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<document ENCODING="UTF8" ID-GBIF-Dataset="f239c787-e431-4a16-8df8-0acf9699b055" checkinTime="1335272175715" checkinUser="thomas" docAuthor="Willmann, C." docDate="1952" docId="A92D03891094432AA76008BF849C37FA" docLanguage="de" docName="Willmann1952_gg1.xml" docOrigin="Veröffentlichungen des Instituts für Meeresforschung Bremerhaven 1" docSource="http://unknown" docTitle="Tyrophagus dimidiatus" docType="treatment" docVersion="12" lastPageNumber="169" masterDocId="F8BE7A25B0C648D91CCA128A88CEA007" masterDocTitle="Die Milbenfauna der Nordseeinsel Wangerooge" masterLastPageNumber="186" masterPageNumber="139" pageNumber="169" updateTime="1661611762921" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Die Milbenfauna der Nordseeinsel Wangerooge</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Willmann, C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Veröffentlichungen des Instituts für Meeresforschung Bremerhaven</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>1952</mods:date>
<mods:detail type="volume">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>186</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://unknown</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="Plazi-MODS">ORI11037</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6286990" ID-GBIF-Taxon="100110468" ID-Zenodo-Dep="6286990" LSID="urn:lsid:plazi:treatment:A92D03891094432AA76008BF849C37FA" httpUri="http://treatment.plazi.org/id/A92D03891094432AA76008BF849C37FA" lastPageNumber="169" pageId="30" pageNumber="169">
<subSubSection pageId="30" pageNumber="169" type="nomenclature">
<paragraph pageId="30" pageNumber="169">
93.
<taxonomicName class="Arachnida" family="Acaridae" genus="Tyrophagus" kingdom="Animalia" order="Astigmata" pageId="30" pageNumber="169" phylum="Arthropoda" rank="species" species="dimidiatus">Tyrophagus dimidiatus</taxonomicName>
(Hermann 1804).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="169" type="materials_examined">
<paragraph pageId="30" pageNumber="169">
Fundorte: Sedumpolster an
<normalizedToken originalValue="zerstörtem" pageId="30" pageNumber="169">zerstoertem</normalizedToken>
Bunker, 9. VI. 49 - Altes
<normalizedToken originalValue="Anspülicht" pageId="30" pageNumber="169">Anspuelicht</normalizedToken>
von Winterhochfluten, 18. VI. 49 - Ruderalplatz am &quot;Haus Friesland&quot;, 22. VIII. 49.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1196.118698" ID-Pensoft-Pub="1313-2970-1196-303" ID-Pensoft-UUID="7DCE9F4064D25B2CAE7EE0EEC7CAC17B" ID-ZooBank="BACE4334CEC348F2A9A1A77D1F106FD3" ModsDocID="1313-2970-1196-303" checkinTime="1711997986401" checkinUser="pensoft" docAuthor="Bedard, Nicolas, Brunke, Adam, Bloin, Pierrick &amp; Leclerc, Ludovic" docDate="2024" docId="A92D0A630E49564ABC14E44911D4996C" docLanguage="en" docName="ZooKeys 1196: 303-329" docOrigin="ZooKeys 1196" docPubDate="2024-04-01" docSource="http://dx.doi.org/10.3897/zookeys.1196.118698" docTitle="Gauropterus fulgidus *" docType="treatment" docVersion="1" id="7DCE9F4064D25B2CAE7EE0EEC7CAC17B" lastPageNumber="303" masterDocId="7DCE9F4064D25B2CAE7EE0EEC7CAC17B" masterDocTitle="New records of rove beetles from the Province of Quebec, and additional provincial records in Canada (Coleoptera, Staphylinidae)" masterLastPageNumber="329" masterPageNumber="303" pageNumber="303" updateTime="1711997986401" updateUser="pensoft">
<mods:mods id="8900CFC5607FE554F48B1F939D10A724" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="49206D69B3A9D4EE1E09F5370B8B36E6">
<mods:title id="6E7A67BFE77EF796197B3201CA733200">New records of rove beetles from the Province of Quebec, and additional provincial records in Canada (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="6811BF1613EBB98CB54730A6FEC6B0B4" type="personal">
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<mods:roleTerm id="B169909C8861C985C51F7D9D391ECAFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC80BD4AC1C0D889B3DB398B12F96570">Bedard, Nicolas</mods:namePart>
<mods:nameIdentifier id="E0A1A356D3EF93908CABE154FD472A46" type="ORCID">https://orcid.org/0009-0004-7649-100X</mods:nameIdentifier>
<mods:affiliation id="A0D8F2D3C76F56DCD67BC4462AA5F4ED">Natural Resources Canada, Canadian Forestry Service, Laurentian Forestry Centre, 1055, rue du P. E. P. S., C. P. 10380, Quebec, QC G 1 V 4 C 7, Canada</mods:affiliation>
<mods:nameIdentifier id="FBF19B8DAA2435461214E3CDD4AAF581" type="email">nicolas.bedard@nrcan-rncan.gc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="4DA1A117D466695AE4603DBCF4870DAE" type="personal">
<mods:role id="D0C5A4221C6673C9D3AE429BACD6C6BC">
<mods:roleTerm id="59702133C4DBFBF52171362FE4FF30C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACDEA2EBF4FB8FB34D191A896C01473A">Brunke, Adam</mods:namePart>
<mods:nameIdentifier id="E494EC6E71272F94F7F6B051358A0B30" type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
<mods:affiliation id="E92C6EA81CE12D43354C0923E99E6025">Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON, K 1 A 0 C 6, Canada</mods:affiliation>
</mods:name>
<mods:name id="58AC316887E341CBCD0123230C46DC2C" type="personal">
<mods:role id="1B5AB7A66978E9D79A940729780528B9">
<mods:roleTerm id="816B4C6576640649FD45ED31C304457A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="030D93FC6F0777A0418EFCDD6C36FAF2">Bloin, Pierrick</mods:namePart>
<mods:nameIdentifier id="A39433C59C84EB1B076B5A3E5AEE0EBE" type="ORCID">https://orcid.org/0009-0009-3260-9017</mods:nameIdentifier>
<mods:affiliation id="87DBE0753510589EEC0B513E50A257C9">Natural Resources Canada, Canadian Forestry Service, Laurentian Forestry Centre, 1055, rue du P. E. P. S., C. P. 10380, Quebec, QC G 1 V 4 C 7, Canada</mods:affiliation>
</mods:name>
<mods:name id="8723299C48C0C0A528D49C4969C8A4CE" type="personal">
<mods:role id="5CF0E718E97391E66CF2F521B01B83E7">
<mods:roleTerm id="A9B9436C39ABBE67F1CB58F80DB75E11">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F6EFC56EF75F022AE0DAB31A5C505E4">Leclerc, Ludovic</mods:namePart>
<mods:nameIdentifier id="0E279907A1B87D829637AC70B4B0CCDE" type="ORCID">https://orcid.org/0009-0006-2381-9627</mods:nameIdentifier>
<mods:affiliation id="047903DEE788B6580CD38DD39F2A6F0B">Laval University, 12325, rue de l'Universite, Quebec, QC G 1 V 0 A 6, Canada &amp; Natural Resources Canada, Canadian Forestry Service, Laurentian Forestry Centre, 1055, rue du P. E. P. S., C. P. 10380, Quebec, QC G 1 V 4 C 7, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5E432B90EA4FE205FFB9C281C467A8E1">text</mods:typeOfResource>
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<mods:titleInfo id="53DED5A9B3412BF8E63A67496FDD1881">
<mods:title id="ED317F1B864CDF42D0D61581821AB7A0">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="9DFBD5B7CB88A93273FB302847836349">
<mods:date id="B1A870B685C5C0A4DB9C9166B63E051F">2024</mods:date>
<mods:detail id="B734C3592060C0EE3A081E2750EA5660" type="pubDate">
<mods:number id="34503F8856495CCD3953348CB5F6C10E">2024-04-01</mods:number>
</mods:detail>
<mods:detail id="625300FADFAE52267A2CDABAB30CE3F1" type="volume">
<mods:number id="B8F37528B273E1AAFAB2E46D1048DB90">1196</mods:number>
</mods:detail>
<mods:extent id="C223FA864AC39DB00B54EFEB442AC6AC" unit="page">
<mods:start id="11887FCC025BBBD18ABECD42A69598DA">303</mods:start>
<mods:end id="4170C24BF66D35245D659CE68638323E">329</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="862BE8955D2BE91F783BB522B932DF8D">
<mods:url id="C03EDCF5755DA77D618827EADEE3D9D0">http://dx.doi.org/10.3897/zookeys.1196.118698</mods:url>
</mods:location>
<mods:classification id="F4CFF9806B957DA8BE1295058E181997">journal article</mods:classification>
<mods:identifier id="BB364DB5A9E3E48B5BEACE9E1828D58B" type="DOI">http://dx.doi.org/10.3897/zookeys.1196.118698</mods:identifier>
<mods:identifier id="7DA303FE081873DB2FABB8AC5EA0509A" type="Pensoft-Pub">1313-2970-1196-303</mods:identifier>
<mods:identifier id="623D33863BFE901BE3EDB5CB76833021" type="ZooBank">BACE4334CEC348F2A9A1A77D1F106FD3</mods:identifier>
<mods:identifier id="5128B92EBA1620FDEE39E66B79606ED5" type="Pensoft-UUID">7DCE9F4064D25B2CAE7EE0EEC7CAC17B</mods:identifier>
</mods:mods>
<treatment id="A92D0A630E49564ABC14E44911D4996C" LSID="urn:lsid:plazi:treatment:A92D0A630E49564ABC14E44911D4996C" httpUri="http://treatment.plazi.org/id/A92D0A630E49564ABC14E44911D4996C" lastPageNumber="303" pageId="0" pageNumber="303">
<subSubSection id="E40B2E7EE7551CD991FF01C19B5EFC1D" pageId="0" pageNumber="303" type="nomenclature">
<paragraph id="FDB7B5CBB255D7141C864369351F05B0" pageId="0" pageNumber="303">
<taxonomicName id="48D4EBCA09E35EE89A29F88208DB4A0B" ID-CoL="3FFDL" LSID="A92D0A63-0E49-564A-BC14-E44911D4996C" authority="(Fabricius, 1787) *" authorityName="*" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Staphylinidae" genus="Gauropterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gauropterus fulgidus" order="Coleoptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="fulgidus">Gauropterus fulgidus (Fabricius, 1787)*</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CC481A3B378D0DB2EF511CCEE4A108F4" pageId="0" pageNumber="303" type="description">
<paragraph id="A5FC634CFA8F052766C6D50ACAD1EE58" pageId="0" pageNumber="303">
<figureCitation id="5C08F6EF73C4A2FDDAB07A02D5AABABE" captionStart="Figures 10, 11" captionStartId="F6" captionText="Figures 10, 11. Habitus of 10 Gauropterus fulgidus (Fabricius, 1787) 11 Phacophallus pallidipennis (Motschulsky, 1858)." figureDoi="10.3897/zookeys.1196.118698.figures10-11" httpUri="https://binary.pensoft.net/fig/1013258" pageId="0" pageNumber="303">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A63386697A88D46CB0FDDA5362758E80" pageId="0" pageNumber="303" type="notes">
<paragraph id="5F56B9BF4FD984E2D1DEB81D0FF0D50F" pageId="0" pageNumber="303">Note.</paragraph>
<paragraph id="69BE883442EC6AB9EBC2790F46696F10" pageId="0" pageNumber="303">
See
<bibRefCitation id="6267A00A06D901B96149F796C09CAA0A" DOI="https://doi.org/10.4039/entm114120fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="303" refId="B56" refString="Smetana, A, 1982. Revision of the subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 114(S120): 1-389. https://doi.org/10.4039/entm114120fv" title="Revision of the subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 114 (S 120): 1 - 389." url="https://doi.org/10.4039/entm114120fv" year="1982">Smetana (1982)</bibRefCitation>
for identification. This very characteristic and large xantholinine beetle was accidentally introduced to North America from Europe in the 19th century and is now found in both the western and eastern parts of North America (
<bibRefCitation id="907869C1ACC1552BDC17B3A05982E323" DOI="https://doi.org/10.4039/entm114120fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="303" refId="B56" refString="Smetana, A, 1982. Revision of the subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 114(S120): 1-389. https://doi.org/10.4039/entm114120fv" title="Revision of the subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 114 (S 120): 1 - 389." url="https://doi.org/10.4039/entm114120fv" year="1982">Smetana 1982</bibRefCitation>
). We report here the first occurrences of this species in the province of Quebec.
