id="8281EAB2FFD40E7AFF2BFF7CFEA6FF30" bold="true" box="[158,295,193,216]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFD40E7AFF2BFF54FD6AFEE8" blockId="38.[158,781,193,1938]" box="[158,747,233,256]" pageId="38" pageNumber="39">Figure 15. Tajikistan (Gissar and Khozretishi Ranges).</paragraph>
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<emphasis id="8281EAB2FFE90E47FCF8FDC4FA80FD78" bold="true" box="[845,1281,633,656]" italics="true" pageId="27" pageNumber="28">Canthophorus melanopterus contrarius</emphasis>
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Figures 1
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<bibRefCitation id="D4644B51FFE90E47FB23FD7FFACFFD3F" author="Wagner E." box="[1174,1358,706,727]" journalOrPublisher="Bulletin de la Societe entomologique d' Egypte." pageId="27" pageNumber="28" pagination="187 - 197" part="40" refId="ref40976" refString="Wagner E. 1956. Zur Systematik von Sehirus dubius Scop. (Hemiptera - Heteroptera: Cydnidae). Bulletin de la Societe entomologique d' Egypte. 40: 187 - 197." title="Zur Systematik von Sehirus dubius Scop. (Hemiptera - Heteroptera: Cydnidae)" type="journal article" year="1956">Wagner 1956: 195</bibRefCitation>
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<emphasis id="8281EAB2FFE90E47FCF9FD45FB49FCE5" box="[844,1224,760,781]" italics="true" pageId="27" pageNumber="28">Canthophorus melanopterus contrarius</emphasis>
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;
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.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFE90E47FCF9FBE9FBB3F9F9" pageId="27" pageNumber="28" type="diagnosis">
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<heading id="EB0281CCFFE90E47FCF9FBE9FC38FB83" bold="true" box="[844,953,1108,1131]" fontSize="10" level="2" pageId="27" pageNumber="28" reason="6">
<emphasis id="8281EAB2FFE90E47FCF9FBE9FC38FB83" bold="true" box="[844,953,1108,1131]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</heading>
</paragraph>
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Membrane whitish, subtransparent or slightly smokybrownish (
<figureCitation id="28CE2A25FFE90E47FC71FB21FBB7FB5B" box="[964,1078,1180,1203]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="27" pageNumber="28">
Figure 1
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). Yellowish rib on lateral margins of pronotum and corium broader than base of antennal segment II. Lateral branch of vesica short or very short (
<figureCitation id="28CE2A25FFE90E47FCE0FB46FC47FAFA" box="[853,966,1275,1298]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="27" pageNumber="28">
Figure 4
<collectionCode id="D6E4AE65FFE90E47FC07FB46FC47FAFA" box="[946,966,1275,1298]" pageId="27" pageNumber="28">H</collectionCode>
</figureCitation>
). Spicule basal plate at dorsolateral lobes of conjunctiva with narrow distal denticle (
<figureCitation id="28CE2A25FFE90E47FA81FAA6FA28FADA" box="[1332,1449,1307,1330]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="27" pageNumber="28">
Figure 7
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). Posterior pouches of gynatrial sac approximately as long as wide and rather long posteriorly (beyond dorsal inner margin); sclerites inside pouches large, symmetrical, hemispherical, and strongly sclerotised (
<figureCitation id="28CE2A25FFE90E47FCE0FA06FC58FA3A" box="[853,985,1467,1490]" captionStart="Figure 14" captionStartId="20.[158,225,1738,1759]" captionTargetBox="[300,1351,177,1709]" captionTargetId="figure-80@20.[283,1370,164,1722]" captionTargetPageId="20" captionText="Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927045" httpUri="https://zenodo.org/record/14927045/files/figure.png" pageId="27" pageNumber="28">
Figure 14
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). Tubular proximal part of spermathecal capsule 2–3 times as long as transverse diameter of spherical distal part.
</paragraph>
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<heading id="EB0281CCFFE90E47FCF9F9EEFC70F982" bold="true" box="[844,1009,1619,1642]" fontSize="10" level="2" pageId="27" pageNumber="28" reason="6">
<emphasis id="8281EAB2FFE90E47FCF9F9EEFC70F982" bold="true" box="[844,1009,1619,1642]" pageId="27" pageNumber="28">Measurements</emphasis>
</heading>
</paragraph>
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Body 5.35–8.00 in length,
<quantity id="770D9B45FFE90E47FBC6F9C6FB7AF97A" box="[1139,1275,1658,1682]" metricMagnitude="-2" metricUnit="m" metricValue="9.779" metricValueMax="11.684" metricValueMin="7.874" pageId="27" pageNumber="28" unit="in" value="3.8499999999999996" valueMax="4.6" valueMin="3.1">3.10–4.60 in</quantity>
width; pronotum
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width; head
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width; ocular index 3.53–4.00; length of antennal segments I–V: 0.28–0.38: 0.52–0.64: 0.52–0.67: 0.78–0.91: 0.81–0.95.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFE90E40FCF9F88EFE9FFCFE" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
<paragraph id="B04A36A0FFE90E47FCF9F88EFC32F8A2" blockId="27.[844,1466,194,1938]" box="[844,947,1843,1866]" pageId="27" pageNumber="28">
<heading id="EB0281CCFFE90E47FCF9F88EFC32F8A2" bold="true" box="[844,947,1843,1866]" fontSize="10" level="2" pageId="27" pageNumber="28" reason="6">
<emphasis id="8281EAB2FFE90E47FCF9F88EFC32F8A2" bold="true" box="[844,947,1843,1866]" pageId="27" pageNumber="28">Remarks</emphasis>
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</paragraph>
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<bibRefCitation id="D4644B51FFE90E47FCF9F8E6FC6BF89A" author="Wagner E." box="[844,1002,1883,1906]" journalOrPublisher="Bulletin de la Societe entomologique d' Egypte." pageId="27" pageNumber="28" pagination="187 - 197" part="40" refId="ref40976" refString="Wagner E. 1956. Zur Systematik von Sehirus dubius Scop. (Hemiptera - Heteroptera: Cydnidae). Bulletin de la Societe entomologique d' Egypte. 40: 187 - 197." title="Zur Systematik von Sehirus dubius Scop. (Hemiptera - Heteroptera: Cydnidae)" type="journal article" year="1956">Wagner (1956)</bibRefCitation>
distinguished his new subspecies from the nominotypical one by the smaller body size (body length 5.00–
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males and
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females), slenderer proportions (body length twice greater than body width), whitish membrane, smaller parameres with smaller, narrower hyphophysis and wider sensory process, and thinner, more acute spicule (“grosse Haken”) at dorsolateral lobes of conjunctiva. I could not find any significant differences of
<emphasis id="8281EAB2FFEE0E40FD61FEDCFE86FE70" italics="true" pageId="28" pageNumber="29">C. m. contrarius</emphasis>
in the structure of parameres, and the spicules in the specimens examined by me are slightly thicker than in most of the examined specimens belonging to the nominotypical subspecies. The thicker spicule of
<emphasis id="8281EAB2FFEE0E40FDE8FE5DFC8AFE1F" box="[605,779,480,503]" italics="true" pageId="28" pageNumber="29">C. m. contrarius</emphasis>
can be seen also in the drawing of Wagner. Perhaps he meant the denticles (“kleine Zaehne”) on the spicule basal plate; in his figure 23, they are shown to be shorter and thinner, but the length and shape of these denticles actually vary somewhat, and the specimens I have studied from Cyprus have virtually the same denticles as in specimens of
<emphasis id="8281EAB2FFEE0E40FD3AFD1DFF77FD3F" italics="true" pageId="28" pageNumber="29">
C. m.
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</emphasis>
. The dimensions and proportions of the body, as seen on the material examined, also do not have diagnostic significance.
</paragraph>
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<emphasis id="8281EAB2FFEE0E40FF2BFCE5FEA6FC87" bold="true" box="[158,295,856,879]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FF2BFC3DFF73FC7F" blockId="28.[158,780,193,1938]" box="[158,242,896,919]" pageId="28" pageNumber="29">Cyprus.</paragraph>
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(
<figureCitation id="28CE2A25FFEC0E42FC1CFACAFBA0FA66" box="[937,1057,1399,1422]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="30" pageNumber="31">
Figures 1
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<figureCitation id="28CE2A25FFEC0E42FB84FACAFBD2FA66" box="[1073,1107,1399,1422]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="30" pageNumber="31">
3
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,
<figureCitation id="28CE2A25FFEC0E42FBD7FACAFB03FA66" box="[1122,1154,1399,1422]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="30" pageNumber="31">
5
<collectionCode id="D6E4AE65FFEC0E42FBDAFACAFB03FA66" box="[1135,1154,1399,1422]" pageId="30" pageNumber="31">A</collectionCode>
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,
<figureCitation id="28CE2A25FFEC0E42FB27FACAFB35FA66" box="[1170,1204,1399,1422]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="30" pageNumber="31">
7
<collectionCode id="D6E4AE65FFEC0E42FB2AFACAFB35FA66" box="[1183,1204,1399,1422]" pageId="30" pageNumber="31">H</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFEC0E42FB76FACAFB63FA66" box="[1219,1250,1399,1422]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="30" pageNumber="31">
8
<collectionCode id="D6E4AE65FFEC0E42FB65FACAFB63FA66" box="[1232,1250,1399,1422]" pageId="30" pageNumber="31">L</collectionCode>
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,
<figureCitation id="28CE2A25FFEC0E42FB47FACAFA9EFA66" box="[1266,1311,1399,1422]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="30" pageNumber="31">
10
<collectionCode id="D6E4AE65FFEC0E42FAB9FACAFA9EFA66" box="[1292,1311,1399,1422]" pageId="30" pageNumber="31">C</collectionCode>
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,
<figureCitation id="28CE2A25FFEC0E42FA9AFACAFADDFA66" box="[1327,1372,1399,1422]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="30" pageNumber="31">
12
<collectionCode id="D6E4AE65FFEC0E42FAFCFACAFADDFA66" box="[1353,1372,1399,1422]" pageId="30" pageNumber="31">D</collectionCode>
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)
</emphasis>
</paragraph>
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<paragraph id="B04A36A0FFEC0E42FCF9FA68FC6AFA04" blockId="30.[844,1466,194,1938]" box="[844,1003,1493,1516]" pageId="30" pageNumber="31">
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<typeStatus id="6F4E8802FFEC0E42FCF9FA68FC02FA04" box="[844,899,1493,1516]" pageId="30" pageNumber="31">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
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<paragraph id="B04A36A0FFEC0E43FCF9FA40FDEAFCDE" blockId="30.[844,1466,194,1938]" lastBlockId="31.[158,781,193,1938]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<materialsCitation id="009D3CFDFFEC0E42FCF9FA40FAFFF99B" collectionCode="ZISP" country="Armenia" latitude="40.270084" location="Gekhmakhan or Ziarat-Kizyl Mt." longLatPrecision="1" longitude="44.894306" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Aragatsotn Prov" typeStatus="holotype">
<typeStatus id="6F4E8802FFEC0E42FCF9FA40FC33F9FC" box="[844,946,1533,1556]" pageId="30" pageNumber="31">Holotype</typeStatus>
:
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<collectingCountry id="C8E27630FFEC0E42FC74FA40FBA5F9FC" box="[961,1060,1533,1556]" name="Armenia" pageId="30" pageNumber="31" value="Armenia">Armenia</collectingCountry>
</emphasis>
:
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<collectingRegion id="7231F842FFEC0E42FB8DFA40FB6AF9FC" box="[1080,1259,1533,1556]" pageId="30" pageNumber="31">Aragatsotn Prov</collectingRegion>
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., “gora Kizilziarat [
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,
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40°16
<emphasis id="8281EAB2FFEC0E42FA85F9A1FAB7F9DC" bold="true" box="[1328,1334,1564,1588]" italics="true" pageId="30" pageNumber="31">ʹ</emphasis>
12.3
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N
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<geoCoordinate id="D5C15067FFEC0E42FA21F9A0FC37F9BB" degrees="44" direction="east" minutes="53" orientation="longitude" pageId="30" pageNumber="31" precision="1" seconds="39.5" value="44.894306">
44° 53
<emphasis id="8281EAB2FFEC0E42FCD3F981FCEDF9BC" bold="true" box="[870,876,1596,1620]" italics="true" pageId="30" pageNumber="31">ʹ</emphasis>
39.5
<emphasis id="8281EAB2FFEC0E42FC2EF981FC24F9BC" bold="true" box="[923,933,1596,1620]" italics="true" pageId="30" pageNumber="31">ʺ</emphasis>
E
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]/Novobayaz.[etskiy] u.[ezd] Eriv.[anskoy] g. [ubernii]/[Ya.G.] Efremov. [
<date id="C44B1060FFEC0E42FBC3F9E1FB37F99B" box="[1142,1206,1628,1651]" pageId="30" pageNumber="31">1.VII.</date>
1]909.”,
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(
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)
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.
<materialsCitation id="009D3CFDFFEC0E42FCC1F9C1FC12F91B" collectingDate="1922-07-22" collectionCode="ZISP" country="Georgia" location="North-Caucasian Regional Plant Protection Station" municipality="Lebarde" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="paratype">
<typeStatus id="6F4E8802FFEC0E42FCC1F9C1FC5BF97B" box="[884,986,1660,1683]" pageId="30" pageNumber="31">Paratypes</typeStatus>
:
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<collectingCountry id="C8E27630FFEC0E42FC53F9C1FBBCF97C" box="[998,1085,1660,1684]" name="Georgia" pageId="30" pageNumber="31" value="Georgia">Georgia</collectingCountry>
</emphasis>
:
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<collectingRegion id="7231F842FFEC0E42FBFAF9C1FACEF97B" box="[1103,1359,1660,1683]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="30" pageNumber="31">Samegrelo-Zemo Svaneti</collectingRegion>
</emphasis>
,
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, “Sev.-Kav. Kraystazra” [
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],
<date id="C44B1060FFEC0E42FB93F901FB23F93B" box="[1062,1186,1724,1747]" pageId="30" pageNumber="31" value="1922-07-22">
<collectingDate id="D40FE988FFEC0E42FB93F901FB23F93B" box="[1062,1186,1724,1747]" pageId="30" pageNumber="31" value="1922-07-22">22.VII.1922</collectingDate>
</date>
(N. Arkhangel’ skiy),
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(
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)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFEC0E42FC1FF961FAC0F8FB" collectingDate="1909-06-26" collectionCode="ZISP" country="Georgia" location="Tambovka" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Samtskhe-Javakheti">
<emphasis id="8281EAB2FFEC0E42FC1FF961FBF3F91B" box="[938,1138,1756,1779]" italics="true" pageId="30" pageNumber="31">
<collectingRegion id="7231F842FFEC0E42FC1FF961FBF3F91B" box="[938,1138,1756,1779]" pageId="30" pageNumber="31">Samtskhe-Javakheti</collectingRegion>
</emphasis>
:
<location id="B52A607BFFEC0E42FB38F961FB78F91B" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFEC0E42FB38F961FB78F91B" box="[1165,1273,1756,1779]" country="Georgia" name="Tambovka" pageId="30" pageNumber="31" stateProvince="Samtskhe-Javakheti">Tambovka</location>
<locationDeviation id="A1F82F82FFEC0E42FAB9F961FC72F8FB" pageId="30" pageNumber="31">nr “Toporovan” [Paravani] Lake</locationDeviation>
,
<date id="C44B1060FFEC0E42FBB4F941FBF4F8FB" box="[1025,1141,1788,1811]" pageId="30" pageNumber="31" value="1909-06-26">
<collectingDate id="D40FE988FFEC0E42FBB4F941FBF4F8FB" box="[1025,1141,1788,1811]" pageId="30" pageNumber="31" value="1909-06-26">26.VI.1909</collectingDate>
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(Berg),
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(
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)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFEC0E42FAFAF941FB66F8DA" collectingDate="1910-06-11" collectionCode="ZISP" country="Georgia" location="Bakuriani" municipality="Bakuriani" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Samtskhe-javakheti">
<collectingMunicipality id="502EACDAFFEC0E42FAFAF941FA34F8FB" box="[1359,1461,1788,1811]" pageId="30" pageNumber="31">Bakuriani</collectingMunicipality>
,
<date id="C44B1060FFEC0E42FCF9F8A6FC3EF8DA" box="[844,959,1819,1842]" pageId="30" pageNumber="31" value="1910-06-11">
<collectingDate id="D40FE988FFEC0E42FCF9F8A6FC3EF8DA" box="[844,959,1819,1842]" pageId="30" pageNumber="31" value="1910-06-11">11.VI.1910</collectingDate>
</date>
(Mlakosevich),
<specimenCount id="A6F3FD29FFEC0E42FBC7F8A6FB12F8DB" box="[1138,1171,1819,1843]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFEC0E42FB12F8A6FB61F8DA" box="[1191,1248,1819,1842]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="30" pageNumber="31">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFEC0E42FB4DF8A6FABFF89A" collectingDate="1980-07-24" collectionCode="ZISP" collectorName="M. G. Volkovitsh" country="Georgia" elevation="1950" location="Akhalkalaki" municipality="Paravani Riv." pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Samtskhe-javakheti">
gorge of
<collectingMunicipality id="502EACDAFFEC0E42FAD4F8A6FCF9F8BA" pageId="30" pageNumber="31">Paravani Riv.</collectingMunicipality>
,
<locationDeviation id="A1F82F82FFEC0E42FC3CF886FBD2F8BA" box="[905,1107,1851,1874]" pageId="30" pageNumber="31">
<quantity id="770D9B45FFEC0E42FC3CF886FC47F8BA" box="[905,966,1851,1874]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="5.0">5 km</quantity>
upstream of
</locationDeviation>
<location id="B52A607BFFEC0E42FBD7F886FB61F8BA" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFEC0E42FBD7F886FB61F8BA" box="[1122,1248,1851,1874]" country="Georgia" municipality="Paravani Riv." name="Akhalkalaki" pageId="30" pageNumber="31" stateProvince="Samtskhe-javakheti">Akhalkalaki</location>
,
<quantity id="770D9B45FFEC0E42FB44F886FAC6F8BA" box="[1265,1351,1851,1874]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="30" pageNumber="31" unit="m" value="1950.0">
<elevation id="3BD8D193FFEC0E42FB44F886FAC6F8BA" box="[1265,1351,1851,1874]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="30" pageNumber="31" unit="m" value="1950.0">1950 m</elevation>
</quantity>
amsl,
<date id="C44B1060FFEC0E42FA2CF886FC29F89A" pageId="30" pageNumber="31" value="1980-07-24">
<collectingDate id="D40FE988FFEC0E42FA2CF886FC29F89A" pageId="30" pageNumber="31" value="1980-07-24">24. VII.1980</collectingDate>
</date>
(
<collectorName id="1D005376FFEC0E42FC75F8E6FBF6F89A" box="[960,1143,1883,1906]" pageId="30" pageNumber="31">M.G. Volkovitsh</collectorName>
), 2J,
<specimenCount id="A6F3FD29FFEC0E42FB72F8E6FB69F89B" box="[1223,1256,1883,1907]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFEC0E42FB4AF8E6FAB9F89A" box="[1279,1336,1883,1906]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="30" pageNumber="31">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFEC0E43FAE7F8E6FDBEFEF0" collectionCode="ZISP, MNHN" country="Armenia" lastPageId="31" lastPageNumber="32" latitude="40.270084" location="Gekhmakhan or Ziarat-Kizyl Mt." longLatPrecision="1" longitude="44.894306" pageId="30" pageNumber="31" specimenCount="8" specimenCount-female="8" stateProvince="Aragatsotn Prov">
<emphasis id="8281EAB2FFEC0E42FAE7F8E6FA33F89A" bold="true" box="[1362,1458,1883,1906]" pageId="30" pageNumber="31">
<collectingCountry id="C8E27630FFEC0E42FAE7F8E6FA33F89A" box="[1362,1458,1883,1906]" name="Armenia" pageId="30" pageNumber="31" value="Armenia">Armenia</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFEC0E42FCF9F8C6FC72F87A" box="[844,1011,1915,1938]" italics="true" pageId="30" pageNumber="31">
<collectingRegion id="7231F842FFEC0E42FCF9F8C6FC72F87A" box="[844,1011,1915,1938]" pageId="30" pageNumber="31">Aragatsotn Prov</collectingRegion>
</emphasis>
.: “gora Kizilziarat [
<location id="B52A607BFFEC0E43FB76F8C6FE88FF30" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFEC0E43FB76F8C6FE88FF30" country="Armenia" lastPageId="31" lastPageNumber="32" latitude="40.270084" longLatPrecision="1" longitude="44.894306" name="Gekhmakhan or Ziarat-Kizyl Mt." pageId="30" pageNumber="31" stateProvince="Aragatsotn Prov">Gekhmakhan or Ziarat-Kizyl Mt.</location>
,
<geoCoordinate id="D5C15067FFED0E43FEADFF7CFE22FF30" box="[280,419,193,217]" degrees="40" direction="north" minutes="16" orientation="latitude" pageId="31" pageNumber="32" precision="1" seconds="12.3" value="40.270084">
40°16
<emphasis id="8281EAB2FFED0E43FEE3FF7CFEDAFF31" bold="true" box="[342,347,193,217]" italics="true" pageId="31" pageNumber="32">ʹ</emphasis>
12.3
<emphasis id="8281EAB2FFED0E43FE32FF7CFE11FF31" bold="true" box="[391,400,193,217]" italics="true" pageId="31" pageNumber="32">ʺ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="D5C15067FFED0E43FE1AFF7CFDB6FF30" box="[431,567,193,217]" degrees="44" direction="east" minutes="53" orientation="longitude" pageId="31" pageNumber="32" precision="1" seconds="39.5" value="44.894306">
44°53
<emphasis id="8281EAB2FFED0E43FE59FF7CFE70FF31" bold="true" box="[492,497,193,217]" italics="true" pageId="31" pageNumber="32">ʹ</emphasis>
39.5
<emphasis id="8281EAB2FFED0E43FDABFF7CFDA6FF31" bold="true" box="[542,551,193,217]" italics="true" pageId="31" pageNumber="32">ʺ</emphasis>
E
</geoCoordinate>
]/Novobayaz.[etskiy] u.[ezd] Eriv.[anskoy] g.[ubernii]/[Ya.G.] Efremov. [
<date id="C44B1060FFED0E43FD0FFF5CFD7AFF10" box="[698,763,225,248]" pageId="31" pageNumber="32">1.VII.</date>
1] 909.”, 4J,
<specimenCount id="A6F3FD29FFED0E43FE97FEBCFEC2FEF1" box="[290,323,257,281]" count="7" pageId="31" pageNumber="32" type="female">7♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FEECFEBCFE10FEF0" box="[345,401,257,280]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
),
<specimenCount id="A6F3FD29FFED0E43FE19FEBCFE4CFEF1" box="[428,461,257,281]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FE51FEBCFDB9FEF0" box="[484,568,257,280]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="31" pageNumber="32">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFED0E43FDE7FEBCFEEDFE90" collectingDate="1934-06-16" collectionCode="ZISP, MNHN" country="Armenia" elevation="2000" location="Aragats Mt." municipality="Mantash Riv." pageId="31" pageNumber="32" specimenCount="3" specimenCount-female="3" stateProvince="Aragatsotn Prov" typeStatus="paratype">
upper reaches of
<collectingMunicipality id="502EACDAFFED0E43FF2BFE9CFEADFED0" box="[158,300,289,312]" pageId="31" pageNumber="32">Mantash Riv.</collectingMunicipality>
, “Alagez” [
<location id="B52A607BFFED0E43FE12FE9CFDABFED0" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FE12FE9CFDABFED0" box="[423,554,289,312]" country="Armenia" municipality="Mantash Riv." name="Aragats Mt." pageId="31" pageNumber="32" stateProvince="Aragatsotn Prov">Aragats Mt.</location>
],
<emphasis id="8281EAB2FFED0E43FD8AFE9CFDCBFED0" box="[575,586,289,312]" italics="true" pageId="31" pageNumber="32">c</emphasis>
.
<quantity id="770D9B45FFED0E43FDF8FE9CFD20FED0" box="[589,673,289,312]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="2000.0">
<elevation id="3BD8D193FFED0E43FDF8FE9CFD20FED0" box="[589,673,289,312]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="2000.0">2000 m</elevation>
</quantity>
amsl,
<date id="C44B1060FFED0E43FD59FE9CFF72FEB0" pageId="31" pageNumber="32" value="1934-06-16">
<collectingDate id="D40FE988FFED0E43FD59FE9CFF72FEB0" pageId="31" pageNumber="32" value="1934-06-16">16. VI.1934</collectingDate>
</date>
(Ter-Minassjan), 1J,
<specimenCount id="A6F3FD29FFED0E43FE65FEFCFE70FEB1" box="[464,497,321,345]" count="2" pageId="31" pageNumber="32" type="female">2♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FE48FEFCFDB4FEB0" box="[509,565,321,344]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
),
<specimenCount id="A6F3FD29FFED0E43FDF3FEFCFDE7FEB1" box="[582,614,321,345]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FDC6FEFCFD44FEB0" box="[627,709,321,344]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="31" pageNumber="32">MNHN</collectionCode>
) [
<typeStatus id="6F4E8802FFED0E43FD63FEFCFF55FE90" pageId="31" pageNumber="32">paratypes</typeStatus>
of
<taxonomicName id="77F54D23FFED0E43FF4AFEDCFEE5FE90" authorityName="Asanova" authorityYear="1964" box="[255,356,353,376]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFED0E43FF4AFEDCFEE5FE90" box="[255,356,353,376]" italics="true" pageId="31" pageNumber="32">C. mixtus</emphasis>
</taxonomicName>
]
</materialsCitation>
;
<materialsCitation id="009D3CFDFFED0E43FECBFEDCFF68FE50" collectingDate="1935-07-29" collectionCode="ZISP" country="Armenia" elevation="3000" location="Aragats Mt." municipality="Alagez" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Aragatsotn Prov" typeStatus="paratype">
NW slope of “
<collectingMunicipality id="502EACDAFFED0E43FD95FEDCFDE6FE90" box="[544,615,353,376]" pageId="31" pageNumber="32">Alagez</collectingMunicipality>
” [
<location id="B52A607BFFED0E43FD31FEDCFC87FE90" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FD31FEDCFC87FE90" box="[644,774,353,376]" country="Armenia" municipality="Alagez" name="Aragats Mt." pageId="31" pageNumber="32" stateProvince="Aragatsotn Prov">Aragats Mt.</location>
]
<quantity id="770D9B45FFED0E43FF2BFE3CFF73FE70" box="[158,242,385,408]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="m" value="3000.0">
<elevation id="3BD8D193FFED0E43FF2BFE3CFF73FE70" box="[158,242,385,408]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="m" value="3000.0">3000 m</elevation>
</quantity>
,
<date id="C44B1060FFED0E43FEB5FE3CFEFCFE70" box="[256,381,385,408]" pageId="31" pageNumber="32" value="1935-07-29">
<collectingDate id="D40FE988FFED0E43FEB5FE3CFEFCFE70" box="[256,381,385,408]" pageId="31" pageNumber="32" value="1935-07-29">29.VII.1935</collectingDate>
</date>
(Rikhter),
<specimenCount id="A6F3FD29FFED0E43FE47FE3CFD92FE70" box="[498,531,384,408]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FD91FE3CFDDAFE70" box="[548,603,385,408]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
) [
<typeStatus id="6F4E8802FFED0E43FDC4FE3CFD4BFE70" box="[625,714,385,408]" pageId="31" pageNumber="32">paratype</typeStatus>
of
<taxonomicName id="77F54D23FFED0E43FD41FE3CFF61FE50" authorityName="Asanova" authorityYear="1964" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFED0E43FD41FE3CFF61FE50" italics="true" pageId="31" pageNumber="32">C. mixtus</emphasis>
</taxonomicName>
]
</materialsCitation>
;
<materialsCitation id="009D3CFDFFED0E43FF40FE1CFE54FE1F" collectionCode="ZISP" collectorName="A. Schelk" country="Armenia" location="Agmagan [- Karadag" municipality="Agmagan [- Karadag" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Kotayk Prov">
<emphasis id="8281EAB2FFED0E43FF40FE1CFEF2FE50" box="[245,371,417,440]" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="7231F842FFED0E43FF40FE1CFEF2FE50" box="[245,371,417,440]" pageId="31" pageNumber="32">Kotayk Prov</collectingRegion>
</emphasis>
.,
<collectingMunicipality id="502EACDAFFED0E43FE30FE1CFDD2FE50" box="[389,595,417,440]" pageId="31" pageNumber="32">Agmagan[-Karadag</collectingMunicipality>
], “pr. alp. Šiš- kara [?] 10,000”,
<date id="C44B1060FFED0E43FEA9FE7CFEE6FE30" box="[284,359,449,472]" pageId="31" pageNumber="32">16.VII.</date>
[19]26, “pr. alp. Ciċagio [?] 9000”,
<date id="C44B1060FFED0E43FD72FE7CFC8DFE30" box="[711,780,449,472]" pageId="31" pageNumber="32">1.VIII.</date>
[19]26 (
<collectorName id="1D005376FFED0E43FF45FE5DFED6FE1F" box="[240,343,480,503]" pageId="31" pageNumber="32">A. Schelk</collectorName>
), 2J (
<collectionCode id="D6E4AE65FFED0E43FE20FE5DFE4FFE1F" box="[405,462,480,503]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFED0E43FE6AFE5DFE9EFDDF" collectingDate="1980-07-30" collectionCode="ZISP" collectorName="M. G. Volkovitsh" country="Armenia" elevation="2100" location="Lchashen Settlm. env." pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Gegharkunik Prov">
<emphasis id="8281EAB2FFED0E43FE6AFE5DFD19FE1F" box="[479,664,480,503]" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="7231F842FFED0E43FE6AFE5DFD19FE1F" box="[479,664,480,503]" pageId="31" pageNumber="32">Gegharkunik Prov</collectingRegion>
</emphasis>
.,
<location id="B52A607BFFED0E43FD1FFE5DFE9DFDFF" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FD1FFE5DFE9DFDFF" country="Armenia" name="Lchashen Settlm. env." pageId="31" pageNumber="32" stateProvince="Gegharkunik Prov">Lchashen Settlm. env.</location>
,
<quantity id="770D9B45FFED0E43FE9CFDBDFEF8FDFF" box="[297,377,512,535]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="32" unit="m" value="2100.0">
<elevation id="3BD8D193FFED0E43FE9CFDBDFEF8FDFF" box="[297,377,512,535]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="32" unit="m" value="2100.0">2100 m</elevation>
</quantity>
amsl,
<date id="C44B1060FFED0E43FE76FDBDFDBEFDFF" box="[451,575,512,535]" pageId="31" pageNumber="32" value="1980-07-30">
<collectingDate id="D40FE988FFED0E43FE76FDBDFDBEFDFF" box="[451,575,512,535]" pageId="31" pageNumber="32" value="1980-07-30">30.VII.1980</collectingDate>
</date>
(
<collectorName id="1D005376FFED0E43FDE5FDBDFC80FDFF" box="[592,769,512,535]" pageId="31" pageNumber="32">M.G. Volkovitsh</collectorName>
),
<specimenCount id="A6F3FD29FFED0E43FF2BFD9DFF3EFDD0" box="[158,191,544,568]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FF55FD9DFE98FDDF" box="[224,281,544,567]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="009D3CFDFFED0E43FE88FD9DFD02FD9F" collectingDate="1959-08-17" collectionCode="ZISP" collectorName="V. Zaitzev" country="Armenia" elevation="2375" location="Kaputjugh] Mt." pageId="31" pageNumber="32" specimenCount="1" stateProvince="Syunik Prov.">
<collectingRegion id="7231F842FFED0E43FE88FD9DFE4EFDDF" box="[317,463,544,567]" pageId="31" pageNumber="32">
<emphasis id="8281EAB2FFED0E43FE88FD9DFE48FDDF" box="[317,457,544,567]" italics="true" pageId="31" pageNumber="32">Syunik Prov</emphasis>
.
</collectingRegion>
, E slopes “Karutdzhukh” [
<location id="B52A607BFFED0E43FF11FDFDFEC7FDBF" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FF11FDFDFEC7FDBF" box="[164,326,576,599]" country="Armenia" name="Kaputjugh] Mt." pageId="31" pageNumber="32" stateProvince="Syunik Prov.">Kaputjugh] Mt.</location>
,
<quantity id="770D9B45FFED0E43FEFAFDFDFE5DFDBF" box="[335,476,576,600]" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="2.7" metricValueMin="2.05" pageId="31" pageNumber="32" unit="m" value="2375.0" valueMax="2700.0" valueMin="2050.0">
<elevation id="3BD8D193FFED0E43FEFAFDFDFE5DFDBF" box="[335,476,576,600]" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="2.7" metricValueMin="2.05" pageId="31" pageNumber="32" unit="m" value="2375.0" valueMax="2700.0" valueMin="2050.0">2050–2700 m</elevation>
</quantity>
amsl, mountain steppe, alpine meadows,
<date id="C44B1060FFED0E43FEBFFDDDFE0EFD9F" box="[266,399,608,631]" pageId="31" pageNumber="32" value="1959-08-17">
<collectingDate id="D40FE988FFED0E43FEBFFDDDFE0EFD9F" box="[266,399,608,631]" pageId="31" pageNumber="32" value="1959-08-17">17.VIII.1959</collectingDate>
</date>
(
<collectorName id="1D005376FFED0E43FE2EFDDDFD87FD9F" box="[411,518,608,631]" pageId="31" pageNumber="32">V. Zaitzev</collectorName>
), 1J (
<collectionCode id="D6E4AE65FFED0E43FDF6FDDDFDFDFD9F" box="[579,636,608,631]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFED0E43FD39FDDDFDC4FD7F" collectionCode="ZISP" collectorName="V. Jakovlev" country="Azerbaijan" location="Ordubad" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Ordubad">
<emphasis id="8281EAB2FFED0E43FD39FDDDFC84FD9F" bold="true" box="[652,773,608,631]" pageId="31" pageNumber="32">
<collectingCountry id="C8E27630FFED0E43FD39FDDDFC84FD9F" box="[652,773,608,631]" name="Azerbaijan" pageId="31" pageNumber="32" value="Azerbaijan">Azerbaijan</collectingCountry>
</emphasis>
:
<collectingRegion id="7231F842FFED0E43FF2BFD3DFF79FD7F" box="[158,248,640,663]" country="Azerbaijan" name="Ordubad" pageId="31" pageNumber="32">Ordubad</collectingRegion>
(
<collectorName id="1D005376FFED0E43FEBCFD3DFE05FD7F" box="[265,388,640,663]" pageId="31" pageNumber="32">V. Jakovlev</collectorName>
coll.),
<specimenCount id="A6F3FD29FFED0E43FE66FD3DFE75FD7F" box="[467,500,639,663]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FDB0FD3DFDBFFD7F" box="[517,574,640,663]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFED0E43FDE6FD3DFD30FD5F" collectingDate="1910-06-30" collectionCode="ZISP" country="Turkey" location="Ardahan" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Ardahan Prov">
<emphasis id="8281EAB2FFED0E43FDE6FD3DFD20FD7F" bold="true" box="[595,673,640,663]" pageId="31" pageNumber="32">
<collectingCountry id="C8E27630FFED0E43FDE6FD3DFD20FD7F" box="[595,673,640,663]" name="Turkey" pageId="31" pageNumber="32" value="Turkey">Turkey</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFED0E43FD06FD3DFF4CFD5F" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="7231F842FFED0E43FD06FD3DFF4CFD5F" pageId="31" pageNumber="32">Ardahan Prov</collectingRegion>
</emphasis>
.,
<location id="B52A607BFFED0E43FF57FD1DFEBDFD5F" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FF57FD1DFEBDFD5F" box="[226,316,672,695]" country="Turkey" name="Ardahan" pageId="31" pageNumber="32" stateProvince="Ardahan Prov">
<collectingRegion id="7231F842FFED0E43FF57FD1DFEBDFD5F" box="[226,316,672,695]" country="Turkey" name="Ardahan" pageId="31" pageNumber="32">Ardahan</collectingRegion>
</location>
,
<date id="C44B1060FFED0E43FEFCFD1DFE3CFD5F" box="[329,445,672,695]" pageId="31" pageNumber="32" value="1910-06-30">
<collectingDate id="D40FE988FFED0E43FEFCFD1DFE3CFD5F" box="[329,445,672,695]" pageId="31" pageNumber="32" value="1910-06-30">30.VI.1910</collectingDate>
</date>
(Nesterov),
<specimenCount id="A6F3FD29FFED0E43FDF5FD1DFDE0FD5F" box="[576,609,671,695]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FDC4FD1DFD2BFD5F" box="[625,682,672,695]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFED0E43FD08FD1DFF64FCFE" collectionCode="ZISP" country="Russia" location="Sikhote-Alin" municipality="Khor Riv." pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Khabarovsk Terr">
<emphasis id="8281EAB2FFED0E43FD08FD1DFC84FD5F" bold="true" box="[701,773,672,695]" pageId="31" pageNumber="32">
<collectingCountry id="C8E27630FFED0E43FD08FD1DFC84FD5F" box="[701,773,672,695]" name="Russia" pageId="31" pageNumber="32" value="Russia">Russia</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFED0E43FF2BFD7DFED0FD3F" box="[158,337,704,727]" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="7231F842FFED0E43FF2BFD7DFED0FD3F" box="[158,337,704,727]" pageId="31" pageNumber="32">Khabarovsk Terr</collectingRegion>
</emphasis>
.:
<collectingMunicipality id="502EACDAFFED0E43FEDBFD7DFE5FFD3F" box="[366,478,704,727]" pageId="31" pageNumber="32">Khor Riv.</collectingMunicipality>
, spurs of
<location id="B52A607BFFED0E43FDEFFD7DFD5EFD3F" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FDEFFD7DFD5EFD3F" box="[602,735,704,727]" country="Russia" municipality="Khor Riv." name="Sikhote-Alin" pageId="31" pageNumber="32" stateProvince="Khabarovsk Terr">Sikhote-Alin</location>
, 56 “verst” [
<locationDeviation id="A1F82F82FFED0E43FF4EFD62FD96FD1E" box="[251,535,735,758]" pageId="31" pageNumber="32">
<quantity id="770D9B45FFED0E43FF4EFD62FEC8FD1E" box="[251,329,735,758]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="km" value="60.0">60 km</quantity>
] from
<collectingRegion id="7231F842FFED0E43FE2AFD62FE5AFD1E" box="[415,475,735,758]" country="Russia" name="Amur" pageId="31" pageNumber="32">Amur</collectingRegion>
Riv.
