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<document id="7FB3B6369793DB8F1CC4343116CBA760" ID-DOI="10.50826/bnmnszool.48.4_139" ID-ISSN="2434-091X" ID-Zenodo-Dep="12760064" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721224785302" checkinUser="felipe" docAuthor="Xiang, Ke, Pan, Jinbin, Yu, Jiaojiao, Xiao, Lehui, Sun, Shao-Kai &amp; Cheng, Ran" docDate="2022" docId="561C87BEC160FFC880EF69DD5971FA56" docLanguage="en" docName="BullNatlMusNatSciSerA.48.4.139-146.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology (Zoology) 48 (4)" docSource="http://dx.doi.org/10.1039/d4ay00658e" docStyle="DocumentStyle:A6FFB80BFE6276686563735C80A89639.2:BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleId="A6FFB80BFE6276686563735C80A89639" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleVersion="2" docTitle="Callobius cavernarius Xiang, Pan, Yu, Xiao, Sun &amp; Cheng, 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="144" masterDocId="AA25FFC6C163FFCD80146B6A5A3CFF99" masterDocTitle="Two New Species of the Spider Family Amaurobiidae from the Ryukyu Islands, Japan" masterLastPageNumber="146" masterPageNumber="139" pageNumber="142" updateTime="1721225202872" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="64C2F5C8BF334C6C0B7500D36C312D51">Two New Species of the Spider Family Amaurobiidae from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="8BB810B65482160022002FD7E60B7CEF">Xiang, Ke</mods:namePart>
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<mods:namePart id="D084DB68A52EF081101B718E373433E0">Pan, Jinbin</mods:namePart>
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<mods:namePart id="FA053D468B7560B610EBF346ECC50BCB">Yu, Jiaojiao</mods:namePart>
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<mods:namePart id="EA12801A202AAF3B57ABFE6288AD6EBD">Xiao, Lehui</mods:namePart>
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<mods:namePart id="8B3BD7ADD3478AA8D7881F8D516AD1EF">Sun, Shao-Kai</mods:namePart>
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<mods:date id="1BBD1B3DDBD076B74217607EC4322DA9">2022</mods:date>
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<mods:number id="1B9846FE0B8D8287E9CCD9FB2E045D37">2022-11-22</mods:number>
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<treatment id="561C87BEC160FFC880EF69DD5971FA56" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56" httpUri="http://treatment.plazi.org/id/561C87BEC160FFC880EF69DD5971FA56" lastPageId="5" lastPageNumber="144" pageId="3" pageNumber="142">
<subSubSection id="96AF6523C160FFCE80EF69DD5827FC8E" pageId="3" pageNumber="142" type="nomenclature">
<paragraph id="DE0A36A8C160FFCE80EF69DD5812FD56" blockId="3.[167,642,695,791]" box="[251,558,695,719]" pageId="3" pageNumber="142">
<heading id="854281C4C160FFCE80EF69DD5812FD56" box="[251,558,695,719]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="142" reason="4">
<taxonomicName id="19B54D2BC160FFCE80EF69DD5BD1FD56" ID-CoL="C6JWN" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[251,493,695,719]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="cavernarius" status="sp. nov.">
<emphasis id="ECC1EABAC160FFCE80EF69DD5BD1FD56" bold="true" box="[251,493,695,719]" italics="true" pageId="3" pageNumber="142">Callobius cavernarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C160FFCE81E069D25812FD56" box="[500,558,696,719]" pageId="3" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="DE0A36A8C160FFCE80B369B05827FC8E" blockId="3.[167,642,695,791]" pageId="3" pageNumber="142">
[Japanese name: Erabuhoraana-gakejigumo] (
<figureCitation id="468E2A2DC160FFCE8100686B5BBCFC8F" box="[276,384,769,791]" captionStart="Fig" captionStartId="1.[163,196,1748,1770]" captionTargetBox="[161,1217,187,1714]" captionTargetId="figure-8@1.[157,1218,185,1715]" captionTargetPageId="1" captionText="Fig. 1. Comparison of male palps of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Male palp (left), dorsal view (A, C, E); same, ventral view (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760066" httpUri="https://zenodo.org/record/12760066/files/figure.png" pageId="3" pageNumber="142">Figs. 1A–B</figureCitation>
,
<figureCitation id="468E2A2DC160FFCE819D686B5BFEFC8F" box="[393,450,769,790]" captionStart="Fig" captionStartId="2.[163,196,966,988]" captionTargetBox="[159,1217,366,930]" captionTargetId="figure-49@2.[157,1218,365,933]" captionTargetPageId="2" captionText="Fig. 2. Comparison of female genital organs of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Epigyne, ventral view (A, C, E); female internal genitalia (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760068" httpUri="https://zenodo.org/record/12760068/files/figure.png" pageId="3" pageNumber="142">2A–B</figureCitation>
,
<figureCitation id="468E2A2DC160FFCE81DF686B5BD4FC8F" box="[459,488,769,790]" captionStart="Fig" captionStartId="2.[163,196,1748,1770]" captionTargetBox="[182,1216,1098,1712]" captionTargetId="figure-89@2.[157,1218,1097,1714]" captionTargetPageId="2" captionText="Fig. 3. Comparison of body appearance of female specimens. Callobius cavernarius sp. n. (A); C. shimojanai sp. n. (B); and C. breviprocessus (C). Scale bars=10mm." figureDoi="http://doi.org/10.5281/zenodo.12760070" httpUri="https://zenodo.org/record/12760070/files/figure.png" pageId="3" pageNumber="142">3A</figureCitation>
,
<figureCitation id="468E2A2DC160FFCE81E6686B5BC3FC8F" box="[498,511,769,790]" captionStart="Fig" captionStartId="3.[729,762,962,984]" captionTargetBox="[744,1210,235,922]" captionTargetId="figure-450@3.[713,1230,213,928]" captionTargetPageId="3" captionText="Fig. 4. Callobius cavernarius sp. n.; one specimen of female paratypes. Scale bar=5mm." figureDoi="http://doi.org/10.5281/zenodo.12760074" httpUri="https://zenodo.org/record/12760074/files/figure.png" pageId="3" pageNumber="142">4</figureCitation>
,
<figureCitation id="468E2A2DC160FFCE821C686B5828FC8E" box="[520,532,769,791]" captionStart="Fig" captionStartId="4.[163,196,1304,1326]" captionTargetBox="[158,1215,214,1271]" captionTargetId="figure-159@4.[157,1218,213,1271]" captionTargetPageId="4" captionText="Fig. 5. Copulatory organs of Callobius cavernarius sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760076" httpUri="https://zenodo.org/record/12760076/files/figure.png" pageId="3" pageNumber="142">5</figureCitation>
)
</paragraph>
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<uri id="AA243AAAC160FFCE8098685B5B06FCFD" pageId="3" pageNumber="142">
urn:lsid:zoobank.org:act:
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<emphasis id="ECC1EABAC160FFCE80B368E75B7DFC3C" box="[167,321,909,933]" italics="true" pageId="3" pageNumber="142">Type material</emphasis>
.
