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<document id="FE32540138393E7078C51105FDF43688" ID-DOI="10.3897/fr.27.133914" ID-ZooBank="A30F4FFF-547A-4323-80C0-771663DF6FB7" ID-publisher-id="133914" URI-arpha="66B94E5B-EC0A-579D-B7ED-F85ED4A51F96" URI-zoobank="http://zoobank.org/A30F4FFF-547A-4323-80C0-771663DF6FB7" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1730736172305" checkinUser="pensoft" docAuthor="Abbassi, Nasrollah, Salehi Tinooni, Mohammad, Ghorbani Dehnavi, Mahdi, Shakeri, Safoora & Eshaghi, Ali" docDate="2024" docId="065C0489DA765CFE8AE1414C6AEA184E" docLanguage="en" docName="Fossil Record 27(2): 265-287" docOrigin="Fossil Record 27 (2)" docSource="https://fr.pensoft.net/article/133914/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Moropopus kashanensis Abbassi, Tinooni, Dehnavi, Shakeri & Eshaghi, 2024, isp. nov." docType="treatment" docUuid="0E24AF8D-52D9-4C29-A514-4FF98C339401" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="287" masterDocId="66B94E5BEC0A579DB7EDF85ED4A51F96" masterDocTitle="Oligocene vertebrate footprints from the Lower Red Formation, Central Iran" masterLastPageNumber="287" masterPageNumber="265" pageNumber="265" updateTime="1730752961050" updateUser="admin">
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<mods:mods id="134F5DD53C1808F06E8C1E758E76351F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BBA4B2257583ABD6CCFA4F9C4FF3AA55">
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<mods:title id="7A3C6F79BC4AC4A1AC7ADFE131ABFF3A">Oligocene vertebrate footprints from the Lower Red Formation, Central Iran</mods:title>
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<mods:name id="55562E531CB2CEFBC197917A0AA7F19B" type="personal">
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<mods:namePart id="6EEA19B7C2DAE60BC79D55F2AD60D577">Abbassi, Nasrollah</mods:namePart>
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<mods:nameIdentifier id="F8DD2220687BEE432783D57B2910547C" type="ORCID">0000-0002-9994-9122</mods:nameIdentifier>
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<mods:affiliation id="F8DC7833C1C394417D81D1C58AE811D5">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran</mods:affiliation>
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</mods:name>
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<mods:name id="E6FD43C7DF9A0ED9134E1834A886DC0B" type="personal">
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<mods:roleTerm id="0DA7833208FE274FEAD82968E741D261">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8F64EEA1486C51B2AA3F1AF39F4A3879">Salehi Tinooni, Mohammad</mods:namePart>
|
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<mods:affiliation id="902B8221F443DEE4D160B96251BA2445">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="E1A612AD9AA18911633B25C184E0FA43">Ghorbani Dehnavi, Mahdi</mods:namePart>
|
||||
<mods:affiliation id="4EB6521E4808709B99FCD4ECE5418708">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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</mods:role>
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<mods:namePart id="D8ED5A2C4074053C4EAE9BE469CEAD72">Shakeri, Safoora</mods:namePart>
|
||||
<mods:affiliation id="723C9FF449B8435CB5EFDAAB1D2B4F24">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran & Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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</mods:role>
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<mods:namePart id="1615FE8B8EA3C4A66FC28FC0284DA886">Eshaghi, Ali</mods:namePart>
|
||||
<mods:affiliation id="51CC414D1EF2F5F5F640A22C8609FDE0">Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="7F68CD5C941479171B9D744FFEA74292">text</mods:typeOfResource>
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<mods:titleInfo id="7B90206CFEF1494C34D711B86332B421">
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<mods:title id="9720FB30A4764A9E4104CA0B5401C1FF">Fossil Record</mods:title>
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</mods:titleInfo>
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<mods:part id="FEFF49C21890712778AC43DC01180D01">
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<mods:date id="8AAA7BF0DBE91C48B32E61E49B4680C0">2024</mods:date>
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<mods:detail id="AAC9478D1B23B32484CF546A760543C0" type="pubDate">
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<mods:number id="3AFFE0EA2AA06BCC411BFC589D6F4F62">2024-11-04</mods:number>
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<mods:detail id="06FFB39F9311560E3CF6EDE77E4253D8" type="volume">
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<mods:number id="4695F4B860AF3FA7C6C4B9FC556DC900">27</mods:number>
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</mods:detail>
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<mods:detail id="43BCC58808252FF1EE5F27A391DC7959" type="issue">