</locationDeviation>
, 1909 (Efremov),
<specimenCount id="A6F3FD29FFED0E43FD5EFD62FC8DFD1F" box="[747,780,735,759]" count="1" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFED0E43FF10FD42FF5FFCFE" box="[165,222,767,790]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="009D3CFDFFED0E43FF47FD42FDE7FCDE" collectionCode="ZISP" country="China" location="Kangding" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Sichuan">
<emphasis id="8281EAB2FFED0E43FF47FD42FEB5FCFE" bold="true" box="[242,308,767,790]" pageId="31" pageNumber="32">
<collectingCountry id="C8E27630FFED0E43FF47FD42FEB5FCFE" box="[242,308,767,790]" name="China" pageId="31" pageNumber="32" value="China">China</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFED0E43FEF1FD42FE17FCFE" box="[324,406,767,790]" italics="true" pageId="31" pageNumber="32">
<collectingRegion id="7231F842FFED0E43FEF1FD42FE17FCFE" box="[324,406,767,790]" country="China" name="Sichuan" pageId="31" pageNumber="32">Sichuan</collectingRegion>
</emphasis>
, “Sztschwah, Ta-tsien-lu” [
<collectingRegion id="7231F842FFED0E43FD07FD42FC89FCFE" box="[690,776,767,790]" country="China" name="Sichuan" pageId="31" pageNumber="32">Sichuan</collectingRegion>
,
<location id="B52A607BFFED0E43FF2BFCA2FE84FCDE" LSID="urn:lsid:plazi:treatment:385C87B6FFEC0E43FC3DFAEAFB49F9DF:B52A607BFFED0E43FF2BFCA2FE84FCDE" box="[158,261,799,822]" country="China" name="Kangding" pageId="31" pageNumber="32" stateProvince="Sichuan">Kangding</location>
] (“St. Breuning ded.”), 1J (
<collectionCode id="D6E4AE65FFED0E43FD93FCA2FDDEFCDE" box="[550,607,799,822]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E43FF2BFCC3FDF9FB74" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="B04A36A0FFED0E43FF2BFCC3FE8AFC7D" blockId="31.[158,781,193,1938]" box="[158,267,894,917]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FF2BFCC3FE8AFC7D" bold="true" box="[158,267,894,917]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FF2BFCC3FE8AFC7D" bold="true" box="[158,267,894,917]" pageId="31" pageNumber="32">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FF2BFC1BFDF9FB74" blockId="31.[158,781,193,1938]" pageId="31" pageNumber="32">
Membrane brownish, opaque (
<figureCitation id="28CE2A25FFED0E43FE56FC1BFDCAFC55" box="[483,587,934,957]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="31" pageNumber="32">
Figure 1
<collectionCode id="D6E4AE65FFED0E43FD8CFC1BFDCAFC55" box="[569,587,934,957]" pageId="31" pageNumber="32">F</collectionCode>
</figureCitation>
). Sensory process of paramere rather widely rounded at apex (
<figureCitation id="28CE2A25FFED0E43FD38FC7BFD7CFC35" box="[653,765,966,989]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="31" pageNumber="32">
Figure 3
<collectionCode id="D6E4AE65FFED0E43FD5CFC7BFD7CFC35" box="[745,765,966,989]" pageId="31" pageNumber="32">G</collectionCode>
</figureCitation>
). Spicule at dorsolateral lobes of conjunctiva smoothly curved in middle (
<figureCitation id="28CE2A25FFED0E43FEA2FBBBFE07FBF5" box="[279,390,1030,1053]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="31" pageNumber="32">
Figure 7
<collectionCode id="D6E4AE65FFED0E43FEC7FBBBFE07FBF5" box="[370,390,1030,1053]" pageId="31" pageNumber="32">H</collectionCode>
</figureCitation>
). Posterior pouches of gynatrial sac shorter than in nominotypical subspecies, as long as wide or slightly longer than wide, not narrowed at base, lying on posterior margin of gynatrial sac, with inner margins passing into the posterior margin (
<figureCitation id="28CE2A25FFED0E43FE1BFB38FDAFFB74" box="[430,558,1157,1180]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="31" pageNumber="32">
Figures 10
<collectionCode id="D6E4AE65FFED0E43FDA9FB38FDAFFB74" box="[540,558,1157,1180]" pageId="31" pageNumber="32">C</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFED0E43FD89FB38FDE8FB74" box="[572,617,1157,1180]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="31" pageNumber="32">
12
<collectionCode id="D6E4AE65FFED0E43FDE0FB38FDE8FB74" box="[597,617,1157,1180]" pageId="31" pageNumber="32">D</collectionCode>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E43FF2BFB54FE76F92E" pageId="31" pageNumber="32" type="description">
<paragraph id="B04A36A0FFED0E43FF2BFB54FE96FAE8" blockId="31.[158,781,193,1938]" box="[158,279,1257,1280]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FF2BFB54FE96FAE8" bold="true" box="[158,279,1257,1280]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FF2BFB54FE96FAE8" bold="true" box="[158,279,1257,1280]" pageId="31" pageNumber="32">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FF2BFAACFE76F92E" blockId="31.[158,781,193,1938]" pageId="31" pageNumber="32">
Proximal part of spermathecal duct shorter than or as long as, and sometimes (several specimens from Gegharkunik Prov.) longer than median part. In
<specimenCount id="A6F3FD29FFED0E43FE7FFAECFDCBFA80" box="[458,586,1361,1384]" count="2" pageId="31" pageNumber="32" type="female">two females</specimenCount>
from Aragats Mt. and Lchashen, posterior pouches of gynatrial sac only slightly shorter than in nominotypical subspecies; in remaining specimens they much shorter; in another female from Aragats Mt., posterior pouches extremely small. Both posterior pouches in
<specimenCount id="A6F3FD29FFED0E43FE88FA4DFE3DF9EF" box="[317,444,1520,1543]" count="2" pageId="31" pageNumber="32" type="female">two females</specimenCount>
from Tambovka and Lchashen, and only right pouch in female from Bakuriani with small shapeless patches of sclerotisation inside.
<specimenCount id="A6F3FD29FFED0E43FDFBF98DFD44F9AF" box="[590,709,1584,1607]" count="1" pageId="31" pageNumber="32" type="female">One female</specimenCount>
examined from China having brownish membrane corresponding to diagnosis of the new subspecies, and at the same time rather long posterior pouches with inner margins passing into posterior margin of gynatrial sac.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E43FF2BF8A9FBCEFC1E" pageId="31" pageNumber="32" type="discussion">
<paragraph id="B04A36A0FFED0E43FF2BF8A9FE84F8C3" blockId="31.[158,781,193,1938]" box="[158,261,1812,1835]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FF2BF8A9FE84F8C3" bold="true" box="[158,261,1812,1835]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FF2BF8A9FE84F8C3" bold="true" box="[158,261,1812,1835]" pageId="31" pageNumber="32">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FF2BF886FBCEFC1E" blockId="31.[158,781,193,1938]" lastBlockId="31.[844,1466,193,1938]" pageId="31" pageNumber="32">
The single female examined from Khabarovsk Territory completely matches the diagnosis of the new Transcaucasian subspecies. The taxonomic position of Far Eastern populations of
<taxonomicName id="77F54D23FFED0E43FCF9FF7FFC55FF31" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[844,980,194,217]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFED0E43FCF9FF7FFC55FF31" box="[844,980,194,217]" italics="true" pageId="31" pageNumber="32">C. impressus</emphasis>
</taxonomicName>
needs to be clarified, but as yet I place this specimen together with the Chinese female, partially corresponding to the subspecific diagnosis, in the new subspecies that can be polytopic. On the other hand, it is impossible to exclude erroneous labelling of these
<specimenCount id="A6F3FD29FFED0E43FB74FEFCFAD9FEB0" box="[1217,1368,321,344]" count="2" pageId="31" pageNumber="32" type="generic">two specimens</specimenCount>
collected more than 100 years ago. The following data may be indirect evidence of mislabelling of these
<specimenCount id="A6F3FD29FFED0E43FB16FE3CFABFFE70" box="[1187,1342,385,408]" count="2" pageId="31" pageNumber="32" type="generic">two specimens</specimenCount>
. Judging by the original labels from cotton pads, specimens from the vicinity of “Kizilziarat” Mt. and from the banks of the Khor Riv. were collected by different persons, by Ya.G. Efremov in the first case and by N.E. Efremov in the second case; based on the registration numbers on the original labels, these specimens were received in the
<collectionCode id="D6E4AE65FFED0E43FBC3FDFDFB2BFDBF" box="[1142,1194,576,599]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="31" pageNumber="32">ZISP</collectionCode>
in 1910; judging by the identity of the paper, handwriting and ink of the labels under all these specimens, they were mounted simultaneously by one preparator who may have confused the labels, and pinned under
<specimenCount id="A6F3FD29FFED0E43FC2CFD7DFBB0FD3F" box="[921,1073,704,727]" count="1" pageId="31" pageNumber="32" type="generic">one specimen</specimenCount>
from Armenia the label for the Khabarovsk Territory, because of the identical surnames of the collectors. The doubt about the correctness of labelling in the case of the specimen from Sichuan is based on the fact that no more specimens of
<taxonomicName id="77F54D23FFED0E43FB89FC82FB43FCBE" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[1084,1218,831,854]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFED0E43FB89FC82FB43FCBE" box="[1084,1218,831,854]" italics="true" pageId="31" pageNumber="32">C. impressus</emphasis>
</taxonomicName>
are known from northwestern, northern or north-eastern territories of China, and the distances to the nearest localities where the species was found in the north-east of Mongolia and in eastern Kazakhstan and Kyrgyzstan are more than 2000 and about
<quantity id="770D9B45FFED0E43FA97FC02FA03FC3E" box="[1314,1410,959,982]" metricMagnitude="6" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="km" value="2500.0">2500 km</quantity>
in a straight line, respectively.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E43FCF9FB85FAACFB3E" pageId="31" pageNumber="32" type="description">
<paragraph id="B04A36A0FFED0E43FCF9FB85FC70FBA7" blockId="31.[844,1466,193,1938]" box="[844,1009,1080,1103]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FCF9FB85FC70FBA7" bold="true" box="[844,1009,1080,1103]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FCF9FB85FC70FBA7" bold="true" box="[844,1009,1080,1103]" pageId="31" pageNumber="32">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FCF9FBDDFAACFB3E" blockId="31.[844,1466,193,1938]" pageId="31" pageNumber="32">
Body
<quantity id="770D9B45FFED0E43FC3AFBDDFB96FB9F" box="[911,1047,1119,1143]" metricMagnitude="-1" metricUnit="m" metricValue="1.50495" metricValueMax="1.8161" metricValueMin="1.1938" pageId="31" pageNumber="32" unit="in" value="5.925" valueMax="7.15" valueMin="4.7">4.70–7.15 in</quantity>
length,
<quantity id="770D9B45FFED0E43FBC6FBDDFB7AFB9F" box="[1139,1275,1119,1143]" metricMagnitude="-2" metricUnit="m" metricValue="9.398000000000001" metricValueMax="10.921999999999999" metricValueMin="7.8740000000000006" pageId="31" pageNumber="32" unit="in" value="3.7" valueMax="4.3" valueMin="3.1">3.10–4.30 in</quantity>
width; pronotum
<quantity id="770D9B45FFED0E43FCF9FB3DFC50FB7F" box="[844,977,1151,1175]" metricMagnitude="-2" metricUnit="m" metricValue="8.064499999999999" metricValueMax="9.652" metricValueMin="6.476999999999999" pageId="31" pageNumber="32" unit="in" value="3.175" valueMax="3.8" valueMin="2.55">2.55–3.80 in</quantity>
width; head
<quantity id="770D9B45FFED0E43FBE8FB3DFB62FB7F" box="[1117,1251,1151,1175]" metricMagnitude="-2" metricUnit="m" metricValue="3.5814" metricValueMax="3.9878" metricValueMin="3.175" pageId="31" pageNumber="32" unit="in" value="1.41" valueMax="1.57" valueMin="1.25">1.25–1.57 in</quantity>
width; ocular index 3.27–3.60; length of antennal segments I–V: 0.27–0.35: 0.35–0.55: 0.38–0.57: 0.53–0.77: 0.64–0.87.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E43FCF9FAA5FA3BFA7F" pageId="31" pageNumber="32" type="etymology">
<paragraph id="B04A36A0FFED0E43FCF9FAA5FC47FAC7" blockId="31.[844,1466,193,1938]" box="[844,966,1304,1327]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FCF9FAA5FC47FAC7" bold="true" box="[844,966,1304,1327]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FCF9FAA5FC47FAC7" bold="true" box="[844,966,1304,1327]" pageId="31" pageNumber="32">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FCF9FAFDFA3BFA7F" blockId="31.[844,1466,193,1938]" pageId="31" pageNumber="32">
The name is derived from the Latin
<emphasis id="8281EAB2FFED0E43FB6DFAFDFACCFABF" box="[1240,1357,1344,1367]" italics="true" pageId="31" pageNumber="32">Hajastania</emphasis>
, denoting Armenia which included the largest part of the subspecies range in antiquity after the fall of the Achaemenid Empire.
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<emphasis id="8281EAB2FFED0E43FCF9FA64FC54FA18" bold="true" box="[844,981,1497,1520]" pageId="31" pageNumber="32">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E43FCF9F9BDFB49F9DF" blockId="31.[844,1466,193,1938]" pageId="31" pageNumber="32">Figure 15. Georgia, Armenia, Azerbaijan, Turkey, Russian Far East[?], and China, Sichuan[?].</paragraph>
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<emphasis id="8281EAB2FFED0E43FC19F9C5FB2BF967" bold="true" box="[940,1194,1656,1679]" italics="true" pageId="31" pageNumber="32">Canthophorus wagneri</emphasis>
<bibRefCitation id="D4644B51FFED0E43FB06F9C5FAD8F967" author="Asanova RB" box="[1203,1369,1656,1679]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="31" pageNumber="32" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
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(
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Figures 3
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5
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6
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<figureCitation id="28CE2A25FFED0E43FB18F925FA83F947" box="[1197,1282,1688,1711]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="31" pageNumber="32">
8
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<location id="B52A607BFFED0E43FBD7F961FB24F919" LSID="urn:lsid:plazi:treatment:385C87B6FFED0E7CFC19F9C5FB41FF11:B52A607BFFED0E43FBD7F961FB24F919" box="[1122,1189,1756,1777]" country="Russia" name="Anapa" pageId="31" pageNumber="32" stateProvince="Krasnodar Terr">Anapa</location>
]
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.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFED0E7CFCF9F88EFB41FF11" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
<paragraph id="B04A36A0FFED0E43FCF9F88EFC4FF8A2" blockId="31.[844,1466,193,1938]" box="[844,974,1843,1866]" pageId="31" pageNumber="32">
<heading id="EB0281CCFFED0E43FCF9F88EFC4FF8A2" bold="true" box="[844,974,1843,1866]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="6">
<emphasis id="8281EAB2FFED0E43FCF9F88EFC4FF8A2" bold="true" box="[844,974,1843,1866]" pageId="31" pageNumber="32">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFED0E7CFCF9F8E6FEA1FEF0" blockId="31.[844,1466,193,1938]" lastBlockId="32.[158,781,193,1939]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">Lateral margin of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metapleural evaporatorium with long and curved peritreme. Membrane brownish, opaque.</paragraph>
<paragraph id="B04A36A0FFD20E7CFF15FE8CFDC5FE80" blockId="32.[158,781,193,1939]" pageId="32" pageNumber="33">
<emphasis id="8281EAB2FFD20E7CFF15FE8CFE9BFEA0" bold="true" box="[160,282,305,328]" pageId="32" pageNumber="33">Pygophore</emphasis>
slightly longer than wide, with dorsal infolding and dorsal wall as in
<taxonomicName id="77F54D23FFD20E7CFE06FEECFDBCFE80" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[435,573,337,360]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFD20E7CFE06FEECFDBCFE80" box="[435,573,337,360]" italics="true" pageId="32" pageNumber="33">C. impressus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFD20E7CFF15FE3CFD43FE50" blockId="32.[158,781,193,1939]" pageId="32" pageNumber="33">
<emphasis id="8281EAB2FFD20E7CFF15FE3CFE8EFE70" bold="true" box="[160,271,385,408]" pageId="32" pageNumber="33">Paramere</emphasis>
as in previous species, with rather short, apically widely rounded sensory process (
<figureCitation id="28CE2A25FFD20E7CFDF1FE1CFD33FE50" box="[580,690,417,440]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="32" pageNumber="33">
Figure 3
<collectionCode id="D6E4AE65FFD20E7CFD28FE1CFD33FE50" box="[669,690,417,440]" pageId="32" pageNumber="33">H</collectionCode>
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).
</paragraph>
<paragraph id="B04A36A0FFD20E7CFF15FE6CFD11FA6C" blockId="32.[158,781,193,1939]" pageId="32" pageNumber="33">
<emphasis id="8281EAB2FFD20E7CFF15FE6CFE8CFE00" bold="true" box="[160,269,465,488]" pageId="32" pageNumber="33">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFD20E7CFE9FFE6CFE20FE00" box="[298,417,465,488]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="32" pageNumber="33">
Figure 5
<collectionCode id="D6E4AE65FFD20E7CFE3AFE6CFE20FE00" box="[399,417,465,488]" pageId="32" pageNumber="33">B</collectionCode>
</figureCitation>
). Ventral lobe of conjunctiva extending to but not protruding beyond apical margin of theca; in ventral view, lobe narrowed at base, widening apically, at extreme apex rather strongly or slightly widened, rounded, with short carina on dorsal surface and impressions at sides of it (
<figureCitation id="28CE2A25FFD20E7CFE4DFDCDFD11FD6F" box="[504,656,624,647]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="32" pageNumber="33">
Figure 6
<collectionCode id="D6E4AE65FFD20E7CFDE6FDCDFDE0FD6F" box="[595,609,624,647]" pageId="32" pageNumber="33">P</collectionCode>
, 6
<collectionCode id="D6E4AE65FFD20E7CFDCBFDCDFD11FD6F" box="[638,656,624,647]" pageId="32" pageNumber="33">R</collectionCode>
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); in lateral view, lobe narrow (
<figureCitation id="28CE2A25FFD20E7CFECFFD2DFD9BFD4F" box="[378,538,656,679]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="32" pageNumber="33">
Figure 6
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, 6
<collectionCode id="D6E4AE65FFD20E7CFDB9FD2DFD9BFD4F" box="[524,538,656,679]" pageId="32" pageNumber="33">S</collectionCode>
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). Apical part of conjunctiva slightly shorter than in previous species. Apical lobes of conjunctiva short, lying in dorsoventral plane of apical part of conjunctiva, at bases directed laterad and curved posteriorly in middle of length. Vesica slightly shorter than in previous species, without lateral branch. Dorsolateral lobes long and rather wide, straight, spaced basally, diverging in a V-shape, directed dorsolaterad and slightly posteriad. Anterior walls of dorsolateral lobes with weak sclerotised areas before apices. Each spicule extending from posterior wall of lobe apex, at base directed anteriad and slightly dorsad, apically directed mediad. Spicule long, sharply curved C-shaped and rather widened at base, having almost parallel margins slightly narrowing apicad in remaining part, and arcuately curving in varying degrees, with weak S-shaped bend before apex (outer margin of spicule here slightly concave, inner margin slightly convex); extreme apex of spicule sharply curved inwards and narrowly rounded (
<figureCitation id="28CE2A25FFD20E7CFDB9FB73FD21FB0D" box="[524,672,1230,1253]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="32" pageNumber="33">
Figure
<date id="C44B1060FFD20E7CFDEAFB73FDF5FB0D" box="[607,628,1230,1253]" pageId="32" pageNumber="33">
7
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</date>
, 7
<collectionCode id="D6E4AE65FFD20E7CFD20FB73FD21FB0D" box="[661,672,1230,1253]" pageId="32" pageNumber="33">J</collectionCode>
</figureCitation>
). Spicule basal plate long, occupying entire apex of lobe, at anterior end with one large, laterally flattened denticle curved at right angle and directed mediad by rectangularly rounded apex (
<figureCitation id="28CE2A25FFD20E7CFF56FAF0FEC9FA8C" box="[227,328,1357,1380]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="32" pageNumber="33">
Figure
<date id="C44B1060FFD20E7CFE87FAF0FEC9FA8C" box="[306,328,1357,1380]" pageId="32" pageNumber="33">
7
<collectionCode id="D6E4AE65FFD20E7CFE8AFAF0FEC9FA8C" box="[319,328,1357,1380]" pageId="32" pageNumber="33">I</collectionCode>
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</figureCitation>
) or with more or less spear-shaped apex (
<figureCitation id="28CE2A25FFD20E7CFF12FAD0FE8DFA6C" box="[167,268,1389,1412]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="32" pageNumber="33">
Figure 7
<collectionCode id="D6E4AE65FFD20E7CFEB4FAD0FE8DFA6C" box="[257,268,1389,1412]" pageId="32" pageNumber="33">J</collectionCode>
</figureCitation>
) directed dorsolaterad and anteriad.
</paragraph>
<paragraph id="B04A36A0FFD20E7CFF15FA20FB41FF11" blockId="32.[158,781,193,1939]" lastBlockId="32.[843,1471,194,1938]" pageId="32" pageNumber="33">
<emphasis id="8281EAB2FFD20E7CFF15FA20FE9FFA5C" bold="true" box="[160,286,1437,1460]" pageId="32" pageNumber="33">Gynatrium</emphasis>
(
<figureCitation id="28CE2A25FFD20E7CFE87FA20FE30FA5C" box="[306,433,1437,1460]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="32" pageNumber="33">
Figure 10
<collectionCode id="D6E4AE65FFD20E7CFE28FA20FE30FA5C" box="[413,433,1437,1460]" pageId="32" pageNumber="33">D</collectionCode>
</figureCitation>
). Posterior pouches of gynatrial sac lying beyond posterior margin of latter, wide, of same width all over and rather long, about 2.5–3 times as long as wide; they longer and wider than in
<taxonomicName id="77F54D23FFD20E7CFDBEFA40FD0FF9FC" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[523,654,1533,1556]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFD20E7CFDBEFA40FD0FF9FC" box="[523,654,1533,1556]" italics="true" pageId="32" pageNumber="33">C. impressus</emphasis>
</taxonomicName>
, shorter and wider than in
<taxonomicName id="77F54D23FFD20E7CFE85F9A0FE13F9DC" authorityName="Asanova" authorityYear="1964" box="[304,402,1565,1588]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFD20E7CFE85F9A0FE13F9DC" box="[304,402,1565,1588]" italics="true" pageId="32" pageNumber="33">C. mixtus</emphasis>
</taxonomicName>
. Pouches directed posterolaterad; in dorsal view, they are straight at all lengths, but sometimes lyrately curved at apices (in female from Gokcha,
<figureCitation id="28CE2A25FFD20E7CFF2BF9C1FE93F97B" box="[158,274,1660,1683]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="32" pageNumber="33">
Figure 10
<collectionCode id="D6E4AE65FFD20E7CFEB4F9C1FE93F97B" box="[257,274,1660,1683]" pageId="32" pageNumber="33">E</collectionCode>
</figureCitation>
). In posterolateral view, their dorsal wall convex at base (
<figureCitation id="28CE2A25FFD20E7CFF48F921FEF4F95B" box="[253,373,1692,1715]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="32" pageNumber="33">
Figure 12
<collectionCode id="D6E4AE65FFD20E7CFED1F921FEF4F95B" box="[356,373,1692,1715]" pageId="32" pageNumber="33">E</collectionCode>
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). Posterior wall of gynatrial sac near outer margin of ring sclerites with convex areas clearly visible near base of each pouch in dorsal and slightly posterior view. Anterior part of gynatrial sac large, slightly smaller than in
<taxonomicName id="77F54D23FFD20E7CFF45F8A1FED7F8DB" box="[240,342,1820,1843]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="dubius">
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. Basal part of spermathecal duct short, distal part slightly shorter than half of middle part of duct. Proximal part of spermathecal capsule about 1–2 times as long as transverse diameter of distal part. First rami strongly widened anteriorly, with obliquely truncated anterior end (
<figureCitation id="28CE2A25FFD20E7CFBA1FF5FFB31FF11" box="[1044,1200,226,249]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="32" pageNumber="33">
Figure 8
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).
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<emphasis id="8281EAB2FFEE0E40FF50FC60FDF5FC1C" bold="true" box="[229,628,989,1012]" italics="true" pageId="28" pageNumber="29">Canthophorus melanopterus mariae</emphasis>
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,
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amsl,
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,
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(Gussakovskij), 1J,
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),
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(
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)
</materialsCitation>
;
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.,
<collectingMunicipality id="502EACDAFFEE0E40FE04FA80FDC5FABC" box="[433,580,1341,1364]" pageId="28" pageNumber="29">Gissar Range</collectingMunicipality>
,
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, vicinity of “Romt” [
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],
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(
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), 3J,
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(
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),
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(
<collectionCode id="D6E4AE65FFEE0E40FE34FAC1FE59FA7B" box="[385,472,1404,1427]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEE0E40FF2BFA68FBB8FEF0" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="B04A36A0FFEE0E40FF2BFA68FE8AFA04" blockId="28.[158,780,193,1938]" box="[158,267,1493,1516]" pageId="28" pageNumber="29">
<heading id="EB0281CCFFEE0E40FF2BFA68FE8AFA04" bold="true" box="[158,267,1493,1516]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="6">
<emphasis id="8281EAB2FFEE0E40FF2BFA68FE8AFA04" bold="true" box="[158,267,1493,1516]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FF2BFA40FBB8FEF0" blockId="28.[158,780,193,1938]" lastBlockId="28.[844,1466,194,1938]" pageId="28" pageNumber="29">
Yellowish rib on lateral margins of pronotum and hemelytra thin, narrower than base of antennal segment II; proximal tubular part of spermathecal capsule approximately 3–4 times as long as transverse diameter of distal spherical part of capsule (
<figureCitation id="28CE2A25FFEE0E40FEA8F9C1FE0BF97B" box="[285,394,1660,1683]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="28" pageNumber="29">
Figure 9
<collectionCode id="D6E4AE65FFEE0E40FEC2F9C1FE0BF97B" box="[375,394,1660,1683]" pageId="28" pageNumber="29">D</collectionCode>
</figureCitation>
); posterior pouches of gynatrial sac posteriorly strongly shortened; sclerotisation inside posterior pouches shaped as narrow, arcuately curved band with irregular margins, one end directed lateroposteriad and opposite end directed medioanteriad (
<figureCitation id="28CE2A25FFEE0E40FE2CF941FD84F8FB" box="[409,517,1788,1811]" captionStart="Figure 14" captionStartId="20.[158,225,1738,1759]" captionTargetBox="[300,1351,177,1709]" captionTargetId="figure-80@20.[283,1370,164,1722]" captionTargetPageId="20" captionText="Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927045" httpUri="https://zenodo.org/record/14927045/files/figure.png" pageId="28" pageNumber="29">
Figure
<date id="C44B1060FFEE0E40FE56F941FD84F8FB" box="[483,517,1788,1811]" pageId="28" pageNumber="29" value="2014-01">
14
<collectionCode id="D6E4AE65FFEE0E40FE49F941FD84F8FB" box="[508,517,1788,1811]" pageId="28" pageNumber="29">I</collectionCode>
</date>
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), or sclerotisation almost lost except for very thin, weakly sclerotised plate along anterior inner wall of pouch and visible as very thin band in dorsal view (as in form 6 of
<emphasis id="8281EAB2FFEE0E40FECAF8E6FDC6F89A" box="[383,583,1883,1906]" italics="true" pageId="28" pageNumber="29">
C. m.
<taxonomicName id="77F54D23FFEE0E40FE0BF8E6FDC6F89A" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[446,583,1883,1906]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="melanopterus">melanopterus</taxonomicName>
</emphasis>
). The new subspecies is also distinguished from
<emphasis id="8281EAB2FFEE0E40FDA1F8C6FD55F87A" box="[532,724,1915,1938]" italics="true" pageId="28" pageNumber="29">C. m. contrarius</emphasis>
and Mediterranean specimens of
<emphasis id="8281EAB2FFEE0E40FB31FF7FFAD9FF31" box="[1156,1368,194,217]" italics="true" pageId="28" pageNumber="29">
C. m.
<taxonomicName id="77F54D23FFEE0E40FB7BFF7FFAD9FF31" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[1230,1368,194,217]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="melanopterus">melanopterus</taxonomicName>
</emphasis>
by long lateral branch of vesica, and from
<emphasis id="8281EAB2FFEE0E40FB16FF5CFAC8FF10" box="[1187,1353,225,248]" italics="true" pageId="28" pageNumber="29">C. m. contrarius</emphasis>
by brownish, opaque membrane.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEE0E40FCF9FED0FAACFB15" pageId="28" pageNumber="29" type="description">
<paragraph id="B04A36A0FFEE0E40FCF9FED0FC44FE6C" blockId="28.[844,1466,194,1938]" box="[844,965,365,388]" pageId="28" pageNumber="29">
<heading id="EB0281CCFFEE0E40FCF9FED0FC44FE6C" bold="true" box="[844,965,365,388]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="6">
<emphasis id="8281EAB2FFEE0E40FCF9FED0FC44FE6C" bold="true" box="[844,965,365,388]" pageId="28" pageNumber="29">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9FE28FB92FBE2" blockId="28.[844,1466,194,1938]" pageId="28" pageNumber="29">
In
<specimenCount id="A6F3FD29FFEE0E40FCC5FE28FC65FE44" box="[880,996,405,428]" count="2" pageId="28" pageNumber="29" type="male">two males</specimenCount>
from Kondara Gorge, distal denticle of sclerite basal plate at dorsolateral lobes of conjunctiva wider than in other examined specimens of the species (
<figureCitation id="28CE2A25FFEE0E40FCE1FE49FC43FDE3" box="[852,962,500,523]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="28" pageNumber="29">
Figure 7
<collectionCode id="D6E4AE65FFEE0E40FC07FE49FC43FDE3" box="[946,962,500,523]" pageId="28" pageNumber="29">F</collectionCode>
</figureCitation>
). Among specimens from vicinity of Romit,
<specimenCount id="A6F3FD29FFEE0E40FCF9FDA9FC44FDC3" box="[844,965,532,555]" count="1" pageId="28" pageNumber="29" type="female">one female</specimenCount>
with posterior pouches having in addition to arcuate sclerites, also a pair of large, formless longitudinal sclerites lying along anterodorsal margin of pouch and connected with posterior ends of arcuate sclerites; in another female, arcuate sclerites very narrow, connected by posterior end with very narrow sclerotised band lying along anterodorsal margin of pouch; in remaining females, arcuate sclerites rather wide and sclerotised band along anterodorsal margin of pouch poorly noticeable; in
<specimenCount id="A6F3FD29FFEE0E40FB13FC8EFAA3FCA2" box="[1190,1314,819,842]" count="1" pageId="28" pageNumber="29" type="female">one female</specimenCount>
, right arcuate sclerite shorter and narrower than left sclerite; in
<specimenCount id="A6F3FD29FFEE0E40FA24FCEEFC3EFC62" count="2" pageId="28" pageNumber="29" type="generic">two specimens</specimenCount>
, right arcuate sclerite large but weakly sclerotised, left arcuate sclerite absent. In
<specimenCount id="A6F3FD29FFEE0E40FA84FC2EFA38FC42" box="[1329,1465,915,938]" count="2" pageId="28" pageNumber="29" type="female">two females</specimenCount>
from Kondara Gorge, arcuate sclerites absent and sclerotised bands along anterodorsal margins of pouches poorly noticeable.
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9FBE3FC70FB9D" blockId="28.[844,1466,194,1938]" box="[844,1009,1118,1141]" pageId="28" pageNumber="29">
<heading id="EB0281CCFFEE0E40FCF9FBE3FC70FB9D" bold="true" box="[844,1009,1118,1141]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="6">
<emphasis id="8281EAB2FFEE0E40FCF9FBE3FC70FB9D" bold="true" box="[844,1009,1118,1141]" pageId="28" pageNumber="29">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9FB3BFAACFB15" blockId="28.[844,1466,194,1938]" pageId="28" pageNumber="29">
Body
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length,
<quantity id="770D9B45FFEE0E40FBCEFB3BFB79FB75" box="[1147,1272,1158,1182]" metricMagnitude="-1" metricUnit="m" metricValue="1.03505" metricValueMax="1.2192" metricValueMin="0.8509" pageId="28" pageNumber="29" unit="in" value="4.075" valueMax="4.8" valueMin="3.35">3.35–4.8 in</quantity>
width; pronotum
<quantity id="770D9B45FFEE0E40FCF9FB1BFC50FB55" box="[844,977,1189,1213]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.033" metricValueMin="7.747" pageId="28" pageNumber="29" unit="in" value="3.5" valueMax="3.95" valueMin="3.05">3.05–3.95 in</quantity>
width; head
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width; ocular index 3.76–5.64; length of antennal segments I–V: 0.31–0.39: 0.49–0.60: 0.56–0.70: 0.80–0.98: 0.91–1.06.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEE0E40FCF9FAECFAE8FA18" pageId="28" pageNumber="29" type="etymology">
<paragraph id="B04A36A0FFEE0E40FCF9FAECFC47FA80" blockId="28.[844,1466,194,1938]" box="[844,966,1361,1384]" pageId="28" pageNumber="29">
<heading id="EB0281CCFFEE0E40FCF9FAECFC47FA80" bold="true" box="[844,966,1361,1384]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="6">
<emphasis id="8281EAB2FFEE0E40FCF9FAECFC47FA80" bold="true" box="[844,966,1361,1384]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9FAC4FAE8FA18" blockId="28.[844,1466,194,1938]" pageId="28" pageNumber="29">
The subspecies is named after Maria Berlina, the senior custodian of the Rhynchota collection in
<collectionCode id="D6E4AE65FFEE0E40FB4BFA24FAB5FA58" box="[1278,1332,1433,1456]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="28" pageNumber="29">ZISP</collectionCode>
, who helped me in dissection of the female terminalia and was the first to see the diagnostic characters of this new taxon.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEE0E40FCF9F9F9FB5DF96B" box="[844,1244,1604,1667]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="B04A36A0FFEE0E40FCF9F9F9FC54F9B3" blockId="28.[844,1466,194,1938]" box="[844,981,1604,1627]" pageId="28" pageNumber="29">
<heading id="EB0281CCFFEE0E40FCF9F9F9FC54F9B3" bold="true" box="[844,981,1604,1627]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="6">
<emphasis id="8281EAB2FFEE0E40FCF9F9F9FC54F9B3" bold="true" box="[844,981,1604,1627]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9F9D1FB5DF96B" blockId="28.[844,1466,194,1938]" box="[844,1244,1644,1667]" pageId="28" pageNumber="29">
<figureCitation id="28CE2A25FFEE0E40FCF9F9D1FC35F96B" box="[844,948,1644,1667]" captionStart="Figure 16" captionStartId="24.[133,200,1361,1382]" captionTargetBox="[137,1877,713,1324]" captionTargetId="figure-1@24.[137,1878,713,1324]" captionTargetPageId="24" captionText="Figure 16. Geographical variability of male and female terminalia in Canthophorus melanopterus (Herrich-Schaeffer, 1835). 1–6 in legend correspond to numbers of forms 1–6 by female terminalia in C. m. melanopterus; 7, females of C. m. mariae n. ssp.; 8, males with long lateral branch of vesica; 9, males with short lateral branch of vesica." figureDoi="http://doi.org/10.5281/zenodo.14927049" httpUri="https://zenodo.org/record/14927049/files/figure.png" pageId="28" pageNumber="29">Figure 16</figureCitation>
. Tajikistan (Gissar Range).
</paragraph>
</subSubSection>
</treatment>
</document>

274
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<mods:date id="DE198EA51B98B34F0C310C9EB5233AE4">2018</mods:date>
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<taxonomicName id="77F54D23FFEE0E40FC2FF979FAEAF933" authority="(Horvath, 1880)" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[922,1387,1732,1755]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="impressus">
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<emphasis id="8281EAB2FFEE0E40FC2FF979FB2DF933" bold="true" box="[922,1196,1732,1755]" italics="true" pageId="28" pageNumber="29">Canthophorus impressus</emphasis>
(
<bibRefCitation id="D4644B51FFEE0E40FB08F979FAE3F933" author="Horvath G." box="[1213,1378,1732,1755]" journalOrPublisher="Termeszetrajzi Fuzetek." pageId="28" pageNumber="29" pagination="184 - 192" part="4" refId="ref40056" refString="Horvath G. 1880. Hemipterologiai kozlemenyek. Termeszetrajzi Fuzetek. 4: 184 - 192." title="Hemipterologiai kozlemenyek" type="journal article" year="1880">Horváth, 1880</bibRefCitation>
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<paragraph id="B04A36A0FFEE0E40FCE0F959FB78F8F3" blockId="28.[844,1466,194,1938]" pageId="28" pageNumber="29">
<emphasis id="8281EAB2FFEE0E40FCE0F959FB78F8F3" bold="true" pageId="28" pageNumber="29">
(
<figureCitation id="28CE2A25FFEE0E40FCE8F959FB85F913" box="[861,1028,1764,1787]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="28" pageNumber="29">
Figures 1
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<figureCitation id="28CE2A25FFEE0E40FBA1F959FBE1F913" box="[1044,1120,1764,1787]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="28" pageNumber="29">
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<figureCitation id="28CE2A25FFEE0E40FBDAF959FB0EF913" box="[1135,1167,1764,1787]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="28" pageNumber="29">
5
<collectionCode id="D6E4AE65FFEE0E40FBC9F959FB0EF913" box="[1148,1167,1764,1787]" pageId="28" pageNumber="29">A</collectionCode>
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<figureCitation id="28CE2A25FFEE0E40FB2AF959FB70F913" box="[1183,1265,1764,1787]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="28" pageNumber="29">
6
<collectionCode id="D6E4AE65FFEE0E40FB19F959FB41F913" box="[1196,1216,1764,1787]" pageId="28" pageNumber="29">N</collectionCode>
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<figureCitation id="28CE2A25FFEE0E40FAB5F959FAD5F913" box="[1280,1364,1764,1787]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="28" pageNumber="29">
7
<collectionCode id="D6E4AE65FFEE0E40FABBF959FAA2F913" box="[1294,1315,1764,1787]" pageId="28" pageNumber="29">G</collectionCode>
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<figureCitation id="28CE2A25FFEE0E40FAD6F959FA2BF913" box="[1379,1450,1763,1787]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="28" pageNumber="29">
8
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<figureCitation id="28CE2A25FFEE0E40FBB8F8B9FBF4F8F3" box="[1037,1141,1795,1819]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="28" pageNumber="29">
10
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<figureCitation id="28CE2A25FFEE0E40FB30F8B9FB6EF8F3" box="[1157,1263,1796,1819]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="28" pageNumber="29">
12
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)
</emphasis>
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<taxonomicName id="77F54D23FFEE0E40FCF9F890FB30F8AA" authority="Horvath 1880: 184" authorityName="Horvath" authorityPageNumber="184" authorityYear="1880" box="[844,1201,1837,1858]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFEE0E40FCF9F890FC76F8AA" box="[844,1015,1837,1858]" italics="true" pageId="28" pageNumber="29">Sehirus impressus</emphasis>
<bibRefCitation id="D4644B51FFEE0E40FC48F890FB30F8AA" author="Horvath G." box="[1021,1201,1837,1858]" journalOrPublisher="Termeszetrajzi Fuzetek." pageId="28" pageNumber="29" pagination="184 - 192" part="4" refId="ref40056" refString="Horvath G. 1880. Hemipterologiai kozlemenyek. Termeszetrajzi Fuzetek. 4: 184 - 192." title="Hemipterologiai kozlemenyek" type="journal article" year="1880">Horváth 1880: 184</bibRefCitation>
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[
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:
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,
<collectingCountry id="C8E27630FFEE0E40FADBF890FA37F8AA" box="[1390,1462,1837,1858]" name="Austria" pageId="28" pageNumber="29" value="Austria">Austria</collectingCountry>
, Carinthia].