<materialsCitation id="6EDD3CF5C160FFCE817768E75AE6FB94" collectingDate="2004-04-28" collectionCode="T" collectorName="Tamura" country="Japan" location="Okinoerabu Island" municipality="Ginsuido Cave" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<typeStatus id="010E880AC160FFCE817768E75BFAFC3C" box="[355,454,909,933]" pageId="3" pageNumber="142" type="holotype">Holotype</typeStatus>
ˁ,
<collectingCountry id="A6A27638C160FFCE821168E75866FC3C" box="[517,602,909,933]" name="Japan" pageId="3" pageNumber="142">JAPAN</collectingCountry>
:
<collectingMunicipality id="3E6EACD2C160FFCE827868E75B29FC5E" pageId="3" pageNumber="142">Ginsuido Cave</collectingMunicipality>
,
<location id="DB6A6073C160FFCE812468C55835FC5E" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE812468C55835FC5E" box="[304,521,943,967]" country="Japan" municipality="Ginsuido Cave" name="Okinoerabu Island" pageId="3" pageNumber="142" stateProvince="Kagoshima">Okinoerabu Island</location>
,
<collectingRegion id="1C71F84AC160FFCE823768C55AFEFC73" country="Japan" name="Kagoshima" pageId="3" pageNumber="142">Kagoshima Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE80C468B85B52FC73" box="[208,366,978,1002]" pageId="3" pageNumber="142" value="2004-04-28">
<collectingDate id="BA4FE980C160FFCE80C468B85B52FC73" box="[208,366,978,1002]" pageId="3" pageNumber="142" value="2004-04-28">28 April 2004</collectingDate>
</date>
,
<collectionCode id="B8A4AE6DC160FFCE816868B85BB2FC70" box="[380,398,978,1001]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="142">T</collectionCode>
.
<collectorName id="7340537EC160FFCE818F68B85BD0FC70" box="[411,492,978,1001]" pageId="3" pageNumber="142">Tamura</collectorName>
leg. (NSMT-Ar 22142)
</materialsCitation>
.
<materialsCitation id="6EDD3CF5C160FFCE80FD689F5BEBFB45" collectingDate="2004-04-30" collectingDateMax="2021-12-19" collectingDateMin="2004-04-30" collectionCode="T" collectorName="Tamura &amp; Ogimi &amp; Ogimi-son &amp; Kunigami-gun &amp; Y. Suzuki" country="Japan" county="Suirendo cave" location="Okinawajima Island" municipality="Okinoerabu Island" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
<typeStatus id="010E880AC160FFCE80FD689F5B69FB94" box="[233,341,1013,1037]" pageId="3" pageNumber="142" type="paratype">Paratypes</typeStatus>
: 2˂, same data as the holotype (NSMT-Ar 22143); 1ˁ,
<collectingCounty id="376B4E24C160FFCE81E16F7D58A4FBB6" box="[501,664,1047,1071]" pageId="3" pageNumber="142">Suirendo cave</collectingCounty>
,
<collectingMunicipality id="3E6EACD2C160FFCE80986F505B60FBCB" box="[140,348,1082,1106]" pageId="3" pageNumber="142">Okinoerabu Island</collectingMunicipality>
,
<collectingRegion id="1C71F84AC160FFCE817A6F505810FBCB" box="[366,556,1082,1106]" country="Japan" name="Kagoshima" pageId="3" pageNumber="142">Kagoshima Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE82296F505AF8FBEC" pageId="3" pageNumber="142" value="2004-04-30">
<collectingDate id="BA4FE980C160FFCE82296F505AF8FBEC" pageId="3" pageNumber="142" value="2004-04-30">30 April 2004</collectingDate>
</date>
,
<collectionCode id="B8A4AE6DC160FFCE80C06F375ADAFBED" box="[212,230,1117,1140]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="142">T</collectionCode>
.
<collectorName id="7340537EC160FFCE80E26F375B7BFBED" box="[246,327,1117,1140]" pageId="3" pageNumber="142">Tamura</collectorName>
leg. (NSMT-Ar 22144); 1 ˂,
<collectorName id="7340537EC160FFCE80986F155AEFFB0E" box="[140,211,1151,1175]" pageId="3" pageNumber="142">Ogimi</collectorName>
,
<collectorName id="7340537EC160FFCE80F46F155B69FB0E" box="[224,341,1151,1175]" pageId="3" pageNumber="142">Ogimi-son</collectorName>
,
<collectorName id="7340537EC160FFCE81766F15583CFB0E" box="[354,512,1151,1175]" pageId="3" pageNumber="142">Kunigami-gun</collectorName>
,
<location id="DB6A6073C160FFCE82196F155AECFB23" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE82196F155AECFB23" county="Suirendo cave" municipality="Okinoerabu Island" name="Okinawajima Island" pageId="3" pageNumber="142" stateProvince="Kagoshima">Okinawajima Island</location>
,
<collectingRegion id="1C71F84AC160FFCE80F66FC85BBBFB23" box="[226,391,1186,1210]" country="Japan" name="Okinawa" pageId="3" pageNumber="142">Okinawa Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE818C6FC85849FB23" box="[408,629,1186,1210]" pageId="3" pageNumber="142" value="2021-12-19">
<collectingDate id="BA4FE980C160FFCE818C6FC85849FB23" box="[408,629,1186,1210]" pageId="3" pageNumber="142" value="2021-12-19">19 December 2021</collectingDate>
</date>
,
<collectorName id="7340537EC160FFCE82936FC85AEAFB44" pageId="3" pageNumber="142">Y. Suzuki</collectorName>
leg. (NSMT- Ar 22145)
</materialsCitation>
</paragraph>
<caption id="8ACA6620C160FFCE82CD68A85E54FC6D" ID-DOI="http://doi.org/10.5281/zenodo.12760074" ID-Zenodo-Dep="12760074" httpUri="https://zenodo.org/record/12760074/files/figure.png" pageId="3" pageNumber="142" startId="3.[729,762,962,984]" targetBox="[744,1210,235,922]" targetPageId="3" targetType="figure">
<paragraph id="DE0A36A8C160FFCE82CD68A85E54FC6D" blockId="3.[729,1214,962,1013]" pageId="3" pageNumber="142">
Fig. 4.
<taxonomicName id="19B54D2BC160FFCE832468A859C7FC4E" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[816,1019,962,983]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="cavernarius" status="sp. nov.">
<emphasis id="ECC1EABAC160FFCE832468A859C7FC4E" box="[816,1019,962,983]" italics="true" pageId="3" pageNumber="142">Callobius cavernarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C160FFCE841668A85E0AFC4E" box="[1026,1078,962,983]" pageId="3" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
; one specimen of female paratypes. Scale bar=5 mm.
</paragraph>
</caption>
<paragraph id="DE0A36A8C160FFCE80B36F8D5BEBF9AE" blockId="3.[140,670,906,1799]" pageId="3" pageNumber="142">
<emphasis id="ECC1EABAC160FFCE80B36F8D5BC9FB66" box="[167,501,1255,1279]" italics="true" pageId="3" pageNumber="142">Other materials examined</emphasis>
.