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<mods:number id="F5078491BDA1295E01B4114730848888">2</mods:number>
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<mods:extent id="95E3585E8B8A97539642AB855FE3F31C" unit="page">
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<mods:start id="2D89F63CCFC64F3512D715BC6570C988">265</mods:start>
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<mods:end id="8495251D07A653B4F03D36C8D4B9F47A">287</mods:end>
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<mods:classification id="409F8328FBCE99B052BE1271A521250D">journal article</mods:classification>
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<mods:identifier id="BC58A2B36EBDE64731AEF692987DAF83" type="DOI">10.3897/fr.27.133914</mods:identifier>
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<mods:identifier id="B9F5A9AC69BE7DD4FD5718F9B20C7377" type="ZooBank">A30F4FFF-547A-4323-80C0-771663DF6FB7</mods:identifier>
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</mods:mods>
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<treatment id="065C0489DA765CFE8AE1414C6AEA184E" ID-arpha="065C0489-DA76-5CFE-8AE1-414C6AEA184E" ID-zoobank="https://zoobank.org/0E24AF8D-52D9-4C29-A514-4FF98C339401" LSID="urn:lsid:zoobank.org:act:0E24AF8D-52D9-4C29-A514-4FF98C339401" LSID-ZBK="urn:lsid:zoobank.org:act:0E24AF8D-52D9-4C29-A514-4FF98C339401" httpUri="http://treatment.plazi.org/id/065C0489DA765CFE8AE1414C6AEA184E">
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<subSubSection id="4C638DECA269D6368CA8E36997EEF4D5" type="nomenclature">
|
||||
<paragraph id="B6F91FB562112FF5109F850D9FA21873">
|
||||
<heading id="BAE24DB52D8D6688AFA8BF1E221CD52A" reason="label">Ichnospecies</heading>
|
||||
<taxonomicName id="618BDEFB3DF4D50C7946456B0A3B388C" authorityName="Abbassi & Salehi Tinooni & Ghorbani Dehnavi & Shakeri & Eshaghi" authorityYear="2024" class="Mammalia" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="kashanensis" status="sp. nov.">
|
||||
<emphasis id="D67C24204B24579882D5B8E3E3A49E62" italics="true">Moropopus kashanensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D03CD4582BB37C31C8AF6D386A6314F9" rank="species">isp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C1D670B5F740A6F037C004A51964AB37" type="description">
|
||||
<paragraph id="29E6302FE229C73165D765489F7F34F0">
|
||||
<figureCitation id="8C1AEAD63DFD8979795922791D707D95" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Moropopus kashanensis nov. in the studied slab (A); B – D. Holotype of pes imprint with outline sketch and photogrammetry photo (IFMI - 686 / 1); E – I. Holotype of manus imprint with its outline sketch (F) and photogrammetric false-color depth maps, (G, H) and closeup of digit claw imprint (I arrow), (IFMI - 686 / 2)." figureDoi="10.3897/fr.27.133914.figure8" httpUri="https://binary.pensoft.net/fig/1167590">Fig. 8</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E42AI" type="material">
|
||||
<paragraph id="6E44983B3814C1B09F5C2101A42977A8">
|
||||
<heading id="EF0D4952DBE352A5CFC6467ECA7D8F8A" reason="title">Materials.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4E7D8959320EF402495FC5B3A09FDB24">
|
||||
<materialsCitation id="ED2F804651AE2B1C5A392712CE2D728E" collectionCode="IFMI" specimenCode="IFMI-686" specimenCount="1">
|
||||
Two footprints of one trackway, sampled (
|
||||
<specimenCode id="1F508DA966FD1B8FE63008AAEC887FE5">
|
||||
<abbrev content-type="institution" id="ABBRID0ED3AI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
|
||||
-686
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||||
</specimenCode>
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||||
/ 1-2)
|
||||
</materialsCitation>
|
||||
.
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||||
</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection id="SECID0EI3AI" type="Holotype">
|
||||
<paragraph id="810B4C22D3C9BD2A58F4142ABB8D32C1">
|
||||
<heading id="70D0B4D3B0B9B5D64709F2593504631C" reason="title">
|
||||
<typeStatus id="282E30E6FDDB3E4581A12B5B18BFAC2B" type="holotype">Holotype</typeStatus>
|
||||
.
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||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B10811488860274AD5C640C6CF625721">
|
||||
<materialsCitation id="8836976B715A6EEBF2D50E19AE253A1B" collectionCode="IFMI" specimenCode="IFMI-686" specimenCount="1" typeStatus="holotype">
|
||||
specimen
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||||
<specimenCode id="906E7CE296127571BAFABD378FA37428">
|
||||
<abbrev content-type="institution" id="ABBRID0EO3AI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
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||||
-686
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||||
</specimenCode>
|
||||
</materialsCitation>
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||||
.