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<paragraph id="B04A36A0FFEE0E40FCF9F8DFFA8BF89F" blockId="28.[844,1466,194,1938]" box="[844,1290,1890,1911]" pageId="28" pageNumber="29">
<taxonomicName id="77F54D23FFEE0E40FCF9F8DFFBBBF89F" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[844,1082,1890,1911]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="8281EAB2FFEE0E40FCF9F8DFFBBBF89F" box="[844,1082,1890,1911]" italics="true" pageId="28" pageNumber="29">Canthophorus impressus</emphasis>
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:
<bibRefCitation id="D4644B51FFEE0E40FBFCF8DFFA87F89F" author="Signoret V." box="[1097,1286,1890,1911]" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="28" pageNumber="29" pagination="621 - 657" part="16" refId="ref40727" refString="Signoret V. 1881. Revue des Cydnides contenns dans la collection du Musee Civique d' Histoire Naturelle de Genes. Annali del Museo Civico di Storia Naturale di Genova. 16: 621 - 657." title="Revue des Cydnides contenns dans la collection du Musee Civique d' Histoire Naturelle de Genes" type="book chapter" year="1881">Signoret 1881: 657</bibRefCitation>
.
</paragraph>
<paragraph id="B04A36A0FFEE0E40FCF9F8C0FA02F87A" blockId="28.[844,1466,194,1938]" box="[844,1411,1917,1938]" pageId="28" pageNumber="29">
?
<taxonomicName id="77F54D23FFEE0E40FCE3F8C0FA01F87A" authority="Signoret 1884: 56" authorityName="Signoret" authorityPageNumber="56" authorityYear="1884" box="[854,1408,1917,1938]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="variety" species="dubius" variety="brevipennis">
<emphasis id="8281EAB2FFEE0E40FCE3F8C0FBA4F87A" box="[854,1061,1917,1938]" italics="true" pageId="28" pageNumber="29">Canthophorus dubius</emphasis>
var.
<emphasis id="8281EAB2FFEE0E40FBECF8C0FB46F87A" box="[1113,1223,1917,1938]" italics="true" pageId="28" pageNumber="29">brevipennis</emphasis>
<bibRefCitation id="D4644B51FFEE0E40FB7BF8C0FA01F87A" author="Signoret V." box="[1230,1408,1917,1938]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="28" pageNumber="29" pagination="45 - 62" part="4" refId="ref40764" refString="Signoret V. 1884. Revision du Groupe des Cydnides de la Famille des Pentatomides. 13 e et derniere partie. Annales de la Societe entomologique de France (Ser. 6). 4: 45 - 62." title="Revision du Groupe des Cydnides de la Famille des Pentatomides. 13 e et derniere partie" type="book chapter" year="1884">Signoret 1884: 56</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEF0E41FF2BFF7CFAD8FD7F" pageId="29" pageNumber="30" type="description">
<paragraph id="B04A36A0FFEF0E41FF2BFF7CFEA1FF30" blockId="29.[158,781,193,1892]" box="[158,288,193,216]" pageId="29" pageNumber="30">
<heading id="EB0281CCFFEF0E41FF2BFF7CFEA1FF30" bold="true" box="[158,288,193,216]" fontSize="10" level="2" pageId="29" pageNumber="30" reason="6">
<emphasis id="8281EAB2FFEF0E41FF2BFF7CFEA1FF30" bold="true" box="[158,288,193,216]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEF0E41FF2BFF54FF71FE68" blockId="29.[158,781,193,1892]" pageId="29" pageNumber="30">Lateral margins of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metapleural evaporatorium with long and curved peritreme. Membrane brownish or whitish.</paragraph>
<paragraph id="B04A36A0FFEF0E41FF15FE24FDF4FE07" blockId="29.[158,781,193,1892]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF15FE24FE94FE58" bold="true" box="[160,277,409,432]" italics="true" pageId="29" pageNumber="30">Pygophore</emphasis>
as long as wide, with lateral walls less concave, with shorter, less concave dorsal infolding, and with longer dorsal wall than in
<taxonomicName id="77F54D23FFEF0E41FE08FE65FDEFFE07" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[445,622,472,495]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFEF0E41FE08FE65FDEFFE07" box="[445,622,472,495]" italics="true" pageId="29" pageNumber="30">C. melanopterus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFEF0E41FF2BFDB1FECFFD8A" blockId="29.[158,781,193,1892]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF2BFDB1FE91FDCB" bold="true" box="[158,272,524,547]" italics="true" pageId="29" pageNumber="30">Paramere.</emphasis>
Sensory process rather shorter and slightly more curved dorsally than in two previous species (
<figureCitation id="28CE2A25FFEF0E41FF12FDF6FEBEFD8A" box="[167,319,587,610]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="29" pageNumber="30">
Figure 3
<collectionCode id="D6E4AE65FFEF0E41FF4AFDF6FE8FFD8A" box="[255,270,587,610]" pageId="29" pageNumber="30">F</collectionCode>
, 3
<collectionCode id="D6E4AE65FFEF0E41FE9FFDF6FEBEFD8A" box="[298,319,587,610]" pageId="29" pageNumber="30">G</collectionCode>
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).
</paragraph>
<paragraph id="B04A36A0FFEF0E41FF2BFDC3FE34FA9B" blockId="29.[158,781,193,1892]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF2BFDC3FE18FD7D" bold="true" box="[158,409,638,661]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF2BFDC3FE89FD7D" bold="true" box="[158,264,638,661]" italics="true" pageId="29" pageNumber="30">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFEF0E41FEADFDC3FE08FD7D" box="[280,393,638,661]" captionStart="Figure 5" captionStartId="9.[158,225,1628,1649]" captionTargetBox="[190,1436,174,1608]" captionTargetId="figure-150@9.[190,1436,174,1610]" captionTargetPageId="9" captionText="Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927027" httpUri="https://zenodo.org/record/14927027/files/figure.png" pageId="29" pageNumber="30">
Figure 5
<collectionCode id="D6E4AE65FFEF0E41FEC0FDC3FE08FD7D" box="[373,393,638,661]" pageId="29" pageNumber="30">A</collectionCode>
</figureCitation>
).
</emphasis>
Ventral lobe of conjunctiva extending to but not protruding beyond apical margin of theca, rather broad, gradually and slightly narrowing apically in ventral view, at extreme apex rather broadly widened, with widely rounded lateral margins and sharply rounded lateroapical angles; apical margin of lobe with rather deep triangular notch and ventral surface of lobe near this notch with small triangular impression (
<figureCitation id="28CE2A25FFEF0E41FDD2FCE3FD58FC9D" box="[615,729,862,885]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="29" pageNumber="30">
Figure 6
<collectionCode id="D6E4AE65FFEF0E41FD70FCE3FD58FC9D" box="[709,729,862,885]" pageId="29" pageNumber="30">N</collectionCode>
</figureCitation>
); in lateral view, lobe narrow (
<figureCitation id="28CE2A25FFEF0E41FE0BFCC3FDAAFC7D" box="[446,555,894,917]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="29" pageNumber="30">
Figure 6
<collectionCode id="D6E4AE65FFEF0E41FDADFCC3FDAAFC7D" box="[536,555,894,917]" pageId="29" pageNumber="30">O</collectionCode>
</figureCitation>
). Curved apical part of conjunctiva slightly shorter than in two previous species. Apical lobes long, entirely lying in dorsoventral plane of apical part of conjunctiva, basally directed laterad and curved posteriad before middle. Vesica without lateral branch. Dorsolateral lobes short and rather narrow, with slightly spaced bases, separated by shallow depression, directed dorsolaterad and curved towards base of aedeagus. Walls of dorsolateral lobes without areas of sclerotisation at sides. Spicules at dorsolateral lobes moderately long, rather longer than in two previous species, but shorter than in remaining species, thin, slightly thickened at base and tapering to acute apices, smoothly or rather sharply arcuately curved in middle (
<figureCitation id="28CE2A25FFEF0E41FE14FAA1FDBFFADB" box="[417,574,1308,1331]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="29" pageNumber="30">
Figure 7
<collectionCode id="D6E4AE65FFEF0E41FE4FFAA1FD8FFADB" box="[506,526,1308,1331]" pageId="29" pageNumber="30">G</collectionCode>
, 7
<collectionCode id="D6E4AE65FFEF0E41FD9FFAA1FDBFFADB" box="[554,574,1308,1331]" pageId="29" pageNumber="30">H</collectionCode>
</figureCitation>
), and directed dorsoposteriad. Spicule basal plate rather short, without denticles (
<figureCitation id="28CE2A25FFEF0E41FEBFFAE1FE27FA9B" box="[266,422,1372,1395]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="29" pageNumber="30">
Figure 7
<collectionCode id="D6E4AE65FFEF0E41FED6FAE1FEF6FA9B" box="[355,375,1372,1395]" pageId="29" pageNumber="30">G</collectionCode>
, 7
<collectionCode id="D6E4AE65FFEF0E41FE27FAE1FE27FA9B" box="[402,422,1372,1395]" pageId="29" pageNumber="30">H</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="B04A36A0FFEF0E41FF2BFA32FAB6FDF0" blockId="29.[158,781,193,1892]" lastBlockId="29.[843,1466,194,1892]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF2BFA32FE37FA4E" bold="true" box="[158,438,1423,1446]" pageId="29" pageNumber="30">
<emphasis id="8281EAB2FFEF0E41FF2BFA32FE94FA4E" bold="true" box="[158,277,1423,1446]" italics="true" pageId="29" pageNumber="30">Gynatrium</emphasis>
(
<figureCitation id="28CE2A25FFEF0E41FE93FA32FE26FA4E" box="[294,423,1423,1446]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="29" pageNumber="30">
Figure 10
<collectionCode id="D6E4AE65FFEF0E41FE26FA32FE26FA4E" box="[403,423,1423,1446]" pageId="29" pageNumber="30">A</collectionCode>
</figureCitation>
).
</emphasis>
Posterior pouches of gynatrial sac lying anterior to or exactly on posterior margin of gynatrial sac, rather small, no more than 2 times longer than wide or as long as wide, with thick, widely rounded apical wall, and directed posteriad or lateroposteriad (
<figureCitation id="28CE2A25FFEF0E41FF47F993FE35F9AD" box="[242,436,1582,1605]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="29" pageNumber="30">
Figure 10
<collectionCode id="D6E4AE65FFEF0E41FED4F993FEF2F9AD" box="[353,371,1582,1605]" pageId="29" pageNumber="30">B</collectionCode>
, 10
<collectionCode id="D6E4AE65FFEF0E41FE17F993FE35F9AD" box="[418,436,1582,1605]" pageId="29" pageNumber="30">C</collectionCode>
</figureCitation>
). Dorsal walls of pouches at base not convex; interior walls without distinct sclerites, very rarely with only small shapeless areas of sclerotisation (as shown at
<figureCitation id="28CE2A25FFEF0E41FE33F933FD85F94D" box="[390,516,1678,1701]" captionStart="Figure 10" captionStartId="16.[158,225,1715,1736]" captionTargetBox="[218,1405,170,1688]" captionTargetId="figure-180@16.[203,1418,163,1698]" captionTargetPageId="16" captionText="Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="29" pageNumber="30">
Figure 10
<collectionCode id="D6E4AE65FFEF0E41FE47F933FD85F94D" box="[498,516,1678,1701]" pageId="29" pageNumber="30">B</collectionCode>
</figureCitation>
) [probably, these areas of sclerotisation are the result of injuring the walls of the posterior pouches by the aedeagal spicules during copulation]. Posterior wall of gynatrial sac near bases of pouches without convex areas (
<figureCitation id="28CE2A25FFEF0E41FD9FF8B3FD7DF8CD" box="[554,764,1806,1829]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="29" pageNumber="30">
Figure 12
<collectionCode id="D6E4AE65FFEF0E41FD2AF8B3FD30F8CD" box="[671,689,1806,1829]" pageId="29" pageNumber="30">C</collectionCode>
, 12
<collectionCode id="D6E4AE65FFEF0E41FD5DF8B3FD7DF8CD" box="[744,764,1806,1829]" pageId="29" pageNumber="30">D</collectionCode>
</figureCitation>
). Anterior part of gynatrial sac relatively shorter and narrower than in two previous species. Proximal part of spermathecal duct slightly longer than shortened median part or latter rather long and proximal part as long as or slightly shorter, less often strongly shorter than half of median part; distal part of duct approximately as long as half of median part or slightly longer than it. Proximal tubular part of spermathecal capsule 2–3 times as long as transverse diameter of its distal spherical part, curved, but not gyrose. First rami rather widened anteriorly, widely and evenly rounded (
<figureCitation id="28CE2A25FFEF0E41FCE1FE5CFC3CFE10" box="[852,957,481,504]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="29" pageNumber="30">
Figure 8
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), less often acute-angled (
<figureCitation id="28CE2A25FFEF0E41FB5CFE5CFADBFE10" box="[1257,1370,481,504]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="29" pageNumber="30">
Figure 8
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) of narrowly rounded (
<figureCitation id="28CE2A25FFEF0E41FBB4FDBCFBEEFDF0" box="[1025,1135,513,536]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="29" pageNumber="30">
Figure 8
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) at anterior ends.
</paragraph>
<paragraph id="B04A36A0FFEF0E41FCC6FD9CFAD8FD7F" blockId="29.[843,1466,194,1892]" pageId="29" pageNumber="30">
In
<specimenCount id="A6F3FD29FFEF0E41FC25FD9CFB85FDD0" box="[912,1028,545,568]" count="1" pageId="29" pageNumber="30" type="female">one female</specimenCount>
from Tuva, posterior pouches formlessly sclerotised apically, dorsal wall of left pouch with narrow longitudinal sclerotised line, dorsal wall of right pouch with isolated triangular sclerotised patch at base.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFEF0E41FCF9FD58FBD4FC8B" pageId="29" pageNumber="30" type="distribution">
<paragraph id="B04A36A0FFEF0E41FCF9FD58FBBCFD14" blockId="29.[843,1466,194,1892]" box="[844,1085,741,764]" pageId="29" pageNumber="30">
<heading id="EB0281CCFFEF0E41FCF9FD58FBBCFD14" bold="true" box="[844,1085,741,764]" fontSize="10" level="2" pageId="29" pageNumber="30" reason="6">
<emphasis id="8281EAB2FFEF0E41FCF9FD58FBBCFD14" bold="true" box="[844,1085,741,764]" pageId="29" pageNumber="30">Notes on distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFEF0E41FCF9FCB0FBD4FC8B" blockId="29.[843,1466,194,1892]" pageId="29" pageNumber="30">The species is recorded for the first time for Georgia, Azerbaijan, Turkey, Kyrgyzstan, Russian Far East [?], and China (Sichuan) [?].</paragraph>
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</treatment>
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Figures 1
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3
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<figureCitation id="28CE2A25FFFC0E52FB6AFEC4FB7EFE78" box="[1247,1279,377,400]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="14" pageNumber="15">
7
<collectionCode id="D6E4AE65FFFC0E52FB59FEC4FB7EFE78" box="[1260,1279,377,400]" pageId="14" pageNumber="15">A</collectionCode>
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,
<figureCitation id="28CE2A25FFFC0E52FABAFEC4FAB0FE78" box="[1295,1329,377,400]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="14" pageNumber="15">
8
<collectionCode id="D6E4AE65FFFC0E52FAA9FEC4FAB0FE78" box="[1308,1329,377,400]" pageId="14" pageNumber="15">H</collectionCode>
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,
<figureCitation id="28CE2A25FFFC0E52FAF4FEC4FAEFFE78" box="[1345,1390,377,400]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="14" pageNumber="15">
12
<collectionCode id="D6E4AE65FFFC0E52FAEEFEC4FAEFFE78" box="[1371,1390,377,400]" pageId="14" pageNumber="15">A</collectionCode>
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)
</subSubSection>
</emphasis>
</paragraph>
<subSubSection id="F8EF652BFFFC0E52FCFEFE6EFB9BFCA3" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="B04A36A0FFFC0E52FCFEFE6EFBA3FE02" blockId="14.[843,1466,194,1938]" box="[843,1058,467,490]" pageId="14" pageNumber="15">
<heading id="EB0281CCFFFC0E52FCFEFE6EFBA3FE02" bold="true" box="[843,1058,467,490]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="8281EAB2FFFC0E52FCFEFE6EFBA3FE02" bold="true" box="[843,1058,467,490]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFC0E52FCF9FE4AFB9BFCA3" blockId="14.[843,1466,194,1938]" pageId="14" pageNumber="15">
<materialsCitation id="009D3CFDFFFC0E52FCF9FE4AFAC4FDE4" box="[844,1349,502,524]" collectionCode="ZISP" country="Germany" location="Hercynian Forest" municipality="Hercynia" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFFC0E52FCF9FE4AFC2BFDE4" bold="true" box="[844,938,503,524]" pageId="14" pageNumber="15">
<collectingCountry id="C8E27630FFFC0E52FCF9FE4AFC2BFDE4" box="[844,938,503,524]" name="Germany" pageId="14" pageNumber="15">Germany</collectingCountry>
</emphasis>
: “
<collectingMunicipality id="502EACDAFFFC0E52FC75FE4AFB94FDE4" box="[960,1045,503,524]" pageId="14" pageNumber="15">Hercynia</collectingMunicipality>
” [
<location id="B52A607BFFFC0E52FB9CFE4AFB4EFDE4" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FB9CFE4AFB4EFDE4" box="[1065,1231,503,524]" country="Germany" municipality="Hercynia" name="Hercynian Forest" pageId="14" pageNumber="15">Hercynian Forest</location>
],
<specimenCount id="A6F3FD29FFFC0E52FB68FE4AFB7BFDE4" box="[1245,1274,502,524]" count="1" pageId="14" pageNumber="15" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FAB3FE4AFABBFDE4" box="[1286,1338,503,524]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="14" pageNumber="15">ZISP</collectionCode>
).
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FAFCFE4AFA3BFDCE" collectionCode="MNHN" country="France" location="Foret de Fontainebleau" municipality="Ilede-France" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Seine-et-Marne">
<emphasis id="8281EAB2FFFC0E52FAFCFE4AFA0EFDE4" bold="true" box="[1353,1423,503,524]" pageId="14" pageNumber="15">
<collectingCountry id="C8E27630FFFC0E52FAFCFE4AFA0EFDE4" box="[1353,1423,503,524]" name="France" pageId="14" pageNumber="15">France</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFFC0E52FA2FFE4AFC2DFDCE" italics="true" pageId="14" pageNumber="15">
<collectingMunicipality id="502EACDAFFFC0E52FA2FFE4AFC2DFDCE" pageId="14" pageNumber="15">Îlede-France</collectingMunicipality>
</emphasis>
:
<collectingRegion id="7231F842FFFC0E52FC0CFDACFBCAFDCE" box="[953,1099,529,550]" pageId="14" pageNumber="15">Seine-et-Marne</collectingRegion>
,
<location id="B52A607BFFFC0E52FBE0FDACFAAEFDCE" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FBE0FDACFAAEFDCE" box="[1109,1327,529,550]" country="France" municipality="Ilede-France" name="Foret de Fontainebleau" pageId="14" pageNumber="15" stateProvince="Seine-et-Marne">Forêt de Fontainebleau</location>
, 1J (
<collectionCode id="D6E4AE65FFFC0E52FAD6FDACFA31FDCE" box="[1379,1456,529,550]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FCF9FD91FC50FDB3" collectingDate="1936-06" collectionCode="MNHN" location="Moloy" municipality="Cote-d' Or" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Region Bourgogne-Franche-Comte">
<emphasis id="8281EAB2FFFC0E52FCF9FD91FB18FDA9" box="[844,1177,556,577]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="7231F842FFFC0E52FCF9FD91FB18FDA9" box="[844,1177,556,577]" pageId="14" pageNumber="15">Région Bourgogne-Franche-Comté</collectingRegion>
</emphasis>
:
<collectingMunicipality id="502EACDAFFFC0E52FB1EFD91FA8AFDA9" box="[1195,1291,555,577]" pageId="14" pageNumber="15">Côte-d’ Or</collectingMunicipality>
,
<location id="B52A607BFFFC0E52FAACFD91FAD9FDA9" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FAACFD91FAD9FDA9" box="[1305,1368,556,577]" municipality="Cote-d' Or" name="Moloy" pageId="14" pageNumber="15" stateProvince="Region Bourgogne-Franche-Comte">Moloy</location>
,
<date id="C44B1060FFFC0E52FAD3FD91FA37FDA9" box="[1382,1462,556,577]" pageId="14" pageNumber="15" value="1936-06">
<collectingDate id="D40FE988FFFC0E52FAD3FD91FA37FDA9" box="[1382,1462,556,577]" pageId="14" pageNumber="15" value="1936-06">VI.1936</collectingDate>
</date>
,
<specimenCount id="A6F3FD29FFFC0E52FCF9FDFBFCE8FDB4" box="[844,873,582,604]" count="1" pageId="14" pageNumber="15" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FCCFFDFBFC46FDB3" box="[890,967,582,603]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FC6EFDFBFC28FD79" collectionCode="MNHN" collectorName="A. Jung" location="Foret Domaniale des Morieres" municipality="Provence-Alpes-Cote d' Azur" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1">
<emphasis id="8281EAB2FFFC0E52FC6EFDFBFB6BFDB3" box="[987,1258,582,604]" italics="true" pageId="14" pageNumber="15">
<collectingMunicipality id="502EACDAFFFC0E52FC6EFDFBFB6BFDB3" box="[987,1258,582,604]" pageId="14" pageNumber="15">Provence-Alpes-Côte d’ Azur</collectingMunicipality>
</emphasis>
: “FOR. MORIERES [
<location id="B52A607BFFFC0E52FCE4FDDCFB01FD9E" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FCE4FDDCFB01FD9E" box="[849,1152,609,630]" municipality="Provence-Alpes-Cote d' Azur" name="Foret Domaniale des Morieres" pageId="14" pageNumber="15">Forêt Domaniale des Morières</location>
] VAR”, coll.
<collectorName id="1D005376FFFC0E52FABBFDDCFADEFD9E" box="[1294,1375,609,630]" pageId="14" pageNumber="15">A. Jung</collectorName>
, 1J,
<specimenCount id="A6F3FD29FFFC0E52FA29FDDCFA38FD9F" box="[1436,1465,609,631]" count="1" pageId="14" pageNumber="15" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FCE7FDC1FC1EFD79" box="[850,927,636,657]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FC04FDC1FA3BFD79" box="[945,1466,635,657]" collectingDate="1973-07-25" collectionCode="MNHN" location="chaine de l' Etoile" municipality="Mimet" pageId="14" pageNumber="15" specimenCount="1">
<collectingMunicipality id="502EACDAFFFC0E52FC04FDC1FC73FD79" box="[945,1010,636,657]" pageId="14" pageNumber="15">Mimet</collectingMunicipality>
,
<location id="B52A607BFFFC0E52FC4BFDC1FB28FD79" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FC4BFDC1FB28FD79" box="[1022,1193,635,657]" municipality="Mimet" name="chaine de l' Etoile" pageId="14" pageNumber="15">chaîne de l’ Étoile</location>
,
<date id="C44B1060FFFC0E52FB00FDC1FAA8FD79" box="[1205,1321,636,657]" pageId="14" pageNumber="15" value="1973-07-25">
<collectingDate id="D40FE988FFFC0E52FB00FDC1FAA8FD79" box="[1205,1321,636,657]" pageId="14" pageNumber="15" value="1973-07-25">25.VII.1973</collectingDate>
</date>
, 1J (
<collectionCode id="D6E4AE65FFFC0E52FAD6FDC1FA31FD79" box="[1379,1456,636,657]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FCF9FD2BFAECFD43" box="[844,1389,662,684]" collectingDate="1983-07-26" collectionCode="MNHN" location="prairie" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Roquefort-les-Pins">
<collectingRegion id="7231F842FFFC0E52FCF9FD2BFC7CFD43" box="[844,1021,662,683]" pageId="14" pageNumber="15">Roquefort-les-Pins</collectingRegion>
,
<location id="B52A607BFFFC0E52FBBBFD2BFBCDFD43" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FBBBFD2BFBCDFD43" box="[1038,1100,662,683]" name="prairie" pageId="14" pageNumber="15" stateProvince="Roquefort-les-Pins">prairie</location>
,
<date id="C44B1060FFFC0E52FBEBFD2BFB53FD43" box="[1118,1234,662,683]" pageId="14" pageNumber="15" value="1983-07-26">
<collectingDate id="D40FE988FFFC0E52FBEBFD2BFB53FD43" box="[1118,1234,662,683]" pageId="14" pageNumber="15" value="1983-07-26">26.VII.1983</collectingDate>
</date>
,
<specimenCount id="A6F3FD29FFFC0E52FB56FD2BFA80FD44" box="[1251,1281,662,684]" count="1" pageId="14" pageNumber="15" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FAA3FD2BFAE2FD43" box="[1302,1379,662,683]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
).
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FACFFD2BFC4EFD08" collectingDate="1981-05-28" collectionCode="MNHN" country="Spain" location="Los Mallos [de Riglos" municipality="Province of Huesca" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Aragon">
<emphasis id="8281EAB2FFFC0E52FACFFD2BFA32FD43" bold="true" box="[1402,1459,662,683]" pageId="14" pageNumber="15">
<collectingCountry id="C8E27630FFFC0E52FACFFD2BFA32FD43" box="[1402,1459,662,683]" name="Spain" pageId="14" pageNumber="15">Spain</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFFC0E52FCF9FD0CFC13FD2E" box="[844,914,689,710]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="7231F842FFFC0E52FCF9FD0CFC13FD2E" box="[844,914,689,710]" country="Spain" name="Aragon" pageId="14" pageNumber="15">Aragon</collectingRegion>
</emphasis>
:
<collectingMunicipality id="502EACDAFFFC0E52FC15FD0CFBDCFD2E" box="[928,1117,689,710]" pageId="14" pageNumber="15">Province of Huesca</collectingMunicipality>
,
<location id="B52A607BFFFC0E52FBDDFD0CFAC2FD2E" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FBDDFD0CFAC2FD2E" box="[1128,1347,689,710]" country="Spain" municipality="Province of Huesca" name="Los Mallos [de Riglos" pageId="14" pageNumber="15" stateProvince="Aragon">Los Mallos [de Riglos</location>
],
<date id="C44B1060FFFC0E52FAE1FD0CFA37FD2E" box="[1364,1462,689,710]" pageId="14" pageNumber="15" value="1981-05-28">
<collectingDate id="D40FE988FFFC0E52FAE1FD0CFA37FD2E" box="[1364,1462,689,710]" pageId="14" pageNumber="15" value="1981-05-28">28.V.1981</collectingDate>
</date>
,
<specimenCount id="A6F3FD29FFFC0E52FCF9FD76FCE8FD09" box="[844,873,715,737]" count="1" pageId="14" pageNumber="15" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FCC2FD76FC45FD08" box="[887,964,715,736]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
).
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FC60FD76FC10FD13" collectingDate="1934-05-28" collectionCode="ZISP" country="Italy" location="Lecco" municipality="Lecco" pageId="14" pageNumber="15" specimenCount="6" specimenCount-female="6" stateProvince="Lombardy">
<emphasis id="8281EAB2FFFC0E52FC60FD76FB82FD08" bold="true" box="[981,1027,715,736]" pageId="14" pageNumber="15">
<collectingCountry id="C8E27630FFFC0E52FC60FD76FB82FD08" box="[981,1027,715,736]" name="Italy" pageId="14" pageNumber="15">Italy</collectingCountry>
</emphasis>
:
<emphasis id="8281EAB2FFFC0E52FBA7FD76FBF0FD08" box="[1042,1137,715,736]" italics="true" pageId="14" pageNumber="15">
<collectingRegion id="7231F842FFFC0E52FBA7FD76FBF0FD08" box="[1042,1137,715,736]" country="Italy" name="Lombardia" pageId="14" pageNumber="15">Lombardy</collectingRegion>
</emphasis>
:
<collectingMunicipality id="502EACDAFFFC0E52FBCAFD76FB3BFD08" box="[1151,1210,715,736]" pageId="14" pageNumber="15">Lecco</collectingMunicipality>
,
<date id="C44B1060FFFC0E52FB70FD76FAA7FD08" box="[1221,1318,715,736]" pageId="14" pageNumber="15" value="1934-05-28">
<collectingDate id="D40FE988FFFC0E52FB70FD76FAA7FD08" box="[1221,1318,715,736]" pageId="14" pageNumber="15" value="1934-05-28">28.V.1934</collectingDate>
</date>
(Passauro),
<specimenCount id="A6F3FD29FFFC0E52FA29FD76FA38FD09" box="[1436,1465,715,737]" count="6" pageId="14" pageNumber="15" type="female">6♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFC0E52FCE7FD5BFC07FD13" box="[850,902,742,763]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="14" pageNumber="15">ZISP</collectionCode>
).
</materialsCitation>
<materialsCitation id="009D3CFDFFFC0E52FC13FD5BFB02FCFE" collectingDate="1897-06-21" collectionCode="ZISP" country="Ukraine" location="vicinity of Zhytomyr" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Zhytomyr Prov.">
<emphasis id="8281EAB2FFFC0E52FC13FD5BFC76FD13" bold="true" box="[934,1015,742,763]" pageId="14" pageNumber="15">
<collectingCountry id="C8E27630FFFC0E52FC13FD5BFC76FD13" box="[934,1015,742,763]" name="Ukraine" pageId="14" pageNumber="15">Ukraine</collectingCountry>
</emphasis>
:
<collectingRegion id="7231F842FFFC0E52FBA1FD5BFB35FD13" box="[1044,1204,742,763]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FBA1FD5BFB2EFD13" box="[1044,1199,742,763]" italics="true" pageId="14" pageNumber="15">Zhytomyr Prov</emphasis>
.
</collectingRegion>
:
<location id="B52A607BFFFC0E52FB64FD5BFA37FD13" LSID="urn:lsid:plazi:treatment:385C87B6FFFC0E53FC3CFEE4FAD2FF10:B52A607BFFFC0E52FB64FD5BFA37FD13" box="[1233,1462,742,763]" country="Ukraine" name="vicinity of Zhytomyr" pageId="14" pageNumber="15" stateProvince="Zhytomyr Prov.">
vicinity of
<collectingRegion id="7231F842FFFC0E52FAE2FD5BFA37FD13" box="[1367,1462,742,763]" country="Ukraine" name="Zhytomyr" pageId="14" pageNumber="15">Zhytomyr</collectingRegion>
</location>
,
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(Birula),
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(
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);
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<emphasis id="8281EAB2FFFC0E52FB3EFCBCFA94FCFE" box="[1163,1301,769,790]" italics="true" pageId="14" pageNumber="15">Vinnytsia Prov</emphasis>
.
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, “Podol’ sk.[aya] gub.[erniya], Mogil.[evskiy uezd]”,
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.,
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<collectingDate id="D40FE988FFFC0E52FAEEFCA6FA3BFCD8" box="[1371,1466,795,816]" pageId="14" pageNumber="15" value="1901-06-08">8.VI.1901</collectingDate>
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(Chekini), 1J (
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).
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</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFC0E52FCF9FC30FBBBFAE3" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="B04A36A0FFFC0E52FCF9FC30FC38FC4C" blockId="14.[843,1466,194,1938]" box="[844,953,909,932]" pageId="14" pageNumber="15">
<heading id="EB0281CCFFFC0E52FCF9FC30FC38FC4C" bold="true" box="[844,953,909,932]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="8281EAB2FFFC0E52FCF9FC30FC38FC4C" bold="true" box="[844,953,909,932]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFC0E52FCF9FC08FBBBFAE3" blockId="14.[843,1466,194,1938]" pageId="14" pageNumber="15">
Larger. Membrane whitish, subtransparent (
<figureCitation id="28CE2A25FFFC0E52FA82FC08FA28FC24" box="[1335,1449,949,972]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="14" pageNumber="15">
Figure 1
<collectionCode id="D6E4AE65FFFC0E52FA27FC08FA28FC24" box="[1426,1449,949,972]" pageId="14" pageNumber="15">A</collectionCode>
</figureCitation>
). Apex of sensory process of paramere truncate, trapezoidally or rectangularly rounded (
<figureCitation id="28CE2A25FFFC0E52FB43FC48FA28FBE4" box="[1270,1449,1013,1036]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="14" pageNumber="15">
Figure 3
<collectionCode id="D6E4AE65FFFC0E52FAEEFC48FAF1FBE4" box="[1371,1392,1013,1036]" pageId="14" pageNumber="15">A</collectionCode>
, 3
<collectionCode id="D6E4AE65FFFC0E52FA23FC48FA28FBE4" box="[1430,1449,1013,1036]" pageId="14" pageNumber="15">B</collectionCode>
</figureCitation>
). Apex of conjunctival ventral lobe widened, rhomboid, with lateral denticles clearly visible in ventral view; posterior margin of lobe dorsal wall clearly visible in ventral view at extreme apex (
<figureCitation id="28CE2A25FFFC0E52FB1BFBC8FAD1FB64" box="[1198,1360,1140,1164]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="14" pageNumber="15">
Figure 6
<collectionCode id="D6E4AE65FFFC0E52FABBFBC8FAA3FB64" box="[1294,1314,1141,1164]" pageId="14" pageNumber="15">A</collectionCode>
–6
<collectionCode id="D6E4AE65FFFC0E52FA8BFBC8FAD1FB64" box="[1342,1360,1141,1164]" pageId="14" pageNumber="15">C</collectionCode>
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). Length of denticle on spicule basal plate at dorsolateral lobes greater relative to length of spicule: distance from apex of denticle to apex of spicule less than length of denticle (
<figureCitation id="28CE2A25FFFC0E52FC03FB49FBA8FAE3" box="[950,1065,1268,1291]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="14" pageNumber="15">
Figure 7
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).