</paragraph>
<caption id="9390F17823855EDEDC62216FE1CAE53A" doi="10.3897/zookeys.1196.118698.figures10-11" httpUri="https://binary.pensoft.net/fig/1013258" pageId="0" pageNumber="303" start="Figures 10, 11" startId="F6">
<paragraph id="6EF6CD17F82B2530D57D6E2C431733D5" pageId="0" pageNumber="303">
<emphasis id="68C005BC30139ACC0745FD90A166A6B2" bold="true" pageId="0" pageNumber="303">Figures 10, 11.</emphasis>
Habitus of
<emphasis id="0C94F55349BF59FA5063504219918D00" bold="true" pageId="0" pageNumber="303">10</emphasis>
<taxonomicName id="C4A5AD9A3632483B7249AF7A4188CEBE" authorityName="*" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Staphylinidae" genus="Gauropterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gauropterus fulgidus" order="Coleoptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="fulgidus">
<emphasis id="A8DED3439C770555914603ACC138AF2E" italics="true" pageId="0" pageNumber="303">Gauropterus fulgidus</emphasis>
</taxonomicName>
(Fabricius, 1787)
<emphasis id="F2F9320A487A8D85D6043A6A9B10D823" bold="true" pageId="0" pageNumber="303">11</emphasis>
<taxonomicName id="8B669ABB4DEB8CC0EAC76A7400DFAB8D" authorityName="*" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" class="Insecta" family="Staphylinidae" genus="Phacophallus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phacophallus pallidipennis" order="Coleoptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="pallidipennis">
<emphasis id="4CAB7C216FF54648BB292B72550A82C9" italics="true" pageId="0" pageNumber="303">Phacophallus pallidipennis</emphasis>
</taxonomicName>
(Motschulsky, 1858).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7C440E6D1F2E9B8AA617124BCB39D742" pageId="0" pageNumber="303" type="specimen data">
<paragraph id="3363EBCF8D7140B4447321E38A501D91" pageId="0" pageNumber="303">Specimen data.</paragraph>
<paragraph id="58A5ED6FA69B8DC3CCAD74AEBEC8C8A5" pageId="0" pageNumber="303">
<emphasis id="B450B74E388C70E9F8E7616B9D889DBE" bold="true" pageId="0" pageNumber="303">Canada: QUEBEC - MRC des Deux-Montagnes</emphasis>
, Parc National
<normalizedToken id="B521259EC0B856AC2707C1BEF34E9341" originalValue="d’Oka">d'Oka</normalizedToken>
(
<geoCoordinate id="B73B93101E61D18BA99ABE1534DE7EDB" degrees="45.476714" direction="north" orientation="latitude" precision="1" value="45.476715">45.476714</geoCoordinate>
,
<geoCoordinate id="F3735EA8810F3157FA2A3877C1396AF7" degrees="74.053690" direction="west" orientation="longitude" precision="1" value="-74.05369">-74.053690</geoCoordinate>
), 27.V.2023, R. Vigneault, white tulle fabric interception trap in a compost site [Collected with permit] (1, NBC). -
<emphasis id="7E8E3E499AE11208EAE554496C5407B9" bold="true" pageId="0" pageNumber="303">
Ville de
<normalizedToken id="FE3231237B560B40C7116E0B392311F1" originalValue="Québec">Quebec</normalizedToken>
</emphasis>
, Beauport (
<geoCoordinate id="7BC341E9FCB45C044C2ED604AEAE0A42" degrees="46.9421" direction="north" orientation="latitude" precision="5" value="46.9421">46.9421</geoCoordinate>
,
<geoCoordinate id="E0F084EEE0B9683A5108D4E1A82AE23F" degrees="71.1987" direction="west" orientation="longitude" precision="5" value="-71.1987">-71.1987</geoCoordinate>
), 21.V.2021, N.
<normalizedToken id="0D2EFFC0E8D2DAA8ECC3730835F5B6BA" originalValue="Bédard">Bedard</normalizedToken>
, white tulle fabric interception trap (1, NBC); Sainte-Foy (
<geoCoordinate id="3B90E01315CDA9E7156B36827031883D" degrees="46.7921" direction="north" orientation="latitude" precision="5" value="46.7921">46.7921</geoCoordinate>
,
<geoCoordinate id="48285F62B2B8FF4E61B0C2D8883CCCF2" degrees="71.2806" direction="west" orientation="longitude" precision="5" value="-71.2806">-71.2806</geoCoordinate>
), 15.X.2022, L. Leclerc, sifted from dried vegetal debris (2, LLC);
<normalizedToken id="5309CDAAF2614A81516543EC40D282CC" originalValue="Cité-Universitaire">Cite-Universitaire</normalizedToken>
(
<geoCoordinate id="18654C7EE4757F73D1523F98A4BF50AB" degrees="46.7863" direction="north" orientation="latitude" precision="5" value="46.7863">46.7863</geoCoordinate>
,
<geoCoordinate id="916029D33CB53A442E1DAE58F6838581" degrees="71.2686" direction="west" orientation="longitude" precision="5" value="-71.2686">-71.2686</geoCoordinate>
), 26.X.2022, P. Bloin, sifted from wood chips (2, PBC).
</paragraph>
</subSubSection>
<subSubSection id="A8C90180A7452FB539D24DBC2B5AD0D2" pageId="0" pageNumber="303" type="distribution">
<paragraph id="757CF6862F476648E4F93D5B8E8176E0" pageId="0" pageNumber="303">Distribution in Canada.</paragraph>
<paragraph id="13819E1A1EC5BAB7E07EC59042A1C09A" pageId="0" pageNumber="303">
ON,
<emphasis id="F87080E51FB76D3E8F35FAA2CB5B9E2E" bold="true" pageId="0" pageNumber="303">QC</emphasis>
(
<bibRefCitation id="43B8066A2B89852B6FACE4B90D9A13F6" DOI="https://doi.org/10.3897/zookeys.360.4742" author="Bousquet, Y" journalOrPublisher="109, Sofia-Moscow" pageId="0" pageNumber="303" refId="B4" refString="Bousquet, Y, Bouchard, P, Davies, AE, Sikes, D, 2013. Checklist of beetles (Coleoptera) of Canada and Alaska. Pensoft Series Faunistica No. 109, Sofia-Moscow, DOI: https://doi.org/10.3897/zookeys.360.4742" title="Checklist of beetles (Coleoptera) of Canada and Alaska. Pensoft Series Faunistica No." url="https://doi.org/10.3897/zookeys.360.4742" year="2013">Bousquet et al. 2013</bibRefCitation>
) -
<emphasis id="6885267EFCDF909C3113EB183285AF79" bold="true" pageId="0" pageNumber="303">New to Quebec</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The ground beetles (Coleoptera: Carabidae) of the Strandzha Mountain and adjacent coastal territories (Bulgaria and Turkey)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kostova, Rumyana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>8135</mods:start>
<mods:end>8135</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.4.e8135</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e8135</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--8135</mods:identifier>
</mods:mods>
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<subSubSection pageId="0" pageNumber="8135" type="nomenclature">
<paragraph pageId="0" pageNumber="8135">
<taxonomicName authority="Paykull, 1790" authorityName="Paykull" authorityYear="1790" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limodromus assimilis" order="Coleoptera" pageId="0" pageNumber="8135" phylum="Arthropoda" rank="species" species="assimilis">Limodromus assimilis (Paykull, 1790)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8135" type="materials_examined">
<paragraph pageId="0" pageNumber="8135">Materials</paragraph>
<paragraph pageId="0" pageNumber="8135">
<materialsCitation collectingDate="2009-04-19" collectorName="R. Kostova" latitude="42.03086" location="Stoilovo Vill., &quot; Sredoka &quot; Reserve, along Mechi dol River" longLatPrecision="1" longitude="27.513916" pageId="0" pageNumber="8135" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">R. Kostova</collectorName>
; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
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,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.513916">E27°30'50.1&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135" value="2009-04-19">19/04/2009</collectingDate>
</materialsCitation>
<materialsCitation collectingDate="2009-07-05" collectorName="R. Bekchiev" latitude="41.877583" location="Along Demirkoey - Igneada road" longLatPrecision="1" longitude="27.941444" pageId="0" pageNumber="8135" specimenCount="3" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">R. Bekchiev</collectorName>
; individualCount:
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; Location: countryCode: TR; locality:
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Along
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; verbatimElevation:
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; verbatimCoordinates:
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,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.941444">E27°56'29.2&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
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</materialsCitation>
<materialsCitation collectingDate="2010-05-27" collectorName="R. Kostova" latitude="42.03086" location="Stoilovo Vill., &quot; Sredoka &quot; Reserve, along Mechi dol River" longLatPrecision="1" longitude="27.513916" pageId="0" pageNumber="8135" specimenCount="4" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
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,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.513916">E27°30'50.1&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
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</materialsCitation>
<materialsCitation collectingDate="2010-05-27" collectorName="R. Kostova" latitude="41.983807" location="Malko Tarnovo surroundings" longLatPrecision="1" longitude="27.497055" pageId="0" pageNumber="8135" specimenCount="2" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:0011A0D042A9AB7345DFFAD7B775EB7C" latitude="41.983807" longLatPrecision="1" longitude="27.497055" name="Malko Tarnovo surroundings" pageId="0" pageNumber="8135">Malko Tarnovo surroundings</location>
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.983807">N41°59'01.7&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.497055">E27°29'49.4&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
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</materialsCitation>
<materialsCitation collectingDate="09.06 - 02.07.2009" collectorName="R. Kostova" latitude="41.980446" location="Malko Tarnovo, &quot; Propada &quot; Place" longLatPrecision="1" longitude="27.490473" pageId="0" pageNumber="8135" specimenCount="3" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.980446">N41°58'49.6&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.490473">E27°29'25.7&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
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</materialsCitation>
<materialsCitation collectingDate="09.05 - 08.06.2009" collectorName="R. Kostova" latitude="41.960503" location="Slivarovo Vill., &quot; Shafariitsa &quot; Place" longLatPrecision="1" longitude="27.659666" pageId="0" pageNumber="8135" specimenCount="2" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.960503">N41°57'37.8&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.659666">E27°39'34.8&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135">09.05-08.06.2009</collectingDate>
; habitat: black alder-oak forest
</materialsCitation>
<materialsCitation collectingDate="03.07 - 02.08.2009" collectorName="R. Kostova" latitude="41.960503" location="Slivarovo Vill., &quot; Shafariitsa &quot; Place" longLatPrecision="1" longitude="27.659666" pageId="0" pageNumber="8135" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.960503">N41°57'37.8&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.659666">E27°39'34.8&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135">03.07-02.08.2009</collectingDate>
; habitat: black alder-oak forest
</materialsCitation>
<materialsCitation collectingDate="15.04 - 08.05.2009" collectorName="R. Kostova" latitude="42.056442" location="Kosti Vill., &quot; St. Ilia &quot; Place" longLatPrecision="1" longitude="27.764334" pageId="0" pageNumber="8135" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="42.056442">N42°03'23.2&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.764334">E27°45'51.6&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
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; habitat: meadow with single trees
</materialsCitation>
<materialsCitation collectingDate="03.07 - 02.08.2009" collectorName="R. Kostova" latitude="42.056442" location="Kosti Vill., &quot; St. Ilia &quot; Place" longLatPrecision="1" longitude="27.764334" pageId="0" pageNumber="8135" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; Location: countryCode: BG; locality:
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; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="42.056442">N42°03'23.2&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.764334">E27°45'51.6&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135">03.07-02.08.2009</collectingDate>
; habitat: meadow with single trees
</materialsCitation>
<materialsCitation collectingDate="15.04 - 07.09.2009" collectorName="R. Kostova" latitude="42.061" location="Sinemorets Vill., PA &quot; Estuary of Veleka river &quot;" longLatPrecision="1" longitude="27.965334" pageId="0" pageNumber="8135" specimenCount="27" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">R. Kostova</collectorName>
; individualCount:
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; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:10BDB9F0CF6806666E0261D6A00C485F" latitude="42.061" longLatPrecision="1" longitude="27.965334" name="Sinemorets Vill., PA &quot; Estuary of Veleka river &quot;" pageId="0" pageNumber="8135">Sinemorets Vill., PA &quot;Estuary of Veleka river&quot;</location>
; verbatimElevation:
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; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="42.061">N42°03'39.6&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.965334">E27°57'55.2&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135">15.04-07.09.2009</collectingDate>
; habitat: swamp forest
</materialsCitation>
<materialsCitation collectingDate="06.07 - 29.09.2009" collectorName="R. Kostova" latitude="41.864166" location="Igneada surroundings" longLatPrecision="1" longitude="27.94836" pageId="0" pageNumber="8135" specimenCount="7" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">R. Kostova</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="8135">7</specimenCount>
; Location: countryCode: TR; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:81FB79F90B3C55F33F4A63F0DBE137FE" latitude="41.864166" longLatPrecision="1" longitude="27.94836" name="Igneada surroundings" pageId="0" pageNumber="8135">Igneada surroundings</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="8135">15</elevation>
; verbatimCoordinates:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.864166">N41°51'51.0&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="27.94836">E27°56'54.1&quot;</geoCoordinate>
; geodeticDatum: WGS84; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135">06.07-29.09.2009</collectingDate>
; habitat: swamp forest
</materialsCitation>
<materialsCitation collectingDate="1934-07" collectionCode="NMP" collectorName="K. Taborsky" location="Maslen nos Cape" pageId="0" pageNumber="8135" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">K. Taborsky</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="8135">1</specimenCount>
; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:E0EFFABF6EC611FD4251FBEA04657872" name="Maslen nos Cape" pageId="0" pageNumber="8135">Maslen nos Cape</location>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135" value="1934-07">VII.1934</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="8135">NMP</collectionCode>
</materialsCitation>
<materialsCitation collectingDate="1995-05-25" collectionCode="NMNHS" collectorName="B. Gueorguiev" location="Veleka River near Mladezko Village" pageId="0" pageNumber="8135" specimenCount="6" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">
B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="8135">6</specimenCount>
; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:7FEAB445139AC54D1694494B8C10395F" name="Veleka River near Mladezko Village" pageId="0" pageNumber="8135">Veleka River near Mladezko Village</location>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135" value="1995-05-25">25/05/1995</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="8135">NMNHS</collectionCode>
</materialsCitation>
<materialsCitation collectingDate="1995-05-26" collectionCode="NMNHS" collectorName="B. Gueorguiev" location="Aydere River" pageId="0" pageNumber="8135" specimenCount="6" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">
B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="8135">6</specimenCount>
; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:4DD1DCA2D70D04BBFF57F81C5B56D28E" name="Aydere River" pageId="0" pageNumber="8135">Aydere River</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="8135">350-450</elevation>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135" value="1995-05-26">26/05/1995</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="8135">NMNHS</collectionCode>
</materialsCitation>
<materialsCitation collectingDate="1995-05-29" collectionCode="NMNHS" collectorName="B. Gueorguiev" location="Nature Park Ropotamo" pageId="0" pageNumber="8135" specimenCount="2" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="8135">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="8135">
B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="8135">2</specimenCount>
; Location: countryCode: BG; locality:
<location LSID="urn:lsid:plazi:treatment:A92D14A92DFA5842FF95138907C0F797:67718108FD468F61078543999B5F099B" name="Nature Park Ropotamo" pageId="0" pageNumber="8135">Nature Park Ropotamo</location>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="8135" value="1995-05-29">29/05/1995</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="8135">NMNHS</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="42BCD675E67F199DA65DE1D7AFE3945D" pageId="67" pageNumber="68">Original source.</paragraph>
<paragraph id="70E71C9A573BEBF03E20FCC078320A51" pageId="67" pageNumber="68">
<bibRefCitation id="B828E2B42199D01DA3DE210CADE6EC83" author="Gentiluomo, C" journalOrPublisher="Bullettino Malacologico Italiano" pageId="311" pageNumber="312" pagination="81 - 100" publicationUrl="http://www.biodiversitylibrary.org/item/122154page/131/mode/1up" refId="B180" refString="Gentiluomo, C, 1868. Catalogo dei Molluschi terrestri e fluviatili della Toscana. Bullettino Malacologico Italiano 1 (6): 81 - 100, http://www.biodiversitylibrary.org/item/122154page/131/mode/1up" title="Catalogo dei Molluschi terrestri e fluviatili della Toscana." url="http://www.biodiversitylibrary.org/item/122154page/131/mode/1up" volume="1" year="1868">Gentiluomo 1868</bibRefCitation>
: 97, pl. 6, figs 10-11.