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ET3AI" type="etymology">
|
||||
<paragraph id="04B0AE89BC3F589870F69ECCF7C3F890">
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||||
<heading id="85BE4ABF255FA68529811BE58EE89283" reason="title">Etymology.</heading>
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||||
</paragraph>
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||||
<paragraph id="F65582ED7E6DCC695385D274A009B97A">
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<emphasis id="1ACF7B934F56C5D1A1A2D46AAE948A40" italics="true">kashanensis</emphasis>
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refers to Kashan city where the traces were found.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="SECID0ED4AI" type="Type-locality">
|
||||
<paragraph id="500AFB08F1D81D96357AFFFAC105F497">
|
||||
<heading id="9872D160B0DC734C79CE2019A4A3D400" reason="title">Type-locality.</heading>
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||||
</paragraph>
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||||
<paragraph id="68312CF5B9DA8A2F94EA2DF9B68389CC">
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||||
<collectingCountry id="0CB5EAA1AA7E9B82341831835C1F7B5E" name="Iran">Iran</collectingCountry>
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||||
,
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<collectingRegion id="F5DFC8EE2191E8BB24A8CD01A2E5E511" country="Iran" name="Esfahan">Esfahan</collectingRegion>
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||||
(=
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<collectingRegion id="6B86B2655A6D88B3851143FFB5C43C9E" country="Iran" name="Esfahan">Isfahan</collectingRegion>
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) province, Kashan township, west Ab Shirin, northwest Deh Nar,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EM4AI" specific-use="{"type":"Point","coordinates":[51.200000,34.316667]}">
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||||
<geoCoordinate id="41112A8B2C1224371B921AFE66B0B581" degrees="34" direction="north" minutes="19" orientation="latitude" precision="925" value="34.316666">34 ° 19 ' N</geoCoordinate>
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,
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<geoCoordinate id="E147A73DB2AD76CBEA0386220BE74F66" degrees="51" direction="east" minutes="12" orientation="longitude" precision="925" value="51.2">51 ° 12 ' E</geoCoordinate>
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</named-content>
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.
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</paragraph>
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</subSubSection>
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||||
<subSubSection id="SECID0ER4AI" type="Type horizon">
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||||
<paragraph id="A4624A60C772FDE7481967EE80E43F71">
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<heading id="0EA32C7A407FC53970E37936060176B8" reason="title">Type horizon.</heading>
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</paragraph>
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<paragraph id="69FE5441F94B404A598E337909AAF55B">lower Oligocene (Rupelian).</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="SECID0EW4AI" type="diagnosis">
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||||
<paragraph id="1DCEC139118D2C8909F89E224AA10CDC">
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||||
<heading id="7D36DE8CA99A5F1A2BAA3AB4E4E589B4" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FD787CCBBEB116611ABD4928146F017F">
|
||||
<taxonomicName id="D3DE99F3D3B30C14BFBC8E59CCC5005A" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
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||||
<emphasis id="D2FFADAAC08219711CE00C9ED1C618C0" italics="true">Moropopus</emphasis>
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||||
</taxonomicName>
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with tridactyl footprint in pes and manus imprints. Pes has elongated digit imprints; digit
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||||
<collectionCode id="B8482EF43F8C55F600C387899D9C55F4">III</collectionCode>
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||||
is larger than the lateral digits
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||||
<collectionCode id="65B64B9C9AC676B1D841DD3CFBACB667">II</collectionCode>
|
||||
and
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||||
<collectionCode id="2850B93C4F073CF6D8D9ACBA5297E192">IV</collectionCode>
|
||||
. Tips of digits are completely round. The manus imprint has a digit
|
||||
<collectionCode id="1797E9689FC2D67F0AF447CCF1DB5678">III</collectionCode>
|
||||
that is longer than the lateral digits. The tip of digit
|
||||
<collectionCode id="1960231BDCB63781B2A78A91C8732398">III</collectionCode>
|
||||
is round, but lateral digits end with a distally sharp claw-like imprint.