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFC0E52FCF9FA8EFAACFA3A" pageId="14" pageNumber="15" type="description">
<paragraph id="B04A36A0FFFC0E52FCF9FA8EFC70FAA2" blockId="14.[843,1466,194,1938]" box="[844,1009,1331,1354]" pageId="14" pageNumber="15">
<heading id="EB0281CCFFFC0E52FCF9FA8EFC70FAA2" bold="true" box="[844,1009,1331,1354]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="8281EAB2FFFC0E52FCF9FA8EFC70FAA2" bold="true" box="[844,1009,1331,1354]" pageId="14" pageNumber="15">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFC0E52FCF9FAE6FAACFA3A" blockId="14.[843,1466,194,1938]" pageId="14" pageNumber="15">
Body
<quantity id="770D9B45FFFC0E52FC3AFAE6FB96FA9A" box="[911,1047,1371,1395]" metricMagnitude="-1" metricUnit="m" metricValue="1.96215" metricValueMax="2.1843999999999997" metricValueMin="1.7399" pageId="14" pageNumber="15" unit="in" value="7.725" valueMax="8.6" valueMin="6.85">6.85–8.60 in</quantity>
length,
<quantity id="770D9B45FFFC0E52FBC6FAE6FB7AFA9A" box="[1139,1275,1371,1395]" metricMagnitude="-1" metricUnit="m" metricValue="1.2065" metricValueMax="1.3716" metricValueMin="1.0413999999999999" pageId="14" pageNumber="15" unit="in" value="4.75" valueMax="5.4" valueMin="4.1">4.10–5.40 in</quantity>
width; pronotum
<quantity id="770D9B45FFFC0E52FCF9FAC6FC50FA7A" box="[844,977,1402,1426]" metricMagnitude="-1" metricUnit="m" metricValue="1.1049" metricValueMax="1.2573" metricValueMin="0.9525" pageId="14" pageNumber="15" unit="in" value="4.35" valueMax="4.95" valueMin="3.75">3.75–4.95 in</quantity>
width; head
<quantity id="770D9B45FFFC0E52FBE8FAC6FB62FA7A" box="[1117,1251,1402,1426]" metricMagnitude="-2" metricUnit="m" metricValue="4.5339" metricValueMax="4.8768" metricValueMin="4.191" pageId="14" pageNumber="15" unit="in" value="1.785" valueMax="1.92" valueMin="1.65">1.65–1.92 in</quantity>
width; ocular index 2.98–3.66; length of antennal segments I–V: 0.35–0.42: 0.53–0.78: 0.64–0.94: 0.91–1.18: 1.01–1.18.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFC0E53FCF9F9A9FAD2FF10" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="distribution">
<paragraph id="B04A36A0FFFC0E52FCF9F9A9FC54F9C3" blockId="14.[843,1466,194,1938]" box="[844,981,1556,1579]" pageId="14" pageNumber="15">
<heading id="EB0281CCFFFC0E52FCF9F9A9FC54F9C3" bold="true" box="[844,981,1556,1579]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="8281EAB2FFFC0E52FCF9F9A9FC54F9C3" bold="true" box="[844,981,1556,1579]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFC0E53FCF9F981FAD2FF10" blockId="14.[843,1466,194,1938]" lastBlockId="15.[844,1466,193,1854]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Southern (except Portugal), Western (except Belgium and the Netherlands), Eastern (except Belarus and Russia), and Northern (probably Lithuania) Europe. The subspecies was recorded also from Asian Turkey, Lebanon, Iran, Israel, Morocco and also doubtingly from Cyprus, Syria, Algeria and Egypt (
<bibRefCitation id="D4644B51FFFC0E52FB97F961FB02F91B" author="Lis JA" box="[1058,1155,1756,1779]" editor="Aukema B &amp; Rieger C" journalOrPublisher="Amsterdam: Netherlands Entomological Society" pageId="14" pageNumber="15" pagination="119 - 147" refId="ref40328" refString="Lis JA. 2006. Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs). In: Aukema B and Rieger C, editors. Catalogue of the Heteroptera of the Palaearctic Region Vol. 5, Amsterdam: Netherlands Entomological Society; p. 119 - 147." title="Family Cydnidae Billberg, 1820 - burrowing bugs (burrower bugs)" type="book chapter" volumeTitle="Catalogue of the Heteroptera of the Palaearctic Region" year="2006">Lis 2006</bibRefCitation>
). Apparently, the subspecies is absent in North Africa, and its distribution in Asia Minor and the Near East is very questionable and needs to be clarified. For example, modern records of
<taxonomicName id="77F54D23FFFC0E52FAE7F886FA3BF8BA" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[1362,1466,1851,1874]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFFC0E52FAE7F886FA3BF8BA" box="[1362,1466,1851,1874]" italics="true" pageId="14" pageNumber="15">C. dubius</emphasis>
</taxonomicName>
for Turkey are based on the identification of specimens without studying the genitalia; this follows from the fact that special notes about studying the genitalia are given only for
<taxonomicName id="77F54D23FFFD0E53FEB2F89BFDEDF8D5" authority="(Fallen, 1807)" baseAuthorityName="Fallen" baseAuthorityYear="1807" box="[263,620,1830,1853]" class="Insecta" family="Bombyliidae" genus="Legnotus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="8281EAB2FFFD0E53FEB2F89BFE40F8D5" box="[263,449,1830,1853]" italics="true" pageId="15" pageNumber="16">Legnotus picipes</emphasis>
(Fallén, 1807)
</taxonomicName>
(
<bibRefCitation id="D4644B51FFFD0E53FD35F89BFC00FF30" author="Yazici G &amp; Avar Y &amp; Yildirim E." box="[640,897,193,1853]" pageId="15" pageNumber="16" pagination="604 - 609" refId="ref41043" refString="Yazici G, Avar Y, Yildirim E. 2015. Contribution to the knowledge of the Cydnidae (Hemiptera, Heteroptera, Pentatomoidea) fauna of Turkey. Turkish Journal of Zoology. 39: 604 - 609." type="journal article" year="2015">Yazici et al. 2015</bibRefCitation>
) and the species of
<taxonomicName id="77F54D23FFFD0E53FBD5FF7CFA0BFF30" authority="Laporte, 1833" authorityName="Laporte" authorityYear="1833" box="[1120,1418,193,216]" class="Insecta" family="Carabidae" genus="Odontoscelis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="8281EAB2FFFD0E53FBD5FF7CFB6BFF30" box="[1120,1258,193,216]" italics="true" pageId="15" pageNumber="16">Odontoscelis</emphasis>
Laporte, 1833
</taxonomicName>
and
<taxonomicName id="77F54D23FFFD0E53FCF9FF5CFBE1FF10" authority="Laporte, 1833" authorityName="Laporte" authorityYear="1833" box="[844,1120,225,248]" class="Insecta" family="Scutelleridae" genus="Eurygaster" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="8281EAB2FFFD0E53FCF9FF5CFC40FF10" box="[844,961,225,248]" italics="true" pageId="15" pageNumber="16">Eurygaster</emphasis>
Laporte, 1833
</taxonomicName>
(
<bibRefCitation id="D4644B51FFFD0E53FBC5FF5CFAC2FF10" author="Fent M &amp; Aktac N." box="[1136,1347,225,248]" journalOrPublisher="Turkiye Entomoloji Dernegi." pageId="15" pageNumber="16" pagination="193 - 204" part="33" refId="ref39677" refString="Fent M, Aktac N. 2009. Trakya bolgesi acanthosomatidae, thyreocoridae, cydnidae, plataspidae, scutelleridae (Pentatomoidea: Heteroptera) faunasina katkilari. Turkiye Entomoloji Dernegi. 33: 193 - 204. Turkish." title="Trakya bolgesi acanthosomatidae, thyreocoridae, cydnidae, plataspidae, scutelleridae (Pentatomoidea: Heteroptera) faunasina katkilari" type="journal article" year="2009">Fent &amp; Aktaç 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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767
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<taxonomicName id="77F54D23FFFD0E53FC16FE84FA91FEB8" authority="Gapon, 2018" authorityName="Gapon" authorityYear="2018" box="[931,1296,313,336]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="dubius" status="subsp. nov." subSpecies="sanigarum">
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(
<figureCitation id="28CE2A25FFFD0E53FCD4FEE4FC5CFE98" box="[865,989,345,368]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="15" pageNumber="16">
Figures 1
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<figureCitation id="28CE2A25FFFD0E53FC58FEE4FBBCFE98" box="[1005,1085,345,368]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="15" pageNumber="16">
3
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<figureCitation id="28CE2A25FFFD0E53FBF9FEE4FB1DFE98" box="[1100,1180,345,368]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="15" pageNumber="16">
4
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, 4
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<figureCitation id="28CE2A25FFFD0E53FB1EFEE4FB71FE98" box="[1195,1264,344,368]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="15" pageNumber="16">
6
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<figureCitation id="28CE2A25FFFD0E53FAB5FEE4FA9EFE98" box="[1280,1311,345,368]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="15" pageNumber="16">
7
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<figureCitation id="28CE2A25FFFD0E53FA9BFEE4FAC4FE98" box="[1326,1349,345,368]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="15" pageNumber="16">
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8
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<figureCitation id="28CE2A25FFFD0E53FAE0FEE4FA25FE98" box="[1365,1444,345,368]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="15" pageNumber="16">
9
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43°40
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40°12
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[?] (Glusow [?], coll. Kiritshenko),
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],
<date id="C44B1060FFFD0E53FBE8FD0EFB56FD22" box="[1117,1239,691,714]" pageId="15" pageNumber="16" value="1925-08-08">8.VIII.1925</date>
(
<collectorName id="1D005376FFFD0E53FB5CFD0EFA05FD22" box="[1257,1412,691,714]" pageId="15" pageNumber="16">M. Korsakova</collectorName>
), 1J (
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);
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<materialsCitation id="009D3CFDFFFD0E53FC16FD6EFA38FD02" box="[931,1465,722,746]" collectingDate="1913-07-11" collectionCode="ZISP" location="Ashe" municipality="Ashe" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="2">
<collectingMunicipality id="502EACDAFFFD0E53FC16FD6EFC5AFD02" box="[931,987,723,746]" pageId="15" pageNumber="16">Ashe</collectingMunicipality>
,
<date id="C44B1060FFFD0E53FC5CFD6EFBACFD02" box="[1001,1069,723,746]" pageId="15" pageNumber="16" value="1913-06">23.VI.</date>
,
<date id="C44B1060FFFD0E53FB8EFD6EFB38FD02" box="[1083,1209,723,746]" pageId="15" pageNumber="16" value="1913-07-11">11.VII.1913</date>
(Nasonov),
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(
<collectionCode id="D6E4AE65FFFD0E53FAC1FD6EFA2FFD02" box="[1396,1454,723,746]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="15" pageNumber="16">ZISP</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFD0E53FCF9FD4EFB4FFCC2" collectingDate="1916-05-02" collectionCode="ZISP, MNHN" location="Krasnaya Polyana" municipality="Krasnaya Polyana" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="2">
<collectingMunicipality id="502EACDAFFFD0E53FCF9FD4EFB8DFCE2" box="[844,1036,755,778]" pageId="15" pageNumber="16">Krasnaya Polyana</collectingMunicipality>
, “07.. ... .. V.” [
<date id="C44B1060FFFD0E53FB02FD4EFA98FCE2" box="[1207,1305,755,778]" pageId="15" pageNumber="16" value="1916-05-02">2.V.1916</date>
] (Kiritshenko), 4J,
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(
<collectionCode id="D6E4AE65FFFD0E53FC01FCAEFC6FFCC2" box="[948,1006,787,810]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="15" pageNumber="16">ZISP</collectionCode>
), 1J,
<specimenCount id="A6F3FD29FFFD0E53FB8CFCAEFBDBFCC2" box="[1081,1114,786,810]" count="1" pageId="15" pageNumber="16" type="female">1♀</specimenCount>
(
<collectionCode id="D6E4AE65FFFD0E53FBD8FCAEFB42FCC2" box="[1133,1219,787,810]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="009D3CFDFFFD0E53FB6FFCAEFBD6FCA2" collectingDate="1924-08-16" collectionCode="ZISP" location="Nebug" pageId="15" pageNumber="16" specimenCount="1">
<location id="B52A607BFFFD0E53FB6FFCAEFAA3FCC2" LSID="urn:lsid:plazi:treatment:385C87B6FFFD0E4AFC16FE84FDBDFCFE:B52A607BFFFD0E53FB6FFCAEFAA3FCC2" box="[1242,1314,787,810]" name="Nebug" pageId="15" pageNumber="16">Nebug</location>
,
<date id="C44B1060FFFD0E53FA87FCAEFA3BFCC2" box="[1330,1466,787,810]" pageId="15" pageNumber="16" value="1924-08-16">16.VIII.1924</date>
(Gittermann), 1J (
<collectionCode id="D6E4AE65FFFD0E53FBA4FC8EFBCAFCA2" box="[1041,1099,819,842]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="15" pageNumber="16">ZISP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFD0E53FCF9FC2DFAEDFB06" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="B04A36A0FFFD0E53FCF9FC2DFC38FC4F" blockId="15.[844,1466,193,1854]" box="[844,953,912,935]" pageId="15" pageNumber="16">
<heading id="EB0281CCFFFD0E53FCF9FC2DFC38FC4F" bold="true" box="[844,953,912,935]" fontSize="10" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="8281EAB2FFFD0E53FCF9FC2DFC38FC4F" bold="true" box="[844,953,912,935]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFD0E53FCF9FC05FAEDFB06" blockId="15.[844,1466,193,1854]" pageId="15" pageNumber="16">
Smaller. Membrane brownish, not transparent (
<figureCitation id="28CE2A25FFFD0E53FA8AFC05FA2BFC27" box="[1343,1450,952,975]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="15" pageNumber="16">
Figure 1
<collectionCode id="D6E4AE65FFFD0E53FA20FC05FA2BFC27" box="[1429,1450,952,975]" pageId="15" pageNumber="16">B</collectionCode>
</figureCitation>
). Sensory process of paramere apically not truncated, narrowly or widely smoothly rounded (
<figureCitation id="28CE2A25FFFD0E53FB48FC45FA28FBE7" box="[1277,1449,1016,1039]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="15" pageNumber="16">
Figure 3
<collectionCode id="D6E4AE65FFFD0E53FAEAFC45FAF3FBE7" box="[1375,1394,1016,1039]" pageId="15" pageNumber="16">C</collectionCode>
, 3
<collectionCode id="D6E4AE65FFFD0E53FA20FC45FA28FBE7" box="[1429,1449,1016,1039]" pageId="15" pageNumber="16">D</collectionCode>
</figureCitation>
). Conjunctival ventral lobe at apex slightly widened, subrhomboid or widely rounded; denticles on its lateral walls hardly visible or absent, dorsal wall of lobe at its apex not visible in ventral view (
<figureCitation id="28CE2A25FFFD0E53FBF9FBCAFB5CFB66" box="[1100,1245,1143,1167]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="15" pageNumber="16">
Figure 6
<collectionCode id="D6E4AE65FFFD0E53FB13FBCAFB3BFB66" box="[1190,1210,1143,1166]" pageId="15" pageNumber="16">D</collectionCode>
–
<date id="C44B1060FFFD0E53FB72FBCAFB5CFB66" box="[1223,1245,1143,1166]" pageId="15" pageNumber="16">
6
<collectionCode id="D6E4AE65FFFD0E53FB61FBCAFB5CFB66" box="[1236,1245,1143,1166]" pageId="15" pageNumber="16">I</collectionCode>
</date>
</figureCitation>
). Length of denticle on spicule basal plate at dorsolateral lobes less relative to length of spicule: distance from apex of denticle to apex of spicule longer than length of denticle (
<figureCitation id="28CE2A25FFFD0E53FB5AFB6AFADDFB06" box="[1263,1372,1239,1262]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="15" pageNumber="16">
Figure 7
<collectionCode id="D6E4AE65FFFD0E53FAFFFB6AFADDFB06" box="[1354,1372,1239,1262]" pageId="15" pageNumber="16">B</collectionCode>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFD0E53FCF9FA88FA3BF971" pageId="15" pageNumber="16" type="description">
<paragraph id="B04A36A0FFFD0E53FCF9FA88FC44FAA4" blockId="15.[844,1466,193,1854]" box="[844,965,1333,1356]" pageId="15" pageNumber="16">
<heading id="EB0281CCFFFD0E53FCF9FA88FC44FAA4" bold="true" box="[844,965,1333,1356]" fontSize="10" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="8281EAB2FFFD0E53FCF9FA88FC44FAA4" bold="true" box="[844,965,1333,1356]" pageId="15" pageNumber="16">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFD0E53FCF9FAE0FBD5FA5B" blockId="15.[844,1466,193,1854]" pageId="15" pageNumber="16">
In specimen from Tuapse, ventral lobe wider, its apex convoluted as tube, without lateral denticles (
<figureCitation id="28CE2A25FFFD0E53FAAFFAC1FA2BFA7B" box="[1306,1450,1404,1428]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="15" pageNumber="16">
Figure 6
<collectionCode id="D6E4AE65FFFD0E53FAC0FAC1FA09FA7B" box="[1397,1416,1404,1427]" pageId="15" pageNumber="16">G</collectionCode>
–
<date id="C44B1060FFFD0E53FA20FAC1FA2BFA7B" box="[1429,1450,1404,1427]" pageId="15" pageNumber="16">
6
<collectionCode id="D6E4AE65FFFD0E53FA17FAC1FA2BFA7B" box="[1442,1450,1404,1427]" pageId="15" pageNumber="16">I</collectionCode>
</date>
</figureCitation>
); perhaps this is a teratism.
</paragraph>
<caption id="E48A6628FFFD0E53FF2BFA66FF7DF978" ID-DOI="http://doi.org/10.5281/zenodo.14927035" ID-Zenodo-Dep="14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="15" pageNumber="16" startId="15.[158,225,1499,1520]" targetBox="[180,761,175,1475]" targetPageId="15" targetType="figure">
<paragraph id="B04A36A0FFFD0E53FF2BFA66FF7DF978" blockId="15.[158,781,1499,1680]" pageId="15" pageNumber="16">
<emphasis id="8281EAB2FFFD0E53FF2BFA66FF7FFA18" bold="true" box="[158,254,1499,1520]" pageId="15" pageNumber="16">Figure 9.</emphasis>
Female terminalia in
<taxonomicName id="77F54D23FFFD0E53FE5BFA66FD44FA18" box="[494,709,1499,1520]" class="Insecta" family="Cydnidae" genus="Canthophorus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="8281EAB2FFFD0E53FE5BFA66FDF7FA18" box="[494,630,1499,1520]" italics="true" pageId="15" pageNumber="16">Canthophorus</emphasis>
spp.
<emphasis id="8281EAB2FFFD0E53FD06FA66FD64FA18" bold="true" box="[691,741,1499,1520]" pageId="15" pageNumber="16">A, B</emphasis>
</taxonomicName>
,
<taxonomicName id="77F54D23FFFD0E53FD43FA66FE07F9E3" authority="(Scopoli, 1763)" baseAuthorityName="Scopoli" baseAuthorityYear="1763" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFFD0E53FD43FA66FF5CF9E3" italics="true" pageId="15" pageNumber="16">C. dubius</emphasis>
(
<bibRefCitation id="D4644B51FFFD0E53FF44FA4BFEFCF9E3" author="Scopoli JA" box="[241,381,1526,1547]" journalOrPublisher="Vindobonae: Trattner" pageId="15" pageNumber="16" refId="ref40650" refString="Scopoli JA. 1763. Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana. Vindobonae: Trattner." title="Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana" type="book" year="1763">Scopoli, 1763</bibRefCitation>
)
</taxonomicName>
, Russia: Krasnaya Polyana (A), Italy: Lecco (B);
<emphasis id="8281EAB2FFFD0E53FE9EF9ACFEBDF9CE" bold="true" box="[299,316,1553,1574]" pageId="15" pageNumber="16">C</emphasis>
,
<taxonomicName id="77F54D23FFFD0E53FEECF9ACFD1FF9CE" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[345,670,1553,1574]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="melanopterus" subSpecies="melanopterus">
<emphasis id="8281EAB2FFFD0E53FEECF9ACFD1FF9CE" box="[345,670,1553,1574]" italics="true" pageId="15" pageNumber="16">C. melanopterus melanopterus</emphasis>
</taxonomicName>
(HerrichSchaeffer, 1835), Azerbaijan: Ganja;
<emphasis id="8281EAB2FFFD0E53FDB0F996FD97F9A8" bold="true" box="[517,534,1579,1600]" pageId="15" pageNumber="16">D</emphasis>
,
<taxonomicName id="77F54D23FFFD0E53FD96F996FC8DF9A8" authorityName="Gapon" authorityYear="2018" box="[547,780,1579,1600]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="melanopterus" subSpecies="mariae">
<emphasis id="8281EAB2FFFD0E53FD96F996FC8DF9A8" box="[547,780,1579,1600]" italics="true" pageId="15" pageNumber="16">C. melanopterus mariae</emphasis>
</taxonomicName>
<emphasis id="8281EAB2FFFD0E53FF2BF9FBFF61F9B3" bold="true" box="[158,224,1606,1627]" pageId="15" pageNumber="16">n. ssp</emphasis>
., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph id="B04A36A0FFFD0E53FCF9FA47FC70F9F9" blockId="15.[844,1466,193,1854]" box="[844,1009,1530,1553]" pageId="15" pageNumber="16">
<heading id="EB0281CCFFFD0E53FCF9FA47FC70F9F9" bold="true" box="[844,1009,1530,1553]" fontSize="10" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="8281EAB2FFFD0E53FCF9FA47FC70F9F9" bold="true" box="[844,1009,1530,1553]" pageId="15" pageNumber="16">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFD0E53FCF9F99FFA3BF971" blockId="15.[844,1466,193,1854]" pageId="15" pageNumber="16">
Body
<quantity id="770D9B45FFFD0E53FC27F99FFBA3F9D1" box="[914,1058,1569,1593]" metricMagnitude="-1" metricUnit="m" metricValue="1.8415" metricValueMax="2.1209" metricValueMin="1.5621" pageId="15" pageNumber="16" unit="in" value="7.25" valueMax="8.35" valueMin="6.15">6.15–8.35 in</quantity>
length,
<quantity id="770D9B45FFFD0E53FB31F99FFA92F9D1" box="[1156,1299,1569,1593]" metricMagnitude="-1" metricUnit="m" metricValue="1.13665" metricValueMax="1.3081" metricValueMin="0.9652" pageId="15" pageNumber="16" unit="in" value="4.475" valueMax="5.15" valueMin="3.8">3.80–5.15 in</quantity>
width; pronotum
<quantity id="770D9B45FFFD0E53FCCAF9FFFB8DF9B1" box="[895,1036,1601,1625]" metricMagnitude="-2" metricUnit="m" metricValue="9.9695" metricValueMax="11.43" metricValueMin="8.509" pageId="15" pageNumber="16" unit="in" value="3.925" valueMax="4.5" valueMin="3.35">3.35–4.50 in</quantity>
width; head
<quantity id="770D9B45FFFD0E53FB16F9FFFAB1F9B1" box="[1187,1328,1601,1625]" metricMagnitude="-2" metricUnit="m" metricValue="4.1148" metricValueMax="4.4704" metricValueMin="3.7592" pageId="15" pageNumber="16" unit="in" value="1.62" valueMax="1.76" valueMin="1.48">1.48–1.76 in</quantity>
width; ocular index 3.44–4.00; length of antennal segments (
<collectionCode id="D6E4AE65FFFD0E53FAC8F9DFFA07F991" box="[1405,1414,1634,1657]" pageId="15" pageNumber="16">I</collectionCode>
–
<collectionCode id="D6E4AE65FFFD0E53FA20F9DFFA2DF991" box="[1429,1452,1634,1657]" pageId="15" pageNumber="16">V</collectionCode>
): 0.34–0.41: 0.50–0.66: 0.56–0.76: 0.84–1.01: 0.94–1.09.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFD0E4EFCF9F962FDF2FBF9" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16" type="etymology">
<paragraph id="B04A36A0FFFD0E53FCF9F962FC47F91E" blockId="15.[844,1466,193,1854]" box="[844,966,1759,1782]" pageId="15" pageNumber="16">
<heading id="EB0281CCFFFD0E53FCF9F962FC47F91E" bold="true" box="[844,966,1759,1782]" fontSize="10" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="8281EAB2FFFD0E53FCF9F962FC47F91E" bold="true" box="[844,966,1759,1782]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFD0E4EFCF9F8BAFDF2FBF9" blockId="15.[844,1466,193,1854]" lastBlockId="18.[158,781,986,1936]" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
The subspecies epithet is given by the name of the tribe
<taxonomicName id="77F54D23FFFD0E53FCF9F89AFC15F8D6" box="[844,916,1831,1854]" pageId="15" pageNumber="16" rank="tribe" tribe="Sanigs">Sanigs</taxonomicName>
, Σάνηγας or Σανίγες in the ancient Greek spelling, which was inhabiting the
<typeStatus id="6F4E8802FFE20E4CFE05F88CFE5CF8A0" box="[432,477,1841,1864]" pageId="16" pageNumber="17">type</typeStatus>
locality of the subspecies in the Antiquity period: “...Σάνηγας δὲ καὶ Λαζοὺς ἐποίκους εἶχον τὰ χωρία...” [Memnon of Heraclea. History of Heracleia, 37]; “Ἐνθένδε εἰς Ἀχαιοῦντα στάδιοι ἑξήκοντα, ὅσπερ ποταμὸς διορίζει Ζιλχοὺς καὶ Σανίγας” (“From here [Masaitica Riv.], it is sixty stadia to Achaeunte [Ashe or Shakhe Riv.], where the river separates the Zygii and the
<taxonomicName id="77F54D23FFE00E4EFF2BFC47FF62FBF9" box="[158,227,1018,1041]" pageId="18" pageNumber="16" rank="tribe" tribe="Sanigs">Sanigs</taxonomicName>
”) [Arrian. Periplus Ponti Euxini, 15].
</paragraph>
</subSubSection>
<caption id="E48A6628FFE20E4CFF2BF90EFA3BF916" ID-DOI="http://doi.org/10.5281/zenodo.14927037" ID-Zenodo-Dep="14927037" httpUri="https://zenodo.org/record/14927037/files/figure.png" pageId="16" pageNumber="17" startId="16.[158,225,1715,1736]" targetBox="[218,1405,170,1688]" targetPageId="16" targetType="figure">
<paragraph id="B04A36A0FFE20E4CFF2BF90EFA3BF916" blockId="16.[158,1466,1715,1790]" pageId="16" pageNumber="17">
<emphasis id="8281EAB2FFE20E4CFF2BF90EFE87F920" bold="true" box="[158,262,1715,1736]" pageId="16" pageNumber="17">Figure 10.</emphasis>
Female terminalia in
<taxonomicName id="77F54D23FFE20E4CFE5FF90EFD3BF920" box="[490,698,1715,1736]" class="Insecta" family="Cydnidae" genus="Canthophorus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="8281EAB2FFE20E4CFE5FF90EFDF3F920" box="[490,626,1715,1736]" italics="true" pageId="16" pageNumber="17">Canthophorus</emphasis>
spp.
<emphasis id="8281EAB2FFE20E4CFD1DF90EFD57F920" bold="true" box="[680,726,1715,1736]" pageId="16" pageNumber="17">A, B</emphasis>
</taxonomicName>
,
<taxonomicName id="77F54D23FFE20E4CFD56F90EFBE7F920" authority="(Horvath, 1880)" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[739,1126,1715,1736]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="impressus">
<emphasis id="8281EAB2FFE20E4CFD56F90EFC42F920" box="[739,963,1715,1736]" italics="true" pageId="16" pageNumber="17">C. impressus impressus</emphasis>
(
<bibRefCitation id="D4644B51FFE20E4CFC64F90EFBDCF920" author="Horvath G." box="[977,1117,1715,1736]" journalOrPublisher="Termeszetrajzi Fuzetek." pageId="16" pageNumber="17" pagination="184 - 192" part="4" refId="ref40056" refString="Horvath G. 1880. Hemipterologiai kozlemenyek. Termeszetrajzi Fuzetek. 4: 184 - 192." title="Hemipterologiai kozlemenyek" type="journal article" year="1880">Horváth, 1880</bibRefCitation>
)
</taxonomicName>
, Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B);
<emphasis id="8281EAB2FFE20E4CFE79F973FE5CF90B" bold="true" box="[460,477,1742,1763]" pageId="16" pageNumber="17">C</emphasis>
,
<taxonomicName id="77F54D23FFE20E4CFE5FF973FD67F90B" authorityName="Gapon" authorityYear="2018" box="[490,742,1742,1763]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="hajastanicus">
<emphasis id="8281EAB2FFE20E4CFE5FF973FD67F90B" box="[490,742,1742,1763]" italics="true" pageId="16" pageNumber="17">C. impressus hajastanicus</emphasis>
</taxonomicName>
<emphasis id="8281EAB2FFE20E4CFD58F973FCA9F90B" bold="true" box="[749,808,1742,1763]" pageId="16" pageNumber="17">n. ssp</emphasis>
., Armenia: Gekhmakhan Mt., paratype;
<emphasis id="8281EAB2FFE20E4CFB04F973FB61F90B" bold="true" box="[1201,1248,1742,1763]" pageId="16" pageNumber="17">D, E</emphasis>
,
<taxonomicName id="77F54D23FFE20E4CFB58F973FF4FF916" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis id="8281EAB2FFE20E4CFB58F973FAD8F90B" box="[1261,1369,1742,1763]" italics="true" pageId="16" pageNumber="17">C. wagneri</emphasis>
<bibRefCitation id="D4644B51FFE20E4CFAD5F973FF4FF916" author="Asanova RB" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="16" pageNumber="17" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.
</paragraph>
</caption>
<caption id="E48A6628FFE30E4DFF2BF954FD28F8A5" ID-DOI="http://doi.org/10.5281/zenodo.14927039" ID-Zenodo-Dep="14927039" httpUri="https://zenodo.org/record/14927039/files/figure.png" pageId="17" pageNumber="18" startId="17.[158,225,1769,1790]" targetBox="[198,1415,169,1738]" targetPageId="17" targetType="figure">
<paragraph id="B04A36A0FFE30E4DFF2BF954FD28F8A5" blockId="17.[158,1466,1769,1869]" pageId="17" pageNumber="18">
<emphasis id="8281EAB2FFE30E4DFF2BF954FE85F916" bold="true" box="[158,260,1769,1790]" pageId="17" pageNumber="18">Figure 11.</emphasis>
Female terminalia in
<taxonomicName id="77F54D23FFE30E4DFE50F954FD32F916" box="[485,691,1769,1790]" class="Insecta" family="Cydnidae" genus="Canthophorus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="8281EAB2FFE30E4DFE50F954FDECF916" box="[485,621,1769,1790]" italics="true" pageId="17" pageNumber="18">Canthophorus</emphasis>
spp.
<emphasis id="8281EAB2FFE30E4DFD14F954FD4FF916" bold="true" box="[673,718,1769,1790]" pageId="17" pageNumber="18">A, B</emphasis>
</taxonomicName>
,
<taxonomicName id="77F54D23FFE30E4DFD6FF954FB8CF916" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[730,1037,1769,1790]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="mixtus" subSpecies="mixtus">
<emphasis id="8281EAB2FFE30E4DFD6FF954FCF9F916" box="[730,888,1769,1790]" italics="true" pageId="17" pageNumber="18">C. mixtus mixtus</emphasis>
<bibRefCitation id="D4644B51FFE30E4DFCCBF954FB8CF916" author="Asanova RB" box="[894,1037,1769,1790]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="17" pageNumber="18" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B);
<emphasis id="8281EAB2FFE30E4DFEB6F8BEFE95F8F0" bold="true" box="[259,276,1795,1816]" pageId="17" pageNumber="18">C</emphasis>
,
<taxonomicName id="77F54D23FFE30E4DFE91F8BEFD07F8F0" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Kiritshenko" baseAuthorityYear="1964" box="[292,646,1795,1816]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="mixtus" subSpecies="hissaricus">
<emphasis id="8281EAB2FFE30E4DFE91F8BEFE6BF8F0" box="[292,490,1795,1816]" italics="true" pageId="17" pageNumber="18">C. mixtus hissaricus</emphasis>
<bibRefCitation id="D4644B51FFE30E4DFE41F8BEFD07F8F0" author="Asanova RB" box="[500,646,1795,1816]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="17" pageNumber="18" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
;
<emphasis id="8281EAB2FFE30E4DFD23F8BEFD59F8F0" bold="true" box="[662,728,1795,1816]" pageId="17" pageNumber="18">n. stat</emphasis>
., Tajikistan: urochishche Ruydasht, paratype;
<emphasis id="8281EAB2FFE30E4DFB15F8BEFB30F8F0" bold="true" box="[1184,1201,1795,1816]" pageId="17" pageNumber="18">D</emphasis>
,
<taxonomicName id="77F54D23FFE30E4DFB74F8BEFF4FF8DB" authority="Scott, 1874" authorityName="Scott" authorityYear="1874" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="niveimarginatus">
<emphasis id="8281EAB2FFE30E4DFB74F8BEFAF8F8F0" box="[1217,1401,1795,1816]" italics="true" pageId="17" pageNumber="18">C. niveimarginatus</emphasis>
<bibRefCitation id="D4644B51FFE30E4DFA36F8BEFF4FF8DB" author="Scott J." journalOrPublisher="Annals and Magazine of Natural History." pageId="17" pageNumber="18" pagination="289 - 304" part="14" refId="ref40679" refString="Scott J. 1874. On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species. Annals and Magazine of Natural History. 14 (ser. 4): 289 - 304; 360 - 365; 426 - 452." title="On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species" type="journal article" year="1874">Scott, 1874</bibRefCitation>
</taxonomicName>
; Russia: Kamen’-Rybolov;
<emphasis id="8281EAB2FFE30E4DFE51F8A3FE75F8DB" bold="true" box="[484,500,1822,1843]" pageId="17" pageNumber="18">E</emphasis>
,
<taxonomicName id="77F54D23FFE30E4DFDB1F8A3FC98F8DB" authority="(Reuter, 1902)" baseAuthorityName="Reuter" baseAuthorityYear="1902" box="[516,793,1822,1843]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="8281EAB2FFE30E4DFDB1F8A3FD01F8DB" box="[516,640,1822,1843]" italics="true" pageId="17" pageNumber="18">C. coeruleus</emphasis>
(
<bibRefCitation id="D4644B51FFE30E4DFD24F8A3FC91F8DB" author="Reuter OM" box="[657,784,1822,1843]" journalOrPublisher="Ofversigt af Finska Vetenskaps-Societetens Forhandlingar." pageId="17" pageNumber="18" pagination="141 - 188" part="44" refId="ref40588" refString="Reuter OM. 1902. Miscellanea Hemipterologica. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar. 44: 141 - 188." title="Miscellanea Hemipterologica" type="journal article" year="1902">Reuter, 1902</bibRefCitation>
)
</taxonomicName>
, Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.
</paragraph>
</caption>
<caption id="E48A6628FFE00E4EFF2BFCABFE72FC7D" ID-DOI="http://doi.org/10.5281/zenodo.14927041" ID-Zenodo-Dep="14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="18" pageNumber="19" startId="18.[158,225,790,811]" targetBox="[179,1446,162,769]" targetPageId="18">
<paragraph id="B04A36A0FFE00E4EFF2BFCABFE72FC7D" blockId="18.[158,1466,790,917]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFF2BFCABFE87FCC3" bold="true" box="[158,262,790,811]" pageId="18" pageNumber="19">Figure 12.</emphasis>
Posterior pouches of gynatrial sac in
<taxonomicName id="77F54D23FFE00E4EFD3DFCABFCC5FCC3" box="[648,836,790,811]" class="Insecta" family="Cydnidae" genus="Canthophorus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8281EAB2FFE00E4EFD3DFCABFC92FCC3" box="[648,787,790,811]" italics="true" pageId="18" pageNumber="19">Canthophorus</emphasis>
spp.
</taxonomicName>
, posterio-inner view.
<emphasis id="8281EAB2FFE00E4EFBA2FCABFBA9FCC3" bold="true" box="[1047,1064,790,811]" pageId="18" pageNumber="19">A</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFB80FCABFAB2FCC3" authority="(Scopoli, 1763)" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[1077,1331,790,811]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFE00E4EFB80FCABFB15FCC3" box="[1077,1172,790,811]" italics="true" pageId="18" pageNumber="19">C. dubius</emphasis>
(
<bibRefCitation id="D4644B51FFE00E4EFB14FCABFAABFCC3" author="Scopoli JA" box="[1185,1322,790,811]" journalOrPublisher="Vindobonae: Trattner" pageId="18" pageNumber="19" refId="ref40650" refString="Scopoli JA. 1763. Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana. Vindobonae: Trattner." title="Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana" type="book" year="1763">Scopoli, 1763</bibRefCitation>
)
</taxonomicName>
, Italy: Lecco;
<emphasis id="8281EAB2FFE00E4EFF2BFC8DFF2FFCAD" bold="true" box="[158,174,816,837]" pageId="18" pageNumber="19">B</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFF0EFC8DFDDDFCAD" authority="(Herrich-Schaeffer, 1835)" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[187,604,816,837]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFE00E4EFF0EFC8DFEDAFCAD" box="[187,347,816,837]" italics="true" pageId="18" pageNumber="19">C. melanopterus</emphasis>
(
<bibRefCitation id="D4644B51FFE00E4EFED2FC8DFDD3FCAD" author="Herrich-Schaeffer GAW" box="[359,594,816,837]" journalOrPublisher="Regensburg: Pustet" pageId="18" pageNumber="19" refId="ref40016" refString="Herrich-Schaeffer GAW. 1835. Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie. Regensburg: Pustet." title="Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie" type="journal volume" year="1835">Herrich-Schaeffer, 1835</bibRefCitation>
)
</taxonomicName>
, Ukraine: Alma Riv.;
<emphasis id="8281EAB2FFE00E4EFC87FC8DFCC2FCAD" bold="true" box="[818,835,816,837]" pageId="18" pageNumber="19">C</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFCE5FC8DFB57FCAD" authority="(Horvath, 1880)" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[848,1238,816,837]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="impressus">
<emphasis id="8281EAB2FFE00E4EFCE5FC8DFBB2FCAD" box="[848,1075,816,837]" italics="true" pageId="18" pageNumber="19">C. impressus impressus</emphasis>
(
<bibRefCitation id="D4644B51FFE00E4EFB8AFC8DFB4CFCAD" author="Horvath G." box="[1087,1229,816,837]" journalOrPublisher="Termeszetrajzi Fuzetek." pageId="18" pageNumber="19" pagination="184 - 192" part="4" refId="ref40056" refString="Horvath G. 1880. Hemipterologiai kozlemenyek. Termeszetrajzi Fuzetek. 4: 184 - 192." title="Hemipterologiai kozlemenyek" type="journal article" year="1880">Horváth, 1880</bibRefCitation>
)
</taxonomicName>
, Ukraine: “yayla Besh-tekme”;
<emphasis id="8281EAB2FFE00E4EFF46FCF6FE85FC88" bold="true" box="[243,260,843,864]" pageId="18" pageNumber="19">D</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFEA6FCF6FD97FC88" authorityName="Gapon" authorityYear="2018" box="[275,534,843,864]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="hajastanicus">
<emphasis id="8281EAB2FFE00E4EFEA6FCF6FD97FC88" box="[275,534,843,864]" italics="true" pageId="18" pageNumber="19">C. impressus hajastanicus</emphasis>
</taxonomicName>
<emphasis id="8281EAB2FFE00E4EFDAAFCF6FDDAFC88" bold="true" box="[543,603,843,864]" pageId="18" pageNumber="19">n. ssp</emphasis>
., Armenia: Gekhmakhan Mt., paratype;
<emphasis id="8281EAB2FFE00E4EFC44FCF6FB80FC88" bold="true" box="[1009,1025,843,864]" pageId="18" pageNumber="19">E</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFBA5FCF6FA9DFC88" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[1040,1308,843,864]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis id="8281EAB2FFE00E4EFBA5FCF6FBFFFC88" box="[1040,1150,843,864]" italics="true" pageId="18" pageNumber="19">C. wagneri</emphasis>
<bibRefCitation id="D4644B51FFE00E4EFB32FCF6FA9DFC88" author="Asanova RB" box="[1159,1308,843,864]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="18" pageNumber="19" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Russia: Anapa (some diagnostic characters is indicated by arrows);
<emphasis id="8281EAB2FFE00E4EFD75FCD8FD4CFC92" bold="true" box="[704,717,869,890]" pageId="18" pageNumber="19">F</emphasis>
,
<taxonomicName id="77F54D23FFE00E4EFD6AFCD8FC62FC92" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[735,995,869,890]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="8281EAB2FFE00E4EFD6AFCD8FCC1FC92" box="[735,832,869,890]" italics="true" pageId="18" pageNumber="19">C. mixtus</emphasis>
<bibRefCitation id="D4644B51FFE00E4EFCFEFCD8FC62FC92" author="Asanova RB" box="[843,995,869,890]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="18" pageNumber="19" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.
</paragraph>
</caption>
<subSubSection id="F8EF652BFFE00E4EFF2BFBEAFD03F9EA" pageId="18" pageNumber="19" type="distribution">
<paragraph id="B04A36A0FFE00E4EFF2BFBEAFEB4FB5E" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<heading id="EB0281CCFFE00E4EFF2BFBEAFEA6FB86" bold="true" box="[158,295,1111,1134]" fontSize="10" level="2" pageId="18" pageNumber="19" reason="6">
<emphasis id="8281EAB2FFE00E4EFF2BFBEAFEA6FB86" bold="true" box="[158,295,1111,1134]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
Figure 15. Krasnodar Territory of Russia (Black Sea coast and foothills).