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<paragraph id="B33FFD0EA268B51B3B6D0C2DEF6712D7" pageId="67" pageNumber="68">Type locality.</paragraph>
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&quot;Lago
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, Toscana&quot;, Italy.
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<paragraph id="37F61D288320499E6E9DE2A071E3429C" pageId="67" pageNumber="68">Remarks.</paragraph>
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Junior homonym of
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<emphasis id="A733B9BBF2205A18A0694D222B6F8879" italics="true" pageId="67" pageNumber="68">Melanopsis carinata</emphasis>
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Sowerby, 1826.
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<paragraph id="3EE201CAEA3F10A24BD752B9398721B8" pageNumber="390">&quot;Habitat in Europa, Canada.&quot; RCN: 7231.</paragraph>
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<treatment id="A92D421AFFEBFFE4FF6A06BFFC46FE93" ID-DOI="http://doi.org/10.5281/zenodo.6082350" ID-GBIF-Taxon="124483809" ID-Zenodo-Dep="6082350" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93" httpUri="http://treatment.plazi.org/id/A92D421AFFEBFFE4FF6A06BFFC46FE93" lastPageId="5" lastPageNumber="386" pageId="0" pageNumber="381">
<subSubSection id="699EA087FFEBFFE1FF6A06BFFE85F8AC" pageId="0" pageNumber="381" type="nomenclature">
<paragraph id="213BF30CFFEBFFE1FF6A06BFFD8BF88E" blockId="0.[151,553,1766,1826]" box="[151,553,1766,1792]" pageId="0" pageNumber="381">
<heading id="7A734460FFEBFFE1FF6A06BFFD8BF88E" bold="true" box="[151,553,1766,1792]" fontSize="11" level="1" pageId="0" pageNumber="381" reason="1">
<taxonomicName id="E684888FFFEBFFE1FF6A06BFFD8BF88E" ID-CoL="73V85" authority="Okamoto, 1922" authorityName="Okamoto" authorityYear="1922" box="[151,553,1766,1792]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="381" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEBFFE1FF6A06BFFD8BF88E" bold="true" box="[151,553,1766,1792]" pageId="0" pageNumber="381">
<emphasis id="13F02F1EFFEBFFE1FF6A06BFFEC1F88E" bold="true" box="[151,355,1766,1792]" italics="true" pageId="0" pageNumber="381">Nemoura papilla</emphasis>
<bibRefCitation id="45158EFDFFEBFFE1FE9706BFFD8BF88E" author="Okamoto" box="[362,553,1766,1792]" pageId="0" pageNumber="381" refString="Okamoto, H. (1922) Zweiter Beitrag zur Kenntnis der Japanischen Plecopteren. Bulletin of the Ehima Agricultural Experiment Station, 1, 1 - 46. [Gov. - Gen. Chosen, Suigen]" type="journal article" year="1922">Okamoto, 1922</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="213BF30CFFEBFFE1FF6A0750FE85F8AC" blockId="0.[151,553,1766,1826]" box="[151,295,1801,1826]" pageId="0" pageNumber="381">
(
<figureCitation id="B9BFEF89FFEBFFE1FF620750FF4AF8AC" box="[159,232,1801,1826]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="0" pageNumber="381">Figs. 1</figureCitation>
–12).
</paragraph>
</subSubSection>
<subSubSection id="699EA087FFEBFFE1FF6A0708FAFAF82A" pageId="0" pageNumber="381" type="description">
<paragraph id="213BF30CFFEBFFE1FF6A0708FAFAF82A" blockId="0.[151,1436,1873,1956]" pageId="0" pageNumber="381">
<bibRefCitation id="45158EFDFFEBFFE1FF6A0708FE97F8E9" author="Okamoto" box="[151,309,1873,1895]" pageId="0" pageNumber="381" refString="Okamoto, H. (1922) Zweiter Beitrag zur Kenntnis der Japanischen Plecopteren. Bulletin of the Ehima Agricultural Experiment Station, 1, 1 - 46. [Gov. - Gen. Chosen, Suigen]" type="journal article" year="1922">Okamoto 1922</bibRefCitation>
. Bull. Agric. Exp. Stn. Chosen, Suigen 1(1):1–46 + Tafe l I–VI (original description of male and female);
<bibRefCitation id="45158EFDFFEBFFE1FF3A0729FEFCF808" author="Shimizu" box="[199,350,1904,1926]" pageId="0" pageNumber="381" refString="Shimizu, T. (1994) Taxonomic Changes and Synonyms for the East Asian Species of the Genus Nemoura (Plecoptera: Nemouridae). Aquatic Insects, 16 (4), 213 - 225. http: // dx. doi. org / 10.1080 / 01650429409361558" type="journal article" year="1994">Shimizu, 1994</bibRefCitation>
, Aquatic Insects, 16(4): 221–223 (
<taxonomicName id="E684888FFFEBFFE1FD3E0728FBB5F808" authority="Okamoto, 1922" authorityName="Okamoto" authorityYear="1922" box="[707,1047,1904,1926]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="381" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEBFFE1FD3E0728FCD2F80B" box="[707,880,1904,1926]" italics="true" pageId="0" pageNumber="381">Nemoura papilla</emphasis>
<bibRefCitation id="45158EFDFFEBFFE1FC850729FBB5F808" author="Okamoto" box="[888,1047,1904,1926]" pageId="0" pageNumber="381" refString="Okamoto, H. (1922) Zweiter Beitrag zur Kenntnis der Japanischen Plecopteren. Bulletin of the Ehima Agricultural Experiment Station, 1, 1 - 46. [Gov. - Gen. Chosen, Suigen]" type="journal article" year="1922">Okamoto, 1922</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="E684888FFFEBFFE1FBC90728FA3AF808" authority="Kawai, 1954" authorityName="Kawai" authorityYear="1954" box="[1076,1432,1904,1926]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="381" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="13F02F1EFFEBFFE1FBC90728FAAEF80B" box="[1076,1292,1904,1926]" italics="true" pageId="0" pageNumber="381">Nemoura denticulata</emphasis>
Kawai, 1954
</taxonomicName>
,
<taxonomicNameLabel id="08C39265FFEBFFE1FF3A07D6FE80F82A" box="[199,290,1935,1956]" pageId="0" pageNumber="381">syn. nov.</taxonomicNameLabel>
and
<taxonomicName id="E684888FFFEBFFE1FEAF07D6FDD1F82A" authority="Zwick, 1974" authorityName="Zwick" authorityYear="1974" box="[338,627,1934,1956]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="381" phylum="Arthropoda" rank="species" species="levanidovae">
<emphasis id="13F02F1EFFEBFFE1FEAF07D6FE48F82A" box="[338,490,1935,1956]" italics="true" pageId="0" pageNumber="381">N. levanidovae</emphasis>
Zwick, 1974
</taxonomicName>
,
<taxonomicNameLabel id="08C39265FFEBFFE1FD8007D6FD78F82A" box="[637,730,1935,1956]" pageId="0" pageNumber="381">syn. nov.</taxonomicNameLabel>
) (supplementary description of the epiproct, male and female).
</paragraph>
</subSubSection>
<subSubSection id="699EA087FFEBFFE0FF6A0788FAAAF932" lastPageId="1" lastPageNumber="382" pageId="0" pageNumber="381" type="materials_examined">
<paragraph id="213BF30CFFEBFFE0FF6A0788FD9EFF76" blockId="0.[151,1436,2001,2026]" lastBlockId="1.[151,1437,151,2013]" lastPageId="1" lastPageNumber="382" pageId="0" pageNumber="381">
<materialsCitation id="91ECF951FFEBFFE0FF6A0788FEF5FF5B" ID-GBIF-Occurrence="1306554148" collectorName="V. Teslenko" county="Material" lastPageId="1" lastPageNumber="382" latitude="43.407932" location="Anan'evka River" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" pageId="0" pageNumber="381" specimenCount="67" specimenCount-female="26" specimenCount-larva="35" specimenCount-male="6" stateProvince="Primorsky">
<emphasis id="13F02F1EFFEBFFE1FF6A0788FE27F864" bold="true" box="[151,389,2001,2026]" pageId="0" pageNumber="381">
<collectingCounty id="C85A8B80FFEBFFE1FF6A0788FF5DF864" box="[151,255,2001,2026]" pageId="0" pageNumber="381">Material</collectingCounty>
examined.
</emphasis>
<collectingMunicipality id="C15F6976FFEBFFE1FE6C0788FDF8F867" box="[401,602,2001,2026]" pageId="0" pageNumber="381">Russian Far East</collectingMunicipality>
,
<collectingRegion id="E3403DEEFFEBFFE1FD960788FD7DF864" box="[619,735,2001,2026]" country="Russia" name="Primorskiy" pageId="0" pageNumber="381">Primorsky</collectingRegion>
Region:
<specimenCount id="37823885FFEBFFE1FCAD0788FC0DF864" box="[848,943,2001,2026]" pageId="0" pageNumber="381" type="male">6 males</specimenCount>
,
<specimenCount id="37823885FFEBFFE1FC420788FBE3F864" box="[959,1089,2001,2026]" pageId="0" pageNumber="381" type="female">26 females</specimenCount>
,
<specimenCount id="37823885FFEBFFE1FBAC0788FB62F864" box="[1105,1216,2001,2026]" pageId="0" pageNumber="381" type="larva">35 larvae</specimenCount>
,
<location id="245BA5D7FFEBFFE1FB2C0788FA3AF864" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEBFFE1FB2C0788FA3AF864" box="[1233,1432,2001,2026]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="Anan'evka River" pageId="0" pageNumber="381" stateProvince="Primorsky">Anan’evka River</location>
,
<location id="245BA5D7FFEAFFE0FF6A00CEFE17FF3E" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEAFFE0FF6A00CEFE17FF3E" box="[151,437,151,176]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="Razdol'naya River Basin" pageId="1" pageNumber="382" stateProvince="Primorsky">
<location id="245BA5D7FFEAFFE0FF6A00CEFEC9FF3E" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEAFFE0FF6A00CEFEC9FF3E" box="[151,363,151,176]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="Razdol'naya River" pageId="1" pageNumber="382" stateProvince="Primorsky">
Razdol’naya
<location id="245BA5D7FFEAFFE0FED000CEFEC9FF3E" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEAFFE0FED000CEFEC9FF3E" box="[301,363,151,176]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="River Basin" pageId="1" pageNumber="382" stateProvince="Primorsky">River</location>
</location>
Basin
</location>
,
<location id="245BA5D7FFEAFFE0FE3F00C1FC81FF3E" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEAFFE0FE3F00C1FC81FF3E" box="[450,803,151,176]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="upper Venevitinovo Settlement" pageId="1" pageNumber="382" stateProvince="Primorsky">upper Venevitinovo Settlement</location>
,
<geoCoordinate id="44B095CBFFEAFFE0FCD200C1FC64FF3E" box="[815,966,151,176]" direction="north" orientation="latitude" pageId="1" pageNumber="382" precision="1" value="43.407932">N 43º24.476’</geoCoordinate>
<geoCoordinate id="44B095CBFFEAFFE0FC3200C1FBC9FF3E" box="[975,1131,152,176]" direction="east" orientation="longitude" pageId="1" pageNumber="382" precision="1" value="131.73837">E131º44.302’</geoCoordinate>
, 27.04−4.05.2015, rearing, coll.
<collectorName id="8C7196DAFFEAFFE0FF3200E4FEF5FF5B" box="[207,343,188,213]" pageId="1" pageNumber="382">V. Teslenko</collectorName>
</materialsCitation>
;
<materialsCitation id="91ECF951FFEAFFE0FE9E00E5FD9AFF76" ID-GBIF-Occurrence="1306554146" collectingDate="2015-08-16" collectorName="Meortvaya Pad Stream, N &amp; V. Teslenko" location="Vladivostok" pageId="1" pageNumber="382" specimenCount="15" specimenCount-larva="15" stateProvince="Primorsky">
<specimenCount id="37823885FFEAFFE0FE9E00E5FE6FFF5B" box="[355,461,188,213]" pageId="1" pageNumber="382" type="larva">15 larvae</specimenCount>
,
<location id="245BA5D7FFEAFFE0FE2400E5FDC7FF5B" LSID="urn:lsid:plazi:treatment:A92D421AFFEBFFE4FF6A06BFFC46FE93:245BA5D7FFEAFFE0FE2400E5FDC7FF5B" box="[473,613,188,213]" name="Vladivostok" pageId="1" pageNumber="382" stateProvince="Primorsky">Vladivostok</location>
, Rybachiy Sett.,
<collectorName id="8C7196DAFFEAFFE0FCD900E4FBEAFF5A" box="[804,1096,188,213]" pageId="1" pageNumber="382">Meortvaya Pad Stream, N</collectorName>
42°080.417’ E 132°021.346’, 28.07.−
<date id="553AD5CCFFEAFFE0FF160086FECCFF76" box="[235,366,223,248]" pageId="1" pageNumber="382" value="2015-08-16">
<collectingDate id="457E2C24FFEAFFE0FF160086FECCFF76" box="[235,366,223,248]" pageId="1" pageNumber="382" value="2015-08-16">16.08.2015</collectingDate>
</date>
, coll.