<materialsCitation id="6EDD3CF5C160FFCE82026F8D5810FADC" collectingDate="2021-12-09" collectorName="Y. Suzuki" location="Okinoerabu Island" municipality="Shoryudo Cave" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Kagoshima">
1 ˁ, 6 ˂,
<collectingMunicipality id="3E6EACD2C160FFCE80986E605B04FABB" box="[140,312,1290,1314]" pageId="3" pageNumber="142">Shoryudo Cave</collectingMunicipality>
,
<location id="DB6A6073C160FFCE81536E605828FABB" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE81536E605828FABB" box="[327,532,1290,1314]" municipality="Shoryudo Cave" name="Okinoerabu Island" pageId="3" pageNumber="142" stateProvince="Kagoshima">Okinoerabu Island</location>
,
<collectingRegion id="1C71F84AC160FFCE82376E605AFEFADC" country="Japan" name="Kagoshima" pageId="3" pageNumber="142">Kagoshima Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE80D86E475BB1FADC" box="[204,397,1325,1349]" pageId="3" pageNumber="142" value="2021-12-09">
<collectingDate id="BA4FE980C160FFCE80D86E475BB1FADC" box="[204,397,1325,1349]" pageId="3" pageNumber="142" value="2021-12-09">9 December 2021</collectingDate>
</date>
,
<collectorName id="7340537EC160FFCE81826E475BC2FADC" box="[406,510,1325,1349]" pageId="3" pageNumber="142">Y. Suzuki</collectorName>
leg.
</materialsCitation>
<materialsCitation id="6EDD3CF5C160FFCE82266E475866FA13" collectingDate="1982-03-06" collectorName="M. Shimojana" location="Okinoerabu Island" municipality="Shoryudo Cave" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Kagoshima">
2 ˁ, 2 ˂,
<collectingMunicipality id="3E6EACD2C160FFCE80986E255B04FAFE" box="[140,312,1359,1383]" pageId="3" pageNumber="142">Shoryudo Cave</collectingMunicipality>
,
<location id="DB6A6073C160FFCE81536E255828FAFE" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE81536E255828FAFE" box="[327,532,1359,1383]" municipality="Shoryudo Cave" name="Okinoerabu Island" pageId="3" pageNumber="142" stateProvince="Kagoshima">Okinoerabu Island</location>
,
<collectingRegion id="1C71F84AC160FFCE82376E255AFEFA13" country="Japan" name="Kagoshima" pageId="3" pageNumber="142">Kagoshima Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE80C76E185B45FA13" box="[211,377,1394,1418]" pageId="3" pageNumber="142" value="1982-03-06">
<collectingDate id="BA4FE980C160FFCE80C76E185B45FA13" box="[211,377,1394,1418]" pageId="3" pageNumber="142" value="1982-03-06">6 March 1982</collectingDate>
</date>
,
<collectorName id="7340537EC160FFCE819E6E185814FA13" box="[394,552,1394,1418]" pageId="3" pageNumber="142">M. Shimojana</collectorName>
leg
</materialsCitation>
:
<materialsCitation id="6EDD3CF5C160FFCE827F6E185A88FA6B" collectingDate="1982-03-06" collectorName="M. Shimojana" location="Okinoerabu Island" municipality="Suirendo Cave" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Kagoshima">
5 ˁ, 3˂,
<collectingMunicipality id="3E6EACD2C160FFCE80C96EFF5BA9FA34" box="[221,405,1429,1453]" pageId="3" pageNumber="142">Suirendo Cave</collectingMunicipality>
,
<location id="DB6A6073C160FFCE81AC6EFF58A5FA34" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE81AC6EFF58A5FA34" box="[440,665,1429,1453]" municipality="Suirendo Cave" name="Okinoerabu Island" pageId="3" pageNumber="142" stateProvince="Kagoshima">Okinoerabu Island</location>
,
<collectingRegion id="1C71F84AC160FFCE80986EDD5B7AFA56" box="[140,326,1463,1487]" country="Japan" name="Kagoshima" pageId="3" pageNumber="142">Kagoshima Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE81406EDD5BC8FA56" box="[340,500,1463,1487]" pageId="3" pageNumber="142" value="1982-03-06">
<collectingDate id="BA4FE980C160FFCE81406EDD5BC8FA56" box="[340,500,1463,1487]" pageId="3" pageNumber="142" value="1982-03-06">6 March 1982</collectingDate>
</date>
,
<collectorName id="7340537EC160FFCE82166ED258A1FA56" box="[514,669,1463,1487]" pageId="3" pageNumber="142">M. Shimojana</collectorName>
leg.
</materialsCitation>
<materialsCitation id="6EDD3CF5C160FFCE80D76EB05BEBF9AE" collectingDate="2021-12-19" collectorName="Y. Suzuki" county="Ogimi" location="Kunigami-gun" municipality="Ogimi-son" pageId="3" pageNumber="142" specimenCount="1" stateProvince="Okinawa">
2ˁ, 3˂,
<collectingCounty id="376B4E24C160FFCE81526EB05BB1FA6B" box="[326,397,1498,1522]" pageId="3" pageNumber="142">Ogimi</collectingCounty>
,
<collectingMunicipality id="3E6EACD2C160FFCE81B56EB0582AFA6B" box="[417,534,1498,1522]" pageId="3" pageNumber="142">Ogimi-son</collectingMunicipality>
,
<location id="DB6A6073C160FFCE823D6EB05A8BF98C" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE823D6EB05A8BF98C" county="Ogimi" municipality="Ogimi-son" name="Kunigami-gun" pageId="3" pageNumber="142" stateProvince="Okinawa">Kunigami-gun</location>
,
<location id="DB6A6073C160FFCE80D86E975B8DF98C" LSID="urn:lsid:plazi:treatment:561C87BEC160FFC880EF69DD5971FA56:DB6A6073C160FFCE80D86E975B8DF98C" box="[204,433,1533,1557]" county="Ogimi" municipality="Ogimi-son" name="Okinawajima Island" pageId="3" pageNumber="142" stateProvince="Okinawa">Okinawajima Island</location>
,
<collectingRegion id="1C71F84AC160FFCE81D26E975852F98C" box="[454,622,1533,1557]" country="Japan" name="Okinawa" pageId="3" pageNumber="142">Okinawa Pref.</collectingRegion>
,
<date id="AA0B1068C160FFCE82966E975B04F9AE" pageId="3" pageNumber="142" value="2021-12-19">
<collectingDate id="BA4FE980C160FFCE82966E975B04F9AE" pageId="3" pageNumber="142" value="2021-12-19">19 December 2021</collectingDate>
</date>
,
<collectorName id="7340537EC160FFCE81566D4A5B95F9AE" box="[322,425,1567,1591]" pageId="3" pageNumber="142">Y. Suzuki</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="DE0A36A8C160FFCE80B26D285B5CF958" blockId="3.[140,670,906,1799]" pageId="3" pageNumber="142">
<emphasis id="ECC1EABAC160FFCE80B26D285B24F9C3" box="[166,280,1602,1626]" italics="true" pageId="3" pageNumber="142">Etymology</emphasis>
. The specific name is derived from the Latin adjective
<taxonomicName id="19B54D2BC160FFCE81826D0F5829F9E5" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[406,533,1637,1660]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="cavernarius">
<emphasis id="ECC1EABAC160FFCE81826D0F5829F9E5" box="[406,533,1637,1660]" italics="true" pageId="3" pageNumber="142">cavernarius</emphasis>
</taxonomicName>
lconcerning caves,z referring to the discovery of the
<typeStatus id="010E880AC160FFCE827B6DE258A1F939" box="[623,669,1672,1696]" pageId="3" pageNumber="142">type</typeStatus>
specimens in caves.