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF11FB43FD41FABD" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFF11FB43FD41FABD" bold="true" pageId="18" pageNumber="19">
<taxonomicName id="77F54D23FFE00E4EFF11FB43FC86FAFD" authority="(Herrich-Schaeffer, 1835)" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[164,775,1278,1301]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFE00E4EFF11FB43FE5FFAFD" bold="true" box="[164,478,1278,1301]" italics="true" pageId="18" pageNumber="19">Canthophorus melanopterus</emphasis>
(
<bibRefCitation id="D4644B51FFE00E4EFE5BFB43FD7FFAFD" author="Herrich-Schaeffer GAW" box="[494,766,1278,1301]" journalOrPublisher="Regensburg: Pustet" pageId="18" pageNumber="19" refId="ref40016" refString="Herrich-Schaeffer GAW. 1835. Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie. Regensburg: Pustet." title="Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie" type="journal volume" year="1835">Herrich-Schaeffer, 1835</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="28CE2A25FFE00E4EFF1BFAA3FEDDFADD" box="[174,348,1310,1333]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="18" pageNumber="19">
Figures 1
<collectionCode id="D6E4AE65FFE00E4EFEADFAA3FEADFADD" box="[280,300,1310,1333]" pageId="18" pageNumber="19">C</collectionCode>
, 1
<collectionCode id="D6E4AE65FFE00E4EFEF2FAA3FEDDFADD" box="[327,348,1310,1333]" pageId="18" pageNumber="19">D</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFE00E4EFEDEFAA3FE3AFADD" box="[363,443,1310,1333]" captionStart="Figure 2" captionStartId="6.[158,225,1725,1746]" captionTargetBox="[209,1426,182,1704]" captionTargetId="figure-102@6.[209,1426,182,1704]" captionTargetPageId="6" captionText="Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927021" httpUri="https://zenodo.org/record/14927021/files/figure.png" pageId="18" pageNumber="19">
2
<collectionCode id="D6E4AE65FFE00E4EFECDFAA3FE0AFADD" box="[376,395,1310,1333]" pageId="18" pageNumber="19">A</collectionCode>
, 2
<collectionCode id="D6E4AE65FFE00E4EFE1CFAA3FE3AFADD" box="[425,443,1310,1333]" pageId="18" pageNumber="19">B</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFE00E4EFE7FFAA3FE68FADD" box="[458,489,1310,1333]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="18" pageNumber="19">
3
<collectionCode id="D6E4AE65FFE00E4EFE62FAA3FE68FADD" box="[471,489,1310,1333]" pageId="18" pageNumber="19">E</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFE00E4EFE4DFAA3FDC9FADD" box="[504,584,1310,1334]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="18" pageNumber="19">
4
<collectionCode id="D6E4AE65FFE00E4EFDB3FAA3FD98FADD" box="[518,537,1310,1333]" pageId="18" pageNumber="19">C</collectionCode>
–4
<collectionCode id="D6E4AE65FFE00E4EFD86FAA3FDC9FADD" box="[563,584,1310,1333]" pageId="18" pageNumber="19">H</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFE00E4EFDE2FAA3FD27FADD" box="[599,678,1310,1334]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="18" pageNumber="19">
6
<collectionCode id="D6E4AE65FFE00E4EFDD1FAA3FDF3FADD" box="[612,626,1310,1333]" pageId="18" pageNumber="19">J</collectionCode>
–6
<collectionCode id="D6E4AE65FFE00E4EFD39FAA3FD27FADD" box="[652,678,1310,1333]" pageId="18" pageNumber="19">M</collectionCode>
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,
<figureCitation id="28CE2A25FFE00E4EFD00FAA3FD7FFADD" box="[693,766,1310,1334]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="18" pageNumber="19">
7
<collectionCode id="D6E4AE65FFE00E4EFD77FAA3FD54FADD" box="[706,725,1310,1333]" pageId="18" pageNumber="19">C</collectionCode>
–7
<collectionCode id="D6E4AE65FFE00E4EFD5AFAA3FD7FFADD" box="[751,766,1310,1333]" pageId="18" pageNumber="19">F</collectionCode>
</figureCitation>
,
<figureCitation id="28CE2A25FFE00E4EFF5FFA83FEBBFABD" box="[234,314,1342,1366]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="18" pageNumber="19">
8
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–8
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,
<figureCitation id="28CE2A25FFE00E4EFEFCFA83FE18FABD" box="[329,409,1342,1365]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="18" pageNumber="19">
9
<collectionCode id="D6E4AE65FFE00E4EFEE3FA83FEE8FABD" box="[342,361,1342,1365]" pageId="18" pageNumber="19">C</collectionCode>
, 9
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,
<figureCitation id="28CE2A25FFE00E4EFE1CFA83FE57FABD" box="[425,470,1342,1365]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="18" pageNumber="19">
12
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<figureCitation id="28CE2A25FFE00E4EFE50FA83FDC9FABD" box="[485,584,1342,1366]" captionStart="Figure 13" captionStartId="19.[158,225,1739,1760]" captionTargetBox="[281,1360,179,1715]" captionTargetId="figure-79@19.[281,1360,179,1715]" captionTargetPageId="19" captionText="Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927043" httpUri="https://zenodo.org/record/14927043/files/figure.png" pageId="18" pageNumber="19">
13
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–13
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,
<figureCitation id="28CE2A25FFE00E4EFDE2FA83FD36FABD" box="[599,695,1342,1366]" captionStart="Figure 14" captionStartId="20.[158,225,1738,1759]" captionTargetBox="[300,1351,177,1709]" captionTargetId="figure-80@20.[283,1370,164,1722]" captionTargetPageId="20" captionText="Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927045" httpUri="https://zenodo.org/record/14927045/files/figure.png" pageId="18" pageNumber="19">
14
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–
<date id="C44B1060FFE00E4EFD27FA83FD36FABD" box="[658,695,1342,1365]" pageId="18" pageNumber="19" value="2014-01">
14
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)
</emphasis>
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF2BFAD5FEB9FA7F" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<taxonomicName id="77F54D23FFE00E4EFF2BFAD5FD1CFA95" authority="Herrich Schaeffer 1835: 55" authorityName="Herrich Schaeffer" authorityPageNumber="55" authorityYear="1835" box="[158,669,1384,1405]" class="Insecta" family="Cydnidae" genus="Cydnus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFE00E4EFF2BFAD5FEF4FA95" box="[158,373,1384,1405]" italics="true" pageId="18" pageNumber="19">Cydnus melanopterus</emphasis>
Herrich Schaeffer 1835: 55
</taxonomicName>
[
<typeStatus id="6F4E8802FFE00E4EFD05FAD5FC88FA95" box="[688,777,1384,1405]" pageId="18" pageNumber="19">syntypes</typeStatus>
: Spain; lost].
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF2BFA20FDFCFA5A" blockId="18.[158,781,986,1936]" box="[158,637,1437,1458]" pageId="18" pageNumber="19">
<taxonomicName id="77F54D23FFE00E4EFF2BFA20FDAFFA5A" authority="Vidal 1950: 43" authorityName="Vidal" authorityPageNumber="43" authorityYear="1950" box="[158,558,1437,1458]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="variety" species="dubius" status="syn. nov." variety="nigra">
<emphasis id="8281EAB2FFE00E4EFF2BFA20FEADFA5A" box="[158,300,1437,1458]" italics="true" pageId="18" pageNumber="19">Sehirus dubius</emphasis>
var.
<emphasis id="8281EAB2FFE00E4EFEEAFA20FE12FA5A" box="[351,403,1437,1458]" italics="true" pageId="18" pageNumber="19">nigra</emphasis>
<bibRefCitation id="D4644B51FFE00E4EFE2EFA20FDAFFA5A" author="Vidal JP" box="[411,558,1437,1458]" journalOrPublisher="Memoires de la Societe des Sciences Naturelles du Maroc." pageId="18" pageNumber="19" pagination="1 - 238" part="48" refId="ref40914" refString="Vidal JP. 1950. Hemipteres de l' Afrique du Nord et des pays circum-mediterraneens. Memoires de la Societe des Sciences Naturelles du Maroc. 48: [1949]: 1 - 238." title="Hemipteres de l' Afrique du Nord et des pays circum-mediterraneens" type="journal article" year="1950">Vidal 1950: 43</bibRefCitation>
</taxonomicName>
,
<emphasis id="8281EAB2FFE00E4EFD8FFA20FDFCFA5A" bold="true" box="[570,637,1437,1458]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="99B257C9FFE00E4EFD8FFA20FDFCFA5A" box="[570,637,1437,1458]" pageId="18" pageNumber="19">n. syn.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF2BFA0AFDD5FA0F" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<taxonomicName id="77F54D23FFE00E4EFF2BFA0AFE4DFA24" box="[158,460,1463,1484]" class="Amphibia" family="Cryptobranchidae" genus="Tritomegas" kingdom="Animalia" order="Caudata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="melanopterus" subSpecies="niger">
<emphasis id="8281EAB2FFE00E4EFF2BFA0AFE4DFA24" box="[158,460,1463,1484]" italics="true" pageId="18" pageNumber="19">Tritomegas melanopterus niger</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFE00E4EFE6FFA0AFD08FA24" author="Stichel W." box="[474,649,1463,1484]" journalOrPublisher="Berlin- Hermsdorf: Stichel" pageId="18" pageNumber="19" pagination="545 - 768" refId="ref40805" refString="Stichel W. 1961. Illustrierte Bestimmungstabellen der Wanzen. II. Europa. (Hemiptera - Heteroptera Europae). Vol. 4. Berlin- Hermsdorf: Stichel; p. 545 - 768." title="Illustrierte Bestimmungstabellen der Wanzen. II. Europa. (Hemiptera - Heteroptera Europae). Vol. 4" type="book chapter" year="1961">Stichel 1961: 675</bibRefCitation>
.
<taxonomicName id="77F54D23FFE00E4EFF2BFA6FFE6BFA0F" baseAuthorityName="Vidal" baseAuthorityYear="1950" box="[158,490,1490,1511]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="melanopterus" subSpecies="niger">
<emphasis id="8281EAB2FFE00E4EFF2BFA6FFE6BFA0F" box="[158,490,1490,1511]" italics="true" pageId="18" pageNumber="19">Canthophorus melanopterus niger</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFE00E4EFE4DFA6FFDCFFA0F" author="Lis JA" box="[504,590,1490,1511]" journalOrPublisher="Genus." pageId="18" pageNumber="19" pagination="165 - 249" part="10" refId="ref40295" refString="Lis JA. 1999. Burrower bugs of the Old World - a catalogue (Hemiptera: Heteroptera: Cydnidae). Genus. 10 (2): 165 - 249." title="Burrower bugs of the Old World - a catalogue (Hemiptera: Heteroptera: Cydnidae)" type="journal article" year="1999">Lis 1999</bibRefCitation>
.
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF2BFA50FD03F9EA" blockId="18.[158,781,986,1936]" box="[158,642,1517,1538]" pageId="18" pageNumber="19">
<taxonomicName id="77F54D23FFE00E4EFF2BFA50FE31F9EA" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[158,432,1517,1538]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFE00E4EFF2BFA50FE31F9EA" box="[158,432,1517,1538]" italics="true" pageId="18" pageNumber="19">Canthophorus melanopterus</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFE00E4EFE0BFA50FDFFF9EA" author="Asanova RB" box="[446,638,1517,1538]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="18" pageNumber="19" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova 1964: 143</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFE00E4AFF2BF9E0FDBDFCFE" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19" type="description">
<paragraph id="B04A36A0FFE00E4EFF2BF9E0FD49F914" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<heading id="EB0281CCFFE00E4EFF2BF9E0FEA1F99C" bold="true" box="[158,288,1629,1652]" fontSize="10" level="2" pageId="18" pageNumber="19" reason="6">
<emphasis id="8281EAB2FFE00E4EFF2BF9E0FEA1F99C" bold="true" box="[158,288,1629,1652]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
Lateral margin of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metapleural evaporatorium with long and curved peritreme. Membrane brownish or whitish.
</paragraph>
<paragraph id="B04A36A0FFE00E4EFF15F8A7FE5EF878" blockId="18.[158,781,986,1936]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFF15F8A7FE94F8D9" bold="true" box="[160,277,1818,1841]" italics="true" pageId="18" pageNumber="19">Pygophore</emphasis>
slightly longer than wide; its lateral walls deeply concave in middle of length (
<figureCitation id="28CE2A25FFE00E4EFD8AF887FD31F8B9" box="[575,688,1850,1873]" captionStart="Figure 2" captionStartId="6.[158,225,1725,1746]" captionTargetBox="[209,1426,182,1704]" captionTargetId="figure-102@6.[209,1426,182,1704]" captionTargetPageId="6" captionText="Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927021" httpUri="https://zenodo.org/record/14927021/files/figure.png" pageId="18" pageNumber="19">
Figure 2
<collectionCode id="D6E4AE65FFE00E4EFD29F887FD31F8B9" box="[668,688,1850,1873]" pageId="18" pageNumber="19">A</collectionCode>
</figureCitation>
); dorsal infolding long, strongly concave in middle of width; dorsal wall short (
<figureCitation id="28CE2A25FFE00E4EFED6F8C4FE4EF878" box="[355,463,1913,1936]" captionStart="Figure 2" captionStartId="6.[158,225,1725,1746]" captionTargetBox="[209,1426,182,1704]" captionTargetId="figure-102@6.[209,1426,182,1704]" captionTargetPageId="6" captionText="Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927021" httpUri="https://zenodo.org/record/14927021/files/figure.png" pageId="18" pageNumber="19">
Figure 2
<collectionCode id="D6E4AE65FFE00E4EFE0BF8C4FE4EF878" box="[446,463,1913,1936]" pageId="18" pageNumber="19">B</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="B04A36A0FFE00E4EFCF9FC67FB37FBD9" blockId="18.[844,1466,986,1938]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFCF9FC67FC3CFC19" bold="true" box="[844,957,986,1009]" italics="true" pageId="18" pageNumber="19">Paramere.</emphasis>
Hypophysis curved dorsad rather less strongly than in previous species, so that apex of hypophysis not visible in outer view (
<figureCitation id="28CE2A25FFE00E4EFB8EFBA7FB26FBD9" box="[1083,1191,1050,1073]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="18" pageNumber="19">
Figure 3
<collectionCode id="D6E4AE65FFE00E4EFB20FBA7FB26FBD9" box="[1173,1191,1050,1073]" pageId="18" pageNumber="19">E</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="B04A36A0FFE00E4EFCF9FBF1FA31F8A8" blockId="18.[844,1466,986,1938]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFCF9FBF1FBC7FB8B" bold="true" box="[844,1094,1100,1123]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFCF9FBF1FC37FB8B" bold="true" box="[844,950,1100,1123]" italics="true" pageId="18" pageNumber="19">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFE00E4EFC70FBF1FBB6FB8B" box="[965,1079,1100,1123]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="18" pageNumber="19">
Figure 4
<collectionCode id="D6E4AE65FFE00E4EFB91FBF1FBB6FB8B" box="[1060,1079,1100,1123]" pageId="18" pageNumber="19">C</collectionCode>
</figureCitation>
).
</emphasis>
Ventral lobe of conjunctiva slightly protruding beyond apical margin of theca, slightly tapering or slightly widening at apex in ventral view; apex of lobe rounded, without lateral denticles; ventral wall of lobe rooflike, strongly elevated in apical part, with longitudinal narrow carina (
<figureCitation id="28CE2A25FFE00E4EFC44FB56FB04FAEA" box="[1009,1157,1259,1282]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="18" pageNumber="19">
Figure 6
<collectionCode id="D6E4AE65FFE00E4EFBF9FB56FBD6FAEA" box="[1100,1111,1259,1282]" pageId="18" pageNumber="19">J</collectionCode>
, 6
<collectionCode id="D6E4AE65FFE00E4EFBC1FB56FB04FAEA" box="[1140,1157,1259,1282]" pageId="18" pageNumber="19">L</collectionCode>
</figureCitation>
); in lateral view, apical part of lobe wide, carina on its ventral wall protruding beyond apical margin of lobe as denticle (
<figureCitation id="28CE2A25FFE00E4EFB0AFA96FAE0FAAA" box="[1215,1377,1323,1346]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="18" pageNumber="19">
Figure 6
<collectionCode id="D6E4AE65FFE00E4EFAADFA96FAADFAAA" box="[1304,1324,1323,1346]" pageId="18" pageNumber="19">K</collectionCode>
, 6
<collectionCode id="D6E4AE65FFE00E4EFAFCFA96FAE0FAAA" box="[1353,1377,1323,1346]" pageId="18" pageNumber="19">M</collectionCode>
</figureCitation>
); sometimes apical margin of lobe with two small notches on either side of carina (
<figureCitation id="28CE2A25FFE00E4EFBF3FAD6FB6FFA6A" box="[1094,1262,1387,1410]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="18" pageNumber="19">
Figure 6
<collectionCode id="D6E4AE65FFE00E4EFB11FAD6FB34FA6A" box="[1188,1205,1387,1410]" pageId="18" pageNumber="19">L</collectionCode>
, 6
<collectionCode id="D6E4AE65FFE00E4EFB63FAD6FB6FFA6A" box="[1238,1262,1387,1410]" pageId="18" pageNumber="19">M</collectionCode>
</figureCitation>
). Apical lobes of conjunctiva long, protruding beyond dorsoventral plane of apical half of conjunctiva, basally directed dorsolaterad, in middle curved posteroventrad. Vesica with more or less long and thin membranous lateral branch originating before middle of vesica at acute angle to its longitudinal axis and lying most often on its right side; pointed apex of lateral brunch directed to apex of vesica (
<figureCitation id="28CE2A25FFE00E4EFAB3F9F7FA2BF989" box="[1286,1450,1610,1634]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="18" pageNumber="19">
Figure 4
<collectionCode id="D6E4AE65FFE00E4EFADFF9F7FAFCF989" box="[1386,1405,1610,1633]" pageId="18" pageNumber="19">D</collectionCode>
–4
<collectionCode id="D6E4AE65FFE00E4EFA22F9F7FA2BF989" box="[1431,1450,1610,1633]" pageId="18" pageNumber="19">H</collectionCode>
</figureCitation>
). Dorsolateral lobes of conjunctiva long, with bases not separated from each other by impression; they directed dorsoanteriad and curved to each other by apices. Inner wall of each lobe before apex with sclerotised plate passing into spicule basal plate. Apices of spicules directed to each other and dorsad. Spicule basal plate on inner end with two triangular denticles (
<figureCitation id="28CE2A25FFE00E4EFB91F894FB36F8A8" box="[1060,1207,1833,1857]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="18" pageNumber="19">
Figure 7
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) directed dorsomediad.
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<paragraph id="B04A36A0FFE00E4AFCF9F8E6FDBDFCFE" blockId="18.[844,1466,986,1938]" lastBlockId="22.[158,781,193,1892]" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFCF9F8E6FB10F89A" bold="true" box="[844,1169,1883,1906]" pageId="18" pageNumber="19">
<emphasis id="8281EAB2FFE00E4EFCF9F8E6FC42F89A" bold="true" box="[844,963,1883,1906]" italics="true" pageId="18" pageNumber="19">Gynatrium</emphasis>
(
<figureCitation id="28CE2A25FFE00E4EFC6DF8E6FB03F89A" box="[984,1154,1883,1906]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="18" pageNumber="19">
Figure 8
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).
</emphasis>
Posterior pouches located behind posterior margin of gynatrial sac, approximately as long as wide or slightly wider than long in dorsal view, with rounded posterior margin. Walls of pouches thicker and more rigid than in
<taxonomicName id="77F54D23FFE40E4AFE24FEBCFE79FEF0" box="[401,504,257,280]" class="Insecta" family="Cydnidae" genus="Sehirus" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFE40E4AFE24FEBCFE79FEF0" box="[401,504,257,280]" italics="true" pageId="22" pageNumber="23">C. dubius</emphasis>
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; inner wall of each pouch sloped and slightly concave. Each pouch containing shaped sclerite inside (
<figureCitation id="28CE2A25FFE40E4AFE2CFEFCFD90FEB0" box="[409,529,321,344]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="22" pageNumber="23">
Figures 8
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,
<figureCitation id="28CE2A25FFE40E4AFD96FEFCFDCEFEB0" box="[547,591,321,344]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="22" pageNumber="23">
12
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,
<figureCitation id="28CE2A25FFE40E4AFDD5FEFCFD41FEB0" box="[608,704,321,345]" captionStart="Figure 13" captionStartId="19.[158,225,1739,1760]" captionTargetBox="[281,1360,179,1715]" captionTargetId="figure-79@19.[281,1360,179,1715]" captionTargetPageId="19" captionText="Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927043" httpUri="https://zenodo.org/record/14927043/files/figure.png" pageId="22" pageNumber="23">
13
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,
<figureCitation id="28CE2A25FFE40E4AFD64FEFCFEBCFE90" captionStart="Figure 14" captionStartId="20.[158,225,1738,1759]" captionTargetBox="[300,1351,177,1709]" captionTargetId="figure-80@20.[283,1370,164,1722]" captionTargetPageId="20" captionText="Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927045" httpUri="https://zenodo.org/record/14927045/files/figure.png" pageId="22" pageNumber="23">
14
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– 14
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), rarely poorly visible as very thin plate along anterior inner margin of pouch (
<figureCitation id="28CE2A25FFE40E4AFDCDFE3CFD7DFE70" box="[632,764,385,408]" captionStart="Figure 14" captionStartId="20.[158,225,1738,1759]" captionTargetBox="[300,1351,177,1709]" captionTargetId="figure-80@20.[283,1370,164,1722]" captionTargetPageId="20" captionText="Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927045" httpUri="https://zenodo.org/record/14927045/files/figure.png" pageId="22" pageNumber="23">
Figure 14
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). Anterior part of gynatrial sac long and wide. Spermathecal duct with very short basal part, relatively long, porrect median part, slightly more than twice as long as distal part; sometimes proximal part elongated, attaining one third of length of shortened medial part having almost spherical shape. Tubular proximal part of spermathecal capsule gyrose and clearly longer than transverse diameter of spherical distal part (
<figureCitation id="28CE2A25FFE40E4AFDAFFD3DFD41FD7F" box="[538,704,640,663]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="22" pageNumber="23">
Figure 9
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, 9
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). First rami widely varying in shape and width without any geographical regularity: sharply or smoothly widening anteriorly, with rectangularly or smoothly rounded or skewed anterior ends (
<figureCitation id="28CE2A25FFE40E4AFE27FD42FDACFCFE" box="[402,557,767,791]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="22" pageNumber="23">
Figure 8
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).
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<mods:identifier id="7A2B199F2D8661D38B69CF9D998D3629" type="DOI">10.1080/00379271.2018.1467788</mods:identifier>
<mods:identifier id="B1B3BFF65F1E0FF5F1BE197BA0348C6E" type="Zenodo-Dep">14927017</mods:identifier>
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<paragraph id="B04A36A0FFFE0E50FC07F944FAD2F8F8" blockId="12.[844,1466,194,1938]" box="[946,1363,1785,1808]" pageId="12" pageNumber="13">
<taxonomicName id="77F54D23FFFE0E50FC07F944FAD2F8F8" authority="(Scopoli, 1763)" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[946,1363,1785,1808]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFFE0E50FC07F944FAD2F8F8" bold="true" box="[946,1363,1785,1808]" pageId="12" pageNumber="13">
<emphasis id="8281EAB2FFFE0E50FC07F944FB23F8F8" bold="true" box="[946,1186,1785,1808]" italics="true" pageId="12" pageNumber="13">Canthophorus dubius</emphasis>
(
<bibRefCitation id="D4644B51FFFE0E50FB07F944FACBF8F8" author="Scopoli JA" box="[1202,1354,1785,1808]" journalOrPublisher="Vindobonae: Trattner" pageId="12" pageNumber="13" refId="ref40650" refString="Scopoli JA. 1763. Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana. Vindobonae: Trattner." title="Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana" type="book" year="1763">Scopoli, 1763</bibRefCitation>
)
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<paragraph id="B04A36A0FFFE0E50FCF9F8A4FB4CF8B8" blockId="12.[844,1466,194,1938]" pageId="12" pageNumber="13">
<emphasis id="8281EAB2FFFE0E50FCF9F8A4FB4CF8B8" bold="true" pageId="12" pageNumber="13">
(
<figureCitation id="28CE2A25FFFE0E50FCE1F8A4FC7DF8D8" box="[852,1020,1817,1840]" captionStart="Figure 1" captionStartId="5.[158,225,1575,1596]" captionTargetBox="[320,1303,193,1541]" captionTargetId="figure-79@5.[320,1303,193,1542]" captionTargetPageId="5" captionText="Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency." figureDoi="http://doi.org/10.5281/zenodo.14927019" httpUri="https://zenodo.org/record/14927019/files/figure.png" pageId="12" pageNumber="13">
Figures 1
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<figureCitation id="28CE2A25FFFE0E50FBBFF8A4FBD6F8D8" box="[1034,1111,1816,1840]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="12" pageNumber="13">
3
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,
<figureCitation id="28CE2A25FFFE0E50FBD0F8A4FB33F8D8" box="[1125,1202,1817,1840]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="12" pageNumber="13">
4
<collectionCode id="D6E4AE65FFFE0E50FBC7F8A4FB04F8D8" box="[1138,1157,1817,1840]" pageId="12" pageNumber="13">A</collectionCode>
, 4
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<figureCitation id="28CE2A25FFFE0E50FB0AF8A4FA85F8D8" box="[1215,1284,1816,1840]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="12" pageNumber="13">
6
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<date id="C44B1060FFFE0E50FB58F8A4FA85F8D8" box="[1261,1284,1817,1840]" pageId="12" pageNumber="13">
6
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<figureCitation id="28CE2A25FFFE0E50FAA7F8A4FADEF8D8" box="[1298,1375,1817,1840]" captionStart="Figure 7" captionStartId="11.[158,225,1745,1766]" captionTargetBox="[199,1420,162,1729]" captionTargetId="figure-213@11.[199,1420,159,1729]" captionTargetPageId="11" captionText="Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927031" httpUri="https://zenodo.org/record/14927031/files/figure.png" pageId="12" pageNumber="13">
7
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, 7
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<figureCitation id="28CE2A25FFFE0E50FAD9F8A4FA33F8D8" box="[1388,1458,1817,1840]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="12" pageNumber="13">
8
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8
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<figureCitation id="28CE2A25FFFE0E50FB8DF884FB06F8B8" box="[1080,1159,1849,1872]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="12" pageNumber="13">
9
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, 9
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<figureCitation id="28CE2A25FFFE0E50FB22F884FB45F8B8" box="[1175,1220,1849,1872]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="12" pageNumber="13">
12
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)
</emphasis>
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<paragraph id="B04A36A0FFFE0E50FCF9F8DFFA29F89F" blockId="12.[844,1466,194,1938]" box="[844,1448,1890,1911]" pageId="12" pageNumber="13">
<materialsCitation id="009D3CFDFFFE0E50FCF9F8DFFA29F89F" box="[844,1448,1890,1911]" country="Italy" location="Triest" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Triest" typeStatus="syntype">
<taxonomicName id="77F54D23FFFE0E50FCF9F8DFFB0BF89F" authority="Scopoli 1763: 121" authorityName="Scopoli" authorityPageNumber="121" authorityYear="1763" box="[844,1162,1890,1911]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="8281EAB2FFFE0E50FCF9F8DFFC4EF89F" box="[844,975,1890,1911]" italics="true" pageId="12" pageNumber="13">Cimex dubius</emphasis>
<bibRefCitation id="D4644B51FFFE0E50FC62F8DFFB0BF89F" author="Scopoli JA" box="[983,1162,1890,1911]" journalOrPublisher="Vindobonae: Trattner" pageId="12" pageNumber="13" refId="ref40650" refString="Scopoli JA. 1763. Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana. Vindobonae: Trattner." title="Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana" type="book" year="1763">Scopoli 1763: 121</bibRefCitation>
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[
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:
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,
<collectingRegion id="7231F842FFFE0E50FA87F8DFFAEAF89F" box="[1330,1387,1890,1911]" pageId="12" pageNumber="13">Triest</collectingRegion>
; lost].
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<taxonomicName id="77F54D23FFFE0E50FCF9F8C0FB5CF87A" authority="Schrank 1776: 80" authorityName="Schrank" authorityPageNumber="80" authorityYear="1776" box="[844,1245,1917,1938]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="albomarginatus">
<emphasis id="8281EAB2FFFE0E50FCF9F8C0FBA7F87A" box="[844,1062,1917,1938]" italics="true" pageId="12" pageNumber="13">Cimex albomarginatus</emphasis>
<bibRefCitation id="D4644B51FFFE0E50FB9BF8C0FB5CF87A" author="Schrank FVP" box="[1070,1245,1917,1938]" journalOrPublisher="Leipzig: Fritsch" pageId="12" pageNumber="13" refId="ref40637" refString="Schrank FVP. 1776. Beytrage zur Naturgeschichte. Leipzig: Fritsch." title="Beytrage zur Naturgeschichte" type="book" year="1776">Schrank 1776: 80</bibRefCitation>
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.
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<emphasis id="8281EAB2FFFF0E51FF2BF9D7FF7FF997" bold="true" box="[158,254,1642,1663]" pageId="13" pageNumber="14">Figure 8.</emphasis>
Female terminalia in
<taxonomicName id="77F54D23FFFF0E51FE46F9D7FD4CF997" box="[499,717,1642,1663]" class="Insecta" family="Cydnidae" genus="Canthophorus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="8281EAB2FFFF0E51FE46F9D7FDFFF997" box="[499,638,1642,1663]" italics="true" pageId="13" pageNumber="14">Canthophorus</emphasis>
spp.
<emphasis id="8281EAB2FFFF0E51FD09F9D7FD6DF997" bold="true" box="[700,748,1642,1664]" pageId="13" pageNumber="14">A–G</emphasis>
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,
<taxonomicName id="77F54D23FFFF0E51FD4BF9D7FB2EF997" authority="(Herrich-Schaeffer, 1835)" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[766,1199,1642,1663]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFFF0E51FD4BF9D7FC22F997" box="[766,931,1642,1663]" italics="true" pageId="13" pageNumber="14">C. melanopterus</emphasis>
(
<bibRefCitation id="D4644B51FFFF0E51FC00F9D7FB24F997" author="Herrich-Schaeffer GAW" box="[949,1189,1642,1663]" journalOrPublisher="Regensburg: Pustet" pageId="13" pageNumber="14" refId="ref40016" refString="Herrich-Schaeffer GAW. 1835. Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie. Regensburg: Pustet." title="Nomenclator entomologicus. Verzeichniss der europaischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 1; Lepidoptera und Hemiptera, letztere synoptisch bearbeitet und mit vollstandiger Synonymie" type="journal volume" year="1835">Herrich-Schaeffer, 1835</bibRefCitation>
)
</taxonomicName>
, Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’ skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G);
<emphasis id="8281EAB2FFFF0E51FE44F922FD82F95C" bold="true" box="[497,515,1695,1716]" pageId="13" pageNumber="14">H</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FDA7F922FCDDF95C" authority="(Scopoli, 1763)" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[530,860,1695,1716]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="dubius" subSpecies="dubius">
<emphasis id="8281EAB2FFFF0E51FDA7F922FD3BF95C" box="[530,698,1695,1716]" italics="true" pageId="13" pageNumber="14">C. dubius dubius</emphasis>
(
<bibRefCitation id="D4644B51FFFF0E51FD7DF922FCD2F95C" author="Scopoli JA" box="[712,851,1695,1716]" journalOrPublisher="Vindobonae: Trattner" pageId="13" pageNumber="14" refId="ref40650" refString="Scopoli JA. 1763. Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana. Vindobonae: Trattner." title="Entomologia carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietatis. Methodo Linnaeana" type="book" year="1763">Scopoli, 1763</bibRefCitation>
)
</taxonomicName>
, France: Forêt des Morières;
<emphasis id="8281EAB2FFFF0E51FB37F922FB0AF95C" bold="true" box="[1154,1163,1695,1716]" pageId="13" pageNumber="14">I</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FB2FF922FAE9F95C" box="[1178,1384,1695,1716]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="dubius" subSpecies="sanigarum">
<emphasis id="8281EAB2FFFF0E51FB2FF922FAE9F95C" box="[1178,1384,1695,1716]" italics="true" pageId="13" pageNumber="14">C. dubius sanigarum</emphasis>
</taxonomicName>
<emphasis id="8281EAB2FFFF0E51FAC4F922FA2CF95C" bold="true" box="[1393,1453,1695,1716]" pageId="13" pageNumber="14">n. ssp</emphasis>
., Russia: Krasnaya Polyana, paratype;
<emphasis id="8281EAB2FFFF0E51FDBDF907FDB2F927" bold="true" box="[520,563,1722,1743]" pageId="13" pageNumber="14">J, K</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FDF5F907FC49F927" authority="(Horvath, 1880)" baseAuthorityName="Horvath" baseAuthorityYear="1880" box="[576,968,1722,1743]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="impressus">
<emphasis id="8281EAB2FFFF0E51FDF5F907FCA2F927" box="[576,803,1722,1743]" italics="true" pageId="13" pageNumber="14">C. impressus impressus</emphasis>
(
<bibRefCitation id="D4644B51FFFF0E51FC84F907FC3EF927" author="Horvath G." box="[817,959,1722,1743]" journalOrPublisher="Termeszetrajzi Fuzetek." pageId="13" pageNumber="14" pagination="184 - 192" part="4" refId="ref40056" refString="Horvath G. 1880. Hemipterologiai kozlemenyek. Termeszetrajzi Fuzetek. 4: 184 - 192." title="Hemipterologiai kozlemenyek" type="journal article" year="1880">Horváth, 1880</bibRefCitation>
)
</taxonomicName>
, Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K);
<emphasis id="8281EAB2FFFF0E51FF49F969FE8DF901" bold="true" box="[252,268,1748,1769]" pageId="13" pageNumber="14">L</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FEA9F969FDA0F901" authorityName="Gapon" authorityYear="2018" box="[284,545,1748,1769]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="impressus" subSpecies="hajastanicus">
<emphasis id="8281EAB2FFFF0E51FEA9F969FDA0F901" box="[284,545,1748,1769]" italics="true" pageId="13" pageNumber="14">C. impressus hajastanicus</emphasis>
</taxonomicName>
<emphasis id="8281EAB2FFFF0E51FD9FF969FDE9F901" bold="true" box="[554,616,1748,1769]" pageId="13" pageNumber="14">n. ssp</emphasis>
., Armenia: Gekhmakhan Mt., paratype;
<emphasis id="8281EAB2FFFF0E51FBB1F969FBBAF901" bold="true" box="[1028,1083,1748,1769]" pageId="13" pageNumber="14">M, N</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FBFEF969FA32F901" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[1099,1459,1748,1769]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="wagneri" subSpecies="wagneri">
<emphasis id="8281EAB2FFFF0E51FBFEF969FA95F901" box="[1099,1300,1748,1769]" italics="true" pageId="13" pageNumber="14">C. wagneri wagneri</emphasis>
<bibRefCitation id="D4644B51FFFF0E51FAABF969FA32F901" author="Asanova RB" box="[1310,1459,1748,1769]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="13" pageNumber="14" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Russia: Anapa (M), Armenia: Sevan Lake (N);
<emphasis id="8281EAB2FFFF0E51FD38F952FD38F8EC" bold="true" box="[653,697,1775,1797]" pageId="13" pageNumber="14">O–S</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FD7EF952FB96F8EC" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" box="[715,1047,1775,1796]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="mixtus" subSpecies="mixtus">
<emphasis id="8281EAB2FFFF0E51FD7EF952FCF4F8EC" box="[715,885,1775,1796]" italics="true" pageId="13" pageNumber="14">C. mixtus mixtus</emphasis>
<bibRefCitation id="D4644B51FFFF0E51FC34F952FB96F8EC" author="Asanova RB" box="[897,1047,1775,1796]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="13" pageNumber="14" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S);
<emphasis id="8281EAB2FFFF0E51FA93F8B7FAD0F8F7" bold="true" box="[1318,1361,1802,1823]" pageId="13" pageNumber="14">T, U</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FAEBF8B7FE18F8D1" authority="Asanova, 1964" authorityName="Asanova" authorityYear="1964" baseAuthorityName="Kiritshenko" baseAuthorityYear="1964" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="mixtus" subSpecies="hissaricus">
<emphasis id="8281EAB2FFFF0E51FAEBF8B7FE81F8D1" italics="true" pageId="13" pageNumber="14">C. mixtus hissaricus</emphasis>
<bibRefCitation id="D4644B51FFFF0E51FEB2F899FE18F8D1" author="Asanova RB" box="[263,409,1828,1849]" journalOrPublisher="Entomologicheskoe Obozrenie." pageId="13" pageNumber="14" pagination="138 - 144" part="43" refId="ref39567" refString="Asanova RB. 1964. Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR. Entomologicheskoe Obozrenie. 43: 138 - 144. Russian." title="Genus Canthophorus Muls. et Rey (Heteroptera, Cydnidae) in the fauna of the USSR" type="journal article" year="1964">Asanova, 1964</bibRefCitation>
</taxonomicName>
;
<emphasis id="8281EAB2FFFF0E51FE10F899FE65F8D1" bold="true" box="[421,484,1828,1849]" pageId="13" pageNumber="14">n. stat</emphasis>
., Tajikistan: urochishche Ruydasht [paratypes];
<emphasis id="8281EAB2FFFF0E51FC0CF899FC6BF8D1" bold="true" box="[953,1002,1828,1849]" pageId="13" pageNumber="14">V, W</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FC43F899FAA5F8D1" authority="Scott, 1874" authorityName="Scott" authorityYear="1874" box="[1014,1316,1828,1849]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="niveimarginatus">
<emphasis id="8281EAB2FFFF0E51FC43F899FB2EF8D1" box="[1014,1199,1828,1849]" italics="true" pageId="13" pageNumber="14">C. niveimarginatus</emphasis>
<bibRefCitation id="D4644B51FFFF0E51FB00F899FAA5F8D1" author="Scott J." box="[1205,1316,1828,1849]" journalOrPublisher="Annals and Magazine of Natural History." pageId="13" pageNumber="14" pagination="289 - 304" part="14" refId="ref40679" refString="Scott J. 1874. On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species. Annals and Magazine of Natural History. 14 (ser. 4): 289 - 304; 360 - 365; 426 - 452." title="On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species" type="journal article" year="1874">Scott, 1874</bibRefCitation>
</taxonomicName>
; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W);
<emphasis id="8281EAB2FFFF0E51FDE5F882FDFEF8BC" bold="true" box="[592,639,1855,1876]" pageId="13" pageNumber="14">X, Y</emphasis>
,
<taxonomicName id="77F54D23FFFF0E51FD3AF882FC26F8BC" authority="(Reuter, 1902)" baseAuthorityName="Reuter" baseAuthorityYear="1902" box="[655,935,1855,1876]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="8281EAB2FFFF0E51FD3AF882FC8CF8BC" box="[655,781,1855,1876]" italics="true" pageId="13" pageNumber="14">C. coeruleus</emphasis>
(
<bibRefCitation id="D4644B51FFFF0E51FCA8F882FC1CF8BC" author="Reuter OM" box="[797,925,1855,1876]" journalOrPublisher="Ofversigt af Finska Vetenskaps-Societetens Forhandlingar." pageId="13" pageNumber="14" pagination="141 - 188" part="44" refId="ref40588" refString="Reuter OM. 1902. Miscellanea Hemipterologica. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar. 44: 141 - 188." title="Miscellanea Hemipterologica" type="journal article" year="1902">Reuter, 1902</bibRefCitation>
)
</taxonomicName>
, Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y).