<collectorName id="8C7196DAFFEAFFE0FE4C00B9FD9AFF76" box="[433,568,223,248]" pageId="1" pageNumber="382">V. Teslenko</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="213BF30CFFEAFFE0FF3A015DFF17FC62" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="382">
<emphasis id="13F02F1EFFEAFFE0FF3A015DFECBFE93" bold="true" box="[199,361,260,285]" pageId="1" pageNumber="382">Mature larva</emphasis>
(
<figureCitation id="B9BFEF89FFEAFFE0FE86015DFE65FE92" box="[379,455,260,284]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1</figureCitation>
−12). Body relatively slender, length in males,
<quantity id="E67C5EE9FFEAFFE0FC12015DFBDEFE95" box="[1007,1148,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" pageId="1" pageNumber="382" unit="mm" value="5.95" valueMax="6.4" valueMin="5.5">5.5–6.4 mm</quantity>
; in females
<quantity id="E67C5EE9FFEAFFE0FAF6015DFA3BFE95" box="[1291,1433,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="1" pageNumber="382" unit="mm" value="7.3" valueMax="7.8" valueMin="6.8">6.8–7.8 mm</quantity>
. General color a mottled brown with specific pattern consisting of pale and dark brown patches on head, legs, and cerci, and a pale mesal stripe on thorax (
<figureCitation id="B9BFEF89FFEAFFE0FD930115FD42FEEA" box="[622,736,332,357]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
). Head with pale, tentorial callosities outlined by contrasting unconnected dark brown lines towards posterior ocelli, a rectangular brown interocellar patch with thin longitudinal pale line medially, a diffuse brownish patch in front of the anterior ocellus extending to clypeus (
<figureCitation id="B9BFEF89FFEAFFE0FA9201CDFF0AFE5E" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Fig. 2</figureCitation>
). Epicranial suture appear as pale Y mark between the lateral ocelli; an oval, oblique pale patch to the inside of each compound eye. Occiput with pair of oval oblique pale patches surrounded by thread-like brown loops, posterior margin darkened (
<figureCitation id="B9BFEF89FFEAFFE0FE270259FD81FD96" box="[474,547,512,536]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Fig. 2</figureCitation>
). Antenna brown, long, length reaches 85−100% of body length; scape and pedicel dark brown. Pronotum oval, approximately 1.2 X wider than long, pattern diffuse, with small dark brown spots and pale patches of different size and form; narrow pale mesal stripe on pronotum continues over meso-, metanotum and abdominal segments (
<figureCitation id="B9BFEF89FFEAFFE0FDB90235FD13FD0A" box="[580,689,620,645]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
). Meso- and metanotum with U-shaped brown pattern and diffuse pale patches anteromedially. Legs slender, long with contrasting brown and dark brown bands and pale patches. Femora brown with diffuse dark brown band in the distal ⅓ and one pale patch apically, sometimes with a bald median pale line (
<figureCitation id="B9BFEF89FFEAFFE0FE99028EFE70FD7E" box="[356,466,727,752]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
). Fore femur 3.3 X longer than wide (
<figureCitation id="B9BFEF89FFEAFFE0FC7A028EFC6EFD7E" box="[903,972,727,752]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Fig. 6</figureCitation>
) and hind femur about 4.1−5.0 X longer than wide. Tibia brown with two diffuse dark brown bands in the basal and distal ⅓ (
<figureCitation id="B9BFEF89FFEAFFE0FB7E02A5FB56FC9A" box="[1155,1268,764,789]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
) and two pale patches basally and distally. Third tarsal segment darkened basally and distally, pale medially. Abdomen relatively slender, integument light, matte in appearance, terga brownish with a mesal row of pale small patches forming a medial longitudinal band (
<figureCitation id="B9BFEF89FFEAFFE0FE3D033EFD8FFC0E" box="[448,557,871,896]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
). Cerci pale, very long, length reaches 100−133% of body length, with 31−33 segments, each segment darkened basally and distally, pale medially; the segment sides are nearly parallel in basal part of cercus, club-shaped in middle and apical parts, apical segments very long,
<date id="553AD5CCFFEAFFE0FBD703F6FBF8FC46" box="[1066,1114,943,968]" pageId="1" pageNumber="382" value="1925-10">25X</date>
longer than wide (
<figureCitation id="B9BFEF89FFEAFFE0FAD103F6FA35FC46" box="[1324,1431,943,968]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,2006,2029]" captionTargetBox="[153,1436,194,1984]" captionTargetId="figure@2.[150,1436,193,1985]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Larva of Nemoura papilla, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="1" pageNumber="382">Figs. 1, 2</figureCitation>
, 9).
</paragraph>
<paragraph id="213BF30CFFEAFFE0FF3A03AEFAAAF932" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="382">
Setation weakly visible. Setal fringe around the pronotum relatively spaced, consisting of club-shaped bristles of different length (
<figureCitation id="B9BFEF89FFEAFFE0FE8C0445FE7EFBBA" box="[369,476,1052,1077]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Figs. 3, 4</figureCitation>
). Few longest club-shaped bristles present at the posterior corners, the longest bristles approximately equal 4.9−5.5% of pronotum width. Pronotum disc bears tiny oval setae with widened apex, short club-shaped setae, and occasional long hairs (
<figureCitation id="B9BFEF89FFEAFFE0FD69043DFCA2FBF2" box="[660,768,1124,1149]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Figs. 3, 4</figureCitation>
). Outer wing pad margin covered with club-shaped bristles; tiny club-shaped setae in the lines on wing-pads (
<figureCitation id="B9BFEF89FFEAFFE0FD3604DEFCB2FB2E" box="[715,784,1159,1184]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Fig. 5</figureCitation>
). Legs with markedly heterogeneous chaetotaxy (
<figureCitation id="B9BFEF89FFEAFFE0FABA04DEFF05FB4A" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Figs. 6, 7</figureCitation>
). All femora covered with spatulate bristles of different length, occasionally fine procumbent setae and seldom thin long hairs (
<figureCitation id="B9BFEF89FFEAFFE0FEAD0496FE63FB66" box="[336,449,1231,1256]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Figs. 6, 7</figureCitation>
). Setal fringe on outer femur margin in the apical half consisting of three long, clubshaped bristles in an irregular arrangement on fore leg, and with five long, club-shaped bristles on hind leg (
<figureCitation id="B9BFEF89FFEAFFE0FAB504ADFF05FABE" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Figs. 6, 7</figureCitation>
). The longest club-shaped bristles reach 37% of the width of femur on fore leg, and 26% of the hind leg. Tibia with sporadic fine procumbent setae, dense and strong spine-like bristles along the inner edge, the bristles not longer than width of tibia; the outer edge bears tiny club-shaped setae (
<figureCitation id="B9BFEF89FFEAFFE0FC2D0506FBB5FAF6" box="[976,1047,1375,1400]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1860,1883]" captionTargetBox="[150,1436,193,1830]" captionTargetId="figure@3.[150,1436,193,1840]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Larva of Nemoura papilla. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3 – 5 = 0.1; 6 = 0.3; 7 = 0.05." httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="1" pageNumber="382">Fig. 6</figureCitation>
). Terga covered with sparse short club-shaped bristles. The posterior tergal margins feature mostly short club-shaped bristles and one pair of conspicuous relatively long slightly curved club-shaped bristles (Fig. 8). Length of the longest bristles on terga 5−6 reaches 34% of segment length (Fig. 8). Cercal chaetotaxy heterogeneous, bristles of different form and size in the apical whorl; intercalary setation very sparse and almost invisible (Figs. 10−12). Apical whorl comprises a set of club-shaped bristles on basal cercal segments; the longest club-shaped bristles reach 67% of segment length on cercal segments 8−10 (Fig. 10). In the middle part of cerci the apical whorl bears tiny setae with rounded apex mixed with long, acute bristles, which do not exceed 59% of the segment’s length on cercal segments 16−17 (Fig. 11). On apical cercal segments the apical whorls include tiny setae with rounded apex and one or two thin acute bristles (like hairs), bristle length is only about 16% of segment length at cercal segment 31 (Fig. 12).
</paragraph>
</subSubSection>
<subSubSection id="699EA087FFEAFFE0FF3A069EFC9AF853" pageId="1" pageNumber="382" type="diagnosis">
<paragraph id="213BF30CFFEAFFE0FF3A069EFC9AF853" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="382">
<emphasis id="13F02F1EFFEAFFE0FF3A069EFEE3F96E" bold="true" box="[199,321,1735,1760]" pageId="1" pageNumber="382">Diagnosis.</emphasis>
The larvae of
<taxonomicName id="E684888FFFEAFFE0FE110690FDFCF96E" box="[492,606,1736,1760]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEAFFE0FE110690FDFCF96E" box="[492,606,1736,1760]" italics="true" pageId="1" pageNumber="382">N. papilla</emphasis>
</taxonomicName>
can be separated from known larvae of East-Asian
<taxonomicName id="E684888FFFEAFFE0FB490690FABEF96E" box="[1204,1308,1737,1760]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="13F02F1EFFEAFFE0FB490690FABEF96E" box="[1204,1308,1737,1760]" italics="true" pageId="1" pageNumber="382">Nemoura</emphasis>
</taxonomicName>
species by mottled contrasting body pattern, long antenna, cerci exceeding the body length, weak but conspicuous variable setation on the legs and cerci. Setal fringe of the pronotum sparse with club-shaped bristles, club-shaped bristles of different size present also in setal fringes along outer wing pad margin, outer femur margin, posterior margins of terga and in the apical whorls on basal cercal segments. Setation of femur also includes a few long club-shaped bristles and special spatulate bristles. The medial apical whorl of the cerci differs from the apical whorl on basal segments having long acute bristles not exceeding 59% of segment length at cercal segments 16−17. Intercalary setation on cercal segments sparse and almost indiscernible.
</paragraph>
</subSubSection>
<caption id="75FBA384FFE9FFE3FF6A078FFC05F862" box="[151,935,2006,2029]" httpUri="https://zenodo.org/record/256791/files/figure.png" pageId="2" pageNumber="383" targetBox="[153,1436,194,1984]" targetPageId="2">
<paragraph id="213BF30CFFE9FFE3FF6A078FFC05F862" blockId="2.[151,935,2006,2029]" box="[151,935,2006,2029]" pageId="2" pageNumber="383">
<emphasis id="13F02F1EFFE9FFE3FF6A078FFE98F862" bold="true" box="[151,314,2006,2029]" pageId="2" pageNumber="383">FIGURES 1–2.</emphasis>
Larva of
<emphasis id="13F02F1EFFE9FFE3FE60078EFDF2F862" box="[413,592,2007,2028]" italics="true" pageId="2" pageNumber="383">
<taxonomicName id="E684888FFFE9FFE3FE60078EFDE9F862" box="[413,587,2007,2028]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="papilla">Nemoura papilla</taxonomicName>
,
</emphasis>
habitus, dorsal view, not to scale.
</paragraph>
</caption>
<caption id="75FBA384FFE8FFE2FF6A071DFE8FF839" httpUri="https://zenodo.org/record/256792/files/figure.png" pageId="3" pageNumber="384" targetBox="[150,1436,193,1830]" targetPageId="3">
<paragraph id="213BF30CFFE8FFE2FF6A071DFE8FF839" blockId="3.[151,1436,1860,1975]" pageId="3" pageNumber="384">
<emphasis id="13F02F1EFFE8FFE2FF6A071DFE99F8D4" bold="true" box="[151,315,1860,1883]" pageId="3" pageNumber="384">FIGURES 3–7.</emphasis>
Larva of
<taxonomicName id="E684888FFFE8FFE2FE5F071FFDEDF8D4" box="[418,591,1861,1883]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFE8FFE2FE5F071FFDEDF8D4" box="[418,591,1861,1883]" italics="true" pageId="3" pageNumber="384">Nemoura papilla</emphasis>
</taxonomicName>
. 3. Сlub-shaped bristles near midline on anterior margin of pronotum. 4. Setae on disc and setal fringe of right anterior corner of pronotum. 5. Bristles on outer margin and rows of tiny setae on wing pad. 6. Right fore leg. 7. Setal fringe on outer femur margin on hind leg with five long club-shaped bristles Scale (mm): 3–5=0.1; 6=0.3; 7=0.05.
</paragraph>
</caption>
<paragraph id="213BF30CFFEFFFE5FF6A070DFF7AF826" pageId="4" pageNumber="385">
<table id="538401ACFFEF001EFF6A070DFA3EF826" box="[151,1436,1876,1960]" gridcols="4" gridrows="3" pageId="4" pageNumber="385">
<tr id="9FB4F14EFFEF001EFF6A070DFA3EF8E5" box="[151,1436,1876,1899]" gridrow="0" pageId="4" pageNumber="385">
<th id="DC659832FFEF001EFF6A070DFE04F8E5" box="[151,422,1876,1899]" gridcol="0" gridrow="0" pageId="4" pageNumber="385">
<emphasis id="13F02F1EFFEFFFE5FF6A070DFEEAF8E7" bold="true" box="[151,328,1876,1899]" pageId="4" pageNumber="385">FIGURES 8–12.</emphasis>
Larva of
</th>
<emphasis id="13F02F1EFFEFFFE5FE50070CFDF8F8E7" box="[429,602,1876,1898]" italics="true" pageId="4" pageNumber="385">
<th id="DC659832FFEF001EFE50070DFDA9F8E5" box="[429,523,1876,1899]" gridcol="1" gridrow="0" pageId="4" pageNumber="385">
<taxonomicName id="E684888FFFEFFFE5FE50070CFDA9F8E4" box="[429,523,1877,1898]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="13F02F1EFFEFFFE5FE50070CFDA9F8E4" box="[429,523,1877,1898]" italics="true" pageId="4" pageNumber="385">Nemoura</emphasis>
</taxonomicName>
</th>
<th id="DC659832FFEF001EFDEF070DFBBAF8E5" box="[530,1048,1876,1899]" gridcol="2" gridrow="0" pageId="4" pageNumber="385">
<taxonomicName id="E684888FFFEFFFE5FDEF070DFDF8F8E7" box="[530,602,1876,1897]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="385" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEFFFE5FDEF070DFDF8F8E7" box="[530,602,1876,1897]" italics="true" pageId="4" pageNumber="385">papilla</emphasis>
</taxonomicName>
. 8. Chaetotaxy of terga 5–6. 9. Left cercus.