</paragraph>
<paragraph id="DE0A36A8C160FFCE80B36DA75EB2FA6B" blockId="3.[140,670,906,1799]" lastBlockId="3.[707,1237,1047,1799]" pageId="3" pageNumber="142">
<emphasis id="ECC1EABAC160FFCE80B36DA75B2FF97D" box="[167,275,1741,1764]" italics="true" pageId="3" pageNumber="142">Diagnosis</emphasis>
. Male specimens of
<taxonomicName id="19B54D2BC160FFCE82216DA75B37F89E" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="cavernarius" status="sp. nov.">
<emphasis id="ECC1EABAC160FFCE82216DA75B37F89E" italics="true" pageId="3" pageNumber="142">Callobius cavernarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C160FFCE810F6D9A5B62F89E" box="[283,350,1776,1799]" pageId="3" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from other congeners by the narrow cymbium, the long tibia and the trapezoidal MP with an extremely small projection in the palp. Female can be distinguished by the triangular lateral lobes located distant from each other in the epigyne and the semicircular spermathecae of the internal genitalia. As an example, the differences from
<taxonomicName id="19B54D2BC160FFCE83116E6059F5FABB" box="[773,969,1290,1314]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="breviprocessus">
<emphasis id="ECC1EABAC160FFCE83116E6059F5FABB" box="[773,969,1290,1314]" italics="true" pageId="3" pageNumber="142">C. breviprocessus</emphasis>
</taxonomicName>
that lives in the most neighboring areas are shown (
<figureCitation id="468E2A2DC160FFCE84116E475E73FADC" box="[1029,1103,1325,1349]" captionStart="Fig" captionStartId="1.[163,196,1748,1770]" captionTargetBox="[161,1217,187,1714]" captionTargetId="figure-8@1.[157,1218,185,1715]" captionTargetPageId="1" captionText="Fig. 1. Comparison of male palps of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Male palp (left), dorsal view (A, C, E); same, ventral view (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760066" httpUri="https://zenodo.org/record/12760066/files/figure.png" pageId="3" pageNumber="142">Figs. 1</figureCitation>
,
<figureCitation id="468E2A2DC160FFCE84496E475E51FADC" box="[1117,1133,1325,1349]" captionStart="Fig" captionStartId="2.[163,196,966,988]" captionTargetBox="[159,1217,366,930]" captionTargetId="figure-49@2.[157,1218,365,933]" captionTargetPageId="2" captionText="Fig. 2. Comparison of female genital organs of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Epigyne, ventral view (A, C, E); female internal genitalia (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760068" httpUri="https://zenodo.org/record/12760068/files/figure.png" pageId="3" pageNumber="142">2</figureCitation>
). For differences from the other two species,
<taxonomicName id="19B54D2BC160FFCE844C6E3A5930FA13" authorityName="Okumura" authorityYear="2010" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="yakushimensis">
<emphasis id="ECC1EABAC160FFCE844C6E3A5930FA13" italics="true" pageId="3" pageNumber="142">C. yakushimensis</emphasis>
</taxonomicName>
and
<taxonomicName id="19B54D2BC160FFCE83546E1859E1FA13" box="[832,989,1394,1418]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="amamiensis">
<emphasis id="ECC1EABAC160FFCE83546E1859E1FA13" box="[832,989,1394,1418]" italics="true" pageId="3" pageNumber="142">C. amamiensis</emphasis>
</taxonomicName>
in the Ryukyu Islands, refer to
<bibRefCitation id="BA244B59C160FFCE83096EFF5E31FA34" author="Okumura, K. &amp; Y. Suzuki &amp; R. Serita" box="[797,1037,1429,1453]" pageId="3" pageNumber="142" pagination="596 - 601" refId="ref3254" refString="Okumura, K., Y. Suzuki and R. Serita 2020. Two new species of spiders of the families Agelenidae and Amaurobiidae from Okinawa Island, Japan. Arachnology 18: 596 - 601." type="journal article" year="2020">
Okumura
<emphasis id="ECC1EABAC160FFCE839F6EFF59F8FA34" box="[907,964,1429,1453]" italics="true" pageId="3" pageNumber="142">et al.</emphasis>
, 2020
</bibRefCitation>
. The body size of the species is also much larger than the other species living on Okinawajima Island (
<figureCitation id="468E2A2DC160FFCE842B6EB05EBCFA6B" box="[1087,1152,1498,1522]" captionStart="Fig" captionStartId="2.[163,196,1748,1770]" captionTargetBox="[182,1216,1098,1712]" captionTargetId="figure-89@2.[157,1218,1097,1714]" captionTargetPageId="2" captionText="Fig. 3. Comparison of body appearance of female specimens. Callobius cavernarius sp. n. (A); C. shimojanai sp. n. (B); and C. breviprocessus (C). Scale bars=10mm." figureDoi="http://doi.org/10.5281/zenodo.12760070" httpUri="https://zenodo.org/record/12760070/files/figure.png" pageId="3" pageNumber="142">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="DE0A36A8C160FFC982C96E975BD4FA6B" blockId="3.[707,1237,1047,1799]" lastBlockId="4.[140,670,1428,1800]" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142">
<emphasis id="ECC1EABAC160FFCE82C96E975967F98D" box="[733,859,1533,1556]" italics="true" pageId="3" pageNumber="142">Description</emphasis>
. Male (
<typeStatus id="010E880AC160FFCE83D86E975E0CF98C" box="[972,1072,1533,1557]" pageId="3" pageNumber="142" type="holotype">holotype</typeStatus>
). Total length 9.8, carapace 5.5 long, 3.8 wide; abdomen 4.3 long, 2.7 wide; sternum 2.3 long, 1.9 wide. Eye sizes and interdistances AME 0.19, ALE 0.26, PME 0.21, PLE 0.24; AME-AME 0.09, AME- ALE 0.11, PME-PME 0.21, PME-PLE 0.30, AME-PME 0.31, ALE-PLE 0.10. MOA; anterior width 0.47, posterior width 0.63, length 0.71. Leg measurements: I: 27.7 (7.1, 9.2, 7.5, 3.9); II: 19.2 (5.3, 6.6, 4.8, 2.5); III: 16.5 (4.8, 5.4, 4.3, 2.0); IV: 21.8 (6.1, 7.0, 6.0, 2.7).
</paragraph>
<caption id="8ACA6620C167FFC980B76E725816FAF1" ID-DOI="http://doi.org/10.5281/zenodo.12760076" ID-Zenodo-Dep="12760076" httpUri="https://zenodo.org/record/12760076/files/figure.png" pageId="4" pageNumber="143" startId="4.[163,196,1304,1326]" targetBox="[158,1215,214,1271]" targetPageId="4" targetType="figure">
<paragraph id="DE0A36A8C167FFC980B76E725816FAF1" blockId="4.[163,1214,1304,1385]" pageId="4" pageNumber="143">
Fig. 5. Copulatory organs of
<taxonomicName id="19B54D2BC167FFC981D66E7258B1FAB7" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[450,653,1304,1326]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="cavernarius" status="sp. nov.">
<emphasis id="ECC1EABAC167FFC981D66E7258B1FAB7" box="[450,653,1304,1326]" italics="true" pageId="4" pageNumber="143">Callobius cavernarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C167FFC982806E7358FAFAB7" box="[660,710,1305,1326]" pageId="4" pageNumber="143" rank="species">sp. n.</taxonomicNameLabel>
holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm.