<emphasis id="8281EAB2FFFF0E51FE46F8C9FD93F861" box="[499,530,1908,1929]" italics="true" pageId="13" pageNumber="14">a.g</emphasis>
– anterior part of gynatrial sac;
<emphasis id="8281EAB2FFFF0E51FCEBF8C9FCFCF861" box="[862,893,1908,1929]" italics="true" pageId="13" pageNumber="14">d.d</emphasis>
– distal part of spermathecal duct;
<emphasis id="8281EAB2FFFF0E51FB52F8C9FB7FF861" box="[1255,1278,1908,1929]" italics="true" pageId="13" pageNumber="14">f.r</emphasis>
– first rami;
<emphasis id="8281EAB2FFFF0E51FA32F8C9FA25F861" box="[1415,1444,1908,1929]" italics="true" pageId="13" pageNumber="14">g.c</emphasis>
– gynatrial cone;
<emphasis id="8281EAB2FFFF0E51FE88F833FED4F84B" box="[317,341,1934,1955]" italics="true" pageId="13" pageNumber="14">gn</emphasis>
– gonangulum;
<emphasis id="8281EAB2FFFF0E51FE4AF833FDA7F84B" box="[511,550,1934,1955]" italics="true" pageId="13" pageNumber="14">gp.I</emphasis>
– gonapophyses I;
<emphasis id="8281EAB2FFFF0E51FD47F833FCA1F84B" box="[754,800,1934,1955]" italics="true" pageId="13" pageNumber="14">gp.II</emphasis>
– gonapophyses II;
<emphasis id="8281EAB2FFFF0E51FC41F833FBA1F84B" box="[1012,1056,1934,1955]" italics="true" pageId="13" pageNumber="14">gx.II</emphasis>
– gonocoxites II;
<emphasis id="8281EAB2FFFF0E51FB54F833FA84F84B" box="[1249,1285,1934,1955]" italics="true" pageId="13" pageNumber="14">m.d</emphasis>
– median part of spermathecal duct;
<emphasis id="8281EAB2FFFF0E51FEE9F814FEFBF856" box="[348,378,1961,1982]" italics="true" pageId="13" pageNumber="14">p.d</emphasis>
– proximal part of spermathecal duct;
<emphasis id="8281EAB2FFFF0E51FD4FF814FC99F856" box="[762,792,1961,1982]" italics="true" pageId="13" pageNumber="14">p.g</emphasis>
– posterior part of gynatrial sac;
<emphasis id="8281EAB2FFFF0E51FBD7F814FB04F856" box="[1122,1157,1961,1982]" italics="true" pageId="13" pageNumber="14">p.m</emphasis>
– posterior margin of gynatrial sac;
<emphasis id="8281EAB2FFFF0E51FF79F879FF6BF831" box="[204,234,1988,2009]" italics="true" pageId="13" pageNumber="14">p.p</emphasis>
– posterior pouches of gynatrial sac;
<emphasis id="8281EAB2FFFF0E51FDD6F879FDFBF831" box="[611,634,1988,2009]" italics="true" pageId="13" pageNumber="14">r.s</emphasis>
– ring sclerites;
<emphasis id="8281EAB2FFFF0E51FC97F879FCB3F831" box="[802,818,1988,2009]" italics="true" pageId="13" pageNumber="14">tr</emphasis>
– triangulum. Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph id="B04A36A0FFFC0E52FF2BFF7CFE70FF3E" blockId="14.[158,781,193,1938]" box="[158,497,193,214]" pageId="14" pageNumber="15">
<taxonomicName id="77F54D23FFFC0E52FF2BFF7CFE6CFF3E" authority="Goeze" authorityName="Goeze" authorityPageNumber="276" authorityYear="1778" box="[158,493,193,214]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="histriolus">
<emphasis id="8281EAB2FFFC0E52FF2BFF7CFEBDFF3E" box="[158,316,193,214]" italics="true" pageId="14" pageNumber="15">Cimex histriolus</emphasis>
<bibRefCitation id="D4644B51FFFC0E52FEF1FF7CFE6CFF3E" author="Goeze JAE" box="[324,493,193,214]" journalOrPublisher="Leipzig: Weidmanns Erben &amp; Reich" pageId="14" pageNumber="15" refId="ref39989" refString="Goeze JAE. 1778. Entomologische Beytrage zu des Ritter Linne zwolften Ausgabe des Natursystems. Vol. 2, Leipzig: Weidmanns Erben &amp; Reich." title="Entomologische Beytrage zu des Ritter Linne zwolften Ausgabe des Natursystems" type="book" year="1778">Goeze 1778: 276</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFF61FD90FF19" blockId="14.[158,781,193,1938]" box="[158,529,220,241]" pageId="14" pageNumber="15">
<taxonomicName id="77F54D23FFFC0E52FF2BFF61FD8CFF19" authority="Gmelin 1790: 2159" authorityName="Gmelin" authorityPageNumber="2159" authorityYear="1790" box="[158,525,220,241]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="viennensis">
<emphasis id="8281EAB2FFFC0E52FF2BFF61FEC7FF19" box="[158,326,220,241]" italics="true" pageId="14" pageNumber="15">Cimex viennensis</emphasis>
<bibRefCitation id="D4644B51FFFC0E52FEF8FF61FD8CFF19" author="Gmelin JF" box="[333,525,220,241]" journalOrPublisher="Lipsiae: Beer" pageId="14" pageNumber="15" part="1" refId="ref39942" refString="Gmelin JF. 1790. Caroli a Linne, systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (edn 13). 1 (4). Lipsiae: Beer." title="Caroli a Linne, systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" type="journal volume" year="1790">Gmelin 1790: 2159</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFF4AFDB0FEE4" blockId="14.[158,781,193,1938]" box="[158,561,247,268]" pageId="14" pageNumber="15">
<taxonomicName id="77F54D23FFFC0E52FF2BFF4AFDACFEE4" authority="Gmelin 1790: 2161" authorityName="Gmelin" authorityPageNumber="2161" authorityYear="1790" box="[158,557,247,268]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="melanoleucos">
<emphasis id="8281EAB2FFFC0E52FF2BFF4AFEE4FEE4" box="[158,357,247,268]" italics="true" pageId="14" pageNumber="15">Cimex melanoleucos</emphasis>
<bibRefCitation id="D4644B51FFFC0E52FED8FF4AFDACFEE4" author="Gmelin JF" box="[365,557,247,268]" journalOrPublisher="Lipsiae: Beer" pageId="14" pageNumber="15" part="1" refId="ref39942" refString="Gmelin JF. 1790. Caroli a Linne, systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (edn 13). 1 (4). Lipsiae: Beer." title="Caroli a Linne, systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" type="journal volume" year="1790">Gmelin 1790: 2161</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFEACFDCFFECE" blockId="14.[158,781,193,1938]" box="[158,590,273,294]" pageId="14" pageNumber="15">
<taxonomicName id="77F54D23FFFC0E52FF2BFEACFDCBFECE" authority="Fabricius 1794: 123" authorityName="Fabricius" authorityPageNumber="123" authorityYear="1794" box="[158,586,273,294]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="albomarginellus">
<emphasis id="8281EAB2FFFC0E52FF2BFEACFEFFFECE" box="[158,382,273,294]" italics="true" pageId="14" pageNumber="15">Cimex albomarginellus</emphasis>
<bibRefCitation id="D4644B51FFFC0E52FE30FEACFDCBFECE" author="Fabricius JC" box="[389,586,273,294]" journalOrPublisher="Hafniae: Proft" pageId="14" pageNumber="15" part="4" refId="ref39642" refString="Fabricius JC. 1794. Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Vol. 4, Hafniae: Proft." title="Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus" type="journal volume" year="1794">Fabricius 1794: 123</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFE91FC8CFEA9" blockId="14.[158,781,193,1938]" box="[158,781,300,321]" pageId="14" pageNumber="15">
<taxonomicName id="77F54D23FFFC0E52FF2BFE91FD8BFEA9" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[158,522,300,321]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dubius" subGenus="Canthophorus">
<emphasis id="8281EAB2FFFC0E52FF2BFE91FEA7FEA9" box="[158,294,300,321]" italics="true" pageId="14" pageNumber="15">Canthophorus</emphasis>
(
<emphasis id="8281EAB2FFFC0E52FE80FE91FE3CFEA9" box="[309,445,300,321]" italics="true" pageId="14" pageNumber="15">Canthophorus</emphasis>
)
<emphasis id="8281EAB2FFFC0E52FE7EFE91FD8BFEA9" box="[459,522,300,321]" italics="true" pageId="14" pageNumber="15">dubius</emphasis>
</taxonomicName>
:
<bibRefCitation id="D4644B51FFFC0E52FDADFE91FC88FEA9" author="Mulsant E &amp; Rey C." box="[536,777,300,321]" journalOrPublisher="Paris: Savy &amp; Deyrolle" pageId="14" pageNumber="15" refId="ref40479" refString="Mulsant E, Rey C. 1866. Histoire naturelle des punaises de France. Pentatomides. Paris: Savy &amp; Deyrolle." title="Histoire naturelle des punaises de France. Pentatomides" type="book" year="1866">Mulsant &amp; Rey 1866: 63</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8EF652BFFFC0E52FF2BFE3AFA84FEF1" pageId="14" pageNumber="15" type="description">
<paragraph id="B04A36A0FFFC0E52FF2BFE3AFEA1FE76" blockId="14.[158,781,193,1938]" box="[158,288,391,414]" pageId="14" pageNumber="15">
<heading id="EB0281CCFFFC0E52FF2BFE3AFEA1FE76" bold="true" box="[158,288,391,414]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="6">
<emphasis id="8281EAB2FFFC0E52FF2BFE3AFEA1FE76" bold="true" box="[158,288,391,414]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFE12FF71FDAE" blockId="14.[158,781,193,1938]" pageId="14" pageNumber="15">Lateral margins of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metasternal evaporatorium with long and curved peritreme. Membrane brownish or whitish.</paragraph>
<paragraph id="B04A36A0FFFC0E52FF15FDE2FE77FD3D" blockId="14.[158,781,193,1938]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF15FDE2FE94FD9E" bold="true" box="[160,277,607,630]" italics="true" pageId="14" pageNumber="15">Pygophore</emphasis>
slightly longer than wide; lateral walls very slightly concave in middle of length; dorsal infolding slightly shorter, without impression, dorsal wall slightly longer than in
<taxonomicName id="77F54D23FFFC0E52FE8BFD03FE6EFD3D" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1835" box="[318,495,702,725]" class="Insecta" family="Cydnidae" genus="Canthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="melanopterus">
<emphasis id="8281EAB2FFFC0E52FE8BFD03FE6EFD3D" box="[318,495,702,725]" italics="true" pageId="14" pageNumber="15">C. melanopterus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFD49FD4BFCC3" blockId="14.[158,781,193,1938]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFD49FE4EFCE3" bold="true" box="[158,463,755,779]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFD49FE88FCE3" bold="true" box="[158,265,756,779]" italics="true" pageId="14" pageNumber="15">Paramere</emphasis>
(
<figureCitation id="28CE2A25FFFC0E52FEA8FD49FE3EFCE3" box="[285,447,755,779]" captionStart="Figure 3" captionStartId="7.[158,225,1667,1688]" captionTargetBox="[208,1412,184,1636]" captionTargetId="figure-177@7.[200,1419,169,1651]" captionTargetPageId="7" captionText="Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant &amp; Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927023" httpUri="https://zenodo.org/record/14927023/files/figure.png" pageId="14" pageNumber="15">
Figure 3
<collectionCode id="D6E4AE65FFFC0E52FECBFD49FE10FCE3" box="[382,401,756,779]" pageId="14" pageNumber="15">A</collectionCode>
–3
<collectionCode id="D6E4AE65FFFC0E52FE19FD49FE3EFCE3" box="[428,447,756,779]" pageId="14" pageNumber="15">D</collectionCode>
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).
</emphasis>
Hypophysis strongly curved dorsally, so that its apex is visible from outer view.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFCF4FD44FA56" blockId="14.[158,781,193,1938]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFCF4FE4DFC88" bold="true" box="[158,460,841,864]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFCF4FE89FC88" bold="true" box="[158,264,841,864]" italics="true" pageId="14" pageNumber="15">Aedeagus</emphasis>
(
<figureCitation id="28CE2A25FFFC0E52FEAFFCF4FE3CFC88" box="[282,445,841,864]" captionStart="Figure 4" captionStartId="8.[158,225,1769,1790]" captionTargetBox="[198,1428,182,1738]" captionTargetId="figure-124@8.[195,1428,168,1753]" captionTargetPageId="8" captionText="Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927025" httpUri="https://zenodo.org/record/14927025/files/figure.png" pageId="14" pageNumber="15">
Figure 4
<collectionCode id="D6E4AE65FFFC0E52FECFFCF4FE0CFC88" box="[378,397,841,864]" pageId="14" pageNumber="15">A</collectionCode>
, 4
<collectionCode id="D6E4AE65FFFC0E52FE1EFCF4FE3CFC88" box="[427,445,841,864]" pageId="14" pageNumber="15">B</collectionCode>
</figureCitation>
).
</emphasis>
Ventral lobe of conjunctiva barely reaching apical margin of theca and narrowed before widened apex; latter with two small lateral denticles directed laterad and with two nariform impressions dorsal to them (
<figureCitation id="28CE2A25FFFC0E52FE45FC74FC82FC08" box="[496,771,969,992]" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="14" pageNumber="15">
Figure 6
<collectionCode id="D6E4AE65FFFC0E52FDE7FC74FDE7FC08" box="[594,614,969,992]" pageId="14" pageNumber="15">A</collectionCode>
, 6
<collectionCode id="D6E4AE65FFFC0E52FD3CFC74FD1AFC08" box="[649,667,969,992]" pageId="14" pageNumber="15">C</collectionCode>
, 6
<collectionCode id="D6E4AE65FFFC0E52FD08FC74FD50FC08" box="[701,721,969,992]" pageId="14" pageNumber="15">D</collectionCode>
, 6
<collectionCode id="D6E4AE65FFFC0E52FD46FC74FC82FC08" box="[755,771,969,992]" pageId="14" pageNumber="15">F</collectionCode>
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) or rarely without denticles and impressions (
<figureCitation id="28CE2A25FFFC0E52FD26FC54FF35FBC8" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="14" pageNumber="15">
Figure 6
<collectionCode id="D6E4AE65FFFC0E52FD45FC54FC85FBE8" box="[752,772,1001,1024]" pageId="14" pageNumber="15">G</collectionCode>
,
<date id="C44B1060FFFC0E52FF2BFBB4FF35FBC8" box="[158,180,1033,1056]" pageId="14" pageNumber="15">
6
<collectionCode id="D6E4AE65FFFC0E52FF1EFBB4FF35FBC8" box="[171,180,1033,1056]" pageId="14" pageNumber="15">I</collectionCode>
</date>
</figureCitation>
); ventral surface of lobe slightly convex, without carina; in lateral view, lobe rather narrow (
<figureCitation id="28CE2A25FFFC0E52FD27FB94FF72FBB7" captionStart="Figure 6" captionStartId="10.[158,225,1240,1261]" captionTargetBox="[204,1413,190,1211]" captionTargetId="figure-445@10.[199,1418,186,1219]" captionTargetPageId="10" captionText="Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927029" httpUri="https://zenodo.org/record/14927029/files/figure.png" pageId="14" pageNumber="15">
Figure 6
<collectionCode id="D6E4AE65FFFC0E52FD46FB94FC84FBA8" box="[755,773,1065,1088]" pageId="14" pageNumber="15">B</collectionCode>
, 6
<collectionCode id="D6E4AE65FFFC0E52FF1EFBF5FF3DFBB7" box="[171,188,1096,1119]" pageId="14" pageNumber="15">E</collectionCode>
, 6
<collectionCode id="D6E4AE65FFFC0E52FF6AFBF5FF72FBB7" box="[223,243,1096,1119]" pageId="14" pageNumber="15">H</collectionCode>
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). Apical lobes of conjunctiva short, directed ventroposteriad, lying completely in dorsoventral plane of apical half of conjunctiva and curved one towards other in middle. Vesica without lateral branch. Dorsolateral lobes of conjunctiva long, basally divided by impression, divergent in a V-shape, directed dorsolaterad, and curved anteriad at apices. Anterior wall of each dorsolateral lobe with area of weak sclerotisation before apex. Spicule broad at base, tapering to pointed apex, basally directed anteriad and dorsad, curved mediad before apex. Inner end of spicule basal plate with one short conical denticle directed mediad.
</paragraph>
<paragraph id="B04A36A0FFFC0E52FF2BFA60FA84FEF1" blockId="14.[158,781,193,1938]" lastBlockId="14.[843,1466,194,1938]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFA60FE27FA1C" bold="true" box="[158,422,1501,1524]" pageId="14" pageNumber="15">
<emphasis id="8281EAB2FFFC0E52FF2BFA60FE94FA1C" bold="true" box="[158,277,1501,1524]" italics="true" pageId="14" pageNumber="15">Gynatrium</emphasis>
(
<figureCitation id="28CE2A25FFFC0E52FE91FA60FE16FA1C" box="[292,407,1501,1524]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="14" pageNumber="15">
Figure 9
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</figureCitation>
).
</emphasis>
Posterior pouches located behind posterior margin of gynatrial sac; line of posterior margin laterally curved posteriad and extending on dorsal walls of pouches (
<figureCitation id="28CE2A25FFFC0E52FEB9F981FE0CF9BB" box="[268,397,1596,1619]" captionStart="Figure 12" captionStartId="18.[158,225,790,811]" captionTargetBox="[179,1446,162,769]" captionTargetId="figure-569@18.[179,1446,162,769]" captionTargetPageId="18" captionText="Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927041" httpUri="https://zenodo.org/record/14927041/files/figure.png" pageId="14" pageNumber="15">
Figure 12
<collectionCode id="D6E4AE65FFFC0E52FECCF981FE0CF9BB" box="[377,397,1596,1619]" pageId="14" pageNumber="15">A</collectionCode>
</figureCitation>
). They are rather large, approximately as long as wide, with wide base and widely rounded posterior margin in dorsal view. Their walls rather soft, inner walls evenly convex, not sloped. Pouches subtransparent, without any sclerotisations inside. Anterior part of gynatrial sac very long and wide. Proximal part of spermathecal duct very short; median part elongated; distal part as long as half of median part or slightly shorter. Tubular part of spermathecal capsule approximately as long as transverse diameter of its spherical part and curved, but not gyrose (
<figureCitation id="28CE2A25FFFC0E52FDE4F8C6FD40F87A" box="[593,705,1915,1938]" captionStart="Figure 9" captionStartId="15.[158,225,1499,1520]" captionTargetBox="[180,761,175,1475]" captionTargetId="figure-435@15.[175,766,163,1481]" captionTargetPageId="15" captionText="Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927035" httpUri="https://zenodo.org/record/14927035/files/figure.png" pageId="14" pageNumber="15">
Figure 9
<collectionCode id="D6E4AE65FFFC0E52FD1AF8C6FD40F87A" box="[687,705,1915,1938]" pageId="14" pageNumber="15">B</collectionCode>
</figureCitation>
). First rami (
<figureCitation id="28CE2A25FFFC0E52FC38FF7FFB9EFF31" box="[909,1055,194,217]" captionStart="Figure 8" captionStartId="13.[158,225,1642,1663]" captionTargetBox="[217,1399,160,1624]" captionTargetId="figure-308@13.[217,1399,154,1624]" captionTargetPageId="13" captionText="Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14927033" httpUri="https://zenodo.org/record/14927033/files/figure.png" pageId="14" pageNumber="15">
Figure 8
<collectionCode id="D6E4AE65FFFC0E52FC53FF7FFC7AFF31" box="[998,1019,194,217]" pageId="14" pageNumber="15">H</collectionCode>
, 8
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) rather broadly widened anteriorly, at anterior ends truncate at acute angle, rarely slightly widening and with rounded anterior end.
</paragraph>
</subSubSection>
</treatment>
</document>

74
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@ -1,74 +0,0 @@
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Populations of
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Males vocalize after rains at night in shallow puddles at the edge of shallow streams, lakes, or in areas of low and sparse vegetation near pasture or cultivated fields. The calls are composed of a sequence of approximately 153 notes with a duration of around 5 seconds, emitted at a rate of 4 calls per minute and 28 notes per second. The calls reach maximum amplitude rapidly and remain constant until the end (
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Populations of
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Males vocalize after rains at night in shallow puddles at the edge of shallow streams, lakes, or in areas of low and sparse vegetation near pasture or cultivated fields. The calls are composed of a sequence of approximately 153 notes with a duration of around 5 seconds, emitted at a rate of 4 calls per minute and 28 notes per second. The calls reach maximum amplitude rapidly and remain constant until the end (
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<emphasis id="5F8FC85EFFBCFFEEA0B8FBAAFECDFBCD" bold="true" box="[151,275,1100,1124]" pageId="8" pageNumber="113">FIGURE 3.</emphasis>
Audio spectrogram and corresponding waveform of the advertisement call of a topotype
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Considering a 614-base pair sequences of the mitochondrial 16S rRNA fragment, the
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<mods:namePart id="CC650CA324EB8B2BF4917C5EF383F5BA">Giaretta, Ariovaldo Antonio</mods:namePart>
<mods:affiliation id="816AEE5BCAC8AA4F101AED9DE63008C1">Laboratório de Taxonomia e Sistemática de Anuros Neotropicais. Faculdades Integradas do Pontal. Universidade Federal de Uberlândia. Rua 20 Nº 1600 - Bairro Tupã 38304 - 402. Ituiutaba (CEP: 38302 - 000). Minas Gerais, Brasil</mods:affiliation>
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<mods:title id="ACE4996ABDC0441C7013E48B00D941E5">Zootaxa</mods:title>
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<mods:date id="A90EA851CB1BAA66CF496011C9C47C7D">2025</mods:date>
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<mods:number id="F7E7239F8C03188ECB751ACEECCC0072">5588</mods:number>
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<subSubSection id="25E147C7FFB6FFE4A0B8FAB9FD25FAD0" box="[151,763,1375,1402]" pageId="2" pageNumber="107" type="nomenclature">
<paragraph id="6D44144CFFB6FFE4A0B8FAB9FD25FAD0" blockId="2.[151,763,1375,1402]" box="[151,763,1375,1402]" pageId="2" pageNumber="107">
<heading id="360CA320FFB6FFE4A0B8FAB9FD25FAD0" bold="true" box="[151,763,1375,1402]" fontSize="11" level="1" pageId="2" pageNumber="107" reason="1">
<emphasis id="5F8FC85EFFB6FFE4A0B8FAB9FD25FAD0" bold="true" box="[151,763,1375,1402]" pageId="2" pageNumber="107">
<taxonomicName id="AAFB6FCFFFB6FFE4A0B8FAB9FE19FAD3" ID-CoL="4S7P3" baseAuthorityName="Gallardo" baseAuthorityYear="1965" box="[151,455,1375,1402]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="107" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB6FFE4A0B8FAB9FE19FAD3" bold="true" box="[151,455,1375,1402]" italics="true" pageId="2" pageNumber="107">Rhinella mirandaribeiroi</emphasis>
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(Cerrado vs.
<typeStatus id="B240AAEEFFB6FFE4A25DFAB9FD2AFAD0" box="[626,756,1375,1401]" pageId="2" pageNumber="107" type="topotype">Topotypes</typeStatus>
)
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</paragraph>
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<subSubSection id="25E147C7FFB6FFE0A0B8FA41FE69FA0B" lastPageId="6" lastPageNumber="111" pageId="2" pageNumber="107" type="description">
<paragraph id="6D44144CFFB6FFE4A0B8FA41FDF0FA68" blockId="2.[151,558,1447,1473]" box="[151,558,1447,1473]" pageId="2" pageNumber="107">
<heading id="360CA320FFB6FFE4A0B8FA41FDF0FA68" bold="true" box="[151,558,1447,1473]" fontSize="11" level="1" pageId="2" pageNumber="107" reason="1">
<emphasis id="5F8FC85EFFB6FFE4A0B8FA41FDF0FA68" bold="true" box="[151,558,1447,1473]" pageId="2" pageNumber="107">Morphologic and Morphometric</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFB6FFE4A0B8FA09FA47F948" blockId="2.[151,1437,1519,1762]" pageId="2" pageNumber="107">
Morphometric variable values can be found in
<tableCitation id="207921F7FFB6FFE4A280FA09FCDAF9A0" box="[687,772,1519,1545]" captionStart="TABLE 1" captionStartId="5.[152,244,151,175]" captionTargetBox="[163,1335,228,1929]" captionText="TABLE 1. Morphometric variable values for three populations of Rhinella mirandaribeiroi. N = number of analyzed adult males; data is presented as mean ± standard deviation (minimum–maximum)." httpUri="http://table.plazi.org/id/398444C4FFB1FFE3A0B7FF71FBCEFF7A" pageId="2" pageNumber="107" tableUuid="398444C4FFB1FFE3A0B7FF71FBCEFF7A">Table 1</tableCitation>
. The
<typeStatus id="B240AAEEFFB6FFE4A369FA16FC6DF9A3" box="[838,947,1520,1546]" pageId="2" pageNumber="107" type="topotype">topotypes</typeStatus>
of
<taxonomicName id="AAFB6FCFFFB6FFE4A3F3FA16FB71F9A0" baseAuthorityName="Gallardo" baseAuthorityYear="1965" box="[988,1199,1519,1545]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="2" pageNumber="107" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB6FFE4A3F3FA16FB71F9A0" box="[988,1199,1519,1545]" italics="true" pageId="2" pageNumber="107">R. mirandaribeiroi</emphasis>
</taxonomicName>
are morphologically similar to
<emphasis id="5F8FC85EFFB6FFE4A122F9F2FEA2F984" box="[269,380,1555,1581]" italics="true" pageId="2" pageNumber="107">B. g. lutzi</emphasis>
and
<typeStatus id="B240AAEEFFB6FFE4A199F9F5FDB0F984" box="[438,622,1555,1581]" isOtherTypeStatus="true" pageId="2" pageNumber="107">other specimens</typeStatus>
from the
<location id="68244297FFB6FFE4A2CEF9F5FCE0F984" LSID="urn:lsid:plazi:treatment:E552A55AFFB6FFE0A0B8FAB9FE69FA0B:68244297FFB6FFE4A2CEF9F5FCE0F984" box="[737,830,1555,1581]" county="Neither" municipality="Loreal region" name="Cerrado" pageId="2" pageNumber="107">Cerrado</location>
, and they can be diagnosed from most species in the group by: granular or continuous cranial crests, low supraorbital crest, infraorbital crest extending usually beyond the postorbital crest, generally discontinuous reduced maxillary crest, and absence of a parietal crest (
<figureCitation id="F5C008C9FFB6FFE4A52AF9BDFA51F9DC" box="[1285,1423,1627,1653]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[152,255,1467,1491]" captionStartId-1="4.[151,250,1909,1933]" captionTargetBox-0="[161,1426,191,1433]" captionTargetBox-1="[295,1292,188,1878]" captionTargetId-0="figure-17@3.[151,1436,181,1444]" captionTargetId-1="figure-24@4.[288,1299,181,1885]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Dorsal and lateral view of the head of adult males of. R. mirandaribeiroi and R. granulosa: (A) Soure, Ilha de Marajó, Pará state (topotype; AAG-UFU 6564). (B) Pirapora, Minas Gerais state (topotype of B. g. lutzi; AAG-UFU 7133). R. granulosa: (C) Salinas, Minas Gerais state (CFBH 47633). Note the absence of distinctive elements between populations and species in supraorbital crest (Sc), infraorbital crest (Ic) (extending or not beyond the postorbital crest), tympanum near the postorbital crest (Tp), and the presence of a clear longitudinal vertebral line (Lv)." captionText-1="FIGURE 2. Dorsal and ventral view of adult males of R. mirandaribeiroi and R. granulosa. (A) R. mirandaribeiroi, Soure, Ilha de Marajó, Pará state (topotypes; AAG-UFU 6542). (B) (topotypes of B. g. lutzi; AAG-UFU 7134 Pirapora, Minas de Gerais state). (C) R. granulosa: Salinas, Minas Gerais state (CFBH 47633). Note the lack of distinctive elements between populations and species." figureDoi-0="http://doi.org/10.5281/zenodo.14896120" figureDoi-1="http://doi.org/10.5281/zenodo.14896122" httpUri-0="https://zenodo.org/record/14896120/files/figure.png" httpUri-1="https://zenodo.org/record/14896122/files/figure.png" pageId="2" pageNumber="107">Figures 1–2</figureCitation>
).
<collectingMunicipality id="8D208E36FFB6FFE4A0B8F999FEEFF930" box="[151,305,1663,1689]" pageId="2" pageNumber="107">Loreal region</collectingMunicipality>
clearly visible in dorsal view, tympanum very close to the postorbital crest (DTP 0.4–1.0% CRC), a usually present clear longitudinal vertebral line, and a light cream or yellowish ventral side, with or without pigmentation.
<collectingCounty id="84256CC0FFB6FFE4A116F921FE53F948" box="[313,397,1735,1761]" pageId="2" pageNumber="107">Neither</collectingCounty>
morphological nor morphometric characters allowed for the diagnosis between populations.
</paragraph>
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<paragraph id="6D44144CFFB7FFE5A0B7FA5DFC11F9CD" blockId="3.[151,1437,1467,1636]" pageId="3" pageNumber="108">
<emphasis id="5F8FC85EFFB7FFE5A0B7FA5DFEC4FA7A" bold="true" box="[152,282,1467,1491]" pageId="3" pageNumber="108">FIGURE 1.</emphasis>
Dorsal and lateral view of the head of adult males of.
<taxonomicName id="AAFB6FCFFFB7FFE5A378FA5AFBC9FA7A" baseAuthorityName="Gallardo" baseAuthorityYear="1965" box="[855,1047,1467,1491]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="108" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB7FFE5A378FA5AFBC9FA7A" box="[855,1047,1467,1491]" italics="true" pageId="3" pageNumber="108">R. mirandaribeiroi</emphasis>
</taxonomicName>
and
<taxonomicName id="AAFB6FCFFFB7FFE5A463FA5AFB0FFA7A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1100,1233,1467,1491]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="108" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFB7FFE5A463FA5AFB0FFA7A" box="[1100,1233,1467,1491]" italics="true" pageId="3" pageNumber="108">R. granulosa</emphasis>
</taxonomicName>
: (A) Soure, Ilha de Marajó, Pará state (topotype; AAG-UFU 6564). (B) Pirapora, Minas Gerais state (topotype of
<emphasis id="5F8FC85EFFB7FFE5A442FA06FB0DFA5E" box="[1133,1235,1503,1527]" italics="true" pageId="3" pageNumber="108">B. g. lutzi</emphasis>
; AAG-UFU 7133).
<taxonomicName id="AAFB6FCFFFB7FFE5A0B8F9E2FEC2F9B2" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[151,284,1539,1563]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="108" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFB7FFE5A0B8F9E2FEC2F9B2" box="[151,284,1539,1563]" italics="true" pageId="3" pageNumber="108">R. granulosa</emphasis>
</taxonomicName>
: (C) Salinas, Minas Gerais state (CFBH 47633). Note the absence of distinctive elements between populations and species in supraorbital crest (Sc), infraorbital crest (Ic) (extending or not beyond the postorbital crest), tympanum near the postorbital crest (Tp), and the presence of a clear longitudinal vertebral line (Lv).
</paragraph>
</caption>
<caption id="398444C4FFB0FFE2A0B8F893FECDF850" ID-DOI="http://doi.org/10.5281/zenodo.14896122" ID-Zenodo-Dep="14896122" httpUri="https://zenodo.org/record/14896122/files/figure.png" pageId="4" pageNumber="109" startId="4.[151,250,1909,1933]" targetBox="[295,1292,188,1878]" targetPageId="4" targetType="figure">
<paragraph id="6D44144CFFB0FFE2A0B8F893FECDF850" blockId="4.[151,1436,1909,2041]" pageId="4" pageNumber="109">
<emphasis id="5F8FC85EFFB0FFE2A0B8F893FECCF824" bold="true" box="[151,274,1909,1933]" pageId="4" pageNumber="109">FIGURE 2.</emphasis>
Dorsal and ventral view of adult males of
<taxonomicName id="AAFB6FCFFFB0FFE2A291F890FCA2F824" baseAuthorityName="Gallardo" baseAuthorityYear="1965" box="[702,892,1909,1933]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="109" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB0FFE2A291F890FCA2F824" box="[702,892,1909,1933]" italics="true" pageId="4" pageNumber="109">R. mirandaribeiroi</emphasis>
</taxonomicName>
and
<taxonomicName id="AAFB6FCFFFB0FFE2A383F890FBF1F824" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[940,1071,1909,1933]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="109" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFB0FFE2A383F890FBF1F824" box="[940,1071,1909,1933]" italics="true" pageId="4" pageNumber="109">R. granulosa</emphasis>
</taxonomicName>
. (A)
<taxonomicName id="AAFB6FCFFFB0FFE2A44BF890FAB2F824" authority=", Soure" authorityName="Soure" box="[1124,1388,1909,1933]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="109" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB0FFE2A44BF890FAFFF824" box="[1124,1313,1909,1933]" italics="true" pageId="4" pageNumber="109">R. mirandaribeiroi</emphasis>
, Soure
</taxonomicName>
, Ilha de Marajó, Pará state (topotypes; AAG-UFU 6542). (B) (topotypes of
<emphasis id="5F8FC85EFFB0FFE2A344F87CFC11F818" box="[875,975,1945,1969]" italics="true" pageId="4" pageNumber="109">B. g. lutzi</emphasis>
; AAG-UFU 7134 Pirapora, Minas de Gerais state). (C)
<emphasis id="5F8FC85EFFB0FFE2A12CF858FE50F87C" box="[259,398,1981,2005]" italics="true" pageId="4" pageNumber="109">
<taxonomicName id="AAFB6FCFFFB0FFE2A12CF858FE57F87C" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[259,393,1981,2005]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="109" phylum="Chordata" rank="species" species="granulosa">R. granulosa</taxonomicName>
:
</emphasis>
Salinas, Minas Gerais state (CFBH 47633). Note the lack of distinctive elements between populations and species.
</paragraph>
</caption>
<caption id="398444C4FFB1FFE3A0B7FF71FBCEFF7A" ID-Table-UUID="398444C4FFB1FFE3A0B7FF71FBCEFF7A" httpUri="http://table.plazi.org/id/398444C4FFB1FFE3A0B7FF71FBCEFF7A" pageId="5" pageNumber="110" startId="5.[152,244,151,175]" targetBox="[163,1335,228,1929]" targetIsTable="true" targetPageId="5">
<paragraph id="6D44144CFFB1FFE3A0B7FF71FBCEFF7A" blockId="5.[151,1436,151,211]" pageId="5" pageNumber="110">
<emphasis id="5F8FC85EFFB1FFE3A0B7FF71FECFFF06" bold="true" box="[152,273,151,175]" pageId="5" pageNumber="110">TABLE 1.</emphasis>
Morphometric variable values for three populations of
<taxonomicName id="AAFB6FCFFFB1FFE3A35CFF71FB5FFF06" box="[883,1153,151,175]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="110" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB1FFE3A35CFF71FB5FFF06" box="[883,1153,151,175]" italics="true" pageId="5" pageNumber="110">Rhinella mirandaribeiroi</emphasis>
</taxonomicName>
. N = number of analyzed adult males; data is presented as mean ± standard deviation (minimum–maximum).