</th>
</emphasis>
<th id="DC659832FFEF001EFBE2070DFA3EF8E5" box="[1055,1436,1876,1899]" gridcol="3" gridrow="0" pageId="4" pageNumber="385">
10. Cercus segments 8−10 with
<tr id="9FB4F14EFFEF001EFF6A072AFA3EF807" box="[151,1436,1907,1929]" gridrow="1" pageId="4" pageNumber="385">
<th id="DC659832FFEF001EFF6A072AFE04F807" box="[151,422,1907,1929]" gridcol="0" gridrow="1" pageId="4" pageNumber="385">clubshaped bristles. 11. Cercus</th>
<td id="DC659832FFEF001EFE50072AFDA9F807" box="[429,523,1907,1929]" gridcol="1" gridrow="1" pageId="4" pageNumber="385">segments</td>
<td id="DC659832FFEF001EFDEF072AFBBAF807" box="[530,1048,1907,1929]" gridcol="2" gridrow="1" pageId="4" pageNumber="385">15 &amp;16 with long, acute bristles. 12. Apical cercus</td>
<td id="DC659832FFEF001EFBE2072AFA3EF807" box="[1055,1436,1907,1929]" gridcol="3" gridrow="1" pageId="4" pageNumber="385">segments. Scale (mm): 8, 10–12=0.1;</td>
</tr>
</th>
</tr>
<tr id="9FB4F14EFFEF001EFF6A07CBFA3EF826" box="[151,1436,1938,1960]" gridrow="2" pageId="4" pageNumber="385" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="DC659832FFEF001EFF6A07CBFE04F826" box="[151,422,1938,1960]" gridcol="0" gridrow="2" pageId="4" pageNumber="385">9=0.3.</th>
</tr>
</table>
</paragraph>
<subSubSection id="699EA087FFEEFFE4FF3A00CEFC46FE93" pageId="5" pageNumber="386" type="distribution">
<paragraph id="213BF30CFFEEFFE4FF3A00CEFC46FE93" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="386">
<emphasis id="13F02F1EFFEEFFE4FF3A00CEFEFBFF3E" bold="true" box="[199,345,151,176]" pageId="5" pageNumber="386">Distribution</emphasis>
.
<taxonomicName id="E684888FFFEEFFE4FE9700C0FD89FF3E" box="[362,555,152,176]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEEFFE4FE9700C0FD89FF3E" box="[362,555,152,176]" italics="true" pageId="5" pageNumber="386">Nemoura papilla</emphasis>
</taxonomicName>
is an East Asian mainland and insular, widely distributed species originally described from
<collectingCountry id="5993B39CFFEEFFE4FEBA00E4FE28FF5A" box="[327,394,189,212]" name="Japan" pageId="5" pageNumber="386">Japan</collectingCountry>
, but also known from the southern Kuril Islands,
<collectingRegion id="E3403DEEFFEEFFE4FC4800E5FBB5FF5B" box="[949,1047,188,213]" country="Russia" name="Sakhalin" pageId="5" pageNumber="386">Sakhalin</collectingRegion>
Island, mainland Russian Far East (RFE),
<collectingCountry id="5993B39CFFEEFFE4FF120086FE96FF76" box="[239,308,223,248]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
, and
<collectingCountry id="5993B39CFFEEFFE4FE8A00B9FE19FF79" box="[375,443,224,247]" name="South Korea" pageId="5" pageNumber="386">Korea</collectingCountry>
(Teslenko, 2015). The flight period extends from May to July. Larvae of
<taxonomicName id="E684888FFFEEFFE4FAD500B8FA39FF76" box="[1320,1435,224,248]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="papilla">
<emphasis id="13F02F1EFFEEFFE4FAD500B8FA39FF76" box="[1320,1435,224,248]" italics="true" pageId="5" pageNumber="386">N. papilla</emphasis>
</taxonomicName>
inhabit mountain streams and rivers, occurring in pools with silty substrate.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="699EA087FFEEFFE4FF6A0136FE95FE25" pageId="5" pageNumber="386" type="nomenclature">
<paragraph id="213BF30CFFEEFFE4FF6A0136FC79FE04" blockId="5.[151,987,367,427]" box="[151,987,367,394]" pageId="5" pageNumber="386">
<heading id="7A734460FFEEFFE4FF6A0136FC79FE04" bold="true" box="[151,987,367,394]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
<taxonomicName id="E684888FFFEEFFE4FF6A0136FC79FE04" ID-CoL="469YL" authority="Zhiltzova, 1997 in Teslenko &amp; Zhiltzova, 1997" authorityName="Zhiltzova, 1997 in Teslenko &amp; Zhiltzova" authorityYear="1997" box="[151,987,367,394]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFEEFFE4FF6A0136FC79FE04" bold="true" box="[151,987,367,394]" pageId="5" pageNumber="386">
<emphasis id="13F02F1EFFEEFFE4FF6A0136FE35FE07" bold="true" box="[151,407,367,393]" italics="true" pageId="5" pageNumber="386">Nemoura ussuriensis</emphasis>
<bibRefCitation id="45158EFDFFEEFFE4FE630129FDFDFE04" author="Zhiltzova" box="[414,607,368,394]" pageId="5" pageNumber="386" refString="Teslenko, V. A. &amp; Zhiltzova, L. A. (1997) New and little known species of stoneflies (Plecoptera: Nemouridae and Perlodidae) from the Russian Far East. Aquatic Insects, 19 (1), 23 - 35. http: // dx. doi. org / 10.1080 / 01650429709361633" type="journal article" year="1997">
Zhiltzova,
<quantity id="E67C5EE9FFEEFFE4FDD90129FDFDFE04" box="[548,607,368,394]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="5" pageNumber="386" unit="in" value="1997.0">1997</quantity>
</bibRefCitation>
in
<bibRefCitation id="45158EFDFFEEFFE4FD7B0129FC79FE04" author="Teslenko" box="[646,987,368,394]" pageId="5" pageNumber="386" refString="Teslenko, V. A. &amp; Zhiltzova, L. A. (1997) New and little known species of stoneflies (Plecoptera: Nemouridae and Perlodidae) from the Russian Far East. Aquatic Insects, 19 (1), 23 - 35. http: // dx. doi. org / 10.1080 / 01650429709361633" type="journal article" year="1997">Teslenko &amp; Zhiltzova, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="213BF30CFFEEFFE4FF6A01CBFE95FE25" blockId="5.[151,987,367,427]" box="[151,311,402,427]" pageId="5" pageNumber="386">
(
<figureCitation id="B9BFEF89FFEEFFE4FF6201CBFE88FE25" box="[159,298,402,427]" captionStart-0="FIGURES 13 – 14" captionStart-1="FIGURE 15 – 18" captionStart-2="FIGURES 19 – 23" captionStartId-0="6.[151,264,1952,1975]" captionStartId-1="7.[151,250,1518,1540]" captionStartId-2="8.[151,264,1901,1924]" captionTargetBox-0="[182,1417,201,1931]" captionTargetBox-1="[172,1406,193,1485]" captionTargetBox-2="[172,1402,339,1875]" captionTargetId-0="figure@6.[170,1417,193,1932]" captionTargetId-1="figure@7.[172,1407,193,1494]" captionTargetId-2="figure@8.[172,1412,339,1884]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 13 – 14. Larva of Nemoura ussuriensis, habitus, dorsal view, not to scale." captionText-1="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." captionText-2="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri-0="https://zenodo.org/record/256793/files/figure.png" httpUri-1="https://zenodo.org/record/256794/files/figure.png" httpUri-2="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Figs. 13−23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="699EA087FFEEFFE4FF6A0183FB9FF88D" pageId="5" pageNumber="386" type="materials_examined">
<paragraph id="213BF30CFFEEFFE4FF6A0183FB61FDD1" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="386">
<materialsCitation id="91ECF951FFEEFFE4FF6A0183FF49FDB5" ID-GBIF-Occurrence="1306554145" collectingDate="2004-04-27" collectorName="I. Tiunov" county="Material" latitude="43.407932" location="Anan'evka River" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" pageId="5" pageNumber="386" specimenCount="1" stateProvince="Primorsky">
<emphasis id="13F02F1EFFEEFFE4FF6A0183FE29FE7D" bold="true" box="[151,395,474,499]" pageId="5" pageNumber="386">
<collectingCounty id="C85A8B80FFEEFFE4FF6A0183FF5DFE7D" box="[151,255,474,499]" pageId="5" pageNumber="386">Material</collectingCounty>
examined.
</emphasis>
<collectingMunicipality id="C15F6976FFEEFFE4FE610183FDCDFE7C" box="[412,623,474,499]" pageId="5" pageNumber="386">Russian Far East</collectingMunicipality>
,
<collectingRegion id="E3403DEEFFEEFFE4FD780183FD5BFE7D" box="[645,761,474,499]" country="Russia" name="Primorskiy" pageId="5" pageNumber="386">Primorsky</collectingRegion>
Region: 4 mature larvae of males,
<location id="245BA5D7FFEEFFE4FB310183FA3AFE7D" LSID="urn:lsid:plazi:treatment:A92D421AFFEEFFE9FF6A0136FCFCFE92:245BA5D7FFEEFFE4FB310183FA3AFE7D" box="[1228,1432,474,499]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="Anan'evka River" pageId="5" pageNumber="386" stateProvince="Primorsky">Anan’evka River</location>
,
<location id="245BA5D7FFEEFFE4FF6A01A7FECEFD99" LSID="urn:lsid:plazi:treatment:A92D421AFFEEFFE9FF6A0136FCFCFE92:245BA5D7FFEEFFE4FF6A01A7FECEFD99" box="[151,364,510,535]" county="Material" latitude="43.407932" longLatPrecision="1" longitude="131.73837" municipality="Russian Far East" name="Razdol'naya River" pageId="5" pageNumber="386" stateProvince="Primorsky">Razdol’naya River</location>
. Basin, near road bridge, at Venevitinovo Sett.,
<geoCoordinate id="44B095CBFFEEFFE4FC7F01A6FBBAFD99" box="[898,1048,510,535]" direction="north" orientation="latitude" pageId="5" pageNumber="386" precision="1" value="43.407932">N 43º24.476’</geoCoordinate>
<geoCoordinate id="44B095CBFFEEFFE4FBDD01A6FB1EFD99" box="[1056,1212,511,535]" direction="east" orientation="longitude" pageId="5" pageNumber="386" precision="1" value="131.73837">E131º44.302’</geoCoordinate>
,
<date id="553AD5CCFFEEFFE4FB3501A6FAEBFD99" box="[1224,1353,511,535]" pageId="5" pageNumber="386" value="2004-04-27">
<collectingDate id="457E2C24FFEEFFE4FB3501A6FAEBFD99" box="[1224,1353,511,535]" pageId="5" pageNumber="386" value="2004-04-27">27.04.2004</collectingDate>
</date>
, coll.
<collectorName id="8C7196DAFFEEFFE4FA7101A6FF49FDB5" pageId="5" pageNumber="386">I. Tiunov</collectorName>
</materialsCitation>
;
<materialsCitation id="91ECF951FFEEFFE4FF0B027AFF49FDD1" ID-GBIF-Occurrence="1306554147" collectorName="I. Tiunov" location="Razdol'naya River" municipality="Vtoraya Rechka River" pageId="5" pageNumber="386" specimenCount="2" specimenCount-larva="2" stateProvince="Primorsky">
<specimenCount id="37823885FFEEFFE4FF0B027AFEF2FDB5" box="[246,336,546,571]" pageId="5" pageNumber="386" type="larva">2 larvae</specimenCount>
of females,
<collectingMunicipality id="C15F6976FFEEFFE4FE27027AFD75FDB5" box="[474,727,546,571]" pageId="5" pageNumber="386">Vtoraya Rechka River</collectingMunicipality>
,
<location id="245BA5D7FFEEFFE4FD1F027BFC1AFDB5" LSID="urn:lsid:plazi:treatment:A92D421AFFEEFFE9FF6A0136FCFCFE92:245BA5D7FFEEFFE4FD1F027BFC1AFDB5" box="[738,952,546,571]" municipality="Vtoraya Rechka River" name="Razdol'naya River" pageId="5" pageNumber="386" stateProvince="Primorsky">Razdol’naya River</location>
. Basin, near road bridge, 3.05.2013, coll.
<collectorName id="8C7196DAFFEEFFE4FA71027AFF49FDD1" pageId="5" pageNumber="386">I. Tiunov</collectorName>
</materialsCitation>
;
<materialsCitation id="91ECF951FFEEFFE4FF08021EFB1DFDD1" ID-GBIF-Occurrence="1306554144" box="[245,1215,582,607]" collectingDate="2013-04-26" collectorName="I. Tiunov" location="Gladkaya River" pageId="5" pageNumber="386" specimenCount="1" stateProvince="Primorsky">
1 mature male larva,
<location id="245BA5D7FFEEFFE4FE1E021FFD37FDD1" LSID="urn:lsid:plazi:treatment:A92D421AFFEEFFE9FF6A0136FCFCFE92:245BA5D7FFEEFFE4FE1E021FFD37FDD1" box="[483,661,582,607]" name="Gladkaya River" pageId="5" pageNumber="386" stateProvince="Primorsky">Gladkaya River</location>
, near Gvozdevo Sett.,
<date id="553AD5CCFFEEFFE4FC6F021FFBB1FDD1" box="[914,1043,582,607]" pageId="5" pageNumber="386" value="2013-04-26">
<collectingDate id="457E2C24FFEEFFE4FC6F021FFBB1FDD1" box="[914,1043,582,607]" pageId="5" pageNumber="386" value="2013-04-26">26.04.2013</collectingDate>
</date>
, coll.
