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<mods:title id="74D3FE1E24DE340331DCD4C8D6123ECA">Oligocene vertebrate footprints from the Lower Red Formation, Central Iran</mods:title>
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<mods:namePart id="97799FDBB51D8C9DD5A0BE70B98D6DDB">Abbassi, Nasrollah</mods:namePart>
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<mods:affiliation id="001D22A462C99FBECB85D1B047788D2B">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran</mods:affiliation>
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<mods:namePart id="CB8D6CB85256022EF70A1056D14CB6EE">Salehi Tinooni, Mohammad</mods:namePart>
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<mods:affiliation id="79727004088BF7E56CFD19CEB56F0020">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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<mods:namePart id="40FF40C026A7CE3705DAA933F221719B">Ghorbani Dehnavi, Mahdi</mods:namePart>
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<mods:affiliation id="F01CC3928DD5685ADADEC303DB2679E0">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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<mods:namePart id="A92E3C76B052E82D7C5A867208B40BCE">Shakeri, Safoora</mods:namePart>
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<mods:affiliation id="D62E937335C823070684292FB5F424AA">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran & Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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<mods:namePart id="D518E534DED39BB30425D189EC98AD47">Eshaghi, Ali</mods:namePart>
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<mods:affiliation id="26139C1F0CEA9C0FDD87C0E3232EA301">Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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<mods:title id="27831C617DE57F55617116468ED35A2D">Fossil Record</mods:title>
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<mods:part id="49ECB245345589FE06C672DCC8437607">
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<mods:date id="E5ECB7290643210E09E9BB23B5794EE8">2024</mods:date>
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<mods:number id="F0B9E3B9D56F33E2EDF576D8F84FA65A">2024-11-04</mods:number>
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<mods:number id="24699339A4C291FF9D3EBD6B7D06A0A9">27</mods:number>
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<mods:classification id="2840DC7E6AFE155BF49023C3B86DF09F">journal article</mods:classification>
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<mods:identifier id="9DB27E21A7FB5C829B6866F53E91A5D8" type="DOI">10.3897/fr.27.133914</mods:identifier>
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<treatment id="B73803F124285A358E52BCF1C54A7A67" ID-arpha="B73803F1-2428-5A35-8E52-BCF1C54A7A67" LSID="urn:lsid:plazi:treatment:B73803F124285A358E52BCF1C54A7A67" httpUri="http://treatment.plazi.org/id/B73803F124285A358E52BCF1C54A7A67">
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<subSubSection id="E7B8B780F1C2D073E23F34849BFD46A5" type="nomenclature">
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<paragraph id="16B9BB8D229E2203D5360E324AF372DC">
|
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<heading id="6E040DF8AD999D03CAFD8F6C94BADE89" reason="label">Ichnogenus</heading>
|
||||
<taxonomicName id="85D18862DD4F13FA2BA3BFA3CCC2C510" authority="Abbassi, Alinasiri & Lucas, 2017" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" class="Mammalia" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
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<emphasis id="E74C8CB2713CC57D571EE8C562BF1936" italics="true">Moropopus</emphasis>
|
||||
<bibRefCitation id="7AAC43F3C0DFB874CDFC6C3B94759524" DOI="10.1080/08912963.2016.1267162" author="Abbassi N & Alinasiri S & Lucas SG" journalOrPublisher="Historical Biology" pagination="987-1006" refId="B4" refString="Abbassi N, Alinasiri S, Lucas SG (2017) New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran. Historical Biology 29: 987 - 1006. https: // doi. org / 10.1080 / 08912963.2016. 1267162" title="New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran." volume="29" year="2017">Abbassi, Alinasiri & Lucas, 2017</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EQTAI" type="type_taxon">
|
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<paragraph id="6B441B5A50C0C90DFE3638C0F973D893">
|
||||
<heading id="8E712940F339C04B3D0853E5AD0F9FB0" reason="title">
|
||||
<typeStatus id="3B6C013123AF81361F9B4627335DCD68">Type</typeStatus>
|
||||
ichnospecies.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7A9FB6F4A1FB5820613676D14F84E876">
|
||||
<taxonomicName id="AFB1A5688E543F61691AC565B8CF0A3E" authority="Abbassi, Alinasiri & Lucas, 2017" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="elongatus">
|
||||
<emphasis id="B22AD131D4DB3A73CEC53C436A671800" italics="true">Moropopus elongatus</emphasis>
|
||||
<bibRefCitation id="E7CAB0DFB0068EB9E8F1D73C24D89AFB" DOI="10.1080/08912963.2016.1267162" author="Abbassi N & Alinasiri S & Lucas SG" journalOrPublisher="Historical Biology" pagination="987-1006" refId="B4" refString="Abbassi N, Alinasiri S, Lucas SG (2017) New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran. Historical Biology 29: 987 - 1006. https: // doi. org / 10.1080 / 08912963.2016. 1267162" title="New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran." volume="29" year="2017">Abbassi, Alinasiri & Lucas, 2017</bibRefCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBUAI" type="Emended diagnosis">