</paragraph>
</caption>
<paragraph id="6D44144CFFB1FFE3A08CFF02FAF4F820" pageId="5" pageNumber="110">
<table id="1FFBE6ECFFB10019A08CFF02FAE9F820" box="[163,1335,228,1929]" colsContinueIn="6.[163,1330,202,684]" gridcols="4" gridrows="23" pageId="5" pageNumber="110">
<tr id="D3CB160EFFB10019A08CFF02FAE9FE89" box="[163,1335,228,288]" gridrow="0" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFF02FE5BFE89" box="[163,389,228,288]" gridcol="0" gridrow="0" pageId="5" pageNumber="110">Population-Variables</th>
<th id="901A7F72FFB10019A211FF02FD6AFE89" box="[574,692,228,288]" gridcol="1" gridrow="0" pageId="5" pageNumber="110">
<emphasis id="5F8FC85EFFB1FFE3A26DFF02FD72FE89" bold="true" box="[578,688,228,288]" pageId="5" pageNumber="110">Topotypes Soure PA</emphasis>
</th>
<th id="901A7F72FFB10019A354FF02FC24FE89" box="[891,1018,228,288]" gridcol="2" gridrow="0" pageId="5" pageNumber="110">
<emphasis id="5F8FC85EFFB1FFE3A3A2FF02FC24FE89" bold="true" pageId="5" pageNumber="110">Cerrado populations</emphasis>
</th>
<th id="901A7F72FFB10019A4EFFF02FAE9FE89" box="[1216,1335,228,288]" gridcol="3" gridrow="0" pageId="5" pageNumber="110">
<emphasis id="5F8FC85EFFB1FFE3A4EBFF02FAF0FEB6" bold="true" box="[1220,1330,228,287]" pageId="5" pageNumber="110">
Topotypes
<emphasis id="5F8FC85EFFB1FFE3A4E5FEEEFAF0FEB6" bold="true" box="[1226,1326,263,287]" italics="true" pageId="5" pageNumber="110">B. g. lutzi</emphasis>
</emphasis>
</th>
</tr>
<tr id="D3CB160EFFB10019A08CFED6FAE9FEE1" box="[163,1335,304,328]" gridrow="1" pageId="5" pageNumber="110" rowspan-0="1">
<td id="901A7F72FFB10019A211FED6FD6AFEE1" box="[574,692,304,328]" gridcol="1" gridrow="1" pageId="5" pageNumber="110">N = 23</td>
<td id="901A7F72FFB10019A354FED6FC24FEE1" box="[891,1018,304,328]" gridcol="2" gridrow="1" pageId="5" pageNumber="110">N = 45</td>
<td id="901A7F72FFB10019A4EFFED6FAE9FEE1" box="[1216,1335,304,328]" gridcol="3" gridrow="1" pageId="5" pageNumber="110">N = 2</td>
</tr>
<tr id="D3CB160EFFB10019A08CFEBEFAE9FE3D" box="[163,1335,344,404]" gridrow="2" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFEBEFE5BFE3D" box="[163,389,344,404]" gridcol="0" gridrow="2" pageId="5" pageNumber="110">Snout-vent length</th>
<td id="901A7F72FFB10019A211FEBEFD6AFE3D" box="[574,692,344,404]" gridcol="1" gridrow="2" pageId="5" pageNumber="110">44.4 ± 2.5 (38.3–49.9)</td>
<td id="901A7F72FFB10019A354FEBEFC24FE3D" box="[891,1018,344,404]" gridcol="2" gridrow="2" pageId="5" pageNumber="110">48 ± 3.8 (41.2–59.4)</td>
<td id="901A7F72FFB10019A4EFFEBEFAE9FE3D" box="[1216,1335,344,404]" gridcol="3" gridrow="2" pageId="5" pageNumber="110">43.5 ± 7.5 (43.5–53.3)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFE42FAE9FE48" box="[163,1335,420,481]" gridrow="3" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFE42FE5BFE48" box="[163,389,420,481]" gridcol="0" gridrow="3" pageId="5" pageNumber="110">Thigh length</th>
<td id="901A7F72FFB10019A211FE42FD6AFE48" box="[574,692,420,481]" gridcol="1" gridrow="3" pageId="5" pageNumber="110">15.7 ± 1.1 (13.8–7.9)</td>
<td id="901A7F72FFB10019A354FE42FC24FE48" box="[891,1018,420,481]" gridcol="2" gridrow="3" pageId="5" pageNumber="110">16.8 ± 2 (12.2–20.8)</td>
<td id="901A7F72FFB10019A4EFFE42FAE9FE48" box="[1216,1335,420,481]" gridcol="3" gridrow="3" pageId="5" pageNumber="110">17.1 ± 5.5 (15.8–18.3)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFE17FAE9FD84" box="[163,1335,497,557]" gridrow="4" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFE17FE5BFD84" box="[163,389,497,557]" gridcol="0" gridrow="4" pageId="5" pageNumber="110">Tibia length</th>
<td id="901A7F72FFB10019A211FE17FD6AFD84" box="[574,692,497,557]" gridcol="1" gridrow="4" pageId="5" pageNumber="110">15.1 ± 0.9 (13.8–17)</td>
<td id="901A7F72FFB10019A354FE17FC24FD84" box="[891,1018,497,557]" gridcol="2" gridrow="4" pageId="5" pageNumber="110">16.5 ± 1.5 (13.0–19.9)</td>
<td id="901A7F72FFB10019A4EFFE17FAE9FD84" box="[1216,1335,497,557]" gridcol="3" gridrow="4" pageId="5" pageNumber="110">16.4 ± 5.1 (15.2–17.5)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFDDBFAE9FDD0" box="[163,1335,573,633]" gridrow="5" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFDDBFE5BFDD0" box="[163,389,573,633]" gridcol="0" gridrow="5" pageId="5" pageNumber="110">Tarsus length</th>
<td id="901A7F72FFB10019A211FDDBFD6AFDD0" box="[574,692,573,633]" gridcol="1" gridrow="5" pageId="5" pageNumber="110">9.4 ± 0.7 (8.2–10.8)</td>
<td id="901A7F72FFB10019A354FDDBFC24FDD0" box="[891,1018,573,633]" gridcol="2" gridrow="5" pageId="5" pageNumber="110">10.2 ± 1 (8.09–12.0)</td>
<td id="901A7F72FFB10019A4EFFDDBFAE9FDD0" box="[1216,1335,573,633]" gridcol="3" gridrow="5" pageId="5" pageNumber="110">11.3 ± 4.8 (10.6–12)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFD6FFAE9FD6C" box="[163,1335,649,709]" gridrow="6" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFD6FFE5BFD6C" box="[163,389,649,709]" gridcol="0" gridrow="6" pageId="5" pageNumber="110">Hand length</th>
<td id="901A7F72FFB10019A211FD6FFD6AFD6C" box="[574,692,649,709]" gridcol="1" gridrow="6" pageId="5" pageNumber="110">9.8 ± 0.6 (8.8–11)</td>
<td id="901A7F72FFB10019A354FD6FFC24FD6C" box="[891,1018,649,709]" gridcol="2" gridrow="6" pageId="5" pageNumber="110">10 ± 0.9 (8.44–12.2)</td>
<td id="901A7F72FFB10019A4EFFD6FFAE9FD6C" box="[1216,1335,649,709]" gridcol="3" gridrow="6" pageId="5" pageNumber="110">11 ± 4.7 (10.4–11.6)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFD33FAE9FCBB" box="[163,1335,725,786]" gridrow="7" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFD33FE5BFCBB" box="[163,389,725,786]" gridcol="0" gridrow="7" pageId="5" pageNumber="110">Foot length</th>
<td id="901A7F72FFB10019A211FD33FD6AFCBB" box="[574,692,725,786]" gridcol="1" gridrow="7" pageId="5" pageNumber="110">15.8 ± 1.4 (12.9–18.3)</td>
<td id="901A7F72FFB10019A354FD33FC24FCBB" box="[891,1018,725,786]" gridcol="2" gridrow="7" pageId="5" pageNumber="110">16.5 ± 1.5 (13.4–19.8)</td>
<td id="901A7F72FFB10019A4EFFD33FAE9FCBB" box="[1216,1335,725,786]" gridcol="3" gridrow="7" pageId="5" pageNumber="110">17.8 ± 4.7 (17.2–18.4)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFCC4FAE9FCF7" box="[163,1335,802,862]" gridrow="8" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFCC4FE5BFCF7" box="[163,389,802,862]" gridcol="0" gridrow="8" pageId="5" pageNumber="110">Head height</th>
<td id="901A7F72FFB10019A211FCC4FD6AFCF7" box="[574,692,802,862]" gridcol="1" gridrow="8" pageId="5" pageNumber="110">6.4 ± 0.4 (5.8–7.2)</td>
<td id="901A7F72FFB10019A354FCC4FC24FCF7" box="[891,1018,802,862]" gridcol="2" gridrow="8" pageId="5" pageNumber="110">6.5 ± 0.5 (5.5–8.2)</td>
<td id="901A7F72FFB10019A4EFFCC4FAE9FCF7" box="[1216,1335,802,862]" gridcol="3" gridrow="8" pageId="5" pageNumber="110">6.3 ± 3.9 (5.9–6.7)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFC88FAE9FC03" box="[163,1335,878,938]" gridrow="9" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFC88FE5BFC03" box="[163,389,878,938]" gridcol="0" gridrow="9" pageId="5" pageNumber="110">Head width</th>
<td id="901A7F72FFB10019A211FC88FD6AFC03" box="[574,692,878,938]" gridcol="1" gridrow="9" pageId="5" pageNumber="110">14.3 ± 0.59 (12.8–15.7)</td>
<td id="901A7F72FFB10019A354FC88FC24FC03" box="[891,1018,878,938]" gridcol="2" gridrow="9" pageId="5" pageNumber="110">15.2 ± 0.9 (13.4–17.7)</td>
<td id="901A7F72FFB10019A4EFFC88FAE9FC03" box="[1216,1335,878,938]" gridcol="3" gridrow="9" pageId="5" pageNumber="110">15.4 ± 3.9 (14.5–16.3)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFC5CFAE9FC5F" box="[163,1335,954,1014]" gridrow="10" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFC5CFE5BFC5F" box="[163,389,954,1014]" gridcol="0" gridrow="10" pageId="5" pageNumber="110">Head length</th>
<td id="901A7F72FFB10019A211FC5CFD6AFC5F" box="[574,692,954,1014]" gridcol="1" gridrow="10" pageId="5" pageNumber="110">12.8 ± 0.8 (11.4–14.3)</td>
<td id="901A7F72FFB10019A354FC5CFC24FC5F" box="[891,1018,954,1014]" gridcol="2" gridrow="10" pageId="5" pageNumber="110">13.3 ± 0.9 (11.3–15)</td>
<td id="901A7F72FFB10019A4EFFC5CFAE9FC5F" box="[1216,1335,954,1014]" gridcol="3" gridrow="10" pageId="5" pageNumber="110">13.4 ± 3.1 (12.8–14.1)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFBE0FAE9FBEA" box="[163,1335,1030,1091]" gridrow="11" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFBE0FE5BFBEA" box="[163,389,1030,1091]" gridcol="0" gridrow="11" pageId="5" pageNumber="110">Eye diameter</th>
<td id="901A7F72FFB10019A211FBE0FD6AFBEA" box="[574,692,1030,1091]" gridcol="1" gridrow="11" pageId="5" pageNumber="110">3.6 ± 0.4 (2.9–4.5)</td>
<td id="901A7F72FFB10019A354FBE0FC24FBEA" box="[891,1018,1030,1091]" gridcol="2" gridrow="11" pageId="5" pageNumber="110">4.2 ± 0.5 (3–5.1)</td>
<td id="901A7F72FFB10019A4EFFBE0FAE9FBEA" box="[1216,1335,1030,1091]" gridcol="3" gridrow="11" pageId="5" pageNumber="110">4.2 ± 2.1 (3.7–4.7)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFBB5FAE9FB26" box="[163,1335,1107,1167]" gridrow="12" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFBB5FE5BFB26" box="[163,389,1107,1167]" gridcol="0" gridrow="12" pageId="5" pageNumber="110">Tympanum diameter</th>
<td id="901A7F72FFB10019A211FBB5FD6AFB26" box="[574,692,1107,1167]" gridcol="1" gridrow="12" pageId="5" pageNumber="110">2.3 ± 0.3 (1.6–2.8)</td>
<td id="901A7F72FFB10019A354FBB5FC24FB26" box="[891,1018,1107,1167]" gridcol="2" gridrow="12" pageId="5" pageNumber="110">2.7 ± 0.37 (2–3.4)</td>
<td id="901A7F72FFB10019A4EFFBB5FAE9FB26" box="[1216,1335,1107,1167]" gridcol="3" gridrow="12" pageId="5" pageNumber="110">2.3 ± 2.1 (2.3–2.4)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFB79FAE9FB72" box="[163,1335,1183,1243]" gridrow="13" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFB79FE5BFB72" box="[163,389,1183,1243]" gridcol="0" gridrow="13" pageId="5" pageNumber="110">Tympanum height</th>
<td id="901A7F72FFB10019A211FB79FD6AFB72" box="[574,692,1183,1243]" gridcol="1" gridrow="13" pageId="5" pageNumber="110">2.9 ± 0.4 (2.3–3.7)</td>
<td id="901A7F72FFB10019A354FB79FC24FB72" box="[891,1018,1183,1243]" gridcol="2" gridrow="13" pageId="5" pageNumber="110">3.4 ± 0.3 (2.8–4)</td>
<td id="901A7F72FFB10019A4EFFB79FAE9FB72" box="[1216,1335,1183,1243]" gridcol="3" gridrow="13" pageId="5" pageNumber="110">3 ± 2.2 (3–3.1)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFB0DFAE9FA8E" box="[163,1335,1259,1319]" gridrow="14" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFB0DFE5BFA8E" box="[163,389,1259,1319]" gridcol="0" gridrow="14" pageId="5" pageNumber="110">Snout width</th>
<td id="901A7F72FFB10019A211FB0DFD6AFA8E" box="[574,692,1259,1319]" gridcol="1" gridrow="14" pageId="5" pageNumber="110">5.2 ± 0.4 (4.5–5.7)</td>
<td id="901A7F72FFB10019A354FB0DFC24FA8E" box="[891,1018,1259,1319]" gridcol="2" gridrow="14" pageId="5" pageNumber="110">5.5 ± 0.4 (4.2–6.3)</td>
<td id="901A7F72FFB10019A4EFFB0DFAE9FA8E" box="[1216,1335,1259,1319]" gridcol="3" gridrow="14" pageId="5" pageNumber="110">5.1 ± 2.3 (5–5.3)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFAD1FAE9FADD" box="[163,1335,1335,1396]" gridrow="15" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFAD1FE5BFADD" box="[163,389,1335,1396]" gridcol="0" gridrow="15" pageId="5" pageNumber="110">Parotoid gland width</th>
<td id="901A7F72FFB10019A211FAD1FD6AFADD" box="[574,692,1335,1396]" gridcol="1" gridrow="15" pageId="5" pageNumber="110">7.4 ± 1.2 (4.9–9.8)</td>
<td id="901A7F72FFB10019A354FAD1FC24FADD" box="[891,1018,1335,1396]" gridcol="2" gridrow="15" pageId="5" pageNumber="110">8.2 ± 1.4 (5.2–10.9)</td>
<td id="901A7F72FFB10019A4EFFAD1FAE9FADD" box="[1216,1335,1335,1396]" gridcol="3" gridrow="15" pageId="5" pageNumber="110">43.5 ± 7.5 (43.5–53.3)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFA65FAE9FA69" box="[163,1335,1411,1472]" gridrow="16" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFA65FE5BFA69" box="[163,389,1411,1472]" gridcol="0" gridrow="16" pageId="5" pageNumber="110">Parotoid gland height</th>
<td id="901A7F72FFB10019A211FA65FD6AFA69" box="[574,692,1411,1472]" gridcol="1" gridrow="16" pageId="5" pageNumber="110">5.6 ± 0.4 (4.83–6.59)</td>
<td id="901A7F72FFB10019A354FA65FC24FA69" box="[891,1018,1411,1472]" gridcol="2" gridrow="16" pageId="5" pageNumber="110">6.3 ± 0.9 (4.7–9.4)</td>
<td id="901A7F72FFB10019A4EFFA65FAE9FA69" box="[1216,1335,1411,1472]" gridcol="3" gridrow="16" pageId="5" pageNumber="110">5.7 ± 2 (5.4–6.0)</td>
</tr>
<tr id="D3CB160EFFB10019A08CFA36FAE9F9A5" box="[163,1335,1488,1548]" gridrow="17" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CFA36FE5BF9A5" box="[163,389,1488,1548]" gridcol="0" gridrow="17" pageId="5" pageNumber="110">Internarial distance</th>
<td id="901A7F72FFB10019A211FA36FD6AF9A5" box="[574,692,1488,1548]" gridcol="1" gridrow="17" pageId="5" pageNumber="110">2.43 ± 0.2 (1.9–2.7)</td>
<td id="901A7F72FFB10019A354FA36FC24F9A5" box="[891,1018,1488,1548]" gridcol="2" gridrow="17" pageId="5" pageNumber="110">2.59 ± 0.19 (2.2–3.1)</td>
<td id="901A7F72FFB10019A4EFFA36FAE9F9A5" box="[1216,1335,1488,1548]" gridcol="3" gridrow="17" pageId="5" pageNumber="110">2.45 ± 1.9 (2.1–2.8)</td>
</tr>
<tr id="D3CB160EFFB10019A08CF9FAFAE9F9F1" box="[163,1335,1564,1624]" gridrow="18" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CF9FAFE5BF9F1" box="[163,389,1564,1624]" gridcol="0" gridrow="18" pageId="5" pageNumber="110">Eye-snout distance</th>
<td id="901A7F72FFB10019A211F9FAFD6AF9F1" box="[574,692,1564,1624]" gridcol="1" gridrow="18" pageId="5" pageNumber="110">2.32 ± 0.4 (1.6–3)</td>
<td id="901A7F72FFB10019A354F9FAFC24F9F1" box="[891,1018,1564,1624]" gridcol="2" gridrow="18" pageId="5" pageNumber="110">2.85 ± 0.36 (2–3.6)</td>
<td id="901A7F72FFB10019A4EFF9FAFAE9F9F1" box="[1216,1335,1564,1624]" gridcol="3" gridrow="18" pageId="5" pageNumber="110">3.2 ± 2 (2.8–3.6)</td>
</tr>
<tr id="D3CB160EFFB10019A08CF98EFAE9F90D" box="[163,1335,1640,1700]" gridrow="19" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CF98EFE5BF90D" box="[163,389,1640,1700]" gridcol="0" gridrow="19" pageId="5" pageNumber="110">Eye–nostril distance</th>
<td id="901A7F72FFB10019A211F98EFD6AF90D" box="[574,692,1640,1700]" gridcol="1" gridrow="19" pageId="5" pageNumber="110">4.28 ± 0.5 (3.5–5.3)</td>
<td id="901A7F72FFB10019A354F98EFC24F90D" box="[891,1018,1640,1700]" gridcol="2" gridrow="19" pageId="5" pageNumber="110">5.02 ± 0.57 (3.9–6)</td>
<td id="901A7F72FFB10019A4EFF98EFAE9F90D" box="[1216,1335,1640,1700]" gridcol="3" gridrow="19" pageId="5" pageNumber="110">5.4 ± 2.1 (4.9–5.9)</td>
</tr>
<tr id="D3CB160EFFB10019A08CF952FAE9F958" box="[163,1335,1716,1777]" gridrow="20" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CF952FE5BF958" box="[163,389,1716,1777]" gridcol="0" gridrow="20" pageId="5" pageNumber="110">Interorbital distance</th>
<td id="901A7F72FFB10019A211F952FD6AF958" box="[574,692,1716,1777]" gridcol="1" gridrow="20" pageId="5" pageNumber="110">4.96 ± 0.4 (4.1–5.8)</td>
<td id="901A7F72FFB10019A354F952FC24F958" box="[891,1018,1716,1777]" gridcol="2" gridrow="20" pageId="5" pageNumber="110">5.65 ± 0.47 (4.9–6.6)</td>
<td id="901A7F72FFB10019A4EFF952FAE9F958" box="[1216,1335,1716,1777]" gridcol="3" gridrow="20" pageId="5" pageNumber="110">5.35 ± 2 (5.1–5.6)</td>
</tr>
<tr id="D3CB160EFFB10019A08CF8E7FAE9F894" box="[163,1335,1793,1853]" gridrow="21" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CF8E7FE5BF894" box="[163,389,1793,1853]" gridcol="0" gridrow="21" pageId="5" pageNumber="110">Interorbital distance II</th>
<td id="901A7F72FFB10019A211F8E7FD6AF894" box="[574,692,1793,1853]" gridcol="1" gridrow="21" pageId="5" pageNumber="110">4.81 ± 0.4 (4–5.6)</td>
<td id="901A7F72FFB10019A354F8E7FC24F894" box="[891,1018,1793,1853]" gridcol="2" gridrow="21" pageId="5" pageNumber="110">4.94 ± 0.33 (4.2–5.8)</td>
<td id="901A7F72FFB10019A4EFF8E7FAE9F894" box="[1216,1335,1793,1853]" gridcol="3" gridrow="21" pageId="5" pageNumber="110">4.55 ± 1.8 (4.4–4.7)</td>
</tr>
<tr id="D3CB160EFFB10019A08CF8ABFAE9F820" box="[163,1335,1869,1929]" gridrow="22" pageId="5" pageNumber="110">
<th id="901A7F72FFB10019A08CF8ABFE5BF820" box="[163,389,1869,1929]" gridcol="0" gridrow="22" pageId="5" pageNumber="110">Eyelid width</th>
<td id="901A7F72FFB10019A211F8ABFD6AF820" box="[574,692,1869,1929]" gridcol="1" gridrow="22" pageId="5" pageNumber="110">3.69 ± 0.2 (3.3–4.1)</td>
<td id="901A7F72FFB10019A354F8ABFC24F820" box="[891,1018,1869,1929]" gridcol="2" gridrow="22" pageId="5" pageNumber="110">4.38 ± 0.35 (3.5–5)</td>
<td id="901A7F72FFB10019A4EFF8ABFAE9F820" box="[1216,1335,1869,1929]" gridcol="3" gridrow="22" pageId="5" pageNumber="110">4.25 ± 1.5 (3.7–4.8)</td>
</tr>
</table>
</paragraph>
<footnote id="0EE00842FFB1FFE3A475F87CFA4FF81B" box="[1114,1425,1946,1970]" pageId="5" pageNumber="110">
<paragraph id="6D44144CFFB1FFE3A475F87CFA4FF81B" blockId="5.[1114,1425,1946,1970]" box="[1114,1425,1946,1970]" pageId="5" pageNumber="110">
<emphasis id="5F8FC85EFFB1FFE3A475F87CFA4FF81B" box="[1114,1425,1946,1970]" italics="true" pageId="5" pageNumber="110">......continued on the next page</emphasis>
</paragraph>
</footnote>
<caption id="398444C4FFB2FFE0A08CFF44FE4FFF13" box="[163,401,162,186]" isContinuationCaption="true" pageId="6" pageNumber="111" startId="6.[163,247,162,186]" targetType="table">
<paragraph id="6D44144CFFB2FFE0A08CFF44FE4FFF13" blockId="6.[163,401,162,186]" box="[163,401,162,186]" pageId="6" pageNumber="111">
<emphasis id="5F8FC85EFFB2FFE0A08CFF44FECEFF13" bold="true" box="[163,272,162,186]" pageId="6" pageNumber="111">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="6D44144CFFB2FFE0A08CFF2CFAF4FD05" pageId="6" pageNumber="111">
<table id="1FFBE6ECFFB20019A08CFF2CFAECFD05" box="[163,1330,202,684]" colsContinueFrom="5.[163,1335,228,1929]" gridcols="4" gridrows="12" pageId="6" pageNumber="111">
<tr id="D3CB160EFFB20019A08CFF2CFAECFEAF" box="[163,1330,202,262]" gridrow="0" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFF2CFE1EFEAF" box="[163,448,202,262]" gridcol="0" gridrow="0" pageId="6" pageNumber="111">Population-Variables</th>
<th id="901A7F72FFB20019A26DFF2CFD6EFEAF" box="[578,688,202,262]" gridcol="1" gridrow="0" pageId="6" pageNumber="111">
<emphasis id="5F8FC85EFFB2FFE0A26DFF2CFD6EFF4B" bold="true" box="[578,688,202,226]" pageId="6" pageNumber="111">Topotypes</emphasis>
<emphasis id="5F8FC85EFFB2FFE0A268FF08FD72FEAF" bold="true" box="[583,684,238,262]" pageId="6" pageNumber="111">Soure PA</emphasis>
</th>
<th id="901A7F72FFB20019A354FF2CFC24FEAF" box="[891,1018,202,262]" gridcol="2" gridrow="0" pageId="6" pageNumber="111">
<emphasis id="5F8FC85EFFB2FFE0A3A2FF2CFC36FF4B" bold="true" box="[909,1000,202,226]" pageId="6" pageNumber="111">Cerrado</emphasis>
<emphasis id="5F8FC85EFFB2FFE0A354FF08FC24FEAF" bold="true" box="[891,1018,238,262]" pageId="6" pageNumber="111">populations</emphasis>
</th>
<th id="901A7F72FFB20019A4EBFF2CFAECFEAF" box="[1220,1330,202,262]" gridcol="3" gridrow="0" pageId="6" pageNumber="111">
<emphasis id="5F8FC85EFFB2FFE0A4EBFF2CFAECFF4B" bold="true" box="[1220,1330,202,226]" pageId="6" pageNumber="111">Topotypes</emphasis>
<emphasis id="5F8FC85EFFB2FFE0A4E5FF08FAF0FEAF" bold="true" box="[1226,1326,238,262]" italics="true" pageId="6" pageNumber="111">B. g. lutzi</emphasis>
</th>
</tr>
<tr id="D3CB160EFFB20019A08CFEF0FAECFE86" box="[163,1330,278,303]" gridrow="1" pageId="6" pageNumber="111" rowspan-0="1">
<td id="901A7F72FFB20019A26DFEF0FD6EFE86" box="[578,688,278,303]" gridcol="1" gridrow="1" pageId="6" pageNumber="111">N = 23</td>
<td id="901A7F72FFB20019A354FEF0FC24FE86" box="[891,1018,278,303]" gridcol="2" gridrow="1" pageId="6" pageNumber="111">N = 45</td>
<td id="901A7F72FFB20019A4EBFEF0FAECFE86" box="[1220,1330,278,303]" gridcol="3" gridrow="1" pageId="6" pageNumber="111">N = 2</td>
</tr>
<tr id="D3CB160EFFB20019A08CFED9FAECFEFE" box="[163,1330,319,343]" gridrow="2" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFED9FE1EFEFE" box="[163,448,319,343]" gridcol="0" gridrow="2" pageId="6" pageNumber="111">Supratympanic crest length</th>
<td id="901A7F72FFB20019A26DFED9FD6EFEFE" box="[578,688,319,343]" gridcol="1" gridrow="2" pageId="6" pageNumber="111">1.93 ± 0.4</td>
<td id="901A7F72FFB20019A354FED9FC24FEFE" box="[891,1018,319,343]" gridcol="2" gridrow="2" pageId="6" pageNumber="111">1.98 ± 0.55</td>
<td id="901A7F72FFB20019A4EBFED9FAECFEFE" box="[1220,1330,319,343]" gridcol="3" gridrow="2" pageId="6" pageNumber="111">2 ± 1.1</td>
</tr>
<tr id="D3CB160EFFB20019A08CFE85FAECFED2" box="[163,1330,355,379]" gridrow="3" pageId="6" pageNumber="111" rowspan-0="1">
<td id="901A7F72FFB20019A26DFE85FD6EFED2" box="[578,688,355,379]" gridcol="1" gridrow="3" pageId="6" pageNumber="111">(1–2.5)</td>
<td id="901A7F72FFB20019A354FE85FC24FED2" box="[891,1018,355,379]" gridcol="2" gridrow="3" pageId="6" pageNumber="111">(0.8–3.2)</td>
<td id="901A7F72FFB20019A4EBFE85FAECFED2" box="[1220,1330,355,379]" gridcol="3" gridrow="3" pageId="6" pageNumber="111">(1.5–2.5)</td>
</tr>
<tr id="D3CB160EFFB20019A08CFE6DFAECFE0D" box="[163,1330,395,420]" gridrow="4" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFE6DFE1EFE0D" box="[163,448,395,420]" gridcol="0" gridrow="4" pageId="6" pageNumber="111">Postorbital space</th>
<td id="901A7F72FFB20019A26DFE6DFD6EFE0D" box="[578,688,395,420]" gridcol="1" gridrow="4" pageId="6" pageNumber="111">3.06 ± 0.3</td>
<td id="901A7F72FFB20019A354FE6DFC24FE0D" box="[891,1018,395,420]" gridcol="2" gridrow="4" pageId="6" pageNumber="111">3.46 ± 0.66</td>
<td id="901A7F72FFB20019A4EBFE6DFAECFE0D" box="[1220,1330,395,420]" gridcol="3" gridrow="4" pageId="6" pageNumber="111">2.85 ± 1.1</td>
</tr>
<tr id="D3CB160EFFB20019A08CFE49FAECFE61" box="[163,1330,431,456]" gridrow="5" pageId="6" pageNumber="111" rowspan-0="1">
<td id="901A7F72FFB20019A26DFE49FD6EFE61" box="[578,688,431,456]" gridcol="1" gridrow="5" pageId="6" pageNumber="111">(2.3–3.6)</td>
<td id="901A7F72FFB20019A354FE49FC24FE61" box="[891,1018,431,456]" gridcol="2" gridrow="5" pageId="6" pageNumber="111">(0.3–4.3)</td>
<td id="901A7F72FFB20019A4EBFE49FAECFE61" box="[1220,1330,431,456]" gridcol="3" gridrow="5" pageId="6" pageNumber="111">(2.6–3.1)</td>
</tr>
<tr id="D3CB160EFFB20019A08CFE3EFAECFE59" box="[163,1330,472,496]" gridrow="6" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFE3EFE1EFE59" box="[163,448,472,496]" gridcol="0" gridrow="6" pageId="6" pageNumber="111">Distance from tympanum to</th>
<td id="901A7F72FFB20019A26DFE3EFD6EFE59" box="[578,688,472,496]" gridcol="1" gridrow="6" pageId="6" pageNumber="111">0.27 ± 0.1</td>
<td id="901A7F72FFB20019A354FE3EFC24FE59" box="[891,1018,472,496]" gridcol="2" gridrow="6" pageId="6" pageNumber="111">0.26 ± 0.12</td>
<td id="901A7F72FFB20019A4EBFE3EFAECFE59" box="[1220,1330,472,496]" gridcol="3" gridrow="6" pageId="6" pageNumber="111">0.3 ± 0.3</td>
</tr>
<tr id="D3CB160EFFB20019A08CFE1AFAECFDBD" box="[163,1330,508,532]" gridrow="7" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFE1AFE1EFDBD" box="[163,448,508,532]" gridcol="0" gridrow="7" pageId="6" pageNumber="111">postorbital crest</th>
<td id="901A7F72FFB20019A26DFE1AFD6EFDBD" box="[578,688,508,532]" gridcol="1" gridrow="7" pageId="6" pageNumber="111">(0.2–0.4)</td>
<td id="901A7F72FFB20019A354FE1AFC24FDBD" box="[891,1018,508,532]" gridcol="2" gridrow="7" pageId="6" pageNumber="111">(0.1–0.5)</td>
<td id="901A7F72FFB20019A4EBFE1AFAECFDBD" box="[1220,1330,508,532]" gridcol="3" gridrow="7" pageId="6" pageNumber="111">(0.1–0.5)</td>
</tr>
<tr id="D3CB160EFFB20019A08CFDC2FAECFD94" box="[163,1330,548,573]" gridrow="8" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFDC2FE1EFD94" box="[163,448,548,573]" gridcol="0" gridrow="8" pageId="6" pageNumber="111">Distance from tympanum to</th>
<td id="901A7F72FFB20019A26DFDC2FD6EFD94" box="[578,688,548,573]" gridcol="1" gridrow="8" pageId="6" pageNumber="111">0.97 ± 0.2</td>
<td id="901A7F72FFB20019A354FDC2FC24FD94" box="[891,1018,548,573]" gridcol="2" gridrow="8" pageId="6" pageNumber="111">0.8 ± 0.18</td>
<td id="901A7F72FFB20019A4EBFDC2FAECFD94" box="[1220,1330,548,573]" gridcol="3" gridrow="8" pageId="6" pageNumber="111">0.55 ± 0.2</td>
</tr>
<tr id="D3CB160EFFB20019A08CFDAEFAECFDC9" box="[163,1330,584,608]" gridrow="9" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFDAEFE1EFDC9" box="[163,448,584,608]" gridcol="0" gridrow="9" pageId="6" pageNumber="111">supra tympanic crest</th>
<td id="901A7F72FFB20019A26DFDAEFD6EFDC9" box="[578,688,584,608]" gridcol="1" gridrow="9" pageId="6" pageNumber="111">(0.5–1.4)</td>
<td id="901A7F72FFB20019A354FDAEFC24FDC9" box="[891,1018,584,608]" gridcol="2" gridrow="9" pageId="6" pageNumber="111">(0.4–1.2)</td>
<td id="901A7F72FFB20019A4EBFDAEFAECFDC9" box="[1220,1330,584,608]" gridcol="3" gridrow="9" pageId="6" pageNumber="111">(0.3–0.8)</td>
</tr>
<tr id="D3CB160EFFB20019A08CFD96FAECFD20" box="[163,1330,624,649]" gridrow="10" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFD96FE1EFD20" box="[163,448,624,649]" gridcol="0" gridrow="10" pageId="6" pageNumber="111">Distance from tympanum to</th>
<td id="901A7F72FFB20019A26DFD96FD6EFD20" box="[578,688,624,649]" gridcol="1" gridrow="10" pageId="6" pageNumber="111">0.71 ± 0.3</td>
<td id="901A7F72FFB20019A354FD96FC24FD20" box="[891,1018,624,649]" gridcol="2" gridrow="10" pageId="6" pageNumber="111">0.86 ± 0.34</td>
<td id="901A7F72FFB20019A4EBFD96FAECFD20" box="[1220,1330,624,649]" gridcol="3" gridrow="10" pageId="6" pageNumber="111">0.6 ± 0.1</td>
</tr>
<tr id="D3CB160EFFB20019A08CFD72FAECFD05" box="[163,1330,660,684]" gridrow="11" pageId="6" pageNumber="111">
<th id="901A7F72FFB20019A08CFD72FE1EFD05" box="[163,448,660,684]" gridcol="0" gridrow="11" pageId="6" pageNumber="111">parotoid gland</th>
<td id="901A7F72FFB20019A26DFD72FD6EFD05" box="[578,688,660,684]" gridcol="1" gridrow="11" pageId="6" pageNumber="111">(0.2–1.25)</td>
<td id="901A7F72FFB20019A354FD72FC24FD05" box="[891,1018,660,684]" gridcol="2" gridrow="11" pageId="6" pageNumber="111">(0.3–1.5)</td>
<td id="901A7F72FFB20019A4EBFD72FAECFD05" box="[1220,1330,660,684]" gridcol="3" gridrow="11" pageId="6" pageNumber="111">(0.5–0.7)</td>
</tr>
</table>
</paragraph>
<paragraph id="6D44144CFFB2FFE0A0B8FCE6FED0FCB3" blockId="6.[151,270,768,794]" box="[151,270,768,794]" pageId="6" pageNumber="111">
<heading id="360CA320FFB2FFE0A0B8FCE6FED0FCB3" bold="true" box="[151,270,768,794]" fontSize="11" level="1" pageId="6" pageNumber="111" reason="1">
<emphasis id="5F8FC85EFFB2FFE0A0B8FCE6FED0FCB3" bold="true" box="[151,270,768,794]" pageId="6" pageNumber="111">Acoustics</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFB2FFE0A0B8FCAEFCBFFC5B" blockId="6.[151,1436,840,1010]" pageId="6" pageNumber="111">We found conserved parameters of call features among samples. All evaluated specimens have advertisement calls consisting of regular series of pulsed notes (trills) that can last up to 11 seconds. The call has ascending amplitude modulation, with the initial notes generally having low amplitude and then quickly reaching the maximum, which is maintained up the end. The notes consist of three to five concatenated pulses (without intervals of silence). No acoustic feature was different among the evaluated populations.</paragraph>
<paragraph id="6D44144CFFB2FFE0A0B8FBA2FC37FBF4" blockId="6.[151,1001,1091,1118]" box="[151,1001,1091,1118]" pageId="6" pageNumber="111">
<heading id="360CA320FFB2FFE0A0B8FBA2FC37FBF4" bold="true" box="[151,1001,1091,1118]" fontSize="11" level="1" pageId="6" pageNumber="111" reason="1">
<emphasis id="5F8FC85EFFB2FFE0A0B8FBA2FC37FBF4" bold="true" box="[151,1001,1091,1118]" pageId="6" pageNumber="111">
Topotypical population of
<taxonomicName id="AAFB6FCFFFB2FFE0A1CFFBA2FD1CFBF7" box="[480,706,1091,1118]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFB2FFE0A1CFFBA2FD1CFBF7" bold="true" box="[480,706,1091,1118]" italics="true" pageId="6" pageNumber="111">R. mirandaribeiroi</emphasis>
</taxonomicName>
(Soure, Marajó Island)
</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFB2FFE0A0B8FB6AFE69FA0B" blockId="6.[151,1437,1164,1442]" pageId="6" pageNumber="111">
In the
<typeStatus id="B240AAEEFFB2FFE0A0CEFB6AFECFFB0F" box="[225,273,1164,1190]" pageId="6" pageNumber="111">type</typeStatus>
locality, males vocalize after rains at night in shallow puddles formed along dirt roads on the outskirts of the city. The immediate surroundings of these vocalization sites were open anthropogenic environments with pasture or cultivated areas. Syntopic anurans include
<emphasis id="5F8FC85EFFB2FFE0A328FB32FD48FABB" italics="true" pageId="6" pageNumber="111">
<taxonomicName id="AAFB6FCFFFB2FFE0A328FB32FC26FB47" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[775,1016,1236,1262]" class="Amphibia" family="Leptodactylidae" genus="Leptodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="fuscus">Leptodactylus fuscus</taxonomicName>
,
<taxonomicName id="AAFB6FCFFFB2FFE0A428FB32FB0CFB47" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1031,1234,1236,1262]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="x-signatus">Scinax x-signatus</taxonomicName>
,
<taxonomicName id="AAFB6FCFFFB2FFE0A4CEFB33FA46FB47" baseAuthorityName="Cope" baseAuthorityYear="1862" box="[1249,1432,1237,1262]" class="Amphibia" family="Hylidae" genus="Boana" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="raniceps">Boana raniceps</taxonomicName>
,
<taxonomicName id="AAFB6FCFFFB2FFE0A0B8FB1EFDCBFABB" authority=", Physalaemus" authorityName="Physalaemus" box="[151,533,1272,1298]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="fuscomarginatus">S. fuscomarginatus, Physalaemus</taxonomicName>
ephippifer,
</emphasis>
and
<taxonomicName id="AAFB6FCFFFB2FFE0A2FDFB1EFC32FABB" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[722,1004,1272,1298]" class="Amphibia" family="Leptodactylidae" genus="Adenomera" kingdom="Animalia" order="Anura" pageId="6" pageNumber="111" phylum="Chordata" rank="species" species="hylaedactyla">
<emphasis id="5F8FC85EFFB2FFE0A2FDFB1EFC32FABB" box="[722,1004,1272,1298]" italics="true" pageId="6" pageNumber="111">Adenomera hylaedactyla</emphasis>
</taxonomicName>
. The advertisement call consists of a sequence of approximately 126 notes and lasts around 4 seconds, emitted at a rate of 8 calls per minute with 33 notes per second. The calls reach maximum amplitude rapidly and remain constant until the end (
<figureCitation id="F5C008C9FFB2FFE0A4D9FAA6FA84FAF3" box="[1270,1370,1344,1370]" captionStart="FIGURE 3" captionStartId="8.[151,250,1100,1124]" captionTargetBox="[157,1430,188,1069]" captionTargetId="figure-24@8.[151,1436,181,1076]" captionTargetPageId="8" captionText="FIGURE 3. Audio spectrogram and corresponding waveform of the advertisement call of a topotype Rhinella mirandaribeiroi (Soure, Marajó Island, Pará state). Voucher AAG-UFU 6542. (A) Complete advertisement call. (B) Detail of seven notes from the middle portion of the call. Note the 4-pulsed structure, the first and last being of lower amplitude. File = Rhinella_ mirandaribeiroiSourePA1aAAGm661MK2. Air temperature 28°C and water temperature 28°C." figureDoi="http://doi.org/10.5281/zenodo.14896124" httpUri="https://zenodo.org/record/14896124/files/figure.png" pageId="6" pageNumber="111">Figure 3</figureCitation>
). The notes have a duration of about 20 ms and contain 3−4 pulses per note. The dominant frequency is around 2800 Hz. Further details in
<tableCitation id="207921F7FFB2FFE0A172FA6EFE6DFA0B" box="[349,435,1416,1442]" captionStart="TABLE 2" captionStartId="6.[152,244,1489,1513]" captionTargetPageId="6" captionText="TABLE 2. Advertisement call variables for three populations of Rhinella mirandaribeiroi. N = number of individuals and analyzed calls; data presented as mean ± standard deviation (minimum–maximum)." httpUri="http://table.plazi.org/id/398444C4FFB2FFE0A0B7FA37FBCAF9A7" pageId="6" pageNumber="111" tableUuid="398444C4FFB2FFE0A0B7FA37FBCAF9A7">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="25768507799AA12AB8158588F1E6C5F4">The identity of Rhinella mirandaribeiroi (Gallardo, 1965) (Anura: Bufonidae) and its diagnosis in relation to Rhinella granulosa (Spix, 1824)</mods:title>
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<mods:identifier id="A143E9530AE897DAAB2EA6BCA160DFDB" type="ZooBank">AC95B339-E90D-4898-8ECB-4F590CF25F55</mods:identifier>
</mods:mods>
<treatment id="E552A55AFFBCFFEDA0B8F9F9FE16F99F" LSID="urn:lsid:plazi:treatment:E552A55AFFBCFFEDA0B8F9F9FE16F99F" httpUri="http://treatment.plazi.org/id/E552A55AFFBCFFEDA0B8F9F9FE16F99F" lastPageId="11" lastPageNumber="116" pageId="8" pageNumber="113">
<subSubSection id="25E147C7FFBCFFEEA0B8F9F9FD98F990" box="[151,582,1566,1593]" pageId="8" pageNumber="113" type="nomenclature">
<paragraph id="6D44144CFFBCFFEEA0B8F9F9FD98F990" blockId="8.[151,582,1566,1593]" box="[151,582,1566,1593]" pageId="8" pageNumber="113">
<heading id="360CA320FFBCFFEEA0B8F9F9FD98F990" bold="true" box="[151,582,1566,1593]" fontSize="11" level="1" pageId="8" pageNumber="113" reason="1">
<emphasis id="5F8FC85EFFBCFFEEA0B8F9F9FD98F990" bold="true" box="[151,582,1566,1593]" pageId="8" pageNumber="113">
<taxonomicName id="AAFB6FCFFFBCFFEEA0B8F9F9FEA7F990" ID-CoL="4S7P3" box="[151,377,1566,1593]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="113" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBCFFEEA0B8F9F9FEA7F990" bold="true" box="[151,377,1566,1593]" italics="true" pageId="8" pageNumber="113">R. mirandaribeiroi</emphasis>
</taxonomicName>
vs.