<collectorName id="8C7196DAFFEEFFE4FBAB021EFB1DFDD1" box="[1110,1215,582,607]" pageId="5" pageNumber="386">I. Tiunov</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="213BF30CFFEEFFE4FF3A0233FABAFB81" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="386">
<emphasis id="13F02F1EFFEEFFE4FF3A0233FECDFD0D" bold="true" box="[199,367,618,643]" pageId="5" pageNumber="386">Mature larva.</emphasis>
Mature larvae are large, body length
<quantity id="E67C5EE9FFEEFFE4FCDB0232FC0DFD0C" box="[806,943,619,643]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="7.7" metricValueMin="7.2" pageId="5" pageNumber="386" unit="mm" value="7.45" valueMax="7.7" valueMin="7.2">7.2–7.7 mm</quantity>
in males;
<quantity id="E67C5EE9FFEEFFE4FBD70233FB16FD0C" box="[1066,1204,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.6" metricValueMin="8.3" pageId="5" pageNumber="386" unit="mm" value="8.95" valueMax="9.6" valueMin="8.3">8.3–9.6 mm</quantity>
in females. General color brown with conspicuous head pattern: tentorial callosities brown, contoured by narrow dark bands extending to the clypeus, resembling the Russian letter П with mesal notch; clypeus with prominent rounded anterior corners. The interocellar area bears a diffuse dark spot attaining the posterior ocelli (
<figureCitation id="B9BFEF89FFEEFFE4FC13028FFBDAFD61" box="[1006,1144,726,751]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[182,1417,201,1931]" captionTargetId="figure@6.[170,1417,193,1932]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Larva of Nemoura ussuriensis, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256793/files/figure.png" pageId="5" pageNumber="386">Figs. 13, 14</figureCitation>
). Occiput with three pairs of pale spots surrounded by thread-like brown loops, posterior margin darkened. Antenna brown, long, up to 85−100% of body length, scape paler than both head or pedicle and flagellum. Pronotum approximately 1.5 X wider than long, pattern indistinct, with small dark brown spots together resembling the letter Х (
<figureCitation id="B9BFEF89FFEEFFE4FAFC031BFA2DFCD5" box="[1281,1423,834,859]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[182,1417,201,1931]" captionTargetId="figure@6.[170,1417,193,1932]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Larva of Nemoura ussuriensis, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256793/files/figure.png" pageId="5" pageNumber="386">Figs. 13, 14</figureCitation>
). Mesonotum and metanotum with U-shaped brown band and diffuse pale patch anteromedially. Legs brown, outer margin of femora darker than inner, the fore femur is 2.6 X longer than wide (
<figureCitation id="B9BFEF89FFEEFFE4FC0003D3FBEDFC2D" box="[1021,1103,906,931]" captionStart="FIGURE 15 – 18" captionStartId="7.[151,250,1518,1540]" captionTargetBox="[172,1406,193,1485]" captionTargetId="figure@7.[172,1407,193,1494]" captionTargetPageId="7" captionText="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="5" pageNumber="386">Fig. 18</figureCitation>
), the hind femur 3.3 X longer than wide, hind legs are long. Abdomen relatively slender, integument light, matte in appearance, terga brown with a mesal row of dark brown patches which forms a longitudinal band on 1−8 segments (
<figureCitation id="B9BFEF89FFEEFFE4FB82038BFAA9FC65" box="[1151,1291,978,1003]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[182,1417,201,1931]" captionTargetId="figure@6.[170,1417,193,1932]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Larva of Nemoura ussuriensis, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256793/files/figure.png" pageId="5" pageNumber="386">Figs. 13, 14</figureCitation>
). Cerci pale, strongly hairy, long, with 31−32 segments, length reaches 100−118% of body length (
<figureCitation id="B9BFEF89FFEEFFE4FBA403AFFB42FB81" box="[1113,1248,1014,1039]" captionStart="FIGURES 13 – 14" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[182,1417,201,1931]" captionTargetId="figure@6.[170,1417,193,1932]" captionTargetPageId="6" captionText="FIGURES 13 – 14. Larva of Nemoura ussuriensis, habitus, dorsal view, not to scale." httpUri="https://zenodo.org/record/256793/files/figure.png" pageId="5" pageNumber="386">Figs. 13, 14</figureCitation>
,
<figureCitation id="B9BFEF89FFEEFFE4FB1103AEFAAEFB81" box="[1260,1292,1015,1039]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">20</figureCitation>
).
</paragraph>
<paragraph id="213BF30CFFEEFFE4FF3A0443FB9FF88D" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="386">
Setation distinct. Setal fringe around the pronotum consisting of occasional long hairs, and cylindrical bristles, the apices rounded and often slightly wider than the base; length of bristles on lateral margins and anterior and posterior corners greater than on anterior and posterior margins; fringe on lateral pronotal margins relatively regular and denser than on anterior and posterior margins, because of bristles stand in two or three rows (
<figureCitation id="B9BFEF89FFEEFFE4FAC404DFFF14FB4D" captionStart="FIGURE 15 – 18" captionStartId="7.[151,250,1518,1540]" captionTargetBox="[172,1406,193,1485]" captionTargetId="figure@7.[172,1407,193,1494]" captionTargetPageId="7" captionText="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="5" pageNumber="386">Figs. 15, 16</figureCitation>
). Length of the longest cylindrical bristles at the anterior and posterior pronotal corners reaches 3.4% of pronotal width. Pronotal disc with occasional long hairs, short cylindrical setae and tiny procumbent setae with short acute hooked apices. These tiny setae are concentrated on the lateral fields of pronotum (
<figureCitation id="B9BFEF89FFEEFFE4FB4B04ABFAABFA85" box="[1206,1289,1266,1291]" captionStart="FIGURE 15 – 18" captionStartId="7.[151,250,1518,1540]" captionTargetBox="[172,1406,193,1485]" captionTargetId="figure@7.[172,1407,193,1494]" captionTargetPageId="7" captionText="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="5" pageNumber="386">Fig. 16</figureCitation>
). Outer wing pad margin covered with slender club-shaped bristles, and occasional long hairs; cylindrical setae and occasional long hairs in the lines on the wing-pads (
<figureCitation id="B9BFEF89FFEEFFE4FDA00563FD12FADD" box="[605,688,1338,1363]" captionStart="FIGURE 15 – 18" captionStartId="7.[151,250,1518,1540]" captionTargetBox="[172,1406,193,1485]" captionTargetId="figure@7.[172,1407,193,1494]" captionTargetPageId="7" captionText="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="5" pageNumber="386">Fig. 17</figureCitation>
). All femora bear fine numerous procumbent setae and dense short blunt bristles. Setal fringe on the outer femur margin short, including a few long thin hairs and short blunt bristles in irregular arrangement (
<figureCitation id="B9BFEF89FFEEFFE4FE4105DBFDB3FA15" box="[444,529,1410,1435]" captionStart="FIGURE 15 – 18" captionStartId="7.[151,250,1518,1540]" captionTargetBox="[172,1406,193,1485]" captionTargetId="figure@7.[172,1407,193,1494]" captionTargetPageId="7" captionText="FIGURE 15 – 18. Larva of Nemoura ussuriensis. 15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15 – 17 = 0.1; 18 = 0.3." httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="5" pageNumber="386">Fig. 18</figureCitation>
); the longest bristle reaches approximately 22% of femur width on the fore leg and 20% on the hind leg. Setation of tibia and tarsi very similar to femur. Terga covered mainly with short cylindrical bristles and sparse tiny procumbent setae (
<figureCitation id="B9BFEF89FFEEFFE4FCC70593FC35FA6D" box="[826,919,1482,1507]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Fig. 19</figureCitation>
). Setae along the posterior tergal margin heterogeneous: most apically rounded cylindrical bristles are relatively short (
<figureCitation id="B9BFEF89FFEEFFE4FBFC05B7FBF6F989" box="[1025,1108,1518,1543]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Fig. 19</figureCitation>
), while one conspicuous pair of long bristles turns apically into fine, thin hairs that are easily overlooked; on the posterior margin of terga 5−6 hairs reach 47% of segment length (
<figureCitation id="B9BFEF89FFEEFFE4FDCD066FFD19F9C1" box="[560,699,1590,1615]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Figs. 19, 23</figureCitation>
). Cerci densely covered with acute bristles of different length: in apical whorl on basal cercal segments the bristles shorter the segment; on cercal segment 8 the longest acute bristles reach 148% of segment’s length, intercalary setation visible (
<figureCitation id="B9BFEF89FFEEFFE4FC650627FC4EF919" box="[920,1004,1662,1687]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Fig. 21</figureCitation>
); in cercal segments 16–17 the longest bristles do not exceed 95% of segment length, the intercalary setae reach 128% of the segment’s length (
<figureCitation id="B9BFEF89FFEEFFE4FAC606FBFA32F935" box="[1339,1424,1698,1723]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="5" pageNumber="386">Fig. 22</figureCitation>
). Setal ring of apical cercal segments sparse, intercalary hairs noticeable. Additionally to the long thin acute bristles there are tiny setae with rounded apex and a few thin long hairs in the apical whorl.
</paragraph>
</subSubSection>
<subSubSection id="699EA087FFEEFFE6FF3A0757FEFEF8B8" lastPageId="7" lastPageNumber="388" pageId="5" pageNumber="386" type="diagnosis">
<paragraph id="213BF30CFFEEFFE6FF3A0757FEFEF8B8" blockId="5.[151,1437,474,2011]" lastBlockId="7.[151,1436,1641,2026]" lastPageId="7" lastPageNumber="388" pageId="5" pageNumber="386">
<emphasis id="13F02F1EFFEEFFE4FF3A0757FEE3F8A9" bold="true" box="[199,321,1806,1831]" pageId="5" pageNumber="386">Diagnosis.</emphasis>
The larvae of
<taxonomicName id="E684888FFFEEFFE4FE1A0756FD27F8A8" box="[487,645,1807,1830]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFEEFFE4FE1A0756FD27F8A8" box="[487,645,1807,1830]" italics="true" pageId="5" pageNumber="386">N. ussuriensis</emphasis>
</taxonomicName>
can be easily distinguished from the known larvae of
<taxonomicName id="E684888FFFEEFFE4FB110756FAF6F8A8" box="[1260,1364,1807,1830]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="13F02F1EFFEEFFE4FB110756FAF6F8A8" box="[1260,1364,1807,1830]" italics="true" pageId="5" pageNumber="386">Nemoura</emphasis>
</taxonomicName>
in the RFE by the head pattern, long and the “fuzzy-like” appearance of the cerci exceeding the body length with distinctive setation. Setal fringe around pronotum, outer wing pad margin and posterior tergal margins consisting of cylindrical bristles with apices rounded and often a little wider than the base, but not club-shaped. Fringe on lateral pronotal margins regular and dense, cylindrical bristles placed in two or three rows, tiny procumbent setae on pronotal lateral fields with short acute hooked apexes. The outer femur margin bears bristles of another
<typeStatus id="FE3F4DAEFFEEFFE4FADC079AFAF6F855" box="[1313,1364,1987,2011]" pageId="5" pageNumber="386">type</typeStatus>
, they are short and blunt. The posterior tergal margins 5–6 feature conspicuous paired long, thin cylindrical bristles (like hairs) reaching 47% of the length of segment (
<figureCitation id="B9BFEF89FFECFFE6FD5D06D4FC85F928" box="[672,807,1677,1702]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[172,1402,339,1875]" captionTargetId="figure@8.[172,1412,339,1884]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Larva of Nemoura ussuriensis. 19. Chaetotaxy of terga 5 – 6. 20. Left cercus. 21. Cercus segments 8 − 11 with acute bristles in whorl cercus and acute intercalary bristles. 22. Cercus segments 16 &amp; 17 with long, acute bristles in cercus whorl and intercalary bristles. 23. Cylindrical bristles and one of the paired long slightly curved thin bristle on the posterior margin of terga 5 − 6. Scale (mm): 19, 21, 22 = 0.1; 20 = 0.3; 23 = 0.05." httpUri="https://zenodo.org/record/256795/files/figure.png" pageId="7" pageNumber="388">Figs. 19, 23</figureCitation>
) among the mostly short cylindrical bristles. Cerci have homogeneous setation of long acute bristles. In the apical whorl of cercal segment 8 the longest acute bristles reach 148% of the segment’s length, intercalary setation visible; in apical whorl of 16−17 cercal segments the longest acute bristles do not exceed 95% of the segment’s length and the longest intercalary setae reach 128% of the segment’s length.
</paragraph>
</subSubSection>
<caption id="75FBA384FFEDFFE7FF6A07F9FC4BF839" box="[151,1001,1952,1975]" httpUri="https://zenodo.org/record/256793/files/figure.png" pageId="6" pageNumber="387" targetBox="[182,1417,201,1931]" targetPageId="6">
<paragraph id="213BF30CFFEDFFE7FF6A07F9FC4BF839" blockId="6.[151,1001,1952,1975]" box="[151,1001,1952,1975]" pageId="6" pageNumber="387">
<emphasis id="13F02F1EFFEDFFE7FF6A07F9FEF1F838" bold="true" box="[151,339,1952,1975]" pageId="6" pageNumber="387">FIGURES 13–14.</emphasis>
Larva of
<taxonomicName id="E684888FFFEDFFE7FE4B07FBFD29F839" box="[438,651,1954,1975]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFEDFFE7FE4B07FBFD29F839" box="[438,651,1954,1975]" italics="true" pageId="6" pageNumber="387">Nemoura ussuriensis</emphasis>
</taxonomicName>
, habitus, dorsal view, not to scale.
</paragraph>
</caption>
<caption id="75FBA384FFECFFE6FF6A05B7FAE7F9CC" httpUri="https://zenodo.org/record/256794/files/figure.png" pageId="7" pageNumber="388" targetBox="[172,1406,193,1485]" targetPageId="7">
<paragraph id="213BF30CFFECFFE6FF6A05B7FAE7F9CC" blockId="7.[151,1437,1518,1602]" pageId="7" pageNumber="388">
<emphasis id="13F02F1EFFECFFE6FF6A05B7FEE4F98A" bold="true" box="[151,326,1518,1540]" pageId="7" pageNumber="388">FIGURE 15–18.</emphasis>
Larva of
<emphasis id="13F02F1EFFECFFE6FE5605B6FD2AF98A" box="[427,648,1519,1540]" italics="true" pageId="7" pageNumber="388">
<taxonomicName id="E684888FFFECFFE6FE5605B6FD26F98A" box="[427,644,1519,1540]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="ussuriensis">Nemoura ussuriensis</taxonomicName>
.
</emphasis>
15. Cylindrical bristles with rounded apices on anterior margin of pronotum. 16. Chaetotaxy on disc, right anterior pronotal corner and lateral margin with dense cylindrical bristles. 17. Cylindrical and club-shaped bristles on outer margin and in the rows on wing pad. 18. Right fore leg. Scale (mm): 15–17=0.1; 18=0.3.
</paragraph>
</caption>
<subSubSection id="699EA087FFECFFE9FF3A0718FCFCFE92" lastPageId="8" lastPageNumber="389" pageId="7" pageNumber="388" type="distribution">
<paragraph id="213BF30CFFECFFE9FF3A0718FCFCFE92" blockId="7.[151,1436,1641,2026]" lastBlockId="8.[151,1436,151,285]" lastPageId="8" lastPageNumber="389" pageId="7" pageNumber="388">
<emphasis id="13F02F1EFFECFFE6FF3A0718FEC2F8D4" bold="true" box="[199,352,1857,1882]" pageId="7" pageNumber="388">Distribution.</emphasis>
<taxonomicName id="E684888FFFECFFE6FE97071BFDFBF8D7" box="[362,601,1858,1881]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFECFFE6FE97071BFDFBF8D7" box="[362,601,1858,1881]" italics="true" pageId="7" pageNumber="388">Nemoura ussuriensis</emphasis>
</taxonomicName>
is currently considered an East Asian mainland endemic species with a restricted distribution south of
<collectingRegion id="E3403DEEFFECFFE6FE06073CFDCDF8F0" box="[507,623,1893,1918]" country="Russia" name="Primorskiy" pageId="7" pageNumber="388">Primorsky</collectingRegion>
Region of the RFE in the Razdol’naya River Basin and small salmon streams flowing into Peter the Great Bay of the Sea of
<collectingCountry id="5993B39CFFECFFE6FCD907D3FCC6F82F" box="[804,868,1930,1953]" name="Japan" pageId="7" pageNumber="388">Japan</collectingCountry>
(
<bibRefCitation id="45158EFDFFECFFE6FC8807D0FB18F82C" author="Teslenko" box="[885,1210,1929,1954]" pageId="7" pageNumber="388" refString="Teslenko, V. A. &amp; Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs. Dalnauka, Vladivostok, 382 pp." type="book" year="2009">Teslenko &amp; Zhiltzova 2009</bibRefCitation>
).