|
||||
<paragraph id="43765F9BA2987BDD9A64FD527EA3F911">
|
||||
<heading id="6847D275BE70F5C01FECED8E64FB47EF" reason="title">Emended diagnosis.</heading>
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||||
</paragraph>
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<paragraph id="9B9DFF28F8CDC1507B08137224512336">
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Medium-sized tridactyl footprints with elongated oval to ellipsoidal to fusiform digit imprints. Two digital pads may be present in lateral digits. Digit imprints connect to metatarsus by a narrower proximal imprint. Lateral digits (digit
|
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<collectionCode id="BAE05E8E52C46DE0B9C0B998D94EA7DF">II</collectionCode>
|
||||
and
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<collectionCode id="C915FABECE6358BDD0DD0E15ACF4186F">IV</collectionCode>
|
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) are smaller than the middle digit (digit
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<collectionCode id="EE2B25E6E69342E097B550077AB18E07">III</collectionCode>
|
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) and are curved toward the front. Digit
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<collectionCode id="3CEBE04C80430C3C2EB1429F00827F32">III</collectionCode>
|
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imprint is wider in the front. Digit
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<collectionCode id="0DEFB0257DB1A9959D62D8BD160F2B28">III</collectionCode>
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tip is completely curved, and the tips of lateral digits may curve or end in elongate sharp tips, like the claw imprints. Metatarsus imprints include two metatarsal pads with a complete rounded back, or with two lobes.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="SECID0EGUAI" type="discussion">
|
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<paragraph id="991E718D1CAF303658C21206F3960646">
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<heading id="5759C21881ED2552267E653BF172925B" reason="title">Discussion.</heading>
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</paragraph>
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<paragraph id="475A35D0679DC433E585CCE747D6E0A3">
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The narrow proximal part of digit imprints is a distinctive feature of
|
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<taxonomicName id="E8F629A2EBB1C63B5D39D6783DE0039D" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="D805D34A8DED07DFE527E73CA3FC3A42" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
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||||
, which distinguishes it from the other tridactyl ichnogenera of perissodactyls. The presence of sharp tips on the lateral digits of the pes imprint of the Deh Nar footprints caused an emended diagnosis of
|
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<taxonomicName id="0EA3849A74CA36F746092941A5FD95E8" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="2ED0A86C31E22A6819CD28C05FD5B622" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
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and introduced a second ichnospecies of
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<taxonomicName id="3CF6E1DD13231A5251289F5970F8350D" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="6D5C56F685218A3133A66CC89BB8E11F" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
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</paragraph>
|
||||
<paragraph id="1894C2D7C597168835FA27F0FF6CE27A">
|
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In Asia, mammal tridactyl ungulate footprints have been reported from Early Tertiary of
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<collectingCountry id="846505F4B35659213A8AFCC16BACCEE5" name="China">China</collectingCountry>
|
||||
(Lockley et al. 1999), Oligoene of
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<collectingCountry id="661556F1BE1226277AE8DCEB19DFC5D6" name="India">India</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="CA8BA8FC9B43F66A15F7F97CAE46A9B5" DOI="10.1007/s12040-013-0396-6" author="Rajkumar and Klein" firstAuthor="Rajkumar" issue="2" journalOrPublisher="Journal of Earth System Science" pagination="413-420" refId="B82" refString="Rajkumar HS, Klein H (2014) First perissodactyl footprints from Flysch deposits of the Barail Group (Lower Oligocene) of Manipur, India. Journal of Earth System Science 123 (2): 413–420. https://doi.org/10.1007/s12040-013-0396-6" title="First perissodactyl footprints from Flysch deposits of the Barail Group (Lower Oligocene) of Manipur, India." volume="123" year="2014">Rajkumar and Klein 2014</bibRefCitation>
|
||||
) and Eocene-Oligocene of
|
||||