<taxonomicName id="AAFB6FCFFFBCFFEEA185F9F9FD98F990" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[426,582,1566,1593]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="113" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBCFFEEA185F9F9FD98F990" bold="true" box="[426,582,1566,1593]" italics="true" pageId="8" pageNumber="113">R. granulosa</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25E147C7FFBCFFEDA0B8F980FE16F99F" lastPageId="11" lastPageNumber="116" pageId="8" pageNumber="113" type="description">
<paragraph id="6D44144CFFBCFFEEA0B8F980FDEAF929" blockId="8.[151,564,1638,1664]" box="[151,564,1638,1664]" pageId="8" pageNumber="113">
<heading id="360CA320FFBCFFEEA0B8F980FDEAF929" bold="true" box="[151,564,1638,1664]" fontSize="11" level="1" pageId="8" pageNumber="113" reason="1">
<emphasis id="5F8FC85EFFBCFFEEA0B8F980FDEAF929" bold="true" box="[151,564,1638,1664]" pageId="8" pageNumber="113">Morphometrics and Morphology</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFBCFFEEA0B8F948FB97F8D5" blockId="8.[151,1437,1710,1917]" pageId="8" pageNumber="113">
Both species share the following morphological features: 1) granular or continuous cranial crests; 2) low supraorbital crest; 3) infraorbital extending usually beyond the postorbital crest; 4) typically reduced and discontinuous maxillary crest; 5) absent parietal crest; 6) visible loreal region in dorsal view; and 7) light cream or yellowish ventral coloration with or without pigmentation. However, a clear dorsal vertebral line is present in 95% of the individuals from the Cerrado (
<taxonomicName id="AAFB6FCFFFBCFFEEA12CF8D9FE09F8F1" box="[259,471,1854,1880]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="113" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBCFFEEA12CF8D9FE09F8F1" box="[259,471,1854,1880]" italics="true" pageId="8" pageNumber="113">R. mirandaribeiroi</emphasis>
</taxonomicName>
) and only in 5% from the Caatinga and Mata Atlântica (
<taxonomicName id="AAFB6FCFFFBCFFEEA44EF8D9FB2AF8F1" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1121,1268,1854,1880]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="113" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBCFFEEA44EF8D9FB2AF8F1" box="[1121,1268,1854,1880]" italics="true" pageId="8" pageNumber="113">R. granulosa</emphasis>
</taxonomicName>
) (
<figureCitation id="F5C008C9FFBCFFEEA522F8D8FAB0F8F1" box="[1293,1390,1854,1880]" captionStart="FIGURE 1" captionStartId="3.[152,255,1467,1491]" captionTargetBox="[161,1426,191,1433]" captionTargetId="figure-17@3.[151,1436,181,1444]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal and lateral view of the head of adult males of. R. mirandaribeiroi and R. granulosa: (A) Soure, Ilha de Marajó, Pará state (topotype; AAG-UFU 6564). (B) Pirapora, Minas Gerais state (topotype of B. g. lutzi; AAG-UFU 7133). R. granulosa: (C) Salinas, Minas Gerais state (CFBH 47633). Note the absence of distinctive elements between populations and species in supraorbital crest (Sc), infraorbital crest (Ic) (extending or not beyond the postorbital crest), tympanum near the postorbital crest (Tp), and the presence of a clear longitudinal vertebral line (Lv)." figureDoi="http://doi.org/10.5281/zenodo.14896120" httpUri="https://zenodo.org/record/14896120/files/figure.png" pageId="8" pageNumber="113">Figure 1</figureCitation>
,
<figureCitation id="F5C008C9FFBCFFEEA551F8D8FA51F8F1" box="[1406,1423,1854,1880]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[295,1292,188,1878]" captionTargetId="figure-24@4.[288,1299,181,1885]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral view of adult males of R. mirandaribeiroi and R. granulosa. (A) R. mirandaribeiroi, Soure, Ilha de Marajó, Pará state (topotypes; AAG-UFU 6542). (B) (topotypes of B. g. lutzi; AAG-UFU 7134 Pirapora, Minas de Gerais state). (C) R. granulosa: Salinas, Minas Gerais state (CFBH 47633). Note the lack of distinctive elements between populations and species." figureDoi="http://doi.org/10.5281/zenodo.14896122" httpUri="https://zenodo.org/record/14896122/files/figure.png" pageId="8" pageNumber="113">2</figureCitation>
). Overall, the morphometric variables were similar between the two species (
<tableCitation id="207921F7FFBCFFEEA3CBF884FBE5F8D5" box="[996,1083,1890,1916]" captionStart="TABLE 3" captionStartId="9.[152,244,1309,1333]" captionTargetPageId="9" captionText="TABLE 3. Values of morphometric variables for the species R. mirandaribeiroi and R. granulosa. N = number of analyzed adult males; data presented as mean ± standard deviation (minimum–maximum)." httpUri="http://table.plazi.org/id/398444C4FFBDFFEFA0B7FAFBFC26FAF3" pageId="8" pageNumber="113" tableUuid="398444C4FFBDFFEFA0B7FAFBFC26FAF3">Table 3</tableCitation>
).
</paragraph>
<caption id="398444C4FFBDFFEFA0B8FBA3FBCCFB63" ID-DOI="http://doi.org/10.5281/zenodo.14896126" ID-Zenodo-Dep="14896126" httpUri="https://zenodo.org/record/14896126/files/figure.png" pageId="9" pageNumber="114" startId="9.[151,250,1093,1117]" targetBox="[158,1429,188,1063]" targetPageId="9" targetType="figure">
<paragraph id="6D44144CFFBDFFEFA0B8FBA3FBCCFB63" blockId="9.[151,1436,1093,1226]" pageId="9" pageNumber="114">
<emphasis id="5F8FC85EFFBDFFEFA0B8FBA3FECEFBF7" bold="true" box="[151,272,1093,1118]" pageId="9" pageNumber="114">FIGURE 4</emphasis>
. Audio spectrogram and corresponding waveform of the advertisement call of a topotype
<emphasis id="5F8FC85EFFBDFFEFA4EFFBA0FAF9FBF7" box="[1216,1319,1094,1118]" italics="true" pageId="9" pageNumber="114">B. g. lutzi</emphasis>
(=
<taxonomicName id="AAFB6FCFFFBDFFEFA569FBA0FEE7FB2B" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBDFFEFA569FBA0FEE7FB2B" italics="true" pageId="9" pageNumber="114">Rhinella mirandaribeiroi</emphasis>
</taxonomicName>
) from Pirapora. The individual was collected. (A) Complete advertisement call. (B) Seven notes from the middle portion of the advertisement call. Note the 4-pulsed structure, the first and last being of lower amplitude. File =
<taxonomicName id="AAFB6FCFFFBDFFEFA515FB68FA4EFB0F" authorityName="Fitzinger" authorityYear="1826" box="[1338,1424,1166,1190]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="genus">Rhinella</taxonomicName>
_ lutziPiraporaMG2aAAGm661MK2. Air temperature 25°C and water temperature 26°C.
</paragraph>
</caption>
<caption id="398444C4FFBDFFEFA0B7FAFBFC26FAF3" ID-Table-UUID="398444C4FFBDFFEFA0B7FAFBFC26FAF3" httpUri="http://table.plazi.org/id/398444C4FFBDFFEFA0B7FAFBFC26FAF3" pageId="9" pageNumber="114" startId="9.[152,244,1309,1333]" targetBox="[163,1290,1385,1983]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="6D44144CFFBDFFEFA0B7FAFBFC26FAF3" blockId="9.[151,1436,1309,1370]" pageId="9" pageNumber="114">
<emphasis id="5F8FC85EFFBDFFEFA0B7FAFBFED3FA9F" bold="true" box="[152,269,1309,1334]" pageId="9" pageNumber="114">TABLE 3.</emphasis>
Values of morphometric variables for the species
<emphasis id="5F8FC85EFFBDFFEFA33BFAF8FB4BFA9F" box="[788,1173,1310,1334]" italics="true" pageId="9" pageNumber="114">
<taxonomicName id="AAFB6FCFFFBDFFEFA33BFAF8FC05FA9F" box="[788,987,1310,1334]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="species" species="mirandaribeiroi">R. mirandaribeiroi</taxonomicName>
and
<taxonomicName id="AAFB6FCFFFBDFFEFA422FAF8FB4BFA9F" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1037,1173,1310,1334]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="species" species="granulosa">R. granulosa</taxonomicName>
</emphasis>
. N = number of analyzed adult males; data presented as mean ± standard deviation (minimum–maximum).
</paragraph>
</caption>
<paragraph id="6D44144CFFBDFFEFA08CFA8CFB2AF816" pageId="9" pageNumber="114">
<table id="1FFBE6ECFFBD0019A08CFA8FFAD4F816" box="[163,1290,1385,1983]" colsContinueIn="10.[163,1290,201,1715]" gridcols="3" gridrows="9" pageId="9" pageNumber="114">
<tr id="D3CB160EFFBD0019A08CFA8FFAD4FA2B" box="[163,1290,1385,1410]" gridrow="0" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CFA8FFE83FA2B" box="[163,349,1385,1410]" gridcol="0" gridrow="0" pageId="9" pageNumber="114">Species -Variables</th>
<th id="901A7F72FFBD0019A297FA8FFCA5FA2B" box="[696,891,1385,1410]" gridcol="1" gridrow="0" pageId="9" pageNumber="114">
<taxonomicName id="AAFB6FCFFFBDFFEFA297FA8CFCA5FA28" box="[696,891,1385,1409]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBDFFEFA297FA8CFCA5FA28" bold="true" box="[696,891,1385,1409]" italics="true" pageId="9" pageNumber="114">R. mirandaribeiroi</emphasis>
</taxonomicName>
</th>
<th id="901A7F72FFBD0019A4ADFA8FFAD4FA2B" box="[1154,1290,1385,1410]" gridcol="2" gridrow="0" pageId="9" pageNumber="114">
<taxonomicName id="AAFB6FCFFFBDFFEFA4ADFA8CFAD4FA28" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1154,1290,1385,1409]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="114" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBDFFEFA4ADFA8CFAD4FA28" bold="true" box="[1154,1290,1385,1409]" italics="true" pageId="9" pageNumber="114">R. granulosa</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="D3CB160EFFBD0019A08CFA74FAD4FA03" box="[163,1290,1426,1450]" gridrow="1" pageId="9" pageNumber="114" rowspan-0="1">
<td id="901A7F72FFBD0019A297FA74FCA5FA03" box="[696,891,1426,1450]" gridcol="1" gridrow="1" pageId="9" pageNumber="114">N =70</td>
<td id="901A7F72FFBD0019A4ADFA74FAD4FA03" box="[1154,1290,1426,1450]" gridcol="2" gridrow="1" pageId="9" pageNumber="114">N = 61</td>
</tr>
<tr id="D3CB160EFFBD0019A08CFA5CFAD4FA5F" box="[163,1290,1466,1526]" gridrow="2" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CFA5CFE83FA5F" box="[163,349,1466,1526]" gridcol="0" gridrow="2" pageId="9" pageNumber="114">Snout–vent length</th>
<td id="901A7F72FFBD0019A297FA5CFCA5FA5F" box="[696,891,1466,1526]" gridcol="1" gridrow="2" pageId="9" pageNumber="114">47.0 ± 3.9 (38.3–59.3)</td>
<td id="901A7F72FFBD0019A4ADFA5CFAD4FA5F" box="[1154,1290,1466,1526]" gridcol="2" gridrow="2" pageId="9" pageNumber="114">48.7 ± 4.8 (37–59.4)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF9E0FAD4F9EB" box="[163,1290,1542,1602]" gridrow="3" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF9E0FE83F9EB" box="[163,349,1542,1602]" gridcol="0" gridrow="3" pageId="9" pageNumber="114">Thigh length</th>
<td id="901A7F72FFBD0019A297F9E0FCA5F9EB" box="[696,891,1542,1602]" gridcol="1" gridrow="3" pageId="9" pageNumber="114">16.5 ± 1.8 (12.2–20.8)</td>
<td id="901A7F72FFBD0019A4ADF9E0FAD4F9EB" box="[1154,1290,1542,1602]" gridcol="2" gridrow="3" pageId="9" pageNumber="114">17.4 ± 2.1 (13.3–22.3)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF9B4FAD4F926" box="[163,1290,1618,1679]" gridrow="4" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF9B4FE83F926" box="[163,349,1618,1679]" gridcol="0" gridrow="4" pageId="9" pageNumber="114">Tibia length</th>
<td id="901A7F72FFBD0019A297F9B4FCA5F926" box="[696,891,1618,1679]" gridcol="1" gridrow="4" pageId="9" pageNumber="114">16.1 ± 1.5 (13–19.9)</td>
<td id="901A7F72FFBD0019A4ADF9B4FAD4F926" box="[1154,1290,1618,1679]" gridcol="2" gridrow="4" pageId="9" pageNumber="114">17.8 ± 1.9 (14.5–21.8)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF979FAD4F972" box="[163,1290,1695,1755]" gridrow="5" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF979FE83F972" box="[163,349,1695,1755]" gridcol="0" gridrow="5" pageId="9" pageNumber="114">Tarsus length</th>
<td id="901A7F72FFBD0019A297F979FCA5F972" box="[696,891,1695,1755]" gridcol="1" gridrow="5" pageId="9" pageNumber="114">10 ± 1 (8.1–12)</td>
<td id="901A7F72FFBD0019A4ADF979FAD4F972" box="[1154,1290,1695,1755]" gridcol="2" gridrow="5" pageId="9" pageNumber="114">11.1 ± 1.3 (8.4–13.9)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF90DFAD4F88E" box="[163,1290,1771,1831]" gridrow="6" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF90DFE83F88E" box="[163,349,1771,1831]" gridcol="0" gridrow="6" pageId="9" pageNumber="114">Hand length</th>
<td id="901A7F72FFBD0019A297F90DFCA5F88E" box="[696,891,1771,1831]" gridcol="1" gridrow="6" pageId="9" pageNumber="114">10 ± 0.8 (8.4–12.2)</td>
<td id="901A7F72FFBD0019A4ADF90DFAD4F88E" box="[1154,1290,1771,1831]" gridcol="2" gridrow="6" pageId="9" pageNumber="114">11.3 ± 1.2 (8.5–14.4)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF8D1FAD4F8DA" box="[163,1290,1847,1907]" gridrow="7" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF8D1FE83F8DA" box="[163,349,1847,1907]" gridcol="0" gridrow="7" pageId="9" pageNumber="114">Foot length</th>
<td id="901A7F72FFBD0019A297F8D1FCA5F8DA" box="[696,891,1847,1907]" gridcol="1" gridrow="7" pageId="9" pageNumber="114">16.4 ± 1.5 (12.9–19.8)</td>
<td id="901A7F72FFBD0019A4ADF8D1FAD4F8DA" box="[1154,1290,1847,1907]" gridcol="2" gridrow="7" pageId="9" pageNumber="114">18.1 ± 2.0 (13.6–22.2)</td>
</tr>
<tr id="D3CB160EFFBD0019A08CF865FAD4F816" box="[163,1290,1923,1983]" gridrow="8" pageId="9" pageNumber="114">
<th id="901A7F72FFBD0019A08CF865FE83F816" box="[163,349,1923,1983]" gridcol="0" gridrow="8" pageId="9" pageNumber="114">Head height</th>
<td id="901A7F72FFBD0019A297F865FCA5F816" box="[696,891,1923,1983]" gridcol="1" gridrow="8" pageId="9" pageNumber="114">6.5 ± 0.5 (5.5–8.2)</td>
<td id="901A7F72FFBD0019A4ADF865FAD4F816" box="[1154,1290,1923,1983]" gridcol="2" gridrow="8" pageId="9" pageNumber="114">6.7 ± 0.6 (5.5–8.2)</td>
</tr>
</table>
</paragraph>
<footnote id="0EE00842FFBDFFEFA475F836FA4FF841" box="[1114,1425,2000,2024]" pageId="9" pageNumber="114">
<paragraph id="6D44144CFFBDFFEFA475F836FA4FF841" blockId="9.[1114,1425,2000,2024]" box="[1114,1425,2000,2024]" pageId="9" pageNumber="114">
<emphasis id="5F8FC85EFFBDFFEFA475F836FA4FF841" box="[1114,1425,2000,2024]" italics="true" pageId="9" pageNumber="114">......continued on the next page</emphasis>
</paragraph>
</footnote>
<caption id="398444C4FFBEFFECA08CFF44FE4FFF13" box="[163,401,162,186]" isContinuationCaption="true" pageId="10" pageNumber="115" startId="10.[163,247,162,186]" targetType="table">
<paragraph id="6D44144CFFBEFFECA08CFF44FE4FFF13" blockId="10.[163,401,162,186]" box="[163,401,162,186]" pageId="10" pageNumber="115">
<emphasis id="5F8FC85EFFBEFFECA08CFF44FECEFF13" bold="true" box="[163,272,162,186]" pageId="10" pageNumber="115">TABLE 3.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="6D44144CFFBEFFECA08CFF2CFB2AF91A" pageId="10" pageNumber="115">
<table id="1FFBE6ECFFBE0019A08CFF2FFAD4F91A" box="[163,1290,201,1715]" colsContinueFrom="9.[163,1290,1385,1983]" gridcols="3" gridrows="21" pageId="10" pageNumber="115">
<tr id="D3CB160EFFBE0019A08CFF2FFAD4FF4B" box="[163,1290,201,226]" gridrow="0" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFF2FFDECFF4B" box="[163,562,201,226]" gridcol="0" gridrow="0" pageId="10" pageNumber="115">Species -Variables</th>
<th id="901A7F72FFBE0019A297FF2FFCA5FF4B" box="[696,891,201,226]" gridcol="1" gridrow="0" pageId="10" pageNumber="115">
<taxonomicName id="AAFB6FCFFFBEFFECA297FF2CFCA5FF48" box="[696,891,201,225]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="115" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBEFFECA297FF2CFCA5FF48" bold="true" box="[696,891,201,225]" italics="true" pageId="10" pageNumber="115">R. mirandaribeiroi</emphasis>
</taxonomicName>
</th>
<th id="901A7F72FFBE0019A4ADFF2FFAD4FF4B" box="[1154,1290,201,226]" gridcol="2" gridrow="0" pageId="10" pageNumber="115">
<taxonomicName id="AAFB6FCFFFBEFFECA4ADFF2CFAD4FF48" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1154,1290,201,225]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="115" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBEFFECA4ADFF2CFAD4FF48" bold="true" box="[1154,1290,201,225]" italics="true" pageId="10" pageNumber="115">R. granulosa</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="D3CB160EFFBE0019A08CFF14FAD4FEA3" box="[163,1290,242,266]" gridrow="1" pageId="10" pageNumber="115" rowspan-0="1">
<td id="901A7F72FFBE0019A297FF14FCA5FEA3" box="[696,891,242,266]" gridcol="1" gridrow="1" pageId="10" pageNumber="115">N =70</td>
<td id="901A7F72FFBE0019A4ADFF14FAD4FEA3" box="[1154,1290,242,266]" gridcol="2" gridrow="1" pageId="10" pageNumber="115">N = 61</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFEFDFAD4FEFE" box="[163,1290,283,343]" gridrow="2" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFEFDFDECFEFE" box="[163,562,283,343]" gridcol="0" gridrow="2" pageId="10" pageNumber="115">Head width</th>
<td id="901A7F72FFBE0019A297FEFDFCA5FEFE" box="[696,891,283,343]" gridcol="1" gridrow="2" pageId="10" pageNumber="115">14.9 ± 0.9 (12.8–17.7)</td>
<td id="901A7F72FFBE0019A4ADFEFDFAD4FEFE" box="[1154,1290,283,343]" gridcol="2" gridrow="2" pageId="10" pageNumber="115">15.6 ± 1.3 (13–18.3)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFE81FAD4FE0A" box="[163,1290,359,419]" gridrow="3" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFE81FDECFE0A" box="[163,562,359,419]" gridcol="0" gridrow="3" pageId="10" pageNumber="115">Head length</th>
<td id="901A7F72FFBE0019A297FE81FCA5FE0A" box="[696,891,359,419]" gridcol="1" gridrow="3" pageId="10" pageNumber="115">13.2 ± 0.9 (11.3–15)</td>
<td id="901A7F72FFBE0019A4ADFE81FAD4FE0A" box="[1154,1290,359,419]" gridcol="2" gridrow="3" pageId="10" pageNumber="115">13.8 ± 1.2 (11.4–16.4)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFE55FAD4FE46" box="[163,1290,435,495]" gridrow="4" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFE55FDECFE46" box="[163,562,435,495]" gridcol="0" gridrow="4" pageId="10" pageNumber="115">Eye diameter</th>
<td id="901A7F72FFBE0019A297FE55FCA5FE46" box="[696,891,435,495]" gridcol="1" gridrow="4" pageId="10" pageNumber="115">4 ± 0.5 (2.9–5.1)</td>
<td id="901A7F72FFBE0019A4ADFE55FAD4FE46" box="[1154,1290,435,495]" gridcol="2" gridrow="4" pageId="10" pageNumber="115">4.6 ± 0.5 (3.5–5.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFDE6FAD4FD95" box="[163,1290,512,572]" gridrow="5" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFDE6FDECFD95" box="[163,562,512,572]" gridcol="0" gridrow="5" pageId="10" pageNumber="115">Tympanum diameter</th>
<td id="901A7F72FFBE0019A297FDE6FCA5FD95" box="[696,891,512,572]" gridcol="1" gridrow="5" pageId="10" pageNumber="115">2.6 ± 0.4 (1.6–3.4)</td>
<td id="901A7F72FFBE0019A4ADFDE6FAD4FD95" box="[1154,1290,512,572]" gridcol="2" gridrow="5" pageId="10" pageNumber="115">2.7 ± 0.4 (1.9–3.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFDAAFAD4FD21" box="[163,1290,588,648]" gridrow="6" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFDAAFDECFD21" box="[163,562,588,648]" gridcol="0" gridrow="6" pageId="10" pageNumber="115">Tympanum height</th>
<td id="901A7F72FFBE0019A297FDAAFCA5FD21" box="[696,891,588,648]" gridcol="1" gridrow="6" pageId="10" pageNumber="115">3.3 ± 0.4 (2.3–4)</td>
<td id="901A7F72FFBE0019A4ADFDAAFAD4FD21" box="[1154,1290,588,648]" gridcol="2" gridrow="6" pageId="10" pageNumber="115">3.3 ± 0.4 (2.4–4.1)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFD7EFAD4FD7D" box="[163,1290,664,724]" gridrow="7" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFD7EFDECFD7D" box="[163,562,664,724]" gridcol="0" gridrow="7" pageId="10" pageNumber="115">Snout width</th>
<td id="901A7F72FFBE0019A297FD7EFCA5FD7D" box="[696,891,664,724]" gridcol="1" gridrow="7" pageId="10" pageNumber="115">5.4 ± 0.4 (4.2–6.3)</td>
<td id="901A7F72FFBE0019A4ADFD7EFAD4FD7D" box="[1154,1290,664,724]" gridcol="2" gridrow="7" pageId="10" pageNumber="115">5.3 ± 0.5 (4.1–6.5)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFD02FAD4FC89" box="[163,1290,740,800]" gridrow="8" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFD02FDECFC89" box="[163,562,740,800]" gridcol="0" gridrow="8" pageId="10" pageNumber="115">Parotid gland width</th>
<td id="901A7F72FFBE0019A297FD02FCA5FC89" box="[696,891,740,800]" gridcol="1" gridrow="8" pageId="10" pageNumber="115">8.0 ± 1.4 (4.9–10.9)</td>
<td id="901A7F72FFBE0019A4ADFD02FAD4FC89" box="[1154,1290,740,800]" gridcol="2" gridrow="8" pageId="10" pageNumber="115">7.2 ± 1.1 (5–10.9)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFCD7FAD4FCC4" box="[163,1290,817,877]" gridrow="9" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFCD7FDECFCC4" box="[163,562,817,877]" gridcol="0" gridrow="9" pageId="10" pageNumber="115">Parotid gland height</th>
<td id="901A7F72FFBE0019A297FCD7FCA5FCC4" box="[696,891,817,877]" gridcol="1" gridrow="9" pageId="10" pageNumber="115">6.1 ± 0.9 (4.7–9.4)</td>
<td id="901A7F72FFBE0019A4ADFCD7FAD4FCC4" box="[1154,1290,817,877]" gridcol="2" gridrow="9" pageId="10" pageNumber="115">6.1 ± 0.9 (4.3–8.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFC9BFAD4FC10" box="[163,1290,893,953]" gridrow="10" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFC9BFDECFC10" box="[163,562,893,953]" gridcol="0" gridrow="10" pageId="10" pageNumber="115">Internarial distance</th>
<td id="901A7F72FFBE0019A297FC9BFCA5FC10" box="[696,891,893,953]" gridcol="1" gridrow="10" pageId="10" pageNumber="115">2.53 ± 0.2 (1.9–3.1)</td>
<td id="901A7F72FFBE0019A4ADFC9BFAD4FC10" box="[1154,1290,893,953]" gridcol="2" gridrow="10" pageId="10" pageNumber="115">2.6 ± 0.3 (2–3.5)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFC2FFAD4FBAC" box="[163,1290,969,1029]" gridrow="11" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFC2FFDECFBAC" box="[163,562,969,1029]" gridcol="0" gridrow="11" pageId="10" pageNumber="115">Eye–snout distance</th>
<td id="901A7F72FFBE0019A297FC2FFCA5FBAC" box="[696,891,969,1029]" gridcol="1" gridrow="11" pageId="10" pageNumber="115">2.7 ± 0.5 (1.6–3.6)</td>
<td id="901A7F72FFBE0019A4ADFC2FFAD4FBAC" box="[1154,1290,969,1029]" gridcol="2" gridrow="11" pageId="10" pageNumber="115">3.1 ± 0.5 (2.4–4.4)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFBF3FAD4FBF8" box="[163,1290,1045,1105]" gridrow="12" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFBF3FDECFBF8" box="[163,562,1045,1105]" gridcol="0" gridrow="12" pageId="10" pageNumber="115">Eye–nostril distance</th>
<td id="901A7F72FFBE0019A297FBF3FCA5FBF8" box="[696,891,1045,1105]" gridcol="1" gridrow="12" pageId="10" pageNumber="115">4.8 ± 0.7 (3.5–6)</td>
<td id="901A7F72FFBE0019A4ADFBF3FAD4FBF8" box="[1154,1290,1045,1105]" gridcol="2" gridrow="12" pageId="10" pageNumber="115">5.3 ± 0.8 (4–7.5)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFB87FAD4FB34" box="[163,1290,1121,1181]" gridrow="13" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFB87FDECFB34" box="[163,562,1121,1181]" gridcol="0" gridrow="13" pageId="10" pageNumber="115">Interorbital distance</th>
<td id="901A7F72FFBE0019A297FB87FCA5FB34" box="[696,891,1121,1181]" gridcol="1" gridrow="13" pageId="10" pageNumber="115">5.4 ± 0.6 (4.1–6.6)</td>
<td id="901A7F72FFBE0019A4ADFB87FAD4FB34" box="[1154,1290,1121,1181]" gridcol="2" gridrow="13" pageId="10" pageNumber="115">5.3 ± 0.6 (4.5–7.1)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFB48FAD4FB43" box="[163,1290,1198,1258]" gridrow="14" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFB48FDECFB43" box="[163,562,1198,1258]" gridcol="0" gridrow="14" pageId="10" pageNumber="115">Interorbital distance II</th>
<td id="901A7F72FFBE0019A297FB48FCA5FB43" box="[696,891,1198,1258]" gridcol="1" gridrow="14" pageId="10" pageNumber="115">4.9 ± 0.3 (4–5.8)</td>
<td id="901A7F72FFBE0019A4ADFB48FAD4FB43" box="[1154,1290,1198,1258]" gridcol="2" gridrow="14" pageId="10" pageNumber="115">4.9 ± 0.7 (3.8–7.1)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFB1CFAD4FA9F" box="[163,1290,1274,1334]" gridrow="15" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFB1CFDECFA9F" box="[163,562,1274,1334]" gridcol="0" gridrow="15" pageId="10" pageNumber="115">Eyelid width</th>
<td id="901A7F72FFBE0019A297FB1CFCA5FA9F" box="[696,891,1274,1334]" gridcol="1" gridrow="15" pageId="10" pageNumber="115">4.1 ± 0.5 (3.3–5)</td>
<td id="901A7F72FFBE0019A4ADFB1CFAD4FA9F" box="[1154,1290,1274,1334]" gridcol="2" gridrow="15" pageId="10" pageNumber="115">4.4 ± 0.5 (3.5–5.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFAA0FAD4FA2B" box="[163,1290,1350,1410]" gridrow="16" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFAA0FDECFA2B" box="[163,562,1350,1410]" gridcol="0" gridrow="16" pageId="10" pageNumber="115">Supratympanic crest length</th>
<td id="901A7F72FFBE0019A297FAA0FCA5FA2B" box="[696,891,1350,1410]" gridcol="1" gridrow="16" pageId="10" pageNumber="115">2 ± 0.5 (0.8–3.2)</td>
<td id="901A7F72FFBE0019A4ADFAA0FAD4FA2B" box="[1154,1290,1350,1410]" gridcol="2" gridrow="16" pageId="10" pageNumber="115">2.3 ± 0.3 (1.6–3)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFA74FAD4FA66" box="[163,1290,1426,1487]" gridrow="17" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFA74FDECFA66" box="[163,562,1426,1487]" gridcol="0" gridrow="17" pageId="10" pageNumber="115">Postorbital space</th>
<td id="901A7F72FFBE0019A297FA74FCA5FA66" box="[696,891,1426,1487]" gridcol="1" gridrow="17" pageId="10" pageNumber="115">3.3 ± 0.5 (2.3–4.3)</td>
<td id="901A7F72FFBE0019A4ADFA74FAD4FA66" box="[1154,1290,1426,1487]" gridcol="2" gridrow="17" pageId="10" pageNumber="115">4.2 ± 0.6 (3.2–5.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CFA39FAD4F9B2" box="[163,1290,1503,1563]" gridrow="18" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CFA39FDECF9B2" box="[163,562,1503,1563]" gridcol="0" gridrow="18" pageId="10" pageNumber="115">Distance from tympanum to postorbital crest</th>
<td id="901A7F72FFBE0019A297FA39FCA5F9B2" box="[696,891,1503,1563]" gridcol="1" gridrow="18" pageId="10" pageNumber="115">0.3 ± 0.1 (0.1–0.5)</td>
<td id="901A7F72FFBE0019A4ADFA39FAD4F9B2" box="[1154,1290,1503,1563]" gridcol="2" gridrow="18" pageId="10" pageNumber="115">0.4 ± 0.2 (0.1–0.8)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CF9CDFAD4F9CE" box="[163,1290,1579,1639]" gridrow="19" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CF9CDFDECF9CE" box="[163,562,1579,1639]" gridcol="0" gridrow="19" pageId="10" pageNumber="115">Distance from tympanum to supra tympanic crest</th>
<td id="901A7F72FFBE0019A297F9CDFCA5F9CE" box="[696,891,1579,1639]" gridcol="1" gridrow="19" pageId="10" pageNumber="115">0.9 ± 0.2 (0.4–1.4)</td>
<td id="901A7F72FFBE0019A4ADF9CDFAD4F9CE" box="[1154,1290,1579,1639]" gridcol="2" gridrow="19" pageId="10" pageNumber="115">1 ± 0.2 (0.4–1.5)</td>
</tr>
<tr id="D3CB160EFFBE0019A08CF991FAD4F91A" box="[163,1290,1655,1715]" gridrow="20" pageId="10" pageNumber="115">
<th id="901A7F72FFBE0019A08CF991FDECF91A" box="[163,562,1655,1715]" gridcol="0" gridrow="20" pageId="10" pageNumber="115">Distance from tympanum to parotoid gland</th>
<td id="901A7F72FFBE0019A297F991FCA5F91A" box="[696,891,1655,1715]" gridcol="1" gridrow="20" pageId="10" pageNumber="115">0.8 ± 0.3 (0.2–1.5)</td>
<td id="901A7F72FFBE0019A4ADF991FAD4F91A" box="[1154,1290,1655,1715]" gridcol="2" gridrow="20" pageId="10" pageNumber="115">1.3 ± 0.3 (0.7–2.2)</td>
</tr>
</table>
</paragraph>
<paragraph id="6D44144CFFBEFFECA0B8F8E1FE52F888" blockId="10.[151,396,1799,1825]" box="[151,396,1799,1825]" pageId="10" pageNumber="115">
<heading id="360CA320FFBEFFECA0B8F8E1FE52F888" bold="true" box="[151,396,1799,1825]" fontSize="11" level="1" pageId="10" pageNumber="115" reason="1">
<emphasis id="5F8FC85EFFBEFFECA0B8F8E1FE52F888" bold="true" box="[151,396,1799,1825]" pageId="10" pageNumber="115">Molecular Analyses</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFBEFFECA0B8F8A9FBBBF824" blockId="10.[151,1436,1871,1933]" pageId="10" pageNumber="115">
Both species showed expressive genetic divergences (3.8–6.1%) in the mitochondrial fragment. The intraspecific genetic distance was on average 0.09 for
<taxonomicName id="AAFB6FCFFFBEFFECA24AF892FCE9F824" box="[613,823,1907,1933]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="115" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBEFFECA24AF892FCE9F824" box="[613,823,1907,1933]" italics="true" pageId="10" pageNumber="115">R. mirandaribeiroi</emphasis>
</taxonomicName>
and 0.06 for
<taxonomicName id="AAFB6FCFFFBEFFECA3E1F892FB81F824" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[974,1119,1907,1933]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="115" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBEFFECA3E1F892FB81F824" box="[974,1119,1907,1933]" italics="true" pageId="10" pageNumber="115">R. granulosa</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="398444C4FFBFFFEDA0B8FBAFFB69FB67" ID-DOI="http://doi.org/10.5281/zenodo.14896128" ID-Zenodo-Dep="14896128" httpUri="https://zenodo.org/record/14896128/files/figure.png" pageId="11" pageNumber="116" startId="11.[151,250,1097,1121]" targetBox="[158,1429,188,1066]" targetPageId="11" targetType="figure">
<paragraph id="6D44144CFFBFFFEDA0B8FBAFFB69FB67" blockId="11.[151,1436,1097,1230]" pageId="11" pageNumber="116">
<emphasis id="5F8FC85EFFBFFFEDA0B8FBAFFEC4FBCB" bold="true" box="[151,282,1097,1122]" pageId="11" pageNumber="116">FIGURE 5.</emphasis>
Audio spectrogram and corresponding waveform of the advertisement call from the Cerrado of
<taxonomicName id="AAFB6FCFFFBFFFEDA569FBAFFEE7FB2C" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="116" phylum="Chordata" rank="species" species="mirandaribeiroi">
<emphasis id="5F8FC85EFFBFFFEDA569FBAFFEE7FB2C" italics="true" pageId="11" pageNumber="116">Rhinella mirandaribeiroi</emphasis>
</taxonomicName>
(Uberlândia, Minas Gerais state). Individual not collected. (A) Complete advertisement call. (B) Six notes from the middle portion of the call. Note the 5-pulsed structure; the first and last being of lower amplitude. File =
<taxonomicName id="AAFB6FCFFFBFFFEDA515FB77FA4EFB00" authorityName="Fitzinger" authorityYear="1826" box="[1338,1424,1169,1193]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="116" phylum="Chordata" rank="genus">Rhinella</taxonomicName>
_ mirandaribeiroiUberlandiaMG14aAAGGm661Mk2. Air temperature 21°C and water temperature 21°C.
</paragraph>
</caption>
<paragraph id="6D44144CFFBFFFEDA0B8FAF9FED0FA90" blockId="11.[151,270,1311,1337]" box="[151,270,1311,1337]" pageId="11" pageNumber="116">
<heading id="360CA320FFBFFFEDA0B8FAF9FED0FA90" bold="true" box="[151,270,1311,1337]" fontSize="11" level="1" pageId="11" pageNumber="116" reason="1">
<emphasis id="5F8FC85EFFBFFFEDA0B8FAF9FED0FA90" bold="true" box="[151,270,1311,1337]" pageId="11" pageNumber="116">Acoustics</emphasis>
</heading>
</paragraph>
<paragraph id="6D44144CFFBFFFEDA0B8FA8EFE16F99F" blockId="11.[151,1437,1384,1590]" pageId="11" pageNumber="116">
We found conserved call parameter features between the two species. The advertisement call of
<taxonomicName id="AAFB6FCFFFBFFFEDA528FA8EFA43FA2B" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1287,1437,1384,1410]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="116" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="5F8FC85EFFBFFFEDA528FA8EFA43FA2B" box="[1287,1437,1384,1410]" italics="true" pageId="11" pageNumber="116">R. granulosa</emphasis>
</taxonomicName>
consists of a sequence of approximately 170 notes and lasts about 5 seconds, emitted at a rate of 5 calls per minute with 32 notes per second. The notes had a duration of approximately 22 ms and with 4 to 5 pulses per note. The dominant frequency is around 2700 Hz. The calls reach maximum amplitude quickly and remain constant until the end (
<figureCitation id="F5C008C9FFBFFFEDA0E2FA1EFEF0F9BB" box="[205,302,1528,1554]" captionStart="FIGURE 6" captionStartId="13.[151,250,1402,1426]" captionTargetBox="[157,1430,497,1372]" captionTargetId="figure-73@13.[151,1436,490,1378]" captionTargetPageId="13" captionText="FIGURE 6. Audio spectrogram and its respective oscillogram of the advertisement call of Rhinella granulosa (Andarai, BA). The individual was collected (AAG-UFU5663). (A) Complete advertisement call. (B) Seven notes from the middle portion of the advertisement call. Note the 4-pulsed structure; the first and last ones being of lower amplitude. File = Rhinella_ granulosaAndaraiBA1bAAGm671. Air temperature 22°C." figureDoi="http://doi.org/10.5281/zenodo.14896130" httpUri="https://zenodo.org/record/14896130/files/figure.png" pageId="11" pageNumber="116">Figure 6</figureCitation>
). Further details can be found in
<tableCitation id="207921F7FFBFFFEDA2B8FA1EFD32F9BB" box="[663,748,1528,1554]" captionStart="TABLE 4" captionStartId="11.[152,244,1637,1661]" captionTargetPageId="11" captionText="TABLE 4. Advertisement call variables for the species R. mirandaribeiroi and R. granulosa. N = number of individuals and analyzed calls; data presented as mean ± standard deviation (minimum–maximum)." httpUri="http://table.plazi.org/id/398444C4FFBFFFEDA0B7F983FB9FF908" pageId="11" pageNumber="116" tableUuid="398444C4FFBFFFEDA0B7F983FB9FF908">Table 4</tableCitation>
. Our results are consistent with the published analysis of these data (Giarretta
<emphasis id="5F8FC85EFFBFFFEDA16EF9FAFEA7F99F" box="[321,377,1564,1590]" italics="true" pageId="11" pageNumber="116">et al.</emphasis>
2018).
</paragraph>
</subSubSection>
</treatment>
</document>