</paragraph>
</caption>
<paragraph id="DE0A36A8C167FFC980B36E9758C8FA34" blockId="4.[140,670,1428,1800]" lastBlockId="4.[707,1236,1429,1799]" pageId="4" pageNumber="143">
Palp (
<figureCitation id="468E2A2DC167FFC980F86E975B4DF98C" box="[236,369,1533,1557]" captionStart="Fig" captionStartId="1.[163,196,1748,1770]" captionTargetBox="[161,1217,187,1714]" captionTargetId="figure-8@1.[157,1218,185,1715]" captionTargetPageId="1" captionText="Fig. 1. Comparison of male palps of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Male palp (left), dorsal view (A, C, E); same, ventral view (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760066" httpUri="https://zenodo.org/record/12760066/files/figure.png" pageId="4" pageNumber="143">Figs. 1A–B</figureCitation>
,
<figureCitation id="468E2A2DC167FFC981916E965BF5F98D" box="[389,457,1532,1556]" captionStart="Fig" captionStartId="4.[163,196,1304,1326]" captionTargetBox="[158,1215,214,1271]" captionTargetId="figure-159@4.[157,1218,213,1271]" captionTargetPageId="4" captionText="Fig. 5. Copulatory organs of Callobius cavernarius sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760076" httpUri="https://zenodo.org/record/12760076/files/figure.png" pageId="4" pageNumber="143">5A–C</figureCitation>
): three projections present in the lateral to dorsal portion of long tibia; RTA thick and has a small process in the tip, DP slightly curved and pointed in the tip, MP trapezoid with an extremely small projection, cymbial excavation clear; CO thin and semitransparent, median apophysis large and claw-shaped, embolus short and conglutinates with the tegulum.
</paragraph>
<paragraph id="DE0A36A8C167FFC982C96EDD594DFA6B" blockId="4.[707,1236,1429,1799]" pageId="4" pageNumber="143">Chelicerae: promargin with 5 teeth, retromargin with 5 teeth.</paragraph>
<paragraph id="DE0A36A8C167FFC982C96E9759E1F89E" blockId="4.[707,1236,1429,1799]" pageId="4" pageNumber="143">Colouration: carapace yellowish brown with indistinct radial flecks, dorsum of abdomen blackish brown but grayish brown only in the anteromedian part, and venter blackish brown with white lines in both lateral sides, sternum yellowish brown, chelicerae blackish brown, maxillae and labium brown, legs yellowish brown without ring flecks.</paragraph>
<caption id="8ACA6620C166FFC880B768375B5FFC52" ID-DOI="http://doi.org/10.5281/zenodo.12760078" ID-Zenodo-Dep="12760078" httpUri="https://zenodo.org/record/12760078/files/figure.png" pageId="5" pageNumber="144" startId="5.[163,196,861,883]" targetBox="[147,662,214,827]" targetPageId="5" targetType="figure">
<paragraph id="DE0A36A8C166FFC880B768375B5FFC52" blockId="5.[163,647,861,971]" pageId="5" pageNumber="144">
Fig. 6. Distribution records of
<taxonomicName id="19B54D2BC166FFC881DB68375AD6FC09" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="cavernarius" status="sp. nov.">
<emphasis id="ECC1EABAC166FFC881DB68375AD6FC09" italics="true" pageId="5" pageNumber="144">Callobius cavernarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C166FFC880E168115B17FC09" box="[245,299,891,912]" pageId="5" pageNumber="144" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="19B54D2BC166FFC8817568115BDAFC16" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[353,486,890,912]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="shimojanai" status="sp. n.">
<emphasis id="ECC1EABAC166FFC8817568115BDAFC16" box="[353,486,890,912]" italics="true" pageId="5" pageNumber="144">C. shimojanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C166FFC881E46811581BFC09" box="[496,551,891,912]" pageId="5" pageNumber="144" rank="species">sp. n.</taxonomicNameLabel>
including three closely related species living on the Ryukyu Islands.
</paragraph>
</caption>
<caption id="8ACA6620C166FFC882CD68DA5E54FC7A" ID-DOI="http://doi.org/10.5281/zenodo.12760080" ID-Zenodo-Dep="12760080" httpUri="https://zenodo.org/record/12760080/files/figure.png" pageId="5" pageNumber="144" startId="5.[729,762,944,966]" targetBox="[714,1230,215,910]" targetPageId="5" targetType="figure">
<paragraph id="DE0A36A8C166FFC882CD68DA5E54FC7A" blockId="5.[729,1214,944,996]" pageId="5" pageNumber="144">
Fig. 7.
<taxonomicName id="19B54D2BC166FFC8832468DA59CAFC5C" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[816,1014,944,965]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="shimojanai" status="sp. nov.">
<emphasis id="ECC1EABAC166FFC8832468DA59CAFC5C" box="[816,1014,944,965]" italics="true" pageId="5" pageNumber="144">Callobius shimojanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C166FFC883E968DB5E0FFC5F" box="[1021,1075,945,966]" pageId="5" pageNumber="144" rank="species">sp. n.</taxonomicNameLabel>
; one specimen of female paratypes. Scale bar=2 mm.
</paragraph>
</caption>
<paragraph id="DE0A36A8C166FFC880B3689F5BD4FAFE" blockId="5.[140,670,1013,1799]" pageId="5" pageNumber="144">
Female (one of
<typeStatus id="010E880AC166FFC8814E689F5BF9FB94" box="[346,453,1013,1037]" pageId="5" pageNumber="144" type="paratype">paratypes</typeStatus>
). Total length 10.4, carapace 3.9 long, 2.4 wide; abdomen 6.5 long, 4.7 wide; sternum 1.7 long, 1.6 wide. Eye sizes; AME 0.13, ALE 0.20, PME 0.16, PLE 0.18. Distances between eyes; AME-AME 0.11, AME- ALE 0.10, PME-PME 0.15, PME-PLE 0.28, AME-PME 0.21, ALE-PLE 0.08. MOA; anterior width 0.37, posterior width 0.47, length 0.50. Leg measurements: I: 14.3 (4.0, 4.9, 3.4, 2.0); II: 11.5 (3.3, 3.9, 2.7, 1.6); III: 10.2 (3.0, 3.4, 2.5, 1.3); IV: 13.0 (3.8, 4.6, 3.4, 1.2).