<bibRefCitation id="45158EFDFFECFFE6FB3207D0FACEF82C" author="Zwick" box="[1231,1388,1929,1954]" pageId="7" pageNumber="388" refString="Zwick, P. (2010) New species and new records of Plecoptera from Korea and the Russian Far East. Illiesia, 6 (9), 75 - 97. Avaliable from: http: // www 2. pms-lj. si / illiesia / Illiesia 06 - 09. pdf (Accessed 7 Sept. 2016)" type="journal article" year="2010">Zwick (2010)</bibRefCitation>
has suggested that the description of the Korean species,
<taxonomicName id="E684888FFFECFFE6FD1207F7FB8DF848" authority="Ham &amp; Lee, 1999" authorityName="Ham &amp; Lee" authorityYear="1999" box="[751,1071,1966,1990]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="espera">
<emphasis id="13F02F1EFFECFFE6FD1207F7FCA0F84B" box="[751,770,1966,1989]" italics="true" pageId="7" pageNumber="388">N</emphasis>
.
<emphasis id="13F02F1EFFECFFE6FCEC07F7FCFEF84B" box="[785,860,1966,1989]" italics="true" pageId="7" pageNumber="388">espera</emphasis>
Ham &amp; Lee, 1999
</taxonomicName>
is very similar to
<taxonomicName id="E684888FFFECFFE6FB0307F7FA39F84B" box="[1278,1435,1966,1989]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFECFFE6FB0307F7FA39F84B" box="[1278,1435,1966,1989]" italics="true" pageId="7" pageNumber="388">N. ussuriensis</emphasis>
</taxonomicName>
and may be a junior synonym. Certainly, this suggestion requires further study of comparative material. However, it seems that
<taxonomicName id="E684888FFFE3FFE9FEC900C0FE76FF21" box="[308,468,152,176]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="13F02F1EFFE3FFE9FEC900C0FE76FF21" box="[308,468,152,176]" italics="true" pageId="8" pageNumber="389">N. ussuriensis</emphasis>
</taxonomicName>
should be encountered in streams of
<collectingCountry id="5993B39CFFE3FFE9FC7300CEFC73FF3E" box="[910,977,151,176]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
and the Korean Peninsula, because the transboundary Razdol’naya River flows through
<collectingCountry id="5993B39CFFE3FFE9FD4700E5FD5FFF5B" box="[698,765,188,213]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
and
<collectingCountry id="5993B39CFFE3FFE9FCCF00E5FCDDFF5B" box="[818,895,188,213]" name="Russia" pageId="8" pageNumber="389">Russia</collectingCountry>
, and small salmon streams originate in the spurs of the Black Mountains of the Changbai Mountain Range shared with
<collectingCountry id="5993B39CFFE3FFE9FC250086FBB9FF76" box="[984,1051,223,248]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
and
<collectingCountry id="5993B39CFFE3FFE9FBA700B9FB3CFF79" box="[1114,1182,224,247]" name="South Korea" pageId="8" pageNumber="389">Korea</collectingCountry>
Pennisula. The flight period extended from the end of April to the beginning of June.
</paragraph>
</subSubSection>
</treatment>
</document>

104
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@ -0,0 +1,104 @@
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<mods:namePart>Rosa, Paolo</mods:namePart>
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<mods:namePart>Vardal, Hege</mods:namePart>
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<treatment ID-GBIF-Taxon="152059382" LSID="urn:lsid:plazi:treatment:A92D58A07F5325FCD337FEA26DC86222" httpUri="http://treatment.plazi.org/id/A92D58A07F5325FCD337FEA26DC86222" lastPageNumber="82" pageId="3" pageNumber="82">
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<taxonomicName LSID="A92D58A0-7F53-25FC-D337-FEA26DC86222" authority="Bohart, 1982" authorityName="Bohart" authorityYear="1982" class="Insecta" family="Chrysididae" genus="Argochrysis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Argochrysis albicornis" order="Hymenoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="albicornis">Argochrysis albicornis Bohart, 1982</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="reference_group">
<paragraph pageId="3" pageNumber="82">
<taxonomicName authorityName="Bohart" authorityYear="1982" class="Insecta" family="Chrysididae" genus="Argochrysis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Argochrysis albicornis" order="Hymenoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="albicornis">Argochrysis albicornis</taxonomicName>
: Bohart (in Bohart &amp; Kimsey)
<bibRefCitation author="Bohart, RM" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="43" pageNumber="122" refId="B10" refString="Bohart, RM, Kimsey, LS, 1982. A synopsis of the Chrysididae in America North of Mexico. Memoirs of the American Entomological Institute 33" title="A synopsis of the Chrysididae in America North of Mexico." volume="33" year="1982">1982</bibRefCitation>
: 189.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="type locality">
<paragraph pageId="3" pageNumber="82">Type locality.</paragraph>
<paragraph pageId="3" pageNumber="82">U.S.A. (holotype from Borrego Valley, San Diego Co., California; paratypes: 44 ♂♂ and 58 ♀♀ form California and Nevada).</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="paratype 1 ♂">
<paragraph pageId="3" pageNumber="82">Paratype 1 ♂.</paragraph>
<paragraph pageId="3" pageNumber="82">
[1,000 Palms Cyn., Cal. Riverside Co. IV-9-1964] [R.M. Bohart collector] [Paratype
<taxonomicName authorityName="Bohart" authorityYear="1982" class="Insecta" family="Chrysididae" genus="Argochrysis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Argochrysis albicornis" order="Hymenoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="albicornis">Argochrysis albicornis</taxonomicName>
♂
<emphasis italics="true" pageId="3" pageNumber="82">R.</emphasis>
M. Bohart] &lt;red label&gt; [NHRS-HEVA000001057].
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="paratype 1 ♀">
<paragraph pageId="3" pageNumber="82">Paratype 1 ♀.</paragraph>
<paragraph pageId="3" pageNumber="82">
[Calif
<emphasis italics="true" pageId="3" pageNumber="82">2</emphasis>
mi
<emphasis italics="true" pageId="3" pageNumber="82">E</emphasis>
Lone Pine Inyo Co. V-19-1970] [E.E. Grissell Colr] [Paratype
<taxonomicName authorityName="Bohart" authorityYear="1982" class="Insecta" family="Chrysididae" genus="Argochrysis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Argochrysis albicornis" order="Hymenoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="albicornis">Argochrysis albicornis</taxonomicName>
♀ R.M. Bohart] &lt;red label&gt; [NHRS-HEVA000001058].
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="remarks">
<paragraph pageId="3" pageNumber="82">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="82">The holotype is deposited at the BME.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="82" type="current status">
<paragraph pageId="3" pageNumber="82">Current status.</paragraph>
<paragraph pageId="3" pageNumber="82">
<taxonomicName authorityName="Bohart" authorityYear="1982" class="Insecta" family="Chrysididae" genus="Argochrysis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Argochrysis albicornis" order="Hymenoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="albicornis">
<emphasis italics="true" pageId="3" pageNumber="82">Argochrysis albicornis</emphasis>
</taxonomicName>
Bohart, 1982.
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Figs 82
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22
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<paragraph pageId="260" pageNumber="261">Type locality.</paragraph>
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FRENCH GUIANA: Montagne des Chevaux [
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,
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].
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<paragraph pageId="260" pageNumber="261">Type material.</paragraph>
<paragraph pageId="260" pageNumber="261">
Holotype female: &quot;GUYANE
<normalizedToken originalValue="FRANÇAISE">FRANCAISE</normalizedToken>
: Montagne des Chevaux,
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22.i.2011, SEAG leg.&quot;/ &quot;Caterino/Tishechkin Exosternini Voucher EXO-02501&quot; (MNHN). Paratypes (2): FRENCH GUIANA: 1:
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de
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,
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,
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,14.ii.2011, FIT, SEAG (CHND). 1: GUYANA: Region 8, Iwokrama Forest, Kabocalli Field Stn., 60m,
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,
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, 3-5.vi.2001, R. Brooks, Z. Falin (SEMC).
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<paragraph pageId="260" pageNumber="261">Diagnostic description.</paragraph>
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Length: 2.0-2.2mm, width: 1.7-1.9mm; body elongate oval, sides subparallel, weakly depressed, glabrous; pronotum and elytra uniformly metallic violet-bronze, head, pygidia and venter rufo-piceous; frons weakly elevated above antennal bases, slightly depressed at middle, interocular margins strongly convergent dorsad, ground punctation fine, secondary punctures small, sparse, slightly
<pageBreakToken pageId="261" pageNumber="262" start="start">denser</pageBreakToken>
toward vertex, frontal stria present along inner margin of eye, curving mediad but absent across front; supraorbital stria absent; epistoma weakly convex along apical margin; labrum about 4
<normalizedToken originalValue="×wider">xwider</normalizedToken>
than long, weakly bisinuate apically; each mandible with acute basal tooth; antennal scape short, club asymmetrically oblong; pronotal sides subparallel in basal half, abruptly arcuate to apex, depressed in anterior corners, marginal stria complete along lateral and anterior margins, slightly crenulate in front, lateral submarginal stria absent, ground punctation of pronotal disk very fine, secondary punctures uniformly interspersed in lateral thirds; elytra with 2-3 complete epipleural striae, the outer variably abbreviated apically, outer subhumeral stria absent, inner subhumeral stria present as short basal fragment, dorsal striae 1-2 more or less complete, 3rd stria nearly complete to significantly abbreviated apically, 4th stria ab
<pageBreakToken pageId="262" pageNumber="263" start="start">breviated</pageBreakToken>
from apex, but generally more nearly complete than 3rd stria, 5th stria absent, sutural stria present in apical two-thirds, elytral disk with few coarse punctures in apical fourth; prosternum rather narrow, distinctly convex, keel shallowly emarginate at base, carinal striae complete, convergent at middle but separate throughout; prosternal lobe about half keel length, apical margin broadly rounded, marginal stria nearly complete, faintly obsolete at sides; mesoventrite sinuate to weakly produced, marginal stria complete, mesometaventral stria weakly arched forward, crenulate, continuous at sides with lateral metaventral stria, which curves posterolaterad toward middle of metacoxa, outer lateral metaventral stria present, short, divergent from base of inner stria, metaventral disk impunctate at middle; abdominal ventrite 1 with single lateral stria more or less complete, middle portion of disk impunctate; protibia 4-5 dentate, basal denticles weak, outer margin serrulate between teeth; mesotibia with two marginal spines; outer metatibial margin smooth; propygidium without basal stria, with ground punctation very fine, inconspicuous, discal punctures rather small, ocellate, separated by their diameters or slightly less in basal half, smaller and sparser apically; propygidial gland openings conspicuous nearly one-half from anterior and one-fourth from lateral margins; pygidium with ground punctation fine, secondary punctures small, very uniformly interspersed throughout. Male: not known.
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<paragraph pageId="262" pageNumber="263">
Figure 82.
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incertae sedis spp. A Dorsal habitus of
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B Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Baconia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baconia purpurata" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="purpurata">Baconia purpurata</taxonomicName>
C Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Baconia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baconia aenea" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="aenea">Baconia aenea</taxonomicName>
D Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Baconia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baconia aenea" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="aenea">Baconia aenea</taxonomicName>
E Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Baconia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baconia clemens" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="clemens">Baconia clemens</taxonomicName>
F Frons of male
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.
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<paragraph pageId="262" pageNumber="263">Remarks.</paragraph>
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This
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color pattern is distinctive, with strongly violet metallic pronotum and elytra (Fig. 82A), but with the head and pygidium similarly colored to non-metallic venter. It is similar and apparently closely related to
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, which is much more commonly collected in French Guiana - both lack the frontal stria, and may be generally violet in color. However,
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is more richly violet in color, lacks the transverse basal propygidial stria, has a narrower frons, a rather small labrum (Fig. 82B), and lacks a basal arch representing the 5th elytral stria.
</paragraph>
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<paragraph pageId="262" pageNumber="263">Etymology.</paragraph>
<paragraph pageId="262" pageNumber="263">
This
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name refers to its metallic violet coloration.
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Walker, 1837
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<mods:namePart id="DCCE7627D7DA2D6314A09ABC90E493AF">Helen K. Larson</mods:namePart>
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<treatment id="A92DCF0694E4C7E790987AE1F2B833CE" ID-DOI="http://doi.org/10.5281/zenodo.6264613" ID-GBIF-Taxon="100121598" ID-Zenodo-Dep="6264613" LSID="urn:lsid:plazi:treatment:A92DCF0694E4C7E790987AE1F2B833CE" httpUri="http://treatment.plazi.org/id/A92DCF0694E4C7E790987AE1F2B833CE" lastPageNumber="4" pageNumber="1">
<subSubSection id="120EDEFAAC189A892C7E0FD7FE6695F7" type="nomenclature">
<paragraph id="B346F3622434BB8609140C7A3E81EF04">
[[ Genus
<taxonomicName id="E61AE691277FB4D5F08BD51A24006288" ID-CoL="62V3V" ID-ENA="2067727" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
]]
</paragraph>
</subSubSection>
<subSubSection id="0CB2A2EB2AEB38F0F0107DD3445DE745" type="discussion">
<paragraph id="B156E30B5459B2F0C53291387F2783F5" pageNumber="1">
The gobiid genus
<taxonomicName id="16398D3137C1623AAC6616BEF2FA840D" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
is speciose in the western Indian Ocean. Smith (1958) placed a number of species in
<taxonomicName id="A6FB02F36C37EAA421B951F362DFAD89" LSID-ZBK="urn:lsid:zoobank.org:act:E59F9F13-515E-4908-BF3D-E8A0D425C382" family="Gobiidae" genus="Monishia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Monishia Smith 1959:206" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Monishia</taxonomicName>
or in various genera, now regarded by Akihito &amp; Meguro (1981) and Hoese (1986) as junior synonyms of
<taxonomicName id="FCC4FE30B9E48222649CF373ED0AC0C1" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
. Hoese &amp; Winterbottom (1979) treated some species from South Africa, while Hoese (1986) revised the genus and recognized 13 species and synonymized several nominal genera. Subsequently Gill (1998) described an additional species from Mauritius. The present paper describes two new species from the south Pacific and redescribes
<taxonomicName id="6606D44BE1CE24FCCFD7B4A23403CB34" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="caudilinea">Hetereleotris caudilinea (Randall)</taxonomicName>
, in order to compare its characteristics with the two new taxa. Previously only a single species of
<taxonomicName id="22E438A34099797F56A597295B291667" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
had been known from outside the Indian Ocean.