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</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="SECID0ED5AI" type="description">
|
||||
<paragraph id="08AD06C14BE953DD90A149E782B54F8B">
|
||||
<heading id="06B01341FC0717943EE9147925A328A6" reason="title">Description.</heading>
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||||
</paragraph>
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<paragraph id="4C947D08EBA3C01A644E80B779113499">The smaller posterior footprint is considered as a pes imprint and it includes two digit imprints and one of the lateral digit and most parts of the metapodial imprints are omitted by weathering. Digits include one pedal circular imprint with round tips, and connect by a narrower imprint to the metapodial imprint. The anterior larger imprint is the manus imprint and it is distinguished by the sharp tip of the lateral digit imprints and it seems that the middle digit imprint has a round tip. The metacarpal imprint is circular, with a round proximal outline.</paragraph>
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<caption id="850518BED7ED51E40FCAABCAD363BB70" ID-DOI="10.3897/fr.27.133914.figure8" ID-arpha="00C37C2B-DA70-5216-A49C-16F43ECFA328" httpUri="https://binary.pensoft.net/fig/1167590" startId="F8">
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||||
<paragraph id="ACD61BC3B87433B06BE3FDAFF6517358">
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<label id="BCDE68E4EB9D220811F476C3EB5FEA85">Figure 8.</label>
|
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</paragraph>
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<paragraph id="474245CD4970E53D35D2711090E9FBE2">
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<taxonomicName id="3B055796CA04537114DC27BB5A5322C9" authorityName="Abbassi & Salehi Tinooni & Ghorbani Dehnavi & Shakeri & Eshaghi" authorityYear="2024" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="kashanensis">
|
||||
<emphasis id="A36EBFE6BD3415F8AE628C4D960B7DB8" italics="true">Moropopus kashanensis</emphasis>
|
||||
</taxonomicName>
|
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nov. in the studied slab (
|
||||
<emphasis id="C7CEE2CC4EC1CDD90180762AE758F18D" bold="true">A</emphasis>
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||||
);
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||||
<emphasis id="5D95983E9D521136476EE57A6C2C3B8C" bold="true">B – D.</emphasis>
|
||||
Holotype of pes imprint with outline sketch and photogrammetry photo (
|
||||
<abbrev content-type="institution" id="ABBRID0E65AI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
|
||||
- 686 / 1);
|
||||
<emphasis id="DA31E1B80C33F30398CED6E557C55BFC" bold="true">E – I.</emphasis>
|
||||
Holotype of manus imprint with its outline sketch (
|
||||
<emphasis id="F9F66CEA9383AA8E7AFE517BD0250F3B" bold="true">F</emphasis>
|
||||
) and photogrammetric false-color depth maps, (
|
||||
<emphasis id="B17417F00FB00120D604DE9021040AFA" bold="true">G, H</emphasis>
|
||||
) and closeup of digit claw imprint (
|
||||
<emphasis id="4CBCF6B5EF4912326BA74900C69C7451" bold="true">I</emphasis>
|
||||
arrow), (
|
||||
<abbrev content-type="institution" id="ABBRID0EM6AI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
|
||||
- 686 / 2).
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EW6AI" type="discussion">
|
||||
<paragraph id="14DDD8E0D4B1FEA3470BF9C1C157D517">
|
||||
<heading id="3FB3D1F688E081D7832DD6F3A3E89A99" reason="title">Discussion.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="31EA37655B2C68AA15B09C7609532975">
|
||||
The general morphology of
|
||||
<taxonomicName id="9146D292D6F3C8ED474AE9FDD7C49FCE" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="88DE1186BD2413716B5A8804CF62B993" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
|
||||
with elongated distally swollen digit imprints is traceable in these footprints, but the distinctive feature is the sharp tips of lateral digits. Usually, perissodactyls have round distal outlines in hooves, and there are no records of sharp tips. The skeletal remains of pes and manus of perissodactyls anatomically comprise distally round phalanges (ungula) with or without medial fissure and covered by fat digital pads or round unguals. Instead, the unguals of artiodactyls usually have sharp tips.
|
||||
</paragraph>
|
||||
<paragraph id="4C545AE5A30BFC9022216AB3DEB12381">
|
||||
<taxonomicName id="D3DB002ADB91407A115D07BE8EF5AA72" authorityName="Abbassi & Salehi Tinooni & Ghorbani Dehnavi & Shakeri & Eshaghi" authorityYear="2024" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="kashanensis">
|
||||
<emphasis id="2B676CF3D76E266A9DA7169467A19D9C" italics="true">Moropopus kashanensis</emphasis>
|
||||
</taxonomicName>
|
||||
however shows strongly sharp tips in lateral digits of the manus imprint. No secondary deformations such as drag marks or combination with non-biogenic structures were included in the detailed analysis of the tips of
|
||||
<taxonomicName id="275297CE0E328BBFC3270492B34A88E8" authorityName="Abbassi & Salehi Tinooni & Ghorbani Dehnavi & Shakeri & Eshaghi" authorityYear="2024" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="kashanensis">
|
||||
<emphasis id="EC050CB50109CA6B6A87ECF5F25732DC" italics="true">Moropopus kashanensis</emphasis>
|
||||
</taxonomicName>
|
||||
. Thus, we believe these footprints are a new ichnospecies of
|
||||
<taxonomicName id="FFF7C108FA8EBC545D2EFCB513294DFC" authorityName="Abbassi, Alinasiri & Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="genus">
|
||||
<emphasis id="EB31BC7F4760DB7DBC1BD4DCE282FF09" italics="true">Moropopus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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