</paragraph>
<paragraph id="DE0A36A8C166FFC880B36E185ACBF9C3" blockId="5.[140,670,1013,1799]" pageId="5" pageNumber="144">
Epigyne and internal genitalia (
<figureCitation id="468E2A2DC166FFC882076E1858A5FA13" box="[531,665,1394,1418]" captionStart="Fig" captionStartId="2.[163,196,966,988]" captionTargetBox="[159,1217,366,930]" captionTargetId="figure-49@2.[157,1218,365,933]" captionTargetPageId="2" captionText="Fig. 2. Comparison of female genital organs of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Epigyne, ventral view (A, C, E); female internal genitalia (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760068" httpUri="https://zenodo.org/record/12760068/files/figure.png" pageId="5" pageNumber="144">Figs. 2A–B</figureCitation>
,
<figureCitation id="468E2A2DC166FFC880986EFE5AF3FA35" box="[140,207,1428,1452]" captionStart="Fig" captionStartId="4.[163,196,1304,1326]" captionTargetBox="[158,1215,214,1271]" captionTargetId="figure-159@4.[157,1218,213,1271]" captionTargetPageId="4" captionText="Fig. 5. Copulatory organs of Callobius cavernarius sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760076" httpUri="https://zenodo.org/record/12760076/files/figure.png" pageId="5" pageNumber="144">5D–E</figureCitation>
): lateral lobes almost semicircular, swollen and distant from each other, median lobe almost circular from the posterior view, ectal lobe and posterior lobe absent. SP semicircular and distant from each other, fertilization ducts small and indistinct.
</paragraph>
<paragraph id="DE0A36A8C166FFC880B36D0F5BDEF95B" blockId="5.[140,670,1013,1799]" pageId="5" pageNumber="144">Chelicerae: promargin with 6 teeth on the left, and 5 on the right, retromargin with 6 teeth on the left, and 5 teeth on the right.</paragraph>
<paragraph id="DE0A36A8C166FFC880B36DA75ACDF89E" blockId="5.[140,670,1013,1799]" pageId="5" pageNumber="144">
Colouration: almost same as that of the male
<typeStatus id="010E880AC166FFC880986D855AD1F89E" box="[140,237,1775,1799]" pageId="5" pageNumber="144" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="DE0A36A8C166FFC882C96F7D5966FBCB" blockId="5.[707,1236,1047,1487]" pageId="5" pageNumber="144">
Distribution: Okinoerabu Island, Okinawajima Island (
<figureCitation id="468E2A2DC166FFC883076F50596EFBCB" box="[787,850,1082,1106]" captionStart="Fig" captionStartId="5.[163,196,861,883]" captionTargetBox="[147,662,214,827]" captionTargetId="figure-299@5.[146,663,213,828]" captionTargetPageId="5" captionText="Fig. 6. Distribution records of Callobius cavernarius sp. n. and C. shimojanai sp. n. including three closely related species living on the Ryukyu Islands." figureDoi="http://doi.org/10.5281/zenodo.12760078" httpUri="https://zenodo.org/record/12760078/files/figure.png" pageId="5" pageNumber="144">Fig. 6</figureCitation>
)
</paragraph>
<paragraph id="DE0A36A8C166FFC882C96F375971FA56" blockId="5.[707,1236,1047,1487]" pageId="5" pageNumber="144">
Remarks. Although the number of the marginal teeth in the female
<typeStatus id="010E880AC166FFC883C56FEA59C3FB01" box="[977,1023,1152,1176]" pageId="5" pageNumber="144">type</typeStatus>
specimen using for the description is different between the right che- licera and the left one, we considered this is a case of variation because the variation of the number of the marginal teeth can easily occurs (
<bibRefCitation id="BA244B59C166FFC882DE6E47598AFADC" author="Okumura, K. &amp; Y. Suzuki &amp; R. Serita" box="[714,950,1325,1349]" pageId="5" pageNumber="144" pagination="596 - 601" refId="ref3254" refString="Okumura, K., Y. Suzuki and R. Serita 2020. Two new species of spiders of the families Agelenidae and Amaurobiidae from Okinawa Island, Japan. Arachnology 18: 596 - 601." type="journal article" year="2020">
Okumura
<emphasis id="ECC1EABAC166FFC8832E6E47594FFADC" box="[826,883,1325,1349]" italics="true" pageId="5" pageNumber="144">et al.</emphasis>
2020
</bibRefCitation>
). The main habitat of this new species is in caves in Okinoerabu Island, but some specimens have also been collected from the ground in the mountainous areas on Okinawajima Island.
</paragraph>
</subSubSection>
</treatment>
</document>

234
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<emphasis id="ECC1EABAC166FFC883226D635E1DF9B8" bold="true" box="[822,1057,1545,1569]" italics="true" pageId="5" pageNumber="144">Callobius shimojanai</emphasis>
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<paragraph id="DE0A36A8C166FFC883516D365EA8F916" blockId="5.[771,1172,1628,1679]" pageId="5" pageNumber="144">
(
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<figureCitation id="468E2A2DC166FFC884166D365E22F9E8" box="[1026,1054,1628,1649]" captionStart="Fig" captionStartId="2.[163,196,1748,1770]" captionTargetBox="[182,1216,1098,1712]" captionTargetId="figure-89@2.[157,1218,1097,1714]" captionTargetPageId="2" captionText="Fig. 3. Comparison of body appearance of female specimens. Callobius cavernarius sp. n. (A); C. shimojanai sp. n. (B); and C. breviprocessus (C). Scale bars=10mm." figureDoi="http://doi.org/10.5281/zenodo.12760070" httpUri="https://zenodo.org/record/12760070/files/figure.png" pageId="5" pageNumber="144">3B</figureCitation>
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<figureCitation id="468E2A2DC166FFC8843C6D365E09F9E8" box="[1064,1077,1628,1649]" captionStart="Fig" captionStartId="5.[729,762,944,966]" captionTargetBox="[714,1230,215,910]" captionTargetId="figure-327@5.[713,1230,213,911]" captionTargetPageId="5" captionText="Fig. 7. Callobius shimojanai sp. n.; one specimen of female paratypes. Scale bar=2mm." figureDoi="http://doi.org/10.5281/zenodo.12760080" httpUri="https://zenodo.org/record/12760080/files/figure.png" pageId="5" pageNumber="144">7</figureCitation>
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<figureCitation id="468E2A2DC166FFC8842B6D365E76F9E8" box="[1087,1098,1628,1649]" captionStart="Fig" captionStartId="6.[163,196,1318,1340]" captionTargetBox="[158,1215,214,1284]" captionTargetId="figure-160@6.[157,1218,213,1285]" captionTargetPageId="6" captionText="Fig. 8. Copulatory organs of Callobius shimojanai sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760082" httpUri="https://zenodo.org/record/12760082/files/figure.png" pageId="5" pageNumber="144">8</figureCitation>
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ˁ,
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leg. (NSMT-Ar 22146)
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: 4 ˁ3 ˂, same data as the holotype (NSMT-Ar 22147); 1˂,
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,
<collectingRegion id="1C71F84AC165FFCB81666D75582CF9AE" box="[370,528,1567,1591]" country="Japan" name="Okinawa" pageId="6" pageNumber="145">Okinawa Pref.</collectingRegion>
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<date id="AA0B1068C165FFCB820F6D755AF8F9C3" pageId="6" pageNumber="145" value="2022-02-10">
<collectingDate id="BA4FE980C165FFCB820F6D755AF8F9C3" pageId="6" pageNumber="145" value="2022-02-10">10 February 2022</collectingDate>
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<collectorName id="7340537EC165FFCB80DA6D285B09F9C3" box="[206,309,1602,1626]" pageId="6" pageNumber="145">Y. Suzuki</collectorName>
leg. (NSMT-Ar 22148)
</materialsCitation>
</paragraph>
<caption id="8ACA6620C165FFCB80B76E4C5816FAEF" ID-DOI="http://doi.org/10.5281/zenodo.12760082" ID-Zenodo-Dep="12760082" httpUri="https://zenodo.org/record/12760082/files/figure.png" pageId="6" pageNumber="145" startId="6.[163,196,1318,1340]" targetBox="[158,1215,214,1284]" targetPageId="6" targetType="figure">
<paragraph id="DE0A36A8C165FFCB80B76E4C5816FAEF" blockId="6.[163,1214,1318,1399]" pageId="6" pageNumber="145">
Fig. 8. Copulatory organs of
<taxonomicName id="19B54D2BC165FFCB81D06E4C58B4FAA2" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[452,648,1318,1339]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="shimojanai" status="sp. nov.">
<emphasis id="ECC1EABAC165FFCB81D06E4C58B4FAA2" box="[452,648,1318,1339]" italics="true" pageId="6" pageNumber="145">Callobius shimojanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C165FFCB829B6E4D58FDFAA5" box="[655,705,1319,1340]" pageId="6" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm.