<collectingCountry id="DA42D6C60D256EE8968B71AFCCF39F34" name="Iran">Iran</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="CCC257CB8F8EE5EAB77D93E7F590776D" DOI="10.1016/j.palaeo.2015.08.039" author="Abbassi" etAl="et al." firstAuthor="Abbassi" journalOrPublisher="Palaeogeography, Palaeoclimatology, Palaeoecology" pagination="78-89" refId="B7" refString="Abbassi N, Lucas SG, Zaare GR (2015) First Report of Oligocene Vertebrate Footprints from Iran. Palaeogeography, Palaeoclimatology, Palaeoecology 440: 78–89. https://doi.org/10.1016/j.palaeo.2015.08.039" title="First Report of Oligocene Vertebrate Footprints from Iran." volume="440" year="2015">Abbassi et al. 2015</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="7DA82A45DFD1275F8B9A781BA9878A65" DOI="10.1080/08912963.2016.1267162" author="Abbassi" firstAuthor="Abbassi" journalOrPublisher="Historical Biology" pagination="987-1006" refId="B4" refString="Abbassi N, Alinasiri S, Lucas SG (2017) New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran. Historical Biology 29: 987–1006. https://doi.org/10.1080/08912963.2016.1267162" title="New localities of Late Eocene vertebrate footprints from the Tarom Mountains, Northwestern Iran." volume="29" year="2017">2017</bibRefCitation>
|
||||
). Oligoene tridactyl footprints of
|
||||
<collectingCountry id="9A961B4B6B1D0BE9FDFD7A68BD72693A" name="India">India</collectingCountry>
|
||||
strongly resemble those described from
|
||||
<collectingCountry id="6A30CD8C7321580D1A7A6DEE4646A0C6" name="China">China</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="6E0F5D3AD1DD8BBA199621FB9E3DD5A4" DOI="10.1007/s12040-013-0396-6" author="Rajkumar and Klein" firstAuthor="Rajkumar" issue="2" journalOrPublisher="Journal of Earth System Science" pagination="413-420" refId="B82" refString="Rajkumar HS, Klein H (2014) First perissodactyl footprints from Flysch deposits of the Barail Group (Lower Oligocene) of Manipur, India. Journal of Earth System Science 123 (2): 413–420. https://doi.org/10.1007/s12040-013-0396-6" title="First perissodactyl footprints from Flysch deposits of the Barail Group (Lower Oligocene) of Manipur, India." volume="123" year="2014">Rajkumar and Klein 2014</bibRefCitation>
|
||||
) and differ from Iran’s Oligocene tridactyl footprints in morphology and size.
|
||||
<bibRefCitation id="DF6740145901F1C60DCB9BD2F288CB89" author="Demathieu G & Ginsburg L & Guerin C & Truc G" etAl="et al." firstAuthor="Demathieu" refId="B28" refString="Demathieu G, Ginsburg L, Guerin C, Truc G (1984) Étude paléontologique, ichnologique et paléoécologique du gisement oligocène de Saignon (bassin d’Apt, Vaucluse). Bulletin du Museum National d’Histoire Naturelle, 4 e série, section C 2: 153–183." year="1984">Demathieu et al. (1984)</bibRefCitation>
|
||||
reported
|
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<taxonomicName id="7ADC098263C32251B3D3143B5FAFD37B" authorityName="Demathieu, Ginsburg, Guérin & Truc" authorityYear="1984" class="Mammalia" genus="Ronzotherichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
|
||||
<emphasis id="F7082D92E29044E534832C8EA4D3B552" italics="true">Ronzotherichnus</emphasis>
|
||||
</taxonomicName>
|
||||
with short digit imprints not connected to the metatarsus imprint and thus differing from
|
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<taxonomicName id="8CEF8CBFC67A75B52C2ACB443B135768" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="F1435F43F81437B4F0FD7F755E107458" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
|
||||
. Usually, tridactyl footprints in perissodactyls show round outlines in the distal part of digits, however some ichnogenera of tridactyls have sharp tips in the lateral digits, such as
|
||||
<taxonomicName id="D0E0C623C11861FDB82F059C7E32FF50" authority="Santamaria et al. (1989 - 1990" authorityName="Santamaria et al. (1989 -" authorityYear="1990" class="Mammalia" genus="Plagiolophustipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B9FB7ED0B1C91783BD4B195923859D61" italics="true">Plagiolophustipus</emphasis>
|
||||
<bibRefCitation id="A6F8939DBD99B3CC14513342FB285332" author="Santamaria R & López G & Casanovas-Cladellas ML" journalOrPublisher="Paleontologia i Evolució" pagination="141-152" refId="B86" refString="Santamaria R, López G, Casanovas-Cladellas ML (1989–1990) Nuevos yacimientos con icnitas de mamíferos del Oligoceno de los alrededores de Agramunt (Lleida, España). Paleontologia i Evolució 23: 141–152." title="Nuevos yacimientos con icnitas de mamíferos del Oligoceno de los alrededores de Agramunt (Lleida, España)." volume="23">Santamaria et al. (1989–1990</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
<taxonomicName id="18FEBA4DBE7D6F8CC042843D92AE8ACB" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="E68DDD889D86795F3FF1419181C74CF9" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="B8DBB5DBD0A68DE95F6F71318F2630A8" authorityName="Santamaria et al. (1989 -" authorityYear="1990" class="Mammalia" genus="Plagiolophustipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
|
||||
<emphasis id="CCF75A6DEF9B4FB1B81CAA68C70831F5" italics="true">Plagiolophustipus</emphasis>
|
||||
</taxonomicName>
|
||||
by its narrow digit
|
||||
<collectionCode id="B5E80A2A4273B508C00204682D95F4DE">III</collectionCode>
|
||||
imprint.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
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