</paragraph>
<paragraph id="C73BD75ACD1C4C38B7EE413C77A6BBBE" pageNumber="2">
Tentatively the species are placed here in
<taxonomicName id="7351615433E1CDB438A62F9F56BB2657" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
based on the presence of 27 vertebrae and the closure of the first gill slit. Other genera with 27 vertebrae from the Indo-Pacific region include
<taxonomicName id="D427E282108AFABEF653B670BA780BF8" LSID-ZBK="urn:lsid:zoobank.org:act:ADAF83BF-42AD-4A07-8D23-0365879A2423" family="Gobiidae" genus="Psammogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Psammogobius Smith 1935:215" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Psammogobius</taxonomicName>
,
<taxonomicName id="1C6D7B6E8617D82422B409B56DD2A5EA" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
,
<taxonomicName id="671FD70EB70F6B07B98EC6B3D12532F0" LSID-ZBK="urn:lsid:zoobank.org:act:E608AAC1-2634-4D90-B4F5-3F6F4BD78FBB" family="Gobiidae" genus="Bathygobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bathygobius Bleeker 1878:54" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Bathygobius</taxonomicName>
,
<taxonomicName id="34C2F5D3DE9396A18199B6FCC00E0B2D" LSID-ZBK="urn:lsid:zoobank.org:act:97FDC767-1FD7-483E-8184-8BED94A7B9AD" family="Gobiidae" genus="Palutrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Palutrus Smith 1959:208" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Palutrus</taxonomicName>
and
<taxonomicName id="F44627070B4D489B699AC654F1B7CF21" LSID-ZBK="urn:lsid:zoobank.org:act:2B81E71A-6677-4E07-9CEB-8947612B603B" family="Gobiidae" genus="Coryogalops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Coryogalops Smith 1958:144" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Coryogalops</taxonomicName>
. The vertebral counts are highly consistent within these genera, except for some dwarf species of
<taxonomicName id="C88C4968BD4D9CE092946446347D7A96" LSID-ZBK="urn:lsid:zoobank.org:act:97FDC767-1FD7-483E-8184-8BED94A7B9AD" family="Gobiidae" genus="Palutrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Palutrus Smith 1959:208" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Palutrus</taxonomicName>
and
<taxonomicName id="FB8EAC06E7BBE1F9982BCA8E25FC64A6" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
which have 26 vertebrae and some species of
<taxonomicName id="D033286C1780A6E9E81DE13A488275B3" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
, known from high altitudes of greater than 500m from New Guinea, which have 28 or 29 vertebrae. All of these also share a lobed mental frenum (highly modified and sometimes absent in
<taxonomicName id="A3B94F04BFC03B8E39EDB99896E41C6D" LSID-ZBK="urn:lsid:zoobank.org:act:4DC3E52C-04FA-4AEB-B7F4-9941C410D753" family="Gobiidae" genus="Glossogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glossogobius Gill 1859:46" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Glossogobius</taxonomicName>
).
</paragraph>
<paragraph id="8FD2BEC7ED494C8B77A5D8E5F7B3CADB" pageNumber="2">
All three species described here share a number of characters not commonly found in
<taxonomicName id="FE5A012CE1D883A9A8C7CBBC30E10465" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
, particularly the flattened and elongate urogenital papilla of the males, the modified basicaudal scales (Fig. 1), the posterior nostril a simple pore or with only a slightly elevated margin anteriorly, two papillae just behind the mental frenum and the reduced transverse papilla pattern (Fig. 2). The modified urogenital papilla of males is similar to that described as the nonfimbriate condition for many species of
<taxonomicName id="DFB7835D584C55CDB059BB4E3CBC45E4" LSID-ZBK="urn:lsid:zoobank.org:act:8AC2C2C8-4045-4563-9189-6C090E61ACAB" family="Gobiidae" genus="Eviota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Eviota Jenkins 1903:501" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Eviota</taxonomicName>
(Lachner &amp; Karnella 1980), a genus which has 25 or 26 vertebrae. The modified basicaudal scale is found in a number of species of New World gobies (
<normalizedToken id="74DA85B072B66766EF1DFD61215250B5" originalValue="Böhlke">Boehlke</normalizedToken>
&amp; Robins 1968), which also have 27 vertebrae, but which have 11 precaudal vertebrae, rather than 10 as in the Indo-Pacific species. The closure of the first gill slit is rare, but also found in the eleotrid
<taxonomicName id="DDE51A61935330621AE4D806AB40A1D2" LSID-ZBK="urn:lsid:zoobank.org:act:86A153A0-8DED-4BB0-91E8-8E3D7E7B92A0" family="Thalasseleotrididae" genus="Thalasseleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Thalasseleotris Hoese &amp; Larson 1987:44" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Thalasseleotris</taxonomicName>
(Hoese &amp; Larson, 1987), which bears a remarkable overall external similarity to the present species, but has 6 branchiostegal rays. Other species of
<taxonomicName id="DE778C97A165C8A8DC11F224ACDA1DEB" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
usually have the posterior nostril with an elevated rim or at the end of a short tube, two parallel rows of papillae behind the mental frenum, although in
<taxonomicName id="8898C8A57AF30617023229B9A2BE3B8A" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="zanzibariensis">H. zanzibariensis</taxonomicName>
, the number of papillae in each row can be as few as 3. A transverse and reduced transverse papilla pattern, similar to the new
<taxonomicName id="BF4A9E93946C550560C7C8E7DECA3430" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
species, also occur in
<taxonomicName id="C1C2F7ED072DF448A52E52C39A710582" family="Gobiidae" genus="Priolepis" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Priolepis</taxonomicName>
(Winterbottom &amp; Burridge, 1992, 1993).
</paragraph>
<paragraph id="304259D08FED7DBCB6A660E5917E4B75" pageNumber="3">
Despite these differences from typical
<taxonomicName id="3B954C423E0EE45171894AC106313C79" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
, the species are included here in that genus until a more thorough phylogenetic analysis of the Indo-Pacific genera (particularly those with 27 vertebrae) can be carried out. Examination of the osteology of one species described below shows it to be typical of other small gobiids and consistent with that described for
<taxonomicName id="F0307FC7D4CB50A2CE998FC04FC550AE" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
by Hoese (1986). Randall (2005) defined
<taxonomicName id="CDFCCA411F538502C77B3D2054575D1F" LSID-ZBK="urn:lsid:zoobank.org:act:A528131D-AC68-4546-B0D0-6E94AC65DA5B" family="Gobiidae" genus="Pascua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pascua Randall 2005:20" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Pascua</taxonomicName>
based on several characters. These are:
</paragraph>
<paragraph id="1DBE5410B8C6B21C8953C8978F3A2BFB" pageNumber="3">
1 head width less than body depth (common to many gobioids and some species of
<taxonomicName id="B5248E73090DE50DC68218B5849EDAD2" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
, such as
<taxonomicName id="1A7ACC3477A47808388EB416C3AE87DF" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="tentaculata">H. tentaculata</taxonomicName>
, and therefore not useful as a generic character);
</paragraph>
<paragraph id="38FC9596153D2E7BA3265049D8041748" pageNumber="3">
2 1st gill slit closed by membrane - shared with all
<taxonomicName id="16D310803D4DE5B7E9503989CB881970" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
;
</paragraph>
<paragraph id="3DB315F7F07C031E05BB4C2E19B01B43" pageNumber="3">
3 gill opening ending at level of ventral edge of pectoral-fin base - shared with all species of
<taxonomicName id="C71CCDD4E8F308D7F72D78ED3525E98C" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
and numerous other gobiid genera;
</paragraph>
<paragraph id="3D40E9C1A99AFA01F2230CD100CD9047" pageNumber="3">
4 posterior nostril not tubular as in other
<taxonomicName id="C18FD879092E776D68873603DD3A1F06" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
;
</paragraph>
<paragraph id="29E28A4EFA41BDDDB8917E4C2B757B46" pageNumber="3">
5 a complete cephalic sensory system - complete is a misleading term, because many gobiid species have more head pores than this species and all
<taxonomicName id="05509FF09120EEC293D034616638C9DC" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
lack the posterior portion of the oculoscapular canal (above the operculum) and some species lack pores entirely;
</paragraph>
<paragraph id="B9A01C7B88964C876F49FCF532893856" pageNumber="3">
6 reduced sensory papillae on cheek - species in other genera such as
<taxonomicName id="2A69F3C5C69B53558DCD048608AF2F69" family="Gobiidae" genus="Priolepis" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Priolepis</taxonomicName>
show a similar variation from transverse to reduced transverse;
</paragraph>
<paragraph id="9ACC0DF8A1DDD76675628AF6569C3760" lastPageNumber="4" pageNumber="3">
7 dorsal fins not completely separate - common to some other species of
<taxonomicName id="BAD59962E93B91D1E0DAC1E73CAC1793" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
; in material of the species treated here the first dorsal fin connects to the base of the second dorsal fin, varying from attaching to the back just before the second dorsal fin to the basal one fifth of the spine of the second dorsal fin, which is similar to species of
<taxonomicName id="1186C39BEE17C4E37E8A36D945D1F9D7" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
, such as
<taxonomicName id="347C32DF1292E0F90D284209BC1D9B02" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="vulgaris">H. vulgaris</taxonomicName>
and
<taxonomicName id="7E3ED8B49CAA52DBA7C7331D7ABAD164" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="aporia">H. aporia</taxonomicName>
as reported by Hoese and Winterbottom, 1979.
</paragraph>
<paragraph id="3434F6D4CF40E4A05201348B5E35F24F" pageNumber="4">
8 dorsal fin pterygiophores as in
<taxonomicName id="4D356F5A89FC5A1215381EC9B43FDF9C" LSID-ZBK="urn:lsid:zoobank.org:act:E608AAC1-2634-4D90-B4F5-3F6F4BD78FBB" family="Gobiidae" genus="Bathygobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bathygobius Bleeker 1878:54" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Bathygobius</taxonomicName>
- common to most gobiines;
</paragraph>
<paragraph id="49C8B92FF11C3197400AB43E4BC9C637" pageNumber="4">
9 17 segmented caudal rays (14 branched) - 17 segmented rays is common to most gobioid fishes, we find 13-15 branched caudal rays for
<taxonomicName id="CEFF11C3DE1CB2A298D9FEFB0680D6B0" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">caudilinea</taxonomicName>
and the species described here;
</paragraph>
<paragraph id="F4F8C5011A52B2A09486C9320CB980CC" pageNumber="4">
10 pectoral rays 20 or 21 - we find 18-21 rays for
<taxonomicName id="7A05A490A45FB972E243697B04B3FB20" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">caudilinea</taxonomicName>
and 17-21 for other species treated here and Hoese (1986) reported counts of 15 to 19 pectoral rays for Indian Ocean species of
<taxonomicName id="4B770751818A6E62DD955BCDEE4B1FA9" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
;
</paragraph>
<paragraph id="1509EA25DC0376A3B76F604F8408E66E" pageNumber="4">
11 pelvic rays I,5 - common to most gobioid fishes, but some species of
<taxonomicName id="C7F40AF26F032E56677918D942920722" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
have I,4;
</paragraph>
<paragraph id="4AA516F681D4E507B6785E4B778C8ACC" pageNumber="4">
12 fins completely separate, without frenum - common to most
<taxonomicName id="C2164550F2BB86BFF2D7C698F8CE5676" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
;
</paragraph>
<paragraph id="25CFF6DB12C8DA8E205D14A36509F7D5" pageNumber="4">
13 fifth pelvic ray unbranched and clearly shorter than fourth ray - we find the fifth ray sometimes with a branch at the tip, the shorter fifth ray is common to all
<taxonomicName id="0F823443538CBF9AB3F98AD60ECC3D04" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
that have the fifth ray;
</paragraph>
<paragraph id="F759236307A911BA2CC3597FF2EB6037" pageNumber="4">
14 scales ctenoid, except a few anteriorly and on abdomen - common to numerous gobioid genera and some species of
<taxonomicName id="4DDACCE5B7541E8B1B0FA1F85B4ED078" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
;
</paragraph>
<paragraph id="2477CB26C9AFD41266B7C7890F1FE069" pageNumber="4">
15 scales 24-29 in longitudinal series - we find 23-26 for
<taxonomicName id="55C1BB5F841111D9E57307153165E403" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="caudilinea">H. caudilinea</taxonomicName>
and 22-26 for all species studied here,
<taxonomicName id="5E3D8A13A027C595926F8E842EF6865F" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
species vary from no scales to 52 in the longitudinal series;
</paragraph>
<paragraph id="56DDF3BDFA07BC55F7E98CFF5CDA6086" pageNumber="4">
16 no scales on head, nape, prepectoral area or chest - common to numerous gobiine genera and all other species of
<taxonomicName id="C222D2C7171551289C0EF4DB28081A39" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
.
</paragraph>
<paragraph id="1CD813C086E0625581E3E85CF78D9DCB" pageNumber="4">
Consequently only one of the characters reported appears to be consistent for the south Pacific species (no elevated posterior nasal tube). Evidence is presented below for other specialisations, but because of the variation of characteristics in
<taxonomicName id="638268DC5ED2360B0F640A31221427F4" LSID-ZBK="urn:lsid:zoobank.org:act:D5E53DC5-5480-4F01-9C4C-2D9BF239B442" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hetereleotris Bleeker 1874:306" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
we do not consider that the Pacific species warrant generic separation. The genus needs to be thoroughly revised to take into account the range of characters and species.
</paragraph>
</subSubSection>
</treatment>
</document>