</paragraph>
</caption>
<paragraph id="DE0A36A8C165FFCB80B36D0F5BA8F95B" blockId="6.[140,670,1463,1800]" pageId="6" pageNumber="145">
<emphasis id="ECC1EABAC165FFCB80B36D0F5B25F9E4" box="[167,281,1637,1661]" italics="true" pageId="6" pageNumber="145">Etymology</emphasis>
. The specific name is dedicated to Dr. Matsuei Shimojana who collected the new species for the first time.
</paragraph>
<paragraph id="DE0A36A8C165FFCB80B36DA75914F97C" blockId="6.[140,670,1463,1800]" lastBlockId="6.[707,1236,1463,1799]" pageId="6" pageNumber="145">
<emphasis id="ECC1EABAC165FFCB80B36DA75B2FF97D" box="[167,275,1741,1764]" italics="true" pageId="6" pageNumber="145">Diagnosis</emphasis>
.
<taxonomicName id="19B54D2BC165FFCB81206DA7581CF97C" authorityName="Xiang &amp; Pan &amp; Yu &amp; Xiao &amp; Sun &amp; Cheng" authorityYear="2022" box="[308,544,1741,1765]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="shimojanai" status="sp. nov.">
<emphasis id="ECC1EABAC165FFCB81206DA7581CF97C" box="[308,544,1741,1765]" italics="true" pageId="6" pageNumber="145">Callobius shimojanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F257C1C165FFCB82386DA75857F97D" box="[556,619,1741,1764]" pageId="6" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from other congeners by the mutually crossing DP and MP in the tibia of male palp and the tip of the lateral lobes located close to each other in the epigyne. The new species clearly different from
<taxonomicName id="19B54D2BC165FFCB83A96D4A5EBCF9AE" box="[957,1152,1567,1591]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="breviprocessus">
<emphasis id="ECC1EABAC165FFCB83A96D4A5EBCF9AE" box="[957,1152,1567,1591]" italics="true" pageId="6" pageNumber="145">C. breviprocessus</emphasis>
</taxonomicName>
, which inhabit the same island, by the above characteristics (
<figureCitation id="468E2A2DC165FFCB82EF6D0F5974F9E4" box="[763,840,1637,1661]" captionStart="Fig" captionStartId="1.[163,196,1748,1770]" captionTargetBox="[161,1217,187,1714]" captionTargetId="figure-8@1.[157,1218,185,1715]" captionTargetPageId="1" captionText="Fig. 1. Comparison of male palps of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Male palp (left), dorsal view (A, C, E); same, ventral view (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760066" httpUri="https://zenodo.org/record/12760066/files/figure.png" pageId="6" pageNumber="145">Figs. 1</figureCitation>
,
<figureCitation id="468E2A2DC165FFCB834F6D0F5957F9E4" box="[859,875,1637,1661]" captionStart="Fig" captionStartId="2.[163,196,966,988]" captionTargetBox="[159,1217,366,930]" captionTargetId="figure-49@2.[157,1218,365,933]" captionTargetPageId="2" captionText="Fig. 2. Comparison of female genital organs of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Epigyne, ventral view (A, C, E); female internal genitalia (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760068" httpUri="https://zenodo.org/record/12760068/files/figure.png" pageId="6" pageNumber="145">2</figureCitation>
). For differences from the other two species,
<taxonomicName id="19B54D2BC165FFCB83466DE25E32F906" authorityName="Okumura" authorityYear="2010" box="[850,1038,1671,1695]" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="yakushimensis">
<emphasis id="ECC1EABAC165FFCB83466DE25E32F906" box="[850,1038,1671,1695]" italics="true" pageId="6" pageNumber="145">C. yakushimensis</emphasis>
</taxonomicName>
and
<taxonomicName id="19B54D2BC165FFCB84536DE258E3F95B" class="Arachnida" family="Amaurobiidae" genus="Callobius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="amamiensis">
<emphasis id="ECC1EABAC165FFCB84536DE258E3F95B" italics="true" pageId="6" pageNumber="145">C. amamiensis</emphasis>
</taxonomicName>
in the Ryukyu Islands, refer to
<bibRefCitation id="BA244B59C165FFCB84456DC05919F97C" author="Okumura, K. &amp; Y. Suzuki &amp; R. Serita" pageId="6" pageNumber="145" pagination="596 - 601" refId="ref3254" refString="Okumura, K., Y. Suzuki and R. Serita 2020. Two new species of spiders of the families Agelenidae and Amaurobiidae from Okinawa Island, Japan. Arachnology 18: 596 - 601." type="journal article" year="2020">
Okumura
<emphasis id="ECC1EABAC165FFCB84D56DC158E3F97C" italics="true" pageId="6" pageNumber="145">et al.</emphasis>
, 2020
</bibRefCitation>
.
</paragraph>
<paragraph id="DE0A36A8C165FFCA82C96D9A5B9EFDBE" blockId="6.[707,1236,1463,1799]" lastBlockId="7.[140,670,215,1799]" lastPageId="7" lastPageNumber="146" pageId="6" pageNumber="145">
<emphasis id="ECC1EABAC165FFCB82C96D9A5967F89E" box="[733,859,1776,1799]" italics="true" pageId="6" pageNumber="145">Description</emphasis>
. Male (
<typeStatus id="010E880AC165FFCB83D86D855E0CF89E" box="[972,1072,1775,1799]" pageId="6" pageNumber="145" type="holotype">holotype</typeStatus>
). Total length 4.6, carapace 2.5 long, 1.8 wide; abdomen 2.1 long, 1.5 wide; sternum 1.2 long, 1.0 wide. Eye sizes and interdistances AME 0.06, ALE 0.11, PME 0.13, PLE 0.13; AME-AME 0.04, AME- ALE 0.06, PME-PME 0.09, PME-PLE 0.11, AME-PME 0.23, ALE-PLE 0.05. MOA; anterior width 0.16, posterior width 0.35, length 0.42. Leg measurements: I: 11.3 (2.9, 3.8, 2.8, 1.8); II: 7.1 (2.1, 2.5, 1.5, 1.0); III: 6.0 (1.9, 2.0, 1.2, 0.9); IV: 7.9 (2.2, 2.6, 2.0, 1.1).
</paragraph>
<paragraph id="DE0A36A8C164FFCA80B369585B74FCC6" blockId="7.[140,670,215,1799]" pageId="7" pageNumber="146">
Palp (
<figureCitation id="468E2A2DC164FFCA80F869585B4DFDD3" box="[236,369,562,586]" captionStart="Fig" captionStartId="1.[163,196,1748,1770]" captionTargetBox="[161,1217,187,1714]" captionTargetId="figure-8@1.[157,1218,185,1715]" captionTargetPageId="1" captionText="Fig. 1. Comparison of male palps of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Male palp (left), dorsal view (A, C, E); same, ventral view (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760066" httpUri="https://zenodo.org/record/12760066/files/figure.png" pageId="7" pageNumber="146">Figs. 1C–D</figureCitation>
,
<figureCitation id="468E2A2DC164FFCA819169585BF5FDD3" box="[389,457,562,586]" captionStart="Fig" captionStartId="6.[163,196,1318,1340]" captionTargetBox="[158,1215,214,1284]" captionTargetId="figure-160@6.[157,1218,213,1285]" captionTargetPageId="6" captionText="Fig. 8. Copulatory organs of Callobius shimojanai sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760082" httpUri="https://zenodo.org/record/12760082/files/figure.png" pageId="7" pageNumber="146">8A–C</figureCitation>
): three projections present in the lateral to dorsal portion of tibia; RTA thick and has a minute process in the tip, DP somewhat small and pointed in the tip, MP long, the two processes mentioned above cross each other. cymbial excavation clear; CO thin and semitransparent, median apophysis thick and claw-shaped, embolus short and conglutinates with the tegulum.
</paragraph>
<paragraph id="DE0A36A8C164FFCA80B368005B06FC3C" blockId="7.[140,670,215,1799]" pageId="7" pageNumber="146">Chelicerae: promargin with 6 teeth, retromargin with 6 teeth.</paragraph>
<paragraph id="DE0A36A8C164FFCA80B368C55BC8FB0E" blockId="7.[140,670,215,1799]" pageId="7" pageNumber="146">Colouration: carapace yellowish brown with indistinct radial flecks, dorsum of abdomen grayish brown with five pairs of white markings, and venter with two pairs of white lines in both lateral sides, sternum grayish brown, chelicerae, maxillae and labium blackish brown, legs yellowish brown without ring flecks.</paragraph>
<paragraph id="DE0A36A8C164FFCA80B36FC85B9EF98C" blockId="7.[140,670,215,1799]" pageId="7" pageNumber="146">
Female (one of
<typeStatus id="010E880AC164FFCA81756FC85BF0FB23" box="[353,460,1186,1210]" pageId="7" pageNumber="146" type="paratype">paratypes</typeStatus>
). Total length 5.6, carapace 2.6 long, 1.8 wide; abdomen 3.0 long, 2.3 wide; sternum 1.2 long, 1.0 wide. Eye sizes; AME 0.05, ALE 0.13, PME 0.11, PLE 0.11. Distances between eyes; AME-AME 0.06, AME- ALE 0.08, PME-PME 0.10, PME-PLE 0.15, AME-PME 0.20, ALE-PLE 0.06. MOA; anterior width 0.16, posterior width 0.32, length 0.36. Leg measurements: I: 6.8 (1.9, 2.4, 1.4, 1.1); II: 5.4 (1.6, 1.9, 1.1, 0.8); III: 4.8 (1.4, 1.6, 1.1, 0.7); IV: 6.3 (1.8, 2.2, 1.5, 0.8).
</paragraph>
<paragraph id="DE0A36A8C164FFCA80B36D75581AF89E" blockId="7.[140,670,215,1799]" pageId="7" pageNumber="146">
Epigyne and internal genitalia (
<figureCitation id="468E2A2DC164FFCA82076D7558A5F9AE" box="[531,665,1567,1591]" captionStart="Fig" captionStartId="2.[163,196,966,988]" captionTargetBox="[159,1217,366,930]" captionTargetId="figure-49@2.[157,1218,365,933]" captionTargetPageId="2" captionText="Fig. 2. Comparison of female genital organs of Callobius cavernarius sp. n. (A–B); C. shimojanai sp. n. (C–D); and C. breviprocessus (E–F). Epigyne, ventral view (A, C, E); female internal genitalia (B, D, F). Scale bars=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12760068" httpUri="https://zenodo.org/record/12760068/files/figure.png" pageId="7" pageNumber="146">Figs. 2C–D</figureCitation>
,
<figureCitation id="468E2A2DC164FFCA80986D285AF3F9C3" box="[140,207,1602,1626]" captionStart="Fig" captionStartId="6.[163,196,1318,1340]" captionTargetBox="[158,1215,214,1284]" captionTargetId="figure-160@6.[157,1218,213,1285]" captionTargetPageId="6" captionText="Fig. 8. Copulatory organs of Callobius shimojanai sp. n. holotype (A, B, C), paratypes (D, E). Male palp (left), ventral view (A); same, retrolateral view (B); same, dorsal view (C); epigyne, ventral view (D); female internal genitalia (E). Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12760082" httpUri="https://zenodo.org/record/12760082/files/figure.png" pageId="7" pageNumber="146">8D–E</figureCitation>
): lateral lobes ligulate and close to each other at the tip, median lobe quite small and indistinct, ectal lobe and posterior lobe absent. SP anteroposteriorly elongated, slender toward the posterior margin and distant from each other, fertilization ducts small and indistinct.
</paragraph>
<paragraph id="DE0A36A8C164FFCA82C96BBD594DFE8B" blockId="7.[707,1236,215,378]" pageId="7" pageNumber="146">Chelicerae: promargin with 5 teeth, retromargin with 6 teeth.</paragraph>
<paragraph id="DE0A36A8C164FFCA82C96A775914FECE" blockId="7.[707,1236,215,378]" pageId="7" pageNumber="146">
Colouration: almost same as that of the male
<typeStatus id="010E880AC164FFCA82D76A555918FECE" box="[707,804,319,343]" pageId="7" pageNumber="146" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="DE0A36A8C164FFCA82C96A085EA0FEE3" blockId="7.[707,1236,215,378]" box="[733,1180,354,378]" pageId="7" pageNumber="146">
<emphasis id="ECC1EABAC164FFCA82C96A085963FEE3" box="[733,863,354,378]" italics="true" pageId="7" pageNumber="146">Distribution</emphasis>
: Okinawajima Island (
<figureCitation id="468E2A2DC164FFCA84406A085EA8FEE3" box="[1108,1172,354,378]" captionStart="Fig" captionStartId="5.[163,196,861,883]" captionTargetBox="[147,662,214,827]" captionTargetId="figure-299@5.[146,663,213,828]" captionTargetPageId="5" captionText="Fig. 6. Distribution records of Callobius cavernarius sp. n. and C. shimojanai sp. n. including three closely related species living on the Ryukyu Islands." figureDoi="http://doi.org/10.5281/zenodo.12760078" httpUri="https://zenodo.org/record/12760078/files/figure.png" pageId="7" pageNumber="146">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>